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<mods:title id="F36B275AC0CE057E36A889A63E048493">Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
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<treatment id="CC91D7ABA56E59DEA00AF50EDEA0A505" ID-GBIF-Taxon="199468678" LSID="urn:lsid:zoobank.org:act:F9403230-DD0E-4B48-B597-1954F056BF7E" httpUri="http://treatment.plazi.org/id/CC91D7ABA56E59DEA00AF50EDEA0A505" lastPageNumber="309" pageId="0" pageNumber="309" scope_family="Metasarcidae" scope_order="Opiliones" scope_subOrder="Laniatores">
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3.40.
<taxonomicName LSID="http://zoobank.org/F9403230-DD0E-4B48-B597-1954F056BF7E" authority="Benedetti &amp; Pinto-da-Rocha, 2022" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus beni" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="beni" status="sp. nov.">Metasarcus beni</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="309">sp. nov.</taxonomicNameLabel>
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<taxonomicNameLabel id="A58F136A2F0A13AB8BED684F7DECF7F4" pageId="0" pageNumber="309">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">Figs 5A</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">, 10A, B</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">, 14C, D</figureCitation>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">, 31</figureCitation>
<subSubSection id="20C2FC8526651EEF710372F7BB64057A" pageId="0" pageNumber="309" type="description">
<paragraph id="F30DD3F311BB3923E0725E032B3080FA" pageId="0" pageNumber="309">
<figureCitation id="5280B8D37A71D6F49C28A3B6DF510C46" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">Figs 5A</figureCitation>
<figureCitation id="390BFB3A255219E3030305FBDBE157AC" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">, 10A, B</figureCitation>
<figureCitation id="978268ADA2552ADD128EDC170B3EE67C" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">, 14C, D</figureCitation>
<figureCitation id="3B39AD1185EBE51777051C3AE3FF0471" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">, 31</figureCitation>
</paragraph>
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<paragraph pageId="0" pageNumber="309">Description.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
<subSubSection id="27BD0A21BC68041196D52C691BCFD020" pageId="0" pageNumber="309" type="description">
<paragraph id="83799969E84F9D36E23E55AD51A9EDD8" pageId="0" pageNumber="309">Description.</paragraph>
<paragraph id="887ECAA303FC5192D6B37AE378257646" pageId="0" pageNumber="309">
<emphasis id="0DE5C00F3DE43081BA27FBEDFFA285D3" bold="true" pageId="0" pageNumber="309">
MALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
<emphasis id="77BD0A8B01D0BE709FBA52242F59B006" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
<emphasis id="FC92EF9444BC151D3C93DF4933A90067" italics="true" pageId="0" pageNumber="309">n</emphasis>
=2) DSW: 5.0-5.1 (5.1); DSL: 5.0-5.2 (5.2); CL: 1.2-1.3 (1.3). FIVL: 16.0-17.0 (17). ChL: 3.3-3.4 (3.4).
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
<emphasis id="38EFD36774489CEE70CC9C66271720A3" bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
(in ethanol): Yellow.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
<emphasis id="0534D115FD1D1C089024C9AF72F3E64B" bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
<figureCitation id="1BD606A6D97F2857F2EF2A0B6973C684" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
) Gamma-P-type DSS. Anterior margin of carapace with median elevation, with few granules. Carapace with granules concentrated in the lateral regions to ocularium and sparse behind ocularium. Ocularium with pronounced median depression; granulate and unarmed. Areas I-II granulate; III-IV with sparse granulation. Area I divided by longitudinal groove; with one pair of median tubercles. Areas II and IV unarmed. Area III with two long spines, directed posteriorly. Posterior margin of dorsal scutum and free tergites with few granules distributed. Lateral margins of dorsal scutum with granules along the posterior ⅔.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
<emphasis id="FEC1CEE0650CF02E84EDFFE87153E0DB" bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
<figureCitation id="98DB83436A7A481FBBCBE23D698D77E2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
) Swollen. Segment I granular. Segment II with few granules; finger with seven teeth. Segment III with four teeth.
