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<mods:namePart>Villinger, Jandouwe</mods:namePart>
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<taxonomicName authority="Austen, 1907" authorityName="Austen" authorityYear="1907" class="Insecta" family="Tabanidae" genus="Chrysops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysops brucei" order="Diptera" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="brucei">Chrysops brucei Austen, 1907</taxonomicName>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Key morphological features of sampled tabanid species. A Ancala fasciata B Tabanus donaldsoni green eyes (in fresh sample) C Tabanus donaldsoni, wing with distinct R 4 appendix (px), indistict hair tuft (hf) D Atylotus nigromaculatus, wing with short, clear appendix on R 4 E Haematopota duttoni, mottled wing with a right angled white thick line (in black circle) between vein A 1 and the wing margin, the R 4 has a long appendix F Haematopota fenestralis, shiny black callus on frons, eyes have bright coloured bands G Chrysops brucei, lateral view of head showing the antenna, black mouth parts and brown legs (partial)." figureDoi="10.3897/zookeys.769.21144.figure4" httpUri="https://binary.pensoft.net/fig/212726" pageId="0" pageNumber="117">Fig. 4G</figureCitation>
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<taxonomicName id="29E5F37E9D88DBCF01DC52D27AA65530" ID-CoL="5Z48R" authority="Austen, 1907" authorityName="Austen" authorityYear="1907" class="Insecta" family="Tabanidae" genus="Chrysops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chrysops brucei" order="Diptera" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="brucei">Chrysops brucei Austen, 1907</taxonomicName>
<figureCitation id="4BF3199FC3F1504CFF02F99004E593EC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Key morphological features of sampled tabanid species. A Ancala fasciata B Tabanus donaldsoni green eyes (in fresh sample) C Tabanus donaldsoni, wing with distinct R 4 appendix (px), indistict hair tuft (hf) D Atylotus nigromaculatus, wing with short, clear appendix on R 4 E Haematopota duttoni, mottled wing with a right angled white thick line (in black circle) between vein A 1 and the wing margin, the R 4 has a long appendix F Haematopota fenestralis, shiny black callus on frons, eyes have bright coloured bands G Chrysops brucei, lateral view of head showing the antenna, black mouth parts and brown legs (partial)." figureDoi="10.3897/zookeys.769.21144.figure4" httpUri="https://binary.pensoft.net/fig/212726" pageId="0" pageNumber="117">Fig. 4G</figureCitation>
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<paragraph pageId="0" pageNumber="117">Location.</paragraph>
<paragraph pageId="0" pageNumber="117">Wakiso, Uganda</paragraph>
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<paragraph id="DEDDD17D741735AF405CB53226419429" pageId="0" pageNumber="117">Location.</paragraph>
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<paragraph pageId="0" pageNumber="117">Descriprion.</paragraph>
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<paragraph id="66CDEDB23B33F2FE9C49B14176DDF4C5" pageId="0" pageNumber="117">Descriprion.</paragraph>
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Head. Head as wide as thorax with black eyes separated by dark brown frons. Frons widened slightly towards antennae and has black and brown long-standing hair. Callus black and glossy. Lower callus wider than long and oval shaped; upper callus with three ocelli arranged in triangular pattern (two lateral and one median ocellus) (Suppl. material 2: Figure S12A). Antennae black with long black hair. Scape and pedicel almost of equal length although scapes slightly longer. Flagellum equally slender with indistinctly marked annulations. Second segment of the palpus brown, slender with black hair; labellum black (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Key morphological features of sampled tabanid species. A Ancala fasciata B Tabanus donaldsoni green eyes (in fresh sample) C Tabanus donaldsoni, wing with distinct R 4 appendix (px), indistict hair tuft (hf) D Atylotus nigromaculatus, wing with short, clear appendix on R 4 E Haematopota duttoni, mottled wing with a right angled white thick line (in black circle) between vein A 1 and the wing margin, the R 4 has a long appendix F Haematopota fenestralis, shiny black callus on frons, eyes have bright coloured bands G Chrysops brucei, lateral view of head showing the antenna, black mouth parts and brown legs (partial)." figureDoi="10.3897/zookeys.769.21144.figure4" httpUri="https://binary.pensoft.net/fig/212726" pageId="0" pageNumber="117">4G</figureCitation>
<figureCitation id="43E9981B3A184244530800E329E3E91A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Key morphological features of sampled tabanid species. A Ancala fasciata B Tabanus donaldsoni green eyes (in fresh sample) C Tabanus donaldsoni, wing with distinct R 4 appendix (px), indistict hair tuft (hf) D Atylotus nigromaculatus, wing with short, clear appendix on R 4 E Haematopota duttoni, mottled wing with a right angled white thick line (in black circle) between vein A 1 and the wing margin, the R 4 has a long appendix F Haematopota fenestralis, shiny black callus on frons, eyes have bright coloured bands G Chrysops brucei, lateral view of head showing the antenna, black mouth parts and brown legs (partial)." figureDoi="10.3897/zookeys.769.21144.figure4" httpUri="https://binary.pensoft.net/fig/212726" pageId="0" pageNumber="117">4G</figureCitation>
; Suppl. material 2: Figure S12B).
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<paragraph pageId="0" pageNumber="117">Thorax. Black thorax with black and brown hair; sub-lateral stripes brown and reach posterior border of mesonotum. Basicosta of wing black with prominent thorn-like projection, while the costa is slender and long without longitudinal groove. Clear wing with longitudinal dark brown band that does not reach posterior margin. Fore leg dark brown with dark brown hair (Suppl. material 2: Figure S12B). Black halteres and stalk.</paragraph>
<paragraph pageId="0" pageNumber="117">Abdomen. Abdomen black with black and white hair and parallel lateral sides that taper slightly at fifth segment. Each segment ashy grey at posterior margin with predominantly white hair. Indistinct greyish median stripe. Seventh segment rounded; ventral abdominal surface black with long fine whitish hair.</paragraph>
<paragraph id="995F3EE3A0340A30366E3DE42919A2E0" pageId="0" pageNumber="117">Thorax. Black thorax with black and brown hair; sub-lateral stripes brown and reach posterior border of mesonotum. Basicosta of wing black with prominent thorn-like projection, while the costa is slender and long without longitudinal groove. Clear wing with longitudinal dark brown band that does not reach posterior margin. Fore leg dark brown with dark brown hair (Suppl. material 2: Figure S12B). Black halteres and stalk.</paragraph>
<paragraph id="CD4C3B0885D7245D1B79833FDEC3A849" pageId="0" pageNumber="117">Abdomen. Abdomen black with black and white hair and parallel lateral sides that taper slightly at fifth segment. Each segment ashy grey at posterior margin with predominantly white hair. Indistinct greyish median stripe. Seventh segment rounded; ventral abdominal surface black with long fine whitish hair.</paragraph>
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<paragraph id="7C505B1A8186D40D2AC97E6C98044D39" pageId="2" pageNumber="51">Taxon classification Animalia Araneae Agelenidae</paragraph>
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@ -77,7 +77,7 @@ Wang and
: 46 (synonymized 2 genera).
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<taxonomicName id="0F5464F2D9C090ADDDC31BBAB7F48ADD" class="Arachnida" family="Agelenidae" genus="Paracoelotes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Paracoelotes" order="Araneae" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">Paracoelotes</taxonomicName>
<taxonomicName id="0F5464F2D9C090ADDDC31BBAB7F48ADD" genus="Paracoelotes" lsidName="Paracoelotes" pageId="2" pageNumber="51" rank="genus">Paracoelotes</taxonomicName>
Brignoli, 1982: 348. Type species
<taxonomicName id="EAB0CF79BB36E2E1867FA651B6571067" class="Arachnida" family="Agelenidae" genus="Coelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coelotes armeniacus" order="Araneae" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="armeniacus">Coelotes armeniacus</taxonomicName>
Brignoli, 1978 from Turkey;
@ -110,7 +110,7 @@ sp. n., male holotype. A Prolateral view B Ventral view C Retrolateral view. Sca
Described in
<bibRefCitation id="58AE4E7AE169A1BCA3AE05DC0F7BBE78" author="Wang, XP" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="17" pageNumber="66" pagination="1 - 150" title="A generic-level revision of the spider subfamily Coelotinae (Araneae, Amaurobiidae)." url="10.1206/0003-0090(2002)269&lt;0001:AGLROT&gt;2.0.CO;2" volume="269" year="2002">Wang (2002</bibRefCitation>
, sub
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<taxonomicName id="ADB56AB240DACBDFA0C025F88AD57CF4" genus="Paracoelotes" lsidName="Paracoelotes" pageId="2" pageNumber="51" rank="genus">Paracoelotes</taxonomicName>
).
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View file

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<treatment id="2D4B8EC9FE405B6FB5F0E2187DAAC615" ID-DOI="http://doi.org/10.5281/zenodo.14232109" ID-GBIF-Taxon="247601126" ID-Zenodo-Dep="14232109" ID-arpha="2D4B8EC9-FE40-5B6F-B5F0-E2187DAAC615" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615" httpUri="http://treatment.plazi.org/id/2D4B8EC9FE405B6FB5F0E2187DAAC615">
<subSubSection id="0DAC4F268E8ED4A2CE48810C45CE2D89" type="nomenclature">
<paragraph id="22DD3C693B1A395EE11D869B18BB2B3F">
<taxonomicName id="0E1AC907BF9E8DB4262224789AA07A20" authority="Cheek ex Mackinder, Wieringa &amp; Burgt" authorityName="Mackinder, Wieringa &amp; Burgt" class="Magnoliopsida" family="Leguminosae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="6F88C63E5974C8F451034E5734D977E4" italics="true">Talbotiella bakossiensis</emphasis>
Cheek ex Mackinder, Wieringa &amp; Burgt
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="181248ACE03C1DDC57ADFEF66CEDF0AB" type="description">
<paragraph id="4A050C7EEBA65B11C64F8018A3CF25F3">
<figureCitation id="4D4732FEB41D6BF784CEAC3743F8DF6E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Talbotiella couteronii and T. bakossiensis in Cameroon." figureDoi="10.5091/plecevo.133256.figure1" httpUri="https://binary.pensoft.net/fig/1179774">Fig. 1</figureCitation>
,
<tableCitation id="14511D933C1FF232CE0A39DDC6A23493" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Leaf characteristics of the 10 species of Talbotiella. The species are sorted by number of leaflet pairs. Leaflet length and leaflet width were measured on the largest pair of leaflets because leaflet size varies considerably depending on the position of the leaflet. Adapted from Table 1 in Mackinder et al. (2010 b); the leaflet characteristics were now measured only on leaves of fertile branches; and the measurements of several newly determined collections are included." httpUri="http://table.plazi.org/id/398F80273458B3F1B8C43F7A4E6F1957" tableUuid="398F80273458B3F1B8C43F7A4E6F1957">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EALAE" type="material">
<paragraph id="41E83B1BFD0CE158F561F026685A5BB4">
<heading id="6F8C9FEAEF858E497E178AFCD611A3EB" reason="title">Type.</heading>
</paragraph>
<paragraph id="09E9516193D49B8534F5C20E84B0E15D">
<materialsCitation id="1D126285794067A2C6AD11922A4BBDC8" ID-GBIF-Occurrence="4985319332" collectingDate="1998-11-18" collectionCode="K, P, WAG, YA" collectorName="Cheek" country="Cameroon" elevation="1350" latitude="4.9833336" location="Mwanzum to Kodmin between hut 1 and 2" longLatPrecision="1306" longitude="9.666667" municipality="Bakossi Mts" specimenCode="WAG 0248233" specimenCount="2" stateProvince="Southwest Region" typeStatus="holotype">
<collectingCountry id="398AE2CDD66C7410FE2A481CBB5D0C92" name="Cameroon">CAMEROON</collectingCountry>
<emphasis id="36431036C55C17E930FDA20E2203E520" bold="true">
<collectingRegion id="3467BA342A6D836B054B1757AE8B1D33" country="Cameroon" name="South-West">Southwest Region</collectingRegion>
</emphasis>
<collectingMunicipality id="448934D199D3D8BF21ADD3D9B3FCC2A2">Bakossi Mts</collectingMunicipality>
,
<location id="5C976FAFB4CDA4F982A88841F88F8D7A" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:5C976FAFB4CDA4F982A88841F88F8D7A" country="Cameroon" county="Bakossi Mts" latitude="4.9833336" longLatPrecision="1306" longitude="9.666667" municipality="Bakossi Mts" name="Mwanzum to Kodmin between hut 1 and 2" stateProvince="Southwest Region">Mwanzum to Kodmin between hut 1 and 2</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ELLAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.666667,4.983333]}">
<geoCoordinate id="39A864ADA78638424DDE302D95AF5044" degrees="4" direction="north" minutes="59" orientation="latitude" precision="925" value="4.9833336">4°59N</geoCoordinate>
,
<geoCoordinate id="B1F131BE52727B5638A3D0A6C5B6A154" degrees="9" direction="east" minutes="40" orientation="longitude" precision="925" value="9.666667">9°40E</geoCoordinate>
</named-content>
;
<quantity id="3CA16B045E4B66EAD2DA19FA81C47426" metricMagnitude="3" metricUnit="m" metricValue="1.35" unit="m" value="1350.0">
<elevation id="4E5AA49B388FF78BAC5FC2249BB95178" metricMagnitude="3" metricUnit="m" metricValue="1.35" unit="m" value="1350.0">1350 m</elevation>
</quantity>
;
<date id="0DA0928770A2957691BCF82C3F5E4114" value="1998-11-18">
<collectingDate id="726219FDFA72DA06C1DF1DA36D2B4724" value="1998-11-18">18 Nov. 1998</collectingDate>
</date>
;
<emphasis id="0A0F1DD1A1D80972C31DAC05D5A08460" italics="true">
<collectorName id="6908FAC4AD1044C999252F9F24E403C8">Cheek</collectorName>
9688
</emphasis>
;
<typeStatus id="B8B9F84C5A249A9D2ED48F015AAB1C06" type="holotype">holotype</typeStatus>
<collectionCode id="ABA8675F33BDEB61A9AF96808271C43A">K</collectionCode>
n. v.;
<typeStatus id="2E86BB83BAF8FF444D40AD270EF8DA5C" type="isotype">isotypes</typeStatus>
:
<collectionCode id="A2DF182252245F2735AF4CCD628BE0F8">P</collectionCode>
n. v.,
<collectionCode id="4CE89519C51DE30F8E1891390D5F5D6A" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
[
<specimenCode id="BBD8077714E16E3389E2853B6C658650" collectionCode="WAG" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG 0248233</specimenCode>
] image,
<collectionCode id="0535E2374CEDE4B70D269CED34B8725A" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
n. v
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ESLAE" type="description">
<paragraph id="704525020647570807E14D04792CBB2E">
<heading id="1A63E7EE2C312BE59938F0C68570B3B5" reason="title">Description.</heading>
</paragraph>
<paragraph id="FEA9EFCA9AEC89C5CE10D0D527F632F3">
<underline id="5263B8CE2FF8924F0218AA68B7515E33">Tree</underline>
up to
<quantity id="398246BE562E9005C3EB714FABC32EAF" metricMagnitude="1" metricUnit="m" metricValue="2.9" unit="m" value="29.0">29 m</quantity>
high, stem up to
<quantity id="2F9A1BED413415DE80E82BEA6F7C2009" metricMagnitude="-1" metricUnit="m" metricValue="7.4" unit="cm" value="74.0">74 cm</quantity>
diameter; bole weakly 5 - fluted (
<emphasis id="544D098FB80D49D01BC76DE2B8EACB5C" italics="true">Cheek 10382</emphasis>
; see original description), base with small buttresses; bark smooth to slightly rough; outer slash thick with dark outer line, dull orange, granular-sandy; inner slash pale orange-yellow, fibrous; crown dense.
<underline id="5DD21D582E7675D7FAE2A7382DDB0C7D">Twigs</underline>
moderately to densely pubescent, hairs
<quantity id="36A953B49ECF3A7E089726478BDCD9AC" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
long.
<underline id="3A61FE85D05600D8384DCF8DDBBB2C30">Bud scales</underline>
510, caducous, distichous, brown, coriaceous, not keeled, outer and inner surface glabrous, proximal scale orbicular, ca
<quantity id="1B7FA0ED7F9248EA2C5972854A9A1F0E" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
diam.; distal scales becoming progressively longer and relatively narrower; apical scale obovate, 12 ×
<quantity id="87F30CA48D182CF7249C011FF47C9243" metricMagnitude="-3" metricUnit="m" metricValue="8.0" unit="mm" value="8.0">8 mm</quantity>
, leafy, not keeled, pubescent along midrib.
<underline id="3A311E58329DE1483DE0FB43E313A7BB">Stipules</underline>
early caducous, in pairs, free, auriculate at base, the auricle suborbicular, up to
<quantity id="9BFD5B254774E44C697D71B3B492FFFF" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
long, with 26 acute triangular lobes, glabrous on both surfaces but margins ciliate; upper part of stipule narrowly lanceolate, 817 ×
<quantity id="D4BB719E395426827AA5A8A0550151EB" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
; outer surface densely pubescent along the midrib, veins sparsely pubescent, margins ciliate, inner surface glabrous, apex acute.
<underline id="9C68D3755BB408E78D3A342D8A2FAFBD">Leaves</underline>
alternate, paripinnate, 4.58.5 ×
<quantity id="42B596EE0FE55285D4C820DE7759829B" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" unit="cm" value="1.8" valueMax="2.0" valueMin="1.6">1.62 cm</quantity>
; petiole
<quantity id="AC96A40344C089FF5F028E5070F155F0" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
; leaf rachis
<quantity id="5D97E2E110F4C778AE97C3D9E5F157A1" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" unit="cm" value="6.0" valueMax="8.0" valueMin="4.0">48 cm</quantity>
long (up to
<quantity id="7446F234C35040E0E4FF50578C0D4BA8" metricMagnitude="-1" metricUnit="m" metricValue="1.2" unit="cm" value="12.0">12 cm</quantity>
in saplings), moderately pubescent, with a tuft of longer hairs at the insertion of leaflets; leaflets sessile, in 2127 pairs, upper and middle pairs opposite, lower 35 pairs subopposite,
<quantity id="F6B46CE74718FB18D4994F8E21B80854" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" unit="mm" value="11.0" valueMax="14.0" valueMin="8.0">814 mm</quantity>
×
<quantity id="F6AAD94AE264CC188EDFB273AEEC050B" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
(up to 22 ×
<quantity id="64D15F342471C74521FEA74D4D95D22B" metricMagnitude="-3" metricUnit="m" metricValue="6.0" unit="mm" value="6.0">6 mm</quantity>
in saplings), narrowly oblong, base asymmetric, proximal margin tapering towards the apex, glabrous above, lower surface appearing glabrous with a hand lens but moderately appressed puberulous under a microscope, fringing hairs
<quantity id="E490644B9B09B93653F1C104F922F9CC" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" unit="mm" value="0.625" valueMax="0.75" valueMin="0.5">0.50.75 mm</quantity>
long, present particularly on the lower margins on young leaflets but scattered or absent on mature foliage, midvein central, prominent above and below for most of its length, becoming obscure just before the apex; small glands 12 ( 3), not visible with a hand lens but visible under a microscope, present on the distal half of the lower surface, positioned about midway between midvein and margin, always present on basal pair, present or absent on middle pairs, not seen on uppermost pair.
<underline id="3005C47E989E2FAEF9F09453C7D506B0">Inflorescence</underline>
a 715 - flowered raceme, axis
<quantity id="A21A4E25DD7E129FAB77D4FB40C172F2" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.5" metricValueMin="2.3" unit="cm" value="2.9" valueMax="3.5" valueMin="2.3">2.33.5 cm</quantity>
long, with a moderate to dense indumentum of golden patent hairs, ca
<quantity id="D720AAD3FC83F133451DBE4E370E961A" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" unit="mm" value="1.05" valueMax="1.5" valueMin="0.6">0.61.5 mm</quantity>
long, floral bracts slightly spathulate, 610 ×
<quantity id="D7612F077D6AC249D273C56E56415DBB" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
, outer surface densely appressed tomentose, inner surface glabrous, persistent; pedicels with moderately spreading hairs
<quantity id="BD19A5165A32205FAF15ADB96BFE720C" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.30.5 mm</quantity>
long, portion of pedicel below bracteoles
<quantity id="BABA3B90C50EA01DACC676FEE54BD5BC" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">24 mm</quantity>
long, bracteoles subopposite to opposite, narrowly oblong, ca 5 ×
<quantity id="16499487CB8B2B8613826413FD843A20" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 mm</quantity>
, tapering towards the apex in upper third, mostly glabrous but with margins ciliate at least in the lower half and along the basal part of the midvein, with a tuft of hairs at the apex, portion of pedicels above bracteoles ca
<quantity id="49433E2808667BAC0D64F12BCF04DE47" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
.
<underline id="92932564245449D477F8DB3FE81AED63">Hypanthium</underline>
narrowly campanulate,
<quantity id="7D4BC76B50F3B54B921F37F338B2B8F4" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
deep, puberulous at the mouth, otherwise glabrous on both surfaces.
<underline id="D5D69728CA45C9860B186B49ACFBA02D">Sepals</underline>
4, obovate, white, subequal, cucullate, 57 ×
<quantity id="C5E1A29E9E20200802BDB3741093C9D9" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.53 mm</quantity>
, glabrous on both surfaces except for scattered marginal pubescence on upper third with tuft of hairs 0.2
<quantity id="18C834B75327FCE4D5A0F51249DD7CCB" metricMagnitude="-4" metricUnit="m" metricValue="3.0" unit="mm" value="0.3">0. 3 mm</quantity>
long at the rounded apex.
<underline id="2D79498BD22EAFEEDE4974BEF392937E">Petals</underline>
absent.
<underline id="D7752E23125C79AF3273F3185172CDCC">Stamens</underline>
10; filaments free,
<quantity id="C939F7EF956DA6BB202E980128707B1E" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">79 mm</quantity>
long, glabrous; anthers elliptic, ca
<quantity id="EAE7F47D43B51EE5A4AABDB777084927" metricMagnitude="-3" metricUnit="m" metricValue="1.2" unit="mm" value="1.2">1.2 mm</quantity>
long.
<underline id="8F921ABAD355D3AB4D3F14C9AE3006BD">Ovary</underline>
oblong, 2.32.7 ×
<quantity id="5EB42C0D1E3031F3580A0310A47F6631" metricMagnitude="-3" metricUnit="m" metricValue="1.3" metricValueMax="1.5" metricValueMin="1.1" unit="mm" value="1.3" valueMax="1.5" valueMin="1.1">1.11.5 mm</quantity>
, densely golden to red-brown tomentose, on a glabrous stipe
<quantity id="C6326BBA55AC9B1EC985FC335429CB1E" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
long, fused with the hypanthium near its apex, free part of stipe very short; ovules 2; style ca
<quantity id="85D65B92BDFB2AE6A649035FE21D30B8" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
long, moderately to densely pubescent for most of its length, becoming glabrous just under the stigma, stigma capitate.
<underline id="76B310A2689158DAD822ACB76C0B6D8E">Pod</underline>
obovate-triangular, 4.56.7 ×
<quantity id="25EE141035A8F58BC5AC4B7D2176EA81" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" unit="cm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 cm</quantity>
, upper suture slightly broadened, surfaces and suture moderately golden puberulous.
<underline id="FBD9C0DE91CD0FA3C8C35BD867FFDA9A">Seeds</underline>
discoid, 12 per fruit, 910 × 910 ×
<quantity id="38948B41E1D48796004EBF9411131172" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUMAE" type="distribution">
<paragraph id="5D36DBFBEBBF9CA044552668578F1377">
<heading id="5F36618AA3A956AB3B5EC6A324721F2D" reason="title">Distribution and chorology.</heading>
</paragraph>
<paragraph id="51CD64708CD19B820D63CBECB813BA15">
<taxonomicName id="B1622826D0455170CE6916D7C846B07C" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="F571FB2EE52E74DF746ADBFD6180E2F3" italics="true">Talbotiella bakossiensis</emphasis>
</taxonomicName>
is currently known only from the Southwest and Littoral Regions in
<collectingCountry id="F8384580C030E10B7154F51337E5FD71" name="Cameroon">Cameroon</collectingCountry>
(Fig.
<figureCitation id="462D266FF3F1BD54B7B877ABEE78C1CE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of Talbotiella couteronii and T. bakossiensis in Cameroon." figureDoi="10.5091/plecevo.133256.figure1" httpUri="https://binary.pensoft.net/fig/1179774">1</figureCitation>
). Lower
<collectingCountry id="38E56B6468CA87E5E08B170D9E7FAD23" name="Guinea-Bissau">Guinea</collectingCountry>
Domain of the Guineo-Congolian Region.
