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<figureCitation captionStart="Figures 12" captionStartId="F1" captionText="Figures 1 - 2. Madagopsina parvapollina, live photographs by Gernot Kunz, Mahajanga, Boeny (www. inaturalist. org / observations / 20766277) 1 anterior view 2 dorsal view." figureDoi="10.3897/zookeys.1057.67433.figures1-2" httpUri="https://binary.pensoft.net/fig/580793" pageId="0" pageNumber="1">Figures 1-2</figureCitation>
<figureCitation captionStart="Figures 34" captionStartId="F2" captionText="Figures 3 - 4. Madagopsina makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 3 habitus, dorsolateral view 4 thorax, dorsolateral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1057.67433.figures3-4" httpUri="https://binary.pensoft.net/fig/580794" pageId="0" pageNumber="1">, 3-4</figureCitation>
<figureCitation captionStart="Figures 58" captionStartId="F3" captionText="Figures 5 - 8. Madagopsina makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 5 head, anterior view 6 central head, anterior view 7 scutellar spine and apical seta, inner view (seta not in natural position in line with spine) 8 abdomen, dorsal view. Scale bars: 1 mm (5); 0.5 mm (6 - 8)." figureDoi="10.3897/zookeys.1057.67433.figures5-8" httpUri="https://binary.pensoft.net/fig/580795" pageId="0" pageNumber="1">, 5-8</figureCitation>
<figureCitation captionStart="Figures 911" captionStartId="F4" captionText="Figures 9 - 11. Madagopsina apollo species group, dorsal view of wings 9 M. apollo, ♂, Ambohitra 10 M. makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 11 M. parvapollina, ♀, paratype, Ankarana. Scale bars: 0.5 mm. Figures 9, 11 (Feijen et al. 2018, figures 6, 7)." figureDoi="10.3897/zookeys.1057.67433.figures9-11" httpUri="https://binary.pensoft.net/fig/580796" pageId="0" pageNumber="1">, 9-11</figureCitation>
<figureCitation captionStart="Figures 1214" captionStartId="F5" captionText="Figures 12 - 14. Madagopsina apographica species group, dorsal view of wings 12 M. apographica, ♀, Fianarantsoa 13 M. freidbergi, ♀, paratype, Vohimana 14 M. tschirnhausi, ♂, holotype, Mount Ambre. Scale bars 0.5 mm. Figures 12 - 14 (Feijen et al. 2018, figures 8 - 10)." figureDoi="10.3897/zookeys.1057.67433.figures12-14" httpUri="https://binary.pensoft.net/fig/580797" pageId="0" pageNumber="1">, 12-14</figureCitation>
<figureCitation captionStart="Figures 1517" captionStartId="F6" captionText="Figures 15 - 17. Madagopsina makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 15 basal section of abdomen with intersternite 1 - 2, ventral view 16 sternites 5 and 6, ventral view 17 synsternite 7 + 8, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1057.67433.figures15-17" httpUri="https://binary.pensoft.net/fig/580798" pageId="0" pageNumber="1">, 15-17</figureCitation>
<figureCitation captionStart="Figures 1822" captionStartId="F7" captionText="Figures 18 - 22. Madagopsina makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 18 epandrium with surstyli and cerci, posterior view 19 surstylus, inner view 20 hypandrial clasper, lateral view 21 phallapodeme and phallus, lateral view 22 ejaculatory apodeme and sac. Scale bars: 0.5 mm (18, 21); 0.1 mm (19, 20, 22)." figureDoi="10.3897/zookeys.1057.67433.figures18-22" httpUri="https://binary.pensoft.net/fig/580799" pageId="0" pageNumber="1">, 18-22</figureCitation>
<figureCitation captionStart="Figures 2328" captionStartId="F8" captionText="Figures 23 - 28. Posterior view of Madagopsina surstyli 23 - 25 M. apollo species group 26 - 28 M. apographica species group 23 M. apollo 24 M. makayensis Feijen, Feijen &amp; Feijen, sp. nov. 25 M. parvapollina 26 M. apographica 27 M. freidbergi 28 M. tschirnhausi. Scale bar 0.1 mm (all drawn to the same scale). Figures 25 - 28 (Feijen et al. 2018, figures 187, 184, 186, 188)." figureDoi="10.3897/zookeys.1057.67433.figures23-28" httpUri="https://binary.pensoft.net/fig/580800" pageId="0" pageNumber="1">, 23-28</figureCitation>
<figureCitation captionStart="Figure 29" captionStartId="F9" captionText="Figure 29. Principal component analysis of wing venation morphometry for the two species of Gracilopsina and six species of Madagopsina: biplot of the first two PCA axes." figureDoi="10.3897/zookeys.1057.67433.figure29" httpUri="https://binary.pensoft.net/fig/580801" pageId="0" pageNumber="1">, 29</figureCitation>
<figureCitation captionStart="Figure 30" captionStartId="F10" captionText="Figure 30. Cluster dendrogram for the Euclidian distance in wing morphometry PCA for the Gracilopsina and Madagopsina species using the complete clustering method. Branch labels give the approximately unbiased p - value (AU) and bootstrap probability (BP) values (%)." figureDoi="10.3897/zookeys.1057.67433.figure30" httpUri="https://binary.pensoft.net/fig/580802" pageId="0" pageNumber="1">, 30</figureCitation>
<figureCitation captionStart="Figure 31" captionStartId="F11" captionText="Figure 31. Eye span plotted against body length for the three species in the Madagopsina apollo species group: M. apollo, M. parvapollina and M. makayensis Feijen, Feijen &amp; Feijen, sp. nov. Note the position of the single data point for the ♂ of the latter species, in line with the ♂ data points for M. parvapollina." figureDoi="10.3897/zookeys.1057.67433.figure31" httpUri="https://binary.pensoft.net/fig/580803" pageId="0" pageNumber="1">, 31</figureCitation>
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<figureCitation id="545DD0DE531EBC8FBE7050C7F25E7294" captionStart="Figures 12" captionStartId="F1" captionText="Figures 1 - 2. Madagopsina parvapollina, live photographs by Gernot Kunz, Mahajanga, Boeny (www. inaturalist. org / observations / 20766277) 1 anterior view 2 dorsal view." figureDoi="10.3897/zookeys.1057.67433.figures1-2" httpUri="https://binary.pensoft.net/fig/580793" pageId="0" pageNumber="1">Figures 1-2</figureCitation>
<figureCitation id="760ECEFFB976C5829ED4890BC19EABE6" captionStart="Figures 34" captionStartId="F2" captionText="Figures 3 - 4. Madagopsina makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 3 habitus, dorsolateral view 4 thorax, dorsolateral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1057.67433.figures3-4" httpUri="https://binary.pensoft.net/fig/580794" pageId="0" pageNumber="1">, 3-4</figureCitation>
<figureCitation id="4555DBB8CE1BA337DFD1DACA5742E2B9" captionStart="Figures 58" captionStartId="F3" captionText="Figures 5 - 8. Madagopsina makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 5 head, anterior view 6 central head, anterior view 7 scutellar spine and apical seta, inner view (seta not in natural position in line with spine) 8 abdomen, dorsal view. Scale bars: 1 mm (5); 0.5 mm (6 - 8)." figureDoi="10.3897/zookeys.1057.67433.figures5-8" httpUri="https://binary.pensoft.net/fig/580795" pageId="0" pageNumber="1">, 5-8</figureCitation>
<figureCitation id="A894F533D3CAA372CE66360DAC3EBC6A" captionStart="Figures 911" captionStartId="F4" captionText="Figures 9 - 11. Madagopsina apollo species group, dorsal view of wings 9 M. apollo, ♂, Ambohitra 10 M. makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 11 M. parvapollina, ♀, paratype, Ankarana. Scale bars: 0.5 mm. Figures 9, 11 (Feijen et al. 2018, figures 6, 7)." figureDoi="10.3897/zookeys.1057.67433.figures9-11" httpUri="https://binary.pensoft.net/fig/580796" pageId="0" pageNumber="1">, 9-11</figureCitation>
<figureCitation id="2CC1E6499F261805DC01BBFF18FBE80F" captionStart="Figures 1214" captionStartId="F5" captionText="Figures 12 - 14. Madagopsina apographica species group, dorsal view of wings 12 M. apographica, ♀, Fianarantsoa 13 M. freidbergi, ♀, paratype, Vohimana 14 M. tschirnhausi, ♂, holotype, Mount Ambre. Scale bars 0.5 mm. Figures 12 - 14 (Feijen et al. 2018, figures 8 - 10)." figureDoi="10.3897/zookeys.1057.67433.figures12-14" httpUri="https://binary.pensoft.net/fig/580797" pageId="0" pageNumber="1">, 12-14</figureCitation>
<figureCitation id="6E9EB7487F7C0814ACBCA16FC073637F" captionStart="Figures 1517" captionStartId="F6" captionText="Figures 15 - 17. Madagopsina makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 15 basal section of abdomen with intersternite 1 - 2, ventral view 16 sternites 5 and 6, ventral view 17 synsternite 7 + 8, ventral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1057.67433.figures15-17" httpUri="https://binary.pensoft.net/fig/580798" pageId="0" pageNumber="1">, 15-17</figureCitation>
<figureCitation id="78C8766E9F460C86B6F1F2667E3EB4D1" captionStart="Figures 1822" captionStartId="F7" captionText="Figures 18 - 22. Madagopsina makayensis Feijen, Feijen &amp; Feijen, sp. nov., ♂, holotype, Makay 18 epandrium with surstyli and cerci, posterior view 19 surstylus, inner view 20 hypandrial clasper, lateral view 21 phallapodeme and phallus, lateral view 22 ejaculatory apodeme and sac. Scale bars: 0.5 mm (18, 21); 0.1 mm (19, 20, 22)." figureDoi="10.3897/zookeys.1057.67433.figures18-22" httpUri="https://binary.pensoft.net/fig/580799" pageId="0" pageNumber="1">, 18-22</figureCitation>
<figureCitation id="6404F117677786CFDF592D0D4910B9B5" captionStart="Figures 2328" captionStartId="F8" captionText="Figures 23 - 28. Posterior view of Madagopsina surstyli 23 - 25 M. apollo species group 26 - 28 M. apographica species group 23 M. apollo 24 M. makayensis Feijen, Feijen &amp; Feijen, sp. nov. 25 M. parvapollina 26 M. apographica 27 M. freidbergi 28 M. tschirnhausi. Scale bar 0.1 mm (all drawn to the same scale). Figures 25 - 28 (Feijen et al. 2018, figures 187, 184, 186, 188)." figureDoi="10.3897/zookeys.1057.67433.figures23-28" httpUri="https://binary.pensoft.net/fig/580800" pageId="0" pageNumber="1">, 23-28</figureCitation>
<figureCitation id="FC9462BA07D9479C05F59A12C6B5B846" captionStart="Figure 29" captionStartId="F9" captionText="Figure 29. Principal component analysis of wing venation morphometry for the two species of Gracilopsina and six species of Madagopsina: biplot of the first two PCA axes." figureDoi="10.3897/zookeys.1057.67433.figure29" httpUri="https://binary.pensoft.net/fig/580801" pageId="0" pageNumber="1">, 29</figureCitation>
<figureCitation id="0C8AD34686062926E9710130C1A46F3E" captionStart="Figure 30" captionStartId="F10" captionText="Figure 30. Cluster dendrogram for the Euclidian distance in wing morphometry PCA for the Gracilopsina and Madagopsina species using the complete clustering method. Branch labels give the approximately unbiased p - value (AU) and bootstrap probability (BP) values (%)." figureDoi="10.3897/zookeys.1057.67433.figure30" httpUri="https://binary.pensoft.net/fig/580802" pageId="0" pageNumber="1">, 30</figureCitation>
<figureCitation id="304BDA689BCD172E1FCFD8A94B90FDA6" captionStart="Figure 31" captionStartId="F11" captionText="Figure 31. Eye span plotted against body length for the three species in the Madagopsina apollo species group: M. apollo, M. parvapollina and M. makayensis Feijen, Feijen &amp; Feijen, sp. nov. Note the position of the single data point for the ♂ of the latter species, in line with the ♂ data points for M. parvapollina." figureDoi="10.3897/zookeys.1057.67433.figure31" httpUri="https://binary.pensoft.net/fig/580803" pageId="0" pageNumber="1">, 31</figureCitation>
</paragraph>
</subSubSection>
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<paragraph pageId="0" pageNumber="1">
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<subSubSection id="C7C8C1DD3D017CDA247956C39BAB9C66" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="AC218CD1D40D1704A50EDC9A22834E60" pageId="0" pageNumber="1">
<taxonomicName id="CEA73EE223BCB7B4479109D9E0A0DDE8" authorityName="Feijen, Feijen &amp; Feijen" authorityYear="2018" class="Insecta" family="Diopsidae" genus="Madagopsina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagopsina" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Madagopsina</taxonomicName>
Feijen, Feijen &amp; Feijen, 2018:145. Type species
<taxonomicName authorityName="Brunetti" authorityYear="1928" class="Insecta" family="Diopsidae" genus="Diopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diopsis apollo" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="apollo">Diopsis apollo</taxonomicName>
<taxonomicName id="C76CE9EB5F9DA1F7DAC40D71D6187E38" authorityName="Brunetti" authorityYear="1928" class="Insecta" family="Diopsidae" genus="Diopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diopsis apollo" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="apollo">Diopsis apollo</taxonomicName>
Brunetti, 1928.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Frey" authorityYear="1928" class="Insecta" family="Diopsidae" genus="Eurydiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurydiopsis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Eurydiopsis</taxonomicName>
<paragraph id="C5FE12B86C6921BDFEB33AA410FF9B62" pageId="0" pageNumber="1">
<taxonomicName id="890EF51CF96E3728D4612C2F63640334" authorityName="Frey" authorityYear="1928" class="Insecta" family="Diopsidae" genus="Eurydiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eurydiopsis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Eurydiopsis</taxonomicName>
sensu
<normalizedToken originalValue="Séguy">Seguy</normalizedToken>
<normalizedToken id="AAE72FA192DD4AF765E265A49C3F8450" originalValue="Séguy">Seguy</normalizedToken>
&amp; Vanschuytbroeck (nec Frey) - in part;
<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1971.tb00763.x" author="Shillito, JF" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="287 - 295" refId="B14" refString="Shillito, JF, 1971. The genera of Diopsidae (Insecta: Diptera). Zoological Journal of the Linnean Society 50 (3): 287 - 295, DOI: https://doi.org/10.1111/j.1096-3642.1971.tb00763.x" title="The genera of Diopsidae (Insecta: Diptera)." url="https://doi.org/10.1111/j.1096-3642.1971.tb00763.x" volume="50" year="1971">Shillito 1971</bibRefCitation>
<bibRefCitation id="F5FCBF127FECFDC65624B78936C3DCEF" DOI="https://doi.org/10.1111/j.1096-3642.1971.tb00763.x" author="Shillito, JF" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="287 - 295" refId="B14" refString="Shillito, JF, 1971. The genera of Diopsidae (Insecta: Diptera). Zoological Journal of the Linnean Society 50 (3): 287 - 295, DOI: https://doi.org/10.1111/j.1096-3642.1971.tb00763.x" title="The genera of Diopsidae (Insecta: Diptera)." url="https://doi.org/10.1111/j.1096-3642.1971.tb00763.x" volume="50" year="1971">Shillito 1971</bibRefCitation>
: 287;
<bibRefCitation author="Feijen, HR" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="1" pagination="465 - 482" publicationUrl="https://journals.co.za/content/annals/24/2/AJA03040798_549" refId="B5" refString="Feijen, HR, 1981. A review of Diopsina Curran, 1928 (Diptera: Diopsidae), with a note on Cyrtodiopsis. Annals of the Natal Museum 24: 465 - 482, https://journals.co.za/content/annals/24/2/AJA03040798_549" title="A review of Diopsina Curran, 1928 (Diptera: Diopsidae), with a note on Cyrtodiopsis." url="https://journals.co.za/content/annals/24/2/AJA03040798_549" volume="24" year="1981">Feijen 1981</bibRefCitation>
<bibRefCitation id="AFBD5B3EEA66E1B72149540B9A950C77" author="Feijen, HR" journalOrPublisher="Annals of the Natal Museum" pageId="0" pageNumber="1" pagination="465 - 482" publicationUrl="https://journals.co.za/content/annals/24/2/AJA03040798_549" refId="B5" refString="Feijen, HR, 1981. A review of Diopsina Curran, 1928 (Diptera: Diopsidae), with a note on Cyrtodiopsis. Annals of the Natal Museum 24: 465 - 482, https://journals.co.za/content/annals/24/2/AJA03040798_549" title="A review of Diopsina Curran, 1928 (Diptera: Diopsidae), with a note on Cyrtodiopsis." url="https://journals.co.za/content/annals/24/2/AJA03040798_549" volume="24" year="1981">Feijen 1981</bibRefCitation>
: 482;
<bibRefCitation author="Feijen, HR" editor="Griffiths, GCD" journalOrPublisher="Schweizerbart, Stuttgart" pageId="0" pageNumber="1" pagination="1 - 122" refId="B6" refString="Feijen, HR, 1989. Diopsidae. Volume IX, Cyclorrhapha III (Schizophora other than Calyptratae), Part 12. In: Griffiths, GCD, Ed., Flies of the Nearctic Region. Schweizerbart, Stuttgart: 1 - 122" title="Diopsidae. Volume IX, Cyclorrhapha III (Schizophora other than Calyptratae), Part 12." volumeTitle="Flies of the Nearctic Region." year="1989">Feijen 1989</bibRefCitation>
<bibRefCitation id="331F2CFBF8D27E8727F8943557883C4F" author="Feijen, HR" editor="Griffiths, GCD" journalOrPublisher="Schweizerbart, Stuttgart" pageId="0" pageNumber="1" pagination="1 - 122" refId="B6" refString="Feijen, HR, 1989. Diopsidae. Volume IX, Cyclorrhapha III (Schizophora other than Calyptratae), Part 12. In: Griffiths, GCD, Ed., Flies of the Nearctic Region. Schweizerbart, Stuttgart: 1 - 122" title="Diopsidae. Volume IX, Cyclorrhapha III (Schizophora other than Calyptratae), Part 12." volumeTitle="Flies of the Nearctic Region." year="1989">Feijen 1989</bibRefCitation>
: 63;
<bibRefCitation DOI="https://doi.org/10.1163/22119434-00002029" author="Feijen, HR" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="1" pagination="161 - 189" refId="B8" refString="Feijen, HR, Feijen, C, 2013. A revision of the genus Diopsina Curran (Diptera, Diopsidae) with description of a new species from Guinea-Bissau. Tijdschrift voor Entomologie 156: 161 - 189, DOI: https://doi.org/10.1163/22119434-00002029" title="A revision of the genus Diopsina Curran (Diptera, Diopsidae) with description of a new species from Guinea-Bissau." url="https://doi.org/10.1163/22119434-00002029" volume="156" year="2013">Feijen and Feijen 2013</bibRefCitation>
<bibRefCitation id="4C7146A2E2D2AF5E22C2A8308F990890" DOI="https://doi.org/10.1163/22119434-00002029" author="Feijen, HR" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="1" pagination="161 - 189" refId="B8" refString="Feijen, HR, Feijen, C, 2013. A revision of the genus Diopsina Curran (Diptera, Diopsidae) with description of a new species from Guinea-Bissau. Tijdschrift voor Entomologie 156: 161 - 189, DOI: https://doi.org/10.1163/22119434-00002029" title="A revision of the genus Diopsina Curran (Diptera, Diopsidae) with description of a new species from Guinea-Bissau." url="https://doi.org/10.1163/22119434-00002029" volume="156" year="2013">Feijen and Feijen 2013</bibRefCitation>
: 182, 185.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="5AC4420535B628E2D2FD1299B718441F" pageId="0" pageNumber="1" type="remarks">
<paragraph id="EAA1064C109F38B0CFF0D29C13CAF666" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="1F1A4DEF22AE00CE950CC3351C57830E" pageId="0" pageNumber="1">
A concise catalogue for the genus is given below. For details on the type series, records, and combinations to various other genera of the earlier described species can be referred to
<bibRefCitation DOI="https://doi.org/10.1163/22119434-00002069" author="Feijen, HR" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="1" pagination="141 - 215" refId="B10" refString="Feijen, HR, Feijen, FAA, Feijen, C, 2018. Madagopsina gen. n. and Gracilopsina gen. n. (Diptera: Diopsidae) from Madagascar with description of four new species. Tijdschrift voor Entomologie 160 (3): 141 - 215, DOI: https://doi.org/10.1163/22119434-00002069" title="Madagopsina gen. n. and Gracilopsina gen. n. (Diptera: Diopsidae) from Madagascar with description of four new species." url="https://doi.org/10.1163/22119434-00002069" volume="160" year="2018">Feijen et al. (2018)</bibRefCitation>
<bibRefCitation id="24DC76EB2E8DD1B8CE7FEADC25F4045F" DOI="https://doi.org/10.1163/22119434-00002069" author="Feijen, HR" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="1" pagination="141 - 215" refId="B10" refString="Feijen, HR, Feijen, FAA, Feijen, C, 2018. Madagopsina gen. n. and Gracilopsina gen. n. (Diptera: Diopsidae) from Madagascar with description of four new species. Tijdschrift voor Entomologie 160 (3): 141 - 215, DOI: https://doi.org/10.1163/22119434-00002069" title="Madagopsina gen. n. and Gracilopsina gen. n. (Diptera: Diopsidae) from Madagascar with description of four new species." url="https://doi.org/10.1163/22119434-00002069" volume="160" year="2018">Feijen et al. (2018)</bibRefCitation>
. Reference is now made to new records which appeared after this publication. The new species
<taxonomicName authorityName="Feijen, Feijen &amp; Feijen" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Madagopsina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Madagopsina makayensis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makayensis">
<emphasis italics="true" pageId="0" pageNumber="1">Madagopsina makayensis</emphasis>
<taxonomicName id="ACE68FA2765EC64DB0FE00F063EBF13D" authorityName="Feijen, Feijen &amp; Feijen" authorityYear="2021" class="Insecta" family="Diopsidae" genus="Madagopsina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagopsina makayensis" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="makayensis">
<emphasis id="E84E7BBA35DF53AC72C2CF45E1699325" italics="true" pageId="0" pageNumber="1">Madagopsina makayensis</emphasis>
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Feijen, Feijen &amp; Feijen, sp. nov. is added.
</paragraph>

