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<mods:title id="1A632416157A022C30577C69F39D5E59">Review of the genus Tersilochus Holmgren (Hymenoptera, Ichneumonidae, Tersilochinae) from South Korea</mods:title>
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<mods:namePart>Khalaim, Andrey I.</mods:namePart>
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<mods:namePart>Balueva, Ekaterina N.</mods:namePart>
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<taxonomicName LSID="C4D4CB97-F4F2-58B5-B27D-E4C5517B2FDE" authority="Khalaim, 2011" authorityName="Khalaim" authorityYear="2011" class="Hexapoda" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tersilochus (Tersilochus) granulatus" order="Hymenoptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="granulatus" subGenus="Tersilochus">Tersilochus (Tersilochus) granulatus Khalaim, 2011</taxonomicName>
<figureCitation captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">Figs 15-17</figureCitation>
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<figureCitation id="BF6819F728DA7CD846C806A3ABD755DB" captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">Figs 15-17</figureCitation>
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<paragraph pageId="7" pageNumber="34">Comparison.</paragraph>
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<paragraph id="5C5812F4B4A02F41FF297B9FBBD878C6" pageId="7" pageNumber="34">Comparison.</paragraph>
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Differs from other Korean species of the genus by an evenly granulate propodeum with basal keel (
<figureCitation captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">Fig. 17</figureCitation>
<figureCitation id="E8094537153E4DBB8CA191B9DDAF3AF2" captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">Fig. 17</figureCitation>
), very slender brown first tergite (
<figureCitation captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">Figs 15</figureCitation>
<figureCitation id="230EF65681129013D9820179338A4F1D" captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">Figs 15</figureCitation>
,
<figureCitation captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">17</figureCitation>
<figureCitation id="819CB3DB31049A411035FCD5664442EE" captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">17</figureCitation>
), and distinctly elongate second tergite (
<figureCitation captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">Fig. 17</figureCitation>
<figureCitation id="DDC7E7CCE828E53BE30107F6CCF9A5CD" captionStart="Figures 1522" captionStartId="F3" captionText="Figures 15 - 22. Tersilochus granulatus Khalaim, female, holotype (Figs 15, 16) and paratype (Fig. 17): 15 habitus (without apices of wings and ovipositor), lateral view 16 head and anterior part of mesoscutum, dorsal view 17 propodeum and base of metasoma, dorsal view. Tersilochus iracundus sp. n., female, holotype: 18 habitus (without wings), lateral view 19 head and anterior part of mesoscutum, dorsal view 20 antenna, lateral view 21 base of antenna, lateral view 22 propodeum and hind coxae, dorsal view." httpUri="https://binary.pensoft.net/fig/11869" pageId="7" pageNumber="34">Fig. 17</figureCitation>
).
</paragraph>
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<emphasis bold="true" pageId="7" pageNumber="34">Figures 15-22.</emphasis>
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<emphasis italics="true" pageId="7" pageNumber="34">Tersilochus granulatus</emphasis>
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Khalaim, female, holotype (Figs
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,
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<emphasis id="3DA772C3E4DE3CA5C9C1A060EB08C646" bold="true" pageId="7" pageNumber="34">16</emphasis>
) and paratype (Fig.
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):
<emphasis bold="true" pageId="7" pageNumber="34">15</emphasis>
<emphasis id="FEB5AE7B8BB1A8353868F4F554A8C04B" bold="true" pageId="7" pageNumber="34">15</emphasis>
habitus (without apices of wings and ovipositor), lateral view
<emphasis bold="true" pageId="7" pageNumber="34">16</emphasis>
<emphasis id="77DD02EFF28A586A57FDD81C7BD16BC0" bold="true" pageId="7" pageNumber="34">16</emphasis>
head and anterior part of mesoscutum, dorsal view
<emphasis bold="true" pageId="7" pageNumber="34">17</emphasis>
<emphasis id="9F18DAE1E0E86104B3302C52C95F3A62" bold="true" pageId="7" pageNumber="34">17</emphasis>
propodeum and base of metasoma, dorsal view.
<taxonomicName authorityName="Khalaim &amp; Lee" authorityYear="2014" class="Hexapoda" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="iracundus">
<emphasis italics="true" pageId="7" pageNumber="34">Tersilochus iracundus</emphasis>
<taxonomicName id="D56403B34383DDD4CAE781C3F134C27F" authorityName="Khalaim &amp; Lee" authorityYear="2014" class="Hexapoda" family="Ichneumonidae" genus="Tersilochus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="iracundus">
<emphasis id="1CADB8B0A0DDC53DFC41BAD044278FCA" italics="true" pageId="7" pageNumber="34">Tersilochus iracundus</emphasis>
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sp. n., female, holotype:
<emphasis bold="true" pageId="7" pageNumber="34">18</emphasis>
<emphasis id="4AC88EA64BA71232C715F217B76E322B" bold="true" pageId="7" pageNumber="34">18</emphasis>
habitus (without wings), lateral view
<emphasis bold="true" pageId="7" pageNumber="34">19</emphasis>
<emphasis id="4A14B51046B0378208E187DDDFBF82AF" bold="true" pageId="7" pageNumber="34">19</emphasis>
head and anterior part of mesoscutum, dorsal view
<emphasis bold="true" pageId="7" pageNumber="34">20</emphasis>
<emphasis id="07EDCA951902125635DD74B8F41DF47F" bold="true" pageId="7" pageNumber="34">20</emphasis>
antenna, lateral view
<emphasis bold="true" pageId="7" pageNumber="34">21</emphasis>
<emphasis id="7F013A3C10904242481FD91C2D2E532D" bold="true" pageId="7" pageNumber="34">21</emphasis>
base of antenna, lateral view
<emphasis bold="true" pageId="7" pageNumber="34">22</emphasis>
<emphasis id="C631AEA9491FCA8C80DC4CB15E768483" bold="true" pageId="7" pageNumber="34">22</emphasis>
propodeum and hind coxae, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="7" pageNumber="34" type="remarks">
<paragraph pageId="7" pageNumber="34">Remarks.</paragraph>
<paragraph pageId="7" pageNumber="34">Flagellum with finger-shaped subapical structures present on outer surface of flagellomeres 4 to 6; these structures in this species are very small, inconspicuous, and hardly visible under a light microscope.</paragraph>
<subSubSection id="E7AA40F8E926E3072F91A969459EE1E8" pageId="7" pageNumber="34" type="remarks">
<paragraph id="60903DF88636995656DA426F7EE8DF49" pageId="7" pageNumber="34">Remarks.</paragraph>
<paragraph id="C971C80A112A0BCA79D0A7F15D9D959B" pageId="7" pageNumber="34">Flagellum with finger-shaped subapical structures present on outer surface of flagellomeres 4 to 6; these structures in this species are very small, inconspicuous, and hardly visible under a light microscope.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="34" type="distribution">
<paragraph pageId="7" pageNumber="34">Distribution.</paragraph>
<paragraph pageId="7" pageNumber="34">South Korea: Gyeongsangnam-do (GN).</paragraph>
<subSubSection id="B2E9B637B203D06E9CBB6352747E3746" pageId="7" pageNumber="34" type="distribution">
<paragraph id="7B201539B24BB4CE67B4BC8BC5730A99" pageId="7" pageNumber="34">Distribution.</paragraph>
<paragraph id="77E58D6BBA34CFB4898A4EFEBA0F7DFC" pageId="7" pageNumber="34">South Korea: Gyeongsangnam-do (GN).</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,49 +1,50 @@
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@ -118,7 +119,7 @@ is a member of Taxonina of Allantini,
<emphasis id="73F86635FFEAFFCFFB43F8E5FAB9EBED" box="[1195,1338,1908,1932]" italics="true" pageId="1" pageNumber="2">Emphytopsis</emphasis>
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is close to
<taxonomicName id="868CC1A4FFEAFFCFFF50F809FD8AEBD1" authority="Cameron, 1876" authorityName="Cameron" authorityYear="1876" box="[184,521,1944,1968]" class="Insecta" family="Tenthredinidae" genus="Allomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<taxonomicName id="868CC1A4FFEAFFCFFF50F809FD8AEBD1" authority="Cameron, 1876" authorityName="Cameron" authorityYear="1876" box="[184,521,1944,1968]" class="Insecta" genus="Allomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="73F86635FFEAFFCFFF50F809FEC6EBD1" box="[184,325,1944,1968]" italics="true" pageId="1" pageNumber="2">Allomorpha</emphasis>
<bibRefCitation id="251DC7D6FFEAFFCFFEA6F809FD8AEBD1" author="Cameron" box="[334,521,1944,1968]" pageId="1" pageNumber="2" refString="Cameron, P. (1876) Descriptions of new genera and species of Tenthredinidae and Siricidae, chiefly from the East Indies, in the Collection of the British Museum. Transactions of the Entomological Society of London, P. III, 459 - 471." type="book chapter" year="1876">
<collectingCountry id="399BFAB7FFEAFFCFFEA6F809FE3EEBD1" box="[334,445,1944,1968]" name="Cameroon" pageId="1" pageNumber="2">Cameron</collectingCountry>
@ -130,11 +131,11 @@ is close to
<emphasis id="73F86635FFEAFFCFFDFFF809FD25EBD1" box="[535,678,1944,1968]" italics="true" pageId="1" pageNumber="2">Emphytopsis</emphasis>
</taxonomicName>
differs from
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<taxonomicName id="868CC1A4FFEAFFCFFCA8F809FC4FEBD1" box="[832,972,1944,1968]" class="Insecta" genus="Allomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="73F86635FFEAFFCFFCA8F809FC4FEBD1" box="[832,972,1944,1968]" italics="true" pageId="1" pageNumber="2">Allomorpha</emphasis>
</taxonomicName>
in antenna short and slender, more or less tapering toward apex, middle white ring absent; hind wing with cell Rs open, anal cell petiolate; each lobe of the first abdominal tergite subtriangular, membranous blotch large; serrulae each with a large inner subbasal tooth; lateral lobes of clypeus asymmetric; body yellow with conspicuous black maculae. In
<taxonomicName id="868CC1A4FFE9FFCCFA89FF2DFF71EC99" class="Insecta" family="Tenthredinidae" genus="Allomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<taxonomicName id="868CC1A4FFE9FFCCFA89FF2DFF71EC99" class="Insecta" genus="Allomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="73F86635FFE9FFCCFA89FF2DFF71EC99" italics="true" pageId="2" pageNumber="3">Allomorpha</emphasis>
</taxonomicName>
, the antenna stout with a white ring at middle, middle and subapical antennomeres distinctly compressed and dilated; hind wing with cell Rs closed, anal cell sessile; lobes of the first abdominal tergite trapezoidal, membranous blotch small; serrulae without inner subbasal tooth; lateral lobes of clypeus subsymmetric; body usually black with pale maculae.
@ -155,7 +156,7 @@ Both
<emphasis id="73F86635FFE9FFCCFEE2FE05FE1AEDCD" box="[266,409,404,428]" italics="true" pageId="2" pageNumber="3">Emphytopsis</emphasis>
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and
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<taxonomicName id="868CC1A4FFE9FFCCFE3DFE05FDE1EDCD" box="[469,610,404,428]" class="Insecta" genus="Allomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="73F86635FFE9FFCCFE3DFE05FDE1EDCD" box="[469,610,404,428]" italics="true" pageId="2" pageNumber="3">Allomorpha</emphasis>
</taxonomicName>
are endemic to E. Asia. The two genera have a similar distribution pattern (
@ -198,7 +199,7 @@ comprehensive and maybe polyphyletic:
<emphasis id="73F86635FFE9FFCCFD52FDD9FC9DEE01" box="[698,798,584,608]" italics="true" pageId="2" pageNumber="3">hirasana</emphasis>
<bibRefCitation id="251DC7D6FFE9FFCCFCC2FDD6FC5FEE3E" author="Takeuchi" box="[810,988,583,607]" pageId="2" pageNumber="3" refString="Takeuchi, K. (1929) Descriptions of new sawflies from the Japanese Empire. (I). Transaction of the Natural History Society of Formosa, 105, 495 - 520." type="journal article" year="1929">Takeuchi, 1929</bibRefCitation>
(a species of
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<taxonomicName id="868CC1A4FFE9FFCCFB64FDD9FF67EEE5" authority="Takeuchi, 1929" authorityName="Takeuchi" authorityYear="1929" class="Insecta" genus="Allomorpha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="73F86635FFE9FFCCFB64FDD9FA91EE01" box="[1164,1298,584,608]" italics="true" pageId="2" pageNumber="3">Allomorpha</emphasis>
,
<bibRefCitation id="251DC7D6FFE9FFCCFACDFDD6FF67EEE5" author="Takeuchi" pageId="2" pageNumber="3" refString="Takeuchi, K. (1929) Descriptions of new sawflies from the Japanese Empire. (I). Transaction of the Natural History Society of Formosa, 105, 495 - 520." type="journal article" year="1929">(Takeuchi, 1929))</bibRefCitation>

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@ -1,96 +1,96 @@
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<taxonomicName id="D7612147A58C11A9363E5566DE2ECBC2" authorityName="Friese" authorityYear="1895" class="Insecta" family="Apidae" genus="Epeolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epeolus fasciatus" order="Hymenoptera" pageId="0" pageNumber="191" phylum="Arthropoda" rank="species" species="fasciatus">Epeolus fasciatus</taxonomicName>
Friese, 1895: 208, ♀, ♂ (type locality: Hungary, Budapest [Pest]; Museum
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Naturkunde, Berlin;
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National
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Naturelle, Paris; ZISP).
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<paragraph pageId="0" pageNumber="191">Material examined.</paragraph>
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<paragraph id="A3D4A7D79D2764302969841BCE181FC8" pageId="0" pageNumber="191">Material examined.</paragraph>
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<materialsCitation id="138D9200B65919AEFEAB50B89CE79300" collectingDate="1886-07-02" country="Hungary" location="Pest" specimenCount="1" specimenCount-male="1" stateProvince="Pest" typeStatus="syntype">
<collectingCountry id="EE59B869778819A4B31C5F38206BE680" name="Hungary">Hungary</collectingCountry>
,
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[
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<collectingRegion id="5EA3BD7234F432BF1F20515434294D6A" country="Hungary" name="Budapest">Budapest</collectingRegion>
],
<collectingDate value="1886-07-02">2.VII.1886</collectingDate>
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(
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), Friese [ZISP]
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.
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<paragraph pageId="0" pageNumber="191">Distribution.</paragraph>
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<paragraph id="8354EDE11B3B27AED44649DD497877DA" pageId="0" pageNumber="191">Distribution.</paragraph>
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Southern and Central Europe, Turkey (
<bibRefCitation DOI="https://doi.org/10.2478/aemnp-2018-0010" author="Bogusch, P" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="191" pagination="127 - 135" refId="B6" refString="Bogusch, P, 2018. Three new species and new records of cuckoo bees of the genus Epeolus in Turkey (Hymenoptera: Apidae: Nomadinae). Acta Entomologica Musei Nationalis Pragae 58: 127 - 135, DOI: https://doi.org/10.2478/aemnp-2018-0010" title="Three new species and new records of cuckoo bees of the genus Epeolus in Turkey (Hymenoptera: Apidae: Nomadinae)." url="https://doi.org/10.2478/aemnp-2018-0010" volume="58" year="2018">Bogusch 2018</bibRefCitation>
<bibRefCitation id="367592255FE0CB3E489EB54F23DC1F0B" DOI="https://doi.org/10.2478/aemnp-2018-0010" author="Bogusch, P" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="191" pagination="127 - 135" refId="B6" refString="Bogusch, P, 2018. Three new species and new records of cuckoo bees of the genus Epeolus in Turkey (Hymenoptera: Apidae: Nomadinae). Acta Entomologica Musei Nationalis Pragae 58: 127 - 135, DOI: https://doi.org/10.2478/aemnp-2018-0010" title="Three new species and new records of cuckoo bees of the genus Epeolus in Turkey (Hymenoptera: Apidae: Nomadinae)." url="https://doi.org/10.2478/aemnp-2018-0010" volume="58" year="2018">Bogusch 2018</bibRefCitation>
).
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@ -234,7 +234,7 @@ Unknown.
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sp. nov. is known from Saraburi, Thailand (Fig.

View file

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<paragraph id="8145D07D11DBEC9D2B4EDEFCD962C251" pageId="21" pageNumber="52">Distribution.</paragraph>
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The species is distributed in central part of Russian sector of Arctic. From the Far East it is known only in Chaunski district (Chaun Bay) of Chukotka, as &quot;cf. britannica&quot; by
<bibRefCitation author="MacLean, SF" journalOrPublisher="Arctic and Alpine Research" pageId="22" pageNumber="53" pagination="559 - 568" title="Soil acari and Collembola from Chaun Bay, Northern Chukotka." url="https://doi.org/10.2307/1550679" volume="10" year="1978">MacLean et al. (1978)</bibRefCitation>
<bibRefCitation id="91C92231BFFC0C4A16FB8ABF04AC892E" author="MacLean, SF" journalOrPublisher="Arctic and Alpine Research" pageId="22" pageNumber="53" pagination="559 - 568" title="Soil acari and Collembola from Chaun Bay, Northern Chukotka." url="https://doi.org/10.2307/1550679" volume="10" year="1978">MacLean et al. (1978)</bibRefCitation>
, which is the most eastern record of the species so far.
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View file

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<taxonomicName id="8CBD510B27DB0C81BFBF81ADFE61CC04" authorityName="Sars" authorityYear="1869" class="Polychaeta" family="Amphinomidae" genus="Paramphinome" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paramphinome pulchella" order="Amphinomida" pageId="0" pageNumber="33" phylum="Annelida" rank="species" species="pulchella">
<emphasis id="969DBF0A2D7FEE112AF3FB104D282ECE" italics="true" pageId="0" pageNumber="33">Paramphinome pulchella</emphasis>
</taxonomicName>
M. Sars in G. Sars, 1872.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="diagnosis">
<paragraph pageId="0" pageNumber="33">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="33">Small but long long-bodied forms. Prostomium posteriorly with Y-shaped or elongated caruncle. Branchiae comb-shaped, limited to the anterior chaetigers. First chaetiger with curved hooks in notopodia.</paragraph>
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<paragraph id="9D1EB7BECDB9ADA040CCF603F4FADC37" pageId="0" pageNumber="33">Diagnosis.</paragraph>
<paragraph id="3F76B19D32087B9C064839EDF54C6AC2" pageId="0" pageNumber="33">Small but long long-bodied forms. Prostomium posteriorly with Y-shaped or elongated caruncle. Branchiae comb-shaped, limited to the anterior chaetigers. First chaetiger with curved hooks in notopodia.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,169 +1,169 @@
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Rhizoplaca occulta S. Leavitt, F.
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, Lumbsch, Sohrabi &amp; L. St. Clair
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<paragraph pageId="7" pageNumber="8">Type.</paragraph>
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<paragraph id="7529E809E6737FBAFDD4D2F08E92EE76" pageId="7" pageNumber="8">Type.</paragraph>
<paragraph id="41034D47EA5F649F15BAB1F25DD8DFC3" pageId="7" pageNumber="8">
USA, Nevada, White Pine County, on Cave Mountain, north of antenna site, Humboldt-Toiyabe National Forest,
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,
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<geoCoordinate id="347BEBCC100637B173E3A3FF33AB2492" direction="west" orientation="longitude" precision="5" value="-114.613">114.6130°W</geoCoordinate>
, on basalt, 3150 m alt., July 2010, S. D. Leavitt &amp; Larry L. St. Clair BRY-C55076 (holotype BRY).
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<paragraph pageId="7" pageNumber="8">Description.</paragraph>
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<paragraph id="9A746E2B3C207192EB5D6C845B5754E8" pageId="7" pageNumber="8">Description.</paragraph>
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Consists of specimens recovered within 'clade
<normalizedToken originalValue="IVa">IVa'</normalizedToken>
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<bibRefCitation author="Leavitt, SD" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="16" pageNumber="17" pagination="587 - 602" title="Complex patterns of speciation in cosmopolitan '' rock posy' ' lichens - Discovering and delimiting cryptic fungal species in the lichen-forming Rhizoplaca melanophthalma species-complex (Lecanoraceae, Ascomycota)." url="10.1016/j.ympev.2011.03.020" volume="59" year="2011 a">Leavitt et al. (2011a)</bibRefCitation>
<bibRefCitation id="553DB73825F2EC4FAE882229520B0718" author="Leavitt, SD" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="16" pageNumber="17" pagination="587 - 602" title="Complex patterns of speciation in cosmopolitan '' rock posy' ' lichens - Discovering and delimiting cryptic fungal species in the lichen-forming Rhizoplaca melanophthalma species-complex (Lecanoraceae, Ascomycota)." url="10.1016/j.ympev.2011.03.020" volume="59" year="2011 a">Leavitt et al. (2011a)</bibRefCitation>
, which is supported as a lineage distinct from all other populations according to coalescent-based genetic analysis of multiple genetic loci. This species is morphologically variable. While some individuals are morphologically similar to
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca melanophthalma" order="Lecanorales" pageId="7" pageNumber="8" phylum="Ascomycota" rank="species" species="melanophthalma">Rhizoplaca melanophthalma</taxonomicName>
<taxonomicName id="59BFFCFAD0E52579D95F064EC552B908" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca melanophthalma" order="Lecanorales" pageId="7" pageNumber="8" phylum="Ascomycota" rank="species" species="melanophthalma">Rhizoplaca melanophthalma</taxonomicName>
sensu stricto, vagrant forms, including
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca cerebriformis" order="Lecanorales" pageId="7" pageNumber="8" phylum="Ascomycota" rank="species" species="cerebriformis">Rhizoplaca cerebriformis</taxonomicName>
<taxonomicName id="9FF44985428BB65E75F863A743A6BC3B" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca cerebriformis" order="Lecanorales" pageId="7" pageNumber="8" phylum="Ascomycota" rank="species" species="cerebriformis">Rhizoplaca cerebriformis</taxonomicName>
ined. and
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca subidahoensis" order="Lecanorales" pageId="7" pageNumber="8" phylum="Ascomycota" rank="species" species="subidahoensis">Rhizoplaca subidahoensis</taxonomicName>
<taxonomicName id="7E170D7EA88D462BE9D167CF3EF3FD48" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca subidahoensis" order="Lecanorales" pageId="7" pageNumber="8" phylum="Ascomycota" rank="species" species="subidahoensis">Rhizoplaca subidahoensis</taxonomicName>
ined. which have been suggested to be distinct species based on morphology (
<bibRefCitation author="Arup, U" journalOrPublisher="The Lichenologist" pageId="14" pageNumber="15" title="Is Rhizoplaca (Lecanorales, lichenized Ascomycota) a monophyletic genus? Canadian Journal of Botany 78: 318 - 327." url="10.1139/b00-006" year="2000">Arup and Grube 2000</bibRefCitation>
<bibRefCitation id="039F6BF5651F32ECEEFF305203867271" author="Arup, U" journalOrPublisher="The Lichenologist" pageId="14" pageNumber="15" title="Is Rhizoplaca (Lecanorales, lichenized Ascomycota) a monophyletic genus? Canadian Journal of Botany 78: 318 - 327." url="10.1139/b00-006" year="2000">Arup and Grube 2000</bibRefCitation>
; figs 7 &amp; 9), also belong within this species. The mean genetic distance among ITS haplotypes was estimated to be 0.003
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0.001.
</paragraph>
<paragraph pageId="7" pageNumber="8">Chemistry - Usnic (major), constipatic (minor), dehydroconstipatic (minor), 2'-O-demethylpsoromic (minor), and 2'-O-demethylsubpsoromic (minor or trace) acids; usually with psoromic acid (major); and occasionally with dehydroprotocetraric acid (minor).</paragraph>
<paragraph id="621865587196E058A7DF8D15591ECE74" pageId="7" pageNumber="8">Chemistry - Usnic (major), constipatic (minor), dehydroconstipatic (minor), 2'-O-demethylpsoromic (minor), and 2'-O-demethylsubpsoromic (minor or trace) acids; usually with psoromic acid (major); and occasionally with dehydroprotocetraric acid (minor).</paragraph>
</subSubSection>
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<paragraph pageId="7" pageNumber="8">Reference phylogeny.</paragraph>
<paragraph pageId="7" pageNumber="8">
<bibRefCitation author="Leavitt, SD" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="16" pageNumber="17" pagination="587 - 602" title="Complex patterns of speciation in cosmopolitan '' rock posy' ' lichens - Discovering and delimiting cryptic fungal species in the lichen-forming Rhizoplaca melanophthalma species-complex (Lecanoraceae, Ascomycota)." url="10.1016/j.ympev.2011.03.020" volume="59" year="2011 a">Leavitt et al. 2011a</bibRefCitation>
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<paragraph id="D4F1CAF9172AF9520CAEFDF31BCD1B21" pageId="7" pageNumber="8">Reference phylogeny.</paragraph>
<paragraph id="056F98DF0ABB3232AA0DD91FEAFC88A6" pageId="7" pageNumber="8">
<bibRefCitation id="C66A0E8624B5843B3C81B47C4F6A9A25" author="Leavitt, SD" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="16" pageNumber="17" pagination="587 - 602" title="Complex patterns of speciation in cosmopolitan '' rock posy' ' lichens - Discovering and delimiting cryptic fungal species in the lichen-forming Rhizoplaca melanophthalma species-complex (Lecanoraceae, Ascomycota)." url="10.1016/j.ympev.2011.03.020" volume="59" year="2011 a">Leavitt et al. 2011a</bibRefCitation>
(fig. 5, 'clade
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</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="reference sequences">
<paragraph pageId="7" pageNumber="8">Reference sequences.</paragraph>
<paragraph pageId="7" pageNumber="8">GenBank Nos. HM577307 (ITS), HM577081 (IGS), HM577210 (group I intron), HM577441 (MCM 7), and HM576952 (β-tubulin).</paragraph>
<paragraph pageId="8" pageNumber="9">
<pageBreakToken pageId="8" pageNumber="9" start="start">Phylogenetic</pageBreakToken>
<subSubSection id="0A6303A65501CC9578A6FAB10D97DE04" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="reference sequences">
<paragraph id="8BD771D82F740F9EBEE7CA38CD9AD16B" pageId="7" pageNumber="8">Reference sequences.</paragraph>
<paragraph id="1FBE2F1CA6CF43FF22512CC93619CCE6" pageId="7" pageNumber="8">GenBank Nos. HM577307 (ITS), HM577081 (IGS), HM577210 (group I intron), HM577441 (MCM 7), and HM576952 (β-tubulin).</paragraph>
<paragraph id="E1FB9572CCFB480D13E34A62FF0291D3" pageId="8" pageNumber="9">
<pageBreakToken id="A0C284874D6B3E8CB74AA4DD50E655A7" pageId="8" pageNumber="9" start="start">Phylogenetic</pageBreakToken>
notes: Strongly supported as monophyletic lineage in both concatenated multilocus gene tree (ML bootstrap = 100%: posterior probability = 1.0) and the ITS gene topology (ML bootstrap = 99%, this study); and strong speciation probability inferred from multiple loci (BPP speciation probability ≥ 0.98).
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca occulta" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="occulta">Rhizoplaca occulta</taxonomicName>
<taxonomicName id="2BF46A7996C3CFB62616A91F06186808" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca occulta" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="occulta">Rhizoplaca occulta</taxonomicName>
belongs to a closely related, and well-supported, monophyletic lineage including
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca paralis" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="paralis">Rhizoplaca paralis</taxonomicName>
<taxonomicName id="6D6740ADE8A30BA3266122D1A275D0BA" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca paralis" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="paralis">Rhizoplaca paralis</taxonomicName>
,
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca polymorpha" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="polymorpha">Rhizoplaca polymorpha</taxonomicName>
<taxonomicName id="6F29A9B6BAB1EAFE19E95FBA16E7825D" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca polymorpha" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="polymorpha">Rhizoplaca polymorpha</taxonomicName>
,
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca porterii" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="porterii">Rhizoplaca porterii</taxonomicName>
<taxonomicName id="B4BA9D5331D4AB102AA20FA6A598FF07" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca porterii" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="porterii">Rhizoplaca porterii</taxonomicName>
, and the obligatory vagrant species
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca haydenii" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="haydenii">Rhizoplaca haydenii</taxonomicName>
<taxonomicName id="8C2F4F8698A1821EF9B72DDE7BDC8BA6" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca haydenii" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="haydenii">Rhizoplaca haydenii</taxonomicName>
and
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca idahoensis" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="idahoensis">Rhizoplaca idahoensis</taxonomicName>
<taxonomicName id="3C523FB20FFB691E5602813CD6CA4F42" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca idahoensis" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="idahoensis">Rhizoplaca idahoensis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="ecology and distribution">
<paragraph pageId="8" pageNumber="9">Ecology and distribution.</paragraph>
<paragraph pageId="8" pageNumber="9">
<subSubSection id="34B39CDCECEFF7BBBEF77FCEC791FB0F" pageId="8" pageNumber="9" type="ecology and distribution">
<paragraph id="A4B06BF378B98FB2B906A30BEC76D84A" pageId="8" pageNumber="9">Ecology and distribution.</paragraph>
<paragraph id="1E681D350804547F386879A1B39358FA" pageId="8" pageNumber="9">
Growing usually on exposed calcium-poor rock (e.g. basalt, granite, schist) in pinyon-juniper woodlands but also occurs free on soil. So far known only from collections in western North America.
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca occulta" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="occulta">Rhizoplaca occulta</taxonomicName>
<taxonomicName id="2AB07DA79E5FAC62A76F03D6FC3F887B" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca occulta" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="occulta">Rhizoplaca occulta</taxonomicName>
included a total of five individuals from Idaho (3 individuals), Nevada (1), and Utah (1), USA, and included GenBank accessions identified as
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca cerebriformis" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="cerebriformis">Rhizoplaca cerebriformis</taxonomicName>
<taxonomicName id="F2EE319992DD47B54D38277B2C30F8DA" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca cerebriformis" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="cerebriformis">Rhizoplaca cerebriformis</taxonomicName>
ined. (AF159942) and
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca subidahoensis" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="subidahoensis">Rhizoplaca subidahoensis</taxonomicName>
<taxonomicName id="65F22547EAF7EB9519E248710D4E0A0F" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca subidahoensis" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="subidahoensis">Rhizoplaca subidahoensis</taxonomicName>
ined. (AF159944).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="etymology">
<paragraph pageId="8" pageNumber="9">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="9">
<subSubSection id="7CBDA08C60957CE21D2E52C8093FA806" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8440EAA6485208942C95C36F979DDC76" pageId="8" pageNumber="9">Etymology.</paragraph>
<paragraph id="910EAB11CCF1C65E7DD77C75F3EBD19A" pageId="8" pageNumber="9">
The name is derived from the Latin
<normalizedToken originalValue="“occultus,”">&quot;occultus,&quot;</normalizedToken>
<normalizedToken id="43CB797D80B9781D49BB45CB7D37E151" originalValue="“occultus,”">&quot;occultus,&quot;</normalizedToken>
meaning hidden, and refers to the fact that this species was hidden within the phenotypically circumscribed taxon
<taxonomicName class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca melanophthalma" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="melanophthalma">Rhizoplaca melanophthalma</taxonomicName>
<taxonomicName id="81FD7472350B52AC3AF671E3703922AF" class="Lecanoromycetes" family="Lecanoraceae" genus="Rhizoplaca" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Rhizoplaca melanophthalma" order="Lecanorales" pageId="8" pageNumber="9" phylum="Ascomycota" rank="species" species="melanophthalma">Rhizoplaca melanophthalma</taxonomicName>
sensu lato.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="specimens examined">
<paragraph pageId="8" pageNumber="9">Specimens examined.</paragraph>
<paragraph pageId="8" pageNumber="9">See supplementary file 1.</paragraph>
<subSubSection id="36C5269C27E99CF154AD7C29F97C9750" pageId="8" pageNumber="9" type="specimens examined">
<paragraph id="DF8A14A7269947461855DD1AA297A6A3" pageId="8" pageNumber="9">Specimens examined.</paragraph>
<paragraph id="8A5F1C13AE3274694E4344834B215F57" pageId="8" pageNumber="9">See supplementary file 1.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,93 +1,92 @@
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<paragraph pageId="3" pageNumber="126">Taxon classification Animalia Lepidoptera Zygaenidae</paragraph>
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<paragraph id="5BC0F990E77FD65079FE102B2EDE7303" pageId="3" pageNumber="126">Taxon classification Animalia Lepidoptera Zygaenidae</paragraph>
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<taxonomicName authority="Esper, [1789]" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Zygaena) rhadamanthus" order="Lepidoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="rhadamanthus" subGenus="Zygaena">Zygaena (Zygaena) rhadamanthus (Esper, [1789])</taxonomicName>
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<taxonomicName id="4DBB8DAA628EC5CAD528B54082FD9349" ID-CoL="93SHQ" authority="Esper, [1789]" class="Insecta" family="Zygaenidae" genus="Zygaena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zygaena (Zygaena) rhadamanthus" order="Lepidoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="rhadamanthus" subGenus="Zygaena">Zygaena (Zygaena) rhadamanthus (Esper, [1789])</taxonomicName>
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<subSubSection pageId="3" pageNumber="126" type="reference_group">
<paragraph pageId="3" pageNumber="126">
<taxonomicName class="Insecta" family="Sphingidae" genus="Sphinx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphinx rhadamanthus" order="Lepidoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="rhadamanthus">Sphinx rhadamanthus</taxonomicName>
<subSubSection id="07AFC465B32FAE3D6195F46337E7E5A8" pageId="3" pageNumber="126" type="reference_group">
<paragraph id="9AF81CED123D85DCC185E96C491EB447" pageId="3" pageNumber="126">
<taxonomicName id="F479BE8B6DF4DB5C53AB1A3369288954" class="Insecta" family="Sphingidae" genus="Sphinx" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphinx rhadamanthus" order="Lepidoptera" pageId="3" pageNumber="126" phylum="Arthropoda" rank="species" species="rhadamanthus">Sphinx rhadamanthus</taxonomicName>
Esper, [1789], Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen 2 (Supplement): pl. 40, figs 1, 2; 1794, ibidem 2: 13. Type-locality. France: Gard (Languedoc),
<normalizedToken originalValue="Nîmes">Nimes</normalizedToken>
<normalizedToken id="C394DF09C218400FA618322D7E9814C6" originalValue="Nîmes">Nimes</normalizedToken>
.
</paragraph>
</subSubSection>
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<paragraph pageId="3" pageNumber="126">Remarks.</paragraph>
<paragraph pageId="3" pageNumber="126">
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<paragraph id="41BBF05B51289AE4F2D8752319A15BEC" pageId="3" pageNumber="126">Remarks.</paragraph>
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In stark contrast to the splitting and naming of geographical populations of other
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species,
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: 98) lumps together a number of subspecific taxa of
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(Esper, [1789]), all but one of which were considered valid by
<bibRefCitation author="Hofmann, A" journalOrPublisher="Harley Books, Colchester" pageId="7" pageNumber="130" title="A systematic Catalogue of the Zygaeninae (Lepidoptera: Zygaenidae)." year="1996">Hofmann and Tremewan (1996</bibRefCitation>
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: 130-131). The latter authors accept that less variation is exhibited in the populations occurring in France (east of the Pyrenees) compared to those in the Iberian Peninsula where they exhibit a mosaic of extreme geographical variation, with phenotypes ranging from
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and melanistic to individuals with the forewing ground colour powdered with white scales (griseoid) and reminiscent of
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. The subspecific nominal taxa relevant to their treatment by
<bibRefCitation author="Leraut, P" journalOrPublisher="NAP Editions, Verrieres-le Buisson" pageId="7" pageNumber="130" title="Volume 3: Moths of Europe, Zygaenids, Pyralids 1 and Brachodids." year="2012">Leraut (2012</bibRefCitation>
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: 98) are discussed in detail below.
</paragraph>
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@ -1,86 +1,86 @@
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Genus
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Simon, 1872.
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Figure 4
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Monro, 1937:304-305, textfig. 22.
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<paragraph pageId="8" pageNumber="9">Type material.</paragraph>
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Central Indian Ocean, South Arabian Sea. Holotype (NHML-1937.9.2.455), John Murray Expedition, H.E.M.S. Mabahiss, Stat. 133(
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).
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Figure 4.
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<taxonomicName id="FD897521D3D33C25E8B627197BB0FDE1" class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus hirsutus" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="hirsutus">Ilyphagus hirsutus</taxonomicName>
Monro, 1937. Holotype (NHML-1937.9.2.455). A Dorsal view B Ventral view C Anterior end, dorsal view D Chaetiger, basal, medial and distal notochaetal regions E Chaetiger, neurochaetae F Same, basal region and two hirsute tips. Bars.- A, B 5 mm C 0.2 mm D 45
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="394444C9FE14F53407D510BBF6256932" originalValue="µm">µm</normalizedToken>
E 0.4 mm F 65
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="79CC1B85F68F8771B78CEAD895B06CF5" originalValue="µm">µm</normalizedToken>
.
