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<taxonomicName LSID="20070E29-AF8A-5A3B-BDFB-F018D2BB4A2B" authority="Ng &amp; Takeda, 1992" authorityName="Ng &amp; Takeda" authorityYear="1992" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora hoiseni" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hoiseni">Johora hoiseni Ng &amp; Takeda, 1992</taxonomicName>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">Figures 1D</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">, 2D</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">, 3D</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">, 4D</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">, 5D</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">, 8E-H</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">, 12D</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">, 13D</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">, 14D</figureCitation>
<treatment id="20070E29AF8A5A3BBDFBF018D2BB4A2B" ID-GBIF-Taxon="169094346" LSID="urn:lsid:plazi:treatment:20070E29AF8A5A3BBDFBF018D2BB4A2B" httpUri="http://treatment.plazi.org/id/20070E29AF8A5A3BBDFBF018D2BB4A2B" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Potamidae" scope_infraOrder="Brachyura">
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<taxonomicName id="1F3472151BEE43A38B958EF36D1D69BE" ID-CoL="6NG6X" LSID="20070E29-AF8A-5A3B-BDFB-F018D2BB4A2B" authority="Ng &amp; Takeda, 1992" authorityName="Ng &amp; Takeda" authorityYear="1992" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora hoiseni" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hoiseni">Johora hoiseni Ng &amp; Takeda, 1992</taxonomicName>
<figureCitation id="ADE9CE869469BF286B48F1EFB9C32550" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">Figures 1D</figureCitation>
<figureCitation id="1B5A1B7E206AE905D28E821E47145FC5" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">, 2D</figureCitation>
<figureCitation id="1D9BD1464547B8A98C4675474FC78DA9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">, 3D</figureCitation>
<figureCitation id="3EC42DD4E78D5F8EE71C3C308D6F3BEF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">, 4D</figureCitation>
<figureCitation id="E434FA5059508A3D6CCB4333D8C43F2E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">, 5D</figureCitation>
<figureCitation id="E32C7F0DF648A30E28098F7E6A89B9F2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">, 8E-H</figureCitation>
<figureCitation id="38CE35A5DBF4E2564541D5648972BA42" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">, 12D</figureCitation>
<figureCitation id="36A9EAFE52780231F8F8E26F7C6D6288" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">, 13D</figureCitation>
<figureCitation id="A263C71C8A5D6A0557FDFE5E5BF6343B" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">, 14D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Potamiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamiscus (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Potamiscus (Johora) johorensis tahanensis</taxonomicName>
<subSubSection id="DC21BCFB0A562C5576B076554865BCEE" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="1CB438DB87BD3FA5B60ADDD59E5F989C" pageId="0" pageNumber="1">
<taxonomicName id="41D17FB5EB77F1629E33B14DFDD0766D" authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Potamiscus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamiscus (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Potamiscus (Johora) johorensis tahanensis</taxonomicName>
Bott, 1966: 495 (part), fig. 32 (not
<taxonomicName class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
<taxonomicName id="62FDD7FE319A54DE12CF5D87EB4E0E81" class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
Bott, 1966 s. str.).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
<paragraph id="95DE049C9B100300098A284697344CD9" pageId="0" pageNumber="1">
<taxonomicName id="BC053057C647B949CFDD22E4C72DEE64" class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
-
<bibRefCitation author="Bott, R" journalOrPublisher="Abhandlungen der senckenbergischen naturforschenden Gesellschaft" pageId="0" pageNumber="1" pagination="1 - 338" refId="B2" refString="Bott, R, 1970. Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der senckenbergischen naturforschenden Gesellschaft 526: 1 - 338" title="Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda)." volume="526" year="1970">Bott 1970</bibRefCitation>
<bibRefCitation id="7D8E0367E3218D549E1BC2EB5781BDAD" author="Bott, R" journalOrPublisher="Abhandlungen der senckenbergischen naturforschenden Gesellschaft" pageId="0" pageNumber="1" pagination="1 - 338" refId="B2" refString="Bott, R, 1970. Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der senckenbergischen naturforschenden Gesellschaft 526: 1 - 338" title="Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda)." volume="526" year="1970">Bott 1970</bibRefCitation>
: 181 (part), pl. 40 fig. 57;
<bibRefCitation author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="167 - 174" refId="B20" refString="Ng, PKL, Tan, LWH, 1984. A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia. Malayan Nature Journal 37 (3): 167 - 174" title="A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia." volume="37" year="1984">Ng and Tan 1984</bibRefCitation>
<bibRefCitation id="1476CFE03BC2F0F7AB975A9203011231" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="167 - 174" refId="B20" refString="Ng, PKL, Tan, LWH, 1984. A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia. Malayan Nature Journal 37 (3): 167 - 174" title="A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia." volume="37" year="1984">Ng and Tan 1984</bibRefCitation>
: 172, fig. 7 (not
<taxonomicName class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
<taxonomicName id="E50B237DFC7076CEF962BD853A3B1BB7" class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
Bott, 1966).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tahanensis">Johora tahanensis</taxonomicName>
<paragraph id="2EB774FF1949BFCEC44079B9806DCE3B" pageId="0" pageNumber="1">
<taxonomicName id="2CBBA8AAB60E0E3A17BE1E4DF2B39A4A" authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tahanensis">Johora tahanensis</taxonomicName>
-
<bibRefCitation author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">Ng 1987</bibRefCitation>
<bibRefCitation id="3B47D1ECFA951C842101EFA7C513D6D4" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">Ng 1987</bibRefCitation>
: 33 (part), fig. 9B-K;
<bibRefCitation author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">Ng 1988</bibRefCitation>
<bibRefCitation id="5D2033AEE9E7AB6050AFA47CAEF37A47" author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">Ng 1988</bibRefCitation>
: 42 (part), fig. 18A, B, D, E, G (not
<taxonomicName class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
<taxonomicName id="DC6D6200FC99FDAB94F3B52588BE148B" class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
Bott, 1966).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ng &amp; Takeda" authorityYear="1992" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora hoiseni" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hoiseni">Johora hoiseni</taxonomicName>
<paragraph id="0F0AE23578DF01FB9FE4D584CB68085D" pageId="0" pageNumber="1">
<taxonomicName id="926C6BEAC0BB2E8D0047060CCAC81861" authorityName="Ng &amp; Takeda" authorityYear="1992" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora hoiseni" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hoiseni">Johora hoiseni</taxonomicName>
Ng &amp; Takeda, 1992: 108, figs 2, 3;
<bibRefCitation author="Ng, PKL" editor="Yule, C" journalOrPublisher="Malaysian Academy of Sciences, city" pageId="0" pageNumber="1" pagination="311 - 336" refId="B14" refString="Ng, PKL, 2004. Crustacea: Decapoda, Brachyura. In: Yule, C, Yong, HS, Eds., Freshwater Invertebrates of the Malaysian Region. Malaysian Academy of Sciences, city: 311 - 336" title="Crustacea: Decapoda, Brachyura." volumeTitle="Freshwater Invertebrates of the Malaysian Region." year="2004">Ng 2004</bibRefCitation>
<bibRefCitation id="F4BB7DFD2DA9119A63A3C5704BD9F2DD" author="Ng, PKL" editor="Yule, C" journalOrPublisher="Malaysian Academy of Sciences, city" pageId="0" pageNumber="1" pagination="311 - 336" refId="B14" refString="Ng, PKL, 2004. Crustacea: Decapoda, Brachyura. In: Yule, C, Yong, HS, Eds., Freshwater Invertebrates of the Malaysian Region. Malaysian Academy of Sciences, city: 311 - 336" title="Crustacea: Decapoda, Brachyura." volumeTitle="Freshwater Invertebrates of the Malaysian Region." year="2004">Ng 2004</bibRefCitation>
: 321;
<bibRefCitation author="Ng, PKL" editor="Chua, L" journalOrPublisher="28 - 30 June 2005, Forest Research Institute Malaysia, Kepong" pageId="0" pageNumber="1" pagination="95 - 120" refId="B21" refString="Ng, PKL, Yeo, DCJ, 2007. Malaysian freshwater crabs: conservation prospects and challenges. In: Chua, L, Ed., Proceedings of the Seminar on the Status of Biological Diversity in Malaysia &amp; Threat Assessment of Plant Species in Malaysia. 28-30 June 2005, Forest Research Institute Malaysia, Kepong: 95 - 120" title="Malaysian freshwater crabs: conservation prospects and challenges." volumeTitle="Proceedings of the Seminar on the Status of Biological Diversity in Malaysia &amp; Threat Assessment of Plant Species in Malaysia." year="2007">Ng and Yeo 2007</bibRefCitation>
<bibRefCitation id="E07163620845C81C0C3DFB65BC99BFBC" author="Ng, PKL" editor="Chua, L" journalOrPublisher="28 - 30 June 2005, Forest Research Institute Malaysia, Kepong" pageId="0" pageNumber="1" pagination="95 - 120" refId="B21" refString="Ng, PKL, Yeo, DCJ, 2007. Malaysian freshwater crabs: conservation prospects and challenges. In: Chua, L, Ed., Proceedings of the Seminar on the Status of Biological Diversity in Malaysia &amp; Threat Assessment of Plant Species in Malaysia. 28-30 June 2005, Forest Research Institute Malaysia, Kepong: 95 - 120" title="Malaysian freshwater crabs: conservation prospects and challenges." volumeTitle="Proceedings of the Seminar on the Status of Biological Diversity in Malaysia &amp; Threat Assessment of Plant Species in Malaysia." year="2007">Ng and Yeo 2007</bibRefCitation>
: 100;
<bibRefCitation author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="1" pagination="1 - 286" refId="B18" refString="Ng, PKL, Guinot, D, Davie, PJF, 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286" title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world." volume="17" year="2008">Ng et al. 2008</bibRefCitation>
<bibRefCitation id="A4A1B8A5D4A07ED23B848215F25D45E9" author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="0" pageNumber="1" pagination="1 - 286" refId="B18" refString="Ng, PKL, Guinot, D, Davie, PJF, 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286" title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world." volume="17" year="2008">Ng et al. 2008</bibRefCitation>
: 163;
<bibRefCitation DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. 2009</bibRefCitation>
<bibRefCitation id="A61C67C1AD6DB9C5A7B28F75D69BFD10" DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. 2009</bibRefCitation>
: table.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
<subSubSection id="2AB0568BDD4AF697FB92A608040A5B97" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="0AD7092C7C388CCAEDE96DA3F84F59E8" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="B0F34BD071B607EA256E8D729BEC5D70" pageId="0" pageNumber="1">
<emphasis id="2066E5951B8E3BDE9163762F171D4DC4" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: male (25.1
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="40B5014419EE58A7EA4A349FB205E682" originalValue="×">x</normalizedToken>
21.1 mm) (ZRC 1984.6673), stream entering Sungei Galas, near Gua Madir, Taman Negara National Park, Kelantan, ca.
<geoCoordinate degrees="4" direction="north" minutes="51" orientation="latitude" precision="15" seconds="30" value="4.858333">4°51'30&quot;N</geoCoordinate>
<geoCoordinate id="82B18C91D86640310E37050716673F34" degrees="4" direction="north" minutes="51" orientation="latitude" precision="15" seconds="30" value="4.858333">4°51'30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="102" direction="east" minutes="03" orientation="longitude" precision="15" seconds="23" value="102.05639">102°03'23&quot;E</geoCoordinate>
<geoCoordinate id="2C1453B18AE4EBC5611F9CE1C3E78716" degrees="102" direction="east" minutes="03" orientation="longitude" precision="15" seconds="23" value="102.05639">102°03'23&quot;E</geoCoordinate>
, coll. Tweedie MWF, August 1939. Paratypes: 53 males, 13 females (ZRC 1984.6674-6755), same data as holotype. Others: 3 males, 18 females (ZRC 1984.7683-7687), stream entering Sungei Galas, near Gua Madir, Taman Negara National Park, Kelantan, ca.
<geoCoordinate degrees="4" direction="north" minutes="51" orientation="latitude" precision="15" seconds="30" value="4.858333">4°51'30&quot;N</geoCoordinate>
<geoCoordinate id="65A85F6C3B52AEA9BA2FD242803B87D8" degrees="4" direction="north" minutes="51" orientation="latitude" precision="15" seconds="30" value="4.858333">4°51'30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="102" direction="east" minutes="03" orientation="longitude" precision="15" seconds="23" value="102.05639">102°03'23&quot;E</geoCoordinate>
<geoCoordinate id="F0B8DD18C1FC200292F050DE15C1C418" degrees="102" direction="east" minutes="03" orientation="longitude" precision="15" seconds="23" value="102.05639">102°03'23&quot;E</geoCoordinate>
, coll. Tweedie MWF, August 1939; 2 males, 5 females (ZRC 1989.3617-3623), Pahang or Kelantan?, no other data. All locations in Peninsular Malaysia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="885A7E0E8E069CC6CC7EE50D7103B67B" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="44B1013B1C4457C66C396973A5613B84" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="DD189A7720D09D7295C6DBE437280CAC" pageId="0" pageNumber="1">
Adult carapace width to length ratio 1.16-1.19 (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1D</figureCitation>
<figureCitation id="E8152709EB99F8573241852AA7CDE78D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1D</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
<figureCitation id="69B705550F49A924481485F34103A636" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12D</figureCitation>
<figureCitation id="ABD9A74E404DE93DA03D62D3AA1C85FA" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12D</figureCitation>
); dorsal surface gently convex in frontal view, not inflated (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3D</figureCitation>
<figureCitation id="FFDECB557FB069DAD5EC3CFF3FB26BEF" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3D</figureCitation>
); frontal margin slightly sinuous to almost straight (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
<figureCitation id="6619DC244994CC96167351945814724B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
); suborbital, pterygostomial and sub-branchial regions rugose, pterygostomial region covered with dense setae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3D</figureCitation>
<figureCitation id="A45B9CCCEE56B0C9F8850177C2108BF4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3D</figureCitation>
); epigastric cristae distinct, distinctly anterior to sharp postorbital cristae, postorbital cristae with lateral edges low, joining lateral margin through oblique striae (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
<figureCitation id="4A6B31EF54A15723DB3D141C171395F2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
); external orbital tooth separated from epibranchial tooth by distinct cleft, epibranchial tooth sharp, distinct (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
<figureCitation id="8050CDA22201CB432E671FD5ECC2C09C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
); anterolateral margin gently convex (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
<figureCitation id="3FBDBFB3AEBDF960DEFA02718D76D49F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
); posterolateral margin medially concave to sinuous, gently converging towards gently convex, entire posterior carapace margin (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
<figureCitation id="C701704638FE1437E3DE1D35A2467A52" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2D</figureCitation>
); posterior margin of epistome with triangular median triangle, lateral margin obliquely sloping (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3D</figureCitation>
<figureCitation id="A949D6E7829835BC90469797248653E3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3D</figureCitation>
); outer surfaces of third maxillipeds with dense, long stiff setae; ischium subrectangular, with shallow median oblique groove (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3D</figureCitation>
<figureCitation id="DD2A54A863EC3363E24A4436FFEE6D4E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3D</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">4D</figureCitation>
<figureCitation id="8F9A73CEE5C934F07D91E3F009C73F3E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">4D</figureCitation>
); ambulatory legs not elongate, length to width ratio of merus of fourth ambulatory leg 2.7-2.8 (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1D</figureCitation>
<figureCitation id="4EF5F4322334BB1B63316B2154D14432" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1D</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12D</figureCitation>
<figureCitation id="56A3B7568B856985C1069D6174C9E993" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12D</figureCitation>
); G1 subterminal segment with broad proximal part, tapering suddenly to slender distal part, with distinct shelf-like structure along outer margin; terminal segment straight, ca. two-thirds length of subterminal segment, surfaces with numerous short setae (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8E-G</figureCitation>
<figureCitation id="617993CB1C3E15C2643BCC50F7B37892" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8E-G</figureCitation>
); G2 slightly longer than G1, distal segment long, about half length of basal segment (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8H</figureCitation>
<figureCitation id="6DF1D379D18A537773C4C64036646CFB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8H</figureCitation>
). Female pleon ovate; somite 3 less wide than somite 4, somites 4-6 progressively narrower; telson semicircular (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">13D</figureCitation>
<figureCitation id="4CD324D4DB51F3D718EF09DAA0DB9256" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">13D</figureCitation>
). Vulvae large, on anterior half of sternite 6, slightly pushing into suture with sternite 5, lateral sternal vulvar cover semicircular (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14D</figureCitation>
<figureCitation id="A7EA13211C3DB4F7A56C23FED66E29FE" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda (1992)</bibRefCitation>
<subSubSection id="6BD5AE391730ABBA974FB5842CE14F0A" pageId="0" pageNumber="1" type="remarks">
<paragraph id="2525E695B6FCC9619507FE9EC4DFC732" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="EE4146734B613BFF1C44F5F9ED717209" pageId="0" pageNumber="1">
<bibRefCitation id="00D11D64ADA34F44283BE1EA636F9237" author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda (1992)</bibRefCitation>
showed that the G1 differences of the material that had been identified as &quot;
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
<taxonomicName id="2B34B6D7544A769ADDD7596BECA66324" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis id="DA9EE79EBE33E17984DE223BED003E90" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
&quot; by
<bibRefCitation author="Bott, R" journalOrPublisher="Senckenbergiana biologica" pageId="0" pageNumber="1" pagination="469 - 509" refId="B1" refString="Bott, R, 1966. Potamiden aus Asien (Potamon Savigny and Potamiscus Alcock) (Crustacea, Decapoda). Senckenbergiana biologica 47: 469 - 509" title="Potamiden aus Asien (Potamon Savigny and Potamiscus Alcock) (Crustacea, Decapoda)." volume="47" year="1966">Bott (1966</bibRefCitation>
<bibRefCitation id="1D3115D56CBAEC050764F4B311BE2A55" author="Bott, R" journalOrPublisher="Senckenbergiana biologica" pageId="0" pageNumber="1" pagination="469 - 509" refId="B1" refString="Bott, R, 1966. Potamiden aus Asien (Potamon Savigny and Potamiscus Alcock) (Crustacea, Decapoda). Senckenbergiana biologica 47: 469 - 509" title="Potamiden aus Asien (Potamon Savigny and Potamiscus Alcock) (Crustacea, Decapoda)." volume="47" year="1966">Bott (1966</bibRefCitation>
,
<bibRefCitation author="Bott, R" journalOrPublisher="Abhandlungen der senckenbergischen naturforschenden Gesellschaft" pageId="0" pageNumber="1" pagination="1 - 338" refId="B2" refString="Bott, R, 1970. Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der senckenbergischen naturforschenden Gesellschaft 526: 1 - 338" title="Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda)." volume="526" year="1970">1970</bibRefCitation>
<bibRefCitation id="603385B844DD3AC9DFCE7D37C5AE585D" author="Bott, R" journalOrPublisher="Abhandlungen der senckenbergischen naturforschenden Gesellschaft" pageId="0" pageNumber="1" pagination="1 - 338" refId="B2" refString="Bott, R, 1970. Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der senckenbergischen naturforschenden Gesellschaft 526: 1 - 338" title="Die Suesswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda)." volume="526" year="1970">1970</bibRefCitation>
;
<bibRefCitation author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">Ng 1987</bibRefCitation>
<bibRefCitation id="E0DFDDFFA8C3EEEE2718DDAE3676B8A5" author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">Ng 1987</bibRefCitation>
,
<bibRefCitation author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">1988</bibRefCitation>
<bibRefCitation id="EC552B90D07D5C74F106C5E526765515" author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">1988</bibRefCitation>
) belong to a distinct species,
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
<taxonomicName id="11C5E3633264D29942BF3EDC401EF012" lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis id="6A25D396F202083BEA543AD19557F805" italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
(see remarks for
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
<taxonomicName id="439A26AC52DD7DF5DAD2668938F72D88" lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis id="9A83F430BFE0E2D1FEF8EAF9A40035E8" italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ng &amp; Takeda" authorityYear="1992" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora hoiseni" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hoiseni">
<emphasis italics="true" pageId="0" pageNumber="1">Johora hoiseni</emphasis>
<subSubSection id="2A03B8364686295DA64971D9AA527A07" pageId="0" pageNumber="1" type="distribution">
<paragraph id="BCE3C601B3A1EF50D39965D82BC22F6F" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="7C0C7EA121D071599EAA4FFBAB931B23" pageId="0" pageNumber="1">
<taxonomicName id="DC444330FD50C4514FC69140E0B0FA4A" authorityName="Ng &amp; Takeda" authorityYear="1992" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora hoiseni" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hoiseni">
<emphasis id="BF1AAA4B0F32FE790C6F3DEA7B477D21" italics="true" pageId="0" pageNumber="1">Johora hoiseni</emphasis>
</taxonomicName>
is known from the drainages near Gua Musang in Kelantan, in the northwestern part of Taman Negara National Park (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of species of the Johora tahanensis species group 1 J. tahanensis (Bott, 1966) 2 J. thoi Ng, 1990 3 J. hoiseni Ng &amp; Takeda, 1992 4 J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005 5 J. booliati sp. nov. 6 J. erici sp. nov. 7 J. michaeli sp. nov. Map data 2020 Google." figureDoi="10.3897/zookeys.994.56810.figure15" httpUri="https://binary.pensoft.net/fig/476326" pageId="0" pageNumber="1">15</figureCitation>
<figureCitation id="6C113D3681FB91B2A704B0168F7556BA" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of species of the Johora tahanensis species group 1 J. tahanensis (Bott, 1966) 2 J. thoi Ng, 1990 3 J. hoiseni Ng &amp; Takeda, 1992 4 J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005 5 J. booliati sp. nov. 6 J. erici sp. nov. 7 J. michaeli sp. nov. Map data 2020 Google." figureDoi="10.3897/zookeys.994.56810.figure15" httpUri="https://binary.pensoft.net/fig/476326" pageId="0" pageNumber="1">15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="conservation">
<paragraph pageId="0" pageNumber="1">Conservation.</paragraph>
<paragraph pageId="0" pageNumber="1">
<subSubSection id="8707D250988F39FE6D530FEDD3B6DD9A" pageId="0" pageNumber="1" type="conservation">
<paragraph id="2806CD2DFD65C4EAE053DC397ADBBBD7" pageId="0" pageNumber="1">Conservation.</paragraph>
<paragraph id="20477DE17E51A82E92D5FB6D3C64717B" pageId="0" pageNumber="1">
The species is not under any immediate threat as it is found in
<normalizedToken originalValue="Malaysia’s">Malaysia's</normalizedToken>
<normalizedToken id="9F4A9CD643E7A755C758BB67765F4F81" originalValue="Malaysia’s">Malaysia's</normalizedToken>
oldest and largest national park.
<bibRefCitation author="Ng, PKL" editor="Chua, L" journalOrPublisher="28 - 30 June 2005, Forest Research Institute Malaysia, Kepong" pageId="0" pageNumber="1" pagination="95 - 120" refId="B21" refString="Ng, PKL, Yeo, DCJ, 2007. Malaysian freshwater crabs: conservation prospects and challenges. In: Chua, L, Ed., Proceedings of the Seminar on the Status of Biological Diversity in Malaysia &amp; Threat Assessment of Plant Species in Malaysia. 28-30 June 2005, Forest Research Institute Malaysia, Kepong: 95 - 120" title="Malaysian freshwater crabs: conservation prospects and challenges." volumeTitle="Proceedings of the Seminar on the Status of Biological Diversity in Malaysia &amp; Threat Assessment of Plant Species in Malaysia." year="2007">Ng and Yeo (2007)</bibRefCitation>
<bibRefCitation id="CD6945F7EC54946177964C0C48052A32" author="Ng, PKL" editor="Chua, L" journalOrPublisher="28 - 30 June 2005, Forest Research Institute Malaysia, Kepong" pageId="0" pageNumber="1" pagination="95 - 120" refId="B21" refString="Ng, PKL, Yeo, DCJ, 2007. Malaysian freshwater crabs: conservation prospects and challenges. In: Chua, L, Ed., Proceedings of the Seminar on the Status of Biological Diversity in Malaysia &amp; Threat Assessment of Plant Species in Malaysia. 28-30 June 2005, Forest Research Institute Malaysia, Kepong: 95 - 120" title="Malaysian freshwater crabs: conservation prospects and challenges." volumeTitle="Proceedings of the Seminar on the Status of Biological Diversity in Malaysia &amp; Threat Assessment of Plant Species in Malaysia." year="2007">Ng and Yeo (2007)</bibRefCitation>
treated the species as endangered as it is only known from a relatively small geographic area but
<bibRefCitation DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. (2009)</bibRefCitation>
<bibRefCitation id="81663422EC4FBA9C5C10BA62699FC608" DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. (2009)</bibRefCitation>
listed it as of least concern as it is in a protected area.
</paragraph>
</subSubSection>

116
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: 38 [general description], lectotype female (ANSP) and paralectotype male designated, Plate 4, figs 35-35a; type-locality: &quot;Dutch Guiana, S. America&quot;.
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: 22 [synonymic catalogue of
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<bibRefCitation id="EB1BFE728B32B43E9EE7D073BF85FDCA" author="Bruner, L" journalOrPublisher="Annals of the Carnegie Museum" pageId="0" pageNumber="45" pagination="89 - 143" refId="B3" refString="Bruner, L, 1910. South American Tetrigidae. Annals of the Carnegie Museum 7: 89 - 143" title="South American Tetrigidae." volume="7" year="1910">Bruner 1910</bibRefCitation>
[key, new localities].
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: 263 [new localities].
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: 331 [catalogue of Brazilian
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]. Otte D. 1979 (1978): 39 [catalogue of primary types at ANSP].
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281
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<mods:detail id="B379ABDB0ABFD373BB6A81B963751E59" type="volume">
<mods:number id="F95FD787BF4C38BCB4B04622A1B03204">596</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>128</mods:end>
<mods:extent id="D9BC7B10FDD497D66A3BDCBA406C69C0" unit="page">
<mods:start id="B7231B4DD8EC86FA72C85D6DEDD4BB62">87</mods:start>
<mods:end id="AD11876E46EF8B5AAEA0152F53293C7E">128</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.596.8725</mods:url>
<mods:location id="4F683E480D7B714CCAF5428020983A2D">
<mods:url id="85DC2D7532575CE6956184469A2B06D5">http://dx.doi.org/10.3897/zookeys.596.8725</mods:url>
</mods:location>
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</mods:mods>
<treatment ID-GBIF-Taxon="127885602" LSID="urn:lsid:plazi:treatment:20230174E5DFBEFA9E27B22779B10D07" httpUri="http://treatment.plazi.org/id/20230174E5DFBEFA9E27B22779B10D07" lastPageId="30" lastPageNumber="117" pageId="26" pageNumber="113">
<subSubSection pageId="26" pageNumber="113" type="multiple">
<paragraph pageId="26" pageNumber="113">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
<treatment id="20230174E5DFBEFA9E27B22779B10D07" ID-GBIF-Taxon="127885602" LSID="urn:lsid:plazi:treatment:20230174E5DFBEFA9E27B22779B10D07" httpUri="http://treatment.plazi.org/id/20230174E5DFBEFA9E27B22779B10D07" lastPageId="30" lastPageNumber="117" pageId="26" pageNumber="113" scope_family="Chrysomelidae" scope_order="Coleoptera" scope_subFamily="Chrysomelinae">
<subSubSection id="38871C72AEAE01E0DC667D640C44ED5C" pageId="26" pageNumber="113" type="multiple">
<paragraph id="B931F4BB9272EB1A96C13072313E2AA9" pageId="26" pageNumber="113">Taxon classification Animalia Coleoptera Chrysomelidae</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="113" type="nomenclature">
<paragraph pageId="26" pageNumber="113">
<taxonomicName authority="Bechyne, 1948" authorityName="Bechyne" authorityYear="1948" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="26" pageNumber="113" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">
<subSubSection id="762B5A19F996ED8B5A9F3A6B11BABD33" pageId="26" pageNumber="113" type="nomenclature">
<paragraph id="9B0CDCE3EE6EC44AC862BA41717F87DE" pageId="26" pageNumber="113">
<taxonomicName id="A0CE379CA5DB65B0BA9C331EA8FF3AD4" authority="Bechyne, 1948" authorityName="Bechyne" authorityYear="1948" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="26" pageNumber="113" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">
Gonioctena (Gonioctena) springlovae (
<normalizedToken originalValue="Bechyně">Bechyne</normalizedToken>
<normalizedToken id="AAE9F070FE77D379A67A9FAA67D533DD" originalValue="Bechyně">Bechyne</normalizedToken>
, 1948)
</taxonomicName>
Figs 11-12, 16, 63-65
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="114" type="reference_group">
<paragraph pageId="27" pageNumber="114">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae">
<pageBreakToken pageId="27" pageNumber="114" start="start">Phytodecta</pageBreakToken>
<subSubSection id="D565F212052EC9B43835B9806B00B917" pageId="27" pageNumber="114" type="reference_group">
<paragraph id="0DE812291E7F30365F554837EFD0877E" pageId="27" pageNumber="114">
<taxonomicName id="1ABB5E8C3D584B1C14D43957ECE1C09E" genus="Phytodecta" lsidName="Phytodecta springlovae" pageId="27" pageNumber="114" rank="species" species="springlovae">
<pageBreakToken id="0584F89478C6EEA65F9AF537B6EAF898" pageId="27" pageNumber="114" start="start">Phytodecta</pageBreakToken>
springlovae
</taxonomicName>
<normalizedToken originalValue="Bechyně">Bechyne</normalizedToken>
<normalizedToken id="53B42BC3B7C33E5C78715E38CD6E84C3" originalValue="Bechyně">Bechyne</normalizedToken>
, 1948: 115, 116 (type locality: Japonia, Kioto).
