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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<treatment ID-GBIF-Taxon="193377865" LSID="urn:lsid:plazi:treatment:E31DF58F032D5606BEE0D71033F3C7BE" httpUri="http://treatment.plazi.org/id/E31DF58F032D5606BEE0D71033F3C7BE" lastPageNumber="1" pageId="0" pageNumber="1">
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<taxonomicName LSID="E31DF58F-032D-5606-BEE0-D71033F3C7BE" authority="(Moellendorff, 1894)" authorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" authorityYear="2022" baseAuthorityName="Moellendorff" baseAuthorityYear="1894" class="Gastropoda" family="Camaenidae" genus="Bouchetcamaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bouchetcamaena platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platytropis" status="comb. nov.">
<treatment id="E31DF58F032D5606BEE0D71033F3C7BE" ID-GBIF-Taxon="193377865" LSID="urn:lsid:plazi:treatment:E31DF58F032D5606BEE0D71033F3C7BE" httpUri="http://treatment.plazi.org/id/E31DF58F032D5606BEE0D71033F3C7BE" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Camaenidae" scope_order="Stylommatophora">
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<taxonomicName id="AE5925722BD1B19FEC9673C1B76DEC01" ID-CoL="8VQ83" LSID="E31DF58F-032D-5606-BEE0-D71033F3C7BE" authority="(Moellendorff, 1894)" authorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" authorityYear="2022" baseAuthorityName="Moellendorff" baseAuthorityYear="1894" class="Gastropoda" family="Camaenidae" genus="Bouchetcamaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bouchetcamaena platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platytropis" status="comb. nov.">
Bouchetcamaena platytropis (
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
<normalizedToken id="33475292B2B2523CAC31CE717509B284" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1894)
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<taxonomicNameLabel pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="9FE7A6182FF76B1280DE8E54239365B8" pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bouchetcamaena platytropis Moellendorff, 1894 A-D anatomically examined specimen (ZMH 51934) E-H lectotype (SMF 8526). All photos: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77180.figure2" httpUri="https://binary.pensoft.net/fig/646356" pageId="0" pageNumber="1">Figures 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reproductive anatomy of Bouchetcamaena platytropis Moellendorff, 1894 (ZMH 51934) A entire genitalia B penis before cutting the weak fibres connecting proximal end of penis to distal part of epiphallus; C after removing the weak fibres. Scales represent 1 mm." figureDoi="10.3897/zookeys.1086.77180.figure3" httpUri="https://binary.pensoft.net/fig/646357" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inner wall of the penis of Bouchetcamaena platytropis Moellendorff, 1894 (ZMH 51934) A entire penis B penis-epiphallus transition." figureDoi="10.3897/zookeys.1086.77180.figure4" httpUri="https://binary.pensoft.net/fig/646358" pageId="0" pageNumber="1">, 4</figureCitation>
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<figureCitation id="B1017F2DF9ED7EEE8B386594FA728AB9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Bouchetcamaena platytropis Moellendorff, 1894 A-D anatomically examined specimen (ZMH 51934) E-H lectotype (SMF 8526). All photos: B. Pall-Gergely." figureDoi="10.3897/zookeys.1086.77180.figure2" httpUri="https://binary.pensoft.net/fig/646356" pageId="0" pageNumber="1">Figures 2</figureCitation>
<figureCitation id="D28E160EA7A1C39357EE1126CB240E21" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Reproductive anatomy of Bouchetcamaena platytropis Moellendorff, 1894 (ZMH 51934) A entire genitalia B penis before cutting the weak fibres connecting proximal end of penis to distal part of epiphallus; C after removing the weak fibres. Scales represent 1 mm." figureDoi="10.3897/zookeys.1086.77180.figure3" httpUri="https://binary.pensoft.net/fig/646357" pageId="0" pageNumber="1">, 3</figureCitation>
<figureCitation id="636EB070807B82C97214F56318BC746D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Inner wall of the penis of Bouchetcamaena platytropis Moellendorff, 1894 (ZMH 51934) A entire penis B penis-epiphallus transition." figureDoi="10.3897/zookeys.1086.77180.figure4" httpUri="https://binary.pensoft.net/fig/646358" pageId="0" pageNumber="1">, 4</figureCitation>
</paragraph>
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<taxonomicName authorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" authorityYear="2022" baseAuthorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platytropis" status="comb. nov.">Chloritis platytropis</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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<paragraph id="ADF91FE5F606699EA6D46EDE05AC2983" pageId="0" pageNumber="1">
<taxonomicName id="83FAF82D239A43D5833E36D1F1A2A4ED" authorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" authorityYear="2022" baseAuthorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platytropis" status="comb. nov.">Chloritis platytropis</taxonomicName>
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, 1894: 150.
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<taxonomicName authority="platytropis" authorityName="platytropis" authorityYear="2022" baseAuthorityName="platytropis" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis (Trichochloritis) platytropis subsp. platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="platytropis" status="comb. nov." subGenus="Trichochloritis" subSpecies="platytropis">Chloritis (Trichochloritis) platytropis platytropis</taxonomicName>
<paragraph id="BB4137365379B7879349AB1FD8F676C7" pageId="0" pageNumber="1">
<taxonomicName id="D6D32BAB40A75ABC164B027AB6451F04" authority="platytropis" authorityName="platytropis" authorityYear="2022" baseAuthorityName="platytropis" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis (Trichochloritis) platytropis subsp. platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="platytropis" status="comb. nov." subGenus="Trichochloritis" subSpecies="platytropis">Chloritis (Trichochloritis) platytropis platytropis</taxonomicName>
Zilch, 1966: 304, pl. 10, fig. 34.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="platytropis" authorityName="platytropis" authorityYear="2022" baseAuthorityName="platytropis" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis subsp. platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="platytropis" status="comb. nov." subSpecies="platytropis">Chloritis platytropis platytropis</taxonomicName>
<paragraph id="614DDACE351B2ADA0759892B1F9AF334" pageId="0" pageNumber="1">
<taxonomicName id="C4E17ADE0191B1BAC1F968E5F4A8CDA5" authority="platytropis" authorityName="platytropis" authorityYear="2022" baseAuthorityName="platytropis" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis subsp. platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="platytropis" status="comb. nov." subSpecies="platytropis">Chloritis platytropis platytropis</taxonomicName>
Maassen, 2001: 121.
</paragraph>
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<paragraph pageId="0" pageNumber="1">Types examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
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<paragraph id="FFE9A57051B1D67A6975B3DE56F82965" pageId="0" pageNumber="1">Types examined.</paragraph>
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Siam, Tschaya, coll. O.
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<normalizedToken id="815BBCDBA56037BAE8CA3812F6BBE296" originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Roebelen, SMF 8526/1 (lectotype of
<taxonomicName authorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" authorityYear="2022" baseAuthorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platytropis">
<emphasis italics="true" pageId="0" pageNumber="1">Chloritis platytropis</emphasis>
<taxonomicName id="3590E6A5A4C8B4B12FA83D6730CBC6EF" authorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" authorityYear="2022" baseAuthorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platytropis">
<emphasis id="E1DA13527A070A55DAACB429A6F4B3DC" italics="true" pageId="0" pageNumber="1">Chloritis platytropis</emphasis>
</taxonomicName>
); same data, SMF 8527/1 (paralectotype of
<taxonomicName authorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" authorityYear="2022" baseAuthorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platytropis">
<emphasis italics="true" pageId="0" pageNumber="1">Chloritis platytropis</emphasis>
<taxonomicName id="E79C300CCFA8B9D8B5556542EA7C4B21" authorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" authorityYear="2022" baseAuthorityName="Páll-Gergely &amp; Ablett &amp; Szabó &amp; Neubert" baseAuthorityYear="2022" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platytropis">
<emphasis id="01BBE905ED612CEE6C50F70711D989A1" italics="true" pageId="0" pageNumber="1">Chloritis platytropis</emphasis>
</taxonomicName>
); Golf von Siam, Insel Samui, coll. O.
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
<normalizedToken id="C8C442E5E50BD95CC1959C3885A5A3E6" originalValue="Möllendorff">Moellendorff</normalizedToken>
ex coll. Roebelen, SMF 8524/1 (lectotype of
<taxonomicName authorityName="Mollendorff" authorityYear="1894" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis subsp. var. var. samuiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="platytropis" subSpecies="var." variety="samuiana">Chloritis platytropis var. samuiana</taxonomicName>
<taxonomicName id="078EDFB1E92FCE04C2F8186501BE68C7" authorityName="Mollendorff" authorityYear="1894" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis subsp. var. var. samuiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="platytropis" subSpecies="var." variety="samuiana">Chloritis platytropis var. samuiana</taxonomicName>
); same data, SMF 8525/1 (paralectotype of
<taxonomicName authorityName="Mollendorff" authorityYear="1894" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis subsp. var. var. samuiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="platytropis" subSpecies="var." variety="samuiana">Chloritis platytropis var. samuiana</taxonomicName>
<taxonomicName id="FE3A50D29740FBE1E9729ABE17390A90" authorityName="Mollendorff" authorityYear="1894" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis platytropis subsp. var. var. samuiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="platytropis" subSpecies="var." variety="samuiana">Chloritis platytropis var. samuiana</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation accessionNumber="ZMH 51934" collectingDate="2010-08-02" collectorName="Hausdorf" country="Thailand" elevation="90" latitude="8.436389" location="Ton Prai" longLatPrecision="21" longitude="98.309166" municipality="Thai Mueang" specimenCount="1" stateProvince="Phangnga">
<collectingCountry name="Thailand">Thailand</collectingCountry>
<subSubSection id="29FE47C20F56DE084CCBE92AA69FE229" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="2830F11FA7597C88CFE51AC61D291EDB" pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph id="F3B8C613BCAF1765FCDD354E65412615" pageId="0" pageNumber="1">
<materialsCitation id="A3B74ACC7207DF53088DB071D805C06B" accessionNumber="ZMH 51934" collectingDate="2010-08-02" collectorName="Hausdorf" country="Thailand" elevation="90" latitude="8.436389" location="Ton Prai" longLatPrecision="21" longitude="98.309166" municipality="Thai Mueang" specimenCount="1" stateProvince="Phangnga">
<collectingCountry id="98D0E2D3C03CCC5A475BE600E724D6A7" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion country="Thailand" name="Phangnga">Phangnga</collectingRegion>
<collectingRegion id="43487A300378B7FCD0014C4DCF424132" country="Thailand" name="Phangnga">Phangnga</collectingRegion>
:
<collectingMunicipality>Thai Mueang</collectingMunicipality>
<collectingMunicipality id="10BE9A7F91FF2C4340E9F6265527A0B1">Thai Mueang</collectingMunicipality>
:
<location LSID="urn:lsid:plazi:treatment:E31DF58F032D5606BEE0D71033F3C7BE:B99150993057A9CE8882B84E8F4476C9" country="Thailand" latitude="8.436389" longLatPrecision="21" longitude="98.309166" municipality="Thai Mueang" name="Ton Prai" stateProvince="Phangnga">Ton Prai</location>
<location id="B99150993057A9CE8882B84E8F4476C9" LSID="urn:lsid:plazi:treatment:E31DF58F032D5606BEE0D71033F3C7BE:B99150993057A9CE8882B84E8F4476C9" country="Thailand" latitude="8.436389" longLatPrecision="21" longitude="98.309166" municipality="Thai Mueang" name="Ton Prai" stateProvince="Phangnga">Ton Prai</location>
waterfall,
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">90 m</elevation>
<quantity id="08AB9A3781C6AED8074D61BC84F27BC3" metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">
<elevation id="ED2243A346C0593BB474A132F972EB97" metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">90 m</elevation>
</quantity>
,
<geoCoordinate degrees="08" direction="north" minutes="26" orientation="latitude" precision="15" seconds="11" value="8.436389">08°26'11&quot;N</geoCoordinate>
<geoCoordinate id="298D0DBC6CDD7EEAD1669A30CC7CA882" degrees="08" direction="north" minutes="26" orientation="latitude" precision="15" seconds="11" value="8.436389">08°26'11&quot;N</geoCoordinate>
,
<geoCoordinate degrees="098" direction="east" minutes="18" orientation="longitude" precision="15" seconds="33" value="98.309166">098°18'33&quot;E</geoCoordinate>
<geoCoordinate id="6B9EDB697F51D0C902D4A10988FF751E" degrees="098" direction="east" minutes="18" orientation="longitude" precision="15" seconds="33" value="98.309166">098°18'33&quot;E</geoCoordinate>
, leg.
<collectorName>Hausdorf</collectorName>
<collectorName id="D797EF6E878F8E0A0F5313BB3008854A">Hausdorf</collectorName>
,
<collectingDate value="2010-08-02">02.08.2010</collectingDate>
<collectingDate id="CF75E377F1D3F701706A920218F08C75" value="2010-08-02">02.08.2010</collectingDate>
,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH51934">ZMH 51934</accessionNumber>
<accessionNumber id="AE290E84FA2AB34E973CB83C4FE665A0" httpUri="http://www.ncbi.nlm.nih.gov/protein/ZMH51934">ZMH 51934</accessionNumber>
(1 dry shell + ethanol-preserved body)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description of the genitalia.</paragraph>
<paragraph pageId="0" pageNumber="1">Penis long, cylindrical, with swollen proximal part connected to adjacent epiphallic area by weak fibres; internally with ca. 6 wide, longitudinal, low folds; no penial verge present, although the folds form a circular ring with slightly elongated margin; epiphallus slightly longer than penis; retractor muscle slender, inserting on the distal end of epiphallus near its joint with penis; flagellum long, slender, pointed; no penial caecum present; vagina shorter and thicker than penis; spermoviduct elongated, stalk of bursa copulatrix very long, with some central swelling, bursa small, rounded; albumen gland long, banana-shaped, talon small.</paragraph>
<subSubSection id="9BF1E7173C242C1004A33B0613F5A2A6" pageId="0" pageNumber="1" type="description">
<paragraph id="92191329540976FF02ABA7295134DF13" pageId="0" pageNumber="1">Description of the genitalia.</paragraph>
<paragraph id="8D23D8808F9347D3D2496AA798771FBC" pageId="0" pageNumber="1">Penis long, cylindrical, with swollen proximal part connected to adjacent epiphallic area by weak fibres; internally with ca. 6 wide, longitudinal, low folds; no penial verge present, although the folds form a circular ring with slightly elongated margin; epiphallus slightly longer than penis; retractor muscle slender, inserting on the distal end of epiphallus near its joint with penis; flagellum long, slender, pointed; no penial caecum present; vagina shorter and thicker than penis; spermoviduct elongated, stalk of bursa copulatrix very long, with some central swelling, bursa small, rounded; albumen gland long, banana-shaped, talon small.</paragraph>
</subSubSection>
</treatment>
</document>

124
data/E3/21/0D/E3210D8091C554569636F9F6B82A3631.xml
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@ -1,97 +1,97 @@
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<taxonomicName id="44D46A74A03441CCF0D86C32CF093CC7" class="Sordariomycetes" family="Phialemoniopsidaceae" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="" order="Thyridiales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="family">Phialemoniopsidaceae</taxonomicName>
K.D. Hyde &amp; Hongsanan, [as
<taxonomicName class="Sordariomycetes" family="Phialemoniopsaceae" higherTaxonomySource="treatment-meta" kingdom="Fungi" lsidName="" order="Thyridiales" pageId="0" pageNumber="147" rank="family">Phialemoniopsaceae</taxonomicName>
<taxonomicName id="27E1999FB0187296B78431AD33305476" class="Sordariomycetes" family="Phialemoniopsaceae" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="" order="Thyridiales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="family">Phialemoniopsaceae</taxonomicName>
] Fungal Divers. 107: 95 (2021).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="type genus">
<paragraph pageId="0" pageNumber="147">Type genus.</paragraph>
<paragraph pageId="0" pageNumber="147">
<taxonomicName authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Thyridiaceae" genus="Thyridium" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Thyridium" order="Thyridiales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Thyridium</emphasis>
<subSubSection id="2FA0AC5C1A76AA9C241BBC749E77A315" pageId="0" pageNumber="147" type="type genus">
<paragraph id="D044318D1E4FD0CB3248293A5749B21A" pageId="0" pageNumber="147">Type genus.</paragraph>
<paragraph id="8BA2D78E7A29CA16FE9218FE5FB764B1" pageId="0" pageNumber="147">
<taxonomicName id="3209CFCA9159F35EA9B3896EE0270F75" authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Thyridiaceae" genus="Thyridium" kingdom="Fungi" lsidName="Thyridium" order="Thyridiales" pageId="0" pageNumber="147" rank="genus">
<emphasis id="8719CA96CD2DF28AF362F9CEAFD37363" italics="true" pageId="0" pageNumber="147">Thyridium</emphasis>
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Nitschke, Pyrenomyc. Germ. 1: 110 (1867).
