Add updates up until 2024-07-08 18:25:09

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<document id="67C9BB0AC127AA3632967F7977D24653" ID-DOI="10.5281/zenodo.12648807" ID-ISSN="1407-8953" ID-Zenodo-Dep="12648807" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720073718367" checkinUser="felipe" docAuthor="Anichtchenko, Alexander &amp; Medina, Milton Norman" docDate="2019" docId="D9505061500DFFD7FF95FFE32D71C691" docLanguage="en" docName="ActaBiolUnivDaugavp.19.1.13-15.pdf" docOrigin="Acta Biologica Universitatis Daugavpiliensis 19 (1)" docStyle="DocumentStyle:DD7D35B4ACFEF3C20AEB77B8656EEADF.5:ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleId="DD7D35B4ACFEF3C20AEB77B8656EEADF" docStyleName="ActaBiolUnivDaugavp.2009-.journal_article.type1" docStyleVersion="5" docTitle="Neocollyris (Heterocollyris) chaudoiri subsp. sciakyi Anichtchenko &amp; Medina 2019" docType="treatment" docVersion="2" lastPageNumber="14" masterDocId="25692819500CFFD6FFCEFF902C6DC77A" masterDocTitle="A New Neocollyris (Heterocollyris) Subspecies From Mindanao, Philippines (Coleoptera, Carabidae, Cicindelinae)" masterLastPageNumber="15" masterPageNumber="13" pageNumber="14" updateTime="1720086936683" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="76035908C6240B5384E00EA91F8F1F00">A New Neocollyris (Heterocollyris) Subspecies From Mindanao, Philippines (Coleoptera, Carabidae, Cicindelinae)</mods:title>
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<emphasis id="638D3D65500DFFD7FF9AFF032CD6C7D1" bold="true" box="[84,187,147,171]" pageId="1" pageNumber="14">ssp. nov.</emphasis>
(Figs. 1-4)
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del N /
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del N /
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<mods:title id="51626420A8327F8A9F5695CD3EB9C98C">Four new species of the planthopper genus Catonidia Uhler, 1896 from China (Hemiptera: Fulgoromorpha: Achilidae)</mods:title>
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<mods:namePart id="105DCFBBE4C89C7C267A6CBF6A3D5B65">Lv, Sha-Sha</mods:namePart>
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<mods:affiliation id="B7020FF819A87EA9D67B34D8EBF2308D">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China.</mods:affiliation>
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<mods:affiliation id="98E99E4B8EB23FBA83DC7DEE56B69378">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China.</mods:affiliation>
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<mods:title id="25ADC0B55A2E0BE314C1E39957018045">European Journal of Taxonomy</mods:title>
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<mods:part id="0F3BDA4457B46777691080F44F8DA2A3">
<mods:date id="2AB403C2F843CDBAB3CC886140830EE5">2024</mods:date>
<mods:detail id="939F0C60BA5F2A558717A2480E97938E" type="pubDate">
<mods:number id="77DA0549651733FFBE19D67713176B40">2024-07-01</mods:number>
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<mods:number id="C320055C5F8BA292B8E8EDE627CE6025">943</mods:number>
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<mods:start id="585B9F8F080D2648DFEF837239E0CA89">127</mods:start>
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<mods:url id="849D298826F2AE4F5BA5BBF8388BFF08">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2589/11801</mods:url>
</mods:location>
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<mods:identifier id="45485FFD8A9A29731C9789FA354D09AB" type="CLB-Dataset">299624</mods:identifier>
<mods:identifier id="5A312668A9EC137FD76C92CCE9740E25" type="DOI">10.5852/ejt.2024.943.2589</mods:identifier>
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<treatment id="D96C878CFFC1FF89FDF4FEB0B283FC2A" ID-DOI="http://doi.org/10.5281/zenodo.12656371" ID-GBIF-Taxon="235006929" ID-Zenodo-Dep="12656371" LSID="urn:lsid:zoobank.org:act:A75B7B43-404C-48D9-B340-40E15AD5A905" httpUri="http://treatment.plazi.org/id/D96C878CFFC1FF89FDF4FEB0B283FC2A" lastPageId="6" lastPageNumber="133" pageId="4" pageNumber="131">
<subSubSection id="19DF6511FFC1FF8BFDF4FEB0B5DDFEA0" box="[624,964,264,291]" pageId="4" pageNumber="131" type="nomenclature">
<paragraph id="517A369AFFC1FF8BFDF4FEB0B5DDFEA0" blockId="4.[365,1223,264,361]" box="[624,964,264,291]" pageId="4" pageNumber="131">
<heading id="0A3281F6FFC1FF8BFDF4FEB0B5DDFEA0" box="[624,964,264,291]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="131" reason="2">
<taxonomicName id="96C54D19FFC1FF8BFDF4FEB0B57BFEA0" authorityName="Lv &amp; Long &amp; Yang &amp; Chen" authorityYear="2024" box="[624,866,264,291]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="131" phylum="Arthropoda" rank="species" species="triangula" status="sp. nov.">
<emphasis id="63B1EA88FFC1FF8BFDF4FEB0B57BFEA0" bold="true" box="[624,866,264,291]" italics="true" pageId="4" pageNumber="131">Catonidia triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="788257F3FFC1FF8BFCEDFEB1B5DDFEA0" box="[873,964,265,291]" pageId="4" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC1FF8BFEE9FE94B573FEEA" pageId="4" pageNumber="131" type="description">
<paragraph id="517A369AFFC1FF8BFEE9FE94B2DEFEC4" blockId="4.[365,1223,264,361]" box="[365,1223,300,327]" pageId="4" pageNumber="131">
<uri id="25543A98FFC1FF8BFEE9FE94B2DEFEC4" box="[365,1223,300,327]" pageId="4" pageNumber="131">
urn:lsid:zoobank.org:act:
<uuid id="25630C4FFFC1FF8BFD0BFE94B2DEFEC4" box="[655,1223,300,327]" pageId="4" pageNumber="131">A75B7B43-404C-48D9-B340-40E15AD5A905</uuid>
</uri>
</paragraph>
<paragraph id="517A369AFFC1FF8BFD4DFEF7B573FEEA" blockId="4.[365,1223,264,361]" box="[713,874,335,361]" pageId="4" pageNumber="131">
<figureCitation id="C9FE2A1FFFC1FF8BFD4DFEF7B559FEEA" box="[713,832,335,361]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Figs 112</figureCitation>
,
<figureCitation id="C9FE2A1FFFC1FF8BFCC9FEF7B573FEEA" box="[845,874,335,361]" captionStart="Fig" captionStartId="13.[189,232,1998,2024]" captionTargetBox="[192,1388,1015,1936]" captionTargetId="figure-224@13.[189,1398,996,1960]" captionTargetPageId="13" captionText="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi="http://doi.org/10.5281/zenodo.12656834" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="4" pageNumber="131">47</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC1FF8BFF39FE28B466FD0A" pageId="4" pageNumber="131" type="diagnosis">
<paragraph id="517A369AFFC1FF8BFF39FE28B72EFE29" blockId="4.[189,311,400,426]" box="[189,311,400,426]" pageId="4" pageNumber="131">
<heading id="0A3281F6FFC1FF8BFF39FE28B72EFE29" bold="true" box="[189,311,400,426]" fontSize="11" level="3" pageId="4" pageNumber="131" reason="3">
<emphasis id="63B1EA88FFC1FF8BFF39FE28B72EFE29" bold="true" box="[189,311,400,426]" pageId="4" pageNumber="131">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC1FF8BFF39FE07B466FD0A" blockId="4.[187,1399,447,650]" pageId="4" pageNumber="131">
Salient features of new species include: forewing (
<figureCitation id="C9FE2A1FFFC1FF8BFD7BFE07B57CFE5A" box="[767,869,447,473]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Figs 12</figureCitation>
) grayish white, many grayish brown to brown spots distributed along veins, veins grayish white; medioventral process of pygofer (
<figureCitation id="C9FE2A1FFFC1FF8BFB20FE5AB2F3FE7E" box="[1188,1258,482,509]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Fig. 8</figureCitation>
) triangular, lateral margins straight in ventral view; aedeagus (
<figureCitation id="C9FE2A1FFFC1FF8BFC88FDBEB588FDA3" box="[780,913,518,544]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Figs 1011</figureCitation>
) asymmetrical at end, in left lateral view, apical part of phallobasal conjunctival processes nearly angular, lateral side with small ear-like process at subapical part; in right lateral view, apical part of phallobasal conjunctival processes wide, rounded at apex, ear-like process broad and large.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC1FF8BFF39FD09B411FC9F" pageId="4" pageNumber="131" type="etymology">
<paragraph id="517A369AFFC1FF8BFF39FD09B75CFD48" blockId="4.[189,325,689,715]" box="[189,325,689,715]" pageId="4" pageNumber="131">
<heading id="0A3281F6FFC1FF8BFF39FD09B75CFD48" bold="true" box="[189,325,689,715]" fontSize="11" level="3" pageId="4" pageNumber="131" reason="3">
<emphasis id="63B1EA88FFC1FF8BFF39FD09B75CFD48" bold="true" box="[189,325,689,715]" pageId="4" pageNumber="131">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC1FF8BFF39FD67B411FC9F" blockId="4.[188,1399,735,797]" pageId="4" pageNumber="131">
The specific epithet is derived from the Latin word
<emphasis id="63B1EA88FFC1FF8BFC98FD67B58FFD7A" box="[796,918,735,761]" italics="true" pageId="4" pageNumber="131">triangular</emphasis>
, referring to the triangular medioventral process of the male pygofer.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC1FF8BFF39FCFCB498FB22" pageId="4" pageNumber="131" type="materials_examined">
<paragraph id="517A369AFFC1FF8BFF39FCFCB774FCDD" blockId="4.[189,365,836,862]" box="[189,365,836,862]" pageId="4" pageNumber="131">
<heading id="0A3281F6FFC1FF8BFF39FCFCB774FCDD" bold="true" box="[189,365,836,862]" fontSize="11" level="3" pageId="4" pageNumber="131" reason="3">
<emphasis id="63B1EA88FFC1FF8BFF39FCFCB774FCDD" bold="true" box="[189,365,836,862]" pageId="4" pageNumber="131">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC1FF8BFF6EFCCBB744FC0E" blockId="4.[189,1399,883,979]" box="[234,349,883,909]" pageId="4" pageNumber="131">
<emphasis id="63B1EA88FFC1FF8BFF6EFCCBB744FC0E" bold="true" box="[234,349,883,909]" pageId="4" pageNumber="131">
<typeStatus id="8E7E8838FFC1FF8BFF6EFCCBB744FC0E" box="[234,349,883,909]" pageId="4" pageNumber="131" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="517A369AFFC1FF8BFF39FC2EB41BFC50" blockId="4.[189,1399,883,979]" pageId="4" pageNumber="131">
<materialsCitation id="E1AD3CC7FFC1FF8BFF39FC2EB41BFC50" ID-GBIF-Occurrence="4903684306" collectingDate="2013-08-16" collectionCode="IEGU" collectorName="Yuan Liu" country="China" latitude="27.688" location="Wuyishan National Natural Reserve" longitude="117.742" pageId="4" pageNumber="131" specimenCount="1" specimenCount-male="1" stateProvince="Fujian Province" typeStatus="holotype">
<collectingCountry id="29D2760AFFC1FF8BFF39FC2EB701FC33" box="[189,280,918,944]" name="China" pageId="4" pageNumber="131">CHINA</collectingCountry>
<emphasis id="63B1EA88FFC1FF8BFEBFFC2DB41EFC2C" bold="true" box="[315,519,917,943]" pageId="4" pageNumber="131">
<collectingRegion id="9301F878FFC1FF8BFEBFFC2DB41EFC2C" box="[315,519,917,943]" country="China" name="Fujian" pageId="4" pageNumber="131">Fujian Province</collectingRegion>
</emphasis>
<specimenCount id="47C3FD13FFC1FF8BFDA2FC2EB426FC33" box="[550,575,918,944]" count="1" pageId="4" pageNumber="131" type="male"></specimenCount>
;
<location id="541A6041FFC1FF8BFDCAFC2EB218FC33" LSID="urn:lsid:plazi:treatment:D96C878CFFC1FF89FDF4FEB0B283FC2A:541A6041FFC1FF8BFDCAFC2EB218FC33" box="[590,1025,918,944]" country="China" name="Wuyishan National Natural Reserve" pageId="4" pageNumber="131" stateProvince="Fujian Province">Wuyishan National Natural Reserve</location>
;
<geoCoordinate id="34F1505DFFC1FF8BFB94FC2EB29DFC33" box="[1040,1156,916,944]" degrees="27" direction="north" minutes="68" orientation="latitude" pageId="4" pageNumber="131" precision="925" value="27.688">27°67 N,</geoCoordinate>
<geoCoordinate id="34F1505DFFC1FF8BFB0AFC2EB31EFC33" box="[1166,1287,916,944]" degrees="27" direction="north" minutes="68" orientation="latitude" pageId="4" pageNumber="131" precision="925" value="27.688">117°96 E</geoCoordinate>
;
<date id="257B105AFFC1FF8BFA93FC2EB6E2FC50" pageId="4" pageNumber="131" value="2013-08-16">
<collectingDate id="353FE9B2FFC1FF8BFA93FC2EB6E2FC50" pageId="4" pageNumber="131" value="2013-08-16">16 Aug. 2013</collectingDate>
</date>
;
<collectorName id="FC30534CFFC1FF8BFE82FC01B76AFC50" box="[262,371,953,979]" pageId="4" pageNumber="131">Yuan Liu</collectorName>
leg.;
<collectionCode id="37D4AE5FFFC1FF8BFE30FC01B7E7FC50" box="[436,510,953,979]" pageId="4" pageNumber="131">IEGU</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="517A369AFFC1FF8BFF6EFC42B770FB97" blockId="4.[189,1398,1018,1186]" box="[234,361,1018,1044]" pageId="4" pageNumber="131">
<emphasis id="63B1EA88FFC1FF8BFF6EFC42B770FB97" bold="true" box="[234,361,1018,1044]" pageId="4" pageNumber="131">
<typeStatus id="8E7E8838FFC1FF8BFF6EFC42B770FB97" box="[234,361,1018,1044]" pageId="4" pageNumber="131" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="517A369AFFC1FF8BFF39FBA6B498FB22" blockId="4.[189,1398,1018,1186]" pageId="4" pageNumber="131">
<materialsCitation id="E1AD3CC7FFC1FF8BFF39FBA6B36FFBBB" ID-GBIF-Occurrence="4903684305" box="[189,1398,1053,1080]" collectingDate="2009-08-05" collectionCode="IEGU" collectorName="Yan-Li Zheng" country="China" latitude="27.688" location="Fujian Province" longitude="117.742" pageId="4" pageNumber="131" specimenCount="1" specimenCount-male="1" stateProvince="Fujian Province" typeStatus="paratype">
<collectingCountry id="29D2760AFFC1FF8BFF39FBA6B701FBBB" box="[189,280,1054,1080]" name="China" pageId="4" pageNumber="131">CHINA</collectingCountry>
<emphasis id="63B1EA88FFC1FF8BFEBFFBA5B41CFBB4" bold="true" box="[315,517,1053,1079]" pageId="4" pageNumber="131">
<collectingRegion id="9301F878FFC1FF8BFEBFFBA5B41CFBB4" box="[315,517,1053,1079]" country="China" name="Fujian" pageId="4" pageNumber="131">Fujian Province</collectingRegion>
</emphasis>
<specimenCount id="47C3FD13FFC1FF8BFD9BFBA6B449FBBB" box="[543,592,1054,1080]" count="1" pageId="4" pageNumber="131" type="male">1 ♂</specimenCount>
; same data as for holotype;
<date id="257B105AFFC1FF8BFC26FBA5B220FBBB" box="[930,1081,1053,1080]" pageId="4" pageNumber="131" value="2009-08-05">
<collectingDate id="353FE9B2FFC1FF8BFC26FBA5B220FBBB" box="[930,1081,1053,1080]" pageId="4" pageNumber="131" value="2009-08-05">5 Aug. 2009</collectingDate>
</date>
;
<collectorName id="FC30534CFFC1FF8BFBC1FBA5B2F2FBB4" box="[1093,1259,1053,1079]" pageId="4" pageNumber="131">Yan-Li Zheng</collectorName>
leg.;
<collectionCode id="37D4AE5FFFC1FF8BFAB4FBA6B36FFBBB" box="[1328,1398,1054,1080]" pageId="4" pageNumber="131">IEGU</collectionCode>
</materialsCitation>
<materialsCitation id="E1AD3CC7FFC1FF8BFF55FBF9B71EFBFD" ID-GBIF-Occurrence="4903684313" collectingDate="2013-08-16" collectingDateMax="2013-08-19" collectingDateMin="2013-08-16" collectionCode="IEGU" collectorName="Ying-Jian Wang &amp; Yang-Yang Liu" country="China" location="Wuyishan National Natural Reserve" pageId="4" pageNumber="131" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Fujian" typeStatus="paratype">
<specimenCount id="47C3FD13FFC1FF8BFF55FBF9B70EFBD8" box="[209,279,1089,1115]" count="4" pageId="4" pageNumber="131" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="47C3FD13FFC1FF8BFEA1FBF9B74DFBD8" box="[293,340,1089,1115]" count="1" pageId="4" pageNumber="131" type="female">1 ♀</specimenCount>
; same data as for holotype;
<date id="257B105AFFC1FF8BFD2FFBF9B564FBD8" box="[683,893,1089,1115]" pageId="4" pageNumber="131" value="2013-08-16" valueMax="2013-08-19" valueMin="2013-08-16">
<collectingDate id="353FE9B2FFC1FF8BFD2FFBF9B564FBD8" box="[683,893,1089,1115]" pageId="4" pageNumber="131" value="2013-08-16" valueMax="2013-08-19" valueMin="2013-08-16">1619 Aug. 2013</collectingDate>
</date>
;
<collectorName id="FC30534CFFC1FF8BFC0EFBF9B250FBD8" box="[906,1097,1089,1115]" pageId="4" pageNumber="131">Ying-Jian Wang</collectorName>
and
<collectorName id="FC30534CFFC1FF8BFB01FBF9B323FBD8" box="[1157,1338,1089,1115]" pageId="4" pageNumber="131">Yang-Yang Liu</collectorName>
leg.;
<collectionCode id="37D4AE5FFFC1FF8BFF39FBDCB71EFBFD" box="[189,263,1124,1150]" pageId="4" pageNumber="131">IEGU</collectionCode>
</materialsCitation>
.
<materialsCitation id="E1AD3CC7FFC1FF8BFEABFBDCB464FB22" ID-GBIF-Occurrence="4903684312" collectingDate="2010-08-16" collectionCode="IEGU" collectorName="Xiao-Hui Hou" country="China" latitude="27.688" location="Kuankuoshui National Natural Reserve" longitude="107.19" pageId="4" pageNumber="131" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou Province" typeStatus="paratype">
<emphasis id="63B1EA88FFC1FF8BFEABFBDCB40BFBFD" bold="true" box="[303,530,1124,1150]" pageId="4" pageNumber="131">
<collectingRegion id="9301F878FFC1FF8BFEABFBDCB40BFBFD" box="[303,530,1124,1150]" country="China" name="Guizhou" pageId="4" pageNumber="131">Guizhou Province</collectingRegion>
</emphasis>
<specimenCount id="47C3FD13FFC1FF8BFDA9FBDCB446FBFD" box="[557,607,1124,1150]" count="1" pageId="4" pageNumber="131" type="male">1 ♂</specimenCount>
;
<location id="541A6041FFC1FF8BFDE9FBDCB25FFBFD" LSID="urn:lsid:plazi:treatment:D96C878CFFC1FF89FDF4FEB0B283FC2A:541A6041FFC1FF8BFDE9FBDCB25FFBFD" box="[621,1094,1124,1150]" country="China" longitude="107.316666" name="Kuankuoshui National Natural Reserve" pageId="4" pageNumber="131" stateProvince="Guizhou Province">Kuankuoshui National Natural Reserve</location>
;
<geoCoordinate id="34F1505DFFC1FF8BFBD0FBDCB2DEFBFD" box="[1108,1223,1122,1150]" degrees="107" direction="east" minutes="19" orientation="longitude" pageId="4" pageNumber="131" precision="925" value="107.19">27°96 N,</geoCoordinate>
<geoCoordinate id="34F1505DFFC1FF8BFB54FBDCB353FBFD" box="[1232,1354,1122,1150]" degrees="107" direction="east" minutes="19" orientation="longitude" pageId="4" pageNumber="131" precision="925" value="107.19">107°19 E</geoCoordinate>
;
<date id="257B105AFFC1FF8BFADDFBDCB725FB22" pageId="4" pageNumber="131" value="2010-08-16">
<collectingDate id="353FE9B2FFC1FF8BFADDFBDCB725FB22" pageId="4" pageNumber="131" value="2010-08-16">16 Aug. 2010</collectingDate>
</date>
;
<collectorName id="FC30534CFFC1FF8BFECCFB3FB7E8FB21" box="[328,497,1159,1186]" pageId="4" pageNumber="131">Xiao-Hui Hou</collectorName>
leg.;
<collectionCode id="37D4AE5FFFC1FF8BFDB7FB3FB464FB22" box="[563,637,1159,1185]" pageId="4" pageNumber="131">IEGU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC1FF89FF39FB70B4B6FDE3" lastPageId="6" lastPageNumber="133" pageId="4" pageNumber="131" type="description">
<paragraph id="517A369AFFC1FF8BFF39FB70B756FB61" blockId="4.[189,335,1224,1250]" box="[189,335,1224,1250]" pageId="4" pageNumber="131">
<heading id="0A3281F6FFC1FF8BFF39FB70B756FB61" bold="true" box="[189,335,1224,1250]" fontSize="11" level="3" pageId="4" pageNumber="131" reason="3">
<emphasis id="63B1EA88FFC1FF8BFF39FB70B756FB61" bold="true" box="[189,335,1224,1250]" pageId="4" pageNumber="131">Description</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC1FF8BFF39FB40B34DFA92" blockId="4.[189,1364,1271,1298]" box="[189,1364,1271,1298]" pageId="4" pageNumber="131">
<smallCapsWord id="579CA046FFC1FF8BFF39FB40B777FA93" baselines="1291,1291" box="[189,366,1272,1298]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="4" pageNumber="131">MEASUREMENTS</smallCapsWord>
. Body length including forewing: male
<quantity id="963D9B7FFFC1FF8BFCC1FB4FB5CBFA91" box="[837,978,1271,1298]" metricMagnitude="-3" metricUnit="m" metricValue="8.85" metricValueMax="9.2" metricValueMin="8.5" pageId="4" pageNumber="131" unit="mm" value="8.85" valueMax="9.2" valueMin="8.5">8.59.2 mm</quantity>
(N = 7), female
<quantity id="963D9B7FFFC1FF8BFB11FB4FB2F6FA91" box="[1173,1263,1271,1298]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="4" pageNumber="131" unit="mm" value="9.8">9.8 mm</quantity>
(N = 1).
</paragraph>
<paragraph id="517A369AFFC1FF8BFF39FA81B5E9FA5C" blockId="4.[187,1399,1336,1504]" pageId="4" pageNumber="131">
<smallCapsWord id="579CA046FFC1FF8BFF39FA81B751FAD2" baselines="1356,1356" box="[189,328,1337,1363]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="4" pageNumber="131">COLORATION</smallCapsWord>
. General color grayish white to yellowish brown (
<figureCitation id="C9FE2A1FFFC1FF8BFC18FA80B21EFAD0" box="[924,1031,1336,1363]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Figs 12</figureCitation>
). Vertex pale yellowish brown. Eyes reddish-brown, ocelli light reddish-brown. Genae pale brown. Pronotum and tegula yellowish white to yellowish brown. Mesonotum yellowish brown to fuscous. Forewings grayish white, semihyaline, many grayish brown to brown spots distributed along veins, veins grayish white. Hindwings nearly hyaline, gray, veins light brown. Legs and abdomen yellowish brown.
