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<mods:title>Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:title id="35F0545E74E2917B02614B87BFC1F4DD">Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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<mods:namePart>Surbakti, Suriani</mods:namePart>
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<mods:namePart>Sumoked, Bob</mods:namePart>
<mods:affiliation>Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
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<mods:namePart>Balke, Michael</mods:namePart>
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<treatment ID-GBIF-Taxon="180680999" LSID="urn:lsid:zoobank.org:act:44511064-4B35-4242-8018-898B0BAB4B0C" httpUri="http://treatment.plazi.org/id/367E9AA59B905821889868232A82AAEA" lastPageNumber="45" pageId="0" pageNumber="45">
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<taxonomicName LSID="http://zoobank.org/44511064-4B35-4242-8018-898B0BAB4B0C" authority="Shaverdo, Surbakti &amp; Balke" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina riberai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberai" status="sp. nov.">Exocelina riberai Shaverdo, Surbakti &amp; Balke</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">Figures 3</figureCitation>
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">, 7</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">, 15</figureCitation>
<treatment id="367E9AA59B905821889868232A82AAEA" ID-GBIF-Taxon="180680999" LSID="urn:lsid:zoobank.org:act:44511064-4B35-4242-8018-898B0BAB4B0C" httpUri="http://treatment.plazi.org/id/367E9AA59B905821889868232A82AAEA" lastPageNumber="45" pageId="0" pageNumber="45" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
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<paragraph id="F63B469AD780C348E972BF6C7DEB96A5" pageId="0" pageNumber="45">
<taxonomicName id="64FDBC4FF2A80EE1CE943E219FC6F04F" ID-CoL="8LZFG" LSID="http://zoobank.org/44511064-4B35-4242-8018-898B0BAB4B0C" authority="Shaverdo, Surbakti &amp; Balke" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina riberai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberai" status="sp. nov.">Exocelina riberai Shaverdo, Surbakti &amp; Balke</taxonomicName>
<taxonomicNameLabel id="D68492835755E8157FB1ADB4CA8FF6D4" pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation id="17CFCE7972AE2E36D3ECE2C47EFA74E4" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">Figures 3</figureCitation>
<figureCitation id="733D8C4214BAB4AE3BE0561B8E33B7EE" captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">, 7</figureCitation>
<figureCitation id="510B00500F1E64D49D956C442017B42C" captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">, 15</figureCitation>
</paragraph>
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<taxonomicName authority="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke, 2021" authorityName="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus" status="sp. nov.">Exocelina</taxonomicName>
<normalizedToken originalValue="“Foja”">&quot;Foja&quot;</normalizedToken>
<subSubSection id="BE7CCA99DD549EDAFC5EB46BDA234F5D" pageId="0" pageNumber="45" type="reference_group">
<paragraph id="61823CCBEC7760CA7F9419D4488523C0" pageId="0" pageNumber="45">
<taxonomicName id="35B852C3765C613B305268A9E8881439" authority="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke, 2021" authorityName="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus" status="sp. nov.">Exocelina</taxonomicName>
<normalizedToken id="A0003DEEEA648A2842ECD0BDF59F7632" originalValue="“Foja”">&quot;Foja&quot;</normalizedToken>
#7282:
<bibRefCitation DOI="https://doi.org/10.1186/s12862-021-01764-2" author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" refId="B10" refString="Toussaint, EFA, White, LT, Shaverdo, H, Lam, A, Surbakti, S, Panjaitan, R, Sumoked, B, von Rintelen, T, Sagata, K, Balke, M, 2021. [in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology. https://doi.org/10.1186/s12862-021-01764-2" title="[in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology." url="https://doi.org/10.1186/s12862-021-01764-2" year="2021">Toussaint et al. 2021</bibRefCitation>
<bibRefCitation id="3017BAB5D3B47430CD5451CD1EF06977" DOI="https://doi.org/10.1186/s12862-021-01764-2" author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" refId="B10" refString="Toussaint, EFA, White, LT, Shaverdo, H, Lam, A, Surbakti, S, Panjaitan, R, Sumoked, B, von Rintelen, T, Sagata, K, Balke, M, 2021. [in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology. https://doi.org/10.1186/s12862-021-01764-2" title="[in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology." url="https://doi.org/10.1186/s12862-021-01764-2" year="2021">Toussaint et al. 2021</bibRefCitation>
: figs 3-6.
