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10.
<taxonomicName LSID="http://zoobank.org/27045083-198A-4614-B95A-7786EDDA7842" class="Insecta" family="Tenebrionidae" genus="Charisius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Charisius punctatus" order="Coleoptera" pageId="17" pageNumber="286" phylum="Arthropoda" rank="species" species="punctatus">Charisius punctatus</taxonomicName>
<taxonomicNameLabel pageId="17" pageNumber="286">sp. n.</taxonomicNameLabel>
<taxonomicName id="C1285E132E3EA282227741B75ABF9EC0" LSID="http://zoobank.org/27045083-198A-4614-B95A-7786EDDA7842" genus="Charisius" lsidName="Charisius punctatus" pageId="17" pageNumber="286" rank="species" species="punctatus">Charisius punctatus</taxonomicName>
<taxonomicNameLabel id="8816E0E304E576967D90259C63EFD4B0" pageId="17" pageNumber="286">sp. n.</taxonomicNameLabel>
Figs 12, 24, 28
</paragraph>
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<subSubSection pageId="17" pageNumber="286" type="description">
<paragraph pageId="17" pageNumber="286">Description.</paragraph>
<paragraph pageId="17" pageNumber="286">Body reddish-brown; elytra testaceus with each elytron having piceous markings as follows (Fig. 12): a small circle near middle, a narrow, crescent-shaped band at apical fourth and extreme apex piceous to black (Fig. 12). Length 7.0-10.9 mm.</paragraph>
<paragraph pageId="17" pageNumber="286">Head moderately coarsely punctate on vertex; punctures separated by distance subequal to diameter of a puncture. Eyes moderately small, mean OI of 5 males 32.7 (range 30-35) and of five females 38.6 (range 37-40). Pronotum distinctly wider than long, mean PI of 11 specimens 84.6 (range 83 to 88); surface smooth, shining; punctures coarse, moderately densely, evenly distributed, separated by average distance distinctly greater than diameter of a puncture; sides sinuate near base, widest near middle than evenly convex to just before apex; disc evenly convex in cross section. Prosternum and hypomeron coarsely, densely punctate; punctures separated by distance less than diameter of a puncture. Metaventrite finely, sparsely punctate medially, punctures becoming coarser and denser approaching sides. Ventrites with punctures moderately fine, sparsely distributed; last two visible ventrites more coarsely and densely punctate. Elytra with striae moderately deeply, evenly impressed throughout; strial interstices moderately convex.</paragraph>
<paragraph pageId="17" pageNumber="286">Male. Anterior tibia triangularly widened on inner side near middle. Fifth ventrite broadly, deeply, triangularly impressed medially. Lobes of eighth sternum (Fig. 24) broad, with apical third broadly expanded, apices obliquely truncate; viewed laterally, distinctly deflexed apically. Lobes of ninth sternum short, moderately broad, with apices evenly convex. Aedeagus with apicale (Fig. 28) short, moderately narrow; sides narrowed from base to narrowly, evenly convex apex, slightly constricted medially; viewed laterally, apicale strongly curved dorsally; basale 7.1-7.9 times longer than apicale.</paragraph>
<paragraph pageId="17" pageNumber="286">Female. Elytra with sutural margin and apex entire.</paragraph>
<subSubSection id="E9076E6126720B7DAFB4EAB84E7801B7" pageId="17" pageNumber="286" type="description">
<paragraph id="8FD5A11B58CA305A8BF2F8A3AD932CB1" pageId="17" pageNumber="286">Description.</paragraph>
<paragraph id="A4C947A72BACC4321CB8684DABFF9E93" pageId="17" pageNumber="286">Body reddish-brown; elytra testaceus with each elytron having piceous markings as follows (Fig. 12): a small circle near middle, a narrow, crescent-shaped band at apical fourth and extreme apex piceous to black (Fig. 12). Length 7.0-10.9 mm.</paragraph>
<paragraph id="4EA2D4F9DDA5AFDC99A1381B86B0816D" pageId="17" pageNumber="286">Head moderately coarsely punctate on vertex; punctures separated by distance subequal to diameter of a puncture. Eyes moderately small, mean OI of 5 males 32.7 (range 30-35) and of five females 38.6 (range 37-40). Pronotum distinctly wider than long, mean PI of 11 specimens 84.6 (range 83 to 88); surface smooth, shining; punctures coarse, moderately densely, evenly distributed, separated by average distance distinctly greater than diameter of a puncture; sides sinuate near base, widest near middle than evenly convex to just before apex; disc evenly convex in cross section. Prosternum and hypomeron coarsely, densely punctate; punctures separated by distance less than diameter of a puncture. Metaventrite finely, sparsely punctate medially, punctures becoming coarser and denser approaching sides. Ventrites with punctures moderately fine, sparsely distributed; last two visible ventrites more coarsely and densely punctate. Elytra with striae moderately deeply, evenly impressed throughout; strial interstices moderately convex.</paragraph>
<paragraph id="020DABDF08C17E34EFA21FA403DDFB89" pageId="17" pageNumber="286">Male. Anterior tibia triangularly widened on inner side near middle. Fifth ventrite broadly, deeply, triangularly impressed medially. Lobes of eighth sternum (Fig. 24) broad, with apical third broadly expanded, apices obliquely truncate; viewed laterally, distinctly deflexed apically. Lobes of ninth sternum short, moderately broad, with apices evenly convex. Aedeagus with apicale (Fig. 28) short, moderately narrow; sides narrowed from base to narrowly, evenly convex apex, slightly constricted medially; viewed laterally, apicale strongly curved dorsally; basale 7.1-7.9 times longer than apicale.</paragraph>
<paragraph id="DFCF7A4E1529AD6AFA7419CFFA74921D" pageId="17" pageNumber="286">Female. Elytra with sutural margin and apex entire.</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="286" type="type">
<paragraph pageId="17" pageNumber="286">Type.</paragraph>
<paragraph pageId="17" pageNumber="286">Holotype, male, with labels as follows: GUAT. Depto. Zacapa, San Lorenzo, 1740 m, Sierra de las Minas, 18.VII.1986, J.M.Campbell/ beating mixed vegetation in pine-oak forest/ HOLOTYPE _, Charisius punctatus, desig. 2013, JM Campbell. The specimen is deposited in the CNCI.</paragraph>
<paragraph pageId="17" pageNumber="286">Paratypes, 44 in the CNCI, CMNC, JMCC, and the JEWC.</paragraph>
<subSubSection id="36A64183D6BA265B242EE238EF0EE097" pageId="17" pageNumber="286" type="type">
<paragraph id="7DE0E7F7FA8F87056010E2F20B53F173" pageId="17" pageNumber="286">Type.</paragraph>
<paragraph id="5DA07228297C2D0DB58A4F9BF4CEC5AF" pageId="17" pageNumber="286">Holotype, male, with labels as follows: GUAT. Depto. Zacapa, San Lorenzo, 1740 m, Sierra de las Minas, 18.VII.1986, J.M.Campbell/ beating mixed vegetation in pine-oak forest/ HOLOTYPE _, Charisius punctatus, desig. 2013, JM Campbell. The specimen is deposited in the CNCI.</paragraph>
<paragraph id="99DA3F3704832D58D0706ABE483AD7D0" pageId="17" pageNumber="286">Paratypes, 44 in the CNCI, CMNC, JMCC, and the JEWC.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="287" type="distribution and records">
<paragraph pageId="18" pageNumber="287">
<pageBreakToken pageId="18" pageNumber="287" start="start">Distribution</pageBreakToken>
<subSubSection id="F508E6566035A5B84B643292BE42A773" pageId="18" pageNumber="287" type="distribution and records">
<paragraph id="5F899210640D15B0A975F2083EB19A82" pageId="18" pageNumber="287">
<pageBreakToken id="08BCF66072BFAB4A33F1732C6720C0EB" pageId="18" pageNumber="287" start="start">Distribution</pageBreakToken>
and records.
</paragraph>
<paragraph pageId="18" pageNumber="287">This species is known only from Guatemala between 1400 and 1740 meters in elevation.</paragraph>
<paragraph pageId="18" pageNumber="287">
<paragraph id="26CC3E9E0079B0FD316CCFDE4A0947FC" pageId="18" pageNumber="287">This species is known only from Guatemala between 1400 and 1740 meters in elevation.</paragraph>
<paragraph id="8A309041BC1589AF2AF91DDC6B5EAA43" pageId="18" pageNumber="287">
GUATEMALA: Alta Verapaz: Jnct. Rds. Coban &amp;
<normalizedToken originalValue="Purulhá">Purulha</normalizedToken>
<normalizedToken id="A7610F36E24C2212691CBE42479A27ED" originalValue="Purulhá">Purulha</normalizedToken>
, 5500 ft, 24.VI.1966, JMC (JMCC) 1. Baja Verapaz: Pantin Rd., 9 km N
<normalizedToken originalValue="Salamá">Salama</normalizedToken>
<normalizedToken id="D320FE9ADC3D5E57E778A050CB12C6BC" originalValue="Salamá">Salama</normalizedToken>
, 20-21.VI.1993, JMC (JMCC) 1; 6-9 km E
<normalizedToken originalValue="Purulhá">Purulha</normalizedToken>
<normalizedToken id="A619DE3EAD8F2C3DFF1ACD9757459E32" originalValue="Purulhá">Purulha</normalizedToken>
, 5000 ft, 15-24.IV.1990, E Geisbert (JEWC) 1. Chimaltenango: Yepocapa, 4600 ft, 10.V.1966, JMC (JMCC) 1. Zacapa: San Lorenzo, Sierra de las Minas, 1740 m, 8-11.1986, JMC (CNCI, JMCC) 10; same locality, 11.VII. and 17.VII.1986, L LeSage (CNCI) 2; same locality, 5200 ft, 17-19.VI.1986, JMC (CNCI, JMCC) 11; nr. San Lorenzo, 13.IV.1990, JE.Wappes (JEWC) 1; vic. San Lorenzo, 5800 ft, 10-15.VI.1991, E Geisbert (JEWC) 1; 13 km N Hwy, San Lorenzo Rd., 3000 ft, 17.VI.1993, JMC (CNCI) 1.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="287" type="etymology">
<paragraph pageId="18" pageNumber="287">Etymology.</paragraph>
<paragraph pageId="18" pageNumber="287">The species name punctatus refers to the coarse punctation of the pronotum.</paragraph>
<subSubSection id="51DFFF72B045E7C6349A73213F86E910" pageId="18" pageNumber="287" type="etymology">
<paragraph id="5539127441B12C496DDFFF97B1D85C82" pageId="18" pageNumber="287">Etymology.</paragraph>
<paragraph id="18A230E62F071B232FC2391E7786E72B" pageId="18" pageNumber="287">The species name punctatus refers to the coarse punctation of the pronotum.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="287" type="remarks">
<paragraph pageId="18" pageNumber="287">Remarks.</paragraph>
<paragraph pageId="18" pageNumber="287">
<subSubSection id="61D79BCFF80051699CAC56E833FC9A46" pageId="18" pageNumber="287" type="remarks">
<paragraph id="C0E9CB8150592ABD87B69927B3B1178A" pageId="18" pageNumber="287">Remarks.</paragraph>
<paragraph id="A300505B83C402C8E81A1A0FC2723D7E" pageId="18" pageNumber="287">
Adults of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Charisius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Charisius punctatus" order="Coleoptera" pageId="18" pageNumber="287" phylum="Arthropoda" rank="species" species="punctatus">Charisius punctatus</taxonomicName>
<taxonomicName id="C3C09349EFC4FFC04DD9252F621BEB7A" genus="Charisius" lsidName="Charisius punctatus" pageId="18" pageNumber="287" rank="species" species="punctatus">Charisius punctatus</taxonomicName>
are very similar to those of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Charisius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Charisius salvini" order="Coleoptera" pageId="18" pageNumber="287" phylum="Arthropoda" rank="species" species="salvini">Charisius salvini</taxonomicName>
<taxonomicName id="126B1331F885D3070C9A521966C7EA23" genus="Charisius" lsidName="Charisius salvini" pageId="18" pageNumber="287" rank="species" species="salvini">Charisius salvini</taxonomicName>
, but may be easily distinguished by the additional piceous markings on the elytra (Fig. 9). Males may also be distinguished by having the basal four segments of the anterior tarsi lobed ventrally, by having a triangular expansion on the inner margin of the anterior tibia; by the broader and deeper median impression of the fifth abdominal ventrite, and by the more broadly expanded apical portion of the eighth sternal lobes (Fig. 24) which are obliquely truncate.
</paragraph>
<paragraph pageId="18" pageNumber="287">The species has been collected beating mixed dead and herbaceous vegetation in a pine-oak forest.</paragraph>
<paragraph id="5FB8FC1F1DA71CDB11B1DF62BD74458E" pageId="18" pageNumber="287">The species has been collected beating mixed dead and herbaceous vegetation in a pine-oak forest.</paragraph>
</subSubSection>
</treatment>
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<treatment id="814A5D5EA29F55638D883268E35F7146" ID-GBIF-Taxon="225766417" LSID="urn:lsid:plazi:treatment:814A5D5EA29F55638D883268E35F7146" httpUri="http://treatment.plazi.org/id/814A5D5EA29F55638D883268E35F7146" lastPageNumber="111" pageId="0" pageNumber="111" scope_family="Amphientomidae" scope_subFamily="Amphientominae">
<subSubSection id="462D4A734DC0EF7822D393B625B5EE33" pageId="0" pageNumber="111" type="nomenclature">
<paragraph id="5930A38B7D000BD9D3B5C50C87E23BC0" pageId="0" pageNumber="111">
Genus
@ -142,17 +142,17 @@ Genus
</subSubSection>
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<paragraph id="69EAFE43778D598478968AAEC0F1511B" pageId="0" pageNumber="111">
<taxonomicName id="D63DC963EAF3D7F87B9DD0385FA9450C" authorityName="Hagen" authorityYear="1859" family="Amphientomidae" genus="Amphicetomum" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphicetomum" order="Psocodea" pageId="0" pageNumber="111" rank="genus">Amphicetomum</taxonomicName>
<taxonomicName id="D63DC963EAF3D7F87B9DD0385FA9450C" authorityName="Hagen" authorityYear="1859" class="Insecta" family="Amphientomidae" genus="Amphicetomum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphicetomum" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">Amphicetomum</taxonomicName>
=
<taxonomicName id="30361F55A178C3B84DBEDD062CFE60C3" authorityName="Hagen" authorityYear="1859" family="Amphientomidae" genus="Amphicetomum" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphicetomum" order="Psocodea" pageId="0" pageNumber="111" rank="genus">Amphicetomum</taxonomicName>
<taxonomicName id="30361F55A178C3B84DBEDD062CFE60C3" authorityName="Hagen" authorityYear="1859" class="Insecta" family="Amphientomidae" genus="Amphicetomum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphicetomum" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">Amphicetomum</taxonomicName>
Hagen, 1859.
</paragraph>
<paragraph id="5A69AED2DC03DEAE6272918E264D0B95" pageId="0" pageNumber="111">
<taxonomicName id="E2069BC96D2291B3B3F74FB8E89A631C" authorityName="Enderlein" authorityYear="1925" class="Insecta" family="Amphientomidae" genus="Palaeoseopis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoseopis" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">Palaeoseopis</taxonomicName>
<taxonomicName id="E2069BC96D2291B3B3F74FB8E89A631C" authorityName="Enderlein" authorityYear="1925" class="Insecta" family="Amphientomidae" genus="Palaeoseopis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Palaeoseopis" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">Palaeoseopis</taxonomicName>
=
<taxonomicName id="1084765D70886D39188D292306293189" authorityName="Enderlein" authorityYear="1925" class="Insecta" family="Amphientomidae" genus="Palaeoseopis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Palaeoseopis" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">Palaeoseopis</taxonomicName>
<taxonomicName id="1084765D70886D39188D292306293189" authorityName="Enderlein" authorityYear="1925" class="Insecta" family="Amphientomidae" genus="Palaeoseopis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Palaeoseopis" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">Palaeoseopis</taxonomicName>
Enderlein, 1925 syn. nov. (Type species: †
<taxonomicName id="0C92CD293973F748D0B16BAAF5693915" family="Amphientomidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Am. colpolepis" order="Psocodea" pageId="0" pageNumber="111" rank="species" species="colpolepis">Am. colpolepis</taxonomicName>
<taxonomicName id="0C92CD293973F748D0B16BAAF5693915" class="Insecta" family="Amphientomidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Am. colpolepis" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="colpolepis">Am. colpolepis</taxonomicName>
Enderlein, 1905
<emphasis id="4F30DD7AE49182E2D52A797D05BC75E0" italics="true" pageId="0" pageNumber="111">by original designation.</emphasis>
).
@ -162,7 +162,7 @@ Enderlein, 1905
<paragraph id="0B036CD84DE86796511FCD1D66A2B4B1" pageId="0" pageNumber="111">Type species.</paragraph>
<paragraph id="86C0D74E6651FB7B1D932E795ADCA014" pageId="0" pageNumber="111">
<taxonomicName id="A9D85FDFDE01541C148BA349F12FDB56" family="Amphientomidae" kingdom="Animalia" lsidName="Am. (Amphientomum) paradoxum" order="Psocodea" pageId="0" pageNumber="111" rank="species" species="paradoxum" subGenus="Amphientomum">Am. (Amphientomum) paradoxum</taxonomicName>
<taxonomicName id="A9D85FDFDE01541C148BA349F12FDB56" class="Insecta" family="Amphientomidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Am. (Amphientomum) paradoxum" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="paradoxum" subGenus="Amphientomum">Am. (Amphientomum) paradoxum</taxonomicName>
Pictet, 1854.
</paragraph>
</subSubSection>
@ -200,9 +200,9 @@ provided morphological characters to define previously established groupings inc
Enderlein, 1925 as a subgenus of
<taxonomicName id="93DA3B364EDF4A950F2E31C85F676296" authorityName="Pictet" authorityYear="1854" class="Insecta" family="Amphientomidae" genus="Amphientomum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphientomum" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">Amphientomum</taxonomicName>
. As such,
<taxonomicName id="73D5F97678B34052C2390074DC477098" authority=")" family="Amphientomidae" kingdom="Animalia" lsidName="Am. (Palaeoseopsis)" order="Psocodea" pageId="0" pageNumber="111" rank="subGenus" subGenus="Palaeoseopsis">Am. (Palaeoseopsis)</taxonomicName>
<taxonomicName id="73D5F97678B34052C2390074DC477098" authority=")" class="Insecta" family="Amphientomidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Am. (Palaeoseopsis)" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="subGenus" subGenus="Palaeoseopsis">Am. (Palaeoseopsis)</taxonomicName>
is supposed to differ from
<taxonomicName id="4DADFF8D9854D7B3663848F691BF9E92" authority=")" family="Amphientomidae" kingdom="Animalia" lsidName="Am. (Amphientomum)" order="Psocodea" pageId="0" pageNumber="111" rank="subGenus" subGenus="Amphientomum">Am. (Amphientomum)</taxonomicName>
<taxonomicName id="4DADFF8D9854D7B3663848F691BF9E92" authority=")" class="Insecta" family="Amphientomidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Am. (Amphientomum)" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="subGenus" subGenus="Amphientomum">Am. (Amphientomum)</taxonomicName>
by the open basiradial cell, the lack of the basal section of Rs in the hindwing, and the emarginate scale tips (
<bibRefCitation id="ADC5890F5C4C14988CEC4337A5046FBC" author="Enderlein, G" journalOrPublisher="Konowia (Vienna)" pageId="0" pageNumber="111" pagination="97 - 108" refId="B100" refString="Enderlein, G, 1925. Beitraege zur Kenntnis der Copeognathen IX. Konowia (Vienna) 4: 97 - 108" title="Beitraege zur Kenntnis der Copeognathen IX." volume="4" year="1925">Enderlein 1925</bibRefCitation>
;
@ -210,13 +210,13 @@ by the open basiradial cell, the lack of the basal section of Rs in the hindwing
).
<bibRefCitation id="244A977499728DF945F7CCD9852D4892" author="Badonnel, A" journalOrPublisher="Publicacoes Culturais da Companhia de Diamantes de Angola" pageId="0" pageNumber="111" pagination="1 - 267" refId="B16" refString="Badonnel, A, 1955. Psocopteres de l'Angola. Publicacoes Culturais da Companhia de Diamantes de Angola 26: 1 - 267" title="Psocopteres de l'Angola." volume="26" year="1955">Badonnel (1955)</bibRefCitation>
went a step further and proposed that the
<taxonomicName id="E9EB46F75C9FAA0683A843CB195F6C38" authority="Palaeoseopsis" family="Amphientomidae" kingdom="Animalia" lsidName="(Palaeoseopsis) subsp. subgenus" order="Psocodea" pageId="0" pageNumber="111" rank="subGenus" subGenus="Palaeoseopsis">subgenus Palaeoseopsis</taxonomicName>
<taxonomicName id="E9EB46F75C9FAA0683A843CB195F6C38" authority="Palaeoseopsis" class="Insecta" family="Amphientomidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="(Palaeoseopsis) subsp. subgenus" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="subGenus" subGenus="Palaeoseopsis">subgenus Palaeoseopsis</taxonomicName>
can be removed entirely but did not follow through on this action. It should be noted that an open basiradial cell in the hindwing occurs in all species of the genus that are outside the subgenus
<taxonomicName id="06A4CFFB7820BB2CB0984C344626F045" authority=")" family="Amphientomidae" kingdom="Animalia" lsidName="Am. (Amphientomum)" order="Psocodea" pageId="0" pageNumber="111" rank="subGenus" subGenus="Amphientomum">Am. (Amphientomum)</taxonomicName>
<taxonomicName id="06A4CFFB7820BB2CB0984C344626F045" authority=")" class="Insecta" family="Amphientomidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Am. (Amphientomum)" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="subGenus" subGenus="Amphientomum">Am. (Amphientomum)</taxonomicName>
, as in most species only a short spur vein of the basal section of Rs is present or the basal section of Rs is entirely missing. Phylogenetic studies on this genus are lacking and the monophyly of the subgenera
<taxonomicName id="33F9F9B9B6998C85760B2E1FBDDC9D63" authority=")" family="Amphientomidae" kingdom="Animalia" lsidName="Am. (Palaeoseopsis)" order="Psocodea" pageId="0" pageNumber="111" rank="subGenus" subGenus="Palaeoseopsis">Am. (Palaeoseopsis)</taxonomicName>
<taxonomicName id="33F9F9B9B6998C85760B2E1FBDDC9D63" authority=")" class="Insecta" family="Amphientomidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Am. (Palaeoseopsis)" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="subGenus" subGenus="Palaeoseopsis">Am. (Palaeoseopsis)</taxonomicName>
and
<taxonomicName id="15932B10E61B6995FF17EDF9728C0D92" authority=")" family="Amphientomidae" kingdom="Animalia" lsidName="Am. (Amphientomum)" order="Psocodea" pageId="0" pageNumber="111" rank="subGenus" subGenus="Amphientomum">Am. (Amphientomum)</taxonomicName>
<taxonomicName id="15932B10E61B6995FF17EDF9728C0D92" authority=")" class="Insecta" family="Amphientomidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Am. (Amphientomum)" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="subGenus" subGenus="Amphientomum">Am. (Amphientomum)</taxonomicName>
is therefore questionable, as they are neither supported by morphological apomorphies, nor by molecular data. We therefore formalize the synonymy of
<taxonomicName id="CC0D1EB7141D2084AF258FAB8517E7A2" authorityName="Pictet" authorityYear="1854" class="Insecta" family="Amphientomidae" genus="Amphientomum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphientomum" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="1800C064FCE6FDCEC64592E67A9B8BB1" italics="true" pageId="0" pageNumber="111">Amphientomum</emphasis>
@ -232,7 +232,7 @@ j. syn., syn. nov. The diagnostic characters of the genus
are as follows after the identification key by
<bibRefCitation id="60A21CFDF9370B5BED600421AEB172F8" DOI="https://doi.org/10.18195/issn.0313-122x.83.2013.245-252" author="Taylor, CK" journalOrPublisher="Records of the Western Australian Museum" pageId="0" pageNumber="111" pagination="245 - 252" refId="B307" refString="Taylor, CK, 2013. The genus Lithoseopsis (Psocodea: Amphientomidae) in the Western Australian fauna, with description of the male of Lithoseopsis humphreysi from Barrow Island. Records of the Western Australian Museum 83: 245 - 252, DOI: https://doi.org/10.18195/issn.0313-122x.83.2013.245-252" title="The genus Lithoseopsis (Psocodea: Amphientomidae) in the Western Australian fauna, with description of the male of Lithoseopsis humphreysi from Barrow Island." url="https://doi.org/10.18195/issn.0313-122x.83.2013.245-252" volume="83" year="2013">Taylor (2013)</bibRefCitation>
: presence of wings, the vein M in the hindwing simple, presence of three ocelli, the lateral ocelli closer to each other than to compound eyes, the vein R1 reaching the wing margin in the hindwing, and the distal section of the vein Sc in the forewing present. See also the Remarks section for †
<taxonomicName id="191CCF73D7DB1372D1C3F7901A329332" family="Amphientomidae" kingdom="Animalia" lsidName="Am. knorrei" order="Psocodea" pageId="0" pageNumber="111" rank="species" species="knorrei">
<taxonomicName id="191CCF73D7DB1372D1C3F7901A329332" class="Insecta" family="Amphientomidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Am. knorrei" order="Psocodea" pageId="0" pageNumber="111" phylum="Arthropoda" rank="species" species="knorrei">
<emphasis id="3C1494ABE416B6B3FD8BF7D443EC5EC3" italics="true" pageId="0" pageNumber="111">Am. knorrei</emphasis>
</taxonomicName>
sp. nov.

View file

@ -1,259 +1,259 @@
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. STM photographs of live specimens of several Terebellides species (non-type specimens) A, B Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 (species 1; A ZMBN 116171 B ZMBN 116181) C Terebellides lavesquei sp. nov. (species 5; GNM 15112) D, E Terebellides williamsae Jirkov, 1989 (species 2; D GNM 15108 E GNM 15109) F Terebellides gracilis Malm, 1874 (species 3; GNM 15111). Abbreviations: bdl - branchial dorsal lobe; bf - branchial filament; bvl - branchial ventral lobe; ooc - oocytes; wTC - white thoracic chaetiger." figureDoi="10.3897/zookeys.1132.91244.figure2" httpUri="https://binary.pensoft.net/fig/775018" pageId="0" pageNumber="85">Figs 2D</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. STM photographs of several Terebellides species (A, C-F non-type specimens) A Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 (species 1; ZMBN 116186) B Terebellides lavesquei sp. nov. (species 5; holotype, ZMBN 116322) C Terebellides atlantis Williams, 1984 (species 16; ZMBN 116472) D Terebellides irinae Gagaev, 2009 (species 24; ZMBN 116498) E Terebellides williamsae Jirkov, 1989 (species 2; ZMBN 116269) F Terebellides gracilis Malm, 1874 (species 3; ZMBN 116283). Abbreviations: bdl - branchial dorsal lobe; bf - branchial filament; bvl - branchial ventral lobe; ooc - oocytes; wTC - white thoracic chaetiger." figureDoi="10.3897/zookeys.1132.91244.figure3" httpUri="https://binary.pensoft.net/fig/775019" pageId="0" pageNumber="85">, 3E</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Geographic distribution of species of Terebellides in Northeast Atlantic Ocean A Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 B Terebellides lavesquei sp. nov. C Terebellides atlantis Williams, 1984 D Terebellides irinae Gagaev, 2009 E Terebellides williamsae Jirkov, 1989 F Terebellides gracilis Malm, 1874. Pink star denotes the type locality of each taxon." figureDoi="10.3897/zookeys.1132.91244.figure10" httpUri="https://binary.pensoft.net/fig/775026" pageId="0" pageNumber="85">, 10E</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Bathymetric distribution of Terebellides species studied in this work." figureDoi="10.3897/zookeys.1132.91244.figure11" httpUri="https://binary.pensoft.net/fig/775027" pageId="0" pageNumber="85">, 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Body MG staining patterns in ventral view of Terebellides species. Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016, Terebellides lavesquei sp. nov., Terebellides atlantis Williams, 1984, Terebellides irinae Gagaev, 2009, Terebellides williamsae Jirkov, 1989 and Terebellides gracilis Malm, 1874. Segments indicated in Arabic numbers." figureDoi="10.3897/zookeys.1132.91244.figure12" httpUri="https://binary.pensoft.net/fig/775028" pageId="0" pageNumber="85">, 12</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">, 14</figureCitation>
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<figureCitation id="CE470550336C39677C731C6BAF7172B7" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">, 14</figureCitation>
<figureCitation id="B3678BF9653627003BEAF2D74202362E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimen, ZMBN 116249), SEM micrographs A TC 5 - 7, lateral view B TC 6 (TU 1), geniculate chaetae C row of thoracic uncini D thoracic uncinus E abdominal neuropodium F abdominal uncini. Abbreviations: cap - capitium; gc - geniculate chaetae; nc - notochaetae; TC - thoracic chaetiger; TU - thoracic unciniger." figureDoi="10.3897/zookeys.1132.91244.figure15" httpUri="https://binary.pensoft.net/fig/775031" pageId="0" pageNumber="85">, 15</figureCitation>
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Jirkov, 1989: 124.
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Species 2 -
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<bibRefCitation id="9A1B6529B384989B42FA3D7511B8D4F9" DOI="https://doi.org/10.1371/journal.pone.0198356" author="Nygren, A" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="85" refId="B26" refString="Nygren, A, Parapar, J, Pons, J, Meissner, K, Bakken, T, Kongsrud, JA, Oug, E, Gaeva, D, Sikorski, A, Johansen, RA, Hutchings, PA, Lavesque, N, Capa, M, 2018. A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13(6): e0198356. https://doi.org/10.1371/journal.pone.0198356" title="A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13 (6): e 0198356." url="https://doi.org/10.1371/journal.pone.0198356" year="2018">Nygren et al. 2018</bibRefCitation>
: 18-22, figs 6, 10.