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<paragraph pageId="11" pageNumber="36">
<taxonomicName authority="Gerstaecker, 1868" authorityName="Gerstaecker" authorityYear="1868" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="11" pageNumber="36" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis Gerstaecker, 1868</taxonomicName>
<treatment id="4D0A3636ADEAFF23B61BB777ABF70091" ID-GBIF-Taxon="159360973" LSID="urn:lsid:plazi:treatment:4D0A3636ADEAFF23B61BB777ABF70091" httpUri="http://treatment.plazi.org/id/4D0A3636ADEAFF23B61BB777ABF70091" lastPageId="16" lastPageNumber="41" pageId="11" pageNumber="36" scope_class="Insecta" scope_family="Mydidae" scope_order="Diptera">
<subSubSection id="B221FC7190B9D799A6B1E6BF64745F7C" pageId="11" pageNumber="36" type="nomenclature">
<paragraph id="BD775B3EC35EDFF336F8C758D802CDF2" pageId="11" pageNumber="36">
<taxonomicName id="EE35BCC44FBD7E34E63BDD6F9AD57372" ID-CoL="38RWS" authority="Gerstaecker, 1868" authorityName="Gerstaecker" authorityYear="1868" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="11" pageNumber="36" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis Gerstaecker, 1868</taxonomicName>
Figs 20-22, 29313745
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="37" pageId="11" pageNumber="36" type="reference_group">
<paragraph pageId="11" pageNumber="36">
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="11" pageNumber="36" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<bibRefCitation author="Gerstaecker, A" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="26" pageNumber="51" pagination="65 - 103" title="Systematische Uebersicht der bis jetzt bekannt gewordenen Mydaiden (Mydasii Latr.)" url="http://www.biodiversitylibrary.org/bibliography/8630" year="1868">Gerstaecker 1868</bibRefCitation>
<subSubSection id="02A32181184157948893B12DE0F5235C" lastPageId="12" lastPageNumber="37" pageId="11" pageNumber="36" type="reference_group">
<paragraph id="6FD0A5375FCF5236240786F978FF527A" pageId="11" pageNumber="36">
<taxonomicName id="F1E2B08AB8B5FC3A1ED33F12982E65A6" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="11" pageNumber="36" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<bibRefCitation id="3277B8C9F11B1D254AE2B71A65646748" author="Gerstaecker, A" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="26" pageNumber="51" pagination="65 - 103" title="Systematische Uebersicht der bis jetzt bekannt gewordenen Mydaiden (Mydasii Latr.)" url="http://www.biodiversitylibrary.org/bibliography/8630" year="1868">Gerstaecker 1868</bibRefCitation>
: 92;
<bibRefCitation author="Bezzi, M" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="191 - 234" title="The South African Mydaidae (Diptera) as represented in the South African Museum" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="19" year="1924">Bezzi 1924</bibRefCitation>
<bibRefCitation id="5890BA94789D07E649DA8BBEB7A8EF0A" author="Bezzi, M" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="191 - 234" title="The South African Mydaidae (Diptera) as represented in the South African Museum" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="19" year="1924">Bezzi 1924</bibRefCitation>
: 196;
<bibRefCitation author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
<bibRefCitation id="811E30EF925EDB796C048EB47A38A604" author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
: 369;
<bibRefCitation author="Bowden, J" editor="Crosskey, RW" journalOrPublisher="British Museum of Natural History, London" pageId="26" pageNumber="51" pagination="325 - 333" title="27. Family Mydidae" volumeTitle="Catalogue of the Diptera of the Afrotropical Region" year="1980">Bowden 1980</bibRefCitation>
<bibRefCitation id="B5853D73705171DF57A7FC4F6C466FBD" author="Bowden, J" editor="Crosskey, RW" journalOrPublisher="British Museum of Natural History, London" pageId="26" pageNumber="51" pagination="325 - 333" title="27. Family Mydidae" volumeTitle="Catalogue of the Diptera of the Afrotropical Region" year="1980">Bowden 1980</bibRefCitation>
: 326.
</paragraph>
<paragraph pageId="12" pageNumber="37">
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="pinguis">
<pageBreakToken pageId="12" pageNumber="37" start="start">Ectyphus</pageBreakToken>
<paragraph id="C50F6E83ECB33A687DD77DA7916E1502" pageId="12" pageNumber="37">
<taxonomicName id="A657D0A5D6B47C4FA1BB4453E7DD8D8A" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="pinguis">
<pageBreakToken id="45A737E40760BEB4F73ECCBC392F4576" pageId="12" pageNumber="37" start="start">Ectyphus</pageBreakToken>
pinguis
</taxonomicName>
var. litoralis
<bibRefCitation author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
<bibRefCitation id="E546EE4CB3B452DC9E4314001AFEFBA1" author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
: 372. unavailable name
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="74C33474B0E43DCD7F3D4E66742715C8" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. karooensis
<bibRefCitation author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
<bibRefCitation id="842841259FDFFE41CE456B701BC0877C" author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
: 374. unavailable name
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="B82BAD0B34FCEBE3B9A2F47566C9A8F4" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. ceramiiformis
<bibRefCitation author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
<bibRefCitation id="E44F778410E52CEA08C312DAB9899AAE" author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
: 375. unavailable name
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus bitaeniatus" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="bitaeniatus">Ectyphus bitaeniatus</taxonomicName>
<bibRefCitation author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
<taxonomicName id="2FF71178DFAE7F02B0B0CAD671A012E1" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus bitaeniatus" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="bitaeniatus">Ectyphus bitaeniatus</taxonomicName>
<bibRefCitation id="EB731B50B326508CA2ECB7CEBDD7B1C8" author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
: 380. syn. n.
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
<bibRefCitation author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
<taxonomicName id="F9FCC2F6CD57E7305BCCD748389339BC" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="12" pageNumber="37" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
<bibRefCitation id="F5A2EEF7503C90C3F5432F6C5A0D4829" author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse 1969</bibRefCitation>
: 378. syn. n.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="37" type="diagnosis">
<paragraph pageId="12" pageNumber="37">Diagnosis:</paragraph>
<paragraph pageId="12" pageNumber="37">The species is distinguished from congeners by the enlarged yellow facial gibbosity, the yellow posterior margin of the abdominal tergites that are widened laterally and interrupted medially, and the distinctly yellow metepimeron.</paragraph>
<subSubSection id="7235A3A987AE1B82C3225E1B17326C50" pageId="12" pageNumber="37" type="diagnosis">
<paragraph id="04204F71C28112A5BDC85D9E423CB8DF" pageId="12" pageNumber="37">Diagnosis:</paragraph>
<paragraph id="F031EB16EFD67C3A40500BE381FF2D63" pageId="12" pageNumber="37">The species is distinguished from congeners by the enlarged yellow facial gibbosity, the yellow posterior margin of the abdominal tergites that are widened laterally and interrupted medially, and the distinctly yellow metepimeron.</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="39" pageId="12" pageNumber="37" type="re-description male">
<paragraph pageId="12" pageNumber="37">Re-description male:</paragraph>
<paragraph pageId="12" pageNumber="37">
<subSubSection id="A95D1B3075A01563DF254155F4EB9480" lastPageId="14" lastPageNumber="39" pageId="12" pageNumber="37" type="re-description male">
<paragraph id="8176BF9EF56149DD5C14843CD4BBB40E" pageId="12" pageNumber="37">Re-description male:</paragraph>
<paragraph id="31D49E21F41763DFFCD3A5C1AFE3EBFE" pageId="12" pageNumber="37">
Head: black, facial gibbosity yellow, in general lightly silver pubescent; width distinctly greater than thorax, interocular distance on vertex larger than at ventral eye margin, vertex between compound eyes slightly depressed, parafacial area very narrow, facial gibbosity nearly touching median eye margin; facial gibbosity distinct, well-developed and discernible in lateral view; mystax white, covering only lateral facial gibbosity (asetose medially); frons medially apubescent, laterally grey pubescent, vertex apubescent, postgena lightly silver pubescent; setation: vertex white, frons white, ocp setae white, pocl setae white; ocellar triangle apubescent; proboscis light brown, short, about
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="3F9BD1CEE0E6364B7C8A322CB6DB70DF" originalValue="½">1/2</normalizedToken>
length of oral cavity; labellum small, as wide as prementum, as long as prementum, unsclerotised laterally; maxillary palpus cylindrical, brown, longer than
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="B75D6C0071080DF6537B992099BCAACF" originalValue="½">1/2</normalizedToken>
length of proboscis.
</paragraph>
<paragraph pageId="12" pageNumber="37">
<paragraph id="20F114F89A5EB150C8F2D57A0DAAA118" pageId="12" pageNumber="37">
Antenna: brown, scape and pedicel white and yellow setose dorsally and ventrally; postpedicel cylindrical in proximal
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="A50E297C0E203ADFFFDDBE04248288CC" originalValue="½">1/2</normalizedToken>
, symmetrically bulbous in distal
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="C2DAF406658C4F7A84A8349199757C3E" originalValue="½">1/2</normalizedToken>
, ≥ 8.0 times as long as combined length of scape and pedicel; apical
<normalizedToken originalValue="seta-like">'seta-like'</normalizedToken>
<normalizedToken id="E8E4D07B417774A6310D4F1F950C6B56" originalValue="seta-like">'seta-like'</normalizedToken>
sensory element situated apically in cavity on postpedicel.
</paragraph>
<paragraph pageId="12" pageNumber="37">Thorax: dark brown to bluish-black, predominantly grey pubescent; scutum medially brown, laterally dark yellow, surface entirely smooth, lightly grey pubescent, scutal setation comprised of distinct rows of short dorsocentral setae and lateral scutal setae; dc setae pre- and postsuturally white, acr setae absent, lateral scutal setae white, npl, spal, and pal setae absent; postpronotal lobe yellow, partly white pubescent; proepisternum, lateral postpronotum, and postpronotal lobe long white setose; scutellum apubescent, asetose medially, laterally yellow setose, apical scutellar setae absent; mesopostnotum, anatergite, and katatergite silver pubescent, asetose; katatergite elevated and smoothly convex; anterior anepisternum asetose, supero-posterior anepisternum asetose; posterior anepimeron long white setose, katepimeron asetose; metepimeron evenly elevated, yellow, lightly silver pubescent, asetose; metepisternum silver pubescent, asetose.</paragraph>
<paragraph pageId="12" pageNumber="37">
<paragraph id="89FB02A301C79B48744BF88D91F9108B" pageId="12" pageNumber="37">Thorax: dark brown to bluish-black, predominantly grey pubescent; scutum medially brown, laterally dark yellow, surface entirely smooth, lightly grey pubescent, scutal setation comprised of distinct rows of short dorsocentral setae and lateral scutal setae; dc setae pre- and postsuturally white, acr setae absent, lateral scutal setae white, npl, spal, and pal setae absent; postpronotal lobe yellow, partly white pubescent; proepisternum, lateral postpronotum, and postpronotal lobe long white setose; scutellum apubescent, asetose medially, laterally yellow setose, apical scutellar setae absent; mesopostnotum, anatergite, and katatergite silver pubescent, asetose; katatergite elevated and smoothly convex; anterior anepisternum asetose, supero-posterior anepisternum asetose; posterior anepimeron long white setose, katepimeron asetose; metepimeron evenly elevated, yellow, lightly silver pubescent, asetose; metepisternum silver pubescent, asetose.</paragraph>
<paragraph id="EB5C4FABC462ECF1FADE5169F5E26950" pageId="12" pageNumber="37">
Leg: light brown, setation predominantly white; pro, mes, and met coxa apubescent, long white setose; met trochanter macrosetose medially; femur brown, met femur evenly clubbed in distal
<normalizedToken originalValue="¾">3/4</normalizedToken>
<normalizedToken id="4F2ADAFFC1C7E8E71BB724BE7BCDAED8" originalValue="¾">3/4</normalizedToken>
, in distal
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="535EEA41E43603F42C808CDD69E84AB5" originalValue="½">1/2</normalizedToken>
macrosetose, 1 antero-ventral and 1 postero-ventral row of macrosetae; pro, mes, and met tibia straight, met tibia cylindrical with distinct ventral keel terminating into a sharp spine; pro and mes tarsomere 1 longer than tarsomere 2, but less than combined length of tarsomeres 2-3, met tarsomere 1 as long as combined length of tarsomeres 2-3; pulvillus well-developed, as long as well-developed claw, and as wide as base of claw; empodium absent.
</paragraph>
<paragraph lastPageId="14" lastPageNumber="39" pageId="13" pageNumber="38">
<pageBreakToken pageId="13" pageNumber="38" start="start">Wing</pageBreakToken>
<paragraph id="612DC784C37CC6B13329C51369D64869" lastPageId="14" lastPageNumber="39" pageId="13" pageNumber="38">
<pageBreakToken id="D0AB2D865A92FAB1DBEA4B053693F4E6" pageId="13" pageNumber="38" start="start">Wing</pageBreakToken>
: length = 10.2
<normalizedToken originalValue="13.3(">-13.3(-</normalizedToken>
<normalizedToken id="0F90C4058958BCF5E7FA79B8485CC6E9" originalValue="13.3(">-13.3(-</normalizedToken>
14.2) mm; hyaline throughout, slightly brown stained along veins, veins light brown, microtrichia absent; cells r1, r4, r5, m3, + cup closed; C well-developed, around entire wing; R4 terminates in R1; R5 terminates in R1; stump vein (R3) at base of R4 present, long but not reaching R2; R4 and R5 widest apart med
<pageBreakToken pageId="14" pageNumber="39" start="start">ially</pageBreakToken>
<pageBreakToken id="0991BAF67377CFB0EA5C549FB1C4F8CB" pageId="14" pageNumber="39" start="start">ially</pageBreakToken>
; r-m distinct, R4+5 and M1 apart, connected by crossvein; M1 straight at r-m (not curving anteriorly), M1 (or M1+M2) terminates in C; CuA1 and CuA2 split proximally to m-cu (cell m3 narrow proximally); M3+CuA1 terminate together in C; A1 undulating, cell a1 wide, A1 and wing margin further apart proximally than distally; alula well-developed; halter light yellow.
</paragraph>
<paragraph pageId="14" pageNumber="39">Abdomen: brown; setation comprised of scattered white setae, surface entirely smooth; T1 brown, narrow yellow posterior margin, T2-7 brown, broad yellow posterior margin, expanding antero-laterally particularly on T2-3; T1 long white setose, T2-3 sparsely white setose; T predominantly apubescent; S1 yellow, S2-7 yellow with brown areas medially and laterally; S1-3 asetose; S predominantly apubescent; T2-4 parallel-sided and not constricted waist-like; bullae on T2 black, transversely elongate, surface entirely smooth, T2 surface anterior to bullae smooth.</paragraph>
<paragraph pageId="14" pageNumber="39">
<paragraph id="4274732030AD672F1DEA3935DB54F3C4" pageId="14" pageNumber="39">Abdomen: brown; setation comprised of scattered white setae, surface entirely smooth; T1 brown, narrow yellow posterior margin, T2-7 brown, broad yellow posterior margin, expanding antero-laterally particularly on T2-3; T1 long white setose, T2-3 sparsely white setose; T predominantly apubescent; S1 yellow, S2-7 yellow with brown areas medially and laterally; S1-3 asetose; S predominantly apubescent; T2-4 parallel-sided and not constricted waist-like; bullae on T2 black, transversely elongate, surface entirely smooth, T2 surface anterior to bullae smooth.</paragraph>
<paragraph id="9507FA8D22F643F21E4CF3A23877C4AC" pageId="14" pageNumber="39">
Male terminalia: T1-7 well-developed, entirely sclerotised, T8 postero-medially weakly sclerotised, with anterior transverse sclerotised bridge connecting lateral sclerites; T7-8 anteriorly with 2 lateral apodemes; S6 regular, without any special setation postero-medially, S8 well-developed and simple, fused to T8 dorso-laterally, entire (undivided) ventro-medially; epandrium formed by single sclerite (fused medially
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="DF7D47B91707B444D7841117ACFEFCF7" originalValue="±">+/-</normalizedToken>
entirely), pointed postero-laterally; subepandrial sclerite without lateral or median protuberances; hypandrium
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="716EE36277EC9E0BEE5E70A0CBE70269" originalValue="±">+/-</normalizedToken>
flat, rectangular to square sclerite, entirely fused with gonocoxite, forming a gonocoxite-hypandrial complex; gonocoxite dorso-ventrally flattened in distal
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="1BCF49D74A0AEE179314CEF49BF11CDF" originalValue="½">1/2</normalizedToken>
, higher in proximal
<normalizedToken originalValue="½">1/2</normalizedToken>
<normalizedToken id="F3E177D43F08C438A346F6F0D5C1C683" originalValue="½">1/2</normalizedToken>
, with palp-like lateral appendage, gonocoxal apodeme present, short (at most slightly extending hypopygium anteriorly); 1 functional aedeagal prong, aedeagal epimere absent; lateral ejaculatory process absent; ejaculatory apodeme formed by single dorso-ventrally oriented plate; ventro-median margin of dorsal aedeagal sheath heavily sclerotised (appearing entirely closed); dorsal aedeagal sheath long, sperm sac entirely covered; sperm sac appearing
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="9BCE8B5F6BEA4EF906981532BAC85EDA" originalValue="±">+/-</normalizedToken>
heavily sclerotised.
</paragraph>
<caption pageId="14" pageNumber="39">
<paragraph pageId="14" pageNumber="39">
<caption id="5F5ADE493F15ADCFA923BE82C16666EC" pageId="14" pageNumber="39">
<paragraph id="AF0F783908C4EFDA94045BEEF6D320AA" pageId="14" pageNumber="39">
Figures 20-30. Male terminalia and female genitalia of
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="genus">Ectyphus</taxonomicName>
<taxonomicName id="1B78DD8CA7DCADD9A7F415D77406DECF" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="genus">Ectyphus</taxonomicName>
and
<taxonomicName class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="genus">Parectyphus</taxonomicName>
<taxonomicName id="B3B7843CDCBACE199B5FC3F192BDDBDF" class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="genus">Parectyphus</taxonomicName>
species. Male terminalia: 20-22
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="E5C78CA508CAB48DAF7D6435F5B3EEA7" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
20 lateral 21 dorsal 22 ventral; 23-25
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pretoriensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pretoriensis">Ectyphus pretoriensis</taxonomicName>
<taxonomicName id="8381DC2AC052CC47B62F12EF0EE11DBC" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pretoriensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pretoriensis">Ectyphus pretoriensis</taxonomicName>
23 lateral 24 dorsal 25 ventral; 26-28
<taxonomicName class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus namibiensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="namibiensis">Parectyphus namibiensis</taxonomicName>
<taxonomicName id="4A8D75FB9A6A9C42854D6F6FF67554D2" class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus namibiensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="namibiensis">Parectyphus namibiensis</taxonomicName>
26 lateral 27 dorsal 28 ventral; Female genitalia in dorsal view: 29
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="944FBEDE342BE47F1C6ABA3B277A58E2" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
30
<taxonomicName class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus namibiensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="namibiensis">Parectyphus namibiensis</taxonomicName>
<taxonomicName id="12C3991C9C12807D020939E5F730582B" class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus namibiensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="namibiensis">Parectyphus namibiensis</taxonomicName>
. Scale lines = 1 mm.
</paragraph>
</caption>
<caption pageId="14" pageNumber="39">
<paragraph pageId="14" pageNumber="39">
<caption id="BF4D85769A950BF99D4FB36FDC21B2FE" pageId="14" pageNumber="39">
<paragraph id="608136368CEE870E9119033ED9E4DA65" pageId="14" pageNumber="39">
Figures 31-34. Photographs of
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="4BDB414DC71B76A15D9623ACA694619F" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
specimens. 31
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="486ADE8FB20C3E26CD4D2E215EFE9F66" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
(♂, AAM-003496, AMGS, Morphbank) 32
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="B776DEF51B0EAFB4F29723CEDE807C15" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
(♂, AAM-003487, SAMC, Morphbank) 33
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="68114A0269620F73BA8D8AEE0F9D499C" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
(♀, AAM-003509, SAMC, Morphbank) 34
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="BAA34E30AF0D07595A722DA9CA4AC247" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
(♂ holotype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="F9667179C9DA483C1C80877C14E28B5D" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. ceramiiformis, NMSA, Morphbank). Scale lines = 5 mm.
</paragraph>
</caption>
<caption pageId="14" pageNumber="39">
<paragraph pageId="14" pageNumber="39">
<caption id="0AD92EED0990D53C513EAE42DCDDD890" pageId="14" pageNumber="39">
<paragraph id="E3216E64105D36786A36C3CC7C330D40" pageId="14" pageNumber="39">
Figures 35-38. Photographs of
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="596A833DE2731025832E3D0AA0717897" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
specimens and wings of
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="462B9F3559DBDAE9DED08BCD688A8973" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
and
<taxonomicName class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus namibiensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="namibiensis">Parectyphus namibiensis</taxonomicName>
<taxonomicName id="0F6EE652737CBA961BBFF818CE551729" class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus namibiensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="namibiensis">Parectyphus namibiensis</taxonomicName>
. 35
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="080490DC2A55B035272EB60BB0BE9C18" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
(♀ holotype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus bitaeniatus" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="bitaeniatus">Ectyphus bitaeniatus</taxonomicName>
<taxonomicName id="46AEC875D5EBA46331DF855F13B77F14" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus bitaeniatus" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="bitaeniatus">Ectyphus bitaeniatus</taxonomicName>
, NMSA, Morphbank) 36
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="DA362D3B40EB07042C9967726ACCC8A0" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
(♀ holotype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
<taxonomicName id="7F8A3003ED01014B911838EF0375E100" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
, NMSA, Morphbank); Wings 37
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="3EB7AEA738E177147C683C03E41F3421" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
(AAM-003463, NMSA, Morphbank) 38
<taxonomicName class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus namibiensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="namibiensis">Parectyphus namibiensis</taxonomicName>
<taxonomicName id="93E87F675D8854935FBCE02AB4E0D84C" class="Insecta" family="Mydidae" genus="Parectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parectyphus namibiensis" order="Diptera" pageId="14" pageNumber="39" phylum="Arthropoda" rank="species" species="namibiensis">Parectyphus namibiensis</taxonomicName>
(AAM-003506, NMNW, Morphbank). Scale lines = 5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="14" pageNumber="39" type="re-description female">
<paragraph pageId="14" pageNumber="39">Re-description female:</paragraph>
<paragraph pageId="14" pageNumber="39">Head: brown, facial gibbosity yellow; mystax white, sparse short setae covering entire facial gibbosity; vertex predominantly apubescent, only lateral margin grey pubescent; maxillary palpus light brown.</paragraph>
<paragraph pageId="14" pageNumber="39">Thorax: brown; scutum surface entirely smooth, scutal setation comprised of distinct rows of long dorsocentral setae and lateral scutal setae.</paragraph>
<paragraph pageId="14" pageNumber="39">Leg: femur light brown.</paragraph>
<paragraph pageId="14" pageNumber="39">Wing: length = 12.0-13.7 mm.</paragraph>
<paragraph pageId="14" pageNumber="39">
<subSubSection id="0954D85E77ABE557CCB0548E86E294B6" pageId="14" pageNumber="39" type="re-description female">
<paragraph id="671BC010125F31A1220AB1FE8F87E4E2" pageId="14" pageNumber="39">Re-description female:</paragraph>
<paragraph id="DE20F402E7F69F0E1DE7C9712C3ED5D3" pageId="14" pageNumber="39">Head: brown, facial gibbosity yellow; mystax white, sparse short setae covering entire facial gibbosity; vertex predominantly apubescent, only lateral margin grey pubescent; maxillary palpus light brown.</paragraph>
<paragraph id="D2CEF270CE0651969A5FE423C406DA16" pageId="14" pageNumber="39">Thorax: brown; scutum surface entirely smooth, scutal setation comprised of distinct rows of long dorsocentral setae and lateral scutal setae.</paragraph>
<paragraph id="2CD9141201B509DEF99802495BB0F1A2" pageId="14" pageNumber="39">Leg: femur light brown.</paragraph>
<paragraph id="50EDE2F349257DE406596550F1CAC281" pageId="14" pageNumber="39">Wing: length = 12.0-13.7 mm.</paragraph>
<paragraph id="29541B649F2868F04E0372B7F7497F5D" pageId="14" pageNumber="39">
Female genitalia: densely arranged anteriorly directed setae absent, only few on T7-8 and S7-8; T8 with broad anterior rectangular apodeme; T9 formed by wide, rectangular sclerite with median protuberance; T9+10 entirely fused, T10 divided into 2 heavily sclerotised acanthophorite plates, 11 acanthophorite spurs per plate; 3 spermathecae, all equally large, formed by
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="38668E4BD19E8E1EF6112F7538C52745" originalValue="±">+/-</normalizedToken>
expanded weakly sclerotised ducts; individual spermathecal duct long; S9 (furca) formed by 2 sclerites, separated anteriorly and posteriorly, anterior furcal apodeme present, 2 lateral projections forming divided apodeme, lateral furcal apodeme absent, median furcal bridge absent.
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="41" pageId="14" pageNumber="39" type="material examined">
<paragraph pageId="14" pageNumber="39">Material examined:</paragraph>
<paragraph lastPageId="16" lastPageNumber="41" pageId="14" pageNumber="39">
<subSubSection id="E28F03FA5E16CFED39063C7979DEB8D5" lastPageId="16" lastPageNumber="41" pageId="14" pageNumber="39" type="material examined">
<paragraph id="40E609A31EE6370666CB3C10D50076A8" pageId="14" pageNumber="39">Material examined:</paragraph>
<paragraph id="B3E4CCD4C171C59D120C0D06FEB435BB" lastPageId="16" lastPageNumber="41" pageId="14" pageNumber="39">
South Africa: 1♂ 'Africa
<normalizedToken originalValue="australis">australis'</normalizedToken>
<normalizedToken id="E78B2A483966C892B67EDC01965733F7" originalValue="australis">australis'</normalizedToken>
,
<normalizedToken originalValue="Drège">Drege</normalizedToken>
<normalizedToken id="37346CF8507414C18398B882D09A9F66" originalValue="Drège">Drege</normalizedToken>
(lectotype, ZMHB); 1
<normalizedToken originalValue="F">F'</normalizedToken>
<normalizedToken id="808B6171D3F3079C74024B1D0DA4D0CC" originalValue="F">F'</normalizedToken>
Capland,
<normalizedToken originalValue="Drège">Drege</normalizedToken>
<normalizedToken id="0F64A6DBA59EB1095FA33E4F72867C8F" originalValue="Drège">Drege</normalizedToken>
(paralectotype, ZMHB); Eastern Cape Province: 1♂ Cuylerville,
<pageBreakToken pageId="15" pageNumber="40" start="start">31</pageBreakToken>
<pageBreakToken id="8CD00CC352EA959782A4341EC4F7A7EA" pageId="15" pageNumber="40" start="start">31</pageBreakToken>
°47'00&quot;S; 26°56'00&quot;E, 20.xii.1920, H. Cronwright (AAM-003522, SAMC); 1♀ Resolution, 33°10'00&quot;S; 26°37'00&quot;E, 23.xii.1927, A. Walton (holotype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus bitaeniatus" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="bitaeniatus">Ectyphus bitaeniatus</taxonomicName>
<taxonomicName id="FD171DBBC794AAFED1D3400F90F052B0" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus bitaeniatus" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="bitaeniatus">Ectyphus bitaeniatus</taxonomicName>
, NMSA); 1♂ Willowmore, 33°17'00&quot;S; 23°29'00&quot;E, -.-.-, J. Brauns (holotype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="C1A7D780BC9A50EC85F6D0D684C669A8" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. ceramiiformis, NMSA); 2♂ Willowmore, -.-.-, J. Brauns (holotype and paratype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="A04128CAB72FD8D8CAAF5296093281D8" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. karooensis, NMSA); 1♀ Willowmore, -.-.-, J. Brauns (AAM-003492, USNM); 1♀ Willowmore, 20.xii.1909, J. Brauns (paratype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
<taxonomicName id="1BBFBCD395D1F9324A8A79131FC5FADD" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
, NMSA); 1♀ Albany District, 33°24'00&quot;S; 26°32'00&quot;E, -.xii.1949, R. Phillips (AAM-003473, BMNH); 1♀ Georgida, 33°27'00&quot;S; 23°17'00&quot;E, 5.i.1927, J. Brauns (holotype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
<taxonomicName id="044C5FF828CD484BEF0AC8BE0EDE3ADD" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
, NMSA); 1♀ 1♂ Dunbrody, 33°28'00&quot;S; 25°33'00&quot;E, -.-.1897,
<normalizedToken originalValue="ONeil">O'Neil</normalizedToken>
<normalizedToken id="17B0785EC40F793ED538D3AF32444A97" originalValue="ONeil">O'Neil</normalizedToken>
(AAM-003509-AAM-003510, SAMC); 1♂ Kleinemonde, 33°31'39&quot;S; 27°03'00&quot;E, -.-.-, J. Cooper (AAM-003487, SAMC); 2♂ Kleinemonde, -.i.1891, M. White (AAM-003520-AAM-003521, SAMC); 1♀ Port Alfred, 33°36'00&quot;S; 26°54'00&quot;E, 17.ii.1955, F. Junor (paratype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="C893107126EB909E0C36BA523C581A58" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="15" pageNumber="40" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. litoralis, SAMC); 1♂ Port Alfred, 9.i.1971, J. Londt (AAM-003503, AMGS); 1♀ Port Alfred (AAM-003508, SAMC); 1♀ 2♂ Riet River mouth, 33°36'00&quot;S; 26°54'00&quot;E, 17.xii.1971, D. Greathead (AAM-003470-AAM-003472, BMNH); 1♀ 3♂ Rietrivier mouth, near Port Alfred, 17.xii.1974, F. Gess (AAM-003493-AAM-003496, AMGS); 5♂ Rietrivier mouth, near Port Alfred, 29.xii.1973, F. Gess (AAM-003497-AAM-003501, AMGS); 2♂ Algoa Bay, 33°41'34&quot;S; 25°56'39&quot;E, 15.xii.1892, J. Brauns
<pageBreakToken pageId="16" pageNumber="41" start="start">(</pageBreakToken>
<pageBreakToken id="ACE0E816F35DFB64BC8F46094BE91CC7" pageId="16" pageNumber="41" start="start">(</pageBreakToken>
ZSMC); 2♀ 2♂ Swartkops, Algoa Bay, 33°51'49&quot;S; 25°36'06&quot;E, 25.xi.1921, J. Brauns (AAM-003447, AAM-003459-AAM-003461, NMSA); 1♂ Swartkops, Algoa Bay, 25.xi.1921, J. Brauns (AAM-003466, MZLU); 1♂ Swartkops, Algoa Bay, -.i.1919, B.
<normalizedToken originalValue="Krüger">Krueger</normalizedToken>
<normalizedToken id="50BC9E9C9F3E732370569B05D93D5758" originalValue="Krüger">Krueger</normalizedToken>
(DEIC); 3♀ Swartkops, Port Elizabeth, 20.xi.1919, B.
<normalizedToken originalValue="Krüger">Krueger</normalizedToken>
<normalizedToken id="BAF65A577FE5C43809BADAF66ACE1C33" originalValue="Krüger">Krueger</normalizedToken>
(AAM-003448, AAM-003462-AAM-03463, NMSA); 1♀ Swartkops, Port Elizabeth, -.ii.1919, B.
<normalizedToken originalValue="Krüger">Krueger</normalizedToken>
<normalizedToken id="6F6F0F7D2531539CE7636DF669F5CD71" originalValue="Krüger">Krueger</normalizedToken>
(AAM-003465, NMSA); 1♂ Swartkops, Port Elizabeth, 25.xi.1919, B.
<normalizedToken originalValue="Krüger">Krueger</normalizedToken>
<normalizedToken id="9E8E35B1AF1607B7DE4456C8D5D20545" originalValue="Krüger">Krueger</normalizedToken>
(AAM-003464, NMSA); 1♀ 1♂ Swartkops, Port Elizabeth, 25.xi.1919, B.
<normalizedToken originalValue="Krüger">Krueger</normalizedToken>
<normalizedToken id="62BEF45E0AD9E59EEECDC4023043F44C" originalValue="Krüger">Krueger</normalizedToken>
(AAM-003467-AAM-003468, ISNB); 4♀ 6♂ Gamtoos River Mouth, Papiesfontein, 33°57'47&quot;S; 25°01'46&quot;E, -.i.1960, SAM Museum Staff (paratype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="4D344C42E9CE4E807284E368D6375312" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. litoralis, SAMC); 1♂ Van
<normalizedToken originalValue="Stadens">Staden's</normalizedToken>
<normalizedToken id="0E08AE2974B6352E3C3FA9854EC4F0C4" originalValue="Stadens">Staden's</normalizedToken>
River Mouth, 33°58'00&quot;S; 25°13'00&quot;E, -.i.1960, SAM Museum Staff (paratype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="0F4F10C325BDBABC04534A924DA104D8" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. litoralis, SAMC); 1♀ 1♂ Port Elizabeth, 33°58'00&quot;S; 25°35'00&quot;E, 1.ii.1950, A. Brown (AAM-003450-AAM-003451, NMSA); 1♀ Port Elizabeth, 1.i.1971, M. Strydom (AAM-003034, SANC); 2♀ 3♂ Port Elizabeth, Cape Recife area, 34°01'14&quot;S; 25°40'60&quot;E, 22-27.xii.1985, J. Londt (AAM-003442-AAM-003446, NMSA); 1♂
<normalizedToken originalValue="Jeffreys">Jeffrey's</normalizedToken>
<normalizedToken id="D19D78C12A32EFB3ED6610C0DD3173C3" originalValue="Jeffreys">Jeffrey's</normalizedToken>
Bay, Humansdorp, 34°02'00&quot;S; 24°46'00&quot;E, 23.xii.1922, J. Brauns (AAM-003469, BMNH); 1♀ The Willows, Port Elizabeth, 34°02'00&quot;S; 25°36'00&quot;E, 28.xii.1970, M. Strydom (AAM-003489, SANC); 1♂
<normalizedToken originalValue="Jeffreys">Jeffrey's</normalizedToken>
<normalizedToken id="5BCBA66B856A7478DA6864553C877732" originalValue="Jeffreys">Jeffrey's</normalizedToken>
Bay, 34°03'00&quot;S; 24°55'00&quot;E, 19.xii.1922, (DEIC); 9♀ 3♂
<normalizedToken originalValue="Jeffreys">Jeffrey's</normalizedToken>
<normalizedToken id="0573D95D19E6FA553F38B91F4034C204" originalValue="Jeffreys">Jeffrey's</normalizedToken>
Bay, -.i.1960, SAM Museum Staff (holotype and paratype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="69AD7ADDF9F4F75EB75E8440E5B87D8A" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. litoralis, SAMC); 1♀ 1♂
<normalizedToken originalValue="Jeffreys">Jeffrey's</normalizedToken>
<normalizedToken id="EC988C52FC475253D6F4F06F226273FE" originalValue="Jeffreys">Jeffrey's</normalizedToken>
Bay, -.i.1960, SAM Museum Staff (paratype
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="247A51698D45A80E0FFE154292769A62" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. litoralis, ISNB); 1♀ 1♂
<normalizedToken originalValue="Jeffreys">Jeffrey's</normalizedToken>
<normalizedToken id="672CBB76AE0E6A12789F75B92828300F" originalValue="Jeffreys">Jeffrey's</normalizedToken>
Bay, -.-.-, (AAM-003828-AAM-003829, SMNS); Western Cape Province: 1♀ Tulbagh, 33°17'00&quot;S; 19°09'00&quot;E, 10.xii.1924, J. Brauns (AAM-003449, NMSA); 1♂ 'Cape of Good
<normalizedToken originalValue="Hope">Hope'</normalizedToken>
<normalizedToken id="CB58FF5B7D713E04D8E433FDA97AA174" originalValue="Hope">Hope'</normalizedToken>
, 33°48'03&quot;S; 19°00'36&quot;E, -.-.1835, J. Verreaux (MNHN).
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="41" type="type locality, distribution, and biodiversity hotspots">
<paragraph pageId="16" pageNumber="41">Type locality, distribution, and biodiversity hotspots:</paragraph>
<paragraph pageId="16" pageNumber="41">
<subSubSection id="F97166FF51734A116AB75294741AF61F" pageId="16" pageNumber="41" type="type locality, distribution, and biodiversity hotspots">
<paragraph id="37BE4B39495025D6A3F72B0F44C7CFA5" pageId="16" pageNumber="41">Type locality, distribution, and biodiversity hotspots:</paragraph>
<paragraph id="50C31009530C01DC5335F3A49CCE0868" pageId="16" pageNumber="41">
The original type locality is 'Africa
<normalizedToken originalValue="australis">australis'</normalizedToken>
<normalizedToken id="5746E0A1BD5F8D55ABF7F82BCD118BBB" originalValue="australis">australis'</normalizedToken>
(South Africa). Following recommendation 76A.1.4. of the International Code of Zoological Nomenclature (4th edition) a new type locality is selected from within the range of the species. We hereby designate the Riet River mouth (33°36'00&quot;S; 026°54'00&quot;E), near Port Alfred, Eastern Cape, South Africa as the new type locality. Cape Floristic Region and Maputaland-Pondoland-Albany biodiversity hotspots.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="41" type="remarks">
<paragraph pageId="16" pageNumber="41">Remarks:</paragraph>
<paragraph pageId="16" pageNumber="41">
<subSubSection id="72E05021EBC393E75E8F033422ABD7AA" pageId="16" pageNumber="41" type="remarks">
<paragraph id="9721E3752131ED88EBDB2AF5FBC10F92" pageId="16" pageNumber="41">Remarks:</paragraph>
<paragraph id="A6F1EEC8A69F6104F427586D12B91116" pageId="16" pageNumber="41">
Specimens identified as belonging to the three varieties of
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="9EAE3B55420D9B7CFEB3EE0508A7CBB5" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
were examined, including a large series of paratypes of
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="D24003880682F17EC85C9F2CE5D4660B" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. litoralis. These specimens were determined to represent colour and vestiture variation in
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="B89BA1CB10961AD9DC79DD0CDF466C33" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
, rather than belonging to distinct subspecies. The three names,
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="E33BCE68BA36DACE643220D9C5F5A101" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. litoralis,
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="069259807A8F5192B144A0B0F4BBCF0E" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. karooensis, and
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="38638E15E73AF428493E3C08D1F2369C" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
var. ceramiiformis, were proposed by Hesse to delimit infrasubspecific entities. Because these names were never adopted as valid for a species or subspecies, as was already pointed out by
<bibRefCitation author="Bowden, J" editor="Crosskey, RW" journalOrPublisher="British Museum of Natural History, London" pageId="26" pageNumber="51" pagination="325 - 333" title="27. Family Mydidae" volumeTitle="Catalogue of the Diptera of the Afrotropical Region" year="1980">Bowden (1980: 326)</bibRefCitation>
<bibRefCitation id="C920AB268B9467887630ACC139CA7545" author="Bowden, J" editor="Crosskey, RW" journalOrPublisher="British Museum of Natural History, London" pageId="26" pageNumber="51" pagination="325 - 333" title="27. Family Mydidae" volumeTitle="Catalogue of the Diptera of the Afrotropical Region" year="1980">Bowden (1980: 326)</bibRefCitation>
, the names are unavailable following the International Code of Zoological Nomenclature (4th edition, Article 45.6.4. and 45.6.4.1.). The unique female holotype of
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus bitaeniatus" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="bitaeniatus">Ectyphus bitaeniatus</taxonomicName>
<taxonomicName id="5D2C1FDF8B4CBC1FA1CE49B6664F0489" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus bitaeniatus" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="bitaeniatus">Ectyphus bitaeniatus</taxonomicName>
and the two female type specimens of
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
<taxonomicName id="2B1E77E74527C6A20008E7F52A25AC29" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus flavidorsalis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="flavidorsalis">Ectyphus flavidorsalis</taxonomicName>
were also examined. These specimens represent
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="83EE24AF6AF493374E15A77F92FE8559" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
in our view and are here synonymised with this species. Similar colour variation as exhibited by these two species was observed in female specimens of
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="0A1D90E943B72FEAAAF1B8A048B76BC0" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
and therefore does not characterise distinct species.
<bibRefCitation author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse (1969: 372)</bibRefCitation>
<bibRefCitation id="3CCC2B030D0B4B83CE5266DC3E34F958" author="Hesse, AJ" journalOrPublisher="Annals of the South African Museum" pageId="26" pageNumber="51" pagination="1 - 388" title="The Mydaidae (Diptera) of Southern Africa" url="http://www.biodiversitylibrary.org/bibliography/6928" volume="54" year="1969">Hesse (1969: 372)</bibRefCitation>
mentioned the ♀ specimen from Tulbagh in the Western Cape as probably being mislabelled. We have studied the specimen and agree with his identification as
<taxonomicName class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
<taxonomicName id="3BCD53C1B0AC65A42311F66E88EC6A15" class="Insecta" family="Mydidae" genus="Ectyphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectyphus pinguis" order="Diptera" pageId="16" pageNumber="41" phylum="Arthropoda" rank="species" species="pinguis">Ectyphus pinguis</taxonomicName>
, but cannot add any information whether the locality is correct or not. This locality is far removed from any other locality in the eastern Western Cape and in the western Eastern Cape provinces (Fig. 45).
</paragraph>
</subSubSection>

