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<mods:title>Two new species of the millipede genus Tylopus Jeekel, 1968 from Shan State, Myanmar (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:title id="BC821D289E687018853B43069FCFD612">Two new species of the millipede genus Tylopus Jeekel, 1968 from Shan State, Myanmar (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation>Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai, 50290, Thailand & Biodiversity and Utilization Research Center of Maejo University, Maejo University, Chiang Mai, 50290, Thailand</mods:affiliation>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:namePart>Srisonchai, Ruttapon</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, 40002, Thailand</mods:affiliation>
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<mods:namePart id="EDD49E630DEADAF26F94803B0035116B">Srisonchai, Ruttapon</mods:namePart>
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<mods:affiliation id="FC6C1043BB73E20173A67F1F19E83768">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, 40002, Thailand</mods:affiliation>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<taxonomicName LSID="http://zoobank.org/FD497FBD-67B7-4171-9905-ED6D2011DF51" authority="Likhitrakarn & Golovatch & Srisonchai & Panha, 2021" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola" status="sp. nov.">Tylopus monticola</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="167">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">Figs 1A</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tylopus monticola sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, lateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure3" httpUri="https://binary.pensoft.net/fig/549196" pageId="0" pageNumber="167">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and stout process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167">, 4</figureCitation>
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<treatment id="21DBC0B8DDB250D3AC18055CCDC0A131" ID-GBIF-Taxon="182408862" LSID="urn:lsid:zoobank.org:act:FD497FBD-67B7-4171-9905-ED6D2011DF51" httpUri="http://treatment.plazi.org/id/21DBC0B8DDB250D3AC18055CCDC0A131" lastPageNumber="167" pageId="0" pageNumber="167" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida" scope_subFamily="Paradoxosomatinae" scope_tribe="Sulciferini">
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<taxonomicName id="308FAC236764A45B3137AF387008EDA5" ID-CoL="8KR6X" LSID="http://zoobank.org/FD497FBD-67B7-4171-9905-ED6D2011DF51" authority="Likhitrakarn & Golovatch & Srisonchai & Panha, 2021" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola" status="sp. nov.">Tylopus monticola</taxonomicName>
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<taxonomicNameLabel id="A31E468BC159D9C553233C8695C996B7" pageId="0" pageNumber="167">sp. nov.</taxonomicNameLabel>
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<figureCitation id="1453F2B505C99EF2AE372871888ED668" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">Figs 1A</figureCitation>
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<figureCitation id="226D728C35CC183B3F1A7A197D41AC12" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">, 2</figureCitation>
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<figureCitation id="C3181572EDC6E2329E1DE99F91E8775B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tylopus monticola sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, lateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure3" httpUri="https://binary.pensoft.net/fig/549196" pageId="0" pageNumber="167">, 3</figureCitation>
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<figureCitation id="E5E8A9279549606CCB8F711E99CDDF7C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and stout process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167">, 4</figureCitation>
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</paragraph>
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<subSubSection pageId="0" pageNumber="167" type="materials_examined">
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<paragraph pageId="0" pageNumber="167">Material examined.</paragraph>
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||||
<paragraph pageId="0" pageNumber="167">
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<materialsCitation collectingDate="2016-09-21" collectorName="J. Sutcharit, R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Montawa Cave" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="♂" stateProvince="Shan State" typeStatus="Holotype">
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<typeStatus>Holotype</typeStatus>
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<paragraph id="9786D1BFD0986E47B3033825516DC850" pageId="0" pageNumber="167">Material examined.</paragraph>
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<materialsCitation id="DF7431D61593A72A9E1F086EACAA8ED6" collectingDate="2016-09-21" collectorName="J. Sutcharit, R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Montawa Cave" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="♂" stateProvince="Shan State" typeStatus="Holotype">
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<typeStatus id="1C3DC8EBFBC8A9D210FB1595C5266B56">Holotype</typeStatus>
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:
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<collectingCountry name="Myanmar">Myanmar</collectingCountry>
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<collectingCountry id="8196459B11CE99825F9342EFCCBD7E2D" name="Myanmar">Myanmar</collectingCountry>
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-
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<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||||
<emphasis id="402E31AFA64D4AB40BF7E28092F643FE" bold="true" pageId="0" pageNumber="167">
|
||||
<collectingRegion id="63D55196CDB689CD5329F9D7B48EB318" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||||
</emphasis>
|
||||
•
|
||||
<specimenCount>♂</specimenCount>
|
||||
<specimenCount id="1B8BC784F6C11B129E4C9C7EE7C7B562">♂</specimenCount>
|
||||
;
|
||||
<collectingMunicipality>Taunggyi District</collectingMunicipality>
|
||||
<collectingMunicipality id="8841462A7CACB734F22818BF81B16F44">Taunggyi District</collectingMunicipality>
|
||||
, near
|
||||
<location LSID="urn:lsid:plazi:treatment:21DBC0B8DDB250D3AC18055CCDC0A131:4DE8D71E7C578BDD24BCCC52EF630F38" country="Myanmar" latitude="20.754417" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" name="Montawa Cave" stateProvince="Shan State">Montawa Cave</location>
|
||||
<location id="4DE8D71E7C578BDD24BCCC52EF630F38" LSID="urn:lsid:plazi:treatment:21DBC0B8DDB250D3AC18055CCDC0A131:4DE8D71E7C578BDD24BCCC52EF630F38" country="Myanmar" latitude="20.754417" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" name="Montawa Cave" stateProvince="Shan State">Montawa Cave</location>
|
||||
; elev.
