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<mods:title id="A0ECD490819473437A46A917846F4DB7">New records of Pauropoda (Myriapoda) from north-western Thailand</mods:title>
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<mods:date>2011</mods:date>
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<treatment ID-GBIF-Taxon="152016694" LSID="urn:lsid:zoobank.org:act:CC141A91-BC0A-4E8B-B550-BDBE60F9D981" httpUri="http://treatment.plazi.org/id/869BB2FAE9A6620B7CA252495DC90A9A" lastPageId="20" lastPageNumber="71" pageId="18" pageNumber="69">
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<subSubSection pageId="18" pageNumber="69" type="nomenclature">
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||||
<paragraph pageId="18" pageNumber="69">
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||||
<taxonomicName LSID="urn:lsid:zoobank.org:act:CC141A91-BC0A-4E8B-B550-BDBE60F9D981" class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus choanephorus" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="Choanephorus">Samarangopus choanephorus</taxonomicName>
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||||
<taxonomicNameLabel pageId="18" pageNumber="69">sp. n.</taxonomicNameLabel>
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||||
<treatment id="869BB2FAE9A6620B7CA252495DC90A9A" ID-GBIF-Taxon="152016694" LSID="urn:lsid:zoobank.org:act:CC141A91-BC0A-4E8B-B550-BDBE60F9D981" httpUri="http://treatment.plazi.org/id/869BB2FAE9A6620B7CA252495DC90A9A" lastPageId="20" lastPageNumber="71" pageId="18" pageNumber="69" scope_class="Pauropoda">
|
||||
<subSubSection id="33667376047A8DFE69AE8B910121B470" pageId="18" pageNumber="69" type="nomenclature">
|
||||
<paragraph id="01D9E396E5C3A0746E912A444982CD0B" pageId="18" pageNumber="69">
|
||||
<taxonomicName id="7BB91BCAB331C6D1F303A5BEA3F1B42B" ID-CoL="9B8R8" LSID="urn:lsid:zoobank.org:act:CC141A91-BC0A-4E8B-B550-BDBE60F9D981" class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus choanephorus" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="choanephorus">Samarangopus choanephorus</taxonomicName>
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||||
<taxonomicNameLabel id="CBB404AB9CC648D95C234924DC0FFD65" pageId="18" pageNumber="69">sp. n.</taxonomicNameLabel>
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||||
Figs 72-83
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</paragraph>
|
||||
</subSubSection>
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||||
<subSubSection pageId="18" pageNumber="69" type="material">
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<paragraph pageId="18" pageNumber="69">Material.</paragraph>
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<paragraph pageId="18" pageNumber="69">Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, below the top, primary rain forest, soil, alt. 2400 m, 1991.vii.4, loc. CM-195. - 1 specimen.</paragraph>
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||||
<subSubSection id="355F388FBA2AEC5FD569D82F63E7219E" pageId="18" pageNumber="69" type="material">
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||||
<paragraph id="8512E00D9A2B678613B01950265941F7" pageId="18" pageNumber="69">Material.</paragraph>
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<paragraph id="9E977A9DD1C097DF5A9C38049313DE8F" pageId="18" pageNumber="69">Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, below the top, primary rain forest, soil, alt. 2400 m, 1991.vii.4, loc. CM-195. - 1 specimen.</paragraph>
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</subSubSection>
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||||
<subSubSection pageId="18" pageNumber="69" type="etymology">
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||||
<paragraph pageId="18" pageNumber="69">Etymology.</paragraph>
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||||
<paragraph pageId="18" pageNumber="69">From the Greek choane, funnel, and phero, bear, carry (referring to the funnel-shaped organs on the tergites).</paragraph>
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||||
<subSubSection id="F96E712D3C3023220F9205C531C2BF98" pageId="18" pageNumber="69" type="etymology">
|
||||
<paragraph id="D3425D5D9D90E27BA7899D391B7B512B" pageId="18" pageNumber="69">Etymology.</paragraph>
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||||
<paragraph id="2A53DBD11871F627E74E5EE9F9EDDDF2" pageId="18" pageNumber="69">From the Greek choane, funnel, and phero, bear, carry (referring to the funnel-shaped organs on the tergites).</paragraph>
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||||
</subSubSection>
|
||||
<subSubSection lastPageId="19" lastPageNumber="70" pageId="18" pageNumber="69" type="diagnosis">
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||||
<paragraph pageId="18" pageNumber="69">Diagnosis.</paragraph>
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||||
<paragraph lastPageId="19" lastPageNumber="70" pageId="18" pageNumber="69">
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<taxonomicName class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus choanephorus" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="Choanephorus">Samarangopus choanephorus</taxonomicName>
|
||||
<subSubSection id="64D3D697ABC03E68FB38D493613605A5" lastPageId="19" lastPageNumber="70" pageId="18" pageNumber="69" type="diagnosis">
|
||||
<paragraph id="CA3774A22DDD36CC2B57480C99770C58" pageId="18" pageNumber="69">Diagnosis.</paragraph>
|
||||
<paragraph id="337D1B5579E770C431111ABA2D1ADFA6" lastPageId="19" lastPageNumber="70" pageId="18" pageNumber="69">
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||||
<taxonomicName id="97632F27C43B2A9B73352AA0D0C4EEA2" class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus choanephorus" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="choanephorus">Samarangopus choanephorus</taxonomicName>
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sp. n. may be grouped together with
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<taxonomicName class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus umbonifer" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="Umbonifer">Samarangopus umbonifer</taxonomicName>
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<taxonomicName id="579980594B21BFB1AD86D1F97A0831AB" class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus umbonifer" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="umbonifer">Samarangopus umbonifer</taxonomicName>
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Scheller from Thailand (
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<bibRefCitation author="Scheller, U" journalOrPublisher="Tropical Zoology" pageId="22" pageNumber="73" pagination="7 - 41" title="Pauropoda (Pauropodidae Eurypauropodidae) from north-western Thailand." volume="8" year="1995">Scheller 1995</bibRefCitation>
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||||
<bibRefCitation id="73DD90BDD8B9EF13C8B14E74B9C5FF07" author="Scheller, U" journalOrPublisher="Tropical Zoology" pageId="22" pageNumber="73" pagination="7 - 41" title="Pauropoda (Pauropodidae Eurypauropodidae) from north-western Thailand." volume="8" year="1995">Scheller 1995</bibRefCitation>