</paragraph>
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<subSubSection pageId="0" pageNumber="147" type="notes">
<paragraph pageId="0" pageNumber="147">Notes.</paragraph>
<paragraph pageId="0" pageNumber="147">
<taxonomicName authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Phialemoniopsidaceae" kingdom="Fungi" lsidName="" order="Thyridiales" pageId="0" pageNumber="147" rank="family">Phialemoniopsidaceae</taxonomicName>
<subSubSection id="160318B21D45C28D33AF5AFD38B90893" pageId="0" pageNumber="147" type="notes">
<paragraph id="3F9D65FCD1972A7E46E891644AEE4BEE" pageId="0" pageNumber="147">Notes.</paragraph>
<paragraph id="211F276B81A1E044F3D72BBE1337B3AB" pageId="0" pageNumber="147">
<taxonomicName id="88B3B28D0CDCB0E6AFA70158A7AB7312" authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Phialemoniopsidaceae" kingdom="Fungi" lsidName="" order="Thyridiales" pageId="0" pageNumber="147" rank="family">Phialemoniopsidaceae</taxonomicName>
is considered a synonym of
<taxonomicName authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Thyridiaceae" kingdom="Fungi" lsidName="" order="Thyridiales" pageId="0" pageNumber="147" rank="family">Thyridiaceae</taxonomicName>
<taxonomicName id="7F62A1039188370FFF73ADF9BB8117A9" authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Thyridiaceae" kingdom="Fungi" lsidName="" order="Thyridiales" pageId="0" pageNumber="147" rank="family">Thyridiaceae</taxonomicName>
because
<taxonomicName class="Sordariomycetes" family="Thyridiaceae" genus="Phialemoniopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phialemoniopsis" order="Boliniales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Phialemoniopsis</emphasis>
<taxonomicName id="E5782DE70B671BCA2F555832D1E0C950" class="Sordariomycetes" family="Thyridiaceae" genus="Phialemoniopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phialemoniopsis" order="Boliniales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="genus">
<emphasis id="2A458AF57B08D644E652F967CD8E90AE" italics="true" pageId="0" pageNumber="147">Phialemoniopsis</emphasis>
</taxonomicName>
, the type genus of
<taxonomicName class="Sordariomycetes" family="Phialemoniopsidaceae" kingdom="Fungi" lsidName="" order="Boliniales" pageId="0" pageNumber="147" rank="family">Phialemoniopsidaceae</taxonomicName>
<taxonomicName id="F8EA6B41BCEA6E81EBCDC292B50C8910" class="Sordariomycetes" family="Phialemoniopsidaceae" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="" order="Boliniales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="family">Phialemoniopsidaceae</taxonomicName>
, was revealed congeneric with
<taxonomicName authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Thyridiaceae" genus="Thyridium" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Thyridium" order="Thyridiales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Thyridium</emphasis>
<taxonomicName id="1958498FE1F5C37C710DA7F2E1DC9042" authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Thyridiaceae" genus="Thyridium" kingdom="Fungi" lsidName="Thyridium" order="Thyridiales" pageId="0" pageNumber="147" rank="genus">
<emphasis id="1223F1AE02C82F8B61DD4512F49D219F" italics="true" pageId="0" pageNumber="147">Thyridium</emphasis>
</taxonomicName>
and is placed in the synonymy of the latter genus in this study. The type genera of both families, that is,
<taxonomicName authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Thyridiaceae" genus="Thyridium" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Thyridium" order="Thyridiales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Thyridium</emphasis>
<taxonomicName id="51E12606D4CA0B4B87F4E92B5C27C2D1" authorityName="Nitschke, Pyrenomyc. Germ. 1: 110" authorityYear="1867" class="Ascomycetes" family="Thyridiaceae" genus="Thyridium" kingdom="Fungi" lsidName="Thyridium" order="Thyridiales" pageId="0" pageNumber="147" rank="genus">
<emphasis id="CA8AC52B72C072F698ED859FBD39C3DF" italics="true" pageId="0" pageNumber="147">Thyridium</emphasis>
</taxonomicName>
and
<taxonomicName class="Sordariomycetes" family="Thyridiaceae" genus="Phialemoniopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phialemoniopsis" order="Boliniales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Phialemoniopsis</emphasis>
<taxonomicName id="A7CFA49711D421ABB67EDA1694622D7F" class="Sordariomycetes" family="Thyridiaceae" genus="Phialemoniopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phialemoniopsis" order="Boliniales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="genus">
<emphasis id="FA982128778BE07A722A2216B844745F" italics="true" pageId="0" pageNumber="147">Phialemoniopsis</emphasis>
</taxonomicName>
, share many morphological features in their asexual states, as noted below.
</paragraph>

220
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<figureCitation captionStart="Figures 18, 19" captionStartId="F8" captionText="Figures 18, 19. 18 Gibbidessus kangarooensis sp. nov. 19 G. pederzanii sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures18-19" httpUri="https://binary.pensoft.net/fig/462667" pageId="0" pageNumber="11">, 19</figureCitation>
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<taxonomicNameLabel id="96BDC27FBAA26F05D4A8464C5549A755" pageId="0" pageNumber="11">sp. nov.</taxonomicNameLabel>
<figureCitation id="09A7B3CE82E6169E770025EE72C41546" captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Habitus of 5 Gibbidessus drikdrikensis sp. nov. 6 G. kangarooensis sp. nov. 7 G. pederzanii sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures5-7" httpUri="https://binary.pensoft.net/fig/462661" pageId="0" pageNumber="11">Figs 7</figureCitation>
<figureCitation id="EEDEF9F8A3ACB7EE3497FCA4CADBBC47" captionStart="Figure 12" captionStartId="F4" captionText="Figure 12. Head of G. davidi sp. nov. A with cervical line (red arrow), and head of G. pederzanii sp. nov. B without cervical line but rather a few punctures instead (red arrow)." figureDoi="10.3897/zookeys.975.55456.figure12" httpUri="https://binary.pensoft.net/fig/462663" pageId="0" pageNumber="11">, 12</figureCitation>
<figureCitation id="2E0A25EE82B4F18C4EFDD4F0D527206A" captionStart="Figures 18, 19" captionStartId="F8" captionText="Figures 18, 19. 18 Gibbidessus kangarooensis sp. nov. 19 G. pederzanii sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures18-19" httpUri="https://binary.pensoft.net/fig/462667" pageId="0" pageNumber="11">, 19</figureCitation>
<figureCitation id="CF1D572241541695E60CF338FF633649" captionStart="Figures 24, 25" captionStartId="F11" captionText="Figures 24, 25. Distribution of 24 Gibbidessus pictipes (black dots) 25 G. pederzanii sp. nov. (red dot), G. rottnestensis sp. nov. (blue dots)." figureDoi="10.3897/zookeys.975.55456.figures24-25" httpUri="https://binary.pensoft.net/fig/462670" pageId="0" pageNumber="11">, 25</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="type locality">
<paragraph pageId="0" pageNumber="11">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="11">
<subSubSection id="40B44D987464C0345665FB0B7B2DB244" pageId="0" pageNumber="11" type="type locality">
<paragraph id="050F7C711A6D2EDBA0D0C3ECB3FCC90C" pageId="0" pageNumber="11">Type locality.</paragraph>
<paragraph id="45CD375FB2091B23FACCE5C93C56B816" pageId="0" pageNumber="11">
Australia, Western Australia, creek around Nannup [
<geoCoordinate degrees="33" direction="south" minutes="58" orientation="latitude" precision="925" value="-33.966667">33°58'S</geoCoordinate>
<geoCoordinate id="E3C46908040B172FAA8B13B49435C7B9" degrees="33" direction="south" minutes="58" orientation="latitude" precision="925" value="-33.966667">33°58'S</geoCoordinate>
,
<geoCoordinate degrees="115" direction="east" minutes="45" orientation="longitude" precision="925" value="115.75">115°45'E</geoCoordinate>
<geoCoordinate id="736958FF47B1AC6EE55AB7EBC3662258" degrees="115" direction="east" minutes="45" orientation="longitude" precision="925" value="115.75">115°45'E</geoCoordinate>
].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="type material">
<paragraph pageId="0" pageNumber="11">Type material.</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Holotype</emphasis>
<subSubSection id="2AC1CBAEDE07349100B1427CEDEC8D68" pageId="0" pageNumber="11" type="type material">
<paragraph id="168B510A2D9F33F5E228A61300372A96" pageId="0" pageNumber="11">Type material.</paragraph>
<paragraph id="2A32171721ED9F39DF2249B3CA0BB25A" pageId="0" pageNumber="11">
<emphasis id="8D28003FDFE71E93D2EF824813FC7E5A" bold="true" pageId="0" pageNumber="11">
<emphasis id="4545863A1B54C4F0A3190395E0E7F8EA" italics="true" pageId="0" pageNumber="11">Holotype</emphasis>
:
</emphasis>
Male, &quot;Australia (WA) Nannup env. roadside creeks 1/12/98 Pederzani&quot;, &quot;Holotype
<taxonomicName authorityName="Hendrich &amp; Watts &amp; Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus pederzanii" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="pederzanii">
<emphasis italics="true" pageId="0" pageNumber="11">Gibbidessus pederzanii</emphasis>
<taxonomicName id="F3592239FFC261E38B6A81FCAD58CE28" authorityName="Hendrich &amp; Watts &amp; Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Gibbidessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbidessus pederzanii" order="Coleoptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="pederzanii">
<emphasis id="4A7DB2763A0340C5744D0E0708EEF8A2" italics="true" pageId="0" pageNumber="11">Gibbidessus pederzanii</emphasis>
</taxonomicName>
Hendrich, Watts &amp; Balke des. 2020&quot; [red printed label] (SAMA).
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Paratypes</emphasis>
<emphasis id="EF5BA2623932AD930FE0C2FE240D413C" bold="true" pageId="0" pageNumber="11">
<emphasis id="18212AC937E07A8139F88200E1C4C36B" italics="true" pageId="0" pageNumber="11">Paratypes</emphasis>
(13 exs.):
</emphasis>
All specimens with same data as holotype. Two specimens with &quot; SAMA Database No 25-001593&quot; and one with a yellow printed label
<normalizedToken originalValue="“photographed”">&quot;photographed&quot;</normalizedToken>
<normalizedToken id="1C4A2885B0F7A06A41C4A567F1610AB7" originalValue="“photographed”">&quot;photographed&quot;</normalizedToken>
(CFP, CLH, SAMA, ZSM). All paratypes are provided with printed red paratype labels.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="diagnosis">
<paragraph pageId="0" pageNumber="11">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="11">
<subSubSection id="C39E1CB618189DB3E0498B3AE69FE331" pageId="0" pageNumber="11" type="diagnosis">
<paragraph id="4789A483A77B058C9FCDD8CB2F53835D" pageId="0" pageNumber="11">Diagnosis.</paragraph>
<paragraph id="F20C2EF0F291F47031B6CB88B2183261" pageId="0" pageNumber="11">
Medium-sized species which externally is characterised by a rounded habitus, without disruption between pronotum and elytron, and shiny, non-microreticulate dorsal surface with testaceous markings on elytra. Dorsoventrally rather arched. Without cervical line but rather a few punctures instead (Fig.
<figureCitation captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Habitus of 5 Gibbidessus drikdrikensis sp. nov. 6 G. kangarooensis sp. nov. 7 G. pederzanii sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures5-7" httpUri="https://binary.pensoft.net/fig/462661" pageId="0" pageNumber="11">7</figureCitation>
<figureCitation id="E785326712D30E073D911B8B2D9937F8" captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Habitus of 5 Gibbidessus drikdrikensis sp. nov. 6 G. kangarooensis sp. nov. 7 G. pederzanii sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures5-7" httpUri="https://binary.pensoft.net/fig/462661" pageId="0" pageNumber="11">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="description">
<paragraph pageId="0" pageNumber="11">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="11">Holotype: TL = 1.5 mm, TL-H = 1.3 mm, MW = 0.85 mm. Paratypes: TL = 1.5-1.6 mm, TL-H = 1.3-1.4 mm, MW = 0.85-0.95 mm.</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Head</emphasis>
<subSubSection id="618B089E7043FA16B535317465A9C540" pageId="0" pageNumber="11" type="description">
<paragraph id="FFDC996D3CF3878393407ED249311073" pageId="0" pageNumber="11">Measurements.</paragraph>
<paragraph id="8E06FD2C8D3C41446B1DE5AB9D3F60AF" pageId="0" pageNumber="11">Holotype: TL = 1.5 mm, TL-H = 1.3 mm, MW = 0.85 mm. Paratypes: TL = 1.5-1.6 mm, TL-H = 1.3-1.4 mm, MW = 0.85-0.95 mm.</paragraph>
<paragraph id="5B030A055381164857E8A02CB13C7176" pageId="0" pageNumber="11">
<emphasis id="0214F4FBB2D10CA37D9062DF50A420C6" bold="true" pageId="0" pageNumber="11">
<emphasis id="4E8DCC998FB5F7877E21A50B9EC70E02" italics="true" pageId="0" pageNumber="11">Head</emphasis>
:
</emphasis>
Ferruginous, around eyes almost black, without cervical line but rather a few punctures instead (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F4" captionText="Figure 12. Head of G. davidi sp. nov. A with cervical line (red arrow), and head of G. pederzanii sp. nov. B without cervical line but rather a few punctures instead (red arrow)." figureDoi="10.3897/zookeys.975.55456.figure12" httpUri="https://binary.pensoft.net/fig/462663" pageId="0" pageNumber="11">12B</figureCitation>
<figureCitation id="B673988F02FECED2FB1A1402C970AB0F" captionStart="Figure 12" captionStartId="F4" captionText="Figure 12. Head of G. davidi sp. nov. A with cervical line (red arrow), and head of G. pederzanii sp. nov. B without cervical line but rather a few punctures instead (red arrow)." figureDoi="10.3897/zookeys.975.55456.figure12" httpUri="https://binary.pensoft.net/fig/462663" pageId="0" pageNumber="11">12B</figureCitation>
). Coarsely punctate, rather shiny, weak microreticulation visible. Punctures weakly anteriorly and strongly posteriorly between eyes. Antennae relatively short, stout. Antennomeres 1 and 2 ferruginous, 3-11 darkened anteriorly.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Pronotum</emphasis>
<paragraph id="C1FE56A55665963F791F3791877A4447" pageId="0" pageNumber="11">
<emphasis id="9877E2D47F4EB981C28236D628B2AF3F" bold="true" pageId="0" pageNumber="11">
<emphasis id="D5041565A00761D3336F7E269487A373" italics="true" pageId="0" pageNumber="11">Pronotum</emphasis>
:
</emphasis>
Ferruginous, anterior and posterior margins, between striae, slightly darker, broadest at posterior corners. Punctation very weak almost evenly distributed, shiny and microsculpture absent. Sides of pronotum margined and almost evenly rounded. Angle between pronotum and elytra less pronounced, basal pronotal plicae present. Striae moderately defined, almost 1/2 length of pronotum, slightly incurved.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Elytra</emphasis>
<paragraph id="B84F0770F44390C630AEBFD0F5673B5A" pageId="0" pageNumber="11">
<emphasis id="C61A7B11B1BEBB5968506AE4F6DDBF73" bold="true" pageId="0" pageNumber="11">
<emphasis id="E039DA30D24137A361B53C4C6CAFA84F" italics="true" pageId="0" pageNumber="11">Elytra</emphasis>
:
</emphasis>
Dark brown with vague basolateral and apical area ferruginous (Fig.
<figureCitation captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Habitus of 5 Gibbidessus drikdrikensis sp. nov. 6 G. kangarooensis sp. nov. 7 G. pederzanii sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures5-7" httpUri="https://binary.pensoft.net/fig/462661" pageId="0" pageNumber="11">7</figureCitation>
<figureCitation id="FA7F1BB5706B0E9950BBFFDAF1A6B7A9" captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Habitus of 5 Gibbidessus drikdrikensis sp. nov. 6 G. kangarooensis sp. nov. 7 G. pederzanii sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures5-7" httpUri="https://binary.pensoft.net/fig/462661" pageId="0" pageNumber="11">7</figureCitation>
). Coarsely and densely punctate, shiny, microsculpture absent. Striae deeply impressed, straight but shorter than basal pronotal striae.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Ventral side</emphasis>
<paragraph id="958F24835D05AABF1615F01D8CC52408" pageId="0" pageNumber="11">
<emphasis id="48A5E695D6BF6BC1D5066520B3FD1592" bold="true" pageId="0" pageNumber="11">
<emphasis id="C4C6960EDA975E548D0BAB6613C11DED" italics="true" pageId="0" pageNumber="11">Ventral side</emphasis>
:
</emphasis>
Ferruginous. Prothorax and abdomen paler than other parts. Metacoxae and metaventrite covered with larger punctures, surface shiny, without microreticulation. Abdominal ventrites with dense and finer punctures, shiny, microreticulation absent. Metacoxal lines almost straight, anteriorly not divergent. Epipleuron testaceous, with few coarse punctures, shiny, lacking microsculpture. Legs ferruginous with meta- and mesotarsi somewhat darkened.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">
<emphasis italics="true" pageId="0" pageNumber="11">Male</emphasis>
<paragraph id="E5F3C92B575336CEB42EEDC05AF4E869" pageId="0" pageNumber="11">
<emphasis id="99413D52AAC43456E50C15D2EB1A1AD9" bold="true" pageId="0" pageNumber="11">
<emphasis id="98F648B933C3C1A851C66870B4D242B5" italics="true" pageId="0" pageNumber="11">Male</emphasis>
.
</emphasis>
Dorsal surface with coarse punctures but otherwise with shiny surface (Fig.
<figureCitation captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Habitus of 5 Gibbidessus drikdrikensis sp. nov. 6 G. kangarooensis sp. nov. 7 G. pederzanii sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures5-7" httpUri="https://binary.pensoft.net/fig/462661" pageId="0" pageNumber="11">7</figureCitation>
<figureCitation id="4215B562701C7FC1D3696C16B8EBB589" captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Habitus of 5 Gibbidessus drikdrikensis sp. nov. 6 G. kangarooensis sp. nov. 7 G. pederzanii sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures5-7" httpUri="https://binary.pensoft.net/fig/462661" pageId="0" pageNumber="11">7</figureCitation>
). Median lobe of aedeagus as in Fig.