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@ -1,287 +1,286 @@
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<mods:title>A new family Lepidocharontidae with description of Lepidocharon gen. n., from the Great Barrier Reef, Australia, and redefinition of the Microparasellidae (Isopoda, Asellota)</mods:title>
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<mods:title id="D92ADBBBA066D570F517A3998AA41367">A new family Lepidocharontidae with description of Lepidocharon gen. n., from the Great Barrier Reef, Australia, and redefinition of the Microparasellidae (Isopoda, Asellota)</mods:title>
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<mods:namePart>Bruce, Niel L.</mods:namePart>
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<mods:namePart>Fiasca, Barbara</mods:namePart>
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<mods:namePart>Dole-Olivier, Marie-Jose</mods:namePart>
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<paragraph pageId="1" pageNumber="12">Taxon classification Animalia Isopoda Lepidocharontidae</paragraph>
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<paragraph id="0F3B5213BBDF9E8AD5A5186BD31A5E48" pageId="1" pageNumber="12">Taxon classification Animalia Isopoda Lepidocharontidae</paragraph>
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<paragraph pageId="1" pageNumber="12">
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<taxonomicNameLabel pageId="1" pageNumber="12">fam. n.</taxonomicNameLabel>
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<paragraph id="7D3C78C67350D1E49421CA1EF94494EE" pageId="1" pageNumber="12">
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<paragraph pageId="1" pageNumber="12">Diagnosis.</paragraph>
<paragraph lastPageId="2" lastPageNumber="13" pageId="1" pageNumber="12">
<subSubSection id="8AE0AB068388CFDDB3F6D764140F1AB0" lastPageId="2" lastPageNumber="13" pageId="1" pageNumber="12" type="diagnosis">
<paragraph id="AA400408E2A028C67CAA281CC7CF0D5A" pageId="1" pageNumber="12">Diagnosis.</paragraph>
<paragraph id="70A85C82D9D297639CC7029D548AB305" lastPageId="2" lastPageNumber="13" pageId="1" pageNumber="12">
Male. Body dorsally flat, slender, ~4
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long as wide, without chromatophores; somites all subsimilar in width, somites sub-rectangular or trapezoidal, lateral margins of head and pereionites sub-parallel. Pleon of one segment, with free lateral margins. Head with weak or absent rostrum, without pseudorostrum. Eyes absent. Antennula with maximally 4 flagellar articles. Antenna flagellum longer than podomeres. Antennal scale (rudimentary exopod) present, even if more or less developed among genera. Mandible incisor with 2 to 8 cusps; molar process subconical, without grinding surface, with apical unequal smooth and pinnate setae; spine row and lacinia mobilis
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<pageBreakToken id="BE148EE4CD557249D9432BB102D09580" pageId="2" pageNumber="13" start="start">present</pageBreakToken>
, the latter only on left mandible. Maxilliped slender, covering entire mouthpart field, endite distal margin narrowly rounded; epipod slender, quadrate or distally acute; palp composed of 5 articles; 2 stiff pectinate setae always present on maxilliped distal article. Pereiopods 1-7 subsimilar, always without subchela; all pereiopods with 2 dactylar claws; pereiopods articulating dorso-laterally or laterally, and projecting ventrally (in
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
<taxonomicName id="8905D2D9AD023A30F315EF020E9BAD77" class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
and
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<taxonomicName id="D7A308BD0BF2C5ABA363F1ACB4F4D0A6" class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
) or outwards (in
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<taxonomicName id="24EFC5D1F5623088DB4425014EBF1A68" class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
). Penial processes with openings coalescent and medial. Male pleopods 1 and 2 not operculate; male pleopod 1 distally rounded or subtruncate, with or without acute distolateral lobes; proximal part of the pleopod with or without scale-like elements on postero-lateral margins; stylet-guiding grooves running parallel to the lateral free distal margin of pleopod 1 and folded by a hyaline lamella (transversal and unfolded in
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
<taxonomicName id="C5AB2D89C6A9B5CB712D31DC4A95B90B" class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
); pleopod 3 endopod with 3 plumose setae (marine taxa) or without plumose setae (freshwater taxa), exopod slender. Pleopod 4 globular, pleopod 5 absent. Uropods biramous, ventrally inserted on pleotelson, protopod large, c. 0.5-1.3 as long as pleotelson; protopod length/width ratio c. 2.5-4.5; rami slender with exopod articulating anteriorly to endopod. Anus terminal, not covered by pleopods. Anus outside pleopodal chamber, between bases of uropodal protopods.
</paragraph>
<paragraph pageId="2" pageNumber="13">Female. Operculum (pleopod 2) from sub-quadrate (as long as wide) to more than 2 times longer than wide, with free distal margin deeply incised medially, faintly incised, or without medial incision, armed with 4 or 2 setae, or unarmed.</paragraph>
<paragraph id="9380670FEB6CD8CC9D4551ADB26FCD9A" pageId="2" pageNumber="13">Female. Operculum (pleopod 2) from sub-quadrate (as long as wide) to more than 2 times longer than wide, with free distal margin deeply incised medially, faintly incised, or without medial incision, armed with 4 or 2 setae, or unarmed.</paragraph>
</subSubSection>
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<paragraph pageId="2" pageNumber="13">Genera included.</paragraph>
<paragraph pageId="2" pageNumber="13">
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<paragraph id="9690DE204E4A9236833709994095771C" pageId="2" pageNumber="13">Genera included.</paragraph>
<paragraph id="D28520364664ECC015CE14C359AB906A" pageId="2" pageNumber="13">
<taxonomicName id="E0059CF60A93260A0D061EA0D594A6D6" class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
Karaman, 1934;
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<taxonomicName id="15A73F20DEF622A396E042B9CF2D3D8E" class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
Albuquerque, Boulanouar &amp; Coineau, 2014;
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<taxonomicName id="2E6FFD6CE13608B7EBEAFAFBC04AF1D0" class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
Galassi &amp; Bruce, gen. n.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="13" type="genus incertae sedis">
<paragraph pageId="2" pageNumber="13">Genus incertae sedis.</paragraph>
<paragraph pageId="2" pageNumber="13">
<taxonomicName class="Malacostraca" genus="Angeliera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angeliera" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Angeliera</taxonomicName>
<subSubSection id="E01152C6F2DA1C54141A3EBB09ED7F28" pageId="2" pageNumber="13" type="genus incertae sedis">
<paragraph id="991C24BEFD754867B71845E619D23B22" pageId="2" pageNumber="13">Genus incertae sedis.</paragraph>
<paragraph id="3D9ABA1A76E0DEEC73014FADD5239E6D" pageId="2" pageNumber="13">
<taxonomicName id="9A2F901A43CECC067A91F1546AD1680B" class="Malacostraca" genus="Angeliera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angeliera" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Angeliera</taxonomicName>
Chappuis &amp; Delamare Deboutteville, 1952.
</paragraph>
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<subSubSection lastPageId="4" lastPageNumber="15" pageId="2" pageNumber="13" type="remarks">
<paragraph pageId="2" pageNumber="13">Remarks.</paragraph>
<paragraph pageId="2" pageNumber="13">
<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="683 - 747" title="A systematic review of the family Janiridae (Crustacea: Isopoda: Asellota)." url="10.1071/IT9940683" volume="8" year="1994">
<subSubSection id="6CA62136CBB0C522B52A37896805C33D" lastPageId="4" lastPageNumber="15" pageId="2" pageNumber="13" type="remarks">
<paragraph id="2989CCAAF28824AA1D4CF3ABFF64B8FE" pageId="2" pageNumber="13">Remarks.</paragraph>
<paragraph id="4F2818CADC3E6C6E64EBFED15BAA36EC" pageId="2" pageNumber="13">
<bibRefCitation id="F1C378B128BA659A7D533537B045D0A9" author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="683 - 747" title="A systematic review of the family Janiridae (Crustacea: Isopoda: Asellota)." url="10.1071/IT9940683" volume="8" year="1994">
Wilson and
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(1994)
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and
<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="749 - 766" title="A phylogenetic analysis of the isopod family Janiridae (Crustacea)." url="10.1071/IT9940749" volume="8" year="1994">Wilson (1994)</bibRefCitation>
<bibRefCitation id="828C0D07F1DD0DE5A53A652F16B5B660" author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="749 - 766" title="A phylogenetic analysis of the isopod family Janiridae (Crustacea)." url="10.1071/IT9940749" volume="8" year="1994">Wilson (1994)</bibRefCitation>
critically discussed the status of the
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in relation to the
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<taxonomicName id="807191DE6D8E158DCA461F454BDF60A1" class="Malacostraca" family="Janiridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Janiridae</taxonomicName>
on a cladistic phylogenetic basis.
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<bibRefCitation id="270E8BA57A4E65C68EB52520C23988EC" author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="683 - 747" title="A systematic review of the family Janiridae (Crustacea: Isopoda: Asellota)." url="10.1071/IT9940683" volume="8" year="1994">
Wilson and
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(1994)
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in their review of the
<taxonomicName family="Janiridae" lsidName="" pageId="2" pageNumber="13" rank="family">Janiridae</taxonomicName>
<taxonomicName id="4AE0E29729649BBEE8A6C6FAA92C42C0" class="Malacostraca" family="Janiridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Janiridae</taxonomicName>
analysed the status of the family
<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="9E97890BD2A1F3CD49AE4B4ED09B0E93" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
in detail, reviewing the history of the debate over the status of the family.
<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="749 - 766" title="A phylogenetic analysis of the isopod family Janiridae (Crustacea)." url="10.1071/IT9940749" volume="8" year="1994">Wilson (1994)</bibRefCitation>
<bibRefCitation id="FEC72F7895E2F5BDEBF701B125C393DA" author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="749 - 766" title="A phylogenetic analysis of the isopod family Janiridae (Crustacea)." url="10.1071/IT9940749" volume="8" year="1994">Wilson (1994)</bibRefCitation>
also included the
<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="F17D1D50C591E18D5A8FDA8CEF12A2B4" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
in his cladistic analysis of the phylogeny of the
<taxonomicName family="Janiridae" lsidName="" pageId="2" pageNumber="13" rank="family">Janiridae</taxonomicName>
<taxonomicName id="E3BFF95E9235EB8B0472E160AE869647" class="Malacostraca" family="Janiridae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Janiridae</taxonomicName>
.
<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="683 - 747" title="A systematic review of the family Janiridae (Crustacea: Isopoda: Asellota)." url="10.1071/IT9940683" volume="8" year="1994">
<bibRefCitation id="A2B3930B18A1A5DE72FA8C154B6623F4" author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="683 - 747" title="A systematic review of the family Janiridae (Crustacea: Isopoda: Asellota)." url="10.1071/IT9940683" volume="8" year="1994">
Wilson and
<normalizedToken originalValue="Wägele">Waegele</normalizedToken>
<normalizedToken id="51FEE0E24B6A22BC6B19EFFDFA740A45" originalValue="Wägele">Waegele</normalizedToken>
(1994
</bibRefCitation>
: page 721) stated &quot;The family concept of the
<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="D27E03E778897A188FF64C97E60B620D" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
may be open to challenge because
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
<taxonomicName id="8D992DC93EA2BEFC6F3683E16B93839E" class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
is distinct from the other three genera in these autapomorphies&quot;, these being the differences in the somatic and uropodal morphology. These authors went on to say &quot;the composition of this family will require further study.&quot; In
<bibRefCitation author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="749 - 766" title="A phylogenetic analysis of the isopod family Janiridae (Crustacea)." url="10.1071/IT9940749" volume="8" year="1994">
<normalizedToken originalValue="Wilsons">Wilson's</normalizedToken>
<bibRefCitation id="9D24420309CDC86A19A6610CCD21F15A" author="Wilson, GDF" journalOrPublisher="Invertebrate Taxonomy" pageId="24" pageNumber="35" pagination="749 - 766" title="A phylogenetic analysis of the isopod family Janiridae (Crustacea)." url="10.1071/IT9940749" volume="8" year="1994">
<normalizedToken id="8CF531F332CE4856852AFF49F5BF08A5" originalValue="Wilsons">Wilson's</normalizedToken>
(1994)
</bibRefCitation>
analysis, there were no supporting apomorphies for the
<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="7EE364F7DAB2C3E0F1D56B158551ECA1" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
as then constituted, but there were separate supporting apomorphies for the genus
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
<taxonomicName id="D3BDB6329CAB21F6230A61F8111541ED" class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
and the clade holding the remaining genera, strongly suggesting that potentially these were two monophyletic clades, albeit the
<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="EE4D8C96135E8A811202B5164CA2DA84" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
being monogeneric. The principle basis for this is that each group had unique and derived uropod morphology and substantial differences in body morphology.
</paragraph>
<paragraph lastPageId="3" lastPageNumber="14" pageId="2" pageNumber="13">
<paragraph id="D25B00225B0FAE67BC6403E1B466F161" lastPageId="3" lastPageNumber="14" pageId="2" pageNumber="13">
The description of the new genus
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
<taxonomicName id="82F7E60B5971EB748BD23658BD87576A" class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
Galassi &amp; Bruce, gen. n. led to a re-appraisal of the taxonomic status of the
<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="77B4F686E29BB49025D6E8A65CB2A392" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
and its constituent genera. We conclude that the
<taxonomicName family="Microparasellidae" lsidName="" pageId="2" pageNumber="13" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="F2867A54F7BE86AAEA108C2AE503FD91" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="2" pageNumber="13" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
is a mono-generic family supported by a prominent acute or narrowly rounded rostrum, the antennal flagellum shorter than podomeres,
<pageBreakToken pageId="3" pageNumber="14" start="start">all</pageBreakToken>
<pageBreakToken id="4752CD2166C6C50B0B27CAEC3BF3468A" pageId="3" pageNumber="14" start="start">all</pageBreakToken>
somites with straight lateral margins that also have scales, an indisputable ventral position of the pereiopods, the unique uniramous and short uropods (see Appendix 1). The remaining genera are housed in the new family
<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
<taxonomicName id="53834EA4CEDB9787169ED15ED13836D7" class="Malacostraca" family="Lepidocharontidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Lepidocharontidae</taxonomicName>
fam. n., the diagnostic characters being the elongate body (up to 10
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="FE1D389957BF556B5AF567E9E150E91A" originalValue="×">x</normalizedToken>
as long as maximum width), a weak or absent rostrum, the antennal flagellum longer than podomeres, the pereionites rectangular or trapezoidal in dorsal view, with sub-parallel lateral margins, a lateral or dorso-lateral position of the pereiopods, a tendency to reduction of pereiopodal coxal plates, and the uropod with a large protopod with the exopod articulating anteriorly and separately to the endopod.
</paragraph>
<paragraph pageId="3" pageNumber="14">
<paragraph id="6A2C7313D10C7BCA1BEED4BF2EBD0503" pageId="3" pageNumber="14">
Within the family
<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
<taxonomicName id="C7C2BD8E4356DF4ED67B016F0EB3B338" class="Malacostraca" family="Lepidocharontidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Lepidocharontidae</taxonomicName>
there is great uniformity of the diagnostic characters among all the genera. The genus
<taxonomicName class="Malacostraca" genus="Angeliera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angeliera" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Angeliera</taxonomicName>
<taxonomicName id="5CC2BBD45B6F43D4E6F7A550AAF123B3" class="Malacostraca" genus="Angeliera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Angeliera" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Angeliera</taxonomicName>
has been placed incertae sedis in the
<taxonomicName lsidName="Asellota" pageId="3" pageNumber="14" rank="suborder" suborder="Asellota">Asellota</taxonomicName>
<taxonomicName id="2B96962ADDEFE9F0AE6DBCF49A8CDD3D" lsidName="Asellota" pageId="3" pageNumber="14" rank="suborder" suborder="Asellota">Asellota</taxonomicName>
on the basis of marked differences in several morphological features that set this genus far away the basic body plan observed in
<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
<taxonomicName id="F12234D7D9A9430B0963CB3EE4278D08" class="Malacostraca" family="Lepidocharontidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Lepidocharontidae</taxonomicName>
.
</paragraph>
<paragraph pageId="3" pageNumber="14">
<paragraph id="F81945667A30B2821317E6592E5EAFBF" pageId="3" pageNumber="14">
The largest genus in the
<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
<taxonomicName id="2AAD14C8255384EE5B4CFDFDA68BC8F6" class="Malacostraca" family="Lepidocharontidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Lepidocharontidae</taxonomicName>
is
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
<taxonomicName id="16328E30357F965170B6D53BBD41D8B1" class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
with 77 species, both marine and freshwater. Many species lack full descriptions, and there are inconsistencies in the distribution of certain characters within the genus. A dorsal view of the head is not routinely figured; when figured, it can be seen that some species do have a rostral point or rostrum, while others have the anterior margin of the head weakly concave.
</paragraph>
<paragraph pageId="3" pageNumber="14">
<paragraph id="A679D1C0B411C8CDD50502640F6410D5" pageId="3" pageNumber="14">
The generic name
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
<taxonomicName id="222210F75BEF64A76DE7D29FDB940A59" class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
is unavailable under the
<bibRefCitation author="Harrison, K" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="22" pageNumber="33" title="International Code of Zoological Nomenclature" year="1999">ICZN (1999)</bibRefCitation>
<bibRefCitation id="2B71AB58219478011FFA98448D3EC1B4" author="Harrison, K" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="22" pageNumber="33" title="International Code of Zoological Nomenclature" year="1999">ICZN (1999)</bibRefCitation>
's rules because the genus was established by
<bibRefCitation author="Karaman, S" journalOrPublisher="Mitteilungen ueber Hohlen- und Karstforschung" pageId="23" pageNumber="34" pagination="42 - 44" title="Beitrage zur Kenntnis des Isopoden-Familie Microparasellidae." volume="1934" year="1934">Karaman (1934)</bibRefCitation>
<bibRefCitation id="DDED2F834B58C5367FFC001B4E1653B6" author="Karaman, S" journalOrPublisher="Mitteilungen ueber Hohlen- und Karstforschung" pageId="23" pageNumber="34" pagination="42 - 44" title="Beitrage zur Kenntnis des Isopoden-Familie Microparasellidae." volume="1934" year="1934">Karaman (1934)</bibRefCitation>
without type species designation. This prevents the use of
<normalizedToken originalValue="“Microcharontidae”">&quot;Microcharontidae&quot;</normalizedToken>
<normalizedToken id="73E856432BD8F1750082D9BFF9E115B1" originalValue="“Microcharontidae”">&quot;Microcharontidae&quot;</normalizedToken>
(from the most speciose and well-known genus
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
<taxonomicName id="664919CDC239FF939A4530CA11AAB624" class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
) as the family name because
<bibRefCitation author="Harrison, K" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="22" pageNumber="33" title="International Code of Zoological Nomenclature" year="1999">ICZN (1999)</bibRefCitation>
<bibRefCitation id="1EB4FD89F223551BB429314A54A07725" author="Harrison, K" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="22" pageNumber="33" title="International Code of Zoological Nomenclature" year="1999">ICZN (1999)</bibRefCitation>
' article 13.2 unambiguously states: &quot;To be available, every new family-group name published after 1930 must satisfy the provisions of Article 13.1 and must be formed from an available genus-group name then used as valid by the author in the family-group taxon [Arts. 11.7.1.1, 29]&quot;. Therefore we here propose the name
<taxonomicName family="Lepidocharontidae" lsidName="" pageId="3" pageNumber="14" rank="family">Lepidocharontidae</taxonomicName>
<taxonomicName id="54B53A55C0642B92D7F7C5328B07038F" class="Malacostraca" family="Lepidocharontidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Lepidocharontidae</taxonomicName>
fam. n.
</paragraph>
<paragraph pageId="3" pageNumber="14">
<paragraph id="35EE8AF95736D9D8A6E2FFACBCA02A5C" pageId="3" pageNumber="14">
The family name
<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="F80395D17BB1F0FF217C21B259239189" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
was first introduced by
<bibRefCitation author="Karaman, S" journalOrPublisher="Mitteilungen ueber Hohlen- und Karstforschung" pageId="23" pageNumber="34" pagination="42 - 44" title="Beitrage zur Kenntnis des Isopoden-Familie Microparasellidae." volume="1934" year="1934">Karaman (1934</bibRefCitation>
<bibRefCitation id="1CC7370D6250C564DDD7BBD25C7443E2" author="Karaman, S" journalOrPublisher="Mitteilungen ueber Hohlen- und Karstforschung" pageId="23" pageNumber="34" pagination="42 - 44" title="Beitrage zur Kenntnis des Isopoden-Familie Microparasellidae." volume="1934" year="1934">Karaman (1934</bibRefCitation>
: page 44) when describing the genus
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
<taxonomicName id="BBB15027FC85BBE6BEF54592F2B79FB2" class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
, although the family had been earlier diagnosed by
<bibRefCitation author="Karaman, S" journalOrPublisher="Zoologischer Anzeiger" pageId="23" pageNumber="34" pagination="16 - 22" title="Neue Isopoden aus unterirdischen Gewaessern Jugoslawiens." volume="102" year="1933">Karaman (1933</bibRefCitation>
<bibRefCitation id="9498F72647BEA629179E86CABAA9D110" author="Karaman, S" journalOrPublisher="Zoologischer Anzeiger" pageId="23" pageNumber="34" pagination="16 - 22" title="Neue Isopoden aus unterirdischen Gewaessern Jugoslawiens." volume="102" year="1933">Karaman (1933</bibRefCitation>
: page 17) with the accompanying statement &quot;
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
<taxonomicName id="8CF7879DF84608941E534C03A6A18099" class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
n. fam., n. gen.&quot;, but without type-species designation for the genus
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
<taxonomicName id="AF8EBC603A7B5E1A63F71ECBE4214F93" class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
(see
<bibRefCitation author="Karaman, S" journalOrPublisher="Zoologischer Anzeiger" pageId="23" pageNumber="34" pagination="16 - 22" title="Neue Isopoden aus unterirdischen Gewaessern Jugoslawiens." volume="102" year="1933">Karaman 1933</bibRefCitation>
<bibRefCitation id="2A6A58E6E677524E0C2BD64047F63616" author="Karaman, S" journalOrPublisher="Zoologischer Anzeiger" pageId="23" pageNumber="34" pagination="16 - 22" title="Neue Isopoden aus unterirdischen Gewaessern Jugoslawiens." volume="102" year="1933">Karaman 1933</bibRefCitation>
). According to the
<bibRefCitation author="Harrison, K" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="22" pageNumber="33" title="International Code of Zoological Nomenclature" year="1999">ICZN (1999</bibRefCitation>
<bibRefCitation id="747062F447B13BBF17BD2A2EB4168575" author="Harrison, K" journalOrPublisher="The International Trust for Zoological Nomenclature, London, UK" pageId="22" pageNumber="33" title="International Code of Zoological Nomenclature" year="1999">ICZN (1999</bibRefCitation>
: Article 13.2) the family name
<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="09FBD0B51F05D12F8B1826CEEB12EF27" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
proposed by
<bibRefCitation author="Karaman, S" journalOrPublisher="Mitteilungen ueber Hohlen- und Karstforschung" pageId="23" pageNumber="34" pagination="42 - 44" title="Beitrage zur Kenntnis des Isopoden-Familie Microparasellidae." volume="1934" year="1934">Karaman (1934)</bibRefCitation>
<bibRefCitation id="908AD6577EA0504EEF730BA5110AB2A5" author="Karaman, S" journalOrPublisher="Mitteilungen ueber Hohlen- und Karstforschung" pageId="23" pageNumber="34" pagination="42 - 44" title="Beitrage zur Kenntnis des Isopoden-Familie Microparasellidae." volume="1934" year="1934">Karaman (1934)</bibRefCitation>
is thus a nomen nudum, because the family was erected on the unavailable generic name
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
<taxonomicName id="1496C03E48FB43A0DE0D5E62AD5DC750" class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
Karaman, 1933 that lacked type-species designation. Nevertheless, Article 13.2.1 states that &quot;A family group name first published after 1930 and before 1961 which does not satisfy the provisions of Article 13.1 is available from its original publication only if it was used as valid before 2000, and also was not rejected by an author who, after 1960 and before 2000, expressly applied Article 13 of the then current editions of the Code&quot;. The family name was considered valid until 2000, and for this reason it is an available name as
<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="4AB95525C7E49DBFF37A8AB4E1971709" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
Karaman, 1934. The family name
<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="3DDE072D59FAAC9C0733862AD8218415" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
is then valid.
</paragraph>
<paragraph lastPageId="4" lastPageNumber="15" pageId="3" pageNumber="14">
<paragraph id="27D780FED1C17054B9C4D5EC60694E96" lastPageId="4" lastPageNumber="15" pageId="3" pageNumber="14">
Conversely, we provisionally maintain current and common usage of the names
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<taxonomicName id="D3E938B5999125CBE404A5B877030E86" class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
and
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
<taxonomicName id="801D21BB8CB8C4005559ECCCE50A2F0D" class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
, and this is discussed in more detail together with a new diagnosis for the
<taxonomicName family="Microparasellidae" lsidName="" pageId="3" pageNumber="14" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="6E33BE7DE85D7F33A19742C1EA0BD78B" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="3" pageNumber="14" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
(see Appendix 1). As the nomenclature within
<pageBreakToken pageId="4" pageNumber="15" start="start">the</pageBreakToken>
<pageBreakToken id="6F7E2D540A51B624EE2FE3D8A8A05F0C" pageId="4" pageNumber="15" start="start">the</pageBreakToken>
family
<taxonomicName family="Microparasellidae" lsidName="" pageId="4" pageNumber="15" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="9BC1B320D573FDA8632AF7DE3AF0C3EE" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
is well established and widely used, a proposition (Galassi and Bruce in preparation) will be submitted to the ICZN Commission for maintaining the stability of the current nomenclature and related authorities.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="15" type="key">
<paragraph pageId="4" pageNumber="15">
<subSubSection id="AA893DF6687C031A075D5D23E1FBF974" pageId="4" pageNumber="15" type="key">
<paragraph id="6E31ED5BBAE602F6ED0A9636BC72A09C" pageId="4" pageNumber="15">
Key to
<taxonomicName family="Microparasellidae" lsidName="" pageId="4" pageNumber="15" rank="family">Microparasellidae</taxonomicName>
<taxonomicName id="E8B17E3AE1129DA4E1E38183508BF31A" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
and genera of
<taxonomicName family="Lepidocharontidae" lsidName="" pageId="4" pageNumber="15" rank="family">Lepidocharontidae</taxonomicName>
<taxonomicName id="0FE9CD635A7688A77D006D589E99B35F" class="Malacostraca" family="Lepidocharontidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="family">Lepidocharontidae</taxonomicName>
fam. n.
</paragraph>
<paragraph pageId="4" pageNumber="15">
<table pageId="4" pageNumber="15">
<tr pageId="4" pageNumber="15">
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">
<taxonomicName family="Microparasellidae" lsidName="" pageId="4" pageNumber="15" rank="family">Microparasellidae</taxonomicName>
<paragraph id="726D13B3E63B2E4B671151672247AEB3" pageId="4" pageNumber="15">
<table id="4635F74B5D36B829C417207E5C2106D1" pageId="4" pageNumber="15">
<tr id="81611EBE1337D25B2746744EA5E9B09C" pageId="4" pageNumber="15">
<td id="997CA10A91F3A50A83EA7D2ADA69E08D" colspan="1" pageId="4" pageNumber="15" rowspan="1">
<taxonomicName id="CEEDC231045FDFB25FB75D9DF8606CC5" class="Malacostraca" family="Microparasellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="family">Microparasellidae</taxonomicName>
(
<taxonomicName class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
<taxonomicName id="C991834A9606923258A309005C499EDE" class="Malacostraca" family="Microparasellidae" genus="Microparasellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microparasellus" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Microparasellus</taxonomicName>
)
</td>
</tr>
<tr pageId="4" pageNumber="15">
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">2</td>
<tr id="4B0040D72053E593A4F08BF2C430CB76" pageId="4" pageNumber="15">
<td id="4127D7F1CB81DD28C29411F38AC678FD" colspan="1" pageId="4" pageNumber="15" rowspan="1">2</td>
</tr>
<tr pageId="4" pageNumber="15">
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
<tr id="ECFDBB282E7367BE507DF37160D9AED6" pageId="4" pageNumber="15">
<td id="A31469F517D9C2AAF6D00199D020A317" colspan="1" pageId="4" pageNumber="15" rowspan="1">
<taxonomicName id="7B6F1174BFC9EA53C27A4BA416B0DCC4" class="Malacostraca" family="Lepidocharontidae" genus="Microcharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microcharon" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Microcharon</taxonomicName>
</td>
</tr>
<tr pageId="4" pageNumber="15">
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">4</td>
<tr id="D76C983D0A1F1D2930B3207D94416948" pageId="4" pageNumber="15">
<td id="8ED335F0658DAD489B5E4517B62564F5" colspan="1" pageId="4" pageNumber="15" rowspan="1">4</td>
</tr>
<tr pageId="4" pageNumber="15">
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
<tr id="CB5588AE3CD891CF23EAF75689A21A6B" pageId="4" pageNumber="15">
<td id="CCA6625744BD6EA33084DE4F5B1CF57D" colspan="1" pageId="4" pageNumber="15" rowspan="1">
<taxonomicName id="4FDD0901C525A8118D23419B80ECA2F8" class="Malacostraca" family="Lepidocharontidae" genus="Janinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Janinella" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Janinella</taxonomicName>
</td>
</tr>
<tr pageId="4" pageNumber="15">
<td colspan="1" pageId="4" pageNumber="15" rowspan="1">
<taxonomicName class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
<tr id="C2260DDF9333E379EBF74865D7AA6643" pageId="4" pageNumber="15">
<td id="BF3C6F6306527C879AAE4006EEC63722" colspan="1" pageId="4" pageNumber="15" rowspan="1">
<taxonomicName id="8AC894C0E05E29CF05EB911F594C44E6" class="Malacostraca" family="Lepidocharontidae" genus="Lepidocharon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lepidocharon" order="Isopoda" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">Lepidocharon</taxonomicName>
gen. n.
</td>
</tr>

View file

@ -1,186 +1,186 @@
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Glaucotrema thailandicum Naksuwankul,
<normalizedToken originalValue="Lücking">Luecking</normalizedToken>
<normalizedToken id="162AA4B81BD1E556683A7747DD20A09B" originalValue="Lücking">Luecking</normalizedToken>
&amp; Lumbsch
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Figure 3
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Figure 3
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THAILAND, Ubon Ratchathani Province, Pha Tam National Park, Sang Chan waterfall,
<geoCoordinate direction="north" orientation="latitude" precision="925" value="15.5">15°30'N</geoCoordinate>
<geoCoordinate id="74BFB7AAE4F2E3CE7F82C60B5790A830" direction="north" orientation="latitude" precision="925" value="15.5">15°30'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="105.583336">105°35'E</geoCoordinate>
<geoCoordinate id="56B56A16286917FDCA2A64762DD960A6" direction="east" orientation="longitude" precision="925" value="105.583336">105°35'E</geoCoordinate>
, 124 m, dry evergreen forest, on bark; 12 April 2013, K. Papong 8560 (holotype: MSUT; isotype: F).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="51" type="diagnosis">
<paragraph pageId="4" pageNumber="51">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="51">Characterized within the genus by having submuriform ascospores.</paragraph>
<subSubSection id="EFF4370028F3AE2FC482FC0E10158496" pageId="4" pageNumber="51" type="diagnosis">
<paragraph id="6F65F282F0FAF9C6C8A34C8ADC6BE2B7" pageId="4" pageNumber="51">Diagnosis.</paragraph>
<paragraph id="8E58720ACD743513C7DC513577A67223" pageId="4" pageNumber="51">Characterized within the genus by having submuriform ascospores.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="51" type="etymology">
<paragraph pageId="4" pageNumber="51">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="51">The specific epithet refers to the country where the type specimen was collected.</paragraph>
<subSubSection id="AA0EA3E415565DD1F2D4D13951865801" pageId="4" pageNumber="51" type="etymology">
<paragraph id="B701BA5B824889940B17105369AF19CD" pageId="4" pageNumber="51">Etymology.</paragraph>
<paragraph id="C4A7C8519CB9CBFECF46E2EB80BFB8A6" pageId="4" pageNumber="51">The specific epithet refers to the country where the type specimen was collected.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="51" type="description">
<paragraph pageId="4" pageNumber="51">Description.</paragraph>
<paragraph pageId="4" pageNumber="51">
<subSubSection id="354A02148942506909D738E101407A91" pageId="4" pageNumber="51" type="description">
<paragraph id="D6EC176134586BFE0F69340C3D65C6AE" pageId="4" pageNumber="51">Description.</paragraph>
<paragraph id="4A4D5BA3195DAD49B9A81F73DE9FB95F" pageId="4" pageNumber="51">
Thallus endophloeodal to epiphloeodal, up to c. 120
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="D04A274E8038AEF7650D979AF2ABEB8E" originalValue="µm">µm</normalizedToken>
thick, pale green to yellowish green, smooth. True cortex
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="18B45896DE0CEC6829F2E70FC01B4B75" originalValue="±">+/-</normalizedToken>
continuous, to c. 25
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="D71AA4B661337CA67CD7FBBD7CE790B4" originalValue="µm">µm</normalizedToken>
thick. Algal layer poorly to well developed,
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="5646B485DA1C82A026F6FB79E8B62491" originalValue="±">+/-</normalizedToken>
continuous; calcium oxalate crystals sparse to abundant, large and clustered; medulla usually distinct. Vegetative propagules not seen. Ascomata conspicuous, to c. 0.8-1.2 mm diam., often larger when fused,
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="4A423B9A3A5BE988B0D8646DC77FA43A" originalValue="±">+/-</normalizedToken>
rounded to irregular, apothecioid to somewhat chroodiscoid, solitary to more often fused, becoming slightly to distinctly emergent, mostly irregularly or regularly urceolate. Disc usually partly visible from above, rarely completely exposed, pale yellowish to whitish green. Pores broad to gaping, to c. 0.6-08 mm wide,
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="8FE55BD65E9F99DBE835E47E15880DD4" originalValue="±">+/-</normalizedToken>
rounded to irregular, entire to slightly ragged; thalline exciple often becoming apically visible, rarely completely visible from above,
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="FEFDB1858D71B144816DC4D3C626BEFA" originalValue="±">+/-</normalizedToken>
free, whitish. Thalline rim margin broad to gaping,
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="5E5F9555ABBD8FC05B053467822D3260" originalValue="±">+/-</normalizedToken>
rounded, more commonly irregular, thick, entire, concolorous to whitish. Thalline exciple fused to partly or entirely free, thick, hyaline internally, pale yellowish or greenish marginally, with calcium oxalate crystals. Hymenium to c. 120
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="ED634DA7BF94EF7D46A1BCA4A4B76340" originalValue="µm">µm</normalizedToken>
thick, clear, strongly conglutinated; paraphyses thick, irregular and often distoseptate,
<normalizedToken originalValue="±">+/-</normalizedToken>
<normalizedToken id="FC549B597CA1326CAD587B55ED75E361" originalValue="±">+/-</normalizedToken>
interwoven, with thickened irregular tips; lateral paraphyses absent; columella whitish and reticulate. Epihymenium hyaline, with fine crystals. Asci 8-spored; tholus initially thick, thin when mature, 100-110
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="784FC254A86B58D2270CB42CD76722C8" originalValue="×">x</normalizedToken>
10-12
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="26403AF31D8AEB47A68C7FD684658FDD" originalValue="µm">µm</normalizedToken>
. Ascospores submuriform with 3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="0FCA9D6307B219E9D7F764D0E8EF9476" originalValue="×">x</normalizedToken>
0-1 septa, hyaline, slightly amyloid, 15-20
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="168F2E29479761B594A37A20DBF7BB68" originalValue="×">x</normalizedToken>
7.5
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="DFBA4F5450FD1FF542734BCAAA918A5F" originalValue="µm">µm</normalizedToken>
. Pycnidia not seen.
</paragraph>
<caption pageId="4" pageNumber="51">
<paragraph pageId="4" pageNumber="51">
<caption id="64F2B15F21F82E372BCDDC2DA4085568" pageId="4" pageNumber="51">
<paragraph id="7B586DD65AD0AD92EDAC9904CC9E76FE" pageId="4" pageNumber="51">
Figure 3. Morphology and anatomy of
<taxonomicName class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema thailandicum" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="thailandicum">Glaucotrema thailandicum</taxonomicName>
<taxonomicName id="C020A9DC7A8C34EA2A5AE9C5CA54EDCF" class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema thailandicum" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="thailandicum">Glaucotrema thailandicum</taxonomicName>
(
<normalizedToken originalValue="AE">A-E</normalizedToken>
<normalizedToken id="8B0BA094E7CBB24DBF4C83441245F55F" originalValue="AE">A-E</normalizedToken>
)
<normalizedToken originalValue="AB">A-B</normalizedToken>
<normalizedToken id="EEB89FB8AB82C12BAAF413BE460BD61C" originalValue="AB">A-B</normalizedToken>
habitat of ascoma C cross-section of ascoma show whitish and reticulate columella D asci with spores and E submuriform ascospores (holotype),
<taxonomicName class="Lecanoromycetes" family="Graphidaceae" genus="Ocellularia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ocellularia klinhomii" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="klinhomii">Ocellularia klinhomii</taxonomicName>
<taxonomicName id="55AA1064447EA9161C5299CF2DA68B8C" class="Lecanoromycetes" family="Graphidaceae" genus="Ocellularia" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Ocellularia klinhomii" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="klinhomii">Ocellularia klinhomii</taxonomicName>
(
<normalizedToken originalValue="FK">F-K</normalizedToken>
<normalizedToken id="F3A9655E7115D68E105673D95EC6D8DD" originalValue="FK">F-K</normalizedToken>
)
<normalizedToken originalValue="FG">F-G</normalizedToken>
<normalizedToken id="E7CB9A9DD6891E48E2451CC815E7A3DB" originalValue="FG">F-G</normalizedToken>
ascomata immersed in verrucae and surrounded by a black ring H cross-section of ascoma with carbonized columella and apically carbonized exiple
<normalizedToken originalValue="IK">I-K</normalizedToken>
<normalizedToken id="487376EC3BE7FDBC5EC56434BBEED095" originalValue="IK">I-K</normalizedToken>
ascus and ascospores (holotype). Scale bar
<normalizedToken originalValue="AB">A-B</normalizedToken>
<normalizedToken id="31DB3B311E8747FF25F5C5C607B71C0F" originalValue="AB">A-B</normalizedToken>
,
<normalizedToken originalValue="FG">F-G</normalizedToken>
<normalizedToken id="E0B0DF1613A213801182F16E9BA57312" originalValue="FG">F-G</normalizedToken>
= 1 mm, H = 100
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="F2B0E57CDF49761719B7335B5D94BF52" originalValue="µm">µm</normalizedToken>
,
<normalizedToken originalValue="CD">C-D</normalizedToken>
<normalizedToken id="53648D3764A5AC8DD5170AA7FEA99A99" originalValue="CD">C-D</normalizedToken>
, I = 50
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="01E6D2EC6C8B72995F9A03B84FF34CD1" originalValue="µm">µm</normalizedToken>
, E,
<normalizedToken originalValue="JK">J-K</normalizedToken>
<normalizedToken id="1CC4F9327F6E904DB6692B5440E0BA19" originalValue="JK">J-K</normalizedToken>
= 20
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="E0EC8BDD4F94E1F9D545AF6AA48425B7" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="51" type="secondary chemistry">
<paragraph pageId="4" pageNumber="51">Secondary chemistry.</paragraph>
<paragraph pageId="4" pageNumber="51">
<subSubSection id="56FF97F7C1A6F2BE45175D88654A18E7" pageId="4" pageNumber="51" type="secondary chemistry">
<paragraph id="E408E3CD77DAE72EA7C4AB196459AF3A" pageId="4" pageNumber="51">Secondary chemistry.</paragraph>
<paragraph id="797D97EDD5FAFFD34450A2CE7BC93609" pageId="4" pageNumber="51">
Thallus K+ yellowish,
<normalizedToken originalValue="C">C-</normalizedToken>
<normalizedToken id="88B5138070B35F852270828B1AB8EF12" originalValue="C">C-</normalizedToken>
, P+ yellow; containing psoromic acid.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="51" type="distribution">
<paragraph pageId="4" pageNumber="51">Distribution and ecology.</paragraph>
<paragraph pageId="4" pageNumber="51">The new species was found in northeastern Thailand, growing on bark in a dry evergreen forest. It is known only from the type locality.</paragraph>
<subSubSection id="2F904082CC1A4358173888A0A5C6C567" pageId="4" pageNumber="51" type="distribution">
<paragraph id="A82F17D5969FB74CC36A4A595FA55783" pageId="4" pageNumber="51">Distribution and ecology.</paragraph>
<paragraph id="7013B26108E45906B2439DA178101922" pageId="4" pageNumber="51">The new species was found in northeastern Thailand, growing on bark in a dry evergreen forest. It is known only from the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="51" type="remarks">
<paragraph pageId="4" pageNumber="51">Remarks.</paragraph>
<paragraph pageId="4" pageNumber="51">
<subSubSection id="10F1141606315F057328F9BB02091D1B" pageId="4" pageNumber="51" type="remarks">
<paragraph id="41B1E3D56603C7BD67A8C1FD49BF17BE" pageId="4" pageNumber="51">Remarks.</paragraph>
<paragraph id="E1C5B21DC0AE7ADC093D35F6E834072E" pageId="4" pageNumber="51">
This new species is unique within the genus in having submuriform ascospores, whereas all other described species have transversely septate ascospores. In addition, the ascospores in
<taxonomicName class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema bahianum" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="bahianum">Glaucotrema bahianum</taxonomicName>
<taxonomicName id="AC8D87F1E14F81FE5597FC4C52BCEDB8" class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema bahianum" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="bahianum">Glaucotrema bahianum</taxonomicName>
,
<taxonomicName class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema costaricense" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="costaricense">Glaucotrema costaricense</taxonomicName>
<taxonomicName id="AF58F29011EC1866123D4DF57D417493" class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema costaricense" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="costaricense">Glaucotrema costaricense</taxonomicName>
and
<taxonomicName class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema stegoboloides" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="stegoboloides">Glaucotrema stegoboloides</taxonomicName>
<taxonomicName id="EE6D1DE64FC7BF61EA172E8BD9682C6D" class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema stegoboloides" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="stegoboloides">Glaucotrema stegoboloides</taxonomicName>
are smaller than in the new species. Molecular data support the distinction of the new taxon. In morphology it resembles
<taxonomicName class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema bahianum" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="bahianum">Glaucotrema bahianum</taxonomicName>
<taxonomicName id="739F3C96815846880C1AA19FB4DCD20D" class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema bahianum" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="bahianum">Glaucotrema bahianum</taxonomicName>
and
<taxonomicName class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema stegoboloides" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="stegoboloides">Glaucotrema stegoboloides</taxonomicName>
<taxonomicName id="D908C94FB167A8A94F5B48A6333A76BA" class="Lecanoromycetes" family="Graphidaceae" genus="Glaucotrema" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Glaucotrema stegoboloides" order="Ostropales" pageId="4" pageNumber="51" phylum="Ascomycota" rank="species" species="stegoboloides">Glaucotrema stegoboloides</taxonomicName>
.
</paragraph>
</subSubSection>

