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<mods:title>Pyralis cardinalis, a charismatic new species related to P. regalis [Denis & Schiffermueller], 1775, first recognized in Finland (Lepidoptera, Pyralidae)</mods:title>
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<mods:title id="F78AE94CB2FA22BE372A191024911A3F">Pyralis cardinalis, a charismatic new species related to P. regalis [Denis & Schiffermueller], 1775, first recognized in Finland (Lepidoptera, Pyralidae)</mods:title>
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<mods:namePart>Wikstroem, Bo</mods:namePart>
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<mods:affiliation>Yli-Haakkointie 13, 03100 Nummela, Finland e-mail: bo. r. wikstrom @ gmail. com</mods:affiliation>
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<mods:namePart>Huemer, Peter</mods:namePart>
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<mods:namePart>Mutanen, Marko</mods:namePart>
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<mods:affiliation>Ecology and Genetics Research Unit, PO BOX 3000, FI- 90014 University of Oulu, Finland; e-mail: marko. mutanen @ oulu. fi</mods:affiliation>
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<mods:namePart>Tyllinen, Juha</mods:namePart>
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<mods:namePart>Kaila, Lauri</mods:namePart>
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<mods:affiliation>Finnish Museum of Natural History, Zoology Unit, University of Helsinki, Finland; e-mail: Lauri. Kaila @ Helsinki. fi (corresponding author)</mods:affiliation>
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<mods:affiliation id="F45B1E84C9D8FAFA2774AAF346F1BE77">Finnish Museum of Natural History, Zoology Unit, University of Helsinki, Finland; e-mail: Lauri. Kaila @ Helsinki. fi (corresponding author)</mods:affiliation>
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<treatment ID-GBIF-Taxon="170296335" LSID="urn:lsid:plazi:treatment:BDA8601C15E6525CB246D374EFDA8E7B" httpUri="http://treatment.plazi.org/id/BDA8601C15E6525CB246D374EFDA8E7B" lastPageNumber="337" pageId="0" pageNumber="337">
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<taxonomicName LSID="BDA8601C-15E6-525C-B246-D374EFDA8E7B" authority="Butler, 1889" authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">Pyralis princeps Butler, 1889</taxonomicName>
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<figureCitation captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Pyralis spp., habitus. 18. P. princeps Butler, ♂; Nepal, Godawari nr. Kathmandu, leg. exp. Albrecht et al. 19. P. joannisi Leraut, ♂; Vietnam, Bac Kan province, leg. A. Wikstroem. 20, 21. P. perversalis (Herrich-Schaeffer), habitus. 20. ♂, Czech Republic, Bohemia sept., leg. J. Liska; 21. ♀, S. Ural, Cheliabinsk district, leg. T. Nupponen." figureDoi="10.3897/nl.43.54916.figures18-21" httpUri="https://binary.pensoft.net/fig/473651" pageId="0" pageNumber="337">Figs 18</figureCitation>
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<figureCitation captionStart="Figures 25–27" captionStartId="F8" captionText="Figures 25 - 27. ♂ genitalia of Pyralis spp. 25. P. kacheticalis (Christoph), GPBW 7942, Greece, Lesvos, Anaxos. 26. P. princeps Butler, L. Kaila prep. 6296, Nepal, Godawari, 27. P. joannisi Leraut, L. Kaila prep. 6297, Vietnam, Bac Kan Prov." figureDoi="10.3897/nl.43.54916.figures25-27" httpUri="https://binary.pensoft.net/fig/473653" pageId="0" pageNumber="337">, 26</figureCitation>
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<treatment id="BDA8601C15E6525CB246D374EFDA8E7B" ID-GBIF-Taxon="170296335" LSID="urn:lsid:plazi:treatment:BDA8601C15E6525CB246D374EFDA8E7B" httpUri="http://treatment.plazi.org/id/BDA8601C15E6525CB246D374EFDA8E7B" lastPageNumber="337" pageId="0" pageNumber="337" scope_family="Pyralidae" scope_order="Lepidoptera">
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<subSubSection id="F1BBDD13D0BC2814356305F85E4A637D" pageId="0" pageNumber="337" type="nomenclature">
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<paragraph id="76DF3205BDD2E6F63C49C28A8E934562" pageId="0" pageNumber="337">
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<taxonomicName id="6EA0982BA61C544B43A6E750F01A5316" ID-CoL="986JP" LSID="BDA8601C-15E6-525C-B246-D374EFDA8E7B" authority="Butler, 1889" authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">Pyralis princeps Butler, 1889</taxonomicName>
