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<treatment ID-GBIF-Taxon="152053528" LSID="urn:lsid:plazi:treatment:340399A97289CFD1C96059391D8DF2EE" httpUri="http://treatment.plazi.org/id/340399A97289CFD1C96059391D8DF2EE" lastPageId="11" lastPageNumber="64" pageId="9" pageNumber="62">
<subSubSection pageId="9" pageNumber="62" type="nomenclature">
<paragraph pageId="9" pageNumber="62">
<treatment id="340399A97289CFD1C96059391D8DF2EE" ID-GBIF-Taxon="152053528" LSID="urn:lsid:plazi:treatment:340399A97289CFD1C96059391D8DF2EE" httpUri="http://treatment.plazi.org/id/340399A97289CFD1C96059391D8DF2EE" lastPageId="11" lastPageNumber="64" pageId="9" pageNumber="62" scope_family="Tenebrionidae" scope_order="Coleoptera">
<subSubSection id="2586006EE0D73CF6DD448E898644A2D9" pageId="9" pageNumber="62" type="nomenclature">
<paragraph id="7A0F7285CDADD3F40E772633A6D95A99" pageId="9" pageNumber="62">
(5) Subgenus
<taxonomicName authority="Gay &amp; Solier in Solier, 1840" authorityName="Gay &amp; Solier in Solier" authorityYear="1840" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis (Filotarsus)" order="Coleoptera" pageId="9" pageNumber="62" phylum="Arthropoda" rank="subGenus" subGenus="Filotarsus">Praocis (Filotarsus) Gay &amp; Solier in Solier, 1840</taxonomicName>
<taxonomicName id="FEBDFF352A1B6C4598C968AE590372AB" authority="Gay &amp; Solier in Solier, 1840" authorityName="Gay &amp; Solier in Solier" authorityYear="1840" genus="Praocis" lsidName="Praocis (Filotarsus)" pageId="9" pageNumber="62" rank="subGenus" subGenus="Filotarsus">Praocis (Filotarsus) Gay &amp; Solier in Solier, 1840</taxonomicName>
Figs 10, 23-24
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="62" type="type species">
<paragraph pageId="9" pageNumber="62">Type species.</paragraph>
<paragraph pageId="9" pageNumber="62">
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Filotarsus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Filotarsus tenuicornis" order="Coleoptera" pageId="9" pageNumber="62" phylum="Arthropoda" rank="species" species="tenuicornis">Filotarsus tenuicornis</taxonomicName>
<subSubSection id="B1269C1D884A357DB3152E8B23D1F8B8" pageId="9" pageNumber="62" type="type species">
<paragraph id="D1576D7B62846B6A2DAF2FC3CFD36C30" pageId="9" pageNumber="62">Type species.</paragraph>
<paragraph id="F9DA7CA92FBB68A90C736FF85A4D021B" pageId="9" pageNumber="62">
<taxonomicName id="8FDCFA315AC7748818F348A9F28F2F83" genus="Filotarsus" lsidName="Filotarsus tenuicornis" pageId="9" pageNumber="62" rank="species" species="tenuicornis">Filotarsus tenuicornis</taxonomicName>
Gay &amp; Solier in Solier, 1840, monotypy and original designation by
<bibRefCitation author="Solier, AJJ" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="23" pageNumber="76" pagination="207 - 370" title="Essai sur les Collapterides. 11 e Tribu. Praocites." volume="9" year="1840">Solier (1840</bibRefCitation>
<bibRefCitation id="A2A382C083C8C5B1C2975273726774C0" author="Solier, AJJ" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="23" pageNumber="76" pagination="207 - 370" title="Essai sur les Collapterides. 11 e Tribu. Praocites." volume="9" year="1840">Solier (1840</bibRefCitation>
: 241).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="62" type="diagnosis">
<paragraph pageId="9" pageNumber="62">Diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="62">Clypeus with anterior margin extending beyond to lateral expansion of frons or at same level as lateral expansion of frons, width of anterior margin not exceeding half the interocular width or width of anterior margin same as interocular width, clypeal suture as a vertical groove, not covered by frons, clypeus and frons at same level or clypeal suture as a horizontal groove not covered by frons, clypeus lower than frons; antennomere 9 longer than antennomere 10, antennomere 11 longer than antennomere 10; apical tomentose sensory patches on antennomere 10 in a dorsally continuous semicircle, on antennomere 11 on distal half; prosternum with a narrow edge on anterior margin; lateral margin of elytron not defined; ventral surface of profemora without a row of setae on anterior edge, protibiae explanate.</paragraph>
<subSubSection id="EA9A5FF83DED0D34B16D8130F13EEE6F" pageId="9" pageNumber="62" type="diagnosis">
<paragraph id="ED1DDF12092D20ADE026BC649DB15400" pageId="9" pageNumber="62">Diagnosis.</paragraph>
<paragraph id="D9EADA5B6CF3B0737246B1036290F0D5" pageId="9" pageNumber="62">Clypeus with anterior margin extending beyond to lateral expansion of frons or at same level as lateral expansion of frons, width of anterior margin not exceeding half the interocular width or width of anterior margin same as interocular width, clypeal suture as a vertical groove, not covered by frons, clypeus and frons at same level or clypeal suture as a horizontal groove not covered by frons, clypeus lower than frons; antennomere 9 longer than antennomere 10, antennomere 11 longer than antennomere 10; apical tomentose sensory patches on antennomere 10 in a dorsally continuous semicircle, on antennomere 11 on distal half; prosternum with a narrow edge on anterior margin; lateral margin of elytron not defined; ventral surface of profemora without a row of setae on anterior edge, protibiae explanate.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="62" type="distribution">
<paragraph pageId="9" pageNumber="62">Distribution.</paragraph>
<paragraph pageId="9" pageNumber="62">
<subSubSection id="F752D27F58C336EABB8ED9841C6064F5" pageId="9" pageNumber="62" type="distribution">
<paragraph id="B21534A72FC98DDB2D91F94F64473A54" pageId="9" pageNumber="62">Distribution.</paragraph>
<paragraph id="A5FDAA73246BD6E82088887735709BAF" pageId="9" pageNumber="62">
Species of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis (Filotarsus)" order="Coleoptera" pageId="9" pageNumber="62" phylum="Arthropoda" rank="subGenus" subGenus="Filotarsus">Praocis (Filotarsus)</taxonomicName>
<taxonomicName id="341E90BFC8BCA411AC53CCD4F1E5B9FC" genus="Praocis" lsidName="Praocis (Filotarsus)" pageId="9" pageNumber="62" rank="subGenus" subGenus="Filotarsus">Praocis (Filotarsus)</taxonomicName>
inhabit central and northern Chile, western and northern Argentina, estern Bolivia and southern Peru. They occur from 12°South (Cuzco, Peru) to 39°South (
<normalizedToken originalValue="Neuquén">Neuquen</normalizedToken>
<normalizedToken id="63005B8E5C13999ECE3315CF968C2620" originalValue="Neuquén">Neuquen</normalizedToken>
, Argentina) in the biogeographic provinces of Puna, Atacama, Coquimbo, Santiago, Prepuna, Monte, and Central Patagonia (
<bibRefCitation author="Morrone, JJ" journalOrPublisher="Annual Review of Entomology" pageId="22" pageNumber="75" pagination="467 - 494" title="Biogeographic areas and transition zones of Latin America and the Caribbean islands based on panbiogeographic and cladistics analyses of the entomofauna." url="10.1146/annurev.ento.50.071803.130447" volume="51" year="2006">Morrone 2006</bibRefCitation>
<bibRefCitation id="7BD9F240F8F78D4F06AB9D648BD4AD6F" author="Morrone, JJ" journalOrPublisher="Annual Review of Entomology" pageId="22" pageNumber="75" pagination="467 - 494" title="Biogeographic areas and transition zones of Latin America and the Caribbean islands based on panbiogeographic and cladistics analyses of the entomofauna." url="10.1146/annurev.ento.50.071803.130447" volume="51" year="2006">Morrone 2006</bibRefCitation>
) (Fig. 23).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="63" type="diversity">
<paragraph pageId="10" pageNumber="63">
<pageBreakToken pageId="10" pageNumber="63" start="start">Diversity</pageBreakToken>
<subSubSection id="CB8AA2F884DFA0421B910D07D856DF0E" pageId="10" pageNumber="63" type="diversity">
<paragraph id="DDEA59773D1756EF54EC798F509C47E2" pageId="10" pageNumber="63">
<pageBreakToken id="792DE2F64975029CC658BC24C69912B1" pageId="10" pageNumber="63" start="start">Diversity</pageBreakToken>
.
</paragraph>
<paragraph pageId="10" pageNumber="63">
<paragraph id="44C6CF44ED3ED7B4322F77F7D21F6D59" pageId="10" pageNumber="63">
This subgenus contains 14 species (
<bibRefCitation author="Kulzer, H" journalOrPublisher="Entomologische Arbeiten aus dem Museum George Frey" pageId="22" pageNumber="75" pagination="1 - 105" title="Monographie der suedamerikanischen Tribus Praocini (Col.) (16 Beitrag zur Kenntnis der Tenebrioniden)." volume="9" year="1958">Kulzer 1958</bibRefCitation>
<bibRefCitation id="EAF131831E5E07CA9CC5C44FFB6A96B0" author="Kulzer, H" journalOrPublisher="Entomologische Arbeiten aus dem Museum George Frey" pageId="22" pageNumber="75" pagination="1 - 105" title="Monographie der suedamerikanischen Tribus Praocini (Col.) (16 Beitrag zur Kenntnis der Tenebrioniden)." volume="9" year="1958">Kulzer 1958</bibRefCitation>
;
<bibRefCitation author="Flores, GE" journalOrPublisher="Zootaxa" pageId="21" pageNumber="74" pagination="21 - 33" title="Revision of some types of the South American tribes Nycteliini, Praocini, and Scotobiini (Coleoptera: Tenebrionidae), with new synonymies." volume="1985" year="2009">Flores 2009</bibRefCitation>
<bibRefCitation id="DADF52C3949096A562D0AEEAE8287606" author="Flores, GE" journalOrPublisher="Zootaxa" pageId="21" pageNumber="74" pagination="21 - 33" title="Revision of some types of the South American tribes Nycteliini, Praocini, and Scotobiini (Coleoptera: Tenebrionidae), with new synonymies." volume="1985" year="2009">Flores 2009</bibRefCitation>
) plus 6 species to be described, 20 species in total, with a 43 percent increase in the number of species (Fig. 33).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="63" type="habitat">
<paragraph pageId="10" pageNumber="63">Habitat.</paragraph>
<paragraph pageId="10" pageNumber="63">
<subSubSection id="40A47368357DD76B081353AB0FEC4931" pageId="10" pageNumber="63" type="habitat">
<paragraph id="07CEF86302CBBED9332B86A4BD1D3633" pageId="10" pageNumber="63">Habitat.</paragraph>
<paragraph id="FDBCA149EF2577908BA24D725F53CD71" pageId="10" pageNumber="63">
Species of
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis (Filotarsus)" order="Coleoptera" pageId="10" pageNumber="63" phylum="Arthropoda" rank="subGenus" subGenus="Filotarsus">Praocis (Filotarsus)</taxonomicName>
<taxonomicName id="689CA4363F30E5776F26A4E8742CC97F" genus="Praocis" lsidName="Praocis (Filotarsus)" pageId="10" pageNumber="63" rank="subGenus" subGenus="Filotarsus">Praocis (Filotarsus)</taxonomicName>
have nocturnal habits, remaining during the day under stones or plants. In central Chile they can be observed in gullies and Coastal and Andean mountain ranges from 400 m to an altitude of 2500 m. In Argentina, Bolivia, northern Chile and Peru they occur from 1600 m in high altitudinal valleys associated with the Andes mountain range to an altitude of 5200 m in the high Puna plateau, in clayey, poorly permeable soils (
<bibRefCitation author="Ferru, MA" journalOrPublisher="Boletin del Museo Nacional de Historia Natural, Chile" pageId="21" pageNumber="74" pagination="9 - 61" title="Lista de Coleopteros (Insecta: Coleoptera) de las Regiones de Arica y Parinacota y de Tarapaca, Chile." volume="60" year="2011">
<normalizedToken originalValue="Ferrú">Ferru</normalizedToken>
<bibRefCitation id="A827CC815B7F0533B30250138B8F997F" author="Ferru, MA" journalOrPublisher="Boletin del Museo Nacional de Historia Natural, Chile" pageId="21" pageNumber="74" pagination="9 - 61" title="Lista de Coleopteros (Insecta: Coleoptera) de las Regiones de Arica y Parinacota y de Tarapaca, Chile." volume="60" year="2011">
<normalizedToken id="435C78339DD364D18F89AFE2D6317011" originalValue="Ferrú">Ferru</normalizedToken>
and Elgueta 2011
</bibRefCitation>
; collection data FMNH, IADIZA, LEULS, and pers. obs.) (Fig. 24).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="64" type="species included">
<paragraph pageId="11" pageNumber="64">
<pageBreakToken pageId="11" pageNumber="64" start="start">Species</pageBreakToken>
<subSubSection id="B50DF951386E54ACE6BEAD952C708315" pageId="11" pageNumber="64" type="species included">
<paragraph id="941298F7B8FAB1B0CCFED04A9EF568F0" pageId="11" pageNumber="64">
<pageBreakToken id="11E9FE25B18C5BA4FF8D5849E12282E6" pageId="11" pageNumber="64" start="start">Species</pageBreakToken>
included.
</paragraph>
<paragraph pageId="11" pageNumber="64">
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis tenuicornis" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="tenuicornis">Praocis tenuicornis</taxonomicName>
<paragraph id="5CB3ABFA6FB00545F3A2C9384AE994A2" pageId="11" pageNumber="64">
<taxonomicName id="5046F5D9EBD12388D3565AF3A6BC6FE8" genus="Praocis" lsidName="Praocis tenuicornis" pageId="11" pageNumber="64" rank="species" species="tenuicornis">Praocis tenuicornis</taxonomicName>
Gay &amp; Solier in Solier, 1840;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis castanea" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="castanea">Praocis castanea</taxonomicName>
<taxonomicName id="8A13901F09D2E683E14C8182C387CF6E" genus="Praocis" lsidName="Praocis castanea" pageId="11" pageNumber="64" rank="species" species="castanea">Praocis castanea</taxonomicName>
Germain, 1855;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis rufilabris" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="rufilabris">Praocis rufilabris</taxonomicName>
<taxonomicName id="99D3EEA38DB289745B92A970C528D749" genus="Praocis" lsidName="Praocis rufilabris" pageId="11" pageNumber="64" rank="species" species="rufilabris">Praocis rufilabris</taxonomicName>
Gay &amp; Solier in Solier, 1840;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis uretai" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="uretai">Praocis uretai</taxonomicName>
<taxonomicName id="E3DA8E14A911E6D112B0E566B851C7E8" genus="Praocis" lsidName="Praocis uretai" pageId="11" pageNumber="64" rank="species" species="uretai">Praocis uretai</taxonomicName>
Kulzer, 1958 (=
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis freyi" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="freyi">Praocis freyi</taxonomicName>
<taxonomicName id="0A23852146B97D7529F28A272EE8B0AE" genus="Praocis" lsidName="Praocis freyi" pageId="11" pageNumber="64" rank="species" species="freyi">Praocis freyi</taxonomicName>
Marcuzzi, 1977, synonymy by
<bibRefCitation author="Flores, GE" journalOrPublisher="Zootaxa" pageId="21" pageNumber="74" pagination="21 - 33" title="Revision of some types of the South American tribes Nycteliini, Praocini, and Scotobiini (Coleoptera: Tenebrionidae), with new synonymies." volume="1985" year="2009">Flores 2009</bibRefCitation>
<bibRefCitation id="F115FA7F790A53DEB5E2F5D9E9D42F14" author="Flores, GE" journalOrPublisher="Zootaxa" pageId="21" pageNumber="74" pagination="21 - 33" title="Revision of some types of the South American tribes Nycteliini, Praocini, and Scotobiini (Coleoptera: Tenebrionidae), with new synonymies." volume="1985" year="2009">Flores 2009</bibRefCitation>
);
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis reedi" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="reedi">Praocis reedi</taxonomicName>
<taxonomicName id="B4F25CB8B06E667E2DF3FFA1D540876A" genus="Praocis" lsidName="Praocis reedi" pageId="11" pageNumber="64" rank="species" species="reedi">Praocis reedi</taxonomicName>
Kulzer, 1958;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis oblonga" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="oblonga">Praocis oblonga</taxonomicName>
<taxonomicName id="250578D89EAE8E22E666CC9BDC54A424" genus="Praocis" lsidName="Praocis oblonga" pageId="11" pageNumber="64" rank="species" species="oblonga">Praocis oblonga</taxonomicName>
Solier, 1851;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis peltata" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="peltata">Praocis peltata</taxonomicName>
<taxonomicName id="9976CCE7EC470B69075F0720595FC8BC" genus="Praocis" lsidName="Praocis peltata" pageId="11" pageNumber="64" rank="species" species="peltata">Praocis peltata</taxonomicName>
Erichson, 1834;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis forsteri" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="forsteri">Praocis forsteri</taxonomicName>
<taxonomicName id="38E5B33F24AD2454B632B55B2778052E" genus="Praocis" lsidName="Praocis forsteri" pageId="11" pageNumber="64" rank="species" species="forsteri">Praocis forsteri</taxonomicName>
Kulzer, 1958;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis obesa" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="obesa">Praocis obesa</taxonomicName>
<taxonomicName id="02A343A87E8880E865DD2C3F9A8DB703" genus="Praocis" lsidName="Praocis obesa" pageId="11" pageNumber="64" rank="species" species="obesa">Praocis obesa</taxonomicName>
Kulzer, 1958;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis titschacki" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="titschacki">Praocis titschacki</taxonomicName>
<taxonomicName id="32B6075A64DE33E383BFDBC6E2F1A4EC" genus="Praocis" lsidName="Praocis titschacki" pageId="11" pageNumber="64" rank="species" species="titschacki">Praocis titschacki</taxonomicName>
Kulzer, 1958;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis brevicornis" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="brevicornis">Praocis brevicornis</taxonomicName>
<taxonomicName id="D479DCC3E6379C86F579F189A15081FD" genus="Praocis" lsidName="Praocis brevicornis" pageId="11" pageNumber="64" rank="species" species="brevicornis">Praocis brevicornis</taxonomicName>
Kulzer, 1958;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis weyrauchi" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="weyrauchi">Praocis weyrauchi</taxonomicName>
<taxonomicName id="9908A82D6EAE8E741D0E0EA9B025E56F" genus="Praocis" lsidName="Praocis weyrauchi" pageId="11" pageNumber="64" rank="species" species="weyrauchi">Praocis weyrauchi</taxonomicName>
Kulzer, 1958;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis peruana" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="peruana">Praocis peruana</taxonomicName>
<taxonomicName id="464BAE2F72006CA1582D8E1267980400" genus="Praocis" lsidName="Praocis peruana" pageId="11" pageNumber="64" rank="species" species="peruana">Praocis peruana</taxonomicName>
Fairmaire, 1902;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Praocis grossa" order="Coleoptera" pageId="11" pageNumber="64" phylum="Arthropoda" rank="species" species="grossa">Praocis grossa</taxonomicName>
<taxonomicName id="53C5B4F6EA84245A2FB51807E0DAB7DF" genus="Praocis" lsidName="Praocis grossa" pageId="11" pageNumber="64" rank="species" species="grossa">Praocis grossa</taxonomicName>
Kulzer, 1958.
</paragraph>
</subSubSection>

View file

@ -1,183 +1,183 @@
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4.2.5.