<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
<taxonomicName id="AACFD4CD17830CFC8421593B478C12AE" genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis id="316C41348D0A8C51931435A4954D737A" bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: Femur-tibia with small tubercles on dorsal surface. Trochanter with a ventroapical setiferous tubercle. Femur a ventral row of 5-6 large spines; a large retroapical spine. Patella with a large retroapical and smaller proapical spines. Tibia: prolateral IIII, retrolateral IIII. Tarsus: prolateral IIi, retrolateral iIi.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
<emphasis id="36B7599AE59F6E68FA4DD121C836C9B5" bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxa I with a row of 5-6 tubercles. Coxae II-IV with sparse granules. Rows of four tiny tubercles between coxae II-III and six tubercles between coxae III-IV. Genital area, free sternites and anal operculum few granulate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
<emphasis id="C426E173D3D6F3A51CE9294525A0D072" bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
<figureCitation id="AA2471AA1E70AA0CB903A73CD706970E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10A, B</figureCitation>
<figureCitation id="42092CB0619A0125CADA7AAC4D23AEDA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10A, B</figureCitation>
) Coxae I-II each with a prolateral and a retrolateral apophysis. Coxa III unarmed. Coxa IV with few sparse granules. Trochanters I-III unarmed and smooth. Trochanter IV unarmed and granular. Femora I-III unarmed and with small granules. Femur IV granular; a distal retrodorsal small apophysis; a distal retroventral row of 11-12 small tubercles. Patella-tibiae I-IV unarmed, with few granules. Tarsal segmentation (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
<emphasis id="C68A1649EED395E650120C954D7FBDE3" italics="true" pageId="0" pageNumber="309">n</emphasis>
=2) 7, 16-17 (17), 10, 11-12 (12).
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
<emphasis id="D6D696FC741F90B58120BDE9CAF680EE" bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">14C, D</figureCitation>
<figureCitation id="5667A5203B17A117A8B11481BDB656FF" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">14C, D</figureCitation>
) VP rectangular; distal margin slightly straight; straight in lateral view, with apex slightly inclined. MS C1-C4(C5) subapical long and straight (some with curved apex); MS A1-A2(A3) median long and straight, shorter than MS C. Lateral sacs long, with acuminate apex; with long T3-like microsetae. Stylus with apical region cylindrical, not swollen; with many apical small projections. Dorsal process conical, with acuminate apex, smaller than stylus. Promontory convex. -
<emphasis bold="true" pageId="0" pageNumber="309">
<emphasis id="EC1775329E3E866272E1FA9631F74664" bold="true" pageId="0" pageNumber="309">
FEMALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
<emphasis id="24DCEE49A41B2C74EFBAADE1CDA6C4C3" italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
<emphasis id="8ABE60E489D2CD89BF8E856349DD45B7" italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) DSW: 5.2-5.5; DSL: 4.5-4.7; CL: 1.7-1.8. FIVL: 11.8-12.4. ChL: 2.4-2.5. Chelicerae not swollen. Femur IV unarmed. Tarsal segmentation: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
<emphasis id="9651E8DF5E191EE9EA5A5EC04676C93F" italics="true" pageId="0" pageNumber="309">n</emphasis>
=3) 7, 13, 10, 11.
</paragraph>
<caption doi="10.3897/asp.80.e73829.figure26" httpUri="https://binary.pensoft.net/fig/729900" pageId="0" pageNumber="309" start="Figure 26" startId="F26">
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">Figure 26.</emphasis>
<caption id="17830B92F50F6250AEBF3C6D734035EF" doi="10.3897/asp.80.e73829.figure26" httpUri="https://binary.pensoft.net/fig/729900" pageId="0" pageNumber="309" start="Figure 26" startId="F26">
<paragraph id="67B105A17D18F9346E62759E217770DD" pageId="0" pageNumber="309">
<emphasis id="8B3A803EDF9747B3C8E7CB26D01317FE" bold="true" pageId="0" pageNumber="309">Figure 26.</emphasis>
Live specimens of
<taxonomicName authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="309">Metasarcus</emphasis>
<taxonomicName id="031D81B668537457E520B71E0358F447" authorityName="Roewer" authorityYear="1913" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="094B8AA98EBBA9E7FD61D49036BB7012" italics="true" pageId="0" pageNumber="309">Metasarcus</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="309">A</emphasis>