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<figureCitation id="9C88E3F7D51E12A323713993ABFE9E08" captionStart="Figures 18–21" captionStartId="F6" captionText="Figures 18 - 21. Pyralis spp., habitus. 18. P. princeps Butler, ♂; Nepal, Godawari nr. Kathmandu, leg. exp. Albrecht et al. 19. P. joannisi Leraut, ♂; Vietnam, Bac Kan province, leg. A. Wikstroem. 20, 21. P. perversalis (Herrich-Schaeffer), habitus. 20. ♂, Czech Republic, Bohemia sept., leg. J. Liska; 21. ♀, S. Ural, Cheliabinsk district, leg. T. Nupponen." figureDoi="10.3897/nl.43.54916.figures18-21" httpUri="https://binary.pensoft.net/fig/473651" pageId="0" pageNumber="337">Figs 18</figureCitation>
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<figureCitation id="EC9F244A807B3DBB985583413FC9F511" captionStart="Figures 25–27" captionStartId="F8" captionText="Figures 25 - 27. ♂ genitalia of Pyralis spp. 25. P. kacheticalis (Christoph), GPBW 7942, Greece, Lesvos, Anaxos. 26. P. princeps Butler, L. Kaila prep. 6296, Nepal, Godawari, 27. P. joannisi Leraut, L. Kaila prep. 6297, Vietnam, Bac Kan Prov." figureDoi="10.3897/nl.43.54916.figures25-27" httpUri="https://binary.pensoft.net/fig/473653" pageId="0" pageNumber="337">, 26</figureCitation>
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</paragraph>
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<subSubSection pageId="0" pageNumber="337" type="reference_group">
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<paragraph pageId="0" pageNumber="337">
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<taxonomicName authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">Pyralis princeps</taxonomicName>
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<subSubSection id="9FCE0A32704BAFB66A098D6BC4D65C79" pageId="0" pageNumber="337" type="reference_group">
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<paragraph id="55C95D7EBD07C06E9DFDED69F3CA9992" pageId="0" pageNumber="337">
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<taxonomicName id="5E649C303DD5B0F772D2348156E543AF" authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">Pyralis princeps</taxonomicName>
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||||
Butler, 1889(7): 91, pl. 134, fig. 12.
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</paragraph>
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<subSubSection pageId="0" pageNumber="337" type="materials_examined">
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<paragraph pageId="0" pageNumber="337">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="337">
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<subSubSection id="C2BF1B67F9C75A083A228B970A83D508" pageId="0" pageNumber="337" type="materials_examined">
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<paragraph id="E1D63476D4E49EF26490AC83914BA484" pageId="0" pageNumber="337">Material examined.</paragraph>
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<paragraph id="884F02A5AA671280587500342F40416F" pageId="0" pageNumber="337">
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Type material.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="337">Lectotype</emphasis>
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<emphasis id="F173FC15872CF32C40EED999A8A9572E" bold="true" italics="true" pageId="0" pageNumber="337">Lectotype</emphasis>
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, here designated: India, Dharamshala, Photograph of
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<emphasis bold="true" italics="true" pageId="0" pageNumber="337">lectotype</emphasis>
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of
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<emphasis italics="true" pageId="0" pageNumber="337">Pyralis princeps</emphasis>
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<emphasis id="CEA7E5B1D781ABC5FD6FDA4D26B83DFE" italics="true" pageId="0" pageNumber="337">Pyralis princeps</emphasis>
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|
||||
made available by David Lees, labelled: LECTO-TYPE [blue round label]; MANUSCRIPT LECTOTYPE; Type locality India Dharmsala; Lectotype
|
||||
<taxonomicName authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">Pyralis princeps</emphasis>
|
||||
<taxonomicName id="DC70125ABDDD24926219C12C42041E6B" authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">
|
||||
<emphasis id="ED06420B453B63CE96C835B71EED4D3E" italics="true" pageId="0" pageNumber="337">Pyralis princeps</emphasis>
|
||||
</taxonomicName>
|
||||
det. M. Shaffer 1985[]; stat. n. [abdomen in capsule] (NHMUK).