</paragraph>
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<typeStatus>Type</typeStatus>
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<typeStatus id="B2B84A1A2ADEC24D8A58319DE36CB616">Type</typeStatus>
locality.
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<collectingCountry name="Indonesia">Indonesia</collectingCountry>
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<materialsCitation id="62E21F98CBB46AD13BB3A93B9FF0C374" ID-GBIF-Occurrence="3062138312" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Foja Mts, N" country="Indonesia" elevation="115" latitude="-2.374889" location="Waaf Village" longLatPrecision="1" longitude="138.73886" municipality="Foja Mts" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<collectingCountry id="FAB21224BDF2F39828920A51F10F2C3A" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
<collectingRegion id="A1DE4546FDCBEEE349AC6EB5676C3DAB" country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
, Sarmi Regency,
<collectorName>
<collectingMunicipality>Foja Mts</collectingMunicipality>
<collectorName id="33923D747E24E68E3F72CBE4F209FEC5">
<collectingMunicipality id="BC14D656F98D0387A009C422E6EACFB4">Foja Mts</collectingMunicipality>
, N
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,
<geoCoordinate degrees="02" direction="south" minutes="22" orientation="latitude" precision="1" seconds="29.6" value="-2.374889">02°22'29.6&quot;S</geoCoordinate>
<geoCoordinate id="3119FD5B92E281EBD18CB09C270A4C8B" degrees="02" direction="south" minutes="22" orientation="latitude" precision="1" seconds="29.6" value="-2.374889">02°22'29.6&quot;S</geoCoordinate>
,
<geoCoordinate degrees="138" direction="east" minutes="44" orientation="longitude" precision="1" seconds="19.9" value="138.73886">138°44'19.9&quot;E</geoCoordinate>
<geoCoordinate id="83B5A5B497F8FD286D8B3E2E51D69DDD" degrees="138" direction="east" minutes="44" orientation="longitude" precision="1" seconds="19.9" value="138.73886">138°44'19.9&quot;E</geoCoordinate>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">115 m</quantity>
<elevation id="D5C5AA80674632BE56CCE037F3D4CEDF" metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">
<quantity id="68A1E8E40C93BC08D38E134B9E980B26" metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">115 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type material">
<paragraph pageId="0" pageNumber="45">
<typeStatus>Type</typeStatus>
<subSubSection id="733EFB42D5645568DA7955BABDCF5C15" pageId="0" pageNumber="45" type="type material">
<paragraph id="260159A28EAF886EC2A7CA5386A2324D" pageId="0" pageNumber="45">
<typeStatus id="99CCF41439021B822A24B45BF2D2E1BF">Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Holotype</typeStatus>
<paragraph id="7D399D12226AD63F31D1DA5524380170" pageId="0" pageNumber="45">
<emphasis id="8FEFD3BA7797BBB1B50F26932F01D906" bold="true" pageId="0" pageNumber="45">
<emphasis id="312CA5A35C4D0B1B37C1B51CF248FAE2" italics="true" pageId="0" pageNumber="45">
<typeStatus id="F89480964726C1F630D5533F8B5F4186">Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3062138310" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB" country="Indonesia" elevation="115" latitude="-2.374874" location="Foja Mountains" longLatPrecision="1" longitude="138.73886" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<materialsCitation id="18C9B7F9AAA76C0A859A46D4A0D00D30" ID-GBIF-Occurrence="3062138310" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB" country="Indonesia" elevation="115" latitude="-2.374874" location="Foja Mountains" longLatPrecision="1" longitude="138.73886" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
male &quot;
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
<collectingCountry id="75F3DDA81CCD78D9EC6E5503EFC01379" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
<collectingRegion id="FD7D6F2BAD3B0087914EF7E2F1B64ED8" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:367E9AA59B905821889868232A82AAEA:5E767422BE173026BDC5B2210F22D670" country="Indonesia" latitude="-2.374874" longLatPrecision="1" longitude="138.73886" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
