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<mods:title>Diversity and phylogeny of the extinct wasp subfamily Lancepyrinae (Hymenoptera, Bethylidae) revealed by mid-Cretaceous Burmese amber</mods:title>
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<mods:title id="CD942551EECE27AEB93C23966D4F50C6">Diversity and phylogeny of the extinct wasp subfamily Lancepyrinae (Hymenoptera, Bethylidae) revealed by mid-Cretaceous Burmese amber</mods:title>
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<mods:namePart>Brazidec, Manuel</mods:namePart>
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<mods:affiliation>Univ. Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000 Rennes, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:affiliation id="33EFA0FA70D56EF64416841705D36078">Univ. Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000 Rennes, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:namePart>Legendre, Frederic</mods:namePart>
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<mods:affiliation>Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, Sorbonne Universite, EPHE, Universite des Antilles, CP 50, 57, rue Cuvier, 75005 Paris, France</mods:affiliation>
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<mods:namePart id="65DFBAC1410E45718FCF1A5A3B7652A8">Legendre, Frederic</mods:namePart>
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<mods:affiliation id="AA37D53E7E00EF634CD6A3498379ABD0">Institut de Systematique, Evolution, Biodiversite (ISYEB), Museum national d'Histoire naturelle, CNRS, Sorbonne Universite, EPHE, Universite des Antilles, CP 50, 57, rue Cuvier, 75005 Paris, France</mods:affiliation>
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<mods:affiliation>Univ. Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000 Rennes, France</mods:affiliation>
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<treatment LSID="urn:lsid:zoobank.org:act:EB5CF079-C86A-4D97-9631-4B6BCF3E52B7" httpUri="http://treatment.plazi.org/id/9E272910B84F5F08A9776831020B0869" lastPageNumber="345" pageId="0" pageNumber="345">
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<taxonomicName LSID="https://zoobank.org/EB5CF079-C86A-4D97-9631-4B6BCF3E52B7" authority="Brazidec & Legendre & Perrichot, 2023" authorityName="Brazidec & Legendre & Perrichot" authorityYear="2023" family="Bethylidae" genus="Azepyris" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Azepyris delamarrei" order="Hymenoptera" pageId="0" pageNumber="345" rank="species" species="delamarrei" status="sp. nov.">Azepyris delamarrei</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="345">sp. nov.</taxonomicNameLabel>
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||||
<treatment id="9E272910B84F5F08A9776831020B0869" ID-GBIF-Taxon="209487366" LSID="urn:lsid:zoobank.org:act:EB5CF079-C86A-4D97-9631-4B6BCF3E52B7" httpUri="http://treatment.plazi.org/id/9E272910B84F5F08A9776831020B0869" lastPageNumber="345" pageId="0" pageNumber="345" scope_family="Bethylidae" scope_order="Hymenoptera" scope_subFamily="Lancepyrinae" scope_superFamily="Chrysidoidea">
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<subSubSection id="06642073F6C4EDF7B2941F7F857177F5" pageId="0" pageNumber="345" type="nomenclature">
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<paragraph id="A900E2266AD35D16CF8C70B2BC6BEC65" pageId="0" pageNumber="345">
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<taxonomicName id="EB40D89ED34C17CD5097B0770D3D5DF3" LSID="https://zoobank.org/EB5CF079-C86A-4D97-9631-4B6BCF3E52B7" authority="Brazidec & Legendre & Perrichot, 2023" authorityName="Brazidec & Legendre & Perrichot" authorityYear="2023" class="Insecta" family="Bethylidae" genus="Azepyris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Azepyris delamarrei" order="Hymenoptera" pageId="0" pageNumber="345" phylum="Arthropoda" rank="species" species="delamarrei" status="sp. nov.">Azepyris delamarrei</taxonomicName>
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<taxonomicNameLabel id="9669989B7EB7A3884AFB7E3F7E79A6C3" pageId="0" pageNumber="345">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection pageId="0" pageNumber="345" type="description">
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<paragraph pageId="0" pageNumber="345">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Azepyris delamarrei gen. et sp. nov., holotype female number SNHM- 6001. A habitus in lateral view; B head and anterior mesosoma in lateral view (arrow = fore arolium); C habitus in dorsal view; D anterior mesosoma in dorsal view; E posterior mesosoma in dorsal view (arrow = lateral projection of posterior corner of metapectal-propodeal complex); F line drawing of fore wing. Scale bars: 1 mm (A, C, F); 0.5 mm (B, D, E)." figureDoi="10.3897/asp.81.e96737.figure1" httpUri="https://binary.pensoft.net/fig/833456" pageId="0" pageNumber="345">Fig. 1</figureCitation>