</paragraph>
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</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="description">
<paragraph pageId="8" pageNumber="9">Description.</paragraph>
<paragraph pageId="8" pageNumber="9">Holotype pale, globose, widened in the posterior half (Fig. 4A, B); 37 mm long, 10 mm wide, cephalic cage chaetae broken, 19 chaetigers. Body surface densely papillated, with abundant fine sediment particles; papillae long, cylindrical, each covered by a thin layer of fine sediment particles.</paragraph>
<paragraph pageId="8" pageNumber="9">Cephalic hood not exposed; specimen not dissected to avoid further damage. Cephalic cage chaetae length unknown. Chaetiger 1 involved in the cephalic cage; chaetae arranged in short rows, notochaetae dorsal, 5-7 per bundle (bases damaged, difficult to count); neurochaetae ventrolateral, 3-4 per bundle.</paragraph>
<paragraph pageId="8" pageNumber="9">Anterior dorsal margin of first chaetiger projected anteriorly (Fig. 4C), large rounded lobe, bent ventrally. Anterior chaetigers without especially long papillae. Chaetiger 1 largest, chaetigers 2-3 of about the same size. Chaetal transition from cephalic cage to body chaetae abrupt; chaetiger 2 with shorter neurochaetae, directed ventrolaterally. Gonopodial lobes present in chaetiger 5, low rounded dark, displaced ventrally, and positioned towards posterior segmental margin.</paragraph>
<paragraph pageId="8" pageNumber="9">Parapodia lateral; median neuropodia ventrolateral. Notopodia scarcely noticed; neuropodia thick low muscular lobes, without especially longer papillae. Noto- and neuropodia close to each other.</paragraph>
<paragraph pageId="8" pageNumber="9">Median notochaetae mostly broken; one anterior notopodia with 3 notochaetae, arranged in a tuft, median notochaetae as long as 1/5 body width, about 3 per fascicle; all notochaetae multiarticulated capillaries, articles short basally, medium-sized medially, long distally (Fig. 4D). Neurochaetae probably multiarticulated capillaries in chaetiger 1; from chaetiger 2, anchylosed aristate spines (Fig. 4E), arranged in transverse rows, in two series, with about 8 chaetae per fascicle. Neurospines basally cylindrical with very short articles, medially flat, distally tapering with slightly longer articles; neurospines basally smooth, subdistally and distally with fibers separated from the main stem, giving hirsute appearance to chaetal surface (Fig. 4F).</paragraph>
<paragraph pageId="8" pageNumber="9">Posterior end globose, damaged; pygidium with anus terminal, without anal cirri.</paragraph>
<subSubSection id="B2811D62E2FDD1F52E5AA545BC3F3F91" pageId="8" pageNumber="9" type="description">
<paragraph id="2E153917B76D11FDD1BFBB425FC37AC2" pageId="8" pageNumber="9">Description.</paragraph>
<paragraph id="9A3B24E494A561050710FF904299F335" pageId="8" pageNumber="9">Holotype pale, globose, widened in the posterior half (Fig. 4A, B); 37 mm long, 10 mm wide, cephalic cage chaetae broken, 19 chaetigers. Body surface densely papillated, with abundant fine sediment particles; papillae long, cylindrical, each covered by a thin layer of fine sediment particles.</paragraph>
<paragraph id="6AFFB6DAD8A25959AE67EBB1758817A5" pageId="8" pageNumber="9">Cephalic hood not exposed; specimen not dissected to avoid further damage. Cephalic cage chaetae length unknown. Chaetiger 1 involved in the cephalic cage; chaetae arranged in short rows, notochaetae dorsal, 5-7 per bundle (bases damaged, difficult to count); neurochaetae ventrolateral, 3-4 per bundle.</paragraph>
<paragraph id="E98978C5EFDD4A261C25C462AABC63AA" pageId="8" pageNumber="9">Anterior dorsal margin of first chaetiger projected anteriorly (Fig. 4C), large rounded lobe, bent ventrally. Anterior chaetigers without especially long papillae. Chaetiger 1 largest, chaetigers 2-3 of about the same size. Chaetal transition from cephalic cage to body chaetae abrupt; chaetiger 2 with shorter neurochaetae, directed ventrolaterally. Gonopodial lobes present in chaetiger 5, low rounded dark, displaced ventrally, and positioned towards posterior segmental margin.</paragraph>
<paragraph id="CB135192A094832CF7C73F672C87C475" pageId="8" pageNumber="9">Parapodia lateral; median neuropodia ventrolateral. Notopodia scarcely noticed; neuropodia thick low muscular lobes, without especially longer papillae. Noto- and neuropodia close to each other.</paragraph>
<paragraph id="B8DF8AC0E23129BAB63273EB32C7DD59" pageId="8" pageNumber="9">Median notochaetae mostly broken; one anterior notopodia with 3 notochaetae, arranged in a tuft, median notochaetae as long as 1/5 body width, about 3 per fascicle; all notochaetae multiarticulated capillaries, articles short basally, medium-sized medially, long distally (Fig. 4D). Neurochaetae probably multiarticulated capillaries in chaetiger 1; from chaetiger 2, anchylosed aristate spines (Fig. 4E), arranged in transverse rows, in two series, with about 8 chaetae per fascicle. Neurospines basally cylindrical with very short articles, medially flat, distally tapering with slightly longer articles; neurospines basally smooth, subdistally and distally with fibers separated from the main stem, giving hirsute appearance to chaetal surface (Fig. 4F).</paragraph>
<paragraph id="EE0C5EEFB5F9940C7468E272AC09C92E" pageId="8" pageNumber="9">Posterior end globose, damaged; pygidium with anus terminal, without anal cirri.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="remarks">
<paragraph pageId="8" pageNumber="9">Remarks.</paragraph>
<paragraph pageId="8" pageNumber="9">
<taxonomicName class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus hirsutus" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="hirsutus">Ilyphagus hirsutus</taxonomicName>
<subSubSection id="50767E77B5E467EF37B9A164CF5C3585" pageId="8" pageNumber="9" type="remarks">
<paragraph id="A97A37204EA9F9C6D86AFB270BDDF4B3" pageId="8" pageNumber="9">Remarks.</paragraph>
<paragraph id="E3B32BE7DA01A8DA1BC80ACE2AEE63A8" pageId="8" pageNumber="9">
<taxonomicName id="72B0C9D30E2135FFC2B5245EBA2940A9" class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus hirsutus" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="hirsutus">Ilyphagus hirsutus</taxonomicName>
Monro, 1937 resembles
<taxonomicName class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus bythincola" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="bythincola">Ilyphagus bythincola</taxonomicName>
<taxonomicName id="4994E6E7775F519291A715A61205CFCB" class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus bythincola" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="bythincola">Ilyphagus bythincola</taxonomicName>
because of their dense coverage with fine, long papillae. They differ because
<taxonomicName class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus hirsutus" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="hirsutus">Ilyphagus hirsutus</taxonomicName>
<taxonomicName id="05AF3825F113DDC6971D8CA88E2E6A53" class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus hirsutus" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="hirsutus">Ilyphagus hirsutus</taxonomicName>
has a projected lobe in the first neuropodia, which is not present in
<taxonomicName class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus bythincola" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="bythincola">Ilyphagus bythincola</taxonomicName>
<taxonomicName id="32138B9F52ABA248358230A0CC90D5E0" class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus bythincola" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="bythincola">Ilyphagus bythincola</taxonomicName>
, and because the neurochaetae of the former are markedly hirsute distally, whereas those in
<taxonomicName class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus bythincola" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="bythincola">Ilyphagus bythincola</taxonomicName>
<taxonomicName id="C90BF3A6C5EE6E35B58E3A8E0817006B" class="Polychaeta" family="Flabelligeridae" genus="Ilyphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ilyphagus bythincola" order="Terebellida" pageId="8" pageNumber="9" phylum="Annelida" rank="species" species="bythincola">Ilyphagus bythincola</taxonomicName>
are either distally hyaline or slightly hirsute.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="distribution">
<paragraph pageId="8" pageNumber="9">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="9">Originally described from the Central Indian Ocean, in deep water (3385 m); it has not been recorded since.</paragraph>
<subSubSection id="589950047B78E56466119A65E426FE1E" pageId="8" pageNumber="9" type="distribution">
<paragraph id="6ED8E24EB26776078E39F5C63DBD65FA" pageId="8" pageNumber="9">Distribution.</paragraph>
<paragraph id="3361DA4E4841339CB7450D4AD7576A57" pageId="8" pageNumber="9">Originally described from the Central Indian Ocean, in deep water (3385 m); it has not been recorded since.</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName LSID="urn:lsid:zoobank.org:act:B802BBE3-E4BF-4971-9569-96266500DBE4" authority="Pujade-Villar &amp; Melika" class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella grahami" order="Hymenoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="grahami">Zapatella grahami Pujade-Villar &amp; Melika</taxonomicName>
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<paragraph pageId="3" pageNumber="78">Type material.</paragraph>
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<paragraph id="6526C7BDDD62D8D867327302DCD69083" pageId="3" pageNumber="78">Type material.</paragraph>
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HOLOTYPE female (deposited in UB): &quot;COSTA RICA, Cartago-Jose, Cerro de la Muerte, 3000 m, 2.X.1988. Col. Hanson&quot; (white label), &quot;
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus costaricensis" order="Fagales" pageId="3" pageNumber="78" phylum="Tracheophyta" rank="species" species="costaricensis">Quercus costaricensis</taxonomicName>
<taxonomicName id="203A5AF29A8D039B93CF9519F838FDD5" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus costaricensis" order="Fagales" pageId="3" pageNumber="78" phylum="Tracheophyta" rank="species" species="costaricensis">Quercus costaricensis</taxonomicName>
, fruit (acorn) galls&quot; (white label), Holotype of
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella grahami" order="Hymenoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="grahami">Zapatella grahami</taxonomicName>
<taxonomicName id="B2E8804A57742751F86797DB93296907" genus="Zapatella" lsidName="Zapatella grahami" pageId="3" pageNumber="78" rank="species" species="grahami">Zapatella grahami</taxonomicName>
♀ Pujade
<pageBreakToken pageId="4" pageNumber="79" start="start">-</pageBreakToken>
<pageBreakToken id="78C9D7FA127D94865AE6A39AE2029DE4" pageId="4" pageNumber="79" start="start">-</pageBreakToken>
Villar &amp; Melika n. gen &amp; n. sp. design. JP-V 2012&quot; (red label). PARATYPES (5 males and 20 females): 3 males and 14 females with the same data as the holotype and 1 male and 1 female with the similar data, only the collecting date is II.1988. 2 males and 10 females are deposited in UB, 1 male and 5 females in PDL, 1 male and 3 females in MZUCR, 1 male and 2 females USNM.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79" type="diagnosis">
<paragraph pageId="4" pageNumber="79">Diagnosis.</paragraph>
<paragraph lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79">
<subSubSection id="642F697F0C338DDABFC156CE2539A824" lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79" type="diagnosis">
<paragraph id="EB16FAF2B556EB7E9E23BDBD8C66A1DF" pageId="4" pageNumber="79">Diagnosis.</paragraph>
<paragraph id="981D6E0D105F30C47547FC6647BCBAAA" lastPageId="5" lastPageNumber="80" pageId="4" pageNumber="79">
In
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella" order="Hymenoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="genus">Zapatella</taxonomicName>
<taxonomicName id="1579AA43B8346C0BA6429409313B7F86" genus="Zapatella" lsidName="Zapatella" pageId="4" pageNumber="79" rank="genus">Zapatella</taxonomicName>
three species,
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella oblata" order="Hymenoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="oblata">Zapatella oblata</taxonomicName>
<taxonomicName id="16DA80EF41151DF59742513A629440A5" genus="Zapatella" lsidName="Zapatella oblata" pageId="4" pageNumber="79" rank="species" species="oblata">Zapatella oblata</taxonomicName>
,
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella grahami" order="Hymenoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="grahami">Zapatella grahami</taxonomicName>
<taxonomicName id="C7443898E8B7A4F665402F914CC3B205" genus="Zapatella" lsidName="Zapatella grahami" pageId="4" pageNumber="79" rank="species" species="grahami">Zapatella grahami</taxonomicName>
sp. n. and
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella nievesaldreyi" order="Hymenoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="nievesaldreyi">Zapatella nievesaldreyi</taxonomicName>
<taxonomicName id="9FDEAC43CA9AEA3CA92BBFE7265823DB" genus="Zapatella" lsidName="Zapatella nievesaldreyi" pageId="4" pageNumber="79" rank="species" species="nievesaldreyi">Zapatella nievesaldreyi</taxonomicName>
sp. n., have the head and mesosoma partially dark brown to black.
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella oblata" order="Hymenoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="oblata">Zapatella oblata</taxonomicName>
<taxonomicName id="CCB12BDE0D8B4CDF1581DE2DC57BEFD2" genus="Zapatella" lsidName="Zapatella oblata" pageId="4" pageNumber="79" rank="species" species="oblata">Zapatella oblata</taxonomicName>
differs from the two other mentioned species by a very long median mesoscutal line which extending to 2/3 of the mesoscutum length, while in
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella grahami" order="Hymenoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="grahami">Zapatella grahami</taxonomicName>
<taxonomicName id="845D8AC0BF64086FF15DC3CF3A9BF824" genus="Zapatella" lsidName="Zapatella grahami" pageId="4" pageNumber="79" rank="species" species="grahami">Zapatella grahami</taxonomicName>
and
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella nievesaldreyi" order="Hymenoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="nievesaldreyi">Zapatella nievesaldreyi</taxonomicName>
<taxonomicName id="F7C6093853F393D6B5FCC3B4C0DE5580" genus="Zapatella" lsidName="Zapatella nievesaldreyi" pageId="4" pageNumber="79" rank="species" species="nievesaldreyi">Zapatella nievesaldreyi</taxonomicName>
the median mesoscutal line is absent or present in a form of a very short triangle. In
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella grahami" order="Hymenoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="grahami">Zapatella grahami</taxonomicName>
<taxonomicName id="BEAE0635FC25B58353B53CCED93690C7" genus="Zapatella" lsidName="Zapatella grahami" pageId="4" pageNumber="79" rank="species" species="grahami">Zapatella grahami</taxonomicName>
the females are much darker, POL 1.4 times as broad as OOL (Fig. 2), bottom of scutellar foveae with rugae (Fig. 11), and the prominent part of the ventral spine of the hypopygium 7.5-8.5 times as long as broad (Figs 14, 16). In
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella nievesaldreyi" order="Hymenoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="nievesaldreyi">
<pageBreakToken pageId="5" pageNumber="80" start="start">Zapatella</pageBreakToken>
<taxonomicName id="E646A5FD4A5449FAE3639038A535059D" genus="Zapatella" lsidName="Zapatella nievesaldreyi" pageId="5" pageNumber="80" rank="species" species="nievesaldreyi">
<pageBreakToken id="06B7DB00D8D7F9A6C94527F7D6924AD1" pageId="5" pageNumber="80" start="start">Zapatella</pageBreakToken>
nievesaldreyi
</taxonomicName>
the females are lighter, POL equal OOL (Fig. 20), bottom of scutellar foveae smooth and without rugae (Fig. 22), and the prominent part of the ventral spine of the hypopygium 6.0-7.0 times as long as broad (Figs 28-29).
</paragraph>
<caption pageId="5" pageNumber="80">
<paragraph pageId="5" pageNumber="80">
<caption id="04977DB2E7BC5B0E8E50DB3DC85EBD8E" pageId="5" pageNumber="80">
<paragraph id="6358608926C13E56F4F93D4E96F38C71" pageId="5" pageNumber="80">
Figures 9-13.
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella grahami" order="Hymenoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="grahami">Zapatella grahami</taxonomicName>
<taxonomicName id="4DAB8D0220701F170E6CA3F34B94DF64" genus="Zapatella" lsidName="Zapatella grahami" pageId="5" pageNumber="80" rank="species" species="grahami">Zapatella grahami</taxonomicName>
,female 9 mesosoma (lateral view) 10 mesoscutum (dorsal view) 11 mesoscutellum (dorsal view) 12 metascutellum and propodeum (posterodorsal view) 13 forewing.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="82" pageId="5" pageNumber="80" type="description">
<paragraph pageId="5" pageNumber="80">Description.</paragraph>
<paragraph pageId="5" pageNumber="80">Female (Figs 1-3, 6, 8, 9-16).</paragraph>
<paragraph pageId="5" pageNumber="80">Length. 2.6-3.2 mm (n=15).</paragraph>
<paragraph pageId="5" pageNumber="80">Coloration. Body, including antennae and legs predominantly dark to chestnut brown. Head, brown with more or less extensive black areas on lower face, basal part of genae, central part of frons and vertex; posteriorly head dark brown to black. Antenna uniformly dark brown, F1-F5 lighter; mesosoma laterally black, except brown dorsolateral area of pronotum; propleura black; mesoscutum brown, with black stripes along anterior parallel and parapsidal lines; mesoscutellum dark brown to black, with slightly lighter scutellar foveae. Propodeum uniformly black; axillula yellowish; legs uniformly brown, with darker hind legs; metasoma brown, anterodorsally darker.</paragraph>
<paragraph pageId="5" pageNumber="80">Head (Figs 1-3). Uniformly and delicately reticulated, with few white setae, 1.8-2.0 times as broad as long from above, 1.3-1.5 times as broad as high in frontal view and slightly broader than mesosoma. Gena broadened behind eye, as broad as transverse diameter of eye; malar space 0.35-0.4 times as long as height of eye, with delicate striae radiating from clypeus and nearly reaching eye margin, malar sulcus absent. POL 1.4 times as long as OOL; OOL 2.5 times as long as length of lateral ocellus and 1.8 times as long as LOL. Transfacial distance nearly 1.2 times as broad as height of eye; diameter of antennal torulus around 3.8 times as great as distance between them, distance between torulus and inner margin of eye equal to or slightly longer than diameter of torulus; inner margins of eyes parallel; lower face delicately coriaceous, with dense white setae, the median elevated area smooth. Clypeus small, squared, smooth, impressed in basal part, ventrally straight; anterior tentorial pits, epistomal sulcus and clypeo-pleurostomal line indistinct. Frons, vertex, interocellar area and occiput delicately reticulate. Postocciput alutaceous and shiny, smooth and impressed around occipital foramen; posterior tentorial pits large; height of occipital foramen as long as height of postgenal bridge; hypostomal carina emarginate, not going around oral foramen, continuing into gular sulcus. Labial palpus 3-segmented, terminal peg distinct, all three segments densely setose; maxillary palpus 5-segmented, terminal peg distinct, three terminal segments densely setose.</paragraph>
<caption pageId="5" pageNumber="80">
<paragraph pageId="5" pageNumber="80">
<subSubSection id="B8D216BDFB85F9845541F28170C1F954" lastPageId="7" lastPageNumber="82" pageId="5" pageNumber="80" type="description">
<paragraph id="47ED31B68773FFF1129A7675A43F70BA" pageId="5" pageNumber="80">Description.</paragraph>
<paragraph id="D4347885C92BEDA292BD62A3FBD3DBF9" pageId="5" pageNumber="80">Female (Figs 1-3, 6, 8, 9-16).</paragraph>
<paragraph id="BF2A8FB77D11F3F1EE21B5FB51B78E8D" pageId="5" pageNumber="80">Length. 2.6-3.2 mm (n=15).</paragraph>
<paragraph id="73C3934243267EC86BE89AE3CDB94411" pageId="5" pageNumber="80">Coloration. Body, including antennae and legs predominantly dark to chestnut brown. Head, brown with more or less extensive black areas on lower face, basal part of genae, central part of frons and vertex; posteriorly head dark brown to black. Antenna uniformly dark brown, F1-F5 lighter; mesosoma laterally black, except brown dorsolateral area of pronotum; propleura black; mesoscutum brown, with black stripes along anterior parallel and parapsidal lines; mesoscutellum dark brown to black, with slightly lighter scutellar foveae. Propodeum uniformly black; axillula yellowish; legs uniformly brown, with darker hind legs; metasoma brown, anterodorsally darker.</paragraph>
<paragraph id="BB7BFB006A684D75C2D88C1C4016D617" pageId="5" pageNumber="80">Head (Figs 1-3). Uniformly and delicately reticulated, with few white setae, 1.8-2.0 times as broad as long from above, 1.3-1.5 times as broad as high in frontal view and slightly broader than mesosoma. Gena broadened behind eye, as broad as transverse diameter of eye; malar space 0.35-0.4 times as long as height of eye, with delicate striae radiating from clypeus and nearly reaching eye margin, malar sulcus absent. POL 1.4 times as long as OOL; OOL 2.5 times as long as length of lateral ocellus and 1.8 times as long as LOL. Transfacial distance nearly 1.2 times as broad as height of eye; diameter of antennal torulus around 3.8 times as great as distance between them, distance between torulus and inner margin of eye equal to or slightly longer than diameter of torulus; inner margins of eyes parallel; lower face delicately coriaceous, with dense white setae, the median elevated area smooth. Clypeus small, squared, smooth, impressed in basal part, ventrally straight; anterior tentorial pits, epistomal sulcus and clypeo-pleurostomal line indistinct. Frons, vertex, interocellar area and occiput delicately reticulate. Postocciput alutaceous and shiny, smooth and impressed around occipital foramen; posterior tentorial pits large; height of occipital foramen as long as height of postgenal bridge; hypostomal carina emarginate, not going around oral foramen, continuing into gular sulcus. Labial palpus 3-segmented, terminal peg distinct, all three segments densely setose; maxillary palpus 5-segmented, terminal peg distinct, three terminal segments densely setose.</paragraph>
<caption id="CC65E34890639CC6BDBECC4B230559D5" pageId="5" pageNumber="80">
<paragraph id="352CE69989F55F85084CFD0357F942A7" pageId="5" pageNumber="80">
Figures 14-18.
<taxonomicName class="Insecta" family="Cynipidae" genus="Zapatella" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Zapatella grahami" order="Hymenoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="grahami">Zapatella grahami</taxonomicName>
<taxonomicName id="0C0AF5564A4FC2422968E275F5946663" genus="Zapatella" lsidName="Zapatella grahami" pageId="5" pageNumber="80" rank="species" species="grahami">Zapatella grahami</taxonomicName>
14 metasoma, female (lateral view) 15 metasoma, male (lateral view) 16 ventral spine of hypopygium (ventral view) 17-18 gall.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="80">
<paragraph id="05ADD897BD1B776DEFEEA0A9BB500080" pageId="5" pageNumber="80">
Antenna (Fig. 6). With 11 flagellomeres (14: 9
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="AE7D9542C753A650D9AB2F7786A08327" originalValue="×">x</normalizedToken>
8: 15
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="6E0F58C0EB65D8C781D5F810F7CD93A0" originalValue="×">x</normalizedToken>
8: 19: 15: 14: 13: 11: 10: 10: 9: 8: 16); longer than head+mesosoma (48:34); pedicel globose, as long as broad; F1 as long as scapus; F2 1.2-1.3 times as long as F1; F1=F3; F4-F5 subequal and shorter, F6-F10 shorter and progressively shortening in length; F11 twice as long as F10; placodeal sensilla distinct on F6-F11, indistinct but present on F4-F5, absent on F1-F3.
</paragraph>
<paragraph lastPageId="7" lastPageNumber="82" pageId="5" pageNumber="80">
<paragraph id="59184D11C8E588D0D6E8FF4AFFDB1115" lastPageId="7" lastPageNumber="82" pageId="5" pageNumber="80">
Mesosoma (Figs 9-12). 1.2-1.3 times as long as high in lateral view, with few white setae. Mesoscutum as long as broad, or only slightly longer than broad in dorsal view; with sparse scattered setae and transverse, delicate, interrupted striae which connect with longitudinally orientated weak striae, forming an irregular network of striae, and an irregularly reticulate surface sculpture. Notauli incomplete, extending at most to half length of mesoscutum; converging, deep and broad posteriorly. Anterior parallel lines extending to
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="6C98CA0BB0C641876A9C4705FAA09ABB" originalValue="½">1/2</normalizedToken>
length of mesoscutum; parapsidal
<pageBreakToken pageId="6" pageNumber="81" start="start">lines</pageBreakToken>
<pageBreakToken id="9574E30FEEADAB52263A81112F27B4AB" pageId="6" pageNumber="81" start="start">lines</pageBreakToken>
distinct and broad, starting from posterior margin and extending to
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="A605334A608A3B988CAA3D322AD8F846" originalValue="½">1/2</normalizedToken>
length of mesoscutum; median mesoscutal line very short or absent. Mesoscutellum 0.5 times as long as mesoscutum, as long as broad, not overhanging metanotum, center of disk reticulate, sides and posterior 1/3 dull rugose; scutellar foveae present, ovate, not delimited posteriorly, bottom shiny, with some rugae; median carina broad. Mesopleuron uniformly delicately reticulate, smooth and shiny basally; mesopleural triangle conspicuously setose; dorsal axillar area coriaceous with numerous setae, lateral axillar area reticulo-carinate, without setae; axillula smooth, with white setae; subaxillular bar smooth, shiny, narrower than height of metanotal trough; postalar process long, strong, reticulate; metapleural sulcus reaching mesopleuron in upper 2/3 of its height. Metascutellum strongly reticulate, rectangular. Metanotal trough with short white setae; ventral impressed area at least twice as narrow as height of metascutellum, delicately reticulate. Propodeum setose lateraly, glabrous centrally;
<pageBreakToken pageId="7" pageNumber="82" start="start">central</pageBreakToken>
<pageBreakToken id="CED3107E3BB45210D4C05BB72D2F51DF" pageId="7" pageNumber="82" start="start">central</pageBreakToken>
propodeal area smooth, shiny, with many irregular wrinkles and rugae, lateral propodeal carinae weak, diverging anteriorly and converging in posterior 1/3. Nucha with irregular wrinkles and rugae.
</paragraph>
<paragraph pageId="7" pageNumber="82">Legs (Fig. 8). Tarsal claws simple, without basal lobe; hind coxae with dense white setae on the dorsoposterior surface.</paragraph>
<paragraph pageId="7" pageNumber="82">Forewing (Fig. 13). Longer than body, hyaline, without cilia on margin; radial cell 3.3 times as long as broad; 2r distinct; R1 absent or hardly visible, Rs very inconspicuous, nearly straight; areolet absent or very indistinct. Rs+M indistinct, reaching basalis at half of its height.</paragraph>
<paragraph pageId="7" pageNumber="82">Metasoma (Figs 14, 16). Shorter than head+mesosoma, slightly higher than long in lateral view; base of 2nd metasomal tergite with felt-like dense ring of white setae, interrupted dorsally and few setae scattered on lateral surface of tergite. Narrow posterior band on 2nd metasomal tergite and all subsequent tergites with very delicate dense micropunctures. Prominent part of ventral spine of hypopygium needle-like, tapering to apex, 7.5-8.5 times as long as broad, with two parallel rows of short white scattered setae which do not extend beyond the apex of spine.</paragraph>
<paragraph pageId="7" pageNumber="82">
<paragraph id="C6EC2213827AFDD170E29D84326FB9DA" pageId="7" pageNumber="82">Legs (Fig. 8). Tarsal claws simple, without basal lobe; hind coxae with dense white setae on the dorsoposterior surface.</paragraph>
<paragraph id="F6C8F87B0FEE85C3949D0D6BF71098A4" pageId="7" pageNumber="82">Forewing (Fig. 13). Longer than body, hyaline, without cilia on margin; radial cell 3.3 times as long as broad; 2r distinct; R1 absent or hardly visible, Rs very inconspicuous, nearly straight; areolet absent or very indistinct. Rs+M indistinct, reaching basalis at half of its height.</paragraph>
<paragraph id="9E5FBEAC51D65696A9FA92C441C9D23B" pageId="7" pageNumber="82">Metasoma (Figs 14, 16). Shorter than head+mesosoma, slightly higher than long in lateral view; base of 2nd metasomal tergite with felt-like dense ring of white setae, interrupted dorsally and few setae scattered on lateral surface of tergite. Narrow posterior band on 2nd metasomal tergite and all subsequent tergites with very delicate dense micropunctures. Prominent part of ventral spine of hypopygium needle-like, tapering to apex, 7.5-8.5 times as long as broad, with two parallel rows of short white scattered setae which do not extend beyond the apex of spine.</paragraph>
<paragraph id="3F35063C236BE4E70502318E2F72A993" pageId="7" pageNumber="82">
Male (Figs 4-5, 7, 15). Length 2.3-2.5 mm (n=4). Similar to female, except in the following characters: predominantly black with few brown areas; head 2.0 times as broad as long from above, 1.2 times as broad as high and broader than mesosoma in frontal view; malar space 0.3 times as long as height of eye; POL 2.0 times as broad as OOL; OOL 2.0 times as long as length of lateral ocellus and 1.3 times as long as LOL. Antennae with 13 flagellomeres (6: 4
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="A785A1FB322BD8D9E9261909674992B7" originalValue="×">x</normalizedToken>
4: 11
<normalizedToken originalValue="×">x</normalizedToken>
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3.5: 10: 9: 9: 8: 8: 7: 7: 6: 6: 6: 6: 7); longer than body (101:93); pedicel as long as broad; F1 slightly longer than F2, distinctly curved, dorsally flattened and excavate; subsequent flagellomeres progressively shorter in length; F13 longer than F12; placodeal sensilla on all flagellomeres.
</paragraph>
<paragraph pageId="7" pageNumber="82">Gall (Figs 17-18). Acorn galls. Individual chambers located in the acorn cup, often between the cup and the seed. Usually there is one gall per acorn, but sometime two or three.</paragraph>
<paragraph pageId="7" pageNumber="82">
<paragraph id="26DDF6D6B523CFF9981FE84DC84ECE56" pageId="7" pageNumber="82">Gall (Figs 17-18). Acorn galls. Individual chambers located in the acorn cup, often between the cup and the seed. Usually there is one gall per acorn, but sometime two or three.</paragraph>
<paragraph id="EE787DC044B971D0C7F1FC7CB807AE83" pageId="7" pageNumber="82">
Biology. Only the sexual generation is known and it induces galls on
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus costaricensis" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="costaricensis">Quercus costaricensis</taxonomicName>
<taxonomicName id="1427AA751A5ED99323A1828D3D5DA8E3" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus costaricensis" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="costaricensis">Quercus costaricensis</taxonomicName>
. Galls were collected in February and later in October in forests located above 3000 m altitude, adults emerged immediately after the galls were collected, in February and October. This very unusual emergence of adults in two periods may be due to the sporadic nature of the collecting and to the peculiar phenology of
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus costaricensis" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="costaricensis">Quercus costaricensis</taxonomicName>
<taxonomicName id="1372A55080266D4606952E82A68BFD48" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus costaricensis" order="Fagales" pageId="7" pageNumber="82" phylum="Tracheophyta" rank="species" species="costaricensis">Quercus costaricensis</taxonomicName>
. In the area where the galls were collected,
<bibRefCitation author="Camacho, M" journalOrPublisher="Revista de Biologia Tropical" pageId="21" pageNumber="96" pagination="533 - 542" title="Patrones fenologicos de doce especies arboreas del bosque montano de la Cordillera de Talamanca, Costa Rica." volume="46" year="1998">Camacho and Orozco (1998)</bibRefCitation>
<bibRefCitation id="A19DED52A8A5AFDE68B0BA1BEF5F97A9" author="Camacho, M" journalOrPublisher="Revista de Biologia Tropical" pageId="21" pageNumber="96" pagination="533 - 542" title="Patrones fenologicos de doce especies arboreas del bosque montano de la Cordillera de Talamanca, Costa Rica." volume="46" year="1998">Camacho and Orozco (1998)</bibRefCitation>
observed the flowering and fruiting phenology for a four year period (July 1986 to July 1990. The female flowers were present for ten months of the year, starting in the rainy season, with a flowering peak in the dry season. Male flowers were present for seven months, with a flowering peak from October to January, the period from the end of the rainy season and continuing to the beginning of the dry season. During the four years of observation there was only one fruiting period, which was synchronous, very productive, and extended for eight months (August 1988 to March 1989); this is one year after the initial production of female flowers and six months after the end of male flower production.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="83" type="distribution">
<paragraph pageId="8" pageNumber="83">
<pageBreakToken pageId="8" pageNumber="83" start="start">Distribution</pageBreakToken>
<subSubSection id="F2F8C9F5B9C3E89B61ED6D92560A1310" pageId="8" pageNumber="83" type="distribution">
<paragraph id="2E211E1770C3A8C9268497F4FB989156" pageId="8" pageNumber="83">
<pageBreakToken id="723808F614786CC580730403E52E44BD" pageId="8" pageNumber="83" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="8" pageNumber="83">Currently known only from Costa Rica (Cerro de la Muerte).</paragraph>
<paragraph id="15070245AABAD5590B86B6B6F2EBC856" pageId="8" pageNumber="83">Currently known only from Costa Rica (Cerro de la Muerte).</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="83" type="etymology">
<paragraph pageId="8" pageNumber="83">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="83">In recognition of the continuing contribution of our friend, prof. Graham N. Stone (Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, Scotland) to research on oak gallwasps.</paragraph>
<subSubSection id="FE245E4EA6053679B8CA8446A188A43F" pageId="8" pageNumber="83" type="etymology">
<paragraph id="43FC6730804F783D5E7D48D6ABA79AB3" pageId="8" pageNumber="83">Etymology.</paragraph>
<paragraph id="6FDE7828F81487465967D1A5DFD99AED" pageId="8" pageNumber="83">In recognition of the continuing contribution of our friend, prof. Graham N. Stone (Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, Scotland) to research on oak gallwasps.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,368 +1,368 @@
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<treatment ID-GBIF-Taxon="159932957" LSID="urn:lsid:plazi:treatment:C9C05D9D73FC536987E8091315C6F4CB" httpUri="http://treatment.plazi.org/id/C9C05D9D73FC536987E8091315C6F4CB" lastPageNumber="107" pageId="0" pageNumber="107">
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<taxonomicName LSID="C9C05D9D-73FC-5369-87E8-091315C6F4CB" authority="Becker, 1912" authorityName="Becker" authorityYear="1912" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">Enderleiniella Becker, 1912</taxonomicName>
<treatment id="C9C05D9D73FC536987E8091315C6F4CB" ID-GBIF-Taxon="159932957" LSID="urn:lsid:plazi:treatment:C9C05D9D73FC536987E8091315C6F4CB" httpUri="http://treatment.plazi.org/id/C9C05D9D73FC536987E8091315C6F4CB" lastPageNumber="107" pageId="0" pageNumber="107" scope_family="Chloropidae" scope_order="Diptera">
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<paragraph id="6CFC8D60D05CF798C86CFF59B125B51E" pageId="0" pageNumber="107">
<taxonomicName id="245D8756620AE61B14D1B802A59392D2" ID-CoL="4BL7" LSID="C9C05D9D-73FC-5369-87E8-091315C6F4CB" authority="Becker, 1912" authorityName="Becker" authorityYear="1912" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">Enderleiniella Becker, 1912</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="reference_group">
<paragraph pageId="0" pageNumber="107">
<taxonomicName class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">Enderleiniella</taxonomicName>
<subSubSection id="C43472F7225E52685BBE4DBB76A3A08A" pageId="0" pageNumber="107" type="reference_group">
<paragraph id="FCB44D25C7A8B8E92849AB64F65AA08F" pageId="0" pageNumber="107">
<taxonomicName id="876A3F75491DE4960BED497145FEC585" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">Enderleiniella</taxonomicName>
Becker, 1912: 192. Type species:
<taxonomicName class="Insecta" family="Chloropidae" genus="Tricimba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tricimba longiventris" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="longiventris">Tricimba longiventris</taxonomicName>
<taxonomicName id="F4EFF8909F13D165AACC3E915D4FCA8B" class="Insecta" family="Chloropidae" genus="Tricimba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tricimba longiventris" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="longiventris">Tricimba longiventris</taxonomicName>
Enderlein (original description).
</paragraph>
<paragraph pageId="0" pageNumber="107">
<taxonomicName class="Insecta" family="Chloropidae" genus="Anoscinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoscinella" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">Anoscinella</taxonomicName>
<paragraph id="7CB5D9FFA1513DE08B56B1EBD617D730" pageId="0" pageNumber="107">
<taxonomicName id="E9887C8C068D2DEBE79A4CA618D8F83E" class="Insecta" family="Chloropidae" genus="Anoscinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoscinella" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">Anoscinella</taxonomicName>
Becker, 1916: 448. Type species:
<taxonomicName class="Insecta" family="Chloropidae" genus="Anoscinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoscinella tripunctata" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="tripunctata">Anoscinella tripunctata</taxonomicName>
<taxonomicName id="7ECC4C0DE62F32580E9694644A9A9123" class="Insecta" family="Chloropidae" genus="Anoscinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anoscinella tripunctata" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="tripunctata">Anoscinella tripunctata</taxonomicName>
Becker (monotypy);
<bibRefCitation author="Duda, O" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="107" refId="B4" refString="Duda, O, 1930. Die neotropischen Chloropiden (Dipt.). Folia Zoologica et Hydrobiologica 2: 46-128, v-xix." title="Die neotropischen Chloropiden (Dipt.). Folia Zoologica et Hydrobiologica 2: 46 - 128, v-xix." year="1930">Duda 1930</bibRefCitation>
<bibRefCitation id="1602EDE5F673E2BA2B982F4CBD879B8B" author="Duda, O" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="107" refId="B4" refString="Duda, O, 1930. Die neotropischen Chloropiden (Dipt.). Folia Zoologica et Hydrobiologica 2: 46-128, v-xix." title="Die neotropischen Chloropiden (Dipt.). Folia Zoologica et Hydrobiologica 2: 46 - 128, v-xix." year="1930">Duda 1930</bibRefCitation>
: 70 (synonymy).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="diagnosis">
<paragraph pageId="0" pageNumber="107">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="107">
<subSubSection id="35BCC16A062141807250DC9D2D1B7ED0" pageId="0" pageNumber="107" type="diagnosis">
<paragraph id="373294B5BF7DA2561DA22C2324290626" pageId="0" pageNumber="107">Diagnosis.</paragraph>
<paragraph id="8286EA6B99B4FA0E193004F9E2647ED2" pageId="0" pageNumber="107">
Small to medium
<taxonomicName lsidName="" pageId="0" pageNumber="107" rank="subfamily" subfamily="Oscinellinae">Oscinellinae</taxonomicName>
<taxonomicName id="DFC2A365F11910C89F10657BF16343A4" lsidName="" pageId="0" pageNumber="107" rank="subfamily" subfamily="Oscinellinae">Oscinellinae</taxonomicName>
with head as wide as or slightly wider than scutum in dorsal view, occiput posteriorly convex in dorsal view, eye usually hairy, gena thin always parallel with ventral portion of eye, scutum elongate, with three parallel lines of shallow incised punctures, wing long and slender, broadest in distal half, with reduced alula and anal angle, legs long and slender, abdomen long and narrow, male genitalia with sternite 6 present, in some species epandrium enlarged and cercus elongate.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="description">
<paragraph pageId="0" pageNumber="107">Description.</paragraph>
<paragraph pageId="0" pageNumber="107">
<subSubSection id="781C2BA2F0384EDD7E0B095D9BF13A53" pageId="0" pageNumber="107" type="description">
<paragraph id="2C4625CE4D32BF67CE6693E9D3D0E4E4" pageId="0" pageNumber="107">Description.</paragraph>
<paragraph id="ED06422EBAD80E0D4F9F78684A4E58F0" pageId="0" pageNumber="107">
(
<figureCitation captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. 1 Enderleiniella longiventris, head lateral view. 2 Enderleiniella longiventris, scutum and scutellum 3 Enderleiniella flavida, scutum and scutellum 4 Enderleiniella maculata, wing. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.36154.figures1-4" httpUri="https://binary.pensoft.net/fig/351394" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures1-4">Figs 1-4</figureCitation>
<figureCitation id="9E6ACF3A4361165A6041BF1D7CDF4819" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. 1 Enderleiniella longiventris, head lateral view. 2 Enderleiniella longiventris, scutum and scutellum 3 Enderleiniella flavida, scutum and scutellum 4 Enderleiniella maculata, wing. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.884.36154.figures1-4" httpUri="https://binary.pensoft.net/fig/351394" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures1-4">Figs 1-4</figureCitation>
):
<taxonomicName family="Chloropidae" lsidName="" pageId="0" pageNumber="107" rank="family">Chloropidae</taxonomicName>
<taxonomicName id="E77CAABA57B05318E8EE20DA603A9E5D" class="Insecta" family="Chloropidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="family">Chloropidae</taxonomicName>
,
<taxonomicName lsidName="" pageId="0" pageNumber="107" rank="subfamily" subfamily="Oscinellinae">Oscinellinae</taxonomicName>
<taxonomicName id="EFBF60DCC62F5467C2CD64F5BDD73556" lsidName="" pageId="0" pageNumber="107" rank="subfamily" subfamily="Oscinellinae">Oscinellinae</taxonomicName>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Head.</emphasis>
<emphasis id="B0978F272BCDED27CCE02DD13D5BAD2C" bold="true" italics="true" pageId="0" pageNumber="107">Head.</emphasis>
Frontal triangle glossy or microtomentose, occiput convex; frons with many interfrontal setulae; cephalic setae short, 6-12 reclinate fronto-orbitals with most dorsal seta proclinate; ocellars and postocellars erect, convergent, vertical setae stronger than other cephalic setae, inner vertical setae as long as outer vertical setae, in line with posterior ocelli; ocelli in most species large; eye large, oval, usually densely haired; postgena clearly visible; gena linear, densely pruinose, very pale except for brown ventral margin; vibrissal angle not projecting, vibrissa present but small; face flat, narrow, pruinose, very pale, facial carina short and small; antenna with scape and pedicel short, first flagellomere large, subquadrate; arista sparsely pubescent, aristal setulae usually longer than width of arista at base; proboscis variable from small to geniculate, but never long, palpus short.