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="337">
|
||||
<emphasis bold="true" pageId="0" pageNumber="337">Other material.</emphasis>
|
||||
<paragraph id="9309D8ABFE0D07D21215DAA6408D52D2" pageId="0" pageNumber="337">
|
||||
<emphasis id="1873EBC75C00E7AB9308A030532EBC18" bold="true" pageId="0" pageNumber="337">Other material.</emphasis>
|
||||
Nepal. 2♂,
|
||||
<geoCoordinate degrees="27" direction="north" minutes="40" orientation="latitude" precision="925" value="27.666666">27°40'N</geoCoordinate>
|
||||
<geoCoordinate id="7F1B68E29084B3FA546B5B51D9E3D396" degrees="27" direction="north" minutes="40" orientation="latitude" precision="925" value="27.666666">27°40'N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="85" direction="east" minutes="25" orientation="longitude" precision="925" value="85.416664">85°25'E</geoCoordinate>
|
||||
<geoCoordinate id="F7C904BC7907374859A631FDF9AD1E0A" degrees="85" direction="east" minutes="25" orientation="longitude" precision="925" value="85.416664">85°25'E</geoCoordinate>
|
||||
, Godavari, 15 km SE Kathmandu, 1500 m, 8.-9.v. 1996 leg. Exp. A. Albrecht, O.
|
||||
<normalizedToken originalValue="Biström">Bistroem</normalizedToken>
|
||||
<normalizedToken id="BC37912E2B78B698206CB071F519A06E" originalValue="Biström">Bistroem</normalizedToken>
|
||||
, K. Mikkola & A. Wikberg, L. Kaila prep. 6296, BOLD sample ID: TLMF Lep 05678, BOLD sample ID: TLMF Lep 05679, http://id.luomus.fi/HV.369 (MZH).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="337" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="337">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="337">
|
||||
<subSubSection id="3FBF5C333061A898F3626DA387A40B60" pageId="0" pageNumber="337" type="diagnosis">
|
||||
<paragraph id="53EB90F9D8C9BC1564872DB3219C0B32" pageId="0" pageNumber="337">Diagnosis.</paragraph>
|
||||
<paragraph id="8EF348B7DF96962054CCA5D12D7B9650" pageId="0" pageNumber="337">
|
||||
The separation of this species from other species is explained in the key. In the genitalia, the dense bush of long spines at the posterior end of vesica is characteristic, and the genital characters readily separate it from the externally similar but smaller
|
||||
<taxonomicName genus="P." lsidName="P. joannisi" pageId="0" pageNumber="337" rank="species" species="joannisi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. joannisi</emphasis>
|
||||
<taxonomicName id="66EA1A5CEB44ABD33C9133FF206D8311" lsidName="P. joannisi" pageId="0" pageNumber="337" rank="species" species="joannisi">
|
||||
<emphasis id="31773EC3A8B44A228074E4562926D364" italics="true" pageId="0" pageNumber="337">P. joannisi</emphasis>
|
||||
</taxonomicName>
|
||||
; see the key for further details.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="337" type="molecular data">
|
||||
<paragraph pageId="0" pageNumber="337">Molecular data.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="337">
|
||||
<subSubSection id="FE58E397028F8FB088DA219BBBE5D927" pageId="0" pageNumber="337" type="molecular data">
|
||||
<paragraph id="1AFFDD3219EFEC6E3803B26784EA2978" pageId="0" pageNumber="337">Molecular data.</paragraph>
|
||||
<paragraph id="1BCCE39BA9E4B9DA8C9772763EBA4EB6" pageId="0" pageNumber="337">