</paragraph>
<paragraph pageId="27" pageNumber="114">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">Gonioctena (Gonioctena) springlovae</taxonomicName>
<paragraph id="5393FBCB88958FCB6821940B6CC82718" pageId="27" pageNumber="114">
<taxonomicName id="BAAB7B2917E6E03ACAD7A80D9EB0C86B" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">Gonioctena (Gonioctena) springlovae</taxonomicName>
:
<bibRefCitation author="Chujo, M" journalOrPublisher="Niponius, Acta Coleopterologica" pageId="31" pageNumber="118" pagination="1 - 10" title="Descriptions of three new genera and a new species of chrysomelid-beetles from Japan, with some notes on the Japanese species." volume="1" year="1960">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
<bibRefCitation id="8FC781A5EF5126E21DA116FFEA800CAB" author="Chujo, M" journalOrPublisher="Niponius, Acta Coleopterologica" pageId="31" pageNumber="118" pagination="1 - 10" title="Descriptions of three new genera and a new species of chrysomelid-beetles from Japan, with some notes on the Japanese species." volume="1" year="1960">
<normalizedToken id="D1838FFD2DD60BB3B7875F2E90A3C0DC" originalValue="Chûjô">Chujo</normalizedToken>
and Kimoto 1960
</bibRefCitation>
: 5,
<bibRefCitation author="Chujo, M" journalOrPublisher="Pacific Insects" pageId="31" pageNumber="118" pagination="117 - 202" title="Systematic catalog of Japanese Chrysomelidae (Coleoptera)." volume="3" year="1961">1961</bibRefCitation>
<bibRefCitation id="8AB6DE3183255E7F34F52CB47A6DA1E0" author="Chujo, M" journalOrPublisher="Pacific Insects" pageId="31" pageNumber="118" pagination="117 - 202" title="Systematic catalog of Japanese Chrysomelidae (Coleoptera)." volume="3" year="1961">1961</bibRefCitation>
: 153;
<bibRefCitation author="Gressitt, JL" journalOrPublisher="Pacific Insects Monograph" pageId="32" pageNumber="119" pagination="301 - 1026" title="The Chrysomelidae (Coleopt.) of China and Korea, Part 2." volume="1 B" year="1963">Gressitt and Kimoto 1963</bibRefCitation>
<bibRefCitation id="E18947E0BBECD7D9CE5C9756B4338A90" author="Gressitt, JL" journalOrPublisher="Pacific Insects Monograph" pageId="32" pageNumber="119" pagination="301 - 1026" title="The Chrysomelidae (Coleopt.) of China and Korea, Part 2." volume="1 B" year="1963">Gressitt and Kimoto 1963</bibRefCitation>
: 358, 362;
<bibRefCitation author="Kimoto, S" journalOrPublisher="Fragmenta Coleopterologica" pageId="32" pageNumber="119" pagination="13 - 18" title="Descriptions of new species of the Chrysomelidae from Japan, with notes on some known species." volume="3 / 4" year="1963">Kimoto 1963</bibRefCitation>
<bibRefCitation id="8FE8AD75D8B6A53AA93EDEDB039E250D" author="Kimoto, S" journalOrPublisher="Fragmenta Coleopterologica" pageId="32" pageNumber="119" pagination="13 - 18" title="Descriptions of new species of the Chrysomelidae from Japan, with notes on some known species." volume="3 / 4" year="1963">Kimoto 1963</bibRefCitation>
: 17,
<bibRefCitation author="Kimoto, S" journalOrPublisher="Journal of the Faculty of Agriculture, Kyushu University" pageId="32" pageNumber="119" pagination="263 - 286" title="The Chrysomelidae of Japan and the Ryukyu Islands V. Subfamily Chrysomelinae." volume="13" year="1964">1964</bibRefCitation>
<bibRefCitation id="DF6E87164D472B0084BDE61ABC735779" author="Kimoto, S" journalOrPublisher="Journal of the Faculty of Agriculture, Kyushu University" pageId="32" pageNumber="119" pagination="263 - 286" title="The Chrysomelidae of Japan and the Ryukyu Islands V. Subfamily Chrysomelinae." volume="13" year="1964">1964</bibRefCitation>
: 280, 282;
<bibRefCitation author="Takizawa, H" journalOrPublisher="Kontyu" pageId="36" pageNumber="123" pagination="444 - 468" title="Larvae of the genus Gonioctena Chevrolat (Coleoptera, Chrysomelidae): descriptions of Japanese species and the implications of larval characters for the phylogeny." volume="44" year="1976">Takizawa 1976</bibRefCitation>
<bibRefCitation id="5B53B7257D8AA2473A413F0B505B1276" author="Takizawa, H" journalOrPublisher="Kontyu" pageId="36" pageNumber="123" pagination="444 - 468" title="Larvae of the genus Gonioctena Chevrolat (Coleoptera, Chrysomelidae): descriptions of Japanese species and the implications of larval characters for the phylogeny." volume="44" year="1976">Takizawa 1976</bibRefCitation>
: 454 (larva, pupa, biology); L. N.
<bibRefCitation author="Medvedev, LN" editor="Ler, PA" journalOrPublisher="Nauka, St. Petersburg" pageId="33" pageNumber="120" pagination="533 - 602" title="Family Chrysomelidae." volumeTitle="Opredelitel' Nasekomykh Dal'nego Vostoka SSSR Tom 3 - Zheskokrylye, ili zhuki Chast 2" year="1992">Medvedev 1992</bibRefCitation>
<bibRefCitation id="53189531833FD2E3D17357A0219021A7" author="Medvedev, LN" editor="Ler, PA" journalOrPublisher="Nauka, St. Petersburg" pageId="33" pageNumber="120" pagination="533 - 602" title="Family Chrysomelidae." volumeTitle="Opredelitel' Nasekomykh Dal'nego Vostoka SSSR Tom 3 - Zheskokrylye, ili zhuki Chast 2" year="1992">Medvedev 1992</bibRefCitation>
: 575 (as synonym of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena affinis" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="affinis">Gonioctena affinis</taxonomicName>
<taxonomicName id="911BCFF6F2D881ADD2B7CC2E872C6712" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena affinis" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="affinis">Gonioctena affinis</taxonomicName>
);
<bibRefCitation author="Kimoto, S" journalOrPublisher="Tokai University Press, Tokyo" pageId="32" pageNumber="119" title="Leaf beetles (Chrysomelidae) of Japan." year="1994">Kimoto and Takizawa 1994</bibRefCitation>
<bibRefCitation id="28B03FDF7DBEA0B1BDAE1810F1D781B6" author="Kimoto, S" journalOrPublisher="Tokai University Press, Tokyo" pageId="32" pageNumber="119" title="Leaf beetles (Chrysomelidae) of Japan." year="1994">Kimoto and Takizawa 1994</bibRefCitation>
: 139, 229, 302, 452, 498 (incl. larva and pupa); V. L.
<bibRefCitation author="Medvedev, VL" journalOrPublisher="Russian Entomological Journal" pageId="34" pageNumber="121" pagination="41 - 42" title="New species of the genus Gonioctena Chevrolat, 1837 (Coleoptera: Chrysomelidae) from China." volume="13" year="2004">Medvedev 2004</bibRefCitation>
<bibRefCitation id="792F46F918FC48625C8D78F18D7632F2" author="Medvedev, VL" journalOrPublisher="Russian Entomological Journal" pageId="34" pageNumber="121" pagination="41 - 42" title="New species of the genus Gonioctena Chevrolat, 1837 (Coleoptera: Chrysomelidae) from China." volume="13" year="2004">Medvedev 2004</bibRefCitation>
: 41;
<bibRefCitation author="Takizawa, H" journalOrPublisher="Insecta Matsumurana, new series" pageId="36" pageNumber="123" pagination="35 - 50" title="A revision of the genus Gonioctena Chevrolat in Japan (Coleoptera: Chrysomelidae)." volume="63" year="2007">Takizawa 2007</bibRefCitation>
<bibRefCitation id="188D8BCCFD5E83A469A7ECA2CFB6ED0C" author="Takizawa, H" journalOrPublisher="Insecta Matsumurana, new series" pageId="36" pageNumber="123" pagination="35 - 50" title="A revision of the genus Gonioctena Chevrolat in Japan (Coleoptera: Chrysomelidae)." volume="63" year="2007">Takizawa 2007</bibRefCitation>
: 38, 42;
<bibRefCitation author="Kippenberg, H" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="32" pageNumber="119" pagination="67 - 73" title="New acts and comments, Chrysomelidae: Chrysomelinae." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 6, Chrysomeloidea" year="2010">Kippenberg 2010</bibRefCitation>
<bibRefCitation id="405984A82F1B0A4FD48207135DAD9AE0" author="Kippenberg, H" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="32" pageNumber="119" pagination="67 - 73" title="New acts and comments, Chrysomelidae: Chrysomelinae." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 6, Chrysomeloidea" year="2010">Kippenberg 2010</bibRefCitation>
: 434.
</paragraph>
<paragraph pageId="27" pageNumber="114">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae">Gonioctena springlovae</taxonomicName>
<paragraph id="F5E2F7465CE3D2984731ABBBC36E9046" pageId="27" pageNumber="114">
<taxonomicName id="9472E76A1259B605206B60A3FA053009" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae">Gonioctena springlovae</taxonomicName>
:
<bibRefCitation author="Takizawa, H" journalOrPublisher="Kontyu" pageId="35" pageNumber="122" pagination="172 - 176" title="A list of chrysomelid beetles from Sakhalin in the collection of the Entomological Institute, Hokkaido University (Coleoptera)." volume="39" year="1971">Takizawa 1971</bibRefCitation>
<bibRefCitation id="5A2C7EBDE98948820837041BA0C1CA5C" author="Takizawa, H" journalOrPublisher="Kontyu" pageId="35" pageNumber="122" pagination="172 - 176" title="A list of chrysomelid beetles from Sakhalin in the collection of the Entomological Institute, Hokkaido University (Coleoptera)." volume="39" year="1971">Takizawa 1971</bibRefCitation>
: 173;
<bibRefCitation author="Cox, ML" editor="Jolivet, PHA" journalOrPublisher="SPB Academic Publishing, Amsterdam" pageId="31" pageNumber="118" pagination="119 - 265" title="The Pupae of Chrysomeloidea." volumeTitle="Chrysomelidae Biology. Vol. 1: The Classification, Phylogeny and Genetics" year="1996">Cox 1996</bibRefCitation>
<bibRefCitation id="0C6E940253D19CA3896380FA2CBB6DFB" author="Cox, ML" editor="Jolivet, PHA" journalOrPublisher="SPB Academic Publishing, Amsterdam" pageId="31" pageNumber="118" pagination="119 - 265" title="The Pupae of Chrysomeloidea." volumeTitle="Chrysomelidae Biology. Vol. 1: The Classification, Phylogeny and Genetics" year="1996">Cox 1996</bibRefCitation>
: 146 (pupa);
<bibRefCitation author="Kudo, S" editor="Jolivet, P" journalOrPublisher="SPB Academic Publishing, The Hague" pageId="33" pageNumber="120" pagination="727 - 738" title="Diversified reproductive strategies in Gonioctena (Chrysomelinae) leaf beetles." volumeTitle="New Developments in the Biology of Chrysomelidae" year="2003">Kudo and Hasegawa 2003</bibRefCitation>
<bibRefCitation id="3A5661E3A7C4EE998A81AA0E7764A73A" author="Kudo, S" editor="Jolivet, P" journalOrPublisher="SPB Academic Publishing, The Hague" pageId="33" pageNumber="120" pagination="727 - 738" title="Diversified reproductive strategies in Gonioctena (Chrysomelinae) leaf beetles." volumeTitle="New Developments in the Biology of Chrysomelidae" year="2003">Kudo and Hasegawa 2003</bibRefCitation>
: 729 (biology);
<bibRefCitation author="Takahashi, S" editor="Shiyake, S" journalOrPublisher="Osaka Museum of Natural History, Osaka" pageId="35" pageNumber="122" pagination="239 - 372" title="Chrysomelidae (exclusive of Bruchinae &amp; Donaciinae)." volumeTitle="Specimen list of Coleoptera in the Insect Collection of the Osaka Museum of Natural History (Part 2)" year="2012">Takahashi 2012</bibRefCitation>
<bibRefCitation id="ED1DE60EEC7CABAB89088330FA513463" author="Takahashi, S" editor="Shiyake, S" journalOrPublisher="Osaka Museum of Natural History, Osaka" pageId="35" pageNumber="122" pagination="239 - 372" title="Chrysomelidae (exclusive of Bruchinae &amp; Donaciinae)." volumeTitle="Specimen list of Coleoptera in the Insect Collection of the Osaka Museum of Natural History (Part 2)" year="2012">Takahashi 2012</bibRefCitation>
: 289.
</paragraph>
<paragraph pageId="27" pageNumber="114">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta gracilicornis" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="gracilicornis">Phytodecta gracilicornis</taxonomicName>
<paragraph id="FE350CDB952A4427E64B385179857FB5" pageId="27" pageNumber="114">
<taxonomicName id="B527EC89613846EDC926B1224B0EEFF7" genus="Phytodecta" lsidName="Phytodecta gracilicornis" pageId="27" pageNumber="114" rank="species" species="gracilicornis">Phytodecta gracilicornis</taxonomicName>
?:
<bibRefCitation author="Jacoby, M" journalOrPublisher="Proceedings of the Scientific Meetings of the Zoological Society of London" pageId="32" pageNumber="119" pagination="190 - 211" title="Descriptions of the phytophagous Coleoptera of Japan, obtained by Mr. George Lewis during his Second Journey, from February 1880 to September 1881. Part I." url="10.1111/j.1096-3642.1885.tb02894.x" volume="1885" year="1885">Jacoby 1885</bibRefCitation>
<bibRefCitation id="D112EAB14B1EBC50B78CCC60DBF44E09" author="Jacoby, M" journalOrPublisher="Proceedings of the Scientific Meetings of the Zoological Society of London" pageId="32" pageNumber="119" pagination="190 - 211" title="Descriptions of the phytophagous Coleoptera of Japan, obtained by Mr. George Lewis during his Second Journey, from February 1880 to September 1881. Part I." url="10.1111/j.1096-3642.1885.tb02894.x" volume="1885" year="1885">Jacoby 1885</bibRefCitation>
: 210 (misidentification).
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="114" type="type material">
<paragraph pageId="27" pageNumber="114">Type material.</paragraph>
<paragraph pageId="27" pageNumber="114">
<subSubSection id="1A5F3D136E921B724D4A38FAC94F77F4" pageId="27" pageNumber="114" type="type material">
<paragraph id="41C99BC64C057365036BAD1A5CDE75B0" pageId="27" pageNumber="114">Type material.</paragraph>
<paragraph id="2986A74DB805C595452B3279EDC65A2F" pageId="27" pageNumber="114">
Holotype: ♂ (NMPC), Japon, Kioto // TYPUS //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phytodecta" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="genus">Phytodecta</taxonomicName>
<taxonomicName id="857F531081DDF01A27ED3873EE10DC47" genus="Phytodecta" lsidName="Phytodecta" pageId="27" pageNumber="114" rank="genus">Phytodecta</taxonomicName>
TYPE
<taxonomicName lsidName="springlovae" pageId="27" pageNumber="114" rank="species" species="springlovae">springlovae</taxonomicName>
<taxonomicName id="BC2CA2CE5087AC46EC18E4464168AB53" lsidName="springlovae" pageId="27" pageNumber="114" rank="species" species="springlovae">springlovae</taxonomicName>
n. sp. 1945 Det. J.
<normalizedToken originalValue="Bechyně">Bechyne</normalizedToken>
<normalizedToken id="F67C818027977280AD9BEC224B8B6ABC" originalValue="Bechyně">Bechyne</normalizedToken>
. // Coll. Achard Mus. Pragense // Mus. Nat. Pragae Inv. 18 960 //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">Gonioctena (Gonioctena) springlovae</taxonomicName>
<taxonomicName id="4F2991C48C3E3A30DE74B019A6E4D5D8" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">Gonioctena (Gonioctena) springlovae</taxonomicName>
(
<normalizedToken originalValue="Bechyně">Bechyne</normalizedToken>
<normalizedToken id="F8E1D154C6A9274B973F79A7F3755972" originalValue="Bechyně">Bechyne</normalizedToken>
) Det. S. GE 2004.
</paragraph>
</subSubSection>
<subSubSection lastPageId="28" lastPageNumber="115" pageId="27" pageNumber="114" type="materials_examined">
<paragraph pageId="27" pageNumber="114">Other material.</paragraph>
<paragraph lastPageId="28" lastPageNumber="115" pageId="27" pageNumber="114">
<subSubSection id="DFE362D92AB82411E27A298747C025E0" lastPageId="28" lastPageNumber="115" pageId="27" pageNumber="114" type="materials_examined">
<paragraph id="04B5B04D2B659E7FDC70B044EF8EA19D" pageId="27" pageNumber="114">Other material.</paragraph>
<paragraph id="F6FB585CC10C489F4BC9250CA471DDFD" lastPageId="28" lastPageNumber="115" pageId="27" pageNumber="114">
Japan (Hokkaido): 1♂ (NMPC), Japon, Kioto // TYPUS //
<taxonomicName aberration="graduata" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" infraspecific-rank="ab." kingdom="Animalia" lsidName="Phytodecta springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="aberration" species="springlovae">Phytodecta springlovae ab. graduata</taxonomicName>
<taxonomicName id="CB650AC0C3BF6326FF9A8B0E5E6F4EA6" aberration="graduata" genus="Phytodecta" infraspecific-rank="ab." lsidName="Phytodecta springlovae" pageId="27" pageNumber="114" rank="aberration" species="springlovae">Phytodecta springlovae ab. graduata</taxonomicName>
n. ab. TYPE 1945 Det. J.
<normalizedToken originalValue="Bechyně">Bechyne</normalizedToken>
<normalizedToken id="739104298F6C72E6D1C485EA6F4605C3" originalValue="Bechyně">Bechyne</normalizedToken>
. //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">Gonioctena (Gonioctena) springlovae</taxonomicName>
<taxonomicName id="4436D8FF716D1AC2F186B34751BAD7FA" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">Gonioctena (Gonioctena) springlovae</taxonomicName>
(
<normalizedToken originalValue="Bechyně">Bechyne</normalizedToken>
<normalizedToken id="E4E5D6A03A1C26F9BDEA955FF00FD3C0" originalValue="Bechyně">Bechyne</normalizedToken>
) Det. S. GE 2004; 1♂ (NMPC), Japon, Kioto // coll. Achard Mus. Pragense // TYPUS //
<taxonomicName aberration="graduata" class="Insecta" family="Chrysomelidae" genus="Phytodecta" higherTaxonomySource="GBIF" infraspecific-rank="ab." kingdom="Animalia" lsidName="Phytodecta springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="aberration" species="springlovae">Phytodecta springlovae ab. graduata</taxonomicName>
<taxonomicName id="5992009859F151E3FC1F3E7C741DE430" aberration="graduata" genus="Phytodecta" infraspecific-rank="ab." lsidName="Phytodecta springlovae" pageId="27" pageNumber="114" rank="aberration" species="springlovae">Phytodecta springlovae ab. graduata</taxonomicName>
n. ab. PARATYPE 1945 Det. J.
<normalizedToken originalValue="Bechyně">Bechyne</normalizedToken>
<normalizedToken id="D67A959D482013EAF609F201E05D3439" originalValue="Bechyně">Bechyne</normalizedToken>
. // Mus. Nat. Pragae Inv. 18 961 //
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">Gonioctena (Gonioctena) springlovae</taxonomicName>
<taxonomicName id="F8DBA4886389AD48142786E542F1CB90" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena (Gonioctena) springlovae" order="Coleoptera" pageId="27" pageNumber="114" phylum="Arthropoda" rank="species" species="springlovae" subGenus="Gonioctena">Gonioctena (Gonioctena) springlovae</taxonomicName>
(
<normalizedToken originalValue="Bechyně">Bechyne</normalizedToken>
<normalizedToken id="D2F777B2D295FD0B9CACFA247F353AF7" originalValue="Bechyně">Bechyne</normalizedToken>
) Det. S. GE 2004; 3♂♂ (JBC), Japan, Hokkaido, Eniwa Mt. 30 km S from Sapporo, Shikotsu-Toya N.P. 500 m, 6.VII.1997, lgt. V. Kostal; 1♀ (BMNH), Japan, G. Lewis. 1910-320. // Chiuzenji; 5♂♂, 1♀ (HCC), Japan, Hokkaido, Sapporo, Kannon-zawa, 29.V.1995, S. Kudo; 1♂, 1♀ (HCC), Japan, Hokkaido, Sapporo, Jozankei, 29.VIII.2011, H. Suenaga leg.; 1♂ (SEHU), Jozankei, Hokkaido, 22.VII.1955, M. Konishi; 1♂, 1♀ (SEHU), Hidaka, Hokkaido, 1955, S. Watanabe; 1♂ (AWC), JAPAN, Takahiro Parking Area, Bifuka, Hokkaido, 4.VII.2002, Y. Komiya lgt. Russia (Sakhalin): 1♂ (NHMB), Sakhalin, riv. Naiba, VIII.1991; 2♂♂ (BMNH), Russia,
<pageBreakToken pageId="28" pageNumber="115" start="start">Saghalien</pageBreakToken>
<pageBreakToken id="1D5194AA3372C2105B7CB53AFD183925" pageId="28" pageNumber="115" start="start">Saghalien</pageBreakToken>
, Central Expt. Sta.; 1♂ (SEHU), Saghalien, 16.VII.1933, Uchida, Okada, Sawamoto &amp; Hoye legs; 4♂♂ (ZIN), Russia, Sakhalin Oblast, Sakhalin Island, Holmsky pass (approx. 47°N, 142°E), 1.VII.1982, Smirnov leg.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="115" type="diagnosis">
<paragraph pageId="28" pageNumber="115">Diagnosis.</paragraph>
<paragraph pageId="28" pageNumber="115">
<subSubSection id="055878EC3035660593B75F98126FBA12" pageId="28" pageNumber="115" type="diagnosis">
<paragraph id="6388EEA790EB4C8085777354C95CBD7A" pageId="28" pageNumber="115">Diagnosis.</paragraph>
<paragraph id="7E3167577DF17AFF9AACCE3E4008C80A" pageId="28" pageNumber="115">
See diagnosis of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena gracilicornis" order="Coleoptera" pageId="28" pageNumber="115" phylum="Arthropoda" rank="species" species="gracilicornis">Gonioctena gracilicornis</taxonomicName>
<taxonomicName id="B6820E318A2DA5B8D6CBC5A5A88F7D8C" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena gracilicornis" order="Coleoptera" pageId="28" pageNumber="115" phylum="Arthropoda" rank="species" species="gracilicornis">Gonioctena gracilicornis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="29" lastPageNumber="116" pageId="28" pageNumber="115" type="redescription">
<paragraph pageId="28" pageNumber="115">Redescription.</paragraph>
<paragraph lastPageId="29" lastPageNumber="116" pageId="28" pageNumber="115">
<subSubSection id="AD4D8A67FDA36834C6EDFFA3113A58D4" lastPageId="29" lastPageNumber="116" pageId="28" pageNumber="115" type="redescription">
<paragraph id="446D9AD1C1F09DF6219D5B792AD8F140" pageId="28" pageNumber="115">Redescription.</paragraph>
<paragraph id="A62DDBC92335D1D0151CB391F531333F" lastPageId="29" lastPageNumber="116" pageId="28" pageNumber="115">
Measurements in mm (n = 5): length of body: 5.70-6.20 (mean 6.00); width of body: 3.20-3.60 (mean 3.40); height of body: 2.30-2.50 (mean 2.42); width of head: 1.60-1.75 (mean 1.66); interocular distance: 1.10-1.20 (mean 1.13); width of apex of pronotum: 1.87-2.05 (mean 1.95); width of base of pronotum: 2.70-3.05 (mean 2.84); maximum width of pronotum: 2.72-3.07 (mean 2.87); length of
<pageBreakToken pageId="29" pageNumber="116" start="start">pronotum</pageBreakToken>
<pageBreakToken id="E1B802DED6DB465BC38CF4B96B9179B7" pageId="29" pageNumber="116" start="start">pronotum</pageBreakToken>
along midline: 1.35-1.50 (mean 1.41); length of elytra along suture: 4.10-4.60 (mean 4.42).
</paragraph>
<paragraph pageId="29" pageNumber="116">Body oblong oval and moderately convex (Fig. 11). Coloration variable. Head black, with dark reddish brown band near apex of mandibles. Antennomeres 1-5 yellowish brown, generally darkened, 6-7 dark brown to blackish brown, 8-11 black. Pronotum entirely black. Scutellum black. Elytra reddish brown or yellowish brown, with 5 pairs of black spots, generally connected with each other, rarely elytra entirely black (Fig. 65). Venter black, with lateral margins of last abdominal ventrite reddish brown. Legs black, with tarsi blackish brown, sometimes tibiae largely dark brown to reddish brown.</paragraph>
<caption pageId="29" pageNumber="116">
<paragraph pageId="29" pageNumber="116">
<paragraph id="05EFDFC08B10B96B9B1BE7952ABBC69E" pageId="29" pageNumber="116">Body oblong oval and moderately convex (Fig. 11). Coloration variable. Head black, with dark reddish brown band near apex of mandibles. Antennomeres 1-5 yellowish brown, generally darkened, 6-7 dark brown to blackish brown, 8-11 black. Pronotum entirely black. Scutellum black. Elytra reddish brown or yellowish brown, with 5 pairs of black spots, generally connected with each other, rarely elytra entirely black (Fig. 65). Venter black, with lateral margins of last abdominal ventrite reddish brown. Legs black, with tarsi blackish brown, sometimes tibiae largely dark brown to reddish brown.</paragraph>
<caption id="CDF2A7588CDD38B2E8E87AA7C8F85B2B" pageId="29" pageNumber="116">
<paragraph id="CBA56FEC5767A4B63463CBD989D6F4D3" pageId="29" pageNumber="116">
Figures 63-65.