</paragraph>
<paragraph id="517A369AFFC1FF8BFF39F9BFB436F802" blockId="4.[189,1400,1543,1921]" pageId="4" pageNumber="131">
<smallCapsWord id="579CA046FFC1FF8BFF39F9BFB6E5F99C" baselines="1562,1562" box="[189,252,1543,1569]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="4" pageNumber="131">HEAD</smallCapsWord>
<smallCapsWord id="579CA046FFC1FF8BFE81F9B4B72BF99C" baselines="1562" box="[261,306,1548,1567]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="4" pageNumber="131">AND</smallCapsWord>
<smallCapsWord id="579CA046FFC1FF8BFEBDF9BFB78CF99C" baselines="1562,1562" box="[313,405,1543,1569]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="4" pageNumber="131">THORAX</smallCapsWord>
. Head (
<figureCitation id="C9FE2A1FFFC1FF8BFE75F9BFB440F9A2" box="[497,601,1543,1569]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Figs 1, 3</figureCitation>
) including eyes narrower than pronotum (0.6: 1). Vertex wider than long in middle line (2.6: 1), apex concave, median carina absent, anterior margin broadly rounded, posterior margin rounded concavely, lateral margins ridged. Frons (
<figureCitation id="C9FE2A1FFFC1FF8BFC60F9F5B234F9E4" box="[996,1069,1613,1639]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Fig. 4</figureCitation>
) longer in middle line than maximum width (1.9: 1), width at base narrower than width at apex (0.8: 1), median carina distinct, lateral margins concave. Postclypeus (
<figureCitation id="C9FE2A1FFFC1FF8BFDF8F92BB4DAF92D" box="[636,707,1683,1710]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Fig. 4</figureCitation>
) shorter than frons in midline (about 0.5: 1), median carina distinct, lateral margins excavate. Pronotum (
<figureCitation id="C9FE2A1FFFC1FF8BFD53F90FB55BF952" box="[727,834,1719,1745]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Figs 1, 3</figureCitation>
) longer in midline than vertex (1.8: 1), median carina distinct, lateral carinae raised, not extending to posterior margin, confluent with midline at anterior margin to form angular process. Mesonotum (
<figureCitation id="C9FE2A1FFFC1FF8BFD61F945B548F894" box="[741,849,1789,1815]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Figs 1, 3</figureCitation>
) about 4.2 × as long as pronotum in midline, longer than pronotum and vertex combined (2.7: 1), apical part of median carina clear, lateral carinae unclear. Forewings (
<figureCitation id="C9FE2A1FFFC1FF8BFE2AF8FBB7EAF8DD" box="[430,499,1859,1886]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Fig. 5</figureCitation>
) broadened apically, longer than maximal width (2.2: 1). Hindwings (
<figureCitation id="C9FE2A1FFFC1FF8BFAADF8FBB376F8DE" box="[1321,1391,1859,1885]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Fig. 6</figureCitation>
) longer than widest part (1.4: 1).
</paragraph>
<paragraph id="517A369AFFC1FF89FF39F811B4B6FDE3" blockId="4.[189,1401,1960,2022]" lastBlockId="6.[188,1399,265,608]" lastPageId="6" lastPageNumber="133" pageId="4" pageNumber="131">
<smallCapsWord id="579CA046FFC1FF8BFF39F811B6E6F842" baselines="1980,1980" box="[189,255,1961,1987]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="4" pageNumber="131">MALE</smallCapsWord>
<smallCapsWord id="579CA046FFC1FF8BFE8DF816B76EF842" baselines="1980" box="[265,375,1966,1985]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="4" pageNumber="131">GENITALIA</smallCapsWord>
. Pygofer (
<figureCitation id="C9FE2A1FFFC1FF8BFE7CF810B47BF841" box="[504,610,1960,1986]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="4" pageNumber="131">Figs 79</figureCitation>
) in lateral view with dorsal margin distinctly shorter than ventral margin, ⅓ of posterior margin strongly angulated caudad; in ventral view, medioventral process triangular, lateral margins straight. Anal tube (
<figureCitation id="C9FE2A1FFFC3FF89FD75FEB1B579FEA0" box="[753,864,265,291]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="6" pageNumber="133">Figs 7, 9</figureCitation>
) longer than widest part by about 1.4:
<quantity id="963D9B7FFFC3FF89FAC2FEB1B36EFEA0" box="[1350,1399,265,291]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="133" unit="in" value="1.0">1 in</quantity>
dorsal view, apical margin rounded and convex, basal margin slightly truncate. Aedeagus (
<figureCitation id="C9FE2A1FFFC3FF89FB6CFE94B377FEC5" box="[1256,1390,300,326]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="6" pageNumber="133">Figs 1011</figureCitation>
) asymmetrical at end, in left lateral view, phallobase broad at base, dorsal lobe stout finger-like, apical margin rounded; lateral lobes pointed apically, proximal part of ventral margin slightly rolled up; basal part of ventral lobe constricted into a stalk, bends and extends ventrad, middle part narrowing apically, constricted abruptly at apex; apical part of phallobasal conjunctival processes nearly angular, lateral side with small ear-like process at subapical part; in right lateral view, apical part of phallobasal conjunctival processes wide, rounded, ear-like process broad and large. Gonostyli (
<figureCitation id="C9FE2A1FFFC3FF89FC6FFE47B279FD9A" box="[1003,1120,511,538]" captionStart="Figs 112" captionStartId="5.[189,242,1891,1917]" captionTargetBox="[211,1388,376,1833]" captionTargetId="figure-11@5.[189,1398,367,1853]" captionTargetPageId="5" captionText="Figs 112. Catonidia triangula sp. nov., ♂, holotype (IEGU). 1. Habitus, dorsal view. 2. Habitus, lateral view. 3. Head and thorax, dorsal view.4. Frons, ventral view.5. Forewing.6. Hindwing.7. Pygofer, lateral view. 8. Pygofer, ventral view. 9. Pygofer, dorsal view. 10. Aedeagus, left lateral view. 11. Aedeagus, right lateral view. 12. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="6" pageNumber="133">Figs 7, 12</figureCitation>
) longer than wide, base narrow, apical margin broadly rounded, dorsal margin with stout twisted auricular process near middle, inner side of base with finger-like process.
</paragraph>
</subSubSection>
<caption id="05BA6612FFC0FF8AFF39F8DBB20BF864" ID-DOI="http://doi.org/10.5281/zenodo.12656824" ID-Zenodo-Dep="12656824" httpUri="https://zenodo.org/record/12656824/files/figure.png" pageId="5" pageNumber="132" startId="5.[189,242,1891,1917]" targetBox="[211,1388,376,1833]" targetPageId="5">
<paragraph id="517A369AFFC0FF8AFF39F8DBB20BF864" blockId="5.[186,1398,1891,2024]" pageId="5" pageNumber="132">
<emphasis id="63B1EA88FFC0FF8AFF39F8DBB722F8FE" bold="true" box="[189,315,1891,1917]" pageId="5" pageNumber="132">Figs 112.</emphasis>
<taxonomicName id="96C54D19FFC0FF8AFEC4F8DBB432F8FE" authorityName="Lv &amp; Long &amp; Yang &amp; Chen" authorityYear="2024" box="[320,555,1891,1917]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="species" species="triangula" status="sp. nov.">
<emphasis id="63B1EA88FFC0FF8AFEC4F8DBB432F8FE" box="[320,555,1891,1917]" italics="true" pageId="5" pageNumber="132">Catonidia triangula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="788257F3FFC0FF8AFDB7F8DCB496F8FD" box="[563,655,1892,1918]" pageId="5" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (IEGU).
<emphasis id="63B1EA88FFC0FF8AFC15F8DBB5B9F8FE" bold="true" box="[913,928,1891,1917]" pageId="5" pageNumber="132">1</emphasis>
. Habitus, dorsal view.
<emphasis id="63B1EA88FFC0FF8AFB23F8DBB2AFF8FE" bold="true" box="[1191,1206,1891,1917]" pageId="5" pageNumber="132">2</emphasis>
. Habitus, lateral view.
<emphasis id="63B1EA88FFC0FF8AFE84F83EB716F823" bold="true" box="[256,271,1926,1952]" pageId="5" pageNumber="132">3</emphasis>
. Head and thorax, dorsal view.
<emphasis id="63B1EA88FFC0FF8AFDF5F83EB499F823" bold="true" box="[625,640,1926,1952]" pageId="5" pageNumber="132">4</emphasis>
. Frons, ventral view.
<emphasis id="63B1EA88FFC0FF8AFCF0F83EB59AF823" bold="true" box="[884,899,1926,1952]" pageId="5" pageNumber="132">5</emphasis>
. Forewing.
<emphasis id="63B1EA88FFC0FF8AFB88F83EB202F823" bold="true" box="[1036,1051,1926,1952]" pageId="5" pageNumber="132">6</emphasis>
. Hindwing.
<emphasis id="63B1EA88FFC0FF8AFB2CF83EB2AEF823" bold="true" box="[1192,1207,1926,1952]" pageId="5" pageNumber="132">7</emphasis>
. Pygofer, lateral view.
<emphasis id="63B1EA88FFC0FF8AFE81F811B70DF840" bold="true" box="[261,276,1961,1987]" pageId="5" pageNumber="132">8</emphasis>
. Pygofer, ventral view.
<emphasis id="63B1EA88FFC0FF8AFDB0F811B458F840" bold="true" box="[564,577,1961,1987]" pageId="5" pageNumber="132">9</emphasis>
. Pygofer, dorsal view.
<emphasis id="63B1EA88FFC0FF8AFCDCF811B56DF840" bold="true" box="[856,884,1961,1987]" pageId="5" pageNumber="132">10</emphasis>
. Aedeagus, left lateral view.
<emphasis id="63B1EA88FFC0FF8AFB56F811B2F5F840" bold="true" box="[1234,1260,1961,1987]" pageId="5" pageNumber="132">11</emphasis>
. Aedeagus, right lateral view.
<emphasis id="63B1EA88FFC0FF8AFE16F875B7B6F864" bold="true" box="[402,431,1997,2023]" pageId="5" pageNumber="132">12</emphasis>
. Left gonostyli, inner surface. Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="19DF6511FFC3FF89FF39FD3FB4D2FD53" box="[189,715,647,720]" pageId="6" pageNumber="133" type="distribution">
<paragraph id="517A369AFFC3FF89FF39FD3FB741FD22" blockId="6.[189,344,647,673]" box="[189,344,647,673]" pageId="6" pageNumber="133">
<heading id="0A3281F6FFC3FF89FF39FD3FB741FD22" bold="true" box="[189,344,647,673]" fontSize="11" level="3" pageId="6" pageNumber="133" reason="3">
<emphasis id="63B1EA88FFC3FF89FF39FD3FB741FD22" bold="true" box="[189,344,647,673]" pageId="6" pageNumber="133">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC3FF89FF39FD0EB4D2FD53" blockId="6.[189,715,694,720]" box="[189,715,694,720]" pageId="6" pageNumber="133">
<collectingCountry id="29D2760AFFC3FF89FF39FD0EB71AFD53" box="[189,259,694,720]" name="China" pageId="6" pageNumber="133">China</collectingCountry>
(
<collectingRegion id="9301F878FFC3FF89FE96FD0EB77BFD53" box="[274,354,694,720]" country="China" name="Fujian" pageId="6" pageNumber="133">Fujian</collectingRegion>
,
<collectingRegion id="9301F878FFC3FF89FEEAFD0EB7CAFD53" box="[366,467,694,720]" country="China" name="Guizhou" pageId="6" pageNumber="133">Guizhou</collectingRegion>
provinces) (
<figureCitation id="C9FE2A1FFFC3FF89FDE2FD0EB4A7FD53" box="[614,702,694,720]" captionStart="Fig" captionStartId="13.[189,232,1998,2024]" captionTargetBox="[192,1388,1015,1936]" captionTargetId="figure-224@13.[189,1398,996,1960]" captionTargetPageId="13" captionText="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi="http://doi.org/10.5281/zenodo.12656834" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="6" pageNumber="133">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC3FF89FF39FD4FB283FC2A" pageId="6" pageNumber="133" type="discussion">
<paragraph id="517A369AFFC3FF89FF39FD4FB729FC92" blockId="6.[189,304,759,785]" box="[189,304,759,785]" pageId="6" pageNumber="133">
<heading id="0A3281F6FFC3FF89FF39FD4FB729FC92" bold="true" box="[189,304,759,785]" fontSize="11" level="3" pageId="6" pageNumber="133" reason="3">
<emphasis id="63B1EA88FFC3FF89FF39FD4FB729FC92" bold="true" box="[189,304,759,785]" pageId="6" pageNumber="133">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC3FF89FF39FC9EB283FC2A" blockId="6.[189,1399,806,938]" pageId="6" pageNumber="133">
This species is similar to
<taxonomicName id="96C54D19FFC3FF89FE76FC9EB5B6FCC3" authority="Long &amp; Chen, 2012" authorityName="Long &amp; Chen" authorityYear="2012" box="[498,943,806,832]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="133" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="63B1EA88FFC3FF89FE76FC9EB4ADFCC3" box="[498,692,806,832]" italics="true" pageId="6" pageNumber="133">Catonidia choui</emphasis>
<bibRefCitation id="35544B6BFFC3FF89FD3FFC9EB5B6FCC3" author="Long J. K. &amp; Chen X. S." box="[699,943,806,832]" pageId="6" pageNumber="133" pagination="215 - 221" refId="ref7925" refString="Long J. K. &amp; Chen X. S. 2012. A new species of Catonidia Uhler (Hemiptera: Fulgoromorpha: Achilidae) from China. Entomotaxonomia 34 (2): 215 - 221." type="journal article" year="2012">Long &amp; Chen, 2012</bibRefCitation>
</taxonomicName>
, but differs from the latter as follows: (1) forewing grayish white (forewing yellowish brown in
<taxonomicName id="96C54D19FFC3FF89FCE4FCF1B5DDFCE0" authorityName="Long &amp; Chen" authorityYear="2012" box="[864,964,841,867]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="133" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="63B1EA88FFC3FF89FCE4FCF1B5DDFCE0" box="[864,964,841,867]" italics="true" pageId="6" pageNumber="133">C. choui</emphasis>
</taxonomicName>
); (2) medioventral process in ventral view triangular, lateral margins straight (medioventral process in ventral view rounded, lateral margins curved in
<taxonomicName id="96C54D19FFC3FF89FEB6FC28B78EFC2A" authorityName="Long &amp; Chen" authorityYear="2012" box="[306,407,911,938]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="133" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="63B1EA88FFC3FF89FEB6FC28B78EFC2A" box="[306,407,911,938]" italics="true" pageId="6" pageNumber="133">C. choui</emphasis>
</taxonomicName>
); (3) aedeagus asymmetrical (aedeagus symmetrical in
<taxonomicName id="96C54D19FFC3FF89FBA1FC28B293FC2A" authorityName="Long &amp; Chen" authorityYear="2012" box="[1061,1162,911,938]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="133" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="63B1EA88FFC3FF89FBA1FC28B293FC2A" box="[1061,1162,911,938]" italics="true" pageId="6" pageNumber="133">C. choui</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,314 @@
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<mods:title id="96334DC72131C754B45D2F435CE93050">Four new species of the planthopper genus Catonidia Uhler, 1896 from China (Hemiptera: Fulgoromorpha: Achilidae)</mods:title>
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<taxonomicName id="96C54D19FFC3FF89FDF8FC5FB54CFB81" authorityName="Lv &amp; Long &amp; Yang &amp; Chen" authorityYear="2024" box="[636,853,999,1026]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="133" phylum="Arthropoda" rank="species" species="saccata" status="sp. nov.">
<emphasis id="63B1EA88FFC3FF89FDF8FC5FB54CFB81" bold="true" box="[636,853,999,1026]" italics="true" pageId="6" pageNumber="133">Catonidia saccata</emphasis>
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<paragraph id="517A369AFFC3FF89FD46FB96B568FBCB" blockId="6.[365,1223,999,1096]" box="[706,881,1070,1096]" pageId="6" pageNumber="133">
<figureCitation id="C9FE2A1FFFC3FF89FD46FB96B551FBCB" box="[706,840,1070,1096]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="6" pageNumber="133">Figs 1323</figureCitation>
,
<figureCitation id="C9FE2A1FFFC3FF89FCD0FB96B568FBCB" box="[852,881,1070,1096]" captionStart="Fig" captionStartId="13.[189,232,1998,2024]" captionTargetBox="[192,1388,1015,1936]" captionTargetId="figure-224@13.[189,1398,996,1960]" captionTargetPageId="13" captionText="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi="http://doi.org/10.5281/zenodo.12656834" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="6" pageNumber="133">47</figureCitation>
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<emphasis id="63B1EA88FFC3FF89FF39FBD7B72EFB0A" bold="true" box="[189,311,1135,1161]" pageId="6" pageNumber="133">Diagnosis</emphasis>
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Salient features of new species include: forewings (
<figureCitation id="C9FE2A1FFFC3FF89FC93FB26B5DEFB3B" box="[791,967,1182,1208]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="6" pageNumber="133">Figs 1314, 17</figureCitation>
) milk-white, clavus apex ½ to costal margin with broad brown marking, medial aera with 5 small round brown spots; lateral margins of medioventral process (
<figureCitation id="C9FE2A1FFFC3FF89FE43FB5CB454FB7D" box="[455,589,1252,1278]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="6" pageNumber="133">Figs 1920</figureCitation>
) nearly parallel in lateral view, apex with dentate process outward in ventral view; apical part of ventral lobe (
<figureCitation id="C9FE2A1FFFC3FF89FD21FABFB4E4FAA1" box="[677,765,1287,1314]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="6" pageNumber="133">Fig. 22</figureCitation>
) sac-like, lateral sides with 2 lamellar processes near middle; phallobasal conjunctival processes (
<figureCitation id="C9FE2A1FFFC3FF89FD41FA93B502FAC6" box="[709,795,1323,1349]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="6" pageNumber="133">Fig. 22</figureCitation>
) bent nearly 90 degrees toward back at apical part.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC3FF89FF39FAD4B439FA5B" pageId="6" pageNumber="133" type="etymology">
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<emphasis id="63B1EA88FFC3FF89FF39FAD4B75CFA05" bold="true" box="[189,325,1388,1414]" pageId="6" pageNumber="133">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC3FF89FF39FA22B439FA5B" blockId="6.[189,1399,1434,1496]" pageId="6" pageNumber="133">
The specific epithet is derived from the Latin word
<emphasis id="63B1EA88FFC3FF89FCC7FA23B5B0FA37" box="[835,937,1435,1460]" italics="true" pageId="6" pageNumber="133">saccatus</emphasis>
, referring to the sac-like apex of the ventral lobe of the phallobase.
</paragraph>
</subSubSection>
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<paragraph id="517A369AFFC3FF89FF39FA47B774F99A" blockId="6.[189,365,1535,1561]" box="[189,365,1535,1561]" pageId="6" pageNumber="133">
<heading id="0A3281F6FFC3FF89FF39FA47B774F99A" bold="true" box="[189,365,1535,1561]" fontSize="11" level="3" pageId="6" pageNumber="133" reason="3">
<emphasis id="63B1EA88FFC3FF89FF39FA47B774F99A" bold="true" box="[189,365,1535,1561]" pageId="6" pageNumber="133">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC3FF89FF6EF996B744F9CB" blockId="6.[189,1399,1582,1679]" box="[234,349,1582,1608]" pageId="6" pageNumber="133">
<emphasis id="63B1EA88FFC3FF89FF6EF996B744F9CB" bold="true" box="[234,349,1582,1608]" pageId="6" pageNumber="133">
<typeStatus id="8E7E8838FFC3FF89FF6EF996B744F9CB" box="[234,349,1582,1608]" pageId="6" pageNumber="133" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="517A369AFFC3FF89FF39F9E9B40CF90D" blockId="6.[189,1399,1582,1679]" pageId="6" pageNumber="133">
<materialsCitation id="E1AD3CC7FFC3FF89FF39F9E9B40CF90D" ID-GBIF-Occurrence="4903684310" collectingDate="2013-08-11" collectionCode="IEGU" collectorName="Wei-Cheng Yang" country="China" county="Tengchong County" latitude="25.05" location="Jietou Town" longLatPrecision="1248" longitude="98.833336" pageId="6" pageNumber="133" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
<collectingCountry id="29D2760AFFC3FF89FF39F9E9B701F9E8" box="[189,280,1617,1643]" name="China" pageId="6" pageNumber="133">CHINA</collectingCountry>
<emphasis id="63B1EA88FFC3FF89FEBCF9E9B409F9E8" bold="true" box="[312,528,1617,1643]" pageId="6" pageNumber="133">
<collectingRegion id="9301F878FFC3FF89FEBCF9E9B409F9E8" box="[312,528,1617,1643]" country="China" name="Yunnan" pageId="6" pageNumber="133">Yunnan Province</collectingRegion>
</emphasis>
<specimenCount id="47C3FD13FFC3FF89FDAEF9E9B45AF9E8" box="[554,579,1617,1643]" count="1" pageId="6" pageNumber="133" type="male"></specimenCount>
;
<collectingCounty id="B81B4E16FFC3FF89FDD4F9E9B52BF9E8" box="[592,818,1617,1643]" pageId="6" pageNumber="133">Tengchong County</collectingCounty>
,
<location id="541A6041FFC3FF89FCBAF9E9B5CBF9E8" LSID="urn:lsid:plazi:treatment:D96C878CFFC3FF87FDF8FC5FB525F94A:541A6041FFC3FF89FCBAF9E9B5CBF9E8" box="[830,978,1617,1643]" country="China" county="Tengchong County" latitude="25.05" longLatPrecision="1248" longitude="98.833336" name="Jietou Town" pageId="6" pageNumber="133" stateProvince="Yunnan Province">Jietou Town</location>
;
<geoCoordinate id="34F1505DFFC3FF89FC5AF9E9B252F9E8" box="[990,1099,1615,1644]" degrees="27" direction="north" minutes="68" orientation="latitude" pageId="6" pageNumber="133" precision="925" value="27.668">25°03 N</geoCoordinate>
,
<geoCoordinate id="34F1505DFFC3FF89FBDCF9E9B2DBF9E8" box="[1112,1218,1615,1643]" degrees="98" direction="east" minutes="50" orientation="longitude" pageId="6" pageNumber="133" precision="925" value="98.833336">98°50 E</geoCoordinate>
;
<date id="257B105AFFC3FF89FB4BF9E9B36BF9E8" box="[1231,1394,1617,1643]" pageId="6" pageNumber="133" value="2013-08-11">
<collectingDate id="353FE9B2FFC3FF89FB4BF9E9B36BF9E8" box="[1231,1394,1617,1643]" pageId="6" pageNumber="133" value="2013-08-11">11 Aug. 2013</collectingDate>
</date>
;
<collectorName id="FC30534CFFC3FF89FF39F9CCB79CF90C" box="[189,389,1652,1679]" pageId="6" pageNumber="133">Wei-Cheng Yang</collectorName>
leg.;
<collectionCode id="37D4AE5FFFC3FF89FE43F9CCB408F90D" box="[455,529,1652,1678]" pageId="6" pageNumber="133">IEGU</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="517A369AFFC3FF89FF6EF90EB770F953" blockId="6.[189,1003,1718,1779]" box="[234,361,1718,1744]" pageId="6" pageNumber="133">
<emphasis id="63B1EA88FFC3FF89FF6EF90EB770F953" bold="true" box="[234,361,1718,1744]" pageId="6" pageNumber="133">
<typeStatus id="8E7E8838FFC3FF89FF6EF90EB770F953" box="[234,361,1718,1744]" pageId="6" pageNumber="133" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="517A369AFFC3FF89FF39F961B5F2F970" blockId="6.[189,1003,1718,1779]" box="[189,1003,1752,1779]" pageId="6" pageNumber="133">
<materialsCitation id="E1AD3CC7FFC3FF89FF39F961B5FEF970" ID-GBIF-Occurrence="4903684307" box="[189,999,1752,1779]" collectingDate="2013-08-11" collectionCode="IEGU" collectorName="Wei-Cheng Yang" country="China" county="Tengchong County" latitude="25.05" location="Yunnan Province" longLatPrecision="1248" longitude="98.833336" pageId="6" pageNumber="133" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="paratype">
<collectingCountry id="29D2760AFFC3FF89FF39F961B701F970" box="[189,280,1753,1779]" name="China" pageId="6" pageNumber="133">CHINA</collectingCountry>
<emphasis id="63B1EA88FFC3FF89FEB2F961B414F971" bold="true" box="[310,525,1752,1779]" pageId="6" pageNumber="133">
<collectingRegion id="9301F878FFC3FF89FEB2F961B414F971" box="[310,525,1752,1779]" country="China" name="Yunnan" pageId="6" pageNumber="133">Yunnan Province</collectingRegion>
</emphasis>
<specimenCount id="47C3FD13FFC3FF89FDA2F961B44CF970" box="[550,597,1753,1779]" count="1" pageId="6" pageNumber="133" type="male">1 ♂</specimenCount>
; same data as for holotype;
<collectionCode id="37D4AE5FFFC3FF89FC19F961B5FEF970" box="[925,999,1753,1779]" pageId="6" pageNumber="133">IEGU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC3FF87FF39F8A2B704FAB9" lastPageId="8" lastPageNumber="135" pageId="6" pageNumber="133" type="description">
<paragraph id="517A369AFFC3FF89FF39F8A2B756F8B7" blockId="6.[189,335,1818,1844]" box="[189,335,1818,1844]" pageId="6" pageNumber="133">
<heading id="0A3281F6FFC3FF89FF39F8A2B756F8B7" bold="true" box="[189,335,1818,1844]" fontSize="11" level="3" pageId="6" pageNumber="133" reason="3">
<emphasis id="63B1EA88FFC3FF89FF39F8A2B756F8B7" bold="true" box="[189,335,1818,1844]" pageId="6" pageNumber="133">Description</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC3FF89FF39F8F1B22EF8E0" blockId="6.[189,1079,1865,1892]" box="[189,1079,1865,1892]" pageId="6" pageNumber="133">
<smallCapsWord id="579CA046FFC3FF89FF39F8F1B777F8E2" baselines="1884,1884" box="[189,366,1865,1891]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="6" pageNumber="133">MEASUREMENTS</smallCapsWord>
. Body length including forewing: male
<quantity id="963D9B7FFFC3FF89FCC1F8F1B5CBF8E0" box="[837,978,1865,1892]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="6" pageNumber="133" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.59.5 mm</quantity>
(N = 2).
