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Sm. in Sibthorp &amp; Smith, Fl. Graec. Prodr. 1: 283. 1809.
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not designated; original material: &quot;In
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minori&quot;,
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<emphasis id="BB92AB0826E89D4EACF7ADDD789920FE" italics="true" pageId="0" pageNumber="91">Sibthorp s.n.</emphasis>
(OXF Sib-FG Sib-0954!, K000077455!).
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6148996" ID-GBIF-Taxon="125135988" ID-Zenodo-Dep="6148996" LSID="urn:lsid:plazi:treatment:BB2C9CAF4C1EE10D744A074D0D74675C" httpUri="http://treatment.plazi.org/id/BB2C9CAF4C1EE10D744A074D0D74675C" lastPageId="19" lastPageNumber="119" pageId="7" pageNumber="108">
<subSubSection pageId="7" pageNumber="108" type="nomenclature">
<paragraph box="[118,420,1317,1392]" pageId="7" pageNumber="108" type="heading">
<taxonomicName authority="Forel" authorityName="Forel" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei (Forel)</taxonomicName>
<treatment id="BB2C9CAF4C1EE10D744A074D0D74675C" ID-DOI="http://doi.org/10.5281/zenodo.6148996" ID-GBIF-Taxon="125135988" ID-Zenodo-Dep="6148996" LSID="urn:lsid:plazi:treatment:BB2C9CAF4C1EE10D744A074D0D74675C" httpUri="http://treatment.plazi.org/id/BB2C9CAF4C1EE10D744A074D0D74675C" lastPageId="19" lastPageNumber="119" pageId="7" pageNumber="108" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="E979942F066A8CA7A268798E8D207410" pageId="7" pageNumber="108" type="nomenclature">
<paragraph id="AF214336794BD32FB30AFA861A4A644E" box="[118,420,1317,1392]" pageId="7" pageNumber="108" type="heading">
<taxonomicName id="7AA6B12C6E762D3BAD7E5298BEBD0E8A" ID-CoL="4BQTK" authority="Forel" authorityName="Forel" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei (Forel)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="108" type="description">
<paragraph box="[118,400,1400,1419]" pageId="7" pageNumber="108" type="mainText">(Figures 6, 8, 10, 14-28, 44-57)</paragraph>
<subSubSection id="14FBAA18CC53F0C1E9C0E3B44C1F8F28" pageId="7" pageNumber="108" type="description">
<paragraph id="46F242345D2593309800EDB0D47B00F2" box="[118,400,1400,1419]" pageId="7" pageNumber="108" type="mainText">(Figures 6, 8, 10, 14-28, 44-57)</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="108" type="materials_examined">
<paragraph box="[118,1123,1456,1569]" pageId="7" pageNumber="108" type="mainText">
<taxonomicName authority="Forel, 1891" authorityName="Forel," authorityYear="1891" class="Hexapoda" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="cambouei">
<subSubSection id="D377E89E894C3ADF304B406F5A5C94C8" pageId="7" pageNumber="108" type="materials_examined">
<paragraph id="7C59B7EA2D8B0F097D2FDFAA35C19C39" box="[118,1123,1456,1569]" pageId="7" pageNumber="108" type="mainText">
<taxonomicName id="48D6C6F26AED6EBAA0D33346C5982F57" authority="Forel, 1891" authorityName="Forel," authorityYear="1891" genus="Bothroponera" pageId="7" pageNumber="108" rank="species" species="cambouei">
Bothroponera cambouei
<bibRefCitation author="Forel, A." box="[118,1122,502,543]" editor="Grandidier, A." journalOrPublisher="Hachette et Cie, Paris" pageId="39" pageNumber="140" pagination="v + 237 pp" title="Les Formicides. [part]" type="book" volumeTitle="1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule)" year="1891">Forel, 1891</bibRefCitation>
<bibRefCitation id="4F471AF696DE5C77C022F5CEBA7CB28E" author="Forel, A." box="[118,1122,502,543]" editor="Grandidier, A." journalOrPublisher="Hachette et Cie, Paris" pageId="39" pageNumber="140" pagination="v + 237 pp" title="Les Formicides. [part]" type="book" volumeTitle="1891. Histoire physique, naturelle, et politique de Madagascar. Volume XX. Histoire naturelle des Hymenopteres. Deuxieme partie (28 e fascicule)" year="1891">Forel, 1891</bibRefCitation>
</taxonomicName>
: 133, pl. 4, fig. 7 Lectotype worker, present designation, Madagascar, Imerina, Antananarivo (
<normalizedToken originalValue="Camboué" pageId="7" pageNumber="108">Camboue</normalizedToken>
<normalizedToken id="E1CD3A2631E7EDDCAB87AA8ED4AA4458" originalValue="Camboué" pageId="7" pageNumber="108">Camboue</normalizedToken>
) AntWeb CASENT0101770 (MHNG) [examined]. Paralectotype worker and queens with same data as lectotype but with specimen codes: CASENT0101028, CASENT0101029 (MHNG) and CASENT0101404, CASENT0101402 (MNHN) [examined]. [Combination in
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="subGenus" subGenus="Bothroponera">Pachycondyla (Bothroponera)</taxonomicName>
:
<bibRefCitation author="Emery, C." box="[118,1122,1464,1505]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="38" pageNumber="139" pagination="32 - 54" part="45" title="Notes sur les sous-familles des Dorylines et Ponerines (Famille des Formicides)" type="journal article" year="1901">Emery, 1901</bibRefCitation>
: 45,
<bibRefCitation author="Emery, C." box="[118,938,1512,1530]" journalOrPublisher="Genera Insectorum" pageId="38" pageNumber="139" pagination="1 - 125" part="118" title="Hymenoptera. Fam. Formicidae. Subfam. Ponerinae" type="journal article" year="1911">1911</bibRefCitation>
: 78; Combination in
<taxonomicName authority="Mayr" authorityName="Mayr" class="Hexapoda" family="Formicidae" genus="Bothroponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">Bothroponera</taxonomicName>
<taxonomicName id="21E704B2F30BD326AA9D0C7C8A90F610" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="subGenus" subGenus="Bothroponera">Pachycondyla (Bothroponera)</taxonomicName>
:
<bibRefCitation author="Wheeler, W. M." box="[118,1122,1005,1046]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="141" pagination="1005 - 1055" part="45" title="Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region" type="journal article" year="1922">Wheeler, 1922</bibRefCitation>
<bibRefCitation id="D1E4D7C6A9975DA5AEEA78992D604DB0" author="Emery, C." box="[118,1122,1464,1505]" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="38" pageNumber="139" pagination="32 - 54" part="45" title="Notes sur les sous-familles des Dorylines et Ponerines (Famille des Formicides)" type="journal article" year="1901">Emery, 1901</bibRefCitation>
: 45,
<bibRefCitation id="1FF365EF7592E165D89067D5900EB188" author="Emery, C." box="[118,938,1512,1530]" journalOrPublisher="Genera Insectorum" pageId="38" pageNumber="139" pagination="1 - 125" part="118" title="Hymenoptera. Fam. Formicidae. Subfam. Ponerinae" type="journal article" year="1911">1911</bibRefCitation>
: 78; Combination in
<taxonomicName id="3D604689AF562C629837B063F1DFCD3C" genus="Bothroponera" pageId="7" pageNumber="108" rank="genus">Bothroponera</taxonomicName>
:
<bibRefCitation id="C845421248298833419F3464BA98DAA4" author="Wheeler, W. M." box="[118,1122,1005,1046]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="40" pageNumber="141" pagination="1005 - 1055" part="45" title="Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. IX. A synonymic list of the ants of the Malagasy region" type="journal article" year="1922">Wheeler, 1922</bibRefCitation>
: 1007; in
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">Pachycondyla</taxonomicName>
<taxonomicName id="8E1FA044C40ACB02552DE6BF64D4B9E8" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">Pachycondyla</taxonomicName>
by
<bibRefCitation author="Bolton, B." box="[118,1122,913,954]" journalOrPublisher="Harvard University Press, Cambridge, Massachusetts" pageId="38" pageNumber="139" pagination="504 pp" title="A New General Catalogue of the Ants of the World" type="book" year="1995">Bolton, 1995</bibRefCitation>
<bibRefCitation id="D9E72EADD044A352DD4E699D1F6AA96B" author="Bolton, B." box="[118,1122,913,954]" journalOrPublisher="Harvard University Press, Cambridge, Massachusetts" pageId="38" pageNumber="139" pagination="504 pp" title="A New General Catalogue of the Ants of the World" type="book" year="1995">Bolton, 1995</bibRefCitation>
: 303].
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="109" type="diagnosis">
<paragraph box="[118,290,118,137]" pageId="8" pageNumber="109" type="mainText">
<pageBreakToken pageId="8" pageNumber="109" start="start">Worker</pageBreakToken>
<subSubSection id="4F6ACCBB7CE551461858B25C3C9EC0EC" pageId="8" pageNumber="109" type="diagnosis">
<paragraph id="EE4627F23DE25AD4E9F875EEAFA02A41" box="[118,290,118,137]" pageId="8" pageNumber="109" type="mainText">
<pageBreakToken id="9856C08DE8DF482E210F18AB5B8C21ED" pageId="8" pageNumber="109" start="start">Worker</pageBreakToken>
diagnosis:
</paragraph>
<paragraph box="[118,1120,147,278]" pageId="8" pageNumber="109" type="mainText">With head in full-face view, antennal scape not surpassing posterior cephalic margin; dorsum of head and mesosoma not finely striate; dorsum of the body with erect slender hairs and pubescence; basal half of antennal scape rounded; dorsal outline of mesosoma not forming a continuous convexity; distinct angle discernable between propodeal dorsum and declivitous surface; petiole nodiform, not anteroposteriorly flattened; anterior half of fourth abdominal tergite (gastral tergite 2) mostly smooth and shiny between large punctures.</paragraph>
<paragraph box="[118,1121,287,334]" pageId="8" pageNumber="109" type="mainText">Worker measurements (n=116): HL: 1.38-2.29, HW: 1.19-2.01, CI: 85-95, SL: 0.92-1.69, SI: 72-87, PW: 0.95-1.71, WL: 1.64-2.89, NL: 0.65-1.05, NW: 0.79-1.26, NH: 0.95-1.54, DNI: 110-137, LNI: 130-169.</paragraph>
<paragraph id="0985BCB10BD8E8E0FEF412E848D50976" box="[118,1120,147,278]" pageId="8" pageNumber="109" type="mainText">With head in full-face view, antennal scape not surpassing posterior cephalic margin; dorsum of head and mesosoma not finely striate; dorsum of the body with erect slender hairs and pubescence; basal half of antennal scape rounded; dorsal outline of mesosoma not forming a continuous convexity; distinct angle discernable between propodeal dorsum and declivitous surface; petiole nodiform, not anteroposteriorly flattened; anterior half of fourth abdominal tergite (gastral tergite 2) mostly smooth and shiny between large punctures.</paragraph>
<paragraph id="DC067EC05244AB9900007CA43EB2B2CD" box="[118,1121,287,334]" pageId="8" pageNumber="109" type="mainText">Worker measurements (n=116): HL: 1.38-2.29, HW: 1.19-2.01, CI: 85-95, SL: 0.92-1.69, SI: 72-87, PW: 0.95-1.71, WL: 1.64-2.89, NL: 0.65-1.05, NW: 0.79-1.26, NH: 0.95-1.54, DNI: 110-137, LNI: 130-169.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="109" type="description">
<paragraph box="[118,233,371,390]" pageId="8" pageNumber="109" type="mainText">Description:</paragraph>
<paragraph box="[118,1121,400,700]" pageId="8" pageNumber="109" type="mainText">Worker. In full-face view head widest at or slightly behind level of eyes; sides somewhat convex, but weakly converging in front of eyes; shape of posterior margin variable, ranging from straight to strongly medially excised. Compound eyes either large and moderately protruding, or small with diameter less than half of maximum diameter of scape; with head in full-face view, eyes breaking lateral margins of the head. Antennal scape subcylindrical, with rounded leading edge, and not surpassing posterior margin of head. Anterior clypeal margin truncate and either straight or slightly concave. Mandibles triangular, apical margins armed with seven to nine teeth and denticles. With mesosoma in profile, the dorsal outline not forming a continuous convexity, propodeal dorsum meeting declivity at a distinct angle; mesopleural sulcus usually indistinct and incomplete. Petiole nodiform; in lateral view, anterior face, dorsum, and posterior face meet in a rounded angle. With petiole node in dorsal view, posterior margin straight or medially excised. Hind basitarsus nearly rounded and not flattened dorsoventrally, basal half of inside surface not concave.</paragraph>
<paragraph box="[118,1120,709,1066]" pageId="8" pageNumber="109" type="mainText">Mandibles longitudinally striate and covered with sparse piligerous punctures. Dorsum of head finely reticulate-punctate or reticulate rugulose, superimposed by punctures or setae-bearing foveolae; sides with less defined sculpture, but finely reticulate-punctate or reticulate-rugulose, interspersed with small piligerous pits. Mesosoma and petiole node with variable sculpture, dorsally ranging from coarsely densely punctate to reticulate-rugose or irregularly punctate, interspersed with quite closely spaced smaller punctures; generally fine striations running to bottom of large punctures. For lateral portions, sculpturing varies from finely rugulose to transversely striate with scattered punctures. Propodeal declivity smooth and shiny or with granular surface. Third and fourth abdominal tergites covered with large, shallow punctures, interspaces either smooth and shiny or covered with fine, small punctures; in some specimen s the large punctures are irregularly spaced and much more accentuated. Dorsum of body covered with erect slender or suberect thick hairs, which are reduced in number or absent on the antennal scape and legs. Pubescence on the third and fourth abdominal tergites either dense, sparse, or lacking. Body colors range from ferruginous red to reddish brown through dark brown to black with lighter appendages.</paragraph>
<paragraph box="[118,1121,1075,1206]" pageId="8" pageNumber="109" type="mainText">
<subSubSection id="5B944CFB13FBDE1DA6835E6F9C513A8B" pageId="8" pageNumber="109" type="description">
<paragraph id="5C458596A8BA5C44FE11E23C7C471F6D" box="[118,233,371,390]" pageId="8" pageNumber="109" type="mainText">Description:</paragraph>
<paragraph id="91CAE7F5728539CEE2AE5584696C22D6" box="[118,1121,400,700]" pageId="8" pageNumber="109" type="mainText">Worker. In full-face view head widest at or slightly behind level of eyes; sides somewhat convex, but weakly converging in front of eyes; shape of posterior margin variable, ranging from straight to strongly medially excised. Compound eyes either large and moderately protruding, or small with diameter less than half of maximum diameter of scape; with head in full-face view, eyes breaking lateral margins of the head. Antennal scape subcylindrical, with rounded leading edge, and not surpassing posterior margin of head. Anterior clypeal margin truncate and either straight or slightly concave. Mandibles triangular, apical margins armed with seven to nine teeth and denticles. With mesosoma in profile, the dorsal outline not forming a continuous convexity, propodeal dorsum meeting declivity at a distinct angle; mesopleural sulcus usually indistinct and incomplete. Petiole nodiform; in lateral view, anterior face, dorsum, and posterior face meet in a rounded angle. With petiole node in dorsal view, posterior margin straight or medially excised. Hind basitarsus nearly rounded and not flattened dorsoventrally, basal half of inside surface not concave.</paragraph>
<paragraph id="2FDEE520BE0248948E0F5205D2B2E290" box="[118,1120,709,1066]" pageId="8" pageNumber="109" type="mainText">Mandibles longitudinally striate and covered with sparse piligerous punctures. Dorsum of head finely reticulate-punctate or reticulate rugulose, superimposed by punctures or setae-bearing foveolae; sides with less defined sculpture, but finely reticulate-punctate or reticulate-rugulose, interspersed with small piligerous pits. Mesosoma and petiole node with variable sculpture, dorsally ranging from coarsely densely punctate to reticulate-rugose or irregularly punctate, interspersed with quite closely spaced smaller punctures; generally fine striations running to bottom of large punctures. For lateral portions, sculpturing varies from finely rugulose to transversely striate with scattered punctures. Propodeal declivity smooth and shiny or with granular surface. Third and fourth abdominal tergites covered with large, shallow punctures, interspaces either smooth and shiny or covered with fine, small punctures; in some specimen s the large punctures are irregularly spaced and much more accentuated. Dorsum of body covered with erect slender or suberect thick hairs, which are reduced in number or absent on the antennal scape and legs. Pubescence on the third and fourth abdominal tergites either dense, sparse, or lacking. Body colors range from ferruginous red to reddish brown through dark brown to black with lighter appendages.</paragraph>
<paragraph id="7355FFCEF5655C30F8DD2A32EB33D45C" box="[118,1121,1075,1206]" pageId="8" pageNumber="109" type="mainText">
Queen. Measurements (n=14): HL: 1.53-2.18, HW: 1.41-2.09, CI: 89-98, EL: 0.28-0.42, OI: 17-21, SL: 1.02-1.66, SI: 72-82, PW: 1.21-1.87, WL: 2.36-3.45, NL: 0.63-1.02, NW: 0.94-1.34, NH: 0.92-1.35, DNI: 130-153, LNI: 129-146. Winged queens and ergatoids are present in
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="cambouei">cambouei</taxonomicName>
<taxonomicName id="1F47C940EB29A3831BC60F8358FB5662" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="cambouei">cambouei</taxonomicName>
and its variants. Ergatoid queens and workers look very similar, but the former have ocelli and incomplete thoracic sclerites while the body of alate queens is noticeably larger in general.
</paragraph>
<paragraph box="[118,211,1239,1258]" pageId="8" pageNumber="109" type="mainText">Variation:</paragraph>
<paragraph box="[118,1120,1267,1432]" pageId="8" pageNumber="109" type="mainText">
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei</taxonomicName>
<paragraph id="548303E981D874548A9B73E3A94AEBEB" box="[118,211,1239,1258]" pageId="8" pageNumber="109" type="mainText">Variation:</paragraph>
<paragraph id="5E573007C20C359A15ADCD1BFF30E115" box="[118,1120,1267,1432]" pageId="8" pageNumber="109" type="mainText">
<taxonomicName id="230339C608EE08D0082E8A9C865CC818" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei</taxonomicName>
shows a remarkable range of phenotypic diversity. To faciliate discussion of this diversity, the morphological characters of the worker caste are divided into seven forms on the basis of the shape of the occipital corner, the posterior margin of the head, the size of the eyes, the form of the mesosoma, and the abundance of pubescence on the fourth abdominal tergites. However, there is no simple pattern to the degree of variation of these forms. Specifically, a few forms have very restricted geographic boundaries while others show a wide distribution along the humid forests of Madagascar. Each form of
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="67307A586524BAA43B6304D37E67FA0B" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
is described below:
</paragraph>
<paragraph box="[118,1121,1442,1519]" pageId="8" pageNumber="109" type="mainText">Form 1 (Figs 10, 15, 18, 22, 24-25, 53). Worker measurements (n=35 workers): HL: 1.75-2.17, HW: 1.59-1.99, CI: 88-95, SL: 1.24-1.57, SI: 75-85, PW: 1.23-1.67, WL: 2.15-2.89, NL: 0.82-1.02, NW: 0.96-1.24, NH: 1.16-1.54, DNI: 110-130, LNI: 130-157.</paragraph>
<paragraph id="BB94FBB0F51D87622A099206B07AB797" box="[118,1121,1442,1519]" pageId="8" pageNumber="109" type="mainText">Form 1 (Figs 10, 15, 18, 22, 24-25, 53). Worker measurements (n=35 workers): HL: 1.75-2.17, HW: 1.59-1.99, CI: 88-95, SL: 1.24-1.57, SI: 75-85, PW: 1.23-1.67, WL: 2.15-2.89, NL: 0.82-1.02, NW: 0.96-1.24, NH: 1.16-1.54, DNI: 110-130, LNI: 130-157.</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="110" pageId="8" pageNumber="109" type="discussion">
<paragraph box="[118,1124,1530,1578][118,1124,118,312]" lastPageId="9" lastPageNumber="110" pageId="8" pageNumber="109" type="mainText">
<subSubSection id="0EBB6FF121B9EF8D0B076751B54544B5" lastPageId="9" lastPageNumber="110" pageId="8" pageNumber="109" type="discussion">
<paragraph id="4B323364FAEC9FB6AF875D47C57BF159" box="[118,1124,1530,1578][118,1124,118,312]" lastPageId="9" lastPageNumber="110" pageId="8" pageNumber="109" type="mainText">
This form corresponds to the typical
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="06520BC15628E9B0335E58DB157EE0AE" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
, which is distinguished by much larger eyes, medially excised posterior cephalic margin and normal shape of head and mesosoma, medially excised posterior margin of petiole, and
<pageBreakToken pageId="9" pageNumber="110" start="start">smooth</pageBreakToken>
<pageBreakToken id="76E91C29414C04480CD647D4CDE55AC7" pageId="9" pageNumber="110" start="start">smooth</pageBreakToken>
or sparsely punctate fourth abdominal tergites. Some characters are variable, such as the smooth or shagreenate propodeal declivity, and slender erect hairs with reduced pubescence or shorter and much more robust erect hairs with more pubescence. A few larger worker specimens (HW: 1.86-1.99 mm) from the Masoala area in northeastern Madagascar are included in this form. These larger workers have less defined sculpture and are covered with elongate and slender erect hairs and less pubescence. At first glance, these differences are suggestive of a distinct species, but other specimens from various localities also show patterns of less enhanced sculpture, slender and long erect hairs, and a larger-sized body than a more typical
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="44FFB7963318D6A7FCFBB7D1267E70F1" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
specimen.
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="113" pageId="9" pageNumber="110" type="description">
<paragraph box="[118,1120,323,399]" pageId="9" pageNumber="110" type="mainText">Form 2 (Figs 8, 16, 26-27, 54). Worker measurements (n= 8): HL: 1.83-2.19, HW: 1.66-1.96, CI: 88-91, SL: 1.33-1.51, SI: 77-81, PW: 1.26-1.48, WL: 2.35-2.69, NL: 0.82-0.97, NW: 0.98-1.18, NH: 1.19-1.44, DNI: 114-121, LNI: 138-148.</paragraph>
<paragraph box="[118,1128,410,546]" pageId="9" pageNumber="110" type="mainText">
<subSubSection id="1AD8A6ABA6411823AEF8A935E1803478" lastPageId="12" lastPageNumber="113" pageId="9" pageNumber="110" type="description">
<paragraph id="CCE363660ACC0625F8D86B7FDD8B142C" box="[118,1120,323,399]" pageId="9" pageNumber="110" type="mainText">Form 2 (Figs 8, 16, 26-27, 54). Worker measurements (n= 8): HL: 1.83-2.19, HW: 1.66-1.96, CI: 88-91, SL: 1.33-1.51, SI: 77-81, PW: 1.26-1.48, WL: 2.35-2.69, NL: 0.82-0.97, NW: 0.98-1.18, NH: 1.19-1.44, DNI: 114-121, LNI: 138-148.</paragraph>
<paragraph id="76B8CA4AC1A356B3C01F3B25E1356C19" box="[118,1128,410,546]" pageId="9" pageNumber="110" type="mainText">
Known only from workers collected in the isolated rainforest of the RS Ambohijanahary. This variant is characterized by a backward extension into a lobe of each angle of the posterior margin of the head (Fig. 16). However, workers with more typical characters of
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="FCBC48F269D6D0F7324A073730D2CAD8" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
are also found within a few colonies of this variant, and workers that show intermediate levels of protrusion between this form and form 3 (see below) also occur in FC Didy, FC Andriantantely and PN Mantadia. Workers with intermediate levels of protrusion are included in the latter form.
</paragraph>
<paragraph box="[118,1121,556,633]" pageId="9" pageNumber="110" type="mainText">Form 3 (Figs 6, 17, 28-29, 55). Worker measurements (n=25): HL: 1.92-2.22, HW: 1.72-2.03, CI: 86-93, SL: 1.40-1.58, SI: 77-82, PW: 1.29-1.55, AL: 2.37-3.00, NL: 0.91-0.99, NW: 1.04-1.17, NH: 1.3-1.43, DNI: 112-125, LNI: 140-152.</paragraph>
<paragraph box="[118,1122,643,808]" pageId="9" pageNumber="110" type="mainText">
<paragraph id="84F34C47285FCDBFBE310193D542332F" box="[118,1121,556,633]" pageId="9" pageNumber="110" type="mainText">Form 3 (Figs 6, 17, 28-29, 55). Worker measurements (n=25): HL: 1.92-2.22, HW: 1.72-2.03, CI: 86-93, SL: 1.40-1.58, SI: 77-82, PW: 1.29-1.55, AL: 2.37-3.00, NL: 0.91-0.99, NW: 1.04-1.17, NH: 1.3-1.43, DNI: 112-125, LNI: 140-152.</paragraph>
<paragraph id="94F0204F93DEE5F5CD74CE64CFB087D1" box="[118,1122,643,808]" pageId="9" pageNumber="110" type="mainText">
This variant is distinguished by the fact that the posterolateral margin of the head at each occipital corner protrudes posteriorly into a hornlike tooth. Individual workers with intermediate degrees of expansion of the occipital lobes are known from Ivohibe and PN Mantadia, and this trait varies gradually along its geographical distribution to match the general diagnosis of typical
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="8CF38B7617279CC0F1524F5275575073" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
(form 1). Interestingly, worker specimens in some colonies from the PN Ranomafana also present this gradual form of hornlike tooth, which for this form may be indicative of some biological meaning in terms of colony behavior.
</paragraph>
<paragraph box="[118,1120,818,895]" pageId="9" pageNumber="110" type="mainText">Form 4 (Figs 19, 30-31, 54). Worker measurements (n=10): HL: 1.74-1.97, HW: 1.55-1.82, CI: 89-92, SL: 1.25-1.48, SI: 79-86, PW: 1.21-1.38, WL: 2.14-2.55, NL: 0.84-0.99, NW: 0.97-1.14, NH: 1.22-1.34, DNI: 111-126, LNI: 135-151.</paragraph>
<paragraph box="[118,1124,906,1071]" pageId="9" pageNumber="110" type="mainText">
<paragraph id="DC3E2FE77F2F9140C96A760D4F86DA8C" box="[118,1120,818,895]" pageId="9" pageNumber="110" type="mainText">Form 4 (Figs 19, 30-31, 54). Worker measurements (n=10): HL: 1.74-1.97, HW: 1.55-1.82, CI: 89-92, SL: 1.25-1.48, SI: 79-86, PW: 1.21-1.38, WL: 2.14-2.55, NL: 0.84-0.99, NW: 0.97-1.14, NH: 1.22-1.34, DNI: 111-126, LNI: 135-151.</paragraph>
<paragraph id="8CD662C9C13AD5C59BD69CC0788FBC74" box="[118,1124,906,1071]" pageId="9" pageNumber="110" type="mainText">
Workers of this form are differentiated by the absence of erect hairs and the absence or great reduction in abundance of pubescence on the dorsum of the propodeum. Numerous specimens that show an intermediate abundance of erect hairs and pubescence on the propodeal dorsum also are known across the range of
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="8360CEC4B780E7B2F072E9EF588D336F" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
. Specimens of this form could probably be misidentified as P.
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="masoala">masoala</taxonomicName>
<taxonomicName id="05D4B0A210EAD17D1A2AA5D976124A07" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="masoala">masoala</taxonomicName>
due to the absence of erect hairs on propodeal dorsum and less defined propodeal sculpture. However, the thickness of the petiolar node, which is about as broad as long when viewed laterally, combined with the gradient of characters, are those of a true
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="A59D04E1582F008933BFF020DCA10F63" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
.
</paragraph>
<paragraph box="[118,1120,1081,1158]" pageId="9" pageNumber="110" type="mainText">Form 5 (Figs 20, 32-33, 54). Worker measurements (n=5): HL: 2.10-2.29, HW: 1.82-2.01, CI: 87-88, SL: 1.54-1.69, SI: 83-85, PW: 1.49-1.71, WL: 2.71-2.88, NL: 0.92-1.05, NW: 1.15-1.26, NH: 1.48-1.52, DNI: 118-126, LNI: 141-163.</paragraph>
<paragraph box="[118,1122,1168,1450]" pageId="9" pageNumber="110" type="mainText">
<paragraph id="F7754F45BED3D7D9288B7A90F3FC36FE" box="[118,1120,1081,1158]" pageId="9" pageNumber="110" type="mainText">Form 5 (Figs 20, 32-33, 54). Worker measurements (n=5): HL: 2.10-2.29, HW: 1.82-2.01, CI: 87-88, SL: 1.54-1.69, SI: 83-85, PW: 1.49-1.71, WL: 2.71-2.88, NL: 0.92-1.05, NW: 1.15-1.26, NH: 1.48-1.52, DNI: 118-126, LNI: 141-163.</paragraph>
<paragraph id="3D1F08667A259A0608E231D876E51666" box="[118,1122,1168,1450]" pageId="9" pageNumber="110" type="mainText">
Two different altitudes in PN Masoala in northeast Madagascar yielded a few workers (4) in which both anterodorsal angles of the pronotum are bidentate or tuberculate where they meet the cervical shield, and the ventral surface of the head near the occipital angles is distinctly compressed; the integument of the entire body is ferruginous red. At first glance this form appears to be a morphologically distinct species, but among the four workers, there is one specimen with a pronotum almost angulate in dorsal view, with less developed teeth, the head apparently not strongly flattened ventrally near the occipital corners. An additional worker collected from the same national park presents a rounded anterodorsal angle of the pronotum when viewed dorsally and rounded occipital angles with the head in profile. The presence of these intermediate characters seems to link this variant to the other forms of
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="811C7E1F7E18DA300A4EC6A88798E2FE" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
. Colonies with all castes of this form should be collected in order to investigate how this character variation relates to the biology of the population and to clarify relationships with the other forms.
</paragraph>
<paragraph box="[118,1120,1460,1537]" pageId="9" pageNumber="110" type="mainText">Form 6 (Figs 14, 21, 23, 34-35, 56). Worker measurements (n=30 workers): HL: 1.60-2.07, HW: 1.46-1.87, CI: 86-93, SL: 1.11-1.57, SI: 74-87, PW: 1.12-1.47, WL: 2.02-2.74, NL: 0.76-1.02, NW: 0.93-1.18, NH: 1.08-1.41, DNI: 112-135, LNI: 132-169.</paragraph>
<caption box="[118,1124,1428,1590]" pageId="10" pageNumber="111" type="caption">
<paragraph box="[118,1124,1428,1590]" pageId="10" pageNumber="111" type="caption">
<pageBreakToken pageId="10" pageNumber="111" start="start">FIGURES</pageBreakToken>
<paragraph id="A0C002DFA08AFC0B47CBD0EA2FB0CF8C" box="[118,1120,1460,1537]" pageId="9" pageNumber="110" type="mainText">Form 6 (Figs 14, 21, 23, 34-35, 56). Worker measurements (n=30 workers): HL: 1.60-2.07, HW: 1.46-1.87, CI: 86-93, SL: 1.11-1.57, SI: 74-87, PW: 1.12-1.47, WL: 2.02-2.74, NL: 0.76-1.02, NW: 0.93-1.18, NH: 1.08-1.41, DNI: 112-135, LNI: 132-169.</paragraph>
<caption id="D45E452A1694D0091AB16DF83C72FA31" box="[118,1124,1428,1590]" pageId="10" pageNumber="111" type="caption">
<paragraph id="C9F849B7F7C3C1E58F8957FD9B5CEA68" box="[118,1124,1428,1590]" pageId="10" pageNumber="111" type="caption">
<pageBreakToken id="9AE7E6C864ECA01BF72E49414C6D0760" pageId="10" pageNumber="111" start="start">FIGURES</pageBreakToken>
15-23. Morphological character variation within
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="67AC5D35E207E5BC7408E52074499B1B" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
. Figs 15-17: Head in profile showing the shape of the occipital corner. Fig. 15: Normally rounded in form 1; Fig. 16: Protruding as a lobe in form 2; Fig. 17: Hornlike extension in form 3. Figs 18-19: Mesosoma in lateral view showing hairs on propodeal dorsum. Fig. 18: Hairs present in form 1; Fig. 19: Hairs absent in form 4. Figs 20-21: Anterodorsal angles of pronotum. Fig. 20: Bidentate in form 5; Fig. 21: Smoothly rounded in form 6. Figs 22-23: Sculpture and pubescence on fourth abdominal tergite. Fig. 22: Smooth and shiny between large punctures and pubescence absent (form 1); Fig. 23: With closely-spaced small punctures between large punctures and abundant pubescence (form 6).
</paragraph>
</caption>
<caption box="[118,1122,1186,1249]" pageId="11" pageNumber="112" type="caption">
<paragraph box="[118,1122,1186,1249]" pageId="11" pageNumber="112" type="caption">
<pageBreakToken pageId="11" pageNumber="112" start="start">FIGURES</pageBreakToken>
<caption id="E766D1C8DC285322C39A5842229E5075" box="[118,1122,1186,1249]" pageId="11" pageNumber="112" type="caption">
<paragraph id="8A047BC0CB95ABDB7A145C6483B119F1" box="[118,1122,1186,1249]" pageId="11" pageNumber="112" type="caption">
<pageBreakToken id="8981DF21072DC7F4C84B03132717582C" pageId="11" pageNumber="112" start="start">FIGURES</pageBreakToken>
24-29. Full-face view of the head and lateral view of the body of the worker caste of
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei</taxonomicName>
<taxonomicName id="3ADA16DF02960474D88D9C23B8071CF5" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei</taxonomicName>
and its various forms; Figs 24-25: Form 1, CASENT0101404; Figs 26-27: Form 2, CASENT0029098; Figs 28-29: Form 3, CASENT0034270.
</paragraph>
</caption>
<paragraph box="[118,1121,1278,1466]" pageId="11" pageNumber="112" type="mainText">
<paragraph id="F3CD4AC83CCBE88DF512220527546A8E" box="[118,1121,1278,1466]" pageId="11" pageNumber="112" type="mainText">
This is one of the forms most similar to typical
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="CD2A43143E89EC3CFAFFEB6F387BD681" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
. Workers can be recognized by the combination of the following characters: smaller eyes, straight posterior cephalic margin, slightly concave anterior clypeal margin, flagellar segments as long as broad, shagreened propodeal declivity, and less accentuated sculpture. Abdominal tergite 4 densely covered with appressed hairs when viewed dorsally; petiole node with lateral margins strongly converging anteriorly to form a bluntly rounded tip; posterior margin slightly concave with a weak median excision; body color reddish brown. This form is dominant in the humid, high altitude forests of southern Madagascar.
</paragraph>
<paragraph box="[118,1120,1475,1550]" pageId="11" pageNumber="112" type="mainText">Form 7 (Figs 36-37, 57). Worker measurements (n=20): HL: 1.38-1.65, HW: 1.19-1.44, CI: 85-92, SL: 0.92-1.13, SI: 72-80, PW: 0.95-1.18, WL: 1.64-2.07, NL: 0.65-0.81, NW: 0.79-1.05, NH: 0.95-1.17, DNI: 119-137, LNI: 139-152.</paragraph>
<caption box="[118,1122,1242,1307]" pageId="12" pageNumber="113" type="caption">
<paragraph box="[118,1122,1242,1307]" pageId="12" pageNumber="113" type="caption">
<pageBreakToken pageId="12" pageNumber="113" start="start">FIGURES</pageBreakToken>
<paragraph id="D8207598929C6A06EA874415CCB93013" box="[118,1120,1475,1550]" pageId="11" pageNumber="112" type="mainText">Form 7 (Figs 36-37, 57). Worker measurements (n=20): HL: 1.38-1.65, HW: 1.19-1.44, CI: 85-92, SL: 0.92-1.13, SI: 72-80, PW: 0.95-1.18, WL: 1.64-2.07, NL: 0.65-0.81, NW: 0.79-1.05, NH: 0.95-1.17, DNI: 119-137, LNI: 139-152.</paragraph>
<caption id="A99B2F0AF0E8E1F1247FE3FFC0257FF2" box="[118,1122,1242,1307]" pageId="12" pageNumber="113" type="caption">
<paragraph id="F70475AEEF70E6D69F17461C41059C2F" box="[118,1122,1242,1307]" pageId="12" pageNumber="113" type="caption">
<pageBreakToken id="F30613C7524477518D0210B727ADC388" pageId="12" pageNumber="113" start="start">FIGURES</pageBreakToken>
30-39. Full-face view of the head and lateral view of the body of the worker caste of
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei</taxonomicName>
<taxonomicName id="160B7D391F2FA76A392192530E2765BA" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei</taxonomicName>
and its various forms; Figs 30-31: Form 4, CASENT0034474; Figs 32-33: Form 5, BLF0976(40)-2; Figs 34-35: Form 6, CASENT0027983.