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<paragraph pageId="0" pageNumber="85">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="85">
<materialsCitation accessionNumber="GNM14639, GNM15107, GNM15108" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" specimenCount="20">
<specimenCount type="generic">20 specimens</specimenCount>
<subSubSection id="44A6DC7C29C83E897B327A68491E08FA" pageId="0" pageNumber="85" type="materials_examined">
<paragraph id="BFAD4AA089EE570C8289D03E1B7CC7D7" pageId="0" pageNumber="85">Material examined.</paragraph>
<paragraph id="AA67B6C5D7DC97F990CC201B71E6689E" pageId="0" pageNumber="85">
<materialsCitation id="1786AA60C4099F0201BBB5B3360572ED" accessionNumber="GNM14639, GNM15107, GNM15108" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" specimenCount="20">
<specimenCount id="B8FAD0731CC3F3166B4B09436A47EE15" type="generic">20 specimens</specimenCount>
(Suppl. material 1), Skagerrak (
<accessionNumber isEnumeration="true">GNM14639, GNM15107, GNM15108</accessionNumber>
<accessionNumber id="C84D99571E0D1E9331210F5DB35AF249" isEnumeration="true">GNM14639, GNM15107, GNM15108</accessionNumber>
); Barents Sea (ZMBN116246, ZMBN116247, ZMBN116248, ZMBN116249, ZMBN116251, ZMBN116252, ZMBN116253, ZMBN116254, ZMBN116255, ZMBN116257, ZMBN116260, ZMBN116262, ZMBN116263, ZMBN116266, ZMBN116269, ZMBN116270, ZMBN116271)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="materials_examined">
<paragraph pageId="0" pageNumber="85">GenBank accession numbers of material examined (COI).</paragraph>
<paragraph pageId="0" pageNumber="85">
<materialsCitation accessionNumber="MG024957, MG024958, MG024959, MG024960, MG024961, MG024962, MG024963, MG024964, MG024965, MG024966, MG024967, MG024968, MG024969, MG024970, MG024971, MG024972, MG024973, MG024974, MG024975, MG024976, MG024977, MG024978, MG024979, MG024980, MG024981, MG024982, MG024983, MG024984, MG024985, MG024986, MG024987, MG024988" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" specimenCount="1">
<accessionNumber isEnumeration="true">MG024957, MG024958, MG024959, MG024960, MG024961, MG024962, MG024963, MG024964, MG024965, MG024966, MG024967, MG024968, MG024969, MG024970, MG024971, MG024972, MG024973, MG024974, MG024975, MG024976, MG024977, MG024978, MG024979, MG024980, MG024981, MG024982, MG024983, MG024984, MG024985, MG024986, MG024987, MG024988</accessionNumber>
<subSubSection id="D06DB1C5A389A17F7A85D14C604B1F13" pageId="0" pageNumber="85" type="materials_examined">
<paragraph id="5BC7839794759D70BAA61827468EA0DE" pageId="0" pageNumber="85">GenBank accession numbers of material examined (COI).</paragraph>
<paragraph id="79A708B399BFB17EAE265B49BB86AD7D" pageId="0" pageNumber="85">
<materialsCitation id="3E7B11E86D40061F49E4F5BF4BDA0AE4" accessionNumber="MG024957, MG024958, MG024959, MG024960, MG024961, MG024962, MG024963, MG024964, MG024965, MG024966, MG024967, MG024968, MG024969, MG024970, MG024971, MG024972, MG024973, MG024974, MG024975, MG024976, MG024977, MG024978, MG024979, MG024980, MG024981, MG024982, MG024983, MG024984, MG024985, MG024986, MG024987, MG024988" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" specimenCount="1">
<accessionNumber id="1272FB4936A6229AD9EC084CF62BED3A" isEnumeration="true">MG024957, MG024958, MG024959, MG024960, MG024961, MG024962, MG024963, MG024964, MG024965, MG024966, MG024967, MG024968, MG024969, MG024970, MG024971, MG024972, MG024973, MG024974, MG024975, MG024976, MG024977, MG024978, MG024979, MG024980, MG024981, MG024982, MG024983, MG024984, MG024985, MG024986, MG024987, MG024988</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="materials_examined">
<paragraph pageId="0" pageNumber="85">Diagnostic features of studied material.</paragraph>
<paragraph pageId="0" pageNumber="85">
<subSubSection id="7FA3013CF129F78ED301841EF0DFFC33" pageId="0" pageNumber="85" type="materials_examined">
<paragraph id="C08C3C115ACD3184D160F3DB5632797E" pageId="0" pageNumber="85">Diagnostic features of studied material.</paragraph>
<paragraph id="494077FB1A5BF82D72D90A7700DDABC8" pageId="0" pageNumber="85">
Complete individuals ranging from 9.0-34.0 mm in length (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Relationship between number of abdominal chaetigers and body length (complete specimens considered except for T. irinae)." figureDoi="10.3897/zookeys.1132.91244.figure9" httpUri="https://binary.pensoft.net/fig/775025" pageId="0" pageNumber="85">9</figureCitation>
<figureCitation id="AF0B0DABDA15B7287104A931C5578D6D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Relationship between number of abdominal chaetigers and body length (complete specimens considered except for T. irinae)." figureDoi="10.3897/zookeys.1132.91244.figure9" httpUri="https://binary.pensoft.net/fig/775025" pageId="0" pageNumber="85">9</figureCitation>
). Branchial dorsal lobes lamellae provided with well-developed papillary projections and branchial ventral lobes provided with short posterior filaments, 50.0
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="C4760C450578385D34F67D136E2A4B98" originalValue="µm">µm</normalizedToken>
in length (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. STM photographs of several Terebellides species (A, C-F non-type specimens) A Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 (species 1; ZMBN 116186) B Terebellides lavesquei sp. nov. (species 5; holotype, ZMBN 116322) C Terebellides atlantis Williams, 1984 (species 16; ZMBN 116472) D Terebellides irinae Gagaev, 2009 (species 24; ZMBN 116498) E Terebellides williamsae Jirkov, 1989 (species 2; ZMBN 116269) F Terebellides gracilis Malm, 1874 (species 3; ZMBN 116283). Abbreviations: bdl - branchial dorsal lobe; bf - branchial filament; bvl - branchial ventral lobe; ooc - oocytes; wTC - white thoracic chaetiger." figureDoi="10.3897/zookeys.1132.91244.figure3" httpUri="https://binary.pensoft.net/fig/775019" pageId="0" pageNumber="85">3E</figureCitation>
<figureCitation id="F27475371EEC61E731E8DEBCFE20283D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. STM photographs of several Terebellides species (A, C-F non-type specimens) A Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 (species 1; ZMBN 116186) B Terebellides lavesquei sp. nov. (species 5; holotype, ZMBN 116322) C Terebellides atlantis Williams, 1984 (species 16; ZMBN 116472) D Terebellides irinae Gagaev, 2009 (species 24; ZMBN 116498) E Terebellides williamsae Jirkov, 1989 (species 2; ZMBN 116269) F Terebellides gracilis Malm, 1874 (species 3; ZMBN 116283). Abbreviations: bdl - branchial dorsal lobe; bf - branchial filament; bvl - branchial ventral lobe; ooc - oocytes; wTC - white thoracic chaetiger." figureDoi="10.3897/zookeys.1132.91244.figure3" httpUri="https://binary.pensoft.net/fig/775019" pageId="0" pageNumber="85">3E</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14A</figureCitation>
<figureCitation id="3A8E824CC06A703B1DEBF71BB387C50B" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14A</figureCitation>
). Between 16-18 lamellae on dorsal lobes (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14A, B</figureCitation>
<figureCitation id="D629C248C25B5AFD38477DC436D79011" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14A, B</figureCitation>
). Ciliary tufts present in inner face of lamellae (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14B, C</figureCitation>
<figureCitation id="817EBFE1014D83C008415CA14AAB2918" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14B, C</figureCitation>
). Ventral branchial lobes hidden in between dorsal ones but sometimes discernible below (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14A</figureCitation>
<figureCitation id="682A10245E622CA7AE9726EEA187B4B4" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14A</figureCitation>
). Lateral lappets present on TC 1-4; dorsal projection of thoracic notopodia on TC 2-4 (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14A</figureCitation>
<figureCitation id="BC9FF4DD7A1BE7C937DDCB25D2EDAA17" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14A</figureCitation>
). White ventral colouration present on TC 1-4 (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. STM photographs of live specimens of several Terebellides species (non-type specimens) A, B Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 (species 1; A ZMBN 116171 B ZMBN 116181) C Terebellides lavesquei sp. nov. (species 5; GNM 15112) D, E Terebellides williamsae Jirkov, 1989 (species 2; D GNM 15108 E GNM 15109) F Terebellides gracilis Malm, 1874 (species 3; GNM 15111). Abbreviations: bdl - branchial dorsal lobe; bf - branchial filament; bvl - branchial ventral lobe; ooc - oocytes; wTC - white thoracic chaetiger." figureDoi="10.3897/zookeys.1132.91244.figure2" httpUri="https://binary.pensoft.net/fig/775018" pageId="0" pageNumber="85">2D</figureCitation>
<figureCitation id="074F10C098FCAFC0D9CC5B62E67CC733" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. STM photographs of live specimens of several Terebellides species (non-type specimens) A, B Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 (species 1; A ZMBN 116171 B ZMBN 116181) C Terebellides lavesquei sp. nov. (species 5; GNM 15112) D, E Terebellides williamsae Jirkov, 1989 (species 2; D GNM 15108 E GNM 15109) F Terebellides gracilis Malm, 1874 (species 3; GNM 15111). Abbreviations: bdl - branchial dorsal lobe; bf - branchial filament; bvl - branchial ventral lobe; ooc - oocytes; wTC - white thoracic chaetiger." figureDoi="10.3897/zookeys.1132.91244.figure2" httpUri="https://binary.pensoft.net/fig/775018" pageId="0" pageNumber="85">2D</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. STM photographs of several Terebellides species (A, C-F non-type specimens) A Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 (species 1; ZMBN 116186) B Terebellides lavesquei sp. nov. (species 5; holotype, ZMBN 116322) C Terebellides atlantis Williams, 1984 (species 16; ZMBN 116472) D Terebellides irinae Gagaev, 2009 (species 24; ZMBN 116498) E Terebellides williamsae Jirkov, 1989 (species 2; ZMBN 116269) F Terebellides gracilis Malm, 1874 (species 3; ZMBN 116283). Abbreviations: bdl - branchial dorsal lobe; bf - branchial filament; bvl - branchial ventral lobe; ooc - oocytes; wTC - white thoracic chaetiger." figureDoi="10.3897/zookeys.1132.91244.figure3" httpUri="https://binary.pensoft.net/fig/775019" pageId="0" pageNumber="85">3E</figureCitation>
<figureCitation id="814621DF003AAD3FCEB058003BEC4B33" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. STM photographs of several Terebellides species (A, C-F non-type specimens) A Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 (species 1; ZMBN 116186) B Terebellides lavesquei sp. nov. (species 5; holotype, ZMBN 116322) C Terebellides atlantis Williams, 1984 (species 16; ZMBN 116472) D Terebellides irinae Gagaev, 2009 (species 24; ZMBN 116498) E Terebellides williamsae Jirkov, 1989 (species 2; ZMBN 116269) F Terebellides gracilis Malm, 1874 (species 3; ZMBN 116283). Abbreviations: bdl - branchial dorsal lobe; bf - branchial filament; bvl - branchial ventral lobe; ooc - oocytes; wTC - white thoracic chaetiger." figureDoi="10.3897/zookeys.1132.91244.figure3" httpUri="https://binary.pensoft.net/fig/775019" pageId="0" pageNumber="85">3E</figureCitation>
). Geniculate chaetae in TC 5, acutely bent, with well-marked capitium (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimen, ZMBN 116249), SEM micrographs A TC 5 - 7, lateral view B TC 6 (TU 1), geniculate chaetae C row of thoracic uncini D thoracic uncinus E abdominal neuropodium F abdominal uncini. Abbreviations: cap - capitium; gc - geniculate chaetae; nc - notochaetae; TC - thoracic chaetiger; TU - thoracic unciniger." figureDoi="10.3897/zookeys.1132.91244.figure15" httpUri="https://binary.pensoft.net/fig/775031" pageId="0" pageNumber="85">15B</figureCitation>
<figureCitation id="E6F5F73FF3E587E2EF7FEC51E5EAD593" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimen, ZMBN 116249), SEM micrographs A TC 5 - 7, lateral view B TC 6 (TU 1), geniculate chaetae C row of thoracic uncini D thoracic uncinus E abdominal neuropodium F abdominal uncini. Abbreviations: cap - capitium; gc - geniculate chaetae; nc - notochaetae; TC - thoracic chaetiger; TU - thoracic unciniger." figureDoi="10.3897/zookeys.1132.91244.figure15" httpUri="https://binary.pensoft.net/fig/775031" pageId="0" pageNumber="85">15B</figureCitation>
). Ciliated papilla dorsal to thoracic notopodia observed in TC 7 (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14D, E</figureCitation>
<figureCitation id="3158DEA4832702850245D9AB85180E46" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimens, ZMBN 116249 and ZMBN 116251), SEM micrographs A anterior end, left lateral view B branchial lamellae, detail C ciliary tufts, detail D TC and thoracic dorsal papilla E thoracic dorsal papilla, detail. Abbreviations: abl - anterior branchial lobe; bdl - branchial dorsal lobe; bvl - branchial ventral lobe; ct - ciliary tuft; dpn - dorsal projection of notopodium; TC - thoracic chaetiger; tdp - thoracic dorsal papilla; tm - tentacular membrane." figureDoi="10.3897/zookeys.1132.91244.figure14" httpUri="https://binary.pensoft.net/fig/775030" pageId="0" pageNumber="85">14D, E</figureCitation>
). From TC 7, neuropodia with one row of
<typeStatus>type</typeStatus>
<typeStatus id="0A23245D95EC04282C92942F1B93F1E1">type</typeStatus>
1 thoracic uncini per torus, with rostrum/capitium length ratio of ~ 2:1 and capitium with a first row of two or three large teeth, followed by many smaller teeth (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimen, ZMBN 116249), SEM micrographs A TC 5 - 7, lateral view B TC 6 (TU 1), geniculate chaetae C row of thoracic uncini D thoracic uncinus E abdominal neuropodium F abdominal uncini. Abbreviations: cap - capitium; gc - geniculate chaetae; nc - notochaetae; TC - thoracic chaetiger; TU - thoracic unciniger." figureDoi="10.3897/zookeys.1132.91244.figure15" httpUri="https://binary.pensoft.net/fig/775031" pageId="0" pageNumber="85">15C, D</figureCitation>
<figureCitation id="8CCD97DE7A918DD8EFE061404FCA89C8" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimen, ZMBN 116249), SEM micrographs A TC 5 - 7, lateral view B TC 6 (TU 1), geniculate chaetae C row of thoracic uncini D thoracic uncinus E abdominal neuropodium F abdominal uncini. Abbreviations: cap - capitium; gc - geniculate chaetae; nc - notochaetae; TC - thoracic chaetiger; TU - thoracic unciniger." figureDoi="10.3897/zookeys.1132.91244.figure15" httpUri="https://binary.pensoft.net/fig/775031" pageId="0" pageNumber="85">15C, D</figureCitation>
). Abdomen with 38-44 pairs of neuropodia with
<typeStatus>type</typeStatus>
<typeStatus id="8D3F21E7D751CA36ECF12BF01FE0FC0D">type</typeStatus>
1A uncini (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimen, ZMBN 116249), SEM micrographs A TC 5 - 7, lateral view B TC 6 (TU 1), geniculate chaetae C row of thoracic uncini D thoracic uncinus E abdominal neuropodium F abdominal uncini. Abbreviations: cap - capitium; gc - geniculate chaetae; nc - notochaetae; TC - thoracic chaetiger; TU - thoracic unciniger." figureDoi="10.3897/zookeys.1132.91244.figure15" httpUri="https://binary.pensoft.net/fig/775031" pageId="0" pageNumber="85">15E, F</figureCitation>
<figureCitation id="B628733227906E04B95759673A3F3203" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Terebellides williamsae Jirkov, 1989 (species 2; non-type specimen, ZMBN 116249), SEM micrographs A TC 5 - 7, lateral view B TC 6 (TU 1), geniculate chaetae C row of thoracic uncini D thoracic uncinus E abdominal neuropodium F abdominal uncini. Abbreviations: cap - capitium; gc - geniculate chaetae; nc - notochaetae; TC - thoracic chaetiger; TU - thoracic unciniger." figureDoi="10.3897/zookeys.1132.91244.figure15" httpUri="https://binary.pensoft.net/fig/775031" pageId="0" pageNumber="85">15E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="colour pattern">
<paragraph pageId="0" pageNumber="85">Colour pattern.</paragraph>
<paragraph pageId="0" pageNumber="85">
<subSubSection id="9FFFA0A6138F0A82122EBF532261DB57" pageId="0" pageNumber="85" type="colour pattern">
<paragraph id="E31FDE4F804B019BFA7E8269795B9349" pageId="0" pageNumber="85">Colour pattern.</paragraph>
<paragraph id="06A4A3E53A67A31090D1FA9B2486A492" pageId="0" pageNumber="85">
MG staining pattern characterised by compact green colourant in SG 1-5 and SG 7-13, SG 6 white and SG 14 striped, J-shaped glandular regions in SG 3-5 (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Body MG staining patterns in ventral view of Terebellides species. Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016, Terebellides lavesquei sp. nov., Terebellides atlantis Williams, 1984, Terebellides irinae Gagaev, 2009, Terebellides williamsae Jirkov, 1989 and Terebellides gracilis Malm, 1874. Segments indicated in Arabic numbers." figureDoi="10.3897/zookeys.1132.91244.figure12" httpUri="https://binary.pensoft.net/fig/775028" pageId="0" pageNumber="85">12</figureCitation>
<figureCitation id="1D40A23B4A5087F298E8778E36864296" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Body MG staining patterns in ventral view of Terebellides species. Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016, Terebellides lavesquei sp. nov., Terebellides atlantis Williams, 1984, Terebellides irinae Gagaev, 2009, Terebellides williamsae Jirkov, 1989 and Terebellides gracilis Malm, 1874. Segments indicated in Arabic numbers." figureDoi="10.3897/zookeys.1132.91244.figure12" httpUri="https://binary.pensoft.net/fig/775028" pageId="0" pageNumber="85">12</figureCitation>
). Similar to pattern 2.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="nucleotide diagnostic features">
<paragraph pageId="0" pageNumber="85">Nucleotide diagnostic features.</paragraph>
<paragraph pageId="0" pageNumber="85">
<subSubSection id="8E8BED6626E817ABE1FF9A6AB0914C83" pageId="0" pageNumber="85" type="nucleotide diagnostic features">
<paragraph id="247B70A950A2ECB0CC2FB891B32F9E9E" pageId="0" pageNumber="85">Nucleotide diagnostic features.</paragraph>
<paragraph id="9E5B5B72A6ED357AA9FB59AC6507F676" pageId="0" pageNumber="85">
All sequences of
<taxonomicName authorityName="Jirkov" authorityYear="1989" class="Polychaeta" family="Trichobranchidae" genus="Terebellides" kingdom="Animalia" lsidName="Terebellides williamsae" order="Canalipalpata" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis italics="true" pageId="0" pageNumber="85">Terebellides williamsae</emphasis>
<taxonomicName id="AC06B4902F919D15F3DECC428FF5E823" authorityName="Jirkov" authorityYear="1989" class="Polychaeta" family="Trichobranchidae" genus="Terebellides" kingdom="Animalia" lsidName="Terebellides williamsae" order="Canalipalpata" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis id="1BD80F035D2C6097A03BFFC25D74850A" italics="true" pageId="0" pageNumber="85">Terebellides williamsae</emphasis>
</taxonomicName>
share and are distinguished from other available
<taxonomicName class="Polychaeta" family="Trichobranchidae" genus="Terebellides" kingdom="Animalia" lsidName="Terebellides" order="Canalipalpata" pageId="0" pageNumber="85" phylum="Annelida" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="85">Terebellides</emphasis>
<taxonomicName id="392B3F4BC6E9AA7626FBE2F276281149" class="Polychaeta" family="Trichobranchidae" genus="Terebellides" kingdom="Animalia" lsidName="Terebellides" order="Canalipalpata" pageId="0" pageNumber="85" phylum="Annelida" rank="genus">
<emphasis id="67461F0B12F7D64BAFC634D3D08E69D0" italics="true" pageId="0" pageNumber="85">Terebellides</emphasis>
</taxonomicName>
sequences in unique combinations of nucleotides (underlined) at the given position of our alignment: 59-62: TATC, 75-96: TGGACAACCTGGGGCATTCCTG, 132-144: TCATGCTTTTTTA, 153-157: TTTCC, 216-234: TGCTCCTGATATAGCTTTC, 264-277: CCTCCCTCCAGCTT, 315-318: GGTT, 327-342: CTGAACAGTATACCCC, 381-399: AGATTTGGCTATTTTTTCT, 414-432: TATCTCCTCTATTCTTGGC, 450-454: TACA, 515-529: AAAAATCACTACCA, 543-573: TTCACTTCCTGTATTAGCAGGAGCTATTACA, 600-609: CACTTCCTTT, 630-640: CGACCCAATTT.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="type locality">
<paragraph pageId="0" pageNumber="85">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="85">
<subSubSection id="699E91C3CB6A9DDCC81576049B46A8F3" pageId="0" pageNumber="85" type="type locality">
<paragraph id="E1C52221F60D4E52E61C6F0BAC3B57E9" pageId="0" pageNumber="85">Type locality.</paragraph>
<paragraph id="9C1435E53ABE61E20A86182BCC312394" pageId="0" pageNumber="85">
Barents Sea, Norway,
<geoCoordinate degrees="74" direction="north" minutes="30" orientation="latitude" precision="925" value="74.5">74°30'N</geoCoordinate>
<geoCoordinate id="932F4D07C88CB5AC78EEBB53F6703C1E" degrees="74" direction="north" minutes="30" orientation="latitude" precision="925" value="74.5">74°30'N</geoCoordinate>
,
<geoCoordinate degrees="28" direction="east" minutes="00" orientation="longitude" precision="925" value="28.0">28°00'E</geoCoordinate>
<geoCoordinate id="80BB4A9A462A10232AE81B2E026C9406" degrees="28" direction="east" minutes="00" orientation="longitude" precision="925" value="28.0">28°00'E</geoCoordinate>
(
<bibRefCitation author="Jirkov, IA" journalOrPublisher="Moscow State University Press, Moscow" pageId="0" pageNumber="85" refId="B16" refString="Jirkov, IA, 1989. Bottom fauna of the USSR. Polychaeta. Moscow State University Press, Moscow" title="Bottom fauna of the USSR. Polychaeta." year="1989">Jirkov 1989</bibRefCitation>
<bibRefCitation id="AC2A26A84364E8540416EFFA53A3CE2C" author="Jirkov, IA" journalOrPublisher="Moscow State University Press, Moscow" pageId="0" pageNumber="85" refId="B16" refString="Jirkov, IA, 1989. Bottom fauna of the USSR. Polychaeta. Moscow State University Press, Moscow" title="Bottom fauna of the USSR. Polychaeta." year="1989">Jirkov 1989</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="distribution">
<paragraph pageId="0" pageNumber="85">Distribution and bathymetry.</paragraph>
<paragraph pageId="0" pageNumber="85">
<subSubSection id="EC4F082E7EBB7C00F5601D1BDACB842C" pageId="0" pageNumber="85" type="distribution">
<paragraph id="535B7DE646EFC12BA86BBE6F18CA30B2" pageId="0" pageNumber="85">Distribution and bathymetry.</paragraph>
<paragraph id="8CE768C6F27BABA4512B861A132E64AA" pageId="0" pageNumber="85">
Barents Sea, Greenland Sea, Norwegian coast and shelf, Skagerrak; at depths of 178-612 m but most of the specimens (97%) were collected above 200 m (Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Geographic distribution of species of Terebellides in Northeast Atlantic Ocean A Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 B Terebellides lavesquei sp. nov. C Terebellides atlantis Williams, 1984 D Terebellides irinae Gagaev, 2009 E Terebellides williamsae Jirkov, 1989 F Terebellides gracilis Malm, 1874. Pink star denotes the type locality of each taxon." figureDoi="10.3897/zookeys.1132.91244.figure10" httpUri="https://binary.pensoft.net/fig/775026" pageId="0" pageNumber="85">10E</figureCitation>
<figureCitation id="B0C46F7AFB674BBD5BE4A36E71954EC0" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Geographic distribution of species of Terebellides in Northeast Atlantic Ocean A Terebellides shetlandica Parapar, Moreira &amp; O'Reilly, 2016 B Terebellides lavesquei sp. nov. C Terebellides atlantis Williams, 1984 D Terebellides irinae Gagaev, 2009 E Terebellides williamsae Jirkov, 1989 F Terebellides gracilis Malm, 1874. Pink star denotes the type locality of each taxon." figureDoi="10.3897/zookeys.1132.91244.figure10" httpUri="https://binary.pensoft.net/fig/775026" pageId="0" pageNumber="85">10E</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Bathymetric distribution of Terebellides species studied in this work." figureDoi="10.3897/zookeys.1132.91244.figure11" httpUri="https://binary.pensoft.net/fig/775027" pageId="0" pageNumber="85">11</figureCitation>
<figureCitation id="AD74680AA6AC60D7BDC6261134839187" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Bathymetric distribution of Terebellides species studied in this work." figureDoi="10.3897/zookeys.1132.91244.figure11" httpUri="https://binary.pensoft.net/fig/775027" pageId="0" pageNumber="85">11</figureCitation>
, Suppl. material 1).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="85" type="remarks">
<paragraph pageId="0" pageNumber="85">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="85">
<taxonomicName authorityName="Jirkov" authorityYear="1989" class="Polychaeta" family="Trichobranchidae" genus="Terebellides" kingdom="Animalia" lsidName="Terebellides williamsae" order="Canalipalpata" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis italics="true" pageId="0" pageNumber="85">Terebellides williamsae</emphasis>
<subSubSection id="7E2F8922EC1EC44435092F3C57ACA6DF" pageId="0" pageNumber="85" type="remarks">
<paragraph id="AD9AB6F3968CDC01B132344BE805543A" pageId="0" pageNumber="85">Remarks.</paragraph>
<paragraph id="EF82F0271F010ED1B98E0A001522E176" pageId="0" pageNumber="85">
<taxonomicName id="797295C8E35460A323EDBA45BDD96066" authorityName="Jirkov" authorityYear="1989" class="Polychaeta" family="Trichobranchidae" genus="Terebellides" kingdom="Animalia" lsidName="Terebellides williamsae" order="Canalipalpata" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis id="CD395751795105AEAD092C466CA57A41" italics="true" pageId="0" pageNumber="85">Terebellides williamsae</emphasis>
</taxonomicName>
is a medium-sized species, reaching up to 34 mm in length; it is characterised by the lack of papillae on margins of branchial lamellae and by having branchiae of type 2 and posterior filaments in ventral branchial lobes, thoracic uncini of type 1 and abdominal uncini of type 1A (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of discriminating taxonomic characters of the species studied in this work. Cells in italics show discriminatory characters of each subgroup. (1) sensu Parapar et al. (2016 a); (2) sensu Parapar et al. (2020 b); (3) sensu Parapar et al. (2020 a); (4) dominant trend in bold; (5) Skagerrak." httpUri="http://table.plazi.org/id/D35598DC856676ACF7BB2895D92EA844" pageId="0" pageNumber="85" tableUuid="D35598DC856676ACF7BB2895D92EA844">1</tableCitation>
<tableCitation id="25EF932D214D6CD162E431554B7C423C" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of discriminating taxonomic characters of the species studied in this work. Cells in italics show discriminatory characters of each subgroup. (1) sensu Parapar et al. (2016 a); (2) sensu Parapar et al. (2020 b); (3) sensu Parapar et al. (2020 a); (4) dominant trend in bold; (5) Skagerrak." httpUri="http://table.plazi.org/id/D35598DC856676ACF7BB2895D92EA844" pageId="0" pageNumber="85" tableUuid="D35598DC856676ACF7BB2895D92EA844">1</tableCitation>
). All these features are shared with
<taxonomicName class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
<taxonomicName id="82789F66E6B5ACAEB43DB2831ABA078E" class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis id="11CF5F7B2EADD9A3218D159E6AE49C2B" italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
</taxonomicName>
; in fact,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2983.1.1" author="Parapar, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="85" pagination="1 - 20" refId="B29" refString="Parapar, J, Moreira, J, Helgason, GV, 2011. Taxonomy and distribution of Terebellides (Polychaeta, Trichobranchidae) in Icelandic waters, with the description of a new species. Zootaxa 2983 (1): 1 - 20, DOI: https://doi.org/10.11646/zootaxa.2983.1.1" title="Taxonomy and distribution of Terebellides (Polychaeta, Trichobranchidae) in Icelandic waters, with the description of a new species." url="https://doi.org/10.11646/zootaxa.2983.1.1" volume="2983" year="2011">Parapar et al. (2011)</bibRefCitation>
<bibRefCitation id="CAD7C15A4704EDE77203CEF2873D5F21" DOI="https://doi.org/10.11646/zootaxa.2983.1.1" author="Parapar, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="85" pagination="1 - 20" refId="B29" refString="Parapar, J, Moreira, J, Helgason, GV, 2011. Taxonomy and distribution of Terebellides (Polychaeta, Trichobranchidae) in Icelandic waters, with the description of a new species. Zootaxa 2983 (1): 1 - 20, DOI: https://doi.org/10.11646/zootaxa.2983.1.1" title="Taxonomy and distribution of Terebellides (Polychaeta, Trichobranchidae) in Icelandic waters, with the description of a new species." url="https://doi.org/10.11646/zootaxa.2983.1.1" volume="2983" year="2011">Parapar et al. (2011)</bibRefCitation>
suggested this species as a synonym to
<taxonomicName class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
<taxonomicName id="3FC830C317C2F03AE7F46278B5D2D105" class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis id="A8485784932051C95F51B59D2C512BDF" italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
</taxonomicName>
after examining specimens from Iceland.
<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0198356" author="Nygren, A" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="85" refId="B26" refString="Nygren, A, Parapar, J, Pons, J, Meissner, K, Bakken, T, Kongsrud, JA, Oug, E, Gaeva, D, Sikorski, A, Johansen, RA, Hutchings, PA, Lavesque, N, Capa, M, 2018. A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13(6): e0198356. https://doi.org/10.1371/journal.pone.0198356" title="A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13 (6): e 0198356." url="https://doi.org/10.1371/journal.pone.0198356" year="2018">Nygren et al. (2018)</bibRefCitation>
<bibRefCitation id="AFB696930761751F9A9C88F0D28A3A32" DOI="https://doi.org/10.1371/journal.pone.0198356" author="Nygren, A" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="85" refId="B26" refString="Nygren, A, Parapar, J, Pons, J, Meissner, K, Bakken, T, Kongsrud, JA, Oug, E, Gaeva, D, Sikorski, A, Johansen, RA, Hutchings, PA, Lavesque, N, Capa, M, 2018. A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13(6): e0198356. https://doi.org/10.1371/journal.pone.0198356" title="A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13 (6): e 0198356." url="https://doi.org/10.1371/journal.pone.0198356" year="2018">Nygren et al. (2018)</bibRefCitation>
pointed out that there were no morphological differences between both species, but their molecular analyses indicate that specimens from the Barents Sea (&quot;Species 2&quot;) would correspond to
<taxonomicName class="Polychaeta" kingdom="Animalia" lsidName="T. williamsae" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis italics="true" pageId="0" pageNumber="85">T. williamsae</emphasis>
<taxonomicName id="4D3C7CE15D26FD896187ED2A0D6B27FE" class="Polychaeta" kingdom="Animalia" lsidName="T. williamsae" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis id="A54A854F552EB3FD1B71C1A598D1B8F7" italics="true" pageId="0" pageNumber="85">T. williamsae</emphasis>
</taxonomicName>
.
<bibRefCitation DOI="https://doi.org/10.1371/journal.pone.0198356" author="Nygren, A" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="85" refId="B26" refString="Nygren, A, Parapar, J, Pons, J, Meissner, K, Bakken, T, Kongsrud, JA, Oug, E, Gaeva, D, Sikorski, A, Johansen, RA, Hutchings, PA, Lavesque, N, Capa, M, 2018. A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13(6): e0198356. https://doi.org/10.1371/journal.pone.0198356" title="A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13 (6): e 0198356." url="https://doi.org/10.1371/journal.pone.0198356" year="2018">Nygren et al. (2018)</bibRefCitation>
<bibRefCitation id="BA250170170DDA5CC5E83D86B8D0BFFB" DOI="https://doi.org/10.1371/journal.pone.0198356" author="Nygren, A" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="85" refId="B26" refString="Nygren, A, Parapar, J, Pons, J, Meissner, K, Bakken, T, Kongsrud, JA, Oug, E, Gaeva, D, Sikorski, A, Johansen, RA, Hutchings, PA, Lavesque, N, Capa, M, 2018. A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13(6): e0198356. https://doi.org/10.1371/journal.pone.0198356" title="A mega-cryptic species complex hidden among one of the most common annelids in the North-East Atlantic. PLoS ONE 13 (6): e 0198356." url="https://doi.org/10.1371/journal.pone.0198356" year="2018">Nygren et al. (2018)</bibRefCitation>
suggested therefore that
<taxonomicName class="Polychaeta" kingdom="Animalia" lsidName="T. williamsae" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis italics="true" pageId="0" pageNumber="85">T. williamsae</emphasis>
<taxonomicName id="B630876EF687EEDCB11ABE65F9821BBD" class="Polychaeta" kingdom="Animalia" lsidName="T. williamsae" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis id="C4A7B30AA21B9489EBF610A7C5F8AAEC" italics="true" pageId="0" pageNumber="85">T. williamsae</emphasis>
</taxonomicName>
might be a valid species and different from
<taxonomicName class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
<taxonomicName id="B0052FEAD85F3080D9F301E114D0F38D" class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis id="D6E273FC68E847C7A476BFC94782059D" italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
</taxonomicName>
(&quot;Species 3&quot;, see below). Here, examination of specimens of
<taxonomicName class="Polychaeta" kingdom="Animalia" lsidName="T. williamsae" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis italics="true" pageId="0" pageNumber="85">T. williamsae</emphasis>
<taxonomicName id="8A579F4A04CA8A261E0CC94CE0A81165" class="Polychaeta" kingdom="Animalia" lsidName="T. williamsae" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis id="A5EB7B9109084714ECD0E27F73560CFD" italics="true" pageId="0" pageNumber="85">T. williamsae</emphasis>
</taxonomicName>
show that they differ from
<taxonomicName class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
<taxonomicName id="7F357133D403CF262C8D30EE0EB61665" class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis id="280326F2030796A26426D796420D7B63" italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
</taxonomicName>
in the number of chaetigers with white ventral colouration, i.e., in
<taxonomicName class="Polychaeta" kingdom="Animalia" lsidName="T. williamsae" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis italics="true" pageId="0" pageNumber="85">T. williamsae</emphasis>
<taxonomicName id="BD4C2B4980AA47CCEF56610219899052" class="Polychaeta" kingdom="Animalia" lsidName="T. williamsae" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="williamsae">
<emphasis id="8087256DF5F6C92BB36B20AA6599EACF" italics="true" pageId="0" pageNumber="85">T. williamsae</emphasis>
</taxonomicName>
white colouration is present in TC 1-4 while in
<taxonomicName class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
<taxonomicName id="570067423D6AA6BB11BA06EBCADEA314" class="Polychaeta" kingdom="Animalia" lsidName="T. gracilis" pageId="0" pageNumber="85" phylum="Annelida" rank="species" species="gracilis">
<emphasis id="CDCCFA4391CF730C1EDEEF17B84F182F" italics="true" pageId="0" pageNumber="85">T. gracilis</emphasis>
</taxonomicName>
it is only present on TC 4.
</paragraph>

View file

@ -1,157 +1,158 @@
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4818170" ID-GBIF-Taxon="182400795" ID-Zenodo-Dep="4818170" LSID="urn:lsid:plazi:treatment:817987A05774B00A799AB7120AFFFDE2" httpUri="http://treatment.plazi.org/id/817987A05774B00A799AB7120AFFFDE2" lastPageId="6" lastPageNumber="37" pageId="3" pageNumber="34">
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<paragraph blockId="3.[741,1145,533,591]" pageId="3" pageNumber="34">
<taxonomicName authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" box="[741,1145,533,559]" class="Actinopterygii" family="Cyprinidae" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="macrurus">
<emphasis box="[741,973,533,559]" italics="true" pageId="3" pageNumber="34">Tarsichthys macrurus</emphasis>
<treatment id="817987A05774B00A799AB7120AFFFDE2" ID-DOI="http://doi.org/10.5281/zenodo.4818170" ID-GBIF-Taxon="182400795" ID-Zenodo-Dep="4818170" LSID="urn:lsid:plazi:treatment:817987A05774B00A799AB7120AFFFDE2" httpUri="http://treatment.plazi.org/id/817987A05774B00A799AB7120AFFFDE2" lastPageId="6" lastPageNumber="37" pageId="3" pageNumber="34" scope_family="Cyprinidae">
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<paragraph id="096F36B65774B00F799AB71208C3FDA2" blockId="3.[741,1145,533,591]" pageId="3" pageNumber="34">
<taxonomicName id="CED04D355774B00F799AB7120F5EFDC2" authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" box="[741,1145,533,559]" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="3BA4EAA45774B00F799AB71208EAFDC3" box="[741,973,533,559]" italics="true" pageId="3" pageNumber="34">Tarsichthys macrurus</emphasis>
(Agassiz, 1839)
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(
<figureCitation box="[897,987,564,591]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="4.[112,123,700,718]" captionStartId-1="4.[677,688,1199,1217]" captionStartId-2="6.[677,688,1054,1072]" captionStartId-3="6.[677,688,1410,1428]" captionTargetBox-0="[113,1208,214,659]" captionTargetBox-1="[681,1202,787,1157]" captionTargetBox-2="[677,1206,215,933]" captionTargetBox-3="[678,1209,1168,1373]" captionTargetId-0="figure-352@4.[113,1208,214,659]" captionTargetId-1="figure-380@4.[678,1207,732,1176]" captionTargetId-2="figure-346@6.[677,1206,214,1012]" captionTargetId-3="figure-400@6.[677,1210,1167,1374]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="6" captionText-0="FIG. 3.— Tarsichthys macrurus (Agassiz,1839).Squelette découvert par André Nel au nord-est des Figons (MNHN.F.AIX224).Échelle: 1 cm." captionText-1="FIG. 4. — Tarsichthys cf. macrurus (Agassiz, 1839). Région operculaire du spécimen des Figons. Échelle: 2 mm." captionText-2="FIG. 5. — Tarsichthys cf. macrurus (Agassiz, 1839). Dent pharyngienne prélevée sur le squelette des Figons (MNHN.F.AIX224) Échelle: 1 mm." captionText-3="FIG. 6. — Tarsichthys cf. macrurus (Agassiz, 1839). Premier rayon de nageoire pelvienne du spécimen des Figons. Échelle: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4537935" figureDoi-1="http://doi.org/10.5281/zenodo.4537937" figureDoi-2="http://doi.org/10.5281/zenodo.4537939" figureDoi-3="http://doi.org/10.5281/zenodo.4537943" httpUri-0="https://zenodo.org/record/4537935/files/figure.png" httpUri-1="https://zenodo.org/record/4537937/files/figure.png" httpUri-2="https://zenodo.org/record/4537939/files/figure.png" httpUri-3="https://zenodo.org/record/4537943/files/figure.png" pageId="3" pageNumber="34">Figs 3-6</figureCitation>
<figureCitation id="91EB2A335774B00F78FEB73308FCFDA2" box="[897,987,564,591]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="4.[112,123,700,718]" captionStartId-1="4.[677,688,1199,1217]" captionStartId-2="6.[677,688,1054,1072]" captionStartId-3="6.[677,688,1410,1428]" captionTargetBox-0="[113,1208,214,659]" captionTargetBox-1="[681,1202,787,1157]" captionTargetBox-2="[677,1206,215,933]" captionTargetBox-3="[678,1209,1168,1373]" captionTargetId-0="figure-352@4.[113,1208,214,659]" captionTargetId-1="figure-380@4.[678,1207,732,1176]" captionTargetId-2="figure-346@6.[677,1206,214,1012]" captionTargetId-3="figure-400@6.[677,1210,1167,1374]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="6" captionText-0="FIG. 3.— Tarsichthys macrurus (Agassiz,1839).Squelette découvert par André Nel au nord-est des Figons (MNHN.F.AIX224).Échelle: 1 cm." captionText-1="FIG. 4. — Tarsichthys cf. macrurus (Agassiz, 1839). Région operculaire du spécimen des Figons. Échelle: 2 mm." captionText-2="FIG. 5. — Tarsichthys cf. macrurus (Agassiz, 1839). Dent pharyngienne prélevée sur le squelette des Figons (MNHN.F.AIX224) Échelle: 1 mm." captionText-3="FIG. 6. — Tarsichthys cf. macrurus (Agassiz, 1839). Premier rayon de nageoire pelvienne du spécimen des Figons. Échelle: 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4537935" figureDoi-1="http://doi.org/10.5281/zenodo.4537937" figureDoi-2="http://doi.org/10.5281/zenodo.4537939" figureDoi-3="http://doi.org/10.5281/zenodo.4537943" httpUri-0="https://zenodo.org/record/4537935/files/figure.png" httpUri-1="https://zenodo.org/record/4537937/files/figure.png" httpUri-2="https://zenodo.org/record/4537939/files/figure.png" httpUri-3="https://zenodo.org/record/4537943/files/figure.png" pageId="3" pageNumber="34">Figs 3-6</figureCitation>
)
</paragraph>
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<paragraph blockId="3.[675,1210,628,844]" pageId="3" pageNumber="34">
<subSubSection id="41CA653D5774B00A79DAB773091BFEFC" lastPageId="6" lastPageNumber="37" pageId="3" pageNumber="34" type="description">
<paragraph id="096F36B65774B00F79DAB7730851FCA1" blockId="3.[675,1210,628,844]" pageId="3" pageNumber="34">
Un spécimen pratiquement complet à lexception de la partie postérieure de la nageoire caudale de
<taxonomicName box="[676,796,691,717]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
<taxonomicName id="CED04D355774B00F79DBB7B4083BFD20" box="[676,796,691,717]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
a été découvert par André Nel au nordest du village des Figons (
<figureCitation box="[952,1016,723,749]" captionStart="FIG" captionStartId="4.[112,123,700,718]" captionTargetBox="[113,1208,214,659]" captionTargetId="figure-352@4.[113,1208,214,659]" captionTargetPageId="4" captionText="FIG. 3.— Tarsichthys macrurus (Agassiz,1839).Squelette découvert par André Nel au nord-est des Figons (MNHN.F.AIX224).Échelle: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4537935" httpUri="https://zenodo.org/record/4537935/files/figure.png" pageId="3" pageNumber="34">Fig. 3</figureCitation>
<figureCitation id="91EB2A335774B00F78C7B7D408DFFD00" box="[952,1016,723,749]" captionStart="FIG" captionStartId="4.[112,123,700,718]" captionTargetBox="[113,1208,214,659]" captionTargetId="figure-352@4.[113,1208,214,659]" captionTargetPageId="4" captionText="FIG. 3.— Tarsichthys macrurus (Agassiz,1839).Squelette découvert par André Nel au nord-est des Figons (MNHN.F.AIX224).Échelle: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4537935" httpUri="https://zenodo.org/record/4537935/files/figure.png" pageId="3" pageNumber="34">Fig. 3</figureCitation>
). Il est conservé à Paris, dans les collections paléontologiques du Muséum national dHistoire naturelle, sous le numéro MNHN.F.AIX224.
</paragraph>
<paragraph blockId="3.[675,1212,882,1700]" box="[677,971,882,908]" pageId="3" pageNumber="34">DESCRIPTION ANATOMIQUE</paragraph>
<paragraph blockId="3.[675,1212,882,1700]" pageId="3" pageNumber="34">
<paragraph id="096F36B65774B00F79DAB67508ECFC67" blockId="3.[675,1212,882,1700]" box="[677,971,882,908]" pageId="3" pageNumber="34">DESCRIPTION ANATOMIQUE</paragraph>
<paragraph id="096F36B65774B00F79DAB69608ADFBE7" blockId="3.[675,1212,882,1700]" pageId="3" pageNumber="34">
Cest un poisson au corps allongé dont la longueur standard mesure
<quantity box="[855,948,945,971]" metricMagnitude="-1" metricUnit="m" metricValue="1.0699999999999998" pageId="3" pageNumber="34" unit="mm" value="107.0">107 mm</quantity>
<quantity id="CE289B535774B00F7828B6B60893FC26" box="[855,948,945,971]" metricMagnitude="-1" metricUnit="m" metricValue="1.0699999999999998" pageId="3" pageNumber="34" unit="mm" value="107.0">107 mm</quantity>
et dont la hauteur maximale du corps est comprise environ 3,5 fois dans la longueur standard.