</paragraph>
<caption id="E6083CFB629F07C814E1420E09932DA5" ID-DOI="10.5091/plecevo.133256.figure1" ID-arpha="49A32D5F-FFA4-5A44-B4AF-6E8A42764AA2" httpUri="https://binary.pensoft.net/fig/1179774" startId="F1">
<paragraph id="65BC6EB34C66FF56A2F76CF9CC2B5C52">
<label id="0FBA714F7A12E2EA1D0A8010E5FC0021">Figure 1.</label>
</paragraph>
<paragraph id="2899F9B7A17281D25B50BDB7F1116180">
Distribution of
<taxonomicName id="CECB6F5BC92B844FD27A1AD283F5EC1A" authorityName="Sonke, M. Simo &amp; Burgt" authorityYear="2024" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="couteronii">
<emphasis id="E9E10EC3F79F93F8CF61169329463FBF" italics="true">Talbotiella couteronii</emphasis>
</taxonomicName>
and
<taxonomicName id="983EC0A729EDF1550D58FAA0D44ED134" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="D5467338E076B60EC412D83FD96E82D1" italics="true">T. bakossiensis</emphasis>
</taxonomicName>
in Cameroon.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EMOAE" type="habitat">
<paragraph id="165C59ED2B78037CB49FC7620D17C34A">
<heading id="AAEBAB396BE4D34E5214FB3760EF4840" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="AB5D786ED2AB66BBDFC735F641848860">
<taxonomicName id="7805BFF9455FCCC447DD7CC49D37BE7D" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="14420214EC538B3DF71DA0E7B4AD858E" italics="true">T. bakossiensis</emphasis>
</taxonomicName>
occurs in lowland to submontane forest at
<quantity id="E48B98C2EF5D9AE81932773EED4768CE" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="13.5" metricValueMin="1.0" unit="m" value="725.0" valueMax="1350.0" valueMin="100.0">1001350 m</quantity>
altitude.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4OAE" type="phenology">
<paragraph id="C755D44D296CDC1E1A859F6AF06D49F0">
<heading id="49699EF3547684DA34826F355A0189DD" reason="title">Phenology.</heading>
</paragraph>
<paragraph id="BCF192BD891BAAF8702D4FD319AC46E1">Flowering specimens were collected in February, October, and November, and fruiting specimens in February and December.</paragraph>
</subSubSection>
<subSubSection id="SECID0ECPAE" type="Preliminary IUCN conservation assessment">
<paragraph id="20B4C1B63E2155B4D0882CD1D5C46785">
<heading id="767B422D7959C75C9033BB7DBBDB4CB3" reason="title">Preliminary IUCN conservation assessment.</heading>
</paragraph>
<paragraph id="DAD66A901A46D75943683D43BB44D924">
<taxonomicName id="A7ABEE6585CF559E9442A92608167A45" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="8C24E032B33967F3342A602A88FE2B39" italics="true">Talbotiella bakossiensis</emphasis>
</taxonomicName>
is assessed as Endangered B 1 ab (i, ii, iii, v) + 2 ab (i, ii, iii, v) on the IUCN Red List (
<bibRefCitation id="5835F8C457A57050B1D3CB8BE1C996F9" DOI="10.2305/IUCN.UK.2022-1.RLTS.T172492710A172492722.en" author="Cheek" etAl="et al." firstAuthor="Cheek" refId="B6" refString="Cheek M, Rotton H, van der Burgt X (2022) Talbotiella bakossiensis. The IUCN Red List of Threatened Species 2022: e. T 172492710 A 172492722. https://doi.org/10.2305/IUCN.UK.2022-1.RLTS.T172492710A172492722.en [accessed 26. 07. 2024]" year="2022">Cheek et al. 2022</bibRefCitation>
). Nevertheless, this published assessment does not take into account five collections recently named as this species. The main threats mentioned by the published assessment include shifting agriculture and logging. Including the five recently named collections, the extent of occurrence is now calculated at
<quantity id="FE5EECA6457C5D166714E118A00E9428" metricMagnitude="6" metricUnit="m" metricValue="2.58" unit="km" value="2580.0">2580 km</quantity>
<superScript id="27CFEED5C3296C5473BD11B50CF718E5">2</superScript>
and the estimated area of occupancy as
<quantity id="52D09013229DBCC564EE7348E4FEBB0E" metricMagnitude="4" metricUnit="m" metricValue="3.2" unit="km" value="32.0">32 km</quantity>
<superScript id="E89405E24AAF589938DC2B5455160DDB">2</superScript>
, on the basis of a 2 ×
<quantity id="9984651CE49F3DFB2BC6000C71633955" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
grid size. We believe the threats mentioned by the original assessment are still applicable, and the total of estimated locations reaches five when including the five collections recently named as this species. As a consequence, the assessment of
<taxonomicName id="5C937FB95755D8BFEA0FA91DD43BD189" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="1A48923179A22891655A9EA4C56F009F" italics="true">T. bakossiensis</emphasis>
</taxonomicName>
remains Endangered B 1 ab (i, ii, iii, v) + 2 ab (i, ii, iii, v).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGQAE" type="material">
<paragraph id="D73FD6E8B25639924237CF7A5A6396A6">
<heading id="614ABEB57C569CCFB107DF6EE4A1EA2C" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="D171754743CC40C254FE8243E135498B">
<materialsCitation id="152B42E45D422C007D8E43C643E59923" ID-GBIF-Occurrence="4985319335" collectingDate="2015-10-10" collectionCode="BM, BR, C, E, EA, ETH, F, FI, FHO, G, GH, H, HBG, K, LBV, LE, LISC, MA, MO, MPU, NY, O, P, PRE, S, SCA, SING, US, W, WAG, YA" collectorName="van der Burgt" country="Cameroon" county="Along" elevation="100" latitude="4.323972" location="Along the road from Djoukou to Yabassi" longLatPrecision="1" longitude="10.071806" municipality="Djoukou" specimenCode="BR 0000015256874" specimenCount="1" stateProvince="Littoral Region">
<collectingCountry id="E98E23DB6EE207B62BE1E86E2BAE8DFE" name="Cameroon">CAMEROON</collectingCountry>
<emphasis id="9350ED7D070ABA34ADCD9A8260DF63AD" bold="true">
<collectingRegion id="FBC58D25B97D2547B405D5EA3E2F4EB7" country="Cameroon" name="Littoral">Littoral Region</collectingRegion>
</emphasis>
<location id="8E510D75A13A751E248D24EF8595BD95" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:8E510D75A13A751E248D24EF8595BD95" country="Cameroon" county="Along" latitude="4.323972" longLatPrecision="1" longitude="10.071806" municipality="Djoukou" name="Along the road from Djoukou to Yabassi" stateProvince="Littoral Region">Along the road from Djoukou to Yabassi</location>
,
<locationDeviation id="11071E491AA7AF527CA24D06EA261340">
<quantity id="863A5DEA2833185B9F662D7F8E64DF0E" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
on a side road to Yingui
</locationDeviation>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERQAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[10.071806,4.323972]}">
<geoCoordinate id="514915D38877D8C471838EB233324C0D" degrees="4" direction="north" minutes="19" orientation="latitude" precision="1" seconds="26.3" value="4.323972">4°1926.3”N</geoCoordinate>
,
<geoCoordinate id="CC3DFB245BDB32D434313947FC8F3C68" degrees="10" direction="east" minutes="04" orientation="longitude" precision="1" seconds="18.5" value="10.071806">10°0418.5”E</geoCoordinate>
</named-content>
;
<quantity id="4AC7ACCAEDD609917821DB2262431C4D" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation id="50052B1CB7D2F1B3474ACADB51FCA44F" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
;
<date id="A0BC29BB039B3C139CF8D6C0D9AEBDE2" value="2015-10-10">
<collectingDate id="3DD460E23E9041720EC91AB7CF858E95" value="2015-10-10">10 Oct. 2015</collectingDate>
</date>
;
<emphasis id="DC119AB4A536538F5141EB54E26DD1FA" italics="true">
<collectorName id="A298D6DF1137E4A5A32AEE78613DF10A">van der Burgt</collectorName>
1889
</emphasis>
; B,
<collectionCode id="B6AA61F97F078223BF64F4DA7A42CEAC" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
,
<collectionCode id="762CBACEE3EB00684B14682F86CF7243" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
[
<specimenCode id="53CEBF0AB99C130E446ABF54049FA224" collectionCode="BR" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR 0000015256874</specimenCode>
V] n. v.,
<collectionCode id="9BADB7806BAF8280ADAB3EA6E4015C81">C</collectionCode>
n. v.,
<collectionCode id="A8D267ECDDE3171204E74CE9C43396B4">E</collectionCode>
n. v.,
<collectionCode id="7AAF5B0ABA98C09880E241B6972FBF32">EA</collectionCode>
n. v.,
<collectionCode id="B1C6E7AA31A581B0D59F239B99115B19">ETH</collectionCode>
n. v.,
<collectionCode id="58E050461D50E563FAFB4803C876621B">F</collectionCode>
n. v.,
<collectionCode id="3CBF9B9E41DA62533D42CAF202F52C2D">FI</collectionCode>
n. v.,
<collectionCode id="F32C45C03D78A7CE34B27209AD95E99D">FHO</collectionCode>
n. v.,
<collectionCode id="D915590AFE93FBF20B258179368B5BCD">G</collectionCode>
n. v.,
<collectionCode id="42107ACCDF4B412D851E8F4544153632">GH</collectionCode>
n. v.,
<collectionCode id="B46FB322ADB4866C09F301372FFDE0DC">H</collectionCode>
n. v.,
<collectionCode id="53AAD0DEE31E5AA51F0F5C1D9828D940">HBG</collectionCode>
n. v.,
<collectionCode id="46ACCB5C168660F54C0DBD3588BEBD9A">K</collectionCode>
,
<collectionCode id="AC3CA80BA8F5C78D095D93001D01DACF">LBV</collectionCode>
n. v.,
<collectionCode id="191D1EF712B48681EDD28D3592E32620">LE</collectionCode>
n. v.,
<collectionCode id="7C09145AE9E717B059558B0D9CDF5302">LISC</collectionCode>
n. v.,
<collectionCode id="345F082C25AF12660DF5F2A30E56C77E" country="Spain" lsid="urn:lsid:biocol.org:col:15682" name="Real Jardín Botánico" type="Herbarium">MA</collectionCode>
n. v.,
<collectionCode id="E685753FDA515DAF0B1842ED699FE9B1" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" type="Herbarium">MO</collectionCode>
n. v.,
<collectionCode id="481CA2C32AE1AF44FA8B7125E2179B52">MPU</collectionCode>
n. v.,
<collectionCode id="27748CBF6071A4A39613359E19ED2006">NY</collectionCode>
n. v.,
<collectionCode id="4122DB97E388A2186B3D8216A2A40DF4">O</collectionCode>
n. v.,
<collectionCode id="8650BD457321C81616226F05D72277FC">P</collectionCode>
n. v.,
<collectionCode id="2C355B5A7E98BA935EEAE555ACAC8BDE">PRE</collectionCode>
n. v.,
<collectionCode id="B69424F0658900F15ECC05B1DE88E513">S</collectionCode>
n. v.,
<collectionCode id="8F885E59B7E0DAC527FFC418D3BB316F">SCA</collectionCode>
n. v.,
<collectionCode id="222A081BA47AE11A7503BED77AD2ABEB">SING</collectionCode>
n. v.,
<collectionCode id="CB0A0500AA4C14C741ABB1A23E482DDB">US</collectionCode>
n. v.,
<collectionCode id="CE7686C39BEC54C86436EFF93736219F">W</collectionCode>
n. v.,
<collectionCode id="F804A9D28FF005EBFB487371D59594D0" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
n. v.,
<collectionCode id="2E93B0BCF6F0FEE76D4A1AE4C0EAF7D9" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
n. v
</materialsCitation>
. •
<materialsCitation id="A3CE0209751D09D2C1D3C0794D13DB21" ID-GBIF-Occurrence="4985319318" collectedFrom="same tree" collectingDate="2019-12-02" collectionCode="B, BR, G, K, LISC, MO, P, WAG, YA" collectorName="van der Burgt" country="United States of America" county="Along" elevation="100" latitude="4.323972" location="Along the road from Djoukou to Yabassi" longLatPrecision="1" longitude="10.071806" municipality="Djoukou" specimenCount="1" stateProvince="Littoral Region">
<location id="208C49BF0CDD33F1C36E15E7552C6EBD" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:208C49BF0CDD33F1C36E15E7552C6EBD" country="United States of America" county="Along" latitude="4.323972" longLatPrecision="1" longitude="10.071806" municipality="Djoukou" name="Along the road from Djoukou to Yabassi" stateProvince="Littoral Region">Along the road from Djoukou to Yabassi</location>
,
<locationDeviation id="6C987E339DF68D167CBCC974F07D2BFF">
<quantity id="27A633E4E15C87CE05D0FB98ADEBA709" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
on a side road to Yingui
</locationDeviation>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2QAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[10.071806,4.323972]}">
<geoCoordinate id="FD8C594A45F9B5825919E5CD8DB0A900" degrees="4" direction="north" minutes="19" orientation="latitude" precision="1" seconds="26.3" value="4.323972">4°1926.3”N</geoCoordinate>
,
<geoCoordinate id="14A5F1E36201EB1C5B26D83C1512EB07" degrees="10" direction="east" minutes="04" orientation="longitude" precision="1" seconds="18.5" value="10.071806">10°0418.5”E</geoCoordinate>
</named-content>
;
<quantity id="74E42B5374C85EE48187455C213ED51A" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">
<elevation id="EF3EBF0DFA80D530AC1128AB1BF01079" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</elevation>
</quantity>
;
<collectedFrom id="BCB718D3E5C8DB34C931BB226748C995">same tree</collectedFrom>
;
<date id="B8F40110A68F3E161D6A503FF3866D15" value="2019-12-02">
<collectingDate id="E756EBB2CA02B08A13EC791046A93ADC" value="2019-12-02">2 Dec. 2019</collectingDate>
</date>
;
<emphasis id="0CE85DBCB86F8AD0D07F3723F10BEC46" italics="true">
<collectorName id="EDED1C7A823C7AF59CCDAE8C1F2204F5">van der Burgt</collectorName>
2356
</emphasis>
;
<collectionCode id="E4B31B43E707541AF26731947E1F5573">B</collectionCode>
n. v.,
<collectionCode id="E3BDF37DBF117E9E21CCB99AB8554491" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
n. v.,
<collectionCode id="1288AB33D7C2CA3288D66179655B4BA0">G</collectionCode>
n. v.,
<collectionCode id="B454C2D8726FA7B721A8BD49734D79C6">K</collectionCode>
,
<collectionCode id="5465E64909D3529100CF41EC1D7DFD24">LISC</collectionCode>
n. v.,
<collectionCode id="D99DE3C9C426603E845C04529E8EC303" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" type="Herbarium">MO</collectionCode>
n. v.,
<collectionCode id="8061ADB058EF1AA30C6DBF9110A631FF">P</collectionCode>
n. v.,
<collectionCode id="C5E1546A5234614A5F71F8F3389CCE68" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
n. v.,
<collectionCode id="AC7AFF5E1ECDA68433E3D1F59F08CEFE" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
n. v.
</materialsCitation>
<materialsCitation id="D25B688E87E0789B8E1AAB3D8D894DCE" ID-GBIF-Occurrence="4985319329" collectingDate="1998-11-18" collectionCode="K, WAG, YA" collectorName="Cheek" country="United States of America" latitude="5.0" location="Nyale" longLatPrecision="1305" longitude="9.633333" municipality="Kupe-Mwanenguba" specimenCount="1" stateProvince="Southwest Region">
<emphasis id="03AE2C83B2FC2A818D65B57B5B632BF5" bold="true">
<collectingRegion id="A4474832FB7F805908151DCE26DBD2F3" inferredBySuffix="true">Southwest Region</collectingRegion>
</emphasis>
<collectingMunicipality id="ABBCC512336722B1F0AA8962EE1771FF">Kupe-Mwanenguba</collectingMunicipality>
,
<location id="B75A90D0368B8B2E225CBC262CCF6572" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:B75A90D0368B8B2E225CBC262CCF6572" country="United States of America" latitude="5.0" longLatPrecision="1305" longitude="9.633333" municipality="Kupe-Mwanenguba" name="Nyale" stateProvince="Southwest Region">Nyale</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EHRAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.633333,5.000000]}">
<geoCoordinate id="FACBCA5E400A2A5038CD8624ABDC1AFE" degrees="5" direction="north" minutes="00" orientation="latitude" precision="925" value="5.0">5°00N</geoCoordinate>
,
<geoCoordinate id="FDC88D81AFB4BD21579DA5AA306B3866" degrees="9" direction="east" minutes="38" orientation="longitude" precision="925" value="9.633333">9°38E</geoCoordinate>
</named-content>
;
<date id="FCBACFF13EB29A80CBE635D6F3F76E0A" value="1998-11-18">
<collectingDate id="68335CE79E13C85C9293B38FC981252C" value="1998-11-18">18 Nov. 1998</collectingDate>
</date>
;
<emphasis id="B1682B3757699B6A5B23261BE1946843" italics="true">
<collectorName id="6A55A920275D4405C923EEAC6807766D">Cheek</collectorName>
9678
</emphasis>
;
<collectionCode id="6F287E8B99E917D7A2AECB8B64F95A22">K</collectionCode>
n. v.,
<collectionCode id="342AE78403515FD3D9D4400C3D08C3D3" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
n. v.,
<collectionCode id="21471A4771A0973FB7086A6272A2DDB9" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
n. v
</materialsCitation>
. •
<materialsCitation id="0A474767457F8BA61516749F2038AA90" ID-GBIF-Occurrence="4985319328" collectingDate="1999-12-11" collectionCode="K, YA" collectorName="Cheek" country="United States of America" county="Kupe-Mwanenguba" latitude="4.9166665" location="Abang road and then right to forest" longLatPrecision="1306" longitude="9.733334" municipality="Ngomboku" specimenCount="1">
<collectingCounty id="154C0CB42C26B4BB3F7206CE0779F119">Kupe-Mwanenguba</collectingCounty>
,
<collectingMunicipality id="7B978E27F518EDB31AD3BE7E2B450126">Ngomboku</collectingMunicipality>
,
<location id="2E4E080A43235BD64E6406470D83C722" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:2E4E080A43235BD64E6406470D83C722" country="United States of America" county="Kupe-Mwanenguba" latitude="4.9166665" longLatPrecision="1306" longitude="9.733334" municipality="Ngomboku" name="Abang road and then right to forest">Abang road and then right to forest</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERRAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.733333,4.916667]}">
<geoCoordinate id="740E716C60A05F3831B93927EB2C3F9D" degrees="4" direction="north" minutes="55" orientation="latitude" precision="925" value="4.9166665">4°55N</geoCoordinate>
,
<geoCoordinate id="80024993177C0E73E1192967768E2000" degrees="9" direction="east" minutes="44" orientation="longitude" precision="925" value="9.733334">9°44E</geoCoordinate>
</named-content>
;
<date id="4E435C7A1B04AEC09E2D33FE9E7AA8A4" value="1999-12-11">
<collectingDate id="B7E11ABC5301F27DFB16EDDB2BCD48AA" value="1999-12-11">11 Dec. 1999</collectingDate>
</date>
;
<emphasis id="163F4559FE16B4C780370DF5E0FB8303" italics="true">
<collectorName id="C6D048EC463AFA0EFF25713792DC76D7">Cheek</collectorName>
10326
</emphasis>
;
<collectionCode id="A167D703612894D7EAD48539CFA2A163">K</collectionCode>
n. v.,
<collectionCode id="2860FDE2308F712F7E2F5A8A5C93BEC8" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
n. v
</materialsCitation>
. •
<materialsCitation id="79092D3E3D39DFEDFAA0BC1EFDB2A1E0" ID-GBIF-Occurrence="4985319305" collectingDate="1999-12-15" collectionCode="YA" collectorName="Cheek" country="United States of America" county="Kupe-Mwanenguba" latitude="4.9166665" location="forest ridge S of Abang track" longLatPrecision="1306" longitude="9.733334" municipality="Ngomboku" specimenCount="1">
<collectingCounty id="BEDB519DEA222D2E69E3C44B728BD002">Kupe-Mwanenguba</collectingCounty>
,
<collectingMunicipality id="8C78E99CEF7E517FA234C58163B4CE2F">Ngomboku</collectingMunicipality>
,
<location id="127A461F85BCCB7B177E44F38146F1F3" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:127A461F85BCCB7B177E44F38146F1F3" country="United States of America" county="Kupe-Mwanenguba" latitude="4.9166665" longLatPrecision="1306" longitude="9.733334" municipality="Ngomboku" name="forest ridge S of Abang track">forest ridge S of Abang track</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2RAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.733333,4.916667]}">
<geoCoordinate id="80E2C2A22E3E5BFF717DC66A9D7A41FB" degrees="4" direction="north" minutes="55" orientation="latitude" precision="925" value="4.9166665">4°55N</geoCoordinate>
,
<geoCoordinate id="0198D4DDF2EDCEF8FE0CC9E6BFD7C69C" degrees="9" direction="east" minutes="44" orientation="longitude" precision="925" value="9.733334">9°44E</geoCoordinate>
</named-content>
;
<date id="2B00E21ADE0871F210044A0121A8C436" value="1999-12-15">
<collectingDate id="2419D859B2380A2EF39A07DBD0F78CD7" value="1999-12-15">15 Dec. 1999</collectingDate>
</date>
;
<emphasis id="9979061D68D5E729EA1B01A302D26D7D" italics="true">
<collectorName id="F1CA4738472A1ADCD462ED29025362E1">Cheek</collectorName>
10382
</emphasis>
; K n. v.,
<collectionCode id="B045BC45C863CB53869FB07A2B732B77" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
n. v
</materialsCitation>
. •
<materialsCitation id="C38BD03E86D3DDF521793EE0DFD82F87" ID-GBIF-Occurrence="4985319333" collectingDate="1976-03-24" collectionCode="BR, HBG, K, P, WAG, YA" collectorName="Letouzey" country="United States of America" county="Rumpi Mts" latitude="4.9" location="Mt Rata" longLatPrecision="1306" longitude="9.233334" municipality="Rumpi Mts" specimenCount="1">
<locationDeviation id="B388FA6B497C51B2F8729B36EA52286F" location="Kumba">
<quantity id="6A47B10127DEDC52A6DCF42B4CA4EA68" metricMagnitude="4" metricUnit="m" metricValue="3.5" unit="km" value="35.0">35 km</quantity>
NNW Kumba
</locationDeviation>
,
<locationDeviation id="2B0AE85930CDF00465F835FF8B65FE5B">
<quantity id="010695E0D3ACE5AAD33DD939C44DB15A" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
SW Dikome Balue
</locationDeviation>
,
<collectingCounty id="D1F4C11D595977223EE5FF02ED7E0DEE">Rumpi Mts</collectingCounty>
,
<location id="0605162895C78C0ABBBDFF4D00458671" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:0605162895C78C0ABBBDFF4D00458671" country="United States of America" county="Rumpi Mts" latitude="4.9" longLatPrecision="1306" longitude="9.233334" municipality="Rumpi Mts" name="Mt Rata">Mt Rata</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFSAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.233333,4.900000]}">
<geoCoordinate id="6F334E808EED633C1E1A13BF9F0E0B59" degrees="4" direction="north" minutes="54" orientation="latitude" precision="925" value="4.9">4°54N</geoCoordinate>
,
<geoCoordinate id="FB1CEC8CD68FE2520247EF5EB28F0E1A" degrees="9" direction="east" minutes="14" orientation="longitude" precision="925" value="9.233334">9°14E</geoCoordinate>
</named-content>
;
<date id="A27FD898AB30F341747A03649B43E147" value="1976-03-24">
<collectingDate id="5609A0810133C6560061375495EB54CE" value="1976-03-24">24 Mar. 1976</collectingDate>
</date>
;
<emphasis id="5E4C0A90D05A4E7F0673715463AB70E0" italics="true">
<collectorName id="6F920A242D6BA56C2C1946CC4EECEE34">Letouzey</collectorName>
14546
</emphasis>
;
<collectionCode id="CE285F896C3230057D9E0FA4141F6E5B" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
n. v.,
<collectionCode id="A0C978D481408D33AB9D7652AFCBFEBA">HBG</collectionCode>
n. v.,
<collectionCode id="25CEC485F72EEE11FEE44947A786384A">K</collectionCode>
n. v.,
<collectionCode id="1F4482508B9127CA1F17A402D873E0D4">P</collectionCode>
n. v.,
<collectionCode id="D8AF69D074CFBC34CF450E586FCEAC67" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
n. v.,
<collectionCode id="5B3B1596DA09B1389A67C19707783DA6" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
n. v
</materialsCitation>
. •
<materialsCitation id="B248EEB00207DC9A9368C506D1E6EA99" ID-GBIF-Occurrence="4985319310" collectingDate="1976-03-24" collectionCode="BR, K, P, WAG, YA" collectorName="Letouzey" country="United States of America" county="Dikome Balue" latitude="4.9" location="Letouzey" longLatPrecision="1306" longitude="9.233334" municipality="Same" specimenCount="1">
Same data as for preceding;
<emphasis id="006A5C3CC5FF89ADB3C1C6EBEE54FFC5" italics="true">
<collectorName id="B27C981334D22F26219DA339FADEEB5D">Letouzey</collectorName>
15185
</emphasis>
;
<collectionCode id="73290B448D731AF7AB184C41A52FF4C7" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
n. v.,
<collectionCode id="A9664DF745408F81A3FD0FF00097DE08">K</collectionCode>
n. v.,
<collectionCode id="C5C30AFF611D6D097807F28F0B230CB9">P</collectionCode>
n. v.,
<collectionCode id="1EFE3A9E6F748C16E6002840ECB4DAE0" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
n. v.,
<collectionCode id="0FDDFC700AD7B6B5D0E077CA749474F7" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
</materialsCitation>
<materialsCitation id="5735003C1329410BC3AF659BEAF45523" ID-GBIF-Occurrence="4985319312" collectingDate="1991-11" collectionCode="K, WAG" collectorName="McKey Rh" country="United States of America" county="Rumpi Hills" latitude="4.9" location="near Dikome (Bakume) Balue" longLatPrecision="1306" longitude="9.266666" municipality="Dikome" specimenCode="K 000093017" specimenCount="1">
<collectingCounty id="E4D180E81BC230EC6DF77F881EF968E0">Rumpi Hills</collectingCounty>
,
<location id="33F930EED6C3D50CF277D978B8B39010" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:33F930EED6C3D50CF277D978B8B39010" country="United States of America" county="Rumpi Hills" latitude="4.9" longLatPrecision="1306" longitude="9.266666" municipality="Dikome" name="near Dikome (Bakume) Balue">near Dikome (Bakume) Balue</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ERSAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.266667,4.900000]}">
<geoCoordinate id="E9253F215A863D7B0722BA30A7A7B632" degrees="4" direction="north" minutes="54" orientation="latitude" precision="925" value="4.9">4°54N</geoCoordinate>
,
<geoCoordinate id="218593F579D6BC0BC61BE98A362BBBBA" degrees="9" direction="east" minutes="16" orientation="longitude" precision="925" value="9.266666">9°16E</geoCoordinate>
</named-content>
;
<date id="007B782B3F3845122413398B7E5D722B" value="1991-11">
<collectingDate id="7763724630ED756D197FE13356E73BE3" value="1991-11">Nov. 1991</collectingDate>
</date>
;
<emphasis id="50982B4804C15DE0D005679881142511" italics="true">
<collectorName id="54BFC9ED773AAD4311CD5622836DCF7E">McKey Rh</collectorName>
2 / 43
</emphasis>
;
<collectionCode id="0D99243AD850D1EC064B630534A81C9D">K</collectionCode>
[
<specimenCode id="9A2598F126D69AB01E3082A45BB43B8A">K 000093017</specimenCode>
] n. v.,
<collectionCode id="BB92396D8CC7CA658290310FDE736445" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
n. v
</materialsCitation>
. •
<materialsCitation id="AA4BF6575F0F9F5515B4C6C1925CBAFB" ID-GBIF-Occurrence="4985319307" collectingDate="1987-05-07" collectionCode="K, MO" collectorName="Thomas" country="United States of America" elevation="1100" latitude="4.8333335" location="Rumpi Hills" longLatPrecision="1306" longitude="9.25" specimenCount="1">
<location id="340F91AA80D25FD0FF6A7EFD5280D77F" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:340F91AA80D25FD0FF6A7EFD5280D77F" country="United States of America" latitude="4.8333335" longLatPrecision="1306" longitude="9.25" name="Rumpi Hills">Rumpi Hills</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2SAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.250000,4.833333]}">
<geoCoordinate id="6A0866EB5A7BC93110F23747760FDF19" degrees="4" direction="north" minutes="50" orientation="latitude" precision="925" value="4.8333335">4°50N</geoCoordinate>
,
<geoCoordinate id="46695185FFDD6A0229841BF8DB771B27" degrees="9" direction="east" minutes="15" orientation="longitude" precision="925" value="9.25">9°15E</geoCoordinate>
</named-content>
;
<quantity id="80DC73401542BEC25088D53BC211F657" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
<elevation id="1EB127B7B39F35F4FEE43427D46702D0" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">10001200 m</elevation>
</quantity>
;
<date id="E32373ED0238267D2B7605A9C78DB19A" value="1987-05-07">
<collectingDate id="BB0BF8B6921F800FECED1701EC60707E" value="1987-05-07">7 May 1987</collectingDate>
</date>
;
<emphasis id="73D12DD50FC3E15524B09F343DAF9D09" italics="true">
<collectorName id="B2F6052329231A38BAEB94856FE6BD9A">Thomas</collectorName>
7449
</emphasis>
;
<collectionCode id="346A405A0FF7C45F28CD94946E4AFD08">K</collectionCode>
,
<collectionCode id="0C6C3CAB384796DBEF5D5CA2649DA184" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" type="Herbarium">MO</collectionCode>
n. v
</materialsCitation>
. •
<materialsCitation id="04913C00CE09B6D4089085DD0FC745CC" ID-GBIF-Occurrence="4985319308" collectingDate="1995-02-10" collectionCode="K, WAG, YA" collectorName="Thomas" country="United States of America" latitude="4.9166665" location="Rumpi Hills" longLatPrecision="1306" longitude="9.25" specimenCount="1">
<location id="06474F2E54CB932C5925E2A84283F045" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:06474F2E54CB932C5925E2A84283F045" country="United States of America" latitude="4.9166665" longLatPrecision="1306" longitude="9.25" name="Rumpi Hills">Rumpi Hills</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EFTAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.250000,4.916667]}">
<geoCoordinate id="8F64502188FB4C9C9E9AB31156AC376E" degrees="4" direction="north" minutes="55" orientation="latitude" precision="925" value="4.9166665">4°55N</geoCoordinate>
,
<geoCoordinate id="E7403C1E75CE08CE1B3FD3F6C0FE1537" degrees="9" direction="east" minutes="15" orientation="longitude" precision="925" value="9.25">9°15E</geoCoordinate>
</named-content>
;
<date id="6EF9572BB7288FAED1E58AAE45ADA77A" value="1995-02-10">
<collectingDate id="C395917599EAC7722C47D57701A7B3A2" value="1995-02-10">10 Feb. 1995</collectingDate>
</date>
;
<emphasis id="C4D05EE2FCFBA9DC6775085F56BC4365" italics="true">
<collectorName id="455305623C28B66253F89DF2D150B93D">Thomas</collectorName>
10302
</emphasis>
;
<collectionCode id="A69D3B5A0DA32B4C0AAF98634E15EE5C">K</collectionCode>
,
<collectionCode id="17BE6FB4D7BBF3A392B97C30B04DD152" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
n. v.,
<collectionCode id="E370167C90F5B8F2A26FAAAE98449C0F" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
n. v
</materialsCitation>
. •
<materialsCitation id="F876BF54DE691405C419CA2FE714F8F1" ID-GBIF-Occurrence="4985319331" collectingDate="1995-02-22" collectionCode="BR, K, MO, P, WAG, YA" collectorName="Thomas" country="United States of America" latitude="4.9666667" location="Rumpi Hills near Madie River" longLatPrecision="1306" longitude="9.25" municipality="Rumpi Hills" specimenCount="1">
<location id="3510CFA40056F3D5917A001C690C3083" LSID="urn:lsid:plazi:treatment:2D4B8EC9FE405B6FB5F0E2187DAAC615:3510CFA40056F3D5917A001C690C3083" country="United States of America" latitude="4.9666667" longLatPrecision="1306" longitude="9.25" municipality="Rumpi Hills" name="Rumpi Hills near Madie River">Rumpi Hills near Madie River</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPTAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[9.250000,4.966667]}">
<geoCoordinate id="788F185C7B45B6705103E0D9FF9D0C12" degrees="4" direction="north" minutes="58" orientation="latitude" precision="925" value="4.9666667">4°58N</geoCoordinate>
,
<geoCoordinate id="C8094BB58C03FD2A33A6A8E268E16CFE" degrees="9" direction="east" minutes="15" orientation="longitude" precision="925" value="9.25">9°15E</geoCoordinate>
</named-content>
;
<date id="13EF5B44928CC01100E713B5808B2787" value="1995-02-22">
<collectingDate id="D588E964245C505E57A08007228A4D08" value="1995-02-22">22 Feb. 1995</collectingDate>
</date>
;
<emphasis id="77DD8EE36E0DCB738EE929B2601C5677" italics="true">
<collectorName id="84BEF3660E3B455C743410B81A450AF7">Thomas</collectorName>
10443
</emphasis>
;
<collectionCode id="6AD69FFB89C2C8A2B0E11FDB188E07E1" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
n. v.,
<collectionCode id="6061FE2A32D5DC581267E68907A764E4">K</collectionCode>
,
<collectionCode id="48D1E1135D51309AD2B352E1C654DEB4" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" type="Herbarium">MO</collectionCode>
n. v.,
<collectionCode id="7CA3E1347DCA8099D9B3A06310FA2BA9">P</collectionCode>
n. v.,
<collectionCode id="7269FF970D084A3FC2235A4122ED3EB1" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15578" lsid="urn:lsid:biocol.org:col:15578" name="Wageningen University" type="Herbarium">WAG</collectionCode>
n. v.,
<collectionCode id="56D2B1BCAEB7330C9FD7E02D9044A66C" country="Cameroon" lsid="urn:lsid:biocol.org:col:15278" name="National Herbarium of Cameroon" type="Herbarium">YA</collectionCode>
n. v
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWTAE" type="notes">
<paragraph id="302F862B3B090200D76DCD464B1C3324">
<heading id="0C119A267958A947288E8C235D791561" reason="title">Notes.</heading>
</paragraph>
<paragraph id="64070F80898F8D32C52C72ED2F7068BD">
<taxonomicName id="DA56D07723E0D208DF71635C2FD27ED1" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="15DE4D09A35BE51B568D26F47B53C017" italics="true">Talbotiella bakossiensis</emphasis>
</taxonomicName>
resembles
<taxonomicName id="C024B61F253C747E6F3280434FE71B23" authority="J. Leonard" authorityName="J. Leonard" class="Magnoliopsida" family="Fabaceae" genus="Hymenostegia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="neoaubrevillei">
<emphasis id="971EF0A8C57A85F7267937C900CB15D7" italics="true">Hymenostegia neoaubrevillei</emphasis>
J. Léonard
</taxonomicName>
, a species endemic to
<collectingCountry id="41B6CE0B202F9EC99673DE468FC60437" name="Gabon">Gabon</collectingCountry>
. The most important morphological difference is that petals are absent in
<taxonomicName id="DAA57D18739746380D130A47C1DDCBF8" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="C0F09AD8DABFEB2AF46B2F47CD28632E" italics="true">T. bakossiensis</emphasis>
</taxonomicName>
, while there are 2 petals in
<taxonomicName id="E9A86EAA9D580CDD16228BECB1DFD4D0" class="Magnoliopsida" family="Fabaceae" genus="Hymenostegia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="neoaubrevillei">
<emphasis id="79F5B028C5367E6711ED345DEE1226E3" italics="true">H. neoaubrevillei</emphasis>
</taxonomicName>
. However, the Flore du
<collectingCountry id="86FCB8EA2C3444665D484E53CFE51CE5" name="Gabon">Gabon</collectingCountry>
(
<bibRefCitation id="2DFA10ADE641445F8B1ECB8CBC373140" DOI="10.5281/zenodo.11032174" author="Aubreville" firstAuthor="Aubreville" refId="B1" refString="Aubréville A (1968) Flore du Gabon, Volume 15, Légumineuses Caesalpinioidées. Muséum national dHistoire naturelle, Paris. https://doi.org/10.5281/zenodo.11032174" year="1968">Aubréville 1968</bibRefCitation>
) described the petals of
<taxonomicName id="BAC110990ADA94F086E782B8AC6443A3" class="Magnoliopsida" family="Fabaceae" genus="Hymenostegia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="neoaubrevillei">
<emphasis id="0CCA4B5D8A1137E75D273277A6F6903A" italics="true">H. neoaubrevillei</emphasis>
</taxonomicName>
as “ petals 2, sometimes very reduced or even none at all ”. Many of the collections of
<taxonomicName id="8F4920F5D70300DB4666C0780A222752" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="AC36E136A370B455DE0E5BE71989BBF2" italics="true">T. bakossiensis</emphasis>
</taxonomicName>
, including the
<typeStatus id="371B202F55F0849A7D97EA6AD6EBAE72" type="holotype">holotype</typeStatus>
, have not been seen by us (see Introduction). Our study of the available material reveals several additional differences between the two species.
