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: 291, fig. 1, pl. 1;
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: 303 (revised diagnosis);
<bibRefCitation author="Grimaldi, DA" journalOrPublisher="Bulletin of the American Museum of Natural History, 408, New York" pageId="0" pageNumber="295" refId="B12" refString="Grimaldi, DA, 2016. Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History, 408, New York" title="Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII." year="2016">Grimaldi 2016</bibRefCitation>
<bibRefCitation id="D1F468294AB3CF9DD292D9F26C73E1C1" author="Grimaldi, DA" journalOrPublisher="Bulletin of the American Museum of Natural History, 408, New York" pageId="0" pageNumber="295" refId="B12" refString="Grimaldi, DA, 2016. Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History, 408, New York" title="Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII." year="2016">Grimaldi 2016</bibRefCitation>
: 45, fig. 16A (emended diagnosis).
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<subSubSection pageId="0" pageNumber="295" type="diagnosis">
<paragraph pageId="0" pageNumber="295">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="295">Ocellar triangle developed with visible ocelli; basal callus absent; postspiracular scale absent; R5 slightly curved; hind tibia with 2 spurs; tergite X undivided; cercus two-segmented.</paragraph>
<subSubSection id="023FF0277B4DF7FBF9FA2D9CA5AF5AB4" pageId="0" pageNumber="295" type="diagnosis">
<paragraph id="5A86B16DE6E1A069FB1BFF654103B97F" pageId="0" pageNumber="295">Diagnosis.</paragraph>
<paragraph id="69598BF0583D0D038CF25C836E11602E" pageId="0" pageNumber="295">Ocellar triangle developed with visible ocelli; basal callus absent; postspiracular scale absent; R5 slightly curved; hind tibia with 2 spurs; tergite X undivided; cercus two-segmented.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="295" type="comments">
<paragraph pageId="0" pageNumber="295">Comments.</paragraph>
<paragraph pageId="0" pageNumber="295">
<subSubSection id="8B7094F28CE912C3EDB5531D28D7787C" pageId="0" pageNumber="295" type="comments">
<paragraph id="5BADCA3F4FF66A87B97C3F04FA866F6F" pageId="0" pageNumber="295">Comments.</paragraph>
<paragraph id="E65AC2D825845A490E8F58A12A1F075E" pageId="0" pageNumber="295">
The genus
<taxonomicName class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
<taxonomicName id="DAC5E541759F477C94718A09F8DA79E9" class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="520C5CD08526DBD435DF8759CC1F1252" italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
</taxonomicName>
differs from extant horse flies by the presence of two-segmented cercus and by the absence of the postspiracular scale, a one-segmented cercus and a postspiracular scale is shared by all extant
<taxonomicName authorityName="Latreille" authorityYear="1802" family="Tabanidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="295" rank="family">Tabanidae</taxonomicName>
<taxonomicName id="7278C72219B0511A34EDC9F56259AFD2" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Tabanidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="family">Tabanidae</taxonomicName>
(
<bibRefCitation DOI="https://www.doi.org/10.1071/ZO9540431" author="Mackerras, IM" journalOrPublisher="Australian Journal of Zoology" pageId="0" pageNumber="295" pagination="43 - 454" refId="B19" refString="Mackerras, IM, 1954. The classification and distribution of Tabanidae (Diptera). I. General Review. Australian Journal of Zoology 2: 43 - 454, DOI: https://www.doi.org/10.1071/ZO9540431" title="The classification and distribution of Tabanidae (Diptera). I. General Review." url="https://www.doi.org/10.1071/ZO9540431" volume="2" year="1954">Mackerras 1954</bibRefCitation>
<bibRefCitation id="33500EA44F1CED3778288E63E3080C88" DOI="https://www.doi.org/10.1071/ZO9540431" author="Mackerras, IM" journalOrPublisher="Australian Journal of Zoology" pageId="0" pageNumber="295" pagination="43 - 454" refId="B19" refString="Mackerras, IM, 1954. The classification and distribution of Tabanidae (Diptera). I. General Review. Australian Journal of Zoology 2: 43 - 454, DOI: https://www.doi.org/10.1071/ZO9540431" title="The classification and distribution of Tabanidae (Diptera). I. General Review." url="https://www.doi.org/10.1071/ZO9540431" volume="2" year="1954">Mackerras 1954</bibRefCitation>
).
<bibRefCitation author="Grimaldi, DA" journalOrPublisher="Bulletin of the American Museum of Natural History, 408, New York" pageId="0" pageNumber="295" refId="B12" refString="Grimaldi, DA, 2016. Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History, 408, New York" title="Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII." year="2016">Grimaldi (2016)</bibRefCitation>
<bibRefCitation id="D3CB10A9FEF0AB7150B4E3FCCA801FC7" author="Grimaldi, DA" journalOrPublisher="Bulletin of the American Museum of Natural History, 408, New York" pageId="0" pageNumber="295" refId="B12" refString="Grimaldi, DA, 2016. Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History, 408, New York" title="Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII." year="2016">Grimaldi (2016)</bibRefCitation>
also pointed out that the length of the discal cell in
<taxonomicName class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
<taxonomicName id="22DD5D6E07122648B31ADBBCAEAAE1E4" class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="5779982014968EF7E55CA1537F7BEAE5" italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
</taxonomicName>
is nearly the same as the length of the vein M1. However, this character is variable in the extant species. The genus can be attributed to
<taxonomicName authorityName="Latreille" authorityYear="1802" family="Tabanidae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="295" rank="family">Tabanidae</taxonomicName>
<taxonomicName id="2A9EFD13FB50327BF6C1E24F776F4B2E" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Tabanidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="family">Tabanidae</taxonomicName>
by the general disposition of the wing veins, the general shape of the body, the clypeus format and the presence of an enlarged calypter (
<bibRefCitation author="Grimaldi, DA" journalOrPublisher="Bulletin of the American Museum of Natural History, 408, New York" pageId="0" pageNumber="295" refId="B12" refString="Grimaldi, DA, 2016. Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History, 408, New York" title="Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII." year="2016">Grimaldi 2016</bibRefCitation>
<bibRefCitation id="2753263AAB58CBB865A3AFB54B1FCB73" author="Grimaldi, DA" journalOrPublisher="Bulletin of the American Museum of Natural History, 408, New York" pageId="0" pageNumber="295" refId="B12" refString="Grimaldi, DA, 2016. Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History, 408, New York" title="Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII." year="2016">Grimaldi 2016</bibRefCitation>
, and our Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cratotabanus stonemyomorphus. Martins-Neto and Santos. A, B Holotype female. A habitus preserved in ventral view. B Detail of head. C Male, preserved in dorsal view. Scale bars: 2 mm (A, C); 1 mm (B)." figureDoi="10.3897/asp.80.e86673.figure1" httpUri="https://binary.pensoft.net/fig/718026" pageId="0" pageNumber="295">1</figureCitation>
<figureCitation id="E128A321BCA4F282DE6305E810E51CF8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cratotabanus stonemyomorphus. Martins-Neto and Santos. A, B Holotype female. A habitus preserved in ventral view. B Detail of head. C Male, preserved in dorsal view. Scale bars: 2 mm (A, C); 1 mm (B)." figureDoi="10.3897/asp.80.e86673.figure1" httpUri="https://binary.pensoft.net/fig/718026" pageId="0" pageNumber="295">1</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cratotabanus stonemyomorphus. A Holotype female, illustration of dorsal habitus. B Male, illustration of ventral habitus. C Holotype female, wing. D Male, Wing. Scale bars: 2 mm (A, C); 1 mm (C, D)." figureDoi="10.3897/asp.80.e86673.figure2" httpUri="https://binary.pensoft.net/fig/718027" pageId="0" pageNumber="295">2</figureCitation>
<figureCitation id="1C8C7BD3563666AD23613847780574CB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Cratotabanus stonemyomorphus. A Holotype female, illustration of dorsal habitus. B Male, illustration of ventral habitus. C Holotype female, wing. D Male, Wing. Scale bars: 2 mm (A, C); 1 mm (C, D)." figureDoi="10.3897/asp.80.e86673.figure2" httpUri="https://binary.pensoft.net/fig/718027" pageId="0" pageNumber="295">2</figureCitation>
).
</paragraph>
<caption doi="10.3897/asp.80.e86673.figure1" httpUri="https://binary.pensoft.net/fig/718026" pageId="0" pageNumber="295" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="295">
<emphasis bold="true" pageId="0" pageNumber="295">Figure 1.</emphasis>
<taxonomicName class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus stonemyomorphus." order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="stonemyomorphus.">
<emphasis italics="true" pageId="0" pageNumber="295">Cratotabanus stonemyomorphus.</emphasis>
<caption id="DE75FF8FE5A4D3D2DAFDC271E5855AFB" doi="10.3897/asp.80.e86673.figure1" httpUri="https://binary.pensoft.net/fig/718026" pageId="0" pageNumber="295" start="Figure 1" startId="F1">
<paragraph id="573ADFD73A128D1BB4C467D384C9F703" pageId="0" pageNumber="295">
<emphasis id="F076174EBB7152573A078ACB04EB85A0" bold="true" pageId="0" pageNumber="295">Figure 1.</emphasis>
<taxonomicName id="E10667A7E65E5253174B8ABC99B2C1DD" class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus stonemyomorphus." order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="stonemyomorphus.">
<emphasis id="5F1AC464E9EAB26D8AB429CB49FFD24D" italics="true" pageId="0" pageNumber="295">Cratotabanus stonemyomorphus.</emphasis>
</taxonomicName>
Martins-Neto and Santos.
<emphasis bold="true" pageId="0" pageNumber="295">A</emphasis>
<emphasis id="ECA4CB488581D69BCDB3DFF388FB054D" bold="true" pageId="0" pageNumber="295">A</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="295">B</emphasis>
<emphasis id="3F8F22D2FA34040EC10512F43C9E53C5" bold="true" pageId="0" pageNumber="295">B</emphasis>
Holotype female.
<emphasis bold="true" pageId="0" pageNumber="295">A</emphasis>
<emphasis id="B4709571D3F436ACE25429FEB6D73E10" bold="true" pageId="0" pageNumber="295">A</emphasis>
habitus preserved in ventral view.
<emphasis bold="true" pageId="0" pageNumber="295">B</emphasis>
<emphasis id="2A02119016C4BEF974D279753F3226DB" bold="true" pageId="0" pageNumber="295">B</emphasis>
Detail of head.
<emphasis bold="true" pageId="0" pageNumber="295">C</emphasis>
<emphasis id="B677A68150D1E4A0EC755B235F846BF7" bold="true" pageId="0" pageNumber="295">C</emphasis>
Male, preserved in dorsal view. Scale bars: 2 mm (A, C); 1 mm (B).
</paragraph>
</caption>
<caption doi="10.3897/asp.80.e86673.figure2" httpUri="https://binary.pensoft.net/fig/718027" pageId="0" pageNumber="295" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="295">
<emphasis bold="true" pageId="0" pageNumber="295">Figure 2.</emphasis>
<taxonomicName class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus stonemyomorphus." order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="stonemyomorphus.">
<emphasis italics="true" pageId="0" pageNumber="295">Cratotabanus stonemyomorphus.</emphasis>
<caption id="0EAD6E6316DEFC01C14C875CE95F3F5E" doi="10.3897/asp.80.e86673.figure2" httpUri="https://binary.pensoft.net/fig/718027" pageId="0" pageNumber="295" start="Figure 2" startId="F2">
<paragraph id="629415F4BE436FDF2065AD6AF36D30FA" pageId="0" pageNumber="295">
<emphasis id="AD16CE8C3EF6F47D17410733D3C72C2B" bold="true" pageId="0" pageNumber="295">Figure 2.</emphasis>
<taxonomicName id="A29AFCCCBC5257B3AF745D825A2D56EF" class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus stonemyomorphus." order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="stonemyomorphus.">
<emphasis id="0FF7B710115E057801697A21E9260EB3" italics="true" pageId="0" pageNumber="295">Cratotabanus stonemyomorphus.</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="295">A</emphasis>
<emphasis id="52C11769D0538DBAC5A196370581AAD2" bold="true" pageId="0" pageNumber="295">A</emphasis>
Holotype female, illustration of dorsal habitus.
<emphasis bold="true" pageId="0" pageNumber="295">B</emphasis>
<emphasis id="07514DF671703D22FE53C26E866D322E" bold="true" pageId="0" pageNumber="295">B</emphasis>
Male, illustration of ventral habitus.
<emphasis bold="true" pageId="0" pageNumber="295">C Holotype female</emphasis>
<emphasis id="F167524C6858815CE4DFE2E619736C42" bold="true" pageId="0" pageNumber="295">C Holotype female</emphasis>
, wing.
<emphasis bold="true" pageId="0" pageNumber="295">D</emphasis>
<emphasis id="72704D77BF816268563B34E27749FBCD" bold="true" pageId="0" pageNumber="295">D</emphasis>
Male, Wing. Scale bars: 2 mm (A, C); 1 mm (C, D).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="295">
<taxonomicName class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
<paragraph id="3BCFCEF1616DDEA45E0E34431C5C5A35" pageId="0" pageNumber="295">
<taxonomicName id="575E702B4660E560B6EE825941D032C4" class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="66F2A3A5012299B126BC11857B8DD145" italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Zhang" authorityYear="2012" class="Insecta" family="Tabanidae" genus="Laiyangitabanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Laiyangitabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="295">Laiyangitabanus</emphasis>
<taxonomicName id="DEA4095A699D0422288345A23010D4AF" authorityName="Zhang" authorityYear="2012" class="Insecta" family="Tabanidae" genus="Laiyangitabanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Laiyangitabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="81934A774AF39421F1C54AD952B557F9" italics="true" pageId="0" pageNumber="295">Laiyangitabanus</emphasis>
</taxonomicName>
Zhang and
<taxonomicName class="Insecta" family="Tabanidae" genus="Eotabanoid" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eotabanoid" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="295">Eotabanoid</emphasis>
<taxonomicName id="232299AE8EA37FDA893D9D5AB0D16220" class="Insecta" family="Tabanidae" genus="Eotabanoid" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eotabanoid" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="E4C865A50F63701DAA2DFD253FF1240F" italics="true" pageId="0" pageNumber="295">Eotabanoid</emphasis>
</taxonomicName>
Mostovski et al. by the presence of vein R2+3 curved only at the apex. From
<taxonomicName authorityName="Zhang" authorityYear="2012" class="Insecta" family="Tabanidae" genus="Laiyangitabanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Laiyangitabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="295">Laiyangitabanus</emphasis>
<taxonomicName id="3FF241E9850C412DE1D31710EF9CDA02" authorityName="Zhang" authorityYear="2012" class="Insecta" family="Tabanidae" genus="Laiyangitabanus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Laiyangitabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="D9ACB200E59F3998BAE386988ED113C7" italics="true" pageId="0" pageNumber="295">Laiyangitabanus</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
<taxonomicName id="23F3F7F425E7E3E66B59B33AE8BE2A53" class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="2EC57ECFC3424A89C01AB2FB87F12E90" italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
</taxonomicName>
further differs by the presence of a well-developed ocellar triangle, and r-m closer to the base of the discal cell (
<bibRefCitation author="Grimaldi, DA" journalOrPublisher="Bulletin of the American Museum of Natural History, 408, New York" pageId="0" pageNumber="295" refId="B12" refString="Grimaldi, DA, 2016. Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History, 408, New York" title="Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII." year="2016">Grimaldi 2016</bibRefCitation>
<bibRefCitation id="CAD0F4487189CA7986E0B93AF14DB711" author="Grimaldi, DA" journalOrPublisher="Bulletin of the American Museum of Natural History, 408, New York" pageId="0" pageNumber="295" refId="B12" refString="Grimaldi, DA, 2016. Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History, 408, New York" title="Diverse Orthorrhapham flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII." year="2016">Grimaldi 2016</bibRefCitation>
).
<taxonomicName class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
<taxonomicName id="6991000CC764D65F9C31B910E17F280B" class="Insecta" family="Tabanidae" genus="Cratotabanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratotabanus" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="BD961DAFD9F02BC0D2AE4D9DF5ED64DD" italics="true" pageId="0" pageNumber="295">Cratotabanus</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Ren" authorityYear="1998" class="Insecta" family="Tabanidae" genus="Eopangonius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eopangonius" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="295">Eopangonius</emphasis>
<taxonomicName id="30FFC94A03D1EAEBD03061C5BA2EC9FF" authorityName="Ren" authorityYear="1998" class="Insecta" family="Tabanidae" genus="Eopangonius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eopangonius" order="Diptera" pageId="0" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="20623CCDDC1DE2A83C466347D94C1154" italics="true" pageId="0" pageNumber="295">Eopangonius</emphasis>
</taxonomicName>
Ren by the absence of basal callus and a not projected clypeus.
</paragraph>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1117.86984" ID-Pensoft-Pub="1313-2970-1117-95" ID-Pensoft-UUID="54DB7D7318935384856B59FD7D22DFF4" ID-ZooBank="C583CC7AB2AD42048FA083C49BB088EA" ModsDocID="1313-2970-1117-95" checkinTime="1660292277777" checkinUser="pensoft" docAuthor="Holzenthal, Ralph W., Blahnik, Roger J. &amp; Rios-Touma, Blanca" docDate="2022" docId="A162FCCECD7B55ECA58B05239A27A2DC" docLanguage="en" docName="ZooKeys 1117: 95-122" docOrigin="ZooKeys 1117" docPubDate="2022-08-11" docSource="http://dx.doi.org/10.3897/zookeys.1117.86984" docTitle="Wormaldia natalis Holzenthal &amp; Blahnik &amp; Ríos-Touma 2022, sp. nov." docType="treatment" docUuid="84769896-75D5-4104-ABAF-24B17AF6CDAC" docUuidSource="ZooBank" docVersion="1" id="54DB7D7318935384856B59FD7D22DFF4" lastPageNumber="95" masterDocId="54DB7D7318935384856B59FD7D22DFF4" masterDocTitle="A new genus and new species of Ecuadorian Philopotamidae (Trichoptera)" masterLastPageNumber="122" masterPageNumber="95" pageNumber="95" updateTime="1660292277777" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title>A new genus and new species of Ecuadorian Philopotamidae (Trichoptera)</mods:title>
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<mods:title id="2B1A71AEC7EA797B20626FCCBCD91376">A new genus and new species of Ecuadorian Philopotamidae (Trichoptera)</mods:title>
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<figureCitation id="A24DC4F01392FF54AFA1A68BE8AEE089" captionStart="Figure 14" captionStartId="F12" captionText="Figure 14. Wormaldia natalis sp. nov. Male genitalia A segments VIII-X, lateral B segments VIII-X, dorsal C segments VIII, IX, inferior appendage, ventral D phallus, lateral, detail: endothecal spines." figureDoi="10.3897/zookeys.1117.86984.figure14" httpUri="https://binary.pensoft.net/fig/726441" pageId="0" pageNumber="95">Fig. 14</figureCitation>
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<paragraph pageId="0" pageNumber="95">Type material.</paragraph>
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<emphasis italics="true" pageId="0" pageNumber="95">Holotype</emphasis>
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<paragraph id="AF9D3F5A8DE52AD5E2D85CBD28E8B7D3" pageId="0" pageNumber="95">Type material.</paragraph>
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Male (pinned). Ecuador: Napo: Wildsumaco Lodge, small stream, Coati Trail @ wooden bridge,
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, 1420 m a.s.l., 10.iii.2020,
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, Holzenthal, Frandsen, Pauls, Amigo, UMSP000500642 (UMSP).
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Ecuador: same data as holotype, 1 male (pinned) (MECN).
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<paragraph pageId="0" pageNumber="95">Diagnosis.</paragraph>
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<paragraph id="FEE620E2116C0B7BDE2128BC487E8102" pageId="0" pageNumber="95">Diagnosis.</paragraph>
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is undoubtedly most closely related to
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&amp; Holzenthal, 2015, described from Bolivia and resembling it in several respects, but particularly in the short, broad posteromesal projections from the posterior margin of tergum VIII (Fig.
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) and the strongly tapering apical segment of the inferior appendage (Fig.
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). However, it differs in the shorter, more basally inflated preanal appendages (Fig.
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), less angular projection of segment IX below the inferior appendages (Fig.
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), details of the apex and shape of the lateral projections of tergum X (Fig.
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), development of the apical segment of the inferior appendages, and in having a longer phallic spine (Fig.
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<figureCitation id="747C0384F1252C9AE134BAEAE2A74F62" captionStart="Figure 14" captionStartId="F12" captionText="Figure 14. Wormaldia natalis sp. nov. Male genitalia A segments VIII-X, lateral B segments VIII-X, dorsal C segments VIII, IX, inferior appendage, ventral D phallus, lateral, detail: endothecal spines." figureDoi="10.3897/zookeys.1117.86984.figure14" httpUri="https://binary.pensoft.net/fig/726441" pageId="0" pageNumber="95">14D</figureCitation>
). Additionally, the two short projections from the posterior margin of tergum VIII have a more V-shaped than U-shaped basal separation (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F12" captionText="Figure 14. Wormaldia natalis sp. nov. Male genitalia A segments VIII-X, lateral B segments VIII-X, dorsal C segments VIII, IX, inferior appendage, ventral D phallus, lateral, detail: endothecal spines." figureDoi="10.3897/zookeys.1117.86984.figure14" httpUri="https://binary.pensoft.net/fig/726441" pageId="0" pageNumber="95">14B</figureCitation>
<figureCitation id="6DD7F2E60EE842070358F73287125B09" captionStart="Figure 14" captionStartId="F12" captionText="Figure 14. Wormaldia natalis sp. nov. Male genitalia A segments VIII-X, lateral B segments VIII-X, dorsal C segments VIII, IX, inferior appendage, ventral D phallus, lateral, detail: endothecal spines." figureDoi="10.3897/zookeys.1117.86984.figure14" httpUri="https://binary.pensoft.net/fig/726441" pageId="0" pageNumber="95">14B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1117.86984.figure14" httpUri="https://binary.pensoft.net/fig/726441" pageId="0" pageNumber="95" start="Figure 14" startId="F12">
<paragraph pageId="0" pageNumber="95">
<emphasis bold="true" pageId="0" pageNumber="95">Figure 14.</emphasis>
<taxonomicName authorityName="Holzenthal &amp; Blahnik &amp; Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia natalis" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="natalis">
<emphasis italics="true" pageId="0" pageNumber="95">Wormaldia natalis</emphasis>
<caption id="CDD69721FFF4BB3715D618230542D3A3" doi="10.3897/zookeys.1117.86984.figure14" httpUri="https://binary.pensoft.net/fig/726441" pageId="0" pageNumber="95" start="Figure 14" startId="F12">
<paragraph id="2FE48A973A3D09399ED218524C4EC903" pageId="0" pageNumber="95">
<emphasis id="432AF9CEDF330310834D19B355B74E8B" bold="true" pageId="0" pageNumber="95">Figure 14.</emphasis>
<taxonomicName id="BD913209E20B59F8F4806C866BAC604F" authorityName="Holzenthal &amp; Blahnik &amp; Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia natalis" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="natalis">
<emphasis id="A0AFB5B30D6DB719A94439C6845C3DC6" italics="true" pageId="0" pageNumber="95">Wormaldia natalis</emphasis>
</taxonomicName>
sp. nov. Male genitalia
<emphasis bold="true" pageId="0" pageNumber="95">A</emphasis>
<emphasis id="3189D1AFDEA330835432DBC57C413B39" bold="true" pageId="0" pageNumber="95">A</emphasis>
segments VIII-X, lateral
<emphasis bold="true" pageId="0" pageNumber="95">B</emphasis>
<emphasis id="B9C3BEF367E0D13B5CC41C700A19B612" bold="true" pageId="0" pageNumber="95">B</emphasis>
segments VIII-X, dorsal
<emphasis bold="true" pageId="0" pageNumber="95">C</emphasis>
<emphasis id="F1B8D7D55FC26B0E7B71825825E1338A" bold="true" pageId="0" pageNumber="95">C</emphasis>
segments VIII, IX, inferior appendage, ventral
<emphasis bold="true" pageId="0" pageNumber="95">D</emphasis>
<emphasis id="30F10B2B963ED73EDDE99DE4BBD9BD1A" bold="true" pageId="0" pageNumber="95">D</emphasis>
phallus, lateral, detail: endothecal spines.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="95" type="description">
<paragraph pageId="0" pageNumber="95">Description.</paragraph>
<paragraph pageId="0" pageNumber="95">
<emphasis bold="true" pageId="0" pageNumber="95">Adult.</emphasis>
<subSubSection id="A800FF40627A7D6C79162A2DE6DFFA33" pageId="0" pageNumber="95" type="description">
<paragraph id="062DE2E4FC91D1CAD36216D644AA5795" pageId="0" pageNumber="95">Description.</paragraph>
<paragraph id="C41C4DA61CEA7981D340FC76D1D63D20" pageId="0" pageNumber="95">
<emphasis id="D60D00FEB019052C8DD3E020CAD325A1" bold="true" pageId="0" pageNumber="95">Adult.</emphasis>
Forewing length male 5.2 mm (
<emphasis italics="true" pageId="0" pageNumber="95">n</emphasis>
<emphasis id="6CCC0B6E8C919285F9BE8D93D117A14F" italics="true" pageId="0" pageNumber="95">n</emphasis>
= 1). Head brown, with yellowish setae. Antenna with overall color yellowish, indistinctly annulated with light brown, somewhat flattened setae. Palps very short, with dark brown setae. Dorsum of thorax brown. Legs medium brown, spurs slightly darker; hind tibiae with sparse brush of long setae. Forewing covered with dark brown setae; apical forks I, II, III, IV, and V present. Hind wing translucent, with very fine, small, brown setae; apical forks I, II, III, and V present.
</paragraph>
<paragraph pageId="0" pageNumber="95">
<emphasis bold="true" pageId="0" pageNumber="95">Male.</emphasis>
<paragraph id="985825E3BDADEF921A6E5B0F5E47E8F3" pageId="0" pageNumber="95">
<emphasis id="13D422E725C30AF7B079DCF76128ACFD" bold="true" pageId="0" pageNumber="95">Male.</emphasis>
Segment VIII slightly shorter than segment IX, tergum with pair of short, diverging, subtruncate posteromesal projections. Segment IX, as viewed laterally, with anterior margin weakly angularly produced at ca. mid-height, posterior margin nearly linear, angularly narrowing just below preanal appendage, dorsal margin, as viewed dorsally, very short, with short projecting anterolateral apodemes. Segment X, in lateral view, elongate narrow, somewhat down-curved, apex with short, angular apicodorsal projection, sensilla confined to apex; as viewed dorsally, elongate, tapering, subtriangular, apex subacute, with scattered sensilla, apex continuous laterally with short, diverging, rounded projections, each with several sensilla. Preanal appendage elongate, irregular in shape, with distinct basodorsal expansion, narrowing apically. Inferior appendage bi-segmented, both segments tapering from base to apex, basal segment very wide basally, ca. half height of segment IX, apical segment with apex subacute and slightly down-turned; as viewed ventrally, with basal segment bulbous, ~ 2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="FFCC73ABAC4450C4E437E1F0E22B0A20" originalValue="×">x</normalizedToken>
width of apical segment, apical segment with dense apicomesal pad of short spines. Phallus, when viewed laterally, with phallobase very short, with basodorsal expansion, weakly sclerotized, tapering apically, endotheca with two spines, one short and strongly curved, the other elongate, narrow, more sinuously curved.
</paragraph>
<paragraph pageId="0" pageNumber="95">
<emphasis bold="true" pageId="0" pageNumber="95">Female.</emphasis>
<paragraph id="3E67EE6AEF5D9645BBDAEA9FBA7938EE" pageId="0" pageNumber="95">
<emphasis id="48FBEC2783BCC05C25205902322ED8EA" bold="true" pageId="0" pageNumber="95">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="95" type="etymology">
<paragraph pageId="0" pageNumber="95">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="95">
<subSubSection id="ED1079FA833CD73CB2E5FB33678FAC51" pageId="0" pageNumber="95" type="etymology">
<paragraph id="9518FAA78A5A71E00918CBCD3138B5B0" pageId="0" pageNumber="95">Etymology.</paragraph>
<paragraph id="BB0A14DD644A7BFBCFF59DB8DC080574" pageId="0" pageNumber="95">
From the Latin
<taxonomicName authorityName="Holzenthal &amp; Blahnik &amp; Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia natalis" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="natalis">
<emphasis italics="true" pageId="0" pageNumber="95">Wormaldia natalis</emphasis>
<taxonomicName id="1CDD5BC2B6840ACC863D7B427FB42E1B" authorityName="Holzenthal &amp; Blahnik &amp; Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia natalis" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="natalis">
<emphasis id="661CC18CAA7E1EB948B155EEBA6E5226" italics="true" pageId="0" pageNumber="95">Wormaldia natalis</emphasis>
</taxonomicName>
, meaning birthday in reference to the date when the species was collected, March 10th, the shared birthday of collectors Frandsen and Holzenthal.
</paragraph>

View file

@ -1,162 +1,162 @@
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Genus
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<bibRefCitation author="Haliday, AH" journalOrPublisher="Entomological Magazine" pageId="8" pageNumber="57" pagination="458 - 468" title="Essay on parasitic Hymenoptera of the Ichneumones Adsciti." volume="2" year="1835">Haliday 1835</bibRefCitation>
<subSubSection id="8AA7448B4CA6E6DBE58A50AD849AB6AA" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50" type="reference_group">
<paragraph id="449D80FEA1919FBB9996491062CD4442" pageId="1" pageNumber="50">
<taxonomicName id="D172AD1096BCA31FB5E0FD17A75CC8F6" genus="Ismarus" lsidName="Ismarus" pageId="1" pageNumber="50" rank="genus">Ismarus</taxonomicName>
<bibRefCitation id="17BBE7A5648142898BF923C041F7DE63" author="Haliday, AH" journalOrPublisher="Entomological Magazine" pageId="8" pageNumber="57" pagination="458 - 468" title="Essay on parasitic Hymenoptera of the Ichneumones Adsciti." volume="2" year="1835">Haliday 1835</bibRefCitation>
, Entomological Magazine 2: 467. Type:
<taxonomicName class="Insecta" family="Diapriidae" genus="Cinetus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cinetus dorsiger" order="Hymenoptera" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="dorsiger">Cinetus dorsiger</taxonomicName>
<taxonomicName id="07AFC86236CC3534D32BA4450A574176" genus="Cinetus" lsidName="Cinetus dorsiger" pageId="1" pageNumber="50" rank="species" species="dorsiger">Cinetus dorsiger</taxonomicName>
Haliday, by monotypy.
</paragraph>
<paragraph pageId="1" pageNumber="50">
<taxonomicName genus="Entomia" lsidName="Entomia" pageId="1" pageNumber="50" rank="genus">Entomia</taxonomicName>
<normalizedToken originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
<paragraph id="9F338E3D22B1083602E3B394D7C6F618" pageId="1" pageNumber="50">
<taxonomicName id="4BDA5F2EE52E1CF755879167D464EFAF" genus="Entomia" lsidName="Entomia" pageId="1" pageNumber="50" rank="genus">Entomia</taxonomicName>
<normalizedToken id="7B3263FD4D4B77CA60EC3B09DFA98D81" originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
1840, Nomenclator Entomologicus 2: 127. Type:
<taxonomicName genus="Entomia" lsidName="Entomia campanulata" pageId="1" pageNumber="50" rank="species" species="campanulata">Entomia campanulata</taxonomicName>
<normalizedToken originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
<taxonomicName id="0FDE05E7202A72FB78CAF1EA61161B71" genus="Entomia" lsidName="Entomia campanulata" pageId="1" pageNumber="50" rank="species" species="campanulata">Entomia campanulata</taxonomicName>
<normalizedToken id="71E4870222DDA7C09BB03C9A709ECD6E" originalValue="Herrich-Schäffer">Herrich-Schaeffer</normalizedToken>
, 1840, by monotypy.
