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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.177.2426" ID-GBIF-Dataset="3e03f4b2-ac3b-491e-ad1a-ea3124a4bddf" ID-PMC="PMC3317615" ID-Pensoft-Pub="1313-2970-177-15" ID-PubMed="22532783" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-177-15" ModsDocOrigin="ZooKeys 177" ModsDocTitle="Six new species of the genus Laena Dejean from China (Coleoptera, Tenebrionidae, Lagriinae)" checkinTime="1451249244682" checkinUser="pensoft" docAuthor="Xiao-Lin, Zhao &amp; Guo-Dong, Ren" docDate="2012" docId="C229662A2053D104FFE2CA0FAEB69BCC" docLanguage="en" docName="ZooKeys 177: 15-36" docOrigin="ZooKeys 177" docSource="http://dx.doi.org/10.3897/zookeys.177.2426" docTitle="Laena dentata Xiao-Lin &amp; Guo-Dong, 2012, sp. n." docType="treatment" docVersion="4" lastPageNumber="30" masterDocId="FFF0FF0BE902FFC25465FFC8107DFF8E" masterDocTitle="Six new species of the genus Laena Dejean from China (Coleoptera, Tenebrionidae, Lagriinae)" masterLastPageNumber="36" masterPageNumber="15" pageNumber="28" updateTime="1668153451917" updateUser="ExternalLinkService">
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<mods:title>Six new species of the genus Laena Dejean from China (Coleoptera, Tenebrionidae, Lagriinae)</mods:title>
<document id="09947C0E81AD81923B9287F88A67CB5B" ID-CLB-Dataset="35211" ID-DOI="http://dx.doi.org/10.3897/zookeys.177.2426" ID-GBIF-Dataset="3e03f4b2-ac3b-491e-ad1a-ea3124a4bddf" ID-PMC="PMC3317615" ID-Pensoft-Pub="1313-2970-177-15" ID-PubMed="22532783" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-177-15" ModsDocOrigin="ZooKeys 177" ModsDocTitle="Six new species of the genus Laena Dejean from China (Coleoptera, Tenebrionidae, Lagriinae)" checkinTime="1451249244682" checkinUser="pensoft" docAuthor="Xiao-Lin, Zhao &amp; Guo-Dong, Ren" docDate="2012" docId="C229662A2053D104FFE2CA0FAEB69BCC" docLanguage="en" docName="ZooKeys 177: 15-36" docOrigin="ZooKeys 177" docSource="http://dx.doi.org/10.3897/zookeys.177.2426" docTitle="Laena dentata Xiao-Lin &amp; Guo-Dong, 2012, sp. n." docType="treatment" docVersion="6" lastPageNumber="30" masterDocId="FFF0FF0BE902FFC25465FFC8107DFF8E" masterDocTitle="Six new species of the genus Laena Dejean from China (Coleoptera, Tenebrionidae, Lagriinae)" masterLastPageNumber="36" masterPageNumber="15" pageNumber="28" updateTime="1732753707853" updateUser="ExternalLinkService">
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<mods:title id="0B6CCA6F91888C3969109D6D9BA2F26B">Six new species of the genus Laena Dejean from China (Coleoptera, Tenebrionidae, Lagriinae)</mods:title>
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<mods:namePart>Xiao-Lin, Zhao</mods:namePart>
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<mods:namePart>Guo-Dong, Ren</mods:namePart>
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<treatment ID-GBIF-Taxon="152033618" LSID="urn:lsid:zoobank.org:act:D27A8819-4E08-41AE-AE46-C57D4913B28E" httpUri="http://treatment.plazi.org/id/C229662A2053D104FFE2CA0FAEB69BCC" lastPageId="15" lastPageNumber="30" pageId="13" pageNumber="28">
<subSubSection pageId="13" pageNumber="28" type="nomenclature">
<paragraph pageId="13" pageNumber="28">
<taxonomicName LSID="urn:lsid:zoobank.org:act:D27A8819-4E08-41AE-AE46-C57D4913B28E" class="Insecta" family="Tenebrionidae" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Laena dentata" order="Coleoptera" pageId="13" pageNumber="28" phylum="Arthropoda" rank="species" species="dentata">Laena dentata</taxonomicName>
<taxonomicNameLabel pageId="13" pageNumber="28">sp. n.</taxonomicNameLabel>
<treatment id="C229662A2053D104FFE2CA0FAEB69BCC" ID-GBIF-Taxon="152033618" LSID="urn:lsid:zoobank.org:act:D27A8819-4E08-41AE-AE46-C57D4913B28E" httpUri="http://treatment.plazi.org/id/C229662A2053D104FFE2CA0FAEB69BCC" lastPageId="15" lastPageNumber="30" pageId="13" pageNumber="28" scope_family="Tenebrionidae" scope_order="Coleoptera">
<subSubSection id="9F6F936730DC7008AAE6F4F77ACC5E5A" pageId="13" pageNumber="28" type="nomenclature">
<paragraph id="5EB9A4083EDCCC50F38F2FA2ABE12689" pageId="13" pageNumber="28">