<figureCitation captionStart="Figures 18, 19" captionStartId="F8" captionText="Figures 18, 19. 18 Gibbidessus kangarooensis sp. nov. 19 G. pederzanii sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures18-19" httpUri="https://binary.pensoft.net/fig/462667" pageId="0" pageNumber="11">19A, B</figureCitation>
<figureCitation id="10F0DF6511D0E5F11B8859299EA57ECB" captionStart="Figures 18, 19" captionStartId="F8" captionText="Figures 18, 19. 18 Gibbidessus kangarooensis sp. nov. 19 G. pederzanii sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures18-19" httpUri="https://binary.pensoft.net/fig/462667" pageId="0" pageNumber="11">19A, B</figureCitation>
. Shape of median lobe, almost straight and fairly uniform in lateral view, in ventral view rounded at apex. Parameres bi-segmented, elongated, and without setae at apex (Fig.
<figureCitation captionStart="Figures 18, 19" captionStartId="F8" captionText="Figures 18, 19. 18 Gibbidessus kangarooensis sp. nov. 19 G. pederzanii sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures18-19" httpUri="https://binary.pensoft.net/fig/462667" pageId="0" pageNumber="11">19C, D</figureCitation>
<figureCitation id="A4AC75B7B374CA2BC410ECD08FFED07B" captionStart="Figures 18, 19" captionStartId="F8" captionText="Figures 18, 19. 18 Gibbidessus kangarooensis sp. nov. 19 G. pederzanii sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures18-19" httpUri="https://binary.pensoft.net/fig/462667" pageId="0" pageNumber="11">19C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="affinities">
<paragraph pageId="0" pageNumber="11">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="11">
<subSubSection id="83CF29FB5781F38DB78EF8851506EF50" pageId="0" pageNumber="11" type="affinities">
<paragraph id="3E48F10BAAEA507FB27671B2F79E3E3E" pageId="0" pageNumber="11">Affinities.</paragraph>
<paragraph id="F3833332174988D76C2615FC512DE09D" pageId="0" pageNumber="11">
This species is similar to
<taxonomicName lsidName="G. davidi" pageId="0" pageNumber="11" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="11">G. davidi</emphasis>
<taxonomicName id="BBBD698C9538298E725639F18B461026" lsidName="G. davidi" pageId="0" pageNumber="11" rank="species" species="davidi">
<emphasis id="B57470550D706BEE4CA1F20AC421653C" italics="true" pageId="0" pageNumber="11">G. davidi</emphasis>
</taxonomicName>
sp. nov. but readily separated by the different colour pattern, the more roundish body (Figs
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">4</figureCitation>
<figureCitation id="24BCBC15486D27695A85EEA32F5C7AAA" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus of 1 Gibbidessus atomus sp. nov. 2 G. chipi, 3 G. davidi, male 4 G. davidi female. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures1-4" httpUri="https://binary.pensoft.net/fig/462660" pageId="0" pageNumber="11">4</figureCitation>
,
<figureCitation captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Habitus of 5 Gibbidessus drikdrikensis sp. nov. 6 G. kangarooensis sp. nov. 7 G. pederzanii sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures5-7" httpUri="https://binary.pensoft.net/fig/462661" pageId="0" pageNumber="11">7</figureCitation>
<figureCitation id="1A802004A9E9BFA836960684EF97C333" captionStart="Figures 5–7" captionStartId="F2" captionText="Figures 5 - 7. Habitus of 5 Gibbidessus drikdrikensis sp. nov. 6 G. kangarooensis sp. nov. 7 G. pederzanii sp. nov. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.975.55456.figures5-7" httpUri="https://binary.pensoft.net/fig/462661" pageId="0" pageNumber="11">7</figureCitation>
), the larger punctation on elytra, and the form of the median lobe and parameres (Figs
<figureCitation captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. 16 Gibbidessus davidi sp. nov. 17 G. drikdrikensis sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view, D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures16-17" httpUri="https://binary.pensoft.net/fig/462666" pageId="0" pageNumber="11">16</figureCitation>
<figureCitation id="C784895525F9869C048ACAA933F7BF8C" captionStart="Figures 16, 17" captionStartId="F7" captionText="Figures 16, 17. 16 Gibbidessus davidi sp. nov. 17 G. drikdrikensis sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view, D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures16-17" httpUri="https://binary.pensoft.net/fig/462666" pageId="0" pageNumber="11">16</figureCitation>
,
<figureCitation captionStart="Figures 18, 19" captionStartId="F8" captionText="Figures 18, 19. 18 Gibbidessus kangarooensis sp. nov. 19 G. pederzanii sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures18-19" httpUri="https://binary.pensoft.net/fig/462667" pageId="0" pageNumber="11">19</figureCitation>
<figureCitation id="6E4E2D1F418A57A773A41081179840E1" captionStart="Figures 18, 19" captionStartId="F8" captionText="Figures 18, 19. 18 Gibbidessus kangarooensis sp. nov. 19 G. pederzanii sp. nov. A median lobe in ventral view B median lobe in lateral view, left side C left paramere in lateral view D right paramere in lateral view. Scale bar: 0.2 mm." figureDoi="10.3897/zookeys.975.55456.figures18-19" httpUri="https://binary.pensoft.net/fig/462667" pageId="0" pageNumber="11">19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="etymology">
<paragraph pageId="0" pageNumber="11">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="11">The species is named after our colleague, the dytiscid specialist Fernando Pederzani (Ravenna, Italy), who collected the type material. The specific epithet is a substantive in the genitive case.</paragraph>
<subSubSection id="1F47869A876C108AA4061801DFC16247" pageId="0" pageNumber="11" type="etymology">
<paragraph id="3B666722E282750C9FC236A152A3E992" pageId="0" pageNumber="11">Etymology.</paragraph>
<paragraph id="F85E051FF8F71B1ABA7A1C1A59CA8410" pageId="0" pageNumber="11">The species is named after our colleague, the dytiscid specialist Fernando Pederzani (Ravenna, Italy), who collected the type material. The specific epithet is a substantive in the genitive case.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="distribution">
<paragraph pageId="0" pageNumber="11">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="11">
<subSubSection id="57FEF7E977FBBF008152449124E76D47" pageId="0" pageNumber="11" type="distribution">
<paragraph id="D69D13C2FBA9066D9B3944F041B4E92A" pageId="0" pageNumber="11">Distribution.</paragraph>
<paragraph id="EEB186804312085F37D39F9C2612FE6A" pageId="0" pageNumber="11">
South-western Australia. A rare species, which is only known from the type locality somewhere around Nannup in south-western Australia. Most probably a more inland species and restricted to forested areas and not in heathland or coastal sedge swamps (Fig.
<figureCitation captionStart="Figures 24, 25" captionStartId="F11" captionText="Figures 24, 25. Distribution of 24 Gibbidessus pictipes (black dots) 25 G. pederzanii sp. nov. (red dot), G. rottnestensis sp. nov. (blue dots)." figureDoi="10.3897/zookeys.975.55456.figures24-25" httpUri="https://binary.pensoft.net/fig/462670" pageId="0" pageNumber="11">25</figureCitation>
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</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera brevirostris" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="brevirostris">Phalacrocera brevirostris</taxonomicName>
<paragraph id="51C1C2936E76147405C32E2A193621FF" pageId="0" pageNumber="13">
<taxonomicName id="23B3D6F9CBAE184C7D523747277D824D" baseAuthorityName="Zetterstedt" baseAuthorityYear="1838" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera brevirostris" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="brevirostris">Phalacrocera brevirostris</taxonomicName>
(Zetterstedt, 1838)
</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName baseAuthorityName="Walker" baseAuthorityYear="1856" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera diversa" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="diversa">Phalacrocera diversa</taxonomicName>
<paragraph id="8E0D702CE3EC58C4A908ED3BE608121D" pageId="0" pageNumber="13">
<taxonomicName id="8AFDEAA43641D4B5A70CD357CB004113" baseAuthorityName="Walker" baseAuthorityYear="1856" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera diversa" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="diversa">Phalacrocera diversa</taxonomicName>
(Walker, 1856)
</paragraph>
<paragraph pageId="0" pageNumber="13">
<taxonomicName authorityName="Alexander" authorityYear="1914" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera neoxena" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="neoxena">Phalacrocera neoxena</taxonomicName>
<paragraph id="36D11DD015630177FF291ADCB93BCCBE" pageId="0" pageNumber="13">
<taxonomicName id="58B81E181FD68D9DC1CAE392FD359B21" authorityName="Alexander" authorityYear="1914" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera neoxena" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="neoxena">Phalacrocera neoxena</taxonomicName>
Alexander, 1914.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="type material">
<paragraph pageId="0" pageNumber="13">Non-type material examined.</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Canada</emphasis>
<subSubSection id="F6A0D3B50C815B6545E8393E9E589A4A" pageId="0" pageNumber="13" type="type material">
<paragraph id="F5C9694EA169EA6A9F71F1FBD2FEAD6F" pageId="0" pageNumber="13">Non-type material examined.</paragraph>
<paragraph id="18FA4F4245A215A87B9FFD9B3363FE26" pageId="0" pageNumber="13">
<emphasis id="783C193CAA37FC6E286C1A728F0529B3" bold="true" pageId="0" pageNumber="13">Canada</emphasis>
• 2 ♂, 1 ♀; Ontario, Ottawa, Stony Swamp;
<geoCoordinate degrees="45.3" direction="north" orientation="latitude" precision="5555" value="45.3">45.3°N</geoCoordinate>
<geoCoordinate id="B8F83C79009C0CEEDFAE1FD900F632E2" degrees="45.3" direction="north" orientation="latitude" precision="5555" value="45.3">45.3°N</geoCoordinate>
,
<geoCoordinate degrees="75.83" direction="west" orientation="longitude" precision="555" value="-75.83">75.83° W</geoCoordinate>
<geoCoordinate id="E2F2D7E3B8920DBBC7E5C0389A66DE40" degrees="75.83" direction="west" orientation="longitude" precision="555" value="-75.83">75.83° W</geoCoordinate>
; alt. 115 m; 10 May. 2017; F. Brodo leg.; CKLP.
<emphasis bold="true" pageId="0" pageNumber="13">Finland</emphasis>
<emphasis id="9B2B7A078DB56B322870C8EF80BFFA20" bold="true" pageId="0" pageNumber="13">Finland</emphasis>
• 2 ♂, 1 ♀; Kirkkonummi, Stormossen.
<geoCoordinate degrees="60.07901" direction="north" orientation="latitude" precision="1" value="60.07901">60.07901°N</geoCoordinate>
<geoCoordinate id="392271A15E5239BA3471C01E116F3F60" degrees="60.07901" direction="north" orientation="latitude" precision="1" value="60.07901">60.07901°N</geoCoordinate>
,
<geoCoordinate degrees="24.57980" direction="east" orientation="longitude" precision="1" value="24.5798">24.57980°E</geoCoordinate>
<geoCoordinate id="0F55EAF1FC8BF36B95486319EF7C4F4E" degrees="24.57980" direction="east" orientation="longitude" precision="1" value="24.5798">24.57980°E</geoCoordinate>
; alt. 7 m; 2 Jun. 2016; E. Viitanen leg.; CKLP. • 1 ♂, 2 ♀; Kaarina, Jarvela;
<geoCoordinate degrees="60.46157" direction="north" orientation="latitude" precision="1" value="60.46157">60.46157°N</geoCoordinate>
<geoCoordinate id="92412F1CE7A296828A46E90E9B366A7F" degrees="60.46157" direction="north" orientation="latitude" precision="1" value="60.46157">60.46157°N</geoCoordinate>
,
<geoCoordinate degrees="22.37418" direction="east" orientation="longitude" precision="1" value="22.37418">22.37418°E</geoCoordinate>
<geoCoordinate id="DA333C8B12925659EA20E7B5DA30C7B3" degrees="22.37418" direction="east" orientation="longitude" precision="1" value="22.37418">22.37418°E</geoCoordinate>
; alt. 38 m; 18 May. 2016 - 1 Jun. 2016; E. Viitanen leg.; Malaise trap; CKLP. • 1 ♂; Tervola, Karhakkamaanjanka;
<geoCoordinate degrees="66.19764" direction="north" orientation="latitude" precision="1" value="66.19764">66.19764°N</geoCoordinate>
<geoCoordinate id="4DEFBA6444C6D78BB7EF2C25C9B0D5D2" degrees="66.19764" direction="north" orientation="latitude" precision="1" value="66.19764">66.19764°N</geoCoordinate>
,
<geoCoordinate degrees="25.12660" direction="east" orientation="longitude" precision="1" value="25.1266">25.12660°E</geoCoordinate>
<geoCoordinate id="FB4A92657A8137583B72166457CD8A63" degrees="25.12660" direction="east" orientation="longitude" precision="1" value="25.1266">25.12660°E</geoCoordinate>
; alt. 58 m; 25 May. 2004 - 28 Jun. 2004; J. Salmela leg.; CKLP.
<emphasis bold="true" pageId="0" pageNumber="13">Russia</emphasis>
<emphasis id="823BBF4E238A350D06A974C6985A67F3" bold="true" pageId="0" pageNumber="13">Russia</emphasis>
• 1 ♂; Krasnoyarsk Krai, Turukhansky District, Igarka City, within the settlement, swampy lake shore in the city;
<geoCoordinate degrees="67.466" direction="north" orientation="latitude" precision="55" value="67.466">67.466°N</geoCoordinate>
<geoCoordinate id="F24D7D1192B3338B0B70FB1573646E93" degrees="67.466" direction="north" orientation="latitude" precision="55" value="67.466">67.466°N</geoCoordinate>
,
<geoCoordinate degrees="86.581" direction="east" orientation="longitude" precision="55" value="86.581">86.581°E</geoCoordinate>
<geoCoordinate id="BEEB6C5A2E50EE119CF4B9A7BE8D4507" degrees="86.581" direction="east" orientation="longitude" precision="55" value="86.581">86.581°E</geoCoordinate>
; alt. 23 m; 30 Jun. 1967; K.B. Gorodkov leg.; CKLP. • 1 ♀; Krasnoyarsk Krai, Igarka City, within the settlement, sedge swamp;
<geoCoordinate degrees="67.466" direction="north" orientation="latitude" precision="55" value="67.466">67.466°N</geoCoordinate>
<geoCoordinate id="D48D47594E905781D5305FAA8C9C76DD" degrees="67.466" direction="north" orientation="latitude" precision="55" value="67.466">67.466°N</geoCoordinate>
,
<geoCoordinate degrees="86.581" direction="east" orientation="longitude" precision="55" value="86.581">86.581°E</geoCoordinate>
<geoCoordinate id="1E0677DAD09AA4232F50642559DE1298" degrees="86.581" direction="east" orientation="longitude" precision="55" value="86.581">86.581°E</geoCoordinate>
; alt. 23 m; 1 Jul. 1967; K.B. Gorodkov leg.; CKLP.