</paragraph>
<paragraph id="517A369AFFC3FF87FF39F832B361FE33" blockId="6.[189,1399,1930,1992]" lastBlockId="8.[189,1400,263,432]" lastPageId="8" lastPageNumber="135" pageId="6" pageNumber="133">
<smallCapsWord id="579CA046FFC3FF89FF39F832B751F820" baselines="1949,1950" box="[189,328,1930,1956]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="6" pageNumber="133">COLORATION</smallCapsWord>
. General color milk-white to brown (
<figureCitation id="C9FE2A1FFFC3FF89FCA7F832B5AAF827" box="[803,947,1930,1956]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="6" pageNumber="133">Figs 1314</figureCitation>
). Head and pronotum fuscous. Eyes reddish-brown, ocelli light red. Genae pale brown. Tegula and mesonotum light brown. Forewings milk-white, semihyaline, clavus apex ½ to costal margin with broad brown marking, ⅓ and apical part of costal cell each with large spot (former smaller than latter), with brown spot near MP bifurcation, apex of radial aera with markings, medial aera with 5 small round brown spots, between MP4 and MP5 with light brown stripe at terminal transverse vein, several irregular light brown to brown spots along veins near middle, veins gray. Hindwings nearly hyaline, veins pale grayish brown. Legs and abdomen brown.
</paragraph>
<caption id="05BA6612FFC2FF88FF39F8D9B56EF865" ID-DOI="http://doi.org/10.5281/zenodo.12656826" ID-Zenodo-Dep="12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="7" pageNumber="134" startId="7.[189,242,1889,1915]" targetBox="[208,1385,374,1829]" targetPageId="7">
<paragraph id="517A369AFFC2FF88FF39F8D9B56EF865" blockId="7.[187,1399,1889,2022]" pageId="7" pageNumber="134">
<emphasis id="63B1EA88FFC2FF88FF39F8D9B754F8FF" bold="true" box="[189,333,1889,1916]" pageId="7" pageNumber="134">Figs 1323.</emphasis>
<taxonomicName id="96C54D19FFC2FF88FEDCF8DAB42BF8FF" authorityName="Lv &amp; Long &amp; Yang &amp; Chen" authorityYear="2024" box="[344,562,1890,1916]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="134" phylum="Arthropoda" rank="species" species="saccata" status="sp. nov.">
<emphasis id="63B1EA88FFC2FF88FEDCF8DAB42BF8FF" box="[344,562,1890,1916]" italics="true" pageId="7" pageNumber="134">Catonidia saccata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="788257F3FFC2FF88FDB9F8DAB487F8FF" box="[573,670,1890,1916]" pageId="7" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (IEGU).
<emphasis id="63B1EA88FFC2FF88FC2BF8DAB5D5F8FF" bold="true" box="[943,972,1890,1916]" pageId="7" pageNumber="134">13</emphasis>
. Habitus, dorsal view.
<emphasis id="63B1EA88FFC2FF88FB61F8DAB31BF8FF" bold="true" box="[1253,1282,1890,1916]" pageId="7" pageNumber="134">14</emphasis>
. Habitus, lateral view.
<emphasis id="63B1EA88FFC2FF88FEDEF83DB76CF81C" bold="true" box="[346,373,1925,1951]" pageId="7" pageNumber="134">15</emphasis>
. Head and thorax, dorsal view.
<emphasis id="63B1EA88FFC2FF88FD7DF83DB50DF81C" bold="true" box="[761,788,1925,1951]" pageId="7" pageNumber="134">16</emphasis>
. Frons, ventral view.
<emphasis id="63B1EA88FFC2FF88FB9AF83DB220F81C" bold="true" box="[1054,1081,1925,1951]" pageId="7" pageNumber="134">17</emphasis>
. Forewing.
<emphasis id="63B1EA88FFC2FF88FB49F83DB2F1F81C" bold="true" box="[1229,1256,1925,1951]" pageId="7" pageNumber="134">18</emphasis>
. Hindwing.
<emphasis id="63B1EA88FFC2FF88FF3BF810B6C3F841" bold="true" box="[191,218,1960,1986]" pageId="7" pageNumber="134">19</emphasis>
. Pygofer, lateral view.
<emphasis id="63B1EA88FFC2FF88FE6DF810B41FF841" bold="true" box="[489,518,1960,1986]" pageId="7" pageNumber="134">20</emphasis>
. Pygofer, ventral view.
<emphasis id="63B1EA88FFC2FF88FC99F810B523F841" bold="true" box="[797,826,1960,1986]" pageId="7" pageNumber="134">21</emphasis>
. Left gonostyli, inner surface.
<emphasis id="63B1EA88FFC2FF88FB27F810B2D9F841" bold="true" box="[1187,1216,1960,1986]" pageId="7" pageNumber="134">22</emphasis>
. Aedeagus, left lateral view.
<emphasis id="63B1EA88FFC2FF88FED7F873B769F866" bold="true" box="[339,368,1995,2021]" pageId="7" pageNumber="134">23</emphasis>
. Pygofer, dorsal view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="517A369AFFCDFF87FF39FE60B484FCAD" blockId="8.[187,1401,471,815]" pageId="8" pageNumber="135">
<smallCapsWord id="579CA046FFCDFF87FF39FE60B6E5FE73" baselines="491,491" box="[189,252,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="8" pageNumber="135">HEAD</smallCapsWord>
<smallCapsWord id="579CA046FFCDFF87FE82FE65B72AFE73" baselines="491" box="[262,307,477,496]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="8" pageNumber="135">AND</smallCapsWord>
<smallCapsWord id="579CA046FFCDFF87FEB8FE60B781FE73" baselines="491,491" box="[316,408,472,498]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="8" pageNumber="135">THORAX</smallCapsWord>
. Head (
<figureCitation id="C9FE2A1FFFCDFF87FE7DFE6FB49BFE71" box="[505,642,471,498]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Figs 13, 15</figureCitation>
) including eyes narrower than pronotum (0.5: 1). Vertex wider than long in middle line (3.2: 1), vertex concave, median carina absent, anterior margin angular convex in middle, posterior margin curved. Frons (
<figureCitation id="C9FE2A1FFFCDFF87FD43FDA6B507FDB4" box="[711,798,541,568]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Fig. 16</figureCitation>
) longer in midline than maximum width (1.9: 1), width at base narrower than width at apex (0.8: 1), median carina distinct, lateral margins concave. Postclypeus (
<figureCitation id="C9FE2A1FFFCDFF87FEDDFDDCB7B7FDFD" box="[345,430,612,638]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Fig. 16</figureCitation>
) shorter in middle line than frons (about 0.66: 1), with weak median carina, lateral margins excavate. Pronotum (
<figureCitation id="C9FE2A1FFFCDFF87FDA1FD3FB4ABFD21" box="[549,690,647,674]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Figs 13, 15</figureCitation>
) longer in middle line than vertex (2.5: 1), median carina distinct, lateral carinae raised, not extending to posterior margin. Mesonotum (
<figureCitation id="C9FE2A1FFFCDFF87FBC1FD12B2D1FD46" box="[1093,1224,682,709]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Figs 13, 15</figureCitation>
) about 3.9 × as long as pronotum in midline, longer than pronotum and vertex combined (2.8: 1), with weak median and lateral carinae. Forewings (
<figureCitation id="C9FE2A1FFFCDFF87FE7FFD49B449FC88" box="[507,592,753,779]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Fig. 17</figureCitation>
) broadened apically, longer than maximal width (2.1: 1). Hindwings (
<figureCitation id="C9FE2A1FFFCDFF87FF42FCACB703FCAD" box="[198,282,788,814]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Fig. 18</figureCitation>
) longer than widest part (1.4: 1).
</paragraph>
<paragraph id="517A369AFFCDFF87FF39FCEEB704FAB9" blockId="8.[188,1401,853,1338]" pageId="8" pageNumber="135">
<smallCapsWord id="579CA046FFCDFF87FF39FCEEB6E6FCED" baselines="873,873" box="[189,255,854,880]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="8" pageNumber="135">MALE</smallCapsWord>
<smallCapsWord id="579CA046FFCDFF87FE8FFCE3B760FCED" baselines="873" box="[267,377,859,878]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="8" pageNumber="135">GENITALIA</smallCapsWord>
. Pygofer (
<figureCitation id="C9FE2A1FFFCDFF87FE7FFCEDB4AAFCF3" box="[507,691,853,880]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Figs 1920, 23</figureCitation>
) in lateral view with dorsal margin distinctly shorter than ventral margin, ⅓ of posterior margin strongly angulated caudad; in lateral view, lateral margins of medioventral process nearly parallel, apical margin truncate; in ventral view, apex of medioventral process with dentate process outward, apical margin wide and curved. Anal tube (
<figureCitation id="C9FE2A1FFFCDFF87FB14FC07B300FC5A" box="[1168,1305,959,985]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Figs 19, 23</figureCitation>
) longer than widest part by about 1.1:
<quantity id="963D9B7FFFCDFF87FDA8FC5AB443FC7F" box="[556,602,994,1020]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="135" unit="in" value="1.0">1 in</quantity>
dorsal view, apical margin broadly rounded, basal margin concave medially. Aedeagus (
<figureCitation id="C9FE2A1FFFCDFF87FE35FBBDB41DFBA3" box="[433,516,1029,1056]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Fig. 22</figureCitation>
) symmetrical, phallobase in lateral view broad at base, dorsal lobe vestigial, lateral lobes narrowing apically, curving dorsad, with labial lobule near middle, dorsoventral margin of subapical segment slightly furled, angular at apex; ventral lobe shorter, with basal part constricted into stalk, apical part sac-like, lateral sides with 2 lamellar processes near middle. Phallobasal conjunctival processes bent nearly 90 degrees toward back at apical part, apical margin rounded, dorsal margin with lamellar process at curved region, lateral side with long and large ear-like process along ventral margin at apex. Gonostyli (
<figureCitation id="C9FE2A1FFFCDFF87FE2EFB61B42DFB70" box="[426,564,1241,1267]" captionStart="Figs 1323" captionStartId="7.[189,242,1889,1915]" captionTargetBox="[208,1385,374,1829]" captionTargetId="figure-12@7.[189,1398,365,1851]" captionTargetPageId="7" captionText="Figs 1323. Catonidia saccata sp. nov., ♂, holotype (IEGU). 13. Habitus, dorsal view. 14. Habitus, lateral view. 15. Head and thorax, dorsal view. 16. Frons, ventral view. 17. Forewing. 18. Hindwing. 19. Pygofer, lateral view. 20. Pygofer, ventral view. 21. Left gonostyli, inner surface. 22. Aedeagus, left lateral view. 23. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656826" httpUri="https://zenodo.org/record/12656826/files/figure.png" pageId="8" pageNumber="135">Figs 19, 21</figureCitation>
) longer than wide, base narrow, apical margin broadly rounded, with stout twisted process rising from apical third of dorsal margin, inner side of base with slender, finger-like process.
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<emphasis id="63B1EA88FFCDFF87FF39FAD8B741FAF9" bold="true" box="[189,344,1376,1402]" pageId="8" pageNumber="135">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCDFF87FF39FA37B442FA2A" blockId="8.[189,603,1423,1449]" box="[189,603,1423,1449]" pageId="8" pageNumber="135">
<collectingCountry id="29D2760AFFCDFF87FF39FA37B71AFA2A" box="[189,259,1423,1449]" name="China" pageId="8" pageNumber="135">China</collectingCountry>
(
<collectingRegion id="9301F878FFCDFF87FE96FA37B7C7FA2A" box="[274,478,1423,1449]" country="China" name="Yunnan" pageId="8" pageNumber="135">Yunnan province</collectingRegion>
) (
<figureCitation id="C9FE2A1FFFCDFF87FE72FA37B454FA2A" box="[502,589,1423,1449]" captionStart="Fig" captionStartId="13.[189,232,1998,2024]" captionTargetBox="[192,1388,1015,1936]" captionTargetId="figure-224@13.[189,1398,996,1960]" captionTargetPageId="13" captionText="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi="http://doi.org/10.5281/zenodo.12656834" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="8" pageNumber="135">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCDFF87FF39FA69B525F94A" pageId="8" pageNumber="135" type="discussion">
<paragraph id="517A369AFFCDFF87FF39FA69B729FA68" blockId="8.[189,304,1489,1515]" box="[189,304,1489,1515]" pageId="8" pageNumber="135">
<heading id="0A3281F6FFCDFF87FF39FA69B729FA68" bold="true" box="[189,304,1489,1515]" fontSize="11" level="3" pageId="8" pageNumber="135" reason="3">
<emphasis id="63B1EA88FFCDFF87FF39FA69B729FA68" bold="true" box="[189,304,1489,1515]" pageId="8" pageNumber="135">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCDFF87FF39FA47B525F94A" blockId="8.[188,1399,1535,1738]" pageId="8" pageNumber="135">
This species is similar to
<taxonomicName id="96C54D19FFCDFF87FE73FA47B561F99A" authority="Chen &amp; He, 2009" authorityName="Chen &amp; He" authorityYear="2009" box="[503,888,1535,1562]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="lii">
<emphasis id="63B1EA88FFCDFF87FE73FA47B488F99A" box="[503,657,1535,1561]" italics="true" pageId="8" pageNumber="135">Catonidia lii</emphasis>
<bibRefCitation id="35544B6BFFCDFF87FD18FA47B561F99A" author="Chen X. S. &amp; He T. T." box="[668,888,1535,1562]" pageId="8" pageNumber="135" pagination="43 - 52" refId="ref7528" refString="Chen X. S. &amp; He T. T. 2009. Two new species of Catonidia Uhler (Hemiptera: Fulgoromorpha: Achilidae) from southwestern China, with the first description of the male of Catonidia wuyishanana Wang &amp; Huang. Zootaxa 2197: 43 - 52. https: // doi. org / 10.11646 / zootaxa. 2197.1.4" type="journal article" year="2009">Chen &amp; He, 2009</bibRefCitation>
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, but differs from the latter as follows: (1) clavus of forewings apex ½ to costal margin with a broad brown marking (clavus of forewings apex ½ to costal margin without a broad brown marking in
<taxonomicName id="96C54D19FFCDFF87FC8BF9FEB550F9DC" authorityName="Chen &amp; He" authorityYear="2009" box="[783,841,1605,1632]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="lii">
<emphasis id="63B1EA88FFCDFF87FC8BF9FEB550F9DC" box="[783,841,1605,1632]" italics="true" pageId="8" pageNumber="135">C. lii</emphasis>
</taxonomicName>
); (2) medioventral process in lateral view stout, apical margin truncate (medioventral process in lateral view slender, apical margin rounded in
<taxonomicName id="96C54D19FFCDFF87FAADF9D1B37DF901" authorityName="Chen &amp; He" authorityYear="2009" box="[1321,1380,1640,1667]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="lii">
<emphasis id="63B1EA88FFCDFF87FAADF9D1B37DF901" box="[1321,1380,1640,1667]" italics="true" pageId="8" pageNumber="135">C. lii</emphasis>
</taxonomicName>
); (3) lateral sides of ventral lobe with 2 lamellar processes near the middle (lateral sides of ventral lobe without 2 lamellar processes near the middle in
<taxonomicName id="96C54D19FFCDFF87FD6BF917B532F94A" authorityName="Chen &amp; He" authorityYear="2009" box="[751,811,1711,1737]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="135" phylum="Arthropoda" rank="species" species="lii">
<emphasis id="63B1EA88FFCDFF87FD6BF917B532F94A" box="[751,811,1711,1737]" italics="true" pageId="8" pageNumber="135">C. lii</emphasis>
</taxonomicName>
).
</paragraph>
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</treatment>
</document>

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<mods:title id="00A2D070EB95D4F12587DECCE818E088">Four new species of the planthopper genus Catonidia Uhler, 1896 from China (Hemiptera: Fulgoromorpha: Achilidae)</mods:title>
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<figureCitation id="C9FE2A1FFFCCFF86FD46FEF7B551FEE9" box="[706,840,335,362]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Figs 2435</figureCitation>
,
<figureCitation id="C9FE2A1FFFCCFF86FCD0FEF7B568FEEA" box="[852,881,335,361]" captionStart="Fig" captionStartId="13.[189,232,1998,2024]" captionTargetBox="[192,1388,1015,1936]" captionTargetId="figure-224@13.[189,1398,996,1960]" captionTargetPageId="13" captionText="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi="http://doi.org/10.5281/zenodo.12656834" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="9" pageNumber="136">47</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCCFF86FF39FE28B744FD09" pageId="9" pageNumber="136" type="diagnosis">
<paragraph id="517A369AFFCCFF86FF39FE28B72EFE29" blockId="9.[189,311,400,426]" box="[189,311,400,426]" pageId="9" pageNumber="136">
<heading id="0A3281F6FFCCFF86FF39FE28B72EFE29" bold="true" box="[189,311,400,426]" fontSize="11" level="3" pageId="9" pageNumber="136" reason="3">
<emphasis id="63B1EA88FFCCFF86FF39FE28B72EFE29" bold="true" box="[189,311,400,426]" pageId="9" pageNumber="136">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCCFF86FF39FE07B744FD09" blockId="9.[189,1399,446,650]" pageId="9" pageNumber="136">
Salient features of new species include: mesonotum (
<figureCitation id="C9FE2A1FFFCCFF86FCB3FE07B5D8FE5A" box="[823,961,447,473]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Figs 24, 26</figureCitation>
) light yellowish brown at the top ⅓, ⅔ yellowish brown; apical margin of medioventral process (
<figureCitation id="C9FE2A1FFFCCFF86FCF1FE5AB5D1FE7E" box="[885,968,482,509]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Fig. 31</figureCitation>
) angular concave medially in ventral view; aedeagus (
<figureCitation id="C9FE2A1FFFCCFF86FEFAFDBEB41AFDA3" box="[382,515,518,544]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Figs 3334</figureCitation>
) asymmetrical at end, in left lateral view, dorsal lobe long finger-like, lateral lobes divide into three lamellar processes apically; basal lateral ⅓ of ventral lobe with wing-shaped process; in right lateral view, lateral lobes without three lamellar processes apically, narrowing apically, acute at apex.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCCFF86FF39FD09B207FC9E" pageId="9" pageNumber="136" type="etymology">
<paragraph id="517A369AFFCCFF86FF39FD09B75CFD48" blockId="9.[189,325,689,715]" box="[189,325,689,715]" pageId="9" pageNumber="136">
<heading id="0A3281F6FFCCFF86FF39FD09B75CFD48" bold="true" box="[189,325,689,715]" fontSize="11" level="3" pageId="9" pageNumber="136" reason="3">
<emphasis id="63B1EA88FFCCFF86FF39FD09B75CFD48" bold="true" box="[189,325,689,715]" pageId="9" pageNumber="136">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCCFF86FF39FD67B207FC9E" blockId="9.[189,1399,735,797]" pageId="9" pageNumber="136">
The specific epithet is derived from the Latin word
<emphasis id="63B1EA88FFCCFF86FCC5FD67B5ADFD7A" box="[833,948,735,761]" italics="true" pageId="9" pageNumber="136">trilobatus</emphasis>
, referring to the lateral lobes of the phallobase in left lateral view being divided into three lamellar processes.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCCFF86FF39FCFCB2AAFB47" pageId="9" pageNumber="136" type="materials_examined">
<paragraph id="517A369AFFCCFF86FF39FCFCB774FCDD" blockId="9.[189,365,836,862]" box="[189,365,836,862]" pageId="9" pageNumber="136">
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<emphasis id="63B1EA88FFCCFF86FF39FCFCB774FCDD" bold="true" box="[189,365,836,862]" pageId="9" pageNumber="136">Type material</emphasis>
</heading>
</paragraph>
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<typeStatus id="8E7E8838FFCCFF86FF6EFCCBB744FC0E" box="[234,349,883,909]" pageId="9" pageNumber="136" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="517A369AFFCCFF86FF39FC2EB792FC50" blockId="9.[189,1399,883,979]" pageId="9" pageNumber="136">
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;
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leg.;
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.
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<paragraph id="517A369AFFCCFF86FF6EFC42B770FB97" blockId="9.[189,1399,1018,1221]" box="[234,361,1018,1044]" pageId="9" pageNumber="136">
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<typeStatus id="8E7E8838FFCCFF86FF6EFC42B770FB97" box="[234,361,1018,1044]" pageId="9" pageNumber="136" type="paratype">Paratypes</typeStatus>
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</paragraph>
<paragraph id="517A369AFFCCFF86FF39FBA6B2AAFB47" blockId="9.[189,1399,1018,1221]" pageId="9" pageNumber="136">
<materialsCitation id="E1AD3CC7FFCCFF86FF39FBA6B435FBD8" ID-GBIF-Occurrence="4903684302" collectingDate="1998-05-30" collectionCode="IEGU" collectorName="Zi-Zhong Li" country="China" latitude="25.4" location="Maolan National Natural Reserve" longitude="117.742" pageId="9" pageNumber="136" specimenCount="1" specimenCount-female="1" stateProvince="Guizhou Province" typeStatus="paratype">
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<collectingRegion id="9301F878FFCCFF86FEBEFBA5B404FBB4" box="[314,541,1053,1079]" country="China" name="Guizhou" pageId="9" pageNumber="136">Guizhou Province</collectingRegion>
</emphasis>
<specimenCount id="47C3FD13FFCCFF86FDBCFBA6B47FFBBB" box="[568,614,1054,1080]" count="1" pageId="9" pageNumber="136" type="female">1 ♀</specimenCount>
;
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;
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;
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;
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leg.;
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;
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;
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,
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<subSubSection id="19DF6511FFCCFF84FF39FB53B28FFC88" lastPageId="11" lastPageNumber="138" pageId="9" pageNumber="136" type="description">
<paragraph id="517A369AFFCCFF86FF39FB53B756FA86" blockId="9.[189,335,1259,1285]" box="[189,335,1259,1285]" pageId="9" pageNumber="136">
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<emphasis id="63B1EA88FFCCFF86FF39FB53B756FA86" bold="true" box="[189,335,1259,1285]" pageId="9" pageNumber="136">Description</emphasis>
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. Body length including forewing: male
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(N = 3), female
<quantity id="963D9B7FFFCCFF86FB4EFAA2B36EFAB6" box="[1226,1399,1306,1333]" metricMagnitude="-2" metricUnit="m" metricValue="1.105" metricValueMax="1.17" metricValueMin="1.04" pageId="9" pageNumber="136" unit="mm" value="11.05" valueMax="11.7" valueMin="10.4">10.411.7 mm</quantity>
(N = 3).