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena springlovae" order="Coleoptera" pageId="29" pageNumber="116" phylum="Arthropoda" rank="species" species="springlovae">Gonioctena springlovae</taxonomicName>
<taxonomicName id="75214B64C16D1F1B6817FEADC29C8C1D" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena springlovae" order="Coleoptera" pageId="29" pageNumber="116" phylum="Arthropoda" rank="species" species="springlovae">Gonioctena springlovae</taxonomicName>
. 63 Antenna (♂, ♀) 64 Aedeagus 65 Color variation. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption pageId="29" pageNumber="116">
<paragraph pageId="29" pageNumber="116">
<caption id="D5DAC5A53A81A326326B9D2460399DF1" pageId="29" pageNumber="116">
<paragraph id="403836DAE4EF5086187E74F777278020" pageId="29" pageNumber="116">
Figures 66-67. Ovoviviparous species. 66 Newly laid larvae of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena gracilicornis" order="Coleoptera" pageId="29" pageNumber="116" phylum="Arthropoda" rank="species" species="gracilicornis">Gonioctena gracilicornis</taxonomicName>
<taxonomicName id="1FE1C8A5CE757881025051901D5C8DD8" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena gracilicornis" order="Coleoptera" pageId="29" pageNumber="116" phylum="Arthropoda" rank="species" species="gracilicornis">Gonioctena gracilicornis</taxonomicName>
67 Larvae dissected from a female of
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena nivosa" order="Coleoptera" pageId="29" pageNumber="116" phylum="Arthropoda" rank="species" species="nivosa">Gonioctena nivosa</taxonomicName>
<taxonomicName id="2B30626FAA30DD969A2F0966BE13C458" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena nivosa" order="Coleoptera" pageId="29" pageNumber="116" phylum="Arthropoda" rank="species" species="nivosa">Gonioctena nivosa</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="29" pageNumber="116">Head. Vertex weakly convex, covered with sparse punctures, becoming coarser and denser toward sides. Frontal suture V-shaped, coronal suture absent. Frons flat, strongly depressed anteriorly, covered with moderately dense punctures. Clypeus narrow and trapezoidal. Anterior margin of labrum distinctly concave. Mandibles with 2 sharp apical teeth and a deep excavation for apical maxillary palpomere at outer side. Maxillary palps 4-segmented, with apical palpomere distinctly widened, truncate apically in male; slightly widened in female. Antennae in male longer than half length of body; antennomere 1 robust; antennomere 2 shorter than 3; antennomere 3 longer than 4; antennomeres 7-11 elongate; antennomere 11 longest, about 3.95 times as long as wide (Fig. 63). Antennae in female reaching elytral humeri; antennomere 11 about 2.58 times as long as wide.</paragraph>
<paragraph pageId="29" pageNumber="116">Pronotum. Lateral sides widest near base, feebly rounded, slightly narrowed anteriorly, anterior angles strongly produced (Fig. 12). Anterior and lateral margins bordered, lateral margins well visible in dorsal view. Trichobothria present on posterior angles. Disc covered with sparse punctures; lateral sides covered with much coarser and denser punctures, becoming larger toward base, partially confluent near basal margin; interspaces covered with fine and sparse punctures. Scutellum variable in length, as long as wide, longer than wide or wider than long.</paragraph>
<paragraph pageId="29" pageNumber="116">Elytra. Lateral sides slightly widened posteriorly, widest beyond middle, thence roundly narrowed posteriorly. Humeral calli well developed. Disc covered with 11 regular rows of large punctures, including a short scutellar row; interspaces shagreened, covered with fine and sparse punctures. Epipleura wholly visible in lateral view. Hind wings well developed.</paragraph>
<paragraph pageId="29" pageNumber="116">Venter. Hypomera weakly rugose, with dense punctures on anterior side. Prosternum covered with coarse and dense punctures bearing long setae; prosternal process enlarged apically, bordered laterally, with sparse punctures. Metasternum covered with small and sparse punctures in median region, large and dense punctures in lateral region. Abdominal ventrites covered with dense punctures bearing short setae.</paragraph>
<paragraph pageId="29" pageNumber="116">Legs. Moderately robust. Tibiae widened apically, with a tooth-like projection. Fore legs with tarsomere 1 enlarged, slightly wider than 3 in male; distinctly narrower than 3 in female. Tarsal claws appendiculate.</paragraph>
<paragraph pageId="29" pageNumber="116">Genitalia. Aedeagus thin, distinctly narrowed apically, with apical process thin in dorsal view; moderately curved, with apical process pointed and slightly bent downward at apex in lateral view (Fig. 64). Spermatheca absent.</paragraph>
<paragraph id="EC3342439B1F09CDFA6788B567326B62" pageId="29" pageNumber="116">Head. Vertex weakly convex, covered with sparse punctures, becoming coarser and denser toward sides. Frontal suture V-shaped, coronal suture absent. Frons flat, strongly depressed anteriorly, covered with moderately dense punctures. Clypeus narrow and trapezoidal. Anterior margin of labrum distinctly concave. Mandibles with 2 sharp apical teeth and a deep excavation for apical maxillary palpomere at outer side. Maxillary palps 4-segmented, with apical palpomere distinctly widened, truncate apically in male; slightly widened in female. Antennae in male longer than half length of body; antennomere 1 robust; antennomere 2 shorter than 3; antennomere 3 longer than 4; antennomeres 7-11 elongate; antennomere 11 longest, about 3.95 times as long as wide (Fig. 63). Antennae in female reaching elytral humeri; antennomere 11 about 2.58 times as long as wide.</paragraph>
<paragraph id="FB64D28A369D5F1951E8076D45884E15" pageId="29" pageNumber="116">Pronotum. Lateral sides widest near base, feebly rounded, slightly narrowed anteriorly, anterior angles strongly produced (Fig. 12). Anterior and lateral margins bordered, lateral margins well visible in dorsal view. Trichobothria present on posterior angles. Disc covered with sparse punctures; lateral sides covered with much coarser and denser punctures, becoming larger toward base, partially confluent near basal margin; interspaces covered with fine and sparse punctures. Scutellum variable in length, as long as wide, longer than wide or wider than long.</paragraph>
<paragraph id="9B8805BD92A52A4CBF5B7A26A3B3F4D6" pageId="29" pageNumber="116">Elytra. Lateral sides slightly widened posteriorly, widest beyond middle, thence roundly narrowed posteriorly. Humeral calli well developed. Disc covered with 11 regular rows of large punctures, including a short scutellar row; interspaces shagreened, covered with fine and sparse punctures. Epipleura wholly visible in lateral view. Hind wings well developed.</paragraph>
<paragraph id="B49E9B86FD393D2A3FB08899C892744B" pageId="29" pageNumber="116">Venter. Hypomera weakly rugose, with dense punctures on anterior side. Prosternum covered with coarse and dense punctures bearing long setae; prosternal process enlarged apically, bordered laterally, with sparse punctures. Metasternum covered with small and sparse punctures in median region, large and dense punctures in lateral region. Abdominal ventrites covered with dense punctures bearing short setae.</paragraph>
<paragraph id="87197A2CBA8278592D7AD9C88BF02E09" pageId="29" pageNumber="116">Legs. Moderately robust. Tibiae widened apically, with a tooth-like projection. Fore legs with tarsomere 1 enlarged, slightly wider than 3 in male; distinctly narrower than 3 in female. Tarsal claws appendiculate.</paragraph>
<paragraph id="909B92123D8456EDE8FE9B65EE3414A4" pageId="29" pageNumber="116">Genitalia. Aedeagus thin, distinctly narrowed apically, with apical process thin in dorsal view; moderately curved, with apical process pointed and slightly bent downward at apex in lateral view (Fig. 64). Spermatheca absent.</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="117" type="distribution">
<paragraph pageId="30" pageNumber="117">
<pageBreakToken pageId="30" pageNumber="117" start="start">Distribution</pageBreakToken>
<subSubSection id="489BDC474410B242E2B8F2D245CD5E99" pageId="30" pageNumber="117" type="distribution">
<paragraph id="9A03612F0AE67B328D51E4AED74D050F" pageId="30" pageNumber="117">
<pageBreakToken id="3EBF74370F0A659DFC087FF9ED3AAB61" pageId="30" pageNumber="117" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="30" pageNumber="117">Russia (Sakhalin), Japan (Hokkaido) (Fig. 16).</paragraph>
<paragraph id="624991AC114FC341329759CCC48ECC86" pageId="30" pageNumber="117">Russia (Sakhalin), Japan (Hokkaido) (Fig. 16).</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="117" type="host">
<paragraph pageId="30" pageNumber="117">Host plant.</paragraph>
<paragraph pageId="30" pageNumber="117">
<taxonomicName family="Salicaceae" lsidName="" pageId="30" pageNumber="117" rank="family">Salicaceae</taxonomicName>
<subSubSection id="30198B9351150AEA6B81A6A4E2F84B4D" pageId="30" pageNumber="117" type="host">
<paragraph id="331BF2A5333DC8F5D9DB7E44ABC1C263" pageId="30" pageNumber="117">Host plant.</paragraph>
<paragraph id="AB0BE857CA4DF627AB8A1726486D313B" pageId="30" pageNumber="117">
<taxonomicName id="3CCAC6F8F4CAF1D650ED06853A14E2BB" class="Magnoliopsida" family="Salicaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Malpighiales" pageId="30" pageNumber="117" phylum="Tracheophyta" rank="family">Salicaceae</taxonomicName>
:
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="30" pageNumber="117" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
<taxonomicName id="41B337C233B24F33210B5FA7C366D7BA" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus" order="Malpighiales" pageId="30" pageNumber="117" phylum="Tracheophyta" rank="genus">Populus</taxonomicName>
spp.,
<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="30" pageNumber="117" phylum="Chordata" rank="genus">Salix</taxonomicName>
<taxonomicName id="3EA98BD6AC60933E1A8D9C1A9E8BB7F9" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="30" pageNumber="117" phylum="Chordata" rank="genus">Salix</taxonomicName>
spp. (
<bibRefCitation author="Chujo, M" journalOrPublisher="Pacific Insects" pageId="31" pageNumber="118" pagination="117 - 202" title="Systematic catalog of Japanese Chrysomelidae (Coleoptera)." volume="3" year="1961">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
<bibRefCitation id="9061A439CBF79B20FD8E456B6E9CF9DD" author="Chujo, M" journalOrPublisher="Pacific Insects" pageId="31" pageNumber="118" pagination="117 - 202" title="Systematic catalog of Japanese Chrysomelidae (Coleoptera)." volume="3" year="1961">
<normalizedToken id="13FFD5392F679CC83D5CD1BC5A1E3FF7" originalValue="Chûjô">Chujo</normalizedToken>
and Kimoto 1961
</bibRefCitation>
,
<bibRefCitation author="Kimoto, S" journalOrPublisher="Journal of the Faculty of Agriculture, Kyushu University" pageId="32" pageNumber="119" pagination="263 - 286" title="The Chrysomelidae of Japan and the Ryukyu Islands V. Subfamily Chrysomelinae." volume="13" year="1964">Kimoto 1964</bibRefCitation>
<bibRefCitation id="66AD16088926768AB992B148051FBB86" author="Kimoto, S" journalOrPublisher="Journal of the Faculty of Agriculture, Kyushu University" pageId="32" pageNumber="119" pagination="263 - 286" title="The Chrysomelidae of Japan and the Ryukyu Islands V. Subfamily Chrysomelinae." volume="13" year="1964">Kimoto 1964</bibRefCitation>
);
<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="30" pageNumber="117" phylum="Chordata" rank="genus">Salix</taxonomicName>
<taxonomicName id="15DA3D9F4F5B0E1BBF9692F7D93E1BEF" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="30" pageNumber="117" phylum="Chordata" rank="genus">Salix</taxonomicName>
spp. (
<bibRefCitation author="Takizawa, H" journalOrPublisher="Kontyu" pageId="36" pageNumber="123" pagination="444 - 468" title="Larvae of the genus Gonioctena Chevrolat (Coleoptera, Chrysomelidae): descriptions of Japanese species and the implications of larval characters for the phylogeny." volume="44" year="1976">Takizawa 1976</bibRefCitation>
<bibRefCitation id="CFC776DF2F3D919EEE4685A929EC8233" author="Takizawa, H" journalOrPublisher="Kontyu" pageId="36" pageNumber="123" pagination="444 - 468" title="Larvae of the genus Gonioctena Chevrolat (Coleoptera, Chrysomelidae): descriptions of Japanese species and the implications of larval characters for the phylogeny." volume="44" year="1976">Takizawa 1976</bibRefCitation>
,
<bibRefCitation author="Kudo, S" editor="Jolivet, P" journalOrPublisher="SPB Academic Publishing, The Hague" pageId="33" pageNumber="120" pagination="727 - 738" title="Diversified reproductive strategies in Gonioctena (Chrysomelinae) leaf beetles." volumeTitle="New Developments in the Biology of Chrysomelidae" year="2003">Kudo and Hasegawa 2003</bibRefCitation>
<bibRefCitation id="D31764A28173DC8659183B21002D4B28" author="Kudo, S" editor="Jolivet, P" journalOrPublisher="SPB Academic Publishing, The Hague" pageId="33" pageNumber="120" pagination="727 - 738" title="Diversified reproductive strategies in Gonioctena (Chrysomelinae) leaf beetles." volumeTitle="New Developments in the Biology of Chrysomelidae" year="2003">Kudo and Hasegawa 2003</bibRefCitation>
);
<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus maximowiczii" order="Malpighiales" pageId="30" pageNumber="117" phylum="Tracheophyta" rank="species" species="maximowiczii">Populus maximowiczii</taxonomicName>
<taxonomicName id="7CFC1C6F59231263AEFE68B062420886" class="Magnoliopsida" family="Salicaceae" genus="Populus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Populus maximowiczii" order="Malpighiales" pageId="30" pageNumber="117" phylum="Tracheophyta" rank="species" species="maximowiczii">Populus maximowiczii</taxonomicName>
,
<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="30" pageNumber="117" phylum="Chordata" rank="genus">Salix</taxonomicName>
<taxonomicName id="C8CBB5E68C69988EC60C7238BAC4102C" class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="30" pageNumber="117" phylum="Chordata" rank="genus">Salix</taxonomicName>
spp. (
<bibRefCitation author="Takizawa, H" journalOrPublisher="Insecta Matsumurana, new series" pageId="36" pageNumber="123" pagination="35 - 50" title="A revision of the genus Gonioctena Chevrolat in Japan (Coleoptera: Chrysomelidae)." volume="63" year="2007">Takizawa 2007</bibRefCitation>
<bibRefCitation id="09476B88EA12C343344AD82D29755D32" author="Takizawa, H" journalOrPublisher="Insecta Matsumurana, new series" pageId="36" pageNumber="123" pagination="35 - 50" title="A revision of the genus Gonioctena Chevrolat in Japan (Coleoptera: Chrysomelidae)." volume="63" year="2007">Takizawa 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="117" type="remarks">
<paragraph pageId="30" pageNumber="117">Remarks.</paragraph>
<paragraph pageId="30" pageNumber="117">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena springlovae" order="Coleoptera" pageId="30" pageNumber="117" phylum="Arthropoda" rank="species" species="springlovae">Gonioctena springlovae</taxonomicName>
<subSubSection id="50FC7D0CCC8D3BD6DC60990E773CC583" pageId="30" pageNumber="117" type="remarks">
<paragraph id="9C760268B4AA4FB53B784D16C8988B71" pageId="30" pageNumber="117">Remarks.</paragraph>
<paragraph id="D46ACAB4AADB19188255AC3DAAEB59DB" pageId="30" pageNumber="117">
<taxonomicName id="06E3636030E57E17C8449E6DB07FA138" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena springlovae" order="Coleoptera" pageId="30" pageNumber="117" phylum="Arthropoda" rank="species" species="springlovae">Gonioctena springlovae</taxonomicName>
is restricted to Hokkaido and Sakhalin, whereas its closely related species
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena gracilicornis" order="Coleoptera" pageId="30" pageNumber="117" phylum="Arthropoda" rank="species" species="gracilicornis">Gonioctena gracilicornis</taxonomicName>
<taxonomicName id="608FE14DF97997450D8C703071DC9AC9" class="Insecta" family="Chrysomelidae" genus="Gonioctena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonioctena gracilicornis" order="Coleoptera" pageId="30" pageNumber="117" phylum="Arthropoda" rank="species" species="gracilicornis">Gonioctena gracilicornis</taxonomicName>
is widely distributed in the Northeastern Palearctic region. The distributions of these two species overlap only in southern Sakhalin (Fig. 16). The type locality &quot;Kioto [= Kyoto in Honshu]&quot; is probably in error. As
<bibRefCitation author="Chujo, M" journalOrPublisher="Niponius, Acta Coleopterologica" pageId="31" pageNumber="118" pagination="1 - 10" title="Descriptions of three new genera and a new species of chrysomelid-beetles from Japan, with some notes on the Japanese species." volume="1" year="1960">
<normalizedToken originalValue="Chûjô">Chujo</normalizedToken>
<bibRefCitation id="012FE424B49F383CB561D6E3D710CF4B" author="Chujo, M" journalOrPublisher="Niponius, Acta Coleopterologica" pageId="31" pageNumber="118" pagination="1 - 10" title="Descriptions of three new genera and a new species of chrysomelid-beetles from Japan, with some notes on the Japanese species." volume="1" year="1960">
<normalizedToken id="3A46F7C3763D2B12642A0FD16324030D" originalValue="Chûjô">Chujo</normalizedToken>
and Kimoto (1960)
</bibRefCitation>
mentioned, no single specimen has been collected again in Honshu whereas many specimens have been collected in Hokkaido. This species is ovoviviparous (
<bibRefCitation author="Takizawa, H" journalOrPublisher="Kontyu" pageId="36" pageNumber="123" pagination="444 - 468" title="Larvae of the genus Gonioctena Chevrolat (Coleoptera, Chrysomelidae): descriptions of Japanese species and the implications of larval characters for the phylogeny." volume="44" year="1976">Takizawa 1976</bibRefCitation>
<bibRefCitation id="6ED431B59EC27CDABC2F8DC14833533A" author="Takizawa, H" journalOrPublisher="Kontyu" pageId="36" pageNumber="123" pagination="444 - 468" title="Larvae of the genus Gonioctena Chevrolat (Coleoptera, Chrysomelidae): descriptions of Japanese species and the implications of larval characters for the phylogeny." volume="44" year="1976">Takizawa 1976</bibRefCitation>
,
<bibRefCitation author="Kimoto, S" journalOrPublisher="Tokai University Press, Tokyo" pageId="32" pageNumber="119" title="Leaf beetles (Chrysomelidae) of Japan." year="1994">Kimoto and Takizawa 1994</bibRefCitation>
<bibRefCitation id="393BA22ABA1574C3718C1A11BCDE7628" author="Kimoto, S" journalOrPublisher="Tokai University Press, Tokyo" pageId="32" pageNumber="119" title="Leaf beetles (Chrysomelidae) of Japan." year="1994">Kimoto and Takizawa 1994</bibRefCitation>
,
<bibRefCitation author="Kudo, S" editor="Jolivet, P" journalOrPublisher="SPB Academic Publishing, The Hague" pageId="33" pageNumber="120" pagination="727 - 738" title="Diversified reproductive strategies in Gonioctena (Chrysomelinae) leaf beetles." volumeTitle="New Developments in the Biology of Chrysomelidae" year="2003">Kudo and Hasegawa 2003</bibRefCitation>
<bibRefCitation id="9A4A5301D6B3DFBD256C25474AC74CE9" author="Kudo, S" editor="Jolivet, P" journalOrPublisher="SPB Academic Publishing, The Hague" pageId="33" pageNumber="120" pagination="727 - 738" title="Diversified reproductive strategies in Gonioctena (Chrysomelinae) leaf beetles." volumeTitle="New Developments in the Biology of Chrysomelidae" year="2003">Kudo and Hasegawa 2003</bibRefCitation>
).
</paragraph>
</subSubSection>

194
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</paragraph>
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<paragraph pageId="0" pageNumber="27">Type.</paragraph>
<paragraph pageId="0" pageNumber="27">
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<paragraph id="E310A862CE6D6B89D3C7B1D1CF2FABDF" pageId="0" pageNumber="27">Type.</paragraph>
<paragraph id="8AB9B575F296573F71945DCFCFA148D4" pageId="0" pageNumber="27">
Habitat in Europae pascuis, no date, Hudson 16 (neotype, designated by Soreng in
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: 256]: LINN (LINN-87.9!)).
</paragraph>
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<subSubSection pageId="0" pageNumber="27" type="many heterotypic synonyms">
<paragraph pageId="0" pageNumber="27">Many heterotypic synonyms.</paragraph>
<paragraph pageId="0" pageNumber="27">
<subSubSection id="F7BDCD1F4C2ABD3D199D31B3EA9EAEE8" pageId="0" pageNumber="27" type="many heterotypic synonyms">
<paragraph id="2C3E1B744C92FFC7D69AACDE3ACC6D7D" pageId="0" pageNumber="27">Many heterotypic synonyms.</paragraph>
<paragraph id="38881D0BD20E843FBCB9129E8CFCEF42" pageId="0" pageNumber="27">
-
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<taxonomicName id="8F9CA3A5648ABAE63BEBAB79C2160A37" lsidName="P. subsp. sect." pageId="0" pageNumber="27" rank="section" section="Pandemos">P. sect. Pandemos</taxonomicName>
Asch. &amp; Graebn., Syn. Mitteleur. Fl. 2: 425. 1900. Type,
<taxonomicName lsidName="P. trivialis" pageId="0" pageNumber="27" rank="species" species="trivialis">
<emphasis italics="true" pageId="0" pageNumber="27">P. trivialis</emphasis>
<taxonomicName id="DBD38B0DC4F04704B215FDC8D6A23E21" lsidName="P. trivialis" pageId="0" pageNumber="27" rank="species" species="trivialis">
<emphasis id="91209C149E900ADA9AE113B02599B7B5" italics="true" pageId="0" pageNumber="27">P. trivialis</emphasis>
</taxonomicName>
L.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="distribution">
<paragraph pageId="0" pageNumber="27">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="27">South Africa, Gautan Province. Introduced, native to western Eurasia and North Africa, introduced to sub-Saharan Africa in Zimbabwe and South Africa.</paragraph>
<subSubSection id="6D872CC66B562DCDE607D46DD95FB9FE" pageId="0" pageNumber="27" type="distribution">
<paragraph id="C7E996615E78BCF6CDCAE577CC5F14F6" pageId="0" pageNumber="27">Distribution.</paragraph>
<paragraph id="669379A33C27C3736027C0B04373C5DF" pageId="0" pageNumber="27">South Africa, Gautan Province. Introduced, native to western Eurasia and North Africa, introduced to sub-Saharan Africa in Zimbabwe and South Africa.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="ecology">
<paragraph pageId="0" pageNumber="27">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="27">ruderal of temperate climates.</paragraph>
<subSubSection id="43D5B4C535894C25CA013BEFA8C7C9BD" pageId="0" pageNumber="27" type="ecology">
<paragraph id="640A7B9FBFE286D0546BCAC0C69E79F0" pageId="0" pageNumber="27">Ecology.</paragraph>
<paragraph id="ED597A40C36B303A5040018940F10608" pageId="0" pageNumber="27">ruderal of temperate climates.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="flowering">
<paragraph pageId="0" pageNumber="27">Flowering.</paragraph>
<paragraph pageId="0" pageNumber="27">spring.</paragraph>
<subSubSection id="DC6B185DE4519570836C6BF95CCBF672" pageId="0" pageNumber="27" type="flowering">
<paragraph id="8E68510BE041A68E4AE0A0641B44DF5F" pageId="0" pageNumber="27">Flowering.</paragraph>
<paragraph id="57C3C0922C0E059257424BEBB36F2797" pageId="0" pageNumber="27">spring.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="economics">
<paragraph pageId="0" pageNumber="27">Economics.</paragraph>
<paragraph pageId="0" pageNumber="27">rarely collected. Sometimes seeded for pastures, invasive.</paragraph>
<subSubSection id="0693405616583E0FC5DACD97E3A1CB31" pageId="0" pageNumber="27" type="economics">
<paragraph id="7E98A717B5485AD85A5BD8BF67CF4E64" pageId="0" pageNumber="27">Economics.</paragraph>
<paragraph id="94240A0FCEB6DEEAC974816C200EA8D4" pageId="0" pageNumber="27">rarely collected. Sometimes seeded for pastures, invasive.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="vouchers">
<paragraph pageId="0" pageNumber="27">Vouchers.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">South Africa</emphasis>
<subSubSection id="EA7B29A6BB5EBF35574F07851751CE1F" pageId="0" pageNumber="27" type="vouchers">
<paragraph id="E1728ACDFEF95B028870AFFFA7FB9C92" pageId="0" pageNumber="27">Vouchers.</paragraph>
<paragraph id="8D571C34F04A99244699FD6070EB673A" pageId="0" pageNumber="27">
<emphasis id="4B692CE255904025A69EA33DAB0E04BE" bold="true" pageId="0" pageNumber="27">South Africa</emphasis>
. Gauteng: Johannesburg, Rosebank 50 Bath Ave., 28 Dec 1962, Meredith s.n. (PRE0021311-0); Gauteng: Johannesburg, Hort. Rosebank, Mar 1965, Meredith s.n. (PRE0029743-0).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="notes">
<paragraph pageId="0" pageNumber="27">Notes.</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName authority="subsp. trivialis" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa trivialis subsp. trivialis" order="Poales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="trivialis" subSpecies="trivialis">Poa trivialis subsp. trivialis</taxonomicName>
<subSubSection id="C471E7F74AE058206F52A29BB49D48C4" pageId="0" pageNumber="27" type="notes">
<paragraph id="16407F8D2149E5049CF61749549211BF" pageId="0" pageNumber="27">Notes.</paragraph>
<paragraph id="6C1DAF0926E7905F5FDA60A04E04A163" pageId="0" pageNumber="27">
<taxonomicName id="856D4352712AE9F780127A46586CCA30" authority="subsp. trivialis" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa trivialis subsp. trivialis" order="Poales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="subSpecies" species="trivialis" subSpecies="trivialis">Poa trivialis subsp. trivialis</taxonomicName>
is reputedly self-incompatible and sexually reproducing (
<bibRefCitation DOI="https://doi.org/10.1080/0028825X.1979.10432571" author="Connor, HE" journalOrPublisher="New Zealand Journal of Botany" pageId="0" pageNumber="27" pagination="547 - 574" refId="B7" refString="Connor, HE, 1979. Breeding systems in the grasses: A survey. New Zealand Journal of Botany 17 (4): 547 - 574, DOI: https://doi.org/10.1080/0028825X.1979.10432571" title="Breeding systems in the grasses: A survey." url="https://doi.org/10.1080/0028825X.1979.10432571" volume="17" year="1979">Connor 1979</bibRefCitation>
<bibRefCitation id="F926A52A2C63B7B4159B888E16FC1C5D" DOI="https://doi.org/10.1080/0028825X.1979.10432571" author="Connor, HE" journalOrPublisher="New Zealand Journal of Botany" pageId="0" pageNumber="27" pagination="547 - 574" refId="B7" refString="Connor, HE, 1979. Breeding systems in the grasses: A survey. New Zealand Journal of Botany 17 (4): 547 - 574, DOI: https://doi.org/10.1080/0028825X.1979.10432571" title="Breeding systems in the grasses: A survey." url="https://doi.org/10.1080/0028825X.1979.10432571" volume="17" year="1979">Connor 1979</bibRefCitation>
). It can be quite invasive in temperate climates with a cool wet season. Aesthetically, it makes a poor lawn grass due to its sprawling habit when mown.
<normalizedToken originalValue="Valdés">Valdes</normalizedToken>
<normalizedToken id="6E42223314539427FB0D4C9AF171245E" originalValue="Valdés">Valdes</normalizedToken>
and Scholz (2009) recorded it only for Algeria in North Africa. The second major subspecies,
<taxonomicName lsidName="P. t. subsp. sylvicola" pageId="0" pageNumber="27" rank="subSpecies" species="t." subSpecies="sylvicola">P. t. subsp. sylvicola</taxonomicName>
<taxonomicName id="C84B14B8EDE94A46A436F6AABB1F714A" lsidName="P. t. subsp. sylvicola" pageId="0" pageNumber="27" rank="subSpecies" species="t." subSpecies="sylvicola">P. t. subsp. sylvicola</taxonomicName>
(Guss.) H. Lindb., has bead-like swellings along the stolons and more hair on the lemma marginal veins and is more tolerant of drought: It is infrequently found outside of the Mediterranean basin and Irano-Turainian floristic region, but is reported across northern Africa (
<bibRefCitation author="Valdes, B" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="0" pageNumber="27" publicationUrl="http://www.emplantbase.org/home.html" refId="B28" refString="Valdes, B, Scholz, H, Raab-Straube von, E, Parolly, G, 2009. Poaceae (pro parte majore). Euro+Med Plantbase - the information resource for Euro-Mediterranean plant diversity. http://www.emplantbase.org/home.html" title="Poaceae (pro parte majore). Euro + Med Plantbase - the information resource for Euro-Mediterranean plant diversity." url="http://www.emplantbase.org/home.html" year="2009">
<normalizedToken originalValue="Valdés">Valdes</normalizedToken>
<bibRefCitation id="B3E13D3C6859C70758DAB834EDBFE24C" author="Valdes, B" journalOrPublisher="Annals of the Missouri Botanical Garden" pageId="0" pageNumber="27" publicationUrl="http://www.emplantbase.org/home.html" refId="B28" refString="Valdes, B, Scholz, H, Raab-Straube von, E, Parolly, G, 2009. Poaceae (pro parte majore). Euro+Med Plantbase - the information resource for Euro-Mediterranean plant diversity. http://www.emplantbase.org/home.html" title="Poaceae (pro parte majore). Euro + Med Plantbase - the information resource for Euro-Mediterranean plant diversity." url="http://www.emplantbase.org/home.html" year="2009">
<normalizedToken id="1FAF4D998266DF34FDA1E9FC996CA261" originalValue="Valdés">Valdes</normalizedToken>
and Scholz 2009
</bibRefCitation>
). 2
<emphasis italics="true" pageId="0" pageNumber="27">n</emphasis>
<emphasis id="4B17645D8D0BF9F0AAE399D7B9C09889" italics="true" pageId="0" pageNumber="27">n</emphasis>
=
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">14</emphasis>
<emphasis id="7431C3E8A0DA9D1CF66BE180099D788B" bold="true" italics="true" pageId="0" pageNumber="27">14</emphasis>
, 14 + 1 - 2B, 15, 27, 28 (27 and 28 counts not confirmed to subspecies may represent subsp.
<taxonomicName lsidName="subsp. sylvicola" pageId="0" pageNumber="27" rank="subSpecies" species="t." subSpecies="sylvicola">sylvicola</taxonomicName>
<taxonomicName id="8244A03EA7E73EA03C6E04EA8E581709" lsidName="subsp. sylvicola" pageId="0" pageNumber="27" rank="subSpecies" species="t." subSpecies="sylvicola">sylvicola</taxonomicName>
). -
<emphasis bold="true" pageId="0" pageNumber="27">Vv</emphasis>
<emphasis id="37A7354FDA099860EBB98E50529A1ADA" bold="true" pageId="0" pageNumber="27">Vv</emphasis>
genotype.
</paragraph>
<caption doi="10.3897/phytokeys.165.55948.figure8" httpUri="https://binary.pensoft.net/fig/468636" pageId="0" pageNumber="27" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figure 8.</emphasis>
<taxonomicName class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa trivialis" order="Poales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="trivialis">
<emphasis italics="true" pageId="0" pageNumber="27">Poa trivialis</emphasis>
<caption id="24ABDECB7A1A14DCB474A9539FA7EB9A" doi="10.3897/phytokeys.165.55948.figure8" httpUri="https://binary.pensoft.net/fig/468636" pageId="0" pageNumber="27" start="Figure 8" startId="F8">
<paragraph id="D19F221E6D9A3A2F2BCEB43D410F70BE" pageId="0" pageNumber="27">
<emphasis id="1620302CB1350723DF3B6570AE1531D3" bold="true" pageId="0" pageNumber="27">Figure 8.</emphasis>
<taxonomicName id="CC09A83FD7CA3560A81A24A07D05E047" class="Liliopsida" family="Poaceae" genus="Poa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Poa trivialis" order="Poales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="trivialis">
<emphasis id="95D669B0CF6D11F3803B19180FCF190E" italics="true" pageId="0" pageNumber="27">Poa trivialis</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="27">A</emphasis>
<emphasis id="025987EDE098C019142388DB7CF09D53" bold="true" pageId="0" pageNumber="27">A</emphasis>
whole plant
<emphasis bold="true" pageId="0" pageNumber="27">B</emphasis>
<emphasis id="A82D5E45DDB155C0D602AA01D2773FF8" bold="true" pageId="0" pageNumber="27">B</emphasis>
part of inflorescence with spikelets mostly in lateral view. Images of Meredith s.n. (PRE0021311-0).