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) and
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<taxonomicName class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus condylus" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="Condylus">Samarangopus condylus</taxonomicName>
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<taxonomicName id="08141F99CB11356BE7C09CF4AF37895B" class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus condylus" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="condylus">Samarangopus condylus</taxonomicName>
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Scheller from the Philippines (
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<bibRefCitation author="Scheller, U" journalOrPublisher="International Journal of Myriapodology" pageId="22" pageNumber="73" pagination="69 - 148" title="Records of Pauropoda (Pauropodidae, Brachypauropodidae, Eurypauropodidae) from Indonesia and the Philippines with description of a new genus and 26 new species." url="doi:10.1163/187525409X12577705044548" volume="2" year="2009">Scheller 2009</bibRefCitation>
|
||||
<bibRefCitation id="BA53A8929774B739EFFD16F128C624F4" author="Scheller, U" journalOrPublisher="International Journal of Myriapodology" pageId="22" pageNumber="73" pagination="69 - 148" title="Records of Pauropoda (Pauropodidae, Brachypauropodidae, Eurypauropodidae) from Indonesia and the Philippines with description of a new genus and 26 new species." url="doi:10.1163/187525409X12577705044548" volume="2" year="2009">Scheller 2009</bibRefCitation>
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), for certain more to the former than to the latter. Good distinguishing characters in relation to
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<taxonomicName class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus umbonifer" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="Umbonifer">Samarangopus umbonifer</taxonomicName>
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||||
<taxonomicName id="FDD9C2B9FDEF786810485E64C77BB0E3" class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus umbonifer" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="umbonifer">Samarangopus umbonifer</taxonomicName>
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||||
are the shape of the antennal globulus
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<pageBreakToken pageId="19" pageNumber="70" start="start">g</pageBreakToken>
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<pageBreakToken id="50068C9E6201DE370CE1654E3A36C0ED" pageId="19" pageNumber="70" start="start">g</pageBreakToken>
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, stalk much longer than globulus in
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<taxonomicName class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus choanephorus" order="Tetramerocerata" pageId="19" pageNumber="70" phylum="Arthropoda" rank="species" species="Choanephorus">Borneopauropus choanephorus</taxonomicName>
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<taxonomicName id="D4399285FD8FB86F54AD9DCFE5906550" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus choanephorus" order="Tetramerocerata" pageId="19" pageNumber="70" phylum="Arthropoda" rank="species" species="choanephorus">Borneopauropus choanephorus</taxonomicName>
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, shorter in
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<taxonomicName class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus umbonifer" order="Tetramerocerata" pageId="19" pageNumber="70" phylum="Arthropoda" rank="species" species="Umbonifer">Borneopauropus umbonifer</taxonomicName>
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||||
<taxonomicName id="F3B0DBA0E2F94EF624E19FD08D72AD5E" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus umbonifer" order="Tetramerocerata" pageId="19" pageNumber="70" phylum="Arthropoda" rank="species" species="umbonifer">Borneopauropus umbonifer</taxonomicName>
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, the large funnel-shaped organs of the tergites, with central rod, not without, the protuberances of the posterior margin of the tergite VI, with evenly convex lateral margins, not with a small knob on each side, the shape of the setae on coxa and trochanter of leg 9, simple, not furcate, and the length of the pygidial setae b3, almost as long as the b2, not distinctly shorter.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="20" lastPageNumber="71" pageId="19" pageNumber="70" type="description">
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<paragraph pageId="19" pageNumber="70">Description.</paragraph>
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<paragraph pageId="19" pageNumber="70">
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<subSubSection id="E6530B6B037030B14033D145D56541E7" lastPageId="20" lastPageNumber="71" pageId="19" pageNumber="70" type="description">
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<paragraph id="4974C413AE80C0B7FE6CDDFFAC8EB47C" pageId="19" pageNumber="70">Description.</paragraph>
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<paragraph id="2985B6F43FC2BEE3B97C0C17FA666083" pageId="19" pageNumber="70">
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Length. 0.74 mm. Antennae (Fig. 72). Chaetotaxy of segments 1-4: 2/2/2/2; no
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<normalizedToken originalValue="g´">g'</normalizedToken>
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<normalizedToken id="82267B9F2F11A78F49AFEC080032BD11" originalValue="g´">g'</normalizedToken>
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. Setae thin striate, on segment 4 only two proved, their relative lengths p=10,
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<normalizedToken originalValue="p´or">p'or</normalizedToken>
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<normalizedToken originalValue="p´´">p''</normalizedToken>
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<normalizedToken id="62BE2138A96135AD22FEC8A534B67F2A" originalValue="p´or">p'or</normalizedToken>
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<normalizedToken id="5941F1A7793855A9ECA168F11925B2F8" originalValue="p´´">p''</normalizedToken>
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=6. Tergal branch t almost cylindrical, 3.1 times as wide as greatest diameter and as long as sternal branch s. That branch twice longer than greatest diameter, anterodistal corner distinctly truncate. Seta q as setae of 4th segment, 0.5 of the length of s. Relative lengths of flagella (base segments included) and base segments: F1=100, bs1=18, F2=?, bs2~10, F3=85, bs3=17. F1 5.6 times as long as t, F3 3.3 times as long as s. Calyces conical; flagella axes inconsiderably widened below calyx. Globulus g 1.6 times as long as greatest width; 10 thin bracts, capsule spherical, diameter of g 0.8 of the greatest diameter of t. Antenna glabrous.