</paragraph>
<paragraph id="517A369AFFCCFF86FF39FAC7B70BF9CA" blockId="9.[188,1399,1407,1609]" pageId="9" pageNumber="136">
<smallCapsWord id="579CA046FFCCFF86FF39FAC7B751FA14" baselines="1426,1426" box="[189,328,1407,1433]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="9" pageNumber="136">COLORATION</smallCapsWord>
. General color yellowish brown (
<figureCitation id="C9FE2A1FFFCCFF86FD54FAC7B540FA19" box="[720,857,1407,1434]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Figs 2425</figureCitation>
). Vertex and frons pale yellowish brown. Eyes reddish brown, ocelli light red. Genae pale brown. Pronotum and tegula yellowish brown. Mesonotum light yellowish brown at top ⅓, ⅔ yellowish brown. Forewings yellowish brown, large number of unclear grayish brown and dark spots distributed along veins, clavus terminally with dark brown marking, veins yellowish brown. Hindwings nearly hyaline, veins pale grayish brown. Legs and abdomen yellowish brown.
</paragraph>
<paragraph id="517A369AFFCCFF84FF39F9C9B5CCFEC5" blockId="9.[188,1401,1648,1956]" lastBlockId="11.[189,1399,265,327]" lastPageId="11" lastPageNumber="138" pageId="9" pageNumber="136">
<smallCapsWord id="579CA046FFCCFF86FF39F9C9B6E5F90A" baselines="1668,1668" box="[189,252,1649,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="9" pageNumber="136">HEAD</smallCapsWord>
<smallCapsWord id="579CA046FFCCFF86FE82F9CEB72AF90A" baselines="1668" box="[262,307,1654,1673]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="9" pageNumber="136">AND</smallCapsWord>
<smallCapsWord id="579CA046FFCCFF86FEB8F9C9B781F90A" baselines="1668,1668" box="[316,408,1649,1675]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="9" pageNumber="136">THORAX</smallCapsWord>
. Head (
<figureCitation id="C9FE2A1FFFCCFF86FE7DF9C8B49AF909" box="[505,643,1648,1674]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Figs 24, 26</figureCitation>
) including eyes narrower than pronotum (0.6: 1). Vertex wider than long in middle line (1.9: 1), vertex concave, median carina present at base, anterior margin angular, posterior margin subangular concave, lateral margins ridged. Frons (
<figureCitation id="C9FE2A1FFFCCFF86FC64F90FB22FF952" box="[992,1078,1719,1745]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Fig. 27</figureCitation>
) longer in middle line than maximum width (2.1: 1), width at base narrower than width at apex (0.8: 1), median carina distinct, lateral margins concave. Postclypeus (
<figureCitation id="C9FE2A1FFFCCFF86FD06F945B4C0F894" box="[642,729,1789,1815]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Fig. 27</figureCitation>
) shorter in middle line than frons (about 0.5: 1), median carina distinct, lateral margins excavate. Pronotum (
<figureCitation id="C9FE2A1FFFCCFF86FCB8F898B5D2F8B9" box="[828,971,1824,1850]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Figs 24, 26</figureCitation>
) longer in middle line than vertex (1.4: 1), median carina distinct, lateral carinae raised, not extending to posterior margin, confluent with midline at anterior margin to form angular process. Mesonotum (
<figureCitation id="C9FE2A1FFFCCFF86FC49F8DFB241F802" box="[973,1112,1895,1921]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="9" pageNumber="136">Figs 24, 26</figureCitation>
) about 3.9 × as long as pronotum in midline, longer than pronotum and vertex combined (2.3: 1), apex of median carina unclear, lateral carinae reaching posterior margin. Forewings (
<figureCitation id="C9FE2A1FFFCEFF84FCB7FEB1B590FEA0" box="[819,905,265,291]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="11" pageNumber="138">Fig. 28</figureCitation>
) broadened apically, longer than maximal width (2.4: 1). Hindwings (
<figureCitation id="C9FE2A1FFFCEFF84FE79FE94B44AFEC5" box="[509,595,300,326]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="11" pageNumber="138">Fig. 29</figureCitation>
) longer than widest part (1.6: 1).
</paragraph>
<caption id="05BA6612FFCFFF85FF39F8D9B31CF865" ID-DOI="http://doi.org/10.5281/zenodo.12656830" ID-Zenodo-Dep="12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="10" pageNumber="137" startId="10.[189,242,1889,1915]" targetBox="[204,1392,372,1843]" targetPageId="10">
<paragraph id="517A369AFFCFFF85FF39F8D9B31CF865" blockId="10.[189,1399,1889,2022]" pageId="10" pageNumber="137">
<emphasis id="63B1EA88FFCFFF85FF39F8D9B755F8FF" bold="true" box="[189,332,1889,1916]" pageId="10" pageNumber="137">Figs 2435.</emphasis>
<taxonomicName id="96C54D19FFCFFF85FEDCF8DAB426F8FF" authorityName="Lv &amp; Long &amp; Yang &amp; Chen" authorityYear="2024" box="[344,575,1890,1916]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="137" phylum="Arthropoda" rank="species" species="trilobata" status="sp. nov.">
<emphasis id="63B1EA88FFCFFF85FEDCF8DAB426F8FF" box="[344,575,1890,1916]" italics="true" pageId="10" pageNumber="137">Catonidia trilobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="788257F3FFCFFF85FDC3F8DAB4BFF8FF" box="[583,678,1890,1916]" pageId="10" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (IEGU).
<emphasis id="63B1EA88FFCFFF85FC30F8DAB5C8F8FF" bold="true" box="[948,977,1890,1916]" pageId="10" pageNumber="137">24</emphasis>
. Habitus, dorsal view.
<emphasis id="63B1EA88FFCFFF85FB6CF8DAB31DF8FF" bold="true" box="[1256,1284,1890,1916]" pageId="10" pageNumber="137">25</emphasis>
. Habitus, lateral view.
<emphasis id="63B1EA88FFCFFF85FEDEF83DB76CF81C" bold="true" box="[346,373,1925,1951]" pageId="10" pageNumber="137">26</emphasis>
. Head and thorax, dorsal view.
<emphasis id="63B1EA88FFCFFF85FD7DF83DB50DF81C" bold="true" box="[761,788,1925,1951]" pageId="10" pageNumber="137">27</emphasis>
. Frons, ventral view.
<emphasis id="63B1EA88FFCFFF85FB9AF83DB220F81C" bold="true" box="[1054,1081,1925,1951]" pageId="10" pageNumber="137">28</emphasis>
. Forewing.
<emphasis id="63B1EA88FFCFFF85FB49F83DB2F1F81C" bold="true" box="[1229,1256,1925,1951]" pageId="10" pageNumber="137">29</emphasis>
. Hindwing.
<emphasis id="63B1EA88FFCFFF85FF3BF810B6C3F841" bold="true" box="[191,218,1960,1986]" pageId="10" pageNumber="137">30</emphasis>
. Pygofer, lateral view.
<emphasis id="63B1EA88FFCFFF85FE68F810B410F841" bold="true" box="[492,521,1960,1986]" pageId="10" pageNumber="137">31</emphasis>
. Pygofer, ventral view.
<emphasis id="63B1EA88FFCFFF85FCA7F810B559F841" bold="true" box="[803,832,1960,1986]" pageId="10" pageNumber="137">32</emphasis>
. Pygofer, dorsal view.
<emphasis id="63B1EA88FFCFFF85FBD5F810B277F841" bold="true" box="[1105,1134,1960,1986]" pageId="10" pageNumber="137">33</emphasis>
. Aedeagus, left lateral view.
<emphasis id="63B1EA88FFCFFF85FE87F873B739F866" bold="true" box="[259,288,1995,2021]" pageId="10" pageNumber="137">34</emphasis>
. Aedeagus, right lateral view.
<emphasis id="63B1EA88FFCFFF85FD01F873B4BBF866" bold="true" box="[645,674,1995,2021]" pageId="10" pageNumber="137">35</emphasis>
. Left gonostyli, inner surface. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="517A369AFFCEFF84FF39FED6B28FFC88" blockId="11.[187,1401,366,779]" pageId="11" pageNumber="138">
<smallCapsWord id="579CA046FFCEFF84FF39FED6B6E6FE05" baselines="385,385" box="[189,255,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="11" pageNumber="138">MALE</smallCapsWord>
<smallCapsWord id="579CA046FFCEFF84FE83FECBB76CFE05" baselines="385" box="[263,373,371,390]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="11" pageNumber="138">GENITALIA</smallCapsWord>
. Pygofer (
<figureCitation id="C9FE2A1FFFCEFF84FE75FED6B460FE0B" box="[497,633,366,392]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="11" pageNumber="138">Figs 3032</figureCitation>
) in lateral view with dorsal margin distinctly shorter than ventral margin, ⅓ of posterior margin strongly angulated caudad; in ventral view, apical margin of medioventral process angular concave medially. Anal tube (
<figureCitation id="C9FE2A1FFFCEFF84FD61FE0CB569FE4D" box="[741,880,436,462]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="11" pageNumber="138">Figs 30, 32</figureCitation>
) longer than widest part by about 1.6:
<quantity id="963D9B7FFFCEFF84FAC3FE0CB36EFE4D" box="[1351,1399,436,462]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="138" unit="in" value="1.0">1 in</quantity>
dorsal view, apical margin rounded and convex, basal margin slightly truncate. Aedeagus (
<figureCitation id="C9FE2A1FFFCEFF84FB6EFE6FB368FE72" box="[1258,1393,471,497]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="11" pageNumber="138">Figs 3334</figureCitation>
) asymmetrical at end, in left lateral view, phallobase broad at base, dorsal lobe long finger-like, lateral lobes divide into three lamellar processes apically, apical part curved dorsad; ventral lobe slender, bends and extends ventrad, pointed apically, basal lateral ⅓ with wing shaped process; apical part of phallobasal conjunctival processes curved dorsad, apical margin rounded, lateral side with long and narrow ear-like process at subapical part; in right lateral view, lateral lobes without three lamellar processes apically, narrowing apically, acute at apex; ear-like process of phallobasal conjunctival processes broad. Gonostyli (
<figureCitation id="C9FE2A1FFFCEFF84FF47FD76B750FD6B" box="[195,329,717,744]" captionStart="Figs 2435" captionStartId="10.[189,242,1889,1915]" captionTargetBox="[204,1392,372,1843]" captionTargetId="figure-19@10.[189,1398,365,1851]" captionTargetPageId="10" captionText="Figs 2435. Catonidia trilobata sp. nov., ♂, holotype (IEGU). 24. Habitus, dorsal view. 25. Habitus, lateral view. 26. Head and thorax, dorsal view. 27. Frons, ventral view. 28. Forewing. 29. Hindwing. 30. Pygofer, lateral view. 31. Pygofer, ventral view. 32. Pygofer, dorsal view. 33. Aedeagus, left lateral view. 34. Aedeagus, right lateral view. 35. Left gonostyli, inner surface. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656830" httpUri="https://zenodo.org/record/12656830/files/figure.png" pageId="11" pageNumber="138">Figs 30, 35</figureCitation>
) longer than wide, base narrow, apical margin broadly rounded, with stout twisted process rising from apical third of dorsal margin, inner side of base with finger-like process.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCEFF84FF39FC8AB4CCFCF8" box="[189,725,818,891]" pageId="11" pageNumber="138" type="distribution">
<paragraph id="517A369AFFCEFF84FF39FC8AB741FCCF" blockId="11.[189,344,818,844]" box="[189,344,818,844]" pageId="11" pageNumber="138">
<heading id="0A3281F6FFCEFF84FF39FC8AB741FCCF" bold="true" box="[189,344,818,844]" fontSize="11" level="3" pageId="11" pageNumber="138" reason="3">
<emphasis id="63B1EA88FFCEFF84FF39FC8AB741FCCF" bold="true" box="[189,344,818,844]" pageId="11" pageNumber="138">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCEFF84FF39FCD9B4CCFCF8" blockId="11.[189,725,865,891]" box="[189,725,865,891]" pageId="11" pageNumber="138">
<collectingCountry id="29D2760AFFCEFF84FF39FCD9B71AFCF8" box="[189,259,865,891]" name="China" pageId="11" pageNumber="138">China</collectingCountry>
(
<collectingRegion id="9301F878FFCEFF84FE96FCD9B765FCF8" box="[274,380,865,891]" country="China" name="Guizhou" pageId="11" pageNumber="138">Guizhou</collectingRegion>
,
<collectingRegion id="9301F878FFCEFF84FE0CFCD9B7C4FCF8" box="[392,477,865,891]" country="China" name="Hainan" pageId="11" pageNumber="138">Hainan</collectingRegion>
provinces) (
<figureCitation id="C9FE2A1FFFCEFF84FDF4FCD9B4D1FCF8" box="[624,712,865,891]" captionStart="Fig" captionStartId="13.[189,232,1998,2024]" captionTargetBox="[192,1388,1015,1936]" captionTargetId="figure-224@13.[189,1398,996,1960]" captionTargetPageId="13" captionText="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi="http://doi.org/10.5281/zenodo.12656834" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="11" pageNumber="138">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCEFF84FF39FC1AB4CFFB18" pageId="11" pageNumber="138" type="discussion">
<paragraph id="517A369AFFCEFF84FF39FC1AB729FC3F" blockId="11.[189,304,930,956]" box="[189,304,930,956]" pageId="11" pageNumber="138">
<heading id="0A3281F6FFCEFF84FF39FC1AB729FC3F" bold="true" box="[189,304,930,956]" fontSize="11" level="3" pageId="11" pageNumber="138" reason="3">
<emphasis id="63B1EA88FFCEFF84FF39FC1AB729FC3F" bold="true" box="[189,304,930,956]" pageId="11" pageNumber="138">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCEFF84FF39FC69B4CFFB18" blockId="11.[188,1399,977,1179]" pageId="11" pageNumber="138">
This species is similar to
<taxonomicName id="96C54D19FFCEFF84FE67FC69B58FFC68" authority="Long &amp; Chen, 2012" authorityName="Long &amp; Chen" authorityYear="2012" box="[483,918,977,1003]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="138" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="63B1EA88FFCEFF84FE67FC69B4BAFC68" box="[483,675,977,1003]" italics="true" pageId="11" pageNumber="138">Catonidia choui</emphasis>
<bibRefCitation id="35544B6BFFCEFF84FD2EFC69B58FFC68" author="Long J. K. &amp; Chen X. S." box="[682,918,977,1003]" pageId="11" pageNumber="138" pagination="215 - 221" refId="ref7925" refString="Long J. K. &amp; Chen X. S. 2012. A new species of Catonidia Uhler (Hemiptera: Fulgoromorpha: Achilidae) from China. Entomotaxonomia 34 (2): 215 - 221." type="journal article" year="2012">Long &amp; Chen, 2012</bibRefCitation>
</taxonomicName>
, but differs from the latter as follows: (1) anterior margin of vertex angular (anterior margin of vertex rounded in
<taxonomicName id="96C54D19FFCEFF84FB89FC4CB26BFB8D" authorityName="Long &amp; Chen" authorityYear="2012" box="[1037,1138,1012,1038]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="138" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="63B1EA88FFCEFF84FB89FC4CB26BFB8D" box="[1037,1138,1012,1038]" italics="true" pageId="11" pageNumber="138">C. choui</emphasis>
</taxonomicName>
); (2) apical margin of medioventral process concave medially in ventral view (apical margin of medioventral process convex medially in ventral view in
<taxonomicName id="96C54D19FFCEFF84FD82FB83B472FBD7" authorityName="Long &amp; Chen" authorityYear="2012" box="[518,619,1082,1109]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="138" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="63B1EA88FFCEFF84FD82FB83B472FBD7" box="[518,619,1082,1109]" italics="true" pageId="11" pageNumber="138">C. choui</emphasis>
</taxonomicName>
); (3) aedeagus asymmetrical (aedeagus symmetrical in
<taxonomicName id="96C54D19FFCEFF84FA84FB83B37CFBD7" authorityName="Long &amp; Chen" authorityYear="2012" box="[1280,1381,1082,1109]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="138" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="63B1EA88FFCEFF84FA84FB83B37CFBD7" box="[1280,1381,1082,1109]" italics="true" pageId="11" pageNumber="138">C. choui</emphasis>
</taxonomicName>
); (4) lateral lobes divide into three lamellar processes apically in left lateral view (lateral lobes without three lamellar processes apically in
<taxonomicName id="96C54D19FFCEFF84FDE4FB39B4DCFB18" authorityName="Long &amp; Chen" authorityYear="2012" box="[608,709,1153,1179]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="138" phylum="Arthropoda" rank="species" species="choui">
<emphasis id="63B1EA88FFCEFF84FDE4FB39B4DCFB18" box="[608,709,1153,1179]" italics="true" pageId="11" pageNumber="138">C. choui</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,333 @@
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<mods:title id="F7B3365754C135BFFCCFCCA66032B964">Four new species of the planthopper genus Catonidia Uhler, 1896 from China (Hemiptera: Fulgoromorpha: Achilidae)</mods:title>
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<mods:affiliation id="076A79B452BD1A543532BBEF47E0EF64">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University, Guiyang, Guizhou 550025, P. R. China.</mods:affiliation>
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<subSubSection id="19DF6511FFCEFF84FDF7FB60B5D8FB70" box="[627,961,1240,1267]" pageId="11" pageNumber="138" type="nomenclature">
<paragraph id="517A369AFFCEFF84FDF7FB60B5D8FB70" blockId="11.[349,1240,1240,1337]" box="[627,961,1240,1267]" pageId="11" pageNumber="138">
<heading id="0A3281F6FFCEFF84FDF7FB60B5D8FB70" box="[627,961,1240,1267]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="138" reason="2">
<taxonomicName id="96C54D19FFCEFF84FDF7FB60B546FB70" authorityName="Lv &amp; Long &amp; Yang &amp; Chen" authorityYear="2024" box="[627,863,1240,1267]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="138" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="63B1EA88FFCEFF84FDF7FB60B546FB70" bold="true" box="[627,863,1240,1267]" italics="true" pageId="11" pageNumber="138">Catonidia uncinata</emphasis>
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<taxonomicNameLabel id="788257F3FFCEFF84FCE3FB61B5D8FB70" box="[871,961,1241,1267]" pageId="11" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="517A369AFFCEFF84FD5CFAA7B542FABA" blockId="11.[349,1240,1240,1337]" box="[728,859,1311,1337]" pageId="11" pageNumber="138">
<figureCitation id="C9FE2A1FFFCEFF84FD5CFAA7B542FABA" box="[728,859,1311,1337]" captionStart-0="Figs 3646" captionStart-1="Fig" captionStartId-0="12.[189,242,1889,1915]" captionStartId-1="13.[189,232,1998,2024]" captionTargetBox-0="[209,1391,374,1834]" captionTargetBox-1="[192,1388,1015,1936]" captionTargetId-0="figure-19@12.[189,1398,365,1851]" captionTargetId-1="figure-224@13.[189,1398,996,1960]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." captionText-1="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi-0="http://doi.org/10.5281/zenodo.12656832" figureDoi-1="http://doi.org/10.5281/zenodo.12656834" httpUri-0="https://zenodo.org/record/12656832/files/figure.png" httpUri-1="https://zenodo.org/record/12656834/files/figure.png" pageId="11" pageNumber="138">Figs 3647</figureCitation>
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<paragraph id="517A369AFFCEFF84FF39FAD8B72EFAF9" blockId="11.[189,311,1376,1402]" box="[189,311,1376,1402]" pageId="11" pageNumber="138">
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<emphasis id="63B1EA88FFCEFF84FF39FAD8B72EFAF9" bold="true" box="[189,311,1376,1402]" pageId="11" pageNumber="138">Diagnosis</emphasis>
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Salient features of new species include: forewings (
<figureCitation id="C9FE2A1FFFCEFF84FC9AFA37B5BCFA2A" box="[798,933,1423,1449]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="11" pageNumber="138">Figs 3637</figureCitation>
) transverse veins of costal area grayish white, radial and medial areas with 6 clear brown spots; dorsal lobe of phallobase (
<figureCitation id="C9FE2A1FFFCEFF84FB14FA0AB2FFFA4E" box="[1168,1254,1458,1485]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="11" pageNumber="138">Fig. 45</figureCitation>
) long spiny, lateral lobes transected at apex, curled into inverted hooked process along ventral margin at end, ventral lobe short, middle to apex knife shaped, apical margin blunt, lateral sides with a lamellar process at the basal ⅓.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCEFF84FF39F9E6B2C0F949" pageId="11" pageNumber="138" type="etymology">
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<heading id="0A3281F6FFCEFF84FF39F9E6B75CF9FB" bold="true" box="[189,325,1630,1656]" fontSize="11" level="3" pageId="11" pageNumber="138" reason="3">
<emphasis id="63B1EA88FFCEFF84FF39F9E6B75CF9FB" bold="true" box="[189,325,1630,1656]" pageId="11" pageNumber="138">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCEFF84FF39F934B2C0F949" blockId="11.[189,1399,1676,1738]" pageId="11" pageNumber="138">
The specific epithet is derived from the Latin word
<emphasis id="63B1EA88FFCEFF84FCC6F935B5ACF925" box="[834,949,1677,1702]" italics="true" pageId="11" pageNumber="138">uncinatus</emphasis>
, referring to the lateral lobes of the phallobase being curled into an inverted hooked process along the ventral margin at apex.
</paragraph>
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<paragraph id="517A369AFFCEFF84FF39F949B774F889" blockId="11.[189,365,1776,1803]" box="[189,365,1776,1803]" pageId="11" pageNumber="138">
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<emphasis id="63B1EA88FFCEFF84FF39F949B774F889" bold="true" box="[189,365,1776,1803]" pageId="11" pageNumber="138">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCEFF84FF6EF8A7B744F8BA" blockId="11.[189,1398,1823,1920]" box="[234,349,1823,1849]" pageId="11" pageNumber="138">
<emphasis id="63B1EA88FFCEFF84FF6EF8A7B744F8BA" bold="true" box="[234,349,1823,1849]" pageId="11" pageNumber="138">
<typeStatus id="8E7E8838FFCEFF84FF6EF8A7B744F8BA" box="[234,349,1823,1849]" pageId="11" pageNumber="138" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="517A369AFFCEFF84FF39F8FAB496F803" blockId="11.[189,1398,1823,1920]" pageId="11" pageNumber="138">
<materialsCitation id="E1AD3CC7FFCEFF84FF39F8FAB496F803" ID-GBIF-Occurrence="4903684304" collectingDate="2012-07-28" collectionCode="IEGU" collectorName="Wei-Cheng Yang" country="China" latitude="22.033333" location="Xishuangbanna National Natural Reserve" longitude="100.9" pageId="11" pageNumber="138" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="holotype">
<collectingCountry id="29D2760AFFCEFF84FF39F8FAB701F8DF" box="[189,280,1858,1884]" name="China" pageId="11" pageNumber="138">CHINA</collectingCountry>
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<collectingRegion id="9301F878FFCEFF84FEBFF8FAB40CF8DF" box="[315,533,1858,1884]" country="China" name="Yunnan" pageId="11" pageNumber="138">Yunnan Province</collectingRegion>
</emphasis>
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<date id="257B105AFFCEFF84FADDF8FAB735F803" pageId="11" pageNumber="138" value="2012-07-28">
<collectingDate id="353FE9B2FFCEFF84FADDF8FAB735F803" pageId="11" pageNumber="138" value="2012-07-28">28 Jul. 2012</collectingDate>
</date>
;
<collectorName id="FC30534CFFCEFF84FEB3F8DDB419F803" box="[311,512,1893,1920]" pageId="11" pageNumber="138">Wei-Cheng Yang</collectorName>
leg.;
<collectionCode id="37D4AE5FFFCEFF84FDC5F8DEB492F803" box="[577,651,1894,1920]" pageId="11" pageNumber="138">IEGU</collectionCode>
.