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Scutum.</emphasis>
<emphasis id="CFC9C5743BB25E3975230B873C4E1886" bold="true" italics="true" pageId="0" pageNumber="107">Scutum.</emphasis>
Pronotum elongate and visible in dorsal view, sulcus between postpronotum and scutum deep and well defined; scutum usually glossy, rarely pruinose, with three parallel lines of finely incised punctures (treated as grooves by
<bibRefCitation DOI="https://doi.org/10.1071/IT9930297" author="Ismay, JW" journalOrPublisher="Invertebrate Taxonomy" pageId="0" pageNumber="107" pagination="297 - 499" refId="B9" refString="Ismay, JW, 1993. Revision of Tricimba Lioy and Aprometopis Becker (Diptera: Chloropidae) from Australia and the Papuan region. Invertebrate Taxonomy 7: 297 - 499" title="Revision of Tricimba Lioy and Aprometopis Becker (Diptera: Chloropidae) from Australia and the Papuan region." url="https://doi.org/10.1071/IT9930297" volume="7" year="1993">Ismay (1993)</bibRefCitation>
<bibRefCitation id="10C27F3B15B455D6E8ED03CD3D699B25" DOI="https://doi.org/10.1071/IT9930297" author="Ismay, JW" journalOrPublisher="Invertebrate Taxonomy" pageId="0" pageNumber="107" pagination="297 - 499" refId="B9" refString="Ismay, JW, 1993. Revision of Tricimba Lioy and Aprometopis Becker (Diptera: Chloropidae) from Australia and the Papuan region. Invertebrate Taxonomy 7: 297 - 499" title="Revision of Tricimba Lioy and Aprometopis Becker (Diptera: Chloropidae) from Australia and the Papuan region." url="https://doi.org/10.1071/IT9930297" volume="7" year="1993">Ismay (1993)</bibRefCitation>
), less distinct in some species, with one anterior and either one (
<taxonomicName class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella flavida" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="flavida">
<emphasis italics="true" pageId="0" pageNumber="107">Enderleiniella flavida</emphasis>
<taxonomicName id="B6734B2519A176FED45C6FE8EFFD2FAA" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella flavida" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="17D1F8EE727251F471452B0AC1F2C462" italics="true" pageId="0" pageNumber="107">Enderleiniella flavida</emphasis>
</taxonomicName>
) or two posterior notopleural setae, one longer outer postalar bristle and one weak inner postalar setae, one postsutural dorsocentral bristle, all dark; postpronotal seta and other scutal setae weak; scutal setulae short, weak, evenly arranged in three distinct rows; in certain species scattered setulae present between and outside the three distinct rows; scutellum flattened, trapezoidal or rectangular dorsally, with two small marginal projections, bearing one strong bristle, lateral marginal scutellar setae present; Thoracic pleurites bare.
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Wing.</emphasis>
<emphasis id="675798F53E04D47D614EF8B98F32E31C" bold="true" italics="true" pageId="0" pageNumber="107">Wing.</emphasis>
Hyaline (a darkened spot in
<taxonomicName class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella maculata" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="107">Enderleiniella maculata</emphasis>
<taxonomicName id="BF144FF2815FE31308E595ED72BA97E9" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella maculata" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="6A58CF7C40FF3AD2C82FBE682B019134" italics="true" pageId="0" pageNumber="107">Enderleiniella maculata</emphasis>
</taxonomicName>
), long and slender, broadest in distal half, alula and anal angle reduced, cell c normal, second costal sector longer than third; cell r1 normal and long, R2+3 and R4+5 divergent at base, cell br narrow, crossvein r-m near middle of cell dm, crossvein dm-cu fused with M4 in a right angle.
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Legs.</emphasis>
<emphasis id="0B4DDF3E18B60D73648F4CF191081A49" bold="true" italics="true" pageId="0" pageNumber="107">Legs.</emphasis>
Slender without outstanding setae or spurs; femoral organ present as a row or small patch of three or four sensillae, tibial organ long oval, velvety, with single longitudinal row of setulae along midline.
<emphasis bold="true" italics="true" pageId="0" pageNumber="107">Abdomen.</emphasis>
<emphasis id="25573FE741AB5B01253363E83F3F485E" bold="true" italics="true" pageId="0" pageNumber="107">Abdomen.</emphasis>
Narrow, cylindrical; syntergites 1+2 elongate; tergites broad; sternites small and narrow; spiracles 3-5 in abdominal membrane ventral to lateral margins of tergites;
</paragraph>
<caption doi="10.3897/zookeys.884.36154.figures1-4" httpUri="https://binary.pensoft.net/fig/351394" pageId="0" pageNumber="107" start="Figures 14" startId="F1">
<paragraph pageId="0" pageNumber="107">
<emphasis bold="true" pageId="0" pageNumber="107">Figures 1-4.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="107">1</emphasis>
<taxonomicName class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella longiventris" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="longiventris">
<emphasis italics="true" pageId="0" pageNumber="107">Enderleiniella longiventris</emphasis>
<caption id="98A10D6A91DE987008F0019B4257E673" doi="10.3897/zookeys.884.36154.figures1-4" httpUri="https://binary.pensoft.net/fig/351394" pageId="0" pageNumber="107" start="Figures 14" startId="F1">
<paragraph id="AC32DDA36C1EEF1BF1F2C1FFDC15E0DC" pageId="0" pageNumber="107">
<emphasis id="770826329C61B4E559C89D9DA62EE9AC" bold="true" pageId="0" pageNumber="107">Figures 1-4.</emphasis>
<emphasis id="8A54E7AF87379F1ADB0ABB5606CD95DC" bold="true" pageId="0" pageNumber="107">1</emphasis>
<taxonomicName id="62A03D250F0636DD0652F8008A33F4B8" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella longiventris" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="longiventris">
<emphasis id="FD4F90643B2F3A50A16F15B5EDA323AC" italics="true" pageId="0" pageNumber="107">Enderleiniella longiventris</emphasis>
</taxonomicName>
, head lateral view.
<emphasis bold="true" pageId="0" pageNumber="107">2</emphasis>
<taxonomicName class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella longiventris" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="longiventris">
<emphasis italics="true" pageId="0" pageNumber="107">Enderleiniella longiventris</emphasis>
<emphasis id="693D63498B0260926F731022C7233CBC" bold="true" pageId="0" pageNumber="107">2</emphasis>
<taxonomicName id="05D87A4611DB985E56D7781B6B27FB12" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella longiventris" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="longiventris">
<emphasis id="2EF8733980EBEAE1A33B36ED40CCE528" italics="true" pageId="0" pageNumber="107">Enderleiniella longiventris</emphasis>
</taxonomicName>
, scutum and scutellum
<emphasis bold="true" pageId="0" pageNumber="107">3</emphasis>
<taxonomicName class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella flavida" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="flavida">
<emphasis italics="true" pageId="0" pageNumber="107">Enderleiniella flavida</emphasis>
<emphasis id="CA764D4EAA523DFEDCA34EEC68C5DC2E" bold="true" pageId="0" pageNumber="107">3</emphasis>
<taxonomicName id="BB81F977213F52E8540DBD2FC346A06B" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella flavida" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="9FCDFC7A4FB8A0885BBA31714ACA3468" italics="true" pageId="0" pageNumber="107">Enderleiniella flavida</emphasis>
</taxonomicName>
, scutum and scutellum
<emphasis bold="true" pageId="0" pageNumber="107">4</emphasis>
<taxonomicName class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella maculata" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="107">Enderleiniella maculata</emphasis>
<emphasis id="FF945F4E0ADDFBAE083C120248D04169" bold="true" pageId="0" pageNumber="107">4</emphasis>
<taxonomicName id="EE0E267EF1A133C4C954A928EFF30AD4" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella maculata" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="6BEB4A34E1E4F72BC829E09639CC063A" italics="true" pageId="0" pageNumber="107">Enderleiniella maculata</emphasis>
</taxonomicName>
, wing. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="description">
<paragraph pageId="0" pageNumber="107">Male specific characters.</paragraph>
<paragraph pageId="0" pageNumber="107">Sternite 6 present; dorsal pregenital sclerite symmetrical, left and right spiracles 6 and 7 within sclerite close to lateral margin; epandrium variable from small and simple to enlarged and inflated. Surstylus simple, straight, elongate; hypandrium open posteriorly; pregonite and postgonite simple, round and elongate; cercus variable from simple and elongate to multiple lobed; subepandrial sclerite usually small.</paragraph>
<subSubSection id="2ADAA615C41EBFBA39D82024DEBA19CD" pageId="0" pageNumber="107" type="description">
<paragraph id="09FE7BFBB69C64C5C88D4F684E504BF2" pageId="0" pageNumber="107">Male specific characters.</paragraph>
<paragraph id="18976F04E1F6DD13112484C823F77132" pageId="0" pageNumber="107">Sternite 6 present; dorsal pregenital sclerite symmetrical, left and right spiracles 6 and 7 within sclerite close to lateral margin; epandrium variable from small and simple to enlarged and inflated. Surstylus simple, straight, elongate; hypandrium open posteriorly; pregonite and postgonite simple, round and elongate; cercus variable from simple and elongate to multiple lobed; subepandrial sclerite usually small.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="description">
<paragraph pageId="0" pageNumber="107">Female specific characters.</paragraph>
<paragraph pageId="0" pageNumber="107">
<subSubSection id="45AA84154D700F71CC2CB9FC64D3CFF1" pageId="0" pageNumber="107" type="description">
<paragraph id="D6C2549975118E4EADAA971801A3525A" pageId="0" pageNumber="107">Female specific characters.</paragraph>
<paragraph id="42FAD881A168D2E48CB7D356F8C535CE" pageId="0" pageNumber="107">
Terminalia unmodified, typical of
<taxonomicName lsidName="" pageId="0" pageNumber="107" rank="subfamily" subfamily="Oscinellinae">Oscinellinae</taxonomicName>
<taxonomicName id="949F89FA0405129139C411E5D2C8E517" lsidName="" pageId="0" pageNumber="107" rank="subfamily" subfamily="Oscinellinae">Oscinellinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="107" type="comments">
<paragraph pageId="0" pageNumber="107">Comments.</paragraph>
<paragraph pageId="0" pageNumber="107">
<subSubSection id="6874E3BFBD2DADFDD326B4CE6D922B35" pageId="0" pageNumber="107" type="comments">
<paragraph id="D909A14B94B7CBCBF4E0DA653B7AC201" pageId="0" pageNumber="107">Comments.</paragraph>
<paragraph id="8318CDB601811FBB0ABC18364F03829E" pageId="0" pageNumber="107">
The type species,
<taxonomicName class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella longiventris" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="longiventris">
<emphasis italics="true" pageId="0" pageNumber="107">Enderleiniella longiventris</emphasis>
<taxonomicName id="9AAE79C99BC8EFCA5AFDA89C64453785" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella longiventris" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="longiventris">
<emphasis id="A551FEBB1CD9C52199D59D307463C092" italics="true" pageId="0" pageNumber="107">Enderleiniella longiventris</emphasis>
</taxonomicName>
, is atypical in having a densely pruinose scutum and a long, rectangular scutellum.
</paragraph>
<subSection pageId="0" pageNumber="107" type="key to described species of enderleiniella">
<paragraph pageId="0" pageNumber="107">
<subSection id="BE5B25CD51E549ABDA16F08D7A3344F5" pageId="0" pageNumber="107" type="key to described species of enderleiniella">
<paragraph id="1EA32FFD37BA783AAA2E66B9BE69F6F9" pageId="0" pageNumber="107">
Key to described species of
<taxonomicName class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="107">Enderleiniella</emphasis>
<taxonomicName id="3B5456ACC136E9C3F508E3918416B8E4" class="Insecta" family="Chloropidae" genus="Enderleiniella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Enderleiniella" order="Diptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="8C72C3231FE52D1050E6CC1E0332263A" italics="true" pageId="0" pageNumber="107">Enderleiniella</emphasis>
</taxonomicName>
</paragraph>
<paragraph pageId="0" pageNumber="107">
<table pageId="0" pageNumber="107">
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">1</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<paragraph id="DD84F66424F2BDE56A32C618B0D0EB73" pageId="0" pageNumber="107">
<table id="BA2BD9336FF5A3B02EA0FC8636E1BBBB" pageId="0" pageNumber="107">
<tr id="0E89986AB7AA978850D0DFE03969B281" pageId="0" pageNumber="107">
<td id="C70C80937F15390676293751B6B06FC0" colspan="1" pageId="0" pageNumber="107" rowspan="1">1</td>
<td id="E44DB292EBA09040D7E864CB0A19036C" colspan="1" pageId="0" pageNumber="107" rowspan="1">
Scutellum pale yellow (
<figureCitation captionStart="Figures 1519" captionStartId="F4" captionText="Figures 15 - 19. Enderleiniella flavida. 15 Lateral habitus 16 dorsal habitus 17 male genitalia (lateral) 18 male genitalia (posterior) 19 geographic distribution." figureDoi="10.3897/zookeys.884.36154.figures15-19" httpUri="https://binary.pensoft.net/fig/351397" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures15-19">Fig. 16</figureCitation>
<figureCitation id="058B77BC9E9613270F4C512373829276" captionStart="Figures 1519" captionStartId="F4" captionText="Figures 15 - 19. Enderleiniella flavida. 15 Lateral habitus 16 dorsal habitus 17 male genitalia (lateral) 18 male genitalia (posterior) 19 geographic distribution." figureDoi="10.3897/zookeys.884.36154.figures15-19" httpUri="https://binary.pensoft.net/fig/351397" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures15-19">Fig. 16</figureCitation>
), contrasting strongly with dark scutum; occiput with single strong bristle behind eye; one anterior and one posterior notopleural bristle
</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. flavida" pageId="0" pageNumber="107" rank="species" species="flavida">
<emphasis italics="true" pageId="0" pageNumber="107">E. flavida</emphasis>
<td id="188FD45704D9BE13BB7B3BC1EC46C666" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="AD09BB665915440FA4FE948120D8CF46" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="3B9A942FEC884E8BD3EC25F46BF97728" lsidName="E. flavida" pageId="0" pageNumber="107" rank="species" species="flavida">
<emphasis id="56EC3E1480811F4553092103DD387FDC" italics="true" pageId="0" pageNumber="107">E. flavida</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum brown or grey, not contrasting strongly with scutum; occiput without outstanding bristle behind eye; one anterior and two posterior notopleural setae</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">2</emphasis>
<tr id="BBE67AB2DD8B0819D49A10CB7849A83C" pageId="0" pageNumber="107">
<td id="5C86FE703474468AE281227CBF5A92DE" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="A4B8827D2B5634EB3200B00DB60540DA" colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum brown or grey, not contrasting strongly with scutum; occiput without outstanding bristle behind eye; one anterior and two posterior notopleural setae</td>
<td id="A02F96699D9D792079F819106FA755CE" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="F7A9F589E5E24FBC5EE8F367917AC64F" bold="true" pageId="0" pageNumber="107">2</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">2</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<tr id="CD036CA0FA7461A3B14B973185DD347B" pageId="0" pageNumber="107">
<td id="C82916B40AE3422F1F8A3A4CF225D8AF" colspan="1" pageId="0" pageNumber="107" rowspan="1">2</td>
<td id="292DD09694F9563FEA76B12D310029E6" colspan="1" pageId="0" pageNumber="107" rowspan="1">
Scutum and frontal triangle densely pollinose (dull) (
<figureCitation captionStart="Figures 2327" captionStartId="F6" captionText="Figures 23 - 27. Enderleiniella longiventris. 23 Male abdomen (lateral) 24 male genitalia (lateral) 25 male genitalia (posterior) 26 male genitalia (ventral) 27 male genitalia (posteroventral). Abbreviations: cer - cercus; hyp - hypandrium; phal - phallus; phap - phallapodeme; pog - postgonite; S 6 - remnant sternite 6; T 7 + 8 - tergite 7 + 8." figureDoi="10.3897/zookeys.884.36154.figures23-27" httpUri="https://binary.pensoft.net/fig/351399" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures23-27">Figs 26</figureCitation>
<figureCitation id="A698BF46FDE95898314ACFBE7040EA25" captionStart="Figures 2327" captionStartId="F6" captionText="Figures 23 - 27. Enderleiniella longiventris. 23 Male abdomen (lateral) 24 male genitalia (lateral) 25 male genitalia (posterior) 26 male genitalia (ventral) 27 male genitalia (posteroventral). Abbreviations: cer - cercus; hyp - hypandrium; phal - phallus; phap - phallapodeme; pog - postgonite; S 6 - remnant sternite 6; T 7 + 8 - tergite 7 + 8." figureDoi="10.3897/zookeys.884.36154.figures23-27" httpUri="https://binary.pensoft.net/fig/351399" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures23-27">Figs 26</figureCitation>
,
<figureCitation captionStart="Figures 3134" captionStartId="F8" captionText="Figures 31 - 34. Enderleiniella maculata. 31 Male genitalia (lateral) 32 male genitalia (posterior) 33 male genitalia (ventral) 34 male genitalia (posteroventral)." figureDoi="10.3897/zookeys.884.36154.figures31-34" httpUri="https://binary.pensoft.net/fig/351401" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures31-34">33</figureCitation>
<figureCitation id="69AF88652C1690F22E422883410B198C" captionStart="Figures 3134" captionStartId="F8" captionText="Figures 31 - 34. Enderleiniella maculata. 31 Male genitalia (lateral) 32 male genitalia (posterior) 33 male genitalia (ventral) 34 male genitalia (posteroventral)." figureDoi="10.3897/zookeys.884.36154.figures31-34" httpUri="https://binary.pensoft.net/fig/351401" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures31-34">33</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">3</emphasis>
<td id="EE132D6430F6E95E51CC59F354ACE468" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="B85A99E658124198B1D96580B17A29C7" bold="true" pageId="0" pageNumber="107">3</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<tr id="91DB3428EE125BA3C2EB244BC795022F" pageId="0" pageNumber="107">
<td id="7608EDFB284F80BF27CB493B2E6CC73D" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="7193C665C87834809A14F1935F4FCECE" colspan="1" pageId="0" pageNumber="107" rowspan="1">
Scutum and frontal triangle glossy or sparsely pollinose (
<figureCitation captionStart="Figures 2022" captionStartId="F5" captionText="Figures 20 - 22. Enderleiniella longiventris. 20 Lateral habitus 21 dorsal habitus 22 geographic distribution." figureDoi="10.3897/zookeys.884.36154.figures20-22" httpUri="https://binary.pensoft.net/fig/351398" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures20-22">Figs 21</figureCitation>
<figureCitation id="47108F85838DC0CBF860791A3689FA7A" captionStart="Figures 2022" captionStartId="F5" captionText="Figures 20 - 22. Enderleiniella longiventris. 20 Lateral habitus 21 dorsal habitus 22 geographic distribution." figureDoi="10.3897/zookeys.884.36154.figures20-22" httpUri="https://binary.pensoft.net/fig/351398" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures20-22">Figs 21</figureCitation>
,
<figureCitation captionStart="Figures 5357" captionStartId="F13" captionText="Figures 53 - 57. Enderleiniella tripunctata. 53 Lateral habitus 54 dorsal habitus 55 male genitalia (lateral) 56 male genitalia (posterior) 57 geographic distribution." figureDoi="10.3897/zookeys.884.36154.figures53-57" httpUri="https://binary.pensoft.net/fig/351406" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures53-57">53</figureCitation>
<figureCitation id="1718850028CEF58C167A6A9A77E6EBE5" captionStart="Figures 5357" captionStartId="F13" captionText="Figures 53 - 57. Enderleiniella tripunctata. 53 Lateral habitus 54 dorsal habitus 55 male genitalia (lateral) 56 male genitalia (posterior) 57 geographic distribution." figureDoi="10.3897/zookeys.884.36154.figures53-57" httpUri="https://binary.pensoft.net/fig/351406" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures53-57">53</figureCitation>
,
<figureCitation captionStart="Figures 5863" captionStartId="F14" captionText="Figures 58 - 63. Enderleiniella tumescens. 58 Lateral habitus 59 dorsal habitus 60 head (dorsal) 61 male genitalia (lateral) 62 male genitalia (posterior) 63 geographic distribution." figureDoi="10.3897/zookeys.884.36154.figures58-63" httpUri="https://binary.pensoft.net/fig/351407" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures58-63">59</figureCitation>
<figureCitation id="6F780370B290768A065F03E31245F785" captionStart="Figures 5863" captionStartId="F14" captionText="Figures 58 - 63. Enderleiniella tumescens. 58 Lateral habitus 59 dorsal habitus 60 head (dorsal) 61 male genitalia (lateral) 62 male genitalia (posterior) 63 geographic distribution." figureDoi="10.3897/zookeys.884.36154.figures58-63" httpUri="https://binary.pensoft.net/fig/351407" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures58-63">59</figureCitation>
)
</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">4</emphasis>
<td id="B77C139B83DBBB0655B95EBACFF1F1EA" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="829DC5751161F1E064C0844E6AC69419" bold="true" pageId="0" pageNumber="107">4</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">3</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Wing hyaline; male epandrium expanded, broader and tall than pre-epandrium; large species, body length 2.7-3.6mm</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. longiventris" pageId="0" pageNumber="107" rank="species" species="longiventris">
<emphasis italics="true" pageId="0" pageNumber="107">E. longiventris</emphasis>
<tr id="354FC9331D535636104B5B10BD17C608" pageId="0" pageNumber="107">
<td id="2D5A98EB7AE1A01EA3361D27E74B875D" colspan="1" pageId="0" pageNumber="107" rowspan="1">3</td>
<td id="859E1F7A4E750115B003C69BCDA45979" colspan="1" pageId="0" pageNumber="107" rowspan="1">Wing hyaline; male epandrium expanded, broader and tall than pre-epandrium; large species, body length 2.7-3.6mm</td>
<td id="FDAD605094EEB06A559F74F8C434E246" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="8C9ED1BEED7C84322569843D443C3B6A" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="C714DC88B2DFE7DF21185BA81A9BA660" lsidName="E. longiventris" pageId="0" pageNumber="107" rank="species" species="longiventris">
<emphasis id="CE09D1E9BAFA53A3BBF10F35F36565ED" italics="true" pageId="0" pageNumber="107">E. longiventris</emphasis>
</taxonomicName>
(Enderlein)
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<tr id="F0E7D7846860C0B283465E21C16104C3" pageId="0" pageNumber="107">
<td id="3D891DDD7649AEF2A9E11C3CCB299319" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="3136DF3389A3EF431DBACBCCADF9A3B2" colspan="1" pageId="0" pageNumber="107" rowspan="1">
Wing with a dark apical spot (
<figureCitation captionStart="Figures 3134" captionStartId="F8" captionText="Figures 31 - 34. Enderleiniella maculata. 31 Male genitalia (lateral) 32 male genitalia (posterior) 33 male genitalia (ventral) 34 male genitalia (posteroventral)." figureDoi="10.3897/zookeys.884.36154.figures31-34" httpUri="https://binary.pensoft.net/fig/351401" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures31-34">Fig. 33</figureCitation>
<figureCitation id="08EB585AA0138D87C300F282F50CD0BB" captionStart="Figures 3134" captionStartId="F8" captionText="Figures 31 - 34. Enderleiniella maculata. 31 Male genitalia (lateral) 32 male genitalia (posterior) 33 male genitalia (ventral) 34 male genitalia (posteroventral)." figureDoi="10.3897/zookeys.884.36154.figures31-34" httpUri="https://binary.pensoft.net/fig/351401" pageId="0" pageNumber="107" tableDoi="10.3897/zookeys.884.36154.figures31-34">Fig. 33</figureCitation>
); male epandrium smaller, not excessively higher than pre-epandrium; smaller species, body length 2.4-2.9mm
</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. maculata" pageId="0" pageNumber="107" rank="species" species="maculata">
<emphasis italics="true" pageId="0" pageNumber="107">E. maculata</emphasis>
<td id="BD68D3B47B7B9265CEA6C5457FAFC902" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="9F85F8A7E8D806874CF88A3901BA992C" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="526F9A489502014D3E3C4339081322A6" lsidName="E. maculata" pageId="0" pageNumber="107" rank="species" species="maculata">
<emphasis id="A39AE63AE8A5588AA8868844D21F8BC4" italics="true" pageId="0" pageNumber="107">E. maculata</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Occiput with dorsolateral pubescent swelling just behind outer vertical bristle (in males because females currently unknown); mouthparts not geniculate (the only species from Venezuela)</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. tumescens" pageId="0" pageNumber="107" rank="species" species="tumescens">
<emphasis italics="true" pageId="0" pageNumber="107">E. tumescens</emphasis>
<tr id="F855502BE51E82CCFD6C436E9555B641" pageId="0" pageNumber="107">
<td id="2813771B7A18F70D3CEAEAE87E0CE25D" colspan="1" pageId="0" pageNumber="107" rowspan="1">4</td>
<td id="5EBC3E87008DDD12D1FFDD4EB48BC661" colspan="1" pageId="0" pageNumber="107" rowspan="1">Occiput with dorsolateral pubescent swelling just behind outer vertical bristle (in males because females currently unknown); mouthparts not geniculate (the only species from Venezuela)</td>
<td id="793E41CCA7D4407A756424665BEE5A57" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="7A25E6768697FB8515D02C720B5EDA0B" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="A02BD8B917BAF66993878FBF94AE9476" lsidName="E. tumescens" pageId="0" pageNumber="107" rank="species" species="tumescens">
<emphasis id="42ACEC4F1AA4D3FEAB366B9301FA5E0D" italics="true" pageId="0" pageNumber="107">E. tumescens</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Occiput not modified, swelling absent; mouthparts geniculate or not (species not known from Venezuela)</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">5</emphasis>
<tr id="BCFB6D2E6A132B70A1828F82C71F0230" pageId="0" pageNumber="107">
<td id="0D8BB623003B9043928EA742B8C59B41" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="7BB9DB5FC45ACC54C69E08E178A350CC" colspan="1" pageId="0" pageNumber="107" rowspan="1">Occiput not modified, swelling absent; mouthparts geniculate or not (species not known from Venezuela)</td>
<td id="04E8354E29DF24046176B989C0F8CD05" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="FBCD613E983EAE3694DFB03D429F0330" bold="true" pageId="0" pageNumber="107">5</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutum 1.2 times as long as wide; Abdominal tergite 3 with setae arising from enlarged punctate sockets (in males because females currently unknown)</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. punctata" pageId="0" pageNumber="107" rank="species" species="punctata">
<emphasis italics="true" pageId="0" pageNumber="107">E. punctata</emphasis>
<tr id="E73EB8EACA7A314EF62F4753DD50C6C2" pageId="0" pageNumber="107">
<td id="91B653A262AC1225654235FA1B5A8E11" colspan="1" pageId="0" pageNumber="107" rowspan="1">5</td>
<td id="39E65C7623C6E12DEFD7D36DEEBC1FFD" colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutum 1.2 times as long as wide; Abdominal tergite 3 with setae arising from enlarged punctate sockets (in males because females currently unknown)</td>
<td id="D3C2B746753B8A70CF7437B1B2F0A84F" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="F0F7A49BED606D297B3CA802E3D0B2B7" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="3E210FB2DD10B7B84F47AD92C52C8238" lsidName="E. punctata" pageId="0" pageNumber="107" rank="species" species="punctata">
<emphasis id="36B3529FA0AAA812CA689F57CF54D89C" italics="true" pageId="0" pageNumber="107">E. punctata</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutum at least 1.5 times as long as wide. Abdominal tergite 3 unmodified</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">6</emphasis>
<tr id="E755DA54C8F8E810421129464D05CFB5" pageId="0" pageNumber="107">
<td id="D1237BB7C4E837F8E77D82EC6B14EBC5" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="EE6F79AF277B704C29CE8A26EA854300" colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutum at least 1.5 times as long as wide. Abdominal tergite 3 unmodified</td>
<td id="CD05D28D4AEA8BCF3A8275A105057BF3" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="5B72FEE2A7FE9353753DFCAEEEC7A13C" bold="true" pageId="0" pageNumber="107">6</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum rectangular; apical scutellar tubercles long at least 1/5 the length of the scutellum</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. maya" pageId="0" pageNumber="107" rank="species" species="maya">
<emphasis italics="true" pageId="0" pageNumber="107">E. maya</emphasis>
<tr id="B33DD12CE19D59BBF1F348D725EC5AD4" pageId="0" pageNumber="107">
<td id="D3A2D0F1A6E71528190D231EA6BC15EE" colspan="1" pageId="0" pageNumber="107" rowspan="1">6</td>
<td id="EA9EBC4807D00D19A9F97603443BBE44" colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum rectangular; apical scutellar tubercles long at least 1/5 the length of the scutellum</td>
<td id="F9547911B945FD9C87D37FB119D7581D" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="7810E5668F39FFDBAB1059FA2819BE65" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="13D93F663F168D56A7AED3A7EADF8082" lsidName="E. maya" pageId="0" pageNumber="107" rank="species" species="maya">
<emphasis id="1D80A4F7242BC093CA4BCFFFB3A0049B" italics="true" pageId="0" pageNumber="107">E. maya</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum trapezoidal; apical scutellar tubercles short or long at most 1/8 the length of scutellum</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">7</emphasis>
<tr id="2DA0C0E7697343925656289F1D8ADBCC" pageId="0" pageNumber="107">
<td id="9F83645F53B28CD2B3074B3CF14F6C23" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="E29B56C64F5E360F1FF481F1F023308B" colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum trapezoidal; apical scutellar tubercles short or long at most 1/8 the length of scutellum</td>
<td id="E33ADA730494E842DB6FF33C44E89A48" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="69294635D14B50663E8F09021003B1CF" bold="true" pageId="0" pageNumber="107">7</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutum sparsely pollinose</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. marshalli" pageId="0" pageNumber="107" rank="species" species="marshalli">
<emphasis italics="true" pageId="0" pageNumber="107">E. marshalli</emphasis>
<tr id="36F439A266B50E38E8210F4C2A43BA8C" pageId="0" pageNumber="107">
<td id="667F47B4B4FAFC70B71B5B4193124930" colspan="1" pageId="0" pageNumber="107" rowspan="1">7</td>
<td id="EE42C2328A7FD59FAA948B0DAAE972EF" colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutum sparsely pollinose</td>
<td id="999A5EA4DD739161D0552180C042586D" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="C61703D12F69F59AA6777AA642873341" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="257BF7106DDA7BA6C57A123B8E701D26" lsidName="E. marshalli" pageId="0" pageNumber="107" rank="species" species="marshalli">
<emphasis id="E6BB2D18DC21BFEDB7BAD89623A83946" italics="true" pageId="0" pageNumber="107">E. marshalli</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutum polished</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">8</emphasis>
<tr id="5A3D679D93330DE84403AEAE9339F8A3" pageId="0" pageNumber="107">
<td id="DA322C2E01623CFD0F9ACC1DE6D55F6E" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="F4AFE480E9FCF145E167D182C1385AB6" colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutum polished</td>
<td id="83453926D1B63F26409F18FF47F40469" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="8B2B88A2C0CAEF3E2D89904C3CD567D9" bold="true" pageId="0" pageNumber="107">8</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Mouthparts geniculate</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">9</emphasis>
<tr id="B1403AEBDE2A01C43756EFA176B64644" pageId="0" pageNumber="107">
<td id="852C4F0C65A7787E4C191E77B10E52DE" colspan="1" pageId="0" pageNumber="107" rowspan="1">8</td>
<td id="7C64ECF8C7E056339E3FEE5054A6B045" colspan="1" pageId="0" pageNumber="107" rowspan="1">Mouthparts geniculate</td>
<td id="052243BD04140A9B3E3375DEF14405A2" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="509502A2753A92247221812B9D2F55D7" bold="true" pageId="0" pageNumber="107">9</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Mouthparts not geniculate or elongate</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">10</emphasis>
<tr id="52D749F95CED691F68F6BDAC54DDDF80" pageId="0" pageNumber="107">
<td id="FE750695700D1225A1098354C22FB0A3" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="F08A49378A2C57C93E1E5E1888034554" colspan="1" pageId="0" pageNumber="107" rowspan="1">Mouthparts not geniculate or elongate</td>
<td id="99C151707625A1E395ACC57B4C167CB3" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="6FC4B1B06B72AF136B81C2093E6A6BFB" bold="true" pageId="0" pageNumber="107">10</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum 0.8 times as wide as long, with distinct tubercles, Male epandrium large, surstylus parallel sided with round tip</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. cryptica" pageId="0" pageNumber="107" rank="species" species="cryptica">
<emphasis italics="true" pageId="0" pageNumber="107">E. cryptica</emphasis>
<tr id="D9DB28917921C338B44C9ADBCAA61B93" pageId="0" pageNumber="107">
<td id="5AD889979874E405368E079C753A0656" colspan="1" pageId="0" pageNumber="107" rowspan="1">9</td>
<td id="F18355F38AA20CF70E0BCD8DA3344779" colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum 0.8 times as wide as long, with distinct tubercles, Male epandrium large, surstylus parallel sided with round tip</td>
<td id="C81E73F4F62F5C40E8B720913100BFD6" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="12CDCA4371D01A7FCFB09B4549941F72" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="14491A0CE72CC51430E59B79B080D58F" lsidName="E. cryptica" pageId="0" pageNumber="107" rank="species" species="cryptica">
<emphasis id="6BDE621E4BF3275371D32398E4BAAC8B" italics="true" pageId="0" pageNumber="107">E. cryptica</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum 0.6 times as long as wide, with very small tubercles; Male genitalia small, surstylus triangular with pointed tip</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. wheeleri" pageId="0" pageNumber="107" rank="species" species="wheeleri">
<emphasis italics="true" pageId="0" pageNumber="107">E. wheeleri</emphasis>
<tr id="081132480DA66AC7B834816CD9F359CD" pageId="0" pageNumber="107">
<td id="F7C686A23096F0416D9A605AD5570ED0" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="71ECA4FAB2F215A8E1E81E8E4C0D9F3B" colspan="1" pageId="0" pageNumber="107" rowspan="1">Scutellum 0.6 times as long as wide, with very small tubercles; Male genitalia small, surstylus triangular with pointed tip</td>
<td id="40767685E396999382434346B584C517" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="5FD4DA543BE5BFE349DBB347D4F0C857" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="B939EC177F3BD4F7A1F1C7CB0518CC18" lsidName="E. wheeleri" pageId="0" pageNumber="107" rank="species" species="wheeleri">
<emphasis id="3971706636A5B86CDA70532B50B89A42" italics="true" pageId="0" pageNumber="107">E. wheeleri</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">10</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Hairs on scutum in well-defined rows between punctate dorsocentral rows (grooves). In male, epandrium very wide compared to high, cerci in lateral view directed posteroventrally</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. caerulea" pageId="0" pageNumber="107" rank="species" species="caerulea">
<emphasis italics="true" pageId="0" pageNumber="107">E. caerulea</emphasis>
<tr id="9A162D9003B04BF79EB8A0AD6E7F15C3" pageId="0" pageNumber="107">
<td id="C03C2D7C58F2924872943A09DC4C19E3" colspan="1" pageId="0" pageNumber="107" rowspan="1">10</td>
<td id="DDD50A1C2A65387921671ADCC3A858CE" colspan="1" pageId="0" pageNumber="107" rowspan="1">Hairs on scutum in well-defined rows between punctate dorsocentral rows (grooves). In male, epandrium very wide compared to high, cerci in lateral view directed posteroventrally</td>
<td id="56020CCE8D229202AD87D4B7CC4651D2" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="065F34A7C1338F4DB9FFF445DB9A0EC0" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="1BCD2332A8BD6D3C854A9C4ED92106CD" lsidName="E. caerulea" pageId="0" pageNumber="107" rank="species" species="caerulea">
<emphasis id="5A6020CC8B7E80B72C5E9BDA7ACCA714" italics="true" pageId="0" pageNumber="107">E. caerulea</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="107">
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">Hairs on scutum scattered between punctate rows. In male, epandrium square, cerci in lateral view directed ventrally</td>
<td colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="107">
<taxonomicName lsidName="E. tripunctata" pageId="0" pageNumber="107" rank="species" species="tripunctata">
<emphasis italics="true" pageId="0" pageNumber="107">E. tripunctata</emphasis>
<tr id="FEE25C1241B38B4332D207B6972432F8" pageId="0" pageNumber="107">
<td id="88CF4879F6F41A410DEF7ADDFF2CCB0F" colspan="1" pageId="0" pageNumber="107" rowspan="1">-</td>
<td id="582228B78B6236F5978088D8BF4E0E9F" colspan="1" pageId="0" pageNumber="107" rowspan="1">Hairs on scutum scattered between punctate rows. In male, epandrium square, cerci in lateral view directed ventrally</td>
<td id="9E0C93B47DFD459D62C638DD395577CC" colspan="1" pageId="0" pageNumber="107" rowspan="1">
<emphasis id="729C1AC10DB308A8B0D08122E0BCFCA0" bold="true" pageId="0" pageNumber="107">
<taxonomicName id="65690B784EAB4F7023CEA49FF352FBF3" lsidName="E. tripunctata" pageId="0" pageNumber="107" rank="species" species="tripunctata">
<emphasis id="A0DAE5D4115C38765015B479DFBA4779" italics="true" pageId="0" pageNumber="107">E. tripunctata</emphasis>
</taxonomicName>
(Becker)
</emphasis>

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@ -1,171 +1,171 @@
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<mods:title>Review of the world species of Paroplitis Mason, 1981 (Hymenoptera, Braconidae, Microgastrinae), with description of three new species</mods:title>
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<mods:title id="AF9399825E293754F62AE8C1FC089D17">Review of the world species of Paroplitis Mason, 1981 (Hymenoptera, Braconidae, Microgastrinae), with description of three new species</mods:title>
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<mods:namePart>Fujie, Shunpei</mods:namePart>
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<mods:namePart>Japoshvili, George</mods:namePart>
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<taxonomicName LSID="http://zoobank.org/6A970B8F-A95B-43BF-9B46-35A1F1BF21C9" authority="Japoshvili, Fujie &amp; Fernandez-Triana" authorityName="Japoshvili, Fujie &amp; Fernandez-Triana" authorityYear="2021" class="Insecta" family="Braconidae" genus="Paroplitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroplitis kakhetiensis" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="kakhetiensis" status="sp. nov.">Paroplitis kakhetiensis Japoshvili, Fujie &amp; Fernandez-Triana</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="33">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 2027" captionStartId="F5" captionText="Figures 20 - 27. Paroplitis kakhetiensis, holotype. 20. Habitus; 21. Fore wing; 22. Dorsal view of mesoscutellar disc, propodeum and mediotergites; 23. Apical segments of antenna; 24. Basal segments of antenna; 25. Frontal view of head; 26. Dorsal view of head; 27. Detail of T 1 and T 2, dorsal view." figureDoi="10.3897/dez.68.59641.figures20-27" httpUri="https://binary.pensoft.net/fig/494140" pageId="0" pageNumber="33">Figs 20-27</figureCitation>
<treatment id="C9C8F128DD4455B19C655C184D0E7D2B" ID-GBIF-Taxon="176435555" LSID="urn:lsid:zoobank.org:act:6A970B8F-A95B-43BF-9B46-35A1F1BF21C9" httpUri="http://treatment.plazi.org/id/C9C8F128DD4455B19C655C184D0E7D2B" lastPageNumber="33" pageId="0" pageNumber="33" scope_family="Braconidae" scope_order="Hymenoptera">
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<taxonomicName id="8D50DD14C9CB895B9AEDB105A45B86B6" LSID="http://zoobank.org/6A970B8F-A95B-43BF-9B46-35A1F1BF21C9" authority="Japoshvili, Fujie &amp; Fernandez-Triana" authorityName="Japoshvili, Fujie &amp; Fernandez-Triana" authorityYear="2021" class="Insecta" family="Braconidae" genus="Paroplitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroplitis kakhetiensis" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="kakhetiensis" status="sp. nov.">Paroplitis kakhetiensis Japoshvili, Fujie &amp; Fernandez-Triana</taxonomicName>
<taxonomicNameLabel id="0630E7E30E476FE21211254BA89850E7" pageId="0" pageNumber="33">sp. nov.</taxonomicNameLabel>
<figureCitation id="81C846E6125142209C685301501F9E64" captionStart="Figures 2027" captionStartId="F5" captionText="Figures 20 - 27. Paroplitis kakhetiensis, holotype. 20. Habitus; 21. Fore wing; 22. Dorsal view of mesoscutellar disc, propodeum and mediotergites; 23. Apical segments of antenna; 24. Basal segments of antenna; 25. Frontal view of head; 26. Dorsal view of head; 27. Detail of T 1 and T 2, dorsal view." figureDoi="10.3897/dez.68.59641.figures20-27" httpUri="https://binary.pensoft.net/fig/494140" pageId="0" pageNumber="33">Figs 20-27</figureCitation>
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<paragraph pageId="0" pageNumber="33">Holotype.</paragraph>
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<paragraph id="7AA8750E7B4ADBFE16AE06DFAC210D32" pageId="0" pageNumber="33">Holotype.</paragraph>
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Female, AUG. Holotype labels: Georgia: Lagodekhi Reserve, Mt. Kudigora,
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,
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, 1,841 m alt., 5-15. V. 2014, Malaise Trap, G. Japoshvili, CNC507801.
