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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.930.47742" ID-GBIF-Dataset="9223ceda-3eb6-4d4d-91a1-a08d3dc443d7" ID-PMC="PMC7200893" ID-Pensoft-Pub="1313-2970-930-37" ID-Pensoft-UUID="255C174A2BB75F5B8259DCBB698296D2" ID-PubMed="32390747" ID-ZooBank="FABF3E569A1544CD9A5072DCF25AF87D" ModsDocID="1313-2970-930-37" checkinTime="1588165340049" checkinUser="pensoft" docAuthor="Semenyuk, Irina, Golovatch, Sergei I. & Wesener, Thomas" docDate="2020" docId="4FCE290506CA53B5B8B78EFAD5029C4F" docLanguage="en" docName="ZooKeys 930: 37-60" docOrigin="ZooKeys 930" docSource="http://dx.doi.org/10.3897/zookeys.930.47742" docTitle="Sphaeropoeus honbaensis Semenyuk & Golovatch & Wesener 2020, sp. nov." docType="treatment" docUuid="8C6B0566-6BEE-4A06-B59F-BDA14A90892B" docUuidSource="ZooBank" docVersion="5" id="255C174A2BB75F5B8259DCBB698296D2" lastPageNumber="37" masterDocId="255C174A2BB75F5B8259DCBB698296D2" masterDocTitle="Some new or poorly-known Zephroniidae (Diplopoda, Sphaerotheriida) from Vietnam" masterLastPageNumber="60" masterPageNumber="37" pageNumber="37" updateTime="1668168797197" updateUser="ExternalLinkService">
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<mods:title>Some new or poorly-known Zephroniidae (Diplopoda, Sphaerotheriida) from Vietnam</mods:title>
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<document ID-CLB-Dataset="22483" ID-DOI="http://dx.doi.org/10.3897/zookeys.930.47742" ID-GBIF-Dataset="9223ceda-3eb6-4d4d-91a1-a08d3dc443d7" ID-PMC="PMC7200893" ID-Pensoft-Pub="1313-2970-930-37" ID-Pensoft-UUID="255C174A2BB75F5B8259DCBB698296D2" ID-PubMed="32390747" ID-ZooBank="FABF3E569A1544CD9A5072DCF25AF87D" ModsDocID="1313-2970-930-37" checkinTime="1588165340049" checkinUser="pensoft" docAuthor="Semenyuk, Irina, Golovatch, Sergei I. & Wesener, Thomas" docDate="2020" docId="4FCE290506CA53B5B8B78EFAD5029C4F" docLanguage="en" docName="ZooKeys 930: 37-60" docOrigin="ZooKeys 930" docSource="http://dx.doi.org/10.3897/zookeys.930.47742" docTitle="Sphaeropoeus honbaensis Semenyuk & Golovatch & Wesener 2020, sp. nov." docType="treatment" docUuid="8C6B0566-6BEE-4A06-B59F-BDA14A90892B" docUuidSource="ZooBank" docVersion="7" id="255C174A2BB75F5B8259DCBB698296D2" lastPageNumber="37" masterDocId="255C174A2BB75F5B8259DCBB698296D2" masterDocTitle="Some new or poorly-known Zephroniidae (Diplopoda, Sphaerotheriida) from Vietnam" masterLastPageNumber="60" masterPageNumber="37" pageNumber="37" updateTime="1732754218732" updateUser="ExternalLinkService">
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<mods:title id="86690E51612ACCB879C0AC4F05CF9139">Some new or poorly-known Zephroniidae (Diplopoda, Sphaerotheriida) from Vietnam</mods:title>
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<mods:namePart>Semenyuk, Irina</mods:namePart>
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<mods:affiliation>A. N. Severtsov Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia & Joint Russian-Vietnamese Tropical Center, Street 3 Thang 2, 3, Q 10, Ho Chi Minh City, Vietnam</mods:affiliation>
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<mods:namePart id="D86AE7C407EE45BBD897C43001F0716D">Semenyuk, Irina</mods:namePart>
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<mods:affiliation id="39E872D078C46AB5C2CF2CA8D750ECF8">A. N. Severtsov Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia & Joint Russian-Vietnamese Tropical Center, Street 3 Thang 2, 3, Q 10, Ho Chi Minh City, Vietnam</mods:affiliation>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation>A. N. Severtsov Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:namePart id="D7EED413945AF75FFF1EF1450C9D7BB7">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="417FC382A648039578C9D05BEE57F2F2">A. N. Severtsov Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:affiliation>Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">twesener@uni-bonn.de</mods:nameIdentifier>
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<mods:namePart id="A1009FBF0CBC7255E173433B1A6D9BE7">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="69FFD325C157833A13722E36416820A2">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany</mods:affiliation>
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<mods:nameIdentifier id="9FC7571011F5C3707F2658CE1139C991" type="email">twesener@uni-bonn.de</mods:nameIdentifier>
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<mods:date>2020</mods:date>
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<treatment ID-GBIF-Taxon="163875344" LSID="urn:lsid:zoobank.org:act:8C6B0566-6BEE-4A06-B59F-BDA14A90892B" httpUri="http://treatment.plazi.org/id/4FCE290506CA53B5B8B78EFAD5029C4F" lastPageNumber="37" pageId="0" pageNumber="37">