<taxonomicName id="415FB2A1AC69E2BD5B23E71832FD85C6" ID-CoL="8JKZL" LSID="urn:lsid:zoobank.org:act:D27A8819-4E08-41AE-AE46-C57D4913B28E" genus="Laena" lsidName="Laena dentata" pageId="13" pageNumber="28" rank="species" species="dentata">Laena dentata</taxonomicName>
<taxonomicNameLabel id="8EFFFC4D43A79D1966CE4DB8BB4B0C49" pageId="13" pageNumber="28">sp. n.</taxonomicNameLabel>
Figs 432-40
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="28" type="type material">
<paragraph pageId="13" pageNumber="28">Type material.</paragraph>
<paragraph pageId="13" pageNumber="28">Holotype ♂ (MHUB): China, Yunnan, Dali, Cangshan E slope, 3400 m, 19 August 2008, J. S. Xu leg.</paragraph>
<paragraph pageId="13" pageNumber="28">Paratype: 1♀ (MHUB): labelled as the holotype.</paragraph>
<subSubSection id="3FEF25101EE69F3A652B051F0DCD37E9" pageId="13" pageNumber="28" type="type material">
<paragraph id="12BC5CE9E0FDCA66AB2E276E033247AE" pageId="13" pageNumber="28">Type material.</paragraph>
<paragraph id="F4FB0E6D98FF4C26F73EE5C3CBE504D7" pageId="13" pageNumber="28">Holotype ♂ (MHUB): China, Yunnan, Dali, Cangshan E slope, 3400 m, 19 August 2008, J. S. Xu leg.</paragraph>
<paragraph id="AF2BC9F8A2C947BEA33A0A15913D4CCC" pageId="13" pageNumber="28">Paratype: 1♀ (MHUB): labelled as the holotype.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="28" type="etymology">
<paragraph pageId="13" pageNumber="28">Etymology.</paragraph>
<paragraph pageId="13" pageNumber="28">Named after anterior tibiae of male with a medial tooth.</paragraph>
<subSubSection id="4C2849EDCE77CE796744F75413F299F1" pageId="13" pageNumber="28" type="etymology">
<paragraph id="889A42069B13C049AF09432B1048AB4C" pageId="13" pageNumber="28">Etymology.</paragraph>
<paragraph id="2F4BE6BEB97C51B70675764B3CE5A808" pageId="13" pageNumber="28">Named after anterior tibiae of male with a medial tooth.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="29" type="diagnosis">
<paragraph pageId="14" pageNumber="29">
<pageBreakToken pageId="14" pageNumber="29" start="start">Diagnosis</pageBreakToken>
<subSubSection id="878059C44C74342D62E0C054F96A5BBF" pageId="14" pageNumber="29" type="diagnosis">
<paragraph id="3A39688F9AF29D8D4DADAC4901AF68A4" pageId="14" pageNumber="29">
<pageBreakToken id="536A49FD299DAC24355B229BBAC18C7A" pageId="14" pageNumber="29" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="14" pageNumber="29">
<paragraph id="442D4809707EE3E96A1145CDEE8C5561" pageId="14" pageNumber="29">
The new species shares with
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Laena schusteri" order="Coleoptera" pageId="14" pageNumber="29" phylum="Arthropoda" rank="species" species="schusteri">Laena schusteri</taxonomicName>
<taxonomicName id="0AE08296A1E7C2188A94F6A50B3AE374" genus="Laena" lsidName="Laena schusteri" pageId="14" pageNumber="29" rank="species" species="schusteri">Laena schusteri</taxonomicName>
Schawaller, 2001 the body shape, and the medial tooth of anterior tibia, but can be separated by the teeth of all femora, and middle and posterior tibiae of male with finely hooked inner apex.
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="30" pageId="14" pageNumber="29" type="description">
<paragraph pageId="14" pageNumber="29">Description.</paragraph>
<paragraph pageId="14" pageNumber="29">
<subSubSection id="E638384A378EA41966BB0CF33B4C9443" lastPageId="15" lastPageNumber="30" pageId="14" pageNumber="29" type="description">
<paragraph id="385242833ADA7EC09C5191EFEB5CB519" pageId="14" pageNumber="29">Description.</paragraph>
<paragraph id="7EA7B15B7D6422E073678929929D9917" pageId="14" pageNumber="29">
Male. Eyes (Fig. 4) elliptical, moderately prominent. Antennae (Fig. 32) extending to base of pronotum, ratio of length (width) of antennomeres
<normalizedToken originalValue="II–XI">II-XI</normalizedToken>
<normalizedToken id="D526AD8DA27944C2F629C434EDA9C989" originalValue="II–XI">II-XI</normalizedToken>
as follows: 7.8 (10.8): 22.1 (11.5): 15.5 (12.8): 16.8 (12.0): 15.0 (13.9): 16.5 (13.9): 15.5 (13.0): 17.8 (13.1): 16.9 (14.2): 20.8 (13.9).