</paragraph>
<paragraph blockId="3.[675,1212,882,1700]" pageId="3" pageNumber="34">Ses principales dimensions (en millimètres) se présentent comme suit:</paragraph>
<paragraph blockId="3.[675,1212,882,1700]" pageId="3" pageNumber="34"> Longueur standard ……………………. 107 Hauteur maximale du corps …………... 28,5 Longueur de la tête ……………………. 30 Distance anté-dorsale ………………….. 50 Distance anté-anale ……………………. 74,5 Distance anté-pectorale ……………… 31 Distance anté-pelvienne ……………… 52 Longueur de la dorsale ………………. 23,5 Longueur de lanale ………………….. 17,5 Longueur des pectorales ……………… 19 Longueur des pelviennes ………………. 23 Longueur basale de la dorsale …………. 15 Longueur basale de lanale …………….. 9,5 Longueur du pédicule caudal ………….. 18,5 Hauteur du pédicule caudal ……………. 15,5 La tête est conservée en double empreinte.Relativement massive, sa hauteur égale approximativement les ¾ de sa longueur. Lorbite est relativement petite, son diamètre horizontal égalant environ un quart de</paragraph>
<paragraph blockId="4.[827,1187,77,95]" box="[827,1187,77,95]" pageId="4" pageNumber="35">
<taxonomicName box="[827,907,78,95]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="35" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
<paragraph id="096F36B65774B00F79C4B117088DFBA7" blockId="3.[675,1212,882,1700]" pageId="3" pageNumber="34">Ses principales dimensions (en millimètres) se présentent comme suit:</paragraph>
<paragraph id="096F36B65774B00F79DAB1480F9DF94E" blockId="3.[675,1212,882,1700]" pageId="3" pageNumber="34"> Longueur standard ……………………. 107 Hauteur maximale du corps …………... 28,5 Longueur de la tête ……………………. 30 Distance anté-dorsale ………………….. 50 Distance anté-anale ……………………. 74,5 Distance anté-pectorale ……………… 31 Distance anté-pelvienne ……………… 52 Longueur de la dorsale ………………. 23,5 Longueur de lanale ………………….. 17,5 Longueur des pectorales ……………… 19 Longueur des pelviennes ………………. 23 Longueur basale de la dorsale …………. 15 Longueur basale de lanale …………….. 9,5 Longueur du pédicule caudal ………….. 18,5 Hauteur du pédicule caudal ……………. 15,5 La tête est conservée en double empreinte.Relativement massive, sa hauteur égale approximativement les ¾ de sa longueur. Lorbite est relativement petite, son diamètre horizontal égalant environ un quart de</paragraph>
<paragraph id="096F36B65773B0087844B5490F84FFB2" blockId="4.[827,1187,77,95]" box="[827,1187,77,95]" pageId="4" pageNumber="35">
<taxonomicName id="CED04D355773B0087844B54908ACFFB2" box="[827,907,78,95]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="35" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
oligocènes dAix-en-Provence (
<collectingCountry box="[1132,1182,77,95]" name="France" pageId="4" pageNumber="35">France</collectingCountry>
<collectingCountry id="71C776265773B0087F13B54A0FB9FFB2" box="[1132,1182,77,95]" name="France" pageId="4" pageNumber="35">France</collectingCountry>
)
</paragraph>
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<paragraph id="096F36B65773B0087B0FB7BB0F9CFD23" blockId="4.[112,1211,697,718]" box="[112,1211,697,718]" pageId="4" pageNumber="35">
FIG. 3.—
<taxonomicName authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" box="[186,481,700,718]" class="Pisces (awaiting allocation)" family="Pisces (awaiting allocation)" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pisces (awaiting allocation)" pageId="4" pageNumber="35" phylum="Chordata" rank="species" species="macrurus">
<emphasis box="[186,354,700,718]" italics="true" pageId="4" pageNumber="35">Tarsichthys macrurus</emphasis>
<taxonomicName id="CED04D355773B0087BC5B7BB0AC6FD23" authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" box="[186,481,700,718]" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="35" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="3BA4EAA45773B0087BC5B7BB0A45FD23" box="[186,354,700,718]" italics="true" pageId="4" pageNumber="35">Tarsichthys macrurus</emphasis>
(Agassiz,1839)
</taxonomicName>
.Squelette découvert par André Nel au nord-est des Figons (MNHN.F.AIX224).Échelle: 1 cm.
</paragraph>
</caption>
<paragraph blockId="4.[111,648,786,1700]" pageId="4" pageNumber="35">
<paragraph id="096F36B65773B0087B0EB6150A84FA4B" blockId="4.[111,648,786,1700]" pageId="4" pageNumber="35">
la longueur de la tête. Elle est délimitée par la série infraorbitaire caractérisée par le grand développement du lacrymal qui est étiré longitudinalement.La cavité buccale est courte, larticulation de la mandibule avec le crâne prenant place en avant de la verticale passant par le bord antérieur de lorbite. De ce fait, la longueur de la mandibule est sensiblement inférieure à la moitié de la longueur de la tête. En arrière de lorbite sobservent les restes de lhyomandibulaire, ainsi que le préopercule dont la branche verticale est à peine plus longue que la branche horizontale, ces deux branches déterminant entre elles un angle denviron130° mesuré le long des deux branches du canal préoperculaire. Lopercule est relativement large: sa largeur maximale égale environ le tiers de la longueur de la tête. Son bord antérieur subrectiligne se termine dorsalement par un angle antéro-dorsal saillant. Le bord dorsal de los, faiblement concave, se termine vers larrière par un angle postéro-dorsal saillant à partir duquel le contour postéro-dorsal sabaisse régulièrement jusquà langle antéro-ventral (
<figureCitation box="[342,406,1420,1446]" captionStart="FIG" captionStartId="4.[677,688,1199,1217]" captionTargetBox="[681,1202,787,1157]" captionTargetId="figure-380@4.[678,1207,732,1176]" captionTargetPageId="4" captionText="FIG. 4. — Tarsichthys cf. macrurus (Agassiz, 1839). Région operculaire du spécimen des Figons. Échelle: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4537937" httpUri="https://zenodo.org/record/4537937/files/figure.png" pageId="4" pageNumber="35">Fig. 4</figureCitation>
<figureCitation id="91EB2A335773B0087A29B08B0AB1FA4B" box="[342,406,1420,1446]" captionStart="FIG" captionStartId="4.[677,688,1199,1217]" captionTargetBox="[681,1202,787,1157]" captionTargetId="figure-380@4.[678,1207,732,1176]" captionTargetPageId="4" captionText="FIG. 4. — Tarsichthys cf. macrurus (Agassiz, 1839). Région operculaire du spécimen des Figons. Échelle: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4537937" httpUri="https://zenodo.org/record/4537937/files/figure.png" pageId="4" pageNumber="35">Fig. 4</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4537937" ID-Zenodo-Dep="4537937" httpUri="https://zenodo.org/record/4537937/files/figure.png" pageId="4" pageNumber="35" startId="4.[677,688,1199,1217]" targetBox="[681,1202,787,1157]" targetPageId="4">
<paragraph blockId="4.[677,1209,1198,1239]" pageId="4" pageNumber="35">
<caption id="5DAF663E5773B00879DAB1A80F1DFB3A" ID-DOI="http://doi.org/10.5281/zenodo.4537937" ID-Zenodo-Dep="4537937" httpUri="https://zenodo.org/record/4537937/files/figure.png" pageId="4" pageNumber="35" startId="4.[677,688,1199,1217]" targetBox="[681,1202,787,1157]" targetPageId="4">
<paragraph id="096F36B65773B00879DAB1A80F1DFB3A" blockId="4.[677,1209,1198,1239]" pageId="4" pageNumber="35">
FIG. 4. —
<taxonomicName authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" box="[757,1094,1199,1217]" class="Pisces (awaiting allocation)" family="Pisces (awaiting allocation)" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pisces (awaiting allocation)" pageId="4" pageNumber="35" phylum="Chordata" rank="species" species="macrurus">
<emphasis box="[757,849,1199,1217]" italics="true" pageId="4" pageNumber="35">Tarsichthys</emphasis>
<taxonomicName id="CED04D355773B008798AB1A80F61FB2C" authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" box="[757,1094,1199,1217]" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="35" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="3BA4EAA45773B008798AB1A80876FB2C" box="[757,849,1199,1217]" italics="true" pageId="4" pageNumber="35">Tarsichthys</emphasis>
cf.
<emphasis box="[881,959,1199,1217]" italics="true" pageId="4" pageNumber="35">macrurus</emphasis>
<emphasis id="3BA4EAA45773B008780EB1A80898FB2C" box="[881,959,1199,1217]" italics="true" pageId="4" pageNumber="35">macrurus</emphasis>
(Agassiz, 1839)
</taxonomicName>
. Région operculaire du spécimen des Figons. Échelle: 2 mm.
</paragraph>
</caption>
<paragraph blockId="4.[111,648,786,1700]" pageId="4" pageNumber="35">
<materialsCitation ID-GBIF-Occurrence="3119716301" collectorName="de Rutte" location="Hakenzahn de Rutte" pageId="4" pageNumber="35" specimenCount="1">
<paragraph id="096F36B65773B0087BF7B0AB0A32F949" blockId="4.[111,648,786,1700]" pageId="4" pageNumber="35">
<materialsCitation id="B9B83CEB5773B0087BF7B0AB0ACAF9C9" ID-GBIF-Occurrence="3119716301" collectorName="de Rutte" location="Hakenzahn de Rutte" pageId="4" pageNumber="35" specimenCount="1">
Une dent pharyngienne gauche du
<typeStatus box="[507,552,1452,1478]" pageId="4" pageNumber="35">type</typeStatus>
<typeStatus id="D66B88145773B0087A84B0AB090FFA2B" box="[507,552,1452,1478]" pageId="4" pageNumber="35">type</typeStatus>
«en crochet» (
<location LSID="urn:lsid:plazi:treatment:817987A05774B00A799AB7120AFFFDE2:0C0F606D5773B0087BC8B0CC0AA9FA0B" box="[183,398,1483,1510]" name="Hakenzahn de Rutte" pageId="4" pageNumber="35">
<location id="0C0F606D5773B0087BC8B0CC0AA9FA0B" LSID="urn:lsid:plazi:treatment:817987A05774B00A799AB7120AFFFDE2:0C0F606D5773B0087BC8B0CC0AA9FA0B" box="[183,398,1483,1510]" name="Hakenzahn de Rutte" pageId="4" pageNumber="35">
Hakenzahn
<collectorName box="[310,398,1483,1510]" pageId="4" pageNumber="35">de Rutte</collectorName>
<collectorName id="A42553605773B0087A49B0CC0AA9FA0B" box="[310,398,1483,1510]" pageId="4" pageNumber="35">de Rutte</collectorName>
</location>
1962) était fossilisée sur ce squelette (
<figureCitation box="[245,307,1514,1541]" captionStart="FIG" captionStartId="6.[677,688,1054,1072]" captionTargetBox="[677,1206,215,933]" captionTargetId="figure-346@6.[677,1206,214,1012]" captionTargetPageId="6" captionText="FIG. 5. — Tarsichthys cf. macrurus (Agassiz, 1839). Dent pharyngienne prélevée sur le squelette des Figons (MNHN.F.AIX224) Échelle: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4537939" httpUri="https://zenodo.org/record/4537939/files/figure.png" pageId="4" pageNumber="35">Fig. 5</figureCitation>
<figureCitation id="91EB2A335773B0087B8AB0EC0A14F9E8" box="[245,307,1514,1541]" captionStart="FIG" captionStartId="6.[677,688,1054,1072]" captionTargetBox="[677,1206,215,933]" captionTargetId="figure-346@6.[677,1206,214,1012]" captionTargetPageId="6" captionText="FIG. 5. — Tarsichthys cf. macrurus (Agassiz, 1839). Dent pharyngienne prélevée sur le squelette des Figons (MNHN.F.AIX224) Échelle: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4537939" httpUri="https://zenodo.org/record/4537939/files/figure.png" pageId="4" pageNumber="35">Fig. 5</figureCitation>
). Cette dent, de forme allongée, possède une base relativement grêle
</materialsCitation>
; sa couronne sélargit fortement au-dessous de lextrémité de larea masticatrice. Celle-ci, en forme de gouttière, détermine un angle denviron 35° avec laxe longitudinal de la couronne.
</paragraph>
<paragraph blockId="4.[675,1212,1293,1700]" lastBlockId="5.[111,649,216,1700]" lastPageId="5" lastPageNumber="36" pageId="4" pageNumber="35">La colonne vertébrale est formée de 37 vertèbres; quatre dentre elles sont modifiées pour constituer lappareil de Weber dont la présence se déduit de lobservation dune lame osseuse disposée dans le plan sagittal en avant de larc neural court porté par le composant postérieur de lappareil de Weber, et de lexistence dune paire de longues parapophyses en avant de la première paire de côtes pleurales. On dénombre 17 vertèbres postabdominales. Les centra vertébraux sont tous sensiblement allongés. Les sept centra abdominaux libres antérieurs supportent des neurapophyses très longues dont lextrémité distale atteint presque le bord dorsal du corps. Cinq dorsospinalia de forme plus ou moins triangulaire sintercalent entre les neurapophyses situées en avant de la nageoire dorsale.</paragraph>
<paragraph blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">Dans la région postabdominale, les neurapophyses, de forme arquée, sont un peu moins longues puisque leur extrémité distale prend place approximativement aux ⅔ de la distance séparant la région dorsale des centra du bord dorsal du corps. Ventralement, les hémapophyses font pendant aux neurapophyses. Leur extrémité distale atteint environ les ¾ de la distance séparant la région ventrale des centra du bord dorsal du corps.</paragraph>
<paragraph blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">
<paragraph id="096F36B65773B00979C4B00A0936FEDC" blockId="4.[675,1212,1293,1700]" lastBlockId="5.[111,649,216,1700]" lastPageId="5" lastPageNumber="36" pageId="4" pageNumber="35">La colonne vertébrale est formée de 37 vertèbres; quatre dentre elles sont modifiées pour constituer lappareil de Weber dont la présence se déduit de lobservation dune lame osseuse disposée dans le plan sagittal en avant de larc neural court porté par le composant postérieur de lappareil de Weber, et de lexistence dune paire de longues parapophyses en avant de la première paire de côtes pleurales. On dénombre 17 vertèbres postabdominales. Les centra vertébraux sont tous sensiblement allongés. Les sept centra abdominaux libres antérieurs supportent des neurapophyses très longues dont lextrémité distale atteint presque le bord dorsal du corps. Cinq dorsospinalia de forme plus ou moins triangulaire sintercalent entre les neurapophyses situées en avant de la nageoire dorsale.</paragraph>
<paragraph id="096F36B65772B0097BF7B4300A70FDA2" blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">Dans la région postabdominale, les neurapophyses, de forme arquée, sont un peu moins longues puisque leur extrémité distale prend place approximativement aux ⅔ de la distance séparant la région dorsale des centra du bord dorsal du corps. Ventralement, les hémapophyses font pendant aux neurapophyses. Leur extrémité distale atteint environ les ¾ de la distance séparant la région ventrale des centra du bord dorsal du corps.</paragraph>
<paragraph id="096F36B65772B0097BF7B75209A0FD20" blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">
Des
<emphasis box="[181,291,596,622]" italics="true" pageId="5" pageNumber="36">epineuralia</emphasis>
<emphasis id="3BA4EAA45772B0097BCAB7530A04FD83" box="[181,291,596,622]" italics="true" pageId="5" pageNumber="36">epineuralia</emphasis>
très développés, bifides vers lavant, sont présents tout le long de la colonne vertébrale contre la base des neurapophyses. En outre, des
<emphasis box="[115,234,691,717]" italics="true" pageId="5" pageNumber="36">epipleuralia</emphasis>
<emphasis id="3BA4EAA45772B0097B0CB7B40BCDFD20" box="[115,234,691,717]" italics="true" pageId="5" pageNumber="36">epipleuralia</emphasis>
existent dans la région postabdominale.
</paragraph>
<paragraph blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">Les côtes pleurales, au nombre denviron 13 paires, sont longues et robustes. Lextrémité distale des côtes situées en avant des nageoires pelviennes atteint pratiquement le bord ventral de la cavité abdominale.</paragraph>
<paragraph blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">La nageoire caudale nest quincomplètement conservée car lextrémité distale de ses rayons manque. Il apparaît toutefois quelle était fourchue. Elle est composée de 19 rayons principaux dont 17 sont à la fois articulés et bifurqués, auxquels sajoutent, dorsalement et ventralement, dix rayons marginaux.</paragraph>
<paragraph blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">Le squelette caudal axial est composé de quatre éléments. À larrière prend place le complexe uroterminal, prolongé par un uroneural qui est redressé à 140° par rapport à laxe de la colonne vertébrale. Il supporte ventralement cinq hypuraux en avant desquels prend place le parhypural. Dorsalement, le complexe uro-terminal est surmonté par un arc neural court en arrière duquel est disposé un épural unique très long. Les trois centra préuraux libres portent des neurapophyses et des hémapophyses très longues qui supportent les rayons marginaux dorsaux et ventraux.</paragraph>
<paragraph blockId="5.[111,649,216,1700]" lastBlockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">La nageoire dorsale est insérée un peu en avant du milieu de la longueur du corps, mesurée de la pointe du museau au bord postérieur des hypuraux. Elle débute par deux courts rayons. Le troisième rayon, qui est seulement articulé, est le plus long de la nageoire. Sa longueur égale approximativement les ⁴⁄₅ de la hauteur maximale du corps. Plus en arrière prennent place huit rayons à la fois articulés et bifurqués dont la longueur diminue progressivement vers larrière.</paragraph>
<paragraph blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">Lendosquelette de la dorsale est composé de neuf ptérygiophores.Le ptérygiophore antérieur, bifide, est très long. Son axe principal sengage très profondé - ment entre les septième et huitième neurapophyses portées par les centra abdominaux libres.Lextrémité proximale des deuxième et troisième ptérygiophores sintercale assez profondément entre les neurapophyses suivantes. Les axonostes suivants sont beaucoup plus courts, au point que leur extrémité proximale atteint à peine, ou parfois natteint pas, lextrémité distale des neurapophyses.</paragraph>
<paragraph blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">La nageoire anale occupe une position assez reculée: elle débute nettement en arrière de la verticale passant par la base du dernier rayon de la dorsale. On y distingue à lavant deux courts rayons. Ensuite, prend place un long rayon articulé dont la longueur égale les ⁴⁄₅ de la hauteur du corps mesurée au niveau de son point dinsertion. Plus en arrière prennent place sept rayons à la fois articulés et bifurqués dont la longueur diminue progressivement vers larrière.</paragraph>
<paragraph blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">Lendosquelette de lanale est composé de huit ptérygiophores. Les six premiers sont suffisamment longs pour que leur extrémité proximale sintercale plus ou moins profondément entre les extrémités distales des premières hémapophyses.</paragraph>
<paragraph blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">Les nageoires pectorales, insérées en position basse sur le flanc, sont grandes: lextrémité distale de leur plus long rayon atteint les ¾ de la distance séparant sa base de lorigine des pelviennes. On y dénombre une quinzaine de rayons.</paragraph>
<paragraph blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">
<paragraph id="096F36B65772B0097BF7B7D40A59FC86" blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">Les côtes pleurales, au nombre denviron 13 paires, sont longues et robustes. Lextrémité distale des côtes situées en avant des nageoires pelviennes atteint pratiquement le bord ventral de la cavité abdominale.</paragraph>
<paragraph id="096F36B65772B0097BF7B67509A1FBC4" blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">La nageoire caudale nest quincomplètement conservée car lextrémité distale de ses rayons manque. Il apparaît toutefois quelle était fourchue. Elle est composée de 19 rayons principaux dont 17 sont à la fois articulés et bifurqués, auxquels sajoutent, dorsalement et ventralement, dix rayons marginaux.</paragraph>
<paragraph id="096F36B65772B0097BF7B1370A6BFA4B" blockId="5.[111,649,216,1700]" pageId="5" pageNumber="36">Le squelette caudal axial est composé de quatre éléments. À larrière prend place le complexe uroterminal, prolongé par un uroneural qui est redressé à 140° par rapport à laxe de la colonne vertébrale. Il supporte ventralement cinq hypuraux en avant desquels prend place le parhypural. Dorsalement, le complexe uro-terminal est surmonté par un arc neural court en arrière duquel est disposé un épural unique très long. Les trois centra préuraux libres portent des neurapophyses et des hémapophyses très longues qui supportent les rayons marginaux dorsaux et ventraux.</paragraph>
<paragraph id="096F36B65772B0097BF7B0AB0849FEFC" blockId="5.[111,649,216,1700]" lastBlockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">La nageoire dorsale est insérée un peu en avant du milieu de la longueur du corps, mesurée de la pointe du museau au bord postérieur des hypuraux. Elle débute par deux courts rayons. Le troisième rayon, qui est seulement articulé, est le plus long de la nageoire. Sa longueur égale approximativement les ⁴⁄₅ de la hauteur maximale du corps. Plus en arrière prennent place huit rayons à la fois articulés et bifurqués dont la longueur diminue progressivement vers larrière.</paragraph>
<paragraph id="096F36B65772B00979C4B410089BFD83" blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">Lendosquelette de la dorsale est composé de neuf ptérygiophores.Le ptérygiophore antérieur, bifide, est très long. Son axe principal sengage très profondé - ment entre les septième et huitième neurapophyses portées par les centra abdominaux libres.Lextrémité proximale des deuxième et troisième ptérygiophores sintercale assez profondément entre les neurapophyses suivantes. Les axonostes suivants sont beaucoup plus courts, au point que leur extrémité proximale atteint à peine, ou parfois natteint pas, lextrémité distale des neurapophyses.</paragraph>
<paragraph id="096F36B65772B00979C4B77308C2FC46" blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">La nageoire anale occupe une position assez reculée: elle débute nettement en arrière de la verticale passant par la base du dernier rayon de la dorsale. On y distingue à lavant deux courts rayons. Ensuite, prend place un long rayon articulé dont la longueur égale les ⁴⁄₅ de la hauteur du corps mesurée au niveau de son point dinsertion. Plus en arrière prennent place sept rayons à la fois articulés et bifurqués dont la longueur diminue progressivement vers larrière.</paragraph>
<paragraph id="096F36B65772B00979C4B6B70F11FBA4" blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">Lendosquelette de lanale est composé de huit ptérygiophores. Les six premiers sont suffisamment longs pour que leur extrémité proximale sintercale plus ou moins profondément entre les extrémités distales des premières hémapophyses.</paragraph>
<paragraph id="096F36B65772B00979C4B1480F00FB05" blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">Les nageoires pectorales, insérées en position basse sur le flanc, sont grandes: lextrémité distale de leur plus long rayon atteint les ¾ de la distance séparant sa base de lorigine des pelviennes. On y dénombre une quinzaine de rayons.</paragraph>
<paragraph id="096F36B65772B00979C4B1E90F64F94E" blockId="5.[674,1212,216,1700]" pageId="5" pageNumber="36">
Les nageoires pelviennes, qui sont insérées approximativement au milieu de lespace séparant la base des pectorales de lorigine de lanale, sont grandes, au point que lextrémité distale de leurs plus longs rayons atteint pratiquement lorigine de lanale. On y distingue un rayon très épaissi et sept rayons articulés et bifurqués. La présence de ce rayon épaissi dont les articles sont beaucoup plus larges que hauts indique que nous sommes en présence dun individu mâle dune espèce caractérisée par lexistence dun dimorphisme sexuel (
<figureCitation box="[1127,1193,1578,1605]" captionStart="FIG" captionStartId="6.[677,688,1410,1428]" captionTargetBox="[678,1209,1168,1373]" captionTargetId="figure-400@6.[677,1210,1167,1374]" captionTargetPageId="6" captionText="FIG. 6. — Tarsichthys cf. macrurus (Agassiz, 1839). Premier rayon de nageoire pelvienne du spécimen des Figons. Échelle: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4537943" httpUri="https://zenodo.org/record/4537943/files/figure.png" pageId="5" pageNumber="36">Fig. 6</figureCitation>
<figureCitation id="91EB2A335772B0097F18B32D0F8EF9A8" box="[1127,1193,1578,1605]" captionStart="FIG" captionStartId="6.[677,688,1410,1428]" captionTargetBox="[678,1209,1168,1373]" captionTargetId="figure-400@6.[677,1210,1167,1374]" captionTargetPageId="6" captionText="FIG. 6. — Tarsichthys cf. macrurus (Agassiz, 1839). Premier rayon de nageoire pelvienne du spécimen des Figons. Échelle: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4537943" httpUri="https://zenodo.org/record/4537943/files/figure.png" pageId="5" pageNumber="36">Fig. 6</figureCitation>
). De même, les os pelviens sont fortement développés: leur longueur égale approximativement les ⅔ de celle des plus longs rayons pelviens.
</paragraph>
<paragraph blockId="6.[113,646,216,274]" pageId="6" pageNumber="37">Le corps est couvert décailles cycloïdes très fines dont le champ postérieur est orné de radii.</paragraph>
<paragraph id="096F36B65771B00A7BF7B5DF091BFEFC" blockId="6.[113,646,216,274]" pageId="6" pageNumber="37">Le corps est couvert décailles cycloïdes très fines dont le champ postérieur est orné de radii.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="37" type="discussion">
<paragraph blockId="6.[112,649,311,527]" box="[113,243,311,337]" pageId="6" pageNumber="37">DISCUSSION</paragraph>
<paragraph blockId="6.[112,649,311,527]" pageId="6" pageNumber="37">
<subSubSection id="41CA653D5771B00A7B0EB4300AFFFDE2" pageId="6" pageNumber="37" type="discussion">
<paragraph id="096F36B65771B00A7B0EB4300BD4FEA2" blockId="6.[112,649,311,527]" box="[113,243,311,337]" pageId="6" pageNumber="37">DISCUSSION</paragraph>
<paragraph id="096F36B65771B00A7B0EB4500AFFFDE2" blockId="6.[112,649,311,527]" pageId="6" pageNumber="37">
Tous les caractères méristiques de ce spécimen sintègrent parfaitement dans les limites de variabilité de lespèce
<taxonomicName authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" class="Pisces (awaiting allocation)" family="Pisces (awaiting allocation)" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pisces (awaiting allocation)" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="macrurus">
<emphasis box="[310,539,406,432]" italics="true" pageId="6" pageNumber="37">Tarsichthys macrurus</emphasis>
<taxonomicName id="CED04D355771B00A7A49B4910B9FFE3D" authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="3BA4EAA45771B00A7A49B491093CFE5D" box="[310,539,406,432]" italics="true" pageId="6" pageNumber="37">Tarsichthys macrurus</emphasis>
(Agassiz, 1839)
</taxonomicName>
, de lOligocène supérieur de Stösschen et Rott (
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<collectingCountry id="71C776265771B00A7BCFB4D20A02FE1D" box="[176,293,469,496]" name="Germany" pageId="6" pageNumber="37">Allemagne</collectingCountry>
) (
<bibRefCitation author="GAUDANT J." box="[312,473,469,495]" pageId="6" pageNumber="37" pagination="121 - 177" refId="ref6960" refString="GAUDANT J. 2002. - Nouvelles recherches sur l'ichthyofaune des lignites feuilletes oligocenes de Rott, Stosschen am Minderberg et Orsberg (Siebengebirge, Allemagne). Palaeontographica (A) 265: 121 - 177." type="journal article" year="2002">Gaudant 2002</bibRefCitation>
<bibRefCitation id="6D414B475771B00A7A47B4D20AFEFE02" author="GAUDANT J." box="[312,473,469,495]" pageId="6" pageNumber="37" pagination="121 - 177" refId="ref6960" refString="GAUDANT J. 2002. - Nouvelles recherches sur l'ichthyofaune des lignites feuilletes oligocenes de Rott, Stosschen am Minderberg et Orsberg (Siebengebirge, Allemagne). Palaeontographica (A) 265: 121 - 177." type="journal article" year="2002">Gaudant 2002</bibRefCitation>
). Cest pourquoi nous le rapportons à cette espèce.
</paragraph>
</subSubSection>

View file

@ -1,133 +1,134 @@
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<taxonomicName authorityName="Troschel" authorityYear="1854" box="[303,419,1483,1509]" class="Actinopterygii" family="Cyprinidae" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="33" phylum="Chordata" rank="genus">
<emphasis box="[303,419,1483,1509]" italics="true" pageId="2" pageNumber="33">Tarsichthys</emphasis>
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<emphasis id="3BA4EAA45775B00E7A50B0CC0A84FA08" box="[303,419,1483,1509]" italics="true" pageId="2" pageNumber="33">Tarsichthys</emphasis>
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sp.
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<paragraph blockId="2.[339,420,1515,1541]" box="[339,420,1515,1541]" pageId="2" pageNumber="33">
<subSubSection id="41CA653D5775B00F7A2CB0EC0F3EFE9C" lastPageId="3" lastPageNumber="34" pageId="2" pageNumber="33" type="description">
<paragraph id="096F36B65775B00E7A2CB0EC0A83F9E8" blockId="2.[339,420,1515,1541]" box="[339,420,1515,1541]" pageId="2" pageNumber="33">
(
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<figureCitation id="91EB2A335775B00E7A24B0EC0ABBF9E8" box="[347,412,1515,1541]" captionStart="FIG" captionStartId="2.[113,124,1161,1179]" captionTargetBox="[177,1145,215,1124]" captionTargetId="figure-173@2.[176,1145,214,1124]" captionTargetPageId="2" captionText="FIG. 2. — Tarsichthys sp. Débris dun individu découvert près de léchangeur de la Chevalière (MNHN.F.AIX226): A, moitié postérieure du corps dépourvue de restes de la colonne vertébrale; B, dent pharyngienne isolée; C, écaille isolée. Échelles: A, 1 cm; B, C, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4537933" httpUri="https://zenodo.org/record/4537933/files/figure.png" pageId="2" pageNumber="33">Fig. 2</figureCitation>
)
</paragraph>
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<paragraph id="096F36B65775B00E7B0EB32D0F2CFAAA" blockId="2.[113,649,1578,1700]" lastBlockId="2.[675,1211,1261,1351]" pageId="2" pageNumber="33">
Lunique spécimen de
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<taxonomicName id="CED04D355775B00E7A18B32D0974F9A9" authority="decouvert" authorityName="DECOUVERT" box="[359,595,1578,1604]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="34" phylum="Chordata" rank="family">Cyprinidae découvert</taxonomicName>
près de léchangeur de la Chevalière est conservé en double empreinte dans une marne blanche. Il est à la fois très incomplet et médiocrement conservé. Il sagit dun fragment de peau couvert décailles. On a recueilli en outre à proximité quelques écailles isolées et une dent pharyngienne.
</paragraph>
<paragraph blockId="2.[675,1212,1389,1700]" box="[677,971,1389,1415]" pageId="2" pageNumber="33">DESCRIPTION ANATOMIQUE</paragraph>
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<paragraph id="096F36B65775B00E79DAB06A08ECFA68" blockId="2.[675,1212,1389,1700]" box="[677,971,1389,1415]" pageId="2" pageNumber="33">DESCRIPTION ANATOMIQUE</paragraph>
<paragraph id="096F36B65775B00F79DAB08B0958FDE2" blockId="2.[675,1212,1389,1700]" lastBlockId="3.[111,648,216,1415]" lastPageId="3" lastPageNumber="34" pageId="2" pageNumber="33">
Le fragment de tronc (
<figureCitation box="[917,998,1420,1446]" captionStart="FIG" captionStartId="2.[113,124,1161,1179]" captionTargetBox="[177,1145,215,1124]" captionTargetId="figure-173@2.[176,1145,214,1124]" captionTargetPageId="2" captionText="FIG. 2. — Tarsichthys sp. Débris dun individu découvert près de léchangeur de la Chevalière (MNHN.F.AIX226): A, moitié postérieure du corps dépourvue de restes de la colonne vertébrale; B, dent pharyngienne isolée; C, écaille isolée. Échelles: A, 1 cm; B, C, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4537933" httpUri="https://zenodo.org/record/4537933/files/figure.png" pageId="2" pageNumber="33">Fig. 2A</figureCitation>
<figureCitation id="91EB2A335775B00E78EAB08B08C1FA4B" box="[917,998,1420,1446]" captionStart="FIG" captionStartId="2.[113,124,1161,1179]" captionTargetBox="[177,1145,215,1124]" captionTargetId="figure-173@2.[176,1145,214,1124]" captionTargetPageId="2" captionText="FIG. 2. — Tarsichthys sp. Débris dun individu découvert près de léchangeur de la Chevalière (MNHN.F.AIX226): A, moitié postérieure du corps dépourvue de restes de la colonne vertébrale; B, dent pharyngienne isolée; C, écaille isolée. Échelles: A, 1 cm; B, C, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4537933" httpUri="https://zenodo.org/record/4537933/files/figure.png" pageId="2" pageNumber="33">Fig. 2A</figureCitation>
) correspond à la région postérieure du corps dun grand poisson dont la longueur de la partie conservée égale environ
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. Le fait que la distance entre linsertion des pelviennes, qui occupent une position abdominale, et la base des rayons de la nageoire caudale égale
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permet destimer que la longueur standard de ce poisson devait être voisine de
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. De lanale ne subsistent que la région distale des rayons. On y distingue un long rayon non bifurqué, formé darticles relativement courts par rapport à leur largeur, et cinq rayons à la fois articulés et bifurqués dont la longueur diminue rapidement vers larrière. La nageoire caudale est très incomplète, la base de ses rayons nétant pas conservée. On peut toutefois remarquer que son contour postérieur est assez faiblement échancré et quelle comportait 17 rayons principaux à la fois articulés et bifurqués, chacun deux étant divisé en huit à son extrémité distale.
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Aucun reste de vertèbre nest observable sur le corps dont le mauvais état de conservation résulte probablement du degré de putréfaction avancé auquel avait été soumis le cadavre avant son enfouissement.On remarquera cependant quà lexception de celles portées par le pédoncule caudal, les écailles, cycloïdes, de grande taille, sont fossilisées en place. Lexamen des écailles isolées montre (
<figureCitation box="[272,350,754,780]" captionStart="FIG" captionStartId="2.[113,124,1161,1179]" captionTargetBox="[177,1145,215,1124]" captionTargetId="figure-173@2.[176,1145,214,1124]" captionTargetPageId="2" captionText="FIG. 2. — Tarsichthys sp. Débris dun individu découvert près de léchangeur de la Chevalière (MNHN.F.AIX226): A, moitié postérieure du corps dépourvue de restes de la colonne vertébrale; B, dent pharyngienne isolée; C, écaille isolée. Échelles: A, 1 cm; B, C, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4537933" httpUri="https://zenodo.org/record/4537933/files/figure.png" pageId="3" pageNumber="34">Fig.2C</figureCitation>
<figureCitation id="91EB2A335774B00F7A6FB7F50A79FCE1" box="[272,350,754,780]" captionStart="FIG" captionStartId="2.[113,124,1161,1179]" captionTargetBox="[177,1145,215,1124]" captionTargetId="figure-173@2.[176,1145,214,1124]" captionTargetPageId="2" captionText="FIG. 2. — Tarsichthys sp. Débris dun individu découvert près de léchangeur de la Chevalière (MNHN.F.AIX226): A, moitié postérieure du corps dépourvue de restes de la colonne vertébrale; B, dent pharyngienne isolée; C, écaille isolée. Échelles: A, 1 cm; B, C, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4537933" httpUri="https://zenodo.org/record/4537933/files/figure.png" pageId="3" pageNumber="34">Fig.2C</figureCitation>
) quelles sont de forme ovale. Leur surface est ornée de circuli qui recoupent sur le champ postérieur des radii au nombre dune vingtaine.