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<taxonomicNameLabel pageId="0" pageNumber="57">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the genus Typoderus. Map designed using the online SimpleMappr tool (Shorthouse 2010)." figureDoi="10.3897/asp.79.e66021.figure1" httpUri="https://binary.pensoft.net/fig/533979" pageId="0" pageNumber="57">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neighbour Joining clustering of 20 DNA barcodes of Cameroonian Typoderus (analysis A 1). Terminal names consist of BOLD sample ID (its last four digits correspond to specimen number), litter sifting sample number (Table 1), length of the DNA barcode (number of ambiguous nucleotides is in square brackets), BIN three-letter four-digit number and GenBank accession number. Names next to clusters are those of the new species described in section 4.2. HT denote holotypes." figureDoi="10.3897/asp.79.e66021.figure2" httpUri="https://binary.pensoft.net/fig/533980" pageId="0" pageNumber="57">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood inference phylogram of Typoderus weevils rooted on Lupangus (analysis A 3, root is not shown). Cameroonian species form a clade. Digits at internodes are bootstrap values of 50 % and above. Non-Cameroonian (= Tanzanian) terminals are collapsed in species. Terminal names consist of specimen number (Table 3) and sifting sample number (Table 1). Habitus images are to scale; their numbers indicate exact Cameroonian specimens imaged. Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure3" httpUri="https://binary.pensoft.net/fig/533981" pageId="0" pageNumber="57">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Maximum clade credibility Cameroonian Typoderus tree with median ages derived from the BEAST analysis A 4. The 95 % credibility interval for each node is given with a horizontal bar. Digits at nodes are million years before present. Terminal names consist of specimen number (Table 3), GenBank accession number and sifting sample number (Table 1). Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure4" httpUri="https://binary.pensoft.net/fig/533982" pageId="0" pageNumber="57">, 4</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male holotype of T. iphitus sp. nov., habitus (A: dorsal, B: left lateral, C: ventral, D: left fronto-lateral), aedeagus (E: dorsal, F: ventral, G: right lateral) and H: sternite 9." figureDoi="10.3897/asp.79.e66021.figure8" httpUri="https://binary.pensoft.net/fig/533986" pageId="0" pageNumber="57">, 8A-H.</figureCitation>
<taxonomicName id="AF9C2BF1360C17DB42CA5E4CAE6DD765" LSID="http://zoobank.org/83775EEA-6CA3-47FB-99BB-0B6EEC752B35" authority="Grebennikov, 2021" authorityName="Grebennikov" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Typoderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Typoderus iphitus" order="Coleoptera" pageId="0" pageNumber="57" phylum="Arthropoda" rank="species" species="iphitus" status="sp. nov.">Typoderus iphitus</taxonomicName>
<taxonomicNameLabel id="0FE4C25EC55575BB9E3CB8F35B2D3313" pageId="0" pageNumber="57">sp. nov.</taxonomicNameLabel>
<figureCitation id="C7A78688C9EFEC29B22A0C5280C13B95" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the genus Typoderus. Map designed using the online SimpleMappr tool (Shorthouse 2010)." figureDoi="10.3897/asp.79.e66021.figure1" httpUri="https://binary.pensoft.net/fig/533979" pageId="0" pageNumber="57">Figs 1</figureCitation>
<figureCitation id="49618D2942A8F20E207EF8A3E25DFE90" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neighbour Joining clustering of 20 DNA barcodes of Cameroonian Typoderus (analysis A 1). Terminal names consist of BOLD sample ID (its last four digits correspond to specimen number), litter sifting sample number (Table 1), length of the DNA barcode (number of ambiguous nucleotides is in square brackets), BIN three-letter four-digit number and GenBank accession number. Names next to clusters are those of the new species described in section 4.2. HT denote holotypes." figureDoi="10.3897/asp.79.e66021.figure2" httpUri="https://binary.pensoft.net/fig/533980" pageId="0" pageNumber="57">, 2</figureCitation>
<figureCitation id="CFA507F458B6282A52CF5B1A866850FC" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood inference phylogram of Typoderus weevils rooted on Lupangus (analysis A 3, root is not shown). Cameroonian species form a clade. Digits at internodes are bootstrap values of 50 % and above. Non-Cameroonian (= Tanzanian) terminals are collapsed in species. Terminal names consist of specimen number (Table 3) and sifting sample number (Table 1). Habitus images are to scale; their numbers indicate exact Cameroonian specimens imaged. Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure3" httpUri="https://binary.pensoft.net/fig/533981" pageId="0" pageNumber="57">, 3</figureCitation>
<figureCitation id="523AD02DDB5DB7A6C1C44534B40578C8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Maximum clade credibility Cameroonian Typoderus tree with median ages derived from the BEAST analysis A 4. The 95 % credibility interval for each node is given with a horizontal bar. Digits at nodes are million years before present. Terminal names consist of specimen number (Table 3), GenBank accession number and sifting sample number (Table 1). Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure4" httpUri="https://binary.pensoft.net/fig/533982" pageId="0" pageNumber="57">, 4</figureCitation>
<figureCitation id="F4CE6DBDF9548CBA5AF11A073FC88B29" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male holotype of T. iphitus sp. nov., habitus (A: dorsal, B: left lateral, C: ventral, D: left fronto-lateral), aedeagus (E: dorsal, F: ventral, G: right lateral) and H: sternite 9." figureDoi="10.3897/asp.79.e66021.figure8" httpUri="https://binary.pensoft.net/fig/533986" pageId="0" pageNumber="57">, 8A-H.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="description">
<paragraph pageId="0" pageNumber="57">Species diagnosis and description.</paragraph>
<paragraph pageId="0" pageNumber="57">
<subSubSection id="45C05CF6A5E486CBBC7729AC40BC82F8" pageId="0" pageNumber="57" type="description">
<paragraph id="D73AF07046250C55EB543E65F599D0AF" pageId="0" pageNumber="57">Species diagnosis and description.</paragraph>
<paragraph id="FC239ACCB3E529381C92A163E775219A" pageId="0" pageNumber="57">
Holotype male (Figs
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male holotype of T. iphitus sp. nov., habitus (A: dorsal, B: left lateral, C: ventral, D: left fronto-lateral), aedeagus (E: dorsal, F: ventral, G: right lateral) and H: sternite 9." figureDoi="10.3897/asp.79.e66021.figure8" httpUri="https://binary.pensoft.net/fig/533986" pageId="0" pageNumber="57">8A-H</figureCitation>
<figureCitation id="59BC6261FD1FDE3144B337430466603E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male holotype of T. iphitus sp. nov., habitus (A: dorsal, B: left lateral, C: ventral, D: left fronto-lateral), aedeagus (E: dorsal, F: ventral, G: right lateral) and H: sternite 9." figureDoi="10.3897/asp.79.e66021.figure8" httpUri="https://binary.pensoft.net/fig/533986" pageId="0" pageNumber="57">8A-H</figureCitation>
), length between anterior edge of pronotum and elytral apex 4.6 mm, DNA barcode: MH917891. Diagnostic combination of morphological characters as in Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Character matrix with discrete morphological characters for diagnostics of new Cameroonian Typoderus weevils (see also Discussion). - Characters: 1. - Body, colour of darkest specimens (Fig. 3, that is of a beetle at least a few days old, because specimens freshly emerged from the pupa are much paler): greyish: 0; brownish: 1; blackish: 2. 2. - Elytra at middle in cross-section: triangular (Figs 5 A-H, 6 A-H); elytral disk flat, ridge in interstriae 6 - 7 forming elytral lateral contour in dorsal view: 0; cylindrical (Figs 7 A-H, 8 A-H, 9 A-H); elytral disk evenly rounded laterally, ridge in interstriae 6 - 7 positioned inside lateral contour in dorsal view: 1. 3. - Hind tibiae, additional apical projection near inner apical spur (in at least some males): absent (Figs 7 B, 8 B, 9 B): 0; present (Figs 5 B, 6 B): 1. 4. - Pilosity on elytral declivity (excluding that on elytral protuberances), whether similar to that on the rest of elytra; similar: 0; dissimilar, notably denser and longer (Fig. 9 A, except abraded specimens): 1. 5. - Ratio of length to width of sclerotized part of aedeagus, dorsal view: one (aedeagus subquadrate as in Figs 5 E, 6 E, 9 E or notably unsclerotized, as in Fig. 8 E): 1; two (aedeagus elongate, Fig. 7 E): 2. 6. - Numerous (about 10) &quot; teeth &quot; inside internal sac of aedeagus: absent (Figs 5 E-F, 6 E-F, 8 E-F, 9 E-F): 0; present (Figs 7 E-F): 1. 7. - Tegminal apodeme: absent (Figs 6 F, 8 F) or indistinguishably short (Figs 5 F, 9 F): 0; present, consisting of two rods (Fig. 7 F): 1." httpUri="http://table.plazi.org/id/9CBF17CBDB7A16608D6711C763B03F8C" pageId="0" pageNumber="57" tableUuid="9CBF17CBDB7A16608D6711C763B03F8C">4</tableCitation>
<tableCitation id="8DEABF1388189099640F76BB81CD7C23" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Character matrix with discrete morphological characters for diagnostics of new Cameroonian Typoderus weevils (see also Discussion). - Characters: 1. - Body, colour of darkest specimens (Fig. 3, that is of a beetle at least a few days old, because specimens freshly emerged from the pupa are much paler): greyish: 0; brownish: 1; blackish: 2. 2. - Elytra at middle in cross-section: triangular (Figs 5 A-H, 6 A-H); elytral disk flat, ridge in interstriae 6 - 7 forming elytral lateral contour in dorsal view: 0; cylindrical (Figs 7 A-H, 8 A-H, 9 A-H); elytral disk evenly rounded laterally, ridge in interstriae 6 - 7 positioned inside lateral contour in dorsal view: 1. 3. - Hind tibiae, additional apical projection near inner apical spur (in at least some males): absent (Figs 7 B, 8 B, 9 B): 0; present (Figs 5 B, 6 B): 1. 4. - Pilosity on elytral declivity (excluding that on elytral protuberances), whether similar to that on the rest of elytra; similar: 0; dissimilar, notably denser and longer (Fig. 9 A, except abraded specimens): 1. 5. - Ratio of length to width of sclerotized part of aedeagus, dorsal view: one (aedeagus subquadrate as in Figs 5 E, 6 E, 9 E or notably unsclerotized, as in Fig. 8 E): 1; two (aedeagus elongate, Fig. 7 E): 2. 6. - Numerous (about 10) &quot; teeth &quot; inside internal sac of aedeagus: absent (Figs 5 E-F, 6 E-F, 8 E-F, 9 E-F): 0; present (Figs 7 E-F): 1. 7. - Tegminal apodeme: absent (Figs 6 F, 8 F) or indistinguishably short (Figs 5 F, 9 F): 0; present, consisting of two rods (Fig. 7 F): 1." httpUri="http://table.plazi.org/id/9CBF17CBDB7A16608D6711C763B03F8C" pageId="0" pageNumber="57" tableUuid="9CBF17CBDB7A16608D6711C763B03F8C">4</tableCitation>
. Sister clade to
<taxonomicName genus="T." lsidName="T. amphion" pageId="0" pageNumber="57" rank="species" species="amphion">
<emphasis italics="true" pageId="0" pageNumber="57">T. amphion</emphasis>
<taxonomicName id="DD172B5DEAF0CD485C3FFDBBE8747536" lsidName="T. amphion" pageId="0" pageNumber="57" rank="species" species="amphion">
<emphasis id="618328C86D7499BAF20725FAD3455B78" italics="true" pageId="0" pageNumber="57">T. amphion</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
<emphasis id="4E23C8BF611833E80E5E48BAE5CA9759" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
,
<taxonomicName genus="T." lsidName="T. telamon" pageId="0" pageNumber="57" rank="species" species="telamon">
<emphasis italics="true" pageId="0" pageNumber="57">T. telamon</emphasis>
<taxonomicName id="374828F1EC1742A229391B34830E67D0" lsidName="T. telamon" pageId="0" pageNumber="57" rank="species" species="telamon">
<emphasis id="720842CB96AF5BC373A04BE303B7BA6E" italics="true" pageId="0" pageNumber="57">T. telamon</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
<emphasis id="AC2D79C9B4587685C4371D0F002DA72C" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
and
<taxonomicName genus="T." lsidName="T. clytius" pageId="0" pageNumber="57" rank="species" species="clytius">
<emphasis italics="true" pageId="0" pageNumber="57">T. clytius</emphasis>
<taxonomicName id="82637EE943CE712176A46CDE2DD1EAD5" lsidName="T. clytius" pageId="0" pageNumber="57" rank="species" species="clytius">
<emphasis id="7F0336F4EFFC105BAEE28C4007256588" italics="true" pageId="0" pageNumber="57">T. clytius</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
<emphasis id="95D6D01AEAED6F9E81A5BB467562F89C" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
(Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood inference phylogram of Typoderus weevils rooted on Lupangus (analysis A 3, root is not shown). Cameroonian species form a clade. Digits at internodes are bootstrap values of 50 % and above. Non-Cameroonian (= Tanzanian) terminals are collapsed in species. Terminal names consist of specimen number (Table 3) and sifting sample number (Table 1). Habitus images are to scale; their numbers indicate exact Cameroonian specimens imaged. Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure3" httpUri="https://binary.pensoft.net/fig/533981" pageId="0" pageNumber="57">3</figureCitation>
<figureCitation id="D15E0640616A8D18EB157B2A220A4F68" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood inference phylogram of Typoderus weevils rooted on Lupangus (analysis A 3, root is not shown). Cameroonian species form a clade. Digits at internodes are bootstrap values of 50 % and above. Non-Cameroonian (= Tanzanian) terminals are collapsed in species. Terminal names consist of specimen number (Table 3) and sifting sample number (Table 1). Habitus images are to scale; their numbers indicate exact Cameroonian specimens imaged. Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure3" httpUri="https://binary.pensoft.net/fig/533981" pageId="0" pageNumber="57">3</figureCitation>
).
</paragraph>
<caption doi="10.3897/asp.79.e66021.figure8" httpUri="https://binary.pensoft.net/fig/533986" pageId="0" pageNumber="57" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="57">
<emphasis bold="true" pageId="0" pageNumber="57">Figure 8.</emphasis>
<caption id="7E0D3F4BFD6ADDC723C76ECEC170C102" doi="10.3897/asp.79.e66021.figure8" httpUri="https://binary.pensoft.net/fig/533986" pageId="0" pageNumber="57" start="Figure 8" startId="F8">
<paragraph id="E4F243A452A308F43BB2AE4609EF6819" pageId="0" pageNumber="57">
<emphasis id="136CC036905B77F307CEB00F704BBE55" bold="true" pageId="0" pageNumber="57">Figure 8.</emphasis>
Male holotype of
<taxonomicName genus="T." lsidName="T. iphitus" pageId="0" pageNumber="57" rank="species" species="iphitus">
<emphasis italics="true" pageId="0" pageNumber="57">T. iphitus</emphasis>
<taxonomicName id="947791EA73D76DCC7E7613480596C532" lsidName="T. iphitus" pageId="0" pageNumber="57" rank="species" species="iphitus">
<emphasis id="86652D846D3C60648BD27517E0C9D3BA" italics="true" pageId="0" pageNumber="57">T. iphitus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
<emphasis id="9FC99FAD066EF3F9F77A9182CFB8EF88" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
, habitus (
<emphasis bold="true" pageId="0" pageNumber="57">A</emphasis>
<emphasis id="F09BC2EDB031636B5A482DA4DEFC8B8E" bold="true" pageId="0" pageNumber="57">A</emphasis>
: dorsal,
<emphasis bold="true" pageId="0" pageNumber="57">B</emphasis>
<emphasis id="06DD74A32F841E6414D6F39EBD599CC5" bold="true" pageId="0" pageNumber="57">B</emphasis>
: left lateral,
<emphasis bold="true" pageId="0" pageNumber="57">C</emphasis>
<emphasis id="B03448AE65A932FAB9DDFD2AEC783A27" bold="true" pageId="0" pageNumber="57">C</emphasis>
: ventral,
<emphasis bold="true" pageId="0" pageNumber="57">D</emphasis>
<emphasis id="338F62E7FCE13ACA889557CBDEC0D9C7" bold="true" pageId="0" pageNumber="57">D</emphasis>
: left fronto-lateral), aedeagus (
<emphasis bold="true" pageId="0" pageNumber="57">E</emphasis>
<emphasis id="253811760A3009A3D39A1F844FB7A5AF" bold="true" pageId="0" pageNumber="57">E</emphasis>
: dorsal,
<emphasis bold="true" pageId="0" pageNumber="57">F</emphasis>
<emphasis id="040AAB414C0EBCD833F9716C551D0230" bold="true" pageId="0" pageNumber="57">F</emphasis>
: ventral,
<emphasis bold="true" pageId="0" pageNumber="57">G</emphasis>
<emphasis id="B87EBC9114EA59FA23B39E480B374FB1" bold="true" pageId="0" pageNumber="57">G</emphasis>
: right lateral) and
<emphasis bold="true" pageId="0" pageNumber="57">H</emphasis>
<emphasis id="9DB60A15318E7417DE77277DD4057E16" bold="true" pageId="0" pageNumber="57">H</emphasis>
: sternite 9.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="materials_examined">
<paragraph pageId="0" pageNumber="57">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="57">
<materialsCitation ID-GBIF-Occurrence="3080451310" collectingDate="2015-12-29" collectorName="V. Grebennikov" country="CAMEROON" elevation="1977" latitude="4.8017" location="Mt. Kupe" longLatPrecision="7" longitude="9.7015" specimenCount="1" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
male (CNC): &quot;
<collectingCountry name="Cameroon">CAMEROON</collectingCountry>
<subSubSection id="F3C2F8CF44C35ED4D2703B0A9E5DDC46" pageId="0" pageNumber="57" type="materials_examined">
<paragraph id="799D77B522CD4EE67E54956E8F20AA75" pageId="0" pageNumber="57">Material examined.</paragraph>
<paragraph id="74CD9DFBB0A067C95521EA87B40B5A2B" pageId="0" pageNumber="57">
<materialsCitation id="51B392D834AD3865720E9EAD1979CEE8" ID-GBIF-Occurrence="3080451310" collectingDate="2015-12-29" collectorName="V. Grebennikov" country="CAMEROON" elevation="1977" latitude="4.8017" location="Mt. Kupe" longLatPrecision="7" longitude="9.7015" specimenCount="1" typeStatus="Holotype">
<typeStatus id="82C55013E4B38FD585FBE223207A12B5">Holotype</typeStatus>
male (CNC): &quot;
<collectingCountry id="857FA6B803A219A85C9E8D725B207CEE" name="Cameroon">CAMEROON</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:340A628155E5598B88B266968BEBDA27:4D18AF91B6C6FE2A8C86886E6C8DA884" country="CAMEROON" latitude="4.8017" longLatPrecision="7" longitude="9.7015" name="Mt. Kupe">Mt. Kupe</location>
<location id="4D18AF91B6C6FE2A8C86886E6C8DA884" LSID="urn:lsid:plazi:treatment:340A628155E5598B88B266968BEBDA27:4D18AF91B6C6FE2A8C86886E6C8DA884" country="CAMEROON" latitude="4.8017" longLatPrecision="7" longitude="9.7015" name="Mt. Kupe">Mt. Kupe</location>
,
<geoCoordinate degrees="4.8017" direction="north" orientation="latitude" precision="5" value="4.8017">4.8017</geoCoordinate>
<geoCoordinate id="410B82CC8A6E751741F78084CA989FD8" degrees="4.8017" direction="north" orientation="latitude" precision="5" value="4.8017">4.8017</geoCoordinate>
,
<geoCoordinate degrees="9.7015" direction="east" orientation="longitude" precision="5" value="9.7015">9.7015</geoCoordinate>
<geoCoordinate id="E33278BE69CC447463DEE0AB262E7849" degrees="9.7015" direction="east" orientation="longitude" precision="5" value="9.7015">9.7015</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">1977m</elevation>
<quantity id="35EFF6B5AE37DCDF3310BBBC0AA27579" metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">
<elevation id="6F8FA290CAFF8C6C37DCF5FEF52488B0" metricMagnitude="3" metricUnit="m" metricValue="1.9769999999999999" unit="m" value="1977.0">1977m</elevation>
</quantity>
,
<collectingDate value="2015-12-29">29.xii.2015</collectingDate>
<collectingDate id="5FE1E983970059597F126F1FA20C5843" value="2015-12-29">29.xii.2015</collectingDate>
, sift.CM15,
<collectorName>V.Grebennikov</collectorName>
<collectorName id="F0787456B0C16571C75D058D29C04EE7">V.Grebennikov</collectorName>
&quot;,
<normalizedToken originalValue="“CNCCOLVG00009722”">&quot;CNCCOLVG00009722&quot;</normalizedToken>
<normalizedToken id="7CD125E08AE0F140B48E009244275F90" originalValue="“CNCCOLVG00009722”">&quot;CNCCOLVG00009722&quot;</normalizedToken>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3080451303" collectingDate="2015-12-29" collectorName="V. Grebennikov" country="CAMEROON" elevation="1525" latitude="4.8193" location="Mt. Kupe" longLatPrecision="7" longitude="9.7075" specimenCount="1" typeStatus="Paratypes">