<location id="5E767422BE173026BDC5B2210F22D670" LSID="urn:lsid:plazi:treatment:367E9AA59B905821889868232A82AAEA:5E767422BE173026BDC5B2210F22D670" country="Indonesia" latitude="-2.374874" longLatPrecision="1" longitude="138.73886" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
N foot, N Waaf vill,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">115m</elevation>
<quantity id="E738519444A64C5289C518A1462AD931" metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">
<elevation id="29EAEFE62036171F0897FCB8FDD375AD" metricMagnitude="2" metricUnit="m" metricValue="1.15" unit="m" value="115.0">115m</elevation>
</quantity>
,
<date value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
<date id="7EFD3F42815FC35296E5DBEE085EBA57" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate id="BF0562A420775FD070A2EE5EE9AA8C6F" value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
,
<geoCoordinate degrees="2.374874" direction="south" orientation="latitude" precision="1" value="-2.374874">-2.374874</geoCoordinate>
<geoCoordinate degrees="138.738855" direction="east" orientation="longitude" precision="1" value="138.73886">138.738855</geoCoordinate>
<geoCoordinate id="ABE320F26C858CCD7B8208A704CA54B5" degrees="2.374874" direction="south" orientation="latitude" precision="1" value="-2.374874">-2.374874</geoCoordinate>
<geoCoordinate id="68A26F0A8652899FFE32DA296555FB8A" degrees="138.738855" direction="east" orientation="longitude" precision="1" value="138.73886">138.738855</geoCoordinate>
, Sumoked (Pap060)&quot; (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
<collectionCode id="51A08DE57FD6DAEB7E8FEF8866802EC2" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138306" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="MZB, KSP" specimenCount="8" specimenCount-female="5" specimenCount-male="3" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Paratypes</typeStatus>
<paragraph id="550EAC41E09AE20149D107EC8468B240" pageId="0" pageNumber="45">
<materialsCitation id="8A281BCEEAC5238A8C17CCA6FE4A640D" ID-GBIF-Occurrence="3062138306" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="MZB, KSP" specimenCount="8" specimenCount-female="5" specimenCount-male="3" typeStatus="paratype">
<emphasis id="3FCB0E8B70EF7B5BBA881714E7C40242" bold="true" pageId="0" pageNumber="45">
<emphasis id="ACFC672ECD8BD4E070AD69BC556C5EC6" italics="true" pageId="0" pageNumber="45">
<typeStatus id="66559508B3DC1E080F5F7240BBA7C068">Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount type="male">3 males</specimenCount>
<specimenCount id="E05E5C789B6C265E174744C455768374" type="male">3 males</specimenCount>
,
<specimenCount type="female">5 females</specimenCount>
<specimenCount id="A56FF77881C6BE1F2510877966CDB34F" type="female">5 females</specimenCount>
with the same label as the
<typeStatus>holotype</typeStatus>
<typeStatus id="89175A9F1293888BBC7C483351E8D929">holotype</typeStatus>
(
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
<collectionCode id="A0B8B5CCEF7E4B0BC4C5F4D7C77BC566" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
,
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
<collectionCode id="5222234811978E75A29D4A7D2FB541CA" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3062138307" collectingDate="2016-06-04" collectingDateMax="2016-06-07" collectingDateMin="2016-06-04" collectionCode="KSP" country="Indonesia" elevation="150" latitude="-2.406142" location="Foja Mountains" longLatPrecision="1" longitude="138.74399" specimenCount="2" specimenCount-male="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">2 males</specimenCount>
<materialsCitation id="A460110E43A5A66C3B4FD98A5077F54C" ID-GBIF-Occurrence="3062138307" collectingDate="2016-06-04" collectingDateMax="2016-06-07" collectingDateMin="2016-06-04" collectionCode="KSP" country="Indonesia" elevation="150" latitude="-2.406142" location="Foja Mountains" longLatPrecision="1" longitude="138.74399" specimenCount="2" specimenCount-male="2" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="2EA4083FCDDE4FD01098DDA744585889" type="male">2 males</specimenCount>
&quot;