</materialsCitation>
</paragraph>
<caption id="05BA6612FFC9FF83FF39F8D9B56EF865" ID-DOI="http://doi.org/10.5281/zenodo.12656832" ID-Zenodo-Dep="12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="12" pageNumber="139" startId="12.[189,242,1889,1915]" targetBox="[209,1391,374,1834]" targetPageId="12">
<paragraph id="517A369AFFC9FF83FF39F8D9B56EF865" blockId="12.[187,1399,1889,2022]" pageId="12" pageNumber="139">
<emphasis id="63B1EA88FFC9FF83FF39F8D9B755F8FF" bold="true" box="[189,332,1889,1916]" pageId="12" pageNumber="139">Figs 3646.</emphasis>
<taxonomicName id="96C54D19FFC9FF83FEDCF8DAB459F8FF" authorityName="Lv &amp; Long &amp; Yang &amp; Chen" authorityYear="2024" box="[344,576,1890,1916]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="139" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="63B1EA88FFC9FF83FEDCF8DAB459F8FF" box="[344,576,1890,1916]" italics="true" pageId="12" pageNumber="139">Catonidia uncinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="788257F3FFC9FF83FDC3F8DAB4BFF8FF" box="[583,678,1890,1916]" pageId="12" pageNumber="139" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (IEGU).
<emphasis id="63B1EA88FFC9FF83FC30F8DAB5C8F8FF" bold="true" box="[948,977,1890,1916]" pageId="12" pageNumber="139">36</emphasis>
. Habitus, dorsal view.
<emphasis id="63B1EA88FFC9FF83FB6CF8DAB31DF8FF" bold="true" box="[1256,1284,1890,1916]" pageId="12" pageNumber="139">37</emphasis>
. Habitus, lateral view.
<emphasis id="63B1EA88FFC9FF83FEDEF83DB76CF81C" bold="true" box="[346,373,1925,1951]" pageId="12" pageNumber="139">38</emphasis>
. Head and thorax, dorsal view.
<emphasis id="63B1EA88FFC9FF83FD7DF83DB50DF81C" bold="true" box="[761,788,1925,1951]" pageId="12" pageNumber="139">39</emphasis>
. Frons, ventral view.
<emphasis id="63B1EA88FFC9FF83FB9AF83DB220F81C" bold="true" box="[1054,1081,1925,1951]" pageId="12" pageNumber="139">40</emphasis>
. Forewing.
<emphasis id="63B1EA88FFC9FF83FB49F83DB2F1F81C" bold="true" box="[1229,1256,1925,1951]" pageId="12" pageNumber="139">41</emphasis>
. Hindwing.
<emphasis id="63B1EA88FFC9FF83FF3BF810B6C3F841" bold="true" box="[191,218,1960,1986]" pageId="12" pageNumber="139">42</emphasis>
. Pygofer, lateral view.
<emphasis id="63B1EA88FFC9FF83FE6DF810B41FF841" bold="true" box="[489,518,1960,1986]" pageId="12" pageNumber="139">43</emphasis>
. Pygofer, ventral view.
<emphasis id="63B1EA88FFC9FF83FC99F810B523F841" bold="true" box="[797,826,1960,1986]" pageId="12" pageNumber="139">44</emphasis>
. Left gonostyli, inner surface.
<emphasis id="63B1EA88FFC9FF83FB27F810B2D9F841" bold="true" box="[1187,1216,1960,1986]" pageId="12" pageNumber="139">45</emphasis>
. Aedeagus, left lateral view.
<emphasis id="63B1EA88FFC9FF83FED7F873B769F866" bold="true" box="[339,368,1995,2021]" pageId="12" pageNumber="139">46</emphasis>
. Pygofer, dorsal view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="517A369AFFC8FF82FF6EFEB1B770FEA0" blockId="13.[189,1398,265,361]" box="[234,361,265,291]" pageId="13" pageNumber="140">
<emphasis id="63B1EA88FFC8FF82FF6EFEB1B770FEA0" bold="true" box="[234,361,265,291]" pageId="13" pageNumber="140">
<typeStatus id="8E7E8838FFC8FF82FF6EFEB1B770FEA0" box="[234,361,265,291]" pageId="13" pageNumber="140" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="517A369AFFC8FF82FF39FE94B5BDFEEA" blockId="13.[189,1398,265,361]" pageId="13" pageNumber="140">
<materialsCitation id="E1AD3CC7FFC8FF82FF39FE94B5B9FEEA" ID-GBIF-Occurrence="4903684303" collectingDate="2012-07-28" collectionCode="IEGU" collectorName="Wei-Cheng Yang &amp; Zhi-Min Chang" country="China" latitude="22.033333" location="Xishuangbanna National Natural Reserve" longitude="100.9" pageId="13" pageNumber="140" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan Province" typeStatus="paratype">
<collectingCountry id="29D2760AFFC8FF82FF39FE94B701FEC5" box="[189,280,300,326]" name="China" pageId="13" pageNumber="140">CHINA</collectingCountry>
<emphasis id="63B1EA88FFC8FF82FEB7FE94B413FEC6" bold="true" box="[307,522,299,326]" pageId="13" pageNumber="140">
<collectingRegion id="9301F878FFC8FF82FEB7FE94B413FEC6" box="[307,522,299,326]" country="China" name="Yunnan" pageId="13" pageNumber="140">Yunnan Province</collectingRegion>
</emphasis>
<specimenCount id="47C3FD13FFC8FF82FDA4FE94B457FEC5" box="[544,590,300,326]" count="1" pageId="13" pageNumber="140" type="male">1 ♂</specimenCount>
,
<specimenCount id="47C3FD13FFC8FF82FDDDFE94B49CFEC5" box="[601,645,300,326]" count="1" pageId="13" pageNumber="140" type="female">1 ♀</specimenCount>
;
<location id="541A6041FFC8FF82FD16FE94B263FEC5" LSID="urn:lsid:plazi:treatment:D96C878CFFCEFF81FDF7FB60B233FAAB:541A6041FFC8FF82FD16FE94B263FEC5" box="[658,1146,300,326]" country="China" latitude="22.033333" name="Xishuangbanna National Natural Reserve" pageId="13" pageNumber="140" stateProvince="Yunnan Province">Xishuangbanna National Natural Reserve</location>
;
<geoCoordinate id="34F1505DFFC8FF82FB01FE92B2ECFEC5" box="[1157,1269,298,326]" degrees="22" direction="north" minutes="02" orientation="latitude" pageId="13" pageNumber="140" precision="925" value="22.033333">22°02 N,</geoCoordinate>
<geoCoordinate id="34F1505DFFC8FF82FB7FFE92B368FEC5" box="[1275,1393,298,326]" degrees="22" direction="north" minutes="02" orientation="latitude" pageId="13" pageNumber="140" precision="925" value="22.033333">100°90 E</geoCoordinate>
;
<date id="257B105AFFC8FF82FF39FEF7B748FEEA" box="[189,337,335,361]" pageId="13" pageNumber="140" value="2012-07-28">
<collectingDate id="353FE9B2FFC8FF82FF39FEF7B748FEEA" box="[189,337,335,361]" pageId="13" pageNumber="140" value="2012-07-28">28 Jul. 2012</collectingDate>
</date>
;
<collectorName id="FC30534CFFC8FF82FED8FEF7B43DFEEA" box="[348,548,335,361]" pageId="13" pageNumber="140">Wei-Cheng Yang</collectorName>
and
<collectorName id="FC30534CFFC8FF82FDD9FEF7B50CFEEA" box="[605,789,335,361]" pageId="13" pageNumber="140">Zhi-Min Chang</collectorName>
leg.;
<collectionCode id="37D4AE5FFFC8FF82FCD2FEF7B5B9FEEA" box="[854,928,335,361]" pageId="13" pageNumber="140">IEGU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFC8FF81FF39FE28B56BFC1B" lastPageId="14" lastPageNumber="141" pageId="13" pageNumber="140" type="description">
<paragraph id="517A369AFFC8FF82FF39FE28B756FE29" blockId="13.[189,335,400,426]" box="[189,335,400,426]" pageId="13" pageNumber="140">
<heading id="0A3281F6FFC8FF82FF39FE28B756FE29" bold="true" box="[189,335,400,426]" fontSize="11" level="3" pageId="13" pageNumber="140" reason="3">
<emphasis id="63B1EA88FFC8FF82FF39FE28B756FE29" bold="true" box="[189,335,400,426]" pageId="13" pageNumber="140">Description</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFC8FF82FF39FE07B34DFE5A" blockId="13.[189,1364,447,473]" box="[189,1364,447,473]" pageId="13" pageNumber="140">
<smallCapsWord id="579CA046FFC8FF82FF39FE07B777FE54" baselines="466,466" box="[189,366,447,473]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="13" pageNumber="140">MEASUREMENTS</smallCapsWord>
. Body length including forewing: male
<quantity id="963D9B7FFFC8FF82FCC1FE07B5CBFE5A" box="[837,978,447,473]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="9.3" metricValueMin="8.1" pageId="13" pageNumber="140" unit="mm" value="8.7" valueMax="9.3" valueMin="8.1">8.19.3 mm</quantity>
(N = 2), female
<quantity id="963D9B7FFFC8FF82FB11FE07B2F6FE5A" box="[1173,1263,447,473]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="13" pageNumber="140" unit="mm" value="8.7">8.7 mm</quantity>
(N = 1).
</paragraph>
<paragraph id="517A369AFFC8FF82FF39FDB8B790FD49" blockId="13.[189,1401,512,714]" pageId="13" pageNumber="140">
<smallCapsWord id="579CA046FFC8FF82FF39FDB8B751FD9A" baselines="531,532" box="[189,328,512,538]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="13" pageNumber="140">COLORATION</smallCapsWord>
. General color yellowish brown (
<figureCitation id="C9FE2A1FFFC8FF82FD57FDB8B545FD99" box="[723,860,512,538]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="13" pageNumber="140">Figs 3637</figureCitation>
). Vertex, base of frons and pronotum fuscous. Eyes reddish brown, ocelli light red. Posterior of pronotum milk-white. Tegula light yellowish brown. Forewings yellowish brown, large number of unclear grayish brown and dark spots distributed along veins, transverse veins of costal area grayish white, radial and medial areas with 6 clear brown spots, veins yellowish brown. Hindwings nearly hyaline, veins pale grayish brown. Legs and abdomen yellowish brown.
</paragraph>
<paragraph id="517A369AFFC8FF81FF39FD4AB5CDFE50" blockId="13.[188,1401,754,921]" lastBlockId="14.[189,1400,265,468]" lastPageId="14" lastPageNumber="141" pageId="13" pageNumber="140">
<smallCapsWord id="579CA046FFC8FF82FF39FD4AB6E5FC89" baselines="773,773" box="[189,252,754,780]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="13" pageNumber="140">HEAD</smallCapsWord>
<smallCapsWord id="579CA046FFC8FF82FE82FD4FB72AFC89" baselines="773" box="[262,307,759,778]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="13" pageNumber="140">AND</smallCapsWord>
<smallCapsWord id="579CA046FFC8FF82FEB8FD4AB781FC89" baselines="773,773" box="[316,408,754,780]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="13" pageNumber="140">THORAX</smallCapsWord>
. Head (
<figureCitation id="C9FE2A1FFFC8FF82FE7DFD4AB49BFC8F" box="[505,642,754,780]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="13" pageNumber="140">Figs 36, 38</figureCitation>
) including eyes narrower than pronotum (0.6: 1). Vertex wider than long in middle line (2.7: 1), apex concave, median carina absent, anterior margin angular, posterior margin subangular concave, lateral margins ridged, nearly parallel. Frons (
<figureCitation id="C9FE2A1FFFC8FF82FBC6FC80B282FCD1" box="[1090,1179,824,850]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="13" pageNumber="140">Fig. 39</figureCitation>
) longer in middle line than maximum width (1.5: 1), width at base narrower than width at apex (0.6: 1), median carina distinct, lateral margins concave. Postclypeus (
<figureCitation id="C9FE2A1FFFC8FF82FD63FCC6B527FC1A" box="[743,830,894,921]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="13" pageNumber="140">Fig. 39</figureCitation>
) shorter in middle line than frons (about 0.7: 1), median carina distinct, lateral margins excavate. Pronotum (
<figureCitation id="C9FE2A1FFFCBFF81FC13FEB1B23DFEA0" box="[919,1060,265,291]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 36, 38</figureCitation>
) longer in middle line than vertex (2.4: 1), median carina distinct, lateral carinae raised, not extending to posterior margin, confluent with midline at anterior margin to form angular process. Mesonotum (
<figureCitation id="C9FE2A1FFFCBFF81FC7FFEF7B29BFEEA" box="[1019,1154,335,361]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 36, 38</figureCitation>
) about 3.7 × as long as pronotum in midline, longer than pronotum and vertex combined (2.4: 1), basal of median carina present clearly, lateral carinae unclear. Forewings (
<figureCitation id="C9FE2A1FFFCBFF81FCA6FE2EB562FE33" box="[802,891,406,432]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Fig. 40</figureCitation>
) broadened apically, longer than maximal width (2.1: 1). Hindwings (
<figureCitation id="C9FE2A1FFFCBFF81FE79FE01B44BFE50" box="[509,594,441,467]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Fig. 41</figureCitation>
) longer than widest part (1.5: 1).
</paragraph>
<caption id="05BA6612FFC8FF82FF39F876B5E0F86A" ID-DOI="http://doi.org/10.5281/zenodo.12656834" ID-Zenodo-Dep="12656834" box="[189,1017,1998,2025]" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="13" pageNumber="140" startId="13.[189,232,1998,2024]" targetBox="[192,1388,1015,1936]" targetPageId="13">
<paragraph id="517A369AFFC8FF82FF39F876B5E0F86A" blockId="13.[189,1017,1998,2025]" box="[189,1017,1998,2025]" pageId="13" pageNumber="140">
<emphasis id="63B1EA88FFC8FF82FF39F876B702F86B" bold="true" box="[189,283,1998,2024]" pageId="13" pageNumber="140">Fig. 47.</emphasis>
Geographic distributions of species of
<taxonomicName id="96C54D19FFC8FF82FD63F876B5EBF86B" authority="Uhler, 1896" authorityName="Uhler" authorityYear="1896" box="[743,1010,1998,2024]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="63B1EA88FFC8FF82FD63F876B547F86B" box="[743,862,1998,2024]" italics="true" pageId="13" pageNumber="140">Catonidia</emphasis>
<bibRefCitation id="35544B6BFFC8FF82FCE2F876B5EBF86B" author="Uhler P. R." box="[870,1010,1998,2024]" pageId="13" pageNumber="140" pagination="255 - 297" refId="ref7987" refString="Uhler P. R. 1896. Summary of the Hemiptera of Japan, presented to the United States National Museum by Professor Mitzukuri. Proceedings of the United States National Museum 19 (1108): 255 - 297. https: // doi. org / 10.5479 / si. 00963801.1108.255" type="journal article" year="1896">Uhler, 1896</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="517A369AFFCBFF81FF39FE43B56BFC1B" blockId="14.[187,1399,506,920]" pageId="14" pageNumber="141">
<smallCapsWord id="579CA046FFCBFF81FF39FE43B6E6FD90" baselines="526,526" box="[189,255,507,533]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Male" pageId="14" pageNumber="141">MALE</smallCapsWord>
<smallCapsWord id="579CA046FFCBFF81FE8FFDB8B760FD90" baselines="526" box="[267,377,512,531]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genitalia" pageId="14" pageNumber="141">GENITALIA</smallCapsWord>
. Pygofer (
<figureCitation id="C9FE2A1FFFCBFF81FE7FFE42B4ADFD97" box="[507,692,506,533]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 4243, 46</figureCitation>
) in lateral view with dorsal margin distinctly shorter than ventral margin, ⅓ of posterior margin strongly angulated caudad; in ventral view, apical margin of medioventral process concave medially. Anal tube (
<figureCitation id="C9FE2A1FFFCBFF81FC9FFDF9B5B8FDD8" box="[795,929,577,603]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 42, 46</figureCitation>
) longer than widest part by about 1.5:
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dorsal view, apical margin broadly rounded, basal margin narrow, slightly constricted near base. Aedeagus (
<figureCitation id="C9FE2A1FFFCBFF81FEC0FD3FB787FD21" box="[324,414,647,674]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Fig. 45</figureCitation>
) symmetrical, in lateral view, phallobase narrow at base, dorsal lobe long spiny; lateral lobes transected at apex, curled into inverted hooked process along ventral margin at apex, ventral margin of hook process with small process in the middle; ventral lobe shorter, basal part constricted into stalk, middle to end knife shaped, apical margin blunt, lateral sides with lamellar processes at basal ⅓; apical margin of phallobasal conjunctival processes wide and rounded, lateral sides each with long and large lamellar process along dorsal and ventral margin at subapical area. Gonostyli (
<figureCitation id="C9FE2A1FFFCBFF81FB12FC8FB305FCD2" box="[1174,1308,823,849]" captionStart="Figs 3646" captionStartId="12.[189,242,1889,1915]" captionTargetBox="[209,1391,374,1834]" captionTargetId="figure-19@12.[189,1398,365,1851]" captionTargetPageId="12" captionText="Figs 3646. Catonidia uncinata sp. nov., ♂, holotype (IEGU). 36. Habitus, dorsal view. 37. Habitus, lateral view. 38. Head and thorax, dorsal view. 39. Frons, ventral view. 40. Forewing. 41. Hindwing. 42. Pygofer, lateral view. 43. Pygofer, ventral view. 44. Left gonostyli, inner surface. 45. Aedeagus, left lateral view. 46. Pygofer, dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12656832" httpUri="https://zenodo.org/record/12656832/files/figure.png" pageId="14" pageNumber="141">Figs 42, 44</figureCitation>
) longer than wide, base narrow, apical margin broadly rounded, dorsal margin with stout twisted process near middle, inner side of base with slender, finger-like process.
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<emphasis id="63B1EA88FFCBFF81FF39FC07B741FC5A" bold="true" box="[189,344,959,985]" pageId="14" pageNumber="141">Distribution</emphasis>
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</paragraph>
<paragraph id="517A369AFFCBFF81FF39FC55B442FB8B" blockId="14.[189,603,1005,1032]" box="[189,603,1005,1032]" pageId="14" pageNumber="141">
<collectingCountry id="29D2760AFFCBFF81FF39FC55B71AFB84" box="[189,259,1005,1031]" name="China" pageId="14" pageNumber="141">China</collectingCountry>
(
<collectingRegion id="9301F878FFCBFF81FE96FC56B7C7FB84" box="[274,478,1005,1032]" country="China" name="Yunnan" pageId="14" pageNumber="141">Yunnan province</collectingRegion>
) (
<figureCitation id="C9FE2A1FFFCBFF81FE72FC55B454FB8B" box="[502,589,1005,1032]" captionStart="Fig" captionStartId="13.[189,232,1998,2024]" captionTargetBox="[192,1388,1015,1936]" captionTargetId="figure-224@13.[189,1398,996,1960]" captionTargetPageId="13" captionText="Fig. 47. Geographic distributions of species of Catonidia Uhler, 1896." figureDoi="http://doi.org/10.5281/zenodo.12656834" httpUri="https://zenodo.org/record/12656834/files/figure.png" pageId="14" pageNumber="141">Fig. 47</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="19DF6511FFCBFF81FF39FB97B233FAAB" pageId="14" pageNumber="141" type="discussion">
<paragraph id="517A369AFFCBFF81FF39FB97B729FBCA" blockId="14.[189,304,1071,1097]" box="[189,304,1071,1097]" pageId="14" pageNumber="141">
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<emphasis id="63B1EA88FFCBFF81FF39FB97B729FBCA" bold="true" box="[189,304,1071,1097]" pageId="14" pageNumber="141">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="517A369AFFCBFF81FF39FBE5B233FAAB" blockId="14.[189,1401,1117,1320]" pageId="14" pageNumber="141">
This species is similar to
<taxonomicName id="96C54D19FFCBFF81FE6BFBE5B203FBFB" authority="Wang &amp; Huang, 1990" authorityName="Wang &amp; Huang" authorityYear="1990" box="[495,1050,1117,1144]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="141" phylum="Arthropoda" rank="species" species="wuyishanana">
<emphasis id="63B1EA88FFCBFF81FE6BFBE5B512FBF4" box="[495,779,1117,1143]" italics="true" pageId="14" pageNumber="141">Catonidia wuyishanana</emphasis>
<bibRefCitation id="35544B6BFFCBFF81FC96FBE6B203FBFB" author="Wang S. Z. &amp; Huang J. &amp; Song J. Z." box="[786,1050,1118,1144]" pageId="14" pageNumber="141" pagination="120 - 123" refId="ref8040" refString="Wang S. Z., Huang J. &amp; Song J. Z. 1990. Notes on the Catonidia from China (Homoptera: Achilidae). Acta Agriculturae Boreali-Sinica 5 (4): 120 - 123." type="journal article" year="1990">Wang &amp; Huang, 1990</bibRefCitation>
</taxonomicName>
, but differs from the latter as follows: (1) radial and medial areas of forewing with 6 clear brown spots (radial and medial areas of forewing without 6 clear brown spots in
<taxonomicName id="96C54D19FFCBFF81FD2CFB1CB57EFB3D" authorityName="Wang &amp; Huang" authorityYear="1990" box="[680,871,1188,1214]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="141" phylum="Arthropoda" rank="species" species="wuyishanana">
<emphasis id="63B1EA88FFCBFF81FD2CFB1CB57EFB3D" box="[680,871,1188,1214]" italics="true" pageId="14" pageNumber="141">C. wuyishanana</emphasis>
</taxonomicName>
); (2) lateral lobes transected at apex, curled into an inverted hooked process along the ventral margin at apex (lateral lobes acute at apex, without inverted hooked process along the ventral margin at apex in
<taxonomicName id="96C54D19FFCBFF81FC20FB53B27FFA87" authorityName="Wang &amp; Huang" authorityYear="1990" box="[932,1126,1258,1285]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="141" phylum="Arthropoda" rank="species" species="wuyishanana">
<emphasis id="63B1EA88FFCBFF81FC20FB53B27FFA87" box="[932,1126,1258,1285]" italics="true" pageId="14" pageNumber="141">C. wuyishanana</emphasis>
</taxonomicName>
); (3) apical margin of ventral lobe blunt (apical margin of ventral lobe acute in
<taxonomicName id="96C54D19FFCBFF81FCDEFAB6B200FAA4" authorityName="Wang &amp; Huang" authorityYear="1990" box="[858,1049,1293,1320]" class="Insecta" family="Achilidae" genus="Catonidia" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="141" phylum="Arthropoda" rank="species" species="wuyishanana">
<emphasis id="63B1EA88FFCBFF81FCDEFAB6B200FAA4" box="[858,1049,1293,1320]" italics="true" pageId="14" pageNumber="141">C. wuyishanana</emphasis>
</taxonomicName>
).