<taxonomicName id="5C62D61CA4A5CF8EF636278919476640" class="Magnoliopsida" family="Fabaceae" genus="Talbotiella" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="bakossiensis">
<emphasis id="37E9E1614966161CB0747BC08C01A010" italics="true">Talbotiella bakossiensis</emphasis>
</taxonomicName>
has the leaf rachis slightly winged, with flattened, patent margins; the lower leaflets are alternate or subopposite and the bud scales pubescent outside.
<taxonomicName id="3020E9C2A95AB053E27AD5C426F9EBDD" class="Magnoliopsida" family="Fabaceae" genus="Hymenostegia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="neoaubrevillei">
<emphasis id="C405356CBFAA0D36FB0D386531D98C57" italics="true">Hymenostegia neoaubrevillei</emphasis>
</taxonomicName>
has the leaf rachis canaliculate with margins erect, the leaflets opposite and the bud scales mostly glabrous, pubescent only at the base.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:affiliation>Swedish Museum of Natural History, Department of Zoology, Box 50007, SE- 10405 Stockholm, Sweden</mods:affiliation>
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<taxonomicName LSID="2D56F90E-7240-520C-90A6-666B313140D3" authority="Guignot, 1936" authorityName="Guignot" authorityYear="1936" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus ruwenzoricus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ruwenzoricus">Agabus ruwenzoricus Guignot, 1936</taxonomicName>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11H</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">, 13</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">, 14</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Small forest stream at 1900 m in remaining fragment of afromontane cloud forest on Vuria, the highest mountain in the Taita Hills, Taita-Taveta district, northeastern part of Eastern Arc Mountains, Kenya. Albeit minute, the small pools seen in the photo were inhabited by tens of individuals of Agabus ruwenzoricus. January 19, 2016." figureDoi="10.3897/zookeys.963.53470.figure15" httpUri="https://binary.pensoft.net/fig/444412" pageId="0" pageNumber="45">, 15</figureCitation>
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<taxonomicName id="BED4914651A30C765AFFDB06E26894C6" ID-CoL="7QDPX" LSID="2D56F90E-7240-520C-90A6-666B313140D3" authority="Guignot, 1936" authorityName="Guignot" authorityYear="1936" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus ruwenzoricus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ruwenzoricus">Agabus ruwenzoricus Guignot, 1936</taxonomicName>
<figureCitation id="DF967EB69CF25A72E1B2AFD9AC6CCEA4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of the Agabus raffrayi group. Relief is expressed in meters above sea level." figureDoi="10.3897/zookeys.963.53470.figure1" httpUri="https://binary.pensoft.net/fig/444398" pageId="0" pageNumber="45">Figures 1</figureCitation>
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<figureCitation id="74DB5B7F54DE39FFA4246CBD01CEB325" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">, 8C</figureCitation>
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<figureCitation id="592E0C300915D59DD38D398E7960006A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">, 11H</figureCitation>
<figureCitation id="67D238299FA2B70D374C32B368B2F1FE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">, 12</figureCitation>
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<figureCitation id="E604A6A604B095D43049E7B86DDFDFFF" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">, 14</figureCitation>
<figureCitation id="2C0F018FC9D574174DF45F9665B88EE1" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Small forest stream at 1900 m in remaining fragment of afromontane cloud forest on Vuria, the highest mountain in the Taita Hills, Taita-Taveta district, northeastern part of Eastern Arc Mountains, Kenya. Albeit minute, the small pools seen in the photo were inhabited by tens of individuals of Agabus ruwenzoricus. January 19, 2016." figureDoi="10.3897/zookeys.963.53470.figure15" httpUri="https://binary.pensoft.net/fig/444412" pageId="0" pageNumber="45">, 15</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="45">
<taxonomicName class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus (Agabinectes) pallidus subsp. var. var. ruwenzoricus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="variety" species="pallidus" subGenus="Agabinectes" subSpecies="var." variety="ruwenzoricus">Agabus (Agabinectes) pallidus var. ruwenzoricus</taxonomicName>
<subSubSection id="109CA153486C3109E8F266E9BD0D42C5" pageId="0" pageNumber="45" type="reference_group">
<paragraph id="454B736A154E3E20C76E3F79B50D58DF" pageId="0" pageNumber="45">
<taxonomicName id="22D530760E28A325F6DB3A387D15AEE9" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus (Agabinectes) pallidus subsp. var. var. ruwenzoricus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="variety" species="pallidus" subGenus="Agabinectes" subSpecies="var." variety="ruwenzoricus">Agabus (Agabinectes) pallidus var. ruwenzoricus</taxonomicName>
Guignot, 1936: 49
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type locality">
<paragraph pageId="0" pageNumber="45">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="D2881729AC09CA9B668961A1227AE5DD" pageId="0" pageNumber="45" type="type locality">
<paragraph id="DE2C529E5908BB3618BE03FDE7AB12F1" pageId="0" pageNumber="45">Type locality.</paragraph>
<paragraph id="A3810851F7AAE03EDAC1C15654ACE67B" pageId="0" pageNumber="45">
&quot;Uganda. Mons Ruwenzori, versant est, 3.000
<normalizedToken originalValue="à">a</normalizedToken>
<normalizedToken id="A74B05903E185798BA59C3A082CC0570" originalValue="à">a</normalizedToken>
4.000 m.&quot; [Uganda, Mount Ruwenzori, eastern slope, 3000 to 4000 m].
</paragraph>
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<subSubSection pageId="0" pageNumber="45" type="type material">
<paragraph pageId="0" pageNumber="45">Type material.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="034ECD6E4C1841013E5589FAFB8D2C81" pageId="0" pageNumber="45" type="type material">
<paragraph id="7D3506179FBC77F5E888C851F669C591" pageId="0" pageNumber="45">Type material.</paragraph>
<paragraph id="83A13B59A06F0517D68C073BB086080E" pageId="0" pageNumber="45">
[Not examined]: Information about type specimens from
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson (1992a)</bibRefCitation>
<bibRefCitation id="9A0EB714C876A89D6E00CBFFA5691462" DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson (1992a)</bibRefCitation>
: &quot;Lectotype here designated in NMNH (coll. Guignot) labelled: 'Monts Ruwenzori versant est zone alpine 3000 4000 m Ch. Alluaud I 1909', '3000
<normalizedToken originalValue="m">m'</normalizedToken>
<normalizedToken id="A611894EBEE34046FF77B3220063EDE3" originalValue="m">m'</normalizedToken>
,
<normalizedToken originalValue="‘♂’">'♂'</normalizedToken>
<normalizedToken id="3F9A745F0AC051AF686E901B107F7DD2" originalValue="‘♂’">'♂'</normalizedToken>
, and my lectotype label; paralectotype ♂ with same original labels and my paralectotype label.&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="diagnosis">
<paragraph pageId="0" pageNumber="45">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="986C56681CD279115D8FC413EAB79800" pageId="0" pageNumber="45" type="diagnosis">
<paragraph id="44A43C896FB7E34A4D0574207DAE6A26" pageId="0" pageNumber="45">Diagnosis.</paragraph>
<paragraph id="D178B10E7D4465ED5E420B10B98404B1" pageId="0" pageNumber="45">
Most similar to
<taxonomicName lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
<taxonomicName id="D19793E9D8FC6846FADA2FFA1A8574A4" lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="B84FFE1828BFA55E8AA7D91A10141A91" italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
<taxonomicName id="ADEB4FD12EDF17BF05B3854CC7CB2C08" lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis id="CD64B673E2AC52B7F4E2D985947C8967" italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
but separated from these taxa by the presence of distinct interocular spots on head (compare Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agabus spp., head and pronotum in dorsal view A A. dytiscoides B A. ruwenzoricus C A. sjostedti D A. pallidus. Red dashed lines indicate measurements used for interocular distance and pronotal width." figureDoi="10.3897/zookeys.963.53470.figure4" httpUri="https://binary.pensoft.net/fig/444401" pageId="0" pageNumber="45">4B and D</figureCitation>
<figureCitation id="0E4E7067C82286B87A5AB3795AF1902E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Agabus spp., head and pronotum in dorsal view A A. dytiscoides B A. ruwenzoricus C A. sjostedti D A. pallidus. Red dashed lines indicate measurements used for interocular distance and pronotal width." figureDoi="10.3897/zookeys.963.53470.figure4" httpUri="https://binary.pensoft.net/fig/444401" pageId="0" pageNumber="45">4B and D</figureCitation>
). The metasternal wing is rather narrow; the WC/WS frequency distribution being intermediate between
<taxonomicName lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
<taxonomicName id="D8ED3B321B6C0C5B0C317D80E21F1560" lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="50FC351BF60B6E49B6B1BB5AB5B3A52F" italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
<taxonomicName id="64BA558F7B0576B8F47E08FD424E05D7" lsidName="A. raffrayi" pageId="0" pageNumber="45" rank="species" species="raffrayi">
<emphasis id="3ADA74B73C2A51C905A8E199F5AFD28D" italics="true" pageId="0" pageNumber="45">A. raffrayi</emphasis>
</taxonomicName>
, but most similar to
<taxonomicName lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
<taxonomicName id="93BEF75C9C4F36DCA8A20334933B420C" lsidName="A. pallidus" pageId="0" pageNumber="45" rank="species" species="pallidus">
<emphasis id="A910DF6422D5C5A7D8CBBD2890690D0D" italics="true" pageId="0" pageNumber="45">A. pallidus</emphasis>
</taxonomicName>
(see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
<tableCitation id="A0A7F4870E6586386CCEAB3F8FB6BC20" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
, Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
<figureCitation id="2372B22233CFAF377A13CB4E65A24247" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
). The aedeagus has an extended portion between the subapical broadening and the apical teeth, and the pronotal hypomeron is not visible in lateral view.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="0D7D3ED488F9A370AA8E23D6015EE2F5" pageId="0" pageNumber="45" type="description">
<paragraph id="38518E0FA1DC20ECD2CFA306A40EAEE8" pageId="0" pageNumber="45">Description.</paragraph>
<paragraph id="2129660AA5C0A127760648B2E98BCAC4" pageId="0" pageNumber="45">
Habitus as in Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">11C, H</figureCitation>
<figureCitation id="4DF5182A30399C30DE1CE3B6BB43E2A9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Agabus spp., habitus of males in dorsal view A, F A. sjostedti B, G A. dytiscoides; C, H A. ruwenzoricus D, I A. raffrayi E, J A. pallidus K, O A. anguluverpus sp. nov. L, P A. austellus sp. nov. M, Q A. riberae sp. nov. N, R A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure11" httpUri="https://binary.pensoft.net/fig/444408" pageId="0" pageNumber="45">11C, H</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Colour</emphasis>
<paragraph id="9C7AB7402BE1BD6A9CDA13490AF7F971" pageId="0" pageNumber="45">
<emphasis id="D00C9301641151CC50C52F1555B9302D" bold="true" italics="true" pageId="0" pageNumber="45">Colour</emphasis>
: Head black with rufous anterior area; rufous interocular spots present. Pronotum black with minute to well-defined rufous margin. Elytra ferrugineous to rufopiceous. Ventral surface black, hypomeron rufotestaceous to rufous and epipleuron testaceous to rufotestaceous. Legs rufous to black. Antennae and palpi testaceous.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName genus="Microreticulation" lsidName="Microreticulation" pageId="0" pageNumber="45" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Microreticulation</emphasis>
<paragraph id="6453B6999380ACD22A3C33CC1F0E9C76" pageId="0" pageNumber="45">
<taxonomicName id="F0A66C81763E30FBE56792C0C9D29DBD" genus="Microreticulation" lsidName="Microreticulation" pageId="0" pageNumber="45" rank="genus">
<emphasis id="4A5DD27BC033E14B4D6E983862E13FAB" bold="true" italics="true" pageId="0" pageNumber="45">Microreticulation</emphasis>
</taxonomicName>
: Medium impressed on head, pronotum and elytra, and rather similar in both sexes. Composed of a mixture of small and somewhat larger, uneven meshes.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<taxonomicName genus="Structural" lsidName="Structural features" pageId="0" pageNumber="45" rank="species" species="features">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Structural features</emphasis>
<paragraph id="582AF85E4909DE38EDBA1EF520CD89E0" pageId="0" pageNumber="45">
<taxonomicName id="27F0A728241A11680B10DEBC031B25CB" genus="Structural" lsidName="Structural features" pageId="0" pageNumber="45" rank="species" species="features">
<emphasis id="FFC3A08AA65B1727CC448AE1EEA4C618" bold="true" italics="true" pageId="0" pageNumber="45">Structural features</emphasis>
</taxonomicName>
: Body length: 7.36-8.08 mm (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
<tableCitation id="4F2BD1A367122001945ED0DA920DF77F" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
). Hypomeron not visible in strict lateral view (as in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agabus spp., hypomeron in lateral view A A. sjostedti B A. dytiscoides C A. anguluverpus sp. nov. D A. raffrayi. White arrows indicate hypomeron." figureDoi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45">10C, D</figureCitation>
<figureCitation id="542DEAC38533749B2940D6BC2F06F320" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agabus spp., hypomeron in lateral view A A. sjostedti B A. dytiscoides C A. anguluverpus sp. nov. D A. raffrayi. White arrows indicate hypomeron." figureDoi="10.3897/zookeys.963.53470.figure10" httpUri="https://binary.pensoft.net/fig/444407" pageId="0" pageNumber="45">10C, D</figureCitation>
, compare with 10A, B), lateral bead of pronotum narrow and well defined. Metasternal wing narrow, WC/WS 3.0 or more in most specimens (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
<tableCitation id="9A947CAF48099E807B4526CD3C504C8B" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
, Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
<figureCitation id="DA3401D8A3FDE13616877958C5B29447" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
). Pronotum broad, more than twice as broad as interocular distance (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
<tableCitation id="D576092B0C481DA9F1337E7A5DED8C09" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological characters in the Agabus raffrayi group. WC / WS = width of metacoxal plate / width of metasternal wing, PW / ID = width of pronotum / interocular distance, TL = total body length, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/EC20EA9B30576BE179C66F15DBC0C3C6" pageId="0" pageNumber="45" tableUuid="EC20EA9B30576BE179C66F15DBC0C3C6">1</tableCitation>
, Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">13</figureCitation>
<figureCitation id="1DC850F07DD7164B2A3D072B77D1C312" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. The ratio between pronotal width and interocular distance in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the very narrow pronotum of A. dytiscoides, a distinguishing feature of this species." figureDoi="10.3897/zookeys.963.53470.figure13" httpUri="https://binary.pensoft.net/fig/444410" pageId="0" pageNumber="45">13</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Legs</emphasis>
<paragraph id="0FC7600CEDBF2C9CB14EF325A78BCA79" pageId="0" pageNumber="45">
<emphasis id="68B2EDF32DA110FC8F879FB271A4DEF5" bold="true" italics="true" pageId="0" pageNumber="45">Legs</emphasis>
: Protarsal claws long,&gt; 1.6
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="9E42AFAE94D4460F83319FB2D34D7EDE" originalValue="×">x</normalizedToken>
as long as protarsomere 4 in most specimens (see Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
<tableCitation id="CD4053A49B464BA9F9507928245FE2E6" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
, Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">14</figureCitation>
<figureCitation id="D8107BA3D2B3D61776EA82B715F27D64" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. The ratio between length of protarsal claw and length of protarsomere 4 in males of the Agabus raffrayi group. Symbols as in Fig. 12. Note the short claws of A. sjostedti and A. dytiscoides." figureDoi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45">14</figureCitation>
). Metatarsomeres short and broad; metatarsomere 2 &lt;1.8
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="221BBF5E01461F29181D80D43CF077D8" originalValue="×">x</normalizedToken>
as long as broad (see Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
<tableCitation id="477CA4E64C55C6D664F166DDB8211726" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
), metatarsomere 5 &lt;3.3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="FB8FD5E0FFC890A0AE064BE253468722" originalValue="×">x</normalizedToken>
as long as broad (see Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
<tableCitation id="FEB1AA4CFBC6B0AC5F5DAE49C594CF2D" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological characters in the Agabus raffrayi group. MT 2 L / MT 2 W = length / width of metatarsomere 2, MT 5 L / MT 5 W = length / width of metatarsomere 5, CL / PT 4 L = length of protarsal claw / length of protarsomere 4, N = number of specimens, Min = minimum value, Max = maximum value, SD = standard deviation, NA = not applicable." httpUri="http://table.plazi.org/id/59FBA533E2ADB4142B7B5363E75795AF" pageId="0" pageNumber="45" tableUuid="59FBA533E2ADB4142B7B5363E75795AF">2</tableCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" italics="true" pageId="0" pageNumber="45">Male genitalia</emphasis>
<paragraph id="DC8B7D47D96951831940EE282AF205C1" pageId="0" pageNumber="45">
<emphasis id="490B57A6E6439D0D0E3E354A032308C0" bold="true" italics="true" pageId="0" pageNumber="45">Male genitalia</emphasis>
: Subapically broadened and prolonged between the subapical broadening and the apical and subapical teeth (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">8C</figureCitation>
<figureCitation id="BABB9FDEFADFDBF19067FE802B78A796" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agabus spp., aedeagus in lateral view A A. sjostedti B A. dytiscoides C A. ruwenzoricus D A. raffrayi E A. pallidus F A. anguluverpus sp. nov. G A. austellus sp. nov. H A. riberae sp. nov. I A. agulhas sp. nov." figureDoi="10.3897/zookeys.963.53470.figure8" httpUri="https://binary.pensoft.net/fig/444405" pageId="0" pageNumber="45">8C</figureCitation>
). Subapical tooth with varying appearance (similar to magnitude in variation seen in Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agabus spp., lateral view of tip of aedeagus A-C A. raffrayi, different specimens showing the variation in shape of the subapical tooth D A. austellus sp. nov. Note the prolonged section between subapical broadening and base of apical and subapical teeth in A. raffrayi (indicated with orange in the smaller outline illustrations)." figureDoi="10.3897/zookeys.963.53470.figure5" httpUri="https://binary.pensoft.net/fig/444402" pageId="0" pageNumber="45">5A-C</figureCitation>
<figureCitation id="105975D492E9AFF5CFB864B1D060F8F0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Agabus spp., lateral view of tip of aedeagus A-C A. raffrayi, different specimens showing the variation in shape of the subapical tooth D A. austellus sp. nov. Note the prolonged section between subapical broadening and base of apical and subapical teeth in A. raffrayi (indicated with orange in the smaller outline illustrations)." figureDoi="10.3897/zookeys.963.53470.figure5" httpUri="https://binary.pensoft.net/fig/444402" pageId="0" pageNumber="45">5A-C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Female</emphasis>
<paragraph id="23C84B641738334F201453A6E8475348" pageId="0" pageNumber="45">
<emphasis id="FB16BC4DA6EA518667E725452D9DFCBB" bold="true" pageId="0" pageNumber="45">Female</emphasis>
: Externally similar to males.
</paragraph>
<caption doi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45" start="Figure 14" startId="F14">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 14.</emphasis>
<caption id="5A55A4F2E23CBB0AA9A8820CC83F4532" doi="10.3897/zookeys.963.53470.figure14" httpUri="https://binary.pensoft.net/fig/444411" pageId="0" pageNumber="45" start="Figure 14" startId="F14">
<paragraph id="AB65CA7BB8C7182B272D0BC40F8D94E8" pageId="0" pageNumber="45">
<emphasis id="BD4888AA05142659AA793696707B4781" bold="true" pageId="0" pageNumber="45">Figure 14.</emphasis>
The ratio between length of protarsal claw and length of protarsomere 4 in males of the
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus raffrayi</emphasis>
<taxonomicName id="4AAD34FD882F4B2F1EADB60BF6E976D5" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus raffrayi" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="1C0CDFD43A3E8BDE04AD40C83A18D530" italics="true" pageId="0" pageNumber="45">Agabus raffrayi</emphasis>
</taxonomicName>
group. Symbols as in Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
<figureCitation id="9C1C628BD5AAF2A7B712831DFE7BADDE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. The ratio between width of the metacoxal plate and metasternal wing (WC / WS) in the A. raffrayi group (incl. specimens of both sexes). Thick black line inside boxes represents medians, left and right box borders 25 th (Q 1) and 75 th (Q 3) percentiles respectively. Whiskers were calculated with the boxplot. stats function in R using the default coefficient value of 1.5 (drawn to the highest and lowest value within 1.5 * IQD (Inter Quartile Distance = Q 3 - Q 1) away from the 75 th and 25 th percentiles respectively). Note that this character fully separates A. raffrayi from A. pallidus." figureDoi="10.3897/zookeys.963.53470.figure12" httpUri="https://binary.pensoft.net/fig/444409" pageId="0" pageNumber="45">12</figureCitation>
. Note the short claws of
<taxonomicName lsidName="A. sjostedti" pageId="0" pageNumber="45" rank="species" species="sjostedti">
<emphasis italics="true" pageId="0" pageNumber="45">A. sjostedti</emphasis>
<taxonomicName id="A4F2E367C8B2F20BE95845B0EFB1F7E8" lsidName="A. sjostedti" pageId="0" pageNumber="45" rank="species" species="sjostedti">
<emphasis id="D024F11CDE5E72B507FE19226D249207" italics="true" pageId="0" pageNumber="45">A. sjostedti</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="A. dytiscoides" pageId="0" pageNumber="45" rank="species" species="dytiscoides">
<emphasis italics="true" pageId="0" pageNumber="45">A. dytiscoides</emphasis>
<taxonomicName id="5C20594BEB72DDCB52C4B604E00ABAB2" lsidName="A. dytiscoides" pageId="0" pageNumber="45" rank="species" species="dytiscoides">
<emphasis id="4FF6174D1F0618C6D77270237B4EA131" italics="true" pageId="0" pageNumber="45">A. dytiscoides</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="82E6BB99B9124522EAB83DF47FDB3314" pageId="0" pageNumber="45" type="distribution">
<paragraph id="DA1B2A0E9465C2285EC577B66BC34CCE" pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph id="C5B3FF2D1B8646A4F2CDBA4B703CF2F8" pageId="0" pageNumber="45">
Kenya, Rwanda, Uganda and the Democratic Republic of the Congo.
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson (1992a)</bibRefCitation>
<bibRefCitation id="91051CAC019B4F7493C98A6DB1A1FDE8" DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson (1992a)</bibRefCitation>
and
<bibRefCitation author="Omer-Cooper, J" journalOrPublisher="South African Animal Life" pageId="0" pageNumber="45" pagination="59 - 214" refId="B20" refString="Omer-Cooper, J, 1965. Chapter 2. Coleoptera: Dytiscidae. A review of the Dytiscidae of Southern Africa being the results of the Lund University Expedition 1950-1951, with which are incorporated all other records known to the author. South African Animal Life 11: 59 - 214" title="Chapter 2. Coleoptera: Dytiscidae. A review of the Dytiscidae of Southern Africa being the results of the Lund University Expedition 1950 - 1951, with which are incorporated all other records known to the author." volume="11" year="1965">Omer-Cooper (1965)</bibRefCitation>
<bibRefCitation id="B758577814796BACC7FB3ABFE79BD58E" author="Omer-Cooper, J" journalOrPublisher="South African Animal Life" pageId="0" pageNumber="45" pagination="59 - 214" refId="B20" refString="Omer-Cooper, J, 1965. Chapter 2. Coleoptera: Dytiscidae. A review of the Dytiscidae of Southern Africa being the results of the Lund University Expedition 1950-1951, with which are incorporated all other records known to the author. South African Animal Life 11: 59 - 214" title="Chapter 2. Coleoptera: Dytiscidae. A review of the Dytiscidae of Southern Africa being the results of the Lund University Expedition 1950 - 1951, with which are incorporated all other records known to the author." volume="11" year="1965">Omer-Cooper (1965)</bibRefCitation>
also give South Africa and Zimbabwe but these records are likely to belong to other species (see below).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="habitat">
<paragraph pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="45">
<subSubSection id="AE515A32C01E6FE5EB62F2E1F634BA8F" pageId="0" pageNumber="45" type="habitat">
<paragraph id="0FE8A63116C4635A31B2F61298B0863E" pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph id="B66AE06488F244B7666D92596C96B3F4" pageId="0" pageNumber="45">
Most records are from small mountain streams and rivers at elevations of 1900 to 3100 m, but it has also been found in stagnant waterbodies (
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson 1992a</bibRefCitation>
<bibRefCitation id="6B4A28A9D115D7D4160A866B47A68D9C" DOI="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" author="Nilsson, AN" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="45" pagination="155 - 179" refId="B15" refString="Nilsson, AN, 1992a. A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists. Systematic Entomology 17: 155 - 179, DOI: https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" title="A revision of Afrotropical Agabus Leach (Coleoptera, Dytiscidae), and the evolution of tropicoalpine super specialists." url="https://doi.org/10.1111/j.1365-3113.1992.tb00328.x" volume="17" year="1992 a">Nilsson 1992a</bibRefCitation>
). We found the species in a very small cold-water forest stream at an elevation of 1900 m in the Taita Hills, Kenya (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Small forest stream at 1900 m in remaining fragment of afromontane cloud forest on Vuria, the highest mountain in the Taita Hills, Taita-Taveta district, northeastern part of Eastern Arc Mountains, Kenya. Albeit minute, the small pools seen in the photo were inhabited by tens of individuals of Agabus ruwenzoricus. January 19, 2016." figureDoi="10.3897/zookeys.963.53470.figure15" httpUri="https://binary.pensoft.net/fig/444412" pageId="0" pageNumber="45">15</figureCitation>
<figureCitation id="5FB8B381B66BE8B4853579C4A7A60475" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Small forest stream at 1900 m in remaining fragment of afromontane cloud forest on Vuria, the highest mountain in the Taita Hills, Taita-Taveta district, northeastern part of Eastern Arc Mountains, Kenya. Albeit minute, the small pools seen in the photo were inhabited by tens of individuals of Agabus ruwenzoricus. January 19, 2016." figureDoi="10.3897/zookeys.963.53470.figure15" httpUri="https://binary.pensoft.net/fig/444412" pageId="0" pageNumber="45">15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">The name refers to the locality where the species was first collected, Mount Ruwenzori.</paragraph>
<caption doi="10.3897/zookeys.963.53470.figure15" httpUri="https://binary.pensoft.net/fig/444412" pageId="0" pageNumber="45" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figure 15.</emphasis>
<subSubSection id="C44F3F25335DD3799DA657B8373A1658" pageId="0" pageNumber="45" type="etymology">
<paragraph id="7B59466D3B97AC41C21FFB74D7409F0C" pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph id="0F5E0510D66E15024D75A01758561C6D" pageId="0" pageNumber="45">The name refers to the locality where the species was first collected, Mount Ruwenzori.</paragraph>
<caption id="25CB96FFDEF0F79D57588F58E53CFB7E" doi="10.3897/zookeys.963.53470.figure15" httpUri="https://binary.pensoft.net/fig/444412" pageId="0" pageNumber="45" start="Figure 15" startId="F15">
<paragraph id="CE0E7A278267AADF5AC9F57D14843B78" pageId="0" pageNumber="45">
<emphasis id="2A3DB19D69A1FF1AC879A19F861DD410" bold="true" pageId="0" pageNumber="45">Figure 15.</emphasis>
Small forest stream at 1900 m in remaining fragment of afromontane cloud forest on Vuria, the highest mountain in the Taita Hills, Taita-Taveta district, northeastern part of Eastern Arc Mountains, Kenya. Albeit minute, the small pools seen in the photo were inhabited by tens of individuals of
<taxonomicName authorityName="Guignot" authorityYear="1936" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus ruwenzoricus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ruwenzoricus">
<emphasis italics="true" pageId="0" pageNumber="45">Agabus ruwenzoricus</emphasis>
<taxonomicName id="5C1EB6CADB8E19F5890177C4658C8A54" authorityName="Guignot" authorityYear="1936" class="Insecta" family="Dytiscidae" genus="Agabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agabus ruwenzoricus" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ruwenzoricus">
<emphasis id="72D4238BB48FD3FCA2B1A294512F9167" italics="true" pageId="0" pageNumber="45">Agabus ruwenzoricus</emphasis>
</taxonomicName>
. January 19, 2016.
</paragraph>

View file

@ -1,47 +1,48 @@
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@ -384,7 +385,7 @@
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<taxonomicName id="62F54ABD315D7B30FF76F427FD39FCA0" authority="Motschulsky, 1857: 37" authorityName="Motschulsky" authorityPageNumber="37" authorityYear="1857" box="[151,760,780,802]" class="Insecta" genus="Raphidopalpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="femoralis">
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Motschulsky, 1857: 37 (
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/ labelled by MNHUB 2014 [r, p]”.
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: not studied. They should be deposited at the ZMUM.

