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<document id="4E72D91E8B8B7EBA2DFA41F9B18B2D63" ID-DOI="10.3897/zookeys.1228.143007" ID-ZooBank="3A5C895C-ACF6-4E44-BD53-6296E1F9EAB1" ID-publisher-id="143007" URI-arpha="C30A29E7-DD94-5DA1-B8D1-F45BB72750DE" URI-zoobank="http://zoobank.org/3A5C895C-ACF6-4E44-BD53-6296E1F9EAB1" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1740162329141" checkinUser="pensoft" docAuthor="Popovici, George &amp; Edgecombe, Gregory D." docDate="2025" docId="18A70C29CAA05FD888E215C1A621B2A3" docLanguage="en" docName="ZooKeys 1228: 225-273" docOrigin="ZooKeys 1228" docSource="https://zookeys.pensoft.net/article/143007/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Lamyctes tristani" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="273" masterDocId="C30A29E7DD945DA1B8D1F45BB72750DE" masterDocTitle="Centipedes (Myriapoda, Chilopoda) of Aldabra Atoll (Seychelles)" masterLastPageNumber="273" masterPageNumber="225" pageNumber="225" updateTime="1740173442479" updateUser="admin">
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<mods:title id="5F20426189C67F044885CF2BA10ED9E6">Centipedes (Myriapoda, Chilopoda) of Aldabra Atoll (Seychelles)</mods:title>
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<mods:namePart id="DBB82DBBBA49030CBF237CEA56BB1447">Popovici, George</mods:namePart>
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<mods:affiliation id="06E176F86B1AFBF1F7977A5AFD20CE7A">Department of Life Sciences, Imperial College London, London SW 7 2 AZ, UK &amp; &quot; Grigore Antipa &quot; National Museum of Natural History, Pavel Dimitrievici Kiseleff St. 1, 011341, Bucharest, Romania</mods:affiliation>
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<mods:namePart id="2052C4B23FC6E626F886C626ECC7193D">Edgecombe, Gregory D.</mods:namePart>
<mods:nameIdentifier id="3D4778C462098A979B4866BAE2B659C6" type="ORCID">0000-0002-9591-8011</mods:nameIdentifier>
<mods:affiliation id="4EF490C64F3FD9971248919EBB7F1E0D">The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<treatment id="18A70C29CAA05FD888E215C1A621B2A3" ID-arpha="18A70C29-CAA0-5FD8-88E2-15C1A621B2A3" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3" httpUri="http://treatment.plazi.org/id/18A70C29CAA05FD888E215C1A621B2A3" scope_class="Chilopoda" scope_family="Henicopidae" scope_order="Lithobiomorpha">
<subSubSection id="7C94D0A5545870D7CDFB3A27EED66FB4" type="nomenclature">
<paragraph id="95196D75531D89EDACE821127475BEE5">
<taxonomicName id="3AA6EC826A989EB0992C1AC74A038CA5" ID-CoL="3S67V" authority="(Pocock, 1893)" baseAuthorityName="Pocock" baseAuthorityYear="1893" class="Chilopoda" family="Henicopidae" genus="Lamyctes" kingdom="Animalia" order="Lithobiomorpha" phylum="Arthropoda" rank="species" species="tristani">
<emphasis id="6A47898102A665EEC3633F5F6E724105" italics="true">Lamyctes tristani</emphasis>
(
<bibRefCitation id="0EB6628951E5A9A8C652FD8777720CFE" DOI="10.1080/00222939308677479" author="Pocock RI" issue="6" journalOrPublisher="Annals and Magazine of Natural History" pagination="121-142" refId="B83" refString="Pocock RI (1893) Report upon the Myriopoda of the ' Challenger' Expedition, with remarks upon the fauna of Bermuda. Annals and Magazine of Natural History 6 (6): 121-142. https://doi.org/10.1080/00222939308677479" title="Report upon the Myriopoda of the Challenger Expedition, with remarks upon the fauna of Bermuda." volume="6" year="1893">Pocock, 1893</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="061097DA50486D0472BFD3AA130A8120" type="description">
<paragraph id="D5AAC429467D2741B4672541EA33C97E">
<figureCitation id="A4A7D8DF002FE4079A3C9E6F872DE43B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lamyctes tristani (Pocock, 1893) A, B, D NHMUK 015991452 A forcipular segment, ventral view B anterior margin of forcipular coxosternite, ventral view C NHMUK 015991458, distal end of tibia of leg pair 12, lateral view D distal end of tibia of leg pair 12, lateral view E NHMUK 015991456, Female gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure1" httpUri="https://binary.pensoft.net/fig/1257820">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECDAE" type="material">
<paragraph id="7B67CE8F898FA5D45E59611771C42D13">
<heading id="21F85422A7D9455121C45C78F16C8ED9" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="40E5DF4DCE7A3D0DC5FDF01176C3EC08">
<specimenCount id="B471C7A270D7DE7AC1F0784CCB6A49E3" count="41" type="generic">41 specimens</specimenCount>
:
<materialsCitation id="AE7BEDF839C6018F17FB6093F34D4B08" collectingDate="1974-03-16" collectionCode="NHMUK" location="Cinq Cases" specimenCode="NHMUK 015991449" specimenCount="1" specimenCount-female="1">
<specimenCode id="F75C925F90FC6174B0E6DF6C6EC6FEE3">NHMUK 015991449</specimenCode>
,
<specimenCount id="299FC19DF1EE3DA45DC3C426070C2AD8" count="1" type="female">1 ♀</specimenCount>
,
<location id="A04047BBE3784353F6C562FC0962947F" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:A04047BBE3784353F6C562FC0962947F" name="Cinq Cases">Cinq Cases</location>
,
<date id="FCF5341629CD2685B0CB74F86EE394B7" value="1974-03-16">
<collectingDate id="7461DEB62ADC6EDCE56BF2BBAC185607" value="1974-03-16">16. 03. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="4DFF71D89C9FE821B4F090C32B843444" collectingDate="1975-02-14" collectionCode="NHMUK" location="Malabar" specimenCode="NHMUK 015991450" specimenCount="2" specimenCount-female="2">
<specimenCode id="8E1E277D563A52B8D41BE60FAD08A4EB">NHMUK 015991450</specimenCode>
,
<specimenCount id="7EC26719B7F8EAEFE9A289577510F78D" count="2" type="female">2 ♀</specimenCount>
,
<location id="802B013D01DCD1A027FB0B6C8F72E665" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:802B013D01DCD1A027FB0B6C8F72E665" name="Malabar">Malabar</location>
,
<date id="AD7518A0E89C5DE18BCA868D50337575" value="1975-02-14">
<collectingDate id="420C4779D2B8E29C6AD34CE2B9CF7A17" value="1975-02-14">14. 02. 1975</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="2CA5C5DA768CBD3E0E5107ACC7337008" collectingDate="1975-02-28" collectionCode="NHMUK" location="Picard" specimenCode="NHMUK 015991451" specimenCount="1" specimenCount-female="1">
<specimenCode id="1C691A89A9C946014CA908C5DCD4633D">NHMUK 015991451</specimenCode>
<specimenCount id="BDD4766C5E69E408EE0E18A09A46F39B" count="1" type="female">1 ♀</specimenCount>
,
<location id="EED73BBBFC6E5A805895A2ABED571553" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:EED73BBBFC6E5A805895A2ABED571553" name="Picard">Picard</location>
,
<date id="66FDF1D05DABF1E85B6833537315E154" value="1975-02-28">
<collectingDate id="095FF46927647952CF3F0E4C73C473E7" value="1975-02-28">28. 02. 1975</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="686375DD406F1E6CD3BB2C1081EEAA57" collectingDate="1974-03-20" collectionCode="NHMUK" location="Cinq Cases B. F." specimenCode="NHMUK 015991452" specimenCount="4" specimenCount-female="3" specimenCount-juvenile="1">
<specimenCode id="1A3BC7FED5148C7F2C31FFF3EBFB9DF6">NHMUK 015991452</specimenCode>
,
<specimenCount id="DD0BC89C5B3958A98E44ED75524AF5C6" count="3" type="female">3 ♀</specimenCount>
,
<specimenCount id="1650433E5A8F32A063A2724EB12FCFD1" count="1" type="juvenile">1 juvenile</specimenCount>
,
<location id="C72465340621407A61EBC79DDF77DEFE" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:C72465340621407A61EBC79DDF77DEFE" name="Cinq Cases B. F.">Cinq Cases B. F.</location>
,
<date id="20489401CC339694E0B4BFDDAFC9723C" value="1974-03-20">
<collectingDate id="59060BFEA1EBDA56DD53EFA1C45C8238" value="1974-03-20">20. 03. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="79E417899DC7C444ABA55EE9CA84F7F9" collectingDate="1974-02-23" collectionCode="NHMUK" location="Picard" specimenCode="NHMUK 015991453" specimenCount="2" specimenCount-female="2">
<specimenCode id="C03727E60F3047F15CA26E70CE055D25">NHMUK 015991453</specimenCode>
,
<specimenCount id="9707856D7FCA4F57379D751AF9808032" count="2" type="female">2 ♀</specimenCount>
,
<location id="F05F5289D869DA329E61197A39496D05" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:F05F5289D869DA329E61197A39496D05" name="Picard">Picard</location>
,
<date id="749F94A8DB9694A7972D91DE9CD510A1" value="1974-02-23">
<collectingDate id="939D6DE8D43B3F7D7341346D1D97B192" value="1974-02-23">23. 02. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="79A0494D3D4E23D5288A23864D2DBF2B" collectingDate="1974-03-24" collectionCode="NHMUK" location="Cinq Cases" specimenCode="NHMUK 015991454" specimenCount="2" specimenCount-female="2">
<specimenCode id="129919F3F0D873E2AF85F90895A6C790">NHMUK 015991454</specimenCode>
,
<specimenCount id="9A21196F7F7103193EBDD8911991EFEB" count="2" type="female">2 ♀</specimenCount>
,
<location id="1F1B8639A94C5B215EBD4389DE483845" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:1F1B8639A94C5B215EBD4389DE483845" name="Cinq Cases">Cinq Cases</location>
,
<date id="DC25D3E8DBDD6079A71C89CF368C0207" value="1974-03-24">
<collectingDate id="833ACC2BE44D0C21B5668A2CBDD811A6" value="1974-03-24">24. 03. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="2F059C9410853C2C7C6197C05FAD6348" collectingDate="1974-06-08" collectionCode="NHMUK" location="Malabar" specimenCode="NHMUK 015991458" specimenCount="3" specimenCount-female="3">
<specimenCode id="7B13F8A170F5625834236FF4850D9E1C">NHMUK 015991458</specimenCode>
,
<specimenCount id="0D50B0BA93F9C425BC0D605A20257585" count="3" type="female">3 ♀</specimenCount>
,
<location id="2D39CD06142210DD4270C112C987AE57" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:2D39CD06142210DD4270C112C987AE57" name="Malabar">Malabar</location>
,
<date id="685AD8D628630903E24D513D7545CD52" value="1974-06-08">
<collectingDate id="42F196D42A210555BBDB44DC9112CFE2" value="1974-06-08">08. 06. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="2394A9CB125B8622C3AA5283620B3990" collectingDate="1975-01-30" collectionCode="NHMUK" location="Aldabra" specimenCode="NHMUK 015991455" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1">
<specimenCode id="1D9791C0DE586D99C507BE16C122A623">NHMUK 015991455</specimenCode>
,
<specimenCount id="5C3663340E4AD76188F87751AA9187B0" count="1" type="female">1 ♀</specimenCount>
,
<specimenCount id="1F7EEF61CF2D7DE46FBA5C476718F670" count="1" type="juvenile">1 juvenile</specimenCount>
,
<location id="51BD51331D92D512FB9D2D78EC061CED" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:51BD51331D92D512FB9D2D78EC061CED" name="Aldabra">Aldabra</location>
,
<date id="EDE951670A32EEEAE4ABB2B2C88BF17B" value="1975-01-30">
<collectingDate id="6C747D3B803D4D03738FF3F19139248D" value="1975-01-30">30. 01. 1975</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="2CC86DB355B0B3144AEF3F202564349B" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Picard" specimenCode="NHMUK 015991456" specimenCount="21" specimenCount-female="21">
<specimenCode id="7BBB8FC58B75A2FE7C1E01FA35A681B3">NHMUK 015991456</specimenCode>
,
<specimenCount id="85FC067CE27C09A6626F6C015B8DE085" count="21" type="female">21 ♀</specimenCount>
,
<location id="25362C2700317BC3BD968C5E78F3B756" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:25362C2700317BC3BD968C5E78F3B756" name="Picard">Picard</location>
, Summer 1975, leg.
<collectorName id="36090166E5F0C0F4F9D58EE2795CF8A2">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="0CDF16951954336FD0414C21AA0A0213" collectedFrom="Ochna litter" collectingDate="1974-03-23" collectionCode="NHMUK" location="Cinq Cases B. F." specimenCode="NHMUK 015991457" specimenCount="3" specimenCount-female="2" specimenCount-juvenile="1">
<specimenCode id="3CCA096D96ABC08B73B8E10A190595DE">NHMUK 015991457</specimenCode>
,
<specimenCount id="FCAE411AC23EB76E0D5E2AD2EAFBCF93" count="2" type="female">2 ♀</specimenCount>
,
<specimenCount id="8C10C15B15C04ECFF25BA21BE5873903" count="1" type="juvenile">1 juvenile</specimenCount>
,
<collectedFrom id="FD20ADF09F8B654F873344DF2731B111">
<taxonomicName id="24C5E7584F47BC4621F8ABBA3C95C97A" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="genus">
<emphasis id="0CFE46C433BAE2A7A1BF841397904527" italics="true">Ochna</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="384F106DC20CB015D6082D3D9A9F5F2A" LSID="urn:lsid:plazi:treatment:18A70C29CAA05FD888E215C1A621B2A3:384F106DC20CB015D6082D3D9A9F5F2A" name="Cinq Cases B. F.">Cinq Cases B. F.</location>
,
<date id="1C328D8631167E8C295F6888DA2E6A40" value="1974-03-23">
<collectingDate id="87A0944A486A566C040E63A0DC133212" value="1974-03-23">23. 03. 1974</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EODAE" type="remarks">
<paragraph id="A90CBDE4681B70427874109EC8530EAD">
<heading id="0DFEA3C0862D5EB3E6F1CE7A538F311C" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="43DC6C2A62317FE85BB54BD19C92320B">
The
<specimenCount id="B39488C5A5A420A2ACF59871CACF0738" count="38" type="generic">38 specimens</specimenCount>
from the Aldabra Atoll (from Cinq Cases, Malabar, and Picard) for which sex can reliably be determined are all females, such that the population is parthenogenetic. They are likely conspecific with
<taxonomicName id="2206E1C1D503583FB2F7157471162FCF" baseAuthorityName="Pocock" baseAuthorityYear="1893" class="Chilopoda" family="Henicopidae" genus="Lamyctes" kingdom="Animalia" order="Lithobiomorpha" phylum="Arthropoda" rank="species" species="tristani">
<emphasis id="504C08DAB652EB33B3E2C962CB6034F1" italics="true">L. tristani</emphasis>
</taxonomicName>
as described from the Chagos Archipelago (
<bibRefCitation id="1AB19AD280C9F150B56CC63D453E31AB" DOI="10.1080/00222933.2024.2395903" author="Popovici" etAl="et al." firstAuthor="Popovici" issue="41-44" journalOrPublisher="Journal of Natural History" pagination="1885-1915" refId="B87" refString="Popovici G, Edgecombe GD, Hall DW (2024) New Chilopoda from the Chagos Archipelago. Journal of Natural History 58 (4144): 18851915. https://doi.org/10.1080/00222933.2024.2395903" title="New Chilopoda from the Chagos Archipelago." volume="58" year="2024">Popovici et al. 2024</bibRefCitation>
), sharing the distinctly reduced spinous process on the tibia of leg pair 12, often taking the form of a rounded bump (Fig.
<figureCitation id="8A96712FB4E5A3998D9B7C26A37DB7D6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lamyctes tristani (Pocock, 1893) A, B, D NHMUK 015991452 A forcipular segment, ventral view B anterior margin of forcipular coxosternite, ventral view C NHMUK 015991458, distal end of tibia of leg pair 12, lateral view D distal end of tibia of leg pair 12, lateral view E NHMUK 015991456, Female gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure1" httpUri="https://binary.pensoft.net/fig/1257820">1 C</figureCitation>
). The stability of this character is however uncertain as specimens from localities in the Aldabra Atoll exhibit conspicuous variation in the shape of the process (Fig.
<figureCitation id="FC7EABCCC6C914B0DB83F240E1D224A9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lamyctes tristani (Pocock, 1893) A, B, D NHMUK 015991452 A forcipular segment, ventral view B anterior margin of forcipular coxosternite, ventral view C NHMUK 015991458, distal end of tibia of leg pair 12, lateral view D distal end of tibia of leg pair 12, lateral view E NHMUK 015991456, Female gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure1" httpUri="https://binary.pensoft.net/fig/1257820">1 D</figureCitation>
, Table
<tableCitation id="405B1D1609D57E2AC8BF9915B47ADAFB" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological variability of putative L. tristani specimens from Western Indian Ocean localities. Asterisk indicating regenerated appendage." httpUri="http://table.plazi.org/id/E7BCCC7F334CDE655A61EA104D28DA1B" tableUuid="E7BCCC7F334CDE655A61EA104D28DA1B">1</tableCitation>
). Additional similarities between these two populations include the morphology of the forcipular coxosternite (Fig.
<figureCitation id="B4E47BD5771FE39390451284F98195E4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lamyctes tristani (Pocock, 1893) A, B, D NHMUK 015991452 A forcipular segment, ventral view B anterior margin of forcipular coxosternite, ventral view C NHMUK 015991458, distal end of tibia of leg pair 12, lateral view D distal end of tibia of leg pair 12, lateral view E NHMUK 015991456, Female gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure1" httpUri="https://binary.pensoft.net/fig/1257820">1 A, B</figureCitation>
), the number of antennal articles, and the number and shape of the spurs on the female gonopods (Fig.
<figureCitation id="C3916C2AFEF06AA05C7BE76642ED4975" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Lamyctes tristani (Pocock, 1893) A, B, D NHMUK 015991452 A forcipular segment, ventral view B anterior margin of forcipular coxosternite, ventral view C NHMUK 015991458, distal end of tibia of leg pair 12, lateral view D distal end of tibia of leg pair 12, lateral view E NHMUK 015991456, Female gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure1" httpUri="https://binary.pensoft.net/fig/1257820">1 E</figureCitation>
). Variability in the characters of antennal article number, number, and distribution of coxal pores and shape of the distal spinous process on the tibia of leg pair 12 are listed in Table
<tableCitation id="A0B1CBB5F5EB1CBC3036E33F1E78755B" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological variability of putative L. tristani specimens from Western Indian Ocean localities. Asterisk indicating regenerated appendage." httpUri="http://table.plazi.org/id/E7BCCC7F334CDE655A61EA104D28DA1B" tableUuid="E7BCCC7F334CDE655A61EA104D28DA1B">1</tableCitation>
and compared with putatively conspecific individuals from the Chagos Archipelago, excluding anamorphic stages.
</paragraph>
<caption id="E7BCCC7F334CDE655A61EA104D28DA1B" ID-Table-UUID="E7BCCC7F334CDE655A61EA104D28DA1B" httpUri="http://table.plazi.org/id/E7BCCC7F334CDE655A61EA104D28DA1B" startId="T1" targetIsTable="true">
<paragraph id="066353F428D5D24880E086D63221EAD8">
<label id="C560287E2AC0E73B5866FF195666A258">Table 1.</label>
</paragraph>
<paragraph id="F4234E43F56688149213D21D88AAE9C1">
Morphological variability of putative
<taxonomicName id="F83A805D465A360851A5315B5DF94052" baseAuthorityName="Pocock" baseAuthorityYear="1893" class="Chilopoda" family="Henicopidae" genus="Lamyctes" kingdom="Animalia" order="Lithobiomorpha" phylum="Arthropoda" rank="species" species="tristani">
<emphasis id="468DB75C434FF474F3E3AE3833D61CA0" italics="true">L. tristani</emphasis>
</taxonomicName>
specimens from Western Indian Ocean localities. Asterisk indicating regenerated appendage.