<taxonomicName class="Arachnida" genus="M." kingdom="Animalia" lsidName="M. limachii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="limachii">
<emphasis italics="true" pageId="0" pageNumber="309">M. limachii</emphasis>
<emphasis id="FCA8A9C6B54EC0D3983C3F5A91248C52" bold="true" pageId="0" pageNumber="309">A</emphasis>
<taxonomicName id="D221F445F8715904BDE1DCD58227B524" class="Arachnida" kingdom="Animalia" lsidName="M. limachii" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="limachii">
<emphasis id="B719E0D79C904EEAFC66E30221498B47" italics="true" pageId="0" pageNumber="309">M. limachii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
<emphasis id="AE99CC31B605B87E09960B01BC579042" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male;
<emphasis bold="true" pageId="0" pageNumber="309">B</emphasis>
<emphasis id="7A96C7F31837C6149360970222F02C07" bold="true" pageId="0" pageNumber="309">B</emphasis>
female;
<emphasis bold="true" pageId="0" pageNumber="309">C</emphasis>
<taxonomicName authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus trispinosus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis italics="true" pageId="0" pageNumber="309">Metasarcus trispinosus</emphasis>
<emphasis id="30D644AAC19893DB74A3F51BADC29A98" bold="true" pageId="0" pageNumber="309">C</emphasis>
<taxonomicName id="819A4FBFB8CFA7FF6E28E4498E9BD341" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus trispinosus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="trispinosus">
<emphasis id="2E4D283CA1024CDA006DA5B03F275A7F" italics="true" pageId="0" pageNumber="309">Metasarcus trispinosus</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
<emphasis id="1B23FC70E2825294F48975C0A61378C2" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male;
<emphasis bold="true" pageId="0" pageNumber="309">D</emphasis>
<emphasis id="B5B7799DFF52A7BFFED48150E952F99D" bold="true" pageId="0" pageNumber="309">D</emphasis>
female;
<emphasis bold="true" pageId="0" pageNumber="309">E</emphasis>
<taxonomicName class="Arachnida" genus="M." kingdom="Animalia" lsidName="M. vacafloresae" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="vacafloresae">
<emphasis italics="true" pageId="0" pageNumber="309">M. vacafloresae</emphasis>
<emphasis id="8D518D969FFD263C04A2026529D5C885" bold="true" pageId="0" pageNumber="309">E</emphasis>
<taxonomicName id="565CDB7D3C5085AE7894B07FBE6552E9" class="Arachnida" kingdom="Animalia" lsidName="M. vacafloresae" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="vacafloresae">
<emphasis id="4CB5073B6F20E4A695ECCF5CF4D40DED" italics="true" pageId="0" pageNumber="309">M. vacafloresae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
<emphasis id="2F2237D9963DC2C46FAEF812FB39F070" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
, male;
<emphasis bold="true" pageId="0" pageNumber="309">F</emphasis>
<emphasis id="C31F786D0537F36906291C2775F95DEC" bold="true" pageId="0" pageNumber="309">F</emphasis>
female.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="diagnosis">
<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="309">
<subSubSection id="72D1DC036C07A9DBEEB8B4E968F147E0" pageId="0" pageNumber="309" type="diagnosis">
<paragraph id="32CFAF4542EDDDC10E6B87A31A45ADD8" pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph id="4225A2C7D00AA8F1572B1465746C698C" pageId="0" pageNumber="309">
Similar to
<taxonomicName authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus kurosawai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kurosawai">
<emphasis italics="true" pageId="0" pageNumber="309">Metasarcus kurosawai</emphasis>
<taxonomicName id="27AE0584C5310ED2FF34A7AB25565252" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Metasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metasarcus kurosawai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kurosawai">
<emphasis id="F70208447B1F56C27B7A85AF7AD9DB7D" italics="true" pageId="0" pageNumber="309">Metasarcus kurosawai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
<emphasis id="65A02E96A935D450268FA4821A013208" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
because ocularium unarmed, area I divided, area III with a pair of long spines (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
<figureCitation id="2A1DBFEBCDD83C884A408571A83A68B8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