<typeStatus>Paratypes</typeStatus>
<materialsCitation id="3997BF93F6669723A6012AC797203C9F" ID-GBIF-Occurrence="3080451303" collectingDate="2015-12-29" collectorName="V. Grebennikov" country="CAMEROON" elevation="1525" latitude="4.8193" location="Mt. Kupe" longLatPrecision="7" longitude="9.7075" specimenCount="1" typeStatus="Paratypes">
<typeStatus id="A479A0F5442D9F2C742AA72B5BF51A6D">Paratypes</typeStatus>
(CNC): 4, as in Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood inference phylogram of Typoderus weevils rooted on Lupangus (analysis A 3, root is not shown). Cameroonian species form a clade. Digits at internodes are bootstrap values of 50 % and above. Non-Cameroonian (= Tanzanian) terminals are collapsed in species. Terminal names consist of specimen number (Table 3) and sifting sample number (Table 1). Habitus images are to scale; their numbers indicate exact Cameroonian specimens imaged. Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure3" httpUri="https://binary.pensoft.net/fig/533981" pageId="0" pageNumber="57">3</figureCitation>
; specimens 9556, 9723 and 9724: same data as holotype; specimen 9531: &quot;
<collectingCountry name="Cameroon">CAMEROON</collectingCountry>
<figureCitation id="2DC85D8189E68B797B8BB534D4BD08E7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Maximum Likelihood inference phylogram of Typoderus weevils rooted on Lupangus (analysis A 3, root is not shown). Cameroonian species form a clade. Digits at internodes are bootstrap values of 50 % and above. Non-Cameroonian (= Tanzanian) terminals are collapsed in species. Terminal names consist of specimen number (Table 3) and sifting sample number (Table 1). Habitus images are to scale; their numbers indicate exact Cameroonian specimens imaged. Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure3" httpUri="https://binary.pensoft.net/fig/533981" pageId="0" pageNumber="57">3</figureCitation>
; specimens 9556, 9723 and 9724: same data as holotype; specimen 9531: &quot;
<collectingCountry id="91089A3E9AAF736B89DF7511A5F3345C" name="Cameroon">CAMEROON</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:340A628155E5598B88B266968BEBDA27:6A962D642815F16FFF8ADD215A0AC3DA" country="CAMEROON" latitude="4.8193" longLatPrecision="7" longitude="9.7075" name="Mt. Kupe">Mt. Kupe</location>
<location id="6A962D642815F16FFF8ADD215A0AC3DA" LSID="urn:lsid:plazi:treatment:340A628155E5598B88B266968BEBDA27:6A962D642815F16FFF8ADD215A0AC3DA" country="CAMEROON" latitude="4.8193" longLatPrecision="7" longitude="9.7075" name="Mt. Kupe">Mt. Kupe</location>
,
<geoCoordinate degrees="4.8193" direction="north" orientation="latitude" precision="5" value="4.8193">4.8193</geoCoordinate>
<geoCoordinate id="86F8F293BD964ED0AA9407536323C373" degrees="4.8193" direction="north" orientation="latitude" precision="5" value="4.8193">4.8193</geoCoordinate>
,
<geoCoordinate degrees="9.7075" direction="east" orientation="longitude" precision="5" value="9.7075">9.7075</geoCoordinate>
<geoCoordinate id="309B524E0F26BED2217569D768BC7BC8" degrees="9.7075" direction="east" orientation="longitude" precision="5" value="9.7075">9.7075</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.525" unit="m" value="1525.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.525" unit="m" value="1525.0">1525m</elevation>
<quantity id="CCF46309EE9373CB8522317B3889F3DB" metricMagnitude="3" metricUnit="m" metricValue="1.525" unit="m" value="1525.0">
<elevation id="D62FE8559A27272067450AEBF0FFF826" metricMagnitude="3" metricUnit="m" metricValue="1.525" unit="m" value="1525.0">1525m</elevation>
</quantity>
,
<collectingDate value="2015-12-29">29.xii.2015</collectingDate>
<collectingDate id="D96CAC95355ED048EC203F6CBAA27DA3" value="2015-12-29">29.xii.2015</collectingDate>
, sift.CM13,
<collectorName>V.Grebennikov</collectorName>
<collectorName id="EAEE8FA6ED738E170956D9B5619F8058">V.Grebennikov</collectorName>
&quot;
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="distribution">
<paragraph pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="57">
<subSubSection id="A7C81B2DCE56B897EEC71E35294525EA" pageId="0" pageNumber="57" type="distribution">
<paragraph id="D3D9176463ECFD9A8E7896CCA3C4D265" pageId="0" pageNumber="57">Distribution.</paragraph>
<paragraph id="3C893F16663B5EBB669371B89B1889B7" pageId="0" pageNumber="57">
Known only from Mt. Kupe in Cameroon, in sympatry with
<taxonomicName genus="T." lsidName="T. telamon" pageId="0" pageNumber="57" rank="species" species="telamon">
<emphasis italics="true" pageId="0" pageNumber="57">T. telamon</emphasis>
<taxonomicName id="7C6691AB6BEB27ED1A1E9BAEDF281E10" lsidName="T. telamon" pageId="0" pageNumber="57" rank="species" species="telamon">
<emphasis id="51643BCF2976CB44B17D1BBA5B5C9E47" italics="true" pageId="0" pageNumber="57">T. telamon</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
<emphasis id="4D8D7EF05D10E5567A08768430E59EC6" bold="true" pageId="0" pageNumber="57">sp. nov.</emphasis>
Elevation: 1,525-1,977 m.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="etymology">
<paragraph pageId="0" pageNumber="57">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="57">Iphitus, from ancient Greek mythology, one of the Argonauts, brother of Clytius; noun in apposition.</paragraph>
<subSubSection id="D0F67D7F1AACC6BC3B254CBCFE9F11BB" pageId="0" pageNumber="57" type="etymology">
<paragraph id="4BEA8D1DD31F47C9B82327DCAE93F4AA" pageId="0" pageNumber="57">Etymology.</paragraph>
<paragraph id="FBC3E7AE3CDBB992EF7E20AD37661485" pageId="0" pageNumber="57">Iphitus, from ancient Greek mythology, one of the Argonauts, brother of Clytius; noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="comments">
<paragraph pageId="0" pageNumber="57">Comment.</paragraph>
<paragraph pageId="0" pageNumber="57">
<subSubSection id="5A9EB9E35AF8BF29C7195FC78F415EE4" pageId="0" pageNumber="57" type="comments">
<paragraph id="CF2D317ABF69026FBF0A5CFACCFA7A3B" pageId="0" pageNumber="57">Comment.</paragraph>
<paragraph id="29C5B79531A7A15F1A5CF15145271819" pageId="0" pageNumber="57">
Being morphologically indistinguishable and sympatric, it is more parsimonious to treat all specimens assigned to this species as the same biological entity, despite a relatively deep intraspecific divergence (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neighbour Joining clustering of 20 DNA barcodes of Cameroonian Typoderus (analysis A 1). Terminal names consist of BOLD sample ID (its last four digits correspond to specimen number), litter sifting sample number (Table 1), length of the DNA barcode (number of ambiguous nucleotides is in square brackets), BIN three-letter four-digit number and GenBank accession number. Names next to clusters are those of the new species described in section 4.2. HT denote holotypes." figureDoi="10.3897/asp.79.e66021.figure2" httpUri="https://binary.pensoft.net/fig/533980" pageId="0" pageNumber="57">2</figureCitation>
<figureCitation id="EEE6F2A1BC133006F8A8CD2D312910ED" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Neighbour Joining clustering of 20 DNA barcodes of Cameroonian Typoderus (analysis A 1). Terminal names consist of BOLD sample ID (its last four digits correspond to specimen number), litter sifting sample number (Table 1), length of the DNA barcode (number of ambiguous nucleotides is in square brackets), BIN three-letter four-digit number and GenBank accession number. Names next to clusters are those of the new species described in section 4.2. HT denote holotypes." figureDoi="10.3897/asp.79.e66021.figure2" httpUri="https://binary.pensoft.net/fig/533980" pageId="0" pageNumber="57">2</figureCitation>
-
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Maximum clade credibility Cameroonian Typoderus tree with median ages derived from the BEAST analysis A 4. The 95 % credibility interval for each node is given with a horizontal bar. Digits at nodes are million years before present. Terminal names consist of specimen number (Table 3), GenBank accession number and sifting sample number (Table 1). Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure4" httpUri="https://binary.pensoft.net/fig/533982" pageId="0" pageNumber="57">4</figureCitation>
<figureCitation id="6EDD599153C7A3D28EC3B684734F14F1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Maximum clade credibility Cameroonian Typoderus tree with median ages derived from the BEAST analysis A 4. The 95 % credibility interval for each node is given with a horizontal bar. Digits at nodes are million years before present. Terminal names consist of specimen number (Table 3), GenBank accession number and sifting sample number (Table 1). Names next to clades are those of the new species described in section 4.2." figureDoi="10.3897/asp.79.e66021.figure4" httpUri="https://binary.pensoft.net/fig/533982" pageId="0" pageNumber="57">4</figureCitation>
).
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@ -1,56 +1,56 @@
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<taxonomicName id="CDB3001336911D0751A6A73413829A77" LSID="3427682B-4FA8-5BF9-B950-67825F08C12E" authority="Itagaki &amp; Hosoya" authorityName="Itagaki &amp; Hosoya" authorityYear="2023" class="Leotiomycetes" family="Mollisiaceae" genus="Neobelonopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Neobelonopsis obtusa" order="Helotiales" pageId="0" pageNumber="45" phylum="Ascomycota" rank="species" species="obtusa" status="sp. nov.">Neobelonopsis obtusa Itagaki &amp; Hosoya</taxonomicName>
<taxonomicNameLabel id="92F2AC78B0068024BE0584F6E160D49B" pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
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@ -83,7 +83,7 @@ and
<paragraph id="6855A594F33BA9378BB82211795034FB" pageId="0" pageNumber="45">Holotype.</paragraph>
<paragraph id="950AD732DC2539F9BE54FABC550A0010" pageId="0" pageNumber="45">
TNS-F-15602, Iryuda, Odawara City, Kanagawa Pref., 12 April 2007, on decaying wood of
<taxonomicName id="44A792DBD40C7B9A55FAEF35ED69D34B" authority="Thunb. var. japonica" class="Equisetopsida" family="Lauraceae" genus="Aucuba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aucuba japonica subsp. var. var. japonica" order="Garryales" pageId="0" pageNumber="45" phylum="Magnoliophyta" rank="variety" species="japonica" subSpecies="var." variety="japonica">Aucuba japonica Thunb. var. japonica</taxonomicName>
<taxonomicName id="44A792DBD40C7B9A55FAEF35ED69D34B" authority="Thunb. var. japonica" class="Equisetopsida" family="Lauraceae" genus="Aucuba" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Aucuba japonica subsp. var. var. japonica" order="Laurales" pageId="0" pageNumber="45" phylum="Magnoliophyta" rank="variety" species="japonica" subSpecies="var." variety="japonica">Aucuba japonica Thunb. var. japonica</taxonomicName>
, ex-holotype culture NBRC 115381.
</paragraph>
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@ -142,7 +142,7 @@ diam., hyaline, thin-walled.
<caption id="2B5A6B756CBB46DCB7121C277E853720" doi="10.3897/mycokeys.99.90117.figure9" httpUri="https://binary.pensoft.net/fig/892425" pageId="0" pageNumber="45" start="Figure 9" startId="F9">
<paragraph id="900C8DEAD133246F649F69F8D455B72B" pageId="0" pageNumber="45">
<emphasis id="2B6ADF54289D7ED1EBDB6D2B8924C924" bold="true" pageId="0" pageNumber="45">Figure 9.</emphasis>
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<emphasis id="D0FE4B3B9BB06B47672B50DA4537E14F" italics="true" pageId="0" pageNumber="45">Neobelonopsis obtusa</emphasis>
</taxonomicName>
(TNS-F-15602, holotype)
@ -227,7 +227,7 @@ sp.; TNS-F-86658, Yugashima, Izu City, Shizuoka Pref.,
<paragraph id="9DC9FFA135AF53EE0B212EE17B2E82E2" pageId="0" pageNumber="45">Notes.</paragraph>
<paragraph id="B146D614DADBF18F5233B3B806272F13" pageId="0" pageNumber="45">
The ectal excipulum consisting of closely packed brownish cells of
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<taxonomicName id="1952E106FBD67AFAA9D4D2BC16943D5F" authorityName="Itagaki &amp; Hosoya" authorityYear="2023" class="Leotiomycetes" family="Mollisiaceae" genus="Neobelonopsis" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Neobelonopsis obtusa" order="Helotiales" pageId="0" pageNumber="45" phylum="Ascomycota" rank="species" species="obtusa">
<emphasis id="DECB9FAD0D303ABC2D73E2453A4D03FC" italics="true" pageId="0" pageNumber="45">Neobelonopsis obtusa</emphasis>
</taxonomicName>
is similar to that of
@ -235,7 +235,7 @@ is similar to that of
<emphasis id="ED7BB129DBB25F1A7D8B7853A4009A38" italics="true" pageId="0" pageNumber="45">N. didymospora</emphasis>
</taxonomicName>
. However, the two species can be easily distinguished by the stable number of septa of ascospores (3-septate vs. 1-septate).
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<emphasis id="C485D9373F97ACA3D7B308B8B09D761C" italics="true" pageId="0" pageNumber="45">Neobelonopsis obtusa</emphasis>
</taxonomicName>
forms an asexual stage on CMA (Figs

View file

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<paragraph id="62211A4F092E11DD02CE2CD06CAE4B9D" pageId="15" pageNumber="34">
<taxonomicName id="A43ADBFFD42866C4734DB34192E1873A" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella" order="Opiliones" pageId="15" pageNumber="34" phylum="Arthropoda" rank="genus">Acuclavella</taxonomicName>
Shear, 1986: 13. Type species
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella cosmetoides" order="Opiliones" pageId="15" pageNumber="34" phylum="Arthropoda" rank="species" species="cosmetoides">Acuclavella cosmetoides</taxonomicName>
<taxonomicName id="1BE786679EA32BCD0367A2535C340E63" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella cosmetoides" order="Opiliones" pageId="15" pageNumber="34" phylum="Arthropoda" rank="species" species="cosmetoides">Acuclavella cosmetoides</taxonomicName>
Shear, 1986.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="34" type="description">
<paragraph pageId="15" pageNumber="34">Description.</paragraph>
<paragraph pageId="15" pageNumber="34">
<subSubSection id="DAB5E994C89F782D890D484B8B1690CD" pageId="15" pageNumber="34" type="description">
<paragraph id="902F3BFE4EAB54ACD19B6530BE466BF6" pageId="15" pageNumber="34">Description.</paragraph>
<paragraph id="0C0839D2A08110F67F3FF8EADC4E91D8" pageId="15" pageNumber="34">
The simple, distally tapering penis (Appendix VIII, Figure I) and short ovipositor (Appendix III, Figure 2) morphologies are conserved across species, with intraspecific variation seemingly as great as interspecific variation. Penis sheath with two sclerotized bands. Metatarsus of leg II with or without false leg articulations (Appendix IX). Distitarsi with three segments on legs I and II; distal end of legs III and IV with two constrictions, each comprised of two segments. All males with raised, glandular (
<bibRefCitation author="Shear, WA" journalOrPublisher="American Museum Novitates" pageId="41" pageNumber="60" pagination="1 - 29" title="A cladistic analysis of the opilionid superfamily Ischyropsalidoidea, with description of the new family Ceratolasmatidae, the new genus Acuclavella, and four new species." volume="2844" year="1986">Shear 1986</bibRefCitation>
<bibRefCitation id="E3FDF113A2A1BEF44C9604A49472C434" author="Shear, WA" journalOrPublisher="American Museum Novitates" pageId="41" pageNumber="60" pagination="1 - 29" title="A cladistic analysis of the opilionid superfamily Ischyropsalidoidea, with description of the new family Ceratolasmatidae, the new genus Acuclavella, and four new species." volume="2844" year="1986">Shear 1986</bibRefCitation>
), setose mound dorsally on basal article of chelicerae, setae often capped with secretions (MorphBank image 822802); mounds lacking in females. Chela teeth diaphanous, with two heavily sclerotized teeth distally on fingers (MorphBank image 822817). All leg coxae with prolateral and retrolateral tubercles in the form of distended clusters of warts (MorphBank image 828524). Epistome of stomotheca horn-like, projecting outward (MorphBank image 828520) to strongly decurved (MorphBank image 82215), without apparent interspecific trends. Pseudotrachea of pedipalpal coxaphysis sclerotized ctenoid (MorphBank image 822815).
</paragraph>
</subSubSection>
<subSubSection lastPageId="16" lastPageNumber="35" pageId="15" pageNumber="34" type="key to the species of acuclavella">
<paragraph pageId="15" pageNumber="34">
<subSubSection id="926A17957FD88140AB78BC933A12716D" lastPageId="16" lastPageNumber="35" pageId="15" pageNumber="34" type="key to the species of acuclavella">
<paragraph id="A29007BD2FBC99ACF3FAE7037F85BFF4" pageId="15" pageNumber="34">
Key to the species of
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella" order="Opiliones" pageId="15" pageNumber="34" phylum="Arthropoda" rank="genus">Acuclavella</taxonomicName>
<taxonomicName id="BF92C124D72264D898A765C1D55F9550" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella" order="Opiliones" pageId="15" pageNumber="34" phylum="Arthropoda" rank="genus">Acuclavella</taxonomicName>
.
</paragraph>
<paragraph pageId="15" pageNumber="34">
<paragraph id="BB288A05677BAFE414E726D47335A5B6" pageId="15" pageNumber="34">
This dichotomous key should allow users to identify the new species described herein. However, discovery of new morphologies in northern Idaho not described by
<bibRefCitation author="Shear, WA" journalOrPublisher="American Museum Novitates" pageId="41" pageNumber="60" pagination="1 - 29" title="A cladistic analysis of the opilionid superfamily Ischyropsalidoidea, with description of the new family Ceratolasmatidae, the new genus Acuclavella, and four new species." volume="2844" year="1986">Shear (1986)</bibRefCitation>
<bibRefCitation id="E0FE0E832238152F8A9EF98C4ECFD932" author="Shear, WA" journalOrPublisher="American Museum Novitates" pageId="41" pageNumber="60" pagination="1 - 29" title="A cladistic analysis of the opilionid superfamily Ischyropsalidoidea, with description of the new family Ceratolasmatidae, the new genus Acuclavella, and four new species." volume="2844" year="1986">Shear (1986)</bibRefCitation>
, and not ascribed to species herein, are currently of uncertain placement at the species level. These morphologies include females with pairs of spines on scute areas I-IV, individuals with three pairs of spines on scute areas I-III, and individuals with pairs of spines on scute areas I-II which are geographically and genetically distinct, but morphologically similar to
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella quattuor" order="Opiliones" pageId="15" pageNumber="34" phylum="Arthropoda" rank="species" species="quattuor">Acuclavella quattuor</taxonomicName>
<taxonomicName id="2BD31AC1E50D4D2E440304E786CE9368" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella quattuor" order="Opiliones" pageId="15" pageNumber="34" phylum="Arthropoda" rank="species" species="quattuor">Acuclavella quattuor</taxonomicName>
. For the purposes of this work, the four species named by
<bibRefCitation author="Shear, WA" journalOrPublisher="American Museum Novitates" pageId="41" pageNumber="60" pagination="1 - 29" title="A cladistic analysis of the opilionid superfamily Ischyropsalidoidea, with description of the new family Ceratolasmatidae, the new genus Acuclavella, and four new species." volume="2844" year="1986">Shear (1986)</bibRefCitation>
<bibRefCitation id="313F4CD0643AA0558E9DDC4505C6AAE7" author="Shear, WA" journalOrPublisher="American Museum Novitates" pageId="41" pageNumber="60" pagination="1 - 29" title="A cladistic analysis of the opilionid superfamily Ischyropsalidoidea, with description of the new family Ceratolasmatidae, the new genus Acuclavella, and four new species." volume="2844" year="1986">Shear (1986)</bibRefCitation>
are treated sensu stricto, and follow his diagnoses.