</paragraph>
</caption>
<paragraph id="FA7F70EF27FC3B31660E49FEE138BA75">
<table id="TID0ERJCI" rules="all">
<tbody id="B2F87EB49B7E4F3A548AAB1ED932C51C">
<tr id="2BDCCA2995B07D0D20CF7C91061696ED">
<th id="C0EAF14009C3B513E9DC45C2E42D65CE" colspan="1" rowspan="1">Locality</th>
<th id="DA491D5670058A30CBEE8E8F733B3045" colspan="1" rowspan="1">Body length (mm)</th>
<th id="78FA3A6E0DB8CAE383D9C2391846E696" colspan="1" rowspan="1">Antennal article number</th>
<th id="B4A89AD5E02B090B7E9D8B31B1F4895D" colspan="1" rowspan="1">Coxal pore formula (legs 12 / 12 13 / 13 14 / 14 15 / 15)</th>
<th id="50A918D039D65FAC13A1B9C7C2B7F6EB" colspan="1" rowspan="1">Shape of tibial spinous process on leg pair 12</th>
</tr>
<tr id="4CB804C3E8A6D8C1ACA94B579329A7DC">
<td id="EBE159F0AAF6176C62CB0671E047678A" colspan="1" rowspan="1">Cinq Cases (Aldabra)</td>
<td id="456B5DFAF7304CAF95E47C6A5B32E885" colspan="1" rowspan="1">3.5</td>
<td id="8C49D688CE670EAB65279B73D8A4E8F4" colspan="1" rowspan="1">20</td>
<td id="FAAA4AA735B9405B3A0CEE161EE2D154" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="037467812D3B467F366E213E59FDA6A3" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="6FB5E6138189E90A778D451BED8EECBF">
<td id="27E4FBB86EAF407C2A81C89143EA56EF" colspan="1" rowspan="1">Cinq Cases (Aldabra)</td>
<td id="805E6E01007B965FB4FD54E2CD498612" colspan="1" rowspan="1">4</td>
<td id="1DE1A551B3E7C7722672266868992264" colspan="1" rowspan="1">20</td>
<td id="0B21BD0714D556BA4A9726458002C186" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="E25039FB0459D156B48004B8DEB74476" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="49F0B6AF4692D6B1B1B94E7CD1428ECC">
<td id="A92E083949C2A085EE077205CA49F71A" colspan="1" rowspan="1">Cinq Cases (Aldabra)</td>
<td id="8DF64C57847C5479C424A9504E2CA3AD" colspan="1" rowspan="1">5</td>
<td id="E5F065C2450AED375842E6FA1ED26BC9" colspan="1" rowspan="1">17</td>
<td id="8405B28C22A23AAC8B20009962845053" colspan="1" rowspan="1">2 / 2 2 / 2 3 / 3 2 / 2</td>
<td id="98D9F127DE423900ACF948CB6BC05CBD" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="60F61C9D172F07C0B47C0DAB21E8B869">
<td id="3E61CF20FBA2CD34EFCC0B91602E6254" colspan="1" rowspan="1">Cinq Cases (Aldabra)</td>
<td id="F8BD267185836C373BED1F83AF0854D9" colspan="1" rowspan="1">5</td>
<td id="FB0DFB66F2C7CDB9D4083B3999071658" colspan="1" rowspan="1">22</td>
<td id="BF24E53281371FF409D27EC6278AC6F6" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 2 / 2</td>
<td id="E3C854F82736297EEFBC3F099FC11B2F" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="F603B75182186E6F70584C9AA14212B2">
<td id="3629B2A87771FCEE7196350ADA113A8E" colspan="1" rowspan="1">Cinq Cases (Aldabra)</td>
<td id="CF1B7201F530D2DADE1030F2F9A533B6" colspan="1" rowspan="1">5</td>
<td id="22677B0303DAE1EF42232FB9CB770F20" colspan="1" rowspan="1">-</td>
<td id="7318DFAC3FE1B8A8368A16F09EBE6702" colspan="1" rowspan="1">2 / 2 3 / 3 3 / 3 2 / 2</td>
<td id="75169E7A8D852D9AA2D91915FCD959CE" colspan="1" rowspan="1">-</td>
</tr>
<tr id="973D170F513925E1F1790764FC6B2A9A">
<td id="D37548C180606C5C9FC41469A69234C0" colspan="1" rowspan="1">Cinq Cases (Aldabra)</td>
<td id="1AC9045B50059AF0F853D260C0D8BEB4" colspan="1" rowspan="1">5</td>
<td id="A85737EF68DE62C8884EB0D8559A5E6B" colspan="1" rowspan="1">22</td>
<td id="65B5A0F7E070B5535463996D5EB65E96" colspan="1" rowspan="1">2 / 2 3 / 3 3 / 3 2 / 2</td>
<td id="BEA4960D72053DD8DCE1F28951EDD6A6" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="0952FDC901E6EDBE26CA8E6334194462">
<td id="E0D60E9D6B5B8DC8C14350F871B39CF2" colspan="1" rowspan="1">Cinq Cases (Aldabra)</td>
<td id="9708FBBF125F485C725598B8CB05497C" colspan="1" rowspan="1">6</td>
<td id="36A6C2AFE3D64A6A24C89D24B484AF40" colspan="1" rowspan="1">23</td>
<td id="D3BD1B81BFF4E5675A71DEBFC4B0E3B1" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 3 / 3</td>
<td id="56C52C09D28E58D43A9745430B98CBFA" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="4B642743C89A0A7E090D52BE4C765590">
<td id="0F7E6638B985F8B044DD006326A37826" colspan="1" rowspan="1">Malabar (Aldabra)</td>
<td id="2EDEC8E3A184BCCCCEC7EA1D0FE08D1F" colspan="1" rowspan="1">5.5</td>
<td id="C1F0561937B7B36C2095AF8B71F5A1AE" colspan="1" rowspan="1">24</td>
<td id="4E731FB77913E8959C309FF717429B51" colspan="1" rowspan="1">2 / 2 2 / 2 3 / 3 2 / 2</td>
<td id="55F860ABE516A554DA5B65A4004856F1" colspan="1" rowspan="1">-</td>
</tr>
<tr id="ADC95CAAE1FC16FC03FBA202F239981D">
<td id="1D4F070F520618240168FF433B43E503" colspan="1" rowspan="1">Malabar (Aldabra)</td>
<td id="5DD8ADE9920633FEAEC387C1366FF23D" colspan="1" rowspan="1">5.5</td>
<td id="658EE31514CD116499020AAB91391345" colspan="1" rowspan="1">24</td>
<td id="73B47CFABBC001128DF216A938C3DA52" colspan="1" rowspan="1">2 / 2 2 / 2 3 / 3 2 / 2</td>
<td id="3C7158BBCDC166DA7B161481A5CD2615" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="0E16D99BDDB3CA4F29B0A1A130D7A1E4">
<td id="9A7A49C2498E0ECC9422F00E1953091C" colspan="1" rowspan="1">Malabar (Aldabra)</td>
<td id="69DADC059FE5924C46E81CA12A4FD14E" colspan="1" rowspan="1">6</td>
<td id="0EBC50EB2F9505C19C374CD94CBF52F3" colspan="1" rowspan="1">(23) 24</td>
<td id="35F2D08EA9CAC48F6F80C3E610ABD93B" colspan="1" rowspan="1">2 / 2 2 / 2 2 / 3 2 / 2</td>
<td id="19F9C67C19D2A898D396CA8E9114814D" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="5156EA99DD929115632960161167DFC1">
<td id="C2A7CFCB4D3A22F70C45F3106BE86E7B" colspan="1" rowspan="1">Malabar (Aldabra)</td>
<td id="4CBD5FADEC92216F2E39128E7BAF19A1" colspan="1" rowspan="1">6</td>
<td id="02FD3782A6946AFE537636F4E1079025" colspan="1" rowspan="1">15 (20)</td>
<td id="5443B6341BDE7DE27F75720768C241FC" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 3 / 3</td>
<td id="1157BE13D9DE5DD5C68844B643A565A6" colspan="1" rowspan="1">Asymmetrical, rounded bump on left tibia, acuminate spur on right tibia</td>
</tr>
<tr id="DDDB6F677CCBD48ADDB88F5FA0983CD0">
<td id="9318A50EF0D790DE0A94F3A27C09C863" colspan="1" rowspan="1">Malabar (Aldabra)</td>
<td id="D3242D90EC01337DBE2E22FC7E65C347" colspan="1" rowspan="1">6</td>
<td id="09E9CF620603777786C169D2F9EAE213" colspan="1" rowspan="1">24</td>
<td id="CE473D8F115B99FD22B128018FEB4A44" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 3 / 3</td>
<td id="985A1B974AB20D60497A157C3E11BD41" colspan="1" rowspan="1">-</td>
</tr>
<tr id="292ECCB12F25F1001B7696AAC7DDF03B">
<td id="8AF964E3042F3ACA9EC91DF86D1305BD" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="92E60F2E8071F36462AB4EC0D9D74111" colspan="1" rowspan="1">3.5</td>
<td id="15363F9FDFDA50934775B7588FDA297E" colspan="1" rowspan="1">19</td>
<td id="7DE6D9D8733D76F6447F2E24C963CD2D" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="85695BBD8AFBFC24702DDB83C49A8827" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="9C028A7F155AF8263ECD7A75CDC631D2">
<td id="EBCD223F62937BB81378C8CDEE405C3A" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="948441FE6D8C25319A2648587DBB506C" colspan="1" rowspan="1">3.5</td>
<td id="FB2620FFF55A959614D7BB6D35832992" colspan="1" rowspan="1">20</td>
<td id="AD440F7B78B2477E58C51765E9E16DE9" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="C0220A186FCA8DDE481985597FFFB86B" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="92D0828FBA6578CC6D45CD8E3F269355">
<td id="A5198E95D9F31ED5B84C75B8CFC06778" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="EC687181AE901207FD9A71B27EACF2E3" colspan="1" rowspan="1">3.5</td>
<td id="16EEE3ACA28E38787CB307C9FE4C1F8F" colspan="1" rowspan="1">20</td>
<td id="BA06CB21CE1AB413BEF30361F8B29154" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="16CAC6F46D04AC2E842F8E0D74B17615" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="B4AEE5CD9E09670CD964F815D651D4E4">
<td id="FF4C028614F571E0B1CE026D98A00773" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="546CFF884B19AF6E8C17202939D2C065" colspan="1" rowspan="1">4</td>
<td id="B98F82E755AB6E224753EC844021C0C5" colspan="1" rowspan="1">20</td>
<td id="A17BB636765F1E26A3DE0B8880475B2B" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="AFCFF68A6C8E785B7558CE855BA85FE8" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="6E7B5C2E6D3B47888D44C14E7380F0FD">
<td id="EBF4F4D1766C46874610D157C3954C5E" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="8079750A711C87B861C9889410E822B5" colspan="1" rowspan="1">4</td>
<td id="9D89CBDC948A7096FE1E4D77E4E98618" colspan="1" rowspan="1">20</td>
<td id="1337A6D6DA8C55996950FA530FABC40E" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="22314337FA531007C2A9456D8C91A93E" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="98E925CB745255973167B87455C48F43">
<td id="0DDC4CDE7FB4C8A06E7638EFF9CA55D3" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="0E311C56DF4B0E73E77FE49F4805BB36" colspan="1" rowspan="1">4</td>
<td id="0D3FDAF64ABCCEF9B997A56D3982913E" colspan="1" rowspan="1">20</td>
<td id="DA4FB40B0D61FEE9E9DD956F274C8B57" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="BBDB1F6D6B0ED6A1E17A63F1503FD9AC" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="1B20AC9DE02380D4B1CCC57394BCFD9E">
<td id="9E0EB37A6A6D3282E307EFF1C3F1EDB0" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="B746AA9051DC75EA77D64B1397154D66" colspan="1" rowspan="1">4</td>
<td id="1ECC5326DFBC89419446BAAE6C5125FC" colspan="1" rowspan="1">20</td>
<td id="D8F31249B69D223390D486EF79B0CB64" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="9ABA732D6280C72C86093DE7DBD7467C" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="11F032DC6047154BBD8C0FF7C5DCC4C2">
<td id="CE945ACFF4C300E68A4A24B3BE828A9B" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="08CAA842FF0E2521C3792923D10C996A" colspan="1" rowspan="1">4</td>
<td id="4050EED61EF5E19B648612DA07AE20C0" colspan="1" rowspan="1">21</td>
<td id="58C6D713A33475DE9BBE60C43DE35E1B" colspan="1" rowspan="1">2 / 2 2 / 2 2 / 2 1 / 1</td>
<td id="2123DD92CA5EFA974129308AD9E0FB5A" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="E0E737ACC5A62AA697FCDA11C75705DA">
<td id="B6E996C079DE02BB80D8D9C283CA94C0" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="BE0B0EB08C546B16F281F8C8114C1FA0" colspan="1" rowspan="1">4</td>
<td id="CF64A6A1B2668FF6BD274F6B9A677A24" colspan="1" rowspan="1">21</td>
<td id="BA68FE8E63C622D024F2C37985A0B756" colspan="1" rowspan="1">2 / 2 2 / 2 2 / 2 1 / 1</td>
<td id="E7C3A5AE1CCF1E9DC4866E8F504E25CE" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="7553B4C16BB19DE8D8C05E10B097A421">
<td id="F3D6E7B066F83F650899C1DFF5FEB4CE" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="9CAB508EF71D5F15F972DB1244FE4695" colspan="1" rowspan="1">4</td>
<td id="DB53DBF5037E8C8009E67F074D4F9DFD" colspan="1" rowspan="1">20</td>
<td id="6EA209C24E802F5129A07AAC19BD7CFA" colspan="1" rowspan="1">1 / 1 1 / 1 1 / 1 1 / 1</td>
<td id="5A78CA4BDF5B28CF6BF102FB46521368" colspan="1" rowspan="1">-</td>
</tr>
<tr id="6469EED188BB4B3C9285D045A2B311EE">
<td id="5C2AE4A50871299C326BDE6FBD6C6B98" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="A739B4A000DCF8D57AB8C5D1F6224426" colspan="1" rowspan="1">4.5</td>
<td id="C8D12BCA6D04C0E22722509C36B9644C" colspan="1" rowspan="1">24</td>
<td id="E08E76C09DA552F6AC0F77AA53D947CA" colspan="1" rowspan="1">2 / 2 2 / 2 3 / 3 2 / 2</td>
<td id="1912CCEAF586E9138FB3E785A63E094F" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="8932B6A151FE985F1102BD1195BCBF76">
<td id="3B6172B17011D1716D5A7376F26D61B5" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="B885CF08ECFC4A0BBFBF80FCEEF78E8D" colspan="1" rowspan="1">4.5</td>
<td id="112521876BA2094FCFC7AEAD748A7DB3" colspan="1" rowspan="1">(22) 23</td>
<td id="78BA561C149535BA9B9ECB875F9E0D90" colspan="1" rowspan="1">2 / 2 2 / 2 3 / 3 2 / 2</td>
<td id="A175DC1B2C854D00466B81DEFC429B18" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="DEF8F84B1151C85196888E8EB2F04507">
<td id="987DDB393D2C84C2D8F49F910B13EC9E" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="C1E3B75525AB0E5BDD52B480066ED78F" colspan="1" rowspan="1">4.5</td>
<td id="452A504186A22DF208B2837462193F1F" colspan="1" rowspan="1">21</td>
<td id="D0C40ECF409F260B5D1593F7215968DA" colspan="1" rowspan="1">1 / 1 2 / 2 2 / 2 2 / 2</td>
<td id="78C3EBE098BC1B9A777E4ED6CA56CCE5" colspan="1" rowspan="1">-</td>
</tr>
<tr id="16D0851A9E2EE8DDA3C91C56F627A46A">
<td id="8F0D6E160B2A3322F6A541AF2FE92C31" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="06FEB4253DFC956E7B208557565D5C8B" colspan="1" rowspan="1">5</td>
<td id="EC89ECC81E466112B8C8FA5EC44287F1" colspan="1" rowspan="1">(24) 25</td>
<td id="EC5193F5AD2F82420BA7E7F82BCF6BC4" colspan="1" rowspan="1">2 / 2 2 / 2 2 / 2 2 / 2</td>
<td id="6055B2802C06DB19DCF7C4EAD1B38F30" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="AB2401E1A4EE0AFC040CB61F49963C78">
<td id="BF57C3A42FF4BE52EAA986CC3A31FB97" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="9E907D2040A7C11BF35CCB09F15718A9" colspan="1" rowspan="1">5</td>
<td id="6412FEC1C54F9A23F418627D6F3E58AC" colspan="1" rowspan="1">(22) 23</td>
<td id="E426633A1B4059B2922FBC0B8FCE08E5" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 2 / 2</td>
<td id="70CE6D2AE8CD078545DE31EA5349F606" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="3457C2B79D9C642654BF6D8249A3A2D9">
<td id="358060BAAF86C87903E6E7194A239C4D" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="9ABD30BFD171ED528B952FD94B6671F7" colspan="1" rowspan="1">5</td>
<td id="85526F671E1F14FBBF94F8A928D63755" colspan="1" rowspan="1">15 *</td>
<td id="E3DA4AB24879548BC1BE11214E9AEC82" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 2 / 2</td>
<td id="DDBCC6C189E8A9E705AACECAAA235583" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="29828BEA19B9EDA584E5D0A9519DE141">
<td id="1F11962986756034F03C2C6EC1F7E507" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="664C871A7C3E304331A965BC4D6B5E15" colspan="1" rowspan="1">5</td>
<td id="87A27922E29B0F6D2DD54D9BEC019C8C" colspan="1" rowspan="1">24</td>
<td id="97808F5C47551F3C4D84A02C7746B9D2" colspan="1" rowspan="1">2 / 2 3 / 3 3 / 3 2 / 2</td>
<td id="7044144E5EA4F300469640AEF6019EA6" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="C92135AD721033021023D9119798DEFA">
<td id="0AAF9E7FC932905C7CE5E79B5D21B7DC" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="98BCD84E43410C5B41954744C4CCEBEC" colspan="1" rowspan="1">5</td>
<td id="C733926C99953E4415781D08269D17B0" colspan="1" rowspan="1">(21) 24</td>
<td id="644852449B4FAE04324CEAE5F8B5953D" colspan="1" rowspan="1">2 / 2 3 / 3 3 / 3 2 / 2</td>
<td id="505F15BF234882853681A73814CF115D" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="A6B818A74A9F3ED151994CDB7283A806">
<td id="220AAD16904B6EB2B22866C444BBE9AB" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="B8DC8F3A82E274341E10D5D644B57D6B" colspan="1" rowspan="1">5</td>
<td id="076D0B32B28DAAD0E1F2B23DDDC1E44A" colspan="1" rowspan="1">-</td>
<td id="79ADE70602E64249026B6CC92A03F986" colspan="1" rowspan="1">2 / 2 3 / 3 3 / 3 2 / 2</td>
<td id="7F8121350627B064EF21465BE0CF71AD" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="D37D67E79CA060DDEB1502EE7A92C9ED">
<td id="666689776658F8A8E792A70E89933840" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="F28170775260E0F6077AD6FB7047306B" colspan="1" rowspan="1">5</td>
<td id="08754A3FE4463689D046BDC09AAA158C" colspan="1" rowspan="1">24</td>
<td id="F9DBA4E45574B752545547FE2B062F77" colspan="1" rowspan="1">2 / 2 3 / 3 3 / 3 2 / 2</td>
<td id="ABCE5EE5FC6BCFA70186AD46EF992479" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="DC472D02FCF52B8761002FEA2DAE4B46">
<td id="D77E9653C090B548A0B4011CF0A41C34" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="6F002A5A6F55B72970FA423C3EEF7BA7" colspan="1" rowspan="1">5</td>
<td id="EFD0889E14DCB93B778BF76A7D9EF260" colspan="1" rowspan="1">20</td>
<td id="60E0BDA3F88193C38FA21931100BF32D" colspan="1" rowspan="1">2 / 2 2 / 2 3 / 3 2 / 2</td>
<td id="F3F21FC1591D0845E041A20322A8CC61" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="BFD3C316C6DD3556FEEE1AB1D770D659">
<td id="AF2831AAF37E8A49DADA4FB1CD52D88F" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="4A3A65FEA4FD9F6BA9753CAC2615D98A" colspan="1" rowspan="1">5.5</td>
<td id="C2674E179E3C322B1E64E11AE9BE75BF" colspan="1" rowspan="1">24</td>
<td id="8ED1F3C299006B0DBAD15A93B3D76BCC" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 3 / 3</td>
<td id="15F4C8041095B78F1311FA5E3730E5E1" colspan="1" rowspan="1">Acuminate, minute</td>
</tr>
<tr id="24265A2030024D21FAA4BF86A888AD33">
<td id="B4450546D5C7F1F59034179B5D0CF523" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="886F06B0354465FACAD1D747F3D1B7E7" colspan="1" rowspan="1">6</td>
<td id="A9B218E3E6DD3817D6ECD859A3842EE5" colspan="1" rowspan="1">24</td>
<td id="9D13A8660E9349B9D9FC36AFC9998916" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 3 / 3</td>
<td id="9F8D898F2D28F27CB2B53A6CCA5BB54F" colspan="1" rowspan="1">Acuminate, reduced in size</td>
</tr>
<tr id="236BB69670522D17FD6130B336D87846">
<td id="F0A04793E313377356934CA8300FF860" colspan="1" rowspan="1">Picard (Aldabra)</td>
<td id="227FD702B4B628CC05F55B22717A8F49" colspan="1" rowspan="1">7</td>
<td id="6814335599DF91EE51F6542C5FBDD5DD" colspan="1" rowspan="1">(24) 25</td>
<td id="7C067CE754A5BF47ABD95B5E71CEE1C1" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 3 / 3</td>
<td id="4913B11BB90FB6563B48AAF16886F8AD" colspan="1" rowspan="1">Acuminate, reduced in size</td>
</tr>
<tr id="9A2A870AC0C5295549657B5FAA6E44FC">
<td id="CE334E3C1E02AC3E1207E29073C7CF4A" colspan="1" rowspan="1">Diego Garcia (Chagos)</td>
<td id="2976036874E2037CA6F080BB4321F2FF" colspan="1" rowspan="1">6</td>
<td id="2AB707CB775D880BD80D8FC49E84B565" colspan="1" rowspan="1">27</td>
<td id="02A4D6E537374F175FD96204FA5F296E" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 3 / 3</td>
<td id="8656CBC896270AE7A42E1F5F1E8C2C7E" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="AC568B7124E7E2D6CEA57C28CD77C23E">
<td id="66FC014358C6425163B376764DA3DE6E" colspan="1" rowspan="1">Diego Garcia (Chagos)</td>
<td id="B3A8F8DA7D9DE4613669E8DC1899F0C6" colspan="1" rowspan="1">6.2</td>
<td id="B8D8DECF707217B9DCFC89FCDAF21C64" colspan="1" rowspan="1">(18) 20</td>
<td id="10B8D13F121C3433482089EEDF5C3C84" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 3 3 / 3</td>
<td id="D36CE52078C3DC1C4D68598AAF302355" colspan="1" rowspan="1">Rounded bump</td>
</tr>
<tr id="3B00E944C21D2C1063B1C4760AE4E200">
<td id="621B49823185C2E9D741FC08A9FE1604" colspan="1" rowspan="1">Diego Garcia (Chagos)</td>
<td id="2341CD539B9CD7EBF3F3B06C834D5D42" colspan="1" rowspan="1">6.5</td>
<td id="33EBEDB08AF5BE1537A89CD1F1CCBB32" colspan="1" rowspan="1">(21) 24</td>
<td id="25BEABD76480B1ABE6ED2BCAA5FC97AF" colspan="1" rowspan="1">3 / 3 3 / 3 3 / 4 4 / 4</td>
<td id="F666FB1A79BC6A6B599A88C06957AC7E" colspan="1" rowspan="1">Rounded bump</td>
</tr>
</tbody>
</table>
</paragraph>
<caption id="D5CC719813573C9653A89C4B603E0FBE" ID-DOI="10.3897/zookeys.1228.143007.figure1" ID-arpha="2DD18A5C-80F5-5260-9F3B-B864739B6324" httpUri="https://binary.pensoft.net/fig/1257820" startId="F1">
<paragraph id="9134790D605951BBF0F7AD6814B3C869">
<label id="329B29240F4A29D432ACD0A79030E3AF">Figure 1.</label>
</paragraph>
<paragraph id="1E0C7B780AF27C2F97AE652B77ACFF54">
<taxonomicName id="47F6CEC5843C462D9668FCFED02BA12F" authority="(Pocock, 1893)" baseAuthorityName="Pocock" baseAuthorityYear="1893" class="Chilopoda" family="Henicopidae" genus="Lamyctes" kingdom="Animalia" order="Lithobiomorpha" phylum="Arthropoda" rank="species" species="tristani">
<emphasis id="20A9AB4984367B6C43D761D37A4E20BC" italics="true">Lamyctes tristani</emphasis>
(
<bibRefCitation id="8D0B0204578EC2A0D7570D6C61810A45" DOI="10.1080/00222939308677479" author="Pocock RI" issue="6" journalOrPublisher="Annals and Magazine of Natural History" pagination="121-142" refId="B83" refString="Pocock RI (1893) Report upon the Myriopoda of the ' Challenger' Expedition, with remarks upon the fauna of Bermuda. Annals and Magazine of Natural History 6 (6): 121-142. https://doi.org/10.1080/00222939308677479" title="Report upon the Myriopoda of the Challenger Expedition, with remarks upon the fauna of Bermuda." volume="6" year="1893">Pocock, 1893</bibRefCitation>
)
</taxonomicName>
<emphasis id="D3C513E59575F23B72794F28A23A48EA" bold="true">A, B, D</emphasis>
<specimenCode id="9DE56E048EB9E89A75B357E1E1AE6267">NHMUK 015991452</specimenCode>
<emphasis id="8682F530221840B2FCA5601A19A4AD15" bold="true">A</emphasis>
forcipular segment, ventral view
<emphasis id="C777AF019EB3DE073C1F9D974B7372BE" bold="true">B</emphasis>
anterior margin of forcipular coxosternite, ventral view
<emphasis id="08D705B1B09F305667E2B551AA7AE093" bold="true">C</emphasis>
<specimenCode id="EAF8E8FABE99331CBD8F3C2BE4DC97D3">NHMUK 015991458</specimenCode>
, distal end of tibia of leg pair 12, lateral view
<emphasis id="363933497F2972255596994EF1FA1D9A" bold="true">D</emphasis>
distal end of tibia of leg pair 12, lateral view
<emphasis id="5AD3E42371869F54D4D9E9A8E16A0A8C" bold="true">E</emphasis>
<specimenCode id="5F099A2986FC8D8D13C9542A981BDF8F">NHMUK 015991456</specimenCode>
, Female gonopods, ventral view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,566 @@
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<paragraph id="F2E93AFFCA29782EB51A56F81A3BD327">
<taxonomicName id="BCBB067002A1E93EA546D4E17D7B0311" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="92EE5E57446316C189B4D0EF2A63B958" italics="true">Orphnaeus dekanius</emphasis>
<bibRefCitation id="A35861C2CDD54C0D7B6A1EC979352FB6" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbücher, Abteilung für Systematik" pagination="339-388" refId="B107" refString="Verhoeff KW (1938) Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik 71: 339-388." title="Chilopoden-Studien zur Kenntnis der Epimorphen." volume="71" year="1938">Verhoeff, 1938</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="34A206EEB6084DB4D74437A3626502EB" type="description">
<paragraph id="D441014BD62DE2F34CD3935AB7337867">
<figureCitation id="8B75FD38BA703E91F2F54DE6C2FF7500" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orphnaeus dekanius Verhoeff, 1938 A, B, D NHMUK 015991470 A, C NHMUK 015991474. Head and antennae, dorsal view B antennal article XIV, dorsal view D forcipular segment, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure12" httpUri="https://binary.pensoft.net/fig/1257831">Figs 12</figureCitation>
,
<figureCitation id="C985498E4BA053FBC396AA7F2C2E8E59" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Orphnaeus dekanius Verhoeff, 1938. NHMUK 015991470 A clypeus and labrum, ventral view B labrum, ventral view C, D Mandible, lateral and dorsal views." figureDoi="10.3897/zookeys.1228.143007.figure13" httpUri="https://binary.pensoft.net/fig/1257832">13</figureCitation>
,
<figureCitation id="73A1D55365D00B3F9AA16C0E111F7568" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Orphnaeus dekanius Verhoeff, 1938 NHMUK 015991470 A first and second maxillae, ventral view B first maxillary right telopodal lappet, ventral view C first maxillary left telopodal and coxosternal lappets, dorsal view D, E left and right second maxillary pretarsi, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure14" httpUri="https://binary.pensoft.net/fig/1257833">14</figureCitation>
,
<figureCitation id="EFA18D8F9B0DC72A6EB8D3B08A12EA53" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Orphnaeus dekanius Verhoeff, 1938 A, D NHMUK 015991474 B, C NHMUK 015991470 A metasternites of leg-bearing segments 79 80, ventral view B walking leg, pair 74, lateral view C Walking leg pair 74 pretarsus, lateral view D Leg-bearing segments 79 81, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure15" httpUri="https://binary.pensoft.net/fig/1257834">15</figureCitation>
,
<figureCitation id="5FC5CF73519136D6BF6ADB0383ECAA00" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Orphnaeus dekanius Verhoeff, 1938 A, C, D NHMUK 015991471 B, E NHMUK 015991470 A, B ultimate leg-bearing and postpedal segments, ventral view C ultimate leg pair metatarsus, lateral view D female gonopods, ventral view E male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure16" httpUri="https://binary.pensoft.net/fig/1257835">16</figureCitation>
,
<figureCitation id="11147CAAD12C2328753CB4B813D04A97" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Female gonopods of Orphnaeus Meinert, 1870 species A, B, C Orphnaeus brevilabiatus (Newport, 1845). (A = NHMUK 015991423 (Myanmar), B = NHMUK 015991421 (Thailand), C = NHMUK 015991420 (Singapore )) D, E, F, G, H Orphnaeus dekanius Verhoeff, 1938. (D = NHMUK 015991415 (Sri Lanka), E = NHMUK 015991413 (Sri Lanka), F = NHMUK 015991417 (Singapore), G = NHMUK 015991418 (Singapore), H = NHMUK 015991416 (Sri Lanka )). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1228.143007.figure18" httpUri="https://binary.pensoft.net/fig/1257837">18 D H</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQOBI" type="material">
<paragraph id="538E7B7A0A633BEC413C6B029F743EDD">
<heading id="1FE7941C16C00EB55DD4668C4AF92C93" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="55DBE174226196AFD05870E7CE67D889">
<specimenCount id="85B195E3099E428F27692DB9BE11B964" count="6" type="generic">6 specimens</specimenCount>
:
<materialsCitation id="E40EF9379CB14A931784618E695F1A53" collectingDate="1974-10-08" collectionCode="NHMUK" location="Picard" municipality="Aldabra" specimenCode="NHMUK 015991469" specimenCount="1" specimenCount-male="1">
<specimenCode id="456885F59E6CDA62ACCBA5A3925AB80B">NHMUK 015991469</specimenCode>
,
<specimenCount id="671D26FF0C6FBF98EFC91151555E1D06" count="1" type="male">1 ♂</specimenCount>
,
<quantity id="5C2381CAAC9CB15601989DC372274341" metricMagnitude="-2" metricUnit="m" metricValue="4.0" unit="mm" value="40.0">40 mm</quantity>
, 73 leg-bearing segments,
<location id="3CE84A522858D33122D99782ACAD2ED7" LSID="urn:lsid:plazi:treatment:2D319CB7E06954D08FCB7D9035EF2638:3CE84A522858D33122D99782ACAD2ED7" municipality="Aldabra" name="Picard">Picard</location>
,
<collectingMunicipality id="62E6AA504487CD150CE0FF73D59EC98B">Aldabra</collectingMunicipality>
,
<date id="822DA82AA7992348A0D013E296E4BDAB" value="1974-10-08">
<collectingDate id="57602AFEB2C750E1766FF98DEFB6E0DD" value="1974-10-08">08. 10. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="70D85829C15869540A28292C891C6EA1" collectingDate="1974-04-09" collectionCode="NHMUK" location="Picard" specimenCode="NHMUK 015991470" specimenCount="1" specimenCount-male="1">
<specimenCode id="61AE663A82C39BF129BE2E85829CB2BA">NHMUK 015991470</specimenCode>
,
<specimenCount id="39A1CC1B58DAA74BD5DDCD3EB0EE6339" count="1" type="male">1 ♂</specimenCount>
,
<quantity id="45D667A7A2E103973AB9C761BBE5F230" metricMagnitude="-2" metricUnit="m" metricValue="3.5" unit="mm" value="35.0">35 mm</quantity>
, 75 leg-bearing segments,
<location id="6B8CFD8891FB6BF4CC6D9407DF35A428" LSID="urn:lsid:plazi:treatment:2D319CB7E06954D08FCB7D9035EF2638:6B8CFD8891FB6BF4CC6D9407DF35A428" name="Picard">Picard</location>
,
<date id="8BADB9201E4E18B8F9154222F3CC8733" value="1974-04-09">
<collectingDate id="6E4699407002CD76A0E01D56E3934A08" value="1974-04-09">09. 04. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="B06A75F9DD1855FCDA92E5EB4F136679" collectionCode="NHMUK" collectorName="J. Wilson" location="Grande Terre" municipality="Aldabra" specimenCode="NHMUK 015991471" specimenCount="1" specimenCount-female="1">
<specimenCode id="4A80D369FDD77AF227AAF9816ADC5E6A">NHMUK 015991471</specimenCode>
,
<specimenCount id="5ACB66AF5A04A203CF21C8D08E191253" count="1" type="female">1 ♀</specimenCount>
,
<quantity id="3A7EAD7B7D80F8401C73B7D5A59BE5EC" metricMagnitude="-2" metricUnit="m" metricValue="2.4" unit="mm" value="24.0">24 mm</quantity>
, 81 leg-bearing segments,
<location id="42E5DE464DB130AFB546A77DC0B832BC" LSID="urn:lsid:plazi:treatment:2D319CB7E06954D08FCB7D9035EF2638:42E5DE464DB130AFB546A77DC0B832BC" municipality="Aldabra" name="Grande Terre">Grande Terre</location>
,
<collectingMunicipality id="4FD36F447FD4FAE5E679490DBC0CBACF">Aldabra</collectingMunicipality>
, 03.1974, leg.
<collectorName id="B59526106823620934B4E8E6B2B8BDDD">J. Wilson</collectorName>
</materialsCitation>
;
<materialsCitation id="336F59EAF13B86A09210DDD8196CBF3B" collectionCode="NHMUK" collectorName="J. Wilson" location="Grande Terre" municipality="Aldabra" specimenCode="NHMUK 015991472" specimenCount="1" specimenCount-juvenile="1">
<specimenCode id="693634B72BD7CCA436F7A97C287EDE77">NHMUK 015991472</specimenCode>
,
<specimenCount id="29AA5941C6A46365A15F74F79D3BB3E2" count="1" type="juvenile">1 juvenile</specimenCount>
,
<quantity id="18E2102CD45ED87215D3D6EEB8B2DD0C" metricMagnitude="-2" metricUnit="m" metricValue="1.3" unit="mm" value="13.0">13 mm</quantity>
, 81 leg-bearing segments,
<location id="E1D811EF5BD9BBE3137B1042030B4A87" LSID="urn:lsid:plazi:treatment:2D319CB7E06954D08FCB7D9035EF2638:E1D811EF5BD9BBE3137B1042030B4A87" municipality="Aldabra" name="Grande Terre">Grande Terre</location>
,
<collectingMunicipality id="D71736E1F4734A7DA229724E3BABCB20">Aldabra</collectingMunicipality>
, 05.1974, leg.