View file

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<treatment ID-GBIF-Taxon="161840392" LSID="urn:lsid:zoobank.org:act:939FDB9B-0561-473C-921D-D9AD409E87BF" httpUri="http://treatment.plazi.org/id/4D3EDE20AB6F57C3B79E98D70B15486C" lastPageNumber="131" pageId="0" pageNumber="131">
<subSubSection pageId="0" pageNumber="131" type="nomenclature">
<paragraph pageId="0" pageNumber="131">
<taxonomicName LSID="4D3EDE20-AB6F-57C3-B79E-98D70B15486C" authority="Will, 2020" authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis" status="sp. nov.">Tiferonia leytensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="131">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tiferonia leytensis sp. nov. head, left lateral view. Arrows indicate a post-ocular sulcus and b posterior edge of the orbit." figureDoi="10.3897/zookeys.906.48255.figure1" httpUri="https://binary.pensoft.net/fig/374830" pageId="0" pageNumber="131">Figs 1</figureCitation>
<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Tiferonia species aedeagi, right lateral view (A) and dorsal view (B). 2 Tiferonia leytensis sp. nov. 3 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures2-3" httpUri="https://binary.pensoft.net/fig/374831" pageId="0" pageNumber="131">, 2</figureCitation>
<figureCitation captionStart="Figures 4, 5" captionStartId="F3" captionText="Figures 4, 5. Tiferonia species pronota. 4 Tiferonia leytensis sp. nov. 5 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures4-5" httpUri="https://binary.pensoft.net/fig/374832" pageId="0" pageNumber="131">, 4</figureCitation>
<figureCitation captionStart="Figures 69" captionStartId="F4" captionText="Figures 6 - 9. Tiferonia species dorsal habitus. 6 Tiferonia parva, male paratype 7 Tiferonia leytensis male holotype 8 Tiferonia schoutedeni 9 Tiferonia brunnea, male holotype." figureDoi="10.3897/zookeys.906.48255.figures6-9" httpUri="https://binary.pensoft.net/fig/374833" pageId="0" pageNumber="131">, 7</figureCitation>
<treatment id="4D3EDE20AB6F57C3B79E98D70B15486C" ID-GBIF-Taxon="161840392" LSID="urn:lsid:zoobank.org:act:939FDB9B-0561-473C-921D-D9AD409E87BF" httpUri="http://treatment.plazi.org/id/4D3EDE20AB6F57C3B79E98D70B15486C" lastPageNumber="131" pageId="0" pageNumber="131" scope_family="Carabidae" scope_tribe="Abacetini">
<subSubSection id="33E531A51ED2C9950F889DCB19CC2E4C" pageId="0" pageNumber="131" type="nomenclature">
<paragraph id="411B6E9E089D5DF973958DF5DE04B867" pageId="0" pageNumber="131">
<taxonomicName id="8EC51555525342929E524A6827B5414D" ID-CoL="8MCR8" LSID="4D3EDE20-AB6F-57C3-B79E-98D70B15486C" authority="Will, 2020" authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis" status="sp. nov.">Tiferonia leytensis</taxonomicName>
<taxonomicNameLabel id="328451CFEE3D977FC10505E5972324B3" pageId="0" pageNumber="131">sp. nov.</taxonomicNameLabel>
<figureCitation id="C0D71C3C44FF52F72347ECAD351450C2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Tiferonia leytensis sp. nov. head, left lateral view. Arrows indicate a post-ocular sulcus and b posterior edge of the orbit." figureDoi="10.3897/zookeys.906.48255.figure1" httpUri="https://binary.pensoft.net/fig/374830" pageId="0" pageNumber="131">Figs 1</figureCitation>
<figureCitation id="40C97DE9B08617DD07DC88826CF1CFAF" captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Tiferonia species aedeagi, right lateral view (A) and dorsal view (B). 2 Tiferonia leytensis sp. nov. 3 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures2-3" httpUri="https://binary.pensoft.net/fig/374831" pageId="0" pageNumber="131">, 2</figureCitation>
<figureCitation id="8095EA7A00F98CBCF454487F9C0EFE99" captionStart="Figures 4, 5" captionStartId="F3" captionText="Figures 4, 5. Tiferonia species pronota. 4 Tiferonia leytensis sp. nov. 5 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures4-5" httpUri="https://binary.pensoft.net/fig/374832" pageId="0" pageNumber="131">, 4</figureCitation>
<figureCitation id="64DAFDC516689184332646D6663AC016" captionStart="Figures 69" captionStartId="F4" captionText="Figures 6 - 9. Tiferonia species dorsal habitus. 6 Tiferonia parva, male paratype 7 Tiferonia leytensis male holotype 8 Tiferonia schoutedeni 9 Tiferonia brunnea, male holotype." figureDoi="10.3897/zookeys.906.48255.figures6-9" httpUri="https://binary.pensoft.net/fig/374833" pageId="0" pageNumber="131">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="materials_examined">
<paragraph pageId="0" pageNumber="131">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Holotype</emphasis>
<subSubSection id="97262BE22D48D3CA6C098D5A86B45BD4" pageId="0" pageNumber="131" type="materials_examined">
<paragraph id="93BFA752644CDF8DED5EC6448D20287F" pageId="0" pageNumber="131">Material examined.</paragraph>
<paragraph id="4FFCF7DBD8E246DAD447858E44C8B4D1" pageId="0" pageNumber="131">
<emphasis id="BCE69A2A6BBDB9C0AF20C8873CB31808" bold="true" italics="true" pageId="0" pageNumber="131">Holotype</emphasis>
: Philippines • ♂, // &quot;Plains of NE Leyte Is.,P.I. Nov
<normalizedToken originalValue="44-Jan">'44-Jan'</normalizedToken>
<normalizedToken id="70B533D0729FDA5343D02520443FA69B" originalValue="44-Jan">'44-Jan'</normalizedToken>
45 Darlington&quot;// &quot;MCZ Holotype 36215&quot;// [deposited MCZ].
</paragraph>
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Paratypes</emphasis>
<paragraph id="35CA213824BDD08EE6174E0F11E8BF3C" pageId="0" pageNumber="131">
<emphasis id="264A4BF52DD009FF60B0064C7F7D1A13" bold="true" italics="true" pageId="0" pageNumber="131">Paratypes</emphasis>
: Philippines • 1♂, 1 ♀, same data as holotype [MCZ].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="type locality">
<paragraph pageId="0" pageNumber="131">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="131">
<subSubSection id="4636FC4CE033D659D85E9435979ED56A" pageId="0" pageNumber="131" type="type locality">
<paragraph id="B9829A4090C9D5AB355A00EF736E7B26" pageId="0" pageNumber="131">Type locality.</paragraph>
<paragraph id="2C6BBE7DEAF605372A34016E2574CB83" pageId="0" pageNumber="131">
As listed on locality label, type locality is estimated to be roughly centered on
<geoCoordinate degrees="11.221" direction="north" orientation="latitude" precision="55" value="11.221">11.221N</geoCoordinate>
<geoCoordinate id="CE3897A3D83A3DD63BFC9FEC52BF464D" degrees="11.221" direction="north" orientation="latitude" precision="55" value="11.221">11.221N</geoCoordinate>
,
<geoCoordinate degrees="124.828" direction="east" orientation="longitude" precision="55" value="124.828">124.828E</geoCoordinate>
<geoCoordinate id="E51552370AD9306CFF0D13DDEFA40D1B" degrees="124.828" direction="east" orientation="longitude" precision="55" value="124.828">124.828E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="diagnosis">
<paragraph pageId="0" pageNumber="131">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="131">
<subSubSection id="AF38E1F72F17D80E09AA26B46895CBBE" pageId="0" pageNumber="131" type="diagnosis">
<paragraph id="24BD23985A080210E7C0940416D9ACEE" pageId="0" pageNumber="131">Diagnosis.</paragraph>
<paragraph id="55466CA528244806BD0A0DA171AEDEFB" pageId="0" pageNumber="131">
The combination of two pairs of supraorbital setae, all elytral striae impressed, and the pronotum (Fig.
<figureCitation captionStart="Figures 4, 5" captionStartId="F3" captionText="Figures 4, 5. Tiferonia species pronota. 4 Tiferonia leytensis sp. nov. 5 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures4-5" httpUri="https://binary.pensoft.net/fig/374832" pageId="0" pageNumber="131">4</figureCitation>
<figureCitation id="3064AE98BFC0B4CF53B0D644281E30B5" captionStart="Figures 4, 5" captionStartId="F3" captionText="Figures 4, 5. Tiferonia species pronota. 4 Tiferonia leytensis sp. nov. 5 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures4-5" httpUri="https://binary.pensoft.net/fig/374832" pageId="0" pageNumber="131">4</figureCitation>
) basally narrowed with slightly sinuate lateral margins separates this species from all other species of
<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
<taxonomicName id="80EBB6AD50F0AE94C7AF77E1ABA576A4" authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="004136259B024934ADBE77724A4C29A6" italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
</taxonomicName>
.
</paragraph>
<caption doi="10.3897/zookeys.906.48255.figures4-5" httpUri="https://binary.pensoft.net/fig/374832" pageId="0" pageNumber="131" start="Figures 4, 5" startId="F3">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figures 4, 5.</emphasis>
<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
<caption id="79F15C6898D7169E7D2510BE4926850C" doi="10.3897/zookeys.906.48255.figures4-5" httpUri="https://binary.pensoft.net/fig/374832" pageId="0" pageNumber="131" start="Figures 4, 5" startId="F3">
<paragraph id="96FA2EF47741302D1DAC54FEABB618E3" pageId="0" pageNumber="131">
<emphasis id="447D1FE772020402BADB74E699F470A0" bold="true" pageId="0" pageNumber="131">Figures 4, 5.</emphasis>
<taxonomicName id="CAD0233E4F99E15C8D6995D5FEBFC394" authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="A587E2E444A3BD2772EE3C4CC41B1EF1" italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
</taxonomicName>
species pronota.
<emphasis bold="true" pageId="0" pageNumber="131">4</emphasis>
<taxonomicName authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia leytensis</emphasis>
<emphasis id="1546CE607402726308C57C9501DE0B6B" bold="true" pageId="0" pageNumber="131">4</emphasis>
<taxonomicName id="F11ED862F54D11CF1192A4F1A55EBC32" authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis">
<emphasis id="515AB49AFBAFD2C41E1412564030382C" italics="true" pageId="0" pageNumber="131">Tiferonia leytensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="131">5</emphasis>
<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia parva" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia parva</emphasis>
<emphasis id="FB05741DC81FB0A6D5447AC12E6B543A" bold="true" pageId="0" pageNumber="131">5</emphasis>
<taxonomicName id="90A8E7C777874EDECF81332B4EEFB33B" authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia parva" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="80EE3AF76E4CB98971452D0862889398" italics="true" pageId="0" pageNumber="131">Tiferonia parva</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="description">
<paragraph pageId="0" pageNumber="131">Description.</paragraph>
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Size.</emphasis>
<subSubSection id="1403D37A9AA436E28913BD36D1DC0A7B" pageId="0" pageNumber="131" type="description">
<paragraph id="17132E46A8424322A56909056C1DB09B" pageId="0" pageNumber="131">Description.</paragraph>
<paragraph id="C5CC6F3692FE9554DDE3A468E984651D" pageId="0" pageNumber="131">
<emphasis id="A9F1040445D1DBF948945F51D0AA16FF" bold="true" italics="true" pageId="0" pageNumber="131">Size.</emphasis>
Overall length (sbl) 3.96 mm [3.96-4.21 mm]; greatest width over elytra 1.65 mm [1.62-1.79 mm].
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Color.</emphasis>
<emphasis id="368F3FC9535F86916BA9949031ED9245" bold="true" italics="true" pageId="0" pageNumber="131">Color.</emphasis>
Dorsal and ventral surfaces piceous to dark rufous, head slightly darker, elytral interval 1 and apical fourth of elytra paler; legs, mouthparts, and antennae pale brunneous, tibiae darker rufous.
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Luster.</emphasis>
<emphasis id="52CB2EFC5B91E41DDDAEF4C5DD52ECB3" bold="true" italics="true" pageId="0" pageNumber="131">Luster.</emphasis>
Dorsally and ventrally distinctly shiny. Iridescence not evident on head, spectral iridescence slightly apparent on pronotum, distinctly evident on elytra, slightly apparent on ventral surface of body.
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Head.</emphasis>
<emphasis id="E96111DA63939ECA5DB515AC67150B14" bold="true" italics="true" pageId="0" pageNumber="131">Head.</emphasis>
Dorsal microsculpture evident nearly isodiametric mesh. Clypeal-ocular impressions clearly impressed, narrow, divergent, and extended to anterior supraocular seta. Ocular ratio 1.61 [1.55-1.58]. Eyes moderate size, prominent, with large posterior orbital area; gena slightly narrower than width of antennomere 1. Labrum with anterior margin straight. Mentum median tooth entire, triangular. Antennae, overall length moderately long, antennomeres 10-11 surpassing pronotal base.
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Thorax.</emphasis>
<emphasis id="DAF7C3DD3D954C02AB45116F758DD28B" bold="true" italics="true" pageId="0" pageNumber="131">Thorax.</emphasis>
Pronotum dorsally impunctate, microsculpture not evident at 50
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="7B7D33C765EAA9F2743D2D4E64561C47" originalValue="×">x</normalizedToken>
magnification in anterior half, irregular, very transverse mesh slightly evident near base; widest at middle, lateral margins with short, slight sinuation near hind angles; lateral marginal bead uniformly evident and narrow throughout; basal margin smooth, not beaded; anterior angles moderately produced; anterior submarginal sulcus broadly interrupted medially; hind angles right angled, denticulate; basal impressions linear, deeply impressed, slightly crenulate, shallowly reaching basal margin; seta at hind angle touching marginal bead. Elytra parallel sided; plica large and externally visible. Elytral striae well impressed, deeply, densely crenulate-punctate; intervals nearly flat, slightly convex basally. Elytral microsculpture scarcely evident as very transversely stretched sculpticells. Prosternal process rounded, margin not marked with bead; prosterna and proepisterna smooth. Mesosterna with few, coarse punctures. Metasternum laterally and metepisternum with shallow, coarse punctures.
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Abdomen.</emphasis>
<emphasis id="2F07802EC7DB9ADCC0E50AA73D4269DD" bold="true" italics="true" pageId="0" pageNumber="131">Abdomen.</emphasis>
Abdominal ventrites irregularly, coarsely punctate laterally, impunctate medially.
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Male genitalia</emphasis>
<emphasis id="95AFEF2F414843C209CBF6B466BD9B29" bold="true" italics="true" pageId="0" pageNumber="131">Male genitalia</emphasis>
(Fig.
<figureCitation captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Tiferonia species aedeagi, right lateral view (A) and dorsal view (B). 2 Tiferonia leytensis sp. nov. 3 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures2-3" httpUri="https://binary.pensoft.net/fig/374831" pageId="0" pageNumber="131">2</figureCitation>
<figureCitation id="3B61EA1C1C6F6E4EA3B5EB62572A5B1B" captionStart="Figures 2, 3" captionStartId="F2" captionText="Figures 2, 3. Tiferonia species aedeagi, right lateral view (A) and dorsal view (B). 2 Tiferonia leytensis sp. nov. 3 Tiferonia parva." figureDoi="10.3897/zookeys.906.48255.figures2-3" httpUri="https://binary.pensoft.net/fig/374831" pageId="0" pageNumber="131">2</figureCitation>
), ostium dorsal, endophallus with light spine field in left apical position in repose.
<emphasis bold="true" italics="true" pageId="0" pageNumber="131">Female ovipositor</emphasis>
<emphasis id="8BF78173163D1989048BF9C1BA1C03B1" bold="true" italics="true" pageId="0" pageNumber="131">Female ovipositor</emphasis>
moderately long, slightly curved, two large ensiform setae, one dorsal one ventral, two long nematoform setae in well-developed groove.
</paragraph>
<caption doi="10.3897/zookeys.906.48255.figures6-9" httpUri="https://binary.pensoft.net/fig/374833" pageId="0" pageNumber="131" start="Figures 69" startId="F4">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figures 6-9.</emphasis>
<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
<caption id="DECD151AB4B1992E660A79603F9FC79A" doi="10.3897/zookeys.906.48255.figures6-9" httpUri="https://binary.pensoft.net/fig/374833" pageId="0" pageNumber="131" start="Figures 69" startId="F4">
<paragraph id="F8DCBF33E9AE6F8B786B679E19BC8DFF" pageId="0" pageNumber="131">
<emphasis id="37C67C2585507384CB8D6D056671BF2F" bold="true" pageId="0" pageNumber="131">Figures 6-9.</emphasis>
<taxonomicName id="3F777831B502D0840D41695C4F4BF4B1" authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="62E0057EA43B042054EF4E19D9432335" italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
</taxonomicName>
species dorsal habitus.
<emphasis bold="true" pageId="0" pageNumber="131">6</emphasis>
<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia parva" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia parva</emphasis>
<emphasis id="F9676BA5949502731B228E3772BA1FDE" bold="true" pageId="0" pageNumber="131">6</emphasis>
<taxonomicName id="95B44CA1BB92056861913DB4153EF089" authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia parva" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="A5E6F7B5AF74D6764B35654D94DE0054" italics="true" pageId="0" pageNumber="131">Tiferonia parva</emphasis>
</taxonomicName>
, male paratype
<emphasis bold="true" pageId="0" pageNumber="131">7</emphasis>
<taxonomicName authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia leytensis</emphasis>
<emphasis id="50D507C4EC36702A3A422374F017CCF2" bold="true" pageId="0" pageNumber="131">7</emphasis>
<taxonomicName id="6992EED67DB797699776F10FF6990116" authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis">
<emphasis id="C2CB74513F598FB3C9D841A065E14FF9" italics="true" pageId="0" pageNumber="131">Tiferonia leytensis</emphasis>
</taxonomicName>
male holotype
<emphasis bold="true" pageId="0" pageNumber="131">8</emphasis>
<taxonomicName authorityName="Will" authorityYear="2020" baseAuthorityName="Straneo" baseAuthorityYear="1943" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia schoutedeni" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="schoutedeni">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia schoutedeni</emphasis>
<emphasis id="6DA1A2014FB136530A183C28B9878809" bold="true" pageId="0" pageNumber="131">8</emphasis>
<taxonomicName id="8C3CAA5F93A43D27AC382E03BF23B0BA" authorityName="Will" authorityYear="2020" baseAuthorityName="Straneo" baseAuthorityYear="1943" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia schoutedeni" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="schoutedeni">
<emphasis id="7BB6DDB649F6EC2A8BCF0FEAA713F1ED" italics="true" pageId="0" pageNumber="131">Tiferonia schoutedeni</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="131">9</emphasis>
<taxonomicName baseAuthorityName="Jedlicka" baseAuthorityYear="1935" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia brunnea" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia brunnea</emphasis>
<emphasis id="34FA396098695BB09B2BA5F9A347D1DA" bold="true" pageId="0" pageNumber="131">9</emphasis>
<taxonomicName id="DC406C46D6EF6B5267218411CCAA5E33" baseAuthorityName="Jedlicka" baseAuthorityYear="1935" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia brunnea" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis id="6CA6B4DB178C315829023408CDCF4EBD" italics="true" pageId="0" pageNumber="131">Tiferonia brunnea</emphasis>
</taxonomicName>
, male holotype.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="etymology">
<paragraph pageId="0" pageNumber="131">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="131">
<subSubSection id="7CF536F43889E57E701C0B3E1B96AD53" pageId="0" pageNumber="131" type="etymology">
<paragraph id="05E36D8ED158F753E21283CC8E5576C0" pageId="0" pageNumber="131">Etymology.</paragraph>
<paragraph id="53D20C4115C063BACDE058A0237208AC" pageId="0" pageNumber="131">
The specific epithet
<taxonomicName authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis">
<emphasis italics="true" pageId="0" pageNumber="131">leytensis</emphasis>
<taxonomicName id="AF9CCF87E68968FD7921F88B668AD59E" authorityName="Will" authorityYear="2020" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="leytensis" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="leytensis">
<emphasis id="4702B339F1B6CBAFC8FE945AE43A984D" italics="true" pageId="0" pageNumber="131">leytensis</emphasis>
</taxonomicName>
is based on the type locality and is treated as an adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="notes">
<paragraph pageId="0" pageNumber="131">Notes.</paragraph>
<paragraph pageId="0" pageNumber="131">
<subSubSection id="BD8182FDC895857B8EEED90E79AA2CA3" pageId="0" pageNumber="131" type="notes">
<paragraph id="E08EF292275C667DA8C38300A8B99F8B" pageId="0" pageNumber="131">Notes.</paragraph>
<paragraph id="948E63C436DB6F13CD99A79E21D42F94" pageId="0" pageNumber="131">
In his discussion of the species of
<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
<taxonomicName id="2A1B5264B69C057D852C6C0C67A6FE93" authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="E3ACF06BBFAFEC63959AC34C89CC1797" italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
</taxonomicName>
,
<bibRefCitation author="Darlington, PJ" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="0" pageNumber="131" pagination="321 - 564" refId="B5" refString="Darlington, PJ, 1962. The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini. Bulletin of the Museum of Comparative Zoology at Harvard College 126: 321 - 564" title="The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini." volume="126" year="1962">Darlington (1962</bibRefCitation>
<bibRefCitation id="84A3A3EB0360032BC87E01449608F0E4" author="Darlington, PJ" journalOrPublisher="Bulletin of the Museum of Comparative Zoology at Harvard College" pageId="0" pageNumber="131" pagination="321 - 564" refId="B5" refString="Darlington, PJ, 1962. The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini. Bulletin of the Museum of Comparative Zoology at Harvard College 126: 321 - 564" title="The Carabid beetles of New Guinea Part I. Cicindelinae, Carabinae, Harpalinae through Pterostichini." volume="126" year="1962">Darlington (1962</bibRefCitation>
: 561) states that
<taxonomicName lsidName="T. brunnea" pageId="0" pageNumber="131" rank="species" species="brunnea">
<emphasis italics="true" pageId="0" pageNumber="131">T. brunnea</emphasis>
<taxonomicName id="8DFDA8928301EFE94FED88EA7D1CF6B4" lsidName="T. brunnea" pageId="0" pageNumber="131" rank="species" species="brunnea">
<emphasis id="56182709C77EACD85EF85D8697C5EEF7" italics="true" pageId="0" pageNumber="131">T. brunnea</emphasis>
</taxonomicName>
from the Philippines is distinguished by having only a single pair of supraorbital setae and then states that he has a series of that species from Leyte Island. However, this appears to be an error. Among
<normalizedToken originalValue="Darlingtons">Darlington's</normalizedToken>
<normalizedToken id="F294544F95B499E942E3A3FA62914D32" originalValue="Darlingtons">Darlington's</normalizedToken>
specimens at the MCZ he has a series from Leyte Island, but they all have two pairs of supraorbital setae and are otherwise distinctly different from
<taxonomicName lsidName="T. brunnea" pageId="0" pageNumber="131" rank="species" species="brunnea">
<emphasis italics="true" pageId="0" pageNumber="131">T. brunnea</emphasis>
<taxonomicName id="565BBB293E2E1BE03C1A10FCC78BCBB5" lsidName="T. brunnea" pageId="0" pageNumber="131" rank="species" species="brunnea">
<emphasis id="D816FC7771F4037E95E5B04A962A862E" italics="true" pageId="0" pageNumber="131">T. brunnea</emphasis>
</taxonomicName>
. These specimens comprise the holotype and paratypes of
<taxonomicName lsidName="T. leytensis" pageId="0" pageNumber="131" rank="species" species="leytensis">
<emphasis italics="true" pageId="0" pageNumber="131">T. leytensis</emphasis>
<taxonomicName id="E31FB88A377A60443A30DA78CD05EA2C" lsidName="T. leytensis" pageId="0" pageNumber="131" rank="species" species="leytensis">
<emphasis id="FD474EB308280DCFA1DC889B52A9AA42" italics="true" pageId="0" pageNumber="131">T. leytensis</emphasis>
</taxonomicName>
.
</paragraph>

