Add updates up until 2024-08-15 12:42:58

This commit is contained in:
ggserver 2024-08-15 12:49:04 +00:00
parent dbfd872fd0
commit f7f1009883
4 changed files with 2511 additions and 0 deletions

View file

@ -0,0 +1,613 @@
<document id="C1C19417F5B9CC28B5BFC2C4B0662D96" ID-DOI="10.1371/journal.pone.0209792" ID-ISSN="1932-6203" ID-PMC="PMC6334993" ID-PubMed="30650102" ID-Zenodo-Dep="12630921" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1719975332143" checkinUser="felipe" docAuthor="Barboza, Gloria E., García, Carolina Carrizo, González, Segundo Leiva, Scaldaferro, Marisel &amp; Reyes, Ximena" docDate="2019" docId="03F887D6FFAAFFEA305EFD896314F9D8" docLanguage="en" docName="PLoSONE.14.1.e0209792.pdf" docOrigin="PLoS ONE (e 0209792) 14 (1)" docSource="http://dx.doi.org/10.1371/journal.pone.0209792" docStyle="DocumentStyle:55318D21378FF0528C342C9740A1E1BE.2:PLoSONE.2014-.journal_article" docStyleId="55318D21378FF0528C342C9740A1E1BE" docStyleName="PLoSONE.2014-.journal_article" docStyleVersion="2" docTitle="Capsicum neei Barboza &amp; X. Reyes 2019, sp. nov." docType="treatment" docVersion="1" lastPageNumber="23" masterDocId="FFC1FFAEFFB8FFFC326BFF8E615BFFE8" masterDocTitle="Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus" masterLastPageNumber="26" masterPageNumber="1" pageNumber="19" updateTime="1723725765965" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
<mods:mods id="49597F65D6EADEEEF3A6A0EF742013D9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5DA717CF86DDED8B284055362AE3D0D0">
<mods:title id="63B4EEEB315FF20C275495C6E816DD64">Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus</mods:title>
</mods:titleInfo>
<mods:name id="2335F1920BA174F479ABAC1705202029" type="personal">
<mods:role id="E464C6629A163105AE3EE3AD8CEEB3F2">
<mods:roleTerm id="810B2C70790B775A6212B6AA8DB57FB7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83156E73FAD8DD6924C37F9CA880E18F">Barboza, Gloria E.</mods:namePart>
</mods:name>
<mods:name id="9C3D7702A1E290D964F43437DD5B64BE" type="personal">
<mods:role id="DBC052AE57CE8739FD21B0FDE21AB922">
<mods:roleTerm id="081AF515468CA21CCE8569EC490BE872">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6C3AE6376336CA191520E8BC5C1E1423">García, Carolina Carrizo</mods:namePart>
<mods:affiliation id="38C015A00DD6F5A8C4FF5D4ED84DDCE3">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Department of Botany and Biodiversity Research, University of Vienna, Vienna, Austria,</mods:affiliation>
</mods:name>
<mods:name id="5316713DA48B0AF54E54A94CA44BF2FC" type="personal">
<mods:role id="0D9005F08BB98E0A9B0EBC2034C33512">
<mods:roleTerm id="CCDC602BF81F16FCF5EEC8B98FBC310C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="10699B2ABB109DEAD04EDAD7C7DED120">González, Segundo Leiva</mods:namePart>
</mods:name>
<mods:name id="AC950DA56C29C509092395BD57A37D4A" type="personal">
<mods:role id="46C0CB008F3C6B046CDB7293FBD0FFA9">
<mods:roleTerm id="4CA6404BC04BFA6AA32CA8D58BA8088F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B1870CB582C81CFFA4F80791D4AF453D">Scaldaferro, Marisel</mods:namePart>
<mods:affiliation id="B509D5310C87B40F21D26A5CAF0D9A89">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Córdoba, Argentina,</mods:affiliation>
</mods:name>
<mods:name id="D8EA3B6347F73932F1CB213056C93B05" type="personal">
<mods:role id="F3CF503154ACF518CD9496AF957AFE48">
<mods:roleTerm id="00E0D2800048CC430C25BC6F6F2C9582">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E4FA4AEF0D36BBCD125E494DD483F775">Reyes, Ximena</mods:namePart>
<mods:affiliation id="E3939DAC9FF182AC766BA1F2536CA478">Centro de Investigaciones Fitoecogenéticas de Pairumani, Cochabamba, Bolivia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E6B3FBA7D69E5DF595D457FEDEE1312E">text</mods:typeOfResource>
<mods:relatedItem id="0E9234C3624D312C8285888D8D8EDCFB" type="host">
<mods:titleInfo id="36C24CC5F425ED95B83407019E5BD781">
<mods:title id="3FE5DD2F743D8E1200EF2D5129303E99">PLoS ONE</mods:title>
</mods:titleInfo>
<mods:part id="F66F3E36430B870F4CC65F3AA16BCC92">
<mods:date id="5FE8A984397C1C7205CECA1EB6E7A4B8">2019</mods:date>
<mods:detail id="11A68985B744A9A896EAAF67318EAA5F" type="series">
<mods:title id="F503F059D1E98A39712B93B81AC85503">e 0209792</mods:title>
</mods:detail>
<mods:detail id="867AC1B9A296FB73D8AC0154E74F0228" type="pubDate">
<mods:number id="2E347369C3C4022B273228625B2F5C10">2019-01-16</mods:number>
</mods:detail>
<mods:detail id="C9F567F94B341999FFFC8C1C91038A81" type="volume">
<mods:number id="E168CFFCEFE132C3258F17F9E4B2891A">14</mods:number>
</mods:detail>
<mods:detail id="473FADF196C856113F696C6381392D01" type="issue">
<mods:number id="1AAE118834CB0AF2E2DBEE96F84C19E5">1</mods:number>
</mods:detail>
<mods:extent id="361DDEFCCEF283AE9F832BA099DB2499" unit="page">
<mods:start id="193C7FA565DE6DC9A51F6AB838BD67CA">1</mods:start>
<mods:end id="45B8A914C439319DD5BCDF5DA90F9D90">26</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="D747003A7B575E04BC67C7E78C1A26E0">
<mods:url id="D8C8BE85D479F740EDF1E987ECB9881D">http://dx.doi.org/10.1371/journal.pone.0209792</mods:url>
</mods:location>
<mods:classification id="79B1A6F020506278CE6F46E5C696084D">journal article</mods:classification>
<mods:identifier id="D006896199B7B1719378A2C0D1C38EAE" type="DOI">10.1371/journal.pone.0209792</mods:identifier>
<mods:identifier id="44894AED03BB2467A05D5C60D379D0BA" type="ISSN">1932-6203</mods:identifier>
<mods:identifier id="1FF6E9BF4675D22306CA77AB440C60CA" type="PMC">PMC6334993</mods:identifier>
<mods:identifier id="F15F3BD1914688B60B992F27515133EB" type="PubMed">30650102</mods:identifier>
<mods:identifier id="AF0B37B3F47323700CC0424C2415C39B" type="Zenodo-Dep">12630921</mods:identifier>
</mods:mods>
<treatment id="03F887D6FFAAFFEA305EFD896314F9D8" LSID="urn:lsid:plazi:treatment:03F887D6FFAAFFEA305EFD896314F9D8" httpUri="http://treatment.plazi.org/id/03F887D6FFAAFFEA305EFD896314F9D8" lastPageId="22" lastPageNumber="23" pageId="18" pageNumber="19">
<subSubSection id="C34B654BFFAAFFEE305EFD89654FFDC8" box="[565,1044,519,544]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BEE36C0FFAAFFEE305EFD89654FFDC8" blockId="18.[533,1536,207,1896]" box="[565,1044,519,544]" pageId="18" pageNumber="19">
<emphasis id="B925EAD2FFAAFFEE305EFD89654FFDC8" bold="true" box="[565,1044,519,544]" pageId="18" pageNumber="19">
<taxonomicName id="4C514D43FFAAFFEE305EFD8962EFFDC8" ID-CoL="7TBQQ" authority="Barboza &amp; X. Reyes, 2019" authorityName="Barboza &amp; X. Reyes" authorityYear="2019" box="[565,948,519,544]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="neei" status="sp. nov.">
<emphasis id="B925EAD2FFAAFFEE305EFD896388FDF7" bold="true" box="[565,723,519,543]" italics="true" pageId="18" pageNumber="19">Capsicum neei</emphasis>
Barboza &amp; X. Reyes
</taxonomicName>
,
<taxonomicNameLabel id="A21657A9FFAAFFEE31D4FD86654FFDC8" box="[959,1044,520,544]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFAAFFEE3671FD866396FD25" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BEE36C0FFAAFFEE3671FD866311FD42" blockId="18.[533,1536,207,1896]" pageId="18" pageNumber="19">
[
<materialsCitation id="3B393C9DFFAAFFEE364BFD866251FD60" collectingDate="2017-12-16" collectorName="de Monteagudo &amp; G. E. Barboza" country="Bolivia" latitude="-19.804617" location="Monteagudo" longLatPrecision="1" longitude="-64.01992" municipality="Type" pageId="18" pageNumber="19" specimenCount="2" stateProvince="Chuquisaca" typeStatus="holotype">
urn:lsid:ipni.org:names: 771925601].
<collectingMunicipality id="6B8AACBAFFAAFFEE37D0FD8664A8FDC8" box="[1467,1523,520,544]" pageId="18" pageNumber="19">Type</collectingMunicipality>
:
<collectingCountry id="F3467650FFAAFFEE307EFDA4633AFDAA" box="[533,609,554,578]" name="Bolivia" pageId="18" pageNumber="19">Bolivia</collectingCountry>
.
<collectingRegion id="4995F822FFAAFFEE3000FDA463B1FDAA" box="[619,746,554,578]" country="Bolivia" name="Chuquisaca" pageId="18" pageNumber="19">Chuquisaca</collectingRegion>
: Prov. Hernando Siles, a
<quantity id="4CA99B25FFAAFFEE3199FDA46561FDAA" box="[1010,1082,554,578]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="18" pageNumber="19" unit="km" value="4.1">4.1 km</quantity>
del puente nuevo
<collectorName id="26A45316FFAAFFEE3696FDA464FEFDAA" box="[1277,1445,554,578]" pageId="18" pageNumber="19">de Monteagudo</collectorName>
viniendo desde
<location id="8E8E601BFFAAFFEE30D7FDC3621CFD8D" LSID="urn:lsid:plazi:treatment:03F887D6FFAAFFEA305EFD896314F9D8:8E8E601BFFAAFFEE30D7FDC3621CFD8D" box="[700,839,589,613]" country="Bolivia" latitude="-19.804617" longLatPrecision="1" longitude="-64.01992" municipality="Type" name="Monteagudo" pageId="18" pageNumber="19" stateProvince="Chuquisaca">Monteagudo</location>
, sobre mano derecha, -
<geoCoordinate id="EE655007FFAAFFEE3651FDC365E7FD8D" box="[1082,1212,589,613]" degrees="19.804617" direction="south" orientation="latitude" pageId="18" pageNumber="19" precision="1" value="-19.804617">19.804617 S</geoCoordinate>
, -
<geoCoordinate id="EE655007FFAAFFEE36A5FDC36438FD8D" box="[1230,1379,589,613]" degrees="64, 019923" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="1" value="-64.01992">64, 019923 W</geoCoordinate>
,
<date id="FFEF1000FFAAFFEE3706FDC364B4FD8D" box="[1389,1519,589,613]" pageId="18" pageNumber="19" value="2017-12-16">
<collectingDate id="EFABE9E8FFAAFFEE3706FDC364B4FD8D" box="[1389,1519,589,613]" pageId="18" pageNumber="19" value="2017-12-16">16 Dec 2017</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFAAFFEE3029FDFE638AFD6F" box="[578,721,623,648]" pageId="18" pageNumber="19">
<emphasis id="B925EAD2FFAAFFEE3029FDFE630FFD6F" box="[578,596,624,647]" italics="true" pageId="18" pageNumber="19">G</emphasis>
.
<emphasis id="B925EAD2FFAAFFEE300BFDFE6334FD6F" box="[608,623,624,647]" italics="true" pageId="18" pageNumber="19">E</emphasis>
.
<emphasis id="B925EAD2FFAAFFEE3011FDE16251FD60" box="[634,778,623,648]" italics="true" pageId="18" pageNumber="19">Barboza 4927</emphasis>
</collectorName>
</materialsCitation>
<materialsCitation id="3B393C9DFFAAFFEE317BFDFE62EFFD60" box="[784,948,624,648]" collectingDate="2017-12-16" collectorName="de Monteagudo &amp; G. E. Barboza" country="Bolivia" latitude="-19.804617" location="Monteagudo" longLatPrecision="1" longitude="-64.01992" municipality="Type" pageId="18" pageNumber="19" specimenCount="2" stateProvince="Chuquisaca" typeStatus="holotype">
(
<typeStatus id="54EA8862FFAAFFEE317CFDFE6223FD60" box="[791,888,624,648]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
, LPB;
</materialsCitation>
<materialsCitation id="3B393C9DFFAAFFEE31D1FDFE631EFD42" collectingDate="2017-12-16" collectorName="de Monteagudo &amp; G. E. Barboza" country="Bolivia" latitude="-19.804617" location="Monteagudo" longLatPrecision="1" longitude="-64.01992" municipality="Type" pageId="18" pageNumber="19" specimenCount="2" stateProvince="Chuquisaca" typeStatus="holotype">
<typeStatus id="54EA8862FFAAFFEE31D1FDFE654AFD60" box="[954,1041,624,648]" pageId="18" pageNumber="19" type="isotype">isotypes</typeStatus>
, CORD [CORD-00006935, CORD-00006956], NY)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36C0FFAAFFEE305EFD3B6396FD25" blockId="18.[533,1536,207,1896]" box="[565,717,693,717]" pageId="18" pageNumber="19">
<figureCitation id="136A2A45FFAAFFEE305EFD3B6324FD25" box="[565,639,693,717]" captionStart="Fig 10" captionStartId="19.[388,416,1825,1844]" captionTargetBox="[388,1533,208,1809]" captionTargetId="figure-1@19.[388,1536,208,1810]" captionTargetPageId="19" captionText="Fig 10. Capsicum neei Barboza &amp; X. Reyes. (A) Flowering branch. (B) Inflorescence. (C) Flower bud. (D) Flower. (E) Calyx. (F) Opened corolla.(G) Gynoecium (H) Fruit. (I) Glandular trichome of the inside calyx. (J) Glandular trichome of the pedicels.(K)" figureDoi="http://doi.org/10.5281/zenodo.12630935" httpUri="https://zenodo.org/record/12630935/files/figure.png" pageId="18" pageNumber="19">Figs 10</figureCitation>
and
<figureCitation id="136A2A45FFAAFFEE30DFFD3B6396FD25" box="[692,717,693,717]" captionStart="Fig 11" captionStartId="20.[533,561,1818,1837]" captionTargetBox="[533,1386,517,1797]" captionTargetId="figure-104@20.[533,1386,517,1797]" captionTargetPageId="20" captionText="Fig 11. Capsicum neei Barboza &amp; X. Reyes. (A) Plant. (B) Flower bud. (C) Flowering branch. Photos by G. E. Barboza. https://doi.org/10.1371/journal.pone.0209792.g011" figureDoi="http://doi.org/10.5281/zenodo.12630937" httpUri="https://zenodo.org/record/12630937/files/figure.png" pageId="18" pageNumber="19">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFAAFFEE305EFD5664F2FCFA" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BEE36C0FFAAFFEE305EFD5664F2FCFA" blockId="18.[533,1536,207,1896]" pageId="18" pageNumber="19">
<emphasis id="B925EAD2FFAAFFEE305EFD5663F8FD18" bold="true" box="[565,675,728,752]" pageId="18" pageNumber="19">Diagnosis</emphasis>
. Like
<taxonomicName id="4C514D43FFAAFFEE3088FD5665DAFD18" authority="Rusby (Hunz.)" box="[739,1153,727,752]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="minutiflorum">
<emphasis id="B925EAD2FFAAFFEE3088FD566284FD07" box="[739,991,727,751]" italics="true" pageId="18" pageNumber="19">Capsicum minutiflorum</emphasis>
Rusby (Hunz.)
</taxonomicName>
but differing in the non-geniculate pendent flowering pedicels and the strongly nerved calyx with 10 unequal appendages.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFAAFFE8305EFC936240FECD" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
<paragraph id="8BEE36C0FFAAFFEE305EFC93642EF880" blockId="18.[533,1536,207,1896]" pageId="18" pageNumber="19">
<emphasis id="B925EAD2FFAAFFEE305EFC9363E3FCDD" bold="true" box="[565,696,797,821]" pageId="18" pageNumber="19">Description</emphasis>
. Small shrubs 0.702 (3) m tall, thin, erect, laxly branched above. Young stems green, slim, fragile, glabrescent, and slightly striate, without lenticels; bark of older stems light brown, glabrous, with a few oblong lenticels. Sympodial units difoliate, geminate, leaf pair not markedly anisophyllous in size and shape. Leaves simple, membranaceous, glabrescent on both surfaces and margins with 4-7-celled non glandular trichomes
<quantity id="4CA99B25FFAAFFEE3688FC26643AFC28" box="[1251,1377,936,960]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="18" pageNumber="19" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.20.5 mm</quantity>
long; the larger leaves with blades (5.5)
<quantity id="4CA99B25FFAAFFEE3139FC44629BFC0A" box="[850,960,970,994]" metricMagnitude="-2" metricUnit="m" metricValue="8.85" metricValueMax="11.0" metricValueMin="6.7" pageId="18" pageNumber="19" unit="cm" value="8.85" valueMax="11.0" valueMin="6.7">6.711 cm</quantity>
long, 2.14 (4.5) cm wide, elliptic or ovate, major veins 34 on each side of midvein, base attenuate, margin entire, apex acute; petioles 0.30.8 (1.5) cm long; the minor leaves 2.74.6 (6) cm long, 1.21.8 (2.3) cm wide, elliptic or ovate, major veins 23 on each side of midvein, base attenuate, margin entire, apex obtuse or acute; petioles 0.20.5 (0.8) cm long, with similar pubescence as in larger leaves. Flowers 24 per axil, rarely solitary; flowering pedicels green, filiform, striate, pendent, slightly curved, not geniculate at anthesis, (0.65)
<quantity id="4CA99B25FFAAFFEE316BFB146228FB5A" box="[768,883,1178,1202]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="18" pageNumber="19" unit="cm" value="1.15" valueMax="1.5" valueMin="0.8">0.81.5 cm</quantity>
long, with sparse 5-6-celled non-glandular trichomes and tiny dark glandular trichomes (stalk unicellular, head multicellular). Flower buds ovoid, greenish pale yellow. Calyx
<quantity id="4CA99B25FFAAFFEE3095FB6E6227FB10" box="[766,892,1248,1272]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.5" metricValueMin="1.7" pageId="18" pageNumber="19" unit="mm" value="2.1" valueMax="2.5" valueMin="1.7">1.72.5 mm</quantity>
long,
<quantity id="4CA99B25FFAAFFEE31D6FB6E654DFB10" box="[957,1046,1248,1272]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="18" pageNumber="19" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
wide, cup-shaped, green, with 10 nerves clearly evident, the margin truncate, pubescent, with non-glandular trichomes
<quantity id="4CA99B25FFAAFFEE373CFA8C648EFAF2" box="[1367,1493,1282,1306]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="18" pageNumber="19" unit="mm" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">0.30.6 mm</quantity>
long outside and dense glandular pubescence inside (head multicellular, stalk unicellular), 10 unequal linear appendages, green, the five longer appendages (0.7) 0.91.75 (2) mm long, emerging almost from the margin, the five shorter 0.20.8 (1.2) mm long, emerging
<quantity id="4CA99B25FFAAFFEE37E4FAE464A0FA6A" box="[1423,1531,1386,1410]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="18" pageNumber="19" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.81 mm</quantity>
below the margin, with the same non glandular trichomes of the calyx tube. Corolla (6)
<quantity id="4CA99B25FFAAFFEE37D8FA03631AFA20" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="18" pageNumber="19" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
long,
<quantity id="4CA99B25FFAAFFEE30E9FA3E6380FA20" box="[642,731,1456,1480]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="18" pageNumber="19" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
diam, stellate, delicate, entirely yellow or with small brownish green spots in the base of the lobes and tube inside, with a thin interpetalar tissue; tube
<quantity id="4CA99B25FFAAFFEE3744FA5C64C0FA02" box="[1327,1435,1490,1514]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="18" pageNumber="19" unit="mm" value="3.75" valueMax="4.5" valueMin="3.0">34.5 mm</quantity>
long, with tiny glandular trichomes (head and stalk one celled each) inside and glabrescent outside; lobes
<quantity id="4CA99B25FFAAFFEE3039F996638BF9D8" box="[594,720,1560,1584]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.5" metricValueMin="3.5" pageId="18" pageNumber="19" unit="mm" value="4.5" valueMax="5.5" valueMin="3.5">3.55.5 mm</quantity>
long, ca.
<quantity id="4CA99B25FFAAFFEE3158F9966229F9D8" box="[819,882,1560,1584]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="mm" value="2.0">2 mm</quantity>
wide, ovate, erect, glabrous adaxially and with sparse non-glandular trichomes abaxially, the tips papillose and cucullate. Stamens 5, subequal, filaments
<quantity id="4CA99B25FFAAFFEE307EF9D363FBF99D" box="[533,672,1629,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.75" metricValueMin="1.4" pageId="18" pageNumber="19" unit="mm" value="1.575" valueMax="1.75" valueMin="1.4">1.41.75 mm</quantity>
long, cream, glabrous, inserted on the corolla ca.
<quantity id="4CA99B25FFAAFFEE36C7F9D365A5F99D" box="[1196,1278,1629,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="19" unit="mm" value="1.2">1.2 mm</quantity>
from the base, with inconspicuous auricles at point of insertion; anthers (1.5)
<quantity id="4CA99B25FFAAFFEE3613F90E65BFF970" box="[1144,1252,1664,1688]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="18" pageNumber="19" unit="mm" value="1.9" valueMax="2.0" valueMin="1.8">1.82 mm</quantity>
long, not connivent, elliptic, light yellow. Ovary ca.
<quantity id="4CA99B25FFAAFFEE3147F92C6226F952" box="[812,893,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="19" unit="mm" value="1.2">1.2 mm</quantity>
long,
<quantity id="4CA99B25FFAAFFEE31D5F92C6554F952" box="[958,1039,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="19" unit="mm" value="1.3">1.3 mm</quantity>
diam, ovoid or subglobose, light green, glabrous; nectary ca.
<quantity id="4CA99B25FFAAFFEE30BEF94B627DF935" box="[725,806,1733,1757]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="mm" value="0.3">0.3 mm</quantity>
tall, style
<quantity id="4CA99B25FFAAFFEE31E5F94B62B7F935" box="[910,1004,1733,1757]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" pageId="18" pageNumber="19" unit="mm" value="3.75">3.75 mm</quantity>
long, cream, widening distally, glabrous; stigma ca.
<quantity id="4CA99B25FFAAFFEE3053F96663D2F8E8" box="[568,649,1768,1792]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="mm" value="0.2">0.2 mm</quantity>
long,
<quantity id="4CA99B25FFAAFFEE30A1F9666240F8E8" box="[714,795,1768,1792]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="19" unit="mm" value="0.3">0.3 mm</quantity>
wide, light green, somewhat bilobed. Berry
<quantity id="4CA99B25FFAAFFEE3680F9666430F8E8" box="[1259,1387,1768,1792]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="7.5" metricValueMin="4.0" pageId="18" pageNumber="19" unit="cm" value="0.575" valueMax="0.75" valueMin="0.4">0.40.75 cm</quantity>
diam, globose, green when immature, red at maturity, glabrous; fruiting pedicels (1.3)
<quantity id="4CA99B25FFAAFFEE3728F88464ECF8CA" box="[1347,1463,1802,1826]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="18" pageNumber="19" unit="cm" value="2.05" valueMax="2.3" valueMin="1.8">1.82.3 cm</quantity>
long, pendent, striate and widened distally; the fruiting calyx persistent, non-accrescent, ca.
<quantity id="4CA99B25FFAAFFEE37CFF8A364B8F8AD" box="[1444,1507,1837,1861]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="19" unit="mm" value="4.0">4 mm</quantity>
diam, discoid, the appendages spreading
<quantity id="4CA99B25FFAAFFEE31A3F8DE657AF880" box="[968,1057,1872,1896]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="18" pageNumber="19" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
long, subulate. Seeds unknown.
</paragraph>
<caption id="DF2E6648FFABFFEF33EFF8AF64E6F8A6" ID-DOI="http://doi.org/10.5281/zenodo.12630935" ID-Zenodo-Dep="12630935" httpUri="https://zenodo.org/record/12630935/files/figure.png" pageId="19" pageNumber="20" startId="19.[388,416,1825,1844]" targetBox="[388,1533,208,1809]" targetPageId="19" targetType="figure">
<paragraph id="8BEE36C0FFABFFEF33EFF8AF64E6F8A6" blockId="19.[388,1470,1825,1870]" pageId="19" pageNumber="20">
<emphasis id="B925EAD2FFABFFEF33EFF8AF63A0F8DC" bold="true" box="[388,763,1825,1844]" pageId="19" pageNumber="20">
Fig 10.
<taxonomicName id="4C514D43FFABFFEF33ADF8AF63A0F8DC" authority="Barboza &amp; X. Reyes." authorityName="Barboza &amp; X. Reyes." box="[454,763,1825,1844]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFABFFEF33ADF8AF631EF8DC" bold="true" box="[454,581,1825,1844]" italics="true" pageId="19" pageNumber="20">Capsicum neei</emphasis>
Barboza &amp; X. Reyes.
</taxonomicName>
</emphasis>
(A) Flowering branch. (B) Inflorescence. (C) Flower bud. (D) Flower. (E) Calyx. (F) Opened corolla.(G) Gynoecium (H) Fruit. (I) Glandular trichome of the inside calyx. (J) Glandular trichome of the pedicels.(K)
</paragraph>
</caption>
<paragraph id="8BEE36C0FFACFFE833EFFF5E6387FF14" blockId="20.[388,1516,208,293]" pageId="20" pageNumber="21">Glandular trichome of the inside corolla. (L) Non-glandular trichome of the outside calyx. (M) Non-glandular trichome of the outside corolla lobes. Drawn by S. Montecchiesi.</paragraph>
<paragraph id="8BEE36C0FFACFFE833EFFE9C6240FECD" blockId="20.[388,1516,208,293]" box="[388,795,274,293]" pageId="20" pageNumber="21">https://doi.org/10.1371/journal.pone.0209792.g010</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFACFFE9305EFEC065CCFABB" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BEE36C0FFACFFE9305EFEC063C9FAD9" blockId="20.[533,1510,334,462]" lastBlockId="21.[533,1536,1235,1883]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="B925EAD2FFACFFE8305EFEC0620AFE8E" bold="true" box="[565,849,334,358]" pageId="20" pageNumber="21">Distribution and ecology.</emphasis>
Endemic to southeastern
<collectingCountry id="F3467650FFACFFE8360FFEC065F6FE8E" box="[1124,1197,334,358]" name="Bolivia" pageId="20" pageNumber="21">Bolivia</collectingCountry>
(
<figureCitation id="136A2A45FFACFFE836D6FEC065A6FE8E" box="[1213,1277,334,358]" captionStart="Fig 12" captionStartId="21.[533,561,1141,1160]" captionTargetBox="[533,1320,208,1118]" captionTargetId="figure-259@21.[533,1320,208,1119]" captionTargetPageId="21" captionText="Fig 12. Distribution of Capsicum neei Barboza &amp; X. Reyes. https://doi.org/10.1371/journal.pone.0209792.g012" figureDoi="http://doi.org/10.5281/zenodo.12630939" httpUri="https://zenodo.org/record/12630939/files/figure.png" pageId="20" pageNumber="21">Fig 12</figureCitation>
), mainly in the Serranías Iñao, Yahuañanca and Khaskha Orkho (Dpt.
<collectingRegion id="4995F822FFACFFE8364CFEFF65F3FE61" box="[1063,1192,369,393]" country="Bolivia" name="Chuquisaca" pageId="20" pageNumber="21">Chuquisaca</collectingRegion>
). A few collections have been recorded from the Yungas (Dpt.
<collectingRegion id="4995F822FFACFFE8311BFE1D62B2FE43" box="[880,1001,403,427]" country="Bolivia" name="Santa Cruz" pageId="20" pageNumber="21">Santa Cruz</collectingRegion>
).
<taxonomicName id="4C514D43FFACFFE83191FE1A65C9FE43" box="[1018,1170,404,427]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFACFFE83191FE1A65C9FE43" box="[1018,1170,404,427]" italics="true" pageId="20" pageNumber="21">Capsicum neei</emphasis>
</taxonomicName>
is most commonly collected in the Boliviano-Tucumano Forest in both Departments [
<bibRefCitation id="EFC04B31FFACFFE83637FE38652EFE26" author="Navarro G &amp; Ferreira W." box="[1116,1141,438,462]" pageId="20" pageNumber="21" pagination="25 - 56" refId="ref14129" refString="35. Navarro G, Ferreira W. 2011. Clasificacion y caracterizacion de la vegetacion del Departamento de Chuquisaca. In: Carretero A, Serrano M, Borchsenius F, Balslev H, editores. Pueblos y Plantas de Chuquisaca. Estado del conocimiento de los pueblos, la flora, uso y conservacion. Sucre: Herbario del Sur de Bolivia-Universidad Mayor, Real y Pontificia de San Francisco Xavier de Chuquisaca; 2011. pp. 25 - 56." type="journal article" year="2011">35</bibRefCitation>
] from understories at the foot of cloud forest hillsides and deciduous forests, between
<quantity id="4CA99B25FFADFFE9362EFB5D658FFB03" box="[1093,1236,1235,1259]" metricMagnitude="3" metricUnit="m" metricValue="1.425" metricValueMax="1.75" metricValueMin="1.1" pageId="21" pageNumber="22" unit="m" value="1425.0" valueMax="1750.0" valueMin="1100.0">11001750 m</quantity>
elevation. It grows associated with members of
<taxonomicName id="4C514D43FFADFFE93169FB7862D4FAE6" box="[770,911,1270,1294]" class="Magnoliopsida" family="Juglandaceae" kingdom="Plantae" order="Fagales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="family">Juglandaceae</taxonomicName>
,
<taxonomicName id="4C514D43FFADFFE931F2FB78655DFAE6" box="[921,1030,1270,1294]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
,
<taxonomicName id="4C514D43FFADFFE93664FB786441FAE6" authority=", Leguminosae" authorityName="Leguminosae" box="[1039,1306,1270,1294]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="family">Myrtaceae, Leguminosae</taxonomicName>
, ferns and bryophytes.
</paragraph>
<caption id="DF2E6648FFACFFE8307EF89462F6F898" ID-DOI="http://doi.org/10.5281/zenodo.12630937" ID-Zenodo-Dep="12630937" httpUri="https://zenodo.org/record/12630937/files/figure.png" pageId="20" pageNumber="21" startId="20.[533,561,1818,1837]" targetBox="[533,1386,517,1797]" targetPageId="20" targetType="figure">
<paragraph id="8BEE36C0FFACFFE8307EF894633AF8AF" blockId="20.[533,1461,1818,1904]" pageId="20" pageNumber="21">
<emphasis id="B925EAD2FFACFFE8307EF89462D6F8C5" bold="true" box="[533,909,1818,1837]" pageId="20" pageNumber="21">
Fig 11.
<taxonomicName id="4C514D43FFACFFE8303CF89462D6F8C5" authority="Barboza &amp; X. Reyes." authorityName="Barboza &amp; X. Reyes." box="[599,909,1818,1837]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFACFFE8303CF894638DF8C5" bold="true" box="[599,726,1818,1837]" italics="true" pageId="20" pageNumber="21">Capsicum neei</emphasis>
Barboza &amp; X. Reyes.
</taxonomicName>
</emphasis>
(A) Plant. (B) Flower bud. (C) Flowering branch. Photos by G. E. Barboza.
</paragraph>
<paragraph id="8BEE36C0FFACFFE8307EF8D362F6F898" blockId="20.[533,1461,1818,1904]" box="[533,941,1885,1904]" pageId="20" pageNumber="21">https://doi.org/10.1371/journal.pone.0209792.g011</paragraph>
</caption>
<caption id="DF2E6648FFADFFE9307EFBFB62F6FB59" ID-DOI="http://doi.org/10.5281/zenodo.12630939" ID-Zenodo-Dep="12630939" httpUri="https://zenodo.org/record/12630939/files/figure.png" pageId="21" pageNumber="22" startId="21.[533,561,1141,1160]" targetBox="[533,1320,208,1118]" targetPageId="21" targetType="figure">
<paragraph id="8BEE36C0FFADFFE9307EFBFB654CFB60" blockId="21.[533,1047,1140,1201]" box="[533,1047,1140,1160]" pageId="21" pageNumber="22">
<emphasis id="B925EAD2FFADFFE9307EFBFB654CFB60" bold="true" box="[533,1047,1140,1160]" pageId="21" pageNumber="22">
Fig 12. Distribution of
<taxonomicName id="4C514D43FFADFFE93089FBFA654CFB60" authority="Barboza &amp; X. Reyes." authorityName="Barboza &amp; X. Reyes." box="[738,1047,1140,1160]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFADFFE93089FBFA623AFB6F" bold="true" box="[738,865,1140,1159]" italics="true" pageId="21" pageNumber="22">Capsicum neei</emphasis>
Barboza &amp; X. Reyes.
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BEE36C0FFADFFE9307EFB1062F6FB59" blockId="21.[533,1047,1140,1201]" box="[533,941,1182,1201]" pageId="21" pageNumber="22">https://doi.org/10.1371/journal.pone.0209792.g012</paragraph>
</caption>
<paragraph id="8BEE36C0FFADFFE9305EFAB565CCFABB" blockId="21.[533,1536,1235,1883]" box="[565,1175,1339,1363]" pageId="21" pageNumber="22">
<emphasis id="B925EAD2FFADFFE9305EFAB563F1FABB" bold="true" box="[565,682,1339,1363]" pageId="21" pageNumber="22">Phenology</emphasis>
. Flowering and fruiting from October to May.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFADFFE9305EFAD064E7F939" pageId="21" pageNumber="22" type="etymology">
<paragraph id="8BEE36C0FFADFFE9305EFAD0651FFA53" blockId="21.[533,1536,1235,1883]" pageId="21" pageNumber="22">
<emphasis id="B925EAD2FFADFFE9305EFAD063EFFA9E" bold="true" box="[565,692,1374,1398]" pageId="21" pageNumber="22">Etymology.</emphasis>
The epithet is in honor to Dr. Mike Nee (
<collectionCode id="ED40AE05FFADFFE93600FAD065CFFA9E" box="[1131,1172,1374,1398]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="21" pageNumber="22" type="Herbarium">NY</collectionCode>
), a solanaceous specialist who carried out extensive explorations in the Bolivian territory and separated specimens of this species as a rare or probable new species in various herbaria.