<collectorName id="5884AC82249AFC2AE6C868641A68CDD3">J. Wilson</collectorName>
</materialsCitation>
;
<materialsCitation id="F2BFAF0076E5D98E49172FFEAA6830C4" collectedFrom="Pandanus litter" collectingDate="1974-03-22" collectionCode="NHMUK" location="Aldabra" specimenCode="NHMUK 015991473" specimenCount="1" specimenCount-male="1">
<specimenCode id="C09E1B02329B997A6BCDCDD11C73CFC3">NHMUK 015991473</specimenCode>
,
<specimenCount id="047EA8E6D255CF9510FD8E3CA3735A09" count="1" type="male">1 ♂</specimenCount>
,
<quantity id="0F9DB8502B78A40EDF9AA3EE56039771" metricMagnitude="-2" metricUnit="m" metricValue="3.7" unit="mm" value="37.0">37 mm</quantity>
, 75 leg-bearing segments,
<collectedFrom id="8C4E9FD03622E7187D609B22DBB0A587">
<taxonomicName id="AE52A69AB3CB9C87230160FEC93200BF" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" phylum="Tracheophyta" rank="genus">
<emphasis id="936CF4305FB10D490F021C06A80D1035" italics="true">Pandanus</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="797E1FAA49530990ED6EEABE44B459ED" LSID="urn:lsid:plazi:treatment:2D319CB7E06954D08FCB7D9035EF2638:797E1FAA49530990ED6EEABE44B459ED" name="Aldabra">Aldabra</location>
,
<date id="52545BA855518B4966F37AA982822612" value="1974-03-22">
<collectingDate id="2B43546D878BB15E4CE8F5F38148B870" value="1974-03-22">22. 03. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="7EEA66BBF20BF2B9EE3D4EA2AB9A5EFA" collectingDate="1968-02-23" collectingDateMax="1968-02-27" collectingDateMin="1968-02-23" collectionCode="NHMUK" collectorName="B. Cogan &amp; A. Hutson" location="Takamaka" specimenCode="NHMUK 015991474" specimenCount="1" specimenCount-female="1" stateProvince="Anse Takamaka">
<specimenCode id="11CAD6D2995713823FD130FF370F2D74">NHMUK 015991474</specimenCode>
,
<specimenCount id="1A7C6BD30E630C5AFA81ED19DECF7DB8" count="1" type="female">1 ♀</specimenCount>
,
<quantity id="48048D6B3B79545C8BB4899CA7A77425" metricMagnitude="-2" metricUnit="m" metricValue="5.1" unit="mm" value="51.0">51 mm</quantity>
,
<location id="3EE943FEE09639A83B67B3EE574312A6" LSID="urn:lsid:plazi:treatment:2D319CB7E06954D08FCB7D9035EF2638:3EE943FEE09639A83B67B3EE574312A6" name="Takamaka" stateProvince="Anse Takamaka">Takamaka</location>
(
<collectingRegion id="6A6ECDBB3535BFBDD65A35419F8ADC8D">Anse Takamaka</collectingRegion>
),
<date id="307FCE2C71541634DEF7CD88AACD92BB" value="1968-02-23" valueMax="1968-02-27" valueMin="1968-02-23">
<collectingDate id="D72FCE06842A90B89FF4A839FF39AABC" value="1968-02-23" valueMax="1968-02-27" valueMin="1968-02-23">2327.02. 1968</collectingDate>
</date>
, leg.
<collectorName id="40EFB65CF826ADD8114BFF57BFA122AD">B. Cogan &amp; A. Hutson</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="3E9A570BB1FF0B9EF9F3ED4EC03D1BAF">
<emphasis id="385433FCB837C1B5CD2CB538BEA72B1F" bold="true">Diagnosis.</emphasis>
Medium to large size
<taxonomicName id="2160A02D504C9CEA9391257613FD5361">
<emphasis id="4B9C6D63120C7D8B79CE16E6269A91D5" italics="true">Orphnaeus</emphasis>
species
</taxonomicName>
, with 7381 leg-bearing segments and variable but generally present longitudinal bands of dark pigment flanking the central vessel. Mandible with three or four pectinate lamellae. First maxillae with both telopodal and coxosternal lappets present and uniarticulate telopodite. Second maxillary pretarsus spatulate, fringed by acuminate hyaline projections. Posterior trunk metasternites with paired pore fields at the posterior end. Pore fields on posterior metasternites, procoxae and metacoxae bordered by dense groups of setae-like projections. Female gonopods uniarticulate, medially overlapping, with angled, rounded external margin.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIPBI" type="description">
<paragraph id="9E437851FE5E419161839FBAF620B906">
<heading id="33DC00C27F508830DFBDF6267712C20A" reason="title">Description.</heading>
</paragraph>
<paragraph id="D30EB7BDB1817DDE2748D9EE60A2A8BC">
<emphasis id="7BA317EEE61DB8B08B7B64E52A58B146" bold="true">
<emphasis id="968314CC50B9E06A0C390CB9242041CE" bold="true" italics="true">Head and antennae</emphasis>
.
</emphasis>
Cephalic plate with broadly rounded anterior margin and straight posterior margin, overlapping the forcipular tergite. Head approximately as broad as long (
<specimenCode id="5D4A088739422B774AB5FE866EF567E4">
<collectionCode id="BC0D7B2C5027BF08E9D1C3D4CC4E7D89" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991473
</specimenCode>
) to 1.2 × broader than long (
<specimenCode id="A18A751BE828CCEBB0AA936DF54216A2">
<collectionCode id="57D1FB48798864F5D678BBC845C67D5E" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991474
</specimenCode>
). Antennae approximately 2.5 × longer than head, weakly tapering distally (Fig.
<figureCitation id="CE5199ACC8F7F7C7D0A4F39CE7CC7EFE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orphnaeus dekanius Verhoeff, 1938 A, B, D NHMUK 015991470 A, C NHMUK 015991474. Head and antennae, dorsal view B antennal article XIV, dorsal view D forcipular segment, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure12" httpUri="https://binary.pensoft.net/fig/1257831">12 A</figureCitation>
). In older specimens, the tapering of antennae and dorsoventral compression of proximal antennal articles are more clearly visible (Fig.
<figureCitation id="6CCBB71FE99566BBED94BC0FE8C369DE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orphnaeus dekanius Verhoeff, 1938 A, B, D NHMUK 015991470 A, C NHMUK 015991474. Head and antennae, dorsal view B antennal article XIV, dorsal view D forcipular segment, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure12" httpUri="https://binary.pensoft.net/fig/1257831">12 C</figureCitation>
). Antennal article XIV 1.9 × longer than the penultimate, with two clusters of sensilla basiconica arranged in lateral pits. Small, spear-like sensilla present at apical end.
</paragraph>
<caption id="802950A231F9A9D0D93CC5CC9D798464" ID-DOI="10.3897/zookeys.1228.143007.figure12" ID-arpha="790B73DB-FD81-56AF-B1F9-D12B61B681D8" httpUri="https://binary.pensoft.net/fig/1257831" startId="F12">
<paragraph id="0493E90C0EC46447E19EE804616E57A8">
<label id="89C0D437D14308D9DBB84091F20FC5F6">Figure 12.</label>
</paragraph>
<paragraph id="5C982F1EAAB294EC5DD45C1EA7605643">
<taxonomicName id="6E6A6246C098AA2A7929689B717AF7B2" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="3214B8A0EAB2D75D9DBA7BA67DC4C4A9" italics="true">Orphnaeus dekanius</emphasis>
<bibRefCitation id="0BE90588C36D43FBAA9F83D69E4D6697" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbücher, Abteilung für Systematik" pagination="339-388" refId="B107" refString="Verhoeff KW (1938) Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik 71: 339-388." title="Chilopoden-Studien zur Kenntnis der Epimorphen." volume="71" year="1938">Verhoeff, 1938</bibRefCitation>
</taxonomicName>
<emphasis id="CF7CE0AA0DFBE76C80CFA33645885E2F" bold="true">A, B, D</emphasis>
<specimenCode id="EC586F482084CA4C5DFD397EFF38857C">NHMUK 015991470</specimenCode>
<emphasis id="18550F9A801599104AE00A748D9E8783" bold="true">A, C</emphasis>
<specimenCode id="A0A596A3A0309D69DCFB0E897AA13DAA">NHMUK 015991474</specimenCode>
. Head and antennae, dorsal view
<emphasis id="1E48E925EC419E388B7266DAA8A0902D" bold="true">B</emphasis>
antennal article XIV, dorsal view
<emphasis id="2AD9E8B4839FD5F22D864C9DB8931CE5" bold="true">D</emphasis>
forcipular segment, ventral view.
</paragraph>
</caption>
<paragraph id="B448C7150C4BC790888C419BF36905A3">
<emphasis id="37630078E7AA73247C22E210259466FD" bold="true">
<emphasis id="AAE2632E410AD07424F512FD809F32F2" bold="true" italics="true">Mandibles</emphasis>
.
</emphasis>
Of typical aspect for the genus (Fig.
<figureCitation id="9319173B6B1784B9BF68E98C17578652" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Orphnaeus dekanius Verhoeff, 1938. NHMUK 015991470 A clypeus and labrum, ventral view B labrum, ventral view C, D Mandible, lateral and dorsal views." figureDoi="10.3897/zookeys.1228.143007.figure13" httpUri="https://binary.pensoft.net/fig/1257832">13 C, D</figureCitation>
). Four conspicuous pectinate lamellae evident, arranged concentrically around distal edge. Proximal to the outermost lamella, isolated projections resembling those on the lamellae are present. Two minute sensilla present laterally.
</paragraph>
<caption id="9FAD26AC24003AFD9AA0B8C5F20BFE05" ID-DOI="10.3897/zookeys.1228.143007.figure13" ID-arpha="36D0760B-C803-5C37-A128-DB8E7A7DB780" httpUri="https://binary.pensoft.net/fig/1257832" startId="F13">
<paragraph id="9456D616AD3A1ECA3ED15EBCC75F2EE9">
<label id="803570E2B945E55CF67A4DDFB0664924">Figure 13.</label>
</paragraph>
<paragraph id="72F8EAA0D0EB9FFC74885783FB6EBFBE">
<taxonomicName id="5C47819F0AC39394603ACDD824E6DB39" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="334146B409E9CD5F35392C5B7F87C1E0" italics="true">Orphnaeus dekanius</emphasis>
<bibRefCitation id="54A46277F1269BE2695FE69677C4B304" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbücher, Abteilung für Systematik" pagination="339-388" refId="B107" refString="Verhoeff KW (1938) Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik 71: 339-388." title="Chilopoden-Studien zur Kenntnis der Epimorphen." volume="71" year="1938">Verhoeff, 1938</bibRefCitation>
</taxonomicName>
.
<specimenCode id="48BC4B160E229A786D6ECC38CFCA9F6C">NHMUK 015991470</specimenCode>
<emphasis id="84D60A93CB5D94AF2A2E2BC4BA7C4BA6" bold="true">A</emphasis>
clypeus and labrum, ventral view
<emphasis id="6DC06DB2E413A93A94010C32388FD6F8" bold="true">B</emphasis>
labrum, ventral view
<emphasis id="EE3B9312282720F4AC1C9894EF050476" bold="true">C, D</emphasis>
Mandible, lateral and dorsal views.
</paragraph>
</caption>
<paragraph id="5D961D7FD3E0AF1A2BED58E1FB083345">
<emphasis id="EE2FB674F74B52B8F3803886AE83695E" bold="true">
<emphasis id="B01B08CAC577BE9C6E89350799D84CC4" bold="true" italics="true">Labrum and clypeus</emphasis>
.
</emphasis>
Labrum of typical oryid aspect, with short hairlike hyaline projections on middle part (Fig.
<figureCitation id="0750AFD24945B12D22A65B4E064A87ED" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Orphnaeus dekanius Verhoeff, 1938. NHMUK 015991470 A clypeus and labrum, ventral view B labrum, ventral view C, D Mandible, lateral and dorsal views." figureDoi="10.3897/zookeys.1228.143007.figure13" httpUri="https://binary.pensoft.net/fig/1257832">13 B</figureCitation>
). Lateral parts incompletely separated from middle part and clypeus by evident sutures. Clypeus with two pairs of postantennal sensilla, a median field of sensilla spanning its mediolateral axis and one pair of prelabral sensilla posterior to these (Fig.
<figureCitation id="A4AC68FF4673ED733F63BD80162C53DA" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Orphnaeus dekanius Verhoeff, 1938. NHMUK 015991470 A clypeus and labrum, ventral view B labrum, ventral view C, D Mandible, lateral and dorsal views." figureDoi="10.3897/zookeys.1228.143007.figure13" httpUri="https://binary.pensoft.net/fig/1257832">13 A</figureCitation>
). Polygonal reticulation evident.
</paragraph>
<paragraph id="F0E40B3D153BC92454F173B096ECC218">
<emphasis id="95DFDC8762A61D3B91FB30C7706F1ADA" bold="true">
<emphasis id="28E3AA30CC2500EB10D8B86D95212749" bold="true" italics="true">Maxillae</emphasis>
.
</emphasis>
First maxillae with apically rounded, short coxal projections, bearing 811 sensilla (Fig.
<figureCitation id="EEEB07F0F337E6EBA61CBC7DD88ECBD5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Orphnaeus dekanius Verhoeff, 1938 NHMUK 015991470 A first and second maxillae, ventral view B first maxillary right telopodal lappet, ventral view C first maxillary left telopodal and coxosternal lappets, dorsal view D, E left and right second maxillary pretarsi, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure14" httpUri="https://binary.pensoft.net/fig/1257833">14 A</figureCitation>
). Telopodite broadly rounded, uniarticulated, of similar size to the coxal projection, bearing 712 sensilla. Both telopodal and coxosternal pairs of lappets present, with distinct spinous reticulation (Fig.
<figureCitation id="D5362C026F4FCEF9C488DA0700006252" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Orphnaeus dekanius Verhoeff, 1938 NHMUK 015991470 A first and second maxillae, ventral view B first maxillary right telopodal lappet, ventral view C first maxillary left telopodal and coxosternal lappets, dorsal view D, E left and right second maxillary pretarsi, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure14" httpUri="https://binary.pensoft.net/fig/1257833">14 C</figureCitation>
). Coxosternal pair of lappets completely obscured by second maxillary telopodite in ventral view. Second maxillary coxosternite with shallow, concave, rounded anterior margin bordered by a row of trichoid sensilla and two groups of trichoid sensilla proximally (Fig.
<figureCitation id="924912C7A069C459D391CF6B908C6E62" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Orphnaeus dekanius Verhoeff, 1938 NHMUK 015991470 A first and second maxillae, ventral view B first maxillary right telopodal lappet, ventral view C first maxillary left telopodal and coxosternal lappets, dorsal view D, E left and right second maxillary pretarsi, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure14" httpUri="https://binary.pensoft.net/fig/1257833">14 A</figureCitation>
). Metameric pore conspicuous, surrounded by sclerotised rim. Telopodal articles stout. Pretarsus with spine-like projections around entire exterior margin and two pores opening on its dorsal surface (Fig.
<figureCitation id="16DCB2A403ABB9D423EED654AD289B61" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Orphnaeus dekanius Verhoeff, 1938 NHMUK 015991470 A first and second maxillae, ventral view B first maxillary right telopodal lappet, ventral view C first maxillary left telopodal and coxosternal lappets, dorsal view D, E left and right second maxillary pretarsi, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure14" httpUri="https://binary.pensoft.net/fig/1257833">14 D, E</figureCitation>
).
</paragraph>
<caption id="4A0006EFF4007CAAF0ABAC70FA589FB1" ID-DOI="10.3897/zookeys.1228.143007.figure14" ID-arpha="9BC505BB-5928-518E-BD3A-4ADC5BB254AB" httpUri="https://binary.pensoft.net/fig/1257833" startId="F14">
<paragraph id="4BBCED98C89057EF6062CFA9AD075E50">
<label id="F1CCF68A6030F990707773603715D9DC">Figure 14.</label>
</paragraph>
<paragraph id="1438942D5F7543B5FB21D740BBD528E5">
<taxonomicName id="E7F1765B1CDCD3B3C329F6161125EB0A" authority="Verhoeff, 1938 NHMUK" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="DBE604D4656C2938F7F1F1B86DDFCD72" italics="true">Orphnaeus dekanius</emphasis>
<bibRefCitation id="508A8CBC07B2E023897C99535DD7A155" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbücher, Abteilung für Systematik" pagination="339-388" refId="B107" refString="Verhoeff KW (1938) Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik 71: 339-388." title="Chilopoden-Studien zur Kenntnis der Epimorphen." volume="71" year="1938">Verhoeff, 1938</bibRefCitation>
<specimenCode id="7983AE3961D38D760F9E782ED9C78E78">NHMUK 015991470</specimenCode>
</taxonomicName>
<emphasis id="3952AAA7F75C13C89DDF3272ED86997D" bold="true">A</emphasis>
first and second maxillae, ventral view
<emphasis id="B981602BF774BDADA022C41C240346FB" bold="true">B</emphasis>
first maxillary right telopodal lappet, ventral view
<emphasis id="7D593C6FC089366B82CE2CCE1FF7EB9A" bold="true">C</emphasis>
first maxillary left telopodal and coxosternal lappets, dorsal view
<emphasis id="58980144220A5F1E9C43EA72AE12C0BB" bold="true">D, E</emphasis>
left and right second maxillary pretarsi, ventral view.
</paragraph>
</caption>
<paragraph id="7CF0B1163923FE1DDCC50D7F9C6AD391">
<emphasis id="566E4F7C2FDFB36B7370CE392607BC8E" bold="true">
<emphasis id="DAE9126CFD955CFEF923E87FD44FEB37" bold="true" italics="true">Forcipular segment</emphasis>
.
</emphasis>
Forcipular tergite 3.2 × broader than long. Exposed surface of forcipular coxosternite 2.2 × broader than long (Fig.
<figureCitation id="6F380A499FAE51BDAACD5D44A0330A10" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Orphnaeus dekanius Verhoeff, 1938 A, B, D NHMUK 015991470 A, C NHMUK 015991474. Head and antennae, dorsal view B antennal article XIV, dorsal view D forcipular segment, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure12" httpUri="https://binary.pensoft.net/fig/1257831">12 D</figureCitation>
). Chitin lines absent. Anterior margin rounded, deeply concave. Pleural sutures strongly converging posteriorly. Trochanteroprefemur ~ 1.5 × broader than long, with evidently rounded external face. Tarsungulum stout, large, entirely covered by anterior edge of cephalic plate, with smooth inner concavity. Opening of venom gland channel immediately proximal to tip of tarsungulum. All forcipular articles without denticles.
</paragraph>
<paragraph id="620F245CBDF20A0BBFC0364E4BB68CFA">
<emphasis id="4A1F4AD247C7A5EB48EED6E607EAC84B" bold="true">
<emphasis id="E94B9DEA8A631CD05721BDE010A09B08" bold="true" italics="true">Trunk</emphasis>
.
</emphasis>
Last five or six trunk metasternites with two posteriorly located pore fields (Fig.
<figureCitation id="73F557A18142F7BF8360D07894B8CE32" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Orphnaeus dekanius Verhoeff, 1938 A, D NHMUK 015991474 B, C NHMUK 015991470 A metasternites of leg-bearing segments 79 80, ventral view B walking leg, pair 74, lateral view C Walking leg pair 74 pretarsus, lateral view D Leg-bearing segments 79 81, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure15" httpUri="https://binary.pensoft.net/fig/1257834">15 D</figureCitation>
), anteriorly bordered by dense clusters of hairs (Fig.
<figureCitation id="E2A50A63EE9E836AB1AC48A8EDCD2BB1" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Orphnaeus dekanius Verhoeff, 1938 A, D NHMUK 015991474 B, C NHMUK 015991470 A metasternites of leg-bearing segments 79 80, ventral view B walking leg, pair 74, lateral view C Walking leg pair 74 pretarsus, lateral view D Leg-bearing segments 79 81, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure15" httpUri="https://binary.pensoft.net/fig/1257834">15 A</figureCitation>
). All other trunk metasternites with four pore fields, two anterior and two posterior, of equal size on metasternites 246, the anterior gradually decreasing in size until disappearing on metasternites 7476. One single row of paratergites present beginning from the second leg-bearing segment, becoming very conspicuous on the eighth leg-bearing segment. General setation of sclerites sparse.
</paragraph>
<caption id="3B403ED87CE3529AB9AE31D288E0EA9B" ID-DOI="10.3897/zookeys.1228.143007.figure15" ID-arpha="14CD1EC9-C155-50AC-A23C-59AE448B092E" httpUri="https://binary.pensoft.net/fig/1257834" startId="F15">
<paragraph id="5BF892787751C67EBFB0C43E37422E2F">
<label id="6ABC4CA5E51AE2D452149123998772FD">Figure 15.</label>
</paragraph>
<paragraph id="EDF39883746A6FD4A0DFDE65702D80E1">
<taxonomicName id="AB9C77A802F08B2CD483CA9E84ED4A6C" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="8C9FBF0479417BA2C4621548B8EF9E57" italics="true">Orphnaeus dekanius</emphasis>
<bibRefCitation id="03CAD634A22E3666B3224C4E8F6B9952" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbücher, Abteilung für Systematik" pagination="339-388" refId="B107" refString="Verhoeff KW (1938) Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik 71: 339-388." title="Chilopoden-Studien zur Kenntnis der Epimorphen." volume="71" year="1938">Verhoeff, 1938</bibRefCitation>
</taxonomicName>
<emphasis id="4FDC639250DBFC207AC7A0212D79D23C" bold="true">A, D</emphasis>
<specimenCode id="ACFF6CAF3CBAAE8D2E81F382F41FA30F">NHMUK 015991474</specimenCode>
<emphasis id="A2B46FCC84C126F98212E1029F54EC0E" bold="true">B, C</emphasis>
<specimenCode id="D4733929B45835040FB8F9F1570EA73F">NHMUK 015991470</specimenCode>
<emphasis id="B7644F9C6B0ED547C98195B1AADE3767" bold="true">A</emphasis>
metasternites of leg-bearing segments 79 80, ventral view
<emphasis id="0359D6587A3FC01F499AB9809D0394D2" bold="true">B</emphasis>
walking leg, pair 74, lateral view
<emphasis id="03B6831ECF4FE30DC8BA10B9680B5F33" bold="true">C</emphasis>
Walking leg pair 74 pretarsus, lateral view
<emphasis id="F374787BECC3D0E5541E63F98F00BBC7" bold="true">D</emphasis>
Leg-bearing segments 79 81, ventral view.
</paragraph>
</caption>
<paragraph id="E7642D290DF625884D299BBB142D7C64">
<emphasis id="DFF296FCEE18A2FE069135A17820F51D" bold="true">
<emphasis id="4A8B6B8986EA6B6755A197B23845BF51" bold="true" italics="true">Ultimate leg</emphasis>
-
<emphasis id="B231B522C3AAAB80CB4920B342FD6A8A" bold="true" italics="true">bearing and postpedal segments</emphasis>
.
</emphasis>
Ultimate leg-bearing segment metasternite variably trapeziform, 2.3 × broader than long (Fig.
<figureCitation id="53314C9859CECAD6FF0E14E5B03BAF52" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Orphnaeus dekanius Verhoeff, 1938 A, C, D NHMUK 015991471 B, E NHMUK 015991470 A, B ultimate leg-bearing and postpedal segments, ventral view C ultimate leg pair metatarsus, lateral view D female gonopods, ventral view E male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure16" httpUri="https://binary.pensoft.net/fig/1257835">16 A, B</figureCitation>
). Posterior edge with dense field of hairs and occasionally small clusters of pores. Coxopleuron stout, without coxal organs. Telopodite of ultimate leg-pair only moderately inflated in both males (Fig.
<figureCitation id="406FABF4AAC5CC2BFDA8BCA2B7C503B8" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Orphnaeus dekanius Verhoeff, 1938 A, C, D NHMUK 015991471 B, E NHMUK 015991470 A, B ultimate leg-bearing and postpedal segments, ventral view C ultimate leg pair metatarsus, lateral view D female gonopods, ventral view E male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure16" httpUri="https://binary.pensoft.net/fig/1257835">16 B</figureCitation>
) and females (Fig.
<figureCitation id="DD0A6F7205FB852438AF8E51B8060F07" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Orphnaeus dekanius Verhoeff, 1938 A, C, D NHMUK 015991471 B, E NHMUK 015991470 A, B ultimate leg-bearing and postpedal segments, ventral view C ultimate leg pair metatarsus, lateral view D female gonopods, ventral view E male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure16" httpUri="https://binary.pensoft.net/fig/1257835">16 A</figureCitation>
), with dense fields of hairs present on the ventral side of all articles. Pretarsus absent (Fig.
<figureCitation id="7752AB16418474DD09A53AC6DD1ED781" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Orphnaeus dekanius Verhoeff, 1938 A, C, D NHMUK 015991471 B, E NHMUK 015991470 A, B ultimate leg-bearing and postpedal segments, ventral view C ultimate leg pair metatarsus, lateral view D female gonopods, ventral view E male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure16" httpUri="https://binary.pensoft.net/fig/1257835">16 C</figureCitation>
). Metatarsus with small hair-like projections at its apical edge.
</paragraph>
<caption id="8B30A6CECB290E60A40699209175CD3B" ID-DOI="10.3897/zookeys.1228.143007.figure16" ID-arpha="D779F08E-0CDD-598A-8CF2-C4C014BF3CE8" httpUri="https://binary.pensoft.net/fig/1257835" startId="F16">
<paragraph id="8FEC5E96D75900D59FB91C46E86E880E">
<label id="3A75EB7310A30485719247480948262A">Figure 16.</label>
</paragraph>
<paragraph id="0F672D3EC2FD0DEB6735C2AD702D9DE8">
<taxonomicName id="F92E59D568E3DE9FE9C54E15874B6B2E" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="0ED94EBFC3233D075AE186E5FC9EB034" italics="true">Orphnaeus dekanius</emphasis>
<bibRefCitation id="24DA99B57E39CD78615D1C25FB1E472F" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbücher, Abteilung für Systematik" pagination="339-388" refId="B107" refString="Verhoeff KW (1938) Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik 71: 339-388." title="Chilopoden-Studien zur Kenntnis der Epimorphen." volume="71" year="1938">Verhoeff, 1938</bibRefCitation>
</taxonomicName>
<emphasis id="DDD21FE8C4D8CF4BAB9A53DF7FC75038" bold="true">A, C, D</emphasis>
<specimenCode id="EF033BF8AB83F989AA83DD154E7EB51C">NHMUK 015991471</specimenCode>
<emphasis id="A206788B0661FB2575145497C512C6AE" bold="true">B, E</emphasis>
<specimenCode id="57AEFAE196719DC322655E70B73B6A7E">NHMUK 015991470</specimenCode>
<emphasis id="4A79E98BC45B32CAC1047ABE13EF27DF" bold="true">A, B</emphasis>
ultimate leg-bearing and postpedal segments, ventral view
<emphasis id="6F29605E1E7662C59FAE321831B728A3" bold="true">C</emphasis>
ultimate leg pair metatarsus, lateral view
<emphasis id="870E4B1A20952008B05DBFD8B383BEE3" bold="true">D</emphasis>
female gonopods, ventral view
<emphasis id="E7DB30A3473C7E38BBBB9C8BD570CA7E" bold="true">E</emphasis>
male gonopods, ventral view.