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
<collectingCountry id="8EBE78192B44F9F41404668C90A119CB" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
<collectingRegion id="1288947B0E32401CC590623A284C1FDD" country="Indonesia" name="Papua Barat">Papua</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:367E9AA59B905821889868232A82AAEA:3AE5B416A2D1ACC1802AC8BA55545365" country="Indonesia" latitude="-2.406142" longLatPrecision="1" longitude="138.74399" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
<location id="3AE5B416A2D1ACC1802AC8BA55545365" LSID="urn:lsid:plazi:treatment:367E9AA59B905821889868232A82AAEA:3AE5B416A2D1ACC1802AC8BA55545365" country="Indonesia" latitude="-2.406142" longLatPrecision="1" longitude="138.74399" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
N foot, N Waaf vill, pondok,
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150m</elevation>
<quantity id="DC63E727C310E8AC4ACCA8FAC78A70EE" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">
<elevation id="E73AD14124CFB16F5EEA7EEDEF53548D" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150m</elevation>
</quantity>
,
<date value="2016-06-04" valueMax="2016-06-07" valueMin="2016-06-04">
<collectingDate value="2016-06-04" valueMax="2016-06-07" valueMin="2016-06-04">4.-7.vi.2016</collectingDate>
<date id="2AE625ADE498633F37B1C1202AADFB19" value="2016-06-04" valueMax="2016-06-07" valueMin="2016-06-04">
<collectingDate id="8021A22E31CD3E1C3BA0256683D36138" value="2016-06-04" valueMax="2016-06-07" valueMin="2016-06-04">4.-7.vi.2016</collectingDate>
</date>
,
<geoCoordinate degrees="2.406142" direction="south" orientation="latitude" precision="1" value="-2.406142">-2.406142</geoCoordinate>
<geoCoordinate degrees="138.74399" direction="east" orientation="longitude" precision="1" value="138.74399">138.74399</geoCoordinate>
<geoCoordinate id="52981D04ECEEDB782FBD4854D8F63755" degrees="2.406142" direction="south" orientation="latitude" precision="1" value="-2.406142">-2.406142</geoCoordinate>
<geoCoordinate id="D7DCA6C5EEEFD31BE09A540260C596BA" degrees="138.74399" direction="east" orientation="longitude" precision="1" value="138.74399">138.74399</geoCoordinate>
, Sumoked (Pap061)&quot;, one male with an additional green text label
<normalizedToken originalValue="“7282”">&quot;7282&quot;</normalizedToken>
<normalizedToken id="CA282D01B9AE0CF89F49321B21314586" originalValue="“7282”">&quot;7282&quot;</normalizedToken>
(
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
<collectionCode id="9DDFE6F34CEA92774066DF16EEAA51EE" country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Body size and form</emphasis>
<subSubSection id="6A0F2F39EE7AB6ECD9F1CC382529387E" pageId="0" pageNumber="45" type="description">
<paragraph id="C7B1FB892C152502CB177810C2C2D853" pageId="0" pageNumber="45">Description.</paragraph>
<paragraph id="407BD343EB3AD512A9853C912C2B9C01" pageId="0" pageNumber="45">
<emphasis id="1FF6D2F485AC0D990A6F84383BC1FB26" bold="true" pageId="0" pageNumber="45">
<emphasis id="7DECA0C13B52567F0B73B47CED90DB88" italics="true" pageId="0" pageNumber="45">Body size and form</emphasis>
:
</emphasis>
Beetle small to medium-sized: TL-H
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.85" metricValueMin="3.45" unit="mm" value="3.65" valueMax="3.85" valueMin="3.45">3.45-3.85 mm</quantity>
<quantity id="26B05615A6CDC98DD55629C159493B26" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.85" metricValueMin="3.45" unit="mm" value="3.65" valueMax="3.85" valueMin="3.45">3.45-3.85 mm</quantity>
, TL
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" unit="mm" value="4.05" valueMax="4.3" valueMin="3.8">3.8-4.3 mm</quantity>
<quantity id="7039459A608D0F60DBDBF3E121C5DD81" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" unit="mm" value="4.05" valueMax="4.3" valueMin="3.8">3.8-4.3 mm</quantity>
, MW
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.975" metricValueMax="2.1" metricValueMin="1.85" unit="mm" value="1.975" valueMax="2.1" valueMin="1.85">1.85-2.1 mm</quantity>