|
||||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.204" unit="m" value="1204.0">
|
||||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.204" unit="m" value="1204.0">1204 m</elevation>
|
||||
<quantity id="D03622F222A4161B0DC3ADD315016CF8" metricMagnitude="3" metricUnit="m" metricValue="1.204" unit="m" value="1204.0">
|
||||
<elevation id="B833868F2F4ADD731F5BF07BECDCB7D7" metricMagnitude="3" metricUnit="m" metricValue="1.204" unit="m" value="1204.0">1204 m</elevation>
|
||||
</quantity>
|
||||
;
|
||||
<geoCoordinate degrees="20" direction="north" minutes="45" orientation="latitude" precision="1" seconds="15.9" value="20.754417">20°45'15.9"N</geoCoordinate>
|
||||
<geoCoordinate id="A6FE7493F04ECD01678983089AB70D18" degrees="20" direction="north" minutes="45" orientation="latitude" precision="1" seconds="15.9" value="20.754417">20°45'15.9"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="97" direction="east" minutes="01" orientation="longitude" precision="1" seconds="03.4" value="97.01762">97°01'03.4"E</geoCoordinate>
|
||||
<geoCoordinate id="D66363FFFC538A67A101EBC829BF76C1" degrees="97" direction="east" minutes="01" orientation="longitude" precision="1" seconds="03.4" value="97.01762">97°01'03.4"E</geoCoordinate>
|
||||
;
|
||||
<collectingDate value="2016-09-21">21 Sep. 2016</collectingDate>
|
||||
<collectingDate id="DF4E77C754240A742665875309662A23" value="2016-09-21">21 Sep. 2016</collectingDate>
|
||||
;
|
||||
<collectorName>J. Sutcharit</collectorName>
|
||||
<collectorName id="8617AC452B514834570AD3F40E6BF01D">J. Sutcharit</collectorName>
|
||||
,
|
||||
<collectorName>R. Srisonchai</collectorName>
|
||||
<collectorName id="6C0AB6CDDA725AF5519A9E63EB000F38">R. Srisonchai</collectorName>
|
||||
leg.; CUMZ
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Sutcharit & R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Shan State" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="3" specimenCount-female="3" stateProvince="Shan State" typeStatus="Paratypes">
|
||||
<emphasis bold="true" italics="true" pageId="0" pageNumber="167">
|
||||
<typeStatus>Paratypes</typeStatus>
|
||||
<materialsCitation id="C5A6C79C4ECB433746F2665B62EF8539" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Sutcharit & R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Shan State" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="3" specimenCount-female="3" stateProvince="Shan State" typeStatus="Paratypes">
|
||||
<emphasis id="F0C89E7EDB18FB6D572E7F2B9033AB4B" bold="true" italics="true" pageId="0" pageNumber="167">
|
||||
<typeStatus id="EF853B5D783D0AFD94B75470C6EDC9C6">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
:
|
||||
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
|
||||
<collectingCountry id="FC0810DADD27E3749B172F73A02CB204" name="Myanmar">Myanmar</collectingCountry>
|
||||
-
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">
|
||||
<collectingRegion country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||||
<emphasis id="C51AA21F9503ED90AEAFED26B2E22CA0" bold="true" pageId="0" pageNumber="167">
|
||||
<collectingRegion id="B4D7B56CCB46C1579BB9529774087B02" country="Myanmar" name="Shan">Shan State</collectingRegion>
|
||||
</emphasis>
|
||||
•
|
||||
<specimenCount type="female">3 ♀♀</specimenCount>
|
||||
<specimenCount id="F2F14591EBD8DFF31DF10E027744A477" type="female">3 ♀♀</specimenCount>
|
||||
; same collection data as holotype; CUMZ
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Sutcharit & R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Montawa Cave" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="1" specimenCount-female="1" stateProvince="Shan State" typeStatus="paratype">
|
||||
<specimenCount type="female">1 ♀</specimenCount>
|
||||
<materialsCitation id="F5CAF6A29F9DB18CBD55671ACF31EA18" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Sutcharit & R. Srisonchai" country="Myanmar" elevation="1204" latitude="20.754417" location="Montawa Cave" longLatPrecision="1" longitude="97.01762" municipality="Taunggyi District" specimenCount="1" specimenCount-female="1" stateProvince="Shan State" typeStatus="paratype">
|
||||
<specimenCount id="41D5C1C154AEAE051BB5C3A4BA1C84B6" type="female">1 ♀</specimenCount>
|
||||
; same collection data as holotype; ZMUM
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation collectingDate="2016-09-21" collectorName="J. Sutcharit, R. Srisonchai" country="Myanmar" elevation="1583" latitude="20.791527" location="Aye Say Tee Cave" longLatPrecision="1" longitude="97.050446" specimenCount="3" specimenCount-female="3" stateProvince="Shan State" typeStatus="paratype">
|
||||
<specimenCount type="female">3 ♀♀</specimenCount>
|
||||
<materialsCitation id="412B3B9680E59E28E09BCB16132BEC8F" collectingDate="2016-09-21" collectorName="J. Sutcharit, R. Srisonchai" country="Myanmar" elevation="1583" latitude="20.791527" location="Aye Say Tee Cave" longLatPrecision="1" longitude="97.050446" specimenCount="3" specimenCount-female="3" stateProvince="Shan State" typeStatus="paratype">
|
||||
<specimenCount id="4AE2B7A6C1637BF3FEF10E05A4D7FE7A" type="female">3 ♀♀</specimenCount>
|
||||
; near
|
||||
<location LSID="urn:lsid:plazi:treatment:21DBC0B8DDB250D3AC18055CCDC0A131:04E6D43684EC6FFF2CE39AB64DBDB9B9" country="Myanmar" latitude="20.791527" longLatPrecision="1" longitude="97.050446" name="Aye Say Tee Cave" stateProvince="Shan State">Aye Say Tee Cave</location>
|
||||
<location id="04E6D43684EC6FFF2CE39AB64DBDB9B9" LSID="urn:lsid:plazi:treatment:21DBC0B8DDB250D3AC18055CCDC0A131:04E6D43684EC6FFF2CE39AB64DBDB9B9" country="Myanmar" latitude="20.791527" longLatPrecision="1" longitude="97.050446" name="Aye Say Tee Cave" stateProvince="Shan State">Aye Say Tee Cave</location>
|
||||
; elev.