<emphasis bold="true" pageId="0" pageNumber="13">Usa</emphasis>
<emphasis id="501C6E351C55516449727DCB98EA44A6" bold="true" pageId="0" pageNumber="13">Usa</emphasis>
• 2 ♂; Michigan, Cheboygan, hard wood swamp;
<geoCoordinate degrees="45.29277" direction="north" orientation="latitude" precision="1" value="45.29277">45.29277°N</geoCoordinate>
<geoCoordinate id="EBB660A85E8E4F5D4291C019B88F0BE7" degrees="45.29277" direction="north" orientation="latitude" precision="1" value="45.29277">45.29277°N</geoCoordinate>
,
<geoCoordinate degrees="84.42805" direction="west" orientation="longitude" precision="1" value="-84.42805">84.42805° W</geoCoordinate>
<geoCoordinate id="28E4F1F9CEFA1983D850EB8BE3BF9777" degrees="84.42805" direction="west" orientation="longitude" precision="1" value="-84.42805">84.42805° W</geoCoordinate>
, alt. 280 m; 20 May. 2015; F. Brodo leg.; CKLP.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="description">
<paragraph pageId="0" pageNumber="13">Supplementary description.</paragraph>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Male terminalia</emphasis>
<subSubSection id="FD46074353DF45703A04C93946B85634" pageId="0" pageNumber="13" type="description">
<paragraph id="88F0FDD5CD6421B1BFBB84E93B489E2E" pageId="0" pageNumber="13">Supplementary description.</paragraph>
<paragraph id="7475224381B4C50423F356C485F4D52F" pageId="0" pageNumber="13">
<emphasis id="19738E2684AAB24CDE0F9F35B6EF1CCE" bold="true" pageId="0" pageNumber="13">Male terminalia</emphasis>
directed dorsally. Tergite 9 fused with gonocoxite and sternite 9 (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32C</figureCitation>
<figureCitation id="9472CD7BED071AF71027C3F583EC6ADB" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32C</figureCitation>
); caudal margin of tergite 9 with small, rounded lateral lobe both ventral and lateral view (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32A, C</figureCitation>
<figureCitation id="CF582E245030545414F31A7CF6860AB0" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32A, C</figureCitation>
); posterior margin U- or V-shaped. Sternite 9 reduced to narrow band (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32B, C</figureCitation>
<figureCitation id="FF408469EA7DE176973ABFCE7B1D8994" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32B, C</figureCitation>
). Gonocoxite 1.2-1.3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="B676489C1A06DA20B7906859158B9E37" originalValue="×">x</normalizedToken>
longer than tergite 9; ventral lobe relatively small, rounded (in dry specimen looks more finger-like) (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32B</figureCitation>
<figureCitation id="E6CFE1CD153705CB4A7D2D1A28E7F170" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32B</figureCitation>
); apical lobe indistinct; inner surface of gonocoxite sclerotised, with few hairs, without evident modifications. Gonostylus comparable large and complex, with a subapical tooth on outer margin; additional two or three smaller teeth basally (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32A, C</figureCitation>
<figureCitation id="854F61C0F49A84AEE7C022BD3931A946" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32A, C</figureCitation>
). Aedeagus complex 1.2-1.3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9B21559749908B7CF4A63C152D80E479" originalValue="×">x</normalizedToken>
longer than gonocoxite in lateral view (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32C</figureCitation>
<figureCitation id="DA135DD2797EAA80C49C559FA9020F0D" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32C</figureCitation>
); ejaculator apodeme large, not covered by paramere in lateral view (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32F</figureCitation>
<figureCitation id="0AEE1192EB76FC33360824EF6D9EE2F9" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32F</figureCitation>
); interbase seems flat and wide, with a small dorsal tooth in lateral view (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32F</figureCitation>
<figureCitation id="8BD5F4313187F237F37A80BF79A7E852" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32F</figureCitation>
); interbase directed dorso-laterally, with a deep notch on tip in dorsal view (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D, E</figureCitation>
<figureCitation id="DCE2D493F8D0F435DA53140C0DCDB453" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D, E</figureCitation>
); dorsal lobe between interbases indistinct or absent (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D</figureCitation>
<figureCitation id="1351126CA9B3631EC4C0B71A75547FBF" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D</figureCitation>
); aedeagus half as long as entry aedeagus complex; aedeagus trifid, straight; median branch longer than lateral branches (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D-F</figureCitation>
<figureCitation id="A93C2587D56FDF68A27329505BDBF426" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D-F</figureCitation>
), with a triangular dorsal outgrowth; tip slightly bifid or trifid, depend on angle (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32A</figureCitation>
<figureCitation id="FF1B02686729217FA4E013E75F2659F9" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32A</figureCitation>
); lateral branches slightly curved laterally in dorsal and ventral view (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D, E</figureCitation>
<figureCitation id="915ED1D36CD8AAC61BEF61125CED40FE" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D, E</figureCitation>
); tips of branch widened dorsally; sperm ducts branching from short elongation of sperm pump; branching area dark (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D-F</figureCitation>
<figureCitation id="239DDEF2BD9C4DBA9345058D41D2B8B6" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Male genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, ventral view; C Terminalia, lateral view D aedeagus complex, dorsal view E aedeagus complex, ventral view F aedeagus complex, lateral view." figureDoi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13">32D-F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13" start="Figure 32" startId="F32">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 32.</emphasis>
<caption id="4A50ECE0B6DC1180EE7B1918C9B6B537" doi="10.3897/zookeys.1083.75624.figure32" httpUri="https://binary.pensoft.net/fig/637505" pageId="0" pageNumber="13" start="Figure 32" startId="F32">
<paragraph id="0A35962E1C5D0C01DB8D74A3CC1772B8" pageId="0" pageNumber="13">
<emphasis id="7FAB907C0A75A3C50757579AD6DCD814" bold="true" pageId="0" pageNumber="13">Figure 32.</emphasis>
Male genital structures of
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera replicata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="replicata">
<emphasis italics="true" pageId="0" pageNumber="13">Phalacrocera replicata</emphasis>
<taxonomicName id="F04317C953766CBCA810D98130364955" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera replicata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="replicata">
<emphasis id="56A1C9DE4944B556DD53D19287052340" italics="true" pageId="0" pageNumber="13">Phalacrocera replicata</emphasis>
</taxonomicName>
(Linnaeus, 1758)
<emphasis bold="true" pageId="0" pageNumber="13">A</emphasis>
<emphasis id="2CF2DC1F47D31C4023C5B2DCE3299D06" bold="true" pageId="0" pageNumber="13">A</emphasis>
terminalia, dorsal view
<emphasis bold="true" pageId="0" pageNumber="13">B</emphasis>
<emphasis id="F134EA3F3E47FA419E0754D1AE6C2ADC" bold="true" pageId="0" pageNumber="13">B</emphasis>
terminalia, ventral view;
<emphasis bold="true" pageId="0" pageNumber="13">C</emphasis>
<emphasis id="5F97F0F97C9AF3B69DA1BC6B5DE2DAD9" bold="true" pageId="0" pageNumber="13">C</emphasis>
Terminalia, lateral view
<emphasis bold="true" pageId="0" pageNumber="13">D</emphasis>
<emphasis id="4B30431CA6EBC3DD5BFF39980050BCE0" bold="true" pageId="0" pageNumber="13">D</emphasis>
aedeagus complex, dorsal view
<emphasis bold="true" pageId="0" pageNumber="13">E</emphasis>
<emphasis id="9ADE5920C06548DFDC8BBE5AF2A23FF2" bold="true" pageId="0" pageNumber="13">E</emphasis>
aedeagus complex, ventral view
<emphasis bold="true" pageId="0" pageNumber="13">F</emphasis>
<emphasis id="04474F5776D1D02B47786624CF04A698" bold="true" pageId="0" pageNumber="13">F</emphasis>
aedeagus complex, lateral view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Female terminalia</emphasis>
<paragraph id="C334C7B00D52382FACAA0E358DA43154" pageId="0" pageNumber="13">
<emphasis id="03A848748E8C71B4BFB7EE8C6B149C79" bold="true" pageId="0" pageNumber="13">Female terminalia</emphasis>
: Tergite 8 posterior part membranous, with few hairs, but not divided medially (Fig.
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<figureCitation id="742F1FC9DFE32386911AB55CED780B2E" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33A</figureCitation>
); wider than tergite 9 in lateral view (Fig.
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<figureCitation id="DD3449FD28A23321E40D411C8F0E3D72" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33B</figureCitation>
). Tergite 10 with small, slightly separated median lobe in middle of posterior margin (Fig.
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<figureCitation id="A967D6D8C12CBD8B83B5B79D29B9FB77" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33A, B</figureCitation>
); covered with short hairs. Triangular sclerite widely fused with tip of tergite 10; tergite 10 without lateral lobe (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33B</figureCitation>
<figureCitation id="0285D07987869B9AE57A62C03C98F4A5" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33B</figureCitation>
). Cercus elongated blade, with dorsal margin with weakly serrate margin (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33B</figureCitation>
<figureCitation id="84FAB48007E1E54A79D16DB089D1ED08" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33B</figureCitation>
). Hypogynial valve wide and long, dorsal margin close to tip with tooth-like lobe, directed caudally (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33B, C</figureCitation>
<figureCitation id="9437268660EF07E647C357CD5821F821" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33B, C</figureCitation>
). Common spermathecal duct short or indistinct; sperm ducts simple, without distinct pattern, tapering proximally (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33D</figureCitation>
<figureCitation id="08C7A1262D30EB1CD2F1B492D8D3AC36" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33D</figureCitation>
); three spermathecae elongated, with straight duct (Fig.
<figureCitation captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33E</figureCitation>
<figureCitation id="485A911B037C5155D6FC9A34D029ADA6" captionStart="Figure 33" captionStartId="F33" captionText="Figure 33. Female genital structures of Phalacrocera replicata (Linnaeus, 1758) A terminalia, dorsal view B terminalia, lateral view C sternite 8, hypogynial valve, and genital fork, inner dorsal view D sperm ducts E spermathecae." figureDoi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13">33E</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13" start="Figure 33" startId="F33">
<paragraph pageId="0" pageNumber="13">
<emphasis bold="true" pageId="0" pageNumber="13">Figure 33.</emphasis>
<caption id="1B1D4BED73AE3B68568BBE1B5D13FE61" doi="10.3897/zookeys.1083.75624.figure33" httpUri="https://binary.pensoft.net/fig/637506" pageId="0" pageNumber="13" start="Figure 33" startId="F33">
<paragraph id="D766742A527417CF1669F2CAA36BADF2" pageId="0" pageNumber="13">
<emphasis id="51C2F2B2D4B78F0FA4FA34910CDA2277" bold="true" pageId="0" pageNumber="13">Figure 33.</emphasis>
Female genital structures of
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera replicata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="replicata">
<emphasis italics="true" pageId="0" pageNumber="13">Phalacrocera replicata</emphasis>
<taxonomicName id="A679702F0B0DE61C435ADFDE28143C6A" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Insecta" family="Cylindrotomidae" genus="Phalacrocera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalacrocera replicata" order="Diptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="replicata">
<emphasis id="BEACA6F4FD3DE1412E9BA5CE269362BA" italics="true" pageId="0" pageNumber="13">Phalacrocera replicata</emphasis>
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(Linnaeus, 1758)
<emphasis bold="true" pageId="0" pageNumber="13">A</emphasis>
<emphasis id="907D4409AD6219E62E903C370A54EACD" bold="true" pageId="0" pageNumber="13">A</emphasis>
terminalia, dorsal view
<emphasis bold="true" pageId="0" pageNumber="13">B</emphasis>
<emphasis id="7DE1737D9C573E6E4C47730BC92AD9EE" bold="true" pageId="0" pageNumber="13">B</emphasis>
terminalia, lateral view
<emphasis bold="true" pageId="0" pageNumber="13">C</emphasis>
<emphasis id="F8B77920CBBDA2B285FF3595C377B9AE" bold="true" pageId="0" pageNumber="13">C</emphasis>
sternite 8, hypogynial valve, and genital fork, inner dorsal view
<emphasis bold="true" pageId="0" pageNumber="13">D</emphasis>
<emphasis id="883276CF94587DF4946F5E61C23022A0" bold="true" pageId="0" pageNumber="13">D</emphasis>
sperm ducts
<emphasis bold="true" pageId="0" pageNumber="13">E</emphasis>
<emphasis id="8FD7E78D93E6BC81B5156BC1D8DA2227" bold="true" pageId="0" pageNumber="13">E</emphasis>
spermathecae.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="13" type="distribution">
<paragraph pageId="0" pageNumber="13">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="13">
<subSubSection id="06BCB087730945087B66D977F20DA8F2" pageId="0" pageNumber="13" type="distribution">
<paragraph id="C6212D84CEA8AFDFAD6AF41F4155B7FA" pageId="0" pageNumber="13">Distribution.</paragraph>
<paragraph id="5C572208E1DCB0629F6521C5AF203B2A" pageId="0" pageNumber="13">
Widely distributed in Holarctic. It has been reported from the Nearctic: Canada and USA from Ontario and Quebec, south to Michigan, Pennsylvania and Massachusetts. Palearctic: Austria, Belarus, Belgium, China (Heilongjiang), Czech Republic, Denmark, Estonia, Finland, France, Germany, Great Britain, Ireland, Italy (north), Lithuania, Mongolia, Netherlands, Norway, Poland, Russia: North European Russia, Central European Russia, East Siberia (Irkutsk Oblast), Far East (Republic of Sakha (Yakutia), Spain (Zamoro), Sweden, Switzerland, Ukraine (
<bibRefCitation author="Oosterbroek, P" journalOrPublisher="Bulletin de la Societe Neuchateloise des Sciences Naturelles" pageId="0" pageNumber="13" publicationUrl="https://ccw.naturalis.nl/index.php" refId="B39" refString="Oosterbroek, P, 2021. Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea). https://ccw.naturalis.nl/index.php" title="Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea)." url="https://ccw.naturalis.nl/index.php" year="2021">Oosterbroek 2021</bibRefCitation>
<bibRefCitation id="28529EA556BFA0A18D91E54A70BD7522" author="Oosterbroek, P" journalOrPublisher="Bulletin de la Societe Neuchateloise des Sciences Naturelles" pageId="0" pageNumber="13" publicationUrl="https://ccw.naturalis.nl/index.php" refId="B39" refString="Oosterbroek, P, 2021. Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea). https://ccw.naturalis.nl/index.php" title="Catalogue of the Crane flies of the World (Insecta, Diptera, Nematocera, Tipuloidea)." url="https://ccw.naturalis.nl/index.php" year="2021">Oosterbroek 2021</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.15298/rusentj.30.2.13" author="Paramonov, NM" journalOrPublisher="Russian Entomological Journal" pageId="0" pageNumber="13" pagination="182 - 188" refId="B46" refString="Paramonov, NM, Pilipenko, VE, 2021. New synonym and new records of craneflies (Diptera: Tipuloidea) in China. Russian Entomological Journal 30 (2): 182 - 188, DOI: https://doi.org/10.15298/rusentj.30.2.13" title="New synonym and new records of craneflies (Diptera: Tipuloidea) in China." url="https://doi.org/10.15298/rusentj.30.2.13" volume="30" year="2021">Paramonov and Pilipenko 2021</bibRefCitation>
<bibRefCitation id="0163984955C79806A5EF3472839E1BCD" DOI="https://doi.org/10.15298/rusentj.30.2.13" author="Paramonov, NM" journalOrPublisher="Russian Entomological Journal" pageId="0" pageNumber="13" pagination="182 - 188" refId="B46" refString="Paramonov, NM, Pilipenko, VE, 2021. New synonym and new records of craneflies (Diptera: Tipuloidea) in China. Russian Entomological Journal 30 (2): 182 - 188, DOI: https://doi.org/10.15298/rusentj.30.2.13" title="New synonym and new records of craneflies (Diptera: Tipuloidea) in China." url="https://doi.org/10.15298/rusentj.30.2.13" volume="30" year="2021">Paramonov and Pilipenko 2021</bibRefCitation>
).. Here we record the species for the first time from Krasnoyarsk Krai, East Siberia, Russia.