</paragraph>
<paragraph id="8BEE36C0FFADFFE9305EFA4864E7F939" blockId="21.[533,1536,1235,1883]" pageId="21" pageNumber="22">
<emphasis id="B925EAD2FFADFFE9305EFA4862F5FA36" bold="true" box="[565,942,1478,1502]" pageId="21" pageNumber="22">Species Conservation Assessment.</emphasis>
According to IUCN criteria [
<bibRefCitation id="EFC04B31FFADFFE93682FA486459FA36" author="Using the IUCN Red List Categories" box="[1257,1282,1478,1502]" pageId="21" pageNumber="22" refId="ref13303" refString="15. IUCN. Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. 2017 [cited 3 May 2018]. Available from http: // www. iucnredlist. org / documents / RedListGuidelines. pdf" type="book" year="2017">15</bibRefCitation>
],
<taxonomicName id="4C514D43FFADFFE9377CFA486405FA35" box="[1303,1374,1478,1502]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFADFFE9377CFA486473FA35" box="[1303,1320,1478,1501]" italics="true" pageId="21" pageNumber="22">C</emphasis>
.
<emphasis id="B925EAD2FFADFFE9375FFA486405FA35" box="[1332,1374,1478,1501]" italics="true" pageId="21" pageNumber="22">neei</emphasis>
</taxonomicName>
is proposed as Near Threatened species. The species meets the area requirements under criterion B for threatened (EOO:
<quantity id="4CA99B25FFADFFE930F1F985625EF9CB" box="[666,773,1547,1571]" metricMagnitude="7" metricUnit="m" metricValue="1.6911999999999998" pageId="21" pageNumber="22" unit="km" value="16912.0">16912 km</quantity>
2, B1 &lt;
<quantity id="4CA99B25FFADFFE93133F9856299F9CB" box="[856,962,1547,1571]" metricMagnitude="7" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="22" unit="km" value="20000.0">20000 km</quantity>
2, Vulnerable; AOO:
<quantity id="4CA99B25FFADFFE936CBF98565BEF9CB" box="[1184,1253,1547,1571]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="21" pageNumber="22" unit="km" value="44.0">44 km</quantity>
2, B2 &lt;
<quantity id="4CA99B25FFADFFE93753F98564D2F9CB" box="[1336,1417,1547,1571]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="22" unit="km" value="500.0">500 km</quantity>
2, Endangered) and is declining, but the population is not severely fragmented and occurs in more than 10 locations.
<taxonomicName id="4C514D43FFADFFE930CBF9DF6263F980" box="[672,824,1617,1640]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFADFFE930CBF9DF6263F980" box="[672,824,1617,1640]" italics="true" pageId="21" pageNumber="22">Capsicum neei</emphasis>
</taxonomicName>
has been collected many times in the last 23 years in a recently Protected Area: National Park and Integrated Management Natural Area “Serranía Iñao” [
<bibRefCitation id="EFC04B31FFADFFE937B9F9FD64B0F963" author="Castro L." box="[1490,1515,1651,1675]" pageId="21" pageNumber="22" pagination="345 - 351" refId="ref14213" refString="36. Castro L. Gestion y conservacion in situ de la biodiversidad en Chuquisaca. In: Carretero A, Serrano M, Borchsenius F, Balslev H, editores. Pueblos y Plantas de Chuquisaca. Estado del conocimiento de los pueblos, la flora, uso y conservacion. Sucre: Herbario del Sur de Bolivia-Universidad Mayor, Real y Pontificia de San Francisco Xavier de Chuquisaca; 2011. pp. 345 - 351." type="book chapter" year="2011">36</bibRefCitation>
], and in nearby areas which suggests that both the decline in its geographic range (EOO and AOO) and the population size will not be significantly affected in the forthcoming years.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFADFFE9305EF95563B1F8FE" pageId="21" pageNumber="22" type="description">
<paragraph id="8BEE36C0FFADFFE9305EF95563B1F8FE" blockId="21.[533,1536,1235,1883]" pageId="21" pageNumber="22">
<emphasis id="B925EAD2FFADFFE9305EF95563C5F91B" bold="true" box="[565,670,1755,1779]" pageId="21" pageNumber="22">Affinities</emphasis>
.
<taxonomicName id="4C514D43FFADFFE930C1F9526219F91B" box="[682,834,1756,1779]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFADFFE930C1F9526219F91B" box="[682,834,1756,1779]" italics="true" pageId="21" pageNumber="22">Capsicum neei</emphasis>
</taxonomicName>
is nested within the Bolivian clade, strongly resolved as sister to
<taxonomicName id="4C514D43FFADFFE9307EF97063C3F8FE" box="[533,664,1790,1814]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="caballeroi">
<emphasis id="B925EAD2FFADFFE9307EF970637DF8FD" box="[533,550,1790,1813]" italics="true" pageId="21" pageNumber="22">C</emphasis>
.
<emphasis id="B925EAD2FFADFFE93059F97063C3F8FE" box="[562,664,1790,1814]" italics="true" pageId="21" pageNumber="22">caballeroi</emphasis>
</taxonomicName>
(
<figureCitation id="136A2A45FFADFFE930CCF9706380F8FE" box="[679,731,1790,1814]" captionStart="Fig 6" captionStartId="12.[133,161,1434,1453]" captionTargetBox="[134,1535,208,1411]" captionTargetId="figure-188@12.[133,1536,208,1412]" captionTargetPageId="12" captionText="Fig 6. Bayesian majority-rule consensus tree of Capsicum. Posterior probabilities values indicated by each branch. New species are highlighted in bold-colored letters and the clade to which they belong is indicated. https://doi.org/10.1371/journal.pone.0209792.g006" figureDoi="http://doi.org/10.5281/zenodo.12630927" httpUri="https://zenodo.org/record/12630927/files/figure.png" pageId="21" pageNumber="22">Fig 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFADFFEA305EF8AF6314F9D8" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BEE36C0FFADFFEA305EF8AF64D7FC75" blockId="21.[533,1536,1235,1883]" lastBlockId="22.[533,1536,207,1584]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="B925EAD2FFADFFE9305EF8AF65FAF8D1" bold="true" box="[565,1185,1825,1849]" pageId="21" pageNumber="22">
Additional specimens examined.
<materialsCitation id="3B393C9DFFADFFEA31CEF8AF6491FE5F" collectingDate="2000-10-04" collectingDateMax="2006-12-15" collectingDateMin="2000-10-04" collectionCode="HSB" country="Bolivia" county="Hernando Siles" elevation="1127" lastPageId="22" lastPageNumber="23" location="Heredia" municipality="Monteagudo" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Chuquisaca">
<collectingCountry id="F3467650FFADFFE931CEF8AF654FF8D1" box="[933,1044,1825,1849]" name="Bolivia" pageId="21" pageNumber="22">BOLIVIA</collectingCountry>
.
<collectingRegion id="4995F822FFADFFE93674F8AF65FAF8D1" box="[1055,1185,1825,1849]" country="Bolivia" name="Chuquisaca" pageId="21" pageNumber="22">Chuquisaca</collectingRegion>
:
<collectingCounty id="628F4E4CFFADFFE936C7F8AF6409F8D1" box="[1196,1362,1825,1849]" pageId="21" pageNumber="22">Hernando Siles</collectingCounty>
, ca. 7 km
<collectorName id="26A45316FFADFFE937D1F8AF63FBF8B3" pageId="21" pageNumber="22">
de
<collectingMunicipality id="6B8AACBAFFADFFE9307EF8CD63FBF8B3" box="[533,672,1859,1883]" pageId="21" pageNumber="22">Monteagudo</collectingMunicipality>
</collectorName>
, inicio del cañón
<location id="8E8E601BFFADFFE93131F8CD62E9F8B3" LSID="urn:lsid:plazi:treatment:03F887D6FFAAFFEA305EFD896314F9D8:8E8E601BFFADFFE93131F8CD62E9F8B3" box="[858,946,1859,1883]" country="Bolivia" county="Hernando Siles" municipality="Monteagudo" name="Heredia" pageId="21" pageNumber="22" stateProvince="Chuquisaca">Heredia</location>
, 19˚4717S, 64˚0208W, 1127 m,
<date id="FFEF1000FFADFFE9374FF8CD64FDF8B3" box="[1316,1446,1859,1883]" pageId="21" pageNumber="22" value="2006-12-13">
<collectingDate id="EFABE9E8FFADFFE9374FF8CD64FDF8B3" box="[1316,1446,1859,1883]" pageId="21" pageNumber="22" value="2006-12-13">13 Dec 2006</collectingDate>
</date>
(fl, fr),
<collectorName id="26A45316FFAEFFEA307EFF5E63D1FF0F" box="[533,650,207,232]" pageId="22" pageNumber="23">
<emphasis id="B925EAD2FFAEFFEA307EFF5E6372FF0F" box="[533,553,208,231]" italics="true" pageId="22" pageNumber="23">H</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA305EFF4163E0FF0F" box="[565,699,207,231]" italics="true" pageId="22" pageNumber="23">Huaylla et al</emphasis>
</collectorName>
.
<emphasis id="B925EAD2FFAEFFEA30ACFF5E63A1FF00" box="[711,762,208,232]" italics="true" pageId="22" pageNumber="23">2178</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA316CFF5E6260FF00" box="[775,827,208,232]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO); Parque Nacional y área natural de manejo integrado
<collectorName id="26A45316FFAEFFEA37DFFF5E6335FEE2" pageId="22" pageNumber="23">
de la
<location id="8E8E601BFFAEFFEA307EFF7C6397FEE2" LSID="urn:lsid:plazi:treatment:03F887D6FFAAFFEA305EFD896314F9D8:8E8E601BFFAEFFEA307EFF7C6397FEE2" box="[533,716,242,266]" country="Bolivia" county="Hernando Siles" municipality="Monteagudo" name="Serrania del Inao" pageId="22" pageNumber="23" stateProvince="Chuquisaca">Serranía del Iñao</location>
</collectorName>
, cuenca del
<location id="8E8E601BFFAEFFEA3127FF7C62E1FEE2" LSID="urn:lsid:plazi:treatment:03F887D6FFAAFFEA305EFD896314F9D8:8E8E601BFFAEFFEA3127FF7C62E1FEE2" box="[844,954,242,266]" country="Bolivia" county="Hernando Siles" municipality="Monteagudo" name="rio Limon" pageId="22" pageNumber="23" stateProvince="Chuquisaca">río Limón</location>
, 19˚4401&quot;S, 63˚5452&quot;W, 1247 m,
<date id="FFEF1000FFAEFFEA374DFF7C64F3FEE2" box="[1318,1448,242,266]" pageId="22" pageNumber="23" value="2006-12-15">
<collectingDate id="EFABE9E8FFAEFFEA374DFF7C64F3FEE2" box="[1318,1448,242,266]" pageId="22" pageNumber="23" value="2006-12-15">15 Dec 2006</collectingDate>
</date>
(fr),
<collectorName id="26A45316FFAEFFEA37B6FF7D630EFEC5" pageId="22" pageNumber="23">
<emphasis id="B925EAD2FFAEFFEA37B6FF7D64B7FEE2" box="[1501,1516,243,266]" italics="true" pageId="22" pageNumber="23">E</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA307EFE9B63DCFEC5" box="[533,647,277,301]" italics="true" pageId="22" pageNumber="23">Portal et al</emphasis>
</collectorName>
.
<emphasis id="B925EAD2FFAEFFEA30F8FE9B63E2FEC5" box="[659,697,277,301]" italics="true" pageId="22" pageNumber="23">108</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA30ACFE9B63A0FEC5" box="[711,763,277,301]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO); foot of Cerro Urkhal path before 2nd river crossing, 19˚48S 63˚ 57W, 1300 m,
<date id="FFEF1000FFAEFFEA30DFFEB6627CFEB8" box="[692,807,312,336]" pageId="22" pageNumber="23" value="2000-10-04">
<collectingDate id="EFABE9E8FFAEFFEA30DFFEB6627CFEB8" box="[692,807,312,336]" pageId="22" pageNumber="23" value="2000-10-04">4 Oct 2000</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFAEFFEA3132FEB6655CFEA7" box="[857,1031,311,336]" pageId="22" pageNumber="23">
<emphasis id="B925EAD2FFAEFFEA3132FEB66231FEA7" box="[857,874,312,335]" italics="true" pageId="22" pageNumber="23">K</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA311DFEB96568FEB8" box="[886,1075,311,336]" italics="true" pageId="22" pageNumber="23">Wendelberger 170</emphasis>
</collectorName>
(
<collectionCode id="ED40AE05FFAEFFEA362BFEB6652FFEB8" box="[1088,1140,312,336]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO); Luis Calvo, Ticucha, serranía del Iñao, 12 km al NO de la comunidad
<collectorName id="26A45316FFAEFFEA31D6FED4656FFE9A" box="[957,1076,346,370]" pageId="22" pageNumber="23">de Ticucha</collectorName>
, 19˚350.4”S, 63˚5312.7”W, 1431 m,
<date id="FFEF1000FFAEFFEA37D7FED46323FE7D" pageId="22" pageNumber="23" value="2003-04-11">
<collectingDate id="EFABE9E8FFAEFFEA37D7FED46323FE7D" pageId="22" pageNumber="23" value="2003-04-11">11 Apr 2003</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFAEFFEA30C1FEF36270FE7C" box="[682,811,381,405]" pageId="22" pageNumber="23">
<emphasis id="B925EAD2FFAEFFEA30C1FEF363E7FE7C" box="[682,700,381,404]" italics="true" pageId="22" pageNumber="23">A</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA30ACFEF36206FE7D" box="[711,861,381,405]" italics="true" pageId="22" pageNumber="23">Carretero et al</emphasis>
</collectorName>
.
<emphasis id="B925EAD2FFAEFFEA3101FEF362CBFE7D" box="[874,912,381,405]" italics="true" pageId="22" pageNumber="23">824</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA31F6FEF3628AFE7D" box="[925,977,381,405]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO, NY); Entierrillos, aprox. a 5 km de la escuela
<collectorName id="26A45316FFAEFFEA307EFE2E63F5FE50" box="[533,686,416,440]" pageId="22" pageNumber="23">de Entierrillos</collectorName>
, serranía del Iñao, 19˚31S, 63˚52W, 1700 m,
<date id="FFEF1000FFAEFFEA36FFFE2E644DFE50" box="[1172,1302,416,440]" pageId="22" pageNumber="23" value="2003-12-18">
<collectingDate id="EFABE9E8FFAEFFEA36FFFE2E644DFE50" box="[1172,1302,416,440]" pageId="22" pageNumber="23" value="2003-12-18">18 Dec 2003</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFAEFFEA3723FE2E6491FE5F" box="[1352,1482,416,440]" pageId="22" pageNumber="23">
<emphasis id="B925EAD2FFAEFFEA3723FE2E6401FE5F" box="[1352,1370,416,439]" italics="true" pageId="22" pageNumber="23">A</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA370DFE2E6491FE5F" box="[1382,1482,416,439]" italics="true" pageId="22" pageNumber="23">Carretero</emphasis>
</collectorName>
</materialsCitation>
</emphasis>
<emphasis id="B925EAD2FFAEFFEA307EFE4D631AFE32" box="[533,577,450,474]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA3026FE4C6328FE32" box="[589,627,450,474]" italics="true" pageId="22" pageNumber="23">939</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA30EBFE4C63EFFE32" box="[640,692,450,474]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO); Serranía del Iñao, pasando la Laguna, 19˚31S, 63˚52W,
<date id="FFEF1000FFAEFFEA373EFE4C6483FE32" box="[1365,1496,450,474]" pageId="22" pageNumber="23" value="2003-12-18">18 Dec 2003</date>
(fl),
<emphasis id="B925EAD2FFAEFFEA3029FE6B630FFE14" box="[578,596,485,508]" italics="true" pageId="22" pageNumber="23">A</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA3034FE6B63AEFE15" box="[607,757,485,509]" italics="true" pageId="22" pageNumber="23">Carretero et al</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA316AFE6B627CFE15" box="[769,807,485,509]" italics="true" pageId="22" pageNumber="23">998</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA315FFE6B6233FE15" box="[820,872,485,509]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO, NY); Las Frías, ca. a la la cima de la serranía de Ñahuañanca, 19˚0930.6&quot;S, 63˚5040.6&quot;W,
<quantity id="4CA99B25FFAEFFEA31BEFD86657EFDC8" box="[981,1061,520,544]" metricMagnitude="3" metricUnit="m" metricValue="1.93" pageId="22" pageNumber="23" unit="m" value="1930.0">1930 m</quantity>
,
<date id="FFEF1000FFAEFFEA3644FD8665EAFDC8" box="[1071,1201,520,544]" pageId="22" pageNumber="23" value="2003-12-22">22 Dec 2003</date>
(fl, fr),
<emphasis id="B925EAD2FFAEFFEA376AFD866448FDF7" box="[1281,1299,520,543]" italics="true" pageId="22" pageNumber="23">A</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA3775FD8664EFFDF7" box="[1310,1460,519,543]" italics="true" pageId="22" pageNumber="23">Carretero et al</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA37ABFD8964A8FDF7" box="[1472,1523,519,543]" italics="true" pageId="22" pageNumber="23">1067</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA3076FDA4630AFDAA" box="[541,593,554,578]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO, NY); Las Frías, ca. 1/
<quantity id="4CA99B25FFAEFFEA3107FDA462F8FDAA" box="[876,931,554,578]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="23" unit="km" value="2.0">2 km</quantity>
de la vivienda de Sr. Severino Daza, hacia la cima de la serranía de Yahuañanca, 19˚0931&quot;S, 63˚5023&quot;W,
<quantity id="4CA99B25FFAEFFEA364CFDC36523FD8D" box="[1063,1144,589,613]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="23" unit="m" value="1600.0">1600 m</quantity>
,
<date id="FFEF1000FFAEFFEA36E9FDC3645FFD8D" box="[1154,1284,589,613]" pageId="22" pageNumber="23" value="2004-12-23">23 Dec 2004</date>
(fl),
<emphasis id="B925EAD2FFAEFFEA375DFDC36413FD8C" box="[1334,1352,589,612]" italics="true" pageId="22" pageNumber="23">A</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA3738FDC364B2FD8D" box="[1363,1513,589,613]" italics="true" pageId="22" pageNumber="23">Carretero et al</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA307EFDFE6313FD60" box="[533,584,624,648]" italics="true" pageId="22" pageNumber="23">1085</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA303EFDFE63D2FD60" box="[597,649,624,648]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO, NY); Sud Cinti, ca. 3 horas en caballo al NW de la comunidad de Orocote entre los ríos Limonal y Cochayo, 20˚47S, 64˚21W,
<quantity id="4CA99B25FFAEFFEA3628FD1C65CFFD42" box="[1091,1172,658,682]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="22" pageNumber="23" unit="m" value="1650.0">1650 m</quantity>
,
<date id="FFEF1000FFAEFFEA36F6FD1C6444FD42" box="[1181,1311,658,682]" pageId="22" pageNumber="23" value="2005-04-29">29 Apr 2005</date>
(fr),
<emphasis id="B925EAD2FFAEFFEA373EFD1D643EFD42" box="[1365,1381,659,682]" italics="true" pageId="22" pageNumber="23">R</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA371BFD1D64ADFD42" box="[1392,1526,658,682]" italics="true" pageId="22" pageNumber="23">Lozano 1207</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA3076FD3B630AFD25" box="[541,593,693,717]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO); Tomina, aprox.
<specimenCount id="9D57FD49FFAEFFEA312DFD3B62D5FD25" box="[838,910,693,717]" count="800" pageId="22" pageNumber="23" type="generic">
<quantity id="4CA99B25FFAEFFEA312DFD3B62D2FD25" box="[838,905,693,717]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="23" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
antes de llegar a Llantoj, de
<taxonomicName id="4C514D43FFAEFFEA36DCFD3B647FFD25" box="[1207,1316,693,717]" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="florida">La Florida</taxonomicName>
subiendo hacia el E de la Serranía de Kaska Orcko, 19˚0946&quot;S, 64˚0342&quot;W,
<quantity id="4CA99B25FFAEFFEA3603FD5665E3FD18" box="[1128,1208,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="22" pageNumber="23" unit="m" value="1750.0">1750 m</quantity>
,
<date id="FFEF1000FFAEFFEA36A9FD56641AFD18" box="[1218,1345,728,752]" pageId="22" pageNumber="23" value="2004-10-11">11 Oct 2004</date>
(fl, fr),
<emphasis id="B925EAD2FFAEFFEA37FAFD5664C2FD07" box="[1425,1433,728,751]" italics="true" pageId="22" pageNumber="23">J</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA37CEFD566311FCFA" italics="true" pageId="22" pageNumber="23">Gutiérrez R</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA303DFD7463D2FCFA" box="[598,649,762,786]" italics="true" pageId="22" pageNumber="23">1004</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA30FDFD746391FCFA" box="[662,714,762,786]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO); Llantoj, aprox.
<quantity id="4CA99B25FFAEFFEA31DEFD7462ACFCFA" box="[949,1015,762,786]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="22" pageNumber="23" unit="m" value="800.0">800 m</quantity>
antes de llegar a Llantoj, de
<taxonomicName id="4C514D43FFAEFFEA374AFD7464DDFCFA" box="[1313,1414,762,786]" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="florida">la Florida</taxonomicName>
subiendo hacia el E de la Serranía de Kaska Orcko, 19˚0946&quot;S, 64˚0342&quot;W,
<quantity id="4CA99B25FFAEFFEA36B9FC936478FCDD" box="[1234,1315,797,821]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="22" pageNumber="23" unit="m" value="1750.0">1750 m</quantity>
,
<date id="FFEF1000FFAEFFEA3747FC9364F4FCDD" box="[1324,1455,797,821]" pageId="22" pageNumber="23" value="2004-12-15">15 Dec 2004</date>
(fr),
<emphasis id="B925EAD2FFAEFFEA378FFC9364B7FCDC" box="[1508,1516,797,820]" italics="true" pageId="22" pageNumber="23">J</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA307EFCCE63D5FCBF" box="[533,654,832,855]" italics="true" pageId="22" pageNumber="23">Gutiérrez R</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA30F1FCCE6396FCB0" box="[666,717,832,856]" italics="true" pageId="22" pageNumber="23">1072</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA30B1FCCE6255FCB0" box="[730,782,832,856]" country="Bolivia" lsid="urn:lsid:biocol.org:col:14349" name="Universidad Mayor Real y Pontifícia de San Francisco Xavier de Chuquisaca" pageId="22" pageNumber="23" type="Herbarium">HSB</collectionCode>
, MO); Rio Limón Valley between Padilla and Monteagudo,
<quantity id="4CA99B25FFAEFFEA37E2FCCE6482FCB0" box="[1417,1497,832,856]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="23" unit="m" value="1500.0">1500 m</quantity>
,
<date id="FFEF1000FFAEFFEA3788FCCE632AFC92" pageId="22" pageNumber="23" value="1995-01-01">1 Jan 1995</date>
(fl),
<emphasis id="B925EAD2FFAEFFEA30C8FCED63F0FC92" box="[675,683,867,890]" italics="true" pageId="22" pageNumber="23">J</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA30DCFCED639CFC92" box="[695,711,867,890]" italics="true" pageId="22" pageNumber="23">R</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA30B8FCED6387FC92" box="[723,732,867,890]" italics="true" pageId="22" pageNumber="23">I</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA3083FCEC623BFC92" box="[744,864,866,890]" italics="true" pageId="22" pageNumber="23">Wood 9104</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA3106FCEC62CCFC92" box="[877,919,866,890]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="22" pageNumber="23" type="Herbarium">NY</collectionCode>
).
<emphasis id="B925EAD2FFAEFFEA31C3FCEC6571FC92" bold="true" box="[936,1066,866,890]" pageId="22" pageNumber="23">
<collectingRegion id="4995F822FFAEFFEA31C3FCEC657EFC92" box="[936,1061,866,890]" country="Bolivia" name="Santa Cruz" pageId="22" pageNumber="23">Santa Cruz</collectingRegion>
.
</emphasis>
Prov.
<taxonomicName id="4C514D43FFAEFFEA3605FCEC6366FC75" authority=", Mairana, La Yunga de Mairana" authorityName="Mairana, La Yunga de Mairana" class="Magnoliopsida" family="Solanaceae" genus="Lycianthes" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="florida">Florida, Mairana, La Yunga de Mairana</taxonomicName>
, 18˚0413”S, 63˚5508”W,
<quantity id="4CA99B25FFAEFFEA3138FC0B62F8FC75" box="[851,931,901,925]" metricMagnitude="3" metricUnit="m" metricValue="2.19" pageId="22" pageNumber="23" unit="m" value="2190.0">2190 m</quantity>
,
<date id="FFEF1000FFAEFFEA31C6FC0B6569FC75" box="[941,1074,901,925]" pageId="22" pageNumber="23" value="2004-11-15">15 Nov 2004</date>
(fl),
<emphasis id="B925EAD2FFAEFFEA360FFC0B6520FC74" box="[1124,1147,901,924]" italics="true" pageId="22" pageNumber="23">M</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA36ECFC0B6451FC75" box="[1159,1290,901,925]" italics="true" pageId="22" pageNumber="23">Serrano et al</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA377CFC0B6411FC75" box="[1303,1354,901,925]" italics="true" pageId="22" pageNumber="23">5482</emphasis>
(
<collectionCode id="ED40AE05FFAEFFEA373CFC0B64DAFC75" box="[1367,1409,901,925]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="22" pageNumber="23" type="Herbarium">NY</collectionCode>
).
</paragraph>
<paragraph id="8BEE36C0FFAEFFEA305EFC26651AFB10" blockId="22.[533,1536,207,1584]" pageId="22" pageNumber="23">
<taxonomicName id="4C514D43FFAEFFEA305EFC266396FC57" box="[565,717,936,959]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFAEFFEA305EFC266396FC57" box="[565,717,936,959]" italics="true" pageId="22" pageNumber="23">Capsicum neei</emphasis>
</taxonomicName>
is morphologically most similar to the Bolivian
<taxonomicName id="4C514D43FFAEFFEA36A2FC26642FFC57" box="[1225,1396,935,960]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="minutiflorum">
<emphasis id="B925EAD2FFAEFFEA36A2FC266581FC57" box="[1225,1242,936,959]" italics="true" pageId="22" pageNumber="23">C</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA368DFC29642FFC57" box="[1254,1396,935,959]" italics="true" pageId="22" pageNumber="23">minutiflorum</emphasis>
</taxonomicName>
in having stellate yellow corolla and red fruit at maturity. It can be distinguished by the non-geniculate pendent flowering pedicels and the strongly nerved calyx with 10 unequal appendages (
<figureCitation id="136A2A45FFAEFFEA37C4FC63632FFBC0" captionStart="Fig 10" captionStartId="19.[388,416,1825,1844]" captionTargetBox="[388,1533,208,1809]" captionTargetId="figure-1@19.[388,1536,208,1810]" captionTargetPageId="19" captionText="Fig 10. Capsicum neei Barboza &amp; X. Reyes. (A) Flowering branch. (B) Inflorescence. (C) Flower bud. (D) Flower. (E) Calyx. (F) Opened corolla.(G) Gynoecium (H) Fruit. (I) Glandular trichome of the inside calyx. (J) Glandular trichome of the pedicels.(K)" figureDoi="http://doi.org/10.5281/zenodo.12630935" httpUri="https://zenodo.org/record/12630935/files/figure.png" pageId="22" pageNumber="23">Figs 10C, 10E</figureCitation>
and
<figureCitation id="136A2A45FFAEFFEA30C2FB9E638AFBC0" box="[681,721,1040,1064]" captionStart="Fig 11" captionStartId="20.[533,561,1818,1837]" captionTargetBox="[533,1386,517,1797]" captionTargetId="figure-104@20.[533,1386,517,1797]" captionTargetPageId="20" captionText="Fig 11. Capsicum neei Barboza &amp; X. Reyes. (A) Plant. (B) Flower bud. (C) Flowering branch. Photos by G. E. Barboza. https://doi.org/10.1371/journal.pone.0209792.g011" figureDoi="http://doi.org/10.5281/zenodo.12630937" httpUri="https://zenodo.org/record/12630937/files/figure.png" pageId="22" pageNumber="23">11B</figureCitation>
) versus the geniculate and erect flowering pedicels and the calyx weakly nerved and with 5 equal short appendages in
<taxonomicName id="4C514D43FFAEFFEA319FFBBD641AFBA2" authority="(Rusby) Hunz." authorityName="Hunz." baseAuthorityName="Rusby" box="[1012,1345,1074,1098]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="minutiflorum">
<emphasis id="B925EAD2FFAEFFEA319FFBBD655EFBA2" box="[1012,1029,1075,1098]" italics="true" pageId="22" pageNumber="23">C</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA367AFBBC65C4FBA2" box="[1041,1183,1074,1098]" italics="true" pageId="22" pageNumber="23">minutiflorum</emphasis>
(Rusby) Hunz.
</taxonomicName>
The flowers in
<taxonomicName id="4C514D43FFAEFFEA378FFBBD6364FB84" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFAEFFEA378FFBBD64AEFBA2" box="[1508,1525,1075,1098]" italics="true" pageId="22" pageNumber="23">C</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA307EFBDB6364FB84" box="[533,575,1109,1132]" italics="true" pageId="22" pageNumber="23">neei</emphasis>
</taxonomicName>
often appear to be solitary but the remains of 23 early deciduous bud or flower scars can be seen in the axils. Fruit features as pungency, presence of giant cells and sclerotic granules in the pericarp and mature seeds are unknown at present but it is probable that the fruits are pungent and have giant cells in the innermost layer of the pericarp as occur in the remaining species of the Bolivian clade where
<taxonomicName id="4C514D43FFAEFFEA310DFB6E62F6FB1F" box="[870,941,1248,1272]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFAEFFEA310DFB6E622CFB1F" box="[870,887,1248,1271]" italics="true" pageId="22" pageNumber="23">C</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA31E8FB6E62F6FB1F" box="[899,941,1248,1271]" italics="true" pageId="22" pageNumber="23">neei</emphasis>
</taxonomicName>
is positioned.
</paragraph>
<paragraph id="8BEE36C0FFAEFFEA305EFA8C64CEFA88" blockId="22.[533,1536,207,1584]" pageId="22" pageNumber="23">
This new species is sympatric with
<taxonomicName id="4C514D43FFAEFFEA31CCFA8D6571FAF2" box="[935,1066,1282,1306]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="baccatum">
<emphasis id="B925EAD2FFAEFFEA31CCFA8D62E3FAF2" box="[935,952,1283,1306]" italics="true" pageId="22" pageNumber="23">C</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA31AFFA8C6571FAF2" box="[964,1066,1282,1306]" italics="true" pageId="22" pageNumber="23">baccatum</emphasis>
</taxonomicName>
L.
<taxonomicName id="4C514D43FFAEFFEA3621FA8C6585FAF2" box="[1098,1246,1282,1306]" pageId="22" pageNumber="23" rank="variety" variety="baccatum">
var.
<emphasis id="B925EAD2FFAEFFEA3613FA8C6585FAF2" box="[1144,1246,1282,1306]" italics="true" pageId="22" pageNumber="23">baccatum</emphasis>
</taxonomicName>
, a taxon with a much wider distribution in South America, that has geniculate pedicels, calyx with 5 equal appendages, white corollas with greenish yellow spots inside and ovoid or globose red fruits.
</paragraph>
<paragraph id="8BEE36C0FFAEFFEA305EFAE56314F9D8" blockId="22.[533,1536,207,1584]" pageId="22" pageNumber="23">
<taxonomicName id="4C514D43FFAEFFEA305EFAE56396FA6A" box="[565,717,1387,1410]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="neei">
<emphasis id="B925EAD2FFAEFFEA305EFAE56396FA6A" box="[565,717,1387,1410]" italics="true" pageId="22" pageNumber="23">Capsicum neei</emphasis>
</taxonomicName>
has been resolved as a new member of the Bolivian clade, which is coherent with its geographic range and the main common feature recognized for the clade, the yellow corollas [
<bibRefCitation id="EFC04B31FFAEFFEA301CFA3E63DFFA20" author="Carrizo Garcia C &amp; Sehr EM &amp; Barfuss MHJ &amp; Barboza GE &amp; Samuel R &amp; Moscone EA" box="[631,644,1456,1480]" pageId="22" pageNumber="23" pagination="35 - 51" refId="ref12646" refString="1. Carrizo Garcia C, Sehr EM, Barfuss MHJ, Barboza GE, Samuel R, Moscone EA et al. Phylogenetic relationships, diversification and expansion of chili peppers (Capsicum, Solanaceae). Ann Bot. 2016; 118: 35 - 51. https: // doi. org / 10.1093 / aob / mcw 079 PMID: 27245634" type="journal article" year="2016">1</bibRefCitation>
]. However, the Bolivian clade has a weak support, most likely due to the apparent divergence of
<taxonomicName id="4C514D43FFAEFFEA30C1FA5D626FFA02" box="[682,820,1490,1514]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="coccineum">
<emphasis id="B925EAD2FFAEFFEA30C1FA5D63E0FA02" box="[682,699,1491,1514]" italics="true" pageId="22" pageNumber="23">C</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA30ADFA5D626FFA02" box="[710,820,1491,1514]" italics="true" pageId="22" pageNumber="23">coccineum</emphasis>
</taxonomicName>
from the rest of the species; indeed,
<taxonomicName id="4C514D43FFAEFFEA36DEFA5D641BFA02" box="[1205,1344,1490,1514]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="coccineum">
<emphasis id="B925EAD2FFAEFFEA36DEFA5D659DFA02" box="[1205,1222,1491,1514]" italics="true" pageId="22" pageNumber="23">C</emphasis>
.