|
||||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.583" unit="m" value="1583.0">
|
||||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.583" unit="m" value="1583.0">1583 m</elevation>
|
||||
<quantity id="42ED45BE6D0A1A8173E406F3BA094007" metricMagnitude="3" metricUnit="m" metricValue="1.583" unit="m" value="1583.0">
|
||||
<elevation id="AAADAD6C694E3627FFBE104FB78A3596" metricMagnitude="3" metricUnit="m" metricValue="1.583" unit="m" value="1583.0">1583 m</elevation>
|
||||
</quantity>
|
||||
;
|
||||
<geoCoordinate degrees="20" direction="north" minutes="47" orientation="latitude" precision="1" seconds="29.5" value="20.791527">20°47'29.5"N</geoCoordinate>
|
||||
<geoCoordinate id="D5A8CCDA60E2844019052ADCBDF502CF" degrees="20" direction="north" minutes="47" orientation="latitude" precision="1" seconds="29.5" value="20.791527">20°47'29.5"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="97" direction="east" minutes="03" orientation="longitude" precision="1" seconds="01.6" value="97.050446">97°03'01.6"E</geoCoordinate>
|
||||
<geoCoordinate id="F9C0488F03BB72579AA80928348C26AF" degrees="97" direction="east" minutes="03" orientation="longitude" precision="1" seconds="01.6" value="97.050446">97°03'01.6"E</geoCoordinate>
|
||||
;
|
||||
<collectingDate value="2016-09-21">21 Sep. 2016</collectingDate>
|
||||
<collectingDate id="F6A20A9690C87594D54F5ABF56D72F5E" value="2016-09-21">21 Sep. 2016</collectingDate>
|
||||
;
|
||||
<collectorName>J. Sutcharit</collectorName>
|
||||
<collectorName id="69D1B32FCE9D0BC3B2596AEA15AE1E3E">J. Sutcharit</collectorName>
|
||||
,
|
||||
<collectorName>R. Srisonchai</collectorName>
|
||||
<collectorName id="652262ACCBF5F727D1A6A651005A57BF">R. Srisonchai</collectorName>
|
||||
leg.; CUMZ
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation collectingDate="2016-09-23" collectorName="J. Sutcharit, R. Srisonchai" country="Myanmar" elevation="1159" latitude="20.251028" location="Parpant area" longLatPrecision="1" longitude="97.23997" specimenCount="1" specimenCount-male="1" stateProvince="Shan State" typeStatus="paratype">
|
||||
<specimenCount type="male">1 ♂</specimenCount>
|
||||
<materialsCitation id="5FD03AA7B33CE29E590F749DA33A5172" collectingDate="2016-09-23" collectorName="J. Sutcharit, R. Srisonchai" country="Myanmar" elevation="1159" latitude="20.251028" location="Parpant area" longLatPrecision="1" longitude="97.23997" specimenCount="1" specimenCount-male="1" stateProvince="Shan State" typeStatus="paratype">
|
||||
<specimenCount id="A3E3641590EBEE1DE9580A351F6F99EB" type="male">1 ♂</specimenCount>
|
||||
;
|
||||
<location LSID="urn:lsid:plazi:treatment:21DBC0B8DDB250D3AC18055CCDC0A131:35B10F0159ABBDE04FDD17035F8E6D04" country="Myanmar" latitude="20.251028" longLatPrecision="1" longitude="97.23997" name="Parpant area" stateProvince="Shan State">Parpant area</location>
|
||||
<location id="35B10F0159ABBDE04FDD17035F8E6D04" LSID="urn:lsid:plazi:treatment:21DBC0B8DDB250D3AC18055CCDC0A131:35B10F0159ABBDE04FDD17035F8E6D04" country="Myanmar" latitude="20.251028" longLatPrecision="1" longitude="97.23997" name="Parpant area" stateProvince="Shan State">Parpant area</location>
|
||||
, outside the cave; elev.
|
||||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.159" unit="m" value="1159.0">
|
||||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.159" unit="m" value="1159.0">1159 m</elevation>
|
||||
<quantity id="86EFAD4B5DC3A9DE1C22F328A88676D6" metricMagnitude="3" metricUnit="m" metricValue="1.159" unit="m" value="1159.0">
|
||||
<elevation id="132AB791FB4F3FA2BC4AA16F74B11E2C" metricMagnitude="3" metricUnit="m" metricValue="1.159" unit="m" value="1159.0">1159 m</elevation>
|
||||
</quantity>
|
||||
;
|
||||
<geoCoordinate degrees="20" direction="north" minutes="15" orientation="latitude" precision="1" seconds="03.7" value="20.251028">20°15'03.7"N</geoCoordinate>
|
||||
<geoCoordinate id="BFDD7B06484A80B1C08F1340598FA455" degrees="20" direction="north" minutes="15" orientation="latitude" precision="1" seconds="03.7" value="20.251028">20°15'03.7"N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate degrees="97" direction="east" minutes="14" orientation="longitude" precision="1" seconds="23.9" value="97.23997">97°14'23.9"E</geoCoordinate>
|
||||
<geoCoordinate id="636418F7EABB69F451BC41435E0D7817" degrees="97" direction="east" minutes="14" orientation="longitude" precision="1" seconds="23.9" value="97.23997">97°14'23.9"E</geoCoordinate>
|
||||
;
|
||||
<collectingDate value="2016-09-23">23 Sep. 2016</collectingDate>
|
||||
<collectingDate id="EEF302D8D729DDD07F38BA33E13E315C" value="2016-09-23">23 Sep. 2016</collectingDate>
|
||||
;
|
||||
<collectorName>J. Sutcharit</collectorName>
|
||||
<collectorName id="A912C40C3BCC46E6906757D1FDFD0E93">J. Sutcharit</collectorName>
|
||||
,
|
||||
<collectorName>R. Srisonchai</collectorName>
|
||||
<collectorName id="3070F7BB1F502FF77B5FB03E9B3D945E">R. Srisonchai</collectorName>
|
||||
leg.; CUMZ
|
||||
</materialsCitation>
|
||||
•
|
||||
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Sutcharit & R. Srisonchai" country="Myanmar" elevation="1159" latitude="20.251028" location="Parpant area" longLatPrecision="1" longitude="97.23997" specimenCount="1" specimenCount-male="1" stateProvince="Shan State" typeStatus="paratype">
|
||||
<specimenCount type="male">1 ♂</specimenCount>
|
||||
<materialsCitation id="23ADC5EAF6A4B4C12FA8EB2F7FCE0F72" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Sutcharit & R. Srisonchai" country="Myanmar" elevation="1159" latitude="20.251028" location="Parpant area" longLatPrecision="1" longitude="97.23997" specimenCount="1" specimenCount-male="1" stateProvince="Shan State" typeStatus="paratype">
|
||||
<specimenCount id="5C45739B1D7E71D726AD54BD5D8D7C13" type="male">1 ♂</specimenCount>
|
||||
; same collection data as previous; ZMUM
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="167" type="diagnosis">
|
||||
<paragraph pageId="0" pageNumber="167">Diagnosis.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<subSubSection id="90E629B14D6FC47572666B8C7809908F" pageId="0" pageNumber="167" type="diagnosis">
|
||||
<paragraph id="DADCF19FA82B75D99DF8EFC63CB0C52C" pageId="0" pageNumber="167">Diagnosis.</paragraph>
|
||||
<paragraph id="D3058D9B06F873A298210FF8DA8249C3" pageId="0" pageNumber="167">
|
||||
Using the latest key to
|
||||
<taxonomicName authorityName="Jeekel" authorityYear="1968" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus</emphasis>
|
||||
<taxonomicName id="CA4754FB3C35DBCD794F246EC24C764F" authorityName="Jeekel" authorityYear="1968" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="62A43F8DF64576AFAA6351348872D34A" italics="true" pageId="0" pageNumber="167">Tylopus</emphasis>
|
||||
</taxonomicName>
|
||||
species (