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</paragraph>
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<paragraph pageId="19" pageNumber="70">Trunk (Figs 73-77). Setae of collum segment (Fig. 73) similar thin furcate, main branch pointed striate, secondary branch rudimentary pointed; sublateral setae 1.3 times as long as submedian setae. Sternite process broad triangular pointed anteriorly. Appendages wide cylindrical, caps flat. All parts of collum segment glabrous.</paragraph>
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<paragraph pageId="19" pageNumber="70">Tergites with four main types of protuberances (Figs 74-77): 1. stalked campanulate marginal protuberances (Figs 74-77), short on anterior margin of tergite I, larger on lateral margins of tergites; 2. fungiform organs (Figs 74-77) with hat in the shape of an upside down transparent funnel with a central rod, foot subcylindrical, these distributed over whole the surface of tergites; 3. small subcylindrical organs with upside down funnel at top (Fig. 77); 4. many small conical structures in between the different protuberances. Number of marginal protuberances: (holotype only): I - 36, II, 1 small - T1 - 11; III, 8 - T2 - 8; IV, 8 - T3 - 6; V, 9 - T4 - 4, VI, 7 - T5 - 1.</paragraph>
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<paragraph pageId="19" pageNumber="70">
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<paragraph id="23C3102CEFD248ADDCFFB9E52120FB1D" pageId="19" pageNumber="70">Trunk (Figs 73-77). Setae of collum segment (Fig. 73) similar thin furcate, main branch pointed striate, secondary branch rudimentary pointed; sublateral setae 1.3 times as long as submedian setae. Sternite process broad triangular pointed anteriorly. Appendages wide cylindrical, caps flat. All parts of collum segment glabrous.</paragraph>
|
||||
<paragraph id="84CF2A40FD6F0F5D1D3192932AC52EEA" pageId="19" pageNumber="70">Tergites with four main types of protuberances (Figs 74-77): 1. stalked campanulate marginal protuberances (Figs 74-77), short on anterior margin of tergite I, larger on lateral margins of tergites; 2. fungiform organs (Figs 74-77) with hat in the shape of an upside down transparent funnel with a central rod, foot subcylindrical, these distributed over whole the surface of tergites; 3. small subcylindrical organs with upside down funnel at top (Fig. 77); 4. many small conical structures in between the different protuberances. Number of marginal protuberances: (holotype only): I - 36, II, 1 small - T1 - 11; III, 8 - T2 - 8; IV, 8 - T3 - 6; V, 9 - T4 - 4, VI, 7 - T5 - 1.</paragraph>
|
||||
<paragraph id="8946B6D5A3CEC0959A584642B608E091" pageId="19" pageNumber="70">
|
||||
Bothriotricha. All but T3 with very thin axes, curved distally. T3 (Fig. 78) with thicker axes distally forming a subcylindrical, almost glabrous, swelling, length
|
||||
<normalizedToken originalValue="¼">1/4</normalizedToken>
|
||||
<normalizedToken id="5A1D93630950D8A589815EAC2E7FF0B9" originalValue="¼">1/4</normalizedToken>
|
||||
of the length of bothriotrix. Relative lengths of bothriotricha (holotype only): T1=100, T2=95, T3=42, T4≈90, T5=89.