</paragraph>
<paragraph lastPageId="16" lastPageNumber="35" pageId="15" pageNumber="34">
<table lastPageId="16" lastPageNumber="35" pageId="15" pageNumber="34">
<tr pageId="15" pageNumber="34">
<td colspan="1" pageId="15" pageNumber="34" rowspan="1">Fig. 6</td>
<paragraph id="D9375B6DB7BCE716A4886F7462D41D15" lastPageId="16" lastPageNumber="35" pageId="15" pageNumber="34">
<table id="3641CE9BE077C4F398D9530E4499CBA4" lastPageId="16" lastPageNumber="35" pageId="15" pageNumber="34">
<tr id="0C9AB145905773452F0ADA75C9BA3635" pageId="15" pageNumber="34">
<td id="477401E5879714C094C7C364C9761E9F" colspan="1" pageId="15" pageNumber="34" rowspan="1">Fig. 6</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<pageBreakToken pageId="16" pageNumber="35" start="start">Fig</pageBreakToken>
<tr id="1FCC5753D64653131FF1D20F2DDFAA3C" pageId="16" pageNumber="35">
<td id="5EA89E945B9720D2E928D219EC4C27CC" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<pageBreakToken id="CF9817CEED9FCEAEC7E5FD10A7325C68" pageId="16" pageNumber="35" start="start">Fig</pageBreakToken>
. 9
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">Fig. 11</td>
<tr id="49DE57AEB22DD016215259192E58F00A" pageId="16" pageNumber="35">
<td id="823FA29A979222E5625C57E8A9754BAD" colspan="1" pageId="16" pageNumber="35" rowspan="1">Fig. 11</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella sheari" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="sheari">Acuclavella sheari</taxonomicName>
<tr id="3A89BEB8D25F472B59EF731BF24A75F8" pageId="16" pageNumber="35">
<td id="A3D499D08514F404E875C2DAE32DBB83" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="086D5FF7F6C8F687B976CBD59F092FB2" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella sheari" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="sheari">Acuclavella sheari</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella merickeli" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="merickeli">Acuclavella merickeli</taxonomicName>
<tr id="1280E019FDC3E7CAFC81019517CF531C" pageId="16" pageNumber="35">
<td id="446C5B4D8306A6F35235BA1958606C8F" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="77D11C0743D4C466628F8FD356E8619F" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella merickeli" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="merickeli">Acuclavella merickeli</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella leonardi" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="leonardi">Acuclavella leonardi</taxonomicName>
<tr id="784F721436204B02D15E6EEE54AA1BD5" pageId="16" pageNumber="35">
<td id="0EFEF07134FB947085F5CAEF1047C764" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="139109F4D9F1158E7B0CB9A94F75787B" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella leonardi" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="leonardi">Acuclavella leonardi</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella makah" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="makah">Acuclavella makah</taxonomicName>
<tr id="583869CAA8584FE38BD9F0818527042A" pageId="16" pageNumber="35">
<td id="8529A4C3EF955530F354E10A993219DA" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="38EE207D095B8A1572BD71F51659EA55" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella makah" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="makah">Acuclavella makah</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">Fig. 11</td>
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella quattuor" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="quattuor">Acuclavella quattuor</taxonomicName>
<tr id="02534D59D8CF94BD5448115243DDE96D" pageId="16" pageNumber="35">
<td id="091A9048AFB3A4B4BF071D28EF707726" colspan="1" pageId="16" pageNumber="35" rowspan="1">Fig. 11</td>
<td id="7A59796EFE7580BC1104DF60ECA97D89" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="0D8C2493743A07C1348D9A038EE64FF6" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella quattuor" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="quattuor">Acuclavella quattuor</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">Figure 1</td>
<tr id="3761E07073DA11E125B960CBA2A55919" pageId="16" pageNumber="35">
<td id="DF3079CB02D6DE831D1DBD10AB4C646D" colspan="1" pageId="16" pageNumber="35" rowspan="1">Figure 1</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella cosmetoides" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="cosmetoides">Acuclavella cosmetoides</taxonomicName>
<tr id="AC32C76497387C8914B658FA6A6B9053" pageId="16" pageNumber="35">
<td id="3F5D53BF696ED98B9C082F19F56F35C0" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="EF0241B82F1965333071FB3FDEC95279" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella cosmetoides" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="cosmetoides">Acuclavella cosmetoides</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella shoshone" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="shoshone">Acuclavella shoshone</taxonomicName>
<tr id="D91AC8F463CB75A58EAEE9BF378742EA" pageId="16" pageNumber="35">
<td id="613FE72184C1D9FAB294C1F26E68D88D" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="6F6B2307E48847D87A020917515C19E5" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella shoshone" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="shoshone">Acuclavella shoshone</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">Fig. 9</td>
<tr id="04577E76FB9B910F2002DA89B67A390E" pageId="16" pageNumber="35">
<td id="F89DF4005AF8CE5C051C724091728DE2" colspan="1" pageId="16" pageNumber="35" rowspan="1">Fig. 9</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella merickeli" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="merickeli">Acuclavella merickeli</taxonomicName>
<tr id="FB841D88A59FEC3179E5A6B37A62A0B1" pageId="16" pageNumber="35">
<td id="268AE0AD3613469880BCCF00EB349289" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="BA963DFD54BE412ABBB9D273B85E701B" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella merickeli" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="merickeli">Acuclavella merickeli</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella leonardi" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="leonardi">Acuclavella leonardi</taxonomicName>
<tr id="DBE8A002245D0818FBD85E5C8A1F3C2C" pageId="16" pageNumber="35">
<td id="61BA4727A94CE74826517F395B30BE9D" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="0E92AF6A90D5E48A65261A3641E66507" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella leonardi" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="leonardi">Acuclavella leonardi</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella makah" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="makah">Acuclavella makah</taxonomicName>
<tr id="C64B0CE49D6989D8D7E603FC08CAC1EB" pageId="16" pageNumber="35">
<td id="96D13C28DCD80CB639A3B1E05BA47776" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="A892C3730D6D3F9C311EC2538DF99AD7" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella makah" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="makah">Acuclavella makah</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella quattuor" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="quattuor">Acuclavella quattuor</taxonomicName>
<tr id="6BA5C1AFD710FA794021D9BC4001DD2F" pageId="16" pageNumber="35">
<td id="969BE84FBD4FE74A87179092670B070D" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="CC13DCFAEF5FDFC0D1CE6FD67E5C9381" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella quattuor" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="quattuor">Acuclavella quattuor</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella cosmetoides" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="cosmetoides">Acuclavella cosmetoides</taxonomicName>
<tr id="FBF5A00C5B124903A27238E0E57B77C9" pageId="16" pageNumber="35">
<td id="D56AD4E9A7C31A802FBEF57AB0F76487" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="DE83092BFE2E7CCF8B5A8D55729D0D95" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella cosmetoides" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="cosmetoides">Acuclavella cosmetoides</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">Fig. 9</td>
<tr id="06935799C2462DAF6576DDCFBE224566" pageId="16" pageNumber="35">
<td id="C84203FAA3F81613229291106F45F901" colspan="1" pageId="16" pageNumber="35" rowspan="1">Fig. 9</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella sheari" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="sheari">Acuclavella sheari</taxonomicName>
<tr id="3052ECBDABE5BF4338793175F6605988" pageId="16" pageNumber="35">
<td id="5E15EF6C290F9412077B36553072C05D" colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName id="529B9CFDA47A1B27A4CC07198F971AD2" class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella sheari" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="sheari">Acuclavella sheari</taxonomicName>
</td>
</tr>
<tr pageId="16" pageNumber="35">
<td colspan="1" pageId="16" pageNumber="35" rowspan="1">
<taxonomicName class="Arachnida" family="Ischyropsalididae" genus="Acuclavella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuclavella shoshone" order="Opiliones" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="shoshone">Acuclavella shoshone</taxonomicName>
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@ -1,295 +1,295 @@
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Agabus raffrayi group. Relief is expressed in meters above sea level." figureDoi="10.3897/zookeys.963.53470.figure1" httpUri="https://binary.pensoft.net/fig/444398" pageId="0" pageNumber="45">Figures 1</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. South African Agabus spp., aedeagal apices, lateral view A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov., Gydo Pass, Western Cape C A. austellus sp. nov. Groote Swartberg, Western Cape D A. austellus sp. nov. Sentinel Peak, KZN Drakensberg E A. riberae sp. nov. holotype, Kamiesberg, Northern Cape; F A. riberae sp. nov. paratype, Kamiesberg, Northern Cape G A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure6" httpUri="https://binary.pensoft.net/fig/444403" pageId="0" pageNumber="45">, 6E, F</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">, 8H</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. South African Agabus spp., microreticulation of male elytral disc A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov. Groote Swartberg, Western Cape C A. austellus sp. nov. Sentinel Peak, KZN Drakensberg D A. austellus sp. nov. Harkerville Forest, Western Cape E A. riberae sp. nov. Kamiesberg, Northern Cape F A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure9" httpUri="https://binary.pensoft.net/fig/444406" pageId="0" pageNumber="45">, 9E</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11M</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11Q</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">, 13</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">, 14</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Stream at 1100 m on Witwater-Langkloof road ca 1 km S. of junction, Kamiesberg, Northern Cape Province, South Africa. Type locality of Agabus riberae sp. nov., with DTB. September 19, 2010. Photo Rebecca Bilton." figureDoi="10.3897/zookeys.963.53470.figure18" httpUri="https://binary.pensoft.net/fig/444415" pageId="0" pageNumber="45">, 18</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Tra-Tra River at 485 m at Wupperthal, Cederberg range, Western Cape Province, South Africa. Locality for Agabus riberae sp. nov. September 28, 2018. Photo Stacey DeAmicis." figureDoi="10.3897/zookeys.963.53470.figure19" httpUri="https://binary.pensoft.net/fig/444416" pageId="0" pageNumber="45">, 19</figureCitation>
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<figureCitation id="ABDFC4F4DB2BCAAE7917643DD3F05C58" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. South African Agabus spp., aedeagal apices, lateral view A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov., Gydo Pass, Western Cape C A. austellus sp. nov. Groote Swartberg, Western Cape D A. austellus sp. nov. Sentinel Peak, KZN Drakensberg E A. riberae sp. nov. holotype, Kamiesberg, Northern Cape; F A. riberae sp. nov. paratype, Kamiesberg, Northern Cape G A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure6" httpUri="https://binary.pensoft.net/fig/444403" pageId="0" pageNumber="45">, 6E, F</figureCitation>
<figureCitation id="51A8740E2B7032D0B34D58EB9657C3A5" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">, 8H</figureCitation>
<figureCitation id="0A068C2EFACD0EF8EC7FAA167CB356AB" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. South African Agabus spp., microreticulation of male elytral disc A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov. Groote Swartberg, Western Cape C A. austellus sp. nov. Sentinel Peak, KZN Drakensberg D A. austellus sp. nov. Harkerville Forest, Western Cape E A. riberae sp. nov. Kamiesberg, Northern Cape F A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure9" httpUri="https://binary.pensoft.net/fig/444406" pageId="0" pageNumber="45">, 9E</figureCitation>
<figureCitation id="ACCA2C3EC232F70A72DE5B181898BC3A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11M</figureCitation>
<figureCitation id="EDE53046132C5163AD3D3655B2116D7D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11Q</figureCitation>
<figureCitation id="8036498EFEC8915F9667AE9082ACDA54" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">, 12</figureCitation>
<figureCitation id="28348786B9F52753F9F1E5381FC89451" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">, 13</figureCitation>
<figureCitation id="4246B72A1E11D2193071BAF2C9E8C918" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">, 14</figureCitation>
<figureCitation id="36B457AA24621DAD33297CCE9AFAE437" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Stream at 1100 m on Witwater-Langkloof road ca 1 km S. of junction, Kamiesberg, Northern Cape Province, South Africa. Type locality of Agabus riberae sp. nov., with DTB. September 19, 2010. Photo Rebecca Bilton." figureDoi="10.3897/zookeys.963.53470.figure18" httpUri="https://binary.pensoft.net/fig/444415" pageId="0" pageNumber="45">, 18</figureCitation>
<figureCitation id="D392247EC3921D68019FD5EA19BD67AB" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Tra-Tra River at 485 m at Wupperthal, Cederberg range, Western Cape Province, South Africa. Locality for Agabus riberae sp. nov. September 28, 2018. Photo Stacey DeAmicis." figureDoi="10.3897/zookeys.963.53470.figure19" httpUri="https://binary.pensoft.net/fig/444416" pageId="0" pageNumber="45">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type locality">
<paragraph pageId="0" pageNumber="45">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="45">South Africa, Northern Cape Province, Kamiesberg, stream on Witwater-Langkloof Road ca 1 km S. of junction, 30°23'41.30&quot;S 18°08'07.95&quot;E, 1100 m.</paragraph>
<subSubSection id="8B055E37A67EDBE07DAE2A768CC1E9BA" pageId="0" pageNumber="45" type="type locality">
<paragraph id="22BE0CA40BF58D821507665A83B1ADCF" pageId="0" pageNumber="45">Type locality.</paragraph>
<paragraph id="77A26E34BF234E5AC93096F071CF7D79" pageId="0" pageNumber="45">South Africa, Northern Cape Province, Kamiesberg, stream on Witwater-Langkloof Road ca 1 km S. of junction, 30°23'41.30&quot;S 18°08'07.95&quot;E, 1100 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type material">
<paragraph pageId="0" pageNumber="45">Type material.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Holotype</emphasis>
<subSubSection id="96E3A99D80D216B369725C94E0E33DC1" pageId="0" pageNumber="45" type="type material">
<paragraph id="9406A02F69C29D721C555DFEB8093C79" pageId="0" pageNumber="45">Type material.</paragraph>
<paragraph id="31A67064C6B21262A87F6CA54E07583B" pageId="0" pageNumber="45">
<emphasis id="A8E7D194AEECF5CA4731F69D832EF528" bold="true" italics="true" pageId="0" pageNumber="45">Holotype</emphasis>
♂ (AMG) labelled: &quot;19/ix/2010 South Africa NC Kamiesberg Stream on Witwater-Langkloof road ca 1 km S. of junction. D.T. Bilton leg.&quot;.
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Paratypes</emphasis>
<emphasis id="75A8F062ACA467205D1E58D1217C9E16" bold="true" italics="true" pageId="0" pageNumber="45">Paratypes</emphasis>
: 5 ♂ 5 ♀ (CBP, SANC, ZSM) labelled: &quot;19/ix/2010 South Africa NC Kamiesberg Stream on Witwater-Langkloof road ca 1 km S. of junction. D.T. Bilton leg.&quot;; paratype 4 ♂ 4 ♀ (AMG, CBP, ISAM, NHRS) labelled: &quot;19/1x/2010 South Africa NC Kamiesberg - stream above Studer Pass ca 5 km W of Witwater
<geoCoordinate degrees="30" direction="south" minutes="23" orientation="latitude" precision="1" seconds="13.49" value="-30.387081">30°23'13.49&quot;S</geoCoordinate>
<geoCoordinate id="9A58198598BE54AB5017E7EB83276097" degrees="30" direction="south" minutes="23" orientation="latitude" precision="1" seconds="13.49" value="-30.387081">30°23'13.49&quot;S</geoCoordinate>
,
<geoCoordinate degrees="18" direction="east" minutes="07" orientation="longitude" precision="1" seconds="05.78" value="18.118273">18°07'05.78&quot;E</geoCoordinate>
<geoCoordinate id="56CFA40F5F1E57835F44935B5A6A5789" degrees="18" direction="east" minutes="07" orientation="longitude" precision="1" seconds="05.78" value="18.118273">18°07'05.78&quot;E</geoCoordinate>
1,105 m D T Bilton leg.&quot;; paratype 1 ♀ (IBE) labelled: &quot;22/viii/2004 South Africa NC Kamiesberg - stream at top of Studer Pass G. Challet leg.&quot;; paratype ♂ (CBP) labelled: &quot;19/ix/2014 South Africa NC Kamiesberg temporary stream in Kamiesberg Pass D T Bilton leg.&quot;; paratypes 2 ♂ (CBP) labelled: &quot;18/ix/2014 South Africa NC Kamiesberg spring pool on Rondefontein Farm
<geoCoordinate degrees="30" direction="south" minutes="30" orientation="latitude" precision="1" seconds="05.59" value="-30.501553">30°30'05.59&quot;S</geoCoordinate>
<geoCoordinate id="13BC27EE1B681E2E0627D7C920813909" degrees="30" direction="south" minutes="30" orientation="latitude" precision="1" seconds="05.59" value="-30.501553">30°30'05.59&quot;S</geoCoordinate>
,
<geoCoordinate degrees="18" direction="east" minutes="08" orientation="longitude" precision="1" seconds="56.35" value="18.148987">18°08'56.35&quot;E</geoCoordinate>
<geoCoordinate id="F3707F8D25D1B2FCF6700FD4F91E27EB" degrees="18" direction="east" minutes="08" orientation="longitude" precision="1" seconds="56.35" value="18.148987">18°08'56.35&quot;E</geoCoordinate>
1023 m D T Bilton leg.&quot;; paratypes 4 ♂ 1 ♀ (CBP, NHRS) labelled: &quot;17/ix/2014 South Africa NC Kamiesberg stream nr. Damsland on N. side of Rooiberg
<geoCoordinate degrees="30" direction="south" minutes="23" orientation="latitude" precision="1" seconds="36.33" value="-30.393425">30°23'36.33&quot;S</geoCoordinate>
<geoCoordinate id="651E51E8D2B0AD1366E229BFC253ED8A" degrees="30" direction="south" minutes="23" orientation="latitude" precision="1" seconds="36.33" value="-30.393425">30°23'36.33&quot;S</geoCoordinate>
,
<geoCoordinate degrees="18" direction="east" minutes="06" orientation="longitude" precision="1" seconds="32.12" value="18.108923">18°06'32.12&quot;E</geoCoordinate>
<geoCoordinate id="7995C0E06BEC1CB856272D44DA3BCAAB" degrees="18" direction="east" minutes="06" orientation="longitude" precision="1" seconds="32.12" value="18.108923">18°06'32.12&quot;E</geoCoordinate>
1111 m D T Bilton leg.&quot;; paratype 1 ♂ (CBP) labelled: 19/ix/2014 South Africa NC Kamiesberg stream nr. De Kuilen 30°10'44.94&quot;S, 18°04'37.71&quot;E 940 m D T Bilton leg.&quot;; paratypes 2 ♂ (CBP) labelled: &quot;18/ix/2014 South Africa NC Kamiesberg - stream at bottom of Langkloof
<geoCoordinate degrees="30" direction="south" minutes="33" orientation="latitude" precision="1" seconds="16.98" value="-30.554716">30°33'16.98&quot;S</geoCoordinate>
<geoCoordinate id="3F08F6B9EDFB581F7F773FE0341E49BD" degrees="30" direction="south" minutes="33" orientation="latitude" precision="1" seconds="16.98" value="-30.554716">30°33'16.98&quot;S</geoCoordinate>
,
<geoCoordinate degrees="18" direction="east" minutes="08" orientation="longitude" precision="1" seconds="19.13" value="18.138647">18°08'19.13&quot;E</geoCoordinate>
<geoCoordinate id="2F0621D9ACB59E34D5F037DF7BAE2EB8" degrees="18" direction="east" minutes="08" orientation="longitude" precision="1" seconds="19.13" value="18.138647">18°08'19.13&quot;E</geoCoordinate>
594 m D T Bilton leg.&quot;; paratypes 2 ♂ 4 ♀ (AMG, CBP) labelled: &quot;28/ix/2018 South Africa WC Cederberg Tra-Tra river @ Wupperthal
<geoCoordinate degrees="32" direction="south" minutes="16" orientation="latitude" precision="1" seconds="45.35" value="-32.279263">32°16'45.35&quot;S</geoCoordinate>
<geoCoordinate id="5C90A3EC80F101AEFA96CD656E9A0872" degrees="32" direction="south" minutes="16" orientation="latitude" precision="1" seconds="45.35" value="-32.279263">32°16'45.35&quot;S</geoCoordinate>
,
<geoCoordinate degrees="19" direction="east" minutes="13" orientation="longitude" precision="1" seconds="04.32" value="19.217867">19°13'04.32&quot;E</geoCoordinate>
<geoCoordinate id="926E7C9703B3280BA27229E53E7FD216" degrees="19" direction="east" minutes="13" orientation="longitude" precision="1" seconds="04.32" value="19.217867">19°13'04.32&quot;E</geoCoordinate>
485 m D T Bilton leg.&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="diagnosis">
<paragraph pageId="0" pageNumber="45">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="C91E53F82EBD63A71CA58CAAA995E943" pageId="0" pageNumber="45" type="diagnosis">
<paragraph id="2A89FF09350F228A655D96D684C5A394" pageId="0" pageNumber="45">Diagnosis.</paragraph>
<paragraph id="B470ABA260768C9CC1F5421C3AB8ACD0" pageId="0" pageNumber="45">
Very similar to
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
<taxonomicName id="795D1CA65A3600CEEF277254F7F55E2C" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="4A80439AF7F41ED8EE0463B191E8E3D5" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov., differing from this species in having a relatively broad metasternal wing (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
<tableCitation id="00E225A0541424A3BBF2A97300EF2109" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
), and an elytral microreticulation dominated by larger, more irregular meshes than seen in most
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
<taxonomicName id="489E8225AAB941948EFB12AD65E99A9D" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="3777F29836F012A20AC93322F293EBB3" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. specimens (see above). Metatarsomere 5 is also somewhat longer in this species than in most
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
<taxonomicName id="91EE01C90F8E8747900369C3D58B6158" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="09C923C0B4AA41B69D4A52A3C20FC247" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="CD542A55C2DE821FA582C8CFBE22699E" pageId="0" pageNumber="45" type="description">
<paragraph id="D57A5093BE042E975CCA5E30BFE21D75" pageId="0" pageNumber="45">Description.</paragraph>
<paragraph id="72C7A636900C302C7BF82D8255A016F4" pageId="0" pageNumber="45">
Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">11M, Q</figureCitation>
<figureCitation id="4B0A082D481A9C5BC11557F00DA95887" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">11M, Q</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Colour</emphasis>
<paragraph id="08421A2ABC1B715C2C04AB00FC12D33D" pageId="0" pageNumber="45">
<emphasis id="A1193F7B4FDDAB1E6F88D913B066364A" bold="true" italics="true" pageId="0" pageNumber="45">Colour</emphasis>
: Head black with rufous interocular spots and an anterior rufous area. Pronotum black with rufous borders. Elytra rufopiceous to black. Ventral surface black, testaceous lines on abdominal segments rarely present, hypomeron and epipleuron rufotestaceous to rufous. Legs rufous to rufopiceous. Antennae and palpi testaceous.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName genus="Microreticulation" lsidName="Microreticulation" pageId="0" pageNumber="45" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Microreticulation</emphasis>
<paragraph id="8F4D39FD8E53947F9FF315AA410777CF" pageId="0" pageNumber="45">
<taxonomicName id="0019C7CC77E056610193558072295B13" genus="Microreticulation" lsidName="Microreticulation" pageId="0" pageNumber="45" rank="genus">
<emphasis id="2566C309E24DF13F74BBD9EDFE237A07" bold="true" italics="true" pageId="0" pageNumber="45">Microreticulation</emphasis>
</taxonomicName>
: Relatively fine on both pronotum and elytra, and rather similarly impressed in both sexes. The microreticulation of the elytral disc is typically dominated by relatively large, somewhat irregular meshes (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. South African Agabus spp., microreticulation of male elytral disc A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov. Groote Swartberg, Western Cape C A. austellus sp. nov. Sentinel Peak, KZN Drakensberg D A. austellus sp. nov. Harkerville Forest, Western Cape E A. riberae sp. nov. Kamiesberg, Northern Cape F A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure9" httpUri="https://binary.pensoft.net/fig/444406" pageId="0" pageNumber="45">9E</figureCitation>