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@ -1,219 +1,219 @@
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<mods:namePart id="BEA9E114C272DBF45132479D03BA9203">Gastineau, Romain</mods:namePart>
<mods:affiliation id="B9576C82872DB599A934A747CDCF5075">University of Szczecin, Institute of Marine and Environmental Sciences, Mickiewicza 16 A, 70 - 383 Szczecin, Poland &amp; Natural Sciences Education and Research Centre, University of Szczecin, Mickiewicza 16 a, 70 - 383 Szczecin, Poland</mods:affiliation>
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</mods:role>
<mods:namePart>Duong, Thi Thuy</mods:namePart>
<mods:affiliation>Institute of Environmental Technology, Vietnam Academy of Sciences and Technology, 18 Hoang Quoc Viet Road, Cau Giay, Hanoi, Vietnam</mods:affiliation>
<mods:namePart id="26AB133FEAE74E886D5F084CC2F3A81F">Duong, Thi Thuy</mods:namePart>
<mods:affiliation id="D7ECA0117A8452EBCF83B8F0CD35C8BA">Institute of Environmental Technology, Vietnam Academy of Sciences and Technology, 18 Hoang Quoc Viet Road, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart>Rosa, Philippe</mods:namePart>
<mods:affiliation>Universite de Nantes, EA 2160 Mer - Molecules - Sante 2, Rue de la Houssiniere, 44322 Nantes, France</mods:affiliation>
<mods:namePart id="9F85E8B73486249EE9799FF3971FB40B">Rosa, Philippe</mods:namePart>
<mods:affiliation id="0C8F85847C1EFC6184670756F37BF538">Universite de Nantes, EA 2160 Mer - Molecules - Sante 2, Rue de la Houssiniere, 44322 Nantes, France</mods:affiliation>
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</mods:role>
<mods:namePart>Meleder, Vona</mods:namePart>
<mods:affiliation>Universite de Nantes, EA 2160 Mer - Molecules - Sante 2, Rue de la Houssiniere, 44322 Nantes, France</mods:affiliation>
<mods:namePart id="3EF73BA11D906B857D7A4535F0AA175C">Meleder, Vona</mods:namePart>
<mods:affiliation id="B35858C831407139F48DE329234F989A">Universite de Nantes, EA 2160 Mer - Molecules - Sante 2, Rue de la Houssiniere, 44322 Nantes, France</mods:affiliation>
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<mods:title id="D27AB00991D115BF19B25DDB2292F126">PhytoKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-10-22</mods:number>
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<mods:date id="A937729C6BDCAB8DB46ABEDA0A07C45C">2021</mods:date>
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<mods:number id="3ADD6E48086AF762A232EC4ABBFF46F0">2021-10-22</mods:number>
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<mods:number>183</mods:number>
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<mods:number id="B0717C12B49FA23D7C4C7EC924381BAE">183</mods:number>
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<mods:start>115</mods:start>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.183.71049</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.183.71049</mods:identifier>
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<treatment ID-GBIF-Taxon="189468321" LSID="urn:lsid:plazi:treatment:2D953ADA039A58A3A7C84099BC9D9D90" httpUri="http://treatment.plazi.org/id/2D953ADA039A58A3A7C84099BC9D9D90" lastPageNumber="115" pageId="0" pageNumber="115">
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<paragraph pageId="0" pageNumber="115">
<taxonomicName LSID="2D953ADA-039A-58A3-A7C8-4099BC9D9D90" authority="(Hustedt) D. G. Mann" authorityName="(Hustedt) D. G. Mann" class="Bacillariophyceae" family="Diadesmidaceae" genus="Luticola" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Luticola inserata" order="Naviculales" pageId="0" pageNumber="115" phylum="Bacillariophyta" rank="species" species="inserata">Luticola inserata (Hustedt) D.G.Mann</taxonomicName>
<treatment id="2D953ADA039A58A3A7C84099BC9D9D90" ID-GBIF-Taxon="189468321" LSID="urn:lsid:plazi:treatment:2D953ADA039A58A3A7C84099BC9D9D90" httpUri="http://treatment.plazi.org/id/2D953ADA039A58A3A7C84099BC9D9D90" lastPageNumber="115" pageId="0" pageNumber="115" scope_class="Bacillariophyceae" scope_order="Naviculales" scope_phylum="Bacillariophyta">
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<taxonomicName id="CCE72339ED08E53D1CB685C6F37B74A5" LSID="2D953ADA-039A-58A3-A7C8-4099BC9D9D90" authority="(Hustedt) D. G. Mann" authorityName="(Hustedt) D. G. Mann" class="Bacillariophyceae" family="Diadesmidaceae" genus="Luticola" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Luticola inserata" order="Naviculales" pageId="0" pageNumber="115" phylum="Bacillariophyta" rank="species" species="inserata">Luticola inserata (Hustedt) D.G.Mann</taxonomicName>
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<paragraph pageId="0" pageNumber="115">
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. LM micrographs of Luticola inserata (Hustedt) D. G. Mann in size diminution series (A-R). Scale bar: 10 µm." figureDoi="10.3897/phytokeys.183.71049.figure8" httpUri="https://binary.pensoft.net/fig/601479" pageId="0" pageNumber="115">Figures 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM micrographs of Luticola inserata (Hustedt) D. G. Mann. External view of valve (A, B). Detailed external views showing proximal raphe endings with irregular shallow grooves C distal raphe ending, areolae D stigma opening E internal view of valve F detailed internal view showing proximal raphe endings and stigma opening G and distal raphe ending and hymenate structure H open copulae I. Scale bars: 10 µm (A, F, I), 5 µm (B-D, G), 3 µm (E), 4 µm (H)." figureDoi="10.3897/phytokeys.183.71049.figure9" httpUri="https://binary.pensoft.net/fig/601480" pageId="0" pageNumber="115">, 9</figureCitation>
<subSubSection id="E649FC81D446DF947CE7ED53BF9087DF" pageId="0" pageNumber="115" type="description">
<paragraph id="1D4142C429EC8AC12842B9E8D612BF0D" pageId="0" pageNumber="115">
<figureCitation id="9CDFC8E48743ED6523B63A8AD99DD806" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. LM micrographs of Luticola inserata (Hustedt) D. G. Mann in size diminution series (A-R). Scale bar: 10 µm." figureDoi="10.3897/phytokeys.183.71049.figure8" httpUri="https://binary.pensoft.net/fig/601479" pageId="0" pageNumber="115">Figures 8</figureCitation>
<figureCitation id="8AB96AD38A38BE2C467CB5586B8B1418" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. SEM micrographs of Luticola inserata (Hustedt) D. G. Mann. External view of valve (A, B). Detailed external views showing proximal raphe endings with irregular shallow grooves C distal raphe ending, areolae D stigma opening E internal view of valve F detailed internal view showing proximal raphe endings and stigma opening G and distal raphe ending and hymenate structure H open copulae I. Scale bars: 10 µm (A, F, I), 5 µm (B-D, G), 3 µm (E), 4 µm (H)." figureDoi="10.3897/phytokeys.183.71049.figure9" httpUri="https://binary.pensoft.net/fig/601480" pageId="0" pageNumber="115">, 9</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="115">Description LM.</paragraph>
<paragraph pageId="0" pageNumber="115">
<subSubSection id="AC3D5D12FB2C8ADF35217C1A2077155E" pageId="0" pageNumber="115" type="description">
<paragraph id="5DA1C07EDC7A59CACAADE33DD25CFC2F" pageId="0" pageNumber="115">Description LM.</paragraph>
<paragraph id="3B3215D8BB693519D73F66335184F51A" pageId="0" pageNumber="115">
Valves lanceolate-elliptical to broadly elliptic with weakly undulated margins with rounded, rostrate to capitate apices, 12.2-33.5
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="506E797FA74B1E3948CDE7E5EF5ADE2B" originalValue="μm">μm</normalizedToken>
in length, 8.2-14.0
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="06DC5A5492A876EB844B86BBD177FE1B" originalValue="μm">μm</normalizedToken>
in width (n = 20). Axial area narrow, gradually broadening towards valve center, central area rectangular, asymmetrical, bordered by two or three shortened striae with slit-like stigma located close to valve margin. Transapical striae radiate, becoming strongly radiate toward apices, 15-20 in 10
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="099D6EBB6FD3E7C147343F597F6FA57A" originalValue="μm">μm</normalizedToken>
. Copulae open.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="description">
<paragraph pageId="0" pageNumber="115">Description SEM.</paragraph>
<paragraph pageId="0" pageNumber="115">Valve face flat, raphe branches filiform and straight. External proximal raphe endings clearly bent to the site opposite the stigma with irregularly-shaped grooves expanded opposite the stigma. External distal raphe endings terminate on apices, strongly hooked. Internally, raphe branches straight, only proximal endings slightly bent, distal raphe endings terminating in small helictoglossae. Transapical striae composed of 4-5 round to elongate areolae. Single row of elongate areolae occurs also on valve mantle. Both areolae on mantle and valve face with small spines on edges. Internally, areolae covered with hymen forming continuous strips. Ghost areolae rarely present within central area. External opening of stigma small and rounded, positioned very close to valve margin. Internal stigma opening with large-lipped structure positioned mid-way between valve margin and valve center. Internally, longitudinal channel present along the valve margin, with small silica flap on side opposite the stigma.</paragraph>
<caption doi="10.3897/phytokeys.183.71049.figure8" httpUri="https://binary.pensoft.net/fig/601479" pageId="0" pageNumber="115" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="115">
<emphasis bold="true" pageId="0" pageNumber="115">Figure 8.</emphasis>
<subSubSection id="2ECAE2F854DD35222E33C105758B806B" pageId="0" pageNumber="115" type="description">
<paragraph id="A60E2A93B054A1D10112E11EF58AF60D" pageId="0" pageNumber="115">Description SEM.</paragraph>
<paragraph id="D3151071A6F7DABC4057AB9A9C39D73F" pageId="0" pageNumber="115">Valve face flat, raphe branches filiform and straight. External proximal raphe endings clearly bent to the site opposite the stigma with irregularly-shaped grooves expanded opposite the stigma. External distal raphe endings terminate on apices, strongly hooked. Internally, raphe branches straight, only proximal endings slightly bent, distal raphe endings terminating in small helictoglossae. Transapical striae composed of 4-5 round to elongate areolae. Single row of elongate areolae occurs also on valve mantle. Both areolae on mantle and valve face with small spines on edges. Internally, areolae covered with hymen forming continuous strips. Ghost areolae rarely present within central area. External opening of stigma small and rounded, positioned very close to valve margin. Internal stigma opening with large-lipped structure positioned mid-way between valve margin and valve center. Internally, longitudinal channel present along the valve margin, with small silica flap on side opposite the stigma.</paragraph>
<caption id="920E930D62E4B49B89946563C19CBB76" doi="10.3897/phytokeys.183.71049.figure8" httpUri="https://binary.pensoft.net/fig/601479" pageId="0" pageNumber="115" start="Figure 8" startId="F8">
<paragraph id="CC8D9AAD496628434F149683D97F85A3" pageId="0" pageNumber="115">
<emphasis id="BA292271EB8DA1E5E3F82E5145A24CBE" bold="true" pageId="0" pageNumber="115">Figure 8.</emphasis>
LM micrographs of
<taxonomicName class="Bacillariophyceae" family="Diadesmidaceae" genus="Luticola" kingdom="Plantae" lsidName="Luticola inserata" order="Naviculales" pageId="0" pageNumber="115" phylum="Bacillariophyta" rank="species" species="inserata">
<emphasis italics="true" pageId="0" pageNumber="115">Luticola inserata</emphasis>
<taxonomicName id="A8A8525C374D146BCDAB2A7E0E1EFF9E" class="Bacillariophyceae" family="Diadesmidaceae" genus="Luticola" kingdom="Plantae" lsidName="Luticola inserata" order="Naviculales" pageId="0" pageNumber="115" phylum="Bacillariophyta" rank="species" species="inserata">
<emphasis id="F912FC0906D817F80A34C6ECBB702DFC" italics="true" pageId="0" pageNumber="115">Luticola inserata</emphasis>
</taxonomicName>
(Hustedt) D.G.Mann in size diminution series (
<emphasis bold="true" pageId="0" pageNumber="115">A-R</emphasis>
<emphasis id="55C7E378AF404AB74A39C2735E961A94" bold="true" pageId="0" pageNumber="115">A-R</emphasis>
). Scale bar: 10
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="8DB8C58D595EDC0359FDC705E94FE0A5" originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.183.71049.figure9" httpUri="https://binary.pensoft.net/fig/601480" pageId="0" pageNumber="115" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="115">
<emphasis bold="true" pageId="0" pageNumber="115">Figure 9.</emphasis>
<caption id="5B74047A1BAB20A2A60E46E17D6FA014" doi="10.3897/phytokeys.183.71049.figure9" httpUri="https://binary.pensoft.net/fig/601480" pageId="0" pageNumber="115" start="Figure 9" startId="F9">
<paragraph id="C583179101BAED277A0E4A3A13E027BF" pageId="0" pageNumber="115">
<emphasis id="38F56155468FB2DAF5ABE62879953AE2" bold="true" pageId="0" pageNumber="115">Figure 9.</emphasis>
SEM micrographs of
<taxonomicName class="Bacillariophyceae" family="Diadesmidaceae" genus="Luticola" kingdom="Plantae" lsidName="Luticola inserata" order="Naviculales" pageId="0" pageNumber="115" phylum="Bacillariophyta" rank="species" species="inserata">
<emphasis italics="true" pageId="0" pageNumber="115">Luticola inserata</emphasis>
<taxonomicName id="10990EF2C198C469FDE9613817596CCE" class="Bacillariophyceae" family="Diadesmidaceae" genus="Luticola" kingdom="Plantae" lsidName="Luticola inserata" order="Naviculales" pageId="0" pageNumber="115" phylum="Bacillariophyta" rank="species" species="inserata">
<emphasis id="F0DF86CCBC8F689ADF4F86387AEB21F6" italics="true" pageId="0" pageNumber="115">Luticola inserata</emphasis>
</taxonomicName>
(Hustedt) D.G.Mann. External view of valve (
<emphasis bold="true" pageId="0" pageNumber="115">A, B</emphasis>
<emphasis id="1E975D11A99A265B764574B1C44B3F8B" bold="true" pageId="0" pageNumber="115">A, B</emphasis>
). Detailed external views showing proximal raphe endings with irregular shallow grooves
<emphasis bold="true" pageId="0" pageNumber="115">C</emphasis>
<emphasis id="87F8B9342CE3033757C9BCF077AAFD74" bold="true" pageId="0" pageNumber="115">C</emphasis>
distal raphe ending, areolae
<emphasis bold="true" pageId="0" pageNumber="115">D</emphasis>
<emphasis id="3F4362E806BDB5C07EA2764692F4F5CD" bold="true" pageId="0" pageNumber="115">D</emphasis>
stigma opening
<emphasis bold="true" pageId="0" pageNumber="115">E</emphasis>
<emphasis id="E0AB450C4F3D55C20EE8A01B6A7114B6" bold="true" pageId="0" pageNumber="115">E</emphasis>
internal view of valve
<emphasis bold="true" pageId="0" pageNumber="115">F</emphasis>
<emphasis id="352E7C980A27210DFDD79F3CEC5551DE" bold="true" pageId="0" pageNumber="115">F</emphasis>
detailed internal view showing proximal raphe endings and stigma opening
<emphasis bold="true" pageId="0" pageNumber="115">G</emphasis>
<emphasis id="3AC812C3ACFACCAF53D2DCBFFD6D77FD" bold="true" pageId="0" pageNumber="115">G</emphasis>
and distal raphe ending and hymenate structure
<emphasis bold="true" pageId="0" pageNumber="115">H</emphasis>
<emphasis id="D0403080B43AEF50A8F3DC508B2E676C" bold="true" pageId="0" pageNumber="115">H</emphasis>
open copulae
<emphasis bold="true" pageId="0" pageNumber="115">I</emphasis>
<emphasis id="090B82A8352E3C9566DB5CC28378E079" bold="true" pageId="0" pageNumber="115">I</emphasis>
. Scale bars: 10
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="E108AFBBCC5A1975396CA520D4A74609" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="115">A, F, I</emphasis>
<emphasis id="7C0E5772F1C39CB14C86BB3CBEE32935" bold="true" pageId="0" pageNumber="115">A, F, I</emphasis>
), 5
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="060650BCE976C5C1A5C98ACAE470BC1C" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="115">B-D, G</emphasis>
<emphasis id="B643B70AD454A695AB66DE670E68D1D2" bold="true" pageId="0" pageNumber="115">B-D, G</emphasis>
), 3
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="4AB583446A5DC64281BF79F201E18DE2" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="115">E</emphasis>
<emphasis id="58775A67E16AA66BD710646109AC71CB" bold="true" pageId="0" pageNumber="115">E</emphasis>
), 4
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="945275DC0FDD585954CF7F0B620124CC" originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="115">H</emphasis>
<emphasis id="4DABECB79DCA25334B25AB67EF5B776E" bold="true" pageId="0" pageNumber="115">H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="distribution">
<paragraph pageId="0" pageNumber="115">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="115">
<subSubSection id="49C3A4AF705ADBD2CC2D99FCD46C35F6" pageId="0" pageNumber="115" type="distribution">
<paragraph id="F5E474FE9F69D9C91D589282E5400A8E" pageId="0" pageNumber="115">Distribution.</paragraph>
<paragraph id="17551EBE800606A56BBB7C932A0F9261" pageId="0" pageNumber="115">
This species was observed only in sample SZCZ26472 from Western South China Sea,
<normalizedToken originalValue="Quáng">Quang</normalizedToken>
<normalizedToken originalValue="Yên">Yen</normalizedToken>
<normalizedToken id="CFE97C348DB39BB79F6A2DE367D10DF8" originalValue="Quáng">Quang</normalizedToken>
<normalizedToken id="35F8934D0109CB411125E755EF51E815" originalValue="Yên">Yen</normalizedToken>
, in Halong region of NE Vietnam, shell scrape from oysters in offshore aquaculture area.
</paragraph>
</subSubSection>

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<mods:title>Three new species of Fancy Case caterpillars from threatened forests of Hawaii (Lepidoptera, Cosmopterigidae, Hyposmocoma)</mods:title>
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<treatment ID-GBIF-Taxon="152033292" LSID="urn:lsid:plazi:treatment:2DA78A31658CD905A9223157CD984E78" httpUri="http://treatment.plazi.org/id/2DA78A31658CD905A9223157CD984E78" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
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<paragraph pageId="4" pageNumber="5">
<taxonomicName LSID="http://species-id.net/wiki/Hyposmocoma_inversella" authority="Walsingham, 1907" authorityName="Walsingham" authorityYear="1907" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyposmocoma inversella" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="inversella">
<pageBreakToken pageId="4" pageNumber="5" start="start">Hyposmocoma</pageBreakToken>
<treatment id="2DA78A31658CD905A9223157CD984E78" ID-GBIF-Taxon="152033292" LSID="urn:lsid:plazi:treatment:2DA78A31658CD905A9223157CD984E78" httpUri="http://treatment.plazi.org/id/2DA78A31658CD905A9223157CD984E78" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" scope_family="Cosmopterigidae" scope_order="Lepidoptera">
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<paragraph id="9A70962AE648A640D3A06AC1747D04DC" pageId="4" pageNumber="5">
<taxonomicName id="37AD5A1BDA858751EB15C979DF62095C" ID-CoL="943NY" LSID="http://species-id.net/wiki/Hyposmocoma_inversella" authority="Walsingham, 1907" authorityName="Walsingham" authorityYear="1907" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyposmocoma inversella" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="inversella">
<pageBreakToken id="7F40932B0F884DCF49050B7DEB42CAD2" pageId="4" pageNumber="5" start="start">Hyposmocoma</pageBreakToken>
inversella Walsingham, 1907
</taxonomicName>
Figs 217
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="diagnosis">
<paragraph pageId="4" pageNumber="5">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="5">
<subSubSection id="9A0F29D557BB5AEA11722285009E82CA" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="EA4959BCBF32EAC3014FD9C353F1DADE" pageId="4" pageNumber="5">Diagnosis.</paragraph>
<paragraph id="31761A6645E7AB20DCD22ACB18D6E74A" pageId="4" pageNumber="5">
This species has a large, orange,
<normalizedToken originalValue="“V”">&quot;V&quot;</normalizedToken>
<normalizedToken id="28322B4B84E0D5DF8877C30562F7ED36" originalValue="“V”">&quot;V&quot;</normalizedToken>
-shaped marking on the forewing found in no other described species of
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<taxonomicName id="F0579724C3AEB7539953B427E2F69C16" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyposmocoma" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Hyposmocoma</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="re-description">
<paragraph pageId="4" pageNumber="5">Re-description.</paragraph>
<paragraph lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="16DA7F9349ED291F6C80322D646FE5B5" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="re-description">
<paragraph id="3F12F50052DAE865CF37E8C9C89511C4" pageId="4" pageNumber="5">Re-description.</paragraph>
<paragraph id="D923D3DB7CEB042B1BC857DF6D581793" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Male. (n = 1; Fig 2). Forewing length 4.9 mm.Head with copper-colored scales arranged radially from compound eye. Haustellum with light brown
<pageBreakToken pageId="5" pageNumber="6" start="start">scales</pageBreakToken>
<pageBreakToken id="6D268642478091EE8CB5073A10685D87" pageId="5" pageNumber="6" start="start">scales</pageBreakToken>
. Maxillary palpus reduced. Labial palpus curved with copper colored scales, scale color similar on all surfaces of palpus. Antenna brown with a mosaic of copper scales. Thorax mostly copper; dark brown scales present along anterior margin. Foreleg coxa with brown and copper scales; femur, tibia, and tarsomeres mostly dark brown with light brown ring at distal margin of femur, tibia, and tarsomeres I-V. Midleg as foreleg, but spurs covered in a mixture of dark and light-brown scales. Hindleg as midleg, but longer with long hairs on dorsal margin of tibia. Forewing brown, two transverse orange fascia form a
<normalizedToken originalValue="“V”">&quot;V&quot;</normalizedToken>
<normalizedToken id="8B81DA97381AA7DEE718D551EFF87671" originalValue="“V”">&quot;V&quot;</normalizedToken>
. Proximal orange band widens along anal margin, distal band uniform in width. Fringe orange and brown, longer scales tending to be brown, shorter ones orange. Hindwing brown with brown fringe. Abdomen dorsally dark brown; ventrally copper, with tuft of long dark brown scales covering lateral surface of genitalia.
</paragraph>
<paragraph pageId="5" pageNumber="6">Male genitalia. Right brachium of uncus sickle shaped, slender, heavily sclerotized, widening at 1/2 length, slightly twisted to left. Left brachium small, not sclerotized. Tegumen wide and sclerotized. Four long thin setae along dorsal margin of left valva, three shorter setae at same location of right valva. Left valva slightly wider than right. Dense row of fine, hair-like setae disposed along inner ventral margin of valva. Phallus stout, blunt tipped, heavily sclerotized, open ventrally, and bent ventrad at about 1/2 of length; vesica without spines or cornuti. Anellus with two symmetrical rounded lobes with short setae.</paragraph>
<paragraph pageId="5" pageNumber="6">Female. (n = 1). Externally as male, forewing length 4.7 mm.</paragraph>
<paragraph pageId="5" pageNumber="6">Female genitalia. (Fig 17). Papillae anales short and setose. Anterior and posterior apophyses slender, long, anterior apophyses slightly longer than posterior apophyses. Ostium bursae small, heavily sclerotized, externally protruding, roughly triangular, not snail-shaped. Corpus bursae oval and elongate, with very light scobination; signum absent. Ductus bursae long and narrow, slightly twisted, about equal in length of corpus bursae. Apical margin of tergum VIII with median emargination.</paragraph>
<paragraph pageId="5" pageNumber="6">
<paragraph id="57FCCCA085B85053A0DC771BC507D84C" pageId="5" pageNumber="6">Male genitalia. Right brachium of uncus sickle shaped, slender, heavily sclerotized, widening at 1/2 length, slightly twisted to left. Left brachium small, not sclerotized. Tegumen wide and sclerotized. Four long thin setae along dorsal margin of left valva, three shorter setae at same location of right valva. Left valva slightly wider than right. Dense row of fine, hair-like setae disposed along inner ventral margin of valva. Phallus stout, blunt tipped, heavily sclerotized, open ventrally, and bent ventrad at about 1/2 of length; vesica without spines or cornuti. Anellus with two symmetrical rounded lobes with short setae.</paragraph>
<paragraph id="A818295C7C868A202A2082504792A0E3" pageId="5" pageNumber="6">Female. (n = 1). Externally as male, forewing length 4.7 mm.</paragraph>
<paragraph id="FF3CDCA2E0EFF6908B7DBEE583BF5E7A" pageId="5" pageNumber="6">Female genitalia. (Fig 17). Papillae anales short and setose. Anterior and posterior apophyses slender, long, anterior apophyses slightly longer than posterior apophyses. Ostium bursae small, heavily sclerotized, externally protruding, roughly triangular, not snail-shaped. Corpus bursae oval and elongate, with very light scobination; signum absent. Ductus bursae long and narrow, slightly twisted, about equal in length of corpus bursae. Apical margin of tergum VIII with median emargination.</paragraph>
<paragraph id="9E2B9D3B2B7F93FEEB92AA66628A2120" pageId="5" pageNumber="6">
Larval case. (n = 2).Dark brown, smooth, 7.8-8.5 mm in length and 1.8-2 mm wide. The larval case is very similar to that of
<taxonomicName class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyposmocoma auropurpurea" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="auropurpurea">Hyposmocoma auropurpurea</taxonomicName>
<taxonomicName id="FF507B7AF5F8B9DA60A5AEBA34DC8A48" class="Insecta" family="Cosmopterigidae" genus="Hyposmocoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyposmocoma auropurpurea" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="auropurpurea">Hyposmocoma auropurpurea</taxonomicName>
. Though both species are found in the Waianae Mountains, adult morphology and DNA sequence confirms these are distinct species.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="material examined">
<paragraph pageId="5" pageNumber="6">Material examined.</paragraph>
<paragraph pageId="5" pageNumber="6">
<subSubSection id="A5529B41582DDEFD5B104C8E4FE36023" pageId="5" pageNumber="6" type="material examined">
<paragraph id="02A8B2137915D145D331F853B01DE8CB" pageId="5" pageNumber="6">Material examined.</paragraph>
<paragraph id="9F14763FC9125B96FFBA76D3B81AAA47" pageId="5" pageNumber="6">
(1♂, 2♀). 1♂: HI: Oahu, Waianae Range, Palikea Trail,
<normalizedToken originalValue="“purse”">&quot;purse&quot;</normalizedToken>
<normalizedToken id="A94FAE5F857F4704599670D6C93A4ED6" originalValue="“purse”">&quot;purse&quot;</normalizedToken>
case, 15-I-2009, emergence. 16-III-2009, #DR09A3I, coll. P. Schmitz, D. Rubinoff, W. Haines, J. Eiben. Specimen in perfect condition. Male genitalia slide #AK100. 1♀: same data as male. Female genitalia slide #AK101. 1♀: HI: Oahu, Pahole NAR. Northern Waianae Mts. Elev. 500 m, on Pipturus albidus (
<taxonomicName genus="Urticaceae" lsidName="Urticaceae" pageId="5" pageNumber="6" rank="genus">Urticaceae</taxonomicName>
<taxonomicName id="E01C6C219A69971A998CC6595FED6C5C" genus="Urticaceae" lsidName="Urticaceae" pageId="5" pageNumber="6" rank="genus">Urticaceae</taxonomicName>
). 14-XII-2010, #DR10L1C, em. 24-III-2011, coll. P. Krushelnycky. Additional voucher collection #AR0803, spec/lot #PKSP11751. Molecular ID: AK-259-11. All specimens from the UHIM.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="specimens sequenced">
<paragraph pageId="5" pageNumber="6">Specimens sequenced.</paragraph>
<paragraph pageId="5" pageNumber="6">Two specimens from Oahu, Hawaii, USA (UH log number DR09A3I), extraction codes DN1314A, DN1314B.</paragraph>
<subSubSection id="1975BE6F0CF564E7F1F1E16D6A5A9938" pageId="5" pageNumber="6" type="specimens sequenced">
<paragraph id="CA5B0EA18203CCF06058E82374AD59EF" pageId="5" pageNumber="6">Specimens sequenced.</paragraph>
<paragraph id="2CE76A6C0D6E3F4ACC817CFFF64D0BCB" pageId="5" pageNumber="6">Two specimens from Oahu, Hawaii, USA (UH log number DR09A3I), extraction codes DN1314A, DN1314B.</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="biology">
<paragraph pageId="5" pageNumber="6">Biology.</paragraph>
<paragraph lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<subSubSection id="7C52D34E98E6F4C85A18626D84FFEB71" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="biology">
<paragraph id="972E0531AE71EBEC7ACDA5CEF2E012DB" pageId="5" pageNumber="6">Biology.</paragraph>
<paragraph id="E296691F1F193F27EBDAE5151B102723" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Case-making larvae were collected in the Waianae Mountains of Oahu on the Palikea Trail during December and January. Larvae were found in leaf litter and on rotting logs. Adult emergence occurred between March and April. Because we have
<pageBreakToken pageId="6" pageNumber="7" start="start">collected</pageBreakToken>
<pageBreakToken id="934B610DFAD7D16E63FB7BFD72115323" pageId="6" pageNumber="7" start="start">collected</pageBreakToken>
fairly extensively in the Waianae Mountains, we predict that this species univoltine, with larvae active during the winter months and adults emerging in the spring.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="distribution">
<paragraph pageId="6" pageNumber="7">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="7">Known only from the Waianae Mountain Range, Oahu.</paragraph>
<subSubSection id="C1CF59BB8EAA8E2CF6E1A5D1AEDF6BED" pageId="6" pageNumber="7" type="distribution">
<paragraph id="EE684FA15DE7B97A7316E0C721B65C13" pageId="6" pageNumber="7">Distribution.</paragraph>
<paragraph id="DA22B3FA4BBB0DB23F511FEE8D8D32C1" pageId="6" pageNumber="7">Known only from the Waianae Mountain Range, Oahu.</paragraph>
</subSubSection>
</treatment>
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<mods:title id="8508CCF5A174301E4962BE35F04A9BA9">A new, widespread genus of Baetidae from South Asia (Insecta, Ephemeroptera)</mods:title>
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<mods:title id="C70709C0B247429C495913EEF69B5EFD">A new, widespread genus of Baetidae from South Asia (Insecta, Ephemeroptera)</mods:title>
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<mods:namePart id="87D6983D45A4E915022AB2FE3251C203">Kaltenbach, Thomas</mods:namePart>
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<mods:affiliation id="AF4D6F3C681B25FD77B3CBF0D675CF90">Department of Entomology, Biological Faculty, Saint-Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia</mods:affiliation>
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<mods:affiliation id="F3A25C0CED6A63706E70AA1DA0ED332C">Department of Entomology, Biological Faculty, Saint-Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia</mods:affiliation>
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<taxonomicNameLabel id="F85B2E6ABB6A1A4360E5A1E959BDECF0" pageId="0" pageNumber="231">sp. nov.</taxonomicNameLabel>
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@ -108,7 +108,7 @@ cottage, UN11;
<emphasis id="1AE757EF04A4F6FBD7084E2284FF6BCD" italics="true" pageId="0" pageNumber="231">A. sumatrensis</emphasis>
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sp. nov. from other species of
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<taxonomicName id="CC626F23680D233BB480A734A5EE76D8" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Arcobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arcobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="1B64BC2F28EAB42C0342E687AF85089C" italics="true" pageId="0" pageNumber="231">Arcobaetis</emphasis>
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gen. nov.: A) distal margins of segments in middle part of flagellum without enlarged spines; B) labial palp segment III sub-rectangular, at base smaller than distal margin of segment II (Fig.
@ -124,7 +124,7 @@ gen. nov.: A) distal margins of segments in middle part of flagellum without enl
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<paragraph id="03C4048FE3587C3C7E4DD98E65D7B22C" pageId="0" pageNumber="231">
<emphasis id="7EC4815F9A2610C19C94E3536B2FA2C6" bold="true" pageId="0" pageNumber="231">Figure 6.</emphasis>
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<emphasis id="765AFF6BC9ED3F91805EF671FC85F7E2" italics="true" pageId="0" pageNumber="231">Arcobaetis sumatrensis</emphasis>
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sp. nov., larva, habitus (holotype)
@ -139,7 +139,7 @@ ventral view. Scale bar: 1 mm
<caption id="D0716DFE41A6D371288C5F1337B1E1DD" doi="10.3897/zookeys.1168.104844.figure7" httpUri="https://binary.pensoft.net/fig/872209" pageId="0" pageNumber="231" start="Figure 7" startId="F7">
<paragraph id="98707F250CA5E86C604041DC66937EC7" pageId="0" pageNumber="231">
<emphasis id="112243FF9736B704DB9A2782A7A52634" bold="true" pageId="0" pageNumber="231">Figure 7.</emphasis>
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<emphasis id="A9B8C4642705F73FE7C10042855E078D" italics="true" pageId="0" pageNumber="231">Arcobaetis sumatrensis</emphasis>
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sp. nov., larva
@ -168,7 +168,7 @@ apex of paraglossa (dorsal view).