</paragraph>
<paragraph pageId="1" pageNumber="50">
<taxonomicName genus="Entomius" lsidName="Entomius" pageId="1" pageNumber="50" rank="genus">Entomius</taxonomicName>
<bibRefCitation author="Haliday, AH" journalOrPublisher=")" pageId="8" pageNumber="57" pagination="166 - 174" title="Notes on the peculiar form of the ovaries observed in a hymenopterous insect, constituting a new genus and species of the family Diapriidae. Natural History Review (Proc." volume="4" year="1857">Haliday 1857</bibRefCitation>
<paragraph id="0F09DAAA0A9FBA03306D60F91F30E993" pageId="1" pageNumber="50">
<taxonomicName id="688FC42CC901697F70792BECF4D4B23D" genus="Entomius" lsidName="Entomius" pageId="1" pageNumber="50" rank="genus">Entomius</taxonomicName>
<bibRefCitation id="82E17D8FE2CFDA54B3E69F0121009361" author="Haliday, AH" journalOrPublisher=")" pageId="8" pageNumber="57" pagination="166 - 174" title="Notes on the peculiar form of the ovaries observed in a hymenopterous insect, constituting a new genus and species of the family Diapriidae. Natural History Review (Proc." volume="4" year="1857">Haliday 1857</bibRefCitation>
, Nature History Review 4: 169. Emendation.
</paragraph>
<paragraph pageId="2" pageNumber="51">
<taxonomicName class="Insecta" family="Diapriidae" genus="Agonophorus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Agonophorus" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="2" pageNumber="51" start="start">Agonophorus</pageBreakToken>
<paragraph id="2E738E738A619FF594BEA34CB452A460" pageId="2" pageNumber="51">
<taxonomicName id="4476A212ABE69DD7EAAE0A4903819C7D" genus="Agonophorus" lsidName="Agonophorus" pageId="2" pageNumber="51" rank="genus">
<pageBreakToken id="A6832A7EF8959630F51013FAE8894932" pageId="2" pageNumber="51" start="start">Agonophorus</pageBreakToken>
</taxonomicName>
Dahlbom 1858,
<normalizedToken originalValue="Öfversigt">Oefversigt</normalizedToken>
<normalizedToken id="13923E42E58CC85D513D2487C373CBD0" originalValue="Öfversigt">Oefversigt</normalizedToken>
af Kongliga Ventenskaps-Akadamiens
<normalizedToken originalValue="Förhandlingar">Foerhandlingar</normalizedToken>
<normalizedToken id="5CFD8EC7F46374343818B62EBA453F98" originalValue="Förhandlingar">Foerhandlingar</normalizedToken>
14: 289. Synonymized by
<bibRefCitation author="Dessart, P" journalOrPublisher="Records of South Australian Museum" pageId="8" pageNumber="57" pagination="551 - 559" title="Corrections in the systematic position of certain taxa of the Chalcidoidea and Proctotrupoidea (s. l.), with notes on synonymy." volume="15" year="1967">Dessart (1967)</bibRefCitation>
<bibRefCitation id="C43D7CE063142F0D5564FC99E78E7148" author="Dessart, P" journalOrPublisher="Records of South Australian Museum" pageId="8" pageNumber="57" pagination="551 - 559" title="Corrections in the systematic position of certain taxa of the Chalcidoidea and Proctotrupoidea (s. l.), with notes on synonymy." volume="15" year="1967">Dessart (1967)</bibRefCitation>
. Type:
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus rugulosus" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="rugulosus">Ismarus rugulosus</taxonomicName>
<taxonomicName id="0BE9D3E2C212AAC1DF3F9EE38C6A01D5" genus="Ismarus" lsidName="Ismarus rugulosus" pageId="2" pageNumber="51" rank="species" species="rugulosus">Ismarus rugulosus</taxonomicName>
Foerster, designated by
<bibRefCitation author="Muesebeck, CFW" journalOrPublisher="Proceedings of Entomological Society of Washington" pageId="8" pageNumber="57" title="On the identity of Agonophorus Dahlbom (Hymenoptera: Diapriidae)." volume="74" year="1972">Muesebeck (1972)</bibRefCitation>
<bibRefCitation id="EAF2E5168E16D0A082AF77EF95FCA1FC" author="Muesebeck, CFW" journalOrPublisher="Proceedings of Entomological Society of Washington" pageId="8" pageNumber="57" title="On the identity of Agonophorus Dahlbom (Hymenoptera: Diapriidae)." volume="74" year="1972">Muesebeck (1972)</bibRefCitation>
.
</paragraph>
<paragraph pageId="2" pageNumber="51">
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">Ismarus</taxonomicName>
<paragraph id="436BF683EF81C311D61599A5C4200A91" pageId="2" pageNumber="51">
<taxonomicName id="7B81A13B91C3BFFC046D58BB5D63BE86" genus="Ismarus" lsidName="Ismarus" pageId="2" pageNumber="51" rank="genus">Ismarus</taxonomicName>
Haliday, 1835:
<bibRefCitation author="Masner, L" journalOrPublisher="Canadian Entomologist" pageId="8" pageNumber="57" pagination="1243 - 1266" title="A revision of the Ismarinae of the New World (Hymenoptera, Proctotrupoidea, Diapriidae)." url="doi:10.4039/Ent1081243-11" volume="108" year="1976">Masner 1976</bibRefCitation>
<bibRefCitation id="8BD9AE0CF0C8087A250BC34FC163012F" author="Masner, L" journalOrPublisher="Canadian Entomologist" pageId="8" pageNumber="57" pagination="1243 - 1266" title="A revision of the Ismarinae of the New World (Hymenoptera, Proctotrupoidea, Diapriidae)." url="doi:10.4039/Ent1081243-11" volume="108" year="1976">Masner 1976</bibRefCitation>
, Canadian Entomologist 108: 1251.
</paragraph>
<paragraph pageId="2" pageNumber="51">
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">Ismarus</taxonomicName>
<paragraph id="B1F3B93D15956CEE41D62D179CEA275F" pageId="2" pageNumber="51">
<taxonomicName id="AB708A6AF7183A6090E85E1894BBA050" genus="Ismarus" lsidName="Ismarus" pageId="2" pageNumber="51" rank="genus">Ismarus</taxonomicName>
Haliday, 1835:
<bibRefCitation author="Johnson, NF" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="8" pageNumber="57" pagination="1 - 825" title="Catalog of Worlds Species of Proctotrupoidea, Exclusive of Platygastridae (Hymenoptera)." volume="51" year="1992">Johnson 1992</bibRefCitation>
<bibRefCitation id="9F574D67F4328639A6AE31047C0E5367" author="Johnson, NF" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="8" pageNumber="57" pagination="1 - 825" title="Catalog of Worlds Species of Proctotrupoidea, Exclusive of Platygastridae (Hymenoptera)." volume="51" year="1992">Johnson 1992</bibRefCitation>
, Memoirs of the American Entomological Institute 51: 259.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="51" type="description">
<paragraph pageId="2" pageNumber="51">Description.</paragraph>
<paragraph pageId="2" pageNumber="51">Body stout; body colour usually black, but brown in a few species. Head transverse in dorsal view, with sparse setae on face, frons and occiput; labrum exposed and sclerotized; mandible bidentate; palpal formula 4-3 or 5-3; antennae inserted low on face, close to clypeus; face with a distinct transverse carina below antennal sockets; antennal shelf not prominent; antenna of female 15-segmented, of male 14-segmented; modified male sex-segment is second flagellomere, rarely both first and second flagellomeres modified; eyes bare; occipital carina complete; pronotum dorsally and along anterior margin with long setae; mesoscutum convex, smooth; notauli reduced to anterior pits; humeral sulcus developed; anterior scutellar pit with a weak but distinct median longitudinal carina; scutellum posteriorly raised, with posterior margin truncate or round; metanotum crenulated, with median carina distinct; propodeum with distinct transverse and longitudinal carinae; metapleuron reticulate rugose, with dense setosity; fore wing with radial cell closed; fore tibia with regular spur and one false spur, hind tibia strongly incrassate; petiole short and transverse; base of second tergite with longitudinal furrows; sutures between tergites distinct or absent; sternite with fine setae.</paragraph>
<subSubSection id="1A4919451BEF154CA574AE196BCC2FC8" pageId="2" pageNumber="51" type="description">
<paragraph id="C2AB725E38F75B34AF6AEFA24F6E0ABF" pageId="2" pageNumber="51">Description.</paragraph>
<paragraph id="A9C3B47DD52C4F08201C5D06BCA129FB" pageId="2" pageNumber="51">Body stout; body colour usually black, but brown in a few species. Head transverse in dorsal view, with sparse setae on face, frons and occiput; labrum exposed and sclerotized; mandible bidentate; palpal formula 4-3 or 5-3; antennae inserted low on face, close to clypeus; face with a distinct transverse carina below antennal sockets; antennal shelf not prominent; antenna of female 15-segmented, of male 14-segmented; modified male sex-segment is second flagellomere, rarely both first and second flagellomeres modified; eyes bare; occipital carina complete; pronotum dorsally and along anterior margin with long setae; mesoscutum convex, smooth; notauli reduced to anterior pits; humeral sulcus developed; anterior scutellar pit with a weak but distinct median longitudinal carina; scutellum posteriorly raised, with posterior margin truncate or round; metanotum crenulated, with median carina distinct; propodeum with distinct transverse and longitudinal carinae; metapleuron reticulate rugose, with dense setosity; fore wing with radial cell closed; fore tibia with regular spur and one false spur, hind tibia strongly incrassate; petiole short and transverse; base of second tergite with longitudinal furrows; sutures between tergites distinct or absent; sternite with fine setae.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="51" type="biology">
<paragraph pageId="2" pageNumber="51">Biology.</paragraph>
<paragraph pageId="2" pageNumber="51">
<subSubSection id="5EC89219AA841260525B096B2A9316A2" pageId="2" pageNumber="51" type="biology">
<paragraph id="28EA15A03611A7B4F79F42ECDDF79632" pageId="2" pageNumber="51">Biology.</paragraph>
<paragraph id="0D49D1DC6241350645D89F691EBEE63F" pageId="2" pageNumber="51">
Some species of the genus are hyperparasitoids of Dryinidae (Hymenoptera).
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus flavicornis" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="flavicornis">Ismarus flavicornis</taxonomicName>
<taxonomicName id="85E32E677D508FE780B613E58C8497F9" genus="Ismarus" lsidName="Ismarus flavicornis" pageId="2" pageNumber="51" rank="species" species="flavicornis">Ismarus flavicornis</taxonomicName>
(Thomson, 1859) was reared from
<taxonomicName class="Insecta" family="Dryinidae" genus="Anteon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anteon flavicorne" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="flavicorne">Anteon flavicorne</taxonomicName>
<taxonomicName id="0E77FD33F0ED0FCEA367E378A6AEA894" genus="Anteon" lsidName="Anteon flavicorne" pageId="2" pageNumber="51" rank="species" species="flavicorne">Anteon flavicorne</taxonomicName>
(Dalman, 1818),
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus halidayi" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="halidayi">Ismarus halidayi</taxonomicName>
<taxonomicName id="EC57C62C7016E3C12FC272D9DD42B378" genus="Ismarus" lsidName="Ismarus halidayi" pageId="2" pageNumber="51" rank="species" species="halidayi">Ismarus halidayi</taxonomicName>
was reared from an
<taxonomicName class="Insecta" family="Dryinidae" genus="Anteon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anteon" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">Anteon</taxonomicName>
<taxonomicName id="D63575E89025685967EC4D78BF6B5B4A" genus="Anteon" lsidName="Anteon" pageId="2" pageNumber="51" rank="genus">Anteon</taxonomicName>
sp. and
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus dorsiger" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="dorsiger">Ismarus dorsiger</taxonomicName>
<taxonomicName id="60DB5C6B02133B0D7771E9311CBA46CB" genus="Ismarus" lsidName="Ismarus dorsiger" pageId="2" pageNumber="51" rank="species" species="dorsiger">Ismarus dorsiger</taxonomicName>
was recorded to attack an
<taxonomicName class="Insecta" family="Dryinidae" genus="Aphelopus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aphelopus" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">Aphelopus</taxonomicName>
<taxonomicName id="55E3A903E0EF07DF50CCF58C2E124E88" genus="Aphelopus" lsidName="Aphelopus" pageId="2" pageNumber="51" rank="genus">Aphelopus</taxonomicName>
sp. (
<bibRefCitation author="Chambers, VH" journalOrPublisher="Entomologist's Monthly Magazine" pageId="8" pageNumber="57" pagination="114 - 115" title="Some hosts of Anteon spp. (Hym. Dryinidae) and a hyperparasite Ismarus (Hymenoptera. Belytidae)." volume="91" year="1955">Chambers 1955</bibRefCitation>
<bibRefCitation id="5959D035CA874E53446A43971B4EB85E" author="Chambers, VH" journalOrPublisher="Entomologist's Monthly Magazine" pageId="8" pageNumber="57" pagination="114 - 115" title="Some hosts of Anteon spp. (Hym. Dryinidae) and a hyperparasite Ismarus (Hymenoptera. Belytidae)." volume="91" year="1955">Chambers 1955</bibRefCitation>
;
<bibRefCitation author="Chambers, VH" journalOrPublisher="Entomologist's Monthly Magazine" pageId="8" pageNumber="57" title="A host for Ismarus halidayi Foerst (Hym, Diapriidae)." volume="117" year="1981">1981</bibRefCitation>
<bibRefCitation id="835288BFB7459AC3961B9351729C9DD5" author="Chambers, VH" journalOrPublisher="Entomologist's Monthly Magazine" pageId="8" pageNumber="57" title="A host for Ismarus halidayi Foerst (Hym, Diapriidae)." volume="117" year="1981">1981</bibRefCitation>
;
<bibRefCitation author="Nixon, GEL" journalOrPublisher="Handbooks for the Identification of British Insects" pageId="8" pageNumber="57" pagination="1 - 107" title="Hymenoptera, Proctotrupoidea, Diapriidae subfamily Belytinae." volume="8" year="1957">Nixon 1957</bibRefCitation>
<bibRefCitation id="D27E4CAEC647F0FA95F9D3AB6682DA48" author="Nixon, GEL" journalOrPublisher="Handbooks for the Identification of British Insects" pageId="8" pageNumber="57" pagination="1 - 107" title="Hymenoptera, Proctotrupoidea, Diapriidae subfamily Belytinae." volume="8" year="1957">Nixon 1957</bibRefCitation>
;
<bibRefCitation author="Jervis, MA" journalOrPublisher="Entomologist's Gazette" pageId="8" pageNumber="57" pagination="127 - 129" title="Parasitism of Aphelopus species (Hymenoptera: Dryinidae) by Ismarus dorsiger (Curtis) (Hymenoptera: Diapriidae)." volume="30" year="1979">Jervis 1979</bibRefCitation>
<bibRefCitation id="9CA58B87729D653201C4A21BE7A91DED" author="Jervis, MA" journalOrPublisher="Entomologist's Gazette" pageId="8" pageNumber="57" pagination="127 - 129" title="Parasitism of Aphelopus species (Hymenoptera: Dryinidae) by Ismarus dorsiger (Curtis) (Hymenoptera: Diapriidae)." volume="30" year="1979">Jervis 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="51" type="distribution">
<paragraph pageId="2" pageNumber="51">Distribution.</paragraph>
<paragraph pageId="2" pageNumber="51">Palaearctic, Oriental, Ethiopian, Nearctic, Neotropical and Australian regions.</paragraph>
<subSubSection id="6220FA85E6CC205BB3B416C2EF4702A2" pageId="2" pageNumber="51" type="distribution">
<paragraph id="A941AFED63744CBBE3B82A2B8F4D3F9E" pageId="2" pageNumber="51">Distribution.</paragraph>
<paragraph id="C6ACF670EE3AA7E0802D419A9D4258E4" pageId="2" pageNumber="51">Palaearctic, Oriental, Ethiopian, Nearctic, Neotropical and Australian regions.</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="51" type="discussion">
<paragraph pageId="2" pageNumber="51">Discussion.</paragraph>
<paragraph pageId="2" pageNumber="51">
<bibRefCitation author="Masner, L" journalOrPublisher="Canadian Entomologist" pageId="8" pageNumber="57" pagination="1243 - 1266" title="A revision of the Ismarinae of the New World (Hymenoptera, Proctotrupoidea, Diapriidae)." url="doi:10.4039/Ent1081243-11" volume="108" year="1976">Masner (1976)</bibRefCitation>
<subSubSection id="B7B205A7BDD8B6E8630FAEEE0CD5679F" pageId="2" pageNumber="51" type="discussion">
<paragraph id="738C0CA3186CDA1B6893311B4F1CD83C" pageId="2" pageNumber="51">Discussion.</paragraph>
<paragraph id="589B3B8998F2BCDAAC3BC2334B943B6C" pageId="2" pageNumber="51">
<bibRefCitation id="9A92B94E10FBDEE271614009C7C786EA" author="Masner, L" journalOrPublisher="Canadian Entomologist" pageId="8" pageNumber="57" pagination="1243 - 1266" title="A revision of the Ismarinae of the New World (Hymenoptera, Proctotrupoidea, Diapriidae)." url="doi:10.4039/Ent1081243-11" volume="108" year="1976">Masner (1976)</bibRefCitation>
divided
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">Ismarus</taxonomicName>
<taxonomicName id="D69AB27B7B9033E27C2EBC6552AE6321" genus="Ismarus" lsidName="Ismarus" pageId="2" pageNumber="51" rank="genus">Ismarus</taxonomicName>
into four species groups:
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus rugulosus" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="rugulosus">Ismarus rugulosus</taxonomicName>
<taxonomicName id="3264EE12B64C1668E00A81C808C8629B" genus="Ismarus" lsidName="Ismarus rugulosus" pageId="2" pageNumber="51" rank="species" species="rugulosus">Ismarus rugulosus</taxonomicName>
-,
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus halidayi" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="halidayi">Ismarus halidayi</taxonomicName>
<taxonomicName id="AB369579F5AF0B1B888219C9C5DF4050" genus="Ismarus" lsidName="Ismarus halidayi" pageId="2" pageNumber="51" rank="species" species="halidayi">Ismarus halidayi</taxonomicName>
-,
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus rex" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="rex">Ismarus rex</taxonomicName>
<taxonomicName id="79E51C8229981155019DE20A998481C0" genus="Ismarus" lsidName="Ismarus rex" pageId="2" pageNumber="51" rank="species" species="rex">Ismarus rex</taxonomicName>
- and
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus dorsiger" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="dorsiger">Ismarus dorsiger</taxonomicName>
<taxonomicName id="B0C87687C7047E0344A7FCB4934519E1" genus="Ismarus" lsidName="Ismarus dorsiger" pageId="2" pageNumber="51" rank="species" species="dorsiger">Ismarus dorsiger</taxonomicName>
-groups. The species of the present study belong to the
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus halidayi" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="halidayi">Ismarus halidayi</taxonomicName>
<taxonomicName id="789D0595ABF49DE48E72A9943B89E9E9" genus="Ismarus" lsidName="Ismarus halidayi" pageId="2" pageNumber="51" rank="species" species="halidayi">Ismarus halidayi</taxonomicName>
- and
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus dorsiger" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="dorsiger">Ismarus dorsiger</taxonomicName>
<taxonomicName id="9164F0B0128A9C17840A54C64E2F3CE5" genus="Ismarus" lsidName="Ismarus dorsiger" pageId="2" pageNumber="51" rank="species" species="dorsiger">Ismarus dorsiger</taxonomicName>
-groups. Compared to the diagnosis of
<taxonomicName class="Insecta" family="Diapriidae" genus="Ismarus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ismarus" order="Hymenoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">Ismarus</taxonomicName>
<taxonomicName id="FC27CE26C85E69FD2AD8E7F28BD79B23" genus="Ismarus" lsidName="Ismarus" pageId="2" pageNumber="51" rank="genus">Ismarus</taxonomicName>
given by
<bibRefCitation author="Masner, L" journalOrPublisher="Canadian Entomologist" pageId="8" pageNumber="57" pagination="1243 - 1266" title="A revision of the Ismarinae of the New World (Hymenoptera, Proctotrupoidea, Diapriidae)." url="doi:10.4039/Ent1081243-11" volume="108" year="1976">Masner (1976)</bibRefCitation>
<bibRefCitation id="F2C3CEFC1F04A56E79412B24B6E19275" author="Masner, L" journalOrPublisher="Canadian Entomologist" pageId="8" pageNumber="57" pagination="1243 - 1266" title="A revision of the Ismarinae of the New World (Hymenoptera, Proctotrupoidea, Diapriidae)." url="doi:10.4039/Ent1081243-11" volume="108" year="1976">Masner (1976)</bibRefCitation>
, we found that all species from China have the labrum exposed and possess a transverse carina on the lower face just below the antennal sockets.
</paragraph>
</subSubSection>

View file

@ -1,227 +1,227 @@
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Gryonoides mirabilicornis Masner &amp;
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<taxonomicNameLabel pageId="0" pageNumber="523">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Gryonoides mirabilicornis Masner &amp; Miko, sp. nov. A A 4 and A 5 showing elongate release and spread structures (rss) B release and spread structures (tyloids) correspond to class 3 gland cells (Noirot and Quennedey 1976) and are characterised by the presence of cuticular canals (arrows). Release and spread structures are present only on the fifth flagellomere in most Platygastroidea (except Sparasionini)." figureDoi="10.3897/jhr.87.72931.figure22" httpUri="https://binary.pensoft.net/fig/627303" pageId="0" pageNumber="523">Figs 22</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Gryonoides mirabilicornis Masner &amp; Miko, sp. nov. A head, anterior view B head and mesosoma, lateral view." figureDoi="10.3897/jhr.87.72931.figure23" httpUri="https://binary.pensoft.net/fig/627304" pageId="0" pageNumber="523">, 23</figureCitation>
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<paragraph id="EE52E2817ED79D4D3C690EA5B1A43341" pageId="0" pageNumber="523">
<figureCitation id="C84C6C5BA89E52936FF59AA572B6BA19" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Gryonoides mirabilicornis Masner &amp; Miko, sp. nov. A A 4 and A 5 showing elongate release and spread structures (rss) B release and spread structures (tyloids) correspond to class 3 gland cells (Noirot and Quennedey 1976) and are characterised by the presence of cuticular canals (arrows). Release and spread structures are present only on the fifth flagellomere in most Platygastroidea (except Sparasionini)." figureDoi="10.3897/jhr.87.72931.figure22" httpUri="https://binary.pensoft.net/fig/627303" pageId="0" pageNumber="523">Figs 22</figureCitation>
<figureCitation id="94608FCDD1DF039758ED4AEAD103EA97" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Gryonoides mirabilicornis Masner &amp; Miko, sp. nov. A head, anterior view B head and mesosoma, lateral view." figureDoi="10.3897/jhr.87.72931.figure23" httpUri="https://binary.pensoft.net/fig/627304" pageId="0" pageNumber="523">, 23</figureCitation>
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<paragraph pageId="0" pageNumber="523">Diagnosis.</paragraph>
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<emphasis italics="true" pageId="0" pageNumber="523">Gryonoides mirabilicornis</emphasis>
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<paragraph id="15CF37B648E12C7A146E0D9A67C16440" pageId="0" pageNumber="523">Diagnosis.</paragraph>
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<emphasis id="A44270533576087316B4524C97478DEC" italics="true" pageId="0" pageNumber="523">Gryonoides mirabilicornis</emphasis>
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differs from all other teleasine species in having tyloids (keel like release and spread structures,
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.28120" author="Isidoro, N" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="523" pagination="206 - 239" refId="B19" refString="Isidoro, N, Bin, F, Colazza, S, Vinson, SB, 1996. Morphology of antennal gustatory sensilla and glands in some parasitoid Hymenoptera with hypothesis on their role in sex and host recognition. Journal of Hymenoptera Research 5: 206 - 239, DOI: https://doi.org/10.5962/bhl.part.28120" title="Morphology of antennal gustatory sensilla and glands in some parasitoid Hymenoptera with hypothesis on their role in sex and host recognition." url="https://doi.org/10.5962/bhl.part.28120" volume="5" year="1996">Isidoro et al. 1996</bibRefCitation>
<bibRefCitation id="6CF775CCFBC373FC67A8973BB7840BDE" DOI="https://doi.org/10.5962/bhl.part.28120" author="Isidoro, N" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="523" pagination="206 - 239" refId="B19" refString="Isidoro, N, Bin, F, Colazza, S, Vinson, SB, 1996. Morphology of antennal gustatory sensilla and glands in some parasitoid Hymenoptera with hypothesis on their role in sex and host recognition. Journal of Hymenoptera Research 5: 206 - 239, DOI: https://doi.org/10.5962/bhl.part.28120" title="Morphology of antennal gustatory sensilla and glands in some parasitoid Hymenoptera with hypothesis on their role in sex and host recognition." url="https://doi.org/10.5962/bhl.part.28120" volume="5" year="1996">Isidoro et al. 1996</bibRefCitation>
) on A5 and A6.
</paragraph>
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<paragraph pageId="0" pageNumber="523">Description.</paragraph>
<paragraph pageId="0" pageNumber="523">
<subSubSection id="9499DFD6E095A3C8F7D58905CEFB9AD1" pageId="0" pageNumber="523" type="description">
<paragraph id="A49C03F706F5CFC366F49C3A1C534A9C" pageId="0" pageNumber="523">Description.</paragraph>
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Body length: 2620-2700
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="1881D766594F81C9F8C26D875605B6B4" originalValue="µm">µm</normalizedToken>
. Antenna color: scape and pedicel yellow, flagellum dark brown. Color of head: dark brown. Color of mesosoma: legs, tegula, lateral mesoscutellar spines, metascutellar spine yellowish, axillae, metanotum, lateral region of pronotal rim reddish, rest of mesosoma black. Color of metasoma: dark brown. Length of setae on male flagellomeres: more than 2 times as long as flagellomere width. Torular triangle: absent. Transverse setal fields on upper face: absent. Upper face sculpture: punctate dorsally. Upper face concavity dorsal view: convex. Central keel: absent. Head shape anterior view: head rounded in anterior view (longest head width in horizontal midline of head). Occipital carina structure dorsomedially: not crenulate. Facial striae dorsal end: not reaching midlevel of eye. Vertex sculpture: smooth. Notaulus: present. Notaulus anterior end: anterior to the transscutal line. Dorsal metapleural area: glabrous. Anteromedial pits of propodeum: absent. Area between plica and lateral propodeal carina sculpture: carinate (1 or 2 carinae present). Number of longitudinal carinae between plica and longitudinal (dorsal) section of lateral propodeal carina: 1. Medial region of lateral propodeal area pilosity: glabrous. Posterior propodeal projection length: more than two times as long as wide. Rugulose sculpture on T3: absent. T3 posterior 4/5th: smooth.
</paragraph>
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<subSubSection pageId="0" pageNumber="523" type="comments">
<paragraph pageId="0" pageNumber="523">Comments.</paragraph>
<paragraph pageId="0" pageNumber="523">
<subSubSection id="B8F23F7B598FBC61349AE04F4CBB313A" pageId="0" pageNumber="523" type="comments">
<paragraph id="C7BE183BBB8E5F0F01471328EDEED8D7" pageId="0" pageNumber="523">Comments.</paragraph>
<paragraph id="1CCCF9924A7D0938F84F5B5584BB702D" pageId="0" pageNumber="523">
We are not aware of a similar RSS pattern in
<taxonomicName lsidName="" pageId="0" pageNumber="523" rank="subFamily" subFamily="Teleasinae">Teleasinae</taxonomicName>
<taxonomicName id="0BDFF37434AEA13698DE474A0181F79C" lsidName="" pageId="0" pageNumber="523" rank="subFamily" subFamily="Teleasinae">Teleasinae</taxonomicName>
,
<taxonomicName lsidName="" pageId="0" pageNumber="523" rank="subFamily" subFamily="Telenominae">Telenominae</taxonomicName>
<taxonomicName id="92A05E431A5A847F75F48B28C9FE986D" lsidName="" pageId="0" pageNumber="523" rank="subFamily" subFamily="Telenominae">Telenominae</taxonomicName>
and more derived
<taxonomicName authorityName="Haliday" authorityYear="1839" lsidName="" pageId="0" pageNumber="523" rank="subFamily" subFamily="Scelioninae">Scelioninae</taxonomicName>
<taxonomicName id="02BEC17F8CBDF1B8A8481432676E055A" authorityName="Haliday" authorityYear="1839" lsidName="" pageId="0" pageNumber="523" rank="subFamily" subFamily="Scelioninae">Scelioninae</taxonomicName>
. The geometry of the female antenna and the distribution pattern of uniporous olfactory sensilla have been reported to correspond to the male RSS (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.28120" author="Isidoro, N" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="523" pagination="206 - 239" refId="B19" refString="Isidoro, N, Bin, F, Colazza, S, Vinson, SB, 1996. Morphology of antennal gustatory sensilla and glands in some parasitoid Hymenoptera with hypothesis on their role in sex and host recognition. Journal of Hymenoptera Research 5: 206 - 239, DOI: https://doi.org/10.5962/bhl.part.28120" title="Morphology of antennal gustatory sensilla and glands in some parasitoid Hymenoptera with hypothesis on their role in sex and host recognition." url="https://doi.org/10.5962/bhl.part.28120" volume="5" year="1996">Isidoro et al. 1996</bibRefCitation>
<bibRefCitation id="381C0B4F3831D7F47A515554B79B5FE6" DOI="https://doi.org/10.5962/bhl.part.28120" author="Isidoro, N" journalOrPublisher="Journal of Hymenoptera Research" pageId="0" pageNumber="523" pagination="206 - 239" refId="B19" refString="Isidoro, N, Bin, F, Colazza, S, Vinson, SB, 1996. Morphology of antennal gustatory sensilla and glands in some parasitoid Hymenoptera with hypothesis on their role in sex and host recognition. Journal of Hymenoptera Research 5: 206 - 239, DOI: https://doi.org/10.5962/bhl.part.28120" title="Morphology of antennal gustatory sensilla and glands in some parasitoid Hymenoptera with hypothesis on their role in sex and host recognition." url="https://doi.org/10.5962/bhl.part.28120" volume="5" year="1996">Isidoro et al. 1996</bibRefCitation>
) in some parasitic
<taxonomicName kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="523" rank="order">Hymenoptera</taxonomicName>
<taxonomicName id="978FB192A9C46BC2FA74D86AAE2C519E" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="0" pageNumber="523" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
.