</paragraph>
</caption>
<paragraph id="3668B3C297EAE62EEF7DB89F82C095B5">
Female gonopods usually uni-articulated (Figs
<figureCitation id="909C1D0603524B54ECD7AB6C620A54C0" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Orphnaeus dekanius Verhoeff, 1938 A, D NHMUK 015991474 B, C NHMUK 015991470 A metasternites of leg-bearing segments 79 80, ventral view B walking leg, pair 74, lateral view C Walking leg pair 74 pretarsus, lateral view D Leg-bearing segments 79 81, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure15" httpUri="https://binary.pensoft.net/fig/1257834">15 D</figureCitation>
,
<figureCitation id="285FBA756C88931700FE78FBF61DA095" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Orphnaeus dekanius Verhoeff, 1938 A, C, D NHMUK 015991471 B, E NHMUK 015991470 A, B ultimate leg-bearing and postpedal segments, ventral view C ultimate leg pair metatarsus, lateral view D female gonopods, ventral view E male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure16" httpUri="https://binary.pensoft.net/fig/1257835">16 D</figureCitation>
), occasionally with an anterior notch or asymmetrical articulation. Lateral edge smooth, rounded, forming strongly acute angle with posterior edge of first genital sternite. Male gonopods biarticulated, bearing 16 setae (Fig.
<figureCitation id="FEB58A2C73145B9C11F722688C6A5DE4" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Orphnaeus dekanius Verhoeff, 1938 A, C, D NHMUK 015991471 B, E NHMUK 015991470 A, B ultimate leg-bearing and postpedal segments, ventral view C ultimate leg pair metatarsus, lateral view D female gonopods, ventral view E male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure16" httpUri="https://binary.pensoft.net/fig/1257835">16 E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETYBI" type="remarks">
<paragraph id="6A3C478C63DB8A1F97704B139063F122">
<heading id="0A4CF99B0518992182CABDBCC2A7C453" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="244DD16DA058E88F5826D5EB813288FB">
<taxonomicName id="803153F97389B4CE922270CE6B9A8D7D" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="F8D3AAAFE6482A04F0484B1D25D05359" italics="true">Orphnaeus dekanius</emphasis>
</taxonomicName>
was originally described from Trivandrum (Thiruvanathapuram),
<collectingCountry id="47AF48069DF56B33BCFC0454C16DA680" name="India">India</collectingCountry>
(
<bibRefCitation id="FA802DF1F8DD5932B563B4A4A9EB7F2C" author="Verhoeff" firstAuthor="Verhoeff" journalOrPublisher="Zoologische Jahrbücher, Abteilung für Systematik" pagination="339-388" refId="B107" refString="Verhoeff KW (1938) Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbücher, Abteilung für Systematik 71: 339388." title="Chilopoden-Studien zur Kenntnis der Epimorphen." volume="71" year="1938">Verhoeff 1938</bibRefCitation>
), and maintained as a valid species under
<taxonomicName id="F761E500A45E3C5A76C2E97DAE37C48A" authorityName="Meinert" authorityYear="1870" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="8BDF0B4E135A6CC61803D8B39017C8DE" italics="true">Orphnaeus</emphasis>
</taxonomicName>
until its reassignment to
<taxonomicName id="6C0BF02973738320FCC16A9440218096" authorityName="Crabill" authorityYear="1959" class="Chilopoda" family="Oryidae" genus="Nycternyssa" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="D02E153077DC622882CAFA375A54F3D2" italics="true">Nycternyssa</emphasis>
</taxonomicName>
, justified by the description of the female gonopods as “ uni-articulate ”. A detailed re-evaluation of the status of
<taxonomicName id="407CF25C03CFF569CDCD58ED95477806" authorityName="Crabill" authorityYear="1959" class="Chilopoda" family="Oryidae" genus="Nycternyssa" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="4942645CDBBAA654C92D0016DF2A44CC" italics="true">Nycternyssa</emphasis>
</taxonomicName>
is provided below.
</paragraph>
<paragraph id="443F699B1A9952A7D12C9E718C243B26">
Subsequent to its original description, there is no evidence that other specimens had been assigned to either
<taxonomicName id="C32CDF8508BB214CB10C02D99863AA8A" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="A6107121E3F72337C48F61517AFEB6E0" italics="true">O. dekanius</emphasis>
</taxonomicName>
or
<taxonomicName id="1C71EC062B7D4681992FC27005C99814" authorityName="Verhoeff" authorityYear="1937" class="Chilopoda" family="Oryidae" genus="Nycternyssa" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekania">
<emphasis id="1DD0F0D7ED4ADC4D8238639C33B2AB0E" italics="true">N. dekania</emphasis>
</taxonomicName>
prior to recent records from the Chagos Archipelago (
<bibRefCitation id="D55875E94F7EE11D988A8A770F4DE2D2" DOI="10.1080/00222933.2024.2395903" author="Popovici" etAl="et al." firstAuthor="Popovici" issue="41-44" journalOrPublisher="Journal of Natural History" pagination="1885-1915" refId="B87" refString="Popovici G, Edgecombe GD, Hall DW (2024) New Chilopoda from the Chagos Archipelago. Journal of Natural History 58 (4144): 18851915. https://doi.org/10.1080/00222933.2024.2395903" title="New Chilopoda from the Chagos Archipelago." volume="58" year="2024">Popovici et al. 2024</bibRefCitation>
). Based on morphology alone, the present specimens are considered conspecific with those collected in the Chagos Archipelago and match the original description of
<taxonomicName id="F9E22488D38279612C7F82686BF128C1" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="5002F014C63C0E86930A8F362353A000" italics="true">O. dekanius</emphasis>
</taxonomicName>
. The most salient diagnostic trait allowing for reliable differentiation of
<taxonomicName id="E9BD677AE211F9B5E18A2886993B4F2C" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="B8F3C5BA3AEBDD6C790CF4272858F3CE" italics="true">O. dekanius</emphasis>
</taxonomicName>
from
<taxonomicName id="7EE597A733FAF6B8215168E8AF8FCB7D" authorityName="Newport" authorityYear="1845" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="brevilabiatus">
<emphasis id="E57F5074CC4814A45B36A1AC62658E2D" italics="true">O. brevilabiatus</emphasis>
</taxonomicName>
is the presence of dense clusters of minute setae bordering the posterior pore fields of the former (Fig.
<figureCitation id="DCCF87B9816CB8029F870B3AE0F646FD" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Orphnaeus dekanius Verhoeff, 1938 A, D NHMUK 015991474 B, C NHMUK 015991470 A metasternites of leg-bearing segments 79 80, ventral view B walking leg, pair 74, lateral view C Walking leg pair 74 pretarsus, lateral view D Leg-bearing segments 79 81, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure15" httpUri="https://binary.pensoft.net/fig/1257834">15 A</figureCitation>
). Although this character was illustrated by Verhoeff in the original description of
<taxonomicName id="09DBF5C403F020513AED52DFFADFF6D2" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="972183DF2FB18EC09264916B9FA346B1" italics="true">O. dekanius</emphasis>
</taxonomicName>
(
<bibRefCitation id="83112B17E5C9B23CEFB4AE6C4AC2DFB4" author="Verhoeff" firstAuthor="Verhoeff" journalOrPublisher="Zoologische Jahrbücher, Abteilung für Systematik" pagination="339-388" refId="B107" refString="Verhoeff KW (1938) Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbücher, Abteilung für Systematik 71: 339388." title="Chilopoden-Studien zur Kenntnis der Epimorphen." volume="71" year="1938">Verhoeff 1938</bibRefCitation>
: tafel 8, fig. 61), no mention was made of it in the text of the description. It is unambiguously shared by all specimens studied here from near the type locality of
<taxonomicName id="742EAB471B3BC777501EDB58D6159A09" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="D283711C3A8A2DDB64C322082B97A5E9" italics="true">O. dekanius</emphasis>
</taxonomicName>
(specimens from
<collectingCountry id="78BB61B8F837B2071CBB8623DA681FB4" name="Sri Lanka">Sri Lanka</collectingCountry>
listed above under “ Specimen data ”), and from other localities in the Western Indian Ocean, and is consistent in both sexes in specimens. Setae are sparse in the smallest studied Aldabra specimen (
<quantity id="CA0B53CABFFB994CD62FCD947AA87EBF" metricMagnitude="-2" metricUnit="m" metricValue="1.3" unit="mm" value="13.0">13 mm</quantity>
) but are clustered by a body length of
<quantity id="8C3D7656C5A179513201330A55812D92" metricMagnitude="-2" metricUnit="m" metricValue="2.4" unit="mm" value="24.0">24 mm</quantity>
. As these clusters of setae are completely absent in specimens identified as
<taxonomicName id="ED9A45FEB1DCB6BA409EE4836A250B20" authorityName="Newport" authorityYear="1845" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="brevilabiatus">
<emphasis id="03B62279E8DC1B56E6C8DCBF55721798" italics="true">O. brevilabiatus</emphasis>
</taxonomicName>
from near its type locality in
<collectingCountry id="27F5C38A9696011BCCF83ABF3B60557E" name="Myanmar">Myanmar</collectingCountry>
(Fig.
<figureCitation id="9CA9DCBE080F28E6C05170C9A8DE0A56" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Orphnaeus brevilabiatus (Newport, 1845). NHMUK 015991423 A leg bearing segments 77 79, ventral view B metasternite of leg-bearing segment 78, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure17" httpUri="https://binary.pensoft.net/fig/1257836">17</figureCitation>
) and other localities in mainland and maritime Southeast Asia, we maintain the validity of
<taxonomicName id="664E0576F989BA1CD167F72C68EA2034" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="1EB562572B55E9FF88FD875FAE751193" italics="true">O. dekanius</emphasis>
</taxonomicName>
and
<taxonomicName id="EFDE4C86AAB6C8D14733013482BCC770" authorityName="Newport" authorityYear="1845" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="brevilabiatus">
<emphasis id="A27411A879B6760D3317F2DB6CA9E21C" italics="true">O. brevilabiatus</emphasis>
</taxonomicName>
even in light of the variability of female gonopodal articulation in the latter (Fig.
<figureCitation id="86AAF532CA43E3F95DB14263D381A3C2" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Female gonopods of Orphnaeus Meinert, 1870 species A, B, C Orphnaeus brevilabiatus (Newport, 1845). (A = NHMUK 015991423 (Myanmar), B = NHMUK 015991421 (Thailand), C = NHMUK 015991420 (Singapore )) D, E, F, G, H Orphnaeus dekanius Verhoeff, 1938. (D = NHMUK 015991415 (Sri Lanka), E = NHMUK 015991413 (Sri Lanka), F = NHMUK 015991417 (Singapore), G = NHMUK 015991418 (Singapore), H = NHMUK 015991416 (Sri Lanka )). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1228.143007.figure18" httpUri="https://binary.pensoft.net/fig/1257837">18 A C</figureCitation>
). The incomplete description of
<taxonomicName id="89CF41AC3F653C243AE49AD3DB4E5CB9" authority="Attems, 1909" authorityName="Attems" authorityYear="1909" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="meruinus">
<emphasis id="0EC325E7D0D5592BEB57D61514116F0E" italics="true">Orphnaeus meruinus</emphasis>
<bibRefCitation id="A46DC00B760B392530888B9A6314BF4F" author="Attems C" editor="Sjöstedt Y" journalOrPublisher="Palmquists, Stockholm" pagination="1-64" refId="B3" refString="Attems C (1909) Myriopoda. In: Sjostedt Y (Ed.) Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906. Palmquists, Stockholm 19: 1-64." title="Myriopoda." volume="19" volumeTitle="Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den umgebenden Massaisteppen Deutsch-Ostafrikas 1905-1906." year="1909">Attems, 1909</bibRefCitation>
</taxonomicName>
does not allow for reliable separation from
<taxonomicName id="479FC04CD608B9691288AEC0CCFFF30C" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="2294E57F30ECE932B14A041F87FA818D" italics="true">O. dekanius</emphasis>
</taxonomicName>
, given the inconsistencies in how diagnostic characters for these two taxa are coded in past literature. One salient difference from all
<taxonomicName id="09835685315095836D96FCD7AF091325" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="1924908A9D1FF39E16889FA7317A06F4" italics="true">O. dekanius</emphasis>
</taxonomicName>
specimens in our sample is the greatly inflated ultimate leg telopodite in males assigned to
<taxonomicName id="9AB6EECBFBACBC54A334E7B1DE13A45E" authorityName="Attems" authorityYear="1909" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="meruinus">
<emphasis id="8CB4608BB82F875A4E8E385CF07770F9" italics="true">O. meruinus</emphasis>
</taxonomicName>
collected in
<collectingCountry id="8AFD6330D85668E264D953C977FBA32C" name="Oman">Oman</collectingCountry>
(
<bibRefCitation id="3217F0462490D183EAEA3CC6C0F952A0" author="Lewis and Gallagher" firstAuthor="Lewis" journalOrPublisher="Fauna of Saudi Arabia" pagination="55-62" refId="B63" refString="Lewis JGE, Gallagher MD (1993) Scolopendromorph and Geophilomorph centipedes from Oman and the United Arab Emirates. Fauna of Saudi Arabia 13: 5562." title="Scolopendromorph and Geophilomorph centipedes from Oman and the United Arab Emirates." volume="13" year="1993">Lewis and Gallagher 1993</bibRefCitation>
), a character that separates these species even when accounting for body size.
</paragraph>
<caption id="4A08F1A02F8CFC56D94FCAAE95F65A80" ID-DOI="10.3897/zookeys.1228.143007.figure17" ID-arpha="005A522F-6D5C-56CB-B289-4DD2B1500FDE" httpUri="https://binary.pensoft.net/fig/1257836" startId="F17">
<paragraph id="68C04D9CC04308586CB4D476D5D7C79C">
<label id="8058D02B2114954E3A8496DCB5FD8F8D">Figure 17.</label>
</paragraph>
<paragraph id="82811FB82FDB6F582126F6BE5B86A220">
<taxonomicName id="8217ADAE79D218E5B7F6B1BB73D7DEC6" authorityName="Newport" authorityYear="1845" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="brevilabiatus">
<emphasis id="73902A27A2D32AD68850C14088DE3780" italics="true">Orphnaeus brevilabiatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="158DBEE6186BE1D69F000F0B52254B14" DOI="10.1111/j.1096-3642.1842.tb00370.x" author="Newport G" issue="4" journalOrPublisher="Transactions of the Linnean Society of London" pagination="349-439" refId="B72" refString="Newport G (1845) XXVIII. Monograph of the Class Myriapoda, Order Chilopoda. Transactions of the Linnean Society of London 19 (4): 349-439. https://doi.org/10.1111/j.1096-3642.1842.tb00370.x" title="XXVIII. Monograph of the Class Myriapoda, Order Chilopoda." volume="19" year="1845">Newport, 1845</bibRefCitation>
).
<specimenCode id="8EDCED169EC3AEAEDE85FC746197E82C">NHMUK 015991423</specimenCode>
<emphasis id="4C791D999E977CC5C4D65A22321D4006" bold="true">A</emphasis>
leg bearing segments 77 79, ventral view
<emphasis id="572FACAE6E7B93F745D937F96E4C4C2E" bold="true">B</emphasis>
metasternite of leg-bearing segment 78, ventral view.
</paragraph>
</caption>
<caption id="335C17B00FD242252F50DEAF3B6213B9" ID-DOI="10.3897/zookeys.1228.143007.figure18" ID-arpha="5221D50F-B5A9-55CF-886A-C1988EE3D1A6" httpUri="https://binary.pensoft.net/fig/1257837" startId="F18">
<paragraph id="17F1F423BE84EEEFFC8DFEEC7E78B053">
<label id="CDBFFF87AA1724870315827E13C58828">Figure 18.</label>
</paragraph>
<paragraph id="804E14599702B799900D3501F8937D3B">
Female gonopods of
<taxonomicName id="6D0F47A0DDC50F4E9C3B8BBC746CED6D" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="undefined-A">
<emphasis id="B1464A599869CF8272726FEBC59308EF" italics="true">Orphnaeus</emphasis>
<bibRefCitation id="5F0030F01C32C81E541EBF080839C74C" author="Meinert F" journalOrPublisher="Naturhistorisk Tidsskrift" pagination="1-128" refId="B68" refString="Meinert F (1870) Myriapoda Musei Hauniensis. Bidrag til myriapodernes morphologi og systematik. I. Geophili. Naturhistorisk Tidsskrift 7: 1-128." title="Myriapoda Musei Hauniensis. Bidrag til myriapodernes morphologi og systematik. I. Geophili." volume="7" year="1870">Meinert, 1870</bibRefCitation>
species
<emphasis id="0385B63280A7355426C2BF766F67A4C5" bold="true">A, B, C</emphasis>
</taxonomicName>
<taxonomicName id="1F89E829E01DC7FC76B998FBB02BE71B" authorityName="Newport" authorityYear="1845" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="brevilabiatus">
<emphasis id="FBF8B4EAC64ED88257CD5C9218E063F0" italics="true">Orphnaeus brevilabiatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="2CFB0886099532F89B663E45FC180980" DOI="10.1111/j.1096-3642.1842.tb00370.x" author="Newport G" issue="4" journalOrPublisher="Transactions of the Linnean Society of London" pagination="349-439" refId="B72" refString="Newport G (1845) XXVIII. Monograph of the Class Myriapoda, Order Chilopoda. Transactions of the Linnean Society of London 19 (4): 349-439. https://doi.org/10.1111/j.1096-3642.1842.tb00370.x" title="XXVIII. Monograph of the Class Myriapoda, Order Chilopoda." volume="19" year="1845">Newport, 1845</bibRefCitation>
). (A =
<specimenCode id="00B5B0D9B29B860B1883CCD14230CF42">NHMUK 015991423</specimenCode>
(Myanmar), B =
<specimenCode id="8A6CAE5F071BE65D921D2CEB971A5341">NHMUK 015991421</specimenCode>
(Thailand), C =
<specimenCode id="74AED256EC19C5F8E3E6708BAE15C6D1">NHMUK 015991420</specimenCode>
(Singapore ))
<emphasis id="5868528F6446FEF286238DBD7B6B94B8" bold="true">D, E, F, G, H</emphasis>
<taxonomicName id="E96A1A0D59635AAAAE101F09BB74E760" authority="Verhoeff, 1938" authorityName="Verhoeff" authorityYear="1938" class="Chilopoda" family="Oryidae" genus="Orphnaeus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="dekanius">
<emphasis id="52DF9C8B136FA0808893036AE8DFCDA7" italics="true">Orphnaeus dekanius</emphasis>
<bibRefCitation id="324F5CBE925D69B537B76E8F6C3C52BF" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbücher, Abteilung für Systematik" pagination="339-388" refId="B107" refString="Verhoeff KW (1938) Chilopoden-Studien zur Kenntnis der Epimorphen. Zoologische Jahrbucher, Abteilung fur Systematik 71: 339-388." title="Chilopoden-Studien zur Kenntnis der Epimorphen." volume="71" year="1938">Verhoeff, 1938</bibRefCitation>
</taxonomicName>
. (D =
<specimenCode id="57229FC653AD827DC78C8038FF6CF565">NHMUK 015991415</specimenCode>
(Sri Lanka), E =
<specimenCode id="AF93FB564A1FC25E12C4CA52CE75C8FA">NHMUK 015991413</specimenCode>
(Sri Lanka), F =
<specimenCode id="CB602F18FEF72F88162C45F00B715CA7">NHMUK 015991417</specimenCode>
(Singapore), G =
<specimenCode id="18AE126BA83F77657C102485B89D6345">NHMUK 015991418</specimenCode>
(Singapore), H =
<specimenCode id="99E778362AAEB880AF59990B2553E277">NHMUK 015991416</specimenCode>
(Sri Lanka )). Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,623 @@
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<treatment id="4FD2EFF6F9BE5603832C3466D1B6B45B" ID-arpha="4FD2EFF6-F9BE-5603-832C-3466D1B6B45B" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B" httpUri="http://treatment.plazi.org/id/4FD2EFF6F9BE5603832C3466D1B6B45B" scope_class="Chilopoda" scope_family="Geophilidae" scope_order="Geophilomorpha">
<subSubSection id="89D68ED7A2DC9B13F1A5C32A874B7D21" type="nomenclature">
<paragraph id="442531CC31B5637E502778F3AFB11890">
<taxonomicName id="47E27FA87516CCFC9C63A59DEC21D79D" ID-CoL="59LW8" authority="(Pocock, 1891 a)" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="DFF93AF17039EE71392F12D958AE44B5" italics="true">Tuoba sydneyensis</emphasis>
(Pocock, 1891 a)
</taxonomicName>
</paragraph>
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<paragraph id="5FA0C72083B8621F787C41F8B0A0C3B1">
<figureCitation id="AF7356FDA20089F03F1F7C2FEAFA3D9A" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tuoba sydneyensis (Pocock, 1891 a). NHMUK 015991484 A head and antennae, dorsal view B cephalic shield, dorsal view C antennal article XIV, dorsal view." figureDoi="10.3897/zookeys.1228.143007.figure6" httpUri="https://binary.pensoft.net/fig/1257825">Figs 6</figureCitation>
,
<figureCitation id="8D97DCFAB54011D90BD4BE22AF02F873" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Tuoba sydneyensis (Pocock, 1891 a) B, C NHMUK 015991478 A NHMUK 015991484 B clypeus and labrum, ventral view C labrum, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure7" httpUri="https://binary.pensoft.net/fig/1257826">7</figureCitation>
,
<figureCitation id="21040F346044C6D2866DB4750CF93FB9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tuoba sydneyensis (Pocock, 1891 a). NHMUK 015991484 A Mandible, lateral view B First and second maxillae, ventral view C second maxillary article III and pretarsus, ventral view D forcipular segment, ventral view E right forcipule, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure8" httpUri="https://binary.pensoft.net/fig/1257827">8</figureCitation>
,
<figureCitation id="033E66D33CEB8863FCDF3C18A514C433" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Tuoba sydneyensis (Pocock, 1891 a) A NHMUK 015991484, female ultimate leg-bearing and postpedal segments, ventral view B NHMUK 015991478, juvenile ultimate leg-bearing and postpedal segments, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure9" httpUri="https://binary.pensoft.net/fig/1257828">9</figureCitation>
,
<figureCitation id="13E33FFFB0A0AE5A3632FE707B987DE7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Tuoba sydneyensis (Pocock, 1891 a), NHMUK 01591483 A male ultimate leg-bearing and postpedal segments, ventral view B male gonopods, ventral view C penis, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure10" httpUri="https://binary.pensoft.net/fig/1257829">10</figureCitation>
</paragraph>
</subSubSection>
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<paragraph id="F311E9DA8ECFAA7A5CC8DF01129DF2F7">
<treatmentCitationGroup id="281CA559AD2057AD65DFD7DC34FCD0EE">
<treatmentCitation id="25CC60879C1B021D087529D37C55D615">
<taxonomicName id="275984B2928B7486E68F658BD575AF7B" ID-CoL="3FR4F" authority="Verhoeff, 1939" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lemuricus" status="syn. nov." subGenus="Bothrogeophilus">
<emphasis id="A561D178D81132827F31E780DFB9F8E9" italics="true">Geophilus</emphasis>
(
<emphasis id="2965525C2059876046D0A08DDAB06691" italics="true">Bothrogeophilus</emphasis>
)
<emphasis id="A67B2E11D45E20208D5A481F23417D7E" italics="true">lemuricus</emphasis>
<bibRefCitation id="8093A4866D545DA436738E1371359AEB" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 72: 71-98." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff, 1939</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<taxonomicNameLabel id="F887DDC261DE9C7A62F3EAAFF4705B25">syn. nov.</taxonomicNameLabel>
</treatmentCitationGroup>
</paragraph>
<paragraph id="B25AE2C70DDF2761CC28EEFFFB0F999B">
<treatmentCitationGroup id="79DD629C56DB6A04DEBE2175BE5FAD99">
<treatmentCitation id="98746EF2B9214BDA13672A190304EB55">
<taxonomicName id="9F7F25D4B9D6314F82B7472E27173DB8" ID-CoL="3MR8J" authority="Lawrence, 1960" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="waterloti" status="syn. nov.">
<emphasis id="012C57228807F0BEAAD922482BFFB61D" italics="true">Hovanyx waterloti</emphasis>
<bibRefCitation id="6B9D1BFEFCC54F70637F1E262455B598" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<taxonomicNameLabel id="EA4736AF5E8C2A8D40045EEAFB0EEC03">syn. nov.</taxonomicNameLabel>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJLAI" type="material">
<paragraph id="63DD0AEB48B04C228CC59CED2581BE9E">
<heading id="653ADF872EA10DD6B95DA661E4ADC3C9" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="F5033063C15750F038BBD8E705E62F10">
<specimenCount id="153A46AACA312AAB0800E5FAAF6B86DF" count="19" type="generic">19 specimens</specimenCount>
:
<materialsCitation id="E6B0232E1DCD7D30BD708D21D185BA9B" collectingDate="1968-03-13" collectingDateMax="1968-03-20" collectingDateMin="1968-03-13" collectionCode="NHMUK" collectorName="B. Cogan &amp; A. Hutson" location="South Island" municipality="Aldabra" specimenCode="NHMUK 015991475" specimenCount="1">
<specimenCode id="535D1C1C08C44B6FBCC22AECDAC9AB11">NHMUK 015991475</specimenCode>
,
<specimenCount id="6E58B40B8F6DB49FDAD7ECFE2EBC3A62" count="1" type="generic">1 specimen</specimenCount>
,
<location id="EB1764C94CA641EA87D1B9CDFD2EBBDE" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:EB1764C94CA641EA87D1B9CDFD2EBBDE" municipality="Aldabra" name="South Island">South Island</location>
,
<collectingMunicipality id="8FAC84648DB9850DB6084D77DE3AD4DE">Aldabra</collectingMunicipality>
,
<date id="8FFC9A9147DD99BE882B5BAA929D4AA8" value="1968-03-13" valueMax="1968-03-20" valueMin="1968-03-13">
<collectingDate id="81C9D70FDE757BF5A8F44B5B148FCB24" value="1968-03-13" valueMax="1968-03-20" valueMin="1968-03-13">1320.03. 1968</collectingDate>
</date>
, leg.
<collectorName id="FAC3EF76908B02D88390DE42A719E2A1">B. Cogan</collectorName>
&amp;
<collectorName id="46F6EB93C0C62C7F0BA6A01D61E709B5">A. Hutson</collectorName>
</materialsCitation>
;
<materialsCitation id="609C1131DC1A5D40AAA8DA17CA55FB25" collectingDate="1974-03-27" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Cinq Cases / Point Hodoul Arga" specimenCode="NHMUK 015991476" specimenCount="1" specimenCount-juvenile="1">
<specimenCode id="80CEF648FE411DBED747B3044AFE80FA">NHMUK 015991476</specimenCode>
,
<specimenCount id="13A071ED6FC54146E81BC198C08F853C" count="1" type="juvenile">1 juvenile</specimenCount>
,
<location id="24B0ADC90DE717F1D1CB0D8F18C59E97" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:24B0ADC90DE717F1D1CB0D8F18C59E97" name="Cinq Cases / Point Hodoul Arga">Cinq Cases / Point Hodoul Arga</location>
,
<date id="B54DD3697FFB9148B7D694CC45C84BB9" value="1974-03-27">
<collectingDate id="0D23FC10762F67ED0249E9CA8E8078A8" value="1974-03-27">27. 03. 1974</collectingDate>
</date>
, leg.
<collectorName id="CF4779F17E3AC235252ECD3C04AF8FCC">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="FAD97E7BDF3AFD417068071A5E6580F1" collectedFrom="inside fallen dead coconut tree" collectingDate="1974-02-23" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Picard" municipality="Aldabra" specimenCode="NHMUK 015991477" specimenCount="1" specimenCount-female="1">
<specimenCode id="FA57DCE28F5D94A5FB9C665C1CF6FA76">NHMUK 015991477</specimenCode>
,
<specimenCount id="4028FC33F23F33947AA9D3B40F463E70" count="1" type="female">1 ♀</specimenCount>
,
<collectedFrom id="E5997F0060A384C685F8128C4D9D1243">inside fallen dead coconut tree</collectedFrom>
,
<location id="BFC826191B7ED7535D8D3F8815F40EDD" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:BFC826191B7ED7535D8D3F8815F40EDD" municipality="Aldabra" name="Picard">Picard</location>
,
<collectingMunicipality id="C2006806342561B61C5F570026587584">Aldabra</collectingMunicipality>
,
<date id="FD23DD4F3F9EFD8714BAA7A564CAF3AE" value="1974-02-23">
<collectingDate id="B58B7A7507AA11F3C44AB4EEFA1D06B1" value="1974-02-23">23. 02. 1974</collectingDate>
</date>
, leg.