</paragraph>
</caption>

112
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Figs 47-50
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<paragraph pageId="12" pageNumber="73">Description.</paragraph>
<paragraph pageId="12" pageNumber="73">Body length 1.94-2.16 mm, width 1.18-1.29 mm. Color chestnut brown with appendages lighter (Fig. 47). Head surface shiny with few small punctures (Fig. 48), supraorbital pore much larger than a few small punctures on vertex. Supracallinal sulcus clearly separates antennal calli and vertex medially. Frontal ridge wide, longer than antennal calli. Anterofrontal ridge making long denticle about as long as half clypeus length. Pronotum and elytron with fine punctures. Interspaces of elytron flat. Proportions of tarsomeres of male as follows: protarsomeres 7:4:4:9; mesotarsomeres 7:4:4:9; metatarsomeres 10:4:4:9. Median lobe of aedeagus more or less parallel sided in ventral, median lobe view, with ridge in middle being wider at base, narrowing towards middle and widening and disappearing towards apex. In lateral, median lobe view slightly curved with bump on ventral side beyond middle (Fig. 50).</paragraph>
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<paragraph id="B11F5BB81C8B64FB4D347ADBA4587B01" pageId="12" pageNumber="73">Description.</paragraph>
<paragraph id="61F867482B296B75363FD54C85599EB7" pageId="12" pageNumber="73">Body length 1.94-2.16 mm, width 1.18-1.29 mm. Color chestnut brown with appendages lighter (Fig. 47). Head surface shiny with few small punctures (Fig. 48), supraorbital pore much larger than a few small punctures on vertex. Supracallinal sulcus clearly separates antennal calli and vertex medially. Frontal ridge wide, longer than antennal calli. Anterofrontal ridge making long denticle about as long as half clypeus length. Pronotum and elytron with fine punctures. Interspaces of elytron flat. Proportions of tarsomeres of male as follows: protarsomeres 7:4:4:9; mesotarsomeres 7:4:4:9; metatarsomeres 10:4:4:9. Median lobe of aedeagus more or less parallel sided in ventral, median lobe view, with ridge in middle being wider at base, narrowing towards middle and widening and disappearing towards apex. In lateral, median lobe view slightly curved with bump on ventral side beyond middle (Fig. 50).</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="73" type="etymology">
<paragraph pageId="12" pageNumber="73">Etymology.</paragraph>
<paragraph pageId="12" pageNumber="73">The specific epithet is a noun in apposition based on the type locality.</paragraph>
<subSubSection id="F55BFE66E091C738F5258EF3664B926A" pageId="12" pageNumber="73" type="etymology">
<paragraph id="400EDE01C72F3DF00C9F0FC24549FF24" pageId="12" pageNumber="73">Etymology.</paragraph>
<paragraph id="CF249857CA711449BD4BD1F13761280B" pageId="12" pageNumber="73">The specific epithet is a noun in apposition based on the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="73" type="diagnosis and comparison">
<paragraph pageId="12" pageNumber="73">Diagnosis and comparison.</paragraph>
<paragraph pageId="12" pageNumber="73">
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica eltoro" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
<subSubSection id="EF0BFE835F54691AC3131A781321590E" pageId="12" pageNumber="73" type="diagnosis and comparison">
<paragraph id="D08D5B2D3632DED77B09112916D49A65" pageId="12" pageNumber="73">Diagnosis and comparison.</paragraph>
<paragraph id="30C5EB452AB3D05B76B74552DF7B85E4" pageId="12" pageNumber="73">
<taxonomicName id="290841D567048E0BC1E2EBFD45715B29" genus="Ulrica" lsidName="Ulrica eltoro" pageId="12" pageNumber="73" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
can be easily differentiated from
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica iviei" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="iviei">Ulrica iviei</taxonomicName>
<taxonomicName id="0BC583E6EF3260475FC264BFF3CAAF92" genus="Ulrica" lsidName="Ulrica iviei" pageId="12" pageNumber="73" rank="species" species="iviei">Ulrica iviei</taxonomicName>
based on the key below.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="73" type="ecology">
<paragraph pageId="12" pageNumber="73">Ecology.</paragraph>
<paragraph pageId="12" pageNumber="73">
<subSubSection id="7352A97C09C74C2FC4A9D219CF615714" pageId="12" pageNumber="73" type="ecology">
<paragraph id="E7B430A23A8ED7BE512AD276829E62DA" pageId="12" pageNumber="73">Ecology.</paragraph>
<paragraph id="C059EF8DC4974A3D13A4A6AEC2F052C5" pageId="12" pageNumber="73">
Unidentified moss samples that contained
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica eltoro" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
<taxonomicName id="26C4F2D1E86426015551532D24F4D6B1" genus="Ulrica" lsidName="Ulrica eltoro" pageId="12" pageNumber="73" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
were collected in the forest from a variety of substrates (rocks, tree stumps, trunks and branches) (Figs 59, 60).
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="73" type="type material">
<paragraph pageId="12" pageNumber="73">Type material.</paragraph>
<paragraph pageId="12" pageNumber="73">
<subSubSection id="5F2C94AA5354D884B7BA88A6945BBC9C" pageId="12" pageNumber="73" type="type material">
<paragraph id="BC173A5D753ADDCFE7AE6157C47CE897" pageId="12" pageNumber="73">Type material.</paragraph>
<paragraph id="E1DF64A6B12DD4392A9E3FE72E9A7391" pageId="12" pageNumber="73">
Holotype: ♂, Puerto Rico: El Yunque, El Toro trail,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="18.280834">18°16.850'N</geoCoordinate>
<geoCoordinate id="8026D6A389880177DF039ACB19C8907F" direction="north" orientation="latitude" precision="1" value="18.280834">18°16.850'N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-65.829216">65°49.753'W</geoCoordinate>
<geoCoordinate id="5D42F2C8F8FCE524C0AD5E7F43C93858" direction="west" orientation="longitude" precision="1" value="-65.829216">65°49.753'W</geoCoordinate>
, 1066m, 14.VI.2008, moss (unsifted) leg. A. Konstantinov (USNM). Paratype ♂, same label as holotype (USNM).
</paragraph>
<caption pageId="12" pageNumber="73">
<paragraph pageId="12" pageNumber="73">
<caption id="DF79237ECF29EA1AABF4CF853126030E" pageId="12" pageNumber="73">
<paragraph id="FC54A5FACFCC4C137F4E1FC2AE267A67" pageId="12" pageNumber="73">
Figures 47-50.
<taxonomicName class="Insecta" family="Chrysomelidae" genus="Ulrica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ulrica eltoro" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
<taxonomicName id="F2DC3F5457B62DFE7380AD01DAC6D3CA" genus="Ulrica" lsidName="Ulrica eltoro" pageId="12" pageNumber="73" rank="species" species="eltoro">Ulrica eltoro</taxonomicName>
: 47 habitus, dorsal view 48 habitus, frontalview 49 habitus, lateral view 50 median lobe of aedeagus, ventral, lateral and dorsal views.
</paragraph>
</caption>

236
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Campanicola anguilliformis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-H female epigynum (D ventral view, not dissected E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95">Figs 7</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Collection localities. C. anguilliformis sp. nov. (1); C. campanulata (2 - 15); C. falciformis sp. nov. (2 - 5, 16); C. ferrumequina (1, 11); C. heteroidea sp. nov. (17); C. tauricornis sp. nov. (18); C. volubilis sp. nov. (19); P. ducta (20)." figureDoi="10.3897/zookeys.1055.67620.figure16" httpUri="https://binary.pensoft.net/fig/575812" pageId="0" pageNumber="95">, 16</figureCitation>
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<paragraph id="63DB070F9155C4FD684EA660FDDFB7FB" pageId="0" pageNumber="95">
<figureCitation id="850821B3BBD683EA166E91AA78794E23" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Campanicola anguilliformis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-H female epigynum (D ventral view, not dissected E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95">Figs 7</figureCitation>
<figureCitation id="A8B1342F729AED23C4E5936C3D44651C" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Collection localities. C. anguilliformis sp. nov. (1); C. campanulata (2 - 15); C. falciformis sp. nov. (2 - 5, 16); C. ferrumequina (1, 11); C. heteroidea sp. nov. (17); C. tauricornis sp. nov. (18); C. volubilis sp. nov. (19); P. ducta (20)." figureDoi="10.3897/zookeys.1055.67620.figure16" httpUri="https://binary.pensoft.net/fig/575812" pageId="0" pageNumber="95">, 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="95" type="type material">
<paragraph pageId="0" pageNumber="95">Type material.</paragraph>
<paragraph pageId="0" pageNumber="95">
<emphasis bold="true" pageId="0" pageNumber="95">
<emphasis italics="true" pageId="0" pageNumber="95">Holotype</emphasis>
<subSubSection id="D2053B249D765FF8E9859A1D281C487B" pageId="0" pageNumber="95" type="type material">
<paragraph id="F5B9F5DE16695F23C2AAEE12303468AE" pageId="0" pageNumber="95">Type material.</paragraph>
<paragraph id="074262C580E36599663F84D4DF0EA4F9" pageId="0" pageNumber="95">
<emphasis id="46DBF5C87F75AFD3EC421123C96D34CE" bold="true" pageId="0" pageNumber="95">
<emphasis id="D39657DA339EC347821AF550EC88C83E" italics="true" pageId="0" pageNumber="95">Holotype</emphasis>
: ♀, China, Fujian Province
</emphasis>
: Wuyishan City, Wuyi Mountain Natural Reserves (
<geoCoordinate degrees="26" direction="north" minutes="39" orientation="latitude" precision="15" seconds="42" value="26.661667">26°39'42&quot;N</geoCoordinate>
<geoCoordinate id="F2B95B5C4D6EE96006A7C9E7DA016E38" degrees="26" direction="north" minutes="39" orientation="latitude" precision="15" seconds="42" value="26.661667">26°39'42&quot;N</geoCoordinate>
,
<geoCoordinate degrees="117" direction="east" minutes="56" orientation="longitude" precision="15" seconds="24" value="117.94">117°56'24&quot;E</geoCoordinate>
<geoCoordinate id="DB764C627EC55A3A11321617E5FC4FE2" degrees="117" direction="east" minutes="56" orientation="longitude" precision="15" seconds="24" value="117.94">117°56'24&quot;E</geoCoordinate>
, 399 m), 28 August 2019, Y. Zhong and F.J. Liu leg.
<emphasis bold="true" pageId="0" pageNumber="95">Paratypes</emphasis>
<emphasis id="422345929C74F48C77935331A2C812F4" bold="true" pageId="0" pageNumber="95">Paratypes</emphasis>
: 1 ♀, same data as holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="95" type="diagnosis">
<paragraph pageId="0" pageNumber="95">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="95">
<subSubSection id="1274F8A56197D0B62A8A3634DF3FECB6" pageId="0" pageNumber="95" type="diagnosis">
<paragraph id="C168799325100FAAEEB84464B234B683" pageId="0" pageNumber="95">Diagnosis.</paragraph>
<paragraph id="0D928AB90AFDBE94D0F6771C4E0D0D07" pageId="0" pageNumber="95">
This new species is similar to
<taxonomicName genus="C." lsidName="C. ferrumequina" pageId="0" pageNumber="95" rank="species" species="ferrumequina">
<emphasis italics="true" pageId="0" pageNumber="95">C. ferrumequina</emphasis>
<taxonomicName id="F15C986CAAD4B7151778BD1E69BB6A25" lsidName="C. ferrumequina" pageId="0" pageNumber="95" rank="species" species="ferrumequina">
<emphasis id="483D1366F4AB10CAAEF14A70D2CF5B85" italics="true" pageId="0" pageNumber="95">C. ferrumequina</emphasis>
</taxonomicName>
,
<taxonomicName genus="C." lsidName="C. tauricornis" pageId="0" pageNumber="95" rank="species" species="tauricornis">
<emphasis italics="true" pageId="0" pageNumber="95">C. tauricornis</emphasis>
<taxonomicName id="C79A100FFF0EB03ACC5A2986A7130B11" lsidName="C. tauricornis" pageId="0" pageNumber="95" rank="species" species="tauricornis">
<emphasis id="0983430378F56CD79809530C92CF4B76" italics="true" pageId="0" pageNumber="95">C. tauricornis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName genus="C." lsidName="C. volubilis" pageId="0" pageNumber="95" rank="species" species="volubilis">
<emphasis italics="true" pageId="0" pageNumber="95">C. volubilis</emphasis>
<taxonomicName id="587E14A512724A1E6DD6EF399081C1EB" lsidName="C. volubilis" pageId="0" pageNumber="95" rank="species" species="volubilis">
<emphasis id="212F6146BAFE4C21683B8114098EAC80" italics="true" pageId="0" pageNumber="95">C. volubilis</emphasis>
</taxonomicName>
sp. nov. in having spherical spermathecae, and long and looped copulatory ducts, but can be distinguished from them by the direction of the copulatory duct: it starts from the middle and anterior part of the atrium, extends posteriorly, then curves several times and enters the spermathecae laterally and ventrally in
<taxonomicName genus="C." lsidName="C. anguilliformis" pageId="0" pageNumber="95" rank="species" species="anguilliformis">
<emphasis italics="true" pageId="0" pageNumber="95">C. anguilliformis</emphasis>
<taxonomicName id="04CF89574CAEEF1C62BB422D37287A6C" lsidName="C. anguilliformis" pageId="0" pageNumber="95" rank="species" species="anguilliformis">
<emphasis id="E0B3F2C58CB362738E232DE5A48969DA" italics="true" pageId="0" pageNumber="95">C. anguilliformis</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Campanicola anguilliformis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-H female epigynum (D ventral view, not dissected E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95">7E-H</figureCitation>
<figureCitation id="179ACE772335546CCE211AF1308D65C9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Campanicola anguilliformis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-H female epigynum (D ventral view, not dissected E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95">7E-H</figureCitation>
); it starts from the lateral and anterior part of the atrium, extends anteriorly, then curves several times and enters the spermathecae laterally and dorsally in
<taxonomicName genus="C." lsidName="C. ferrumequina" pageId="0" pageNumber="95" rank="species" species="ferrumequina">
<emphasis italics="true" pageId="0" pageNumber="95">C. ferrumequina</emphasis>
<taxonomicName id="145105FFA9AA7E4F1B1F6EDC316DE209" lsidName="C. ferrumequina" pageId="0" pageNumber="95" rank="species" species="ferrumequina">
<emphasis id="E5F5CC37912F42A7B7C60F8695E54F6D" italics="true" pageId="0" pageNumber="95">C. ferrumequina</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Campanicola ferrumequina (Boesenberg &amp; Strand, 1906) A-C female habitus (A dorsal view B lateral view C ventral view) D-H female epigynum (D ventral view, uncleared E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure4" httpUri="https://binary.pensoft.net/fig/575800" pageId="0" pageNumber="95">4E-H</figureCitation>
<figureCitation id="C6201C2BAC5ABEF5306D9F97583111B5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Campanicola ferrumequina (Boesenberg &amp; Strand, 1906) A-C female habitus (A dorsal view B lateral view C ventral view) D-H female epigynum (D ventral view, uncleared E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure4" httpUri="https://binary.pensoft.net/fig/575800" pageId="0" pageNumber="95">4E-H</figureCitation>
); it starts from the lateral and anterior part of the atrium, extends anteriorly, then curves several times and enters the spermathecae laterally and ventrally in
<taxonomicName genus="C." lsidName="C. tauricornis" pageId="0" pageNumber="95" rank="species" species="tauricornis">
<emphasis italics="true" pageId="0" pageNumber="95">C. tauricornis</emphasis>
<taxonomicName id="965C6FB3E6AB4719AF1F56FB9210B3D3" lsidName="C. tauricornis" pageId="0" pageNumber="95" rank="species" species="tauricornis">
<emphasis id="64D4E72D98036D7C3ADDE1EAB5836CAC" italics="true" pageId="0" pageNumber="95">C. tauricornis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName genus="C." lsidName="C. volubilis" pageId="0" pageNumber="95" rank="species" species="volubilis">
<emphasis italics="true" pageId="0" pageNumber="95">C. volubilis</emphasis>
<taxonomicName id="E2C0E6BE04025DF1F2E761BC36F6D3E2" lsidName="C. volubilis" pageId="0" pageNumber="95" rank="species" species="volubilis">
<emphasis id="AE3CBA6B2734F69A95F7D6F8189216B2" italics="true" pageId="0" pageNumber="95">C. volubilis</emphasis>
</taxonomicName>
sp. nov. (Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Campanicola tauricornis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-G female epigynum (D ventral view, cleared E dorsal view, cleared F ventral view, cleared and embedded in Arabic gum G dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-G)." figureDoi="10.3897/zookeys.1055.67620.figure10" httpUri="https://binary.pensoft.net/fig/575806" pageId="0" pageNumber="95">10D-G</figureCitation>
<figureCitation id="DF805EA0DA40C410E86D9A6438EC436A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Campanicola tauricornis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-G female epigynum (D ventral view, cleared E dorsal view, cleared F ventral view, cleared and embedded in Arabic gum G dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-G)." figureDoi="10.3897/zookeys.1055.67620.figure10" httpUri="https://binary.pensoft.net/fig/575806" pageId="0" pageNumber="95">10D-G</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Campanicola volubilis sp. nov. A-C female habitus (A dorsal view B lateral view C ventral view) D-G female epigynum (D ventral view, cleared E dorsal view, cleared F ventral view, cleared and embedded in Arabic gum G dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-G)." figureDoi="10.3897/zookeys.1055.67620.figure11" httpUri="https://binary.pensoft.net/fig/575807" pageId="0" pageNumber="95">11D-G</figureCitation>
<figureCitation id="1B5B4757DB604DA22F1950901DE68E45" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Campanicola volubilis sp. nov. A-C female habitus (A dorsal view B lateral view C ventral view) D-G female epigynum (D ventral view, cleared E dorsal view, cleared F ventral view, cleared and embedded in Arabic gum G dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-G)." figureDoi="10.3897/zookeys.1055.67620.figure11" httpUri="https://binary.pensoft.net/fig/575807" pageId="0" pageNumber="95">11D-G</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="95">
<emphasis bold="true" pageId="0" pageNumber="95">Figure 7.</emphasis>
<taxonomicName authorityName="Li &amp; Liu" authorityYear="2021" class="Arachnida" family="Theridiidae" genus="Campanicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Campanicola anguilliformis" order="Araneae" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="anguilliformis">
<emphasis italics="true" pageId="0" pageNumber="95">Campanicola anguilliformis</emphasis>
<caption id="2A001322A9F88FDF141CD12C1BEDA42F" doi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95" start="Figure 7" startId="F7">
<paragraph id="D16CB8A32F12667C6694378D18A30819" pageId="0" pageNumber="95">
<emphasis id="F435FE4EEAA5E3761D52946D76C7E4A3" bold="true" pageId="0" pageNumber="95">Figure 7.</emphasis>
<taxonomicName id="749C38CD516C918C984CE53AF1E8CA49" authorityName="Li &amp; Liu" authorityYear="2021" class="Arachnida" family="Theridiidae" genus="Campanicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Campanicola anguilliformis" order="Araneae" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="anguilliformis">
<emphasis id="B51F697FF1E4B767739EB9AAEA92BAAE" italics="true" pageId="0" pageNumber="95">Campanicola anguilliformis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="95">A-C</emphasis>
<emphasis id="A9622DDFA309E540D781BAD72FFA37CE" bold="true" pageId="0" pageNumber="95">A-C</emphasis>
female habitus (
<emphasis bold="true" pageId="0" pageNumber="95">A</emphasis>
<emphasis id="8620300AB78D8BF72BB3EDE392463302" bold="true" pageId="0" pageNumber="95">A</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="95">B</emphasis>
<emphasis id="830D296D89EFEF7F7F2C098C8D6BDC1C" bold="true" pageId="0" pageNumber="95">B</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="95">C</emphasis>
<emphasis id="1EBA65CB75BA831B8B443B2092F677CD" bold="true" pageId="0" pageNumber="95">C</emphasis>
lateral view)
<emphasis bold="true" pageId="0" pageNumber="95">D-H</emphasis>
<emphasis id="DD8D5D2C49603D28713F2CFA2AC71FA9" bold="true" pageId="0" pageNumber="95">D-H</emphasis>
female epigynum (
<emphasis bold="true" pageId="0" pageNumber="95">D</emphasis>
<emphasis id="29ED5395C46F7082A0A4ACBCF69255F8" bold="true" pageId="0" pageNumber="95">D</emphasis>
ventral view, not dissected
<emphasis bold="true" pageId="0" pageNumber="95">E</emphasis>
<emphasis id="5054DF5495EAFC0A5E3223C6628C31FC" bold="true" pageId="0" pageNumber="95">E</emphasis>
ventral view, cleared
<emphasis bold="true" pageId="0" pageNumber="95">F</emphasis>
<emphasis id="1E6F9EBEEF1C89047B2B96E7E11B7686" bold="true" pageId="0" pageNumber="95">F</emphasis>
dorsal view, cleared
<emphasis bold="true" pageId="0" pageNumber="95">G</emphasis>
<emphasis id="A7C2E0884C4A35463C11C095B0A18CE5" bold="true" pageId="0" pageNumber="95">G</emphasis>
ventral view, cleared and embedded in Arabic gum
<emphasis bold="true" pageId="0" pageNumber="95">H</emphasis>
<emphasis id="D565841A0A13F25585103F2033C3CE7C" bold="true" pageId="0" pageNumber="95">H</emphasis>
dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="95">A-C</emphasis>
<emphasis id="B1E0E178BB5F170221A2BEAFD5BBBF9E" bold="true" pageId="0" pageNumber="95">A-C</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="95">D-H</emphasis>
<emphasis id="D5331AB1F92E1D1664447D054D99DE78" bold="true" pageId="0" pageNumber="95">D-H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="95" type="etymology">
<paragraph pageId="0" pageNumber="95">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="95">
<subSubSection id="F08BD2E592B3491C6A338345D48E7120" pageId="0" pageNumber="95" type="etymology">
<paragraph id="3C45E953997A066A0684784D8C884E15" pageId="0" pageNumber="95">Etymology.</paragraph>
<paragraph id="761E8108C29515DFB726FC57E5DAF2C9" pageId="0" pageNumber="95">
The specific name is derived from the Latin adjective
<taxonomicName authorityName="Li &amp; Liu" authorityYear="2021" class="Arachnida" family="Theridiidae" genus="Campanicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Campanicola anguilliformis" order="Araneae" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="anguilliformis">
<emphasis italics="true" pageId="0" pageNumber="95">Campanicola anguilliformis</emphasis>
<taxonomicName id="7181EB64DACA6FC0A42E4E47C047FC3F" authorityName="Li &amp; Liu" authorityYear="2021" class="Arachnida" family="Theridiidae" genus="Campanicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Campanicola anguilliformis" order="Araneae" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="anguilliformis">
<emphasis id="30D25014FB014A0F0D9314E54BA85C08" italics="true" pageId="0" pageNumber="95">Campanicola anguilliformis</emphasis>
</taxonomicName>
, meaning eel-like, referring to the eel-like copulatory ducts; adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="95" type="description">
<paragraph pageId="0" pageNumber="95">Description.</paragraph>
<paragraph pageId="0" pageNumber="95">
<emphasis bold="true" pageId="0" pageNumber="95">Male</emphasis>
<subSubSection id="2B3230B7B3FA10886866825B33FDBB88" pageId="0" pageNumber="95" type="description">
<paragraph id="90F3C0BB76BF9307B5279DF9429614C0" pageId="0" pageNumber="95">Description.</paragraph>
<paragraph id="7B55C8E35776E20379182F0C54CC4601" pageId="0" pageNumber="95">
<emphasis id="4AD982CB3B9C47FA7400EF56EDA4233D" bold="true" pageId="0" pageNumber="95">Male</emphasis>
unknown.
<emphasis bold="true" pageId="0" pageNumber="95">Female (holotype)</emphasis>
<emphasis id="3FB963F2885592ECB2665FE69EEDA24B" bold="true" pageId="0" pageNumber="95">Female (holotype)</emphasis>
: Total length 2.06. Prosoma 0.91 long, 0.80 wide, brownish yellow, with black margin. Sternum 0.55 long, 0.51 wide, yellow, peltate, with sparse setae. Opisthosoma 1.45 long, 1.16 wide, dorsum black, with two pairs of transverse patterns in the shape of mustache and a longitudinal stripe, irregular white spots distributed in the patterns and the stripe; venter yellowish, with few bright white spots. Anal tubercle yellow. Spinnerets surrounded with black ring, without colulus (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Campanicola anguilliformis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-H female epigynum (D ventral view, not dissected E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95">7A-C</figureCitation>
<figureCitation id="5A976595EE7787BA2423525E92AF610F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Campanicola anguilliformis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-H female epigynum (D ventral view, not dissected E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95">7A-C</figureCitation>
). Diameters of eyes: AME 0.07, ALE 0.08, PME 0.09, PLE 0.07. Interdistances of eyes: AME-AME 0.06, AME-ALE 0.03, PME-PME 0.07, PME-PLE 0.05. Clypeus 0.16 high, yellow. Chelicerae yellow, promargin with 2 teeth. Endites yellow. Labium 0.19 long, 0.05 wide, yellow and rectangular, fused with sternum. Measurements of legs: I 3.58 (1.12, 0.29, 0.80, 0.89, 0.48), II 2.50 (0.77, 0.29, 0.47, 0.56, 0.41), III 1.90 (0.60, 0.22, 0.32, 0.42, 0.34), IV 3.00 (0.98, 0.32, 0.62, 0.66, 0.42). Leg formula: I-IV-II-III. Femur and patella light yellow, tibia, metatarsus and tarsus orange yellow. Femur, tibia and metatarsus with dark brown ring. Epigynum: atrium depression, small and oval; copulatory pores distinguishable, close to each other, located on the middle and anterior part of atrium; copulatory ducts long, winding, lightly sclerotized, connected with spermathecae from anterior-ventral part; spermathecae spherical; fertilization ducts short, curved and tapering (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Campanicola anguilliformis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-H female epigynum (D ventral view, not dissected E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95">7E, F</figureCitation>
<figureCitation id="9CEB8CAF38DC9F99B74BF06B96B6417E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Campanicola anguilliformis sp. nov. A-C female habitus (A dorsal view B ventral view C lateral view) D-H female epigynum (D ventral view, not dissected E ventral view, cleared F dorsal view, cleared G ventral view, cleared and embedded in Arabic gum H dorsal view, cleared and embedded in Arabic gum). Scale bars: 0.5 mm (A-C); 0.1 mm (D-H)." figureDoi="10.3897/zookeys.1055.67620.figure7" httpUri="https://binary.pensoft.net/fig/575803" pageId="0" pageNumber="95">7E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="95" type="distribution">
<paragraph pageId="0" pageNumber="95">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="95">
<subSubSection id="8E2965AD24EA111800F34D13F7CC81F1" pageId="0" pageNumber="95" type="distribution">
<paragraph id="DDB739C092942267649D81F1D37F38B4" pageId="0" pageNumber="95">Distribution.</paragraph>
<paragraph id="152DDDF3064C06C93A53D89DC88961D6" pageId="0" pageNumber="95">
China (Sichuan) (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Collection localities. C. anguilliformis sp. nov. (1); C. campanulata (2 - 15); C. falciformis sp. nov. (2 - 5, 16); C. ferrumequina (1, 11); C. heteroidea sp. nov. (17); C. tauricornis sp. nov. (18); C. volubilis sp. nov. (19); P. ducta (20)." figureDoi="10.3897/zookeys.1055.67620.figure16" httpUri="https://binary.pensoft.net/fig/575812" pageId="0" pageNumber="95">16</figureCitation>
<figureCitation id="91881E5C8C9A1573AABF86B51740BE86" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Collection localities. C. anguilliformis sp. nov. (1); C. campanulata (2 - 15); C. falciformis sp. nov. (2 - 5, 16); C. ferrumequina (1, 11); C. heteroidea sp. nov. (17); C. tauricornis sp. nov. (18); C. volubilis sp. nov. (19); P. ducta (20)." figureDoi="10.3897/zookeys.1055.67620.figure16" httpUri="https://binary.pensoft.net/fig/575812" pageId="0" pageNumber="95">16</figureCitation>
).
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134
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@ -1,108 +1,108 @@
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<taxonomicName LSID="2050F2FC-108F-593E-A517-CDFC629B8897" authority="(Lindl.) Rchb. f., Annales Botanices Systematicae 6: 480. 1861." authorityName="(Lindl.) Rchb. f., Annales Botanices Systematicae 6: 480. 1861." class="Liliopsida" family="Orchidaceae" genus="Elleanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Elleanthus furfuraceus" order="Asparagales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="furfuraceus">Elleanthus furfuraceus (Lindl.) Rchb.f., Annales Botanices Systematicae 6: 480. 1861.</taxonomicName>
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<paragraph pageId="0" pageNumber="93">
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Evelyna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Evelyna furfuracea" order="Asparagales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="furfuracea">Evelyna furfuracea</taxonomicName>
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<taxonomicName id="D2C3C8E65CBC1498325C640E8CF51C4F" class="Liliopsida" family="Orchidaceae" genus="Evelyna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Evelyna furfuracea" order="Asparagales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="furfuracea">Evelyna furfuracea</taxonomicName>
Basionym:
<taxonomicName class="Liliopsida" family="Orchidaceae" genus="Evelyna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Evelyna furfuracea" order="Asparagales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="furfuracea">Evelyna furfuracea</taxonomicName>
<taxonomicName id="02CCD65A1782A50D6212FF24347A3D47" class="Liliopsida" family="Orchidaceae" genus="Evelyna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Evelyna furfuracea" order="Asparagales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="furfuracea">Evelyna furfuracea</taxonomicName>
Lindl.,
<taxonomicName family="Orchidaceae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="" order="Asparagales" pageId="0" pageNumber="93" rank="family">Orchidaceae</taxonomicName>
<taxonomicName id="5450322A8D3AFD190E6ADC00C5524008" class="Liliopsida" family="Orchidaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asparagales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="family">Orchidaceae</taxonomicName>
Lindenianae 12. no. 65. 1846. Type: Venezuela, Trujillo &quot;Agua de Obispo, at the height of 9000 feet [2743 m], May&quot;,
<emphasis italics="true" pageId="0" pageNumber="93">Linden 627</emphasis>
<emphasis id="215B8021B8882288633CD8BF8EFD4992" italics="true" pageId="0" pageNumber="93">Linden 627</emphasis>
; Lectotype (designated here): P (P00389698); isolectotypes: P (P00389697), BR (BR0000013083588), W-R (W-R17083); Syntype: Venezuela, Merida &quot;forest of Merida, at the height of 5500 feet [1676 m], June&quot;,
<emphasis italics="true" pageId="0" pageNumber="93">Linden 619</emphasis>
<emphasis id="29A42BD32488B7D8411E1332796D913D" italics="true" pageId="0" pageNumber="93">Linden 619</emphasis>
(unknown location).