<quantity id="74648C3F2A03596112E6C62CEE73C554" metricMagnitude="-3" metricUnit="m" metricValue="1.975" metricValueMax="2.1" metricValueMin="1.85" unit="mm" value="1.975" valueMax="2.1" valueMin="1.85">1.85-2.1 mm</quantity>
(
<typeStatus>holotype</typeStatus>
<typeStatus id="960E7FA857FF336781E481FA325A8E05">holotype</typeStatus>
: TL-H
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.6" unit="mm" value="3.6">3.6 mm</quantity>
<quantity id="C1A4106055BC410ED5CD3F5578AC9EF3" metricMagnitude="-3" metricUnit="m" metricValue="3.6" unit="mm" value="3.6">3.6 mm</quantity>
, TL
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.1" unit="mm" value="4.1">4.1 mm</quantity>
<quantity id="34A3FBCD2EEC294434BC616ED512C6E0" metricMagnitude="-3" metricUnit="m" metricValue="4.1" unit="mm" value="4.1">4.1 mm</quantity>
,
<collectionCode country="Netherlands" name="Museum Wasmann">MW</collectionCode>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.9" unit="mm" value="1.9">1.9 mm</quantity>
<collectionCode id="36A2410657CE767A935AA2A8EA7282D3" country="Netherlands" name="Museum Wasmann">MW</collectionCode>
<quantity id="8A775DA594B6BC4319F6A26423F6F6ED" metricMagnitude="-3" metricUnit="m" metricValue="1.9" unit="mm" value="1.9">1.9 mm</quantity>
), with oblong-oval habitus (Fig.
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">3</figureCitation>
<figureCitation id="090BE2314582AA1553325BFB911D32A6" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">3</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Colouration</emphasis>
<paragraph id="B92EC62E9E190A37AF44706B780CBA56" pageId="0" pageNumber="45">
<emphasis id="EDA7A622F3F3CFA786DDA3D953533405" bold="true" pageId="0" pageNumber="45">
<emphasis id="F415746A18BE0EC515E67FF82709CF6E" italics="true" pageId="0" pageNumber="45">Colouration</emphasis>
:
</emphasis>
Dorsally dark brown to piceous, usually with paler, reddish brown, head and pronotum (Fig.
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">3</figureCitation>
<figureCitation id="34A0896C0E35DA3068804596FB094AE8" captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">3</figureCitation>
). Head reddish brown, reddish anteriorly, dark brown around eyes; pronotum reddish brown to brown, with darker, to dark brown, disc; elytra dark brown, sometimes with reddish brown sutural lines; head appendages yellow, legs yellowish red to reddish brown. One specimen (from locality Pap061) piceous, with dark brown pronotal lateral sides. Most specimens from locality Pap060 teneral, therefore, paler.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Surface sculpture</emphasis>
<paragraph id="48A5641ABB2B00FA5B0D857E34F24CA6" pageId="0" pageNumber="45">
<emphasis id="11241DB13B132374774A0749D645457C" bold="true" pageId="0" pageNumber="45">
<emphasis id="7A8BA3F2ACD19EC6631CD9F3AF9DE709" italics="true" pageId="0" pageNumber="45">Surface sculpture</emphasis>
:
</emphasis>
Shiny dorsally, with inconspicuous, almost invisible elytral punctation and weakly impressed microreticulation. Head with relatively fine and sparse punctation (spaces between punctures 1-4 times size of punctures); diameter of punctures smaller than diameter of cells of microreticulation. Pronotum with finer, sparser, and more evenly distributed punctation than on head, often inconspicuous. Elytra with very sparse and fine punctation, almost invisible. Elytra with weakly impressed microreticulation; pronotum and especially head with stronger microreticulation. Metaventrite and metacoxa distinctly but weakly microreticulate, metacoxal plates with longitudinal strioles and very weak transverse wrinkles. Abdominal ventrites with weak microreticulation, strioles, and punctation visible only on two last abdominal ventrites.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Structures</emphasis>
<paragraph id="FBA64834600DDCD6BA0D5932F84CD862" pageId="0" pageNumber="45">
<emphasis id="05CAD4F842580B22969989E99FDBDD8C" bold="true" pageId="0" pageNumber="45">
<emphasis id="8C9326F577D418ADC4D66F301D271771" italics="true" pageId="0" pageNumber="45">Structures</emphasis>
:
</emphasis>