</paragraph>
</subSubSection>

288
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<mods:affiliation id="9450847FE9E28606D143CC2B6A866764">Senckenberg Museum of Natural History Goerlitz, Am Museum 1, 02826 Goerlitz, Germany</mods:affiliation>
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<mods:date>2020</mods:date>
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<mods:number>2020-09-21</mods:number>
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<treatment ID-GBIF-Taxon="182223411" LSID="urn:lsid:zoobank.org:act:632BFBD8-D80E-4AA5-B16B-46D0E2662D07" httpUri="http://treatment.plazi.org/id/E36006BBA46F5A51A4A5D21AFB3660CA" lastPageNumber="183" pageId="0" pageNumber="183">
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<paragraph pageId="0" pageNumber="183">
<taxonomicName LSID="http://zoobank.org/632BFBD8-D80E-4AA5-B16B-46D0E2662D07" authority="Bernhard, 2020" authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens" status="sp. nov.">Plagiolepis invadens</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="183">sp. nov.</taxonomicNameLabel>
<treatment id="E36006BBA46F5A51A4A5D21AFB3660CA" ID-GBIF-Taxon="182223411" LSID="urn:lsid:zoobank.org:act:632BFBD8-D80E-4AA5-B16B-46D0E2662D07" httpUri="http://treatment.plazi.org/id/E36006BBA46F5A51A4A5D21AFB3660CA" lastPageNumber="183" pageId="0" pageNumber="183" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="4B30C8BDD479BCCC0EC7CA7955EDD37A" pageId="0" pageNumber="183" type="nomenclature">
<paragraph id="5880CBFC7ADE5898640151916C986235" pageId="0" pageNumber="183">
<taxonomicName id="CEFE4B05F3A388BFE825921CD6A8ADE8" LSID="http://zoobank.org/632BFBD8-D80E-4AA5-B16B-46D0E2662D07" authority="Bernhard, 2020" authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens" status="sp. nov.">Plagiolepis invadens</taxonomicName>
<taxonomicNameLabel id="4BC95AD3836C251A1621A55445B0AAB7" pageId="0" pageNumber="183">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection pageId="0" pageNumber="183" type="etymology">
<paragraph pageId="0" pageNumber="183">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="183">
<subSubSection id="7B0E472313A2B64BCAA2F4A3D6D6F156" pageId="0" pageNumber="183" type="etymology">
<paragraph id="FC825E7834E09FB46247E02288693D1B" pageId="0" pageNumber="183">Etymology.</paragraph>
<paragraph id="F3F53DBAC574A6B14B63EDF4A12A6800" pageId="0" pageNumber="183">
Meaning
<normalizedToken originalValue="“invasive”">&quot;invasive&quot;</normalizedToken>
<normalizedToken id="BF7E438667CF8FB1CFED4C2D15DB9498" originalValue="“invasive”">&quot;invasive&quot;</normalizedToken>
(from Latin
<emphasis italics="true" pageId="0" pageNumber="183">invado</emphasis>
<emphasis id="3DB4ADFA85ECD5E3D4358F1AEEBB2005" italics="true" pageId="0" pageNumber="183">invado</emphasis>
) The type colony in SW Germany is an anthropogenous introduction from an unknown origin, shows circannual outdoor nesting, but invaded houses in large numbers during the extremely dry summers of 2018 and 2019.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="type material">
<paragraph pageId="0" pageNumber="183">Type material.</paragraph>
<paragraph pageId="0" pageNumber="183">
<subSubSection id="2C8F394AAECEC3D8035BDD2886967F41" pageId="0" pageNumber="183" type="type material">
<paragraph id="9FD2D2CDB3CC4760CC536C6F097AF5B8" pageId="0" pageNumber="183">Type material.</paragraph>
<paragraph id="09814B5E94F17CF4495EF6930FE703D9" pageId="0" pageNumber="183">
Holotype plus one paratype worker on the same pin labelled &quot;GER:
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<geoCoordinate id="5AD6A24C1EEC23F9FFC8A6DDD7BFE9B3" degrees="49.65703" direction="north" orientation="latitude" precision="1" value="49.65703">49.65703°N</geoCoordinate>
,
<geoCoordinate degrees="8.41775" direction="east" orientation="longitude" precision="1" value="8.41775">8.41775°E</geoCoordinate>
<geoCoordinate id="A3D288AC92FA541FD394219162B42298" degrees="8.41775" direction="east" orientation="longitude" precision="1" value="8.41775">8.41775°E</geoCoordinate>
Hofheim, 92 m, supercolony in garden, known since about 4 years, leg. Heller 2019.08&quot; and &quot;Holotype (top) and paratype of
<taxonomicName authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Plagiolepis invadens</emphasis>
<taxonomicName id="A82F226CEE701BA27EBECF32EA624F0C" authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="DC3750B23EC50258F8833B88CBFCFE1B" italics="true" pageId="0" pageNumber="183">Plagiolepis invadens</emphasis>
</taxonomicName>
Seifert&quot;; three paratype workers from the same collecting data; five paratype workers labelled &quot;GER:
<geoCoordinate degrees="49.65703" direction="north" orientation="latitude" precision="1" value="49.65703">49.65703°N</geoCoordinate>
<geoCoordinate id="C4A018F4F5BDA78D36F8B1512F9B17DC" degrees="49.65703" direction="north" orientation="latitude" precision="1" value="49.65703">49.65703°N</geoCoordinate>
,
<geoCoordinate degrees="8.41775" direction="east" orientation="longitude" precision="1" value="8.41775">8.41775°E</geoCoordinate>
<geoCoordinate id="A4A0E464FD558F044EF48E75FBE8A2AF" degrees="8.41775" direction="east" orientation="longitude" precision="1" value="8.41775">8.41775°E</geoCoordinate>
Hofheim, 92 m, supercolony in garden, known since about 5 years, leg. Heller 2020.06&quot;; all material is stored in SMN
<normalizedToken originalValue="Görlitz">Goerlitz</normalizedToken>
<normalizedToken id="9A438740FE0FAA5916E6DC62E841D69D" originalValue="Görlitz">Goerlitz</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="diagnosis">
<paragraph pageId="0" pageNumber="183">Diagnosis and taxonomy</paragraph>
<paragraph pageId="0" pageNumber="183">
<subSubSection id="9D0387ADC7F39D96481FC1C7B196EB54" pageId="0" pageNumber="183" type="diagnosis">
<paragraph id="EA87157F81337AF9C227A8BED1521417" pageId="0" pageNumber="183">Diagnosis and taxonomy</paragraph>
<paragraph id="5DA0D5027207D4715EDF40EFE5F89034" pageId="0" pageNumber="183">
(Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. RAV-corrected morphometric data of worker individuals of the Plagiolepis schmitzii complex given as arithmetic mean + / - standard deviation [minimum, maximum]; n = number of individuals. F values and significance levels p are from a univariate ANOVA and evaluate the differences between Pl. atlantis and Pl. invadens sp. nov." httpUri="http://table.plazi.org/id/D391BECA51134E5ACE149FBAA04D14FB" pageId="0" pageNumber="183" tableUuid="D391BECA51134E5ACE149FBAA04D14FB">1</tableCitation>
<tableCitation id="EDC2B1F35D6C2C2D7407C98988A1DC7C" captionStart="Table 1" captionStartId="T1" captionText="Table 1. RAV-corrected morphometric data of worker individuals of the Plagiolepis schmitzii complex given as arithmetic mean + / - standard deviation [minimum, maximum]; n = number of individuals. F values and significance levels p are from a univariate ANOVA and evaluate the differences between Pl. atlantis and Pl. invadens sp. nov." httpUri="http://table.plazi.org/id/D391BECA51134E5ACE149FBAA04D14FB" pageId="0" pageNumber="183" tableUuid="D391BECA51134E5ACE149FBAA04D14FB">1</tableCitation>
, key, Figures
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head of the holotype worker of Plagiolepis invadens sp. nov." figureDoi="10.3897/dez.67.53199.figure6" httpUri="https://binary.pensoft.net/fig/455224" pageId="0" pageNumber="183">6</figureCitation>
<figureCitation id="9CE6365783B76D2C6AF37AE9FFDE2AE6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Head of the holotype worker of Plagiolepis invadens sp. nov." figureDoi="10.3897/dez.67.53199.figure6" httpUri="https://binary.pensoft.net/fig/455224" pageId="0" pageNumber="183">6</figureCitation>
-
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal aspect of the holotype worker of Plagiolepis invadens sp. nov." figureDoi="10.3897/dez.67.53199.figure8" httpUri="https://binary.pensoft.net/fig/455226" pageId="0" pageNumber="183">8</figureCitation>
<figureCitation id="7272612B0E1FB1CB5527CE55198434B0" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Dorsal aspect of the holotype worker of Plagiolepis invadens sp. nov." figureDoi="10.3897/dez.67.53199.figure8" httpUri="https://binary.pensoft.net/fig/455226" pageId="0" pageNumber="183">8</figureCitation>
). With narrowly-spaced basal pits of pubescence hairs (BPdG 15.6
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="003923456A4414D881488A696B8342D6" originalValue="µm">µm</normalizedToken>
), low pubescence distance (sqPDG 2.69) and the 4th funiculus segment not being much longer than the 3rd (F4/F3 1.346),
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
<taxonomicName id="A4D875270AF864ED701E12242CBD1925" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="B394AE1F9F94A8AF1E4588C6A07A6DAB" italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
</taxonomicName>
sp. nov. is clearly separable from the species of the
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. pygmaea" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="pygmaea">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. pygmaea</emphasis>
<taxonomicName id="9B5D6765B68FA8A9B53DF6E240017F77" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. pygmaea" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="537A632FBD93BFACC2881CABA4FC9611" italics="true" pageId="0" pageNumber="183">Pl. pygmaea</emphasis>
</taxonomicName>
and
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. pallescens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="pallescens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. pallescens</emphasis>
<taxonomicName id="F27A180B67FBB177365A7DDA8881C17F" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. pallescens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="428EC4BFBD9096274483A17FAB65BE94" italics="true" pageId="0" pageNumber="183">Pl. pallescens</emphasis>
</taxonomicName>
group and is characterised as a member of the
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. schmitzii" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="schmitzii">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. schmitzii</emphasis>
<taxonomicName id="6FAA8D054DD7D05C662D6C0056ABBF92" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. schmitzii" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="schmitzii">
<emphasis id="C2C994B16EFB51D22074EC96D7088B4B" italics="true" pageId="0" pageNumber="183">Pl. schmitzii</emphasis>
</taxonomicName>
group. It is outstanding within the latter group by the very short 3rd funiculus segment: primary ratios of F3/CS are 7.37
<normalizedToken originalValue="±">+/-</normalizedToken>
group. It is outstanding within the latter group by the very short 3rd funiculus segment: primary ratios of F3/CS are 7.37
<normalizedToken id="968D206D072D2BC0874DDF38FE0C6BAB" originalValue="±">+/-</normalizedToken>
0.43 [6.85, 8.04]% in ten workers of
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
<taxonomicName id="1BAE5BD062CC96669345A4C4FCA96910" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="0D97F00006310929C9D700256BBE3AA9" italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
</taxonomicName>
sp. nov., but 9.78
<normalizedToken originalValue="±">+/-</normalizedToken>
sp. nov., but 9.78
<normalizedToken id="6C057C16926C2A608332791AE1BDD8DD" originalValue="±">+/-</normalizedToken>
0.61 [8.59, 11.59]% in 124 specimens of the other three
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. schmitzii" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="schmitzii">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. schmitzii</emphasis>
<taxonomicName id="427F0F8FDBDA12C7B517A6C49EC28216" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. schmitzii" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="schmitzii">
<emphasis id="4F85658C4C74004A702E84A9BC1095FE" italics="true" pageId="0" pageNumber="183">Pl. schmitzii</emphasis>
</taxonomicName>
group species. The most similar species is
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. atlantis" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="atlantis">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. atlantis</emphasis>
<taxonomicName id="543C5C0ADF9225EC2A13A573DCBACE3D" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. atlantis" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="64F539E7F8F75AD594DA3F50D054A6EC" italics="true" pageId="0" pageNumber="183">Pl. atlantis</emphasis>
</taxonomicName>
and it may be asked if there is a risk of synonymy, considering that the description of
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
<taxonomicName id="27249DC0065E699BE6B4BFC3D4382051" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="D7D1581BB2D9C2C2F1ED056746D2B1E3" italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
</taxonomicName>
sp. nov. is based on workers from only a single supercolony. Running a PCA, considering absolute head size and the 16 RAV-corrected shape, pubescence and surface characters, resulted in a strong separation of all individuals by the 1st principal component:
</paragraph>
<caption doi="10.3897/dez.67.53199.figure6" httpUri="https://binary.pensoft.net/fig/455224" pageId="0" pageNumber="183" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="183">
<emphasis bold="true" pageId="0" pageNumber="183">Figure 6.</emphasis>
<caption id="1A8126EC23BF2E0B9591D3CFFC26CF0F" doi="10.3897/dez.67.53199.figure6" httpUri="https://binary.pensoft.net/fig/455224" pageId="0" pageNumber="183" start="Figure 6" startId="F6">
<paragraph id="9E3A6F743C2B843931F8ACC1ECE3F59C" pageId="0" pageNumber="183">
<emphasis id="BC0E298D6005EE7F5F1DD1EBD089146A" bold="true" pageId="0" pageNumber="183">Figure 6.</emphasis>
Head of the holotype worker of
<taxonomicName authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Plagiolepis invadens</emphasis>
<taxonomicName id="1A3EBE4F45ED9E71F8DE79219116AF90" authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="E286717708B91595F685B1BB369AC5ED" italics="true" pageId="0" pageNumber="183">Plagiolepis invadens</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="183">
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
<paragraph id="AD7D98FCEB1C9E3ADEAD47C3261A416B" pageId="0" pageNumber="183">
<taxonomicName id="85AB8E886E578BD85A645E518A97B075" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="DA8509DC6D174988BDA986A68E51E575" italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
</taxonomicName>
sp. nov. -1.946
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="088A6FDAB9415D213CF8D45D09E6B837" originalValue="±">+/-</normalizedToken>
0.343 [-2.477, -1.368] n = 10
</paragraph>
<paragraph pageId="0" pageNumber="183">
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. atlantis" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="atlantis">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. atlantis</emphasis>
<paragraph id="9C0842F1E096715AF55AB1338FF073B8" pageId="0" pageNumber="183">
<taxonomicName id="AADCB3A3E7D36A369D2A267867E327C4" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. atlantis" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="F1D2C9807B5F61C1A2C053C531AEC4C6" italics="true" pageId="0" pageNumber="183">Pl. atlantis</emphasis>
</taxonomicName>
0.348
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="C3AB8062B1D86485E29EE64521EE7333" originalValue="±">+/-</normalizedToken>
0.592 [-1.020, 1.338] n = 56.
</paragraph>
<paragraph pageId="0" pageNumber="183">This clear result (ANOVA, F1,64 = 140.4, p &lt;0.001), achieved without character selection, is a reasonable indication of heterospecificity. A vector considering the 1st and 3rd principal component with 1.893*PC1 + 0.563*PC3 provides an even stronger separation (ANOVA, F1,64 = 189.6, p &lt;&lt; 0.001):</paragraph>
<paragraph pageId="0" pageNumber="183">
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
<paragraph id="D4013A11D028BEB7194933443024A583" pageId="0" pageNumber="183">This clear result (ANOVA, F1,64 = 140.4, p &lt;0.001), achieved without character selection, is a reasonable indication of heterospecificity. A vector considering the 1st and 3rd principal component with 1.893*PC1 + 0.563*PC3 provides an even stronger separation (ANOVA, F1,64 = 189.6, p &lt;&lt; 0.001):</paragraph>
<paragraph id="085745BA967FEB0326257A17BBCA8FD3" pageId="0" pageNumber="183">
<taxonomicName id="CF59B2E1C523FD86BF2C634107FA766B" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="ABBDD1E4946BDE8D28755689874667FE" italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
</taxonomicName>
sp. nov. -4.010
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="4F2E7E9E2938E8643A59B7F2C27D6C42" originalValue="±">+/-</normalizedToken>
0.649 [-4.962, -2.932] n = 10
</paragraph>
<paragraph pageId="0" pageNumber="183">
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. atlantis" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="atlantis">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. atlantis</emphasis>
<paragraph id="A100657EAEBE875F1BB6A24D3AAADBB1" pageId="0" pageNumber="183">
<taxonomicName id="391BF7184412BBA10154260A699A96B3" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. atlantis" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="3FE2E6A681ED740720835A1B6AAF1D8B" italics="true" pageId="0" pageNumber="183">Pl. atlantis</emphasis>
</taxonomicName>
0.716
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="4F8AB3715E1D1B9B55C46FFEAF84EB8D" originalValue="±">+/-</normalizedToken>
1.046 [-1.597, 2.681] n = 56.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="further descriptive statements">
<paragraph pageId="0" pageNumber="183">Further descriptive statements.</paragraph>
<paragraph pageId="0" pageNumber="183">Head moderately elongated (CL/CW 1.105). Scape shorter than in related species (SL/CS 0.923). Eye medium-sized (EL/CS 0.258). Mesosoma width smaller than in related species (MW/CS 0.615). Cuticular surface of head, mesosoma, coxae and femora brilliantly shining and with a dilute appressed to decumbent pubescence. Scape and tibiae with a more dense decumbent pubescence. Head, scape, femora and mesosoma varying from dark brown with yellowish tinge to almost black. Antennal funiculus, coxae, tibiae and sometimes pronotum pale yellowish-brown.</paragraph>
<caption doi="10.3897/dez.67.53199.figure7" httpUri="https://binary.pensoft.net/fig/455225" pageId="0" pageNumber="183" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="183">
<emphasis bold="true" pageId="0" pageNumber="183">Figure 7.</emphasis>
<subSubSection id="798406580D8934481FCF867EC19D8258" pageId="0" pageNumber="183" type="further descriptive statements">
<paragraph id="2C983D957C1E2DB9F6F90A28C4989B50" pageId="0" pageNumber="183">Further descriptive statements.</paragraph>
<paragraph id="3EDC4094DBC7000DC3A4E07668E78176" pageId="0" pageNumber="183">Head moderately elongated (CL/CW 1.105). Scape shorter than in related species (SL/CS 0.923). Eye medium-sized (EL/CS 0.258). Mesosoma width smaller than in related species (MW/CS 0.615). Cuticular surface of head, mesosoma, coxae and femora brilliantly shining and with a dilute appressed to decumbent pubescence. Scape and tibiae with a more dense decumbent pubescence. Head, scape, femora and mesosoma varying from dark brown with yellowish tinge to almost black. Antennal funiculus, coxae, tibiae and sometimes pronotum pale yellowish-brown.</paragraph>
<caption id="1D2F2BC745D15EAFBD31295D937DFAF2" doi="10.3897/dez.67.53199.figure7" httpUri="https://binary.pensoft.net/fig/455225" pageId="0" pageNumber="183" start="Figure 7" startId="F7">
<paragraph id="B279AF153C1036C0CA5FE53929A8A839" pageId="0" pageNumber="183">
<emphasis id="EC264ACE0742005B8234545524B62924" bold="true" pageId="0" pageNumber="183">Figure 7.</emphasis>
Lateral aspect of the holotype worker of
<taxonomicName authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Plagiolepis invadens</emphasis>
<taxonomicName id="7324D6BB66C3F8A38624AF2447F07829" authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="0567B8A77C3181F715D6712F88854090" italics="true" pageId="0" pageNumber="183">Plagiolepis invadens</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="biology_ecology">
<paragraph pageId="0" pageNumber="183">Biology.</paragraph>
<paragraph pageId="0" pageNumber="183">
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
<subSubSection id="72569D6EAA2A62DF2481E5040C28733A" pageId="0" pageNumber="183" type="biology_ecology">
<paragraph id="A349DC5815FC619E6DD494578B102D79" pageId="0" pageNumber="183">Biology.</paragraph>
<paragraph id="B2756DC117DF06B067FE4AF90566A25B" pageId="0" pageNumber="183">
<taxonomicName id="A7EB48CD3448975DAD38269D5AE30155" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="762443423173E6C13E6FCB6B9C6805D8" italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
</taxonomicName>
sp. nov. is known so far from only a single supercolony in SW Germany in a settlement with about 30% greenery and 70% building or sealed area. Residents became aware of the ants in the gardens in about 2016. Ants were not perceived as plagues inside the houses in the years 2016 and 2017, but masses of workers invaded houses during the extremely dry summers of 2018 and 2019 in such numbers that the residents tried to get rid of the ants by using vacuum cleaners. Gerhard Heller observed in September 2019 and June 2020 the presence of a true supercolony with millions of workers and runways stretching along the roadside of at least two properties. Preferred nest sites were the most humid spots with much greenery where the ants constructed small hills made of soil ejections. The residents also reported that &quot;black ants being clearly bigger&quot; than the
<taxonomicName authorityName="Mayr" authorityYear="1861" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="183">Plagiolepis</emphasis>
<taxonomicName id="AD7DA9E24D87E7E525DF2E4E14C18E63" authorityName="Mayr" authorityYear="1861" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="7BA09744BAC1B581378DC0468FA4A2A6" italics="true" pageId="0" pageNumber="183">Plagiolepis</emphasis>
</taxonomicName>
- presumably
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Hexapoda" family="Formicidae" genus="Lasius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lasius niger" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="niger">
<emphasis italics="true" pageId="0" pageNumber="183">Lasius niger</emphasis>
<taxonomicName id="402A88A38D2E37FB3DA26D67751360F9" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Hexapoda" family="Formicidae" genus="Lasius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lasius niger" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="7851ED9C5C4470CE877705F59AE1E5B3" italics="true" pageId="0" pageNumber="183">Lasius niger</emphasis>
</taxonomicName>
(Linnaeus, 1758) - vanished after the development of the
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
<taxonomicName id="6BBEA1CFC5F64D62F82CAE6FF6DE4BC4" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="39555465195736E829FDBCA6CE384FB8" italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
</taxonomicName>
sp. nov. supercolony. The species is obviously able to long-time survival under outdoor conditions within the current climatic scenario and will have to be considered as established neozoon in Germany if efforts to eradicate the population fail. Reproductive biology, demography and food ecology of
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
<taxonomicName id="61A2AE80B00AFFC65EC3167418CCC115" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="9869501B8FB2B074CFB74142A1115EE8" italics="true" pageId="0" pageNumber="183">Pl. invadens</emphasis>
</taxonomicName>
sp. nov. are not studied so far, but are expected to show the traits described in the concluding chapter of this paper.