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<taxonomicName LSID="4FCE2905-06CA-53B5-B8B7-8EFAD5029C4F" authority="Semenyuk & Golovatch & Wesener, 2020" authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Diplopoda" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="honbaensis" status="sp. nov.">Sphaeropoeus honbaensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="37">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live Sphaeropoeus bidoupensis sp. nov. (A-E) and Sphaeropoeus honbaensis sp. nov. (F-H) in the field A enrolled male (left) and female (right) B adult, ventral view C adult and its excrements in a " living room " in leaf litter D early instar juvenile enrolled in its " living room ", with surrounding dead leaves eaten and excrement used to shape the room E enrolled middle-sized instar juvenile F-H adult, enrolled and walking, respectively. Pictures taken not to scale." figureDoi="10.3897/zookeys.930.47742.figure2" httpUri="https://binary.pensoft.net/fig/404783" pageId="0" pageNumber="37">Figs 2F-H</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endoterga of midbody segments, SEM micrographs. A Sphaerobelum bicorne Attems, 1938, ♂ (ZFMK) B Sphaerobelum pumatense sp. nov., ♂ holotype C Sphaeropoeus maculatus (Verhoeff, 1924), ♂ lectotype D Sphaeropoeus bidoupensis sp. nov., ♂ holotype E Sphaeropoeus honbaensis sp. nov., ♂ holotype. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37">, 3E</figureCitation>
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">, 8</figureCitation>
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<treatment id="4FCE290506CA53B5B8B78EFAD5029C4F" ID-GBIF-Taxon="163875344" LSID="urn:lsid:zoobank.org:act:8C6B0566-6BEE-4A06-B59F-BDA14A90892B" httpUri="http://treatment.plazi.org/id/4FCE290506CA53B5B8B78EFAD5029C4F" lastPageNumber="37" pageId="0" pageNumber="37" scope_class="Diplopoda" scope_order="Sphaerotheriida">
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<subSubSection id="E9C779589121B23EAD49FE9A1B91D31E" pageId="0" pageNumber="37" type="nomenclature">
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<paragraph id="866A6D01DCBA82DD5C5D17D9C9A94A91" pageId="0" pageNumber="37">
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<taxonomicName id="80CE6084641E403C330B33CEC221E433" LSID="4FCE2905-06CA-53B5-B8B7-8EFAD5029C4F" authority="Semenyuk & Golovatch & Wesener, 2020" authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Diplopoda" pageId="0" pageNumber="37" rank="species" species="honbaensis" status="sp. nov.">Sphaeropoeus honbaensis</taxonomicName>
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<taxonomicNameLabel id="3C9EBC03624CD5B143E3D7EC8FA61FEA" pageId="0" pageNumber="37">sp. nov.</taxonomicNameLabel>
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<figureCitation id="44A1F0B51B0CBD299740CF5EB9897319" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live Sphaeropoeus bidoupensis sp. nov. (A-E) and Sphaeropoeus honbaensis sp. nov. (F-H) in the field A enrolled male (left) and female (right) B adult, ventral view C adult and its excrements in a " living room " in leaf litter D early instar juvenile enrolled in its " living room ", with surrounding dead leaves eaten and excrement used to shape the room E enrolled middle-sized instar juvenile F-H adult, enrolled and walking, respectively. Pictures taken not to scale." figureDoi="10.3897/zookeys.930.47742.figure2" httpUri="https://binary.pensoft.net/fig/404783" pageId="0" pageNumber="37">Figs 2F-H</figureCitation>
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||||
<figureCitation id="45799460B04CE88A50AFBF6B37486AE4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endoterga of midbody segments, SEM micrographs. A Sphaerobelum bicorne Attems, 1938, ♂ (ZFMK) B Sphaerobelum pumatense sp. nov., ♂ holotype C Sphaeropoeus maculatus (Verhoeff, 1924), ♂ lectotype D Sphaeropoeus bidoupensis sp. nov., ♂ holotype E Sphaeropoeus honbaensis sp. nov., ♂ holotype. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37">, 3E</figureCitation>
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||||
<figureCitation id="733C13D3D7E4D22B9DA14A37C3488720" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">, 8</figureCitation>
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</paragraph>
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||||
</subSubSection>
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<subSubSection pageId="0" pageNumber="37" type="materials_examined">
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||||
<paragraph pageId="0" pageNumber="37">Material examined.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="37">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Holotype</emphasis>
|
||||
<subSubSection id="12344A4B4116772A7E8D1897C6220A71" pageId="0" pageNumber="37" type="materials_examined">
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<paragraph id="D82A4C3F53BF0CEFC36B263A0F57F2A3" pageId="0" pageNumber="37">Material examined.</paragraph>
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||||
<paragraph id="7FC3BE96AFAE2C8173AFFB0D4DB416D3" pageId="0" pageNumber="37">
|
||||
<emphasis id="9D498554423AC56A2314F4AA2C8EED9E" bold="true" italics="true" pageId="0" pageNumber="37">Holotype</emphasis>
|
||||
♂ (ZMUM Rd 4644), Vietnam, Khanh Hoa Prov., Hon Ba Nature Reserve,
|