</paragraph>
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<paragraph id="096F36B65774B00F7BF9B65509A4FBE7" blockId="3.[111,648,216,1415]" pageId="3" pageNumber="34">
Un opercule isolé dont la région postérieure fait malheureusement défaut permet de constater que le contour dorsal de los,faiblement concave,est pratiquement perpendiculaire au bord antérieur.Cette disposition rappelle celle observée sur les opercules des genres
<taxonomicName authority="Cuvier, 1817" authorityName="Cuvier" authorityYear="1817" box="[111,310,1008,1034]" class="Actinopterygii" family="Cyprinidae" genus="Tinca" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis box="[111,169,1009,1034]" italics="true" pageId="3" pageNumber="34">Tinca</emphasis>
<taxonomicName id="CED04D355774B00F7B10B6F60A11FBE7" authority="Cuvier, 1817" authorityName="Cuvier" authorityYear="1817" box="[111,310,1008,1034]" class="Actinopterygii" family="Cyprinidae" genus="Tinca" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="3BA4EAA45774B00F7B10B6F60B8EFBE7" box="[111,169,1009,1034]" italics="true" pageId="3" pageNumber="34">Tinca</emphasis>
Cuvier, 1817
</taxonomicName>
et
<taxonomicName authority="Obrhelova, 1969" authorityName="Obrhelova" authorityYear="1969" box="[341,639,1008,1034]" class="Actinopterygii" family="Cyprinidae" genus="Palaeotinca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis box="[341,454,1008,1034]" italics="true" pageId="3" pageNumber="34">Palaeotinca</emphasis>
<bibRefCitation author="OBRHELOVA N." box="[460,639,1008,1034]" pageId="3" pageNumber="34" pagination="39 - 52" refId="ref7118" refString="OBRHELOVA N. 1969. - Die Karpfenfische im tschechoslowakischen Susswassertertiar. Casopis pro Mineralogii a Geologii 14 (1): 39 - 52." type="journal article" year="1969">Obrhelová, 1969</bibRefCitation>
<taxonomicName id="CED04D355774B00F7A2AB6F70958FBE7" authority="Obrhelova, 1969" authorityName="Obrhelova" authorityYear="1969" box="[341,639,1008,1034]" class="Actinopterygii" family="Cyprinidae" genus="Palaeotinca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="3BA4EAA45774B00F7A2AB6F70AE1FBE7" box="[341,454,1008,1034]" italics="true" pageId="3" pageNumber="34">Palaeotinca</emphasis>
<bibRefCitation id="6D414B475774B00F7AB3B6F70958FBE7" author="OBRHELOVA N." box="[460,639,1008,1034]" pageId="3" pageNumber="34" pagination="39 - 52" refId="ref7118" refString="OBRHELOVA N. 1969. - Die Karpfenfische im tschechoslowakischen Susswassertertiar. Casopis pro Mineralogii a Geologii 14 (1): 39 - 52." type="journal article" year="1969">Obrhelová, 1969</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<paragraph id="096F36B65774B00F7BF7B11709A3FA6B" blockId="3.[111,648,216,1415]" pageId="3" pageNumber="34">
Une dent pharyngienne isolée a également été recueillie dans le même niveau (
<figureCitation box="[416,494,1071,1097]" captionStart="FIG" captionStartId="2.[113,124,1161,1179]" captionTargetBox="[177,1145,215,1124]" captionTargetId="figure-173@2.[176,1145,214,1124]" captionTargetPageId="2" captionText="FIG. 2. — Tarsichthys sp. Débris dun individu découvert près de léchangeur de la Chevalière (MNHN.F.AIX226): A, moitié postérieure du corps dépourvue de restes de la colonne vertébrale; B, dent pharyngienne isolée; C, écaille isolée. Échelles: A, 1 cm; B, C, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4537933" httpUri="https://zenodo.org/record/4537933/files/figure.png" pageId="3" pageNumber="34">Fig. 2B</figureCitation>
<figureCitation id="91EB2A335774B00F7ADFB1280AC9FBA4" box="[416,494,1071,1097]" captionStart="FIG" captionStartId="2.[113,124,1161,1179]" captionTargetBox="[177,1145,215,1124]" captionTargetId="figure-173@2.[176,1145,214,1124]" captionTargetPageId="2" captionText="FIG. 2. — Tarsichthys sp. Débris dun individu découvert près de léchangeur de la Chevalière (MNHN.F.AIX226): A, moitié postérieure du corps dépourvue de restes de la colonne vertébrale; B, dent pharyngienne isolée; C, écaille isolée. Échelles: A, 1 cm; B, C, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4537933" httpUri="https://zenodo.org/record/4537933/files/figure.png" pageId="3" pageNumber="34">Fig. 2B</figureCitation>
). Il sagit dune dent assez grande provenant très vraisemblablement de lindividu décrit précédemment. La longueur de cette dent égale environ
<quantity box="[356,443,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="34" unit="mm" value="4.5">4,5 mm</quantity>
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. Cette dent, du
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« en crochet» («Hakenzahn»de Rutte 1962) possède une couronne dont la largeur maximale, mesurée près de lextrémité proximale de larea masticatrice, est un peu inférieure à la moitié de la longueur de la couronne. Larea masticatrice, en forme de sillon étroit et allongé, détermine avec laxe longitudinal de la dent un angle denviron 25° (cf.Rutte 1962: fig. 3).
</paragraph>
<paragraph blockId="3.[111,646,1452,1700]" box="[111,243,1452,1478]" pageId="3" pageNumber="34">D ISCUSSION</paragraph>
<paragraph blockId="3.[111,646,1452,1700]" lastBlockId="3.[674,1210,216,369]" pageId="3" pageNumber="34">
<paragraph id="096F36B65774B00F7B10B0AB0BD4FA29" blockId="3.[111,646,1452,1700]" box="[111,243,1452,1478]" pageId="3" pageNumber="34">D ISCUSSION</paragraph>
<paragraph id="096F36B65774B00F7B0EB0CB0F3EFE9C" blockId="3.[111,646,1452,1700]" lastBlockId="3.[674,1210,216,369]" pageId="3" pageNumber="34">
La morphologie de cette dent évoque à la fois celle de certaines dents pharyngiennes de
<taxonomicName authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" class="Actinopterygii" family="Cyprinidae" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="macrurus">
<emphasis box="[445,646,1515,1541]" italics="true" pageId="3" pageNumber="34">Tarsichthys macrurus</emphasis>
<taxonomicName id="CED04D355774B00F7AC2B0EC0A2DF9C8" authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="3BA4EAA45774B00F7AC2B0EC09A1F9E9" box="[445,646,1515,1541]" italics="true" pageId="3" pageNumber="34">Tarsichthys macrurus</emphasis>
(Agassiz,1839)
</taxonomicName>
provenant de lOligocène supérieur de Rott (
<collectingCountry box="[169,282,1578,1605]" name="Germany" pageId="3" pageNumber="34">Allemagne</collectingCountry>
<collectingCountry id="71C776265774B00F7BD6B32D0A3DF9A8" box="[169,282,1578,1605]" name="Germany" pageId="3" pageNumber="34">Allemagne</collectingCountry>
) (
<bibRefCitation author="GAUDANT J." box="[301,456,1578,1604]" pageId="3" pageNumber="34" pagination="121 - 177" refId="ref6960" refString="GAUDANT J. 2002. - Nouvelles recherches sur l'ichthyofaune des lignites feuilletes oligocenes de Rott, Stosschen am Minderberg et Orsberg (Siebengebirge, Allemagne). Palaeontographica (A) 265: 121 - 177." type="journal article" year="2002">Gaudant 2002</bibRefCitation>
<bibRefCitation id="6D414B475774B00F7A52B32D0AEFF9A9" author="GAUDANT J." box="[301,456,1578,1604]" pageId="3" pageNumber="34" pagination="121 - 177" refId="ref6960" refString="GAUDANT J. 2002. - Nouvelles recherches sur l'ichthyofaune des lignites feuilletes oligocenes de Rott, Stosschen am Minderberg et Orsberg (Siebengebirge, Allemagne). Palaeontographica (A) 265: 121 - 177." type="journal article" year="2002">Gaudant 2002</bibRefCitation>
: fig. 28) et celle de certaines dents de lespèce
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<emphasis box="[385,587,1610,1636]" italics="true" pageId="3" pageNumber="34">Palaeotinca egeriana</emphasis>
<bibRefCitation author="OBRHELOVA N." pageId="3" pageNumber="34" pagination="39 - 52" refId="ref7118" refString="OBRHELOVA N. 1969. - Die Karpfenfische im tschechoslowakischen Susswassertertiar. Casopis pro Mineralogii a Geologii 14 (1): 39 - 52." type="journal article" year="1969">Obrhelová, 1969</bibRefCitation>
<taxonomicName id="CED04D355774B00F7AFEB34D0BD0F96E" authority="Obrhelova, 1969" authorityName="Obrhelova" authorityYear="1969" class="Actinopterygii" family="Cyprinidae" genus="Palaeotinca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="egeriana">
<emphasis id="3BA4EAA45774B00F7AFEB34D096CF989" box="[385,587,1610,1636]" italics="true" pageId="3" pageNumber="34">Palaeotinca egeriana</emphasis>
<bibRefCitation id="6D414B475774B00F792EB34D0BD0F96E" author="OBRHELOVA N." pageId="3" pageNumber="34" pagination="39 - 52" refId="ref7118" refString="OBRHELOVA N. 1969. - Die Karpfenfische im tschechoslowakischen Susswassertertiar. Casopis pro Mineralogii a Geologii 14 (1): 39 - 52." type="journal article" year="1969">Obrhelová, 1969</bibRefCitation>
</taxonomicName>
figurées par
<bibRefCitation author="OBRHELOVA N." box="[376,558,1641,1668]" pageId="3" pageNumber="34" pagination="99 - 209" refId="ref7144" refString="OBRHELOVA N. 1970. - Die Osteologie der Vorlaufer vonTinca tinca (Pisces) aus dem Susswasser der CSSR. Abhandlungen des staatlichen Museums fur Mineralogie und Geologie zu Dresden 16: 99 - 209." type="journal article" year="1970">Obrhelová (1970</bibRefCitation>
<bibRefCitation id="6D414B475774B00F7A07B36E0909F969" author="OBRHELOVA N." box="[376,558,1641,1668]" pageId="3" pageNumber="34" pagination="99 - 209" refId="ref7144" refString="OBRHELOVA N. 1970. - Die Osteologie der Vorlaufer vonTinca tinca (Pisces) aus dem Susswasser der CSSR. Abhandlungen des staatlichen Museums fur Mineralogie und Geologie zu Dresden 16: 99 - 209." type="journal article" year="1970">Obrhelová (1970</bibRefCitation>
: fig. 32; pl.10, fig.8). Pour leur part, les écailles ressemblent à celles de lespèce
<taxonomicName authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" box="[845,1210,216,242]" class="Actinopterygii" family="Cyprinidae" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="macrurus">
<emphasis box="[845,1049,216,242]" italics="true" pageId="3" pageNumber="34">Tarsichthys macrurus</emphasis>
<taxonomicName id="CED04D355774B00F7832B5DF0F9DFF1F" authority="(Agassiz, 1839)" baseAuthorityName="Agassiz" baseAuthorityYear="1839" box="[845,1210,216,242]" genus="Tarsichthys" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="3" pageNumber="34" phylum="Chordata" rank="species" species="macrurus">
<emphasis id="3BA4EAA45774B00F7832B5DF0F3EFF1F" box="[845,1049,216,242]" italics="true" pageId="3" pageNumber="34">Tarsichthys macrurus</emphasis>
(Agassiz, 1839)
</taxonomicName>
figurées par
<bibRefCitation author="GAUDANT J." box="[796,959,247,274]" pageId="3" pageNumber="34" pagination="121 - 177" refId="ref6960" refString="GAUDANT J. 2002. - Nouvelles recherches sur l'ichthyofaune des lignites feuilletes oligocenes de Rott, Stosschen am Minderberg et Orsberg (Siebengebirge, Allemagne). Palaeontographica (A) 265: 121 - 177." type="journal article" year="2002">Gaudant (2002</bibRefCitation>
<bibRefCitation id="6D414B475774B00F7863B5F00898FEFF" author="GAUDANT J." box="[796,959,247,274]" pageId="3" pageNumber="34" pagination="121 - 177" refId="ref6960" refString="GAUDANT J. 2002. - Nouvelles recherches sur l'ichthyofaune des lignites feuilletes oligocenes de Rott, Stosschen am Minderberg et Orsberg (Siebengebirge, Allemagne). Palaeontographica (A) 265: 121 - 177." type="journal article" year="2002">Gaudant (2002</bibRefCitation>
: pl. 8, fig.3). En conclusion, en dépit du caractère fragmentaire de ce fossile, il est possible de le considérer comme un représentant du genre
<emphasis box="[771,872,342,368]" italics="true" pageId="3" pageNumber="34">Tarsichthy</emphasis>
<emphasis id="3BA4EAA45774B00F787CB451084FFE9D" box="[771,872,342,368]" italics="true" pageId="3" pageNumber="34">Tarsichthy</emphasis>
s Troschel, 1854.
</paragraph>
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@ -1,185 +1,186 @@
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<taxonomicName authorityName="Laube" authorityYear="1901" box="[732,887,1515,1541]" class="Actinopterygii" family="Cyprinidae" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis box="[732,887,1515,1541]" italics="true" pageId="11" pageNumber="42">Protothymallus</emphasis>
<treatment id="817987A0577CB00379A3B0EC0A36FD83" ID-DOI="http://doi.org/10.5281/zenodo.4818172" ID-GBIF-Taxon="182400790" ID-Zenodo-Dep="4818172" LSID="urn:lsid:plazi:treatment:817987A0577CB00379A3B0EC0A36FD83" httpUri="http://treatment.plazi.org/id/817987A0577CB00379A3B0EC0A36FD83" lastPageId="15" lastPageNumber="45" pageId="11" pageNumber="42" scope_family="Cyprinidae">
<subSubSection id="41CA653D577CB00779A3B0EC08D3F9C9" pageId="11" pageNumber="42" type="nomenclature">
<paragraph id="096F36B6577CB00779A3B0EC08D3F9C9" blockId="11.[732,1155,1515,1572]" pageId="11" pageNumber="42">
<taxonomicName id="CED04D35577CB00779A3B0EC0850F9E8" authorityName="Laube" authorityYear="1901" box="[732,887,1515,1541]" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="3BA4EAA4577CB00779A3B0EC0850F9E8" box="[732,887,1515,1541]" italics="true" pageId="11" pageNumber="42">Protothymallus</emphasis>
</taxonomicName>
sp. vel
<emphasis box="[968,1117,1515,1541]" italics="true" pageId="11" pageNumber="42">Varhostichthys</emphasis>
<emphasis id="3BA4EAA4577CB00778B7B0EC0F7AF9E8" box="[968,1117,1515,1541]" italics="true" pageId="11" pageNumber="42">Varhostichthys</emphasis>
sp. (
<figureCitation box="[881,963,1546,1572]" captionStart="FIG" captionStartId="13.[112,123,749,767]" captionTargetBox="[112,1209,214,711]" captionTargetId="figure-400@13.[112,1209,214,711]" captionTargetPageId="13" captionText="FIG. 13. — Protothymallus sp. vel Varhostichthys sp. Vue générale dun squelette découvert dans lhorizon IV du Centre commercial ouest. Spécimen MNHN.F.AIX229. Échelle: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4537960" httpUri="https://zenodo.org/record/4537960/files/figure.png" pageId="11" pageNumber="42">Figs 13</figureCitation>
<figureCitation id="91EB2A33577CB007780EB30D08E4F9C9" box="[881,963,1546,1572]" captionStart="FIG" captionStartId="13.[112,123,749,767]" captionTargetBox="[112,1209,214,711]" captionTargetId="figure-400@13.[112,1209,214,711]" captionTargetPageId="13" captionText="FIG. 13. — Protothymallus sp. vel Varhostichthys sp. Vue générale dun squelette découvert dans lhorizon IV du Centre commercial ouest. Spécimen MNHN.F.AIX229. Échelle: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4537960" httpUri="https://zenodo.org/record/4537960/files/figure.png" pageId="11" pageNumber="42">Figs 13</figureCitation>
;
<figureCitation box="[975,1003,1546,1572]" captionStart="FIG" captionStartId="14.[678,688,847,865]" captionTargetBox="[684,1196,218,789]" captionTargetId="figure-540@14.[678,1204,197,805]" captionTargetPageId="14" captionText="FIG. 14.— Protothymallus sp.vel Varhostichthys sp.Dents pharyngiennes provenant de lhorizon IV du Centre commercial ouest. Spécimen MNHN.F.AIX230. Échelle: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4537965" httpUri="https://zenodo.org/record/4537965/files/figure.png" pageId="11" pageNumber="42">14</figureCitation>
<figureCitation id="91EB2A33577CB00778B0B30D08CCF9C9" box="[975,1003,1546,1572]" captionStart="FIG" captionStartId="14.[678,688,847,865]" captionTargetBox="[684,1196,218,789]" captionTargetId="figure-540@14.[678,1204,197,805]" captionTargetPageId="14" captionText="FIG. 14.— Protothymallus sp.vel Varhostichthys sp.Dents pharyngiennes provenant de lhorizon IV du Centre commercial ouest. Spécimen MNHN.F.AIX230. Échelle: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4537965" httpUri="https://zenodo.org/record/4537965/files/figure.png" pageId="11" pageNumber="42">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="42" type="description">
<paragraph blockId="11.[675,1209,1610,1700]" box="[677,971,1610,1636]" pageId="11" pageNumber="42">DESCRIPTION ANATOMIQUE</paragraph>
<paragraph blockId="11.[675,1209,1610,1700]" pageId="11" pageNumber="42">Lors des travaux de terrassement réalisés en 1981 pour la construction dun hypermarché du Cen-</paragraph>
<subSubSection id="41CA653D577CB00779DAB34D0F9EF94E" pageId="11" pageNumber="42" type="description">
<paragraph id="096F36B6577CB00779DAB34D08ECF98F" blockId="11.[675,1209,1610,1700]" box="[677,971,1610,1636]" pageId="11" pageNumber="42">DESCRIPTION ANATOMIQUE</paragraph>
<paragraph id="096F36B6577CB00779DAB36D0F9EF94E" blockId="11.[675,1209,1610,1700]" pageId="11" pageNumber="42">Lors des travaux de terrassement réalisés en 1981 pour la construction dun hypermarché du Cen-</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4537957" ID-Zenodo-Dep="4537957" httpUri="https://zenodo.org/record/4537957/files/figure.png" pageId="12" pageNumber="43" startId="12.[113,124,1013,1031]" targetBox="[113,1209,215,971]" targetPageId="12">
<paragraph blockId="12.[113,1210,1013,1054]" pageId="12" pageNumber="43">
<caption id="5DAF663E577BB0007B0EB6F20A44FBF3" ID-DOI="http://doi.org/10.5281/zenodo.4537957" ID-Zenodo-Dep="4537957" httpUri="https://zenodo.org/record/4537957/files/figure.png" pageId="12" pageNumber="43" startId="12.[113,124,1013,1031]" targetBox="[113,1209,215,971]" targetPageId="12">
<paragraph id="096F36B6577BB0007B0EB6F20A44FBF3" blockId="12.[113,1210,1013,1054]" pageId="12" pageNumber="43">
FIG. 12. —
<taxonomicName authorityName="Obrhelova" authorityYear="1969" box="[208,304,1013,1031]" class="Actinopterygii" family="Cyprinidae" genus="Palaeotinca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis box="[208,304,1013,1031]" italics="true" pageId="12" pageNumber="43">Palaeotinca</emphasis>
<taxonomicName id="CED04D35577BB0007BAFB6F20A17FBEA" authorityName="Obrhelova" authorityYear="1969" box="[208,304,1013,1031]" class="Actinopterygii" family="Cyprinidae" genus="Palaeotinca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis id="3BA4EAA4577BB0007BAFB6F20A17FBEA" box="[208,304,1013,1031]" italics="true" pageId="12" pageNumber="43">Palaeotinca</emphasis>
</taxonomicName>
? sp. Région postabdominale dun poisson provenant de lhorizon XIV du chemin du Pin (MNHN.F.AIX228). Échelles: A, 1 cm; B, 1 mm.
</paragraph>
</caption>
<subSubSection pageId="12" pageNumber="43" type="reference_group">
<paragraph blockId="12.[112,648,1134,1700]" pageId="12" pageNumber="43">
<subSubSection id="41CA653D577BB0007B0EB1680908FAEA" pageId="12" pageNumber="43" type="reference_group">
<paragraph id="096F36B6577BB0007B0EB1680908FAEA" blockId="12.[112,648,1134,1700]" pageId="12" pageNumber="43">
tre commercial ouest, nous avons recueilli deux spécimens incomplets attribuables soit au genre
<emphasis box="[114,274,1198,1224]" italics="true" pageId="12" pageNumber="43">Varhostichthys</emphasis>
<bibRefCitation author="OBRHELOVA N." box="[284,488,1198,1224]" pageId="12" pageNumber="43" pagination="39 - 52" refId="ref7118" refString="OBRHELOVA N. 1969. - Die Karpfenfische im tschechoslowakischen Susswassertertiar. Casopis pro Mineralogii a Geologii 14 (1): 39 - 52." type="journal article" year="1969">Obrhelová, 1969</bibRefCitation>
<emphasis id="3BA4EAA4577BB0007B0DB1A90A35FB25" box="[114,274,1198,1224]" italics="true" pageId="12" pageNumber="43">Varhostichthys</emphasis>
<bibRefCitation id="6D414B47577BB0007A63B1A90ACFFB25" author="OBRHELOVA N." box="[284,488,1198,1224]" pageId="12" pageNumber="43" pagination="39 - 52" refId="ref7118" refString="OBRHELOVA N. 1969. - Die Karpfenfische im tschechoslowakischen Susswassertertiar. Casopis pro Mineralogii a Geologii 14 (1): 39 - 52." type="journal article" year="1969">Obrhelová, 1969</bibRefCitation>
, soit à
<taxonomicName authority="Laube, 1901" authorityName="Laube" authorityYear="1901" class="Actinopterygii" family="Cyprinidae" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="12" pageNumber="43">Protothymallus</emphasis>
<taxonomicName id="CED04D35577BB000793EB1A80A5EFB05" authority="Laube, 1901" authorityName="Laube" authorityYear="1901" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="43" phylum="Chordata" rank="genus">
<emphasis id="3BA4EAA4577BB000793EB1A80BFDFB0A" italics="true" pageId="12" pageNumber="43">Protothymallus</emphasis>
Laube, 1901
</taxonomicName>
. Malheureusement, ces deux spécimens sont dépourvus de tête.
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="45" pageId="12" pageNumber="43" type="description">
<paragraph blockId="12.[112,648,1134,1700]" lastBlockId="12.[675,1212,1134,1699]" pageId="12" pageNumber="43">
<subSubSection id="41CA653D577BB0027BF6B00A0A38FA6B" lastPageId="14" lastPageNumber="45" pageId="12" pageNumber="43" type="description">
<paragraph id="096F36B6577BB0007BF6B00A0F9EF94E" blockId="12.[112,648,1134,1700]" lastBlockId="12.[675,1212,1134,1699]" pageId="12" pageNumber="43">
Comme le montre le spécimen le plus complet, le corps est élancé. Sa hauteur maximale devait égaler environ un quart de la longueur standard, laquelle était un peu inférieure à
<quantity box="[544,642,1388,1415]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="43" unit="mm" value="150.0">150 mm</quantity>
<quantity id="CE289B53577BB000795FB06B09A5FA6A" box="[544,642,1388,1415]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="43" unit="mm" value="150.0">150 mm</quantity>
. La colonne vertébrale se compose de 40 ou 41 vertèbres dont 24 ou 25 constituent la région abdominale (y compris les centra modifiés pour former lappareil de Weber). Le nombre de vertèbres postabdominales est de 17 sur lun des spécimens et seulement de 15 sur lautre. Les centra vertébraux supportent des neurapophyses et des hémapophyses bien développées dont lextrémité distale atteint, dans la région abdominale, les ⅔ de la distance séparant les parties dorsale et ventrale des centra, respectivement des bords dorsal et ventral de lanimal. Des epineuralia et des epipleuralia sont observables dans la région postabdominale. Les premiers sont également présents dans la région abdominale. Quelques traces de dorsospinalia sont visibles entre les premières neurapophyses abdominales. On remarque en particulier la présence du premier dentre eux, en forme de lame aplatie dans le plan sagittal, en avant de la neurapophyse portée par le premier centrum abdominal libre. Les côtes pleurales sont au nombre de 15 ou 16 paires. Elles sont très développées puisque, à lexception de celles situées en arrière des nageoires pelviennes, leur extrémité distale atteint pratiquement le bord ventral de la cavité abdominale. En avant de la première paire de côtes pleurales sobserve la
</paragraph>
<paragraph blockId="13.[136,217,78,95]" box="[136,217,78,95]" pageId="13" pageNumber="44">Gaudant J.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4537960" ID-Zenodo-Dep="4537960" httpUri="https://zenodo.org/record/4537960/files/figure.png" pageId="13" pageNumber="44" startId="13.[112,123,749,767]" targetBox="[112,1209,214,711]" targetPageId="13">
<paragraph blockId="13.[112,1203,749,790]" pageId="13" pageNumber="44">
<paragraph id="096F36B6577AB0017BF7B5490BFEFFB3" blockId="13.[136,217,78,95]" box="[136,217,78,95]" pageId="13" pageNumber="44">Gaudant J.</paragraph>
<caption id="5DAF663E577AB0017B0FB7EA0937FCFB" ID-DOI="http://doi.org/10.5281/zenodo.4537960" ID-Zenodo-Dep="4537960" httpUri="https://zenodo.org/record/4537960/files/figure.png" pageId="13" pageNumber="44" startId="13.[112,123,749,767]" targetBox="[112,1209,214,711]" targetPageId="13">
<paragraph id="096F36B6577AB0017B0FB7EA0937FCFB" blockId="13.[112,1203,749,790]" pageId="13" pageNumber="44">
FIG. 13. —
<taxonomicName authorityName="Laube" authorityYear="1901" box="[204,328,749,767]" class="Pisces (awaiting allocation)" family="Pisces (awaiting allocation)" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pisces (awaiting allocation)" pageId="13" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis box="[204,328,749,767]" italics="true" pageId="13" pageNumber="44">Protothymallus</emphasis>
<taxonomicName id="CED04D35577AB0017BB3B7EA0A6FFD12" authorityName="Laube" authorityYear="1901" box="[204,328,749,767]" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="3BA4EAA4577AB0017BB3B7EA0A6FFD12" box="[204,328,749,767]" italics="true" pageId="13" pageNumber="44">Protothymallus</emphasis>
</taxonomicName>
sp. vel
<emphasis box="[392,513,749,767]" italics="true" pageId="13" pageNumber="44">Varhostichthys</emphasis>
<emphasis id="3BA4EAA4577AB0017AF7B7EA0926FD12" box="[392,513,749,767]" italics="true" pageId="13" pageNumber="44">Varhostichthys</emphasis>
sp. Vue générale dun squelette découvert dans lhorizon IV du Centre commercial ouest. Spécimen MNHN.F.AIX229. Échelle: 1 cm.
</paragraph>
</caption>
<paragraph blockId="13.[112,649,849,1699]" pageId="13" pageNumber="44">paire de parapophyses très développées portées par le quatrième centrum modifié pour constituer lappareil de Weber.</paragraph>
<paragraph blockId="13.[112,649,849,1699]" pageId="13" pageNumber="44">La nageoire caudale est assez grande puisque sa longueur devait être comprise un peu moins de cinq fois dans la longueur standard. Elle est profondément bifurquée, la longueur de ses lépidotriches axiaux nexcédant pas la moitié de celle des plus longs rayons de chaque lobe. Elle se compose de 19 rayons principaux, dont 17 bifurqués, en avant desquels prennent place, dorsalement et ventralement, respectivement 9 et 8 rayons marginaux.</paragraph>
<paragraph blockId="13.[112,649,849,1699]" pageId="13" pageNumber="44">La nageoire caudale est soutenue par un squelette caudal axial composé de quatre vertèbres. On dis - tingue à larrière le complexe uro-terminal, prolongé par le pleurostyle qui est redressé denviron 40° par rapport à laxe longitudinal de la colonne vertébrale. Le complexe uro-terminal supporte ventralement le parhypural et cinq hypuraux. Il est surmonté par un arc neural réduit auquel fait suite un épural unique. En avant du complexe uro-terminal, trois centra préuraux libres portent des neurapophyses et hémapophyses allongées qui sont également plus ou moins élargies pour soutenir les rayons marginaux dorsaux et ventraux de la nageoire caudale.</paragraph>
<paragraph blockId="13.[675,1211,849,1700]" pageId="13" pageNumber="44">La nageoire dorsale était probablement insérée un peu en arrière du milieu du corps et légèrement en arrière de la verticale passant par lorigine des pelviennes. Elle débute par deux rayons courts en arrière desquels prend place un long rayon articulé qui est le plus développé de la nageoire et dont la longueur représente approximativement les ¾ de la hauteur maximale du corps. Plus en arrière, on dénombre sept lépidotriches à la fois articulés et bifurqués dont la longueur diminue progressivement vers larrière.</paragraph>
<paragraph blockId="13.[675,1211,849,1700]" pageId="13" pageNumber="44">Lendosquelette de la nageoire dorsale se compose de huit ptérygiophores. À lexception du premier dentre eux qui est bifide, les ptérygiophores sont constitués dun épaississement osseux longitudinal sur lequel prend appui une lamelle osseuse triangulaire disposée dans le plan sagittal.</paragraph>
<paragraph blockId="13.[675,1211,849,1700]" pageId="13" pageNumber="44">La nageoire anale occupe une position reculée. Composée de neuf rayons, elle débute par deux rayons courts en arrière desquels prend place un long rayon articulé qui nest pas bifurqué. La partie postérieure de la nageoire est composée de six (ou sept?) lépidotriches à la fois articulés et bifurqués dont la longueur diminue assez rapidement vers larrière.</paragraph>
<paragraph blockId="13.[675,1211,849,1700]" lastBlockId="14.[112,649,216,1414]" lastPageId="14" lastPageNumber="45" pageId="13" pageNumber="44">Lendosquelette de la nageoire anale est formé de six ou sept ptérygiophores. Les premiers dentre eux, très longs, sintercalent profondément entre les hémapophyses correspondantes.</paragraph>
<paragraph blockId="14.[112,649,216,1414]" pageId="14" pageNumber="45">Les nageoires pectorales, qui sont composées de 17 rayons, sont insérées en position basse sur le flanc. Elles sont de taille modérée puisque lextrémité distale de leurs plus longs rayons ne dépasse quà peine le milieu de lespace séparant la base des pectorales de celle des pelviennes.</paragraph>
<paragraph blockId="14.[112,649,216,1414]" pageId="14" pageNumber="45">Les nageoires pelviennes, situées un peu plus près de lorigine de lanale que de la base des pectorales, sont relativement grandes puisque la longueur de leurs rayons les plus développés est approximativement égale à celle des plus longs rayons des pectorales. Elles semblent avoir été constituées de neuf rayons, dont huit à la fois articulés et bifurqués, auxquels sajoute dorsalement un très court rayon accolé au rayon non bifurqué.</paragraph>
<paragraph blockId="14.[112,649,216,1414]" pageId="14" pageNumber="45">
<paragraph id="096F36B6577AB0017B0EB6560A68FC46" blockId="13.[112,649,849,1699]" pageId="13" pageNumber="44">paire de parapophyses très développées portées par le quatrième centrum modifié pour constituer lappareil de Weber.</paragraph>
<paragraph id="096F36B6577AB0017BF7B6B60A52FB0A" blockId="13.[112,649,849,1699]" pageId="13" pageNumber="44">La nageoire caudale est assez grande puisque sa longueur devait être comprise un peu moins de cinq fois dans la longueur standard. Elle est profondément bifurquée, la longueur de ses lépidotriches axiaux nexcédant pas la moitié de celle des plus longs rayons de chaque lobe. Elle se compose de 19 rayons principaux, dont 17 bifurqués, en avant desquels prennent place, dorsalement et ventralement, respectivement 9 et 8 rayons marginaux.</paragraph>
<paragraph id="096F36B6577AB0017BF7B1E90A4BF94E" blockId="13.[112,649,849,1699]" pageId="13" pageNumber="44">La nageoire caudale est soutenue par un squelette caudal axial composé de quatre vertèbres. On dis - tingue à larrière le complexe uro-terminal, prolongé par le pleurostyle qui est redressé denviron 40° par rapport à laxe longitudinal de la colonne vertébrale. Le complexe uro-terminal supporte ventralement le parhypural et cinq hypuraux. Il est surmonté par un arc neural réduit auquel fait suite un épural unique. En avant du complexe uro-terminal, trois centra préuraux libres portent des neurapophyses et hémapophyses allongées qui sont également plus ou moins élargies pour soutenir les rayons marginaux dorsaux et ventraux de la nageoire caudale.</paragraph>
<paragraph id="096F36B6577AB00179C4B65508D7FB45" blockId="13.[675,1211,849,1700]" pageId="13" pageNumber="44">La nageoire dorsale était probablement insérée un peu en arrière du milieu du corps et légèrement en arrière de la verticale passant par lorigine des pelviennes. Elle débute par deux rayons courts en arrière desquels prend place un long rayon articulé qui est le plus développé de la nageoire et dont la longueur représente approximativement les ¾ de la hauteur maximale du corps. Plus en arrière, on dénombre sept lépidotriches à la fois articulés et bifurqués dont la longueur diminue progressivement vers larrière.</paragraph>
<paragraph id="096F36B6577AB00179C4B1A90F51FA8B" blockId="13.[675,1211,849,1700]" pageId="13" pageNumber="44">Lendosquelette de la nageoire dorsale se compose de huit ptérygiophores. À lexception du premier dentre eux qui est bifide, les ptérygiophores sont constitués dun épaississement osseux longitudinal sur lequel prend appui une lamelle osseuse triangulaire disposée dans le plan sagittal.</paragraph>
<paragraph id="096F36B6577AB00179C4B06A0851F989" blockId="13.[675,1211,849,1700]" pageId="13" pageNumber="44">La nageoire anale occupe une position reculée. Composée de neuf rayons, elle débute par deux rayons courts en arrière desquels prend place un long rayon articulé qui nest pas bifurqué. La partie postérieure de la nageoire est composée de six (ou sept?) lépidotriches à la fois articulés et bifurqués dont la longueur diminue assez rapidement vers larrière.</paragraph>
<paragraph id="096F36B6577AB00279C4B36E0ADCFEFC" blockId="13.[675,1211,849,1700]" lastBlockId="14.[112,649,216,1414]" lastPageId="14" lastPageNumber="45" pageId="13" pageNumber="44">Lendosquelette de la nageoire anale est formé de six ou sept ptérygiophores. Les premiers dentre eux, très longs, sintercalent profondément entre les hémapophyses correspondantes.</paragraph>
<paragraph id="096F36B65779B0027BF7B41F0942FE22" blockId="14.[112,649,216,1414]" pageId="14" pageNumber="45">Les nageoires pectorales, qui sont composées de 17 rayons, sont insérées en position basse sur le flanc. Elles sont de taille modérée puisque lextrémité distale de leurs plus longs rayons ne dépasse quà peine le milieu de lespace séparant la base des pectorales de celle des pelviennes.</paragraph>
<paragraph id="096F36B65779B0027BF7B4D10BFBFCE1" blockId="14.[112,649,216,1414]" pageId="14" pageNumber="45">Les nageoires pelviennes, situées un peu plus près de lorigine de lanale que de la base des pectorales, sont relativement grandes puisque la longueur de leurs rayons les plus développés est approximativement égale à celle des plus longs rayons des pectorales. Elles semblent avoir été constituées de neuf rayons, dont huit à la fois articulés et bifurqués, auxquels sajoute dorsalement un très court rayon accolé au rayon non bifurqué.</paragraph>
<paragraph id="096F36B65779B0027BF7B6140A2FFB65" blockId="14.[112,649,216,1414]" pageId="14" pageNumber="45">
Les écailles, de
<typeStatus box="[309,357,787,813]" pageId="14" pageNumber="45">type</typeStatus>
<typeStatus id="D66B88145779B0027A4AB6140A42FCC0" box="[309,357,787,813]" pageId="14" pageNumber="45">type</typeStatus>
cycloïde, paraissent avoir été relativement grandes. Leur surface porte de nombreux circuli concentriques dont la densité semble avoir été proche de 25 au millimètre. En outre, une douzaine de radii ornent le champ postérieur de lécaille, alors que le champ antérieur de lécaille semble en avoir été dépourvu. La ligne latérale est observable dans la région postabdominale. Elle prend place un peu audessus de la colonne vertébrale quelle rejoint vers lavant au niveau des dernières vertèbres abdominales.
</paragraph>
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<paragraph blockId="14.[677,1211,847,911]" pageId="14" pageNumber="45">
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<paragraph id="096F36B65779B00279D9B6480F36FC62" blockId="14.[677,1211,847,911]" pageId="14" pageNumber="45">
FIG. 14.—
<taxonomicName authorityName="Laube" authorityYear="1901" box="[762,884,847,865]" class="Pisces (awaiting allocation)" family="Pisces (awaiting allocation)" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pisces (awaiting allocation)" pageId="14" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis box="[762,884,847,865]" italics="true" pageId="14" pageNumber="45">Protothymallus</emphasis>
<taxonomicName id="CED04D355779B0027985B6480853FC8C" authorityName="Laube" authorityYear="1901" box="[762,884,847,865]" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="3BA4EAA45779B0027985B6480853FC8C" box="[762,884,847,865]" italics="true" pageId="14" pageNumber="45">Protothymallus</emphasis>
</taxonomicName>
sp.vel
<emphasis box="[943,1062,847,865]" italics="true" pageId="14" pageNumber="45">Varhostichthys</emphasis>
<emphasis id="3BA4EAA45779B00278D0B6480F01FC8C" box="[943,1062,847,865]" italics="true" pageId="14" pageNumber="45">Varhostichthys</emphasis>
sp.Dents pharyngiennes provenant de lhorizon IV du Centre commercial ouest. Spécimen MNHN.F.AIX230. Échelle: 1 mm.
</paragraph>
</caption>
<paragraph blockId="14.[112,649,216,1414]" pageId="14" pageNumber="45">
<paragraph id="096F36B65779B0027BF7B1880A38FA6B" blockId="14.[112,649,216,1414]" pageId="14" pageNumber="45">
Deux dents pharyngiennes isolées du
<typeStatus box="[551,598,1167,1193]" pageId="14" pageNumber="45">type</typeStatus>
<typeStatus id="D66B88145779B0027958B1880971FB44" box="[551,598,1167,1193]" pageId="14" pageNumber="45">type</typeStatus>
« en crochet » (« Hakenzahn » de Rutte 1962) ont été recueillies près des débris de la tête du second spécimen (
<figureCitation box="[231,310,1261,1287]" captionStart="FIG" captionStartId="14.[678,688,847,865]" captionTargetBox="[684,1196,218,789]" captionTargetId="figure-540@14.[678,1204,197,805]" captionTargetPageId="14" captionText="FIG. 14.— Protothymallus sp.vel Varhostichthys sp.Dents pharyngiennes provenant de lhorizon IV du Centre commercial ouest. Spécimen MNHN.F.AIX230. Échelle: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4537965" httpUri="https://zenodo.org/record/4537965/files/figure.png" pageId="14" pageNumber="45">Fig. 14</figureCitation>
<figureCitation id="91EB2A335779B0027B98B1EA0A11FAEA" box="[231,310,1261,1287]" captionStart="FIG" captionStartId="14.[678,688,847,865]" captionTargetBox="[684,1196,218,789]" captionTargetId="figure-540@14.[678,1204,197,805]" captionTargetPageId="14" captionText="FIG. 14.— Protothymallus sp.vel Varhostichthys sp.Dents pharyngiennes provenant de lhorizon IV du Centre commercial ouest. Spécimen MNHN.F.AIX230. Échelle: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4537965" httpUri="https://zenodo.org/record/4537965/files/figure.png" pageId="14" pageNumber="45">Fig. 14</figureCitation>
). Lune est une dent postérieure effilée, de forme sigmoïde. La seconde possède une couronne renflée au-dessous de larea masticatrice qui détermine un angle de 40° avec laxe de la couronne.