</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="33">Figures 20-27.</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="33">Paroplitis kakhetiensis</emphasis>
<caption id="C9DE14AF1859F6B7E47803696C26C2BC" doi="10.3897/dez.68.59641.figures20-27" httpUri="https://binary.pensoft.net/fig/494140" pageId="0" pageNumber="33" start="Figures 2027" startId="F5">
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<emphasis id="D7D94D2D67C893807A15A9E69F8A5785" bold="true" pageId="0" pageNumber="33">Figures 20-27.</emphasis>
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<emphasis id="27535AE1B711F3272FE88D59BB988824" italics="true" pageId="0" pageNumber="33">Paroplitis kakhetiensis</emphasis>
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, holotype.
<emphasis bold="true" pageId="0" pageNumber="33">20.</emphasis>
<emphasis id="322CD52FF04F848639052506C3DE1F83" bold="true" pageId="0" pageNumber="33">20.</emphasis>
Habitus;
<emphasis bold="true" pageId="0" pageNumber="33">21.</emphasis>
<emphasis id="17A377DC168E87ED9556F65499B6C83A" bold="true" pageId="0" pageNumber="33">21.</emphasis>
Fore wing;
<emphasis bold="true" pageId="0" pageNumber="33">22.</emphasis>
<emphasis id="9F8567E97F5F3A5C59DF037745CBBD70" bold="true" pageId="0" pageNumber="33">22.</emphasis>
Dorsal view of mesoscutellar disc, propodeum and mediotergites;
<emphasis bold="true" pageId="0" pageNumber="33">23.</emphasis>
<emphasis id="B251C5F9A7B756F22B09D40D9FFDD4EE" bold="true" pageId="0" pageNumber="33">23.</emphasis>
Apical segments of antenna;
<emphasis bold="true" pageId="0" pageNumber="33">24.</emphasis>
<emphasis id="E16B1F4EB650FCB69F2B5EB8BD1CFC59" bold="true" pageId="0" pageNumber="33">24.</emphasis>
Basal segments of antenna;
<emphasis bold="true" pageId="0" pageNumber="33">25.</emphasis>
<emphasis id="E1D7574E1E71B41058A692627DAD44BC" bold="true" pageId="0" pageNumber="33">25.</emphasis>
Frontal view of head;
<emphasis bold="true" pageId="0" pageNumber="33">26.</emphasis>
<emphasis id="EF366F790D6F6950597022E89B5C0745" bold="true" pageId="0" pageNumber="33">26.</emphasis>
Dorsal view of head;
<emphasis bold="true" pageId="0" pageNumber="33">27.</emphasis>
<emphasis id="7050AE60263B259EF41C5FEA5429EFAA" bold="true" pageId="0" pageNumber="33">27.</emphasis>
Detail of T1 and T2, dorsal view.
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<paragraph pageId="0" pageNumber="33">Description.</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Female</emphasis>
(n = 1). Body length: 2.4 mm. Fore wing length: 2.4 mm. F2 L/W: 1.4
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. F14 L/W: 1.3
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. F15 L/W: 1.3
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. F2 L/F14 L: 1.2
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. OOL/OD: 2.1
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. POL/OD: 1.4
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. Fore wing with vein 2CUa entirely nebulous; vein R1 shorter than pterostigma length and a little longer than distance delimited between end of vein R1 and end of vein 3RSb. Fore wing with areolet triangular and relatively small, its maximum height 0.3
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vein r length, its maximum width 0.5
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vein r length. Propodeum mostly smooth and shiny, with some rugosity longitudinally and along median transverse area, without trace of some transverse carina; median longitudinal carina complete at least on anterior 0.5; propodeal areola present medio-posteriorly. Metafemur L/W: 2.5
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. Anterior 0.5 of T1 coarsely punctate-rugose, rest of T1 and T2 mostly smooth; T1 median length 1.7
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its width at posterior margin; T2 width at posterior margin 1.8
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its median length. Metatibia L: 0.79 mm. Metatibia L/ovipositor sheath L: 2.9
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<paragraph id="721762DD12E3DF5B7CACB91426D4ED36" pageId="0" pageNumber="33">Description.</paragraph>
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<emphasis id="7DED35CE6763EAC6CD24554A468D217B" bold="true" pageId="0" pageNumber="33">Female</emphasis>
(n = 1). Body length: 2.4 mm. Fore wing length: 2.4 mm. F2 L/W: 1.4
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. F14 L/W: 1.3
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. F15 L/W: 1.3
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. F2 L/F14 L: 1.2
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. OOL/OD: 2.1
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. POL/OD: 1.4
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. Fore wing with vein 2CUa entirely nebulous; vein R1 shorter than pterostigma length and a little longer than distance delimited between end of vein R1 and end of vein 3RSb. Fore wing with areolet triangular and relatively small, its maximum height 0.3
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vein r length, its maximum width 0.5
<normalizedToken id="8DC2AB5EC15AE6DA6AA34B11DC7A8F6F" originalValue="×">x</normalizedToken>
vein r length. Propodeum mostly smooth and shiny, with some rugosity longitudinally and along median transverse area, without trace of some transverse carina; median longitudinal carina complete at least on anterior 0.5; propodeal areola present medio-posteriorly. Metafemur L/W: 2.5
<normalizedToken id="289148826943A9AD11CDCE4B993F7263" originalValue="×">x</normalizedToken>
. Anterior 0.5 of T1 coarsely punctate-rugose, rest of T1 and T2 mostly smooth; T1 median length 1.7
<normalizedToken id="DC0BD756FCFAAA36CD23216D194A7CE5" originalValue="×">x</normalizedToken>
its width at posterior margin; T2 width at posterior margin 1.8
<normalizedToken id="22C5DAC9440A32B8E7123DBA3747F779" originalValue="×">x</normalizedToken>
its median length. Metatibia L: 0.79 mm. Metatibia L/ovipositor sheath L: 2.9
<normalizedToken id="1D3FCFF62BE7DC4E7085BABB4590C41D" originalValue="×">x</normalizedToken>
. Ovipositor sheath L: 0.27 mm. Maximum length of setae on ovipositor sheath at most slightly longer than maximum width of ovipositor sheath.
</paragraph>
<paragraph pageId="0" pageNumber="33">Body dark brown to black. Mouth parts, humeral complex, wing veins, trochantellus, apical part of pro- and mesofemora, pro- and mesotibiae and tarsi and anterior 0.2 of metatibia brown to yellowish-brown. Palpi yellow.</paragraph>
<paragraph pageId="0" pageNumber="33">
<emphasis bold="true" pageId="0" pageNumber="33">Male.</emphasis>
<paragraph id="3C2E2178AAA1DBAAB14FF56BA30E063F" pageId="0" pageNumber="33">Body dark brown to black. Mouth parts, humeral complex, wing veins, trochantellus, apical part of pro- and mesofemora, pro- and mesotibiae and tarsi and anterior 0.2 of metatibia brown to yellowish-brown. Palpi yellow.</paragraph>
<paragraph id="7B53D1EA000452381391DE74C1B2B56A" pageId="0" pageNumber="33">
<emphasis id="24C9F04673BCAD71DFD7FEC5495EBAC5" bold="true" pageId="0" pageNumber="33">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="host">
<paragraph pageId="0" pageNumber="33">Hosts.</paragraph>
<paragraph pageId="0" pageNumber="33">Unknown.</paragraph>
<subSubSection id="FC19890E5CE287C2535CD3E325BC4303" pageId="0" pageNumber="33" type="host">
<paragraph id="F79160065CA2F630BEAA79A841ED6E49" pageId="0" pageNumber="33">Hosts.</paragraph>
<paragraph id="56209FDBC9BB80D0F6EB399C1F73BE8B" pageId="0" pageNumber="33">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="distribution">
<paragraph pageId="0" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="33">Western Palaearctic Region: Georgia.</paragraph>
<subSubSection id="3B677AE5616AA0B3863201D65A16DCF3" pageId="0" pageNumber="33" type="distribution">
<paragraph id="F250F873601B3F8E35A2F50BC6AC65CF" pageId="0" pageNumber="33">Distribution.</paragraph>
<paragraph id="F5B15870557C2DB03F9D17A7E00C26A4" pageId="0" pageNumber="33">Western Palaearctic Region: Georgia.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="etymology">
<paragraph pageId="0" pageNumber="33">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="33">The species is named after the region in Georgia (Kakheti), where it was found.</paragraph>
<subSubSection id="85C22D5F20D0C316536AF818A2A33E6A" pageId="0" pageNumber="33" type="etymology">
<paragraph id="E07D8AEC8391E8BA96F712ACCC5EDC1B" pageId="0" pageNumber="33">Etymology.</paragraph>
<paragraph id="2A1F6D99A1C1975CFEA29B9FB3EBEC07" pageId="0" pageNumber="33">The species is named after the region in Georgia (Kakheti), where it was found.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="comments">
<paragraph pageId="0" pageNumber="33">Comments.</paragraph>
<paragraph pageId="0" pageNumber="33">
<subSubSection id="B261CE4B988C716EA6364C3F8F8AE0A0" pageId="0" pageNumber="33" type="comments">
<paragraph id="8A33AAB20D077DE05182CA4C10E6E7B0" pageId="0" pageNumber="33">Comments.</paragraph>
<paragraph id="4CFCA9D1362C31EADE3B42EDB0AB492E" pageId="0" pageNumber="33">
The distribution of
<taxonomicName genus="P." lsidName="P. kakhetiensis" pageId="0" pageNumber="33" rank="species" species="kakhetiensis">
<emphasis italics="true" pageId="0" pageNumber="33">P. kakhetiensis</emphasis>
<taxonomicName id="E740F54FCA859B8472CAB7CCAA31A22C" lsidName="P. kakhetiensis" pageId="0" pageNumber="33" rank="species" species="kakhetiensis">
<emphasis id="97B0EA296E6FE8263C00DCE0E1E0611E" italics="true" pageId="0" pageNumber="33">P. kakhetiensis</emphasis>
</taxonomicName>
seems to overlap with that of
<taxonomicName genus="P." lsidName="P. wesmaeli" pageId="0" pageNumber="33" rank="species" species="wesmaeli">
<emphasis italics="true" pageId="0" pageNumber="33">P. wesmaeli</emphasis>
<taxonomicName id="687BEF59A44B33B3D3663A5457F0A3CE" lsidName="P. wesmaeli" pageId="0" pageNumber="33" rank="species" species="wesmaeli">
<emphasis id="CF558F9BC84121438282825707FE9386" italics="true" pageId="0" pageNumber="33">P. wesmaeli</emphasis>
</taxonomicName>
, although
<taxonomicName genus="P." lsidName="P. kakhetiensis" pageId="0" pageNumber="33" rank="species" species="kakhetiensis">
<emphasis italics="true" pageId="0" pageNumber="33">P. kakhetiensis</emphasis>
<taxonomicName id="2E9E7371DACA03354034C8A5DA570F8F" lsidName="P. kakhetiensis" pageId="0" pageNumber="33" rank="species" species="kakhetiensis">
<emphasis id="0810AFF4D5713BF2CF3D24B91F046F63" italics="true" pageId="0" pageNumber="33">P. kakhetiensis</emphasis>
</taxonomicName>
was collected at a higher altitude (1840 m) than
<taxonomicName baseAuthorityName="Ruthe" baseAuthorityYear="1860" class="Insecta" family="Braconidae" genus="Paroplitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroplitis wesmaeli" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wesmaeli">
<emphasis italics="true" pageId="0" pageNumber="33">Paroplitis wesmaeli</emphasis>
<taxonomicName id="0D581DC9FBD6F98E94E87A7E1D0E20EF" baseAuthorityName="Ruthe" baseAuthorityYear="1860" class="Insecta" family="Braconidae" genus="Paroplitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroplitis wesmaeli" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wesmaeli">
<emphasis id="2B1E678DA3EEDFF6C5B95FC684341A3C" italics="true" pageId="0" pageNumber="33">Paroplitis wesmaeli</emphasis>
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specimens.
</paragraph>

View file

@ -1,183 +1,183 @@
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<taxonomicName LSID="C9C92FC1-C345-57A2-B7A5-1C7FA3FB8613" authority="Seyrig, 1932" authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion acaule" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="acaule">Tetractenion acaule Seyrig, 1932</taxonomicName>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tetractenion acaule Lectotype (MNHN) A habitus, lateral view B habitus, dorsal view C head, anterior view D data labels. Photographs of lectotype RECOLNAT (ANR- 11 - INBS- 0004) - Christophe Herve - 2014. http: // coldb. mnhn. fr / catalognumber / mnhn / ey / ey 9333 (used with permission of Agniele Touret-Alby - Curator of Hymenoptera MNHN)." figureDoi="10.3897/zookeys.1007.55543.figure1" httpUri="https://binary.pensoft.net/fig/492439" pageId="0" pageNumber="49">Fig. 1</figureCitation>
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<paragraph pageId="0" pageNumber="49">Type material.</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Lectotype</emphasis>
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<emphasis id="CEAB0870DAB65733A4AFFABB92DDC4D6" bold="true" italics="true" pageId="0" pageNumber="49">Lectotype</emphasis>
♀: Madagascar, Rogez,
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Cote Est,
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Paris, 1.31. A. Seyrig, EY9333, [White label with TYPE written in red] [Red type label]: Lectotype ♀
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<emphasis italics="true" pageId="0" pageNumber="49">Tectractenion acuale</emphasis>
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, Seyrig, 1932, designated by
<bibRefCitation author="Townes, HK" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="0" pageNumber="49" pagination="1 - 416" refId="B39" refString="Townes, HK, Townes, M, 1973. A catalogue and reclassification of the Ethiopian Ichneumonidae. Memoirs of the American Entomological Institute 19: 1 - 416" title="A catalogue and reclassification of the Ethiopian Ichneumonidae." volume="19" year="1973">Townes and Townes (1973)</bibRefCitation>
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, labeled by T. Yoshida, 2011 (MNHN) (photos of Lectotype examined: http://coldb.mnhn.fr/catalognumber/mnhn/ey/ey9333).
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Non-type</emphasis>
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♀ (examined): Madagascar, Bekily, Reg. Sud. de
<normalizedToken originalValue="Lile">L'ile</normalizedToken>
<normalizedToken id="468FDF940E8C780BECD36C43C83FE8CE" originalValue="Lile">L'ile</normalizedToken>
, Feb 1930 and Jan-Feb 1931, Coll. Mus. Congo, Col. P.L.G. Benoit,
<taxonomicName genus="Tectractenion" lsidName="Tectractenion acuale" pageId="0" pageNumber="49" rank="species" species="acuale">Tectractenion acuale</taxonomicName>
<taxonomicName id="C67EDD59F20A2896F55C7D8A99AC8C9C" genus="Tectractenion" lsidName="Tectractenion acuale" pageId="0" pageNumber="49" rank="species" species="acuale">Tectractenion acuale</taxonomicName>
, det. P.L.G. Benoit, 1953 (RMNH).
<emphasis bold="true" pageId="0" pageNumber="49">
<emphasis italics="true" pageId="0" pageNumber="49">Additional material</emphasis>
<emphasis id="8114E3D79F5D0C0C9C5A7A9D0BE06B2B" bold="true" pageId="0" pageNumber="49">
<emphasis id="A87938509A36DB67331872E53F5F61E1" italics="true" pageId="0" pageNumber="49">Additional material</emphasis>
.
</emphasis>
♀: Madagascar: Majunga Prov., Besalampy District, Marofototra dry forest, 17 km W of Besalampy, 4-11 February 2008,
<geoCoordinate degrees="16" direction="south" minutes="43.30" orientation="latitude" precision="9" value="-16.721666">16°43.30'S</geoCoordinate>
<geoCoordinate id="6D304183CD84743499E6AFFE3B6E19E5" degrees="16" direction="south" minutes="43.30" orientation="latitude" precision="9" value="-16.721666">16°43.30'S</geoCoordinate>
,
<geoCoordinate degrees="44" direction="east" minutes="25.42" orientation="longitude" precision="9" value="44.423668">44°25.42'E</geoCoordinate>
<geoCoordinate id="83D59391BB3100A0E55573EDC6B5BAE5" degrees="44" direction="east" minutes="25.42" orientation="longitude" precision="9" value="44.423668">44°25.42'E</geoCoordinate>
, coll. M. Irwin, R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
<normalizedToken id="F8AB2F0E0C78DC0BBC11677DD665D69C" originalValue="HarinHala">Harin'Hala</normalizedToken>
, Malaise, dry wash in forest, elev. 170 ft MG-42A-20 (CASC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="differential diagnosis">
<paragraph pageId="0" pageNumber="49">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="49">
<taxonomicName authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion acaule" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="acaule">
<emphasis italics="true" pageId="0" pageNumber="49">Tetractenion acaule</emphasis>
<subSubSection id="C3C973DB1D0D2C6C773B6ED3CBE8D161" pageId="0" pageNumber="49" type="differential diagnosis">
<paragraph id="857E09071B93E66EA4532849C5C54CA8" pageId="0" pageNumber="49">Differential diagnosis.</paragraph>
<paragraph id="83087FC1E2E0FD91A10FE922AC7169CF" pageId="0" pageNumber="49">
<taxonomicName id="9E8CC77FFC9E9F0F79B5B0FBB522E824" authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion acaule" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="acaule">
<emphasis id="E4E369C471DABCE158EFBE02AB961B42" italics="true" pageId="0" pageNumber="49">Tetractenion acaule</emphasis>
</taxonomicName>
is immediately distinguishable from all other
<taxonomicName authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="49">Tetractenion</emphasis>
<taxonomicName id="14D66EFFCF8DE81D9B4CCAD2B4C0E611" authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="2708C15E71028AB329B4657D9F404578" italics="true" pageId="0" pageNumber="49">Tetractenion</emphasis>
</taxonomicName>
species by its unique color combination of a red mesosoma and a mostly black metasoma; distinct keels are present on outer mesoscutal lobes, the notauli do not reach the scutellum; metasomal tergite I has a distinct medial compression in the dorso-ventral view, tergite II have distinct gastrocoeli, and tergites IV-VIII are dorso-posteriorly weakly sclerotized, appearing as large membranous white areas on the dorsal surface.
<taxonomicName authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion acaule" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="acaule">
<emphasis italics="true" pageId="0" pageNumber="49">Tetractenion acaule</emphasis>
<taxonomicName id="B8483D271D27725AF87CE56A5A6A8FAF" authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion acaule" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="acaule">
<emphasis id="B11C074E0B6E0EB09F986BB41E84837F" italics="true" pageId="0" pageNumber="49">Tetractenion acaule</emphasis>
</taxonomicName>
closely resembles
<taxonomicName lsidName="T. ibayaensis" pageId="0" pageNumber="49" rank="species" species="ibayaensis">
<emphasis italics="true" pageId="0" pageNumber="49">T. ibayaensis</emphasis>
<taxonomicName id="410050BEBCBAEE1F2BBA3217BF71F563" lsidName="T. ibayaensis" pageId="0" pageNumber="49" rank="species" species="ibayaensis">
<emphasis id="604AC6502FB671A93653B9F4BAAF0072" italics="true" pageId="0" pageNumber="49">T. ibayaensis</emphasis>
</taxonomicName>
as both species are similar in color, having largely fulvous bodies with a white face and the hind femur infuscate, whereas the remaining
<taxonomicName authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="49">Tetractenion</emphasis>
<taxonomicName id="1669E52CC89D2275D54F186FC52CF65F" authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="9D825CF1020ACB762600FB7ED9902925" italics="true" pageId="0" pageNumber="49">Tetractenion</emphasis>
</taxonomicName>
species are largely yellow in color with yellow hind femora.
<taxonomicName authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion acaule" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="acaule">
<emphasis italics="true" pageId="0" pageNumber="49">Tetractenion acaule</emphasis>
<taxonomicName id="543368A9C49B723F408A3105C208204D" authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion acaule" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="acaule">
<emphasis id="C70C10BC0B4975F07F2C57D5E068F4DC" italics="true" pageId="0" pageNumber="49">Tetractenion acaule</emphasis>
</taxonomicName>
can easily be distinguished from
<taxonomicName lsidName="T. ibayaensis" pageId="0" pageNumber="49" rank="species" species="ibayaensis">
<emphasis italics="true" pageId="0" pageNumber="49">T. ibayaensis</emphasis>
<taxonomicName id="22487ADE223058759ADAA8F2944A9363" lsidName="T. ibayaensis" pageId="0" pageNumber="49" rank="species" species="ibayaensis">
<emphasis id="9822A12D41CED1ABD45E40C98633710C" italics="true" pageId="0" pageNumber="49">T. ibayaensis</emphasis>
</taxonomicName>
by having a white gena and weakly sclerotized metasomal tergites IV-VIII; the head is narrow, straight behind the eyes; a distinct carina is present on the pronotal collar; distinct keels are present on the outer mesoscutal lobes, with the notauli not reaching the scutellum; pits on the mesopleuron and propodeum are shallow; metasomal tergite I is distinctly dorso-medially compressed; gastrocoeli on tergite II are distinct; tergites IV-VIII are postero-dorsally weakly sclerotized and white; and tarsal claws on the hind leg are simple. In
<taxonomicName lsidName="T. ibayaensis" pageId="0" pageNumber="49" rank="species" species="ibayaensis">
<emphasis italics="true" pageId="0" pageNumber="49">T. ibayaensis</emphasis>
<taxonomicName id="32B3550488085E073E8694321FDEA584" lsidName="T. ibayaensis" pageId="0" pageNumber="49" rank="species" species="ibayaensis">
<emphasis id="761193CBB253523090B0B94925548BDF" italics="true" pageId="0" pageNumber="49">T. ibayaensis</emphasis>
</taxonomicName>
the gena is brown and tergites IV-VIII are strongly sclerotized; the head is rounded behind the eyes; with no more than a wrinkle present on the pronotal collar; the mesoscutal lobes are hardly present, the notauli reach the scutellum; pits on the mesopleuron and propodeum are deep; metasomal tergite I is stout and indistinctly dorso-ventrally compressed in the medial region; gastrocoeli on tergite II are indistinct; and tarsal claws on the hind legs are pectinate.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="description">
<paragraph pageId="0" pageNumber="49">Description</paragraph>
<paragraph pageId="0" pageNumber="49">
<subSubSection id="212F0F77002A408DAEA5EBC7A021AD99" pageId="0" pageNumber="49" type="description">
<paragraph id="C90581CE18BCC69B9C87C7FBE2A94F46" pageId="0" pageNumber="49">Description</paragraph>
<paragraph id="006E9BF93E164C7AE97DF0B4B2E9DFDF" pageId="0" pageNumber="49">
(updated from
<bibRefCitation author="Seyrig, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="49" refId="B31" refString="Seyrig, A, 1932. Les Ichneumonides de Madagascar. I IchneumonidaePimplinae. Memoires de l'Academie Malgache. Fascicule 11, 183 pp." title="Les Ichneumonides de Madagascar. I IchneumonidaePimplinae. Memoires de l'Academie Malgache. Fascicule 11, 183 pp." year="1932">Seyrig 1932</bibRefCitation>
<bibRefCitation id="00D3380AE22956309F8C9109BBD6241C" author="Seyrig, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="49" refId="B31" refString="Seyrig, A, 1932. Les Ichneumonides de Madagascar. I IchneumonidaePimplinae. Memoires de l'Academie Malgache. Fascicule 11, 183 pp." title="Les Ichneumonides de Madagascar. I IchneumonidaePimplinae. Memoires de l'Academie Malgache. Fascicule 11, 183 pp." year="1932">Seyrig 1932</bibRefCitation>
). Size 9-11 mm.
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Color</emphasis>
<emphasis id="7C430535E38CB37D71CA9EE1DE7279A5" bold="true" italics="true" pageId="0" pageNumber="49">Color</emphasis>
: head white with a large black central area on occiput, reaching eyes on vertex and pointed on frons; antenna black, without pale ring; mesosoma red; metasomal tergites I and II red, though tergite II sometimes brownish, following tergites black with large membranous white areas from tergite IV; legs red, hind femur, tibia and tarsus infuscate; wings with sparse microtrichia, venation brown, pterostigma brown and centrally translucent reddish.
</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Head</emphasis>
<paragraph id="38084BFC6FC7731DCB0D0D883557A86F" pageId="0" pageNumber="49">
<emphasis id="9F831398E202FC45F4179C64EBC6C0DE" bold="true" italics="true" pageId="0" pageNumber="49">Head</emphasis>
narrow, straight behind eyes; occiput deeply and angularly excavated, occipital carina strong, extending to lower gena at base of mandible; eyes very large; malar space almost half as long as mandibular basal width; face and clypeus finely, evenly and rather sparsely punctate on a shiny background; face with three lobes, tentorial pits deep; clypeus small, laterally convex with declivity, apically invaginated, with clypeal edge convex; mandibular teeth triangular, lower tooth longer than upper tooth; antenna long, slender and apically tapered.
</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Mesosoma</emphasis>
<paragraph id="2D190E3BB02E8A664774593B862DE981" pageId="0" pageNumber="49">
<emphasis id="7E23319913EF7AF1812096B343DF1968" bold="true" italics="true" pageId="0" pageNumber="49">Mesosoma</emphasis>
stout; mesonotum deeply punctate, inter-punctuate spaces about as wide as punctures, rather matt, but not coriaceous; keels distinctly raised on outer mesoscutal lobes of mesoscutum, notauli abbreviated, not reaching the scutellum; apex of scutellum rounded; pronotum shining with a distinct thickened carina on collar, sparsely and very finely punctate; mesopleuron higher than wide, sparsely but more deeply punctate, speculum similarly punctate, background hardly shining, epicnemial carina ending at anterior edge of mesopleuron; shallow pits on mesopleuron and propodeum; metapleuron matt and deeply punctate; propodeum weakly convex, roughly punctate dorsally, punctate posteriorly confluently grading into transverse wrinkles, posterior transverse carina reduced, lateral longitudinal carinae present but faint, spiracle roundish-elliptic and small.
</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Metasoma</emphasis>
<paragraph id="8A8F5228DD472D3F16D9B678CA137EB9" pageId="0" pageNumber="49">
<emphasis id="932CE9EA6ED38D951176DCB3FF77BB36" bold="true" italics="true" pageId="0" pageNumber="49">Metasoma</emphasis>
hardly punctate at base of tergite II, indistinctly punctate beyond base; tergite I elongate, more than twice as long as wide, tapered anteriorly, glymma present, spiracle positioned slightly in front of middle and protruding, especially dorsally, with a distinct medial depression dorso-ventrally; tergite II longer than wide or subquadrate with gastrocoeli distinct; tergite III quadrate to transverse; metasomal tergites IV-VIII moderately laterally compressed; ovipositor sheath concealed or hardly protruding.
</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" italics="true" pageId="0" pageNumber="49">Fore wing</emphasis>
<paragraph id="3B54F85E7001D85ACED8571BE1C0FF26" pageId="0" pageNumber="49">
<emphasis id="0473AE53716271B06E296EE0606FE4BE" bold="true" italics="true" pageId="0" pageNumber="49">Fore wing</emphasis>
without ramellus on Rs-M vein; Rs hardly sinuate; areolet large and quadrate with a short stalk receiving 2m-cu at center. Hind wing with Cu1 shorter than cu-a such that Cu2 arises above the middle of these combined veins. Legs very long; hind femur reaching beyond metasomal apex; length of tibia III plus tarsus III as long as body; spurs of tibia III longer than half metatarsal length; tarsal claws on hind leg simple.
</paragraph>
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Male</emphasis>
<paragraph id="7EB5640D6A1B8BA4F31D2C4D1D389923" pageId="0" pageNumber="49">
<emphasis id="820690AD69CD89CC6BD81DBBDB7A92E9" bold="true" pageId="0" pageNumber="49">Male</emphasis>
hardly different: temples a bit less narrowed behind eyes, metasomal tergite II entirely black.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="49" type="distribution">
<paragraph pageId="0" pageNumber="49">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="49">Madagascar.</paragraph>
<caption doi="10.3897/zookeys.1007.55543.figure1" httpUri="https://binary.pensoft.net/fig/492439" pageId="0" pageNumber="49" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="49">
<emphasis bold="true" pageId="0" pageNumber="49">Figure 1.</emphasis>
<taxonomicName authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion acaule" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="acaule">
<emphasis italics="true" pageId="0" pageNumber="49">Tetractenion acaule</emphasis>
<subSubSection id="BB46BF50D8D304E82B5E39D42EE91946" pageId="0" pageNumber="49" type="distribution">
<paragraph id="6C84E2C1CE04ADD6D96C6989F3D405BC" pageId="0" pageNumber="49">Distribution.</paragraph>
<paragraph id="6509794E804300A7F09E3E3CD4A1D71A" pageId="0" pageNumber="49">Madagascar.</paragraph>
<caption id="91D72B8A9FECFAA901FB62DC1E400480" doi="10.3897/zookeys.1007.55543.figure1" httpUri="https://binary.pensoft.net/fig/492439" pageId="0" pageNumber="49" start="Figure 1" startId="F1">
<paragraph id="6DC8A1E91BC41C3B17C9100752C1EBC0" pageId="0" pageNumber="49">
<emphasis id="012C53CB096A6257E6806BFB57C0ADA6" bold="true" pageId="0" pageNumber="49">Figure 1.</emphasis>
<taxonomicName id="F73A3DF94D1F2F99BB5A858FCD17958F" authorityName="Seyrig" authorityYear="1932" class="Insecta" family="Ichneumonidae" genus="Tetractenion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tetractenion acaule" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="acaule">
<emphasis id="0437DCCE1E6D2D149EAE78924ED78517" italics="true" pageId="0" pageNumber="49">Tetractenion acaule</emphasis>
</taxonomicName>
Lectotype (MNHN)
<emphasis bold="true" pageId="0" pageNumber="49">A</emphasis>
<emphasis id="E14B8453F12F5887AD443997B4E5E917" bold="true" pageId="0" pageNumber="49">A</emphasis>
habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="49">B</emphasis>
<emphasis id="C840491E9CF091A4C2B419F5E9C469F7" bold="true" pageId="0" pageNumber="49">B</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="49">C</emphasis>
<emphasis id="8B9DEC77D19E3A6C9D96E3E7548B2F28" bold="true" pageId="0" pageNumber="49">C</emphasis>
head, anterior view
<emphasis bold="true" pageId="0" pageNumber="49">D</emphasis>
<emphasis id="F5112FF3A29D02A781F681923EAD7487" bold="true" pageId="0" pageNumber="49">D</emphasis>
data labels. Photographs of lectotype RECOLNAT (ANR-11-INBS-0004) - Christophe
<normalizedToken originalValue="Hervé">Herve</normalizedToken>
<normalizedToken id="5F836AD0B56A4462C33A2BED218C8715" originalValue="Hervé">Herve</normalizedToken>
- 2014. http://coldb.mnhn.fr/catalognumber/mnhn/ey/ey9333 (used with permission of
<normalizedToken originalValue="Agnièle">Agniele</normalizedToken>
<normalizedToken id="B18039F85D6EF9E367B947978F1F9135" originalValue="Agnièle">Agniele</normalizedToken>
Touret-Alby - Curator of
<taxonomicName authorityName="Seyrig" authorityYear="1932" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="49" rank="order">Hymenoptera</taxonomicName>
<taxonomicName id="BA0975FFA326B47CE91C0AAC3169509E" authorityName="Seyrig" authorityYear="1932" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
MNHN).
</paragraph>
</caption>

View file

@ -1,248 +1,247 @@
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<mods:title>A revision of the new genus Amiga Nakahara, Willmott &amp; Espeland, gen. n., described for Papilioarnaca Fabricius, 1776 (Lepidoptera, Nymphalidae, Satyrinae)</mods:title>
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<paragraph pageId="5" pageNumber="90">Type species.</paragraph>
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<paragraph id="70F119A904A4F24C4923907AE0E3DAAB" pageId="5" pageNumber="90">Type species.</paragraph>
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Fabricius, 1776, by present designation
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<paragraph pageId="5" pageNumber="90">Systematic placement and diagnosis.</paragraph>
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<bibRefCitation author="Espeland, M" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="21" pageNumber="106" pagination="116 - 124" title="Four hundred shades of brown: higher level phylogeny of the problematic Euptychiina (Lepidoptera, Nymphalidae, Satyrinae) based on hybrid enrichment data." url="https://doi.org/10.1016/j.ympev.2018.10.039" volume="131" year="2019">Espeland et al. (2019)</bibRefCitation>
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<paragraph id="CC8E85D534A7DF7B9817E54893687312" pageId="5" pageNumber="90">Systematic placement and diagnosis.</paragraph>
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<bibRefCitation id="4F2FA040D956A93FF6E8A460434F9CF5" author="Espeland, M" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="21" pageNumber="106" pagination="116 - 124" title="Four hundred shades of brown: higher level phylogeny of the problematic Euptychiina (Lepidoptera, Nymphalidae, Satyrinae) based on hybrid enrichment data." url="https://doi.org/10.1016/j.ympev.2018.10.039" volume="131" year="2019">Espeland et al. (2019)</bibRefCitation>
recovered
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comb. n. as sister to the &quot;
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clade&quot;, whose composition partially corresponded to that found in
<bibRefCitation author="Pena, C" journalOrPublisher="Zoologica Scripta" pageId="22" pageNumber="107" pagination="243 - 258" title="Biogeographic history of the butterfly subtribe Euptychiina (Lepidoptera, Nymphalidae, Satyrinae)." url="https://doi.org/10.1111/j.1463-6409.2010.00421.x" volume="39" year="2010">
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, with a high support (BS and PP&gt; 0.95). The &quot;
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clade&quot; itself was also well supported (BS and PP&gt; 0.95), including
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(Bates, 1865),
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<taxonomicName id="B7CD1E4B7D2262C2E4033FE7F2BE8877" class="Insecta" family="Nymphalidae" genus="Magneuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Magneuptychia inani" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="inani">Magneuptychia inani</taxonomicName>
(Staudinger, [1886]),
<taxonomicName class="Insecta" family="Nymphalidae" genus="Euptychoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euptychoides albofasciata" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="Albofasciata">Euptychoides albofasciata</taxonomicName>
<taxonomicName id="0707DB317258963B8A81A0EC838334CA" class="Insecta" family="Nymphalidae" genus="Euptychoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euptychoides albofasciata" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="albofasciata">Euptychoides albofasciata</taxonomicName>
(Hewitson, 1869),
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<taxonomicName id="8AE2CA3B966FC9815F91B2DD2DC9FC1A" class="Insecta" family="Nymphalidae" genus="Neonympha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neonympha areolatus" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="areolatus">Neonympha areolatus</taxonomicName>
(Smith, 1797),
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<taxonomicName id="86273EE09C73E09E105EB8EAC01661C8" class="Insecta" family="Nymphalidae" genus="Erichthodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erichthodes antonina" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="antonina">Erichthodes antonina</taxonomicName>
(C. Felder &amp; R. Felder, 1867),
<taxonomicName class="Insecta" family="Nymphalidae" genus="Pareuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pareuptychia ocirrhoe" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="Ocirrhoe">Pareuptychia ocirrhoe</taxonomicName>
<taxonomicName id="3753F2C8FAA58DA176762BDCE79ABCB4" class="Insecta" family="Nymphalidae" genus="Pareuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pareuptychia ocirrhoe" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="ocirrhoe">Pareuptychia ocirrhoe</taxonomicName>
(Fabricius, 1776),
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<taxonomicName id="71417082DA16B95CFE449A473C7F2F53" class="Insecta" family="Nymphalidae" genus="Megeuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megeuptychia antonoe" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="antonoe">Megeuptychia antonoe</taxonomicName>
(Cramer, 1775),
<taxonomicName class="Insecta" family="Nymphalidae" genus="Splendeuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Splendeuptychia doxes" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="Doxes">Splendeuptychia doxes</taxonomicName>
<taxonomicName id="76EFC972E69936991F208675650744C3" class="Insecta" family="Nymphalidae" genus="Splendeuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Splendeuptychia doxes" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="doxes">Splendeuptychia doxes</taxonomicName>
(Godart, [1824]),
<taxonomicName class="Insecta" family="Nymphalidae" genus="Nhambikuara" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nhambikuara mima" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="Mima">Nhambikuara mima</taxonomicName>
<taxonomicName id="A152F5ECF6090EBD4BF18107267C61CE" genus="Nhambikuara" lsidName="Nhambikuara mima" pageId="5" pageNumber="90" rank="species" species="mima">Nhambikuara mima</taxonomicName>
(Butler, 1867), and
<taxonomicName class="Insecta" family="Nymphalidae" genus="Euptychoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euptychoides eugenia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="Eugenia">Euptychoides eugenia</taxonomicName>
<taxonomicName id="96407E8ABDDDF6E30369FF407AFD78E3" class="Insecta" family="Nymphalidae" genus="Euptychoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euptychoides eugenia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="eugenia">Euptychoides eugenia</taxonomicName>
(C. Felder &amp; R. Felder, 1867).