<emphasis id="B925EAD2FFAEFFEA36B9FA5D641BFA02" box="[1234,1344,1491,1514]" italics="true" pageId="22" pageNumber="23">coccineum</emphasis>
</taxonomicName>
would deserve more attention considering some morphological variability observed in the species (GEB, pers. obs.).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,680 @@
<document id="72D4099B5E213194930FA38E07FF043B" ID-DOI="10.1371/journal.pone.0209792" ID-ISSN="1932-6203" ID-PMC="PMC6334993" ID-PubMed="30650102" ID-Zenodo-Dep="12630921" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1719975332143" checkinUser="felipe" docAuthor="Barboza, Gloria E., García, Carolina Carrizo, González, Segundo Leiva, Scaldaferro, Marisel &amp; Reyes, Ximena" docDate="2019" docId="03F887D6FFB5FFEE305EFED4620DFE15" docLanguage="en" docName="PLoSONE.14.1.e0209792.pdf" docOrigin="PLoS ONE (e 0209792) 14 (1)" docSource="http://dx.doi.org/10.1371/journal.pone.0209792" docStyle="DocumentStyle:55318D21378FF0528C342C9740A1E1BE.2:PLoSONE.2014-.journal_article" docStyleId="55318D21378FF0528C342C9740A1E1BE" docStyleName="PLoSONE.2014-.journal_article" docStyleVersion="2" docTitle="Capsicum piuranum Barboza &amp; S. Leiva 2019, sp. nov." docType="treatment" docVersion="1" lastPageNumber="19" masterDocId="FFC1FFAEFFB8FFFC326BFF8E615BFFE8" masterDocTitle="Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus" masterLastPageNumber="26" masterPageNumber="1" pageNumber="14" updateTime="1723725765965" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
<mods:mods id="E43DA2E61AECAF6B761B6A68D19543C4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CACFD2D05881FBCA26BFF9B903BE1216">
<mods:title id="65A37ADEAD747372E90D06F1302D4E2D">Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus</mods:title>
</mods:titleInfo>
<mods:name id="B29D1363EB7384574696A7DD5F2BB7F8" type="personal">
<mods:role id="F2706906D50C70C644D300CAE70588CB">
<mods:roleTerm id="395A0315488F12D0F88BC6D356E4A0EE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4C743E6E9488AD1B99B34D4EC626C76C">Barboza, Gloria E.</mods:namePart>
</mods:name>
<mods:name id="6CB0110C0D62987F7080017830751FC1" type="personal">
<mods:role id="F9DC1AA38736680F91A3ABEE86282714">
<mods:roleTerm id="09481EB709D8EF6A64F18B8CBFC44112">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="24EA451EF0A271835EE261B292931A93">García, Carolina Carrizo</mods:namePart>
<mods:affiliation id="690F85FBAA9320362AB6D41C5BB1EEA8">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Department of Botany and Biodiversity Research, University of Vienna, Vienna, Austria,</mods:affiliation>
</mods:name>
<mods:name id="2007766E7AC3FF50C0221CC375FF08BD" type="personal">
<mods:role id="CC4F43AAEC382FF384D9DC6D76E6B4DA">
<mods:roleTerm id="4B250A2C27A7A366A74C08F5BCD82D4A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5A99DF2FFCB136CC557F5C6251E1FB1D">González, Segundo Leiva</mods:namePart>
</mods:name>
<mods:name id="5773D2B03B8F434C4FCD5E5663E4AD57" type="personal">
<mods:role id="788BE88CEF62C09B9CE71D757B2AC0BC">
<mods:roleTerm id="266C8791247A3205D185F2C42775108D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F5F25952955B85B737C352A12C12EAB">Scaldaferro, Marisel</mods:namePart>
<mods:affiliation id="C13207DEA42DCD5B7D3F978348518FEA">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Córdoba, Argentina,</mods:affiliation>
</mods:name>
<mods:name id="A358C7716A525F8AEBF8080505C68F8A" type="personal">
<mods:role id="2543E920C1A53F8A8FBF4BD69B51EB30">
<mods:roleTerm id="55FE415C97F412630AA13E10B74A7EBF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3800C2647094651086D8DA4F8ADB5F6">Reyes, Ximena</mods:namePart>
<mods:affiliation id="312E99B3AFAEDA033E9511D3CB5A5FBC">Centro de Investigaciones Fitoecogenéticas de Pairumani, Cochabamba, Bolivia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="CE87308CD1022069B082511960054D61">text</mods:typeOfResource>
<mods:relatedItem id="D7FE4E4284C50D720A2DA2E009416E66" type="host">
<mods:titleInfo id="3563C009B496025E9FDF684A6614E778">
<mods:title id="FA7824CC2F3A8288C77234DB89F4F9A7">PLoS ONE</mods:title>
</mods:titleInfo>
<mods:part id="678B4A834068B3C7CC6563E8C1226D28">
<mods:date id="292580EB50AF0FC3B5A78406A7C09913">2019</mods:date>
<mods:detail id="7F3E6BE61E7835EF70EC35E709210FE5" type="series">
<mods:title id="E570181F94395FF6BCE79854641E6EC8">e 0209792</mods:title>
</mods:detail>
<mods:detail id="AF2B14BBDB5194D458CB1F3B0080CBEC" type="pubDate">
<mods:number id="CBD81D310E2E58E294D7C00038C2C65E">2019-01-16</mods:number>
</mods:detail>
<mods:detail id="906F6E8ED7AD43935CF1B186FAB84833" type="volume">
<mods:number id="F4FA3BCF60DC95AE3B6B9AA188E3A204">14</mods:number>
</mods:detail>
<mods:detail id="2DECD5072989A5339F99EDDB3583C4F3" type="issue">
<mods:number id="EA6EFD75CB41B64AE8C15DCF27A906C5">1</mods:number>
</mods:detail>
<mods:extent id="C9EA9CD831B5A2D9538BC97A39D109D9" unit="page">
<mods:start id="CC7A709186AA84B0ED0CBC40A8DE18E7">1</mods:start>
<mods:end id="9B15A165EAD2705A1BF5A193996984E2">26</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="85912AF41852310B259B186D70C95974">
<mods:url id="FDCED23236C2E03651D567109E420ACF">http://dx.doi.org/10.1371/journal.pone.0209792</mods:url>
</mods:location>
<mods:classification id="28B2007D7BEB25C3F5E9EBBFB84195E8">journal article</mods:classification>
<mods:identifier id="D6C83915AF94E7F5BBFB8B5FC092EA46" type="DOI">10.1371/journal.pone.0209792</mods:identifier>
<mods:identifier id="D79207144C22346CF1CB8B39F9F45AFF" type="ISSN">1932-6203</mods:identifier>
<mods:identifier id="24043230C0DC35E5869DF743F8884292" type="PMC">PMC6334993</mods:identifier>
<mods:identifier id="CD9CC0DF4E121C96A5ACB2AE7DE1FBAA" type="PubMed">30650102</mods:identifier>
<mods:identifier id="F919CEA70E83F945877C94A540CACF0C" type="Zenodo-Dep">12630921</mods:identifier>
</mods:mods>
<treatment id="03F887D6FFB5FFEE305EFED4620DFE15" LSID="urn:lsid:plazi:treatment:03F887D6FFB5FFEE305EFED4620DFE15" httpUri="http://treatment.plazi.org/id/03F887D6FFB5FFEE305EFED4620DFE15" lastPageId="18" lastPageNumber="19" pageId="13" pageNumber="14">
<subSubSection id="C34B654BFFB5FFF1305EFED464BCFE9A" box="[565,1511,346,370]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BEE36C0FFB5FFF1305EFED464BCFE9A" blockId="13.[533,1536,208,1896]" box="[565,1511,346,370]" pageId="13" pageNumber="14">
<emphasis id="B925EAD2FFB5FFF1305EFED46517FE9A" bold="true" box="[565,1100,346,370]" pageId="13" pageNumber="14">
<taxonomicName id="4C514D43FFB5FFF1305EFED462B7FE9A" ID-CoL="7TBQR" authority="Barboza &amp; S. Leiva" authorityName="Barboza &amp; S. Leiva" authorityYear="2019" box="[565,1004,346,370]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="piuranum" status="sp. nov.">
<emphasis id="B925EAD2FFB5FFF1305EFED4624EFE9A" bold="true" box="[565,789,346,370]" italics="true" pageId="13" pageNumber="14">Capsicum piuranum</emphasis>
Barboza &amp; S. Leiva
</taxonomicName>
,
<taxonomicNameLabel id="A21657A9FFB5FFF1319CFED46517FE9A" box="[1015,1100,346,370]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
[urn:lsid:ipni.org:names: 771925591]
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB5FFF1305EFEF363EFFE15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BEE36C0FFB5FFF1305EFEF3629CFE32" blockId="13.[533,1536,208,1896]" pageId="13" pageNumber="14">
<materialsCitation id="3B393C9DFFB5FFF1305EFEF36522FE50" collectingDate="2011-03-22" collectorName="T. Mione" country="Peru" elevation="2385" location="Type" pageId="13" pageNumber="14" specimenCount="2" stateProvince="Piura" typeStatus="holotype">
<location id="8E8E601BFFB5FFF1305EFEF36336FE7D" LSID="urn:lsid:plazi:treatment:03F887D6FFB5FFEE305EFED4620DFE15:8E8E601BFFB5FFF1305EFEF36336FE7D" box="[565,621,381,405]" country="Peru" name="Type" pageId="13" pageNumber="14" stateProvince="Piura">Type</location>
:
<collectingCountry id="F3467650FFB5FFF1301CFEF363F7FE7D" box="[631,684,381,405]" name="Peru" pageId="13" pageNumber="14">Peru</collectingCountry>
.
<collectingRegion id="4995F822FFB5FFF130DEFEF363AAFE7D" box="[693,753,381,405]" country="Peru" name="Piura" pageId="13" pageNumber="14">Piura</collectingRegion>
: Prov. Huancabamba, borde de carretera y riachuelo, 5˚2246”S, 79˚ 3347”W,
<quantity id="4CA99B25FFB5FFF13015FE2E6254FE50" box="[638,783,416,440]" metricMagnitude="3" metricUnit="m" metricValue="2.385" metricValueMax="2.459" metricValueMin="2.311" pageId="13" pageNumber="14" unit="m" value="2385.0" valueMax="2459.0" valueMin="2311.0">23112459 m</quantity>
,
<date id="FFEF1000FFB5FFF13172FE2E62C5FE50" box="[793,926,416,440]" pageId="13" pageNumber="14" value="2011-03-22">
<collectingDate id="EFABE9E8FFB5FFF13172FE2E62C5FE50" box="[793,926,416,440]" pageId="13" pageNumber="14" value="2011-03-22">22 Mar 2011</collectingDate>
</date>
(fl, fr),
<collectorName id="26A45316FFB5FFF13186FE2E6516FE5F" box="[1005,1101,416,440]" pageId="13" pageNumber="14">
<emphasis id="B925EAD2FFB5FFF13186FE2E62A6FE5F" box="[1005,1021,416,439]" italics="true" pageId="13" pageNumber="14">T</emphasis>
.
<emphasis id="B925EAD2FFB5FFF13662FE2E6522FE50" box="[1033,1145,416,440]" italics="true" pageId="13" pageNumber="14">Mione 812</emphasis>
</collectorName>
</materialsCitation>
<materialsCitation id="3B393C9DFFB5FFF13615FE2E63D1FE32" collectingDate="2011-03-22" collectorName="T. Mione" country="Peru" elevation="2385" location="Type" pageId="13" pageNumber="14" specimenCount="2" stateProvince="Piura" typeStatus="holotype">
(
<typeStatus id="54EA8862FFB5FFF136EDFE2E65BCFE50" box="[1158,1255,416,440]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
, CORD [CORD-00006936];
</materialsCitation>
<materialsCitation id="3B393C9DFFB5FFF130FBFE4C6299FE32" box="[656,962,450,474]" collectingDate="2011-03-22" collectorName="T. Mione" country="Peru" elevation="2385" location="Type" pageId="13" pageNumber="14" specimenCount="2" stateProvince="Piura" typeStatus="holotype">
<typeStatus id="54EA8862FFB5FFF130FBFE4C6385FE32" box="[656,734,450,474]" pageId="13" pageNumber="14" type="isotype">isotype</typeStatus>
, NY [NY-03231447])
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36C0FFB5FFF1305EFE6B63EFFE15" blockId="13.[533,1536,208,1896]" box="[565,692,485,509]" pageId="13" pageNumber="14">
<figureCitation id="136A2A45FFB5FFF1305EFE6B6328FE15" box="[565,627,485,509]" captionStart="Fig 7" captionStartId="15.[373,401,203,222]" captionTargetId="figure-1@14.[373,1536,208,1872]" captionTargetPageId="14" captionText="Fig 7. Capsicum piuranum Barboza &amp; S. Leiva. (A) Flowering branch. (B) Calyx. (C) Flower. (D) Opened corolla. (E, F, G). Anther, ventral, dorsal and lateral view, respectively. (H) Gynoecium. (I) Fruit. (J) Seed. Drawn by S. Leiva González." pageId="13" pageNumber="14">Figs 7</figureCitation>
and
<figureCitation id="136A2A45FFB5FFF130CCFE6B63EFFE15" box="[679,692,485,509]" captionStart="Fig 8" captionStartId="16.[527,555,1702,1721]" captionTargetBox="[527,1536,208,1680]" captionTargetId="figure-3@16.[527,1536,208,1680]" captionTargetPageId="16" captionText="Fig 8. Capsicum piuranum Barboza &amp; S. Leiva. (A) Plant. (B) Leaves, abaxial surface. (C) Fruiting branch. (D) Flower bud. (E) Flower and immature fruit. (F) Mature fruit. (G) Fruit, transverse section, showing placenta and seeds. (H) Fruit transverse section, showing a stone cell at the apex (arrow). Photos by S. Leiva González and G. E. Barboza. https://doi.org/10.1371/journal.pone.0209792.g008" figureDoi="http://doi.org/10.5281/zenodo.12630929" httpUri="https://zenodo.org/record/12630929/files/figure.png" pageId="13" pageNumber="14">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB5FFF1305EFD86622EFD8D" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8BEE36C0FFB5FFF1305EFD86622EFD8D" blockId="13.[533,1536,208,1896]" pageId="13" pageNumber="14">
<emphasis id="B925EAD2FFB5FFF1305EFD8663F8FDC8" bold="true" box="[565,675,520,544]" pageId="13" pageNumber="14">Diagnosis</emphasis>
. Like
<taxonomicName id="4C514D43FFB5FFF13088FD866551FDC8" authority="M. Nee" authorityName="M. Nee" box="[739,1034,519,544]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="caballeroi">
<emphasis id="B925EAD2FFB5FFF13088FD8662ECFDF7" box="[739,951,519,543]" italics="true" pageId="13" pageNumber="14">Capsicum caballeroi</emphasis>
M. Nee
</taxonomicName>
but differing in the purple calyx, the 5 equal calyx appendages, the longer tubular-campanulate corolla, the globose orange non-pungent mature fruit, and the black seeds.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB5FFF1305EFDFE64B4F8CA" pageId="13" pageNumber="14" type="description">
<paragraph id="8BEE36C0FFB5FFF1305EFDFE64B4F8CA" blockId="13.[533,1536,208,1896]" pageId="13" pageNumber="14">
<emphasis id="B925EAD2FFB5FFF1305EFDFE63E3FD60" bold="true" box="[565,696,624,648]" pageId="13" pageNumber="14">Description</emphasis>
. Scandent shrubs 22.20 (3) m tall, densely branched. Young stems green, shiny, fragile, flexuous, glabrous, striate; bark of older stems green to dark brown, glabrous, striate; lenticels absent. Sympodial units difoliate, geminate, leaf pair markedly anisophyllous in size and shape. Leaves simple, membranaceous, discolorous, adaxial surface dark green and shiny, abaxial surface light green and opaque, glabrous or with sparse simple antrorse trichomes
<quantity id="4CA99B25FFB5FFF13006FC9363B0FCDD" box="[621,747,797,821]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="13" pageNumber="14" unit="mm" value="0.85" valueMax="1.2" valueMin="0.5">0.51.2 mm</quantity>
long adaxially and abaxially, occasionally trichomes more abundant on main veins and margins; the larger leaves with blades (8)
<quantity id="4CA99B25FFB5FFF13619FCCE65B7FCB0" box="[1138,1260,832,856]" metricMagnitude="-1" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.77" metricValueMin="1.2" pageId="13" pageNumber="14" unit="cm" value="14.85" valueMax="17.7" valueMin="12.0">1217.7 cm</quantity>
long, (2)
<quantity id="4CA99B25FFB5FFF13739FCCE649EFCB0" box="[1362,1477,832,856]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="13" pageNumber="14" unit="cm" value="3.5" valueMax="4.5" valueMin="2.5">2.54.5 cm</quantity>
wide, elliptic, major veins 79 on each side of midvein, base asymmetric and attenuate, margin entire, apex long-acuminate; petioles 0.71.4 (1.7) cm long, slightly winged from the decurrent leaf bases, glabrous or glabrescent with trichomes like those of the leaves; the minor leaves
<quantity id="4CA99B25FFB5FFF137BAFC266307FC0A" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.5" metricValueMin="2.5" pageId="13" pageNumber="14" unit="cm" value="3.5" valueMax="4.5" valueMin="2.5">2.5 4.5 cm</quantity>
long,
<quantity id="4CA99B25FFB5FFF130F6FC44624BFC0A" box="[669,784,970,994]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.6" metricValueMin="1.5" pageId="13" pageNumber="14" unit="cm" value="2.05" valueMax="2.6" valueMin="1.5">1.52.6 cm</quantity>
wide, ovate or elliptic, major veins 34 on each side of midvein, base short attenuate or rounded, asymmetric, margin entire, apex acute or slightly rounded; petioles
<quantity id="4CA99B25FFB5FFF1307EFB9E63D2FBC0" box="[533,649,1040,1064]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="13" pageNumber="14" unit="cm" value="0.35" valueMax="0.5" valueMin="0.2">0.20.5 cm</quantity>
long, glabrescent or pubescent. Flowers solitary or in fascicles of 3; flowering pedicels green, filiform, terete, pendent, slightly curved, not geniculate at anthesis,
<quantity id="4CA99B25FFB5FFF1373BFBBC6498FBA2" box="[1360,1475,1074,1098]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.6" metricValueMin="1.9" pageId="13" pageNumber="14" unit="cm" value="2.25" valueMax="2.6" valueMin="1.9">1.92.6 cm</quantity>
long, glabrous or glabrescent, the trichomes simple, non-glandular, multicellular, antrorse,
<quantity id="4CA99B25FFB5FFF137F5FBDB6328FB78" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="13" pageNumber="14" unit="mm" value="0.375" valueMax="0.45" valueMin="0.3">0.30 0.45 mm</quantity>
long. Flower buds ovoid, yellow or pale yellow. Calyx 1.52.6 (3) mm long,
<quantity id="4CA99B25FFB5FFF137F8FBF664B7FB78" box="[1427,1516,1144,1168]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="13" pageNumber="14" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
wide, cup-shaped, thick, purple or greenish purple, the margin truncate, glabrescent to pubescent, with 5 appendages (0.9)
<quantity id="4CA99B25FFB5FFF1313BFB3362E0FB3D" box="[848,955,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="13" pageNumber="14" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.53 mm</quantity>
long,
<quantity id="4CA99B25FFB5FFF13197FB336521FB3D" box="[1020,1146,1213,1237]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="13" pageNumber="14" unit="mm" value="0.65" valueMax="0.8" valueMin="0.5">0.50.8 mm</quantity>
wide, thick, erect, subulate, inserted close to the margin, glabrous or glabrescent with the same trichomes as pedicels and calyx tube. Corolla
<quantity id="4CA99B25FFB5FFF130CEFA8C6271FAF2" box="[677,810,1282,1306]" metricMagnitude="-2" metricUnit="m" metricValue="1.575" metricValueMax="1.7" metricValueMin="1.45" pageId="13" pageNumber="14" unit="mm" value="15.75" valueMax="17.0" valueMin="14.5">14.517 mm</quantity>
long,
<quantity id="4CA99B25FFB5FFF13100FA8C6286FAF2" box="[875,989,1282,1306]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.7" metricValueMin="1.2" pageId="13" pageNumber="14" unit="mm" value="14.5" valueMax="17.0" valueMin="12.0">1217 mm</quantity>
diam, tubular-campanulate, thick, entirely yellow; tube
<quantity id="4CA99B25FFB5FFF13021FAAB63E7FAD5" box="[586,700,1317,1341]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="13" pageNumber="14" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
long, glabrous inside and outside; lobes
<quantity id="4CA99B25FFB5FFF13603FAAB658FFAD5" box="[1128,1236,1317,1341]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="13" pageNumber="14" unit="mm" value="4.25" valueMax="5.0" valueMin="3.5">3.55 mm</quantity>
long,
<quantity id="4CA99B25FFB5FFF1377FFAAB64DBFAD5" box="[1300,1408,1317,1341]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="13" pageNumber="14" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.55 mm</quantity>
wide, broadly ovate, erect, glabrous adaxially and abaxially, the tips papillose. Stamens 5, equal, filaments subequal,
<quantity id="4CA99B25FFB5FFF130ACFAE4627BFA6A" box="[711,800,1386,1410]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="13" pageNumber="14" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
long, greenish white, glabrous, inserted on the corolla
<quantity id="4CA99B25FFB5FFF13709FAE464E0FA6A" box="[1378,1467,1386,1410]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="13" pageNumber="14" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
from the base, with inconspicuous auricles at point of insertion; anthers 22.5 (2.8) mm long, slightly connivent before anthesis, elliptic, yellowish white. Ovary
<quantity id="4CA99B25FFB5FFF136A5FA3E6402FA20" box="[1230,1369,1456,1480]" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.5" metricValueMin="1.25" pageId="13" pageNumber="14" unit="mm" value="1.375" valueMax="1.5" valueMin="1.25">1.251.5 mm</quantity>
long,
<quantity id="4CA99B25FFB5FFF137F1FA3E64B0FA20" box="[1434,1515,1456,1480]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="14" unit="mm" value="1.5">1.5 mm</quantity>
diam, subglobose, white, glabrous; nectary ca.
<quantity id="4CA99B25FFB5FFF13195FA5C650BFA02" box="[1022,1104,1490,1514]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="0.5">0.5 mm</quantity>
tall, inconspicuous, yellowish white; style
<quantity id="4CA99B25FFB5FFF13020FA7B63ECF9E5" box="[587,695,1525,1549]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="13" pageNumber="14" unit="mm" value="7.75" valueMax="8.0" valueMin="7.5">7.58 mm</quantity>
long, white, widening distally, glabrous; stigma
<quantity id="4CA99B25FFB5FFF136D9FA7B6458F9E5" box="[1202,1283,1525,1549]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="mm" value="0.5">0.5 mm</quantity>
long,
<quantity id="4CA99B25FFB5FFF1372FFA7B64EBF9E5" box="[1348,1456,1525,1549]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="13" pageNumber="14" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.81 mm</quantity>
wide, green, somewhat bilobed. Berry
<quantity id="4CA99B25FFB5FFF13100F9966285F9D8" box="[875,990,1560,1584]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="13" pageNumber="14" unit="cm" value="1.05" valueMax="1.2" valueMin="0.9">0.91.2 cm</quantity>
in diameter, globose, slightly flattened at the apex, green or white when immature, orange to red at maturity, glabrous, not pungent, the pericarp lacking giant cells (endocarp smooth), sclerotic granules 2, polyhedral, yellowish white; fruiting pedicels
<quantity id="4CA99B25FFB5FFF130F3F90E6257F970" box="[664,780,1664,1688]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.6" metricValueMin="2.8" pageId="13" pageNumber="14" unit="cm" value="3.2" valueMax="3.6" valueMin="2.8">2.83.6 cm</quantity>
long, pendent, slightly striate and widened distally; the fruiting calyx persistent, non-accrescent, ca.
<quantity id="4CA99B25FFB5FFF13132F92C62C3F952" box="[857,920,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="14" unit="mm" value="4.0">4 mm</quantity>
diam, discoid, green-purple or green, the reflexed appendages
<quantity id="4CA99B25FFB5FFF130FCF94B6258F935" box="[663,771,1733,1757]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="6.1" metricValueMin="5.0" pageId="13" pageNumber="14" unit="mm" value="5.55" valueMax="6.1" valueMin="5.0">56.1 mm</quantity>
long,
<quantity id="4CA99B25FFB5FFF1312FF94B62F4F935" box="[836,943,1733,1757]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="13" pageNumber="14" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.81 mm</quantity>
wide at base, fleshy and subulate. Seeds ca. 5080 per fruit,
<quantity id="4CA99B25FFB5FFF1303BF96663E7F8E8" box="[592,700,1768,1792]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="13" pageNumber="14" unit="mm" value="2.1" valueMax="2.2" valueMin="2.0">22.2 mm</quantity>
long, ca.
<quantity id="4CA99B25FFB5FFF13174F966622BF8E8" box="[799,880,1768,1792]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="14" unit="mm" value="2.5">2.5 mm</quantity>
wide, somewhat compressed, subreniform or obconic, dark brown, the surface reticulate, cells polygonal in shape, lateral walls straight or slightly sinuate.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB5FFF3305EF8A362EEFE21" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BEE36C0FFB5FFF1305EF8A364A7F880" blockId="13.[533,1536,208,1896]" pageId="13" pageNumber="14">
<emphasis id="B925EAD2FFB5FFF1305EF8A3620AF8AD" bold="true" box="[565,849,1837,1861]" pageId="13" pageNumber="14">Distribution and ecology.</emphasis>
Endemic to a restricted area in northern
<collectingCountry id="F3467650FFB5FFF1376DF8A36463F8AD" box="[1286,1336,1837,1861]" name="Peru" pageId="13" pageNumber="14">Peru</collectingCountry>
(
<collectingRegion id="4995F822FFB5FFF1372DF8A364D8F8AD" box="[1350,1411,1837,1861]" country="Peru" name="Piura" pageId="13" pageNumber="14">Piura</collectingRegion>
,
<figureCitation id="136A2A45FFB5FFF137E6F8A3649AF8AD" box="[1421,1473,1837,1861]" captionStart="Fig 2" captionStartId="6.[533,561,1150,1169]" captionTargetBox="[533,1320,208,1128]" captionTargetId="figure-278@6.[533,1320,208,1129]" captionTargetPageId="6" captionText="Fig 2. Distribution of Capsicum species. https://doi.org/10.1371/journal.pone.0209792.g002" figureDoi="http://doi.org/10.5281/zenodo.12630923" httpUri="https://zenodo.org/record/12630923/files/figure.png" pageId="13" pageNumber="14">Fig 2</figureCitation>
) growing in montane misty rain forests, associated with other
<taxonomicName id="4C514D43FFB5FFF136F6F8DE644AF880" box="[1181,1297,1872,1896]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
shrubs (
<taxonomicName id="4C514D43FFB5FFF13707F8C164A7F880" box="[1388,1532,1871,1896]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B925EAD2FFB5FFF13707F8C16492F88F" box="[1388,1481,1871,1895]" italics="true" pageId="13" pageNumber="14">Solanum</emphasis>
spp.
</taxonomicName>
</paragraph>
<caption id="DF2E6648FFB7FFF3331EFF45645CFF1F" pageId="15" pageNumber="16" startId="15.[373,401,203,222]" targetBox="[374,1529,211,1872]" targetPageId="14" targetType="figure">
<paragraph id="8BEE36C0FFB7FFF3331EFF45645CFF1F" blockId="15.[373,1491,203,248]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF3331EFF456255FF36" bold="true" box="[373,782,203,222]" pageId="15" pageNumber="16">
Fig 7.
<taxonomicName id="4C514D43FFB7FFF333C7FF456255FF36" authority="Barboza &amp; S. Leiva." authorityName="Barboza &amp; S. Leiva." box="[428,782,203,222]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFB7FFF333C7FF456304FF36" bold="true" box="[428,607,203,222]" italics="true" pageId="15" pageNumber="16">Capsicum piuranum</emphasis>
Barboza &amp; S. Leiva.
</taxonomicName>
</emphasis>
(A) Flowering branch. (B) Calyx. (C) Flower. (D) Opened corolla. (E, F, G). Anther, ventral, dorsal and lateral view, respectively. (H) Gynoecium. (I) Fruit. (J) Seed. Drawn by S. Leiva González.
</paragraph>
</caption>
<paragraph id="8BEE36C0FFB7FFF3331EFE806257FEC9" blockId="15.[373,780,270,289]" box="[373,780,270,289]" pageId="15" pageNumber="16">https://doi.org/10.1371/journal.pone.0209792.g007</paragraph>
<paragraph id="8BEE36C0FFB7FFF3307EFEC462EEFE21" blockId="15.[533,1536,329,1879]" pageId="15" pageNumber="16">
and
<taxonomicName id="4C514D43FFB7FFF33028FEC7628AFE8A" authority="(Benth.) Miers" authorityName="Miers" baseAuthorityName="Benth." box="[579,977,329,354]" class="Magnoliopsida" family="Solanaceae" genus="Streptosolen" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="jamesonii">
<emphasis id="B925EAD2FFB7FFF33028FEC76276FE89" box="[579,813,329,353]" italics="true" pageId="15" pageNumber="16">Streptosolen jamesonii</emphasis>
(Benth.) Miers
</taxonomicName>
),
<taxonomicName id="4C514D43FFB7FFF33188FEC4656CFE89" box="[995,1079,330,353]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB7FFF33188FEC4656CFE89" box="[995,1079,330,353]" italics="true" pageId="15" pageNumber="16">Begonia</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C514D43FFB7FFF33635FEC465BCFE8A" box="[1118,1255,330,354]" class="Magnoliopsida" family="Begoniaceae" kingdom="Plantae" order="Cucurbitales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="family">Begoniaceae</taxonomicName>
),
<taxonomicName id="4C514D43FFB7FFF33693FEC764B8FE8A" authority="C. H. Stirt." authorityName="C. H. Stirt." box="[1272,1507,329,354]" class="Magnoliopsida" family="Fabaceae" genus="Otholobium" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB7FFF33693FEC7642CFE89" box="[1272,1399,329,353]" italics="true" pageId="15" pageNumber="16">Otholobium</emphasis>
C.H.Stirt.
</taxonomicName>
(
<taxonomicName id="4C514D43FFB7FFF33076FEE263D8FE6C" box="[541,643,364,388]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
),
<taxonomicName id="4C514D43FFB7FFF330FFFEE26209FE6C" authority="R. Br." authorityName="R. Br." box="[660,850,364,388]" class="Magnoliopsida" family="Acanthaceae" genus="Aphelandra" kingdom="Plantae" order="Lamiales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB7FFF330FFFEE2624BFE6C" box="[660,784,364,388]" italics="true" pageId="15" pageNumber="16">Aphelandra</emphasis>
R. Br.
</taxonomicName>
(
<taxonomicName id="4C514D43FFB7FFF33135FEE262B0FE6C" box="[862,1003,364,388]" class="Magnoliopsida" family="Acanthaceae" kingdom="Plantae" order="Lamiales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="family">Acanthaceae</taxonomicName>
),
<taxonomicName id="4C514D43FFB7FFF33197FEE26513FE6C" box="[1020,1096,364,388]" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB7FFF33197FEE26513FE6C" box="[1020,1096,364,388]" italics="true" pageId="15" pageNumber="16">Juglans</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C514D43FFB7FFF33604FEE265A5FE6C" box="[1135,1278,364,388]" class="Magnoliopsida" family="Juglandaceae" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="family">Juglandaceae</taxonomicName>
) amongst other herbs and shrubs. It grows in margins of forest and near streams, between
<quantity id="4CA99B25FFB7FFF3368CFE01642DFE4F" box="[1255,1398,399,423]" metricMagnitude="3" metricUnit="m" metricValue="2.58" metricValueMax="2.86" metricValueMin="2.3" pageId="15" pageNumber="16" unit="m" value="2580.0" valueMax="2860.0" valueMin="2300.0">23002860 m</quantity>
elevation, in areas of low temperature and rich soils.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB7FFF3305EFE5A6493FE04" box="[565,1480,468,492]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BEE36C0FFB7FFF3305EFE5A6493FE04" blockId="15.[533,1536,329,1879]" box="[565,1480,468,492]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF3305EFE5A63E9FE04" bold="true" box="[565,690,468,492]" pageId="15" pageNumber="16">Phenology.</emphasis>
Flowering from November to May, with a peak of fruiting in MarchMay.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB7FFF3305EFE796587FCE9" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BEE36C0FFB7FFF3305EFE796386FDD9" blockId="15.[533,1536,329,1879]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF3305EFE7963EFFDE7" bold="true" box="[565,692,503,527]" pageId="15" pageNumber="16">Etymology.</emphasis>
The new species is named in allusion to its very restricted habitat in Department
<collectingRegion id="4995F822FFB7FFF33038FD9763D6FDD9" box="[595,653,537,561]" country="Peru" name="Piura" pageId="15" pageNumber="16">Piura</collectingRegion>
(
<collectingCountry id="F3467650FFB7FFF330F1FD97638AFDD9" box="[666,721,537,561]" name="Peru" pageId="15" pageNumber="16">Peru</collectingCountry>
).
</paragraph>
<paragraph id="8BEE36C0FFB7FFF3305EFDB26587FCE9" blockId="15.[533,1536,329,1879]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF3305EFDB262F5FDBC" bold="true" box="[565,942,572,596]" pageId="15" pageNumber="16">Species Conservation Assessment.</emphasis>
According to IUCN criteria (IUCN, 2017),
<taxonomicName id="4C514D43FFB7FFF33715FDB2631DFD9E" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFB7FFF33715FDB264D4FDBB" box="[1406,1423,572,595]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF337F0FDB3631DFD9E" italics="true" pageId="15" pageNumber="16">piuranum</emphasis>
</taxonomicName>
is proposed as Critically Endangered (
<collectionCode id="ED40AE05FFB7FFF331B5FDD1655FFD9F" box="[990,1028,607,631]" country="Costa Rica" lsid="urn:lsid:biocol.org:col:15652" name="Museo Nacional de Costa Rica" pageId="15" pageNumber="16" type="Herbarium">CR</collectionCode>
) species. The extent of occurrence is calculated to be
<quantity id="4CA99B25FFB7FFF33024FD0F63E5FD71" box="[591,702,641,665]" metricMagnitude="4" metricUnit="m" metricValue="1.0195" pageId="15" pageNumber="16" unit="km" value="10.195">10.195 km</quantity>
2 (Criterion B1 &lt;
<quantity id="4CA99B25FFB7FFF33114FD0F628BFD71" box="[895,976,641,665]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="km" value="100.0">100 km</quantity>
2, Critically Endangered), the area of occupancy,
<quantity id="4CA99B25FFB7FFF337BDFD0F6363FD54" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="km" value="8.0">8 km</quantity>
2 (Criterion B2 &lt;
<quantity id="4CA99B25FFB7FFF33092FD2A6265FD54" box="[761,830,676,700]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="km" value="10.0">10 km</quantity>
2, Critically Endangered). The species is known from only three locations (Criterion B2a
<emphasis id="B925EAD2FFB7FFF33170FD49626BFD37" bold="true" box="[795,816,711,735]" pageId="15" pageNumber="16"><EFBFBD></emphasis>
5, Endangered) and the number of mature individuals observed in each subpopulation is
<emphasis id="B925EAD2FFB7FFF33169FD67624CFCE9" bold="true" box="[770,791,745,769]" pageId="15" pageNumber="16"><EFBFBD></emphasis>
50 (Criterion C2a, Critically Endangered).