</paragraph>
</caption>
<paragraph box="[118,1120,1341,1557]" pageId="12" pageNumber="113" type="mainText">
<paragraph id="773659C70A11902E76FBA63503C82675" box="[118,1120,1341,1557]" pageId="12" pageNumber="113" type="mainText">
As in form 6 but workers tend to have smaller body size, broader than long basal flagellar segments, straight anterior clypeal margin, short and robust mesotibiae, and less pubescence on abdominal tergite 4. In dorsal view, petiole node nearly subquadrate, with straight or slightly concave posterior margin and broadly rounded anterior margin; reddish brown to dark brown in color with lighter gaster and appendages. In the north of Madagascar, workers of this form are slightly sculptured and noticeably shiny, with much longer erect hairs, whereas those from the south possess finer, stronger sculpture and shorter hairs. In this variant, morphological diversity occurs and continuously intergrades into that of form 6. Form 7 dominates the rainforests in northern Madagascar but is absent from the PN Montagne
<normalizedToken originalValue="dAmbre" pageId="12" pageNumber="113">d'Ambre</normalizedToken>
<normalizedToken id="D681C89C9669AC8ABC4A2EB642F3BB97" originalValue="dAmbre" pageId="12" pageNumber="113">d'Ambre</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="114" type="discussion">
<paragraph box="[118,224,118,137]" pageId="13" pageNumber="114" type="mainText">
<pageBreakToken pageId="13" pageNumber="114" start="start">Discussion</pageBreakToken>
<subSubSection id="80A2F434D3FDA87146685F361449E480" pageId="13" pageNumber="114" type="discussion">
<paragraph id="74C7861CCC3CC5FB128E499A20A22ED6" box="[118,224,118,137]" pageId="13" pageNumber="114" type="mainText">
<pageBreakToken id="F196505DAA88BCB4B8E90DDFDDC28168" pageId="13" pageNumber="114" start="start">Discussion</pageBreakToken>
:
</paragraph>
<paragraph box="[118,1121,147,306]" pageId="13" pageNumber="114" type="mainText">
<paragraph id="B7E55375F793520FF3835C6DF7CB83A9" box="[118,1121,147,306]" pageId="13" pageNumber="114" type="mainText">
A better understanding of species boundaries in Malagasy ants is made possible by recent improvements in collecting methods as well as intensive sampling across the entire region. Although morphological differences can distinguish several forms within
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="155D0049AB4FDEA6EA92A157E2E47EC6" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
, enough gradual variation in character states exists across their geographical range to prevent us from hypothesizing separate species for each form. For example, differences in the shape of the posterior margin and the posterior corner of the head, density and size of punctures, and other body sculpture characteristics are not consistent across the region.
</paragraph>
<paragraph box="[118,1122,315,587]" pageId="13" pageNumber="114" type="mainText">
<paragraph id="E4BEC5F231F8F3471BC4FDCF767CB436" box="[118,1122,315,587]" pageId="13" pageNumber="114" type="mainText">
Madagascar is characterized by complex topography and habitat heterogeneity. A mountain plateau on the island creates habitats ranging from montane forest at high altitudes and dry forest on its western slopes to lowland rainforest and littoral forests on its eastern side. Continuous environmental gradients exist between each type of habitat. Across these ecological gradients and ecotones, a widespread species like
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="EB38EBEB9ECE4CEFF683D65EA1067375" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
would be expected to show some variation, especially if the suitiable habitats are isolated (
<bibRefCitation author="Raxworthy, C. J. &amp; Nussbaum, R. A." box="[118,1122,142,208]" journalOrPublisher="Journal of Zoology" pageId="40" pageNumber="141" pagination="525 - 558" part="235" publicationUrl="http://dx.doi.org" title="Systematics, speciation and biogeography of the dwarf chameleons (Brookesia; Reptilia, Squamata, Chamaeleontidae) of northern Madagascar" type="journal article" year="1995">Raxworthy &amp; Nussbaum 1995</bibRefCitation>
<bibRefCitation id="149AF76667D919E062C8EB75FC33D5B0" author="Raxworthy, C. J. &amp; Nussbaum, R. A." box="[118,1122,142,208]" journalOrPublisher="Journal of Zoology" pageId="40" pageNumber="141" pagination="525 - 558" part="235" publicationUrl="http://dx.doi.org" title="Systematics, speciation and biogeography of the dwarf chameleons (Brookesia; Reptilia, Squamata, Chamaeleontidae) of northern Madagascar" type="journal article" year="1995">Raxworthy &amp; Nussbaum 1995</bibRefCitation>
;
<bibRefCitation author="Ricklefs, R. E." box="[118,1122,214,256]" journalOrPublisher="Ecological Letters" pageId="40" pageNumber="141" pagination="1 - 15" part="7" publicationUrl="http://dx.doi.org/10.1046/j.1461-0248.2003.00554.x" title="A comprehensive framework for global patterns in biodiversity" type="journal article" year="2004">Ricklefs 2004</bibRefCitation>
<bibRefCitation id="7225EB55EF78001F41C75622CB5626F3" author="Ricklefs, R. E." box="[118,1122,214,256]" journalOrPublisher="Ecological Letters" pageId="40" pageNumber="141" pagination="1 - 15" part="7" publicationUrl="http://dx.doi.org/10.1046/j.1461-0248.2003.00554.x" title="A comprehensive framework for global patterns in biodiversity" type="journal article" year="2004">Ricklefs 2004</bibRefCitation>
;
<bibRefCitation author="Wiens, J. J. &amp; Donoghue, M. J." box="[118,1122,1053,1094]" journalOrPublisher="Trends in Ecology and Evolution" pageId="40" pageNumber="141" pagination="639 - 644" part="19" publicationUrl="http://dx.doi.org/10.1016/j.tree.2004.09.011" title="Historical biogeography, ecology, and species richness" type="journal article" year="2004">Wiens &amp; Donoghue 2004</bibRefCitation>
<bibRefCitation id="E57AB33EE8EBD5F83380C42A42FF865C" author="Wiens, J. J. &amp; Donoghue, M. J." box="[118,1122,1053,1094]" journalOrPublisher="Trends in Ecology and Evolution" pageId="40" pageNumber="141" pagination="639 - 644" part="19" publicationUrl="http://dx.doi.org/10.1016/j.tree.2004.09.011" title="Historical biogeography, ecology, and species richness" type="journal article" year="2004">Wiens &amp; Donoghue 2004</bibRefCitation>
;
<bibRefCitation author="Wilme, L. &amp; Goodman, S. M. &amp; Ganzhorn, J. U." box="[118,1122,1101,1142]" journalOrPublisher="Science" pageId="40" pageNumber="141" pagination="1063 - 1065" part="312" publicationUrl="http://dx.doi.org/10.1126/science.1122806" title="Biogeographic evolution of Madagascar's microendemic biota" type="journal article" year="2006">
<normalizedToken originalValue="Wilmé" pageId="13" pageNumber="114">Wilme</normalizedToken>
<bibRefCitation id="6CB068B8BDEFA1A924E97FED6EBABF67" author="Wilme, L. &amp; Goodman, S. M. &amp; Ganzhorn, J. U." box="[118,1122,1101,1142]" journalOrPublisher="Science" pageId="40" pageNumber="141" pagination="1063 - 1065" part="312" publicationUrl="http://dx.doi.org/10.1126/science.1122806" title="Biogeographic evolution of Madagascar's microendemic biota" type="journal article" year="2006">
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et al. 2006
</bibRefCitation>
;
<bibRefCitation author="Boumans, L. &amp; Vieites, D. R. &amp; Glaw, F. &amp; Vences, M." box="[118,1122,1033,1098]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="38" pageNumber="139" pagination="822 - 839" part="45" publicationUrl="http://dx.doi.org/10.1016/j.ympev.2007.05.028" title="Geographical patterns of deep mitochondrial differentiation in widespread Malagasy reptiles" type="journal article" year="2007">Boumans et al. 2007</bibRefCitation>
<bibRefCitation id="D2CE13565FF95BAE774E65A0B0AB0440" author="Boumans, L. &amp; Vieites, D. R. &amp; Glaw, F. &amp; Vences, M." box="[118,1122,1033,1098]" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="38" pageNumber="139" pagination="822 - 839" part="45" publicationUrl="http://dx.doi.org/10.1016/j.ympev.2007.05.028" title="Geographical patterns of deep mitochondrial differentiation in widespread Malagasy reptiles" type="journal article" year="2007">Boumans et al. 2007</bibRefCitation>
;
<bibRefCitation author="Smith, S. A. &amp; Nieto Montes deOca, A. &amp; Reeder, T. W. &amp; Wiens, J. J." box="[118,1122,406,471]" journalOrPublisher="Evolution" pageId="40" pageNumber="141" pagination="1188 - 1207" part="61" publicationUrl="http://dx.doi.org/10.1111/j.1558-5646.2007.00085.x" title="A phylogenetic perspective on elevational species richness patterns in middle American treefrogs: why so few species in lowland tropical forests" type="journal article" year="2007">Smith et al. 2007</bibRefCitation>
<bibRefCitation id="BE6DF0B7AE1B60DAD43FFABCB7568AE0" author="Smith, S. A. &amp; Nieto Montes deOca, A. &amp; Reeder, T. W. &amp; Wiens, J. J." box="[118,1122,406,471]" journalOrPublisher="Evolution" pageId="40" pageNumber="141" pagination="1188 - 1207" part="61" publicationUrl="http://dx.doi.org/10.1111/j.1558-5646.2007.00085.x" title="A phylogenetic perspective on elevational species richness patterns in middle American treefrogs: why so few species in lowland tropical forests" type="journal article" year="2007">Smith et al. 2007</bibRefCitation>
;
<bibRefCitation author="Vences, M. &amp; Wollenberg, K. C. &amp; Vieites, D. R. &amp; Lees, D. C." box="[118,1121,717,759]" journalOrPublisher="Trends in Ecology and Evolution" pageId="40" pageNumber="141" pagination="456 - 465" part="24" publicationUrl="http://dx.doi.org/10.1016/j.tree.2009.03.011" title="Madagascar as a model region of species diversification" type="journal article" year="2009">Vences et al. 2009</bibRefCitation>
<bibRefCitation id="EA2A2C924DEEDFA411C87FE0B4FAD7CD" author="Vences, M. &amp; Wollenberg, K. C. &amp; Vieites, D. R. &amp; Lees, D. C." box="[118,1121,717,759]" journalOrPublisher="Trends in Ecology and Evolution" pageId="40" pageNumber="141" pagination="456 - 465" part="24" publicationUrl="http://dx.doi.org/10.1016/j.tree.2009.03.011" title="Madagascar as a model region of species diversification" type="journal article" year="2009">Vences et al. 2009</bibRefCitation>
). An example of isolated populations is the form 2 located in the relict montane rainforests of RS Ambohijanahary, which is nested in extensive dry habitats in the western slope of Madagascar. The area may have served as a habitat refugia during periods of climatic fluctuations (
<bibRefCitation author="Wilme, L. &amp; Goodman, S. M. &amp; Ganzhorn, J. U." box="[118,1122,1101,1142]" journalOrPublisher="Science" pageId="40" pageNumber="141" pagination="1063 - 1065" part="312" publicationUrl="http://dx.doi.org/10.1126/science.1122806" title="Biogeographic evolution of Madagascar's microendemic biota" type="journal article" year="2006">
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<bibRefCitation id="A3F6EDB2E49D75AEE40B822A65818B99" author="Wilme, L. &amp; Goodman, S. M. &amp; Ganzhorn, J. U." box="[118,1122,1101,1142]" journalOrPublisher="Science" pageId="40" pageNumber="141" pagination="1063 - 1065" part="312" publicationUrl="http://dx.doi.org/10.1126/science.1122806" title="Biogeographic evolution of Madagascar's microendemic biota" type="journal article" year="2006">
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et al. 2006
</bibRefCitation>
;
<bibRefCitation author="Yoder, A. D. &amp; Heckman, K." box="[118,1122,1220,1285]" editor="Fleagle, J." journalOrPublisher="Kluwer, New York, New York" pageId="40" pageNumber="141" pagination="255 - 268" title="Mouse lemur phylogeography revises a model of ecogeographic constraint in Madagascar" type="book chapter" volumeTitle="Primate Biogeography: Progress and Prospects" year="2006">Yoder and Heckman 2006</bibRefCitation>
<bibRefCitation id="F029DCE7DCFF72745923B104B002C816" author="Yoder, A. D. &amp; Heckman, K." box="[118,1122,1220,1285]" editor="Fleagle, J." journalOrPublisher="Kluwer, New York, New York" pageId="40" pageNumber="141" pagination="255 - 268" title="Mouse lemur phylogeography revises a model of ecogeographic constraint in Madagascar" type="book chapter" volumeTitle="Primate Biogeography: Progress and Prospects" year="2006">Yoder and Heckman 2006</bibRefCitation>
;
<bibRefCitation author="Wollenberg, K. C. &amp; Vieites, D. R. &amp; van der Meijden, A. &amp; Glaw, F. &amp; Cannatella, D. C. &amp; Vences, M." box="[118,1122,1149,1214]" journalOrPublisher="Evolution" pageId="40" pageNumber="141" pagination="1890 - 1907" part="62" publicationUrl="http://dx.doi.org/10.1111/j.1558-5646.2008.00420.x" title="Patterns of endemism and species richness in Malagasy cophyline frogs support a key role of mountainous areas for speciation" type="journal article" year="2008">Wollenberg et al. 2008</bibRefCitation>
<bibRefCitation id="ED7B8AED27C0743F5C60FD5FF0DD8233" author="Wollenberg, K. C. &amp; Vieites, D. R. &amp; van der Meijden, A. &amp; Glaw, F. &amp; Cannatella, D. C. &amp; Vences, M." box="[118,1122,1149,1214]" journalOrPublisher="Evolution" pageId="40" pageNumber="141" pagination="1890 - 1907" part="62" publicationUrl="http://dx.doi.org/10.1111/j.1558-5646.2008.00420.x" title="Patterns of endemism and species richness in Malagasy cophyline frogs support a key role of mountainous areas for speciation" type="journal article" year="2008">Wollenberg et al. 2008</bibRefCitation>
;
<bibRefCitation author="Pearson, R. G. &amp; Raxworthy, C. J." box="[118,1122,1532,1573]" journalOrPublisher="Evolution" pageId="39" pageNumber="140" pagination="959 - 967" part="63" publicationUrl="http://dx.doi.org/10.1111/j.1558-" title="The evolution of local endemism in Madagascar: watershed versus climatic gradient hypotheses evaluated by null biogeographic models" type="journal article" year="2009">Pearson and Raxworthy 2009</bibRefCitation>
<bibRefCitation id="56C35A45A2A7543CBDD647683963559D" author="Pearson, R. G. &amp; Raxworthy, C. J." box="[118,1122,1532,1573]" journalOrPublisher="Evolution" pageId="39" pageNumber="140" pagination="959 - 967" part="63" publicationUrl="http://dx.doi.org/10.1111/j.1558-" title="The evolution of local endemism in Madagascar: watershed versus climatic gradient hypotheses evaluated by null biogeographic models" type="journal article" year="2009">Pearson and Raxworthy 2009</bibRefCitation>
;
<bibRefCitation author="Vences, M. &amp; Wollenberg, K. C. &amp; Vieites, D. R. &amp; Lees, D. C." box="[118,1121,717,759]" journalOrPublisher="Trends in Ecology and Evolution" pageId="40" pageNumber="141" pagination="456 - 465" part="24" publicationUrl="http://dx.doi.org/10.1016/j.tree.2009.03.011" title="Madagascar as a model region of species diversification" type="journal article" year="2009">Vences et al., 2009</bibRefCitation>
<bibRefCitation id="3F96D20013EC96D17B3D1FB11218AF03" author="Vences, M. &amp; Wollenberg, K. C. &amp; Vieites, D. R. &amp; Lees, D. C." box="[118,1121,717,759]" journalOrPublisher="Trends in Ecology and Evolution" pageId="40" pageNumber="141" pagination="456 - 465" part="24" publicationUrl="http://dx.doi.org/10.1016/j.tree.2009.03.011" title="Madagascar as a model region of species diversification" type="journal article" year="2009">Vences et al., 2009</bibRefCitation>
).
</paragraph>
<paragraph box="[118,1121,597,728]" pageId="13" pageNumber="114" type="mainText">
<paragraph id="B591D4A11A4A41B456EA512B803E58C0" box="[118,1121,597,728]" pageId="13" pageNumber="114" type="mainText">
To better understand the morphological diversity of
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="A90D24AF62DA298720B0E03BC0851F41" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
, especially in a historical context, we hope future studies will include molecular sequence data and other morphological data such as male characters which have proven useful in distinguishing cryptic species of ants (
<bibRefCitation author="LaPolla, J. S. &amp; Hawkes, P. G. &amp; Fisher, B. L." box="[118,1122,1196,1238]" journalOrPublisher="Zootaxa" pageId="39" pageNumber="140" pagination="10 - 36" part="3110" title="Monograph of Nylanderia (Hymenoptera: Formicidae) of the World, Part I: Nylanderia in the Afrotropics" type="journal article" year="2011">LaPolla et al. 2011</bibRefCitation>
<bibRefCitation id="68AD7E83E6A0D524472E60CD8C84621B" author="LaPolla, J. S. &amp; Hawkes, P. G. &amp; Fisher, B. L." box="[118,1122,1196,1238]" journalOrPublisher="Zootaxa" pageId="39" pageNumber="140" pagination="10 - 36" part="3110" title="Monograph of Nylanderia (Hymenoptera: Formicidae) of the World, Part I: Nylanderia in the Afrotropics" type="journal article" year="2011">LaPolla et al. 2011</bibRefCitation>
). The use of molecular sequence data of
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="232849B57519D75B3B2C2B4FED3AF95E" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
will also help evaluate the biogeographical mechanisms that underlie the evolutionary diversification of taxa in Madagascar (as reviewed by
<bibRefCitation author="Vences, M. &amp; Wollenberg, K. C. &amp; Vieites, D. R. &amp; Lees, D. C." box="[118,1121,717,759]" journalOrPublisher="Trends in Ecology and Evolution" pageId="40" pageNumber="141" pagination="456 - 465" part="24" publicationUrl="http://dx.doi.org/10.1016/j.tree.2009.03.011" title="Madagascar as a model region of species diversification" type="journal article" year="2009">Vences et al. 2009</bibRefCitation>
<bibRefCitation id="C92AC53B152AD4D78BA6A659B1A00058" author="Vences, M. &amp; Wollenberg, K. C. &amp; Vieites, D. R. &amp; Lees, D. C." box="[118,1121,717,759]" journalOrPublisher="Trends in Ecology and Evolution" pageId="40" pageNumber="141" pagination="456 - 465" part="24" publicationUrl="http://dx.doi.org/10.1016/j.tree.2009.03.011" title="Madagascar as a model region of species diversification" type="journal article" year="2009">Vences et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="114" type="distribution">
<paragraph box="[118,353,761,779]" pageId="13" pageNumber="114" type="mainText">Distribution and biology:</paragraph>
<paragraph box="[118,1122,789,1146]" pageId="13" pageNumber="114" type="mainText">
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei</taxonomicName>
<subSubSection id="80802B0C4492DAF7E8631B3855439F8B" pageId="13" pageNumber="114" type="distribution">
<paragraph id="0FD8FAD1468834CC1F65CDD88A20B91B" box="[118,353,761,779]" pageId="13" pageNumber="114" type="mainText">Distribution and biology:</paragraph>
<paragraph id="05A20C324C51D4A1503BE5F1A8CB7ABD" box="[118,1122,789,1146]" pageId="13" pageNumber="114" type="mainText">
<taxonomicName id="AFE3DAAA57BD2D7734E1B3D2616C3F6E" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">Pachycondyla cambouei</taxonomicName>
is an endemic, widespread species found from the north throughout the center and the south of Madagascar. It occupies mainly mesic forests at higher and lower altitudes, and also occurs in littoral and transitional forest habitats. This species dominates the representatives of the genus
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="genus">Pachycondyla</taxonomicName>
<taxonomicName id="60B850C8D9B28CDBDE58CA4B835132BF" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="genus">Pachycondyla</taxonomicName>
across its distribution range. Two or more different forms within the species occur sympatrically in several localities; these sympatric groupings include forms 2 and 6; forms 3 and 6; forms 3 and 7; forms 6 and 7; forms 1, 4, and 7; and forms 1, 5, and 7. Although this species is generally terrestrial, a few specimens have been found foraging on low vegetation. Forms 2-7 occur generally on the mountaintops, while forms 1 and 4 also can be found in lowland forests. Most forage through leaf litter and rarely on the forest floor. Nest series were discovered for most forms, collected frequently from rotten logs, under stones, in the ground, in rotten tree stumps, and between root mat layers, but seldom in dead twigs or branches above the ground. Workers of
<taxonomicName class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
<taxonomicName id="95C652B0BD5F163C25A521EF4ED42F8F" class="Insecta" family="Formicidae" genus="Pachycondyla" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="cambouei">P. cambouei</taxonomicName>
often play dead when their nests are disturbed. Colonies of forms 5 and 6 have not yet been found, which suggests they may be soil dwellers. The smaller eyes, longer erect hairs, and abundant pubescence in form 6 are also suggestive of an underground life style.
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="120" pageId="13" pageNumber="114" type="materials_examined">
<paragraph box="[118,361,1179,1197]" pageId="13" pageNumber="114" type="mainText">Other material examined:</paragraph>
<paragraph box="[118,1122,1207,1592][118,1122,118,1572][118,1122,118,1572]" lastPageId="15" lastPageNumber="116" pageId="13" pageNumber="114" type="mainText">
<subSubSection id="E9DAC9B28E730B546DDBB9508CBABA8E" lastPageId="19" lastPageNumber="120" pageId="13" pageNumber="114" type="materials_examined">
<paragraph id="140884AE34D3DF1314795D75927E3F5C" box="[118,361,1179,1197]" pageId="13" pageNumber="114" type="mainText">Other material examined:</paragraph>
<paragraph id="2FC9CD0DFD126A0EC53EEAAB55EE5D09" box="[118,1122,1207,1592][118,1122,118,1572][118,1122,118,1572]" lastPageId="15" lastPageNumber="116" pageId="13" pageNumber="114" type="mainText">
Form 1. MADAGASCAR: Antananarivo: Antananarivo (
<normalizedToken originalValue="Camboué" pageId="13" pageNumber="114">Camboue</normalizedToken>
<normalizedToken id="43E2BC8E272F35ADD2D86977D415AB16" originalValue="Camboué" pageId="13" pageNumber="114">Camboue</normalizedToken>
) (MNHN); Tam Perinet; B.M. 1983-201, 27 Apr-3 May 1983 (J.S. Noyes, M.C. Day) (BMNH); Amparihibe (SB); 18; BMNH (E)2003-61, -15.0353, 49.5839 (K.A. Jackson, D. Carpenter) (BMNH); Andrangoloaka (Sikora) (MHNG); Antananarivo; Museum Paris, Grandidier 1893 (
<normalizedToken originalValue="Camboué" pageId="13" pageNumber="114">Camboue</normalizedToken>
<normalizedToken id="4CA8C806181C5848135CE9F4AEB3D18D" originalValue="Camboué" pageId="13" pageNumber="114">Camboue</normalizedToken>
) (MNHN);
<normalizedToken originalValue="Imérina" pageId="13" pageNumber="114">Imerina</normalizedToken>
<normalizedToken id="AB8D7E07558DF9185872B1EFDB7F0E1C" originalValue="Imérina" pageId="13" pageNumber="114">Imerina</normalizedToken>
; Museum Paris, Grandidier (
<normalizedToken originalValue="Camboué" pageId="13" pageNumber="114">Camboue</normalizedToken>
<normalizedToken id="FE7C42F88DC52CB980B2C09FF9FF20EE" originalValue="Camboué" pageId="13" pageNumber="114">Camboue</normalizedToken>
) (MNHN); Imerina (
<normalizedToken originalValue="Camboué" pageId="13" pageNumber="114">Camboue</normalizedToken>
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) (MHNG); (W.M. Wheeler) (MCZC); 25 km NNE Ankazobe, -18.1, 47.18333, 1500 m, montane rainforest (P.S. Ward) (MCZC); 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe, -18.47333, 47.96, 1300 m, montane rainforest (B.L. Fisher et al.) (CASC); Ankazobe, -18.31617, 47.11583, 1241 m, urban/garden (B.L. Fisher et al.) (CASC); Kaloy, -18.58998, 47.65102, 1423 m, disturbed montane rainforest (B.L. Fisher et al.) (CASC); RNI Sohisika, Sohisika 24.6 km NNE Ankazobe, -18.10322, 47.18692, 1464 m, gallery montane forest (B.L. Fisher et al.) (CASC); RS Ambohitantely, F
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, 20.9 km 72° NE Ankazobe, -18.22528, 47.28683, 1410 m, montane rainforest (Fisher, Griswold et al.) (CASC); RS Ambohitantely, F
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, Jardin Botanique, 24.1 km 59° NE Ankazobe, -18.17139, 47.28182, 1620 m, montane rainforest (Fisher, Griswold et al.) (CASC); Tsinjoarivo, E Ambatolampy (W.L. Brown) (MCZC); Angavokely, -18.93333, 47.75 (B. Pettersson) (PSWC); Vanjamanitra, Antananarivo, 1380 m (J.M. Betsch) (MCZC); Bongolava, Pref.
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(A. Peyrieras) (MCZC); Antsiranana: 9.2 km WSW Befingotra, RS Anjanaharibe-Sud, -14.75, 49.46667, 1200 m, montane rainforest (B.L. Fisher) (CASC); Ampasindava, F d'Ambilanivy, 3.9 km 181° S Ambaliha, -13.79861, 48.16167, 600 m, rainforest (Fisher, Griswold et al.) (CASC); Ankobahoba, 32.3 km N Ambanja, -13.39166, 48.48249, 41 m, disturbed littoral rainforest (B.L. Fisher et al.) (CASC); Betaolana Forest, along Bekona River, -14.52996, 49.44039, 880 m, rainforest (B.L. Fisher et al.) (CASC); F Ambanitaza, 26.1 km 347° Antalaha, -14.67933, 50.18367, 240 m, rainforest (B.L. Fisher) (CASC); F d'Anabohazo, 21.6 km 247° WSW Maromandia, -14.30889, 47.91433, 120 m, tropical dry forest (Fisher, Griswold et al.) (CASC); F de Binara, 9.1 km 233° SW Daraina, -13.26333, 49.60333, 650-800 m, rainforest (B.L. Fisher et al.) (CASC); F de Binara, 9.4 km 235° SW Daraina, -13.26333, 49.6, 1100 m, montane rainforest (B.L. Fisher et al.) (CASC); F d'Orangea, 3.6 km 128° SE Remena, -12.25889, 49.37467, 90 m, littoral rainforest (Fisher, Griswold et al.) (CASC); Marojejy
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I. #12, -14.44533, 49.78564, 375 m, rainforest (G.D. Alpert) (MCZC); Marojejy, tributary Manantenina R., - 14.43333, 49.75, 750 m (Quinter &amp; Nguyen) (CASC); Nosy Be, RNI Lokobe, 6.3 km 112° ESE Hellville, - 13.41933, 48.33117, 30 m, rainforest (Fisher, Griswold et al.) (CASC); PN Marojejy, Antranohofa, 26.6 km 31° NNE Andapa, 10.7 km 318° NW Manantenina, -14.44333, 49.74333, 1325 m, montane rainforest (B.L. Fisher) (CASC); PN Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, -14.435, 49.76, 775 m, rainforest (B.L. Fisher) (CASC); PN Marojejy, Manantenina River, 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina, -14.43667, 49.775, 450 m, rainforest (B.L. Fisher et al.) (CASC); PN Montagne d'Ambre, 3.6 km 235° SW Joffreville, -12.53444, 49.1795, 925 m, montane rainforest (Fisher, Griswold et al.) (CASC); PN Montagne d'Ambre, Antomboka, -12.51269, 49.17807, 970 m, montane rainforest (B.L. Fisher et al.) (CASC); PN Montagne d'Ambre,
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, -12.58132, 49.13368, 1110 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Manongarivo, 12.8 km 228° SW Antanambao, -13.97667, 48.42333, 780 m, rainforest (B.L. Fisher) (CASC); Makirovana Forest: -14.1667, 49.95, 715 m; -14.1604, 49.9522, 550 m; -14.1707, 49.9541, 415 m, rainforest (B.L. Fisher et al.) (CASC); Fianarantsoa: 29.5 km WNW Tolanaro, Vasiha Mt., -24.93694, 46.75361, 200 m, rainforest (K.C. Emberton et al) (CASC); 45 km S Ambalavao, -22.21667, 47.01667, 720-785 m, rainforest (B.L. Fisher) (CASC); 8.0 km NE Ivohibe, -22.42167, 46.89833, 1200 m, montane rainforest (Sylvain) (CASC); 9.0 km NE Ivohibe, -22.42667, 46.93833, 900 m, rainforest (Sylvain) (CASC); F d'Ambalagoavy Nord, Ikongo, Ambatombe, -21.8275, 47.33889, 625 m (R. Harin'Hala &amp; M.E. Irwin) (CASC); F d'Atsirakambiaty, 7.6 km 285° WNW Itremo, -20.59333, 46.56333, 1550 m, montane rainforest (Fisher, Griswold et al.) (CASC); F de Vevembe, 66.6 km 293° Farafangana, -22.791, 47.18183, 600 m, rainforest, transition to montane forest (B.L. Fisher et al.) (CASC); PN Ranomafana, Ambatolahy Forest, montane rainforest (Alpert et al.) (MCZC); PN Andringitra, F Ravaro 12.5 km SW Antanifotsy, -22.21167, 46.845, 1500-1800 m (S. Razafimandimby) (CASC); PN Befotaka-Midongy, Papango 28.5 km S Midongy-Sud, Mount Papango, -23.84083, 46.9575, 1250 m, montane rainforest (B.L. Fisher et al.) (CASC); PN Ranomafana, Vatoharanana River, 4.1 km 231° SW Ranomafana, -21.29, 47.43333, 1100 m, montane rainforest (Fisher, Griswold et al.) (CASC); RS Ivohibe: 8.0 km E Ivohibe, -22.48333, 46.96833, 1200 m, montane rainforest (Sylvain) (CASC); RS Ivohibe, 7.5 km ENE Ivohibe, -22.47, 46.96, 900 m, rainforest (Sylvain) (CASC); Ranomafana Nat. Park, Maharira Forest, -21, 47, 1190 m, montane forest (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Miaranony Forest, -21.25, 47.41667, 700 m, montane forest (Emile Rajeriarison) (MCZC); RF d'Agnalazaha, Mahabo, 42.9 km 215° Farafangana, -23.19383, 47.723, 20 m, littoral rainforest (B.L. Fisher et al.) (CASC); RS Manombo 24.5 km 228° Farafangana, -23.01583, 47.719, 30 m, rainforest (B.L. Fisher et al.) (CASC); Mahajanga: 10 km N Mahatsinjo, -17.84028, 47.02472, 1355 m, tropical dry forest (Alpert et al.) (MCZC); RS Marotandrano, Marotandrano 48.3 km S Mandritsara, -16.28322, 48.81443, 865 m, transition humid forest (B.L. Fisher et al.) (CASC); Toamasina: 33 km south of Moramanga, Rte. d'Anosibe, - 19.25, 48.21667, forest humus and litter (A. Peyrieras) (MCZC); 5.3 km SSE Ambanizana, Andranobe, -15.66667, 49.96667, 425 m, rainforest (B.L. Fisher) (CASC); 6.3 km S Ambanizana, Andranobe, -15.68131, 49.958, 25 m, rainforest (B.L. Fisher) (CASC); 6.9 km NE Ambanizana, Ambohitsitondroina, -15.56667, 50, 825 m, rainforest (B.L. Fisher) (CASC); Ambanizana, PN Masoala, -15.57167, 50.00611, 800-897 m, montane rainforest (D. Andriamalala, D. Silva et al.) (CASC); Ambanizana, PN Masoala, -15.57222, 50.00694, 930-1110 m, montane rainforest (D. Andriamalala, D. Silva et al.) (CASC); Amparihibe (SB), 18 and 31, BMNH (E) 2003-61, [-15.0353, 49.5839] (K.A. Jackson, D. Carpenter) (BMNH); Ambatovy, 12.4 km NE Moramanga, -18.84963, 48.2947, 1010 m, montane rainforest (B.L. Fisher et al.) (CASC); Ambatovy, 12.4 km NE Moramanga, -18.84773, 48.29568, 1000 m, grassland (B.L. Fisher et al.) (CASC); Ambatovy, 12.4 km NE Moramanga, -18.85813, 48.28488, 1040 m, grassland (B.L. Fisher et al.) (CASC); Ambatovy, 12.4 km NE Moramanga, -18.84963, 48.2947, 1010 m, montane
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(B.L.Fisher et al.) (CASC); F Ambatovy, 14.3 km 57° Moramanga, -18.85083, 48.32, 1075 m, montane rainforest (B.L. Fisher) (CASC) Analalava, 7.0 km 255° Mahavelona, -17.7095, 49.454, 50 m, littoral rainforest (B.L. Fisher et al.) (CASC); Analamay, -18.80623, 48.33707, 1068 m, montane rainforest (Malagasy ant team) (CASC); Beforona, 29 km east of Perinet, -18.96667, 48.58333, 500 m, forest (A. Peyrieras) (MCZC); Bevolota 17.1 km N Andasibe, -18.77071, 48.43164, 995 m, montane rainforest (B.L. Fisher et al.) (CASC);
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(A. Pauly) (CASC); FC Andriantantely, -18.695, 48.81333, 530 m, rainforest (H.J. Ratsirarson) (CASC); FC Sandranantitra, -18.04833, 49.09167, 450 m, rainforest (H.J. Ratsirarson) (CASC); Ile Sainte Marie, F Kalalao, 9.9 km 34° Ambodifotatra, -16.9225, 49.88733, 100 m, rainforest (B.L. Fisher et al.) (CASC); La Mandraka, 1280 m, montane forest (W.L. &amp; D.E. Brown) (MCZC); Mahavelona (Foulpointe), -17.66667, 49.5, forest (A. Pauly) (CASC); Manakambahiny, near Vavatenina Forest, -17.46667, 49.35 (A. Pauly) (CASC); Montagne d'Akirindro 7.6 km 341° NNW Ambinanitelo, -15.28833, 49.54833, 600 m, rainforest (Fisher, Griswold et al.) (CASC); Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, -15.18833, 49.615, 470 m, rainforest (Fisher, Griswold et al.) (CASC); Morarano-Chrome, 25 km W,
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(A. Pauly) (CASC); PN Mantadia, -18.79167, 48.42667, 895 m, rainforest (H.J. Ratsirarson) (CASC); PN Andasibe-Mantadia, F de Mantadia, 25.7 km 248° Moramanga, -18.81402, 48.43028, 1040 m, rainforest (F.N. Raharimalala, B. Blaimer) (CASC); PN Zahamena, - 17.73359, 48.72625, 950 m, rainforest (B.L. Fisher et al.) (CASC); PN Zahamena, Besaky River, -17.75244, 48.85321, 760 m, rainforest (B.L. Fisher et al.) (CASC); PN Zahamena, Onibe River, -17.75908, 48.85468, 780 m, rainforest (B.L. Fisher et al.) (CASC); PN Zahamena, Sahavorondrano River, -17.75257, 48.85725, 765 m, rainforest (B.L. Fisher et al.) (CASC); PN Zahamena, Tetezambatana Forest, near junction of Nosivola and Manakambahiny Rivers, -17.74298, 48.72936, 860 m, rainforest (B.L. Fisher et al.) (CASC); PN Mananara-Nord, 7.1 km 261° Antanambe, -16.455, 49.7875, 225 m, rainforest (B.L. Fisher et al.) (CASC); Parcelle K7 Tampolo, - 17.28333, 49.41667, 10 m, littoral forest (Malagasy ant team) (CASC); Parcelle K9 Tampolo, -17.175, 49.268, 10 m, littoral forest (Malagasy ant team) (CASC); Perinet (E.S. Ross) (MCZC); Perinet and vic., -18.93333, 48.41667, rainforest (W.L. Brown) (MCZC); Perinet; B.M. 1983-201 (J.S. Noyes, M.C. Day) (BMNH); Res. Ambodiriana, 4.8 km 306° Manompana, along Manompana River, -16.67233, 49.70117, 125 m, rainforest (B.L. Fisher et al.) (CASC); Res. Perinet, rainforest (L.Bartolozzi, S.Tiati &amp; C.Raharimina) (MCZC); RNI Betampona, Camp Rendrirendry 34.1 km 332° Toamasina, -17.924, 49.19967, 390-550 m, rainforest (B.L. Fisher et al.) (CASC); RNI Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina, -17.88667, 49.2025, 520 m, rainforest (B.L. Fisher et al.) (CASC); RNI Betampona, Betampona 35.1 km NW Toamasina, -17.91801, 49.20074, 500 m, rainforest (B.L. Fisher et al.) (CASC); RNI Betampona, 34.08 km 332° Toamasina, -17.91977, 49.20039, 525 m, rainforest (B.L. Fisher) (CASC); RS Ambatovaky, Sandrangato River, -16.81753, 49.29498, 360 m, rainforest (B.L. Fisher et al.) (CASC); RS Ambatovaky, Sandrangato River, -16.77274, 49.26551, 450 m, rainforest (B.L. Fisher et al.) (CASC); RS Ambatovaky, Sandrangato River, -16.7755, 49.26427, 430 m, rainforest (B.L. Fisher et al.) (CASC); RS Ambatovaky, Sandrangato River, -16.7633, 49.26692, 520 m, rainforest (B.L. Fisher et al.) (CASC); RS Ambatovaky, Sandrangato River, -16.76912, 49.26704, 475 m, rainforest (B.L. Fisher et al.) (CASC); SF Tampolo, 10 km NNE Fenoarivo Atn., -17.2825, 49.43, 10 m, littoral rainforest (B.L. Fisher) (CASC); Sahafina forest 11.4 km W Brickaville, -18.81445, 48.96205, 140 m, rainforest (B.L. Fisher et al.) (CASC); SF Analamazaotra, Analamazaotra 1.3 km S Andasibe, -18.38466, 48.41271, 980 m, montane rainforest (B.L. Fisher et al.) (CASC); Torotorofotsy, -18.87082, 48.34737, 1070 m, montane rainforest, marsh edge (Malagasy ant team) (CASC); Torotorofotsy, -18.87467, 48.3725, 960 m, rainforest (Woodhead &amp; Vences) (CASC); 1 km W Andampibe, Cap Masoala, -15.69361, 50.18139, 125 m, lowland forest (G.D. Alpert) (MCZC);
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. Ambodiriana 45 km NW Toamasina (P. Rabeson) (MCZC); vic Andasibe (=Perinet), 950 m, forest (W.L. &amp; D.E. Brown) (MCZC); Toliara: 29.5 km WNW Tolanaro, Vasiha Mt., -24.92306, 46.74083, 700 m, rainforest (K.C. Emberton et al.) (CASC); 10 km NW Enakara,
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Andohahela, -24.56667, 46.81667, 420 m, rainforest (B.L. Fisher) (CASC); 11 km NW Enakara,
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. Andohahela, -24.56667, 46.83333, 800 m, rainforest (B.L. Fisher) (CASC); 13 km NW Enakara,
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. Andohahela, -24.55, 46.8, 1250 m, montane rainforest (B.L. Fisher) (CASC); 13 km NW Enakara,
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Andohahela, -24.55, 46.8, 1280 m, montane rainforest (B.L. Fisher) (CASC); 2.7 km WNW 302° Ste. Luce, - 24.77167, 47.17167, 20 m, littoral rainforest (J.-Baptiste) (CASC); 6 km SSW Eminiminy, Res. Andohahela, - 24.73333, 46.8, 330 m, rainforest (E. Rajeriarison) (MCZC); Andohahela, -24.77639, 46.70528, 320 m, tropical dry forest (P. Rabeson) (MCZC); Andohahela, Parcel #1 versante E., 300 m, rainforest (L. Bartolozzi, S. Tiati &amp; C. Raharimina) (MCZC); Res. Andohahela, 6 km ESE Imonty, -24.84444, 46.75361, 1000 m, rainforest (P. Rabeson) (MCZC); Res.Andohahela, 9 km SSW Eminiminy, -24.75, 46.78333, 500 m, rainforest (E. Rajeriarison) (MCZC);
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Analavelona, 29.2 km 343° NNW Mahaboboka, -22.675, 44.19, 1100 m, montane rainforest (Fisher, Griswold et al.) (CASC); FC Analavelona, 33.2 km 344° NNW Mahaboboka, -22.64333, 44.17167, 1300 m, montane rainforest (Fisher, Griswold et al.) (CASC); F Ivohibe 55.0 km N Tolagnaro, -24.569, 47.204, 200 m, rainforest (B.L. Fisher et al.) (CASC); Manatantely, 8.9 km NW Tolagnaro, -24.9815, 46.92567, 100 m, rainforest (B.L. Fisher et al.) (CASC); PN Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro, -24.7585, 46.85367, 275 m, rainforest (B.L. Fisher et al.) (CASC); PN Andohahela, Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro, -24.76389, 46.75167, 900 m, montane rainforest (Fisher, Griswold Arthropod Team) (CASC); PN d'Andohahela, Manampanihy River, 5.4 km 113° ESE Mahamavo, 36.7 km 343° NNW Tolagnaro, -24.76389, 46.76683, 650 m, rainforest (Fisher, Griswold Arthropod Team) (CASC); Res. Andohahela, 5 km SSW Eminiminy, - 24.73333, 46.8, 200 m (E. Rajeriarison) (MCZC); RS Ambohijanahary, F d'Ankazotsihitafototra, 34.6 km 314° NW Ambaravaranala, -18.26, 45.41833, 1100 m, montane rainforest (Fisher, Griswold et al.) (CASC); RS Ambohijanahary, F d'Ankazotsihitafototra, 35.2 km 312° NW Ambaravaranala, -18.26667, 45.40667, 1050 m, montane rainforest (Fisher, Griswold et al.) (CASC); RS Kalambatritra, Ambinanitelo, -23.4502, 46.45658, 1325 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Kalambatritra, Ampanihy, -23.4635, 46.4631, 1270 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Kalambatritra, Ampanihy, -23.463, 46.47057, 1269 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Kalambatritra, Ampanihy, -23.4635, 46.4631, 1270 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Kalambatritra, Ampanihy, -23.463, 46.47057, 1269 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Kalambatritra, Befarara, -23.4178, 46.4478, 1390 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Kalambatritra, Betanana, -23.4144, 46.459, 1360 m, montane rainforest (B.L. Fisher et al.) (CASC).