</paragraph>
<caption doi="10.3897/dez.67.53199.figure8" httpUri="https://binary.pensoft.net/fig/455226" pageId="0" pageNumber="183" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="183">
<emphasis bold="true" pageId="0" pageNumber="183">Figure 8.</emphasis>
<caption id="92570BC4D452BC2DF82BF6ABCAA6D1CF" doi="10.3897/dez.67.53199.figure8" httpUri="https://binary.pensoft.net/fig/455226" pageId="0" pageNumber="183" start="Figure 8" startId="F8">
<paragraph id="EE748EAF266880C28D352B2FD2CE055E" pageId="0" pageNumber="183">
<emphasis id="D7C45818800F3276DE65DDD68E1AC04C" bold="true" pageId="0" pageNumber="183">Figure 8.</emphasis>
Dorsal aspect of the holotype worker of
<taxonomicName authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis italics="true" pageId="0" pageNumber="183">Plagiolepis invadens</emphasis>
<taxonomicName id="73B346F5D7A2D2F7ECA980FC2C04CF22" authorityName="Bernhard" authorityYear="2020" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis invadens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="invadens">
<emphasis id="7A22A20FC72BA57E5C82F0F503EDC5A7" italics="true" pageId="0" pageNumber="183">Plagiolepis invadens</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="183" type="comments">
<paragraph pageId="0" pageNumber="183">Comments.</paragraph>
<paragraph pageId="0" pageNumber="183">
<subSubSection id="F8923AD9DA7EDA1B8D7A0967C40F64B1" pageId="0" pageNumber="183" type="comments">
<paragraph id="4F70D3709496BAE106B8F5C7E09A3699" pageId="0" pageNumber="183">Comments.</paragraph>
<paragraph id="3B0BFE180579598D3BBB0E1C16950884" pageId="0" pageNumber="183">
There is certainly some risk describing a new species based upon a single colony. Yet, this risk is calculable and apparently low. Firstly, the separation in the PCA is very strong and the next similar species
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. atlantis" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="atlantis">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. atlantis</emphasis>
<taxonomicName id="B1823835D747F86E4B9379C201528BCD" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. atlantis" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="D2F749FA6AB302F4FC12B054E4CEC192" italics="true" pageId="0" pageNumber="183">Pl. atlantis</emphasis>
</taxonomicName>
was available for this PCA in a large sample. Secondly, the reported diagnostic characters, which are homogenously distributed over the colony in space and time, are unlikely to represent a spontaneous mutant. This would require a single founding queen which was homozygous for at least one allele, both determining length of scape and funiculus segments and slenderness of the mesosoma and it would require propagation of this mutant over millions of individuals in the supercolony. Thirdly, considering the Palaearctic region, 10 taxa of the
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. schmitzii" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="schmitzii">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. schmitzii</emphasis>
<taxonomicName id="E485F8D62046C7DA25C88AD2663608CC" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. schmitzii" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="schmitzii">
<emphasis id="4CF5C8A15FFC22FE5A932757D2BF9E1D" italics="true" pageId="0" pageNumber="183">Pl. schmitzii</emphasis>
</taxonomicName>
group (reported here) and 14 taxa of the
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. pallescens" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="pallescens">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. pallescens</emphasis>
<taxonomicName id="878915CEFEBA4B10B46E5063915644F9" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. pallescens" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="pallescens">
<emphasis id="9A7BFE36A1C1020C7BC9BA94B53F9FC8" italics="true" pageId="0" pageNumber="183">Pl. pallescens</emphasis>
</taxonomicName>
and
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. pygmaea" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="pygmaea">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. pygmaea</emphasis>
<taxonomicName id="9E4A0A1AF424DBA6B5C9FF62814A4053" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. pygmaea" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="pygmaea">
<emphasis id="D0837DBEB87B5D2F42D15077BE998FC2" italics="true" pageId="0" pageNumber="183">Pl. pygmaea</emphasis>
</taxonomicName>
group have been checked and excluded as senior synonyms (Kirschner et al. in prep.). There is only one taxon which seems to pose some risk:
<taxonomicName authorityName="Emery" authorityYear="1921" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis barbara subsp. var. var. pyrenaica" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="variety" species="barbara" subSpecies="var." variety="pyrenaica">Plagiolepis barbara var. pyrenaica</taxonomicName>
<taxonomicName id="26109B5E191DADD8E34B6339CF695FA1" authorityName="Emery" authorityYear="1921" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis barbara subsp. var. var. pyrenaica" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="variety" species="barbara" subSpecies="var." variety="pyrenaica">Plagiolepis barbara var. pyrenaica</taxonomicName>
Emery, 1921, collected in the Eastern Pyrenees. Assessing a photo of a type specimen (
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), it seems to have a gastral pubescence density approaching the situation in the
<taxonomicName family="Formicidae" genus="Pl." kingdom="Animalia" lsidName="Pl. schmitzii" order="Hymenoptera" pageId="0" pageNumber="183" rank="species" species="schmitzii">
<emphasis italics="true" pageId="0" pageNumber="183">Pl. schmitzii</emphasis>
<taxonomicName id="E0DC01DAD2DE09E035DC0681A98ACD60" class="Insecta" family="Formicidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pl. schmitzii" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="species" species="schmitzii">
<emphasis id="8E997755C80BC92E0F1265B3411A5C00" italics="true" pageId="0" pageNumber="183">Pl. schmitzii</emphasis>
</taxonomicName>
group, but further conclusions are impossible. A direct investigation of type specimen(s), which should exist in MCSN Genoa, is currently not possible due to long-term effects caused by the COVID19 pandemic. Fourthly, scrutinising the photos of type specimens of 44
<taxonomicName authorityName="Mayr" authorityYear="1861" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="183">Plagiolepis</emphasis>
<taxonomicName id="0AB19909726F21131AB71EA4A83E6B02" authorityName="Mayr" authorityYear="1861" class="Insecta" family="Formicidae" genus="Plagiolepis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plagiolepis" order="Hymenoptera" pageId="0" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="B1489C1B52F9E5373D5BE140B87480DD" italics="true" pageId="0" pageNumber="183">Plagiolepis</emphasis>
</taxonomicName>
taxa from remote zoogeographic regions (
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), there is no apparent candidate taxon for senior synonymy. All these taxa differ clearly. Arranged in alphabetic order of species-level taxon names, their unique specimen identifiers were CASENT0909850, CASENT0132814, CASENT0917582, CASENT0909845, CASENT0217200, CASENT0281160, CASENT0909846, CASENT0909847, CASENT0917579, CASENT0915713, CASENT0903144, CASENT0917578, CASENT0906247, CASENT0912417, CASENT0281161, CASENT0909848, CASENT0917484, CASENT0101305, CASENT0903146, CASENT0916654, CASENT0912418, CASENT0235986, CASENT0913616, CASENT0905142, CASENT0905143, CASENT0909852, CASENT0909861, CASENT0906471, CASENT0912419, CASENT0217737, CASENT0101224, CASENT0909853, CASENT0909854, CASENT0912411, GBIF-D/FoCol 2225, CASENT0787965, CASENT0903145, CASENT0906251, CASENT0905141, CASENT0217738, CASENT0909863, CASENT0912412, CASENT0909864 and CASENT0917485.
</paragraph>
</subSubSection>

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<treatment ID-GBIF-Taxon="189470270" LSID="urn:lsid:plazi:treatment:E3C062861E095A5898A485F2C7ED7F56" httpUri="http://treatment.plazi.org/id/E3C062861E095A5898A485F2C7ED7F56" lastPageNumber="429" pageId="0" pageNumber="429">
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<taxonomicName LSID="E3C06286-1E09-5A58-98A4-85F2C7ED7F56" authority="(Ermolaev, 1927)" baseAuthorityName="Ermolaev" baseAuthorityYear="1927" class="Arachnida" family="Agelenidae" genus="Pireneitega" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pireneitega birulai" order="Araneae" pageId="0" pageNumber="429" phylum="Arthropoda" rank="species" species="birulai">Pireneitega birulai (Ermolaev, 1927)</taxonomicName>
<treatment id="E3C062861E095A5898A485F2C7ED7F56" ID-GBIF-Taxon="189470270" LSID="urn:lsid:plazi:treatment:E3C062861E095A5898A485F2C7ED7F56" httpUri="http://treatment.plazi.org/id/E3C062861E095A5898A485F2C7ED7F56" lastPageNumber="429" pageId="0" pageNumber="429" scope_class="Arachnida" scope_family="Agelenidae">
<subSubSection id="5D21C5D6CE9B9BAE8F37AD44568AA11D" pageId="0" pageNumber="429" type="nomenclature">
<paragraph id="32FB16EC75C0D702CCE88761F1B576EF" pageId="0" pageNumber="429">
<taxonomicName id="F924A3AF52D299C4510B955FEECB492F" ID-CoL="4J7WH" LSID="E3C06286-1E09-5A58-98A4-85F2C7ED7F56" authority="(Ermolaev, 1927)" baseAuthorityName="Ermolaev" baseAuthorityYear="1927" class="Arachnida" family="Agelenidae" genus="Pireneitega" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pireneitega birulai" order="Araneae" pageId="0" pageNumber="429" phylum="Arthropoda" rank="species" species="birulai">Pireneitega birulai (Ermolaev, 1927)</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="429" type="materials_examined">
<paragraph pageId="0" pageNumber="429">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="429">
<specimenCount type="male">1♂</specimenCount>
<subSubSection id="B0B20AFDC794485AC90981F13235D520" pageId="0" pageNumber="429" type="materials_examined">
<paragraph id="51BEAA74A8156305B9D804BF74FF9CAD" pageId="0" pageNumber="429">Material examined.</paragraph>
<paragraph id="002C0D497885EBC9E24E9AFCB0C31F5A" pageId="0" pageNumber="429">
<specimenCount id="C10DB2EF6FEB6B543FB0EBC46FEA05DD" type="male">1♂</specimenCount>
[2a].
</paragraph>
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<subSubSection pageId="0" pageNumber="429" type="distribution">
<paragraph pageId="0" pageNumber="429">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="429">
<subSubSection id="5A0F56F8F1156F1F5DCFFC2FCB4B8DE1" pageId="0" pageNumber="429" type="distribution">
<paragraph id="36C44DA943D0BB4EDC0D5E5CF7B71D2E" pageId="0" pageNumber="429">Distribution.</paragraph>
<paragraph id="F445DAD1288B884354E657DA15B0EFA8" pageId="0" pageNumber="429">
This species has Siberio-Central Asian boreo-montane range (
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<bibRefCitation id="5A8F3D8B7E3E8C1EDE4E7CF7B3E78C0D" author="Marusik, YM" journalOrPublisher="Institute for Biological Problems of the North, Magadan" pageId="0" pageNumber="429" refId="B15" refString="Marusik, YM, Logunov, DV, Koponen, S., 2000. Spiders of Tuva, South Siberia. Institute for Biological Problems of the North, Magadan" title="Spiders of Tuva, South Siberia." year="2000">Marusik et al. 2000</bibRefCitation>
). In Siberia, namely in the neighboring Tomsk Oblast, Altai Republic and Tuva Republic, it inhabits diverse habitats: clay cliffs, rocks, birch-aspen and
<taxonomicName genus="Abies-Pinus" lsidName="Abies-Pinus sibirica" pageId="0" pageNumber="429" rank="species" species="sibirica">
<emphasis italics="true" pageId="0" pageNumber="429">Abies-Pinus sibirica</emphasis>
<taxonomicName id="AE6D3CEE50589787C0577320F6D6C90C" genus="Abies-Pinus" lsidName="Abies-Pinus sibirica" pageId="0" pageNumber="429" rank="species" species="sibirica">
<emphasis id="6D10ADA6369472BA6EB09F829EFD9A4A" italics="true" pageId="0" pageNumber="429">Abies-Pinus sibirica</emphasis>
</taxonomicName>
forests and screes (
<bibRefCitation author="Ermolajev, WN" journalOrPublisher="Yezhegodnik Zoologicheskogo Muzeya Akademii Nauk SSSR Leningrad" pageId="0" pageNumber="429" pagination="347 - 355" refId="B4" refString="Ermolajev, WN, 1927. A new species from the genus Coelotes Blackwall (Araneae, Agelenidae) from Western Siberia. Yezhegodnik Zoologicheskogo Muzeya Akademii Nauk SSSR Leningrad 27: 347 - 355" title="A new species from the genus Coelotes Blackwall (Araneae, Agelenidae) from Western Siberia." volume="27" year="1927">Ermolajev 1927</bibRefCitation>
<bibRefCitation id="F917705DCE4F2747390531A19A920DA9" author="Ermolajev, WN" journalOrPublisher="Yezhegodnik Zoologicheskogo Muzeya Akademii Nauk SSSR Leningrad" pageId="0" pageNumber="429" pagination="347 - 355" refId="B4" refString="Ermolajev, WN, 1927. A new species from the genus Coelotes Blackwall (Araneae, Agelenidae) from Western Siberia. Yezhegodnik Zoologicheskogo Muzeya Akademii Nauk SSSR Leningrad 27: 347 - 355" title="A new species from the genus Coelotes Blackwall (Araneae, Agelenidae) from Western Siberia." volume="27" year="1927">Ermolajev 1927</bibRefCitation>
;
<bibRefCitation author="Marusik, YM" journalOrPublisher="Institute for Biological Problems of the North, Magadan" pageId="0" pageNumber="429" refId="B15" refString="Marusik, YM, Logunov, DV, Koponen, S., 2000. Spiders of Tuva, South Siberia. Institute for Biological Problems of the North, Magadan" title="Spiders of Tuva, South Siberia." year="2000">Marusik et al. 2000</bibRefCitation>
<bibRefCitation id="C5C55E213D405BFFC38D7922C1CD3A94" author="Marusik, YM" journalOrPublisher="Institute for Biological Problems of the North, Magadan" pageId="0" pageNumber="429" refId="B15" refString="Marusik, YM, Logunov, DV, Koponen, S., 2000. Spiders of Tuva, South Siberia. Institute for Biological Problems of the North, Magadan" title="Spiders of Tuva, South Siberia." year="2000">Marusik et al. 2000</bibRefCitation>
;
<bibRefCitation author="Azarkina, GN" pageId="0" pageNumber="429" refId="B1" refString="Azarkina, GN, Trilikauskas, LA, 2012. Spider fauna (Aranei) of the Russian Altai, part I: families Agelenidae, Araneidae, Clubionidae, Corinnidae, Dictynidae and Eresidae. Euroasian Entomological Journal 11 (3): 199-208, 212, pl. I." title="Spider fauna (Aranei) of the Russian Altai, part I: families Agelenidae, Araneidae, Clubionidae, Corinnidae, Dictynidae and Eresidae. Euroasian Entomological Journal 11 (3): 199 - 208, 212, pl. I." year="2012">Azarkina and Trilikauskas 2012</bibRefCitation>
<bibRefCitation id="51BE915E568D1F5D92EB64565A37EDA4" author="Azarkina, GN" pageId="0" pageNumber="429" refId="B1" refString="Azarkina, GN, Trilikauskas, LA, 2012. Spider fauna (Aranei) of the Russian Altai, part I: families Agelenidae, Araneidae, Clubionidae, Corinnidae, Dictynidae and Eresidae. Euroasian Entomological Journal 11 (3): 199-208, 212, pl. I." title="Spider fauna (Aranei) of the Russian Altai, part I: families Agelenidae, Araneidae, Clubionidae, Corinnidae, Dictynidae and Eresidae. Euroasian Entomological Journal 11 (3): 199 - 208, 212, pl. I." year="2012">Azarkina and Trilikauskas 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="429" type="notes">
<paragraph pageId="0" pageNumber="429">Notes.</paragraph>
<paragraph pageId="0" pageNumber="429">
<subSubSection id="849290F24F7902F240D2EE0F99B177F6" pageId="0" pageNumber="429" type="notes">
<paragraph id="54E90B3F1502AED2730872186CC579A8" pageId="0" pageNumber="429">Notes.</paragraph>
<paragraph id="F4C80E7C612DFC31637D05B06F070B73" pageId="0" pageNumber="429">
In the
<bibRefCitation author="WSC" journalOrPublisher="Zoologicheskyi Zhurnal" pageId="0" pageNumber="429" publicationUrl="http://wsc.nmbe.ch" refId="B24" refString="WSC, 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern, online at http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern, online at" url="http://wsc.nmbe.ch" year="2020">WSC (2020)</bibRefCitation>
<taxonomicName genus="P." lsidName="P. birulai" pageId="0" pageNumber="429" rank="species" species="birulai">
<emphasis italics="true" pageId="0" pageNumber="429">P. birulai</emphasis>
<bibRefCitation id="1A4C0BB6F989084971690EA6C4B6D38F" author="WSC" journalOrPublisher="Zoologicheskyi Zhurnal" pageId="0" pageNumber="429" publicationUrl="http://wsc.nmbe.ch" refId="B24" refString="WSC, 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern, online at http://wsc.nmbe.ch" title="World Spider Catalog. Version 21.5. Natural History Museum Bern, online at" url="http://wsc.nmbe.ch" year="2020">WSC (2020)</bibRefCitation>
<taxonomicName id="F964FB61831C176557072773A784C126" lsidName="P. birulai" pageId="0" pageNumber="429" rank="species" species="birulai">
<emphasis id="A01C3DFF47054A6854EF23DBE83EBF36" italics="true" pageId="0" pageNumber="429">P. birulai</emphasis>
</taxonomicName>
is listed as a junior synonym of
<taxonomicName genus="P." lsidName="P. luctuosa" pageId="0" pageNumber="429" rank="species" species="luctuosa">
<emphasis italics="true" pageId="0" pageNumber="429">P. luctuosa</emphasis>
<taxonomicName id="CD16775FE282772C673C75F00BFA3BCC" lsidName="P. luctuosa" pageId="0" pageNumber="429" rank="species" species="luctuosa">
<emphasis id="20C581751289D9CB7CCA105796E38B1C" italics="true" pageId="0" pageNumber="429">P. luctuosa</emphasis>
</taxonomicName>
(L. Koch, 1878).