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB7FFF3305EFC8265E7FBB4" pageId="15" pageNumber="16" type="description">
<paragraph id="8BEE36C0FFB7FFF3305EFC8264AEFBFF" blockId="15.[533,1536,329,1879]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF3305EFC8263F5FCCC" bold="true" box="[565,686,780,804]" pageId="15" pageNumber="16">Karyology.</emphasis>
A somatic chromosome number of 2n = 2x = 26 was found in this species. The karyotype comprises
<quantity id="4CA99B25FFB7FFF33084FCA1624CFCAF" box="[751,791,815,839]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="16" unit="m" value="9.0">9 m</quantity>
pairs of rather similar length (19), 3 sm pairs (1012), and one st pair (13) (
<figureCitation id="136A2A45FFB7FFF33025FCDF63DBFC81" box="[590,640,849,873]" captionStart="Fig 9" captionStartId="17.[533,561,1378,1397]" captionTargetBox="[533,1324,203,1356]" captionTargetId="figure-173@17.[533,1324,203,1356]" captionTargetPageId="17" captionText="Fig 9. Somatic metaphase chromosomes and ideogram of Capsicum piuranum. (A) Methaphase chromosomes. (B) Ideogram. Solid black blocks or dots denote CMA+/DAPI- (NOR) or CMA+/DAPIo (terminal and intercalary) heterochromatic bands. The NOR is indicated as a separate block. https://doi.org/10.1371/journal.pone.0209792.g009" figureDoi="http://doi.org/10.5281/zenodo.12630933" httpUri="https://zenodo.org/record/12630933/files/figure.png" pageId="15" pageNumber="16">Fig 9</figureCitation>
, S
<figureCitation id="136A2A45FFB7FFF330F0FCDF63FEFC81" box="[667,677,849,873]" captionStart="Fig 1" captionStartId="5.[454,482,203,222]" captionTargetId="figure-1@4.[454,1536,208,1872]" captionTargetPageId="4" captionText="Fig 1. Capsicum benoistii Hunz. ex Barboza. (A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury." pageId="15" pageNumber="16">1</figureCitation>
Table). One pair is satellited (10 sm). As in
<taxonomicName id="4C514D43FFB7FFF33608FCDC65A9FC81" box="[1123,1266,849,873]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB7FFF33608FCDC6528FC81" box="[1123,1139,850,873]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF33614FCDF65A9FC81" box="[1151,1266,849,873]" italics="true" pageId="15" pageNumber="16">longifolium</emphasis>
</taxonomicName>
, this species bears
<specimenCount id="9D57FD49FFB7FFF337DAFCDF6311FC64" count="2" pageId="15" pageNumber="16" type="generic" typeStatus="types">two types</specimenCount>
of constitutive heterochromatin, GC-rich heterochromatin (
<collectionCode id="ED40AE05FFB7FFF336DDFCFA65A8FC64" box="[1206,1267,884,908]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12899" name="Crayford Manor House Adult Education Centre" pageId="15" pageNumber="16" type="Herbarium">CMA</collectionCode>
+/DAPI-) located in the large heterochromatic band associated to the NOR in pair 10, and moderately GC-rich heterochromatin (
<collectionCode id="ED40AE05FFB7FFF330E5FC376390FC39" box="[654,715,953,977]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12899" name="Crayford Manor House Adult Education Centre" pageId="15" pageNumber="16" type="Herbarium">CMA</collectionCode>
+/DAPIo), located in the small terminal bands and in the intercalary band on the long arm of pair 3. The fluorescent banding pattern is quite simple and very similar to
<taxonomicName id="4C514D43FFB7FFF337C5FC526306FBFF" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB7FFF337C5FC5264E5FC1B" box="[1454,1470,988,1011]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF337A2FC526306FBFF" italics="true" pageId="15" pageNumber="16">longifolium</emphasis>
</taxonomicName>
, except for the presence of 3 small bands that are not seen in that species (
<figureCitation id="136A2A45FFB7FFF33739FC7164DFFBFF" box="[1362,1412,1023,1047]" captionStart="Fig 9" captionStartId="17.[533,561,1378,1397]" captionTargetBox="[533,1324,203,1356]" captionTargetId="figure-173@17.[533,1324,203,1356]" captionTargetPageId="17" captionText="Fig 9. Somatic metaphase chromosomes and ideogram of Capsicum piuranum. (A) Methaphase chromosomes. (B) Ideogram. Solid black blocks or dots denote CMA+/DAPI- (NOR) or CMA+/DAPIo (terminal and intercalary) heterochromatic bands. The NOR is indicated as a separate block. https://doi.org/10.1371/journal.pone.0209792.g009" figureDoi="http://doi.org/10.5281/zenodo.12630933" httpUri="https://zenodo.org/record/12630933/files/figure.png" pageId="15" pageNumber="16">Fig 9</figureCitation>
, S
<figureCitation id="136A2A45FFB7FFF337F7FC7164F2FBFF" box="[1436,1449,1023,1047]" captionStart="Fig 4" captionStartId="10.[333,361,203,222]" captionTargetId="figure-1@9.[333,1536,208,1872]" captionTargetPageId="9" captionText="Fig 4. Capsicum longifolium Barboza &amp; S. Leiva. (A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza." pageId="15" pageNumber="16">4</figureCitation>
Table).
</paragraph>
<paragraph id="8BEE36C0FFB7FFF3305EFBAF65E7FBB4" blockId="15.[533,1536,329,1879]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF3305EFBAF63C5FBD1" bold="true" box="[565,670,1057,1081]" pageId="15" pageNumber="16">Affinities</emphasis>
.
<taxonomicName id="4C514D43FFB7FFF330C1FBAC62D9FBD1" box="[682,898,1058,1081]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFB7FFF330C1FBAC62D9FBD1" box="[682,898,1058,1081]" italics="true" pageId="15" pageNumber="16">Capsicum piuranum</emphasis>
</taxonomicName>
is resolved within the Andean clade, strongly supported as sister to the
<taxonomicName id="4C514D43FFB7FFF330FEFBCA6260FBB4" box="[661,827,1092,1116]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="lycianthoides">
<emphasis id="B925EAD2FFB7FFF330FEFBCA63FDFBB3" box="[661,678,1092,1115]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF330D9FBCA6260FBB4" box="[690,827,1092,1116]" italics="true" pageId="15" pageNumber="16">lycianthoides</emphasis>
</taxonomicName>
<emphasis id="B925EAD2FFB7FFF33150FBCA621FFBB4" box="[827,836,1092,1116]" italics="true" pageId="15" pageNumber="16">-</emphasis>
<taxonomicName id="4C514D43FFB7FFF3312FFBCA62B7FBB4" box="[836,1004,1092,1116]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="geminifolium">
<emphasis id="B925EAD2FFB7FFF3312FFBCA620EFBB3" box="[836,853,1092,1115]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF3310AFBCA62B7FBB4" box="[865,1004,1092,1116]" italics="true" pageId="15" pageNumber="16">geminifolium</emphasis>
</taxonomicName>
assemblage (
<figureCitation id="136A2A45FFB7FFF33613FBCA65F7FBB4" box="[1144,1196,1092,1116]" captionStart="Fig 6" captionStartId="12.[133,161,1434,1453]" captionTargetBox="[134,1535,208,1411]" captionTargetId="figure-188@12.[133,1536,208,1412]" captionTargetPageId="12" captionText="Fig 6. Bayesian majority-rule consensus tree of Capsicum. Posterior probabilities values indicated by each branch. New species are highlighted in bold-colored letters and the clade to which they belong is indicated. https://doi.org/10.1371/journal.pone.0209792.g006" figureDoi="http://doi.org/10.5281/zenodo.12630927" httpUri="https://zenodo.org/record/12630927/files/figure.png" pageId="15" pageNumber="16">Fig 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB7FFF3305EFBE964ACF8BF" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BEE36C0FFB7FFF3305EFBE96481FA5F" blockId="15.[533,1536,329,1879]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF3305EFBE96561FB97" bold="true" box="[565,1082,1127,1151]" pageId="15" pageNumber="16">
Additional specimens examined.
<materialsCitation id="3B393C9DFFB7FFF331CEFBE965CEFB0F" collectingDate="2015-11-01" collectingDateMax="2017-05-08" collectingDateMin="2015-11-01" collectionCode="CORD, USM, HAO, HUSA" country="Peru" county="Canchaque-Huancabamba" elevation="2343" location="Cuello del Indio" municipality="Huancabamba" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Piura">
<collectingCountry id="F3467650FFB7FFF331CEFBE962B0FB97" box="[933,1003,1127,1151]" name="Peru" pageId="15" pageNumber="16">PERU</collectingCountry>
.
<collectingRegion id="4995F822FFB7FFF3319DFBE9656DFB97" box="[1014,1078,1127,1151]" country="Peru" name="Piura" pageId="15" pageNumber="16">Piura</collectingRegion>
: Prov. Huancabamba, carretera
<collectingCounty id="628F4E4CFFB7FFF337E1FBE963B8FB49" pageId="15" pageNumber="16">Canchaque-Huancabamba</collectingCounty>
, km 9899 (ca. 20 km de Canchaque rumbo a
<collectingMunicipality id="6B8AACBAFFB7FFF336A6FB076437FB49" box="[1229,1388,1161,1185]" pageId="15" pageNumber="16">Huancabamba</collectingMunicipality>
), subiendo a
<location id="8E8E601BFFB7FFF3307EFB226398FB2C" LSID="urn:lsid:plazi:treatment:03F887D6FFB5FFEE305EFED4620DFE15:8E8E601BFFB7FFF3307EFB226398FB2C" box="[533,707,1196,1220]" country="Peru" county="Canchaque-Huancabamba" municipality="Huancabamba" name="Cuello del Indio" pageId="15" pageNumber="16" stateProvince="Piura">Cuello del Indio</location>
, 05˚2243”-05˚2244”S, 79˚3334”-79˚3337”W, 23412346 m,
<date id="FFEF1000FFB7FFF3373FFB226494FB2C" box="[1364,1487,1196,1220]" pageId="15" pageNumber="16" value="2017-05-08">
<collectingDate id="EFABE9E8FFB7FFF3373FFB226494FB2C" box="[1364,1487,1196,1220]" pageId="15" pageNumber="16" value="2017-05-08">8 May 2017</collectingDate>
</date>
(fl, fr),
<collectorName id="26A45316FFB7FFF33057FB416390FB0E" box="[572,715,1230,1255]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF33057FB416315FB0E" box="[572,590,1231,1254]" italics="true" pageId="15" pageNumber="16">G</emphasis>
.
<emphasis id="B925EAD2FFB7FFF33031FB416332FB0E" box="[602,617,1231,1254]" italics="true" pageId="15" pageNumber="16">E</emphasis>
.
<emphasis id="B925EAD2FFB7FFF3301FFB4063A3FB0E" box="[628,760,1230,1254]" italics="true" pageId="15" pageNumber="16">
Barboza &amp;
<collectorName id="26A45316FFB7FFF33087FB4162F8FB0E" box="[748,931,1230,1255]" pageId="15" pageNumber="16">
S.
<emphasis id="B925EAD2FFB7FFF3316FFB416575FB0F" box="[772,1070,1230,1255]" italics="true" pageId="15" pageNumber="16">Leiva González 4841 &amp; 4842</emphasis>
</collectorName>
</emphasis>
</collectorName>
(
<collectionCode id="ED40AE05FFB7FFF33650FB4165D1FB0F" box="[1083,1162,1231,1255]" country="Argentina" lsid="urn:lsid:biocol.org:col:15636" name="Universidad Nacional de Córdoba" pageId="15" pageNumber="16" type="Herbarium">CORD</collectionCode>
);
</materialsCitation>
</emphasis>
<materialsCitation id="3B393C9DFFB7FFF336F7FB4163BDFAC4" collectingDate="2015-11-01" collectingDateMax="2017-05-08" collectingDateMin="2015-11-01" collectionCode="CORD, USM, HAO, HUSA" country="Peru" county="Canchaque-Huancabamba" elevation="2343" location="Cuello del Indio" municipality="Huancabamba" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Piura">
carretera Huancabamba-Canchaque, a una hora del Abra Cruz Blanca, 05˚2225&quot;S, 79˚3402&quot;W, 2860 m,
<date id="FFEF1000FFB7FFF33764FB7C64CAFAE2" box="[1295,1425,1266,1290]" pageId="15" pageNumber="16" value="2017-04-13">
<collectingDate id="EFABE9E8FFB7FFF33764FB7C64CAFAE2" box="[1295,1425,1266,1290]" pageId="15" pageNumber="16" value="2017-04-13">13 Apr 2017</collectingDate>
</date>
(fl, fr),
<collectorName id="26A45316FFB7FFF3378AFB7C630CFAC3" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF3378AFB7C64A3FAE1" box="[1505,1528,1266,1289]" italics="true" pageId="15" pageNumber="16">M</emphasis>
.
<emphasis id="B925EAD2FFB7FFF3307EFA9A63CBFAC4" box="[533,656,1300,1324]" italics="true" pageId="15" pageNumber="16">Cueva 2912</emphasis>
</collectorName>
(
<collectionCode id="ED40AE05FFB7FFF330F6FA9A6381FAC4" box="[669,730,1300,1324]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="15" pageNumber="16">USM</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B393C9DFFB7FFF33087FA9A6552FA9A" collectingDate="2015-11-01" collectingDateMax="2017-05-08" collectingDateMin="2015-11-01" collectionCode="CORD, USM, HAO, HUSA" country="Peru" county="Canchaque-Huancabamba" elevation="2343" location="Cuello del Indio" municipality="Huancabamba" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Piura">
distrito Canchaque, km 9899 (ruta Canchaque-Huancabamba), 5˚ 2244.1”-5˚ 2242.7”S, 79˚ 3337.2”-79˚ 3334.1” W, 23412346 m,
<date id="FFEF1000FFB7FFF336A5FAB96412FAA7" box="[1230,1353,1335,1359]" pageId="15" pageNumber="16" value="2017-05-08">
<collectingDate id="EFABE9E8FFB7FFF336A5FAB96412FAA7" box="[1230,1353,1335,1359]" pageId="15" pageNumber="16" value="2017-05-08">8 May 2017</collectingDate>
</date>
(fl, fr),
<collectorName id="26A45316FFB7FFF337F3FAB9632DFA99" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF337F3FAB964FFFAA6" box="[1432,1444,1335,1358]" italics="true" pageId="15" pageNumber="16">S</emphasis>
.
<emphasis id="B925EAD2FFB7FFF337DBFAB963F3FA99" italics="true" pageId="15" pageNumber="16">
Leiva González &amp;
<collectorName id="26A45316FFB7FFF330FDFAD4627DFA99" box="[662,806,1369,1394]" pageId="15" pageNumber="16">
G.
<emphasis id="B925EAD2FFB7FFF330DFFAD46398FA99" box="[692,707,1370,1393]" italics="true" pageId="15" pageNumber="16">E</emphasis>
.
<emphasis id="B925EAD2FFB7FFF330A4FAD762EAFA99" box="[719,945,1369,1393]" italics="true" pageId="15" pageNumber="16">Barboza 6561 &amp; 6562</emphasis>
</collectorName>
</emphasis>
</collectorName>
(
<collectionCode id="ED40AE05FFB7FFF331D5FAD462A6FA9A" box="[958,1021,1370,1394]" country="Peru" lsid="urn:lsid:biocol.org:col:15351" name="Universidad Privada Antenor Orrego" pageId="15" pageNumber="16" type="Herbarium">HAO</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B393C9DFFB7FFF33664FAD46436FA7C" collectingDate="2015-11-01" collectingDateMax="2017-05-08" collectingDateMin="2015-11-01" collectionCode="CORD, USM, HAO, HUSA" country="Peru" county="Canchaque-Huancabamba" elevation="2343" location="Cuello del Indio" municipality="Huancabamba" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Piura">
Prov. Morropon: Chalaco, Bosque Mijal, 05˚ 0351.1&quot;S, 79˚4325.9&quot;W, 2800 m,
<date id="FFEF1000FFB7FFF33112FAF262A9FA7C" box="[889,1010,1404,1428]" pageId="15" pageNumber="16" value="2015-11-01">
<collectingDate id="EFABE9E8FFB7FFF33112FAF262A9FA7C" box="[889,1010,1404,1428]" pageId="15" pageNumber="16" value="2015-11-01">1 Nov 2015</collectingDate>
</date>
(fl, fr),
<collectorName id="26A45316FFB7FFF3362AFAF365FDFA7B" box="[1089,1190,1404,1428]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF3362AFAF36503FA7C" box="[1089,1112,1405,1428]" italics="true" pageId="15" pageNumber="16">M</emphasis>
.
<emphasis id="B925EAD2FFB7FFF3360FFAF26583FA7C" box="[1124,1240,1404,1428]" italics="true" pageId="15" pageNumber="16">Cueva et al</emphasis>
</collectorName>
.
<emphasis id="B925EAD2FFB7FFF3368FFAF2644CFA7C" box="[1252,1303,1404,1428]" italics="true" pageId="15" pageNumber="16">2655</emphasis>
(
<collectionCode id="ED40AE05FFB7FFF3374FFAF2643AFA7C" box="[1316,1377,1404,1428]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="15" pageNumber="16">USM</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B393C9DFFB7FFF33718FAF2648DFA5F" collectingDate="2015-11-01" collectingDateMax="2017-05-08" collectingDateMin="2015-11-01" collectionCode="CORD, USM, HAO, HUSA" country="Peru" county="Canchaque-Huancabamba" elevation="2343" location="Cuello del Indio" municipality="Huancabamba" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Piura">
same locality and date,
<collectorName id="26A45316FFB7FFF33017FA1163BAFA5E" box="[636,737,1439,1463]" pageId="15" pageNumber="16">
<emphasis id="B925EAD2FFB7FFF33017FA1163C8FA5E" box="[636,659,1439,1462]" italics="true" pageId="15" pageNumber="16">M</emphasis>
.
<emphasis id="B925EAD2FFB7FFF330F4FA116248FA5E" box="[671,787,1438,1462]" italics="true" pageId="15" pageNumber="16">Cueva et al</emphasis>
</collectorName>
.
<emphasis id="B925EAD2FFB7FFF33174FA10620AFA5E" box="[799,849,1438,1462]" italics="true" pageId="15" pageNumber="16">2656</emphasis>
,
<emphasis id="B925EAD2FFB7FFF33135FA1062B8FA5E" box="[862,995,1438,1462]" italics="true" pageId="15" pageNumber="16">2657 &amp; 2658</emphasis>
(
<collectionCode id="ED40AE05FFB7FFF3319BFA116577FA5F" box="[1008,1068,1439,1463]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="15" pageNumber="16">USM</collectionCode>
, duplicates to be sent to CORD,
<collectionCode id="ED40AE05FFB7FFF337E8FA116494FA5F" box="[1411,1487,1439,1463]" country="Peru" lsid="urn:lsid:biocol.org:col:14917" name="Universidad Nacional de San Agustín de Arequipa" pageId="15" pageNumber="16" type="Herbarium">HUSA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36C0FFB7FFF3305EFA4C64ACF8BF" blockId="15.[533,1536,329,1879]" pageId="15" pageNumber="16">
<taxonomicName id="4C514D43FFB7FFF3305EFA4C6256FA31" box="[565,781,1474,1497]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFB7FFF3305EFA4C6256FA31" box="[565,781,1474,1497]" italics="true" pageId="15" pageNumber="16">Capsicum piuranum</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName id="4C514D43FFB7FFF336E8FA4C6402FA32" authority="M. Nee" authorityName="M. Nee" box="[1155,1369,1473,1498]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="caballeroi">
<emphasis id="B925EAD2FFB7FFF336E8FA4C65CFFA31" box="[1155,1172,1474,1497]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF336CBFA4F645DFA31" box="[1184,1286,1473,1497]" italics="true" pageId="15" pageNumber="16">caballeroi</emphasis>
M. Nee
</taxonomicName>
of the Bolivian yungas (
<collectingRegion id="4995F822FFB7FFF33006FA6A63BFFA14" box="[621,740,1508,1532]" country="Bolivia" name="Santa Cruz" pageId="15" pageNumber="16">Santa Cruz</collectingRegion>
and
<collectingRegion id="4995F822FFB7FFF33173FA6A62FDFA14" box="[792,934,1508,1532]" country="Bolivia" name="Cochabamba" pageId="15" pageNumber="16">Cochabamba</collectingRegion>
) based on their campanulate yellow corollas. However, these species can be distinguished in the calyx color, the calyx appendages (number, size, and shape), the position of the corolla lobes at anthesis, the fruit size, shape, color and pungency, the presence of stone cells, and the seed color.
<taxonomicName id="4C514D43FFB7FFF33194F9C2658DF98C" box="[1023,1238,1612,1636]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFB7FFF33194F9C2658DF98C" box="[1023,1238,1612,1636]" italics="true" pageId="15" pageNumber="16">Capsicum piuranum</emphasis>
</taxonomicName>
has a purple or greenish purple calyx with 5 equal subulate appendages (
<figureCitation id="136A2A45FFB7FFF33666F9E165FEF96F" box="[1037,1189,1647,1671]" captionStart="Fig 8" captionStartId="16.[527,555,1702,1721]" captionTargetBox="[527,1536,208,1680]" captionTargetId="figure-3@16.[527,1536,208,1680]" captionTargetPageId="16" captionText="Fig 8. Capsicum piuranum Barboza &amp; S. Leiva. (A) Plant. (B) Leaves, abaxial surface. (C) Fruiting branch. (D) Flower bud. (E) Flower and immature fruit. (F) Mature fruit. (G) Fruit, transverse section, showing placenta and seeds. (H) Fruit transverse section, showing a stone cell at the apex (arrow). Photos by S. Leiva González and G. E. Barboza. https://doi.org/10.1371/journal.pone.0209792.g008" figureDoi="http://doi.org/10.5281/zenodo.12630929" httpUri="https://zenodo.org/record/12630929/files/figure.png" pageId="15" pageNumber="16">Fig 8D and 8E</figureCitation>
), while
<taxonomicName id="4C514D43FFB7FFF33693F9E16420F96E" box="[1272,1403,1646,1671]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="caballeroi">
<emphasis id="B925EAD2FFB7FFF33693F9E16452F96E" box="[1272,1289,1647,1670]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF3377EF9E06420F96E" box="[1301,1403,1646,1670]" italics="true" pageId="15" pageNumber="16">caballeroi</emphasis>
</taxonomicName>
has green calyx with 10 unequal linear appendages. Corolla lobes are erect compared to those of
<taxonomicName id="4C514D43FFB7FFF337C8F91C6318F924" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="caballeroi">
<emphasis id="B925EAD2FFB7FFF337C8F91C64EFF941" box="[1443,1460,1682,1705]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF337ABF91F6318F924" italics="true" pageId="15" pageNumber="16">caballeroi</emphasis>
</taxonomicName>
which are recurved. Mature fruits are smaller (up to
<quantity id="4CA99B25FFB7FFF33618F93A659EF924" box="[1139,1221,1716,1740]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="16" unit="mm" value="1.2">1.2 mm</quantity>
diam), globose, orange and not pungent in
<taxonomicName id="4C514D43FFB7FFF330D1F959621BF906" box="[698,832,1751,1775]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFB7FFF330D1F9596390F906" box="[698,715,1751,1774]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF330BDF959621BF906" box="[726,832,1751,1774]" italics="true" pageId="15" pageNumber="16">piuranum</emphasis>
</taxonomicName>
but are larger (up to
<quantity id="4CA99B25FFB7FFF3364BF959652AF907" box="[1056,1137,1751,1775]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="16" unit="mm" value="1.6">1.6 mm</quantity>
diam), globose-depressed to globose, bright red and pungent in
<taxonomicName id="4C514D43FFB7FFF33145F97462EBF8F9" box="[814,944,1785,1809]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="caballeroi">
<emphasis id="B925EAD2FFB7FFF33145F9746264F8F9" box="[814,831,1786,1809]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF33121F97762EBF8F9" box="[842,944,1785,1809]" italics="true" pageId="15" pageNumber="16">caballeroi</emphasis>
</taxonomicName>
.
<taxonomicName id="4C514D43FFB7FFF331D7F97465CFF8F9" box="[956,1172,1786,1809]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFB7FFF331D7F97465CFF8F9" box="[956,1172,1786,1809]" italics="true" pageId="15" pageNumber="16">Capsicum piuranum</emphasis>
</taxonomicName>
has two stone cells (
<figureCitation id="136A2A45FFB7FFF33700F97764EFF8F9" box="[1387,1460,1785,1809]" captionStart="Fig 8" captionStartId="16.[527,555,1702,1721]" captionTargetBox="[527,1536,208,1680]" captionTargetId="figure-3@16.[527,1536,208,1680]" captionTargetPageId="16" captionText="Fig 8. Capsicum piuranum Barboza &amp; S. Leiva. (A) Plant. (B) Leaves, abaxial surface. (C) Fruiting branch. (D) Flower bud. (E) Flower and immature fruit. (F) Mature fruit. (G) Fruit, transverse section, showing placenta and seeds. (H) Fruit transverse section, showing a stone cell at the apex (arrow). Photos by S. Leiva González and G. E. Barboza. https://doi.org/10.1371/journal.pone.0209792.g008" figureDoi="http://doi.org/10.5281/zenodo.12630929" httpUri="https://zenodo.org/record/12630929/files/figure.png" pageId="15" pageNumber="16">Fig 8H</figureCitation>
) and dark brown smaller seeds (
<quantity id="4CA99B25FFB7FFF3315BF89262C5F8DC" box="[816,926,1820,1844]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="15" pageNumber="16" unit="mm" value="2.1" valueMax="2.2" valueMin="2.0">22.2 mm</quantity>
long, ca.
<quantity id="4CA99B25FFB7FFF3366AF8926509F8DC" box="[1025,1106,1820,1844]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="16" unit="mm" value="2.5">2.5 mm</quantity>
wide) while
<taxonomicName id="4C514D43FFB7FFF336B3F8926400F8DC" box="[1240,1371,1820,1844]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="caballeroi">
<emphasis id="B925EAD2FFB7FFF336B3F89265B2F8DB" box="[1240,1257,1820,1843]" italics="true" pageId="15" pageNumber="16">C</emphasis>
.
<emphasis id="B925EAD2FFB7FFF3369EF8926400F8DC" box="[1269,1371,1820,1844]" italics="true" pageId="15" pageNumber="16">caballeroi</emphasis>
</taxonomicName>
lacks of stone cells and the seeds are pale yellow or light brown and larger (
<quantity id="4CA99B25FFB7FFF336FEF8B16458F8BF" box="[1173,1283,1855,1879]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" metricValueMax="4.0" metricValueMin="3.2" pageId="15" pageNumber="16" unit="mm" value="3.6" valueMax="4.0" valueMin="3.2">3.24 mm</quantity>
long,
<quantity id="4CA99B25FFB7FFF33728F8B164F4F8BF" box="[1347,1455,1855,1879]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="5.0" metricValueMin="3.8" pageId="15" pageNumber="16" unit="mm" value="4.4" valueMax="5.0" valueMin="3.8">3.85 mm</quantity>
wide).
</paragraph>
</subSubSection>
<caption id="DF2E6648FFA8FFEC3064F92862FDF8FD" ID-DOI="http://doi.org/10.5281/zenodo.12630929" ID-Zenodo-Dep="12630929" httpUri="https://zenodo.org/record/12630929/files/figure.png" pageId="16" pageNumber="17" startId="16.[527,555,1702,1721]" targetBox="[527,1536,208,1680]" targetPageId="16" targetType="figure">
<paragraph id="8BEE36C0FFA8FFEC3064F9286499F903" blockId="16.[527,1528,1701,1813]" pageId="16" pageNumber="17">
<emphasis id="B925EAD2FFA8FFEC3064F92862F2F951" bold="true" box="[527,937,1701,1721]" pageId="16" pageNumber="17">
Fig 8.
<taxonomicName id="4C514D43FFA8FFEC302DF92B62F2F951" authority="Barboza &amp; S. Leiva." authorityName="Barboza &amp; S. Leiva." box="[582,937,1701,1721]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFA8FFEC302DF92B63A2F950" bold="true" box="[582,761,1701,1720]" italics="true" pageId="16" pageNumber="17">Capsicum piuranum</emphasis>
Barboza &amp; S. Leiva.
</taxonomicName>
</emphasis>
(A) Plant. (B) Leaves, abaxial surface. (C) Fruiting branch. (D) Flower bud. (E) Flower and immature fruit. (F) Mature fruit. (G) Fruit, transverse section, showing placenta and seeds. (H) Fruit transverse section, showing a stone cell at the apex (arrow). Photos by S. Leiva González and G. E. Barboza.
</paragraph>
<paragraph id="8BEE36C0FFA8FFEC3064F88C62FDF8FD" blockId="16.[527,1528,1701,1813]" box="[527,934,1794,1813]" pageId="16" pageNumber="17">https://doi.org/10.1371/journal.pone.0209792.g008</paragraph>
</caption>
<caption id="DF2E6648FFA9FFED307EFAEC62F6FA39" ID-DOI="http://doi.org/10.5281/zenodo.12630933" ID-Zenodo-Dep="12630933" httpUri="https://zenodo.org/record/12630933/files/figure.png" pageId="17" pageNumber="18" startId="17.[533,561,1378,1397]" targetBox="[533,1324,203,1356]" targetPageId="17" targetType="figure">
<paragraph id="8BEE36C0FFA9FFED307EFAEC651AFA40" blockId="17.[533,1527,1378,1489]" pageId="17" pageNumber="18">
<emphasis id="B925EAD2FFA9FFED307EFAEC6592FA9D" bold="true" box="[533,1225,1378,1397]" pageId="17" pageNumber="18">
Fig 9. Somatic metaphase chromosomes and ideogram of
<taxonomicName id="4C514D43FFA9FFED3679FAEC659EFA9D" box="[1042,1221,1378,1397]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFA9FFED3679FAEC659EFA9D" bold="true" box="[1042,1221,1378,1397]" italics="true" pageId="17" pageNumber="18">Capsicum piuranum</emphasis>
</taxonomicName>
.
</emphasis>
(A) Methaphase chromosomes. (B) Ideogram. Solid black blocks or dots denote CMA+/DAPI- (NOR) or CMA+/DAPIo (terminal and intercalary) heterochromatic bands. The NOR is indicated as a separate block.
</paragraph>
<paragraph id="8BEE36C0FFA9FFED307EFA3062F6FA39" blockId="17.[533,1527,1378,1489]" box="[533,941,1470,1489]" pageId="17" pageNumber="18">https://doi.org/10.1371/journal.pone.0209792.g009</paragraph>
</caption>
<subSubSection id="C34B654BFFA9FFEE305EFA7D620DFE15" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="8BEE36C0FFA9FFED305EFA7D64F4F88E" blockId="17.[533,1532,1523,1894]" pageId="17" pageNumber="18">
<taxonomicName id="4C514D43FFA9FFED305EFA7D6256F9E3" box="[565,781,1523,1547]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFA9FFED305EFA7D6256F9E3" box="[565,781,1523,1547]" italics="true" pageId="17" pageNumber="18">Capsicum piuranum</emphasis>
</taxonomicName>
is sympatric with other two Andean species,
<taxonomicName id="4C514D43FFA9FFED3681FA7D64C9F9E3" box="[1258,1426,1523,1547]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="geminifolium">
<emphasis id="B925EAD2FFA9FFED3681FA7D65A0F9E2" box="[1258,1275,1523,1546]" italics="true" pageId="17" pageNumber="18">C</emphasis>
.
<emphasis id="B925EAD2FFA9FFED376CFA7D64C9F9E3" box="[1287,1426,1523,1547]" italics="true" pageId="17" pageNumber="18">geminifolium</emphasis>
</taxonomicName>
(Dam- mer) Hunz. and
<taxonomicName id="4C514D43FFA9FFED30AEF998622DF9C6" box="[709,886,1558,1582]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="rhomboideum">
<emphasis id="B925EAD2FFA9FFED30AEF998638DF9C5" box="[709,726,1558,1581]" italics="true" pageId="17" pageNumber="18">C</emphasis>
.
<emphasis id="B925EAD2FFA9FFED3089F998622DF9C6" box="[738,886,1558,1582]" italics="true" pageId="17" pageNumber="18">rhomboideum</emphasis>
</taxonomicName>
, both of which have also yellow corollas and non-pungent fruits, but a moderate to dense pubescence on stems and leaves.
<taxonomicName id="4C514D43FFA9FFED36D0F9B764E8F9B8" box="[1211,1459,1592,1616]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="geminifolium">
<emphasis id="B925EAD2FFA9FFED36D0F9B764E8F9B8" box="[1211,1459,1592,1616]" italics="true" pageId="17" pageNumber="18">Capsicum geminifolium</emphasis>
</taxonomicName>
differs in having longer calyx appendages (
<quantity id="4CA99B25FFA9FFED31E5F9D562A0F99B" box="[910,1019,1627,1651]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="6.5" metricValueMin="3.0" pageId="17" pageNumber="18" unit="mm" value="4.75" valueMax="6.5" valueMin="3.0">36.5 mm</quantity>
long) compared to
<taxonomicName id="4C514D43FFA9FFED36A7F9D56409F99B" box="[1228,1362,1627,1651]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFA9FFED36A7F9D56586F99A" box="[1228,1245,1627,1650]" italics="true" pageId="17" pageNumber="18">C</emphasis>
.
<emphasis id="B925EAD2FFA9FFED3683F9D26409F99B" box="[1256,1362,1628,1651]" italics="true" pageId="17" pageNumber="18">piuranum</emphasis>
</taxonomicName>
(
<quantity id="4CA99B25FFA9FFED3734F9D56496F99B" box="[1375,1485,1627,1651]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="17" pageNumber="18" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.53 mm</quantity>
long) and funnel-shaped generally purple spotted yellow corollas (tubular-campanulate and pure yellow in
<taxonomicName id="4C514D43FFA9FFED30D9F92F6262F950" box="[690,825,1697,1721]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFA9FFED30D9F92F6398F950" box="[690,707,1697,1720]" italics="true" pageId="17" pageNumber="18">C</emphasis>
.
<emphasis id="B925EAD2FFA9FFED30A4F92F6262F950" box="[719,825,1697,1720]" italics="true" pageId="17" pageNumber="18">piuranum</emphasis>
</taxonomicName>
,
<figureCitation id="136A2A45FFA9FFED312FF92F62D3F951" box="[836,904,1697,1721]" captionStart="Fig 8" captionStartId="16.[527,555,1702,1721]" captionTargetBox="[527,1536,208,1680]" captionTargetId="figure-3@16.[527,1536,208,1680]" captionTargetPageId="16" captionText="Fig 8. Capsicum piuranum Barboza &amp; S. Leiva. (A) Plant. (B) Leaves, abaxial surface. (C) Fruiting branch. (D) Flower bud. (E) Flower and immature fruit. (F) Mature fruit. (G) Fruit, transverse section, showing placenta and seeds. (H) Fruit transverse section, showing a stone cell at the apex (arrow). Photos by S. Leiva González and G. E. Barboza. https://doi.org/10.1371/journal.pone.0209792.g008" figureDoi="http://doi.org/10.5281/zenodo.12630929" httpUri="https://zenodo.org/record/12630929/files/figure.png" pageId="17" pageNumber="18">Fig 8E</figureCitation>
).