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Form 2. Toliara: RS Ambohijanahary, F d'Ankazotsihitafototra, 35.2 km 312° NW Ambaravaranala, - 18.26667, 45.40667, 1050 m, montane rainforest (Fisher, Griswold et al.) (CASC). Form 3. Antsiranana: PN Montagne d'Ambre [1st campsite], -12.51444, 49.18139, 960 m, rainforest (M.E. Irwin, E.I. Schlinger, R. Harin'Hala) (CASC); PN Montagne d'Ambre [Petit Lac road], -12.52028, 49.17917, 1125 m, rainforest (R. Harin'Hala) (CASC); Fianarantsoa: 14 km W of park headquarters, Ranomafana National Park, - 21.22707, 47.36874, 1100 m, cloud forest (M.E. Irwin &amp; E.I. Schlinger) (CASC); 2 km W Andrambovato, along river Tatamaly, -21.51167, 47.41, 1075 m, montane rainforest (B.L. Fisher et al.) (CASC); 3 km W Ranomafana, nr. Ifandiana, -21.25, 47.41667, 950 m, forest (P.S. Ward) (PSWC); 3 km WNW Ranomafana, nr. Ifanadiana, - 21.25, 47.43333, 840 m, roadside (P.S. Ward) (PSWC); 45 km S. Ambalavao, -22.21667, 47.01667, 785 m, rainforest (B.L. Fisher) (CASC); 7.6 km 122
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Form 2. Toliara: RS Ambohijanahary, F d'Ankazotsihitafototra, 35.2 km 312° NW Ambaravaranala, - 18.26667, 45.40667, 1050 m, montane rainforest (Fisher, Griswold et al.) (CASC). Form 3. Antsiranana: PN Montagne d'Ambre [1st campsite], -12.51444, 49.18139, 960 m, rainforest (M.E. Irwin, E.I. Schlinger, R. Harin'Hala) (CASC); PN Montagne d'Ambre [Petit Lac road], -12.52028, 49.17917, 1125 m, rainforest (R. Harin'Hala) (CASC); Fianarantsoa: 14 km W of park headquarters, Ranomafana National Park, - 21.22707, 47.36874, 1100 m, cloud forest (M.E. Irwin &amp; E.I. Schlinger) (CASC); 2 km W Andrambovato, along river Tatamaly, -21.51167, 47.41, 1075 m, montane rainforest (B.L. Fisher et al.) (CASC); 3 km W Ranomafana, nr. Ifandiana, -21.25, 47.41667, 950 m, forest (P.S. Ward) (PSWC); 3 km WNW Ranomafana, nr. Ifanadiana, - 21.25, 47.43333, 840 m, roadside (P.S. Ward) (PSWC); 45 km S. Ambalavao, -22.21667, 47.01667, 785 m, rainforest (B.L. Fisher) (CASC); 7.6 km 122
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Kianjavato, FC Vatovavy, -21.4, 47.94, 175 m, rainforest (B.L. Fisher et al.) (CASC); 7 km W Ranomafana, 1100 m (C. Kremen, R. Van Epps &amp; L. Rasabo) (MCZC); 7 km W Ranomafana, -21.26667, 47.41667, 1000 m, rainforest (M. Stebbins, W.E. Steiner et al.) (MCZC); 9.0 km NE Ivohibe, -22.42667, 46.93833, 900 m, rainforest (Sylvain) (CASC); Belle Vue trail, Ranomafana National Park, Fianarantsoa Prov., -21.2665, 47.42017, 1020 m, mixed tropical forest (R. Harin'Hala) (CASC); Isalo Nat. Park, Ranohira, Canyon de Sinze, -22.48333, 45.55, 800 m, gallery forest (E. Rajeriarison) (MCZC); Nat. Park Ranomafana, Ambatolahy Forest (E. Rajeriarison) (MCZC); PN Ranomafana (L. Bartolozzi, S. Tiati, &amp; C. Raharimina) (MCZC); PN Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango, -23.83517, 46.96367, 940 m, rainforest (B.L. Fisher et al.) (CASC); PN Befotaka-Midongy, Papango 28.5 km S Midongy-Sud, Mount Papango, -23.84083, 46.9575, 1250 m, montane rainforest (B.L. Fisher et al.) (CASC); PN Ranomafana, Sahamalaotra River, 6.6 km 310° NW Ranomafana, -21.23667, 47.39667, 1150 m, montane rainforest (Fisher, Griswold et al.) (CASC); PN Ranomafana, Vatoharanana River, 4.1 km 231° SW Ranomafana, -21.29, 47.43333, 1100 m, montane rainforest (Fisher, Griswold et al.) (CASC); PN Ranomafana Talatakely, -21.24833, 47.425, 900 m, between main bridge and park entrance (J.S. Schweikert) (CASC); PN Ranomafana: Talatakely, -21.24833, 47.42667, in guava forest (C.E. Griswold, D.H. Kavanaugh, N.D. Penny, M.J. Raherilalao, J.S. Ranorianarisoa, J.S. Schweikert) (CASC); P.N. Ranomafana: Talatakely, -21.24833, 47.42667, in bamboo forest (C.E. Griswold, D.H. Kavanaugh, N.D. Penny, M.J. Raherilalao, J.S. Ranorianarisoa, J.S. Schweikert) (CASC); RS Ivohibe 8.0 km E Ivohibe, -22.48333, 46.96833, 1200 m, montane rainforest (Sylvain) (CASC); RS Ivohibe, 6.5 km ESE Ivohibe, - 22.49667, 46.955, 1575 m, montane rainforest (Sylvain) (CASC); RS Ivohibe, 7.5 km ENE Ivohibe, -22.47, 46.96, 900 m, rainforest (Sylvain) (CASC); radio tower, Ranomafana National Park, Fianarantsoa Prov., -21.25083, 47.40717, 1130 m, forest edge, mixed tropical forest, open area (M. Irwin, R. Harin'Hala) (CASC); Ranomafana, - 21.25, 47.36667 (A. Pauly) (CASC); Ranomafana (A. Kingman) (MCZC); Ranomafana Nat. Park, 700 m, rainforest (W.E. Steiner) (MCZC); Ranomafana Nat. Park, 7 km W, 900 m, montane rainforest (W.E. Steiner) (MCZC); Ranomafana Nat. Park, Fivondronana Ifanadiana Talatakely, 1100 m (J.E. Cadle) (MCZC); Ranomafana
<pageBreakToken pageId="17" pageNumber="118" start="start">Nat</pageBreakToken>
. Park, near Andranoroa, low tree forest (I. Constable) (MCZC); Ranomafana Nat. Park, near research cabin, - 21.25, 47.41667 (V.P. Lee, K.J. Rivardo, leg.) (CASC); Ranomafana Nat. Park, SE research cabin, montane rainforest (P. Rabeson) (MCZC); Ranomafana Nat. Park, Talatakely Forest, montane rainforest (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Valoloaka Forest, 1200 m, montane forest (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Vohiparara, 1160 m, montane rainforest (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Vohiparara Sahamalaotra, -21.24444, 47.39694, rainforest (P. Rabeson) (MCZC); Ranomafana Nat. Park, Mahirira Forest, -21, 47, 1350 m (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Miaranony Forest, -21.25, 47.41667, 800 m, montane forest (E. Rajeriarison) (MCZC); Ranomafana, Miaranony village (A. Kingman) (MCZC); research cabin at Talatakely, Ranomafana National Park, -21.25041, 47.41945, 900 m, mixed tropical forest (M.E. Irwin &amp; E.I. Schlinger) (CASC); Vohiparara broken bridge, Fianarantsoa Prov., -21.22617, 47.36983, 1110 m, high altitude rainforest (R. Harin'Hala) (CASC); Vohiparara, 13 km W Ranomafana, foresta secondaria (L. Bartolozzi, S. Tiati, &amp; C. Raharimina) (MCZC); Toamasina: 19 km ESE Maroantsetra, -15.48333, 49.9, 350 m, rainforest (P.S. Ward) (PSWC); 1 km SSW Andasibe, -18.93333, 48.41667, 420 m, forest (P.S. Ward) (MCZC); 1 km SSW Andasibe, -18.93333, 48.41667, 420 m, forest (P.S. Ward) (PSWC); 1 km SSW Andasibe (=Perinet) -18.93333, 48.41667, 920 m, rainforest (P. S. Ward) (MCZC); 6 km ESE Andasibe (=Perinet), -18.95, 48.46667, 900 m, rainforest (P.S. Ward) (MCZC); 6 km ESE Andasibe(-
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. Park, near Andranoroa, low tree forest (I. Constable) (MCZC); Ranomafana Nat. Park, near research cabin, - 21.25, 47.41667 (V.P. Lee, K.J. Rivardo, leg.) (CASC); Ranomafana Nat. Park, SE research cabin, montane rainforest (P. Rabeson) (MCZC); Ranomafana Nat. Park, Talatakely Forest, montane rainforest (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Valoloaka Forest, 1200 m, montane forest (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Vohiparara, 1160 m, montane rainforest (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Vohiparara Sahamalaotra, -21.24444, 47.39694, rainforest (P. Rabeson) (MCZC); Ranomafana Nat. Park, Mahirira Forest, -21, 47, 1350 m (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Miaranony Forest, -21.25, 47.41667, 800 m, montane forest (E. Rajeriarison) (MCZC); Ranomafana, Miaranony village (A. Kingman) (MCZC); research cabin at Talatakely, Ranomafana National Park, -21.25041, 47.41945, 900 m, mixed tropical forest (M.E. Irwin &amp; E.I. Schlinger) (CASC); Vohiparara broken bridge, Fianarantsoa Prov., -21.22617, 47.36983, 1110 m, high altitude rainforest (R. Harin'Hala) (CASC); Vohiparara, 13 km W Ranomafana, foresta secondaria (L. Bartolozzi, S. Tiati, &amp; C. Raharimina) (MCZC); Toamasina: 19 km ESE Maroantsetra, -15.48333, 49.9, 350 m, rainforest (P.S. Ward) (PSWC); 1 km SSW Andasibe, -18.93333, 48.41667, 420 m, forest (P.S. Ward) (MCZC); 1 km SSW Andasibe, -18.93333, 48.41667, 420 m, forest (P.S. Ward) (PSWC); 1 km SSW Andasibe (=Perinet) -18.93333, 48.41667, 920 m, rainforest (P. S. Ward) (MCZC); 6 km ESE Andasibe (=Perinet), -18.95, 48.46667, 900 m, rainforest (P.S. Ward) (MCZC); 6 km ESE Andasibe(-
<normalizedToken id="DCEDE4C7F891EB3021868C769503A277" originalValue="Périnet" pageId="17" pageNumber="118">Perinet</normalizedToken>
), -18.95, 48.46666667, 900 m, rainforest (P.S. Ward) (PSWC); Andranoaria, 15 km S Anosibe An 'alam. (C. Raharimina) (MCZC); FC Andriantantely, -18.695, 48.81333, 530 m, rainforest (H.J. Ratsirarson) (CASC); FC Didy, -18.19833, 48.57833, 960 m, rainforest (H.J. Ratsirarson) (CASC); FC Sandranantitra, -18.04833, 49.09167, 450 m, rainforest (H.J. Ratsirarson) (CASC); PN Mantadia, -18.79167, 48.42667, 895 m, rainforest (H.J. Ratsirarson) (CASC); Perinet and vic., -18.93333, 48.41667, rainforest (W.L. Brown) (MCZC); Perinet and vic., -18.93333, 48.41667, rainforest (W.L. Brown) (MCZC); Res. Perinet, Andasibe, 870 m (Alpert et al.) (MCZC); vic Andasibe (=Perinet), 950 m (W.L. Brown) (MCZC); Toliara: F Ivohibe 55.6 km N Tolagnaro, -24.56167, 47.20017, 650 m, rainforest (B.L. Fisher et al.) (CASC); PN Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro, -24.7585, 46.85367, 275 m, rainforest (B.L. Fisher et al.) (CASC).
</paragraph>
<paragraph box="[118,1122,822,1066]" pageId="17" pageNumber="118" type="mainText">Form 4. Antsiranana: 6.5 km SSW Befingotra, RS Anjanaharibe-Sud, -14.75, 49.5, 875 m, rainforest (B.L. Fisher) (CASC); 9.2 km WSW Befingotra, RS Anjanaharibe-Sud, -14.75, 49.46667, 1280 m, montane rainforest (B.L. Fisher) (CASC); 9.2 km WSW Befingotra, RS Anjanaharibe-Sud, -14.75, 49.46667, 1200 m, montane rainforest (B.L. Fisher) (CASC); Makirovana Forest: -14.1651, 49.9477, 900 m, montane rainforest; -14.1604, 49.9522, 550 m, rainforest (B.L. Fisher et al.) (CASC); Toamasina: Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, -15.18833, 49.615, 470 m, rainforest (Fisher, Griswold et al.) (CASC); Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, -15.17833, 49.635, 1100 m, montane rainforest (Fisher, Griswold et al.) (CASC); Montagne d'Akirindro 7.6 km 341° NNW Ambinanitelo, -15.28833, 49.54833, 600 m, rainforest (Fisher, Griswold et al.) (CASC).</paragraph>
<paragraph box="[118,1121,1075,1150]" pageId="17" pageNumber="118" type="mainText">Form 5. Toamasina: 6.9 km NE Ambanizana, Ambohitsitondroina, -15.56667, 50, 825 m, rainforest (B.L. Fisher) (CASC); 5.3 km SSE Ambanizana, Andranobe, -15.66667, 49.96667, 425 m, rainforest (B.L. Fisher) (CASC).</paragraph>
<paragraph box="[118,1122,1159,1584]" pageId="17" pageNumber="118" type="mainText">
<paragraph id="9E222935857B91810609555FA9D3A339" box="[118,1122,822,1066]" pageId="17" pageNumber="118" type="mainText">Form 4. Antsiranana: 6.5 km SSW Befingotra, RS Anjanaharibe-Sud, -14.75, 49.5, 875 m, rainforest (B.L. Fisher) (CASC); 9.2 km WSW Befingotra, RS Anjanaharibe-Sud, -14.75, 49.46667, 1280 m, montane rainforest (B.L. Fisher) (CASC); 9.2 km WSW Befingotra, RS Anjanaharibe-Sud, -14.75, 49.46667, 1200 m, montane rainforest (B.L. Fisher) (CASC); Makirovana Forest: -14.1651, 49.9477, 900 m, montane rainforest; -14.1604, 49.9522, 550 m, rainforest (B.L. Fisher et al.) (CASC); Toamasina: Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, -15.18833, 49.615, 470 m, rainforest (Fisher, Griswold et al.) (CASC); Montagne d'Anjanaharibe, 19.5 km 27° NNE Ambinanitelo, -15.17833, 49.635, 1100 m, montane rainforest (Fisher, Griswold et al.) (CASC); Montagne d'Akirindro 7.6 km 341° NNW Ambinanitelo, -15.28833, 49.54833, 600 m, rainforest (Fisher, Griswold et al.) (CASC).</paragraph>
<paragraph id="3CFFEB84D04E55D8214C6E9E36CBCDCD" box="[118,1121,1075,1150]" pageId="17" pageNumber="118" type="mainText">Form 5. Toamasina: 6.9 km NE Ambanizana, Ambohitsitondroina, -15.56667, 50, 825 m, rainforest (B.L. Fisher) (CASC); 5.3 km SSE Ambanizana, Andranobe, -15.66667, 49.96667, 425 m, rainforest (B.L. Fisher) (CASC).</paragraph>
<paragraph id="A8E3C885E1977C4CD48023A60ABD653D" box="[118,1122,1159,1584]" pageId="17" pageNumber="118" type="mainText">
Form 6. Antananarivo: 3 km 41° NE Andranomay, 11.5 km 147° SSE Anjozorobe, -18.47333, 47.96, 1300 m, montane rainforest (Fisher, Griswold et al.) (CASC); 2 km W Ivato (G.D. Alpert) (MCZC); Ambohidratrimo, - 18.821, 47.44183, 1362 m, urban garden (B.L. Fisher et al.) (CASC); F de galerie, Andranorovitra, 24.0 km NNE Ankazobe, -18.11243, 47.19757, 1491 m, disturbed gallery montane forest (B.L. Fisher et al.) (CASC); F de galerie, Telomirahavavy, 23.4 km NNE Ankazobe, -18.12167, 47.20627, 1520 m, disturbed gallery montane forest (B.L. Fisher et al.) CASC); Ilafy, -18.85415, 47.56575, 1385 m, urban garden (B.L. Fisher et al.) (CASC); RNI Sohisika, Sohisika 24.6 km NNE Ankazobe, -18.10322, 47.18692, 1464 m, gallery montane forest (B.L. Fisher et al.) (CASC); RS Ambohitantely, F
<normalizedToken originalValue="dAmbohitantely" pageId="17" pageNumber="118">d'Ambohitantely</normalizedToken>
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, 20.9 km 72° NE Ankazobe, -18.22528, 47.28683, 1410 m, montane rainforest (Fisher, Griswold et al.) (CASC); RS Ambohitantely, F
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, Jardin Botanique, 24.1 km 59° NE d Ankazobe, -18.17139, 47.28182, 1620 m, montane rainforest (Fisher, Griswold et al.) (CASC); Antsiranana: Bemanevika, 10 km N de Sambava souspref. de Bealanana, -14.13333, 50.11667 (A. Peyrieras) (MCZC); PN Marojejy, Manantenina River, 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina, -14.43667, 49.775, 450 m, rainforest (B.L. Fisher et al.) (CASC); PN montagne d'Ambre, 3.6 km 235° SW Joffreville, - 12.53444, 49.1795, 925 m, montane rainforest (Fisher, Griswold et al.) (CASC); RS Fianarantsoa: 2 km W Andrambovato, along River Tatamaly, -21.51167, 47.41, 1075 m, montane rainforest (B.L. Fisher et al.) (CASC); 43 km S Ambalavao,
<normalizedToken originalValue="Rés" pageId="17" pageNumber="118">Res</normalizedToken>
. Andringitra, -22.23333, 47, 825 m, rainforest (B.L. Fisher) (CASC); 7.6 km 122
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. Andringitra, -22.23333, 47, 825 m, rainforest (B.L. Fisher) (CASC); 7.6 km 122
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</paragraph>
<paragraph box="[118,1122,118,1383]" pageId="18" pageNumber="119" type="mainText">
<pageBreakToken pageId="18" pageNumber="119" start="start">Kianjavato</pageBreakToken>
<paragraph id="23227072E7A540D316375598D748D70C" box="[118,1122,118,1383]" pageId="18" pageNumber="119" type="mainText">
<pageBreakToken id="4A32895E6B219D1D4599239818CB5211" pageId="18" pageNumber="119" start="start">Kianjavato</pageBreakToken>
, FC Vatovavy, -21.4, 47.94, 175 m, rainforest (B.L. Fisher et al.) (CASC); 7 km W Ranomafana, 1000 m, rainforest (W.E. Steiner) (MCZC); 8 km E Kianjavato, Vatovavy Forest, -21.39306, 47.94194, 170 m, lowland forest (Alpert et al.) (MCZC); 9.0 km NE Ivohibe, -22.42667, 46.93833, 900 m, rainforest (Sylvain) (CASC); Belle Vue trail, Ranomafana National Park, -21.2665, 47.42017, 1020 m, mixed tropical forest (R. Harin'Hala) (CASC); F d'Ambalagoavy Nord, Ikongo, Ambatombe, -21.8275, 47.33889, 625 m (R. Harin'Hala &amp; M.E. Irwin) (CASC); F d'Atsirakambiaty, 7.6 km 285° WNW Itremo, -20.59333, 46.56333, 1550 m, montane rainforest (Fisher, Griswold et al.) (CASC); F de Vevembe, 66.6 km 293° Farafangana, -22.791, 47.18183, 600 m, rainforest, transition to montane forest (B.L. Fisher et al.) (CASC); Isalo Nat. Park, Ranohira, Canyon de Sinze, -22.48333, 45.55, 800 m, gallery forest (E. Rajeriarison) (MCZC); PN Andringitra, F Ravaro 12.5 km SW Antanifotsy, -22.21167, 46.845, 1500-1800 m (S. Razafimandimby) (CASC); PN Ranomafana, Vatoharanana River, 4.1 km 231° SW Ranomafana, -21.29, 47.43333, 1100 m, montane rainforest (Fisher, Griswold et al.) (CASC); Parc naturel communautaire, 28.1 km SW Ambositra, -20.78493, 47.17328, 1727 m, disturbed montane rainforest along roadside (B.L. Fisher et al.) (CASC); RS Ivohibe 8.0 km E Ivohibe, -22.48333, 46.96833, 1200 m, montane rainforest (Sylvain) (CASC); RS Ivohibe, 7.5 km ENE Ivohibe, -22.47, 46.96, 900 m, rainforest (Sylvain) (CASC); Ranomafana Nat. Park, Talatakely Forest, montane rainforest (E. Rajeriarison) (MCZC); Ranomafana Nat. Park, Miaranony Forest, -21.25, 47.41667, 800 m, montane forest (E. Rajeriarison) (MCZC); RS Manombo 24.5 km 228° Farafangana, -23.01583, 47.719, 30 m, rainforest (B.L. Fisher et al.) (CASC); Mahajanga: RS Marotandrano, Marotandrano 48.3 km S Mandritsara, -16.28322, 48.81443, 865 m, transition humid forest (B.L. Fisher et al.) (CASC); Toamasina: 16 km S Moramanga, -19.08333, 48.23333, 950 m, rainforest (P.S. Ward) (PSWC); 6.9 km NE Ambanizana, Ambohitsitondroina, -15.56667, 50.0095, 825 m, rainforest (B.L. Fisher) (CASC); Ambanizana, PN Masoala, - 15.57167, 50.00611, 900-950 m, montane rainforest (D. Andriamalala, D. Silva et al.) (CASC); FC Andriantantely, -18.695, 48.81333, 530 m, rainforest (H.J. Ratsirarson) (CASC); FC Didy, -18.19833, 48.57833, 960 m, rainforest (H.J. Ratsirarson) (CASC); F Ambatovy, 14.3 km 57° Moramanga, -18.85083, 48.32, 1075 m, montane rainforest (Malagasy ant team) (CASC); F d'Analava Mandrisy, 5.9 km 195 Antanambe, -16.48567, 49.847, 10 m, littoral rainforest (B.L. Fisher et al.) (CASC); Montagne d'Akirindro 7.6 km 341° NNW Ambinanitelo, -15.28833, 49.54833, 600 m, rainforest (Fisher, Griswold et al.) (CASC); Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, -15.18833, 49.615, 470 m, rainforest (Fisher, Griswold et al.) (CASC); PN Mantadia, -18.79167, 48.42667, 895 m, rainforest (H.J. Ratsirarson) (CASC); PN Marojejy, - 14.43817, 49.774, 488 m, rainforest (Rin'Ha, Irwin) (CASC); RNI Betampona, Camp Rendrirendry 34.1 km 332° Toamasina, -17.924, 49.19967, 390 m, rainforest (B.L. Fisher et al.) (CASC); SF Analamazaotra, Analamazaotra 1.3 km S Andasibe, -18.38466, 48.41271, 980 m, montane rainforest (B.L. Fisher et al.) (CASC); Torotorofotsy, - 18.87082, 48.34737, 1070 m, montane rainforest, marsh edge (Malagasy ant team) (CASC); Toliara: 11 km NW Enakara,
<normalizedToken originalValue="Rés" pageId="18" pageNumber="119">Res</normalizedToken>
<normalizedToken id="20DDAE558ADA87AF4D955AE3BA517CFC" originalValue="Rés" pageId="18" pageNumber="119">Res</normalizedToken>
. Andohahela, -24.56667, 46.83333, 800 m, rainforest (B.L. Fisher) (CASC); 13 km NW Enakara, Res. Andohahela, -24.55, 46.8, 1250 m, montane rainforest (B.L. Fisher) (CASC); FC Analavelona, 29.2 km 343° NNW Mahaboboka, -22.675, 44.19, 1100 m, montane rainforest (Fisher, Griswold et al.) (CASC); FC Analavelona, 29.4 km 343° NNW Mahaboboka, -22.675, 44.18667, 1050 m, montane rainforest (Fisher, Griswold et al.) (CASC); F de Petriky, 12.5 km W 272° Tolagnaro, -25.06167, 46.87, 10 m, littoral rainforest (B.L. Fisher) (CASC); Mandena, 8.4 km NNE 30° Tolagnaro, -24.95167, 47.00167, 20 m, littoral rainforest (B.L. Fisher) (CASC); PN Andohahela, Col de Tanatana, 33.3 km NW Tolagnaro, -24.7585, 46.85367, 275 m, rainforest (B.L. Fisher et al.) (CASC); PN Andohahela, Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro, -24.76389, 46.75167, 900 m, montane rainforest (Fisher-Griswold Arthropod Team) (CASC); RS Ambohijanahary, F d'Ankazotsihitafototra, 35.2 km 312° NW Ambaravaranala, -18.26667, 45.40667, 1050 m, montane rainforest (Fisher, Griswold et al.) (CASC); RS Kalambatritra, Ambinanitelo, -23.4502, 46.45658, 1325 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Kalambatritra, Ampanihy, -23.4635, 46.4631, 1270 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Kalambatritra, Ampanihy, -23.463, 46.47057, 1269 m, montane rainforest (B.L. Fisher et al.) (CASC); RS Kalambatritra, Befarara, -23.4178, 46.4478, 1390 m, montane rainforest (B.L. Fisher et al.) (CASC).
</paragraph>
<paragraph box="[118,1122,1391,1573][118,1122,118,760]" lastPageId="19" lastPageNumber="120" pageId="18" pageNumber="119" type="mainText">
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Form 7. Antsiranana: 6.5 km SSW Befingotra, RS. Anjanaharibe-Sud, -14.75, 49.5, 875 m, rainforest (B.L. Fisher) (CASC); 9.2 km WSW Befingotra, RS. Anjanaharibe-Sud, -14.75, 49.46667, 1200 m, montane rainforest (B.L. Fisher) (CASC); Ambondrobe, 41.1 km 175° Vohemar, -13.71533, 50.10167, 10 m, littoral rainforest (B.L. Fisher) (CASC); Bemanevika, 10 km N de Sambava souspref. de Bealanana, -14.13333, 50.11667 (A. Peyrieras) (MCZC); F de Binara, 9.4 km 235° SW Daraina, -13.26333, 49.6, 1100 m, montane rainforest (B.L. Fisher et al.) (CASC); PN Marojejy, Manantenina River, 28.0 km 38° NE Andapa, 8.2 km 333° NNW Manantenina, -14.43667, 49.775, 450 m, rainforest (B.L. Fisher et al.) (CASC); RS Manongarivo, 12.8 km 228° SW Antanambao, -
<pageBreakToken pageId="19" pageNumber="120" start="start">13.97667</pageBreakToken>
<pageBreakToken id="F25303085DB35B622434A301488FDA9A" pageId="19" pageNumber="120" start="start">13.97667</pageBreakToken>
, 48.42333, 780 m, rainforest (B.L. Fisher) (CASC); RS Manongarivo, 14.5 km 220° SW Antanambao, - 13.99833, 48.42833, 1175 m, montane rainforest (B.L. Fisher) (CASC); Ambondrobe, 41.1 km 175° Vohemar, - 13.71533, 50.10167, 10 m, littoral rainforest (B.L. Fisher) (CASC); Makirovana Forest, -14.1707, 49.9541, 415 m, rainforest (B.L. Fisher et al.) (CASC); Fianarantsoa: 9.0 km NE Ivohibe, -22.42667, 46.93833, 900 m, rainforest (Sylvain) (CASC); RS Ivohibe 8.0 km E Ivohibe, -22.48333, 46.96833, 1200 m, montane rainforest (Sylvain) (CASC); PN Ranomafana, Vatoharanana River, 4.1 km 231° SW Ranomafana, -21.29, 47.43333, 1100 m, montane rainforest (Fisher, Griswold et al.) (CASC); Belle Vue trail, Ranomafana National Park, -21.2665, 47.42017, 1020 m, mixed tropical forest (R. Harin'Hala) (CASC); Mahajanga: RS Marotandrano, Marotandrano 48.3 km S Mandritsara, -16.28322, 48.81443, 865 m, transitional humid forest (B.L. Fisher et al.) (CASC); Toamasina: 6.9 km NE Ambanizana, Ambohitsitondroina, -15.56667, 50.000, 825 m, rainforest (B.L.Fisher) (CASC); Ambanizana, PN Masoala, -15.57167, 50.00611, 900-950 m, montane rainforest (D. Andriamalala, D. Silva et al.) (CASC); FC Andriantantely, -18.695, 48.81333, 530 m, rainforest (H.J. Ratsirarson) (CASC); FC Didy, -18.19833, 48.57833, 960 m, rainforest (H.J. Ratsirarson) (CASC); FC Sandranantitra, -18.04833, 49.09167, 450 m, rainforest (H.J. Ratsirarson) (CASC); Montagne d'Akirindro 7.6 km 341° NNW Ambinanitelo, -15.28833, 49.54833, 600 m, rainforest (Fisher, Griswold et al.) (CASC); Montagne d'Anjanaharibe, 18.0 km 21° NNE Ambinanitelo, - 15.18833, 49.615, 470 m, rainforest (Fisher, Griswold et al.) (CASC); PN Mantadia, -18.79167, 48.42667, 895 m, rainforest (H.J. Ratsirarson) (CASC); PN Zahamena, Onibe River, -17.75908, 48.85468, 780 m, rainforest (B.L. Fisher et al.) (CASC); Toliara: 11 km NW Enakara,
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<normalizedToken id="D746FECC39888F654765B0274AF73C00" originalValue="Rés" pageId="19" pageNumber="120">Res</normalizedToken>
. Andohahela, -24.56667, 46.83333, 800 m, rainforest (B.L. Fisher) (CASC); 13 km NW Enakara, Res. Andohahela, -24.55, 46.8, 1250 m, montane rainforest (B.L. Fisher) (CASC); PN Andohahela, Col du Sedro, 3.8 km 113° ESE Mahamavo, 37.6 km 341° NNW Tolagnaro, - 24.76389, 46.75167, 900 m, montane rainforest (Fisher-Griswold Arthropod Team) (CASC); 0.7 km WNW 302° Ste. Luce, -24.77167, 47.17167, 20 m, littoral rainforest (J.-Baptiste) (CASC); F de Petriky, 12.5 km W 272° Tolagnaro, -25.06167, 46.87, 10 m, littoral rainforest (B.L. Fisher) (CASC); Mandena, 8.4 km NNE 30° Tolagnaro, -24.95167, 47.00167, 20 m, littoral rainforest (B.L. Fisher) (CASC).