|
||||
</paragraph>
|
||||
<paragraph pageId="19" pageNumber="70">Genital papillae (Fig. 79). Conical, glabrous, 2.2 times as long as greatest diameter, setae long, 0.8 of the length of papillae, inserted below the middle of papillae.</paragraph>
|
||||
<paragraph pageId="19" pageNumber="70">Legs (Figs 80, 81).All legs 5-segmented. Seta on coxa and trochanter (Fig. 80) of legs 1-9thin simple striate. Tarsi tapering, those of leg 9 (Fig. 81) 3.7 times as long as its greatest diameter; two tergal setae, both pointed glabrous; length of proximal one 0.4 of the length of tarsus and 3 times longer than distal seta. Cuticle of tarsus glabrous. No proximal seta on tarsi of leg 1. All legs with large main claw and small setose anterior secondary claw; in leg 9 the former reaching 0.5 of the length of tarsus. Appendage on femur of leg 1 not studied.</paragraph>
|
||||
<paragraph lastPageId="20" lastPageNumber="71" pageId="19" pageNumber="70">
|
||||
<paragraph id="2541C97F9329DE2BF5AAD2BEF7C5F92C" pageId="19" pageNumber="70">Genital papillae (Fig. 79). Conical, glabrous, 2.2 times as long as greatest diameter, setae long, 0.8 of the length of papillae, inserted below the middle of papillae.</paragraph>
|
||||
<paragraph id="F7CE71A1E07B82559C99A9CD663ED37C" pageId="19" pageNumber="70">Legs (Figs 80, 81).All legs 5-segmented. Seta on coxa and trochanter (Fig. 80) of legs 1-9thin simple striate. Tarsi tapering, those of leg 9 (Fig. 81) 3.7 times as long as its greatest diameter; two tergal setae, both pointed glabrous; length of proximal one 0.4 of the length of tarsus and 3 times longer than distal seta. Cuticle of tarsus glabrous. No proximal seta on tarsi of leg 1. All legs with large main claw and small setose anterior secondary claw; in leg 9 the former reaching 0.5 of the length of tarsus. Appendage on femur of leg 1 not studied.</paragraph>
|
||||
<paragraph id="4F62D068A172CD30CF9D420CE302A8F9" lastPageId="20" lastPageNumber="71" pageId="19" pageNumber="70">
|
||||
Pygidium (Figs 82, 83). Tergum. Posterior margin undulate with larger pentagonal lobe between st. Setae a1 cylindrical, curved inward, converging, a2 short clavate straight
|
||||
<pageBreakToken pageId="20" pageNumber="71" start="start">converging</pageBreakToken>
|
||||
<pageBreakToken id="6435D7CDA997F8922972DF6778969BB1" pageId="20" pageNumber="71" start="start">converging</pageBreakToken>
|
||||
, faintly pubescent distally, a3 long thin tapering pointed diverging, st straight lanceolate, faintly pubescent. Relative lengths of setae: a1=10, a2=6(-7), a3=(20-)21, st=(15-)16. Distance a1-a1 1.3 times as long as a1, distance a1-a2 as long as distance a2 -a3; distance st-st almost as long as st and 1.1 times as long as distance a1-a1. Cuticle glabrous.
|
||||
</paragraph>
|
||||
<paragraph pageId="20" pageNumber="71">Sternum (Fig 83). Posterior margin rounded. Setae long tapering pointed, b2 straight diverging, b3 somewhat curved inward. Relative lengths of setae (pygidial a1=10): b1=(45-)47, b2=(26-)29, b3=24(-25). b1 1.1 times as long as interdistance, b2 1.1 times as long as distance b1-b2, b3 0.6 of interdistance.</paragraph>
|
||||
<paragraph pageId="20" pageNumber="71">Anal plate (Fig. 82) 1.4 times as long as broad, lateral margins convex, distal part of plate cleft by a V-shaped incision into two somewhat tapering cylindrical branches; four appendages: each branch provided with one outer and one inner tooth and in between a straight bladder-shaped appendage, that broadest in distal half, length of appendage 0.4 of the length of plate; two short thin cylindrical appendages protruding outward-backward from lateral margins. Plate glabrous, appendages with faint pubescence.</paragraph>
|
||||
<caption pageId="20" pageNumber="71">
|
||||
<paragraph pageId="20" pageNumber="71">
|
||||
<paragraph id="2D9D1FB91DA54F3AABC4A01C00BE39FD" pageId="20" pageNumber="71">Sternum (Fig 83). Posterior margin rounded. Setae long tapering pointed, b2 straight diverging, b3 somewhat curved inward. Relative lengths of setae (pygidial a1=10): b1=(45-)47, b2=(26-)29, b3=24(-25). b1 1.1 times as long as interdistance, b2 1.1 times as long as distance b1-b2, b3 0.6 of interdistance.</paragraph>
|
||||
<paragraph id="77517E35D21D844B96521B5A4890990B" pageId="20" pageNumber="71">Anal plate (Fig. 82) 1.4 times as long as broad, lateral margins convex, distal part of plate cleft by a V-shaped incision into two somewhat tapering cylindrical branches; four appendages: each branch provided with one outer and one inner tooth and in between a straight bladder-shaped appendage, that broadest in distal half, length of appendage 0.4 of the length of plate; two short thin cylindrical appendages protruding outward-backward from lateral margins. Plate glabrous, appendages with faint pubescence.</paragraph>
|
||||
<caption id="1BAA4FAEAA0128208D881E718729633A" pageId="20" pageNumber="71">
|
||||
<paragraph id="87D69C506AD23773D58855933C207749" pageId="20" pageNumber="71">
|
||||
Figures 72-83.
|
||||
<taxonomicName class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus choanephorus" order="Tetramerocerata" pageId="20" pageNumber="71" phylum="Arthropoda" rank="species" species="Choanephorus">Samarangopus choanephorus</taxonomicName>
|
||||
<taxonomicName id="EEF78777BAB401D7A6ABED2DCAABF560" class="Pauropoda" family="Eurypauropodidae" genus="Samarangopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Samarangopus choanephorus" order="Tetramerocerata" pageId="20" pageNumber="71" phylum="Arthropoda" rank="species" species="choanephorus">Samarangopus choanephorus</taxonomicName>
|
||||
sp. n., holotype ad. 9(♂) 72 left antenna, sternal view 73 collum segment, median and left part 74 tergite I, anterolateral margin with campanulate protuberances and fungiform organs 75 tergite I, posterolateral corner 76 tergite IV, right lateral margin around insertion pit of T3 77 tergite VI, right posterolateral part 78T3 79 genital papillae, anterior view 80 seta on trochanter of leg 9 81 tarsus of leg 9 82 pygidial tergum, posterior part, tergal view 83 pygidial sternum, sternal view. Scale: a: Fig. 79; b: Figs 74-76; c: Figs 72, 73, 77, 78, 80, 81; d: Figs 82, 83.