<taxonomicName id="4C514D43FFA9FFED31F6F92F65C5F950" box="[925,1182,1696,1720]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="rhomboideum">
<emphasis id="B925EAD2FFA9FFED31F6F92F65C5F950" box="[925,1182,1696,1720]" italics="true" pageId="17" pageNumber="18">Capsicum rhomboideum</emphasis>
</taxonomicName>
has ovate or rhomboid-ovate leaves, up to 12 axillary flowers, campanulate-rotate smaller corollas (
<quantity id="4CA99B25FFA9FFED369AF94D6428F933" box="[1265,1395,1731,1755]" metricMagnitude="-3" metricUnit="m" metricValue="7.749999999999999" metricValueMax="9.5" metricValueMin="6.0" pageId="17" pageNumber="18" unit="cm" value="0.7749999999999999" valueMax="0.95" valueMin="0.6">0.60.95 cm</quantity>
long), and smaller (up to
<quantity id="4CA99B25FFA9FFED30C4F96863AEF916" box="[687,757,1766,1790]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="18" unit="cm" value="0.9">0.9 cm</quantity>
diam) bright red to blackish red fruits in contrast to
<taxonomicName id="4C514D43FFA9FFED3748F96864F1F915" box="[1315,1450,1766,1790]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFA9FFED3748F968646FF915" box="[1315,1332,1766,1789]" italics="true" pageId="17" pageNumber="18">C</emphasis>
.
<emphasis id="B925EAD2FFA9FFED372BF96864F1F915" box="[1344,1450,1766,1789]" italics="true" pageId="17" pageNumber="18">piuranum</emphasis>
</taxonomicName>
where leaves are elliptic or narrowly elliptic (sometimes the minor leaves are ovate,
<figureCitation id="136A2A45FFA9FFED372BF887648EF8C9" box="[1344,1493,1801,1825]" captionStart="Fig 8" captionStartId="16.[527,555,1702,1721]" captionTargetBox="[527,1536,208,1680]" captionTargetId="figure-3@16.[527,1536,208,1680]" captionTargetPageId="16" captionText="Fig 8. Capsicum piuranum Barboza &amp; S. Leiva. (A) Plant. (B) Leaves, abaxial surface. (C) Fruiting branch. (D) Flower bud. (E) Flower and immature fruit. (F) Mature fruit. (G) Fruit, transverse section, showing placenta and seeds. (H) Fruit transverse section, showing a stone cell at the apex (arrow). Photos by S. Leiva González and G. E. Barboza. https://doi.org/10.1371/journal.pone.0209792.g008" figureDoi="http://doi.org/10.5281/zenodo.12630929" httpUri="https://zenodo.org/record/12630929/files/figure.png" pageId="17" pageNumber="18">Fig 8B and 8C</figureCitation>
), the flowers are solitary or in fascicles of 3 (
<figureCitation id="136A2A45FFA9FFED31BDF8A56542F8AB" box="[982,1049,1835,1859]" captionStart="Fig 8" captionStartId="16.[527,555,1702,1721]" captionTargetBox="[527,1536,208,1680]" captionTargetId="figure-3@16.[527,1536,208,1680]" captionTargetPageId="16" captionText="Fig 8. Capsicum piuranum Barboza &amp; S. Leiva. (A) Plant. (B) Leaves, abaxial surface. (C) Fruiting branch. (D) Flower bud. (E) Flower and immature fruit. (F) Mature fruit. (G) Fruit, transverse section, showing placenta and seeds. (H) Fruit transverse section, showing a stone cell at the apex (arrow). Photos by S. Leiva González and G. E. Barboza. https://doi.org/10.1371/journal.pone.0209792.g008" figureDoi="http://doi.org/10.5281/zenodo.12630929" httpUri="https://zenodo.org/record/12630929/files/figure.png" pageId="17" pageNumber="18">Fig 8E</figureCitation>
), the corolla is tubular-campanulate and longer (
<quantity id="4CA99B25FFA9FFED302FF8C06390F88E" box="[580,715,1870,1894]" metricMagnitude="-2" metricUnit="m" metricValue="1.575" metricValueMax="1.7" metricValueMin="1.45" pageId="17" pageNumber="18" unit="mm" value="15.75" valueMax="17.0" valueMin="14.5">14.517 mm</quantity>
long), and the fruits are larger (
<quantity id="4CA99B25FFA9FFED3672F8C065D4F88E" box="[1049,1167,1870,1894]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="17" pageNumber="18" unit="cm" value="1.05" valueMax="1.2" valueMin="0.9">0.91.2 cm</quantity>
diam) and orange colored.
</paragraph>
<paragraph id="8BEE36C0FFAAFFEE305EFF5E6232FEB8" blockId="18.[533,1536,207,1896]" pageId="18" pageNumber="19">
This species exhibits the same number of chromosomes as
<taxonomicName id="4C514D43FFAAFFEE36CFFF5E6462FF0F" box="[1188,1337,207,232]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFAAFFEE36CFFF5E65EEFF0F" box="[1188,1205,208,231]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.
<emphasis id="B925EAD2FFAAFFEE36AAFF416462FF0F" box="[1217,1337,207,231]" italics="true" pageId="18" pageNumber="19">longifolium</emphasis>
</taxonomicName>
and the species that belong to the Andean clade [
<bibRefCitation id="EFC04B31FFAAFFEE311EFF7C62D5FEE2" author="Scaldaferro MA &amp; Grabiele M &amp; Moscone EA" box="[885,910,242,266]" pageId="18" pageNumber="19" pagination="693 - 709" refId="ref14020" refString="32. Scaldaferro MA, Grabiele M, Moscone EA. Heterochromatin type, amount and distribution in wild species of chili peppers (Capsicum - Solanaceae). Genet Resour Crop Ev. 2013; 60: 693 - 709." type="journal article" year="2013">32</bibRefCitation>
<bibRefCitation id="EFC04B31FFAAFFEE31F7FF7C62EEFEE2" author="Scaldaferro MA &amp; Barboza GE &amp; Moscone EA" box="[924,949,242,266]" pageId="18" pageNumber="19" refId="ref14095" refString="34. Scaldaferro MA, Barboza GE, Moscone EA. Estudios citogeneticos en especies andinas silvestres de Capsicum (Solanaceae). J Basic Appl Genet. 2011; 41: 31." type="journal volume" year="2011">34</bibRefCitation>
]. In addition,
<taxonomicName id="4C514D43FFAAFFEE3625FF7D658EFEE2" box="[1102,1237,242,266]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFAAFFEE3625FF7D6504FEE2" box="[1102,1119,243,266]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.
<emphasis id="B925EAD2FFAAFFEE3600FF7D658EFEE2" box="[1131,1237,243,266]" italics="true" pageId="18" pageNumber="19">piuranum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C514D43FFAAFFEE3762FF7D64C6FEE2" box="[1289,1437,242,266]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFAAFFEE3762FF7D6441FEE2" box="[1289,1306,243,266]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.
<emphasis id="B925EAD2FFAAFFEE374EFF7C64C6FEE2" box="[1317,1437,242,266]" italics="true" pageId="18" pageNumber="19">longifolium</emphasis>
</taxonomicName>
share the same karyotype formula, little heterochromatin, one only pair of NOR, and the smallest chromosomes in the genus [
<bibRefCitation id="EFC04B31FFAAFFEE312AFEB66201FEB8" author="Scaldaferro MA &amp; Grabiele M &amp; Moscone EA" box="[833,858,312,336]" pageId="18" pageNumber="19" pagination="693 - 709" refId="ref14020" refString="32. Scaldaferro MA, Grabiele M, Moscone EA. Heterochromatin type, amount and distribution in wild species of chili peppers (Capsicum - Solanaceae). Genet Resour Crop Ev. 2013; 60: 693 - 709." type="journal article" year="2013">32</bibRefCitation>
].
</paragraph>
<paragraph id="8BEE36C0FFAAFFEE305EFED4620DFE15" blockId="18.[533,1536,207,1896]" pageId="18" pageNumber="19">
The markedly anisophyllous leaves, the deflexed non-geniculate pedicels, the yellow corollas, the globose orange to red non-pungent fruits, the absence of giant cells and the presence of stone cells in the pericarp, the black seeds, and the chromosome number 2n = 26 place
<taxonomicName id="4C514D43FFAAFFEE37C6FE2E6308FE32" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="piuranum">
<emphasis id="B925EAD2FFAAFFEE37C6FE2E64E5FE5F" box="[1453,1470,416,439]" italics="true" pageId="18" pageNumber="19">C</emphasis>
.
<emphasis id="B925EAD2FFAAFFEE37A2FE2E6308FE32" italics="true" pageId="18" pageNumber="19">piuranum</emphasis>
</taxonomicName>
in the Andean clade proposed by Carrizo García et al. [
<bibRefCitation id="EFC04B31FFAAFFEE36F4FE4C65F7FE32" author="Carrizo Garcia C &amp; Sehr EM &amp; Barfuss MHJ &amp; Barboza GE &amp; Samuel R &amp; Moscone EA" box="[1183,1196,450,474]" pageId="18" pageNumber="19" pagination="35 - 51" refId="ref12646" refString="1. Carrizo Garcia C, Sehr EM, Barfuss MHJ, Barboza GE, Samuel R, Moscone EA et al. Phylogenetic relationships, diversification and expansion of chili peppers (Capsicum, Solanaceae). Ann Bot. 2016; 118: 35 - 51. https: // doi. org / 10.1093 / aob / mcw 079 PMID: 27245634" type="journal article" year="2016">1</bibRefCitation>
], as it has been determined in this work based on DNA data.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,426 @@
<document id="00C9E6555152E2BF46BFC07B9A86FAF6" ID-DOI="10.1371/journal.pone.0209792" ID-ISSN="1932-6203" ID-PMC="PMC6334993" ID-PubMed="30650102" ID-Zenodo-Dep="12630921" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1719975332143" checkinUser="felipe" docAuthor="Barboza, Gloria E., García, Carolina Carrizo, González, Segundo Leiva, Scaldaferro, Marisel &amp; Reyes, Ximena" docDate="2019" docId="03F887D6FFBBFFFA307EF9486265F950" docLanguage="en" docName="PLoSONE.14.1.e0209792.pdf" docOrigin="PLoS ONE (e 0209792) 14 (1)" docSource="http://dx.doi.org/10.1371/journal.pone.0209792" docStyle="DocumentStyle:55318D21378FF0528C342C9740A1E1BE.2:PLoSONE.2014-.journal_article" docStyleId="55318D21378FF0528C342C9740A1E1BE" docStyleName="PLoSONE.2014-.journal_article" docStyleVersion="2" docTitle="Capsicum benoistii Barboza 2019, sp. nov." docType="treatment" docVersion="1" lastPageNumber="7" masterDocId="FFC1FFAEFFB8FFFC326BFF8E615BFFE8" masterDocTitle="Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus" masterLastPageNumber="26" masterPageNumber="1" pageNumber="4" updateTime="1723725765965" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D64AB2E455B8B2A4172CCCEA8007C3BE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8C85FADAC21B70851E81334C7A3EEAC4">
<mods:title id="2EB612522667D4FF190BCA216E4CAA81">Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus</mods:title>
</mods:titleInfo>
<mods:name id="D9836C5CFBD484C8F5C51FA144241F71" type="personal">
<mods:role id="289F6B2748DBD182A3A1B22D8EAA8216">
<mods:roleTerm id="98349AC11FE0191383EA0BBA75DE2961">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EBECCC48CEF04AC4971F81207601ECB5">Barboza, Gloria E.</mods:namePart>
</mods:name>
<mods:name id="4EBBAF8B3A531B71EF20C28D1E1E6311" type="personal">
<mods:role id="67C4797FA6BD7C7ACE8F1E7B3763EB19">
<mods:roleTerm id="11E8DE122B69FB4822A554F5A6235E0B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BCE2242F1F68C9A0B2021CFDE48DCDDC">García, Carolina Carrizo</mods:namePart>
<mods:affiliation id="C00E630218EE510A70180A662E5EB4FD">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Department of Botany and Biodiversity Research, University of Vienna, Vienna, Austria,</mods:affiliation>
</mods:name>
<mods:name id="102A07D780675E01E249567A540F8C8C" type="personal">
<mods:role id="DAB09C1A4C9C21CF68DC2A2BB35DD3F8">
<mods:roleTerm id="82A46354383984238B476CBC02233351">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BF6D131A46D7FDB1C95C61E6FDB87CC">González, Segundo Leiva</mods:namePart>
</mods:name>
<mods:name id="4BBE07E6678E909829D0785E21465840" type="personal">
<mods:role id="25078780F9DF88FB29B6EFEF31106590">
<mods:roleTerm id="32E9F6FCCDED83B5D2276EB02F75C956">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E225782C252D03CC4D1E01E6922B5B46">Scaldaferro, Marisel</mods:namePart>
<mods:affiliation id="E58547C35CBFC2AA4406079FA528E40C">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Córdoba, Argentina,</mods:affiliation>
</mods:name>
<mods:name id="81AD8318817F1CEB5DD4E42C5403FAAB" type="personal">
<mods:role id="E628CD75F5DC88FE593D2CA392FF479A">
<mods:roleTerm id="9ADE36F9170804FFE1CE779E46054F26">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="123237741A85F3719BE939AA4321D2D6">Reyes, Ximena</mods:namePart>
<mods:affiliation id="617FC7B1D01E70AFA8CD8F1675454FD9">Centro de Investigaciones Fitoecogenéticas de Pairumani, Cochabamba, Bolivia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="FAB8A4228FD2C542113A991033531EBA">text</mods:typeOfResource>
<mods:relatedItem id="03508BE65E403B771A53204996D23EDF" type="host">
<mods:titleInfo id="8DC3DF813D9648D785D08392FCBAE692">
<mods:title id="8C545AE8B418AE6623A5951BE23786A0">PLoS ONE</mods:title>
</mods:titleInfo>
<mods:part id="CCE4C695CCB9ECC9E8A8ED4FFE600EC2">
<mods:date id="AEEF63D539682BB9F2E6667FD133E65A">2019</mods:date>
<mods:detail id="8E19888AC9FE7E5881A4E8C3DF5D3B43" type="series">
<mods:title id="0CD556FF1DF24062EC4669D7799DFEBE">e 0209792</mods:title>
</mods:detail>
<mods:detail id="840ABE9E5246223D109D08CD3DBF26AF" type="pubDate">
<mods:number id="DDD0FA98704F5C6C864479DC768B8FBC">2019-01-16</mods:number>
</mods:detail>
<mods:detail id="0A9693D18D446CE522178AB51F222B01" type="volume">
<mods:number id="115883A6DDC0B34AF9FF6406959EF86C">14</mods:number>
</mods:detail>
<mods:detail id="74B475597BA13B0E26785138F7FBACF1" type="issue">
<mods:number id="48004C373FEB04FFF4A74378BD8CCA23">1</mods:number>
</mods:detail>
<mods:extent id="D8B004D4DD9A2BC518E3F99A5E985AF6" unit="page">
<mods:start id="41B8BFD75D93513BB6F7B275C2D67FA2">1</mods:start>
<mods:end id="97AF4547370FDCD1C29A93B5BED970EE">26</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="9DE0974D833FB5484FC67B521FBDA96E">
<mods:url id="1D933BD936034A76E7FFC616A3927164">http://dx.doi.org/10.1371/journal.pone.0209792</mods:url>
</mods:location>
<mods:classification id="0A33BC9FD8DDF5EBDEAFB1F3C45C2053">journal article</mods:classification>
<mods:identifier id="96297D5A009AF3A5EC2067116AA90583" type="DOI">10.1371/journal.pone.0209792</mods:identifier>
<mods:identifier id="3D807052994B76240BC85EC674060EFB" type="ISSN">1932-6203</mods:identifier>
<mods:identifier id="94B461842F7EA390B7DDD3EB78EAF4A5" type="PMC">PMC6334993</mods:identifier>
<mods:identifier id="5B3F88F690E29BCC2174244664CCE3B7" type="PubMed">30650102</mods:identifier>
<mods:identifier id="EAC6D50C79C10197F19E8EF342876488" type="Zenodo-Dep">12630921</mods:identifier>
</mods:mods>
<treatment id="03F887D6FFBBFFFA307EF9486265F950" LSID="urn:lsid:plazi:treatment:03F887D6FFBBFFFA307EF9486265F950" httpUri="http://treatment.plazi.org/id/03F887D6FFBBFFFA307EF9486265F950" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
<heading id="D0A681ACFFBBFFFF307EF9486402F8E9" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
<subSubSection id="C34B654BFFBBFFFF307EF948654BF936" box="[533,1040,1733,1758]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BEE36C0FFBBFFFF307EF948654BF936" blockId="3.[533,1524,1685,1897]" box="[533,1040,1733,1758]" pageId="3" pageNumber="4">
<emphasis id="B925EAD2FFBBFFFF307EF948654BF936" bold="true" box="[533,1040,1733,1758]" pageId="3" pageNumber="4">
<taxonomicName id="4C514D43FFBBFFFF307EF94862EAF936" ID-CoL="7TBQJ" authority="Hunz. ex Barboza" authorityName="Barboza" authorityYear="2019" box="[533,945,1733,1758]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="benoistii" status="sp. nov.">
<emphasis id="B925EAD2FFBBFFFF307EF94863BFF935" bold="true" box="[533,740,1733,1758]" italics="true" pageId="3" pageNumber="4">Capsicum benoistii</emphasis>
Hunz. ex Barboza
</taxonomicName>
,
<taxonomicNameLabel id="A21657A9FFBBFFFF31D0F948654BF936" box="[955,1040,1734,1758]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFBBFFFF367DF9486331F8CB" pageId="3" pageNumber="4" type="description">
<paragraph id="8BEE36C0FFBBFFFF367DF9486402F8E9" blockId="3.[533,1524,1685,1897]" pageId="3" pageNumber="4">
[
<materialsCitation id="3B393C9DFFBBFFFF3677F948640EF8E9" collectingDate="1931-04-03" collectorName="M. R. Benoist" country="Ecuador" location="Banos" municipality="Type" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Tungurahua" typeStatus="holotype">
urn:lsid:ipni.org:names: 771925561].
<collectingMunicipality id="6B8AACBAFFBBFFFF37DCF94864B4F936" box="[1463,1519,1734,1758]" pageId="3" pageNumber="4">Type</collectingMunicipality>
:
<collectingCountry id="F3467650FFBBFFFF307EF967632AF8E9" box="[533,625,1769,1793]" name="Ecuador" pageId="3" pageNumber="4">Ecuador</collectingCountry>
.
<collectingRegion id="4995F822FFBBFFFF3010F9676259F8E9" box="[635,770,1769,1793]" country="Ecuador" name="Tungurahua" pageId="3" pageNumber="4">Tungurahua</collectingRegion>
:
<location id="8E8E601BFFBBFFFF3160F9676215F8E9" LSID="urn:lsid:plazi:treatment:03F887D6FFBBFFFA307EF9486265F950:8E8E601BFFBBFFFF3160F9676215F8E9" box="[779,846,1769,1793]" country="Ecuador" municipality="Type" name="Banos" pageId="3" pageNumber="4" stateProvince="Tungurahua">Baños</location>
,
<date id="FFEF1000FFBBFFFF3133F9676296F8E9" box="[856,973,1769,1793]" pageId="3" pageNumber="4" value="1931-04-03">
<collectingDate id="EFABE9E8FFBBFFFF3133F9676296F8E9" box="[856,973,1769,1793]" pageId="3" pageNumber="4" value="1931-04-03">3 Apr 1931</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFBBFFFF3194F96765D0F8E8" box="[1023,1163,1769,1793]" pageId="3" pageNumber="4">
<emphasis id="B925EAD2FFBBFFFF3194F967654DF8E8" box="[1023,1046,1769,1792]" italics="true" pageId="3" pageNumber="4">M</emphasis>
.
<emphasis id="B925EAD2FFBBFFFF3649F9676569F8E8" box="[1058,1074,1769,1792]" italics="true" pageId="3" pageNumber="4">R</emphasis>
.
<emphasis id="B925EAD2FFBBFFFF3655F9676598F8E9" box="[1086,1219,1769,1793]" italics="true" pageId="3" pageNumber="4">Benoist 4204</emphasis>
</collectorName>
(
<typeStatus id="54EA8862FFBBFFFF36BBF967646AF8E9" box="[1232,1329,1769,1793]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
, P)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36C0FFBBFFFF305EF8856331F8CB" blockId="3.[533,1524,1685,1897]" box="[565,618,1803,1827]" pageId="3" pageNumber="4">
<figureCitation id="136A2A45FFBBFFFF305EF8856331F8CB" box="[565,618,1803,1827]" captionStart="Fig 1" captionStartId="5.[454,482,203,222]" captionTargetId="figure-1@4.[454,1536,208,1872]" captionTargetPageId="4" captionText="Fig 1. Capsicum benoistii Hunz. ex Barboza. (A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury." pageId="3" pageNumber="4">Fig 1</figureCitation>
</paragraph>
</subSubSection>
</heading>
<subSubSection id="C34B654BFFBBFFF9305EF8A06205FED2" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BEE36C0FFBBFFFF305EF8A06484F881" blockId="3.[533,1524,1685,1897]" pageId="3" pageNumber="4">
<emphasis id="B925EAD2FFBBFFFF305EF8A063F8F8AE" bold="true" box="[565,675,1838,1862]" pageId="3" pageNumber="4">Diagnosis</emphasis>
. Similar to
<taxonomicName id="4C514D43FFBBFFFF3176F8A0658CF8AE" authority="(Dammer) Hunz." authorityName="Hunz." baseAuthorityName="Dammer" box="[797,1239,1838,1862]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="geminifolium">
<emphasis id="B925EAD2FFBBFFFF3176F8A0654DF8AE" box="[797,1046,1838,1862]" italics="true" pageId="3" pageNumber="4">Capsicum geminifolium</emphasis>
(Dammer) Hunz.
</taxonomicName>
but differing in the length of the flowering pedicels, the shape of the corolla, and the presence of heterostylous flowers.
</paragraph>
<caption id="DF2E6648FFBDFFF933ADFF4563AAFEF9" pageId="5" pageNumber="6" startId="5.[454,482,203,222]" targetBox="[454,1532,208,1872]" targetPageId="4" targetType="figure">
<paragraph id="8BEE36C0FFBDFFF933ADFF4563AAFEF9" blockId="5.[454,1518,202,273]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF933ADFF456212FF36" bold="true" box="[454,841,202,222]" pageId="5" pageNumber="6">
Fig 1.
<taxonomicName id="4C514D43FFBDFFF93395FF456212FF36" authority="Hunz. ex Barboza." authorityName="Barboza." box="[510,841,202,222]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBDFFF93395FF4563F8FF35" bold="true" box="[510,675,202,222]" italics="true" pageId="5" pageNumber="6">Capsicum benoistii</emphasis>
Hunz. ex Barboza.
</taxonomicName>
</emphasis>
(A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury.
</paragraph>
</caption>
<paragraph id="8BEE36C0FFBDFFF933ADFEA96205FED2" blockId="5.[454,862,295,314]" box="[454,862,295,314]" pageId="5" pageNumber="6">https://doi.org/10.1371/journal.pone.0209792.g001</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFBDFFF9305EFEED64ACFB70" pageId="5" pageNumber="6" type="description">
<paragraph id="8BEE36C0FFBDFFF9305EFEED64ACFB70" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFEED63E3FE93" bold="true" box="[565,696,355,379]" pageId="5" pageNumber="6">Description</emphasis>
. Shrubs few branched. Young stems light brown, glabrous or pubescent, striate; bark of older stems dark brown, glabrous, striate; lenticels absent. Sympodial units difoliate, geminate, leaf pair markedly anisophyllous in size and shape. Leaves simple, membranaceous, discolorous, adaxial surface dark green, abaxial surface light green, glabrous or with simple antrorse trichomes
<quantity id="4CA99B25FFBDFFF9310EFE6362BFFDED" box="[869,996,493,517]" metricMagnitude="-4" metricUnit="m" metricValue="7" metricValueMax="11.0" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="0.7" valueMax="1.1" valueMin="0.3">0.31.1 mm</quantity>
long adaxially and abaxially, trichomes more abundant on main veins; the larger leaves with blades
<quantity id="4CA99B25FFBDFFF9363BFD9E65E6FDC0" box="[1104,1213,528,552]" metricMagnitude="-1" metricUnit="m" metricValue="1.025" metricValueMax="1.2" metricValueMin="0.85" pageId="5" pageNumber="6" unit="cm" value="10.25" valueMax="12.0" valueMin="8.5">8.512 cm</quantity>
long,
<quantity id="4CA99B25FFBDFFF93695FD9E6404FDC0" box="[1278,1375,528,552]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="6.0" metricValueMin="2.8" pageId="5" pageNumber="6" unit="cm" value="4.4" valueMax="6.0" valueMin="2.8">2.86 cm</quantity>
wide, ovate or elliptic, major veins 45 (6) on each side of midvein, base asymmetric and attenuate, margin entire, apex long-acuminate; petioles 0.51.0 cm long, glabrous or glabrescent with trichomes like those of the leaves; the minor leaves
<quantity id="4CA99B25FFBDFFF931AAFDF66579FD78" box="[961,1058,632,656]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="6.0" metricValueMin="2.4" pageId="5" pageNumber="6" unit="cm" value="4.2" valueMax="6.0" valueMin="2.4">2.46 cm</quantity>
long,
<quantity id="4CA99B25FFBDFFF93608FDF66598FD78" box="[1123,1219,632,656]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="4.0" metricValueMin="1.7" pageId="5" pageNumber="6" unit="cm" value="2.85" valueMax="4.0" valueMin="1.7">1.74 cm</quantity>
wide, ovate or elliptic, major veins 34 on each side of midvein, base rounded, asymmetric, margin entire, apex acute or rounded; petioles
<quantity id="4CA99B25FFBDFFF930BFFD33626EFD3D" box="[724,821,701,725]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="10.0" metricValueMin="1.0" pageId="5" pageNumber="6" unit="cm" value="0.55" valueMax="1.0" valueMin="0.1">0.11 cm</quantity>
long, glabrescent or pubescent. Flowers in fascicles of 36; flowering pedicels filiform, striate, pendent, not geniculate at anthesis,
<quantity id="4CA99B25FFBDFFF936D5FD6E6445FD10" box="[1214,1310,736,760]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="5" pageNumber="6" unit="cm" value="1.65" valueMax="2.0" valueMin="1.3">1.32 cm</quantity>
long, moderately to densely pubescent, the trichomes simple, non-glandular, multicellular, antrorse,
<quantity id="4CA99B25FFBDFFF93702FC8D631AFCD5" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="7.5" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="0.525" valueMax="0.75" valueMin="0.3">0.300.75 mm</quantity>
long. Flower buds ovoid. Calyx
<quantity id="4CA99B25FFBDFFF931FEFCAB655AFCD5" box="[917,1025,805,829]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="5" pageNumber="6" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 mm</quantity>
long, ca.
<quantity id="4CA99B25FFBDFFF9360FFCAB65F8FCD5" box="[1124,1187,805,829]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="5.0">5 mm</quantity>
wide, cup-shaped, thick, the margin truncate, pubescent with the same trichomes as pedicels, with 5 appendages
<quantity id="4CA99B25FFBDFFF937E5FCC6631AFC6B" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="5" pageNumber="6" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.53.5 mm</quantity>
long, ca.
<quantity id="4CA99B25FFBDFFF930CEFCE563ADFC6B" box="[677,758,875,899]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
wide, thick, erect, subulate, inserted close to the margin, pubescent with the same trichomes as calyx tube. Corolla ca.
<quantity id="4CA99B25FFBDFFF93198FC03653EFC4D" box="[1011,1125,909,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="5" pageNumber="6" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">1213 mm</quantity>
long, deeply stellate, thick, without interpetalar tissue; tube ca.
<quantity id="4CA99B25FFBDFFF9315DFC3E622FFC20" box="[822,884,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="3.0">3 mm</quantity>
long, glabrous inside and outside; lobes ca.
<quantity id="4CA99B25FFBDFFF93728FC3E64DAFC20" box="[1347,1409,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="6" unit="mm" value="9.0">9 mm</quantity>
long, ca.
<quantity id="4CA99B25FFBDFFF9378FFC3E631AFC03" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="mm" value="2.0">2 mm</quantity>
wide, narrowly triangular, erect, glabrous adaxially and abaxially, the tips and margins pubescent. Stamens 5, equal, filaments equal,
<quantity id="4CA99B25FFBDFFF9319CFC7B6539FBE5" box="[1015,1122,1013,1037]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="3.1" valueMax="3.2" valueMin="3.0">33.2 mm</quantity>
long, glabrous, inserted on the corolla
<quantity id="4CA99B25FFBDFFF9307EFB96633CFBD8" box="[533,615,1048,1072]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="mm" value="1.5">1.5 mm</quantity>
from the base, with inconspicuous auricles at point of insertion; anthers ca.
<quantity id="4CA99B25FFBDFFF937E0FB966491FBD8" box="[1419,1482,1048,1072]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="3.0">3 mm</quantity>
long, not connivent, elliptic. Ovary
<quantity id="4CA99B25FFBDFFF931E7FBB56550FBBB" box="[908,1035,1083,1107]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="5" pageNumber="6" unit="mm" value="1.5" valueMax="1.7" valueMin="1.3">1.31.7 mm</quantity>
long,
<quantity id="4CA99B25FFBDFFF93627FBB56591FBBB" box="[1100,1226,1083,1107]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="5" pageNumber="6" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 mm</quantity>
diam, subglobose, glabrous; nectary ca.
<quantity id="4CA99B25FFBDFFF930E6FBD36385FB9D" box="[653,734,1117,1141]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="0.3">0.3 mm</quantity>
long, inconspicuous; long style ca.
<quantity id="4CA99B25FFBDFFF93639FBD365F8FB9D" box="[1106,1187,1117,1141]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="6" unit="mm" value="6.5">6.5 mm</quantity>
long, short style ca.
<quantity id="4CA99B25FFBDFFF93712FBD36491FB9D" box="[1401,1482,1117,1141]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="6" unit="mm" value="3.6">3.6 mm</quantity>
long, widening distally, glabrous; stigma
<quantity id="4CA99B25FFBDFFF931AEFB0E654DFB70" box="[965,1046,1152,1176]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="0.3">0.3 mm</quantity>
long,
<quantity id="4CA99B25FFBDFFF9363DFB0E65F3FB70" box="[1110,1192,1152,1176]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
wide, globose. Berry unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFBDFFFA305EFB2D6265F950" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BEE36C0FFBDFFF9305EFB2D64AFFB36" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFB2D6211FB53" bold="true" box="[565,842,1187,1211]" pageId="5" pageNumber="6">Distribution and ecology</emphasis>
. Endemic to a restricted area in central-southern
<collectingCountry id="F3467650FFBDFFF9373CFB2D64EAFB53" box="[1367,1457,1187,1211]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
(
<collectingRegion id="4995F822FFBDFFF937D4FB2D6334FB36" country="Ecuador" name="Tungurahua" pageId="5" pageNumber="6">Tungurahua</collectingRegion>
,
<collectingRegion id="4995F822FFBDFFF93012FB4863F1FB36" box="[633,682,1222,1246]" country="Ecuador" name="Loja" pageId="5" pageNumber="6">Loja</collectingRegion>
,
<figureCitation id="136A2A45FFBDFFF930D8FB4863B3FB36" box="[691,744,1222,1246]" captionStart="Fig 2" captionStartId="6.[533,561,1150,1169]" captionTargetBox="[533,1320,208,1128]" captionTargetId="figure-278@6.[533,1320,208,1129]" captionTargetPageId="6" captionText="Fig 2. Distribution of Capsicum species. https://doi.org/10.1371/journal.pone.0209792.g002" figureDoi="http://doi.org/10.5281/zenodo.12630923" httpUri="https://zenodo.org/record/12630923/files/figure.png" pageId="5" pageNumber="6">Fig 2</figureCitation>
) growing in thickets in montane forests, between
<quantity id="4CA99B25FFBDFFF93692FB4864D3FB36" box="[1273,1416,1222,1246]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.6" metricValueMin="1.5" pageId="5" pageNumber="6" unit="m" value="2050.0" valueMax="2600.0" valueMin="1500.0">15002600 m</quantity>
elevation.
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9305EFB66645CFAE8" blockId="5.[533,1536,355,1869]" box="[565,1287,1256,1280]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFB6663F1FAE8" bold="true" box="[565,682,1256,1280]" pageId="5" pageNumber="6">Phenology</emphasis>
. Flowering from March to May. Fruiting time unknown.
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9305EFA8562EFFA80" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFA8563F7FACB" bold="true" box="[565,684,1291,1315]" pageId="5" pageNumber="6">Etymology</emphasis>
—The new species is named in honor to Raymond Benoist (18811970), a French botanist, who collected in French
<collectingCountry id="F3467650FFBDFFF931A7FAA0657AFAAE" box="[972,1057,1326,1350]" name="Guyana" pageId="5" pageNumber="6">Guyana</collectingCountry>
,
<collectingCountry id="F3467650FFBDFFF93640FAA065D7FAAE" box="[1067,1164,1326,1350]" name="Morocco" pageId="5" pageNumber="6">Morocco</collectingCountry>
and
<collectingCountry id="F3467650FFBDFFF936ABFAA06447FAAE" box="[1216,1308,1326,1350]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
; the
<typeStatus id="54EA8862FFBDFFF93726FAA064F1FAAE" box="[1357,1450,1326,1350]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
is a nice specimen collected by her in 1931.