<figureCitation id="98BE08116C187FD6A99334A404090EC2" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. South African Agabus spp., microreticulation of male elytral disc A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov. Groote Swartberg, Western Cape C A. austellus sp. nov. Sentinel Peak, KZN Drakensberg D A. austellus sp. nov. Harkerville Forest, Western Cape E A. riberae sp. nov. Kamiesberg, Northern Cape F A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure9" httpUri="https://binary.pensoft.net/fig/444406" pageId="0" pageNumber="45">9E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName genus="Structural" lsidName="Structural features" pageId="0" pageNumber="45" rank="species" species="features">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Structural features</emphasis>
<paragraph id="29615D4A92EFAB5A49036C26FF9C5E9B" pageId="0" pageNumber="45">
<taxonomicName id="6CF8EC0AA322CC597E16D0857C5A31A4" genus="Structural" lsidName="Structural features" pageId="0" pageNumber="45" rank="species" species="features">
<emphasis id="D41C106AC19F9E58D0611549428EEC2A" bold="true" italics="true" pageId="0" pageNumber="45">Structural features</emphasis>
</taxonomicName>
: Body length: 7.21-8.24 mm (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
<tableCitation id="BDAA02E0A9A71E4F37974C973278A8FA" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
). Hypomeron marginally visible in strict lateral view, lateral bead of pronotum narrow and well defined. Metasternal wing broad, WC/WS 3.0 or less in most specimens (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
<tableCitation id="B9ECD2C0936E09124C10B6391702B8DA" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
and Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
<figureCitation id="A4FD5D7DC482F0849684D7D78EACB1A2" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
). Pronotum broad, more than twice as broad as interocular distance (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
<tableCitation id="AC46DF3502DB80EB7FC7925037F95C41" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
and Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">13</figureCitation>
<figureCitation id="1C8E3CA132D88FD712EF467F5D0DBDC4" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">13</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Legs</emphasis>
<paragraph id="84297763CBA47A442931A3AFF8B10417" pageId="0" pageNumber="45">
<emphasis id="9D1657C8CDDAC681B6BD112FC2DED93D" bold="true" italics="true" pageId="0" pageNumber="45">Legs</emphasis>
: Male protarsal claws long,&gt; 1.6
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="29377FB7912C8E1A388480004162BAE2" originalValue="×">x</normalizedToken>
as long as protarsomere 4 (see Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
<tableCitation id="C4D73AB164AEA6BDE44298583E535291" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
and Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">14</figureCitation>
<figureCitation id="856F17D9AE6BC21A75AE53148F51DD18" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">14</figureCitation>
). Metatarsomere 2 short and broad, &lt;1.8
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="8BB3D1129B83F60DCBE7E355ACF14D02" originalValue="×">x</normalizedToken>
as long as broad (see Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
<tableCitation id="39A3B49A159F4C680F3E9E1292A2F8E8" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
) in most specimens. Metatarsomere 5 long and slender,&gt; 3.3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="EDEA15D47272FDC3989FA5059AAF6C53" originalValue="×">x</normalizedToken>
as long as broad in most males (see Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
<tableCitation id="2EC44B4AD1194EFF419691B08C44CDEE" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Male genitalia</emphasis>
<paragraph id="6F1BC2C8A2167C0CF8411C8317131AC5" pageId="0" pageNumber="45">
<emphasis id="5E31551DD4B8698F63E4C6CE87766533" bold="true" italics="true" pageId="0" pageNumber="45">Male genitalia</emphasis>
: Tip of aedeagus short, lacking the prolongation of the area located between the subapical broadening and the apical and subapical tooth present in some species in the group (see Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">8H</figureCitation>
<figureCitation id="75CB718DA16FB8942B75B96E1D9324FC" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">8H</figureCitation>
). There is some variation in the shape and size of the subapical tooth (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. South African Agabus spp., aedeagal apices, lateral view A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov., Gydo Pass, Western Cape C A. austellus sp. nov. Groote Swartberg, Western Cape D A. austellus sp. nov. Sentinel Peak, KZN Drakensberg E A. riberae sp. nov. holotype, Kamiesberg, Northern Cape; F A. riberae sp. nov. paratype, Kamiesberg, Northern Cape G A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure6" httpUri="https://binary.pensoft.net/fig/444403" pageId="0" pageNumber="45">6E, F</figureCitation>
<figureCitation id="99B07DBB5E2E0754C3E1BAE0097DAD0C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. South African Agabus spp., aedeagal apices, lateral view A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov., Gydo Pass, Western Cape C A. austellus sp. nov. Groote Swartberg, Western Cape D A. austellus sp. nov. Sentinel Peak, KZN Drakensberg E A. riberae sp. nov. holotype, Kamiesberg, Northern Cape; F A. riberae sp. nov. paratype, Kamiesberg, Northern Cape G A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure6" httpUri="https://binary.pensoft.net/fig/444403" pageId="0" pageNumber="45">6E, F</figureCitation>
), this being relatively long and narrow in most specimens examined (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. South African Agabus spp., aedeagal apices, lateral view A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov., Gydo Pass, Western Cape C A. austellus sp. nov. Groote Swartberg, Western Cape D A. austellus sp. nov. Sentinel Peak, KZN Drakensberg E A. riberae sp. nov. holotype, Kamiesberg, Northern Cape; F A. riberae sp. nov. paratype, Kamiesberg, Northern Cape G A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure6" httpUri="https://binary.pensoft.net/fig/444403" pageId="0" pageNumber="45">6F</figureCitation>
<figureCitation id="B01F4897011F2656F8FA1A7762CE8FDE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. South African Agabus spp., aedeagal apices, lateral view A A. austellus sp. nov. holotype, Keurboom, Western Cape B A. austellus sp. nov., Gydo Pass, Western Cape C A. austellus sp. nov. Groote Swartberg, Western Cape D A. austellus sp. nov. Sentinel Peak, KZN Drakensberg E A. riberae sp. nov. holotype, Kamiesberg, Northern Cape; F A. riberae sp. nov. paratype, Kamiesberg, Northern Cape G A. agulhas sp. nov. holotype, Rattelrivier, Western Cape." figureDoi="10.3897/zookeys.963.53470.figure6" httpUri="https://binary.pensoft.net/fig/444403" pageId="0" pageNumber="45">6F</figureCitation>
), but with both narrow and broader teeth being observed within the same population.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Female</emphasis>
<paragraph id="00C98CAB1ED20F5733C5451AB0571970" pageId="0" pageNumber="45">
<emphasis id="46C6548D25F39A22C29BD1873F62B37A" bold="true" pageId="0" pageNumber="45">Female</emphasis>
: Externally similar to males. Some specimens with dorsal microreticulation slightly more strongly impressed.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="65EB789A7059165913C5A1845C5A0BA2" pageId="0" pageNumber="45" type="distribution">
<paragraph id="A19C4E7724265DC6068158D42F99AA3A" pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph id="359EED910E4FA30E792828D86AE85217" pageId="0" pageNumber="45">
To date known only from Kamiesberg Range in the Northern Cape Province, and the eastern fringes of the Cederberg, Western Cape Province, Republic of South Africa (see Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Agabus raffrayi group. Relief is expressed in meters above sea level." figureDoi="10.3897/zookeys.963.53470.figure1" httpUri="https://binary.pensoft.net/fig/444398" pageId="0" pageNumber="45">1</figureCitation>
<figureCitation id="FF99BD4FB81613FAAD3E152350524943" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Agabus raffrayi group. Relief is expressed in meters above sea level." figureDoi="10.3897/zookeys.963.53470.figure1" httpUri="https://binary.pensoft.net/fig/444398" pageId="0" pageNumber="45">1</figureCitation>
), material from both areas being confirmed from COI sequences. The Kamiesberg represents a northerly outlier of Fynbos and Renosterveld vegetation in predominantly arid Namaqualand, and consequently have a diverse flora with a number of localised endemics (
<bibRefCitation author="Helme, N" journalOrPublisher=")" pageId="0" pageNumber="45" publicationUrl="http://www.cepf.net/Documents/helme_desmet.pdf" refId="B8" refString="Helme, N, Desmet, PG, 2006. A description of the endemic flora and vegetation of the Kamiesberg Uplands, Namaqualand, South Africa. Report for CEPF/SKEP. http://www.cepf.net/Documents/helme_desmet.pdf" title="A description of the endemic flora and vegetation of the Kamiesberg Uplands, Namaqualand, South Africa. Report for CEPF / SKEP." url="http://www.cepf.net/Documents/helme_desmet.pdf" year="2006">Helme and Desmet 2006</bibRefCitation>
<bibRefCitation id="3E643AC8D95461B8949C62743F40199B" author="Helme, N" journalOrPublisher=")" pageId="0" pageNumber="45" publicationUrl="http://www.cepf.net/Documents/helme_desmet.pdf" refId="B8" refString="Helme, N, Desmet, PG, 2006. A description of the endemic flora and vegetation of the Kamiesberg Uplands, Namaqualand, South Africa. Report for CEPF/SKEP. http://www.cepf.net/Documents/helme_desmet.pdf" title="A description of the endemic flora and vegetation of the Kamiesberg Uplands, Namaqualand, South Africa. Report for CEPF / SKEP." url="http://www.cepf.net/Documents/helme_desmet.pdf" year="2006">Helme and Desmet 2006</bibRefCitation>
). The mountains support the bulk of the global population of the endemic dytiscid
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Andex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andex insignis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="insignis">
<emphasis italics="true" pageId="0" pageNumber="45">Andex insignis</emphasis>
<taxonomicName id="8FE2487B42755D631A50CEC842B07CF4" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Andex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andex insignis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="4A4C050EF19864C59D1BD6D2E1146F9D" italics="true" pageId="0" pageNumber="45">Andex insignis</emphasis>
</taxonomicName>
Sharp, 1882 and a number of new, apparently endemic, water beetles have been described from the area in recent years (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3626.4.13" author="Bilton, DT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="45" pagination="589 - 592" refId="B1" refString="Bilton, DT, 2013. Crenitis bicolor sp. n. from the Kamiesberg of South Africa (Coleoptera: Hydrophilidae). Zootaxa 3626: 589 - 592, DOI: https://doi.org/10.11646/zootaxa.3626.4.13" title="Crenitis bicolor sp. n. from the Kamiesberg of South Africa (Coleoptera: Hydrophilidae)." url="https://doi.org/10.11646/zootaxa.3626.4.13" volume="3626" year="2013">Bilton 2013</bibRefCitation>
<bibRefCitation id="888D012A86D4FCAF7815937EC8F12B1D" DOI="https://doi.org/10.11646/zootaxa.3626.4.13" author="Bilton, DT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="45" pagination="589 - 592" refId="B1" refString="Bilton, DT, 2013. Crenitis bicolor sp. n. from the Kamiesberg of South Africa (Coleoptera: Hydrophilidae). Zootaxa 3626: 589 - 592, DOI: https://doi.org/10.11646/zootaxa.3626.4.13" title="Crenitis bicolor sp. n. from the Kamiesberg of South Africa (Coleoptera: Hydrophilidae)." url="https://doi.org/10.11646/zootaxa.3626.4.13" volume="3626" year="2013">Bilton 2013</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3957.4.5" author="Bilton, DT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="45" pagination="441 - 454" refId="B2" refString="Bilton, DT, 2015. A review of the Canthyporus exilis group, with the description of two new species (Coleoptera: Dytiscidae). Zootaxa 3957: 441 - 454, DOI: https://doi.org/10.11646/zootaxa.3957.4.5" title="A review of the Canthyporus exilis group, with the description of two new species (Coleoptera: Dytiscidae)." url="https://doi.org/10.11646/zootaxa.3957.4.5" volume="3957" year="2015">2015</bibRefCitation>
<bibRefCitation id="127C3AAB95B5D46C61678A831182F7C0" DOI="https://doi.org/10.11646/zootaxa.3957.4.5" author="Bilton, DT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="45" pagination="441 - 454" refId="B2" refString="Bilton, DT, 2015. A review of the Canthyporus exilis group, with the description of two new species (Coleoptera: Dytiscidae). Zootaxa 3957: 441 - 454, DOI: https://doi.org/10.11646/zootaxa.3957.4.5" title="A review of the Canthyporus exilis group, with the description of two new species (Coleoptera: Dytiscidae)." url="https://doi.org/10.11646/zootaxa.3957.4.5" volume="3957" year="2015">2015</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4137.4.11" author="Bilton, DT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="45" pagination="585 - 591" refId="B3" refString="Bilton, DT, 2016. Two new water beetles from the South African Cape (Coleoptera, Hydraenidae). Zootaxa 4137: 585 - 591, DOI: https://doi.org/10.11646/zootaxa.4137.4.11" title="Two new water beetles from the South African Cape (Coleoptera, Hydraenidae)." url="https://doi.org/10.11646/zootaxa.4137.4.11" volume="4137" year="2016">2016</bibRefCitation>
<bibRefCitation id="FCA45776E7405523D2A4559F40032CEC" DOI="https://doi.org/10.11646/zootaxa.4137.4.11" author="Bilton, DT" journalOrPublisher="Zootaxa" pageId="0" pageNumber="45" pagination="585 - 591" refId="B3" refString="Bilton, DT, 2016. Two new water beetles from the South African Cape (Coleoptera, Hydraenidae). Zootaxa 4137: 585 - 591, DOI: https://doi.org/10.11646/zootaxa.4137.4.11" title="Two new water beetles from the South African Cape (Coleoptera, Hydraenidae)." url="https://doi.org/10.11646/zootaxa.4137.4.11" volume="4137" year="2016">2016</bibRefCitation>
).
<taxonomicName authorityName="Bilton, Englund &amp; Bergsten" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus riberae" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberae">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus riberae</emphasis>
<taxonomicName id="BFFDB76DA2248D44E2121D02B1C46617" authorityName="Bilton, Englund &amp; Bergsten" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus riberae" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberae">
<emphasis id="914E2DD736C86D40DD6DA45D1FA66F12" italics="true" pageId="0" pageNumber="45">Agabus riberae</emphasis>
</taxonomicName>
sp. nov. appears to be the only
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
<taxonomicName id="00FD1AA9EB099B8D92A67A9A1A78A3BC" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="2590BE06171084E7087ACE9DFC0DF69A" italics="true" pageId="0" pageNumber="45">Agabus</emphasis>
</taxonomicName>
present in Kamiesberg, where it is abundant. In the Cederberg the species has been found close to Wupperthal, in the relatively dry northeastern fringes of the range.
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
<taxonomicName id="C5381356C6EA7C37AA62A42C35B2132E" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="98667824522596C2A198E211CBA34583" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. is the only species so far recorded from the wetter central areas of the Cederberg. All sites known to date experience predominantly winter rainfall.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="ecology">
<paragraph pageId="0" pageNumber="45">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="142DC4B3D00D86C865752B6E089FD339" pageId="0" pageNumber="45" type="ecology">
<paragraph id="820AC34D14D4CF0CF8D4B7A5AA13527F" pageId="0" pageNumber="45">Ecology.</paragraph>
<paragraph id="0CFBD3605AAD4C5DC775CB3E388DD526" pageId="0" pageNumber="45">
Found in streams and associated pools in the Kamiesberg and northeastern Cederberg ranges (Figs
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Stream at 1100 m on Witwater-Langkloof road ca 1 km S. of junction, Kamiesberg, Northern Cape Province, South Africa. Type locality of Agabus riberae sp. nov., with DTB. September 19, 2010. Photo Rebecca Bilton." figureDoi="10.3897/zookeys.963.53470.figure18" httpUri="https://binary.pensoft.net/fig/444415" pageId="0" pageNumber="45">18</figureCitation>
<figureCitation id="3C366EA39F46E6685463C2D6148CD981" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Stream at 1100 m on Witwater-Langkloof road ca 1 km S. of junction, Kamiesberg, Northern Cape Province, South Africa. Type locality of Agabus riberae sp. nov., with DTB. September 19, 2010. Photo Rebecca Bilton." figureDoi="10.3897/zookeys.963.53470.figure18" httpUri="https://binary.pensoft.net/fig/444415" pageId="0" pageNumber="45">18</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Tra-Tra River at 485 m at Wupperthal, Cederberg range, Western Cape Province, South Africa. Locality for Agabus riberae sp. nov. September 28, 2018. Photo Stacey DeAmicis." figureDoi="10.3897/zookeys.963.53470.figure19" httpUri="https://binary.pensoft.net/fig/444416" pageId="0" pageNumber="45">19</figureCitation>
<figureCitation id="447C9345602F5595FA53C8C94123B817" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Tra-Tra River at 485 m at Wupperthal, Cederberg range, Western Cape Province, South Africa. Locality for Agabus riberae sp. nov. September 28, 2018. Photo Stacey DeAmicis." figureDoi="10.3897/zookeys.963.53470.figure19" httpUri="https://binary.pensoft.net/fig/444416" pageId="0" pageNumber="45">19</figureCitation>
), in either Fynbos or Renosterveld vegetation between 480 and 1000 m elevation. Typically netted from marginal vegetation, including at the base of tussocks. Also found amongst grasses in a spring pool with cold water. Ecological differences between this species and
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
<taxonomicName id="7A11FC97D7750FAD444DA9F85676FE46" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="199A9B64DC12C11A88DA7AF34DA553D2" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. are unclear, but may relate, at least in part, to rainfall.
</paragraph>
<caption doi="10.3897/zookeys.963.53470.figure18" httpUri="https://binary.pensoft.net/fig/444415" pageId="0" pageNumber="45" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 18.</emphasis>
<caption id="62ECA9F335872B17833C2B13233366DB" doi="10.3897/zookeys.963.53470.figure18" httpUri="https://binary.pensoft.net/fig/444415" pageId="0" pageNumber="45" start="Figure 18" startId="F18">
<paragraph id="1A8DCA1BE0BB3D954635A006770AB851" pageId="0" pageNumber="45">
<emphasis id="60FDB76C14609A0F136B777D879F8EA8" bold="true" pageId="0" pageNumber="45">Figure 18.</emphasis>
Stream at 1100 m on Witwater-Langkloof road ca 1 km S. of junction, Kamiesberg, Northern Cape Province, South Africa. Type locality of
<taxonomicName authorityName="Bilton, Englund &amp; Bergsten" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus riberae" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberae">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus riberae</emphasis>
<taxonomicName id="D2DB9D141CD0D3DCC3C65E8EAB3C004A" authorityName="Bilton, Englund &amp; Bergsten" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus riberae" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberae">
<emphasis id="E8ED63CF07388DA48B983E720BAA0A74" italics="true" pageId="0" pageNumber="45">Agabus riberae</emphasis>
</taxonomicName>
sp. nov., with DTB. September 19, 2010. Photo Rebecca Bilton.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">Named after our late friend and colleague Ignacio Ribera, who will be sorely missed.</paragraph>
<subSubSection id="79F8A2E98299D2975292315532998121" pageId="0" pageNumber="45" type="etymology">
<paragraph id="8710542FFE49512464ED1EB56AC6D586" pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph id="4DCE5E82AB80EAEFD273978BA4500809" pageId="0" pageNumber="45">Named after our late friend and colleague Ignacio Ribera, who will be sorely missed.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="comments">
<paragraph pageId="0" pageNumber="45">Comments.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="8F5A9D7DA35490FC0E1CD5502248366F" pageId="0" pageNumber="45" type="comments">
<paragraph id="546336FF55CA178606F65D55A1D11466" pageId="0" pageNumber="45">Comments.</paragraph>
<paragraph id="728F89A6162C7720DFEA08DA7FF04288" pageId="0" pageNumber="45">
Despite the relatively minor morphological differences between this species and
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
<taxonomicName id="F8516BB3FB4F55D4F4B103B03C2B6753" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="3248FEFD0E0E351372453C4CEBB9F824" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov. we consider these two taxa distinct. COI sequences for four specimens of
<taxonomicName lsidName="A. riberae" pageId="0" pageNumber="45" rank="species" species="riberae">
<emphasis italics="true" pageId="0" pageNumber="45">A. riberae</emphasis>
<taxonomicName id="163021E00C7D4BCE20A03574FC6709D8" lsidName="A. riberae" pageId="0" pageNumber="45" rank="species" species="riberae">
<emphasis id="CD712DE376DEA394458715286DEFD0CD" italics="true" pageId="0" pageNumber="45">A. riberae</emphasis>
</taxonomicName>
sp. nov. investigated differed by 4.5-4.7% from
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
<taxonomicName id="A8A69C584E0024557CD9BE1B6550DE94" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="614F4C85EBE67B46CADA450A38EFB5E0" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov., more than that observed between many well-established species in the subgenus
<taxonomicName lsidName="(Acatodes)" pageId="0" pageNumber="45" rank="subGenus" subGenus="Acatodes">Acatodes</taxonomicName>
<taxonomicName id="F6BE4CE8D68BDE7015249EB67880E373" lsidName="(Acatodes)" pageId="0" pageNumber="45" rank="subGenus" subGenus="Acatodes">Acatodes</taxonomicName>
Thomson, 1859 (I. Ribera pers. comm.). This is supportive of the recognition as a distinct taxon, particularly given the relative uniformity in COI sequence observed across the wide geographical range of
<taxonomicName lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
<taxonomicName id="AF0C5EF214BAAEC29985589F7EC0CCE2" lsidName="A. austellus" pageId="0" pageNumber="45" rank="species" species="austellus">
<emphasis id="9DCE793E9C9FC14889A6B77E9A2CAFB5" italics="true" pageId="0" pageNumber="45">A. austellus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>

View file

@ -1,91 +1,91 @@
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<taxonomicName LSID="34A465E2-0048-5754-9298-AAD3C7566C7D" authority="(Straneo, 1943)" authorityName="Will" authorityYear="2020" baseAuthorityName="Straneo" baseAuthorityYear="1943" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia schoutedeni" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="schoutedeni" status="comb. nov.">Tiferonia schoutedeni (Straneo, 1943)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="131">comb. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 69" captionStartId="F4" captionText="Figures 6 - 9. Tiferonia species dorsal habitus. 6 Tiferonia parva, male paratype 7 Tiferonia leytensis male holotype 8 Tiferonia schoutedeni 9 Tiferonia brunnea, male holotype." figureDoi="10.3897/zookeys.906.48255.figures6-9" httpUri="https://binary.pensoft.net/fig/374833" pageId="0" pageNumber="131">Fig. 8</figureCitation>
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<figureCitation id="B5C61C532A6BF3072263102C948F1449" captionStart="Figures 69" captionStartId="F4" captionText="Figures 6 - 9. Tiferonia species dorsal habitus. 6 Tiferonia parva, male paratype 7 Tiferonia leytensis male holotype 8 Tiferonia schoutedeni 9 Tiferonia brunnea, male holotype." figureDoi="10.3897/zookeys.906.48255.figures6-9" httpUri="https://binary.pensoft.net/fig/374833" pageId="0" pageNumber="131">Fig. 8</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="131">
<taxonomicName authorityName="Straneo" authorityYear="1943" class="Insecta" family="Carabidae" genus="Patellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patellus schoutedeni" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="schoutedeni">Patellus schoutedeni</taxonomicName>
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<paragraph id="595F188EE2D1A6F690BEDC94A223A912" pageId="0" pageNumber="131">
<taxonomicName id="FDE6A9669630E45FE8C1F7E012051D79" authorityName="Straneo" authorityYear="1943" class="Insecta" family="Carabidae" genus="Patellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patellus schoutedeni" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="schoutedeni">Patellus schoutedeni</taxonomicName>
Straneo, 1943: 5.