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<geoCoordinate degrees="12" direction="north" minutes="07" orientation="latitude" precision="925" value="12.116667">12°07'N</geoCoordinate>
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||||
<geoCoordinate id="6DF3FBD7FC66E82AD9449812074586BF" degrees="12" direction="north" minutes="07" orientation="latitude" precision="925" value="12.116667">12°07'N</geoCoordinate>
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||||
,
|
||||
<geoCoordinate degrees="108" direction="east" minutes="56" orientation="longitude" precision="925" value="108.933334">108°56'E</geoCoordinate>
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<geoCoordinate id="FB0B1FF47EF32811A4459D21538EC570" degrees="108" direction="east" minutes="56" orientation="longitude" precision="925" value="108.933334">108°56'E</geoCoordinate>
|
||||
, 1550 m a.s.l., mixed mossy tropical forest on mountain ridge, on forest floor, night time, VI.2018, I.I. Semenyuk leg.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Paratypes</emphasis>
|
||||
<emphasis id="CDD514EC6B565A3155B087E6318D41AB" bold="true" italics="true" pageId="0" pageNumber="37">Paratypes</emphasis>
|
||||
2 ♂ (ZMUM Rd 4633), 1 ♂ (ZFMK MYR8943), same locality as holotype. 1 ♀ (ZMUM Rd 4645), same locality, 1450 m a.s.l., mixed tropical forest on mountain slope, in leaf litter, day time, VI.2018, I.I. Semenyuk leg.
|
||||
</paragraph>
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||||
<caption doi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37" start="Figure 8" startId="F8">
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||||
<paragraph pageId="0" pageNumber="37">
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||||
<emphasis bold="true" pageId="0" pageNumber="37">Figure 8.</emphasis>
|
||||
<taxonomicName authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="honbaensis">
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||||
<emphasis italics="true" pageId="0" pageNumber="37">Sphaeropoeus honbaensis</emphasis>
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||||
<caption id="8CE536374E669C879FA5D64CFA08C95B" doi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37" start="Figure 8" startId="F8">
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<paragraph id="FC23196632AB20A595316D6EE21C7A78" pageId="0" pageNumber="37">
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<emphasis id="38B38D312596AD7F68308A12CC29914E" bold="true" pageId="0" pageNumber="37">Figure 8.</emphasis>
|
||||
<taxonomicName id="CBD9B01E611DF85871FD01FE4D1627CE" authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="honbaensis">
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<emphasis id="D2B55C3E9F8E1869367D09C9442EEEE1" italics="true" pageId="0" pageNumber="37">Sphaeropoeus honbaensis</emphasis>
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||||
</taxonomicName>
|
||||
sp. nov., ♂ holotype (
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">A-G</emphasis>
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||||
<emphasis id="1AFA13E068FE56097AA6D0FBF4B495DA" bold="true" pageId="0" pageNumber="37">A-G</emphasis>
|
||||
) and ♀ paratype (
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">H</emphasis>
|
||||
<emphasis id="AC2DF416883D6D68C252496379E80A90" bold="true" pageId="0" pageNumber="37">H</emphasis>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
|
||||
<emphasis id="7BB70EBD90857032653C86DE6483809A" bold="true" pageId="0" pageNumber="37">A</emphasis>
|
||||
Left antenna,
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
|
||||
<emphasis id="E6567B986EB3B6BF0D736B388D5CCE35" bold="true" pageId="0" pageNumber="37">B</emphasis>
|
||||
left leg 9, anterior view
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">C, D</emphasis>
|
||||
<emphasis id="4D4D46F6C82D9FD80B4A33C43AE7E760" bold="true" pageId="0" pageNumber="37">C, D</emphasis>
|
||||
left posterior telopod, posterior and anterior views, respectively
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">E-G</emphasis>
|
||||
<emphasis id="0944757F7D6A8755A471AB5D9229B4D4" bold="true" pageId="0" pageNumber="37">E-G</emphasis>
|
||||
left anterior telopod, anterior, posterior and lateral views, respectively
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">H</emphasis>
|
||||
<emphasis id="45E67D0E52B56DED89804F36B8D8E3F3" bold="true" pageId="0" pageNumber="37">H</emphasis>
|
||||
left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm,
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">G</emphasis>
|
||||
<emphasis id="C452257D01F771B80E2E6D00068B1BD5" bold="true" pageId="0" pageNumber="37">G</emphasis>
|
||||
drawn not to scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="37" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="37">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="37">
|
||||
<taxonomicName authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="honbaensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">Sphaeropoeus honbaensis</emphasis>
|
||||
<subSubSection id="8087856EC0B6F9EF6286D616AF25BAA8" pageId="0" pageNumber="37" type="diagnosis">
|
||||