|
||||
BIN: replace: BOLD:ABA8505. The intraspecific mean and maximum divergence of the barcode region is 0% (
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">N</emphasis>
|
||||
<emphasis id="0E7FC8CCD4CBBBB063584F54A96A078F" italics="true" pageId="0" pageNumber="337">N</emphasis>
|
||||
= 2). The minimum distance to the nearest neighbour,
|
||||
<taxonomicName genus="P." lsidName="P. kacheticalis" pageId="0" pageNumber="337" rank="species" species="kacheticalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. kacheticalis</emphasis>
|
||||
<taxonomicName id="21747F8E23D1F371F6B477F832574CEC" lsidName="P. kacheticalis" pageId="0" pageNumber="337" rank="species" species="kacheticalis">
|
||||
<emphasis id="A71C0F1EF0821D3F2277659E902027E1" italics="true" pageId="0" pageNumber="337">P. kacheticalis</emphasis>
|
||||
</taxonomicName>
|
||||
, is 8.26%.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="337" type="distribution">
|
||||
<paragraph pageId="0" pageNumber="337">Distribution.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="337">India, Nepal.</paragraph>
|
||||
<subSubSection id="D465FE974FB1EE45703594DD548208DA" pageId="0" pageNumber="337" type="distribution">
|
||||
<paragraph id="18E5C0F552CD55B67E6D9BE4922074AB" pageId="0" pageNumber="337">Distribution.</paragraph>
|
||||
<paragraph id="27DCDAC2D5911896E81FFE22879B24DB" pageId="0" pageNumber="337">India, Nepal.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="337" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="337">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="337">
|
||||
<taxonomicName authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">Pyralis princeps</emphasis>
|
||||
<subSubSection id="EBEC6B237E3C53B8EE9BCEBA8BCFC875" pageId="0" pageNumber="337" type="remarks">
|
||||
<paragraph id="EAC6D34501D444BF9A0F47601C383ECD" pageId="0" pageNumber="337">Remarks.</paragraph>
|
||||
<paragraph id="951431DCB7552EF95C6EEBD8C59E3B09" pageId="0" pageNumber="337">
|
||||
<taxonomicName id="BD813430812CF25E5EAB664EE5923B42" authorityName="Butler" authorityYear="1889" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis princeps" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="species" species="princeps">
|
||||
<emphasis id="F4AF955255711446AD77149C10D6D756" italics="true" pageId="0" pageNumber="337">Pyralis princeps</emphasis>
|
||||
</taxonomicName>
|
||||
was described from four specimens collected in India (Himachal Pradesh, Dharamshala [Dharmsala]) and two from
|
||||
<normalizedToken originalValue="”Yezo”">"Yezo"</normalizedToken>
|
||||
<normalizedToken id="6B1B0EB495521D0974464E88C938E93D" originalValue="”Yezo”">"Yezo"</normalizedToken>
|
||||
[region from Northern Japan to Kamchatka] (Butler, 1889). However, the latter two are misidentifications of
|
||||
<taxonomicName genus="P." lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
|
||||
<taxonomicName id="7884EB2202C8D8566796BF8CA99F153C" lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
|
||||
<emphasis id="42145D94E90FCBA394437855C01F8855" italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
|
||||
</taxonomicName>
|
||||
, this interpretation also supported by our examination of some Japanese and South Korean specimens. Following