|
||||
</paragraph>
|
||||
</caption>
|
||||
|
|
|
|||
|
|
@ -1,104 +1,104 @@
|
|||
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<treatment ID-GBIF-Taxon="152016689" LSID="urn:lsid:zoobank.org:act:786F9B4C-205F-44C5-B98F-99128031C94C" httpUri="http://treatment.plazi.org/id/86E841A1C2CCE00BB6CA40C95074D93F" lastPageId="18" lastPageNumber="69" pageId="15" pageNumber="66">
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||||
<subSubSection pageId="15" pageNumber="66" type="nomenclature">
|
||||
<paragraph pageId="15" pageNumber="66">
|
||||
<taxonomicName LSID="urn:lsid:zoobank.org:act:786F9B4C-205F-44C5-B98F-99128031C94C" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus platylopas" order="Tetramerocerata" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="Platylopas">Borneopauropus platylopas</taxonomicName>
|
||||
<taxonomicNameLabel pageId="15" pageNumber="66">sp. n.</taxonomicNameLabel>
|
||||
<treatment id="86E841A1C2CCE00BB6CA40C95074D93F" ID-GBIF-Taxon="152016689" LSID="urn:lsid:zoobank.org:act:786F9B4C-205F-44C5-B98F-99128031C94C" httpUri="http://treatment.plazi.org/id/86E841A1C2CCE00BB6CA40C95074D93F" lastPageId="18" lastPageNumber="69" pageId="15" pageNumber="66" scope_class="Pauropoda">
|
||||
<subSubSection id="EAB4260377CF6D601A9A057B3354FE67" pageId="15" pageNumber="66" type="nomenclature">
|
||||
<paragraph id="52DB9121BF60921863209DF3BF899AFC" pageId="15" pageNumber="66">
|
||||
<taxonomicName id="29061DEB680BFF7F5EEE0601704B9467" ID-CoL="9B7YN" LSID="urn:lsid:zoobank.org:act:786F9B4C-205F-44C5-B98F-99128031C94C" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus platylopas" order="Tetramerocerata" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="platylopas">Borneopauropus platylopas</taxonomicName>
|
||||
<taxonomicNameLabel id="6C5FB4030ADA65D89173DF3BBBAC83D3" pageId="15" pageNumber="66">sp. n.</taxonomicNameLabel>
|
||||
Figs 61-71
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="15" pageNumber="66" type="material">
|
||||
<paragraph pageId="15" pageNumber="66">Material.</paragraph>
|
||||
<paragraph pageId="15" pageNumber="66">Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1992.vi.30, loc. CM-080. - Paratype, ad. 9(♀), ibidem, loc. CM-194. - 2 specimens.</paragraph>
|
||||
<subSubSection id="16EB2FCDF444700DADADD7EF300927DD" pageId="15" pageNumber="66" type="material">
|
||||
<paragraph id="5978DBA641A9460EFB32B5F048BBF1ED" pageId="15" pageNumber="66">Material.</paragraph>
|
||||
<paragraph id="526D26CCC661CC455593F08CFD034E56" pageId="15" pageNumber="66">Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1992.vi.30, loc. CM-080. - Paratype, ad. 9(♀), ibidem, loc. CM-194. - 2 specimens.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection pageId="15" pageNumber="66" type="etymology">
|
||||
<paragraph pageId="15" pageNumber="66">Etymology.</paragraph>
|
||||
<paragraph pageId="15" pageNumber="66">From the Greek platys, broad, and lopas, plate (referring to the unusually broad anal plate).</paragraph>
|
||||
<subSubSection id="835BE0317E64B838BB7979D22A923D2E" pageId="15" pageNumber="66" type="etymology">
|
||||
<paragraph id="3FB2073AE99495D3B9B8944418C3812F" pageId="15" pageNumber="66">Etymology.</paragraph>
|
||||
<paragraph id="857B99A3FC3007160ADFB41B838E7F67" pageId="15" pageNumber="66">From the Greek platys, broad, and lopas, plate (referring to the unusually broad anal plate).</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection lastPageId="18" lastPageNumber="69" pageId="15" pageNumber="66" type="diagnosis">
|
||||
<paragraph pageId="15" pageNumber="66">Diagnosis.</paragraph>
|
||||
<paragraph lastPageId="17" lastPageNumber="68" pageId="15" pageNumber="66">
|
||||
<subSubSection id="A4F7CB62597D7B0A7035791B832BACBD" lastPageId="18" lastPageNumber="69" pageId="15" pageNumber="66" type="diagnosis">
|
||||
<paragraph id="5C3955B1CDA8D916092A0E02E56AA649" pageId="15" pageNumber="66">Diagnosis.</paragraph>
|
||||
<paragraph id="4BB8CA1F3BD4DD99AE457A89CDC05180" lastPageId="17" lastPageNumber="68" pageId="15" pageNumber="66">
|
||||
Up to now four species have been described in the genus, one from Sulawesi in Indonesia, two from Sabah and one from Tasmania.