<bibRefCitation author="Bin, F" journalOrPublisher="Entomologica" pageId="0" pageNumber="523" pagination="33 - 97" refId="B5" refString="Bin, F, Colazza, S, Isidoro, N, Solinas, M, Vinson, SB, 1989. Antennal chemosensilla and glands, and their possible meaning in the reproductive behavior of Trissolcus basalis (Woll.) (Hym.: Scelionidae). Entomologica 24: 33 - 97" title="Antennal chemosensilla and glands, and their possible meaning in the reproductive behavior of Trissolcus basalis (Woll.) (Hym.: Scelionidae)." volume="24" year="1989">Bin et al. 1989</bibRefCitation>
<bibRefCitation id="D6A118EE75C7B663EA237695E769A0F1" author="Bin, F" journalOrPublisher="Entomologica" pageId="0" pageNumber="523" pagination="33 - 97" refId="B5" refString="Bin, F, Colazza, S, Isidoro, N, Solinas, M, Vinson, SB, 1989. Antennal chemosensilla and glands, and their possible meaning in the reproductive behavior of Trissolcus basalis (Woll.) (Hym.: Scelionidae). Entomologica 24: 33 - 97" title="Antennal chemosensilla and glands, and their possible meaning in the reproductive behavior of Trissolcus basalis (Woll.) (Hym.: Scelionidae)." volume="24" year="1989">Bin et al. 1989</bibRefCitation>
have observed that
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<emphasis italics="true" pageId="0" pageNumber="523">Trissolcus basalis</emphasis>
<taxonomicName id="EBC152A615A19508FEFEA73C8FBC2EE1" baseAuthorityName="Wollaston" baseAuthorityYear="1858" class="Hexapoda" family="Platygastridae" genus="Trissolcus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Trissolcus basalis" order="Hymenoptera" pageId="0" pageNumber="523" phylum="Arthropoda" rank="species" species="basalis">
<emphasis id="67D1AD620A717A3F7A40E774307F0F4F" italics="true" pageId="0" pageNumber="523">Trissolcus basalis</emphasis>
</taxonomicName>
males are touching the female antenna during copulation &quot;entwining his around hers and repeatedly&quot;. Although a the female receptor for the gland extract has not been identified, it is most reasonable to assume that the distinct change in male RSS morphology in
<taxonomicName lsidName="G. mirabilicornis" pageId="0" pageNumber="523" rank="species" species="mirabilicornis">
<emphasis italics="true" pageId="0" pageNumber="523">G. mirabilicornis</emphasis>
<taxonomicName id="08B0B615E6BE02E9992CAAC828D3DE00" lsidName="G. mirabilicornis" pageId="0" pageNumber="523" rank="species" species="mirabilicornis">
<emphasis id="8FF5898803BD8A998607C7D366F0DF8A" italics="true" pageId="0" pageNumber="523">G. mirabilicornis</emphasis>
</taxonomicName>
most likely corresponds to unique modifications on the female antenna of the same species. Specimens of
<taxonomicName lsidName="G. mirabilicornis" pageId="0" pageNumber="523" rank="species" species="mirabilicornis">
<emphasis italics="true" pageId="0" pageNumber="523">G. mirabilicornis</emphasis>
<taxonomicName id="43F0E07C11C66B474AE627DE8EE35E12" lsidName="G. mirabilicornis" pageId="0" pageNumber="523" rank="species" species="mirabilicornis">
<emphasis id="80259B4210230D25103CD947A205F667" italics="true" pageId="0" pageNumber="523">G. mirabilicornis</emphasis>
</taxonomicName>
were collected together along with some female specimens of
<taxonomicName lsidName="G. flaviclavus" pageId="0" pageNumber="523" rank="species" species="flaviclavus">
<emphasis italics="true" pageId="0" pageNumber="523">G. flaviclavus</emphasis>
<taxonomicName id="6CEC3AF95B1D490D993FF22F45268CC7" lsidName="G. flaviclavus" pageId="0" pageNumber="523" rank="species" species="flaviclavus">
<emphasis id="DD3590F04A3996638CBD743BE2E50FC2" italics="true" pageId="0" pageNumber="523">G. flaviclavus</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="G. fuscoclavatus" pageId="0" pageNumber="523" rank="species" species="fuscoclavatus">
<emphasis italics="true" pageId="0" pageNumber="523">G. fuscoclavatus</emphasis>
<taxonomicName id="99F50FD62746A1E458A4C99771234DBD" lsidName="G. fuscoclavatus" pageId="0" pageNumber="523" rank="species" species="fuscoclavatus">
<emphasis id="E605453BE9443BAE709DCCBC5308BB96" italics="true" pageId="0" pageNumber="523">G. fuscoclavatus</emphasis>
</taxonomicName>
. We did not find differences in the antennal morphology and sensilla pattern between the female antenna of these two species and other
<taxonomicName class="Insecta" family="Platygastridae" genus="Gryonoides" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gryonoides" order="Hymenoptera" pageId="0" pageNumber="523" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="523">Gryonoides</emphasis>
<taxonomicName id="E657A432244230F2FA112466782F66E1" genus="Gryonoides" kingdom="Animalia" lsidName="Gryonoides" order="Hymenoptera" pageId="0" pageNumber="523" rank="genus">
<emphasis id="0954357FE85648B4131B2580FB7EB120" italics="true" pageId="0" pageNumber="523">Gryonoides</emphasis>
</taxonomicName>
. Therefore, we consider the 3 male specimens with modified antennae to be a separate species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="523" type="materials_examined">
<paragraph pageId="0" pageNumber="523">Material.</paragraph>
<paragraph pageId="0" pageNumber="523">
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<typeStatus>Holotype</typeStatus>
<subSubSection id="93E2AD6B8A3BDA8589B0005F633AED5D" pageId="0" pageNumber="523" type="materials_examined">
<paragraph id="CC8C7533E67239F2EF8E4FF0452CB6EF" pageId="0" pageNumber="523">Material.</paragraph>
<paragraph id="A1696EAA4BF4768F5940E6AA85ED91C3" pageId="0" pageNumber="523">
<materialsCitation id="AC0DA3A6DAE8E6D63C4997CBEC324AC2" collectingDate="2011-11-28" collectorName="J. Heraty" country="PERU" county="Pilicopata" elevation="518" latitude="-12.884167" location="Manu" longLatPrecision="21" longitude="-71.40445" municipality="Carmen" specimenCount="1" typeStatus="Holotype">
<typeStatus id="DBD612D869F290FA7613D9749293645E">Holotype</typeStatus>
: male, UNHC_0020586,
<collectingCountry name="Peru">PERU</collectingCountry>
<collectingCountry id="4235D4D404B4EAD081ECE37B5A4E7737" name="Peru">PERU</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:A193ED26830D5F4B96A9FFDDB4012BBB:C02391B454466EF8517D2EBAB43DD58F" country="PERU" county="Pilicopata" latitude="-12.884167" longLatPrecision="21" longitude="-71.40445" municipality="Carmen" name="Manu">Manu</location>
<location id="C02391B454466EF8517D2EBAB43DD58F" LSID="urn:lsid:plazi:treatment:A193ED26830D5F4B96A9FFDDB4012BBB:C02391B454466EF8517D2EBAB43DD58F" country="PERU" county="Pilicopata" latitude="-12.884167" longLatPrecision="21" longitude="-71.40445" municipality="Carmen" name="Manu">Manu</location>
,
<location LSID="urn:lsid:plazi:treatment:A193ED26830D5F4B96A9FFDDB4012BBB:249CC5E88FFA3609BCF3BBC201F0F032" country="PERU" county="Pilicopata" latitude="-12.884167" longLatPrecision="21" longitude="-71.40445" municipality="Carmen" name="Villa">Villa</location>
<location id="249CC5E88FFA3609BCF3BBC201F0F032" LSID="urn:lsid:plazi:treatment:A193ED26830D5F4B96A9FFDDB4012BBB:249CC5E88FFA3609BCF3BBC201F0F032" country="PERU" county="Pilicopata" latitude="-12.884167" longLatPrecision="21" longitude="-71.40445" municipality="Carmen" name="Villa">Villa</location>
,
<collectingMunicipality>Carmen</collectingMunicipality>
<collectingMunicipality id="D39D5F448B967069B6B7C18D6B44983A">Carmen</collectingMunicipality>
,
<collectingCounty>Pilicopata</collectingCounty>
<collectingCounty id="B5C9A870CB165386908C02F195ED4789">Pilicopata</collectingCounty>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.18" unit="m" value="518.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.18" unit="m" value="518.0">518m</elevation>
<quantity id="58D13358618791B65A9B912692C65936" metricMagnitude="2" metricUnit="m" metricValue="5.18" unit="m" value="518.0">
<elevation id="B26ACA9A300E0A185A4829F58EF81C05" metricMagnitude="2" metricUnit="m" metricValue="5.18" unit="m" value="518.0">518m</elevation>
</quantity>
,
<geoCoordinate degrees="12" direction="south" minutes="53" orientation="latitude" precision="15" seconds="03" value="-12.884167">
<geoCoordinate id="A99421D0D245EC01B18F6020C17266AD" degrees="12" direction="south" minutes="53" orientation="latitude" precision="15" seconds="03" value="-12.884167">
12°53'03
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S
</geoCoordinate>
,
<geoCoordinate degrees="71" direction="west" minutes="24" orientation="longitude" precision="15" seconds="16" value="-71.40445">
<geoCoordinate id="175343096A93C3DA5EC385E8C7CC4B8C" degrees="71" direction="west" minutes="24" orientation="longitude" precision="15" seconds="16" value="-71.40445">
71°24'16
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W
</geoCoordinate>
,
<collectingDate value="2011-11-28">28.xi.2011</collectingDate>
<collectingDate id="771E465A37B056B0BF7720552245F3B1" value="2011-11-28">28.xi.2011</collectingDate>
,
<collectorName>J. Heraty</collectorName>
<collectorName id="39DB2886CD44A5EC5FA7F5979DFB4364">J. Heraty</collectorName>
, mature forest, sweeping (CNC)
</materialsCitation>
.
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="PERU" location="PERU" specimenCount="2" specimenCount-male="2" typeStatus="Paratypes">
<typeStatus>Paratypes</typeStatus>
<materialsCitation id="E6DC66E87345AE8CD6F0721D22FDB2AD" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="PERU" location="PERU" specimenCount="2" specimenCount-male="2" typeStatus="Paratypes">
<typeStatus id="D6AEB698EDD2FA4D1234D5DF93A668A5">Paratypes</typeStatus>
:
<collectingCountry name="Peru">PERU</collectingCountry>
<collectingCountry id="EC52C1AF319D5DB2790D51FB27E45698" name="Peru">PERU</collectingCountry>
,
<specimenCount type="male">2 males</specimenCount>
<specimenCount id="F5A0BFC93ED81ABD520963F8DFC1AA8A" type="male">2 males</specimenCount>
(CNC)
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</paragraph>

View file

@ -1,431 +1,431 @@
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Schallenberg, Vivianne M.</mods:namePart>
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<mods:namePart id="FF36CDD8A54C24804BD3B38AA9CDEFEA">Schallenberg, Vivianne M.</mods:namePart>
<mods:affiliation id="B5C597C8A12C933508CB3A32E97E227D">Natural History Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland</mods:affiliation>
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<mods:role>
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<mods:namePart>Heim, Rene</mods:namePart>
<mods:affiliation>St. - Karli-Strasse 32 c, CH- 6004 Lucerne, Switzerland</mods:affiliation>
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<mods:affiliation id="A259CF13B3ED051E51F56A74A7013BC9">St. - Karli-Strasse 32 c, CH- 6004 Lucerne, Switzerland</mods:affiliation>
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<mods:namePart>Schneppat, Ulrich E.</mods:namePart>
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<mods:namePart id="F5116ABFF498567810F69804BD669265">Schneppat, Ulrich E.</mods:namePart>
<mods:affiliation id="BFC4C629F708EC890A022099101426A5">Natur Museum Lucerne, Kasernenplatz 6, CH- 6003 Lucerne, Switzerland</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mueller, Peter</mods:namePart>
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<mods:namePart id="1B758D07D358AE1E2D64F182B19497A2">Mueller, Peter</mods:namePart>
<mods:affiliation id="25182799B5C5CEBE0CC4E3C9D60207E1">Sennereiweg 8, CH- 7074 Churwalden-Malix, Switzerland</mods:affiliation>
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<mods:role>
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<mods:namePart>Rueetschi, Joerg</mods:namePart>
<mods:affiliation>Witikonerstrasse 180, CH- 8053 Zurich, Switzerland</mods:affiliation>
<mods:namePart id="09C2EAA1C7A18656E5639D4962F86D51">Rueetschi, Joerg</mods:namePart>
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</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="FBA47EE0052826C8AFA33EA5AC00ADA6" type="personal">
<mods:role id="0BC0839C1CBB383AD0F532D9BDE89634">
<mods:roleTerm id="89526431ECEB8371960505CDF4F184EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neubert, Eike</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0277-2894</mods:nameIdentifier>
<mods:affiliation>Natural History Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland</mods:affiliation>
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<treatment LSID="urn:lsid:plazi:treatment:A19686D831A353DBBFC9397668CD49AC" httpUri="http://treatment.plazi.org/id/A19686D831A353DBBFC9397668CD49AC" lastPageNumber="149" pageId="0" pageNumber="149">
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<taxonomicName LSID="A19686D8-31A3-53DB-BFC9-397668CD49AC" authority="(Millet, 1843)" baseAuthorityName="Millet" baseAuthorityYear="1843" class="Gastropoda" family="Milacidae" genus="Tandonia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tandonia rustica" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica">Tandonia rustica (Millet, 1843)</taxonomicName>
<treatment id="A19686D831A353DBBFC9397668CD49AC" ID-DOI="http://doi.org/10.5281/zenodo.7019329" ID-GBIF-Taxon="198934343" ID-Zenodo-Dep="7019329" LSID="urn:lsid:plazi:treatment:A19686D831A353DBBFC9397668CD49AC" httpUri="http://treatment.plazi.org/id/A19686D831A353DBBFC9397668CD49AC" lastPageNumber="149" pageId="0" pageNumber="149" scope_family="Milacidae" scope_phylum="Mollusca" scope_superFamily="Parmacelloidea" scope_superOrder="Eupulmonata">
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<taxonomicName id="415297D727B5178A976A036D875AF2A2" ID-CoL="54P2K" LSID="A19686D8-31A3-53DB-BFC9-397668CD49AC" authority="(Millet, 1843)" baseAuthorityName="Millet" baseAuthorityYear="1843" class="Gastropoda" family="Milacidae" genus="Tandonia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tandonia rustica" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica">Tandonia rustica (Millet, 1843)</taxonomicName>
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<paragraph pageId="0" pageNumber="149">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of collected specimens." figureDoi="10.3897/zookeys.1116.82762.figure1" httpUri="https://binary.pensoft.net/fig/725583" pageId="0" pageNumber="149">Figs 1</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">, 3</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Tandonia. A, B Tandonia rustica A sequenced specimen NMBE 568974, Wartau, Truebbach B sequenced specimen NMBE 568973, Rovio, Sovaglia C, D Tandonia rustica (two mottled specimens) meeting a pale colour morph specimen of Tandonia nigra, Val Cuerta, Mt. Generoso E, F Tandonia budapestensis E NMBE 571881, Roggwil, F NMBE 571882, Meggen." figureDoi="10.3897/zookeys.1116.82762.figure7" httpUri="https://binary.pensoft.net/fig/725589" pageId="0" pageNumber="149">, 7A-D</figureCitation>
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<figureCitation id="38331449AD25A8D8677273BA2D4E3C3D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of collected specimens." figureDoi="10.3897/zookeys.1116.82762.figure1" httpUri="https://binary.pensoft.net/fig/725583" pageId="0" pageNumber="149">Figs 1</figureCitation>
<figureCitation id="DF7CEDC1DB4DEE91B103132D03CB2590" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">, 3</figureCitation>
<figureCitation id="541D949AB69EB60FFB40A58F292C198E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Tandonia. A, B Tandonia rustica A sequenced specimen NMBE 568974, Wartau, Truebbach B sequenced specimen NMBE 568973, Rovio, Sovaglia C, D Tandonia rustica (two mottled specimens) meeting a pale colour morph specimen of Tandonia nigra, Val Cuerta, Mt. Generoso E, F Tandonia budapestensis E NMBE 571881, Roggwil, F NMBE 571882, Meggen." figureDoi="10.3897/zookeys.1116.82762.figure7" httpUri="https://binary.pensoft.net/fig/725589" pageId="0" pageNumber="149">, 7A-D</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="149">
<taxonomicName authorityName="Millet" authorityYear="1843" class="Gastropoda" family="Milacidae" genus="Limax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limax rustica" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica">Limax rustica</taxonomicName>
<subSubSection id="84E6910A4CF3321D4A7B75F13207C8AA" pageId="0" pageNumber="149" type="reference_group">
<paragraph id="D63049573E87960D660D40C59A37CD2D" pageId="0" pageNumber="149">
<taxonomicName id="BC8D125225FBD82A358BD90C43196421" authorityName="Millet" authorityYear="1843" class="Gastropoda" family="Milacidae" genus="Limax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limax rustica" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica">Limax rustica</taxonomicName>
Millet, 1843, Magasin de Zoologie, ser. 2, 5: 1, plate 63, fig. 1 [nord de
<normalizedToken originalValue="lAnjou">l'Anjou</normalizedToken>
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,
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la Bouillant, commune de la Chapelle-Hullin, ainsi
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, etc.].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="type specimens">
<paragraph pageId="0" pageNumber="149">Type specimens.</paragraph>
<paragraph pageId="0" pageNumber="149">
<subSubSection id="EF45ADB57C47F701F99BC28BC0179E7E" pageId="0" pageNumber="149" type="type specimens">
<paragraph id="51D2B03C3B60700619F4F95EFC863A41" pageId="0" pageNumber="149">Type specimens.</paragraph>
<paragraph id="1244047AD01BB74B22E7D023E07C4235" pageId="0" pageNumber="149">
Probably does not exist, after
<bibRefCitation author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">Wiktor (1987</bibRefCitation>
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: 286). See also remarks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="differential diagnosis">
<paragraph pageId="0" pageNumber="149">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="149">
<subSubSection id="E7A9E059BE4BEBCA59D2A01B98737C5F" pageId="0" pageNumber="149" type="differential diagnosis">
<paragraph id="B1B1AA1E8AEEB218845E4ECBE55FE0D0" pageId="0" pageNumber="149">Differential diagnosis.</paragraph>
<paragraph id="ADAF73B55FC1D28F96BF40F9E1266840" pageId="0" pageNumber="149">
Epiphallus more than twice as long as the penis (almost the same length in
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. nigra" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="nigra">
<emphasis italics="true" pageId="0" pageNumber="149">T. nigra</emphasis>
<taxonomicName id="1989E98D574AA10831857715F5354948" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. nigra" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="nigra">
<emphasis id="7021615CD343C1A9D74A7628999D4598" italics="true" pageId="0" pageNumber="149">T. nigra</emphasis>
</taxonomicName>
and
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. budapestensis" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="budapestensis">
<emphasis italics="true" pageId="0" pageNumber="149">T. budapestensis</emphasis>
<taxonomicName id="A5F26A9168C12D53542E23645A4B008B" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. budapestensis" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="budapestensis">
<emphasis id="C2721E4DD39F1FE689EEAAF8A8E3C58F" italics="true" pageId="0" pageNumber="149">T. budapestensis</emphasis>
</taxonomicName>
); dark stripe above the pneumostome and on the opposite side of the mantle (missing in
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. nigra" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="nigra">
<emphasis italics="true" pageId="0" pageNumber="149">T. nigra</emphasis>
<taxonomicName id="02BC46413BF30FB1007A1004AC4069E8" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. nigra" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="nigra">
<emphasis id="91918D2B5D41EE56B521B2F3E71B7250" italics="true" pageId="0" pageNumber="149">T. nigra</emphasis>
</taxonomicName>
and
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. budapestensis" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="budapestensis">
<emphasis italics="true" pageId="0" pageNumber="149">T. budapestensis</emphasis>
<taxonomicName id="C57242B49FDCCF062FCE10C133D4FE1B" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. budapestensis" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="budapestensis">
<emphasis id="88B019A9C81257D5CC5EEFF9AF6C6BED" italics="true" pageId="0" pageNumber="149">T. budapestensis</emphasis>
</taxonomicName>
); penial papilla short, blunt, with flap-like lobes around the central porus (short and blunt but folded in
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. nigra" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="nigra">
<emphasis italics="true" pageId="0" pageNumber="149">T. nigra</emphasis>
<taxonomicName id="2B3A23DECC323A8672D544FD76957C72" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. nigra" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="nigra">
<emphasis id="43EC0457EC7CF95B7B0D26F8F4AD640B" italics="true" pageId="0" pageNumber="149">T. nigra</emphasis>
</taxonomicName>
; elongated with a stalk in
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. budapestensis" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="budapestensis">
<emphasis italics="true" pageId="0" pageNumber="149">T. budapestensis</emphasis>
<taxonomicName id="DDC6BF34495ACB26DB0A8370647FB15E" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. budapestensis" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="budapestensis">
<emphasis id="49F66C042E228F2C4EC87D68FDC2AC36" italics="true" pageId="0" pageNumber="149">T. budapestensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="description">
<paragraph pageId="0" pageNumber="149">Description.</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">
<emphasis italics="true" pageId="0" pageNumber="149">Colouration</emphasis>
<subSubSection id="22C0B0216B9D230E4EAB5FD256F3D534" pageId="0" pageNumber="149" type="description">
<paragraph id="7538884B5D66A6137140CEE95A90EE03" pageId="0" pageNumber="149">Description.</paragraph>
<paragraph id="5EAF74725671A5CB1EDD01F71310ED82" pageId="0" pageNumber="149">
<emphasis id="B82FD3925F65F515384AE9CAAD46ED0A" bold="true" pageId="0" pageNumber="149">
<emphasis id="0B25AC1F53343B76FF97D15078485F50" italics="true" pageId="0" pageNumber="149">Colouration</emphasis>
.
</emphasis>
Many fully adult specimens (
<emphasis italics="true" pageId="0" pageNumber="149">n</emphasis>
<emphasis id="370C167D7E09C01CB92D54107A63B0CC" italics="true" pageId="0" pageNumber="149">n</emphasis>
= 56) were studied from six different Swiss Cantons (Bern, St Gallen, Grisons, Ticino, Obwalden, Nidwalden) as well as from five regions in Italy (Bolzano, Sondrio, Torino, Lecco, Trento), from the Republic of San Marino and from two Departments of France (Alpes de Haute Provence,
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).
</paragraph>
<paragraph pageId="0" pageNumber="149">
<paragraph id="40607FF2F8BD7FE170D156AC558FE36D" pageId="0" pageNumber="149">
The general pigmentation of Swiss specimens varies from a warm reddish brown to dark reddish or chestnut brown to very dark brown without any reddish brown hue. This pigmentation normally is not fading downwards to the fringe of the sole. However, in some cases in the less dark pigmented populations in Switzerland, little fading downwards can be observed (Fig.
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<figureCitation id="1C076AF20121030E7E68D2964546F37B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Tandonia. A, B Tandonia rustica A sequenced specimen NMBE 568974, Wartau, Truebbach B sequenced specimen NMBE 568973, Rovio, Sovaglia C, D Tandonia rustica (two mottled specimens) meeting a pale colour morph specimen of Tandonia nigra, Val Cuerta, Mt. Generoso E, F Tandonia budapestensis E NMBE 571881, Roggwil, F NMBE 571882, Meggen." figureDoi="10.3897/zookeys.1116.82762.figure7" httpUri="https://binary.pensoft.net/fig/725589" pageId="0" pageNumber="149">7A</figureCitation>
). The mantle is of the same colour as the dorsum, except for the surrounding of the pneumostome, which usually is paler. Many specimens investigated are markedly darker when compared to
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,
<bibRefCitation author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">1987</bibRefCitation>
<bibRefCitation id="920623884799A22AC1F3702AD6CEFEFE" author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">1987</bibRefCitation>
) and
<bibRefCitation author="Rowson, B" journalOrPublisher="Field Studies Council, Telford, UK" pageId="0" pageNumber="149" refId="B31" refString="Rowson, B, Turner, JA, Anderson, R, Symondson, WOC, 2014b. The Slugs of Britain and Ireland: Identification, understanding and control. Field Studies Council, Telford, UK" title="The Slugs of Britain and Ireland: Identification, understanding and control." year="2014 b">Rowson et al. (2014b)</bibRefCitation>
<bibRefCitation id="7C0A8373525AB830F31926404618F618" author="Rowson, B" journalOrPublisher="Field Studies Council, Telford, UK" pageId="0" pageNumber="149" refId="B31" refString="Rowson, B, Turner, JA, Anderson, R, Symondson, WOC, 2014b. The Slugs of Britain and Ireland: Identification, understanding and control. Field Studies Council, Telford, UK" title="The Slugs of Britain and Ireland: Identification, understanding and control." year="2014 b">Rowson et al. (2014b)</bibRefCitation>
. The darkest specimens were found in southern Switzerland (Ticino, Grisons - Val Bregaglia), and in the southwest bordering areas to Italy and France (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Tandonia. A, B Tandonia rustica A sequenced specimen NMBE 568974, Wartau, Truebbach B sequenced specimen NMBE 568973, Rovio, Sovaglia C, D Tandonia rustica (two mottled specimens) meeting a pale colour morph specimen of Tandonia nigra, Val Cuerta, Mt. Generoso E, F Tandonia budapestensis E NMBE 571881, Roggwil, F NMBE 571882, Meggen." figureDoi="10.3897/zookeys.1116.82762.figure7" httpUri="https://binary.pensoft.net/fig/725589" pageId="0" pageNumber="149">7B</figureCitation>
<figureCitation id="19C8B919CF7C99F49E749744B56B406C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Tandonia. A, B Tandonia rustica A sequenced specimen NMBE 568974, Wartau, Truebbach B sequenced specimen NMBE 568973, Rovio, Sovaglia C, D Tandonia rustica (two mottled specimens) meeting a pale colour morph specimen of Tandonia nigra, Val Cuerta, Mt. Generoso E, F Tandonia budapestensis E NMBE 571881, Roggwil, F NMBE 571882, Meggen." figureDoi="10.3897/zookeys.1116.82762.figure7" httpUri="https://binary.pensoft.net/fig/725589" pageId="0" pageNumber="149">7B</figureCitation>
). Specimens from Central Switzerland (Lucerne, Nidwalden, Obwalden), northern and eastern Grisons, St. Gallen, and the eastern border area in Italy (Bolzano, Trento) are of a markedly lighter, pale reddish brown pigmentation (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Tandonia. A, B Tandonia rustica A sequenced specimen NMBE 568974, Wartau, Truebbach B sequenced specimen NMBE 568973, Rovio, Sovaglia C, D Tandonia rustica (two mottled specimens) meeting a pale colour morph specimen of Tandonia nigra, Val Cuerta, Mt. Generoso E, F Tandonia budapestensis E NMBE 571881, Roggwil, F NMBE 571882, Meggen." figureDoi="10.3897/zookeys.1116.82762.figure7" httpUri="https://binary.pensoft.net/fig/725589" pageId="0" pageNumber="149">7A, C, D</figureCitation>
<figureCitation id="07B4F8EEA794A55911BB1AD8675BDEBF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Tandonia. A, B Tandonia rustica A sequenced specimen NMBE 568974, Wartau, Truebbach B sequenced specimen NMBE 568973, Rovio, Sovaglia C, D Tandonia rustica (two mottled specimens) meeting a pale colour morph specimen of Tandonia nigra, Val Cuerta, Mt. Generoso E, F Tandonia budapestensis E NMBE 571881, Roggwil, F NMBE 571882, Meggen." figureDoi="10.3897/zookeys.1116.82762.figure7" httpUri="https://binary.pensoft.net/fig/725589" pageId="0" pageNumber="149">7A, C, D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="149">
<paragraph id="D718833479034EB143AE40DBF577EF48" pageId="0" pageNumber="149">
The whole slug is covered with small black spots on dorsum, flanks, and mantle, but not on the keel. The characteristic black streaks above the pneumostome and on the same location on the left side of the mantle can vary greatly in size and form. Even though all our investigated specimens had spots and the characteristic streaks on the mantle, in some specimens they are almost invisible because of the overall dark pigmentation (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Tandonia. A, B Tandonia rustica A sequenced specimen NMBE 568974, Wartau, Truebbach B sequenced specimen NMBE 568973, Rovio, Sovaglia C, D Tandonia rustica (two mottled specimens) meeting a pale colour morph specimen of Tandonia nigra, Val Cuerta, Mt. Generoso E, F Tandonia budapestensis E NMBE 571881, Roggwil, F NMBE 571882, Meggen." figureDoi="10.3897/zookeys.1116.82762.figure7" httpUri="https://binary.pensoft.net/fig/725589" pageId="0" pageNumber="149">7B</figureCitation>
<figureCitation id="FA28E74F69404CF7D2221C222CADBAA2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Living specimens of Tandonia. A, B Tandonia rustica A sequenced specimen NMBE 568974, Wartau, Truebbach B sequenced specimen NMBE 568973, Rovio, Sovaglia C, D Tandonia rustica (two mottled specimens) meeting a pale colour morph specimen of Tandonia nigra, Val Cuerta, Mt. Generoso E, F Tandonia budapestensis E NMBE 571881, Roggwil, F NMBE 571882, Meggen." figureDoi="10.3897/zookeys.1116.82762.figure7" httpUri="https://binary.pensoft.net/fig/725589" pageId="0" pageNumber="149">7B</figureCitation>
). Additionally spots in darker specimen may be much larger and sometimes irregularly jagged and elongated to streaks.
</paragraph>
<paragraph pageId="0" pageNumber="149">Head, neck and ommatophores are dark brown, while the tentacles are slightly paler. In many cases, the black pigmented ommatophoran retractor is clearly visible through the integument.</paragraph>
<paragraph pageId="0" pageNumber="149">The keel is commonly paler in colour than the dorsum. Rarely the general pigmentation of the keel matches the dorsal colour, this occurs especially in darker specimens.</paragraph>
<paragraph pageId="0" pageNumber="149">The tripartite sole in pale coloured slugs is creamy yellowish, while darker slugs have a pale yellowish brown sole. Many darker coloured specimens have a hue of greyish black at the posterior outer margins of the lateral sole fields formed by very little black dots. Isolated single greyish black spots may occur along the outer edge of the sole.</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">
<emphasis italics="true" pageId="0" pageNumber="149">Mantle structure</emphasis>
<paragraph id="BAE4BC7FF1BCE45A6326C9611EED4186" pageId="0" pageNumber="149">Head, neck and ommatophores are dark brown, while the tentacles are slightly paler. In many cases, the black pigmented ommatophoran retractor is clearly visible through the integument.</paragraph>
<paragraph id="051B6125FFDB29FC0E3D737F7F12A341" pageId="0" pageNumber="149">The keel is commonly paler in colour than the dorsum. Rarely the general pigmentation of the keel matches the dorsal colour, this occurs especially in darker specimens.</paragraph>
<paragraph id="AF4F93F174C6C82049F73CE5256FE552" pageId="0" pageNumber="149">The tripartite sole in pale coloured slugs is creamy yellowish, while darker slugs have a pale yellowish brown sole. Many darker coloured specimens have a hue of greyish black at the posterior outer margins of the lateral sole fields formed by very little black dots. Isolated single greyish black spots may occur along the outer edge of the sole.</paragraph>
<paragraph id="F94E04962F02BE32598123794CAA9CB2" pageId="0" pageNumber="149">
<emphasis id="2ABB4BF64D19D4685CDE2B56AFA0B99A" bold="true" pageId="0" pageNumber="149">
<emphasis id="96EB15D445B0FEDAF5EEDA7A32923EE7" italics="true" pageId="0" pageNumber="149">Mantle structure</emphasis>
.
</emphasis>
The pneumostome is positioned at 2/3 of mantle length, well posterior of the centre of the mantle. The pneumostome is not surrounded by a distinct ring-like structure, like it is the case in
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. nigra" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="nigra">
<emphasis italics="true" pageId="0" pageNumber="149">T. nigra</emphasis>
<taxonomicName id="745960F70C05715552C33F25A99E8B48" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. nigra" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="nigra">
<emphasis id="CEBCA87AC6EEB855EB7655EC2AD1FE59" italics="true" pageId="0" pageNumber="149">T. nigra</emphasis>
</taxonomicName>
. The
<normalizedToken originalValue="“slit”">&quot;slit&quot;</normalizedToken>
<normalizedToken id="F1A98F7EF6DF15D953740EC6DF4F5AD6" originalValue="“slit”">&quot;slit&quot;</normalizedToken>
of the pneumostome in all specimens does not end in the lumen of the pneumostome but runs anteriorly to at least the dorsal edge of pneumostome. In living individuals, the mantle surface is completely smooth besides the horseshoe-shaped sinus groove. The sinus groove is completely developed in all specimens in our series, and it reaches at both sides almost the posterior end of the mantle. In the dark specimens it needs magnification to see it clearly. The posterior margin of the mantle is not tightly attached to the integument and in living and contracted animals smoothly rounded. The posterior free mantle flap covers the anterior integument-tubercles as well as the openings of the postpallial or
<normalizedToken originalValue="Wiktors">Wiktor's</normalizedToken>
<normalizedToken id="C7993F190A21453FB0649FC1182AB923" originalValue="Wiktors">Wiktor's</normalizedToken>
pocket organ.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">
<emphasis italics="true" pageId="0" pageNumber="149">Postpallial pocket organ</emphasis>
<paragraph id="645CEA7A3010B90F8E632339BF542EB4" pageId="0" pageNumber="149">
<emphasis id="E593839F8FBAD5F491348AC641297695" bold="true" pageId="0" pageNumber="149">
<emphasis id="5349E1F2918EA4B7EE564B69F678F315" italics="true" pageId="0" pageNumber="149">Postpallial pocket organ</emphasis>
.