<paragraph id="BC677880D57A63F4B5B9C0B4A5049DF7" pageId="0" pageNumber="37">Diagnosis.</paragraph>
|
||||
<paragraph id="00FB7D6D304012ABDD80D5640D8B7FA3" pageId="0" pageNumber="37">
|
||||
<taxonomicName id="4660942BF5BE38E63968721DAB53D14E" authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="honbaensis">
|
||||
<emphasis id="3D11640A19C6FCACD71C342A8710208E" italics="true" pageId="0" pageNumber="37">Sphaeropoeus honbaensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. differs from all other known continental species of the genus in the presence of very few (<30) apical cones on the ♂ antenna (usually at least>70, often>120). The anterior telopod also shows a very short, almost completely reduced telopoditomere 4, a character only shared with
|
||||
<taxonomicName lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
<taxonomicName id="22F01B60C24208C2B591D23A564C22E0" lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis id="B95096CFEFA101BA9804DEC67A590B96" italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.
|
||||
<taxonomicName authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="honbaensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">Sphaeropoeus honbaensis</emphasis>
|
||||
<taxonomicName id="ED3589E8E26CB0DD0FDB4FFE5052B56C" authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="honbaensis">
|
||||
<emphasis id="5FB833384F4930A277E4BADB4404EC48" italics="true" pageId="0" pageNumber="37">Sphaeropoeus honbaensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. differs from
|
||||
<taxonomicName lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
<taxonomicName id="988EADA22DE314D033F3582280828304" lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis id="782153A4E0D0F472AA4F6D21D917D86A" italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. in the lack of a spine in the inner area of the large telopoditomere 3 (vs. present in
|
||||
<taxonomicName lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
<taxonomicName id="E74780CDE31F7260D90373818345EC4C" lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis id="97CFDB83E3AE4631DDD2FCBFDA31FF79" italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.), in the endotergum, where the distances between the cuticular impressions are wider than the diameter (vs. slightly narrower than diameter in
|
||||
<taxonomicName lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
<taxonomicName id="8611AC5D03BA5C649350551639C23804" lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis id="16B8B06EDACC1C8383574401AEB72AF2" italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.), the ♂ anal shield being weakly bell-shaped (vs. well-rounded in
|
||||
<taxonomicName lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
<taxonomicName id="108A3F5741C272B05EDB6CEEFA539F8E" lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis id="4DC58840A99EF85C6E834614210E2B9E" italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.), and in leg structure, with leg-pair 3 lacking an apical spine (vs. present in
|
||||
<taxonomicName lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
<taxonomicName id="7EBE1B2FFE0A57032307802CE1F0687F" lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis id="1E13CC31A2310EA24F1BE9BEFBC6B3D8" italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.), the prefemur lacking a dentate mesal margin (vs. present in
|
||||
<taxonomicName lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
<taxonomicName id="7BA7951CD77E5EE9AF38C5F9C6A78B23" lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis id="413810F1043D8EDA8364991A27CF1EDB" italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.), and the coxa process being strongly developed and well-rounded (vs. weakly developed and partly sharp in
|
||||
<taxonomicName lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
<taxonomicName id="24B1CC744B7482AEB07409864B0E6607" lsidName="S. bidoupensis" pageId="0" pageNumber="37" rank="species" species="bidoupensis">
|
||||
<emphasis id="E281A3C16D3886D5C3894652CBACA50A" italics="true" pageId="0" pageNumber="37">S. bidoupensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov.).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="37" type="description">
|
||||
<paragraph pageId="0" pageNumber="37">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="37">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Measurements</emphasis>
|
||||
<subSubSection id="0137AC8CB966E35425F65557EBA37D3C" pageId="0" pageNumber="37" type="description">
|
||||
<paragraph id="B23D3608E6FD09422008510292B1FE88" pageId="0" pageNumber="37">Description.</paragraph>
|
||||
<paragraph id="F46D92060AA3E33EA489949C9537B1C3" pageId="0" pageNumber="37">
|
||||
<emphasis id="A647A8D9F1A6EA06158B24F2A6A342ED" bold="true" italics="true" pageId="0" pageNumber="37">Measurements</emphasis>
|
||||
: holotype ♂ ca. 27 mm long, 12.1 mm (2nd), up to 12.2 mm (7th) wide, 7.8 mm (2nd) up to 15.1 mm height (8th the highest); ♂ paratypes 8-11 mm wide. Paratype ♀ ca. 29 mm long, 13.1 mm (2nd), up to 13.3 mm (7th) wide, 7.8 mm (2nd) up to 8.4 mm height (7th the highest).