) and femur IV of males slender and without rows of tubercles (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10A, B</figureCitation>
<figureCitation id="5282F2DC1AC0102BFC051197F8E2898F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10A, B</figureCitation>
). It differs from
<taxonomicName class="Arachnida" genus="M." kingdom="Animalia" lsidName="M. kurosawai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kurosawai">
<emphasis italics="true" pageId="0" pageNumber="309">M. kurosawai</emphasis>
<taxonomicName id="61329B17AB174F2AF57CBF304B316DB3" class="Arachnida" kingdom="Animalia" lsidName="M. kurosawai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="kurosawai">
<emphasis id="58A067CBB82D1B027A947721273409E6" italics="true" pageId="0" pageNumber="309">M. kurosawai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
<emphasis id="16909D277F7BAB1C3DEDE2CDEFC1C8B1" bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
by gamma-P-type DSS; coxa III unarmed (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
<figureCitation id="702DD82DCCFED6C64B9D912269AE6A65" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus, dorsal of Metasarcus. A M. beni sp. nov., male; B M. bergmani sp. nov., male; C M. bolivianus Roewer, 1913, female; D M. clavifemur (Roewer, 1929), male; E M. fellinii sp. nov., male; F M. kurosawai sp. nov., male; G M. limachii sp. nov., male; H M. trispinosus sp. nov., male; I M. vacafloresae sp. nov., male; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure5" httpUri="https://binary.pensoft.net/fig/729879" pageId="0" pageNumber="309">5A</figureCitation>
); male femur IV with a distal retrodorsal small apophysis (Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10A, B</figureCitation>
<figureCitation id="5B1CFF8B24015D4AA10622FEC733D3FE" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Male trochanter-patella / tibia IV of Metasarcus, dorsal and ventral view, respectively. A-B M. beni sp. nov.; C - D M. bergmani sp. nov.; E - F M. clavifemur (Roewer, 1929); G - H M. fellinii sp. nov.; I - J M. kurosawai sp. nov.; K - L M. limachii sp. nov.; M - N M. trispinosus sp. nov.; O-P M. vacafloresae sp. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure10" httpUri="https://binary.pensoft.net/fig/729884" pageId="0" pageNumber="309">10A, B</figureCitation>
) and penis with dorsal process and long lateral sacs (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">14C, D</figureCitation>
<figureCitation id="370D99B0DCE337D3F5C15DD6A5F9D2EC" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">14C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="etymology">
<paragraph pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
<paragraph pageId="0" pageNumber="309">The specific epithet, a noun in apposition, in reference to Beni Department (&quot;El Beni&quot;), Bolivia, Department of the type locality of the species.</paragraph>
<subSubSection id="77BA8104A23C7BF7C2CB007E7922BDAC" pageId="0" pageNumber="309" type="etymology">
<paragraph id="77DBCA8308E5ADDEFBCFF1441A1526C4" pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
<paragraph id="8C000D252891DDE9632070BF46945C8F" pageId="0" pageNumber="309">The specific epithet, a noun in apposition, in reference to Beni Department (&quot;El Beni&quot;), Bolivia, Department of the type locality of the species.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="distribution">
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="309">
<subSubSection id="D2220EEDEE54C01BA8CD913AB2406427" pageId="0" pageNumber="309" type="distribution">
<paragraph id="3C0CE4A3E5C6CD7D5A967EFA48BFE3E3" pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph id="3BF2619A60B86A4DE7A9B9C3DD1BC03A" pageId="0" pageNumber="309">
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">31</figureCitation>
<figureCitation id="4BD54D0A388CFDAFA0803C5BCA2003FA" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Distribution of Metasarcus in Bolivia. A M. beni sp. nov., M. bergmani sp. nov., M. fellinii sp. nov., M. limachii sp. nov., M. vacafloresae; B M. clavifemur, M. kurosawai sp. nov., M. trispinosus sp. nov." figureDoi="10.3897/asp.80.e73829.figure31" httpUri="https://binary.pensoft.net/fig/729905" pageId="0" pageNumber="309">31</figureCitation>
) BOLIVIA. Beni. Southwest of Yucumo.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
<emphasis italics="true" pageId="0" pageNumber="309">Type material</emphasis>
<subSubSection id="418432D0DFB9E223A835B2F3F6BB0B2B" pageId="0" pageNumber="309" type="materials_examined">