Pronotum without lateral bead, in some specimens (especially characteristic for females) with bead traces or even with narrow bead on lateral sides of pronotum. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 broadly rounded.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Male</emphasis>
<paragraph id="83B839B61D31200B9415E0C9752F5B19" pageId="0" pageNumber="45">
<emphasis id="B1D4B3414B33102D8179FAFEB1C3E04C" bold="true" pageId="0" pageNumber="45">
<emphasis id="7F373D67960F7B9381AC774306A2B0F1" italics="true" pageId="0" pageNumber="45">Male</emphasis>
:
</emphasis>
Antenna simple. Pro- and mesotarsomeres 1-3 not dilated, narrow. Protarsomere 4 cylindrical, narrow, with medium-sized, thick, distinctly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of ten and posterior row of five short setae (Fig.
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">7A</figureCitation>
<figureCitation id="780466B46D56F2E084301C1671C153C9" captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">7A</figureCitation>
). Median lobe with distinctly discontinuous outline; in lateral view, almost straight, with apex narrow, curved downwards, and strongly protruding at tip forming a long thin prolongation; in ventral view, with weak submedian constriction, distal part narrower than proximal one, apex slightly and evenly concave, with distinct protruding tip (Fig.
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">7B, C</figureCitation>
<figureCitation id="11395814F33F350EE6F6F9EFB9C3265B" captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">7B, C</figureCitation>
). Paramere with strong notch on dorsal side and subdistal part subquadrate, large and broad; subdistal setae dense, rather short, flattened; proximal setae hair-like, numerous, dense, and long (Fig.
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">7D</figureCitation>
<figureCitation id="E07667B21C79CCB9F52E5A4F4BE0BB2F" captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">7D</figureCitation>
). Abdominal ventrite 6 broadly rounded, with 13-16 lateral striae on each side.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Female</emphasis>
<paragraph id="DE8474A22BEA674138881D25A0C35AAE" pageId="0" pageNumber="45">
<emphasis id="7D000250E7F64FB45848C02E7E137646" bold="true" pageId="0" pageNumber="45">
<emphasis id="4FB83DDA3936FCBB7DF77303A0BD76A8" italics="true" pageId="0" pageNumber="45">Female</emphasis>
:
</emphasis>
Pro- and mesotarsi not modified. Abdominal ventrite 6 without lateral striae. Bead traces or even with narrow bead on lateral margins pronotum present in majority of females.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="affinities">
<paragraph pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="CA7E72B73CBC8E12BB8F69E91A32CF3B" pageId="0" pageNumber="45" type="affinities">
<paragraph id="1C9DF764364E6B2EBB5A18B0F11CAA20" pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph id="832445F3392AF22215FF1E216458D15C" pageId="0" pageNumber="45">
The new species evidently belongs to the
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekari">
<emphasis italics="true" pageId="0" pageNumber="45">E. ekari</emphasis>
<taxonomicName id="6193D63D694600A1DF28C54DBBDF7FA1" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekari">
<emphasis id="83B5C7B0D04D4AE907596FCF8526B309" italics="true" pageId="0" pageNumber="45">E. ekari</emphasis>
</taxonomicName>
group due to the discontinuous outline of its median lobe. The species is similar to
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. pinocchio" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="pinocchio">
<emphasis italics="true" pageId="0" pageNumber="45">E. pinocchio</emphasis>
<taxonomicName id="2C27AA94C531745BC3E702FEA12FD0EE" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. pinocchio" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="pinocchio">
<emphasis id="C762EE7E7FDAA5D14657A55A98E896D6" italics="true" pageId="0" pageNumber="45">E. pinocchio</emphasis>
</taxonomicName>
Shaverdo &amp; Balke,
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.11556" unit="in" value="2014.0">2014 in</quantity>
<quantity id="62D654AB0056969EB7BCD839F60A3EF9" metricMagnitude="1" metricUnit="m" metricValue="5.11556" unit="in" value="2014.0">2014 in</quantity>