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="46" pageId="14" pageNumber="45" type="discussion">
<paragraph blockId="14.[113,647,1452,1699]" box="[113,243,1452,1478]" pageId="14" pageNumber="45">DISCUSSION</paragraph>
<paragraph blockId="14.[113,647,1452,1699]" lastBlockId="15.[112,648,216,622]" lastPageId="15" lastPageNumber="46" pageId="14" pageNumber="45">
<subSubSection id="41CA653D5779B0037B0EB0AB0A36FD83" lastPageId="15" lastPageNumber="46" pageId="14" pageNumber="45" type="discussion">
<paragraph id="096F36B65779B0027B0EB0AB0BD4FA29" blockId="14.[113,647,1452,1699]" box="[113,243,1452,1478]" pageId="14" pageNumber="45">DISCUSSION</paragraph>
<paragraph id="096F36B65779B0037B0EB0CC0A36FD83" blockId="14.[113,647,1452,1699]" lastBlockId="15.[112,648,216,622]" lastPageId="15" lastPageNumber="46" pageId="14" pageNumber="45">
Lensemble des caractères décrits précédemment permet daffirmer que ces deux fossiles appartiennent bien à la famille des
<taxonomicName box="[431,556,1546,1572]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="45" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
<taxonomicName id="CED04D355779B0027AD0B30D090BF9C9" box="[431,556,1546,1572]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="45" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
comme lindiquent notamment la disposition relative des nageoires, la structure du squelette caudal axial, la possession dun appareil de Weber dont témoigne la présence des longues parapophyses en avant de la première paire de côtes pleurales. Au sein de cette famille, la composition de la colonne vertébrale, formée de 24 ou 25 vertèbres abdominales et de 15 à 17 vertèbres postabdominales, conduit à attribuer ces poissons soit au genre
<emphasis box="[780,933,1134,1160]" italics="true" pageId="14" pageNumber="45">Varhostichthys</emphasis>
<bibRefCitation author="OBRHELOVA N." box="[942,1136,1134,1160]" pageId="14" pageNumber="45" pagination="39 - 52" refId="ref7118" refString="OBRHELOVA N. 1969. - Die Karpfenfische im tschechoslowakischen Susswassertertiar. Casopis pro Mineralogii a Geologii 14 (1): 39 - 52." type="journal article" year="1969">Obrhelová, 1969</bibRefCitation>
<emphasis id="3BA4EAA45779B0027873B1690882FB65" box="[780,933,1134,1160]" italics="true" pageId="14" pageNumber="45">Varhostichthys</emphasis>
<bibRefCitation id="6D414B475779B00278D1B1690F57FB65" author="OBRHELOVA N." box="[942,1136,1134,1160]" pageId="14" pageNumber="45" pagination="39 - 52" refId="ref7118" refString="OBRHELOVA N. 1969. - Die Karpfenfische im tschechoslowakischen Susswassertertiar. Casopis pro Mineralogii a Geologii 14 (1): 39 - 52." type="journal article" year="1969">Obrhelová, 1969</bibRefCitation>
, soit à
<taxonomicName authority="Laube, 1901" authorityName="Laube" authorityYear="1901" box="[677,990,1166,1192]" class="Pisces (awaiting allocation)" family="Pisces (awaiting allocation)" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pisces (awaiting allocation)" pageId="14" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis box="[677,838,1166,1192]" italics="true" pageId="14" pageNumber="45">Protothymallus</emphasis>
<taxonomicName id="CED04D355779B00279DAB18908F9FB45" authority="Laube, 1901" authorityName="Laube" authorityYear="1901" box="[677,990,1166,1192]" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="3BA4EAA45779B00279DAB1890861FB45" box="[677,838,1166,1192]" italics="true" pageId="14" pageNumber="45">Protothymallus</emphasis>
Laube, 1901
</taxonomicName>
. On notera en effet que les deux spécimens dAix-en-Provence possèdent un corps plus élancé que les représentants de lespèce
<typeStatus box="[809,858,1262,1288]" pageId="14" pageNumber="45">type</typeStatus>
<emphasis box="[869,1099,1261,1287]" italics="true" pageId="14" pageNumber="45">Varhostichthys brevis</emphasis>
<typeStatus id="D66B88145779B0027856B1E9087DFAE5" box="[809,858,1262,1288]" pageId="14" pageNumber="45">type</typeStatus>
<emphasis id="3BA4EAA45779B002781AB1EA0F6CFAEA" box="[869,1099,1261,1287]" italics="true" pageId="14" pageNumber="45">Varhostichthys brevis</emphasis>
(Agassiz, 1839), de lOligocène inférieur de Valeč (
<collectingCountry box="[1104,1198,1293,1320]" name="Czech Republic" pageId="14" pageNumber="45">Bohême</collectingCountry>
<collectingCountry id="71C776265779B0027F2FB00A0F89FAC5" box="[1104,1198,1293,1320]" name="Czech Republic" pageId="14" pageNumber="45">Bohême</collectingCountry>
). En effet, la hauteur maximale de leur corps égale environ le quart de la longueur standard alors que, chez
<emphasis box="[731,949,1388,1414]" italics="true" pageId="14" pageNumber="45">Varhostichthys brevis</emphasis>
<emphasis id="3BA4EAA45779B00279A4B06B0892FA6B" box="[731,949,1388,1414]" italics="true" pageId="14" pageNumber="45">Varhostichthys brevis</emphasis>
(Agassiz, 1839), elle atteindrait 40 % de celle-ci selon
<bibRefCitation author="BOHME M." box="[1021,1183,1420,1446]" pageId="14" pageNumber="45" pagination="175 - 194" refId="ref6794" refString="BOHME M. 2007. - Revision of the cyprinids from the Early Oligocene of the Ceske Stredohori mountains, and the phylogenetic relationships of Protothymallus Laube, 1901 (Teleostei, Cyprinidae, Gobioninae). Sbornik Narodniho Muzea v Praze (B) 63 (2 - 4): 175 - 194." type="journal article" year="2007">Böhme (2007)</bibRefCitation>
<bibRefCitation id="6D414B475779B0027882B08B0FB8FA4B" author="BOHME M." box="[1021,1183,1420,1446]" pageId="14" pageNumber="45" pagination="175 - 194" refId="ref6794" refString="BOHME M. 2007. - Revision of the cyprinids from the Early Oligocene of the Ceske Stredohori mountains, and the phylogenetic relationships of Protothymallus Laube, 1901 (Teleostei, Cyprinidae, Gobioninae). Sbornik Narodniho Muzea v Praze (B) 63 (2 - 4): 175 - 194." type="journal article" year="2007">Böhme (2007)</bibRefCitation>
et seulement un tiers selon
<bibRefCitation author="GAUDANT J." box="[960,1148,1451,1477]" pageId="14" pageNumber="45" pagination="281 - 312" refId="ref6998" refString="GAUDANT J. 2009. - Note complementaire sur l'ichthyofaune oligocene de Seifhennersdorf (Saxe, Allemagne) et de Varnsdorf, Kundratice, Lbin, Skalice, Knizeci, etc. (Boheme, Republique tcheque). Annalen des Naturhistorischen Museums inWien 111 A: 281 - 312." type="journal article" year="2009">Gaudant (2009)</bibRefCitation>
<bibRefCitation id="6D414B475779B00278BFB0AC0F5BFA28" author="GAUDANT J." box="[960,1148,1451,1477]" pageId="14" pageNumber="45" pagination="281 - 312" refId="ref6998" refString="GAUDANT J. 2009. - Note complementaire sur l'ichthyofaune oligocene de Seifhennersdorf (Saxe, Allemagne) et de Varnsdorf, Kundratice, Lbin, Skalice, Knizeci, etc. (Boheme, Republique tcheque). Annalen des Naturhistorischen Museums inWien 111 A: 281 - 312." type="journal article" year="2009">Gaudant (2009)</bibRefCitation>
. Or, la hauteur maximale du corps est généralement comprise entre 20 et 25 % chez
<taxonomicName authority="(Kramberger, 1885)" baseAuthorityName="Kramberger" baseAuthorityYear="1885" class="Pisces (awaiting allocation)" family="Pisces (awaiting allocation)" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pisces (awaiting allocation)" pageId="14" pageNumber="45" phylum="Chordata" rank="species" species="elongatus">
<emphasis italics="true" pageId="14" pageNumber="45">Protothymallus elongatus</emphasis>
<taxonomicName id="CED04D355779B0027F66B0EC0F21F9C8" authority="(Kramberger, 1885)" baseAuthorityName="Kramberger" baseAuthorityYear="1885" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="45" phylum="Chordata" rank="species" species="elongatus">
<emphasis id="3BA4EAA45779B0027F66B0EC082BF9C9" italics="true" pageId="14" pageNumber="45">Protothymallus elongatus</emphasis>
(Kramberger, 1885)
</taxonomicName>
, une espèce de
<taxonomicName authority="de l'Oligocene" authorityName="de l'Oligocene" box="[677,967,1578,1604]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="45" phylum="Chordata" rank="family">Cyprinidae de lOligocène</taxonomicName>
<taxonomicName id="CED04D355779B00279DAB32D08E0F9A9" authority="de l'Oligocene" authorityName="de l'Oligocene" box="[677,967,1578,1604]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="45" phylum="Chordata" rank="family">Cyprinidae de lOligocène</taxonomicName>
de
<collectingCountry box="[1005,1096,1578,1604]" name="Czech Republic" pageId="14" pageNumber="45">Bohême</collectingCountry>
<collectingCountry id="71C776265779B0027892B32D0F6FF9A9" box="[1005,1096,1578,1604]" name="Czech Republic" pageId="14" pageNumber="45">Bohême</collectingCountry>
et de
<collectingRegion box="[1161,1209,1578,1604]" country="Germany" name="Sachsen" pageId="14" pageNumber="45">Saxe</collectingRegion>
<collectingRegion id="CB14F8545779B0027FF6B32D0F9EF9A9" box="[1161,1209,1578,1604]" country="Germany" name="Sachsen" pageId="14" pageNumber="45">Saxe</collectingRegion>
(
<collectingCountry box="[683,805,1610,1637]" name="Germany" pageId="14" pageNumber="45">Allemagne</collectingCountry>
<collectingCountry id="71C776265779B00279D4B34D0802F988" box="[683,805,1610,1637]" name="Germany" pageId="14" pageNumber="45">Allemagne</collectingCountry>
) qui possède une colonne vertébrale comportant 23-24 vertèbres abdominales et 15- 16 postabdominales, et des nageoires dorsale et anale composées respectivement de (9) 10-11 rayons et de 9-10 rayons (
<bibRefCitation author="GAUDANT J." box="[415,588,247,274]" pageId="15" pageNumber="46" pagination="281 - 312" refId="ref6998" refString="GAUDANT J. 2009. - Note complementaire sur l'ichthyofaune oligocene de Seifhennersdorf (Saxe, Allemagne) et de Varnsdorf, Kundratice, Lbin, Skalice, Knizeci, etc. (Boheme, Republique tcheque). Annalen des Naturhistorischen Museums inWien 111 A: 281 - 312." type="journal article" year="2009">Gaudant 2009</bibRefCitation>
<bibRefCitation id="6D414B475778B0037AE0B5F0096BFEFF" author="GAUDANT J." box="[415,588,247,274]" pageId="15" pageNumber="46" pagination="281 - 312" refId="ref6998" refString="GAUDANT J. 2009. - Note complementaire sur l'ichthyofaune oligocene de Seifhennersdorf (Saxe, Allemagne) et de Varnsdorf, Kundratice, Lbin, Skalice, Knizeci, etc. (Boheme, Republique tcheque). Annalen des Naturhistorischen Museums inWien 111 A: 281 - 312." type="journal article" year="2009">Gaudant 2009</bibRefCitation>
). Les caractères méristiques de cette espèce sont donc très proches de ceux observés sur les deux spécimens dAix-en-Provence. Labsence probable de radii sur le champ antérieur des écailles incite donc à penser que les deux spécimens décrits ci-dessus pourraient être des représentants du genre
<taxonomicName authority="Laube, 1901" authorityName="Laube" authorityYear="1901" box="[185,503,469,495]" class="Pisces (awaiting allocation)" family="Pisces (awaiting allocation)" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pisces (awaiting allocation)" pageId="15" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis box="[185,348,469,495]" italics="true" pageId="15" pageNumber="46">Protothymallus</emphasis>
<taxonomicName id="CED04D355778B0037BC6B4D20AD0FE02" authority="Laube, 1901" authorityName="Laube" authorityYear="1901" box="[185,503,469,495]" genus="Protothymallus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="46" phylum="Chordata" rank="genus">
<emphasis id="3BA4EAA45778B0037BC6B4D20A7BFE02" box="[185,348,469,495]" italics="true" pageId="15" pageNumber="46">Protothymallus</emphasis>
Laube, 1901
</taxonomicName>
car, comme la montré
<bibRefCitation author="BOHME M." box="[231,395,501,527]" pageId="15" pageNumber="46" pagination="175 - 194" refId="ref6794" refString="BOHME M. 2007. - Revision of the cyprinids from the Early Oligocene of the Ceske Stredohori mountains, and the phylogenetic relationships of Protothymallus Laube, 1901 (Teleostei, Cyprinidae, Gobioninae). Sbornik Narodniho Muzea v Praze (B) 63 (2 - 4): 175 - 194." type="journal article" year="2007">Böhme (2007)</bibRefCitation>
<bibRefCitation id="6D414B475778B0037B98B4F20AACFDE2" author="BOHME M." box="[231,395,501,527]" pageId="15" pageNumber="46" pagination="175 - 194" refId="ref6794" refString="BOHME M. 2007. - Revision of the cyprinids from the Early Oligocene of the Ceske Stredohori mountains, and the phylogenetic relationships of Protothymallus Laube, 1901 (Teleostei, Cyprinidae, Gobioninae). Sbornik Narodniho Muzea v Praze (B) 63 (2 - 4): 175 - 194." type="journal article" year="2007">Böhme (2007)</bibRefCitation>
, lespèce
<emphasis italics="true" pageId="15" pageNumber="46">Varhostichthys brevis</emphasis>
<emphasis id="3BA4EAA45778B0037A90B4F20B89FDC2" italics="true" pageId="15" pageNumber="46">Varhostichthys brevis</emphasis>
(Agassiz, 1839) possède des écailles dont les champs antérieur et postérieur sont tous deux ornés de radii.
</paragraph>
</subSubSection>

View file

@ -1,270 +1,270 @@
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<mods:namePart>Stoehr, Sabine</mods:namePart>
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<mods:namePart id="69749191D339D74B5430D6DDD06E4E63">Stoehr, Sabine</mods:namePart>
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<mods:namePart>Zhang, Haibin</mods:namePart>
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<mods:affiliation>Institute of Deep-sea Science and Engineering, Chinese Academy of Sciences, CAS, 57200 Sanya, China</mods:affiliation>
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<mods:namePart id="7EB4D29095EAC4E9DEA9DD54FBD88824">Zhang, Haibin</mods:namePart>
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<mods:affiliation id="8E0AFF40E1190C96967152E2B7109EB9">Institute of Deep-sea Science and Engineering, Chinese Academy of Sciences, CAS, 57200 Sanya, China</mods:affiliation>
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<paragraph id="F3BF83FCB26E8C711144CEFA704DBBA3" pageId="0" pageNumber="155">
<taxonomicName id="B538F413ACB74C7E09A5BF91DFAAE643" LSID="818A238B-B677-5083-8895-050FC5C469A9" authority="(A. H Clark, 1949)" baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">Ophioplinthaca semele (A. H Clark, 1949)</taxonomicName>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Ophioplinthaca semele (A. H Clark, 1949) (IDSSE-EEB-SW 0113: A-H) A dorsal disc B ventral disc C center of the disc D radial shield E oral frame F ventral arm G dorsal arm H lateral arm; Ophioplinthaca defensor Koehler, 1930 (IDSSE-EEB-SW 0112: I-Q) I dorsal disc J ventral disc K center of the disc L disc spine M oral frame N lateral disc O dorsal arm P ventral arm Q lateral arm. Abbreviations: as adoral shield, dap dorsal arm plate, das dorsal arm spine, gs genital slit, j jaw, lap lateral arm plate, m madreporite, os oral shield, rs radial shield, ts tentacle scale, vap ventral arm plate, vas ventral arm spine. Scale bars: 2 mm (I); 1 mm (A, B, F-H, J, N-P); 500 µm (C-E, K, M, Q); 200 µm (L)." figureDoi="10.3897/zookeys.1099.76479.figure14" httpUri="https://binary.pensoft.net/fig/682275" pageId="0" pageNumber="155">Fig. 14</figureCitation>
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<paragraph id="6368E61524F80AA0E8481D0073A7ABEF" pageId="0" pageNumber="155">
<figureCitation id="1C1CDC8E74515FAB68DF56F1F815734D" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Ophioplinthaca semele (A. H Clark, 1949) (IDSSE-EEB-SW 0113: A-H) A dorsal disc B ventral disc C center of the disc D radial shield E oral frame F ventral arm G dorsal arm H lateral arm; Ophioplinthaca defensor Koehler, 1930 (IDSSE-EEB-SW 0112: I-Q) I dorsal disc J ventral disc K center of the disc L disc spine M oral frame N lateral disc O dorsal arm P ventral arm Q lateral arm. Abbreviations: as adoral shield, dap dorsal arm plate, das dorsal arm spine, gs genital slit, j jaw, lap lateral arm plate, m madreporite, os oral shield, rs radial shield, ts tentacle scale, vap ventral arm plate, vas ventral arm spine. Scale bars: 2 mm (I); 1 mm (A, B, F-H, J, N-P); 500 µm (C-E, K, M, Q); 200 µm (L)." figureDoi="10.3897/zookeys.1099.76479.figure14" httpUri="https://binary.pensoft.net/fig/682275" pageId="0" pageNumber="155">Fig. 14</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="155">
<taxonomicName authorityName="A. H. Clark" authorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomitra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiomitra semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">Ophiomitra semele</taxonomicName>
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<paragraph id="9D9B6F6738D49D6BA4B1CDD8270F67D6" pageId="0" pageNumber="155">
<taxonomicName id="6DE44D8D254DD0D1A8B296822D04ACA6" authorityName="A. H. Clark" authorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomitra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiomitra semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">Ophiomitra semele</taxonomicName>
A. H Clark, 1949: 20-23, fig. 8a, b.
</paragraph>
<paragraph pageId="0" pageNumber="155">
<taxonomicName baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">Ophioplinthaca semele</taxonomicName>
<paragraph id="6E78C29FAB966ACABFF054B510E59B65" pageId="0" pageNumber="155">
<taxonomicName id="27244B2E903EAC3D188D4460FAF55B70" baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">Ophioplinthaca semele</taxonomicName>
:
<bibRefCitation author="O'Hara, TD" journalOrPublisher="Acta Oceanologica Sinica" pageId="0" pageNumber="155" publicationUrl="http://cat.inist.fr/?aModele=afficheN&amp;cpsidt=18965845" refId="B41" refString="O'Hara, TD, Stoehr, S, 2006. Deep water Ophiuroidea (Echinodermata) of New Caledonia: Ophiacanthidae and Hemieuryalidae. In: Richer de Forges B et al. (Eds) Tropical Deep-sea Benthos 24. Memoires du Museum national d'Histoire naturelle (1993), 33-141. http://cat.inist.fr/?aModele=afficheN&amp;cpsidt=18965845" title="Deep water Ophiuroidea (Echinodermata) of New Caledonia: Ophiacanthidae and Hemieuryalidae. In: Richer de Forges B et al. (Eds) Tropical Deep-sea Benthos 24. Memoires du Museum national d'Histoire naturelle (1993), 33 - 141." url="http://cat.inist.fr/?aModele=afficheN&amp;cpsidt=18965845" year="2006">
<normalizedToken originalValue="OHara">O'Hara</normalizedToken>
<bibRefCitation id="932C735E902F6BD7CDE5D991C144526B" author="O'Hara, TD" journalOrPublisher="Acta Oceanologica Sinica" pageId="0" pageNumber="155" publicationUrl="http://cat.inist.fr/?aModele=afficheN&amp;cpsidt=18965845" refId="B41" refString="O'Hara, TD, Stoehr, S, 2006. Deep water Ophiuroidea (Echinodermata) of New Caledonia: Ophiacanthidae and Hemieuryalidae. In: Richer de Forges B et al. (Eds) Tropical Deep-sea Benthos 24. Memoires du Museum national d'Histoire naturelle (1993), 33-141. http://cat.inist.fr/?aModele=afficheN&amp;cpsidt=18965845" title="Deep water Ophiuroidea (Echinodermata) of New Caledonia: Ophiacanthidae and Hemieuryalidae. In: Richer de Forges B et al. (Eds) Tropical Deep-sea Benthos 24. Memoires du Museum national d'Histoire naturelle (1993), 33 - 141." url="http://cat.inist.fr/?aModele=afficheN&amp;cpsidt=18965845" year="2006">
<normalizedToken id="5204C1CF1D28BA02DF0A3519E4B251C5" originalValue="OHara">O'Hara</normalizedToken>
and
<normalizedToken originalValue="Stöhr">Stoehr</normalizedToken>
<normalizedToken id="AC5B2B0273F481B92C8167EE08180BF2" originalValue="Stöhr">Stoehr</normalizedToken>
2006
</bibRefCitation>
: 76;
<bibRefCitation DOI="https://doi.org/10.7717/peerj.11566" author="Chen, W" journalOrPublisher="Ecology and Evolution" pageId="0" pageNumber="155" refId="B2" refString="Chen, W, Na, J, Zhang, D, 2021a. Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ. https://doi.org/10.7717/peerj.11566" title="Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ." url="https://doi.org/10.7717/peerj.11566" year="2021 a">Chen et al. 2021a</bibRefCitation>
<bibRefCitation id="22FEF310A7A9C240996CAA25173DE710" DOI="https://doi.org/10.7717/peerj.11566" author="Chen, W" journalOrPublisher="Ecology and Evolution" pageId="0" pageNumber="155" refId="B2" refString="Chen, W, Na, J, Zhang, D, 2021a. Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ. https://doi.org/10.7717/peerj.11566" title="Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ." url="https://doi.org/10.7717/peerj.11566" year="2021 a">Chen et al. 2021a</bibRefCitation>
: 14-18, fig. 6-8.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="materials_examined">
<paragraph pageId="0" pageNumber="155">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="155">
<materialsCitation accessionNumber="OK043835" collectingDate="2019-09-03" country="Guam Island" county="Southeast" elevation="1160" latitude="12.111167" location="Northwest Pacific" longLatPrecision="13" longitude="141.62117" municipality="Mariana Trench" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:818A238BB67750838895050FC5C469A9:0F7B1D7D80303B3FA23E5ED05D161F56" country="Guam Island" county="Southeast" latitude="12.111167" longLatPrecision="13" longitude="141.62117" municipality="Mariana Trench" name="Northwest Pacific">Northwest Pacific</location>
<subSubSection id="9E894A825CEAEEC845EE3E6936D0B01E" pageId="0" pageNumber="155" type="materials_examined">
<paragraph id="F8051DE06283B229F46C2FF51898DD1E" pageId="0" pageNumber="155">Material examined.</paragraph>
<paragraph id="1717AC97384FCF43C0D21B62E2BCA6C7" pageId="0" pageNumber="155">
<materialsCitation id="72B6B55B5D46B4D0EFEAE7FB51CDE93E" accessionNumber="OK043835" collectingDate="2019-09-03" country="Guam Island" county="Southeast" elevation="1160" latitude="12.111167" location="Northwest Pacific" longLatPrecision="13" longitude="141.62117" municipality="Mariana Trench" specimenCount="1">
<location id="0F7B1D7D80303B3FA23E5ED05D161F56" LSID="urn:lsid:plazi:treatment:818A238BB67750838895050FC5C469A9:0F7B1D7D80303B3FA23E5ED05D161F56" country="Guam Island" county="Southeast" latitude="12.111167" longLatPrecision="13" longitude="141.62117" municipality="Mariana Trench" name="Northwest Pacific">Northwest Pacific</location>
<specimenCount type="generic">1 specimen</specimenCount>
<specimenCount id="27E9802A77889304161F937458B85044" type="generic">1 specimen</specimenCount>
; near
<collectingMunicipality>Mariana Trench</collectingMunicipality>
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,
<collectingCounty>Southeast</collectingCounty>
<collectingCounty id="698E466671D94F57981A6B955F48344D">Southeast</collectingCounty>
of
<collectingCountry name="Guam Island">Guam Island</collectingCountry>
<collectingCountry id="DD69BE1578D059197F96E8F0ECDF0789" name="Guam Island">Guam Island</collectingCountry>
, seamount,
<geoCoordinate degrees="12" direction="north" minutes="6.67" orientation="latitude" precision="9" value="12.111167">12°6.67'N</geoCoordinate>
<geoCoordinate id="989D6A48290F2D6F3BB6AAAE1B8DF4A2" degrees="12" direction="north" minutes="6.67" orientation="latitude" precision="9" value="12.111167">12°6.67'N</geoCoordinate>
,
<geoCoordinate degrees="141" direction="east" minutes="37.27" orientation="longitude" precision="9" value="141.62117">141°37.27'E</geoCoordinate>
<geoCoordinate id="B15B4F21BCD6F327809136C5C8A04AC8" degrees="141" direction="east" minutes="37.27" orientation="longitude" precision="9" value="141.62117">141°37.27'E</geoCoordinate>
; depth
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.16" unit="m" value="1160.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.16" unit="m" value="1160.0">1160 m</elevation>
<quantity id="EE96E656F71C9DF4EB9020F8690A5BAA" metricMagnitude="3" metricUnit="m" metricValue="1.16" unit="m" value="1160.0">
<elevation id="00D5F4AEFD33674BBDEDE614D13E89E5" metricMagnitude="3" metricUnit="m" metricValue="1.16" unit="m" value="1160.0">1160 m</elevation>
</quantity>
;
<collectingDate value="2019-09-03">03 September 2019</collectingDate>
<collectingDate id="27C0AC9230B8B3F05CFA93BC389484FE" value="2019-09-03">03 September 2019</collectingDate>
; Collecting event: stn. SC033; Shenhaiyongshi msv leg; preserved in -80 °C; GenBank:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OK043835">OK043835</accessionNumber>
<accessionNumber id="F390C31F477A9A0A3C5D1F3912CFEE0B" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OK043835">OK043835</accessionNumber>
, IDSSE-EEB-SW0113
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="distribution">
<paragraph pageId="0" pageNumber="155">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="155">537-1987 m depth, southwest of Guam Island, Northwest Pacific, Hawaii Islands.</paragraph>
<subSubSection id="4FC1ACB689E8083036CD27A8B4BC9D17" pageId="0" pageNumber="155" type="distribution">
<paragraph id="2EE3C4466E14B10D0FDE5B0D10ECDCE0" pageId="0" pageNumber="155">Distribution.</paragraph>
<paragraph id="4D86B2BC27C56A08A1A4673AC08D85AE" pageId="0" pageNumber="155">537-1987 m depth, southwest of Guam Island, Northwest Pacific, Hawaii Islands.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="155" type="remarks">
<paragraph pageId="0" pageNumber="155">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="155">
<taxonomicName baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">
<emphasis italics="true" pageId="0" pageNumber="155">Ophioplinthaca semele</emphasis>
<subSubSection id="34646E42B5AF53A7A2C2E99144C517D4" pageId="0" pageNumber="155" type="remarks">
<paragraph id="F7798D00335FFB21869AA7F192586DA6" pageId="0" pageNumber="155">Remarks.</paragraph>
<paragraph id="D569E3BC10FFE0D4BAE52DEA40D36029" pageId="0" pageNumber="155">
<taxonomicName id="30E814B923A6272BD7F6ACEC6F0F24BE" baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">
<emphasis id="907A9BB33952443C31628E689F23F79A" italics="true" pageId="0" pageNumber="155">Ophioplinthaca semele</emphasis>
</taxonomicName>
was first described by A. H
<bibRefCitation author="Clark, AH" journalOrPublisher="Bishop Museum" pageId="0" pageNumber="155" pagination="3 - 133" refId="B11" refString="Clark, AH, 1949. Ophiuroidea of the Hawaiian Islands. Bulletin of the Bernice P. Bishop Museum 195: 3 - 133" title="Ophiuroidea of the Hawaiian Islands. Bulletin of the Bernice P." volume="195" year="1949">Clark (1949)</bibRefCitation>
<bibRefCitation id="2C762FE23886F6E187979597D98A493F" author="Clark, AH" journalOrPublisher="Bishop Museum" pageId="0" pageNumber="155" pagination="3 - 133" refId="B11" refString="Clark, AH, 1949. Ophiuroidea of the Hawaiian Islands. Bulletin of the Bernice P. Bishop Museum 195: 3 - 133" title="Ophiuroidea of the Hawaiian Islands. Bulletin of the Bernice P." volume="195" year="1949">Clark (1949)</bibRefCitation>
in the genus
<taxonomicName authorityName="Lyman" authorityYear="1869" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomitra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiomitra" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="155">Ophiomitra</emphasis>
<taxonomicName id="338AD00C2BC54EDB0E4058163ED4783D" authorityName="Lyman" authorityYear="1869" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomitra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophiomitra" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="genus">
<emphasis id="270E41488DFF00A55DFB855ADA3B076E" italics="true" pageId="0" pageNumber="155">Ophiomitra</emphasis>
</taxonomicName>
, then redescribed by
<bibRefCitation DOI="https://doi.org/10.7717/peerj.11566" author="Chen, W" journalOrPublisher="Ecology and Evolution" pageId="0" pageNumber="155" refId="B2" refString="Chen, W, Na, J, Zhang, D, 2021a. Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ. https://doi.org/10.7717/peerj.11566" title="Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ." url="https://doi.org/10.7717/peerj.11566" year="2021 a">Chen et al. (2021a)</bibRefCitation>
<bibRefCitation id="F942742D70995AB82DB99CF543F0957F" DOI="https://doi.org/10.7717/peerj.11566" author="Chen, W" journalOrPublisher="Ecology and Evolution" pageId="0" pageNumber="155" refId="B2" refString="Chen, W, Na, J, Zhang, D, 2021a. Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ. https://doi.org/10.7717/peerj.11566" title="Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ." url="https://doi.org/10.7717/peerj.11566" year="2021 a">Chen et al. (2021a)</bibRefCitation>
, and that recent study included rich morphological information.
<taxonomicName baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">
<emphasis italics="true" pageId="0" pageNumber="155">Ophioplinthaca semele</emphasis>
<taxonomicName id="FEECC756181ECF1CD5216E96A5EBF0C2" baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">
<emphasis id="BB88127E3A566C4826A2DD0C88A6B8E8" italics="true" pageId="0" pageNumber="155">Ophioplinthaca semele</emphasis>
</taxonomicName>
from the present study concurs largely with the holotype description and
<bibRefCitation DOI="https://doi.org/10.7717/peerj.11566" author="Chen, W" journalOrPublisher="Ecology and Evolution" pageId="0" pageNumber="155" refId="B2" refString="Chen, W, Na, J, Zhang, D, 2021a. Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ. https://doi.org/10.7717/peerj.11566" title="Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ." url="https://doi.org/10.7717/peerj.11566" year="2021 a">Chen et al. (2021a)</bibRefCitation>
<bibRefCitation id="DF97B898E0074D6FD6B3270620E89036" DOI="https://doi.org/10.7717/peerj.11566" author="Chen, W" journalOrPublisher="Ecology and Evolution" pageId="0" pageNumber="155" refId="B2" refString="Chen, W, Na, J, Zhang, D, 2021a. Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ. https://doi.org/10.7717/peerj.11566" title="Description of three species of ophioplinthacids, including a new species, from a deep seamount in the Northwest Pacific Ocean. PeerJ." url="https://doi.org/10.7717/peerj.11566" year="2021 a">Chen et al. (2021a)</bibRefCitation>
, but it differs slightly in the disc stumps at the periphery of the disc. According to the holotype description, the disc stumps had a thorny tip or three thorns in the disc center, but at the periphery these stumps were smooth. Our specimen has a crown of somewhat longer thorns, both in the center and periphery of the disc (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Ophioplinthaca semele (A. H Clark, 1949) (IDSSE-EEB-SW 0113: A-H) A dorsal disc B ventral disc C center of the disc D radial shield E oral frame F ventral arm G dorsal arm H lateral arm; Ophioplinthaca defensor Koehler, 1930 (IDSSE-EEB-SW 0112: I-Q) I dorsal disc J ventral disc K center of the disc L disc spine M oral frame N lateral disc O dorsal arm P ventral arm Q lateral arm. Abbreviations: as adoral shield, dap dorsal arm plate, das dorsal arm spine, gs genital slit, j jaw, lap lateral arm plate, m madreporite, os oral shield, rs radial shield, ts tentacle scale, vap ventral arm plate, vas ventral arm spine. Scale bars: 2 mm (I); 1 mm (A, B, F-H, J, N-P); 500 µm (C-E, K, M, Q); 200 µm (L)." figureDoi="10.3897/zookeys.1099.76479.figure14" httpUri="https://binary.pensoft.net/fig/682275" pageId="0" pageNumber="155">14A-H</figureCitation>
<figureCitation id="5A71EBC863728779E9D44BAA8C13BFB9" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Ophioplinthaca semele (A. H Clark, 1949) (IDSSE-EEB-SW 0113: A-H) A dorsal disc B ventral disc C center of the disc D radial shield E oral frame F ventral arm G dorsal arm H lateral arm; Ophioplinthaca defensor Koehler, 1930 (IDSSE-EEB-SW 0112: I-Q) I dorsal disc J ventral disc K center of the disc L disc spine M oral frame N lateral disc O dorsal arm P ventral arm Q lateral arm. Abbreviations: as adoral shield, dap dorsal arm plate, das dorsal arm spine, gs genital slit, j jaw, lap lateral arm plate, m madreporite, os oral shield, rs radial shield, ts tentacle scale, vap ventral arm plate, vas ventral arm spine. Scale bars: 2 mm (I); 1 mm (A, B, F-H, J, N-P); 500 µm (C-E, K, M, Q); 200 µm (L)." figureDoi="10.3897/zookeys.1099.76479.figure14" httpUri="https://binary.pensoft.net/fig/682275" pageId="0" pageNumber="155">14A-H</figureCitation>
).
<taxonomicName baseAuthorityName="Lyman" baseAuthorityYear="1878" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca dipsacos" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="dipsacos">
<emphasis italics="true" pageId="0" pageNumber="155">Ophioplinthaca dipsacos</emphasis>
<taxonomicName id="8CA42A2B42AB27AAD8BDB64488D772CE" baseAuthorityName="Lyman" baseAuthorityYear="1878" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca dipsacos" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="dipsacos">
<emphasis id="F165B56783F705C3775DEFF231DF5B72" italics="true" pageId="0" pageNumber="155">Ophioplinthaca dipsacos</emphasis>
</taxonomicName>
(Lyman, 1878) is one of the species that most closely resemble
<taxonomicName baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">
<emphasis italics="true" pageId="0" pageNumber="155">Ophioplinthaca semele</emphasis>
<taxonomicName id="9058EC2D5D101465F6E5D944A58F67F8" baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">
<emphasis id="B379FFD89ED9A6164A40D6AD370C8B7A" italics="true" pageId="0" pageNumber="155">Ophioplinthaca semele</emphasis>
</taxonomicName>
by having a similar number and shape of arm spines, radial shield characters, number of lateral oral papillae, similar disc spines, and separated dorsal and ventral arm plates, but it differs in the number of tentacle scales at the first to third tentacle pore, and in the shape of the tentacle scale (Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Tabular key to all species of the genus Ophioplinthaca. Abbreviations: ASE arm segment, ASS Adoral shield spine, DAP dorsal arm plate, DAS dorsal arm spines, L length, LOP lateral oral papillae, VAP ventral arm plate, VAS ventral arm spines, VMT ventralmost tooth, W width." httpUri="http://table.plazi.org/id/9F85B69899787A840F669B2BD815C383" pageId="0" pageNumber="155" tableUuid="9F85B69899787A840F669B2BD815C383">3</tableCitation>
<tableCitation id="508FE4795BA140DB8342E19FEC89C6F0" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Tabular key to all species of the genus Ophioplinthaca. Abbreviations: ASE arm segment, ASS Adoral shield spine, DAP dorsal arm plate, DAS dorsal arm spines, L length, LOP lateral oral papillae, VAP ventral arm plate, VAS ventral arm spines, VMT ventralmost tooth, W width." httpUri="http://table.plazi.org/id/9F85B69899787A840F669B2BD815C383" pageId="0" pageNumber="155" tableUuid="9F85B69899787A840F669B2BD815C383">3</tableCitation>
). Moreover,
<taxonomicName genus="O." lsidName="O. dipsacos" pageId="0" pageNumber="155" rank="species" species="dipsacos">
<emphasis italics="true" pageId="0" pageNumber="155">O. dipsacos</emphasis>
<taxonomicName id="17E539CB944214CF10D81833ADBAE499" lsidName="O. dipsacos" pageId="0" pageNumber="155" rank="species" species="dipsacos">
<emphasis id="2BAFB1F381D54E36395D7DD71EA7DAD3" italics="true" pageId="0" pageNumber="155">O. dipsacos</emphasis>
</taxonomicName>
was recorded from the Gulf of Mexico, far from the known distribution of
<taxonomicName genus="O." lsidName="O. semele" pageId="0" pageNumber="155" rank="species" species="semele">
<emphasis italics="true" pageId="0" pageNumber="155">O. semele</emphasis>
<taxonomicName id="3A3F8AEF1698EFB68A2DE85989B9757F" lsidName="O. semele" pageId="0" pageNumber="155" rank="species" species="semele">
<emphasis id="89B8DFF458453DE71570B2AC5A074546" italics="true" pageId="0" pageNumber="155">O. semele</emphasis>
</taxonomicName>
(Lyman, 1878).