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9305EFAFD64ECFA38" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFAFD62F3FA63" bold="true" box="[565,936,1395,1419]" pageId="5" pageNumber="6">Species Conservation Assessment</emphasis>
. Following the IUCN Red List Criteria (IUCN 2017), this species is proposed as Endangered (
<collectionCode id="ED40AE05FFBDFFF931D7FA1862BFFA46" box="[956,996,1430,1454]" pageId="5" pageNumber="6">EN</collectionCode>
). The extent of occurrence is calculated to be
<quantity id="4CA99B25FFBDFFF937AFFA186363FA38" metricMagnitude="6" metricUnit="m" metricValue="2.05" pageId="5" pageNumber="6" unit="km" value="2050.0">2050 km</quantity>
2 (Criterion B1 &lt;
<quantity id="4CA99B25FFBDFFF93092FA36620CFA38" box="[761,855,1464,1488]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="km" value="5000.0">5000 km</quantity>
2, Endangered), the area of occupancy,
<quantity id="4CA99B25FFBDFFF9369DFA366463FA38" box="[1270,1336,1464,1488]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="6" unit="km" value="12.0">12 km</quantity>
2 (Criterion
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9307EFA55648BFA1B" blockId="5.[533,1536,355,1869]" box="[533,1488,1499,1523]" pageId="5" pageNumber="6">
B2 &lt;
<quantity id="4CA99B25FFBDFFF93039FA5563F8FA1B" box="[594,675,1499,1523]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="km" value="500.0">500 km</quantity>
2, Endangered) and the species is known from only three localities (Criterion
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9307EFA7362D3F9B3" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
B1a
<emphasis id="B925EAD2FFBDFFF93028FA736303F9FD" bold="true" box="[579,600,1533,1557]" pageId="5" pageNumber="6"><EFBFBD></emphasis>
5, Endangered). It is possible that its geographic range has declined (EOO and AOO, Criterion B2b i &amp; ii) because the species has not been collected since 1978 despite recent intensive searches in the same locations.
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9305EF9EB65F7F92B" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EF9EB62C3F995" bold="true" box="[565,920,1637,1661]" pageId="5" pageNumber="6">Additional specimens examined</emphasis>
.
<materialsCitation id="3B393C9DFFBDFFF931CFF9EB644AF948" collectingDate="1978-05-21" collectorName="F. Vivar &amp; Estudiantes" country="Ecuador" elevation="2580" location="Pueblo Nuevo" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Loja">
<emphasis id="B925EAD2FFBDFFF931CFF9EB6539F995" bold="true" box="[932,1122,1637,1661]" pageId="5" pageNumber="6">
<collectingCountry id="F3467650FFBDFFF931CFF9EB6573F995" box="[932,1064,1637,1661]" name="Ecuador" pageId="5" pageNumber="6">ECUADOR</collectingCountry>
.
<collectingRegion id="4995F822FFBDFFF93659F9EB6539F995" box="[1074,1122,1637,1661]" country="Ecuador" name="Loja" pageId="5" pageNumber="6">Loja</collectingRegion>
</emphasis>
:
<location id="8E8E601BFFBDFFF93604F9EB645CF995" LSID="urn:lsid:plazi:treatment:03F887D6FFBBFFFA307EF9486265F950:8E8E601BFFBDFFF93604F9EB645CF995" box="[1135,1287,1637,1661]" country="Ecuador" name="Pueblo Nuevo" pageId="5" pageNumber="6" stateProvince="Loja">Pueblo Nuevo</location>
, 04˚0551S, 79˚ 1155W,
<quantity id="4CA99B25FFBDFFF93014F9066394F948" box="[639,719,1672,1696]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="5" pageNumber="6" unit="m" value="2580.0">
<elevation id="007CD1F3FFBDFFF93014F9066394F948" box="[639,719,1672,1696]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="5" pageNumber="6" unit="m" value="2580.0">2580 m</elevation>
</quantity>
,
<date id="FFEF1000FFBDFFF930B2F906623AF948" box="[729,865,1672,1696]" pageId="5" pageNumber="6" value="1978-05-21">
<collectingDate id="EFABE9E8FFBDFFF930B2F906623AF948" box="[729,865,1672,1696]" pageId="5" pageNumber="6" value="1978-05-21">21 May 1978</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFBDFFF931F8F90762B3F948" box="[915,1000,1672,1696]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF931F8F90762FAF948" box="[915,929,1673,1696]" italics="true" pageId="5" pageNumber="6">F</emphasis>
.
<emphasis id="B925EAD2FFBDFFF931C7F90765E1F948" box="[940,1210,1672,1696]" italics="true" pageId="5" pageNumber="6">
Vivar &amp;
<collectorName id="26A45316FFBDFFF93663F90665DAF948" box="[1032,1153,1672,1696]" pageId="5" pageNumber="6">Estudiantes</collectorName>
1066
</emphasis>
</collectorName>
(LOJA)
</materialsCitation>
;
<materialsCitation id="3B393C9DFFBDFFF93770F90665F2F92B" collectingDate="1956-03-30" collectorName="E. Asplund" country="Ecuador" elevation="1500" location="Rio Verde Grande" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Tungurahua">
<emphasis id="B925EAD2FFBDFFF93770F90664FDF948" bold="true" box="[1307,1446,1672,1696]" pageId="5" pageNumber="6">
<collectingRegion id="4995F822FFBDFFF93770F90664FDF948" box="[1307,1446,1672,1696]" country="Ecuador" name="Tungurahua" pageId="5" pageNumber="6">Tungurahua</collectingRegion>
</emphasis>
:
<location id="8E8E601BFFBDFFF937DAF90663F4F92B" LSID="urn:lsid:plazi:treatment:03F887D6FFBBFFFA307EF9486265F950:8E8E601BFFBDFFF937DAF90663F4F92B" country="Ecuador" name="Rio Verde Grande" pageId="5" pageNumber="6" stateProvince="Tungurahua">Río Verde Grande</location>
,
<quantity id="4CA99B25FFBDFFF930D2F9256252F92B" box="[697,777,1707,1731]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="m" value="1500.0">
<elevation id="007CD1F3FFBDFFF930D2F9256252F92B" box="[697,777,1707,1731]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="FFEF1000FFBDFFF93178F92562C3F92B" box="[787,920,1707,1731]" pageId="5" pageNumber="6" value="1956-03-30">
<collectingDate id="EFABE9E8FFBDFFF93178F92562C3F92B" box="[787,920,1707,1731]" pageId="5" pageNumber="6" value="1956-03-30">30 Mar 1956</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFBDFFF931A1F9256566F92A" box="[970,1085,1706,1731]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF931A1F9256282F92A" box="[970,985,1707,1730]" italics="true" pageId="5" pageNumber="6">E</emphasis>
.
<emphasis id="B925EAD2FFBDFFF9318EF92465DAF92B" box="[997,1153,1706,1731]" italics="true" pageId="5" pageNumber="6">Asplund 20070</emphasis>
</collectorName>
(S)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36C0FFBDFFFA305EF940640FFA68" blockId="5.[533,1536,355,1869]" lastBlockId="6.[533,1536,1245,1893]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<taxonomicName id="4C514D43FFBDFFF9305EF94063A7F90D" box="[565,764,1741,1765]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBDFFF9305EF94063A7F90D" box="[565,764,1741,1765]" italics="true" pageId="5" pageNumber="6">Capsicum benoistii</emphasis>
</taxonomicName>
was identified as a new species by the late
<taxonomicName id="4C514D43FFBDFFF936D2F9436476F90D" box="[1209,1325,1741,1765]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
specialist Armando T. Hunziker (
<collectionCode id="ED40AE05FFBDFFF930CFF97E63AAF8E0" box="[676,753,1776,1800]" country="Argentina" lsid="urn:lsid:biocol.org:col:15636" name="Universidad Nacional de Córdoba" pageId="5" pageNumber="6" type="Herbarium">CORD</collectionCode>
) who annotated the epithet name
<taxonomicName id="4C514D43FFBDFFF93630F97E65EFF8E0" box="[1115,1204,1776,1800]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBDFFF93630F97E65EFF8E0" box="[1115,1204,1776,1800]" italics="true" pageId="5" pageNumber="6">benoistii</emphasis>
</taxonomicName>
on the specimen housed at P (Benoist 4204), but this name was never published. It is a poorly known species collected only three times in
<collectingCountry id="F3467650FFBDFFF930C6F8BB6252F8A5" box="[685,777,1845,1869]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
; none of these collections have fruits. Extensive recent field explorations in
<collectingRegion id="4995F822FFBEFFFA30D0FB53621BFB1D" box="[699,832,1245,1269]" country="Ecuador" name="Tungurahua" pageId="6" pageNumber="7">Tungurahua</collectingRegion>
were unsuccessful in finding this species. It is distinctive in its deeply lobed stellate corolla (lobes three times longer than the tube,
<figureCitation id="136A2A45FFBEFFFA368BFA8E647FFAF0" box="[1248,1316,1280,1304]" captionStart="Fig 1" captionStartId="5.[454,482,203,222]" captionTargetId="figure-1@4.[454,1536,208,1872]" captionTargetPageId="4" captionText="Fig 1. Capsicum benoistii Hunz. ex Barboza. (A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury." pageId="6" pageNumber="7">Fig 1B</figureCitation>
) and in the presence of heterostylous flowers (
<figureCitation id="136A2A45FFBEFFFA314BFAAC62E0FAD2" box="[800,955,1314,1338]" captionStart="Fig 1" captionStartId="5.[454,482,203,222]" captionTargetId="figure-1@4.[454,1536,208,1872]" captionTargetPageId="4" captionText="Fig 1. Capsicum benoistii Hunz. ex Barboza. (A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury." pageId="6" pageNumber="7">Fig 1G and 1H</figureCitation>
). These features plus the short flowering pedicels (
<quantity id="4CA99B25FFBEFFFA3076FACB6324FAB5" box="[541,639,1349,1373]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="6" pageNumber="7" unit="cm" value="1.65" valueMax="2.0" valueMin="1.3">1.32 cm</quantity>
long) distinguish
<taxonomicName id="4C514D43FFBEFFFA3154FACB62EEFAB5" box="[831,949,1349,1373]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBEFFFA3154FACB620BFAB4" box="[831,848,1349,1372]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA3137FACB62EEFAB5" box="[860,949,1349,1373]" italics="true" pageId="6" pageNumber="7">benoistii</emphasis>
</taxonomicName>
from
<taxonomicName id="4C514D43FFBEFFFA319EFACB65C7FAB5" box="[1013,1180,1349,1373]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="geminifolium">
<emphasis id="B925EAD2FFBEFFFA319EFACB655DFAB4" box="[1013,1030,1349,1372]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA367AFACB65C7FAB5" box="[1041,1180,1349,1373]" italics="true" pageId="6" pageNumber="7">geminifolium</emphasis>
</taxonomicName>
, which has funnel-shaped corollas lobed about halfway, homostylous flowers, and longer pedicels (
<quantity id="4CA99B25FFBEFFFA36B0FAE66454FA68" box="[1243,1295,1384,1408]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="cm" value="5.0">5 cm</quantity>
long).
</paragraph>
<caption id="DF2E6648FFBEFFFA307EFBF062F6FB53" ID-DOI="http://doi.org/10.5281/zenodo.12630923" ID-Zenodo-Dep="12630923" httpUri="https://zenodo.org/record/12630923/files/figure.png" pageId="6" pageNumber="7" startId="6.[533,561,1150,1169]" targetBox="[533,1320,208,1128]" targetPageId="6" targetType="figure">
<paragraph id="8BEE36C0FFBEFFFA307EFBF0622EFB79" blockId="6.[533,941,1150,1211]" box="[533,885,1150,1169]" pageId="6" pageNumber="7">
<emphasis id="B925EAD2FFBEFFFA307EFBF0622EFB79" bold="true" box="[533,885,1150,1169]" pageId="6" pageNumber="7">
Fig 2. Distribution of
<taxonomicName id="4C514D43FFBEFFFA30BCFBF0622EFB79" box="[727,885,1150,1169]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B925EAD2FFBEFFFA30BCFBF06276FB79" bold="true" box="[727,813,1150,1169]" italics="true" pageId="6" pageNumber="7">Capsicum</emphasis>
species.
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BEE36C0FFBEFFFA307EFB2662F6FB53" blockId="6.[533,941,1150,1211]" box="[533,941,1192,1211]" pageId="6" pageNumber="7">https://doi.org/10.1371/journal.pone.0209792.g002</paragraph>
</caption>
<paragraph id="8BEE36C0FFBEFFFA305EFA046584F9C5" blockId="6.[533,1536,1245,1893]" pageId="6" pageNumber="7">
The presence of heterostylous flowers as in
<taxonomicName id="4C514D43FFBEFFFA366BFA04652EFA4A" box="[1024,1141,1418,1442]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBEFFFA366BFA04654AFA49" box="[1024,1041,1418,1441]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA3677FA04652EFA4A" box="[1052,1141,1418,1442]" italics="true" pageId="6" pageNumber="7">benoistii</emphasis>
</taxonomicName>
is unusual among
<taxonomicName id="4C514D43FFBEFFFA3757FA0464A1FA4A" box="[1340,1530,1418,1442]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B925EAD2FFBEFFFA3757FA0464FFFA49" box="[1340,1444,1418,1441]" italics="true" pageId="6" pageNumber="7">Capsicum</emphasis>
species.
</taxonomicName>
It has been reported in
<taxonomicName id="4C514D43FFBEFFFA3160FA2362D5FA2D" box="[779,910,1453,1477]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="baccatum">
<emphasis id="B925EAD2FFBEFFFA3160FA236247FA2C" box="[779,796,1453,1476]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA3143FA2362D5FA2D" box="[808,910,1453,1477]" italics="true" pageId="6" pageNumber="7">baccatum</emphasis>
</taxonomicName>
L. varieties:
<taxonomicName id="4C514D43FFBEFFFA3679FA2365FEFA2D" box="[1042,1189,1453,1477]" pageId="6" pageNumber="7" rank="variety" variety="baccatum">
var.
<emphasis id="B925EAD2FFBEFFFA3654FA2365FEFA2D" box="[1087,1189,1453,1477]" italics="true" pageId="6" pageNumber="7">baccatum</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFC04B31FFBEFFFA36DFFA236596FA2D" author="Barboza GE" box="[1204,1229,1453,1477]" pageId="6" pageNumber="7" pagination="21 - 25" refId="ref13912" refString="29. Barboza GE. Capsicum. In Barboza GE, coordinator. Solanaceae. Flora Argentina. Buenos Aires: IBODA-IMBIV, CONICET; 2013; 13: 21 - 25." type="journal article" year="2013">29</bibRefCitation>
] and
<taxonomicName id="4C514D43FFBEFFFA3760FA23624DFA00" authority="(Vell.) Hunz. &amp; Barboza" authorityName="Hunz. &amp; Barboza" baseAuthorityName="Vell." pageId="6" pageNumber="7" rank="variety" variety="umbilicatum">
var.
<emphasis id="B925EAD2FFBEFFFA3753FA2364E4FA2D" box="[1336,1471,1453,1477]" italics="true" pageId="6" pageNumber="7">umbilicatum</emphasis>
(Vell.) Hunz. &amp; Barboza
</taxonomicName>
[
<bibRefCitation id="EFC04B31FFBEFFFA314EFA5E6265FA00" author="Hunziker AT &amp; Barboza GE" box="[805,830,1488,1512]" pageId="6" pageNumber="7" pagination="23 - 31" refId="ref13945" refString="30. Hunziker AT, Barboza GE. Estudios sobre Solanaceae XLV. Sobre la presencia de Exodeconus en Argentina y una novedad en Capsicum baccatum. Kurtziana. 1998; 26: 23 - 31." type="journal article" year="1998">30</bibRefCitation>
,
<bibRefCitation id="EFC04B31FFBEFFFA3121FA5E6238FA00" author="Hunziker AT" box="[842,867,1488,1512]" pageId="6" pageNumber="7" refId="ref13982" refString="31. Hunziker AT. Genera Solanacearum. The genera of Solanaceae illustrated, arranged according to a new system. Ruggell: A. R. G. Gantner Verlag K. - G; 2001." type="book" year="2001">31</bibRefCitation>
], and observed in other species (
<taxonomicName id="4C514D43FFBEFFFA36D4FA5E63D6F9E2" authority="Eshbaugh, P. G. Sm. &amp; Nickrent" authorityName="Eshbaugh, P. G. Sm. &amp; Nickrent" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="tovarii">
<emphasis id="B925EAD2FFBEFFFA36D4FA5E658BFA0F" box="[1215,1232,1488,1511]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA36B0FA5E647BFA0F" box="[1243,1312,1488,1511]" italics="true" pageId="6" pageNumber="7">tovarii</emphasis>
Eshbaugh, P.G.Sm. &amp; Nickrent
</taxonomicName>
and
<taxonomicName id="4C514D43FFBEFFFA30AAFA7C6572F9E2" authority="Ruiz &amp; Pav., Barboza" authorityName="Ruiz &amp; Pav., Barboza" box="[705,1065,1522,1546]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="pubescens">
<emphasis id="B925EAD2FFBEFFFA30AAFA7C6389F9E1" box="[705,722,1522,1545]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA30B5FA7C621EF9E2" box="[734,837,1522,1546]" italics="true" pageId="6" pageNumber="7">pubescens</emphasis>
Ruiz &amp; Pav., Barboza
</taxonomicName>
pers. obs.). This character deserves careful field observations to ascertain if short-styled flowers produce fruits.
</paragraph>
<paragraph id="8BEE36C0FFBEFFFA305EF9B66265F950" blockId="6.[533,1536,1245,1893]" pageId="6" pageNumber="7">
As some data are still unknown (e.g. corolla color, fruit and seed characters, and chromosome number) for this species and freshly collected leaf material is not available for DNA extraction, we cannot suggest in which of the different clades of the current phylogeny of
<taxonomicName id="4C514D43FFBEFFFA37ADF9F36309F95F" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFBEFFFA37ADF9F36309F95F" italics="true" pageId="6" pageNumber="7">Capsicum</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFC04B31FFBEFFFA300AF92E6335F950" author="Carrizo Garcia C &amp; Sehr EM &amp; Barfuss MHJ &amp; Barboza GE &amp; Samuel R &amp; Moscone EA" box="[609,622,1696,1720]" pageId="6" pageNumber="7" pagination="35 - 51" refId="ref12646" refString="1. Carrizo Garcia C, Sehr EM, Barfuss MHJ, Barboza GE, Samuel R, Moscone EA et al. Phylogenetic relationships, diversification and expansion of chili peppers (Capsicum, Solanaceae). Ann Bot. 2016; 118: 35 - 51. https: // doi. org / 10.1093 / aob / mcw 079 PMID: 27245634" type="journal article" year="2016">1</bibRefCitation>
] it could be placed.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,792 @@
<document id="8EF0E17E99FB465108BAFFB878A88844" ID-DOI="10.1371/journal.pone.0209792" ID-ISSN="1932-6203" ID-PMC="PMC6334993" ID-PubMed="30650102" ID-Zenodo-Dep="12630921" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1719975332143" checkinUser="felipe" docAuthor="Barboza, Gloria E., García, Carolina Carrizo, González, Segundo Leiva, Scaldaferro, Marisel &amp; Reyes, Ximena" docDate="2019" docId="03F887D6FFBEFFF1305EF94C63B1FEB8" docLanguage="en" docName="PLoSONE.14.1.e0209792.pdf" docOrigin="PLoS ONE (e 0209792) 14 (1)" docSource="http://dx.doi.org/10.1371/journal.pone.0209792" docStyle="DocumentStyle:55318D21378FF0528C342C9740A1E1BE.2:PLoSONE.2014-.journal_article" docStyleId="55318D21378FF0528C342C9740A1E1BE" docStyleName="PLoSONE.2014-.journal_article" docStyleVersion="2" docTitle="Capsicum longifolium Barboza &amp; S. Leiva 2019, sp. nov." docType="treatment" docVersion="1" lastPageNumber="13" masterDocId="FFC1FFAEFFB8FFFC326BFF8E615BFFE8" masterDocTitle="Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus" masterLastPageNumber="26" masterPageNumber="1" pageNumber="7" updateTime="1723725765965" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
<mods:mods id="4DE4CD204BD95CAD31046166235D9358" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="03940AFCD31123DD6A6B491B2A118EEC">
<mods:title id="E943083ACE918F76BB09EC56AA77BD72">Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus</mods:title>
</mods:titleInfo>
<mods:name id="2CB54DD3DB3C0E9ED63EDA104A035B08" type="personal">
<mods:role id="C1EDF8E3BA79BDCF33AE5FEEFF10C479">
<mods:roleTerm id="409AD4838B0072F388A580E13518A5CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="01F372134565E9E20C5A952EEA1476BE">Barboza, Gloria E.</mods:namePart>
</mods:name>
<mods:name id="B52EA4E3B0F839BEC4CEF8035A405176" type="personal">
<mods:role id="89EA96B730F7A5B55743BB166ED97E7B">
<mods:roleTerm id="E11742911D24CE297C7834FF6B97E6AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2E5083AAD44641424637633345EFFAAA">García, Carolina Carrizo</mods:namePart>
<mods:affiliation id="54A50EA95A266C5DD8DF84E3A86B8519">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Department of Botany and Biodiversity Research, University of Vienna, Vienna, Austria,</mods:affiliation>
</mods:name>
<mods:name id="53449B631DFCF9FC18EB0829E39D4B8D" type="personal">
<mods:role id="87923DBD797DBA0D11B5DF2FF576581A">
<mods:roleTerm id="238D6C7893E34CB287E2974E5F1643CF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8967D0F97ABE9F03F3C184EB84CE099">González, Segundo Leiva</mods:namePart>
</mods:name>
<mods:name id="DFFB6BD794E7CAD9F8EDC63CCF632F53" type="personal">
<mods:role id="1BD3A016FD52EB211462DC8DAAFBA507">
<mods:roleTerm id="041AFBC9629114390A383D71F2FC609D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B50DEF3E02486556AE059C9C1CE7187E">Scaldaferro, Marisel</mods:namePart>
<mods:affiliation id="3D3878BEB6F0D5FD40ECE0E9371D35E9">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Córdoba, Argentina,</mods:affiliation>
</mods:name>
<mods:name id="7ABEF0F417988307E436D48B55D0CA03" type="personal">
<mods:role id="70C044CFEBF8F028E86BBFC673C5C6DD">
<mods:roleTerm id="9EFAC5F19D80CA76CAA5565510102CF1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4EEB0B13F95622726F1A73F9F173ABE6">Reyes, Ximena</mods:namePart>
<mods:affiliation id="1B2A71CC6A5C86994D1180E33F108B83">Centro de Investigaciones Fitoecogenéticas de Pairumani, Cochabamba, Bolivia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="2EBEB3B1CDB9E17ECDC0FAE68D8F9034">text</mods:typeOfResource>
<mods:relatedItem id="DB93A8CA278580E814D18CA096ECF482" type="host">
<mods:titleInfo id="3065C1B2C8893582C67B501C128C7F5D">
<mods:title id="27BDFBFDE92D0D13247756878A4F351F">PLoS ONE</mods:title>
</mods:titleInfo>
<mods:part id="E2DC13A2591BC3490846530318BC1FA4">
<mods:date id="24A57C9CA621A794D67C52B8B3802FE4">2019</mods:date>
<mods:detail id="E9EF3FEA085260998417C09804D849E0" type="series">
<mods:title id="8DC72FA67489A82F2C0A8F94DDA621D7">e 0209792</mods:title>
</mods:detail>
<mods:detail id="618C05F271B5D506808D4A24F93EB00F" type="pubDate">
<mods:number id="01F7EAC6B09DC95098D4F25A233BA6B3">2019-01-16</mods:number>
</mods:detail>
<mods:detail id="FB698D0BF0FFA820393811FDCEDD8381" type="volume">
<mods:number id="811873797243854DB9041647D6BB8D52">14</mods:number>
</mods:detail>
<mods:detail id="14006A1A09181E081668174D6ADE4BB7" type="issue">
<mods:number id="F79CF97FF6B08A8B9E3D773C2A7ECD33">1</mods:number>
</mods:detail>
<mods:extent id="3BDDA500C98CC36ACAA459D505FB5ED5" unit="page">
<mods:start id="F4DF4589E2EFC78EA0B1D0E8CCCD2E25">1</mods:start>
<mods:end id="F24436D6B77429D7E2A0BEB93A88ACC4">26</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E953352C8467CFE4E149DB79ED277318">
<mods:url id="8E22B9DAFECEB58C94D6FF55EE3ACDF3">http://dx.doi.org/10.1371/journal.pone.0209792</mods:url>
</mods:location>
<mods:classification id="89805D1D744D9AFBE9E8CEC55FBC1910">journal article</mods:classification>
<mods:identifier id="925C4432F14C9D90DDAF2B086C9586C0" type="DOI">10.1371/journal.pone.0209792</mods:identifier>
<mods:identifier id="32DCEAAD430EDDD618B45508D41BFB40" type="ISSN">1932-6203</mods:identifier>
<mods:identifier id="EA6EA665179A7F44353A85134717307C" type="PMC">PMC6334993</mods:identifier>
<mods:identifier id="6F711D8610AE80A8C385D61B54156D42" type="PubMed">30650102</mods:identifier>
<mods:identifier id="CC59869F5A15264B6B7EC771D014B920" type="Zenodo-Dep">12630921</mods:identifier>
</mods:mods>
<treatment id="03F887D6FFBEFFF1305EF94C63B1FEB8" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8" httpUri="http://treatment.plazi.org/id/03F887D6FFBEFFF1305EF94C63B1FEB8" lastPageId="13" lastPageNumber="13" pageId="6" pageNumber="7">
<subSubSection id="C34B654BFFBEFFFA305EF94C6507F932" box="[565,1116,1730,1754]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BEE36C0FFBEFFFA305EF94C6507F932" blockId="6.[533,1536,1245,1893]" box="[565,1116,1730,1754]" pageId="6" pageNumber="7">
<emphasis id="B925EAD2FFBEFFFA305EF94C6507F932" bold="true" box="[565,1116,1730,1754]" pageId="6" pageNumber="7">
<taxonomicName id="4C514D43FFBEFFFA305EF94C62A0F932" ID-CoL="7TBQP" authority="Barboza &amp; S. Leiva, 2019" authorityName="Barboza &amp; S. Leiva" authorityYear="2019" box="[565,1019,1730,1754]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="longifolium" status="sp. nov.">
<emphasis id="B925EAD2FFBEFFFA305EF94C627EF932" bold="true" box="[565,805,1730,1754]" italics="true" pageId="6" pageNumber="7">Capsicum longifolium</emphasis>
Barboza &amp; S. Leiva
</taxonomicName>
,
<taxonomicNameLabel id="A21657A9FFBEFFFA366DF94C6507F932" box="[1030,1116,1730,1754]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFBEFFF4360AF94C6248FEC9" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="8BEE36C0FFBEFFFA360AF94C64A5F932" blockId="6.[533,1536,1245,1893]" box="[1121,1534,1730,1754]" pageId="6" pageNumber="7">[urn:lsid:ipni.org:names: 771925581].</paragraph>
<paragraph id="8BEE36C0FFBEFFFA307EF96B64D3F8AA" blockId="6.[533,1536,1245,1893]" pageId="6" pageNumber="7">
<collectingCounty id="628F4E4CFFBEFFFA307EF96B6316F915" box="[533,589,1765,1789]" pageId="6" pageNumber="7">Type</collectingCounty>
:
<collectingCountry id="F3467650FFBEFFFA303CF96B63E8F915" box="[599,691,1765,1789]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
.
<collectingRegion id="4995F822FFBEFFFA30D7F96B62D7F915" box="[700,908,1765,1789]" country="Ecuador" name="Zamora-Chinchipe" pageId="6" pageNumber="7">Zamora-Chinchipe</collectingRegion>
:
<collectingMunicipality id="6B8AACBAFFBEFFFA31FEF96B6416F915" box="[917,1357,1765,1789]" pageId="6" pageNumber="7">Area of Estación Científica San Francisco</collectingMunicipality>
, road
<location id="8E8E601BFFBEFFFA307EF8866330F8C8" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFBEFFFA307EF8866330F8C8" box="[533,619,1800,1824]" country="Ecuador" county="Type" municipality="Area of Estacion Cientifica San Francisco" name="Zamora" pageId="6" pageNumber="7" stateProvince="Zamora-Chinchipe">LojaZamora</location>
, ca.
<quantity id="4CA99B25FFBEFFFA30F3F8866380F8C8" box="[664,731,1800,1824]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="km" value="35.0">35 km</quantity>
from
<collectingRegion id="4995F822FFBEFFFA3170F8866217F8C8" box="[795,844,1800,1824]" country="Ecuador" name="Loja" pageId="6" pageNumber="7">Loja</collectingRegion>
, transect Q2, 03˚58S, 79˚04W,
<quantity id="4CA99B25FFBEFFFA36CBF88665AAF8C8" box="[1184,1265,1800,1824]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="7" unit="m" value="1900.0">
<elevation id="007CD1F3FFBEFFFA36CBF88665AAF8C8" box="[1184,1265,1800,1824]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="7" unit="m" value="1900.0">1900 m</elevation>
</quantity>
,
<date id="FFEF1000FFBEFFFA3691F8866423F8C8" box="[1274,1400,1800,1824]" pageId="6" pageNumber="7" value="2005-06-12">
<collectingDate id="EFABE9E8FFBEFFFA3691F8866423F8C8" box="[1274,1400,1800,1824]" pageId="6" pageNumber="7" value="2005-06-12">12 Jun 2005</collectingDate>
</date>
(fl, fr),
<collectorName id="26A45316FFBEFFFA37ACF886633EF8AA" pageId="6" pageNumber="7">
<emphasis id="B925EAD2FFBEFFFA37ACF886648EF8F7" box="[1479,1493,1800,1823]" italics="true" pageId="6" pageNumber="7">F</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA378AF88664A8F8F7" box="[1505,1523,1800,1823]" italics="true" pageId="6" pageNumber="7">A</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA307EF8A563C5F8AA" box="[533,670,1834,1858]" italics="true" pageId="6" pageNumber="7">Werner 1548</emphasis>
</collectorName>
(
<typeStatus id="54EA8862FFBEFFFA30C1F8A46250F8AA" box="[682,779,1834,1858]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
, QCA [QCA-160608];
<typeStatus id="54EA8862FFBEFFFA3195F8A4650EF8AA" box="[1022,1109,1834,1858]" pageId="6" pageNumber="7" type="isotype">isotypes</typeStatus>
,
<collectingRegion id="4995F822FFBEFFFA3634F8A465C4F8AA" box="[1119,1183,1834,1858]" country="Ecuador" name="Loja" pageId="6" pageNumber="7">LOJA</collectingRegion>
, NY [NY-01130066]).
</paragraph>
<paragraph id="8BEE36C0FFBEFFFA305EF8C363EFF88D" blockId="6.[533,1536,1245,1893]" box="[565,692,1869,1893]" pageId="6" pageNumber="7">
<figureCitation id="136A2A45FFBEFFFA305EF8C36328F88D" box="[565,627,1869,1893]" captionStart="Fig 3" captionStartId="8.[379,407,203,222]" captionTargetId="figure-1@7.[379,1535,208,1869]" captionTargetPageId="7" captionText="Fig 3. Capsicum longifolium Barboza &amp; S. Leiva. (A) Flowering and fruiting branch. (B) Flower. (C) Opened corolla. (D, E, F) Anther, ventral, lateral and dorsal view, respectively.(G) Gynoecium.(H) Ovary in cross section. (I) Fruit. (J) Seed. Drawn by S. Leiva González." pageId="6" pageNumber="7">Figs 3</figureCitation>
and
<figureCitation id="136A2A45FFBEFFFA30CCF8C363EFF88D" box="[679,692,1869,1893]" captionStart="Fig 4" captionStartId="10.[333,361,203,222]" captionTargetId="figure-1@9.[333,1536,208,1872]" captionTargetPageId="9" captionText="Fig 4. Capsicum longifolium Barboza &amp; S. Leiva. (A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza." pageId="6" pageNumber="7">4</figureCitation>
</paragraph>
<caption id="DF2E6648FFB0FFF43310FF4564AFFF1F" pageId="8" pageNumber="9" startId="8.[379,407,203,222]" targetBox="[385,1531,210,1867]" targetPageId="7" targetType="figure">
<paragraph id="8BEE36C0FFB0FFF43310FF4564AFFF1F" blockId="8.[379,1524,202,248]" pageId="8" pageNumber="9">
<emphasis id="B925EAD2FFB0FFF43310FF456279FF36" bold="true" box="[379,802,202,222]" pageId="8" pageNumber="9">
Fig 3.
<taxonomicName id="4C514D43FFB0FFF433D8FF456279FF36" authority="Barboza &amp; S. Leiva." authorityName="Barboza &amp; S. Leiva." box="[435,802,202,222]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB0FFF433D8FF456328FF35" bold="true" box="[435,627,202,222]" italics="true" pageId="8" pageNumber="9">Capsicum longifolium</emphasis>
Barboza &amp; S. Leiva.
</taxonomicName>
</emphasis>
(A) Flowering and fruiting branch. (B) Flower. (C) Opened corolla. (D, E, F) Anther, ventral, lateral and dorsal view, respectively.(G) Gynoecium.(H) Ovary in cross section. (I) Fruit. (J) Seed. Drawn by S. Leiva González.