general appearance and shape of median lobe, but differs from it in more straight apical prolongation of the median lobe and in subquadrate, large and broad subdistal part of the paramere (distinctly more elongate in
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. pinocchio" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="pinocchio">
<emphasis italics="true" pageId="0" pageNumber="45">E. pinocchio</emphasis>
<taxonomicName id="9C885D1B43C534789D48F3D07A054264" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. pinocchio" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="pinocchio">
<emphasis id="196EC2DC3D74888BFD1661EE4D17E1CD" italics="true" pageId="0" pageNumber="45">E. pinocchio</emphasis>
</taxonomicName>
) and its setation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138342" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" location="Foja Mountains" municipality="This" specimenCount="1" stateProvince="Papua Barat">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
<subSubSection id="3CD865752E166A86B7986223032F1D2E" pageId="0" pageNumber="45" type="distribution">
<paragraph id="58F18BEDEBE2442DE5884F2D0D695C54" pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph id="99787CBA7C5F3DE660782A6F2ECD2552" pageId="0" pageNumber="45">
<materialsCitation id="E97F2C2BA20BCA12E41F7EDA8D350941" ID-GBIF-Occurrence="3062138342" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" location="Foja Mountains" municipality="This" specimenCount="1" stateProvince="Papua Barat">
<collectingCountry id="A790223498254234C0BE625EF7E97582" name="Indonesia">Indonesia</collectingCountry>
:
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
<collectingRegion id="482865E5B518D2B3453E63411326082E" country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
.
<collectingMunicipality>This</collectingMunicipality>
<collectingMunicipality id="B5936A338BA32782B20B570D469A0553">This</collectingMunicipality>
species is known only from
<location LSID="urn:lsid:plazi:treatment:367E9AA59B905821889868232A82AAEA:56F3846BF2FCDB31AD0F0129D8DE5993" country="Indonesia" municipality="This" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
<location id="56F3846BF2FCDB31AD0F0129D8DE5993" LSID="urn:lsid:plazi:treatment:367E9AA59B905821889868232A82AAEA:56F3846BF2FCDB31AD0F0129D8DE5993" country="Indonesia" municipality="This" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, from and near the
<typeStatus>type</typeStatus>
<typeStatus id="8651D386D8437C7ECED1172B15FEFA06">type</typeStatus>
locality (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">15</figureCitation>
<figureCitation id="0F380235BD5B227BC5E8CDFC51DEB33B" captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">15</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="habitat">
<paragraph pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="45">The specimens were collected from shaded waterholes on a riverbank.</paragraph>
<subSubSection id="E9C27C78E03C0C02F7AFF2E98757F5D4" pageId="0" pageNumber="45" type="habitat">
<paragraph id="F49DFE857459006955D94C63FA04737B" pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph id="CBF4DB39817C2042929D371530A63D3A" pageId="0" pageNumber="45">The specimens were collected from shaded waterholes on a riverbank.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="771586F066156FD221DC066FA9D1427B" pageId="0" pageNumber="45" type="etymology">
<paragraph id="CF46A5FE3F0B878E41EEE092D3A3AB4E" pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph id="99D63B6DB954268DDF92A8EBDFA52158" pageId="0" pageNumber="45">
The species is named to honour Dr Ignacio Ribera
<normalizedToken originalValue="Galán">Galan</normalizedToken>
<normalizedToken id="A02E9DC9F6E0E1546B78A585EC4B918E" originalValue="Galán">Galan</normalizedToken>
, a leading water beetle specialist and our dear colleague who passed away on
<date value="2020-04-15">15 April 2020</date>
<date id="BD21DA459E9276C7D47A5A8EFF997D5E" value="2020-04-15">15 April 2020</date>
. The name is a noun in the genitive case.
</paragraph>
</subSubSection>