|
||||
<bibRefCitation DOI="https://doi.org/10.5852/ejt.2016.195" author="Likhitrakarn, N" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="167" pagination="1 - 47" refId="B23" refString="Likhitrakarn, N, Golovatch, SI, Panha, S, 2016. The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina. European Journal of Taxonomy 195: 1 - 47, DOI: https://doi.org/10.5852/ejt.2016.195" title="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina." url="https://doi.org/10.5852/ejt.2016.195" volume="195" year="2016">Likhitrakarn et al. 2016</bibRefCitation>
|
||||
<bibRefCitation id="6D472EEF5D6E30BB8A56E6F244177397" DOI="https://doi.org/10.5852/ejt.2016.195" author="Likhitrakarn, N" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="167" pagination="1 - 47" refId="B23" refString="Likhitrakarn, N, Golovatch, SI, Panha, S, 2016. The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina. European Journal of Taxonomy 195: 1 - 47, DOI: https://doi.org/10.5852/ejt.2016.195" title="The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina." url="https://doi.org/10.5852/ejt.2016.195" volume="195" year="2016">Likhitrakarn et al. 2016</bibRefCitation>
|
||||
), as well as the information concerning all 12 congeners described since (
|
||||
<bibRefCitation author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="335 - 354" publicationUrl="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" refId="B16" refString="Golovatch, SI, VandenSpiegel, D, Semenyuk, II, 2016. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXI. Arthropoda Selecta 25 (4): 335 - 354, https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXI." url="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" volume="25" year="2016">Golovatch et al. 2016</bibRefCitation>
|
||||
<bibRefCitation id="6571ECC8837E975A57CCC76A64DFFAA5" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="335 - 354" publicationUrl="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" refId="B16" refString="Golovatch, SI, VandenSpiegel, D, Semenyuk, II, 2016. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXI. Arthropoda Selecta 25 (4): 335 - 354, https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXI." url="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/25/25_4_335_354_Golovatch_et_al_for_Inet.pdf" volume="25" year="2016">Golovatch et al. 2016</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="261 - 277" publicationUrl="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_4_261_277_Golovatch.pdf" refId="B10" refString="Golovatch, SI, 2018. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXV. Arthropoda Selecta 27 (4): 261 - 277, https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_4_261_277_Golovatch.pdf" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXV." url="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_4_261_277_Golovatch.pdf" volume="27" year="2018">Golovatch 2018</bibRefCitation>
|
||||
<bibRefCitation id="2B7BE1B466D5CF51F5B5D11DD328D6BC" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="261 - 277" publicationUrl="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_4_261_277_Golovatch.pdf" refId="B10" refString="Golovatch, SI, 2018. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXV. Arthropoda Selecta 27 (4): 261 - 277, https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_4_261_277_Golovatch.pdf" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXV." url="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_4_261_277_Golovatch.pdf" volume="27" year="2018">Golovatch 2018</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.28.4.01" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="459 - 478" refId="B11" refString="Golovatch, SI, 2019. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVII. Arthropoda Selecta 28 (4): 459 - 478, DOI: https://doi.org/10.15298/arthsel.28.4.01" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVII." url="https://doi.org/10.15298/arthsel.28.4.01" volume="28" year="2019">2019</bibRefCitation>
|
||||
<bibRefCitation id="F2A6AB43D54061B3E4A16D1EEBE1BD56" DOI="https://doi.org/10.15298/arthsel.28.4.01" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="459 - 478" refId="B11" refString="Golovatch, SI, 2019. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVII. Arthropoda Selecta 28 (4): 459 - 478, DOI: https://doi.org/10.15298/arthsel.28.4.01" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVII." url="https://doi.org/10.15298/arthsel.28.4.01" volume="28" year="2019">2019</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation DOI="https://doi.org/10.15298/arthsel.29.2.01" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="161 - 172" refId="B12" refString="Golovatch, SI, 2020. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVIII. Arthropoda Selecta 29 (2): 161 - 172, DOI: https://doi.org/10.15298/arthsel.29.2.01" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVIII." url="https://doi.org/10.15298/arthsel.29.2.01" volume="29" year="2020">2020</bibRefCitation>
|
||||
<bibRefCitation id="D34FA144310BF8D2918507D8512596BE" DOI="https://doi.org/10.15298/arthsel.29.2.01" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="161 - 172" refId="B12" refString="Golovatch, SI, 2020. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVIII. Arthropoda Selecta 29 (2): 161 - 172, DOI: https://doi.org/10.15298/arthsel.29.2.01" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XXVIII." url="https://doi.org/10.15298/arthsel.29.2.01" volume="29" year="2020">2020</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="187 - 200" publicationUrl="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_3_187_200_Golovatch_Semenyuk_for_Inet.pdf" refId="B15" refString="Golovatch, SI, Semenyuk, II, 2018. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XX. Arthropoda Selecta 27 (3): 187 - 200, https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_3_187_200_Golovatch_Semenyuk_for_Inet.pdf" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XX." url="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_3_187_200_Golovatch_Semenyuk_for_Inet.pdf" volume="27" year="2018">Golovatch and Semenyuk 2018</bibRefCitation>
|
||||
<bibRefCitation id="77BCD339BEAC9B7B3829BE21960EC7B0" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="167" pagination="187 - 200" publicationUrl="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_3_187_200_Golovatch_Semenyuk_for_Inet.pdf" refId="B15" refString="Golovatch, SI, Semenyuk, II, 2018. On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XX. Arthropoda Selecta 27 (3): 187 - 200, https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_3_187_200_Golovatch_Semenyuk_for_Inet.pdf" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XX." url="https://kmkjournals.com/upload/PDF/ArthropodaSelecta/27/27_3_187_200_Golovatch_Semenyuk_for_Inet.pdf" volume="27" year="2018">Golovatch and Semenyuk 2018</bibRefCitation>