|
||||
<taxonomicName class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus platylopas" order="Tetramerocerata" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="Platylopas">Borneopauropus platylopas</taxonomicName>
|
||||
<taxonomicName id="E8CA16BD756324EC1477B4F4B108A137" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus platylopas" order="Tetramerocerata" pageId="15" pageNumber="66" phylum="Arthropoda" rank="species" species="platylopas">Borneopauropus platylopas</taxonomicName>
|
||||
has most characters
|
||||
<pageBreakToken pageId="16" pageNumber="67" start="start">in</pageBreakToken>
|
||||
<pageBreakToken id="531EA85AAF7121A3BDF1803C01F4DC79" pageId="16" pageNumber="67" start="start">in</pageBreakToken>
|
||||
common with the Indonesian species,
|
||||
<taxonomicName class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus curtipes" order="Tetramerocerata" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="Curtipes">Borneopauropus curtipes</taxonomicName>
|
||||
<taxonomicName id="B223D7D58B807B31FD6463CB233A842A" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus curtipes" order="Tetramerocerata" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="curtipes">Borneopauropus curtipes</taxonomicName>
|
||||
Scheller (
|
||||
<bibRefCitation author="Scheller, U" journalOrPublisher="International Journal of Myriapodology" pageId="22" pageNumber="73" pagination="69 - 148" title="Records of Pauropoda (Pauropodidae, Brachypauropodidae, Eurypauropodidae) from Indonesia and the Philippines with description of a new genus and 26 new species." url="doi:10.1163/187525409X12577705044548" volume="2" year="2009">Scheller 2009</bibRefCitation>
|
||||
<bibRefCitation id="FF658F82E85B91D990BA49CC746E3489" author="Scheller, U" journalOrPublisher="International Journal of Myriapodology" pageId="22" pageNumber="73" pagination="69 - 148" title="Records of Pauropoda (Pauropodidae, Brachypauropodidae, Eurypauropodidae) from Indonesia and the Philippines with description of a new genus and 26 new species." url="doi:10.1163/187525409X12577705044548" volume="2" year="2009">Scheller 2009</bibRefCitation>
|
||||
) and one of the two species from Sabah,
|
||||
<taxonomicName class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus penanorum" order="Tetramerocerata" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="Penanorum">Borneopauropus penanorum</taxonomicName>
|
||||
<taxonomicName id="62E155A7F02D24F8983F910D8E901F80" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus penanorum" order="Tetramerocerata" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="penanorum">Borneopauropus penanorum</taxonomicName>
|
||||
Scheller (
|
||||
<bibRefCitation author="Scheller, U" journalOrPublisher="Stobaeana" pageId="22" pageNumber="73" pagination="1 - 14" title="First record of Pauropoda (Myriapoda) from Borneo." volume="1" year="1994">Scheller et al. 1994</bibRefCitation>
|
||||
<bibRefCitation id="8648290518CC5898EF429F790557FCC8" author="Scheller, U" journalOrPublisher="Stobaeana" pageId="22" pageNumber="73" pagination="1 - 14" title="First record of Pauropoda (Myriapoda) from Borneo." volume="1" year="1994">Scheller et al. 1994</bibRefCitation>
|
||||
). It can be distinguished from both by the shape of the bothriotricha T3, clavate part moderately widened with long distal flagellum in
|
||||
<taxonomicName class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus platylopas" order="Tetramerocerata" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="Platylopas">Borneopauropus platylopas</taxonomicName>
|
||||
<taxonomicName id="8439DC7F0B35976A2B284221421F8CE0" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus platylopas" order="Tetramerocerata" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="platylopas">Borneopauropus platylopas</taxonomicName>
|
||||
, not strongly widened and with shorter flagellum in
|
||||
<taxonomicName class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus curtipes" order="Tetramerocerata" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="Curtipes">Borneopauropus curtipes</taxonomicName>
|
||||
<pageBreakToken pageId="17" pageNumber="68" start="start">and</pageBreakToken>
|
||||
<taxonomicName class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus penanorum" order="Tetramerocerata" pageId="17" pageNumber="68" phylum="Arthropoda" rank="species" species="Penanorum">Borneopauropus penanorum</taxonomicName>
|
||||
<taxonomicName id="586CDECBA7400DFEC5F467E14DA10BDE" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus curtipes" order="Tetramerocerata" pageId="16" pageNumber="67" phylum="Arthropoda" rank="species" species="curtipes">Borneopauropus curtipes</taxonomicName>
|
||||
<pageBreakToken id="F5D99B3F9886BC623942038D9D4A310D" pageId="17" pageNumber="68" start="start">and</pageBreakToken>
|
||||
<taxonomicName id="67C7E0AF317C5B070EB422BF39549862" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus penanorum" order="Tetramerocerata" pageId="17" pageNumber="68" phylum="Arthropoda" rank="species" species="penanorum">Borneopauropus penanorum</taxonomicName>
|
||||
, and by the shape of the pygidial setae a3, long thin tapering, not bladder-shaped, and the shape of the anal plate, broad triangular, not longish linguiform.