<collectorName id="6D4ED35B175258E960A38AD3DFFDAD06">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="AACA324D1128695C207302C122763B2F" collectingDate="1974-03-10" collectingDateMax="1974-03-17" collectingDateMin="1974-03-10" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Cinq Cases" municipality="Aldabra" specimenCode="NHMUK 015991478" specimenCount="5" specimenCount-female="3" specimenCount-juvenile="1" specimenCount-male="1">
<specimenCode id="F114B3CCB795132580EAE98C2B985558">NHMUK 015991478</specimenCode>
,
<specimenCount id="8BB8F3F48E6562177FE14EE5D97D84F4" count="3" type="female">3 ♀</specimenCount>
,
<specimenCount id="3165C02491841AB7B5F7442BE53387DA" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="7A1CA1C118AAED03EE3A25C4BB1D4079" count="1" type="juvenile">1 juvenile</specimenCount>
,
<location id="3D9FAD4D95EE7511C118C81A0EE8E07D" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:3D9FAD4D95EE7511C118C81A0EE8E07D" municipality="Aldabra" name="Cinq Cases">Cinq Cases</location>
,
<collectingMunicipality id="4280E279A8BC2BB03728556131649C07">Aldabra</collectingMunicipality>
,
<date id="009D8E255E24859DA593FB0824C6A5E3" value="1974-03-10" valueMax="1974-03-17" valueMin="1974-03-10">
<collectingDate id="1D48A8ABE997AD9A6962810AD778091A" value="1974-03-10" valueMax="1974-03-17" valueMin="1974-03-10">1017.03. 1974</collectingDate>
</date>
, leg.
<collectorName id="BD14F2154DFAF68E049FB5095B577A72">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="6BEABE708281926D6CC782696ECD0738" collectedFrom="Cyperus ligularis soil and litter" collectingDate="1974-06-05" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Dune Patates" municipality="Aldabra" specimenCode="NHMUK 015991479" specimenCount="1" specimenCount-female="1">
<specimenCode id="609ECDC23A384AFA06FDEB884C0CE696">NHMUK 015991479</specimenCode>
,
<specimenCount id="7CF1ADB4914DAD138A05972725906339" count="1" type="female">1 ♀</specimenCount>
,
<collectedFrom id="2699241C412C306EC1518ED84AD63791">
<taxonomicName id="F6D629DE288A8A9C9356A923DB278C3D" class="Liliopsida" family="Cyperaceae" genus="Cyperus" kingdom="Plantae" order="Poales" phylum="Tracheophyta" rank="species" species="ligularis">
<emphasis id="820FFAD1496E3A0CD8D2A32D9CACFD3A" italics="true">Cyperus ligularis</emphasis>
</taxonomicName>
soil and litter
</collectedFrom>
,
<location id="54C8A4CF787B510080030913E7DEF9FD" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:54C8A4CF787B510080030913E7DEF9FD" municipality="Aldabra" name="Dune Patates">Dune Patates</location>
,
<collectingMunicipality id="D3E2CB67128D2C09778210A4D9455BD2">Aldabra</collectingMunicipality>
,
<date id="8FA121C946BBEAF5F1AF4DE59C09A281" value="1974-06-05">
<collectingDate id="333E9F4B093B54FDCFD6BC1674EEACC4" value="1974-06-05">05. 06. 1974</collectingDate>
</date>
, leg.
<collectorName id="5E2F32800C5325B3C34D2C1E52AFA817">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="7B61984FC760B21196B2548C3B3AE620" collectedFrom="Mystroxylon and Dracaena litter" collectingDate="1974-12-03" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Gionnet" specimenCode="NHMUK 015991480" specimenCount="1" specimenCount-male="1">
<specimenCode id="3F7B10021696367998E423BEB125D13C">NHMUK 015991480</specimenCode>
,
<specimenCount id="699E52DD1F8A22DCBA1408D33E776A9A" count="1" type="male">1 ♂</specimenCount>
, 1 incomplete,
<collectedFrom id="5AA1B5BEC6289C34ED1FCB96344B8125">
<taxonomicName id="F225E18F751591E5A941D08471A192EF" class="Magnoliopsida" family="Celastraceae" genus="Mystroxylon" kingdom="Plantae" order="Celastrales" phylum="Tracheophyta" rank="genus">
<emphasis id="7EAE9AF1970EFA9796A92DB28917EDDB" italics="true">Mystroxylon</emphasis>
</taxonomicName>
and
<taxonomicName id="1A0FB1DAE1CEC59833491365397FE72D" class="Squamata" family="Teiidae" genus="Dracaena" kingdom="Animalia" phylum="Chordata" rank="genus">
<emphasis id="E5FB5E3E7C13398CE3E28177F134260F" italics="true">Dracaena</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="E238F5B23346D6E6CAD735CD9861D03D" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:E238F5B23346D6E6CAD735CD9861D03D" name="Gionnet">Gionnet</location>
,
<date id="02ADFA37AD58ECCCC759002BA1505650" value="1974-12-03">
<collectingDate id="C0D2331A12F3DF501A1CB3AA9A5042BB" value="1974-12-03">03. 12. 1974</collectingDate>
</date>
, leg.
<collectorName id="DA37E171B1D8D98F43688F520CF7BD0C">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="C89392B0B04ECD23861885C51FBD75F4" collectedFrom="Casuarina litter" collectingDate="1974-05-08" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Picard" municipality="Aldabra" specimenCode="NHMUK 015991481" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCode id="6A594E9E6F70DD2A5CFC475621726F99">NHMUK 015991481</specimenCode>
,
<specimenCount id="436E41CF0E96C2259FDD447B80A8ACAD" count="1" type="female">1 ♀</specimenCount>
,
<specimenCount id="291DCA672EAA921063596BD6D425029C" count="1" type="male">1 ♂</specimenCount>
,
<collectedFrom id="50B34B0D33B09E70693B6415514877BA">
<taxonomicName id="27317855BF1F9540148CC51B33A0CC8F" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="genus">
<emphasis id="E08AE3D5B8A7FE1B76A7AC41C44D9DC1" italics="true">Casuarina</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="ACE88BBC6364091D6429CC9F47589D70" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:ACE88BBC6364091D6429CC9F47589D70" municipality="Aldabra" name="Picard">Picard</location>
,
<collectingMunicipality id="754219CD9EAF72F49449680A25680C75">Aldabra</collectingMunicipality>
,
<date id="1F0980A7C55297805F2B2B4A7627DB0A" value="1974-05-08">
<collectingDate id="D19875ABCB36DA152248C135686A12E2" value="1974-05-08">08. 05. 1974</collectingDate>
</date>
, leg.
<collectorName id="F0F0E72E70E4B6AA7076EE680E0E5424">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="F801CA0CC04B1AA86D92B429747739FD" collectedFrom="Pemphis litter" collectingDate="1975-01-29" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Dune D'Messe" municipality="Grande Terre" specimenCode="NHMUK 015991482" specimenCount="1" specimenCount-male="1">
<specimenCode id="7A39A095A5CB842EC8F7A489BC290A45">NHMUK 015991482</specimenCode>
,
<specimenCount id="1009419D2F2C23E46A6A23F2473384FE" count="1" type="male">1 ♂</specimenCount>
,
<collectedFrom id="D23C5059890DBA35C4A3D0F06AC51398">
<taxonomicName id="58D24ECF47E6A98DBF0F3566FC946FEA" class="Magnoliopsida" family="Lythraceae" genus="Pemphis" kingdom="Plantae" order="Myrtales" phylum="Tracheophyta" rank="genus">
<emphasis id="C624135C29F5F27D7FE568CDC4F4F80E" italics="true">Pemphis</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="898818102BFAEF84E073ECBE22C26319" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:898818102BFAEF84E073ECBE22C26319" municipality="Grande Terre" name="Dune D'Messe">Dune DMesse</location>
,
<collectingMunicipality id="8049D53C898A0D14CDCE44D00ED983C6">Grande Terre</collectingMunicipality>
,
<date id="9C70E28A7623C0F2F7FB0D0A2DB2172A" value="1975-01-29">
<collectingDate id="40EF3E8A87EE52F7D28D752A2C276CFD" value="1975-01-29">29. 01. 1975</collectingDate>
</date>
, leg.
<collectorName id="5FE94C716EB98E75655516ED26CC195A">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="DC3E2BED80C0E330F4B2EADBA2ED111E" collectingDate="1974-12-27" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Casuarina" municipality="Picard" specimenCode="NHMUK 015991483" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCode id="124E6B2074CA26ED58D9FFD2CD062FFE">NHMUK 015991483</specimenCode>
,
<specimenCount id="A371CA41C9983F0900473A8E344CE4CB" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="B7C9AF76DC9F410760F8B6573F9FB1A4" count="1" type="female">1 ♀</specimenCount>
,
<taxonomicName id="FD6C8BC02392FFBD938C515454A1EB10" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="genus">
<emphasis id="B654FA0FE9A153A3852718B9139B3172" italics="true">
<location id="9D0822A70381095AA457A06AB36CCEA2" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:9D0822A70381095AA457A06AB36CCEA2" municipality="Picard" name="Casuarina">Casuarina</location>
</emphasis>
</taxonomicName>
,
<collectingMunicipality id="80C219D5F1792158BECE20CDB8C1822C">Picard</collectingMunicipality>
,
<date id="C6B1E6F0C2D8B3D2B5D97FD5FFA5F5EE" value="1974-12-27">
<collectingDate id="0402814B87D14A5AC6B4E32E5F2B78DD" value="1974-12-27">27. 12. 1974</collectingDate>
</date>
, leg.
<collectorName id="12F5BBD5493EC399538C4BFCDD1180E1">V. W. Spaull</collectorName>
.
</materialsCitation>
;
<materialsCitation id="88699C8E6C4FAB12EC7F7604033535BE" collectedFrom="Suriana litter" collectingDate="1974-03-22" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Point Hodoul" municipality="Grande Terre" specimenCode="NHMUK 015991484" specimenCount="2" specimenCount-female="2">
<specimenCode id="2EB6E63F97FD41D28EACEB7FDD5DDEEA">NHMUK 015991484</specimenCode>
,
<specimenCount id="67ABE4F3CE3CF631551B6047B5BC201E" count="2" type="female">2 ♀</specimenCount>
,
<collectedFrom id="E47F7960C7067013E5945952006C1612">
<taxonomicName id="D51FDD8FFD876685CA3CD99CE2684402" class="Magnoliopsida" family="Surianaceae" genus="Suriana" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="genus">
<emphasis id="B6CA388711EB69B99E2983652A0F3AF9" italics="true">Suriana</emphasis>
</taxonomicName>
litter
</collectedFrom>
near
<location id="49339A8BB5F7A99F58EA72A5162A3F06" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:49339A8BB5F7A99F58EA72A5162A3F06" municipality="Grande Terre" name="Point Hodoul">Point Hodoul</location>
,
<collectingMunicipality id="191DAF4A0016A3A239D44DC02A6F123B">Grande Terre</collectingMunicipality>
,
<date id="7CED4B5512E2D818DC98F4902047478C" value="1974-03-22">
<collectingDate id="F79432CB81067B62CCDA6CC7917DF837" value="1974-03-22">22. 03. 1974</collectingDate>
</date>
, leg.
<collectorName id="2D6E2B819DC45A70C204204A97436BF0">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="7CDCA30D71EBD8AB3B4E6505768B92DA" collectedFrom="Cocos litter" collectingDate="1974-12-14" collectorName="V. W. Spaull" location="Esprit" municipality="Aldabra" specimenCount="1" specimenCount-juvenile="1">
<specimenCount id="6BCA6725E347D8A81A9B5CEBEADCDBA1" count="1" type="juvenile">1 juvenile</specimenCount>
,
<collectedFrom id="67BD94459DE777A28D8DBA11DE0F3A60">
<taxonomicName id="35C1DD0AA6979E5ACF6DAD8338A97C42" class="Liliopsida" family="Arecaceae" genus="Cocos" kingdom="Plantae" order="Arecales" phylum="Tracheophyta" rank="genus">
<emphasis id="98FBA7A7EDDA9B77BA5CCBFC2DEF3708" italics="true">Cocos</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="2291FD52F4DB352F4AF709BD7E47A0BA" LSID="urn:lsid:plazi:treatment:4FD2EFF6F9BE5603832C3466D1B6B45B:2291FD52F4DB352F4AF709BD7E47A0BA" municipality="Aldabra" name="Esprit">Esprit</location>
,
<collectingMunicipality id="119809B39BC3D00D4A1C68970FD5E187">Aldabra</collectingMunicipality>
,
<date id="CC97B4E2A2BCD8C746F6431545E89155" value="1974-12-14">
<collectingDate id="CDD352667BFE90B2884B10CF04EC9BE2" value="1974-12-14">14. 12. 1974</collectingDate>
</date>
, leg.
<collectorName id="F32D6F10E7B2578AAA57B5EFE821F8E0">V. W. Spaull</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELNAI" type="remarks">
<paragraph id="B03D913EC1BE60AD27A1D7EB910ABA6F">
<heading id="44F3F43577800C48A679E17F3931075E" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="C2561505E1E3A422A4F1E6F74528AA6A">
<taxonomicName id="EB01364DB9DAA6AC2E892B178A865BCA" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="5AF4B81708CC48A6FE4EB1892BC4BE8B" italics="true">Tuoba sydneyensis</emphasis>
</taxonomicName>
has previously been reported from the
<collectingCountry id="1BC02DBFADA09AE3686F1501095BD223" name="Seychelles">Seychelles</collectingCountry>
(
<bibRefCitation id="0E80F358FFD1FCD6B34D191B01E3FA09" author="Bonato and Minelli" firstAuthor="Bonato" journalOrPublisher="Phelsuma" pagination="9-38" refId="B11" refString="Bonato L, Minelli A (2010) The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha). Phelsuma 18: 938." title="The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha)." volume="18" year="2010">Bonato and Minelli 2010</bibRefCitation>
). Material presently described from the Aldabra Atoll, as well as specimens collected on Serpent Island (
<collectingCountry id="7CDF3BB87D3C55E1C67930870B9FFEFB" name="Mauritius">Mauritius</collectingCountry>
) and other islands of the
<collectingCountry id="12D00A9A59CAAE30F9DECD2FC664E029" name="Seychelles">Seychelles</collectingCountry>
(
<bibRefCitation id="CBCE886A99E225B4E46FCFD58E52486E" author="Bonato and Minelli" firstAuthor="Bonato" journalOrPublisher="Phelsuma" pagination="9-38" refId="B11" refString="Bonato L, Minelli A (2010) The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha). Phelsuma 18: 938." title="The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha)." volume="18" year="2010">Bonato and Minelli 2010</bibRefCitation>
) can be assigned as conspecific on the basis of the low number of leg-bearing segments (4145), elongation of the antennal articles (Fig.
<figureCitation id="201BB13F37280B242F4705641E6DC3FB" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Tuoba sydneyensis (Pocock, 1891 a). NHMUK 015991484 A head and antennae, dorsal view B cephalic shield, dorsal view C antennal article XIV, dorsal view." figureDoi="10.3897/zookeys.1228.143007.figure6" httpUri="https://binary.pensoft.net/fig/1257825">6 A</figureCitation>
) and in the shape of the ultimate leg-bearing segment metasternite (
<bibRefCitation id="06E38B695CB0A293FD70AFE8619693DC" author="Jones" firstAuthor="Jones" journalOrPublisher="Records of the Western Australian Museum" pagination="333-346" refId="B42" refString="Jones RE (1998) On the species of Tuoba (Chilopoda: Geophilomorpha) in Australia, New Zealand, New Caledonia, Solomon Islands and New Britain. Records of the Western Australian Museum 18: 333346." title="On the species of Tuoba (Chilopoda: Geophilomorpha) in Australia, New Zealand, New Caledonia, Solomon Islands and New Britain." volume="18" year="1998">Jones 1998</bibRefCitation>
).
</paragraph>
<caption id="BDF79FD645EBBFD3AA9C66F2A8CB2F96" ID-DOI="10.3897/zookeys.1228.143007.figure6" ID-arpha="48F0A772-F4B4-5F0B-8EB7-03B5E6076C05" httpUri="https://binary.pensoft.net/fig/1257825" startId="F6">
<paragraph id="1D0F88EA40136971248A25149CDC15B3">
<label id="4C52646480E4DB365BFEC1C0E0AB8856">Figure 6.</label>
</paragraph>
<paragraph id="CB89ABCD041BE1CC851C507D74DA854A">
<taxonomicName id="80C949F5C072D560890358B954A01316" authority="(Pocock, 1891 a)" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="FF1B862F86A8A9DEF16137568B03C738" italics="true">Tuoba sydneyensis</emphasis>
(Pocock, 1891 a)
</taxonomicName>
.
<specimenCode id="FC93FB203152D1DD97BBEFB93071BB18">NHMUK 015991484</specimenCode>
<emphasis id="0939A0586FB7243E740A7F9168F6AE1B" bold="true">A</emphasis>
head and antennae, dorsal view
<emphasis id="135504E70CABA248B858FEB4BFFB79C4" bold="true">B</emphasis>
cephalic shield, dorsal view
<emphasis id="A18781FDBF34A046B9DF0D2AADAB6DCB" bold="true">C</emphasis>
antennal article XIV, dorsal view.
</paragraph>
</caption>
<caption id="17647530898925B33AD22A627F3F386F" ID-DOI="10.3897/zookeys.1228.143007.figure7" ID-arpha="6AE3DD88-477E-52C2-8A5B-5ACA94B974AC" httpUri="https://binary.pensoft.net/fig/1257826" startId="F7">
<paragraph id="5205790463A269C5A9488B11C3792164">
<label id="395D3EF28BE90B257B1D4A80BCF3EB1B">Figure 7.</label>
</paragraph>
<paragraph id="107D43C2B8316CA223FAC74A6BEB5059">
<taxonomicName id="119291DA12CEDE826573855B3F78064B" authority="(Pocock, 1891 a)" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="5EF3A791C8FCA3F8256B7DD651B79BC6" italics="true">Tuoba sydneyensis</emphasis>
(Pocock, 1891 a)
</taxonomicName>
<emphasis id="4FC327819E5849BA3BE2C061D8831793" bold="true">B, C</emphasis>
<specimenCode id="B0160CA0CD073470F37F0096A9BF0F43">NHMUK 015991478</specimenCode>
<emphasis id="229D6B67AC972C50F14BB95CDB451C29" bold="true">A</emphasis>
<specimenCode id="547BAFFF27F010388C47F6FD879C8F46">NHMUK 015991484</specimenCode>
<emphasis id="60C856A10049182035AB7A5786B16AA9" bold="true">B</emphasis>
clypeus and labrum, ventral view
<emphasis id="516DC4D1A4EE0DB3CE1BE0DF8CA219A6" bold="true">C</emphasis>
labrum, ventral view.
</paragraph>
</caption>
<paragraph id="0BB6F20C997D636F31F01E7E61E6D50B">
The only other species of
<taxonomicName id="7D4A0250246D15E79957D479ADDF5F36" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="08D1F4F6E3A6698A1C654D7C1E81477A" italics="true">Tuoba</emphasis>
</taxonomicName>
hitherto reported from the East African coast are
<taxonomicName id="D97F5B8F5F61AB9FAD4913BA788FE002" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="poseidonis">
<emphasis id="E2BAEF105038C713435CEEDD9DF6B806" italics="true">Tuoba poseidonis</emphasis>
<bibRefCitation id="D1ADFFF0117C1CBA590413F656308FEE" author="Verhoeff KW" journalOrPublisher="Nova Acta Abhandlungen der Kaiserlichen Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher" pagination="369-465" refId="B104" refString="Verhoeff KW (1901) Beitrage zur Kenntnis palaarktischer Myriopoden. XVI. Aufsatz: zur vergleichenden Morphologie, Systematik und Geographie der Chilopoden. Nova Acta Abhandlungen der Kaiserlichen Leopoldinisch-Carolinisch Deutschen Akademie der Naturforscher 77: 369-465." title="Beiträge zur Kenntnis paläarktischer Myriopoden. XVI. Aufsatz: zur vergleichenden Morphologie, Systematik und Geographie der Chilopoden." volume="77" year="1901">Verhoeff, 1901</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="54C00850D8EFEA84808F62D0EE76D36B" DOI="10.21426/B614110330" author="Zapparoli" firstAuthor="Zapparoli" journalOrPublisher="Biogeographia - The Journal of Integrative Biogeography" pagination="125-147" refId="B114" refString="Zapparoli M (1990) Note sui Chilopodi della Somalia. Biogeographia The Journal of Integrative Biogeography 14: 125147. https://doi.org/10.21426/B614110330" title="Note sui Chilopodi della Somalia." volume="14" year="1990">Zapparoli 1990</bibRefCitation>
) and
<taxonomicName id="C6A2CFD95853A4EC2C7484FA867523D6" authority="Lewis, 1963" authorityName="Lewis" authorityYear="1963" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sudanensis">
<emphasis id="171F3EDCCCCA46227056EC171A20745B" italics="true">Tuoba sudanensis</emphasis>
<bibRefCitation id="C7B4E612FE5693827EFDBD85662B2B44" DOI="10.1080/00222936308651322" author="Lewis JGE" refId="B49" refString="Lewis JGE (1963) On Clinopodes poseidonis (Verhoeff) (Chilopoda: Geophilomorpha: Geophilidae) with a description of a new subspecies from the Red Sea littoral. Annals and Magazine of Natural History 13 (6) (B): 49 - 55. https://doi.org/10.1080/00222936308651322" year="1963">Lewis, 1963</bibRefCitation>
</taxonomicName>
. The small number of diagnostic characters separating these two species casts doubt on the validity of
<taxonomicName id="066B63003E3D6A90F23CE23C8140EA16" authorityName="Lewis" authorityYear="1963" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sudanensis">
<emphasis id="98FBF58D60064DA74B66F6A4D0CBCC09" italics="true">T. sudanensis</emphasis>
</taxonomicName>
or on the Somali record of
<taxonomicName id="B8ED50324C73E988CC872C413639A572" authorityName="Verhoeff" authorityYear="1901" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="poseidonis">
<emphasis id="DDCABA0355E275A2B93CF8338CF709F5" italics="true">T. poseidonis</emphasis>
</taxonomicName>
. Examination of additional material from the East African coast would be necessary to elucidate the diversity of
<taxonomicName id="165BCE56334ECFE4B0518F54C4FF0A26" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="E3615DC01A8A04F5FC19CA2A345849E4" italics="true">Tuoba</emphasis>
</taxonomicName>
in Northern and Eastern Africa.
</paragraph>
<paragraph id="E97230781113099CCDCFD689A87FD519">
Both species can be reliably differentiated from
<taxonomicName id="E3A86E6115C0567DB61151D51812F5EA" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="F054B5B9132C0E1EB6801A86CA7A049E" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
in the Western Indian Ocean by the larger number of leg-bearing segments (
<quantity id="1327183802667351623AF278F8943981" metricMagnitude="0" metricUnit="m" metricValue="1.3208" metricValueMax="1.3462" metricValueMin="1.2953999999999999" unit="in" value="52.0" valueMax="53.0" valueMin="51.0">5153 in</quantity>
<taxonomicName id="BD81AE77899F0F14AA8936C1A6651A5F" authorityName="Lewis" authorityYear="1963" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sudanensis">
<emphasis id="BA8B076CBB7D05FE7785BA3C34BF4F90" italics="true">T. sudanensis</emphasis>
</taxonomicName>
compared to
<quantity id="CBA4C1F4429DBBEF8D2B60824C2FBA8E" metricMagnitude="0" metricUnit="m" metricValue="1.0922" metricValueMax="1.143" metricValueMin="1.0413999999999999" unit="in" value="43.0" valueMax="45.0" valueMin="41.0">4145 in</quantity>
<taxonomicName id="6E8116F47CC070126CA3B032C7870E9E" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="05F6807C60BF8E4EF523BE62B767C7B4" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
), greater elongation of the telopodal lappets of the first maxillae (30 % of the length of the telopodite as illustrated for
<taxonomicName id="D3A6F340294FD468E6DA1136D6DF1336" authorityName="Lewis" authorityYear="1963" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sudanensis">
<emphasis id="AC9471D9F0E9CCA6B75D36BFE30DD8DE" italics="true">T. sudanensis</emphasis>
</taxonomicName>
; compared to minute in
<taxonomicName id="5163DAD8B8A66EC5E28DF567277FD26F" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="D3419C6CFAF92609E85683AC1B893193" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="9658EBC202A40CCB1E9057D0B4E5110C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tuoba sydneyensis (Pocock, 1891 a). NHMUK 015991484 A Mandible, lateral view B First and second maxillae, ventral view C second maxillary article III and pretarsus, ventral view D forcipular segment, ventral view E right forcipule, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure8" httpUri="https://binary.pensoft.net/fig/1257827">8 B</figureCitation>
)), shape of the
<taxonomicName id="47772A11C1FD051B04AD4A82AF647620" authorityName="Leach" authorityYear="1815" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="carpophagus">
<emphasis id="4FFE9309F382B85B961BC7224C73E2E9" italics="true">carpophagus</emphasis>
</taxonomicName>
structure (with a distinct median “ hump ” in
<taxonomicName id="8A9E9CEDDECE80CA86AEBDF747F72E69" authorityName="Lewis" authorityYear="1963" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sudanensis">
<emphasis id="61D150B3976F95D676D8B2E74B16ABB9" italics="true">T. sudanensis</emphasis>
</taxonomicName>
; lacking any “ hump ” medially in
<taxonomicName id="F3747ADC8DEFF7486FB0AF1EAADD4F2B" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="D0EC04E2BE8097803172C5A169356E8E" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
), point of the midbody transition (sternites
<quantity id="B00CF0E46E08D1D5D5D08445D4E35D7F" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.588" metricValueMin="5.08" unit="in" value="21.0" valueMax="22.0" valueMin="20.0">2022 in</quantity>
<taxonomicName id="BB5C28ADF6F95B1B7D3774BB94985286" authorityName="Verhoeff" authorityYear="1901" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="poseidonis">
<emphasis id="40B98FB085F4D698BB66A736E103E90F" italics="true">T. poseidonis</emphasis>
</taxonomicName>
and
<taxonomicName id="61B3A782451079C7E2AC2EEC8C2F5F0A" authorityName="Lewis" authorityYear="1963" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sudanensis">
<emphasis id="E1A582BE818F8B8D29E9D5AC8497590B" italics="true">T. sudanensis</emphasis>
</taxonomicName>
; sternites
<quantity id="76502E5D3169554578C92B0427DA59F8" metricMagnitude="-1" metricUnit="m" metricValue="3.683" metricValueMax="3.81" metricValueMin="3.556" unit="in" value="14.5" valueMax="15.0" valueMin="14.0">1415 in</quantity>
<taxonomicName id="8C773399EE2135DE23CE2C14E9AB7DD5" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="385191F9323C5E5676D4C72B10A3C27B" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
) and the shape of the ultimate leg-bearing segment metasternite (1.6 × wider than long as illustrated for male
<taxonomicName id="9B504B0B73B074C2C3834FB9675442E6" authorityName="Lewis" authorityYear="1963" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sudanensis">
<emphasis id="479707BF1FD44B83E9F0EB43BAF2C129" italics="true">T. sudanensis</emphasis>
</taxonomicName>
; 1.9 × wider than long in male
<taxonomicName id="F4716E0C514C2BBED26120A39889506D" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="04BBC15B3352CAAA086DA37FA7BF0B85" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
, Fig.