<caption id="B8F206684044AA33C34DDC6E311DB653" doi="10.3897/zookeys.1168.104844.figure8" httpUri="https://binary.pensoft.net/fig/872210" pageId="0" pageNumber="231" start="Figure 8" startId="F8">
<paragraph id="F47DBB788ECA984A941E4C6295250895" pageId="0" pageNumber="231">
<emphasis id="D9A89A63D4A65654976475B7588C3323" bold="true" pageId="0" pageNumber="231">Figure 8.</emphasis>
<taxonomicName id="BFFA73ECC910117A2A35CC3335994FAD" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis sumatrensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="sumatrensis">
<taxonomicName id="BFFA73ECC910117A2A35CC3335994FAD" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Arcobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arcobaetis sumatrensis" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="CD0B914EFE30A4EAB80A3BE07205E8AA" italics="true" pageId="0" pageNumber="231">Arcobaetis sumatrensis</emphasis>
</taxonomicName>
sp. nov., larva
@ -210,7 +210,7 @@ paraproct.
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<emphasis id="FF60805EDF0CDF2932463AABED34BCE0" bold="true" pageId="0" pageNumber="231">Figure 9.</emphasis>
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<taxonomicName id="B90F8A1D96CAF46C6AB21284FE583CA9" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" class="Insecta" family="Baetidae" genus="Arcobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Arcobaetis sumatrensis" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="584B6A5D6FF4D0652A7CA8446BDA5ADC" italics="true" pageId="0" pageNumber="231">Arcobaetis sumatrensis</emphasis>
</taxonomicName>
sp. nov., larva

View file

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<mods:title>Generic placement of the Neotropical species of &quot; Phragmatobia &quot; (Erebidae, Arctiinae), with a remarkable matrivorous species from the Peruvian Andes</mods:title>
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<mods:title id="D33890366BEE45A1EF8E9C86E4C3A80B">Generic placement of the Neotropical species of &quot; Phragmatobia &quot; (Erebidae, Arctiinae), with a remarkable matrivorous species from the Peruvian Andes</mods:title>
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<mods:namePart>Freina, Josef J. De</mods:namePart>
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<taxonomicNameLabel pageId="7" pageNumber="76">gen. n.</taxonomicNameLabel>
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<taxonomicNameLabel id="0904B83DD8E8C2CB6BAB7D31D992E4F4" pageId="7" pageNumber="76">gen. n.</taxonomicNameLabel>
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<paragraph pageId="7" pageNumber="76">Type species.</paragraph>
<paragraph pageId="7" pageNumber="76">
<taxonomicName class="Insecta" family="Arctiidae" genus="Turuptiana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turuptiana thursbyi" order="Lepidoptera" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="thursbyi">Turuptiana thursbyi</taxonomicName>
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<paragraph id="4ED9D603940ADFBC6CAA69EB59F2CE63" pageId="7" pageNumber="76">Type species.</paragraph>
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<taxonomicName id="0D9C096E2A11D51A7B369BEE918A7656" class="Insecta" family="Erebidae" genus="Turuptiana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Turuptiana thursbyi" order="Lepidoptera" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="thursbyi">Turuptiana thursbyi</taxonomicName>
Rothschild, 1910.
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<paragraph pageId="7" pageNumber="76">Etymology.</paragraph>
<paragraph pageId="7" pageNumber="76">
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<paragraph id="E587D68171C3E83D97DA372604851AE8" pageId="7" pageNumber="76">Etymology.</paragraph>
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The name is derived from a combination of the words Patagonia and
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.
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<paragraph pageId="7" pageNumber="76">Diagnosis.</paragraph>
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<paragraph id="BDFF52E2AEC091F884712ECEE1AAF13F" pageId="7" pageNumber="76">Diagnosis.</paragraph>
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Although
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shows similarities to the Holarctic
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in some external aspects, it differs in having longer, symmetrical rami of the male antenna, ochre thoracic collar, lack of a male clasper, pale tan forewing pattern (usually), and a restricted distribution to the Chilean Andes of South America. The wing colour and pattern is also similar to
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, but structurally
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has a more robust build with denser thoracic vestiture, equally long posterior and anterior male antennal rami (anterior rami shorter than posterior in
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), male antennal rami up to 3
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longer than antennal segment length (up to 2
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in
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); 2nd labial segment elongate, 1.8
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as long as wide, 1.5
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longer than apical segment; thoracic collar contrastingly paler ochre (conconcolourous with dorsal thoracic vestiture in
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); thoracic vestiture dense and pilose (sparse and shaggy in
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);femur and tibia elongate, 4.5-5.6
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longer than wide (very stout, 3.0-3.5
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longer than wide in
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); metatibia with two pairs of spurs (one pair in
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); forewing medial line present (absent in
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); postmedial line usually double (absent in
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); hindwing discal spot diffuse and elongate (sharp or absent in
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). The male coremata betwen the 7th and 8th sternite are moderately developed in
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, very reduced in
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.
</paragraph>
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<paragraph pageId="7" pageNumber="76">Description.</paragraph>
<paragraph lastPageId="8" lastPageNumber="77" pageId="7" pageNumber="76">
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<paragraph id="4C549F89C581D8A2B1FBDC241C2C854F" pageId="7" pageNumber="76">Description.</paragraph>
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Male (female unknown). Head - vestiture dark brown to black, setae long; antenna bipectinate, ciliate ventrally; longest posterior rami 1.5-3.0
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segment length, longest anterior rami 1.1-3.0
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segment length; rami longest over middle third of antenna, decreasing in length toward base and apex; eye elliptical, 1.2-1.5
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as high as wide; labial palp short, not extending beyond vestiture of frons; haustellum reduced and poorly sclerotized, presumably nonfunctional. Thorax - vestiture of vertex and ventrum of thorax black brown; tegulae entirely black brown or black brown edged with yellowish brown; patagia yellowish brown or rarely black brown; leg vestiture brownish ochre, dorsum of femur yellow or red, co-varying with hindwing and abdomen ground colour; apex of prothoracic tibia with two subequal, blunt, triangular projections; mesotibia with two apical and two subapical spurs, length of apical spurs 1.5
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and supapical spurs 0.6
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tibial width at apex; two metatibial spurs, posterior spur slightly longer than anterior; metepisternum lacking microtymbals. Forewings - forewing length 12.9-13.2 mm (mean 13.1 mm; n = 4), length:width ratio averaging 2.1; ground colour pale ochre yellow but with broad, sometimes entirely confluent dark-brown transverse bands (Fig. 4c); pattern elements consisting of dark basal area and sinuous, diffuse dark-brown transverse lines (Fig. 4a) discal spot indistinct dorsally, but well defined ventrally; ventrally with bands obsolete except for marginal band. Hindwing - ground colour pinkish red or rarely yellow (Fig. 4d)
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with broad dark-brown marginal and costal band; well-defined brown, crescentic discal spot; similar ventrally but with discal spot better defined. Abdomen - Vestiture brownish black and pinkish red or yellow subdorsdally, ventrally with segmental margins yellowish ochre; abdomen entirely dark brown in melanic specimens (Fig. 4c). Coremata between sternites 7-8 in shallow pockets, scent scales approximately 0.5
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as long as sternite length. Genitalia - highly simplified, with large, triangular, dorsoventrally flattened uncus characteristic of subtribe; uncus 1.5
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longer than width of base, broadly joined to wide, band-like tegumen; dorsal margin of tegumen caudally recurved; valve simple and digitate, lacking processes or claspers, 1.5
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as long as uncus-tegumen complex; vinculum semicircular, saccus v-shaped, similar in length to uncus; aedeagus large, 5-6
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longer than wide, 2
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as long as width of genital capsule, curving dorsad slightly; coecum 1/10 length of aedeagus, directed slightly ventrad; vesica extremely large, diameter when inflated 2
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that of genital capsule; vesica directed right-laterad, globose, finely spiculate, with poorly differentiated basal chamber and large apical chamber (Fig. 11b).
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View file

@ -1,60 +1,60 @@
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<paragraph pageId="0" pageNumber="207">Untersuchte Exemplare.</paragraph>
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3 Ex., BE, Rubigen, Hechteloch, Schwarzbach, 608245N/193083E [
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,
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], 520 m, 10.6.2019, leg. C. Germann (cCG).
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<paragraph pageId="0" pageNumber="207">Bemerkung.</paragraph>
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Die letzten Funde lagen aus den 70 er Jahren vor (Aarwangen BE, leg. &amp; coll. A. Linder ETHZ), hier erstmals aktuelle Funde der auf Feuchtgebiete
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Art.
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View file

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<taxonomicName LSID="2DD185E4-FB69-55F2-8CFE-48A8549B49B0" authority="Papp, 1991" authorityName="Papp" authorityYear="1991" class="Insecta" family="Braconidae" genus="Paroplitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroplitis rugosus" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="rugosus">Paroplitis rugosus Papp, 1991</taxonomicName>
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<paragraph pageId="0" pageNumber="33">
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Papp, 1991: 165. Original description.
</paragraph>
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<subSubSection pageId="0" pageNumber="33" type="holotype">
<paragraph pageId="0" pageNumber="33">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="33">Female, HNHM (not examined). Holotype labels: Austria, Tirol, Obergurgl, Belstein, 2400 m alt., 7 September 1970, leg. S. Mahunka.</paragraph>
<subSubSection id="80F53D5E58694DF116DAD482D9AB4653" pageId="0" pageNumber="33" type="holotype">
<paragraph id="355FED03FFD1D7E2EC012C1E24D55BF4" pageId="0" pageNumber="33">Holotype.</paragraph>
<paragraph id="ECAB64E97963062C9138557A64AA0140" pageId="0" pageNumber="33">Female, HNHM (not examined). Holotype labels: Austria, Tirol, Obergurgl, Belstein, 2400 m alt., 7 September 1970, leg. S. Mahunka.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="description">
<paragraph pageId="0" pageNumber="33">Description.</paragraph>
<paragraph pageId="0" pageNumber="33">
<subSubSection id="8EF60CFEA39846870098FE7FEA99334A" pageId="0" pageNumber="33" type="description">
<paragraph id="C73AC2C5305028D9CC2C356361AA6857" pageId="0" pageNumber="33">Description.</paragraph>
<paragraph id="C6FDFC3FA0FE3126E8A55771C01C8CEB" pageId="0" pageNumber="33">
A detailed description and images of the species in
<bibRefCitation author="Papp, J" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="33" pagination="145 - 167" refId="B8" refString="Papp, J, 1991. New Braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 2. Annales Historico-Naturales Musei Nationalis Hungarici 83: 145 - 167" title="New Braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 2." volume="83" year="1991">Papp (1991)</bibRefCitation>
<bibRefCitation id="34F8E83FD98246E4A220AA8A949231C7" author="Papp, J" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="33" pagination="145 - 167" refId="B8" refString="Papp, J, 1991. New Braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 2. Annales Historico-Naturales Musei Nationalis Hungarici 83: 145 - 167" title="New Braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 2." volume="83" year="1991">Papp (1991)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="host">
<paragraph pageId="0" pageNumber="33">Hosts.</paragraph>
<paragraph pageId="0" pageNumber="33">Unknown.</paragraph>
<subSubSection id="F9D4C3531CB5FFC354BECAC15985B800" pageId="0" pageNumber="33" type="host">
<paragraph id="352B7ED3CF15F44F4DE01FAF56E8FC9F" pageId="0" pageNumber="33">Hosts.</paragraph>
<paragraph id="E47245DACD2004E92E469F69DF4F018D" pageId="0" pageNumber="33">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="distribution">
<paragraph pageId="0" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="33">
<subSubSection id="AC85621C7C37F5EE406A3F2E62FB55A1" pageId="0" pageNumber="33" type="distribution">
<paragraph id="C5DF80EDFE2B122C79D29442E8998A24" pageId="0" pageNumber="33">Distribution.</paragraph>
<paragraph id="42D65EEE18C692E149305182F89FD81A" pageId="0" pageNumber="33">
Western Palaearctic Region: Austria (
<bibRefCitation author="Papp, J" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="33" pagination="145 - 167" refId="B8" refString="Papp, J, 1991. New Braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 2. Annales Historico-Naturales Musei Nationalis Hungarici 83: 145 - 167" title="New Braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 2." volume="83" year="1991">Papp 1991</bibRefCitation>
<bibRefCitation id="C6556F2873F264E9E68B80A4ED8B3421" author="Papp, J" journalOrPublisher="Annales Historico-Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="33" pagination="145 - 167" refId="B8" refString="Papp, J, 1991. New Braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 2. Annales Historico-Naturales Musei Nationalis Hungarici 83: 145 - 167" title="New Braconid wasps (Hymenoptera, Braconidae) in the Hungarian Natural History Museum, 2." volume="83" year="1991">Papp 1991</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="33" type="comments">
<paragraph pageId="0" pageNumber="33">Comments.</paragraph>
<paragraph pageId="0" pageNumber="33">
<subSubSection id="48E92894CA4982DD95D19039E87B73BF" pageId="0" pageNumber="33" type="comments">
<paragraph id="B9234357894084462F7D91E25ED3D364" pageId="0" pageNumber="33">Comments.</paragraph>
<paragraph id="1B9E552B272E322A4E424E4152F12089" pageId="0" pageNumber="33">
Only known from the female holotype. Its distribution seems to overlap with that of
<taxonomicName genus="P." lsidName="P. wesmaeli" pageId="0" pageNumber="33" rank="species" species="wesmaeli">
<emphasis italics="true" pageId="0" pageNumber="33">P. wesmaeli</emphasis>
<taxonomicName id="B932722D596B5477D270169D2791899B" lsidName="P. wesmaeli" pageId="0" pageNumber="33" rank="species" species="wesmaeli">
<emphasis id="4E37EAC726DCE981B3DB26ADCF2D0796" italics="true" pageId="0" pageNumber="33">P. wesmaeli</emphasis>
</taxonomicName>
, although
<taxonomicName genus="P." lsidName="P. rugosus" pageId="0" pageNumber="33" rank="species" species="rugosus">
<emphasis italics="true" pageId="0" pageNumber="33">P. rugosus</emphasis>
<taxonomicName id="3BEE741E1A52C5CB196FC7C7C259076C" lsidName="P. rugosus" pageId="0" pageNumber="33" rank="species" species="rugosus">
<emphasis id="8D6FD89D45CAA6769AC30A1AC24CF771" italics="true" pageId="0" pageNumber="33">P. rugosus</emphasis>
</taxonomicName>
was collected in the Alps at a higher altitude (2400 m) than European specimens of
<taxonomicName baseAuthorityName="Ruthe" baseAuthorityYear="1860" class="Insecta" family="Braconidae" genus="Paroplitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroplitis wesmaeli" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wesmaeli">
<emphasis italics="true" pageId="0" pageNumber="33">Paroplitis wesmaeli</emphasis>
<taxonomicName id="4235C9969BD014410AAB6F02154D4D57" baseAuthorityName="Ruthe" baseAuthorityYear="1860" class="Insecta" family="Braconidae" genus="Paroplitis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paroplitis wesmaeli" order="Hymenoptera" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="wesmaeli">
<emphasis id="78E20EE0A106F0CBE86547B1043128BF" italics="true" pageId="0" pageNumber="33">Paroplitis wesmaeli</emphasis>
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.
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@ -1,244 +1,244 @@
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<taxonomicName LSID="http://species-id.net/wiki/Neophyllognathopus_bassoti" authority="Rouch, 1972" authorityName="Rouch" authorityYear="1972" class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti (Rouch, 1972)</taxonomicName>
<taxonomicNameLabel pageId="15" pageNumber="16">comb. n.</taxonomicNameLabel>
<treatment id="2DDFE2EE7722D9A2EA94FC14B77A4DBC" ID-GBIF-Taxon="152029620" LSID="urn:lsid:plazi:treatment:2DDFE2EE7722D9A2EA94FC14B77A4DBC" httpUri="http://treatment.plazi.org/id/2DDFE2EE7722D9A2EA94FC14B77A4DBC" lastPageId="21" lastPageNumber="22" pageId="15" pageNumber="16" scope_class="Copepoda" scope_family="Phyllognathopodidae" scope_order="Harpacticoida">
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<taxonomicNameLabel id="8D529991364F4579F776B5612D428ECD" pageId="15" pageNumber="16">comb. n.</taxonomicNameLabel>
Figs 1725
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<paragraph pageId="15" pageNumber="16">Neotype designation.</paragraph>
<paragraph pageId="15" pageNumber="16">
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<paragraph id="215CE1BE3A49FE5EE618DDEA50467FDE" pageId="15" pageNumber="16">Neotype designation.</paragraph>
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Female neotype completely dissected and mounted in polyvinyl lactophenol, deposited at the Natural History Museum, London (reg. No. NHM.2008. neotype). Other material: 5 ♀♀ and 3 ♂♂ mounted on slides, 5 ♀♀ and 5 ♂♂ processed for SEM; India, 7 January 1999, Y. Ranga Reddy coll.; 1 ♀, slide code 66/49, 1 ♀, slide code 66/53, 1 ♂, slide code 66/55, Santa Fe, Bantayan island, Pooc, Philippines, V. Cottarelli coll. (see
<bibRefCitation author="Bruno, MC" journalOrPublisher="Journal of Crustacean Biology" pageId="38" pageNumber="39" pagination="510 - 529" title="Harpacticoids from groundwaters in the Philippines: Parastenocaris mangyans, new species, Epactophanes philippinus, new species, and redescription of Phyllognathopus bassoti (Copepoda)." url="doi: 10.2307/1549260" volume="19" year="1999">Bruno and Cottarelli 1999</bibRefCitation>
<bibRefCitation id="6C36E36D1771A3EFA1A35B1E89C4661B" author="Bruno, MC" journalOrPublisher="Journal of Crustacean Biology" pageId="38" pageNumber="39" pagination="510 - 529" title="Harpacticoids from groundwaters in the Philippines: Parastenocaris mangyans, new species, Epactophanes philippinus, new species, and redescription of Phyllognathopus bassoti (Copepoda)." url="doi: 10.2307/1549260" volume="19" year="1999">Bruno and Cottarelli 1999</bibRefCitation>
, for locality details).
</paragraph>
<paragraph pageId="15" pageNumber="16">
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<paragraph id="B1F42F9691803439BEA857C1A069AAC1" pageId="15" pageNumber="16">
<taxonomicName id="47BF76F1A5F77FF58DD96DED97526F0A" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
is proposed herein as new combination for
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus bassoti" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bassoti">Phyllognathopus bassoti</taxonomicName>
<taxonomicName id="3B923B2D6A24D161B8FBE04EE9A3C15A" class="Copepoda" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus bassoti" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bassoti">Phyllognathopus bassoti</taxonomicName>
assigned by
<bibRefCitation author="Rouch, R" journalOrPublisher="Archives Zoologie Experimentale &amp; Generale" pageId="38" pageNumber="39" pagination="147 - 164" title="Deux harpacticidae nouveaux de l'ile de Long Island (Territoire de Papouasie et de Nouvelle-Guinee)." url="doi: 10.1023/A:1013148808110" volume="113" year="1972">Rouch (1972)</bibRefCitation>
<bibRefCitation id="3F5C07E674C7BBA89EA89433038CB533" author="Rouch, R" journalOrPublisher="Archives Zoologie Experimentale &amp; Generale" pageId="38" pageNumber="39" pagination="147 - 164" title="Deux harpacticidae nouveaux de l'ile de Long Island (Territoire de Papouasie et de Nouvelle-Guinee)." url="doi: 10.1023/A:1013148808110" volume="113" year="1972">Rouch (1972)</bibRefCitation>
in the original description to the genus
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Phyllognathopus</taxonomicName>
<taxonomicName id="A958146FC8B284F366805BB46460524B" class="Copepoda" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Phyllognathopus</taxonomicName>
. According to
<bibRefCitation author="ICZN," journalOrPublisher="The International Trust for Zoological Nomenclature, London" pageId="38" pageNumber="39" title="International Code of Zoological Nomenclature. Fourth Edition." year="2000">ICZN (2000)</bibRefCitation>
<bibRefCitation id="7D6119A6E39871CCC8D06276B762BB77" author="ICZN," journalOrPublisher="The International Trust for Zoological Nomenclature, London" pageId="38" pageNumber="39" title="International Code of Zoological Nomenclature. Fourth Edition." year="2000">ICZN (2000)</bibRefCitation>
, a neotype may be designated when no name-bearing type specimen (i.e. holotype, lectotype, syntype or prior neotype) is believed to be extant and an author considers that a name-bearing type is necessary to define the nominal taxon objectively (Article 75.1). Article 75.3 asks also for qualifying conditions for the establishment of a neotype; among them:
<normalizedToken originalValue="….">....</normalizedToken>
<normalizedToken id="FC0DC69D811CD8FCB4B8FA8DAA5E66AD" originalValue="….">....</normalizedToken>
a statement that it is designated with the express purpose of clarifying the taxonomic status or the type locality of a nominal taxon (Article 75.3.1), and the
<normalizedToken originalValue="authors">author's</normalizedToken>
<normalizedToken id="CB2D08951B3424FF32B397E4D35C7CAE" originalValue="authors">author's</normalizedToken>
reasons for believing the
<normalizedToken originalValue="namebearing">name-bearing</normalizedToken>
<normalizedToken id="18D7E8DDB5479FD9A0CA8FB7BE154B64" originalValue="namebearing">name-bearing</normalizedToken>
type specimen(s)
<normalizedToken originalValue="…">...</normalizedToken>
<normalizedToken id="88CE3BFC33E4A2D42A04CC5E976BA3AF" originalValue="…">...</normalizedToken>
to be lost or destroyed, and the steps that have been taken to trace it or them (Article 73.3.4).
</paragraph>
<paragraph lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<paragraph id="A42B88D96618EED24EB08A00F8452A71" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
The specimens on which
<bibRefCitation author="Rouch, R" journalOrPublisher="Archives Zoologie Experimentale &amp; Generale" pageId="38" pageNumber="39" pagination="147 - 164" title="Deux harpacticidae nouveaux de l'ile de Long Island (Territoire de Papouasie et de Nouvelle-Guinee)." url="doi: 10.1023/A:1013148808110" volume="113" year="1972">Rouch (1972)</bibRefCitation>
<bibRefCitation id="B57F90C909F2D5C80F8FC29EF8609D68" author="Rouch, R" journalOrPublisher="Archives Zoologie Experimentale &amp; Generale" pageId="38" pageNumber="39" pagination="147 - 164" title="Deux harpacticidae nouveaux de l'ile de Long Island (Territoire de Papouasie et de Nouvelle-Guinee)." url="doi: 10.1023/A:1013148808110" volume="113" year="1972">Rouch (1972)</bibRefCitation>
based the original description of
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus bassoti" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bassoti">Phyllognathopus bassoti</taxonomicName>
<taxonomicName id="FA609DBCDB7B4AF23138ADE2D997536C" class="Copepoda" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus bassoti" order="Harpacticoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bassoti">Phyllognathopus bassoti</taxonomicName>
no longer exist, and most part of the
<normalizedToken originalValue="Rouchs">Rouch's</normalizedToken>
<normalizedToken id="2658A39EFD4D76C823635086F280970E" originalValue="Rouchs">Rouch's</normalizedToken>
collection has been lost (Rouch, in litt.).
<pageBreakToken pageId="16" pageNumber="17" start="start">Consequently</pageBreakToken>
<pageBreakToken id="8C2B419EEB39EFC6AF4917AF53F774A9" pageId="16" pageNumber="17" start="start">Consequently</pageBreakToken>
, the
<bibRefCitation author="ICZN," journalOrPublisher="The International Trust for Zoological Nomenclature, London" pageId="38" pageNumber="39" title="International Code of Zoological Nomenclature. Fourth Edition." year="2000">ICZN (2000)</bibRefCitation>
<bibRefCitation id="B5A7B13FECBF67F52F1F12E2DB078377" author="ICZN," journalOrPublisher="The International Trust for Zoological Nomenclature, London" pageId="38" pageNumber="39" title="International Code of Zoological Nomenclature. Fourth Edition." year="2000">ICZN (2000)</bibRefCitation>
recommendation 75A cannot be met, because no extant paratypes or paralectotypes, nor topotypic specimens are available, in order to select among them a neotype. The need to clarify the taxonomic status of this species, which is ranked herein to a new genus, imposed to follow another formal procedure, which, if
<pageBreakToken pageId="17" pageNumber="18" start="start">not</pageBreakToken>
<pageBreakToken id="FB0943A177B66DC5E535C684F7CB7342" pageId="17" pageNumber="18" start="start">not</pageBreakToken>
completely fulfills the ICZN rules (Article 75.3.6), is accepted by the Code (Article 76.3): the place of origin of the neotype becomes the type locality of the nominal species-group taxon, despite any previously published statement of the type locality. Following these arguments, a consistent population from India has been selected to establish the new genus, together with additional material from Indonesia. It is relevant to observe that other researchers have given consensus (
<bibRefCitation author="Bruno, MC" journalOrPublisher="Journal of Crustacean Biology" pageId="38" pageNumber="39" pagination="510 - 529" title="Harpacticoids from groundwaters in the Philippines: Parastenocaris mangyans, new species, Epactophanes philippinus, new species, and redescription of Phyllognathopus bassoti (Copepoda)." url="doi: 10.2307/1549260" volume="19" year="1999">Bruno and Cottarelli 1999</bibRefCitation>
<bibRefCitation id="6DF1A056E016F2C1D0F4389B6C7D8241" author="Bruno, MC" journalOrPublisher="Journal of Crustacean Biology" pageId="38" pageNumber="39" pagination="510 - 529" title="Harpacticoids from groundwaters in the Philippines: Parastenocaris mangyans, new species, Epactophanes philippinus, new species, and redescription of Phyllognathopus bassoti (Copepoda)." url="doi: 10.2307/1549260" volume="19" year="1999">Bruno and Cottarelli 1999</bibRefCitation>
,
<bibRefCitation author="Karanovic, T" journalOrPublisher="Annales de Limnologie" pageId="38" pageNumber="39" pagination="121 - 132" title="First record of Phyllognathopus bassoti Rouch, 1972 from India, with remarks on the family Phyllognathopodidae Gurney, 1932 (Crustacea, Copepoda, Harpacticoida)." url="doi: 10.1051/limn/2004010" volume="40" year="2004">Karanovic and Ranga Reddy 2004</bibRefCitation>
<bibRefCitation id="325589FADAA209BAD402A387F6F932FF" author="Karanovic, T" journalOrPublisher="Annales de Limnologie" pageId="38" pageNumber="39" pagination="121 - 132" title="First record of Phyllognathopus bassoti Rouch, 1972 from India, with remarks on the family Phyllognathopodidae Gurney, 1932 (Crustacea, Copepoda, Harpacticoida)." url="doi: 10.1051/limn/2004010" volume="40" year="2004">Karanovic and Ranga Reddy 2004</bibRefCitation>
) on the attribution of both populations to the species
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus bassoti" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bassoti">Phyllognathopus bassoti</taxonomicName>
<taxonomicName id="596071BA6285971E69F522F7C6DE02C5" class="Copepoda" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus bassoti" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bassoti">Phyllognathopus bassoti</taxonomicName>
described by
<bibRefCitation author="Rouch, R" journalOrPublisher="Archives Zoologie Experimentale &amp; Generale" pageId="38" pageNumber="39" pagination="147 - 164" title="Deux harpacticidae nouveaux de l'ile de Long Island (Territoire de Papouasie et de Nouvelle-Guinee)." url="doi: 10.1023/A:1013148808110" volume="113" year="1972">Rouch (1972)</bibRefCitation>
<bibRefCitation id="774E40F4953204A50588C56EFC62B9C6" author="Rouch, R" journalOrPublisher="Archives Zoologie Experimentale &amp; Generale" pageId="38" pageNumber="39" pagination="147 - 164" title="Deux harpacticidae nouveaux de l'ile de Long Island (Territoire de Papouasie et de Nouvelle-Guinee)." url="doi: 10.1023/A:1013148808110" volume="113" year="1972">Rouch (1972)</bibRefCitation>
from the Lake Wisdom (New Guinea).
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="18" type="neotype locality">
<paragraph pageId="17" pageNumber="18">Neotype locality.</paragraph>
<paragraph pageId="17" pageNumber="18">
<subSubSection id="992C05E77970754D583B4F65C4DA722B" pageId="17" pageNumber="18" type="neotype locality">
<paragraph id="7F9F7078E69D36AE096C3CF2E9F6B2F2" pageId="17" pageNumber="18">Neotype locality.</paragraph>
<paragraph id="438F4C1101246CA3A1E2CF16E8B43B9B" pageId="17" pageNumber="18">
India, Andhra Pradesh, town of Guntur, Brindavan Gardens, domestic water reservoir filled by a freshwater bore well; coordinates: approx.
<geoCoordinate direction="north" orientation="latitude" precision="925" value="16.3">16°18'N</geoCoordinate>
<geoCoordinate id="F4B91F2748C09330138E0973DC1C75F5" direction="north" orientation="latitude" precision="925" value="16.3">16°18'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="80.48333">80°29'E</geoCoordinate>
<geoCoordinate id="2E0AD62F8941CB695E6E5597D6639E40" direction="east" orientation="longitude" precision="925" value="80.48333">80°29'E</geoCoordinate>
(see
<bibRefCitation author="Karanovic, T" journalOrPublisher="Annales de Limnologie" pageId="38" pageNumber="39" pagination="121 - 132" title="First record of Phyllognathopus bassoti Rouch, 1972 from India, with remarks on the family Phyllognathopodidae Gurney, 1932 (Crustacea, Copepoda, Harpacticoida)." url="doi: 10.1051/limn/2004010" volume="40" year="2004">Karanovic and Ranga Reddy 2004</bibRefCitation>
<bibRefCitation id="AAAD7C233861BB246F3606380F0C0119" author="Karanovic, T" journalOrPublisher="Annales de Limnologie" pageId="38" pageNumber="39" pagination="121 - 132" title="First record of Phyllognathopus bassoti Rouch, 1972 from India, with remarks on the family Phyllognathopodidae Gurney, 1932 (Crustacea, Copepoda, Harpacticoida)." url="doi: 10.1051/limn/2004010" volume="40" year="2004">Karanovic and Ranga Reddy 2004</bibRefCitation>
, for more details).