</emphasis>
In all specimens examined, the posterior part of the mantle covered two slit-like openings, the postpallial pocket organ, which was first detected and described by
<bibRefCitation DOI="https://doi.org/10.2478/v10125-011-0025-4" author="Schneppat, UE" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="149" pagination="225 - 248" refId="B34" refString="Schneppat, UE, Georgiev, DG, Dedov, IK, Wondrak, G, Knechtle, F, 2011. Tandonia totevi (Wiktor, 1975) (Pulmonata: Milacidae) in Bulgaria and north eastern Greece. Folia Malacologica 19 (4): 225 - 248, DOI: https://doi.org/10.2478/v10125-011-0025-4" title="Tandonia totevi (Wiktor, 1975) (Pulmonata: Milacidae) in Bulgaria and north eastern Greece." url="https://doi.org/10.2478/v10125-011-0025-4" volume="19" year="2011">Schneppat et al. (2011)</bibRefCitation>
<bibRefCitation id="5125C6EAB95FB99993EEC2465BFAC94E" DOI="https://doi.org/10.2478/v10125-011-0025-4" author="Schneppat, UE" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="149" pagination="225 - 248" refId="B34" refString="Schneppat, UE, Georgiev, DG, Dedov, IK, Wondrak, G, Knechtle, F, 2011. Tandonia totevi (Wiktor, 1975) (Pulmonata: Milacidae) in Bulgaria and north eastern Greece. Folia Malacologica 19 (4): 225 - 248, DOI: https://doi.org/10.2478/v10125-011-0025-4" title="Tandonia totevi (Wiktor, 1975) (Pulmonata: Milacidae) in Bulgaria and north eastern Greece." url="https://doi.org/10.2478/v10125-011-0025-4" volume="19" year="2011">Schneppat et al. (2011)</bibRefCitation>
for
<taxonomicName baseAuthorityName="Wiktor" baseAuthorityYear="1975" class="Gastropoda" family="Milacidae" genus="Tandonia" kingdom="Animalia" lsidName="Tandonia totevi" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="totevi">
<emphasis italics="true" pageId="0" pageNumber="149">Tandonia totevi</emphasis>
<taxonomicName id="E6E4E319C5DACB7C2AA9E4B639D21EC6" baseAuthorityName="Wiktor" baseAuthorityYear="1975" class="Gastropoda" family="Milacidae" genus="Tandonia" kingdom="Animalia" lsidName="Tandonia totevi" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="totevi">
<emphasis id="C7053629B540BD5E3A1B957B67EB9071" italics="true" pageId="0" pageNumber="149">Tandonia totevi</emphasis>
</taxonomicName>
(
<bibRefCitation author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="77 - 91" refId="B40" refString="Wiktor, A, 1975. New slug species (Pulmonata: Milacidae and Limacidae) from the Balkan Peninsula. Annales Zoologici 33: 77 - 91" title="New slug species (Pulmonata: Milacidae and Limacidae) from the Balkan Peninsula." volume="33" year="1975">Wiktor 1975</bibRefCitation>
<bibRefCitation id="DF4F6EF3350C1C16759EB3871F5C62E1" author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="77 - 91" refId="B40" refString="Wiktor, A, 1975. New slug species (Pulmonata: Milacidae and Limacidae) from the Balkan Peninsula. Annales Zoologici 33: 77 - 91" title="New slug species (Pulmonata: Milacidae and Limacidae) from the Balkan Peninsula." volume="33" year="1975">Wiktor 1975</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">
<emphasis italics="true" pageId="0" pageNumber="149">Integument structures</emphasis>
<paragraph id="2EF66C34B0A2C7E4DD4AC11386E3B3DB" pageId="0" pageNumber="149">
<emphasis id="FFDD72BBBC6C9AB821494B37695613F8" bold="true" pageId="0" pageNumber="149">
<emphasis id="75126D3A071B1D11C3C2D623BB5DB6D7" italics="true" pageId="0" pageNumber="149">Integument structures</emphasis>
.
</emphasis>
The number of tr (
<emphasis italics="true" pageId="0" pageNumber="149">n</emphasis>
<emphasis id="CE7BCD160179BB433324E01E55DE3B2A" italics="true" pageId="0" pageNumber="149">n</emphasis>
= 52; tr 12/13-tr 19,
<normalizedToken originalValue="ø">o</normalizedToken>
<normalizedToken id="0770998F785E714E4385E6F5253B3DEB" originalValue="ø">o</normalizedToken>
tr 15) does not vary much in our specimens, and there is no significant variation between populations. The surface texture and the width of tubercles in live specimens vary from fully straight to torn, depending on their body position. All tr from head to the 7th-9th row posterior to the pneumostome are entirely flat, smooth, wide, and remain so from the head and flank to the peripodial tubercle. The remaining tr reaching to the keel are somewhat crenulated, but are also wide, flat, and lacking ridges. Crenulations can only be seen under magnification. The tubercle rows may be long and undivided down to the peripodial tubercle, especially the ones below the lateral mantle edge. The more dorsal rows usually are divided in several tubercle compartments.
</paragraph>
<paragraph pageId="0" pageNumber="149">The keel is extended from the posterior margin of the mantle to the end of the dorsum. In some specimens, the keel was observed to be even slightly extending over the fringe of the sole, like a very little terminal thorn or knob. The keel has always an entirely smooth surface structure and is therefore clearly discernible from the crenulated neighbouring dorsal tubercles. Live and preserved specimens do not differ in the extension of the keel and its structure.</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">
<emphasis italics="true" pageId="0" pageNumber="149">Sole structure</emphasis>
<paragraph id="6B6121E468073F5B225EA5BC8141C674" pageId="0" pageNumber="149">The keel is extended from the posterior margin of the mantle to the end of the dorsum. In some specimens, the keel was observed to be even slightly extending over the fringe of the sole, like a very little terminal thorn or knob. The keel has always an entirely smooth surface structure and is therefore clearly discernible from the crenulated neighbouring dorsal tubercles. Live and preserved specimens do not differ in the extension of the keel and its structure.</paragraph>
<paragraph id="E2B16E85A4B4E5040E1EBD7EE9552827" pageId="0" pageNumber="149">
<emphasis id="6758A2F97E1BA84EE21902110CF1824B" bold="true" pageId="0" pageNumber="149">
<emphasis id="5264A446FEFADC97489E28ACC5ADE5DE" italics="true" pageId="0" pageNumber="149">Sole structure</emphasis>
.
</emphasis>
The outermost edge or seam of the sole is separated from the dorsum by a longitudinal fold, the peripodial tubercle, which begins left and right of the mouth-flaps and runs posterior around the body. The peripodial tubercle together with the peripodial groove clearly separate the seam of the sole at its outermost posterior end, where the sole is rounded and not pointed. In the central field of the sole, many V-shaped transverse wrinkles exist, which are invisible in animal crawling on a pane of glass but are well visible in preserved specimens.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">
<emphasis italics="true" pageId="0" pageNumber="149">Mucus</emphasis>
<paragraph id="77D656205DF5CCFE6E11CF53F54B09A8" pageId="0" pageNumber="149">
<emphasis id="DE4D2C2CF2C02419E901C505BFB1C234" bold="true" pageId="0" pageNumber="149">
<emphasis id="8CF4E300F3441CDC06F9CC10F97D2A09" italics="true" pageId="0" pageNumber="149">Mucus</emphasis>
.
</emphasis>
The mucus lacks any pigmentation on body, mantle and sole and is extremely sticky. When irritated, some of the animals produced defensive mucus of white to yellowish greenish colour on dorsum, flanks, and mantle, but this could only rarely be observed.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">
<emphasis italics="true" pageId="0" pageNumber="149">Measurements</emphasis>
<paragraph id="66612350C86A0AE0F5461D0A9CA77986" pageId="0" pageNumber="149">
<emphasis id="7248FBCB47869C891C437F4225AE65D7" bold="true" pageId="0" pageNumber="149">
<emphasis id="8B3150B302AC55AE8AF3E196B5427EEB" italics="true" pageId="0" pageNumber="149">Measurements</emphasis>
.
</emphasis>
lw (
<emphasis italics="true" pageId="0" pageNumber="149">n</emphasis>
<emphasis id="209B9A2A2F5125B97FF8FCFDB3796C72" italics="true" pageId="0" pageNumber="149">n</emphasis>
= 56): 1.5-6 g,
<normalizedToken originalValue="ø">o</normalizedToken>
<normalizedToken id="1C2B9A7C0D67C2A1D14C8B9280F1D0FD" originalValue="ø">o</normalizedToken>
3 g; tl (
<emphasis italics="true" pageId="0" pageNumber="149">n</emphasis>
<emphasis id="A1368CB85807A877F428B0437AE81A3A" italics="true" pageId="0" pageNumber="149">n</emphasis>
= 56): 52-92 mm,
<normalizedToken originalValue="ø">o</normalizedToken>
<normalizedToken id="28E4130A437A6FBD0A0FFFC6B2FCB23B" originalValue="ø">o</normalizedToken>
69 mm; ml (
<emphasis italics="true" pageId="0" pageNumber="149">n</emphasis>
<emphasis id="2C16918F2B3F950A8A4E72BC4342DC77" italics="true" pageId="0" pageNumber="149">n</emphasis>
= 50): 18-33 mm,
<normalizedToken originalValue="ø">o</normalizedToken>
<normalizedToken id="1EC18C6240232152CB46C568C9989063" originalValue="ø">o</normalizedToken>
23 mm; sw (
<emphasis italics="true" pageId="0" pageNumber="149">n</emphasis>
<emphasis id="5B1B46893E561A9D6EEB71EB1C4D54D2" italics="true" pageId="0" pageNumber="149">n</emphasis>
= 47): 5-10 mm,
<normalizedToken originalValue="ø">o</normalizedToken>
<normalizedToken id="DFD647E62434829CEC05F798A85F75C9" originalValue="ø">o</normalizedToken>
7.3 mm. Ratio of tl/ml ranges from ca. 52/18 mm to ca. 92/33 mm;
<normalizedToken originalValue="ø">o</normalizedToken>
<normalizedToken id="013A551646007D88DF616D5BBD6B13D5" originalValue="ø">o</normalizedToken>
ml is 1/3 of tl.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">
<emphasis italics="true" pageId="0" pageNumber="149">Genital organs</emphasis>
<paragraph id="53E197EB895F6EED81E9AC5F4B556949" pageId="0" pageNumber="149">
<emphasis id="CFF6685FB56D9EB69BA7A252C662EE2F" bold="true" pageId="0" pageNumber="149">
<emphasis id="783525C41447C48EDDD302BDD9E4BCE9" italics="true" pageId="0" pageNumber="149">Genital organs</emphasis>
.
</emphasis>
Atrium very short and tubular; penis short, with a distal bulb harbouring the penial papilla and a second bulb consisting of the papilla basis marking the boundary to the epiphallus, interior penial walls simple; penis papilla ornamented, apex of papilla with a row of curved crests encircling the complete papilla giving it a flower like appearance (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">3C, F</figureCitation>
<figureCitation id="C2731BAB3DD526D8C7DF11F13E97D134" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">3C, F</figureCitation>
); penis retractor muscle inserting at the epiphallus/penis boundary; epiphallus externally with smooth surface, consistent in diameter, reaching up to 4 times the length of the penis.
</paragraph>
<caption doi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" pageId="0" pageNumber="149">Figure 3.</emphasis>
<taxonomicName baseAuthorityName="Millet" baseAuthorityYear="1843" class="Gastropoda" family="Milacidae" genus="Tandonia" kingdom="Animalia" lsidName="Tandonia rustica." order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica.">
<emphasis italics="true" pageId="0" pageNumber="149">Tandonia rustica.</emphasis>
<caption id="4DF72E25288EAE638A10F2E7D375C34B" doi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149" start="Figure 3" startId="F3">
<paragraph id="7BE7CEB5DCAE61BC362964E906D0AF03" pageId="0" pageNumber="149">
<emphasis id="D65CF34EEAA48CDBF87F6D0EB24FAA4D" bold="true" pageId="0" pageNumber="149">Figure 3.</emphasis>
<taxonomicName id="DB01ED5AE357E1D634EDB1FB6D58591D" baseAuthorityName="Millet" baseAuthorityYear="1843" class="Gastropoda" family="Milacidae" genus="Tandonia" kingdom="Animalia" lsidName="Tandonia rustica." order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica.">
<emphasis id="5EB418C2ACF5925963D0420BE0844636" italics="true" pageId="0" pageNumber="149">Tandonia rustica.</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="149">A-C</emphasis>
<emphasis id="99D3914975AF8B17DBD2D751592C69E9" bold="true" pageId="0" pageNumber="149">A-C</emphasis>
sequenced specimen NMBE 568973, Rovio, Sovaglia
<emphasis bold="true" pageId="0" pageNumber="149">A</emphasis>
<emphasis id="8765516C76078B8EEC849623895FBF6B" bold="true" pageId="0" pageNumber="149">A</emphasis>
preserved animal
<emphasis bold="true" pageId="0" pageNumber="149">B</emphasis>
<emphasis id="A6B8564786589C0A027ADC6FF2B9A76F" bold="true" pageId="0" pageNumber="149">B</emphasis>
situs of genitalia
<emphasis bold="true" pageId="0" pageNumber="149">C</emphasis>
<emphasis id="B551E10E641C6491A81714CDF9E2B503" bold="true" pageId="0" pageNumber="149">C</emphasis>
distal genitalia showing anatomical details
<emphasis bold="true" pageId="0" pageNumber="149">D</emphasis>
<emphasis id="4AAF6CAFAFA3E7F7911AB4D7BF7DE65A" bold="true" pageId="0" pageNumber="149">D</emphasis>
penis papillae of sequenced specimen NMBE 568978, Carona
<emphasis bold="true" pageId="0" pageNumber="149">E-G</emphasis>
<emphasis id="EED0D2A030BAF30575AA6AA3DEB5366B" bold="true" pageId="0" pageNumber="149">E-G</emphasis>
sequenced specimen NMBE 564708, Goldswil
<emphasis bold="true" pageId="0" pageNumber="149">E</emphasis>
<emphasis id="6C8DF6065361D2C6F7633A18694718D8" bold="true" pageId="0" pageNumber="149">E</emphasis>
situs of genitalia
<emphasis bold="true" pageId="0" pageNumber="149">F</emphasis>
<emphasis id="AFC537C7C79EFA4EB4C914DD81855DA6" bold="true" pageId="0" pageNumber="149">F</emphasis>
distorted epiphallus
<emphasis bold="true" pageId="0" pageNumber="149">G</emphasis>
<emphasis id="D277E874D4951CBAF6FF4DE8AC4B5AA7" bold="true" pageId="0" pageNumber="149">G</emphasis>
details of lumina of vagina, atrium and penis, arrows pointing at the &quot;twin papillae&quot; (tp)
<emphasis bold="true" pageId="0" pageNumber="149">H-J</emphasis>
<emphasis id="7734E732FC75646254C9A7EE95E76098" bold="true" pageId="0" pageNumber="149">H-J</emphasis>
spermathophore of specimen NMBE 571414, Orselina
<emphasis bold="true" pageId="0" pageNumber="149">H</emphasis>
<emphasis id="32A39EF003DADC5D3D19A4338E5C724D" bold="true" pageId="0" pageNumber="149">H</emphasis>
genital pore with piece of spermatophore
<emphasis bold="true" pageId="0" pageNumber="149">I, J</emphasis>
<emphasis id="90E19DA14CB4D5DA9AC325847298EC81" bold="true" pageId="0" pageNumber="149">I, J</emphasis>
largest part of the three pieces
<emphasis bold="true" pageId="0" pageNumber="149">J</emphasis>
<emphasis id="37121353B9E30EF5AA11714F11734D10" bold="true" pageId="0" pageNumber="149">J</emphasis>
detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot;twin papillae&quot;; v = vagina; vd = vas deferens; vg = vaginal gland.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="149">Vagina twice the length of the penis, separated from the atrium by a sphincter; accessory glands entering close to the boundary of atrium and vagina; accessory glands digitiform or sac-like, either beige, brown or bright rusty red coloured; vaginal lumen with elongated, waved folds pointing towards the oviduct and the pedunculus of the bursa copulatrix; pedunculus somewhat longer than the vesicle; vesicle may be pointed or rounded.</paragraph>
<paragraph pageId="0" pageNumber="149">
<emphasis bold="true" italics="true" pageId="0" pageNumber="149">Spermatophore</emphasis>
<paragraph id="BC5FA144FDFEA502A84A172E5C9290BA" pageId="0" pageNumber="149">Vagina twice the length of the penis, separated from the atrium by a sphincter; accessory glands entering close to the boundary of atrium and vagina; accessory glands digitiform or sac-like, either beige, brown or bright rusty red coloured; vaginal lumen with elongated, waved folds pointing towards the oviduct and the pedunculus of the bursa copulatrix; pedunculus somewhat longer than the vesicle; vesicle may be pointed or rounded.</paragraph>
<paragraph id="193503544D0C68773C744D53DE90AEB6" pageId="0" pageNumber="149">
<emphasis id="F6914887E61EFCF157D7DEA700F691B2" bold="true" italics="true" pageId="0" pageNumber="149">Spermatophore</emphasis>
. A spermatophore was found in a single specimen, broken into three parts (NMBE 571414, Ticino, Orselina, 14 February 2020). One part still stuck in the lumen of the genital pore of the slug, length 0.2 cm; it is completely covered with simple spines (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">3H</figureCitation>
<figureCitation id="EAA5053F699835A843DDECC37AC4BD39" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">3H</figureCitation>
). The other two parts were found embedded in the vesicle in a white, fibrous mass; digestion had already started. The smaller part is 0.35 cm long, curved and covered with undivided and, further on, bifurcated spines laterally. The largest part of the spermatophore is ~ 0.9 cm long and curled (visualized in Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">3I, J</figureCitation>
<figureCitation id="5F3DB24D3F7615D6426F15473187D715" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">3I, J</figureCitation>
). Its outer surface is covered on the entire length with bifurcated spines. This is the first record of the spermatophore of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="distribution">
<paragraph pageId="0" pageNumber="149">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="149">
<subSubSection id="CF7DD53C7DD0AA2E0DA3D00381731901" pageId="0" pageNumber="149" type="distribution">
<paragraph id="6F89B91D20F41FE75D1E1E4266F18C44" pageId="0" pageNumber="149">Distribution.</paragraph>
<paragraph id="B68E393BBBBD95B338C14BC89D50846F" pageId="0" pageNumber="149">
Our own distribution records are mainly limited to Switzerland, France, northern Italy, and the Republic of San Marino. The species itself is frequently recorded from the western alpine arc, but also from a wide range in Great Britain, Ireland, The Netherlands, Belgium, Luxemburg, France, Germany, Austria, Poland, Czech Republic, Slovakia, and Hungary (
<bibRefCitation author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">Wiktor 1987</bibRefCitation>
<bibRefCitation id="E55611621CC029CE07AE3AE4B58B7F1C" author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">Wiktor 1987</bibRefCitation>
;
<bibRefCitation author="Gavetti, E" journalOrPublisher="Monografie Museo Regionale di Scienze Naturali" pageId="0" pageNumber="149" pagination="1 - 273" refId="B9" refString="Gavetti, E, Birindelli, S, Bodon, M, Manganelli, G, 2008. Molluschi terrestri e d'acqua dolce della Valle di Susa. Monografie Museo Regionale di Scienze Naturali 44: 1 - 273" title="Molluschi terrestri e d'acqua dolce della Valle di Susa." volume="44" year="2008">Gavetti et al. 2008</bibRefCitation>
<bibRefCitation id="91351342D036C0BC305CE9307EB9A6AE" author="Gavetti, E" journalOrPublisher="Monografie Museo Regionale di Scienze Naturali" pageId="0" pageNumber="149" pagination="1 - 273" refId="B9" refString="Gavetti, E, Birindelli, S, Bodon, M, Manganelli, G, 2008. Molluschi terrestri e d'acqua dolce della Valle di Susa. Monografie Museo Regionale di Scienze Naturali 44: 1 - 273" title="Molluschi terrestri e d'acqua dolce della Valle di Susa." volume="44" year="2008">Gavetti et al. 2008</bibRefCitation>
;
<bibRefCitation author="Rowson, B" journalOrPublisher="Bioinformatics" pageId="0" pageNumber="149" refId="B29" refString="Rowson, B, 2017. Tandonia rustica. The IUCN Red List of Threatened Species 2017: e.T171361A1325060." title="Tandonia rustica. The IUCN Red List of Threatened Species 2017: e. T 171361 A 1325060." year="2017">Rowson 2017</bibRefCitation>
<bibRefCitation id="4A96D9BDA2CA0D5E69863224AE0119D8" author="Rowson, B" journalOrPublisher="Bioinformatics" pageId="0" pageNumber="149" refId="B29" refString="Rowson, B, 2017. Tandonia rustica. The IUCN Red List of Threatened Species 2017: e.T171361A1325060." title="Tandonia rustica. The IUCN Red List of Threatened Species 2017: e. T 171361 A 1325060." year="2017">Rowson 2017</bibRefCitation>
). Reports for Romania are likely misidentifications with
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. kusceri" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="kusceri">
<emphasis italics="true" pageId="0" pageNumber="149">T. kusceri</emphasis>
<taxonomicName id="6DA9CF27B2BCE8B0C50D16BEC025F2BD" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. kusceri" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="kusceri">
<emphasis id="717C0D5BD6D5074C4B10B293D52BC1FC" italics="true" pageId="0" pageNumber="149">T. kusceri</emphasis>
</taxonomicName>
. Old reports for Bulgaria are shown to be wrong identifications of the very common
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. kusceri" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="kusceri">
<emphasis italics="true" pageId="0" pageNumber="149">T. kusceri</emphasis>
<taxonomicName id="18D64EDEBB5B0F4255245EF4F3BA94BD" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. kusceri" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="kusceri">
<emphasis id="BB555D3085F6549AD408D0354BC73513" italics="true" pageId="0" pageNumber="149">T. kusceri</emphasis>
</taxonomicName>
(pers. obs.).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="habitat">
<paragraph pageId="0" pageNumber="149">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="149">This species is confined to beech forests on limestone talus.</paragraph>
<subSubSection id="5130CA1FE49615F5915543022C6AB7D0" pageId="0" pageNumber="149" type="habitat">
<paragraph id="9FE28D1DC1A9ED2BE2D573EA3A21EE45" pageId="0" pageNumber="149">Habitat.</paragraph>
<paragraph id="67A77EC9E047D65449632180773668F5" pageId="0" pageNumber="149">This species is confined to beech forests on limestone talus.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="149" type="remarks">
<paragraph pageId="0" pageNumber="149">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="149">
<subSubSection id="A549B66ABEBE30D45A34B557ABC9ABCB" pageId="0" pageNumber="149" type="remarks">
<paragraph id="78FC4EBF2B4DF916E15FC00E428A878B" pageId="0" pageNumber="149">Remarks.</paragraph>
<paragraph id="CB19FD6D0BC9A06CA4D5360E1441EF75" pageId="0" pageNumber="149">
As original name of this species,
<bibRefCitation author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">Wiktor (1987)</bibRefCitation>
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referenced a &quot;
<taxonomicName authorityName="Millet" authorityYear="1843" class="Gastropoda" family="Milacidae" genus="Limax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limax marginatus subsp. rusticus" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="subSpecies" species="marginatus" subSpecies="rusticus">
<emphasis italics="true" pageId="0" pageNumber="149">Limax marginatus rusticus</emphasis>
<taxonomicName id="8A2C2B3A618A8949EC8B23AECD271E25" authorityName="Millet" authorityYear="1843" class="Gastropoda" family="Milacidae" genus="Limax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limax marginatus subsp. rusticus" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="subSpecies" species="marginatus" subSpecies="rusticus">
<emphasis id="0762CD03781E82BCA96CBE485E5F179F" italics="true" pageId="0" pageNumber="149">Limax marginatus rusticus</emphasis>
</taxonomicName>
Millet, 1843&quot;, which is wrong. In his text, Millet states &quot;Je
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au nord de
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,
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la Bouillant, commune de la Chapelle-Hullin, ainsi
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, etc.&quot;. These localities can be found in the larger surrounding north of Angers, Region Pays de la Loire,
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Maine-et-Loire, thus we consider this area the &quot;type area&quot; of
<taxonomicName genus="L." lsidName="L. rusticus" pageId="0" pageNumber="149" rank="species" species="rusticus">
<emphasis italics="true" pageId="0" pageNumber="149">L. rusticus</emphasis>
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. A neotype should be selected from one of the mentioned places to unambiguously fix the use of this name.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<paragraph id="1D0E41AB31E56A273AB59669E4A3C52A" pageId="0" pageNumber="149">
When it comes to morphology
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mentioned 18-19 tr, which is higher than most of our counts, but still in the range of our data. Comparing the exterior of living and preserved specimens with the description by
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: 286) and
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, many differences were found. No specimens of generally &quot;white or white creamy&quot; pigmentation, nor those of &quot;pinkish or violetish&quot; hues could be recorded by us in Switzerland and bordering areas. Most specimens investigated are explicitly darker than described by
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and
<bibRefCitation author="Rowson, B" journalOrPublisher="Field Studies Council, Telford, UK" pageId="0" pageNumber="149" refId="B31" refString="Rowson, B, Turner, JA, Anderson, R, Symondson, WOC, 2014b. The Slugs of Britain and Ireland: Identification, understanding and control. Field Studies Council, Telford, UK" title="The Slugs of Britain and Ireland: Identification, understanding and control." year="2014 b">Rowson et al (2014b)</bibRefCitation>
<bibRefCitation id="A6522CC5BB8D2584E616C2D2488F8AE1" author="Rowson, B" journalOrPublisher="Field Studies Council, Telford, UK" pageId="0" pageNumber="149" refId="B31" refString="Rowson, B, Turner, JA, Anderson, R, Symondson, WOC, 2014b. The Slugs of Britain and Ireland: Identification, understanding and control. Field Studies Council, Telford, UK" title="The Slugs of Britain and Ireland: Identification, understanding and control." year="2014 b">Rowson et al (2014b)</bibRefCitation>
indicating a much higher intraspecific variability of the species in other areas of distribution. Colour morphs exclude each
<normalizedToken originalValue="others">other's</normalizedToken>
<normalizedToken id="F461AFA3ACD5B302E57DF4B665276274" originalValue="others">other's</normalizedToken>
following the geographical pattern as described above. In
<bibRefCitation author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">Wiktor (1987</bibRefCitation>
<bibRefCitation id="5148F4A47E83B209E903C4C11F0C24B9" author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">Wiktor (1987</bibRefCitation>
: figs 196, 197), the pneumostome is positioned on the left side of the body, which most probably is an error or a sinistral specimen.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<paragraph id="321D006AA20D120E24A138DFDF1E585C" pageId="0" pageNumber="149">
Regarding the genital anatomy we found genetically identified
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. rustica" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="rustica">
<emphasis italics="true" pageId="0" pageNumber="149">T. rustica</emphasis>
<taxonomicName id="6450C80878E77A652DCF6FBAD3CFBDD4" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. rustica" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica">
<emphasis id="8FE77C0376A1D8ECFB3D597CDBF9280F" italics="true" pageId="0" pageNumber="149">T. rustica</emphasis>
</taxonomicName>
(NMBE 564708, NMBE 568978), which show a very similar anatomy to
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. kusceri" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="kusceri">
<emphasis italics="true" pageId="0" pageNumber="149">T. kusceri</emphasis>
<taxonomicName id="C2B515FD1C3F849B5693BDA59C05E955" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. kusceri" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="kusceri">
<emphasis id="592EC68C60697AD99ABFB6E50A6FD6DC" italics="true" pageId="0" pageNumber="149">T. kusceri</emphasis>
</taxonomicName>
with a long (in one specimen) coiled epiphallus and in both a round bursa copulatrix (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">3E, F</figureCitation>
<figureCitation id="B37C22A5F4BE83B22836A7E5F35A0744" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tandonia rustica. A-C sequenced specimen NMBE 568973, Rovio, Sovaglia A preserved animal B situs of genitalia C distal genitalia showing anatomical details D penis papillae of sequenced specimen NMBE 568978, Carona E-G sequenced specimen NMBE 564708, Goldswil E situs of genitalia F distorted epiphallus G details of lumina of vagina, atrium and penis, arrows pointing at the &quot; twin papillae &quot; (tp) H-J spermathophore of specimen NMBE 571414, Orselina H genital pore with piece of spermatophore I, J largest part of the three pieces J detail of surface covering spines. Abbreviations: a = atrium; bc = bursa copulatrix; ep = epiphallus; fod = free oviduct; p = penis; ped = pedunculus; pp = penial papilla; spo = spermoviduct; tp = &quot; twin papillae &quot;; v = vagina; vd = vas deferens; vg = vaginal gland." figureDoi="10.3897/zookeys.1116.82762.figure3" httpUri="https://binary.pensoft.net/fig/725585" pageId="0" pageNumber="149">3E, F</figureCitation>
). For the coiled epiphallus it might be a sexually active specimen, very probably after the ejection of the spermatophore. Even though the characters of
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. rustica" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="rustica">
<emphasis italics="true" pageId="0" pageNumber="149">T. rustica</emphasis>
<taxonomicName id="3F21536FE9CC43EC028A51C6734397EA" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. rustica" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica">
<emphasis id="FA7ABC8F449E5AFAB0F46281DC50F23B" italics="true" pageId="0" pageNumber="149">T. rustica</emphasis>
</taxonomicName>
and
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. kusceri" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="kusceri">
<emphasis italics="true" pageId="0" pageNumber="149">T. kusceri</emphasis>
<taxonomicName id="2937E9E8DD3327005F92B620923728D5" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. kusceri" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="kusceri">
<emphasis id="9F33C9AB4FF6623C49A58651D7B00CE1" italics="true" pageId="0" pageNumber="149">T. kusceri</emphasis>
</taxonomicName>
seem to meet, the two species can be distinguished by their habitat.
<taxonomicName baseAuthorityName="H. Wagner" baseAuthorityYear="1931" class="Gastropoda" family="Milacidae" genus="Tandonia" kingdom="Animalia" lsidName="Tandonia kusceri" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="kusceri">
<emphasis italics="true" pageId="0" pageNumber="149">Tandonia kusceri</emphasis>
<taxonomicName id="58D7DEC59BC5A640804AD72D36958073" baseAuthorityName="H. Wagner" baseAuthorityYear="1931" class="Gastropoda" family="Milacidae" genus="Tandonia" kingdom="Animalia" lsidName="Tandonia kusceri" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="kusceri">
<emphasis id="BAF9D1E4F21431729A09E9F8A59C36BC" italics="true" pageId="0" pageNumber="149">Tandonia kusceri</emphasis>
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usually is found in disturbed, synanthropic habitats, whereas
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. rustica" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="rustica">
<emphasis italics="true" pageId="0" pageNumber="149">T. rustica</emphasis>
<taxonomicName id="4B5F17843F0880E3FDEA3B495A103BE2" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. rustica" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica">
<emphasis id="27550817D0A1977745B996F5539BE0EA" italics="true" pageId="0" pageNumber="149">T. rustica</emphasis>
</taxonomicName>
is usually found in untouched more natural woodland habitats.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<bibRefCitation author="Wiktor, A" journalOrPublisher="Monografie Fauny Polski" pageId="0" pageNumber="149" pagination="1 - 182" refId="B39" refString="Wiktor, A, 1973. Die Nacktschnecken Polens. Monografie Fauny Polski 1: 1 - 182" title="Die Nacktschnecken Polens." volume="1" year="1973">Wiktor (1973)</bibRefCitation>
<paragraph id="A32ACE07A4174164650D9827DBE54BC7" pageId="0" pageNumber="149">
<bibRefCitation id="C98B569BE3555E1E9A1818C63CF6590A" author="Wiktor, A" journalOrPublisher="Monografie Fauny Polski" pageId="0" pageNumber="149" pagination="1 - 182" refId="B39" refString="Wiktor, A, 1973. Die Nacktschnecken Polens. Monografie Fauny Polski 1: 1 - 182" title="Die Nacktschnecken Polens." volume="1" year="1973">Wiktor (1973)</bibRefCitation>
and
<bibRefCitation author="Likharev, IM" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="149" refId="B20" refString="Likharev, IM, Wiktor, A, 1980. Slizni fauny SSSR i sopredelnych stran (Gastropoda terrestria nuda) [The Fauna of Slugs of the USSR and adjacent Countries (Gastropoda terrestria nuda)]. In: Fauna SSSR, Molljuski III, 5 [= N.S. 122] Leningrad (Nauka), 1-437. [pl. 1]" title="Slizni fauny SSSR i sopredelnych stran (Gastropoda terrestria nuda) [The Fauna of Slugs of the USSR and adjacent Countries (Gastropoda terrestria nuda)]. In: Fauna SSSR, Molljuski III, 5 [= N. S. 122] Leningrad (Nauka), 1 - 437. [pl. 1]" year="1980">Likharev and Wiktor (1980)</bibRefCitation>
<bibRefCitation id="78D818FB08E3F48575944DA97A193705" author="Likharev, IM" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="149" refId="B20" refString="Likharev, IM, Wiktor, A, 1980. Slizni fauny SSSR i sopredelnych stran (Gastropoda terrestria nuda) [The Fauna of Slugs of the USSR and adjacent Countries (Gastropoda terrestria nuda)]. In: Fauna SSSR, Molljuski III, 5 [= N.S. 122] Leningrad (Nauka), 1-437. [pl. 1]" title="Slizni fauny SSSR i sopredelnych stran (Gastropoda terrestria nuda) [The Fauna of Slugs of the USSR and adjacent Countries (Gastropoda terrestria nuda)]. In: Fauna SSSR, Molljuski III, 5 [= N. S. 122] Leningrad (Nauka), 1 - 437. [pl. 1]" year="1980">Likharev and Wiktor (1980)</bibRefCitation>
report a long stem of the penial papilla, an observation we could not reproduce. Later,
<bibRefCitation author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">Wiktor (1987)</bibRefCitation>
<bibRefCitation id="E941FA02C9350B2AAD0DBAE31994D3C1" author="Wiktor, A" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="149" pagination="153 - 319" refId="B41" refString="Wiktor, A, 1987. Milacidae (Gastropoda, Pulmonata) - systematic monograph. Annales Zoologici 41: 153 - 319" title="Milacidae (Gastropoda, Pulmonata) - systematic monograph." volume="41" year="1987">Wiktor (1987)</bibRefCitation>
reported a slightly reduced length, but even this reduction does not fit to our results. In all our dissected specimen, we had a bulky papilla tightly attached to the constriction of the epiphallus and therefore without a stem.