</paragraph>
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<subSubSection id="19DB7972ED24FB9BD323A0B913DFA3C8" pageId="0" pageNumber="123" type="reference_group">
<paragraph id="D266F560097E5679F48216439C0CCB47" pageId="0" pageNumber="123">
<taxonomicName id="18D3615019380EA830B1560A0857FA58" authorityName="Yang &amp; Yang" authorityYear="1987" class="Insecta" family="Tipulidae" genus="Nephrotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nephrotoma didyma" order="Diptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="didyma">Nephrotoma didyma</taxonomicName>
Yang &amp; Yang, 1987: 131. Type localty: China: Xizang, Mainling Co.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="diagnosis">
<paragraph pageId="0" pageNumber="123">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="123">
<subSubSection id="3376217932F869C4101A0046FA1ACF66" pageId="0" pageNumber="123" type="diagnosis">
<paragraph id="4A30C0768A816BA757D6AEACE72DB4D0" pageId="0" pageNumber="123">Diagnosis.</paragraph>
<paragraph id="E2120E2AD7EBD148C66E8ECF3D1CFDFB" pageId="0" pageNumber="123">
Pronotum yellow with a brownish spot on lateral side. Mediotergite yellow with dark-brown posterior margin (Figs
<figureCitation captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">17</figureCitation>
<figureCitation id="A31B23F2748208FAC4DDC7AB4BEB66DF" captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">17</figureCitation>
,
<figureCitation captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">19</figureCitation>
<figureCitation id="48D480F972F4238649A5796CA31FDCC6" captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">19</figureCitation>
). Posterior margin of tergite 9 with a jaw-shaped protuberance; spade-shaped protuberance with two short horns at posterior margin (Fig.
<figureCitation captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">22</figureCitation>
<figureCitation id="B263E2ECD25AD3FE79AF95E8D0E63246" captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">22</figureCitation>
). Posterior margin of sternite 8 with meniscus-shaped protrusion (Figs
<figureCitation captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">20</figureCitation>
<figureCitation id="6F9F862E1DE9DE81C09FAF0086A7C53C" captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">20</figureCitation>
,
<figureCitation captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">23</figureCitation>
<figureCitation id="D9D5C7E2211B7C246FDEB96D4C588E20" captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">23</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123" start="Figures 1719" startId="F5">
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">Figures 17-19.</emphasis>
<taxonomicName authorityName="Yang &amp; Yang" authorityYear="1987" class="Insecta" family="Tipulidae" genus="Nephrotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nephrotoma didyma" order="Diptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="didyma">
<emphasis italics="true" pageId="0" pageNumber="123">Nephrotoma didyma</emphasis>
<caption id="D2831000924C99352ED91F5E413E0298" doi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123" start="Figures 1719" startId="F5">
<paragraph id="321F4E8AD52E0E73D2F150B6FD21E83A" pageId="0" pageNumber="123">
<emphasis id="3A3DC5F7DBC7DD6F518AAC721A2451C7" bold="true" pageId="0" pageNumber="123">Figures 17-19.</emphasis>
<taxonomicName id="C70F6F9482FB80FB7EFAEEE0BB80A71D" authorityName="Yang &amp; Yang" authorityYear="1987" class="Insecta" family="Tipulidae" genus="Nephrotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nephrotoma didyma" order="Diptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="didyma">
<emphasis id="A0D7A46AD4249CD02DCAF61D5F598AAF" italics="true" pageId="0" pageNumber="123">Nephrotoma didyma</emphasis>
</taxonomicName>
Yang &amp; Yang, 1987.
<emphasis bold="true" pageId="0" pageNumber="123">17</emphasis>
<emphasis id="76D7AC1A9EEFEACF662BB8755E8A3AFA" bold="true" pageId="0" pageNumber="123">17</emphasis>
Male habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="123">18</emphasis>
<emphasis id="FDA4B0184E85010520AF2E5FA3CF3C24" bold="true" pageId="0" pageNumber="123">18</emphasis>
wing
<emphasis bold="true" pageId="0" pageNumber="123">19</emphasis>
<emphasis id="53F8D912E8D8803B09DE741683D90C16" bold="true" pageId="0" pageNumber="123">19</emphasis>
head and thorax, dorsal view. Scale bars: 1.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="materials_examined">
<paragraph pageId="0" pageNumber="123">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="123">4 males (CAU), China: Xizang, Yigong, 2017.VI.7, 2236 m, Qicheng Yang (light trap). 3 males (CAU), China: Xizang, Yigong, 2017.V 6, 2274 m, Qicheng Yang (light trap). 6 males (CAU), China: Xizang, 106K, 2017.V.17, 2289 m, Qicheng Yang (light trap).</paragraph>
<subSubSection id="60031D67979474AEFCFDC05545B05653" pageId="0" pageNumber="123" type="materials_examined">
<paragraph id="77772270B953DFD51221FAD12B29FE8E" pageId="0" pageNumber="123">Material examined.</paragraph>
<paragraph id="AD4F97966302689B16C15C65C7B1B533" pageId="0" pageNumber="123">4 males (CAU), China: Xizang, Yigong, 2017.VI.7, 2236 m, Qicheng Yang (light trap). 3 males (CAU), China: Xizang, Yigong, 2017.V 6, 2274 m, Qicheng Yang (light trap). 6 males (CAU), China: Xizang, 106K, 2017.V.17, 2289 m, Qicheng Yang (light trap).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="description">
<paragraph pageId="0" pageNumber="123">Description.</paragraph>
<paragraph pageId="0" pageNumber="123">
<subSubSection id="8DCB30F3591E5DCA8C59FF0293FCA603" pageId="0" pageNumber="123" type="description">
<paragraph id="0816FD88273D8880391779A912BA6BCF" pageId="0" pageNumber="123">Description.</paragraph>
<paragraph id="082F777991A758E50DE92AF383AB04AB" pageId="0" pageNumber="123">
Male (
<emphasis italics="true" pageId="0" pageNumber="123">n</emphasis>
<emphasis id="2F1956F099EF4B9C8468E2E2B3B2DA2B" italics="true" pageId="0" pageNumber="123">n</emphasis>
= 13): body length 9.5-12.0 mm, wing length 10.0-12.0 mm, antenna length 3.0-4.0 mm.
</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" italics="true" pageId="0" pageNumber="123">Head</emphasis>
<paragraph id="2A5CE27DB4CE029F9E7932FEF1B21170" pageId="0" pageNumber="123">
<emphasis id="676232249CB1706D60A3CA0A91613F2C" bold="true" italics="true" pageId="0" pageNumber="123">Head</emphasis>
(Figs
<figureCitation captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">17</figureCitation>
<figureCitation id="5AD04F48F54B1E73B7301B86131FCCBF" captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">17</figureCitation>
,
<figureCitation captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">19</figureCitation>
<figureCitation id="6E8FE81373E5B19ECD7E52A140BA56AC" captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">19</figureCitation>
). Mainly yellow. Vertex with brown spot near inner margin of eye. Occipital marking dark brown, bell-shaped, extended to top of vertex. Frontal tubercle relatively high. Dorsal part of rostrum including nasus brownish black. Hairs on head dark brown. Antenna dark brown except scape yellow and pedicel dark yellow; first flagellomere 1.4 times longer than second one. Proboscis yellow, with brown hairs. Palpus brownish grey, with brown hairs.
</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" italics="true" pageId="0" pageNumber="123">Thorax</emphasis>
<paragraph id="F018B9E5DE9336F4C99441F995585745" pageId="0" pageNumber="123">
<emphasis id="99845174C8E49B7D19BC421D78F5D26E" bold="true" italics="true" pageId="0" pageNumber="123">Thorax</emphasis>
(Figs
<figureCitation captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">17</figureCitation>
<figureCitation id="8C48B720CAB4D0FD967EB4AD73EEF084" captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">17</figureCitation>
,
<figureCitation captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">19</figureCitation>
<figureCitation id="154A04E890C3830C193EA31CDA374460" captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">19</figureCitation>
). Mainly yellow. Pronotum yellow with a brownish spot on lateral side. Prescutum with three black longitudinal stripes; anterior end of lateral prescutal stripe bent outward, outer part brown. Scutum with two large, dark-brown spots. Scutellum dark yellow. Mediotergite yellow with dark-brown posterior margin. Anepisternum and katepisternum pale yellow with yellow lower part. Anepimeron pale yellow. Parascutellum pale brown; anatergite and katatergite pale yellow. Legs yellow except middle portions of femora, tips of tibiae, and tarsi dark brown; hairs dark brown, except those on coxae and trochanters yellow. Wing subhyaline, tinged with light brown; pterostigma pale brown; cell m1 nearly sessile (Fig.
<figureCitation captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">18</figureCitation>
<figureCitation id="4084EC00EACB92ABE5B815790D367833" captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">18</figureCitation>
). Halter with stem brownish grey; knob yellowish brown.
</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" italics="true" pageId="0" pageNumber="123">Abdomen</emphasis>
<paragraph id="0313B911CC30F72013B7C9D47D60D888" pageId="0" pageNumber="123">
<emphasis id="D4D6F3317B58B73115734751FC5B3657" bold="true" italics="true" pageId="0" pageNumber="123">Abdomen</emphasis>
(Fig.
<figureCitation captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">17</figureCitation>
<figureCitation id="C93916B993952314777BD5C81A8310E4" captionStart="Figures 1719" captionStartId="F5" captionText="Figures 17 - 19. Nephrotoma didyma Yang &amp; Yang, 1987. 17 Male habitus, lateral view 18 wing 19 head and thorax, dorsal view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.973.46384.figures17-19" httpUri="https://binary.pensoft.net/fig/459924" pageId="0" pageNumber="123">17</figureCitation>
). Mainly yellow. Abdominal tergites with three brown longitudinal stripes. Hypopygium mainly dark yellow. Abdomen with yellow hairs.
</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" italics="true" pageId="0" pageNumber="123">Hypopygium</emphasis>
<paragraph id="A58AFA05684632B7B6494AB3A146029A" pageId="0" pageNumber="123">
<emphasis id="685F5A21EC31D8E50141DC6B8D613727" bold="true" italics="true" pageId="0" pageNumber="123">Hypopygium</emphasis>
(Figs
<figureCitation captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">20-25</figureCitation>
<figureCitation id="830809F7E0C4CB05A8F37526D47B27CD" captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">20-25</figureCitation>
) brownish yellow. Tergite 9 ellipse with spade-shaped protuberance bearing two short horns at posterior margin; posterolateral margin of tergite 9 with a large jaw-shaped protuberance (Fig.
<figureCitation captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">22</figureCitation>
<figureCitation id="C31A4FBF615D5E09163B5928E57212A6" captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">22</figureCitation>
). Posterior margin of sternite 8 with obvious appendages, meniscus-shaped in lateral view, covered with dense yellow hairs (Fig.
<figureCitation captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">20</figureCitation>
<figureCitation id="E24469310361D7E574C4BEC8F5C685AD" captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">20</figureCitation>
). Outer gonostylus fleshy, spade-shaped, wide at middle, narrowed toward tip (Fig.
<figureCitation captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">21</figureCitation>
<figureCitation id="0DD4D08CD6577120CBB5B5E6AB669987" captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">21</figureCitation>
). Inner gonostylus with large concavity, basally with a posterior projection (Figs
<figureCitation captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">24</figureCitation>
<figureCitation id="15ACBD71EF3699CB316ACCA54693F458" captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">24</figureCitation>
,
<figureCitation captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">25</figureCitation>
<figureCitation id="39DDDEEABC5FF79A728EF5C227C4DBC2" captionStart="Figures 2025" captionStartId="F6" captionText="Figures 20 - 25. Nephrotoma didyma Yang &amp; Yang, 1987. 20 Hypopygium, lateral view 21 outer gonostylus, lateral external view 22 tergite 9, dorsal view, softened 23 hypopygium, ventral view 24 inner gonostylus, lateral external view, softened 25 inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (20, 22, 23); 0.1 mm (21, 24, 25)." figureDoi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123">25</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123" start="Figures 2025" startId="F6">
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">Figures 20-25.</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="123">Nephrotoma didyma</emphasis>
<caption id="CCBB804CED37F8E56C5FACD1F380780D" doi="10.3897/zookeys.973.46384.figures20-25" httpUri="https://binary.pensoft.net/fig/459925" pageId="0" pageNumber="123" start="Figures 2025" startId="F6">
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<emphasis id="D91D92D80E238021520F0F087CB0FF9D" bold="true" pageId="0" pageNumber="123">Figures 20-25.</emphasis>
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<emphasis id="F6ABB2BF8D161BA248575FE0D4FD6EC1" italics="true" pageId="0" pageNumber="123">Nephrotoma didyma</emphasis>
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Yang &amp; Yang, 1987.
<emphasis bold="true" pageId="0" pageNumber="123">20</emphasis>
<emphasis id="7D4A72E87362F1C0F38307ABBF937105" bold="true" pageId="0" pageNumber="123">20</emphasis>
Hypopygium, lateral view
<emphasis bold="true" pageId="0" pageNumber="123">21</emphasis>
<emphasis id="D15A6ACF358DD1D3CF4B720F053DCFCE" bold="true" pageId="0" pageNumber="123">21</emphasis>
outer gonostylus, lateral external view
<emphasis bold="true" pageId="0" pageNumber="123">22</emphasis>
<emphasis id="8425B7FE677EE610BC309496EBEC5962" bold="true" pageId="0" pageNumber="123">22</emphasis>
tergite 9, dorsal view, softened
<emphasis bold="true" pageId="0" pageNumber="123">23</emphasis>
<emphasis id="FDF18AFB8AD3074EF84ECF962D5B47A2" bold="true" pageId="0" pageNumber="123">23</emphasis>
hypopygium, ventral view
<emphasis bold="true" pageId="0" pageNumber="123">24</emphasis>
<emphasis id="CD90B2613D73DBAC7CA2A9206B924450" bold="true" pageId="0" pageNumber="123">24</emphasis>
inner gonostylus, lateral external view, softened
<emphasis bold="true" pageId="0" pageNumber="123">25</emphasis>
<emphasis id="81F1FEFCF814621B91BF486549A6F8FE" bold="true" pageId="0" pageNumber="123">25</emphasis>
inner gonostylus, lateral external view, before softened. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="123">20, 22, 23</emphasis>
<emphasis id="A2E33DE09F1558D271298C69FA8ACBEB" bold="true" pageId="0" pageNumber="123">20, 22, 23</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="123">21, 24, 25</emphasis>
<emphasis id="EDB220C3937D92D07899FA9ED54EAF29" bold="true" pageId="0" pageNumber="123">21, 24, 25</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="distribution">
<paragraph pageId="0" pageNumber="123">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="123">China (Xizang).</paragraph>
<subSubSection id="259357592F08E4E713B1100D4F717F9A" pageId="0" pageNumber="123" type="distribution">
<paragraph id="1AFD0E7F3B872EFEC7F4562A7707D196" pageId="0" pageNumber="123">Distribution.</paragraph>
<paragraph id="085C045903032A80BE08FA81A7ED3148" pageId="0" pageNumber="123">China (Xizang).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="remarks">
<paragraph pageId="0" pageNumber="123">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="123">Hypopygium of this species is very specific; no similar species.</paragraph>
<subSubSection id="05CF62D30B67131FBA858EC7C044CD21" pageId="0" pageNumber="123" type="remarks">
<paragraph id="20D49C186DA2F372B7E5051489EC38DD" pageId="0" pageNumber="123">Remarks.</paragraph>
<paragraph id="980B96E4FD875B053A5CF23D67BF7807" pageId="0" pageNumber="123">Hypopygium of this species is very specific; no similar species.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,152 +1,152 @@
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larval morphology (head and mandibular tusk) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D angle measurement of mandibular tusk E R. distafurcus Bae &amp; McCafferty, 1991 F R. speciosus Eaton, 1881." figureDoi="10.3897/zookeys.1067.72779.figure10" httpUri="https://binary.pensoft.net/fig/604591" pageId="0" pageNumber="57">Figures 10F</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larval morphology (foreleg) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D R. distafurcus Bae &amp; McCafferty, 1991 E R. speciosus Eaton, 1881. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1067.72779.figure11" httpUri="https://binary.pensoft.net/fig/604592" pageId="0" pageNumber="57">, 11E</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. SEMs of egg structures of Rhoenanthus speciosus Eaton, 1881 A general outline of egg B micropyle." figureDoi="10.3897/zookeys.1067.72779.figure13" httpUri="https://binary.pensoft.net/fig/604594" pageId="0" pageNumber="57">, 13</figureCitation>
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<figureCitation id="84CA09CE3FE007887381DCE0FFAD24F0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larval morphology (head and mandibular tusk) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D angle measurement of mandibular tusk E R. distafurcus Bae &amp; McCafferty, 1991 F R. speciosus Eaton, 1881." figureDoi="10.3897/zookeys.1067.72779.figure10" httpUri="https://binary.pensoft.net/fig/604591" pageId="0" pageNumber="57">Figures 10F</figureCitation>
<figureCitation id="DC2FDEF2AF7695383C3C1F973C268928" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larval morphology (foreleg) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D R. distafurcus Bae &amp; McCafferty, 1991 E R. speciosus Eaton, 1881. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1067.72779.figure11" httpUri="https://binary.pensoft.net/fig/604592" pageId="0" pageNumber="57">, 11E</figureCitation>
<figureCitation id="EC2E01A9EA0BF938EC9A4146D9CAF588" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. SEMs of egg structures of Rhoenanthus speciosus Eaton, 1881 A general outline of egg B micropyle." figureDoi="10.3897/zookeys.1067.72779.figure13" httpUri="https://binary.pensoft.net/fig/604594" pageId="0" pageNumber="57">, 13</figureCitation>
<figureCitation id="D81E43C45675317FCCA3EB9078E539B0" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Distribution map of the family Potamanthidae in Thailand." figureDoi="10.3897/zookeys.1067.72779.figure14" httpUri="https://binary.pensoft.net/fig/604595" pageId="0" pageNumber="57">, 14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="materials_examined">
<paragraph pageId="0" pageNumber="57">Materials examined.</paragraph>
<paragraph pageId="0" pageNumber="57">
<materialsCitation collectingDate="2018-04-22" collectorName="B. Boonsoong" country="Thailand" latitude="5.7960835" location="Klong Aika Ding" longLatPrecision="1" longitude="101.83486" specimenCount="5" specimenCount-larva="5" stateProvince="Narathiwat province">
<specimenCount type="larva">5 larvae</specimenCount>
<subSubSection id="2961C838FA34B0905B33601231932506" pageId="0" pageNumber="57" type="materials_examined">
<paragraph id="92706967483EFB802E73850CF3C60098" pageId="0" pageNumber="57">Materials examined.</paragraph>
<paragraph id="480DA029431F828BFEF9C695B089CE43" pageId="0" pageNumber="57">
<materialsCitation id="033368FB34DEE6E50817E94D99A433BE" collectingDate="2018-04-22" collectorName="B. Boonsoong" country="Thailand" latitude="5.7960835" location="Klong Aika Ding" longLatPrecision="1" longitude="101.83486" specimenCount="5" specimenCount-larva="5" stateProvince="Narathiwat province">
<specimenCount id="AFDCA37CC87B86A66A9F5BDD6819F8DC" type="larva">5 larvae</specimenCount>
,
<collectingCountry name="Thailand">Thailand</collectingCountry>
<collectingCountry id="48614EC06FFB7692D6D262FDCB52442D" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion country="Thailand" name="Narathiwat">Narathiwat province</collectingRegion>
<collectingRegion id="C79D907E7A995C5AFF451A0104C3576E" country="Thailand" name="Narathiwat">Narathiwat province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:BB3B5E97D40A5257B3E946F0D0CDDEE5:9549DD2F836FA425CA1A94FDF2631619" country="Thailand" latitude="5.7960835" longLatPrecision="1" longitude="101.83486" name="Klong Aika Ding" stateProvince="Narathiwat province">Klong Aika Ding</location>
<location id="9549DD2F836FA425CA1A94FDF2631619" LSID="urn:lsid:plazi:treatment:BB3B5E97D40A5257B3E946F0D0CDDEE5:9549DD2F836FA425CA1A94FDF2631619" country="Thailand" latitude="5.7960835" longLatPrecision="1" longitude="101.83486" name="Klong Aika Ding" stateProvince="Narathiwat province">Klong Aika Ding</location>
stream,
<geoCoordinate degrees="5" direction="north" minutes="47" orientation="latitude" precision="1" seconds="45.9" value="5.7960835">5°47'45.9&quot;N</geoCoordinate>
<geoCoordinate id="7D1A4BD7D9FC4EF361613E4A2650C555" degrees="5" direction="north" minutes="47" orientation="latitude" precision="1" seconds="45.9" value="5.7960835">5°47'45.9&quot;N</geoCoordinate>
,
<geoCoordinate degrees="101" direction="east" minutes="50" orientation="longitude" precision="1" seconds="05.5" value="101.83486">101°50'05.5&quot;E</geoCoordinate>
<geoCoordinate id="5339A441FC93F1ECD122AB0A88FC5DEA" degrees="101" direction="east" minutes="50" orientation="longitude" precision="1" seconds="05.5" value="101.83486">101°50'05.5&quot;E</geoCoordinate>
,
<collectingDate value="2018-04-22">22.IV.2018</collectingDate>
<collectingDate id="0F1DCA0CC33626567603E6E26BD23F0D" value="2018-04-22">22.IV.2018</collectingDate>
,
<collectorName>B. Boonsoong</collectorName>
<collectorName id="C16706304D9B021028665466D0B3F7E3">B. Boonsoong</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="diagnosis">
<paragraph pageId="0" pageNumber="57">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="57">
<subSubSection id="71005B6FF0C0C97E3B76F300EADDD703" pageId="0" pageNumber="57" type="diagnosis">
<paragraph id="B7ED8366B2332EE90F5EA04660CBE730" pageId="0" pageNumber="57">Diagnosis.</paragraph>
<paragraph id="DD5901C5FB537E4FC03071DCC9C068C2" pageId="0" pageNumber="57">
The larvae of
<taxonomicName authorityName="Eaton" authorityYear="1881" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus speciosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis italics="true" pageId="0" pageNumber="57">Rhoenanthus speciosus</emphasis>
<taxonomicName id="2C486842D126D451289078B963A77263" authorityName="Eaton" authorityYear="1881" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus speciosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis id="9B93FBFE80F4614E8709025CE5CF7241" italics="true" pageId="0" pageNumber="57">Rhoenanthus speciosus</emphasis>
</taxonomicName>
can be distinguished from those of other
<taxonomicName authority="(Rhoenanthus)" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus (Rhoenanthus)" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="subGenus" subGenus="Rhoenanthus">Rhoenanthus (Rhoenanthus)</taxonomicName>
<taxonomicName id="431190DD18C6F30AAD9D1B1C2467DC6B" authority="(Rhoenanthus)" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus (Rhoenanthus)" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="subGenus" subGenus="Rhoenanthus">Rhoenanthus (Rhoenanthus)</taxonomicName>
species based on the following characteristics: i) lateral subapical spine of the mandibular tusk present (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larval morphology (head and mandibular tusk) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D angle measurement of mandibular tusk E R. distafurcus Bae &amp; McCafferty, 1991 F R. speciosus Eaton, 1881." figureDoi="10.3897/zookeys.1067.72779.figure10" httpUri="https://binary.pensoft.net/fig/604591" pageId="0" pageNumber="57">10F</figureCitation>
<figureCitation id="8049C6746DD0427C29B8E4C252BF44BC" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larval morphology (head and mandibular tusk) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D angle measurement of mandibular tusk E R. distafurcus Bae &amp; McCafferty, 1991 F R. speciosus Eaton, 1881." figureDoi="10.3897/zookeys.1067.72779.figure10" httpUri="https://binary.pensoft.net/fig/604591" pageId="0" pageNumber="57">10F</figureCitation>
), ii) absence of medial rounded setae on the mandibular tusk, iii) length of the mandibular tusks ca 1.7-2.3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="4C0E8085C8A8997EC262F2975241A545" originalValue="×">x</normalizedToken>
length of the head, iv) length of the foretibiae ca 1.2-1.23
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="675123BEE5B4EF34A5FBBB27EA5A5D48" originalValue="×">x</normalizedToken>
length of the forefemora and about 2.72-2.82
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="19B6966FE9F162929483B174432BD859" originalValue="×">x</normalizedToken>
length of the foretarsi, v) leg with colour marking as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larval morphology (foreleg) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D R. distafurcus Bae &amp; McCafferty, 1991 E R. speciosus Eaton, 1881. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1067.72779.figure11" httpUri="https://binary.pensoft.net/fig/604592" pageId="0" pageNumber="57">11E, and vi</figureCitation>
<figureCitation id="CE38AD1431A08AEB4219F827855B377C" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Larval morphology (foreleg) A Potamanthus formosus Eaton, 1892 B Rhoenanthus magnificus Ulmer, 1920 C R. obscurus Navas, 1922 D R. distafurcus Bae &amp; McCafferty, 1991 E R. speciosus Eaton, 1881. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1067.72779.figure11" httpUri="https://binary.pensoft.net/fig/604592" pageId="0" pageNumber="57">11E, and vi</figureCitation>
) 4 or 5 bipectinated lateral setae on the mandibles (
<bibRefCitation author="Bae, YJ" pageId="0" pageNumber="57" refId="B1" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117(3/4): 1-143." title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3 / 4): 1 - 143." year="1991">Bae and McCafferty 1991</bibRefCitation>
<bibRefCitation id="1841F0250B93A6813875198A048DEF91" author="Bae, YJ" pageId="0" pageNumber="57" refId="B1" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117(3/4): 1-143." title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3 / 4): 1 - 143." year="1991">Bae and McCafferty 1991</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="57">
<emphasis bold="true" pageId="0" pageNumber="57">Egg</emphasis>
<paragraph id="73CEE0D0F5CAB386EFF8F728663432BE" pageId="0" pageNumber="57">
<emphasis id="3D5D98B10970BF18EDACFEF4E010F4FE" bold="true" pageId="0" pageNumber="57">Egg</emphasis>
(dissected from mature larva). Oval; with two large conical polar caps, (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. SEMs of egg structures of Rhoenanthus speciosus Eaton, 1881 A general outline of egg B micropyle." figureDoi="10.3897/zookeys.1067.72779.figure13" httpUri="https://binary.pensoft.net/fig/604594" pageId="0" pageNumber="57">13A</figureCitation>
<figureCitation id="CC53BEE3DBACB6CCCA9022D5B349C828" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. SEMs of egg structures of Rhoenanthus speciosus Eaton, 1881 A general outline of egg B micropyle." figureDoi="10.3897/zookeys.1067.72779.figure13" httpUri="https://binary.pensoft.net/fig/604594" pageId="0" pageNumber="57">13A</figureCitation>
); chorion with numerous scattered tubercles, with knob-terminated coiled threads at the equatorial zone; tagenoform micropyle; sperm guide circular (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. SEMs of egg structures of Rhoenanthus speciosus Eaton, 1881 A general outline of egg B micropyle." figureDoi="10.3897/zookeys.1067.72779.figure13" httpUri="https://binary.pensoft.net/fig/604594" pageId="0" pageNumber="57">13B</figureCitation>
<figureCitation id="F24AFA8C5550994B86F426730BFC7788" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. SEMs of egg structures of Rhoenanthus speciosus Eaton, 1881 A general outline of egg B micropyle." figureDoi="10.3897/zookeys.1067.72779.figure13" httpUri="https://binary.pensoft.net/fig/604594" pageId="0" pageNumber="57">13B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="distribution">
<paragraph pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="57">Narathiwat and Songkla provinces.</paragraph>
<subSubSection id="F79FDB1F2E97F66E92564FE888AB260A" pageId="0" pageNumber="57" type="distribution">
<paragraph id="843212C257759D91AC716CE770C0B7D6" pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph id="08943B915C45AF29FD4CFE359C8ACA8F" pageId="0" pageNumber="57">Narathiwat and Songkla provinces.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="remarks">
<paragraph pageId="0" pageNumber="57">Remark.</paragraph>
<paragraph pageId="0" pageNumber="57">
<subSubSection id="C91F04A345E624D6E364036C3465017C" pageId="0" pageNumber="57" type="remarks">
<paragraph id="A1AC90148B81497900555181B7ECF55E" pageId="0" pageNumber="57">Remark.</paragraph>
<paragraph id="F5EDFF736B948913FD8A96733167A9B2" pageId="0" pageNumber="57">
The larvae of
<taxonomicName authorityName="Eaton" authorityYear="1881" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus speciosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis italics="true" pageId="0" pageNumber="57">Rhoenanthus speciosus</emphasis>
<taxonomicName id="18D22F1DB827E29D6251AC200D7960E5" authorityName="Eaton" authorityYear="1881" class="Insecta" family="Potamanthidae" genus="Rhoenanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhoenanthus speciosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis id="0794A85D68FF76665BE22D1D14DE039F" italics="true" pageId="0" pageNumber="57">Rhoenanthus speciosus</emphasis>
</taxonomicName>
were reported by
<bibRefCitation author="Bae, YJ" pageId="0" pageNumber="57" refId="B1" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117(3/4): 1-143." title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3 / 4): 1 - 143." year="1991">Bae and McCafferty (1991)</bibRefCitation>
<bibRefCitation id="C9C80474AFB2C10B634E69439412AB7A" author="Bae, YJ" pageId="0" pageNumber="57" refId="B1" refString="Bae, YJ, McCafferty, WP, 1991. Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117(3/4): 1-143." title="Phylogenetic Systematics of the Potamanthidae (Ephemeroptera). Transactions of the American Entomological Society 117 (3 / 4): 1 - 143." year="1991">Bae and McCafferty (1991)</bibRefCitation>
based on specimens from Indonesia and Malaysia.
<bibRefCitation author="Parnrong, S" journalOrPublisher="Science Asia" pageId="0" pageNumber="57" pagination="407 - 409" refId="B18" refString="Parnrong, S, Buathong, M, Sites, RW, 2002. New records of Behningiidae, Potamanthidae, and Prosopistomatidae (Ephemeroptera) from Thailand. Science Asia 28: 407 - 409" title="New records of Behningiidae, Potamanthidae, and Prosopistomatidae (Ephemeroptera) from Thailand." volume="28" year="2002">Parnrong et al. (2002)</bibRefCitation>
<bibRefCitation id="BFBA453B00337EBC69FB7983422BFBED" author="Parnrong, S" journalOrPublisher="Science Asia" pageId="0" pageNumber="57" pagination="407 - 409" refId="B18" refString="Parnrong, S, Buathong, M, Sites, RW, 2002. New records of Behningiidae, Potamanthidae, and Prosopistomatidae (Ephemeroptera) from Thailand. Science Asia 28: 407 - 409" title="New records of Behningiidae, Potamanthidae, and Prosopistomatidae (Ephemeroptera) from Thailand." volume="28" year="2002">Parnrong et al. (2002)</bibRefCitation>
reported this species from Songkhla province (southern Thailand). In the present study, we found the larva of this species in the nearby Narathiwat province. The distribution of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="R. speciosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis italics="true" pageId="0" pageNumber="57">R. speciosus</emphasis>
<taxonomicName id="B1CAF74F7EF5561E48A488F70EC14386" class="Insecta" kingdom="Animalia" lsidName="R. speciosus" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="speciosus">
<emphasis id="3B150CD31E68ED15AE6561505F94F48F" italics="true" pageId="0" pageNumber="57">R. speciosus</emphasis>
</taxonomicName>
seems to be restricted to the south of the Isthmus of Kra, as was found for another mayfly species,
<taxonomicName authorityName="Lieftinck" authorityYear="1932" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma wouterae" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="wouterae">
<emphasis italics="true" pageId="0" pageNumber="57">Prosopistoma wouterae</emphasis>
<taxonomicName id="6504288D0DFD1240261B5F5D84AB9823" authorityName="Lieftinck" authorityYear="1932" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma wouterae" order="Ephemeroptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="wouterae">
<emphasis id="0B2A35AA89334A1B25F7BD0D5B3346EB" italics="true" pageId="0" pageNumber="57">Prosopistoma wouterae</emphasis>
</taxonomicName>
(Boonsoong &amp; Sartori, 2019). These findings constitute the northern limit of the known distribution of this species.