<taxonomicName class="Insecta" family="Nymphalidae" genus="Amiga" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amiga" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Amiga</taxonomicName>
<taxonomicName id="17714DB60E1CBC6EE6CDAC100310B002" genus="Amiga" lsidName="Amiga" pageId="5" pageNumber="90" rank="genus">Amiga</taxonomicName>
gen. n. is distinguished from all members of the &quot;
<taxonomicName class="Insecta" family="Nymphalidae" genus="Pareuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pareuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Pareuptychia</taxonomicName>
<taxonomicName id="477900008C277E11DFDC534B9F5C24B2" class="Insecta" family="Nymphalidae" genus="Pareuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pareuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Pareuptychia</taxonomicName>
clade&quot; by the presence of bluish-lilac coloration on the dorsal hindwing and by the purplish sheen in the tornal half of the VHW. Furthermore, the absence of cornuti and membranous lamella antevaginalis of
<taxonomicName class="Insecta" family="Nymphalidae" genus="Amiga" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amiga" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Amiga</taxonomicName>
<taxonomicName id="42E938E0BF8DE67AD00FAF5C0D1C9A09" genus="Amiga" lsidName="Amiga" pageId="5" pageNumber="90" rank="genus">Amiga</taxonomicName>
gen. n. appear to be unusual character states among the clade. The type species of
<taxonomicName class="Insecta" family="Nymphalidae" genus="Chloreuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloreuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Chloreuptychia</taxonomicName>
<taxonomicName id="CB4D8CF00B889615922CC292949E516F" class="Insecta" family="Nymphalidae" genus="Chloreuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloreuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Chloreuptychia</taxonomicName>
,
<taxonomicName class="Insecta" family="Papilionidae" genus="Papilio" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papilio chloris" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="Chloris">Papilio chloris</taxonomicName>
<taxonomicName id="4F3D970F6ECB2E0E3E4EAD0D541DFF04" class="Insecta" family="Papilionidae" genus="Papilio" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papilio chloris" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="chloris">Papilio chloris</taxonomicName>
Cramer, 1780 (=
<taxonomicName class="Insecta" family="Nymphalidae" genus="Chloreuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloreuptychia chlorimene" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="Chlorimene">Chloreuptychia chlorimene</taxonomicName>
<taxonomicName id="CC52F6B59AF783FDB29DDBBEF2EE73D2" class="Insecta" family="Nymphalidae" genus="Chloreuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloreuptychia chlorimene" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="species" species="chlorimene">Chloreuptychia chlorimene</taxonomicName>
) was recovered as sister to a moderately supported (BS and PP&gt; 0. 75 &lt;0. 95), clade including the &quot;
<taxonomicName class="Insecta" family="Nymphalidae" genus="Pareuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pareuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Pareuptychia</taxonomicName>
<taxonomicName id="FBFDBB564448264C3632D1DE16C1DC25" class="Insecta" family="Nymphalidae" genus="Pareuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pareuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Pareuptychia</taxonomicName>
clade&quot;, &quot;
<taxonomicName class="Insecta" family="Nymphalidae" genus="Taygetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taygetis" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Taygetis</taxonomicName>
<taxonomicName id="5D95D1F49D5ABFC0CC8EDD2AA4D766F2" class="Insecta" family="Nymphalidae" genus="Taygetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Taygetis" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Taygetis</taxonomicName>
clade&quot;, &quot;
<taxonomicName class="Insecta" family="Nymphalidae" genus="Splendeuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Splendeuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Splendeuptychia</taxonomicName>
<taxonomicName id="55E8D3513989FE02D9CB33D1EAD6704D" class="Insecta" family="Nymphalidae" genus="Splendeuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Splendeuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Splendeuptychia</taxonomicName>
clade&quot; and &quot;
<taxonomicName class="Insecta" family="Nymphalidae" genus="Archeuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archeuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Archeuptychia</taxonomicName>
<taxonomicName id="93B622F0F951F9F6C02ACE2F33B13613" class="Insecta" family="Nymphalidae" genus="Archeuptychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archeuptychia" order="Lepidoptera" pageId="5" pageNumber="90" phylum="Arthropoda" rank="genus">Archeuptychia</taxonomicName>
clade&quot;.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="92" pageId="5" pageNumber="90" type="description">
<paragraph pageId="5" pageNumber="90">Description.</paragraph>
<paragraph pageId="5" pageNumber="90">MALE: Forewing length: 18.5-22 mm (n = 6)</paragraph>
<paragraph pageId="5" pageNumber="90">Head: Eyes with sparse golden hair-like setae, with white scales along margin; frons dark brownish, covered with creamy brownish scales and hair-like scales towards antennae; first segment of labial palpi similar in width to second segment, similar in length to third segment, adorned with white long hair-like scales and brownish long hair-like scales ventrally, second segment longer than eye depth and covered with white hair-like scales, brownish hair-like scales, and white scales laterally and dorsally, with brownish hair-like scales and brownish scales along edge of distal two-thirds of dorsal surface, ventrally adorned with brownish hair-like scales and whitish hair-like scales longer than segment width, third segment roughly one-third of second segment in length and covered with brownish scales dorsally and ventrally, with white scales laterally; antennae approximately half of forewing length, with ca 34 segments (n = 1), scape about as twice as long as pedicel, white scales on each side of base of flagellomeres, distal 10-12 segments composing club.</paragraph>
<paragraph pageId="5" pageNumber="90">Thorax: Brownish, dorsally scattered with grayish scales and lightly colored long hair-like scales; ventrally scattered with white scales and white long hair-like scales; foreleg (Fig. 5c) whitish, femur, tibia and tarsus similar in length; midleg and hindleg with femur white ventrally, tibia and tarsus grayish dorsally, ocher ventrally tibia and tarsus adorned with spines ventrally, pair of tibial spurs present at distal end of tibia.</paragraph>
<paragraph pageId="5" pageNumber="90">Abdomen (Fig. 3a, b): Eighth tergite sclerotized in a narrow anterior band and slightly broader posterior patch, which appear as two separate plates.</paragraph>
<paragraph pageId="6" pageNumber="91">
<pageBreakToken pageId="6" pageNumber="91" start="start">Wing</pageBreakToken>
<subSubSection id="52127A50264DA254614B12EB3CAA8AAB" lastPageId="7" lastPageNumber="92" pageId="5" pageNumber="90" type="description">
<paragraph id="9D6BCF1C2DBD51C3E70B3020D0D49997" pageId="5" pageNumber="90">Description.</paragraph>
<paragraph id="F56CB5FE96E2006292766D4764403703" pageId="5" pageNumber="90">MALE: Forewing length: 18.5-22 mm (n = 6)</paragraph>
<paragraph id="15A1773B7B45C776E25A8CFFBAC3BB97" pageId="5" pageNumber="90">Head: Eyes with sparse golden hair-like setae, with white scales along margin; frons dark brownish, covered with creamy brownish scales and hair-like scales towards antennae; first segment of labial palpi similar in width to second segment, similar in length to third segment, adorned with white long hair-like scales and brownish long hair-like scales ventrally, second segment longer than eye depth and covered with white hair-like scales, brownish hair-like scales, and white scales laterally and dorsally, with brownish hair-like scales and brownish scales along edge of distal two-thirds of dorsal surface, ventrally adorned with brownish hair-like scales and whitish hair-like scales longer than segment width, third segment roughly one-third of second segment in length and covered with brownish scales dorsally and ventrally, with white scales laterally; antennae approximately half of forewing length, with ca 34 segments (n = 1), scape about as twice as long as pedicel, white scales on each side of base of flagellomeres, distal 10-12 segments composing club.</paragraph>
<paragraph id="F59F37B3EEE661AF7DE51882961AE187" pageId="5" pageNumber="90">Thorax: Brownish, dorsally scattered with grayish scales and lightly colored long hair-like scales; ventrally scattered with white scales and white long hair-like scales; foreleg (Fig. 5c) whitish, femur, tibia and tarsus similar in length; midleg and hindleg with femur white ventrally, tibia and tarsus grayish dorsally, ocher ventrally tibia and tarsus adorned with spines ventrally, pair of tibial spurs present at distal end of tibia.</paragraph>
<paragraph id="0F93251446D831F3479B2DE4D69C4414" pageId="5" pageNumber="90">Abdomen (Fig. 3a, b): Eighth tergite sclerotized in a narrow anterior band and slightly broader posterior patch, which appear as two separate plates.</paragraph>
<paragraph id="94F55D6C7D50E6CC7110A02D73EAFA1C" pageId="6" pageNumber="91">
<pageBreakToken id="867A0F00F8280762244F863B328438BE" pageId="6" pageNumber="91" start="start">Wing</pageBreakToken>
venation (Fig. 3): Basal half of forewing subcostal vein swollen; base of cubitus swollen; forewing recurrent vein present as small projection slightly above origin of M2; hindwing humeral vein developed; origin of M2 slightly towards M1 than M3.
</paragraph>
<paragraph pageId="6" pageNumber="91">Wing shape: Forewing triangular, apex rounded, costal margin slightly convex, outer margin almost straight below M2, inner margin almost straight, but curving inwards towards thorax near base; hindwing somewhat rectangular, slightly elongate, costal margin almost straight, angled inwards at base, outer margin slightly undulating, inner margin slightly curved near tornus, anal lobe convex, slightly rounded.</paragraph>
<paragraph pageId="6" pageNumber="91">Dorsal forewing: Ground color light brownish, slightly translucent, thus subtly revealing ventral bands and ocelli in cell M1.</paragraph>
<paragraph pageId="6" pageNumber="91">Dorsal hindwing: Ground color similar to forewing, iridescent bluish lilac reflection covering most of dorsal hindwing, area near costa and area distal to marginal band revealing ground color; slightly translucent, thus subtly revealing ventral bands and ocelli.</paragraph>
<paragraph pageId="6" pageNumber="91">Ventral forewing: Ground color pale grayish brown; pale reddish-brown discal band extending from radial vein, crossing discal cell, passing origin of Cu2, terminating at 2A; concolorous scales present along discocellular vein; pale reddish-brown postdiscal band extending from radial vein towards inner margin until reaching vein 2A, slightly wider than discal band; broad, faint, iridescent bluish-lilac reflection extending just distal of postdiscal band towards outer margin, prominent between postdiscal band and umbra; sinuate, narrow submarginal band, almost concolorous to basal two bands, extending from apex towards tornus, jagged above Cu1, almost straight below this vein; concolorous marginal band, not jagged, appearing narrower than submarginal band, traversing along marginal area from apex to tornus; fringe brownish; ocellus in cell M1, spilling out from veins M1 and M2, black with two whitish pupils in center, ringed in yellowish orange, ocelli present in cell M2 and M3, appearing as two slate-gray patches surrounded by yellowish-orange ring; umbra appearing as broad rather faint band concolorous to four ventral bands, visible around ocelli, extending to cell Cu2.</paragraph>
<paragraph pageId="6" pageNumber="91">Ventral hindwing: Ground color similar to forewing; general wing pattern similar to forewing except as follows: iridescent bluish-lilac reflection extending from base of wing, towards outer and inner margin, especially area where iridescent bluish scales are present on dorsal surface; discal band passing cubital vein area basal to origin of Cu2; postdiscal band passing origin of Cu1, bent inwards in cell Cu2; submarginal band broadening towards tornus after passing Cu1; five submarginal ocelli, those in cells M1 and Cu1 similar to that in VFW cell M1 but with single pupil, those in cells M2 and M3 similar to those in VFW cells Rs, M2 and M3 but slate grayish patch appearing as single patch in middle.</paragraph>
<paragraph lastPageId="7" lastPageNumber="92" pageId="6" pageNumber="91">
<paragraph id="E4368D8636D9F70897F595863B602B18" pageId="6" pageNumber="91">Wing shape: Forewing triangular, apex rounded, costal margin slightly convex, outer margin almost straight below M2, inner margin almost straight, but curving inwards towards thorax near base; hindwing somewhat rectangular, slightly elongate, costal margin almost straight, angled inwards at base, outer margin slightly undulating, inner margin slightly curved near tornus, anal lobe convex, slightly rounded.</paragraph>
<paragraph id="6AEF59A62CE43C63A303E5ABFA6D6C13" pageId="6" pageNumber="91">Dorsal forewing: Ground color light brownish, slightly translucent, thus subtly revealing ventral bands and ocelli in cell M1.</paragraph>
<paragraph id="93AD57D84DB119242561F9D4E90DCF42" pageId="6" pageNumber="91">Dorsal hindwing: Ground color similar to forewing, iridescent bluish lilac reflection covering most of dorsal hindwing, area near costa and area distal to marginal band revealing ground color; slightly translucent, thus subtly revealing ventral bands and ocelli.</paragraph>
<paragraph id="CA74B3928EE2BF4DDA182CF0298365DB" pageId="6" pageNumber="91">Ventral forewing: Ground color pale grayish brown; pale reddish-brown discal band extending from radial vein, crossing discal cell, passing origin of Cu2, terminating at 2A; concolorous scales present along discocellular vein; pale reddish-brown postdiscal band extending from radial vein towards inner margin until reaching vein 2A, slightly wider than discal band; broad, faint, iridescent bluish-lilac reflection extending just distal of postdiscal band towards outer margin, prominent between postdiscal band and umbra; sinuate, narrow submarginal band, almost concolorous to basal two bands, extending from apex towards tornus, jagged above Cu1, almost straight below this vein; concolorous marginal band, not jagged, appearing narrower than submarginal band, traversing along marginal area from apex to tornus; fringe brownish; ocellus in cell M1, spilling out from veins M1 and M2, black with two whitish pupils in center, ringed in yellowish orange, ocelli present in cell M2 and M3, appearing as two slate-gray patches surrounded by yellowish-orange ring; umbra appearing as broad rather faint band concolorous to four ventral bands, visible around ocelli, extending to cell Cu2.</paragraph>
<paragraph id="11382BCBE61EB116683CDDBF1852ECC2" pageId="6" pageNumber="91">Ventral hindwing: Ground color similar to forewing; general wing pattern similar to forewing except as follows: iridescent bluish-lilac reflection extending from base of wing, towards outer and inner margin, especially area where iridescent bluish scales are present on dorsal surface; discal band passing cubital vein area basal to origin of Cu2; postdiscal band passing origin of Cu1, bent inwards in cell Cu2; submarginal band broadening towards tornus after passing Cu1; five submarginal ocelli, those in cells M1 and Cu1 similar to that in VFW cell M1 but with single pupil, those in cells M2 and M3 similar to those in VFW cells Rs, M2 and M3 but slate grayish patch appearing as single patch in middle.</paragraph>
<paragraph id="8403C5D107481860C790F7B4389410A4" lastPageId="7" lastPageNumber="92" pageId="6" pageNumber="91">
Genitalia (Fig. 4
<normalizedToken originalValue="cg">c-g</normalizedToken>
<normalizedToken id="16AD12AF5F8C1F2C12D9C208A42D510C" originalValue="cg">c-g</normalizedToken>
): Tegumen appears semi-circular in lateral view, anteriorly and dorsally convex, ventral margin rather straight; uncus longer than tegumen in lateral view, apparently without setae, middle section somewhat broadening in dorsal view, tapering posteriorly and terminating in single point; brachium tapering towards apex, apical point positioned above uncus in lateral view, parallel to uncus with apical edge curving inwards in dorsal view; combination of ventral arms from tegumen and
<pageBreakToken pageId="7" pageNumber="92" start="start">dorsal</pageBreakToken>
<pageBreakToken id="734E2374682F22BC4DD4B5AA20443527" pageId="7" pageNumber="92" start="start">dorsal</pageBreakToken>
arms from saccus slightly curved distally; appendices angulares present; saccus narrow, concavity at base of ventral margin, anteriorly rounded, similar or shorter than uncus in length; juxta present as plate with deep concavity at dorsal margin in posterior view; valva distally setose, valva appearing roughly parallelogram in lateral view, ventral margin convex, dorsal margin distal of costa curved, costa curved inwards, apical process somewhat curving upwards; phallus roughly straight, similar in length to valva plus saccus, phallobase about one-third of phallus, ductus ejaculatorius visible as illustrated, posterior portion of aedeagus somewhat curved upwards, manica covering more than half of aedeagus, cornuti absent.
</paragraph>
<paragraph pageId="7" pageNumber="92">FEMALE: forewing length: 19-21 mm (n = 6)</paragraph>
<paragraph pageId="7" pageNumber="92">
<paragraph id="0992F988CEB5AF07F9D6BB6B497BAC36" pageId="7" pageNumber="92">FEMALE: forewing length: 19-21 mm (n = 6)</paragraph>
<paragraph id="B10963B2EE60D999D124E30FED9F8A57" pageId="7" pageNumber="92">
Similar to male except as follows: Five tarsomeres present in foretarsus, with spines along some tarsomeres; forewing appearing somewhat rounded and broad; dorsal hindwing submarginal band somewhat more prominent; bluish lilac reflection appearing more purplish, extending to origin of M1 or further anteriorly; feeble pearly reflection present on dorsal forewing (but see below for further information). Female genitalia and abdomen (Figs 4
<normalizedToken originalValue="hk">h-k</normalizedToken>
<normalizedToken id="6C020A78196DE70A736A2348C4592BAF" originalValue="hk">h-k</normalizedToken>
): Inter-segmental membrane between 7th and 8th abdominal segments pleated and expandable, with weakly sclerotized region present; lamella antevaginalis membranous; lateral side of 8th abdominal segment sclerotized (referred to as &quot;lamella postvaginalis&quot; by Willmott et al. 2018), this sclerotized plate fused to lamella antevaginalis at anterior margin; ductus bursae membranous, somewhat inflated around origin of ductus seminalis,
<taxonomicName class="Insecta" family="Nymphalidae" genus="Euptychiina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Euptychiina located" order="Lepidoptera" pageId="7" pageNumber="92" phylum="Arthropoda" rank="species" species="Located">located</taxonomicName>
<taxonomicName id="AEA31293DA6D340B65885B2A4F8B8516" genus="Euptychiina" lsidName="Euptychiina located" pageId="7" pageNumber="92" rank="species" species="located">located</taxonomicName>
at approximately one-fifth distance from ostium bursae to corpus bursae, ductus bursae apparently weakly sclerotized at region posterior to origin of ductus seminalis; corpus bursae roughly oval in dorsal view, with two relatively narrow signa, together with ductus bursae extending to juncture of 4th and 5th sternite.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="93" pageId="7" pageNumber="92" type="variation">
<paragraph pageId="7" pageNumber="92">Variation.</paragraph>
<paragraph lastPageId="8" lastPageNumber="93" pageId="7" pageNumber="92">
<subSubSection id="03CA1F1B2683B0D3E841B084B4F4F0F5" lastPageId="8" lastPageNumber="93" pageId="7" pageNumber="92" type="variation">
<paragraph id="6A4BDADF503F5C93F7FE10B1BE149553" pageId="7" pageNumber="92">Variation.</paragraph>
<paragraph id="BF353C2F5D02BB9AF8668961FD361034" lastPageId="8" lastPageNumber="93" pageId="7" pageNumber="92">
This species exhibits geographic variation in wing pattern, some of which is recognized here with subspecific names. A broad, faint, iridescent bluish-lilac reflection between the ventral forewing postdiscal band and the umbra is present in specimens from the eastern Andes, whereas it is absent in many specimens from west of the Andes. The absence of this bluish-lilac reflection on the ventral forewing seems rather stable in specimens from western Colombia and western Ecuador. However, this character appears in a few specimens from Panama, and is present in some specimens from Costa Rica and Nicaragua, although the degree of reflection is variable. The bluish-lilac reflection seen mainly on both the dorsal forewing and hindwing is variable in color, varying from light blue to purple. Especially in female specimens, the extent of bluish-lilac reflection on the dorsal forewing is variable, being absent in some specimens, whereas covering most of the discal cell and cells Cu1 and Cu2 in others (see below for further information). There exists a feeble pearly reflection on the ventral forewing from the base to the postdiscal band in specimens from southeastern Brazil (see below for further information), although this is absent in a few specimens. The presence or absence of a ventral forewing ocellus in cell M3 is variable, appearing as a trace in some specimens. The size of the ventral hindwing ocellus in cell M1 is variable in comparison with the ventral hindwing ocellus in cell Cu1, ranging from similar in
<pageBreakToken pageId="8" pageNumber="93" start="start">size</pageBreakToken>
<pageBreakToken id="65961855570BF175B5802F768603F37E" pageId="8" pageNumber="93" start="start">size</pageBreakToken>
to almost twice as large. The genitalia appear not to be informative in separating specimens from east and west of the Andes, although specimens from Central America, including
<taxonomicName lsidName="A. arnaca" pageId="8" pageNumber="93" rank="subSpecies" species="Arnaca" subSpecies="sericeella">A. arnaca sericeella</taxonomicName>
<taxonomicName id="B23EAF635A1550AB01EB9ADBE480D458" lsidName="A. arnaca" pageId="8" pageNumber="93" rank="subSpecies" species="arnaca" subSpecies="sericeella">A. arnaca sericeella</taxonomicName>
comb. n. et stat. n., seem to have a rather curved dorsal margin of the uncus in lateral view (Fig. 4e).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="93" type="etymology">
<paragraph pageId="8" pageNumber="93">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="93">
<subSubSection id="2881659B3754A37C647A2418C3FB3101" pageId="8" pageNumber="93" type="etymology">
<paragraph id="E668B7474D8AF8D3BACAD417F5ABB961" pageId="8" pageNumber="93">Etymology.</paragraph>
<paragraph id="2E088ABA85FF01291A5E79AC40D2C297" pageId="8" pageNumber="93">
The new generic name is derived from the feminine Spanish noun
<normalizedToken originalValue="“amiga”">&quot;amiga&quot;</normalizedToken>
<normalizedToken id="CF5444F013F67CD14735C80840AA3CD5" originalValue="“amiga”">&quot;amiga&quot;</normalizedToken>
, meaning &quot;a (female) friend&quot;, alluding to the fact that this is a common, familiar butterfly. The generic name is regarded as feminine.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="94" pageId="8" pageNumber="93" type="biology_ecology">
<paragraph pageId="8" pageNumber="93">Biology.</paragraph>
<paragraph lastPageId="9" lastPageNumber="94" pageId="8" pageNumber="93">
<subSubSection id="D098406C803374FA7F911DF6FE1D163D" lastPageId="9" lastPageNumber="94" pageId="8" pageNumber="93" type="biology_ecology">
<paragraph id="B5D650608B8F5E6BF7EC5C300FCFDB2B" pageId="8" pageNumber="93">Biology.</paragraph>
<paragraph id="6D03DB3EE43676BA5323D44F042D2311" lastPageId="9" lastPageNumber="94" pageId="8" pageNumber="93">
Janzen and Hallwachs (2018) report four grass species (
<taxonomicName family="Poaceae" lsidName="Euptychiina" pageId="8" pageNumber="93" rank="family">Poaceae</taxonomicName>
<taxonomicName id="431CE5B11453FF404BC4BE75F6E45226" class="Liliopsida" family="Poaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Euptychiina" order="Poales" pageId="8" pageNumber="93" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
),
<taxonomicName class="Liliopsida" family="Poaceae" genus="Ichnanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ichnanthus nemorosus" order="Poales" pageId="8" pageNumber="93" phylum="Tracheophyta" rank="species" species="Nemorosus">Ichnanthus nemorosus</taxonomicName>
<taxonomicName id="466E34EF58B62E11B7D7449CF22F9FF1" class="Liliopsida" family="Poaceae" genus="Ichnanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ichnanthus nemorosus" order="Poales" pageId="8" pageNumber="93" phylum="Tracheophyta" rank="species" species="nemorosus">Ichnanthus nemorosus</taxonomicName>
,
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<taxonomicName id="6F14692A5B2B95A323FAE91E953D3B55" class="Liliopsida" family="Poaceae" genus="Ichnanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ichnanthus pallens" order="Poales" pageId="8" pageNumber="93" phylum="Tracheophyta" rank="species" species="pallens">Ichnanthus pallens</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Poaceae" genus="Lasiacis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lasiacis ruscifolia" order="Poales" pageId="8" pageNumber="93" phylum="Tracheophyta" rank="species" species="Ruscifolia">Lasiacis ruscifolia</taxonomicName>
<taxonomicName id="D98257215F09EA88F0E448EBA9273F80" class="Liliopsida" family="Poaceae" genus="Lasiacis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Lasiacis ruscifolia" order="Poales" pageId="8" pageNumber="93" phylum="Tracheophyta" rank="species" species="ruscifolia">Lasiacis ruscifolia</taxonomicName>
, and
<taxonomicName class="Liliopsida" family="Poaceae" genus="Paspalum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paspalum decumbens" order="Poales" pageId="8" pageNumber="93" phylum="Tracheophyta" rank="species" species="Decumbens">Paspalum decumbens</taxonomicName>
<taxonomicName id="81B894DC1EB27BFCE84EF31C6C02A2B7" class="Liliopsida" family="Poaceae" genus="Paspalum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Paspalum decumbens" order="Poales" pageId="8" pageNumber="93" phylum="Tracheophyta" rank="species" species="decumbens">Paspalum decumbens</taxonomicName>
,
<pageBreakToken pageId="9" pageNumber="94" start="start">as</pageBreakToken>
<pageBreakToken id="71F8496A1CEC7F5FFE67C2041BC61624" pageId="9" pageNumber="94" start="start">as</pageBreakToken>
hostplants for
<taxonomicName class="Insecta" family="Nymphalidae" genus="Amiga" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amiga" order="Lepidoptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="genus">Amiga</taxonomicName>
<taxonomicName id="8A0E245ABB8139CB7FEF213F384114B9" genus="Amiga" lsidName="Amiga" pageId="9" pageNumber="94" rank="genus">Amiga</taxonomicName>
gen. n. in Costa Rica. In addition,
<bibRefCitation author="DeVries, PJ" journalOrPublisher="Princeton University Press, Princeton" pageId="21" pageNumber="106" title="The butterflies of Costa Rica and their natural history. Papilionidae, Pieridae, Nymphalidae." year="1987">DeVries (1987)</bibRefCitation>
<bibRefCitation id="07A2174AD3B2439FBA838115F7D26E99" author="DeVries, PJ" journalOrPublisher="Princeton University Press, Princeton" pageId="21" pageNumber="106" title="The butterflies of Costa Rica and their natural history. Papilionidae, Pieridae, Nymphalidae." year="1987">DeVries (1987)</bibRefCitation>
reported three grass genera, namely
<taxonomicName class="Liliopsida" family="Poaceae" genus="Eleusine" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eleusine" order="Poales" pageId="9" pageNumber="94" phylum="Tracheophyta" rank="genus">Eleusine</taxonomicName>
<taxonomicName id="D7F99FDB45ADA2C0CFA1631611FE9F66" class="Liliopsida" family="Poaceae" genus="Eleusine" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eleusine" order="Poales" pageId="9" pageNumber="94" phylum="Tracheophyta" rank="genus">Eleusine</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Poaceae" genus="Ichnanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ichnanthus" order="Poales" pageId="9" pageNumber="94" phylum="Tracheophyta" rank="genus">Ichnanthus</taxonomicName>
<taxonomicName id="BF2636074750D31D7558F4745D325657" class="Liliopsida" family="Poaceae" genus="Ichnanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ichnanthus" order="Poales" pageId="9" pageNumber="94" phylum="Tracheophyta" rank="genus">Ichnanthus</taxonomicName>
, and
<taxonomicName class="Liliopsida" family="Poaceae" genus="Oplismenus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oplismenus" order="Poales" pageId="9" pageNumber="94" phylum="Tracheophyta" rank="genus">Oplismenus</taxonomicName>
<taxonomicName id="DBF88F7BCE5F277360DE6CB4E06D3F7D" class="Liliopsida" family="Poaceae" genus="Oplismenus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Oplismenus" order="Poales" pageId="9" pageNumber="94" phylum="Tracheophyta" rank="genus">Oplismenus</taxonomicName>
, as hostplants in Costa Rica.
<bibRefCitation pageId="9" pageNumber="94">Singer and Ehrlich (1993)</bibRefCitation>
<bibRefCitation id="41D62C750D6643BA168DDC728739BBCD" pageId="9" pageNumber="94">Singer and Ehrlich (1993)</bibRefCitation>
reported
<taxonomicName class="Liliopsida" family="Poaceae" genus="Ichnanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ichnanthus pallens" order="Poales" pageId="9" pageNumber="94" phylum="Tracheophyta" rank="species" species="Pallens">Ichnanthus pallens</taxonomicName>
<taxonomicName id="E8D36A812DE5C099857485EB9928772D" class="Liliopsida" family="Poaceae" genus="Ichnanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ichnanthus pallens" order="Poales" pageId="9" pageNumber="94" phylum="Tracheophyta" rank="species" species="pallens">Ichnanthus pallens</taxonomicName>
as a hostplant in Trinidad and Tobago. The egg, mature larva, and pupa of
<taxonomicName class="Insecta" family="Nymphalidae" genus="Amiga" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amiga" order="Lepidoptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="genus">Amiga</taxonomicName>
<taxonomicName id="68254849F525EBED5B622E4FF5EAE6CD" genus="Amiga" lsidName="Amiga" pageId="9" pageNumber="94" rank="genus">Amiga</taxonomicName>
gen. n. were described and the latter two stages illustrated in
<bibRefCitation author="DeVries, PJ" journalOrPublisher="Princeton University Press, Princeton" pageId="21" pageNumber="106" title="The butterflies of Costa Rica and their natural history. Papilionidae, Pieridae, Nymphalidae." year="1987">DeVries (1987)</bibRefCitation>
<bibRefCitation id="53D36CB8190C3564A53CD5A74D5B01EC" author="DeVries, PJ" journalOrPublisher="Princeton University Press, Princeton" pageId="21" pageNumber="106" title="The butterflies of Costa Rica and their natural history. Papilionidae, Pieridae, Nymphalidae." year="1987">DeVries (1987)</bibRefCitation>
. Various images of the penultimate instar, ultimate instar, prepupa and pupa are figured by Janzen and Hallwachs (2018), based on material reared in Costa Rica. In Costa Rica, adult females of
<taxonomicName class="Insecta" family="Nymphalidae" genus="Amiga" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Amiga" order="Lepidoptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="genus">Amiga</taxonomicName>
<taxonomicName id="8B7933B05B38682117729DAF9D0A0C6E" genus="Amiga" lsidName="Amiga" pageId="9" pageNumber="94" rank="genus">Amiga</taxonomicName>
gen. n. were seen ovipositing late in the afternoon, and some eggs were observed to be parasitized by trichogrammatid wasps (
<bibRefCitation author="DeVries, PJ" journalOrPublisher="Princeton University Press, Princeton" pageId="21" pageNumber="106" title="The butterflies of Costa Rica and their natural history. Papilionidae, Pieridae, Nymphalidae." year="1987">DeVries 1987</bibRefCitation>
<bibRefCitation id="717A5A6DD72E373F8F9EB27AD684DF6B" author="DeVries, PJ" journalOrPublisher="Princeton University Press, Princeton" pageId="21" pageNumber="106" title="The butterflies of Costa Rica and their natural history. Papilionidae, Pieridae, Nymphalidae." year="1987">DeVries 1987</bibRefCitation>
). The species occurs from sea level to at least 1850 m, and it is common, indeed ubiquitous, in undisturbed to heavily disturbed rain and cloud forest. Both sexes fly low (0.5 m) along shady trails throughout the middle of the day (09:00-15:00), and males are often observed perching singly on tops of leaves, maintaining apparent territories, and patrolling for ca 10 m along a trail.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="94" type="distribution">
<paragraph pageId="9" pageNumber="94">Distribution</paragraph>
<paragraph pageId="9" pageNumber="94">(Fig. 6). This genus ranges from southern Mexico throughout virtually all of tropical Central and South America, where its southernmost distribution appears to be southern Brazil.</paragraph>
<subSubSection id="22176844447A67B2C5652F00FF2A2370" pageId="9" pageNumber="94" type="distribution">
<paragraph id="479CEEE360E094624732F25992600F66" pageId="9" pageNumber="94">Distribution</paragraph>
<paragraph id="635D849CC3260EAB6F99F73707BAA69F" pageId="9" pageNumber="94">(Fig. 6). This genus ranges from southern Mexico throughout virtually all of tropical Central and South America, where its southernmost distribution appears to be southern Brazil.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,241 +1,240 @@
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<taxonomicName authority="Lee, 1974" authorityName="Lee" authorityYear="1974" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella liguladorsi" order="Collembola" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="liguladorsi">Ceratophysella liguladorsi (Lee, 1974)</taxonomicName>
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<taxonomicName class="Collembola" family="Hypogastruridae" genus="Hypogastrura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypogastrura liguladorsi" order="Collembola" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="liguladorsi">Hypogastrura liguladorsi</taxonomicName>
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Lee, 1974: 95.
</paragraph>
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<paragraph pageId="5" pageNumber="72">Studied material.</paragraph>
<paragraph pageId="5" pageNumber="72">
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<paragraph id="D8CC5ACA7085EA5FBAA25814EA6D9493" pageId="5" pageNumber="72">Studied material.</paragraph>
<paragraph id="4C14EFB5E52F77D811829FE94DFB933D" pageId="5" pageNumber="72">
China: Zhejiang: Wuyi, on
<taxonomicName class="Agaricomycetes" family="Polyporaceae" genus="Lentinus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Lentinus" order="Polyporales" pageId="5" pageNumber="72" phylum="Basidiomycota" rank="genus">Lentinus</taxonomicName>
<taxonomicName id="0E638B121BCED2B151DBDE1212B9B054" class="Agaricomycetes" family="Polyporaceae" genus="Lentinus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Lentinus" order="Polyporales" pageId="5" pageNumber="72" phylum="Basidiomycota" rank="genus">Lentinus</taxonomicName>
sp., 11 specimens on slides and 80 in alcohol, leg. Xin Sun, 09 Dec 2014.
</paragraph>
<subSection lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73" type="key to the chinese species of ceratophysella">
<paragraph pageId="6" pageNumber="73">
<pageBreakToken pageId="6" pageNumber="73" start="start">Key</pageBreakToken>
<subSection id="38A4B8AA1F8E381F77D682F9BF1301B2" lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73" type="key to the chinese species of ceratophysella">
<paragraph id="6A39FEE4C99DF0A196EBD952F1FAA815" pageId="6" pageNumber="73">
<pageBreakToken id="14F408ABD7397B702FAD07E2FE226CFE" pageId="6" pageNumber="73" start="start">Key</pageBreakToken>
to the Chinese species of
<taxonomicName class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella" order="Poduromorpha" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Ceratophysella</taxonomicName>
<taxonomicName id="1F8673C70CB266514F43AAC0F64BF20B" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella" order="Poduromorpha" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Ceratophysella</taxonomicName>
</paragraph>
<paragraph pageId="6" pageNumber="73">
<paragraph id="AE2B4B1BCC756D7AF89E811DFD3E7F3F" pageId="6" pageNumber="73">
In 2007 Wu and Yin proposed a key to the six species of the genus
<taxonomicName class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella" order="Poduromorpha" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Ceratophysella</taxonomicName>
<taxonomicName id="90F69349DE38DE30BE55C1AED34250A5" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella" order="Poduromorpha" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Ceratophysella</taxonomicName>
known from China.
<bibRefCitation author="Jia, J" journalOrPublisher="Zootaxa" pageId="8" pageNumber="75" pagination="57 - 63" title="Ceratophysellataiguensis sp. nov. (Collembola, Hypogastruridae) from China, with an annotated checklist of Chinese Ceratophysella Boerner, 1932." url="https://doi.org/10.11646/zootaxa.2644.1.4" volume="2644" year="2010">Jia et al. (2010)</bibRefCitation>
<bibRefCitation id="4FAD3FDF4B13B6329AADC8EA9524FF15" author="Jia, J" journalOrPublisher="Zootaxa" pageId="8" pageNumber="75" pagination="57 - 63" title="Ceratophysellataiguensis sp. nov. (Collembola, Hypogastruridae) from China, with an annotated checklist of Chinese Ceratophysella Boerner, 1932." url="https://doi.org/10.11646/zootaxa.2644.1.4" volume="2644" year="2010">Jia et al. (2010)</bibRefCitation>
proposed a list of 14 Chinese species of
<taxonomicName class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella" order="Poduromorpha" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Ceratophysella</taxonomicName>
<taxonomicName id="1972CDEFE037CA125C439167F0E293D4" class="Collembola" family="Hypogastruridae" genus="Ceratophysella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceratophysella" order="Poduromorpha" pageId="6" pageNumber="73" phylum="Arthropoda" rank="genus">Ceratophysella</taxonomicName>
. The present key includes 15 species. Some of the species are not sufficiently described, but the available characters given in the descriptions are sufficient to include them in the key. The characters for
<taxonomicName lsidName="C. adexilis" pageId="6" pageNumber="73" rank="species" species="adexilis">C. adexilis</taxonomicName>
<taxonomicName id="0B73DED4E8AEAC106B7CB3774ADC61DB" lsidName="C. adexilis" pageId="6" pageNumber="73" rank="species" species="adexilis">C. adexilis</taxonomicName>
have been verified on type material, and for
<taxonomicName lsidName="C. communis" pageId="6" pageNumber="73" rank="species" species="communis">C. communis</taxonomicName>
<taxonomicName id="5F48F5EA9FA8758FA8EE2280EF284041" lsidName="C. communis" pageId="6" pageNumber="73" rank="species" species="communis">C. communis</taxonomicName>
on fresh material from the type locality (Tokyo).