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Coloration</emphasis>
|
||||
<emphasis id="891EDC7C893B7123AB28899517884E58" bold="true" italics="true" pageId="0" pageNumber="37">Coloration</emphasis>
|
||||
: both in vivo and in vitro, after>1.5 years of preservation in ethanol, similar, in life uniformly dark violet brown to violet blackish with vague infuscate bands near caudal margin (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live Sphaeropoeus bidoupensis sp. nov. (A-E) and Sphaeropoeus honbaensis sp. nov. (F-H) in the field A enrolled male (left) and female (right) B adult, ventral view C adult and its excrements in a " living room " in leaf litter D early instar juvenile enrolled in its " living room ", with surrounding dead leaves eaten and excrement used to shape the room E enrolled middle-sized instar juvenile F-H adult, enrolled and walking, respectively. Pictures taken not to scale." figureDoi="10.3897/zookeys.930.47742.figure2" httpUri="https://binary.pensoft.net/fig/404783" pageId="0" pageNumber="37">2F-H</figureCitation>
|
||||
<figureCitation id="7F08DA112FBE2E6EDD1C6FCE682DD02A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Live Sphaeropoeus bidoupensis sp. nov. (A-E) and Sphaeropoeus honbaensis sp. nov. (F-H) in the field A enrolled male (left) and female (right) B adult, ventral view C adult and its excrements in a " living room " in leaf litter D early instar juvenile enrolled in its " living room ", with surrounding dead leaves eaten and excrement used to shape the room E enrolled middle-sized instar juvenile F-H adult, enrolled and walking, respectively. Pictures taken not to scale." figureDoi="10.3897/zookeys.930.47742.figure2" httpUri="https://binary.pensoft.net/fig/404783" pageId="0" pageNumber="37">2F-H</figureCitation>
|
||||
), in alcohol dark brown to brown, in places marbled, only dorsalmost part of anal shield sometimes lighter centrally, light brown. Antennae orange. Legs mostly grey- or olive-brown, but tarsi yellow-brown. Tegument mosly dull to poorly shining.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Head</emphasis>
|
||||
<emphasis id="A47BE4B903EAA6A80C091ED64DF539F3" bold="true" italics="true" pageId="0" pageNumber="37">Head</emphasis>
|
||||
: eyes with ca. 65 ocelli. Antennae short (Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8A</figureCitation>
|
||||
<figureCitation id="D080D4790090AC714F1A5BC11121748C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8A</figureCitation>
|
||||
), protruding beyond centre of head. Antennomeres 1-5 with few longer setae, 6th densely pubescent. Antennomere 6 towards disc with single row of sensilla basiconica. Antennomere 6 slightly swollen in ♂, cylindrical in ♀, twice as long as, but only slightly wider than, antennomeres 1-5. ♂ with 26/27, ♀ with 22/24 apical cones. Palpi of gnathochilarium located in a single field.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Collum</emphasis>
|
||||
<emphasis id="D6ECB7C02C7E962EE552D3908084C981" bold="true" italics="true" pageId="0" pageNumber="37">Collum</emphasis>
|
||||
: completely covered with long setae, like the tergites.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Thoracic shield</emphasis>
|
||||
<emphasis id="CC8C6CD6FD81151789333055586AECA2" bold="true" italics="true" pageId="0" pageNumber="37">Thoracic shield</emphasis>
|
||||
: with wide and shallow grooves, 3 or 4 weak crests present at posterior corner.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Tergites</emphasis>
|
||||
<emphasis id="B298177320DEC4FEF79A6703EB0A630C" bold="true" italics="true" pageId="0" pageNumber="37">Tergites</emphasis>
|
||||
: surface covered with longer setae, most innervating in small pits. Midbody paratergite tips projecting posteriorly (Fig.