</paragraph>

View file

@ -1,184 +1,184 @@
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<mods:namePart>Deranja, Maks</mods:namePart>
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<mods:affiliation>University of Zagreb, Faculty of Science, Department of Biology, Division of Zoology, Evolution Lab, Rooseveltov trg 6, HR- 10000 Zagreb, Croatia</mods:affiliation>
<mods:nameIdentifier type="email">maks.deranja@gmail.com</mods:nameIdentifier>
<mods:namePart id="5698871399CCCAD03C141582F8AF7A75">Deranja, Maks</mods:namePart>
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<treatment LSID="urn:lsid:plazi:treatment:BB5D5F6F6AD251C9898C7AA5B34501D8" httpUri="http://treatment.plazi.org/id/BB5D5F6F6AD251C9898C7AA5B34501D8" lastPageNumber="1" pageId="0" pageNumber="1">
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Subfamily
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<paragraph pageId="0" pageNumber="1">Composition and distribution.</paragraph>
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This diverse and polyphyletic family includes altogether 314 species assigned to 74 genera (
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). Most genera require revision and are not adequately classified in the
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system (
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). Within the subfamily
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, several monophyletic tribes exist, such as Caribbean
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(
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<emphasis italics="true" pageId="0" pageNumber="1">Choriphyllum</emphasis>
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<emphasis id="D7EBF6E6F722BF1B79DFBB7C1273798C" italics="true" pageId="0" pageNumber="1">Choriphyllum</emphasis>
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Serville, 1838 and
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<emphasis italics="true" pageId="0" pageNumber="1">Phyllotettix</emphasis>
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<emphasis id="F0C39F370BE2EEA2ED22091596BB8F22" italics="true" pageId="0" pageNumber="1">Phyllotettix</emphasis>
</taxonomicName>
Hancock, 1902), African
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(including leaf-like
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<emphasis italics="true" pageId="0" pageNumber="1">Acmophyllum</emphasis>
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<emphasis id="E857CFBBB35355473A46BFACBA165BA6" italics="true" pageId="0" pageNumber="1">Acmophyllum</emphasis>
</taxonomicName>
Karsch, 1890,
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<emphasis italics="true" pageId="0" pageNumber="1">Seyidotettix</emphasis>
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<emphasis id="405FAF19A1C3E90C01C1EBA65B8DB7EB" italics="true" pageId="0" pageNumber="1">Seyidotettix</emphasis>
</taxonomicName>
Rehn, 1938,
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<emphasis italics="true" pageId="0" pageNumber="1">Trypophyllum</emphasis>
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<emphasis id="992A0BD17EDB9AAF8B7A4AEDFB28FB2B" italics="true" pageId="0" pageNumber="1">Trypophyllum</emphasis>
</taxonomicName>
Karsch, 1890,
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<emphasis italics="true" pageId="0" pageNumber="1">Xerophyllum</emphasis>
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<emphasis id="2C91E08BCE3596803C90C4DF7E9C006E" italics="true" pageId="0" pageNumber="1">Xerophyllum</emphasis>
</taxonomicName>
Fairmaire, 1846), and Asian
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<taxonomicName id="F2B6A8864A6D121584529996CBCF6CBB" authorityName="Bolivar" authorityYear="1887" class="Insecta" family="Tetrigidae" genus="Cladonotini" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladonotini" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Cladonotini</taxonomicName>
(
<taxonomicName class="Insecta" family="Tetrigidae" genus="Hymenotes" kingdom="Animalia" lsidName="Hymenotes" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hymenotes</emphasis>
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<emphasis id="A1950BA25CC581EE1CEF4CCC4B3A62BD" italics="true" pageId="0" pageNumber="1">Hymenotes</emphasis>
</taxonomicName>
Westwood, 1837,
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<emphasis italics="true" pageId="0" pageNumber="1">Holoarcus</emphasis>
<taxonomicName id="E0E14A64F0C4A50932FB2B31019EFD6D" authorityName="Hancock" authorityYear="1909" class="Insecta" family="Tetrigidae" genus="Holoarcus" kingdom="Animalia" lsidName="Holoarcus" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="644CAF60E12D5B414CCF067EF56176B6" italics="true" pageId="0" pageNumber="1">Holoarcus</emphasis>
</taxonomicName>
Hancock, 1909,
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<emphasis italics="true" pageId="0" pageNumber="1">Dolatettix</emphasis>
<taxonomicName id="8E92825484D7459E37030A2614DD9CA2" class="Insecta" family="Tetrigidae" genus="Dolatettix" kingdom="Animalia" lsidName="Dolatettix" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="09EDF1EE7DBFA7DA52F53D5D18189825" italics="true" pageId="0" pageNumber="1">Dolatettix</emphasis>
</taxonomicName>
Hancock, 1907).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diversity in madagascar">
<paragraph pageId="0" pageNumber="1">Diversity in Madagascar.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="B04DBAB086333A70A0CEA57F239E4D1C" pageId="0" pageNumber="1" type="diversity in madagascar">
<paragraph id="E72CD05F652B7BB20368B93F09A3D886" pageId="0" pageNumber="1">Diversity in Madagascar.</paragraph>
<paragraph id="41FC5D7A5EDAF802E7F729E27324B91C" pageId="0" pageNumber="1">
Altogether, 18
<taxonomicName class="Insecta" family="Tetrigidae" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Tetrigidae</taxonomicName>
<taxonomicName id="A3D27C26F5689B1CD89814D30D8D813D" class="Insecta" family="Tetrigidae" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Tetrigidae</taxonomicName>
species of Madagascar are assigned to the subfamily
<taxonomicName authorityName="Bolivar" authorityYear="1887" family="Tetrigidae" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="1" rank="subFamily" subFamily="Cladonotinae">Cladonotinae</taxonomicName>
<taxonomicName id="C4E9EFE8EB16EB5E0505C1690A279A75" authorityName="Bolivar" authorityYear="1887" class="Insecta" family="Tetrigidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subFamily" subFamily="Cladonotinae">Cladonotinae</taxonomicName>
: two members of
<taxonomicName lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Valalyllini">Valalyllini</taxonomicName>
<taxonomicName id="8DB7DB5789AF60CD6B5B44F9B4D1449A" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Valalyllini">Valalyllini</taxonomicName>
trib. nov. (
<taxonomicName class="Insecta" genus="L." kingdom="Animalia" lsidName="L. fuscus" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis italics="true" pageId="0" pageNumber="1">L. fuscus</emphasis>
<taxonomicName id="87E157C71EDB0C730023182B6EE2ED2C" class="Insecta" kingdom="Animalia" lsidName="L. fuscus" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="2050195CFEE660BA25188C514BC92F19" italics="true" pageId="0" pageNumber="1">L. fuscus</emphasis>
</taxonomicName>
and
<taxonomicName genus="V." lsidName="V. folium" pageId="0" pageNumber="1" rank="species" species="folium">
<emphasis italics="true" pageId="0" pageNumber="1">V. folium</emphasis>
<taxonomicName id="F021E2E94AEE82D65DD03A88F429455C" lsidName="V. folium" pageId="0" pageNumber="1" rank="species" species="folium">
<emphasis id="0CC58BC5D1706902B92E0A86EECB853B" italics="true" pageId="0" pageNumber="1">V. folium</emphasis>
</taxonomicName>
); 13 species of the genus
<taxonomicName authorityName="Rehn" authorityYear="1929" class="Insecta" family="Tetrigidae" genus="Thymochares" kingdom="Animalia" lsidName="Thymochares" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Thymochares</emphasis>
<taxonomicName id="631405EA292D4AC221691F8A4DBD2F62" authorityName="Rehn" authorityYear="1929" class="Insecta" family="Tetrigidae" genus="Thymochares" kingdom="Animalia" lsidName="Thymochares" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="AD26E97C2506F87B2A774B6100C5BD8D" italics="true" pageId="0" pageNumber="1">Thymochares</emphasis>
</taxonomicName>
, which is currently not assigned to any of the tribes, and it is questionable whether it represents a genus of
<taxonomicName authorityName="Bolivar" authorityYear="1887" family="Tetrigidae" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="1" rank="subFamily" subFamily="Cladonotinae">Cladonotinae</taxonomicName>
<taxonomicName id="ACA1E0D2AD481D37360A0F8DE84A6C89" authorityName="Bolivar" authorityYear="1887" class="Insecta" family="Tetrigidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subFamily" subFamily="Cladonotinae">Cladonotinae</taxonomicName>
;
<taxonomicName authorityName="Devriese" authorityYear="1991" class="Insecta" family="Tetrigidae" genus="Microthymochares" kingdom="Animalia" lsidName="Microthymochares pullus" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pullus">
<emphasis italics="true" pageId="0" pageNumber="1">Microthymochares pullus</emphasis>
<taxonomicName id="87ADFB2D05A9949AFB7576D8873B8C37" authorityName="Devriese" authorityYear="1991" class="Insecta" family="Tetrigidae" genus="Microthymochares" kingdom="Animalia" lsidName="Microthymochares pullus" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pullus">
<emphasis id="18D92724A7341B8B5073DE02CCEFD1D6" italics="true" pageId="0" pageNumber="1">Microthymochares pullus</emphasis>
</taxonomicName>
, an endemic genus and species with the same taxonomic problem as
<taxonomicName authorityName="Rehn" authorityYear="1929" class="Insecta" family="Tetrigidae" genus="Thymochares" kingdom="Animalia" lsidName="Thymochares" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Thymochares</emphasis>
<taxonomicName id="FA47AE66AE31958522EDC0B976CE989B" authorityName="Rehn" authorityYear="1929" class="Insecta" family="Tetrigidae" genus="Thymochares" kingdom="Animalia" lsidName="Thymochares" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="CBC44584B06E84D69619E279C1393468" italics="true" pageId="0" pageNumber="1">Thymochares</emphasis>
</taxonomicName>
;
<taxonomicName class="Insecta" family="Tetrigidae" genus="Epitettix" kingdom="Animalia" lsidName="Epitettix spheniscus" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="spheniscus">
<emphasis italics="true" pageId="0" pageNumber="1">Epitettix spheniscus</emphasis>
<taxonomicName id="92E1CDE5E168F48403D1802BCEB37A6A" class="Insecta" family="Tetrigidae" genus="Epitettix" kingdom="Animalia" lsidName="Epitettix spheniscus" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="spheniscus">
<emphasis id="F8F06EDF31051F361B0954C44B3BFD32" italics="true" pageId="0" pageNumber="1">Epitettix spheniscus</emphasis>
</taxonomicName>
also with the same problematic assignment; and
<taxonomicName authorityName="Gunther" authorityYear="1979" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Xerophyllini">Xerophyllini</taxonomicName>
<taxonomicName id="871996382834FB182C2C7F554724096D" authorityName="Gunther" authorityYear="1979" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Xerophyllini">Xerophyllini</taxonomicName>
and its member
<taxonomicName class="Insecta" family="Tetrigidae" genus="Morphopoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morphopoides madagascariensis" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis italics="true" pageId="0" pageNumber="1">Morphopoides madagascariensis</emphasis>
<taxonomicName id="B37E458BB35231EE39F180836E66C3C5" class="Insecta" family="Tetrigidae" genus="Morphopoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Morphopoides madagascariensis" order="Orthoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="69A3F24B586C5E1D31352BE33093342C" italics="true" pageId="0" pageNumber="1">Morphopoides madagascariensis</emphasis>
</taxonomicName>
(
<bibRefCitation author="Cigliano, MM" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="1" publicationUrl="http://Orthoptera.SpeciesFile.org" refId="B3" refString="Cigliano, MM, Braun, H, Eades, DC, Otte, D, 2022. Orthoptera Species File. Version 5.0/5.0. http://Orthoptera.SpeciesFile.org" title="Orthoptera Species File. Version 5.0 / 5.0." url="http://Orthoptera.SpeciesFile.org" year="2022">Cigliano et al. 2022</bibRefCitation>
<bibRefCitation id="BC64F0A84149C305A09A826AD84AA50E" author="Cigliano, MM" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="1" publicationUrl="http://Orthoptera.SpeciesFile.org" refId="B3" refString="Cigliano, MM, Braun, H, Eades, DC, Otte, D, 2022. Orthoptera Species File. Version 5.0/5.0. http://Orthoptera.SpeciesFile.org" title="Orthoptera Species File. Version 5.0 / 5.0." url="http://Orthoptera.SpeciesFile.org" year="2022">Cigliano et al. 2022</bibRefCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,63 +1,63 @@
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<subSubSection id="3A1D61A32DA1659287827998605C2A5D" pageId="0" pageNumber="69" type="nomenclature">
<paragraph id="E3CE9712B34C5E5BFF839680A22E863E" pageId="0" pageNumber="69">
<taxonomicName id="CD697462CDFC5216BF1DE0ED3A9AA24C" ID-CoL="7RYT7" LSID="BB5FF791-5B1A-58AE-8389-606581E755AD" authority="latecincta rearing experiments" authorityName="rearing experiments" family="Chrysomelidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ch. latecincta subsp. latecincta" order="Coleoptera" pageId="0" pageNumber="69" rank="subSpecies" species="latecincta" subSpecies="latecincta">Ch. latecincta latecincta rearing experiments</taxonomicName>
<taxonomicName id="CD697462CDFC5216BF1DE0ED3A9AA24C" ID-CoL="7RYT7" LSID="BB5FF791-5B1A-58AE-8389-606581E755AD" authority="latecincta rearing experiments" authorityName="rearing experiments" class="Insecta" family="Chrysomelidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ch. latecincta subsp. latecincta" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="latecincta" subSpecies="latecincta">Ch. latecincta latecincta rearing experiments</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="709704AE4721D38E0455B2268C75B7D4" pageId="0" pageNumber="69" type="notes">
@ -74,7 +74,7 @@
<emphasis id="A2E32D9B5B3FA84ED2C5BDE52920D019" italics="true" pageId="0" pageNumber="69">Linaria vulgaris</emphasis>
</taxonomicName>
was used temporarily), as with
<taxonomicName id="EA93E0DACB84CBD2E400BF34370174F8" family="Chrysomelidae" kingdom="Animalia" lsidName="Ch. kuesteri" order="Coleoptera" pageId="0" pageNumber="69" rank="species" species="kuesteri">
<taxonomicName id="EA93E0DACB84CBD2E400BF34370174F8" class="Insecta" family="Chrysomelidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ch. kuesteri" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="species" species="kuesteri">
<emphasis id="AC271BCDAD300FF60B33227CD68D2762" italics="true" pageId="0" pageNumber="69">Ch. kuesteri</emphasis>
</taxonomicName>
, which was well accepted. In the middle of September 2021 already three eggs were laid, one of the larvae survived and moulted a fist time till 23rd September, and a second time on 3rd October. From middle of December to February 2022 the larva did not feed anymore and hardly moved. Beginning of March the larva died of unknown reason and despite (or because?) of the mild winter. Three females laid more than 20 eggs (Fig.
@ -124,7 +124,7 @@ Pupa (Photos: C. Germann).
et al. (2013)
</bibRefCitation>
reported, that
<taxonomicName id="61C92DB8195E99B6201E2C41F8EF72AD" authority="latecincta" family="Chrysomelidae" kingdom="Animalia" lsidName="Ch. latecincta subsp. latecincta" order="Coleoptera" pageId="0" pageNumber="69" rank="subSpecies" species="latecincta" subSpecies="latecincta">
<taxonomicName id="61C92DB8195E99B6201E2C41F8EF72AD" authority="latecincta" class="Insecta" family="Chrysomelidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ch. latecincta subsp. latecincta" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="latecincta" subSpecies="latecincta">
<emphasis id="BFBDEC0DEECE5579325AFA1E84519433" italics="true" pageId="0" pageNumber="69">Ch. latecincta latecincta</emphasis>
</taxonomicName>
develops on an
@ -140,7 +140,7 @@ and
<emphasis id="EE18BA0E14F82AEF166106954EC1C11C" italics="true" pageId="0" pageNumber="69">Antirrhinum latifolium</emphasis>
</taxonomicName>
Mill. in the French Alps (800-1500 m) and they develop from September to April, with a quiescence inbetween. The same authors reported a larval development from May to June for
<taxonomicName id="5CA9543908E4B8A6DB0A86F1634B2667" family="Chrysomelidae" kingdom="Animalia" lsidName="Ch. latecincta subsp. decipiens" order="Coleoptera" pageId="0" pageNumber="69" rank="subSpecies" species="latecincta" subSpecies="decipiens">
<taxonomicName id="5CA9543908E4B8A6DB0A86F1634B2667" class="Insecta" family="Chrysomelidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ch. latecincta subsp. decipiens" order="Coleoptera" pageId="0" pageNumber="69" phylum="Arthropoda" rank="subSpecies" species="latecincta" subSpecies="decipiens">
<emphasis id="32033FA7E0020A08229E84615DCCF543" italics="true" pageId="0" pageNumber="69">Ch. latecincta decipiens</emphasis>
</taxonomicName>
(Franz, 1938) from the Pyrenees.

View file

@ -1,117 +1,117 @@
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<paragraph id="B0BEF931BB6AABB8C7A540691D3A45EF" pageId="8" pageNumber="9">
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rufithorax Enderlein, 1905
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Figures 13-18
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<subSubSection pageId="8" pageNumber="9" type="reference_group">
<paragraph pageId="8" pageNumber="9">
<taxonomicName class="Insecta" family="Evaniidae" genus="Evaniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Evaniscus rufithorax" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rufithorax">Evaniscus rufithorax</taxonomicName>
<subSubSection id="0E5E64E2CDBB57E27C4538B9D5936157" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="E5474D95C536035129F93B90B7DA7A80" pageId="8" pageNumber="9">
<taxonomicName id="F27B986BEAA1B5D6B315DE6AAE62FA6E" class="Insecta" family="Evaniidae" genus="Evaniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Evaniscus rufithorax" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rufithorax">Evaniscus rufithorax</taxonomicName>
: Enderlein, 1905 (original description) male, deposited at ZMPA, labels: &quot;Bolivien. Mapiri and Peru: Pachita{Pachitea}-
<normalizedToken originalValue="Fluß”">Fluss&quot;</normalizedToken>
<normalizedToken id="B4C19041D6BE4BC59758D2401854EBEC" originalValue="Fluß”">Fluss&quot;</normalizedToken>
, mx_id: 480; syntype male, deposited at ZMHB, labels: &quot;Bolivia Mapiri Staudinger, V.&quot;, mx_id: 15349; syntype male, deposited at ZMHB, labels: &quot;Bolivia Mapiri Staudinger, V.&quot;, mx_id: 15350; syntype male, deposited at ZMHB, labels: &quot;Peru Pachitea-Fluss Staudinger, V.&quot;, mx_id: 15351
</paragraph>
<paragraph pageId="8" pageNumber="9">
<taxonomicName class="Insecta" family="Evaniidae" genus="Evaniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Evaniscus rufithorax" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rufithorax">Evaniscus rufithorax</taxonomicName>
<paragraph id="36CF32792F70BDF923F3B8759BF81C5D" pageId="8" pageNumber="9">
<taxonomicName id="C19D49EBF8C9865540CFF0C46374682A" class="Insecta" family="Evaniidae" genus="Evaniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Evaniscus rufithorax" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rufithorax">Evaniscus rufithorax</taxonomicName>
: Enderlein, G. 1905: 711
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<paragraph pageId="8" pageNumber="9">Diagnosis.</paragraph>
<paragraph pageId="8" pageNumber="9">
<taxonomicName class="Insecta" family="Evaniidae" genus="Evaniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Evaniscus rufithorax" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rufithorax">Evaniscus rufithorax</taxonomicName>
<subSubSection id="EEA07C88982C90ED0086E64850DC13B7" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="1ECEFC557A4835DC6CC3FC514B3ABE1F" pageId="8" pageNumber="9">Diagnosis.</paragraph>
<paragraph id="911C57420110A24C2CA3D8AF6425A1C0" pageId="8" pageNumber="9">
<taxonomicName id="4041CEF63DF7D4D13ECFA05F99F89A17" class="Insecta" family="Evaniidae" genus="Evaniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Evaniscus rufithorax" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="rufithorax">Evaniscus rufithorax</taxonomicName>
is the most commonly collected species of in the genus. This species differs from other
<taxonomicName genus="Evansicus" lsidName="Evansicus" pageId="8" pageNumber="9" rank="genus">Evansicus</taxonomicName>
<taxonomicName id="EA656DA2016B9408A802FFD1E1C89A6F" genus="Evansicus" lsidName="Evansicus" pageId="8" pageNumber="9" rank="genus">Evansicus</taxonomicName>
by a combination of the following character states: pronotal lobe carina presence: present; subantennal carina length: extending ventrally of medial margin of lower face.
</paragraph>
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<subSubSection lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph pageId="8" pageNumber="9">Description.</paragraph>
<paragraph pageId="8" pageNumber="9">Head. Head color: dorsal half of upper face and vertex color black; ventral half of upper face and lower face color red or yellow; orange. Mandible color vs clypeus color: mandible color same as clypeus color. Subantennal carina length: extending ventrally of medial margin of lower face. Preorbital carina length: extending dorsally to ventral margin of the antennal foramen. Upper face sculpture: foveate. Malar space length vs. half compound eye height (male): shorter than half compound eye height. Ocellar ocular line length vs. lateral ocellus diameter: shorter than lateral ocellus diameter. Posterior ocellar length vs. lateral ocellus diameter: 1.5x as long as the diameter of the lateral ocellus. Ventral region of occipital carina curvature in lateral view: straight. Ventral region of the postoccipital carina shape: not raised. Radicle color: yellow; orange. Scape color: yellow; orange. Scape length vs compound eye height: scape shorter than half compound eye height. Female flagellomere 1-8 shape: distinctly longer than wide (length of flagellomere&gt; width of flagellomere). Shape of occiput: as high as wide.</paragraph>
<paragraph lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="A7C344F4F47D9C9A1B0C0BBDCE06726A" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="B54A8417B1BF8383DF46479066D8FD55" pageId="8" pageNumber="9">Description.</paragraph>
<paragraph id="582251C8125809BA7A765E6F0922195A" pageId="8" pageNumber="9">Head. Head color: dorsal half of upper face and vertex color black; ventral half of upper face and lower face color red or yellow; orange. Mandible color vs clypeus color: mandible color same as clypeus color. Subantennal carina length: extending ventrally of medial margin of lower face. Preorbital carina length: extending dorsally to ventral margin of the antennal foramen. Upper face sculpture: foveate. Malar space length vs. half compound eye height (male): shorter than half compound eye height. Ocellar ocular line length vs. lateral ocellus diameter: shorter than lateral ocellus diameter. Posterior ocellar length vs. lateral ocellus diameter: 1.5x as long as the diameter of the lateral ocellus. Ventral region of occipital carina curvature in lateral view: straight. Ventral region of the postoccipital carina shape: not raised. Radicle color: yellow; orange. Scape color: yellow; orange. Scape length vs compound eye height: scape shorter than half compound eye height. Female flagellomere 1-8 shape: distinctly longer than wide (length of flagellomere&gt; width of flagellomere). Shape of occiput: as high as wide.</paragraph>
<paragraph id="FF319781244ADE25E32C5BC4614D037C" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Mesosoma. Mesosoma length: 2.25-2.85 mm (n=29). Antero-dorsal region of mesosoma color: red. Postero-ventral region of mesosoma color: black; red. Median notch of transverse pronotal carina presence: present. Transverse pronotal carina length: long, extending postero-laterally of epomia. Pronotal collar sculpture: foveate. Patch that is part of dorsal region of lateral pronotal area texture: smooth. Pronotal lobe carina presence: present. Mesonotum color: red; black. Mesoscutum shape: as long as wide (length of mesoscutum&gt; width of mesoscutum). Antero-admedian line length vs. lateral ocellus diameter: equal to lateral ocellus diameter. Parapsidal signum conspicuousness: inconspicuous. Foveae on notaulus presence: present. Distance between depressions vs. diameter of depressions on internotaular area: greater than the diameter of one depression. Mesofemoral depression sculpture: smooth. Mesofemoral depression pilosity presence: absent. Ventral area of the mesopectus sculpture: foveate. Medial region of transmetapectal carina presence: absent. Area dorsal of transmetapectal carina sculpture: smooth. Posterior propodeal projection shape in lateral view: not raised.
<pageBreakToken pageId="9" pageNumber="10" start="start">Posterior</pageBreakToken>
<pageBreakToken id="07932AB67EEB0AE74094706ED6B4F56F" pageId="9" pageNumber="10" start="start">Posterior</pageBreakToken>
region of plica presence: absent. Dorsal area of the metapectal-propodeal complex sculpture: foveate. Posterior margin of the metapectal-propodeal complex ventrally of the propodeal foramen curvature in lateral view: curved. Mesosoma color: black posteroventrally, red anterodorsally.
</paragraph>
<paragraph pageId="9" pageNumber="10">
<paragraph id="C8FE0128EA938AFA7CB49174FCD3223E" pageId="9" pageNumber="10">
Legs. Fore leg color: yellow; red. Mid leg color: yellow; red. Hind leg color: red-black. Metafemur length vs. metatibia length: metafemur equal to or shorter than metatibia. Metabasitarsus length vs metatibia length: metabasitarsus 1.2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="B464EE9F18B23E5DE2F3126E20A78088" originalValue="×">x</normalizedToken>
to 1.4
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="12DEEE25D1E2B722FFADEB7ED4F0AD4F" originalValue="×">x</normalizedToken>
as short as metatibia.
</paragraph>
<paragraph pageId="9" pageNumber="10">Wings. Fore wing vein color: black. Setae on proximal region of fore wing color: black.</paragraph>
<paragraph pageId="9" pageNumber="10">Metasoma. Metasoma color: black. Dorsal region of petiole sculpture: foveate.</paragraph>
<paragraph id="CF245D489081C5272E277D5EDF1C6ECB" pageId="9" pageNumber="10">Wings. Fore wing vein color: black. Setae on proximal region of fore wing color: black.</paragraph>
<paragraph id="64FF1FB527EFE9489E48F5103C90831E" pageId="9" pageNumber="10">Metasoma. Metasoma color: black. Dorsal region of petiole sculpture: foveate.</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="material examined">
<paragraph pageId="9" pageNumber="10">Material examined.</paragraph>
<paragraph lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="D71EAAC6D55D6CE8F0D0354907FA118A" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="material examined">
<paragraph id="11D60530B110242DB1EAFDABC1EF70A6" pageId="9" pageNumber="10">Material examined.</paragraph>
<paragraph id="807628B529C3B41E4FBBB25C8177E519" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Lectotype male, current designation: PERU: 1 male. mx_id 15351 (ZMHB). Paralectotypes: 3 males, current designation: BOLIVIA: 3
<pageBreakToken pageId="10" pageNumber="11" start="start">males</pageBreakToken>
<pageBreakToken id="5263A9F7B88D5B36445B6537C06D383C" pageId="10" pageNumber="11" start="start">males</pageBreakToken>
. mx_id 15349-15350 (ZMHB); mx_id 23037 (ZMPA). Other material (21 females, 77 males, 1 sex unknown): BRAZIL: 8 females, 31 males, 1 sex unknown. NCSU 2395-2400, 6967-6968, 41750 (AEIC); NCSU 67257-67273, 67279 (INPA); NCSU 9888-9889 (UCDC); NCSU 67280-67286 (MPEG); NCSU 67287-67290 (MZSP). COLOMBIA: 1 female, 4 males. NCSU 33582, 36699, 41741-41742 (NCSU); mx 2840 (unknown). ECUADOR: 9 females, 23 males. NCSU 6969-6974, 6976-6977, 6986-7000 (AEIC); NCSU 6979-6985 (UCDC); NCSU 6975 (INHS); NCSU 6978 (USNM). GUYANA: 1 male. mx_id 15338 (BMNH). PERU: 3 females, 15 males. NCSU 18398-18399 (MZLU); NCSU 67277-67278 (MIUP); NCSU 9886-9887, 9890-9891 (AEIC); mx_id 15340-15342 (BMNH); NCSU 2391-2394 (INHS); NCSU 67274-67276 (CAS). SURINAME: 1 male. mx_id 15337 (BMNH).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="11" type="comments">
<paragraph pageId="10" pageNumber="11">Comments.</paragraph>
<paragraph pageId="10" pageNumber="11">
<subSubSection id="8B31264C9AFF4D56F549D39A40937890" pageId="10" pageNumber="11" type="comments">
<paragraph id="82AC24B9886B1FFB8D48A5BB9E5E0102" pageId="10" pageNumber="11">Comments.</paragraph>
<paragraph id="B9D356F0977103D755BB7C9D666B240D" pageId="10" pageNumber="11">
A lectotype was designated because syntypes were from specimens with different localities and one syntype varies in color; the male specimen chosen as lectotype from Peru is in very good condition and fits the description well. The male paralectotype from Bolivia that is at ZMPA varies in color (only) from the other type specimens and all known other material; the antero-dorsal region of mesosoma color is red in all
<taxonomicName class="Insecta" family="Evaniidae" genus="Evaniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Evaniscus rufithorax" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="rufithorax">Evaniscus rufithorax</taxonomicName>
<taxonomicName id="29CB676FC8A9C8B6546F9B91C2B917A8" class="Insecta" family="Evaniidae" genus="Evaniscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Evaniscus rufithorax" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="rufithorax">Evaniscus rufithorax</taxonomicName>
specimens, but is black in this type specimen.
</paragraph>
</subSubSection>

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@ -1,122 +1,122 @@
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<paragraph pageId="3" pageNumber="4">
<taxonomicName LSID="http://species-id.net/wiki/Parastratiosphecomyia_stratiosphecomyioides" authority="Brunetti, 1923" authorityName="Brunetti" authorityYear="1923" class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides Brunetti, 1923</taxonomicName>
<treatment id="BB69E706F5362DC3F468E1D565EB200C" ID-GBIF-Taxon="152037704" LSID="urn:lsid:plazi:treatment:BB69E706F5362DC3F468E1D565EB200C" httpUri="http://treatment.plazi.org/id/BB69E706F5362DC3F468E1D565EB200C" lastPageId="7" lastPageNumber="7" pageId="3" pageNumber="4" scope_family="Stratiomyidae" scope_order="Diptera" scope_subFamily="Pachygastrinae">
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<paragraph id="4E67EA059D1CE526F9358F9BE81346A5" pageId="3" pageNumber="4">
<taxonomicName id="FA8BB5381F1B45FF3300013A35A7C0C9" ID-CoL="4DDPZ" LSID="http://species-id.net/wiki/Parastratiosphecomyia_stratiosphecomyioides" authority="Brunetti, 1923" authorityName="Brunetti" authorityYear="1923" class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides Brunetti, 1923</taxonomicName>
Figs 2, 6, 9, 11-15
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="reference_group">
<paragraph pageId="3" pageNumber="4">
<taxonomicName class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides</taxonomicName>
<subSubSection id="C8F5FA2844FCD6067DDCE355B03CFB5A" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="1019B1E1E9C8A5D00178D31AEAFBC73A" pageId="3" pageNumber="4">
<taxonomicName id="120CC448B7F8CAB9B127AC250B56C352" class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides</taxonomicName>
Brunetti, 1923: 67.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="diagnosis">
<paragraph pageId="3" pageNumber="4">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="4">
<taxonomicName class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides</taxonomicName>
<subSubSection id="8F129C73CBF0B550CA10D543C3DB3179" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="5440589C26CB82847B4A7A8135829E5E" pageId="3" pageNumber="4">Diagnosis.</paragraph>
<paragraph id="EE13DE84B2F4E61137C88176EF4A8082" pageId="3" pageNumber="4">
<taxonomicName id="6AFC02DBE3F6C53BA2BF5F203B77BDFF" class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides</taxonomicName>
is easily distinguished from the other three known species in the genus by its remarkably produced ventral side of the antennal scape (Fig. 9).
</paragraph>
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<subSubSection lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="redescription">
<paragraph pageId="3" pageNumber="4">Redescription.</paragraph>
<paragraph pageId="3" pageNumber="4">Male.Head: Brownish black, but lower frons and most of face pale yellow, lower frons with small diffuse brownish spots adjacent to inner margins of antennal sockets, face dark near oral margin, including conical processes; face slightly convex but depressed medially on lower part, not concave on lowest part between conical processes, with upper medial portion moderately striate, and with a tiny conical process at lower, outer margin of each antennal socket; lower frons with very narrow band of pale tomentum at eye margins, similar tomentum along eye margin at gena and postgena; face, gena, and ocellar tubercle with short to moderate length silvery white hair-like setae, longest on genal area; antenna with scape and pedicel dark yellow, scape slightly darker dorsally, both with short, dark, semi-appressed hair-like setae, densest dorsally; scape with ventral surface markedly produced, inner region near base with rounded excavation; flagellum blackish with dense pilosity (most of flagellum missing in males examined, presumably as in female described below); palpus with first segment pale yellow, second dark brownish; proboscis dark yellowish.</paragraph>
<paragraph lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection id="12D6B0D1267E3EE59B7C5C4E8F977626" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="redescription">
<paragraph id="385F3EA695901DF42DB215AA342DABED" pageId="3" pageNumber="4">Redescription.</paragraph>
<paragraph id="89985C5FA9F4F30D6B3544507918AE36" pageId="3" pageNumber="4">Male.Head: Brownish black, but lower frons and most of face pale yellow, lower frons with small diffuse brownish spots adjacent to inner margins of antennal sockets, face dark near oral margin, including conical processes; face slightly convex but depressed medially on lower part, not concave on lowest part between conical processes, with upper medial portion moderately striate, and with a tiny conical process at lower, outer margin of each antennal socket; lower frons with very narrow band of pale tomentum at eye margins, similar tomentum along eye margin at gena and postgena; face, gena, and ocellar tubercle with short to moderate length silvery white hair-like setae, longest on genal area; antenna with scape and pedicel dark yellow, scape slightly darker dorsally, both with short, dark, semi-appressed hair-like setae, densest dorsally; scape with ventral surface markedly produced, inner region near base with rounded excavation; flagellum blackish with dense pilosity (most of flagellum missing in males examined, presumably as in female described below); palpus with first segment pale yellow, second dark brownish; proboscis dark yellowish.</paragraph>
<paragraph id="84234982D8D252A374301BF50BAB5AE9" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Thorax: Prothorax yellow but proepisternum is brownish black; scutum black with a pair of triangle-shaped lateral spots that are mostly on the presutural part, and area around postalar callus similarly yellow; scutellum black with apical half or more yellow; pleura yellow but ventral part of anepisternum, most of katepisternum, ventral part of meron, laterotergite and mediotergite brownish black; scutum with short, semi-appressed pilosity consisting of dark hair-like setae on dark cuticular areas, and pale hair-like setae on yellow areas, except laterally slightly longer hair-like setae present
<pageBreakToken pageId="4" pageNumber="5" start="start">that</pageBreakToken>
<pageBreakToken id="F28A3EF35687F45832CBCBAC437899DD" pageId="4" pageNumber="5" start="start">that</pageBreakToken>
are mostly pale; pilosity of pleura pale, slightly longer than on dorsal part of thorax and more erect, dorsal part of anepisternum bare; legs with coxae and trochanters pale yellow; front femur dark brown at base, gradually becoming dark yellow in basal third, middle and hind femora similar but the basal brown region more extensive, becoming yellow in apical half; front tibia mostly dark yellowish, narrowly dark dorsally; middle and hind tibiae brownish, more yellowish on ventral parts; tarsi yellowish, middle tarsus paler than others; halter with stem yellowish white, knob dark brown; wing (Fig. 6) with subapical cloud of infuscation starting at the proximal edge of discal cell, darkest and most evident in basal two-thirds of cell r5.
</paragraph>
<paragraph pageId="4" pageNumber="5">Abdomen: Blackish brown, first tergite with irregular yellowish medial spot, tergites 3-5 with narrow lateral margins vaguely paler; first tergite with moderately long, pale hair-like setae, tergites 2-5 densely set with short, dark hair-like setae on most of dorsal surface, tergites 2-3 with longer pale hair-like setae laterally (similar to those on first segment) and tergites 4-5 with longer dark hairs along lateral and posterolateral margins; sternites 1-3 yellowish, 4-5 brownish, with short, pale hair-like setae basally which become dark from apical half of sternite 3 posteriorly.</paragraph>
<paragraph pageId="4" pageNumber="5">Terminalia: Gonocoxites (Fig. 11) with lateral margins nearly straight, diverging posteriorly, with a pair of rounded processes posterior to gonocoxal apodemes; gonocoxal apodemes extending anteriorly to about anterior margin of genital capsule; synsternite of genital capsule with posterior process that is slightly bilobed; gonostyli arcuate, without processes (Figs 11, 12); phallic complex (Figs 13, 14) small, trilobed, the medial lobe shorter than lateral lobes; epandrium (Fig. 15) large, evenly convex, slightly indented posterolaterally, posterior margin truncate; hypoproct sclerotized, but not expanded dorsolaterally.</paragraph>
<paragraph pageId="4" pageNumber="5">Length: 10.8 mm.</paragraph>
<paragraph pageId="4" pageNumber="5">Female. Differs from male as follows: Head: Frons 0.27-0.28 head width, upper and lower frons gradually widening ventrally, upper frons with slight medial depression in front of anterior ocellus, junction of upper and lower frons with indistinct elevation on each side of median line; upper frons with pale, appressed hair-like setae except on ocellar tubercle and in medial depression; antenna 3.6 times length of head; first five flagellomeres of antennal flagellum with dense, black velvety vestiture, with scattered fine, erect hair-like setae especially posteriorly, flagellomeres 6-8 with more erect, longer pilosity that is gradually longer toward apex, last flagellomere tapered apically; palpus with second segment more robust than in male.</paragraph>
<paragraph pageId="4" pageNumber="5">Thorax: Scutum with hair-like setae mostly appressed, golden yellow; front femur sometimes more extensively dark on basal two-thirds; front tibia more extensively brownish black on dorsal part.</paragraph>
<paragraph lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<paragraph id="978739D572A6B0B68F4BEE397AD0AC9D" pageId="4" pageNumber="5">Abdomen: Blackish brown, first tergite with irregular yellowish medial spot, tergites 3-5 with narrow lateral margins vaguely paler; first tergite with moderately long, pale hair-like setae, tergites 2-5 densely set with short, dark hair-like setae on most of dorsal surface, tergites 2-3 with longer pale hair-like setae laterally (similar to those on first segment) and tergites 4-5 with longer dark hairs along lateral and posterolateral margins; sternites 1-3 yellowish, 4-5 brownish, with short, pale hair-like setae basally which become dark from apical half of sternite 3 posteriorly.</paragraph>
<paragraph id="6C36948F42389A382198F6F5C7FB2FF8" pageId="4" pageNumber="5">Terminalia: Gonocoxites (Fig. 11) with lateral margins nearly straight, diverging posteriorly, with a pair of rounded processes posterior to gonocoxal apodemes; gonocoxal apodemes extending anteriorly to about anterior margin of genital capsule; synsternite of genital capsule with posterior process that is slightly bilobed; gonostyli arcuate, without processes (Figs 11, 12); phallic complex (Figs 13, 14) small, trilobed, the medial lobe shorter than lateral lobes; epandrium (Fig. 15) large, evenly convex, slightly indented posterolaterally, posterior margin truncate; hypoproct sclerotized, but not expanded dorsolaterally.</paragraph>
<paragraph id="163C1ADB8D29AEABA549E84FAC1CEC71" pageId="4" pageNumber="5">Length: 10.8 mm.</paragraph>
<paragraph id="7AEA85EB33E6DDDFB518E0D88CCFD274" pageId="4" pageNumber="5">Female. Differs from male as follows: Head: Frons 0.27-0.28 head width, upper and lower frons gradually widening ventrally, upper frons with slight medial depression in front of anterior ocellus, junction of upper and lower frons with indistinct elevation on each side of median line; upper frons with pale, appressed hair-like setae except on ocellar tubercle and in medial depression; antenna 3.6 times length of head; first five flagellomeres of antennal flagellum with dense, black velvety vestiture, with scattered fine, erect hair-like setae especially posteriorly, flagellomeres 6-8 with more erect, longer pilosity that is gradually longer toward apex, last flagellomere tapered apically; palpus with second segment more robust than in male.</paragraph>
<paragraph id="94498DA4823A62AE92CD4BE90D79BF6B" pageId="4" pageNumber="5">Thorax: Scutum with hair-like setae mostly appressed, golden yellow; front femur sometimes more extensively dark on basal two-thirds; front tibia more extensively brownish black on dorsal part.</paragraph>
<paragraph id="ED24FBEE2454AB58009DC2D99D4FE8A2" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Abdomen: Tergite 2 and basal part of tergite 3 suffused with yellowish color medially; tergites 4-5 with shorter golden yellow hairs laterally and posterolaterally; sternite 8 with lateral margins evenly rounded, evenly continuous with posterior margin, produced dorsally along sides of terminalia; cercus yellowish brown, with first segment cylindrical, about three times as long as short, ovoid second segment
<pageBreakToken pageId="5" pageNumber="6" start="start">.</pageBreakToken>
<pageBreakToken id="68FC6F0FB3054E3070FFEBCCCF090902" pageId="5" pageNumber="6" start="start">.</pageBreakToken>
</paragraph>
<paragraph pageId="5" pageNumber="6">Length: 10.3-10.4 mm.</paragraph>
<paragraph id="3CFBB82A0C35C7A011F1E30F4F54AE99" pageId="5" pageNumber="6">Length: 10.3-10.4 mm.</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="distribution">
<paragraph pageId="5" pageNumber="6">Distribution.</paragraph>
<paragraph pageId="5" pageNumber="6">Known from peninsular Malaysia and adjacent Thailand.</paragraph>
<subSubSection id="3ECF02C0EC88FE8178B7FC0C8EEC44FE" pageId="5" pageNumber="6" type="distribution">
<paragraph id="EE8732537F38CBC2F305E6D77578B987" pageId="5" pageNumber="6">Distribution.</paragraph>
<paragraph id="E055231AD922026DC2A88743B20BD64F" pageId="5" pageNumber="6">Known from peninsular Malaysia and adjacent Thailand.</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="type material examined">
<paragraph pageId="5" pageNumber="6">Type material examined.</paragraph>
<paragraph pageId="5" pageNumber="6">
<subSubSection id="6B4FBABE940DB4F1C8980A957E66B114" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="type material examined">
<paragraph id="C220F63DFF1AB4D3648808CBE9E4CAE7" pageId="5" pageNumber="6">Type material examined.</paragraph>
<paragraph id="DAAF61D4997589A08DE3BE6C0120BDB3" pageId="5" pageNumber="6">
The four syntypes noted in
<bibRefCitation author="Brunetti, E" journalOrPublisher="Records of the Indian Museum" pageId="15" pageNumber="16" pagination="45 - 180" title="Second revision of the Oriental Stratiomyidae." volume="25" year="1923">Brunetti (1923)</bibRefCitation>
<bibRefCitation id="C9B2A7FECC2B5C416B3DABFB67509495" author="Brunetti, E" journalOrPublisher="Records of the Indian Museum" pageId="15" pageNumber="16" pagination="45 - 180" title="Second revision of the Oriental Stratiomyidae." volume="25" year="1923">Brunetti (1923)</bibRefCitation>
are present in BMNH, three of which were subsequently labeled as syntypes. I am hereby designating the male specimen in the most complete condition as lectotype to stabilize the taxonomic concept of this species and its name. The specimen is labeled: &quot;Siam: Bulsit Besar. H.C.Robinson &amp; N.Annandale. 1916-21./LECTOTYPE ♂
<taxonomicName class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides</taxonomicName>
<taxonomicName id="774324911846939CED4127F5364C2C06" class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides</taxonomicName>
Brunetti, 1923 des. N. E. Woodley 2012&quot;. As can be seen from the label data, this specimen had not previously been labeled as a syntype, but it clearly is one. There is no date on the data label. The specimen (Fig. 2) is in moderately good condition, but is missing the left antennal flagellum, the apical seven flagellomeres of the right antenna, and the right middle leg beyond the trochanter.