<figureCitation id="67840ACC4C7CF6600062814812CE2DD8" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Tuoba sydneyensis (Pocock, 1891 a), NHMUK 01591483 A male ultimate leg-bearing and postpedal segments, ventral view B male gonopods, ventral view C penis, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure10" httpUri="https://binary.pensoft.net/fig/1257829">10 A</figureCitation>
). The combination of diagnostic characters presented clearly unify the Western Indian Ocean populations of
<taxonomicName id="AD1342916850925B8340C92678A48058" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="FE24008B53CD82217FA326C2E790899D" italics="true">Tuoba</emphasis>
</taxonomicName>
under one morphospecies, closely matching the description of
<taxonomicName id="2EA2A444EBFBDCBE3178A67F13AB6EA9" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="AC7504F7E5080198A528379085ECFC3B" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
, which is distinct from
<taxonomicName id="52A6AAB82A23F99F6FA5BE5285BD0C66">
<emphasis id="09D360FDBD4E95F49F773C73D094D35B" italics="true">Tuoba</emphasis>
species
</taxonomicName>
recorded in continental Eastern and Northern Africa.
</paragraph>
<caption id="70190DD442138C443F9B30C8CD0D8890" ID-DOI="10.3897/zookeys.1228.143007.figure8" ID-arpha="D31C64E2-531B-5A5A-9F86-662384D420CA" httpUri="https://binary.pensoft.net/fig/1257827" startId="F8">
<paragraph id="DA2B8184F03A83BF8ACB350EE5F30AE7">
<label id="255ADEB11AC58D976C9508AF5757494E">Figure 8.</label>
</paragraph>
<paragraph id="514CE7D211985BAC3B06AAC34A759E48">
<taxonomicName id="A3D5F72736D24CD717A6CFFDF8635AB2" authority="(Pocock, 1891 a)" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="64173419B5A858A28C9C9A1582D43FBD" italics="true">Tuoba sydneyensis</emphasis>
(Pocock, 1891 a)
</taxonomicName>
.
<specimenCode id="1BC6C9E2116225E09CE3895F51206818">NHMUK 015991484</specimenCode>
<emphasis id="48201FDE5ED0F7DE51AE108FF260D4DB" bold="true">A</emphasis>
Mandible, lateral view
<emphasis id="02BB53AA19BF7FE50E542DE458F8744C" bold="true">B</emphasis>
First and second maxillae, ventral view
<emphasis id="700CBEE3682E1FC262731B0E538DF79E" bold="true">C</emphasis>
second maxillary article III and pretarsus, ventral view
<emphasis id="43161D3F1C8BDDAA69D87A4C40DDD453" bold="true">D</emphasis>
forcipular segment, ventral view
<emphasis id="F4587FCDBE920654EC52029C24E78E98" bold="true">E</emphasis>
right forcipule, ventral view.
</paragraph>
</caption>
<paragraph id="C7D3E392CCAF07C9B96A30CCCC5CE980">
<bibRefCitation id="A4B6FCA54E8CDC1D03641E42D74FF2B5" author="Verhoeff KW" firstAuthor="Verhoeff" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere 72: 7198." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff (1939)</bibRefCitation>
described
<taxonomicName id="5DF2657C993C0A2489F2F4F52CA52A43" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lemuricus" subGenus="Bothrogeophilus">
<emphasis id="AAD6EAD40A322AFF237F1866715279D5" italics="true">Geophilus</emphasis>
(
<emphasis id="080FC405030943118A25F77571547F31" italics="true">Bothrogeophilus</emphasis>
)
<emphasis id="CBC74D1F1B0FA66AA32317434168D6C0" italics="true">lemuricus</emphasis>
</taxonomicName>
from La Ponce,
<collectingCountry id="7CB99F59F0A2D7E75145BA0EBAD16997" name="Mauritius">Mauritius</collectingCountry>
and remarked that it is closely related to (“ […] nahe verwandt […] ”)
<taxonomicName id="3F32C645F65913AB9A0E57EE9340EBCD" authority="Leach, 1815" authorityName="Leach" authorityYear="1815" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="carpophagus">
<emphasis id="635663562D711D2D1C1FE4897D3B8B2D" italics="true">Geophilus carpophagus</emphasis>
Leach, 1815
</taxonomicName>
. Although incomplete, the description provides several characters that allow reliable assignment to
<taxonomicName id="13F6D1BEFCD2A67E27DA0AADFE4BB661" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="84DCF2AF13EF84F04E8FAEC5B152B723" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
, with which it agrees in number of leg-bearing segments (
<quantity id="BD32A4E656A5BF4A2C2D936BE7C385B2" metricMagnitude="0" metricUnit="m" metricValue="1.1938" unit="in" value="47.0">47 in</quantity>
<taxonomicName id="09281EC995A8C1E14F05B029014F8AD9" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lemuricus">
<emphasis id="C0B44120996DA8105D5D77E34A0A82E8" italics="true">G. lemuricus</emphasis>
</taxonomicName>
,
<quantity id="5CACAB4440BCE125D47B99E381FFF779" metricMagnitude="0" metricUnit="m" metricValue="1.0922" metricValueMax="1.143" metricValueMin="1.0413999999999999" unit="in" value="43.0" valueMax="45.0" valueMin="41.0">4145 in</quantity>
<taxonomicName id="08B2567C38F6C6539DC313918150B933" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="A3BDE6C0FB5100C49C69DFE4D2093CE0" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
from the Western Indian Ocean), the apical claw of the second maxillae being reduced in size and not overtaking surrounding setae in length (“ [...], überragt nicht die Nachbarborsten. ”) (Fig.
<figureCitation id="6D86CDC2C5D9B2C42B058AF96BEA72D8" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Tuoba sydneyensis (Pocock, 1891 a). NHMUK 015991484 A Mandible, lateral view B First and second maxillae, ventral view C second maxillary article III and pretarsus, ventral view D forcipular segment, ventral view E right forcipule, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure8" httpUri="https://binary.pensoft.net/fig/1257827">8 B, C</figureCitation>
), and in the arrangement of the coxal organs on the coxopleuron of the last leg-bearing segment, which are arranged into a rosette opening in a pouch near the edge of the metasternite (“ [...] neben dem Endbeinsternit mündet eine Tasche und in diese eine Rosette von Drüsen ”) (Figs
<figureCitation id="C87598C0268EF1E9310B579784110950" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Tuoba sydneyensis (Pocock, 1891 a) A NHMUK 015991484, female ultimate leg-bearing and postpedal segments, ventral view B NHMUK 015991478, juvenile ultimate leg-bearing and postpedal segments, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure9" httpUri="https://binary.pensoft.net/fig/1257828">9 A</figureCitation>
,
<figureCitation id="1C50C36032A8F955591A73211B1BB603" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Tuoba sydneyensis (Pocock, 1891 a), NHMUK 01591483 A male ultimate leg-bearing and postpedal segments, ventral view B male gonopods, ventral view C penis, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure10" httpUri="https://binary.pensoft.net/fig/1257829">10 A</figureCitation>
). We consider
<taxonomicName id="816D42CE251E09766FE6265807844BDB" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Geophilidae" genus="Geophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lemuricus">
<emphasis id="395C3BCE233819762F0DB8AA6304A6B9" italics="true">G. lemuricus</emphasis>
</taxonomicName>
to be a junior subjective synonym of
<taxonomicName id="4ED640C30874A024C5A91C7654796839" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="0AB10C6A1A771390438375BDFE16B5FD" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
, which has been subsequently collected from
<collectingCountry id="F43EBC0F80D631CE139BD959E61FA5A3" name="Mauritius">Mauritius</collectingCountry>
(
<specimenCode id="3199E3031242C6BA15E615BB2BBC80BD">
<collectionCode id="B7930F64DB1305684CAD3700A12D5FB6" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991411
</specimenCode>
).
</paragraph>
<caption id="7E0D4ED8D192682AD3881C495F908507" ID-DOI="10.3897/zookeys.1228.143007.figure9" ID-arpha="AA4A0D42-F358-5CFA-84A7-D6454D2DC624" httpUri="https://binary.pensoft.net/fig/1257828" startId="F9">
<paragraph id="1B61C8B8390AD46DC8729E8DB5CD79BA">
<label id="11D9526785C399A57DF2CEC0E343E7F3">Figure 9.</label>
</paragraph>
<paragraph id="1540224E284686B533821268B016C96F">
<taxonomicName id="8475E48F955780DC236B145B801D9E1C" authority="(Pocock, 1891 a) A NHMUK" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="EE63F8A22D525A84537AE28AC6380F68" italics="true">Tuoba sydneyensis</emphasis>
(Pocock, 1891 a)
<emphasis id="FAB35956CC141ED1FA9B6256AA8E52B6" bold="true">A</emphasis>
<specimenCode id="3CCE3ACDC86A4B2F811EB09F294D3FC6">NHMUK 015991484</specimenCode>
</taxonomicName>
, female ultimate leg-bearing and postpedal segments, ventral view
<emphasis id="8B02A25F100E3A69D041993E379EC4FF" bold="true">B</emphasis>
<specimenCode id="FF72B9F88F79EED51461E039202B4908">NHMUK 015991478</specimenCode>
, juvenile ultimate leg-bearing and postpedal segments, ventral view.
</paragraph>
</caption>
<caption id="45AE2F27C5C21A5D12E9DE88DC0A5674" ID-DOI="10.3897/zookeys.1228.143007.figure10" ID-arpha="5C97EDDB-4CB1-5E8E-9B36-4344C175EAD5" httpUri="https://binary.pensoft.net/fig/1257829" startId="F10">
<paragraph id="C2D56AFD66BC1458803812C0E9D5A5C9">
<label id="1551588B54EEF011CE944876A5A1DBDA">Figure 10.</label>
</paragraph>
<paragraph id="38FB019ACA34550915DF7A6D03CD1A6D">
<taxonomicName id="3A9F7B1F8B233E14C52C63A7FEFC01CF" authority="(Pocock, 1891 a)" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="5A1362CFBE008A2D349343EE42FA10DC" italics="true">Tuoba sydneyensis</emphasis>
(Pocock, 1891 a)
</taxonomicName>
,
<specimenCode id="0E37EA8B2D1DA58AF5E853157DEE4116">NHMUK 01591483</specimenCode>
<emphasis id="3C8C804F6D44A8258885C2B04E9BAD99" bold="true">A</emphasis>
male ultimate leg-bearing and postpedal segments, ventral view
<emphasis id="E9A29E1007B5FD42ED95EFF31C484455" bold="true">B</emphasis>
male gonopods, ventral view
<emphasis id="46669409E4705AF1AD83052FB3060CF0" bold="true">C</emphasis>
penis, ventral view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,449 @@
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<mods:title id="4D7985BB58A650E30A63B50940481A83">Centipedes (Myriapoda, Chilopoda) of Aldabra Atoll (Seychelles)</mods:title>
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<mods:namePart id="989BF889A5ACB359528609D0F92B4D5D">Popovici, George</mods:namePart>
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<mods:affiliation id="27D92EFA22BA7C20548E2D5E6A0E13CA">Department of Life Sciences, Imperial College London, London SW 7 2 AZ, UK &amp; &quot; Grigore Antipa &quot; National Museum of Natural History, Pavel Dimitrievici Kiseleff St. 1, 011341, Bucharest, Romania</mods:affiliation>
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<mods:namePart id="6EA75B1F06D132084B6644CF7FD15995">Edgecombe, Gregory D.</mods:namePart>
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<mods:affiliation id="94332758136F566B2F8E947F0295683C">The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<treatment id="519BCC2A8AB5583CB0F424C9519743F9" ID-arpha="519BCC2A-8AB5-583C-B0F4-24C9519743F9" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9" httpUri="http://treatment.plazi.org/id/519BCC2A8AB5583CB0F424C9519743F9" scope_class="Chilopoda" scope_family="Scolopendridae" scope_order="Scolopendromorpha">
<subSubSection id="87345B5DA0669F27646055CAE10A4ACF" type="nomenclature">
<paragraph id="B51F69242AAB70259F5BBE2B88E1819A">
<taxonomicName id="AFA86AF5C22B3896A472CBC66CE4ECE1" ID-CoL="4VS9X" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="C0B4DD4EB92AA73A2CC43C3A01EAAB27" italics="true">Scolopendra morsitans</emphasis>
<bibRefCitation id="BFD7E6987BE4CA072F135451AE8EB7DC" author="Linnaeus C" refId="B66" refString="Linnaeus C (1758) Systema naturae. Editio Decima. Holmiae: 1 - 824." year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EC1AG" type="material">
<paragraph id="3CDF7135D9DCD294FF3F2C12E8F73340">
<heading id="366065F14DE01D25861D8D564EC732E4" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="3166F5512C966C759E448A324338174D">
<specimenCount id="C1BB13A52AA968923C96058C15C89906" count="52" type="generic">52 specimens</specimenCount>
:
<materialsCitation id="5E33FE0AC215FADB49C2A90F6BB518EA" collectedFrom="Casuarina litter" collectingDate="1974-04-18" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Picard" municipality="Aldabra" specimenCode="NHMUK 015991435" specimenCount="33" specimenCount-juvenile="33">
<specimenCode id="A7FEECFC5D2CD59AF79EAAF660DA811C">NHMUK 015991435</specimenCode>
,
<specimenCount id="EBB762C46F3FA8967BCB286C9ECCD1D8" count="33" type="juvenile">33 juveniles</specimenCount>
;
<collectedFrom id="18B14125E1BC13597583F95B80A4F80C">
<taxonomicName id="A5B3B5730404A3DD8A24C2575549ED22" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="genus">
<emphasis id="B4442FF894481C372D53FB7477D3A427" italics="true">Casuarina</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="E6E11C92EAFA4F5BC4657BBE95B2CCFD" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:E6E11C92EAFA4F5BC4657BBE95B2CCFD" municipality="Aldabra" name="Picard">Picard</location>
,
<collectingMunicipality id="7A47973608CD93CFCDCACEE4FAB39CFD">Aldabra</collectingMunicipality>
,
<date id="68A044C204120A9B2876902ABA62ACF3" value="1974-04-18">
<collectingDate id="1A752D0544ED7AE9549EC50EE236C00F" value="1974-04-18">18. 04. 1974</collectingDate>
</date>
, leg.
<collectorName id="AF2BAB836EA24596399D34DDC93C7B48">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="F9EEF7CE1195AA78824DDFA847FC8892" collectedFrom="Casuarina" collectingDate="1974-02-04" collectionCode="NHMUK" location="Picard" municipality="Aldabra" specimenCode="NHMUK 015991436" specimenCount="1">
<specimenCode id="CD0DD1C3EF883A98A11F028F0F1FAEB2">NHMUK 015991436</specimenCode>
,
<specimenCount id="60F8114C1B2AF685D29E507C04E3A1ED" count="1" type="generic">1 specimen</specimenCount>
,
<emphasis id="BB29B57A2FBFEB7C1D7F19B5C06345D1" italics="true">
<collectedFrom id="A9181B5F499A65D75C9B1F4A6BF8BC23">Casuarina</collectedFrom>
</emphasis>
,
<location id="844DC6BFC12F586B45E2D030655135AA" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:844DC6BFC12F586B45E2D030655135AA" municipality="Aldabra" name="Picard">Picard</location>
,
<collectingMunicipality id="D850CA26979D0F329DE4622A2CD3440F">Aldabra</collectingMunicipality>
,
<date id="1A5184A1DC01651C845C1C41DE522275" value="1974-02-04">
<collectingDate id="2C9D7EE1AFADDF92B5E5336FF1F07D39" value="1974-02-04">04. 02. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="DF6D98AFA40BE325EE999558B399831F" collectedFrom="Ochna soil" collectingDate="1974-02-15" collectionCode="NHMUK" location="Picard" municipality="Aldabra" specimenCode="NHMUK 015991437" specimenCount="1">
<specimenCode id="F81357407B1621FB6CCD3F0FF8081CA2">NHMUK 015991437</specimenCode>
,
<specimenCount id="03988B0789ED73C84EFF2C91619FCA96" count="1" type="generic">1 specimen</specimenCount>
,
<collectedFrom id="E8AC95181806A7264ACD0C948EB57294">
<taxonomicName id="CDA952B7DE6E755EA114312088B849AF" class="Magnoliopsida" family="Ochnaceae" genus="Ochna" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="genus">
<emphasis id="1A3475050E047C97D353DB3DC9FDADDB" italics="true">Ochna</emphasis>
</taxonomicName>
soil
</collectedFrom>
,
<location id="1D87F9D0BD8983CD731E7A8AF572793F" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:1D87F9D0BD8983CD731E7A8AF572793F" municipality="Aldabra" name="Picard">Picard</location>
,
<collectingMunicipality id="49C50CF1870931A5D8B2B787152FE176">Aldabra</collectingMunicipality>
,
<date id="A982C215F98031B15C5AD5427C90B431" value="1974-02-15">
<collectingDate id="22759C22B86AD0C73A68B82DC19D6C0C" value="1974-02-15">15. 02. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="85F6C6F48D5304EB48B77BB068286EED" collectedFrom="mixed scrub" collectingDate="1974-12-24" collectionCode="NHMUK" location="Picard" specimenCode="NHMUK 015991438" specimenCount="2">
<specimenCode id="0F8B75598AA4B2A976703CE90C185DBE">NHMUK 015991438</specimenCode>
;
<specimenCount id="98701AB4C0A38192060B7A47FF106476" count="2" type="generic">2 specimens</specimenCount>
,
<collectedFrom id="25D6D29E575E9A02FEB70D0A73B4EB33">mixed scrub</collectedFrom>
,
<location id="22B1A74C6C7FCAF7D22A7FE5A620A2F5" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:22B1A74C6C7FCAF7D22A7FE5A620A2F5" name="Picard">Picard</location>
,
<date id="9EFD8045BC3BAA796B64D54843D9B7EC" value="1974-12-24">
<collectingDate id="D78A0CB719F0A06637173A887182E53D" value="1974-12-24">24. 12. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="1446BE73865CF4731078DAD5A918CA5D" collectedFrom="Thespesia litter" collectingDate="1973-11-15" collectionCode="NHMUK" location="Cinq Cases" municipality="Aldabra" specimenCode="NHMUK 015991439" specimenCount="1">
<specimenCode id="7C239AF642F04A3C3DF0AF88C73C10F8">NHMUK 015991439</specimenCode>
,
<specimenCount id="1340E100779B550CBF52C01DD015CF4C" count="1" type="generic">1 specimen</specimenCount>
,
<collectedFrom id="2F17CE2F04AAA46428353E999FCC3FB9">
<taxonomicName id="7FF5757A8EF6951555F6D210047FA97E" class="Magnoliopsida" family="Malvaceae" genus="Thespesia" kingdom="Plantae" order="Malvales" phylum="Tracheophyta" rank="genus">
<emphasis id="425BC7A2915E919C0ACFDD908AFE8562" italics="true">Thespesia</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="C285DD64C960F39DE601FE57123AD534" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:C285DD64C960F39DE601FE57123AD534" municipality="Aldabra" name="Cinq Cases">Cinq Cases</location>
,
<collectingMunicipality id="BA829332E606FE2B0FEBEDA1CF036402">Aldabra</collectingMunicipality>
,
<date id="C85FD23903F837745D54A7A6C47A34AE" value="1973-11-15">
<collectingDate id="92861F1692374BC22AD0D72A2DB78AE3" value="1973-11-15">15. 11. 1973</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="4FAE427CBD317DF549A335B0641D719E" collectionCode="NHMUK" location="Black Path" municipality="Picard" specimenCode="NHMUK 015991441" specimenCount="5">
<specimenCode id="6313D903EFB68CE677970DC022B21F9B">NHMUK 015991441</specimenCode>
,
<specimenCount id="1B39C1D47B074550589FAA33AB5B4A13" count="5" type="generic">5 specimens</specimenCount>
,
<location id="96B444076C5E94A31A27B2F3C0F32DFB" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:96B444076C5E94A31A27B2F3C0F32DFB" municipality="Picard" name="Black Path">Black Path</location>
,
<collectingMunicipality id="484DE227EDD3FC9C72555BBE9B6B3A5D">Picard</collectingMunicipality>
, 1975
</materialsCitation>
;
<materialsCitation id="07A978E5D4B129C08C6BB7962090798E" collectingDate="1973-12-03" collectionCode="NHMUK" location="Picard" specimenCode="NHMUK 015991442" specimenCount="1">
<specimenCode id="4D1E3E2BC84A2C6EACA7813DE4DA1C4B">NHMUK 015991442</specimenCode>
,
<specimenCount id="B1CD63238ABB5D51A7FA9A0DF5A7F731" count="1" type="generic">1 specimen</specimenCount>
,
<location id="871FA51AE56BF32F342E164EBAE643A8" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:871FA51AE56BF32F342E164EBAE643A8" name="Picard">Picard</location>
,
<date id="0230F434597DDEF82E499A9BDB139FBD" value="1973-12-03">
<collectingDate id="FA7039B5ED03A836E12B05B9038CB7FC" value="1973-12-03">03. 12. 1973</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="8B3B0708BCB10F99496FD859171E6F4C" collectingDate="1974-01-18" collectionCode="NHMUK" location="Picard" specimenCode="NHMUK 015991443" specimenCount="1">
<specimenCode id="4E8F6DCE4CCCB9A27880BB32B2D9EDBD">NHMUK 015991443</specimenCode>
,
<specimenCount id="6038033659457E338ECD01323E1E4042" count="1" type="generic">1 specimen</specimenCount>
,
<location id="46D67A7C0A716627A699F7E1F7982D44" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:46D67A7C0A716627A699F7E1F7982D44" name="Picard">Picard</location>
,
<date id="3B72B74362770AB9EAC12F847E423723" value="1974-01-18">
<collectingDate id="9767FF4C46C68E095446D3AC2B01E154" value="1974-01-18">18. 01. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="92D7A758289530CB862F082CE855D1F1" collectingDate="1974-11-18" collectionCode="NHMUK" location="Picard" specimenCode="NHMUK 015991440" specimenCount="4">
<specimenCode id="0B6D07DF565227B6E9C85321AE4235E4">NHMUK 015991440</specimenCode>
,
<specimenCount id="60D42678C9570387137ED0B4A88DDDF9" count="4" type="generic">4 specimens</specimenCount>
,
<location id="2F9CC2E6125FC17719F180DC2E3618B5" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:2F9CC2E6125FC17719F180DC2E3618B5" name="Picard">Picard</location>
,
<date id="60C017D233CEC4B8867ED52125CD5D34" value="1974-11-18">
<collectingDate id="7D3258C74D2835A034BE43459073AB3B" value="1974-11-18">18. 11. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="7EF534E3ADCC3EF562F81FD3A0961CD3" collectingDate="1974-12-08" collectionCode="NHMUK" location="Pitfall trap 5" specimenCode="NHMUK 015991444" specimenCount="1">
<specimenCode id="C9C759B3180454B59F561341CBA4B344">NHMUK 015991444</specimenCode>
,
<specimenCount id="F68CE32FF8994A5CF67ADF1BD2FCA971" count="1" type="generic">1 specimen</specimenCount>
,
<location id="417FE5F20B6D9C5ACCC432F9496679E3" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:417FE5F20B6D9C5ACCC432F9496679E3" name="Pitfall trap 5">Pitfall trap 5</location>
,
<date id="234A19EEE12286628089515EAD433B94" value="1974-12-08">
<collectingDate id="21FCD39E69D90567AC02EF984CCA7C96" value="1974-12-08">08. 12. 1974</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="F2C6B66C42022431CF53AA1BE005D9BA" collectingDate="1968-03-13" collectingDateMax="1968-03-20" collectingDateMin="1968-03-13" collectionCode="NHMUK" collectorName="B. Cogan &amp; A. Hutson" location="South Island" municipality="Aldabra" specimenCode="NHMUK 015991447" specimenCount="1">
<specimenCode id="706D3ACD769C5A786CC37D4514691034">NHMUK 015991447</specimenCode>
,
<specimenCount id="13499BFEFC8F4A076FF1C68F5D55DBA7" count="1" type="generic">1 specimen</specimenCount>
,
<location id="C2CEBCB07DFA40EB07C146D0C813AFAC" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:C2CEBCB07DFA40EB07C146D0C813AFAC" municipality="Aldabra" name="South Island">South Island</location>
,
<collectingMunicipality id="6B4E5952BAFB1760097E44EF460851DE">Aldabra</collectingMunicipality>
,
<date id="39D434EAE0566507D8B12BC348AE0EE6" value="1968-03-13" valueMax="1968-03-20" valueMin="1968-03-13">
<collectingDate id="716EE600A2158EF7086AC4EA21081EF4" value="1968-03-13" valueMax="1968-03-20" valueMin="1968-03-13">1320.03. 1968</collectingDate>
</date>
, leg.
<collectorName id="9C8C92E38762B4FF1AC4E6DC117E134F">B. Cogan</collectorName>
&amp;
<collectorName id="0B09899C750C6475D50C913677661E01">A. Hutson</collectorName>
</materialsCitation>
;
<materialsCitation id="1EC61ED4F750AA69FA66C350CDADFD0B" collectionCode="NHMUK" collectorName="B. Cogan &amp; A. Hutson" location="Ile. Michel" specimenCode="NHMUK 015991448" specimenCount="1">
<specimenCode id="5208E2D7CF712D5812E1BDAB29FC9412">NHMUK 015991448</specimenCode>
,
<specimenCount id="C054B8D14669C60AB17F01C599D6B975" count="1" type="generic">1 specimen</specimenCount>
,
<location id="B21EDEEC5D50E67048B6679BCD0D35F5" LSID="urn:lsid:plazi:treatment:519BCC2A8AB5583CB0F424C9519743F9:B21EDEEC5D50E67048B6679BCD0D35F5" name="Ile. Michel">Ile. Michel</location>
, 02.1968, leg.