</paragraph>
</caption>
<paragraph id="8BEE36C0FFB0FFF43310FE806248FEC9" blockId="8.[379,787,270,289]" box="[379,787,270,289]" pageId="8" pageNumber="9">https://doi.org/10.1371/journal.pone.0209792.g003</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB0FFF4305EFEC462A8FE4F" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BEE36C0FFB0FFF4305EFEC462A8FE4F" blockId="8.[533,1536,329,1879]" pageId="8" pageNumber="9">
<emphasis id="B925EAD2FFB0FFF4305EFEC463F8FE8A" bold="true" box="[565,675,330,354]" pageId="8" pageNumber="9">Diagnosis</emphasis>
. Similar to
<taxonomicName id="4C514D43FFB0FFF43176FEC465EBFE8A" authority="(Miers) Kuntze" authorityName="Kuntze" baseAuthorityName="Miers" box="[797,1200,329,354]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="dimorphum">
<emphasis id="B925EAD2FFB0FFF43176FEC46553FE89" box="[797,1032,329,353]" italics="true" pageId="8" pageNumber="9">Capsicum dimorphum</emphasis>
(Miers) Kuntze
</taxonomicName>
but differing in the long and narrow coriaceous leaves, the number of flowers (39), the unequal thick calyx appendages, and the glabrous vegetative organs and calyx.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB0FFF4305EFE3F6333F98C" pageId="8" pageNumber="9" type="description">
<paragraph id="8BEE36C0FFB0FFF4305EFE3F6333F98C" blockId="8.[533,1536,329,1879]" pageId="8" pageNumber="9">
<emphasis id="B925EAD2FFB0FFF4305EFE3F63E3FE21" bold="true" box="[565,696,433,457]" pageId="8" pageNumber="9">Description</emphasis>
. Scandent shrubs (0.60)
<quantity id="4CA99B25FFB0FFF431D5FE3F657BFE21" box="[958,1056,433,457]" metricMagnitude="0" metricUnit="m" metricValue="2.2" metricValueMax="3.0" metricValueMin="1.4" pageId="8" pageNumber="9" unit="m" value="2.2" valueMax="3.0" valueMin="1.4">1.403 m</quantity>
tall, laxly branched, slightly plagiotropic. Young stems green, fragile, glabrous, striate with abundant ovoid and light dark lenticels; bark of older stems dark green, glabrous, striate with sparse white lenticels. Sympodial units difoliate, geminate, the leaf pair markedly anisophyllous in size and shape. Leaves simple, coriaceous, slightly discolorous, adaxial surface dark green and shiny, abaxial surface light green and opaque, glabrous on both surfaces and margins; the larger leaves with blades (7) 8.517 (18) cm long, (0.8)
<quantity id="4CA99B25FFB0FFF4308BFD0F621AFD71" box="[736,833,641,665]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="8" pageNumber="9" unit="cm" value="1.75" valueMax="2.5" valueMin="1.0">12.5 cm</quantity>
wide, narrowly elliptic (ratio 610.8), major veins (11)13-17 on each side of midvein, base asymmetric and attenuate, margin entire, apex acuminate; petioles 0.2 (0.51.4) cm long, glabrous; the minor leaves
<quantity id="4CA99B25FFB0FFF43676FD4965CBFD37" box="[1053,1168,711,735]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="5.7" metricValueMin="2.5" pageId="8" pageNumber="9" unit="cm" value="4.1" valueMax="5.7" valueMin="2.5">2.55.7 cm</quantity>
long,
<quantity id="4CA99B25FFB0FFF436BAFD496444FD37" box="[1233,1311,711,735]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="8" pageNumber="9" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">12 cm</quantity>
wide (ratio 1.784), ovate or broadly elliptic, major veins 45 on each side of midvein, base short attenuate, sometimes asymmetric, margin entire, apex obtuse; petioles
<quantity id="4CA99B25FFB0FFF43630FC826595FCCC" box="[1115,1230,780,804]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="8" pageNumber="9" unit="cm" value="0.3" valueMax="0.5" valueMin="0.1">0.10.5 cm</quantity>
long, glabrous. Flowers in fascicles of 37 (9) on a short shoot leaving evident scars when fallen, rarely solitary; flowering pedicels green, filiform, terete, pendent, slightly curved, not geniculate at anthesis, widening to the apex,
<quantity id="4CA99B25FFB0FFF43012FCFA63B6FC64" box="[633,749,884,908]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="8" pageNumber="9" unit="cm" value="0.55" valueMax="0.8" valueMin="0.3">0.30.8 cm</quantity>
long, glabrous. Flower buds ovoid, yellow or purplish yellow. Calyx
<quantity id="4CA99B25FFB0FFF437D4FCFA631AFC47" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="8" pageNumber="9" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.53 mm</quantity>
long,
<quantity id="4CA99B25FFB0FFF430E9FC1963B5FC47" box="[642,750,919,943]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="8" pageNumber="9" unit="mm" value="2.9" valueMax="3.0" valueMin="2.8">2.83 mm</quantity>
wide, cup-shaped, very thin, transparent, light green or greenish purple, the margin truncate, glabrous, with 23 thick appendages like triangular-compressed wings,
<quantity id="4CA99B25FFB0FFF4307EFC5263DAFC1C" box="[533,641,988,1012]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="8" pageNumber="9" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 mm</quantity>
long,
<quantity id="4CA99B25FFB0FFF430A9FC52621BFC1C" box="[706,832,988,1012]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="8" pageNumber="9" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
wide, green or purple, glabrous. Corolla
<quantity id="4CA99B25FFB0FFF4369BFC526407FC1C" box="[1264,1372,988,1012]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="8.5" metricValueMin="6.0" pageId="8" pageNumber="9" unit="mm" value="7.25" valueMax="8.5" valueMin="6.0">68.5 mm</quantity>
long,
<quantity id="4CA99B25FFB0FFF437F6FC52631AFBFF" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="8" pageNumber="9" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">811 mm</quantity>
diam, stellate-campanulate, thick, entirely yellow or yellow with red-brown coloration at margin lobes or inside, without interpetalar tissue; tube (3)
<quantity id="4CA99B25FFB0FFF436E2FBAF65B9FBD1" box="[1161,1250,1057,1081]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="9" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long, glabrous inside and outside; lobes 33.5 (4) mm long, ca.
<quantity id="4CA99B25FFB0FFF431F4FBCA6286FBB4" box="[927,989,1092,1116]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="mm" value="3.0">3 mm</quantity>
wide, broadly ovate, erect or patent, glabrous adaxially and abaxially, the tips papillose and cucullate. Stamens 5, equal, filaments equal,
<quantity id="4CA99B25FFB0FFF4307EFB0763DAFB49" box="[533,641,1161,1185]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="8" pageNumber="9" unit="mm" value="2.3" valueMax="2.6" valueMin="2.0">22.6 mm</quantity>
long, white or red-brown, glabrous, inserted on the corolla ca.
<quantity id="4CA99B25FFB0FFF43770FB076401FB49" box="[1307,1370,1161,1185]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="9" unit="mm" value="2.0">2 mm</quantity>
from the base, with inconspicuous auricles at point of insertion; anthers
<quantity id="4CA99B25FFB0FFF4361DFB2265B5FB2C" box="[1142,1262,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="2.375" metricValueMax="2.75" metricValueMin="2.0" pageId="8" pageNumber="9" unit="mm" value="2.375" valueMax="2.75" valueMin="2.0">22.75 mm</quantity>
long, not connivent, elliptic, purplish white or brown. Ovary
<quantity id="4CA99B25FFB0FFF431D4FB416566FB0F" box="[959,1085,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="8" pageNumber="9" unit="mm" value="1.7" valueMax="1.8" valueMin="1.6">1.61.8 mm</quantity>
long,
<quantity id="4CA99B25FFB0FFF43615FB416594FB0F" box="[1150,1231,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="mm" value="1.2">1.2 mm</quantity>
diam, subglobose, white or light green, glabrous; nectary
<quantity id="4CA99B25FFB0FFF43125FB7C6296FAE2" box="[846,973,1266,1290]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="8" pageNumber="9" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.30.5 mm</quantity>
tall, white; style
<quantity id="4CA99B25FFB0FFF43610FB7C65BCFAE2" box="[1147,1255,1266,1290]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" metricValueMax="5.8" metricValueMin="5.0" pageId="8" pageNumber="9" unit="mm" value="5.4" valueMax="5.8" valueMin="5.0">55.8 mm</quantity>
long, white and lilac at the apex, widening distally, glabrous; stigma
<quantity id="4CA99B25FFB0FFF43186FA9A6565FAC4" box="[1005,1086,1300,1324]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="mm" value="0.3">0.3 mm</quantity>
long,
<quantity id="4CA99B25FFB0FFF43614FA9A65A6FAC4" box="[1151,1277,1300,1324]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="8" pageNumber="9" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.20.4 mm</quantity>
wide, light green, somewhat bilobed. Berry
<quantity id="4CA99B25FFB0FFF43080FAB96204FAA7" box="[747,863,1335,1359]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="8" pageNumber="9" unit="cm" value="1.05" valueMax="1.3" valueMin="0.8">0.81.3 cm</quantity>
diam, globose, slightly flattened at the apex, green when immature, orange at maturity, glabrous, not pungent, the pericarp lacking giant cells (endocarp smooth) and stone cells; fruiting pedicels
<quantity id="4CA99B25FFB0FFF431A6FAF26576FA7C" box="[973,1069,1404,1428]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="8" pageNumber="9" unit="cm" value="1.3" valueMax="1.6" valueMin="1.0">11.6 cm</quantity>
long, pendent, terete, widened distally; the fruiting calyx persistent, non-accrescent,
<quantity id="4CA99B25FFB0FFF431A2FA11656EFA5F" box="[969,1077,1439,1463]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="8" pageNumber="9" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">45.5 mm</quantity>
diam, discoid, green-purple or green, the appendages spreading or reflexed, short and wide (
<quantity id="4CA99B25FFB0FFF43645FA4C65C7FA32" box="[1070,1180,1474,1498]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.8" metricValueMin="2.0" pageId="8" pageNumber="9" unit="mm" value="2.4" valueMax="2.8" valueMin="2.0">22.8 mm</quantity>
long,
<quantity id="4CA99B25FFB0FFF436B7FA4C6400FA32" box="[1244,1371,1474,1498]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="8" pageNumber="9" unit="mm" value="2.5" valueMax="2.6" valueMin="2.4">2.42.6 mm</quantity>
wide at base) or long and more slender (
<quantity id="4CA99B25FFB0FFF43144FA6A62F4FA14" box="[815,943,1508,1532]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="8" pageNumber="9" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.55.5 mm</quantity>
long, ca.
<quantity id="4CA99B25FFB0FFF43679FA6A6538FA14" box="[1042,1123,1508,1532]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="mm" value="1.5">1.5 mm</quantity>
wide at base), fleshy and subulate. Seeds ca. 24 per fruit,
<quantity id="4CA99B25FFB0FFF43097F9896221F9F7" box="[764,890,1543,1567]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.3" metricValueMin="1.7" pageId="8" pageNumber="9" unit="mm" value="2.0" valueMax="2.3" valueMin="1.7">1.72.3 mm</quantity>
long,
<quantity id="4CA99B25FFB0FFF431D0F9896562F9F7" box="[955,1081,1543,1567]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.2" metricValueMin="1.7" pageId="8" pageNumber="9" unit="mm" value="1.95" valueMax="2.2" valueMin="1.7">1.72.2 mm</quantity>
wide, not compressed, obconic, black, the surface reticulate, cells rectangular or polygonal in shape, lateral walls straight or slightly sinuate.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB0FFF6305EF9E164EFFE7C" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8BEE36C0FFB0FFF4305EF9E16309F8BF" blockId="8.[533,1536,329,1879]" pageId="8" pageNumber="9">
<emphasis id="B925EAD2FFB0FFF4305EF9E16211F96F" bold="true" box="[565,842,1647,1671]" pageId="8" pageNumber="9">Distribution and ecology</emphasis>
. Endemic to northern
<collectingCountry id="F3467650FFB0FFF43657F9E16535F96F" box="[1084,1134,1647,1671]" name="Peru" pageId="8" pageNumber="9">Peru</collectingCountry>
(
<collectingRegion id="4995F822FFB0FFF43610F9E165B4F96F" box="[1147,1263,1647,1671]" country="Peru" name="Amazonas" pageId="8" pageNumber="9">Amazonas</collectingRegion>
,
<collectingRegion id="4995F822FFB0FFF43692F9E16430F96F" box="[1273,1387,1647,1671]" country="Peru" name="Cajamarca" pageId="8" pageNumber="9">Cajamarca</collectingRegion>
and
<collectingRegion id="4995F822FFB0FFF437F4F9E16481F96F" box="[1439,1498,1647,1671]" country="Peru" name="Piura" pageId="8" pageNumber="9">Piura</collectingRegion>
) and southern
<collectingCountry id="F3467650FFB0FFF430C3F91F6259F941" box="[680,770,1681,1705]" name="Ecuador" pageId="8" pageNumber="9">Ecuador</collectingCountry>
(
<collectingRegion id="4995F822FFB0FFF43164F91F6284F941" box="[783,991,1681,1705]" country="Ecuador" name="Zamora-Chinchipe" pageId="8" pageNumber="9">Zamora-Chinchipe</collectingRegion>
) (
<figureCitation id="136A2A45FFB0FFF4319DF91F6571F941" box="[1014,1066,1681,1705]" captionStart="Fig 2" captionStartId="6.[533,561,1150,1169]" captionTargetBox="[533,1320,208,1128]" captionTargetId="figure-278@6.[533,1320,208,1129]" captionTargetPageId="6" captionText="Fig 2. Distribution of Capsicum species. https://doi.org/10.1371/journal.pone.0209792.g002" figureDoi="http://doi.org/10.5281/zenodo.12630923" httpUri="https://zenodo.org/record/12630923/files/figure.png" pageId="8" pageNumber="9">Fig 2</figureCitation>
), growing in montane wet forests at mid elevations (
<quantity id="4CA99B25FFB0FFF430E7F93A6244F924" box="[652,799,1716,1740]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="8" pageNumber="9" unit="m" value="2000.0" valueMax="2200.0" valueMin="1800.0">18002200 m</quantity>
), associated with other
<taxonomicName id="4C514D43FFB0FFF4367CF93A65D0F924" box="[1047,1163,1716,1740]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
shrubs (
<taxonomicName id="4C514D43FFB0FFF4368CF93A6389F907" authority="(Dammer) Hunz." authorityName="Hunz." baseAuthorityName="Dammer" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="geminifolium">
<emphasis id="B925EAD2FFB0FFF4368CF93A64BBF924" box="[1255,1504,1716,1740]" italics="true" pageId="8" pageNumber="9">Capsicum geminifolium</emphasis>
(Dammer) Hunz.
</taxonomicName>
,
<taxonomicName id="4C514D43FFB0FFF430B7F9586237F907" box="[732,876,1750,1775]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B925EAD2FFB0FFF430B7F9586262F906" box="[732,825,1750,1774]" italics="true" pageId="8" pageNumber="9">Solanum</emphasis>
spp.
</taxonomicName>
, and
<taxonomicName id="4C514D43FFB0FFF431CFF9596578F907" box="[932,1059,1751,1775]" class="Magnoliopsida" family="Solanaceae" genus="Deprea" kingdom="Plantae" order="Solanales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B925EAD2FFB0FFF431CFF95962B4F906" box="[932,1007,1751,1774]" italics="true" pageId="8" pageNumber="9">Deprea</emphasis>
spp.
</taxonomicName>
),
<taxonomicName id="4C514D43FFB0FFF4365EF95865E0F907" authority="Sm." authorityName="Sm." box="[1077,1211,1750,1775]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB0FFF4365EF95865D7F906" box="[1077,1164,1750,1774]" italics="true" pageId="8" pageNumber="9">Cyathea</emphasis>
Sm.
</taxonomicName>
(
<taxonomicName id="4C514D43FFB0FFF436ACF959640AF907" box="[1223,1361,1751,1775]" class="Polypodiopsida" family="Cyatheaceae" kingdom="Plantae" order="Cyatheales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Cyatheaceae</taxonomicName>
),
<taxonomicName id="4C514D43FFB0FFF43709F9596300F8F9" authority="Ruiz &amp; Pav." authorityName="Ruiz &amp; Pav." class="Magnoliopsida" family="Melastomataceae" genus="Miconia" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB0FFF43709F95964E2F906" box="[1378,1465,1751,1774]" italics="true" pageId="8" pageNumber="9">Miconia</emphasis>
Ruiz &amp; Pav.
</taxonomicName>
(
<taxonomicName id="4C514D43FFB0FFF43003F977627EF8F9" box="[616,805,1785,1809]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
),
<taxonomicName id="4C514D43FFB0FFF4315EF9746237F8F9" box="[821,876,1786,1809]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB0FFF4315EF9746237F8F9" box="[821,876,1786,1809]" italics="true" pageId="8" pageNumber="9">Piper</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C514D43FFB0FFF431F8F9776550F8F9" box="[915,1035,1785,1809]" class="Magnoliopsida" family="Piperaceae" kingdom="Plantae" order="Piperales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Piperaceae</taxonomicName>
),
<taxonomicName id="4C514D43FFB0FFF43677F97465FDF8F9" authority="Aubl." authorityName="Aubl." box="[1052,1190,1785,1809]" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB0FFF43677F974653EF8F9" box="[1052,1125,1786,1809]" italics="true" pageId="8" pageNumber="9">Ocotea</emphasis>
Aubl.
</taxonomicName>
(
<taxonomicName id="4C514D43FFB0FFF436D8F977647FF8F9" box="[1203,1316,1785,1809]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
),
<taxonomicName id="4C514D43FFB0FFF4375FF97764AFF8F9" authority="Schott" authorityName="Schott" box="[1332,1524,1785,1809]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB0FFF4375FF97764F0F8F9" box="[1332,1451,1785,1809]" italics="true" pageId="8" pageNumber="9">Anthurium</emphasis>
Schott
</taxonomicName>
(
<taxonomicName id="4C514D43FFB0FFF43076F8926323F8DC" box="[541,632,1820,1844]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
), amongst other shrubs and trees. It grows in the interior of primary forest in shady areas.
</paragraph>
<caption id="DF2E6648FFB2FFF63326FF45655AFEC2" pageId="10" pageNumber="11" startId="10.[333,361,203,222]" targetBox="[334,1536,212,1872]" targetPageId="9" targetType="figure">
<paragraph id="8BEE36C0FFB2FFF63326FF45655AFEC2" blockId="10.[333,1533,202,298]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF63326FF4563A8FF36" bold="true" box="[333,755,202,222]" pageId="10" pageNumber="11">
Fig 4.
<taxonomicName id="4C514D43FFB2FFF633EFFF4563A8FF36" authority="Barboza &amp; S. Leiva." authorityName="Barboza &amp; S. Leiva." box="[388,755,202,222]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB2FFF633EFFF45631FFF35" bold="true" box="[388,580,202,222]" italics="true" pageId="10" pageNumber="11">Capsicum longifolium</emphasis>
Barboza &amp; S. Leiva.
</taxonomicName>
</emphasis>
(A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza.
</paragraph>
</caption>
<paragraph id="8BEE36C0FFB2FFF63326FECE63BEFEBB" blockId="10.[333,741,320,339]" box="[333,741,320,339]" pageId="10" pageNumber="11">https://doi.org/10.1371/journal.pone.0209792.g004</paragraph>
<paragraph id="8BEE36C0FFB2FFF6305EFEF264EFFE7C" blockId="10.[533,1535,380,1895]" box="[565,1460,380,404]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF6305EFEF263F1FE7C" bold="true" box="[565,682,380,404]" pageId="10" pageNumber="11">Phenology</emphasis>
. Flowering and fruiting from December to August, and probably all year.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB2FFF6305EFE11639AFD07" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8BEE36C0FFB2FFF6305EFE11651DFE31" blockId="10.[533,1535,380,1895]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF6305EFE1163F7FE5F" bold="true" box="[565,684,415,439]" pageId="10" pageNumber="11">Etymology</emphasis>
—The species epithet refers to the shape of the leaves, which are the longest and narrowest elliptic leaves known thus far in the genus.
</paragraph>
<paragraph id="8BEE36C0FFB2FFF6305EFE6A639AFD07" blockId="10.[533,1535,380,1895]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF6305EFE6A62F3FE14" bold="true" box="[565,936,484,508]" pageId="10" pageNumber="11">Species Conservation Assessment</emphasis>
. Following the IUCN Criteria (IUCN 2017), we suggest
<taxonomicName id="4C514D43FFB2FFF6307EFD8963F1FDF6" box="[533,682,518,543]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB2FFF6307EFD89637DFDF6" box="[533,550,519,542]" italics="true" pageId="10" pageNumber="11">C</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63059FD8863F1FDF6" box="[562,682,518,542]" italics="true" pageId="10" pageNumber="11">longifolium</emphasis>
</taxonomicName>
deserves a status of Endangered. The extent of occurrence is calculated to be
<quantity id="4CA99B25FFB2FFF6307EFDA76325FDA9" box="[533,638,553,577]" metricMagnitude="7" metricUnit="m" metricValue="1.9207" pageId="10" pageNumber="11" unit="km" value="19207.0">19207 km</quantity>
2 (Criterion B1 &lt;
<quantity id="4CA99B25FFB2FFF63154FDA762F2FDA9" box="[831,937,553,577]" metricMagnitude="7" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="km" value="20000.0">20000 km</quantity>
2, Vulnerable), the area of occupancy,
<quantity id="4CA99B25FFB2FFF63756FDA76424FDA9" box="[1341,1407,553,577]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="km" value="24.0">24 km</quantity>
2 (Criterion B2 &lt;
<quantity id="4CA99B25FFB2FFF63039FDC263F8FD8C" box="[594,675,588,612]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="km" value="500.0">500 km</quantity>
2, Endangered). Although the species has been collected many times in the last 12 years, in San Francisco Biological Reserve (SFBR,
<collectingRegion id="4995F822FFB2FFF6364FFDE165A8FD6F" box="[1060,1267,623,647]" country="Ecuador" name="Zamora-Chinchipe" pageId="10" pageNumber="11">Zamora-Chinchipe</collectingRegion>
,
<collectingCountry id="F3467650FFB2FFF63697FDE16402FD6F" box="[1276,1369,623,647]" name="Ecuador" pageId="10" pageNumber="11">Ecuador</collectingCountry>
), it is known from only other 5 locations (Criterion B1a
<emphasis id="B925EAD2FFB2FFF631B0FD1F62ABFD41" bold="true" box="[987,1008,657,681]" pageId="10" pageNumber="11"><EFBFBD></emphasis>
10, Vulnerable), in areas not included in a National System of Protected Areas which would indicate a risk to the quality of its habitat (Criterion B2b).
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB2FFF6305EFD7764FEFB84" pageId="10" pageNumber="11" type="description">
<paragraph id="8BEE36C0FFB2FFF6305EFD776475FBCF" blockId="10.[533,1535,380,1895]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF6305EFD7763FCFCF9" bold="true" box="[565,679,761,785]" pageId="10" pageNumber="11">Karyology</emphasis>
. This taxon possesses a 2n = 2x = 26 karyotype with
<quantity id="4CA99B25FFB2FFF636B8FD7765A0FCF9" box="[1235,1275,761,785]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="11" unit="m" value="9.0">9 m</quantity>
pairs (19) of decreasing but rather similar size, 3 sm pairs (1012), and one st pair (13) (
<figureCitation id="136A2A45FFB2FFF636B5FC926449FCDC" box="[1246,1298,796,820]" captionStart="Fig 5" captionStartId="11.[533,561,1389,1408]" captionTargetBox="[533,1335,208,1366]" captionTargetId="figure-144@11.[533,1335,208,1367]" captionTargetPageId="11" captionText="Fig 5. Somatic metaphase chromosomes and ideogram of Capsicum longifolium. (A) Methaphase chromosomes. (B) Ideogram. Solid black blocks or dots denote CMA+/DAPI- (NOR) or CMA+/DAPIo (terminal and intercalary) heterochromatic bands. The NOR is indicated as a separate block. https://doi.org/10.1371/journal.pone.0209792.g005" figureDoi="http://doi.org/10.5281/zenodo.12630925" httpUri="https://zenodo.org/record/12630925/files/figure.png" pageId="10" pageNumber="11">Fig 5</figureCitation>
, S
<figureCitation id="136A2A45FFB2FFF63746FC92646CFCDC" box="[1325,1335,796,820]" captionStart="Fig 1" captionStartId="5.[454,482,203,222]" captionTargetId="figure-1@4.[454,1536,208,1872]" captionTargetPageId="4" captionText="Fig 1. Capsicum benoistii Hunz. ex Barboza. (A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury." pageId="10" pageNumber="11">1</figureCitation>
Table). Pair 10 (sm) is satellited.
<specimenCount id="9D57FD49FFB2FFF630A6FCB16261FCBF" box="[717,826,831,855]" count="2" pageId="10" pageNumber="11" type="generic" typeStatus="types">Two types</specimenCount>
of constitutive heterochromatin are found in this taxon, GC-rich heterochromatin (
<collectionCode id="ED40AE05FFB2FFF630BDFCEF624EFC91" box="[726,789,865,889]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12899" name="Crayford Manor House Adult Education Centre" pageId="10" pageNumber="11" type="Herbarium">CMA</collectionCode>
+/DAPI-) and moderately GC-rich heterochromatin (
<collectionCode id="ED40AE05FFB2FFF63725FCEF64D7FC91" box="[1358,1420,865,889]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12899" name="Crayford Manor House Adult Education Centre" pageId="10" pageNumber="11" type="Herbarium">CMA</collectionCode>
+/DAPIo). The fluorescent banding pattern is quite simple, with most of the chromosomes having similar-sized small terminal bands, except for pairs 8, 12 and 13, which are not banded, an intercalary band on the long arm of pair 3, all of them carrying moderately GC-rich heterochromatin, and the large heterochromatic nucleolar organizer region (
<collectionCode id="ED40AE05FFB2FFF636EAFC6265E0FBEC" box="[1153,1211,1004,1028]" pageId="10" pageNumber="11">NOR</collectionCode>
)-associated block on the short arm of satellite pair 10, bearing GC-rich heterochromatin (
<figureCitation id="136A2A45FFB2FFF636EEFB8165E2FBCF" box="[1157,1209,1039,1063]" captionStart="Fig 5" captionStartId="11.[533,561,1389,1408]" captionTargetBox="[533,1335,208,1366]" captionTargetId="figure-144@11.[533,1335,208,1367]" captionTargetPageId="11" captionText="Fig 5. Somatic metaphase chromosomes and ideogram of Capsicum longifolium. (A) Methaphase chromosomes. (B) Ideogram. Solid black blocks or dots denote CMA+/DAPI- (NOR) or CMA+/DAPIo (terminal and intercalary) heterochromatic bands. The NOR is indicated as a separate block. https://doi.org/10.1371/journal.pone.0209792.g005" figureDoi="http://doi.org/10.5281/zenodo.12630925" httpUri="https://zenodo.org/record/12630925/files/figure.png" pageId="10" pageNumber="11">Fig 5</figureCitation>
, S
<figureCitation id="136A2A45FFB2FFF636BAFB816585FBCF" box="[1233,1246,1039,1063]" captionStart="Fig 4" captionStartId="10.[333,361,203,222]" captionTargetId="figure-1@9.[333,1536,208,1872]" captionTargetPageId="9" captionText="Fig 4. Capsicum longifolium Barboza &amp; S. Leiva. (A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza." pageId="10" pageNumber="11">4</figureCitation>
Table).
</paragraph>
<paragraph id="8BEE36C0FFB2FFF6305EFBBC64FEFB84" blockId="10.[533,1535,380,1895]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF6305EFBBC63C5FBA2" bold="true" box="[565,670,1074,1098]" pageId="10" pageNumber="11">Affinities</emphasis>
.
<taxonomicName id="4C514D43FFB2FFF630C1FBBC62CBFBA1" box="[682,912,1073,1097]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB2FFF630C1FBBC62CBFBA1" box="[682,912,1073,1097]" italics="true" pageId="10" pageNumber="11">Capsicum longifolium</emphasis>
</taxonomicName>
is strongly resolved within the Andean clade, as the first diverging branch within the clade, sister to the rest of the species included in it (
<figureCitation id="136A2A45FFB2FFF6370AFBDA64CDFB84" box="[1377,1430,1108,1132]" captionStart="Fig 6" captionStartId="12.[133,161,1434,1453]" captionTargetBox="[134,1535,208,1411]" captionTargetId="figure-188@12.[133,1536,208,1412]" captionTargetPageId="12" captionText="Fig 6. Bayesian majority-rule consensus tree of Capsicum. Posterior probabilities values indicated by each branch. New species are highlighted in bold-colored letters and the clade to which they belong is indicated. https://doi.org/10.1371/journal.pone.0209792.g006" figureDoi="http://doi.org/10.5281/zenodo.12630927" httpUri="https://zenodo.org/record/12630927/files/figure.png" pageId="10" pageNumber="11">Fig 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFB2FFF1305EFBF963B1FEB8" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BEE36C0FFB2FFF6305EFBF965EAF951" blockId="10.[533,1535,380,1895]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF6305EFBF962C3FB67" bold="true" box="[565,920,1143,1167]" pageId="10" pageNumber="11">Additional specimens examined</emphasis>
.
<materialsCitation id="3B393C9DFFB2FFF631CFFBF96405FB3C" collectingDate="2008-09-22" collectionCode="HUSA, USM" collectorName="Quipuscoa" country="Peru" county="Rodriguez de Mendoza" elevation="2118" location="Cordillera" municipality="Omia" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Amazonas">
<emphasis id="B925EAD2FFB2FFF631CFFBF96531FB67" bold="true" box="[932,1130,1143,1167]" pageId="10" pageNumber="11">
<collectingCountry id="F3467650FFB2FFF631CFFBF962B1FB67" box="[932,1002,1143,1167]" name="Peru" pageId="10" pageNumber="11">PERU</collectingCountry>
.
<collectingRegion id="4995F822FFB2FFF6319EFBF96531FB67" box="[1013,1130,1143,1167]" country="Peru" name="Amazonas" pageId="10" pageNumber="11">Amazonas</collectingRegion>
</emphasis>
:
<collectingCounty id="628F4E4CFFB2FFF6361DFBF86434FB67" box="[1142,1391,1142,1167]" pageId="10" pageNumber="11">Rodríguez de Mendoza</collectingCounty>
,
<collectingMunicipality id="6B8AACBAFFB2FFF63713FBF964EDFB67" box="[1400,1462,1143,1167]" pageId="10" pageNumber="11">Omia</collectingMunicipality>
, entre la
<location id="8E8E601BFFB2FFF63046FB1463C2FB5A" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFB2FFF63046FB1463C2FB5A" box="[557,665,1178,1202]" country="Peru" county="Rodriguez de Mendoza" municipality="Omia" name="Cordillera" pageId="10" pageNumber="11" stateProvince="Amazonas">Cordillera</location>
y
<location id="8E8E601BFFB2FFF630DAFB1462FBFB5A" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFB2FFF630DAFB1462FBFB5A" box="[689,928,1178,1202]" country="Peru" county="Rodriguez de Mendoza" municipality="Omia" name="Quebrada de Agua Sal" pageId="10" pageNumber="11" stateProvince="Amazonas">Quebrada de Agua Sal</location>
, 06˚2618-06˚254.4S, 77˚1030.8-77˚102.3W, 2457 2118 m,
<date id="FFEF1000FFB2FFF6301BFB3263AEFB3C" box="[624,757,1212,1236]" pageId="10" pageNumber="11" value="2008-09-22">
<collectingDate id="EFABE9E8FFB2FFF6301BFB3263AEFB3C" box="[624,757,1212,1236]" pageId="10" pageNumber="11" value="2008-09-22">22 Sept 2008</collectingDate>
</date>
(fl, fr),
<emphasis id="B925EAD2FFB2FFF6312EFB33620CFB3C" box="[837,855,1213,1236]" italics="true" pageId="10" pageNumber="11">V</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63108FB3262BFFB3B" box="[867,996,1212,1235]" italics="true" pageId="10" pageNumber="11">
<collectorName id="26A45316FFB2FFF63108FB326289FB3B" box="[867,978,1212,1235]" pageId="10" pageNumber="11">Quipuscoa</collectorName>
S
</emphasis>
.
<emphasis id="B925EAD2FFB2FFF6319BFB336547FB3C" box="[1008,1052,1212,1236]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63643FB326500FB3C" box="[1064,1115,1212,1236]" italics="true" pageId="10" pageNumber="11">4374</emphasis>
(
<collectionCode id="ED40AE05FFB2FFF6360CFB3265E9FB3C" box="[1127,1202,1212,1236]" country="Peru" lsid="urn:lsid:biocol.org:col:14917" name="Universidad Nacional de San Agustín de Arequipa" pageId="10" pageNumber="11" type="Herbarium">HUSA</collectionCode>
, HUT, F,
<collectionCode id="ED40AE05FFB2FFF63770FB326403FB3C" box="[1307,1368,1212,1236]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="10" pageNumber="11">USM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B393C9DFFB2FFF63703FB32626EFAF2" collectingDate="2000-03-13" collectionCode="MO" collectorName="Quebrada El Mirador &amp; J. Campos" country="Peru" elevation="2200" location="Huarango" municipality="San Ignacio" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Cajamarca">
<emphasis id="B925EAD2FFB2FFF63703FB326484FB3C" bold="true" box="[1384,1503,1212,1236]" pageId="10" pageNumber="11">
<collectingRegion id="4995F822FFB2FFF63703FB326484FB3C" box="[1384,1503,1212,1236]" country="Peru" name="Cajamarca" pageId="10" pageNumber="11">Cajamarca</collectingRegion>
</emphasis>
:
<collectingMunicipality id="6B8AACBAFFB2FFF6307EFB5163CFFB1F" box="[533,660,1247,1271]" pageId="10" pageNumber="11">San Ignacio</collectingMunicipality>
,
<location id="8E8E601BFFB2FFF630F5FB516257FB1F" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFB2FFF630F5FB516257FB1F" box="[670,780,1247,1271]" country="Peru" municipality="San Ignacio" name="Huarango" pageId="10" pageNumber="11" stateProvince="Cajamarca">Huarango</location>
,
<collectorName id="26A45316FFB2FFF6317DFB5162A6FB1F" box="[790,1021,1247,1271]" pageId="10" pageNumber="11">Quebrada El Mirador</collectorName>
, 05˚1612&quot;S, 78˚4001&quot;W, 2200 m,
<date id="FFEF1000FFB2FFF63702FB5164ABFB1F" box="[1385,1520,1247,1271]" pageId="10" pageNumber="11" value="2000-03-13">
<collectingDate id="EFABE9E8FFB2FFF63702FB5164ABFB1F" box="[1385,1520,1247,1271]" pageId="10" pageNumber="11" value="2000-03-13">13 Mar 2000</collectingDate>
</date>
,
<collectorName id="26A45316FFB2FFF6307EFA8C6324FAF1" box="[533,639,1282,1306]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF6307EFA8C6346FAF1" box="[533,541,1282,1305]" italics="true" pageId="10" pageNumber="11">J</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63041FA8C63EBFAF1" box="[554,688,1281,1305]" italics="true" pageId="10" pageNumber="11">Campos et al</emphasis>
</collectorName>
.
<emphasis id="B925EAD2FFB2FFF630D7FA8F63B4FAF1" box="[700,751,1281,1305]" italics="true" pageId="10" pageNumber="11">6607</emphasis>
(
<collectionCode id="ED40AE05FFB2FFF63097FA8C6277FAF2" box="[764,812,1282,1306]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="10" pageNumber="11" type="Herbarium">MO</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B393C9DFFB2FFF63155FA8C6448FAB7" collectingDate="2003-04-25" collectionCode="USM" collectorName="Carmen de la Frontera &amp; S. M. Baldeon" country="Peru" county="Huancabamba" elevation="2175" location="Santuario Nacional Tabaconas-Namballe" municipality="Rio Samaniego" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Piura">
<emphasis id="B925EAD2FFB2FFF63155FA8C6220FAF2" bold="true" box="[830,891,1282,1306]" pageId="10" pageNumber="11">
<collectingRegion id="4995F822FFB2FFF63155FA8C6220FAF2" box="[830,891,1282,1306]" country="Peru" name="Piura" pageId="10" pageNumber="11">Piura</collectingRegion>
</emphasis>
:
<collectingCounty id="628F4E4CFFB2FFF631E3FA8C657DFAF2" box="[904,1062,1282,1306]" pageId="10" pageNumber="11">Huancabamba</collectingCounty>
, distr.