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<paragraph pageId="0" pageNumber="93">Note.</paragraph>
<paragraph pageId="0" pageNumber="93">
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<paragraph id="5C97625FF84EDABE0B600CCB27E5E565" pageId="0" pageNumber="93">Note.</paragraph>
<paragraph id="6049CA22D6D7D0A3E143387A963F4AAE" pageId="0" pageNumber="93">
In describing
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<emphasis italics="true" pageId="0" pageNumber="93">Evelyna furfuracea</emphasis>
<taxonomicName id="78F15DE4412EC924E1B2602CBAB1309A" class="Liliopsida" family="Orchidaceae" genus="Evelyna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Evelyna furfuracea" order="Asparagales" pageId="0" pageNumber="93" phylum="Tracheophyta" rank="species" species="furfuracea">
<emphasis id="99F004D868CA22A74795CFE4CC175432" italics="true" pageId="0" pageNumber="93">Evelyna furfuracea</emphasis>
</taxonomicName>
,
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.66687" author="Lindley, J" journalOrPublisher="Bradbury and Evans, Whitefriars, London" pageId="0" pageNumber="93" refId="B10" refString="Lindley, J, 1846. Orchidaceae Lindenianae: Notes open a collection of orchids formed in Colombia and Cuba by Mr. J. Linden. Bradbury and Evans, Whitefriars, London, DOI: https://doi.org/10.5962/bhl.title.66687" title="Orchidaceae Lindenianae: Notes open a collection of orchids formed in Colombia and Cuba by Mr. J. Linden." url="https://doi.org/10.5962/bhl.title.66687" year="1846">Lindley (1846</bibRefCitation>
<bibRefCitation id="9C757AF7D74BCAD5BB409D87ED76265B" DOI="https://doi.org/10.5962/bhl.title.66687" author="Lindley, J" journalOrPublisher="Bradbury and Evans, Whitefriars, London" pageId="0" pageNumber="93" refId="B10" refString="Lindley, J, 1846. Orchidaceae Lindenianae: Notes open a collection of orchids formed in Colombia and Cuba by Mr. J. Linden. Bradbury and Evans, Whitefriars, London, DOI: https://doi.org/10.5962/bhl.title.66687" title="Orchidaceae Lindenianae: Notes open a collection of orchids formed in Colombia and Cuba by Mr. J. Linden." url="https://doi.org/10.5962/bhl.title.66687" year="1846">Lindley (1846</bibRefCitation>
: 12) cited two Linden collections:
<emphasis italics="true" pageId="0" pageNumber="93">619</emphasis>
<emphasis id="9213BF093C75E9484030EAC044359A8B" italics="true" pageId="0" pageNumber="93">619</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="93">627</emphasis>
<emphasis id="F1F048C7A36A49BC09A5BA024CA3CC2D" italics="true" pageId="0" pageNumber="93">627</emphasis>
. Unfortunately, we were unable to locate
<emphasis italics="true" pageId="0" pageNumber="93">Linden 619</emphasis>
<emphasis id="34328E1500AF2B00A7DDB37E379858C2" italics="true" pageId="0" pageNumber="93">Linden 619</emphasis>
. However, collection
<emphasis italics="true" pageId="0" pageNumber="93">Linden 627</emphasis>
<emphasis id="A0C0456DF23794E5C35205ED763D9C93" italics="true" pageId="0" pageNumber="93">Linden 627</emphasis>
is stored in the
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National
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<normalizedToken id="E55C4FD7DA189C347710BEA383B4B97F" originalValue="d’Histoire">d'Histoire</normalizedToken>
Naturelle in Paris (two specimens), in Meise Botanic Garden (one specimen) and in the Naturhistorisches Museum in Vienna (one specimen). However, only the specimens in P and BR are marked as the type collection and only one of them is a complete specimen (P00389698) (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Evelyna furfuracea Lindl. Specimen Linden 627 at the Museum National d'Histoire Naturelle, Paris (P 00389698) designated here as lectotype (CC BY 4.0; http: // coldb. mnhn. fr / catalognumber / mnhn / p / p 00389698)." figureDoi="10.3897/phytokeys.182.68782.figure5" httpUri="https://binary.pensoft.net/fig/592417" pageId="0" pageNumber="93">5</figureCitation>
<figureCitation id="C0DDC64DDD3C8E746DD67100112BD0BE" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Evelyna furfuracea Lindl. Specimen Linden 627 at the Museum National d'Histoire Naturelle, Paris (P 00389698) designated here as lectotype (CC BY 4.0; http: // coldb. mnhn. fr / catalognumber / mnhn / p / p 00389698)." figureDoi="10.3897/phytokeys.182.68782.figure5" httpUri="https://binary.pensoft.net/fig/592417" pageId="0" pageNumber="93">5</figureCitation>
). It can be characterised by lanceolate, acuminate and coriaceous leaves; terminal, laxly to subdensely flowered (with few to several) inflorescences; and subrounded-ovate, acute bracts. This specimen has also a lip obovate to suborbicular in outline, concave, unlobed and retuse in the front and saccate at the base with two large, well-separated, ellipsoid/ovoid calli. Therefore, it has been selected as the lectotype (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Evelyna furfuracea Lindl. Specimen Linden 627 at the Museum National d'Histoire Naturelle, Paris (P 00389698) designated here as lectotype (CC BY 4.0; http: // coldb. mnhn. fr / catalognumber / mnhn / p / p 00389698)." figureDoi="10.3897/phytokeys.182.68782.figure5" httpUri="https://binary.pensoft.net/fig/592417" pageId="0" pageNumber="93">5</figureCitation>
<figureCitation id="5504E08F3E712219BE72CBDBA0B98572" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Evelyna furfuracea Lindl. Specimen Linden 627 at the Museum National d'Histoire Naturelle, Paris (P 00389698) designated here as lectotype (CC BY 4.0; http: // coldb. mnhn. fr / catalognumber / mnhn / p / p 00389698)." figureDoi="10.3897/phytokeys.182.68782.figure5" httpUri="https://binary.pensoft.net/fig/592417" pageId="0" pageNumber="93">5</figureCitation>
).
</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="93">Figure 5.</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="93">Evelyna furfuracea</emphasis>
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<emphasis id="4A073B35BB935FBF514F5F52CA937410" bold="true" pageId="0" pageNumber="93">Figure 5.</emphasis>
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<emphasis id="B6F426CD066E5D1F04A25F9611E59154" italics="true" pageId="0" pageNumber="93">Evelyna furfuracea</emphasis>
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Lindl. Specimen
<emphasis italics="true" pageId="0" pageNumber="93">Linden 627</emphasis>
<emphasis id="38116DA67A8332B33055C3B29980AD2B" italics="true" pageId="0" pageNumber="93">Linden 627</emphasis>
at the Museum National
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Naturelle, Paris (P00389698) designated here as lectotype (CC BY 4.0; http://coldb.mnhn.fr/catalognumber/mnhn/p/p00389698).
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306
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<bibRefCitation author="Cottle, TJ" journalOrPublisher="Canada Journal of Industry, Science, and the Arts" pageId="56" pageNumber="57" pagination="216 - 219" title="On the two species of Astacus found in upper Canada." volume="45" year="1863">Cottle 1863</bibRefCitation>
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<bibRefCitation id="265AD5F61E4D2E6F33CFED58826EEF15" author="Cottle, TJ" journalOrPublisher="Canada Journal of Industry, Science, and the Arts" pageId="56" pageNumber="57" pagination="216 - 219" title="On the two species of Astacus found in upper Canada." volume="45" year="1863">Cottle 1863</bibRefCitation>
:217.
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<taxonomicName class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus argillicola" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="argillicola">Cambarus argillicola</taxonomicName>
<bibRefCitation author="Faxon, W" journalOrPublisher="Proceedings of the American Academy of Arts and Science" pageId="56" pageNumber="57" pagination="107 - 158" title="Descriptions of new species of Cambarus; to which is added synonomical list of the known species of Cambarus and Astacus." volume="20" year="1884">Faxon 1884</bibRefCitation>
<paragraph id="F92CE8103F89D299F470DF79858F48B4" pageId="24" pageNumber="25">
<taxonomicName id="84BE605BBBE4868119F637252C2B8C35" class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus argillicola" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="argillicola">Cambarus argillicola</taxonomicName>
<bibRefCitation id="20CDA5B4344EC47400A12F1901C07754" author="Faxon, W" journalOrPublisher="Proceedings of the American Academy of Arts and Science" pageId="56" pageNumber="57" pagination="107 - 158" title="Descriptions of new species of Cambarus; to which is added synonomical list of the known species of Cambarus and Astacus." volume="20" year="1884">Faxon 1884</bibRefCitation>
:115.
</paragraph>
<paragraph pageId="24" pageNumber="25">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus (Bartonius) argillicola" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="argillicola" subGenus="Bartonius">Cambarus (Bartonius) argillicola</taxonomicName>
<bibRefCitation author="Ortmann, AE" journalOrPublisher="Proceedings of the American Philosophical Society" pageId="58" pageNumber="59" pagination="91 - 136" title="The mutual affinities of the species of Cambarus, and their dispersal over the United States." volume="44" year="1905 b">Ortmann 1905b</bibRefCitation>
<paragraph id="03DF7B174F4DB476AEC19059BE00A8C7" pageId="24" pageNumber="25">
<taxonomicName id="DBACC99428E88DBFEF7D7712C2BF8147" class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus (Bartonius) argillicola" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="argillicola" subGenus="Bartonius">Cambarus (Bartonius) argillicola</taxonomicName>
<bibRefCitation id="9B84135EE3D09E8FEE0C4F0F2D8EA177" author="Ortmann, AE" journalOrPublisher="Proceedings of the American Philosophical Society" pageId="58" pageNumber="59" pagination="91 - 136" title="The mutual affinities of the species of Cambarus, and their dispersal over the United States." volume="44" year="1905 b">Ortmann 1905b</bibRefCitation>
:120.
</paragraph>
<paragraph pageId="24" pageNumber="25">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Bartonius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Bartonius argillicola" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="argillicola">Bartonius argillicola</taxonomicName>
<paragraph id="E59FFCE510BB3762566814BED217E192" pageId="24" pageNumber="25">
<taxonomicName id="B46A9909E0F2F664406D27CCB21EDFFB" genus="Bartonius" lsidName="Bartonius argillicola" pageId="24" pageNumber="25" rank="species" species="argillicola">Bartonius argillicola</taxonomicName>
Wiliamson 1907:749.
</paragraph>
<paragraph pageId="24" pageNumber="25">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus (Cambarus) fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens" subGenus="Cambarus">Cambarus (Cambarus) fodiens</taxonomicName>
<bibRefCitation pageId="24" pageNumber="25">Fowler 1912</bibRefCitation>
<paragraph id="28C6655D5911304D6A753C59C8428392" pageId="24" pageNumber="25">
<taxonomicName id="A2322388F9027382885E37B53A6C0EEB" class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus (Cambarus) fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens" subGenus="Cambarus">Cambarus (Cambarus) fodiens</taxonomicName>
<bibRefCitation id="85F9FCA98E226291AC1005BF48F2A151" pageId="24" pageNumber="25">Fowler 1912</bibRefCitation>
:341.
</paragraph>
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<taxonomicName class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens">Cambarus fodiens</taxonomicName>
<bibRefCitation author="Huntsman, AG" journalOrPublisher="Contributions to Canadian Biology" pageId="57" pageNumber="58" pagination="145 - 163" title="The freshwater malacostraca of Ontario" volume="1911 - 1914" year="1915">Huntsman 1915</bibRefCitation>
<paragraph id="5A469BE124A363030C69B3CB8F73429D" pageId="24" pageNumber="25">
<taxonomicName id="1A332CE54D1CDD7BAFAA5E246C5DD6E4" class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens">Cambarus fodiens</taxonomicName>
<bibRefCitation id="E7C86BAA2B530C2CBE4EBD22A43496FB" author="Huntsman, AG" journalOrPublisher="Contributions to Canadian Biology" pageId="57" pageNumber="58" pagination="145 - 163" title="The freshwater malacostraca of Ontario" volume="1911 - 1914" year="1915">Huntsman 1915</bibRefCitation>
:158, figs. 8f, 9d, 10e, 11a, 12e.
<bibRefCitation author="Crocker, DW" journalOrPublisher="University of Toronto Press, Toronto, Ontario" pageId="56" pageNumber="57" title="Handbook of the crayfishes of Ontario" year="1968">Crocker and Barr 1968</bibRefCitation>
<bibRefCitation id="80F5321E033194AA835E5ECF9583EAC4" author="Crocker, DW" journalOrPublisher="University of Toronto Press, Toronto, Ontario" pageId="56" pageNumber="57" title="Handbook of the crayfishes of Ontario" year="1968">Crocker and Barr 1968</bibRefCitation>
:129-135, figs. 28, 37, 46, 57, 62, 85.
</paragraph>
<paragraph pageId="24" pageNumber="25">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus (Bartonius) fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens" subGenus="Bartonius">Cambarus (Bartonius) fodiens</taxonomicName>
<bibRefCitation author="Creaser, EP" journalOrPublisher="Papers of the Michigan Academy of Sciences Arts and Letters" pageId="56" pageNumber="57" pagination="257 - 276" title="The Michigan decapods crustaceans." volume="13" year="1931">Creaser 1931</bibRefCitation>
<paragraph id="1C78310038F86A025F31DA229283C260" pageId="24" pageNumber="25">
<taxonomicName id="93B9C0C2F99EB42F31A288D1357CCB26" class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus (Bartonius) fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens" subGenus="Bartonius">Cambarus (Bartonius) fodiens</taxonomicName>
<bibRefCitation id="FEA0F7840069B9E500C194B6C4CF1D75" author="Creaser, EP" journalOrPublisher="Papers of the Michigan Academy of Sciences Arts and Letters" pageId="56" pageNumber="57" pagination="257 - 276" title="The Michigan decapods crustaceans." volume="13" year="1931">Creaser 1931</bibRefCitation>
:260, fig. 37.
</paragraph>
<paragraph pageId="24" pageNumber="25">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus (Creaserinus) fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens" subGenus="Creaserinus">Fallicambarus (Creaserinus) fodiens</taxonomicName>
<bibRefCitation author="Hobbs, HH Jr." editor="Holt, PC" journalOrPublisher="Virginia Polytechnic Institution, Blacksburg, Virginia" pageId="57" pageNumber="58" pagination="93 - 179" title="On the distribution and phylogeny of the crayfish genus Cambarus" volumeTitle="The distributional history of the biota of the southern Appalachians. Part I: Invertebrates. Research Division Monograph 1" year="1969">Hobbs 1969</bibRefCitation>
<paragraph id="265671212EB509A4E0EB65C4FBC04058" pageId="24" pageNumber="25">
<taxonomicName id="73FF7653EAF75F2E6363BD93CCEBA003" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus (Creaserinus) fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens" subGenus="Creaserinus">Fallicambarus (Creaserinus) fodiens</taxonomicName>
<bibRefCitation id="406A6FE67BEB024510580D81F372D419" author="Hobbs, HH Jr." editor="Holt, PC" journalOrPublisher="Virginia Polytechnic Institution, Blacksburg, Virginia" pageId="57" pageNumber="58" pagination="93 - 179" title="On the distribution and phylogeny of the crayfish genus Cambarus" volumeTitle="The distributional history of the biota of the southern Appalachians. Part I: Invertebrates. Research Division Monograph 1" year="1969">Hobbs 1969</bibRefCitation>
: 111, fig. 20e; 1972:102, figs 83c, 84b, 85b; 1974:23, fig. 82; 1989:29, fig. 116.
<bibRefCitation author="Page, LM" journalOrPublisher="Bulletin Illinois Natural History Survey" pageId="58" pageNumber="59" pagination="335 - 448" title="The crayfishes and shrimp of Illinois." volume="33" year="1985">Page 1985</bibRefCitation>
<bibRefCitation id="A9B1D303D4FB9BC4B15ED46667DB52D4" author="Page, LM" journalOrPublisher="Bulletin Illinois Natural History Survey" pageId="58" pageNumber="59" pagination="335 - 448" title="The crayfishes and shrimp of Illinois." volume="33" year="1985">Page 1985</bibRefCitation>
:422, figs. 155-158.
<bibRefCitation author="Hobbs, HH III" journalOrPublisher="Miscellaneous Publications of the Milwaukee Public Museum" pageId="57" pageNumber="58" pagination="1 - 177" title="The crayfishes and shrimp of Wisconsin" volume="5" year="1988">Hobbs and Jass 1988</bibRefCitation>
<bibRefCitation id="53CAF38E88DBBE0A716010DB0DA86DF4" author="Hobbs, HH III" journalOrPublisher="Miscellaneous Publications of the Milwaukee Public Museum" pageId="57" pageNumber="58" pagination="1 - 177" title="The crayfishes and shrimp of Wisconsin" volume="5" year="1988">Hobbs and Jass 1988</bibRefCitation>
:39-43, figs.30
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30n.
<bibRefCitation author="Jezerinac, RF" journalOrPublisher="Ohio Biological Survey Bulletin" pageId="58" pageNumber="59" pagination="1 - 193" title="The Crayfishes (Decapoda: Cambaridae) of West Virginia" volume="10" year="1995">Jezerinac et al. 1995</bibRefCitation>
<bibRefCitation id="B9186997D7589DB8170115172E9CB68B" author="Jezerinac, RF" journalOrPublisher="Ohio Biological Survey Bulletin" pageId="58" pageNumber="59" pagination="1 - 193" title="The Crayfishes (Decapoda: Cambaridae) of West Virginia" volume="10" year="1995">Jezerinac et al. 1995</bibRefCitation>
:180-187, figs. 88
<normalizedToken originalValue="a–">a-</normalizedToken>
<normalizedToken id="8BCBC25B0C389C9C57583C75817BE3B5" originalValue="a–">a-</normalizedToken>
88h.
<bibRefCitation author="Taylor, CA" journalOrPublisher="Illinois Natural History Survey Bulletin" pageId="59" pageNumber="60" title="Crayfishes of Kentucky" volume="28" year="2004">Taylor and Schuster 2004</bibRefCitation>
<bibRefCitation id="EFE3F6E30B57B7C4DE77495B98B82499" author="Taylor, CA" journalOrPublisher="Illinois Natural History Survey Bulletin" pageId="59" pageNumber="60" title="Crayfishes of Kentucky" volume="28" year="2004">Taylor and Schuster 2004</bibRefCitation>
:131-133, figs. 100, 101
<normalizedToken originalValue="A–">A-</normalizedToken>
<normalizedToken id="EC957168FDC3B3E40FADC41852094CB5" originalValue="A–">A-</normalizedToken>
101H.
</paragraph>
<paragraph pageId="24" pageNumber="25">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<bibRefCitation author="Pflieger, WL" journalOrPublisher="Missouri Department of Conservation, Jefferson City" pageId="58" pageNumber="59" title="The crayfishes of Missouri" year="1996">Pflieger 1996</bibRefCitation>
<paragraph id="09CB19819E0E61E1B7D1A9FC73EE9F7C" pageId="24" pageNumber="25">
<taxonomicName id="722CB87AD46E8235EEBB0DF62481C437" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<bibRefCitation id="F07A9795CBBFEB7180B08319CF200689" author="Pflieger, WL" journalOrPublisher="Missouri Department of Conservation, Jefferson City" pageId="58" pageNumber="59" title="The crayfishes of Missouri" year="1996">Pflieger 1996</bibRefCitation>
:66-70, figs, 8
<normalizedToken originalValue="A–">A-</normalizedToken>
<normalizedToken id="04D129A64D81A79A71761C33E77B550B" originalValue="A–">A-</normalizedToken>
8H.
<bibRefCitation author="Taylor, CA" journalOrPublisher="Fisheries" pageId="59" pageNumber="60" pagination="25 - 38" title="Conservation status of crayfishes of the United States and Canada." volume="21" year="1996">Taylor et al. 1996</bibRefCitation>
<bibRefCitation id="338B7F272A761A2AA69E4CCB53B289D8" author="Taylor, CA" journalOrPublisher="Fisheries" pageId="59" pageNumber="60" pagination="25 - 38" title="Conservation status of crayfishes of the United States and Canada." volume="21" year="1996">Taylor et al. 1996</bibRefCitation>
:30.
<bibRefCitation author="Taylor, CA" journalOrPublisher="Fisheries" pageId="59" pageNumber="60" pagination="372 - 389" title="A reassessment of the conservation status of crayfishes of the United States and Canada after 10 + years of increased awareness." volume="32" year="2007">Taylor et al. 2007</bibRefCitation>
<bibRefCitation id="0ADC4506EA38509557085257D3D64EF9" author="Taylor, CA" journalOrPublisher="Fisheries" pageId="59" pageNumber="60" pagination="372 - 389" title="A reassessment of the conservation status of crayfishes of the United States and Canada after 10 + years of increased awareness." volume="32" year="2007">Taylor et al. 2007</bibRefCitation>
:383.
</paragraph>
</subSubSection>
<subSubSection lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="diagnosis">
<paragraph pageId="24" pageNumber="25">Diagnosis.</paragraph>
<paragraph lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<subSubSection id="B2C2237CF0184D93C58F09DB1A1893BA" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="D8970B6168001C0C24F72CBD76D88155" pageId="24" pageNumber="25">Diagnosis.</paragraph>
<paragraph id="3BD6DE3E1859B7A230A2C2C3E7E786A6" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
Rostrum slightly broad and moderately excavated, deflected ventrally; margins converging to form acumen cephalically with reduced upturned tubercle; postorbital ridge reduced, not terminating in tubercle; cephalothorax dorsolaterally compressed in profile and vaulted; areola obliterated; branchiostegal region devoid of tubercles; chelae diamond shaped; mesial surface of palm with 2 distinct rows of tubercles; dorsalmost row consisting of 6-9 serrate tubercles; second row consisting
<pageBreakToken pageId="25" pageNumber="26" start="start">of</pageBreakToken>
<pageBreakToken id="112BBD8622DD83BD427650AD9FD7FB40" pageId="25" pageNumber="26" start="start">of</pageBreakToken>
3-6 circular tubercles; basiodactyl row consisting of 5-7 punctations; opposable surface of dactyl with distinct basal notch; junction of dactyl and propodus setiferous; first form gonopods basally contiguous, with 2 terminal elements bent 90° to shaft; central projection of populations on the floodplain possessing distinct subapical notch; total length of central projection equal to mesial process length; mesial process bulbous, truncating distally; second form gonopod non-corneous and blunt; subapical notch absent in second form gonopod; annulus ventralis rhomboid in shape with deep S-shaped sinus and C-shaped fossa; embedded shallowly in sternum, and movable.
</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="26" type="color in life">
<paragraph pageId="25" pageNumber="26">Color in life.</paragraph>
<paragraph pageId="25" pageNumber="26">Carapace dorsally and laterally tan, brown, reddish brown, or gray; cephalic and branchial region mottled with black or deep grey spots; chelae tan, deep gray, or gray brown; tubercles on chelae cream or light gray; distal region of dactyl and propodus increasingly orange; perieopods green or light grey; abdomen grey or olivaceous brown, with 2 distinct dorsal stripes; ventral surfaces cream or white.</paragraph>
<subSubSection id="8C7BDBE4E7A4D1E1FA3F37764202E84E" pageId="25" pageNumber="26" type="color in life">
<paragraph id="36DC7DA0FF60F45BBB83911A683C1451" pageId="25" pageNumber="26">Color in life.</paragraph>
<paragraph id="F581150F81D39A4B86C717F84747AEA7" pageId="25" pageNumber="26">Carapace dorsally and laterally tan, brown, reddish brown, or gray; cephalic and branchial region mottled with black or deep grey spots; chelae tan, deep gray, or gray brown; tubercles on chelae cream or light gray; distal region of dactyl and propodus increasingly orange; perieopods green or light grey; abdomen grey or olivaceous brown, with 2 distinct dorsal stripes; ventral surfaces cream or white.</paragraph>
</subSubSection>
<subSubSection pageId="25" pageNumber="26" type="specimens examined">
<paragraph pageId="25" pageNumber="26">Specimens examined.</paragraph>
<paragraph pageId="25" pageNumber="26">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<subSubSection id="E69CA8220D7071F5F34EE5C786ED37F0" pageId="25" pageNumber="26" type="specimens examined">
<paragraph id="7935EC02453B4E6C4E431719A2D98440" pageId="25" pageNumber="26">Specimens examined.</paragraph>
<paragraph id="64F2C3C1B9D24D952461A07C691CF42B" pageId="25" pageNumber="26">
<taxonomicName id="66D50F2172B4BB74BB09E2296B6A419D" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
were collected from two counties at three locations in the current study, as listed below.
</paragraph>
<paragraph pageId="25" pageNumber="26">CABELL COUNTY: Green Bottom Swamp at Hoeft Marsh Wildlife Management Area, 38.58616 -82.24878; 2 April 2009 - (WLU 09040201), 2 I♂, 2 II♂, 1 O♀, 4♀. MASON COUNTY: Vernal pool complex at RT 2/Lighthouse Gospel Church Road intersection, 38.82201 -82.13136; 17 March 2005 - (WLU 05031707), 1 I♂, 1 ♀; 28 March 2005 - (WLU 05032801), 1 I♂, 3 ♀. Pin oak swamp adjacent to Point Pleasant Moose Lodge in Wagner, 38.833603 -82.12227; 26 February 2004 - (WLU 04022601), 1 II♂, 1 O♀; 26, March 2004 - (WLU 04032601), 3 I♂, 2 ♀; 30 March 2004 - (WLU 04033001), 1 I♂; 12 April 2004 - (WLU 04041202), 1 I♂; 28 April 2004 - (WLU 04042801), 1 II♂, 2 ♀.</paragraph>
<paragraph id="E7ECF66B8CB4AFD93DCAC744780C7D92" pageId="25" pageNumber="26">CABELL COUNTY: Green Bottom Swamp at Hoeft Marsh Wildlife Management Area, 38.58616 -82.24878; 2 April 2009 - (WLU 09040201), 2 I♂, 2 II♂, 1 O♀, 4♀. MASON COUNTY: Vernal pool complex at RT 2/Lighthouse Gospel Church Road intersection, 38.82201 -82.13136; 17 March 2005 - (WLU 05031707), 1 I♂, 1 ♀; 28 March 2005 - (WLU 05032801), 1 I♂, 3 ♀. Pin oak swamp adjacent to Point Pleasant Moose Lodge in Wagner, 38.833603 -82.12227; 26 February 2004 - (WLU 04022601), 1 II♂, 1 O♀; 26, March 2004 - (WLU 04032601), 3 I♂, 2 ♀; 30 March 2004 - (WLU 04033001), 1 I♂; 12 April 2004 - (WLU 04041202), 1 I♂; 28 April 2004 - (WLU 04042801), 1 II♂, 2 ♀.</paragraph>
</subSubSection>
<subSubSection lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="distribution">
<paragraph pageId="25" pageNumber="26">Distribution.</paragraph>
<paragraph pageId="25" pageNumber="26">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<subSubSection id="F2F52063A2A5A356A5677DBA6F821B94" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="distribution">
<paragraph id="1DFE7F43F537788F9D4513A50BDE2815" pageId="25" pageNumber="26">Distribution.</paragraph>
<paragraph id="4CE9A1D6870CB6DEEC5287546ECE06A1" pageId="25" pageNumber="26">
<taxonomicName id="71F5B59CA1FAF349266516DA8B5D54D8" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
is a wide-ranging species occurring along the Atlantic Slope in Maryland, Virginia, and North Carolina east to the Ohio and Mississippi river valleys excluding the majority of the Appalachian Mountains. Northern populations occur along the Great Lakes and in southern Ontario (
<bibRefCitation author="Jezerinac, RF" journalOrPublisher="Ohio Biological Survey Bulletin" pageId="58" pageNumber="59" pagination="1 - 193" title="The Crayfishes (Decapoda: Cambaridae) of West Virginia" volume="10" year="1995">Jezerinac et al. 1995</bibRefCitation>
<bibRefCitation id="B130A7107A8AA4CFAB09DC35C066CF47" author="Jezerinac, RF" journalOrPublisher="Ohio Biological Survey Bulletin" pageId="58" pageNumber="59" pagination="1 - 193" title="The Crayfishes (Decapoda: Cambaridae) of West Virginia" volume="10" year="1995">Jezerinac et al. 1995</bibRefCitation>
).
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="8BDE0CF1E6A2E10036C2030D38C1550F" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
were collected from the Lower Ohio Basin and Middle Ohio South Basin (Figure 16). Within the Middle Ohio South, all populations were within one km of the Middle Ohio South/Lower Kanawha basin border. Two previously unknown populations were discovered in Mason County including one adjacent to the Point Pleasant Moose Lodge in Wagner and another in Krodel Park, Point Pleasant. Increased efforts to find this species along the floodplain throughout the Middle Ohio South, Middle Ohio North and Lower Ohio basins were futile.
</paragraph>
<caption pageId="25" pageNumber="26">
<paragraph pageId="25" pageNumber="26">
<caption id="F8DCCBAF5C6BBDF98BC32502E6196764" pageId="25" pageNumber="26">
<paragraph id="8EED9DE0AE6556A5F71D99912B82C1DB" pageId="25" pageNumber="26">
Figure 16.
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="2601EED039B1AE59B75AC68B8CB7E890" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
and
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Procambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Procambarus acutus" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="acutus">Procambarus acutus</taxonomicName>
<taxonomicName id="946CB4237B93F0C28991BE1FF719C920" class="Malacostraca" family="Cambaridae" genus="Procambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Procambarus acutus" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="acutus">Procambarus acutus</taxonomicName>
distribution along the West Virginia portion of the Ohio River floodplain
</paragraph>
</caption>
<paragraph lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<bibRefCitation author="Jezerinac, RF" journalOrPublisher="Ohio Journal of Science" pageId="58" pageNumber="59" pagination="46 - 47" title="Fallicambarus (Creaserinus) fodiens in West Virginia: a new state record." volume="87" year="1987">Jezerinac and Stocker (1987)</bibRefCitation>
<paragraph id="6D19338A431FCB2AE1F0B77D7958CB73" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<bibRefCitation id="E3A3E18CF423DC0F94B52F097ED3EF60" author="Jezerinac, RF" journalOrPublisher="Ohio Journal of Science" pageId="58" pageNumber="59" pagination="46 - 47" title="Fallicambarus (Creaserinus) fodiens in West Virginia: a new state record." volume="87" year="1987">Jezerinac and Stocker (1987)</bibRefCitation>
theorized that
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="3B72C66790417608F49FBB22E3ADD420" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
populations in West Virginia were Marietta River relicts. Currently, the Marietta River Valley is composed of the Kanawha River Valley. This hypothesis does help explain the scarcity of this species along the floodplain. Future survey efforts for
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="66C015FE7B3A330EABCEE27D7694C697" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
should focus on wetlands associated with the Kanawha River floodplain. The Moose Lodge wetland and its associated wetlands include that floodplain, which is 0.8 km from the Kanawha River and Ohio River confluence. The Moose Lodge wetland is composed of diverse bottomland forest with multiple ephemeral pools, which possess stable
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="BA96E7E02F771505D41F8117BD403372" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
populations.