<collectorName id="7D09248881B45C6E1EB4D3230C63F659">B. Cogan</collectorName>
&amp;
<collectorName id="4BCEACAE75A641752385E21FED22889A">A. Hutson</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE2AG" type="remarks">
<paragraph id="9480F02893C22B57947949BEE0814E4F">
<heading id="D9DAA862AEDFD5F5C056AE84FA20D015" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="FFD164DA11DE8D0242BEDEE4A34907D6">
Specimens of
<taxonomicName id="2C19EFE79E5A1C34B65D1E4DBB5F4FC3" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="4019D767A6D59EC336DE1F6D92047128" italics="true">S. morsitans</emphasis>
<bibRefCitation id="5BCAEC734C35D3622A4C1ED8DEB257EC" author="Linnaeus C" refId="B66" refString="Linnaeus C (1758) Systema naturae. Editio Decima. Holmiae: 1 - 824." year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
collected from mainland Africa, originally identified as
<taxonomicName id="F61BF3761D34FC6FABEF0194ACA5C271" authority="Bucherl, 1946" authorityName="Bucherl" authorityYear="1946" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="8A995404DF3317D4B293EE047936A4F4" italics="true">Scolopendra amazonica</emphasis>
<bibRefCitation id="2AF04F8131555D94F02308BDD910358D" author="Bücherl W" journalOrPublisher="Memórias do Instituto Butantan" pagination="135-158" refId="B16" refString="Bucherl W (1946) Novedades sistematicas na ordem Scolopendromorpha. Memorias do Instituto Butantan 19: 135-158" title="Novedades sistematicas na ordem Scolopendromorpha." volume="19" year="1946">Bücherl, 1946</bibRefCitation>
</taxonomicName>
overlap with the specimens from the Aldabra Atoll in several characters (Table
<tableCitation id="FD1363DFCBA49136A9BB6037C70720DD" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological variability in Scolopendra morsitans populations obtained from literature data (Lewis 1966, 1967, 1969) and examined specimens." httpUri="http://table.plazi.org/id/E034164A8132C3275A4AB901AB0487CB" tableUuid="E034164A8132C3275A4AB901AB0487CB">2</tableCitation>
) (
<bibRefCitation id="4CF168F90BA8B4FE29A7FF83D525D13F" DOI="10.1111/j.1469-7998.1966.tb03893.x" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Journal of Zoology, London" pagination="188-203" refId="B50" refString="Lewis JGE (1966) The taxonomy and biology of the centipede Scolopendra amazonica in the Sudan. Journal of Zoology, London 149: 188203. https://doi.org/10.1111/j.1469-7998.1966.tb03893.x" title="The taxonomy and biology of the centipede Scolopendra amazonica in the Sudan." volume="149" year="1966">Lewis 1966</bibRefCitation>
,
<bibRefCitation id="6269E72FFD6947D492BE57A2477D971A" DOI="10.1111/j.1095-8312.1967.tb00973.x" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Proceedings of the Linnean Society of London" pagination="185-207" refId="B51" refString="Lewis JGE (1967) The scolopendromorph centipedes of the Sudan with remarks on taxonomic characters in the Scolopendridae. Proceedings of the Linnean Society of London 178: 185207. https://doi.org/10.1111/j.1095-8312.1967.tb00973.x" title="The scolopendromorph centipedes of the Sudan with remarks on taxonomic characters in the Scolopendridae." volume="178" year="1967">1967</bibRefCitation>
,
<bibRefCitation id="E6207F7BCFD2417708DBCA0E5CAF73E2" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Journal of the Linnean Society (Zoology)" pagination="315-326" refId="B52" refString="Lewis JGE (1968) Individual variation in a population of the centipede Scolopendra amazonica from Nigeria and its implications for methods of taxonomic discrimination in the Scolopendridae. Journal of the Linnean Society (Zoology) 47: 315326." title="Individual variation in a population of the centipede Scolopendra amazonica from Nigeria and its implications for methods of taxonomic discrimination in the Scolopendridae." volume="47" year="1968">1968</bibRefCitation>
,
<bibRefCitation id="246E66D87E0612EE5DAF1EADF5443E0F" DOI="10.1111/j.1096-3642.1969.tb00704.x" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="49-57" refId="B53" refString="Lewis JGE (1969) The variation of the centipede Scolopendra amazonica in Africa. Zoological Journal of the Linnean Society 48: 4957. https://doi.org/10.1111/j.1096-3642.1969.tb00704.x" title="The variation of the centipede Scolopendra amazonica in Africa." volume="48" year="1969">1969</bibRefCitation>
), but generally have a greater number of glabrous basal antennal articles, a character in which specimens from the Aldabra Atoll more closely match Indian specimens of
<taxonomicName id="2E674460E6E8755B069F89951A164D3A" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="7B050D2C7214ADAD61B52A6495CEC922" italics="true">S. morsitans</emphasis>
</taxonomicName>
previously assigned to
<taxonomicName id="50A50BB531BAD6AA70B4FB2077B6F72C" authorityName="Bucherl" authorityYear="1946" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="940BD2C704DE7D7A3166DD08582E8049" italics="true">S. amazonica</emphasis>
</taxonomicName>
(
<bibRefCitation id="3588F26283D5BC19C7BC11E7EFC4C616" DOI="10.1080/00222935508655671" author="Jangi" firstAuthor="Jangi" issue="92" journalOrPublisher="Annals and Magazine of Natural History" pagination="597-607" refId="B40" refString="Jangi BS (1955) Some aspects of the morphology of the centipede Scolopendra morsitans Linn. (Scolopendridae). Annals and Magazine of Natural History 8 (92): 597607. https://doi.org/10.1080/00222935508655671" title="Some aspects of the morphology of the centipede Scolopendra morsitans Linn. (Scolopendridae)." volume="8" year="1955">Jangi 1955</bibRefCitation>
,
<bibRefCitation id="8F6054AFC8A90458036099E74E9826E0" author="Jangi" firstAuthor="Jangi" journalOrPublisher="Entomological News" pagination="253-257" refId="B41" refString="Jangi BS (1959) Further notes on the taxonomy of the centipede Scolopendra morsitans Linnaeus (Scolopendridae). Entomological News 70: 253257." title="Further notes on the taxonomy of the centipede Scolopendra morsitans Linnaeus (Scolopendridae)." volume="70" year="1959">1959</bibRefCitation>
). In the absence of newly collected material from which molecular data can be collected to evaluate possible interspecific delimitation between different population of
<taxonomicName id="0B6B3BAF7D40797FBB2417CF163AF8DA" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="3933F4980D3BE9C990FDDE35D409EAE2" italics="true">S. morsitans</emphasis>
</taxonomicName>
, we assign material collected in the Aldabra Atoll to
<taxonomicName id="2BA390C247E487AD5930EFB1D6DBD157" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="E508ACA4EC826764BC7E11C408CDFDD2" italics="true">S. morsitans</emphasis>
</taxonomicName>
, following the conclusions on interspecific variation within this taxon reached by
<bibRefCitation id="A936ED5088C40D1C359C76B29B5B8016" author="Würmli M" firstAuthor="Wurmli" journalOrPublisher="Deutsche Entomologische Zeitschrift" pagination="201-206" refId="B113" refString="Würmli M (1975) Systematische Kriterien in der Gruppe von Scolopendra morsitans Linné, 1758 (Chilopoda: Scolopendromorpha: Scolopendridae). Deutsche Entomologische Zeitschrift 22: 201206." title="Systematische Kriterien in der Gruppe von Scolopendra morsitans Linné, 1758 (Chilopoda: Scolopendromorpha: Scolopendridae)." volume="22" year="1975">Würmli (1975)</bibRefCitation>
. Subsequent phylogenetic analyses of molecular data for
<taxonomicName id="22BC5348923A13D3BDB0BE1A6F156868" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="83BC3AF4840D684FCB5D987E4546E142" italics="true">S. morsitans</emphasis>
</taxonomicName>
identified multiple lineages within this taxon (
<bibRefCitation id="C41ADAF160E5FEEEF0EDE6AD298823AE" DOI="10.1016/j.ympev.2011.04.024" author="Joshi and Karanth" firstAuthor="Joshi" issue="3" journalOrPublisher="Molecular Phylogenetics and Evolution" pagination="287-294" refId="B43" refString="Joshi J, Karanth KP (2011) Cretaceous-Tertiary diversification among select Scolopendrid centipedes of South India. Molecular Phylogenetics and Evolution 60 (3): 287294. https://doi.org/10.1016/j.ympev.2011.04.024" title="Cretaceous-Tertiary diversification among select Scolopendrid centipedes of South India." volume="60" year="2011">Joshi and Karanth 2011</bibRefCitation>
;
<bibRefCitation id="A53F91F96F6E58A0EAD7D7573CC7D118" DOI="10.3897/zookeys.590.7950" author="Siriwut" etAl="et al." firstAuthor="Siriwut" journalOrPublisher="ZooKeys" pagination="1-124" refId="B95" refString="Siriwut W, Edgecombe GD, Sutcharit C, Tongkerd P, Panha S (2016) A taxonomic review of the centipede genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae) in mainland Southeast Asia, with description of a new species from Laos. ZooKeys 590: 1124. https://doi.org/10.3897/zookeys.590.7950" title="A taxonomic review of the centipede genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae) in mainland Southeast Asia, with description of a new species from Laos." volume="590" year="2016">Siriwut et al. 2016</bibRefCitation>
), potentially indicating the existence of a cryptic species complex as suggested by recent taxonomic review (
<bibRefCitation id="0554BB035D169EA53ADD3DAEF1708789" DOI="10.1163/187525410X12578602960380" author="Lewis JGE" journalOrPublisher="International Journal of Myriapodology" pagination="83-122" refId="B59" refString="Lewis JGE (2010 a) A key and annotated list of the Scolopendra species of the Old World with a reappraisal of Arthrorhabdus (Chilopoda: Scolopendromorpha: Scolopendridae). International Journal of Myriapodology 3: 83122. https://doi.org/10.1163/187525410X12578602960380" title="A key and annotated list of the Scolopendra species of the Old World with a reappraisal of Arthrorhabdus (Chilopoda: Scolopendromorpha: Scolopendridae)." volume="3">Lewis 2010 a</bibRefCitation>
). Until the global taxonomy of
<taxonomicName id="A5ADD9AD51DA8AF02BD39EA4BC9017E3" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="AC142FE0507ACA60E8216F531D00F0FF" italics="true">S. morsitans</emphasis>
</taxonomicName>
is interrogated using molecular data to establish if this is the case, we classify the examined material as
<taxonomicName id="67CF57820AF193567F5C848AC6D72B46" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="B470996A7BB7C3235B7AD8E802787251" italics="true">S. morsitans</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="E034164A8132C3275A4AB901AB0487CB" ID-Table-UUID="E034164A8132C3275A4AB901AB0487CB" httpUri="http://table.plazi.org/id/E034164A8132C3275A4AB901AB0487CB" startId="T2" targetIsTable="true">
<paragraph id="85CCBDABA8808D2AD926ECB6F111178D">
<label id="ACC0668D13ADA6286C49156B54C58C82">Table 2.</label>
</paragraph>
<paragraph id="5881FCA1E1088962AABD0388D44C04F4">
Morphological variability in
<taxonomicName id="DC7077F9D042C8A7C9129A3D603C4106" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="D3ADBE9BC0676CF26AFFFAD612F64ED7" italics="true">Scolopendra morsitans</emphasis>
</taxonomicName>
populations obtained from literature data (
<bibRefCitation id="4AC7612158A7C33F53F00C39A6B4AB1C" DOI="10.1111/j.1469-7998.1966.tb03893.x" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Journal of Zoology, London" pagination="188-203" refId="B50" refString="Lewis JGE (1966) The taxonomy and biology of the centipede Scolopendra amazonica in the Sudan. Journal of Zoology, London 149: 188203. https://doi.org/10.1111/j.1469-7998.1966.tb03893.x" title="The taxonomy and biology of the centipede Scolopendra amazonica in the Sudan." volume="149" year="1966">Lewis 1966</bibRefCitation>
,
<bibRefCitation id="5E76A4E32CC65F5D7F8EFC9E86F81FB2" DOI="10.1111/j.1095-8312.1967.tb00973.x" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Proceedings of the Linnean Society of London" pagination="185-207" refId="B51" refString="Lewis JGE (1967) The scolopendromorph centipedes of the Sudan with remarks on taxonomic characters in the Scolopendridae. Proceedings of the Linnean Society of London 178: 185207. https://doi.org/10.1111/j.1095-8312.1967.tb00973.x" title="The scolopendromorph centipedes of the Sudan with remarks on taxonomic characters in the Scolopendridae." volume="178" year="1967">1967</bibRefCitation>
,
<bibRefCitation id="7EE14FBB7F2A82796DBB5D8A15C8DD9B" DOI="10.1111/j.1096-3642.1969.tb00704.x" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Zoological Journal of the Linnean Society" pagination="49-57" refId="B53" refString="Lewis JGE (1969) The variation of the centipede Scolopendra amazonica in Africa. Zoological Journal of the Linnean Society 48: 4957. https://doi.org/10.1111/j.1096-3642.1969.tb00704.x" title="The variation of the centipede Scolopendra amazonica in Africa." volume="48" year="1969">1969</bibRefCitation>
) and examined specimens.
</paragraph>
</caption>
<paragraph id="E174AAAD1FBB093A6024510EDF64EA54">
<table id="TID0E6KDI" rules="all">
<tbody id="1F003440AD99C9A1179B6A77CCDFC3B5">
<tr id="D23BD46DF89F86CAADD56BC7B08D34E0">
<th id="9B3B785C36BE53E36B0CA70A18F69DE8" colspan="1" rowspan="1">Character</th>
<th id="F5D2124E6276535376574606AF0BFD94" colspan="1" rowspan="1">
<taxonomicName id="D9A48B5A272D3435A0FF5DD0C440F748" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="963F62458E5AE6B5653069F2CCC85B0E" italics="true">Scolopendra morsitans</emphasis>
</taxonomicName>
(India)
</th>
<th id="C30DBA5955230A5C9F08667134ECB1BD" colspan="1" rowspan="1">
<taxonomicName id="AE60797C25AF7C143F161D967D025D8B" authority="sensu Jangi" authorityName="Jangi" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="E41037B0FBF3756BD4C094C21CDDBA1C" italics="true">Scolopendra amazonica sensu</emphasis>
Jangi
</taxonomicName>
(=
<taxonomicName id="1694179788CF55AD9AF8201B4310B3BB" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="559B47024C71E7AEDB742A5FD9E4C812" italics="true">Scolopendra morsitans</emphasis>
</taxonomicName>
) (India)
</th>
<th id="7788A53DF4234F7B98341BA8BACAAD30" colspan="1" rowspan="1">
<taxonomicName id="3206A1B7EF77D30FD1C2A663C6206EC9" authority="sensu Lewis" authorityName="Lewis" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="amazonica">
<emphasis id="7823F7FF7F63EEA5857742484FB03AC5" italics="true">Scolopendra amazonica sensu</emphasis>
Lewis
</taxonomicName>
(=
<taxonomicName id="6F701002675DE28B9CC41665D43A8221" authorityName="Linnaeus" authorityYear="1758" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="7A3648A0DC2D7459BE79F017822F0074" italics="true">Scolopendra morsitans</emphasis>
</taxonomicName>
) (Africa)
</th>
<th id="9AAACB54253F9021E5F5ADC0EB3EDC88" colspan="1" rowspan="1">
<taxonomicName id="0E2D59B022806B3A532F2D5E2E667420" authority="(Aldabra)" baseAuthorityName="Aldabra" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="morsitans">
<emphasis id="8BAF79828F26DCD11CE70FFC4FD332A7" italics="true">Scolopendra morsitans</emphasis>
(Aldabra)
</taxonomicName>
</th>
</tr>
<tr id="841E1A5EBB2AC583BDAD542587AB8B8D">
<td id="EE750A6C377C60F6457315EC7FD660ED" colspan="1" rowspan="1">Subadult and adult body length (mm)</td>
<td id="D862824FAA3389A4EF6FF095EE8E5CAF" colspan="1" rowspan="1">18113</td>
<td id="246952CBE7C77CC510853FC0E7214E52" colspan="1" rowspan="1">1565</td>
<td id="A08AF057E516D15A3F39CC83344237E8" colspan="1" rowspan="1">13100</td>
<td id="DE381DBE5D92C8FEB3B28BE97345050B" colspan="1" rowspan="1">1378</td>
</tr>
<tr id="C378CFA46EC957AA138827A287E3DB0E">
<td id="12DB69278C9B8CAF3916029C2D44549B" colspan="1" rowspan="1">Number of antennal articles</td>
<td id="84BC1A401239E9B38550364CCFC70EF0" colspan="1" rowspan="1">20</td>
<td id="B757651B58B9DE546A576D8C3762012D" colspan="1" rowspan="1">19</td>
<td id="6FFD11BCCC04749EACEFE4CB9BB67D88" colspan="1" rowspan="1">1821</td>
<td id="8BEF362535734B4B15C6E00D3FBFA276" colspan="1" rowspan="1">(17) 1819</td>
</tr>
<tr id="74717B1F71DD32D486E2F8245C330201">
<td id="861CF84356D10B64FCD9F7F26AF56F22" colspan="1" rowspan="1">Number of glabrous antennal articles</td>
<td id="3ECA881CE54B222B153A13804E6D919E" colspan="1" rowspan="1">Can be&gt; 6</td>
<td id="84ACDE32C29FEAA670939EDB390D5B27" colspan="1" rowspan="1">≤ 6</td>
<td id="6DEC8A80B4AA2E858F5F10D3C8C963F1" colspan="1" rowspan="1">57</td>
<td id="42F8392A1950BF208F927F4E0C336316" colspan="1" rowspan="1">(3.5) 4</td>
</tr>
<tr id="1D6C1EB63BDBECC6E2BEB17616D281D3">
<td id="29652B6A2FF5ED7B5E06E8838225C585" colspan="1" rowspan="1">Complete paramedian sutures begin</td>
<td id="6941B2A6C7294F2D90B7905C584953B3" colspan="1" rowspan="1">Mostly T 3</td>
<td id="048F016DEFC2B1BFFFA4C5E0472EE827" colspan="1" rowspan="1">Mostly T 2</td>
<td id="A7CFE3FE89CE6463C8530D7312AD99D1" colspan="1" rowspan="1">TT 24</td>
<td id="A167034E72D9A69F4FCA8F7C438C3BF2" colspan="1" rowspan="1">T 3</td>
</tr>
<tr id="EEB0B1865D4222226E2623C2705FD859">
<td id="0D8D5BCE290593C40B204FC50A12193D" colspan="1" rowspan="1">Lateral margination</td>
<td id="673882C34E9BEAF9A89EDE586C7CE802" colspan="1" rowspan="1">May begin more anteriorly</td>
<td id="32B41B7F62F052C27FC69A1F251A8E76" colspan="1" rowspan="1">Generally last 5 tergites</td>
<td id="886E534ADD3576A103AE577D53C664D3" colspan="1" rowspan="1">Last 215 tergites</td>
<td id="FF1108DB2DD16A5932C20D59033B0BE2" colspan="1" rowspan="1">Last 410 tergites</td>
</tr>
<tr id="3C4A09C36AAD7BA58E730F189AD0CD31">
<td id="07FBB7EE371DF38C36371A5E49BB223D" colspan="1" rowspan="1">Coxopleural process spines</td>
<td id="CD078D766554A6EBD1624857E035C152" colspan="1" rowspan="1">5</td>
<td id="45C909F289CDD4F104C9DD128C06C0B1" colspan="1" rowspan="1">4</td>
<td id="36B1E330F7513A3BD55EE65E81C2A76F" colspan="1" rowspan="1">26</td>
<td id="1A36DEB3FF283297DC90E5891FCDCF42" colspan="1" rowspan="1">(3) 45</td>
</tr>
<tr id="CFB0F2C93028F33EF27A3057FB843416">
<td id="7672E51A50AE9847DA988F7F85E59751" colspan="1" rowspan="1">Lateral coxopleural spine</td>
<td id="E790C46522044AFD300D3744E4905960" colspan="1" rowspan="1">Present</td>
<td id="4009A949605ABD45792D0716A70CE34E" colspan="1" rowspan="1">Present or absent</td>
<td id="2BDB3FB81217B5E1644371B0FD679531" colspan="1" rowspan="1">Present or absent</td>
<td id="ACC4833B0C52E3FF675669499D93A7ED" colspan="1" rowspan="1">Present</td>
</tr>
<tr id="645A8FFAE515A255B4DC4B7401101D4D">
<td id="E1FACD48421021E1D5F9F663D0236219" colspan="1" rowspan="1">Leg 20 tarsal spur</td>
<td id="46D14341B09C82E9138EF1676A8011F0" colspan="1" rowspan="1">Present</td>
<td id="28D327603CF14584448FDB9DBDE6EC54" colspan="1" rowspan="1">Absent</td>
<td id="A56473049125B24FD43399CC950D9DF3" colspan="1" rowspan="1">Absent</td>
<td id="D70C04A6BC072ABE183464D8D4C3674C" colspan="1" rowspan="1">Absent</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="8384F17752A64028CDF460BE0950F7E0">
Additionally, specimens were compared to the original description of the morphologically similar and geographically proximate
<taxonomicName id="CE44CD8099F701A7160A896A31830621" authority="Kronmuller, 2010" authorityName="Kronmuller" authorityYear="2010" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="antananarivoensis">
<emphasis id="A2717F00856563EAADB976B8BBD3A49F" italics="true">Scolopendra antananarivoensis</emphasis>
<bibRefCitation id="72222D035475A75B3E902D3F72AB8AA4" author="Kronmüller C" journalOrPublisher="Spixiana" pagination="281-288" refId="B45" refString="Kronmuller C (2010) Scolopendra antananarivoensis spec. nov. - a new species of Scolopendra Linnaeus, 1758 related to Scolopendra morsitans Linnaeus, 1758 from Madagascar (Myriapoda, Chilopoda, Scolopendridae). Spixiana 33: 281-288." title="Scolopendra antananarivoensis spec. nov. - a new species of Scolopendra Linnaeus, 1758 related to Scolopendra morsitans Linnaeus, 1758 from Madagascar (Myriapoda, Chilopoda, Scolopendridae)." volume="33" year="2010">Kronmüller, 2010</bibRefCitation>
</taxonomicName>
. Material from the Aldabra Atoll did not exhibit the characters given as diagnostic for
<taxonomicName id="85C1C40DE001AA6E974C563070FD47EC" authorityName="Kronmuller" authorityYear="2010" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="antananarivoensis">
<emphasis id="51991B800A6389C0A368653C3AC0659F" italics="true">S. antananarivoensis</emphasis>
</taxonomicName>
, lacking a longitudinal median depression on sternite 21 and not having a distinctly more elongate coxopleural process (
<bibRefCitation id="F53B10B131ED91F2EF75C19F89097070" author="Kronmuller" firstAuthor="Kronmuller" journalOrPublisher="Spixiana" pagination="281-288" refId="B45" refString="Kronmüller C (2010) Scolopendra antananarivoensis spec. nov. a new species of Scolopendra Linnaeus, 1758 related to Scolopendra morsitans Linnaeus, 1758 from Madagascar (Myriapoda, Chilopoda, Scolopendridae). Spixiana 33: 281288." title="Scolopendra antananarivoensis spec. nov. - a new species of Scolopendra Linnaeus, 1758 related to Scolopendra morsitans Linnaeus, 1758 from Madagascar (Myriapoda, Chilopoda, Scolopendridae)." volume="33" year="2010">Kronmüller 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,445 @@
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<treatment id="5BC7E5A96EDC5799B2F09C1E633A4CE6" ID-arpha="5BC7E5A9-6EDC-5799-B2F0-9C1E633A4CE6" LSID="urn:lsid:plazi:treatment:5BC7E5A96EDC5799B2F09C1E633A4CE6" httpUri="http://treatment.plazi.org/id/5BC7E5A96EDC5799B2F09C1E633A4CE6" scope_class="Chilopoda" scope_family="Geophilidae" scope_order="Geophilomorpha">
<subSubSection id="SECID0EJLBG" type="nomenclature">
<paragraph id="FCB8BE9856C6C334F649E56B68AD876F">
Genus
<taxonomicName id="E10CA5C56AA80616A1325C844AA4E35C" ID-CoL="83QR" authority="Chamberlin, 1920" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="2462B38E731AEA8196559E8703A8A016" italics="true">Tuoba</emphasis>
<bibRefCitation id="F15E52937F5556D7653FB4B352E01720" author="Chamberlin RV" journalOrPublisher="Bulletin of the Museum of Comparative Zoology, Harvard College" pagination="1-269" refId="B18" refString="Chamberlin RV (1920) The Myriopoda of the Australian region. Bulletin of the Museum of Comparative Zoology, Harvard College 64: 1-269." title="The Myriopoda of the Australian region." volume="64" year="1920">Chamberlin, 1920</bibRefCitation>
</taxonomicName>
</paragraph>
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<subSubSection id="SECID0EJLBG" type="reference_group">
<paragraph id="D23CEEBEB5752512768D3E3ECFC93318">
<treatmentCitationGroup id="1770FA0420EA825CA5B96B8B2CAF6B56">
<treatmentCitation id="71ECF9CDEAFDC856E341314D1F3B4DF7" author="Lawrence RF" authority="Lawrence, 1960" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" journalOrPublisher="Faune de Madagascar" kingdom="Animalia" order="Geophilomorpha" pagination="1-123" phylum="Arthropoda" rank="genus" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." status="syn. nov." title="Myriapodes - Chilopodes." treatmentTaxon="Hovanyx" volume="12" year="1960">
<taxonomicName id="C6BEA5490E40296C4F1479DA0116FE2A" ID-CoL="4YYV" authority="Lawrence, 1960" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus" status="syn. nov.">
<emphasis id="B0C1447E65D4FAD49708B0FF9DA17C8E" bold="true" italics="true">Hovanyx</emphasis>
<bibRefCitation id="8A9D15FEB8148DD502589D6AFB837AC0" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
</taxonomicName>
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,
<taxonomicNameLabel id="35AC1DB6FB9054FA58623069B4F676AF">syn. nov.</taxonomicNameLabel>
</treatmentCitationGroup>
</paragraph>
<paragraph id="49A73E6A4B616A266CF2FFF205A65858">
<treatmentCitationGroup id="03CF30B6392347BD2E31482E88837586">
<treatmentCitation id="19E2B398FA833D2234797A15ED518E53" DOI="10.26515/rzsi/v31/i4/1929/162537" ID-CoL="5TXT" author="Silvestri F" authority="Silvestri, 1929" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" issue="4" journalOrPublisher="Records of the Indian Museum, Calcutta" kingdom="Animalia" order="Geophilomorpha" pagination="263-267" phylum="Arthropoda" rank="genus" refId="B94" refString="Silvestri F (1929) Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India). Records of the Indian Museum, Calcutta 31 (4): 263-267. https://doi.org/10.26515/rzsi/v31/i4/1929/162537" status="syn. nov." title="Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India)." treatmentTaxon="Mixophilus" volume="31" year="1929">
<taxonomicName id="89726F81C32A1165D4C2D2CFE1650E20" ID-CoL="5TXT" authority="Silvestri, 1929" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus" status="syn. nov.">
<emphasis id="7A783AC82D881491406AB845EE885DB8" italics="true">Mixophilus</emphasis>
<bibRefCitation id="AFC7E93257EC21253D1238DBE6488961" DOI="10.26515/rzsi/v31/i4/1929/162537" author="Silvestri F" issue="4" journalOrPublisher="Records of the Indian Museum, Calcutta" pagination="263-267" refId="B94" refString="Silvestri F (1929) Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India). Records of the Indian Museum, Calcutta 31 (4): 263-267. https://doi.org/10.26515/rzsi/v31/i4/1929/162537" title="Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India)." volume="31" year="1929">Silvestri, 1929</bibRefCitation>
</taxonomicName>
</treatmentCitation>
<taxonomicNameLabel id="283B03290AAE247290B5DA70F54474F6">syn. nov.</taxonomicNameLabel>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EDSBG" type="remarks">
<paragraph id="4FD1B49D2251C71AD9DF27BB7BA91D67">
<heading id="838B3275C53BDD6972CCBC5DD4924C41" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="4ADE348A1E7ED66930F4947E61C49C85">
The monotypic genus
<taxonomicName id="1EAF569F7D30EEDF3B59D3D691BDD96C" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="04C7F72D9E7472CB5267F18C61FEDE3B" italics="true">Mixophilus</emphasis>
</taxonomicName>
was erected by
<bibRefCitation id="09376AA4028570D99FAB823D844C5B8A" DOI="10.26515/rzsi/v31/i4/1929/162537" author="Silvestri F" firstAuthor="Silvestri" issue="4" journalOrPublisher="Records of the Indian Museum, Calcutta" pagination="263-267" refId="B94" refString="Silvestri F (1929) Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India). Records of the Indian Museum, Calcutta 31 (4): 263267. https://doi.org/10.26515/rzsi/v31/i4/1929/162537" title="Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India)." volume="31" year="1929">Silvestri (1929)</bibRefCitation>
to place a new species of geophilomorph sampled from riparian sites in Madras (Chennai), Southern
<collectingCountry id="CBB7601B001E000C02E88331F24AEB74" name="India">India</collectingCountry>
. The original description of