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</document>

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@ -1,496 +1,496 @@
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<subSubSection pageId="0" pageNumber="971" type="description">
<paragraph pageId="0" pageNumber="971">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree obtained in IQ-tree based on the concatenated alignment of the two mitochondrial fragments (1616 bp), depicting the phylogenetic relationships of the genus Bothrocophias. Outgroups were removed for clarity. Values above branches indicate Ultrafast Bootstrap support of the ML tree; values below branches indicate posterior probabilities of the Bayesian tree." figureDoi="10.3897/vz.72.e87313.figure1" httpUri="https://binary.pensoft.net/fig/763279" pageId="0" pageNumber="971">Figs 1</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the head pholidosis among the holotypes of Bothrocophias microphthalmus, Bothrocophias tulitoi sp. nov. and Bothrocophias myrringae sp. nov. A, B, C Dorsal, lateral, and ventral view of the head of B. microphthalmus ANSP 11515. D, E, F Dorsal, lateral, and ventral view of the head of B. tulitoi sp. nov. INSZ 073. G, H, I Dorsal, lateral, and ventral view of the head of B. myrringae sp. nov. Scales: dark green = second / third supralabial scale. yellow = lacunolabial / prelacunal. Blue = internasal scales. Fuchsia = scale separating internasal scales." figureDoi="10.3897/vz.72.e87313.figure6" httpUri="https://binary.pensoft.net/fig/763284" pageId="0" pageNumber="971">, 6G-I</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Bothrocophias myrringae sp. nov. in life (INSZ 0268. male). A Dorsal view of the body. B Ventral view of the body. C Dorsal view of the head. D - E Lateral views (right / left) of the head. F Ventral view of the head. Black bar length = 1 cm." figureDoi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">, 8A-C</figureCitation>
<subSubSection id="95125414782587BD1183D7D768E37AD8" pageId="0" pageNumber="971" type="description">
<paragraph id="1286AED4BB5758CE382C7AE660A3BC0D" pageId="0" pageNumber="971">
<figureCitation id="D8B64FBC127EEFAD35624619E48E517D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree obtained in IQ-tree based on the concatenated alignment of the two mitochondrial fragments (1616 bp), depicting the phylogenetic relationships of the genus Bothrocophias. Outgroups were removed for clarity. Values above branches indicate Ultrafast Bootstrap support of the ML tree; values below branches indicate posterior probabilities of the Bayesian tree." figureDoi="10.3897/vz.72.e87313.figure1" httpUri="https://binary.pensoft.net/fig/763279" pageId="0" pageNumber="971">Figs 1</figureCitation>
<figureCitation id="AB32E792CA5A28A5F1329BB7FB108D10" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the head pholidosis among the holotypes of Bothrocophias microphthalmus, Bothrocophias tulitoi sp. nov. and Bothrocophias myrringae sp. nov. A, B, C Dorsal, lateral, and ventral view of the head of B. microphthalmus ANSP 11515. D, E, F Dorsal, lateral, and ventral view of the head of B. tulitoi sp. nov. INSZ 073. G, H, I Dorsal, lateral, and ventral view of the head of B. myrringae sp. nov. Scales: dark green = second / third supralabial scale. yellow = lacunolabial / prelacunal. Blue = internasal scales. Fuchsia = scale separating internasal scales." figureDoi="10.3897/vz.72.e87313.figure6" httpUri="https://binary.pensoft.net/fig/763284" pageId="0" pageNumber="971">, 6G-I</figureCitation>
<figureCitation id="4A78CB5E7A85C19ECB0BAB6078764C20" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Bothrocophias myrringae sp. nov. in life (INSZ 0268. male). A Dorsal view of the body. B Ventral view of the body. C Dorsal view of the head. D - E Lateral views (right / left) of the head. F Ventral view of the head. Black bar length = 1 cm." figureDoi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971">, 7</figureCitation>
<figureCitation id="A66F9B51A0A8934EDB1143932C6D5FEA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">, 8A-C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="chresonymy">
<paragraph pageId="0" pageNumber="971">Chresonymy.</paragraph>
<paragraph pageId="0" pageNumber="971">
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1875" class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias microphthalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias microphthalmus</emphasis>
<subSubSection id="09C0D8365839DD4A6F99EDA864D7C151" pageId="0" pageNumber="971" type="chresonymy">
<paragraph id="3283043FC4EDD14B29CC3006ABDCD1CC" pageId="0" pageNumber="971">Chresonymy.</paragraph>
<paragraph id="351115632C40309100325915213E11EB" pageId="0" pageNumber="971">
<taxonomicName id="3556278F35D1A9A0F160756CC77C14CD" baseAuthorityName="Cope" baseAuthorityYear="1875" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias microphthalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis id="BB9CCD7F8FF8649710E93FBD3CFED03E" italics="true" pageId="0" pageNumber="971">Bothrocophias microphthalmus</emphasis>
</taxonomicName>
.
<bibRefCitation author="Campbell, JA" journalOrPublisher="Campbell JA, Lamar WW (Eds). Comstock Pub. Associates" pageId="0" pageNumber="971" refId="B17" refString="Campbell, JA, Lamar, WW, 1989. The Venomous Reptiles of Latin America. Campbell JA, Lamar WW (Eds). Comstock Pub. Associates" title="The Venomous Reptiles of Latin America." year="1989">Campbell and Lamar (1989)</bibRefCitation>
<bibRefCitation id="F34AFB4D82B77855F9B24B84619088E3" author="Campbell, JA" journalOrPublisher="Campbell JA, Lamar WW (Eds). Comstock Pub. Associates" pageId="0" pageNumber="971" refId="B17" refString="Campbell, JA, Lamar, WW, 1989. The Venomous Reptiles of Latin America. Campbell JA, Lamar WW (Eds). Comstock Pub. Associates" title="The Venomous Reptiles of Latin America." year="1989">Campbell and Lamar (1989)</bibRefCitation>
: page 255, figure 229;
<bibRefCitation author="Campbell, JA" journalOrPublisher="Comstock Publishing, New York" pageId="0" pageNumber="971" refId="B18" refString="Campbell, JA, Lamar, WW, 2004. The venomous reptiles of the western hemisphere (Vol. 1). Comstock Publishing, New York" title="The venomous reptiles of the western hemisphere (Vol. 1)." year="2004">Campbell and Lamar (2004)</bibRefCitation>
<bibRefCitation id="0280CD96951127A61AECCEF54140981D" author="Campbell, JA" journalOrPublisher="Comstock Publishing, New York" pageId="0" pageNumber="971" refId="B18" refString="Campbell, JA, Lamar, WW, 2004. The venomous reptiles of the western hemisphere (Vol. 1). Comstock Publishing, New York" title="The venomous reptiles of the western hemisphere (Vol. 1)." year="2004">Campbell and Lamar (2004)</bibRefCitation>
: Volume 1, Plate 474.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="holotype">
<paragraph pageId="0" pageNumber="971">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="971">
<subSubSection id="F795798EB4FA5CF49C076B4328E70BEC" pageId="0" pageNumber="971" type="holotype">
<paragraph id="97F640093F1FF2EA9D5E43A413B54265" pageId="0" pageNumber="971">Holotype.</paragraph>
<paragraph id="3D5E00E0ECC6E200A0CE65C5F764F0CD" pageId="0" pageNumber="971">
[
<emphasis bold="true" pageId="0" pageNumber="971">
<emphasis id="15155236973968B00A11AFCB9FE72CDF" bold="true" pageId="0" pageNumber="971">
Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree obtained in IQ-tree based on the concatenated alignment of the two mitochondrial fragments (1616 bp), depicting the phylogenetic relationships of the genus Bothrocophias. Outgroups were removed for clarity. Values above branches indicate Ultrafast Bootstrap support of the ML tree; values below branches indicate posterior probabilities of the Bayesian tree." figureDoi="10.3897/vz.72.e87313.figure1" httpUri="https://binary.pensoft.net/fig/763279" pageId="0" pageNumber="971">1</figureCitation>
<figureCitation id="6B5369D7FAC38C0121697A6F483A97BA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Maximum likelihood tree obtained in IQ-tree based on the concatenated alignment of the two mitochondrial fragments (1616 bp), depicting the phylogenetic relationships of the genus Bothrocophias. Outgroups were removed for clarity. Values above branches indicate Ultrafast Bootstrap support of the ML tree; values below branches indicate posterior probabilities of the Bayesian tree." figureDoi="10.3897/vz.72.e87313.figure1" httpUri="https://binary.pensoft.net/fig/763279" pageId="0" pageNumber="971">1</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the head pholidosis among the holotypes of Bothrocophias microphthalmus, Bothrocophias tulitoi sp. nov. and Bothrocophias myrringae sp. nov. A, B, C Dorsal, lateral, and ventral view of the head of B. microphthalmus ANSP 11515. D, E, F Dorsal, lateral, and ventral view of the head of B. tulitoi sp. nov. INSZ 073. G, H, I Dorsal, lateral, and ventral view of the head of B. myrringae sp. nov. Scales: dark green = second / third supralabial scale. yellow = lacunolabial / prelacunal. Blue = internasal scales. Fuchsia = scale separating internasal scales." figureDoi="10.3897/vz.72.e87313.figure6" httpUri="https://binary.pensoft.net/fig/763284" pageId="0" pageNumber="971">6G-I</figureCitation>
<figureCitation id="9035B0A8A3DBA369691933E0E30C5324" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the head pholidosis among the holotypes of Bothrocophias microphthalmus, Bothrocophias tulitoi sp. nov. and Bothrocophias myrringae sp. nov. A, B, C Dorsal, lateral, and ventral view of the head of B. microphthalmus ANSP 11515. D, E, F Dorsal, lateral, and ventral view of the head of B. tulitoi sp. nov. INSZ 073. G, H, I Dorsal, lateral, and ventral view of the head of B. myrringae sp. nov. Scales: dark green = second / third supralabial scale. yellow = lacunolabial / prelacunal. Blue = internasal scales. Fuchsia = scale separating internasal scales." figureDoi="10.3897/vz.72.e87313.figure6" httpUri="https://binary.pensoft.net/fig/763284" pageId="0" pageNumber="971">6G-I</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Bothrocophias myrringae sp. nov. in life (INSZ 0268. male). A Dorsal view of the body. B Ventral view of the body. C Dorsal view of the head. D - E Lateral views (right / left) of the head. F Ventral view of the head. Black bar length = 1 cm." figureDoi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971">7</figureCitation>
<figureCitation id="7BBE2B3F236F483EF24F5F16DC4722EA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Bothrocophias myrringae sp. nov. in life (INSZ 0268. male). A Dorsal view of the body. B Ventral view of the body. C Dorsal view of the head. D - E Lateral views (right / left) of the head. F Ventral view of the head. Black bar length = 1 cm." figureDoi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971">7</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8A-C</figureCitation>
<figureCitation id="B6CB5E1BA15E0E2B82795AB77AB9BD78" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8A-C</figureCitation>
</emphasis>
] INSZ 0268, an adult male from vereda Mundo Nuevo, municipality of La Calera, department of Cundinamarca, Colombia, coordinates:
<geoCoordinate degrees="4.660602778" direction="north" orientation="latitude" precision="1" value="4.6606026">N 4.660602778</geoCoordinate>
<geoCoordinate id="142F0F9D8685F7B597C6160C07624C48" degrees="4.660602778" direction="north" orientation="latitude" precision="1" value="4.6606026">N 4.660602778</geoCoordinate>
, W -73.88491667; elevation 2,761 m. a.s.l. The specimen was collected by local people and brought to Francisco Javier Ruiz, staff of the INS, on 29th August 2020.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="paratypes">
<paragraph pageId="0" pageNumber="971">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="971">
<subSubSection id="CBE051500241B4C28371B7C58E22EC44" pageId="0" pageNumber="971" type="paratypes">
<paragraph id="7426DABBC07EA664B7E6F8CCA1D20D3B" pageId="0" pageNumber="971">Paratypes.</paragraph>
<paragraph id="714A806141C90058D6D3D611CC036573" pageId="0" pageNumber="971">
COLOMBIA [
<emphasis italics="true" pageId="0" pageNumber="971">n</emphasis>
<emphasis id="392C865E266C19FDE32D8DBFE6E61DC4" italics="true" pageId="0" pageNumber="971">n</emphasis>
=5; Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8C-D</figureCitation>
<figureCitation id="74AD9E81EA0D2DB3F9590CDEEE278092" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8C-D</figureCitation>
]:
<emphasis bold="true" pageId="0" pageNumber="971">Cundinamarca</emphasis>
<emphasis id="12F8A2593E6D52EBF6DA510A89AF24F2" bold="true" pageId="0" pageNumber="971">Cundinamarca</emphasis>
: Municipality of Guayabetal. Locality: unknown, IAvH-R 6877, coordinates
<geoCoordinate degrees="4.17508" direction="north" orientation="latitude" precision="1" value="4.17508">N 4.17508</geoCoordinate>
<geoCoordinate id="885C8C5388DC3AB3060E6677A0E981C7" degrees="4.17508" direction="north" orientation="latitude" precision="1" value="4.17508">N 4.17508</geoCoordinate>
, W -73.88117 (approximate to the town). Municipality of
<normalizedToken originalValue="Choachí">Choachi</normalizedToken>
<normalizedToken id="FF9261C424CE4F53B75EF2DB4DDF89B8" originalValue="Choachí">Choachi</normalizedToken>
. Locality: Palo Alto, IAvH-R6840, coordinates
<geoCoordinate degrees="4.61578" direction="north" orientation="latitude" precision="1" value="4.61578">N 4.61578</geoCoordinate>
<geoCoordinate id="92760585DA43B2A79514A1A59A596FA1" degrees="4.61578" direction="north" orientation="latitude" precision="1" value="4.61578">N 4.61578</geoCoordinate>
, W -73.8904. Municipality of
<normalizedToken originalValue="Fómeque">Fomeque</normalizedToken>
<normalizedToken id="B37F862394F28D02CAB89D2F103DE673" originalValue="Fómeque">Fomeque</normalizedToken>
. Locality: vereda de
<normalizedToken originalValue="Coasavistá">Coasavista</normalizedToken>
<normalizedToken id="3C5D88B1F69CEE7943CEEAA21FA7C29A" originalValue="Coasavistá">Coasavista</normalizedToken>
, INSV-SR-00365, coordinates
<geoCoordinate degrees="4.495001" direction="north" orientation="latitude" precision="1" value="4.495001">N 4.495001</geoCoordinate>
<geoCoordinate id="45D92AE90E52CAD02F88FCF8D7BF90D3" degrees="4.495001" direction="north" orientation="latitude" precision="1" value="4.495001">N 4.495001</geoCoordinate>
, W -73.852056. -
<emphasis bold="true" pageId="0" pageNumber="971">Meta</emphasis>
<emphasis id="7F10643F4D7CCAED2EF81BC9CAD52F8A" bold="true" pageId="0" pageNumber="971">Meta</emphasis>
: Municipality of El Calvario. Locality: unknown, INSVSR-0099, coordinates
<geoCoordinate degrees="4.358925" direction="north" orientation="latitude" precision="1" value="4.358925">N 4.358925</geoCoordinate>
<geoCoordinate id="6A0AB93819224AAE5E10CE6528EFDFC0" degrees="4.358925" direction="north" orientation="latitude" precision="1" value="4.358925">N 4.358925</geoCoordinate>
, W -73.71358889. Municipality of San Juanito. Locality: unknown, IAvH-R7045, coordinates
<geoCoordinate degrees="4.457913889" direction="north" orientation="latitude" precision="1" value="4.457914">N 4.457913889</geoCoordinate>
<geoCoordinate id="C89FEF4D213A5AE201B1BB43A9525D9A" degrees="4.457913889" direction="north" orientation="latitude" precision="1" value="4.457914">N 4.457913889</geoCoordinate>
, W -73.67618889 (approximate to the town).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="diagnosis">
<paragraph pageId="0" pageNumber="971">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="971">
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<subSubSection id="47C0F129597E7C0268D6965D408CDFFC" pageId="0" pageNumber="971" type="diagnosis">
<paragraph id="7E96EB1961DAC29CFA8028E19FD1A0E0" pageId="0" pageNumber="971">Diagnosis.</paragraph>
<paragraph id="54ABE7D12EF1E51BD4D03F602EADF985" pageId="0" pageNumber="971">
<taxonomicName id="775947C779F578344D8F8CC68D1E964F" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="CD01EAD1834D8D24B70CAF0C1FEFF1CD" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="FB68FE95E083E4F55369028705D8FEC7" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
can be distinguished from all its congeners by the following combination of characters: (1) 157-161 ventral scales in females, 152-155 ventral scales in males; (2) internasal scales in contact or separated by a single small scale (3) absence of canthorostral scales; (4) lacunolabial scale usually present; (5) hemipenial lobes slim and cylindrical, moderately capitate distally, weakly ornamented toward the apex with large and scarce calyces with weakly spinulate edges; (8) bifurcation point of the hemipenial lobes about 2-4 sudcaudal scales; (9) hemipenial body ornamented by numerous mesial spines that increase in size from the center to periphery of the hemipenial body and arranged in oblique lines; (10) in sulcate view, lateral and mesial spines of the hemipenial body variable in size; (11)
<emphasis italics="true" pageId="0" pageNumber="971">sulcus spermaticus</emphasis>
<emphasis id="FAE4008E083A5F195E874CFBDE09B6F3" italics="true" pageId="0" pageNumber="971">sulcus spermaticus</emphasis>
walls weakly defined; (12) usually more than 28 dark-brown bands and/or opposite or juxtaposed trapezoid-shaped blotches with paler centers dorsally; and (13) ventral surfaces of the tail with bright reddish or orange-reddish speckles with black spots without a regular pattern and heavily marked with dark pigment towards the base (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Bothrocophias myrringae sp. nov. in life (INSZ 0268. male). A Dorsal view of the body. B Ventral view of the body. C Dorsal view of the head. D - E Lateral views (right / left) of the head. F Ventral view of the head. Black bar length = 1 cm." figureDoi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971">7</figureCitation>
<figureCitation id="00E5008722A65B2C9FDD141AD5E736CC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Bothrocophias myrringae sp. nov. in life (INSZ 0268. male). A Dorsal view of the body. B Ventral view of the body. C Dorsal view of the head. D - E Lateral views (right / left) of the head. F Ventral view of the head. Black bar length = 1 cm." figureDoi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="comparisons">
<paragraph pageId="0" pageNumber="971">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="971">
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<subSubSection id="5E612180D3B5216561D9EDEC4F7695EC" pageId="0" pageNumber="971" type="comparisons">
<paragraph id="A78EB58BC53B98435F0DEDDCA85FB8A3" pageId="0" pageNumber="971">Comparisons.</paragraph>
<paragraph id="426A2A2225B2F124691BB931855C846F" pageId="0" pageNumber="971">
<taxonomicName id="0609C4BF2C4BC684B2169C1A830AD492" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="25D6E9DAA844D23F200D09A63C491FF5" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="2324EB7690D0795767449775DC97C83B" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
can be distinguished from all its congeners by its creamy yellow ventral surfaces and ventral scales mottled with dark brown pigment, becoming creamy white toward the edges in contact with the paraventral scales, forming a white-cream paraventral stripe which proximally fuses with the final edges of the postocular stripe, and distally is interrupted by dark spots without a regular pattern; mesial ventral surfaces become heavily mottled with dark brown pigment toward the tail (versus homogeneously dark brown to black in
<taxonomicName lsidName="B. campbelli" pageId="0" pageNumber="971" rank="species" species="campbelli">
<emphasis italics="true" pageId="0" pageNumber="971">B. campbelli</emphasis>
<taxonomicName id="78BBCC48BEB6B870BE54229D877CCA93" lsidName="B. campbelli" pageId="0" pageNumber="971" rank="species" species="campbelli">
<emphasis id="C86441D0C768A01B1C17E4B85A7002CB" italics="true" pageId="0" pageNumber="971">B. campbelli</emphasis>
</taxonomicName>
; heavily mottled with dark brown pigment, with the pale interspaces between the ventrolateral blotches encroaching on the lateral edges of the ventral scales in
<taxonomicName lsidName="B. colombianus" pageId="0" pageNumber="971" rank="species" species="colombianus">
<emphasis italics="true" pageId="0" pageNumber="971">B. colombianus</emphasis>
<taxonomicName id="5A4E7709F81832A19AFEF0A8EDB33C53" lsidName="B. colombianus" pageId="0" pageNumber="971" rank="species" species="colombianus">
<emphasis id="D12BBF2076DAD725ADC1A4EAB66849E5" italics="true" pageId="0" pageNumber="971">B. colombianus</emphasis>
</taxonomicName>
; greyish brown medially, becoming paler laterally, with or without alternating dark brown spots in
<taxonomicName lsidName="B. hyoprora" pageId="0" pageNumber="971" rank="species" species="hyoprora">
<emphasis italics="true" pageId="0" pageNumber="971">B. hyoprora</emphasis>
<taxonomicName id="E85C09BEEA801FE9E4DEA7DF0C50E684" lsidName="B. hyoprora" pageId="0" pageNumber="971" rank="species" species="hyoprora">
<emphasis id="D3B05580EB1588F9E3C10EF83549C07B" italics="true" pageId="0" pageNumber="971">B. hyoprora</emphasis>
</taxonomicName>
; yellow mottled with pale to dark brown, darker posteriorly in
<taxonomicName lsidName="B. lojanus" pageId="0" pageNumber="971" rank="species" species="lojanus">
<emphasis italics="true" pageId="0" pageNumber="971">B. lojanus</emphasis>
<taxonomicName id="9454E9595E78629F98E89CD9BA955F16" lsidName="B. lojanus" pageId="0" pageNumber="971" rank="species" species="lojanus">
<emphasis id="8EE677D6929CF0031CBEA29B668524D3" italics="true" pageId="0" pageNumber="971">B. lojanus</emphasis>
</taxonomicName>
; pale pink to almost white in
<taxonomicName lsidName="B. myersi" pageId="0" pageNumber="971" rank="species" species="myersi">
<emphasis italics="true" pageId="0" pageNumber="971">B. myersi</emphasis>
<taxonomicName id="F648FA3ADDF7EA1FD26F404287E6121D" lsidName="B. myersi" pageId="0" pageNumber="971" rank="species" species="myersi">
<emphasis id="C27B77C2D25B71828D891248E727CC30" italics="true" pageId="0" pageNumber="971">B. myersi</emphasis>
</taxonomicName>
); ventral tail surface bright reddish or orange-reddish speckles with black spots in an irregular pattern, and the base of tail heavily marked with dark pigment (versus base of tail dark brown and distally yellow or yellow-green in
<taxonomicName lsidName="B. andianus" pageId="0" pageNumber="971" rank="species" species="andianus">
<emphasis italics="true" pageId="0" pageNumber="971">B. andianus</emphasis>
<taxonomicName id="A3418D17D00202C57497B064AF8E5762" lsidName="B. andianus" pageId="0" pageNumber="971" rank="species" species="andianus">
<emphasis id="14E4A7481711103D36539576E7594733" italics="true" pageId="0" pageNumber="971">B. andianus</emphasis>
</taxonomicName>
; bright yellow to tan with diffuse grayish or brown pigment in
<taxonomicName lsidName="B. campbelli" pageId="0" pageNumber="971" rank="species" species="campbelli">
<emphasis italics="true" pageId="0" pageNumber="971">B. campbelli</emphasis>
<taxonomicName id="949208E80D522AEA74CBD7B315835D08" lsidName="B. campbelli" pageId="0" pageNumber="971" rank="species" species="campbelli">
<emphasis id="6A72977C3FABC43F58DC27EC98E63B34" italics="true" pageId="0" pageNumber="971">B. campbelli</emphasis>
</taxonomicName>
; cream or pale yellow with a sparse peppering of brown in
<taxonomicName lsidName="B. colombianus" pageId="0" pageNumber="971" rank="species" species="colombianus">
<emphasis italics="true" pageId="0" pageNumber="971">B. colombianus</emphasis>
<taxonomicName id="A024D1D16DF2ECDC462A9857CC1F671A" lsidName="B. colombianus" pageId="0" pageNumber="971" rank="species" species="colombianus">
<emphasis id="2D65F42BE0AB3793EFA6360806B6F097" italics="true" pageId="0" pageNumber="971">B. colombianus</emphasis>
</taxonomicName>
; whitish with a moderate suffusion of grey in
<taxonomicName lsidName="B. myersi" pageId="0" pageNumber="971" rank="species" species="myersi">
<emphasis italics="true" pageId="0" pageNumber="971">B. myersi</emphasis>
<taxonomicName id="81D0ED0DCFE33DD5E5FD84B2A2EDE42F" lsidName="B. myersi" pageId="0" pageNumber="971" rank="species" species="myersi">
<emphasis id="06FB7C85E08867FCA625360DD557162D" italics="true" pageId="0" pageNumber="971">B. myersi</emphasis>
</taxonomicName>