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18" type="description based on the designed neotype">
<paragraph pageId="17" pageNumber="18">Description based on the designed neotype.</paragraph>
<paragraph lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="729754B48C31081D66ECE3294F66B052" lastPageId="20" lastPageNumber="21" pageId="17" pageNumber="18" type="description based on the designed neotype">
<paragraph id="6083D5B7D8E6721BE1DA902C4EAB4962" pageId="17" pageNumber="18">Description based on the designed neotype.</paragraph>
<paragraph id="076FDF9CF337027F26D910EF2E0430B4" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
FEMALE NEOTYPE. Body length, measured from tip of rostrum to posterior margin of caudal rami, 348
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="2483201E36C25EB0717D1EAE66FC0CC2" originalValue="µm">µm</normalizedToken>
. Habitus slig
<pageBreakToken pageId="18" pageNumber="19" start="start">htly</pageBreakToken>
<pageBreakToken id="77CB640F3CF53FB5C8F3C8C3854A22C1" pageId="18" pageNumber="19" start="start">htly</pageBreakToken>
dorsoventrally flattened (Fig. 17A), with no clear demarcation between prosome and urosome. Body depigmented and eyeless. Integumental dorsal window on cephalosome not confirmed. First pedigerous somite free. Integument without surface pits, moderately sclerotized. Cephalosome rounded; rostrum elongate, clearly articulated to cephalosome. Hyaline frills of cephalosome, somites bearing P1-P4 and urosome plain both dorsally and ventrally (Fig. 17 A, B). Cephalosome and both thoracic and abdominal somites (except fourth urosomite) with cuticular ornamentation represented by dorsal sensilla. P5-bearing somite with lateral paired and large pores (Fig. 18A). Female genital field located between first and second third of genital double-somite. Genital apparatus simplified; copulatory
<pageBreakToken pageId="19" pageNumber="20" start="start">pore</pageBreakToken>
<pageBreakToken id="D5D5B76401E61D2E8E1A44EA7BE6018A" pageId="19" pageNumber="20" start="start">pore</pageBreakToken>
located at half of genital double-somite. Seminal receptacles laterally located and condensed close to the lamellar sixth legs. Three spinular processes on free distal margin of anal operculum (Figs 17
<normalizedToken originalValue="AC">A-C</normalizedToken>
<normalizedToken id="7A61612E28FFBE9FD4942E32890829C9" originalValue="AC">A-C</normalizedToken>
). Caudal rami sub-quadrate, with incomplete setal pattern (6 setae). Dorsal seta inserted close to free distal margin of caudal ramus (Fig. 17C).
</paragraph>
<caption pageId="19" pageNumber="20">
<paragraph pageId="19" pageNumber="20">
<caption id="81EA470F11E30D9DB2537A79115E3AED" pageId="19" pageNumber="20">
<paragraph id="37CB22314D1F749C8C5684D2EA9BA4C3" pageId="19" pageNumber="20">
Figure 17.
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<taxonomicName id="C9D8013A17A2F492D8A131AC38E1EBE1" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
(Rouch, 1972), comb. n. (♀). A habitus, dorsal view B abdomen, ventral view C anal somite and operculum D antennule E antenna (scale bars in
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="4030C28500EF1BFDA01AFAE70FF7E8AA" originalValue="μm">μm</normalizedToken>
).
</paragraph>
</caption>
<caption pageId="19" pageNumber="20">
<paragraph pageId="19" pageNumber="20">
<caption id="5D3CB9A0E90355FD3000E7BA8F7BF2DE" pageId="19" pageNumber="20">
<paragraph id="4C0C4AA76D4605046E87B4D7BD1DBDB7" pageId="19" pageNumber="20">
Figure 18. SEM micrographs of
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<taxonomicName id="1263F043EBFD6264100190D2FDF4C745" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
(Rouch, 1972), comb. n. (♂). A lateral large pore (arrowed) on P5-bearing somite B anal somite and operculum.
</paragraph>
</caption>
<paragraph pageId="19" pageNumber="20">Antennule (Fig. 17D): consisting of 8 segments, segments 1 and 2 with long tube-pores. Armature formula: 1-[1], 2-[8], 3-[5], 4-[1 + (1 + ae)], 5-[1], 6-[3], 7-[4], 8-[6 + (1 + ae)]. Aesthetasc on segment 4 very large and long, well overreaching the last antennulary segment.</paragraph>
<paragraph pageId="20" pageNumber="21">
<pageBreakToken pageId="20" pageNumber="21" start="start">Antenna</pageBreakToken>
<paragraph id="EB63E901EA9264ADC6D621CC1B75DD82" pageId="19" pageNumber="20">Antennule (Fig. 17D): consisting of 8 segments, segments 1 and 2 with long tube-pores. Armature formula: 1-[1], 2-[8], 3-[5], 4-[1 + (1 + ae)], 5-[1], 6-[3], 7-[4], 8-[6 + (1 + ae)]. Aesthetasc on segment 4 very large and long, well overreaching the last antennulary segment.</paragraph>
<paragraph id="033ADC193ECD46D7D25F8F204ED2E1AA" pageId="20" pageNumber="21">
<pageBreakToken id="630D0C7C0684412AD9B42FEFFB300927" pageId="20" pageNumber="21" start="start">Antenna</pageBreakToken>
(Fig. 17E): exopod and armature of the second endopodal segment as in
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Phyllognathopus</taxonomicName>
<taxonomicName id="FBE1C60A398F89724FEA223594887C5B" class="Copepoda" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Phyllognathopus</taxonomicName>
and
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Parbatocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parbatocamptus" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Parbatocamptus</taxonomicName>
<taxonomicName id="7574B1D1CADC1797A6C6177148427037" class="Copepoda" family="Phyllognathopodidae" genus="Parbatocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parbatocamptus" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Parbatocamptus</taxonomicName>
. Exopod 1-segmented, with 3 lateral and 2 apical setae.
</paragraph>
<paragraph pageId="20" pageNumber="21">
<paragraph id="A609FE608DC9EB79D4E6E3EC3C7E17A4" pageId="20" pageNumber="21">
Mandible (Fig. 19A): mandibular palp biramous, basis with inner spinule row, exopod with 1 apical and 1 inner setae; endopod with 1 inner, 1 subapical and 2 apical setae. Armature of maxillule (Fig. 19B) and maxilla (Fig. 19C) as in
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Phyllognathopus</taxonomicName>
<taxonomicName id="D82A893754A6D12AF45A9D857B1709CA" class="Copepoda" family="Phyllognathopodidae" genus="Phyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllognathopus" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Phyllognathopus</taxonomicName>
.
</paragraph>
<caption pageId="20" pageNumber="21">
<paragraph pageId="20" pageNumber="21">
<caption id="435E763FFD72E076FEB45721DD56FA21" pageId="20" pageNumber="21">
<paragraph id="5CCFE25F61C634C7EB8DC5A08D5BDF59" pageId="20" pageNumber="21">
Figure 19.
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<taxonomicName id="E4EF9E225EFC9A52B67C5B70F239E61C" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
(Rouch, 1972), comb. n. (♀) A mandible B maxillule C maxilla D maxilliped E P4 F P5 G P5 (anomaly) H P6 (scale bars in
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="F28121B16193F422A2AD7C5440A87E15" originalValue="μm">μm</normalizedToken>
).
</paragraph>
</caption>
<paragraph pageId="20" pageNumber="21">
<paragraph id="D6FE1D6E6B812DA1117CA7AF1BC7CC98" pageId="20" pageNumber="21">
Maxilliped (Fig. 19D): phyllopodial, lamelliform, 1-segmented. Clear trace of ancestral 2-segmented condition marked by the presence of both outer and inner incisions. Armature consisting of 11 elements: 1 strong spine inserted at inner corner of former segment 1; 4 spines and 1 spiniform short seta inserted along inner margin, 5 bipinnate setae in apical position, armature topology basically referable to that of
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Parbatocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parbatocamptus" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Parbatocamptus</taxonomicName>
<taxonomicName id="9F9B09546E612A6FC8D605E6032383E9" class="Copepoda" family="Phyllognathopodidae" genus="Parbatocamptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parbatocamptus" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Parbatocamptus</taxonomicName>
.
</paragraph>
<paragraph pageId="20" pageNumber="21">
<paragraph id="3F3520F3CE0766D11D69EB5334EA344E" pageId="20" pageNumber="21">
P1-P3 with 3-segmented exopods and endopods. P4 with 2-segmented exopod and endopod. P1-P3 praecoxa well developed. P4 praecoxa absent (Fig. 19E). P1 exopod of about the same length of endopod. P2-P3 exopods longer than endopods, endopod not overreaching exp-2, fitting the original description (
<bibRefCitation author="Rouch, R" journalOrPublisher="Archives Zoologie Experimentale &amp; Generale" pageId="38" pageNumber="39" pagination="147 - 164" title="Deux harpacticidae nouveaux de l'ile de Long Island (Territoire de Papouasie et de Nouvelle-Guinee)." url="doi: 10.1023/A:1013148808110" volume="113" year="1972">Rouch 1972</bibRefCitation>
<bibRefCitation id="3D7B5669EB2A37C19A775EC5BA27013B" author="Rouch, R" journalOrPublisher="Archives Zoologie Experimentale &amp; Generale" pageId="38" pageNumber="39" pagination="147 - 164" title="Deux harpacticidae nouveaux de l'ile de Long Island (Territoire de Papouasie et de Nouvelle-Guinee)." url="doi: 10.1023/A:1013148808110" volume="113" year="1972">Rouch 1972</bibRefCitation>
) and the subsequent ones (
<bibRefCitation author="Bruno, MC" journalOrPublisher="Journal of Crustacean Biology" pageId="38" pageNumber="39" pagination="510 - 529" title="Harpacticoids from groundwaters in the Philippines: Parastenocaris mangyans, new species, Epactophanes philippinus, new species, and redescription of Phyllognathopus bassoti (Copepoda)." url="doi: 10.2307/1549260" volume="19" year="1999">Bruno and Cottarelli 1999</bibRefCitation>
<bibRefCitation id="92BE54E57CFBBBEF2BE3681954F02A33" author="Bruno, MC" journalOrPublisher="Journal of Crustacean Biology" pageId="38" pageNumber="39" pagination="510 - 529" title="Harpacticoids from groundwaters in the Philippines: Parastenocaris mangyans, new species, Epactophanes philippinus, new species, and redescription of Phyllognathopus bassoti (Copepoda)." url="doi: 10.2307/1549260" volume="19" year="1999">Bruno and Cottarelli 1999</bibRefCitation>
,
<bibRefCitation author="Karanovic, T" journalOrPublisher="Annales de Limnologie" pageId="38" pageNumber="39" pagination="121 - 132" title="First record of Phyllognathopus bassoti Rouch, 1972 from India, with remarks on the family Phyllognathopodidae Gurney, 1932 (Crustacea, Copepoda, Harpacticoida)." url="doi: 10.1051/limn/2004010" volume="40" year="2004">Karanovic and Ranga Reddy 2004</bibRefCitation>
<bibRefCitation id="EC48DCBB8379C9EC79D9843765BE6582" author="Karanovic, T" journalOrPublisher="Annales de Limnologie" pageId="38" pageNumber="39" pagination="121 - 132" title="First record of Phyllognathopus bassoti Rouch, 1972 from India, with remarks on the family Phyllognathopodidae Gurney, 1932 (Crustacea, Copepoda, Harpacticoida)." url="doi: 10.1051/limn/2004010" volume="40" year="2004">Karanovic and Ranga Reddy 2004</bibRefCitation>
).
</paragraph>
<paragraph pageId="20" pageNumber="21">P5 (Fig. 19F, G): free, with clear articulation to P5-bearing somite; right and left legs separate; baseoendopod and exopod coalescent, deep incision marking original segmentation between them; endopodal lobe well developed, elongate, rectangular in shape, longer than exopod, bearing 1 long pinnate seta, subdistally inserted, close to outer margin, and 2 spinule rows, the proximal one composed by tiny elements, the distal one of long spinules; exopodal lobe well discernible, with armature consisting of 4 (rarely 5 elements, observed in only one female) elements, the outermost seta inserted in subdistal outer position, the remaining ones in apical position; the outer apical seta slender and bipinnate, the remaining two spiniform. Basipodal outer seta present.</paragraph>
<paragraph pageId="20" pageNumber="21">P6 (Figs 19H, 20): rudimentary, consisting of small paired chitinous lamellar plates not coalescent along medial margin, partially covering seminal receptacles. Armature consisting of 1 short smooth spine with rounded tip on each side.</paragraph>
<caption pageId="20" pageNumber="21">
<paragraph pageId="20" pageNumber="21">
<paragraph id="F09FA29DC502FEADCEF86AB70E7191C4" pageId="20" pageNumber="21">P5 (Fig. 19F, G): free, with clear articulation to P5-bearing somite; right and left legs separate; baseoendopod and exopod coalescent, deep incision marking original segmentation between them; endopodal lobe well developed, elongate, rectangular in shape, longer than exopod, bearing 1 long pinnate seta, subdistally inserted, close to outer margin, and 2 spinule rows, the proximal one composed by tiny elements, the distal one of long spinules; exopodal lobe well discernible, with armature consisting of 4 (rarely 5 elements, observed in only one female) elements, the outermost seta inserted in subdistal outer position, the remaining ones in apical position; the outer apical seta slender and bipinnate, the remaining two spiniform. Basipodal outer seta present.</paragraph>
<paragraph id="68C99EDE0C15A9580DE5B20A680F1B42" pageId="20" pageNumber="21">P6 (Figs 19H, 20): rudimentary, consisting of small paired chitinous lamellar plates not coalescent along medial margin, partially covering seminal receptacles. Armature consisting of 1 short smooth spine with rounded tip on each side.</paragraph>
<caption id="737BF9CE9A4891717B581F44E036B223" pageId="20" pageNumber="21">
<paragraph id="273C02228573D450619CAE6C133B2E71" pageId="20" pageNumber="21">
Figure 20. SEM micrograph of
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<taxonomicName id="4883DA7EF40D56F2D280A071B807BEF0" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
(Rouch, 1972), comb. n. (♀): P6.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="male">
<paragraph pageId="20" pageNumber="21">Male.</paragraph>
<paragraph pageId="20" pageNumber="21">
<subSubSection id="F8C0CB066DC0891A0CC31BA272A3BFCD" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="male">
<paragraph id="835F6406B52D76CB49E82BC6A275D61E" pageId="20" pageNumber="21">Male.</paragraph>
<paragraph id="ADBDDEB5DFE9650FF98D2EC381372D37" pageId="20" pageNumber="21">
No marked sexual dimorphism in body size. Body length, measured from tip of rostrum to posterior margin of caudal rami, 335
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken id="0049F9EF7C18A63F438A9176F09A6757" originalValue="µm">µm</normalizedToken>
. Rostrum and ornamentation of cephalosome as in female (Fig. 21A). Male urosome consisting of 6 segments (Fig. 21A), third and fourth urosomites with deep ventral sockets (Figs 21B, 22A); socket on third urosomite plicate, with smooth free distal margins, and 2 setules laterally inserted close to the socket opening (Fig. 22B); socket on fourth urosomite with ornamented anterior margin, armed by strong spinules covering the opening (Fig. 22C). Anal somite with paired sensilla on dorsal side. Anal operculum protruding free distal margin of anal somite and extruded in 4 strong spinular processes, rarely 5 (in general anal operculum in males more armed than in females) (Fig. 18B). Antennule (Fig. 21C): basically 9-segmented, geniculation between segments 7 and 8; penultimate and last segments, each with suture line marking original segmentation between former segments 8 and 9, and 10 and 11, respectively. Long tube-pores on segments 1 and 2. Armature formula: 1-[1], 2-[8], 3-[8], 4-[1], 5-[7+(1 + ae)], 6-[2], 7-[1], 8-[1], 9-[10 + (1 + ae)].
</paragraph>
<caption pageId="20" pageNumber="21">
<paragraph pageId="20" pageNumber="21">
<caption id="2F146904A1AB771AAF28FADA279FF2D3" pageId="20" pageNumber="21">
<paragraph id="35524982FFAB18BA47434B744CBB6F10" pageId="20" pageNumber="21">
Figure 21.
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<taxonomicName id="1BF1085AE13C2B1C0925418CD501553F" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
(Rouch, 1972), comb. n. (♂). A habitus, dorsal view B abdomen, ventral view C antennule D P5 E P6 (scale bars in
<normalizedToken originalValue="μm">μm</normalizedToken>
<normalizedToken id="48ED731FA32DED9F1A6A2E1CB6B43FA8" originalValue="μm">μm</normalizedToken>
).
</paragraph>
</caption>
<caption pageId="20" pageNumber="21">
<paragraph pageId="20" pageNumber="21">
<caption id="B9BA0A5D329530ECC9F1049F86142A8D" pageId="20" pageNumber="21">
<paragraph id="EA7C4B278ECFF6C0563AD140743DDF77" pageId="20" pageNumber="21">
Figure 22. SEM micrographs of
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<taxonomicName id="39BC2352A61874DCA70B022FFE94F918" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
(Rouch, 1972), comb. n. (♂). A general view of the urosomal sockets B plicate socket on third urosomite (paired lateral setules arrowed) C socket on fourth urosomite covered by strong spinules.
</paragraph>
</caption>
<paragraph lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<paragraph id="D35547C2D518E749A762E1166EE5E74D" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
P5 (Figs 21D, 23A): free, with clear articulation to P5-bearing somite; right and left legs separate, intercoxal sclerite rudimentary but still discernible (Figs 21D, 23B). Basis of each leg well developed, representing most part of each leg; endopod rudimentary, consisting of a sclerotized and strong process articulated to basis. Row of surface spinules inserted near articulation between endopod and basis. Endopodal seta bipinnate, inserted on posterior surface of the endopod, close to its articulation to basis. Exopod distinct, clear articulated to the basis, wide and short, with unusual topology, being placed at the inner free distal margin of basis; exopodal armature consisting of 6 elements, all of which in apical position. Inner spinulose seta short and distinctly curved inward, the remaining setae of about the same length, 2 of which (the second and the fourth, beginning from the inner margin of the exopod) are respectively bipinnate and unipinnate; the remaining 3 smooth and slender, frequently closely adherent to each other and not easily discernible as distinct (Fig. 24). Male P6 (Figs 21E, 25
<normalizedToken originalValue="AB">A-B</normalizedToken>
<normalizedToken id="6F4050C88E7DA93586DDD6D91EC1B93F" originalValue="AB">A-B</normalizedToken>
): right and left legs distinct, closely adherent along inner margin, and symmetrical, each leg consisting of a well developed lamellar plate, with some spinule rows on the anterior surface. A membranous lamell
<pageBreakToken pageId="21" pageNumber="22" start="start">a</pageBreakToken>
<pageBreakToken id="1F0F6F00A28E593D8F1E4BC9923F64A5" pageId="21" pageNumber="22" start="start">a</pageBreakToken>
is observable between right and left P6 (rudimentary intercoxa?) (Fig. 25B); armature consisting of 2 inner spines of different length and 1 outer naked seta.
</paragraph>
<caption pageId="21" pageNumber="22">
<paragraph pageId="21" pageNumber="22">
<caption id="AC2549B0181B20E018647B27DF42FD2E" pageId="21" pageNumber="22">
<paragraph id="51C7BB132B46085914747886FF17190D" pageId="21" pageNumber="22">
Figure 23.
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<taxonomicName id="A9877EA4ABB3213624F70EBBCE2227F7" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
(Rouch, 1972), comb. n. (♂). A SEM micrograph of P5 B contrast phase micrograph of P5 rudimentary intercoxa (framed).
</paragraph>
</caption>
<caption pageId="21" pageNumber="22">
<paragraph pageId="21" pageNumber="22">
<caption id="48C0A7C16B8C7F299EBF43AE2968CA6B" pageId="21" pageNumber="22">
<paragraph id="5DF1A873971F5717DAF5A7881DA215D3" pageId="21" pageNumber="22">
Figure 24. SEM micrograph of
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<taxonomicName id="9706CC4070B43F767BC706714D0F506F" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
(Rouch, 1972), comb. n. (♂) P5, detail (white lines showing setae III and IV, which are closely adherent to each other and hardly discernible as distinct under contrast phase microscope).
</paragraph>
</caption>
<caption pageId="21" pageNumber="22">
<paragraph pageId="21" pageNumber="22">
<caption id="759F5414E58D8F09697E49E6A0F3C214" pageId="21" pageNumber="22">
<paragraph id="7640A042B12516E98D1096A19BC876C2" pageId="21" pageNumber="22">
Figure 25. SEM micrographs of
<taxonomicName class="Hexanauplia" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
<taxonomicName id="AF7E1593039EB4E38D24F410E12A100D" class="Copepoda" family="Phyllognathopodidae" genus="Neophyllognathopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neophyllognathopus bassoti" order="Harpacticoida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="bassoti">Neophyllognathopus bassoti</taxonomicName>
(Rouch, 1972), comb. n. (♂). A P5 and P6 B detail of P6 showing an interconnecting lamella (rudimentary intercoxa?) between right and left legs arrowed.
</paragraph>
</caption>

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@ -1,52 +1,52 @@
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<paragraph id="02AFBFCA7552392ED3C21D1F0DEDA088" pageId="0" pageNumber="211">
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<taxonomicName id="57685B36347E01693FF20B00D6729AED" ID-CoL="KPK" LSID="2DE65E55-8364-5499-B38E-F5BB21C98675" authority="gen. et sp. indet. 1" authorityName="Parry" authorityYear="1843" class="Insecta" family="Lucanidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="211" phylum="Arthropoda" rank="tribe" tribe="Ceratognathini">Ceratognathini gen. et sp. indet. 1</taxonomicName>
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@ -188,19 +188,19 @@ unambiguously based on the available morphology. I could not observe the details
;
<bibRefCitation id="1ADBDFE36482617B0946B37D0D203C42" author="Reid, C" editor="Ślipinski, A" journalOrPublisher="2). Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, Victoria" pageId="0" pageNumber="211" pagination="391 - 416" refId="B68" refString="Reid, C, 2019. Lucanidae Latreille, 1804. In: Ślipinski, A, Lawrence, JF, Eds., Australian Beetles (Vol. 2). Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, Victoria: 391 - 416" title="Lucanidae Latreille, 1804." volumeTitle="Australian Beetles (Vol." year="2019">Reid 2019</bibRefCitation>
). The new specimen lacks modified scales and setae on the dorsal surface, as in
<taxonomicName id="63A4FBEA15CEE11FC4837C1FB146EFCA" authorityName="Li &amp; Cai" authorityYear="2023" family="Lucanidae" genus="Oncelytris" kingdom="Animalia" lsidName="Oncelytris" order="Coleoptera" pageId="0" pageNumber="211" rank="genus">
<taxonomicName id="63A4FBEA15CEE11FC4837C1FB146EFCA" authorityName="Li &amp; Cai" authorityYear="2023" class="Insecta" family="Lucanidae" genus="Oncelytris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncelytris" order="Coleoptera" pageId="0" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="A6658D6D5733FE9231EFB4AFCDE3F4BD" italics="true" pageId="0" pageNumber="211">Oncelytris</emphasis>
</taxonomicName>
and the two other Kachin amber ceratognathins recorded here. The extinct taxon is treated provisionally herein as &quot;
<taxonomicName id="E3EA8B266BE9746E97AEE8A119426D3A" authorityName="Sharp" authorityYear="1899" lsidName="" pageId="0" pageNumber="211" rank="tribe" tribe="Ceratognathini">Ceratognathini</taxonomicName>
gen. et sp. indet. 1&quot; and can be differentiated from
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<taxonomicName id="A74981BBC9A1E33E5BE16879F582AB0C" authorityName="Yamamoto" authorityYear="2023" class="Insecta" family="Lucanidae" genus="Cretognathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cretognathus" order="Coleoptera" pageId="0" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="6A0FAAF220DAAC9397164A06C31C3018" italics="true" pageId="0" pageNumber="211">Cretognathus</emphasis>
</taxonomicName>
gen. nov. by having a much smaller head and antennal club, more slender tarsi, and different protibial edges (slightly different arrangement of subcontiguous tooth-like spines and much wider mid-dorsal teeth). Moreover, this new specimen does not seem to be congeneric with &quot;
<taxonomicName id="D420235AD879C0FC6A1C4DCB3F5FFBA8" authorityName="Sharp" authorityYear="1899" lsidName="" pageId="0" pageNumber="211" rank="tribe" tribe="Ceratognathini">Ceratognathini</taxonomicName>
gen. et sp. indet. 2&quot; by the distinctly smaller head and protibial morphology (shape, size, location, and number of the subequal tooth-like spines). Nevertheless, it is difficult to distinguish it from
<taxonomicName id="978193F214303F39DED10C04BF328D38" authorityName="Li &amp; Cai" authorityYear="2023" family="Lucanidae" genus="Oncelytris" kingdom="Animalia" lsidName="Oncelytris" order="Coleoptera" pageId="0" pageNumber="211" rank="genus">
<taxonomicName id="978193F214303F39DED10C04BF328D38" authorityName="Li &amp; Cai" authorityYear="2023" class="Insecta" family="Lucanidae" genus="Oncelytris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncelytris" order="Coleoptera" pageId="0" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="5E58DE0C3F7C9DD92D8EC8215E3ADF96" italics="true" pageId="0" pageNumber="211">Oncelytris</emphasis>
</taxonomicName>
, due to the poor state of preservation of the fossil. In fact, the dorsal and ventral surfaces of &quot;
@ -208,7 +208,7 @@ gen. et sp. indet. 2&quot; by the distinctly smaller head and protibial morpholo
gen. et sp. indet. 1&quot; are greatly distorted, preventing the accuate observation of the head and elytra whether it possesses tubercles and protuberances on the dorsum or not (cf. Li Y-D et al. 2023). However, it is also noted that the protibiae of &quot;
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gen. et sp. indet. 1&quot; is evidently arcuate, whereas those of
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<taxonomicName id="69579AE2552F025F9BDE8876933CADE6" authorityName="Li &amp; Cai" authorityYear="2023" class="Insecta" family="Lucanidae" genus="Oncelytris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Oncelytris" order="Coleoptera" pageId="0" pageNumber="211" phylum="Arthropoda" rank="genus">
<emphasis id="CF93EF5D848B1FFE351FE4B8859094E5" italics="true" pageId="0" pageNumber="211">Oncelytris</emphasis>
</taxonomicName>
are linear, suggesting that they may belong to different genera.