</paragraph>
<paragraph pageId="0" pageNumber="131">
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<paragraph id="D8E514791B5C29D6C192845291D1A178" pageId="0" pageNumber="131">
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:
<bibRefCitation author="Straneo, SL" journalOrPublisher="Revue de Zoologie et de Botanique africaines" pageId="0" pageNumber="131" pagination="45 - 56" refId="B11" refString="Straneo, SL, 1962. Su alcuni Pterostichini entrati recentemente nelle collezioni del Museo dell'Africa Centrale (Coleoptera, Carabidae). Nota III. Revue de Zoologie et de Botanique africaines 66: 45 - 56" title="Su alcuni Pterostichini entrati recentemente nelle collezioni del Museo dell'Africa Centrale (Coleoptera, Carabidae). Nota III." volume="66" year="1962">Straneo 1962</bibRefCitation>
<bibRefCitation id="D29C1F29DA9477DFF795519D67DC80AD" author="Straneo, SL" journalOrPublisher="Revue de Zoologie et de Botanique africaines" pageId="0" pageNumber="131" pagination="45 - 56" refId="B11" refString="Straneo, SL, 1962. Su alcuni Pterostichini entrati recentemente nelle collezioni del Museo dell'Africa Centrale (Coleoptera, Carabidae). Nota III. Revue de Zoologie et de Botanique africaines 66: 45 - 56" title="Su alcuni Pterostichini entrati recentemente nelle collezioni del Museo dell'Africa Centrale (Coleoptera, Carabidae). Nota III." volume="66" year="1962">Straneo 1962</bibRefCitation>
: 54; Lorenz 2005: 328.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="notes">
<paragraph pageId="0" pageNumber="131">Notes.</paragraph>
<paragraph pageId="0" pageNumber="131">
<subSubSection id="92030AB442AC198A8B9AB05498927094" pageId="0" pageNumber="131" type="notes">
<paragraph id="EEDE0C35BBF7316960E33B9E1C66EC47" pageId="0" pageNumber="131">Notes.</paragraph>
<paragraph id="51A783385363FDEE8F0A5958E888B7B5" pageId="0" pageNumber="131">
According to
<bibRefCitation author="Straneo, SL" journalOrPublisher="Revue de Zoologie et de Botanique africaines" pageId="0" pageNumber="131" pagination="1 - 5" refId="B10" refString="Straneo, SL, 1943. Sur quelques nouveaux Pterostichini (Coleopt. Carabid.) d'Afrique. Revue de Zoologie et de Botanique africaines 37: 1 - 5" title="Sur quelques nouveaux Pterostichini (Coleopt. Carabid.) d'Afrique." volume="37" year="1943">Straneo (1943)</bibRefCitation>
<bibRefCitation id="154ACF767AD8D723F459A4F92E58740D" author="Straneo, SL" journalOrPublisher="Revue de Zoologie et de Botanique africaines" pageId="0" pageNumber="131" pagination="1 - 5" refId="B10" refString="Straneo, SL, 1943. Sur quelques nouveaux Pterostichini (Coleopt. Carabid.) d'Afrique. Revue de Zoologie et de Botanique africaines 37: 1 - 5" title="Sur quelques nouveaux Pterostichini (Coleopt. Carabid.) d'Afrique." volume="37" year="1943">Straneo (1943)</bibRefCitation>
the type was intended to be deposited in Tervuren. However, the specimen was not located (S. Hanot, Musee Royal de
<normalizedToken originalValue="lAfrique">l'Afrique</normalizedToken>
<normalizedToken id="F01F06EAE0064EF6F0297C95740BE3AB" originalValue="lAfrique">l'Afrique</normalizedToken>
Centrale, Tervuren, Belgium, in litt.). Straneo stated that the specimen is labelled &quot;Congo Belge, Eala (17-I-1921, Dr. H. Schouteden).&quot;
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="materials_examined">
<paragraph pageId="0" pageNumber="131">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="131">
<subSubSection id="DD5660137058B2D23D6785D1BAF43905" pageId="0" pageNumber="131" type="materials_examined">
<paragraph id="AD337E881CD18D3E4CA480191602D17F" pageId="0" pageNumber="131">Material examined.</paragraph>
<paragraph id="63C471D241DE42CB3955AF26381F6268" pageId="0" pageNumber="131">
Central African Republic • 1♀, //&quot;R[epublic] C[entral] A[frica], P[ark]. N[ational]. [Dzanga-]Ndoki, Camp1 02 28 51.0N 016 13 04.5E, 9-11.II.2012,
<normalizedToken originalValue="piége">piege</normalizedToken>
<normalizedToken id="599781935059FC44CD9CEF8E0F949C69" originalValue="piége">piege</normalizedToken>
UV
<normalizedToken originalValue="canopée">canopee</normalizedToken>
<normalizedToken id="D4D794A8CA778712E7A728F7EEEF61AC" originalValue="canopée">canopee</normalizedToken>
35m, Exp. Sangha 2012, P. Moretto leg. -70-&quot;// [CSCHM] • ♂ //&quot;Bot. N°69 Humus dans
<normalizedToken originalValue="résidu">residu</normalizedToken>
<normalizedToken id="791EC6A3ED15ACD675AB3491DE5753D6" originalValue="résidu">residu</normalizedToken>
forestier&quot;// &quot;I[nstitut pour la]. R[echerche]. S[cientifique en]. A[frique]. C[entrale]. -Mus. Congo Kwango: terr, de feshi, rive dr. Kwenge III-1959 B. 69 Mme J. Leleup&quot;// &quot;
<taxonomicName baseAuthorityName="Straneo" baseAuthorityYear="1943" class="Insecta" family="Carabidae" genus="Melanchrous" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanchrous schoutedeni" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="schoutedeni">
<emphasis italics="true" pageId="0" pageNumber="131">Melanchrous schoutedeni</emphasis>
<taxonomicName id="ADE478E40A55C4E292BE9FE2998DC11D" baseAuthorityName="Straneo" baseAuthorityYear="1943" class="Insecta" family="Carabidae" genus="Melanchrous" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melanchrous schoutedeni" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="schoutedeni">
<emphasis id="8911B3EEC1E8E603CBC88863A9817608" italics="true" pageId="0" pageNumber="131">Melanchrous schoutedeni</emphasis>
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S.L. Straneo det. 1960&quot;//[CMNH].
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</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="diagnosis">
<paragraph pageId="0" pageNumber="131">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="131">
<subSubSection id="3B4E6E17949509F28C26042F548E9F82" pageId="0" pageNumber="131" type="diagnosis">
<paragraph id="5F52576E449AEC7CF1B781D7670357AF" pageId="0" pageNumber="131">Diagnosis.</paragraph>
<paragraph id="0CD5EBB0DFBA6F4F62220014CDE183F8" pageId="0" pageNumber="131">
Having only the first three elytral intervals impressed and a relatively large eyes (gena is only half the width of antennomere 1) distinguishes this species from all other
<taxonomicName authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
<taxonomicName id="037EF29F6B780B450F16E8DC2F837646" authorityName="Darlington" authorityYear="1962" class="Insecta" family="Carabidae" genus="Tiferonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tiferonia" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis id="A4FA9D6E4745DA03F86ECB1F8251B706" italics="true" pageId="0" pageNumber="131">Tiferonia</emphasis>
</taxonomicName>
.
</paragraph>

View file

@ -1,163 +1,163 @@
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<treatment ID-GBIF-Taxon="152048447" LSID="urn:lsid:plazi:treatment:34B3C04FCC08C812E62CDC3CB5026723" httpUri="http://treatment.plazi.org/id/34B3C04FCC08C812E62CDC3CB5026723" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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<taxonomicName LSID="http://species-id.net/wiki/Hymedesmia_rissoi" authority="Topsent, 1936" authorityName="Topsent" authorityYear="1936" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymedesmia (Hymedesmia) rissoi" order="Poecilosclerida" pageId="11" pageNumber="12" phylum="Porifera" rank="species" species="rissoi" subGenus="Hymedesmia">Hymedesmia (Hymedesmia) rissoi Topsent, 1936</taxonomicName>
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Figs 11
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<taxonomicName class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymedesmia gracilisigma" order="Poecilosclerida" pageId="11" pageNumber="12" phylum="Porifera" rank="species" species="gracilisigma">Hymedesmia gracilisigma</taxonomicName>
<subSubSection id="F36A6BCE1515BBB5D0F6D7109DA6DFDE" pageId="11" pageNumber="12" type="reference_group">
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<taxonomicName id="A420D066C25027099BACEE820FD5C850" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymedesmia gracilisigma" order="Poecilosclerida" pageId="11" pageNumber="12" phylum="Porifera" rank="species" species="gracilisigma">Hymedesmia gracilisigma</taxonomicName>
var. rissoi Topsent, 1936: 35.
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<paragraph pageId="11" pageNumber="12">Material examined.</paragraph>
<paragraph lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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<paragraph id="95571E3F990DECE87BDA72511821957E" pageId="11" pageNumber="12">Material examined.</paragraph>
<paragraph id="F9296A82A13A0510B746C903D4F22B59" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Specimen IG-F-BL3-F18b-spA; Specimen IG-F-BL4-sp9-sciaf.; specimen IG-F-BL4 sp11-fot.; alcohol preserved, Gallinara Island (station 2, Falcon
<pageBreakToken pageId="12" pageNumber="13" start="start">ara</pageBreakToken>
<pageBreakToken id="F9476C6CD1A8052D00D4A67C66397004" pageId="12" pageNumber="13" start="start">ara</pageBreakToken>
)
<geoCoordinate direction="north" orientation="latitude" precision="15" value="44.022778">44°01'22&quot;N</geoCoordinate>
<geoCoordinate id="AFFEC9130A65A417B73EA91E58FC8B87" direction="north" orientation="latitude" precision="15" value="44.022778">44°01'22&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="8.22611">8°13'34&quot;E</geoCoordinate>
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, depth 35 m, collected on 17-06-2009; specimenSSS-BL1-sp11-sciaf.; Santo Stefano Shoals, (station 1),
<geoCoordinate direction="north" orientation="latitude" precision="925" value="43.816666">43°49'N</geoCoordinate>
<geoCoordinate id="603D1D0915F03BFD79B5E67B84665FB6" direction="north" orientation="latitude" precision="925" value="43.816666">43°49'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="7.9">7°54'E</geoCoordinate>
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, depth 35 m, collected on 14-02-2008.
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<subSubSection pageId="12" pageNumber="13" type="description">
<paragraph pageId="12" pageNumber="13">Description.</paragraph>
<paragraph pageId="12" pageNumber="13">Small (0.5 cm2), slimy, coriaceous encrusting sponge, grey in colour after alcohol preservation, recorded both on the surface and inside the coralligenous blocks.</paragraph>
<paragraph pageId="12" pageNumber="13">Skeleton. Not observed.</paragraph>
<paragraph pageId="12" pageNumber="13">
<subSubSection id="8ED0D25BBAE8D293118DC07B5BF678B2" pageId="12" pageNumber="13" type="description">
<paragraph id="D54D1507F3953EE787804E8A43F604DE" pageId="12" pageNumber="13">Description.</paragraph>
<paragraph id="1CC8EA7CD63785FD0A7C196083A125AD" pageId="12" pageNumber="13">Small (0.5 cm2), slimy, coriaceous encrusting sponge, grey in colour after alcohol preservation, recorded both on the surface and inside the coralligenous blocks.</paragraph>
<paragraph id="2D54E71CE790781AEEFF87635E5D1AF1" pageId="12" pageNumber="13">Skeleton. Not observed.</paragraph>
<paragraph id="EBB5D7529907A6CCB0076E8E58771F4E" pageId="12" pageNumber="13">
Spicules. Megascleres: straight or slightly sinuous anisotornotes, sometimes modified in anisotylotes or strongyles (Fig. 11A), 140 (175) 177.5
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2.5 (2.7) 3.75
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; acanthostyles in a single size category, 67.5 (84) 105
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2.5 (3.5) 3.75
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="AC313EEDC6EABA08F8B810203461D028" originalValue="μm">μm</normalizedToken>
, devoid of conspicuous heads. The extremities may be pointed or blunt (Figs 11B, C). Microscleres: arcuate isochelae (Fig. 11D), 25 (25.6) 27.5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="CC67C6736F979B5D90721F25EAB49C60" originalValue="μm">μm</normalizedToken>
long; thin sigmas
<normalizedToken originalValue="“C”">&quot;C&quot;</normalizedToken>
<normalizedToken id="0C7C393E5BFFD046DDBD84D70D87CB9A" originalValue="“C”">&quot;C&quot;</normalizedToken>
(Fig. 11E) and
<normalizedToken originalValue="“S”">&quot;S&quot;</normalizedToken>
<normalizedToken id="ED965CAE353E11846C77A4ED1D00958D" originalValue="“S”">&quot;S&quot;</normalizedToken>
shaped, 32.5 (35) 37.5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="E50A8B2EA526EBA5B3072D25978EA76F" originalValue="×">x</normalizedToken>
1.25
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="881F7C7C5446D10DB0E02FCF9C4422E4" originalValue="μm">μm</normalizedToken>
.
</paragraph>
<caption pageId="12" pageNumber="13">
<paragraph pageId="12" pageNumber="13">
<caption id="2687CB88968E4D1EF3F93FB63E68DA4D" pageId="12" pageNumber="13">
<paragraph id="502C41E9987CDECFE0C0B60FBA85C80A" pageId="12" pageNumber="13">
Figure 11.
<taxonomicName class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymedesmia (Hymedesmia) rissoi" order="Poecilosclerida" pageId="12" pageNumber="13" phylum="Porifera" rank="species" species="rissoi" subGenus="Hymedesmia">Hymedesmia (Hymedesmia) rissoi</taxonomicName>
<taxonomicName id="BA5C8AFB9E5A2BB2F423C816FE252373" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hymedesmia (Hymedesmia) rissoi" order="Poecilosclerida" pageId="12" pageNumber="13" phylum="Porifera" rank="species" species="rissoi" subGenus="Hymedesmia">Hymedesmia (Hymedesmia) rissoi</taxonomicName>
. A Tornote, sometimes modified into subtylotes and strongyles B Acanthostyles C Arcuate isochelae D Thin sigmas.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="12" pageNumber="13" type="distribution">
<paragraph pageId="12" pageNumber="13">Distribution and discussion.</paragraph>
<paragraph pageId="12" pageNumber="13">
<subSubSection id="2876C609B7A24413D546D6C34342201B" pageId="12" pageNumber="13" type="distribution">
<paragraph id="CD80126ABF788B32D46758192B7F80A5" pageId="12" pageNumber="13">Distribution and discussion.</paragraph>
<paragraph id="0AD8D28DB051A4DD0DCD7A0041B6D6BB" pageId="12" pageNumber="13">
In the original description
<bibRefCitation author="Topsent, E" journalOrPublisher="Bulletin de l'Institut oceanographique, Monaco" pageId="20" pageNumber="21" pagination="1 - 70" title="Eponges observees dans les parages de Monaco. (Deuxieme partie)." volume="686" year="1936">Topsent (1936)</bibRefCitation>
<bibRefCitation id="84A11DEB74922B0E64E87A7822E6F9A3" author="Topsent, E" journalOrPublisher="Bulletin de l'Institut oceanographique, Monaco" pageId="20" pageNumber="21" pagination="1 - 70" title="Eponges observees dans les parages de Monaco. (Deuxieme partie)." volume="686" year="1936">Topsent (1936)</bibRefCitation>
distinguished in this species two size classes of acanthostyles similar in shape: the larger were 185-265
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="34AFE2977A9479162DF2097955D52AEC" originalValue="μm">μm</normalizedToken>
in length and the smaller 75-115
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="7B9FE0678EA1031E185739C6657EA175" originalValue="μm">μm</normalizedToken>
. Subtylotes straight or sometimes slightly sinuous, 225-275
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9CC231DBE9DCDC7F402D7DD673249654" originalValue="×">x</normalizedToken>
3.5-4.5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="042BC840101D44EB950BD4292D762918" originalValue="μm">μm</normalizedToken>
, arcuate isochelae 23-25
<normalizedToken originalValue="μm">μm</normalizedToken>
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long and sigmas 40-50
<normalizedToken originalValue="μm">μm</normalizedToken>
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long and less than 1
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="B3F9F32B24274E0666F364AC45A53352" originalValue="μm">μm</normalizedToken>
thick. The specimens here described match with
<normalizedToken originalValue="Topsents">Topsent's</normalizedToken>
<normalizedToken id="312D26BA9EF4F46216ECD458DCDDC6A3" originalValue="Topsents">Topsent's</normalizedToken>
description apart from the presence of a single size class of acanthostyles. However, other authors (
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<normalizedToken originalValue="Sarà">Sara</normalizedToken>
<bibRefCitation id="60DCBEBD591E58A8CCABB22313A6805D" author="Sara, M" journalOrPublisher="Annuario dell'Istituto e Museo di Zoologia dell'Universita di Napoli" pageId="19" pageNumber="20" pagination="1 - 62" title="La fauna di Poriferi delle ' secche' del golfo di Napoli. II. La secca di Benda Palummo." volume="14" year="1962">
<normalizedToken id="87547BA9591C5FF747339A29E7043A31" originalValue="Sarà">Sara</normalizedToken>
and Siribelli 1962
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), recorded a single class of acanthostyles as well. This is a Mediterranean endemic species (Ligurian Sea and Central Tyrrhenian Sea). It was found on
<taxonomicName class="Anthozoa" genus="Cladocora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladocora caespitosa" order="Scleractinia" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="caespitosa">Cladocora caespitosa</taxonomicName>
<taxonomicName id="C63A588F86E0164692128E2006FD4C61" class="Anthozoa" family="Cladocoridae" genus="Cladocora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladocora caespitosa" order="Scleractinia" pageId="12" pageNumber="13" phylum="Cnidaria" rank="species" species="caespitosa">Cladocora caespitosa</taxonomicName>
, at 15-40 m depth (
<bibRefCitation author="Topsent, E" journalOrPublisher="Bulletin de l'Institut oceanographique, Monaco" pageId="20" pageNumber="21" pagination="1 - 70" title="Eponges observees dans les parages de Monaco. (Deuxieme partie)." volume="686" year="1936">Topsent 1936</bibRefCitation>
<bibRefCitation id="EF9E94A8284116199DE3F6355BBD71E6" author="Topsent, E" journalOrPublisher="Bulletin de l'Institut oceanographique, Monaco" pageId="20" pageNumber="21" pagination="1 - 70" title="Eponges observees dans les parages de Monaco. (Deuxieme partie)." volume="686" year="1936">Topsent 1936</bibRefCitation>
) and on coralligenous bottom, at 40-70 m depth (
<bibRefCitation author="Sara, M" journalOrPublisher="Annuario dell'Istituto e Museo di Zoologia dell'Universita di Napoli" pageId="19" pageNumber="20" pagination="1 - 62" title="La fauna di Poriferi delle ' secche' del golfo di Napoli. II. La secca di Benda Palummo." volume="14" year="1962">
<normalizedToken originalValue="Sarà">Sara</normalizedToken>
<bibRefCitation id="6959713D1F94EBE3E29FDC55D5420AC6" author="Sara, M" journalOrPublisher="Annuario dell'Istituto e Museo di Zoologia dell'Universita di Napoli" pageId="19" pageNumber="20" pagination="1 - 62" title="La fauna di Poriferi delle ' secche' del golfo di Napoli. II. La secca di Benda Palummo." volume="14" year="1962">
<normalizedToken id="6D3729E122AC035D65A66FCDD50EB4F4" originalValue="Sarà">Sara</normalizedToken>
and Siribelli 1962
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).

View file

@ -1,138 +1,138 @@
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Subgenus
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Adelung, 1902.
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Annuaire du
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Zoologique de
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des Sciences de St. Petersburg
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7: 62; Kirby 1906. A Synonymic Catalogue of
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2: 125.
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<taxonomicName authority="" class="Insecta" family="Rhaphidophoridae" genus="Diestrammena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diestrammena (Gymnaeta)" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="subGenus" subGenus="Gymnaeta">Diestrammena (Gymnaeta)</taxonomicName>
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<taxonomicName id="963D81FE7DE388BE2F54BF1BFC0241BB" authority="" class="Insecta" family="Rhaphidophoridae" genus="Diestrammena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diestrammena (Gymnaeta)" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="subGenus" subGenus="Gymnaeta">Diestrammena (Gymnaeta)</taxonomicName>
: Jacobson, 1905[1902-1905]. In: Jacobson and Bianchi,
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<emphasis id="889D9665D49392C4E417471E1477F535" italics="true" pageId="0" pageNumber="115">Orthopteroid and Pseudoneuropteroid Insects of Russian Empire and adjacent countries</emphasis>
, 329, 352, 434;
<bibRefCitation author="Gorochov, AV" journalOrPublisher="Proceedings of the Zoological Institute of the Russian Academy of Sciences" pageId="0" pageNumber="115" pagination="37 - 50" refId="B6" refString="Gorochov, AV, 1994. News of systematics and faunistics of Vietnam insects Part 4. Proceedings of the Zoological Institute of the Russian Academy of Sciences 257: 37 - 50" title="News of systematics and faunistics of Vietnam insects Part 4." volume="257" year="1994">Gorochov 1994</bibRefCitation>
<bibRefCitation id="8977E40B94292A3C9F1EBAE3731B696D" author="Gorochov, AV" journalOrPublisher="Proceedings of the Zoological Institute of the Russian Academy of Sciences" pageId="0" pageNumber="115" pagination="37 - 50" refId="B6" refString="Gorochov, AV, 1994. News of systematics and faunistics of Vietnam insects Part 4. Proceedings of the Zoological Institute of the Russian Academy of Sciences 257: 37 - 50" title="News of systematics and faunistics of Vietnam insects Part 4." volume="257" year="1994">Gorochov 1994</bibRefCitation>
. In: Gorochov and Kireichuk [Eds.],
<emphasis italics="true" pageId="0" pageNumber="115">Proceedings of the Zoological Institute of the Russian Academy of Sciences</emphasis>
<emphasis id="C5CE06EF8164BAE679D5A4638F24D362" italics="true" pageId="0" pageNumber="115">Proceedings of the Zoological Institute of the Russian Academy of Sciences</emphasis>
, 257: 49; Otte 2000.