View file

@ -1,224 +1,223 @@
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<mods:title id="D50238363B09039D10C0C434AF909E93">Review of the Neotropical genus Rhyncholepta with descriptions of three new species-group taxa (Hemiptera, Heteroptera, Pentatomidae)</mods:title>
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<taxonomicName LSID="http://zoobank.org/DF08B4F4-0B65-4A2C-98FD-C299E534174B" class="Insecta" family="Pentatomidae" genus="Rhyncholepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhyncholepta henryi" order="Hemiptera" pageId="33" pageNumber="380" phylum="Arthropoda" rank="species" species="henryi">Rhyncholepta henryi</taxonomicName>
<taxonomicNameLabel pageId="33" pageNumber="380">sp. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="0BD5CA0275513E63037F5A19D58D6F65" pageId="33" pageNumber="380">sp. n.</taxonomicNameLabel>
Figs 6, 23, 36-37, 48-49, 60-61, 69, 75, 84-86, 92, 96
</paragraph>
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<subSubSection pageId="33" pageNumber="380" type="type locality">
<paragraph pageId="33" pageNumber="380">Type locality.</paragraph>
<paragraph pageId="33" pageNumber="380">
<subSubSection id="D06E13174B3619AF8E0E544F20960786" pageId="33" pageNumber="380" type="type locality">
<paragraph id="51B131C6D2DB035AD7BCC43A107ACF1C" pageId="33" pageNumber="380">Type locality.</paragraph>
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French Guiana, Roura Commune, Route de Kaw, Camp Caimans,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.569389">4°34'09.8&quot;N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.218193">52°13'05.5&quot;W</geoCoordinate>
<geoCoordinate id="E5CAAB72578A06E853924228A777E337" direction="north" orientation="latitude" precision="1" value="4.569389">4°34'09.8&quot;N</geoCoordinate>
<geoCoordinate id="4C0B10E75B27583BDC7B1436E8544FA9" direction="west" orientation="longitude" precision="1" value="-52.218193">52°13'05.5&quot;W</geoCoordinate>
, 320 m a.s.l.
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<paragraph pageId="33" pageNumber="380">Type material.</paragraph>
<paragraph pageId="33" pageNumber="380">
<subSubSection id="BBB54B6ABBC55F73CA0F2486CCE0CA47" pageId="33" pageNumber="380" type="type material">
<paragraph id="FC2E47E6738ED2050DE9ADBC1312CC63" pageId="33" pageNumber="380">Type material.</paragraph>
<paragraph id="D689FE01CD922F8DCD33C5080743FFA8" pageId="33" pageNumber="380">
Holotype: ♂ (Figs 6, 23, 36-37, 48-49, 60-61, 69, 75), &quot;GUYANE FR., Rt. de Kaw / Camp Caimans, 320 m a.s.l. /
<geoCoordinate direction="north" orientation="latitude" precision="5" value="4.5694">04.5694N</geoCoordinate>
<geoCoordinate id="48E797BCD51BDB73BED6DE3B82E953DF" direction="north" orientation="latitude" precision="5" value="4.5694">04.5694N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-52.2182">52.2182W</geoCoordinate>
<geoCoordinate id="DE19B9D4719CD604AE0A3D56466AD58D" direction="west" orientation="longitude" precision="5" value="-52.2182">52.2182W</geoCoordinate>
/ 11.-19.i.2016, S. MURZIN lgt. [printed, white label] // COLLECTIO / NATIONAL MUSEUM / Praha, Czech Republic [printed, white label] // ♂ [printed, white label] // HOLOTYPUS / RHYNCHOLEPTA / HENRYI / sp. nov. / det. Kment, Eger, Rider 2017 [printed, red label]&quot; (NMPC). The holotype is card-mounted, with the detached left distiflagellum glued on the same piece of card and the detached genital capsule and abdominal segment VIII glued separately on a small piece of card.
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<paragraph id="CAEDBD49EA37E2EDF897BE6C2F746FF4" pageId="33" pageNumber="380">
Paratypes: FRENCH GUIANA: Cayenne Arrondissement: Kourou Commune: Montagne des Signes near Kourou [ca.
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<geoCoordinate id="31EF459A02632EE6B37FB835BC9BE92A" direction="north" orientation="latitude" precision="1" value="5.091899">5.091899°N</geoCoordinate>
<geoCoordinate id="06A7FAF9E454A1887E1D560987B56BF0" direction="west" orientation="longitude" precision="1" value="-52.699482">52.699481°W</geoCoordinate>
], collected at mercury vapor light, 3.vi.1986, 2 ♂♂, E. C. Riley and D. A. Rider lgt. (DARC). Roura Commune: 27 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5686">4°34.116'N</geoCoordinate>
<geoCoordinate id="0451A8875BE7CF29D0D116194B9864C7" direction="north" orientation="latitude" precision="1" value="4.5686">4°34.116'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.21023">52°12.614'W</geoCoordinate>
<geoCoordinate id="7E3519C721588F89D915B8FB7B36AD0F" direction="west" orientation="longitude" precision="1" value="-52.21023">52°12.614'W</geoCoordinate>
, MV Light, 5.ii.2010, 1 ♂, J. E. Eger lgt. (JEEC); 32 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5601997">4°33.612'N</geoCoordinate>
<geoCoordinate id="9CBB6799959A220C76778C79A34BA24D" direction="north" orientation="latitude" precision="1" value="4.5601997">4°33.612'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.189167">52°11.350'W</geoCoordinate>
<geoCoordinate id="14DA7BC9590C203BBC302B0B0B53E1C0" direction="west" orientation="longitude" precision="1" value="-52.189167">52°11.350'W</geoCoordinate>
, 287 m a.s.l., MV Light, 15.ii.2010, 1 ♂, J. E. Eger lgt. (JEEC); 33 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.568917">4°34.135'N</geoCoordinate>
<geoCoordinate id="D60AD8C33CF352A635B06002C5C94ED9" direction="north" orientation="latitude" precision="1" value="4.568917">4°34.135'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.185833">52°11.150'W</geoCoordinate>
<geoCoordinate id="57B9F669C8BDC92FB73BA14F88A8E593" direction="west" orientation="longitude" precision="1" value="-52.185833">52°11.150'W</geoCoordinate>
, 227 m a.s.l., MV Light, 12.-13.iv.2007, 6 ♂♂, D. G. Hall and J. E. Eger lgt. (JEEC); Amazone Nature Lodge, 30 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5594997">4°33.570'N</geoCoordinate>
<geoCoordinate id="6F43C515CBFCD7B7C382CFF4E72B19B3" direction="north" orientation="latitude" precision="1" value="4.5594997">4°33.570'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.20722">52°12.433'W</geoCoordinate>
<geoCoordinate id="D77127B1B634D82DCF433013C5DBC3EF" direction="west" orientation="longitude" precision="1" value="-52.20722">52°12.433'W</geoCoordinate>
, 300 m a.s.l., UV Light, 10.-18.iv.2007, 1 ♂, D. G. Hall and J. E. Eger lgt. (JEEC); the same locality, MV Lights, 4.-15.i.2016, 4 ♂♂, J. Eger, R. Morris and J. Wappes lgt. (JEEC); Camp Caiman,
<geoCoordinate direction="north" orientation="latitude" precision="55" value="4.569">4.569°N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-52.218">52.218°W</geoCoordinate>
<geoCoordinate id="D27B7602AD63D309EBEEC9A921D9DDEE" direction="north" orientation="latitude" precision="55" value="4.569">4.569°N</geoCoordinate>
<geoCoordinate id="8F8E580D9C4B6F1CA24AA2E2846E8E3A" direction="west" orientation="longitude" precision="55" value="-52.218">52.218°W</geoCoordinate>
, 260 m a.s.l., 8.-31.i.2018, 3 ♂♂, S. Murzin lgt. (NMPC); Entomotech Lodge, 30 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5594997">4°33.570'N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.20722">52°12.433'W</geoCoordinate>
<geoCoordinate id="3CD28BA1B18EC9D406218519BBE0450A" direction="north" orientation="latitude" precision="1" value="4.5594997">4°33.570'N</geoCoordinate>
<geoCoordinate id="EEFD650B087B18DD4BF23D7EB28DEAF8" direction="west" orientation="longitude" precision="1" value="-52.20722">52°12.433'W</geoCoordinate>
, 300 m a.s.l., MV Light, 17.xi.2004, 1 ♂, xi.2004-ii.2005, 2 ♂♂, 17.i.2005, 1 ♂, F. Goubert lgt. (JEEC); Highway N2 to Regina, 45 km S of Cayenne [ca.
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.5313888">4°31'53&quot;N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-52.37222">52°22'20&quot;W</geoCoordinate>
<geoCoordinate id="C7601BE974AE362069A48A587D446298" direction="north" orientation="latitude" precision="15" value="4.5313888">4°31'53&quot;N</geoCoordinate>
<geoCoordinate id="017F3521A96D77348B3925EDF9E7DDEF" direction="west" orientation="longitude" precision="15" value="-52.37222">52°22'20&quot;W</geoCoordinate>
], collected at mercury vapor light, 31.v.1986, 2 ♂♂, E. G. Riley and D. A. Rider lgt. (DARC); Montagne des Chevaux RN2 km 22 [ca.
<geoCoordinate direction="north" orientation="latitude" precision="5" value="4.7216">4.7216°N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-52.3073">52.3073°W</geoCoordinate>
<geoCoordinate id="D1289AFE35CD71256491A017974FC8D9" direction="north" orientation="latitude" precision="5" value="4.7216">4.7216°N</geoCoordinate>
<geoCoordinate id="3A16346D0E9489A8A923F7701F784946" direction="west" orientation="longitude" precision="5" value="-52.3073">52.3073°W</geoCoordinate>
], automatic light trap (white LED), 3.i.2013, 1 ♂, SEAG leg. (MNHN). Saint-Laurent-du-Maroni Arrondissement: Mana Commune:
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Trinité">Trinite</normalizedToken>
<normalizedToken id="61F7618C53BD97654613C8ADDF92551F" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="D2C3DE4C51E85C0DF759B3177B09A4E1" originalValue="Trinité">Trinite</normalizedToken>
[ca.
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.0716667">4°04'18&quot;N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-52.555">52°33'18&quot;W</geoCoordinate>
<geoCoordinate id="4A0B0C689EE676AE32C5232C9CE79663" direction="north" orientation="latitude" precision="15" value="4.0716667">4°04'18&quot;N</geoCoordinate>
<geoCoordinate id="5E7D5A7698B50D61A06FFE8FB462A7A8" direction="west" orientation="longitude" precision="15" value="-52.555">52°33'18&quot;W</geoCoordinate>
], Drop Zone Aya, UV light trap, 30.v.2012, 1 ♂, SEAG leg. (RLFF).
<normalizedToken originalValue="Saül">Sauel</normalizedToken>
<normalizedToken id="A6287BEAA314759C4D8619A9CDBB8FDF" originalValue="Saül">Sauel</normalizedToken>
Commune:
<normalizedToken originalValue="Belvédère">Belvedere</normalizedToken>
<normalizedToken id="1D8A9F8C27F080CED6E996B17107BB68" originalValue="Belvédère">Belvedere</normalizedToken>
[ca.
<geoCoordinate direction="north" orientation="latitude" precision="555" value="2.41">2.41°N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="5555" value="-53.1">53.1°W</geoCoordinate>
<geoCoordinate id="5EB8FD5CFDF0017D760E470E0BDDFCC2" direction="north" orientation="latitude" precision="555" value="2.41">2.41°N</geoCoordinate>
<geoCoordinate id="139538EE1B7E7611064A9E1397BA7F69" direction="west" orientation="longitude" precision="5555" value="-53.1">53.1°W</geoCoordinate>
], automatic light trap (blue LED), 5.vii.2017, 1 ♂, SEAG leg. (RLFF). - All the paratypes are bearing the following identification label: &quot;PARATYPUS / RHYNCHOLEPTA / HENRYI / sp. nov. / det. Kment, Eger, Rider 2017 [printed, yellow label]&quot;.
</paragraph>
</subSubSection>
<subSubSection lastPageId="34" lastPageNumber="381" pageId="33" pageNumber="380" type="materials_examined">
<paragraph pageId="33" pageNumber="380">Additional material examined</paragraph>
<paragraph lastPageId="34" lastPageNumber="381" pageId="33" pageNumber="380">
<subSubSection id="D02F30AE68ACEADBE63B3A7EA30BED63" lastPageId="34" lastPageNumber="381" pageId="33" pageNumber="380" type="materials_examined">
<paragraph id="79AD43C35136E9B25BA3E150FD19C26F" pageId="33" pageNumber="380">Additional material examined</paragraph>
<paragraph id="42FD97A2217BD8EE51B0402B9A32C480" lastPageId="34" lastPageNumber="381" pageId="33" pageNumber="380">
(females tentatively identified as
<taxonomicName genus="Rh." lsidName="Rh. henryi" pageId="33" pageNumber="380" rank="species" species="henryi">Rh. henryi</taxonomicName>
<taxonomicName id="C36D279839AA82B0EBA7E96AD6C6E6AE" lsidName="Rh. henryi" pageId="33" pageNumber="380" rank="species" species="henryi">Rh. henryi</taxonomicName>
). FRENCH GUIANA: Cayenne Arrondissement: Kourou Commune: Montagne des Signes near Kourou [ca.
<geoCoordinate direction="north" orientation="latitude" precision="1" value="5.091899">5.091899°N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.699482">52.699481°W</geoCoordinate>
<geoCoordinate id="05A1C7FD6EF580A7AF61BF5CE740C018" direction="north" orientation="latitude" precision="1" value="5.091899">5.091899°N</geoCoordinate>
<geoCoordinate id="8ADC9377B01BFFCE33FD5D0DC95E0EA4" direction="west" orientation="longitude" precision="1" value="-52.699482">52.699481°W</geoCoordinate>
], collected at mercury vapor light, 3.vi.1986, 1 ♀, E. C. Riley and D. A. Rider lgt. (DARC).
<normalizedToken originalValue="Montsinéry-Tonnegrande">Montsinery-Tonnegrande</normalizedToken>
<normalizedToken id="FB30F0D7C714F626319DCD1A2009CE6E" originalValue="Montsinéry-Tonnegrande">Montsinery-Tonnegrande</normalizedToken>
Commune: 8 km W of Risquetout,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.9182835">4°55.097'N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.552017">52°33.121'W</geoCoordinate>
<geoCoordinate id="ECEA9F9B77246600390D5258E2B02FC1" direction="north" orientation="latitude" precision="1" value="4.9182835">4°55.097'N</geoCoordinate>
<geoCoordinate id="D0D9006138C059FB31A61DD60DC4B05F" direction="west" orientation="longitude" precision="1" value="-52.552017">52°33.121'W</geoCoordinate>
, 45 m a.s.l., MV Light, 15.iv.2007, 3 ♀♀, D. G. Hall and J. E. Eger lgt. (JEEC). Roura Commune: 21 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.601917">4°36.115'N</geoCoordinate>
<geoCoordinate id="495CE5F790DE4D7241E7D4739EAD17F5" direction="north" orientation="latitude" precision="1" value="4.601917">4°36.115'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.2662">52°15.972'W</geoCoordinate>
<geoCoordinate id="3C94126F15426B26E1E1FE11E2CD7AAC" direction="west" orientation="longitude" precision="1" value="-52.2662">52°15.972'W</geoCoordinate>
, MV Light, 6.-7.ii.2010, 1 ♀, J. E. Eger lgt. (JEEC); 28 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5708666">4°34.252'N</geoCoordinate>
<geoCoordinate id="0659498628CF7D6649820CC475ED2C6C" direction="north" orientation="latitude" precision="1" value="4.5708666">4°34.252'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.213284">52°12.797'W</geoCoordinate>
<geoCoordinate id="80058B9788A6ADECEE835FAF459595CE" direction="west" orientation="longitude" precision="1" value="-52.213284">52°12.797'W</geoCoordinate>
, 306 m a.s.l., MV Light, 17.ii.2010, 1 ♀, J. E. Eger lgt. (JEEC); 32 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5601997">4°33.612'N</geoCoordinate>
<geoCoordinate id="D6CA2CB08BFB40269D5E02925BAA6BF8" direction="north" orientation="latitude" precision="1" value="4.5601997">4°33.612'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.189167">52°11.350'W</geoCoordinate>
<geoCoordinate id="D98418C74FB4A6DAF6459AEEC94CCC55" direction="west" orientation="longitude" precision="1" value="-52.189167">52°11.350'W</geoCoordinate>
, 287 m a.s.l., MV Light, 15.ii.2010, 1 ♀, J. E. Eger lgt. (JEEC); 33 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.568917">4°34.135'N</geoCoordinate>
<geoCoordinate id="A715D61746D0963A571B33D54AFAFD33" direction="north" orientation="latitude" precision="1" value="4.568917">4°34.135'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.185833">52°11.150'W</geoCoordinate>
<geoCoordinate id="E64016A2E888F34E7B79D95C7EFED447" direction="west" orientation="longitude" precision="1" value="-52.185833">52°11.150'W</geoCoordinate>
, 227 m a.s.l., MV Light, 12.-13.iv.2007, 7 ♀♀, D. G. Hall and J. E. Eger lgt. (JEEC); Amazone Nature Lodge, 30 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5594997">4°33.570'N</geoCoordinate>
<geoCoordinate id="F00DE9D3753DC1CE5BAC34B3C7170416" direction="north" orientation="latitude" precision="1" value="4.5594997">4°33.570'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.20722">52°12.433'W</geoCoordinate>
<geoCoordinate id="95ACDF470B25715B13BE99DB2E6CD13D" direction="west" orientation="longitude" precision="1" value="-52.20722">52°12.433'W</geoCoordinate>
, 300 m a.s.l., UV
<pageBreakToken pageId="34" pageNumber="381" start="start">Light</pageBreakToken>
<pageBreakToken id="A36D7B212EB586BFE2A0F502CE3EB3F3" pageId="34" pageNumber="381" start="start">Light</pageBreakToken>
, 4.-15.i.2016, 1 ♀, J. Eger, R. Morris and J. Wappes lgt. (JEEC); Camp Caiman,
<geoCoordinate direction="north" orientation="latitude" precision="55" value="4.569">4.569°N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-52.218">52.218°W</geoCoordinate>
<geoCoordinate id="6AE3ABB983BE2B16DB853808ECBDF0A2" direction="north" orientation="latitude" precision="55" value="4.569">4.569°N</geoCoordinate>
<geoCoordinate id="BDF52110D30500A958AE5DA05FDE8F86" direction="west" orientation="longitude" precision="55" value="-52.218">52.218°W</geoCoordinate>
, 260 m a.s.l., 8.-31.i.2018, 1 ♀, S. Murzin lgt. (NMPC); Entomotech Lodge, 30 km SE Roura on Kaw Rd.,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="4.5594997">4°33.570'N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-52.20722">52°12.433'W</geoCoordinate>
<geoCoordinate id="A61A8A98E1701DA9D3D200E39472C67E" direction="north" orientation="latitude" precision="1" value="4.5594997">4°33.570'N</geoCoordinate>
<geoCoordinate id="D172BE5519AE5C5B1990FE36B6EFF84C" direction="west" orientation="longitude" precision="1" value="-52.20722">52°12.433'W</geoCoordinate>
, 300 m a.s.l., MV Light, 1.-12.xii.2002, 1 ♀, J. E. Eger lgt. (DARC); Highway D6 to Kaw, 33.5 km SE of Roura [ca.
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.5463886">4°32'47&quot;N</geoCoordinate>
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-52.144726">52°08'41&quot;W</geoCoordinate>
<geoCoordinate id="FB7A74812326DEE910A6BD0E05DEDD9D" direction="north" orientation="latitude" precision="15" value="4.5463886">4°32'47&quot;N</geoCoordinate>
<geoCoordinate id="3514B62E82438DEC4E102C821E663EB0" direction="west" orientation="longitude" precision="15" value="-52.144726">52°08'41&quot;W</geoCoordinate>
], 10.ii.1986, 1 ♀, G. Tavakilian lgt. (DARC).
</paragraph>
</subSubSection>
<subSubSection pageId="34" pageNumber="381" type="diagnosis">
<paragraph pageId="34" pageNumber="381">Diagnosis.</paragraph>
<paragraph pageId="34" pageNumber="381">
<subSubSection id="A35BA9C74E82E2B3BB999558588248A2" pageId="34" pageNumber="381" type="diagnosis">
<paragraph id="E3EBCDF2EE950E7EF9E494467025B90B" pageId="34" pageNumber="381">Diagnosis.</paragraph>
<paragraph id="C36F440BE9B013A0B22FF8C89C3389EA" pageId="34" pageNumber="381">
Coloration, structure of head, thorax and pregenital abdomen, and vestiture as in other species of the genus (see redescription of
<taxonomicName class="Insecta" family="Pentatomidae" genus="Rhyncholepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhyncholepta" order="Hemiptera" pageId="34" pageNumber="381" phylum="Arthropoda" rank="genus">Rhyncholepta</taxonomicName>
<taxonomicName id="B2A6AEA4CE8569D7958FC002AC40F88A" genus="Rhyncholepta" lsidName="Rhyncholepta" pageId="34" pageNumber="381" rank="genus">Rhyncholepta</taxonomicName>
above) except for the following characters.
</paragraph>
<paragraph pageId="34" pageNumber="381">Apex of scutellum with anteapical black V-shaped stripe well developed (Figure 23). Apical V-shaped callosity thin, narrowly delineating margins of scutellum apex, branches forming more than half of length, tip of scutellum lacking conspicuous triangular callosity (Figure 23).</paragraph>
<paragraph pageId="34" pageNumber="381">
<paragraph id="E581EF8624AACE0B14EEE29D2409E97E" pageId="34" pageNumber="381">Apex of scutellum with anteapical black V-shaped stripe well developed (Figure 23). Apical V-shaped callosity thin, narrowly delineating margins of scutellum apex, branches forming more than half of length, tip of scutellum lacking conspicuous triangular callosity (Figure 23).</paragraph>
<paragraph id="34DDA742B2DE3D93CAE37E77A88D3DD1" pageId="34" pageNumber="381">
Male genitalia. Genital capsule in ventral view only slightly constricted lateroapically, posterolateral angles prominent (Figs 36, 60); dorsal wall at base of posterolateral angles shallowly impressed (Figs 60, 69). Ventral rim in ventral view truncate apically, slightly notched medially (Figs 36-37), posterior hypandrial projections visible and caudal, together with ventral rim forming wide T (Figs 36-37); bases of lateral projections also visible (Figure 37). Hypandrium in posterior view with three pairs of projections: posterior ones caudal, short, flat, narrowly rounded apically with small rounded projection anteapically on posterior margin (Figs 48-49); lateral projections very narrow, directed posterolaterad, apices spinose, curved upwards (Figs 48-49); anterior projections steeply sloping downwards, appearing narrowly triangular in dorsal view, acutangulate apically (Figs 48-49). Anterior hypandrial projections in most exposed (dorso-posterolateral) view appearing nearly triangular with dorsal margin nearly straight, apically with sharp, straight, flattened spine (Figure 75: ap); posterior and lateral projections forming acute angle (Figure 75: a); posterior projection appearing straight, wide, widely rounded apically in this view (Figure 75: pp); lateral projection narrowing, shortly bent downwards apically (Figure 75: lp). Anterior hypandrial projections narrowing anteriad towards apices in dorsal view, apices straight, each forming large claw-like spine curved downwards (Figs 60, 61: ap), posterior projections short, flat, apically rounded (Figs 60, 61: pp); lateral projections directed laterad, each narrowing to sharp spine, curved anteriad apically (Figs 60, 61: lp). Phallus (Figs 84-86) with basal conjunctival sclerite drop-shaped (Figure 85: cjs) and aedeagus only slightly sinuate apically (Figure 85: ae), very similar to that of
<taxonomicName genus="Rh." lsidName="Rh. meinanderi" pageId="34" pageNumber="381" rank="species" species="meinanderi">Rh. meinanderi</taxonomicName>
<taxonomicName id="CA227D51D4B0B6D124648A47F8BF6119" lsidName="Rh. meinanderi" pageId="34" pageNumber="381" rank="species" species="meinanderi">Rh. meinanderi</taxonomicName>
.
</paragraph>
<paragraph pageId="34" pageNumber="381">
<paragraph id="95198A563C8497EDF95D8054ED616DC9" pageId="34" pageNumber="381">
Female genitalia. Posterior edges of laterotergites VIII suddenly attenuated apically, posteriorly about as long as laterotergites IX (Figure 92), indistinguishable from
<taxonomicName genus="Rh." lsidName="Rh. grandicallosa" pageId="34" pageNumber="381" rank="species" species="grandicallosa">Rh. grandicallosa</taxonomicName>
<taxonomicName id="7CEA68835C9998D39352C63FFCF3DCCD" lsidName="Rh. grandicallosa" pageId="34" pageNumber="381" rank="species" species="grandicallosa">Rh. grandicallosa</taxonomicName>
.
</paragraph>
<paragraph pageId="34" pageNumber="381">Measurements. Table 1. Measurements of holotype (in mm): Body length 12.39, body length to segment VII 10.91, head length 2.55, head width 2.27, interocular width 1.15, length of each antennomeres: I - 0.86, IIa - 1.34, IIb - 2.61, III - 3.46, IV - 2.95, pronotum length 2.47, pronotum width 8.05, scutellum length 4.37, scutellum width 3.62.</paragraph>
<paragraph id="D0BABC2187DF7E844B28CF392C09543F" pageId="34" pageNumber="381">Measurements. Table 1. Measurements of holotype (in mm): Body length 12.39, body length to segment VII 10.91, head length 2.55, head width 2.27, interocular width 1.15, length of each antennomeres: I - 0.86, IIa - 1.34, IIb - 2.61, III - 3.46, IV - 2.95, pronotum length 2.47, pronotum width 8.05, scutellum length 4.37, scutellum width 3.62.</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="382" type="differential diagnosis">
<paragraph pageId="35" pageNumber="382">
<pageBreakToken pageId="35" pageNumber="382" start="start">Differential</pageBreakToken>
<subSubSection id="31D4D994ACA106782CF6FB639D49BD1E" pageId="35" pageNumber="382" type="differential diagnosis">
<paragraph id="98E420889134DDDD023A27FA49F2DE83" pageId="35" pageNumber="382">
<pageBreakToken id="58C256E892A712B8CE82376362438EF4" pageId="35" pageNumber="382" start="start">Differential</pageBreakToken>
diagnosis.
</paragraph>
<paragraph pageId="35" pageNumber="382">
<paragraph id="D4AD788DDF8AD4DD6C6B6496AA019F5D" pageId="35" pageNumber="382">
See above key. All specimens differing from the majority of specimens of
<taxonomicName genus="Rh." lsidName="Rh. g." pageId="35" pageNumber="382" rank="subSpecies" species="g." subSpecies="grandicallosa">Rh. g. grandicallosa</taxonomicName>
<taxonomicName id="0D9C06FA5B0ECF1DBA182BD18ADE2A4E" lsidName="Rh. g." pageId="35" pageNumber="382" rank="subSpecies" species="g." subSpecies="grandicallosa">Rh. g. grandicallosa</taxonomicName>
by complete black V-shaped band anteapically on scutellum (character does not work for all specimens of
<taxonomicName genus="Rh." lsidName="Rh. g." pageId="35" pageNumber="382" rank="subSpecies" species="g." subSpecies="grandicallosa">Rh. g. grandicallosa</taxonomicName>
<taxonomicName id="111FDC8A1053C72227EC84910DC297A9" lsidName="Rh. g." pageId="35" pageNumber="382" rank="subSpecies" species="g." subSpecies="grandicallosa">Rh. g. grandicallosa</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="382" type="etymology">
<paragraph pageId="35" pageNumber="382">Etymology.</paragraph>
<paragraph pageId="35" pageNumber="382">
<subSubSection id="77A223B9894834276E524DCC0F3377C6" pageId="35" pageNumber="382" type="etymology">
<paragraph id="9D8D8DBFF58E055BB96623F677A47064" pageId="35" pageNumber="382">Etymology.</paragraph>
<paragraph id="322212518BBA77D9733770A30A4C05C6" pageId="35" pageNumber="382">
We are pleased to dedicate the new species to our colleague, Thomas J. Henry, an excellent specialist in
<taxonomicName lsidName="Heteroptera" pageId="35" pageNumber="382" rank="subOrder" subOrder="Heteroptera">Heteroptera</taxonomicName>
<taxonomicName id="C3D4A5CFC5C4DE70C4F14FEB1BC8EB6F" lsidName="Heteroptera" pageId="35" pageNumber="382" rank="subOrder" subOrder="Heteroptera">Heteroptera</taxonomicName>
and curator of the USNM, on the occasion of his 70th birthday. We much appreciate his generosity in helping visitors with literature or specimens in the USNM, including hosting colleagues in his home when they visited the collection.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="382" type="bionomics">
<paragraph pageId="35" pageNumber="382">Bionomics.</paragraph>
<paragraph pageId="35" pageNumber="382">
<subSubSection id="2E30BCDAF7500E470031ED9F76501DDB" pageId="35" pageNumber="382" type="bionomics">
<paragraph id="853066BEB3BA4A556478A5CD59F72227" pageId="35" pageNumber="382">Bionomics.</paragraph>
<paragraph id="6399B9BC92430DC50C8F9204571E1618" pageId="35" pageNumber="382">
Most specimens were collected by various types of light traps (UV light, mercury vapor light, white and blue LED) in or adjacent to dense forests. This species has never been collected by hand catching, sweeping or beating vegetation during the day or night (JE Eger, pers. observ.).
<taxonomicName class="Insecta" family="Pentatomidae" genus="Rhyncholepta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rhyncholepta henryi" order="Hemiptera" pageId="35" pageNumber="382" phylum="Arthropoda" rank="species" species="henryi">Rhyncholepta henryi</taxonomicName>
<taxonomicName id="742B3258ED2D0387BF353BA856245091" genus="Rhyncholepta" lsidName="Rhyncholepta henryi" pageId="35" pageNumber="382" rank="species" species="henryi">Rhyncholepta henryi</taxonomicName>
was found from November to February and from April to June, with most specimens collected in April (Figure 96).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="382" type="distribution">
<paragraph pageId="35" pageNumber="382">Distribution</paragraph>
<paragraph pageId="35" pageNumber="382">(Figs 99-100). French Guiana (present paper).</paragraph>
<subSubSection id="23FAB8714AF72EB0164F835221D4C8E2" pageId="35" pageNumber="382" type="distribution">
<paragraph id="74EDCB064C1975B187106AA070530120" pageId="35" pageNumber="382">Distribution</paragraph>
<paragraph id="D7439CC26869B468B51C2A475C31794F" pageId="35" pageNumber="382">(Figs 99-100). French Guiana (present paper).</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,565 +1,565 @@
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">Figs 10A-D</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">, 11A-F</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution records of Leviana dimidiata comb. nov. and L. cincinnata sp. nov." httpUri="https://binary.pensoft.net/fig/714526" pageId="0" pageNumber="103">, 9</figureCitation>
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<figureCitation id="748D0360B8ABD26D038D82CCE17A1FA6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">Figs 10A-D</figureCitation>
<figureCitation id="7CAED68DC1836ECDE0852ED1B90273F5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">, 11A-F</figureCitation>
<figureCitation id="27D0D60CF4D2C0188EDAC92D931CF1CA" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution records of Leviana dimidiata comb. nov. and L. cincinnata sp. nov." httpUri="https://binary.pensoft.net/fig/714526" pageId="0" pageNumber="103">, 9</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="103">Types material.</paragraph>
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<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Holotype</emphasis>
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<paragraph id="C33D0F54FD1B6C4E5A34F28998817BCC" pageId="0" pageNumber="103">Types material.</paragraph>
<paragraph id="9F9DCA184FCA9898E7EEC0CFDCD0F015" pageId="0" pageNumber="103">
<emphasis id="0980B16A7EBDB03202FA5EF2732D483A" bold="true" italics="true" pageId="0" pageNumber="103">Holotype</emphasis>
: Male, Bellenden Ker Range, North Queensland, Cable Tower 3 (
<geoCoordinate degrees="17" direction="south" minutes="16" orientation="latitude" precision="925" value="-17.266666">17°16'S</geoCoordinate>
<geoCoordinate id="B549C314AD241FDCE212233BD43F6976" degrees="17" direction="south" minutes="16" orientation="latitude" precision="925" value="-17.266666">17°16'S</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="51" orientation="longitude" precision="925" value="145.85">145°51'E</geoCoordinate>
<geoCoordinate id="56DD19D56CE37B2B4B3DC1E4097A3F9E" degrees="145" direction="east" minutes="51" orientation="longitude" precision="925" value="145.85">145°51'E</geoCoordinate>
, Queensland, Australia), 25-31 October 1981, sweeping, Earthwatch/Qld Museum (QM S26364).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="etymology">
<paragraph pageId="0" pageNumber="103">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="103">
<subSubSection id="BE9ACF9CD81D5E88A950B46493C9268A" pageId="0" pageNumber="103" type="etymology">
<paragraph id="A36F83D838533D9C4F079F6BF43228A1" pageId="0" pageNumber="103">Etymology.</paragraph>
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The specific epithet is an adjective in apposition (
<emphasis italics="true" pageId="0" pageNumber="103">cincinnatus</emphasis>
<emphasis id="473F9486D45336D1485B03797C8AD92A" italics="true" pageId="0" pageNumber="103">cincinnatus</emphasis>
Latin - curly) and refers to the leaf-curling behaviour of this species and other species in the genus.
</paragraph>
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<subSubSection pageId="0" pageNumber="103" type="materials_examined">
<paragraph pageId="0" pageNumber="103">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="103">
<materialsCitation accessionNumber="S47286" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Bartle Frere, W Base" country="Australia" latitude="-17.383333" location="Queensland" longLatPrecision="1279" longitude="145.76666" specimenCount="2" specimenCount-female="2" stateProvince="Queensland">
<collectingCountry name="Australia">Australia</collectingCountry>
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<paragraph id="CA517D2CCE34A612FAE1AED1EA3524CB" pageId="0" pageNumber="103">Other material examined.</paragraph>
<paragraph id="3CD35BEAE928E5CFA3AAB60E0C7E2326" pageId="0" pageNumber="103">
<materialsCitation id="FF244F88EA959C526C55F006B3D009FD" accessionNumber="S47286" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Bartle Frere, W Base" country="Australia" latitude="-17.383333" location="Queensland" longLatPrecision="1279" longitude="145.76666" specimenCount="2" specimenCount-female="2" stateProvince="Queensland">
<collectingCountry id="9E3100EA7845270E5F04A5E452B40155" name="Australia">Australia</collectingCountry>
:
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">
<collectingRegion country="Australia" name="Queensland">Queensland</collectingRegion>
<emphasis id="4082296A0C26B0D8184CA01FFBDA6E2C" bold="true" italics="true" pageId="0" pageNumber="103">
<collectingRegion id="389216C12ACFDBE4FC2CE80131A30E11" country="Australia" name="Queensland">Queensland</collectingRegion>
</emphasis>
:
<specimenCount type="female">2 females</specimenCount>
<specimenCount id="802FEC2A4064E64BDD2804D6DDDC5F33" type="female">2 females</specimenCount>
,
<collectorName>Bartle Frere</collectorName>
<collectorName id="2B17620DD8ADDF80083B16C79486608F">Bartle Frere</collectorName>
,
<collectorName>W Base</collectorName>
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,
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,
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, 0.5 km
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7,
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<geoCoordinate id="7621FDF3203899C2241ED6C38922DA76" degrees="17" direction="south" minutes="16" orientation="latitude" precision="925" value="-17.266666">17°16'S</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="51" orientation="longitude" precision="925" value="145.85">145°51'E</geoCoordinate>
<geoCoordinate id="14C5CB33DF54FD0D267235C44CD26FA1" degrees="145" direction="east" minutes="51" orientation="longitude" precision="925" value="145.85">145°51'E</geoCoordinate>
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,
<collectorName>Bellenden Ker</collectorName>
<collectorName id="2CB1AA667B54FDD33CCE31BFE47B7210">Bellenden Ker</collectorName>
,
<collectorName>Cable Tower</collectorName>
<collectorName id="E3AD859EDEFC3C09959F457414A3303B">Cable Tower</collectorName>
3,
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<geoCoordinate id="3DB7CF1274327E476E37A6500DE25A72" degrees="17" direction="south" minutes="16" orientation="latitude" precision="925" value="-17.266666">17°16'S</geoCoordinate>
,
<geoCoordinate degrees="145" direction="east" minutes="51" orientation="longitude" precision="925" value="145.85">145°51'E</geoCoordinate>
<geoCoordinate id="EABB73BD085FBBBFDF12B59AB56AB85D" degrees="145" direction="east" minutes="51" orientation="longitude" precision="925" value="145.85">145°51'E</geoCoordinate>
(QM
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<specimenCount id="99814855244FB00A38881E9CC015450C" type="female">1 female</specimenCount>
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, same, same locality (QM)
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<specimenCount id="39AB187907C48697BC483C68D5D63D27" type="female">1 female</specimenCount>
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<quantity id="414A93F3CE367FE4F798E923EA1F058A" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
W Edmonton
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,
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<geoCoordinate id="754F56CBC7F64805476F05334B65FCCB" degrees="17" direction="south" minutes="02" orientation="latitude" precision="925" value="-17.033333">17°02'S</geoCoordinate>
,
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(QM
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<specimenCount id="BE51D3A3A6A1668E4675CD80A6B5D2A5" type="male">1 male</specimenCount>
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summit,
<locationDeviation location="Mosman">
<quantity metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
<locationDeviation id="D525FF3BAB90AAE4D465C6A9826A0F09" location="Mosman">
<quantity id="C38010715B0301B80AB030F1E82DFEC4" metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
SW Mosman
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,
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<geoCoordinate id="587EE0919B2D98508B488F8BB3FF3E82" degrees="16" direction="south" minutes="30" orientation="latitude" precision="925" value="-16.5">16°30'S</geoCoordinate>
,
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<geoCoordinate id="F3003CD659EF590B1AF61E80FCA8AB8C" degrees="145" direction="east" minutes="19" orientation="longitude" precision="925" value="145.31667">145°19'E</geoCoordinate>
(QM
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<specimenCount type="female">1 female</specimenCount>
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<specimenCount id="8BBD7AB012C27C4DBD2DCCD168F19614" type="female">1 female</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:4D4378045BE959FCBD0B4105A37547C6:6470B0EB5DD5A26A6B1B296E7A6BB191" country="Australia" latitude="-16.716667" longLatPrecision="1281" longitude="145.61667" name="Mt Formatine South" stateProvince="Queensland">Mt Formatine South</location>
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,
<locationDeviation location="Kuranda">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
<locationDeviation id="DC0EBFA289D1772BF614209C9D6E1893" location="Kuranda">
<quantity id="EB1D898216E2A83430A159A2A94246D5" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
N Kuranda
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,
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,
<geoCoordinate degrees="145" direction="east" minutes="37" orientation="longitude" precision="925" value="145.61667">145°37'E</geoCoordinate>
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(QM
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SE, via
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,
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<quantity id="906FEFBFF4579127F1F56B26D5FA75AD" metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
N,
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,
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,
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</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="diagnosis">
<paragraph pageId="0" pageNumber="103">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="103">
<taxonomicName authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" kingdom="Animalia" lsidName="Leviana cincinnata" order="Araneae" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis italics="true" pageId="0" pageNumber="103">Leviana cincinnata</emphasis>
<subSubSection id="E5673FF98026F9DD2A86A8546FF46D74" pageId="0" pageNumber="103" type="diagnosis">
<paragraph id="D1898BBE128E8CB44D1D1D79F98A6556" pageId="0" pageNumber="103">Diagnosis.</paragraph>
<paragraph id="DC63E1735BB3E02104E813E4779B99B7" pageId="0" pageNumber="103">
<taxonomicName id="5AC50F719A1EB10AD6AC4D9A5D45EF70" authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leviana cincinnata" order="Araneae" pageId="0" pageNumber="103" phylum="Arthropoda" rank="species" species="cincinnata">
<emphasis id="05AA24B04A10C8FFF3C2D34A50FF1054" italics="true" pageId="0" pageNumber="103">Leviana cincinnata</emphasis>
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sp. nov. differs distinctly from all other species in the genus by its uniform olive-grey abdominal colouration with only a white patch antero-centrally. All other species of
<taxonomicName authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" kingdom="Animalia" lsidName="Leviana" order="Araneae" pageId="0" pageNumber="103" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="103">Leviana</emphasis>
<taxonomicName id="5579514C6424FA021D639C5522F7A6C2" authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leviana" order="Araneae" pageId="0" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="AF6D7B562D2BC25E6B082B2A70403164" italics="true" pageId="0" pageNumber="103">Leviana</emphasis>
</taxonomicName>
gen. nov. have some dark to black abdominal markings. Male genital morphology is most similar to
<taxonomicName genus="L." lsidName="L. dimidiata" pageId="0" pageNumber="103" rank="species" species="dimidiata">
<emphasis italics="true" pageId="0" pageNumber="103">L. dimidiata</emphasis>
<taxonomicName id="C6F19139B51806D236A501538C7F6298" lsidName="L. dimidiata" pageId="0" pageNumber="103" rank="species" species="dimidiata">
<emphasis id="B832C9368D1746566764052C056E3230" italics="true" pageId="0" pageNumber="103">L. dimidiata</emphasis>
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comb. nov. due to the broadly lobed tip of the median apophysis and the strong spine in the basal arch of the median apophysis, but both species distinctly differ in the strong sclerotization of the conductor in
<taxonomicName genus="L." lsidName="L. dimidiata" pageId="0" pageNumber="103" rank="species" species="dimidiata">
<emphasis italics="true" pageId="0" pageNumber="103">L. dimidiata</emphasis>
<taxonomicName id="C07B35AA58E1172CB97AE01CF4E26BA3" lsidName="L. dimidiata" pageId="0" pageNumber="103" rank="species" species="dimidiata">
<emphasis id="B69922F0A11F4C174BD5FFCBB3544697" italics="true" pageId="0" pageNumber="103">L. dimidiata</emphasis>
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comb. nov., that is absent in
<taxonomicName genus="L." lsidName="L. cincinnata" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis italics="true" pageId="0" pageNumber="103">L. cincinnata</emphasis>
<taxonomicName id="E027A5ACA7400C23616432CBDBD30BE3" lsidName="L. cincinnata" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis id="A9211E0D5DF88F5150EBD2677D706249" italics="true" pageId="0" pageNumber="103">L. cincinnata</emphasis>
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sp. nov. (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Leviana dimidiata comb. nov., male (QM S 111921). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character distinguishing L. dimidiata comb. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714524" pageId="0" pageNumber="103">7C</figureCitation>
<figureCitation id="28F8382C9DE5E65FDB9F2E3F96BBDE95" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Leviana dimidiata comb. nov., male (QM S 111921). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character distinguishing L. dimidiata comb. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714524" pageId="0" pageNumber="103">7C</figureCitation>
vs Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10C</figureCitation>
<figureCitation id="7ACF6B9B0A97A3C8CCC63D5220A02CEF" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10C</figureCitation>
). The epigyne scape is much broader than in any other species of
<taxonomicName authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" kingdom="Animalia" lsidName="Leviana" order="Araneae" pageId="0" pageNumber="103" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="103">Leviana</emphasis>
<taxonomicName id="5A28A5748A9AD18A820559322A20021C" authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leviana" order="Araneae" pageId="0" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="832314768E40D50878E02D6EA2402DC7" italics="true" pageId="0" pageNumber="103">Leviana</emphasis>
</taxonomicName>
comb. nov.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Figure 10.</emphasis>
<taxonomicName authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" kingdom="Animalia" lsidName="Leviana cincinnata" order="Araneae" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis italics="true" pageId="0" pageNumber="103">Leviana cincinnata</emphasis>
<caption id="552DED17825B691AEA5EE50E2D1A292E" httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103" start="Figure 10" startId="F10">
<paragraph id="F3F9B7CF11EBEFFB7D171D13EE12C62A" pageId="0" pageNumber="103">
<emphasis id="3CC9FFABA011EB181F33AED5E2B1BC2C" bold="true" pageId="0" pageNumber="103">Figure 10.</emphasis>
<taxonomicName id="F8BCB238BEDE25D7966652A3AE0AF155" authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leviana cincinnata" order="Araneae" pageId="0" pageNumber="103" phylum="Arthropoda" rank="species" species="cincinnata">
<emphasis id="E152E2B8075B670B7513A923A56D8D0B" italics="true" pageId="0" pageNumber="103">Leviana cincinnata</emphasis>
</taxonomicName>
sp. nov., male holotype (QM S26364).
<emphasis bold="true" pageId="0" pageNumber="103">A.</emphasis>
<emphasis id="242E804DD3EF0D9DDAFB88BB9F8206A6" bold="true" pageId="0" pageNumber="103">A.</emphasis>
Habitus, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="103">B.</emphasis>
<emphasis id="77AF450558A80E292BE85489D217B025" bold="true" pageId="0" pageNumber="103">B.</emphasis>
Habitus, ventral view;
<emphasis bold="true" pageId="0" pageNumber="103">C.</emphasis>
<emphasis id="A74C512CC6D9EC7F60357A1CE3202286" bold="true" pageId="0" pageNumber="103">C.</emphasis>
Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (
<emphasis bold="true" pageId="0" pageNumber="103">A, B</emphasis>
<emphasis id="D1672C9756A380AF7ED6145DF3F7117A" bold="true" pageId="0" pageNumber="103">A, B</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="103">C, D</emphasis>
<emphasis id="3F62F0FAFA217B4F3C6264A795D3DB61" bold="true" pageId="0" pageNumber="103">C, D</emphasis>
). Arrow in (
<emphasis bold="true" pageId="0" pageNumber="103">A</emphasis>
<emphasis id="D5CFE046441296E19982AB04F113E2CD" bold="true" pageId="0" pageNumber="103">A</emphasis>
) points to diagnostic character between
<taxonomicName genus="L." lsidName="L. dimidiata" pageId="0" pageNumber="103" rank="species" species="dimidiata">
<emphasis italics="true" pageId="0" pageNumber="103">L. dimidiata</emphasis>
<taxonomicName id="FFD8EC92BE90D33248B61BF17D062506" lsidName="L. dimidiata" pageId="0" pageNumber="103" rank="species" species="dimidiata">
<emphasis id="691BA6DDB1C92D29E37A9AFD9DB99B3E" italics="true" pageId="0" pageNumber="103">L. dimidiata</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName genus="L." lsidName="L. folium" pageId="0" pageNumber="103" rank="species" species="folium">
<emphasis italics="true" pageId="0" pageNumber="103">L. folium</emphasis>
<taxonomicName id="695425536DFCF081BF26920977A56064" lsidName="L. folium" pageId="0" pageNumber="103" rank="species" species="folium">
<emphasis id="BCAC79D67803C626D839607EBBCD0A57" italics="true" pageId="0" pageNumber="103">L. folium</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="description">
<paragraph pageId="0" pageNumber="103">Description.</paragraph>
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Male</emphasis>
<subSubSection id="67C43C94D4D84B5832854B02020BE4AF" pageId="0" pageNumber="103" type="description">
<paragraph id="21AB228C7C5103427FCDDAABBE37535F" pageId="0" pageNumber="103">Description.</paragraph>
<paragraph id="7B2F501AB84C876D305C390E42D200DC" pageId="0" pageNumber="103">
<emphasis id="106F65B7DEF2F61B905148EDFD1B938B" bold="true" pageId="0" pageNumber="103">Male</emphasis>
(based on holotype; QM S26364). Total length 3.4.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Carapace</emphasis>
<emphasis id="86CF55EA209B655E02EA3C010CADBC6C" bold="true" italics="true" pageId="0" pageNumber="103">Carapace</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10A</figureCitation>
<figureCitation id="3D7A482AC58CA8050CFBE0B2A23AC57B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10A</figureCitation>
) 2.1 long, 1.5 wide; uniformly yellow-brown, somewhat lighter in cephalic area; very few white setae mainly in cephalic area; fovea indistinct broad depression.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Eyes</emphasis>
<emphasis id="B1472FF9EF3C4D02B8B769E27A2C27D1" bold="true" italics="true" pageId="0" pageNumber="103">Eyes</emphasis>
: AME 0.11, ALE 0.10, PME 0.11, PLE 0.10; row of eyes: AME 0.35, PME 0.29, PLE 0.79.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Sternum</emphasis>
<emphasis id="CD7D6EA5661D40057C9350977040374C" bold="true" italics="true" pageId="0" pageNumber="103">Sternum</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10B</figureCitation>
<figureCitation id="0452E03AE9C1FF9292C6D3CF5DD3B694" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10B</figureCitation>
) 1.0 long, 0.8 wide; uniformly yellow-brown with indistinct grey pigmentation; covered with few light brown setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Labium</emphasis>
<emphasis id="49B8E1A143B88D45C03D09BA4BBB70E2" bold="true" italics="true" pageId="0" pageNumber="103">Labium</emphasis>
0.35 long, 0.22 wide; light brown.
<taxonomicName genus="Maxillae" lsidName="Maxillae" pageId="0" pageNumber="103" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Maxillae</emphasis>
<taxonomicName id="A6326D4850E0B85758B341DB58E12A9E" genus="Maxillae" lsidName="Maxillae" pageId="0" pageNumber="103" rank="genus">
<emphasis id="9E9CD89DC4A8722CF20A5CC589A9EAF4" bold="true" italics="true" pageId="0" pageNumber="103">Maxillae</emphasis>
</taxonomicName>
light brown. Chelicerae light brown; few light brown setae medially; two retromarginal teeth of equal size, three promarginal teeth with the median largest.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Legs</emphasis>
<emphasis id="7B01567B5A1588ECBCC7D4F491F7E2FE" bold="true" italics="true" pageId="0" pageNumber="103">Legs</emphasis>
: leg formula I&gt; II&gt; IV&gt; III; uniformly yellowish-brown; lengths of segments (femur + patella + tibia + metatarsus + tarsus = total leg length): pedipalp 0.6 + 0.2 + 0.2 + - + 0.7 = 1.7, I 2.4 + 0.8 + 2.7 + 2.3 + 1.0 = 9.2, II 2.1 + 0.7 + 1.8 +1.6 + 0.8 = 7.0, III 1.5 + 0.6 + 0.8 + 0.6 + 0.5 = 4.0, IV 1.6 + 0.8 + 1.9 + 1.5 + 0.6 = 6.4.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Abdomen</emphasis>
<emphasis id="F12DF2D7D4069B0488E27045BA952666" bold="true" italics="true" pageId="0" pageNumber="103">Abdomen</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10A, B</figureCitation>
<figureCitation id="3229D44EE8819FDBEEFE110CA02A7837" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10A, B</figureCitation>
) 2.6 long, 1.9 wide; ovoid; dorsally uniformly dark greenish-grey; white central marking anteriorly and some white speckles laterally and centrally; three pairs of dark brown sigilla centrally with the two anterior pairs largest; few light bristles and white setae; venter greenish grey with two irregular longitudinal lateral white markings, covered with few light brown setae; spinnerets yellow-brown.
</paragraph>
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Pedipalps</emphasis>
<paragraph id="B9C654FD268BFDC854D929725295F753" pageId="0" pageNumber="103">
<emphasis id="E2D5770CFD01630A4B82B9F549764F04" bold="true" italics="true" pageId="0" pageNumber="103">Pedipalps</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10C, D</figureCitation>
<figureCitation id="591CC4BD96DBA7C39AF684299BE79FF7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10C, D</figureCitation>
): median apophysis apically broadly lobed, basally with strong thorn opposing a hooded base; terminal apophysis elongated, fleshy with double-tip; embolus short and sinuous (not visible behind conductor in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10C</figureCitation>
<figureCitation id="7D67CD4DF5C28E9857ECC164E0C8AB5A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Leviana cincinnata sp. nov., male holotype (QM S 26364). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left male pedipalp, ventral view; left male pedipalp, dorsal view. Scale bars: 2 mm (A, B); 0.2 mm (C, D). Arrow in (A) points to diagnostic character between L. dimidiata sp. nov. and L. folium sp. nov." httpUri="https://binary.pensoft.net/fig/714527" pageId="0" pageNumber="103">10C</figureCitation>
); conductor wider than high in ventral view.
</paragraph>
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Female</emphasis>
<paragraph id="A5FA4548EB060C10177D500451C74C75" pageId="0" pageNumber="103">
<emphasis id="2E852DB29F0374735DBCAFF32448615F" bold="true" pageId="0" pageNumber="103">Female</emphasis>
(based on QM S111931; epigyne variation NHMD 12202). Total length 5.1.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Carapace</emphasis>
<emphasis id="78FD1DDE26707B1352EF46661E438B10" bold="true" italics="true" pageId="0" pageNumber="103">Carapace</emphasis>
(Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">11A</figureCitation>
<figureCitation id="223A3F72BBE9F75562BA9BB11C44A4B6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">11A</figureCitation>
) 2.6 long, 2.1 wide; light yellowish-grey; few white setae in particular in cephalic area; fovea shallow and poorly demarcated.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Eyes</emphasis>
<emphasis id="658A3350428846F4ECE7285C2A600023" bold="true" italics="true" pageId="0" pageNumber="103">Eyes</emphasis>
: AME 0.20, ALE 0.14, PME 0.14, PLE 0.13; row of eyes: AME 0.40, PME 0.31, PLE 0.77.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Sternum</emphasis>
<emphasis id="16D547438B044135DF95CAC17304A027" bold="true" italics="true" pageId="0" pageNumber="103">Sternum</emphasis>
(Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">11B</figureCitation>
<figureCitation id="2882E47A117D534393CEC6A4364B4333" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">11B</figureCitation>
) 1.2 long, 1.0 wide, light yellowish-grey; covered with few white setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Labium</emphasis>
<emphasis id="1D8A971D41B1EFF3D477DDC5FB7380EB" bold="true" italics="true" pageId="0" pageNumber="103">Labium</emphasis>
0.31 long, 0.54 wide, colouration as male.
<taxonomicName genus="Maxillae" lsidName="Maxillae" pageId="0" pageNumber="103" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Maxillae</emphasis>
<taxonomicName id="77F1611CAC5A8549F6FA5DB916C9AF96" genus="Maxillae" lsidName="Maxillae" pageId="0" pageNumber="103" rank="genus">
<emphasis id="5C87BE0554F06AD7154CB892F149CFA1" bold="true" italics="true" pageId="0" pageNumber="103">Maxillae</emphasis>
</taxonomicName>
as male. Chelicerae yellowish-brown; few white setae in basal half, black setae mesally mainly in apical half; three promarginal teeth, with the median smallest; two retromarginal teeth of similar size.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Legs</emphasis>
<emphasis id="4B5223BFE0CC94DE727C3F00B16799B9" bold="true" italics="true" pageId="0" pageNumber="103">Legs</emphasis>
: leg formula I&gt; II&gt; IV&gt; III; uniformly yellowish-grey; lengths of segments (femur + patella + tibia + metatarsus + tarsus = total length of leg): pedipalp 0.7 + 0.3 + 0.7 + - + 0.8 = 2.5, I 2.3+ 1.0 + 1.7 + 1.9 + 0.9 = 7.8, II 1.9 + 0.9 + 1.3 + 1.6 + 0.7 = 7.8, III 1.2 + 0.6 + 0.9 + 0.9 + 0.6 = 4.2, IV 1.8 + 0.9 + 1.3 + 1.4 + 0.7 = 6.1.
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Abdomen</emphasis>
<emphasis id="D9123D5052A9A2F58AFAF073A9743113" bold="true" italics="true" pageId="0" pageNumber="103">Abdomen</emphasis>
(Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">11A, B</figureCitation>
<figureCitation id="F407C6CE78663B1DFB3EDC8263912622" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">11A, B</figureCitation>
) 2.7 long, 2.5 wide; ovoid, slightly dorsoventrally compressed; dorsal colour pattern and setae largely as male, but lacking lateral and median lighter patches; spinnerets brown.
</paragraph>
<caption httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Figure 11.</emphasis>
<taxonomicName authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" kingdom="Animalia" lsidName="Leviana cincinnata" order="Araneae" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis italics="true" pageId="0" pageNumber="103">Leviana cincinnata</emphasis>
<caption id="B482E0D027A9E27D71E672B729FDCF65" httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103" start="Figure 11" startId="F11">
<paragraph id="0690B76C2303E04494A83B94992C2847" pageId="0" pageNumber="103">
<emphasis id="C55AF648884777BF81357076E4EDFCD6" bold="true" pageId="0" pageNumber="103">Figure 11.</emphasis>
<taxonomicName id="A107CB7153C7026B169330D003646358" authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leviana cincinnata" order="Araneae" pageId="0" pageNumber="103" phylum="Arthropoda" rank="species" species="cincinnata">
<emphasis id="7DC292EAA5FF81497140C220C2B95039" italics="true" pageId="0" pageNumber="103">Leviana cincinnata</emphasis>
</taxonomicName>
sp. nov., female (
<emphasis bold="true" pageId="0" pageNumber="103">A-E</emphasis>
<emphasis id="AA68F69BD6CED3DDD6EF3E0FD46C3B48" bold="true" pageId="0" pageNumber="103">A-E</emphasis>
QM S111931;
<emphasis bold="true" pageId="0" pageNumber="103">F</emphasis>
<emphasis id="03FDD8CB0935DA5F0F3FE5AB549F640F" bold="true" pageId="0" pageNumber="103">F</emphasis>
NHMD 12202).
<emphasis bold="true" pageId="0" pageNumber="103">A.</emphasis>
<emphasis id="86BB8C228845AC649B138AA002DFC5E2" bold="true" pageId="0" pageNumber="103">A.</emphasis>
Habitus, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="103">B.</emphasis>
<emphasis id="1290DF991FA7B6942EC060AE0561A9B9" bold="true" pageId="0" pageNumber="103">B.</emphasis>
Habitus, ventral view;
<emphasis bold="true" pageId="0" pageNumber="103">C.</emphasis>
<emphasis id="2BD3E21CED3D46CFAD3786B724A7AA5B" bold="true" pageId="0" pageNumber="103">C.</emphasis>
Epigyne, ventral view;
<emphasis bold="true" pageId="0" pageNumber="103">D.</emphasis>
<emphasis id="806F80139AA2DD0E0617664E92F79251" bold="true" pageId="0" pageNumber="103">D.</emphasis>
Epigyne, lateral view;
<emphasis bold="true" pageId="0" pageNumber="103">E.</emphasis>
<emphasis id="E3C3F8614C82349A6FE28813C56A9C5A" bold="true" pageId="0" pageNumber="103">E.</emphasis>
Epigyne, posterior view;
<emphasis bold="true" pageId="0" pageNumber="103">F.</emphasis>
<emphasis id="55DC3132E59349DEF39AC97DDDEA2B96" bold="true" pageId="0" pageNumber="103">F.</emphasis>
Epigyne, ventral view. Scale bars: 2 mm (
<emphasis bold="true" pageId="0" pageNumber="103">A, B</emphasis>
<emphasis id="D5FADF3C7821353DDE542EB1F6B6A910" bold="true" pageId="0" pageNumber="103">A, B</emphasis>
); 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="103">C, D</emphasis>
<emphasis id="1388645766648FA3F490617A5954946F" bold="true" pageId="0" pageNumber="103">C, D</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Epigyne</emphasis>
<paragraph id="8BEA448BBF2639B9324F4CAD4BEEFDE3" pageId="0" pageNumber="103">
<emphasis id="46D612F9D965CAFD7A43295EA99C8F9E" bold="true" italics="true" pageId="0" pageNumber="103">Epigyne</emphasis>
(Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">11C-F</figureCitation>
<figureCitation id="F10F5639BB4D8C7BCD30A3D39E7F713D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Leviana cincinnata sp. nov., female (A-E QM S 111931; F NHMD 12202). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view; F. Epigyne, ventral view. Scale bars: 2 mm (A, B); 0.2 mm (C, D)." httpUri="https://binary.pensoft.net/fig/714528" pageId="0" pageNumber="103">11C-F</figureCitation>
) base wider than long; atrium elevated, bulging; central division broadly oval with narrow base; scape broadly lipped but broken off in most specimens; spermathecae irregularly ovoid, fertilization duct slightly curved and attaching laterally at atrium (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Leviana gen. nov., cleared epigynes, posterior view. A. L. dimidiata comb. nov. (AM KS. 81993); B. L. cincinnata sp. nov. (QM S 111931). Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/714523" pageId="0" pageNumber="103">6B</figureCitation>
<figureCitation id="7ED40F4251233E3EF4337B878F30F3B2" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Leviana gen. nov., cleared epigynes, posterior view. A. L. dimidiata comb. nov. (AM KS. 81993); B. L. cincinnata sp. nov. (QM S 111931). Scale bars: 0.05 mm." httpUri="https://binary.pensoft.net/fig/714523" pageId="0" pageNumber="103">6B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="variation">
<paragraph pageId="0" pageNumber="103">Variation.</paragraph>
<paragraph pageId="0" pageNumber="103">Male total length 3.3-4.1 (n = 4); females 5.4-7.5 (n = 8). There are additional white speckles on the abdomen and white discontinuous lateral bands on the abdomens of some males and females. The epigyne scape was broken in all females measured here, with the exception of two, including the one illustrated. The colouration of live specimens is not known, but it is possible that this species displays greenish or even reddish shades when alive, which often fade into yellow-brown when preserved in ethanol.</paragraph>
<subSubSection id="668B49596E496212F37ED3E9056E9FFC" pageId="0" pageNumber="103" type="variation">
<paragraph id="D6F273C2530CDA0D443B0B019A161AA9" pageId="0" pageNumber="103">Variation.</paragraph>
<paragraph id="8AD0B8160725E9164F995C1611432323" pageId="0" pageNumber="103">Male total length 3.3-4.1 (n = 4); females 5.4-7.5 (n = 8). There are additional white speckles on the abdomen and white discontinuous lateral bands on the abdomens of some males and females. The epigyne scape was broken in all females measured here, with the exception of two, including the one illustrated. The colouration of live specimens is not known, but it is possible that this species displays greenish or even reddish shades when alive, which often fade into yellow-brown when preserved in ethanol.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="life history and habitat preferences">
<paragraph pageId="0" pageNumber="103">Life history and habitat preferences.</paragraph>
<paragraph pageId="0" pageNumber="103">
<subSubSection id="AA8C21D7C3C95F70D88F814F91459FFB" pageId="0" pageNumber="103" type="life history and habitat preferences">
<paragraph id="8EA059E8EF8291B86C0E94545444F658" pageId="0" pageNumber="103">Life history and habitat preferences.</paragraph>
<paragraph id="59C4539C15E0D5C513198398A4256D0C" pageId="0" pageNumber="103">
Mature males of
<taxonomicName genus="L." lsidName="L. cincinnata" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis italics="true" pageId="0" pageNumber="103">L. cincinnata</emphasis>
<taxonomicName id="532F4FA0517CC88B0BD5C7CF17E77247" lsidName="L. cincinnata" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis id="95049B25CA267D09A993A86489831C2D" italics="true" pageId="0" pageNumber="103">L. cincinnata</emphasis>
</taxonomicName>
sp. nov. were almost exclusively found between October and December. Mature females were found between October and January, although some were found in April and one in July.
</paragraph>
<paragraph pageId="0" pageNumber="103">
<paragraph id="87BE1599CF6F89F36517EC810E573B20" pageId="0" pageNumber="103">
Records of
<taxonomicName genus="L." lsidName="L. cincinnata" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis italics="true" pageId="0" pageNumber="103">L. cincinnata</emphasis>
<taxonomicName id="87FC5885452D974934822D568EC4A586" lsidName="L. cincinnata" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis id="9794AE64BB71A6FBA92FD26A9BD5ECC0" italics="true" pageId="0" pageNumber="103">L. cincinnata</emphasis>
</taxonomicName>
sp. nov. are almost entirely from rainforests at altitudes above 500 m. The only information on its leaf-curling behaviour was found with one specimen (QM S26348): &quot;
<emphasis italics="true" pageId="0" pageNumber="103">spider in folded leaf of Liane with thread to tree</emphasis>
<emphasis id="92F8A7B9BC681F870CCFC3A3ED277A87" italics="true" pageId="0" pageNumber="103">spider in folded leaf of Liane with thread to tree</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="103">daytime</emphasis>
<emphasis id="82CAEB40F748FEDADECC0F4F0F3A4537" italics="true" pageId="0" pageNumber="103">daytime</emphasis>
&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="distribution">
<paragraph pageId="0" pageNumber="103">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="103">
<taxonomicName authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" kingdom="Animalia" lsidName="Leviana cincinnata" order="Araneae" pageId="0" pageNumber="103" rank="species" species="cincinnata">
<emphasis italics="true" pageId="0" pageNumber="103">Leviana cincinnata</emphasis>
<subSubSection id="8AD49C8E8571641BDE769260B617097A" pageId="0" pageNumber="103" type="distribution">
<paragraph id="4DC9D63C79D2BDE9A4C7CBEE7DAB5AD5" pageId="0" pageNumber="103">Distribution.</paragraph>
<paragraph id="9BA763646066C4881333CB9B0CABD706" pageId="0" pageNumber="103">
<taxonomicName id="AF9F419C8590526587054674CF14B581" authorityName="Framenau &amp; Kuntner" authorityYear="2022" class="Arachnida" family="Araneidae" genus="Leviana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leviana cincinnata" order="Araneae" pageId="0" pageNumber="103" phylum="Arthropoda" rank="species" species="cincinnata">
<emphasis id="6FD651C775FF1633ED4D0A6EAC8386CC" italics="true" pageId="0" pageNumber="103">Leviana cincinnata</emphasis>
</taxonomicName>
sp. nov. has so far only been reported from northern Queensland between about 16°20'S and 18°03'S Latitude (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution records of Leviana dimidiata comb. nov. and L. cincinnata sp. nov." httpUri="https://binary.pensoft.net/fig/714526" pageId="0" pageNumber="103">9</figureCitation>
<figureCitation id="637DBBFFC297C80BBDE42E68992481E8" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution records of Leviana dimidiata comb. nov. and L. cincinnata sp. nov." httpUri="https://binary.pensoft.net/fig/714526" pageId="0" pageNumber="103">9</figureCitation>
).
</paragraph>
</subSubSection>