View file

@ -1,356 +1,356 @@
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<treatment ID-GBIF-Taxon="191980495" LSID="urn:lsid:plazi:treatment:2DE6ED23F86D52489E12BEB025803E88" httpUri="http://treatment.plazi.org/id/2DE6ED23F86D52489E12BEB025803E88" lastPageNumber="299" pageId="0" pageNumber="299">
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<paragraph pageId="0" pageNumber="299">
<taxonomicName LSID="2DE6ED23-F86D-5248-9E12-BEB025803E88" authority="(Thunberg, 1792)" baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="pictus">Plectorhinchus pictus (Thunberg, 1792)</taxonomicName>
<treatment id="2DE6ED23F86D52489E12BEB025803E88" ID-GBIF-Taxon="191980495" LSID="urn:lsid:plazi:treatment:2DE6ED23F86D52489E12BEB025803E88" httpUri="http://treatment.plazi.org/id/2DE6ED23F86D52489E12BEB025803E88" lastPageNumber="299" pageId="0" pageNumber="299" scope_family="Haemulidae" scope_order="Perciformes">
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<paragraph id="E5723A2DEE9149CA078F24041A8FB94E" pageId="0" pageNumber="299">
<taxonomicName id="C55FF74E85A2ED61F031D388CF504DF0" ID-CoL="4K3S7" LSID="2DE6ED23-F86D-5248-9E12-BEB025803E88" authority="(Thunberg, 1792)" baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" rank="species" species="pictus">Plectorhinchus pictus (Thunberg, 1792)</taxonomicName>
</paragraph>
</subSubSection>
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<paragraph pageId="0" pageNumber="299">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lateral view of collected specimen, a. Plectorhinchus pictus, subspecies Plectorhinchus pictus cinerascens (F 1804 SM- 22; 105 SL mm), striped juvenile phase; b. Plectorhinchus macrospilus (F 1803 SM- 67; 335 mm SL); c. Plectorhinchus andamanensis, (F 1803 SM- 19; 137 mm SL); d. Pomadasys argenteus (F 1602 Sb- 01; 132 mm SL)." figureDoi="10.3897/aiep.51.67043.figure2" httpUri="https://binary.pensoft.net/fig/587693" pageId="0" pageNumber="299">Local common name: futki datina (Bangla) Fig. 2a</figureCitation>
<subSubSection id="B89C33380D8CFE921055E9C810054E75" pageId="0" pageNumber="299" type="description">
<paragraph id="A7AF286192429FA408933EB18EC59060" pageId="0" pageNumber="299">
<figureCitation id="82E4FDF3E54BD02077775AD5F9E7E16A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lateral view of collected specimen, a. Plectorhinchus pictus, subspecies Plectorhinchus pictus cinerascens (F 1804 SM- 22; 105 SL mm), striped juvenile phase; b. Plectorhinchus macrospilus (F 1803 SM- 67; 335 mm SL); c. Plectorhinchus andamanensis, (F 1803 SM- 19; 137 mm SL); d. Pomadasys argenteus (F 1602 Sb- 01; 132 mm SL)." figureDoi="10.3897/aiep.51.67043.figure2" httpUri="https://binary.pensoft.net/fig/587693" pageId="0" pageNumber="299">Local common name: futki datina (Bangla) Fig. 2a</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="299" type="materials_examined">
<paragraph pageId="0" pageNumber="299">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="299">
<materialsCitation accessionNumber="MK340608" collectingDate="2018-04-20" country="Bangladesh" county="Saint Martin's Island" latitude="20.611" location="Cox's Bazar" longLatPrecision="1" longitude="92.326996" municipality="Bay of Bengal" specimenCount="2">
<collectingCountry name="Bangladesh">Bangladesh</collectingCountry>
<subSubSection id="F823E0407F2F49F698F68BCA959F580D" pageId="0" pageNumber="299" type="materials_examined">
<paragraph id="840394AEC2E9B4EBC3FAE7AB4890DB2E" pageId="0" pageNumber="299">Material examined.</paragraph>
<paragraph id="7AEC2F45E2DBB2320B5395BDEC80F209" pageId="0" pageNumber="299">
<materialsCitation id="A07B3DC7BF7C5177635A2FF420036B77" accessionNumber="MK340608" collectingDate="2018-04-20" country="Bangladesh" county="Saint Martin's Island" latitude="20.611" location="Cox's Bazar" longLatPrecision="1" longitude="92.326996" municipality="Bay of Bengal" specimenCount="2">
<collectingCountry id="80CC6B040941E93701DAE227D9B2223D" name="Bangladesh">Bangladesh</collectingCountry>
<specimenCount type="generic">2 specimens</specimenCount>
<specimenCount id="DBF8B1E8ADD2865518D9E568655D927B" type="generic">2 specimens</specimenCount>
; F1804SM-21 (
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.1" unit="mm" value="110.0">110 mm</quantity>
<quantity id="C6B3ADBA7A2E1EF595C17BBD4710B6FB" metricMagnitude="-1" metricUnit="m" metricValue="1.1" unit="mm" value="110.0">110 mm</quantity>
SL),
<location LSID="urn:lsid:plazi:treatment:2DE6ED23F86D52489E12BEB025803E88:FB818543162AEDF898867330EE9FD255" country="Bangladesh" county="Saint Martin's Island" latitude="20.611" longLatPrecision="1" longitude="92.326996" municipality="Bay of Bengal" name="Cox's Bazar">
<normalizedToken originalValue="Coxs">Cox's</normalizedToken>
<location id="FB818543162AEDF898867330EE9FD255" LSID="urn:lsid:plazi:treatment:2DE6ED23F86D52489E12BEB025803E88:FB818543162AEDF898867330EE9FD255" country="Bangladesh" county="Saint Martin's Island" latitude="20.611" longLatPrecision="1" longitude="92.326996" municipality="Bay of Bengal" name="Cox's Bazar">
<normalizedToken id="B7B1473D739A46F1C90166AE753FDFC8" originalValue="Coxs">Cox's</normalizedToken>
Bazar
</location>
,
<collectingMunicipality>Bay of Bengal</collectingMunicipality>
<collectingMunicipality id="354D54F279E3DC72FCE38C3AF52243E5">Bay of Bengal</collectingMunicipality>
,
<collectingCounty>
<collectingCounty id="DA037EAA5B2576025AA92063B125A104">
Saint
<normalizedToken originalValue="Martins">Martin's</normalizedToken>
<normalizedToken id="BC8CE26AC965CB58A08C4798259A2202" originalValue="Martins">Martin's</normalizedToken>
Island
</collectingCounty>
,
<geoCoordinate degrees="20" direction="north" minutes="36" orientation="latitude" precision="1" seconds="39.6" value="20.611">20°36'39.6&quot;N</geoCoordinate>
<geoCoordinate id="8A533A84942A49AA21F0F8F3A96FC87C" degrees="20" direction="north" minutes="36" orientation="latitude" precision="1" seconds="39.6" value="20.611">20°36'39.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="92" direction="east" minutes="19" orientation="longitude" precision="1" seconds="37.2" value="92.326996">92°19'37.2&quot;E</geoCoordinate>
<geoCoordinate id="BE062727195350220C5BB1818314917F" degrees="92" direction="east" minutes="19" orientation="longitude" precision="1" seconds="37.2" value="92.326996">92°19'37.2&quot;E</geoCoordinate>
,
<collectingDate value="2018-04-20">20 April 2018</collectingDate>
<collectingDate id="DD5024E14AC1C3C49D0C2DD41CFE3CD8" value="2018-04-20">20 April 2018</collectingDate>
, Amit Kumer Neogi, GenBank:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MK340608">MK340608</accessionNumber>
<accessionNumber id="A41735D51F9DAF4BA50907E04414AA08" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MK340608">MK340608</accessionNumber>
</materialsCitation>
;
<materialsCitation accessionNumber="MK340609" collectingDate="2018-04-20" country="Bangladesh" county="Saint Martin's Island" latitude="20.611" location="Cox's Bazar" longLatPrecision="1" longitude="92.326996" municipality="Bay of Bengal" specimenCount="1">
<materialsCitation id="C15C2501DA1FFC6DF6E3269074F292F3" accessionNumber="MK340609" collectingDate="2018-04-20" country="Bangladesh" county="Saint Martin's Island" latitude="20.611" location="Cox's Bazar" longLatPrecision="1" longitude="92.326996" municipality="Bay of Bengal" specimenCount="1">
F1804SM-22(
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.05" unit="mm" value="105.0">105 mm</quantity>
<quantity id="B2F93CD9489F4FC295FE51D070DE6B26" metricMagnitude="-1" metricUnit="m" metricValue="1.05" unit="mm" value="105.0">105 mm</quantity>
SL),
<location LSID="urn:lsid:plazi:treatment:2DE6ED23F86D52489E12BEB025803E88:D58DCE3B0F8CD30EAE8F3C7CF0B36AB0" country="Bangladesh" county="Saint Martin's Island" latitude="20.611" longLatPrecision="1" longitude="92.326996" municipality="Bay of Bengal" name="Cox's Bazar">
<normalizedToken originalValue="Coxs">Cox's</normalizedToken>
<location id="D58DCE3B0F8CD30EAE8F3C7CF0B36AB0" LSID="urn:lsid:plazi:treatment:2DE6ED23F86D52489E12BEB025803E88:D58DCE3B0F8CD30EAE8F3C7CF0B36AB0" country="Bangladesh" county="Saint Martin's Island" latitude="20.611" longLatPrecision="1" longitude="92.326996" municipality="Bay of Bengal" name="Cox's Bazar">
<normalizedToken id="CDE81E81AF06307DB255FCE62AC15247" originalValue="Coxs">Cox's</normalizedToken>
Bazar
</location>
,
<collectingMunicipality>Bay of Bengal</collectingMunicipality>
<collectingMunicipality id="F18580646C6F0E94CC6C7577907B71D1">Bay of Bengal</collectingMunicipality>
,
<collectingCounty>
<collectingCounty id="7A2FE160EC910A1460D0550C728008E7">
Saint
<normalizedToken originalValue="Martins">Martin's</normalizedToken>
<normalizedToken id="B1E0567420D3706FF7FBBD283E358AC2" originalValue="Martins">Martin's</normalizedToken>
Island
</collectingCounty>
,
<geoCoordinate degrees="20" direction="north" minutes="36" orientation="latitude" precision="1" seconds="39.6" value="20.611">20°36'39.6&quot;N</geoCoordinate>
<geoCoordinate id="A64ACF1C298042E95D8BB2F3949E6FEF" degrees="20" direction="north" minutes="36" orientation="latitude" precision="1" seconds="39.6" value="20.611">20°36'39.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="92" direction="east" minutes="19" orientation="longitude" precision="1" seconds="37.2" value="92.326996">92°19'37.2&quot;E</geoCoordinate>
<geoCoordinate id="4929E0AABBE596176CC7301DD322290C" degrees="92" direction="east" minutes="19" orientation="longitude" precision="1" seconds="37.2" value="92.326996">92°19'37.2&quot;E</geoCoordinate>
,
<collectingDate value="2018-04-20">20 April 2018</collectingDate>
<collectingDate id="79A9084CB038481A22EE1218849E4BAB" value="2018-04-20">20 April 2018</collectingDate>
, Amit Kumer Neogi, GenBank:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MK340609">MK340609</accessionNumber>
<accessionNumber id="E35DF9073001D009C5A503CAD5925FD9" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MK340609">MK340609</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="299" type="description">
<paragraph pageId="0" pageNumber="299">Diagnostic characters.</paragraph>
<paragraph pageId="0" pageNumber="299">Meristics: D-X, 23; P1-17; P2-I, 5; A-III, 7; C-18</paragraph>
<paragraph pageId="0" pageNumber="299">
<subSubSection id="43532456A8E3A4AFCEC6BFD15FF617CA" pageId="0" pageNumber="299" type="description">
<paragraph id="C3E0FB7D0D1142C9E269BCCBF5937E68" pageId="0" pageNumber="299">Diagnostic characters.</paragraph>
<paragraph id="BD584A60519B23946200EAE80B13CF3E" pageId="0" pageNumber="299">Meristics: D-X, 23; P1-17; P2-I, 5; A-III, 7; C-18</paragraph>
<paragraph id="BB9452FE08EAC2A8A8FD8B931B515878" pageId="0" pageNumber="299">
Body deep and compressed. Profile of snout steep; lips fleshy; chin with six pores, but no median pit; lower jaw without longitudinal groove at midline. Caudal peduncle slender and long; scales small, ctenoid. Color varying greatly with size. Juveniles dark brown to black above, 4 broad longitudinal black bands on the body, silvery yellow below; dorsal and caudal fins yellow with black patches and broken stripes (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lateral view of collected specimen, a. Plectorhinchus pictus, subspecies Plectorhinchus pictus cinerascens (F 1804 SM- 22; 105 SL mm), striped juvenile phase; b. Plectorhinchus macrospilus (F 1803 SM- 67; 335 mm SL); c. Plectorhinchus andamanensis, (F 1803 SM- 19; 137 mm SL); d. Pomadasys argenteus (F 1602 Sb- 01; 132 mm SL)." figureDoi="10.3897/aiep.51.67043.figure2" httpUri="https://binary.pensoft.net/fig/587693" pageId="0" pageNumber="299">2a</figureCitation>
<figureCitation id="25D0A7F6854C3864E7A5C5FB764E4D1B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Lateral view of collected specimen, a. Plectorhinchus pictus, subspecies Plectorhinchus pictus cinerascens (F 1804 SM- 22; 105 SL mm), striped juvenile phase; b. Plectorhinchus macrospilus (F 1803 SM- 67; 335 mm SL); c. Plectorhinchus andamanensis, (F 1803 SM- 19; 137 mm SL); d. Pomadasys argenteus (F 1602 Sb- 01; 132 mm SL)." figureDoi="10.3897/aiep.51.67043.figure2" httpUri="https://binary.pensoft.net/fig/587693" pageId="0" pageNumber="299">2a</figureCitation>
).
</paragraph>
<caption doi="10.3897/aiep.51.67043.figure2" httpUri="https://binary.pensoft.net/fig/587693" pageId="0" pageNumber="299" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="299">
<emphasis bold="true" pageId="0" pageNumber="299">Figure 2.</emphasis>
<caption id="2AA3B7DCB185BC8F64599BAE945BA1D0" doi="10.3897/aiep.51.67043.figure2" httpUri="https://binary.pensoft.net/fig/587693" pageId="0" pageNumber="299" start="Figure 2" startId="F2">
<paragraph id="8E9BA21F24E942595D6CDECD8F6A002B" pageId="0" pageNumber="299">
<emphasis id="099050C36955FF0E136353169DEF94F3" bold="true" pageId="0" pageNumber="299">Figure 2.</emphasis>
Lateral view of collected specimen,
<emphasis bold="true" pageId="0" pageNumber="299">a.</emphasis>
<taxonomicName baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="pictus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus</emphasis>
<emphasis id="0B8BBE4AD796EB1F50F3DB7630CC1008" bold="true" pageId="0" pageNumber="299">a.</emphasis>
<taxonomicName id="B309DC34A726585BA4F119BEA5AC5015" baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis id="953398841CB6942F69ED46C191A79FEE" italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus</emphasis>
</taxonomicName>
, subspecies
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus pictus subsp. cinerascens" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="subSpecies" species="pictus" subSpecies="cinerascens">Plectorhinchus pictus cinerascens</taxonomicName>
<taxonomicName id="BDBB285C5D792CF60EE98627DC19117B" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus pictus subsp. cinerascens" order="Perciformes" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="cinerascens">Plectorhinchus pictus cinerascens</taxonomicName>
(F1804SM-22; 105 SL mm), striped juvenile phase;
<emphasis bold="true" pageId="0" pageNumber="299">b.</emphasis>
<taxonomicName authorityName="Satapoomin et Randall" authorityYear="2000" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus macrospilus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="macrospilus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus macrospilus</emphasis>
<emphasis id="D451A1F4790243C87D6D4965D0798401" bold="true" pageId="0" pageNumber="299">b.</emphasis>
<taxonomicName id="3A08CC9470895979FE0D7496069EA3E4" authorityName="Satapoomin et Randall" authorityYear="2000" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus macrospilus" order="Perciformes" pageId="0" pageNumber="299" rank="species" species="macrospilus">
<emphasis id="B104FEE6FFACB78A09A002BF4D05D169" italics="true" pageId="0" pageNumber="299">Plectorhinchus macrospilus</emphasis>
</taxonomicName>
(F1803SM-67; 335 mm SL);
<emphasis bold="true" pageId="0" pageNumber="299">c.</emphasis>
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus andamanensis" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="andamanensis">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus andamanensis</emphasis>
<emphasis id="B7C1BD6B3BFADF39656FE0108B20913F" bold="true" pageId="0" pageNumber="299">c.</emphasis>
<taxonomicName id="E9E6474683287D539FA4F7B9C1F6FA4E" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus andamanensis" order="Perciformes" pageId="0" pageNumber="299" rank="species" species="andamanensis">
<emphasis id="91BEE1F79C5CC20A204833A19CA07FF4" italics="true" pageId="0" pageNumber="299">Plectorhinchus andamanensis</emphasis>
</taxonomicName>
, (F1803SM-19; 137 mm SL);
<emphasis bold="true" pageId="0" pageNumber="299">d.</emphasis>
<taxonomicName baseAuthorityName="Forsskal" baseAuthorityYear="1775" class="Actinopterygii" family="Haemulidae" genus="Pomadasys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pomadasys argenteus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="argenteus">
<emphasis italics="true" pageId="0" pageNumber="299">Pomadasys argenteus</emphasis>
<emphasis id="895C45C43EB9F63B434F4F84A5E74296" bold="true" pageId="0" pageNumber="299">d.</emphasis>
<taxonomicName id="CEC92AABEA4EEA8E6DD621981F38C43A" baseAuthorityName="Forsskal" baseAuthorityYear="1775" family="Haemulidae" genus="Pomadasys" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pomadasys argenteus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="argenteus">
<emphasis id="F3872C267F1B9E5A66367C4A798C3478" italics="true" pageId="0" pageNumber="299">Pomadasys argenteus</emphasis>
</taxonomicName>
(F1602Sb-01; 132 mm SL).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="299" type="remarks">
<paragraph pageId="0" pageNumber="299">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="299">
<subSubSection id="663C136CC5DBD7634230886D99571D17" pageId="0" pageNumber="299" type="remarks">
<paragraph id="D6AEC9554463BEC2D5D04B776F12C521" pageId="0" pageNumber="299">Remarks.</paragraph>
<paragraph id="3913CB1A23E8686F20A6E4EB20274145" pageId="0" pageNumber="299">
This species has previously been included by the majority of authors in the genus
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
<taxonomicName id="2D41D05590DE8A016A0C137BB1465106" class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis id="483B31433276711C54A155C09286A451" italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
</taxonomicName>
Oken, 1817, but recent phylogenetic analyses (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2966.1.4" author="Sanciangco, MD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="299" pagination="37 - 50" refId="B41" refString="Sanciangco, MD, Rocha, LA, Carpenter, KE, 2011. A molecular phylogeny of the grunts (Perciformes: Haemulidae) inferred using mitochondrial and nuclear genes. Zootaxa 2966: 37 - 50, DOI: https://doi.org/10.11646/zootaxa.2966.1.4" title="A molecular phylogeny of the grunts (Perciformes: Haemulidae) inferred using mitochondrial and nuclear genes." url="https://doi.org/10.11646/zootaxa.2966.1.4" volume="2966" year="2011">Sanciangco et al. 2011</bibRefCitation>
<bibRefCitation id="E998D4492B63F50B0D0CB944E52A118F" DOI="https://doi.org/10.11646/zootaxa.2966.1.4" author="Sanciangco, MD" journalOrPublisher="Zootaxa" pageId="0" pageNumber="299" pagination="37 - 50" refId="B41" refString="Sanciangco, MD, Rocha, LA, Carpenter, KE, 2011. A molecular phylogeny of the grunts (Perciformes: Haemulidae) inferred using mitochondrial and nuclear genes. Zootaxa 2966: 37 - 50, DOI: https://doi.org/10.11646/zootaxa.2966.1.4" title="A molecular phylogeny of the grunts (Perciformes: Haemulidae) inferred using mitochondrial and nuclear genes." url="https://doi.org/10.11646/zootaxa.2966.1.4" volume="2966" year="2011">Sanciangco et al. 2011</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1186/1471-2148-12-57" author="Tavera, J" journalOrPublisher="Trends in Ecology and Evolution" pageId="0" pageNumber="299" refId="B48" refString="Tavera, J, Acero, A, Balart, E, Bernardi, G, 2012. Molecular phylogeny of grunts (Teleostei, Haemulidae), with an emphasis on the ecology, evolution, and speciation history of New World species. BMC Evolutionary Biology 12: 57. https://doi.org/10.1186/1471-2148-12-57" title="Molecular phylogeny of grunts (Teleostei, Haemulidae), with an emphasis on the ecology, evolution, and speciation history of New World species. BMC Evolutionary Biology 12: 57." url="https://doi.org/10.1186/1471-2148-12-57" year="2012">Tavera et al. 2012</bibRefCitation>
<bibRefCitation id="17336D674E72C83C316277EB2A1D6B15" DOI="https://doi.org/10.1186/1471-2148-12-57" author="Tavera, J" journalOrPublisher="Trends in Ecology and Evolution" pageId="0" pageNumber="299" refId="B48" refString="Tavera, J, Acero, A, Balart, E, Bernardi, G, 2012. Molecular phylogeny of grunts (Teleostei, Haemulidae), with an emphasis on the ecology, evolution, and speciation history of New World species. BMC Evolutionary Biology 12: 57. https://doi.org/10.1186/1471-2148-12-57" title="Molecular phylogeny of grunts (Teleostei, Haemulidae), with an emphasis on the ecology, evolution, and speciation history of New World species. BMC Evolutionary Biology 12: 57." url="https://doi.org/10.1186/1471-2148-12-57" year="2012">Tavera et al. 2012</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2017.12.032" author="Tavera, J" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="299" pagination="213 - 223" refId="B49" refString="Tavera, J, Acero, PA, Wainwright, PC, 2018. Multilocus phylogeny, divergence times, and a major role for the benthic-to-pelagic axis in the diversification of grunts (Haemulidae). Molecular Phylogenetics and Evolution 121: 213 - 223, DOI: https://doi.org/10.1016/j.ympev.2017.12.032" title="Multilocus phylogeny, divergence times, and a major role for the benthic-to-pelagic axis in the diversification of grunts (Haemulidae)." url="https://doi.org/10.1016/j.ympev.2017.12.032" volume="121" year="2018">Tavera et al. 2018</bibRefCitation>
<bibRefCitation id="42DD0B5723067E4685F7950F30230E11" DOI="https://doi.org/10.1016/j.ympev.2017.12.032" author="Tavera, J" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="299" pagination="213 - 223" refId="B49" refString="Tavera, J, Acero, PA, Wainwright, PC, 2018. Multilocus phylogeny, divergence times, and a major role for the benthic-to-pelagic axis in the diversification of grunts (Haemulidae). Molecular Phylogenetics and Evolution 121: 213 - 223, DOI: https://doi.org/10.1016/j.ympev.2017.12.032" title="Multilocus phylogeny, divergence times, and a major role for the benthic-to-pelagic axis in the diversification of grunts (Haemulidae)." url="https://doi.org/10.1016/j.ympev.2017.12.032" volume="121" year="2018">Tavera et al. 2018</bibRefCitation>
) have shown that
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
<taxonomicName id="A807329E781CEA10307655AA51383B53" class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis id="D8E6AFFA40FE2005CACB4EF5E4E4944E" italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
</taxonomicName>
(including type species
<taxonomicName genus="D." lsidName="D. pictus" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis italics="true" pageId="0" pageNumber="299">D. pictus</emphasis>
<taxonomicName id="D1C3790CAC6AAB65E96D31C3EFB28A95" lsidName="D. pictus" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis id="562A552FAC74C297A83106276B12FDDC" italics="true" pageId="0" pageNumber="299">D. pictus</emphasis>
</taxonomicName>
) is deeply nested within
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
<taxonomicName id="E95F44F3CBCB6FC176710BBE380936C8" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" rank="genus">
<emphasis id="1DDEE2D62369DC7BC27B1B44EA0961DF" italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
</taxonomicName>
(and sister to its type species
<taxonomicName authorityName="Lacepede" authorityYear="1801" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus chaetodonoides" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="chaetodonoides">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus chaetodonoides</emphasis>
<taxonomicName id="D819728D921BCA6D1882378F79472050" authorityName="Lacepede" authorityYear="1801" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus chaetodonoides" order="Perciformes" pageId="0" pageNumber="299" rank="species" species="chaetodonoides">
<emphasis id="7AE53ACC4278851B344E0880ED8DB5C5" italics="true" pageId="0" pageNumber="299">Plectorhinchus chaetodonoides</emphasis>
</taxonomicName>
<normalizedToken originalValue="Lacepède">Lacepede</normalizedToken>
<normalizedToken id="A115518C7262AA7849E24C74925E548F" originalValue="Lacepède">Lacepede</normalizedToken>
, 1801), and
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2017.12.032" author="Tavera, J" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="299" pagination="213 - 223" refId="B49" refString="Tavera, J, Acero, PA, Wainwright, PC, 2018. Multilocus phylogeny, divergence times, and a major role for the benthic-to-pelagic axis in the diversification of grunts (Haemulidae). Molecular Phylogenetics and Evolution 121: 213 - 223, DOI: https://doi.org/10.1016/j.ympev.2017.12.032" title="Multilocus phylogeny, divergence times, and a major role for the benthic-to-pelagic axis in the diversification of grunts (Haemulidae)." url="https://doi.org/10.1016/j.ympev.2017.12.032" volume="121" year="2018">Tavera et al. (2018)</bibRefCitation>
<bibRefCitation id="15241E0CDEA337927F91A52E2F7F01CF" DOI="https://doi.org/10.1016/j.ympev.2017.12.032" author="Tavera, J" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="299" pagination="213 - 223" refId="B49" refString="Tavera, J, Acero, PA, Wainwright, PC, 2018. Multilocus phylogeny, divergence times, and a major role for the benthic-to-pelagic axis in the diversification of grunts (Haemulidae). Molecular Phylogenetics and Evolution 121: 213 - 223, DOI: https://doi.org/10.1016/j.ympev.2017.12.032" title="Multilocus phylogeny, divergence times, and a major role for the benthic-to-pelagic axis in the diversification of grunts (Haemulidae)." url="https://doi.org/10.1016/j.ympev.2017.12.032" volume="121" year="2018">Tavera et al. (2018)</bibRefCitation>
have proposed that
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
<taxonomicName id="06D6CACDA946205411CE6DA9BDD4D335" class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis id="7729D9DC599056B12F4898FB7FD5321A" italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
</taxonomicName>
should be considered a junior synonym of
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
<taxonomicName id="FB5ABC90E97A3B23B98B3D621AF8F1CC" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" rank="genus">
<emphasis id="4D7B9D1F1F162D1CBAFB65CA1C4BCB93" italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
</taxonomicName>
<normalizedToken originalValue="Lacepède">Lacepede</normalizedToken>
<normalizedToken id="D115679A2E5327C12D9B322A8D84AA6B" originalValue="Lacepède">Lacepede</normalizedToken>
, 1801. As pointed out by
<bibRefCitation author="Parenti, P" journalOrPublisher="Iranian Journal of Ichthyology" pageId="0" pageNumber="299" pagination="150 - 196" refId="B32" refString="Parenti, P, 2019. An annotated checklist of the fishes of the family Haemulidae (Teleostei: Perciformes). Iranian Journal of Ichthyology 6 (3): 150 - 196" title="An annotated checklist of the fishes of the family Haemulidae (Teleostei: Perciformes)." volume="6" year="2019">Parenti (2019)</bibRefCitation>
<bibRefCitation id="50196C2309376F0C96EA2672E101361B" author="Parenti, P" journalOrPublisher="Iranian Journal of Ichthyology" pageId="0" pageNumber="299" pagination="150 - 196" refId="B32" refString="Parenti, P, 2019. An annotated checklist of the fishes of the family Haemulidae (Teleostei: Perciformes). Iranian Journal of Ichthyology 6 (3): 150 - 196" title="An annotated checklist of the fishes of the family Haemulidae (Teleostei: Perciformes)." volume="6" year="2019">Parenti (2019)</bibRefCitation>
, however, accepting
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
<taxonomicName id="79BDECC56D0D9609013400AD7EB1CB8D" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" rank="genus">
<emphasis id="50A27E7E7AFB173303E229F90AE0FBB5" italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
</taxonomicName>
as the senior synonym of
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
<taxonomicName id="0FB4CAD4686D5584E8DDA86DC78CBD0E" class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis id="ECC7AC0F665CB9A8A031D52FC35BD8CC" italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
</taxonomicName>
creates several nomenclatural problems:
<taxonomicName baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="pictus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus</emphasis>
<taxonomicName id="0B4BD5D8192071CEDFDC97B5CA417EDA" baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis id="5D5508C2AC73DE45D0D343564E7A4087" italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus</emphasis>
</taxonomicName>
(Tortonese, 1936) becomes a secondary homonym of
<taxonomicName genus="P." lsidName="P. pictus" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis italics="true" pageId="0" pageNumber="299">P. pictus</emphasis>
<taxonomicName id="60A70B68148D6E76AA3AC1F3246AB9B8" lsidName="P. pictus" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis id="0EA322496A3AF2CFF25DFE50D3C08E32" italics="true" pageId="0" pageNumber="299">P. pictus</emphasis>
</taxonomicName>
(Thunberg, 1792), and the next available name for
<taxonomicName genus="P." lsidName="P. pictus" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis italics="true" pageId="0" pageNumber="299">P. pictus</emphasis>
<taxonomicName id="78C5B7DE058DF9C3CC150837C0520820" lsidName="P. pictus" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis id="05F205E9C61AF0065DDEEACD3BB3702F" italics="true" pageId="0" pageNumber="299">P. pictus</emphasis>
</taxonomicName>
(Tortonese) is
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus cinctus subsp. punctatatus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="subSpecies" species="cinctus" subSpecies="punctatatus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus cinctus punctatatus</emphasis>
<taxonomicName id="B7C2980ABE466255169D4DD9D332EA85" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus cinctus subsp. punctatatus" order="Perciformes" pageId="0" pageNumber="299" rank="subSpecies" species="cinctus" subSpecies="punctatatus">
<emphasis id="B52DD0AD6C63758C2FE8E793A160967B" italics="true" pageId="0" pageNumber="299">Plectorhinchus cinctus punctatatus</emphasis>
</taxonomicName>
Fang, 1942 (see
<bibRefCitation author="Fricke, R" journalOrPublisher="Molecular Ecology" pageId="0" pageNumber="299" publicationUrl="http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" refId="B10" refString="Fricke, R, Eschmeyer, WN, Van der Laan, R, 2021b. Eschmeyer's catalog of Fishes: genera, species, references. California Academy of Sciences, San Francisco, CA, USA. [Accessed on 23 April 2021] http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" title="Eschmeyer's catalog of Fishes: genera, species, references. California Academy of Sciences, San Francisco, CA, USA. [Accessed on 23 April 2021]" url="http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" year="2021 b">Fricke et al. 2021b</bibRefCitation>
<bibRefCitation id="C8382BF4566E9D9A284F6CD764FD1882" author="Fricke, R" journalOrPublisher="Molecular Ecology" pageId="0" pageNumber="299" publicationUrl="http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" refId="B10" refString="Fricke, R, Eschmeyer, WN, Van der Laan, R, 2021b. Eschmeyer's catalog of Fishes: genera, species, references. California Academy of Sciences, San Francisco, CA, USA. [Accessed on 23 April 2021] http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" title="Eschmeyer's catalog of Fishes: genera, species, references. California Academy of Sciences, San Francisco, CA, USA. [Accessed on 23 April 2021]" url="http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" year="2021 b">Fricke et al. 2021b</bibRefCitation>
) which is secondarily preoccupied in
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
<taxonomicName id="AFE2A7A08B17A7E523C909C656FA9A7A" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" rank="genus">
<emphasis id="8E133455A3EB61EA05C00C3DD702CDC6" italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
</taxonomicName>
by
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma puncatatum" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="puncatatum">
<emphasis italics="true" pageId="0" pageNumber="299">Diagramma puncatatum</emphasis>
<taxonomicName id="A9E60B7B9720C70F564343472F957162" class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma puncatatum" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="puncatatum">
<emphasis id="F687115880AC7B1DF2E67677615B5752" italics="true" pageId="0" pageNumber="299">Diagramma puncatatum</emphasis>
</taxonomicName>
Cuvier, 1830 (see
<bibRefCitation author="Fricke, R" journalOrPublisher="Molecular Ecology" pageId="0" pageNumber="299" publicationUrl="http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" refId="B10" refString="Fricke, R, Eschmeyer, WN, Van der Laan, R, 2021b. Eschmeyer's catalog of Fishes: genera, species, references. California Academy of Sciences, San Francisco, CA, USA. [Accessed on 23 April 2021] http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" title="Eschmeyer's catalog of Fishes: genera, species, references. California Academy of Sciences, San Francisco, CA, USA. [Accessed on 23 April 2021]" url="http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" year="2021 b">Fricke et al. 2021b</bibRefCitation>
<bibRefCitation id="F920C72D94EBE8E07E3DECEEAD9E713B" author="Fricke, R" journalOrPublisher="Molecular Ecology" pageId="0" pageNumber="299" publicationUrl="http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" refId="B10" refString="Fricke, R, Eschmeyer, WN, Van der Laan, R, 2021b. Eschmeyer's catalog of Fishes: genera, species, references. California Academy of Sciences, San Francisco, CA, USA. [Accessed on 23 April 2021] http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" title="Eschmeyer's catalog of Fishes: genera, species, references. California Academy of Sciences, San Francisco, CA, USA. [Accessed on 23 April 2021]" url="http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp" year="2021 b">Fricke et al. 2021b</bibRefCitation>
). This problem was long ago recognized by
<bibRefCitation author="Whitley, GP" journalOrPublisher="Series B, Biological Sciences" pageId="0" pageNumber="299" refId="B55" refString="Whitley, GP, 1951. New fish names and records. Proceedings of the Royal Zoological Society of New South Wales 1949-1950: 61-68." title="New fish names and records. Proceedings of the Royal Zoological Society of New South Wales 1949 - 1950: 61 - 68." year="1951">Whitley (1951)</bibRefCitation>
<bibRefCitation id="DBB8D1A64565B151FFC6B52C0F9E0C54" author="Whitley, GP" journalOrPublisher="Series B, Biological Sciences" pageId="0" pageNumber="299" refId="B55" refString="Whitley, GP, 1951. New fish names and records. Proceedings of the Royal Zoological Society of New South Wales 1949-1950: 61-68." title="New fish names and records. Proceedings of the Royal Zoological Society of New South Wales 1949 - 1950: 61 - 68." year="1951">Whitley (1951)</bibRefCitation>
, who anticipated
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
<taxonomicName id="1EE2E9E762212C02CA802A4F84B9DD5B" class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis id="E73D5663CE281682643E11B57ABD2B97" italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
<taxonomicName id="A431C75FC42640027E8E03D3A47BA522" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus" order="Perciformes" pageId="0" pageNumber="299" rank="genus">
<emphasis id="9332FC6AA9FA3266C4C998D6AE7D9C21" italics="true" pageId="0" pageNumber="299">Plectorhinchus</emphasis>
</taxonomicName>
and accordingly proposed the name
<taxonomicName authorityName="Whitley" authorityYear="1951" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus fangi" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="fangi">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus fangi</emphasis>
<taxonomicName id="9E3FA1615D7FB958F686253293FA9FC9" authorityName="Whitley" authorityYear="1951" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus fangi" order="Perciformes" pageId="0" pageNumber="299" rank="species" species="fangi">
<emphasis id="13FE7A3646A5D645A1CD869252592C67" italics="true" pageId="0" pageNumber="299">Plectorhinchus fangi</emphasis>
</taxonomicName>
Whitley, 1951, as a replacement for
<taxonomicName genus="P." lsidName="P. cinctus subsp. punctatus" pageId="0" pageNumber="299" rank="subSpecies" species="cinctus" subSpecies="punctatus">
<emphasis italics="true" pageId="0" pageNumber="299">P. cinctus punctatus</emphasis>
<taxonomicName id="179202ECD0836217821CE80DA27E8E71" lsidName="P. cinctus subsp. punctatus" pageId="0" pageNumber="299" rank="subSpecies" species="cinctus" subSpecies="punctatus">
<emphasis id="AC11790911891F03E8AC877261A2FC82" italics="true" pageId="0" pageNumber="299">P. cinctus punctatus</emphasis>
</taxonomicName>
Fang (a species not yet recorded from the Bay of Bengal). Thus, we regard
<taxonomicName baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="pictus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus</emphasis>
<taxonomicName id="A50795229FDB82F6BD79F62D4021ED1F" baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis id="6231A5F6DB267605DAB735F9C64E8F1C" italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus</emphasis>
</taxonomicName>
(Thunberg, 1792) as the correct name for the species described here from Bangladesh.