. Comparisons of meristic and hemipenial characters with all its congeners of toadheaded pitvipers are summarized in Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Comparisons of meristic and hemipenial characters between B. myrringae sp. nov., B. tulitoi sp. nov. and all its congeners of toadheaded pitvipers." httpUri="http://table.plazi.org/id/F7FF0F5117D0326916209BBAF0303130" pageId="0" pageNumber="971" tableUuid="F7FF0F5117D0326916209BBAF0303130">4</tableCitation>
<tableCitation id="898498C9ED0FADAED1F6AE686C7E3989" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Comparisons of meristic and hemipenial characters between B. myrringae sp. nov., B. tulitoi sp. nov. and all its congeners of toadheaded pitvipers." httpUri="http://table.plazi.org/id/F7FF0F5117D0326916209BBAF0303130" pageId="0" pageNumber="971" tableUuid="F7FF0F5117D0326916209BBAF0303130">4</tableCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="971">
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<paragraph id="F5A9E1D316BEE1FA0926A4EF3E63D700" pageId="0" pageNumber="971">
<taxonomicName id="621FE4C56BD07FA8B8E6D120418CD023" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="0CE0723F834B8E0F8A007CD88A3F728B" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="335022D3776D99C464D7BE8F55426B54" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
can be distinguished from Ecuadorian, Peruvian, and Bolivian populations of toadheaded pitvipers currently classified as
<taxonomicName lsidName="B. microphthalmus" pageId="0" pageNumber="971" rank="species" species="microphthalmus">
<emphasis italics="true" pageId="0" pageNumber="971">B. microphthalmus</emphasis>
<taxonomicName id="F37448417751895B10E093743AEFCECA" class="Reptilia" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="B. microphthalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis id="50982549AC8AD5648866E557E8DB788B" italics="true" pageId="0" pageNumber="971">B. microphthalmus</emphasis>
</taxonomicName>
by ventral surface of tail with uniformly bright reddish or orange-reddish speckles with black spots without a regular pattern (versus heavily marked proximally with black or dark brown pigment, medially mottled with pale to dark brown, and distally with pale diffuse or pale mottling with interspaces in Bolivian and Peruvian populations). Comparisons of meristic and hemipenial characters with the Ecuadorian, Peruvian, and Bolivian populations of toadheaded pitvipers the complex are summarized in Table
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Meristic variation in Bothrocophias tulitoi sp. nov. and B. myrringae sp. nov. Bold numbers represent mode or mean. Numbers within parenthesis represent minimum and maximum values observed. * = meristic characters undifferenced by sex." httpUri="http://table.plazi.org/id/FB72B71146185B457EDFAA8C76B5AE4B" pageId="0" pageNumber="971" tableUuid="FB72B71146185B457EDFAA8C76B5AE4B">5</tableCitation>
<tableCitation id="9624F4155B916C9A16371C48692B510F" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Meristic variation in Bothrocophias tulitoi sp. nov. and B. myrringae sp. nov. Bold numbers represent mode or mean. Numbers within parenthesis represent minimum and maximum values observed. * = meristic characters undifferenced by sex." httpUri="http://table.plazi.org/id/FB72B71146185B457EDFAA8C76B5AE4B" pageId="0" pageNumber="971" tableUuid="FB72B71146185B457EDFAA8C76B5AE4B">5</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="description">
<paragraph pageId="0" pageNumber="971">Description of holotype.</paragraph>
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">
<subSubSection id="B573D22C33CE5686F2AF8C7944EDD6D2" pageId="0" pageNumber="971" type="description">
<paragraph id="1F7CDED9B4F24A0AAFE2159A823830A8" pageId="0" pageNumber="971">Description of holotype.</paragraph>
<paragraph id="A9E59994F11EBFABD045B1BC747A20A9" pageId="0" pageNumber="971">
<emphasis id="E767E243231C281DFF2A30A6103C1CAC" bold="true" pageId="0" pageNumber="971">
(Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hemipenial architecture. A, B, C Sulcate, lateral, and asulcate views of the hemipenis of the holotype of Bothrocophias tulitoi sp. nov. (INSZ 073) from vereda Cienaga La Valvanera, municipality of Garagoa, department of Boyaca, Colombia. Coordinates: N 5.106535941, W - 73.25888414; elevation 1894 m. asl. D, E, F Sulcate, lateral, and asulcate views of the hemipenis of the holotype of Bothrocophias myrringae sp. nov. (INSZ 0268) from vereda Mundo Nuevo, municipality of La Calera, department of Cundinamarca, Colombia. Coordinates: N 4.660602778, W - 73.88491667; elevation 2,761 m asl." figureDoi="10.3897/vz.72.e87313.figure5" httpUri="https://binary.pensoft.net/fig/763283" pageId="0" pageNumber="971">5</figureCitation>
<figureCitation id="C0BD53C3543C77CDE09E1E10EEDAEC98" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hemipenial architecture. A, B, C Sulcate, lateral, and asulcate views of the hemipenis of the holotype of Bothrocophias tulitoi sp. nov. (INSZ 073) from vereda Cienaga La Valvanera, municipality of Garagoa, department of Boyaca, Colombia. Coordinates: N 5.106535941, W - 73.25888414; elevation 1894 m. asl. D, E, F Sulcate, lateral, and asulcate views of the hemipenis of the holotype of Bothrocophias myrringae sp. nov. (INSZ 0268) from vereda Mundo Nuevo, municipality of La Calera, department of Cundinamarca, Colombia. Coordinates: N 4.660602778, W - 73.88491667; elevation 2,761 m asl." figureDoi="10.3897/vz.72.e87313.figure5" httpUri="https://binary.pensoft.net/fig/763283" pageId="0" pageNumber="971">5</figureCitation>
-
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Bothrocophias myrringae sp. nov. in life (INSZ 0268. male). A Dorsal view of the body. B Ventral view of the body. C Dorsal view of the head. D - E Lateral views (right / left) of the head. F Ventral view of the head. Black bar length = 1 cm." figureDoi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971">7</figureCitation>
<figureCitation id="8AF1C2044E6CD1AEB7DF64747710B04D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Holotype of Bothrocophias myrringae sp. nov. in life (INSZ 0268. male). A Dorsal view of the body. B Ventral view of the body. C Dorsal view of the head. D - E Lateral views (right / left) of the head. F Ventral view of the head. Black bar length = 1 cm." figureDoi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971">7</figureCitation>
, Table
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Species delimitation for Bothrocophias microphthalmus populations hypothesized as candidate species using the integrative taxonomic approach of Sites Jr et al. (2021). Bp = posterior probability values from Bayesian inference tree, UFB, UltraFast Bootstrap support obtained in the ML tree. * = Moderate diagnosable means that there was some overlap in the scutellation characters. Values for genetic distinctiveness correspond to the genetic distance to the closest clade (see Table 2)." httpUri="http://table.plazi.org/id/A6470B4A62615E431F4B5D7829B430A2" pageId="0" pageNumber="971" tableUuid="A6470B4A62615E431F4B5D7829B430A2">3</tableCitation>
<tableCitation id="1E647199EB40F24F0AB7770C6309C3D4" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Species delimitation for Bothrocophias microphthalmus populations hypothesized as candidate species using the integrative taxonomic approach of Sites Jr et al. (2021). Bp = posterior probability values from Bayesian inference tree, UFB, UltraFast Bootstrap support obtained in the ML tree. * = Moderate diagnosable means that there was some overlap in the scutellation characters. Values for genetic distinctiveness correspond to the genetic distance to the closest clade (see Table 2)." httpUri="http://table.plazi.org/id/A6470B4A62615E431F4B5D7829B430A2" pageId="0" pageNumber="971" tableUuid="A6470B4A62615E431F4B5D7829B430A2">3</tableCitation>
)
</emphasis>
Male, small (SVL = 542 mm, TL = 95 mm, ratio 17.5%), head and body strongly differentiated by nuchal constriction; head longer than wide (HW/HL 53.3%); snout prognathous and not upturned; absence of nasorostral and canthorostral scales; two internasal scales in contact; rostral visible from dorsal view and in narrow contact with internasal scales; two canthal scales in broad contact with internasals, the first preocular, and loreal scales in lateral view; three intercanthal scales; two supraocular scales wider than long (SW/SL= 66.4%); seven intersupraocular scales; 26 interrictal scales; two nasal scales: anterior nasal scale in broad contact with rostral, internasal, and first supralabial scale, and posterior nasal scale in contact with loreal, two prefoveal, and subnasal scales; a single subnasal scale; two prefoveal scales on both sides of the head; a single lacunolabial scale in broad contact with the first and second supralabial scales as well as with prefoveal and loreal scales, and in narrow contact with supralacunal scale; a single supralacunal scale in broad contact with first preocular scale and in narrow contact with second preocular and loreal scales; a single sublacunal in broad contact with second supralabial, and in narrow contact with third preocular, lacunolabial, and single posfoveal scale; a single subocular scale; a single posfoveal scale; five interoculolabial scales; two postocular scales; six supralabial scales, second supralabial slightly higher and wider than fourth to seventh supralabial scales, and notably higher and wider than first supralabial scale; eight infralabial scales, first infralabial scales separated by mental scale and in broad contact with first pair of gular scales, second and third infralabial scales in contact with first pair of gular scales; three pairs of gular scales; four preventral scales; dorsal scale rows 21-23-19; 155 ventral scales; anal scale single; 54 subcaudal scales; slender tail not prehensile.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="color of the holotype in life">
<paragraph pageId="0" pageNumber="971">Color of the holotype in life.</paragraph>
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">
<subSubSection id="CA14B1EC82B361700489D4B773047196" pageId="0" pageNumber="971" type="color of the holotype in life">
<paragraph id="1F07BE7BF6D93D883FF264DC39C4C3C6" pageId="0" pageNumber="971">Color of the holotype in life.</paragraph>
<paragraph id="77D72ED49BD6AA08CA3F868B965A0F54" pageId="0" pageNumber="971">
<emphasis id="D0C1C82105C08642A813A9729CFB03FB" bold="true" pageId="0" pageNumber="971">
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8A-B</figureCitation>
<figureCitation id="FCBE368D9851183CDF0EEF5D66F22650" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8A-B</figureCitation>
)
</emphasis>
The dorsal surface of the head has a dark to reddish-brown ground color with diffuse dark brown marks without a distinctive pattern. The lateral surface of the head is scattered pale yellow from the snout to anterior edge of the eye; a conspicuous dark-brown postocular stripe with white edges running obliquely from the posterior edge of the eye to the angle formed by the quadrate and jaw bone joint encompasses the temporal scales, the last two supralabial scales, the last infralabial scale, and the mesial scales located between the preventral and infralabial scales; conspicuous tricolored ocelli present on fourth to sixth infralabial scales with white or white-cream centers, followed by dark-brown or black edge, and an external broader edge yellow or reddish-yellow. Ventral surfaces of the head are tan with a peppering of brown and dark yellow without ocelli. Ground color of the dorsal body surfaces is pale yellow, tan to brown mottled with dark brown-reddish pigment and weak orange speckles; 34 dark-brown dorsal bands and/or opposite or juxtaposed trapezoid-shaped blotches with pale centre ornamented with or without brown spots. Ventral scales are creamish yellow with ventral scales heavily marked, with brown-reddish and dark yellow pigment towards the edges in contact with the paraventral scales and spotted without a regular pattern on the mesial surface with a conspicuous white-cream paraventral stripe which proximally fuses with the final edges of the postocular stripe, and distally is interrupted by dark spots without a regular pattern. Dorsal surfaces of the tail covered by two broad dark brown bands separated by three narrow pale bands that fuse toward the distal end of the tail; ventral tail surface bright reddish or orange-reddish, darkening heavily towards the base, speckled with black spots in an irregular.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="color of the holotype in ethanol">
<paragraph pageId="0" pageNumber="971">Color of the holotype in ethanol.</paragraph>
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">
<subSubSection id="84830FD48E89C3DA0C44A4677FE899E8" pageId="0" pageNumber="971" type="color of the holotype in ethanol">
<paragraph id="847183B2129FBC33DBDB53076D0A0AE1" pageId="0" pageNumber="971">Color of the holotype in ethanol.</paragraph>
<paragraph id="AEC54A20E8E5D9E64A1DD89918C75787" pageId="0" pageNumber="971">
<emphasis id="2FADB277228466CD72D02625C43053CB" bold="true" pageId="0" pageNumber="971">
(Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the head pholidosis among the holotypes of Bothrocophias microphthalmus, Bothrocophias tulitoi sp. nov. and Bothrocophias myrringae sp. nov. A, B, C Dorsal, lateral, and ventral view of the head of B. microphthalmus ANSP 11515. D, E, F Dorsal, lateral, and ventral view of the head of B. tulitoi sp. nov. INSZ 073. G, H, I Dorsal, lateral, and ventral view of the head of B. myrringae sp. nov. Scales: dark green = second / third supralabial scale. yellow = lacunolabial / prelacunal. Blue = internasal scales. Fuchsia = scale separating internasal scales." figureDoi="10.3897/vz.72.e87313.figure6" httpUri="https://binary.pensoft.net/fig/763284" pageId="0" pageNumber="971">6</figureCitation>
<figureCitation id="90F531FE4733BC73D7AF48707D36A1C1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Comparison of the head pholidosis among the holotypes of Bothrocophias microphthalmus, Bothrocophias tulitoi sp. nov. and Bothrocophias myrringae sp. nov. A, B, C Dorsal, lateral, and ventral view of the head of B. microphthalmus ANSP 11515. D, E, F Dorsal, lateral, and ventral view of the head of B. tulitoi sp. nov. INSZ 073. G, H, I Dorsal, lateral, and ventral view of the head of B. myrringae sp. nov. Scales: dark green = second / third supralabial scale. yellow = lacunolabial / prelacunal. Blue = internasal scales. Fuchsia = scale separating internasal scales." figureDoi="10.3897/vz.72.e87313.figure6" httpUri="https://binary.pensoft.net/fig/763284" pageId="0" pageNumber="971">6</figureCitation>
)
</emphasis>
After six months in ethanol, dark brown and brown coloration was maintained, and surfaces with brown-reddish, reddish-yellow, and yellow coloration became pale grey or creamy white.
</paragraph>
<caption doi="10.3897/vz.72.e87313.figure6" httpUri="https://binary.pensoft.net/fig/763284" pageId="0" pageNumber="971" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">Figure 6.</emphasis>
<caption id="64B2B46F918E63E0A195BDE35237FBAD" doi="10.3897/vz.72.e87313.figure6" httpUri="https://binary.pensoft.net/fig/763284" pageId="0" pageNumber="971" start="Figure 6" startId="F6">
<paragraph id="E37D736D673E080A7346E99AFF4F2375" pageId="0" pageNumber="971">
<emphasis id="8EAD31343B93BB8BE439344C70E46C79" bold="true" pageId="0" pageNumber="971">Figure 6.</emphasis>
Comparison of the head pholidosis among the holotypes of
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1875" class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias microphthalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias microphthalmus</emphasis>
<taxonomicName id="FE8ACE2554D03020DE3CE00F2858AE7E" baseAuthorityName="Cope" baseAuthorityYear="1875" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias microphthalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis id="01C1798A43279D41CC05532EC0D623A8" italics="true" pageId="0" pageNumber="971">Bothrocophias microphthalmus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Angarita-Sierra &amp; Cubides-Cubillos &amp; Hurtado-Gómez" authorityYear="2022" class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias tulitoi" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="tulitoi">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias tulitoi</emphasis>
<taxonomicName id="A9440732DDBE8ABC0817FCD10312BCFB" authorityName="Angarita-Sierra &amp; Cubides-Cubillos &amp; Hurtado-Gómez" authorityYear="2022" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias tulitoi" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="tulitoi">
<emphasis id="54FE400D317BB4302CFDB8A307A4D2E5" italics="true" pageId="0" pageNumber="971">Bothrocophias tulitoi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="5D8A1BD0BD739FE2DB6C26DEFFB9D802" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
and
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<taxonomicName id="BE7EAEE510B383370B7A51BD4377CC33" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="65CCF810855A37D0713CFE75DBEE570E" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov. A</emphasis>
<emphasis id="3A04604B0286BBF5C795F0482B9F8C9A" bold="true" pageId="0" pageNumber="971">sp. nov. A</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="971">B</emphasis>
<emphasis id="56F7B3EEA2C61270EC40185DD34DD30A" bold="true" pageId="0" pageNumber="971">B</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="971">C</emphasis>
<emphasis id="CC667D5B6F43A2D85EA3BCF08D681F35" bold="true" pageId="0" pageNumber="971">C</emphasis>
Dorsal, lateral, and ventral view of the head of
<taxonomicName lsidName="B. microphthalmus" pageId="0" pageNumber="971" rank="species" species="microphthalmus">
<emphasis italics="true" pageId="0" pageNumber="971">B. microphthalmus</emphasis>
<taxonomicName id="95E6A264EB52A0679FF98BD21ADFDADC" class="Reptilia" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="B. microphthalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis id="914834A4D3819B05CFA624F9E63C0195" italics="true" pageId="0" pageNumber="971">B. microphthalmus</emphasis>
</taxonomicName>
ANSP 11515.
<emphasis bold="true" pageId="0" pageNumber="971">D</emphasis>
<emphasis id="6CB358B469663FD49358B3E7848904B5" bold="true" pageId="0" pageNumber="971">D</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="971">E</emphasis>
<emphasis id="CAD33DBDD9A097FAD0A827680F2F6704" bold="true" pageId="0" pageNumber="971">E</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="971">F</emphasis>
<emphasis id="2592E899A88F4AB8D811851F396AD288" bold="true" pageId="0" pageNumber="971">F</emphasis>
Dorsal, lateral, and ventral view of the head of
<taxonomicName lsidName="B. tulitoi" pageId="0" pageNumber="971" rank="species" species="tulitoi">
<emphasis italics="true" pageId="0" pageNumber="971">B. tulitoi</emphasis>
<taxonomicName id="1D001A5782539897ADB7D54999E022BD" class="Reptilia" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="B. tulitoi" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="tulitoi">
<emphasis id="9EE28C4B8099E01E06947AE30FD00864" italics="true" pageId="0" pageNumber="971">B. tulitoi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="C0932A17D920B89FDA3B61F5A6234D39" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
INSZ 073.
<emphasis bold="true" pageId="0" pageNumber="971">G</emphasis>
<emphasis id="BCF6F93FE821B966E4027CD95A1328E7" bold="true" pageId="0" pageNumber="971">G</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="971">H</emphasis>
<emphasis id="2F4914A96C2EDCBE607E3A2A587A482F" bold="true" pageId="0" pageNumber="971">H</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="971">I</emphasis>
<emphasis id="C65088777EC517306761EE94C14D3C8C" bold="true" pageId="0" pageNumber="971">I</emphasis>
Dorsal, lateral, and ventral view of the head of
<taxonomicName lsidName="B. myrringae" pageId="0" pageNumber="971" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">B. myrringae</emphasis>
<taxonomicName id="F98DE0CDCA1F246AC0C2FB1C1C3D70CE" class="Reptilia" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="B. myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="CFD9F6F5AA51C1A495B0160915A48DCC" italics="true" pageId="0" pageNumber="971">B. myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="BF198724E9D7F1FFD3C2036049F77AD1" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
Scales: dark green = second/third supralabial scale. yellow = lacunolabial/prelacunal. Blue = internasal scales. Fuchsia = scale separating internasal scales.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="color pattern variation">
<paragraph pageId="0" pageNumber="971">Color pattern variation.</paragraph>
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">
<subSubSection id="3D01AABA1B78059A1DF6C0AEFD3DDDCC" pageId="0" pageNumber="971" type="color pattern variation">
<paragraph id="F3BCBD870CB0992150C636550EF59690" pageId="0" pageNumber="971">Color pattern variation.</paragraph>
<paragraph id="7A28EE8CD30F745A65021AC1A3478E08" pageId="0" pageNumber="971">
<emphasis id="456E1B3EF769266698041D505C538885" bold="true" pageId="0" pageNumber="971">
(Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8C-D</figureCitation>
<figureCitation id="CAE33EBFCA8FD374B0468E8BEB408F53" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8C-D</figureCitation>
)
</emphasis>
Adults of
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<taxonomicName id="015FD48E20597E6E7B236DA8D15D4A15" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="FA3BDCD632BDAD65B26AF035498D5896" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="B785616FDB3CFC2ED5F520BF1F14A65F" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
exhibit sexual dimorphism. The dorsal, lateral, and ventral surfaces of the head and body of males are melanized with conspicuous tricolored ocelli on the third to seventh infralabial scales, and the lateral and ventral surfaces of the head in females exhibit a homogenous creamish yellow to pale yellow coloration without ocelli in the labials and gular scales (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8C-D</figureCitation>
<figureCitation id="5CDB86A1E1B3DF4972D16168C6303095" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Color in life of Bothrocophias myrringae sp. nov., and sexual dimorphism. A, B Lateral and dorsal view of the holotype INSZ 0268. C Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR 009). D Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de Coasavista, municipality of Fomeque, Cundinamarca, coordinates N 4.495001, W - 73.852056. Pictures by Ronald A. Diaz-Flores." figureDoi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971">8C-D</figureCitation>
). In addition, eye color in males is usually homogeneously mottled with dark brown pigment and weak orange speckles around the pupil; in females, it is orange-gold with a peppering of brown. One female (IAvH-R7045: San Juanito, Meta) has 25 dorsal bands, which was an unusual count among our samples.
</paragraph>
<caption doi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">Figure 7.</emphasis>
<caption id="524FB2F61EFD0B12D65EF5B4E16C8B55" doi="10.3897/vz.72.e87313.figure7" httpUri="https://binary.pensoft.net/fig/763285" pageId="0" pageNumber="971" start="Figure 7" startId="F7">
<paragraph id="B95204F3DB8A73602DEDF903C25AB23A" pageId="0" pageNumber="971">
<emphasis id="47CDEE7B70830AE613DCC2DBF7BB89A1" bold="true" pageId="0" pageNumber="971">Figure 7.</emphasis>
Holotype of
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<taxonomicName id="2D19AF58BFED5003B71A1C3CB998604D" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="C2E3AF5A5EF144B3D5E72D82A18BEE4D" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="0731628181DC0CE88FFA553C4FA865B6" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
in life (INSZ 0268. male).
<emphasis bold="true" pageId="0" pageNumber="971">A</emphasis>
<emphasis id="63076B0F8747F2C71A627C0E9F86B866" bold="true" pageId="0" pageNumber="971">A</emphasis>
Dorsal view of the body.
<emphasis bold="true" pageId="0" pageNumber="971">B</emphasis>
<emphasis id="20B6E9917AB50846D6017DFD561D1692" bold="true" pageId="0" pageNumber="971">B</emphasis>
Ventral view of the body.
<emphasis bold="true" pageId="0" pageNumber="971">C</emphasis>
<emphasis id="6245CE6AD0C5EAD760C0247A3324A2D2" bold="true" pageId="0" pageNumber="971">C</emphasis>
Dorsal view of the head.