<pageBreakToken pageId="26" pageNumber="27" start="start">Habitat</pageBreakToken>
<pageBreakToken id="DCC33C6557B630A713F288B710A21A14" pageId="26" pageNumber="27" start="start">Habitat</pageBreakToken>
specialization may explain low
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="50B7BBD602FE1FC2C4702A0765C85D61" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
numbers elsewhere since little mature bottomland forests remain along the Ohio River floodplain. Apparently this habitat is needed for West
<normalizedToken originalValue="Virginia’s">Virginia's</normalizedToken>
<normalizedToken id="17D3790CE63C220A32A723683C3B3DF9" originalValue="Virginia’s">Virginia's</normalizedToken>
disjunct population of
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="4868DCB5E7B7B7C06638B33F58740F50" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
to persist.
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="morphometrics">
<paragraph pageId="26" pageNumber="27">Morphometrics.</paragraph>
<paragraph pageId="26" pageNumber="27">
<subSubSection id="9E175E0BEAFEC18EE988B1654C6B145F" pageId="26" pageNumber="27" type="morphometrics">
<paragraph id="1C1242DD612CAFD48EDD2123BA75BBCE" pageId="26" pageNumber="27">Morphometrics.</paragraph>
<paragraph id="7770BB9DBC8E907C64F9BC29FE0F29B4" pageId="26" pageNumber="27">
Several animals were observed, but not disturbed due to the rarity of this species in West Virginia. The largest individual was a 40.1 mm TCL form I male collected at the Moose Lodge wetland, Mason County. The largest female collected was 37.3 mm TCL from an ephemeral pool complex located inside Krodel Park, Mason County. Mean carapace length for this species was 34.3 mm (n = 26, SE = 8.67). Morphometrics data for
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="A2B3AAC79F58B693A971B03038D4AFEE" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
is presented in Table 7.
</paragraph>
<caption ID-Table-UUID="25B4F1517D62A6793A00732CCD4BF6CD" httpUri="http://table.plazi.org/id/25B4F1517D62A6793A00732CCD4BF6CD" pageId="26" pageNumber="27">
<paragraph pageId="26" pageNumber="27">
<caption id="25B4F1517D62A6793A00732CCD4BF6CD" ID-Table-UUID="25B4F1517D62A6793A00732CCD4BF6CD" httpUri="http://table.plazi.org/id/25B4F1517D62A6793A00732CCD4BF6CD" pageId="26" pageNumber="27">
<paragraph id="5D66B18085C2D97D909B4C84209D72D3" pageId="26" pageNumber="27">
Table 7. West Virginia Ohio River floodplain
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="F0E8EFF79EB5017DD638DA9FDB5173B7" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
morphometrics.
</paragraph>
</caption>
<paragraph pageId="26" pageNumber="27">
<table pageId="26" pageNumber="27">
<tr pageId="26" pageNumber="27">
<th colspan="1" pageId="26" pageNumber="27" rowspan="1">Sex</th>
<th colspan="1" pageId="26" pageNumber="27" rowspan="1">N</th>
<th colspan="1" pageId="26" pageNumber="27" rowspan="1">Minimum</th>
<th colspan="1" pageId="26" pageNumber="27" rowspan="1">Maximum</th>
<th colspan="1" pageId="26" pageNumber="27" rowspan="1">Mean</th>
<th colspan="1" pageId="26" pageNumber="27" rowspan="1">Standard Deviation</th>
<paragraph id="4FE64A433DDB7266433E6D1277E80B1B" pageId="26" pageNumber="27">
<table id="E49E2311078F7B7AA43652542BFF3E3D" pageId="26" pageNumber="27">
<tr id="EB53E01E9395D39EFF3F75E5A963067E" pageId="26" pageNumber="27">
<th id="28861C033DAD5C78D3C7A8DB433824CF" colspan="1" pageId="26" pageNumber="27" rowspan="1">Sex</th>
<th id="2361D82D612E4F5D30ED25BA3264EC28" colspan="1" pageId="26" pageNumber="27" rowspan="1">N</th>
<th id="E3BE5ACE611838C7B7323CBD7ECE4F27" colspan="1" pageId="26" pageNumber="27" rowspan="1">Minimum</th>
<th id="7F14CF871EF1A28A13730D1A0A6278D8" colspan="1" pageId="26" pageNumber="27" rowspan="1">Maximum</th>
<th id="D08FC6EE592A77A802DAA4F7DEE52D88" colspan="1" pageId="26" pageNumber="27" rowspan="1">Mean</th>
<th id="AA1EDA239DAB39727A4733E3E2825376" colspan="1" pageId="26" pageNumber="27" rowspan="1">Standard Deviation</th>
</tr>
</table>
</paragraph>
<caption pageId="26" pageNumber="27">
<paragraph pageId="26" pageNumber="27">
<caption id="A3719F7545A00300B6281CEB080D4051" pageId="26" pageNumber="27">
<paragraph id="EBF75D262F6C97FA37CCF5454FF3A88D" pageId="26" pageNumber="27">
Figure 17.
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="C4BC273A2BDEFBB8F524BC882EE74D8E" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
, Middle Ohio South basin, Mason County, West Virginia - WLU 05031707
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="habitat and natural history">
<paragraph pageId="26" pageNumber="27">Habitat and natural history.</paragraph>
<paragraph lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<subSubSection id="874F555C538F4DB53FFDDB7E2F3BC7A5" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="habitat and natural history">
<paragraph id="5E8F7DD9B97AE85329A6B687E1F108DB" pageId="26" pageNumber="27">Habitat and natural history.</paragraph>
<paragraph id="167362AD2FC5FEFD6816E0917480759D" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<taxonomicName id="A7C07DA813C3388B01793BA4804DB033" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
(Figure 18) in West Virginia is an ephemeral pool specialist. Within these ecosystems, colonies were associated with
<pageBreakToken pageId="27" pageNumber="28" start="start">either</pageBreakToken>
<pageBreakToken id="64B9189F56210311433D9401E8D57409" pageId="27" pageNumber="28" start="start">either</pageBreakToken>
lowland forest environments or open, wet fields. Similar habitat preferences for this species has been observed across its range (
<bibRefCitation author="Guiasu, RC" journalOrPublisher="Crustaceana" pageId="57" pageNumber="58" pagination="207 - 223" title="Conservation and diversity of the crayfishes of the genus Fallicambarus, with an emphasis on the status of Fallicambarus fodiens in Canada." volume="80" year="2007">Guiasu 2007</bibRefCitation>
<bibRefCitation id="EBC11EB934509DEBC98A82D8DD8E1DD3" author="Guiasu, RC" journalOrPublisher="Crustaceana" pageId="57" pageNumber="58" pagination="207 - 223" title="Conservation and diversity of the crayfishes of the genus Fallicambarus, with an emphasis on the status of Fallicambarus fodiens in Canada." volume="80" year="2007">Guiasu 2007</bibRefCitation>
;
<bibRefCitation author="Norrocky, MJ" journalOrPublisher="American Midland Naturalist" pageId="58" pageNumber="59" pagination="75 - 86" title="Observations on the ecology, reproduction and growth of the burrowing crayfish Fallicambarus (C.) fodiens in north-central Ohio." volume="125" year="1991">Norrocky 1991</bibRefCitation>
<bibRefCitation id="44EB13CA581E172E9C2FA58E547DBD84" author="Norrocky, MJ" journalOrPublisher="American Midland Naturalist" pageId="58" pageNumber="59" pagination="75 - 86" title="Observations on the ecology, reproduction and growth of the burrowing crayfish Fallicambarus (C.) fodiens in north-central Ohio." volume="125" year="1991">Norrocky 1991</bibRefCitation>
; Taylor and Schuster, 2004). With the exception of the Greenbottom Swamp population, ephemeral systems are preferred over larger, more permanent water bodies.
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="CF8BDEC95284B965A60E7696927ECF10" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
colonies typically consist of 5-10 burrows for every 1 m2 of substrate.
<pageBreakToken pageId="28" pageNumber="29" start="start">Within</pageBreakToken>
<pageBreakToken id="E4AE5452A6ABA6A60BE854E8963EF711" pageId="28" pageNumber="29" start="start">Within</pageBreakToken>
these colonies, the sex ratio of captured individuals was 1:1 male to female.
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="538CAE44DA1D9DF427B186FE7AED6032" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
burrow morphology is simple, with the majority of excavated burrows consisting of a central shaft ranging in depth from 0.3 m to 1.0 m. One or two short ancillary tunnels often were present radiating from central shafts. These ancillary tunnels often are full of debris. Resting chambers usually were present at the terminus of these shafts, with either few or no ancillary tunnels radiating from them.
</paragraph>
<caption pageId="28" pageNumber="29">
<paragraph pageId="28" pageNumber="29">
<caption id="1CEE5D673F3527FD2D4B467127A39674" pageId="28" pageNumber="29">
<paragraph id="1289E2386EBBA82BBDBB211FE078FD4B" pageId="28" pageNumber="29">
Figure 18. Ovigerous
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="3F4999B7A5B30F03EDD03CF28E4C3D87" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
, Middle Ohio South basin, Mason County, West Virginia. Ovigerous females were prevalent in surface waters from late February through early March. Females with pleopodal instars were observed from mid March through early May. The pictured specimen was released after capture.
</paragraph>
</caption>
<paragraph pageId="28" pageNumber="29">
<paragraph id="3D91AB557C6A2ADB853E4B2CD2514816" pageId="28" pageNumber="29">
Several
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="E07584EEABAAD3EB7B2D38563A34CA9A" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
females (n = 7) were collected carrying instars and eggs in February and March 2004 and 2005 (Figure 18). Egg extrusion along the floodplain occurred throughout the months of February and March. By early April, females were carrying first stage instars, with 4th stage instars observed by late April.
<bibRefCitation author="Jezerinac, RF" journalOrPublisher="Ohio Biological Survey Bulletin" pageId="58" pageNumber="59" pagination="1 - 193" title="The Crayfishes (Decapoda: Cambaridae) of West Virginia" volume="10" year="1995">Jezerinac et al. (1995)</bibRefCitation>
<bibRefCitation id="D4DF91E0D52E70A4F53DE5B62A0E32CF" author="Jezerinac, RF" journalOrPublisher="Ohio Biological Survey Bulletin" pageId="58" pageNumber="59" pagination="1 - 193" title="The Crayfishes (Decapoda: Cambaridae) of West Virginia" volume="10" year="1995">Jezerinac et al. (1995)</bibRefCitation>
also observed ovigerous females in February and March in West Virginia. Ovigerous females have been collected February through April in Illinois (
<bibRefCitation author="Page, LM" journalOrPublisher="Bulletin Illinois Natural History Survey" pageId="58" pageNumber="59" pagination="335 - 448" title="The crayfishes and shrimp of Illinois." volume="33" year="1985">Page 1985</bibRefCitation>
<bibRefCitation id="1FB01A2D843F8F684DFE796450FC9DCA" author="Page, LM" journalOrPublisher="Bulletin Illinois Natural History Survey" pageId="58" pageNumber="59" pagination="335 - 448" title="The crayfishes and shrimp of Illinois." volume="33" year="1985">Page 1985</bibRefCitation>
), Kentucky (
<bibRefCitation author="Taylor, CA" journalOrPublisher="Illinois Natural History Survey Bulletin" pageId="59" pageNumber="60" title="Crayfishes of Kentucky" volume="28" year="2004">Taylor and Schuster 2004</bibRefCitation>
<bibRefCitation id="9DCC76B5DB8E9291E7D811EABD71500B" author="Taylor, CA" journalOrPublisher="Illinois Natural History Survey Bulletin" pageId="59" pageNumber="60" title="Crayfishes of Kentucky" volume="28" year="2004">Taylor and Schuster 2004</bibRefCitation>
), and Michigan (
<bibRefCitation author="Creaser, EP" journalOrPublisher="Papers of the Michigan Academy of Sciences Arts and Letters" pageId="56" pageNumber="57" pagination="257 - 276" title="The Michigan decapods crustaceans." volume="13" year="1931">Creaser 1931</bibRefCitation>
<bibRefCitation id="EB240785A53BABC8D16FEA7CE638935D" author="Creaser, EP" journalOrPublisher="Papers of the Michigan Academy of Sciences Arts and Letters" pageId="56" pageNumber="57" pagination="257 - 276" title="The Michigan decapods crustaceans." volume="13" year="1931">Creaser 1931</bibRefCitation>
). All adult males captured during this study were form I. Given that females were ovigerous in early spring and males enter the winter season as form I, mating in this species likely occurs in the fall or winter along the floodplain.
</paragraph>
<paragraph pageId="28" pageNumber="29">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<paragraph id="55B09939BA17A3B680743CC63D2F1794" pageId="28" pageNumber="29">
<taxonomicName id="C1562670F5162B0827EB6D4B20B23317" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
females carrying instars were observed nocturnally foraging in open water on 18 March 2004 and 4 April 2009 in Greenbottom Wildlife Management Area. Several individuals (n = 8) were resting or grazing at their burrow entrances on periphyton that had colonized submerged canary grass (
<taxonomicName class="Liliopsida" family="Poaceae" genus="Phalaris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Phalaris canariensis" order="Poales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="canariensis">Phalaris canariensis</taxonomicName>
<taxonomicName id="A1D4225927F0B1C56EE74F5640BDA042" class="Liliopsida" family="Poaceae" genus="Phalaris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Phalaris canariensis" order="Poales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="canariensis">Phalaris canariensis</taxonomicName>
L.). Within the colony burrows were flooded by 15-30 cm of standing water. Instars were observed leaving burrow entrances to graze. Upon provocation, disturbed females stopped moving long enough to allow instars to reattach to their abdomens, then retreated to their burrows. Seasonal data for
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="6F8CB94EC60401F78CBD8D89CCBD949B" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
is presented in Table 3. Crayfish associates collected with
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<taxonomicName id="3C57B4853A5D1AAB0FA283EFFF1C4ED8" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
include
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus bartonii subsp. cavatus" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="bartonii" subSpecies="cavatus">Cambarus bartonii cavatus</taxonomicName>
<taxonomicName id="4D39EDDA1DFB0846D2C418B4DB7978C0" class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus bartonii subsp. cavatus" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="bartonii" subSpecies="cavatus">Cambarus bartonii cavatus</taxonomicName>
,
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus thomai" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="thomai">Cambarus thomai</taxonomicName>
<taxonomicName id="DE1B80A6C349B729E8D86065349DF9D9" class="Malacostraca" family="Cambaridae" genus="Cambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cambarus thomai" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="thomai">Cambarus thomai</taxonomicName>
,
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Orconectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orconectes virilis" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="virilis">Orconectes virilis</taxonomicName>
<taxonomicName id="746824AACBC2FDAC80151AC74F1D17ED" class="Malacostraca" family="Cambaridae" genus="Orconectes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orconectes virilis" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="virilis">Orconectes virilis</taxonomicName>
, and
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Procambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Procambarus acutus subsp. acutus" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="acutus" subSpecies="acutus">Procambarus acutus acutus</taxonomicName>
<taxonomicName id="65AC6018BCD55A6E04AC5548B5B10265" class="Malacostraca" family="Cambaridae" genus="Procambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Procambarus acutus subsp. acutus" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="acutus" subSpecies="acutus">Procambarus acutus acutus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="conservation status within study area">
<paragraph pageId="28" pageNumber="29">Conservation status within study area.</paragraph>
<paragraph pageId="28" pageNumber="29">
<taxonomicName class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
<subSubSection id="77227DA3C8F423F706AABFF614432573" pageId="28" pageNumber="29" type="conservation status within study area">
<paragraph id="BB2F73AA41DD26B0A859821D2EAFE214" pageId="28" pageNumber="29">Conservation status within study area.</paragraph>
<paragraph id="11DD29776911EE2C9A0A9B67483387E6" pageId="28" pageNumber="29">
<taxonomicName id="F144C791938536C9E85C1E9B90A0ADCD" class="Malacostraca" family="Cambaridae" genus="Fallicambarus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fallicambarus fodiens" order="Decapoda" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="fodiens">Fallicambarus fodiens</taxonomicName>
warrants conservation attention and is deserving of S1 status. Surveys are needed along the Kanawha River in wetland habitats to determine if this species persists there. Along the floodplain threats to this
<normalizedToken originalValue="species’">species'</normalizedToken>
<normalizedToken id="A7FD342F705CA3198C2488D821653B63" originalValue="species’">species'</normalizedToken>
survival include land use practices and their associated pollutants, hard surface run off, and destruction of bottomland forests.
</paragraph>
</subSubSection>

90
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Lepidium ruderale
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Fam.
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Passerage des
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Stengel 10-40 cm hoch, aufrecht, besonders oben verzweigt, sehr kurz behaart. Untere
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bis auf den Mittelnerv 1 -3fach fiederteilig mit sehr schmalen Abschnitten,
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absterbend. Obere
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Juni-Oktober. Standort:
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, Bahnareale, Schuttstellen, kollin-montan (-subalpin). Sehr vereinzelt in allen Kantonsteilen (3, 7L).
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">Fig. 7A-E</figureCitation>
<taxonomicNameLabel id="B44C99BDAD61E63528A870694D43444D" pageId="0" pageNumber="137">sp. nov.</taxonomicNameLabel>
<figureCitation id="4449B052BD6BBDCBEB210D3EEAA1C6E4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">Fig. 7A-E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="diagnosis">
<paragraph pageId="0" pageNumber="137">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="137">
<taxonomicName authorityName="Padua, Saeaeksjaervi &amp; Spasojevic" authorityYear="2021" class="Insecta" family="Ichneumonidae" genus="Polysphincta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polysphincta pichincha" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="pichincha">
<emphasis italics="true" pageId="0" pageNumber="137">Polysphincta pichincha</emphasis>
<subSubSection id="B90D5DED2AA2898FF716D3B5887D0EE8" pageId="0" pageNumber="137" type="diagnosis">
<paragraph id="9608562D69F2025FAAA6D4021302F7C2" pageId="0" pageNumber="137">Diagnosis.</paragraph>
<paragraph id="93540087C5D86892FF76CE3B257FA001" pageId="0" pageNumber="137">
<taxonomicName id="6CAFFF40847417E79100557F0D210C22" authorityName="Padua, Saeaeksjaervi &amp; Spasojevic" authorityYear="2021" class="Insecta" family="Ichneumonidae" genus="Polysphincta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polysphincta pichincha" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="pichincha">
<emphasis id="ABE2CA0449819391183B66E28C24AEE2" italics="true" pageId="0" pageNumber="137">Polysphincta pichincha</emphasis>
</taxonomicName>
sp. nov. can be distinguished from other species of the
<taxonomicName genus="P." lsidName="P. dizardi" pageId="0" pageNumber="137" rank="species" species="dizardi">
<emphasis italics="true" pageId="0" pageNumber="137">P. dizardi</emphasis>
<taxonomicName id="7983A237D2722C6B35A4B3B5ECB65CF7" lsidName="P. dizardi" pageId="0" pageNumber="137" rank="species" species="dizardi">
<emphasis id="9DBF82C9EF5C441722F6CBC1EC214D88" italics="true" pageId="0" pageNumber="137">P. dizardi</emphasis>
</taxonomicName>
species-group by the combination of the following characters: (1) epomia absent (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7B</figureCitation>
<figureCitation id="1C07C67B289D308813D9384D2A591E72" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7B</figureCitation>
); (2) malar space 0.4 times as long as proximal mandibular width (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7B</figureCitation>
<figureCitation id="EC686D939B7F37E0510230EA0CCE837A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7B</figureCitation>
); (3) fore wing vein 1
<emphasis italics="true" pageId="0" pageNumber="137">cu-a</emphasis>
<emphasis id="24B31A49DD4EBD2322E7A0E856921AC9" italics="true" pageId="0" pageNumber="137">cu-a</emphasis>
more or less interstitial relative to
<emphasis italics="true" pageId="0" pageNumber="137">M&amp;RS</emphasis>
<emphasis id="C62D7A2C10812221F72FB0AE8E601E81" italics="true" pageId="0" pageNumber="137">M&amp;RS</emphasis>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A</figureCitation>
<figureCitation id="999CB2F36E0DA80F9B62B4405318B06A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A</figureCitation>
); (4) mesosoma entirely orange (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A</figureCitation>
<figureCitation id="C2D3AC444A75BF1CCBEE7196549879A4" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A</figureCitation>
); (5) wings yellowish hyaline (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A</figureCitation>
<figureCitation id="0AD6505533857A5E7B1EE6B36CC68697" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A</figureCitation>
); (6) hind leg orange with trochanter, apex distal and proximal of femur, tibia, except longitudinal spot pale in subdistal region of inner and outer margin and tarsus brownish (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A</figureCitation>
<figureCitation id="FFE35E958CCFC9C46170824B4DEB3187" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A</figureCitation>
); (7) metasoma orange, with posterior margins of tergites II-IV narrowly black, tergites V+ black (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A, C</figureCitation>
<figureCitation id="F0D9F4A3012FE6FD72E69EDEEFE62D31" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Polysphincta pichincha sp. nov., ♀, holotype A habitus, lateral view B head and pronotum, lateral view C habitus, dorsal view D face, anterior view E head and shelf-like projection, dorsal view." figureDoi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137">7A, C</figureCitation>
); (8) ovipositor slender, 1.2 times as long as hind tibia.
</paragraph>
<caption doi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Figure 7.</emphasis>
<taxonomicName authorityName="Padua, Saeaeksjaervi &amp; Spasojevic" authorityYear="2021" class="Insecta" family="Ichneumonidae" genus="Polysphincta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polysphincta pichincha" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="pichincha">
<emphasis italics="true" pageId="0" pageNumber="137">Polysphincta pichincha</emphasis>
<caption id="3DB23DF8BC57F0C39CE56B0BEA42126F" doi="10.3897/zookeys.1041.65407.figure7" httpUri="https://binary.pensoft.net/fig/550748" pageId="0" pageNumber="137" start="Figure 7" startId="F7">
<paragraph id="37CB633ED7EE5570E732558AE1D7A862" pageId="0" pageNumber="137">
<emphasis id="1F89D5D6CED1C885C1FB2EB1EBD33FF2" bold="true" pageId="0" pageNumber="137">Figure 7.</emphasis>
<taxonomicName id="B9F0DC57C51F14114FB1166D9ECF2B9B" authorityName="Padua, Saeaeksjaervi &amp; Spasojevic" authorityYear="2021" class="Insecta" family="Ichneumonidae" genus="Polysphincta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polysphincta pichincha" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="pichincha">
<emphasis id="E35A782EE6CFF2F6D66FA028D93B8E48" italics="true" pageId="0" pageNumber="137">Polysphincta pichincha</emphasis>
</taxonomicName>
sp. nov., ♀, holotype
<emphasis bold="true" pageId="0" pageNumber="137">A</emphasis>
<emphasis id="A2B9F1039295D41A97A3BF16C930FEBC" bold="true" pageId="0" pageNumber="137">A</emphasis>
habitus, lateral view
<emphasis bold="true" pageId="0" pageNumber="137">B</emphasis>
<emphasis id="0A77A621E7A40E83EB892B38B93BE33E" bold="true" pageId="0" pageNumber="137">B</emphasis>
head and pronotum, lateral view
<emphasis bold="true" pageId="0" pageNumber="137">C</emphasis>
<emphasis id="D51B65948FB1F6B1154354F36DF32E81" bold="true" pageId="0" pageNumber="137">C</emphasis>
habitus, dorsal view
<emphasis bold="true" pageId="0" pageNumber="137">D</emphasis>
<emphasis id="3C07C50B941A42BDB04D9302E64FC84B" bold="true" pageId="0" pageNumber="137">D</emphasis>
face, anterior view
<emphasis bold="true" pageId="0" pageNumber="137">E</emphasis>
<emphasis id="E252B4F7C398AB3149A51CD82F2EE9F1" bold="true" pageId="0" pageNumber="137">E</emphasis>
head and shelf-like projection, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="description">
<paragraph pageId="0" pageNumber="137">Description.</paragraph>
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Female.</emphasis>
<subSubSection id="053188BE86554402E03470AD7A3CD148" pageId="0" pageNumber="137" type="description">
<paragraph id="C5137D5F49CDDA9181559AEDA62AFA75" pageId="0" pageNumber="137">Description.</paragraph>
<paragraph id="CAC1ADBEAE856E49F6D0E5CFB16366F7" pageId="0" pageNumber="137">
<emphasis id="F2056CE011F024CF05F4D508C4A73769" bold="true" pageId="0" pageNumber="137">Female.</emphasis>
Body [13.5] mm.
<emphasis bold="true" pageId="0" pageNumber="137">
<emphasis italics="true" pageId="0" pageNumber="137">Head</emphasis>
<emphasis id="918DEFC447168DED51AB16D0FF4C44D1" bold="true" pageId="0" pageNumber="137">
<emphasis id="6E19099850CEA69E8BB540BBF3137576" italics="true" pageId="0" pageNumber="137">Head</emphasis>
.
</emphasis>
Clypeus weakly convex, posterior margin thin and flat centrally; malar space [0.4] times as long as proximal mandibular width; lower face about [1.0] times as broad as high, weakly convex centrally, polished, with fine sparse setiferous punctures; head in dorsal view with margin of gena flat behind eyes, and its margin about [0.5] times length of eye; ocelli moderately large, lateral ones separated from compound eyes by [0.85] times their own maximum diameter.
<emphasis bold="true" pageId="0" pageNumber="137">
<emphasis italics="true" pageId="0" pageNumber="137">Mesosoma</emphasis>
<emphasis id="65357E7EB88FCDEE435CC41CC9B74098" bold="true" pageId="0" pageNumber="137">
<emphasis id="61F91A2EAD47C9884A5082E962C03391" italics="true" pageId="0" pageNumber="137">Mesosoma</emphasis>
.
</emphasis>
Pronotum without epomia; shelf-like projection, in dorsal view, more or less bilobed, broader than long, and, in lateral view, slender and with anterolateral corners weakly decurved; mesoscutum slender, in dorsal view, smooth and polished, with notauli weakly impressed anteriorly; scutellum convex, not laterally carinate; mesopleuron highly polished, virtually impunctate; epicnemial carina reaching almost level of lower corner of pronotum; epicnemium with vestigial vertical carina near lower corner of pronotum; metapleuron convex, smooth and polished, with few sparse, fine bristles evenly spaced, without discernible submetapleural carina. Propodeum mediodorsally smooth and polished, with longitudinal carinae present only posteriorly and laterally with scattered fine bristles. Fore wing length [10.0] mm; 1
<emphasis italics="true" pageId="0" pageNumber="137">cu-a</emphasis>
<emphasis id="3C95490E6DAC0B315A87D36654335215" italics="true" pageId="0" pageNumber="137">cu-a</emphasis>
more or less interstitial relative to
<emphasis italics="true" pageId="0" pageNumber="137">M&amp;RS</emphasis>
<emphasis id="BBAF6BA5A1EDA38662478DFA9D21DAC0" italics="true" pageId="0" pageNumber="137">M&amp;RS</emphasis>
; base of 1
<emphasis italics="true" pageId="0" pageNumber="137">m-cu&amp;M</emphasis>
<emphasis id="0CCE7D6E30CAB1B49DB8F60705415011" italics="true" pageId="0" pageNumber="137">m-cu&amp;M</emphasis>
separated from
<emphasis italics="true" pageId="0" pageNumber="137">CU</emphasis>
<emphasis id="AF5BF183CC6CAB0F6F9DEB9E9909A586" italics="true" pageId="0" pageNumber="137">CU</emphasis>
by more than length of 2
<emphasis italics="true" pageId="0" pageNumber="137">cu-a</emphasis>
<emphasis id="FBF7D89BEB8A88B0BD25D9686DF3177F" italics="true" pageId="0" pageNumber="137">cu-a</emphasis>
; hind wing with distal abscissa of
<emphasis italics="true" pageId="0" pageNumber="137">CU</emphasis>
<emphasis id="28207D4C23B3C4906AE43F23DEEC8409" italics="true" pageId="0" pageNumber="137">CU</emphasis>
present and complete; first abscissa of
<emphasis italics="true" pageId="0" pageNumber="137">RS</emphasis>
<emphasis id="E479A49FE1C212B37B5C8E85BD0F11D7" italics="true" pageId="0" pageNumber="137">RS</emphasis>
subequal to
<emphasis italics="true" pageId="0" pageNumber="137">rs-m</emphasis>
<emphasis id="7A248A514DE4EE6739C04D66852B91E7" italics="true" pageId="0" pageNumber="137">rs-m</emphasis>
. Tarsal claw with proximal lobe quadrangular, with claw apex slightly overtaking the distal margin of lobe.
<emphasis bold="true" pageId="0" pageNumber="137">
<emphasis italics="true" pageId="0" pageNumber="137">Metasoma</emphasis>
<emphasis id="9A018166C0A966AECDC6158486BE4FAE" bold="true" pageId="0" pageNumber="137">
<emphasis id="E6C2DAD28F0710468FB9174A0D5DF2C7" italics="true" pageId="0" pageNumber="137">Metasoma</emphasis>
.