|
||||
<bibRefCitation author="ICZN" journalOrPublisher="The International Trust for Zoological Nomenclature, London" pageId="0" pageNumber="337" refId="B10" refString="ICZN, 1999. International Code of Zoological Nomenclature. Fourth edition. The International Trust for Zoological Nomenclature, London" title="International Code of Zoological Nomenclature. Fourth edition." year="1999">ICZN (1999)</bibRefCitation>
|
||||
<bibRefCitation id="00BF8836E4D56DE18FF2004D5E1856C7" author="ICZN" journalOrPublisher="The International Trust for Zoological Nomenclature, London" pageId="0" pageNumber="337" refId="B10" refString="ICZN, 1999. International Code of Zoological Nomenclature. Fourth edition. The International Trust for Zoological Nomenclature, London" title="International Code of Zoological Nomenclature. Fourth edition." year="1999">ICZN (1999)</bibRefCitation>
|
||||
, a lectotype from Dharamshala [Dharmsala], already labelled as such by M. Shaffer in NHMUK, is here designated in order to fix the identity of the species and conserve nomenclatural stability.We have examined photographs of the specimen and its labels which confirm our interpretation. Outside Europe, the name
|
||||
<taxonomicName genus="P." lsidName="P. princeps" pageId="0" pageNumber="337" rank="species" species="princeps">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. princeps</emphasis>
|
||||
<taxonomicName id="8DF95BC6C765A5DFCC0FA28E2EFA187C" lsidName="P. princeps" pageId="0" pageNumber="337" rank="species" species="princeps">
|
||||
<emphasis id="68A4AE7062A95D95583C464D763C449E" italics="true" pageId="0" pageNumber="337">P. princeps</emphasis>
|
||||
</taxonomicName>
|
||||
Butler has been somewhat inconsistently used for the East Palearctic populations. E.g.
|
||||
<bibRefCitation author="Shibuya, J" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="0" pageNumber="337" pagination="149 - 176" refId="B24" refString="Shibuya, J, 1928. The systematic study on the Japanese Pyralinae (Lepid.). Journal of the Faculty of Agriculture, Hokkaido Imperial University 21 (4): 149 - 176" title="The systematic study on the Japanese Pyralinae (Lepid.)." volume="21" year="1928">Shibuya (1928)</bibRefCitation>
|
||||
<bibRefCitation id="E73F8F4DF2B3019144AB3417EE047649" author="Shibuya, J" journalOrPublisher="Journal of the Faculty of Agriculture, Hokkaido Imperial University" pageId="0" pageNumber="337" pagination="149 - 176" refId="B24" refString="Shibuya, J, 1928. The systematic study on the Japanese Pyralinae (Lepid.). Journal of the Faculty of Agriculture, Hokkaido Imperial University 21 (4): 149 - 176" title="The systematic study on the Japanese Pyralinae (Lepid.)." volume="21" year="1928">Shibuya (1928)</bibRefCitation>
|
||||
, in his treatment of Japanese species of
|
||||
<taxonomicName class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">Pyralis</emphasis>
|
||||
<taxonomicName id="2BED4C9A9924D5C55AC8416F4D94407B" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="7CFA64BA4D8B53440F55F9B728F21AC9" italics="true" pageId="0" pageNumber="337">Pyralis</emphasis>
|
||||
</taxonomicName>
|
||||
and related genera, only recognized
|
||||