</paragraph>
<paragraph pageId="0" pageNumber="149">
<bibRefCitation DOI="https://doi.org/10.1007/BF01260999" author="Gerhardt, U" journalOrPublisher="Zeitschrift fuer Morphologie und Oekologie der Tiere" pageId="0" pageNumber="149" pagination="557 - 580" refId="B10" refString="Gerhardt, U, 1940. Neue biologische Nacktschneckenstudien. -. Zeitschrift fuer Morphologie und Oekologie der Tiere 36 (4): 557 - 580, DOI: https://doi.org/10.1007/BF01260999" title="Neue biologische Nacktschneckenstudien. -." url="https://doi.org/10.1007/BF01260999" volume="36" year="1940">Gerhardt (1940)</bibRefCitation>
<paragraph id="25A03E055DBC81FFDCAA3B8E70F3137C" pageId="0" pageNumber="149">
<bibRefCitation id="ED9E975CD31D50DB3DB04DF4CB0B036D" DOI="https://doi.org/10.1007/BF01260999" author="Gerhardt, U" journalOrPublisher="Zeitschrift fuer Morphologie und Oekologie der Tiere" pageId="0" pageNumber="149" pagination="557 - 580" refId="B10" refString="Gerhardt, U, 1940. Neue biologische Nacktschneckenstudien. -. Zeitschrift fuer Morphologie und Oekologie der Tiere 36 (4): 557 - 580, DOI: https://doi.org/10.1007/BF01260999" title="Neue biologische Nacktschneckenstudien. -." url="https://doi.org/10.1007/BF01260999" volume="36" year="1940">Gerhardt (1940)</bibRefCitation>
observed the copulation behaviour of the species in his laboratory; however, he did not illustrate the spermatophore. He observed the first copulation on 17 November 1938 followed by two copulations on the 4 December 1938. His specimens were collected near Porto
<normalizedToken originalValue="dIschia">d'Ischia</normalizedToken>
<normalizedToken id="D6AC0FA7BD56849C2F1385CEF2552888" originalValue="dIschia">d'Ischia</normalizedToken>
, Isola
<normalizedToken originalValue="dIschia">d'Ischia</normalizedToken>
<normalizedToken id="15459E4F11D79267A3AAA42FFB565DDB" originalValue="dIschia">d'Ischia</normalizedToken>
, Italy. It remains unclear whether these specimens were
<taxonomicName family="Milacidae" genus="T." kingdom="Animalia" lsidName="T. rustica" order="Stylommatophora" pageId="0" pageNumber="149" rank="species" species="rustica">
<emphasis italics="true" pageId="0" pageNumber="149">T. rustica</emphasis>
<taxonomicName id="F90D68C38DD8BA4A008A790BD27CC922" class="Gastropoda" family="Milacidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="T. rustica" order="Stylommatophora" pageId="0" pageNumber="149" phylum="Mollusca" rank="species" species="rustica">
<emphasis id="E5F033540ED8BBD0E86DA5F96C044DC6" italics="true" pageId="0" pageNumber="149">T. rustica</emphasis>
</taxonomicName>
; later research on the island yielded no specimens of this species. In case
<normalizedToken originalValue="Gerhards">Gerhard's</normalizedToken>
<normalizedToken id="809EFB401871EA02E884F22A79C561AE" originalValue="Gerhards">Gerhard's</normalizedToken>
determination was correct, the data of the observed copulation and those from our collection with the spermatophore found on 14 February 2020 indicate that this species probably mates during the winter season, a period where malacological research activity is rather low.
</paragraph>
</subSubSection>

View file

@ -1,379 +1,379 @@
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<mods:title id="27FC4CB97274354F95332AE68FD0DDC4">Journal of Hymenoptera Research</mods:title>
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<mods:date>2018</mods:date>
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<mods:number>2018-06-25</mods:number>
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<mods:number>64</mods:number>
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<treatment ID-GBIF-Taxon="182224078" LSID="urn:lsid:zoobank.org:act:68DF4AF7-FA6A-48A4-9305-CC8D6D4EECF7" httpUri="http://treatment.plazi.org/id/A1BDC0007ED7DC02DEBB69D6C8BD3E63" lastPageId="20" lastPageNumber="161" pageId="15" pageNumber="156">
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<paragraph pageId="15" pageNumber="156">
<taxonomicName LSID="http://zoobank.org/68DF4AF7-FA6A-48A4-9305-CC8D6D4EECF7" authority="Ridenbaugh" authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="15" pageNumber="156" phylum="Arthropoda" rank="species" species="spitzer" status="sp. n.">Eadya spitzer Ridenbaugh</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="156">sp. n.</taxonomicNameLabel>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">Figs 12A-C</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">; 13A-C</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">; 14A-E</figureCitation>
<treatment id="A1BDC0007ED7DC02DEBB69D6C8BD3E63" ID-GBIF-Taxon="182224078" LSID="urn:lsid:zoobank.org:act:68DF4AF7-FA6A-48A4-9305-CC8D6D4EECF7" httpUri="http://treatment.plazi.org/id/A1BDC0007ED7DC02DEBB69D6C8BD3E63" lastPageId="20" lastPageNumber="161" pageId="15" pageNumber="156" scope_family="Braconidae" scope_order="Hymenoptera">
<subSubSection id="DE17F6618E640C54314014D7F59971D7" pageId="15" pageNumber="156" type="nomenclature">
<paragraph id="FF72C0CCAC122C5AC0BB50FB7BFC7B78" pageId="15" pageNumber="156">
<taxonomicName id="1131F0126D1E8636D66B6F2F771CB2C4" LSID="http://zoobank.org/68DF4AF7-FA6A-48A4-9305-CC8D6D4EECF7" authority="Ridenbaugh" authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="15" pageNumber="156" phylum="Arthropoda" rank="species" species="spitzer" status="sp. n.">Eadya spitzer Ridenbaugh</taxonomicName>
<taxonomicNameLabel id="084D21A060990FFF2D884573E9CDDFDD" pageId="15" pageNumber="156">sp. n.</taxonomicNameLabel>
<figureCitation id="5F9BB50981D45E9CF0A635114CB2D494" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">Figs 12A-C</figureCitation>
<figureCitation id="6E252E922BBA356F48568D262679F616" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">; 13A-C</figureCitation>
<figureCitation id="ACEB8257FFB6B12B6B40DAFAF0EF2C4A" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">; 14A-E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="156" type="diagnosis">
<paragraph pageId="15" pageNumber="156">Diagnosis.</paragraph>
<paragraph pageId="15" pageNumber="156">
<taxonomicName authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="15" pageNumber="156" phylum="Arthropoda" rank="species" species="spitzer">
<emphasis italics="true" pageId="15" pageNumber="156">Eadya spitzer</emphasis>
<subSubSection id="7B743B74FB1F89BA2B048751EFB21CBE" pageId="15" pageNumber="156" type="diagnosis">
<paragraph id="DCD937581E3D2B3618B0E9E56DDAF33C" pageId="15" pageNumber="156">Diagnosis.</paragraph>
<paragraph id="7A8EFBB6F1472FA0167F1638BE0888AD" pageId="15" pageNumber="156">
<taxonomicName id="F6EE5D37097952F683B50DE355448837" authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="15" pageNumber="156" phylum="Arthropoda" rank="species" species="spitzer">
<emphasis id="840EF34FDED367C9EF36E8A6AA846244" italics="true" pageId="15" pageNumber="156">Eadya spitzer</emphasis>
</taxonomicName>
sp. n. can be distinguished from all other members of
<taxonomicName authorityName="Huddleston &amp; Short" authorityYear="1978" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya" order="Hymenoptera" pageId="15" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="15" pageNumber="156">Eadya</emphasis>
<taxonomicName id="8A0F1E71FE02BBCFA6405250A4B13E09" authorityName="Huddleston &amp; Short" authorityYear="1978" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya" order="Hymenoptera" pageId="15" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="19E369D14296D7FBD5862B5209F49B4D" italics="true" pageId="15" pageNumber="156">Eadya</emphasis>
</taxonomicName>
by the following combination of characters: Clypeus flanged at ventral margin, with two medial tubercles projecting outward (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A</figureCitation>
<figureCitation id="E4DD433F36FF110AB416BB08C3BC149B" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A</figureCitation>
); frons with inter-antennal and lateral carina flanged (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
<figureCitation id="3060F58C73D8BC409E0722B0A6E0A6AF" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
); occipital carina simple (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
<figureCitation id="036BFF07EDF29E11E999982531E4B77C" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
); occiput simple; notaulus impressed towards anterior margin of mesoscutum, foveate at apex (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
<figureCitation id="B0E7261C1452AE5AB8B0A8093F90EF97" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
); scutellar sulcus divided into many deep pits by ridge like longitudinal carinae (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
<figureCitation id="AB5DD5A58E1A9191F720B6B8FDC0FB1B" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
); sternaulus crenulate (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14D</figureCitation>
<figureCitation id="A3EF41EC8AFAA5EEA16272AEC06BF59C" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14D</figureCitation>
); propodeum rounded in appearance from lateral angle (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
<figureCitation id="6690700CCB60DC126BD278BBB2A81DD2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
), without transverse carinae (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14E</figureCitation>
<figureCitation id="FE1E19823781B2F60E43B6906AE38C7F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14E</figureCitation>
), and not creating a distinct posterior face when viewed laterally; propodeal spiracle circular; head orange except for antenna, apex of mandible, and ocellar triangle black, median of clypeus brown (Figs
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B</figureCitation>
<figureCitation id="8F5614FEFA68105660067B412991DF44" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B</figureCitation>
); prothorax orange (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A</figureCitation>
<figureCitation id="85DC44A8BCA5F7D320A3B09E541B656A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
<figureCitation id="22B40982C8435488C1ED6BA08FCE61C7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
<figureCitation id="37F8E294961A21C8C82DDD43BAFE624D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
); hindwing infuscate with dark brown veins except for anal, basal, subbasal, and anterior half of discal cells hyaline (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13C</figureCitation>
<figureCitation id="E1A72E0282C141435D82B084E4BB82EB" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13C</figureCitation>
); legs black except for fore coxa and trochanter orange (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
<figureCitation id="57CAF9482F8C35C985CF0D5F4DB8AEC7" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
); amino acid sequence (112-118) IRNFIGM (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Cytochrome c oxidase subunit 1 amino acid sequences from Peixoto et al. (2018). Boxes indicate diagnostic molecular characters. For each sequence a unique corresponding DNA voucher code is listed as BJS followed by a number." figureDoi="10.3897/jhr.64.24282.figure15" httpUri="https://binary.pensoft.net/fig/212231" pageId="15" pageNumber="156">15</figureCitation>
<figureCitation id="8A686D2A6272CB67F8B3CC046F479809" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Cytochrome c oxidase subunit 1 amino acid sequences from Peixoto et al. (2018). Boxes indicate diagnostic molecular characters. For each sequence a unique corresponding DNA voucher code is listed as BJS followed by a number." figureDoi="10.3897/jhr.64.24282.figure15" httpUri="https://binary.pensoft.net/fig/212231" pageId="15" pageNumber="156">15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="157" pageId="15" pageNumber="156" type="description">
<paragraph pageId="15" pageNumber="156">Description.</paragraph>
<paragraph pageId="15" pageNumber="156">Female. Body length without abdomen 3.30mm. Abdomen 2.86mm. Ovipositor 1.17mm.</paragraph>
<paragraph pageId="15" pageNumber="156">
<emphasis bold="true" pageId="15" pageNumber="156">Color.</emphasis>
<subSubSection id="13E0A9260DCF6E9B0773DEA8FF912BA5" lastPageId="16" lastPageNumber="157" pageId="15" pageNumber="156" type="description">
<paragraph id="32BED16E873F95CB1CDB9B9BEC61BBF3" pageId="15" pageNumber="156">Description.</paragraph>
<paragraph id="E826A9B3DB2C295155715C8A5031F9F9" pageId="15" pageNumber="156">Female. Body length without abdomen 3.30mm. Abdomen 2.86mm. Ovipositor 1.17mm.</paragraph>
<paragraph id="0C48564E98AD47B9667CDB22774E1AB0" pageId="15" pageNumber="156">
<emphasis id="269EC49267DB1D0F8ACCCE72254332BD" bold="true" pageId="15" pageNumber="156">Color.</emphasis>
Head orange except for antenna, apex of mandible, and ocellar triangle black (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A, B</figureCitation>
<figureCitation id="1BD3CA4A3AC4C54774368FBB6CDDE84B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A, B</figureCitation>
;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
<figureCitation id="4F9343B5EF925B2F5130F898977E1813" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
;
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B</figureCitation>
<figureCitation id="ACB15688C897667ACE0C3FAACA86EFBC" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B</figureCitation>
), median of clypeus brown; prothorax orange (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A, B</figureCitation>
<figureCitation id="225187BB354AD4B399B4542D54134D14" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A, B</figureCitation>
;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
<figureCitation id="8914813305768A4A5DE71C27CA63E6E1" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
;
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B, C</figureCitation>
<figureCitation id="8F653B8C461A62064B6004107380E133" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B, C</figureCitation>
); mesoscutum orange (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A, B</figureCitation>
<figureCitation id="406FBF8CB5BD226E525E8228BD1BF48B" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A, B</figureCitation>
;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
<figureCitation id="F55CDA1CF43A8A1EE07A8AE5E6BE3C34" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
;
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B</figureCitation>
<figureCitation id="04D9ECAB5048150284C5771562A315A2" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B</figureCitation>
); mesopleuron black except for anterior dorsal margin orange (Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
<figureCitation id="510C7B705B3185E92B9287AFDBE437BA" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
;
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14D</figureCitation>
<figureCitation id="E87339F839949100BB79067542E94E56" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14D</figureCitation>
); metathorax black (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12B</figureCitation>
<figureCitation id="4A591DCEFE1B3D0DBD57FDD1BB1CF365" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12B</figureCitation>
;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
<figureCitation id="83D66E593976B28D6179A596C768D471" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
;
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14D, E</figureCitation>
<figureCitation id="4688E021511A3901A429ED919D073086" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14D, E</figureCitation>
); forewing infuscate with dark brown veins except for anal, basal, and subbasal cells hyaline (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13C</figureCitation>
<figureCitation id="4E8089141C8114188C3E021916979FAB" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13C</figureCitation>
); hindwing infuscate with dark brown veins except for anal, basal, subbasal, and anterior half of discal cells hyaline (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13C</figureCitation>
<figureCitation id="E90BE296BEEFCB787E7B41FE5B844269" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13C</figureCitation>
); legs black except for fore coxa and trochanter orange (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A</figureCitation>
<figureCitation id="324C1D5DB20A295368D5E085F2400D23" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12A</figureCitation>
;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
<figureCitation id="ABDCC2697BADE51E18EA595D9DD5E02C" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A, B</figureCitation>
); abdomen black except for ovipositor orange (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12C</figureCitation>
<figureCitation id="22E2FBA497D480F86941EFBB19BE82E2" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156">12C</figureCitation>
;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
<figureCitation id="7851E5EAB59459AE1124ACF9812F6536" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="15" pageNumber="156">13A</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="1303478" doi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156" start="Figure 12" startId="F12">
<paragraph pageId="15" pageNumber="156">
<emphasis bold="true" pageId="15" pageNumber="156">Figure 12.</emphasis>
<taxonomicName authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="15" pageNumber="156" phylum="Arthropoda" rank="species" species="spitzer">
<emphasis italics="true" pageId="15" pageNumber="156">Eadya spitzer</emphasis>
<caption id="697F0DB0F5188E54368D5D6FBEAA4D34" ID-Zenodo-Dep="1303478" doi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="15" pageNumber="156" start="Figure 12" startId="F12">
<paragraph id="C23CFD91845D4A66CB21B66FF1194076" pageId="15" pageNumber="156">
<emphasis id="5B1D09B100D0BC135095FF876E0BC99E" bold="true" pageId="15" pageNumber="156">Figure 12.</emphasis>
<taxonomicName id="BA9AA3DDC39B37D2446F6A7B8BE5980E" authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="15" pageNumber="156" phylum="Arthropoda" rank="species" species="spitzer">
<emphasis id="2117E003BCFDCF6FA1ABA8746F98F8B3" italics="true" pageId="15" pageNumber="156">Eadya spitzer</emphasis>
</taxonomicName>
Ridenbaugh, sp. n. holotype.
<emphasis bold="true" pageId="15" pageNumber="156">A</emphasis>
<emphasis id="92479DDC1E7AEA07AC9C22FBA8E1D46C" bold="true" pageId="15" pageNumber="156">A</emphasis>
Lateral habitus
<emphasis bold="true" pageId="15" pageNumber="156">B</emphasis>
<emphasis id="477DC7762442E0017DEE1057633C7D6F" bold="true" pageId="15" pageNumber="156">B</emphasis>
Dorsal habitus
<emphasis bold="true" pageId="15" pageNumber="156">C</emphasis>
<emphasis id="237FCC06D87B8E51F0946B52296DE394" bold="true" pageId="15" pageNumber="156">C</emphasis>
Metasoma, lateral view. All scale bars are 1mm in length.
</paragraph>
</caption>
<paragraph pageId="15" pageNumber="156">
<emphasis bold="true" pageId="15" pageNumber="156">Head.</emphasis>
<paragraph id="E674946B9D8854E2FBEE5F6F76EEBC72" pageId="15" pageNumber="156">
<emphasis id="E5BFE846ECF6636700C09D907B324952" bold="true" pageId="15" pageNumber="156">Head.</emphasis>
Clypeus simple, smooth with scattered setae, flanged as ventral margin, with two medial tubercles projecting outward (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A</figureCitation>
<figureCitation id="0B504030BDA42E0F803BE52107FD35F3" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A</figureCitation>
); mandibles overlapping, dorsal tooth longer than ventral (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A</figureCitation>
<figureCitation id="531273B80F5C4DFF555337D947425DBA" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A</figureCitation>
); face finely punctate with associated setae (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A</figureCitation>
<figureCitation id="40F6D192488CA826E0006541AF6A2180" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A</figureCitation>
); frons rugose, inter-antennal and lateral carina flanged, starting at the toruli and reaching the ocellar triangle (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B</figureCitation>
<figureCitation id="E63EB7EC293DAE3F32375663E71B4571" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14A, B</figureCitation>
); vertex smooth with scattered setae (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
<figureCitation id="7108F1C7B46E6F24B6B3654F8BEF03C9" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
); occipital carina simple (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
<figureCitation id="ED43F5BF976AF4CAF9D0176319B51E52" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
), reaching the hypostomal carina; hypostomal carina strongly flanged, reaching the mandible and bending around to the mandibular condyle; occiput smooth, normal (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
<figureCitation id="D280A1BE44CB1F753522C47B821E3DF1" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
).
</paragraph>
<paragraph lastPageId="16" lastPageNumber="157" pageId="15" pageNumber="156">
<emphasis bold="true" pageId="15" pageNumber="156">Mesosoma.</emphasis>
<paragraph id="365A36612F3AC716FE86106F5EFADCAB" lastPageId="16" lastPageNumber="157" pageId="15" pageNumber="156">
<emphasis id="FF5424244E246CD303C29F9C0B694BEC" bold="true" pageId="15" pageNumber="156">Mesosoma.</emphasis>
Pronotum exposed in dorsal view (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B, C</figureCitation>
<figureCitation id="407CB3955CED30210C171A495A9D2600" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B, C</figureCitation>
); pronope absent, subpronope absent, smooth except for a faint crenulate line extending laterally and rugulose sculpturing along the lateral posterior margin (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
<figureCitation id="FA20B11D13590C07913B662215109EF6" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14B</figureCitation>
); mesoscutum with median mesonotal lobe smooth (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
<figureCitation id="D86D0C26EC3D9A6A21A8F82EDF58B672" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
); notaulus impressed towards anterior margin of mesoscutum, foveate at apex (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
<figureCitation id="2CCEC8ADEDA5BC6E663E44F1AFED411E" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
); scutellar sulcus divided into many deep pits by ridge like longitudinal carinae (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
<figureCitation id="4B47B9874C0B905315989F90838E1DB2" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14C</figureCitation>
); sternaulus crenulate (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14D</figureCitation>
<figureCitation id="A94838DDB0430E61579E4C3D9226AEE1" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="15" pageNumber="156">14D</figureCitation>
); propo
<pageBreakToken pageId="16" pageNumber="157" start="start">deum</pageBreakToken>
<pageBreakToken id="978E7BB65E387FC31C222540E04DE085" pageId="16" pageNumber="157" start="start">deum</pageBreakToken>
rugose and pubescent, rounded in appearance from lateral angle, without transverse carinae and not creating a distinct posterior face when viewed laterally (Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157">13A</figureCitation>
<figureCitation id="352C2C8C78AFC363AFADF54EABCB93E1" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157">13A</figureCitation>
;
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="16" pageNumber="157">14E</figureCitation>
<figureCitation id="45C3B0412495FE2A69B49518DBBD4BE3" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="16" pageNumber="157">14E</figureCitation>
); propodeal spiracle circular; coxa, trochanter, trochantellus, and femur covered in setae, tibia and tarsus pubescent; tarsal claws simple (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="16" pageNumber="157">12A</figureCitation>
<figureCitation id="E981D7F65C96E531A7DDED404F4757A4" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="16" pageNumber="157">12A</figureCitation>
;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157">13A, B</figureCitation>
<figureCitation id="D17EC991DC0749D16A1739109FE486A5" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157">13A, B</figureCitation>
).
</paragraph>
<paragraph pageId="16" pageNumber="157">
<emphasis bold="true" pageId="16" pageNumber="157">Forewing.</emphasis>
<paragraph id="6F7697F7A65FF097CBA149B4AEC7A774" pageId="16" pageNumber="157">
<emphasis id="CC0F5B5856C10582455014C71EE1AECB" bold="true" pageId="16" pageNumber="157">Forewing.</emphasis>
r-m curved slightly towards stigma before reaching the junction of 3RSa and 3RSb (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157">13C</figureCitation>
<figureCitation id="231E98F4915FB1271D9B8C75AF7A6A48" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157">13C</figureCitation>
).
</paragraph>
<paragraph pageId="16" pageNumber="157">
<emphasis bold="true" pageId="16" pageNumber="157">Hindwing.</emphasis>
<paragraph id="219B7144A6AD6B85AB539A8645790F51" pageId="16" pageNumber="157">
<emphasis id="A5365A81B6B1AABC18051B1153B9CBC9" bold="true" pageId="16" pageNumber="157">Hindwing.</emphasis>
R1a with three hamuli.
</paragraph>
<paragraph pageId="16" pageNumber="157">
<emphasis bold="true" pageId="16" pageNumber="157">Metasoma.</emphasis>
<paragraph id="29C424D568EF74E81F68DEB03D16BB9E" pageId="16" pageNumber="157">
<emphasis id="A4ECA4BE9BAD2DD0E1600F673D0F4B66" bold="true" pageId="16" pageNumber="157">Metasoma.</emphasis>
Metasomal tergite 1 petiolate, spiracle protruding as a tubercle at about the middle of the segment, dorsal and lateral surface punctate with associated setae (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="16" pageNumber="157">14E</figureCitation>
<figureCitation id="6CD2A7754398B22B3C65CEAB5E1E1D82" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="16" pageNumber="157">14E</figureCitation>
); ovipositor straight (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="16" pageNumber="157">12C</figureCitation>
<figureCitation id="EAA9C45C130873AF4E6E8629072CFC81" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Eadya spitzer Ridenbaugh, sp. n. holotype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure12" httpUri="https://binary.pensoft.net/fig/212228" pageId="16" pageNumber="157">12C</figureCitation>
;
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157">13A</figureCitation>
<figureCitation id="466035BD666DD22FAAA1303DB3BC42C8" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157">13A</figureCitation>
).
</paragraph>
<paragraph pageId="16" pageNumber="157">
<emphasis bold="true" pageId="16" pageNumber="157">Male.</emphasis>
<paragraph id="ECE3226561955F708A66D04D7CB864A6" pageId="16" pageNumber="157">
<emphasis id="CE91A8EC1FE31485C0AE6E5831FDF761" bold="true" pageId="16" pageNumber="157">Male.</emphasis>
Unknown.
</paragraph>
<caption ID-Zenodo-Dep="1303480" doi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157" start="Figure 13" startId="F13">
<paragraph pageId="16" pageNumber="157">
<emphasis bold="true" pageId="16" pageNumber="157">Figure 13.</emphasis>
<taxonomicName authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="16" pageNumber="157" phylum="Arthropoda" rank="species" species="spitzer">
<emphasis italics="true" pageId="16" pageNumber="157">Eadya spitzer</emphasis>
<caption id="B2BF9B9BD734240C07D9785DFF5EF920" ID-Zenodo-Dep="1303480" doi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="16" pageNumber="157" start="Figure 13" startId="F13">
<paragraph id="77F08CA857ABC0D17E722C7EE706A4E6" pageId="16" pageNumber="157">
<emphasis id="31DCC3E9D03035DB5170ED71F9B4DB34" bold="true" pageId="16" pageNumber="157">Figure 13.</emphasis>
<taxonomicName id="034724BA4646086198F6323D1B25E8E5" authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="16" pageNumber="157" phylum="Arthropoda" rank="species" species="spitzer">
<emphasis id="E6BA569BFC9916E31D6E38D972B8D207" italics="true" pageId="16" pageNumber="157">Eadya spitzer</emphasis>
</taxonomicName>
Ridenbaugh, sp. n. paratype.
<emphasis bold="true" pageId="16" pageNumber="157">A</emphasis>
<emphasis id="08969EACA3F5443E65E0C39FC2CC8230" bold="true" pageId="16" pageNumber="157">A</emphasis>
Lateral habitus
<emphasis bold="true" pageId="16" pageNumber="157">B</emphasis>
<emphasis id="1C327868217383D40C8079262701F308" bold="true" pageId="16" pageNumber="157">B</emphasis>
Dorsal habitus
<emphasis bold="true" pageId="16" pageNumber="157">C</emphasis>
<emphasis id="44994F4C2F7951656954E3915DD0F74E" bold="true" pageId="16" pageNumber="157">C</emphasis>
Metasoma, lateral view. All scale bars are 1mm in length.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="16" pageNumber="157" type="host">
<paragraph pageId="16" pageNumber="157">Host.</paragraph>
<paragraph pageId="16" pageNumber="157">
<taxonomicName authorityName="Stal" authorityYear="1860" class="Insecta" family="Chrysomelidae" genus="Paropsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paropsis charybdis" order="Coleoptera" pageId="16" pageNumber="157" phylum="Arthropoda" rank="species" species="charybdis">
<emphasis italics="true" pageId="16" pageNumber="157">Paropsis charybdis</emphasis>
<subSubSection id="2BBEF906B214AE51AC29D180D10C4862" pageId="16" pageNumber="157" type="host">
<paragraph id="CAC0328967A43C53384346E495F411F9" pageId="16" pageNumber="157">Host.</paragraph>
<paragraph id="07B19B55F1342731F28A9878B453C770" pageId="16" pageNumber="157">
<taxonomicName id="4FF210713EC5E98110624A21EDBF6343" authorityName="Stal" authorityYear="1860" class="Insecta" family="Chrysomelidae" genus="Paropsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paropsis charybdis" order="Coleoptera" pageId="16" pageNumber="157" phylum="Arthropoda" rank="species" species="charybdis">
<emphasis id="218E36B3CA247B214B9C4785C46DF80C" italics="true" pageId="16" pageNumber="157">Paropsis charybdis</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Selman" authorityYear="1983" class="Insecta" family="Chrysomelidae" genus="Paropsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paropsis aegrota subsp. elliotti" order="Coleoptera" pageId="16" pageNumber="157" phylum="Arthropoda" rank="subSpecies" species="aegrota" subSpecies="elliotti">
<emphasis italics="true" pageId="16" pageNumber="157">Paropsis aegrota elliotti</emphasis>
<taxonomicName id="2A2527F821F9F47BD0568C4273B673E5" authorityName="Selman" authorityYear="1983" class="Insecta" family="Chrysomelidae" genus="Paropsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paropsis aegrota subsp. elliotti" order="Coleoptera" pageId="16" pageNumber="157" phylum="Arthropoda" rank="subSpecies" species="aegrota" subSpecies="elliotti">
<emphasis id="BF1B4C4FECF51C267811971F3A652F41" italics="true" pageId="16" pageNumber="157">Paropsis aegrota elliotti</emphasis>
</taxonomicName>
Selman, 1983.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="157" type="variation">
<paragraph pageId="16" pageNumber="157">Variations.</paragraph>
<paragraph pageId="16" pageNumber="157">
<subSubSection id="81F88F27F90962AA59BA48CA8FD76937" pageId="16" pageNumber="157" type="variation">
<paragraph id="12476E8FD496CDDA3D7F55D40DCFD924" pageId="16" pageNumber="157">Variations.</paragraph>
<paragraph id="39FF1BC218028DF9EA8195DD1F15C5E1" pageId="16" pageNumber="157">
Paratype with clypeus orange (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="16" pageNumber="157">14A</figureCitation>
<figureCitation id="9C32D1FC2E3BE021186E347D4A1CF787" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="16" pageNumber="157">14A</figureCitation>
). This variation may be the result of the DNA extraction process of the Holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="158" type="description">
<paragraph pageId="17" pageNumber="158">
<pageBreakToken pageId="17" pageNumber="158" start="start">Diagnostic</pageBreakToken>
<subSubSection id="67159E3BD9619D2991B412326B5EDD8C" pageId="17" pageNumber="158" type="description">
<paragraph id="60C91C120A5D52FFB433F87FF0BF088A" pageId="17" pageNumber="158">
<pageBreakToken id="A24EE4DBCC5A55F6DEA06A0D7C5ADC97" pageId="17" pageNumber="158" start="start">Diagnostic</pageBreakToken>
molecular characters.