|
||||
</paragraph>
|
||||
<paragraph pageId="17" pageNumber="68">Description. Length. (0.51-)0.58 mm. Head (Fig. 61). Tergal and lateral sides with 26 setae arranged as in Fig. 61; transversal rows indistinct laterally. Setae longest in posteromedian part; all but lateral setae clavate, shortly pubescent, lateral group setae cylindrical annulate. Temporal organs small with at least two short uplifted extensions. Head cuticle glabrous.</paragraph>
|
||||
<paragraph pageId="17" pageNumber="68">
|
||||
<paragraph id="A2D9FB5B81BF90032AD8862ED08EB2C4" pageId="17" pageNumber="68">Description. Length. (0.51-)0.58 mm. Head (Fig. 61). Tergal and lateral sides with 26 setae arranged as in Fig. 61; transversal rows indistinct laterally. Setae longest in posteromedian part; all but lateral setae clavate, shortly pubescent, lateral group setae cylindrical annulate. Temporal organs small with at least two short uplifted extensions. Head cuticle glabrous.</paragraph>
|
||||
<paragraph id="EB0C6EFBDC9931A5D78148A18ED69A2D" pageId="17" pageNumber="68">
|
||||
Antennae (Fig. 62). Segment 4 with 4 cylindrical annulate setae; relative lengths: p=10,
|
||||
<normalizedToken originalValue="p´">p'</normalizedToken>
|
||||
<normalizedToken id="F75C550B4D755638560BEC549FEE00E6" originalValue="p´">p'</normalizedToken>
|
||||
=11(-12),
|
||||
<normalizedToken originalValue="p´´">p''</normalizedToken>
|
||||
<normalizedToken id="C56FB564A23E601F218444069D6F4D87" originalValue="p´´">p''</normalizedToken>
|
||||
=4(-6), r=4. Tergal seta p 0.6 of the length of tergal branch t. The latter subcylindrical, (2.6-)2.9 times as long as wide, 0.9 of the length of sternal branch s. That branch thickest in distal third and with anterodistal corner somewhat more truncated than posterodistal corner; s (2.0-)2.1 times as long as greatest diameter, its seta q cylindrical striate, 0.5 of the length of s. Relative lengths of flagella (base segments included) and base segments: F1=100, bs1=8; F2=(65-)66, bs2=6; F3=72(-74), bs3=(7-)8. F1 3.1 times as long as t, F2 and F3 (1.9-)2.2 and (2.0-)2.3 times as long as s respectively. Distal calyces helmet-shaped, glabrous. Globulus g, (1.6-)1.8 times as long as greatest diameter, the latter (0.9-) as long as greatest diameter of t. Antenna glabrous.
|
||||
</paragraph>
|
||||
<paragraph pageId="17" pageNumber="68">Trunk (Figs 63-65). Setae of collum segment (Fig. 63) simple cylindrical blunt annulate; sublateral seta 1.2 times as long as submedian seta. Sternite process blunt-ended; appendages barrel-shaped with low caps; process and appendages glabrous.</paragraph>
|
||||
<paragraph pageId="17" pageNumber="68">Tergites (Figs 64, 65), I, V, VI entire, II, III, IV transversely and weakly 2-parted. Number of setae on tergites (holotype only, if two groups of values anterior and posterior groups of setae respectively): I 30, II 19+20, III 28+21, IV 25+19, V 20+13, VI 4+2. Setae bladder-shaped with short oblique-erect pubescence, setae lengthening posteriorly, those on VI (Fig. 65) about twice longer than those on I. Cuticle of tergites glabrous.</paragraph>
|
||||
<paragraph pageId="17" pageNumber="68">Bothriotricha (Figs 66, 67). Relative lengths: T1=100, T2=108(-110), T3=(122-)146, T4=(89-)113, T5=(140-)144. Axes simple, most proximally glabrous; proximal half of T3 (Fig. 66) strongly clavate with short pubescence arranged in dense whorls, distal half very thin; other bothriotricha (Fig. 67) with thin and curved axes; pubescence short oblique on proximal parts, erect distally.</paragraph>
|
||||
<paragraph pageId="17" pageNumber="68">Genital papillae (Fig. 68). Short, as long as wide, seta 0.5 of the length of papilla.</paragraph>
|
||||
<paragraph pageId="17" pageNumber="68">Legs (Figs 69, 70). All legs 5-segmented. Setae on coxa (Fig. 69) and trochanter of legs 1-9 simple, cylindrical annulate blunt, rudimentary secondary branch only on coxal setae of leg 2 in male. Tarsus of leg 9 (Fig. 70) short, almost cylindrical, (2.9-)3.4 times as long as greatest diameter; setae short cylindrical annulate, proximal one 0.1 of the length of tarsus and (0.5-)0.6 of the length of distal seta. Cuticle of tarsus glabrous.</paragraph>
|
||||
<paragraph pageId="17" pageNumber="68">
|
||||
<paragraph id="F3A47DD144B69356528C6171CB4C6694" pageId="17" pageNumber="68">Trunk (Figs 63-65). Setae of collum segment (Fig. 63) simple cylindrical blunt annulate; sublateral seta 1.2 times as long as submedian seta. Sternite process blunt-ended; appendages barrel-shaped with low caps; process and appendages glabrous.</paragraph>
|
||||
<paragraph id="1E15E9E0720E19DBE6270FA3E334E0B9" pageId="17" pageNumber="68">Tergites (Figs 64, 65), I, V, VI entire, II, III, IV transversely and weakly 2-parted. Number of setae on tergites (holotype only, if two groups of values anterior and posterior groups of setae respectively): I 30, II 19+20, III 28+21, IV 25+19, V 20+13, VI 4+2. Setae bladder-shaped with short oblique-erect pubescence, setae lengthening posteriorly, those on VI (Fig. 65) about twice longer than those on I. Cuticle of tergites glabrous.</paragraph>