|
||||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus photograph of the lectotype (HNHM 2858 / 1) of Sphaeropoeus maculatus (Verhoeff, 1924), lateral view, and a distribution map of all other relevant zephroniid species in Vietnam. A Sphaerobelum bicorne Attems, 1938 (new record) B Sphaerobelum bicorne Attems, 1938 (old record) C Sphaerobelum pumatense sp. nov. D Sphaeropoeus manca (Attems, 1936) E Sphaeropoeus maculatus (Verhoeff, 1924) F Sphaeropoeus honbaensis sp. nov. G Sphaeropoeus bidoupensis sp. nov." figureDoi="10.3897/zookeys.930.47742.figure10" httpUri="https://binary.pensoft.net/fig/404791" pageId="0" pageNumber="37">10</figureCitation>
|
||||
<figureCitation id="3C9F824F6AF1EA32FCBF4D780779640D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Habitus photograph of the lectotype (HNHM 2858 / 1) of Sphaeropoeus maculatus (Verhoeff, 1924), lateral view, and a distribution map of all other relevant zephroniid species in Vietnam. A Sphaerobelum bicorne Attems, 1938 (new record) B Sphaerobelum bicorne Attems, 1938 (old record) C Sphaerobelum pumatense sp. nov. D Sphaeropoeus manca (Attems, 1936) E Sphaeropoeus maculatus (Verhoeff, 1924) F Sphaeropoeus honbaensis sp. nov. G Sphaeropoeus bidoupensis sp. nov." figureDoi="10.3897/zookeys.930.47742.figure10" httpUri="https://binary.pensoft.net/fig/404791" pageId="0" pageNumber="37">10</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Anal shield</emphasis>
|
||||
<emphasis id="7E1AEFE247C243E824EA9ED5ED02C4AA" bold="true" italics="true" pageId="0" pageNumber="37">Anal shield</emphasis>
|
||||
: well-rounded in ♀, weakly bell-shaped in ♂. In both sexes completely covered with longer setae. Locking carina long, twice as long as width of last laterotergite, located close to margin.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Endotergum</emphasis>
|
||||
<emphasis id="101B0515495C085F169305DCB4B0633C" bold="true" italics="true" pageId="0" pageNumber="37">Endotergum</emphasis>
|
||||
: inner section lacking any spines or setae. Middle area with a single row of large, sparse, elliptical, cuticular impressions. Distance between impressions greater than their diameter. Apically, two rows of long marginal bristles, tips of longest setae clearly protruding beyond tergal margin (Fig.
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endoterga of midbody segments, SEM micrographs. A Sphaerobelum bicorne Attems, 1938, ♂ (ZFMK) B Sphaerobelum pumatense sp. nov., ♂ holotype C Sphaeropoeus maculatus (Verhoeff, 1924), ♂ lectotype D Sphaeropoeus bidoupensis sp. nov., ♂ holotype E Sphaeropoeus honbaensis sp. nov., ♂ holotype. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37">3E</figureCitation>
|
||||
<figureCitation id="1F4B4E8CC61831B43C030D3454A1829D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endoterga of midbody segments, SEM micrographs. A Sphaerobelum bicorne Attems, 1938, ♂ (ZFMK) B Sphaerobelum pumatense sp. nov., ♂ holotype C Sphaeropoeus maculatus (Verhoeff, 1924), ♂ lectotype D Sphaeropoeus bidoupensis sp. nov., ♂ holotype E Sphaeropoeus honbaensis sp. nov., ♂ holotype. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37">3E</figureCitation>
|
||||
). Bristles not smooth, but with numerous small spinicles.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Stigmatic plates</emphasis>
|
||||
<emphasis id="70DFB4290B790DBB35AFCBF8D0E629A3" bold="true" italics="true" pageId="0" pageNumber="37">Stigmatic plates</emphasis>
|
||||
: first well-rounded, triangular.
|
||||
<taxonomicName genus="Laterotergites" lsidName="Laterotergites" pageId="0" pageNumber="37" rank="genus">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Laterotergites</emphasis>
|
||||
<taxonomicName id="3CD6AEDAB830356DEB54FB4C1E885689" genus="Laterotergites" lsidName="Laterotergites" pageId="0" pageNumber="37" rank="genus">
|
||||
<emphasis id="B8BA4084A07CD6002347184AB3AB6237" bold="true" italics="true" pageId="0" pageNumber="37">Laterotergites</emphasis>
|
||||
</taxonomicName>
|
||||
: first with a slightly projecting, well-rounded process.
|
||||
<taxonomicName genus="Laterotergites" lsidName="Laterotergites" pageId="0" pageNumber="37" rank="genus">Laterotergites</taxonomicName>
|
||||
<taxonomicName id="40BED2490BF7BCC7016983221A55E27F" genus="Laterotergites" lsidName="Laterotergites" pageId="0" pageNumber="37" rank="genus">Laterotergites</taxonomicName>
|
||||
2 and following not extended, well-rounded.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Legs</emphasis>
|
||||
<emphasis id="8E5143A23A248D865A0449EAB9D9FA69" bold="true" italics="true" pageId="0" pageNumber="37">Legs</emphasis>
|
||||
: first with 2 or 3, second with 3 or 4, third with 8 ventral and lacking an apical spine. Leg-pairs 4-21 each with 10-12 ventral spines and a single apical spine. Coxa process visible, well-rounded (Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8B</figureCitation>
|
||||
<figureCitation id="9169E617467584D8A8042665A2FAC2C5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8B</figureCitation>
|
||||
). Femur 1.7, tarsus 5.2
|
||||
<normalizedToken originalValue="×">x</normalizedToken>
|
||||
<normalizedToken id="4F7FEF2753D3857FF636915448E24D5A" originalValue="×">x</normalizedToken>
|
||||
as long as wide. Femur with a very long ridge (Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8B</figureCitation>
|
||||
<figureCitation id="FB1437383530C6F3C9BB23C7CE6BB48F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8B</figureCitation>
|
||||
). Mesal margin of femur completely extended into 12-14 teeth, lateral margin of prefemur, juxtaposed to coxal process, slightly extended into 2-4 teeth (Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8B</figureCitation>
|
||||
<figureCitation id="D71430BC60C4699113F79AB47E6E6640" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8B</figureCitation>
|
||||
).