</paragraph>
<paragraph pageId="17" pageNumber="158">
<paragraph id="35D7436A099BAE77E0FE3BC34A3A0C0E" pageId="17" pageNumber="158">
(22-27) IWSGII; (32-34) SVL; (41-46) [M or K]LGRLL; (54) S; (67-73) IVIPIII; (81) I; (90) MM; (95-98) INNI; (104-109) PPSLIL; (112-118) IRNFIGM; (126) M; (133-139) NLRHRGI; (143-144) MS; (150) L; (157) I; (167-169) INI; (172-191) LGLNYDNISLLVWSVNITAI (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Cytochrome c oxidase subunit 1 amino acid sequences from Peixoto et al. (2018). Boxes indicate diagnostic molecular characters. For each sequence a unique corresponding DNA voucher code is listed as BJS followed by a number." figureDoi="10.3897/jhr.64.24282.figure15" httpUri="https://binary.pensoft.net/fig/212231" pageId="17" pageNumber="158">15</figureCitation>
<figureCitation id="81E4FDE75D7E38993EB016EEC6FAB00F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Cytochrome c oxidase subunit 1 amino acid sequences from Peixoto et al. (2018). Boxes indicate diagnostic molecular characters. For each sequence a unique corresponding DNA voucher code is listed as BJS followed by a number." figureDoi="10.3897/jhr.64.24282.figure15" httpUri="https://binary.pensoft.net/fig/212231" pageId="17" pageNumber="158">15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="159" type="distribution">
<paragraph pageId="18" pageNumber="159">
<pageBreakToken pageId="18" pageNumber="159" start="start">Distribution</pageBreakToken>
<subSubSection id="66E5EE8C1D9FA48891001FF5DF638137" pageId="18" pageNumber="159" type="distribution">
<paragraph id="3948B351426547D8A1294038B36BC83A" pageId="18" pageNumber="159">
<pageBreakToken id="4E30F3DADD7CD90BE131FD0681341B66" pageId="18" pageNumber="159" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="18" pageNumber="159">Tasmania.</paragraph>
<paragraph id="A53C9C225AF63FF9C32F58D5CEF6E344" pageId="18" pageNumber="159">Tasmania.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="159" type="etymology">
<paragraph pageId="18" pageNumber="159">Etymology.</paragraph>
<paragraph pageId="18" pageNumber="159">
<subSubSection id="5620F1B7E0249EE70ECCF72514F0EDCB" pageId="18" pageNumber="159" type="etymology">
<paragraph id="CCED9895B9559972BD079EC06ABB0CF4" pageId="18" pageNumber="159">Etymology.</paragraph>
<paragraph id="8C4C73D9EE0FA797129D412A05B8DA86" pageId="18" pageNumber="159">
This species is named in honor of Edwin Spitzer, the first
<normalizedToken originalValue="authors">author's</normalizedToken>
<normalizedToken id="1EE119D02931C697178CAAE27B22C779" originalValue="authors">author's</normalizedToken>
(RDR) late grandfather. This is a noun in apposition to the generic name in order to retain integrity of the surname Spitzer.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="160" type="remarks">
<paragraph pageId="19" pageNumber="160">
<pageBreakToken pageId="19" pageNumber="160" start="start">Remarks</pageBreakToken>
<subSubSection id="2114CDEAC177F485A99EE18EA857407D" pageId="19" pageNumber="160" type="remarks">
<paragraph id="2EF35AD51AF7E205DB0410727C825260" pageId="19" pageNumber="160">
<pageBreakToken id="021776E99155CA9D815A34EA356A595E" pageId="19" pageNumber="160" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph pageId="19" pageNumber="160">
<paragraph id="DE0A7ADF18C805BA5DCA8737BF6FB216" pageId="19" pageNumber="160">
The paratype is for this series is badly damaged, missing both antennae, all six legs, and the abdomen excluding metasomal tergite 1. However, the specimen was photographed before destruction and can be seen in Figures
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="19" pageNumber="160">13A-C</figureCitation>
<figureCitation id="56351E1722C9828D812473FA36FF4057" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Eadya spitzer Ridenbaugh, sp. n. paratype. A Lateral habitus B Dorsal habitus C Metasoma, lateral view. All scale bars are 1 mm in length." figureDoi="10.3897/jhr.64.24282.figure13" httpUri="https://binary.pensoft.net/fig/212229" pageId="19" pageNumber="160">13A-C</figureCitation>
and
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="19" pageNumber="160">14A-E</figureCitation>
<figureCitation id="58BEA6B754A58760FAAD1A575987CA3C" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Eadya spitzer Ridenbaugh, sp. n. paratype. A Head, frontal view B Head, dorsal view C Head and mesoscutum, dorsal view D Mesopleuron, lateral view E Propodeum, dorsal view." figureDoi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="19" pageNumber="160">14A-E</figureCitation>
. This species is referred to as
<taxonomicName authorityName="Huddleston &amp; Short" authorityYear="1978" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya" order="Hymenoptera" pageId="19" pageNumber="160" phylum="Arthropoda" rank="genus">Eadya</taxonomicName>
<taxonomicName id="38AD16A9D80A4496B6C4BACA25E28177" authorityName="Huddleston &amp; Short" authorityYear="1978" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya" order="Hymenoptera" pageId="19" pageNumber="160" phylum="Arthropoda" rank="genus">Eadya</taxonomicName>
sp.2 in
<bibRefCitation author="Peixoto, L" journalOrPublisher="Annu Rev Entomol" pageId="24" pageNumber="165" refId="B17" refString="Peixoto, L, Allen, GR, Ridenbaugh, RD, Quarrell, SR, Withers, TM, Sharanowski, BJ, 2018. When taxonomy and biological control researchers unite: species delimitation of Eadya parasitoids (Braconidae) and consequences for classical biological control of invasive paropsine pests of Eucalyptus. PLoS One, Accepted." title="When taxonomy and biological control researchers unite: species delimitation of Eadya parasitoids (Braconidae) and consequences for classical biological control of invasive paropsine pests of Eucalyptus. PLoS One, Accepted." year="2018">Peixoto et al. (2018)</bibRefCitation>
<bibRefCitation id="0AEF491E63D02B4F8B733A9A9CCCB833" author="Peixoto, L" journalOrPublisher="Annu Rev Entomol" pageId="24" pageNumber="165" refId="B17" refString="Peixoto, L, Allen, GR, Ridenbaugh, RD, Quarrell, SR, Withers, TM, Sharanowski, BJ, 2018. When taxonomy and biological control researchers unite: species delimitation of Eadya parasitoids (Braconidae) and consequences for classical biological control of invasive paropsine pests of Eucalyptus. PLoS One, Accepted." title="When taxonomy and biological control researchers unite: species delimitation of Eadya parasitoids (Braconidae) and consequences for classical biological control of invasive paropsine pests of Eucalyptus. PLoS One, Accepted." year="2018">Peixoto et al. (2018)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="161" pageId="19" pageNumber="160" type="type material">
<paragraph pageId="19" pageNumber="160">Type material.</paragraph>
<paragraph lastPageId="20" lastPageNumber="161" pageId="19" pageNumber="160">
<subSubSection id="1ABC84C7E07FED21A0733115C919AA93" lastPageId="20" lastPageNumber="161" pageId="19" pageNumber="160" type="type material">
<paragraph id="70D1A8963E18AE5465BE0E2C59807E9F" pageId="19" pageNumber="160">Type material.</paragraph>
<paragraph id="C99A06AEDA4D4B659090B3F2BC513607" lastPageId="20" lastPageNumber="161" pageId="19" pageNumber="160">
Holotype, Female (ANIC), &quot;The Lea, TAS, 11 Dec 2012, Emerged 26 Dec 2012, G.R. Allen, Field collected in
<taxonomicName genus="P." lsidName="P. charybdis" pageId="19" pageNumber="160" rank="species" species="charybdis">
<emphasis italics="true" pageId="19" pageNumber="160">P. charybdis</emphasis>
<taxonomicName id="75F00F15B6DF163B73C04D5E9C658A3B" lsidName="P. charybdis" pageId="19" pageNumber="160" rank="species" species="charybdis">
<emphasis id="00C7E5DD169063D3BB9FF2FEF1C1CCAE" italics="true" pageId="19" pageNumber="160">P. charybdis</emphasis>
</taxonomicName>
, E135&quot;, &quot;BJS 199&quot;, GenBank
<pageBreakToken pageId="20" pageNumber="161" start="start">accession</pageBreakToken>
<pageBreakToken id="4B2B3B66FD2B68C285EB162AB51FA920" pageId="20" pageNumber="161" start="start">accession</pageBreakToken>
numbers KX989902, and MH107810. Paratype, Female (ANIC), &quot;Runnymede Site #1, TAS, 13 Dec 2015,
<geoCoordinate degrees="42" direction="south" minutes="38" orientation="latitude" precision="1" seconds="11.1" value="-42.636417">42°38'11.1&quot;S</geoCoordinate>
<geoCoordinate id="D43410B705CE0C6A92392E679EAD3A3E" degrees="42" direction="south" minutes="38" orientation="latitude" precision="1" seconds="11.1" value="-42.636417">42°38'11.1&quot;S</geoCoordinate>
,
<geoCoordinate degrees="147" direction="east" minutes="33" orientation="longitude" precision="1" seconds="54.7" value="147.5652">147°33'54.7&quot;E</geoCoordinate>
<geoCoordinate id="0E5D993DA7D21C7CE95EA88B8F836CF3" degrees="147" direction="east" minutes="33" orientation="longitude" precision="1" seconds="54.7" value="147.5652">147°33'54.7&quot;E</geoCoordinate>
, Flying adult, D. Satchell, Female&quot;.
</paragraph>
<caption ID-Zenodo-Dep="1303482" doi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="20" pageNumber="161" start="Figure 14" startId="F14">
<paragraph pageId="20" pageNumber="161">
<emphasis bold="true" pageId="20" pageNumber="161">Figure 14.</emphasis>
<taxonomicName authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="20" pageNumber="161" phylum="Arthropoda" rank="species" species="spitzer">
<emphasis italics="true" pageId="20" pageNumber="161">Eadya spitzer</emphasis>
<caption id="21DF6EDEE099D8E4B502E3B93360DC1B" ID-Zenodo-Dep="1303482" doi="10.3897/jhr.64.24282.figure14" httpUri="https://binary.pensoft.net/fig/212230" pageId="20" pageNumber="161" start="Figure 14" startId="F14">
<paragraph id="6DB978AFF9CC0EC5AE60A9B54B1ABB5D" pageId="20" pageNumber="161">
<emphasis id="E724C3CB35E7C742009E7E06778E7C0E" bold="true" pageId="20" pageNumber="161">Figure 14.</emphasis>
<taxonomicName id="70BBACEB3F3D653A7F0BC444012DEE62" authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya spitzer" order="Hymenoptera" pageId="20" pageNumber="161" phylum="Arthropoda" rank="species" species="spitzer">
<emphasis id="5FB81112EE3F49BA49A92A79C4795C6A" italics="true" pageId="20" pageNumber="161">Eadya spitzer</emphasis>
</taxonomicName>
Ridenbaugh, sp. n. paratype.
<emphasis bold="true" pageId="20" pageNumber="161">A</emphasis>
<emphasis id="88CF006D116718D19064A4D4CB62905F" bold="true" pageId="20" pageNumber="161">A</emphasis>
Head, frontal view
<emphasis bold="true" pageId="20" pageNumber="161">B</emphasis>
<emphasis id="C32DBF0E4DA306AE3AED35F6D0DB0046" bold="true" pageId="20" pageNumber="161">B</emphasis>
Head, dorsal view
<emphasis bold="true" pageId="20" pageNumber="161">C</emphasis>
<emphasis id="E93B6431D6D4C9F0420A6BF350652AEC" bold="true" pageId="20" pageNumber="161">C</emphasis>
Head and mesoscutum, dorsal view
<emphasis bold="true" pageId="20" pageNumber="161">D</emphasis>
<emphasis id="ED2E20118A92401B864EB3461F069D59" bold="true" pageId="20" pageNumber="161">D</emphasis>
Mesopleuron, lateral view
<emphasis bold="true" pageId="20" pageNumber="161">E</emphasis>
<emphasis id="DB0A167032E307D33205401CBE0898F3" bold="true" pageId="20" pageNumber="161">E</emphasis>
Propodeum, dorsal view.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="1303484" doi="10.3897/jhr.64.24282.figure15" httpUri="https://binary.pensoft.net/fig/212231" pageId="20" pageNumber="161" start="Figure 15" startId="F15">
<paragraph pageId="20" pageNumber="161">
<emphasis bold="true" pageId="20" pageNumber="161">Figure 15.</emphasis>
<caption id="4DF4D7C689F5AC98F42AED6D0519F26C" ID-Zenodo-Dep="1303484" doi="10.3897/jhr.64.24282.figure15" httpUri="https://binary.pensoft.net/fig/212231" pageId="20" pageNumber="161" start="Figure 15" startId="F15">
<paragraph id="17567322C2327F091B7CA1E547AC53F4" pageId="20" pageNumber="161">
<emphasis id="C497F62079CA3CD97D4ABF949586FF08" bold="true" pageId="20" pageNumber="161">Figure 15.</emphasis>
Cytochrome c oxidase subunit 1 amino acid sequences from
<bibRefCitation author="Peixoto, L" journalOrPublisher="Annu Rev Entomol" pageId="24" pageNumber="165" refId="B17" refString="Peixoto, L, Allen, GR, Ridenbaugh, RD, Quarrell, SR, Withers, TM, Sharanowski, BJ, 2018. When taxonomy and biological control researchers unite: species delimitation of Eadya parasitoids (Braconidae) and consequences for classical biological control of invasive paropsine pests of Eucalyptus. PLoS One, Accepted." title="When taxonomy and biological control researchers unite: species delimitation of Eadya parasitoids (Braconidae) and consequences for classical biological control of invasive paropsine pests of Eucalyptus. PLoS One, Accepted." year="2018">Peixoto et al. (2018)</bibRefCitation>
<bibRefCitation id="009551F3AB0F521BBAAEB624E8AA1819" author="Peixoto, L" journalOrPublisher="Annu Rev Entomol" pageId="24" pageNumber="165" refId="B17" refString="Peixoto, L, Allen, GR, Ridenbaugh, RD, Quarrell, SR, Withers, TM, Sharanowski, BJ, 2018. When taxonomy and biological control researchers unite: species delimitation of Eadya parasitoids (Braconidae) and consequences for classical biological control of invasive paropsine pests of Eucalyptus. PLoS One, Accepted." title="When taxonomy and biological control researchers unite: species delimitation of Eadya parasitoids (Braconidae) and consequences for classical biological control of invasive paropsine pests of Eucalyptus. PLoS One, Accepted." year="2018">Peixoto et al. (2018)</bibRefCitation>
. Boxes indicate diagnostic molecular characters. For each sequence a unique corresponding DNA voucher code is listed as BJS followed by a number.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="1303486" doi="10.3897/jhr.64.24282.figure16" httpUri="https://binary.pensoft.net/fig/212232" pageId="20" pageNumber="161" start="Figure 16" startId="F16">
<paragraph pageId="20" pageNumber="161">
<emphasis bold="true" pageId="20" pageNumber="161">Figure 16.</emphasis>
<caption id="B1B551D9BD090A642C364C2C5BACD209" ID-Zenodo-Dep="1303486" doi="10.3897/jhr.64.24282.figure16" httpUri="https://binary.pensoft.net/fig/212232" pageId="20" pageNumber="161" start="Figure 16" startId="F16">
<paragraph id="33E8F4573B286B09A4EE7F5CB0E3DA93" pageId="20" pageNumber="161">
<emphasis id="BEAEC3F758BA1CA7754D443B114E59CF" bold="true" pageId="20" pageNumber="161">Figure 16.</emphasis>
Characters used in the morphometric analysis.
<emphasis bold="true" pageId="20" pageNumber="161">A</emphasis>
<emphasis id="6E11BDB005D551681AA8962AB6E6E8C2" bold="true" pageId="20" pageNumber="161">A</emphasis>
Frontal view of the head illustrating the morphometric character malar space (mlr.l),
<taxonomicName authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya annleckieae" order="Hymenoptera" pageId="20" pageNumber="161" phylum="Arthropoda" rank="species" species="annleckieae">
<emphasis italics="true" pageId="20" pageNumber="161">Eadya annleckieae</emphasis>
<taxonomicName id="23A8BC675CF13520D3C104F4A85E05ED" authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya annleckieae" order="Hymenoptera" pageId="20" pageNumber="161" phylum="Arthropoda" rank="species" species="annleckieae">
<emphasis id="E02D52442C4EBFA0E390E1C470223681" italics="true" pageId="20" pageNumber="161">Eadya annleckieae</emphasis>
</taxonomicName>
Ridenbaugh, sp. n. paratype
<emphasis bold="true" pageId="20" pageNumber="161">B</emphasis>
<emphasis id="D1406B351C7AC648BA117D8E550BE34C" bold="true" pageId="20" pageNumber="161">B</emphasis>
Dorsal view of the head illustrating the morphometric characters lateral ocellar line (LOL), ocular ocellar line (OOL), posterior ocellar line (POL), and occipital ocellar line (oci.l),
<taxonomicName authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya annleckieae" order="Hymenoptera" pageId="20" pageNumber="161" phylum="Arthropoda" rank="species" species="annleckieae">
<emphasis italics="true" pageId="20" pageNumber="161">Eadya annleckieae</emphasis>
<taxonomicName id="359EE31C8F2EB17872E952EC7E5A744B" authorityName="Ridenbaugh" authorityYear="2018" class="Insecta" family="Braconidae" genus="Eadya" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eadya annleckieae" order="Hymenoptera" pageId="20" pageNumber="161" phylum="Arthropoda" rank="species" species="annleckieae">
<emphasis id="7B377EBA905F797D67FF2351A640C136" italics="true" pageId="20" pageNumber="161">Eadya annleckieae</emphasis>
</taxonomicName>
Ridenbaugh, sp. n. paratype. All scale bars are 1mm in length.
</paragraph>

View file

@ -1,180 +1,180 @@
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<mods:title>Review of the genus Leptopilina (Hymenoptera, Cynipoidea, Figitidae, Eucoilinae) from the Eastern United States, including three newly described species</mods:title>
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<subSection lastPageId="30" lastPageNumber="65" pageId="29" pageNumber="64" type="results">
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<taxonomicName id="31E96DD73A2ED0D18B52272FA5F269DD" authorityName="Nordlander" authorityYear="1980" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina victoriae" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="victoriae">Leptopilina victoriae</taxonomicName>
Nordlander, 1980: 447 (original description);
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, Mitsui, Suwito &amp; Kimura, 2011: 344 (new distribution record from Japan, Indonesia and Malaysia, host association); van Noort, Buffington &amp; Forshage, 2015: 92 (listed).
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<paragraph pageId="29" pageNumber="64">Diagnosis.</paragraph>
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<paragraph id="1D1EE1FE89E46DBC83A4CD7FCAF52FFF" pageId="29" pageNumber="64">Diagnosis.</paragraph>
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The shape of antenna and metasoma of
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<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina victoriae</emphasis>
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is similar to
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<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina boulardi</emphasis>
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<emphasis id="4772E6F568120FF244B08195E59CAAFC" italics="true" pageId="29" pageNumber="64">Leptopilina boulardi</emphasis>
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. However, the two species can be easily separate by the patterns of the scutellum (Figs
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,
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). In general, the hairy ring in
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<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina victoriae</emphasis>
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<emphasis id="40E94A53DE75DC2AE53807249F954A87" italics="true" pageId="29" pageNumber="64">Leptopilina victoriae</emphasis>
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(Figs
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) is dense and relatively shorter than other
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species (Fig.
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). This species also has a long horizontal ridge across middle of the metapleuron (arrow, Fig.
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). The ridge is parallel the upper ridge of metapleura and that different from other
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<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina</emphasis>
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species the ridge is not display parallel.
</paragraph>
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<emphasis bold="true" pageId="29" pageNumber="64">Figure 46-47.</emphasis>
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<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina victoriae</emphasis>
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<paragraph id="7C147AD6BF37141262A2B653BEE49BB9" pageId="29" pageNumber="64">
<emphasis id="813DC50E89E48077A4C8C8F802BD258D" bold="true" pageId="29" pageNumber="64">Figure 46-47.</emphasis>
<taxonomicName id="12342B2D94F6667F3DF5446114EBB127" authorityName="Nordlander" authorityYear="1980" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina victoriae" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="victoriae">
<emphasis id="D9B866D6F6F71D22C9F1DABA111F4D90" italics="true" pageId="29" pageNumber="64">Leptopilina victoriae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="29" pageNumber="64" type="redescription">
<paragraph pageId="29" pageNumber="64">Redescription.</paragraph>
<paragraph pageId="29" pageNumber="64">Coloration with head, mesosoma, metasoma black to dark brown, legs light brown. Malar sulcus present. Apical segment of maxillary palp more than 1.5 times as long as preceding segment. Terminal flagellomere with three basiconic sensillae. Basiconic sensillae present on F6-F11. Placoidal sensilla present on F7-11. Number of ridges on pronotal plate in lateral view 2. Sculpture on mesoscutum absent, entire surface smooth, shiny. Dorsal surface of scutellum areolet - rugulose. Circumscutellar carina present, complete, delimiting dorsal and ventral halves of scutellum. Latero-ventral margin of scutellum posterior to axillula smooth ventrally, weakly rugulose dorsally. Dorsal part of scutellum entirely areolate. Scutellar plate, in dorsal view, medium sized, exposing about half of scutellum. Posterior impression of metepimeron absent. Anterior impression of metepisternum, immediately beneath anterior end of metapleural carina, present, small and narrow. Wing vein M present but not well defined. Inter propodeal carinae space lightly setose, smooth. Horizontal carina running anteriorly from lateral propodeal carina, absent. Surface of petiole longitudinally costate, ventral keel absent. Setal band (hairy ring) at base of tergum 3 present, interrupted dorsally, ventrally, dense short hair.</paragraph>
<subSubSection id="60A649152052273C0BD1D67820690AA5" pageId="29" pageNumber="64" type="redescription">
<paragraph id="F531F4804E976A07C3F48504009A7216" pageId="29" pageNumber="64">Redescription.</paragraph>
<paragraph id="463C372261A35B03E4F913ED20715C50" pageId="29" pageNumber="64">Coloration with head, mesosoma, metasoma black to dark brown, legs light brown. Malar sulcus present. Apical segment of maxillary palp more than 1.5 times as long as preceding segment. Terminal flagellomere with three basiconic sensillae. Basiconic sensillae present on F6-F11. Placoidal sensilla present on F7-11. Number of ridges on pronotal plate in lateral view 2. Sculpture on mesoscutum absent, entire surface smooth, shiny. Dorsal surface of scutellum areolet - rugulose. Circumscutellar carina present, complete, delimiting dorsal and ventral halves of scutellum. Latero-ventral margin of scutellum posterior to axillula smooth ventrally, weakly rugulose dorsally. Dorsal part of scutellum entirely areolate. Scutellar plate, in dorsal view, medium sized, exposing about half of scutellum. Posterior impression of metepimeron absent. Anterior impression of metepisternum, immediately beneath anterior end of metapleural carina, present, small and narrow. Wing vein M present but not well defined. Inter propodeal carinae space lightly setose, smooth. Horizontal carina running anteriorly from lateral propodeal carina, absent. Surface of petiole longitudinally costate, ventral keel absent. Setal band (hairy ring) at base of tergum 3 present, interrupted dorsally, ventrally, dense short hair.</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="64" type="distribution">
<paragraph pageId="29" pageNumber="64">Distribution in Eastern North America.</paragraph>
<paragraph pageId="29" pageNumber="64">
<subSubSection id="6AF591C60D67FC81C3BC6244C4490CA7" pageId="29" pageNumber="64" type="distribution">
<paragraph id="971B08CA81A8DDFAAE582A2D901D0616" pageId="29" pageNumber="64">Distribution in Eastern North America.</paragraph>
<paragraph id="E06D70144DC4548FDAD9CCE6A6BA72E8" pageId="29" pageNumber="64">
Of the seven species in our identification key,
<taxonomicName authorityName="Nordlander" authorityYear="1980" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina victoriae" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="victoriae">
<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina victoriae</emphasis>
<taxonomicName id="7CDDE774057C0626A33277BA26680F89" authorityName="Nordlander" authorityYear="1980" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina victoriae" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="victoriae">
<emphasis id="284D5AD8328CADD9FCE7398A0B411F13" italics="true" pageId="29" pageNumber="64">Leptopilina victoriae</emphasis>
</taxonomicName>
did not appear in our field collections. However, because this species is commonly used in laboratory experiments we include this species in the identification key to assist with diagnosis of other
<taxonomicName authorityName="Foerster" authorityYear="1869" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina</emphasis>
<taxonomicName id="D6F5C149DEE24AD263B886FB7F882F6E" authorityName="Foerster" authorityYear="1869" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="genus">
<emphasis id="8CC7539731152765C4001254E2C1948F" italics="true" pageId="29" pageNumber="64">Leptopilina</emphasis>
</taxonomicName>
species that are also commonly used as laboratory strains (e.g.,
<taxonomicName baseAuthorityName="Barbotin, Carton &amp; Kelner-Pillault" baseAuthorityYear="1979" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina boulardi" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="boulardi">
<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina boulardi</emphasis>
<taxonomicName id="FFAB79F79C9EE48DA9B3E67C6207F382" baseAuthorityName="Barbotin, Carton &amp; Kelner-Pillault" baseAuthorityYear="1979" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina boulardi" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="boulardi">
<emphasis id="14710003AFACB6F7B40AF155027AA7DE" italics="true" pageId="29" pageNumber="64">Leptopilina boulardi</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Thomson" baseAuthorityYear="1862" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina heterotoma" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="heterotoma">
<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina heterotoma</emphasis>
<taxonomicName id="B286ECF1DDEABFF890A277D642BD87AF" baseAuthorityName="Thomson" baseAuthorityYear="1862" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina heterotoma" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="heterotoma">
<emphasis id="282EC42453CAA2EE645B4F52E52144D6" italics="true" pageId="29" pageNumber="64">Leptopilina heterotoma</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Hartig" baseAuthorityYear="1841" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina clavipes" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis italics="true" pageId="29" pageNumber="64">Leptopilina clavipes</emphasis>
<taxonomicName id="D6AE8AD90465FFA84425AC7DEC5EADC9" baseAuthorityName="Hartig" baseAuthorityYear="1841" class="Insecta" family="Figitidae" genus="Leptopilina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptopilina clavipes" order="Hymenoptera" pageId="29" pageNumber="64" phylum="Arthropoda" rank="species" species="clavipes">
<emphasis id="145C4FC12BF8EE5E1DB5C683FAA21921" italics="true" pageId="29" pageNumber="64">Leptopilina clavipes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="65" type="materials_examined">
<paragraph pageId="30" pageNumber="65">
<pageBreakToken pageId="30" pageNumber="65" start="start">Material</pageBreakToken>
<subSubSection id="41858897B218BA7B66E0A5E84A12010D" pageId="30" pageNumber="65" type="materials_examined">
<paragraph id="9143677DC033AAAD31097984BDE332E8" pageId="30" pageNumber="65">
<pageBreakToken id="7729490C04BF57664D6AB61174B5910B" pageId="30" pageNumber="65" start="start">Material</pageBreakToken>
examined.
</paragraph>
<paragraph pageId="30" pageNumber="65">
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Strain G" country="Seychelles" location="Seychelles" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="Paratypes">
<typeStatus>Paratypes</typeStatus>
<paragraph id="DBAB229408F6D450A8C78C82F16402AE" pageId="30" pageNumber="65">
<materialsCitation id="3860DDAC61E0E4D2CC589B41C97821BC" collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Strain G" country="Seychelles" location="Seychelles" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="Paratypes">
<typeStatus id="DD7B7BCE4A9A81FA412A232A8D14AE87">Paratypes</typeStatus>
(
<specimenCount type="female">2 females</specimenCount>
<specimenCount id="F296E91D7F38EE4DC44BE38CE7BF20D8" type="female">2 females</specimenCount>
,
<specimenCount type="male">2 males</specimenCount>
<specimenCount id="95B91EDB9A447822BCD87D3DD4B61DD2" type="male">2 males</specimenCount>
, BMNH):
<collectorName>Strain G</collectorName>
<collectorName id="5E7DA74CDD366701BA6427504642247E">Strain G</collectorName>
311-1,
<collectingCountry name="Seychelles">Seychelles</collectingCountry>
<collectingCountry id="6A5907AFF5837BBC17DAB924FF12A2A8" name="Seychelles">Seychelles</collectingCountry>
. G.
<bibRefCitation author="Nordlander, G" journalOrPublisher="Entomologica Scandinavica" pageId="34" pageNumber="69" pagination="428 - 453" publicationUrl="10.1163/187631280794710024" refId="B21" refString="Nordlander, G, 1980. Revision of the genus Leptopilina Foerster, 1869, with notes on the status of some other genera (Hymenoptera, Cynipoidea, Eucoilidae). Entomologica Scandinavica 11: 428 - 453, DOI: 10.1163/187631280794710024" title="Revision of the genus Leptopilina Foerster, 1869, with notes on the status of some other genera (Hymenoptera, Cynipoidea, Eucoilidae)." url="10.1163/187631280794710024" volume="11" year="1980">Nordlander. 1980</bibRefCitation>
<bibRefCitation id="E5E97AA43271F582BB67A6EFC1A710D6" author="Nordlander, G" journalOrPublisher="Entomologica Scandinavica" pageId="34" pageNumber="69" pagination="428 - 453" publicationUrl="10.1163/187631280794710024" refId="B21" refString="Nordlander, G, 1980. Revision of the genus Leptopilina Foerster, 1869, with notes on the status of some other genera (Hymenoptera, Cynipoidea, Eucoilidae). Entomologica Scandinavica 11: 428 - 453, DOI: 10.1163/187631280794710024" title="Revision of the genus Leptopilina Foerster, 1869, with notes on the status of some other genera (Hymenoptera, Cynipoidea, Eucoilidae)." url="10.1163/187631280794710024" volume="11" year="1980">Nordlander. 1980</bibRefCitation>
. Es 551-552. BMNH(E) 970160-970161;
<collectorName>Strain G</collectorName>
<collectorName id="88831E821B4BEDC69BDFABDC8116964F">Strain G</collectorName>
314-1,
<collectingCountry name="Seychelles">Seychelles</collectingCountry>
<collectingCountry id="140F4F378A51CE3AFF555AEDC3615384" name="Seychelles">Seychelles</collectingCountry>
. G.
<bibRefCitation author="Nordlander, G" journalOrPublisher="Entomologica Scandinavica" pageId="34" pageNumber="69" pagination="428 - 453" publicationUrl="10.1163/187631280794710024" refId="B21" refString="Nordlander, G, 1980. Revision of the genus Leptopilina Foerster, 1869, with notes on the status of some other genera (Hymenoptera, Cynipoidea, Eucoilidae). Entomologica Scandinavica 11: 428 - 453, DOI: 10.1163/187631280794710024" title="Revision of the genus Leptopilina Foerster, 1869, with notes on the status of some other genera (Hymenoptera, Cynipoidea, Eucoilidae)." url="10.1163/187631280794710024" volume="11" year="1980">Nordlander. 1980</bibRefCitation>
<bibRefCitation id="7E6F7AD2E941DACE8F18A472D98E2AD4" author="Nordlander, G" journalOrPublisher="Entomologica Scandinavica" pageId="34" pageNumber="69" pagination="428 - 453" publicationUrl="10.1163/187631280794710024" refId="B21" refString="Nordlander, G, 1980. Revision of the genus Leptopilina Foerster, 1869, with notes on the status of some other genera (Hymenoptera, Cynipoidea, Eucoilidae). Entomologica Scandinavica 11: 428 - 453, DOI: 10.1163/187631280794710024" title="Revision of the genus Leptopilina Foerster, 1869, with notes on the status of some other genera (Hymenoptera, Cynipoidea, Eucoilidae)." url="10.1163/187631280794710024" volume="11" year="1980">Nordlander. 1980</bibRefCitation>
. Es 553-554
</materialsCitation>
.

View file

@ -1,295 +1,295 @@
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<paragraph id="E6FEC29B9817A8B1A6EE31F151BE450F" pageId="0" pageNumber="147">
5.