<taxonomicName authorityName="Koehler" authorityYear="1922" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca globata" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="globata">
<emphasis italics="true" pageId="0" pageNumber="155">Ophioplinthaca globata</emphasis>
<taxonomicName id="AD1FBD05E1F26D16F5B5DA9FFBABD352" authorityName="Koehler" authorityYear="1922" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca globata" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="globata">
<emphasis id="26883035796F49EF57D4BFBD2E8F200E" italics="true" pageId="0" pageNumber="155">Ophioplinthaca globata</emphasis>
</taxonomicName>
,
<taxonomicName genus="O." lsidName="O. lithosora" pageId="0" pageNumber="155" rank="species" species="lithosora">
<emphasis italics="true" pageId="0" pageNumber="155">O. lithosora</emphasis>
<taxonomicName id="1EEC0CA8BB563CD72704B607B6E4A83E" lsidName="O. lithosora" pageId="0" pageNumber="155" rank="species" species="lithosora">
<emphasis id="2304DB47B1FEBC3FAB109D74299413B1" italics="true" pageId="0" pageNumber="155">O. lithosora</emphasis>
</taxonomicName>
,
<taxonomicName genus="O." lsidName="O. citata" pageId="0" pageNumber="155" rank="species" species="citata">
<emphasis italics="true" pageId="0" pageNumber="155">O. citata</emphasis>
<taxonomicName id="A9DFFCE6693F1A8EB24C3E12280ABAD0" lsidName="O. citata" pageId="0" pageNumber="155" rank="species" species="citata">
<emphasis id="45AB6A473918F91A9AE9A4CC19598A72" italics="true" pageId="0" pageNumber="155">O. citata</emphasis>
</taxonomicName>
, and
<taxonomicName genus="O." lsidName="O. clothilde" pageId="0" pageNumber="155" rank="species" species="clothilde">
<emphasis italics="true" pageId="0" pageNumber="155">O. clothilde</emphasis>
<taxonomicName id="3111F08B40BA3D8DD007DE2679D30D26" lsidName="O. clothilde" pageId="0" pageNumber="155" rank="species" species="clothilde">
<emphasis id="285EB1E65C3686E094CAF8725C87233F" italics="true" pageId="0" pageNumber="155">O. clothilde</emphasis>
</taxonomicName>
show a similar shape of the disc spines, but differ in size and other morphological characters (Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Tabular key to all species of the genus Ophioplinthaca. Abbreviations: ASE arm segment, ASS Adoral shield spine, DAP dorsal arm plate, DAS dorsal arm spines, L length, LOP lateral oral papillae, VAP ventral arm plate, VAS ventral arm spines, VMT ventralmost tooth, W width." httpUri="http://table.plazi.org/id/9F85B69899787A840F669B2BD815C383" pageId="0" pageNumber="155" tableUuid="9F85B69899787A840F669B2BD815C383">3</tableCitation>
<tableCitation id="132C76651FFC66E9652E26E876AC85A8" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Tabular key to all species of the genus Ophioplinthaca. Abbreviations: ASE arm segment, ASS Adoral shield spine, DAP dorsal arm plate, DAS dorsal arm spines, L length, LOP lateral oral papillae, VAP ventral arm plate, VAS ventral arm spines, VMT ventralmost tooth, W width." httpUri="http://table.plazi.org/id/9F85B69899787A840F669B2BD815C383" pageId="0" pageNumber="155" tableUuid="9F85B69899787A840F669B2BD815C383">3</tableCitation>
). Therefore, variations within species from our collection can be considered as intraspecific variation, rather than species delimiting characters.
</paragraph>
<caption doi="10.3897/zookeys.1099.76479.figure14" httpUri="https://binary.pensoft.net/fig/682275" pageId="0" pageNumber="155" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="155">
<emphasis bold="true" pageId="0" pageNumber="155">Figure 14.</emphasis>
<taxonomicName baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">
<emphasis italics="true" pageId="0" pageNumber="155">Ophioplinthaca semele</emphasis>
<caption id="57316A5B5C92FD41F5D7EA08D6F85B88" doi="10.3897/zookeys.1099.76479.figure14" httpUri="https://binary.pensoft.net/fig/682275" pageId="0" pageNumber="155" start="Figure 14" startId="F14">
<paragraph id="0A818C5B477E39F45DA5ADEC7EBE7D23" pageId="0" pageNumber="155">
<emphasis id="E8E42EF1D0453E4BE4BBF2EB17C593C4" bold="true" pageId="0" pageNumber="155">Figure 14.</emphasis>
<taxonomicName id="762362576B08D2C67B314EB7FFC9659E" baseAuthorityName="A. H Clark" baseAuthorityYear="1949" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca semele" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="semele">
<emphasis id="041C389A56153F4FB20D85A669B016CB" italics="true" pageId="0" pageNumber="155">Ophioplinthaca semele</emphasis>
</taxonomicName>
(A. H Clark, 1949) (IDSSE-EEB-SW0113:
<emphasis bold="true" pageId="0" pageNumber="155">A-H</emphasis>
<emphasis id="63F6ABF0A80CD5C3F240C53D54500F4C" bold="true" pageId="0" pageNumber="155">A-H</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="155">A</emphasis>
<emphasis id="61E365FF4CB53CF945FE21E0F031C096" bold="true" pageId="0" pageNumber="155">A</emphasis>
dorsal disc
<emphasis bold="true" pageId="0" pageNumber="155">B</emphasis>
<emphasis id="633FA1696A3F0A3C00C35D9000993631" bold="true" pageId="0" pageNumber="155">B</emphasis>
ventral disc
<emphasis bold="true" pageId="0" pageNumber="155">C</emphasis>
<emphasis id="0569C0F8E29B151C7653A40C4F6B587A" bold="true" pageId="0" pageNumber="155">C</emphasis>
center of the disc
<emphasis bold="true" pageId="0" pageNumber="155">D</emphasis>
<emphasis id="E80E556D17295B6061F33E78D5997243" bold="true" pageId="0" pageNumber="155">D</emphasis>
radial shield
<emphasis bold="true" pageId="0" pageNumber="155">E</emphasis>
<normalizedToken originalValue="оral">oral</normalizedToken>
<emphasis id="FAB5A99422D9538A3050582D062A4A66" bold="true" pageId="0" pageNumber="155">E</emphasis>
<normalizedToken id="6CEC09FAC50CABFA35A328F97CA62CD6" originalValue="оral">oral</normalizedToken>
frame
<emphasis bold="true" pageId="0" pageNumber="155">F</emphasis>
<emphasis id="3C90EECD78F4C673ACE6E8FBD2040896" bold="true" pageId="0" pageNumber="155">F</emphasis>
ventral arm
<emphasis bold="true" pageId="0" pageNumber="155">G</emphasis>
<emphasis id="87CECA90AC013947B6C8CB83C36AF1D2" bold="true" pageId="0" pageNumber="155">G</emphasis>
dorsal arm
<emphasis bold="true" pageId="0" pageNumber="155">H</emphasis>
<emphasis id="210BD3B17146097B74087881B5656530" bold="true" pageId="0" pageNumber="155">H</emphasis>
lateral arm;
<taxonomicName authorityName="Koehler" authorityYear="1930" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca defensor" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="defensor">
<emphasis italics="true" pageId="0" pageNumber="155">Ophioplinthaca defensor</emphasis>
<taxonomicName id="8AC904168350AD9064802D2AAC2CEC18" authorityName="Koehler" authorityYear="1930" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophioplinthaca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ophioplinthaca defensor" order="Amphilepidida" pageId="0" pageNumber="155" phylum="Echinodermata" rank="species" species="defensor">
<emphasis id="8A9FE45B2EDC3C8A76075A31389625F2" italics="true" pageId="0" pageNumber="155">Ophioplinthaca defensor</emphasis>
</taxonomicName>
Koehler, 1930 (IDSSE-EEB-SW0112:
<emphasis bold="true" pageId="0" pageNumber="155">I-Q</emphasis>
<emphasis id="23BC41DC4855FF372E4709AEA1C281DC" bold="true" pageId="0" pageNumber="155">I-Q</emphasis>
)
<emphasis bold="true" pageId="0" pageNumber="155">I</emphasis>
<emphasis id="64BF9A004C17FF7D599F5A01B5EDC5E8" bold="true" pageId="0" pageNumber="155">I</emphasis>
dorsal disc
<emphasis bold="true" pageId="0" pageNumber="155">J</emphasis>
<emphasis id="54C5185120D478DF5D9EFA44E9AC4B6F" bold="true" pageId="0" pageNumber="155">J</emphasis>
ventral disc
<emphasis bold="true" pageId="0" pageNumber="155">K</emphasis>
<emphasis id="50C598158CEFD1966FF7E73651ABF397" bold="true" pageId="0" pageNumber="155">K</emphasis>
center of the disc
<emphasis bold="true" pageId="0" pageNumber="155">L</emphasis>
<emphasis id="9850FA27FD1E8B097A235BF64721BEB9" bold="true" pageId="0" pageNumber="155">L</emphasis>
disc spine
<emphasis bold="true" pageId="0" pageNumber="155">M</emphasis>
<normalizedToken originalValue="оral">oral</normalizedToken>
<emphasis id="3E72C2451E9FF6C81B757E7419FC29DB" bold="true" pageId="0" pageNumber="155">M</emphasis>
<normalizedToken id="D94E37F63099B7949F36FFD747DFB1D9" originalValue="оral">oral</normalizedToken>
frame
<emphasis bold="true" pageId="0" pageNumber="155">N</emphasis>
<emphasis id="8B1EE2EABB40BF416F11470CD80284A7" bold="true" pageId="0" pageNumber="155">N</emphasis>
lateral disc
<emphasis bold="true" pageId="0" pageNumber="155">O</emphasis>
<emphasis id="2C99618A6AC278CB23EDE7331C11EB98" bold="true" pageId="0" pageNumber="155">O</emphasis>
dorsal arm
<emphasis bold="true" pageId="0" pageNumber="155">P</emphasis>
<emphasis id="3FCA3315C675AFC6504ACD514602963C" bold="true" pageId="0" pageNumber="155">P</emphasis>
ventral arm
<emphasis bold="true" pageId="0" pageNumber="155">Q</emphasis>
<emphasis id="855FEF1ECD31F0FB6D3C462C50E375DA" bold="true" pageId="0" pageNumber="155">Q</emphasis>
lateral arm. Abbreviations:
<emphasis bold="true" pageId="0" pageNumber="155">as</emphasis>
<emphasis id="1126BE190EF419791250A68ABA8EB4D2" bold="true" pageId="0" pageNumber="155">as</emphasis>
adoral shield,
<emphasis bold="true" pageId="0" pageNumber="155">dap</emphasis>
<emphasis id="6015357208D6D38A5C4FD9E694C1837C" bold="true" pageId="0" pageNumber="155">dap</emphasis>
dorsal arm plate,
<emphasis bold="true" pageId="0" pageNumber="155">das</emphasis>
<emphasis id="08D1AA8AD313869265AA2D59E54EC36D" bold="true" pageId="0" pageNumber="155">das</emphasis>
dorsal arm spine,
<emphasis bold="true" pageId="0" pageNumber="155">gs</emphasis>
<emphasis id="29C2E3CCE45E27EEDA273026EFFD8E59" bold="true" pageId="0" pageNumber="155">gs</emphasis>
genital slit,
<emphasis bold="true" pageId="0" pageNumber="155">j</emphasis>
<emphasis id="4EF6F56E36BF4B147ABD8D78E2394A05" bold="true" pageId="0" pageNumber="155">j</emphasis>
jaw,
<emphasis bold="true" pageId="0" pageNumber="155">lap</emphasis>
<emphasis id="C964858D8BB3879F59CFD5429DAAF6F7" bold="true" pageId="0" pageNumber="155">lap</emphasis>
lateral arm plate,
<emphasis bold="true" pageId="0" pageNumber="155">m</emphasis>
<emphasis id="584FA9B5E646CC06B32144610C3FABDB" bold="true" pageId="0" pageNumber="155">m</emphasis>
madreporite,
<emphasis bold="true" pageId="0" pageNumber="155">os</emphasis>
<emphasis id="94E50E12935A7C138645BE587D6CBBC5" bold="true" pageId="0" pageNumber="155">os</emphasis>
oral shield,
<emphasis bold="true" pageId="0" pageNumber="155">rs</emphasis>
<emphasis id="C74C1CCA41744A76855D5558097FD9D2" bold="true" pageId="0" pageNumber="155">rs</emphasis>
radial shield,
<emphasis bold="true" pageId="0" pageNumber="155">ts</emphasis>
<emphasis id="ABC26A42BA9394FC7D57EF67DD60A8CD" bold="true" pageId="0" pageNumber="155">ts</emphasis>
tentacle scale,
<emphasis bold="true" pageId="0" pageNumber="155">vap</emphasis>
<emphasis id="357AC156CC09AB3A1824D1D49E1D89CA" bold="true" pageId="0" pageNumber="155">vap</emphasis>
ventral arm plate,
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<emphasis id="42DECEFEE22379DF9A68639A3AF5976B" bold="true" pageId="0" pageNumber="155">C-E, K, M, Q</emphasis>
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<emphasis id="828C071D07E473C6B940CADC4A9982A7" bold="true" pageId="0" pageNumber="155">L</emphasis>
).
</paragraph>
</caption>

View file

@ -1,162 +1,162 @@
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<figureCitation captionStart="Figures 210" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">, 5</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4">, 11</figureCitation>
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<taxonomicNameLabel id="D2BFC57E5037BAC5544E275BC44087FA" pageId="3" pageNumber="4">sp. nov.</taxonomicNameLabel>
<figureCitation id="64FA876F55DA33BB330B390E54F14CAF" captionStart="Figures 210" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">Figs 4</figureCitation>
<figureCitation id="1CD67F4981780BFFC88C216016BA0EE3" captionStart="Figures 210" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="3" pageNumber="4">, 5</figureCitation>
<figureCitation id="2599DEF56E5416D31EEECFEEB04C5C9F" captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="3" pageNumber="4">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="materials_examined">
<paragraph pageId="4" pageNumber="5">
<pageBreakToken pageId="4" pageNumber="5" start="start">Material</pageBreakToken>
<subSubSection id="20113199AE31ADA9FCF1E63C7F365D04" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="737D5A43827B628D96ED7A980DD14AA6" pageId="4" pageNumber="5">
<pageBreakToken id="8D2E8EE9B234B8313FAEC8E8C694B1F2" pageId="4" pageNumber="5" start="start">Material</pageBreakToken>
examined.
</paragraph>
<paragraph pageId="4" pageNumber="5">
<emphasis bold="true" pageId="4" pageNumber="5">
<emphasis italics="true" pageId="4" pageNumber="5">Holotype</emphasis>
<paragraph id="EF9D1AF7ECD82AF60B9DDDB9CFD7B53D" pageId="4" pageNumber="5">
<emphasis id="8EE628DC29D604D989C8C010AE106A67" bold="true" pageId="4" pageNumber="5">
<emphasis id="FDF48E481B5D2C33CFA5DEEF26124027" italics="true" pageId="4" pageNumber="5">Holotype</emphasis>
.
</emphasis>
(Fig.
<figureCitation captionStart="Figures 210" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="4" pageNumber="5">4</figureCitation>
<figureCitation id="5CC39B051A8C5B8A15A68F7ADBDF3B6B" captionStart="Figures 210" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="4" pageNumber="5">4</figureCitation>
) Bhutan • District Thimphu, c. 5 km E of Chhuzom, W of Geneykha; in brooklet with a prayer wheel along the road; 2750 m a.s.l.;
<geoCoordinate degrees="27" direction="north" minutes="18" orientation="latitude" precision="15" seconds="43" value="27.311943">27°18'43&quot;N</geoCoordinate>
<geoCoordinate id="59623E169EEB1709F2F12494FFD528C5" degrees="27" direction="north" minutes="18" orientation="latitude" precision="15" seconds="43" value="27.311943">27°18'43&quot;N</geoCoordinate>
,
<geoCoordinate degrees="89" direction="east" minutes="36" orientation="longitude" precision="15" seconds="10" value="89.602776">89°36'10&quot;E</geoCoordinate>
<geoCoordinate id="7B1D26EDAD1E69349B8EAEBC24B45928" degrees="89" direction="east" minutes="36" orientation="longitude" precision="15" seconds="10" value="89.602776">89°36'10&quot;E</geoCoordinate>
; E. Gittenberger, Choki Gyeltshen &amp; Pema Leda leg. 25.X.2018; NBCB 1059.
<emphasis bold="true" pageId="4" pageNumber="5">
<emphasis italics="true" pageId="4" pageNumber="5">Paratype</emphasis>
<emphasis id="738A2DA94D4A024FE05F65BB60BBC560" bold="true" pageId="4" pageNumber="5">
<emphasis id="A4616A0FB32F8F91A5CC4A0D46F282C3" italics="true" pageId="4" pageNumber="5">Paratype</emphasis>
.
</emphasis>
(Fig.
<figureCitation captionStart="Figures 210" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="4" pageNumber="5">5</figureCitation>
<figureCitation id="0042F9902F8B5F7472F08729FB623FEC" captionStart="Figures 210" captionStartId="F2" captionText="Figures 2 - 10. Erhaia jannei sp. nov., holotype (2 H 2.25 mm) and paratype (3 H 2.02 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia pelkiae sp. nov., holotype (4 H 1.86 mm) and paratype (5 H 1.86 mm). Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photos by J. Goud Erhaia wangchuki Gittenberger, Sherub &amp; Stelbrink, 2017. Shells from the type locality (6 H 1.98 mm) (7 H 2.03 mm); NBCB 1072. Bhutan, district Wangdue Phodrang, Gangchhu, Gangzetem brooklet, 2883 m a. s. l. Photos by J. Goud 8 Erhaia banepaensis Nesemann &amp; S. Sharma, 2007, holotype (H 1.95 mm); NHMW 103319. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 9 Erhaia nainitalensis Davis &amp; Rao, 1997, holotype of E. chandeshwariensis Nesemann &amp; S. Sharma, 2007 (H 1.94 mm); NHMW 103315. Nepal, Central Zone, Kavre district, small forest stream, left tributary of the Chandeswari Khola upstream from Chandeshwari at Banepa. NHMW 10 Erhaia sugurensis Nesemann, Shah &amp; Tachamo, 2007, holotype (H 1.95 mm); NHMW 104172. Nepal, Central Zone, Lalitpur district, Godawari, upper reaches of Sugure Khola forest stream, 1700 m a. s. l. NHMW." httpUri="https://binary.pensoft.net/fig/402717" pageId="4" pageNumber="5">5</figureCitation>
) 1 shell; same data as for holotype; NBCB 1060.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="diagnosis">
<paragraph pageId="4" pageNumber="5">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="5">Shell with a partly reflected peristome, teleoconch with spiral lirae.</paragraph>
<subSubSection id="9CF02D6385514EF711F0D08D161EFA36" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="4ABED5CE50169F479037DF157F0F3FD4" pageId="4" pageNumber="5">Diagnosis.</paragraph>
<paragraph id="411096EC3C04197D54B49700B8B9ED65" pageId="4" pageNumber="5">Shell with a partly reflected peristome, teleoconch with spiral lirae.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="shell">
<paragraph pageId="4" pageNumber="5">Shell.</paragraph>
<paragraph pageId="4" pageNumber="5">
<subSubSection id="2CCD98A0CC6D8E7EA4CB904601E2D7AA" pageId="4" pageNumber="5" type="shell">
<paragraph id="B0775FEF686C398DA93B4074F0188DFE" pageId="4" pageNumber="5">Shell.</paragraph>
<paragraph id="E2F0743843F81A4315A3DDBCB83B9470" pageId="4" pageNumber="5">
Shell elongated ovoid, with
<normalizedToken originalValue="3½">31/2</normalizedToken>
<normalizedToken id="56E8B9ECB70B79B16AA2681DC1801047" originalValue="3½">31/2</normalizedToken>
convex, shouldered whorls that are separated by a deep suture; clearly higher than broad; light yellowish brown with fine growth lines and some brown periostracal ridges. Peristome reflected at the columellar and the basal side. Parietal and columellar side of the aperture thickened by a whitish callus. An irregular umbilical chink only in the paratype might be represent a malformation resulting from repair of the shell wall. Protoconch encrustated; teleoconch with fine spiral lines.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="description">
<paragraph pageId="4" pageNumber="5">Measurements.</paragraph>
<paragraph pageId="4" pageNumber="5">
<subSubSection id="3F148EE6435970F88E7BB2F61569AEAF" pageId="4" pageNumber="5" type="description">
<paragraph id="C3B078A425E226DFCA9404B2A20E1713" pageId="4" pageNumber="5">Measurements.</paragraph>
<paragraph id="B88B63401C5F11513687D79E81E3C779" pageId="4" pageNumber="5">
(
<emphasis italics="true" pageId="4" pageNumber="5">N</emphasis>
<emphasis id="AA7B83A46EA6AE0E91A0E0E86F284BCD" italics="true" pageId="4" pageNumber="5">N</emphasis>
= 2): holotype and paratype H 1.9 mm, B 1.1 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="ecology">
<paragraph pageId="5" pageNumber="6">
<pageBreakToken pageId="5" pageNumber="6" start="start">Ecology</pageBreakToken>
<subSubSection id="B5C1EBA6FB212C769528A9559863F060" pageId="5" pageNumber="6" type="ecology">
<paragraph id="232127952A5A55F6381E74303F688071" pageId="5" pageNumber="6">
<pageBreakToken id="B17DEFBE67CE2E378ACBCB63B5813389" pageId="5" pageNumber="6" start="start">Ecology</pageBreakToken>
</paragraph>
<paragraph pageId="5" pageNumber="6">
<paragraph id="A2DB5253509EF1BF76B1AA3D23ECDA90" pageId="5" pageNumber="6">
(Fig.
<figureCitation captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="5" pageNumber="6">11</figureCitation>
<figureCitation id="C92116728FD2701EE8C720C88B15D7B3" captionStart="Figure 11" captionStartId="F3" captionText="Figure 11. Type locality of both Erhaia jannei and E. pelkiae; Bhutan, district Thimphu, W of Geneykha, brooklet with water powered prayer wheel, 2750 m a. s. l. Photo by EG." httpUri="https://binary.pensoft.net/fig/402718" pageId="5" pageNumber="6">11</figureCitation>
). The snails are rare on the rocks in the shaded streamlet uphill the prayer wheel. See also the data for
<taxonomicName lsidName="E. jannei" pageId="5" pageNumber="6" rank="species" species="jannei">
<emphasis italics="true" pageId="5" pageNumber="6">E. jannei</emphasis>
<taxonomicName id="567371C23BBF98664BEE1D4AAC34841C" lsidName="E. jannei" pageId="5" pageNumber="6" rank="species" species="jannei">
<emphasis id="5BF2BD28CF509CD00D11F1B88632EE8C" italics="true" pageId="5" pageNumber="6">E. jannei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="notes">
<paragraph pageId="5" pageNumber="6">Notes.</paragraph>
<paragraph pageId="5" pageNumber="6">
<subSubSection id="CC52D7FFCB959DA5813D71C74054784B" pageId="5" pageNumber="6" type="notes">
<paragraph id="9965BB08AD2A43FFB260B19C86AA8610" pageId="5" pageNumber="6">Notes.</paragraph>
<paragraph id="4219553545F291A49DC5D5EFFC9230E1" pageId="5" pageNumber="6">
The differences between the sympatric
<taxonomicName lsidName="E. pelkiae" pageId="5" pageNumber="6" rank="species" species="pelkiae">
<emphasis italics="true" pageId="5" pageNumber="6">E. pelkiae</emphasis>
<taxonomicName id="9F831BDC3E97D25C65F9B9B3E747CD57" lsidName="E. pelkiae" pageId="5" pageNumber="6" rank="species" species="pelkiae">
<emphasis id="BBAE4FD40575198ED4B13B71828886E8" italics="true" pageId="5" pageNumber="6">E. pelkiae</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="E. jannei" pageId="5" pageNumber="6" rank="species" species="jannei">
<emphasis italics="true" pageId="5" pageNumber="6">E. jannei</emphasis>
<taxonomicName id="B297B0D2C4ACAC115C7C5FF973028BBD" lsidName="E. jannei" pageId="5" pageNumber="6" rank="species" species="jannei">
<emphasis id="4CDEBC6760291A73E80BC807C8566424" italics="true" pageId="5" pageNumber="6">E. jannei</emphasis>
</taxonomicName>
are too large to regard as sexual dimorphism.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="etymology">
<paragraph pageId="5" pageNumber="6">Etymology.</paragraph>
<paragraph pageId="5" pageNumber="6">
<subSubSection id="82BFB9E0525B9541ADAAB998A4488179" pageId="5" pageNumber="6" type="etymology">
<paragraph id="5E27437A785E9B8F41355D97E5DF0E90" pageId="5" pageNumber="6">Etymology.</paragraph>
<paragraph id="23928ED483340EC169AA675442033082" pageId="5" pageNumber="6">
The epithet
<taxonomicName lsidName="pelkiae" pageId="5" pageNumber="6" rank="species" species="pelkiae">
<emphasis italics="true" pageId="5" pageNumber="6">pelkiae</emphasis>
<taxonomicName id="D72451A57E9533796E2E8643FAF91E53" lsidName="pelkiae" pageId="5" pageNumber="6" rank="species" species="pelkiae">
<emphasis id="D972ECD25C3659DD176CC92C737157FD" italics="true" pageId="5" pageNumber="6">pelkiae</emphasis>
</taxonomicName>
refers to Ms. Pelki Yangdon, the daughter of the fourth author.
</paragraph>

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G.B. Sowerby I, 1823 [=
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Lamarck, 1822], by monotypy.
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<figureCitation captionStart="Fig. 2" captionStartId="F2" captionText="Fig. 2. A-G. Male subgenital plate; H. Thoracic sternites; I-P. Metasternum of Pteranabropsis species: P. carli (Griffini, 1911) (A, J); P. cuspis sp. nov. (B, H); P. copia from type locality (C, M); P. angusta sp. nov. (D, L); P. carnarius Gorochov, 1998 (E, N); P. pusilla sp. nov. (F, O); P. guadun sp. nov. (G, P); P. copia from Hoang Lien (I); P. bavi sp. nov. (K). Abbreviations: c 1 - fore coxa, c 2 - mid coxa, c 3 - hind coxa, s 1 - prosternum, s 2 - mesosternum, s 3 - metasternum, th - throat with lateral hemispherical sclerites and pair of papillae, pr - underside of pronotum. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig2" httpUri="https://binary.pensoft.net/fig/323899" pageId="4" pageNumber="111">, 2I</figureCitation>
<figureCitation captionStart="Fig. 2" captionStartId="F2" captionText="Fig. 2. A-G. Male subgenital plate; H. Thoracic sternites; I-P. Metasternum of Pteranabropsis species: P. carli (Griffini, 1911) (A, J); P. cuspis sp. nov. (B, H); P. copia from type locality (C, M); P. angusta sp. nov. (D, L); P. carnarius Gorochov, 1998 (E, N); P. pusilla sp. nov. (F, O); P. guadun sp. nov. (G, P); P. copia from Hoang Lien (I); P. bavi sp. nov. (K). Abbreviations: c 1 - fore coxa, c 2 - mid coxa, c 3 - hind coxa, s 1 - prosternum, s 2 - mesosternum, s 3 - metasternum, th - throat with lateral hemispherical sclerites and pair of papillae, pr - underside of pronotum. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig2" httpUri="https://binary.pensoft.net/fig/323899" pageId="4" pageNumber="111">, 2M</figureCitation>
<figureCitation captionStart="Fig. 4" captionStartId="F4" captionText="Fig. 4. A-K. Male paraproctal outgrowths in full or only apical area in lateral (A-B, E-F, H, J), apical (C-D, G), ventro-apical (I), and oblique apical view (K); L-O. Female abdominal apex with ovipositor in lateral view. A-D. P. copia (2 males); E-G. P. carnarius Gorochov, 1998 (two males from Tam Dao, in F the hindmost outgrowth is twisted upside down); H-I. P. pusilla sp. nov.; J-K. P. guadun sp. nov.; L-M. P. copia sp. nov. from Copia (L) and Hoang Lien (M); N. P. bavi sp. nov.; O. P. guadun sp. nov. Scales 1 mm, scale for ovipositor 10 mm." figureDoi="10.3897/jor.28.32182.fig4" httpUri="https://binary.pensoft.net/fig/323901" pageId="4" pageNumber="111">, 4A-D</figureCitation>
<figureCitation captionStart="Fig. 4" captionStartId="F4" captionText="Fig. 4. A-K. Male paraproctal outgrowths in full or only apical area in lateral (A-B, E-F, H, J), apical (C-D, G), ventro-apical (I), and oblique apical view (K); L-O. Female abdominal apex with ovipositor in lateral view. A-D. P. copia (2 males); E-G. P. carnarius Gorochov, 1998 (two males from Tam Dao, in F the hindmost outgrowth is twisted upside down); H-I. P. pusilla sp. nov.; J-K. P. guadun sp. nov.; L-M. P. copia sp. nov. from Copia (L) and Hoang Lien (M); N. P. bavi sp. nov.; O. P. guadun sp. nov. Scales 1 mm, scale for ovipositor 10 mm." figureDoi="10.3897/jor.28.32182.fig4" httpUri="https://binary.pensoft.net/fig/323901" pageId="4" pageNumber="111">, 4L-M</figureCitation>
<figureCitation captionStart="Fig. 5" captionStartId="F5" captionText="Fig. 5. Female subgenital plate of A-C. P. carli (Griffini, 1911) from Hoang Lien (A-B) and Tam Dao (C, in oblique view); D-F. P. copia sp. nov. from Hoang Lien (D-E) and Copia (F); G. P. bavi sp. nov.; H. P. pusilla sp. nov.; I. P. guadun sp. nov.; J-K. P. carnarius Gorochov, 1998 from Tam Dao (J) and from BaVi (K, abdomen laterally compressed). The white arrows point at the small lateral grooves, constrictions or faint steps of the subgenital plates. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig5" httpUri="https://binary.pensoft.net/fig/323902" pageId="4" pageNumber="111">, 5D-F</figureCitation>
<figureCitation captionStart="Fig. 6" captionStartId="F6" captionText="Fig. 6. A-E. P. carli (Griffini, 1911) male from Ngo Luong (A-D) and Tam Dao (E); F-G. P. cuspis sp. nov. male; H-I. P. angusta sp. nov. male; J-M. P. carnarius Gorochov, 1998 male (J) and female from Tam Tao (K-L) and from BaVi (M); N-O. P. pusilla sp. nov. female; P-R. P. guadun sp. nov. female; S-V. P. copia sp. nov. female (S-T) and male (U-V) from Copia. - Fore tibia external side (A, F, H, K, N, P, S) and internal side (B, E, G, I-J, L, O, Q, T); thorax in lateral view (C), prothoracic spiracle (D, M, R, V). Abbreviations: c 1 - anterior coxa, ms - mesothoracic pleurites, mt - metathoracic pleurites, s 1 - s 3 - first, second and third thoracic spiracles. Scales for legs and body, 5 mm; for tympana and spiracles, 1 mm." figureDoi="10.3897/jor.28.32182.fig6" httpUri="https://binary.pensoft.net/fig/323903" pageId="4" pageNumber="111">, 6S-V</figureCitation>
<figureCitation captionStart="Fig. 7" captionStartId="F7" captionText="Fig. 7. Male phallic complex of A-F. P. carli (Griffini, 1911) from Tam Dao (A-B), Cuc Phuong (C-D) and Ngo Luong (E-F); G-H. P. copia sp. nov.; I-J. P. cuspis sp. nov.; K-N. P. angusta sp. nov.; O-P. P. carnarius from Tam Tao; Q-R. P. pusilla sp. nov.; S-T. P. guadun sp. nov. in dorsal (A, C, F, G, I, L, M, O, Q, S) and in ventral (D, E, H, J, K, N, O, R, T) view, or in dorsal view with only dorsal lobes flipped proximad, ventral lobe distad (B). Scales 1 mm (all images)." figureDoi="10.3897/jor.28.32182.fig7" httpUri="https://binary.pensoft.net/fig/323904" pageId="4" pageNumber="111">, 7G-H</figureCitation>
<treatment id="81E24FDF8E1758AA856C33FF8B6C6B27" ID-GBIF-Taxon="182224529" LSID="urn:lsid:zoobank.org:act:B29BE0BA-E3E2-4341-8E4F-38204C1640A5" httpUri="http://treatment.plazi.org/id/81E24FDF8E1758AA856C33FF8B6C6B27" lastPageId="6" lastPageNumber="113" pageId="4" pageNumber="111" scope_family="Anostostomatidae" scope_subFamily="Anabropsinae">
<subSubSection id="C7E476B0948FC5C3A0058AF11640B498" pageId="4" pageNumber="111" type="nomenclature">
<paragraph id="DBEE068DFACC4CA2222E65112F5E4AB2" pageId="4" pageNumber="111">
<taxonomicName id="BD258A3A5DE9F0D1895330AA0C9F3426" ID-CoL="8Q3LQ" LSID="http://zoobank.org/B29BE0BA-E3E2-4341-8E4F-38204C1640A5" authority="Ingrisch, 2019" authorityName="Ingrisch" authorityYear="2019" class="Insecta" family="Anostostomatidae" genus="Pteranabropsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteranabropsis copia" order="Orthoptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="copia" status="sp. nov.">Pteranabropsis copia</taxonomicName>
<taxonomicNameLabel id="982AD4DBC802B6AE7438DDCB56B41B74" pageId="4" pageNumber="111">sp. nov.</taxonomicNameLabel>
<figureCitation id="DE781AD520BEAB0A893F20337CB3891B" captionStart="Fig. 1" captionStartId="F1" captionText="Fig. 1. Habitus dorsal view of Pteranabropsis species. A. P. carli (Griffini, 1911) female; B. P. cuspis sp. nov. male; C-E. P. copia sp. nov. male (C), female from Copia (D), and female from Hoang Lien (E); F. P. angusta sp. nov. male; G. P. bavi sp. nov. female; H. P. guadun sp. nov. female; I. P. pusilla sp. nov. male; J. P. carnarius Gorochov, 1998 male. Scales 10 mm." figureDoi="10.3897/jor.28.32182.fig1" httpUri="https://binary.pensoft.net/fig/323898" pageId="4" pageNumber="111">Figs 1C-E</figureCitation>
<figureCitation id="61415562C236BFD2CFDB4FC15C59BC5D" captionStart="Fig. 2" captionStartId="F2" captionText="Fig. 2. A-G. Male subgenital plate; H. Thoracic sternites; I-P. Metasternum of Pteranabropsis species: P. carli (Griffini, 1911) (A, J); P. cuspis sp. nov. (B, H); P. copia from type locality (C, M); P. angusta sp. nov. (D, L); P. carnarius Gorochov, 1998 (E, N); P. pusilla sp. nov. (F, O); P. guadun sp. nov. (G, P); P. copia from Hoang Lien (I); P. bavi sp. nov. (K). Abbreviations: c 1 - fore coxa, c 2 - mid coxa, c 3 - hind coxa, s 1 - prosternum, s 2 - mesosternum, s 3 - metasternum, th - throat with lateral hemispherical sclerites and pair of papillae, pr - underside of pronotum. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig2" httpUri="https://binary.pensoft.net/fig/323899" pageId="4" pageNumber="111">, 2C</figureCitation>
<figureCitation id="8B347262BC8BB8652BE3A9D216586AF7" captionStart="Fig. 2" captionStartId="F2" captionText="Fig. 2. A-G. Male subgenital plate; H. Thoracic sternites; I-P. Metasternum of Pteranabropsis species: P. carli (Griffini, 1911) (A, J); P. cuspis sp. nov. (B, H); P. copia from type locality (C, M); P. angusta sp. nov. (D, L); P. carnarius Gorochov, 1998 (E, N); P. pusilla sp. nov. (F, O); P. guadun sp. nov. (G, P); P. copia from Hoang Lien (I); P. bavi sp. nov. (K). Abbreviations: c 1 - fore coxa, c 2 - mid coxa, c 3 - hind coxa, s 1 - prosternum, s 2 - mesosternum, s 3 - metasternum, th - throat with lateral hemispherical sclerites and pair of papillae, pr - underside of pronotum. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig2" httpUri="https://binary.pensoft.net/fig/323899" pageId="4" pageNumber="111">, 2I</figureCitation>
<figureCitation id="8C8333D02C64DBA598759061291D03F5" captionStart="Fig. 2" captionStartId="F2" captionText="Fig. 2. A-G. Male subgenital plate; H. Thoracic sternites; I-P. Metasternum of Pteranabropsis species: P. carli (Griffini, 1911) (A, J); P. cuspis sp. nov. (B, H); P. copia from type locality (C, M); P. angusta sp. nov. (D, L); P. carnarius Gorochov, 1998 (E, N); P. pusilla sp. nov. (F, O); P. guadun sp. nov. (G, P); P. copia from Hoang Lien (I); P. bavi sp. nov. (K). Abbreviations: c 1 - fore coxa, c 2 - mid coxa, c 3 - hind coxa, s 1 - prosternum, s 2 - mesosternum, s 3 - metasternum, th - throat with lateral hemispherical sclerites and pair of papillae, pr - underside of pronotum. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig2" httpUri="https://binary.pensoft.net/fig/323899" pageId="4" pageNumber="111">, 2M</figureCitation>
<figureCitation id="E9148A92002C4F6EAB64FEAD4B051660" captionStart="Fig. 4" captionStartId="F4" captionText="Fig. 4. A-K. Male paraproctal outgrowths in full or only apical area in lateral (A-B, E-F, H, J), apical (C-D, G), ventro-apical (I), and oblique apical view (K); L-O. Female abdominal apex with ovipositor in lateral view. A-D. P. copia (2 males); E-G. P. carnarius Gorochov, 1998 (two males from Tam Dao, in F the hindmost outgrowth is twisted upside down); H-I. P. pusilla sp. nov.; J-K. P. guadun sp. nov.; L-M. P. copia sp. nov. from Copia (L) and Hoang Lien (M); N. P. bavi sp. nov.; O. P. guadun sp. nov. Scales 1 mm, scale for ovipositor 10 mm." figureDoi="10.3897/jor.28.32182.fig4" httpUri="https://binary.pensoft.net/fig/323901" pageId="4" pageNumber="111">, 4A-D</figureCitation>
<figureCitation id="505AD95B00C90C908F8B29C9FB2CC81C" captionStart="Fig. 4" captionStartId="F4" captionText="Fig. 4. A-K. Male paraproctal outgrowths in full or only apical area in lateral (A-B, E-F, H, J), apical (C-D, G), ventro-apical (I), and oblique apical view (K); L-O. Female abdominal apex with ovipositor in lateral view. A-D. P. copia (2 males); E-G. P. carnarius Gorochov, 1998 (two males from Tam Dao, in F the hindmost outgrowth is twisted upside down); H-I. P. pusilla sp. nov.; J-K. P. guadun sp. nov.; L-M. P. copia sp. nov. from Copia (L) and Hoang Lien (M); N. P. bavi sp. nov.; O. P. guadun sp. nov. Scales 1 mm, scale for ovipositor 10 mm." figureDoi="10.3897/jor.28.32182.fig4" httpUri="https://binary.pensoft.net/fig/323901" pageId="4" pageNumber="111">, 4L-M</figureCitation>
<figureCitation id="1B65E8167C8F1003E794D44411A77136" captionStart="Fig. 5" captionStartId="F5" captionText="Fig. 5. Female subgenital plate of A-C. P. carli (Griffini, 1911) from Hoang Lien (A-B) and Tam Dao (C, in oblique view); D-F. P. copia sp. nov. from Hoang Lien (D-E) and Copia (F); G. P. bavi sp. nov.; H. P. pusilla sp. nov.; I. P. guadun sp. nov.; J-K. P. carnarius Gorochov, 1998 from Tam Dao (J) and from BaVi (K, abdomen laterally compressed). The white arrows point at the small lateral grooves, constrictions or faint steps of the subgenital plates. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig5" httpUri="https://binary.pensoft.net/fig/323902" pageId="4" pageNumber="111">, 5D-F</figureCitation>
<figureCitation id="29BB17B64979A712EB0CD7411261FD1E" captionStart="Fig. 6" captionStartId="F6" captionText="Fig. 6. A-E. P. carli (Griffini, 1911) male from Ngo Luong (A-D) and Tam Dao (E); F-G. P. cuspis sp. nov. male; H-I. P. angusta sp. nov. male; J-M. P. carnarius Gorochov, 1998 male (J) and female from Tam Tao (K-L) and from BaVi (M); N-O. P. pusilla sp. nov. female; P-R. P. guadun sp. nov. female; S-V. P. copia sp. nov. female (S-T) and male (U-V) from Copia. - Fore tibia external side (A, F, H, K, N, P, S) and internal side (B, E, G, I-J, L, O, Q, T); thorax in lateral view (C), prothoracic spiracle (D, M, R, V). Abbreviations: c 1 - anterior coxa, ms - mesothoracic pleurites, mt - metathoracic pleurites, s 1 - s 3 - first, second and third thoracic spiracles. Scales for legs and body, 5 mm; for tympana and spiracles, 1 mm." figureDoi="10.3897/jor.28.32182.fig6" httpUri="https://binary.pensoft.net/fig/323903" pageId="4" pageNumber="111">, 6S-V</figureCitation>
<figureCitation id="7A06D573B1DCD5ACB7D186C387E8A87E" captionStart="Fig. 7" captionStartId="F7" captionText="Fig. 7. Male phallic complex of A-F. P. carli (Griffini, 1911) from Tam Dao (A-B), Cuc Phuong (C-D) and Ngo Luong (E-F); G-H. P. copia sp. nov.; I-J. P. cuspis sp. nov.; K-N. P. angusta sp. nov.; O-P. P. carnarius from Tam Tao; Q-R. P. pusilla sp. nov.; S-T. P. guadun sp. nov. in dorsal (A, C, F, G, I, L, M, O, Q, S) and in ventral (D, E, H, J, K, N, O, R, T) view, or in dorsal view with only dorsal lobes flipped proximad, ventral lobe distad (B). Scales 1 mm (all images)." figureDoi="10.3897/jor.28.32182.fig7" httpUri="https://binary.pensoft.net/fig/323904" pageId="4" pageNumber="111">, 7G-H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="112" type="holotype (male).">
<paragraph pageId="5" pageNumber="112">
<emphasis italics="true" pageId="5" pageNumber="112">
<pageBreakToken pageId="5" pageNumber="112" start="start">Holotype</pageBreakToken>
<subSubSection id="C7B763B04519DF214229E1FA35DE0FAD" pageId="5" pageNumber="112" type="holotype (male).">
<paragraph id="AB1ACE277421EAFF8621F3D784E34700" pageId="5" pageNumber="112">
<emphasis id="358C461457AD98CC9CEF633B241AE362" italics="true" pageId="5" pageNumber="112">
<pageBreakToken id="C7BB327AFC9216D2218AF5787EBD6EFC" pageId="5" pageNumber="112" start="start">Holotype</pageBreakToken>
(male).