View file

@ -1,63 +1,63 @@
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<subSubSection id="7A221490CEB47E7EE0C694ACBAE50390" pageId="0" pageNumber="147" type="nomenclature">
<paragraph id="FBB949DE70746D614A24C1A0C87DF726" pageId="0" pageNumber="147">
<taxonomicName id="B1DC179783E82816D40F48864245F2F7" LSID="4D4CC3F3-BDE6-5305-988F-3DA602F2E28E" authority="(Tsang, Chan, Ip, Ngan, Chen, Lau, Woo) R. Sugita &amp; Kaz. Tanaka" authorityName="R. Sugita &amp; Kaz. Tanaka" authorityYear="2022" baseAuthorityName="Tsang, Chan, Ip, Ngan, Chen, Lau, Woo" baseAuthorityYear="2022" class="Ascomycetes" family="Thyridiaceae" genus="Thyridium" higherTaxonomySource="treatment-meta" kingdom="Fungi" lsidName="Thyridium hongkongense" order="Thyridiales" pageId="0" pageNumber="147" rank="species" species="hongkongense" status="comb. nov.">Thyridium hongkongense (Tsang, Chan, Ip, Ngan, Chen, Lau, Woo) R. Sugita &amp; Kaz. Tanaka</taxonomicName>
<taxonomicNameLabel id="B1501587D0549804576D88E1E79A8F85" pageId="0" pageNumber="147">comb. nov.</taxonomicNameLabel>
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<subSubSection pageId="0" pageNumber="147" type="basionym">
<paragraph pageId="0" pageNumber="147">Basionym.</paragraph>
<paragraph pageId="0" pageNumber="147">
<taxonomicName class="Sordariomycetes" family="Thyridiaceae" genus="Phialemoniopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phialemoniopsis hongkongensis" order="Boliniales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="species" species="hongkongensis">
<emphasis italics="true" pageId="0" pageNumber="147">Phialemoniopsis hongkongensis</emphasis>
<subSubSection id="3488507DA00779FD8D5349B9F5D0D1E0" pageId="0" pageNumber="147" type="basionym">
<paragraph id="729E77F076756DF5A2E1588F0DFAFA99" pageId="0" pageNumber="147">Basionym.</paragraph>
<paragraph id="987881BA2F1F934C9255D69EAEC702FD" pageId="0" pageNumber="147">
<taxonomicName id="92E7E6634F511CECE727383B23B160A4" class="Sordariomycetes" family="Thyridiaceae" genus="Phialemoniopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Phialemoniopsis hongkongensis" order="Boliniales" pageId="0" pageNumber="147" phylum="Ascomycota" rank="species" species="hongkongensis">
<emphasis id="477546EBFC02B6714E4BF9ED26543573" italics="true" pageId="0" pageNumber="147">Phialemoniopsis hongkongensis</emphasis>
</taxonomicName>
Tsang, Chan, Ip, Ngan, Chen, Lau, Woo, J. Clin. Microbiol. 52: 3284 (2014).
</paragraph>

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@ -1,274 +1,274 @@
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<taxonomicNameLabel pageId="0" pageNumber="37">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Live Sphaerobelum pumatense sp. nov. (A-C) and Sphaerobelum bicorne Attems, 1938 (D-H) in the field A-C adult, enrolled, ventral view and walking, respectively D-F adults, ventral view, and enrolled, respectively G, H juvenile, enrolled and unrolling, respectively. Pictures taken not to scale." figureDoi="10.3897/zookeys.930.47742.figure1" httpUri="https://binary.pensoft.net/fig/404782" pageId="0" pageNumber="37">Figs 1A-C</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endoterga of midbody segments, SEM micrographs. A Sphaerobelum bicorne Attems, 1938, ♂ (ZFMK) B Sphaerobelum pumatense sp. nov., ♂ holotype C Sphaeropoeus maculatus (Verhoeff, 1924), ♂ lectotype D Sphaeropoeus bidoupensis sp. nov., ♂ holotype E Sphaeropoeus honbaensis sp. nov., ♂ holotype. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37">, 3B</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">, 6</figureCitation>
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<paragraph id="E09C8B2C2F5B4E7E88314C7A7BD3888A" pageId="0" pageNumber="37">
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<taxonomicNameLabel id="09BAE809F92A88A0272305921D60A663" pageId="0" pageNumber="37">sp. nov.</taxonomicNameLabel>
<figureCitation id="618660C5219604FA9DFAE742015AC30F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Live Sphaerobelum pumatense sp. nov. (A-C) and Sphaerobelum bicorne Attems, 1938 (D-H) in the field A-C adult, enrolled, ventral view and walking, respectively D-F adults, ventral view, and enrolled, respectively G, H juvenile, enrolled and unrolling, respectively. Pictures taken not to scale." figureDoi="10.3897/zookeys.930.47742.figure1" httpUri="https://binary.pensoft.net/fig/404782" pageId="0" pageNumber="37">Figs 1A-C</figureCitation>
<figureCitation id="B295D23F5018C7747FCE0769FAB66A2A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endoterga of midbody segments, SEM micrographs. A Sphaerobelum bicorne Attems, 1938, ♂ (ZFMK) B Sphaerobelum pumatense sp. nov., ♂ holotype C Sphaeropoeus maculatus (Verhoeff, 1924), ♂ lectotype D Sphaeropoeus bidoupensis sp. nov., ♂ holotype E Sphaeropoeus honbaensis sp. nov., ♂ holotype. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37">, 3B</figureCitation>
<figureCitation id="83392D02DE1ED45B040CCA98BB85FEB6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">, 6</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="37">Material examined.</paragraph>
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<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Holotype</emphasis>
<subSubSection id="DD144C230376A067FAF9DB09D30B0A67" pageId="0" pageNumber="37" type="materials_examined">
<paragraph id="BE5477C6B3A64EBD8A6B9E2959472073" pageId="0" pageNumber="37">Material examined.</paragraph>
<paragraph id="409AA533E644330940A57130911C1E0D" pageId="0" pageNumber="37">
<emphasis id="A5197AD1DABAB697E3007EC584A3AC07" bold="true" italics="true" pageId="0" pageNumber="37">Holotype</emphasis>
♂ (ZMUM Rd 4647), Vietnam, Nghe An Prov., Pu Mat National Park,
<geoCoordinate degrees="18" direction="north" minutes="56" orientation="latitude" precision="925" value="18.933332">18°56'N</geoCoordinate>
<geoCoordinate id="AC28488D76521F058B64AF4D2E60E6CB" degrees="18" direction="north" minutes="56" orientation="latitude" precision="925" value="18.933332">18°56'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="38" orientation="longitude" precision="925" value="104.63333">104°38'E</geoCoordinate>
<geoCoordinate id="4FC0AECB21210240760BFFB5438FCB07" degrees="104" direction="east" minutes="38" orientation="longitude" precision="925" value="104.63333">104°38'E</geoCoordinate>
, 200 m a.s.l., mixed tropical forest on steep slopes, on forest floor, daytime, IV.2018, I.I. Semenyuk leg.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Paratypes</emphasis>
<emphasis id="ACC1AB81BDD1E806D1170223E74A7797" bold="true" italics="true" pageId="0" pageNumber="37">Paratypes</emphasis>
1 ♂, 1 ♀ (ZMUM Rd 4632), 1 ♂, 2 ♀ (ZMUM Rd 4648), 1 ♂, 1 ♀ (ZFMK MYR8942), same data as holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="diagnosis">
<paragraph pageId="0" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="37">
<taxonomicName authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum pumatense" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="pumatense">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaerobelum pumatense</emphasis>
<subSubSection id="9D2CA4B1C1223581A23198750A98833E" pageId="0" pageNumber="37" type="diagnosis">
<paragraph id="AE2C0AA421CFB818F7AC8BD5C4F8B6E6" pageId="0" pageNumber="37">Diagnosis.</paragraph>
<paragraph id="260400B681EBFF3B8BABA6E0301C6C70" pageId="0" pageNumber="37">
<taxonomicName id="C9397957006F1FB1F185EDAB9C9127C7" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum pumatense" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="pumatense">
<emphasis id="D9B50817A91D880E4C7F92F17FA18F88" italics="true" pageId="0" pageNumber="37">Sphaerobelum pumatense</emphasis>
</taxonomicName>
sp. nov. belongs to the group of congeners in which the mesal margin of the femur is extended into several teeth (Fig.
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<figureCitation id="70533E7702DF81197D1184ED171B172C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6F</figureCitation>
). In addition,
<taxonomicName lsidName="S. pumatense" pageId="0" pageNumber="37" rank="species" species="pumatense">
<emphasis italics="true" pageId="0" pageNumber="37">S. pumatense</emphasis>
<taxonomicName id="AD8935AD1C0DD563D6F44A276873BA96" lsidName="S. pumatense" pageId="0" pageNumber="37" rank="species" species="pumatense">
<emphasis id="592BCE58F4B68B16D4E549AF65C044F4" italics="true" pageId="0" pageNumber="37">S. pumatense</emphasis>
</taxonomicName>
sp. nov. shares only with
<taxonomicName lsidName="S. spinatum" pageId="0" pageNumber="37" rank="species" species="spinatum">
<emphasis italics="true" pageId="0" pageNumber="37">S. spinatum</emphasis>
<taxonomicName id="E1AD1A53E42985D7CB8375C9CEB462B9" lsidName="S. spinatum" pageId="0" pageNumber="37" rank="species" species="spinatum">
<emphasis id="CB2EC3089A3D958CA1E4CE50E69C34D7" italics="true" pageId="0" pageNumber="37">S. spinatum</emphasis>
</taxonomicName>
Wesener, 2019 and
<taxonomicName lsidName="S. nigrum" pageId="0" pageNumber="37" rank="species" species="nigrum">
<emphasis italics="true" pageId="0" pageNumber="37">S. nigrum</emphasis>
<taxonomicName id="40CE1C9B4FAD44ACA858ADA8AB9CF633" lsidName="S. nigrum" pageId="0" pageNumber="37" rank="species" species="nigrum">
<emphasis id="AE43D6288B7A3E3E0F2BAE81BF802AA8" italics="true" pageId="0" pageNumber="37">S. nigrum</emphasis>
</taxonomicName>
Wesener, 2019, both from Laos, and with
<taxonomicName lsidName="S. cattiense" pageId="0" pageNumber="37" rank="species" species="cattiense">
<emphasis italics="true" pageId="0" pageNumber="37">S. cattiense</emphasis>
<taxonomicName id="FA3EAB6FB7CE846EA27C967710DD580E" lsidName="S. cattiense" pageId="0" pageNumber="37" rank="species" species="cattiense">
<emphasis id="0F3E1BBFCAED00142F99A7CC0595C9D2" italics="true" pageId="0" pageNumber="37">S. cattiense</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4459.3.7" author="Semenyuk, I" journalOrPublisher="Zootaxa" pageId="0" pageNumber="37" pagination="535 - 550" refId="B8" refString="Semenyuk, I, Golovatch, SI, Wesener, T, 2018. Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae). Zootaxa 4459 (3): 535 - 550, DOI: https://doi.org/10.11646/zootaxa.4459.3.7" title="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)." url="https://doi.org/10.11646/zootaxa.4459.3.7" volume="4459" year="2018">Semenyuk et al., 2018</bibRefCitation>
<bibRefCitation id="8EC967F7DDE7E919A665C0D4F8C809FC" DOI="https://doi.org/10.11646/zootaxa.4459.3.7" author="Semenyuk, I" journalOrPublisher="Zootaxa" pageId="0" pageNumber="37" pagination="535 - 550" refId="B8" refString="Semenyuk, I, Golovatch, SI, Wesener, T, 2018. Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae). Zootaxa 4459 (3): 535 - 550, DOI: https://doi.org/10.11646/zootaxa.4459.3.7" title="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)." url="https://doi.org/10.11646/zootaxa.4459.3.7" volume="4459" year="2018">Semenyuk et al., 2018</bibRefCitation>
and
<taxonomicName lsidName="S. konkakinhense" pageId="0" pageNumber="37" rank="species" species="konkakinhense">
<emphasis italics="true" pageId="0" pageNumber="37">S. konkakinhense</emphasis>
<taxonomicName id="BE81E3059518FBB28B2866CFAEB2C66A" lsidName="S. konkakinhense" pageId="0" pageNumber="37" rank="species" species="konkakinhense">
<emphasis id="E80999D872F67F324C3E7368AE92C794" italics="true" pageId="0" pageNumber="37">S. konkakinhense</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4459.3.7" author="Semenyuk, I" journalOrPublisher="Zootaxa" pageId="0" pageNumber="37" pagination="535 - 550" refId="B8" refString="Semenyuk, I, Golovatch, SI, Wesener, T, 2018. Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae). Zootaxa 4459 (3): 535 - 550, DOI: https://doi.org/10.11646/zootaxa.4459.3.7" title="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)." url="https://doi.org/10.11646/zootaxa.4459.3.7" volume="4459" year="2018">Semenyuk et al., 2018</bibRefCitation>
<bibRefCitation id="EC412017A8D2AE3C449CDEE2966EE8A4" DOI="https://doi.org/10.11646/zootaxa.4459.3.7" author="Semenyuk, I" journalOrPublisher="Zootaxa" pageId="0" pageNumber="37" pagination="535 - 550" refId="B8" refString="Semenyuk, I, Golovatch, SI, Wesener, T, 2018. Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae). Zootaxa 4459 (3): 535 - 550, DOI: https://doi.org/10.11646/zootaxa.4459.3.7" title="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)." url="https://doi.org/10.11646/zootaxa.4459.3.7" volume="4459" year="2018">Semenyuk et al., 2018</bibRefCitation>
, both from Vietnam, a straight telopoditomere 4 of the posterior telopod (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6A, B</figureCitation>
<figureCitation id="CD91DCCCB43AE5940543D72D62B3BB2F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6A, B</figureCitation>
).
<taxonomicName authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum pumatense" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="pumatense">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaerobelum pumatense</emphasis>
<taxonomicName id="727AD9A6F9E64012BD7C33F1A6BE644E" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum pumatense" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="pumatense">
<emphasis id="1116B6FBF0C194F580D828F95EB63866" italics="true" pageId="0" pageNumber="37">Sphaerobelum pumatense</emphasis>
</taxonomicName>
sp. nov. differs in several unique characters from the four other species: vulval operculum not projecting mesally into a strong tip (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6H</figureCitation>
<figureCitation id="17CD9751EB83BA610F56106BE2F08857" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6H</figureCitation>
); telopoditomere 4 of posterior telopods exceptionally slender,&gt;5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="04DA810B70A66D9166266298D8F95364" originalValue="×">x</normalizedToken>
as long as wide (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6A, B</figureCitation>
<figureCitation id="FF364FBAFB26528F692381006AAE4D49" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6A, B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Figure 3.</emphasis>
<caption id="9FC427B2FC759DDEA209E61E57022010" doi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37" start="Figure 3" startId="F3">
<paragraph id="16C568D6C927E325E7BE96780D97F266" pageId="0" pageNumber="37">
<emphasis id="3F537F341799ED2F7BDB7519D6126AE7" bold="true" pageId="0" pageNumber="37">Figure 3.</emphasis>
Endoterga of midbody segments, SEM micrographs.
<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
<taxonomicName authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum bicorne" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaerobelum bicorne</emphasis>
<emphasis id="C6B7765E2D0E6F22792533B900FB57FD" bold="true" pageId="0" pageNumber="37">A</emphasis>
<taxonomicName id="F9D9E623614C75E6FAD8500A80E94963" authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum bicorne" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="6E86C5E1BBF9E27D1843CFFD93105A3F" italics="true" pageId="0" pageNumber="37">Sphaerobelum bicorne</emphasis>
</taxonomicName>
Attems, 1938, ♂ (ZFMK)
<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
<taxonomicName authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum pumatense" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="pumatense">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaerobelum pumatense</emphasis>
<emphasis id="F923850B79584362F0ACA74FDC697D84" bold="true" pageId="0" pageNumber="37">B</emphasis>
<taxonomicName id="C17C614570340D6F97DCE4BA950865DA" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum pumatense" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="pumatense">
<emphasis id="6722F334D89B177CD8DCAD5186518A76" italics="true" pageId="0" pageNumber="37">Sphaerobelum pumatense</emphasis>
</taxonomicName>
sp. nov., ♂ holotype
<emphasis bold="true" pageId="0" pageNumber="37">C</emphasis>
<taxonomicName baseAuthorityName="Verhoeff" baseAuthorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus maculatus" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaeropoeus maculatus</emphasis>
<emphasis id="AC3C57B2DCFC5FA3E3DA46D1C61519B8" bold="true" pageId="0" pageNumber="37">C</emphasis>
<taxonomicName id="14F610545CA44EA1208BD626136638E2" baseAuthorityName="Verhoeff" baseAuthorityYear="1924" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus maculatus" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="96D5A503A56D78ACF6D4DC153A6D15DA" italics="true" pageId="0" pageNumber="37">Sphaeropoeus maculatus</emphasis>
</taxonomicName>
(Verhoeff, 1924), ♂ lectotype
<emphasis bold="true" pageId="0" pageNumber="37">D</emphasis>
<taxonomicName authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus bidoupensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bidoupensis">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaeropoeus bidoupensis</emphasis>
<emphasis id="C8E921A23186D8373D8FF6F1B4425449" bold="true" pageId="0" pageNumber="37">D</emphasis>
<taxonomicName id="0497AE77D233F0CEF623864BC91FDFDF" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus bidoupensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bidoupensis">
<emphasis id="A420AB7E4B334A9C73761F701A71FD6D" italics="true" pageId="0" pageNumber="37">Sphaeropoeus bidoupensis</emphasis>
</taxonomicName>
sp. nov., ♂ holotype
<emphasis bold="true" pageId="0" pageNumber="37">E</emphasis>
<taxonomicName authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="honbaensis">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaeropoeus honbaensis</emphasis>
<emphasis id="6AD661EE256513F58A334CA5E64043CD" bold="true" pageId="0" pageNumber="37">E</emphasis>
<taxonomicName id="712FC238DB6778E5686973B2B84D76BE" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus honbaensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="honbaensis">
<emphasis id="55E3B2E381F967E01B5C95CCF608A37D" italics="true" pageId="0" pageNumber="37">Sphaeropoeus honbaensis</emphasis>
</taxonomicName>
sp. nov., ♂ holotype. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="description">
<paragraph pageId="0" pageNumber="37">Description.</paragraph>
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Body length</emphasis>
<subSubSection id="DAD9E53D0964A5DA814ED0E672AC2384" pageId="0" pageNumber="37" type="description">
<paragraph id="AABACC830327FA9170D090170567C53D" pageId="0" pageNumber="37">Description.</paragraph>
<paragraph id="17D4A1F576CF48C7A4DA78773DB27873" pageId="0" pageNumber="37">
<emphasis id="2CB53B2F2BF0E2785ADBFBF044880148" bold="true" italics="true" pageId="0" pageNumber="37">Body length</emphasis>
: holotype ♂ length ca. 27.2 mm, width of thoracic shield 15.61 mm, of tergite 7 (= broadest), 15.5 mm, height of thoracic shield, 7.9 mm, of tergite 7 (= highest), 8.2 mm; ♀ length ca. 34.8 mm, width of thoracic shield, 17.2 mm, of tergite 7, 17.9 mm (= broadest), height of thoracic shield, 9.5 mm, of tergite 7, 9.9 mm (= highest). Width of paratypes 14-17 (♂) or 15-17 mm (♀).
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Coloration</emphasis>
<emphasis id="ECE0739EA25C03FB4BF25718FF84456C" bold="true" italics="true" pageId="0" pageNumber="37">Coloration</emphasis>
: both in vivo and in vitro, after&gt;1.5 years of preservation in ethanol, similar, uniformly dark blackish brown, dark chestnut, dark grey-brown or dark reddish brown, in alcohol with posterior margin usually dark brown, paratergite depressions, groove of thoracic shield and legs dark olive to olive-brown, but several basal podomeres and distal halves of tarsi considerably lighter, orange. Antennae blackish to dark brown, but antennomeres 5 and 6 lighter, light brown to nearly orange. Tegument mosly dull to poorly shining (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Live Sphaerobelum pumatense sp. nov. (A-C) and Sphaerobelum bicorne Attems, 1938 (D-H) in the field A-C adult, enrolled, ventral view and walking, respectively D-F adults, ventral view, and enrolled, respectively G, H juvenile, enrolled and unrolling, respectively. Pictures taken not to scale." figureDoi="10.3897/zookeys.930.47742.figure1" httpUri="https://binary.pensoft.net/fig/404782" pageId="0" pageNumber="37">1A-C</figureCitation>
<figureCitation id="2C9675DDE462C9A4604D72E1D087C0B2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Live Sphaerobelum pumatense sp. nov. (A-C) and Sphaerobelum bicorne Attems, 1938 (D-H) in the field A-C adult, enrolled, ventral view and walking, respectively D-F adults, ventral view, and enrolled, respectively G, H juvenile, enrolled and unrolling, respectively. Pictures taken not to scale." figureDoi="10.3897/zookeys.930.47742.figure1" httpUri="https://binary.pensoft.net/fig/404782" pageId="0" pageNumber="37">1A-C</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Head</emphasis>
<emphasis id="38B7835DB0459EDFB61F150B17B5482E" bold="true" italics="true" pageId="0" pageNumber="37">Head</emphasis>
: eyes with&gt;75 ommatidia. Aberrant ocellus located inside antennal groove. Antennae short (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6G</figureCitation>
<figureCitation id="4FC6A2AEE5FFA97561DDD1A3CFA5465C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6G</figureCitation>
), with rounded joints, protruding posteriorly to leg-pair 3. Lengths of antennomeres: 1 = 2 = 3 = 4 = 5 &lt;&lt; 6. All antennomeres densely pubescent, sensilla basiconica surrounding apical disc. Shape of antennae sexually dimorphic, cylindrical in ♀, thickened, apically broadened and flattened in ♂. Apical disc with ca. 63/67 (♂) or 48/45 (♀) apical cones, respectively. Organ of
<normalizedToken originalValue="Tömösváry">Toemoesvary</normalizedToken>
<normalizedToken id="A7524421D8437CD7F0380108412654A8" originalValue="Tömösváry">Toemoesvary</normalizedToken>
located inside antennal groove.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Gnathochilarium</emphasis>
<emphasis id="9A79BE423F4E260A39DF7166793A5FA6" bold="true" italics="true" pageId="0" pageNumber="37">Gnathochilarium</emphasis>
: palpi with sensory cones arranged in a single field.
<taxonomicName genus="Mandibles" lsidName="Mandibles" pageId="0" pageNumber="37" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Mandibles</emphasis>
<taxonomicName id="E392EDB719EC1C3081940D5FA57F692B" genus="Mandibles" lsidName="Mandibles" pageId="0" pageNumber="37" rank="genus">
<emphasis id="D03E37E6725BF17F67ABD9B5074E35B9" bold="true" italics="true" pageId="0" pageNumber="37">Mandibles</emphasis>
</taxonomicName>
: not dissected.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Stigmatic plates</emphasis>
<emphasis id="A923AF40220A6F41FD0ED96B29F26BAA" bold="true" italics="true" pageId="0" pageNumber="37">Stigmatic plates</emphasis>
: first stigmatic plate rounded, apex well-rounded, straight towards coxa 1.
<taxonomicName genus="Laterotergites" lsidName="Laterotergites" pageId="0" pageNumber="37" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Laterotergites</emphasis>
<taxonomicName id="140223731A9C1CE9450B3EE60D6BCD01" genus="Laterotergites" lsidName="Laterotergites" pageId="0" pageNumber="37" rank="genus">
<emphasis id="8B0EF87D5E9A5AF927C426E40826311A" bold="true" italics="true" pageId="0" pageNumber="37">Laterotergites</emphasis>
</taxonomicName>
: laterotergite 1 strongly elongated into a well-ronded tip. Laterotergite 2 with a broad, stout, much shorter projection.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Collum</emphasis>
<emphasis id="D00D68962FAB030B374276C665DE882A" bold="true" italics="true" pageId="0" pageNumber="37">Collum</emphasis>
: with few setae on surface, anterior and posterior margins with 3-4 rows of short setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Thoracic shield</emphasis>
<emphasis id="34080B053EC1F0D6760C714D12E5E378" bold="true" italics="true" pageId="0" pageNumber="37">Thoracic shield</emphasis>
: surface like tergites, longer setae only in grooves. Slope towards groove without anterior, but with 3 or 4 posterior keels.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Tergites</emphasis>
<emphasis id="7CA84D9DE33E8B3E527962043C303CF6" bold="true" italics="true" pageId="0" pageNumber="37">Tergites</emphasis>
: surface densely setose with short setae standing in pits. Tips of paratergites of midbody tergites slightly projecting posteriorly.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Endotergum</emphasis>
<emphasis id="B63A0DDFF938B0AFCA08FF96043F0C10" bold="true" italics="true" pageId="0" pageNumber="37">Endotergum</emphasis>
: inner section lacking any spines or setae. Middle area with a single row of large, sparse, elliptical, cuticular impressions. Distance between impressions&gt;2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="CB0ABF6747CFEA6C104358005C1FF575" originalValue="×">x</normalizedToken>
their diameter. Apically, two sparse rows of marginal bristles, tips of longest setae slightly protruding beyond tergal margin (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endoterga of midbody segments, SEM micrographs. A Sphaerobelum bicorne Attems, 1938, ♂ (ZFMK) B Sphaerobelum pumatense sp. nov., ♂ holotype C Sphaeropoeus maculatus (Verhoeff, 1924), ♂ lectotype D Sphaeropoeus bidoupensis sp. nov., ♂ holotype E Sphaeropoeus honbaensis sp. nov., ♂ holotype. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37">3B</figureCitation>
<figureCitation id="5DF421A4500A21FD69941248C96AEA09" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Endoterga of midbody segments, SEM micrographs. A Sphaerobelum bicorne Attems, 1938, ♂ (ZFMK) B Sphaerobelum pumatense sp. nov., ♂ holotype C Sphaeropoeus maculatus (Verhoeff, 1924), ♂ lectotype D Sphaeropoeus bidoupensis sp. nov., ♂ holotype E Sphaeropoeus honbaensis sp. nov., ♂ holotype. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.930.47742.figure3" httpUri="https://binary.pensoft.net/fig/404784" pageId="0" pageNumber="37">3B</figureCitation>
). Bristles not smooth, but barbed, with numerous small spinicles.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Anal shield</emphasis>
<emphasis id="84E21B3324C92B8BD6D1739A2CD8EC6A" bold="true" italics="true" pageId="0" pageNumber="37">Anal shield</emphasis>
: large, in both sexes regularly rounded. Surface in both sexes completely covered with tiny setae located in small pits. Underside with a single, long, black, locking carina 2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="3F7C824F1F0A0E87484A5D1B67F2D058" originalValue="×">x</normalizedToken>
as long as width of last laterotergite. Carina located close to last laterotergite.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Legs</emphasis>
<emphasis id="4502D3AE800A52D818C5B3ADA3D6BEAA" bold="true" italics="true" pageId="0" pageNumber="37">Legs</emphasis>
: leg 1 with 6, leg 2 with 7, leg 3 with 7 or 8 ventral spines. First two leg-pairs without an apical spine. Leg-pairs 4-21 with 10-12 ventral spines and one dorso-apical spine. In leg 9, femur 1.6, tarsus 4.5 times longer than wide (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6F</figureCitation>
<figureCitation id="D3201B4DDD07B2D9E2C23013914AA169" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6F</figureCitation>
). All podomeres densely setose. Coxa with a large and process dentate at margins. Coxa process absent from first and sharply projecting in second leg. Prefemur at apical margin with a projection mesally. Femur extended mesally into a dentate margin featuring 12-14 teeth.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Female sexual characters</emphasis>
<emphasis id="9A8B8D3C7568B382DD50397F71119060" bold="true" italics="true" pageId="0" pageNumber="37">Female sexual characters</emphasis>
: vulva large, covering half of coxa, extending mesally to anterior third of prefemur (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6H</figureCitation>
<figureCitation id="70DDA35537237E57A491B5B09C5D8223" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6H</figureCitation>
). Operculum rounded, very slightly invaginated medially, mesal margin slightly projecting into a well-rounded lobe slightly higher than remaining operculum.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Subanal plate</emphasis>
<emphasis id="DB7F488BFC03AD3DB72EE0652D1FFC0D" bold="true" italics="true" pageId="0" pageNumber="37">Subanal plate</emphasis>
: large and wide, centrally recessed.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Male sexual characters</emphasis>
<emphasis id="9C2AEF8526D65322699B6759B18F9F43" bold="true" italics="true" pageId="0" pageNumber="37">Male sexual characters</emphasis>
: gonopore covered with a single, undivided, circular, sclerotized plate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Anterior telopods</emphasis>
<emphasis id="44477EF83D9DA149B2D38BCA55147C2F" bold="true" italics="true" pageId="0" pageNumber="37">Anterior telopods</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6C-E</figureCitation>
<figureCitation id="560C116259CD9191FA1265FE6D691301" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6C-E</figureCitation>
): consisting of 4 telopoditomeres distal to syncoxite. Telopoditomere 1 rectangular, as long as wide. Telopoditomere 2 large, without process, as long as telopoditomere 1. Process of telopoditomere 2 located posteriorly, visible mesally in anterior view. Process wide, well-rounded, projecting to basal part of telopoditomere 3. Telopoditomeres 3 and 4 slightly curved mesally. Telopoditomere 3 small, cylindrical, slightly shorter than telopoditomere 4, with a spine juxtaposed to process of telopoditomere 2. Telopoditomere 4 cylindrical, well-rounded, posterior side with 2-4 small spines. All telopoditomeres covered with long setae.
<emphasis bold="true" italics="true" pageId="0" pageNumber="37">Posterior telopods</emphasis>
<emphasis id="144152A6FD021C368C4AA8ACEDA79C0D" bold="true" italics="true" pageId="0" pageNumber="37">Posterior telopods</emphasis>
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6A, B</figureCitation>
<figureCitation id="17289F2A2C05CDE15F4061D101E824BB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Sphaerobelum pumatense sp. nov., ♂ holotype (A-G) and ♀ paratype (H). A, B Left posterior telopod, anterior and posterior views, respectively C-E left anterior telopod, anterior, lateral and posterior views, respectively F left leg 9, anterior view G left antenna H left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm, D drawn not to scale." figureDoi="10.3897/zookeys.930.47742.figure6" httpUri="https://binary.pensoft.net/fig/404787" pageId="0" pageNumber="37">6A, B</figureCitation>
): telopoditomere 1 narrow, as long as wide. Immovable finger (process of telopoditomere 2) slightly shorter than movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a characteristic, distally swollen apex, clearly rounded apically, apex only slightly wider than base. Swelling only slightly expressed basally. Telopoditomere 3 rectangular, well-rounded, apically with a clearly rounded extension carrying three small spines. Telopoditomere 4 only slightly shorter and much more slender than telopoditomere 3, 5.2
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="577FB0854FD068A1FC0BD2C6593ED2B2" originalValue="×">x</normalizedToken>
as long as wide, apically weakly tapering, straight, not curved, with 23 small spines at margin towards immovable finger close to tip. In anterior view, telopoditomeres 1-3 covered with setae, in posterior view telopoditomeres 2-4 more glabrous except for numerous setae at margins.
</paragraph>
<caption doi="10.3897/zookeys.930.47742.figure4" httpUri="https://binary.pensoft.net/fig/404785" pageId="0" pageNumber="37" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Figure 4.</emphasis>
<caption id="67F1B0301158F70E1D512D89F9674F41" doi="10.3897/zookeys.930.47742.figure4" httpUri="https://binary.pensoft.net/fig/404785" pageId="0" pageNumber="37" start="Figure 4" startId="F4">
<paragraph id="A7900FB0B3355184A62A9DF74AC2B3CB" pageId="0" pageNumber="37">
<emphasis id="367C736E06FB2DF129B17EC7EFFD76CF" bold="true" pageId="0" pageNumber="37">Figure 4.</emphasis>
Tergal surface of midbody segments, SEM micrographs.
<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
<taxonomicName authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum bicorne" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaerobelum bicorne</emphasis>
<emphasis id="357AFEF7AE877DBAE69C235B95C6759F" bold="true" pageId="0" pageNumber="37">A</emphasis>
<taxonomicName id="6BC23A5D1FE1DBC1911BE4027D2C4CE1" authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum bicorne" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="E473FEBF7A7F9A14704DD1563E24E46D" italics="true" pageId="0" pageNumber="37">Sphaerobelum bicorne</emphasis>
</taxonomicName>
Attems, 1938, ♂ (ZFMK)
<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
<taxonomicName authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus bidoupensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bidoupensis">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaeropoeus bidoupensis</emphasis>
<emphasis id="E6E3E759F9E81204FDF179D800AA796D" bold="true" pageId="0" pageNumber="37">B</emphasis>
<taxonomicName id="61E7AEF9F75EBD0B135EDC533E6E4D03" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaeropoeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaeropoeus bidoupensis" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bidoupensis">
<emphasis id="95DAC0467E8BA98297B1A7BDAF309021" italics="true" pageId="0" pageNumber="37">Sphaeropoeus bidoupensis</emphasis>
</taxonomicName>
sp. nov., ♂ holotype. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="etymology">
<paragraph pageId="0" pageNumber="37">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="37">To emphasize the provenance from the Pu Mat National Park, adjective.</paragraph>
<subSubSection id="E3671649CE6E966CE3E990D96A032475" pageId="0" pageNumber="37" type="etymology">
<paragraph id="0F6A3AFB4ED880E16C7D435B3A60F0BF" pageId="0" pageNumber="37">Etymology.</paragraph>
<paragraph id="61F566A0702F7411EB3FD2BE4396BF86" pageId="0" pageNumber="37">To emphasize the provenance from the Pu Mat National Park, adjective.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="37" type="remarks">
<paragraph pageId="0" pageNumber="37">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="37">
<subSubSection id="88D11969847A3B6D9913BFE5B58EF108" pageId="0" pageNumber="37" type="remarks">
<paragraph id="C1A517E1A19F75EF6288F96B1D7F2898" pageId="0" pageNumber="37">Remarks.</paragraph>
<paragraph id="E9C04A75445FCCE4A190EA1DC96A9A8B" pageId="0" pageNumber="37">
In the field, the weather was fluctuating between very dry and hot days and several rainy days, with day temperature above the leaf litter averaging 24 °C, and night temperature averaging 21 °C, not dropping below 18.5 °C.
<taxonomicName authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum pumatense" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="pumatense">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaerobelum pumatense</emphasis>
<taxonomicName id="81ABF44C603F894360A08D419F393375" authorityName="Semenyuk &amp; Golovatch &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum pumatense" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="pumatense">
<emphasis id="FB3316379BC59D38C6710DC8DD2055C8" italics="true" pageId="0" pageNumber="37">Sphaerobelum pumatense</emphasis>
</taxonomicName>
were quite rare in any weather conditions. The millipedes were found in forests ranging from 150 m to 400 m a.s.l., the forest canopy appeared to have been free of this species. Females were mostly hidden in leaf litter in small patches of litter on very steep slopes (60-70°), forming &quot;living rooms&quot;, choosing places with water oozing from beneath shale so that the leaf litter was constantly wet. Males were found mainly on the forest floor at night. No juveniles were seen.
</paragraph>
<caption doi="10.3897/zookeys.930.47742.figure5" httpUri="https://binary.pensoft.net/fig/404786" pageId="0" pageNumber="37" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="37">
<emphasis bold="true" pageId="0" pageNumber="37">Figure 5.</emphasis>
<taxonomicName authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum bicorne" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis italics="true" pageId="0" pageNumber="37">Sphaerobelum bicorne</emphasis>
<caption id="3348FBCA337EEC8BA4948ED923044876" doi="10.3897/zookeys.930.47742.figure5" httpUri="https://binary.pensoft.net/fig/404786" pageId="0" pageNumber="37" start="Figure 5" startId="F5">
<paragraph id="CC2D8272B535463E10308657664A97C4" pageId="0" pageNumber="37">
<emphasis id="AA6AED75691FFE9325857EDCE4F208F3" bold="true" pageId="0" pageNumber="37">Figure 5.</emphasis>
<taxonomicName id="B5B4878265146B40A85DD727C1D7720D" authorityName="Attems" authorityYear="1938" class="Diplopoda" family="Zephroniidae" genus="Sphaerobelum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphaerobelum bicorne" order="Sphaerotheriida" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="bicorne">
<emphasis id="B48ED48E4D2A5B1C07EC67785CD45470" italics="true" pageId="0" pageNumber="37">Sphaerobelum bicorne</emphasis>
</taxonomicName>
Attems, 1938, ♂ (
<emphasis bold="true" pageId="0" pageNumber="37">A-G</emphasis>
<emphasis id="92292ED03267DE49A53B70026A1B5A88" bold="true" pageId="0" pageNumber="37">A-G</emphasis>
) and ♀ (
<emphasis bold="true" pageId="0" pageNumber="37">H</emphasis>
<emphasis id="7311E093EF34BC82F29977701704B581" bold="true" pageId="0" pageNumber="37">H</emphasis>
) from Song Thanh National Park.
<emphasis bold="true" pageId="0" pageNumber="37">A, B</emphasis>
<emphasis id="0BB09AA820CF916F9E9A5685A02AC3C6" bold="true" pageId="0" pageNumber="37">A, B</emphasis>
Right posterior telopod, anterior and posterior views, respectively
<emphasis bold="true" pageId="0" pageNumber="37">C, D</emphasis>
<emphasis id="2849BCA245A8E013837E7AD850FC8F5F" bold="true" pageId="0" pageNumber="37">C, D</emphasis>
right anterior telopod, posterior and anterior views, respectively
<emphasis bold="true" pageId="0" pageNumber="37">E</emphasis>
<emphasis id="0464A5038CC689E6584DD49CDDA98241" bold="true" pageId="0" pageNumber="37">E</emphasis>
telopoditomeres 2 and 3 of right anterior telopod, lateral view
<emphasis bold="true" pageId="0" pageNumber="37">F</emphasis>
<emphasis id="6B7F282434F24756FDF126E666F724D6" bold="true" pageId="0" pageNumber="37">F</emphasis>
left leg 9, anterior view
<emphasis bold="true" pageId="0" pageNumber="37">G</emphasis>
<emphasis id="4BA50566CEAFE15C285E78C75F284551" bold="true" pageId="0" pageNumber="37">G</emphasis>
left antenna
<emphasis bold="true" pageId="0" pageNumber="37">H</emphasis>
<emphasis id="76B3053160A72957EA96218E76B5ED83" bold="true" pageId="0" pageNumber="37">H</emphasis>
left coxa and prefemur 2 with vulva, posterior view. Scale bars: 1.0 mm,
<emphasis bold="true" pageId="0" pageNumber="37">E</emphasis>
<emphasis id="654963B20E41678DE6B737A41BB2BC56" bold="true" pageId="0" pageNumber="37">E</emphasis>
drawn not to scale.
</paragraph>
</caption>