<taxonomicName genus="P." lsidName="P. regalis" pageId="0" pageNumber="337" rank="species" species="regalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. regalis</emphasis>
|
||||
<taxonomicName id="5CB815DF95788AAACC5B0FEDBAAA0B94" lsidName="P. regalis" pageId="0" pageNumber="337" rank="species" species="regalis">
|
||||
<emphasis id="C1EB366069AE92D60B437F057A27E478" italics="true" pageId="0" pageNumber="337">P. regalis</emphasis>
|
||||
</taxonomicName>
|
||||
of the relevant taxa to occur there. Inoue
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">et al.</emphasis>
|
||||
<emphasis id="AE041189AADF879736E2E1A6B19C666F" italics="true" pageId="0" pageNumber="337">et al.</emphasis>
|
||||
(1982), plate 46: Figs
|
||||
<figureCitation captionStart="Figures 4–11" captionStartId="F4" captionText="Figures 4 - 11. Pyralis sp., habitus. 4 - 7. P. cardinalis sp. nov., 4. ♂, Finland, V, Kemioensaari, leg. J. Tyllinen. 5. ♀, A, Lemland, leg. B. Wikstroem & K. Vaalamo; 6. ♂, Finland, U, Kirkkonummi, leg. J. Junnilainen; 7. ♂, Finland, U, Raasepori, leg. L. Kaila. 8 - 11. P. regalis. 8. ♂, Greece, Lesvos, leg. L. Kaila & J. Kullberg. 9. ♀, Slovenia, Granska Gora, leg. B. Wikstroem; 10. ♂, Italy, Tuscany, Cortona, leg. B. Wikstroem. 11. Bulgaria, 5 km S. Blagoevgrad, ♀, leg. B. A. Bengtsson." figureDoi="10.3897/nl.43.54916.figures4-11" httpUri="https://binary.pensoft.net/fig/473649" pageId="0" pageNumber="337">4</figureCitation>
|
||||
<figureCitation id="DE0366C9102A9871A54CA935E277A325" captionStart="Figures 4–11" captionStartId="F4" captionText="Figures 4 - 11. Pyralis sp., habitus. 4 - 7. P. cardinalis sp. nov., 4. ♂, Finland, V, Kemioensaari, leg. J. Tyllinen. 5. ♀, A, Lemland, leg. B. Wikstroem & K. Vaalamo; 6. ♂, Finland, U, Kirkkonummi, leg. J. Junnilainen; 7. ♂, Finland, U, Raasepori, leg. L. Kaila. 8 - 11. P. regalis. 8. ♂, Greece, Lesvos, leg. L. Kaila & J. Kullberg. 9. ♀, Slovenia, Granska Gora, leg. B. Wikstroem; 10. ♂, Italy, Tuscany, Cortona, leg. B. Wikstroem. 11. Bulgaria, 5 km S. Blagoevgrad, ♀, leg. B. A. Bengtsson." figureDoi="10.3897/nl.43.54916.figures4-11" httpUri="https://binary.pensoft.net/fig/473649" pageId="0" pageNumber="337">4</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figures 4–11" captionStartId="F4" captionText="Figures 4 - 11. Pyralis sp., habitus. 4 - 7. P. cardinalis sp. nov., 4. ♂, Finland, V, Kemioensaari, leg. J. Tyllinen. 5. ♀, A, Lemland, leg. B. Wikstroem & K. Vaalamo; 6. ♂, Finland, U, Kirkkonummi, leg. J. Junnilainen; 7. ♂, Finland, U, Raasepori, leg. L. Kaila. 8 - 11. P. regalis. 8. ♂, Greece, Lesvos, leg. L. Kaila & J. Kullberg. 9. ♀, Slovenia, Granska Gora, leg. B. Wikstroem; 10. ♂, Italy, Tuscany, Cortona, leg. B. Wikstroem. 11. Bulgaria, 5 km S. Blagoevgrad, ♀, leg. B. A. Bengtsson." figureDoi="10.3897/nl.43.54916.figures4-11" httpUri="https://binary.pensoft.net/fig/473649" pageId="0" pageNumber="337">5</figureCitation>
|
||||