<collectorName id="26A45316FFB2FFF63607FA8C6404FAF2" box="[1132,1375,1282,1306]" pageId="10" pageNumber="11">Carmen de la Frontera</collectorName>
,
<collectingMunicipality id="6B8AACBAFFB2FFF63702FA8F6309FAD4" pageId="10" pageNumber="11">Río Samaniego</collectingMunicipality>
, margen derecha, zona de amortiguamiento del
<location id="8E8E601BFFB2FFF6363AFAAA6312FAB7" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFB2FFF6363AFAAA6312FAB7" country="Peru" county="Huancabamba" municipality="Rio Samaniego" name="Santuario Nacional Tabaconas-Namballe" pageId="10" pageNumber="11" stateProvince="Piura">Santuario Nacional Tabaconas-Namballe</location>
, 21502200 m,
<date id="FFEF1000FFB2FFF63085FAC9622BFAB7" box="[750,880,1351,1375]" pageId="10" pageNumber="11" value="2003-04-25">
<collectingDate id="EFABE9E8FFB2FFF63085FAC9622BFAB7" box="[750,880,1351,1375]" pageId="10" pageNumber="11" value="2003-04-25">25 Apr 2003</collectingDate>
</date>
(fl, fr),
<collectorName id="26A45316FFB2FFF631D4FAC96514FAB6" box="[959,1103,1350,1375]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF631D4FAC96290FAB6" box="[959,971,1351,1374]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B925EAD2FFB2FFF631BCFAC962B5FAB6" box="[983,1006,1351,1374]" italics="true" pageId="10" pageNumber="11">M</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63191FAC865DAFAB6" box="[1018,1153,1350,1374]" italics="true" pageId="10" pageNumber="11">Baldeón et al</emphasis>
</collectorName>
.
<emphasis id="B925EAD2FFB2FFF636E6FAC8659BFAB6" box="[1165,1216,1350,1374]" italics="true" pageId="10" pageNumber="11">5316</emphasis>
(
<collectionCode id="ED40AE05FFB2FFF636A6FAC96450FAB7" box="[1229,1291,1351,1375]" country="Malaysia" httpUri="http://grbio.org/cool/11pi-mg53" name="Universiti Sains Malaysia" pageId="10" pageNumber="11">USM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B393C9DFFB2FFF63777FAC965FDFA6A" collectionCode="EBSF" collectorName="Estacion Biologica San Francisco" country="Ecuador" location="Zamora-Chinchipe" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Zamora-Chinchipe">
<emphasis id="B925EAD2FFB2FFF63777FAC963B2FA6A" bold="true" pageId="10" pageNumber="11">
<collectingCountry id="F3467650FFB2FFF63777FAC964FBFAB7" box="[1308,1440,1351,1375]" name="Ecuador" pageId="10" pageNumber="11">ECUADOR</collectingCountry>
.
<collectingRegion id="4995F822FFB2FFF6307EFAE463B2FA6A" box="[533,745,1386,1410]" country="Ecuador" name="Zamora-Chinchipe" pageId="10" pageNumber="11">Zamora-Chinchipe</collectingRegion>
</emphasis>
:
<collectorName id="26A45316FFB2FFF6309EFAE76509FA6A" box="[757,1106,1385,1410]" pageId="10" pageNumber="11">Estación Biológica San Francisco</collectorName>
(
<collectionCode id="ED40AE05FFB2FFF63634FAE465C4FA6A" box="[1119,1183,1386,1410]" pageId="10" pageNumber="11">EBSF</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B393C9DFFB2FFF636DBFAE462E1FA2F" collectingDate="2017-05-02" collectionCode="CORD" collectorName="Barboza &amp; Leiva Gonzalez" country="Ecuador" elevation="1830" location="rio San Francisco" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Zamora-Chinchipe">
camino hacia la antena, pasando el
<location id="8E8E601BFFB2FFF630E7FA02621DFA4C" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFB2FFF630E7FA02621DFA4C" box="[652,838,1420,1444]" country="Ecuador" name="rio San Francisco" pageId="10" pageNumber="11" stateProvince="Zamora-Chinchipe">río San Francisco</location>
, 3˚5822.5”S, 79˚0440.9”W, 1830 m,
<date id="FFEF1000FFB2FFF636A5FA026417FA4C" box="[1230,1356,1420,1444]" pageId="10" pageNumber="11" value="2017-05-02">
<collectingDate id="EFABE9E8FFB2FFF636A5FA026417FA4C" box="[1230,1356,1420,1444]" pageId="10" pageNumber="11" value="2017-05-02">2 May 2017</collectingDate>
</date>
(fl),
<emphasis id="B925EAD2FFB2FFF6371EFA0264DCFA4B" box="[1397,1415,1420,1443]" italics="true" pageId="10" pageNumber="11">G</emphasis>
.
<emphasis id="B925EAD2FFB2FFF637F8FA0364F9FA4C" box="[1427,1442,1421,1444]" italics="true" pageId="10" pageNumber="11">E</emphasis>
.
<emphasis id="B925EAD2FFB2FFF637C5FA036328FA2E" italics="true" pageId="10" pageNumber="11">
<collectorName id="26A45316FFB2FFF637C5FA03631DFA2E" pageId="10" pageNumber="11">Barboza</collectorName>
&amp; S
</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63014FA21620CFA2F" box="[639,855,1454,1479]" italics="true" pageId="10" pageNumber="11">
<collectorName id="26A45316FFB2FFF63014FA216245FA2E" box="[639,798,1454,1478]" pageId="10" pageNumber="11">Leiva González</collectorName>
4821
</emphasis>
(
<collectionCode id="ED40AE05FFB2FFF6310FFA2162E8FA2F" box="[868,947,1455,1479]" country="Argentina" lsid="urn:lsid:biocol.org:col:15636" name="Universidad Nacional de Córdoba" pageId="10" pageNumber="11" type="Herbarium">CORD</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B393C9DFFB2FFF631AFFA216493FA2F" box="[964,1480,1454,1479]" collectionCode="HAO" collectorName="Leiva Gonzalez" country="Ecuador" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Zamora-Chinchipe">
at the same place,
<emphasis id="B925EAD2FFB2FFF636E8FA2165D4FA2E" box="[1155,1167,1455,1478]" italics="true" pageId="10" pageNumber="11">S</emphasis>
.
<emphasis id="B925EAD2FFB2FFF636F0FA216428FA2E" box="[1179,1395,1454,1478]" italics="true" pageId="10" pageNumber="11">
<collectorName id="26A45316FFB2FFF636F0FA216460FA2E" box="[1179,1339,1454,1478]" pageId="10" pageNumber="11">Leiva González</collectorName>
6531
</emphasis>
(
<collectionCode id="ED40AE05FFB2FFF637EBFA2164E4FA2F" box="[1408,1471,1455,1479]" country="Peru" lsid="urn:lsid:biocol.org:col:15351" name="Universidad Privada Antenor Orrego" pageId="10" pageNumber="11" type="Herbarium">HAO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B393C9DFFB2FFF6307EFA5F653BF9C7" collectingDate="2017-08-17" collectionCode="HAO" collectorName="G. E. Barboza &amp; S. Leiva Gonzalez" country="Ecuador" elevation="300" location="Atajo" municipality="Rio San Francisco" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Loja">
EBSF, a unos 300 m después del cruzar el
<collectingMunicipality id="6B8AACBAFFB2FFF631A5FA5F65D4FA01" box="[974,1167,1489,1513]" pageId="10" pageNumber="11">Río San Francisco</collectingMunicipality>
, por el camino del
<location id="8E8E601BFFB2FFF6373CFA5F64C8FA01" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFB2FFF6373CFA5F64C8FA01" box="[1367,1427,1489,1513]" country="Ecuador" municipality="Rio San Francisco" name="Atajo" pageId="10" pageNumber="11" stateProvince="Loja">Atajo</location>
, 3˚ 5821.6”S, 79˚0441.4”W, 1839 m,
<date id="FFEF1000FFB2FFF63116FA7A6559F9E4" box="[893,1026,1524,1548]" pageId="10" pageNumber="11" value="2017-08-17">
<collectingDate id="EFABE9E8FFB2FFF63116FA7A6559F9E4" box="[893,1026,1524,1548]" pageId="10" pageNumber="11" value="2017-08-17">17 Aug 2017</collectingDate>
</date>
(fl, fr),
<collectorName id="26A45316FFB2FFF6363AFA7A65BAF9E4" box="[1105,1249,1524,1548]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF6363AFA7A6538F9E3" box="[1105,1123,1524,1547]" italics="true" pageId="10" pageNumber="11">G</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63604FA7B6525F9E4" box="[1135,1150,1525,1548]" italics="true" pageId="10" pageNumber="11">E</emphasis>
.
<emphasis id="B925EAD2FFB2FFF636E1FA7A6456F9E3" box="[1162,1293,1524,1548]" italics="true" pageId="10" pageNumber="11">
Barboza &amp;
<collectorName id="26A45316FFB2FFF6376AFA7A64E2F9E4" box="[1281,1465,1524,1548]" pageId="10" pageNumber="11">
S.
<emphasis id="B925EAD2FFB2FFF63772FA7B6339F9C7" italics="true" pageId="10" pageNumber="11">Leiva González 4846 &amp; 4851</emphasis>
</collectorName>
</emphasis>
</collectorName>
(
<collectingRegion id="4995F822FFB2FFF63004F99963EAF9C7" box="[623,689,1559,1583]" country="Ecuador" name="Loja" pageId="10" pageNumber="11">LOJA</collectingRegion>
, duplicates to be sent to CORD &amp;
<collectionCode id="ED40AE05FFB2FFF63671F9996503F9C7" box="[1050,1112,1559,1583]" country="Peru" lsid="urn:lsid:biocol.org:col:15351" name="Universidad Privada Antenor Orrego" pageId="10" pageNumber="11" type="Herbarium">HAO</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B393C9DFFB2FFF63601F99965FEF99C" collectingDate="2017-08-17" collectionCode="CORD" collectorName="Barboza &amp; Leiva Gonzalez" country="Ecuador" elevation="1888" location="Rio San Francisco" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Loja">
EBSF, después de cruzar el
<location id="8E8E601BFFB2FFF637E2F9996325F9B9" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFB2FFF637E2F9996325F9B9" country="Ecuador" name="Rio San Francisco" pageId="10" pageNumber="11" stateProvince="Loja">Río San Francisco</location>
, 3˚5822.5”S, 79˚0439.3”W, 1888 m,
<date id="FFEF1000FFB2FFF6366CF9B765D7F9B9" box="[1031,1164,1593,1617]" pageId="10" pageNumber="11" value="2017-08-17">
<collectingDate id="EFABE9E8FFB2FFF6366CF9B765D7F9B9" box="[1031,1164,1593,1617]" pageId="10" pageNumber="11" value="2017-08-17">17 Aug 2017</collectingDate>
</date>
(fl, fr),
<emphasis id="B925EAD2FFB2FFF636B0F9B465B6F9B9" box="[1243,1261,1594,1617]" italics="true" pageId="10" pageNumber="11">G</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63692F9B46453F9B9" box="[1273,1288,1594,1617]" italics="true" pageId="10" pageNumber="11">E</emphasis>
.
<emphasis id="B925EAD2FFB2FFF6377FF9B764CCF9B8" box="[1300,1431,1593,1617]" italics="true" pageId="10" pageNumber="11">
<collectorName id="26A45316FFB2FFF6377FF9B76430F9B9" box="[1300,1387,1593,1617]" pageId="10" pageNumber="11">Barboza</collectorName>
&amp; S
</emphasis>
.
<emphasis id="B925EAD2FFB2FFF637C8F9B4625AF99C" italics="true" pageId="10" pageNumber="11">
<collectorName id="26A45316FFB2FFF637C8F9B4632DF99C" pageId="10" pageNumber="11">Leiva González</collectorName>
4849 &amp; 4850
</emphasis>
(
<collectingRegion id="4995F822FFB2FFF63165F9D2620BF99C" box="[782,848,1628,1652]" country="Ecuador" name="Loja" pageId="10" pageNumber="11">LOJA</collectingRegion>
, duplicates to be sent to
<collectionCode id="ED40AE05FFB2FFF6363AF9D265C4F99C" box="[1105,1183,1628,1652]" country="Argentina" lsid="urn:lsid:biocol.org:col:15636" name="Universidad Nacional de Córdoba" pageId="10" pageNumber="11" type="Herbarium">CORD</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B393C9DFFB2FFF636DBF9D2653EF97F" collectingDate="1959-07-14" collectingDateMax="1959-07-19" collectingDateMin="1959-07-14" collectorName="Harling" country="Ecuador" elevation="2100" location="Loxa-Zamora" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Zamora-Chinchipe">
road
<location id="8E8E601BFFB2FFF6368DF9D26421F99C" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFB2FFF6368DF9D26421F99C" box="[1254,1402,1628,1652]" country="Ecuador" name="Loxa-Zamora" pageId="10" pageNumber="11" stateProvince="Zamora-Chinchipe">Loxa-Zamora</location>
,
<locationDeviation id="9A5C2FE2FFB2FFF637EFF9D2633FF97F" location="Tambo" pageId="10" pageNumber="11">5 km W of Tambo</locationDeviation>
, 2100 m,
<date id="FFEF1000FFB2FFF630A3F9F1623EF97F" box="[712,869,1663,1687]" pageId="10" pageNumber="11" value="1959-07-14" valueMax="1959-07-19" valueMin="1959-07-14">
<collectingDate id="EFABE9E8FFB2FFF630A3F9F1623EF97F" box="[712,869,1663,1687]" pageId="10" pageNumber="11" value="1959-07-14" valueMax="1959-07-19" valueMin="1959-07-14">1419 Jul 1959</collectingDate>
</date>
(fl),
<emphasis id="B925EAD2FFB2FFF631FCF9F162F2F97E" box="[919,937,1663,1686]" italics="true" pageId="10" pageNumber="11">G</emphasis>
.
<emphasis id="B925EAD2FFB2FFF631DEF9F06564F97E" box="[949,1087,1662,1686]" italics="true" pageId="10" pageNumber="11">
<collectorName id="26A45316FFB2FFF631DEF9F0655DF97E" box="[949,1030,1662,1686]" pageId="10" pageNumber="11">Harling</collectorName>
5867
</emphasis>
(S)
</materialsCitation>
;
<materialsCitation id="3B393C9DFFB2FFF63604F9F165F6F951" collectingDate="1985-02-02" collectionCode="GB" collectorName="G. Harling &amp; L. Andersson" country="Ecuador" elevation="2700" location="Yangana" municipality="Valladolid" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Zamora-Chinchipe">
above
<collectingMunicipality id="6B8AACBAFFB2FFF636D8F9F1647AF97F" box="[1203,1313,1663,1687]" pageId="10" pageNumber="11">Valladolid</collectingMunicipality>
on road to
<location id="8E8E601BFFB2FFF637F0F9F164A3F97F" LSID="urn:lsid:plazi:treatment:03F887D6FFBEFFF1305EF94C63B1FEB8:8E8E601BFFB2FFF637F0F9F164A3F97F" box="[1435,1528,1663,1687]" country="Ecuador" municipality="Valladolid" name="Yangana" pageId="10" pageNumber="11" stateProvince="Zamora-Chinchipe">Yanganá</location>
, 2700 m,
<date id="FFEF1000FFB2FFF6301BF92F63B9F951" box="[624,738,1697,1721]" pageId="10" pageNumber="11" value="1985-02-02">
<collectingDate id="EFABE9E8FFB2FFF6301BF92F63B9F951" box="[624,738,1697,1721]" pageId="10" pageNumber="11" value="1985-02-02">2 Feb 1985</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFB2FFF6317FF92C62D8F951" box="[788,899,1697,1721]" pageId="10" pageNumber="11">
<emphasis id="B925EAD2FFB2FFF6317FF92C627DF951" box="[788,806,1698,1721]" italics="true" pageId="10" pageNumber="11">G</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63159F92F62EAF951" box="[818,945,1697,1721]" italics="true" pageId="10" pageNumber="11">
Harling &amp;
<collectorName id="26A45316FFB2FFF631C8F92C6570F951" box="[931,1067,1697,1721]" pageId="10" pageNumber="11">
L.
<emphasis id="B925EAD2FFB2FFF631D6F92F6534F951" box="[957,1135,1697,1721]" italics="true" pageId="10" pageNumber="11">Andersson 21464</emphasis>
</collectorName>
</emphasis>
</collectorName>
(
<collectionCode id="ED40AE05FFB2FFF63616F92F65FFF951" box="[1149,1188,1697,1721]" country="Sweden" name="University of Gothenburg" pageId="10" pageNumber="11" type="Herbarium">GB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36C0FFB2FFF7305EF94A6577F8E1" blockId="10.[533,1535,380,1895]" lastBlockId="11.[533,1536,1534,1870]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<taxonomicName id="4C514D43FFB2FFF6305EF94A6240F934" box="[565,795,1732,1756]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB2FFF6305EF94A6240F934" box="[565,795,1732,1756]" italics="true" pageId="10" pageNumber="11">Capsicum longifolium</emphasis>
</taxonomicName>
is unique in the genus in having the longest and narrowest leaves and the striking calyx appendages that arise from the calyx tube as lateral compressed thick expansions or wings (
<figureCitation id="136A2A45FFB2FFF630D7F88762F2F8C9" box="[700,937,1801,1825]" captionStart="Fig 4" captionStartId="10.[333,361,203,222]" captionTargetId="figure-1@9.[333,1536,208,1872]" captionTargetPageId="9" captionText="Fig 4. Capsicum longifolium Barboza &amp; S. Leiva. (A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza." pageId="10" pageNumber="11">Fig 4C4E and 4L4N</figureCitation>
). Apart from that, it is morphologically most similar to
<taxonomicName id="4C514D43FFB2FFF6307EF8A263F4F8AC" box="[533,687,1836,1860]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="dimorphum">
<emphasis id="B925EAD2FFB2FFF6307EF8A2637DF8AB" box="[533,550,1836,1859]" italics="true" pageId="10" pageNumber="11">C</emphasis>
.
<emphasis id="B925EAD2FFB2FFF63059F8A263F4F8AC" box="[562,687,1836,1860]" italics="true" pageId="10" pageNumber="11">dimorphum</emphasis>
</taxonomicName>
with which it shares the shape and color of the corolla, fruit and seeds.
<taxonomicName id="4C514D43FFB2FFF637CBF8A2639BF88E" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB2FFF637CBF8A2639BF88E" italics="true" pageId="10" pageNumber="11">Capsicum longifolium</emphasis>
</taxonomicName>
can be distinguished by having completely glabrous vegetative organs and calyces, long and narrow (ratio 610.8) coriaceous major leaves, flowers in fascicles of 37 (9) on a short shoot and calyces with 23 thick appendages like triangular-compressed wings compared to the pubescent vegetative organs and calyces, the shorter and wider (ratio 45.25) membranaceous major leaves, the solitary or up to 5 axillary flowers, and the toothless calyx or with 3 tiny appendages of
<taxonomicName id="4C514D43FFB3FFF73141F907629FF948" box="[810,964,1672,1697]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="dimorphum">
<emphasis id="B925EAD2FFB3FFF73141F9076260F948" box="[810,827,1673,1696]" italics="true" pageId="11" pageNumber="12">C</emphasis>
.
<emphasis id="B925EAD2FFB3FFF7312CF906629FF948" box="[839,964,1672,1696]" italics="true" pageId="11" pageNumber="12">dimorphum</emphasis>
</taxonomicName>
. Another species of
<taxonomicName id="4C514D43FFB3FFF736F1F9076459F948" box="[1178,1282,1673,1696]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB3FFF736F1F9076459F948" box="[1178,1282,1673,1696]" italics="true" pageId="11" pageNumber="12">Capsicum</emphasis>
</taxonomicName>
sympatric with
<taxonomicName id="4C514D43FFB3FFF737C7F907633AF92B" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB3FFF737C7F90764E6F948" box="[1452,1469,1673,1696]" italics="true" pageId="11" pageNumber="12">C</emphasis>
.
<emphasis id="B925EAD2FFB3FFF737A2F906633AF92B" italics="true" pageId="11" pageNumber="12">longifolium</emphasis>
</taxonomicName>
(especially in SFBR,
<collectingCountry id="F3467650FFB3FFF73156F92562C3F92B" box="[829,920,1707,1731]" name="Ecuador" pageId="11" pageNumber="12">Ecuador</collectingCountry>
) is
<taxonomicName id="4C514D43FFB3FFF731D7F922653FF92B" box="[956,1124,1707,1731]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="geminifolium">
<emphasis id="B925EAD2FFB3FFF731D7F9226296F92B" box="[956,973,1708,1731]" italics="true" pageId="11" pageNumber="12">C</emphasis>
.
<emphasis id="B925EAD2FFB3FFF731B2F925653FF92B" box="[985,1124,1707,1731]" italics="true" pageId="11" pageNumber="12">geminifolium</emphasis>
</taxonomicName>
that has a dense indumentum, long apiculate leaves, longer pedicels (
<quantity id="4CA99B25FFB3FFF73119F94062FDF90E" box="[882,934,1742,1766]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="cm" value="5.0">5 cm</quantity>
long), thin calyx appendages, and funnel-shaped yellow corollas with many purple or maroon spots inside.
</paragraph>
<caption id="DF2E6648FFB3FFF7307EFAE362F6FA34" ID-DOI="http://doi.org/10.5281/zenodo.12630925" ID-Zenodo-Dep="12630925" httpUri="https://zenodo.org/record/12630925/files/figure.png" pageId="11" pageNumber="12" startId="11.[533,561,1389,1408]" targetBox="[533,1335,208,1366]" targetPageId="11" targetType="figure">
<paragraph id="8BEE36C0FFB3FFF7307EFAE3651AFA5A" blockId="11.[533,1510,1388,1500]" pageId="11" pageNumber="12">
<emphasis id="B925EAD2FFB3FFF7307EFAE3658DFA68" bold="true" box="[533,1238,1388,1408]" pageId="11" pageNumber="12">
Fig 5. Somatic metaphase chromosomes and ideogram of
<taxonomicName id="4C514D43FFB3FFF73679FAE2658AFA97" box="[1042,1233,1388,1407]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB3FFF73679FAE2658AFA97" bold="true" box="[1042,1233,1388,1407]" italics="true" pageId="11" pageNumber="12">Capsicum longifolium</emphasis>
</taxonomicName>
.
</emphasis>
(A) Methaphase chromosomes. (B) Ideogram. Solid black blocks or dots denote CMA+/DAPI- (NOR) or CMA+/DAPIo (terminal and intercalary) heterochromatic bands. The NOR is indicated as a separate block.
</paragraph>
<paragraph id="8BEE36C0FFB3FFF7307EFA4762F6FA34" blockId="11.[533,1510,1388,1500]" box="[533,941,1481,1500]" pageId="11" pageNumber="12">https://doi.org/10.1371/journal.pone.0209792.g005</paragraph>
</caption>
<paragraph id="8BEE36C0FFB3FFF0305EF89D63A7F95C" blockId="11.[533,1536,1534,1870]" lastBlockId="12.[533,1536,1554,1890]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Variation in corolla color and length of the fruiting calyx appendages can be observed in the field in individuals growing under the same environmental conditions. The corolla is mainly pure yellow (
<figureCitation id="136A2A45FFB4FFF03084F99C62F1F9C2" box="[751,938,1554,1578]" captionStart="Fig 4" captionStartId="10.[333,361,203,222]" captionTargetId="figure-1@9.[333,1536,208,1872]" captionTargetPageId="9" captionText="Fig 4. Capsicum longifolium Barboza &amp; S. Leiva. (A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza." pageId="12" pageNumber="13">Fig 4E, 4H and 4I</figureCitation>
), but occasional specimens have corolla lobes red-to brown-edged (
<figureCitation id="136A2A45FFB4FFF030D8F9BA6219F9A4" box="[691,834,1588,1612]" captionStart="Fig 4" captionStartId="10.[333,361,203,222]" captionTargetId="figure-1@9.[333,1536,208,1872]" captionTargetPageId="9" captionText="Fig 4. Capsicum longifolium Barboza &amp; S. Leiva. (A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza." pageId="12" pageNumber="13">Fig 4J and 4K</figureCitation>
), or with a red-brown ring inside the corolla limb (
<figureCitation id="136A2A45FFB4FFF0370AF9BA636EF987" captionStart="Fig 4" captionStartId="10.[333,361,203,222]" captionTargetId="figure-1@9.[333,1536,208,1872]" captionTargetPageId="9" captionText="Fig 4. Capsicum longifolium Barboza &amp; S. Leiva. (A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza." pageId="12" pageNumber="13">Fig 4F and 4G</figureCitation>
); in this latter case, the filaments and the style are also red-brown. In general, the fruiting calyx appendages do not enlarge considerably (
<figureCitation id="136A2A45FFB4FFF03663F9F465C4F97A" box="[1032,1183,1658,1682]" captionStart="Fig 4" captionStartId="10.[333,361,203,222]" captionTargetId="figure-1@9.[333,1536,208,1872]" captionTargetPageId="9" captionText="Fig 4. Capsicum longifolium Barboza &amp; S. Leiva. (A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza." pageId="12" pageNumber="13">Fig 4L and 4N</figureCitation>
) but some specimens have long appendages (
<figureCitation id="136A2A45FFB4FFF030CBF91263B6F95C" box="[672,749,1692,1716]" captionStart="Fig 4" captionStartId="10.[333,361,203,222]" captionTargetId="figure-1@9.[333,1536,208,1872]" captionTargetPageId="9" captionText="Fig 4. Capsicum longifolium Barboza &amp; S. Leiva. (A) Plant. (B) Internode with lenticels. (C, D) Flower buds. (E) Flower, longitudinal section. (F) Flowers showing corolla yellow with brownish center. (G) Same flower as in F, lateral view. (H, I) Flowers with completely yellow corollas, upper and lateral view, respectively. (J, K) Flowers with yellow corollas with red-brown edges, upper and lateral view, respectively. (L, M) Immature fruits. (N) Mature fruit. Photos by S. Leiva González and G. E. Barboza." pageId="12" pageNumber="13">Fig 4M</figureCitation>
).
</paragraph>
<caption id="DF2E6648FFB4FFF032EEFA146346FA07" ID-DOI="http://doi.org/10.5281/zenodo.12630927" ID-Zenodo-Dep="12630927" httpUri="https://zenodo.org/record/12630927/files/figure.png" pageId="12" pageNumber="13" startId="12.[133,161,1434,1453]" targetBox="[134,1535,208,1411]" targetPageId="12" targetType="figure">
<paragraph id="8BEE36C0FFB4FFF032EEFA146316FA2E" blockId="12.[133,1486,1433,1519]" pageId="12" pageNumber="13">
<emphasis id="B925EAD2FFB4FFF032EEFA1463DAFA45" bold="true" box="[133,641,1433,1453]" pageId="12" pageNumber="13">
Fig 6. Bayesian majority-rule consensus tree of
<taxonomicName id="4C514D43FFB4FFF0304DFA176327FA44" box="[550,636,1433,1452]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFB4FFF0304DFA176327FA44" bold="true" box="[550,636,1433,1452]" italics="true" pageId="12" pageNumber="13">Capsicum</emphasis>
</taxonomicName>
.
</emphasis>
Posterior probabilities values indicated by each branch. New species are highlighted in bold-colored letters and the clade to which they belong is indicated.
</paragraph>
<paragraph id="8BEE36C0FFB4FFF032EEFA526346FA07" blockId="12.[133,1486,1433,1519]" box="[133,541,1500,1519]" pageId="12" pageNumber="13">https://doi.org/10.1371/journal.pone.0209792.g006</paragraph>
</caption>
<paragraph id="8BEE36C0FFB4FFF1305EF93163B1FEB8" blockId="12.[533,1536,1554,1890]" lastBlockId="13.[533,1536,208,1896]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
The chromosome number 2n = 26 found in
<taxonomicName id="4C514D43FFB4FFF03662F93165C6F93F" box="[1033,1181,1727,1751]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB4FFF03662F9316541F93E" box="[1033,1050,1727,1750]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="B925EAD2FFB4FFF0364EF93165C6F93F" box="[1061,1181,1727,1751]" italics="true" pageId="12" pageNumber="13">longifolium</emphasis>
</taxonomicName>
is the same as that of
<taxonomicName id="4C514D43FFB4FFF037E9F93163A5F912" authority="(Dunal) Kuntze" authorityName="Kuntze" baseAuthorityName="Dunal" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="rhomboideum">
<emphasis id="B925EAD2FFB4FFF037E9F93164C8F93E" box="[1410,1427,1727,1750]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="B925EAD2FFB4FFF037F5F931630BF911" italics="true" pageId="12" pageNumber="13">rhomboideum</emphasis>
(Dunal) Kuntze
</taxonomicName>
[
<bibRefCitation id="EFC04B31FFB4FFF03166F96C627DF912" author="Scaldaferro MA &amp; Grabiele M &amp; Moscone EA" box="[781,806,1762,1786]" pageId="12" pageNumber="13" pagination="693 - 709" refId="ref14020" refString="32. Scaldaferro MA, Grabiele M, Moscone EA. Heterochromatin type, amount and distribution in wild species of chili peppers (Capsicum - Solanaceae). Genet Resour Crop Ev. 2013; 60: 693 - 709." type="journal article" year="2013">32</bibRefCitation>
],
<taxonomicName id="4C514D43FFB4FFF03150F96C641CF912" authority="(Greenm.) C. V. Morton &amp; Standl." authorityName="C. V. Morton &amp; Standl." baseAuthorityName="Greenm." box="[827,1351,1761,1786]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="lanceolatum">
<emphasis id="B925EAD2FFB4FFF03150F96C6217F911" box="[827,844,1762,1785]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="B925EAD2FFB4FFF03133F96F6282F911" box="[856,985,1761,1785]" italics="true" pageId="12" pageNumber="13">lanceolatum</emphasis>
(Greenm.) C.V. Morton &amp; Standl.
</taxonomicName>
[
<bibRefCitation id="EFC04B31FFB4FFF0373EF96C6435F912" author="Tong N &amp; Bosland PW" box="[1365,1390,1762,1786]" pageId="12" pageNumber="13" pagination="42 - 43" refId="ref14062" refString="33. Tong N, Bosland PW. Meiotic chromosome study of Capsicum lanceolatum. Another 13 chromosome species. Capsicum and Eggplant Newsletter. 1997; 16: 42 - 43." type="journal article" year="1997">33</bibRefCitation>
] and
<taxonomicName id="4C514D43FFB4FFF037C0F96C63BBF8F4" authority="Bitter" authorityName="Bitter" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="lycianthoides">
<emphasis id="B925EAD2FFB4FFF037C0F96C64E7F911" box="[1451,1468,1762,1785]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="B925EAD2FFB4FFF0307EF88A63C5F8F4" box="[533,670,1796,1820]" italics="true" pageId="12" pageNumber="13">lycianthoides</emphasis>
Bitter
</taxonomicName>
[
<bibRefCitation id="EFC04B31FFB4FFF03085F88A625CF8F4" author="Scaldaferro MA &amp; Barboza GE &amp; Moscone EA" box="[750,775,1796,1820]" pageId="12" pageNumber="13" refId="ref14095" refString="34. Scaldaferro MA, Barboza GE, Moscone EA. Estudios citogeneticos en especies andinas silvestres de Capsicum (Solanaceae). J Basic Appl Genet. 2011; 41: 31." type="journal volume" year="2011">34</bibRefCitation>
], all belonging to the Andean clade. Their karyotype formulas are quite similar, but that of
<taxonomicName id="4C514D43FFB4FFF030B4F8A9622FF8D7" box="[735,884,1831,1855]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB4FFF030B4F8A963ABF8D6" box="[735,752,1831,1854]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="B925EAD2FFB4FFF03097F8A9622FF8D7" box="[764,884,1831,1855]" italics="true" pageId="12" pageNumber="13">longifolium</emphasis>
</taxonomicName>
is closest to
<taxonomicName id="4C514D43FFB4FFF0319CF8A965C7F8D7" box="[1015,1180,1831,1855]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="lycianthoides">
<emphasis id="B925EAD2FFB4FFF0319CF8A96553F8D6" box="[1015,1032,1831,1854]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="B925EAD2FFB4FFF03678F8A965C7F8D7" box="[1043,1180,1831,1855]" italics="true" pageId="12" pageNumber="13">lycianthoides</emphasis>
</taxonomicName>
(
<quantity id="4CA99B25FFB4FFF036C1F8A9658FF8D7" box="[1194,1236,1831,1855]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="13" unit="m" value="9.0">9 m</quantity>
+ 3 sm + 1 st) than to
<taxonomicName id="4C514D43FFB4FFF037ABF8A963F2F889" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="rhomboideum">
<emphasis id="B925EAD2FFB4FFF037ABF8A9648AF8D6" box="[1472,1489,1831,1854]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="B925EAD2FFB4FFF0307EF8C763F2F889" box="[533,681,1865,1889]" italics="true" pageId="12" pageNumber="13">rhomboideum</emphasis>
</taxonomicName>
(
<quantity id="4CA99B25FFB4FFF030DCF8C463B6F88A" box="[695,749,1866,1890]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="m" value="10.0">10 m</quantity>
+ 1 sm + 2 st). The species of this clade share small amounts of heterochromatin, a single pair of NOR, short karyotype lengths, and small chromosomes in comparison with other species of the genus [
<bibRefCitation id="EFC04B31FFB5FFF13186FF7C655DFEE2" author="Scaldaferro MA &amp; Grabiele M &amp; Moscone EA" box="[1005,1030,242,266]" pageId="13" pageNumber="14" pagination="693 - 709" refId="ref14020" refString="32. Scaldaferro MA, Grabiele M, Moscone EA. Heterochromatin type, amount and distribution in wild species of chili peppers (Capsicum - Solanaceae). Genet Resour Crop Ev. 2013; 60: 693 - 709." type="journal article" year="2013">32</bibRefCitation>
]. The karyotype of
<taxonomicName id="4C514D43FFB5FFF136BEFF7D6431FEE2" box="[1237,1386,242,266]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="longifolium">
<emphasis id="B925EAD2FFB5FFF136BEFF7D65BDFEE2" box="[1237,1254,243,266]" italics="true" pageId="13" pageNumber="14">C</emphasis>
.
<emphasis id="B925EAD2FFB5FFF13699FF7C6431FEE2" box="[1266,1386,242,266]" italics="true" pageId="13" pageNumber="14">longifolium</emphasis>
</taxonomicName>
is almost half the length of
<taxonomicName id="4C514D43FFB5FFF130CBFE9B620AFEC5" box="[672,849,277,301]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="rhomboideum">
<emphasis id="B925EAD2FFB5FFF130CBFE9B63EAFEC4" box="[672,689,277,300]" italics="true" pageId="13" pageNumber="14">C</emphasis>
.
<emphasis id="B925EAD2FFB5FFF130D6FE9B620AFEC5" box="[701,849,277,301]" italics="true" pageId="13" pageNumber="14">rhomboideum</emphasis>
</taxonomicName>
, the latter with the shortest karyotype length known until now for the entire genus.
</paragraph>
</subSubSection>
</treatment>
</document>