|
||||
),
|
||||
<taxonomicName lsidName="T. monticola" pageId="0" pageNumber="167" rank="species" species="monticola">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">T. monticola</emphasis>
|
||||
<taxonomicName id="4AD033C171B2B21050B09988A18FCBF9" lsidName="T. monticola" pageId="0" pageNumber="167" rank="species" species="monticola">
|
||||
<emphasis id="6E618B4A1E59259D896F71AD02FC515B" italics="true" pageId="0" pageNumber="167">T. monticola</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov. keys out to
|
||||
<taxonomicName lsidName="T. rugosus" pageId="0" pageNumber="167" rank="species" species="rugosus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">T. rugosus</emphasis>
|
||||
<taxonomicName id="2E15A7B95F45FA44DCF0B8475F9B5AA5" lsidName="T. rugosus" pageId="0" pageNumber="167" rank="species" species="rugosus">
|
||||
<emphasis id="790AFAD63CBFFA6F1A31FCD98E75AF2F" italics="true" pageId="0" pageNumber="167">T. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
Golovatch & Enghoff, 1993 on account of the particularly strong similarities in the gonopodal structure (Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gonopod structure of Tylopus rugosus Golovatch & Enghoff, 1993, ♂ holotype, left gonopod A-C mesal, lateral and dorsal views, respectively. Abbreviations: h strong hook-shaped process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bar: 0.5 mm (after Golovatch and Enghoff 1993)." figureDoi="10.3897/zookeys.1040.66209.figure8" httpUri="https://binary.pensoft.net/fig/549201" pageId="0" pageNumber="167">8</figureCitation>
|
||||
<figureCitation id="5BFB5532020D4B4F31EB75BEF2B61008" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gonopod structure of Tylopus rugosus Golovatch & Enghoff, 1993, ♂ holotype, left gonopod A-C mesal, lateral and dorsal views, respectively. Abbreviations: h strong hook-shaped process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bar: 0.5 mm (after Golovatch and Enghoff 1993)." figureDoi="10.3897/zookeys.1040.66209.figure8" httpUri="https://binary.pensoft.net/fig/549201" pageId="0" pageNumber="167">8</figureCitation>
|
||||
). Thus, even though they both share most of the somatic and gonopodal features, the new species differs in the large and long process
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">z</emphasis>
|
||||
<emphasis id="2810C6118BF8B29B9A0FA83AA4802BA4" bold="true" pageId="0" pageNumber="167">z</emphasis>
|
||||
with a serrate edge along the dorsal margin, which protrudes beyond the apicolateral lobe (
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
<emphasis id="2F4C916EFEA8E17509461AC2D1BAECAE" bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
) (Figs
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tylopus monticola sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, lateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure3" httpUri="https://binary.pensoft.net/fig/549196" pageId="0" pageNumber="167">3B-D</figureCitation>
|
||||
<figureCitation id="9FFB381A98EC4CFB7FC9DF52BF946325" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tylopus monticola sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, lateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure3" httpUri="https://binary.pensoft.net/fig/549196" pageId="0" pageNumber="167">3B-D</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and stout process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167">4A, B</figureCitation>
|
||||
<figureCitation id="996BDE15E262E13B382E66914A9484C9" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and stout process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167">4A, B</figureCitation>
|
||||
) (vs. smaller and not protruding beyond
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
<emphasis id="BA477B998A14DCC9359B30D082DA8CF0" bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
) (Fig.
|
||||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gonopod structure of Tylopus rugosus Golovatch & Enghoff, 1993, ♂ holotype, left gonopod A-C mesal, lateral and dorsal views, respectively. Abbreviations: h strong hook-shaped process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bar: 0.5 mm (after Golovatch and Enghoff 1993)." figureDoi="10.3897/zookeys.1040.66209.figure8" httpUri="https://binary.pensoft.net/fig/549201" pageId="0" pageNumber="167">8B, C</figureCitation>
|
||||
<figureCitation id="8EAE14C3F8DC9CC399DCCE6228578A72" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Gonopod structure of Tylopus rugosus Golovatch & Enghoff, 1993, ♂ holotype, left gonopod A-C mesal, lateral and dorsal views, respectively. Abbreviations: h strong hook-shaped process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bar: 0.5 mm (after Golovatch and Enghoff 1993)." figureDoi="10.3897/zookeys.1040.66209.figure8" httpUri="https://binary.pensoft.net/fig/549201" pageId="0" pageNumber="167">8B, C</figureCitation>
|
||||
), all ♂ legs with the prefemora swollen laterally except for leg 1 (vs. except for legs 1 and 2), coupled with the pleurosternal carinae complete crests with an evident, sharp, caudal denticle produced past the rear tergal margin on segments 4-7, gradually decreasing in size until segments 15(16) (♂) or 13(12) (♀) (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2B, D, E</figureCitation>
|
||||
<figureCitation id="76E44E6ED306E283E64B155A29B2CD8B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2B, D, E</figureCitation>
|
||||
) (vs. same, but gradually decreasing in size until segment 18).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="167" type="description">
|
||||
<paragraph pageId="0" pageNumber="167">Description.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="167">Length of holotype ca 30 mm; adult paratypes 29-31 (♂) or 32-35 mm (♀), width of midbody pro- and metazonae of holotype, 2.4 and 3.6 mm; adult paratypes 2.4-2.5 and 3.4-3.7 mm (♂) or 2.9-3.5 and 3.7-4.5 mm (♀), respectively.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<subSubSection id="358C302DC5095752048B405B2CC396B3" pageId="0" pageNumber="167" type="description">
|
||||
<paragraph id="72AF4469DF7675D3B6A1DF0453DBBEBD" pageId="0" pageNumber="167">Description.</paragraph>
|
||||
<paragraph id="271E910776558C18D7EA1E16E7480FB8" pageId="0" pageNumber="167">Length of holotype ca 30 mm; adult paratypes 29-31 (♂) or 32-35 mm (♀), width of midbody pro- and metazonae of holotype, 2.4 and 3.6 mm; adult paratypes 2.4-2.5 and 3.4-3.7 mm (♂) or 2.9-3.5 and 3.7-4.5 mm (♀), respectively.</paragraph>
|
||||
<paragraph id="11E21439F09D3C71AAE7F27D46D1AD79" pageId="0" pageNumber="167">
|
||||
Colouration of live animals dark brown (Fig.