<paragraph id="025FCDAF1D04180B42214DAE48BCCE0B" pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph id="B332F95E9578ADF7C3BA70B6CF6297F3" pageId="0" pageNumber="309">
<emphasis id="F1B9D2EB3D30E481B7E27F8FD69E6C25" bold="true" pageId="0" pageNumber="309">
<emphasis id="FEBF2E5EE2612DF66E2545A1A2FBD8BA" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
:
<materialsCitation collectingDate="1989-11-15" collectingDateMax="1989-11-19" collectingDateMin="1989-11-15" collectorName="Coddington, Griswold, Silva, Larcher, Penaranda" country="BOLIVIA" latitude="-15.383333" location="Yucumo" longLatPrecision="1285" longitude="-66.98333" specimenCount="♂" stateProvince="Beni" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
<specimenCount>♂</specimenCount>
<materialsCitation id="FB9602E516BB3DA32937AEAC5A36EC7F" ID-GBIF-Occurrence="3889641322" collectingDate="1989-11-15" collectingDateMax="1989-11-19" collectingDateMin="1989-11-15" collectorName="Coddington, Griswold, Silva, Larcher, Penaranda" country="BOLIVIA" latitude="-15.383333" location="Yucumo" longLatPrecision="1285" longitude="-66.98333" specimenCount="♂" stateProvince="Beni" typeStatus="Holotype">
<typeStatus id="9BCEC2F94D9A54FD236A13F427B7BC84">Holotype</typeStatus>
<specimenCount id="F1FC79542CA999157ECE0A0F793E2967">♂</specimenCount>
, '
<collectingCountry name="Bolivia">BOLIVIA</collectingCountry>
<collectingCountry id="CF096F3EB9E2FCCD434F57BC26844F8C" name="Bolivia">BOLIVIA</collectingCountry>
,
<collectingRegion country="Bolivia" name="Beni">Beni</collectingRegion>
<collectingRegion id="06C72562D4F5471BE554F948C4C0DD4D" country="Bolivia" name="Beni">Beni</collectingRegion>
, southwest of
<location LSID="urn:lsid:plazi:treatment:CC91D7ABA56E59DEA00AF50EDEA0A505:A7C6E507EC84A820A797BE62707F5BFC" country="BOLIVIA" latitude="-15.383333" longLatPrecision="1285" longitude="-66.98333" name="Yucumo" stateProvince="Beni">Yucumo</location>
<location id="A7C6E507EC84A820A797BE62707F5BFC" LSID="urn:lsid:plazi:treatment:CC91D7ABA56E59DEA00AF50EDEA0A505:A7C6E507EC84A820A797BE62707F5BFC" country="BOLIVIA" latitude="-15.383333" longLatPrecision="1285" longitude="-66.98333" name="Yucumo" stateProvince="Beni">Yucumo</location>
|
<geoCoordinate degrees="15" direction="south" minutes="23" orientation="latitude" precision="925" value="-15.383333">15°23′S</geoCoordinate>
<geoCoordinate degrees="66" direction="west" minutes="59" orientation="longitude" precision="925" value="-66.98333">66°59′W</geoCoordinate>
<geoCoordinate id="B673168954C15A3DB8FD72890F9E47F0" degrees="15" direction="south" minutes="23" orientation="latitude" precision="925" value="-15.383333">15°23′S</geoCoordinate>
<geoCoordinate id="B77428B72962F37D0AA64ECF46E6786B" degrees="66" direction="west" minutes="59" orientation="longitude" precision="925" value="-66.98333">66°59′W</geoCoordinate>
|
<collectingDate value="1989-11-15" valueMax="1989-11-19" valueMin="1989-11-15">15-19/XI/1989</collectingDate>
<collectingDate id="07180B706AED9EE08B9B060E066366F0" value="1989-11-15" valueMax="1989-11-19" valueMin="1989-11-15">15-19/XI/1989</collectingDate>
,
<collectorName>Coddington</collectorName>
<collectorName id="A74AF66AB25F4F04CD0AB9D71B6D5704">Coddington</collectorName>
,
<collectorName>Griswold</collectorName>
<collectorName id="12D1532B2F0B4BE0E3D5C62AAE4572E4">Griswold</collectorName>
,
<collectorName>Silva</collectorName>
<collectorName id="A3DD949D3F81EF2DD6A5233087C2C5F1">Silva</collectorName>
,
<collectorName>Larcher</collectorName>
<collectorName id="F6FEC59643BEFA9F91714948B50D6143">Larcher</collectorName>
&amp;
<collectorName>Penaranda</collectorName>
<collectorName id="971E0FC1E85249E287B63CE2857F3AAA">Penaranda</collectorName>
leg. (USNM)
</materialsCitation>
</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratypes</typeStatus>
<emphasis id="B0FAECB1BB6CF8A780BE31D1163A20E4" bold="true" pageId="0" pageNumber="309">
<typeStatus id="50580918774DA80868D08C006FA13E63">Paratypes</typeStatus>
</emphasis>
<specimenCount type="male">1 ♂</specimenCount>
<specimenCount id="CE9CB0215977413862253A2F4F6E3B21" type="male">1 ♂</specimenCount>
,
<specimenCount type="female">3 ♀</specimenCount>
<specimenCount id="9D948B1508ED18441A472DECA0A67020" type="female">3 ♀</specimenCount>
,
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
<normalizedToken id="0E8D34888A09998C6F5007F2C50D9C12" originalValue="‘ditto’">'ditto'</normalizedToken>
(USNM).
</paragraph>
</subSubSection>