</paragraph>
<paragraph pageId="6" pageNumber="7">
<pageBreakToken pageId="6" pageNumber="7" start="start">Lectotype</pageBreakToken>
<paragraph id="D15EA8E12E0B1F8B3204B0603A5236DF" pageId="6" pageNumber="7">
<pageBreakToken id="E154DE00A27AAF29B5DC08FCDE7AC52D" pageId="6" pageNumber="7" start="start">Lectotype</pageBreakToken>
male (BMNH), THAILAND: Pattani Province, Bukit Besar, H.C. Robinson and N. Annandale; paralectotype male (BMNH), same data but 2500 feet, 1.ix.1901; paralectotype male (BMNH), same data but 2500 feet, 30.viii.1901; paralectotype female (BMNH), same data but 2500 feet, 4.ix.1901.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="additional material examined">
<paragraph pageId="6" pageNumber="7">Additional material examined.</paragraph>
<paragraph pageId="6" pageNumber="7">
<subSubSection id="693AF21137F94F6B969D72F55EDE69DA" pageId="6" pageNumber="7" type="additional material examined">
<paragraph id="EE3F7DADEF372F196E548B3B15735C29" pageId="6" pageNumber="7">Additional material examined.</paragraph>
<paragraph id="5504CE5E554385138A729C5FF6A13208" pageId="6" pageNumber="7">
MALAYSIA: 1 female (FSMU), Perak, Cameron Highlands, environs of Batu Village,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="4.366667">4°22'N</geoCoordinate>
<geoCoordinate id="C4D2CA41576FB37E0F0B7C764295D7E7" direction="north" orientation="latitude" precision="925" value="4.366667">4°22'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="101.333336">101°20'E</geoCoordinate>
<geoCoordinate id="1FD34F15DF25219BA055DE43A54767C9" direction="east" orientation="longitude" precision="925" value="101.333336">101°20'E</geoCoordinate>
, 590 meters, v.2009,
<normalizedToken originalValue="Pacholátko">Pacholatko</normalizedToken>
<normalizedToken id="6A4BCC270BCAAD686219768472DC6954" originalValue="Pacholátko">Pacholatko</normalizedToken>
leg.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="remarks">
<paragraph pageId="6" pageNumber="7">Remarks.</paragraph>
<paragraph lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<bibRefCitation author="Brunetti, E" journalOrPublisher="Records of the Indian Museum" pageId="15" pageNumber="16" pagination="45 - 180" title="Second revision of the Oriental Stratiomyidae." volume="25" year="1923">Brunetti (1923)</bibRefCitation>
<subSubSection id="4710A83288D415CC80C6F4262FD16D2E" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="remarks">
<paragraph id="96F6BE7729F663D599AC2B3CD5B1C412" pageId="6" pageNumber="7">Remarks.</paragraph>
<paragraph id="582C5C9139BB5B9239DB58EE58F2D42F" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<bibRefCitation id="079E5597727A39FA6D1B5398C2304A7D" author="Brunetti, E" journalOrPublisher="Records of the Indian Museum" pageId="15" pageNumber="16" pagination="45 - 180" title="Second revision of the Oriental Stratiomyidae." volume="25" year="1923">Brunetti (1923)</bibRefCitation>
cited the type locality for
<taxonomicName class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides</taxonomicName>
<taxonomicName id="BC91930E0CB552A76B1E5DB6A1A0F1E9" class="Insecta" family="Stratiomyidae" genus="Parastratiosphecomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parastratiosphecomyia stratiosphecomyioides" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="stratiosphecomyioides">Parastratiosphecomyia stratiosphecomyioides</taxonomicName>
as &quot;Bukit Besar, Patani, Peninsular Siam&quot;. However, the labels on all four of the orig
<pageBreakToken pageId="7" pageNumber="8" start="start">inal</pageBreakToken>
<pageBreakToken id="CEA70EF1859B926BE1D80CD22582C76E" pageId="7" pageNumber="8" start="start">inal</pageBreakToken>
syntypes read &quot;Bulsit Besar&quot; rather than
<normalizedToken originalValue="“Bukit”">&quot;Bukit&quot;</normalizedToken>
<normalizedToken id="32B751B2CF919E014CF5797F8BAE4B8E" originalValue="“Bukit”">&quot;Bukit&quot;</normalizedToken>
. Using internet searches I found nothing about a possible locality for Bulsit Besar, but there are several localities called Bukit Besar in peninsular Malaysia (it apparently means &quot;Big Hill&quot; in Malay). Although
<normalizedToken originalValue="“Patani”">&quot;Patani&quot;</normalizedToken>
<normalizedToken id="0FFB35892E13E25D94A3C775939E3F9C" originalValue="“Patani”">&quot;Patani&quot;</normalizedToken>
is not part of the actual specimen labels, it seems that by some means Brunetti was aware that the type locality was in Patani. According to
<normalizedToken originalValue="Websters">Webster's</normalizedToken>
<normalizedToken id="935BB782AA28F586031F682CC067C8A4" originalValue="Websters">Webster's</normalizedToken>
Geographical Dictionary (
<bibRefCitation author="Bethel, JP" journalOrPublisher="G. C. Merriam Company, Springfield, Massachusetts" pageId="15" pageNumber="16" title="Webster's Geographical Dictionary." year="1967">Bethel 1967</bibRefCitation>
<bibRefCitation id="35C99AC84ABA0C34C44E0C0A085C8F09" author="Bethel, JP" journalOrPublisher="G. C. Merriam Company, Springfield, Massachusetts" pageId="15" pageNumber="16" title="Webster's Geographical Dictionary." year="1967">Bethel 1967</bibRefCitation>
: 860) Patani was &quot;formerly, a Malay state in the Malay Penin. under Siamese protection, included among the Malay States; now Pattani prov. in Thailand.&quot; Still a province in southeastern Thailand, Pattani has some mountainous areas, so a locality of 2500 feet, as noted on data labels of some of the syntype specimens, is possible. Therefore, I think it is highly likely that the type locality is in Pattani province.
</paragraph>
</subSubSection>

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@ -1,134 +1,134 @@
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Genus
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<paragraph pageId="3" pageNumber="140">
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Trichochloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichochloritis" order="Stylommatophora" pageId="3" pageNumber="140" phylum="Mollusca" rank="genus">Trichochloritis</taxonomicName>
<subSubSection id="108E70A288D796B2564E1CB25C13FFF5" pageId="3" pageNumber="140" type="reference_group">
<paragraph id="525886C244E277D696AE30905588CEA4" pageId="3" pageNumber="140">
<taxonomicName id="D5D4AE1B514368D52B7DFC8CE17ACD70" class="Gastropoda" family="Camaenidae" genus="Trichochloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichochloritis" order="Stylommatophora" pageId="3" pageNumber="140" phylum="Mollusca" rank="genus">Trichochloritis</taxonomicName>
Pilsbry, 1891: 267.
</paragraph>
<paragraph pageId="3" pageNumber="140">
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Trichochloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichochloritis" order="Stylommatophora" pageId="3" pageNumber="140" phylum="Mollusca" rank="genus">Trichochloritis</taxonomicName>
<paragraph id="461A9DEA8A0B2DDFEC909C5BF2BAE424" pageId="3" pageNumber="140">
<taxonomicName id="CB0F3848F8836941BD428599D9986D12" class="Gastropoda" family="Camaenidae" genus="Trichochloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichochloritis" order="Stylommatophora" pageId="3" pageNumber="140" phylum="Mollusca" rank="genus">Trichochloritis</taxonomicName>
:
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="12" pageNumber="149" pagination="2049 - 2209" refId="B20" refString="Schileyko, AA, 2007. Treatise on Recent terrestrial pulmonate molluscs, Part 15 Oopeltidae, Anadenidae, Arionidae, Philomycidae, Succineidae, Athoracophoridae. . Ruthenica, Supplement 2: 2049 - 2209" title="Treatise on Recent terrestrial pulmonate molluscs, Part 15 Oopeltidae, Anadenidae, Arionidae, Philomycidae, Succineidae, Athoracophoridae." volume="2" year="2007">Schileyko 2007</bibRefCitation>
<bibRefCitation id="04A7A76C85D79118606169CD5224A909" author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="12" pageNumber="149" pagination="2049 - 2209" refId="B20" refString="Schileyko, AA, 2007. Treatise on Recent terrestrial pulmonate molluscs, Part 15 Oopeltidae, Anadenidae, Arionidae, Philomycidae, Succineidae, Athoracophoridae. . Ruthenica, Supplement 2: 2049 - 2209" title="Treatise on Recent terrestrial pulmonate molluscs, Part 15 Oopeltidae, Anadenidae, Arionidae, Philomycidae, Succineidae, Athoracophoridae." volume="2" year="2007">Schileyko 2007</bibRefCitation>
: 2113-2114, fig. 2032 (partim).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="140" type="type species">
<paragraph pageId="3" pageNumber="140">Type species.</paragraph>
<paragraph pageId="3" pageNumber="140">
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix breviseta" order="Stylommatophora" pageId="3" pageNumber="140" phylum="Mollusca" rank="species" species="breviseta">
<emphasis italics="true" pageId="3" pageNumber="140">Helix breviseta</emphasis>
<subSubSection id="E99DD351BE46E5CB4319E3627932C78F" pageId="3" pageNumber="140" type="type species">
<paragraph id="37B49A0AF1B7FDB35B03761F4D4940CA" pageId="3" pageNumber="140">Type species.</paragraph>
<paragraph id="213E232CDE45FBE73FCE65AA6E11DBDA" pageId="3" pageNumber="140">
<taxonomicName id="F64B762A179A87E4F1075ECBE677918A" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix breviseta" order="Stylommatophora" pageId="3" pageNumber="140" phylum="Mollusca" rank="species" species="breviseta">
<emphasis id="4019C66C0BE0B1780E4CF865B4F1F21B" italics="true" pageId="3" pageNumber="140">Helix breviseta</emphasis>
</taxonomicName>
L. Pfeiffer, 1862 by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="141" type="included species">
<paragraph pageId="4" pageNumber="141">
<pageBreakToken pageId="4" pageNumber="141" start="start">Included</pageBreakToken>
<subSubSection id="B6D893129B087490D3964A53FAA08443" pageId="4" pageNumber="141" type="included species">
<paragraph id="509919A6DD1CE6DAC67D327D0ACD4431" pageId="4" pageNumber="141">
<pageBreakToken id="B8D0C6BB475CAEE3B64829F08144A8F0" pageId="4" pageNumber="141" start="start">Included</pageBreakToken>
species.
</paragraph>
<paragraph pageId="4" pageNumber="141">
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix breviseta" order="Stylommatophora" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="breviseta">
<emphasis italics="true" pageId="4" pageNumber="141">Helix breviseta</emphasis>
<paragraph id="E36C1A6485E551BD050C249F1299F82E" pageId="4" pageNumber="141">
<taxonomicName id="A079830CE924E6D8AEE785805608CCA3" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix breviseta" order="Stylommatophora" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="breviseta">
<emphasis id="B801063AF78421E6E424EF31250A76DA" italics="true" pageId="4" pageNumber="141">Helix breviseta</emphasis>
</taxonomicName>
L. Pfeiffer, 1862,
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Trachia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachia penangensis" order="Stylommatophora" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="penangensis">
<emphasis italics="true" pageId="4" pageNumber="141">Trachia penangensis</emphasis>
<taxonomicName id="6BA1EDFF2792C5CAD33C0E3FDDC51348" class="Gastropoda" family="Camaenidae" genus="Trachia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trachia penangensis" order="Stylommatophora" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="penangensis">
<emphasis id="C3E3280A117992AF02D2CCEBD4442271" italics="true" pageId="4" pageNumber="141">Trachia penangensis</emphasis>
</taxonomicName>
Stoliczka, 1873.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="141" type="diagnosis">
<paragraph pageId="4" pageNumber="141">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="141">
<subSubSection id="D63D0435B3422A0FD3B65C017CE1E10C" pageId="4" pageNumber="141" type="diagnosis">
<paragraph id="6FC690BD5405A399FC38DEC6A3B04C0E" pageId="4" pageNumber="141">Diagnosis.</paragraph>
<paragraph id="DFF963D6D34FFEF2F886EC8D29056344" pageId="4" pageNumber="141">
Shell depressed globular, apex not sunken, hairs or hair scars cover the entire shell. Penis thickened, probably with penial verge (?) and a slender, relatively long penial caecum; epiphallus slender, shorter than penis; retractor muscle attached at the penis-epiphallus transition; flagellum short; vagina slender, shorter than penis; stalk of bursa copulatrix long, with thickened base and oval bursa (based on the drawings of
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="12" pageNumber="149" pagination="11 - 38" publicationUrl="https://biodiversitylibrary.org/page/35545891" refId="B24" refString="Stoliczka, F, 1873. On the land-shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea. . Journal of the Asiatic Society of Bengal 42: 11 - 38" title="On the land-shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea." url="https://biodiversitylibrary.org/page/35545891" volume="42" year="1873">Stoliczka 1873</bibRefCitation>
<bibRefCitation id="CA7515FD3036C34F7C7E0BDBC2C106CD" author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="12" pageNumber="149" pagination="11 - 38" publicationUrl="https://biodiversitylibrary.org/page/35545891" refId="B24" refString="Stoliczka, F, 1873. On the land-shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea. . Journal of the Asiatic Society of Bengal 42: 11 - 38" title="On the land-shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea." url="https://biodiversitylibrary.org/page/35545891" volume="42" year="1873">Stoliczka 1873</bibRefCitation>
: plate 3, fig. 18 and
<bibRefCitation author="Collinge, WE" journalOrPublisher="Fasciculi Malayenses, Zoology" pageId="11" pageNumber="148" pagination="205 - 218" publicationUrl="https://biodiversitylibrary.org/page/52164094" refId="B7" refString="Collinge, WE, 1903.: Report on the non-operculate land Mollusca. . Fasciculi Malayenses, Zoology 2: 205 - 218" title=": Report on the non-operculate land Mollusca." url="https://biodiversitylibrary.org/page/52164094" volume="2" year="1903">Collinge 1903</bibRefCitation>
<bibRefCitation id="F07EDE336FAE0D586E5D23C1DE2E0E69" author="Collinge, WE" journalOrPublisher="Fasciculi Malayenses, Zoology" pageId="11" pageNumber="148" pagination="205 - 218" publicationUrl="https://biodiversitylibrary.org/page/52164094" refId="B7" refString="Collinge, WE, 1903.: Report on the non-operculate land Mollusca. . Fasciculi Malayenses, Zoology 2: 205 - 218" title=": Report on the non-operculate land Mollusca." url="https://biodiversitylibrary.org/page/52164094" volume="2" year="1903">Collinge 1903</bibRefCitation>
: plate 12, fig. 17.).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="141" type="remarks">
<paragraph pageId="4" pageNumber="141">Remarks.</paragraph>
<paragraph pageId="4" pageNumber="141">
<subSubSection id="3CDC605DF8264A6B34A171061BC086A7" pageId="4" pageNumber="141" type="remarks">
<paragraph id="D20B11854FA483FE8AB669BD79D8DEC1" pageId="4" pageNumber="141">Remarks.</paragraph>
<paragraph id="33A27305E5728C271567F041F78D7813" pageId="4" pageNumber="141">
The anatomy of the genital organs of
<taxonomicName class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Trachia) malayana" order="Stylommatophora" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="malayana" subGenus="Trachia">Helix (Trachia) malayana</taxonomicName>
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
<taxonomicName id="E404A58BD0419C4A300D7691B8764838" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix (Trachia) malayana" order="Stylommatophora" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="malayana" subGenus="Trachia">Helix (Trachia) malayana</taxonomicName>
<normalizedToken id="246DC331F9408F40B571E236165AACD7" originalValue="Möllendorff">Moellendorff</normalizedToken>
, 1887 (=
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<emphasis italics="true" pageId="4" pageNumber="141">Trichochloritis breviseta</emphasis>
<taxonomicName id="EA726E65899AE06FD678D3561328AC82" class="Gastropoda" family="Camaenidae" genus="Trichochloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichochloritis breviseta" order="Stylommatophora" pageId="4" pageNumber="141" phylum="Mollusca" rank="species" species="breviseta">
<emphasis id="5C60AA0CC3722952CE036E24C33FEE63" italics="true" pageId="4" pageNumber="141">Trichochloritis breviseta</emphasis>
</taxonomicName>
; see
<bibRefCitation pageId="4" pageNumber="141" refId="B10">Maassen 2001</bibRefCitation>
<bibRefCitation id="299190104AEDF1CDA7B627C3E38CE348" pageId="4" pageNumber="141" refId="B10">Maassen 2001</bibRefCitation>
) was described by
<bibRefCitation author="Collinge, WE" journalOrPublisher="Fasciculi Malayenses, Zoology" pageId="11" pageNumber="148" pagination="205 - 218" publicationUrl="https://biodiversitylibrary.org/page/52164094" refId="B7" refString="Collinge, WE, 1903.: Report on the non-operculate land Mollusca. . Fasciculi Malayenses, Zoology 2: 205 - 218" title=": Report on the non-operculate land Mollusca." url="https://biodiversitylibrary.org/page/52164094" volume="2" year="1903">Collinge (1903)</bibRefCitation>
<bibRefCitation id="F25DB77F008F4167D3DA8CF0EE2C944B" author="Collinge, WE" journalOrPublisher="Fasciculi Malayenses, Zoology" pageId="11" pageNumber="148" pagination="205 - 218" publicationUrl="https://biodiversitylibrary.org/page/52164094" refId="B7" refString="Collinge, WE, 1903.: Report on the non-operculate land Mollusca. . Fasciculi Malayenses, Zoology 2: 205 - 218" title=": Report on the non-operculate land Mollusca." url="https://biodiversitylibrary.org/page/52164094" volume="2" year="1903">Collinge (1903)</bibRefCitation>
, and that of
<taxonomicName lsidName="T. penangensis" pageId="4" pageNumber="141" rank="species" species="penangensis">
<emphasis italics="true" pageId="4" pageNumber="141">T. penangensis</emphasis>
<taxonomicName id="331EFA1A75C71F35CFB7335BB3C5387D" lsidName="T. penangensis" pageId="4" pageNumber="141" rank="species" species="penangensis">
<emphasis id="5FA51C28A05D60C8BF055F3A328FC1EA" italics="true" pageId="4" pageNumber="141">T. penangensis</emphasis>
</taxonomicName>
is known from
<bibRefCitation author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="12" pageNumber="149" pagination="11 - 38" publicationUrl="https://biodiversitylibrary.org/page/35545891" refId="B24" refString="Stoliczka, F, 1873. On the land-shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea. . Journal of the Asiatic Society of Bengal 42: 11 - 38" title="On the land-shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea." url="https://biodiversitylibrary.org/page/35545891" volume="42" year="1873">Stoliczka (1873)</bibRefCitation>
<bibRefCitation id="C7C3C3DCBF7CA9958556022580C97D18" author="Stoliczka, F" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="12" pageNumber="149" pagination="11 - 38" publicationUrl="https://biodiversitylibrary.org/page/35545891" refId="B24" refString="Stoliczka, F, 1873. On the land-shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea. . Journal of the Asiatic Society of Bengal 42: 11 - 38" title="On the land-shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea." url="https://biodiversitylibrary.org/page/35545891" volume="42" year="1873">Stoliczka (1873)</bibRefCitation>
, here re-drawn and provided in
<figureCitation captionStart="Figures 89" captionStartId="F4" captionText="Figures 8 - 9. Morphology of the genital organs of Trichochloritis species 8 Trichochloritis penangensis (Stoliczka, 1873) (redrawn from Stoliczka 1873) 9 Trichochloritis breviseta (L. Pfeiffer, 1862) (redrawn from Collinge 1903). Not to scale." figureDoi="10.3897/zookeys.865.36296.figures8-9" httpUri="https://binary.pensoft.net/fig/319422" pageId="4" pageNumber="141">Fig. 8</figureCitation>
<figureCitation id="A38C8989C480C2936632FB8F4EAFF9D2" captionStart="Figures 89" captionStartId="F4" captionText="Figures 8 - 9. Morphology of the genital organs of Trichochloritis species 8 Trichochloritis penangensis (Stoliczka, 1873) (redrawn from Stoliczka 1873) 9 Trichochloritis breviseta (L. Pfeiffer, 1862) (redrawn from Collinge 1903). Not to scale." figureDoi="10.3897/zookeys.865.36296.figures8-9" httpUri="https://binary.pensoft.net/fig/319422" pageId="4" pageNumber="141">Fig. 8</figureCitation>
(
<taxonomicName lsidName="penangensis" pageId="4" pageNumber="141" rank="species" species="penangensis">
<emphasis italics="true" pageId="4" pageNumber="141">penangensis</emphasis>
<taxonomicName id="9C00D922FA4493A418CD4122935B8450" lsidName="penangensis" pageId="4" pageNumber="141" rank="species" species="penangensis">
<emphasis id="3620B23952F452B78CC6919BE1AC108B" italics="true" pageId="4" pageNumber="141">penangensis</emphasis>
</taxonomicName>
) and
<figureCitation captionStart="Figures 89" captionStartId="F4" captionText="Figures 8 - 9. Morphology of the genital organs of Trichochloritis species 8 Trichochloritis penangensis (Stoliczka, 1873) (redrawn from Stoliczka 1873) 9 Trichochloritis breviseta (L. Pfeiffer, 1862) (redrawn from Collinge 1903). Not to scale." figureDoi="10.3897/zookeys.865.36296.figures8-9" httpUri="https://binary.pensoft.net/fig/319422" pageId="4" pageNumber="141">Fig. 9</figureCitation>
<figureCitation id="16D9D4B3D782C0613805E722FB8C0841" captionStart="Figures 89" captionStartId="F4" captionText="Figures 8 - 9. Morphology of the genital organs of Trichochloritis species 8 Trichochloritis penangensis (Stoliczka, 1873) (redrawn from Stoliczka 1873) 9 Trichochloritis breviseta (L. Pfeiffer, 1862) (redrawn from Collinge 1903). Not to scale." figureDoi="10.3897/zookeys.865.36296.figures8-9" httpUri="https://binary.pensoft.net/fig/319422" pageId="4" pageNumber="141">Fig. 9</figureCitation>
(
<taxonomicName lsidName="breviseta" pageId="4" pageNumber="141" rank="species" species="breviseta">
<emphasis italics="true" pageId="4" pageNumber="141">breviseta</emphasis>
<taxonomicName id="D765283AEB7E52E3DD441F52CFBC2678" lsidName="breviseta" pageId="4" pageNumber="141" rank="species" species="breviseta">
<emphasis id="E7F0F405D71D36C60E672C7ED531DC36" italics="true" pageId="4" pageNumber="141">breviseta</emphasis>
</taxonomicName>
). Both species possess a penial caecum, which is here considered as a diagnostic trait for the genus. Without knowing the full anatomy, it is uncertain how many of the hairy
<taxonomicName class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis" order="Stylommatophora" pageId="4" pageNumber="141" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="141">Chloritis</emphasis>
<taxonomicName id="27F5CB21134A9E081CD1974136215DB4" class="Gastropoda" family="Camaenidae" genus="Chloritis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chloritis" order="Stylommatophora" pageId="4" pageNumber="141" phylum="Mollusca" rank="genus">
<emphasis id="4DC08800449B197A4053D37AFB8BF45A" italics="true" pageId="4" pageNumber="141">Chloritis</emphasis>
</taxonomicName>
-like species of continental Asia belong to this group.
</paragraph>

View file

@ -1,119 +1,119 @@
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<treatment id="BB91D6E81D5038233C0251653C80F6E1" ID-GBIF-Taxon="152017884" LSID="urn:lsid:plazi:treatment:BB91D6E81D5038233C0251653C80F6E1" httpUri="http://treatment.plazi.org/id/BB91D6E81D5038233C0251653C80F6E1" lastPageNumber="39" pageId="6" pageNumber="39" scope_family="Aphelinidae" scope_order="Hymenoptera" scope_superFamily="Chalcidoidea">
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Encarsia narroi
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&amp;
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, 2000
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<paragraph pageId="6" pageNumber="39">
<taxonomicName authorityName="Gomez &amp; Garcia" authorityYear="2000" class="Insecta" family="Aphelinidae" genus="Encarsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Encarsia narroi" order="Hymenoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="narroi">Encarsia narroi</taxonomicName>
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<taxonomicName id="6E1AE99712B61290B2B35BAC70309356" authorityName="Gomez &amp; Garcia" authorityYear="2000" class="Insecta" family="Aphelinidae" genus="Encarsia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Encarsia narroi" order="Hymenoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="narroi">Encarsia narroi</taxonomicName>
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&amp;
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<normalizedToken id="73749787DA0C475E63F95F77F2B06AA6" originalValue="García">Garcia</normalizedToken>
, 2000: 49. Holotype female. Mexico: Parras, Coahuila, 1500 masl, 11.ii.1995,
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and
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, ex.
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<emphasis italics="true" pageId="6" pageNumber="39">Aleurodicus</emphasis>
<taxonomicName id="B7B16D48598B6D3AF8B037944AA9EC17" authorityName="Douglas" authorityYear="1892" class="Insecta" family="Aphelinidae" genus="Aleurodicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleurodicus" order="Hymenoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="417C739D1A7A9B91064B32C8E0D1DFC0" italics="true" pageId="6" pageNumber="39">Aleurodicus</emphasis>
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sp., on
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<emphasis italics="true" pageId="6" pageNumber="39">Bauhinia variegata</emphasis>
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<emphasis id="4B0E35DE932FFC72A36F63E068BCD9FB" italics="true" pageId="6" pageNumber="39">Bauhinia variegata</emphasis>
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, in NRCBC;
<bibRefCitation author="Schmidt, S" journalOrPublisher="Systematic Entomology" pageId="12" pageNumber="45" pagination="81 - 94" publicationUrl="10.1111/j.1365-3113.2006.00364.x" refId="B36" refString="Schmidt, S, Polaszek, A, 2007. Encarsia or Encarsiella? - redefining generic limits based on morphological and molecular evidence (Hymenoptera, Aphelinidae). Systematic Entomology 32: 81 - 94, DOI: 10.1111/j.1365-3113.2006.00364.x" title="Encarsia or Encarsiella? - redefining generic limits based on morphological and molecular evidence (Hymenoptera, Aphelinidae)." url="10.1111/j.1365-3113.2006.00364.x" volume="32" year="2007">Schmidt and Polaszek 2007</bibRefCitation>
<bibRefCitation id="3A42D259CA8CB78993652F522E29C684" author="Schmidt, S" journalOrPublisher="Systematic Entomology" pageId="12" pageNumber="45" pagination="81 - 94" publicationUrl="10.1111/j.1365-3113.2006.00364.x" refId="B36" refString="Schmidt, S, Polaszek, A, 2007. Encarsia or Encarsiella? - redefining generic limits based on morphological and molecular evidence (Hymenoptera, Aphelinidae). Systematic Entomology 32: 81 - 94, DOI: 10.1111/j.1365-3113.2006.00364.x" title="Encarsia or Encarsiella? - redefining generic limits based on morphological and molecular evidence (Hymenoptera, Aphelinidae)." url="10.1111/j.1365-3113.2006.00364.x" volume="32" year="2007">Schmidt and Polaszek 2007</bibRefCitation>
: 81.
</paragraph>
<paragraph pageId="6" pageNumber="39">
<taxonomicName class="Insecta" family="Aphelinidae" genus="Encarsiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Encarsiella narroi" order="Hymenoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="narroi">Encarsiella narroi</taxonomicName>
<paragraph id="0F4EB26E72A584971F137DA28AB4FCD6" pageId="6" pageNumber="39">
<taxonomicName id="6F73FE36DF56C8AF6A1EF5CA6A968DEE" class="Insecta" family="Aphelinidae" genus="Encarsiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Encarsiella narroi" order="Hymenoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="narroi">Encarsiella narroi</taxonomicName>
(
<normalizedToken originalValue="Gómez">Gomez</normalizedToken>
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&amp;
<normalizedToken originalValue="García">Garcia</normalizedToken>
<normalizedToken id="6A120842DB40D80EAEF246246BD04031" originalValue="García">Garcia</normalizedToken>
);
<bibRefCitation author="Myartseva, SN" journalOrPublisher="Folia Entomologica Mexicana" pageId="12" pageNumber="45" pagination="227 - 232" refId="B25" refString="Myartseva, SN, Coronado-Blanco, JM, 2004. A new species of Encarsiella Hayat (Hymenoptera: Aphelinidae) with a key to Mexican species. Folia Entomologica Mexicana 43 (2): 227 - 232" title="A new species of Encarsiella Hayat (Hymenoptera: Aphelinidae) with a key to Mexican species." volume="43" year="2004">Myartseva and Coronado-Blanco 2004</bibRefCitation>
<bibRefCitation id="06ABD3BB5296A8E92BBD47706F4E8617" author="Myartseva, SN" journalOrPublisher="Folia Entomologica Mexicana" pageId="12" pageNumber="45" pagination="227 - 232" refId="B25" refString="Myartseva, SN, Coronado-Blanco, JM, 2004. A new species of Encarsiella Hayat (Hymenoptera: Aphelinidae) with a key to Mexican species. Folia Entomologica Mexicana 43 (2): 227 - 232" title="A new species of Encarsiella Hayat (Hymenoptera: Aphelinidae) with a key to Mexican species." volume="43" year="2004">Myartseva and Coronado-Blanco 2004</bibRefCitation>
: 624.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="39" type="diagnosis">
<paragraph pageId="6" pageNumber="39">Diagnosis.</paragraph>
<paragraph pageId="6" pageNumber="39">
<subSubSection id="4C3F1D6356B8EEBCD19D1B47CB06A00D" pageId="6" pageNumber="39" type="diagnosis">
<paragraph id="B89344BCC0EB9DE46CE0E09C22EA0D00" pageId="6" pageNumber="39">Diagnosis.</paragraph>
<paragraph id="56D51C69748010B62AAFF8C939D4CCB1" pageId="6" pageNumber="39">
Female: body brown, scutellum pale yellow, antennal funicle and legs pale yellow, hind femora infuscate. Pedicel subequal in length to first segment of funicle (Fig.
<figureCitation captionStart="Figures 16" captionStartId="F1" captionText="Figures 1 - 6. Antennae of Encarsia spp.: 1 Encarsia aleurodici, female 2 male 3 Encarsia andrewi, female 4 male 5 Encarsia magniclava, female 6 Encarsia narroi, female." figureDoi="10.3897/JHR.39.7307.figures1-6" httpUri="https://binary.pensoft.net/fig/32142" pageId="6" pageNumber="39">6</figureCitation>
<figureCitation id="EDBBFF702DF424EA6FA1B4C907C82AF7" captionStart="Figures 16" captionStartId="F1" captionText="Figures 1 - 6. Antennae of Encarsia spp.: 1 Encarsia aleurodici, female 2 male 3 Encarsia andrewi, female 4 male 5 Encarsia magniclava, female 6 Encarsia narroi, female." figureDoi="10.3897/JHR.39.7307.figures1-6" httpUri="https://binary.pensoft.net/fig/32142" pageId="6" pageNumber="39">6</figureCitation>
). Fore wing hyaline, with small asetose area below stigmal vein, about 2.3 times as long as wide, with 10 basal group setae. Mid tibial spur 0.7 times as long as basitarsus. Male: unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="39" type="distribution">
<paragraph pageId="6" pageNumber="39">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="39">Mexico (Coahuila).</paragraph>
<subSubSection id="5CDDCF554F98648899ACCF314E3C7461" pageId="6" pageNumber="39" type="distribution">
<paragraph id="C1EC431FAAAE06E2B992511786397E11" pageId="6" pageNumber="39">Distribution.</paragraph>
<paragraph id="48662892064C3E0C684142029C322996" pageId="6" pageNumber="39">Mexico (Coahuila).</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="39" type="hosts">
<paragraph pageId="6" pageNumber="39">Hosts.</paragraph>
<paragraph pageId="6" pageNumber="39">
<taxonomicName authorityName="Douglas" authorityYear="1892" class="Insecta" family="Aphelinidae" genus="Aleurodicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleurodicus" order="Hemiptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="6" pageNumber="39">Aleurodicus</emphasis>
<subSubSection id="9F75673BDBFE29F07145B9BA614A0C83" pageId="6" pageNumber="39" type="hosts">
<paragraph id="81ADD110C773FA46C4C958DCEA7CADE6" pageId="6" pageNumber="39">Hosts.</paragraph>
<paragraph id="F59A4B257CCF57209B0500F57D69CAD7" pageId="6" pageNumber="39">
<taxonomicName id="1543D87F413E38C4891D1D4391DE20E0" authorityName="Douglas" authorityYear="1892" class="Insecta" family="Aphelinidae" genus="Aleurodicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aleurodicus" order="Hemiptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="8423DC1460142E3205FF551620DFE5ED" italics="true" pageId="6" pageNumber="39">Aleurodicus</emphasis>
</taxonomicName>
sp.