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@ -1,115 +1,115 @@
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<mods:title>Earthworm species occurrence in agroecosystems in the Midlands, KwaZulu-Natal, South Africa</mods:title>
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Pontoscolex corenthrurus (
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<paragraph pageId="0" pageNumber="411">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="411">Brazil</paragraph>
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<paragraph id="29B2DDB473D811CCD3F5729D06B1C8AD" pageId="0" pageNumber="411">Type locality.</paragraph>
<paragraph id="88E53F3387ED576721E8C0B6DDA5B074" pageId="0" pageNumber="411">Brazil</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="411">(Old material). KZN</emphasis>
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<paragraph id="91FDBA5868C0122B07447063CDDDEA8A" pageId="0" pageNumber="411">RSA distribution</paragraph>
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<emphasis id="DAFD3FFBCA2ABF1686DED4CC2F68C69A" bold="true" pageId="0" pageNumber="411">(Old material). KZN</emphasis>
: Port Shepstone area, Oribi Gorge Nature Reserve, Skyline Nature Reserve, Bluff Nature Reserve, Dlinza Forest Nature Reserve, Greater St Lucia Wetland Park. Kosi Bay, Krantzkloof Nature Reserve, Langepa Natural Heritage Site, Mbumbazi Nature Reserve, Ngoye Forest Reserve, Umfolozi Game Reserve, Mapelane Nature Reserve, Melmoth, Empangeni, Richards Bay, Empangeni, Stanger, Entumeni Nature Reserve, Eshowe, Fairfield Farm, Brooklee Farm, KwaMahleka Farm, Benhurst Farm, Harebottle, Mistyridge Farm, Rockyridge Farm, Cathedral Peak, Monks Cowl Nature Reserve, Umlalazi Nature Reserve, Gingindlovu,
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Bluff The
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Farm, Pietermaritzburg area, Durban area, Tongaat, Inanda, Mt Edgecombe, Ichanga, Westville, Stamford Hill, North Park Nature Reserve, Kenneth Stainbank Nature Reserve, Ixopo area, Vernon Crookes Nature Reserve, Amanzimtoti Mission Reserve, Adams Mission, Umzinto area, Hibberdene, Mpenjati Nature Reserve, Kasseepursad, Tugela area and Umgababa.
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<emphasis id="0BAD370B4802379AC5E11873A537162B" bold="true" pageId="0" pageNumber="411">MP</emphasis>
: Kruger National Park, Thalalanati, Nelspruit, White River and Graskop.
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<emphasis id="F99CFDF71BEBE038B5DF8017301D790F" bold="true" pageId="0" pageNumber="411">LP</emphasis>
: New Agatha, Soutpansberg Ratombo, Entabeni State Forest, Letsitele and Tzaneen area.
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: Pretoria. New material:
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<emphasis id="D330AA6C636B3BBDBB8A26985FA5D143" bold="true" pageId="0" pageNumber="411">KZN</emphasis>
: Dalton.
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<paragraph pageId="0" pageNumber="411">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="411">
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<paragraph id="B6C0514AB10CA21DAF08352D2996129E" pageId="0" pageNumber="411">Remarks.</paragraph>
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This species occurs in numerous sites, both natural and agricultural. In natural sites,
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reported this species in grasslands, forests, natural bushes and near rivers. In cultivated fields,
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has been collected under most crops including maize, sugar-cane, banana, avocado, citrus; some specimens were collected in pine and gum plantations, some even from vegetable gardens (
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). This species has been used in experiments at different institutions.
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<emphasis id="6A962247FA2A989DFFCBD4869541C7C4" italics="true" pageId="0" pageNumber="411">Pontoscolex corenthrurus</emphasis>
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has also been collected in polluted sites in KZN (
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). The current collection was in sugarcane on one farm only, although we had expected to collect the species in other areas in the KZN Midlands because of its wide occurrence.
</paragraph>
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<figureCitation captionStart="Fig. 8" captionStartId="F8" captionText="Fig. 8. Cestrophorus spp. male genitalia compared. A. White pronotal maculae of C. paradoxus; B. Parameres of C. paradoxus ' upside-down dancing anteaters'; C. Dorsal aspect C. ditachus parameres; D. Lateral view of C. ditachus paramere; E. Photo of terminalia of C. amplitenuis; F. Lateral aspect of C. amplitenuis paramere." figureDoi="10.3897/jor.31.82306.fig8" httpUri="https://binary.pensoft.net/fig/750937" pageId="0" pageNumber="143">, 8A, B</figureCitation>
<figureCitation captionStart="Fig. 11" captionStartId="F11" captionText="Fig. 11. Song of C. paradoxus is an ongoing sequence of lisp pairs. A. Sequence at low time resolution showing two complete pairs; B. A lisp at higher resolution reveals its composition as a long train of&gt; 25 discrete rapid-decay pulses; C. Three pulses showing stereotyped waveform detail; D. Power spectrum of C. paradoxus with frequencies confined to the audio range in a narrow band 8 - 13 kHz." figureDoi="10.3897/jor.31.82306.fig11" httpUri="https://binary.pensoft.net/fig/750940" pageId="0" pageNumber="143">, 11</figureCitation>
<figureCitation captionStart="Fig. 12" captionStartId="F12" captionText="Fig. 12. Waterfall record of one C. paradoxus 60 - ms pulse train / lisp indicating absence of ultrasonics." figureDoi="10.3897/jor.31.82306.fig12" httpUri="https://binary.pensoft.net/fig/750941" pageId="0" pageNumber="143">, 12</figureCitation>
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<specimenCount id="B4FCD617FA6370931ED9F37DCD2C7079" type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:4D743D417AA95F5AAD83FC6F286225DC:11DDBFD903AB9F1BCE7AEBD10B49346D" country="Ecuador" name="Cosanga">Cosanga</location>
<location id="11DDBFD903AB9F1BCE7AEBD10B49346D" LSID="urn:lsid:plazi:treatment:4D743D417AA95F5AAD83FC6F286225DC:11DDBFD903AB9F1BCE7AEBD10B49346D" country="Ecuador" name="Cosanga">Cosanga</location>
;
<collectingDate value="1988-02-10">10 Feb. 1988</collectingDate>
<collectingDate id="D1D56C5C5B9409454ED520D77AA3DA58" value="1988-02-10">10 Feb. 1988</collectingDate>
;
<collectorName>G.K. Morris</collectorName>
<collectorName id="AD2C52F8ED801CA35CCF8ED8C3ACF257">G.K. Morris</collectorName>
leg.; MLP
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="redescription">
<paragraph pageId="0" pageNumber="143">
<emphasis italics="true" pageId="0" pageNumber="143">Redescription.</emphasis>
<subSubSection id="FDDCF1FD9DB4F65FD619FBF798C6794B" pageId="0" pageNumber="143" type="redescription">
<paragraph id="700CC69A923DB1346D1CA057ED037EF1" pageId="0" pageNumber="143">
<emphasis id="C88CF1FC311A9D6A6EE81548FA52C853" italics="true" pageId="0" pageNumber="143">Redescription.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="143">
<paragraph id="8BFCD626219147A3B3D76F0BF82B9BE7" pageId="0" pageNumber="143">
Robust with fastigium projecting well beyond scapus. Males with contrasting coloration: fastigium blackish, dorsal surface of pronotum medially dark, in prozona and metazona blackish, in metazona broadly trapezoideal and fringed by a pair of ovoid elongate whitish lateral maculae (Fig.
<figureCitation captionStart="Fig. 8" captionStartId="F8" captionText="Fig. 8. Cestrophorus spp. male genitalia compared. A. White pronotal maculae of C. paradoxus; B. Parameres of C. paradoxus ' upside-down dancing anteaters'; C. Dorsal aspect C. ditachus parameres; D. Lateral view of C. ditachus paramere; E. Photo of terminalia of C. amplitenuis; F. Lateral aspect of C. amplitenuis paramere." figureDoi="10.3897/jor.31.82306.fig8" httpUri="https://binary.pensoft.net/fig/750937" pageId="0" pageNumber="143">8A</figureCitation>
<figureCitation id="4779161206DCEF0B40C3266620DCE48F" captionStart="Fig. 8" captionStartId="F8" captionText="Fig. 8. Cestrophorus spp. male genitalia compared. A. White pronotal maculae of C. paradoxus; B. Parameres of C. paradoxus ' upside-down dancing anteaters'; C. Dorsal aspect C. ditachus parameres; D. Lateral view of C. ditachus paramere; E. Photo of terminalia of C. amplitenuis; F. Lateral aspect of C. amplitenuis paramere." figureDoi="10.3897/jor.31.82306.fig8" httpUri="https://binary.pensoft.net/fig/750937" pageId="0" pageNumber="143">8A</figureCitation>
), thin distal portion of hind tibiae blackish. Male tegmina greenish, broadened beyond width of pronotum and covering abdomen completely. Male cerci approximately broadly triangular in dorsal and lateral view, obtuse medial lobes with inconspicuous obtuse and recurved hook, almost touching in the middle, cercus terminating in sharp upcurved spine, and at the base with long and more or less perpendicularly upcurved spine (up to beyond the height of the dorsal hook).
</paragraph>
<caption doi="10.3897/jor.31.82306.fig7" httpUri="https://binary.pensoft.net/fig/750936" pageId="0" pageNumber="143" start="Fig. 7" startId="F7">
<paragraph pageId="0" pageNumber="143">
<emphasis bold="true" pageId="0" pageNumber="143">Fig. 7.</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="C. amplitenuis" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="amplitenuis">
<emphasis italics="true" pageId="0" pageNumber="143">C. amplitenuis</emphasis>
<caption id="E6845B4C955C2CEA3A43C3F43448A8D6" doi="10.3897/jor.31.82306.fig7" httpUri="https://binary.pensoft.net/fig/750936" pageId="0" pageNumber="143" start="Fig. 7" startId="F7">
<paragraph id="2AE1072D2FA6235467A538262B2D76BB" pageId="0" pageNumber="143">
<emphasis id="165F8559ED774085EFA34EDCA39DFF2A" bold="true" pageId="0" pageNumber="143">Fig. 7.</emphasis>
<taxonomicName id="686E75A6887F5D55CF53615450419E18" class="Insecta" kingdom="Animalia" lsidName="C. amplitenuis" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="amplitenuis">
<emphasis id="FC1DA137F816CF4F882038234FB7BF9E" italics="true" pageId="0" pageNumber="143">C. amplitenuis</emphasis>
</taxonomicName>
pair mating in the field; part of a large deme of singers in shrubby roadside vegetation at the type locality near
<normalizedToken originalValue="Ecuadors">Ecuador's</normalizedToken>
<normalizedToken id="1C887CB9122579F14EF439222F464D41" originalValue="Ecuadors">Ecuador's</normalizedToken>
northern border with Colombia. Female lacks wings.
</paragraph>
</caption>
<caption doi="10.3897/jor.31.82306.fig8" httpUri="https://binary.pensoft.net/fig/750937" pageId="0" pageNumber="143" start="Fig. 8" startId="F8">
<paragraph pageId="0" pageNumber="143">
<emphasis bold="true" pageId="0" pageNumber="143">Fig. 8.</emphasis>
<taxonomicName authorityName="Redtenbacher" authorityYear="1891" class="Insecta" family="Tettigoniidae" genus="Cestrophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cestrophorus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="143">Cestrophorus</emphasis>
<caption id="8E894B39274F6171C885720640F44B0B" doi="10.3897/jor.31.82306.fig8" httpUri="https://binary.pensoft.net/fig/750937" pageId="0" pageNumber="143" start="Fig. 8" startId="F8">
<paragraph id="D801B8B06A979E9321111E3451CEC328" pageId="0" pageNumber="143">
<emphasis id="CED48C33B35C80571CBB8210C007A4D0" bold="true" pageId="0" pageNumber="143">Fig. 8.</emphasis>
<taxonomicName id="0BAEE698A1819AE376420770898D47AA" authorityName="Redtenbacher" authorityYear="1891" class="Insecta" family="Tettigoniidae" genus="Cestrophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cestrophorus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
<emphasis id="B943C4EDB171A7E7D47390B0297653DC" italics="true" pageId="0" pageNumber="143">Cestrophorus</emphasis>
</taxonomicName>
spp. male genitalia compared.
<emphasis bold="true" pageId="0" pageNumber="143">A.</emphasis>
<emphasis id="DDE1532DAA041C3F747A8CE6C13A56E6" bold="true" pageId="0" pageNumber="143">A.</emphasis>
White pronotal maculae of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="C. paradoxus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis italics="true" pageId="0" pageNumber="143">C. paradoxus</emphasis>
<taxonomicName id="72884EA1EC7A707EBD52E35EA1952414" class="Insecta" kingdom="Animalia" lsidName="C. paradoxus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="FA5850D7CEBF7CAFF307F7A27742F0C2" italics="true" pageId="0" pageNumber="143">C. paradoxus</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="143">B.</emphasis>
<emphasis id="CAD9B8F70042E539579481944927814A" bold="true" pageId="0" pageNumber="143">B.</emphasis>
Parameres of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="C. paradoxus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis italics="true" pageId="0" pageNumber="143">C. paradoxus</emphasis>
<taxonomicName id="FE36D47278F9809434C46B2FF4866780" class="Insecta" kingdom="Animalia" lsidName="C. paradoxus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="paradoxus">
<emphasis id="D6526BD7BF44A26BA900FA1C43EFCD84" italics="true" pageId="0" pageNumber="143">C. paradoxus</emphasis>
</taxonomicName>
'upside-down dancing
<normalizedToken originalValue="anteaters">anteaters'</normalizedToken>
<normalizedToken id="0DE31DD6F4CE01C72B817F271CF03BDE" originalValue="anteaters">anteaters'</normalizedToken>
;
<emphasis bold="true" pageId="0" pageNumber="143">C.</emphasis>
<emphasis id="BFBFD7629E5BA40A78511D3DF64D37F4" bold="true" pageId="0" pageNumber="143">C.</emphasis>
Dorsal aspect
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="C. ditachus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ditachus">
<emphasis italics="true" pageId="0" pageNumber="143">C. ditachus</emphasis>
<taxonomicName id="0CB0CFE0E492AB4AED2B6FA2E86E0660" class="Insecta" kingdom="Animalia" lsidName="C. ditachus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ditachus">
<emphasis id="58465147E8B7F9567F226216F203C02D" italics="true" pageId="0" pageNumber="143">C. ditachus</emphasis>
</taxonomicName>
parameres;
<emphasis bold="true" pageId="0" pageNumber="143">D.</emphasis>
<emphasis id="FEBE0D236713C2BE789029AF5DFFDBBA" bold="true" pageId="0" pageNumber="143">D.</emphasis>
Lateral view of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="C. ditachus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ditachus">
<emphasis italics="true" pageId="0" pageNumber="143">C. ditachus</emphasis>
<taxonomicName id="BADFD270AB41C50D5944D6247A527796" class="Insecta" kingdom="Animalia" lsidName="C. ditachus" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="ditachus">
<emphasis id="604AFCD1B8D2EFF73CEDFDD5C1F45812" italics="true" pageId="0" pageNumber="143">C. ditachus</emphasis>
</taxonomicName>
paramere;
<emphasis bold="true" pageId="0" pageNumber="143">E.</emphasis>
<emphasis id="7AE881C2C5EAFCB925D072A0E142B351" bold="true" pageId="0" pageNumber="143">E.</emphasis>
Photo of terminalia of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="C. amplitenuis" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="amplitenuis">
<emphasis italics="true" pageId="0" pageNumber="143">C. amplitenuis</emphasis>
<taxonomicName id="AC00F91A61A992E75504CD20A1C97E90" class="Insecta" kingdom="Animalia" lsidName="C. amplitenuis" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="amplitenuis">
<emphasis id="99839CB077F939F82EFC39FF3AE5D78C" italics="true" pageId="0" pageNumber="143">C. amplitenuis</emphasis>
</taxonomicName>
;
<emphasis bold="true" pageId="0" pageNumber="143">F.</emphasis>
<emphasis id="43476984F77A261F329AE2B49F0507A3" bold="true" pageId="0" pageNumber="143">F.</emphasis>
Lateral aspect of
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="C. amplitenuis" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="amplitenuis">
<emphasis italics="true" pageId="0" pageNumber="143">C. amplitenuis</emphasis>
<taxonomicName id="85E3D8291518E43BDB5F3A761FC7E0FD" class="Insecta" kingdom="Animalia" lsidName="C. amplitenuis" order="Orthoptera" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="amplitenuis">
<emphasis id="484E183BDE3452DE898335A729EA4E58" italics="true" pageId="0" pageNumber="143">C. amplitenuis</emphasis>
</taxonomicName>
paramere.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="description">
<paragraph pageId="0" pageNumber="143">
<emphasis italics="true" pageId="0" pageNumber="143">Measurements.</emphasis>
<subSubSection id="5679E9A08F4C1D8595BD5541771C9047" pageId="0" pageNumber="143" type="description">
<paragraph id="6ABDA49654176A2FDE16C4185D12D351" pageId="0" pageNumber="143">
<emphasis id="4AB60B4A6912575BB206BCC24B7FF175" italics="true" pageId="0" pageNumber="143">Measurements.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="143">Male: midline pronotum 4.6 mm, fastigium verticis 1.8 mm, tegmina 9.2 mm; female: midline pronotum 3.8 mm, fastigium verticis 1.7 mm (based on two specimens, no apparent variation in size).</paragraph>
<paragraph id="FAD4B045A74931630C990D11D2140162" pageId="0" pageNumber="143">Male: midline pronotum 4.6 mm, fastigium verticis 1.8 mm, tegmina 9.2 mm; female: midline pronotum 3.8 mm, fastigium verticis 1.7 mm (based on two specimens, no apparent variation in size).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="143" type="note.">
<paragraph pageId="0" pageNumber="143">
<emphasis italics="true" pageId="0" pageNumber="143">Note.</emphasis>
<subSubSection id="CC1401045E5FD94998DBB0CF5DB1D81E" pageId="0" pageNumber="143" type="note.">
<paragraph id="CC95247753B48A34573843F9EDA88AE3" pageId="0" pageNumber="143">
<emphasis id="4F88886AB846B684C6D870B46E2B079F" italics="true" pageId="0" pageNumber="143">Note.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="143">
<paragraph id="727F0BF3D37049A98F94A948FD859041" pageId="0" pageNumber="143">
All specimens-in February 1988 and again in July 2003-were taken from beside a forest path tracing the south bank of the
<normalizedToken originalValue="Río">Rio</normalizedToken>
<normalizedToken id="3684293AAE8C442B8ECC9A34E69B4F41" originalValue="Río">Rio</normalizedToken>
Aliso upstream; the site is reached by a (very) secondary road branching west from the main highway just before the town of Cosanga. About 3 km along this sideroad is the lodge
<normalizedToken originalValue="Cabañas">Cabanas</normalizedToken>
<normalizedToken id="664F29624A3AC3007DCDCC6B44DD22D7" originalValue="Cabañas">Cabanas</normalizedToken>
San Isidro; a few km farther on, a posted sign at an iron bridge identifies the
<normalizedToken originalValue="Río">Rio</normalizedToken>
<normalizedToken id="7325BA3645AD7ED2A494E07DBCF279BA" originalValue="Río">Rio</normalizedToken>
Aliso.
</paragraph>
</subSubSection>

View file

@ -1,74 +1,74 @@
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Figs 41, 42, 44, 46, 49
</paragraph>
@ -234,7 +234,7 @@ is highly diverse in color and pattern, illustrated in
, this species is unlike any other in the genus. Although eye-catching in a museum specimen, the color and pattern might be cryptic against a plant with lacy silver green foliage, such as sage (
<taxonomicName id="6012C84A812DBF1E39991435054364A5" class="Magnoliopsida" family="Asteraceae" genus="Artemisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Artemisia" order="Asterales" pageId="25" pageNumber="226" phylum="Tracheophyta" rank="genus">Artemisia</taxonomicName>
spp.,
<taxonomicName id="A6AF1D5C849FF4E4DDC9D405AF0A19FA" family="Asteraceae" lsidName="Xylenina" pageId="25" pageNumber="226" rank="family">Asteraceae</taxonomicName>
<taxonomicName id="A6AF1D5C849FF4E4DDC9D405AF0A19FA" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Xylenina" order="Asterales" pageId="25" pageNumber="226" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
<caption id="059DC5BA6A259E91DA18AA8A42CA0416" pageId="25" pageNumber="226">