</paragraph>
<paragraph pageId="0" pageNumber="299">
<paragraph id="B1FF70B621AF9D9140BC3330C9C4F157" pageId="0" pageNumber="299">
Specimens of
<taxonomicName genus="P." lsidName="P. pictus" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis italics="true" pageId="0" pageNumber="299">P. pictus</emphasis>
<taxonomicName id="4DA48E718D32380126923AE31D75AD4C" lsidName="P. pictus" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis id="3FFA445121AD902D7BE624F7733E57C7" italics="true" pageId="0" pageNumber="299">P. pictus</emphasis>
</taxonomicName>
from the Bay of Bengal are part of a distinct subspecies,
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus pictus subsp. cinerascens" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="subSpecies" species="pictus" subSpecies="cinerascens">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus cinerascens</emphasis>
<taxonomicName id="F1F926B00D3DFB2B8C22D796C1EAC49A" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus pictus subsp. cinerascens" order="Perciformes" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="cinerascens">
<emphasis id="B6E7EB47B65B82FFB60162EE2010AB08" italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus cinerascens</emphasis>
</taxonomicName>
Cuvier, 1830-one of five geographically separate subspecies recognized by
<bibRefCitation author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. (2001</bibRefCitation>
<bibRefCitation id="86D3F06B34C1C8BDDBA050F6C59C7EDB" author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. (2001</bibRefCitation>
, as
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
<taxonomicName id="A3A3F2C64A721CA44FFA52047189DFD7" class="Actinopterygii" family="Haemulidae" genus="Diagramma" kingdom="Animalia" lsidName="Diagramma" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="genus">
<emphasis id="6F1A4BE6AEE9C0251558E7BC24F6226F" italics="true" pageId="0" pageNumber="299">Diagramma</emphasis>
</taxonomicName>
):
<taxonomicName authority="pictus" genus="P." lsidName="P. pictus subsp. pictus" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="pictus">
<emphasis italics="true" pageId="0" pageNumber="299">P. pictus pictus</emphasis>
<taxonomicName id="18BF388A7005C70A1BAF9BDC9DA19D59" authority="pictus" lsidName="P. pictus subsp. pictus" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="pictus">
<emphasis id="B908E58A0CFD8A8FB9D113E618AAD573" italics="true" pageId="0" pageNumber="299">P. pictus pictus</emphasis>
</taxonomicName>
(Thunberg),
<taxonomicName genus="P." lsidName="P. pictus subsp. labiosum" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="labiosum">
<emphasis italics="true" pageId="0" pageNumber="299">P. pictus labiosum</emphasis>
<taxonomicName id="A11F53FA8B5624A643ED06E7286A448B" lsidName="P. pictus subsp. labiosum" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="labiosum">
<emphasis id="0F329554791D661FD3270FAAD968C12C" italics="true" pageId="0" pageNumber="299">P. pictus labiosum</emphasis>
</taxonomicName>
Macleay,
<taxonomicName genus="P." lsidName="P. pictus subsp. punctatum" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="punctatum">
<emphasis italics="true" pageId="0" pageNumber="299">P. pictus punctatum</emphasis>
<taxonomicName id="54EF10562255F4E3983625D726B49719" lsidName="P. pictus subsp. punctatum" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="punctatum">
<emphasis id="01E437A18BABFE92B9B180067855B93E" italics="true" pageId="0" pageNumber="299">P. pictus punctatum</emphasis>
</taxonomicName>
Cuvier, and
<taxonomicName genus="P." lsidName="P. pictus subsp. centurio" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="centurio">
<emphasis italics="true" pageId="0" pageNumber="299">P. pictus centurio</emphasis>
<taxonomicName id="79C9626A911FA169B38330C8AE34A618" lsidName="P. pictus subsp. centurio" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="centurio">
<emphasis id="D7FD4C1ABB74CEBEC6EE8DF1D855F79C" italics="true" pageId="0" pageNumber="299">P. pictus centurio</emphasis>
</taxonomicName>
Cuvier).
<taxonomicName baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="species" species="pictus">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus</emphasis>
<taxonomicName id="0F5098BB36F91248C5E4AC0AE96860DA" baseAuthorityName="Thunberg" baseAuthorityYear="1792" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus pictus" order="Perciformes" pageId="0" pageNumber="299" rank="species" species="pictus">
<emphasis id="E77AC70342205D46249AF402EF675B04" italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus</emphasis>
</taxonomicName>
, subspecies
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus pictus subsp. cinerascens" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="subSpecies" species="pictus" subSpecies="cinerascens">Plectorhinchus pictus cinerascens</taxonomicName>
<taxonomicName id="DB607732996B98E71C3915681C9BCD9A" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus pictus subsp. cinerascens" order="Perciformes" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="cinerascens">Plectorhinchus pictus cinerascens</taxonomicName>
of
<bibRefCitation author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. (2001)</bibRefCitation>
<bibRefCitation id="FB4A82DBEA8E9CF1406C61DFEC0800BB" author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. (2001)</bibRefCitation>
differs from all others in the configuration of the body stripes of juveniles, size of spots, and nature of the lines on the cheeks and operculum of large juveniles to small adults. Juveniles of about 150 mm TL often have broad, clearly defined body stripes, not yet beginning to break into broken lines or numerous spots; and progress from a striped to a fully spotted phase at 180-240 mm TL (
<bibRefCitation author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. 2001</bibRefCitation>
<bibRefCitation id="EAF8C6EB1F4B18B970F2745BE4F0A3DE" author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. 2001</bibRefCitation>
: fig. 5). A more detailed genetic study is necessary to determine whether this geographically distinct color variety should be afforded separate species recognition (
<bibRefCitation author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. 2001</bibRefCitation>
<bibRefCitation id="702E5D4E6DA8652C6BAEE7F6F693D8A7" author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="299" type="distribution">
<paragraph pageId="0" pageNumber="299">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="299">
<taxonomicName class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plectorhinchus pictus subsp. cinerascens" order="Perciformes" pageId="0" pageNumber="299" phylum="Chordata" rank="subSpecies" species="pictus" subSpecies="cinerascens">
<emphasis italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus cinerascens</emphasis>
<subSubSection id="6A0BF1BC2B0FE5B4704AD5EEB41CA755" pageId="0" pageNumber="299" type="distribution">
<paragraph id="4E75FBC75F8AD5012B3928F975F6D3B0" pageId="0" pageNumber="299">Distribution.</paragraph>
<paragraph id="E175A09DB07D66BF789F35957A91077A" pageId="0" pageNumber="299">
<taxonomicName id="4ECC70B05E1747B98F0DF6AF48E00A81" class="Actinopterygii" family="Haemulidae" genus="Plectorhinchus" kingdom="Animalia" lsidName="Plectorhinchus pictus subsp. cinerascens" order="Perciformes" pageId="0" pageNumber="299" rank="subSpecies" species="pictus" subSpecies="cinerascens">
<emphasis id="9F4E11C992CB48F820525301CCBD5049" italics="true" pageId="0" pageNumber="299">Plectorhinchus pictus cinerascens</emphasis>
</taxonomicName>
, occurs from the Bay of Bengal to the Arabian/Persian Gulf (
<bibRefCitation author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. 2001</bibRefCitation>
<bibRefCitation id="282704526BBFC87A5119E963534795ED" author="Johnson, JW" journalOrPublisher="Memoirs of the Queensland Museum" pageId="0" pageNumber="299" pagination="657 - 676" refId="B19" refString="Johnson, JW, Randall, JE, Chenoweth, SF, 2001. Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review. Memoirs of the Queensland Museum 46 (2): 657 - 676" title="Diagramma melanacrum, new species of haemulid fish from Indonesia, Borneo and the Philippines with a generic review." volume="46" year="2001">Johnson et al. 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="299" type="conservation status">
<paragraph pageId="0" pageNumber="299">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="299">Not yet assessed, not listed in the IUCN Red List of Threatened Species (https://www.iucnredlist.org/species/123439745/123494892).</paragraph>
<subSubSection id="01F6CA7C84EDD3E6279C74105A20248B" pageId="0" pageNumber="299" type="conservation status">
<paragraph id="FC296BEA7FB0ABA7D58F84B0C5DD297F" pageId="0" pageNumber="299">Conservation status.</paragraph>
<paragraph id="0CB140979D1F0F636AD090B868FFDA68" pageId="0" pageNumber="299">Not yet assessed, not listed in the IUCN Red List of Threatened Species (https://www.iucnredlist.org/species/123439745/123494892).</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,141 +1,141 @@
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<mods:title>Review of the Bobekia - group (Braconidae, Alysiinae, Alysiini), with description of a new genus and a new subgenus</mods:title>
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<mods:title id="17BD832E066BAE36BEC30CC2DFE50C7E">Review of the Bobekia - group (Braconidae, Alysiinae, Alysiini), with description of a new genus and a new subgenus</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
<mods:name id="687E8AF3BF96DCA8132B27F7B68A8124" type="personal">
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<mods:namePart>Zhang, Ruo-Nan</mods:namePart>
<mods:affiliation>Shaanxi Key Laboratory for Animal Conservation / Key Laboratory of Resource Biology and Biotechnology in Western China, Ministry of Education, College of Life Sciences, Northwest University, Xi'an 710069, China</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6568-6731</mods:nameIdentifier>
<mods:namePart id="6E3E3F959674147AE06503CF2D9D15F1">Zhang, Ruo-Nan</mods:namePart>
<mods:affiliation id="127537BA939AF1EC5F241E0ACBF8EAD3">Shaanxi Key Laboratory for Animal Conservation / Key Laboratory of Resource Biology and Biotechnology in Western China, Ministry of Education, College of Life Sciences, Northwest University, Xi'an 710069, China</mods:affiliation>
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<mods:namePart>Achterberg, Cornelis van</mods:namePart>
<mods:affiliation>Shaanxi Key Laboratory for Animal Conservation / Key Laboratory of Resource Biology and Biotechnology in Western China, Ministry of Education, College of Life Sciences, Northwest University, Xi'an 710069, China &amp; State Key Laboratory of Rice Biology and Ministry of Agriculture Key Lab of Agricultural Entomology, Institute of Insect Sciences, Zhejiang University, Hangzhou 310058, China</mods:affiliation>
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<mods:namePart id="435D2A356D99704854D58D1DC08ED368">Achterberg, Cornelis van</mods:namePart>
<mods:affiliation id="4E8AC6B037AFC6370C51CFB920F0B4D7">Shaanxi Key Laboratory for Animal Conservation / Key Laboratory of Resource Biology and Biotechnology in Western China, Ministry of Education, College of Life Sciences, Northwest University, Xi'an 710069, China &amp; State Key Laboratory of Rice Biology and Ministry of Agriculture Key Lab of Agricultural Entomology, Institute of Insect Sciences, Zhejiang University, Hangzhou 310058, China</mods:affiliation>
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<mods:namePart>Tian, Xiao-Xia</mods:namePart>
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<mods:namePart id="5898F47B955D244D9D13BE552358ED4C">Tian, Xiao-Xia</mods:namePart>
<mods:affiliation id="9EF1978EF9ECC5B91AC8755FCDCCF4FB">Shaanxi Key Laboratory for Animal Conservation / Key Laboratory of Resource Biology and Biotechnology in Western China, Ministry of Education, College of Life Sciences, Northwest University, Xi'an 710069, China</mods:affiliation>
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<mods:namePart>Tan, Jiang-Li</mods:namePart>
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<treatment ID-GBIF-Taxon="163626740" LSID="urn:lsid:plazi:treatment:2DED895FC0BC51E38B14BC77BDFFE579" httpUri="http://treatment.plazi.org/id/2DED895FC0BC51E38B14BC77BDFFE579" lastPageNumber="25" pageId="0" pageNumber="25">
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<taxonomicName LSID="2DED895F-C0BC-51E3-8B14-BC77BDFFE579" authority="Granger, 1949" authorityName="Granger" authorityYear="1949" class="Insecta" family="Braconidae" genus="Hovalysia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hovalysia" order="Hymenoptera" pageId="0" pageNumber="25" phylum="Arthropoda" rank="genus">Hovalysia Granger, 1949</taxonomicName>
<figureCitation captionStart="Figures 5364" captionStartId="F7" captionText="Figures 53 - 64. Hovalysia seyrigi Granger, ♂, holotype 53 wings 54 hind leg 55 head, dorsal aspect 56 outer hind claw, lateral aspect 57 mandible, full view of first tooth 58 head, anterior aspect 59 first-third metasomal tergites, dorsal aspect 60 mandible, full view of third tooth (fourth tooth arrowed) 61 mesosoma, dorsal aspect 62 apical antennal segments 63 antenna 64 habitus, lateral aspect." figureDoi="10.3897/zookeys.926.47270.figures53-64" httpUri="https://binary.pensoft.net/fig/399413" pageId="0" pageNumber="25">Figs 53-64</figureCitation>
<treatment id="2DED895FC0BC51E38B14BC77BDFFE579" ID-GBIF-Taxon="163626740" LSID="urn:lsid:plazi:treatment:2DED895FC0BC51E38B14BC77BDFFE579" httpUri="http://treatment.plazi.org/id/2DED895FC0BC51E38B14BC77BDFFE579" lastPageNumber="25" pageId="0" pageNumber="25" scope_family="Braconidae">
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<paragraph id="0B46341291FE6A3912F52FCB3E1D485B" pageId="0" pageNumber="25">
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<figureCitation id="57F08E1691CDAB2EC9DC483BC59403E6" captionStart="Figures 5364" captionStartId="F7" captionText="Figures 53 - 64. Hovalysia seyrigi Granger, ♂, holotype 53 wings 54 hind leg 55 head, dorsal aspect 56 outer hind claw, lateral aspect 57 mandible, full view of first tooth 58 head, anterior aspect 59 first-third metasomal tergites, dorsal aspect 60 mandible, full view of third tooth (fourth tooth arrowed) 61 mesosoma, dorsal aspect 62 apical antennal segments 63 antenna 64 habitus, lateral aspect." figureDoi="10.3897/zookeys.926.47270.figures53-64" httpUri="https://binary.pensoft.net/fig/399413" pageId="0" pageNumber="25">Figs 53-64</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<paragraph id="979D9B666C5C18CA53EEC8EBF68C076D" pageId="0" pageNumber="25">
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Granger, 1949: 400;
<bibRefCitation author="Shenefelt, RD" journalOrPublisher="Alysiinae," pageId="0" pageNumber="25" pagination="937 - 1113" refId="B15" refString="Shenefelt, RD, 1974. Hymenopterorum Catalogus (nova editioBraconidae 7. Alysiinae, 7: 937 - 1113" title="Hymenopterorum Catalogus (nova editioBraconidae 7." volume="7" year="1974">Shenefelt 1974</bibRefCitation>
<bibRefCitation id="EAE245BAC1540236245A83E414ADBB86" author="Shenefelt, RD" journalOrPublisher="Alysiinae," pageId="0" pageNumber="25" pagination="937 - 1113" refId="B15" refString="Shenefelt, RD, 1974. Hymenopterorum Catalogus (nova editioBraconidae 7. Alysiinae, 7: 937 - 1113" title="Hymenopterorum Catalogus (nova editioBraconidae 7." volume="7" year="1974">Shenefelt 1974</bibRefCitation>
: 992;
<bibRefCitation author="Fischer, M" journalOrPublisher="Linzer biologische Beitraege" pageId="0" pageNumber="25" pagination="593 - 647" refId="B9" refString="Fischer, M, 1993. Neubeschreibungen und Wiederbeschreibungen von Kieferwespen (Hym., Braconidae, Alysiinae: Tribus Alysiini) der Alten Welt. Linzer biologische Beitraege 25 (2): 593 - 647" title="Neubeschreibungen und Wiederbeschreibungen von Kieferwespen (Hym., Braconidae, Alysiinae: Tribus Alysiini) der Alten Welt." volume="25" year="1993">Fischer 1993</bibRefCitation>
<bibRefCitation id="38CA9E22FEE331123B2D539A8B8E3D79" author="Fischer, M" journalOrPublisher="Linzer biologische Beitraege" pageId="0" pageNumber="25" pagination="593 - 647" refId="B9" refString="Fischer, M, 1993. Neubeschreibungen und Wiederbeschreibungen von Kieferwespen (Hym., Braconidae, Alysiinae: Tribus Alysiini) der Alten Welt. Linzer biologische Beitraege 25 (2): 593 - 647" title="Neubeschreibungen und Wiederbeschreibungen von Kieferwespen (Hym., Braconidae, Alysiinae: Tribus Alysiini) der Alten Welt." volume="25" year="1993">Fischer 1993</bibRefCitation>
: 609-611,
<bibRefCitation author="Fischer, M" journalOrPublisher="Linzer biologische Beitraege" pageId="0" pageNumber="25" pagination="5 - 56" refId="B10" refString="Fischer, M, 1999. Einiges ueber Kieferwespen (Hymenoptera, Braconidae, Alysiinae). Linzer biologische Beitraege 31 (1): 5 - 56" title="Einiges ueber Kieferwespen (Hymenoptera, Braconidae, Alysiinae)." volume="31" year="1999">1999</bibRefCitation>
<bibRefCitation id="4180C1D4D4E03CA8E2B1AA4D88A7E826" author="Fischer, M" journalOrPublisher="Linzer biologische Beitraege" pageId="0" pageNumber="25" pagination="5 - 56" refId="B10" refString="Fischer, M, 1999. Einiges ueber Kieferwespen (Hymenoptera, Braconidae, Alysiinae). Linzer biologische Beitraege 31 (1): 5 - 56" title="Einiges ueber Kieferwespen (Hymenoptera, Braconidae, Alysiinae)." volume="31" year="1999">1999</bibRefCitation>
: 9-10. Type species (by monotypy);
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<taxonomicName id="1A29A6E8180DF51C2F5292D0923FE8DF" authorityName="Granger" authorityYear="1949" class="Insecta" family="Braconidae" genus="Hovalysia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hovalysia seyrigi" order="Hymenoptera" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="seyrigi">Hovalysia seyrigi</taxonomicName>
Granger, 1949 [holotype (MNHN) examined].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="25" type="notes">
<paragraph pageId="0" pageNumber="25">Notes.</paragraph>
<paragraph pageId="0" pageNumber="25">
<subSubSection id="08AF058307145AF69A4278A2CFC74145" pageId="0" pageNumber="25" type="notes">
<paragraph id="890DEE80F7974F9AA10C66C1B21D9D6E" pageId="0" pageNumber="25">Notes.</paragraph>
<paragraph id="C4FE1C569AD92E29F850F0C483F843C0" pageId="0" pageNumber="25">
The main characters are the medial position of vein r at the pterostigma and the aberrant venation of the fore wing in males (Fig.
<figureCitation captionStart="Figures 5364" captionStartId="F7" captionText="Figures 53 - 64. Hovalysia seyrigi Granger, ♂, holotype 53 wings 54 hind leg 55 head, dorsal aspect 56 outer hind claw, lateral aspect 57 mandible, full view of first tooth 58 head, anterior aspect 59 first-third metasomal tergites, dorsal aspect 60 mandible, full view of third tooth (fourth tooth arrowed) 61 mesosoma, dorsal aspect 62 apical antennal segments 63 antenna 64 habitus, lateral aspect." figureDoi="10.3897/zookeys.926.47270.figures53-64" httpUri="https://binary.pensoft.net/fig/399413" pageId="0" pageNumber="25">53</figureCitation>
<figureCitation id="35B3C05A2031F72BFD108F4D6D77C2E4" captionStart="Figures 5364" captionStartId="F7" captionText="Figures 53 - 64. Hovalysia seyrigi Granger, ♂, holotype 53 wings 54 hind leg 55 head, dorsal aspect 56 outer hind claw, lateral aspect 57 mandible, full view of first tooth 58 head, anterior aspect 59 first-third metasomal tergites, dorsal aspect 60 mandible, full view of third tooth (fourth tooth arrowed) 61 mesosoma, dorsal aspect 62 apical antennal segments 63 antenna 64 habitus, lateral aspect." figureDoi="10.3897/zookeys.926.47270.figures53-64" httpUri="https://binary.pensoft.net/fig/399413" pageId="0" pageNumber="25">53</figureCitation>
), which is unknown in other genera. Only two Afrotropical species are known. For their identification, see the key by
<bibRefCitation author="Fischer, M" journalOrPublisher="Linzer biologische Beitraege" pageId="0" pageNumber="25" pagination="5 - 56" refId="B10" refString="Fischer, M, 1999. Einiges ueber Kieferwespen (Hymenoptera, Braconidae, Alysiinae). Linzer biologische Beitraege 31 (1): 5 - 56" title="Einiges ueber Kieferwespen (Hymenoptera, Braconidae, Alysiinae)." volume="31" year="1999">Fischer (1999)</bibRefCitation>
<bibRefCitation id="F4DCA67017BA0019E1D6B0376AC23B44" author="Fischer, M" journalOrPublisher="Linzer biologische Beitraege" pageId="0" pageNumber="25" pagination="5 - 56" refId="B10" refString="Fischer, M, 1999. Einiges ueber Kieferwespen (Hymenoptera, Braconidae, Alysiinae). Linzer biologische Beitraege 31 (1): 5 - 56" title="Einiges ueber Kieferwespen (Hymenoptera, Braconidae, Alysiinae)." volume="31" year="1999">Fischer (1999)</bibRefCitation>
. So far, no males are known in China with similar venation, but Chinese male
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<emphasis italics="true" pageId="0" pageNumber="25">Separatatus</emphasis>
<taxonomicName id="3F07DB606EC014A6A2DA4B46EA9FBC81" authorityName="Chen &amp; Wu" authorityYear="1994" class="Insecta" family="Braconidae" genus="Separatatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Separatatus" order="Hymenoptera" pageId="0" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="0697D9CA33F5802008E120DA09684A05" italics="true" pageId="0" pageNumber="25">Separatatus</emphasis>
</taxonomicName>
with aberrant venation (Figs
<figureCitation captionStart="Figures 1719" captionStartId="F3" captionText="Figures 17 - 19. Separatatus yinshani Zhang &amp; van Achterberg sp. nov., ♂, paratype 17 habitus, lateral aspect 18 head, anterior aspect 19 genitalia, ventral aspect." figureDoi="10.3897/zookeys.926.47270.figures17-19" httpUri="https://binary.pensoft.net/fig/399409" pageId="0" pageNumber="25">17</figureCitation>
<figureCitation id="ACF37FFAEF7952601CEF7D20A6D0AD72" captionStart="Figures 1719" captionStartId="F3" captionText="Figures 17 - 19. Separatatus yinshani Zhang &amp; van Achterberg sp. nov., ♂, paratype 17 habitus, lateral aspect 18 head, anterior aspect 19 genitalia, ventral aspect." figureDoi="10.3897/zookeys.926.47270.figures17-19" httpUri="https://binary.pensoft.net/fig/399409" pageId="0" pageNumber="25">17</figureCitation>
,
<figureCitation captionStart="Figures 2025" captionStartId="F4" captionText="Figures 20 - 25. Separatatus yinshani Zhang &amp; van Achterberg sp. nov., ♂, paratype 20 wings 21 propodeum, first-third metasomal tergites, dorsal aspect 22 basal antennal segments 23 apical antennal segments 24 head, anterior aspect 25 head, lateral aspect." figureDoi="10.3897/zookeys.926.47270.figures20-25" httpUri="https://binary.pensoft.net/fig/399410" pageId="0" pageNumber="25">20</figureCitation>
<figureCitation id="938717721F595280ECC8C7F00804E611" captionStart="Figures 2025" captionStartId="F4" captionText="Figures 20 - 25. Separatatus yinshani Zhang &amp; van Achterberg sp. nov., ♂, paratype 20 wings 21 propodeum, first-third metasomal tergites, dorsal aspect 22 basal antennal segments 23 apical antennal segments 24 head, anterior aspect 25 head, lateral aspect." figureDoi="10.3897/zookeys.926.47270.figures20-25" httpUri="https://binary.pensoft.net/fig/399410" pageId="0" pageNumber="25">20</figureCitation>
) are known. Therefore,
<bibRefCitation DOI="https://doi.org/10.1071/IT01012" author="Wharton, RA" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="25" pagination="7 - 105" refId="B23" refString="Wharton, RA, 2002. Revision of the Australian Alysiini (Hymenoptera: Braconidae). Invertebrate Systematics 16 (1): 7 - 105, DOI: https://doi.org/10.1071/IT01012" title="Revision of the Australian Alysiini (Hymenoptera: Braconidae)." url="https://doi.org/10.1071/IT01012" volume="16" year="2002">
<normalizedToken originalValue="Whartons">Wharton's</normalizedToken>
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<normalizedToken id="22D29D4616F464EDC7639ED6D53F139C" originalValue="Whartons">Wharton's</normalizedToken>
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reference to Taiwanese specimens with widened veins probably refers to a species of
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<emphasis italics="true" pageId="0" pageNumber="25">Separatatus</emphasis>
<taxonomicName id="B64246781770C878CD10284DB80B635F" authorityName="Chen &amp; Wu" authorityYear="1994" class="Insecta" family="Braconidae" genus="Separatatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Separatatus" order="Hymenoptera" pageId="0" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="E5064A34D2F62DA9F3AB4248646E8647" italics="true" pageId="0" pageNumber="25">Separatatus</emphasis>
</taxonomicName>
. The biology is unknown.
</paragraph>
<caption doi="10.3897/zookeys.926.47270.figures53-64" httpUri="https://binary.pensoft.net/fig/399413" pageId="0" pageNumber="25" start="Figures 5364" startId="F7">
<paragraph pageId="0" pageNumber="25">
<emphasis bold="true" pageId="0" pageNumber="25">Figures 53-64.</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="25">Hovalysia seyrigi</emphasis>
<caption id="DD404AA157CD5B1F1E6DA89D80175976" doi="10.3897/zookeys.926.47270.figures53-64" httpUri="https://binary.pensoft.net/fig/399413" pageId="0" pageNumber="25" start="Figures 5364" startId="F7">
<paragraph id="2F4329DCDF5BF19986C6CB400DBD68EB" pageId="0" pageNumber="25">
<emphasis id="CD5F6B7901ED093556C1C5679B42EF44" bold="true" pageId="0" pageNumber="25">Figures 53-64.</emphasis>
<taxonomicName id="391B24B7F1162EE802B846BC9E870AE3" authorityName="Granger" authorityYear="1949" class="Insecta" family="Braconidae" genus="Hovalysia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hovalysia seyrigi" order="Hymenoptera" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="seyrigi">
<emphasis id="07FFC1E48ABA3295EC065547F6843086" italics="true" pageId="0" pageNumber="25">Hovalysia seyrigi</emphasis>
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Granger, ♂, holotype
<emphasis bold="true" pageId="0" pageNumber="25">53</emphasis>
<emphasis id="CCE2A2C915B0388D7169A8621134321E" bold="true" pageId="0" pageNumber="25">53</emphasis>
wings
<emphasis bold="true" pageId="0" pageNumber="25">54</emphasis>
<emphasis id="32B81DFCD87BA22D9F09747197AA620F" bold="true" pageId="0" pageNumber="25">54</emphasis>
hind leg
<emphasis bold="true" pageId="0" pageNumber="25">55</emphasis>
<emphasis id="5E74411C623FD312F1468B9275157230" bold="true" pageId="0" pageNumber="25">55</emphasis>
head, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="25">56</emphasis>
<emphasis id="7005A5A470BDF070B5145FAAB8598CAE" bold="true" pageId="0" pageNumber="25">56</emphasis>
outer hind claw, lateral aspect
<emphasis bold="true" pageId="0" pageNumber="25">57</emphasis>
<emphasis id="2D1FF035E280D5F8687D9F610FEC8BD8" bold="true" pageId="0" pageNumber="25">57</emphasis>
mandible, full view of first tooth
<emphasis bold="true" pageId="0" pageNumber="25">58</emphasis>
<emphasis id="86CDE08B4A63940035C442390E3F3535" bold="true" pageId="0" pageNumber="25">58</emphasis>
head, anterior aspect
<emphasis bold="true" pageId="0" pageNumber="25">59</emphasis>
<emphasis id="5E2C90E95E0C2DECC54ADF734A473CCD" bold="true" pageId="0" pageNumber="25">59</emphasis>
first-third metasomal tergites, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="25">60</emphasis>
<emphasis id="07085BFE83CC2974ECEDFC5E35C2DFB8" bold="true" pageId="0" pageNumber="25">60</emphasis>
mandible, full view of third tooth (fourth tooth arrowed)
<emphasis bold="true" pageId="0" pageNumber="25">61</emphasis>
<emphasis id="79200573209D468419E437E46302E073" bold="true" pageId="0" pageNumber="25">61</emphasis>
mesosoma, dorsal aspect
<emphasis bold="true" pageId="0" pageNumber="25">62</emphasis>
<emphasis id="9B484EA5EB2126C7305F5A7E9C56D296" bold="true" pageId="0" pageNumber="25">62</emphasis>
apical antennal segments
<emphasis bold="true" pageId="0" pageNumber="25">63</emphasis>
<emphasis id="95C9163E74E1C32602695AD89F5FEED9" bold="true" pageId="0" pageNumber="25">63</emphasis>
antenna
<emphasis bold="true" pageId="0" pageNumber="25">64</emphasis>
<emphasis id="A3957EB04F404CBF6DC958C586DF1B93" bold="true" pageId="0" pageNumber="25">64</emphasis>
habitus, lateral aspect.
</paragraph>
</caption>