<taxonomicName authorityName="Brunner von Wattenwyl" authorityYear="1888" class="Insecta" family="Rhaphidophoridae" genus="Diestrammena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
<taxonomicName id="FEBA4A2982006E610056DCF18E894B04" authorityName="Brunner von Wattenwyl" authorityYear="1888" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
Species File 8: 55;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.1917.1.4" author="Jiao, ZJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="55 - 60" refId="B12" refString="Jiao, ZJ, Niu, CY, Liu, XW, Lei, CL, Bi, WX, 2008. Descriptions of Chinese species of the subgenus Diestrammena (Gymnaeta) Adelung (Orthoptera: Rhaphidophoridae). Zootaxa 1917 (1): 55 - 60, DOI: https://doi.org/10.11646/zootaxa.1917.1.4" title="Descriptions of Chinese species of the subgenus Diestrammena (Gymnaeta) Adelung (Orthoptera: Rhaphidophoridae)." url="https://doi.org/10.11646/zootaxa.1917.1.4" volume="1917" year="2008">Jiao et al. 2008</bibRefCitation>
<bibRefCitation id="A6845146C57BFB5924535F581A82A8F2" DOI="https://doi.org/10.11646/zootaxa.1917.1.4" author="Jiao, ZJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="55 - 60" refId="B12" refString="Jiao, ZJ, Niu, CY, Liu, XW, Lei, CL, Bi, WX, 2008. Descriptions of Chinese species of the subgenus Diestrammena (Gymnaeta) Adelung (Orthoptera: Rhaphidophoridae). Zootaxa 1917 (1): 55 - 60, DOI: https://doi.org/10.11646/zootaxa.1917.1.4" title="Descriptions of Chinese species of the subgenus Diestrammena (Gymnaeta) Adelung (Orthoptera: Rhaphidophoridae)." url="https://doi.org/10.11646/zootaxa.1917.1.4" volume="1917" year="2008">Jiao et al. 2008</bibRefCitation>
.
<emphasis italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
<emphasis id="BA4BA83F7AAF0D176D0787CFD73D97F9" italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
1917: 55;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2272.1.2" author="Zhang, F" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="21 - 36" refId="B31" refString="Zhang, F, Liu, XW, 2009. A review of the subgenus Diestrammena (Gymnaeta) from China (Orthoptera: Rhaphidophoridae Aemodogryllinae). Zootaxa 2272 (1): 21 - 36, DOI: https://doi.org/10.11646/zootaxa.2272.1.2" title="A review of the subgenus Diestrammena (Gymnaeta) from China (Orthoptera: Rhaphidophoridae Aemodogryllinae)." url="https://doi.org/10.11646/zootaxa.2272.1.2" volume="2272" year="2009">Zhang and Liu 2009</bibRefCitation>
<bibRefCitation id="454D84EBDD6C4DBE89D27101F47AAD68" DOI="https://doi.org/10.11646/zootaxa.2272.1.2" author="Zhang, F" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="21 - 36" refId="B31" refString="Zhang, F, Liu, XW, 2009. A review of the subgenus Diestrammena (Gymnaeta) from China (Orthoptera: Rhaphidophoridae Aemodogryllinae). Zootaxa 2272 (1): 21 - 36, DOI: https://doi.org/10.11646/zootaxa.2272.1.2" title="A review of the subgenus Diestrammena (Gymnaeta) from China (Orthoptera: Rhaphidophoridae Aemodogryllinae)." url="https://doi.org/10.11646/zootaxa.2272.1.2" volume="2272" year="2009">Zhang and Liu 2009</bibRefCitation>
.
<emphasis italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
<emphasis id="1C9B5E452131AF52166DD22BDB94F5A8" italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
2272: 21.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName authority="" authorityName="Adelung" authorityYear="1902" class="Insecta" family="Rhaphidophoridae" genus="Tachycines" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachycines (Gymnaeta)" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="subGenus" subGenus="Gymnaeta">Tachycines (Gymnaeta)</taxonomicName>
<paragraph id="279FA1C4391AEB98FA33C68EEDB268F2" pageId="0" pageNumber="115">
<taxonomicName id="236C6D5E70447B2E9DA2135EA893A1B4" authority="" authorityName="Adelung" authorityYear="1902" class="Insecta" family="Rhaphidophoridae" genus="Tachycines" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tachycines (Gymnaeta)" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="subGenus" subGenus="Gymnaeta">Tachycines (Gymnaeta)</taxonomicName>
: Karny, 1934.
<taxonomicName authorityName="Brauns" authorityYear="1884" class="Insecta" family="Rhaphidophoridae" genus="Konowia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Konowia" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">Konowia</taxonomicName>
<taxonomicName id="7D639E7CA7B41131ED838E9131226106" authorityName="Brauns" authorityYear="1884" class="Insecta" family="Rhaphidophoridae" genus="Konowia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Konowia" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">Konowia</taxonomicName>
13 (1-3): 218; Karny 1937.
<emphasis italics="true" pageId="0" pageNumber="115">Genera Insectorum</emphasis>
<emphasis id="D401F3D297567DBB38BF8A62B37FADF8" italics="true" pageId="0" pageNumber="115">Genera Insectorum</emphasis>
206: 248; Storozhenko 1990.
<emphasis italics="true" pageId="0" pageNumber="115">Entomologicheskoe Obozrenie</emphasis>
<emphasis id="97070FB2B9A6DC04E4EE434E3A875F55" italics="true" pageId="0" pageNumber="115">Entomologicheskoe Obozrenie</emphasis>
69(4): 845, 847; Qin et al. 2018.
<emphasis italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
<emphasis id="9ACF2A259C3DEE7C26AB75E2AE3689BC" italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
4374(4): 452;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4560.2.3" author="Qin, YY" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="273 - 310" refId="B20" refString="Qin, YY, Liu, XW, Li, K, 2019. Review of the subgenus Tachycines (Gymnaeta) Adelung, 1902 (Orthoptera, Rhaphidophoridae, Aemodogryllinae, Aemodogryllini). Zootaxa 4560 (2): 273 - 310, DOI: https://doi.org/10.11646/zootaxa.4560.2.3" title="Review of the subgenus Tachycines (Gymnaeta) Adelung, 1902 (Orthoptera, Rhaphidophoridae, Aemodogryllinae, Aemodogryllini)." url="https://doi.org/10.11646/zootaxa.4560.2.3" volume="4560" year="2019">Qin et al. 2019</bibRefCitation>
<bibRefCitation id="01090A09C53EF64593E449294CEA9560" DOI="https://doi.org/10.11646/zootaxa.4560.2.3" author="Qin, YY" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="273 - 310" refId="B20" refString="Qin, YY, Liu, XW, Li, K, 2019. Review of the subgenus Tachycines (Gymnaeta) Adelung, 1902 (Orthoptera, Rhaphidophoridae, Aemodogryllinae, Aemodogryllini). Zootaxa 4560 (2): 273 - 310, DOI: https://doi.org/10.11646/zootaxa.4560.2.3" title="Review of the subgenus Tachycines (Gymnaeta) Adelung, 1902 (Orthoptera, Rhaphidophoridae, Aemodogryllinae, Aemodogryllini)." url="https://doi.org/10.11646/zootaxa.4560.2.3" volume="4560" year="2019">Qin et al. 2019</bibRefCitation>
.
<emphasis italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
<emphasis id="BD5E9AD75E2F99A80DC90B10549AC304" italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
4560(2): 274;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4674.4.8" author="Feng, XL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="491 - 495" refId="B4" refString="Feng, XL, Huang, SH, Luo, CQ, 2019. A new species of the subgenus Tachycines (Gymnaeta) (Orthoptera: Rhaphidophoridae) from karst caves of southern Guizhou, China. Zootaxa 4674 (4): 491 - 495, DOI: https://doi.org/10.11646/zootaxa.4674.4.8" title="A new species of the subgenus Tachycines (Gymnaeta) (Orthoptera: Rhaphidophoridae) from karst caves of southern Guizhou, China." url="https://doi.org/10.11646/zootaxa.4674.4.8" volume="4674" year="2019">Feng et al. 2019</bibRefCitation>
<bibRefCitation id="994846C33CF76BE31B262CBDC51ECEB4" DOI="https://doi.org/10.11646/zootaxa.4674.4.8" author="Feng, XL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="491 - 495" refId="B4" refString="Feng, XL, Huang, SH, Luo, CQ, 2019. A new species of the subgenus Tachycines (Gymnaeta) (Orthoptera: Rhaphidophoridae) from karst caves of southern Guizhou, China. Zootaxa 4674 (4): 491 - 495, DOI: https://doi.org/10.11646/zootaxa.4674.4.8" title="A new species of the subgenus Tachycines (Gymnaeta) (Orthoptera: Rhaphidophoridae) from karst caves of southern Guizhou, China." url="https://doi.org/10.11646/zootaxa.4674.4.8" volume="4674" year="2019">Feng et al. 2019</bibRefCitation>
.
<emphasis italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
<emphasis id="6A45DF0885BDB45CACD9BCE8D037DBA6" italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
4674(4): 492;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.937.49173" author="Zhou, XL" journalOrPublisher="ZooKeys" pageId="0" pageNumber="115" pagination="21 - 29" refId="B32" refString="Zhou, XL, Yang, WC, 2020. A new species of Tachycines Adelung, 1902 (Orthoptera, Rhaphidophoridae, Aemodogryllinae, Aemodogryllini) from karst caves in Guizhou, China. ZooKeys 937: 21 - 29, DOI: https://doi.org/10.3897/zookeys.937.49173" title="A new species of Tachycines Adelung, 1902 (Orthoptera, Rhaphidophoridae, Aemodogryllinae, Aemodogryllini) from karst caves in Guizhou, China." url="https://doi.org/10.3897/zookeys.937.49173" volume="937" year="2020">Zhou and Yang 2020</bibRefCitation>
<bibRefCitation id="0C7D0940DC793893BC8DAA9F494EF948" DOI="https://doi.org/10.3897/zookeys.937.49173" author="Zhou, XL" journalOrPublisher="ZooKeys" pageId="0" pageNumber="115" pagination="21 - 29" refId="B32" refString="Zhou, XL, Yang, WC, 2020. A new species of Tachycines Adelung, 1902 (Orthoptera, Rhaphidophoridae, Aemodogryllinae, Aemodogryllini) from karst caves in Guizhou, China. ZooKeys 937: 21 - 29, DOI: https://doi.org/10.3897/zookeys.937.49173" title="A new species of Tachycines Adelung, 1902 (Orthoptera, Rhaphidophoridae, Aemodogryllinae, Aemodogryllini) from karst caves in Guizhou, China." url="https://doi.org/10.3897/zookeys.937.49173" volume="937" year="2020">Zhou and Yang 2020</bibRefCitation>
.
<emphasis italics="true" pageId="0" pageNumber="115">ZooKeys</emphasis>
<emphasis id="320469E865430F81580D3035F30CC8CE" italics="true" pageId="0" pageNumber="115">ZooKeys</emphasis>
937: 21-29;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4809.1.4" author="Zhu, QD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="71 - 94" refId="B34" refString="Zhu, QD, Chen, HM, Shi, FM, 2020. Remarks on the genus Tachycines Adelung, 1902 (Orthoptera: Rhaphidophoridae: Aemodogryllinae) with description of eight new species from caves in southern China. Zootaxa 4809 (1): 71 - 94, DOI: https://doi.org/10.11646/zootaxa.4809.1.4" title="Remarks on the genus Tachycines Adelung, 1902 (Orthoptera: Rhaphidophoridae: Aemodogryllinae) with description of eight new species from caves in southern China." url="https://doi.org/10.11646/zootaxa.4809.1.4" volume="4809" year="2020">Zhu et al. 2020</bibRefCitation>
<bibRefCitation id="1D4EED59E2BEF055D5E743555D54A9C1" DOI="https://doi.org/10.11646/zootaxa.4809.1.4" author="Zhu, QD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="115" pagination="71 - 94" refId="B34" refString="Zhu, QD, Chen, HM, Shi, FM, 2020. Remarks on the genus Tachycines Adelung, 1902 (Orthoptera: Rhaphidophoridae: Aemodogryllinae) with description of eight new species from caves in southern China. Zootaxa 4809 (1): 71 - 94, DOI: https://doi.org/10.11646/zootaxa.4809.1.4" title="Remarks on the genus Tachycines Adelung, 1902 (Orthoptera: Rhaphidophoridae: Aemodogryllinae) with description of eight new species from caves in southern China." url="https://doi.org/10.11646/zootaxa.4809.1.4" volume="4809" year="2020">Zhu et al. 2020</bibRefCitation>
.
<emphasis italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
<emphasis id="EB8FA1E21A2401AB2226CC7522405B60" italics="true" pageId="0" pageNumber="115">Zootaxa</emphasis>
4809(1): 72.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="type species">
<paragraph pageId="0" pageNumber="115">Type species.</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName baseAuthorityName="Adelung" baseAuthorityYear="1902" class="Insecta" family="Rhaphidophoridae" genus="Gymnaeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gymnaeta berezovskii" order="Saltatoria" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="berezovskii">
<emphasis italics="true" pageId="0" pageNumber="115">Gymnaeta berezovskii</emphasis>
<subSubSection id="2A068ED544D2E9B44C64B520B5132FD3" pageId="0" pageNumber="115" type="type species">
<paragraph id="0F7ED59ABF53C2EEC27391D4A263C58D" pageId="0" pageNumber="115">Type species.</paragraph>
<paragraph id="D97A3333565BA468ED66C96614D5BA4C" pageId="0" pageNumber="115">
<taxonomicName id="0007B2BFE0E4717EE5B982A548AFDFDA" baseAuthorityName="Adelung" baseAuthorityYear="1902" family="Rhaphidophoridae" genus="Gymnaeta" kingdom="Animalia" lsidName="Gymnaeta berezovskii" order="Saltatoria" pageId="0" pageNumber="115" rank="species" species="berezovskii">
<emphasis id="7ADDA9BA8A1A9FA038B4A138AE30F29D" italics="true" pageId="0" pageNumber="115">Gymnaeta berezovskii</emphasis>
</taxonomicName>
Adelung, by subsequent designation; authority: Kirby, W.F. 1906. A Synonymic Catalogue of
<taxonomicName authorityName="Adelung" authorityYear="1902" class="Insecta" family="Rhaphidophoridae" genus="Gymnaeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
<taxonomicName id="C0CC7CBA05D1BB3960D27C3CBC9D9632" authorityName="Adelung" authorityYear="1902" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
(
<taxonomicName authorityName="Adelung" authorityYear="1902" class="Insecta" family="Rhaphidophoridae" genus="Gymnaeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
<taxonomicName authorityName="Adelung" authorityYear="1902" class="Insecta" family="Rhaphidophoridae" genus="Gymnaeta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Saltatoria" pageId="0" pageNumber="115" phylum="Arthropoda" rank="order">Saltatoria</taxonomicName>
<taxonomicName id="6415EF092BCB266075D5F6E8D787F9B4" authorityName="Adelung" authorityYear="1902" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
<taxonomicName id="CB63620C12DB1BAA301C585C75CA391A" authorityName="Adelung" authorityYear="1902" kingdom="Animalia" lsidName="" order="Saltatoria" pageId="0" pageNumber="115" rank="order">Saltatoria</taxonomicName>
,
<taxonomicName class="Insecta" family="Locustidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="family">Locustidae</taxonomicName>
<taxonomicName id="C5C38ACF0107C7A3501404B7DC51BD1A" class="Insecta" family="Locustidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="family">Locustidae</taxonomicName>
vel
<taxonomicName class="Insecta" family="Acridiidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="family">Acridiidae</taxonomicName>
<taxonomicName id="D019C8B232F5D5C101A2A4C6E3910C86" class="Insecta" family="Acridiidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="family">Acridiidae</taxonomicName>
) 2: i-viii, 1-562.
</paragraph>
</subSubSection>

View file

@ -1,204 +1,204 @@
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<mods:namePart id="6624C3044EC4C8FC4D276DF28835AC78">Sutcharit, Chirasak</mods:namePart>
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<treatment ID-GBIF-Taxon="183203211" LSID="urn:lsid:plazi:treatment:34F9EB4CC4FD593E9C6E7E720F72E575" httpUri="http://treatment.plazi.org/id/34F9EB4CC4FD593E9C6E7E720F72E575" lastPageNumber="101" pageId="0" pageNumber="101">
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<taxonomicName LSID="34F9EB4C-C4FD-593E-9C6E-7E720F72E575" authority="Maassen, 2006" authorityName="Maassen" authorityYear="2006" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemiplecta esculenta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="esculenta">Hemiplecta esculenta Maassen, 2006</taxonomicName>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Living snail. A Hemiplecta humphreysiana from Singapore (width about 45 mm) B, C Hemiplecta distincta B from Saraburi, Thailand (width about 65 mm) and C mating pairs D, E Hemiplecta funerea from Nan, Thailand (width about 50 mm) D yellow shell form and E dark shell form (width about 50 mm) F Hemiplecta esculenta from Chiang Mai, Thailand (width about 30 mm)." figureDoi="10.3897/zookeys.1047.65735.figure1" httpUri="https://binary.pensoft.net/fig/560488" pageId="0" pageNumber="101">Figures 1F</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">, 7</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Representative SEM images of radula A-C Hemiplecta humphreysiana, specimen CUMZ 4573 from Singapore A central and lateral teeth B transition from lateral teeth to marginal teeth and C outermost marginal teeth D, E Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand D central and lateral teeth and E outermost marginal teeth F, G Hemiplecta funerea, specimen CUMZ 4575 from Nan, Thailand F central and lateral teeth and G outermost marginal teeth H, I Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand H central and lateral teeth I transition from lateral teeth to marginal teeth J Hemiplecta nemorosa sp. nov., paratype CUMZ 5253 from Maehongsorn, Thailand. Central tooth indicated by ' C'. Numbers indicate the tooth order from lateral to marginal end." figureDoi="10.3897/zookeys.1047.65735.figure10" httpUri="https://binary.pensoft.net/fig/560497" pageId="0" pageNumber="101">, 10H, I</figureCitation>
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<figureCitation id="69FE13DA8DB4B6CC0121CF72D947AB9F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">, 7</figureCitation>
<figureCitation id="7990C4554119BADABD4FFBB94B5805CE" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Representative SEM images of radula A-C Hemiplecta humphreysiana, specimen CUMZ 4573 from Singapore A central and lateral teeth B transition from lateral teeth to marginal teeth and C outermost marginal teeth D, E Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand D central and lateral teeth and E outermost marginal teeth F, G Hemiplecta funerea, specimen CUMZ 4575 from Nan, Thailand F central and lateral teeth and G outermost marginal teeth H, I Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand H central and lateral teeth I transition from lateral teeth to marginal teeth J Hemiplecta nemorosa sp. nov., paratype CUMZ 5253 from Maehongsorn, Thailand. Central tooth indicated by ' C'. Numbers indicate the tooth order from lateral to marginal end." figureDoi="10.3897/zookeys.1047.65735.figure10" httpUri="https://binary.pensoft.net/fig/560497" pageId="0" pageNumber="101">, 10H, I</figureCitation>
</paragraph>
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<taxonomicName id="52038FA26B509DCFCCE69CEEA1D02CAD" authorityName="Maassen" authorityYear="2006" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemiplecta esculenta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="esculenta">Hemiplecta esculenta</taxonomicName>
Maassen, 2006: 17, 18, figs 10-12. Type locality: limestone area near village Hang, Pu Luong National Park, Thanh Hoa, Vietnam.
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="101" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
<bibRefCitation id="7AB935EE36A2444DCB444ECFD8DC9482" DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="101" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
: 76, fig. 35d, e.