|
||||
<paragraph id="7CDD9D25670593E0CF7C7A90D692AFB9" pageId="17" pageNumber="68">Bothriotricha (Figs 66, 67). Relative lengths: T1=100, T2=108(-110), T3=(122-)146, T4=(89-)113, T5=(140-)144. Axes simple, most proximally glabrous; proximal half of T3 (Fig. 66) strongly clavate with short pubescence arranged in dense whorls, distal half very thin; other bothriotricha (Fig. 67) with thin and curved axes; pubescence short oblique on proximal parts, erect distally.</paragraph>
|
||||
<paragraph id="7F4E25F2C9BA13DA78E215297B705E7B" pageId="17" pageNumber="68">Genital papillae (Fig. 68). Short, as long as wide, seta 0.5 of the length of papilla.</paragraph>
|
||||
<paragraph id="6B5134577692483FCC6F90CA5CE5D5F4" pageId="17" pageNumber="68">Legs (Figs 69, 70). All legs 5-segmented. Setae on coxa (Fig. 69) and trochanter of legs 1-9 simple, cylindrical annulate blunt, rudimentary secondary branch only on coxal setae of leg 2 in male. Tarsus of leg 9 (Fig. 70) short, almost cylindrical, (2.9-)3.4 times as long as greatest diameter; setae short cylindrical annulate, proximal one 0.1 of the length of tarsus and (0.5-)0.6 of the length of distal seta. Cuticle of tarsus glabrous.</paragraph>
|
||||
<paragraph id="0A1C85A000EF8F14A40E6C9C9AC62939" pageId="17" pageNumber="68">
|
||||
Pygidium (Fig. 71). Tergum. Hind margin rounded. Relative lengths of setae: a1=10, a2=(13-)14, a3=(22-)25, st rudimentary. a1 bladder-shaped, faintly pubescent, a2 longish clavate, distinctly pubescent, somewhat curved inward, a3 thin tapering, faintly striate, curved inward. Distance a1-a1 1.5 times as long as a1, distance a1-a2 twice longer than distance a2-a3; distance
|
||||
<normalizedToken originalValue="st–st">st-st</normalizedToken>
|
||||
<normalizedToken id="290CF65883BA09A0C3E63F684DD69A89" originalValue="st–st">st-st</normalizedToken>
|
||||
≈25 times as long as st and 0.6 of distance a1-a1.
|
||||
</paragraph>
|
||||
<paragraph lastPageId="18" lastPageNumber="69" pageId="17" pageNumber="68">
|
||||
<paragraph id="2CD9EAE4171D41C49BFE6E9C5F0ED068" lastPageId="18" lastPageNumber="69" pageId="17" pageNumber="68">
|
||||
Sternum. Posterior margin between b1 rounded but with large lobe below anal plate; lobe with median indentation and rounded posterolateral corners. Relative
|
||||
<pageBreakToken pageId="18" pageNumber="69" start="start">lengths</pageBreakToken>
|
||||
<pageBreakToken id="EE4857AEFE79F65CCCCD931FBDFAAF67" pageId="18" pageNumber="69" start="start">lengths</pageBreakToken>
|
||||
of setae (pygidial a1=10): b1=(40-)43, b2=13(-16), b3=7. b1 thin tapering, blunt, striate distally, b2 and b3 cylindrical. b1 (1.3-)1.4 times as long as distance b1-b1, b2 0.7(-0.9) of distance b1-b2 and b3 0.3(-0.4) of distance b3-b3.
|
||||
</paragraph>
|
||||
<paragraph pageId="18" pageNumber="69">Anal plate (Fig. 71) narrowest anteriorly, triangular, posterolateral corner turned anteriorly and posterior margin with small median process, two cylindrical blunt faintly pubescent appendages protruding backward from sternal side just nearby the posteromedian process, length of appendages somewhat longer than plate.</paragraph>
|
||||
<caption pageId="18" pageNumber="69">
|
||||
<paragraph pageId="18" pageNumber="69">
|
||||
<paragraph id="D64A25B79CC3B30B31BF3FFE15DD8573" pageId="18" pageNumber="69">Anal plate (Fig. 71) narrowest anteriorly, triangular, posterolateral corner turned anteriorly and posterior margin with small median process, two cylindrical blunt faintly pubescent appendages protruding backward from sternal side just nearby the posteromedian process, length of appendages somewhat longer than plate.</paragraph>
|
||||
<caption id="446CFAB2C3AB50FFC0B1C84A43A0A284" pageId="18" pageNumber="69">
|
||||
<paragraph id="C65CFD2DB38E8B96786C8D9B40FCFF87" pageId="18" pageNumber="69">
|
||||
Figures 61-71.
|
||||
<taxonomicName class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus platylopas" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="Platylopas">Borneopauropus platylopas</taxonomicName>
|
||||
<taxonomicName id="3F5A458A0A6878DCEA694BE90167166B" class="Pauropoda" family="Brachypauropodidae" genus="Borneopauropus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borneopauropus platylopas" order="Tetramerocerata" pageId="18" pageNumber="69" phylum="Arthropoda" rank="species" species="platylopas">Borneopauropus platylopas</taxonomicName>
|
||||
, sp. n., holotype, ad. 9(♂) 61 head, median and right part, tergal view 62 right antenna, tergal view 63 collum segment, median and left part, sternal view 64 tergites I-II, right half 65 tergite VI, right half 66T3 67T2 68 genital papillae and seta on coxa of leg 2, anterior view 69 seta on coxa of leg 9 70 tarsus of leg 9 71 pygidium, posterior part, tergal view. Scale: a: Figs 64, 65; b: Figs 66-70; c: Fig. 61; d: 62, 63; e: 71.
|
||||
</paragraph>
|
||||
</caption>
|
||||
|
|
|
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