<taxonomicName LSID="A1CF27D2-6DE6-54A0-9B78-EDDC0FA12289" class="Insecta" family="Carabidae" genus="Tropopterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropopterus canaliculus" order="Coleoptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="canaliculus">Tropopterus canaliculus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="147">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tropopterus spp., dorsal view; range of standardized body lengths in mm. A. T. montagnei female; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). B. T. giraudyi female; Valdivia Prov., Chaihuin (EMEC). C. T. peckorum male; Quillota Prov., P. N. La Campana (FMNH). D. T. robustus female; Malleco Prov., M. N. Contulmo (FMNH). E. T. canaliculus female; Malleco Prov., P. N. Nahuelbuta (FMNH). F. T. duponchelii male; Arauco Prov., Caramavida (MCZ)." figureDoi="10.3897/dez.66.38022.figure1" httpUri="https://binary.pensoft.net/fig/354024" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure1">Figures 1E</figureCitation>
<taxonomicName id="EB6E49EB675BBDF3C7451EC6DEC9DB01" ID-CoL="8LY4R" LSID="A1CF27D2-6DE6-54A0-9B78-EDDC0FA12289" class="Insecta" family="Carabidae" genus="Tropopterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropopterus canaliculus" order="Coleoptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="canaliculus">Tropopterus canaliculus</taxonomicName>
<taxonomicNameLabel id="8CA64060F13EED12A93BEEAFF76F1D35" pageId="0" pageNumber="147">sp. nov.</taxonomicNameLabel>
<figureCitation id="CE5E5D2B4CCD7DC64A26A4446428477B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tropopterus spp., dorsal view; range of standardized body lengths in mm. A. T. montagnei female; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). B. T. giraudyi female; Valdivia Prov., Chaihuin (EMEC). C. T. peckorum male; Quillota Prov., P. N. La Campana (FMNH). D. T. robustus female; Malleco Prov., M. N. Contulmo (FMNH). E. T. canaliculus female; Malleco Prov., P. N. Nahuelbuta (FMNH). F. T. duponchelii male; Arauco Prov., Caramavida (MCZ)." figureDoi="10.3897/dez.66.38022.figure1" httpUri="https://binary.pensoft.net/fig/354024" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure1">Figures 1E</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagal median lobe, left view, and antecostal apodeme and tergite of abdominal IX, dorsal view. A-D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta. A, B. (FMNH). C, D. (OSAC). E-G. T. duponchelii; Arauco Prov., Caramavida. E. (NMNH). F-G. (MCZ). For abbreviations used to label structures see Table 1." figureDoi="10.3897/dez.66.38022.figure4" httpUri="https://binary.pensoft.net/fig/354027" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure4">
4
<normalizedToken originalValue="AD">A-D</normalizedToken>
<figureCitation id="9AB2A9397C888D0982A9D2B9AAFAC3BA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagal median lobe, left view, and antecostal apodeme and tergite of abdominal IX, dorsal view. A-D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta. A, B. (FMNH). C, D. (OSAC). E-G. T. duponchelii; Arauco Prov., Caramavida. E. (NMNH). F-G. (MCZ). For abbreviations used to label structures see Table 1." figureDoi="10.3897/dez.66.38022.figure4" httpUri="https://binary.pensoft.net/fig/354027" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure4">
4
<normalizedToken id="01E05E8DCE7A3036E10BCFD479F4AB68" originalValue="AD">A-D</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paired sets of left parameres (above) and right parmeres (below), ectal view, for male specimens of Tropopterus spp. Parameres are in anatomically correct orientation with ventral surface toward bottom of drawing. A, B. T. giraudyi. A. Malleco Prov., P. N. Nahuelbuta (FMNH). B. Chili, &quot; Tropopterus nitidus Sol. &quot; [sic], Chaudoir Coll. (MNHN). C. T. peckorum; Quillota Prov., P. N. La Campana (FMNH). D, E. T. robustus. D. Malleco Prov., P. N. Contulmo (CMNH). E. Cautin Prov., Bellavista (CAS). F, G. T. canaliculus. F. Malleco Prov., P. N. Nahuelbuta (FMNH). G. Arauco Prov., Caramavida (NMNH). H-I. T. duponchelii. H. Talca Prov., Altos de Vilches (MNHN). I. Arauco Prov., Caramavida (MCZ). J, K. T. minimucro. J. Cautin Prov., Bellavista (EMEC). K. Llanquihue Prov. P. N. Vicente Perez Rosales (FMNH). L. T. fieldianus; Valdivia, La Union (FSCA)." figureDoi="10.3897/dez.66.38022.figure6" httpUri="https://binary.pensoft.net/fig/354029" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure6">6F, G</figureCitation>
<figureCitation id="33C1899310078ED7161979C1B3B0A7BF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paired sets of left parameres (above) and right parmeres (below), ectal view, for male specimens of Tropopterus spp. Parameres are in anatomically correct orientation with ventral surface toward bottom of drawing. A, B. T. giraudyi. A. Malleco Prov., P. N. Nahuelbuta (FMNH). B. Chili, &quot; Tropopterus nitidus Sol. &quot; [sic], Chaudoir Coll. (MNHN). C. T. peckorum; Quillota Prov., P. N. La Campana (FMNH). D, E. T. robustus. D. Malleco Prov., P. N. Contulmo (CMNH). E. Cautin Prov., Bellavista (CAS). F, G. T. canaliculus. F. Malleco Prov., P. N. Nahuelbuta (FMNH). G. Arauco Prov., Caramavida (NMNH). H-I. T. duponchelii. H. Talca Prov., Altos de Vilches (MNHN). I. Arauco Prov., Caramavida (MCZ). J, K. T. minimucro. J. Cautin Prov., Bellavista (EMEC). K. Llanquihue Prov. P. N. Vicente Perez Rosales (FMNH). L. T. fieldianus; Valdivia, La Union (FSCA)." figureDoi="10.3897/dez.66.38022.figure6" httpUri="https://binary.pensoft.net/fig/354029" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure6">6F, G</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female reproductive tract and gonocoxae of abdominal segment IX, ventral view. A. T. montagnei; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). Spermatheca disassociated from bursal assemblage. B. T. giraudyi; Llanquihue Prov., P. N. Vicente Perez Rosales (FMNH). C. T. robustus; Malleco Prov., M. N. Contulmo (FMNH). D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta (FMNH). E. T. duponchelii; Arauco Prov., Caramavida (MCZ). F. T. minimucro; Valdivia Prov., P. N. Oncol (EMEC). For abbreviations used to label structures see Table 1. Vertical scale bars: 0.5 mm." figureDoi="10.3897/dez.66.38022.figure7" httpUri="https://binary.pensoft.net/fig/354030" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure7">7D</figureCitation>
<figureCitation id="B3F39207CB23E76DE4C1E944E7B7BFDB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female reproductive tract and gonocoxae of abdominal segment IX, ventral view. A. T. montagnei; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). Spermatheca disassociated from bursal assemblage. B. T. giraudyi; Llanquihue Prov., P. N. Vicente Perez Rosales (FMNH). C. T. robustus; Malleco Prov., M. N. Contulmo (FMNH). D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta (FMNH). E. T. duponchelii; Arauco Prov., Caramavida (MCZ). F. T. minimucro; Valdivia Prov., P. N. Oncol (EMEC). For abbreviations used to label structures see Table 1. Vertical scale bars: 0.5 mm." figureDoi="10.3897/dez.66.38022.figure7" httpUri="https://binary.pensoft.net/fig/354030" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure7">7D</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Left gonocoxa (except where noted), ventral view, for Tropopterus spp. Specimen information is parallel to that in Figure 7 A-F. For abbreviations used to label structures see Table 1. A. T. montagnei. B. T. giraudyi. C. T. robustus. D. T. canaliculus. E. T. duponchelii. F. T. minimucro (right gonocoxa)." figureDoi="10.3897/dez.66.38022.figure8" httpUri="https://binary.pensoft.net/fig/354031" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure8">8D</figureCitation>
<figureCitation id="E18345F840AEBBC6A05135D0A1F986CF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Left gonocoxa (except where noted), ventral view, for Tropopterus spp. Specimen information is parallel to that in Figure 7 A-F. For abbreviations used to label structures see Table 1. A. T. montagnei. B. T. giraudyi. C. T. robustus. D. T. canaliculus. E. T. duponchelii. F. T. minimucro (right gonocoxa)." figureDoi="10.3897/dez.66.38022.figure8" httpUri="https://binary.pensoft.net/fig/354031" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure8">8D</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I. Geographic distributions of Tropopterus spp. A. T. montagnei. B. T. giraudyi. C. T. peckorum. D. T. robustus. E. T. canaliculus. F. T. duponchelii. G. T. trisinuatus. H. T. minimucro. I. T. fieldianus. J. Geographical distributions of Tropopterus spp., labeled as per figures above, hierarchically arranged by phylogenetic relationships of the taxon cladogram, Fig. 10." figureDoi="10.3897/dez.66.38022.figure9" httpUri="https://binary.pensoft.net/fig/354032" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure9">9E</figureCitation>
<figureCitation id="984A88BFF6DBC57B4F3A037504857A28" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I. Geographic distributions of Tropopterus spp. A. T. montagnei. B. T. giraudyi. C. T. peckorum. D. T. robustus. E. T. canaliculus. F. T. duponchelii. G. T. trisinuatus. H. T. minimucro. I. T. fieldianus. J. Geographical distributions of Tropopterus spp., labeled as per figures above, hierarchically arranged by phylogenetic relationships of the taxon cladogram, Fig. 10." figureDoi="10.3897/dez.66.38022.figure9" httpUri="https://binary.pensoft.net/fig/354032" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure9">9E</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cladogram returned under all cladistic protocols described in text; tree length = 97, CI = 57, RI = 47, fast character optimization shown. Outgroup choice of Pharetis thayerae based on results in Liebherr (2020). Geographic region abbreviations after taxon name include: eastern Australia (EOZ); Santiagan entomofaunal province (San); Northern Valdivian entomofaunal province (NV); Southern Valdivian Entomofaunal province (SV)." figureDoi="10.3897/dez.66.38022.figure10" httpUri="https://binary.pensoft.net/fig/354033" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure10">10</figureCitation>
<figureCitation id="AB213530AA7DB02F505FE6124F69AC19" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Cladogram returned under all cladistic protocols described in text; tree length = 97, CI = 57, RI = 47, fast character optimization shown. Outgroup choice of Pharetis thayerae based on results in Liebherr (2020). Geographic region abbreviations after taxon name include: eastern Australia (EOZ); Santiagan entomofaunal province (San); Northern Valdivian entomofaunal province (NV); Southern Valdivian Entomofaunal province (SV)." figureDoi="10.3897/dez.66.38022.figure10" httpUri="https://binary.pensoft.net/fig/354033" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure10">10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="diagnosis">
<paragraph pageId="0" pageNumber="147">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="147">
<subSubSection id="DB20C1D75B0ABC706C09FEFB73DCA98C" pageId="0" pageNumber="147" type="diagnosis">
<paragraph id="88C0FC93034962DBAE40253E62021E51" pageId="0" pageNumber="147">Diagnosis</paragraph>
<paragraph id="8C8C7BF45FEEBC8A370E71BA9D12A54B" pageId="0" pageNumber="147">
(
<emphasis italics="true" pageId="0" pageNumber="147">n</emphasis>
<emphasis id="1A27288099C898C1930BCB99A313612F" italics="true" pageId="0" pageNumber="147">n</emphasis>
= 5). Both this species and
<taxonomicName genus="T." lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
<taxonomicName id="F3940FF7341F4CF09B96F13CDD418E58" lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis id="676E0F9FE489C8984521179E78199E19" italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
</taxonomicName>
exhibit subparallel elytra with the sutural stria represented by a series of isolated punctures on the disc, and the second stria much reduced though still visible as a series of minute punctures (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tropopterus spp., dorsal view; range of standardized body lengths in mm. A. T. montagnei female; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). B. T. giraudyi female; Valdivia Prov., Chaihuin (EMEC). C. T. peckorum male; Quillota Prov., P. N. La Campana (FMNH). D. T. robustus female; Malleco Prov., M. N. Contulmo (FMNH). E. T. canaliculus female; Malleco Prov., P. N. Nahuelbuta (FMNH). F. T. duponchelii male; Arauco Prov., Caramavida (MCZ)." figureDoi="10.3897/dez.66.38022.figure1" httpUri="https://binary.pensoft.net/fig/354024" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure1">Fig. 1E, F</figureCitation>
<figureCitation id="2CE5711CE55F41F5626E23C7D36F5E89" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tropopterus spp., dorsal view; range of standardized body lengths in mm. A. T. montagnei female; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). B. T. giraudyi female; Valdivia Prov., Chaihuin (EMEC). C. T. peckorum male; Quillota Prov., P. N. La Campana (FMNH). D. T. robustus female; Malleco Prov., M. N. Contulmo (FMNH). E. T. canaliculus female; Malleco Prov., P. N. Nahuelbuta (FMNH). F. T. duponchelii male; Arauco Prov., Caramavida (MCZ)." figureDoi="10.3897/dez.66.38022.figure1" httpUri="https://binary.pensoft.net/fig/354024" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure1">Fig. 1E, F</figureCitation>
). Individuals of
<taxonomicName genus="T." lsidName="T. canaliculus" pageId="0" pageNumber="147" rank="species" species="canaliculus">
<emphasis italics="true" pageId="0" pageNumber="147">T. canaliculus</emphasis>
<taxonomicName id="5CCBD169369BE96F084301366EA7188F" lsidName="T. canaliculus" pageId="0" pageNumber="147" rank="species" species="canaliculus">
<emphasis id="DD40B94C4E4282EA09DF0A11C74DBE4B" italics="true" pageId="0" pageNumber="147">T. canaliculus</emphasis>
</taxonomicName>
with more reduced second elytral striae could be confused with those of
<taxonomicName genus="T." lsidName="T. minimucro" pageId="0" pageNumber="147" rank="species" species="minimucro">
<emphasis italics="true" pageId="0" pageNumber="147">T. minimucro</emphasis>
<taxonomicName id="8FFF2D07E9B047E230382398FC2B114E" lsidName="T. minimucro" pageId="0" pageNumber="147" rank="species" species="minimucro">
<emphasis id="2ECADFFAEB97B2F4A5BB2EB452071E0E" italics="true" pageId="0" pageNumber="147">T. minimucro</emphasis>
</taxonomicName>
; however, that species is characterized by the absence of a basal elytral groove, the elytral surface smoothly convex inside the humeral tooth at the anterior end of the lateral marginal depression (compare
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tropopterus spp., dorsal view; range of standardized body lengths in mm. A. T. montagnei female; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). B. T. giraudyi female; Valdivia Prov., Chaihuin (EMEC). C. T. peckorum male; Quillota Prov., P. N. La Campana (FMNH). D. T. robustus female; Malleco Prov., M. N. Contulmo (FMNH). E. T. canaliculus female; Malleco Prov., P. N. Nahuelbuta (FMNH). F. T. duponchelii male; Arauco Prov., Caramavida (MCZ)." figureDoi="10.3897/dez.66.38022.figure1" httpUri="https://binary.pensoft.net/fig/354024" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure1">Figs 1E</figureCitation>
<figureCitation id="0B13432F193CB85138F569035B2A65AF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tropopterus spp., dorsal view; range of standardized body lengths in mm. A. T. montagnei female; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). B. T. giraudyi female; Valdivia Prov., Chaihuin (EMEC). C. T. peckorum male; Quillota Prov., P. N. La Campana (FMNH). D. T. robustus female; Malleco Prov., M. N. Contulmo (FMNH). E. T. canaliculus female; Malleco Prov., P. N. Nahuelbuta (FMNH). F. T. duponchelii male; Arauco Prov., Caramavida (MCZ)." figureDoi="10.3897/dez.66.38022.figure1" httpUri="https://binary.pensoft.net/fig/354024" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure1">Figs 1E</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tropopterus spp., dorsal view; range of standardized body lengths in mm. A. T. trisinuatus female; Cautin Prov., Villarrica (CAS). B. T. minimucro female; Valdivia Prov., P. N. Oncol (OSAC). C. T. fieldianus male; Osorno Prov., Maicolpue (FMNH)." figureDoi="10.3897/dez.66.38022.figure2" httpUri="https://binary.pensoft.net/fig/354025" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure2">2B</figureCitation>
<figureCitation id="D4733EEFB3082E3D3B5D9B749A038C0B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tropopterus spp., dorsal view; range of standardized body lengths in mm. A. T. trisinuatus female; Cautin Prov., Villarrica (CAS). B. T. minimucro female; Valdivia Prov., P. N. Oncol (OSAC). C. T. fieldianus male; Osorno Prov., Maicolpue (FMNH)." figureDoi="10.3897/dez.66.38022.figure2" httpUri="https://binary.pensoft.net/fig/354025" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure2">2B</figureCitation>
). Compared to
<taxonomicName genus="T." lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
<taxonomicName id="D3441B838C6E8401B734B4364A1F2EAA" lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis id="B3AA40C177FDB70789B79B53E87007DE" italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
</taxonomicName>
,
<taxonomicName genus="T." lsidName="T. canaliculus" pageId="0" pageNumber="147" rank="species" species="canaliculus">
<emphasis italics="true" pageId="0" pageNumber="147">T. canaliculus</emphasis>
<taxonomicName id="DED12B5CA2D72BB54ED0A50FAE9C9A97" lsidName="T. canaliculus" pageId="0" pageNumber="147" rank="species" species="canaliculus">
<emphasis id="D11E44EF8B81C4AF41543700D91FA738" italics="true" pageId="0" pageNumber="147">T. canaliculus</emphasis>
</taxonomicName>
can be characterized by the prosternum bearing a median, canaliculate depression that extends from the ventral face of the prosternal projection 2/3 the distance toward the anterior pronotal margin, whereas the prosternum of
<taxonomicName genus="T." lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
<taxonomicName id="4918F30B8C1889833DC7CF1B3BE4F767" lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis id="E55B47B0ADDE734B315AF382185C1737" italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
</taxonomicName>
is medially flattened to broadly depressed, not narrowly canaliculate. The two species can also be told by individuals of
<taxonomicName genus="T." lsidName="T. canaliculus" pageId="0" pageNumber="147" rank="species" species="canaliculus">
<emphasis italics="true" pageId="0" pageNumber="147">T. canaliculus</emphasis>
<taxonomicName id="5324A85CBFC881E1B4F448707316B0B3" lsidName="T. canaliculus" pageId="0" pageNumber="147" rank="species" species="canaliculus">
<emphasis id="CD3ED92CA483E4960E3A7F98F534B951" italics="true" pageId="0" pageNumber="147">T. canaliculus</emphasis>
</taxonomicName>
exhibiting a narrower pronotum (MPW/PL = 1.20-1.25 versus 1.25-1.32 for
<taxonomicName genus="T." lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
<taxonomicName id="F143024F776A77F2268471C53588B251" lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis id="09D36728E593D41573B2DC4C4489F443" italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
</taxonomicName>
) and a glossier upper body surface.
<taxonomicName class="Insecta" family="Carabidae" genus="Tropopterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropopterus canaliculus" order="Coleoptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="canaliculus">
<emphasis italics="true" pageId="0" pageNumber="147">Tropopterus canaliculus</emphasis>
<taxonomicName id="2687C808F77862CA9F0A428A54724FAF" class="Insecta" family="Carabidae" genus="Tropopterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropopterus canaliculus" order="Coleoptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="canaliculus">
<emphasis id="4E92E7BB31A88E13DCACC0BD8D7B5937" italics="true" pageId="0" pageNumber="147">Tropopterus canaliculus</emphasis>
</taxonomicName>
has the vertex glossy with indistinct transverse sculpticells over portions of the surface, the pronotum also glossy, and the elytral disc glossy with indistinct transverse sculpticells visible near depressed portions of the surface such as the strial punctures, and minute micropunctures regularly distributed across the glossy surface. The dorsal microsculpture of
<taxonomicName genus="T." lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
<taxonomicName id="A538C0C32569ABB1B3A005F43D948439" lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis id="B493C4FDDAA68FA790184EF9B2D501E2" italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
</taxonomicName>
is much more developed, with both the pronotum and elytra bearing dense transverse lines that cause iridescence of the latter. In this species the pronotum is sparsely punctate in the laterobasal depressions and lateral portions of the median base, versus smooth in
<taxonomicName genus="T." lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
<taxonomicName id="345384C6EAC0556484FA091144E70CBB" lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis id="12CE89BE377C4FB502C299C79972A7A9" italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
</taxonomicName>
, and the anterior pronotal margin is completely beaded, versus smooth and unmargined across the median half in
<taxonomicName genus="T." lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
<taxonomicName id="5A04AF50E515EC555784B9DD0B42D9C0" lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis id="FB50CF33C4F7B0EC2E317B814BABF3C1" italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
</taxonomicName>
. The standardized body length = 5.8-7.5 mm; a smaller though not diagnostic range of sizes compared to
<taxonomicName genus="T." lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
<taxonomicName id="CB1DF190E31BBFF0407DB74F82035AF9" lsidName="T. duponchelii" pageId="0" pageNumber="147" rank="species" species="duponchelii">
<emphasis id="DD7AEBA873839CD05C7E3DDE19C08851" italics="true" pageId="0" pageNumber="147">T. duponchelii</emphasis>
</taxonomicName>
(below).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="description">
<paragraph pageId="0" pageNumber="147">Description.</paragraph>
<paragraph pageId="0" pageNumber="147">
Head capsule broad basally, eyes moderately convex, ocular ratio 1.40-1.50; eyes covering much of abruptly protruded ocular lobe, ocular lobe ratio 0.85-0.90; horizontal diameter of eye crossing 25 ommatidia; antennomeres moderately elongate, length of antennomere 9 2.0
<normalizedToken originalValue="×">x</normalizedToken>
maximal diameter; mandibles moderately elongate, distance from anterior condyle to apex of left mandible 1.81
<normalizedToken originalValue="×">x</normalizedToken>
distance from condyle to lateroapical margin of labrum; mentum basal breadth 2.86
<normalizedToken originalValue="×">x</normalizedToken>
length from lateral apex to base, deep paramedial pits laterad base of mentum tooth; ligular apex broad, truncate, two setae separated by four setal diameters; paraglossae extended as far beyond ligular margin as distance from their base to ligular margin. Pronotal lateral margins convex, hind angles distinctly denticulate; basal margin with complete marginal bead, median base sparsely punctate across breadth, the punctures continuing across laterobasal depressions; median longitudinal impression with lenticular depression at front of median base, narrow and deep on disc; anterior transverse impression broad and shallow medially, traceable to front angles, anterior callosity broadly convex; anterior margin completely beaded, the bead continuous medially; front angle slightly protruded, right angled; lateral marginal depression very narrow in apical half, broadened from lateral seta to laterobasal depression; lateral seta separated from lateral marginal depression by the diameter of setal articulatory socket; laterobasal depression rugosely punctate, surface upraised in middle of depression; surface of proepisternum slightly irregular, as if beaten by a ball-peen hammer. Elytra broad, sides subparallel, humeri broad; basal groove deep, continuous from anterad parascutellar seta to tightly rounded humeral angle; parascutellar striole absent, sutural stria reaching basal margin mesad parascutellar seta; sutural stria a series of isolated punctures on disc, continuous and smooth apically, the striae of both elytra bordering upraised sutural callus associated with conjoined elytra; surface of elytra depressed mesad subapical and apical elytral setae; stria 8 continuous throughout length; lateral marginal depression broad, lined with transverse sculpticells; subapical sinuation broad, moderately excavated, elytral plica evident in dorsolateral view. Mesepisternum with broad vertical depression lined with ~8 punctures in 2 or 3 vertical rows; metasternal process narrowly rounded apically, apex broadly and sides more narrowly upraised; metepisternum somewhat elongate, maximal width 0.78
<normalizedToken originalValue="×">x</normalizedToken>
<subSubSection id="CD35E236A1FE47B3E92C92CB37414296" pageId="0" pageNumber="147" type="description">
<paragraph id="1F938692AEB0E4DD680B526F0BBE239B" pageId="0" pageNumber="147">Description.</paragraph>
<paragraph id="14F23D342F59FB012510A821750F1259" pageId="0" pageNumber="147">
Head capsule broad basally, eyes moderately convex, ocular ratio 1.40-1.50; eyes covering much of abruptly protruded ocular lobe, ocular lobe ratio 0.85-0.90; horizontal diameter of eye crossing 25 ommatidia; antennomeres moderately elongate, length of antennomere 9 2.0
<normalizedToken id="A348F135FAF1793C290BC2AF4C4FB953" originalValue="×">x</normalizedToken>
maximal diameter; mandibles moderately elongate, distance from anterior condyle to apex of left mandible 1.81
<normalizedToken id="1DFE17BCBBF1CD1FDA23A360428B2B06" originalValue="×">x</normalizedToken>
distance from condyle to lateroapical margin of labrum; mentum basal breadth 2.86
<normalizedToken id="119B2D145BCD67142F310A5C49A7FAFD" originalValue="×">x</normalizedToken>
length from lateral apex to base, deep paramedial pits laterad base of mentum tooth; ligular apex broad, truncate, two setae separated by four setal diameters; paraglossae extended as far beyond ligular margin as distance from their base to ligular margin. Pronotal lateral margins convex, hind angles distinctly denticulate; basal margin with complete marginal bead, median base sparsely punctate across breadth, the punctures continuing across laterobasal depressions; median longitudinal impression with lenticular depression at front of median base, narrow and deep on disc; anterior transverse impression broad and shallow medially, traceable to front angles, anterior callosity broadly convex; anterior margin completely beaded, the bead continuous medially; front angle slightly protruded, right angled; lateral marginal depression very narrow in apical half, broadened from lateral seta to laterobasal depression; lateral seta separated from lateral marginal depression by the diameter of setal articulatory socket; laterobasal depression rugosely punctate, surface upraised in middle of depression; surface of proepisternum slightly irregular, as if beaten by a ball-peen hammer. Elytra broad, sides subparallel, humeri broad; basal groove deep, continuous from anterad parascutellar seta to tightly rounded humeral angle; parascutellar striole absent, sutural stria reaching basal margin mesad parascutellar seta; sutural stria a series of isolated punctures on disc, continuous and smooth apically, the striae of both elytra bordering upraised sutural callus associated with conjoined elytra; surface of elytra depressed mesad subapical and apical elytral setae; stria 8 continuous throughout length; lateral marginal depression broad, lined with transverse sculpticells; subapical sinuation broad, moderately excavated, elytral plica evident in dorsolateral view. Mesepisternum with broad vertical depression lined with ~8 punctures in 2 or 3 vertical rows; metasternal process narrowly rounded apically, apex broadly and sides more narrowly upraised; metepisternum somewhat elongate, maximal width 0.78
<normalizedToken id="E029DB85069E0C848EE4FDF473C2FE1C" originalValue="×">x</normalizedToken>
lateral length. Abdomen with ventrites 3-6 broadly depressed laterally, suture between ventrites 1 and 2 nearly straight, ventrite 2 depressed within slight sinuation; anterior margins of ventrites 4-6 depressed, intersegmental membranes punctate; apical female abdominal ventrite with 2 setae each side, plus a median group of two to four setae, subequal in length; if two nearly in line with lateral setae, if four aligned in an apically broader trapezoid. Coloration of dorsal body surface subtly tricolored, head and pronotum dark rufous, elytra darker, rufopiceous, elytral lateral marginal depressions translucent rufobrunneous; antennae, mouthparts, and legs rufobrunneous; ventral body surface rufopiceous, apex of elytral epipleuron slightly paler, apex of abdominal ventrite 6 narrowly paler to concolorous.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<paragraph id="DBF420FD58B965AA8E74587468606C81" pageId="0" pageNumber="147">
Male genitalia (
<emphasis italics="true" pageId="0" pageNumber="147">n</emphasis>
<emphasis id="6F33137345799DBF77095E3A55793603" italics="true" pageId="0" pageNumber="147">n</emphasis>
= 8). Aedeagal median lobe broad dorsoventrally, but with sclerotized ventral surface of shaft narrow and terminated in a narrow, smooth, spoon-like apex (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagal median lobe, left view, and antecostal apodeme and tergite of abdominal IX, dorsal view. A-D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta. A, B. (FMNH). C, D. (OSAC). E-G. T. duponchelii; Arauco Prov., Caramavida. E. (NMNH). F-G. (MCZ). For abbreviations used to label structures see Table 1." figureDoi="10.3897/dez.66.38022.figure4" httpUri="https://binary.pensoft.net/fig/354027" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure4">Fig. 4A, C</figureCitation>
<figureCitation id="3DE988E03C0BF332F3C330681A512D7D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagal median lobe, left view, and antecostal apodeme and tergite of abdominal IX, dorsal view. A-D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta. A, B. (FMNH). C, D. (OSAC). E-G. T. duponchelii; Arauco Prov., Caramavida. E. (NMNH). F-G. (MCZ). For abbreviations used to label structures see Table 1." figureDoi="10.3897/dez.66.38022.figure4" httpUri="https://binary.pensoft.net/fig/354027" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure4">Fig. 4A, C</figureCitation>
); internal sac with flagellum small, apical. Antecostal apodeme of abdominal IX broadly angulate distally, the short distal extension very broad (
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagal median lobe, left view, and antecostal apodeme and tergite of abdominal IX, dorsal view. A-D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta. A, B. (FMNH). C, D. (OSAC). E-G. T. duponchelii; Arauco Prov., Caramavida. E. (NMNH). F-G. (MCZ). For abbreviations used to label structures see Table 1." figureDoi="10.3897/dez.66.38022.figure4" httpUri="https://binary.pensoft.net/fig/354027" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure4">Fig. 4B, D</figureCitation>
<figureCitation id="E40D3906F224264F494D1842A1F62FB9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagal median lobe, left view, and antecostal apodeme and tergite of abdominal IX, dorsal view. A-D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta. A, B. (FMNH). C, D. (OSAC). E-G. T. duponchelii; Arauco Prov., Caramavida. E. (NMNH). F-G. (MCZ). For abbreviations used to label structures see Table 1." figureDoi="10.3897/dez.66.38022.figure4" httpUri="https://binary.pensoft.net/fig/354027" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure4">Fig. 4B, D</figureCitation>