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<subSubSection id="0D61975AC8358D655BD4133E47BA95B2" pageId="0" pageNumber="345" type="description">
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<paragraph id="19C29E83D9185B0FB625C78B20BD784B" pageId="0" pageNumber="345">
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<figureCitation id="3C56CEA0F34538924C0998DB26F5D97E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Azepyris delamarrei gen. et sp. nov., holotype female number SNHM- 6001. A habitus in lateral view; B head and anterior mesosoma in lateral view (arrow = fore arolium); C habitus in dorsal view; D anterior mesosoma in dorsal view; E posterior mesosoma in dorsal view (arrow = lateral projection of posterior corner of metapectal-propodeal complex); F line drawing of fore wing. Scale bars: 1 mm (A, C, F); 0.5 mm (B, D, E)." figureDoi="10.3897/asp.81.e96737.figure1" httpUri="https://binary.pensoft.net/fig/833456" pageId="0" pageNumber="345">Fig. 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="345" type="etymology">
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<paragraph pageId="0" pageNumber="345">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="345">
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<subSubSection id="A8A795E2021885CFAFA03979F55DBD35" pageId="0" pageNumber="345" type="etymology">
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<paragraph id="4BB673564FA1F4FD67BD2EEAB4164D16" pageId="0" pageNumber="345">Etymology.</paragraph>
|
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<paragraph id="1FC6943A3DB4C191798684838BEADAF6" pageId="0" pageNumber="345">
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||||
The specific epithet is a patronym honouring Yann Delamarre, a student and the senior
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<normalizedToken id="7E111EAAA9850A9E51FD2FBB17F6EDD1" originalValue="author’s">author's</normalizedToken>
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||||
fellow from the palaeontology program at the University of Rennes. The specific epithet is to be treated as a noun in the genitive case.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="345" type="materials_examined">
|
||||
<paragraph pageId="0" pageNumber="345">Material studied.</paragraph>
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||||
<paragraph pageId="0" pageNumber="345">
|
||||
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Mueller" country="Germany" location="Germany" specimenCount="1" typeStatus="Holotype">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="345">
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||||
<typeStatus>Holotype</typeStatus>
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<subSubSection id="15A89499D155648E3E267393293A3ECC" pageId="0" pageNumber="345" type="materials_examined">
|
||||
<paragraph id="85D0DD1441E9CE9173743F52C319759B" pageId="0" pageNumber="345">Material studied.</paragraph>
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||||
<paragraph id="4B636476965FA5D5551230C156AE8E37" pageId="0" pageNumber="345">
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||||
<materialsCitation id="C6C7CECCE1DCEE0B67A78993C6DC937C" ID-GBIF-Occurrence="4071676302" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Mueller" country="Germany" location="Germany" specimenCount="1" typeStatus="Holotype">
|
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<emphasis id="1C4BD22A0985B5474E15361BE23C8896" bold="true" italics="true" pageId="0" pageNumber="345">
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<typeStatus id="CDCAB2C03E211D7848FFB3FB20618D3C">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
SNHM-6001, a complete female; housed in the paleontological collection of the Staatliches Naturhistorisches Museum Braunschweig,
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<collectingCountry name="Germany">Germany</collectingCountry>
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||||
<collectingCountry id="CF5040CEC6D4D457820B04DC86DE6757" name="Germany">Germany</collectingCountry>
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(SMNH, coll.
|
||||
<normalizedToken originalValue="Müller">
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||||
<collectorName>Mueller</collectorName>
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<normalizedToken id="9416C7394511B15A7DCADD41CE6B3B6D" originalValue="Müller">
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||||
<collectorName id="F1E5F6B6AE003EBC16AEB91D17E90946">Mueller</collectorName>
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</normalizedToken>
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).