</paragraph>

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<mods:date>2015</mods:date>
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<mods:number>483</mods:number>
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<mods:date id="BC2B33B70326F0A0CF28A8DEA1B6EF61">2015</mods:date>
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<treatment ID-GBIF-Taxon="152058318" LSID="urn:lsid:zoobank.org:act:F4C3293E-92D1-4AA9-9473-BFC285D2E0D8" httpUri="http://treatment.plazi.org/id/BBA18EEBAD2EB23AEF9C028E5E45C798" lastPageNumber="29" pageId="6" pageNumber="29">
<subSubSection pageId="6" pageNumber="29" type="multiple">
<paragraph pageId="6" pageNumber="29">Taxon classification Animalia Hymenoptera Formicidae</paragraph>
<treatment id="BBA18EEBAD2EB23AEF9C028E5E45C798" ID-GBIF-Taxon="152058318" LSID="urn:lsid:zoobank.org:act:F4C3293E-92D1-4AA9-9473-BFC285D2E0D8" httpUri="http://treatment.plazi.org/id/BBA18EEBAD2EB23AEF9C028E5E45C798" lastPageNumber="29" pageId="6" pageNumber="29" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="811C49D595FBE05F1FA9575FE91BB4C3" pageId="6" pageNumber="29" type="multiple">
<paragraph id="26E3F3632AD4EF0DADE71792F5A092A3" pageId="6" pageNumber="29">Taxon classification Animalia Hymenoptera Formicidae</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="nomenclature">
<paragraph pageId="6" pageNumber="29">
<taxonomicName LSID="http://zoobank.org/F4C3293E-92D1-4AA9-9473-BFC285D2E0D8" authority="Prebus" class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax brevidentis" order="Hymenoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="brevidentis">Temnothorax brevidentis Prebus</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="29">sp. n.</taxonomicNameLabel>
<subSubSection id="6BF1CDADB5EC871EF5FA82D1ECCE3CE1" pageId="6" pageNumber="29" type="nomenclature">
<paragraph id="C85662740488D5F926B3D582DBB91B3C" pageId="6" pageNumber="29">
<taxonomicName id="879A245990F8D202F6202CAB3E377FB0" LSID="http://zoobank.org/F4C3293E-92D1-4AA9-9473-BFC285D2E0D8" authority="Prebus" genus="Temnothorax" lsidName="Temnothorax brevidentis" pageId="6" pageNumber="29" rank="species" species="brevidentis">Temnothorax brevidentis Prebus</taxonomicName>
<taxonomicNameLabel id="2A6833ACC1E6E4A00D7193D635605930" pageId="6" pageNumber="29">sp. n.</taxonomicNameLabel>
Figs 4a, 5
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="type material">
<paragraph pageId="6" pageNumber="29">Type material.</paragraph>
<paragraph pageId="6" pageNumber="29">
<subSubSection id="C6CEAC427EE690B23E3F91F1A437D501" pageId="6" pageNumber="29" type="type material">
<paragraph id="266DE6DD9856E2DF89C50F3F64526BDC" pageId="6" pageNumber="29">Type material.</paragraph>
<paragraph id="EDF6CA3CB6779234CC6A1E3EF0CDBA74" pageId="6" pageNumber="29">
Holotype worker, KENYA, Laikipia District, Mpala Research Centre, 1650 m,
<geoCoordinate direction="north" orientation="latitude" precision="555" value="0.29">0.29°N</geoCoordinate>
<geoCoordinate id="FBA142824B2E0B197E495660EBABB083" direction="north" orientation="latitude" precision="555" value="0.29">0.29°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="36.9">36.90°E</geoCoordinate>
<geoCoordinate id="794730FD106A3E95EBB673B98AD59D25" direction="east" orientation="longitude" precision="555" value="36.9">36.90°E</geoCoordinate>
, Acacia woodland, stray in litter, collection code #99-056, 24.IX.1999 (R.R. Snelling) (BMNH: CASENT0712603).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="diagnosis">
<paragraph pageId="6" pageNumber="29">Diagnosis.</paragraph>
<paragraph pageId="6" pageNumber="29">
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax brevidentis" order="Hymenoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="brevidentis">Temnothorax brevidentis</taxonomicName>
<subSubSection id="6842EE4ADD555228A3E3D47CA6BBFD89" pageId="6" pageNumber="29" type="diagnosis">
<paragraph id="FE1BC8AA23227D68D8D647FF7AED7FFA" pageId="6" pageNumber="29">Diagnosis.</paragraph>
<paragraph id="D24AD995F3B172321D62CB32501CB583" pageId="6" pageNumber="29">
<taxonomicName id="3E04DD2C919FD37AE7E80247BD5407C1" genus="Temnothorax" lsidName="Temnothorax brevidentis" pageId="6" pageNumber="29" rank="species" species="brevidentis">Temnothorax brevidentis</taxonomicName>
is easily distinguishable from the other Afrotropical species by the following character combination:
</paragraph>
<paragraph pageId="6" pageNumber="29">Antennal scapes surpassing the posterior margin of the head by the length of the first funicular segment; postpetiole widest at the anterior 1/3 of the segment; setae on the posterior margin of the first gastric tergite separated by about their own length; posterior margin of head rounded; metanotal groove shallowly impressed; compound eyes moderate in size; propodeal spines short.</paragraph>
<paragraph id="843FF3B54687C36AAA6B8EE9E8FFCE2A" pageId="6" pageNumber="29">Antennal scapes surpassing the posterior margin of the head by the length of the first funicular segment; postpetiole widest at the anterior 1/3 of the segment; setae on the posterior margin of the first gastric tergite separated by about their own length; posterior margin of head rounded; metanotal groove shallowly impressed; compound eyes moderate in size; propodeal spines short.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="worker measurements">
<paragraph pageId="6" pageNumber="29">Worker measurements</paragraph>
<paragraph pageId="6" pageNumber="29">(n = 1). EL 0.155; FRS 0.185; HL 0.633; HW 0.479; IOD 0.422; IOcD N/A; MD 0.172; PH 0.186; PL 0.243; PPH 0.196; PPL 0.196; PPW 0.245; PTW 0.144; PW 0.355; SL 0.573; SPST 0.136; WL 0.802.</paragraph>
<paragraph pageId="6" pageNumber="29">Indices: CI 75.7; DPeI 59.3; DPpI 125; LPeI 131; LPpI 100; OI 32.4; PeNI 40.6; PpNI 69; PPI 170; PSLI 21.5; SI 90.5.</paragraph>
<subSubSection id="18A5819102D84E6B0C8869BD6732213D" pageId="6" pageNumber="29" type="worker measurements">
<paragraph id="D28C45217AF9FC1FEF1D8E94EA94B3FE" pageId="6" pageNumber="29">Worker measurements</paragraph>
<paragraph id="6B8BB77388D6024A0C7800AE6F98EFA7" pageId="6" pageNumber="29">(n = 1). EL 0.155; FRS 0.185; HL 0.633; HW 0.479; IOD 0.422; IOcD N/A; MD 0.172; PH 0.186; PL 0.243; PPH 0.196; PPL 0.196; PPW 0.245; PTW 0.144; PW 0.355; SL 0.573; SPST 0.136; WL 0.802.</paragraph>
<paragraph id="66EA98A85DAEC0EDA038AC3D5915607B" pageId="6" pageNumber="29">Indices: CI 75.7; DPeI 59.3; DPpI 125; LPeI 131; LPpI 100; OI 32.4; PeNI 40.6; PpNI 69; PPI 170; PSLI 21.5; SI 90.5.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="worker description">
<paragraph pageId="6" pageNumber="29">Worker description.</paragraph>
<paragraph pageId="6" pageNumber="29">Head longer than wide (CI 76); head sides parallel, but converging toward the mandibular insertions anteriorly beyond the level of the antennal insertions in full-face view; posterior head margin convex and posterior corners of head broadly rounded. Anterior clypeal margin convex and entire, with the median clypeal lobe projecting slightly beyond the lateral clypeal lobes. Frontal carinae developed: extending posteriorly about one-half the length of the compound eye. Antennae 12-segmented,antennal scapes relatively long, extending past the posterior margin of the head by the length of the first funicular segment (SI 120). Eyes moderate in size (OI 32), 8 ommatidia in longest row.</paragraph>
<paragraph pageId="6" pageNumber="29">Mesosoma relatively slender (WL 1.26 times HL); promesonotal suture absent. Metanotal groove narrowly and shallowly impressed. Propodeal spines short and acute (PSLI 21.5); propodeal lobes small and rounded.</paragraph>
<paragraph pageId="6" pageNumber="29">Petiole without a differentiated peduncle. In profile, the anterior face of node forming a shallow concavity anteriorly as it joins the anterior portion of the petiole. Petiolar node in profile relatively low (LPeI 131), and uniformly rounded, without distinct angles between anterior, dorsal, and posterior faces. In dorsal view petiole elongate (DPeI 59). Postpetiole in profile globular (LPpI 100) and roughly the same height as the petiole. Postpetiole in dorsal view transversely elongate-oval, widest at 1/3 of the total postpetiole length from anterior margin (DPpI 125), widest at 1/3 of the total postpetiole length from the anterior margin, and 1.7 times wider than petiole (PPI 170).</paragraph>
<paragraph pageId="6" pageNumber="29">Mandibular sculpture: longitudinally striate along entire length. Clypeus smooth and shiny, bearing 9 longitudinal rugae, with median ruga strongly developed and running posteriorly from the anterior clypeal margin to the level of antennal insertions before weakening. Cephalic dorsum smooth and shining, with overlying well developed, widely spaced longitudinal rugae, which develop cross-linking rugae on the vertex. In profile, head coarsely reticulate; genae with irregular punctures anterior to the compound eye. Sculpture of mesosoma in dorsal view coarsely rugo-reticulate, with interspaces shining. Space between propodeal spines with a single well developed, arcuate transverse carina, which divides the propodeal dorsum from the declivity. Propodeal declivity irregularly shallowly punctate, bordered laterally by weak carinae which run from the ventral margins of the propodeal spines to the propodeal lobes. In profile, mesosoma predominantly finely rugo-reticulate; pronotum with coarse longitudinal rugae. Petiole and postpetiole finely punctate, with fine reticulation present on the dorsal surface of the petiolar node, and faint longitudinal rugae on the postpetiole. Gaster smooth and shining except for small, widely spaced piligerous punctures.</paragraph>
<paragraph pageId="6" pageNumber="29">
<subSubSection id="246DCA85CF93C30CC5D7943CCB25B486" pageId="6" pageNumber="29" type="worker description">
<paragraph id="87DF30D652E9CE94CE676CA3BC7817E2" pageId="6" pageNumber="29">Worker description.</paragraph>
<paragraph id="CA44B4A784CBF053C2895DE5431E7B9A" pageId="6" pageNumber="29">Head longer than wide (CI 76); head sides parallel, but converging toward the mandibular insertions anteriorly beyond the level of the antennal insertions in full-face view; posterior head margin convex and posterior corners of head broadly rounded. Anterior clypeal margin convex and entire, with the median clypeal lobe projecting slightly beyond the lateral clypeal lobes. Frontal carinae developed: extending posteriorly about one-half the length of the compound eye. Antennae 12-segmented,antennal scapes relatively long, extending past the posterior margin of the head by the length of the first funicular segment (SI 120). Eyes moderate in size (OI 32), 8 ommatidia in longest row.</paragraph>
<paragraph id="49CAC7546B78B0FB858BEEC7E1C1941F" pageId="6" pageNumber="29">Mesosoma relatively slender (WL 1.26 times HL); promesonotal suture absent. Metanotal groove narrowly and shallowly impressed. Propodeal spines short and acute (PSLI 21.5); propodeal lobes small and rounded.</paragraph>
<paragraph id="6D680266D19CE0484ECB5191DA539C4D" pageId="6" pageNumber="29">Petiole without a differentiated peduncle. In profile, the anterior face of node forming a shallow concavity anteriorly as it joins the anterior portion of the petiole. Petiolar node in profile relatively low (LPeI 131), and uniformly rounded, without distinct angles between anterior, dorsal, and posterior faces. In dorsal view petiole elongate (DPeI 59). Postpetiole in profile globular (LPpI 100) and roughly the same height as the petiole. Postpetiole in dorsal view transversely elongate-oval, widest at 1/3 of the total postpetiole length from anterior margin (DPpI 125), widest at 1/3 of the total postpetiole length from the anterior margin, and 1.7 times wider than petiole (PPI 170).</paragraph>
<paragraph id="F130C14A61A2A32A6E419183CB312A08" pageId="6" pageNumber="29">Mandibular sculpture: longitudinally striate along entire length. Clypeus smooth and shiny, bearing 9 longitudinal rugae, with median ruga strongly developed and running posteriorly from the anterior clypeal margin to the level of antennal insertions before weakening. Cephalic dorsum smooth and shining, with overlying well developed, widely spaced longitudinal rugae, which develop cross-linking rugae on the vertex. In profile, head coarsely reticulate; genae with irregular punctures anterior to the compound eye. Sculpture of mesosoma in dorsal view coarsely rugo-reticulate, with interspaces shining. Space between propodeal spines with a single well developed, arcuate transverse carina, which divides the propodeal dorsum from the declivity. Propodeal declivity irregularly shallowly punctate, bordered laterally by weak carinae which run from the ventral margins of the propodeal spines to the propodeal lobes. In profile, mesosoma predominantly finely rugo-reticulate; pronotum with coarse longitudinal rugae. Petiole and postpetiole finely punctate, with fine reticulation present on the dorsal surface of the petiolar node, and faint longitudinal rugae on the postpetiole. Gaster smooth and shining except for small, widely spaced piligerous punctures.</paragraph>
<paragraph id="305CD1D965EFB21091415DB11BBDA16D" pageId="6" pageNumber="29">
Head and mandibles nearly uniformly covered in a fine, yellowish pubescence. Dorsal surface of the head, including clypeus, frons and posterior margin of the head equipped with long, blunt-tipped setae. Anterior clypeal margin with two pairs of long setae flanking the median carina. Antennal scapes with short, sharp-tipped suberect pilosity. Pronotal
<normalizedToken originalValue="“neck”">&quot;neck&quot;</normalizedToken>
<normalizedToken id="0D19C3C163B39C7BC964625EC759D83E" originalValue="“neck”">&quot;neck&quot;</normalizedToken>
and pronotal humeri with short, fine yellowish pubescence. Propleurae and procoxae with long, flexous, sharp-tipped pilosity. Dorsal surfaces of mesosoma, waist segments and gaster with uniformly erect, moderately long, abundant and blunt-tipped setae, their bases spaced from each other by the length of the setae or less. Bases of the setae on the posterior margin of first gastral tergite separated by about the length of the setae arising from them. Ventral surface of the gaster with sparse pilosity like that of the propleuron. Ventral surface of the post petiole free of pilosity.
</paragraph>
<caption pageId="6" pageNumber="29">
<paragraph pageId="6" pageNumber="29">
<caption id="420498B55310F442FC736E24FD6A1393" pageId="6" pageNumber="29">
<paragraph id="FD95467A80E6E54A6FFBC058063495E6" pageId="6" pageNumber="29">
Figure 5.
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax brevidentis" order="Hymenoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="brevidentis">Temnothorax brevidentis</taxonomicName>
<taxonomicName id="3BF1FAC1B7B7717BAADB2807A99E5F11" genus="Temnothorax" lsidName="Temnothorax brevidentis" pageId="6" pageNumber="29" rank="species" species="brevidentis">Temnothorax brevidentis</taxonomicName>
sp. n., worker (CASENT0712603) a body in lateral view b body in dorsal view c head in full face view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="queen">
<paragraph pageId="6" pageNumber="29">Queen.</paragraph>
<paragraph pageId="6" pageNumber="29">Unknown</paragraph>
<subSubSection id="C62EB22F30D240BB50DDA140E5D7888B" pageId="6" pageNumber="29" type="queen">
<paragraph id="B91ABC76F2743A723594A3E59B849951" pageId="6" pageNumber="29">Queen.</paragraph>
<paragraph id="585483CE31D563CE1FA54DB6FAE555B1" pageId="6" pageNumber="29">Unknown</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="male">
<paragraph pageId="6" pageNumber="29">Male.</paragraph>
<paragraph pageId="6" pageNumber="29">Unknown</paragraph>
<subSubSection id="FAE06D8F802896D0F754514933D93989" pageId="6" pageNumber="29" type="male">
<paragraph id="C8E2CA608D6C6FFE8906EA4C1B190C9A" pageId="6" pageNumber="29">Male.</paragraph>
<paragraph id="D6697BDE89223784763CA4F44AC70013" pageId="6" pageNumber="29">Unknown</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="color">
<paragraph pageId="6" pageNumber="29">Color.</paragraph>
<paragraph pageId="6" pageNumber="29">Worker: Overall light reddish brown, with head and gaster slightly darker.</paragraph>
<subSubSection id="B7E48D1263FF9667A2CA1EBFCE7C86B0" pageId="6" pageNumber="29" type="color">
<paragraph id="65501AC2178844D4933BBD8E1FBDB470" pageId="6" pageNumber="29">Color.</paragraph>
<paragraph id="8118288E58AB1E7771699A0B77ED32D8" pageId="6" pageNumber="29">Worker: Overall light reddish brown, with head and gaster slightly darker.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="distribution and ecology">
<paragraph pageId="6" pageNumber="29">Distribution and ecology.</paragraph>
<paragraph pageId="6" pageNumber="29">
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax brevidentis" order="Hymenoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="brevidentis">Temnothorax brevidentis</taxonomicName>
<subSubSection id="57A63FE75A4649F5AC501A6BBCB216D7" pageId="6" pageNumber="29" type="distribution and ecology">
<paragraph id="2CB2F5D784A6606A680CFA808BCA252B" pageId="6" pageNumber="29">Distribution and ecology.</paragraph>
<paragraph id="AB38B49E46F9C2093313BE212B565270" pageId="6" pageNumber="29">
<taxonomicName id="8D2D662467B084C4B59BCAE1639A8BB0" genus="Temnothorax" lsidName="Temnothorax brevidentis" pageId="6" pageNumber="29" rank="species" species="brevidentis">Temnothorax brevidentis</taxonomicName>
is known only from acacia woodland leaf litter at the type locality, Mpala Research Centre.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="29" type="taxonomic notes">
<paragraph pageId="6" pageNumber="29">Taxonomic notes.</paragraph>
<paragraph pageId="6" pageNumber="29">
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax brevidentis" order="Hymenoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="brevidentis">Temnothorax brevidentis</taxonomicName>
<subSubSection id="B8C2AF61CCB097FE143817626CDCC8F1" pageId="6" pageNumber="29" type="taxonomic notes">
<paragraph id="97B847B6E00B0803250E06427FFA6E6E" pageId="6" pageNumber="29">Taxonomic notes.</paragraph>
<paragraph id="5AA86C7539A1D682B4A23FD946187AEB" pageId="6" pageNumber="29">
<taxonomicName id="5C25750EBF344AA03AD230E7F85F68E9" genus="Temnothorax" lsidName="Temnothorax brevidentis" pageId="6" pageNumber="29" rank="species" species="brevidentis">Temnothorax brevidentis</taxonomicName>
appears to be very closely related to both
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax cenatus" order="Hymenoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="cenatus">Temnothorax cenatus</taxonomicName>
<taxonomicName id="955E0F74804AC368E78718665055ED85" genus="Temnothorax" lsidName="Temnothorax cenatus" pageId="6" pageNumber="29" rank="species" species="cenatus">Temnothorax cenatus</taxonomicName>
and
<taxonomicName class="Insecta" family="Formicidae" genus="Temnothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Temnothorax rufus" order="Hymenoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="rufus">Temnothorax rufus</taxonomicName>
<taxonomicName id="C716A470648A53B81E5FA14C96947CD8" genus="Temnothorax" lsidName="Temnothorax rufus" pageId="6" pageNumber="29" rank="species" species="rufus">Temnothorax rufus</taxonomicName>
, differing primarily from these in having short propodeal spines.
</paragraph>
</subSubSection>

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<mods:namePart>Clark, Vincent Ralph</mods:namePart>
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<taxonomicName LSID="BBA6CA6E-6839-5FBB-A1DD-169620F20C31" authority="L. Sp. Pl. 1: 70. 1753. var. vivipara Koeler, Descr. Gram. 189. 1802." class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa bulbosa" order="Poales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="bulbosa">Poa bulbosa L. Sp. Pl. 1: 70. 1753. var. vivipara Koeler, Descr. Gram. 189. 1802.</taxonomicName>
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<taxonomicName id="A24550F0B8CB0167F6B3C2AAA77F89B0" ID-CoL="4KLFK" LSID="BBA6CA6E-6839-5FBB-A1DD-169620F20C31" authority="L. Sp. Pl. 1: 70. 1753. var. vivipara Koeler, Descr. Gram. 189. 1802." class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa bulbosa" order="Poales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="bulbosa">Poa bulbosa L. Sp. Pl. 1: 70. 1753. var. vivipara Koeler, Descr. Gram. 189. 1802.</taxonomicName>
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa bulbosa subsp. vivipara" order="Poales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="bulbosa" subSpecies="vivipara">Poa bulbosa subsp. vivipara</taxonomicName>
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<paragraph id="A3FC6DA4658FA015EF4CEE3E340D3677" pageId="0" pageNumber="27">
<taxonomicName id="819037908420BA507C62A233BC809A6B" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa bulbosa subsp. vivipara" order="Poales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="bulbosa" subSpecies="vivipara">Poa bulbosa subsp. vivipara</taxonomicName>
(Koeler) Arcang., Comp. Fl. Ital. 785. 1882.
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<subSubSection pageId="0" pageNumber="27" type="materials_examined">
<paragraph pageId="0" pageNumber="27">Type.</paragraph>
<paragraph pageId="0" pageNumber="27">[Germany. Mainz:] Prope Moguntiam in arenosis (specimen not found).</paragraph>
<paragraph pageId="0" pageNumber="27">
<subSubSection id="6B0A6B82EA3ED79AB20922E92289756E" pageId="0" pageNumber="27" type="materials_examined">
<paragraph id="D0DB8A0502284E83EAF26071CEBD187C" pageId="0" pageNumber="27">Type.</paragraph>
<paragraph id="E963B5D1CFCB7976665702FE1BBAAF05" pageId="0" pageNumber="27">[Germany. Mainz:] Prope Moguntiam in arenosis (specimen not found).</paragraph>
<paragraph id="D7E0A506A530B54943CF7170794B769A" pageId="0" pageNumber="27">
-
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<taxonomicName id="7ACEE3D7BA1B4456D26E26C0B1A3F504" lsidName="P. subsp. sect." pageId="0" pageNumber="27" rank="section" section="Arenariae">P. sect. Arenariae</taxonomicName>
(Hegetschw.) Stapf s.s., Fl. Brit. India 7(22): 338. 1897 [1896]. Type
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<emphasis italics="true" pageId="0" pageNumber="27">P. bulbosa</emphasis>
<taxonomicName id="F0876F020FFEA76C881C98843C5FCD4D" lsidName="P. bulbosa" pageId="0" pageNumber="27" rank="species" species="bulbosa">
<emphasis id="716ECFAE0F634861244C859FF2833A7D" italics="true" pageId="0" pageNumber="27">P. bulbosa</emphasis>
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L.
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<paragraph pageId="0" pageNumber="27">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="27">native to Eurasia and northwest Africa. Introduced/possibly arrived via long-distance-dispersal, but that seems unlikely for the bulbils are bulky and have no special dispersal mechanisms.</paragraph>
<subSubSection id="2078348A05AB0D7A08AC4EB2A5511E5D" pageId="0" pageNumber="27" type="distribution">
<paragraph id="9360E9A7CC124293172711A041D0ED2B" pageId="0" pageNumber="27">Distribution.</paragraph>
<paragraph id="F4509C5BB1C056F1825B6FAB2D4E13FD" pageId="0" pageNumber="27">native to Eurasia and northwest Africa. Introduced/possibly arrived via long-distance-dispersal, but that seems unlikely for the bulbils are bulky and have no special dispersal mechanisms.</paragraph>
</subSubSection>
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<paragraph pageId="0" pageNumber="27">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="27">hemicryptophyte, geophyte, with bulbous based shoots that store hemi-cellulose. Well-adapted to temperate climates with winter rains and dry summers.</paragraph>
<subSubSection id="DD6616C4ADBBC1E21B06F6B1DE24A38B" pageId="0" pageNumber="27" type="ecology">
<paragraph id="44C7E7B8B76E2B41CFB7CD5D5B3AD865" pageId="0" pageNumber="27">Ecology.</paragraph>
<paragraph id="1BC2BF7F0BB43CAAFE1C5F1B592F08B6" pageId="0" pageNumber="27">hemicryptophyte, geophyte, with bulbous based shoots that store hemi-cellulose. Well-adapted to temperate climates with winter rains and dry summers.</paragraph>
</subSubSection>
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<paragraph pageId="0" pageNumber="27">Flowering.</paragraph>
<paragraph pageId="0" pageNumber="27">winter and spring green, flowering in mid-spring and quickly going dormant, flowers mostly forming bulbils. Apomictic.</paragraph>
<subSubSection id="16E7568B7507426106C9BA27B0DB1B0F" pageId="0" pageNumber="27" type="flowering">
<paragraph id="CCFED90D4292A3A123ED08E27EA4F932" pageId="0" pageNumber="27">Flowering.</paragraph>
<paragraph id="89CC70C87B60A28366A2D1DEEF701E01" pageId="0" pageNumber="27">winter and spring green, flowering in mid-spring and quickly going dormant, flowers mostly forming bulbils. Apomictic.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="economics">
<paragraph pageId="0" pageNumber="27">Economics.</paragraph>
<paragraph pageId="0" pageNumber="27">common, excellent early spring forage for sheep, but invasive and can become dominant.</paragraph>
<subSubSection id="08FCAC91A3E703BEC9B3F423DD1C5214" pageId="0" pageNumber="27" type="economics">
<paragraph id="CCB833401D9C36D57C1AA7B1E1BBE695" pageId="0" pageNumber="27">Economics.</paragraph>
<paragraph id="EABB6622C15C5787E082244976154267" pageId="0" pageNumber="27">common, excellent early spring forage for sheep, but invasive and can become dominant.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="vouchers">
<paragraph pageId="0" pageNumber="27">Vouchers.</paragraph>
<paragraph pageId="0" pageNumber="27">no new records.</paragraph>
<subSubSection id="F9D75516D6D2E9924292F2ED052261BA" pageId="0" pageNumber="27" type="vouchers">
<paragraph id="4159ACF3F423801BEDDB58AD9B79FC8F" pageId="0" pageNumber="27">Vouchers.</paragraph>
<paragraph id="6AB316DBA2899A4E2598BBADFEE864C3" pageId="0" pageNumber="27">no new records.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="notes">
<paragraph pageId="0" pageNumber="27">Notes.</paragraph>
<paragraph pageId="0" pageNumber="27">
<subSubSection id="BD47D59DDDB5DF0777AF5D8987C0D0B8" pageId="0" pageNumber="27" type="notes">
<paragraph id="5B644C4AA5038991088E48630A75B3AC" pageId="0" pageNumber="27">Notes.</paragraph>
<paragraph id="0EAA7F92DEEF49DDC0E1D6607DA6F944" pageId="0" pageNumber="27">
All the specimens reviewed at PRE were pseudoviviparous, at least in part. More or less normal-looking lemmas are commonly present in the lower one or two florets of bulbiferous spikelets. The very normal-looking lemmas will have soft hairs on the keel and marginal veins and a tuft of longer hairs on the dorsal side of the callus. The normal florets are thought to be fertile to some degree, although RJS has rarely observed seed in these. Some plants produce more normal florets and more normal spikelets than others, but the main mode of dispersal and establishment is by leafy bulbils that readily root and grow with the next
<normalizedToken originalValue="seasons">seasons'</normalizedToken>
<normalizedToken id="1178CF0C6FCCA62F7D5D382741429F6C" originalValue="seasons">seasons'</normalizedToken>
rains. Some taxonomists decline to recognise infraspecies here, but for purposes of natural history research, it is useful to identify plants with any bulbiferous spikelets as var.
<taxonomicName lsidName="vivipara" pageId="0" pageNumber="27" rank="species" species="vivipara">vivipara</taxonomicName>
<taxonomicName id="F17291A9C05B12AAB93DCD4FCDB375F6" lsidName="vivipara" pageId="0" pageNumber="27" rank="species" species="vivipara">vivipara</taxonomicName>
. Apomictic via bulbifery. 2
<emphasis italics="true" pageId="0" pageNumber="27">n</emphasis>
<emphasis id="FB63DDD5F37FCCCC72611ED64080C2F3" italics="true" pageId="0" pageNumber="27">n</emphasis>
= 21, 28, 29, 31, 32, 33, 34, 35, 37,
<emphasis italics="true" pageId="0" pageNumber="27">42</emphasis>
<emphasis id="27CCB1EF17800F82539F3E72D5BAF4DB" italics="true" pageId="0" pageNumber="27">42</emphasis>
, 44, 46, 48, 49. -
<emphasis bold="true" pageId="0" pageNumber="27">Aa</emphasis>
<emphasis id="FE0BE1E69FF46C735322E7BA94362EA7" bold="true" pageId="0" pageNumber="27">Aa</emphasis>
genotype.
</paragraph>
</subSubSection>

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<taxonomicName LSID="http://species-id.net/wiki/Polymixis_serpentina_minoica" authority="Fibiger, 1992" authorityName="Fibiger" authorityYear="1992" class="Insecta" family="Noctuidae" genus="Polymixis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polymixis serpentina" order="Lepidoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="serpentina" subSpecies="minoica">Polymixis serpentina minoica Fibiger, 1992</taxonomicName>
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Figs 5, 6
</paragraph>
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<paragraph pageId="4" pageNumber="19">Material examined.</paragraph>
<paragraph pageId="4" pageNumber="19">Greece, Crete slide Nos: ♂♂ OP1283m, OP1345m, ♀♀ OP1284f, OP1340f, OP1346f.</paragraph>
<subSubSection id="B572AE9148F4DCAEE8ABDD8D7EEB8DD4" pageId="4" pageNumber="19" type="material examined">
<paragraph id="DC6EE77270123C3A1AAE6EE296835F41" pageId="4" pageNumber="19">Material examined.</paragraph>
<paragraph id="0453659F98A6E0F685A1022EF7D5345D" pageId="4" pageNumber="19">Greece, Crete slide Nos: ♂♂ OP1283m, OP1345m, ♀♀ OP1284f, OP1340f, OP1346f.</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="male genitalia">
<paragraph pageId="4" pageNumber="19">Male genitalia</paragraph>
<paragraph lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<subSubSection id="E273ACB0F021C589F4AA1C5AE047DA91" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="male genitalia">
<paragraph id="5A8ED45BA80805B65DF68C0F2A8D3A60" pageId="4" pageNumber="19">Male genitalia</paragraph>
<paragraph id="D06BD3D8898B6FC6B3AF74FBBD377D58" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
(Figs 21, 22). The genitalia of this taxon were not described in the original description. It was stated that there were no differences between the two subspecies. There are, however, some recognisable differences between the structure of the genitalia of the Cretan and the other populations of
<taxonomicName class="Insecta" family="Noctuidae" genus="Polymixis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polymixis serpentina" order="Lepidoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="serpentina">Polymixis serpentina</taxonomicName>
<taxonomicName id="4959A04EE458864B0F5BFCAB06FBDADD" class="Insecta" family="Noctuidae" genus="Polymixis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polymixis serpentina" order="Lepidoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="species" species="serpentina">Polymixis serpentina</taxonomicName>
. The ground plan of the male genitalia of
<taxonomicName class="Insecta" family="Noctuidae" genus="Polymixis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polymixis serpentina subsp. minoica" order="Lepidoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="serpentina" subSpecies="minoica">Polymixis serpentina minoica</taxonomicName>
<taxonomicName id="42B60B5B0D99AF2EBCB426005DB268C4" class="Insecta" family="Noctuidae" genus="Polymixis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polymixis serpentina subsp. minoica" order="Lepidoptera" pageId="4" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="serpentina" subSpecies="minoica">Polymixis serpentina minoica</taxonomicName>
and the nominotypical subspecies
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the same, but the shape of the juxta is different and all parts of the posterior half of valvae are more massive, being more intensively sclerotized. The posterior extension of juxta is similar to that of
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, but its lateral arms are more elongated, slightly curved, and significantly more divergent from the basal plate. The valvae are wider, and the costa, triangular basal plate, and its extension, appear rough and massive due to their stronger sclerotization;
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has narrower valvae with all parts of their distal sections being finer and thinner.
</paragraph>
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<subSubSection lastPageId="6" lastPageNumber="21" pageId="5" pageNumber="20" type="female genitalia">
<paragraph pageId="5" pageNumber="20">Female genitalia</paragraph>
<paragraph lastPageId="6" lastPageNumber="21" pageId="5" pageNumber="20">
<subSubSection id="449179D35C451982B3C8384ECAF25E56" lastPageId="6" lastPageNumber="21" pageId="5" pageNumber="20" type="female genitalia">
<paragraph id="9243B6B4F9BC6496D2980E6DCB684677" pageId="5" pageNumber="20">Female genitalia</paragraph>
<paragraph id="10E994B64614F345EDF7F85E5985B1FA" lastPageId="6" lastPageNumber="21" pageId="5" pageNumber="20">
(Figs 37-39). The copulatory organ of ssp. minoica differ from those of the ssp. serpentina in the on average shallower concavity on the posterior margin of antrum
<pageBreakToken pageId="6" pageNumber="21" start="start">.</pageBreakToken>
<pageBreakToken id="B5EB33E5EC3AF288CCAB79011DBCA889" pageId="6" pageNumber="21" start="start">.</pageBreakToken>
</paragraph>
<caption pageId="6" pageNumber="21">
<paragraph pageId="6" pageNumber="21">
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Figures 25-30. Female genitalia. 25-27
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<taxonomicName id="AEA8185B05208D2AC23B3AA3D70916B8" class="Insecta" family="Noctuidae" genus="Polymixis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polymixis ivanchiki" order="Lepidoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="ivanchiki">Polymixis ivanchiki</taxonomicName>
sp. n. 25 paratype, Lebanon, slide No. OP0942f 26 paratype, Lebanon, slide No. OP1347f 27 paratype, SE Turkey, slide No. OP1029f 28-30
<taxonomicName class="Insecta" family="Noctuidae" genus="Polymixis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polymixis serpentina subsp. serpentina" order="Lepidoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="serpentina" subSpecies="serpentina">Polymixis serpentina serpentina</taxonomicName>
<taxonomicName id="5C5F96CD3F0BB4BE907D665CBCB6A237" class="Insecta" family="Noctuidae" genus="Polymixis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polymixis serpentina subsp. serpentina" order="Lepidoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="serpentina" subSpecies="serpentina">Polymixis serpentina serpentina</taxonomicName>
28 Slovenia, slide No. OP0940f 29 Croatia, slide No. OP1334f 30 Bulgaria, slide No. OP1335f.
</paragraph>
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<caption pageId="6" pageNumber="21">
<paragraph pageId="6" pageNumber="21">
<caption id="2133143EBE536A6BB8DAF2F94A8B0330" pageId="6" pageNumber="21">
<paragraph id="7B40FD6D5DC6979BA16C564090BEBCCA" pageId="6" pageNumber="21">
Figures 31-36. Female genitalia. 31-36
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<taxonomicName id="439AFF150523A38F10DE6E1BA4479B2A" class="Insecta" family="Noctuidae" genus="Polymixis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polymixis serpentina subsp. serpentina" order="Lepidoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="subSpecies" species="serpentina" subSpecies="serpentina">Polymixis serpentina serpentina</taxonomicName>
31 NE Greece, Kirki, slide No. OP1339f 32 Central Greece, Amfissa, slide No. OP1015f 33 Greece, Peloponnes, slide No. OP1337f 34 Greece, Rhodos, slide No. OP1039f 35 Turkey, Antalya county, slide No. OP1036f 36 Turkey, Icel county, slide No. OP1028f.