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Female sexual characters</emphasis>
|
||||
<emphasis id="DA480C6D4F598670EB0647E980E9D254" bold="true" italics="true" pageId="0" pageNumber="37">Female sexual characters</emphasis>
|
||||
: vulva large, covering 2/3 coxa, a conspicuous operculum extending to basal half of prefemur (Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8H</figureCitation>
|
||||
<figureCitation id="711EA9516DC65FD99CA540EACE981BD6" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8H</figureCitation>
|
||||
). Operculum massive, larger than bursa, wider than prefemur, apically rounded, mesal margin very slightly projecting and slightly higher than remaining operculum.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Subanal plate</emphasis>
|
||||
<emphasis id="818AC115F7A637481D8F1F3430F12757" bold="true" italics="true" pageId="0" pageNumber="37">Subanal plate</emphasis>
|
||||
: large and wide, triangular.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Male gonopore</emphasis>
|
||||
<emphasis id="26DA357338C8A93849271A4CE3EC89C5" bold="true" italics="true" pageId="0" pageNumber="37">Male gonopore</emphasis>
|
||||
: opening covered with a single, apically membranous plate.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Anterior telopod</emphasis>
|
||||
<emphasis id="D7C8F4239A585B09027AE3117E10004F" bold="true" italics="true" pageId="0" pageNumber="37">Anterior telopod</emphasis>
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8E-G</figureCitation>
|
||||
<figureCitation id="41C9FD253EF0DA9AEC911C5FFB915D3F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8E-G</figureCitation>
|
||||
): four podomeres, first three of equal length regardless of the processes, podomere 4 rudimentary, conical. Telopoditomere 2 with a strong, curved process overreaching telopoditomere 4. Telopoditomere 3 posteriorly with a longer process juxtaposed to apex process of telopoditomere 2, clearly protruding above telopoditomere 4, as well as process of telopoditomere 2. Telopoditomere 4 conical, with two spines.
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Posterior telopod</emphasis>
|
||||
<emphasis id="FC9DB81E89C644C14C75DF8B080330BF" bold="true" italics="true" pageId="0" pageNumber="37">Posterior telopod</emphasis>
|
||||
(Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8C, D</figureCitation>
|
||||
<figureCitation id="8AABD1DDBB1C91BF20CA50BD70A7C2C6" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sphaeropoeus honbaensis sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A Left antenna, B left leg 9, anterior view C, D left posterior telopod, posterior and anterior views, respectively E-G left anterior telopod, anterior, posterior and lateral views, respectively H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, G drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure8" httpUri="https://binary.pensoft.net/fig/404789" pageId="0" pageNumber="37">8C, D</figureCitation>
|
||||
): Podomeres 3 and 4 slightly longer than process of podomere 2. Podomere 4 short, conical, with two spines, slightly curved towards immovable finger. Podomere 3 slender, 4.2 times longer than wide. Its excavate inner margin with a membranous lobe and a single spine, posterior aspect with ca. 12 small crenulated teeth. Immovable finger slender, apically tapering, tip curved towards movable finger. Membranous area apically with a large membranous lobe. Podomeres 1-3 in anterior and posterior views with few setae. Podomere 4 in both aspects glabrous.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.930.47742.figure9" httpUri="https://binary.pensoft.net/fig/404790" pageId="0" pageNumber="37" start="Figure 9" startId="F9">
|
||||
<paragraph pageId="0" pageNumber="37">
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">Figure 9.</emphasis>
|
||||
<taxonomicName authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus bidoupensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bidoupensis">
|
||||
<emphasis italics="true" pageId="0" pageNumber="37">Sphaeropoeus bidoupensis</emphasis>
|
||||
<caption id="4C31F7511DACE51770C3A1BCA3EDE872" doi="10.3897/zookeys.930.47742.figure9" httpUri="https://binary.pensoft.net/fig/404790" pageId="0" pageNumber="37" start="Figure 9" startId="F9">
|
||||
<paragraph id="11B438DF1A61EA3C667DFAB1D3C9964E" pageId="0" pageNumber="37">
|
||||
<emphasis id="973FE90F2F2F1CA9BF3A41FF43A18048" bold="true" pageId="0" pageNumber="37">Figure 9.</emphasis>
|
||||
<taxonomicName id="328B709A63E224A8F257783E68413EC4" authorityName="Semenyuk & Golovatch & Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus bidoupensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bidoupensis">
|
||||
<emphasis id="A2C7BC2B084328FC3BB41B87993FE685" italics="true" pageId="0" pageNumber="37">Sphaeropoeus bidoupensis</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., ♂ holotype (
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">A-H</emphasis>
|
||||
<emphasis id="BC8DD064EAE399015FA9AD632871E6C5" bold="true" pageId="0" pageNumber="37">A-H</emphasis>
|
||||
) and ♀ paratype (
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">I</emphasis>
|
||||
<emphasis id="234FDDB8EF6D10D8C5AD41B01CDE63CD" bold="true" pageId="0" pageNumber="37">I</emphasis>
|
||||
).