<taxonomicName baseAuthorityName="L. Koch" baseAuthorityYear="1878" class="Arachnida" family="Agelenidae" genus="Pireneitega" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pireneitega luctuosa" order="Araneae" pageId="0" pageNumber="429" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis italics="true" pageId="0" pageNumber="429">Pireneitega luctuosa</emphasis>
<taxonomicName id="6C40AC27946312A607FC216C1AC61AD7" baseAuthorityName="L. Koch" baseAuthorityYear="1878" class="Arachnida" family="Agelenidae" genus="Pireneitega" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pireneitega luctuosa" order="Araneae" pageId="0" pageNumber="429" phylum="Arthropoda" rank="species" species="luctuosa">
<emphasis id="043F8127426BF31D4EA657B5739FF3F2" italics="true" pageId="0" pageNumber="429">Pireneitega luctuosa</emphasis>
</taxonomicName>
lives in caves in the southern China (
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<bibRefCitation id="0CB6149C00E3C1A0C737671C31EFA9B1" DOI="https://doi.org/10.3897/zookeys.663.11356" author="Zhang, XQ" journalOrPublisher="ZooKeys" pageId="0" pageNumber="429" pagination="45 - 64" refId="B25" refString="Zhang, XQ, Zhao, Z, Zheng, G, Li, SQ, 2017. A survey of five Pireneitega species (Agelenidae, Coelotinae) from China. ZooKeys 663: 45 - 64, DOI: https://doi.org/10.3897/zookeys.663.11356" title="A survey of five Pireneitega species (Agelenidae, Coelotinae) from China." url="https://doi.org/10.3897/zookeys.663.11356" volume="663" year="2017">Zhang et al. 2017</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3969/j.issn.1005-9628.2018.02.001" author="Jiang, XK" journalOrPublisher="Acta Arachnologica Sinica" pageId="0" pageNumber="429" pagination="65 - 95" refId="B8" refString="Jiang, XK, Chen, HM, Zhang, ZS, 2018. Spiders' diversity in Fanjing Mountain Nature Reserve, Guizhou, China, IV: Coelotine spiders (Araneae, Agelenidae). Acta Arachnologica Sinica 27 (2): 65 - 95, DOI: https://doi.org/10.3969/j.issn.1005-9628.2018.02.001" title="Spiders' diversity in Fanjing Mountain Nature Reserve, Guizhou, China, IV: Coelotine spiders (Araneae, Agelenidae)." url="https://doi.org/10.3969/j.issn.1005-9628.2018.02.001" volume="27" year="2018">Jiang et al. 2018</bibRefCitation>
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). Yu.M. Marusik (pers. comm.) studied the type specimens of both taxa and came to a conclusion that these are different species.
</paragraph>
</subSubSection>

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<taxonomicName id="39929EE577B2D7594D983BF8D56DBC82" authorityName="Kang" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus chiangmaiensis" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="chiangmaiensis">
<emphasis id="4DD029959B81A83AEAA57ED668C474ED" italics="true" pageId="0" pageNumber="67">Ophiclypeus chiangmaiensis</emphasis>
</taxonomicName>
Kang, sp. nov.
@ -99,7 +99,7 @@ Kang, sp. nov.
<paragraph id="3E19FF72FE23B0C952CF8C992D1A7EC7" pageId="0" pageNumber="67">Diagnosis.</paragraph>
<paragraph id="9EF55C580C6DA2024B4CDDB8AE3AE180" pageId="0" pageNumber="67">
Members of
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<taxonomicName id="2A0F5C246FC2EE0BE8228481F1AF5F11" authorityName="Kang" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="75F1C7E3DFCCFCF5B5CBD3B387B3CAB3" italics="true" pageId="0" pageNumber="67">Ophiclypeus</emphasis>
</taxonomicName>
gen. nov. are similar to members of
@ -123,7 +123,7 @@ Hedwig, 1957, and
<emphasis id="1FBDEBC4DAB6D954B72D02621C6161E8" italics="true" pageId="0" pageNumber="67">Retusigaster</emphasis>
</taxonomicName>
Dangerfield, Austin &amp; Whitfield, 1999 based on the following shared characters: ovipositor short, thick, and sharply downcurved; hypopygium short and not pointed apically. However, members of
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<taxonomicName id="90B51ABD9DE0EF947733E574A2D83A97" authorityName="Kang" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="0E7D792F0B16EA893949A60030D5E2D4" italics="true" pageId="0" pageNumber="67">Ophiclypeus</emphasis>
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gen. nov. can be distinguished from the members of the other five genera by the following combination of characters: a narrow face (Figs
@ -185,7 +185,7 @@ longer than hind tibia (Figs
<caption id="D78648301B00E01718003AC6F7582613" doi="10.3897/zookeys.1180.100106.figure1" httpUri="https://binary.pensoft.net/fig/906070" pageId="0" pageNumber="67" start="Figure 1" startId="F1">
<paragraph id="C8477A3CC7C29EE8AA68E93F254CFEB1" pageId="0" pageNumber="67">
<emphasis id="6397EF76BAD806787E4C96AF8C4E9647" bold="true" pageId="0" pageNumber="67">Figure 1.</emphasis>
<taxonomicName id="8446271E4FB0CDD492B54CE6ED66CF57" authorityName="Kang" authorityYear="2023" family="Braconidae" genus="Ophiclypeus" kingdom="Animalia" lsidName="Ophiclypeus chiangmaiensis" order="Hymenoptera" pageId="0" pageNumber="67" rank="species" species="chiangmaiensis">
<taxonomicName id="8446271E4FB0CDD492B54CE6ED66CF57" authorityName="Kang" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus chiangmaiensis" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="chiangmaiensis">
<emphasis id="F2F54E9E66549331D1972549885B8AD6" italics="true" pageId="0" pageNumber="67">Ophiclypeus chiangmaiensis</emphasis>
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sp. nov.
@ -208,7 +208,7 @@ lateral metasoma.
<caption id="07DCC20C4CD47D5DE594DE3448146BC8" doi="10.3897/zookeys.1180.100106.figure2" httpUri="https://binary.pensoft.net/fig/906071" pageId="0" pageNumber="67" start="Figure 2" startId="F2">
<paragraph id="F2DD34399FB67A3204011DCDCB267A45" pageId="0" pageNumber="67">
<emphasis id="06BE430772B43E7D8CFD38AD08587814" bold="true" pageId="0" pageNumber="67">Figure 2.</emphasis>
<taxonomicName id="7ABF82FA6B8B1144239A8FD6C12F8465" authorityName="Ghafouri Moghaddam, Quicke &amp; Butcher" authorityYear="2023" family="Braconidae" genus="Ophiclypeus" kingdom="Animalia" lsidName="Ophiclypeus dvaravati" order="Hymenoptera" pageId="0" pageNumber="67" rank="species" species="dvaravati">
<taxonomicName id="7ABF82FA6B8B1144239A8FD6C12F8465" authorityName="Ghafouri Moghaddam, Quicke &amp; Butcher" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus dvaravati" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="dvaravati">
<emphasis id="7761CCD69E4B020785DAA9B799C97F69" italics="true" pageId="0" pageNumber="67">Ophiclypeus dvaravati</emphasis>
</taxonomicName>
sp. nov.
@ -233,7 +233,7 @@ dorsal propodeum and metasoma.
<caption id="5690BD20309F583106F22DEB6AFC97E6" doi="10.3897/zookeys.1180.100106.figure3" httpUri="https://binary.pensoft.net/fig/906072" pageId="0" pageNumber="67" start="Figure 3" startId="F3">
<paragraph id="6C4F805FC559519FB703333979134046" pageId="0" pageNumber="67">
<emphasis id="2DD8CF1259C7AB48995554BE52855F8B" bold="true" pageId="0" pageNumber="67">Figure 3.</emphasis>
<taxonomicName id="D4B9CDA2A8DCB8A33FF894CE30B19E02" authorityName="Kang" authorityYear="2023" family="Braconidae" genus="Ophiclypeus" kingdom="Animalia" lsidName="Ophiclypeus junyani" order="Hymenoptera" pageId="0" pageNumber="67" rank="species" species="junyani">
<taxonomicName id="D4B9CDA2A8DCB8A33FF894CE30B19E02" authorityName="Kang" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus junyani" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="junyani">
<emphasis id="3DE87ED8DBD4B65046735899F5154BD3" italics="true" pageId="0" pageNumber="67">Ophiclypeus junyani</emphasis>
</taxonomicName>
sp. nov.
@ -320,7 +320,7 @@ Oriental region (Taiwan, Thailand) (Fig.
<paragraph id="DEE84A02DFDF3E5B8A3766A39FE780D8" pageId="0" pageNumber="67">
<emphasis id="60F617651CA7C75E6D613E7E4E3434D1" bold="true" pageId="0" pageNumber="67">Figure 4.</emphasis>
Distribution map of
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<taxonomicName id="F5365947626F8EA5EF79D1D5DBE3CCBD" authorityName="Kang" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="A54FFBA0812E144D7BD4F63D50DFF203" italics="true" pageId="0" pageNumber="67">Ophiclypeus</emphasis>
</taxonomicName>
species.
@ -409,7 +409,7 @@ longer than its height and without clypeal tubercles.
</td>
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<emphasis id="2217AD7FC6589ABA00972543EA141DC6" bold="true" pageId="0" pageNumber="67">
<taxonomicName id="20A86A98EBADF7E31B053352199CCF43" authorityName="Kang" authorityYear="2023" family="Braconidae" genus="Ophiclypeus" kingdom="Animalia" lsidName="Ophiclypeus" order="Hymenoptera" pageId="0" pageNumber="67" rank="genus">
<taxonomicName id="20A86A98EBADF7E31B053352199CCF43" authorityName="Kang" authorityYear="2023" class="Insecta" family="Braconidae" genus="Ophiclypeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiclypeus" order="Hymenoptera" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="6B021CF4BB004C54D979EE1A8008D940" italics="true" pageId="0" pageNumber="67">Ophiclypeus</emphasis>
</taxonomicName>
gen. nov.

210
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Holmb., Bot. Not. 1924: 326 (1924).Type Protologue: &quot;Nach ROUY kommt sie in Frankreich hier und da vor,aber ziemlich selten;
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hat sie nur von einem
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in Norddeutschland. In Schweden kommt sie besonders in Schonen vor; ich habe sie mehrmals eingesammelt, wie in der Gegend von Lund und
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an mehreren Orten;
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; Skartofta in
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; Gudmundtorp.
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sah ich Exemplare aus
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(leg. P. Boren 1903), Skelderviken (leg. Sten Selander), Kalmar (leg. N. Blomgren),
<normalizedToken originalValue="Borås">Boras</normalizedToken>
<normalizedToken id="FFC041C5F0BF2BDC1D5583B67C47285C" originalValue="Borås">Boras</normalizedToken>
(leg. C. Sandberg), Fyen, Stenlose (leg. G. Samuelsson). Kommt in Klee- und Grasfeldern, auf Rainen,
<normalizedToken originalValue="Wegrändern">Wegraendern</normalizedToken>
<normalizedToken id="188DC6F210B7574400729AF4E62929DC" originalValue="Wegrändern">Wegraendern</normalizedToken>
etc. vor, oft mit
<taxonomicName family="Poaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="B. mollis" order="Poales" pageId="6" pageNumber="59" rank="species" species="mollis">
<emphasis italics="true" pageId="6" pageNumber="59">B. mollis</emphasis>
<taxonomicName id="64E8346BC5F5CE4721FE77BC96132838" class="Liliopsida" family="Poaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="B. mollis" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="6C2EEFD16873C1E8473CAD152B367913" italics="true" pageId="6" pageNumber="59">B. mollis</emphasis>
</taxonomicName>
und
<taxonomicName family="Poaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="B. commutatus" order="Poales" pageId="6" pageNumber="59" rank="species" species="commutatus">
<emphasis italics="true" pageId="6" pageNumber="59">B. commutatus</emphasis>
<taxonomicName id="BD4F2095885E3AF246B5427ED5104EED" class="Liliopsida" family="Poaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="B. commutatus" order="Poales" pageId="6" pageNumber="59" phylum="Tracheophyta" rank="species" species="commutatus">
<emphasis id="2607863BEC02671F6CB87939AF4A5022" italics="true" pageId="6" pageNumber="59">B. commutatus</emphasis>
</taxonomicName>
zusammen; bei
<normalizedToken originalValue="Malmö">Malmoe</normalizedToken>
<normalizedToken id="26F30D3B162642327F6111944AC80119" originalValue="Malmö">Malmoe</normalizedToken>
auch als Ruderat&quot;. Type: SWEDEN. Scania:
<normalizedToken originalValue="Malmö">Malmoe</normalizedToken>
<normalizedToken id="1F3584D583EEB5D4CBAFD8FAA61616CD" originalValue="Malmö">Malmoe</normalizedToken>
, in ruderatis, 18-06-1920, Otto R.Holmberg, (lectotype, designated here: LD [LD1136595 image!]; isolectotype: K [K000913599!]). (Figure 7)
</paragraph>
<paragraph pageId="7" pageNumber="60">
<taxonomicName authority="Holmb." authorityName="Holmb" authorityYear="1924" class="Liliopsida" family="Poaceae" form="lasiolepis" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="f." kingdom="Plantae" lsidName="Bromus lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="form" species="lepidus">
<pageBreakToken pageId="7" pageNumber="60" start="start">Bromus</pageBreakToken>
<paragraph id="043C9FBDDAC137A743B10194FCD2E2C9" pageId="7" pageNumber="60">
<taxonomicName id="CCF684F44F7D72535B0CBF4F751A878E" authority="Holmb." authorityName="Holmb" authorityYear="1924" class="Liliopsida" family="Poaceae" form="lasiolepis" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="f." kingdom="Plantae" lsidName="Bromus lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="form" species="lepidus">
<pageBreakToken id="EA767BEDC130FCEEED669C6516A14DF9" pageId="7" pageNumber="60" start="start">Bromus</pageBreakToken>
lepidus Holmb. f. lasiolepis
</taxonomicName>
Holmb., Bot. Not. 1924: 326 (1924). Type Protologue: not indicated. Type: Sweden. Scania:
<normalizedToken originalValue="Vallkörra">Vallkoerra</normalizedToken>
<normalizedToken id="8ACEDC4D6FAC5826316DFE2E31913722" originalValue="Vallkörra">Vallkoerra</normalizedToken>
.
<normalizedToken originalValue="Svenenshög">Svenenshoeg</normalizedToken>
<normalizedToken id="2287C9A4C88C94D3FD8A8278F148692E" originalValue="Svenenshög">Svenenshoeg</normalizedToken>
. 4-07-1923. Otto R.Holmberg, (lectotype, designated here: LD [LD 1136235 image!]; isolectotype: K [K000913598!]). (Figure 8)
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="60" type="remarks">
<paragraph pageId="7" pageNumber="60">Remarks.</paragraph>
<paragraph pageId="7" pageNumber="60">
<subSubSection id="3F7153E5909B01562EF3ED8AFE9D08B5" pageId="7" pageNumber="60" type="remarks">
<paragraph id="185B406CC7C4729296A68B45AA8F8C57" pageId="7" pageNumber="60">Remarks.</paragraph>
<paragraph id="1337B2A45BDD429552FF078322C033A3" pageId="7" pageNumber="60">
This taxon has a long history full of nomenclatural problems.