<bibRefCitation DOI="https://doi.org/10.12657/folmal.027.007" author="Pall-Gergely, B" journalOrPublisher="Folia Malacologica" pageId="0" pageNumber="101" pagination="85 - 89" refId="B79" refString="Pall-Gergely, B, 2019. A new species of Hemiplecta Albers, 1850 from Vietnam (Gastropoda: Pulmonata: Ariophantidae). Folia Malacologica 27: 85 - 89, DOI: https://doi.org/10.12657/folmal.027.007" title="A new species of Hemiplecta Albers, 1850 from Vietnam (Gastropoda: Pulmonata: Ariophantidae)." url="https://doi.org/10.12657/folmal.027.007" volume="27" year="2019">
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2019
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: figs 11-13.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="type specimen">
<paragraph pageId="0" pageNumber="101">Type specimen.</paragraph>
<paragraph pageId="0" pageNumber="101">See the species list of Indochinese species.</paragraph>
<subSubSection id="086B3FD26BE26B30C91FA6DA164789B9" pageId="0" pageNumber="101" type="type specimen">
<paragraph id="3451EC804BD217008ABC6CEC29467603" pageId="0" pageNumber="101">Type specimen.</paragraph>
<paragraph id="7476A65FEB17F8B4C7C9BC28CE1F42B2" pageId="0" pageNumber="101">See the species list of Indochinese species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="materials_examined">
<paragraph pageId="0" pageNumber="101">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="101">
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Thailand" location="Tam Chiang Dao" specimenCount="42" stateProvince="Chiang Mai">
<emphasis bold="true" pageId="0" pageNumber="101">
<collectingCountry name="Thailand">Thailand</collectingCountry>
<subSubSection id="45AC9FEC51AD4E66FB4258BFF942EB14" pageId="0" pageNumber="101" type="materials_examined">
<paragraph id="EAB0DC88D2730D8E0472842240EA9B7A" pageId="0" pageNumber="101">Material examined.</paragraph>
<paragraph id="20A857DFC403279ECC13B66800EB1BCF" pageId="0" pageNumber="101">
<materialsCitation id="C02B900EDE787E7ADC1EFB180145839A" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Thailand" location="Tam Chiang Dao" specimenCount="42" stateProvince="Chiang Mai">
<emphasis id="53073E3A9E1207A363AB5FF5C31CBA33" bold="true" pageId="0" pageNumber="101">
<collectingCountry id="C5AC53514DC9821E22DB5DECD8EE21B0" name="Thailand">Thailand</collectingCountry>
</emphasis>
:
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,
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: CUMZ 4553 (
<specimenCount type="generic">5 specimens</specimenCount>
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in ethanol; Fig.
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<figureCitation id="0440AFE007C8B4F3012D6D918677AE9A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Living snail. A Hemiplecta humphreysiana from Singapore (width about 45 mm) B, C Hemiplecta distincta B from Saraburi, Thailand (width about 65 mm) and C mating pairs D, E Hemiplecta funerea from Nan, Thailand (width about 50 mm) D yellow shell form and E dark shell form (width about 50 mm) F Hemiplecta esculenta from Chiang Mai, Thailand (width about 30 mm)." figureDoi="10.3897/zookeys.1047.65735.figure1" httpUri="https://binary.pensoft.net/fig/560488" pageId="0" pageNumber="101">1F</figureCitation>
), CUMZ 4564 (
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in ethanol), CUMZ 4565 (
<specimenCount type="generic">9 specimens</specimenCount>
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in ethanol), CUMZ 4574 (
<specimenCount type="generic">8 specimens</specimenCount>
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in ethanol; Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, B Hemiplecta funerea, specimens CUMZ 4575 from Nan, Thailand A yellow shell form and B dark shell form C Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand." figureDoi="10.3897/zookeys.1047.65735.figure5" httpUri="https://binary.pensoft.net/fig/560492" pageId="0" pageNumber="101">5C</figureCitation>
<figureCitation id="A3FFEBE7B8DDD308B01F8F7EE84840E6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, B Hemiplecta funerea, specimens CUMZ 4575 from Nan, Thailand A yellow shell form and B dark shell form C Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand." figureDoi="10.3897/zookeys.1047.65735.figure5" httpUri="https://binary.pensoft.net/fig/560492" pageId="0" pageNumber="101">5C</figureCitation>
); Tam Tab-Tao, Chaiprakarn,
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
<collectingRegion id="656DFF61D975AD58FD76757DDDFDB511" country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
: CUMZ 4580 (
<specimenCount type="generic">10 specimens</specimenCount>
<specimenCount id="C68363C1F6205BFDEC5153EED43D819B" type="generic">10 specimens</specimenCount>
in ethanol)
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.
</paragraph>
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<emphasis bold="true" italics="true" pageId="0" pageNumber="101">Shell.</emphasis>
<paragraph id="B6C62921CC63CB04B0D41247840312E2" pageId="0" pageNumber="101">
<emphasis id="061BD5EF497FC78D458B6E5317A2D016" bold="true" italics="true" pageId="0" pageNumber="101">Shell.</emphasis>
Shell relatively small (height up to
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<quantity id="46ED073E63E08133C34E25EE91BC8B19" metricMagnitude="-2" metricUnit="m" metricValue="2.5" unit="mm" value="25.0">25 mm</quantity>
, width up to
<quantity metricMagnitude="-2" metricUnit="m" metricValue="3.5" unit="mm" value="35.0">35 mm</quantity>
<quantity id="776CB983C95DB9B41F005726ACCAE0E6" metricMagnitude="-2" metricUnit="m" metricValue="3.5" unit="mm" value="35.0">35 mm</quantity>
), elevated to slightly depressed, upper surface with distinct nodules arranged on growth line, and lower shell surface nearly smooth. Last whorl keeled; aperture large and ovate; lip simple to slightly expanded and dark brown. Umbilicus widely opened and deep (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, B Hemiplecta funerea, specimens CUMZ 4575 from Nan, Thailand A yellow shell form and B dark shell form C Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand." figureDoi="10.3897/zookeys.1047.65735.figure5" httpUri="https://binary.pensoft.net/fig/560492" pageId="0" pageNumber="101">5C</figureCitation>
<figureCitation id="ED3EB8ED68703BCB9CE14BF586D39314" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A, B Hemiplecta funerea, specimens CUMZ 4575 from Nan, Thailand A yellow shell form and B dark shell form C Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand." figureDoi="10.3897/zookeys.1047.65735.figure5" httpUri="https://binary.pensoft.net/fig/560492" pageId="0" pageNumber="101">5C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">Genitalia.</emphasis>
<paragraph id="A9D4585CF0D6EC8F1D4538FE681E02FD" pageId="0" pageNumber="101">
<emphasis id="8D030BEE2C1036B96EA4F023D82ACF1A" bold="true" italics="true" pageId="0" pageNumber="101">Genitalia.</emphasis>
Genital tracts similar to those of
<taxonomicName higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. humphreysiana" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="humphreysiana">
<emphasis italics="true" pageId="0" pageNumber="101">H. humphreysiana</emphasis>
<taxonomicName id="4E7759693E6A3AD5F816981FFE918D44" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. humphreysiana" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="humphreysiana">
<emphasis id="4D06218537DCE8BA60751C54D914AE2C" italics="true" pageId="0" pageNumber="101">H. humphreysiana</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">7A</figureCitation>
<figureCitation id="F1979B2D102BD5DAA41FAFF0784AE0FE" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">7A</figureCitation>
). Internal wall of penis with sculpture encircling penial verge. Penial sculpture (ps) consists of small to large papillary knobs arranged in oblique lines on penial wall; relatively smaller knobs surrounding penial verge. Penial verge (pv) small, short, conic, and with smooth surface (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">7B</figureCitation>
<figureCitation id="07F552C0AD8E7E540028D55DC8581C0A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">7B</figureCitation>
). Gametolytic sac (gs) bulbous with undifferentiated duct. Internal wall of vagina with series of thin longitudinal vaginal pilasters (vp). Dart apparatus (da) relatively short; internal wall of chamber with smooth wall, and papilla of dart apparatus (dp) slightly elongate, conic, and with smooth surface (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">7B</figureCitation>
<figureCitation id="6346B4505D0B5FED7DABF40984279B38" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">7B</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" pageId="0" pageNumber="101">Figure 7.</emphasis>
<caption id="C7B73097F6BEB5C68036F869CA030D8C" doi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101" start="Figure 7" startId="F7">
<paragraph id="C7B7DC3BAB0226F606C2A0764D268B35" pageId="0" pageNumber="101">
<emphasis id="92666D5F314C31274F5BFA8FFEE703A7" bold="true" pageId="0" pageNumber="101">Figure 7.</emphasis>
Genitalia, mantle edge structure, and caudal region of
<taxonomicName authorityName="Maassen" authorityYear="2006" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" lsidName="Hemiplecta esculenta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="esculenta">
<emphasis italics="true" pageId="0" pageNumber="101">Hemiplecta esculenta</emphasis>
<taxonomicName id="44F4AF8547A60515EC5F001C5E6402C6" authorityName="Maassen" authorityYear="2006" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" lsidName="Hemiplecta esculenta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="esculenta">
<emphasis id="F72DD7BE5FF3EC49A46A370B31DCFEBE" italics="true" pageId="0" pageNumber="101">Hemiplecta esculenta</emphasis>
</taxonomicName>
, specimen CUMZ 4553 from
<collectingRegion country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
<collectingRegion id="2ECEB884A5381C91EA6E8BB261DCBF7D" country="Thailand" name="Chiang Mai">Chiang Mai</collectingRegion>
,
<collectingCountry name="Thailand">Thailand</collectingCountry>
<emphasis bold="true" pageId="0" pageNumber="101">A</emphasis>
<collectingCountry id="27AC3D29D55257241EBAC4145F8A3107" name="Thailand">Thailand</collectingCountry>
<emphasis id="2FD408D0C0EEE57305198CE2F947BB63" bold="true" pageId="0" pageNumber="101">A</emphasis>
whole genital organ
<emphasis bold="true" pageId="0" pageNumber="101">B</emphasis>
<emphasis id="E462D7102D3362FA3EE04EE15350B216" bold="true" pageId="0" pageNumber="101">B</emphasis>
internal wall sculpture of penis, vagina and dart chamber
<emphasis bold="true" pageId="0" pageNumber="101">C</emphasis>
<emphasis id="A9028440F0B1B50D0EA43B66094D566D" bold="true" pageId="0" pageNumber="101">C</emphasis>
ventral view of mantle edge and
<emphasis bold="true" pageId="0" pageNumber="101">D</emphasis>
<emphasis id="8E7C454C06229A58BD53BFDC1AC5BCB7" bold="true" pageId="0" pageNumber="101">D</emphasis>
right view of caudal region.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">Radula.</emphasis>
<paragraph id="29F381EE2FA66A23328A06230BCAE643" pageId="0" pageNumber="101">
<emphasis id="0DE316B075A563FB46222989E82344AA" bold="true" italics="true" pageId="0" pageNumber="101">Radula.</emphasis>
Each row possesses about 161 teeth (80-(18-21)-1-(19-21)-80). Central tooth triangular, tricuspid; ectocones small; mesocone large (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Representative SEM images of radula A-C Hemiplecta humphreysiana, specimen CUMZ 4573 from Singapore A central and lateral teeth B transition from lateral teeth to marginal teeth and C outermost marginal teeth D, E Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand D central and lateral teeth and E outermost marginal teeth F, G Hemiplecta funerea, specimen CUMZ 4575 from Nan, Thailand F central and lateral teeth and G outermost marginal teeth H, I Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand H central and lateral teeth I transition from lateral teeth to marginal teeth J Hemiplecta nemorosa sp. nov., paratype CUMZ 5253 from Maehongsorn, Thailand. Central tooth indicated by ' C'. Numbers indicate the tooth order from lateral to marginal end." figureDoi="10.3897/zookeys.1047.65735.figure10" httpUri="https://binary.pensoft.net/fig/560497" pageId="0" pageNumber="101">10H</figureCitation>
<figureCitation id="E5CF8E1CB160A032E5E5326043195E49" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Representative SEM images of radula A-C Hemiplecta humphreysiana, specimen CUMZ 4573 from Singapore A central and lateral teeth B transition from lateral teeth to marginal teeth and C outermost marginal teeth D, E Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand D central and lateral teeth and E outermost marginal teeth F, G Hemiplecta funerea, specimen CUMZ 4575 from Nan, Thailand F central and lateral teeth and G outermost marginal teeth H, I Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand H central and lateral teeth I transition from lateral teeth to marginal teeth J Hemiplecta nemorosa sp. nov., paratype CUMZ 5253 from Maehongsorn, Thailand. Central tooth indicated by ' C'. Numbers indicate the tooth order from lateral to marginal end." figureDoi="10.3897/zookeys.1047.65735.figure10" httpUri="https://binary.pensoft.net/fig/560497" pageId="0" pageNumber="101">10H</figureCitation>
). Lateral teeth tricuspid; endocone small; mesocone large with pointed tip; ectocone large, basal, and with pointed tip. Outer lateral teeth arranged slightly obliquely, bicuspid; endocone very small to absent; mesocone large trapezoid; ectocone basal, relatively small, and pointed tip. Latero-marginal transition starts from tooth no. 18 to 21. Marginal teeth with curved teeth, bicuspid; endocone usually larger than ectocone (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Representative SEM images of radula A-C Hemiplecta humphreysiana, specimen CUMZ 4573 from Singapore A central and lateral teeth B transition from lateral teeth to marginal teeth and C outermost marginal teeth D, E Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand D central and lateral teeth and E outermost marginal teeth F, G Hemiplecta funerea, specimen CUMZ 4575 from Nan, Thailand F central and lateral teeth and G outermost marginal teeth H, I Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand H central and lateral teeth I transition from lateral teeth to marginal teeth J Hemiplecta nemorosa sp. nov., paratype CUMZ 5253 from Maehongsorn, Thailand. Central tooth indicated by ' C'. Numbers indicate the tooth order from lateral to marginal end." figureDoi="10.3897/zookeys.1047.65735.figure10" httpUri="https://binary.pensoft.net/fig/560497" pageId="0" pageNumber="101">10I</figureCitation>
<figureCitation id="E95E6B49DA638B936D67E0BAC30FD38D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Representative SEM images of radula A-C Hemiplecta humphreysiana, specimen CUMZ 4573 from Singapore A central and lateral teeth B transition from lateral teeth to marginal teeth and C outermost marginal teeth D, E Hemiplecta distincta, specimen CUMZ 4560 from Chanthaburi, Thailand D central and lateral teeth and E outermost marginal teeth F, G Hemiplecta funerea, specimen CUMZ 4575 from Nan, Thailand F central and lateral teeth and G outermost marginal teeth H, I Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand H central and lateral teeth I transition from lateral teeth to marginal teeth J Hemiplecta nemorosa sp. nov., paratype CUMZ 5253 from Maehongsorn, Thailand. Central tooth indicated by ' C'. Numbers indicate the tooth order from lateral to marginal end." figureDoi="10.3897/zookeys.1047.65735.figure10" httpUri="https://binary.pensoft.net/fig/560497" pageId="0" pageNumber="101">10I</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="101">
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">External features.</emphasis>
<paragraph id="0A186934763E07D8D92335E151EBF8C9" pageId="0" pageNumber="101">
<emphasis id="A3D507213D6F71164572D1D5354D7DDF" bold="true" italics="true" pageId="0" pageNumber="101">External features.</emphasis>
Living snail exhibits similar soft body morphology, pulmonary cavity and caudal structure (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">7D</figureCitation>
<figureCitation id="F27168FF7E8AED772A379621FF24839A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Genitalia, mantle edge structure, and caudal region of Hemiplecta esculenta, specimen CUMZ 4553 from Chiang Mai, Thailand A whole genital organ B internal wall sculpture of penis, vagina and dart chamber C ventral view of mantle edge and D right view of caudal region." figureDoi="10.3897/zookeys.1047.65735.figure7" httpUri="https://binary.pensoft.net/fig/560494" pageId="0" pageNumber="101">7D</figureCitation>
) to that of
<taxonomicName higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. humphreysiana" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="humphreysiana">
<emphasis italics="true" pageId="0" pageNumber="101">H. humphreysiana</emphasis>
<taxonomicName id="A8A0160FCBE3BBD6261F7C0F3F946FBE" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. humphreysiana" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="humphreysiana">
<emphasis id="9F0F70CAA54116EFDB4AF74485E9DCCE" italics="true" pageId="0" pageNumber="101">H. humphreysiana</emphasis>
</taxonomicName>
. The distinct characters are the brownish to greyish body and mantle edge; right and left shell lobes absent (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Living snail. A Hemiplecta humphreysiana from Singapore (width about 45 mm) B, C Hemiplecta distincta B from Saraburi, Thailand (width about 65 mm) and C mating pairs D, E Hemiplecta funerea from Nan, Thailand (width about 50 mm) D yellow shell form and E dark shell form (width about 50 mm) F Hemiplecta esculenta from Chiang Mai, Thailand (width about 30 mm)." figureDoi="10.3897/zookeys.1047.65735.figure1" httpUri="https://binary.pensoft.net/fig/560488" pageId="0" pageNumber="101">1F</figureCitation>
<figureCitation id="49EDC05EE4AB0F6926B50C1835E91D7A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Living snail. A Hemiplecta humphreysiana from Singapore (width about 45 mm) B, C Hemiplecta distincta B from Saraburi, Thailand (width about 65 mm) and C mating pairs D, E Hemiplecta funerea from Nan, Thailand (width about 50 mm) D yellow shell form and E dark shell form (width about 50 mm) F Hemiplecta esculenta from Chiang Mai, Thailand (width about 30 mm)." figureDoi="10.3897/zookeys.1047.65735.figure1" httpUri="https://binary.pensoft.net/fig/560488" pageId="0" pageNumber="101">1F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="distribution">
<paragraph pageId="0" pageNumber="101">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="101">
<subSubSection id="1656C793807F624CBA04255D5FC1F8DB" pageId="0" pageNumber="101" type="distribution">
<paragraph id="A6A8BCF83E1744B866D1A0546E87D807" pageId="0" pageNumber="101">Distribution.</paragraph>
<paragraph id="792E8165B9FC44613E6F75BF6D7F2F91" pageId="0" pageNumber="101">
Previously known only from the type locality in northern Vietnam (
<bibRefCitation author="Maassen, WJM" journalOrPublisher="Basteria" pageId="0" pageNumber="101" pagination="13 - 18" refId="B62" refString="Maassen, WJM, 2006. Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae). Basteria 70: 13 - 18" title="Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae)." volume="70" year="2006">Maassen 2006</bibRefCitation>
<bibRefCitation id="95937DFCB1ED9370637E5452019DE546" author="Maassen, WJM" journalOrPublisher="Basteria" pageId="0" pageNumber="101" pagination="13 - 18" refId="B62" refString="Maassen, WJM, 2006. Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae). Basteria 70: 13 - 18" title="Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae)." volume="70" year="2006">Maassen 2006</bibRefCitation>
) and Xieng Khaung, northeastern Laos (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="101" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
<bibRefCitation id="9529AC7FE24BE9A86A15649DAA59A5B2" DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="101" pagination="1 - 166" refId="B47" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
). Recently, we have located populations from northern Thailand in Chiang Mai Province.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="101" type="remarks">
<paragraph pageId="0" pageNumber="101">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="101">
<subSubSection id="4A48B50965AC6255D743237AD9384A39" pageId="0" pageNumber="101" type="remarks">
<paragraph id="BCF76D21F1D84AA46EA23B1D74BB8C93" pageId="0" pageNumber="101">Remarks.</paragraph>
<paragraph id="0E4CDE36BAA7D85F4C0880FA5CF54CB5" pageId="0" pageNumber="101">
The shell features were carefully described in
<bibRefCitation author="Maassen, WJM" journalOrPublisher="Basteria" pageId="0" pageNumber="101" pagination="13 - 18" refId="B62" refString="Maassen, WJM, 2006. Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae). Basteria 70: 13 - 18" title="Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae)." volume="70" year="2006">Maassen (2006)</bibRefCitation>
<bibRefCitation id="D729DBE998760D3FBEA9050471037E81" author="Maassen, WJM" journalOrPublisher="Basteria" pageId="0" pageNumber="101" pagination="13 - 18" refId="B62" refString="Maassen, WJM, 2006. Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae). Basteria 70: 13 - 18" title="Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae)." volume="70" year="2006">Maassen (2006)</bibRefCitation>
. The original description of
<taxonomicName higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. esculenta" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="esculenta">
<emphasis italics="true" pageId="0" pageNumber="101">H. esculenta</emphasis>
<taxonomicName id="747EBA9A68359AB5F973AD2279E55FAD" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="H. esculenta" pageId="0" pageNumber="101" phylum="Mollusca" rank="species" species="esculenta">
<emphasis id="C23D1CD34D339F417A2A6D0566196CD3" italics="true" pageId="0" pageNumber="101">H. esculenta</emphasis>
</taxonomicName>
was based on seven shells and placement within
<taxonomicName authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" lsidName="Hemiplecta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="101">Hemiplecta</emphasis>
<taxonomicName id="BB00988A0C1C56C9B85EC005DF7FDD09" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Ariophantidae" genus="Hemiplecta" kingdom="Animalia" lsidName="Hemiplecta" order="Stylommatophora" pageId="0" pageNumber="101" phylum="Mollusca" rank="genus">
<emphasis id="E43DDBB33D5DC3509ACF355D42534831" italics="true" pageId="0" pageNumber="101">Hemiplecta</emphasis>
</taxonomicName>
was provisional (
<bibRefCitation author="Maassen, WJM" journalOrPublisher="Basteria" pageId="0" pageNumber="101" pagination="13 - 18" refId="B62" refString="Maassen, WJM, 2006. Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae). Basteria 70: 13 - 18" title="Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae)." volume="70" year="2006">Maassen 2006</bibRefCitation>
<bibRefCitation id="42AC86B18E3660E203E3CD6CCB540A5A" author="Maassen, WJM" journalOrPublisher="Basteria" pageId="0" pageNumber="101" pagination="13 - 18" refId="B62" refString="Maassen, WJM, 2006. Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae). Basteria 70: 13 - 18" title="Four new species of terrestrial gastropods from Tonkin, North Vietnam (Gastropoda: Diplommatinidae: Strobilopsidae and Ariophantidae)." volume="70" year="2006">Maassen 2006</bibRefCitation>
), and none of the topotypic specimens have subsequently been examined. The samples from Thailand show only minor variations from the type series, in the presence of a narrow brownish spiral band and slightly elevated spire, which we attribute to intraspecific variation. It is important to examine the genitalia of the topotypic material.
</paragraph>
</subSubSection>