|
||||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">1A</figureCitation>
|
||||
<figureCitation id="0FC48A3C348CBE7749F51FB68B083BB1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">1A</figureCitation>
|
||||
); calluses of paraterga, venter and legs lighter brown; colouration of alcohol material after two years of preservation faded to dark brown; head, antennae and tip of epiproct light brown, calluses of paraterga yellowish brown to pallid, venter and legs light brown to light yellowish (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2</figureCitation>
|
||||
<figureCitation id="6B2F423565E724FB3353E7548AB966FF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167" start="Figure 1" startId="F1">
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">Figure 1.</emphasis>
|
||||
<caption id="647545F2C9E0CACAE0507C4ACBC6CA2B" doi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167" start="Figure 1" startId="F1">
|
||||
<paragraph id="00EBB17E7B8F18AFD3296C958CDE156C" pageId="0" pageNumber="167">
|
||||
<emphasis id="04F9B12BFFC25E075B257682126CB7A0" bold="true" pageId="0" pageNumber="167">Figure 1.</emphasis>
|
||||
Habitus, live colouration
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">A</emphasis>
|
||||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
<emphasis id="B22287AF3C6DA7A8FFAB8CBE292691FB" bold="true" pageId="0" pageNumber="167">A</emphasis>
|
||||
<taxonomicName id="AA33D2971340789023AB742B5A770E4C" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis id="C65E9880D442F56A06A964987B2773EE" italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., ♂ paratype (CUMZ)
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">B</emphasis>
|
||||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus sutchariti</emphasis>
|
||||
<emphasis id="837ED11B4DF03AFE12B42A1A099EB7E2" bold="true" pageId="0" pageNumber="167">B</emphasis>
|
||||
<taxonomicName id="F3D577C817A1C30353E439B7C1338C38" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus sutchariti" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="sutchariti">
|
||||
<emphasis id="9CC5A68898456329F0BAB5E357D00C08" italics="true" pageId="0" pageNumber="167">Tylopus sutchariti</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<paragraph id="D2F1EB5D47618ECBAEF75B01A2AE4E0B" pageId="0" pageNumber="167">
|
||||
Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae short (Figs
|
||||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">1A</figureCitation>
|
||||
<figureCitation id="CCFC590E2F2B835DC531B1D45ABED3F1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live colouration A Tylopus monticola sp. nov., ♂ paratype (CUMZ) B Tylopus sutchariti sp. nov., ♂ holotype (CUMZ). Scale bars: 1 cm." figureDoi="10.3897/zookeys.1040.66209.figure1" httpUri="https://binary.pensoft.net/fig/549194" pageId="0" pageNumber="167">1A</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, B</figureCitation>
|
||||
<figureCitation id="4EA3DA068CE55D353819A176050513B3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, B</figureCitation>
|
||||
), reaching body segment 3 (♂) or 2 (♀) when stretched dorsally. In width, head <segment 3 <4 <5 <6 <collum <segment 2 <7-16 (♂, ♀); thereafter body gently and gradually tapering. Collum with three transverse rows of strong setae: 3+3 anterior, 2+2 intermediate, and 3+3 posterior; a small lateral incision near midway; caudal corner of paraterga rounded, slightly declined ventrad, not produced past rear tergal margin (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, B</figureCitation>
|
||||
<figureCitation id="3B286B61999C149FF9F1B61C61611FAE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167" start="Figure 2" startId="F2">
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">Figure 2.</emphasis>
|
||||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
<caption id="43A16196914BBE4F0D854F4149190E4E" doi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167" start="Figure 2" startId="F2">
|
||||
<paragraph id="C8943CC48EC017E3F7B92360ABD8F595" pageId="0" pageNumber="167">
|
||||
<emphasis id="B9B5B1DDF8D167E83648A74B9783AD0F" bold="true" pageId="0" pageNumber="167">Figure 2.</emphasis>
|
||||
<taxonomicName id="0FA8525A8A1C61809556A29E6EEE257F" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis id="72481309CCB5EAAAB6BFE3D6D212732E" italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., ♂ holotype (CUMZ)
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
||||
<emphasis id="58C7CE76AF1DF0E1321392F81C1FB003" bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
||||
anterior part of body, dorsal and lateral views, respectively
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">C, D</emphasis>
|
||||
<emphasis id="2A479E84818EC0C1BB45B299CC5CBA46" bold="true" pageId="0" pageNumber="167">C, D</emphasis>
|
||||
segments 10 and 11, dorsal and lateral views, respectively
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">E-G</emphasis>
|
||||
<emphasis id="860EFD55A2DA21A62395876F22DAF15C" bold="true" pageId="0" pageNumber="167">E-G</emphasis>
|
||||
posterior part of body, lateral, subdorsal and subventral views, respectively
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">H, I</emphasis>
|
||||
<emphasis id="9DD18D8FEC84B6F95D2B7E730A38535F" bold="true" pageId="0" pageNumber="167">H, I</emphasis>
|
||||
sternal cones between coxae 4, subcaudal and sublateral views, respectively.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<paragraph id="DABFFB3638D91A8A158DB71F4E9D66EC" pageId="0" pageNumber="167">
|
||||
Tegument rather smooth and shining, prozonae very finely shagreened, metaterga mainly smooth, but often rugulose; surface below paraterga finely microgranulate (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A-F</figureCitation>
|
||||
<figureCitation id="8DAA6249395F2539F73FAAE3C449CF14" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A-F</figureCitation>
|
||||
). Postcollum metaterga with two transverse rows of rather long setae: 2+2 in anterior and 3+3 in posterior row, the latter often abraded, but then readily traceable as insertion points. Tergal setae long, strong, slender, about 1/3 metatergal length. Axial line visible both on pro- and metazonae.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<paragraph id="657CAD8C08745922F68A1339E054D240" pageId="0" pageNumber="167">
|
||||
Paraterga strongly developed (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A-F</figureCitation>
|
||||
<figureCitation id="0A4F5BB0B31299D969DB040D7FBDE12B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A-F</figureCitation>
|
||||
), especially well so in ♂, subhorizontal, slightly upturned posteriorly, always lying high, at upper 1/3 of midbody height, but remaining below dorsum; anterior edge well-developed, mostly regularly rounded and narrowly bordered, fused to callus; caudal corner narrowly rounded, extending increasingly past rear tergal margin, especially strongly so on segments 15-19; in segments 16-19, tips strongly curved mesad, posterior edge slightly oblique (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, C, F</figureCitation>
|
||||
<figureCitation id="01970223C8218AB998506D832D3BCAA7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, C, F</figureCitation>
|
||||
); paraterga very thin blunt blades in lateral view, a little thicker only on pore-bearing segments (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2D</figureCitation>
|
||||
<figureCitation id="4E2DB62E92DC3E5C5B61DD43128D8FCD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2D</figureCitation>
|
||||
). Calluses on paraterga delimited by a sulcus only dorsally. Paraterga 2 broad, lateral edge with three evident incisions: one in anterior 1/3, one at midway, and one at posterior 1/3; anterior incision particularly evident. Paraterga 3 and 4 with two small incisions at lateral edge (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A</figureCitation>
|
||||
<figureCitation id="153A3E3906225DB66AB764D5E62DBBC1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A</figureCitation>
|
||||
), one in anterior 1/3, the other at posterior 1/3; anterior incision also particularly evident. Lateral edge of paraterga of following segments with two small incisions, one in anterior 1/3, the other at midway, caudal incision being smaller in pore-bearing segments (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2C</figureCitation>
|
||||
<figureCitation id="EF3E8E4486BC9071EBA9FFB692A7B949" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2C</figureCitation>
|
||||
). Ozopores evident, lateral, lying in an ovoid groove at about 1/3 metatergal length in front of posterior edge of metaterga (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2D</figureCitation>
|
||||
<figureCitation id="72BD9BFFB2FABB7AE7E7B13008F4DFBB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2D</figureCitation>
|
||||
). Transverse sulcus usually distinct (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, C, F</figureCitation>
|
||||
<figureCitation id="F44F080EFE279DE96151D72BABA0D814" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, C, F</figureCitation>
|
||||
), slightly incomplete on segment 18, complete and clearly visible on metaterga 5-17, deep, reaching the bases of paraterga, arcuate, faintly beaded at bottom. Stricture between pro- and metazonae narrow, shallow, beaded at bottom down to base of paraterga (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, C, F</figureCitation>
|
||||
<figureCitation id="6E16735C76AF6FACD300DC62049A8A45" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2A, C, F</figureCitation>
|
||||
). Pleurosternal carinae complete crests on segment 2-3(4) (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2B</figureCitation>
|
||||
<figureCitation id="FEE19893560B689F11F989D403EBF9F2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2B</figureCitation>
|
||||
), with an evident and sharp denticle caudally on segments 4(5)-7 (♂, ♀), thereafter increasingly well reduced and remaining only a small sharp caudal tooth until segment 15(16) (♂) or 13(12) (♀), thereafter missing (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2B, D, E</figureCitation>
|
||||
<figureCitation id="77726B1009C181FDAA22B445D9C4593C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2B, D, E</figureCitation>
|
||||
). Epiproct (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2E-G</figureCitation>
|
||||
<figureCitation id="FAB78DF33D095755065BE8319A901F5B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2E-G</figureCitation>
|
||||
) conical, flattened dorsoventrally, subtruncate, with two evident apical papillae directed caudally, both pointed at tip; pre-apical papillae evident, lying close to tip. Hypoproct subtrapeziform (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2G</figureCitation>
|
||||
<figureCitation id="CBF06F240AEFC762ECDD038B40082006" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2G</figureCitation>
|
||||
), small setiferous knobs at caudal edge well-separated and evident.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<paragraph id="04023361E8FCD8871B4A620E466B7810" pageId="0" pageNumber="167">
|
||||
Sterna densely setose, without modifications (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2G</figureCitation>
|
||||
<figureCitation id="0D8369B60ABF6FBC17FE5E261955A29F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2G</figureCitation>
|
||||
); cross-impressions shallow; a deeply notched sternal lobe between ♂ coxae 4 (Fig.