</emphasis>
-
</paragraph>
<paragraph pageId="5" pageNumber="112">
<paragraph id="EC39F7B61890F93EDFA5B1064E44C2DF" pageId="5" pageNumber="112">
Vietnam: Son La prov., Copia Nat. Res.,
<geoCoordinate degrees="21" direction="north" minutes="22" orientation="latitude" precision="15" seconds="12" value="21.37">21°22'12&quot;N</geoCoordinate>
<geoCoordinate id="13C8108B454CDF6FFA1C7D1CEC3591C2" degrees="21" direction="north" minutes="22" orientation="latitude" precision="15" seconds="12" value="21.37">21°22'12&quot;N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="30" orientation="longitude" precision="15" seconds="42" value="103.511665">103°30'42&quot;E</geoCoordinate>
<geoCoordinate id="46AAA383313E3A028CB519AA964E6D30" degrees="103" direction="east" minutes="30" orientation="longitude" precision="15" seconds="42" value="103.511665">103°30'42&quot;E</geoCoordinate>
, 20-23.vii.2016, leg. J. Constant &amp; J. Bresseel (GTI project, I.G.: 33.282) - (Brussels, ISNB).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="112" type="materials_examined">
<paragraph pageId="5" pageNumber="112">
<emphasis italics="true" pageId="5" pageNumber="112">Other specimens examined.</emphasis>
<subSubSection id="B49A5DB40E9375110AED7A2F8013CE7A" pageId="5" pageNumber="112" type="materials_examined">
<paragraph id="182F52FA26B4A6A26DDEB4104A98D161" pageId="5" pageNumber="112">
<emphasis id="8304DAEEEB3B58F309AE56E554F6B682" italics="true" pageId="5" pageNumber="112">Other specimens examined.</emphasis>
-
</paragraph>
<paragraph pageId="5" pageNumber="112">
<materialsCitation collectingDate="2013-07-01" collectingDateMax="2013-07-05" collectingDateMin="2013-07-01" collectorName="Hoang Lien NP, J. Constant, J. Bresseel" country="Vietnam" latitude="22.35" location="Vietnam" longLatPrecision="925" longitude="103.772224" specimenCount="14" specimenCount-female="12" specimenCount-male="2" typeStatus="paratypes">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
<paragraph id="1590F7FCD5046A06D4DCC6D392D97BEB" pageId="5" pageNumber="112">
<materialsCitation id="5FDDCBC04DB5DFEF48B41922E1D03EE3" collectingDate="2013-07-01" collectingDateMax="2013-07-05" collectingDateMin="2013-07-01" collectorName="Hoang Lien NP, J. Constant, J. Bresseel" country="Vietnam" latitude="22.35" location="Vietnam" longLatPrecision="925" longitude="103.772224" specimenCount="14" specimenCount-female="12" specimenCount-male="2" typeStatus="paratypes">
<collectingCountry id="4B8BF8D968EC63447EC4FE8AFF3E84AA" name="Vietnam">Vietnam</collectingCountry>
: same data as holotype -
<specimenCount type="female">4 females</specimenCount>
<specimenCount id="5DAEE323B72386E1473803F2AD42C717" type="female">4 females</specimenCount>
,
<specimenCount type="male">2 males</specimenCount>
<specimenCount id="BA3504DDF029211E7A6A83762768DDFE" type="male">2 males</specimenCount>
(
<typeStatus>paratypes</typeStatus>
<typeStatus id="56E67C1ECC5602C55A464CC95AE5F246">paratypes</typeStatus>
) (Brussels, ISNB);
<collectorName>Hoang Lien NP</collectorName>
<collectorName id="3E461A68E77AC4853CE3C41D00E3384F">Hoang Lien NP</collectorName>
,
<geoCoordinate degrees="22" direction="north" minutes="21" orientation="latitude" precision="925" value="22.35">22°21'N</geoCoordinate>
<geoCoordinate id="633A4EEB6F3AD9557E65E03F882234C2" degrees="22" direction="north" minutes="21" orientation="latitude" precision="925" value="22.35">22°21'N</geoCoordinate>
,
<geoCoordinate degrees="103" direction="east" minutes="46" orientation="longitude" precision="15" seconds="20" value="103.772224">103°46'20&quot;E</geoCoordinate>
<geoCoordinate id="8328D57441B5B9A67839FD100AB741F3" degrees="103" direction="east" minutes="46" orientation="longitude" precision="15" seconds="20" value="103.772224">103°46'20&quot;E</geoCoordinate>
,
<collectingDate value="2013-07-01" valueMax="2013-07-05" valueMin="2013-07-01">1-5.vii.2013</collectingDate>
<collectingDate id="1CB84EFE48320ED5CC81BD71AC4BC304" value="2013-07-01" valueMax="2013-07-05" valueMin="2013-07-01">1-5.vii.2013</collectingDate>
, leg.
<collectorName>J. Constant</collectorName>
<collectorName id="A157311423472094C22C7863E35F4CE3">J. Constant</collectorName>
&amp;
<collectorName>J. Bresseel</collectorName>
<collectorName id="5BF541B322D68A4AE327866C59D1893B">J. Bresseel</collectorName>
(I.G. 32.454) -
<specimenCount type="female">8 females</specimenCount>
<specimenCount id="E186B990CEC7DD757CDB78707CE2336E" type="female">8 females</specimenCount>
(Brussels, ISNB)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="112" type="diagnosis">
<paragraph pageId="5" pageNumber="112">
<emphasis italics="true" pageId="5" pageNumber="112">Diagnosis.</emphasis>
<subSubSection id="381F0EDECDC861EDACB33EFB63E24924" pageId="5" pageNumber="112" type="diagnosis">
<paragraph id="5CF6C08A999478397C4F34FD092FC3A2" pageId="5" pageNumber="112">
<emphasis id="75D8A7EAC7833E8BF52E92175E0C4BBD" italics="true" pageId="5" pageNumber="112">Diagnosis.</emphasis>
-
</paragraph>
<paragraph pageId="5" pageNumber="112">
<paragraph id="7C494D73C10F87D78B208CCCDC2E8BB4" pageId="5" pageNumber="112">
A brachypterous species with wings that only slightly surpass the hind knees. In contrast to other brachypterous species of the genus, the fore wings of
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. copia" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="copia">
<emphasis italics="true" pageId="5" pageNumber="112">P. copia</emphasis>
<taxonomicName id="631281FBFB4061922F78C54A51B14BBE" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. copia" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="copia">
<emphasis id="733CFDF4A6F931F181DCC9BEA6F7C0AD" italics="true" pageId="5" pageNumber="112">P. copia</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="5" pageNumber="112">sp. nov.</emphasis>
<emphasis id="8D5E7E36A89584AB3F6A1FE153CCF74D" bold="true" pageId="5" pageNumber="112">sp. nov.</emphasis>
are rather wide (2.2-2.4
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="F463374D66D0AA099BED323B440CA0CA" originalValue="×">x</normalizedToken>
longer than wide) and the spread hind wings are longer such that they look more like elongated semi-circles, while in most of the other species mentioned they are more ovoid. Moreover,
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. copia" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="copia">
<emphasis italics="true" pageId="5" pageNumber="112">P. copia</emphasis>
<taxonomicName id="9FFC1E08478C55BC2A821C8B593B9FD2" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. copia" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="copia">
<emphasis id="C3F7E476FAD119A2A994EE8DCEA66207" italics="true" pageId="5" pageNumber="112">P. copia</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="5" pageNumber="112">sp. nov.</emphasis>
<emphasis id="C250EB83B68CDDCA031CB57304831996" bold="true" pageId="5" pageNumber="112">sp. nov.</emphasis>
differs from other similar species of the genus by the very narrow conical apical area of the metasternal lobes that arises in an almost step-like constriction from the wide basal area. The male paraproctal outgrowths of the new species resemble those of
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. cuspis" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="cuspis">
<emphasis italics="true" pageId="5" pageNumber="112">P. cuspis</emphasis>
<taxonomicName id="7E88BBD493B6F01D715B030A0FFB28B2" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. cuspis" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="cuspis">
<emphasis id="A302012C65E2373913DC8B55DBFC3FFC" italics="true" pageId="5" pageNumber="112">P. cuspis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="5" pageNumber="112">sp. nov.</emphasis>
<emphasis id="CE2A0FC3F2CAC5CEAAB5153884070BB7" bold="true" pageId="5" pageNumber="112">sp. nov.</emphasis>
, but the long dorsal process ends in a rather wide obtuse tip that carries a narrow stiffened rim, while in
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. cuspis" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="cuspis">
<emphasis italics="true" pageId="5" pageNumber="112">P. cuspis</emphasis>
<taxonomicName id="539B1E71F3E34BEE597FE0E15E88C805" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. cuspis" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="cuspis">
<emphasis id="772A995C4AD6B42093AF16397204B353" italics="true" pageId="5" pageNumber="112">P. cuspis</emphasis>
</taxonomicName>
it ends in a subacute tip with minute stiffened pad. The female subgenital plate of
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. copia" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="copia">
<emphasis italics="true" pageId="5" pageNumber="112">P. copia</emphasis>
<taxonomicName id="3D03BEBCF1F8610FDB5E4FAC64ECD986" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. copia" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="copia">
<emphasis id="3AB34864AF963E320BA544F3BA5D0751" italics="true" pageId="5" pageNumber="112">P. copia</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName class="Insecta" genus="P." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. carli" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="carli">
<emphasis italics="true" pageId="5" pageNumber="112">P. carli</emphasis>
<taxonomicName id="3AAF0CCD1BFBD8A654D1D969F3E1E983" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. carli" order="Orthoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="carli">
<emphasis id="23296B3C54081172CB70E57A3A21C4AE" italics="true" pageId="5" pageNumber="112">P. carli</emphasis>
</taxonomicName>
but provided in the transient zone between the wide basal and the narrow apical area with small elongate lateral grooves such that the ventral surface of the plate in that area is narrower than the distance between the dorsal lateral margins.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112" type="description">
<paragraph pageId="5" pageNumber="112">
<emphasis italics="true" pageId="5" pageNumber="112">Description.</emphasis>
<subSubSection id="9001CED4C8B8D7515DBDB018350E9D3A" lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112" type="description">
<paragraph id="8A7567BB4EC7B4A80E17A4CF11340045" pageId="5" pageNumber="112">
<emphasis id="3A9E82EF2E47D95015C023380CC99345" italics="true" pageId="5" pageNumber="112">Description.</emphasis>
-
</paragraph>
<paragraph pageId="5" pageNumber="112">
<paragraph id="09F9668823547D0B6768E40FC20E9A67" pageId="5" pageNumber="112">
Medium sized to moderately large species; general habitus as genus. Prosternal lobes near base compressed, afterwards long spiniform, thin; mesosternal lobes in basal area moderately wide, afterwards elongate conical to nearly cylindrical with obtuse tip; metasternal lobes compressed, with concave internal and strongly convex external margins; rather suddenly narrowed into nearly sub-cylindrical apical area with obtuse tip (Fig.
<figureCitation captionStart="Fig. 2" captionStartId="F2" captionText="Fig. 2. A-G. Male subgenital plate; H. Thoracic sternites; I-P. Metasternum of Pteranabropsis species: P. carli (Griffini, 1911) (A, J); P. cuspis sp. nov. (B, H); P. copia from type locality (C, M); P. angusta sp. nov. (D, L); P. carnarius Gorochov, 1998 (E, N); P. pusilla sp. nov. (F, O); P. guadun sp. nov. (G, P); P. copia from Hoang Lien (I); P. bavi sp. nov. (K). Abbreviations: c 1 - fore coxa, c 2 - mid coxa, c 3 - hind coxa, s 1 - prosternum, s 2 - mesosternum, s 3 - metasternum, th - throat with lateral hemispherical sclerites and pair of papillae, pr - underside of pronotum. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig2" httpUri="https://binary.pensoft.net/fig/323899" pageId="5" pageNumber="112">2I, M</figureCitation>
<figureCitation id="1A0FF4C45EEB61AA26CC9FCD45E81E50" captionStart="Fig. 2" captionStartId="F2" captionText="Fig. 2. A-G. Male subgenital plate; H. Thoracic sternites; I-P. Metasternum of Pteranabropsis species: P. carli (Griffini, 1911) (A, J); P. cuspis sp. nov. (B, H); P. copia from type locality (C, M); P. angusta sp. nov. (D, L); P. carnarius Gorochov, 1998 (E, N); P. pusilla sp. nov. (F, O); P. guadun sp. nov. (G, P); P. copia from Hoang Lien (I); P. bavi sp. nov. (K). Abbreviations: c 1 - fore coxa, c 2 - mid coxa, c 3 - hind coxa, s 1 - prosternum, s 2 - mesosternum, s 3 - metasternum, th - throat with lateral hemispherical sclerites and pair of papillae, pr - underside of pronotum. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig2" httpUri="https://binary.pensoft.net/fig/323899" pageId="5" pageNumber="112">2I, M</figureCitation>
).
</paragraph>
<paragraph pageId="5" pageNumber="112">
<paragraph id="CEAC0CCF46E8924791EA42FA5FB642D1" pageId="5" pageNumber="112">
Wings slightly surpassing hind knees (Fig.
<figureCitation captionStart="Fig. 1" captionStartId="F1" captionText="Fig. 1. Habitus dorsal view of Pteranabropsis species. A. P. carli (Griffini, 1911) female; B. P. cuspis sp. nov. male; C-E. P. copia sp. nov. male (C), female from Copia (D), and female from Hoang Lien (E); F. P. angusta sp. nov. male; G. P. bavi sp. nov. female; H. P. guadun sp. nov. female; I. P. pusilla sp. nov. male; J. P. carnarius Gorochov, 1998 male. Scales 10 mm." figureDoi="10.3897/jor.28.32182.fig1" httpUri="https://binary.pensoft.net/fig/323898" pageId="5" pageNumber="112">1C-E</figureCitation>
<figureCitation id="DC77BD2A1865D3D79E3E48611D0F8B1F" captionStart="Fig. 1" captionStartId="F1" captionText="Fig. 1. Habitus dorsal view of Pteranabropsis species. A. P. carli (Griffini, 1911) female; B. P. cuspis sp. nov. male; C-E. P. copia sp. nov. male (C), female from Copia (D), and female from Hoang Lien (E); F. P. angusta sp. nov. male; G. P. bavi sp. nov. female; H. P. guadun sp. nov. female; I. P. pusilla sp. nov. male; J. P. carnarius Gorochov, 1998 male. Scales 10 mm." figureDoi="10.3897/jor.28.32182.fig1" httpUri="https://binary.pensoft.net/fig/323898" pageId="5" pageNumber="112">1C-E</figureCitation>
). Fore wings 2.2-2.4
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="186897F3DAE00F853748F2A3D2756E3F" originalValue="×">x</normalizedToken>
longer than wide. Venation: radius with radius sector arising between middle and apical third of tegmen; media two-branched, branching before mid-length; cubitus anterior either three-branched, branching in basal third and behind mid-length, or two-branched, branching only in basal third; cubitus posterior free and undivided; with 5 anal veins, the last one incomplete. Hind wings nearly semicircular, about 1.4-1.55
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="EFDE00FBB91BD3066C9F57E3D27A614A" originalValue="×">x</normalizedToken>
wider than long.
</paragraph>
<paragraph pageId="5" pageNumber="112">Legs. Fore coxa with a strong spine at swollen anterior surface; mid coxa with a smaller spine at anterior margin. Fore femur with 1-4 small spines and mid femur with 0-3 spines at anterior-ventral margins; in one male, also with one spine at posterior margin of one mid femur only. Hind femur with 1-6 external and 1-6 small internal spines on ventral margins; hind tibia with dorsal spines on inner margin slightly larger than on outer margin, ventral margins with few minute spinules; on both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than external counterparts; ventral margin with 2-3 external and 1 internal spinules.</paragraph>
<paragraph pageId="5" pageNumber="112">
<paragraph id="E55FB928C047BCB2389BF96A4F7080BA" pageId="5" pageNumber="112">Legs. Fore coxa with a strong spine at swollen anterior surface; mid coxa with a smaller spine at anterior margin. Fore femur with 1-4 small spines and mid femur with 0-3 spines at anterior-ventral margins; in one male, also with one spine at posterior margin of one mid femur only. Hind femur with 1-6 external and 1-6 small internal spines on ventral margins; hind tibia with dorsal spines on inner margin slightly larger than on outer margin, ventral margins with few minute spinules; on both sides with 4 apical spurs, the dorsal two pairs very large, the following pair medium, the ventral-most pair small; internal spurs larger than external counterparts; ventral margin with 2-3 external and 1 internal spinules.</paragraph>
<paragraph id="A8796A5633E5C1DE967D6FF39BDBDF8A" pageId="5" pageNumber="112">
Male. Paraproctal outgrowths elongate, narrowest in middle, at end with rounded ventral angle, substraight but slightly oblique apical margin and elongate-triangular dorsal process, slightly tilting posteriorly and with truncate tip that carries on top a narrow stiffened lamella (Fig.
<figureCitation captionStart="Fig. 4" captionStartId="F4" captionText="Fig. 4. A-K. Male paraproctal outgrowths in full or only apical area in lateral (A-B, E-F, H, J), apical (C-D, G), ventro-apical (I), and oblique apical view (K); L-O. Female abdominal apex with ovipositor in lateral view. A-D. P. copia (2 males); E-G. P. carnarius Gorochov, 1998 (two males from Tam Dao, in F the hindmost outgrowth is twisted upside down); H-I. P. pusilla sp. nov.; J-K. P. guadun sp. nov.; L-M. P. copia sp. nov. from Copia (L) and Hoang Lien (M); N. P. bavi sp. nov.; O. P. guadun sp. nov. Scales 1 mm, scale for ovipositor 10 mm." figureDoi="10.3897/jor.28.32182.fig4" httpUri="https://binary.pensoft.net/fig/323901" pageId="5" pageNumber="112">4A-D</figureCitation>
<figureCitation id="0F6185CF1220CA464FD67854E6E27EE7" captionStart="Fig. 4" captionStartId="F4" captionText="Fig. 4. A-K. Male paraproctal outgrowths in full or only apical area in lateral (A-B, E-F, H, J), apical (C-D, G), ventro-apical (I), and oblique apical view (K); L-O. Female abdominal apex with ovipositor in lateral view. A-D. P. copia (2 males); E-G. P. carnarius Gorochov, 1998 (two males from Tam Dao, in F the hindmost outgrowth is twisted upside down); H-I. P. pusilla sp. nov.; J-K. P. guadun sp. nov.; L-M. P. copia sp. nov. from Copia (L) and Hoang Lien (M); N. P. bavi sp. nov.; O. P. guadun sp. nov. Scales 1 mm, scale for ovipositor 10 mm." figureDoi="10.3897/jor.28.32182.fig4" httpUri="https://binary.pensoft.net/fig/323901" pageId="5" pageNumber="112">4A-D</figureCitation>
). Subgenital plate in widened basal area only slightly swollen; apical area with subparallel lateral margins that very faintly diverge posteriorly; apical margin substraight but concave in middle; elongate styli inserted at apico-lateral angles (Fig.
<figureCitation captionStart="Fig. 2" captionStartId="F2" captionText="Fig. 2. A-G. Male subgenital plate; H. Thoracic sternites; I-P. Metasternum of Pteranabropsis species: P. carli (Griffini, 1911) (A, J); P. cuspis sp. nov. (B, H); P. copia from type locality (C, M); P. angusta sp. nov. (D, L); P. carnarius Gorochov, 1998 (E, N); P. pusilla sp. nov. (F, O); P. guadun sp. nov. (G, P); P. copia from Hoang Lien (I); P. bavi sp. nov. (K). Abbreviations: c 1 - fore coxa, c 2 - mid coxa, c 3 - hind coxa, s 1 - prosternum, s 2 - mesosternum, s 3 - metasternum, th - throat with lateral hemispherical sclerites and pair of papillae, pr - underside of pronotum. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig2" httpUri="https://binary.pensoft.net/fig/323899" pageId="5" pageNumber="112">2C</figureCitation>
<figureCitation id="EF2F44CF5B65013208B6DD7E11AEAC7D" captionStart="Fig. 2" captionStartId="F2" captionText="Fig. 2. A-G. Male subgenital plate; H. Thoracic sternites; I-P. Metasternum of Pteranabropsis species: P. carli (Griffini, 1911) (A, J); P. cuspis sp. nov. (B, H); P. copia from type locality (C, M); P. angusta sp. nov. (D, L); P. carnarius Gorochov, 1998 (E, N); P. pusilla sp. nov. (F, O); P. guadun sp. nov. (G, P); P. copia from Hoang Lien (I); P. bavi sp. nov. (K). Abbreviations: c 1 - fore coxa, c 2 - mid coxa, c 3 - hind coxa, s 1 - prosternum, s 2 - mesosternum, s 3 - metasternum, th - throat with lateral hemispherical sclerites and pair of papillae, pr - underside of pronotum. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig2" httpUri="https://binary.pensoft.net/fig/323899" pageId="5" pageNumber="112">2C</figureCitation>
). Phallus membranous (Fig.
<figureCitation captionStart="Fig. 7" captionStartId="F7" captionText="Fig. 7. Male phallic complex of A-F. P. carli (Griffini, 1911) from Tam Dao (A-B), Cuc Phuong (C-D) and Ngo Luong (E-F); G-H. P. copia sp. nov.; I-J. P. cuspis sp. nov.; K-N. P. angusta sp. nov.; O-P. P. carnarius from Tam Tao; Q-R. P. pusilla sp. nov.; S-T. P. guadun sp. nov. in dorsal (A, C, F, G, I, L, M, O, Q, S) and in ventral (D, E, H, J, K, N, O, R, T) view, or in dorsal view with only dorsal lobes flipped proximad, ventral lobe distad (B). Scales 1 mm (all images)." figureDoi="10.3897/jor.28.32182.fig7" httpUri="https://binary.pensoft.net/fig/323904" pageId="5" pageNumber="112">7G-H</figureCitation>
<figureCitation id="27933F0A9D56B28C4A583C602EF09357" captionStart="Fig. 7" captionStartId="F7" captionText="Fig. 7. Male phallic complex of A-F. P. carli (Griffini, 1911) from Tam Dao (A-B), Cuc Phuong (C-D) and Ngo Luong (E-F); G-H. P. copia sp. nov.; I-J. P. cuspis sp. nov.; K-N. P. angusta sp. nov.; O-P. P. carnarius from Tam Tao; Q-R. P. pusilla sp. nov.; S-T. P. guadun sp. nov. in dorsal (A, C, F, G, I, L, M, O, Q, S) and in ventral (D, E, H, J, K, N, O, R, T) view, or in dorsal view with only dorsal lobes flipped proximad, ventral lobe distad (B). Scales 1 mm (all images)." figureDoi="10.3897/jor.28.32182.fig7" httpUri="https://binary.pensoft.net/fig/323904" pageId="5" pageNumber="112">7G-H</figureCitation>
).
</paragraph>
<paragraph lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112">
<paragraph id="E6A76C8E0527F4EACB6336ED90D99682" lastPageId="6" lastPageNumber="113" pageId="5" pageNumber="112">
Female. Subgenital plate acute-angled triangular in more than basal half, terminating into a long apical spine with regularly con
<pageBreakToken pageId="6" pageNumber="113" start="start">verging</pageBreakToken>
<pageBreakToken id="3C88044669EAC657CF82C282537B3BF9" pageId="6" pageNumber="113" start="start">verging</pageBreakToken>
margins to acute tip; at transition between basal and apical areas with a small elongate groove on both sides (Fig.
<figureCitation captionStart="Fig. 5" captionStartId="F5" captionText="Fig. 5. Female subgenital plate of A-C. P. carli (Griffini, 1911) from Hoang Lien (A-B) and Tam Dao (C, in oblique view); D-F. P. copia sp. nov. from Hoang Lien (D-E) and Copia (F); G. P. bavi sp. nov.; H. P. pusilla sp. nov.; I. P. guadun sp. nov.; J-K. P. carnarius Gorochov, 1998 from Tam Dao (J) and from BaVi (K, abdomen laterally compressed). The white arrows point at the small lateral grooves, constrictions or faint steps of the subgenital plates. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig5" httpUri="https://binary.pensoft.net/fig/323902" pageId="6" pageNumber="113">5D-F</figureCitation>
<figureCitation id="D5DAC35586500A47768D5886513261EA" captionStart="Fig. 5" captionStartId="F5" captionText="Fig. 5. Female subgenital plate of A-C. P. carli (Griffini, 1911) from Hoang Lien (A-B) and Tam Dao (C, in oblique view); D-F. P. copia sp. nov. from Hoang Lien (D-E) and Copia (F); G. P. bavi sp. nov.; H. P. pusilla sp. nov.; I. P. guadun sp. nov.; J-K. P. carnarius Gorochov, 1998 from Tam Dao (J) and from BaVi (K, abdomen laterally compressed). The white arrows point at the small lateral grooves, constrictions or faint steps of the subgenital plates. Scales 1 mm." figureDoi="10.3897/jor.28.32182.fig5" httpUri="https://binary.pensoft.net/fig/323902" pageId="6" pageNumber="113">5D-F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="113" type="coloration.">
<paragraph pageId="6" pageNumber="113">
<emphasis italics="true" pageId="6" pageNumber="113">Coloration.</emphasis>
<subSubSection id="A8A2A8E847968854CBE60379C103FD79" pageId="6" pageNumber="113" type="coloration.">
<paragraph id="9B3659394DB3A48A26D89DCD19038F84" pageId="6" pageNumber="113">
<emphasis id="B13229D176F90B72F50B1B4C3131D125" italics="true" pageId="6" pageNumber="113">Coloration.</emphasis>
-
</paragraph>
<paragraph pageId="6" pageNumber="113">Different shades of brown with light and dark brown mottling; head dark brown with light spots; pronotum and hind femora of similar color; hind femora towards apical area indistinctly lighter. Head: face blackish-brown with numerous small and few larger whitish-brown dots, also on scapus; clypeus, labrum and inner area of mandibles more uniformly medium or dark brown. Tegmen semi-transparent brown with indistinct dark spots, along anterior margin lighter; hind wings semi-transparent grey-brown, in anterior area clear, subtransparent.</paragraph>
<paragraph id="169B505F97B2F527585460D50D66D6E6" pageId="6" pageNumber="113">Different shades of brown with light and dark brown mottling; head dark brown with light spots; pronotum and hind femora of similar color; hind femora towards apical area indistinctly lighter. Head: face blackish-brown with numerous small and few larger whitish-brown dots, also on scapus; clypeus, labrum and inner area of mandibles more uniformly medium or dark brown. Tegmen semi-transparent brown with indistinct dark spots, along anterior margin lighter; hind wings semi-transparent grey-brown, in anterior area clear, subtransparent.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="113" type="description">
<paragraph pageId="6" pageNumber="113">
<emphasis italics="true" pageId="6" pageNumber="113">Measurements.</emphasis>
<subSubSection id="75A9CC954C78F417198801B7273C0EE1" pageId="6" pageNumber="113" type="description">
<paragraph id="5FEC6CF12C4F7D47520D35048B040B50" pageId="6" pageNumber="113">
<emphasis id="37B9BD677DBB7931D64F78899D400032" italics="true" pageId="6" pageNumber="113">Measurements.</emphasis>
-
</paragraph>
<paragraph pageId="6" pageNumber="113">(3 males, 4 females from Copia). In mm. Body w/wings: male 48-50, female 46-50; body w/o wings: male 27-31, female 28-35; pronotum: male 6.8-8.0, female 7.5-8.5; tegmen: male 36-38, female 32-37; tegmen width: male 15.5-16.5, female 14.0-15.5; hind femur: male 28.5-29.0, female 27-30; ovipositor: female 19.5-23.0. Index tegmen length: width male 2.24-2.45, female 2.21-2.40.</paragraph>
<paragraph id="F316C643F85B359FDB60A256E831381D" pageId="6" pageNumber="113">(3 males, 4 females from Copia). In mm. Body w/wings: male 48-50, female 46-50; body w/o wings: male 27-31, female 28-35; pronotum: male 6.8-8.0, female 7.5-8.5; tegmen: male 36-38, female 32-37; tegmen width: male 15.5-16.5, female 14.0-15.5; hind femur: male 28.5-29.0, female 27-30; ovipositor: female 19.5-23.0. Index tegmen length: width male 2.24-2.45, female 2.21-2.40.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="113" type="description">
<paragraph pageId="6" pageNumber="113">
<emphasis italics="true" pageId="6" pageNumber="113">Measurements.</emphasis>
<subSubSection id="458C40BE0ACA02138A4E94FFE36A8958" pageId="6" pageNumber="113" type="description">
<paragraph id="3AA64E2C92C6999B341B6605417801CE" pageId="6" pageNumber="113">
<emphasis id="0EF5CDA4F3DC3E2BA1458907F2159585" italics="true" pageId="6" pageNumber="113">Measurements.</emphasis>
-
</paragraph>
<paragraph pageId="6" pageNumber="113">(8 females from Hoang Lien). In mm. Body w/wings: female 39-45; body w/o wings: female 28-32; pronotum: female 7.8-8.2; tegmen: female 30-33; tegmen width: female 14; hind femur: female 26-28; ovipositor: female 18-20.</paragraph>
<paragraph id="21DF2397FF2012DA88557DB1B17B98A3" pageId="6" pageNumber="113">(8 females from Hoang Lien). In mm. Body w/wings: female 39-45; body w/o wings: female 28-32; pronotum: female 7.8-8.2; tegmen: female 30-33; tegmen width: female 14; hind femur: female 26-28; ovipositor: female 18-20.</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="113" type="etymology">
<paragraph pageId="6" pageNumber="113">
<emphasis italics="true" pageId="6" pageNumber="113">Etymology.</emphasis>
<subSubSection id="2EA5A5BF2CB7D38D8EE99183AFBF3321" pageId="6" pageNumber="113" type="etymology">
<paragraph id="974FE5433E298205591881F3954BC9CC" pageId="6" pageNumber="113">
<emphasis id="706EC0117EC7014C13492E76EBEE8D22" italics="true" pageId="6" pageNumber="113">Etymology.</emphasis>
-
</paragraph>
<paragraph pageId="6" pageNumber="113">The name of the new species refers to the type locality; noun in apposition.</paragraph>
<paragraph id="28F179BB7D9EE1F392BF156FBE0EFC2D" pageId="6" pageNumber="113">The name of the new species refers to the type locality; noun in apposition.</paragraph>
</subSubSection>
</treatment>
</document>

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Eggers, 1939: 9:
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: fig.2A.
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<paragraph pageId="0" pageNumber="27">Material examined.</paragraph>
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:
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, ALS(S)2600-5FI,
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(
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HGT)
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.