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">A, B</emphasis>
|
||||
<emphasis id="A8B01444961F444B9B361D440A5CE6FC" bold="true" pageId="0" pageNumber="37">A, B</emphasis>
|
||||
Left posterior telopod, posterior and anterior views, respectively
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">C-F</emphasis>
|
||||
<emphasis id="408BDB661B61106EE37F4E5B8BEA905A" bold="true" pageId="0" pageNumber="37">C-F</emphasis>
|
||||
left anterior telopod, posterior, sublateral, lateral and anterior views, respectively
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">G</emphasis>
|
||||
<emphasis id="39EACAAC95D936819F15EC2DE06B05C2" bold="true" pageId="0" pageNumber="37">G</emphasis>
|
||||
left leg 9, anterior view
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">H</emphasis>
|
||||
<emphasis id="959411597C892F2195D05F54E1269975" bold="true" pageId="0" pageNumber="37">H</emphasis>
|
||||
left antenna
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">I</emphasis>
|
||||
<emphasis id="D403CCCFAAAD56B8694A16C6334AC49B" bold="true" pageId="0" pageNumber="37">I</emphasis>
|
||||
left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm,
|
||||
<emphasis bold="true" pageId="0" pageNumber="37">D</emphasis>
|
||||
<emphasis id="35695E884EF8466694A6E99E29309189" bold="true" pageId="0" pageNumber="37">D</emphasis>
|
||||
drawn not to scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="37" type="etymology">
|
||||
<paragraph pageId="0" pageNumber="37">Etymology.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="37">To emphasize the provenance from the Hon Ba Nature Reserve, adjective.</paragraph>
|
||||
<subSubSection id="3CE38B6F93F3A7C1F8C8732EEC4180C8" pageId="0" pageNumber="37" type="etymology">
|
||||
<paragraph id="A5FFCFB7F797751D96772C17F4BC0B38" pageId="0" pageNumber="37">Etymology.</paragraph>
|
||||
<paragraph id="E22ADB8E387A634321E9E05188C822C6" pageId="0" pageNumber="37">To emphasize the provenance from the Hon Ba Nature Reserve, adjective.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="37" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="37">Remarks.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="37">During the expedition, more than ten males of this species were recorded, all walking on the forest floor in the night, mainly on the mountain ridge. Only one adult female was found despite special searching efforts: it was hidden in leaf litter, which is typical of sphaerotheriidans. The female was encountered in a forest at an elevation considerably lower than the where abundant males were observed. Although confusing, this may be accounted for by high population abundance at the onset of a season. During the expedition, it was raining almost every day, the temperature in the daytime above leaf litter averaging 20 °C and dropping down to 17.5 °C (minimum 16.8 °C) at night. Fog was very often seen on the top of the mountain. No juveniles were recorded.</paragraph>
|
||||
<subSubSection id="A1C59E38771524C22DBA677DF887FA93" pageId="0" pageNumber="37" type="remarks">
|
||||
<paragraph id="E2DD9E7FA9FE5246C55A31CF911AEE82" pageId="0" pageNumber="37">Remarks.</paragraph>
|
||||
<paragraph id="7DCBA080CC2806987161517CCCCB9988" pageId="0" pageNumber="37">During the expedition, more than ten males of this species were recorded, all walking on the forest floor in the night, mainly on the mountain ridge. Only one adult female was found despite special searching efforts: it was hidden in leaf litter, which is typical of sphaerotheriidans. The female was encountered in a forest at an elevation considerably lower than the where abundant males were observed. Although confusing, this may be accounted for by high population abundance at the onset of a season. During the expedition, it was raining almost every day, the temperature in the daytime above leaf litter averaging 20 °C and dropping down to 17.5 °C (minimum 16.8 °C) at night. Fog was very often seen on the top of the mountain. No juveniles were recorded.</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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