|
||||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2H, I</figureCitation>
|
||||
<figureCitation id="75342DF5F6C29C31A0976D8EBFED31E7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B anterior part of body, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E-G posterior part of body, lateral, subdorsal and subventral views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively." figureDoi="10.3897/zookeys.1040.66209.figure2" httpUri="https://binary.pensoft.net/fig/549195" pageId="0" pageNumber="167">2H, I</figureCitation>
|
||||
). Legs long and slender, midbody ones ca 1.4-1.5 (♂) or 0.9-1.0 (♀) as long as body height; all ♂ legs except leg 1 with prefemora swollen laterally; femora and tibiae with particularly dense setae and ventral microgranulations; legs on segments 7-18 with an evident adenostyle (tubercle) medially on each postfemur and tibia (Fig.
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and stout process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167">4C</figureCitation>
|
||||
<figureCitation id="D2F6FB0BC5C794FA77C60E3EBC4D7560" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and stout process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167">4C</figureCitation>
|
||||
); tarsal brushes absent.
|
||||
</paragraph>
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<paragraph id="4D75EF4D0D4A0984B6D60D4BDE64CEC8" pageId="0" pageNumber="167">
|
||||
Gonopods (Figs
|
||||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tylopus monticola sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, lateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure3" httpUri="https://binary.pensoft.net/fig/549196" pageId="0" pageNumber="167">3</figureCitation>
|
||||
<figureCitation id="8111AE08C6BC10A931836EBDDEB69679" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Tylopus monticola sp. nov., ♂ holotype (CUMZ), left gonopod A, B mesal and lateral views, respectively C-F distal part, submesal, lateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1040.66209.figure3" httpUri="https://binary.pensoft.net/fig/549196" pageId="0" pageNumber="167">3</figureCitation>
|
||||
,
|
||||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and stout process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167">4A, B</figureCitation>
|
||||
<figureCitation id="60BF5B0C9CE1D599ED0A6292F1C036E0" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tylopus monticola sp. nov., ♂ holotype (CUMZ) A, B left gonopod, mesal and lateral views, respectively C leg of segment 10. Abbreviations: g mesal groove / hollow of femorite, h short and stout process of femorite, l apicolateral lobe of femorite, z serrate process of femorite. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167">4A, B</figureCitation>
|
||||
) simple; coxite slightly curved caudad, sparsely setose distoventrally. Prefemorite densely setose, about 1/3 as long as femorite +
|
||||
<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
||||
<normalizedToken id="827FF424ED74B2B8F82E594871166974" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
||||
part. Femorite rather stout, expanded distad, suberect, showing a distinct mesal groove/hollow (
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">g</emphasis>
|
||||
<emphasis id="FDEA65EDE9451CFF258D406D28DC9619" bold="true" pageId="0" pageNumber="167">g</emphasis>
|
||||
); apicolateral lobe (
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
<emphasis id="029AFC2453C8E00DDF3578A661E5FC05" bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
) simple; process
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">z</emphasis>
|
||||
<emphasis id="0F51026E769186548AA7C6B0BB096253" bold="true" pageId="0" pageNumber="167">z</emphasis>
|
||||
large and long, serrate along dorsal margin and protruding beyond apicolateral lobe (
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
<emphasis id="1C7E66F4210EBA9BD6C7B622FC7A8356" bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
); process
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">h</emphasis>
|
||||
<emphasis id="75657721D4B2721363C5B8C42DF40BAC" bold="true" pageId="0" pageNumber="167">h</emphasis>
|
||||
short and stout, suberect, with a narrowly rounded tip; solenophore long and slender, typically coiled, tip subtruncate.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1040.66209.figure3" httpUri="https://binary.pensoft.net/fig/549196" pageId="0" pageNumber="167" start="Figure 3" startId="F3">
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">Figure 3.</emphasis>
|
||||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
<caption id="124DD33801BB001A3131549ED738176C" doi="10.3897/zookeys.1040.66209.figure3" httpUri="https://binary.pensoft.net/fig/549196" pageId="0" pageNumber="167" start="Figure 3" startId="F3">
|
||||
<paragraph id="E0DCA541DEAB80A1ADFCA16B9A400A01" pageId="0" pageNumber="167">
|
||||
<emphasis id="8BC4C8FCFBE2E3A17711E53204EABA4D" bold="true" pageId="0" pageNumber="167">Figure 3.</emphasis>
|
||||
<taxonomicName id="E23802E3C0E391B0A866C212467B45CC" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis id="B013F69853A14B957E1F20605AD53492" italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., ♂ holotype (CUMZ), left gonopod
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
||||
<emphasis id="5BFB49F3DE03795099D3BF9114F1088B" bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
||||
mesal and lateral views, respectively
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">C-F</emphasis>
|
||||
<emphasis id="4CBB608A6E6ABA8804EE8438FEC082DC" bold="true" pageId="0" pageNumber="167">C-F</emphasis>
|
||||
distal part, submesal, lateral, suboral and subcaudal views, respectively. Scale bars: 0.2 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="167" type="name">
|
||||
<paragraph pageId="0" pageNumber="167">Name.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<subSubSection id="3459A96D2F8EDE5010EA2513E2E114BB" pageId="0" pageNumber="167" type="name">
|
||||
<paragraph id="122D84CF9A7BA9B1B71B11BC4E90D4D0" pageId="0" pageNumber="167">Name.</paragraph>
|
||||
<paragraph id="10C0A69E4938742009FC778C081DC9EC" pageId="0" pageNumber="167">
|
||||
To emphasize the habitats where this new species was discovered; "
|
||||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
<taxonomicName id="9F7EB176737D35E539AAE7B3B4517F2A" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis id="AB06D84EBD93DF3969F7B797B8D5EB94" italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
</taxonomicName>
|
||||
" meaning a mountain-dweller or a highlander; noun in apposition.