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||||
</materialsCitation>
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</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="345" type="type locality">
|
||||
<paragraph pageId="0" pageNumber="345">Type locality and horizon.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="345">Hkamti site, Hkamti district, Sagaing Region, Myanmar; early Albian, ca. 110 Ma, Early Cretaceous.</paragraph>
|
||||
<subSubSection id="A6920750B119B5907096500C3F214B2F" pageId="0" pageNumber="345" type="type locality">
|
||||
<paragraph id="6571CAF38C1089BA8C51C19619A61B47" pageId="0" pageNumber="345">Type locality and horizon.</paragraph>
|
||||
<paragraph id="94388236BD6A19527BB165107BDA16A7" pageId="0" pageNumber="345">Hkamti site, Hkamti district, Sagaing Region, Myanmar; early Albian, ca. 110 Ma, Early Cretaceous.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="345" type="diagnosis">
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||||
<paragraph pageId="0" pageNumber="345">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="345">As for genus.</paragraph>
|
||||
<subSubSection id="435ECBFCBB87A00D9F8CC2259BB5C348" pageId="0" pageNumber="345" type="diagnosis">
|
||||
<paragraph id="CB6DB3DF2B9239F6E903347204685871" pageId="0" pageNumber="345">Diagnosis.</paragraph>
|
||||
<paragraph id="D75978A37867D1B6BDF9CBA55AC10D66" pageId="0" pageNumber="345">As for genus.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="345" type="description">
|
||||
<paragraph pageId="0" pageNumber="345">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="345">
|
||||
<subSubSection id="1C6E00D3FBB415B35F9AA1898D4BBD76" pageId="0" pageNumber="345" type="description">
|
||||
<paragraph id="80F12BD5EADE546E35552EE3F7CA2D67" pageId="0" pageNumber="345">Description.</paragraph>
|
||||
<paragraph id="A6F1ECF8276911AE96D49B4393F20377" pageId="0" pageNumber="345">
|
||||
Body rather depressed, elongate, poorly pubescent (length 5.35 mm). -
|
||||
<emphasis bold="true" pageId="0" pageNumber="345">Head</emphasis>
|
||||
<emphasis id="4CFC854588EAFF52AFC19F3FEB81B06F" bold="true" pageId="0" pageNumber="345">Head</emphasis>
|
||||
prognathous, longer than wide; LH: 1.04 mm, WH: ca. 0.70 mm, HE: 0.60 mm, VOL: 0.20 mm; frons flat, punctate; compound eye elliptical, longer than high, not covering head length, located on anterior half of head, closer to mandible than to occipital carina; clypeus with median lobe rather projecting forward, lateral lobe poorly developed; mandible long, with three teeth, apical tooth longest; antenna filiform; scape 2.17 times as long as pedicel (length 0.26 mm); flagellomeres 1-10 cylindrical, all longer than wide (length 0.09-0.10 mm); flagellomere 11 longest, tapering at apex: ocelli forming short triangle, anterior ocellus not crossing supra-ocular line; occipital carina present, complete, forming weak arch. -
|
||||
<emphasis bold="true" pageId="0" pageNumber="345">Mesosoma</emphasis>
|
||||
<emphasis id="BA9D402C34BFC5F2E07653B522161D7F" bold="true" pageId="0" pageNumber="345">Mesosoma</emphasis>
|
||||
flattened, with dorsum smooth (length 1.62 mm); propleuron slightly visible in dorsal view,
|
||||
<normalizedToken originalValue="‘neck-shaped’">'neck-shaped'</normalizedToken>
|
||||
<normalizedToken id="F73767C82A9F46FF4126AA49ED4FB1D8" originalValue="‘neck-shaped’">'neck-shaped'</normalizedToken>
|
||||