</emphasis>
Tergite I about [1.8] times as long as posteriorly broad, dorsally with lateromedian longitudinal carinae only discernible at extreme anterior part; sternite I with weak swelling near hind rim, and with weak median longitudinal ridge anteriorly; tergite II about [1.6] times as long as posteriorly broad, highly polished, at most with only fine setiferous punctures laterally; tergite III [1.4] times as long as posteriorly broad, highly polished, at most with only fine setiferous punctures laterally; subgenital plate subquadrate. Ovipositor slightly slender, about [1.2] times as long as hind tibia [without apex].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="color">
<paragraph pageId="0" pageNumber="137">Color.</paragraph>
<paragraph pageId="0" pageNumber="137">Head black, except posterior margin of clypeus whitish; antennae brown; mouthparts pale, except apex of mandible brownish. Mesosoma orange. Metasoma orange, with posterior margins of tergites II-IV narrowly black, tergites V+ black. Legs orange, the mid leg with tarsus brownish, hind leg with trochanter, apex distal and proximal of femur, tibia, except longitudinal spot pale in subdistal region of inner and outer margin and tarsus brownish. Wings yellowish hyaline, pterostigma yellow.</paragraph>
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" pageId="0" pageNumber="137">Male.</emphasis>
<subSubSection id="A126556C1B4FA6F6761A3A5B09B6AC79" pageId="0" pageNumber="137" type="color">
<paragraph id="C3D7E373097DB30D0086D3D56B9F2A97" pageId="0" pageNumber="137">Color.</paragraph>
<paragraph id="4AA603402D557B3915B12F925AEAB087" pageId="0" pageNumber="137">Head black, except posterior margin of clypeus whitish; antennae brown; mouthparts pale, except apex of mandible brownish. Mesosoma orange. Metasoma orange, with posterior margins of tergites II-IV narrowly black, tergites V+ black. Legs orange, the mid leg with tarsus brownish, hind leg with trochanter, apex distal and proximal of femur, tibia, except longitudinal spot pale in subdistal region of inner and outer margin and tarsus brownish. Wings yellowish hyaline, pterostigma yellow.</paragraph>
<paragraph id="5737420C4941A61F64AAD6D2C0E5467F" pageId="0" pageNumber="137">
<emphasis id="634DC0D5FCE83268FBB9941420C42631" bold="true" pageId="0" pageNumber="137">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="type material">
<paragraph pageId="0" pageNumber="137">Type material.</paragraph>
<paragraph pageId="0" pageNumber="137">
<emphasis bold="true" italics="true" pageId="0" pageNumber="137">Holotype</emphasis>
<subSubSection id="B745BCC452DFD71EF184B10F91586B8C" pageId="0" pageNumber="137" type="type material">
<paragraph id="E9802DAD350271932F56D111C6DB973B" pageId="0" pageNumber="137">Type material.</paragraph>
<paragraph id="41B18683AF3EE12AA5B5172D7FABA00A" pageId="0" pageNumber="137">
<emphasis id="796A84D550F194BD769BFCFF53352769" bold="true" italics="true" pageId="0" pageNumber="137">Holotype</emphasis>
♀. Ecuador, Pichincha, Nambillo Valley near Mindo, 1450 m, 26.VI.1987 (M. Cooper leg.), #2005-152, BMNH.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="distribution">
<paragraph pageId="0" pageNumber="137">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="137">
<subSubSection id="60CF87B9F21C269E5B70B0A04600814D" pageId="0" pageNumber="137" type="distribution">
<paragraph id="5C319A84516F17B68E538B9268C1F1E3" pageId="0" pageNumber="137">Distribution.</paragraph>
<paragraph id="8B84FEC17913145F6F5B7B292CE3CA43" pageId="0" pageNumber="137">
Ecuador (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of Polysphincta dizardi group in the Neotropical Region." figureDoi="10.3897/zookeys.1041.65407.figure11" httpUri="https://binary.pensoft.net/fig/550752" pageId="0" pageNumber="137">11</figureCitation>
<figureCitation id="82AE2108AF50131F6FAF739C921A302F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of Polysphincta dizardi group in the Neotropical Region." figureDoi="10.3897/zookeys.1041.65407.figure11" httpUri="https://binary.pensoft.net/fig/550752" pageId="0" pageNumber="137">11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="biological note">
<paragraph pageId="0" pageNumber="137">Biological note.</paragraph>
<paragraph pageId="0" pageNumber="137">Host unknown.</paragraph>
<subSubSection id="B982ECD53ADB82454F27E277D2BCD2A4" pageId="0" pageNumber="137" type="biological note">
<paragraph id="98CD2D44A0EF77FF89DA26B96054BD70" pageId="0" pageNumber="137">Biological note.</paragraph>
<paragraph id="AFADD29FEB89665427751C5534F7F72B" pageId="0" pageNumber="137">Host unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="etymology">
<paragraph pageId="0" pageNumber="137">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="137">The specific name (in apposition) refers to type locality of this species, Pichincha province, Ecuador.</paragraph>
<subSubSection id="02A3BD271A179BEF33E1C70D2ECA9739" pageId="0" pageNumber="137" type="etymology">
<paragraph id="542F9EB217154DE4BB30EDE1A75B688F" pageId="0" pageNumber="137">Etymology.</paragraph>
<paragraph id="BCEF5DF28D8F81D5BCDEA5933CDE3D49" pageId="0" pageNumber="137">The specific name (in apposition) refers to type locality of this species, Pichincha province, Ecuador.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="remarks">
<paragraph pageId="0" pageNumber="137">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="137">
<taxonomicName authorityName="Padua, Saeaeksjaervi &amp; Spasojevic" authorityYear="2021" class="Insecta" family="Ichneumonidae" genus="Polysphincta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polysphincta pichincha" order="Hymenoptera" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="pichincha">
<emphasis italics="true" pageId="0" pageNumber="137">Polysphincta pichincha</emphasis>
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<paragraph id="CACD02011E38F940C31802F8087161D6" pageId="0" pageNumber="137">Remarks.</paragraph>
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sp. nov. closely resembles
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<emphasis italics="true" pageId="0" pageNumber="137">A. sinearanea</emphasis>
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<emphasis id="7CA10B65C2884E4D213B38ACD078DB0A" italics="true" pageId="0" pageNumber="137">A. sinearanea</emphasis>
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<normalizedToken originalValue="Pádua">Padua</normalizedToken>
<normalizedToken id="A2B8293C3D92CE1B22587B5A769B15D8" originalValue="Pádua">Padua</normalizedToken>
, 2018 and
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<emphasis italics="true" pageId="0" pageNumber="137">P. inca</emphasis>
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<emphasis id="1300686420C88573EC6C207C94C47EC4" italics="true" pageId="0" pageNumber="137">P. inca</emphasis>
</taxonomicName>
sp. nov. mainly by coloration with body orange and the last tergites black. It differs from
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<emphasis italics="true" pageId="0" pageNumber="137">P. sinearanea</emphasis>
<taxonomicName id="C8F414B2EDDD411FD44B1DF412325196" lsidName="P. sinearanea" pageId="0" pageNumber="137" rank="species" species="sinearanea">
<emphasis id="B4061CFC1B1C8D83248E6B97B9E63206" italics="true" pageId="0" pageNumber="137">P. sinearanea</emphasis>
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by having epomia absent (present in
<taxonomicName genus="P." lsidName="P. sinearanea" pageId="0" pageNumber="137" rank="species" species="sinearanea">
<emphasis italics="true" pageId="0" pageNumber="137">P. sinearanea</emphasis>
<taxonomicName id="AF70B9D0BBFA7E350FC2052F74CC426C" lsidName="P. sinearanea" pageId="0" pageNumber="137" rank="species" species="sinearanea">
<emphasis id="1DA3D5D35102162D4430B9F99ACC2EC5" italics="true" pageId="0" pageNumber="137">P. sinearanea</emphasis>
</taxonomicName>
), and from
<taxonomicName genus="P." lsidName="P. inca" pageId="0" pageNumber="137" rank="species" species="inca">
<emphasis italics="true" pageId="0" pageNumber="137">P. inca</emphasis>
<taxonomicName id="2861D3FB61283CD10C52DBC66C23AF02" lsidName="P. inca" pageId="0" pageNumber="137" rank="species" species="inca">
<emphasis id="69A99B2A5A23DBB1846CFF34A8DC56E7" italics="true" pageId="0" pageNumber="137">P. inca</emphasis>
</taxonomicName>
sp. nov. by having malar space 0.4 times as long as proximal mandibular width (malar space&gt;0.6 times as long as proximal mandibular width in
<taxonomicName genus="P." lsidName="P. inca" pageId="0" pageNumber="137" rank="species" species="inca">
<emphasis italics="true" pageId="0" pageNumber="137">P. inca</emphasis>
<taxonomicName id="D47D1D6D8FCB5BD0492B305BE1EA0D50" lsidName="P. inca" pageId="0" pageNumber="137" rank="species" species="inca">
<emphasis id="A40C16F3FB5A6B8D5549D02AA02B598D" italics="true" pageId="0" pageNumber="137">P. inca</emphasis>
</taxonomicName>
sp. nov.).
</paragraph>

100
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<taxonomicName LSID="http://species-id.net/wiki/Liodessus_noviaffinis" authority="Miller, 1998" authorityName="Miller" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus noviaffinis" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="noviaffinis">Liodessus noviaffinis Miller, 1998</taxonomicName>
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Map 7
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<paragraph pageId="4" pageNumber="5">Material examined.</paragraph>
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New Brunswick, Saint John Co.,
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,
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, 28.V.2010, R. P. Webster, salt marsh, saline tidal pond (23, NBM, RWC).
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7. Collection localities in New Brunswick, Canada of
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.
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<paragraph pageId="5" pageNumber="6">Collection and habitat data.</paragraph>
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<bibRefCitation author="Miller, KB" journalOrPublisher="Entomologica Scandinavica" pageId="6" pageNumber="7" pagination="281 - 314" title="Revision of the Nearctic Liodessus affinis (Say 1823) species group (Coleoptera: Dytiscidae, Hydroporinae, Bidessini)." url="10.1163/187631298X00113" volume="29" year="1998">Miller (1998)</bibRefCitation>
<subSubSection id="A3DA2CA43C0DAD192F4630724191BC79" pageId="5" pageNumber="6" type="collection and habitat data">
<paragraph id="CFD7ABFB6E14952818A941A31036F71E" pageId="5" pageNumber="6">Collection and habitat data.</paragraph>
<paragraph id="164DF57D67AF175978CD7941933313ED" pageId="5" pageNumber="6">
<bibRefCitation id="20E5436BCD185219F35DA787DE4F21F8" author="Miller, KB" journalOrPublisher="Entomologica Scandinavica" pageId="6" pageNumber="7" pagination="281 - 314" title="Revision of the Nearctic Liodessus affinis (Say 1823) species group (Coleoptera: Dytiscidae, Hydroporinae, Bidessini)." url="10.1163/187631298X00113" volume="29" year="1998">Miller (1998)</bibRefCitation>
reported that this species occurs in coastal ponds and may be somewhat halophilic. The specimens from New Brunswick were collected during late May from saline tidal ponds and pools near the margin of a salt marsh. Adults were abundant, along with numerous salt marsh mosquito larvae.
</paragraph>
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<subSubSection pageId="5" pageNumber="6" type="distribution">
<paragraph pageId="5" pageNumber="6">Distribution in Canada and Alaska.</paragraph>
<paragraph pageId="5" pageNumber="6">
<subSubSection id="88AC2AEA566B1CB62A4DBE4066CFB357" pageId="5" pageNumber="6" type="distribution">
<paragraph id="927159C236454D6A0E02B6F22189284B" pageId="5" pageNumber="6">Distribution in Canada and Alaska.</paragraph>
<paragraph id="F40C8835A495E0C2E1049BA297B33224" pageId="5" pageNumber="6">
NB, NS (
<bibRefCitation author="Larson, DJ" journalOrPublisher="Monographs in Biodiversity, NRC Press, Ottawa, Ontario" pageId="6" pageNumber="7" title="Predaceous diving beetles (Coleoptera: Dytiscidae) of the nearctic region, with emphasis on the fauna of Canada and Alaska." year="2000">Larson et al. 2000</bibRefCitation>
<bibRefCitation id="C7A1CBE802A5FF4EB4117F2207652B05" author="Larson, DJ" journalOrPublisher="Monographs in Biodiversity, NRC Press, Ottawa, Ontario" pageId="6" pageNumber="7" title="Predaceous diving beetles (Coleoptera: Dytiscidae) of the nearctic region, with emphasis on the fauna of Canada and Alaska." year="2000">Larson et al. 2000</bibRefCitation>
). The determination was based on dissected individuals. This species was previously known from Canada from coastal habitats on Cape Breton Island, Nova Scotia (
<bibRefCitation author="Larson, DJ" journalOrPublisher="Monographs in Biodiversity, NRC Press, Ottawa, Ontario" pageId="6" pageNumber="7" title="Predaceous diving beetles (Coleoptera: Dytiscidae) of the nearctic region, with emphasis on the fauna of Canada and Alaska." year="2000">Larson et al. 2000</bibRefCitation>
<bibRefCitation id="11D9B8E44D521EB35FAF71FDE4CA1F78" author="Larson, DJ" journalOrPublisher="Monographs in Biodiversity, NRC Press, Ottawa, Ontario" pageId="6" pageNumber="7" title="Predaceous diving beetles (Coleoptera: Dytiscidae) of the nearctic region, with emphasis on the fauna of Canada and Alaska." year="2000">Larson et al. 2000</bibRefCitation>
).
</paragraph>
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72
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@ -65,7 +65,7 @@ Hainan, 3♂♂, 5♀♀, Qiongzhong City, Limu Shan 600 m, 15.V.2007, leg. Zong
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The morphology of
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was redescribed by
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; an updated distribution map was provided by

366
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DC., Prodr. 9: 175. 1845. Type:
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera foveolata" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="foveolata">Pachyptera foveolata</taxonomicName>
<taxonomicName id="259A67CC8DA5A9024CAB41FD740509FA" class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera foveolata" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="foveolata">Pachyptera foveolata</taxonomicName>
DC. (lectotype, designated by Sandwith [1932: 84]) [=
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera kerere" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="kerere">Pachyptera kerere</taxonomicName>
<taxonomicName id="42189C13BED6B10E822C6E2A61E16815" class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera kerere" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="kerere">Pachyptera kerere</taxonomicName>
(Aubl.) Sandwith]
</paragraph>
<paragraph pageId="8" pageNumber="97">
<taxonomicName authorityName="Rafinesque" authorityYear="1838" class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="genus">Sererea</taxonomicName>
<paragraph id="7A886849FC81D2E3608F8C6C3F20B138" pageId="8" pageNumber="97">
<taxonomicName id="DBE0AFE728829A05BA7080C506F0A8DD" authorityName="Rafinesque" authorityYear="1838" class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="genus">Sererea</taxonomicName>
Raf., Sylva Tellur. 107. 1838. Type:
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea heterophylla" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="heterophylla">Sererea heterophylla</taxonomicName>
<taxonomicName id="73DE806AC716A272BEC214F8D21E06FD" class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea heterophylla" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="heterophylla">Sererea heterophylla</taxonomicName>
(Willd.) Raf., Sylva Tellur. 107. 1838.
<emphasis italics="true" pageId="8" pageNumber="97">nom. illeg. superfl.</emphasis>
<emphasis id="92D3D50026FC685DFD2D0F4BB90595F0" italics="true" pageId="8" pageNumber="97">nom. illeg. superfl.</emphasis>
[=
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera kerere" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="kerere">Pachyptera kerere</taxonomicName>
<taxonomicName id="394A402DB389FCCA424531689F15F41E" class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera kerere" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="kerere">Pachyptera kerere</taxonomicName>
(Aubl.) Sandwith]
</paragraph>
<paragraph pageId="8" pageNumber="97">
<taxonomicName authorityName="Barbosa Rodrigues" authorityYear="1891" class="Magnoliopsida" family="Bignoniaceae" genus="Leucocalantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucocalantha" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="genus">Leucocalantha</taxonomicName>
<paragraph id="9A4576D41DC7CB77DE00090C3A497A23" pageId="8" pageNumber="97">
<taxonomicName id="5B02AB4077C85255E5CC6BA3DF6AD16D" authorityName="Barbosa Rodrigues" authorityYear="1891" class="Magnoliopsida" family="Bignoniaceae" genus="Leucocalantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucocalantha" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="genus">Leucocalantha</taxonomicName>
Barb. Rodr., Vellosia, ed. 2. 1: 46, tab. 7. 1891. Type:
<taxonomicName authorityName="Barb Rodr" authorityYear="1891" class="Magnoliopsida" family="Bignoniaceae" genus="Leucocalantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucocalantha aromatica" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="aromatica">Leucocalantha aromatica</taxonomicName>
<taxonomicName id="5DA7602D6D7F144634A92CE8A2F5060C" authorityName="Barb Rodr" authorityYear="1891" class="Magnoliopsida" family="Bignoniaceae" genus="Leucocalantha" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucocalantha aromatica" order="Lamiales" pageId="8" pageNumber="97" phylum="Tracheophyta" rank="species" species="aromatica">Leucocalantha aromatica</taxonomicName>
Barb. Rodr., Vellosia. ed. 2. 1: 47, tab. 7. 1891.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="97" type="description">
<paragraph pageId="8" pageNumber="97">Description.</paragraph>
<paragraph pageId="8" pageNumber="97">
<emphasis italics="true" pageId="8" pageNumber="97">Liana</emphasis>
<subSubSection id="50D5A9CF64402485C5C1E26DFE1DFAF2" pageId="8" pageNumber="97" type="description">
<paragraph id="8E3F16A54A81E497C567387E12274B57" pageId="8" pageNumber="97">Description.</paragraph>
<paragraph id="0B620E7010D7C5ED466855F1EB7FC950" pageId="8" pageNumber="97">
<emphasis id="5E3A3B3336D844E135B8FD6455A291AB" italics="true" pageId="8" pageNumber="97">Liana</emphasis>
; stems with four phloem wedges in cross-section, solid (hollow in some specimens of
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="7A03A3802BE5E6B80C244BDE4828C191" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="F554024459D8C93B52944EC832E38F75" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
), cylindrical to tetragonal, striated, with lenticels (without in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="F48EF9D71C61B1BDF52EAD70D4588BD1" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="0600AB2C808341004DCDE9C341FB0082" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
), with interpetiolar extrafloral nectaries, with a continuous (discontinuous) and transversal interpetiolar ridge, with a papery peeling bark, lepidote, puberulous becoming glabrescent with age; prophylls of the axillary buds 3(-5) seriated (a single series visible to the naked eye in some specimens of
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="3E5F9A0E6FD3206557714E1FB10C3B3D" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="985C19002F64219F77031B78B80A3FB6" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
), flattened and ensiform (triangular and minute in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="FD6D9E0C5BF5C0894FFFB32A47DE196E" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="58A25AF9B878E513770E1E74DF3933DB" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
).
<emphasis italics="true" pageId="8" pageNumber="97">Leaves</emphasis>
<emphasis id="6F9C407DBDAE156CBF98EB242F50F7A5" italics="true" pageId="8" pageNumber="97">Leaves</emphasis>
3-2-foliolate with the terminal leaflet replaced by a trifid tendril; blades discolorous (concolor), chartaceous to coriaceous, usually asymmetric (symetric in some specimens of
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="84515F9E47DCA882AD513557D091516D" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="07BB4F9DFAF594CDE54B98857474E1CB" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
), apex mucronulate, glabrous to puberulous, with simple trichomes covering veins (throughout surface), lepidote, with patelliform trichomes throughout the lamina, venation pinnate, secondary venation brochidromous, tertiary venation percurrent, margin entire, flat or sub-revolute; petioles striated, apices articulated, glabrous to puberulous, lepidote, patelliform glands distributed at petiole apices; petiolules with unequal lengths, striated, apices not-pulvinated (pulvinated in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="9CE711E5150D4229CEB32EDA6DD35909" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="A99CA98A0C13CE314A57A42D3C5EB8E7" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
), puberulous, lepidote, lateral petiolules shorter than the apical ones.
<emphasis italics="true" pageId="8" pageNumber="97">Inflorescence</emphasis>
<emphasis id="F3BD5C9A7794CBE7AD98BF54F4DDA744" italics="true" pageId="8" pageNumber="97">Inflorescence</emphasis>
axillary or terminal, a few-flowered raceme, congested (lax in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="600C8000B2E8DA3E9F5240F60AD8B91C" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="81707913B697DF7EF4C7788C76DBF0D8" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
); axis puberulous, lepidote, patelliform glands grouped at the axis; pedicel puberulous, lepidote; bracts and bracteoles caducous, scarcely evident, puberulous, lepidote.
<emphasis italics="true" pageId="8" pageNumber="97">Calyx</emphasis>
<emphasis id="FE271D4CDD775D353CCA127F417686E0" italics="true" pageId="8" pageNumber="97">Calyx</emphasis>
tubular (cupular in
<taxonomicName lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
<taxonomicName id="2167CFDC7362C34593D4705850BEE3CA" lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis id="1BC165952F97AEC1EC18852C5E786AE5" italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
</taxonomicName>
), coriaceous, smooth, glabrous internally, puberulous externally, lepidote, patelliform glands grouped at the upper portion.
<emphasis italics="true" pageId="8" pageNumber="97">Corolla</emphasis>
<emphasis id="4D0574F8B0DF42624AC295D416D16292" italics="true" pageId="8" pageNumber="97">Corolla</emphasis>
white to cream (orange to red in
<taxonomicName lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
<taxonomicName id="B6526844D8161AD8D18B171B4AD307AE" lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis id="F5ED5DCA474DF1256D254D05E273A7D7" italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
</taxonomicName>
and light pink to pale purple in
<taxonomicName lsidName="P. incarnata" pageId="8" pageNumber="97" rank="species" species="incarnata">
<emphasis italics="true" pageId="8" pageNumber="97">P. incarnata</emphasis>
<taxonomicName id="9ECB591CCEAD8072E634EB8790A20F05" lsidName="P. incarnata" pageId="8" pageNumber="97" rank="species" species="incarnata">
<emphasis id="C2D87D0C6E7B4BCB9304BE6DC36051BA" italics="true" pageId="8" pageNumber="97">P. incarnata</emphasis>
</taxonomicName>
), with yellow nectar guides, infundibuliform, (hypocrateriform in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="D77EE4DC93954CA5B7559A830CDE39BF" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="16B3B211D4D54C510980DB482E8528C6" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
and tubular-campanulate in
<taxonomicName lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
<taxonomicName id="5BE50A30B20C17CA9026B4BDB81C0190" lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis id="4726444B116B38320DB0C43C4EABE854" italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
</taxonomicName>
), straight, dorso-ventrally compressed (not compressed in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="9F782CD6D8859F24750FED7BDE6236F0" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="9F7BF40567A0C6B65DB5B119FCBE02FF" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
), membranaceous, tube puberulous externally, lepidote, glabrous internally, but villous at the region of insertion of stamens and staminode (glabrous in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="84F63CAF7859577523C2EA34C687860E" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="9EA32D384A00805614D637555DBE4EB1" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
); lobes imbricate, with a pair of patelliform glands arranged in lines externally, lepidote internally.
<emphasis italics="true" pageId="8" pageNumber="97">Androecium</emphasis>
<emphasis id="5E39AF6AAE111B2C40A7558761AE7DA5" italics="true" pageId="8" pageNumber="97">Androecium</emphasis>
didynamous, included in two heights, with one staminode, glabrous (puberulous in
<taxonomicName lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
<taxonomicName id="9ABAFA0067D43C0866DB21C9D26FD2B9" lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis id="211C74DECC31B9DE043E82CFDC42B054" italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
</taxonomicName>
); anthers white, becoming darkish with age, included (sub-exserted in
<taxonomicName lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
<taxonomicName id="A32D35ECFD8FA935A4218A7B6F131A3D" lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis id="180F19CCB2DCCDB7185CF396D18B0E6A" italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
</taxonomicName>
), villous (glabrous in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="B0482E1D5C1F5149CC57159A6B5FCA32" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="FCE50B7576C143BFA4F7604556BF1AAF" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
), basifixed, connective thick, round (acute in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="592762A5E5209AF8139B37DD8CE7BB43" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="AEF744997F95C03E5E1EA2BB5672A7F3" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
), with thecae divergent, curved forward (straight in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="D0B5850F0A8AF82F1962FD75796E3EE3" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="F677F127AEED7F641AE37F84B6E819CD" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
); pollen 3 colpate, microreticulate (4 colpate, psilate-foveolate-microreticulate in
<taxonomicName lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
<taxonomicName id="E4695FACF55D0467670268D44C3FEEFE" lsidName="P. aromatica" pageId="8" pageNumber="97" rank="species" species="aromatica">
<emphasis id="F6B1605AD14BA5F6E9BB5F958A4AD5EA" italics="true" pageId="8" pageNumber="97">P. aromatica</emphasis>
</taxonomicName>
).
<emphasis italics="true" pageId="8" pageNumber="97">Gynoecium</emphasis>
<emphasis id="E9500BD300352EF5FDE56F073FD4224F" italics="true" pageId="8" pageNumber="97">Gynoecium</emphasis>
glabrous; ovary cylindrical, not-sulcate (bisulcate in
<taxonomicName lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
<taxonomicName id="2A1AD408B8A479636BD41B246E5C2E93" lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis id="5545BF57C79F624EFEBA9DD0E403C24C" italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
</taxonomicName>
), smooth, pubescent, (densely lepidote in
<taxonomicName lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
<taxonomicName id="E3E24F9C07C9496D5DE6857788868A50" lsidName="P. erythraea" pageId="8" pageNumber="97" rank="species" species="erythraea">
<emphasis id="0C1A5918C983442C2741062FA3254BDB" italics="true" pageId="8" pageNumber="97">P. erythraea</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. incarnata" pageId="8" pageNumber="97" rank="species" species="incarnata">
<emphasis italics="true" pageId="8" pageNumber="97">P. incarnata</emphasis>
<taxonomicName id="611A55B3A7406DFBA64221A5A60B6814" lsidName="P. incarnata" pageId="8" pageNumber="97" rank="species" species="incarnata">
<emphasis id="44487BC17D875EF27CC75E0D364DB7A9" italics="true" pageId="8" pageNumber="97">P. incarnata</emphasis>
</taxonomicName>
); ovules arranged in two series per locule, placentation axial; stigma glabrous; nectar disc well developed, glabrous.
<emphasis italics="true" pageId="8" pageNumber="97">Capsule</emphasis>
<emphasis id="3FDBA404332704B4DD54A604DDC7EA69" italics="true" pageId="8" pageNumber="97">Capsule</emphasis>
linear, flattened (fusiform, inflated in
<taxonomicName lsidName="P. kerere" pageId="8" pageNumber="97" rank="species" species="kerere">
<emphasis italics="true" pageId="8" pageNumber="97">P. kerere</emphasis>
<taxonomicName id="E366FD800E8139CEBFCD71E5DF7AB470" lsidName="P. kerere" pageId="8" pageNumber="97" rank="species" species="kerere">
<emphasis id="ACA60700FD21E974F06C274B4D37E233" italics="true" pageId="8" pageNumber="97">P. kerere</emphasis>
</taxonomicName>
), coriaceous to woody, smooth, puberulous, lepidote, with patelliform glandular trichomes throughout, in higher densities at the margins of valves, without lenticels, each valve with an inconspicuous longitudinal midline (conspicuous and raised in
<taxonomicName lsidName="P. kerere" pageId="8" pageNumber="97" rank="species" species="kerere">
<emphasis italics="true" pageId="8" pageNumber="97">P. kerere</emphasis>
<taxonomicName id="4EDA8369F2593D88AFB6C615241A7D13" lsidName="P. kerere" pageId="8" pageNumber="97" rank="species" species="kerere">
<emphasis id="531C0DADBD414B5A1C865CA22983C172" italics="true" pageId="8" pageNumber="97">P. kerere</emphasis>
</taxonomicName>
), calyx caducous; seeds oblong, thin, not-corky (irregulary circular, obcordate, thick and corky in
<taxonomicName lsidName="P. kerere" pageId="8" pageNumber="97" rank="species" species="kerere">
<emphasis italics="true" pageId="8" pageNumber="97">P. kerere</emphasis>
<taxonomicName id="2C2D4C87AE920DAD7E7705B6F945A734" lsidName="P. kerere" pageId="8" pageNumber="97" rank="species" species="kerere">
<emphasis id="DB92A567C75DCDB3F03D02840F923961" italics="true" pageId="8" pageNumber="97">P. kerere</emphasis>
</taxonomicName>
), chartaceous to coriaceous, glabrous, smooth, striated, winged (wingless in
<taxonomicName lsidName="P. kerere" pageId="8" pageNumber="97" rank="species" species="kerere">
<emphasis italics="true" pageId="8" pageNumber="97">P. kerere</emphasis>
<taxonomicName id="C405FB59C5E979255871FDA2468730E5" lsidName="P. kerere" pageId="8" pageNumber="97" rank="species" species="kerere">
<emphasis id="EA07B9C9DF0BB6ECC5A23CAF5AE4537A" italics="true" pageId="8" pageNumber="97">P. kerere</emphasis>
</taxonomicName>
), with membranaceous (chartaceous) and hyaline wings.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="98" type="nomenclatural note">
<paragraph pageId="9" pageNumber="98">
<pageBreakToken pageId="9" pageNumber="98" start="start">Nomenclatural</pageBreakToken>
<subSubSection id="5A5374ADEF5CFF213D15F069859EAC63" pageId="9" pageNumber="98" type="nomenclatural note">
<paragraph id="6B4BF44966DBB9DC47842A4B3763FD98" pageId="9" pageNumber="98">
<pageBreakToken id="E585398E630F0D08A31E9255C7B85BAF" pageId="9" pageNumber="98" start="start">Nomenclatural</pageBreakToken>
note.