|
||||
</paragraph>
|
||||
<caption doi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167" start="Figure 4" startId="F4">
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">Figure 4.</emphasis>
|
||||
<taxonomicName authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
<caption id="2E0A6954396E733009B13DF78BEFF3E2" doi="10.3897/zookeys.1040.66209.figure4" httpUri="https://binary.pensoft.net/fig/549197" pageId="0" pageNumber="167" start="Figure 4" startId="F4">
|
||||
<paragraph id="2DCFA33CF9639987A56A0E046D88DECC" pageId="0" pageNumber="167">
|
||||
<emphasis id="05840859BA492B171F20D1F8452A6A18" bold="true" pageId="0" pageNumber="167">Figure 4.</emphasis>
|
||||
<taxonomicName id="F6649EAF84DC6ECCDD5A5C655F225A5E" authorityName="Likhitrakarn & Golovatch & Srisonchai & Panha" authorityYear="2021" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus monticola" order="Polydesmida" pageId="0" pageNumber="167" phylum="Arthropoda" rank="species" species="monticola">
|
||||
<emphasis id="2FF7A027283530CF82E1D09ED11935CE" italics="true" pageId="0" pageNumber="167">Tylopus monticola</emphasis>
|
||||
</taxonomicName>
|
||||
sp. nov., ♂ holotype (CUMZ)
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
||||
<emphasis id="9C059FBF818FDDC1ECBF7E345E480B6D" bold="true" pageId="0" pageNumber="167">A, B</emphasis>
|
||||
left gonopod, mesal and lateral views, respectively
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">C</emphasis>
|
||||
<emphasis id="CA2B86A78F5342E8B9A7E0AE80DE72B0" bold="true" pageId="0" pageNumber="167">C</emphasis>
|
||||
leg of segment 10. Abbreviations:
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">g</emphasis>
|
||||
<emphasis id="EC1198BFBF3D75C1D2097320ED8F1076" bold="true" pageId="0" pageNumber="167">g</emphasis>
|
||||
mesal groove/hollow of femorite,
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">h</emphasis>
|
||||
<emphasis id="22BDA579E85D636F749FD7080962C793" bold="true" pageId="0" pageNumber="167">h</emphasis>
|
||||
short and stout process of femorite,
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
<emphasis id="F7ABE54E00001A44851C9408FAAB0243" bold="true" pageId="0" pageNumber="167">l</emphasis>
|
||||
apicolateral lobe of femorite,
|
||||
<emphasis bold="true" pageId="0" pageNumber="167">z</emphasis>
|
||||
<emphasis id="AF26652959C18828069E0CDA7F54573A" bold="true" pageId="0" pageNumber="167">z</emphasis>
|
||||
serrate process of femorite. Scale bars: 0.5 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="0" pageNumber="167" type="remarks">
|
||||
<paragraph pageId="0" pageNumber="167">Remark.</paragraph>
|
||||
<paragraph pageId="0" pageNumber="167">
|
||||
<subSubSection id="92A0184ABA44290DC1BA88EAB98FBDE9" pageId="0" pageNumber="167" type="remarks">
|
||||
<paragraph id="9AC70A3E5F2B899CF8088BC16CAB0ECF" pageId="0" pageNumber="167">Remark.</paragraph>
|
||||
<paragraph id="576F03BD1F241D89A14AE0BFE7F79BD8" pageId="0" pageNumber="167">
|
||||
The species was found quite far away (about 120 air-km) from the type locality of the most similar species,
|
||||
<taxonomicName lsidName="T. rugosus" pageId="0" pageNumber="167" rank="species" species="rugosus">
|
||||
<emphasis italics="true" pageId="0" pageNumber="167">T. rugosus</emphasis>
|
||||
<taxonomicName id="A93F2DDD99C4816E8A3925951A1CC563" lsidName="T. rugosus" pageId="0" pageNumber="167" rank="species" species="rugosus">
|
||||
<emphasis id="F176B6918D16AC30509A56E3E9EF6416" italics="true" pageId="0" pageNumber="167">T. rugosus</emphasis>
|
||||
</taxonomicName>
|
||||
Golovatch & Enghoff, 1993 (Fig.
|
||||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distributions of Tylopus species recorded from Myanmar. Pink circle: T. brehieri Golovatch, VandenSpiegel & Semenyuk, 2016; Red Asterisk: T. punctus Likhitrakarn, Golovatch & Panha, 2016; Yellow inverted triangle: T. monticola sp. nov.; Blue square: T. sutchariti sp. nov. and T. monticola sp. nov.; Orange diamond: T. silvestris (Pocock, 1895); Green triangle: T. doriae (Pocock, 1895)." figureDoi="10.3897/zookeys.1040.66209.figure9" httpUri="https://binary.pensoft.net/fig/549202" pageId="0" pageNumber="167">9</figureCitation>
|
||||
<figureCitation id="11DAC2F6218941E7F8A98DC0FD60C085" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distributions of Tylopus species recorded from Myanmar. Pink circle: T. brehieri Golovatch, VandenSpiegel & Semenyuk, 2016; Red Asterisk: T. punctus Likhitrakarn, Golovatch & Panha, 2016; Yellow inverted triangle: T. monticola sp. nov.; Blue square: T. sutchariti sp. nov. and T. monticola sp. nov.; Orange diamond: T. silvestris (Pocock, 1895); Green triangle: T. doriae (Pocock, 1895)." figureDoi="10.3897/zookeys.1040.66209.figure9" httpUri="https://binary.pensoft.net/fig/549202" pageId="0" pageNumber="167">9</figureCitation>
|
||||
). Both new species described here have been found to occur syntopically.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
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