</paragraph>
<paragraph lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73">
<table lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73">
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Abd</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">2</td>
<paragraph id="D068D4FE08319D91578923B6B93E470E" lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73">
<table id="E31E3B58E73DE371976C5BB039D7F6BC" lastPageId="7" lastPageNumber="74" pageId="6" pageNumber="73">
<tr id="E82292F26F9B204110761ACB3646BADE" pageId="6" pageNumber="73">
<td id="57251D5B54E60DF15DBFBB763B75EF08" colspan="1" pageId="6" pageNumber="73" rowspan="1">Abd</td>
<td id="81EDAF267F2DAC5B6E5BDFAF3092A7BF" colspan="1" pageId="6" pageNumber="73" rowspan="1">2</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Abd</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">9</td>
<tr id="989BCAF04509BF58A604B63964BB3DC9" pageId="6" pageNumber="73">
<td id="28E701EB1895C8D6468D4C8E279BEAB4" colspan="1" pageId="6" pageNumber="73" rowspan="1">Abd</td>
<td id="9DD74B1AEDFD5F7F358F5DAC9B88459B" colspan="1" pageId="6" pageNumber="73" rowspan="1">9</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">3</td>
<tr id="509EF0EB9272ACE5C5CCADBA9298C606" pageId="6" pageNumber="73">
<td id="DFBD5221F8A1204FD943CF741BB0F9DF" colspan="1" pageId="6" pageNumber="73" rowspan="1">3</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">5</td>
<tr id="C14DF316BF8FA312F0F6707DA47A5B6F" pageId="6" pageNumber="73">
<td id="45849C4778F5DB84FF668977D55BF481" colspan="1" pageId="6" pageNumber="73" rowspan="1">5</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">4</td>
<tr id="7E6A8342740A4B7A12C80EF58C583B8C" pageId="6" pageNumber="73">
<td id="C1C2BADD11C157672BE3003F9BEDD7F3" colspan="1" pageId="6" pageNumber="73" rowspan="1">4</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. zhangi" pageId="6" pageNumber="73" rank="species" species="zhangi">C. zhangi</taxonomicName>
<tr id="BA2B67C4E77F8304D6AEF0EB32DB3A5F" pageId="6" pageNumber="73">
<td id="3C7276603540708D59986A4C11ED3625" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="5616CF0874346F9C908CC1B0F4A808C6" lsidName="C. zhangi" pageId="6" pageNumber="73" rank="species" species="zhangi">C. zhangi</taxonomicName>
(Zhao, 1998)1
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Th</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. succinea" pageId="6" pageNumber="73" rank="species" species="succinea">C. succinea</taxonomicName>
<tr id="2F645993418499AFB9A0F33B549864DD" pageId="6" pageNumber="73">
<td id="7F396699F5B5B92169D064A2CA0A23E8" colspan="1" pageId="6" pageNumber="73" rowspan="1">Th</td>
<td id="B302BEFEF7F8CFC496065529134A6B87" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="76D6A3A11ACE430C776D49BA3E874DF3" lsidName="C. succinea" pageId="6" pageNumber="73" rank="species" species="succinea">C. succinea</taxonomicName>
(Gisin, 1949)
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Th</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. taiguensis" pageId="6" pageNumber="73" rank="species" species="taiguensis">C. taiguensis</taxonomicName>
<tr id="B7955374FC90C88A6F9DCBEAABA577DF" pageId="6" pageNumber="73">
<td id="B63F19DBE08A2FCC63A0F1AC01FCF0C8" colspan="1" pageId="6" pageNumber="73" rowspan="1">Th</td>
<td id="DC6434F980057EA46C5F13210CE6AE06" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="7BCAE597A3DAE0DC3E879B972BEAE4E7" lsidName="C. taiguensis" pageId="6" pageNumber="73" rank="species" species="taiguensis">C. taiguensis</taxonomicName>
Jia,
<normalizedToken originalValue="Skarżyński">Skarzynski</normalizedToken>
<normalizedToken id="7488C682E3CB4B98A158CCD5D6D6AB37" originalValue="Skarżyński">Skarzynski</normalizedToken>
&amp; Li, 2010
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">6</td>
<tr id="23B7C0413ACE90788111EB766446A607" pageId="6" pageNumber="73">
<td id="39B19E8531D609A92FFB145863059D1F" colspan="1" pageId="6" pageNumber="73" rowspan="1">6</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">8</td>
<tr id="A84610125C0AC5009641A3A4EF011946" pageId="6" pageNumber="73">
<td id="BDDB7FE2838135F46E27A0B901A9CE24" colspan="1" pageId="6" pageNumber="73" rowspan="1">8</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">7</td>
<tr id="2418214AA12370E8920BE51E9FB624BB" pageId="6" pageNumber="73">
<td id="607F8CBFEE571B8A7B72DD4C16EE4673" colspan="1" pageId="6" pageNumber="73" rowspan="1">7</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. yinae" pageId="6" pageNumber="73" rank="species" species="yinae">C. yinae</taxonomicName>
<tr id="E7365BF3F81F7DD1E6CD5F5577E0473D" pageId="6" pageNumber="73">
<td id="B5054243F8AD008530F3B45D30905BCD" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="A5885E7AFC6AA65F434E31E8ED1948DB" lsidName="C. yinae" pageId="6" pageNumber="73" rank="species" species="yinae">C. yinae</taxonomicName>
(Yue &amp; Fu, 2000)
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Ant</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. baichengensis" pageId="6" pageNumber="73" rank="species" species="baichengensis">C. baichengensis</taxonomicName>
<tr id="45B44411266AF706B57B50C01E09360E" pageId="6" pageNumber="73">
<td id="16088C6754BAEF52677FAEBAA42BEC8D" colspan="1" pageId="6" pageNumber="73" rowspan="1">Ant</td>
<td id="36AE50A5BD41175E528929CCBAC03C46" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="0FC079825AAD3388186D315C73060D99" lsidName="C. baichengensis" pageId="6" pageNumber="73" rank="species" species="baichengensis">C. baichengensis</taxonomicName>
Wu &amp; Yin, 2007
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Ant</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. adexilis" pageId="6" pageNumber="73" rank="species" species="adexilis">C. adexilis</taxonomicName>
<tr id="690145DC971A309341E6D75FA4A2AB0A" pageId="6" pageNumber="73">
<td id="20273410CA8EA5AE04ED1503AE4F1374" colspan="1" pageId="6" pageNumber="73" rowspan="1">Ant</td>
<td id="C719ED4BC9BF2973436BEEFAA50E4D60" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="21AFC74935DAF910140967114F1DDFE1" lsidName="C. adexilis" pageId="6" pageNumber="73" rank="species" species="adexilis">C. adexilis</taxonomicName>
Stach, 1964
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">AbdAntAbd</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. denticulata" pageId="6" pageNumber="73" rank="species" species="denticulata">C. denticulata</taxonomicName>
<tr id="4E3F9C56FC6252310E0509A51FA2247C" pageId="6" pageNumber="73">
<td id="D214D1A0F79A701B4E34A923DB009633" colspan="1" pageId="6" pageNumber="73" rowspan="1">AbdAntAbd</td>
<td id="84AE7B14482BD657D2B37F76A1E3BD1B" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="2CCA496863D30AEAD74D602B914A8EF0" lsidName="C. denticulata" pageId="6" pageNumber="73" rank="species" species="denticulata">C. denticulata</taxonomicName>
(Bagnall, 1941)
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">AbdAntorAbd</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. communis" pageId="6" pageNumber="73" rank="species" species="communis">C. communis</taxonomicName>
<tr id="5D977967D1A725E7EC6B8439D0FEFEDB" pageId="6" pageNumber="73">
<td id="1A4F6CB77D5475DF1705F2141BA15918" colspan="1" pageId="6" pageNumber="73" rowspan="1">AbdAntorAbd</td>
<td id="C4CB104AB5404AF5F2A6E3ED233B7576" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="F5AD78A1B7B92CF2D463D54B9F2DAD2A" lsidName="C. communis" pageId="6" pageNumber="73" rank="species" species="communis">C. communis</taxonomicName>
(Folsom, 1898)
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Abdor</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">10</td>
<tr id="327D88D9D29A3CCD85A911D5464D8ECE" pageId="6" pageNumber="73">
<td id="13DACF9774FB9CDE13E46846705DB4CF" colspan="1" pageId="6" pageNumber="73" rowspan="1">Abdor</td>
<td id="B9C277D2284E67A8E06D6B208042B63A" colspan="1" pageId="6" pageNumber="73" rowspan="1">10</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Abdor</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">11</td>
<tr id="D5A2ED91FD43987EE1B07B03FD3B396C" pageId="6" pageNumber="73">
<td id="9C8F119D135FFE4140C85000F8E7957C" colspan="1" pageId="6" pageNumber="73" rowspan="1">Abdor</td>
<td id="F40085C87C8DD6FC6C2A5D5DFE653403" colspan="1" pageId="6" pageNumber="73" rowspan="1">11</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Abd</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. liguladorsi" pageId="6" pageNumber="73" rank="species" species="liguladorsi">C. liguladorsi</taxonomicName>
<tr id="6FAFCF835112D3186E09023B4C6938FA" pageId="6" pageNumber="73">
<td id="D5B58DC9F9008D28DB666408FF461DAF" colspan="1" pageId="6" pageNumber="73" rowspan="1">Abd</td>
<td id="1DDC3F5AF018C341F0DB0FCC0FEE2BC6" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="211956558921DEEAB216BF404CDD0557" lsidName="C. liguladorsi" pageId="6" pageNumber="73" rank="species" species="liguladorsi">C. liguladorsi</taxonomicName>
(Lee, 1974)
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">Abd</td>
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName lsidName="C. duplicispinosa" pageId="6" pageNumber="73" rank="species" species="duplicispinosa">C. duplicispinosa</taxonomicName>
<tr id="982F9779DFEF912EE0532E1ED80E06D0" pageId="6" pageNumber="73">
<td id="875D57D0435DD18BCC6FF8FA60778809" colspan="1" pageId="6" pageNumber="73" rowspan="1">Abd</td>
<td id="634E528504AE0B1B82250B9EB8A202E2" colspan="1" pageId="6" pageNumber="73" rowspan="1">
<taxonomicName id="53283942DE5A6926B22FF3B8FF8958A0" lsidName="C. duplicispinosa" pageId="6" pageNumber="73" rank="species" species="duplicispinosa">C. duplicispinosa</taxonomicName>
(Yosii, 1954)
</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">12</td>
<tr id="6D47E937D2F4A9AE9D8228146376385C" pageId="6" pageNumber="73">
<td id="3A481BDAE0F89519596F3E7A9F2D0973" colspan="1" pageId="6" pageNumber="73" rowspan="1">12</td>
</tr>
<tr pageId="6" pageNumber="73">
<td colspan="1" pageId="6" pageNumber="73" rowspan="1">13</td>
<tr id="E941A31CCE53402792A1171B74CA7988" pageId="6" pageNumber="73">
<td id="7E0C048CE8A51445CA8B65593EE5515B" colspan="1" pageId="6" pageNumber="73" rowspan="1">13</td>
</tr>
<tr pageId="7" pageNumber="74">
<td colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName lsidName="C. xiaoi" pageId="7" pageNumber="74" rank="species" species="xiaoi">
<pageBreakToken pageId="7" pageNumber="74" start="start">C</pageBreakToken>
<tr id="9CCD1F627141971B6C04131C6EA43C8B" pageId="7" pageNumber="74">
<td id="994CFFAEE5D913966D9720865CFD7DC6" colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName id="518411D613A7D483718B1A8040FA530F" lsidName="C. xiaoi" pageId="7" pageNumber="74" rank="species" species="xiaoi">
<pageBreakToken id="B5C95E38C15450F77C85D6934C786CB4" pageId="7" pageNumber="74" start="start">C</pageBreakToken>
. xiaoi
</taxonomicName>
(Tamura, 1998)
<bibRefCitation author="Tamura, H" journalOrPublisher="Zoological Research" pageId="8" pageNumber="75" pagination="153 - 159" title="Three species of Collembola from Yunnan, Southwest China (Apterygota)." volume="19" year="1998">Tamura and Zhao 1998</bibRefCitation>
<bibRefCitation id="5852A7D9242546EC1A81C45FA7EBB38B" author="Tamura, H" journalOrPublisher="Zoological Research" pageId="8" pageNumber="75" pagination="153 - 159" title="Three species of Collembola from Yunnan, Southwest China (Apterygota)." volume="19" year="1998">Tamura and Zhao 1998</bibRefCitation>
</td>
</tr>
<tr pageId="7" pageNumber="74">
<td colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName lsidName="C. anshanensis" pageId="7" pageNumber="74" rank="species" species="anshanensis">C. anshanensis</taxonomicName>
<tr id="E246C0DED0121326824BCD44CCE56AEE" pageId="7" pageNumber="74">
<td id="30B3B630CB2EA6EDDD14D28262AA2D95" colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName id="F797DDF6AF3A23C9F948F8E21B83C10B" lsidName="C. anshanensis" pageId="7" pageNumber="74" rank="species" species="anshanensis">C. anshanensis</taxonomicName>
(Wu &amp; Xie, 2007)
</td>
</tr>
<tr pageId="7" pageNumber="74">
<td colspan="1" pageId="7" pageNumber="74" rowspan="1">14</td>
<tr id="7BD2223ED255F1FC4439285887A0CAAB" pageId="7" pageNumber="74">
<td id="0D996649CF564F47CD39D113F980FA8C" colspan="1" pageId="7" pageNumber="74" rowspan="1">14</td>
</tr>
<tr pageId="7" pageNumber="74">
<td colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName lsidName="C. sinensis" pageId="7" pageNumber="74" rank="species" species="sinensis">C. sinensis</taxonomicName>
<tr id="6CE1F317612FC892643577E516F47899" pageId="7" pageNumber="74">
<td id="223848C28C88A67C7DEF31199F60CA8A" colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName id="7C7C0B1BAB108BB97A326630BFA69570" lsidName="C. sinensis" pageId="7" pageNumber="74" rank="species" species="sinensis">C. sinensis</taxonomicName>
Stach, 1964
</td>
</tr>
<tr pageId="7" pageNumber="74">
<td colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName lsidName="C. skarzynskii" pageId="7" pageNumber="74" rank="species" species="skarzynskii">C. skarzynskii</taxonomicName>
<tr id="BE32A44F5ACC29F2BB78FE9815E49CD7" pageId="7" pageNumber="74">
<td id="50580A7F4FAA858F0237720D3A38449C" colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName id="A494635E101BDD99DBA52C5782767F54" lsidName="C. skarzynskii" pageId="7" pageNumber="74" rank="species" species="skarzynskii">C. skarzynskii</taxonomicName>
sp. n.
</td>
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<tr pageId="7" pageNumber="74">
<td colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName lsidName="C. flectochaeta" pageId="7" pageNumber="74" rank="species" species="flectochaeta">C. flectochaeta</taxonomicName>
<tr id="4E9AEE63B5B39FFD03B4B51B8F3B9364" pageId="7" pageNumber="74">
<td id="88436326EC935257F2733BE4EDD8C1B9" colspan="1" pageId="7" pageNumber="74" rowspan="1">
<taxonomicName id="849E1DAA4BDC3D7D424FF9E55D6F04BE" lsidName="C. flectochaeta" pageId="7" pageNumber="74" rank="species" species="flectochaeta">C. flectochaeta</taxonomicName>
Lin &amp; Xia, 1983
</td>
</tr>

View file

@ -1,151 +1,151 @@
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<taxonomicName LSID="http://species-id.net/wiki/Philopteroides_beckeri" authority="Mey, 2004" authorityName="Mey" authorityYear="2004" class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides beckeri" order="Psocodea" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="beckeri">Philopteroides beckeri (Mey, 2004)</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="74">comb. n.</taxonomicNameLabel>
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<paragraph id="30B6F9A593713741058AE70C28B7D6C0" pageId="3" pageNumber="74">
<taxonomicName id="DC7705422D51A72AFAED218DA59A9DB4" LSID="http://species-id.net/wiki/Philopteroides_beckeri" authority="Mey, 2004" authorityName="Mey" authorityYear="2004" class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides beckeri" order="Psocodea" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="beckeri">Philopteroides beckeri (Mey, 2004)</taxonomicName>
<taxonomicNameLabel id="E21CC165C22438A05E1D8BAE14DF90C6" pageId="3" pageNumber="74">comb. n.</taxonomicNameLabel>
Figs 1-2, 9, 13-16, 29-31
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="74" type="reference_group">
<paragraph pageId="3" pageNumber="74">
<taxonomicName class="Insecta" family="Philopteridae" genus="Tyranniphilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tyranniphilopterus beckeri" order="Phthiraptera" pageId="3" pageNumber="74" phylum="Arthropoda" rank="species" species="beckeri">Tyranniphilopterus beckeri</taxonomicName>
<bibRefCitation author="Mey, E" journalOrPublisher="Ornithologischer Anzeiger" pageId="15" pageNumber="86" pagination="149 - 203" title="Zur Taxonomie, Verbreitung und parasitophyletischer Evidenz des Philopterus-Komplexes (Insecta, Phthiraptera, Ischnocera)." volume="43" year="2004">Mey 2004</bibRefCitation>
<subSubSection id="EDA45CF73C6BF782265474104421E508" pageId="3" pageNumber="74" type="reference_group">
<paragraph id="4193B1271BD035137E19246CDEBC12DA" pageId="3" pageNumber="74">
<taxonomicName id="0504BE49DF96C4E7503B1D1449EEF844" genus="Tyranniphilopterus" lsidName="Tyranniphilopterus beckeri" pageId="3" pageNumber="74" rank="species" species="beckeri">Tyranniphilopterus beckeri</taxonomicName>
<bibRefCitation id="3F2D55490CACDFBEDE2C5AAFCC989931" author="Mey, E" journalOrPublisher="Ornithologischer Anzeiger" pageId="15" pageNumber="86" pagination="149 - 203" title="Zur Taxonomie, Verbreitung und parasitophyletischer Evidenz des Philopterus-Komplexes (Insecta, Phthiraptera, Ischnocera)." volume="43" year="2004">Mey 2004</bibRefCitation>
: 182;
<bibRefCitation author="Cicchino, AC" journalOrPublisher="Zootaxa" pageId="15" pageNumber="86" pagination="43 - 50" volume="1547" year="2007">Cicchino 2007</bibRefCitation>
<bibRefCitation id="63F761FDD0FAD01910442BD737DF81E8" author="Cicchino, AC" journalOrPublisher="Zootaxa" pageId="15" pageNumber="86" pagination="43 - 50" volume="1547" year="2007">Cicchino 2007</bibRefCitation>
: 49.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="74" type="type host">
<paragraph pageId="3" pageNumber="74">Type host.</paragraph>
<paragraph pageId="3" pageNumber="74">
<taxonomicName class="Aves" family="Platysteiridae" genus="Platysteira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platysteira cyanea subsp. nyansae" order="Passeriformes" pageId="3" pageNumber="74" phylum="Chordata" rank="subSpecies" species="cyanea" subSpecies="nyansae">Platysteira cyanea nyansae</taxonomicName>
<subSubSection id="653AAB4A42921DD5AD46E308363EF719" pageId="3" pageNumber="74" type="type host">
<paragraph id="DDBA261E1FE325E6AE1021E0C60B807B" pageId="3" pageNumber="74">Type host.</paragraph>
<paragraph id="2AD4383B35D94B202280BAB9A4779B00" pageId="3" pageNumber="74">
<taxonomicName id="89DA87D8D0F02D135C8F6591ADFA9F9F" class="Aves" family="Platysteiridae" genus="Platysteira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platysteira cyanea subsp. nyansae" order="Passeriformes" pageId="3" pageNumber="74" phylum="Chordata" rank="subSpecies" species="cyanea" subSpecies="nyansae">Platysteira cyanea nyansae</taxonomicName>
Neumann, 1905 - Brown-throated wattle-eye. (
<taxonomicName family="Platysteiridae" lsidName="" pageId="3" pageNumber="74" rank="family">Platysteiridae</taxonomicName>
<taxonomicName id="8E2B3519DE4191FD569FC54D79372BAB" class="Aves" family="Platysteiridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Passeriformes" pageId="3" pageNumber="74" phylum="Chordata" rank="family">Platysteiridae</taxonomicName>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="76" pageId="3" pageNumber="74" type="distribution">
<paragraph pageId="3" pageNumber="74">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="74">Uganda.</paragraph>
<paragraph pageId="3" pageNumber="74">
<subSubSection id="84FA9A51A17CF0D28A007A46B15697CA" lastPageId="5" lastPageNumber="76" pageId="3" pageNumber="74" type="distribution">
<paragraph id="9BCCF02724DCC852AAA33F35F4C84A70" pageId="3" pageNumber="74">Distribution.</paragraph>
<paragraph id="DEA4727FC80A303BFD044D6A3D91F300" pageId="3" pageNumber="74">Uganda.</paragraph>
<paragraph id="62EDE34792F751893B4C855E17F0FA71" pageId="3" pageNumber="74">
This species was recently described in detail and, therefore, it is not necessary to redescribed it again. We only include habitus images of the holotype male and one paratype female (Figs 1-2), not figured in the original description by
<bibRefCitation author="Mey, E" journalOrPublisher="Ornithologischer Anzeiger" pageId="15" pageNumber="86" pagination="149 - 203" title="Zur Taxonomie, Verbreitung und parasitophyletischer Evidenz des Philopterus-Komplexes (Insecta, Phthiraptera, Ischnocera)." volume="43" year="2004">Mey (2004)</bibRefCitation>
<bibRefCitation id="340E880631ABB2CA3EA018942C91B68F" author="Mey, E" journalOrPublisher="Ornithologischer Anzeiger" pageId="15" pageNumber="86" pagination="149 - 203" title="Zur Taxonomie, Verbreitung und parasitophyletischer Evidenz des Philopterus-Komplexes (Insecta, Phthiraptera, Ischnocera)." volume="43" year="2004">Mey (2004)</bibRefCitation>
. We also include illustrations to support our change of the original generic combination of this species, and have re-drawn only those characters (Figs 9, 13-16, 30) useful to distinguish it from the second species described below. In addition, the second nymphal stage is described from a single specimen (Fig. 29) mounted together with the female paratype, and a re-interpretation of the male genitalia is presented in Fig. 31.
</paragraph>
<paragraph lastPageId="5" lastPageNumber="76" pageId="3" pageNumber="74">
<paragraph id="D6B8FF54BE05858CE1AAB1D0E31E2CD3" lastPageId="5" lastPageNumber="76" pageId="3" pageNumber="74">
All species of the genus
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<pageBreakToken pageId="4" pageNumber="75" start="start">Tyranniphilopterus</pageBreakToken>
<taxonomicName id="77E58B10BC6E61D7D14F58A0225871C5" genus="Tyranniphilopterus" lsidName="Tyranniphilopterus" pageId="4" pageNumber="75" rank="genus">
<pageBreakToken id="1C91AC7DF0881CBF40A28D417292F1FD" pageId="4" pageNumber="75" start="start">Tyranniphilopterus</pageBreakToken>
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Mey, 2004 have the following characters, which are lacking in &quot;
<taxonomicName class="Insecta" family="Philopteridae" genus="Tyranniphilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tyranniphilopterus" order="Phthiraptera" pageId="4" pageNumber="75" phylum="Arthropoda" rank="genus">Tyranniphilopterus</taxonomicName>
<taxonomicName id="67192F27EEC5A70DB50559C907B031AE" genus="Tyranniphilopterus" lsidName="Tyranniphilopterus" pageId="4" pageNumber="75" rank="genus">Tyranniphilopterus</taxonomicName>
&quot; beckeri: (1) hyaline margin arises at a level below the as3; (2) conspicuous antero-lateral projections on anterior dorsal head plate; (3) prothoracic dorsal setae located on the posterior-lateral angles of the seg
<pageBreakToken pageId="5" pageNumber="76" start="start">ment</pageBreakToken>
<pageBreakToken id="C763FFA08FFD20F65F7C55B6778BCFCE" pageId="5" pageNumber="76" start="start">ment</pageBreakToken>
, not on its posterior margin.; (4) pleuro-tergal plates
<normalizedToken originalValue="IIIII">II-III</normalizedToken>
<normalizedToken id="9E2B2C17BF359C87E1B4C30C58736137" originalValue="IIIII">II-III</normalizedToken>
with well developed postero-lateral projections. Instead, &quot;
<taxonomicName class="Insecta" family="Philopteridae" genus="Tyranniphilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tyranniphilopterus" order="Phthiraptera" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Tyranniphilopterus</taxonomicName>
<taxonomicName id="2AA46B12ABF18FEAB83DD8C321D5D9CB" genus="Tyranniphilopterus" lsidName="Tyranniphilopterus" pageId="5" pageNumber="76" rank="genus">Tyranniphilopterus</taxonomicName>
&quot; beckeri has the features which define
<taxonomicName class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides" order="Psocodea" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Philopteroides</taxonomicName>
<taxonomicName id="7FECA4A1BFC4E3755168DA1A8E6BBCD9" class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides" order="Psocodea" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Philopteroides</taxonomicName>
. These morphological features justify placing it in the latter genus, as
<taxonomicName class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides beckeri" order="Psocodea" pageId="5" pageNumber="76" phylum="Arthropoda" rank="species" species="beckeri">Philopteroides beckeri</taxonomicName>
<taxonomicName id="A657ADA9656A3634160138D5471C61CF" class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides beckeri" order="Psocodea" pageId="5" pageNumber="76" phylum="Arthropoda" rank="species" species="beckeri">Philopteroides beckeri</taxonomicName>
(Mey, 2004) new combination.
</paragraph>
<paragraph pageId="5" pageNumber="76">
<paragraph id="9A00FDC6C10A77E36A0E6AD6F1E6409E" pageId="5" pageNumber="76">
Furthermore, the hosts of all species of
<taxonomicName class="Insecta" family="Philopteridae" genus="Tyranniphilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tyranniphilopterus" order="Phthiraptera" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Tyranniphilopterus</taxonomicName>
<taxonomicName id="F49F87B6494EED76E0F8CEF3B0A27856" genus="Tyranniphilopterus" lsidName="Tyranniphilopterus" pageId="5" pageNumber="76" rank="genus">Tyranniphilopterus</taxonomicName>
- except for &quot;
<taxonomicName class="Insecta" family="Philopteridae" genus="Tyranniphilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tyranniphilopterus" order="Phthiraptera" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Tyranniphilopterus</taxonomicName>
<taxonomicName id="16DCF80F8FB4344D7F131F700B2B38C4" genus="Tyranniphilopterus" lsidName="Tyranniphilopterus" pageId="5" pageNumber="76" rank="genus">Tyranniphilopterus</taxonomicName>
&quot; beckeri - belong to the passerine suborders
<taxonomicName genus="Tyranni" lsidName="Tyranni" pageId="5" pageNumber="76" rank="genus">Tyranni</taxonomicName>
<taxonomicName id="8F574AC9EDC6B578BB68C1D8E8F10AA1" genus="Tyranni" lsidName="Tyranni" pageId="5" pageNumber="76" rank="genus">Tyranni</taxonomicName>
and
<taxonomicName genus="Passeri" lsidName="Passeri" pageId="5" pageNumber="76" rank="genus">Passeri</taxonomicName>
<taxonomicName id="946714953C4BB615B6A6F1205F0AC354" genus="Passeri" lsidName="Passeri" pageId="5" pageNumber="76" rank="genus">Passeri</taxonomicName>
, and are confined to the Americas, while the hosts of all species of
<taxonomicName class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides" order="Psocodea" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Philopteroides</taxonomicName>
<taxonomicName id="A58A1D16F97071BA5C4BFF2257AD314E" class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides" order="Psocodea" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Philopteroides</taxonomicName>
belong to the passerine suborders
<taxonomicName genus="Passeri" lsidName="Passeri" pageId="5" pageNumber="76" rank="genus">Passeri</taxonomicName>
<taxonomicName id="A98626DF82E0EBF763072A6D5825CDF2" genus="Passeri" lsidName="Passeri" pageId="5" pageNumber="76" rank="genus">Passeri</taxonomicName>
and
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<taxonomicName id="1E43024E48CF65ED614DC567EA3D0216" genus="Acanthisitti" lsidName="Acanthisitti" pageId="5" pageNumber="76" rank="genus">Acanthisitti</taxonomicName>
distributed in Africa, Asia and Oceania. Hence, the geographical distribution of its host is further evidence that placing &quot;
<taxonomicName class="Insecta" family="Philopteridae" genus="Tyranniphilopterus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tyranniphilopterus" order="Phthiraptera" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Tyranniphilopterus</taxonomicName>
<taxonomicName id="2F5A483B1EFDA4FA6B16ABC46E15CA17" genus="Tyranniphilopterus" lsidName="Tyranniphilopterus" pageId="5" pageNumber="76" rank="genus">Tyranniphilopterus</taxonomicName>
&quot; beckeri in the genus
<taxonomicName class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides" order="Psocodea" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Philopteroides</taxonomicName>
<taxonomicName id="C852597710AC7770EDE043535975C682" class="Insecta" family="Philopteridae" genus="Philopteroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philopteroides" order="Psocodea" pageId="5" pageNumber="76" phylum="Arthropoda" rank="genus">Philopteroides</taxonomicName>
is correct.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="76" type="description">
<paragraph pageId="5" pageNumber="76">Description of second nymphal stage.</paragraph>
<paragraph pageId="5" pageNumber="76">
<subSubSection id="F796801F5C89246FBFD885182992064C" pageId="5" pageNumber="76" type="description">
<paragraph id="371B4FC87D87B3C6973FCBC38C4217A8" pageId="5" pageNumber="76">Description of second nymphal stage.</paragraph>
<paragraph id="6FC0A17B65B63AEC5F02647585A62C63" pageId="5" pageNumber="76">
Head, thorax and abdomen as in Fig. 29. Head sub-conical, marginal carina entire laterally, with well developed anterior dorsal and ventral head plates. Anterior setae 3 (as3) rigid, 0.04 in length. Dorsal head plate with convex lateral margins and almost straight posterior margin, bearing rigid anterior dorsal head setae (ads), as the adults. Anterior ventral plate cordiform. Marginal temporal setae 3 (m.t.s.3) very long, other temporal setae short to minute. Each of the pair of posterior setae on pronotum located half way between the middle of the segment and its lateroposterior angle. Only four long pterothoracic setae, as in the second nymphal stage of most species of
<taxonomicName family="Philopteridae" lsidName="" pageId="5" pageNumber="76" rank="family">Philopteridae</taxonomicName>
<taxonomicName id="CC5BC88450009C121D7F1EDB974F5EE4" class="Insecta" family="Philopteridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Psocodea" pageId="5" pageNumber="76" phylum="Arthropoda" rank="family">Philopteridae</taxonomicName>
(
<bibRefCitation author="Mey, E" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="15" pageNumber="86" pagination="3 - 84" title="Beziehungen zwischen Larvenmorphologie und Systematik der Adulti bei den Vogel-Ischnozeren (Insecta, Phthiraptera, Ischnocera)." url="10.1002/mmnz.19940700102" volume="70" year="1994">Mey 1994</bibRefCitation>
<bibRefCitation id="B17FDD18CDEA64426287599D2DCC64B1" author="Mey, E" journalOrPublisher="Mitteilungen aus dem Zoologischen Museum in Berlin" pageId="15" pageNumber="86" pagination="3 - 84" title="Beziehungen zwischen Larvenmorphologie und Systematik der Adulti bei den Vogel-Ischnozeren (Insecta, Phthiraptera, Ischnocera)." url="10.1002/mmnz.19940700102" volume="70" year="1994">Mey 1994</bibRefCitation>
). Abdomen mostly membranous, except for the tergo-pleural plate of second segment (first visible), and the pleural plates of segments
<normalizedToken originalValue="IIIV">III-V</normalizedToken>
<normalizedToken id="3BC7E739838AF4CC78DC35E4F086A972" originalValue="IIIV">III-V</normalizedToken>
. Segments VI and VII with small patches of light sclerotization. Abdominal chaetotaxy as in Fig. 29, with two long dorsal setae plus the postspiracular seta on each side of segments
<normalizedToken originalValue="IIIVIII">III-VIII</normalizedToken>
<normalizedToken id="2958E843705F34992BE94E4CEBA942DE" originalValue="IIIVIII">III-VIII</normalizedToken>
. Sternites
<normalizedToken originalValue="IIIII">II-III</normalizedToken>
<normalizedToken id="A5C3DB10CF57C8452B4B64EBD61C2976" originalValue="IIIII">II-III</normalizedToken>
with one spine-like setae among three long setae on each side,
<normalizedToken originalValue="IVV">IV-V</normalizedToken>
<normalizedToken id="9130E41F9E71226C953EF92AA73CAF7C" originalValue="IVV">IV-V</normalizedToken>
with a pair of long setae each side, and
<normalizedToken originalValue="VIVIII">VI-VIII</normalizedToken>
<normalizedToken id="6D80957AE0332306D3601A52B66BDF35" originalValue="VIVIII">VI-VIII</normalizedToken>
with only one long setae on each side. One long pleural seta on
<normalizedToken originalValue="IIIVIII">III-VIII</normalizedToken>
<normalizedToken id="4FF022FA68BFE09A658AF89A7AF8638C" originalValue="IIIVIII">III-VIII</normalizedToken>
. Measurements (n=1): HL 0.37, ANW 0.11, POL 0.14, POW 0.27, ADPL 0.12, ADPW 0.14, TRL 0.08, TRW 0.04, TW 0.36, PW 0.21, PTW 0.29, AW 0.43, TL 1.06.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="76" type="type specimens">
<paragraph pageId="5" pageNumber="76">Type specimens.</paragraph>
<paragraph pageId="5" pageNumber="76">
<subSubSection id="6EF639D0F63DB59271914822D9B43B93" pageId="5" pageNumber="76" type="type specimens">
<paragraph id="7FE788F2D9A86A779D1D6EA519229E48" pageId="5" pageNumber="76">Type specimens.</paragraph>
<paragraph id="6EBFFE93A257DF178852919FB6A04DCC" pageId="5" pageNumber="76">
Holotype ♂ (NHMR #4322.c) and 1♀ paratype (NHMR #4322.b), ex
<taxonomicName class="Aves" family="Platysteiridae" genus="Platysteira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platysteira cyanea subsp. nyansae" order="Passeriformes" pageId="5" pageNumber="76" phylum="Chordata" rank="subSpecies" species="cyanea" subSpecies="nyansae">Platysteira cyanea nyansae</taxonomicName>
<taxonomicName id="B1BD9C8E037C0DC443C9958802907C33" class="Aves" family="Platysteiridae" genus="Platysteira" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platysteira cyanea subsp. nyansae" order="Passeriformes" pageId="5" pageNumber="76" phylum="Chordata" rank="subSpecies" species="cyanea" subSpecies="nyansae">Platysteira cyanea nyansae</taxonomicName>
; UGANDA: Paraa, Murchison Falls, National park, 22.X.1998, P. Becker col.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="76" type="non-type specimen">
<paragraph pageId="5" pageNumber="76">Non-type specimen.</paragraph>
<paragraph pageId="5" pageNumber="76">1 nymph II (NHMR #4322.b), mounted on the same slide as the female paratype.</paragraph>
<subSubSection id="15429D4ADDC7EB7F7A920BEB6BFEE61A" pageId="5" pageNumber="76" type="non-type specimen">
<paragraph id="88C762583DE1318E24686AC93DF067AD" pageId="5" pageNumber="76">Non-type specimen.</paragraph>
<paragraph id="639083E519C90131721F2B5686231A38" pageId="5" pageNumber="76">1 nymph II (NHMR #4322.b), mounted on the same slide as the female paratype.</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,231 +1,231 @@
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4.
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<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">Figs 19-22</figureCitation>
<figureCitation captionStart="Figures 2327" captionStartId="F7" captionText="Figures 23 - 27. Tipula (Nippotipula) champasakensis sp. nov., male 23 hypopygium, lateral view 24 ninth tergite, dorsal view 25 eighth sternite, ventral view; 26 lobe of gonostylus, lateral view 27 clasper of gonostylus, lateral view. tg 8 = eighth tergite, tg 9 = ninth tergite, st 8 = eighth sternite, st 9 = ninth sternite, lg = lobe of gonostylus. Scale bars: 1.0 mm (23 - 25); 0.5 mm (26, 27)." figureDoi="10.3897/zookeys.982.49781.figures23-27" httpUri="https://binary.pensoft.net/fig/470383" pageId="0" pageNumber="11">, 23-27</figureCitation>
<figureCitation captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">, 28-33</figureCitation>
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<figureCitation id="EC511FE41C4F7459A64ACBAC4F15F55E" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">Figs 19-22</figureCitation>
<figureCitation id="A1591A52E3CFF19917B92795B12931FF" captionStart="Figures 2327" captionStartId="F7" captionText="Figures 23 - 27. Tipula (Nippotipula) champasakensis sp. nov., male 23 hypopygium, lateral view 24 ninth tergite, dorsal view 25 eighth sternite, ventral view; 26 lobe of gonostylus, lateral view 27 clasper of gonostylus, lateral view. tg 8 = eighth tergite, tg 9 = ninth tergite, st 8 = eighth sternite, st 9 = ninth sternite, lg = lobe of gonostylus. Scale bars: 1.0 mm (23 - 25); 0.5 mm (26, 27)." figureDoi="10.3897/zookeys.982.49781.figures23-27" httpUri="https://binary.pensoft.net/fig/470383" pageId="0" pageNumber="11">, 23-27</figureCitation>
<figureCitation id="143818EC1EBB41E0F7A8AEE15E069A7E" captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">, 28-33</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="diagnosis">
<paragraph pageId="0" pageNumber="11">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="11">Abdomen is reddish brown with thick brownish yellow setae. Eighth sternite extended backward, posterior margin has a deep median notch. Posterior margin of ninth tergite extended with sclerotized depression. Lobe of gonostylus is fleshy with a V-shaped depression at posterior margin. Clasper of gonostylus is duck-shaped.</paragraph>
<subSubSection id="195A16CA3F9BA85E257E8F24601B3050" pageId="0" pageNumber="11" type="diagnosis">
<paragraph id="4E324CE982A56470D425E081535E4E2E" pageId="0" pageNumber="11">Diagnosis.</paragraph>
<paragraph id="0906BB9E5E607A869C5FB0BA7AFBBCBB" pageId="0" pageNumber="11">Abdomen is reddish brown with thick brownish yellow setae. Eighth sternite extended backward, posterior margin has a deep median notch. Posterior margin of ninth tergite extended with sclerotized depression. Lobe of gonostylus is fleshy with a V-shaped depression at posterior margin. Clasper of gonostylus is duck-shaped.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="description">
<paragraph pageId="0" pageNumber="11">Description.</paragraph>
<paragraph pageId="0" pageNumber="11">
<subSubSection id="997376FF09BF92B20636B4D64AA43087" pageId="0" pageNumber="11" type="description">
<paragraph id="0F307E1A71546F7D4DA3E33B7CB20B42" pageId="0" pageNumber="11">Description.</paragraph>
<paragraph id="23D15AEFCE617FC5C1D4D27BFB98FDF4" pageId="0" pageNumber="11">
Male (
<emphasis italics="true" pageId="0" pageNumber="11">n</emphasis>
<emphasis id="F3A3C76FD3FCD6CAE1C75B0E7A663684" italics="true" pageId="0" pageNumber="11">n</emphasis>
= 8): body length 20-23 mm, wing length 16-18 mm, antenna length 5-6 mm.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<paragraph id="E8DA1A0E0F8B659F441BA4BF954816EB" pageId="0" pageNumber="11">
Head (Figs
<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">19</figureCitation>
<figureCitation id="9BAB4A99AB35A9B46B3C7A99B84810D9" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">19</figureCitation>
,
<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">21</figureCitation>
<figureCitation id="7111F6623A36242DE2677A77D7C724A9" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">21</figureCitation>
). Mostly brownish yellow. Dorsal part of rostrum brownish yellow; nasus blacked; eyes dark black. Setae on head black. Antenna dark brown, except scape brownish and pedicel yellow; first flagellomere longest, slightly longer than scape. Proboscis mostly brownish, with black setae. Palpus greyish brown, with black setae.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<paragraph id="714CE8F4980DD628631877A63B69AEEE" pageId="0" pageNumber="11">
Thorax (Figs
<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">19</figureCitation>
<figureCitation id="266654E32CFFAE3F4DDE322A396EA3F1" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">19</figureCitation>
,
<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">21</figureCitation>
<figureCitation id="523A16681B754BDB74B9569BC5AB14E7" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">21</figureCitation>
). Mainly brownish. Prescutum brownish yellow with three pale yellow stripes; median stripe broad basally with a light brown median line, lateral stripes oval, shorter than median stripe. Scutum brownish yellow, each lobe with two pale yellow stripes. Scutellum gray-yellow. Mediotergite yellow with long brownish yellow setae. Thoracic pleuron mostly brownish throughout, except middle of pleuron bright yellow or tinged brown. Setae on thorax brownish yellow. Legs coxae and trochanters grayish yellow, femora light brown, with subterminal dark band, tibiae, and tarsi brownish. Setae on legs black except those on coxae grayish yellow. Wings brownish yellow, with dark brown spots at the origin of Rs, M, and R4+5. Pterostigma dark brown with some macrotrichiae, Rs relatively long; cell m1 petiolate (Fig.