<figureCitation id="F0917AA63A189276EC787C03FB3AA3A3" captionStart="Figures 4–11" captionStartId="F4" captionText="Figures 4 - 11. Pyralis sp., habitus. 4 - 7. P. cardinalis sp. nov., 4. ♂, Finland, V, Kemioensaari, leg. J. Tyllinen. 5. ♀, A, Lemland, leg. B. Wikstroem & K. Vaalamo; 6. ♂, Finland, U, Kirkkonummi, leg. J. Junnilainen; 7. ♂, Finland, U, Raasepori, leg. L. Kaila. 8 - 11. P. regalis. 8. ♂, Greece, Lesvos, leg. L. Kaila & J. Kullberg. 9. ♀, Slovenia, Granska Gora, leg. B. Wikstroem; 10. ♂, Italy, Tuscany, Cortona, leg. B. Wikstroem. 11. Bulgaria, 5 km S. Blagoevgrad, ♀, leg. B. A. Bengtsson." figureDoi="10.3897/nl.43.54916.figures4-11" httpUri="https://binary.pensoft.net/fig/473649" pageId="0" pageNumber="337">5</figureCitation>
|
||||
illustrated Japanese specimens that seem identical to
|
||||
<taxonomicName genus="P." lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
|
||||
<taxonomicName id="492E8AA862A5073D3E107BDFE5644130" lsidName="P. cardinalis" pageId="0" pageNumber="337" rank="species" species="cardinalis">
|
||||
<emphasis id="C5F00D16AA48001006BDFFA03ADD8ABE" italics="true" pageId="0" pageNumber="337">P. cardinalis</emphasis>
|
||||
</taxonomicName>
|
||||
and whose occurrence in Japan is verified in the present study. Neither of these authors mention other species, but they consider
|
||||
<taxonomicName genus="P." lsidName="P. princeps" pageId="0" pageNumber="337" rank="species" species="princeps">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. princeps</emphasis>
|
||||
<taxonomicName id="26E9C758F81124FE9F258CCB51BC392C" lsidName="P. princeps" pageId="0" pageNumber="337" rank="species" species="princeps">
|
||||
<emphasis id="C15877F006836D94F19F6DA304C7B5B1" italics="true" pageId="0" pageNumber="337">P. princeps</emphasis>
|
||||
</taxonomicName>
|
||||
a synonym of
|
||||
<taxonomicName genus="P." lsidName="P. regalis" pageId="0" pageNumber="337" rank="species" species="regalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. regalis</emphasis>
|
||||
<taxonomicName id="6B1371B3537DCB6C5977C5A2745C64F4" lsidName="P. regalis" pageId="0" pageNumber="337" rank="species" species="regalis">
|
||||
<emphasis id="E9017E7691FEF8295F857F6B04405916" italics="true" pageId="0" pageNumber="337">P. regalis</emphasis>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/nl.43.54916.figures18-21" httpUri="https://binary.pensoft.net/fig/473651" pageId="0" pageNumber="337" start="Figures 18–21" startId="F6">
|
||||
<paragraph pageId="0" pageNumber="337">
|
||||
<emphasis bold="true" pageId="0" pageNumber="337">Figures 18-21.</emphasis>
|
||||
<taxonomicName class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">Pyralis</emphasis>
|
||||
<caption id="71750A18FA2ED9074381B59607AD40EB" doi="10.3897/nl.43.54916.figures18-21" httpUri="https://binary.pensoft.net/fig/473651" pageId="0" pageNumber="337" start="Figures 18–21" startId="F6">
|
||||
<paragraph id="133F987D9BF647AAE2E770610B634D25" pageId="0" pageNumber="337">
|
||||
<emphasis id="85A135B2FF880C8FF88BCF6DE505261E" bold="true" pageId="0" pageNumber="337">Figures 18-21.</emphasis>
|
||||
<taxonomicName id="6D820855B6086272980CE83BFAB05E56" class="Insecta" family="Pyralidae" genus="Pyralis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyralis" order="Lepidoptera" pageId="0" pageNumber="337" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="62AECF58BE127FE63DA6A4EEF34F6AC3" italics="true" pageId="0" pageNumber="337">Pyralis</emphasis>
|
||||
</taxonomicName>
|
||||
spp., habitus.