; dorsal pronotal area 1.56 times as long as anteromesoscutum (length 0.50 mm), narrow, anterior flange developed, lateral margin slightly incurved, posterior margin concave; anteromesocutum wider than dorsal pronotal area, posterior margin straight; notaulus deeply impressed, not reaching posterior margin of anteromesoscutum, convergent; parapsidal signum poorly marked; mesoscuto-mesoscutellar suture with reniform sulcus connecting lateral foveae; mesoscutellum subquadrate, posterior margin slightly convex; metanotum rather developed, overlapping mesoscutellum posteriorly
|
||||
<emphasis italics="true" pageId="0" pageNumber="345">sensu</emphasis>
|
||||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4489.1.1" author="Azevedo, CO" journalOrPublisher="Zootaxa" pageId="0" pageNumber="345" pagination="1 - 294" refId="B3" refString="Azevedo, CO, Alencar, IDCC, Ramos, MS, Barbosa, DN, Colombo, WD, Vargas, JM, Lim, J, 2018. Global guide of the flat wasps (Hymenoptera, Bethylidae). Zootaxa 4489: 1 - 294, DOI: https://doi.org/10.11646/zootaxa.4489.1.1" title="Global guide of the flat wasps (Hymenoptera, Bethylidae)." url="https://doi.org/10.11646/zootaxa.4489.1.1" volume="4489" year="2018">Azevedo et al. (2018)</bibRefCitation>
|
||||
<emphasis id="7AE9CE3A1E10B2998EDD340E1770EFDC" italics="true" pageId="0" pageNumber="345">sensu</emphasis>
|
||||
<bibRefCitation id="FB7AB88AD6E3F038B312AC9854F29864" DOI="https://doi.org/10.11646/zootaxa.4489.1.1" author="Azevedo, CO" journalOrPublisher="Zootaxa" pageId="0" pageNumber="345" pagination="1 - 294" refId="B3" refString="Azevedo, CO, Alencar, IDCC, Ramos, MS, Barbosa, DN, Colombo, WD, Vargas, JM, Lim, J, 2018. Global guide of the flat wasps (Hymenoptera, Bethylidae). Zootaxa 4489: 1 - 294, DOI: https://doi.org/10.11646/zootaxa.4489.1.1" title="Global guide of the flat wasps (Hymenoptera, Bethylidae)." url="https://doi.org/10.11646/zootaxa.4489.1.1" volume="4489" year="2018">Azevedo et al. (2018)</bibRefCitation>
|
||||
; metapectal-propodeal complex smooth laterally, lateral marginal carina distinct, dorsal surface of metapectal-propodeal complex hardly distinguishable, apparently smooth, posterior corner with small lateral dentiform projection. Fore wing hyaline, reaching fourth metasomal segment (LFW: 2.48 mm); C, Sc+R, M+Cu, 1A tubular; 1Rs&1M angled at junction with Rs+M; Rs+M poorly pigmented, spectral; cu-a post-furcal to 1M; 2Cu pigmented, then fading; pterostigma rounded; short stub of R1 tubular distal to pterostigma: 2r-rs&Rs arising on distal half of pterostigma, long but not closing [2R1] cell. Proleg with protrochanter originating from apex of procoxa; profemur moderately thickened; tibial spur formula 1-2-2, pro- and metaspurs long, mesospurs shorter; two tarsal claws, only slightly curved; proarolium well-developed; metafemur moderately thickened; first metatarsomere as long as 2-4 combined. -
|
||||
<emphasis bold="true" pageId="0" pageNumber="345">Metasoma</emphasis>
|
||||
<emphasis id="42986E9071386CAE8BADC974338932B2" bold="true" pageId="0" pageNumber="345">Metasoma</emphasis>
|
||||
longer than mesosoma (length 2.69 mm); fusiform, tapering at apex; petiole conspicuous and narrow: six exposed tergites; T2 longest, laterally covering sternites; T1, T3, T4 and T5 subequal in length; short and narrow sting exserted.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
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