<emphasis bold="true" pageId="0" pageNumber="971">D</emphasis>
<emphasis id="A387EF3C0E0D1DD0AE0A32671D347FD6" bold="true" pageId="0" pageNumber="971">D</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="971">E</emphasis>
<emphasis id="862677C3CB82F0E5BA636DD55B0C07F6" bold="true" pageId="0" pageNumber="971">E</emphasis>
Lateral views (right/left) of the head.
<emphasis bold="true" pageId="0" pageNumber="971">F</emphasis>
<emphasis id="19DFDC8C42453FF87AAA9B82EA360841" bold="true" pageId="0" pageNumber="971">F</emphasis>
Ventral view of the head. Black bar length = 1cm.
</paragraph>
</caption>
<caption doi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">Figure 8.</emphasis>
<caption id="713C0F3A71D7E137DF33AD2D74527219" doi="10.3897/vz.72.e87313.figure8" httpUri="https://binary.pensoft.net/fig/763286" pageId="0" pageNumber="971" start="Figure 8" startId="F8">
<paragraph id="F6F8213D605CAEE16C336E756887D36A" pageId="0" pageNumber="971">
<emphasis id="E220ECF87C5E7352112AB53C5D308F93" bold="true" pageId="0" pageNumber="971">Figure 8.</emphasis>
Color in life of
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<taxonomicName id="C75371351C36CB6DF4CF2D079CD0FCFE" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="88D36141853726F6714315547DC302C2" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="C870BC075345F5C0F691EB2A6E28B003" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
, and sexual dimorphism.
<emphasis bold="true" pageId="0" pageNumber="971">A</emphasis>
<emphasis id="BC93F9978163C6C1DC40B9FB3DFA5B1B" bold="true" pageId="0" pageNumber="971">A</emphasis>
,
<emphasis bold="true" pageId="0" pageNumber="971">B</emphasis>
<emphasis id="1493AA5FE352A68AACFD6D9E881A08D2" bold="true" pageId="0" pageNumber="971">B</emphasis>
Lateral and dorsal view of the holotype INSZ 0268.
<emphasis bold="true" pageId="0" pageNumber="971">C</emphasis>
<emphasis id="687E8FBD61F52772452200774E8206A0" bold="true" pageId="0" pageNumber="971">C</emphasis>
Male exhibiting melanic coloration on dorsal body and head surfaces, as well as conspicuous tricolored ocelli on the infralabial scales (paratype INSV-SR009).
<emphasis bold="true" pageId="0" pageNumber="971">D</emphasis>
<emphasis id="D17C508AF674692E9F93436C0A23FCD9" bold="true" pageId="0" pageNumber="971">D</emphasis>
Female exhibiting creamish-yellow to tan coloration on the dorsal body and head surfaces, without tricolored ocelli on the infralabial scales. Both C and D specimens are from vereda de
<normalizedToken originalValue="Coasavistá">Coasavista</normalizedToken>
<normalizedToken id="DDA508B259E99BFD697D36F9975E272D" originalValue="Coasavistá">Coasavista</normalizedToken>
, municipality of
<normalizedToken originalValue="Fómeque">Fomeque</normalizedToken>
<normalizedToken id="425C8743C0F0B57CAA4DD9DD2122C33E" originalValue="Fómeque">Fomeque</normalizedToken>
, Cundinamarca, coordinates
<geoCoordinate degrees="4.495001" direction="north" orientation="latitude" precision="1" value="4.495001">N 4.495001</geoCoordinate>
<geoCoordinate id="75FC95C6471606B54CF2867EB6805886" degrees="4.495001" direction="north" orientation="latitude" precision="1" value="4.495001">N 4.495001</geoCoordinate>
, W -73.852056. Pictures by Ronald A.
<normalizedToken originalValue="Díaz-Flores">Diaz-Flores</normalizedToken>
<normalizedToken id="EBB3B43F4EB8BD4890B1A017401C1CD9" originalValue="Díaz-Flores">Diaz-Flores</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="meristic variation">
<paragraph pageId="0" pageNumber="971">Meristic variation.</paragraph>
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">
<subSubSection id="AF056C06D927F394E6AA376B8F5F70CB" pageId="0" pageNumber="971" type="meristic variation">
<paragraph id="A3D67758E594290426274898459CFCDF" pageId="0" pageNumber="971">Meristic variation.</paragraph>
<paragraph id="4FB4C6714B7770286DB4F07CCE4DA85D" pageId="0" pageNumber="971">
<emphasis id="89E4E0636DDC7445DBB286B275D03FA3" bold="true" pageId="0" pageNumber="971">
(Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Comparisons of meristic and hemipenial characters between B. myrringae sp. nov., B. tulitoi sp. nov. and all its congeners of toadheaded pitvipers." httpUri="http://table.plazi.org/id/F7FF0F5117D0326916209BBAF0303130" pageId="0" pageNumber="971" tableUuid="F7FF0F5117D0326916209BBAF0303130">4</tableCitation>
<tableCitation id="4E993744479F84CCB5E951AB4E71F6A1" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Comparisons of meristic and hemipenial characters between B. myrringae sp. nov., B. tulitoi sp. nov. and all its congeners of toadheaded pitvipers." httpUri="http://table.plazi.org/id/F7FF0F5117D0326916209BBAF0303130" pageId="0" pageNumber="971" tableUuid="F7FF0F5117D0326916209BBAF0303130">4</tableCitation>
)
</emphasis>
Female and male adults of
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<taxonomicName id="A7DFCB05F3103E60E3FC63010A9B8841" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="24957B5C18DFC496367B5808C8B099FC" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="6FEFFDCB4AC502EE52926AA40F33C7CC" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
exhibit sexual dimorphism in ventral scale counts (Table
<tableCitation captionStart="Table 4" captionStartId="T4" captionText="Table 4. Comparisons of meristic and hemipenial characters between B. myrringae sp. nov., B. tulitoi sp. nov. and all its congeners of toadheaded pitvipers." httpUri="http://table.plazi.org/id/F7FF0F5117D0326916209BBAF0303130" pageId="0" pageNumber="971" tableUuid="F7FF0F5117D0326916209BBAF0303130">4</tableCitation>
<tableCitation id="6EDEEFE3DC71440D9D45893070E4022F" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Comparisons of meristic and hemipenial characters between B. myrringae sp. nov., B. tulitoi sp. nov. and all its congeners of toadheaded pitvipers." httpUri="http://table.plazi.org/id/F7FF0F5117D0326916209BBAF0303130" pageId="0" pageNumber="971" tableUuid="F7FF0F5117D0326916209BBAF0303130">4</tableCitation>
). One female (IAvH-R6877: Guayabetal, Cundinamarca) has a single canthorostral scale on the right side of the head. One male (INSVSR-00099: El Calvario, Meta) lacks a lacunolabial scale on both sides of the head, and this is the only specimen showing this condition.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="hemipenial morphology">
<paragraph pageId="0" pageNumber="971">Hemipenial morphology.</paragraph>
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">
<subSubSection id="67063881C9A46C1B4A94FF7D3134EAC0" pageId="0" pageNumber="971" type="hemipenial morphology">
<paragraph id="02B634425BF138BCACD074678A8E0D2D" pageId="0" pageNumber="971">Hemipenial morphology.</paragraph>
<paragraph id="EBA8F7FB0FD8E26AC0FAE5D77B4186E5" pageId="0" pageNumber="971">
<emphasis id="D1C1269DA8A08AA3E3691B170C5F3A1F" bold="true" pageId="0" pageNumber="971">
(
<emphasis italics="true" pageId="0" pageNumber="971">n</emphasis>
<emphasis id="638850AE653D45F2C4418AD30F361E03" italics="true" pageId="0" pageNumber="971">n</emphasis>
=2, Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hemipenial architecture. A, B, C Sulcate, lateral, and asulcate views of the hemipenis of the holotype of Bothrocophias tulitoi sp. nov. (INSZ 073) from vereda Cienaga La Valvanera, municipality of Garagoa, department of Boyaca, Colombia. Coordinates: N 5.106535941, W - 73.25888414; elevation 1894 m. asl. D, E, F Sulcate, lateral, and asulcate views of the hemipenis of the holotype of Bothrocophias myrringae sp. nov. (INSZ 0268) from vereda Mundo Nuevo, municipality of La Calera, department of Cundinamarca, Colombia. Coordinates: N 4.660602778, W - 73.88491667; elevation 2,761 m asl." figureDoi="10.3897/vz.72.e87313.figure5" httpUri="https://binary.pensoft.net/fig/763283" pageId="0" pageNumber="971">5D-F</figureCitation>
<figureCitation id="F5A3255330EF64CC9C3A5151F42944AC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Hemipenial architecture. A, B, C Sulcate, lateral, and asulcate views of the hemipenis of the holotype of Bothrocophias tulitoi sp. nov. (INSZ 073) from vereda Cienaga La Valvanera, municipality of Garagoa, department of Boyaca, Colombia. Coordinates: N 5.106535941, W - 73.25888414; elevation 1894 m. asl. D, E, F Sulcate, lateral, and asulcate views of the hemipenis of the holotype of Bothrocophias myrringae sp. nov. (INSZ 0268) from vereda Mundo Nuevo, municipality of La Calera, department of Cundinamarca, Colombia. Coordinates: N 4.660602778, W - 73.88491667; elevation 2,761 m asl." figureDoi="10.3897/vz.72.e87313.figure5" httpUri="https://binary.pensoft.net/fig/763283" pageId="0" pageNumber="971">5D-F</figureCitation>
)
</emphasis>
Hemipenes
<emphasis italics="true" pageId="0" pageNumber="971">in situ</emphasis>
<emphasis id="2EE810069C6E437A7E521AA333E1E8B9" italics="true" pageId="0" pageNumber="971">in situ</emphasis>
extend to the level of the 8th or 12th subcaudal scale, bilobation point ranges between the 2rd and 4th subcaudal scale. Everted and inflated, the organ is deeply bilobed, bicalyculate and moderately capitate distally; hemipenial lobes slim and cylindrical; in sulcate view, lobe crotch ornamented with scattered spinules;
<emphasis italics="true" pageId="0" pageNumber="971">sulcus spermaticus</emphasis>
<emphasis id="6ED6596B66D8F68FC774D3E5222575B3" italics="true" pageId="0" pageNumber="971">sulcus spermaticus</emphasis>
cetrolineal, bifurcate and with walls weakly defined, bifurcation occurs below bilobation point and proximal to the midpoint of the hemipenial body;
<emphasis italics="true" pageId="0" pageNumber="971">sulcus spermaticus</emphasis>
<emphasis id="CC9184E95FB09E9F19348DCEE7D00772" italics="true" pageId="0" pageNumber="971">sulcus spermaticus</emphasis>
branch runs to lobe tips; intrasulcar area barely covered with spines that increase in size distally; second half of each hemipenal lobe scarcely ornamented by large calyces with weakly defined edges;
<emphasis italics="true" pageId="0" pageNumber="971">sulcus spermaticus</emphasis>
<emphasis id="7C9D6FF3759D0B2E011B3A1DCFB04E49" italics="true" pageId="0" pageNumber="971">sulcus spermaticus</emphasis>
walls weakly defined. In sulcate view, hemipenial body barely covered with small spines proximally; hemipenial body ornamented medially by few large and strongly calcified mesial spines arranged in oblique rows, with lateral and mesial spines of the hemipenial body variable in size; the spines in each lobe are replaced distally by large calyces with weakly spinulate edges. In lateral view, hemipenial body barely covered with small spines proximally; lobes ornamented medially by scattered, large, and strongly calcified lateral spines that increase in size distally; and the large spines replaced distally by large, weakly defined calyces. In asulcate view, the hemipenial body is barely covered with small spines proximally; hemipenial body ornamented medially by scattered, large, and strongly calcified lateral spines arranged in oblique rows which increase in size distally; and the spines in each lobe replaced by large and poorly developed calyces through an oblique cut.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="etymology">
<paragraph pageId="0" pageNumber="971">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="971">
<subSubSection id="240AAB2DD7C69A186A39D4AC08471EAE" pageId="0" pageNumber="971" type="etymology">
<paragraph id="1FBFA0D029ECEDF23ADC18377AB63490" pageId="0" pageNumber="971">Etymology.</paragraph>
<paragraph id="0F165F01EDA92B020CB816165D56DBA6" pageId="0" pageNumber="971">
The specific epithet
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<taxonomicName id="7B737D48A824E448D54236441C171E39" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="010547FD6D6CE7F0C1DFE788B6B83F5E" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
is the Latin translation of the Spanish nickname
<normalizedToken originalValue="“Mirringa,”">&quot;Mirringa,&quot;</normalizedToken>
<normalizedToken id="3EAFBCEF49F8A0B89E35076AB4E175DE" originalValue="“Mirringa,”">&quot;Mirringa,&quot;</normalizedToken>
which means
<normalizedToken originalValue="“pinch”">&quot;pinch&quot;</normalizedToken>
<normalizedToken id="BDBB3DEDE9CAB728558F4B08F0C98AF4" originalValue="“pinch”">&quot;pinch&quot;</normalizedToken>
or something very small. The word
<normalizedToken originalValue="“Mirringa”">&quot;Mirringa&quot;</normalizedToken>
<normalizedToken id="03237CE0C4A9E51507C345FB55213BEB" originalValue="“Mirringa”">&quot;Mirringa&quot;</normalizedToken>
was popularized by Rafael Pombo (1833-1912), a Colombian poet and writer who wrote a popular fable titled &quot;Mirringa Mirronga.&quot; Given the popularity of the fable, as well as the homophonic similarity of
<normalizedToken originalValue="“Mirringa”">&quot;Mirringa&quot;</normalizedToken>
<normalizedToken id="EFB9393DF66443F2209BB2FB60091CA8" originalValue="“Mirringa”">&quot;Mirringa&quot;</normalizedToken>
and the name
<normalizedToken originalValue="“Myriam,”">&quot;Myriam,&quot;</normalizedToken>
<normalizedToken id="57ED517FF75F7496852B3F4D9F0CB7A9" originalValue="“Myriam,”">&quot;Myriam,&quot;</normalizedToken>
the nickname
<normalizedToken originalValue="“Myrringa”">&quot;Myrringa&quot;</normalizedToken>
<normalizedToken id="E5A63E16B2FBA3E740C3963016D42B95" originalValue="“Myrringa”">&quot;Myrringa&quot;</normalizedToken>
began to be used as a term of endearment. The name of the new species is in honor of the educator Myriam Sierra Guerrero (mother of the first author). She was the philosophical and conceptual advisor of professor Tulio Manuel Angarita Serrano and contributed to the development of the modern Colombian education model that all schools within Colombia currently employ. Professor Sierra Guerrero also helped develop the theoretical framework for the implementation of institutional educational projects in Colombian public and private schools (see Angarita-Serrano 1990; Angarita-Serrano 1994; Angarita-Serrano and Chaves 1995; Angarita-Serran 1996; Angarita-Serrano 2000).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="distribution">
<paragraph pageId="0" pageNumber="971">Distribution and natural history.</paragraph>
<paragraph pageId="0" pageNumber="971">
<subSubSection id="92881E770E5B084BE4FE798E915B572F" pageId="0" pageNumber="971" type="distribution">
<paragraph id="EF5E152FF7F96588EFE5616FDCA459F9" pageId="0" pageNumber="971">Distribution and natural history.</paragraph>
<paragraph id="9F9043EF8B2F4B6E95B605B23F42E1EC" pageId="0" pageNumber="971">
The known localities of
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<taxonomicName id="3BB0CFBF64D15DDA5CB51E2C16404BC3" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="6DD008C3D4E7040CE18D7C9287E51851" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="2348F686036E3B081DBB9115C74C5B52" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
are from 1754 to 2761 m a.s.l. in both the central mountains and eastern slopes of the Cordillera Oriental of Colombia in the municipalities of La Calera,
<normalizedToken originalValue="Choachí">Choachi</normalizedToken>
<normalizedToken id="F20D1F044046E8A1EA31DC8FEE81E4D9" originalValue="Choachí">Choachi</normalizedToken>
,
<normalizedToken originalValue="Fómeque">Fomeque</normalizedToken>
<normalizedToken id="80AD1027E8586EC6F5AC2C512EA6A5ED" originalValue="Fómeque">Fomeque</normalizedToken>
, and Guayabetal (Cundinamarca), and El Calvario and San Juanito (Meta, Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Geographic distribution of Bothrocophias microphtalmus sensu lato, B. tulitoi sp. nov., and B. myrringae sp. nov., including type localities. The background map was obtained from the Esri open database through the following sources: DeLorme. USDS. NPS; USGS. NOAA." figureDoi="10.3897/vz.72.e87313.figure9" httpUri="https://binary.pensoft.net/fig/763287" pageId="0" pageNumber="971">9</figureCitation>
<figureCitation id="07782EB0908576D493B7E53373990BC2" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Geographic distribution of Bothrocophias microphtalmus sensu lato, B. tulitoi sp. nov., and B. myrringae sp. nov., including type localities. The background map was obtained from the Esri open database through the following sources: DeLorme. USDS. NPS; USGS. NOAA." figureDoi="10.3897/vz.72.e87313.figure9" httpUri="https://binary.pensoft.net/fig/763287" pageId="0" pageNumber="971">9</figureCitation>
).
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
<taxonomicName id="0E430C5F3E56909D03F5C6E2A238F003" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="94D94E2A026D3E9D15FF2C654847ABF9" italics="true" pageId="0" pageNumber="971">Bothrocophias myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="3C80199EF2F4D63D16643624DBCBC12D" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
appears to be associated with cloud montane, high Andean Forest, and subparamos but is also tolerant of disturbed or transformed habitats such as livestock pastures and agricultural fields. The new species was found in sympatry with
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Viperidae" genus="Bothrops" kingdom="Animalia" lsidName="Bothrops atrox" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="atrox">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrops atrox</emphasis>
<taxonomicName id="6468197710CFF94635D49CFB5A37CC04" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Viperidae" genus="Bothrops" kingdom="Animalia" lsidName="Bothrops atrox" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="atrox">
<emphasis id="DDC57181587B45637985F202C6A8671D" italics="true" pageId="0" pageNumber="971">Bothrops atrox</emphasis>
</taxonomicName>
in localities from
<normalizedToken originalValue="Fómeque">Fomeque</normalizedToken>
<normalizedToken id="E613DFD573A32FAEFF9B5A821F9409D8" originalValue="Fómeque">Fomeque</normalizedToken>
. No natural history data are available.
</paragraph>
<caption doi="10.3897/vz.72.e87313.figure9" httpUri="https://binary.pensoft.net/fig/763287" pageId="0" pageNumber="971" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">Figure 9.</emphasis>
<caption id="EBEEF9EA3B0F0C3147BF33667DA2F4CA" doi="10.3897/vz.72.e87313.figure9" httpUri="https://binary.pensoft.net/fig/763287" pageId="0" pageNumber="971" start="Figure 9" startId="F9">
<paragraph id="8A7D4DD9AF3FBCAC9071F0666AA38B5E" pageId="0" pageNumber="971">
<emphasis id="7FFCA21AEC5C3A7794DB2D84C6B666F2" bold="true" pageId="0" pageNumber="971">Figure 9.</emphasis>
Geographic distribution of
<taxonomicName class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias microphtalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphtalmus">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias microphtalmus</emphasis>
<taxonomicName id="BB3E609EF94F48D0726CBCBA6CA8805C" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias microphtalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphtalmus">
<emphasis id="D99AB2AA78E94177C552F87F6FD6457B" italics="true" pageId="0" pageNumber="971">Bothrocophias microphtalmus</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="0" pageNumber="971">sensu lato</emphasis>
<emphasis id="8462B4C78CD5FCBA08AF9660E5E5793D" italics="true" pageId="0" pageNumber="971">sensu lato</emphasis>
,
<taxonomicName lsidName="B. tulitoi" pageId="0" pageNumber="971" rank="species" species="tulitoi">
<emphasis italics="true" pageId="0" pageNumber="971">B. tulitoi</emphasis>
<taxonomicName id="D4163C6C6EC84E3F3869346B2325C66F" class="Reptilia" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="B. tulitoi" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="tulitoi">
<emphasis id="86BD58A71FFC27AB172181663EA7DA81" italics="true" pageId="0" pageNumber="971">B. tulitoi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="36A20F10F5A43651E66093D8608D9464" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
, and
<taxonomicName lsidName="B. myrringae" pageId="0" pageNumber="971" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">B. myrringae</emphasis>
<taxonomicName id="2D7DEE1C91D26565FC64AC0FB34746A4" class="Reptilia" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="B. myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="F1F44860C02EBAF02F4F3B79C35290C1" italics="true" pageId="0" pageNumber="971">B. myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="35726B18F36DC8AB7090F485EA41014C" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
, including type localities. The background map was obtained from the Esri open database through the following sources: DeLorme. USDS. NPS; USGS. NOAA.
</paragraph>
</caption>
<caption doi="10.3897/vz.72.e87313.figure10" httpUri="https://binary.pensoft.net/fig/763288" pageId="0" pageNumber="971" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="971">
<emphasis bold="true" pageId="0" pageNumber="971">Figure 10.</emphasis>
<caption id="38FCEB64C85A6C95850AC427C4E25431" doi="10.3897/vz.72.e87313.figure10" httpUri="https://binary.pensoft.net/fig/763288" pageId="0" pageNumber="971" start="Figure 10" startId="F10">
<paragraph id="43DFFD68EF46ADB0582F2851E5026FAB" pageId="0" pageNumber="971">
<emphasis id="126D19D839D2ECE428B72DA5179443C8" bold="true" pageId="0" pageNumber="971">Figure 10.</emphasis>
Elevational segregation in the
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1875" class="Aves" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrocophias microphthalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis italics="true" pageId="0" pageNumber="971">Bothrocophias microphthalmus</emphasis>
<taxonomicName id="AB0018FD9A9E3E7976AD4F4E43737FD0" baseAuthorityName="Cope" baseAuthorityYear="1875" class="Reptilia" family="Viperidae" genus="Bothrocophias" kingdom="Animalia" lsidName="Bothrocophias microphthalmus" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis id="7615D876FF391485253C3E1B29423D96" italics="true" pageId="0" pageNumber="971">Bothrocophias microphthalmus</emphasis>
</taxonomicName>
species complex across its entire distribution.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="971" type="envenomation">
<paragraph pageId="0" pageNumber="971">Envenomation.</paragraph>
<paragraph pageId="0" pageNumber="971">
<subSubSection id="CFB6D491BDB88F729B96A642C9EBDBC0" pageId="0" pageNumber="971" type="envenomation">
<paragraph id="017BA59EF209E2B39EF4DCA1D3155FA0" pageId="0" pageNumber="971">Envenomation.</paragraph>
<paragraph id="2DBDC30DB52FE8FBB109A64B79461F59" pageId="0" pageNumber="971">
A total of three snakebite events in the last decade might have potentially been caused by
<taxonomicName lsidName="B. myrringae" pageId="0" pageNumber="971" rank="species" species="myrringae">
<emphasis italics="true" pageId="0" pageNumber="971">B. myrringae</emphasis>
<taxonomicName id="39D7CDA4311F4342C183F90C7ADBBD1B" class="Reptilia" family="Viperidae" genus="Bothrocophias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="B. myrringae" order="Squamata" pageId="0" pageNumber="971" phylum="Chordata" rank="species" species="myrringae">
<emphasis id="5A916FAA86FDCE900D8732132EAFD530" italics="true" pageId="0" pageNumber="971">B. myrringae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
<emphasis id="BD85AEE0A403609DF3E008FD752A418A" bold="true" pageId="0" pageNumber="971">sp. nov.</emphasis>
, all of which were from the municipality of El Calvario (Meta). Each case was categorized as mild, moderate, and severe, respectively, and one fatality was reported. Local symptoms reported included oedema (100%), pain (100%), erythema (66%), and phlyctens (33%); systemic symptoms included respiratory failure (33%) and muscular weakness (33%). Symptoms such as paresthesia ecchymosis, bruising, sickness, vomit, vertigo, gingivorrhagia, hematuria, and altered vision were not reported.
</paragraph>
</subSubSection>