<bibRefCitation author="Duval-Jouve, MJ" journalOrPublisher="Bulletin de la Societe Botanique de France" pageId="9" pageNumber="62" pagination="196 - 211" refId="B9" refString="Duval-Jouve, MJ, 1865. Variations paralelles des types congeneres. Bulletin de la Societe Botanique de France 12 (4): 196 - 211" title="Variations paralelles des types congeneres." volume="12" year="1865">Duval-Jouve (1865</bibRefCitation>
<bibRefCitation id="5DE0E615F11724BF1B4CEDF5D0E42D1C" author="Duval-Jouve, MJ" journalOrPublisher="Bulletin de la Societe Botanique de France" pageId="9" pageNumber="62" pagination="196 - 211" refId="B9" refString="Duval-Jouve, MJ, 1865. Variations paralelles des types congeneres. Bulletin de la Societe Botanique de France 12 (4): 196 - 211" title="Variations paralelles des types congeneres." volume="12" year="1865">Duval-Jouve (1865</bibRefCitation>
: 208), who only lists, without naming them, the variations in spikelet size and hairiness of
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="7" pageNumber="60">Bromus</emphasis>
<taxonomicName id="AB58BB57AECB3479779FB6E48237B52A" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="genus">
<emphasis id="F76A8E257C115B4EFBC3D446DE3C5519" italics="true" pageId="7" pageNumber="60">Bromus</emphasis>
</taxonomicName>
species, e.g.
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus mollis" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis italics="true" pageId="7" pageNumber="60">Bromus mollis</emphasis>
<taxonomicName id="F80041A143CEA08F8453E24CE0F2F438" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus mollis" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="34D248BDB50ED1A83C8128F67B870FDF" italics="true" pageId="7" pageNumber="60">Bromus mollis</emphasis>
</taxonomicName>
&quot;
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="microstachys" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="microstachys">microstachys</taxonomicName>
<taxonomicName id="662D1C182CD4BE8CA7CE987AD38291B3" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="microstachys" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="microstachys">microstachys</taxonomicName>
glabre&quot; and
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus mollis" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis italics="true" pageId="7" pageNumber="60">Bromus mollis</emphasis>
<taxonomicName id="8205BC2EBB0D1B0FAEA3060A1A8FCF18" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus mollis" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="mollis">
<emphasis id="419CE0FCCF9F0BC3D2FFCFFD62F2F865" italics="true" pageId="7" pageNumber="60">Bromus mollis</emphasis>
</taxonomicName>
&quot;
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="microstachys" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="microstachys">microstachys</taxonomicName>
<taxonomicName id="715DD3B7DC25718E1C76E49287A35DA8" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="microstachys" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="microstachys">microstachys</taxonomicName>
pubescent&quot;. This is the first mention of this taxon. Later
<bibRefCitation author="Rouy, GCH" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="10" pageNumber="63" refId="B26" refString="Rouy, GCH, 1913. Flore de France XIV. M.M. Deslis freres, Paris. http://bibdigital.rjb.csic.es/spa/Libro.php?Libro=876&amp;Pagina=1" title="Flore de France XIV. M. M. Deslis freres, Paris. http: // bibdigital. rjb. csic. es / spa / Libro. php? Libro = 876 &amp; Pagina = 1" year="1913">Rouy (1913</bibRefCitation>
<bibRefCitation id="5F7C8F6AC1040BF69E5C017BFF9DE38A" author="Rouy, GCH" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="10" pageNumber="63" refId="B26" refString="Rouy, GCH, 1913. Flore de France XIV. M.M. Deslis freres, Paris. http://bibdigital.rjb.csic.es/spa/Libro.php?Libro=876&amp;Pagina=1" title="Flore de France XIV. M. M. Deslis freres, Paris. http: // bibdigital. rjb. csic. es / spa / Libro. php? Libro = 876 &amp; Pagina = 1" year="1913">Rouy (1913</bibRefCitation>
: 236) proposed a named variety under the genus
<taxonomicName class="Liliopsida" family="Poaceae" genus="Serrafalcus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Serrafalcus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="7" pageNumber="60">Serrafalcus</emphasis>
<taxonomicName id="A6E22C8635181AC422F5EECEA9990A5E" class="Liliopsida" family="Poaceae" genus="Serrafalcus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Serrafalcus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="genus">
<emphasis id="ADF4222E2188B3B246A7E8C938A81615" italics="true" pageId="7" pageNumber="60">Serrafalcus</emphasis>
</taxonomicName>
Parl.:
<taxonomicName lsidName="S. mollis" pageId="7" pageNumber="60" rank="species" species="mollis">
<emphasis italics="true" pageId="7" pageNumber="60">S. mollis</emphasis>
<taxonomicName id="D3EF643FFA20F46D698C941C65098ED4" lsidName="S. mollis" pageId="7" pageNumber="60" rank="species" species="mollis">
<emphasis id="1F54096209C36101162BF497149E72A4" italics="true" pageId="7" pageNumber="60">S. mollis</emphasis>
</taxonomicName>
<normalizedToken originalValue="ß">ss</normalizedToken>
<taxonomicName lsidName="microstachys" pageId="7" pageNumber="60" rank="species" species="microstachys">
<emphasis italics="true" pageId="7" pageNumber="60">microstachys</emphasis>
<normalizedToken id="A6CFCE31BB2CC2F9A1DA9F6D2C40029D" originalValue="ß">ss</normalizedToken>
<taxonomicName id="9BB4A2C9B69E8857EEF25FB5DBC61253" lsidName="microstachys" pageId="7" pageNumber="60" rank="species" species="microstachys">
<emphasis id="AC01479BBD0C7DD502D0AD950BA1F5C0" italics="true" pageId="7" pageNumber="60">microstachys</emphasis>
</taxonomicName>
Rouy, giving &quot;
<taxonomicName class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus microstachys" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="microstachys">
<emphasis italics="true" pageId="7" pageNumber="60">Bromus microstachys</emphasis>
<taxonomicName id="2A5612AE0BAE2A6EBFD97A61DAC4C6C5" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus microstachys" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="microstachys">
<emphasis id="7940FC960967DD0B2112C7AD9FF4D402" italics="true" pageId="7" pageNumber="60">Bromus microstachys</emphasis>
</taxonomicName>
<bibRefCitation author="Duval-Jouve, MJ" journalOrPublisher="Bulletin de la Societe Botanique de France" pageId="9" pageNumber="62" pagination="196 - 211" refId="B9" refString="Duval-Jouve, MJ, 1865. Variations paralelles des types congeneres. Bulletin de la Societe Botanique de France 12 (4): 196 - 211" title="Variations paralelles des types congeneres." volume="12" year="1865">Duval-Jouve&quot; (1865</bibRefCitation>
<bibRefCitation id="9FEBEF3388DFBA8C01ECEB0A0DBFBCD9" author="Duval-Jouve, MJ" journalOrPublisher="Bulletin de la Societe Botanique de France" pageId="9" pageNumber="62" pagination="196 - 211" refId="B9" refString="Duval-Jouve, MJ, 1865. Variations paralelles des types congeneres. Bulletin de la Societe Botanique de France 12 (4): 196 - 211" title="Variations paralelles des types congeneres." volume="12" year="1865">Duval-Jouve&quot; (1865</bibRefCitation>
: 207) as a synonym and adding a diagnosis. Therefore, there is no doubt this is the same plant cited by Duval-Jouve (l. c.). Afterwards,
<bibRefCitation author="Kroesche, E" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="10" pageNumber="63" pagination="329 - 332" refId="B19" refString="Kroesche, E, 1924. Neue Arten und Formen aus Nordwest-Deutschland. Repertorium Specierum Novarum Regni Vegetabilis 19: 329 - 332" title="Neue Arten und Formen aus Nordwest-Deutschland." volume="19" year="1924">
<normalizedToken originalValue="Krösche">Kroesche</normalizedToken>
<bibRefCitation id="A29A8A64A5967A6FAAFC1DEBF9B72FFE" author="Kroesche, E" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="10" pageNumber="63" pagination="329 - 332" refId="B19" refString="Kroesche, E, 1924. Neue Arten und Formen aus Nordwest-Deutschland. Repertorium Specierum Novarum Regni Vegetabilis 19: 329 - 332" title="Neue Arten und Formen aus Nordwest-Deutschland." volume="19" year="1924">
<normalizedToken id="0D1B0950615F9A8649E7AE73A3010374" originalValue="Krösche">Kroesche</normalizedToken>
(1924
</bibRefCitation>
: 329) described
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. gracilis" order="Poales" pageId="7" pageNumber="60" rank="species" species="gracilis">
<emphasis italics="true" pageId="7" pageNumber="60">B. gracilis</emphasis>
<taxonomicName id="2A229423BEC702060D358AC27D3DE9DA" class="Liliopsida" family="Poaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="B. gracilis" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="520DF6D5230E212270AF5321F6022FB9" italics="true" pageId="7" pageNumber="60">B. gracilis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Krösche">Kroesche</normalizedToken>
<normalizedToken id="579AB311BBFF0ED06256E8830E94D598" originalValue="Krösche">Kroesche</normalizedToken>
, which unfortunately is a posterior homonym as
<bibRefCitation author="Weigel, CE" journalOrPublisher="Journal of Botany, British and Foreign" pageId="10" pageNumber="63" refId="B34" refString="Weigel, CE, 1772. Observationes Botanicae. Ant. Ferd. Roese. Gryphiae. https://doi.org/10.5962/bhl.title.123718" title="Observationes Botanicae. Ant. Ferd. Roese. Gryphiae. https: // doi. org / 10.5962 / bhl. title. 123718" year="1772">Weigel (1772</bibRefCitation>
<bibRefCitation id="9A30A15C923D0E4112043BD25404992A" author="Weigel, CE" journalOrPublisher="Journal of Botany, British and Foreign" pageId="10" pageNumber="63" refId="B34" refString="Weigel, CE, 1772. Observationes Botanicae. Ant. Ferd. Roese. Gryphiae. https://doi.org/10.5962/bhl.title.123718" title="Observationes Botanicae. Ant. Ferd. Roese. Gryphiae. https: // doi. org / 10.5962 / bhl. title. 123718" year="1772">Weigel (1772</bibRefCitation>
: 15) previously proposed this name for a different plant.
</paragraph>
<paragraph pageId="7" pageNumber="60">
<paragraph id="DD2E524EF4807DBD39D745448DE84AC4" pageId="7" pageNumber="60">
Finally,
<bibRefCitation author="Holmberg, OR" journalOrPublisher="Botaniska Notiser" pageId="9" pageNumber="62" pagination="313 - 328" refId="B14" refString="Holmberg, OR, 1924. Bromi molles, eine nomenklatorische und systematische Untersuchung. Botaniska Notiser 1924: 313 - 328" title="Bromi molles, eine nomenklatorische und systematische Untersuchung." volume="1924" year="1924">Holmberg (1924</bibRefCitation>
<bibRefCitation id="894C015345DE2A30D0DC694DD546E958" author="Holmberg, OR" journalOrPublisher="Botaniska Notiser" pageId="9" pageNumber="62" pagination="313 - 328" refId="B14" refString="Holmberg, OR, 1924. Bromi molles, eine nomenklatorische und systematische Untersuchung. Botaniska Notiser 1924: 313 - 328" title="Bromi molles, eine nomenklatorische und systematische Untersuchung." volume="1924" year="1924">Holmberg (1924</bibRefCitation>
: 326) solves this unfortunate situation and names simultaneously the new taxa
<taxonomicName authorityName="Holmb., Bot. Not. 1924: 326" authorityYear="1924" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="lepidus">
<emphasis italics="true" pageId="7" pageNumber="60">Bromus lepidus</emphasis>
<taxonomicName id="C98B20D9895FF443E96D0913A6222753" authorityName="Holmb., Bot. Not. 1924: 326" authorityYear="1924" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="lepidus">
<emphasis id="3271600236A9F708EE92A84B1AF6F965" italics="true" pageId="7" pageNumber="60">Bromus lepidus</emphasis>
</taxonomicName>
Holmb. and
<taxonomicName family="Poaceae" form="lasiolepis" infraspecific-rank="f." kingdom="Plantae" lsidName="B. lepidus" order="Poales" pageId="7" pageNumber="60" rank="form" species="lepidus">B. lepidus f. lasiolepis</taxonomicName>
<taxonomicName id="F3EFE204A84502289D3547C0C55D9CDB" class="Liliopsida" family="Poaceae" form="lasiolepis" higherTaxonomySource="CoL" infraspecific-rank="f." kingdom="Plantae" lsidName="B. lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="form" species="lepidus">B. lepidus f. lasiolepis</taxonomicName>
Holmb. The herbaria having Holmberg material are K, LD and S. There are 13 sheets in LD collected before 1924, five in S and three in K. It is reasonably certain that Holmberg studied all those plants before the description of the species and form and that all of them are original material. Therefore, we decided to limit our choices to sheets with the annotations &quot;
<taxonomicName family="Poaceae" kingdom="Plantae" lsidName="B. lepidus" order="Poales" pageId="7" pageNumber="60" rank="species" species="lepidus">
<emphasis italics="true" pageId="7" pageNumber="60">B. lepidus</emphasis>
<taxonomicName id="93A377E3AD382D8C157D9FDE3BA6A213" class="Liliopsida" family="Poaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="B. lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="lepidus">
<emphasis id="8681552A02490D25F9CD7982B5034E97" italics="true" pageId="7" pageNumber="60">B. lepidus</emphasis>
</taxonomicName>
mihi&quot; and &quot;
<taxonomicName family="Poaceae" form="lasiolepis" infraspecific-rank="f." kingdom="Plantae" lsidName="B. lepidus" order="Poales" pageId="7" pageNumber="60" rank="form" species="lepidus">B. lepidus f. lasiolepis</taxonomicName>
<taxonomicName id="57CDB5FA28E282FD2F07443864DA6410" class="Liliopsida" family="Poaceae" form="lasiolepis" higherTaxonomySource="CoL" infraspecific-rank="f." kingdom="Plantae" lsidName="B. lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="form" species="lepidus">B. lepidus f. lasiolepis</taxonomicName>
Holmb.&quot;, as this annotation indicates that Holmberg is interpreting them as the new taxa he is going to describe.
</paragraph>
<caption ID-Zenodo-Dep="2702127" doi="10.3897/phytokeys.119.30254.figure7" httpUri="https://binary.pensoft.net/fig/297554" pageId="7" pageNumber="60" start="Figure 7" startId="F7">
<paragraph pageId="7" pageNumber="60">
<emphasis bold="true" pageId="7" pageNumber="60">Figure 7.</emphasis>
<caption id="684D35E87D887F86C74C1C719FC58E90" ID-Zenodo-Dep="2702127" doi="10.3897/phytokeys.119.30254.figure7" httpUri="https://binary.pensoft.net/fig/297554" pageId="7" pageNumber="60" start="Figure 7" startId="F7">
<paragraph id="5AB547BAC600730A3550C3B9F2D29796" pageId="7" pageNumber="60">
<emphasis id="9A9BE95BA75C3421668307FCDC96A698" bold="true" pageId="7" pageNumber="60">Figure 7.</emphasis>
Lectotype of
<taxonomicName authorityName="Holmb., Bot. Not. 1924: 326" authorityYear="1924" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="lepidus">
<emphasis italics="true" pageId="7" pageNumber="60">Bromus lepidus</emphasis>
<taxonomicName id="AE82C61C4998825C68714372D337D78E" authorityName="Holmb., Bot. Not. 1924: 326" authorityYear="1924" class="Liliopsida" family="Gramineae" genus="Bromus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bromus lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="species" species="lepidus">
<emphasis id="CABE9CB49D87516B7E9C2CF25953A18A" italics="true" pageId="7" pageNumber="60">Bromus lepidus</emphasis>
</taxonomicName>
Holmberg, preserved at the Herbarium of the biological museum of the Lund University (LD1136295), reproduced with permission.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="2702131" doi="10.3897/phytokeys.119.30254.figure8" httpUri="https://binary.pensoft.net/fig/297555" pageId="7" pageNumber="60" start="Figure 8" startId="F8">
<paragraph pageId="7" pageNumber="60">
<emphasis bold="true" pageId="7" pageNumber="60">Figure 8.</emphasis>
<caption id="35CF9A249547DDB33638C3B92E889228" ID-Zenodo-Dep="2702131" doi="10.3897/phytokeys.119.30254.figure8" httpUri="https://binary.pensoft.net/fig/297555" pageId="7" pageNumber="60" start="Figure 8" startId="F8">
<paragraph id="46D4D3B0914AB5B1C7A8DC2B3C3B75DF" pageId="7" pageNumber="60">
<emphasis id="FCF46E0A0380D92110ECF18F36818403" bold="true" pageId="7" pageNumber="60">Figure 8.</emphasis>
Lectotype of
<taxonomicName authorityName="Holmb" authorityYear="1924" class="Liliopsida" family="Gramineae" form="lasiolepis" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="f." kingdom="Plantae" lsidName="Bromus lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="form" species="lepidus">Bromus lepidus f. lasiolepis</taxonomicName>
<taxonomicName id="FB0BD42FDB61DACFCA5D11209DF96CA6" authorityName="Holmb" authorityYear="1924" class="Liliopsida" family="Gramineae" form="lasiolepis" genus="Bromus" higherTaxonomySource="CoL" infraspecific-rank="f." kingdom="Plantae" lsidName="Bromus lepidus" order="Poales" pageId="7" pageNumber="60" phylum="Tracheophyta" rank="form" species="lepidus">Bromus lepidus f. lasiolepis</taxonomicName>
Holmberg, preserved at the Herbarium of the Biological Museum of Lund University (LD 1136235). Reproduced with permission.
</paragraph>
</caption>

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