<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">22</figureCitation>
<figureCitation id="F2E20FDCF31BBD2A106B94A824B0DECC" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">22</figureCitation>
). Halter length approximately 2.5 mm; halter stem brownish yellow, with brownish yellow setae; halter knob pale yellow (Fig.
<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">21</figureCitation>
<figureCitation id="5B76EE4187C9E4BFBD637CADE8BDAF05" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">21</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="11">
<paragraph id="1E836A7473AB04418837BF049B8E1E58" pageId="0" pageNumber="11">
Abdomen (Fig.
<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">19</figureCitation>
<figureCitation id="26AB69AB9D2481AD64C959E21D397ECF" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">19</figureCitation>
). Mainly brownish yellow. Abdominal segments 1-6 brownish yellow with yellow setae; segments 7 and 8 with brown setae. Hypopygium reddish brown, with brown setae.
</paragraph>
<caption doi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11" start="Figures 1922" startId="F6">
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">Figures 19-22.</emphasis>
<taxonomicName authorityName="Zhang &amp; Ren &amp; Li &amp; Yang" authorityYear="2020" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Nippotipula) champasakensis" order="Diptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="champasakensis" subGenus="Nippotipula">Tipula (Nippotipula) champasakensis</taxonomicName>
<caption id="4E3868CB2D91F45BB390EC050AC725BD" doi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11" start="Figures 1922" startId="F6">
<paragraph id="A14F365DD25E5CFFE68C3CF2CD901D8D" pageId="0" pageNumber="11">
<emphasis id="5E9AFCA17E67A500F735B4846D59060F" bold="true" pageId="0" pageNumber="11">Figures 19-22.</emphasis>
<taxonomicName id="4CF17F88660A8F980CC41B776E5D3671" authorityName="Zhang &amp; Ren &amp; Li &amp; Yang" authorityYear="2020" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Nippotipula) champasakensis" order="Diptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="champasakensis" subGenus="Nippotipula">Tipula (Nippotipula) champasakensis</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="11">19</emphasis>
<emphasis id="C5F1B42FA97E0593641BF2EA5607F224" bold="true" pageId="0" pageNumber="11">19</emphasis>
male habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="11">20</emphasis>
<emphasis id="B2D49B2536A7E95F958BDF4195D6AD5B" bold="true" pageId="0" pageNumber="11">20</emphasis>
female habitus, lateral view;
<emphasis bold="true" pageId="0" pageNumber="11">21</emphasis>
<emphasis id="DDC6F6666DB7E9127ED20282BFD52213" bold="true" pageId="0" pageNumber="11">21</emphasis>
male head and thorax, dorsal view
<emphasis bold="true" pageId="0" pageNumber="11">22</emphasis>
<emphasis id="1F89D079886FD8C93611E6F40507362E" bold="true" pageId="0" pageNumber="11">22</emphasis>
male right wing. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="11">
<paragraph id="194D1BD5B043C2EED4BABD77DA2765A4" pageId="0" pageNumber="11">
Hypopygium (Figs
<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">19</figureCitation>
<figureCitation id="F5D139D77CFDCBCF186F2D0F8999B00F" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">19</figureCitation>
,
<figureCitation captionStart="Figures 2327" captionStartId="F7" captionText="Figures 23 - 27. Tipula (Nippotipula) champasakensis sp. nov., male 23 hypopygium, lateral view 24 ninth tergite, dorsal view 25 eighth sternite, ventral view; 26 lobe of gonostylus, lateral view 27 clasper of gonostylus, lateral view. tg 8 = eighth tergite, tg 9 = ninth tergite, st 8 = eighth sternite, st 9 = ninth sternite, lg = lobe of gonostylus. Scale bars: 1.0 mm (23 - 25); 0.5 mm (26, 27)." figureDoi="10.3897/zookeys.982.49781.figures23-27" httpUri="https://binary.pensoft.net/fig/470383" pageId="0" pageNumber="11">23-27</figureCitation>
<figureCitation id="39865FE5A2D07001101EA1A534D8A124" captionStart="Figures 2327" captionStartId="F7" captionText="Figures 23 - 27. Tipula (Nippotipula) champasakensis sp. nov., male 23 hypopygium, lateral view 24 ninth tergite, dorsal view 25 eighth sternite, ventral view; 26 lobe of gonostylus, lateral view 27 clasper of gonostylus, lateral view. tg 8 = eighth tergite, tg 9 = ninth tergite, st 8 = eighth sternite, st 9 = ninth sternite, lg = lobe of gonostylus. Scale bars: 1.0 mm (23 - 25); 0.5 mm (26, 27)." figureDoi="10.3897/zookeys.982.49781.figures23-27" httpUri="https://binary.pensoft.net/fig/470383" pageId="0" pageNumber="11">23-27</figureCitation>
). Eighth sternite extended backward, posterior margin with a deep median notch and setae; two small, triangular processes present at bottom of notch. Posterior margin of ninth tergite extended with ossified depression; middle of extension with a small spiny protrusion. Ninth sternite with dark-brown setae laterally. Lobe of gonostylus fleshy with dense, dark-brown setae on outer side and dense, black, obtuse spinules on inner side, posterior margin with a V-shaped depression. Clasper of gonostylus duck-shaped; posterior crest with longer setae.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<paragraph id="12FD93522B3CDEF1D141989FBC5DB892" pageId="0" pageNumber="11">
Semen pump (Figs
<figureCitation captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">28-30</figureCitation>
<figureCitation id="BC2BC2BDE5614B230477DC262A27D202" captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">28-30</figureCitation>
). Posterior immovable apodeme (pia) reddish brown, rod-like, and directed backward, symmetrical on both sides. Compressor apodeme (ca) fan-shaped and directed ventrally. Anterior immovable apodeme (aia) wing-shaped. Other appendages complex, as shown in Figures
<figureCitation captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">28-30</figureCitation>
<figureCitation id="4447D9F1F38BEA0013D0184AC87ED213" captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">28-30</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<paragraph id="895B8B807D21053FE889B75D0383C99E" pageId="0" pageNumber="11">
Female (
<emphasis italics="true" pageId="0" pageNumber="11">n</emphasis>
<emphasis id="3FBAF1A89550F034710D7D2B91B1F4A5" italics="true" pageId="0" pageNumber="11">n</emphasis>
= 2): body length 22-23 mm, wing length 17-18 mm, antenna length 5-6 mm.
</paragraph>
<paragraph pageId="0" pageNumber="11">
<paragraph id="A500163257FEEBA99B7AEAEE2465769A" pageId="0" pageNumber="11">
Female resembles male in head and thorax, except abdomen plump (Fig.
<figureCitation captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">20</figureCitation>
<figureCitation id="E34DE76717EC03020C0210200F007A08" captionStart="Figures 1922" captionStartId="F6" captionText="Figures 19 - 22. Tipula (Nippotipula) champasakensis sp. nov. 19 male habitus, lateral view 20 female habitus, lateral view; 21 male head and thorax, dorsal view 22 male right wing. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.982.49781.figures19-22" httpUri="https://binary.pensoft.net/fig/470382" pageId="0" pageNumber="11">20</figureCitation>
). Eighth tergite and eighth sternite reddish brown throughout. Ninth tergite, ninth sternite, and basal region of tenth tergite dark reddish brown (Figs
<figureCitation captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">31-33</figureCitation>
<figureCitation id="38DD1E3B3A21B5C099DD0E93E84965AE" captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">31-33</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="11">
<paragraph id="AF30205503F0D0C288B3426927222DA0" pageId="0" pageNumber="11">
Ovipositor reddish brown (Figs
<figureCitation captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">31-33</figureCitation>
<figureCitation id="8C5CF692323732C887AAB0D620AD6304" captionStart="Figures 2833" captionStartId="F8" captionText="Figures 28 - 33. Tipula (Nippotipula) champasakensis sp. nov. 28 - 30 male 28 semen pump, ventral view 29 semen pump, dorsal view 30 semen pump, lateral view 31 - 33 female 31 ovipositor, lateral view 32 ovipositor, dorsal view 33 ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (28 - 30); 1.0 mm (31 - 33)." figureDoi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11">31-33</figureCitation>
). Cercus narrowed toward tip. Hypovalve curved, broad apically.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="type material">
<paragraph pageId="0" pageNumber="11">Type material.</paragraph>
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" italics="true" pageId="0" pageNumber="11">Holotype</emphasis>
<subSubSection id="B04E0728A9B6580DBA781AC09B708970" pageId="0" pageNumber="11" type="type material">
<paragraph id="BF04608BF18D71127435D2B3602C6512" pageId="0" pageNumber="11">Type material.</paragraph>
<paragraph id="500708AB724678E2D6C63FBFCE616918" pageId="0" pageNumber="11">
<emphasis id="5AB1F248A4FF37653D12C9197FB12A02" bold="true" italics="true" pageId="0" pageNumber="11">Holotype</emphasis>
: male (CAU), Laos: Champasak, Soukhouma, Dond Hua Sao N. B., 2017.VI.19, Liang Wang (light trap).
<emphasis bold="true" italics="true" pageId="0" pageNumber="11">Paratypes</emphasis>
<emphasis id="12713DD32BC0E9ABB4EAC54F0AB13091" bold="true" italics="true" pageId="0" pageNumber="11">Paratypes</emphasis>
: 2 males 2 females (CAU), Laos: Champasak, Sabaidee Valley, 2017.VI.16, Liang Wang (light trap); 5 males (CAU), Laos: Champasak, Soukhouma, Dond Hua Sao N. B., 2017.VI.19, Liang Wang (light trap).
</paragraph>
<caption doi="10.3897/zookeys.982.49781.figures23-27" httpUri="https://binary.pensoft.net/fig/470383" pageId="0" pageNumber="11" start="Figures 2327" startId="F7">
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">Figures 23-27.</emphasis>
<taxonomicName authorityName="Zhang &amp; Ren &amp; Li &amp; Yang" authorityYear="2020" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Nippotipula) champasakensis" order="Diptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="champasakensis" subGenus="Nippotipula">Tipula (Nippotipula) champasakensis</taxonomicName>
<caption id="3ECE0EA0C147D7671C96516C89DA8729" doi="10.3897/zookeys.982.49781.figures23-27" httpUri="https://binary.pensoft.net/fig/470383" pageId="0" pageNumber="11" start="Figures 2327" startId="F7">
<paragraph id="432029D3C8F231DE734F85457F87F57A" pageId="0" pageNumber="11">
<emphasis id="0AB52CEDAD11063F7278ECBC90DB9E66" bold="true" pageId="0" pageNumber="11">Figures 23-27.</emphasis>
<taxonomicName id="9CD44D7C72CBC9C0331D5865973AE8B9" authorityName="Zhang &amp; Ren &amp; Li &amp; Yang" authorityYear="2020" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Nippotipula) champasakensis" order="Diptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="champasakensis" subGenus="Nippotipula">Tipula (Nippotipula) champasakensis</taxonomicName>
sp. nov., male
<emphasis bold="true" pageId="0" pageNumber="11">23</emphasis>
<emphasis id="7114CBC5DE2DF991FBA4B64B24155607" bold="true" pageId="0" pageNumber="11">23</emphasis>
hypopygium, lateral view
<emphasis bold="true" pageId="0" pageNumber="11">24</emphasis>
<emphasis id="FFEB9FC8054C5CF16E0A9B078AC453A8" bold="true" pageId="0" pageNumber="11">24</emphasis>
ninth tergite, dorsal view
<emphasis bold="true" pageId="0" pageNumber="11">25</emphasis>
<emphasis id="962B6BDC5D0CC8EA4C6E5B693B712B31" bold="true" pageId="0" pageNumber="11">25</emphasis>
eighth sternite, ventral view;
<emphasis bold="true" pageId="0" pageNumber="11">26</emphasis>
<emphasis id="6F88A744F95C6A37AE49B2081086FF16" bold="true" pageId="0" pageNumber="11">26</emphasis>
lobe of gonostylus, lateral view
<emphasis bold="true" pageId="0" pageNumber="11">27</emphasis>
<emphasis id="6F5ACFD4B3DE253F0474DE3620B2FC83" bold="true" pageId="0" pageNumber="11">27</emphasis>
clasper of gonostylus, lateral view. tg 8 = eighth tergite, tg 9 = ninth tergite, st 8 = eighth sternite, st 9 = ninth sternite, lg = lobe of gonostylus. Scale bars: 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="11">23-25</emphasis>
<emphasis id="17043B9E22F872E372503893A3882F45" bold="true" pageId="0" pageNumber="11">23-25</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="11">26, 27</emphasis>
<emphasis id="DE92BACDFD7CC558C3B99AB06F1A82B2" bold="true" pageId="0" pageNumber="11">26, 27</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="distribution">
<paragraph pageId="0" pageNumber="11">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="11">Laos (Champasak).</paragraph>
<subSubSection id="5192C405850C6E10387D7621F05327C7" pageId="0" pageNumber="11" type="distribution">
<paragraph id="C9822BB3FA814E04093AE1A33CFFACEA" pageId="0" pageNumber="11">Distribution.</paragraph>
<paragraph id="BE463523710AB1FF5CCD553A03CE7009" pageId="0" pageNumber="11">Laos (Champasak).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="etymology">
<paragraph pageId="0" pageNumber="11">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="11">The species is named after Champasak Province, where the type locality is located.</paragraph>
<subSubSection id="6C89224C8DC69CA860BA4863ABADB7FA" pageId="0" pageNumber="11" type="etymology">
<paragraph id="00FF45A3E2F7686E5A3F67C2E950AE61" pageId="0" pageNumber="11">Etymology.</paragraph>
<paragraph id="8EDCFF6B9F3B1B9BD0F4A1ED6ED6A77F" pageId="0" pageNumber="11">The species is named after Champasak Province, where the type locality is located.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="11" type="remarks">
<paragraph pageId="0" pageNumber="11">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="11">
<subSubSection id="D70AFE4F3A48012D6727DD2F1EB9A1A1" pageId="0" pageNumber="11" type="remarks">
<paragraph id="4CE0913FA216193B8F56C270589FAA20" pageId="0" pageNumber="11">Remarks.</paragraph>
<paragraph id="D474D6D62C99BBFF117DAB9C8BA4E22A" pageId="0" pageNumber="11">
This new species is somewhat similar to
<taxonomicName lsidName="T. (N.) coquilletti" pageId="0" pageNumber="11" rank="species" species="coquilletti" subGenus="N.">T. (N.) coquilletti</taxonomicName>
<taxonomicName id="6DB91CCE3B4C5AD13E1A6E30373731BD" lsidName="T. (N.) coquilletti" pageId="0" pageNumber="11" rank="species" species="coquilletti" subGenus="N.">T. (N.) coquilletti</taxonomicName>
Enderlein, 1912 from Japan and
<taxonomicName lsidName="T. (N.) sinica" pageId="0" pageNumber="11" rank="species" species="sinica" subGenus="N.">T. (N.) sinica</taxonomicName>
<taxonomicName id="01F17435AEA42D18BBC73A290C86ED04" lsidName="T. (N.) sinica" pageId="0" pageNumber="11" rank="species" species="sinica" subGenus="N.">T. (N.) sinica</taxonomicName>
Alexander, 1935 from China (Zhejiang) in having a similarly shaped hypopygium, but it can be separated from these species by the shape of the eighth sternite and ninth tergite, posterior margin of eighth sternite with V-shaped notch which with two long triangular processes at bottom, and posterior margin of ninth tergite with a small spiny protrusion at middle. In
<taxonomicName lsidName="T. (N.) coquilletti" pageId="0" pageNumber="11" rank="species" species="coquilletti" subGenus="N.">T. (N.) coquilletti</taxonomicName>
<taxonomicName id="306E0A430F85691887E6EE1869C06E52" lsidName="T. (N.) coquilletti" pageId="0" pageNumber="11" rank="species" species="coquilletti" subGenus="N.">T. (N.) coquilletti</taxonomicName>
, the posterior margin of eighth sternite is without processes and V -shaped depression, and the posterior margin of the ninth tergite has a V-shaped notch. In
<taxonomicName lsidName="T. (N.) sinica" pageId="0" pageNumber="11" rank="species" species="sinica" subGenus="N.">T. (N.) sinica</taxonomicName>
<taxonomicName id="0FBBF6A66F18AC415A594FA702B996F0" lsidName="T. (N.) sinica" pageId="0" pageNumber="11" rank="species" species="sinica" subGenus="N.">T. (N.) sinica</taxonomicName>
, the eighth sternite is without processes and V-shaped depression, and the posterior margin of the ninth tergite is extended with a sclerotized U-shaped depression.
</paragraph>
<caption doi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11" start="Figures 2833" startId="F8">
<paragraph pageId="0" pageNumber="11">
<emphasis bold="true" pageId="0" pageNumber="11">Figures 28-33.</emphasis>
<taxonomicName authorityName="Zhang &amp; Ren &amp; Li &amp; Yang" authorityYear="2020" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Nippotipula) champasakensis" order="Diptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="champasakensis" subGenus="Nippotipula">Tipula (Nippotipula) champasakensis</taxonomicName>
<caption id="3DBD42B1D1125DB9DF4A3E0A5899595B" doi="10.3897/zookeys.982.49781.figures28-33" httpUri="https://binary.pensoft.net/fig/470384" pageId="0" pageNumber="11" start="Figures 2833" startId="F8">
<paragraph id="DD85EBDCF3079D5FE92603C49D532E29" pageId="0" pageNumber="11">
<emphasis id="7D81D3816A8C69980C7E87CC67C223B5" bold="true" pageId="0" pageNumber="11">Figures 28-33.</emphasis>
<taxonomicName id="0C9E92E9B3F994498125A98253BBF625" authorityName="Zhang &amp; Ren &amp; Li &amp; Yang" authorityYear="2020" class="Insecta" family="Tipulidae" genus="Tipula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tipula (Nippotipula) champasakensis" order="Diptera" pageId="0" pageNumber="11" phylum="Arthropoda" rank="species" species="champasakensis" subGenus="Nippotipula">Tipula (Nippotipula) champasakensis</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="11">28</emphasis>
<emphasis id="862D0C0FBC5C9663854EC747C73E52CE" bold="true" pageId="0" pageNumber="11">28</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="11">30</emphasis>
<emphasis id="1B50AE4151CC39D92E840E1FE39FA4AE" bold="true" pageId="0" pageNumber="11">30</emphasis>
male
<emphasis bold="true" pageId="0" pageNumber="11">28</emphasis>
<emphasis id="8B5189407E85AA146D2C6CC44809DE0E" bold="true" pageId="0" pageNumber="11">28</emphasis>
semen pump, ventral view
<emphasis bold="true" pageId="0" pageNumber="11">29</emphasis>
<emphasis id="E3F7A0912F2E962C2686A5A28E70084F" bold="true" pageId="0" pageNumber="11">29</emphasis>
semen pump, dorsal view
<emphasis bold="true" pageId="0" pageNumber="11">30</emphasis>
<emphasis id="4F8B9C2E42E43CBA6F5FD5737BB3BE71" bold="true" pageId="0" pageNumber="11">30</emphasis>
semen pump, lateral view
<emphasis bold="true" pageId="0" pageNumber="11">31-33</emphasis>
<emphasis id="224C2A426AEC66B1034579971E6C85C8" bold="true" pageId="0" pageNumber="11">31-33</emphasis>
female
<emphasis bold="true" pageId="0" pageNumber="11">31</emphasis>
<emphasis id="DDDB007BCF47FD33A2BDFFAC6C50D889" bold="true" pageId="0" pageNumber="11">31</emphasis>
ovipositor, lateral view
<emphasis bold="true" pageId="0" pageNumber="11">32</emphasis>
<emphasis id="FF7292D3A2E0553D6FDC128A1CA19831" bold="true" pageId="0" pageNumber="11">32</emphasis>
ovipositor, dorsal view
<emphasis bold="true" pageId="0" pageNumber="11">33</emphasis>
<emphasis id="7CE8942E938F143323EE7AFB136CA9EE" bold="true" pageId="0" pageNumber="11">33</emphasis>
ovipositor, ventral view. tg 8 = eighth tergite, tg 9 = ninth tergite, tg 10 = tenth tergite, st 8 = eighth sternite, st 9 = ninth sternite, aia = anterior immovable apodeme, pia = posterior immovable apodeme, ca = compressor apodeme, ce = cercus, hy = hypovalva. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="11">28-30</emphasis>
<emphasis id="8BD1DB3BAB31209CE40F14D6AE87D379" bold="true" pageId="0" pageNumber="11">28-30</emphasis>
); 1.0 mm (
<emphasis bold="true" pageId="0" pageNumber="11">31-33</emphasis>
<emphasis id="AD465461C87B6AAC8DEF0B62AA928B1F" bold="true" pageId="0" pageNumber="11">31-33</emphasis>
).
</paragraph>
</caption>

View file

@ -1,103 +1,103 @@
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<paragraph id="9761D8E01AF568F3C0A67D50908B18B1" pageId="6" pageNumber="41">
<taxonomicName id="39F0EB1A9FF0E0726ABB8936A9CB5065" LSID="http://species-id.net/wiki/Languropilus" authority="Pic, 1940" authorityName="Pic" authorityYear="1940" genus="Languropilus" lsidName="Languropilus" pageId="6" pageNumber="41" rank="genus">Languropilus Pic, 1940</taxonomicName>
Figs 15245162
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="41" type="type species">
<paragraph pageId="6" pageNumber="41">Type species:</paragraph>
<paragraph pageId="6" pageNumber="41">
<taxonomicName class="Insecta" family="Cleridae" genus="Languropilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Languropilus fortipes" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="fortipes">Languropilus fortipes</taxonomicName>
<subSubSection id="8F2AF56C30EA88CEEF82D948D5AEEE0A" pageId="6" pageNumber="41" type="type species">
<paragraph id="8E02AF7324AE035B2470045831D8468B" pageId="6" pageNumber="41">Type species:</paragraph>
<paragraph id="836A0BCC792B0BF33EDCBAB65FB1FB85" pageId="6" pageNumber="41">
<taxonomicName id="C53012AEFB3E83292D23E696018CBAFA" genus="Languropilus" lsidName="Languropilus fortipes" pageId="6" pageNumber="41" rank="species" species="fortipes">Languropilus fortipes</taxonomicName>
Pic, 1940.
<bibRefCitation author="Pic, M" journalOrPublisher="L'Echange, Rev. Linn." pageId="13" pageNumber="48" pagination="2 - 4" title="Diagnoses de Coleopteres exotiques." volume="56" year="1940 a">Pic 1940a</bibRefCitation>
<bibRefCitation id="D96C845D866971500CA63D6DA01E359E" author="Pic, M" journalOrPublisher="L'Echange, Rev. Linn." pageId="13" pageNumber="48" pagination="2 - 4" title="Diagnoses de Coleopteres exotiques." volume="56" year="1940 a">Pic 1940a</bibRefCitation>
: 3.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="41" type="distribution">
<paragraph pageId="6" pageNumber="41">Distribution:</paragraph>
<paragraph pageId="6" pageNumber="41">Aethiopian region (East Africa).</paragraph>
<subSubSection id="AB18BC77203A428EB34C74A34E6018F0" pageId="6" pageNumber="41" type="distribution">
<paragraph id="69AB3B50F9E1764ACE6D4AE2461D1339" pageId="6" pageNumber="41">Distribution:</paragraph>
<paragraph id="B8A382654D053EE9DC34B2E91241FE4B" pageId="6" pageNumber="41">Aethiopian region (East Africa).</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="41" type="material examined">
<paragraph pageId="6" pageNumber="41">Material examined:</paragraph>
<paragraph pageId="6" pageNumber="41">
<taxonomicName class="Insecta" family="Cleridae" genus="Languropilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Languropilus fortipes" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="fortipes">Languropilus fortipes</taxonomicName>
<subSubSection id="13FE30E248C2CEDE847CAD011BD28E23" pageId="6" pageNumber="41" type="material examined">
<paragraph id="8EAF8D3C83E0501916169C02AEA0205A" pageId="6" pageNumber="41">Material examined:</paragraph>
<paragraph id="5001E8DCA93CC2B82D8A22E5CAAA8967" pageId="6" pageNumber="41">
<taxonomicName id="74F48AE4536A401ECA4CD7F55739FCBB" genus="Languropilus" lsidName="Languropilus fortipes" pageId="6" pageNumber="41" rank="species" species="fortipes">Languropilus fortipes</taxonomicName>
(females), Tanzania, Shinyanga Prov., Serengeti Sopa L., 19.XI.93, LF., Heiss (RGCM).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="41" type="description">
<paragraph pageId="6" pageNumber="41">Description</paragraph>
<subSection pageId="6" pageNumber="41" type="head">
<paragraph pageId="6" pageNumber="41">Head:</paragraph>
<paragraph pageId="6" pageNumber="41">With weakly protruding eyes, only very slightly emarginate at antennal insertion; interocular space about two times one eye width; gular sutures long, converging, gular process broad; antennae short, A1 large, stout, almost two times longer than A2, A2=A3 or A2&gt;A3, A3-A8 filiform, antennomeres becoming shorter, A8 almost spherical, A9 and A10 transverse, A11 ovate, terminal three antennomeres forming a distinct club.</paragraph>
<subSubSection id="3A840F044C845F5057605A8E8CA66CAA" pageId="6" pageNumber="41" type="description">
<paragraph id="95F06564A732B7B06C77986EA9E1FF31" pageId="6" pageNumber="41">Description</paragraph>
<subSection id="9FEEE0A165E3BF044C824FA14AA27485" pageId="6" pageNumber="41" type="head">
<paragraph id="77D1FEDDA53D7FD866AE19F9F3509C80" pageId="6" pageNumber="41">Head:</paragraph>
<paragraph id="83ADDAEB7EBEE52F5EDF8F20A7A037DD" pageId="6" pageNumber="41">With weakly protruding eyes, only very slightly emarginate at antennal insertion; interocular space about two times one eye width; gular sutures long, converging, gular process broad; antennae short, A1 large, stout, almost two times longer than A2, A2=A3 or A2&gt;A3, A3-A8 filiform, antennomeres becoming shorter, A8 almost spherical, A9 and A10 transverse, A11 ovate, terminal three antennomeres forming a distinct club.</paragraph>
</subSection>
<subSection pageId="6" pageNumber="41" type="thorax">
<paragraph pageId="6" pageNumber="41">Thorax:</paragraph>
<paragraph pageId="6" pageNumber="41">Proepimeron medium-sized, more rounded than acute; anterior mesosternal process absent; metendosternite with normal furcal stalk length, stalk slender, base almost straight, furcal arms of more or less normal length, acute distally (Fig. 15). Elytra long, broadest behind middle, apices broadly rounded, elytral punctation arranged into ten striae.</paragraph>
<subSection id="325F72811891988B629E4A3AA0E5E247" pageId="6" pageNumber="41" type="thorax">
<paragraph id="A49D2E8D979DAF47CF75E0207914E240" pageId="6" pageNumber="41">Thorax:</paragraph>
<paragraph id="E1E55A90E8BEA4466FC113295F7AFBD9" pageId="6" pageNumber="41">Proepimeron medium-sized, more rounded than acute; anterior mesosternal process absent; metendosternite with normal furcal stalk length, stalk slender, base almost straight, furcal arms of more or less normal length, acute distally (Fig. 15). Elytra long, broadest behind middle, apices broadly rounded, elytral punctation arranged into ten striae.</paragraph>
</subSection>
<subSection pageId="6" pageNumber="41" type="legs">
<paragraph pageId="6" pageNumber="41">Legs:</paragraph>
<paragraph pageId="6" pageNumber="41">Of normal size, stout; tarsal pulvillar formula 4-4-3, tibial spur formula 1-2-2; tibiae without longitudinal carinae; claws simple.</paragraph>
<caption pageId="6" pageNumber="41">
<paragraph pageId="6" pageNumber="41">
<subSection id="7E73781A963C5AF5D2FDD18A2C2BA33D" pageId="6" pageNumber="41" type="legs">
<paragraph id="D71C9B501E6B28A3867CCB4B67232C17" pageId="6" pageNumber="41">Legs:</paragraph>
<paragraph id="48273D8B808E212D3396ACCE9818D2C0" pageId="6" pageNumber="41">Of normal size, stout; tarsal pulvillar formula 4-4-3, tibial spur formula 1-2-2; tibiae without longitudinal carinae; claws simple.</paragraph>
<caption id="04143309F4DD23F9E8504AED7403BC4C" pageId="6" pageNumber="41">
<paragraph id="6F5CB0B404CC664C0F6F4DA4BD5B4A68" pageId="6" pageNumber="41">
Figures 38-46. Aedeagi of 38
<taxonomicName class="Insecta" family="Cleridae" genus="Aphelochroa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aphelochroa" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">Aphelochroa</taxonomicName>
<taxonomicName id="26208EFC788077F1DD04265CEC0B6005" genus="Aphelochroa" lsidName="Aphelochroa" pageId="6" pageNumber="41" rank="genus">Aphelochroa</taxonomicName>
sp. 39
<taxonomicName class="Insecta" family="Cleridae" genus="Caridopus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Caridopus" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">Caridopus</taxonomicName>
<taxonomicName id="10CF36C4CE7BFF30346E5EBEF20E73E5" genus="Caridopus" lsidName="Caridopus" pageId="6" pageNumber="41" rank="genus">Caridopus</taxonomicName>
sp. 40
<taxonomicName class="Insecta" family="Cleridae" genus="Dozocolletus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Dozocolletus discophorus" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="discophorus">Dozocolletus discophorus</taxonomicName>
<taxonomicName id="2CBEE5623277A9234CCA8D22B1CD8DEB" genus="Dozocolletus" lsidName="Dozocolletus discophorus" pageId="6" pageNumber="41" rank="species" species="discophorus">Dozocolletus discophorus</taxonomicName>
41
<taxonomicName class="Insecta" family="Cleridae" genus="Gyponyx" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gyponyx" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">Gyponyx</taxonomicName>
<taxonomicName id="EDB934B8287A0B5298DD7B8BFB2B8C51" genus="Gyponyx" lsidName="Gyponyx" pageId="6" pageNumber="41" rank="genus">Gyponyx</taxonomicName>
sp. 42
<taxonomicName class="Insecta" family="Cleridae" genus="Neorthrius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Neorthrius" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">Neorthrius</taxonomicName>
<taxonomicName id="5D82145E611D73AFB76E4E3E24424D9F" genus="Neorthrius" lsidName="Neorthrius" pageId="6" pageNumber="41" rank="genus">Neorthrius</taxonomicName>
sp. 43
<taxonomicName class="Insecta" family="Cleridae" genus="Nonalatus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nonalatus brevis" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="brevis">Nonalatus brevis</taxonomicName>
<taxonomicName id="5E43A0DD6C08E3DF7115B6E892DB9C89" genus="Nonalatus" lsidName="Nonalatus brevis" pageId="6" pageNumber="41" rank="species" species="brevis">Nonalatus brevis</taxonomicName>
44
<taxonomicName class="Insecta" family="Cleridae" genus="Orthrius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Orthrius sepulcralis" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="sepulcralis">Orthrius sepulcralis</taxonomicName>
<taxonomicName id="B4FE86013D99B4D8087F815DCB93FA90" genus="Orthrius" lsidName="Orthrius sepulcralis" pageId="6" pageNumber="41" rank="species" species="sepulcralis">Orthrius sepulcralis</taxonomicName>
45
<taxonomicName class="Insecta" family="Cleridae" genus="Pseudoastigmus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pseudoastigmus pygidialis" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="pygidialis">Pseudoastigmus pygidialis</taxonomicName>
<taxonomicName id="C0E6D942BCD19FD4622406E2DBBBA687" genus="Pseudoastigmus" lsidName="Pseudoastigmus pygidialis" pageId="6" pageNumber="41" rank="species" species="pygidialis">Pseudoastigmus pygidialis</taxonomicName>
46
<taxonomicName class="Insecta" family="Cleridae" genus="Xenorthrius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xenorthrius simplex" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="simplex">Xenorthrius simplex</taxonomicName>
<taxonomicName id="04B1993CCE60299BC876551632E67BDB" genus="Xenorthrius" lsidName="Xenorthrius simplex" pageId="6" pageNumber="41" rank="species" species="simplex">Xenorthrius simplex</taxonomicName>
. Scale bars 1mm.
</paragraph>
</caption>

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@ -1,302 +1,302 @@
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<taxonomicName LSID="C9F8FF17-8AC7-5B21-A3CA-9EEF8A6AFD13" authority="J. Balfour-Browne, 1939" authorityName="J. Balfour-Browne" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">Copelatus bakewelli J. Balfour-Browne, 1939</taxonomicName>
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<paragraph id="5C28E2BF90BB5607CC316AE07745CF97" pageId="0" pageNumber="81">
<taxonomicName id="8BD107F2247CC998E8C0D9500DBD79AF" ID-CoL="Y346" LSID="C9F8FF17-8AC7-5B21-A3CA-9EEF8A6AFD13" authority="J. Balfour-Browne, 1939" authorityName="J. Balfour-Browne" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">Copelatus bakewelli J. Balfour-Browne, 1939</taxonomicName>
<figureCitation id="864D81569AD45C3AC0B18F21980D6F77" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Figures 1</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Copelatus bakewelli A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure11" httpUri="https://binary.pensoft.net/fig/356480" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure11">11</figureCitation>
<figureCitation id="BFB96806A4E10F37304571E37C48BCAD" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Copelatus bakewelli A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure11" httpUri="https://binary.pensoft.net/fig/356480" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure11">11</figureCitation>
,
<figureCitation captionStart="Figures 20, 21" captionStartId="F20" captionText="Figures 20, 21. Distribution of 20 Copelatus bakewelli 21 C. daemeli." figureDoi="10.3897/zookeys.889.39090.figures20-21" httpUri="https://binary.pensoft.net/fig/356489" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figures20-21">20</figureCitation>
<figureCitation id="2B852AAF2A44F4E28350785F847195EA" captionStart="Figures 20, 21" captionStartId="F20" captionText="Figures 20, 21. Distribution of 20 Copelatus bakewelli 21 C. daemeli." figureDoi="10.3897/zookeys.889.39090.figures20-21" httpUri="https://binary.pensoft.net/fig/356489" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figures20-21">20</figureCitation>
,
<figureCitation captionStart="Figure 30" captionStartId="F28" captionText="Figure 30. Habitat of Copelatus bakewelli, rocky pool, with rotten leaves near Gunlom Waterfall, Northern Territory. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure30" httpUri="https://binary.pensoft.net/fig/356497" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure30">30</figureCitation>
<figureCitation id="58C55217DAF7449D36FAB1BFD8FE68B4" captionStart="Figure 30" captionStartId="F28" captionText="Figure 30. Habitat of Copelatus bakewelli, rocky pool, with rotten leaves near Gunlom Waterfall, Northern Territory. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure30" httpUri="https://binary.pensoft.net/fig/356497" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure30">30</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="reference_group">
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">Copelatus bakewelli</taxonomicName>
<subSubSection id="7C6BFB4B8173A339BFD9211B3639307F" pageId="0" pageNumber="81" type="reference_group">
<paragraph id="4A512028CE9AC700AB80BE924AB15BB0" pageId="0" pageNumber="81">
<taxonomicName id="32BA7B1FC22FA876E3D40E9BE1564854" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">Copelatus bakewelli</taxonomicName>
J.