</paragraph>
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<mods:affiliation id="8E4CC17BD0EE0B45DA2FB5C787059C40">Musee cantonal de zoologie, Palais de Rumine, Place de la Riponne 6, CH- 1014, Lausanne, Switzerland &amp; Department of Ecology and Evolution, Biophore, University of Lausanne, CH- 1015, Lausanne, Switzerland &amp; School of Zoology, Tel Aviv University, Tel Aviv 6997801, Israel &amp; The Steinhardt Museum of Natural History, Tel Aviv University, Tel Aviv 6997801, Israel</mods:affiliation>
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<treatment ID-GBIF-Taxon="168815851" LSID="urn:lsid:plazi:treatment:BBCCDF446EE95FB48339751CFB7FB531" httpUri="http://treatment.plazi.org/id/BBCCDF446EE95FB48339751CFB7FB531" lastPageNumber="15" pageId="0" pageNumber="15">
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<taxonomicName LSID="BBCCDF44-6EE9-5FB4-8339-751CFB7FB531" authority="Crass, 1947" authorityName="Crass" authorityYear="1947" class="Insecta" family="Oligoneuriidae" genus="Oligoneuriopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligoneuriopsis lawrencei" order="Ephemeroptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="lawrencei">Oligoneuriopsis lawrencei Crass, 1947</taxonomicName>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oligoneuriopsis lawrencei, fore- and hind wings. Scale bar 0.5 mm." figureDoi="10.3897/zookeys.985.56649.figure2" httpUri="https://binary.pensoft.net/fig/471480" pageId="0" pageNumber="15">Figures 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligoneuriopsis lawrencei, ventral view of lower abdomen of adult male, showing lateral extensions of segment IX, gonostyli and penes. Black arrow: Apex of the lateral longitudinal lobe of penis. Grey arrow: Proximal process of penis. Scale bar 0.5 mm." figureDoi="10.3897/zookeys.985.56649.figure3" httpUri="https://binary.pensoft.net/fig/471481" pageId="0" pageNumber="15">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Oligoneuriopsis lawrencei A whole nymph, dorsal view B dorsal view of head. Scale bar: 5 mm (A)." figureDoi="10.3897/zookeys.985.56649.figure4" httpUri="https://binary.pensoft.net/fig/471482" pageId="0" pageNumber="15">, 4</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Hind femora of representative Oligoneuriella (A-C) and Oligoneuriopsis (D-F) species A Oligoneuriella rhenana B Oligoneuriella pallida C Oligoneuriella skoura D Oligoneuriopsis lawrencei E Oligoneuriopsis skhounate F Oligoneuriopsis orontensis." figureDoi="10.3897/zookeys.985.56649.figure14" httpUri="https://binary.pensoft.net/fig/471492" pageId="0" pageNumber="15">, 14D</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Outline of gill plate III of representative Oligoneuriella (A-C) and Oligoneuriopsis (D-F) species A Oligoneuriella rhenana B Oligoneuriella pallida C Oligoneuriella skoura D Oligoneuriopsis lawrencei E Oligoneuriopsis skhounate F Oligoneuriopsis villosus." figureDoi="10.3897/zookeys.985.56649.figure15" httpUri="https://binary.pensoft.net/fig/471493" pageId="0" pageNumber="15">, 15D</figureCitation>
<treatment id="BBCCDF446EE95FB48339751CFB7FB531" ID-GBIF-Taxon="168815851" LSID="urn:lsid:plazi:treatment:BBCCDF446EE95FB48339751CFB7FB531" httpUri="http://treatment.plazi.org/id/BBCCDF446EE95FB48339751CFB7FB531" lastPageNumber="15" pageId="0" pageNumber="15" scope_family="Oligoneuriidae" scope_order="Ephemeroptera">
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<paragraph id="A14F8359164334DA93572617B09E60E7" pageId="0" pageNumber="15">
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<figureCitation id="60547E111F929927A7E49598080514E6" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oligoneuriopsis lawrencei, fore- and hind wings. Scale bar 0.5 mm." figureDoi="10.3897/zookeys.985.56649.figure2" httpUri="https://binary.pensoft.net/fig/471480" pageId="0" pageNumber="15">Figures 2</figureCitation>
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<figureCitation id="DC7E8F474B3C81636275DD3CF8E770C7" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Hind femora of representative Oligoneuriella (A-C) and Oligoneuriopsis (D-F) species A Oligoneuriella rhenana B Oligoneuriella pallida C Oligoneuriella skoura D Oligoneuriopsis lawrencei E Oligoneuriopsis skhounate F Oligoneuriopsis orontensis." figureDoi="10.3897/zookeys.985.56649.figure14" httpUri="https://binary.pensoft.net/fig/471492" pageId="0" pageNumber="15">, 14D</figureCitation>
<figureCitation id="6C8D946FF69FB9ADB9167C8532287E58" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Outline of gill plate III of representative Oligoneuriella (A-C) and Oligoneuriopsis (D-F) species A Oligoneuriella rhenana B Oligoneuriella pallida C Oligoneuriella skoura D Oligoneuriopsis lawrencei E Oligoneuriopsis skhounate F Oligoneuriopsis villosus." figureDoi="10.3897/zookeys.985.56649.figure15" httpUri="https://binary.pensoft.net/fig/471493" pageId="0" pageNumber="15">, 15D</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="15">
<taxonomicName authorityName="Crass" authorityYear="1947" class="Insecta" family="Oligoneuriidae" genus="Oligoneuriopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligoneuriopsis lawrencei" order="Ephemeroptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="lawrencei">Oligoneuriopsis lawrencei</taxonomicName>
<subSubSection id="F6BA5B1DC29677577DF3651E31111A22" pageId="0" pageNumber="15" type="reference_group">
<paragraph id="2AB16AD8D5319D74587061724E133AAA" pageId="0" pageNumber="15">
<taxonomicName id="9DEB8A98BCD6E29F5EBBD4A2D30DA9C9" authorityName="Crass" authorityYear="1947" class="Insecta" family="Oligoneuriidae" genus="Oligoneuriopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligoneuriopsis lawrencei" order="Ephemeroptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="lawrencei">Oligoneuriopsis lawrencei</taxonomicName>
Crass, 1947: 53, figs 3, 4.
</paragraph>
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<paragraph pageId="0" pageNumber="15">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" italics="true" pageId="0" pageNumber="15">Neotype</emphasis>
<subSubSection id="9BD2C551002E44D4BA24A9892451ECAC" pageId="0" pageNumber="15" type="materials_examined">
<paragraph id="A46A6A1A396C058C7D5A5047364F0AC5" pageId="0" pageNumber="15">Material examined.</paragraph>
<paragraph id="B8639477AAA868271E041BF1126A4CDA" pageId="0" pageNumber="15">
<emphasis id="CB6E8A1695E6AD48699FB8EAB741DC7F" bold="true" italics="true" pageId="0" pageNumber="15">Neotype</emphasis>
: South Africa • 1N; Eastern Cape Province, Tributary of Tyume River, below Tor Doone, Hogsback;
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,
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; alt. 1445 m a.s.l.; 29 Feb. 1992; F.C. de Moor leg.; AMGS; GEN 1097A; H.M. Barber-James design., 2020. This specimen was chosen because it comes from one of
<bibRefCitation author="Crass, RS" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="15" pagination="37 - 110" refId="B9" refString="Crass, RS, 1947. The May-flies (Ephemeroptera) of Natal and the Eastern Cape. Annals of the Natal Museum 11: 37 - 110" title="The May-flies (Ephemeroptera) of Natal and the Eastern Cape." volume="11" year="1947">Crass (1947)</bibRefCitation>
<bibRefCitation id="98EEB03D7336804748EC3710C66A29C6" author="Crass, RS" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="15" pagination="37 - 110" refId="B9" refString="Crass, RS, 1947. The May-flies (Ephemeroptera) of Natal and the Eastern Cape. Annals of the Natal Museum 11: 37 - 110" title="The May-flies (Ephemeroptera) of Natal and the Eastern Cape." volume="11" year="1947">Crass (1947)</bibRefCitation>
's original localities.
</paragraph>
<caption doi="10.3897/zookeys.985.56649.figure2" httpUri="https://binary.pensoft.net/fig/471480" pageId="0" pageNumber="15" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" pageId="0" pageNumber="15">Figure 2.</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="15">Oligoneuriopsis lawrencei</emphasis>
<caption id="C7011EC4F6476577DA95ADAB39E6071A" doi="10.3897/zookeys.985.56649.figure2" httpUri="https://binary.pensoft.net/fig/471480" pageId="0" pageNumber="15" start="Figure 2" startId="F2">
<paragraph id="2D8E627B30C7A9C0C468CBC6B6D5CD4C" pageId="0" pageNumber="15">
<emphasis id="4FBF3EE782D39E9B7E4B1F434A7743B1" bold="true" pageId="0" pageNumber="15">Figure 2.</emphasis>
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<emphasis id="8855F638407340B987DD510B7885BB32" italics="true" pageId="0" pageNumber="15">Oligoneuriopsis lawrencei</emphasis>
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, fore- and hind wings. Scale bar 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.985.56649.figure3" httpUri="https://binary.pensoft.net/fig/471481" pageId="0" pageNumber="15" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" pageId="0" pageNumber="15">Figure 3.</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="15">Oligoneuriopsis lawrencei</emphasis>
<caption id="F7F9CA28D53D287DA07B78B455190B59" doi="10.3897/zookeys.985.56649.figure3" httpUri="https://binary.pensoft.net/fig/471481" pageId="0" pageNumber="15" start="Figure 3" startId="F3">
<paragraph id="F0E7B167C5230A166AA8D4DF015C72CC" pageId="0" pageNumber="15">
<emphasis id="F8875718FADBF034EE1495F07A827183" bold="true" pageId="0" pageNumber="15">Figure 3.</emphasis>
<taxonomicName id="D79B038D42BC87E45E598EF1E18E20FD" authorityName="Crass" authorityYear="1947" class="Insecta" family="Oligoneuriidae" genus="Oligoneuriopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligoneuriopsis lawrencei" order="Ephemeroptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="0BBB2DAA6D1ECECFC6E47A9C23101FEC" italics="true" pageId="0" pageNumber="15">Oligoneuriopsis lawrencei</emphasis>
</taxonomicName>
, ventral view of lower abdomen of adult male, showing lateral extensions of segment IX, gonostyli and penes. Black arrow: Apex of the lateral longitudinal lobe of penis. Grey arrow: Proximal process of penis. Scale bar 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" italics="true" pageId="0" pageNumber="15">Other material</emphasis>
<paragraph id="165AC103A30A974A7443DE35FF81E753" pageId="0" pageNumber="15">
<emphasis id="A325D3DDE3C4384B936F9C51A036FE46" bold="true" italics="true" pageId="0" pageNumber="15">Other material</emphasis>
: South Africa • 1N; Eastern Cape Province, Tsitsa River, at &quot;The Falls&quot;;
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,
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; alt. 1140 m a.s.l.; 26 Mar. 1991; F.C. de Moor &amp; H.M. Barber leg.; AMGS; ECR 92AE • 6N; same locality; 28 Mar. 1993; F.C. de Moor &amp; H.M. Barber-James &amp; K. Martens leg.; AMGS; ECR 134A • 12N; Eastern Cape Province, Nqancule River, at Waterval;
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,
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; alt. 1220 m a.s.l.; 24 Mar. 1991; F.C. de Moor &amp; H.M. Barber leg.; AMGS; ECR 86A • 6N; Eastern Cape Province, Nqancule River, at Albany;
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,
<geoCoordinate degrees="28.2153" direction="east" orientation="longitude" precision="5" value="28.2153">28.2153°E</geoCoordinate>
<geoCoordinate id="A56B796BC82BD1F722B97F6E3D6330B1" degrees="28.2153" direction="east" orientation="longitude" precision="5" value="28.2153">28.2153°E</geoCoordinate>
; alt. 1240 m a.s.l.; 5 Mar. 1991; F.C. de Moor &amp; H.M. Barber leg.; AMGS; ECR 88A • 1N; Eastern Cape Province, Kettle Spout waterfall, Tyume River tributary, Hogsback;
<geoCoordinate degrees="32.5500" direction="south" orientation="latitude" precision="5" value="-32.55">32.5500°S</geoCoordinate>
<geoCoordinate id="247ABE8CFAF7EB9E5D2AA89BA0386218" degrees="32.5500" direction="south" orientation="latitude" precision="5" value="-32.55">32.5500°S</geoCoordinate>
,
<geoCoordinate degrees="26.9500" direction="east" orientation="longitude" precision="5" value="26.95">26.9500°E</geoCoordinate>
<geoCoordinate id="8751D5CBE39788B2D2D7C7B33A5A7145" degrees="26.9500" direction="east" orientation="longitude" precision="5" value="26.95">26.9500°E</geoCoordinate>
; alt. 1835 m a.sl.; 19 May 2007; F.C. de Moor and N. Phaliso leg.; AMGS; GEN 1845A • 3♀, 4♂, 1N; KwaZulu-Natal, Klein Mooi River, at Durleigh Farm;
<geoCoordinate degrees="29.2283" direction="south" orientation="latitude" precision="5" value="-29.2283">29.2283°S</geoCoordinate>
<geoCoordinate id="23A2D5A72D8B57BC3B998F3A0A30E021" degrees="29.2283" direction="south" orientation="latitude" precision="5" value="-29.2283">29.2283°S</geoCoordinate>
,
<geoCoordinate degrees="29.8997" direction="east" orientation="longitude" precision="5" value="29.8997">29.8997°E</geoCoordinate>
<geoCoordinate id="10D2E6D90E586744A7E316B310A7C6F3" degrees="29.8997" direction="east" orientation="longitude" precision="5" value="29.8997">29.8997°E</geoCoordinate>
; alt. 1392 m a.s.l.; 15 Mar. 1995; C. Dickens; AMGS; MOI 29BS • 3♀ 4♂; same locality; 3 Apr. 1995; F.C. de Moor leg; AMGS; MOI 35B.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="male imago">
<paragraph pageId="0" pageNumber="15">Male imago.</paragraph>
<paragraph pageId="0" pageNumber="15">Lengths. Body: up to 14.8 mm; forewing: up to 14.9 mm; cerci: up to 17.0 mm; caudal filament: up to 12.8 mm.</paragraph>
<paragraph pageId="0" pageNumber="15">
<subSubSection id="143598729066C684943464E6EBBB16E6" pageId="0" pageNumber="15" type="male imago">
<paragraph id="2D9BC9ED54DAB819411BCF658BE6DB9F" pageId="0" pageNumber="15">Male imago.</paragraph>
<paragraph id="3861B2BF3BB083C9445AF832F76BB674" pageId="0" pageNumber="15">Lengths. Body: up to 14.8 mm; forewing: up to 14.9 mm; cerci: up to 17.0 mm; caudal filament: up to 12.8 mm.</paragraph>
<paragraph id="C4AA8F11BCCC5F58B0E0B1EEF89EC5DA" pageId="0" pageNumber="15">
Vertex light brown, frontoclypeus pale cream, broadly rounded apically, compound eyes black, base of ocelli black, ocelli whitish, antennae with scape and pedicel pale cream, first segment and flagellum light brown. Pronotum light brown, margins suffused with dark brown pigmentation. Pterothorax light brown, with pale cream-coloured unsclerotized line between meso and metanotal plate. Mesoscutellar filaments present. Forelegs shorter than mid or hind legs, with outer margin of femora, tibiae and tarsi dark brown, otherwise uniform pale brown colour; mid- and hindlegs cream to light brown, no distinct markings. All three pairs of legs appear to be functional. Tarsal claws paired, blunt. Wings (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oligoneuriopsis lawrencei, fore- and hind wings. Scale bar 0.5 mm." figureDoi="10.3897/zookeys.985.56649.figure2" httpUri="https://binary.pensoft.net/fig/471480" pageId="0" pageNumber="15">2</figureCitation>
<figureCitation id="D8E7D125A6E0B2222F9F116D661D3525" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Oligoneuriopsis lawrencei, fore- and hind wings. Scale bar 0.5 mm." figureDoi="10.3897/zookeys.985.56649.figure2" httpUri="https://binary.pensoft.net/fig/471480" pageId="0" pageNumber="15">2</figureCitation>
), when folded, light brown, almost whitish when unfolded. Forewing typical of the genus, with five groups of veins: Sc+RA, RSa+iRS, RSp+MA1, MA2+MP1, and a forked MP2+CuA - CuP vein. Subcostal field with numerous transversal veins, those issued from RA not reaching iRS in the distal forth of the length, those between iRS and MA1 only present in the proximal half.
</paragraph>
<paragraph pageId="0" pageNumber="15">
<paragraph id="D039933912BCEC91119C223EF8368F43" pageId="0" pageNumber="15">
Abdominal segments uniformly creamish, without distinct patterns, except tergites VII, IX, and X light brown; lateral margins with spine-shaped extensions from segment III to IX, of increasing length towards the posterior. Gonostyli whitish to grey, cerci whitish. Gonostyli 4-segmented, the basal one ca. 3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="C9E32A968EE950E6F421817C0E1354EC" originalValue="×">x</normalizedToken>
the length of segments 2 to 4 combined. Penis lobes almost triangular, with characteristic sclerotized proximal process ending in a simple projection; apex of the lateral longitudinal lobe of penis in a small club-shaped sclerite (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligoneuriopsis lawrencei, ventral view of lower abdomen of adult male, showing lateral extensions of segment IX, gonostyli and penes. Black arrow: Apex of the lateral longitudinal lobe of penis. Grey arrow: Proximal process of penis. Scale bar 0.5 mm." figureDoi="10.3897/zookeys.985.56649.figure3" httpUri="https://binary.pensoft.net/fig/471481" pageId="0" pageNumber="15">3</figureCitation>
<figureCitation id="CAA419E6BE3B66CA32EC32ACB46CFEED" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Oligoneuriopsis lawrencei, ventral view of lower abdomen of adult male, showing lateral extensions of segment IX, gonostyli and penes. Black arrow: Apex of the lateral longitudinal lobe of penis. Grey arrow: Proximal process of penis. Scale bar 0.5 mm." figureDoi="10.3897/zookeys.985.56649.figure3" httpUri="https://binary.pensoft.net/fig/471481" pageId="0" pageNumber="15">3</figureCitation>
). Cerci with whorls of long setae at each junction (not figured).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="female subimago">
<paragraph pageId="0" pageNumber="15">Female subimago.</paragraph>
<paragraph pageId="0" pageNumber="15">Lengths. Body: up to 17 mm; forewing: up to 18.8 mm; cerci: up to 7 mm; caudal filament: up to 4.5 mm. Colouration as in the male; tibiae and tarsi of all legs appear to be functional. Cerci light to medium brown. Posterior margin of sternite IX deeply concave and rounded.</paragraph>
<subSubSection id="5E47B8CEEB7F0377C0FCFDCEF3EECEFC" pageId="0" pageNumber="15" type="female subimago">
<paragraph id="1149FECD134038EE12E48155200276A8" pageId="0" pageNumber="15">Female subimago.</paragraph>
<paragraph id="C435CA1AF378703826BF7C43DD935EDB" pageId="0" pageNumber="15">Lengths. Body: up to 17 mm; forewing: up to 18.8 mm; cerci: up to 7 mm; caudal filament: up to 4.5 mm. Colouration as in the male; tibiae and tarsi of all legs appear to be functional. Cerci light to medium brown. Posterior margin of sternite IX deeply concave and rounded.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="nymph">
<paragraph pageId="0" pageNumber="15">Nymph.</paragraph>
<paragraph pageId="0" pageNumber="15">
<subSubSection id="9CE4C7FAC5A65EA2031B6AB8877F56E4" pageId="0" pageNumber="15" type="nymph">
<paragraph id="3249EEF5B2691BE1301B527785536038" pageId="0" pageNumber="15">Nymph.</paragraph>
<paragraph id="B1A3B5B0EA433BF4B7DBDAD9427B11FC" pageId="0" pageNumber="15">
Lengths. Body up to 15 mm and 18.5 mm in male and female nymphs respectively; cerci (and caudal filament) up to 9.2 mm (4.0 mm) and 10.1 mm (3.5 mm) in male and female nymphs respectively. General colouration light to medium yellow-brown (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Oligoneuriopsis lawrencei A whole nymph, dorsal view B dorsal view of head. Scale bar: 5 mm (A)." figureDoi="10.3897/zookeys.985.56649.figure4" httpUri="https://binary.pensoft.net/fig/471482" pageId="0" pageNumber="15">4A</figureCitation>
<figureCitation id="75F923E425A92153A21DFA4348335696" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Oligoneuriopsis lawrencei A whole nymph, dorsal view B dorsal view of head. Scale bar: 5 mm (A)." figureDoi="10.3897/zookeys.985.56649.figure4" httpUri="https://binary.pensoft.net/fig/471482" pageId="0" pageNumber="15">4A</figureCitation>
), with dark brown dorso-medial markings, better developed in mature male nymphs than in immatures or females. Head (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Oligoneuriopsis lawrencei A whole nymph, dorsal view B dorsal view of head. Scale bar: 5 mm (A)." figureDoi="10.3897/zookeys.985.56649.figure4" httpUri="https://binary.pensoft.net/fig/471482" pageId="0" pageNumber="15">4B</figureCitation>
<figureCitation id="12E35E08B0CE110AAB69FA6E42AAAE84" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Oligoneuriopsis lawrencei A whole nymph, dorsal view B dorsal view of head. Scale bar: 5 mm (A)." figureDoi="10.3897/zookeys.985.56649.figure4" httpUri="https://binary.pensoft.net/fig/471482" pageId="0" pageNumber="15">4B</figureCitation>
) medium brown, with maculation between the compound eyes. Ventrally, head a uniform pale cream colour. Gills at base of maxillae forming a
<normalizedToken originalValue="“beard”">&quot;beard&quot;</normalizedToken>
<normalizedToken id="EDBF57282F969A80FE5466F4CBD33EB8" originalValue="“beard”">&quot;beard&quot;</normalizedToken>
ventrally at base of head, of similar colour to head in Hogsback specimens, orientated in one plane, parallel to length of body. Pro and mesonotum medium brown, with pale cream-coloured maculae. Legs light brown, femoro-tibial articulation darker, setae of forelegs noticeably darker brown than the legs. Femur and tibia of foreleg shorter than those of mid or hind leg, in all cases, femora and tibiae subequal in length. Setae on the outer margin of mid and hind femora well developed, slightly decreasing in size and reaching the apex (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Hind femora of representative Oligoneuriella (A-C) and Oligoneuriopsis (D-F) species A Oligoneuriella rhenana B Oligoneuriella pallida C Oligoneuriella skoura D Oligoneuriopsis lawrencei E Oligoneuriopsis skhounate F Oligoneuriopsis orontensis." figureDoi="10.3897/zookeys.985.56649.figure14" httpUri="https://binary.pensoft.net/fig/471492" pageId="0" pageNumber="15">14D</figureCitation>
<figureCitation id="23FBF7AB91B84FD30C11B3F58829751A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Hind femora of representative Oligoneuriella (A-C) and Oligoneuriopsis (D-F) species A Oligoneuriella rhenana B Oligoneuriella pallida C Oligoneuriella skoura D Oligoneuriopsis lawrencei E Oligoneuriopsis skhounate F Oligoneuriopsis orontensis." figureDoi="10.3897/zookeys.985.56649.figure14" httpUri="https://binary.pensoft.net/fig/471492" pageId="0" pageNumber="15">14D</figureCitation>
). Tibiae and tarsi with long, even fringe of setae along entire dorsal margin, interspersed with occasional short spine-like setae. Abdominal tergites uniformly medium brown, each with darker brown marking medially; sternites uniform pale brown, with no markings. Dense patch of posteriorly orientated setae ventromedially on abdominal sternites II-IV, much reduced patch on segments V, further reduced on VI. Gills II-VII almost subequal in size, gill I smaller. On all gills except for gill I, fibrillae shorter than lamella length. Lamella of gill I less than half the length of the fibrillar portion. Lamellae II-VII with long and thin setae on their distal inner margin (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Outline of gill plate III of representative Oligoneuriella (A-C) and Oligoneuriopsis (D-F) species A Oligoneuriella rhenana B Oligoneuriella pallida C Oligoneuriella skoura D Oligoneuriopsis lawrencei E Oligoneuriopsis skhounate F Oligoneuriopsis villosus." figureDoi="10.3897/zookeys.985.56649.figure15" httpUri="https://binary.pensoft.net/fig/471493" pageId="0" pageNumber="15">15D</figureCitation>
<figureCitation id="4A842B9FC990B4597FCFA9120F2FDD0F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Outline of gill plate III of representative Oligoneuriella (A-C) and Oligoneuriopsis (D-F) species A Oligoneuriella rhenana B Oligoneuriella pallida C Oligoneuriella skoura D Oligoneuriopsis lawrencei E Oligoneuriopsis skhounate F Oligoneuriopsis villosus." figureDoi="10.3897/zookeys.985.56649.figure15" httpUri="https://binary.pensoft.net/fig/471493" pageId="0" pageNumber="15">15D</figureCitation>
). Posterolateral spines of the abdomen increasing in size posteriorly. Whole nymph (dorsal aspect) and gills as illustrated by
<bibRefCitation DOI="https://doi.org/10.1007/978-1-4613-3066-0_29" author="Agnew, JD" editor="Flannagan, JF" journalOrPublisher="Plenum Press, New York" pageId="0" pageNumber="15" pagination="353 - 366" refId="B3" refString="Agnew, JD, 1980. Ultrastructural studies on Oligoneuriidae - taxonomic applications. In: Flannagan, JF, Marshall, KE, Eds., Advances in Ephemeroptera Biology. Plenum Press, New York: 353 - 366, DOI: https://doi.org/10.1007/978-1-4613-3066-0_29" title="Ultrastructural studies on Oligoneuriidae - taxonomic applications." url="https://doi.org/10.1007/978-1-4613-3066-0_29" volumeTitle="Advances in Ephemeroptera Biology." year="1980">Agnew (1980)</bibRefCitation>
<bibRefCitation id="A31714572C44A82662ED167A9ED97BD8" DOI="https://doi.org/10.1007/978-1-4613-3066-0_29" author="Agnew, JD" editor="Flannagan, JF" journalOrPublisher="Plenum Press, New York" pageId="0" pageNumber="15" pagination="353 - 366" refId="B3" refString="Agnew, JD, 1980. Ultrastructural studies on Oligoneuriidae - taxonomic applications. In: Flannagan, JF, Marshall, KE, Eds., Advances in Ephemeroptera Biology. Plenum Press, New York: 353 - 366, DOI: https://doi.org/10.1007/978-1-4613-3066-0_29" title="Ultrastructural studies on Oligoneuriidae - taxonomic applications." url="https://doi.org/10.1007/978-1-4613-3066-0_29" volumeTitle="Advances in Ephemeroptera Biology." year="1980">Agnew (1980)</bibRefCitation>
. Cerci uniformly medium brown, caudal filament paler brown.
</paragraph>
<caption doi="10.3897/zookeys.985.56649.figure4" httpUri="https://binary.pensoft.net/fig/471482" pageId="0" pageNumber="15" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="15">
<emphasis bold="true" pageId="0" pageNumber="15">Figure 4.</emphasis>
<taxonomicName authorityName="Crass" authorityYear="1947" class="Insecta" family="Oligoneuriidae" genus="Oligoneuriopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligoneuriopsis lawrencei" order="Ephemeroptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis italics="true" pageId="0" pageNumber="15">Oligoneuriopsis lawrencei</emphasis>
<caption id="D0BC58B77F85A1488EC343423BFAC1F2" doi="10.3897/zookeys.985.56649.figure4" httpUri="https://binary.pensoft.net/fig/471482" pageId="0" pageNumber="15" start="Figure 4" startId="F4">
<paragraph id="A576223ACBEE85F0A62828AC6B054446" pageId="0" pageNumber="15">
<emphasis id="BE9A4E5A9EDE08D6B475AF06B79A404F" bold="true" pageId="0" pageNumber="15">Figure 4.</emphasis>
<taxonomicName id="B0D884C52441CF7712EFD2A0647F9291" authorityName="Crass" authorityYear="1947" class="Insecta" family="Oligoneuriidae" genus="Oligoneuriopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oligoneuriopsis lawrencei" order="Ephemeroptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="385C874AA80141B1733212D56058498E" italics="true" pageId="0" pageNumber="15">Oligoneuriopsis lawrencei</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="15">A</emphasis>
<emphasis id="A0DFB74F79B66F66F5F15040CA0D618D" bold="true" pageId="0" pageNumber="15">A</emphasis>
whole nymph, dorsal view
<emphasis bold="true" pageId="0" pageNumber="15">B</emphasis>
<emphasis id="D3FC61CCD4A436E97906A24FA69C124D" bold="true" pageId="0" pageNumber="15">B</emphasis>
dorsal view of head. Scale bar: 5 mm (
<emphasis bold="true" pageId="0" pageNumber="15">A</emphasis>
<emphasis id="4B9A26FEA8F41EB4BF8D91A39E9F8E86" bold="true" pageId="0" pageNumber="15">A</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="intraspecific variation">
<paragraph pageId="0" pageNumber="15">Intraspecific variation.</paragraph>
<paragraph pageId="0" pageNumber="15">Specimens from KwaZulu-Natal are darker brown in colour than specimens with more southerly distribution, and first three abdominal segments darker brown in colour dorsally than remaining segments; faint paired median spots visible on the last five abdominal tergites in some KwaZulu-Natal nymphs. The head also possesses a dark brown marking present on the frons between the ocelli, and maxillary gills are much paler in colour relative to head capsule in KwaZulu-Natal specimens.</paragraph>
<subSubSection id="579B5A7A2C7368BA416E12FE1B329317" pageId="0" pageNumber="15" type="intraspecific variation">
<paragraph id="0614A165BDBE0BE8165F12F3E76BF2E7" pageId="0" pageNumber="15">Intraspecific variation.</paragraph>
<paragraph id="60DE80514B386EB7AADD52247A3E5282" pageId="0" pageNumber="15">Specimens from KwaZulu-Natal are darker brown in colour than specimens with more southerly distribution, and first three abdominal segments darker brown in colour dorsally than remaining segments; faint paired median spots visible on the last five abdominal tergites in some KwaZulu-Natal nymphs. The head also possesses a dark brown marking present on the frons between the ocelli, and maxillary gills are much paler in colour relative to head capsule in KwaZulu-Natal specimens.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="affinities">
<paragraph pageId="0" pageNumber="15">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="15">
<subSubSection id="A6E6E860B064A45D0E1D9B7240765941" pageId="0" pageNumber="15" type="affinities">
<paragraph id="AED456D96EA12CC10DBBAFF577D0E1D4" pageId="0" pageNumber="15">Affinities.</paragraph>
<paragraph id="00E0959C205844C0781533B48A77554A" pageId="0" pageNumber="15">
Winged stages were collected from the same locality as the nymphs in one instance, allowing association of the life stages. Imagos were seldom flying at the times of collection. Nymphs of
<taxonomicName lsidName="O. lawrencei" pageId="0" pageNumber="15" rank="species" species="lawrencei">
<emphasis italics="true" pageId="0" pageNumber="15">O. lawrencei</emphasis>
<taxonomicName id="59BF6BB526B08E6144BD6B218A53C89F" lsidName="O. lawrencei" pageId="0" pageNumber="15" rank="species" species="lawrencei">
<emphasis id="5DD8CE19080249704A7D7D73C061E4EF" italics="true" pageId="0" pageNumber="15">O. lawrencei</emphasis>
</taxonomicName>
have a broadly rounded fronto-clypeal region, which easily distinguishes them from
<taxonomicName lsidName="O. jessicae" pageId="0" pageNumber="15" rank="species" species="jessicae">
<emphasis italics="true" pageId="0" pageNumber="15">O. jessicae</emphasis>
<taxonomicName id="DA65C35B3BD8EED2CB29322C62BBF76B" lsidName="O. jessicae" pageId="0" pageNumber="15" rank="species" species="jessicae">
<emphasis id="97F1D89A549175BC6EF87E0A41DD9FA6" italics="true" pageId="0" pageNumber="15">O. jessicae</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="O. elisabethae" pageId="0" pageNumber="15" rank="species" species="elisabethae">
<emphasis italics="true" pageId="0" pageNumber="15">O. elisabethae</emphasis>
<taxonomicName id="DF90054AF013ECDF366046BD476C3D4C" lsidName="O. elisabethae" pageId="0" pageNumber="15" rank="species" species="elisabethae">
<emphasis id="463002E4749CBF55EADCD6B8082DB3A4" italics="true" pageId="0" pageNumber="15">O. elisabethae</emphasis>
</taxonomicName>
, both of which are more pointed in shape. Head very slightly carinate, less so than in
<taxonomicName lsidName="O. elisabethae" pageId="0" pageNumber="15" rank="species" species="elisabethae">
<emphasis italics="true" pageId="0" pageNumber="15">O. elisabethae</emphasis>
<taxonomicName id="8D0A71410FB910066C524275E55FB39E" lsidName="O. elisabethae" pageId="0" pageNumber="15" rank="species" species="elisabethae">
<emphasis id="BDEFF88ACFA55504B3A7AAD33F2C186D" italics="true" pageId="0" pageNumber="15">O. elisabethae</emphasis>
</taxonomicName>
. Note that this is unlike the strong carination seen in
<taxonomicName authorityName="Eaton" authorityYear="1881" class="Insecta" family="Oligoneuriidae" genus="Elassoneuria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elassoneuria" order="Ephemeroptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="15">Elassoneuria</emphasis>
<taxonomicName id="513BF3EEC81BB05488924E30D9C72F4A" authorityName="Eaton" authorityYear="1881" class="Insecta" family="Oligoneuriidae" genus="Elassoneuria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Elassoneuria" order="Ephemeroptera" pageId="0" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B829DD8C939D9397EA35C2CC1FDACCA4" italics="true" pageId="0" pageNumber="15">Elassoneuria</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="habitat">
<paragraph pageId="0" pageNumber="15">Habitat preference.</paragraph>
<paragraph pageId="0" pageNumber="15">Found under large boulders (400-500 mm diameter) in swift current, often in rivers with bedrock substrate.</paragraph>
<subSubSection id="E34599DB477932FC87BC4A1E03C987A5" pageId="0" pageNumber="15" type="habitat">
<paragraph id="A7D3BB445085D5DD04369DAE641C6BA6" pageId="0" pageNumber="15">Habitat preference.</paragraph>
<paragraph id="F3BE871B556D0B9F020E1AD91C1A2FA3" pageId="0" pageNumber="15">Found under large boulders (400-500 mm diameter) in swift current, often in rivers with bedrock substrate.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="15" type="distribution">
<paragraph pageId="0" pageNumber="15">Known distribution.</paragraph>
<paragraph pageId="0" pageNumber="15">South Africa.</paragraph>
<subSubSection id="799AD1271F2BE6D82B2EB5E9E3200A2A" pageId="0" pageNumber="15" type="distribution">
<paragraph id="AD96481A21104A8C672776111F82854E" pageId="0" pageNumber="15">Known distribution.</paragraph>
<paragraph id="A127AA4C0D0EC9A1969D0FAFC37F9827" pageId="0" pageNumber="15">South Africa.</paragraph>
</subSubSection>
</treatment>
</document>

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Genus
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<taxonomicNameLabel id="0D31C5C489DC9A3C9544D3838C0C7648" pageId="0" pageNumber="49">gen. nov.</taxonomicNameLabel>
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@ -79,7 +79,7 @@ Genus
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<paragraph id="23B217DE91AA3DAD49F32D52496E9291" pageId="0" pageNumber="49">Type species.</paragraph>
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<taxonomicName id="97CEDD8B7A596E29F473B6F43071CB63" authorityName="Kang, Hu &amp; Luo" authorityYear="2024" class="Insecta" family="Apteronotidae" genus="Apteronotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Apteronotus indigus" order="Hymenoptera" pageId="0" pageNumber="49" phylum="Arthropoda" rank="species" species="indigus">
<emphasis id="B7AE566EB54168CD24C09A688558E700" italics="true" pageId="0" pageNumber="49">Apteronotus indigus</emphasis>
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Kang, Hu &amp; Luo, sp. nov.