|
||||
<emphasis bold="true" pageId="0" pageNumber="337">18.</emphasis>
|
||||
<taxonomicName genus="P." lsidName="P. princeps" pageId="0" pageNumber="337" rank="species" species="princeps">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. princeps</emphasis>
|
||||
<emphasis id="E6AC4F39CA004F0BE8587E367AD30850" bold="true" pageId="0" pageNumber="337">18.</emphasis>
|
||||
<taxonomicName id="7E9B504A888FF3B4C1F006A1410A66D4" lsidName="P. princeps" pageId="0" pageNumber="337" rank="species" species="princeps">
|
||||
<emphasis id="7530949A41A213633C3C1B3133C688C8" italics="true" pageId="0" pageNumber="337">P. princeps</emphasis>
|
||||
</taxonomicName>
|
||||
Butler, ♂; Nepal, Godawari nr. Kathmandu, leg. exp. Albrecht et al.
|
||||
<emphasis bold="true" pageId="0" pageNumber="337">19.</emphasis>
|
||||
<taxonomicName genus="P." lsidName="P. joannisi" pageId="0" pageNumber="337" rank="species" species="joannisi">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. joannisi</emphasis>
|
||||
<emphasis id="1C4EA26F0DAA06A4A1287AD2979566B7" bold="true" pageId="0" pageNumber="337">19.</emphasis>
|
||||
<taxonomicName id="18E6A9EBA7891CDA5974FF72E43D82CD" lsidName="P. joannisi" pageId="0" pageNumber="337" rank="species" species="joannisi">
|
||||
<emphasis id="160C912BDBFA22CC2432E7C8E0AD08F0" italics="true" pageId="0" pageNumber="337">P. joannisi</emphasis>
|
||||
</taxonomicName>
|
||||
Leraut, ♂; Vietnam, Bac Kan province, leg. A.
|
||||
<normalizedToken originalValue="Wikström">Wikstroem</normalizedToken>
|
||||
<normalizedToken id="EBB47CB18A268A0D80A400A89DEF72D1" originalValue="Wikström">Wikstroem</normalizedToken>
|
||||
.
|
||||
<emphasis bold="true" pageId="0" pageNumber="337">20, 21.</emphasis>
|
||||
<taxonomicName genus="P." lsidName="P. perversalis" pageId="0" pageNumber="337" rank="species" species="perversalis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="337">P. perversalis</emphasis>
|
||||
<emphasis id="A84F51C557277B8F99FE221D792B8A8F" bold="true" pageId="0" pageNumber="337">20, 21.</emphasis>
|
||||
<taxonomicName id="5F1E5755901B638ED05BB88B16445B05" lsidName="P. perversalis" pageId="0" pageNumber="337" rank="species" species="perversalis">
|
||||
<emphasis id="DC44E0C64FE726C251ECF373B172D939" italics="true" pageId="0" pageNumber="337">P. perversalis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<normalizedToken originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
|
||||
<normalizedToken id="F446A19AAA0EB811567E61E781187763" originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
|
||||
), habitus.
|
||||
<emphasis bold="true" pageId="0" pageNumber="337">20.</emphasis>
|
||||
<emphasis id="E96DB8E0135D89477B80023D98DFBC78" bold="true" pageId="0" pageNumber="337">20.</emphasis>
|
||||
♂, Czech Republic, Bohemia sept., leg. J.
|
||||
<normalizedToken originalValue="Liška">Liska</normalizedToken>
|
||||
<normalizedToken id="A3EE0E9C83F051CF95C40D8D6D55AFE2" originalValue="Liška">Liska</normalizedToken>
|
||||
;
|
||||
<emphasis bold="true" pageId="0" pageNumber="337">21.</emphasis>
|
||||
<emphasis id="56EF1F01FDAC72E8184C5498575D0A8F" bold="true" pageId="0" pageNumber="337">21.</emphasis>
|
||||
♀, S. Ural, Cheliabinsk district, leg. T. Nupponen.
|
||||
</paragraph>
|
||||
</caption>
|
||||
|
|
|
|||
Loading…
Add table
Add a link
Reference in a new issue