</paragraph>
<paragraph pageId="9" pageNumber="98">
<taxonomicName authorityName="Rafinesque" authorityYear="1838" class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="98">Sererea</emphasis>
<paragraph id="D0C6F3B967CC938563A66875ED49E455" pageId="9" pageNumber="98">
<taxonomicName id="91BAD1CBE076B48AC82C588DB90E3328" authorityName="Rafinesque" authorityYear="1838" class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis id="290479B3F80E0160BDBAEED1F69B866D" italics="true" pageId="9" pageNumber="98">Sererea</emphasis>
</taxonomicName>
Raf. was described to accommodate one species,
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea heterophyla" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="species" species="heterophyla">
<emphasis italics="true" pageId="9" pageNumber="98">Sererea heterophyla</emphasis>
<taxonomicName id="37528C6D7EF000BBBC8C8ACE6DE6D800" class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea heterophyla" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="species" species="heterophyla">
<emphasis id="77083294D84DFD954A4761D7369E267C" italics="true" pageId="9" pageNumber="98">Sererea heterophyla</emphasis>
</taxonomicName>
Raf., using a wrong spelling. However,
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea heterophylla" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis italics="true" pageId="9" pageNumber="98">Sererea heterophylla</emphasis>
<taxonomicName id="11C7BAF4969D7600363C69A396F17E71" class="Magnoliopsida" family="Bignoniaceae" genus="Sererea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sererea heterophylla" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis id="957A61E34DD9A05959B6EA9551C5553C" italics="true" pageId="9" pageNumber="98">Sererea heterophylla</emphasis>
</taxonomicName>
(Willd.) Raf. was actually based on
<taxonomicName authorityName="Willd" authorityYear="1800" class="Magnoliopsida" family="Bignoniaceae" genus="Bignonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bignonia heterophylla" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis italics="true" pageId="9" pageNumber="98">Bignonia heterophylla</emphasis>
<taxonomicName id="5983DF6D0B68111EACB41579BC46FF99" authorityName="Willd" authorityYear="1800" class="Magnoliopsida" family="Bignoniaceae" genus="Bignonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bignonia heterophylla" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis id="0A8776C8CA642BD6CFC4AE5C1A98B016" italics="true" pageId="9" pageNumber="98">Bignonia heterophylla</emphasis>
</taxonomicName>
Willd, a superfluous name for
<taxonomicName authorityName="Aubl" authorityYear="1775" class="Magnoliopsida" family="Bignoniaceae" genus="Bignonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bignonia kerere" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="species" species="kerere">
<emphasis italics="true" pageId="9" pageNumber="98">Bignonia kerere</emphasis>
<taxonomicName id="8231C4D7BD9272A3AA896C19F247525B" authorityName="Aubl" authorityYear="1775" class="Magnoliopsida" family="Bignoniaceae" genus="Bignonia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Bignonia kerere" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="species" species="kerere">
<emphasis id="7B0B0753CA169C9D2B4D00779850603A" italics="true" pageId="9" pageNumber="98">Bignonia kerere</emphasis>
</taxonomicName>
Aubl.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="98" type="distribution">
<paragraph pageId="9" pageNumber="98">Number of species, distribution and habitat.</paragraph>
<paragraph pageId="9" pageNumber="98">
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="98">Pachyptera</emphasis>
<subSubSection id="089D922C314FAB846290E42CB7CB6E94" pageId="9" pageNumber="98" type="distribution">
<paragraph id="ADF3FE6DC166FB70028BB4E0918E2FD6" pageId="9" pageNumber="98">Number of species, distribution and habitat.</paragraph>
<paragraph id="79BA974A2CF41F2E3793548DAF183167" pageId="9" pageNumber="98">
<taxonomicName id="52A5C4BD7D5D133A8910D48C64E78213" class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis id="C894B11EF98CBC140CA518E12BF69219" italics="true" pageId="9" pageNumber="98">Pachyptera</emphasis>
</taxonomicName>
comprises five species found in wet and flooded forest vegetation from Belize to
<normalizedToken originalValue="Bolívia">Bolivia</normalizedToken>
<normalizedToken id="73F5C650C45D2ECA8A20ACBF1C6D6C81" originalValue="Bolívia">Bolivia</normalizedToken>
and Brazil.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="98" type="key">
<paragraph pageId="9" pageNumber="98">
<subSubSection id="3EC548F13F5A50C042E805AE39D76B8A" pageId="9" pageNumber="98" type="key">
<paragraph id="E72E12B17EF5D54FF392F649B588D782" pageId="9" pageNumber="98">
Key to species of
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="98">Pachyptera</emphasis>
<taxonomicName id="D66A7210B6AA7E58850EDCC7718637CA" class="Magnoliopsida" family="Bignoniaceae" genus="Pachyptera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pachyptera" order="Lamiales" pageId="9" pageNumber="98" phylum="Tracheophyta" rank="genus">
<emphasis id="1CE8890652D48BDD4B0A72D1D7AA0309" italics="true" pageId="9" pageNumber="98">Pachyptera</emphasis>
</taxonomicName>
</paragraph>
<paragraph pageId="9" pageNumber="98">
<table pageId="9" pageNumber="98">
<tr pageId="9" pageNumber="98">
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">1</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">Stems cylindrical (if tetragonal, then only on older portions), without lenticels; prophylls of the axillary buds triangular and minute; corolla hypocrateriform</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName lsidName="P. aromatica" pageId="9" pageNumber="98" rank="species" species="aromatica">
<emphasis bold="true" italics="true" pageId="9" pageNumber="98">P. aromatica</emphasis>
<paragraph id="453F54C40A0745622640206506E25E49" pageId="9" pageNumber="98">
<table id="AFD0EAA7B691D42605EFC81462A58217" pageId="9" pageNumber="98">
<tr id="C36BA63BADD501272AE0A3876F878E16" pageId="9" pageNumber="98">
<td id="0CE4C52EB0BD988C973CCAA0A814695F" colspan="1" pageId="9" pageNumber="98" rowspan="1">1</td>
<td id="3566A28D39AB0BB2D2B383F8730D43F6" colspan="1" pageId="9" pageNumber="98" rowspan="1">Stems cylindrical (if tetragonal, then only on older portions), without lenticels; prophylls of the axillary buds triangular and minute; corolla hypocrateriform</td>
<td id="B34754475309C4FAEB2A8FD1989B4725" colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName id="8D8C960E35F6AFAE222FA935D876EE27" lsidName="P. aromatica" pageId="9" pageNumber="98" rank="species" species="aromatica">
<emphasis id="1EE2F274D17D25935E0E51E686DA47FB" bold="true" italics="true" pageId="9" pageNumber="98">P. aromatica</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="9" pageNumber="98">
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">-</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">Stems tetragonal (if cylindrical, then only on younger portions), with lenticels; prophylls of the axillary buds ensiform and flattened; corolla infundibuliform or tubular-campanulate</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">
<emphasis bold="true" pageId="9" pageNumber="98">2</emphasis>
<tr id="B6E94E01C5ED42181DFEAEC26A57189F" pageId="9" pageNumber="98">
<td id="F172BA98AF145CA24722CDAFBF514ED8" colspan="1" pageId="9" pageNumber="98" rowspan="1">-</td>
<td id="086893BFB6710BFF842DBFAF1E93DF8E" colspan="1" pageId="9" pageNumber="98" rowspan="1">Stems tetragonal (if cylindrical, then only on younger portions), with lenticels; prophylls of the axillary buds ensiform and flattened; corolla infundibuliform or tubular-campanulate</td>
<td id="DF2DE3DD2AE93BC62562C9EE2BBEB802" colspan="1" pageId="9" pageNumber="98" rowspan="1">
<emphasis id="ED33C8A6E07ED5CAB3CE730BB377228E" bold="true" pageId="9" pageNumber="98">2</emphasis>
</td>
</tr>
<tr pageId="9" pageNumber="98">
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">2</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">Corolla light pink to pale purple or orange to red; ovary densely lepidote</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">
<emphasis bold="true" pageId="9" pageNumber="98">3</emphasis>
<tr id="C7C417630E7B0C65672AF6CAEF2A6BD8" pageId="9" pageNumber="98">
<td id="CD443058DEFC0867421DFEFAEDBC6244" colspan="1" pageId="9" pageNumber="98" rowspan="1">2</td>
<td id="7B69A2FE72B6F5C061149A57115636AE" colspan="1" pageId="9" pageNumber="98" rowspan="1">Corolla light pink to pale purple or orange to red; ovary densely lepidote</td>
<td id="9857391A8DC9D870C74D28E19C3E2B14" colspan="1" pageId="9" pageNumber="98" rowspan="1">
<emphasis id="CAA061A5056E65597EE720218525813F" bold="true" pageId="9" pageNumber="98">3</emphasis>
</td>
</tr>
<tr pageId="9" pageNumber="98">
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">-</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">Corolla white to cream; ovary pubescent</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">
<emphasis bold="true" pageId="9" pageNumber="98">4</emphasis>
<tr id="70F640578738D76375F6F87596E97FC8" pageId="9" pageNumber="98">
<td id="FC59911189E4405DFE8B04B2D4485FDD" colspan="1" pageId="9" pageNumber="98" rowspan="1">-</td>
<td id="D3C6B06381150DCEADB70680C29DEE3E" colspan="1" pageId="9" pageNumber="98" rowspan="1">Corolla white to cream; ovary pubescent</td>
<td id="B99F8E5A73FD13740F5D66BAB13D8A84" colspan="1" pageId="9" pageNumber="98" rowspan="1">
<emphasis id="85297006BDB4B494EC1D62608FD68EBC" bold="true" pageId="9" pageNumber="98">4</emphasis>
</td>
</tr>
<tr pageId="9" pageNumber="98">
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">3</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">Calyx reddish-wine throughout, cupular; corolla orange to red, tubular campanulate; stamens sub-exserted; capsule linear, 34.0-41.0 cm long, ≥ 2.7 cm wide</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName lsidName="P. erythraea" pageId="9" pageNumber="98" rank="species" species="erythraea">
<emphasis bold="true" italics="true" pageId="9" pageNumber="98">P. erythraea</emphasis>
<tr id="D46583713C3AD2F797245A45D7C961ED" pageId="9" pageNumber="98">
<td id="DD19E53B0E99412DBACEED3D5046173A" colspan="1" pageId="9" pageNumber="98" rowspan="1">3</td>
<td id="E67D9D89B599AA9D1DAE352AB37DD54C" colspan="1" pageId="9" pageNumber="98" rowspan="1">Calyx reddish-wine throughout, cupular; corolla orange to red, tubular campanulate; stamens sub-exserted; capsule linear, 34.0-41.0 cm long, ≥ 2.7 cm wide</td>
<td id="BF6BAC55A31846012E61362AD47D6A34" colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName id="EB9EA313E399369C92F9583AADCE8ADE" lsidName="P. erythraea" pageId="9" pageNumber="98" rank="species" species="erythraea">
<emphasis id="B783C6DC78DF78C9DFF4A3FECC0F7C56" bold="true" italics="true" pageId="9" pageNumber="98">P. erythraea</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="9" pageNumber="98">
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">-</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">Calyx green, light pink at the apex, tubular; corolla light pink to pale purple, infundibuliform; stamens included; capsule linear, 10.5-42.6 cm long, ≤ 2.6 cm wide</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName lsidName="P. incarnata" pageId="9" pageNumber="98" rank="species" species="incarnata">
<emphasis bold="true" italics="true" pageId="9" pageNumber="98">P. incarnata</emphasis>
<tr id="52A4A8CF90ED251F4368B6D919561DD4" pageId="9" pageNumber="98">
<td id="96CB04B77933FC93D87E3F90DDD222ED" colspan="1" pageId="9" pageNumber="98" rowspan="1">-</td>
<td id="140B2AA519983B8072C8466B254F564B" colspan="1" pageId="9" pageNumber="98" rowspan="1">Calyx green, light pink at the apex, tubular; corolla light pink to pale purple, infundibuliform; stamens included; capsule linear, 10.5-42.6 cm long, ≤ 2.6 cm wide</td>
<td id="70D34300EA0B14D1C323626F848DC558" colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName id="12BDAE8C1083E0DEDC626716753E192F" lsidName="P. incarnata" pageId="9" pageNumber="98" rank="species" species="incarnata">
<emphasis id="61FA0A97B09C03CCA0DA71E6DF987A54" bold="true" italics="true" pageId="9" pageNumber="98">P. incarnata</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="9" pageNumber="98">
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">4</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">Ovary densely pubescent; capsule fusiform, inflated, each valve with a conspicuous midline; seeds thick, corky and wingless</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName lsidName="P. kerere" pageId="9" pageNumber="98" rank="species" species="kerere">
<emphasis bold="true" italics="true" pageId="9" pageNumber="98">P. kerere</emphasis>
<tr id="19C31977A4DD63F7F9F6BA4AFA61A121" pageId="9" pageNumber="98">
<td id="908DBFD8DC790C5686CF6F00059FE0AB" colspan="1" pageId="9" pageNumber="98" rowspan="1">4</td>
<td id="FCB4F867B1B1CAA676FE8E1C931E26FD" colspan="1" pageId="9" pageNumber="98" rowspan="1">Ovary densely pubescent; capsule fusiform, inflated, each valve with a conspicuous midline; seeds thick, corky and wingless</td>
<td id="651BDEA52480319FB03D95FAD4679D12" colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName id="534D43E1E7B78B455DDF9FA225B25401" lsidName="P. kerere" pageId="9" pageNumber="98" rank="species" species="kerere">
<emphasis id="E49953C7C232EBFC7FE7171FEFB4261F" bold="true" italics="true" pageId="9" pageNumber="98">P. kerere</emphasis>
</taxonomicName>
</td>
</tr>
<tr pageId="9" pageNumber="98">
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">-</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">Ovary sparsely to moderately pubescent; capsule linear, flat, each valve with an inconspicuous midline; seeds thin, coriaceous and winged</td>
<td colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName lsidName="P. linearis" pageId="9" pageNumber="98" rank="species" species="linearis">
<emphasis bold="true" italics="true" pageId="9" pageNumber="98">P. linearis</emphasis>
<tr id="139761943838A0494BE1CB21222B014A" pageId="9" pageNumber="98">
<td id="23425615A34EC23251EADBF0D54B70B4" colspan="1" pageId="9" pageNumber="98" rowspan="1">-</td>
<td id="278E75D9E7279085C467B8FC67F51C66" colspan="1" pageId="9" pageNumber="98" rowspan="1">Ovary sparsely to moderately pubescent; capsule linear, flat, each valve with an inconspicuous midline; seeds thin, coriaceous and winged</td>
<td id="B482B3301F3287D32F3DF88BDE7043A7" colspan="1" pageId="9" pageNumber="98" rowspan="1">
<taxonomicName id="936D8F090B2734DDECEE5B616EF11C03" lsidName="P. linearis" pageId="9" pageNumber="98" rank="species" species="linearis">
<emphasis id="58D0549B0C5E86B22BA4FD55B550A3ED" bold="true" italics="true" pageId="9" pageNumber="98">P. linearis</emphasis>
</taxonomicName>
</td>
</tr>

142
data/20/E1/B7/20E1B7B63D6E5803A70E990EA4551A91.xml
View file

@ -1,113 +1,113 @@
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Cameron, 1932
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<taxonomicName authorityName="Cameron" authorityYear="1932" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius lineipennis" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="lineipennis">Quedius lineipennis</taxonomicName>
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Cameron:
<bibRefCitation author="Smetana, A" journalOrPublisher="Quaestiones Entomologicae" pageId="0" pageNumber="109" pagination="163 - 464" refId="B19" refString="Smetana, A, 1988. Revision of the tribes Quediini and Atanygnathinini. Part II. The Himalayan region (Coleoptera: Staphylinidae). Quaestiones Entomologicae 24: 163 - 464" title="Revision of the tribes Quediini and Atanygnathinini. Part II. The Himalayan region (Coleoptera: Staphylinidae)." volume="24" year="1988">Smetana 1988</bibRefCitation>
<bibRefCitation id="6B5A52199ED3B4DCC961615829028BFE" author="Smetana, A" journalOrPublisher="Quaestiones Entomologicae" pageId="0" pageNumber="109" pagination="163 - 464" refId="B19" refString="Smetana, A, 1988. Revision of the tribes Quediini and Atanygnathinini. Part II. The Himalayan region (Coleoptera: Staphylinidae). Quaestiones Entomologicae 24: 163 - 464" title="Revision of the tribes Quediini and Atanygnathinini. Part II. The Himalayan region (Coleoptera: Staphylinidae)." volume="24" year="1988">Smetana 1988</bibRefCitation>
</paragraph>
<paragraph pageId="0" pageNumber="109">
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<taxonomicName id="0BAC0AA9CBE7ED897CA597C3CE858B6E" baseAuthorityName="Cameron" baseAuthorityYear="1932" class="Insecta" family="Staphylinidae" genus="Alesiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alesiella lineipennis" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="species" species="lineipennis">Alesiella lineipennis</taxonomicName>
(Cameron):
<bibRefCitation DOI="https://doi.org/10.1111/syen.12021" author="Brunke, A" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="109" pagination="689 - 707" refId="B3" refString="Brunke, A, Solodovnikov, A, 2013. Alesiella gen.n. and a newly discovered relict lineage of Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology 38: 689 - 707, DOI: https://doi.org/10.1111/syen.12021" title="Alesiella gen. n. and a newly discovered relict lineage of Staphylinini (Coleoptera: Staphylinidae)." url="https://doi.org/10.1111/syen.12021" volume="38" year="2013">Brunke and Solodovnikov 2013</bibRefCitation>
<bibRefCitation id="A5DE4160D902725400E7CD6E6B8FA600" DOI="https://doi.org/10.1111/syen.12021" author="Brunke, A" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="109" pagination="689 - 707" refId="B3" refString="Brunke, A, Solodovnikov, A, 2013. Alesiella gen.n. and a newly discovered relict lineage of Staphylinini (Coleoptera: Staphylinidae). Systematic Entomology 38: 689 - 707, DOI: https://doi.org/10.1111/syen.12021" title="Alesiella gen. n. and a newly discovered relict lineage of Staphylinini (Coleoptera: Staphylinidae)." url="https://doi.org/10.1111/syen.12021" volume="38" year="2013">Brunke and Solodovnikov 2013</bibRefCitation>
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<subSubSection pageId="0" pageNumber="109" type="type locality">
<paragraph pageId="0" pageNumber="109">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="109">Mogok [= Ruby Mines], Mandalay, Myanmar</paragraph>
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<paragraph id="5601BC9AE6ED0925BF73FF844FEF032A" pageId="0" pageNumber="109">Type locality.</paragraph>
<paragraph id="9031372420941C3FF50342130054A81F" pageId="0" pageNumber="109">Mogok [= Ruby Mines], Mandalay, Myanmar</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="109" type="type material">
<paragraph pageId="0" pageNumber="109">Non-type material.</paragraph>
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<emphasis bold="true" pageId="0" pageNumber="109">Thailand: Chiang Rai</emphasis>
<subSubSection id="4B675EE8E990DA0A32D6C198204DCB77" pageId="0" pageNumber="109" type="type material">
<paragraph id="EAB87C73F0F898FBC9E42EA008E99D56" pageId="0" pageNumber="109">Non-type material.</paragraph>
<paragraph id="829D45D31E6D54A2605D67EDAD154102" pageId="0" pageNumber="109">
<emphasis id="5953AD1A2D132A8CEEE968510D901C2D" bold="true" pageId="0" pageNumber="109">Thailand: Chiang Rai</emphasis>
: Wiang Pa Pao District [no specific locality], 17-21.V.2015, K. Takahashi (1 male, aedeagus missing, cHay).
</paragraph>
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<paragraph pageId="0" pageNumber="109">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="109">
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<paragraph id="AD9C0C5CCDBE5FA5044B3ED9AC629F7F" pageId="0" pageNumber="109">Diagnosis.</paragraph>
<paragraph id="B5D8ADCA14824DD824FFB185023CBAF1" pageId="0" pageNumber="109">
Only one species of
<taxonomicName authorityName="Brunke &amp; Solodovnikov" authorityYear="2013" class="Insecta" family="Staphylinidae" genus="Alesiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alesiella" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Alesiella</emphasis>
<taxonomicName id="304510E5D77D67A418A37014B6C0E5F3" authorityName="Brunke &amp; Solodovnikov" authorityYear="2013" class="Insecta" family="Staphylinidae" genus="Alesiella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alesiella" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="7E9C3B8769876CBFA7F00A103F51558B" italics="true" pageId="0" pageNumber="109">Alesiella</emphasis>
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is known and can be recognized by characters in the above key to genera. The specimen from Thailand does not differ from the type material (previously studied by the author), though the aedeagus was lost during mounting (Y. Hayashi, pers. comm.).
</paragraph>
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<paragraph pageId="0" pageNumber="109">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="109">Myanmar and Thailand (new country record).</paragraph>
<subSubSection id="EE673B0BAEBAA139CDB2B74A316B0835" pageId="0" pageNumber="109" type="distribution">
<paragraph id="C72B0E327355CB2614D20B5523A0A79F" pageId="0" pageNumber="109">Distribution.</paragraph>
<paragraph id="E356E5B62BFD2F55080BF1947452E68B" pageId="0" pageNumber="109">Myanmar and Thailand (new country record).</paragraph>
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<paragraph pageId="0" pageNumber="109">Bionomics.</paragraph>
<paragraph pageId="0" pageNumber="109">
<subSubSection id="51E01A91878D0E39FE90739C2D176E01" pageId="0" pageNumber="109" type="bionomics">
<paragraph id="63CA0E7F75F1D7D09CC24B1C68CE3089" pageId="0" pageNumber="109">Bionomics.</paragraph>
<paragraph id="05A1A6D02DDE491A21FD5CF7659F7960" pageId="0" pageNumber="109">
Nothing is known about this
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microhabitat preferences but it probably occurs under the bark of dead trees in the earlier fermentation states of decay, as does its sister group
<taxonomicName authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Quediomacrus" kingdom="Animalia" lsidName="Quediomacrus" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="109">Quediomacrus</emphasis>
<taxonomicName id="92A30F1FC7AB77D002F2B3971DB1286F" authorityName="Sharp" authorityYear="1884" class="Insecta" family="Staphylinidae" genus="Quediomacrus" kingdom="Animalia" lsidName="Quediomacrus" order="Coleoptera" pageId="0" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="C0208AECC612A58F6A9601BA83C07E61" italics="true" pageId="0" pageNumber="109">Quediomacrus</emphasis>
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.
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<subSubSection pageId="0" pageNumber="109" type="comments">
<paragraph pageId="0" pageNumber="109">Comments.</paragraph>
<paragraph pageId="0" pageNumber="109">
<subSubSection id="88A39061FBCD18EB6A354BA009F56DE4" pageId="0" pageNumber="109" type="comments">
<paragraph id="6459398396DF2A904385047F0413E85D" pageId="0" pageNumber="109">Comments.</paragraph>
<paragraph id="5A25C5A5A151AF70E0CCC3E516F43A6F" pageId="0" pageNumber="109">
The above specimen is a new record of the genus and species from Thailand, and represents the first available material in more than 130 years (since 1890). The above record also indicates that the species is certainly still extant and rather widespread, though its elevational range remains unknown. Although the type series only bears the information 'Ruby Mines,
<normalizedToken originalValue="Doherty’">Doherty'</normalizedToken>
<normalizedToken id="7514E2FFBD7F35345644C5A65426DD80" originalValue="Doherty’">Doherty'</normalizedToken>
, the diaries of William Doherty (reproduced in
<bibRefCitation author="Hartert, E" journalOrPublisher="Novitates Zoologicae" pageId="0" pageNumber="109" pagination="494 - 506" refId="B11" refString="Hartert, E, 1901. William Doherty, Obituary. Novitates Zoologicae 8: 494 - 506" title="William Doherty, Obituary." volume="8" year="1901">Hartert 1901</bibRefCitation>
<bibRefCitation id="457B213546BB72C7E437061017AB0858" author="Hartert, E" journalOrPublisher="Novitates Zoologicae" pageId="0" pageNumber="109" pagination="494 - 506" refId="B11" refString="Hartert, E, 1901. William Doherty, Obituary. Novitates Zoologicae 8: 494 - 506" title="William Doherty, Obituary." volume="8" year="1901">Hartert 1901</bibRefCitation>
) indicate that they were collected somewhere along the route (1150-1800 m) between the towns of Mogok (= 'Ruby
<normalizedToken originalValue="Mines’">Mines'</normalizedToken>
<normalizedToken id="4B290D3D2F1D63DD35E8B0F3365FBDFC" originalValue="Mines’">Mines'</normalizedToken>
) and Bernardmyo, during 1890. This montane rainforest elevational range is compatible with the general locality of the specimen from Thailand.
</paragraph>
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116
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Yakovlev, 2006.
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<paragraph pageId="2" pageNumber="37">Small dark colored moths with dark hair densely covering the body. Antennae bipectinate. Forewing with a scarcely seen streaky pattern; hindwing dark without pattern; fringe evenly dark on both wings. Sexual dimorphism weakly expressed but female somewhat larger than male with wider wings and non-pectinate antennae.</paragraph>
<paragraph id="0FCD1E1DCC8F8ED69A8C7477FDE006AB" pageId="2" pageNumber="37">Small dark colored moths with dark hair densely covering the body. Antennae bipectinate. Forewing with a scarcely seen streaky pattern; hindwing dark without pattern; fringe evenly dark on both wings. Sexual dimorphism weakly expressed but female somewhat larger than male with wider wings and non-pectinate antennae.</paragraph>
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<paragraph pageId="2" pageNumber="37">Papillae anales elongated with rounded apices; apophyses posteriores thin, twice as long as anteriores; ostium opening immersed, fissure-like, surrounded by cordate rim; ductus bursae membranous, long and narrow; bursa elongate, gradually inflating to apex; ductus seminalis thin, enters bursa near its junction with ductus bursae.</paragraph>
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<paragraph id="CCCCD01EA98605910814A3EBA8CBD74C" pageId="2" pageNumber="37">Distribution.</paragraph>
<paragraph id="30ECC0DC58D5CDB91DA99DC2C5CF7A1C" pageId="2" pageNumber="37">Three species distributed in NE India (Assam), Nepal, SE China (Sichuan).</paragraph>
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Subgenus
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</paragraph>
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<subSubSection pageId="0" pageNumber="95" type="reference_group">
<paragraph pageId="0" pageNumber="95">
<taxonomicName class="Insecta" family="Carabidae" genus="Feroperis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feroperis" order="Coleoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="genus">Feroperis</taxonomicName>
<subSubSection id="4AB469E1023176EA4A2C4671D03C9189" pageId="0" pageNumber="95" type="reference_group">
<paragraph id="5A6D8242E8B3D5A7E762B69129A6EAEF" pageId="0" pageNumber="95">
<taxonomicName id="E39A21137081F0CC9FA4FA9DD1239890" class="Insecta" family="Carabidae" genus="Feroperis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feroperis" order="Coleoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="genus">Feroperis</taxonomicName>
Lafer, 1979: 5. Type species:
<taxonomicName class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia jugens" order="Coleoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="jugens">Feronia jugens</taxonomicName>
<normalizedToken originalValue="Tschitschérine">Tschitscherine</normalizedToken>
<taxonomicName id="9F5E6606E3EBFEA186539D4DCB7FEE0B" class="Insecta" family="Carabidae" genus="Feronia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Feronia jugens" order="Coleoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="jugens">Feronia jugens</taxonomicName>
<normalizedToken id="76AF09E1F8853CE20FF81F8B1C00261D" originalValue="Tschitschérine">Tschitscherine</normalizedToken>
, 1893, by original designation (
<bibRefCitation pageId="0" pageNumber="95">Lafer 1979</bibRefCitation>
<bibRefCitation id="12C51638609F621C5602ABB62F106598" pageId="0" pageNumber="95">Lafer 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="95" type="diagnosis">
<paragraph pageId="0" pageNumber="95">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="95">
<subSubSection id="A7971E63369FFCB10220DE88EB1E96D2" pageId="0" pageNumber="95" type="diagnosis">
<paragraph id="87E6CDA7CB2C9BEDA78C05C96FB08BF8" pageId="0" pageNumber="95">Diagnosis.</paragraph>
<paragraph id="8079A0A8EC067B3026DE8CEBEBCA5B57" pageId="0" pageNumber="95">
Body of medium size (13.0-17.5 mm), surface completely black and polished, palpi brown, legs and antennae black or dark brown. Head with very weak punctures; eyes large and convex; antennae reaching the base of pronotum. Pronotum more or less round, 1.3-1.6 times wider than the head; anterior angles rounded, moderately or strongly protruding; lateral border gradually widened and then narrowed towards the base, maximum width near anterior third; lateral channel narrow in front but expanding towards the base in the posterior half; basal margin slightly concave in the middle, either rectilinear or obliqued on the sides; basal foveae usually slightly punctate, outer basal foveal groove deep, reaching the basal margin, the inner basal foveal groove shallower and not reaching the basal margin; carinae between lateral margin and outer basal foveae clearly expressed, perpendicular or inclined towards basal margin, well-separated from the lateral channel and approaching to it near the basal margin; pronotum with two lateral setae, one at the posterior angle, the other one near the pronotal maximum width; pronotum disc with transverse wrinkles. Elytra wide, 1.10-1.35 times wider than pronotum, 1.5-1.6 times wider than elytral width; lateral margins subparallel, widest in the middle; basal ridge continuous, forming an indistinct obtuse angle with the elytral lateral margin; humeral teeth small but distinct; striae deeply incised, without or with very fine punctures; intervals moderately convex; parascutellar pore present; scutellar striae present; third interval with three or more setigerous pores, usually 3-4, rarely 5-6, usually adjacent to the second striae; seventh interval with two preapical pores; umbilical setigerous series on the ninth interval, interrupted in the middle. Hind wings very small, not functional. Metepisternum short, its length along inner margin subequal to the width of anterior margin; sterna
<normalizedToken originalValue="IV–VI">IV-VI</normalizedToken>
<normalizedToken id="70B8D17AED7074322B0DA682F7AC28BE" originalValue="IV–VI">IV-VI</normalizedToken>
with a pair of central setae; males with one pair, females with two pairs of marginal setae on sternum VII, slightly removed from the apical margin. Mesofemora and metafemora with two setae on posterior margin; metatrochanters without setae; metacoxae with two setae; fifth tarsomeres setose beneath. Median lobe of aedeagus slender, bent usually at about 90 degrees; median lobe almost straight in median portion (between the basal bend and apical lamella); apical orifice opened to the dorsal-left side; the shape of the apical portion of apical lamella shows species-specific differences. Stylomere 2 with two ensiform setae at the outer margin and one ensiform seta in the middle part of its inner margin; two nematiform setae in a short fovea near the apex of inner surface (i.e.
<bibRefCitation pageId="0" pageNumber="95">Lafer 1979</bibRefCitation>
<bibRefCitation id="B38C4601CBBD6CB5A0199157833123A9" pageId="0" pageNumber="95">Lafer 1979</bibRefCitation>
).
</paragraph>
</subSubSection>