). Right paramere of variable length, length not associated with length of left paramere (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paired sets of left parameres (above) and right parmeres (below), ectal view, for male specimens of Tropopterus spp. Parameres are in anatomically correct orientation with ventral surface toward bottom of drawing. A, B. T. giraudyi. A. Malleco Prov., P. N. Nahuelbuta (FMNH). B. Chili, &quot; Tropopterus nitidus Sol. &quot; [sic], Chaudoir Coll. (MNHN). C. T. peckorum; Quillota Prov., P. N. La Campana (FMNH). D, E. T. robustus. D. Malleco Prov., P. N. Contulmo (CMNH). E. Cautin Prov., Bellavista (CAS). F, G. T. canaliculus. F. Malleco Prov., P. N. Nahuelbuta (FMNH). G. Arauco Prov., Caramavida (NMNH). H-I. T. duponchelii. H. Talca Prov., Altos de Vilches (MNHN). I. Arauco Prov., Caramavida (MCZ). J, K. T. minimucro. J. Cautin Prov., Bellavista (EMEC). K. Llanquihue Prov. P. N. Vicente Perez Rosales (FMNH). L. T. fieldianus; Valdivia, La Union (FSCA)." figureDoi="10.3897/dez.66.38022.figure6" httpUri="https://binary.pensoft.net/fig/354029" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure6">Fig. 6F, G</figureCitation>
<figureCitation id="0E6EEE120B301E6BA86E7349B7A44D8E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paired sets of left parameres (above) and right parmeres (below), ectal view, for male specimens of Tropopterus spp. Parameres are in anatomically correct orientation with ventral surface toward bottom of drawing. A, B. T. giraudyi. A. Malleco Prov., P. N. Nahuelbuta (FMNH). B. Chili, &quot; Tropopterus nitidus Sol. &quot; [sic], Chaudoir Coll. (MNHN). C. T. peckorum; Quillota Prov., P. N. La Campana (FMNH). D, E. T. robustus. D. Malleco Prov., P. N. Contulmo (CMNH). E. Cautin Prov., Bellavista (CAS). F, G. T. canaliculus. F. Malleco Prov., P. N. Nahuelbuta (FMNH). G. Arauco Prov., Caramavida (NMNH). H-I. T. duponchelii. H. Talca Prov., Altos de Vilches (MNHN). I. Arauco Prov., Caramavida (MCZ). J, K. T. minimucro. J. Cautin Prov., Bellavista (EMEC). K. Llanquihue Prov. P. N. Vicente Perez Rosales (FMNH). L. T. fieldianus; Valdivia, La Union (FSCA)." figureDoi="10.3897/dez.66.38022.figure6" httpUri="https://binary.pensoft.net/fig/354029" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure6">Fig. 6F, G</figureCitation>
), ranging from shorter and broader to more elongate with an attenuated apical extension, the ventral surface bearing from 8-20 setae, two longer setae present near or at apex; left paramere much broader basally, with apical extension ranging from an evenly narrowed projection (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paired sets of left parameres (above) and right parmeres (below), ectal view, for male specimens of Tropopterus spp. Parameres are in anatomically correct orientation with ventral surface toward bottom of drawing. A, B. T. giraudyi. A. Malleco Prov., P. N. Nahuelbuta (FMNH). B. Chili, &quot; Tropopterus nitidus Sol. &quot; [sic], Chaudoir Coll. (MNHN). C. T. peckorum; Quillota Prov., P. N. La Campana (FMNH). D, E. T. robustus. D. Malleco Prov., P. N. Contulmo (CMNH). E. Cautin Prov., Bellavista (CAS). F, G. T. canaliculus. F. Malleco Prov., P. N. Nahuelbuta (FMNH). G. Arauco Prov., Caramavida (NMNH). H-I. T. duponchelii. H. Talca Prov., Altos de Vilches (MNHN). I. Arauco Prov., Caramavida (MCZ). J, K. T. minimucro. J. Cautin Prov., Bellavista (EMEC). K. Llanquihue Prov. P. N. Vicente Perez Rosales (FMNH). L. T. fieldianus; Valdivia, La Union (FSCA)." figureDoi="10.3897/dez.66.38022.figure6" httpUri="https://binary.pensoft.net/fig/354029" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure6">Fig. 6F</figureCitation>
<figureCitation id="2417A73FA55C97F5758CBDA49B1EBD99" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paired sets of left parameres (above) and right parmeres (below), ectal view, for male specimens of Tropopterus spp. Parameres are in anatomically correct orientation with ventral surface toward bottom of drawing. A, B. T. giraudyi. A. Malleco Prov., P. N. Nahuelbuta (FMNH). B. Chili, &quot; Tropopterus nitidus Sol. &quot; [sic], Chaudoir Coll. (MNHN). C. T. peckorum; Quillota Prov., P. N. La Campana (FMNH). D, E. T. robustus. D. Malleco Prov., P. N. Contulmo (CMNH). E. Cautin Prov., Bellavista (CAS). F, G. T. canaliculus. F. Malleco Prov., P. N. Nahuelbuta (FMNH). G. Arauco Prov., Caramavida (NMNH). H-I. T. duponchelii. H. Talca Prov., Altos de Vilches (MNHN). I. Arauco Prov., Caramavida (MCZ). J, K. T. minimucro. J. Cautin Prov., Bellavista (EMEC). K. Llanquihue Prov. P. N. Vicente Perez Rosales (FMNH). L. T. fieldianus; Valdivia, La Union (FSCA)." figureDoi="10.3897/dez.66.38022.figure6" httpUri="https://binary.pensoft.net/fig/354029" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure6">Fig. 6F</figureCitation>
) to a narrow, parallel digitiform extension (
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paired sets of left parameres (above) and right parmeres (below), ectal view, for male specimens of Tropopterus spp. Parameres are in anatomically correct orientation with ventral surface toward bottom of drawing. A, B. T. giraudyi. A. Malleco Prov., P. N. Nahuelbuta (FMNH). B. Chili, &quot; Tropopterus nitidus Sol. &quot; [sic], Chaudoir Coll. (MNHN). C. T. peckorum; Quillota Prov., P. N. La Campana (FMNH). D, E. T. robustus. D. Malleco Prov., P. N. Contulmo (CMNH). E. Cautin Prov., Bellavista (CAS). F, G. T. canaliculus. F. Malleco Prov., P. N. Nahuelbuta (FMNH). G. Arauco Prov., Caramavida (NMNH). H-I. T. duponchelii. H. Talca Prov., Altos de Vilches (MNHN). I. Arauco Prov., Caramavida (MCZ). J, K. T. minimucro. J. Cautin Prov., Bellavista (EMEC). K. Llanquihue Prov. P. N. Vicente Perez Rosales (FMNH). L. T. fieldianus; Valdivia, La Union (FSCA)." figureDoi="10.3897/dez.66.38022.figure6" httpUri="https://binary.pensoft.net/fig/354029" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure6">Fig. 6G</figureCitation>
<figureCitation id="1E11824EB8B6ABBC0D058EA9839DCE55" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Paired sets of left parameres (above) and right parmeres (below), ectal view, for male specimens of Tropopterus spp. Parameres are in anatomically correct orientation with ventral surface toward bottom of drawing. A, B. T. giraudyi. A. Malleco Prov., P. N. Nahuelbuta (FMNH). B. Chili, &quot; Tropopterus nitidus Sol. &quot; [sic], Chaudoir Coll. (MNHN). C. T. peckorum; Quillota Prov., P. N. La Campana (FMNH). D, E. T. robustus. D. Malleco Prov., P. N. Contulmo (CMNH). E. Cautin Prov., Bellavista (CAS). F, G. T. canaliculus. F. Malleco Prov., P. N. Nahuelbuta (FMNH). G. Arauco Prov., Caramavida (NMNH). H-I. T. duponchelii. H. Talca Prov., Altos de Vilches (MNHN). I. Arauco Prov., Caramavida (MCZ). J, K. T. minimucro. J. Cautin Prov., Bellavista (EMEC). K. Llanquihue Prov. P. N. Vicente Perez Rosales (FMNH). L. T. fieldianus; Valdivia, La Union (FSCA)." figureDoi="10.3897/dez.66.38022.figure6" httpUri="https://binary.pensoft.net/fig/354029" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure6">Fig. 6G</figureCitation>
), two longer setae present at apex, plus a variable number of subapical and ventral setae.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<paragraph id="A05EEB6D1E37D7BC77A4FD3ADAC61811" pageId="0" pageNumber="147">
Female reproductive tract (
<emphasis italics="true" pageId="0" pageNumber="147">n</emphasis>
= 1). Bursa copulatrix columnar, broadly rounded apically, length 1.75
<normalizedToken originalValue="×">x</normalizedToken>
<emphasis id="AB91D3BC8E968BBEC9FF65DEBD664341" italics="true" pageId="0" pageNumber="147">n</emphasis>
= 1). Bursa copulatrix columnar, broadly rounded apically, length 1.75
<normalizedToken id="8FA51629C08AFC6B469032483931FAFD" originalValue="×">x</normalizedToken>
breadth, compressed under microslide cover slip (
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female reproductive tract and gonocoxae of abdominal segment IX, ventral view. A. T. montagnei; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). Spermatheca disassociated from bursal assemblage. B. T. giraudyi; Llanquihue Prov., P. N. Vicente Perez Rosales (FMNH). C. T. robustus; Malleco Prov., M. N. Contulmo (FMNH). D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta (FMNH). E. T. duponchelii; Arauco Prov., Caramavida (MCZ). F. T. minimucro; Valdivia Prov., P. N. Oncol (EMEC). For abbreviations used to label structures see Table 1. Vertical scale bars: 0.5 mm." figureDoi="10.3897/dez.66.38022.figure7" httpUri="https://binary.pensoft.net/fig/354030" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure7">Fig. 7D</figureCitation>
<figureCitation id="C86096C77FC622D187FAC06219745DE9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Female reproductive tract and gonocoxae of abdominal segment IX, ventral view. A. T. montagnei; Petorca Prov., Q. Tigre Zapallar, Catapilco (MNHN). Spermatheca disassociated from bursal assemblage. B. T. giraudyi; Llanquihue Prov., P. N. Vicente Perez Rosales (FMNH). C. T. robustus; Malleco Prov., M. N. Contulmo (FMNH). D. T. canaliculus; Malleco Prov., P. N. Nahuelbuta (FMNH). E. T. duponchelii; Arauco Prov., Caramavida (MCZ). F. T. minimucro; Valdivia Prov., P. N. Oncol (EMEC). For abbreviations used to label structures see Table 1. Vertical scale bars: 0.5 mm." figureDoi="10.3897/dez.66.38022.figure7" httpUri="https://binary.pensoft.net/fig/354030" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure7">Fig. 7D</figureCitation>
), bursal walls thin, nearly transparent; spermatheca globose; spermathecal gland elongate-ovoid; spermathecal gland duct entering spermathecal duct basad spermathecal reservoir; basal gonocoxite 1 with one large apical fringe seta, a second small seta also present unilaterally (
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Left gonocoxa (except where noted), ventral view, for Tropopterus spp. Specimen information is parallel to that in Figure 7 A-F. For abbreviations used to label structures see Table 1. A. T. montagnei. B. T. giraudyi. C. T. robustus. D. T. canaliculus. E. T. duponchelii. F. T. minimucro (right gonocoxa)." figureDoi="10.3897/dez.66.38022.figure8" httpUri="https://binary.pensoft.net/fig/354031" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure8">Fig. 8D</figureCitation>
<figureCitation id="750F5BB47259C2908066C5DDF1332864" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Left gonocoxa (except where noted), ventral view, for Tropopterus spp. Specimen information is parallel to that in Figure 7 A-F. For abbreviations used to label structures see Table 1. A. T. montagnei. B. T. giraudyi. C. T. robustus. D. T. canaliculus. E. T. duponchelii. F. T. minimucro (right gonocoxa)." figureDoi="10.3897/dez.66.38022.figure8" httpUri="https://binary.pensoft.net/fig/354031" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure8">Fig. 8D</figureCitation>
); apical gonocoxite 2 falciform, base extended laterally, with two lateral ensiform setae, one dorsal ensiform setae, and two apical nematiform setae.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<paragraph id="BF8E9CAE367BA8B2753283129511EB55" pageId="0" pageNumber="147">
Holotype male dissected (CAS): Crest of Sierra / Nahuelbuta / Elev. 1200 m // W. of Angol / CHILE, I-3-51 // Ross and / Michelbacher / Collectors // dissection vial // Tropopterus / canaliculus / Liebherr ♂4 // HOLOTYPE / Tropopterus / canaliculus / J.K. Liebherr 2019 (black-margined red label). The type locality is in Malleco Prov., with geographic coordinates estimated to be
<geoCoordinate degrees="37" direction="south" minutes="47.76" orientation="latitude" precision="9" value="-37.796">37°47.76'S</geoCoordinate>
<geoCoordinate id="4F8846209246E35A9A9854B68791FDC3" degrees="37" direction="south" minutes="47.76" orientation="latitude" precision="9" value="-37.796">37°47.76'S</geoCoordinate>
,
<geoCoordinate degrees="73" direction="west" minutes="02.27" orientation="longitude" precision="9" value="-73.037834">73°02.27'W</geoCoordinate>
<geoCoordinate id="37704A79B20AD482C7EFDDDE7BFC31E6" degrees="73" direction="west" minutes="02.27" orientation="longitude" precision="9" value="-73.037834">73°02.27'W</geoCoordinate>
.
</paragraph>
<paragraph pageId="0" pageNumber="147">
<paragraph id="D9EEE1FD7D800F04E7ACC77C55FC2CAF" pageId="0" pageNumber="147">
Paratypes. Chile: Arauco Prov., San Alfonso, Caramavida, above,
<geoCoordinate degrees="37" direction="south" minutes="42.75" orientation="latitude" precision="9" value="-37.7125">37°42.75'S</geoCoordinate>
<geoCoordinate id="504E1AC31C79D2C6D88179B1C6A02945" degrees="37" direction="south" minutes="42.75" orientation="latitude" precision="9" value="-37.7125">37°42.75'S</geoCoordinate>
,
<geoCoordinate degrees="73" direction="west" minutes="09.00" orientation="longitude" precision="9" value="-73.15">73°09.00'W</geoCoordinate>
<geoCoordinate id="19F62CF758B0457E76944FF1233A8986" degrees="73" direction="west" minutes="09.00" orientation="longitude" precision="9" value="-73.15">73°09.00'W</geoCoordinate>
, 16-17-x-1969, Flint &amp; Barria (NMNH, 1). Malleco Prov., P. N. Nahuelbuta, lot DRM 06.046, DRM DNA2200,
<geoCoordinate degrees="37" direction="south" minutes="48.98" orientation="latitude" precision="9" value="-37.816334">37°48.98'S</geoCoordinate>
<geoCoordinate id="0074F364A07E2D049EA04C4C199678EE" degrees="37" direction="south" minutes="48.98" orientation="latitude" precision="9" value="-37.816334">37°48.98'S</geoCoordinate>
,
<geoCoordinate degrees="73" direction="west" minutes="05.7" orientation="longitude" precision="92" value="-73.095">73°05.7'W</geoCoordinate>
<geoCoordinate id="DE36CD466979444F5AED6E2CA89FF14D" degrees="73" direction="west" minutes="05.7" orientation="longitude" precision="92" value="-73.095">73°05.7'W</geoCoordinate>
, 1312 m el., 7-i-2006, Maddison (OSAC, 1), Los Portones entrance, 4.5 km W, emergent
<taxonomicName class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Nothofagus</emphasis>
<taxonomicName id="673071DD8CB087235C878C73AB12D121" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="BB92A7E1251646B7B0591ABBE50DDF36" italics="true" pageId="0" pageNumber="147">Nothofagus</emphasis>
</taxonomicName>
spp.,
<taxonomicName class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria aurucana" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="aurucana">
<emphasis italics="true" pageId="0" pageNumber="147">Araucaria aurucana</emphasis>
<taxonomicName id="FD32D2B41B84B62037BB44B6C3D42968" class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria aurucana" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="aurucana">
<emphasis id="6FE488300B44FE007BD55398BEA57654" italics="true" pageId="0" pageNumber="147">Araucaria aurucana</emphasis>
</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea" order="Poales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Chusquea</emphasis>
<taxonomicName id="0987F7FED62834B7E3FD375B0F9C93EC" class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea" order="Poales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="B77A81030B395BF043C5C0EBF856295C" italics="true" pageId="0" pageNumber="147">Chusquea</emphasis>
</taxonomicName>
understory, lot 975, pyr.-fogging old logs,
<geoCoordinate degrees="37" direction="south" minutes="49.25" orientation="latitude" precision="9" value="-37.820835">37°49.25'S</geoCoordinate>
<geoCoordinate id="CA885D665E16B4A11E3C53A25FC0B2E6" degrees="37" direction="south" minutes="49.25" orientation="latitude" precision="9" value="-37.820835">37°49.25'S</geoCoordinate>
,
<geoCoordinate degrees="72" direction="west" minutes="59.82" orientation="longitude" precision="9" value="-72.997">72°59.82'W</geoCoordinate>
<geoCoordinate id="301A91084A2BBDBD7ACC7AC5147BA004" degrees="72" direction="west" minutes="59.82" orientation="longitude" precision="9" value="-72.997">72°59.82'W</geoCoordinate>
, 1300 m el., 7-ii-1997, Newton &amp; Thayer (FMNH etoh, 1), Guarderia Pichinahuel, E of,
<taxonomicName genus="Araucaria-Nothofagus" lsidName="Araucaria-Nothofagus dombeyi" pageId="0" pageNumber="147" rank="species" species="dombeyi">
<emphasis italics="true" pageId="0" pageNumber="147">Araucaria-Nothofagus dombeyi</emphasis>
<taxonomicName id="7C629E89A5BA81C5F4D7CE35B351D681" genus="Araucaria-Nothofagus" lsidName="Araucaria-Nothofagus dombeyi" pageId="0" pageNumber="147" rank="species" species="dombeyi">
<emphasis id="F7595A1AB92E5BB9ABC52E34C574BD1D" italics="true" pageId="0" pageNumber="147">Araucaria-Nothofagus dombeyi</emphasis>
</taxonomicName>
w/
<taxonomicName class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea" order="Poales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Chusquea</emphasis>
<taxonomicName id="DCA824E6584D305D36A83A555B9B930D" class="Liliopsida" family="Poaceae" genus="Chusquea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Chusquea" order="Poales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="3162AD4C6A4E344BE267428228639CEE" italics="true" pageId="0" pageNumber="147">Chusquea</emphasis>
</taxonomicName>
bamboo, lot 1054, in rotten logs,
<geoCoordinate degrees="37" direction="south" minutes="48.20" orientation="latitude" precision="9" value="-37.803333">37°48.20'S</geoCoordinate>
<geoCoordinate id="248EF589DD47C36984570D227563FC09" degrees="37" direction="south" minutes="48.20" orientation="latitude" precision="9" value="-37.803333">37°48.20'S</geoCoordinate>
,
<geoCoordinate degrees="73" direction="west" minutes="01.41" orientation="longitude" precision="9" value="-73.0235">73°01.41'W</geoCoordinate>
<geoCoordinate id="5F3997682CC6C73AA4E328FBF6E0C32E" degrees="73" direction="west" minutes="01.41" orientation="longitude" precision="9" value="-73.0235">73°01.41'W</geoCoordinate>
, 1290 m el., 7-xii-2002, Clarke &amp; Solodovnikov (FMNH, 5), lot 1054, pyr. fogging live
<taxonomicName class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Araucaria</emphasis>
<taxonomicName id="90413EE10EDD8F23FC14A4E9DB01B77C" class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="6E09C8365D10125E4320A8BDF0A9F65F" italics="true" pageId="0" pageNumber="147">Araucaria</emphasis>
</taxonomicName>
,
<geoCoordinate degrees="37" direction="south" minutes="48.20" orientation="latitude" precision="9" value="-37.803333">37°48.20'S</geoCoordinate>
<geoCoordinate id="FAAF89A88E9473FB0D868474F96B1246" degrees="37" direction="south" minutes="48.20" orientation="latitude" precision="9" value="-37.803333">37°48.20'S</geoCoordinate>
,
<geoCoordinate degrees="73" direction="west" minutes="01.41" orientation="longitude" precision="9" value="-73.0235">73°01.41'W</geoCoordinate>
<geoCoordinate id="61AE60BDBCFB861A3568F701942A6033" degrees="73" direction="west" minutes="01.41" orientation="longitude" precision="9" value="-73.0235">73°01.41'W</geoCoordinate>
, 1290 m el., 24-xii-2002, Newton &amp; Solodovnikov (FMNH, 1), lot 1054, FMHD#2002-094, Berlese debris under bark large
<taxonomicName class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Araucaria</emphasis>
<taxonomicName id="551D13F19FD49907EC9882CAEB1778A6" class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="380B766CBC7FFAEE93A30B0168D701DB" italics="true" pageId="0" pageNumber="147">Araucaria</emphasis>
</taxonomicName>
log,
<geoCoordinate degrees="37" direction="south" minutes="48.20" orientation="latitude" precision="9" value="-37.803333">37°48.20'S</geoCoordinate>
<geoCoordinate id="6581607932F38AF50AAAED9E22131CE2" degrees="37" direction="south" minutes="48.20" orientation="latitude" precision="9" value="-37.803333">37°48.20'S</geoCoordinate>
,
<geoCoordinate degrees="73" direction="west" minutes="01.41" orientation="longitude" precision="9" value="-73.0235">73°01.41'W</geoCoordinate>
<geoCoordinate id="8107D8A857D7A6F8920F0EAA8B0523FB" degrees="73" direction="west" minutes="01.41" orientation="longitude" precision="9" value="-73.0235">73°01.41'W</geoCoordinate>
, 1290 m el., 24-xii-2002, Newton &amp; Solodovnikov (FMNH, 4), Los Portones entrance, 2.3 km W,
<taxonomicName class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus dombeyi" order="Fagales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="dombeyi">
<emphasis italics="true" pageId="0" pageNumber="147">Nothofagus dombeyi</emphasis>
<taxonomicName id="BF257EAE069B6F0F282A1910CAA9662F" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus dombeyi" order="Fagales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="species" species="dombeyi">
<emphasis id="7D30CCAC108A16C3F63A1EEB416F4447" italics="true" pageId="0" pageNumber="147">Nothofagus dombeyi</emphasis>
</taxonomicName>
+
<taxonomicName lsidName="antarctica" pageId="0" pageNumber="147" rank="species" species="antarctica">? antarctica</taxonomicName>
<taxonomicName id="53FE750F9B47E6F0112142EA38D249A5" lsidName="antarctica" pageId="0" pageNumber="147" rank="species" species="antarctica">? antarctica</taxonomicName>
, mostly open understory, lot 1057, pyr. fogging old
<taxonomicName class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Nothofagus</emphasis>
<taxonomicName id="2675FC3E2342E255EAE12C95002958B8" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="1D324798CB9BC79428EB4DD445AD33EA" italics="true" pageId="0" pageNumber="147">Nothofagus</emphasis>
</taxonomicName>
logs,
<geoCoordinate degrees="37" direction="south" minutes="49.41" orientation="latitude" precision="9" value="-37.8235">37°49.41'S</geoCoordinate>
<geoCoordinate id="B8D9304E606C50D0B36E5A75457F53A1" degrees="37" direction="south" minutes="49.41" orientation="latitude" precision="9" value="-37.8235">37°49.41'S</geoCoordinate>
,
<geoCoordinate degrees="72" direction="west" minutes="58.95" orientation="longitude" precision="9" value="-72.9825">72°58.95'W</geoCoordinate>
<geoCoordinate id="5E486BBAC2A0B642DA5EE5D438DB81E4" degrees="72" direction="west" minutes="58.95" orientation="longitude" precision="9" value="-72.9825">72°58.95'W</geoCoordinate>
, 1150 m el., 25-xii-2002, Newton, Thayer &amp; Chani (FMNH, 6).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="etymology">
<paragraph pageId="0" pageNumber="147">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="147">
<subSubSection id="445C99C574CD7DD9F92831DF2BDCEF34" pageId="0" pageNumber="147" type="etymology">
<paragraph id="C1CA9F9FAAC7FC21FBA627FBDD4D9FA6" pageId="0" pageNumber="147">Etymology.</paragraph>
<paragraph id="0EC519ECCCD13816234650CDADCC84E7" pageId="0" pageNumber="147">
The species epithet
<taxonomicName lsidName="canaliculus" pageId="0" pageNumber="147" rank="species" species="canaliculus">canaliculus</taxonomicName>
<taxonomicName id="3561020FD58BDFE4FAD527981714BC56" lsidName="canaliculus" pageId="0" pageNumber="147" rank="species" species="canaliculus">canaliculus</taxonomicName>
references the narrow, deep median prosternal depression present in these beetles. The epithet is to be treated as a noun.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="147" type="distribution">
<paragraph pageId="0" pageNumber="147">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="147">
<subSubSection id="CB211C8B94279355F39C67F489B55994" pageId="0" pageNumber="147" type="distribution">
<paragraph id="DD97482C58501FBDA9E0986A9F2F0827" pageId="0" pageNumber="147">Distribution and habitat.</paragraph>
<paragraph id="4945B885CEDB84534B1D2910CA220676" pageId="0" pageNumber="147">
This species' distribution lies near 38°S (
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I. Geographic distributions of Tropopterus spp. A. T. montagnei. B. T. giraudyi. C. T. peckorum. D. T. robustus. E. T. canaliculus. F. T. duponchelii. G. T. trisinuatus. H. T. minimucro. I. T. fieldianus. J. Geographical distributions of Tropopterus spp., labeled as per figures above, hierarchically arranged by phylogenetic relationships of the taxon cladogram, Fig. 10." figureDoi="10.3897/dez.66.38022.figure9" httpUri="https://binary.pensoft.net/fig/354032" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure9">Fig. 9E</figureCitation>
<figureCitation id="CE5440EBC77CE1C2519269334D3D4D4F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-I. Geographic distributions of Tropopterus spp. A. T. montagnei. B. T. giraudyi. C. T. peckorum. D. T. robustus. E. T. canaliculus. F. T. duponchelii. G. T. trisinuatus. H. T. minimucro. I. T. fieldianus. J. Geographical distributions of Tropopterus spp., labeled as per figures above, hierarchically arranged by phylogenetic relationships of the taxon cladogram, Fig. 10." figureDoi="10.3897/dez.66.38022.figure9" httpUri="https://binary.pensoft.net/fig/354032" pageId="0" pageNumber="147" tableDoi="10.3897/dez.66.38022.figure9">Fig. 9E</figureCitation>
) in the Northern Valdivian entomofaunal province (
<bibRefCitation author="O'Brien, CW" journalOrPublisher="Entomological News" pageId="0" pageNumber="147" pagination="197 - 207" refId="B53" refString="O'Brien, CW, 1971. The biogeography of Chile through entomofaunal regions. Entomological News 82: 197 - 207" title="The biogeography of Chile through entomofaunal regions." volume="82" year="1971">
<normalizedToken originalValue="OBrien">O'Brien</normalizedToken>
<bibRefCitation id="19440080B3084B46B64CE774CE5C6D52" author="O'Brien, CW" journalOrPublisher="Entomological News" pageId="0" pageNumber="147" pagination="197 - 207" refId="B53" refString="O'Brien, CW, 1971. The biogeography of Chile through entomofaunal regions. Entomological News 82: 197 - 207" title="The biogeography of Chile through entomofaunal regions." volume="82" year="1971">
<normalizedToken id="86A6F9A4C2897FA1EBD977852D5C4D1E" originalValue="OBrien">O'Brien</normalizedToken>
1971
</bibRefCitation>
). Most records are from forest-floor microhabitats, either by pyrethrin fogging of old, rotten
<taxonomicName class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Nothofagus</emphasis>
<taxonomicName id="7023B8A483B279A6C1AD242423A60E54" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Nothofagus" order="Fagales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="562982629B31CF2D4646AB8375EE65D5" italics="true" pageId="0" pageNumber="147">Nothofagus</emphasis>
</taxonomicName>
logs, or in Berlese siftate from under bark of a large
<taxonomicName class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Araucaria</emphasis>
<taxonomicName id="7DD442366D1040C06ED448A0E18A67E5" class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="7362FA50CAAD4D1D0AEE5BDF5CAE04F4" italics="true" pageId="0" pageNumber="147">Araucaria</emphasis>
</taxonomicName>
log. One specimen was collected via pyrethrin spraying onto a live
<taxonomicName class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="147">Araucaria</emphasis>
<taxonomicName id="34C4AC54F93D725B81C9B857646F9E79" class="Pinopsida" family="Araucariaceae" genus="Araucaria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Araucaria" order="Pinales" pageId="0" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="33BAF5A6202438641916FD21531C57F2" italics="true" pageId="0" pageNumber="147">Araucaria</emphasis>
</taxonomicName>
trunk. This species was collected with
<taxonomicName class="Insecta" family="Leiodidae" genus="Anaballetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anaballetus chilensis" order="Coleoptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis italics="true" pageId="0" pageNumber="147">Anaballetus chilensis</emphasis>
<taxonomicName id="A48FA993ED4CA31F8EEB7AC3B1269D69" class="Insecta" family="Leiodidae" genus="Anaballetus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anaballetus chilensis" order="Coleoptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="chilensis">
<emphasis id="D07A2FE2BF53FDC70453BC0C066BDE76" italics="true" pageId="0" pageNumber="147">Anaballetus chilensis</emphasis>
</taxonomicName>
at two sites (
<bibRefCitation DOI="https://doi.org/10.1515/aemnp-2017-0061" author="Newton, AF" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="147" pagination="121 - 140" refId="B49" refString="Newton, AF, Svec, Z, Fikacek, M, 2017. A new genus and two new species of Leiodinae from Chile, with keys to world genera of Sogdini and Leiodinae from Chile and Argentina (Coleoptera: Leiodidae). Acta Entomologica Musei Nationalis Pragae 57: 121 - 140" title="A new genus and two new species of Leiodinae from Chile, with keys to world genera of Sogdini and Leiodinae from Chile and Argentina (Coleoptera: Leiodidae)." url="https://doi.org/10.1515/aemnp-2017-0061" volume="57" year="2017">Newton et al. 2017</bibRefCitation>
<bibRefCitation id="5BA13C3530214469319E7A9DEBB9BC79" DOI="https://doi.org/10.1515/aemnp-2017-0061" author="Newton, AF" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="147" pagination="121 - 140" refId="B49" refString="Newton, AF, Svec, Z, Fikacek, M, 2017. A new genus and two new species of Leiodinae from Chile, with keys to world genera of Sogdini and Leiodinae from Chile and Argentina (Coleoptera: Leiodidae). Acta Entomologica Musei Nationalis Pragae 57: 121 - 140" title="A new genus and two new species of Leiodinae from Chile, with keys to world genera of Sogdini and Leiodinae from Chile and Argentina (Coleoptera: Leiodidae)." url="https://doi.org/10.1515/aemnp-2017-0061" volume="57" year="2017">Newton et al. 2017</bibRefCitation>
: Chile: Malleco Prov., P. N. Nahuelbuta, 2.3 km W Los Portones entrance, FMNH, and P. N. Nahuelbuta, E of Guarderia Pichinahuel, FMNH).
</paragraph>
<caption doi="10.3897/dez.66.38022.figure10" httpUri="https://binary.pensoft.net/fig/354033" pageId="0" pageNumber="147" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="147">
<emphasis bold="true" pageId="0" pageNumber="147">Figure 10.</emphasis>
<caption id="586AF83AD8FD5CD1D7AD35A3C5922585" doi="10.3897/dez.66.38022.figure10" httpUri="https://binary.pensoft.net/fig/354033" pageId="0" pageNumber="147" start="Figure 10" startId="F10">
<paragraph id="8A881932232CEA8E72BB12AC4B4E2A26" pageId="0" pageNumber="147">
<emphasis id="17225A0E81326CEE88A8C161400269FD" bold="true" pageId="0" pageNumber="147">Figure 10.</emphasis>
Cladogram returned under all cladistic protocols described in text; tree length = 97, CI = 57, RI = 47, fast character optimization shown. Outgroup choice of
<taxonomicName authority="Liebherr, 2020" authorityName="Liebherr" authorityYear="2020" class="Insecta" family="Carabidae" genus="Pharetis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pharetis thayerae" order="Coleoptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="thayerae">
<emphasis italics="true" pageId="0" pageNumber="147">Pharetis thayerae</emphasis>
<taxonomicName id="5077793F23EDDDBBE66BE36AE5C282B8" class="Insecta" family="Carabidae" genus="Pharetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pharetis thayerae" order="Coleoptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="thayerae">
<emphasis id="189584FF5B12649F864EE0E9EF68A8F7" italics="true" pageId="0" pageNumber="147">Pharetis thayerae</emphasis>
</taxonomicName>
based on results in
<bibRefCitation author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="147" refId="B39" refString="Liebherr, JK, 2020. Phylogenetic placement of the Australian Pharetis, gen. nov. and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera : Carabidae). Invertebrate Systematics. [in press]" title="Phylogenetic placement of the Australian Pharetis, gen. nov. and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera: Carabidae). Invertebrate Systematics. [in press]" year="2020">Liebherr (2020)</bibRefCitation>
<bibRefCitation id="AB8CB1B0BE0E5ECA7CAFE4C9A7BC988C" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="147" refId="B39" refString="Liebherr, JK, 2020. Phylogenetic placement of the Australian Pharetis, gen. nov. and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera : Carabidae). Invertebrate Systematics. [in press]" title="Phylogenetic placement of the Australian Pharetis, gen. nov. and Spherita, gen. nov., in a revised classification of the circum-Antarctic Moriomorphini (Coleoptera: Carabidae). Invertebrate Systematics. [in press]" year="2020">Liebherr (2020)</bibRefCitation>
. Geographic region abbreviations after taxon name include: eastern Australia (EOZ); Santiagan entomofaunal province (San); Northern Valdivian entomofaunal province (NV); Southern Valdivian Entomofaunal province (SV).
</paragraph>
</caption>