View file

@ -1,159 +1,159 @@
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<paragraph pageId="3" pageNumber="70">
<taxonomicName LSID="http://species-id.net/wiki/Cyphopisthes_descarpentriesi" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi Paulian, 1977</taxonomicName>
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<subSubSection id="52BE1D090EB8AD17EF5D71D504149B9D" pageId="3" pageNumber="70" type="nomenclature">
<paragraph id="E3B6EBE47AD440D43BD43FCAEFE7141D" pageId="3" pageNumber="70">
<taxonomicName id="D560FA6899D319C3FA45048CB8167D7E" ID-CoL="33G2C" LSID="http://species-id.net/wiki/Cyphopisthes_descarpentriesi" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi Paulian, 1977</taxonomicName>
Figs 1
<normalizedToken originalValue="AD">A-D</normalizedToken>
<normalizedToken id="10B92022D780F387E7F632BFD748B88E" originalValue="AD">A-D</normalizedToken>
, 5B, 6
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="70" type="reference_group">
<paragraph pageId="3" pageNumber="70">
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
<subSubSection id="8DB0C4F081BFCE57493FFF04CF967000" pageId="3" pageNumber="70" type="reference_group">
<paragraph id="FCDA5945F9CB7CC584EDFB7A01BAEF04" pageId="3" pageNumber="70">
<taxonomicName id="521AA68BFC3455D51245CAB07350145E" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
Paulian, 1977: 263 (description, distribution, biology);
<bibRefCitation author="Paulian, R" journalOrPublisher="Annales de la Societe Entomologique de France (N. S.)" pageId="23" pageNumber="90" pagination="479 - 514" title="Revision des Ceratocanthidae [Col. Scarabaeoidea] II - Les especes orientales et australiennes." volume="14" year="1978">Paulian 1978</bibRefCitation>
<bibRefCitation id="68D543D732DEAFA487D7AC2263E4FE5B" author="Paulian, R" journalOrPublisher="Annales de la Societe Entomologique de France (N. S.)" pageId="23" pageNumber="90" pagination="479 - 514" title="Revision des Ceratocanthidae [Col. Scarabaeoidea] II - Les especes orientales et australiennes." volume="14" year="1978">Paulian 1978</bibRefCitation>
(key, distribution);
<bibRefCitation author="Cassis, G" editor="Houston, WWK" journalOrPublisher="Canberra, AGPS" pageId="23" pageNumber="90" pagination="65 - 66" title="Ceratocanthidae." volume="9" volumeTitle="Zoological catalogue of Australia. Coleoptera: Scarabaeoidea." year="1992">Cassis and Weir 1992</bibRefCitation>
<bibRefCitation id="2B2395913ED8435FCDBC452F2771D8E1" author="Cassis, G" editor="Houston, WWK" journalOrPublisher="Canberra, AGPS" pageId="23" pageNumber="90" pagination="65 - 66" title="Ceratocanthidae." volume="9" volumeTitle="Zoological catalogue of Australia. Coleoptera: Scarabaeoidea." year="1992">Cassis and Weir 1992</bibRefCitation>
(catalogue);
<bibRefCitation author="Grebennikov, VV" journalOrPublisher="Australian Journal of Entomology" pageId="23" pageNumber="90" pagination="367 - 374" title="Larva and pupa of Cyphopisthes descarpentriesi Paulian (Coleoptera: Scarabaeoidea: Ceratocanthidae) and their phylogenetic implications." url="10.1046/j.1440-6055.2002.00307.x" volume="41" year="2002">Grebennikov et al. 2002</bibRefCitation>
<bibRefCitation id="F103BB66B8471775F47DEE501FA49183" author="Grebennikov, VV" journalOrPublisher="Australian Journal of Entomology" pageId="23" pageNumber="90" pagination="367 - 374" title="Larva and pupa of Cyphopisthes descarpentriesi Paulian (Coleoptera: Scarabaeoidea: Ceratocanthidae) and their phylogenetic implications." url="10.1046/j.1440-6055.2002.00307.x" volume="41" year="2002">Grebennikov et al. 2002</bibRefCitation>
(description of larva and pupa);
<bibRefCitation author="Grebennikov, VV" journalOrPublisher="Systematic Entomology" pageId="23" pageNumber="90" pagination="524 - 543" title="Larvae of Ceratocanthidae and Hybosoridae (Coleoptera: Scarabaeoidea): study of morphology, phylogenetic analysis and evidence of paraphyly of Hybosoridae." url="10.1111/j.0307-6970.2004.00257.x" volume="29" year="2004">Grebennikov et al. 2004</bibRefCitation>
<bibRefCitation id="4C3B1C0F323990D5DB2F04ABC3900130" author="Grebennikov, VV" journalOrPublisher="Systematic Entomology" pageId="23" pageNumber="90" pagination="524 - 543" title="Larvae of Ceratocanthidae and Hybosoridae (Coleoptera: Scarabaeoidea): study of morphology, phylogenetic analysis and evidence of paraphyly of Hybosoridae." url="10.1111/j.0307-6970.2004.00257.x" volume="29" year="2004">Grebennikov et al. 2004</bibRefCitation>
(key to larva);
<bibRefCitation author="Hawkeswood, TJ" journalOrPublisher="Calodema" pageId="23" pageNumber="90" pagination="8 - 11" title="Review of literature and the biology of the Australian Ceratocanthidae (Insecta: Coleoptera)." volume="7" year="2006">Hawkeswood 2006</bibRefCitation>
<bibRefCitation id="D5F3ABE427799847D9CDFCF63AD1186F" author="Hawkeswood, TJ" journalOrPublisher="Calodema" pageId="23" pageNumber="90" pagination="8 - 11" title="Review of literature and the biology of the Australian Ceratocanthidae (Insecta: Coleoptera)." volume="7" year="2006">Hawkeswood 2006</bibRefCitation>
(summary of published data);
<bibRefCitation author="Ocampo, FC" journalOrPublisher="Bulletin of the University of Nebraska State Museum" pageId="23" pageNumber="90" pagination="178 - 209" title="Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae (Coleoptera: Scarabaeoidea). 4. Catalog of the subfamilies Anaidinae, Ceratocanthinae, Hybosorinae, Liparochrinae, and Pachyplectrinae (Scarabaeoidea: Hybosoridae)." volume="19" year="2006">Ocampo and Ballerio 2006</bibRefCitation>
<bibRefCitation id="D3024B4FF54B00D5E4B534A8512CDB36" author="Ocampo, FC" journalOrPublisher="Bulletin of the University of Nebraska State Museum" pageId="23" pageNumber="90" pagination="178 - 209" title="Phylogenetic analysis of the scarab family Hybosoridae and monographic revision of the New World subfamily Anaidinae (Coleoptera: Scarabaeoidea). 4. Catalog of the subfamilies Anaidinae, Ceratocanthinae, Hybosorinae, Liparochrinae, and Pachyplectrinae (Scarabaeoidea: Hybosoridae)." volume="19" year="2006">Ocampo and Ballerio 2006</bibRefCitation>
(cheklist).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="70" type="material examined">
<paragraph pageId="3" pageNumber="70">Material examined.</paragraph>
<paragraph pageId="3" pageNumber="70">
<subSubSection id="C3EBDA0749239C47DE6ABE8DC79D3951" pageId="3" pageNumber="70" type="material examined">
<paragraph id="3595DC2BA650AAB60C7CA79B47D8105D" pageId="3" pageNumber="70">Material examined.</paragraph>
<paragraph id="67B2CE26E8979235E4B1712C7F43E459" pageId="3" pageNumber="70">
Holotype, sex undetermined, (ANIC) [enrolled specimen, glued on a point]: Queensland,
<geoCoordinate direction="south" orientation="latitude" precision="555" value="-19.4">19.40S</geoCoordinate>
<geoCoordinate id="4CF9367E3806EEFF1C36BF78AB3648AA" direction="south" orientation="latitude" precision="555" value="-19.4">19.40S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="555" value="146.51">146.51E</geoCoordinate>
<geoCoordinate id="21E2C9CA6B17387CECDD1E17FCF008B8" direction="east" orientation="longitude" precision="555" value="146.51">146.51E</geoCoordinate>
, Lansdown Station, Woodstock, 3 July 1974, #54, J.A.L. Watson in gallery of
<taxonomicName class="Insecta" family="Mastotermitidae" genus="Mastotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mastotermes" order="Blattodea" pageId="3" pageNumber="70" phylum="Arthropoda" rank="genus">Mastotermes</taxonomicName>
<taxonomicName id="305B3413A83630E27BC56BA5F9AECC6E" class="Insecta" family="Mastotermitidae" genus="Mastotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mastotermes" order="Blattodea" pageId="3" pageNumber="70" phylum="Arthropoda" rank="genus">Mastotermes</taxonomicName>
nest / Holotype / Holotype /
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
<taxonomicName id="B1C30127A84B1BF304369F88E8F96592" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
sp. n. R. Paulian det. / ANIC Database no. 25 062056. Examined paratypes: 2 exx., Lansdown Station, via Woodstock, 10 July 1979, R. A. Barrett, with
<taxonomicName class="Insecta" family="Mastotermitidae" genus="Mastotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mastotermes darwiniensis" order="Blattodea" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="darwiniensis">Mastotermes darwiniensis</taxonomicName>
<taxonomicName id="BE102FF452DBFFAB9D9A739A0225C4E6" class="Insecta" family="Mastotermitidae" genus="Mastotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mastotermes darwiniensis" order="Blattodea" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="darwiniensis">Mastotermes darwiniensis</taxonomicName>
(ANIC); 2 exx., Queensland, Pallarenda. Townsville, 1.VII.1974, J. A. L. Watson, in galleries of
<taxonomicName class="Insecta" family="Mastotermitidae" genus="Mastotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mastotermes" order="Blattodea" pageId="3" pageNumber="70" phylum="Arthropoda" rank="genus">Mastotermes</taxonomicName>
<taxonomicName id="48A1560A9891EB62C0D15091DC6782FB" class="Insecta" family="Mastotermitidae" genus="Mastotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mastotermes" order="Blattodea" pageId="3" pageNumber="70" phylum="Arthropoda" rank="genus">Mastotermes</taxonomicName>
(MNHN, ANIC).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="71" pageId="3" pageNumber="70" type="description">
<paragraph pageId="3" pageNumber="70">Description.</paragraph>
<paragraph pageId="3" pageNumber="70">Size: HL = 0.98 mm; HW = 1.54 mm; PL = 1.63 mm; PW = 2.63 mm; EL = 2.87 mm; EW = 2.63 mm. Overall morphology as in generic description. Reddish-brown, shiny, glabrous (very fine short yellowish setation visible at 50x magnification), sternum, tarsi and antennae reddish-brown.</paragraph>
<paragraph pageId="3" pageNumber="70">Head: interocular distance about six times maximum width of dorsal ocular area, punctation dense and impressed, disc with very short transverse comma shaped punctures, each one having a simple small puncture at its interior side, sides of disc with large comma-shaped punctures centrifugally oriented, with opening facing internally, each one having a simple fine puncture internally, anterior portion of clypeus with three to four irregular anastomosing transverse lines.</paragraph>
<paragraph lastPageId="4" lastPageNumber="71" pageId="3" pageNumber="70">
<subSubSection id="7DD7281F2A42A3ED39FD289C88F58443" lastPageId="4" lastPageNumber="71" pageId="3" pageNumber="70" type="description">
<paragraph id="8D66C1C4F23B3FD02B1412DE4E281BB1" pageId="3" pageNumber="70">Description.</paragraph>
<paragraph id="F26283F0B27573850EA00443B2184BE8" pageId="3" pageNumber="70">Size: HL = 0.98 mm; HW = 1.54 mm; PL = 1.63 mm; PW = 2.63 mm; EL = 2.87 mm; EW = 2.63 mm. Overall morphology as in generic description. Reddish-brown, shiny, glabrous (very fine short yellowish setation visible at 50x magnification), sternum, tarsi and antennae reddish-brown.</paragraph>
<paragraph id="0DF5E75AECD1BAD62FB6A2A6999FF400" pageId="3" pageNumber="70">Head: interocular distance about six times maximum width of dorsal ocular area, punctation dense and impressed, disc with very short transverse comma shaped punctures, each one having a simple small puncture at its interior side, sides of disc with large comma-shaped punctures centrifugally oriented, with opening facing internally, each one having a simple fine puncture internally, anterior portion of clypeus with three to four irregular anastomosing transverse lines.</paragraph>
<paragraph id="29D5DFA97A34CC494C216F4980E3F4DB" lastPageId="4" lastPageNumber="71" pageId="3" pageNumber="70">
Pronotum: margin completely bordered, anterior angles angulate, completely covered by large, almost closed, horseshoe-shaped punctures, on disc with a small opening
<pageBreakToken pageId="4" pageNumber="71" start="start">directed</pageBreakToken>
<pageBreakToken id="3767AC7D75A703A255C1C5AA123D9C12" pageId="4" pageNumber="71" start="start">directed</pageBreakToken>
anteriad, at sides punctures larger than on disc, with a small opening directed laterad, each puncture having inside a small setigerous pore. Punctation dense: interpunctural distance being less than puncture diameter.
</paragraph>
<paragraph pageId="4" pageNumber="71">Scutellum: covered by dense horseshoe-shaped punctures with posterior openings.</paragraph>
<paragraph pageId="4" pageNumber="71">Elytra: W/L: 0,93. Humeral callus indistinct, two short longitudinal lines starting at humerus and occupying proximal third, sutural interstria indistinct, completely and uniformly covered by impressed large horseshoe-shaped punctures with a small posterior openings, each one bearing a setigerous pore in the middle. Pseudepipleura with longitudinally oriented anastomosing horseshoe-shaped punctures mixed with comma-shaped punctures.</paragraph>
<caption pageId="4" pageNumber="71">
<paragraph pageId="4" pageNumber="71">
<paragraph id="E09535456060B07D88196BA0A0ADE177" pageId="4" pageNumber="71">Scutellum: covered by dense horseshoe-shaped punctures with posterior openings.</paragraph>
<paragraph id="BF5A4CECA3C675F2471BAFD64AAB4424" pageId="4" pageNumber="71">Elytra: W/L: 0,93. Humeral callus indistinct, two short longitudinal lines starting at humerus and occupying proximal third, sutural interstria indistinct, completely and uniformly covered by impressed large horseshoe-shaped punctures with a small posterior openings, each one bearing a setigerous pore in the middle. Pseudepipleura with longitudinally oriented anastomosing horseshoe-shaped punctures mixed with comma-shaped punctures.</paragraph>
<caption id="D1087A58D66528557C1F5392CA7C5429" pageId="4" pageNumber="71">
<paragraph id="7A3722C568D7D60B6612BCD738F435C3" pageId="4" pageNumber="71">
Figure 1.
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
<taxonomicName id="B06C32E79FA2E86911D92DD747ABAB2D" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
Paulian, 1977. A Extended Paratype, dorsal view B enrolled Paratype, ventral view C enrolled Paratype, dorsal view D enrolled Paratype, lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="4" pageNumber="71" type="diagnosis">
<paragraph pageId="4" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="71">
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
<subSubSection id="1E7DA3E8FF0B89B5EFE6FEB8B7AE7206" pageId="4" pageNumber="71" type="diagnosis">
<paragraph id="5897793E8DD8BD303C0316B011980144" pageId="4" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="750C144A940AD6372C502176E429F4C6" pageId="4" pageNumber="71">
<taxonomicName id="9EC4FAD3BA8C3B7C1C763E010E4CB36E" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
is unique among all other known Australian
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="genus">Cyphopisthes</taxonomicName>
<taxonomicName id="766F6D52B0D6A34D2A0E23A2D9921CDA" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes" order="Coleoptera" pageId="4" pageNumber="71" phylum="Arthropoda" rank="genus">Cyphopisthes</taxonomicName>
in having the pronotum and the elytra completely covered by such large and dense uniform horseshoe-shaped punctures with small posterior openings. All other known members of the genus have remarkably finer punctation and elytral punctation differs from the pronotal one.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="71" type="etymology">
<paragraph pageId="4" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="71">
<subSubSection id="57606A15BD7B35F3F739FA6DEA234BA8" pageId="4" pageNumber="71" type="etymology">
<paragraph id="D525FA32DEBB4324887063F64920B952" pageId="4" pageNumber="71">Etymology.</paragraph>
<paragraph id="06CFE54F5FFB77877284716A0E7B50A1" pageId="4" pageNumber="71">
Dedicated to
<normalizedToken originalValue="André">Andre</normalizedToken>
<normalizedToken id="0B4C6EE5F3BDC3990CBE642430923740" originalValue="André">Andre</normalizedToken>
Descarpentries (1919-1998) of MNHN.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="72" pageId="4" pageNumber="71" type="distribution">
<paragraph pageId="4" pageNumber="71">Distribution and habitat.</paragraph>
<paragraph lastPageId="5" lastPageNumber="72" pageId="4" pageNumber="71">
<subSubSection id="74EC29430937BF53CDDA83EF6F403BAC" lastPageId="5" lastPageNumber="72" pageId="4" pageNumber="71" type="distribution">
<paragraph id="BF3AD27F4D6A42F5A3CE546235597A76" pageId="4" pageNumber="71">Distribution and habitat.</paragraph>
<paragraph id="4F09125C1CE32AD8CB579F68E5BD8F15" lastPageId="5" lastPageNumber="72" pageId="4" pageNumber="71">
Known from north eastern Queensland coast. All specimens have been collected in open eucalypt woodland, in nests of
<taxonomicName class="Insecta" family="Mastotermitidae" genus="Mastotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mastotermes" order="Blattodea" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="5" pageNumber="72" start="start">Mastotermes</pageBreakToken>
<taxonomicName id="AB56B02D4096173969D532CB1A18CC70" class="Insecta" family="Mastotermitidae" genus="Mastotermes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mastotermes" order="Blattodea" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">
<pageBreakToken id="4395322F635D790F8D1D5936D2E8149B" pageId="5" pageNumber="72" start="start">Mastotermes</pageBreakToken>
</taxonomicName>
(
<taxonomicName class="Magnoliopsida" family="Dipterocarpaceae" genus="Isoptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Isoptera" order="Malvales" pageId="5" pageNumber="72" phylum="Tracheophyta" rank="genus">Isoptera</taxonomicName>
<taxonomicName id="C6315B492E29390C93ABE13EC70C13B7" class="Magnoliopsida" family="Dipterocarpaceae" genus="Isoptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Isoptera" order="Malvales" pageId="5" pageNumber="72" phylum="Tracheophyta" rank="genus">Isoptera</taxonomicName>
). Termitophily has already been reported for
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">Cyphopisthes</taxonomicName>
<taxonomicName id="57C4DAA11C3359AB581A3CBFF5C74470" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">Cyphopisthes</taxonomicName>
(
<bibRefCitation author="Ballerio, A" editor="Ratcliffe, B" journalOrPublisher="ZooKeys" pageId="22" pageNumber="89" pagination="77 - 104" title="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)." url="10.3897/zookeys.34.268" volume="34" volumeTitle="Current advances in Scarabaeoidea research." year="2010">Ballerio and Maruyama 2010</bibRefCitation>
<bibRefCitation id="3FFA4A04EC11815C54BD759300364B0F" author="Ballerio, A" editor="Ratcliffe, B" journalOrPublisher="ZooKeys" pageId="22" pageNumber="89" pagination="77 - 104" title="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)." url="10.3897/zookeys.34.268" volume="34" volumeTitle="Current advances in Scarabaeoidea research." year="2010">Ballerio and Maruyama 2010</bibRefCitation>
). In the same nests larvae and pupae were collected, subsequently described by
<bibRefCitation author="Grebennikov, VV" journalOrPublisher="Australian Journal of Entomology" pageId="23" pageNumber="90" pagination="367 - 374" title="Larva and pupa of Cyphopisthes descarpentriesi Paulian (Coleoptera: Scarabaeoidea: Ceratocanthidae) and their phylogenetic implications." url="10.1046/j.1440-6055.2002.00307.x" volume="41" year="2002">Grebennikov et al. (2002)</bibRefCitation>
<bibRefCitation id="E132D824D2CE99B5347CF4940F3B741B" author="Grebennikov, VV" journalOrPublisher="Australian Journal of Entomology" pageId="23" pageNumber="90" pagination="367 - 374" title="Larva and pupa of Cyphopisthes descarpentriesi Paulian (Coleoptera: Scarabaeoidea: Ceratocanthidae) and their phylogenetic implications." url="10.1046/j.1440-6055.2002.00307.x" volume="41" year="2002">Grebennikov et al. (2002)</bibRefCitation>
. Open eucalypt woodland is a very unusual habitat for a
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">Cyphopisthes</taxonomicName>
<taxonomicName id="0FCB3BA9D5EAF0C885DC57FC4AE26394" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="genus">Cyphopisthes</taxonomicName>
, since most species are rainforest dwellers.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="72" type="remarks">
<paragraph pageId="5" pageNumber="72">Remarks.</paragraph>
<paragraph pageId="5" pageNumber="72">
<subSubSection id="1E2BCC13F1598980853CD60E282B41BB" pageId="5" pageNumber="72" type="remarks">
<paragraph id="3274DD77EE6A8819334BE91E125BF273" pageId="5" pageNumber="72">Remarks.</paragraph>
<paragraph id="BAF4BC0910ADE7A72CAA41E534AF2437" pageId="5" pageNumber="72">
The type series contained three different species (two of them new to science and described below). Among paratypes of true
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
<taxonomicName id="1644E47DFDC8E5BAF7C83D2B699143A1" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
in ANIC one specimen had the genal canthus shortened, leaving a distinct gap between its tip and the occipital area.
</paragraph>
<paragraph pageId="5" pageNumber="72">
<bibRefCitation author="Paulian, R" journalOrPublisher="Annales de la Societe Entomologique de France (N. S.)" pageId="23" pageNumber="90" pagination="479 - 514" title="Revision des Ceratocanthidae [Col. Scarabaeoidea] II - Les especes orientales et australiennes." volume="14" year="1978">Paulian (1978)</bibRefCitation>
<paragraph id="C419563E853DD107BDF259CA9438A082" pageId="5" pageNumber="72">
<bibRefCitation id="B3D31C722D292835A4936BFCE9C15198" author="Paulian, R" journalOrPublisher="Annales de la Societe Entomologique de France (N. S.)" pageId="23" pageNumber="90" pagination="479 - 514" title="Revision des Ceratocanthidae [Col. Scarabaeoidea] II - Les especes orientales et australiennes." volume="14" year="1978">Paulian (1978)</bibRefCitation>
identified as
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
<taxonomicName id="85B26ADEA8FA35B53BAB8B05EEBBA3CE" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
also an old specimen in RMNH, ex coll. Pascoe, labelled &quot;Mus. Godeffroy, Peak Down, Austr./10791/
<taxonomicName class="Insecta" family="Hybosoridae" genus="Synarmostes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synarmostes acromialis" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="acromialis">Synarmostes acromialis</taxonomicName>
<taxonomicName id="D3CDEB8F9F0A95ADB679F62C2E242213" class="Insecta" family="Hybosoridae" genus="Synarmostes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Synarmostes acromialis" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="acromialis">Synarmostes acromialis</taxonomicName>
Pascoe&quot;. Probably the correct locality name should be &quot;Peak Downs&quot; (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-22.25">22°15'S</geoCoordinate>
<geoCoordinate id="A0CC22D5EF52B44D9DB054FEA26830AE" direction="south" orientation="latitude" precision="925" value="-22.25">22°15'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="148.18333">148°11'E</geoCoordinate>
<geoCoordinate id="869A36F8CAE79622795CA58F9415FA10" direction="east" orientation="longitude" precision="925" value="148.18333">148°11'E</geoCoordinate>
) in Central Queensland (Federica Turco, pers. comm.). I examined two photographs of that specimen, kindly provided by Hans Huijbregts (RMNH): it is certainly not a
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
<taxonomicName id="01BDCB6A84A5885517E3BF2F6C9507C1" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes descarpentriesi" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="descarpentriesi">Cyphopisthes descarpentriesi</taxonomicName>
, and probably represents a specimen of
<taxonomicName class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes monteithi" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="monteithi">Cyphopisthes monteithi</taxonomicName>
<taxonomicName id="572B38FB67CDB25A860F7B15591C1DD4" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphopisthes monteithi" order="Coleoptera" pageId="5" pageNumber="72" phylum="Arthropoda" rank="species" species="monteithi">Cyphopisthes monteithi</taxonomicName>
sp. n. or another species very close to it.
</paragraph>
</subSubSection>

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@ -1,101 +1,101 @@
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<subSubSection id="303ABE69499A165AA7A0C5493BFFBCD1" pageId="9" pageNumber="24" type="nomenclature">
<paragraph id="DFDB5D0CA3210BF97CC5E0EBE6805DAB" pageId="9" pageNumber="24">
<taxonomicName id="2A91AD28751910D0AAD90B6993FD5E19" ID-CoL="8T7ML" LSID="urn:lsid:zoobank.org:act:5C1328B7-9A79-445A-AFB6-BC5402DFC36C" authority="Bochkov &amp; Literak" class="Arachnida" family="Harpirhynchidae" genus="Neharpyrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neharpyrhynchus tangara" order="Trombidiformes" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="tangara">Neharpyrhynchus tangara Bochkov &amp; Literak</taxonomicName>
<taxonomicNameLabel id="FAD76BAC17166758A67B78050B87ED0A" pageId="9" pageNumber="24">sp. n.</taxonomicNameLabel>
Fig. 5
<normalizedToken originalValue="DF">D-F</normalizedToken>
<normalizedToken id="CD4DFBEC719F6A1B4550D7618C345BA4" originalValue="DF">D-F</normalizedToken>
6C
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="24" type="type material">
<paragraph pageId="9" pageNumber="24">Type material.</paragraph>
<paragraph pageId="9" pageNumber="24">
<subSubSection id="6D7BB9ECBD61E30EA5EDC6D697A25A1E" pageId="9" pageNumber="24" type="type material">
<paragraph id="26968E177732DC014E333DE2BDFE1BA2" pageId="9" pageNumber="24">Type material.</paragraph>
<paragraph id="526D2F00EC836FFB53E96BD76BE03BF1" pageId="9" pageNumber="24">
Female holotype (MZUSP), 20 female paratypes (ZISP AVB 10-1210-003, 1-20) on slides and numerous paratypes preserved in alcohol from
<taxonomicName class="Aves" family="Thraupidae" genus="Tangara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tangara cayana" order="Passeriformes" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="cayana">Tangara cayana</taxonomicName>
<taxonomicName id="3807BC5FA9ACCE8DF457DE12C62B9810" class="Aves" family="Thraupidae" genus="Tangara" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tangara cayana" order="Passeriformes" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="cayana">Tangara cayana</taxonomicName>
(Linnaeus) (
<taxonomicName genus="Passeriformes" lsidName="Passeriformes" pageId="9" pageNumber="24" rank="genus">Passeriformes</taxonomicName>
<taxonomicName id="CADAB407DC95C9D4F17BC8BA5C919803" genus="Passeriformes" lsidName="Passeriformes" pageId="9" pageNumber="24" rank="genus">Passeriformes</taxonomicName>
,
<taxonomicName family="Thraupidae" lsidName="" pageId="9" pageNumber="24" rank="family">Thraupidae</taxonomicName>
<taxonomicName id="42747031031D5D8CDE7F367CC072D3E5" class="Aves" family="Thraupidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Passeriformes" pageId="9" pageNumber="24" phylum="Chordata" rank="family">Thraupidae</taxonomicName>
) [feathers around ear apertures, back of the head and neck], BRAZIL: Minas Gerais, Belo Horizonte, Nova Lima,
<normalizedToken originalValue="Água">Agua</normalizedToken>
<normalizedToken id="67B1465444543393B9491825B8D92D40" originalValue="Água">Agua</normalizedToken>
Limpa,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-20.216667">20°13'S</geoCoordinate>
<geoCoordinate id="907077831A65FE5883D1A43DF1D9A3E7" direction="south" orientation="latitude" precision="925" value="-20.216667">20°13'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-43.933334">43°56'W</geoCoordinate>
<geoCoordinate id="F9330801FBABA06E3D26B05A64C486E3" direction="west" orientation="longitude" precision="925" value="-43.933334">43°56'W</geoCoordinate>
, 31 August 2010, coll. S.V. Mironov et al. (SVM-10-0831-12).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="24" type="type deposition">
<paragraph pageId="9" pageNumber="24">Type deposition.</paragraph>
<paragraph pageId="9" pageNumber="24">Holotype and 10 paratypes deposited in the MZUSP, six paratypes in the ZISP, 2 paratypes in the UMMZ, and 2 paratypes in the IPCR. Alcohol preserved paratypes deposited in the MZUSP and ZISP.</paragraph>
<subSubSection id="31F35A2A492A1C3850CB71C169B1D10D" pageId="9" pageNumber="24" type="type deposition">
<paragraph id="B94EDE152AB2CB2E4D00608CFFEC78D6" pageId="9" pageNumber="24">Type deposition.</paragraph>
<paragraph id="6D49EED549143510C3107972BD093DA6" pageId="9" pageNumber="24">Holotype and 10 paratypes deposited in the MZUSP, six paratypes in the ZISP, 2 paratypes in the UMMZ, and 2 paratypes in the IPCR. Alcohol preserved paratypes deposited in the MZUSP and ZISP.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="24" type="description">
<paragraph pageId="9" pageNumber="24">Description.</paragraph>
<subSection pageId="9" pageNumber="24" type="female (holotype)">
<paragraph pageId="9" pageNumber="24">Female (holotype).</paragraph>
<paragraph pageId="9" pageNumber="24">Idiosoma, including gnathosoma, 600 long (600-620 in 10 paratypes), 380 wide (380-400). Gnathosoma 125 long (125-130), 150 wide (145-150). Palps 60 long, distinctly inflated dorsally. All palpalae distinctly pectinate (Fig. 5D). Lengths of palpalae: dF 35 (35-40), dG 30 (29-33), and l&quot;G 29 (30-32); dF only slightly longer than dG and l&quot;G. Setae vF about 90 long, smooth. Subcapitulum ventrally with setae n and m, about 70 long. Peritremal branch about 110 long. Idiosoma saccate, 500 long (500-520). Anterior region of propodonotum covered by short rounded scales situated irregularly in its posterior half (4E, 5C). Dorsal shield entire, 190 long in midline (180-200), 330 at maximum width (330-350). Anterior margin of dorsal shield almost straight with pair of lateral anteriad directed projections; its posterior margin almost straight (4F). This shield covered by fine rhomboid-like pattern, almost indistinct in anterior half and slightly better discernible in posterior half. Ventral surface of idiosoma with indistinct transverse striations, without scales or verrucosities. Setal lengths: vi, ve, and si - all distinctly barbed, subequal in length, 150-170; se, c2,and 1a 8-12- all smooth; h1 whip-like, 230 (210-240); 1b smooth, 30-35, 3a absent. Legs as in previous species.</paragraph>
<subSubSection id="2A9B76AC621E6B923527DA6AF18B240A" pageId="9" pageNumber="24" type="description">
<paragraph id="B4DAF89E7B3DE172F3030D49BEA984D9" pageId="9" pageNumber="24">Description.</paragraph>
<subSection id="83C5CA4BF45259DF25A3587EB31B5815" pageId="9" pageNumber="24" type="female (holotype)">
<paragraph id="BE8BAED333B5B631535DFF69739854B8" pageId="9" pageNumber="24">Female (holotype).</paragraph>
<paragraph id="C9FD3FDBA075DEF061737F41C40CC23D" pageId="9" pageNumber="24">Idiosoma, including gnathosoma, 600 long (600-620 in 10 paratypes), 380 wide (380-400). Gnathosoma 125 long (125-130), 150 wide (145-150). Palps 60 long, distinctly inflated dorsally. All palpalae distinctly pectinate (Fig. 5D). Lengths of palpalae: dF 35 (35-40), dG 30 (29-33), and l&quot;G 29 (30-32); dF only slightly longer than dG and l&quot;G. Setae vF about 90 long, smooth. Subcapitulum ventrally with setae n and m, about 70 long. Peritremal branch about 110 long. Idiosoma saccate, 500 long (500-520). Anterior region of propodonotum covered by short rounded scales situated irregularly in its posterior half (4E, 5C). Dorsal shield entire, 190 long in midline (180-200), 330 at maximum width (330-350). Anterior margin of dorsal shield almost straight with pair of lateral anteriad directed projections; its posterior margin almost straight (4F). This shield covered by fine rhomboid-like pattern, almost indistinct in anterior half and slightly better discernible in posterior half. Ventral surface of idiosoma with indistinct transverse striations, without scales or verrucosities. Setal lengths: vi, ve, and si - all distinctly barbed, subequal in length, 150-170; se, c2,and 1a 8-12- all smooth; h1 whip-like, 230 (210-240); 1b smooth, 30-35, 3a absent. Legs as in previous species.</paragraph>
</subSection>
<subSection pageId="9" pageNumber="24" type="male">
<paragraph pageId="9" pageNumber="24">Male.</paragraph>
<paragraph pageId="9" pageNumber="24">Unknown.</paragraph>
<subSection id="0EB6D199DFC24535580E1592867E1914" pageId="9" pageNumber="24" type="male">
<paragraph id="DC3BBEFC8A8FB95E6DBE1D4A84B0CED5" pageId="9" pageNumber="24">Male.</paragraph>
<paragraph id="8E0A4AE8D06B94111F600F5F196DEB59" pageId="9" pageNumber="24">Unknown.</paragraph>
</subSection>
</subSubSection>
<subSubSection pageId="9" pageNumber="24" type="etymology">
<paragraph pageId="9" pageNumber="24">Etymology.</paragraph>
<paragraph pageId="9" pageNumber="24">The species name derives from the generic name of the host and is a noun in apposition.</paragraph>
<subSubSection id="57A3837EBC8A6E9C597D34050E8D4399" pageId="9" pageNumber="24" type="etymology">
<paragraph id="5C7DD4851203E79EF822517C44AE7769" pageId="9" pageNumber="24">Etymology.</paragraph>
<paragraph id="F8F535DAF443310F2222E56FF9A10F05" pageId="9" pageNumber="24">The species name derives from the generic name of the host and is a noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="24" type="differential diagnosis">
<paragraph pageId="9" pageNumber="24">Differential diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="24">
<subSubSection id="613122FDD0B2649544D5B5E6C28A5571" pageId="9" pageNumber="24" type="differential diagnosis">
<paragraph id="FAF1C8AD3E8FDB2CB545C6CF8EB1CCB1" pageId="9" pageNumber="24">Differential diagnosis.</paragraph>
<paragraph id="175F159CC9FD05293BD3011C481258CB" pageId="9" pageNumber="24">
This new species is closest to
<taxonomicName family="Harpirhynchidae" genus="Neharpyrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neharpyrhynchus mironovi" order="Trombidiformes" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="mironovi">Neharpyrhynchus mironovi</taxonomicName>
<taxonomicName id="F1C69B32DF477F8B0F4E1871E68F7EAB" class="Arachnida" family="Harpirhynchidae" genus="Neharpyrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neharpyrhynchus mironovi" order="Trombidiformes" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="mironovi">Neharpyrhynchus mironovi</taxonomicName>
and differs by the following characters. In females of
<taxonomicName family="Harpirhynchidae" genus="Neharpyrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neharpyrhynchus tanagra" order="Trombidiformes" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="tanagra">Neharpyrhynchus tanagra</taxonomicName>
<taxonomicName id="18456B6A4B5563A1047C7C640AEB53D0" class="Arachnida" family="Harpirhynchidae" genus="Neharpyrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neharpyrhynchus tanagra" order="Trombidiformes" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="tanagra">Neharpyrhynchus tanagra</taxonomicName>
, setae c2 are 8-12 long, the posterior margin of the dorsal shield is almost straight. In
<taxonomicName family="Harpirhynchidae" genus="Neharpyrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neharpyrhynchus mironovi" order="Trombidiformes" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="mironovi">Neharpyrhynchus mironovi</taxonomicName>
<taxonomicName id="A37760BC06540E08F71D625E7665FBD1" class="Arachnida" family="Harpirhynchidae" genus="Neharpyrhynchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neharpyrhynchus mironovi" order="Trombidiformes" pageId="9" pageNumber="24" phylum="Arthropoda" rank="species" species="mironovi">Neharpyrhynchus mironovi</taxonomicName>
, setae c2 are 50-60 long, the posterior margin of the dorsal shield is widely concave. Both species are collected from the hosts belonging to the family
<taxonomicName family="Thraupidae" lsidName="" pageId="9" pageNumber="24" rank="family">Thraupidae</taxonomicName>
<taxonomicName id="E2097D233D9AD9F4B15C34FCF418D0B8" class="Aves" family="Thraupidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Passeriformes" pageId="9" pageNumber="24" phylum="Chordata" rank="family">Thraupidae</taxonomicName>
.
</paragraph>
</subSubSection>