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" author="Balfour-Browne, J" journalOrPublisher="The Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="81" pagination="57 - 88" refId="B1" refString="Balfour-Browne, J, 1939. On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with descriptions of new species. The Transactions of the Royal Entomological Society of London 88: 57 - 88" title="On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with descriptions of new species." url="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" volume="88" year="1939">Balfour-Browne 1939</bibRefCitation>
<bibRefCitation id="0E8A2F7165B0086ACDAF71FD86AD8D0D" DOI="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" author="Balfour-Browne, J" journalOrPublisher="The Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="81" pagination="57 - 88" refId="B1" refString="Balfour-Browne, J, 1939. On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with descriptions of new species. The Transactions of the Royal Entomological Society of London 88: 57 - 88" title="On Copelatus Er. and Leiopterus Steph. (Col. Dytiscidae) with descriptions of new species." url="https://doi.org/10.1111/j.1365-2311.1939.tb00250.x" volume="88" year="1939">Balfour-Browne 1939</bibRefCitation>
: 86 (original description);
<bibRefCitation DOI="https://doi.org/10.1071/AJZS057" author="Watts, CHS" journalOrPublisher="Australian Journal of Zoology Supplement Series" pageId="0" pageNumber="81" pagination="1 - 166" refId="B36" refString="Watts, CHS, 1978. A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology Supplement Series 57: 1 - 166" title="A revision of the Australian Dytiscidae (Coleoptera)." url="https://doi.org/10.1071/AJZS057" volume="57" year="1978">Watts 1978</bibRefCitation>
<bibRefCitation id="7E003AD890CA2CEE7FB1F6DBC66D515D" DOI="https://doi.org/10.1071/AJZS057" author="Watts, CHS" journalOrPublisher="Australian Journal of Zoology Supplement Series" pageId="0" pageNumber="81" pagination="1 - 166" refId="B36" refString="Watts, CHS, 1978. A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology Supplement Series 57: 1 - 166" title="A revision of the Australian Dytiscidae (Coleoptera)." url="https://doi.org/10.1071/AJZS057" volume="57" year="1978">Watts 1978</bibRefCitation>
: 127 (description);
<bibRefCitation author="Watts, CHS" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="81" pagination="22 - 28" refId="B37" refString="Watts, CHS, 1985. A faunal assessment of Australian Hydradephaga. Proceedings of the Academy of Natural Sciences of Philadelphia 137: 22 - 28" title="A faunal assessment of Australian Hydradephaga." volume="137" year="1985">Watts 1985</bibRefCitation>
<bibRefCitation id="1E1F96026275B31B24776A34AA7DE848" author="Watts, CHS" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="81" pagination="22 - 28" refId="B37" refString="Watts, CHS, 1985. A faunal assessment of Australian Hydradephaga. Proceedings of the Academy of Natural Sciences of Philadelphia 137: 22 - 28" title="A faunal assessment of Australian Hydradephaga." volume="137" year="1985">Watts 1985</bibRefCitation>
: 26 (general distribution);
<bibRefCitation author="Watts, CHS" journalOrPublisher="Identification Ecology Guide" pageId="0" pageNumber="81" pagination="1 - 110" refId="B38" refString="Watts, CHS, 2002. Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia). Identification Ecology Guide 43: 1 - 110" title="Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia)." volume="43" year="2002">Watts 2002</bibRefCitation>
<bibRefCitation id="47310BED9E8DD24DCADDBA9C64D08914" author="Watts, CHS" journalOrPublisher="Identification Ecology Guide" pageId="0" pageNumber="81" pagination="1 - 110" refId="B38" refString="Watts, CHS, 2002. Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia). Identification Ecology Guide 43: 1 - 110" title="Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia)." volume="43" year="2002">Watts 2002</bibRefCitation>
: 42 (checklist);
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="81" publicationUrl="http://www.waterbeetles.eu" refId="B28" refString="Nilsson, AN, Hajek, J, 2019. .http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2019">
<bibRefCitation id="F4F8A29794E7222E8970F829E4575777" author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="81" publicationUrl="http://www.waterbeetles.eu" refId="B28" refString="Nilsson, AN, Hajek, J, 2019. .http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2019">
Nilsson and
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
<normalizedToken id="51820B8ED02574E0707E070BDACB7EC4" originalValue="Hájek">Hajek</normalizedToken>
2019
</bibRefCitation>
: 62 (catalogue).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type locality">
<paragraph pageId="0" pageNumber="81">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="81">&quot;North Australia&quot;.</paragraph>
<subSubSection id="5E372F1DB442DC45076ED6492CF359DE" pageId="0" pageNumber="81" type="type locality">
<paragraph id="D0AEEF61FD23FE72CD5A7AC94E7E2547" pageId="0" pageNumber="81">Type locality.</paragraph>
<paragraph id="505C6DE6E6247574C6054E4AD3B66E76" pageId="0" pageNumber="81">&quot;North Australia&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type material">
<paragraph pageId="0" pageNumber="81">Type material.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Holotype:</emphasis>
<subSubSection id="CEF72D9F239271F6D32A0FA325ED1F94" pageId="0" pageNumber="81" type="type material">
<paragraph id="AEEDBCEA6C5F2DB5DB0960E04EB8A888" pageId="0" pageNumber="81">Type material.</paragraph>
<paragraph id="AB5CD7F6C6C4B1C33E9F61DFB508FF80" pageId="0" pageNumber="81">
<emphasis id="2CDD34481F28A1F93A55B1C5EF692E90" bold="true" italics="true" pageId="0" pageNumber="81">Holotype:</emphasis>
Female,
<normalizedToken originalValue="“Holotype”">&quot;Holotype&quot;</normalizedToken>
<normalizedToken id="69E573910872D991C973D6F9195BBCA2" originalValue="“Holotype”">&quot;Holotype&quot;</normalizedToken>
[round label with red frame],
<normalizedToken originalValue="“Na”">&quot;Na&quot;</normalizedToken>
<normalizedToken id="6B52A6D06C98DE778F1AA03830E7B7AC" originalValue="“Na”">&quot;Na&quot;</normalizedToken>
[hw label],
<normalizedToken originalValue="“6756”">&quot;6756&quot;</normalizedToken>
<normalizedToken id="DEB0582F3517548212D510D14D3B61BB" originalValue="“6756”">&quot;6756&quot;</normalizedToken>
[printed label], &quot;
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">Copelatus bakewelli</taxonomicName>
<taxonomicName id="A90B21D66A1D040E65D2D8DCA59EE4E6" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">Copelatus bakewelli</taxonomicName>
B-B. Type&quot; [hw label] (NHMUK).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="materials_examined">
<paragraph pageId="0" pageNumber="81">Additional material studied (285 specimens).</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Western Australia:</emphasis>
<subSubSection id="B3111D4452C5BF1DD2D1665E5E216C22" pageId="0" pageNumber="81" type="materials_examined">
<paragraph id="715BED6E5702F7A4D30883B3560CD5F0" pageId="0" pageNumber="81">Additional material studied (285 specimens).</paragraph>
<paragraph id="226AC69D30BFB0200C9925183ABDEF95" pageId="0" pageNumber="81">
<emphasis id="3846852279267E849E9CAF63BC6A90FB" bold="true" italics="true" pageId="0" pageNumber="81">Western Australia:</emphasis>
2 exs., &quot;AUSTRALIA WA, Synnot Range 1/9/1969,
<geoCoordinate degrees="16" direction="south" minutes="25" orientation="latitude" precision="925" value="-16.416666">16°25'S</geoCoordinate>
<geoCoordinate id="5279F05A89C670FA83208A83E861CAC8" degrees="16" direction="south" minutes="25" orientation="latitude" precision="925" value="-16.416666">16°25'S</geoCoordinate>
,
<geoCoordinate degrees="125" direction="east" minutes="28" orientation="longitude" precision="925" value="125.46667">125°28'E</geoCoordinate>
<geoCoordinate id="4DF24C978FE98B65FE88B112F71F060F" degrees="125" direction="east" minutes="28" orientation="longitude" precision="925" value="125.46667">125°28'E</geoCoordinate>
, Giuliani D.D. leg.&quot; (WAM); 1 ex., &quot;AUSTRALIA/WA/Shire of Wyndham - East Kimberley, Gibb River Road, Drysdale River Crossing, 380 m, 14.6.1999, Hendrich leg. Loc. 9/109&quot; (CLH); 4 exs., &quot;AUSTRALIA/WA/ Shire of Wyndham-East Kimberley, Gibb River Road, King Edward River Crossing, 280 m, 15.6.1999, Hendrich leg./coll. Loc. 10/110&quot; (CLH); 77 exs., &quot;AUSTRALIA/WA/Shire of Wyndham-East Kimberley, Mitchell Plateau, Little Mertens Falls, 300 m, 15.6.1999, Hendrich leg./coll. Loc. 11/111b&quot; (CLH, ZSM); 8 exs., &quot;AUSTRALIA/WA/ East Kimberley, Mitchell Plateau, Little Mertens Falls, 300 m, 15.6.1999, Hendrich leg./coll. Loc. 11a/111a&quot; (NMW); 2 exs., &quot;AUSTRALIA/WA/Shire of Wyndham - East Kimberley, Mitchell Plateau, Port Warrender Road/Kalumburu Road, Lowya Creek, 290 m, 18.6.1999, Hendrich leg. Loc. 13/113&quot; (CLH); 1 ex., &quot;AUSTRALIA/WA/Shire of Wyndham - East Kimberley, Mitchell Plateau, Kalumburu Road, 25 km NW King Edward Homestead, N.N. Creek, 370 m, 18.6.1999, Hendrich leg. Loc. 114&quot; (CLH); 4 exs., &quot;AUSTRALIA/WA/Shire of Wyndham - East Kimberley, Gibb River Road, 10 km E Hann River, Snake Creek, 470 m, 19.6.1999, Hendrich leg. Loc. 16/116&quot; (CLH).
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Northern Territory</emphasis>
<emphasis id="0C7B992B42A4528483DAB5DABF50EA66" bold="true" italics="true" pageId="0" pageNumber="81">Northern Territory</emphasis>
: 1 ex., &quot;Australia: NT, Cullen River n. Cullen, at Stuart Hwy, 102 m, 23.VIII.2006, 14.02.052S 131.56.561E, L. &amp; E. Hendrich leg. (NT 13)&quot;, &quot;DNA Balke 2213&quot; [green printed label] (ZSM); 1 ex., &quot;Australia: NT, Kakadu Hwy, Harriet Creek at Hwy Cross., 153 m, 24.VIII.2006, 13.44.512S 131.54.012E, L. &amp; E. Hendrich leg. (NT 14)&quot;, &quot;DNA Balke 1614&quot; [green printed label] (ZSM); 2 exs., &quot;Australia: NT, Litchfield NP, Creek near Wanggi Falls, 191 m, 20.VIII.2006, 13.11.221S 130.43.327E, L. &amp; E. Hendrich leg. (NT 4)&quot;, &quot;DNA Balke 2190&quot;, &quot;DNA Balke 2191&quot; [green printed label] (ZSM); 2 exs., &quot;AUSTRALIA NT Kakadu NP Alligator R. Gungaree Rainforest Dec. 22/93 S&amp;J Peck&quot; (ZSM); 1 ex., &quot;NT, Horse Creek fossil site, Camfield Station [
<geoCoordinate degrees="17.1166" direction="south" orientation="latitude" precision="5" value="-17.1166">-17.1166</geoCoordinate>
<geoCoordinate degrees="131.5166" direction="east" orientation="longitude" precision="5" value="131.5166">131.5166</geoCoordinate>
<geoCoordinate id="1E309D1686FF991024275C63450747A9" degrees="17.1166" direction="south" orientation="latitude" precision="5" value="-17.1166">-17.1166</geoCoordinate>
<geoCoordinate id="2EBB0BFAA8455B40B063A9CE3563DE97" degrees="131.5166" direction="east" orientation="longitude" precision="5" value="131.5166">131.5166</geoCoordinate>
] 2/8/1992&quot;,
<normalizedToken originalValue="“1004435”">&quot;1004435&quot;</normalizedToken>
<normalizedToken id="DE206CC5CEB6DCCDEC8E878EE2FF9BBD" originalValue="“1004435”">&quot;1004435&quot;</normalizedToken>
(MAGNT); 1 ex., &quot;NT, Top Humbert Yard, Humbert River [
<geoCoordinate degrees="16.4333" direction="south" orientation="latitude" precision="5" value="-16.4333">-16.4333</geoCoordinate>
<geoCoordinate id="1778E72D5FAD38446E0EA846D471EA36" degrees="16.4333" direction="south" orientation="latitude" precision="5" value="-16.4333">-16.4333</geoCoordinate>
,
<geoCoordinate degrees="130.4666" direction="east" orientation="longitude" precision="5" value="130.4666">130.4666</geoCoordinate>
<geoCoordinate id="3FA4078F0DC2E0E680EF21B2AD535012" degrees="130.4666" direction="east" orientation="longitude" precision="5" value="130.4666">130.4666</geoCoordinate>
] 7/1986&quot;,
<normalizedToken originalValue="“1004443”">&quot;1004443&quot;</normalizedToken>
<normalizedToken id="ACD60A93DDF6AD62B6F5C3C37B7671C3" originalValue="“1004443”">&quot;1004443&quot;</normalizedToken>
(MAGNT); 2 exs., &quot;Bathurst Is. + Melville Is. N.T. XII. 1975 D. Curry&quot;, &quot;QDPC 0-172712&quot;, &quot;QDPC 0-172713&quot; (QDPC); 2 exs., &quot;Australia, NT, Litchfield NP, Florence Falls,
<geoCoordinate degrees="12" direction="south" minutes="51" orientation="latitude" precision="15" seconds="15" value="-12.854167">12°51'15&quot;S</geoCoordinate>
<geoCoordinate id="396EC8E2AA53A3890ABA30CDDDD1E12D" degrees="12" direction="south" minutes="51" orientation="latitude" precision="15" seconds="15" value="-12.854167">12°51'15&quot;S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="45" orientation="longitude" precision="15" seconds="16" value="132.75444">132°45'16&quot;E</geoCoordinate>
<geoCoordinate id="7E55493D1C754CE650C24867387C4092" degrees="132" direction="east" minutes="45" orientation="longitude" precision="15" seconds="16" value="132.75444">132°45'16&quot;E</geoCoordinate>
, 63 m, at light, 12.V.2006, leg. Berger &amp; Dostal (5/06)&quot; (CGW); 8 exs., &quot;Australia, NT, 80 km W Roper Bar,
<geoCoordinate degrees="14" direction="south" minutes="54" orientation="latitude" precision="15" seconds="19" value="-14.905277">14°54'19&quot;S</geoCoordinate>
<geoCoordinate id="F9B9C7226341A886CA8EF03D8750F796" degrees="14" direction="south" minutes="54" orientation="latitude" precision="15" seconds="19" value="-14.905277">14°54'19&quot;S</geoCoordinate>
,
<geoCoordinate degrees="133" direction="east" minutes="57" orientation="longitude" precision="15" seconds="28" value="133.95778">133°57'28&quot;E</geoCoordinate>
<geoCoordinate id="4E6219686F95E99DE4BE319B7278AB9C" degrees="133" direction="east" minutes="57" orientation="longitude" precision="15" seconds="28" value="133.95778">133°57'28&quot;E</geoCoordinate>
, 78 m, at light, 14.V.2006, leg. Berger &amp; Dostal (7/06)&quot; (CGW); 1 ex., &quot;Australia, N.T./Katherine Gorge, Butterfly Gorge Walk, 150 m, 4.7.1999, Hendrich leg. Loc. 33/133&quot; (CLH); 3 exs., &quot;Australia, N.T. Kakadu N.P., Mary River District, 3 km ESE Gunlom Camping Area. South Alligator River, 50 m, 2.11.1996,
<geoCoordinate degrees="13" direction="south" minutes="27.276" orientation="latitude" precision="1" value="-13.4546">13°27.276'S</geoCoordinate>
<geoCoordinate id="6DDF9C32C8D9F169357A17093B2FE396" degrees="13" direction="south" minutes="27.276" orientation="latitude" precision="1" value="-13.4546">13°27.276'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="26.268" orientation="longitude" precision="1" value="132.4378">132°26.268'E</geoCoordinate>
<geoCoordinate id="5B25963DB303505730E74E9E9882466F" degrees="132" direction="east" minutes="26.268" orientation="longitude" precision="1" value="132.4378">132°26.268'E</geoCoordinate>
L. Hendrich leg./Lok. 14&quot; (CLH); 3 exs., &quot;Australia, N.T./Kakadu N.P., Jim Jim District, Gungurul Lookout, 50 m, 1.11.1996,
<geoCoordinate degrees="13" direction="south" minutes="59.359" orientation="latitude" precision="1" value="-13.989317">13°59.359'S</geoCoordinate>
<geoCoordinate id="3779CA25CBAC0D83F624FC93DA83121D" degrees="13" direction="south" minutes="59.359" orientation="latitude" precision="1" value="-13.989317">13°59.359'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="19.904" orientation="longitude" precision="1" value="132.33173">132°19.904'E</geoCoordinate>
<geoCoordinate id="40A51B83B1FBD10609F1098A12D38472" degrees="132" direction="east" minutes="19.904" orientation="longitude" precision="1" value="132.33173">132°19.904'E</geoCoordinate>
L. Hendrich leg./Lok. 11&quot; (CLH); 19 exs., &quot;Australia, N.T./Kakadu N.P., Mary River District, Gunlom Camping Area, 50 m, 3.11.1996,
<geoCoordinate degrees="13" direction="south" minutes="26.082" orientation="latitude" precision="1" value="-13.4347">13°26.082'S</geoCoordinate>
<geoCoordinate id="85C590ED357770B696596D492DBA45EE" degrees="13" direction="south" minutes="26.082" orientation="latitude" precision="1" value="-13.4347">13°26.082'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="24.929" orientation="longitude" precision="1" value="132.41548">132°24.929'E</geoCoordinate>
<geoCoordinate id="BE20454C381F6A4E910F93E65B36E61F" degrees="132" direction="east" minutes="24.929" orientation="longitude" precision="1" value="132.41548">132°24.929'E</geoCoordinate>
L. Hendrich leg./Lok. 15&quot; (CLH); 2 exs., &quot;Australia, N.T./Kakadu N.P., Jim Jim District, Barramundie Gorge, Maguk, 50 m, 31.10.1996,
<geoCoordinate degrees="13" direction="south" minutes="18.823" orientation="latitude" precision="1" value="-13.313717">13°18.823'S</geoCoordinate>
<geoCoordinate id="DBD90C05E84A8914FC50008FE3C42051" degrees="13" direction="south" minutes="18.823" orientation="latitude" precision="1" value="-13.313717">13°18.823'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="26.198" orientation="longitude" precision="1" value="132.43663">132°26.198'E</geoCoordinate>
<geoCoordinate id="AA73A4EB3726D0815082BE7E88C5EEA9" degrees="132" direction="east" minutes="26.198" orientation="longitude" precision="1" value="132.43663">132°26.198'E</geoCoordinate>
. L. Hendrich leg./ Lok. 9&quot; (CLH); 12 exs., &quot;Australia, NT/Kakadu N.P., Mary River District, Gunlom Waterfall Creek, 150 m, 2.11.1996,
<geoCoordinate degrees="13" direction="south" minutes="26.082" orientation="latitude" precision="1" value="-13.4347">13°26.082'S</geoCoordinate>
<geoCoordinate id="07755DF978F5C879FED305C1DCB668E6" degrees="13" direction="south" minutes="26.082" orientation="latitude" precision="1" value="-13.4347">13°26.082'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="24.929" orientation="longitude" precision="1" value="132.41548">132°24.929'E</geoCoordinate>
<geoCoordinate id="099DEA47EC437316E799B6D393B30B54" degrees="132" direction="east" minutes="24.929" orientation="longitude" precision="1" value="132.41548">132°24.929'E</geoCoordinate>
L. Hendrich leg./Lok. 13&quot; (CLH); 51 exs., &quot;Australia, N.T./Kakadu N.P., Jim Jim District, 1 km S Jim Jim Falls, 70 m, 26.10.1996,
<geoCoordinate degrees="13" direction="south" minutes="16.718" orientation="latitude" precision="1" value="-13.278633">13°16.718'S</geoCoordinate>
<geoCoordinate id="B97F06BF7DBB871C9145F54BA8EF49C1" degrees="13" direction="south" minutes="16.718" orientation="latitude" precision="1" value="-13.278633">13°16.718'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="49.490" orientation="longitude" precision="1" value="132.82483">132°49.490'E</geoCoordinate>
<geoCoordinate id="D4703E7C7E610590E70937125A18417E" degrees="132" direction="east" minutes="49.490" orientation="longitude" precision="1" value="132.82483">132°49.490'E</geoCoordinate>
L. Hendrich leg./Lok. 3&quot; (CLH); 39 exs., &quot;Australia, N.T./Kakadu N.P., Jim Jim District, Jim Jim Falls Camping Area, Jim Jim Creek, 60 m, 26. &amp; 27.10.1996
<geoCoordinate degrees="13" direction="south" minutes="16.218" orientation="latitude" precision="1" value="-13.2703">13°16.218'S</geoCoordinate>
<geoCoordinate id="7123C1E32A46E40D489992D265CFF4AB" degrees="13" direction="south" minutes="16.218" orientation="latitude" precision="1" value="-13.2703">13°16.218'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="49.276" orientation="longitude" precision="1" value="132.82127">132°49.276'E</geoCoordinate>
<geoCoordinate id="435DCCD2CE1050D5BEADF5B1AEBFB866" degrees="132" direction="east" minutes="49.276" orientation="longitude" precision="1" value="132.82127">132°49.276'E</geoCoordinate>
L. Hendrich leg./ Lok. 2.&quot; (CLH); 13 exs., &quot;Australia N.T./Kakadu N.P., Nourlangie District, Gubara, 50 m, 25.10.1996,
<geoCoordinate degrees="12" direction="south" minutes="50.101" orientation="latitude" precision="1" value="-12.835016">12°50.101'S</geoCoordinate>
<geoCoordinate id="AA93898711DA08AB3A1C2D7160DF669F" degrees="12" direction="south" minutes="50.101" orientation="latitude" precision="1" value="-12.835016">12°50.101'S</geoCoordinate>
,
<geoCoordinate degrees="132" direction="east" minutes="52.501" orientation="longitude" precision="1" value="132.87502">132°52.501'E</geoCoordinate>
<geoCoordinate id="AD6CB70970485203DDB26190FAFB739A" degrees="132" direction="east" minutes="52.501" orientation="longitude" precision="1" value="132.87502">132°52.501'E</geoCoordinate>
L. Hendrich leg./Lok. 1&quot; (CLH); 1 ex., &quot;Australia, NT, Pine Creek, Roper Gulf (S)Waterfall Ck., 60 mi. E of Pine Ck. [-13.81667, 132.7], 7.08.1964, Carne, P.B. leg. (ANIC); 1 ex., &quot;Australia, NT, Daly Basin, Katherine (T) Tindal,
<geoCoordinate degrees="14.51667" direction="south" orientation="latitude" precision="1" value="-14.51667">-14.51667</geoCoordinate>
<geoCoordinate id="90D6E6026D09B7A3FB22BA74914B7A53" degrees="14.51667" direction="south" orientation="latitude" precision="1" value="-14.51667">-14.51667</geoCoordinate>
,
<geoCoordinate degrees="132.3667" direction="east" orientation="longitude" precision="5" value="132.3667">132.3667</geoCoordinate>
<geoCoordinate id="B3CF5A8F09091B5E7740B317DCF9E142" degrees="132.3667" direction="east" orientation="longitude" precision="5" value="132.3667">132.3667</geoCoordinate>
, 1.12.1967, Vestjens, W.J.M. leg. (ANIC); 10 exs., &quot;Australia: NT, Litchfield NP, TJAYNERA FALLS,
<geoCoordinate degrees="13" direction="south" minutes="15" orientation="latitude" precision="925" value="-13.25">13°15'S</geoCoordinate>
<geoCoordinate id="826D9B3C28DB740F7534AB092A8D3E74" degrees="13" direction="south" minutes="15" orientation="latitude" precision="925" value="-13.25">13°15'S</geoCoordinate>
,
<geoCoordinate degrees="130" direction="east" minutes="44" orientation="longitude" precision="925" value="130.73334">130°44'E</geoCoordinate>
<geoCoordinate id="4C4FF1F50CE2EBB06619F7BA3C7FD321" degrees="130" direction="east" minutes="44" orientation="longitude" precision="925" value="130.73334">130°44'E</geoCoordinate>
, 63 m, S.
<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
<normalizedToken id="59C719CF5B37463716833FF3355C3A1E" originalValue="Jákl">Jakl</normalizedToken>
leg., 20
<normalizedToken originalValue="27.XI.2008”">-27.XI.2008&quot;</normalizedToken>
<normalizedToken id="D9EE72FE5311036CE4D92F83B0C564F8" originalValue="27.XI.2008”">-27.XI.2008&quot;</normalizedToken>
(NMPC).
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Queensland</emphasis>
<emphasis id="354F15C1BACF6D8D6D30B78CDF5C9AB9" bold="true" italics="true" pageId="0" pageNumber="81">Queensland</emphasis>
: 2 exs., &quot;QLD Cook, 2 km NNW Jowalbinna [-15.75, 144.25] 17/01/1994 Zborowski P. &amp; Edwards, E.D.&quot;,
<normalizedToken originalValue="“25-023624-470”">&quot;25-023624-470&quot;</normalizedToken>
<normalizedToken id="7C2570ACD83B7D7DA25EE34BE06BB6A9" originalValue="“25-023624-470”">&quot;25-023624-470&quot;</normalizedToken>
(ANIC); 1 ex., &quot;QLD Cook, 14 km ENE Heathlands [-11.68, 142.7], rainforest, 21/10/1993 Zborowski P. &amp; Rentz D.C.F.&quot;,
<normalizedToken originalValue="“25-022999-353”">&quot;25-022999-353&quot;</normalizedToken>
<normalizedToken id="C6076F4742B197A7F0B6A9C1EC24F332" originalValue="“25-022999-353”">&quot;25-022999-353&quot;</normalizedToken>
(ANIC); 1 ex., &quot;QLD, Townsville [
<geoCoordinate degrees="19.4833" direction="south" orientation="latitude" precision="5" value="-19.4833">-19.4833</geoCoordinate>
<geoCoordinate id="A6A1EA85534E137DCA1D6D379B031870" degrees="19.4833" direction="south" orientation="latitude" precision="5" value="-19.4833">-19.4833</geoCoordinate>
,
<geoCoordinate degrees="146.8167" direction="east" orientation="longitude" precision="5" value="146.8167">146.8167</geoCoordinate>
<geoCoordinate id="331CED66CA5059CB811D5F23231778A3" degrees="146.8167" direction="east" orientation="longitude" precision="5" value="146.8167">146.8167</geoCoordinate>
], 25.3.1996, CH Watts&quot;,
<normalizedToken originalValue="“25-001942”">&quot;25-001942&quot;</normalizedToken>
<normalizedToken id="34761B2F52F48E1361D6452180471E0D" originalValue="“25-001942”">&quot;25-001942&quot;</normalizedToken>
(SAMA); 1 ex., &quot;QLD, Townsville [-19.2, 146.68], 6-11/2/1998, AJ Watts&quot;,
<normalizedToken originalValue="“25-001940”">&quot;25-001940&quot;</normalizedToken>
<normalizedToken id="CE6C13FC419AE0D16FB679484D1E5A94" originalValue="“25-001940”">&quot;25-001940&quot;</normalizedToken>
(SAMA); 1 ex., &quot;Bamaga N. Qld. March 1984 J.W. Turner&quot;, &quot;QDPC 0-172711&quot; (QDPC); 3 exs., &quot;Bamaca [=Bamaga] xii.1983&quot;, &quot;Australian QLD J. Sedlacek&quot; (CLH); 1 ex., &quot;Australia, Queensland, Bald Hills Station 4 km N of Isabella Falls, 15°15' 145°00', 29. December 1984, mv lamp, G. &amp; A. Daniels&quot; (QM).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Description of male.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Body shape</emphasis>
<subSubSection id="B9BC7360BC863FCEA800F57EA6A49150" pageId="0" pageNumber="81" type="description">
<paragraph id="B58B5D6BCF69A0C8B107242ED39625FE" pageId="0" pageNumber="81">Description of male.</paragraph>
<paragraph id="212EB1B5033B5F6B111CBA1EC93C754F" pageId="0" pageNumber="81">
<emphasis id="1E402F13813412E4BC0C3AC10BA7E575" bold="true" pageId="0" pageNumber="81">
<emphasis id="E729F19732E421F8F17CE8E56680FE24" italics="true" pageId="0" pageNumber="81">Body shape</emphasis>
.
</emphasis>
In dorsal view, oblong oval, broadest at midlength of elytra, moderately convex. Body outline continuous, without discontinuity between pronotum and elytra. Head relatively broad; anterior margin of clypeus not bordered. Pronotum broadest between posterior angles, lateral margins moderately curved. Base of elytra as broad as pronotal base; lateral margins of elytra moderately curved (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Fig. 1</figureCitation>
<figureCitation id="EDB857451B3189AE62B24CF8320F2DE2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Fig. 1</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 1.</emphasis>
<caption id="86FE9F916AFECD1E8DB13CC07538062D" doi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" start="Figure 1" startId="F1">
<paragraph id="ECBE06524DEEF35C7C8D2AFC08D97E68" pageId="0" pageNumber="81">
<emphasis id="BA6DF0F3CE83136072D8069AE62A78FC" bold="true" pageId="0" pageNumber="81">Figure 1.</emphasis>
Habitus and colouration of
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus bakewelli</emphasis>
<taxonomicName id="B53BA6840ABE06475DFD1FD986E57DFE" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">
<emphasis id="C10B691FE87ABCE03FDCE09B1A248315" italics="true" pageId="0" pageNumber="81">Copelatus bakewelli</emphasis>
</taxonomicName>
, female. Total length 5.2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Colouration:</emphasis>
<paragraph id="D25E68B2831D114CAECBCBBA583FA2D3" pageId="0" pageNumber="81">
<emphasis id="2125CC0207FB0306C09BCB09799D7207" bold="true" italics="true" pageId="0" pageNumber="81">Colouration:</emphasis>
Body black, clypeus anteriorly, sides of pronotum, base of elytra, and humeral angle of elytra paler, ferruginous; ventral side reddish, appendages testaceous (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Fig. 1</figureCitation>
<figureCitation id="BC4D4D955D81D1ABBB0F8F6DAD2E2B6C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Fig. 1</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Dorsal surface sculpture:</emphasis>
<paragraph id="473FA7564635CE128624A3A8681019FA" pageId="0" pageNumber="81">
<emphasis id="88F9915194D300ECA73177DB2ADCD404" bold="true" italics="true" pageId="0" pageNumber="81">Dorsal surface sculpture:</emphasis>
Whole surface shiny (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Fig. 1</figureCitation>
<figureCitation id="39AF9A39411B66BC8348AFE4CB82E28E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Copelatus bakewelli, female. Total length 5.2 mm." figureDoi="10.3897/zookeys.889.39090.figure1" httpUri="https://binary.pensoft.net/fig/356470" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure1">Fig. 1</figureCitation>
). Head uniformly microreticulated, reticulation composed of moderately deeply impressed isodiametric meshes. Punctation composed of very small punctures, sparsely spread on surface; rows of deep and coarse punctures present around inner margin of eyes and in small depression anterolaterally of eyes. Pronotum with some moderately strong, short striae, more dense and weak laterally; lateral beading of pronotum very thin and indistinct. Microreticulation and punctation similar to that of head; row of coarse setigerous punctures present along anterior margin, basal margin (except for basomedially), and laterally close to sides. Elytra with microreticulation similar to that of head and pronotum, but less impressed. Punctation consisting of very fine sparse punctures. Each elytron with 11 strongly impressed discal and one submarginal longitudinal striae, intervals subequal, alternate striae tending to be shorter apically; subbasal stria reaching from middle of elytron almost to end of stria 10.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Antennae and legs:</emphasis>
<paragraph id="060434104B56F44CF504CFB7143B9DE2" pageId="0" pageNumber="81">
<emphasis id="08F7CD595DDFD9F5380645D1CBD2A7CD" bold="true" italics="true" pageId="0" pageNumber="81">Antennae and legs:</emphasis>
Antenna with antennomeres long and slender. Protibia straight, not modified. Pro- and mesotarsomeres 1-3 distinctly broadened, with adhesive discs on their ventral side; claws simple.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Ventral part:</emphasis>
<paragraph id="14B8AA97F2DB0D9D767206334123081E" pageId="0" pageNumber="81">
<emphasis id="3D03CFC38416CA5C6C53AD1FAD2343D9" bold="true" italics="true" pageId="0" pageNumber="81">Ventral part:</emphasis>
Finely microreticulated, with intermixed, sparsely distributed, very small punctures. Prosternum obtusely keeled medially. Prosternal process shortly lanceolate, apex obtuse; distinctly bordered laterally. Lateral parts of metaventrite tongue-shaped, very slender. Metacoxal lines almost complete and absent only close to metaventrite. Metacoxae with some short and deep striae, and abdominal ventrites 1-3 with numerous longitudinal striae.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Male genitalia:</emphasis>
<paragraph id="9BB37FA1BAC82315371D72484B16210C" pageId="0" pageNumber="81">
<emphasis id="CD18592F7F103086D612916C1E341BFC" bold="true" italics="true" pageId="0" pageNumber="81">Male genitalia:</emphasis>
Median lobe consisting of a few sclerites, closely attached together; apex in lateral view thin and elongate (
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Copelatus bakewelli A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure11" httpUri="https://binary.pensoft.net/fig/356480" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure11">Fig. 11A, B</figureCitation>
<figureCitation id="C34766766A6ED47D4849D79581C794C3" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Copelatus bakewelli A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure11" httpUri="https://binary.pensoft.net/fig/356480" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure11">Fig. 11A, B</figureCitation>
). Shape of paramere narrowly triangular, with thin, elongate subdistal part and weak but evident long setae along dorsal margin (
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Copelatus bakewelli A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure11" httpUri="https://binary.pensoft.net/fig/356480" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure11">Fig. 11D</figureCitation>
<figureCitation id="8AA4B78137289EE14597B104169B7ED3" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Copelatus bakewelli A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure11" httpUri="https://binary.pensoft.net/fig/356480" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure11">Fig. 11D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="female">
<paragraph pageId="0" pageNumber="81">Female.</paragraph>
<paragraph pageId="0" pageNumber="81">Similar to male in habitus. Pro- and mesotarsomeres not broadened, without adhesive setae. Pronotal striae finer and denser.</paragraph>
<subSubSection id="58F3EDA8BCE6903C87F01FF7C142A210" pageId="0" pageNumber="81" type="female">
<paragraph id="5EE9602E45D7E495BFB5C33B3D29C130" pageId="0" pageNumber="81">Female.</paragraph>
<paragraph id="1DBD0882CDC670D2D086885C404331E4" pageId="0" pageNumber="81">Similar to male in habitus. Pro- and mesotarsomeres not broadened, without adhesive setae. Pronotal striae finer and denser.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="81">TL = 5.1-5.25 mm; TL-H = 4.5-4.65 mm; MW = 2.45-2.50 mm.</paragraph>
<subSubSection id="32C4FE791BEB2962AF203580DD8CBB4C" pageId="0" pageNumber="81" type="description">
<paragraph id="E277FDDB9F680C3A3C0757A335A6EF83" pageId="0" pageNumber="81">Measurements.</paragraph>
<paragraph id="FCF2104277FD8546CC52DCABBBD2F1CD" pageId="0" pageNumber="81">TL = 5.1-5.25 mm; TL-H = 4.5-4.65 mm; MW = 2.45-2.50 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="variability">
<paragraph pageId="0" pageNumber="81">Variability.</paragraph>
<paragraph pageId="0" pageNumber="81">All specimens studied are rather uniform in shape and colour, and vary only little in body length.</paragraph>
<subSubSection id="C995DA94724ECEE6C6A85AE1C7B69F8A" pageId="0" pageNumber="81" type="variability">
<paragraph id="2C7E76D85D35B997B15A9DEB609ACF96" pageId="0" pageNumber="81">Variability.</paragraph>
<paragraph id="0EDE7C207E01C13E08A91ED653104B40" pageId="0" pageNumber="81">All specimens studied are rather uniform in shape and colour, and vary only little in body length.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="differential diagnosis">
<paragraph pageId="0" pageNumber="81">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus bakewelli</emphasis>
<subSubSection id="717977BB11B3F9B236B481E10E32BB0A" pageId="0" pageNumber="81" type="differential diagnosis">
<paragraph id="76BCA3F7B984277B1093120279F900CF" pageId="0" pageNumber="81">Differential diagnosis.</paragraph>
<paragraph id="20401A1A69494CEA4E9C7761F4486006" pageId="0" pageNumber="81">
<taxonomicName id="2959262C6F7398789B46813FF31EAD12" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus bakewelli" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bakewelli">
<emphasis id="71AF1662EE8F0EF48FDF934AE6B2643F" italics="true" pageId="0" pageNumber="81">Copelatus bakewelli</emphasis>
</taxonomicName>
is close to the form of
<taxonomicName lsidName="C. daemeli" pageId="0" pageNumber="81" rank="species" species="daemeli">
<emphasis italics="true" pageId="0" pageNumber="81">C. daemeli</emphasis>
<taxonomicName id="C173F2266AA73E01B50FE9E636D30443" lsidName="C. daemeli" pageId="0" pageNumber="81" rank="species" species="daemeli">
<emphasis id="C9002A3C1071BA9D7DC9E5C35D78B4CE" italics="true" pageId="0" pageNumber="81">C. daemeli</emphasis>
</taxonomicName>
with 11 elytral striae and
<taxonomicName lsidName="C. irregularis" pageId="0" pageNumber="81" rank="species" species="irregularis">
<emphasis italics="true" pageId="0" pageNumber="81">C. irregularis</emphasis>
<taxonomicName id="32314B9145FA97697F50E9B7D27178B5" lsidName="C. irregularis" pageId="0" pageNumber="81" rank="species" species="irregularis">
<emphasis id="82B5CE11C89EAEBFE0059ACC6F3852DD" italics="true" pageId="0" pageNumber="81">C. irregularis</emphasis>
</taxonomicName>
but smaller and without testaceous basal and apical markings on the elytra. Pronotal striae are weak (absent in
<taxonomicName lsidName="C. daemeli" pageId="0" pageNumber="81" rank="species" species="daemeli">
<emphasis italics="true" pageId="0" pageNumber="81">C. daemeli</emphasis>
<taxonomicName id="E40AC3062AAD2A6CD0253049AC659AC5" lsidName="C. daemeli" pageId="0" pageNumber="81" rank="species" species="daemeli">
<emphasis id="0AC96F296EB3D7C6D60DB0714AE4AD42" italics="true" pageId="0" pageNumber="81">C. daemeli</emphasis>
</taxonomicName>
) and the male protibia is not bent as in
<taxonomicName lsidName="C. irregularis" pageId="0" pageNumber="81" rank="species" species="irregularis">
<emphasis italics="true" pageId="0" pageNumber="81">C. irregularis</emphasis>
<taxonomicName id="2D02462AB6E59A4AEC2F56E479DF3C9F" lsidName="C. irregularis" pageId="0" pageNumber="81" rank="species" species="irregularis">
<emphasis id="628984C66B059DB91A9A28F9C3BAB60E" italics="true" pageId="0" pageNumber="81">C. irregularis</emphasis>
</taxonomicName>
. Furthermore, all three species can easily be separated by the form of their median lobes.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="81">
<subSubSection id="80DAD84445B97ADAE6067A9EFF79F9CA" pageId="0" pageNumber="81" type="distribution">
<paragraph id="B72A91081D01A9B9932CCD7A414468C9" pageId="0" pageNumber="81">Distribution.</paragraph>
<paragraph id="9BA0B0C1DE37747DA3DF437989926DAF" pageId="0" pageNumber="81">
Endemic. The species occurs from the Kimberley region in north-western Australia and in the Arnhemland in the Northern Territory to northern Queensland. In that area, the species is mainly distributed in rainforest pockets of the stone country (
<figureCitation captionStart="Figures 20, 21" captionStartId="F20" captionText="Figures 20, 21. Distribution of 20 Copelatus bakewelli 21 C. daemeli." figureDoi="10.3897/zookeys.889.39090.figures20-21" httpUri="https://binary.pensoft.net/fig/356489" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figures20-21">Fig. 20</figureCitation>
<figureCitation id="F96A36EB3126E0999C4C54ACD163B644" captionStart="Figures 20, 21" captionStartId="F20" captionText="Figures 20, 21. Distribution of 20 Copelatus bakewelli 21 C. daemeli." figureDoi="10.3897/zookeys.889.39090.figures20-21" httpUri="https://binary.pensoft.net/fig/356489" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figures20-21">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="habitat">
<paragraph pageId="0" pageNumber="81">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="81">
<subSubSection id="FB7BE22EA6CD2F56A9AE6447259123CC" pageId="0" pageNumber="81" type="habitat">
<paragraph id="5028546A53E979C6059FCCA2FD9F65AB" pageId="0" pageNumber="81">Habitat.</paragraph>
<paragraph id="705315D610A5BFB3D97D8DD921301F12" pageId="0" pageNumber="81">
A lentic species, which occurs in isolated pools of different size in otherwise dry riverbeds of seasonal streams, creeks and rivers. The bottom is consisted of gravel, sand and a layer of rotten plant debris. Occasionally, it occurs in protected bays, at the edge of large (25-50 m width), slow flowing and shallow (up to 20 cm depth) rivers, shaded by old riverine Melaleuca trees. There, the adults can be found among floating roots and organic debris of the paperbark trees. In northern Australia, at Gunlom waterfall, a large series was collected in a small (ca. 50 cm², 10 cm depth), water filled and shaded pothole (
<figureCitation captionStart="Figure 30" captionStartId="F28" captionText="Figure 30. Habitat of Copelatus bakewelli, rocky pool, with rotten leaves near Gunlom Waterfall, Northern Territory. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure30" httpUri="https://binary.pensoft.net/fig/356497" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure30">Fig. 30</figureCitation>
<figureCitation id="35E838EB4C988D6D9106FCCA64B275F2" captionStart="Figure 30" captionStartId="F28" captionText="Figure 30. Habitat of Copelatus bakewelli, rocky pool, with rotten leaves near Gunlom Waterfall, Northern Territory. Photographs by L. Hendrich." figureDoi="10.3897/zookeys.889.39090.figure30" httpUri="https://binary.pensoft.net/fig/356497" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure30">Fig. 30</figureCitation>
), filled up with rotten leaves. In Kakadu and the Kimberley region,
<taxonomicName lsidName="C. bakewelli" pageId="0" pageNumber="81" rank="species" species="bakewelli">
<emphasis italics="true" pageId="0" pageNumber="81">C. bakewelli</emphasis>
<taxonomicName id="A1B49135D86185C88373CC93E3134619" lsidName="C. bakewelli" pageId="0" pageNumber="81" rank="species" species="bakewelli">
<emphasis id="7944BC5DC73211EC2454A7488F576F7B" italics="true" pageId="0" pageNumber="81">C. bakewelli</emphasis>
</taxonomicName>
is more or less restricted to the stone country and the monsoon rainforest pockets. The species is also attracted to light.
</paragraph>