</paragraph>
<paragraph pageId="14" pageNumber="29">Pronotum (Fig. 4) elliptical, 1.3 times as wide as long, widest at middle; disc with small scattered punctures, their distance 0.5-3 times as long as puncture diameters, all punctures with short and adpressed setae, disc with a pair of feeble impressions, lateral margins narrowly bordered, basal margin unbordered, and not bent downwards, posterior angles rounded; propleura without punctures and setae.</paragraph>
<paragraph pageId="15" pageNumber="30">
<pageBreakToken pageId="15" pageNumber="30" start="start">Elytra</pageBreakToken>
<paragraph id="8404B4C336B40C32BF2A150AD30EB02A" pageId="14" pageNumber="29">Pronotum (Fig. 4) elliptical, 1.3 times as wide as long, widest at middle; disc with small scattered punctures, their distance 0.5-3 times as long as puncture diameters, all punctures with short and adpressed setae, disc with a pair of feeble impressions, lateral margins narrowly bordered, basal margin unbordered, and not bent downwards, posterior angles rounded; propleura without punctures and setae.</paragraph>
<paragraph id="316C3FB0BC21F792C81803EE46587BC8" pageId="15" pageNumber="30">
<pageBreakToken id="A2F3125B47F8BD8884B912CF8C6A7E92" pageId="15" pageNumber="30" start="start">Elytra</pageBreakToken>
(Fig. 4) nearly parallel-sided from base to middle, 1.9 times as long as wide, widest at middle; elytra with punctural rows of without striae, punctures distinctly larger than those of pronotum, each puncture bearing a short and adpressed seta, intervals with nearly invisible punctures, but with some similar setae, all intervals flat and shagreened, interval IX with 3 setiferous umbilicate pores.
</paragraph>
<paragraph pageId="15" pageNumber="30">All femora each with a tooth, but middle femur (Fig. 34) with a pair of equally sized teeth. Anterior tibia (Fig. 33) with median tooth and dilated apex, anterior and middle tibiae excavate medially and with caespitose setae, middle and posterior tibiae (Fig. 35) with finely hooked inner apex.</paragraph>
<paragraph pageId="15" pageNumber="30">Last abdominal ventrite (Fig. 36) somewhat sharp at apex. Aedeagus see Figs. 38-40.</paragraph>
<paragraph pageId="15" pageNumber="30">Female: Last abdominal ventrite (Fig. 37) somewhat rounded at apex. Middle femora with a pair of unequal teeth.</paragraph>
<paragraph pageId="15" pageNumber="30">Body length: 8.5-9.5 mm.</paragraph>
<caption pageId="15" pageNumber="30">
<paragraph pageId="15" pageNumber="30">
<paragraph id="658B189B04CB2E30A5C3728670C583BA" pageId="15" pageNumber="30">All femora each with a tooth, but middle femur (Fig. 34) with a pair of equally sized teeth. Anterior tibia (Fig. 33) with median tooth and dilated apex, anterior and middle tibiae excavate medially and with caespitose setae, middle and posterior tibiae (Fig. 35) with finely hooked inner apex.</paragraph>
<paragraph id="02C4494DD1F7EB6F3ABB5C9E3B7DDB46" pageId="15" pageNumber="30">Last abdominal ventrite (Fig. 36) somewhat sharp at apex. Aedeagus see Figs. 38-40.</paragraph>
<paragraph id="3F8373CF765E7E7EF3FEC4BB82EB6825" pageId="15" pageNumber="30">Female: Last abdominal ventrite (Fig. 37) somewhat rounded at apex. Middle femora with a pair of unequal teeth.</paragraph>
<paragraph id="B2AAA895365725C94745739613189D18" pageId="15" pageNumber="30">Body length: 8.5-9.5 mm.</paragraph>
<caption id="98B629CF01D01821FBA11890BFD6C56F" pageId="15" pageNumber="30">
<paragraph id="09521122AFA03A44E1E31ED036953698" pageId="15" pageNumber="30">
Figures 32-40.
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Laena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Laena dentata" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="dentata">Laena dentata</taxonomicName>
<taxonomicName id="F3D71773FB1B83B5BE7B530024E44786" genus="Laena" lsidName="Laena dentata" pageId="15" pageNumber="30" rank="species" species="dentata">Laena dentata</taxonomicName>
sp. n. 32 male antenna, dorsal view 33 anterior femur and tibia, male, ventral view 34 middle femur and tibia, male, ventral view 35 posterior femur and tibia, male, ventral view 36 last abdominal ventrite, male, ventral view 37 last abdominal ventrite, female, ventral view 38-40 aedeagus in dorsal, lateral and ventral views.
</paragraph>
</caption>