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<paragraph pageId="0" pageNumber="27">Distribution.</paragraph>
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<paragraph id="C0FA7CF41B81663FF63843992D18C17D" pageId="0" pageNumber="27">Distribution.</paragraph>
<paragraph id="08C7D82F53AC91083117034309B37630" pageId="0" pageNumber="27">
Myanmar, Nepal (
<bibRefCitation DOI="https://doi.org/10.1002/mmnd.200600014" author="Beaver, RA" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="27" pagination="155 - 178" refId="B3" refString="Beaver, RA, Gebhardt, H, 2006. A review of the oriental species of Scolytoplatypus Schaufuss (Coleoptera, Curculionidae, Scolytinae). Deutsche Entomologische Zeitschrift 53 (2): 155 - 178, DOI: https://doi.org/10.1002/mmnd.200600014" title="A review of the oriental species of Scolytoplatypus Schaufuss (Coleoptera, Curculionidae, Scolytinae)." url="https://doi.org/10.1002/mmnd.200600014" volume="53" year="2006">Beaver and Gebhardt 2006</bibRefCitation>
<bibRefCitation id="1BE35525B1E966CD6D36FB7E8BC4BC16" DOI="https://doi.org/10.1002/mmnd.200600014" author="Beaver, RA" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="27" pagination="155 - 178" refId="B3" refString="Beaver, RA, Gebhardt, H, 2006. A review of the oriental species of Scolytoplatypus Schaufuss (Coleoptera, Curculionidae, Scolytinae). Deutsche Entomologische Zeitschrift 53 (2): 155 - 178, DOI: https://doi.org/10.1002/mmnd.200600014" title="A review of the oriental species of Scolytoplatypus Schaufuss (Coleoptera, Curculionidae, Scolytinae)." url="https://doi.org/10.1002/mmnd.200600014" volume="53" year="2006">Beaver and Gebhardt 2006</bibRefCitation>
). New to China (Yunnan).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="diagnosis">
<paragraph pageId="0" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="27">
<subSubSection id="018075BE73E92EA5A6F5FB18498D1299" pageId="0" pageNumber="27" type="diagnosis">
<paragraph id="8683F0F50ED0743F796549A1312A7A8E" pageId="0" pageNumber="27">Diagnosis.</paragraph>
<paragraph id="6DECAEA5211360C54F0CC3161A94E0BF" pageId="0" pageNumber="27">
The species is most closely similar to
<taxonomicName class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" lsidName="S. wugongshanensis" order="Fagales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="wugongshanensis">
<emphasis italics="true" pageId="0" pageNumber="27">S. wugongshanensis</emphasis>
<taxonomicName id="BEA833EAC5AE18352513732EDC56B39E" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" lsidName="S. wugongshanensis" order="Fagales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="wugongshanensis">
<emphasis id="9D65BE74EC468361C1C2F270AD51B8FC" italics="true" pageId="0" pageNumber="27">S. wugongshanensis</emphasis>
</taxonomicName>
. but can be distinguished by the characters given in the key.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="host">
<paragraph pageId="0" pageNumber="27">Host.</paragraph>
<paragraph pageId="0" pageNumber="27">Not known.</paragraph>
<subSubSection id="AD727B397D153F05D5162766B8980672" pageId="0" pageNumber="27" type="host">
<paragraph id="7D64AE348CEEB876051B4ACC93C1EB80" pageId="0" pageNumber="27">Host.</paragraph>
<paragraph id="CBEB1F9F33128989DCE7DA871D60EDEC" pageId="0" pageNumber="27">Not known.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,199 +1,199 @@
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<treatment ID-GBIF-Taxon="152032687" LSID="urn:lsid:zoobank.org:act:1DCF15CA-E09E-4B16-9F13-FBDD84421895" httpUri="http://treatment.plazi.org/id/81FC1A0DDC828B644DE3781491CABB36" lastPageId="15" lastPageNumber="76" pageId="12" pageNumber="73">
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<paragraph pageId="12" pageNumber="73">
<taxonomicName LSID="urn:lsid:zoobank.org:act:1DCF15CA-E09E-4B16-9F13-FBDD84421895" authority="Konstantinov &amp; Konstantinova" class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica iviei" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="iviei">Ulrica iviei Konstantinov &amp; Konstantinova</taxonomicName>
<taxonomicNameLabel pageId="12" pageNumber="73">sp. n.</taxonomicNameLabel>
<treatment id="81FC1A0DDC828B644DE3781491CABB36" ID-GBIF-Taxon="152032687" LSID="urn:lsid:zoobank.org:act:1DCF15CA-E09E-4B16-9F13-FBDD84421895" httpUri="http://treatment.plazi.org/id/81FC1A0DDC828B644DE3781491CABB36" lastPageId="15" lastPageNumber="76" pageId="12" pageNumber="73" scope_family="Chrysomelidae" scope_order="Coleoptera" scope_subFamily="Galerucinae" scope_tribe="Alticini">
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<paragraph id="2B3CC179873DBF25FBA8066B7537078F" pageId="12" pageNumber="73">
<taxonomicName id="77C6A71A1313363EC41EB9F5912EE664" LSID="urn:lsid:zoobank.org:act:1DCF15CA-E09E-4B16-9F13-FBDD84421895" authority="Konstantinov &amp; Konstantinova" genus="Ulrica" lsidName="Ulrica iviei" pageId="12" pageNumber="73" rank="species" species="iviei">Ulrica iviei Konstantinov &amp; Konstantinova</taxonomicName>
<taxonomicNameLabel id="8230C1085694BD7C14733907D770269D" pageId="12" pageNumber="73">sp. n.</taxonomicNameLabel>
Figs 51-58
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="75" pageId="12" pageNumber="73" type="description">
<paragraph pageId="12" pageNumber="73">Description.</paragraph>
<paragraph lastPageId="14" lastPageNumber="75" pageId="12" pageNumber="73">
<subSubSection id="8F0081FE572337C47976E482F22DB281" lastPageId="14" lastPageNumber="75" pageId="12" pageNumber="73" type="description">
<paragraph id="56CC853F26767467E92972EB92F9935D" pageId="12" pageNumber="73">Description.</paragraph>
<paragraph id="95EA2E130F6FD8E4E95B913968133057" lastPageId="14" lastPageNumber="75" pageId="12" pageNumber="73">
Body length 1.89-2.05 mm, width 1.08-1.29 mm. Color chestnut brown to almost black with appendages lighter (Figs 51, 52). Head surface dorsally shiny, ventrally with some wrinkles (Fig. 53). Vertex with several large punctures, supraorbital pore as large as a few punctures on vertex near it. Supracallinal sulcus not separating antennal calli and vertex medially. Frontal ridge wide, about as long as antennal calli (Fig. 53). Anterofrontal ridge making long denticle about as long as
<pageBreakToken pageId="13" pageNumber="74" start="start">entire</pageBreakToken>
<pageBreakToken id="FCB694F8C0B845956C56692E01CD229E" pageId="13" pageNumber="74" start="start">entire</pageBreakToken>
clypeus. Pronotum and elytron with coarse punctures. Interspaces of elytron flat on disc, slightly convex apically. Proportions of tarsomeres of male as follows: protarsomeres 5:4:5:9; mesotarsomeres 5:4:5:9; metatarsomeres 10:5:5:9. Median lobe of aedeagus with more or less curved sides in ventral view, with ridge in middle being
<pageBreakToken pageId="14" pageNumber="75" start="start">wider</pageBreakToken>
<pageBreakToken id="9F57C167B90714035CF4FFFE86CA7D48" pageId="14" pageNumber="75" start="start">wider</pageBreakToken>
at base, narrowing towards middle and widening towards apex. In lateral view slightly curved without bump on ventral side beyond middle (Fig. 55). Spermatheca with pump at base wider than receptacle and duct making coils (Fig. 56). Sternite eight nearly fully sclerotized with tignum sharply bent anteriorly (Fig. 58). Vaginal palpi merged at about apical one third (Fig. 57).
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="75" type="etymology">
<paragraph pageId="14" pageNumber="75">Etymology.</paragraph>
<paragraph pageId="14" pageNumber="75">The specific epithet is a patronym dedicated to Mike Ivie who collected the holotype.</paragraph>
<subSubSection id="0F6A20B088928F8A9F2CE0992FEA0605" pageId="14" pageNumber="75" type="etymology">
<paragraph id="84ABA6E29CBB25588A111E7FE0F53B9F" pageId="14" pageNumber="75">Etymology.</paragraph>
<paragraph id="B98F46D68AEEC8B82046FE2F72EC4756" pageId="14" pageNumber="75">The specific epithet is a patronym dedicated to Mike Ivie who collected the holotype.</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="75" type="diagnosis and comparison">
<paragraph pageId="14" pageNumber="75">Diagnosis and comparison.</paragraph>
<paragraph pageId="14" pageNumber="75">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica iviei" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="iviei">Ulrica iviei</taxonomicName>
<subSubSection id="60CEAE3C0FB248B68BE6D023851A360E" pageId="14" pageNumber="75" type="diagnosis and comparison">
<paragraph id="58E572CB36F49418A2DBCB68E9D3164C" pageId="14" pageNumber="75">Diagnosis and comparison.</paragraph>
<paragraph id="028E1D7B627828A678B5F4C203EAD80E" pageId="14" pageNumber="75">
<taxonomicName id="5F28C0D823922AC842E45F6CA9D50B71" genus="Ulrica" lsidName="Ulrica iviei" pageId="14" pageNumber="75" rank="species" species="iviei">Ulrica iviei</taxonomicName>
can be easily differentiated from
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica eltoro" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
<taxonomicName id="DC1107681D01E27D14775463638EDB32" genus="Ulrica" lsidName="Ulrica eltoro" pageId="14" pageNumber="75" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
based on the key below.
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="76" pageId="14" pageNumber="75" type="type material">
<paragraph pageId="14" pageNumber="75">Type material.</paragraph>
<paragraph pageId="14" pageNumber="75">
<subSubSection id="EFFFE72762099A9E36432D4D1821DFFB" lastPageId="15" lastPageNumber="76" pageId="14" pageNumber="75" type="type material">
<paragraph id="EF3C671007BA1F7B2085EFF07FF0A987" pageId="14" pageNumber="75">Type material.</paragraph>
<paragraph id="BBE523C2DD7259C95AF4F73EF0AB1AA2" pageId="14" pageNumber="75">
Holotype: ♂, Puerto Rico: Caribbean Nat. For. Pico El Yunque, El Toro trail, 975 m, 23 Sept. 1987, leg. M. A. Ivie, dwarf forest litter (WIBF). Paratypes ♂ and ♀, Puerto Rico El Yunque, Mt. Britton Tr. VIII.11.1999, C. W.
<normalizedToken originalValue="OBrien">O'Brien</normalizedToken>
<normalizedToken id="D81E26B8BAD069F3EB59BD453120B344" originalValue="OBrien">O'Brien</normalizedToken>
, P. Kovarik (MLBU, USNM).
</paragraph>
<subSection lastPageId="15" lastPageNumber="76" pageId="14" pageNumber="75" type="key to species of ulrica from puerto rico">
<paragraph pageId="14" pageNumber="75">
<subSection id="37198036818DC2E4C84636F55801F4EB" lastPageId="15" lastPageNumber="76" pageId="14" pageNumber="75" type="key to species of ulrica from puerto rico">
<paragraph id="9814AF87474DAF682231E6D279F9317F" pageId="14" pageNumber="75">
Key to species of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="genus">Ulrica</taxonomicName>
<taxonomicName id="0E4904D2A2F2AD18AAD3A259C80A0367" genus="Ulrica" lsidName="Ulrica" pageId="14" pageNumber="75" rank="genus">Ulrica</taxonomicName>
from Puerto Rico
</paragraph>
<paragraph pageId="14" pageNumber="75">
<table pageId="14" pageNumber="75">
<tr pageId="14" pageNumber="75">
<td colspan="1" pageId="14" pageNumber="75" rowspan="1">Fig. 47</td>
<td colspan="1" pageId="14" pageNumber="75" rowspan="1">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica eltoro" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
<paragraph id="536A362611097122589E4A3265E7A0E4" pageId="14" pageNumber="75">
<table id="12677F8AADF113AE01A7341730F2308B" pageId="14" pageNumber="75">
<tr id="0476832E4B33EE8D64435BF78C8623C4" pageId="14" pageNumber="75">
<td id="2BD413A27DC41CDACAAD4E4C4596D5BF" colspan="1" pageId="14" pageNumber="75" rowspan="1">Fig. 47</td>
<td id="26491F2B1406FA0AA8DEC14B6EC83BBA" colspan="1" pageId="14" pageNumber="75" rowspan="1">
<taxonomicName id="99277DBE3F24CB448825EF5612EDE69B" genus="Ulrica" lsidName="Ulrica eltoro" pageId="14" pageNumber="75" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
</td>
</tr>
<tr pageId="14" pageNumber="75">
<td colspan="1" pageId="14" pageNumber="75" rowspan="1">Fig. 51</td>
<td colspan="1" pageId="14" pageNumber="75" rowspan="1">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica iviei" order="Coleoptera" pageId="14" pageNumber="75" phylum="Arthropoda" rank="species" species="iviei">Ulrica iviei</taxonomicName>
<tr id="94EF190FE96CCDFD620B71A67436AEAB" pageId="14" pageNumber="75">
<td id="DAF91C23EB3C3CD953CF024372A2C780" colspan="1" pageId="14" pageNumber="75" rowspan="1">Fig. 51</td>
<td id="20196AF9982389DD7964F4989D14DFBE" colspan="1" pageId="14" pageNumber="75" rowspan="1">
<taxonomicName id="71268F3D193BCAF8B4D931950227E9A0" genus="Ulrica" lsidName="Ulrica iviei" pageId="14" pageNumber="75" rank="species" species="iviei">Ulrica iviei</taxonomicName>
</td>
</tr>
</table>
</paragraph>
<caption pageId="15" pageNumber="76">
<paragraph pageId="15" pageNumber="76">
<pageBreakToken pageId="15" pageNumber="76" start="start">Figures</pageBreakToken>
<caption id="1E89871C49AF95DB6736C46715B56964" pageId="15" pageNumber="76">
<paragraph id="D3AF23220A7BC8295FD644C440076681" pageId="15" pageNumber="76">
<pageBreakToken id="BF7AE0334262F8FA8727ABCBEBC95CB1" pageId="15" pageNumber="76" start="start">Figures</pageBreakToken>
51-55.
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica iviei" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="species" species="iviei">Ulrica iviei</taxonomicName>
<taxonomicName id="F2A16A90FF0C90B209EB607527A8C9D8" genus="Ulrica" lsidName="Ulrica iviei" pageId="15" pageNumber="76" rank="species" species="iviei">Ulrica iviei</taxonomicName>
: 51 habitus, dorsal view 52 habitus, lateral view 53 head, frontalview 54 abdominal sternites, female 55 median lobe of aedeagus, ventral, lateral and dorsal views.
</paragraph>
</caption>
<caption pageId="15" pageNumber="76">
<paragraph pageId="15" pageNumber="76">
<caption id="590736FB6C69C42B09E388A330A435B3" pageId="15" pageNumber="76">
<paragraph id="F303DEE5973BF03418F5EBB37A1BD063" pageId="15" pageNumber="76">
Figures 56-58.
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica iviei" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="species" species="iviei">Ulrica iviei</taxonomicName>
<taxonomicName id="339C2CB42FD7126603C5F3B6088604CB" genus="Ulrica" lsidName="Ulrica iviei" pageId="15" pageNumber="76" rank="species" species="iviei">Ulrica iviei</taxonomicName>
: 56 spermatheca 57 vaginal palpi 58 tignum.
</paragraph>
</caption>
<caption pageId="15" pageNumber="76">
<paragraph pageId="15" pageNumber="76">Figures 59-60. El Yunque, Puerto Rico: 59 moss on tree trunks 60 forest along El Toro trail where moss occurs.</paragraph>
<caption id="A8BC051BC2205C45B0B99A2C33E79BC1" pageId="15" pageNumber="76">
<paragraph id="52579A8F17AB1DC318814D6626EC4D0F" pageId="15" pageNumber="76">Figures 59-60. El Yunque, Puerto Rico: 59 moss on tree trunks 60 forest along El Toro trail where moss occurs.</paragraph>
</caption>
</subSection>
<subSection pageId="15" pageNumber="76" type="key to monoplatini genera of the west indies">
<paragraph pageId="15" pageNumber="76">Key to Monoplatini genera of the West Indies</paragraph>
<paragraph pageId="15" pageNumber="76">
<subSection id="4D7F97C0B92484089D061B68AFCCDB50" pageId="15" pageNumber="76" type="key to monoplatini genera of the west indies">
<paragraph id="B8792E35C3DBE99743D0AF7037D0179B" pageId="15" pageNumber="76">Key to Monoplatini genera of the West Indies</paragraph>
<paragraph id="F4B735F1888EB4C798E5EF04F6956A03" pageId="15" pageNumber="76">
In addition to
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Distigmoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Distigmoptera" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Distigmoptera</taxonomicName>
<taxonomicName id="ABCFCB37A861018B3F1643A1B5FDBBEF" class="Insecta" family="Chrysomelidae" genus="Distigmoptera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Distigmoptera" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Distigmoptera</taxonomicName>
and
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Ulrica</taxonomicName>
<taxonomicName id="3C6A1754A50C939172C2DDEFBBDCE13E" genus="Ulrica" lsidName="Ulrica" pageId="15" pageNumber="76" rank="genus">Ulrica</taxonomicName>
, six other Monoplatini genera are reported in the West Indies:
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Aedmon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aedmon" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Aedmon</taxonomicName>
<taxonomicName id="7419318D1941CDB6040F7F95AE52DD48" genus="Aedmon" lsidName="Aedmon" pageId="15" pageNumber="76" rank="genus">Aedmon</taxonomicName>
Clark,
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Apleuraltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Apleuraltica" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Apleuraltica</taxonomicName>
<taxonomicName id="BB2C45120E3EABCE8D95989E87D9A0AE" genus="Apleuraltica" lsidName="Apleuraltica" pageId="15" pageNumber="76" rank="genus">Apleuraltica</taxonomicName>
,
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bonfilsus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bonfilsus" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Bonfilsus</taxonomicName>
<taxonomicName id="177569102398FBBBEC79C56CF00F1CFD" genus="Bonfilsus" lsidName="Bonfilsus" pageId="15" pageNumber="76" rank="genus">Bonfilsus</taxonomicName>
Scherer,
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Homotyphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Homotyphus" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Homotyphus</taxonomicName>
<taxonomicName id="AB03D7D07E3C52DE3F09FA4B8E28CB28" genus="Homotyphus" lsidName="Homotyphus" pageId="15" pageNumber="76" rank="genus">Homotyphus</taxonomicName>
Clark,
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Hypolampsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hypolampsis" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Hypolampsis</taxonomicName>
<taxonomicName id="93CC71F0B39F7CDCEEBCE9882A1E01D5" genus="Hypolampsis" lsidName="Hypolampsis" pageId="15" pageNumber="76" rank="genus">Hypolampsis</taxonomicName>
Clark, and
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Physimerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Physimerus" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Physimerus</taxonomicName>
<taxonomicName id="5152BAAA262660B18AAF712DB193B87D" genus="Physimerus" lsidName="Physimerus" pageId="15" pageNumber="76" rank="genus">Physimerus</taxonomicName>
Clark (
<bibRefCitation author="Takizawa, H" journalOrPublisher="Nueva Serie, Num" pageId="15" pageNumber="76" pagination="1 - 125" title="Check list of Chrysomelidae in the West Indies (Coleoptera). Museo Nacional de Historia Natural, Santo Domingo." volume="2" year="2003">Takizawa 2003</bibRefCitation>
<bibRefCitation id="1537BA3387BD991C77951D4475609462" author="Takizawa, H" journalOrPublisher="Nueva Serie, Num" pageId="15" pageNumber="76" pagination="1 - 125" title="Check list of Chrysomelidae in the West Indies (Coleoptera). Museo Nacional de Historia Natural, Santo Domingo." volume="2" year="2003">Takizawa 2003</bibRefCitation>
).
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Aedmon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aedmon" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Aedmon</taxonomicName>
<taxonomicName id="A3C5732E77A93D5FBA1C0BB334E15C57" genus="Aedmon" lsidName="Aedmon" pageId="15" pageNumber="76" rank="genus">Aedmon</taxonomicName>
,
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Apleuraltica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Apleuraltica" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Apleuraltica</taxonomicName>
<taxonomicName id="A96ED82D7089975260C2BEAFCBF1305C" genus="Apleuraltica" lsidName="Apleuraltica" pageId="15" pageNumber="76" rank="genus">Apleuraltica</taxonomicName>
, and
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bonfilsus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bonfilsus" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Bonfilsus</taxonomicName>
<taxonomicName id="AA3A6E537B8CC3C8521FAABC98992212" genus="Bonfilsus" lsidName="Bonfilsus" pageId="15" pageNumber="76" rank="genus">Bonfilsus</taxonomicName>
are West Indian endemics, relatively small (with the exception of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Aedmon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aedmon" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Aedmon</taxonomicName>
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) and relatively well circumscribed. However,
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Homotyphus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Homotyphus" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Homotyphus</taxonomicName>
<taxonomicName id="1DA9527735387926E4E0D8D525D1BD99" genus="Homotyphus" lsidName="Homotyphus" pageId="15" pageNumber="76" rank="genus">Homotyphus</taxonomicName>
(with about 20 species mostly from South America and just one in the West Indies),
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Hypolampsis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hypolampsis" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Hypolampsis</taxonomicName>
<taxonomicName id="48A8F9F4B8D152D56F01C3B6306AEE52" genus="Hypolampsis" lsidName="Hypolampsis" pageId="15" pageNumber="76" rank="genus">Hypolampsis</taxonomicName>
(with about 50 species from North, Central, and South America and just two in the West Indies), and
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Physimerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Physimerus" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Physimerus</taxonomicName>
<taxonomicName id="C411D655934599AD4114C66CA5F7713B" genus="Physimerus" lsidName="Physimerus" pageId="15" pageNumber="76" rank="genus">Physimerus</taxonomicName>
(with about 60 species from Central and South America and one in the West Indies) are poorly understood; their identity, distinguishing characters and composition need extensive review. Below we provide a key for Monoplatini genera of the West Indies based on the West Indian species.
</paragraph>
<paragraph pageId="15" pageNumber="76">
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<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Ulrica</taxonomicName>
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<taxonomicName id="9E9767B1C87FE56DB7BF65C23AF392C6" genus="Ulrica" lsidName="Ulrica" pageId="15" pageNumber="76" rank="genus">Ulrica</taxonomicName>
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<taxonomicName class="Insecta" family="Chrysomelidae" genus="Bonfilsus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bonfilsus" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Bonfilsus</taxonomicName>
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<taxonomicName class="Insecta" family="Chrysomelidae" genus="Physimerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Physimerus" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Physimerus</taxonomicName>
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<taxonomicName class="Insecta" family="Chrysomelidae" genus="Aedmon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aedmon" order="Coleoptera" pageId="15" pageNumber="76" phylum="Arthropoda" rank="genus">Aedmon</taxonomicName>
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@ -1,126 +1,125 @@
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<mods:url id="BD38CADB7196D4BA37E2D43004760BFA">http://dx.doi.org/10.3897/zookeys.359.6057</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier id="D9196975DA6F129842BF6D479183F039" type="DOI">http://dx.doi.org/10.3897/zookeys.359.6057</mods:identifier>
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<treatment ID-GBIF-Taxon="152049967" LSID="urn:lsid:zoobank.org:act:0B55E091-1323-469F-A479-57E7F47388D4" httpUri="http://treatment.plazi.org/id/81FFB0E7BBADD3AD8D0EE196A0893A28" lastPageId="11" lastPageNumber="24" pageId="10" pageNumber="23">
<subSubSection pageId="10" pageNumber="23" type="nomenclature">
<paragraph pageId="10" pageNumber="23">
<taxonomicName LSID="http://zoobank.org/0B55E091-1323-469F-A479-57E7F47388D4" authority="Wang &amp; Ren" class="Insecta" family="Coccinellidae" genus="Microserangium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microserangium semilunatum" order="Coleoptera" pageId="10" pageNumber="23" phylum="Arthropoda" rank="species" species="semilunatum">Microserangium semilunatum Wang &amp; Ren</taxonomicName>
<taxonomicNameLabel pageId="10" pageNumber="23">sp. n.</taxonomicNameLabel>
<treatment id="81FFB0E7BBADD3AD8D0EE196A0893A28" ID-GBIF-Taxon="152049967" LSID="urn:lsid:zoobank.org:act:0B55E091-1323-469F-A479-57E7F47388D4" httpUri="http://treatment.plazi.org/id/81FFB0E7BBADD3AD8D0EE196A0893A28" lastPageId="11" lastPageNumber="24" pageId="10" pageNumber="23" scope_family="Coccinellidae" scope_order="Coleoptera">
<subSubSection id="7BBE2CC0F3C7F47F7B2FFBD99B24428E" pageId="10" pageNumber="23" type="nomenclature">
<paragraph id="C68F51C0EE9F5F845C3617180BA53A0A" pageId="10" pageNumber="23">
<taxonomicName id="E2208D7A561017DD614D2645CA7AE08F" LSID="http://zoobank.org/0B55E091-1323-469F-A479-57E7F47388D4" authority="Wang &amp; Ren" genus="Microserangium" lsidName="Microserangium semilunatum" pageId="10" pageNumber="23" rank="species" species="semilunatum">Microserangium semilunatum Wang &amp; Ren</taxonomicName>
<taxonomicNameLabel id="ADCDAD1C1E304965A125E653F4DC3D2F" pageId="10" pageNumber="23">sp. n.</taxonomicNameLabel>
Figures 16, 53-60, 93
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="24" pageId="10" pageNumber="23" type="diagnosis">
<paragraph pageId="10" pageNumber="23">Diagnosis.</paragraph>
<paragraph lastPageId="11" lastPageNumber="24" pageId="10" pageNumber="23">
<subSubSection id="5F4D1486D7439E5B87EC83815C4BAB95" lastPageId="11" lastPageNumber="24" pageId="10" pageNumber="23" type="diagnosis">
<paragraph id="0975219BC660B5B03E4DD5AD39DA8684" pageId="10" pageNumber="23">Diagnosis.</paragraph>
<paragraph id="5AA2FF4B289C8B619EC6832799492886" lastPageId="11" lastPageNumber="24" pageId="10" pageNumber="23">
The male genitalia of this species are similar to
<taxonomicName class="Insecta" family="Coccinellidae" genus="Microserangium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microserangium erythrinum" order="Coleoptera" pageId="11" pageNumber="24" phylum="Arthropoda" rank="species" species="erythrinum">
<pageBreakToken pageId="11" pageNumber="24" start="start">Microserangium</pageBreakToken>
<taxonomicName id="B10F66AD93C41059198C873BC7DE565E" genus="Microserangium" lsidName="Microserangium erythrinum" pageId="11" pageNumber="24" rank="species" species="erythrinum">
<pageBreakToken id="19166A82765B3B27FC867EFFBFBB5BAD" pageId="11" pageNumber="24" start="start">Microserangium</pageBreakToken>
erythrinum
</taxonomicName>
, but this species can be distinguished from the latter by its uniformly dark elytra and semilunate penis guide (Figs 16, 56-57). In
<taxonomicName class="Insecta" family="Coccinellidae" genus="Microserangium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microserangium erythrinum" order="Coleoptera" pageId="11" pageNumber="24" phylum="Arthropoda" rank="species" species="erythrinum">Microserangium erythrinum</taxonomicName>
<taxonomicName id="9C22648ABF7A03D8EE7DFE724599E12D" genus="Microserangium" lsidName="Microserangium erythrinum" pageId="11" pageNumber="24" rank="species" species="erythrinum">Microserangium erythrinum</taxonomicName>
,theelytral disk is yellow or burgundy with dark brown margins and the penis guide is acutely triangular (Figs 12, 24-25).
</paragraph>
<caption pageId="11" pageNumber="24">
<paragraph pageId="11" pageNumber="24">
<caption id="755F21AA93B637D7D27DAAEC3E050986" pageId="11" pageNumber="24">
<paragraph id="010CA9EB33EA52E8886C17E7FF08A1A9" pageId="11" pageNumber="24">
Figures 53-68. 53-60
<taxonomicName class="Insecta" family="Coccinellidae" genus="Microserangium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microserangium semilunatum" order="Coleoptera" pageId="11" pageNumber="24" phylum="Arthropoda" rank="species" species="semilunatum">Microserangium semilunatum</taxonomicName>
<taxonomicName id="7B92414A7B01CA393FD5A318A34871D3" genus="Microserangium" lsidName="Microserangium semilunatum" pageId="11" pageNumber="24" rank="species" species="semilunatum">Microserangium semilunatum</taxonomicName>
Wang &amp; Ren, sp. n. 53 abdomen 54-57 male genitalia: 54 penis 55 apex of penis 56 tegmen, lateral view 57 tegmen, ventral view 58 mandible 59 antenna 60 hind leg 61-68
<taxonomicName class="Insecta" family="Coccinellidae" genus="Microserangium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Microserangium hainanensis" order="Coleoptera" pageId="11" pageNumber="24" phylum="Arthropoda" rank="species" species="hainanensis">Microserangium hainanensis</taxonomicName>
<taxonomicName id="E5487E5E97AF768051E2CCE2A39DFC0D" genus="Microserangium" lsidName="Microserangium hainanensis" pageId="11" pageNumber="24" rank="species" species="hainanensis">Microserangium hainanensis</taxonomicName>
Miyatake, 1961 61 abdomen 62-65 male genitalia: 62 penis 63 apex of penis 64 tegmen, lateral view 65 tegmen, ventral view 66 mandible 67 antenna 68 hind leg. Scale bars: 0.1mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="11" pageNumber="24" type="description">
<paragraph pageId="11" pageNumber="24">Description.</paragraph>
<paragraph pageId="11" pageNumber="24">TL: 1.42-1.55mm, TW: 1.24-1.35mm, TH: 0.76-0.86mm, TL/TW: 1.14-1.15; PL/PW: 0.40-0.44; EL/EW: 0.93-0.98; HW/TW: 0.44; PW/TW: 0.73.</paragraph>
<paragraph pageId="11" pageNumber="24">Body shiny and glabrous (Fig. 16). Dorsum uniformly black. Head black, clypeus brown. Underside black, legs dark brown, tibiae and tarsi yellow.</paragraph>
<paragraph pageId="11" pageNumber="24">
<subSubSection id="9FE993CEE71B0A31D794CA123C143557" pageId="11" pageNumber="24" type="description">
<paragraph id="9AC129A31583C0CC71294BFC616C0652" pageId="11" pageNumber="24">Description.</paragraph>
<paragraph id="228D6363E9D2B436F3AFBBE34C0AA08F" pageId="11" pageNumber="24">TL: 1.42-1.55mm, TW: 1.24-1.35mm, TH: 0.76-0.86mm, TL/TW: 1.14-1.15; PL/PW: 0.40-0.44; EL/EW: 0.93-0.98; HW/TW: 0.44; PW/TW: 0.73.</paragraph>
<paragraph id="AA12F0086126852DF1DD66B8DD53F567" pageId="11" pageNumber="24">Body shiny and glabrous (Fig. 16). Dorsum uniformly black. Head black, clypeus brown. Underside black, legs dark brown, tibiae and tarsi yellow.</paragraph>
<paragraph id="206040CAFF5343CD550B562D648DDB6C" pageId="11" pageNumber="24">
Head transverse and ventrally flattened; frontal punctures large and densely distributed, 0.3-1.0 diameter apart, associated with long sparsely distributed setae; eyes round, large and coarsely faceted, widest interocular distance 0.50
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="F764E29D6667E80815ED780773DFD3F6" originalValue="×">x</normalizedToken>
width of head. Antennal club oval and flat, apex blunt (Fig. 59).
</paragraph>
<paragraph pageId="11" pageNumber="24">Pronotum short and strongly transverse, anterior corner inconspicuous and blunt, mat and shagreened. Pronotal disk with densely distributed large and fine shallow punctures both associated with long sparsely distributed setae. Elytra smooth and shiny, with sparse row of long setae along margins, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat weakly furrowed. Metaventrite shiny and glabrous, punctures fine and sparsely distributed, 2.0-3.0 diameters apart. Meso- and metatibiae strongly angulate externally beyond middle; tarsus with 3 tarsomeres (Fig. 60).</paragraph>
<paragraph pageId="11" pageNumber="24">Male genitalia. Penis strongly curved along entire length, apex narrowed and acicular, penis capsule with large outer arm and inconspicuous inner one (Figs 54-55). Tegmen rather slender and strongly asymmetrical. Penis guide in lateral view small, almost straight, apex pointed (Fig. 56), in ventral view small and semilunate (Fig. 57). Parameres conspicuous.</paragraph>
<paragraph id="A9D643BD794DC52C7B72D41D8823AF6D" pageId="11" pageNumber="24">Pronotum short and strongly transverse, anterior corner inconspicuous and blunt, mat and shagreened. Pronotal disk with densely distributed large and fine shallow punctures both associated with long sparsely distributed setae. Elytra smooth and shiny, with sparse row of long setae along margins, punctures extremely fine and inconspicuous. Prosternum mat and impunctate. Mesoventrite transverse, very short, surface mat weakly furrowed. Metaventrite shiny and glabrous, punctures fine and sparsely distributed, 2.0-3.0 diameters apart. Meso- and metatibiae strongly angulate externally beyond middle; tarsus with 3 tarsomeres (Fig. 60).</paragraph>
<paragraph id="D0C93F3C1E92D303908DD54558697E0E" pageId="11" pageNumber="24">Male genitalia. Penis strongly curved along entire length, apex narrowed and acicular, penis capsule with large outer arm and inconspicuous inner one (Figs 54-55). Tegmen rather slender and strongly asymmetrical. Penis guide in lateral view small, almost straight, apex pointed (Fig. 56), in ventral view small and semilunate (Fig. 57). Parameres conspicuous.</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="24" type="types">
<paragraph pageId="11" pageNumber="24">Types.</paragraph>
<paragraph pageId="11" pageNumber="24">
<subSubSection id="82F0AB34A684E27614C0C851D0CC02F3" pageId="11" pageNumber="24" type="types">
<paragraph id="707A253A46B33EE6C92286474C7D0A7E" pageId="11" pageNumber="24">Types.</paragraph>
<paragraph id="A1D8DC783720D5DA748A6132B3870328" pageId="11" pageNumber="24">
Holotype ♂: China, Hainan: Bawangling,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="19.094166">19°05.65'N</geoCoordinate>
<geoCoordinate id="21E4648BBAF61C487BEC6A44B74C8B50" direction="north" orientation="latitude" precision="9" value="19.094166">19°05.65'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="109.11217">109°6.73'E</geoCoordinate>
<geoCoordinate id="EB87A879DCA7DC2F092D475D1950AF19" direction="east" orientation="longitude" precision="9" value="109.11217">109°6.73'E</geoCoordinate>
, ca 330m, 21.iii.1996, Peng ZQ leg. Paratypes (12): Hainan: 4♂♂1♀, same data to holotype; 1♂1♀, Bawangling,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="19.094166">19°05.65'N</geoCoordinate>
<geoCoordinate id="D2C076FE6ECE2102447C13FD16DFE23A" direction="north" orientation="latitude" precision="9" value="19.094166">19°05.65'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="109.11217">109°6.73'E</geoCoordinate>
<geoCoordinate id="0172D3B5A31618387473D6AF0275BD1D" direction="east" orientation="longitude" precision="9" value="109.11217">109°6.73'E</geoCoordinate>
, ca 330m, 3.ix.1998, Peng ZQleg.; 1♂, Bawangling,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="19.094166">19°05.65'N</geoCoordinate>
<geoCoordinate id="6171C17E9BB74499168DA5511637CB3F" direction="north" orientation="latitude" precision="9" value="19.094166">19°05.65'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="109.11217">109°6.73'E</geoCoordinate>
<geoCoordinate id="4B7312EBFE90FA6944C8ABC22F6F1843" direction="east" orientation="longitude" precision="9" value="109.11217">109°6.73'E</geoCoordinate>
, ca 330m, 20.iv.2000, Peng ZQleg.; 1♂1♀, Diaoluoshan,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="18.935833">18°56.15'N</geoCoordinate>
<geoCoordinate id="00A46677CA3A709D087D84EC96756091" direction="north" orientation="latitude" precision="9" value="18.935833">18°56.15'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="109.94833">109°56.90'E</geoCoordinate>
<geoCoordinate id="03F8EB9FD210D42B4F6D231FD80EF292" direction="east" orientation="longitude" precision="9" value="109.94833">109°56.90'E</geoCoordinate>
, ca 200m, 26.vii.2006, Wang XM leg.; 1♂, Diaoluoshan,
<geoCoordinate direction="north" orientation="latitude" precision="9" value="18.935833">18°56.15'N</geoCoordinate>
<geoCoordinate id="A834CDBD86240A6B77F8C4BFF5428D76" direction="north" orientation="latitude" precision="9" value="18.935833">18°56.15'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="9" value="109.94833">109°56.90'E</geoCoordinate>
<geoCoordinate id="67A061C9EC2E0F8FA996B2456E9ECA9D" direction="east" orientation="longitude" precision="9" value="109.94833">109°56.90'E</geoCoordinate>
, ca 200m, 7.v.2005, Wang XM leg.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="24" type="distribution">
<paragraph pageId="11" pageNumber="24">Distribution.</paragraph>
<paragraph pageId="11" pageNumber="24">China (Hainan).</paragraph>
<subSubSection id="8A991BDFB96455EEB4AF18EB03FD7250" pageId="11" pageNumber="24" type="distribution">
<paragraph id="DFDFE21140AF1A44A9E46946622F4652" pageId="11" pageNumber="24">Distribution.</paragraph>
<paragraph id="D9DC9E11634EC450EB7422901D90F613" pageId="11" pageNumber="24">China (Hainan).</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="24" type="etymology">
<paragraph pageId="11" pageNumber="24">Etymology.</paragraph>
<paragraph pageId="11" pageNumber="24">The specific epithet is formed from the the Latin adjective semilunatus, referring to the semilunate penis guide.</paragraph>
<subSubSection id="CD51648DD7F961FD0F7EED808D82FBDB" pageId="11" pageNumber="24" type="etymology">
<paragraph id="CDEC7D94ADD5A1C95A4A2F0B0645461D" pageId="11" pageNumber="24">Etymology.</paragraph>
<paragraph id="07CB709F942CA490828B1EB23D092E20" pageId="11" pageNumber="24">The specific epithet is formed from the the Latin adjective semilunatus, referring to the semilunate penis guide.</paragraph>
</subSubSection>
</treatment>
</document>