<taxonomicName id="34C7781F48B78A3964AA0A2C060BC1F9" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="24F6A021864F525E998982F98A5EB8D2" italics="true">Mixophilus indicus</emphasis>
</taxonomicName>
includes illustrations of the head, forcipular apparatus and ultimate leg-bearing segment as well as detailed ecological observations concerning its preferred microhabitats in the type locality. Subsequent physiological investigations revealed a modified tracheal system, comprising possible adaptations to immersion for long periods of time (
<bibRefCitation id="CFAF918A07BF544B93A7EB8A3760CEB1" author="Rajulu" firstAuthor="Rajulu" issue="17" journalOrPublisher="Current Science" pagination="397-398" refId="B88" refString="Rajulu GS (1970) Tracheal pulsation in a marine centipede Mixophilus indicus. Current Science 39 (17): 397398." title="Tracheal pulsation in a marine centipede Mixophilus indicus." volume="39" year="1970">Rajulu 1970</bibRefCitation>
). An affinity to
<taxonomicName id="6CD9EE8D625006D46E30FC8DA9F41502" class="Chilopoda" family="Dignathodontidae" genus="Henia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="764D5553635D3149C6631E591E807375" italics="true">Henia</emphasis>
</taxonomicName>
and
<taxonomicName id="273945D6DBB033E49F6E7B1616F77EEE" authorityName="Meinert" authorityYear="1870" class="Chilopoda" family="Dignathodontidae" genus="Chaetechelyne" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="60DCA1171505DC68A49761C253391244" italics="true">Chaetechelyne</emphasis>
</taxonomicName>
(
<taxonomicName id="7D6C9C1E1B73B7F3DFB1DA1AC6DB143F" class="Chilopoda" family="Dignathodontidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Dignathodontidae</taxonomicName>
) was suggested, however, within the same section
<bibRefCitation id="DD2AED1D1FDD99F4B523D314D1763803" DOI="10.26515/rzsi/v31/i4/1929/162537" author="Silvestri F" firstAuthor="Silvestri" issue="4" journalOrPublisher="Records of the Indian Museum, Calcutta" pagination="263-267" refId="B94" refString="Silvestri F (1929) Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India). Records of the Indian Museum, Calcutta 31 (4): 263267. https://doi.org/10.26515/rzsi/v31/i4/1929/162537" title="Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India)." volume="31" year="1929">Silvestri (1929)</bibRefCitation>
pointed to differences between
<taxonomicName id="E199DDCC1758C574F6A2E9AC1A67B052" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="654BAD8C6479B740E24C971340B1C74E" italics="true">Mixophilus</emphasis>
</taxonomicName>
and both of these genera in “ the structure of the labrum ” (tripartite in
<taxonomicName id="4F9C795F3634033157284F50F64CB38C" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="656E106A25C9E9D3B91C1DB6D2D70B93" italics="true">M. indicus</emphasis>
</taxonomicName>
but unipartite in
<taxonomicName id="449A5229257CAB16DDD4E557A08F0A69" class="Chilopoda" family="Dignathodontidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Dignathodontidae</taxonomicName>
), “ the distribution of sternal pores ” (transverse band in
<taxonomicName id="FD82AF71A1EC1E627A8D8FD17241A5C8" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="FAAC72CB2AF375AC14C2374A850B9D00" italics="true">M. indicus</emphasis>
</taxonomicName>
, medial sub-circular / elliptical field in
<taxonomicName id="511E9546A25157E58A3373EC71DFDB18" class="Chilopoda" family="Dignathodontidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Dignathodontidae</taxonomicName>
), in the last leg-bearing segment (telopodite composed of 7 articles in
<taxonomicName id="6AFA6744C5604CB9E0469A3DD428DAE6" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="21EE8EAEBC919C2F803AF2B1A54C2125" italics="true">M. indicus</emphasis>
</taxonomicName>
but
<quantity id="851064FC79473E565AB7B4CA72EAD779" metricMagnitude="-1" metricUnit="m" metricValue="1.524" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="EDDD6F868521C9ED5C7C002D334ACC1B" class="Chilopoda" family="Dignathodontidae" genus="Henia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="F55901D287B9AD29BD774689276D916E" italics="true">Henia</emphasis>
</taxonomicName>
), and in the elongation of the forcipular pretergite (longer in
<taxonomicName id="D42D3227366590EEE23FA2F442A1D9BC" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="E499D08F53448FF3C74723F7C8964678" italics="true">M. indicus</emphasis>
</taxonomicName>
than in
<taxonomicName id="038A16CAFE039C7DB365F21C978CCA2C" authorityName="Meinert" authorityYear="1870" class="Chilopoda" family="Dignathodontidae" genus="Chaetechelyne" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="5564218D31D765496A90088511C9C6A8" italics="true">Chaetechelyne</emphasis>
</taxonomicName>
). The structure of the labrum described and illustrated for
<taxonomicName id="8D394FAC62925DCC1C4D816EBD6877DD" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="C8FA084B1BBBC9C52521E2C0DE0E87FC" italics="true">M. indicus</emphasis>
</taxonomicName>
, comprising three conspicuous pieces, suggests a close affinity to other members of
<taxonomicName id="109BDBB1F1EEB2AA2AAEF9127E30C980" baseAuthorityName="Bonato" baseAuthorityYear="2011" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Geophilidae</taxonomicName>
s. str. An estuarine habitat preference and multiple morphological characters show a near complete overlap between the diagnoses of
<taxonomicName id="58CEB23CC9830253B665A5B1898B969E" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="19457184F7E941FA95EEA1224252F8B4" italics="true">Tuoba</emphasis>
</taxonomicName>
and
<taxonomicName id="DBF98E01E1C1B803B98B7461468FFDE1" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="3F7F126901683BCF217B08E50E4EDDDE" italics="true">Mixophilus</emphasis>
</taxonomicName>
(Table
<tableCitation id="BF1CDB803B6C5C7E49496DBAA7EA9AF3" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Taxonomically informative invariant and variable morphological characters for the genera Hovanyx, Mixophilus, and Tuoba based on literature data. Characters in boldface represent putative morphological differences." httpUri="http://table.plazi.org/id/B4073602D8AB8AC245E0D42FF98F554B" tableUuid="B4073602D8AB8AC245E0D42FF98F554B">3</tableCitation>
). Complete chitin lines in
<taxonomicName id="B025481DAFCEAC083CF7F37D61366CBE" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="D17AA51278F6D6E34A8249EF8B2BD73F" italics="true">M. indicus</emphasis>
</taxonomicName>
may indicate intraspecific or interspecific variability which has been recorded in
<taxonomicName id="5C49B3732C1AE017710C29239D54ABB8" authority="(Jones 1998)" baseAuthorityName="Jones" baseAuthorityYear="1998" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="7B64F62AA93F046FB60E864B3C726567" italics="true">Tuoba</emphasis>
(
<bibRefCitation id="EB1E83860C72B54E7AED6146EC682103" author="Jones" firstAuthor="Jones" journalOrPublisher="Records of the Western Australian Museum" pagination="333-346" refId="B42" refString="Jones RE (1998) On the species of Tuoba (Chilopoda: Geophilomorpha) in Australia, New Zealand, New Caledonia, Solomon Islands and New Britain. Records of the Western Australian Museum 18: 333346." title="On the species of Tuoba (Chilopoda: Geophilomorpha) in Australia, New Zealand, New Caledonia, Solomon Islands and New Britain." volume="18" year="1998">Jones 1998</bibRefCitation>
)
</taxonomicName>
or artifacts of examination under light microscopy. In light of this re-evaluation of its original description, we propose reassignment of
<taxonomicName id="63069FDAFD531EFB3F4C96C85DA96908" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="31233F516F0BBBD58A0DD165FD002165" italics="true">M. indicus</emphasis>
</taxonomicName>
to
<taxonomicName id="D85AACC68425A02AF33FAAEE6CCD2700" baseAuthorityName="Jones" baseAuthorityYear="1998" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="D98F8C165CA5C0516BE681414BEC5904" italics="true">Tuoba</emphasis>
</taxonomicName>
considering the available data on its morphology, with
<taxonomicName id="300A6228D52A3060CE98B0EAFD92B267" authorityName="Popovici &amp; Edgecombe" authorityYear="2025" baseAuthorityName="Popovici &amp; Edgecombe" baseAuthorityYear="2025" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indica" status="comb. nov.">
<emphasis id="E7676DA272C6249CB5F48D5364D42E59" italics="true">Tuoba indica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5D2BC6B10AC07909B1D642EA05B36D99" rank="species">comb. nov.</taxonomicNameLabel>
as the valid name for specimens on which its original description was based. Consequently, we propose that
<taxonomicName id="3B961BD56A083FF3BE2268B8C656C892" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="B43840038EDEE7649CCF305B7B0CF545" italics="true">Mixophilus</emphasis>
</taxonomicName>
is the junior subjective synonym of
<taxonomicName id="58C5940FA4704E70B825B089E9CEE9D6" baseAuthorityName="Jones" baseAuthorityYear="1998" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus" status="syn. nov.">
<emphasis id="B2FE2040D8CC38F7E9AC3FD78E9F0228" italics="true">Tuoba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C37032B6BA85D74595C5B9B52FC70655">syn. nov.</taxonomicNameLabel>
Similarities between
<taxonomicName id="2BA988D948FDD4497E6EA96CF5B1D76D" authorityName="Popovici &amp; Edgecombe" authorityYear="2025" baseAuthorityName="Popovici &amp; Edgecombe" baseAuthorityYear="2025" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indica" status="comb. nov.">
<emphasis id="A49CFFFA4977A9CD2201FDD199D19D19" italics="true">Tuoba indica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ED1C10440CC1E8CEF678946D3D104740" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="786750C917F2A0039B98543E34991E1F" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="1AC30AA1413190F1911FCACB9EAFE7D7" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
exist in elongation of the ultimate legs and reduction of the second maxillary pretarsus, however these characters are shared by several species of
<taxonomicName id="08B8EDAB2AD391C66F96634E662ABE46" baseAuthorityName="Jones" baseAuthorityYear="1998" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="C79FAAB73CA57A6F336176514507DF3B" italics="true">Tuoba</emphasis>
</taxonomicName>
. The incomplete original description provided by Silvestri does not allow for definitive assignment of
<taxonomicName id="817BFCE7DBA26032EA5B4BE70F0A42D3" authorityName="Popovici &amp; Edgecombe" authorityYear="2025" baseAuthorityName="Popovici &amp; Edgecombe" baseAuthorityYear="2025" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="indica" status="comb. nov.">
<emphasis id="17629FE986E90E6B8817EB388FE141D0" italics="true">Tuoba indica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="36E00815B2D7CF127792614FC8D70F42" rank="species">comb. nov.</taxonomicNameLabel>
to another species of
<taxonomicName id="24F448FB24C657B0E0ECAC93A3C23E65" baseAuthorityName="Jones" baseAuthorityYear="1998" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="D689891D85C8EE6D7278A2B6605A8408" italics="true">Tuoba</emphasis>
</taxonomicName>
until the type material can be adequately re-described.
</paragraph>
<caption id="B4073602D8AB8AC245E0D42FF98F554B" ID-Table-UUID="B4073602D8AB8AC245E0D42FF98F554B" httpUri="http://table.plazi.org/id/B4073602D8AB8AC245E0D42FF98F554B" startId="T3" targetIsTable="true">
<paragraph id="492F6B4EAAF2C539554D9058C6B0A6DB">
<label id="9CB8535F9103A12DFA1B7364AB5F8CB1">Table 3.</label>
</paragraph>
<paragraph id="3AF3793977BF820B3DED3C85302F376D">
Taxonomically informative invariant and variable morphological characters for the genera
<taxonomicName id="40E27632C277EBD6B15CE7C3591F8E41" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="1F00EEBFBD454865D491CED98463916C" italics="true">Hovanyx</emphasis>
</taxonomicName>
,
<taxonomicName id="76AEC4AA62736AE9D9624A707712BB9C" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="FFE7178ECEC717389D4AB5A5C53D2A5C" italics="true">Mixophilus</emphasis>
</taxonomicName>
, and
<taxonomicName id="1DBB676AF3C51076F136807CBB79DBE4" baseAuthorityName="Jones" baseAuthorityYear="1998" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="4004D2E8BF85B8854DB4021AAA4404CD" italics="true">Tuoba</emphasis>
</taxonomicName>
based on literature data. Characters in boldface represent putative morphological differences.
</paragraph>
</caption>
<paragraph id="E99A2C79937F1D68B8F98B07D7ACD0EA">
<table id="TID0EPUDI" rules="all">
<tbody id="1DF52992DF18A5713438F9976F1FEA25">
<tr id="FE66B1652D9FD989621FA5CCEAB564A6">
<th id="3B30A1472DD3F78511E8F9EC1D86FF47" colspan="1" rowspan="1">Character</th>
<th id="F2A3C189C4DE06AB480C12B2660D5120" colspan="1" rowspan="1">
<taxonomicName id="331332BF41E6FD6DF71A1073DFA76453" authority="Chamberlin, 1920" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="F82F6FE2318A6315ECD22F4535D6AED8" italics="true">Tuoba</emphasis>
<bibRefCitation id="E664D487AA5BDC3251CE50B57DE105F8" author="Chamberlin RV" journalOrPublisher="Bulletin of the Museum of Comparative Zoology, Harvard College" pagination="1-269" refId="B18" refString="Chamberlin RV (1920) The Myriopoda of the Australian region. Bulletin of the Museum of Comparative Zoology, Harvard College 64: 1-269." title="The Myriopoda of the Australian region." volume="64" year="1920">Chamberlin, 1920</bibRefCitation>
</taxonomicName>
</th>
<th id="1493509CE9D78B74ADEB5A7315675159" colspan="1" rowspan="1">
<taxonomicName id="C35D8CBA195ED910EF1FFFE5450A1EEE" authority="Lawrence, 1960" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="EA8329FBAE4A842552F5E89A8E07660B" italics="true">Hovanyx</emphasis>
<bibRefCitation id="8E6DC9E8195CFB777F7623C713D3C12A" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
</taxonomicName>
</th>
<th id="772AD760B2D50FDAF076F14722628B0A" colspan="1" rowspan="1">
<taxonomicName id="B0C39830A27B863094076E3D609E8212" authority="Silvestri, 1929" authorityName="Silvestri" authorityYear="1929" class="Chilopoda" family="Geophilidae" genus="Mixophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="94DAE9FC3D28E09F401213B5FAF517F1" italics="true">Mixophilus</emphasis>
<bibRefCitation id="F683B892DCE11E933AD4C3BB7948F32A" DOI="10.26515/rzsi/v31/i4/1929/162537" author="Silvestri F" issue="4" journalOrPublisher="Records of the Indian Museum, Calcutta" pagination="263-267" refId="B94" refString="Silvestri F (1929) Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India). Records of the Indian Museum, Calcutta 31 (4): 263-267. https://doi.org/10.26515/rzsi/v31/i4/1929/162537" title="Description of a new genus and species of Geophilidae (Myriapoda Chilopoda) from Madras (India)." volume="31" year="1929">Silvestri, 1929</bibRefCitation>
</taxonomicName>
</th>
</tr>
<tr id="92459FD09ABAC3E81073DA517FC529A8">
<td id="4BD682B08B7076BDF121626C3F9AE7CB" colspan="1" rowspan="1">Head shape</td>
<td id="FC3171FDA44B567831A4C8E2FE7D32CA" colspan="1" rowspan="1">Subquadrate</td>
<td id="925A727AC9FCC17CF2EA6AA3A6207361" colspan="1" rowspan="1">
<emphasis id="E9463B84F2CB0F4C5D0C4AD1DE182725" bold="true">Longer than wide</emphasis>
</td>
<td id="F796E53D171E4C9E1AFA9764B4BEC89A" colspan="1" rowspan="1">Subquadrate</td>
</tr>
<tr id="01C6AD52E267E173E2F9BC05A13A863B">
<td id="7B682B38C56319AF03D1D937D5DEDB86" colspan="1" rowspan="1">Setation of clypeus</td>
<td id="69EC613B6B478F56FDC5B9045A243EC1" colspan="1" rowspan="1">Three pairs of setae medially, flanked by a group of 24 setae on each side</td>
<td id="D66421E27BA0F02300068FA99256D463" colspan="1" rowspan="1">1011 setae</td>
<td id="5562AFD7A03ABF015AFE9AE38B443C9E" colspan="1" rowspan="1">Four pairs of setae medially, flanked by a group of 3 or 4 setae on each side</td>
</tr>
<tr id="0FC7B14E86407D05299D216B6BB47B2A">
<td id="B6732E9DD09AFE546CD9A20819CB1CB4" colspan="1" rowspan="1">Medial piece of labrum. Orientation of tubercles</td>
<td id="5533FCD3255012DE76353DFDF2187E68" colspan="1" rowspan="1">Anteriorly recurved (variable)</td>
<td id="12834A985CF61F6DE1C30CE56B9057F5" colspan="1" rowspan="1">Anteriorly recurved</td>
<td id="F3106187EC3C25BF0300F2A3FD158A36" colspan="1" rowspan="1">Anteriorly recurved</td>
</tr>
<tr id="803648175CF0F528817BA8301672D950">
<td id="1DADBD506E65B6D66EF848C972F59844" colspan="1" rowspan="1">Side pieces of labrum</td>
<td id="C34A0750DB859EE603D85AC5CD659561" colspan="1" rowspan="1">With variable number of tubercles or plumose setae</td>
<td id="BB8C7AE35D1B23CBEDF196504BF190D2" colspan="1" rowspan="1">-</td>
<td id="467343296FE672BE78446C8E235485FD" colspan="1" rowspan="1">Without tubercles or plumose setae</td>
</tr>
<tr id="20F149E43E8E3E5E1D692F4962C885F2">
<td id="7AA4C487190A6453E203A4848AA89D97" colspan="1" rowspan="1">Maxillae I lappets</td>
<td id="2D37A72E0DFC0C63D2E214BEEE8D492F" colspan="1" rowspan="1">Absent. External corners with spiniform cuticular projections</td>
<td id="9A57E93CA7DDCB9ED175FE9DE8421929" colspan="1" rowspan="1">Absent. External corners with spiniform cuticular projections</td>
<td id="64781D56A937A97081A4E5C59123CEA0" colspan="1" rowspan="1">Absent</td>
</tr>
<tr id="F234BBA798632CBA6E5D89664E6CF064">
<td id="3991A507526DDBD0614EEF2D2C879610" colspan="1" rowspan="1">Maxillae II pretarsus</td>
<td id="A703D2C69AE07C8278ED1DB8A89FBBE5" colspan="1" rowspan="1">Simple, claw-shaped. Variably reduced in size</td>
<td id="F4E547D1B2F7DBE73B7EF573430AFD5C" colspan="1" rowspan="1">
<emphasis id="23BD76D3D21E728AD33780030338A96A" bold="true">Simple, claw-shaped</emphasis>
</td>
<td id="1BDB25360409AA22888CFB0A8442579A" colspan="1" rowspan="1">Simple, claw-shaped. Reduced in size</td>
</tr>
<tr id="E5F10AEC7259F48A22FCE4E4D055140F">
<td id="E070A3BC85A2CA037FE3372DBF8819B5" colspan="1" rowspan="1">Forcipular coxosternite chitin lines</td>
<td id="7D897F1FD0B767F8F487DC7878030AFF" colspan="1" rowspan="1">Complete or nearly complete</td>
<td id="67C750E30E7AA57C505B352B20FC5E8A" colspan="1" rowspan="1">Vanishing before reaching the condyles</td>
<td id="D24E7B6C53308E15C6C4B073C0424D0E" colspan="1" rowspan="1">Complete</td>
</tr>
<tr id="83ADF3DBBEEAB1FCB199CB44127DEFDE">
<td id="EBBE21EBCD9509ABF530BC8C2EEF83BB" colspan="1" rowspan="1">Denticle at the base of the tarsungulum</td>
<td id="2ED7A3EC3977A78A84E88FAA162AA481" colspan="1" rowspan="1">Present</td>
<td id="3301B68209AFEF4542DADBC823CF0145" colspan="1" rowspan="1">Present</td>
<td id="B5ED9FBC9425ECBA95BE3783641CF4CE" colspan="1" rowspan="1">Present</td>
</tr>
<tr id="46B246C017D5F71AE565CD57A9BFC065">
<td id="4886C121E3B182850757DE600163B6A4" colspan="1" rowspan="1">Carpophagus structures</td>
<td id="181EC7E62CBF8D80A185A9740ABF8ABC" colspan="1" rowspan="1">
<emphasis id="3B9EFEEDE1E1661CA70A99492863834D" bold="true">Present</emphasis>
</td>
<td id="A82FF187E4317D3D62F3CFD5A63C028E" colspan="1" rowspan="1">-</td>
<td id="9D7D3F87083121E9C51722917C40E3D9" colspan="1" rowspan="1">-</td>
</tr>
<tr id="8459371C34EA7F0694DE123F27830BC3">
<td id="DDDFF69351606399908E573A6540CFB7" colspan="1" rowspan="1">Metasternal pore field shape</td>
<td id="8D1B0616DA0AF1F58ACFB7E5676A4292" colspan="1" rowspan="1">Transverse band</td>
<td id="0DF64828AD376B12287457465AC12A48" colspan="1" rowspan="1">Transverse band. Divides on LBS XI XII</td>
<td id="4E9EFF59066BE0B16890842A01791D94" colspan="1" rowspan="1">Transverse band (medially constricted)</td>
</tr>
<tr id="5714D293579AAEFCF6ECDBA44C025294">
<td id="4E84641650AB9521EBCE2DA84C7FCD69" colspan="1" rowspan="1">Pretarsus of walking legs</td>
<td id="52C8DDC6C4356B46BA0B8784E96AD6C5" colspan="1" rowspan="1">Distinctly elongate</td>
<td id="831406A58E6F6878295391F0A1B88090" colspan="1" rowspan="1">Distinctly elongate</td>
<td id="707C7E12B3101784DEA20630BF5A0E0F" colspan="1" rowspan="1">Distinctly elongate</td>
</tr>
<tr id="52DB95F0D685D9A53A53429A200E0507">
<td id="7B24CB3F335057D4243B27FD2554D499" colspan="1" rowspan="1">LLBS metasternite shape</td>
<td id="0A78A140EC0C400347194468C839E925" colspan="1" rowspan="1">Wider than long, trapezoidal</td>
<td id="75D53B8A5A8239EE58E7558D2E1A31AF" colspan="1" rowspan="1">Wider than long, trapezoidal</td>
<td id="4A1B0B50249E13C957178782E2CC0F86" colspan="1" rowspan="1">Wider than long, trapezoidal</td>
</tr>
<tr id="D7D7DB511C0A070A3CDE41C7EB3866FE">
<td id="839948C15134A02358AD86FB5B1B9A4B" colspan="1" rowspan="1">Coxal organs</td>
<td id="5EE73C5E0567D54FD477D778729A65C7" colspan="1" rowspan="1">Multiple opening in single pit</td>
<td id="0DCA12356FA2880A4B431D67338136C8" colspan="1" rowspan="1">
<emphasis id="66E68165B6342DC0AC3B714C193661EE" bold="true">Absent</emphasis>
</td>
<td id="D8AAB4C86B76B5934D33E5A186978D8D" colspan="1" rowspan="1">Multiple opening in single pit</td>
</tr>
<tr id="401425E6616D7417637DC66F0CB04C20">
<td id="7F571898B4F08CFB2D2E47BCDDA7C725" colspan="1" rowspan="1">LLBS pretarsus</td>
<td id="25E19B55C4FBCC521919DB27AB2141DA" colspan="1" rowspan="1">Simple, claw-shaped</td>
<td id="3EEDAFD38912A12243B80D825479F070" colspan="1" rowspan="1">Simple, claw-shaped</td>
<td id="5C3F1B3ED96E3835DF3BDF866F598825" colspan="1" rowspan="1">Simple, claw-shaped</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="C2AFEF0E86E3525D4D2F04CFC766FEBC">
Similarly, the genus
<taxonomicName id="60B8ACEA8A41CA2A7AD606CA64379D93" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="51836E15EF17DEAFBB3E5133592551F5" italics="true">Hovanyx</emphasis>
</taxonomicName>
was erected for the species
<taxonomicName id="D74D0F2EA876EFF2CEE89B92B934E57D" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="waterloti">
<emphasis id="4A5945043B1B56BE3518ABF784BBC1FF" italics="true">Hovanyx waterloti</emphasis>
</taxonomicName>
, described in
<bibRefCitation id="15198D6BC775F4E37B99A6D3FABDE629" author="Lawrence RF" firstAuthor="Lawrence's" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes Chilopodes. Faune de Madagascar 12: 1123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrences (1960)</bibRefCitation>
catalogue of Malagasy centipedes. An affinity to
<taxonomicName id="47304B2C08AE7715D8B52B97E147E72B" class="Chilopoda" family="Dignathodontidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Dignathodontidae</taxonomicName>
was again proposed based on similarities in the structure of the labrum, originally described for
<taxonomicName id="5378F7B99E66129462EFF64D791D4870" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="waterloti">
<emphasis id="98474888714580F9AAB0D9E46D648A88" italics="true">H. waterloti</emphasis>
</taxonomicName>
as composed of a single piece and bearing a small number of rudimentary, anteriorly oriented tubercles (“ Labre […] à dents pas très distinctes, quatre courtes dents triangulaires dirigées vers lavant, de chaque côté. ”). The diagnosis of
<taxonomicName id="1D6CB39B6FFD81C4D281A6F64AEBB3CE" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="B673FDFC1421CABBE0464BB4569BC2F0" italics="true">Hovanyx</emphasis>
</taxonomicName>
singled out the absence of coxal organs (“ […] diffère de tous les autres membres de la sous-famille des Dignathodontinæ par labsence de pores aux pattes terminales. ”) as the main distinguishing trait separating it from all other dignathodontid genera. However, both the original description and accompanying illustrations suggest a closer affinity to
<taxonomicName id="942D13DC4799987902D9A269258ADFE9" baseAuthorityName="Bonato" baseAuthorityYear="2011" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Geophilidae</taxonomicName>
s. str., as multiple other morphological characters (shape of head, forcipular coxosternite, metasternal pore fields) are characteristic of the
<taxonomicName id="D8DC45E07E71C6AB2EFB163742824E07" baseAuthorityName="Bonato" baseAuthorityYear="2011" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Geophilidae</taxonomicName>
rather than the
<taxonomicName id="F1CB884DA445ABB17F3C02E9914FCCAE" class="Chilopoda" family="Dignathodontidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Dignathodontidae</taxonomicName>
, and subsequent taxonomic revision of both families placed
<taxonomicName id="140584DEAA6213131017463F17F26D35" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="3225A1E0B674201BB0562B6F963F1134" italics="true">Hovanyx</emphasis>
</taxonomicName>
under
<taxonomicName id="6E6399F8814EAD544AC3623CF83C4224" authority="(Bonato 2011)" baseAuthorityName="Bonato" baseAuthorityYear="2011" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">
Geophilidae (
<bibRefCitation id="9010CCF0213D76A8387C904992905068" DOI="10.1163/9789004188266" author="Bonato" firstAuthor="Bonato" journalOrPublisher="Leiden, The Netherlands: Brill" refId="B9" refString="Bonato L (2011) Chilopoda Taxonomic Overview. Order Geophilomorpha. In: Minelli A (Eds) Treatise on Zoology Anatomy, Taxonomy, Biology. The Myriapoda, Vol. 1. Leiden, The Netherlands: Brill, 538 pp. https://doi.org/10.1163/9789004188266" title="Chilopoda - Taxonomic Overview. Order Geophilomorpha. In: Minelli A (Eds) Treatise on Zoology - Anatomy, Taxonomy, Biology. The Myriapoda, Vol. 1." year="2011">Bonato 2011</bibRefCitation>
)
</taxonomicName>
. Furthermore, the incomplete original description overlaps almost entirely with that of
<taxonomicName id="D431985ABA1F54086D324460191B5073" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="E40E758D2F852ED7AC4DA92771ECA62F" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
, a wide-ranging geophilid encountered in littoral sites in the
<collectingCountry id="739C0DFACAD4C7CD285F489142D8DEF9" name="Seychelles">Seychelles</collectingCountry>
(
<bibRefCitation id="E361836A8AFA9CF340C68920FD9F832A" author="Bonato and Minelli" firstAuthor="Bonato" journalOrPublisher="Phelsuma" pagination="9-38" refId="B11" refString="Bonato L, Minelli A (2010) The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha). Phelsuma 18: 938." title="The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha)." volume="18" year="2010">Bonato and Minelli 2010</bibRefCitation>
) and the Aldabra Atoll (present records) close to the type locality of
<taxonomicName id="414216E77601DE9F4039E01BA092ACE4" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="waterloti">
<emphasis id="F7914B2C193BB60D5C115856982F8221" italics="true">H. waterloti</emphasis>
</taxonomicName>
. Shared characters include range of leg bearing segment number (4143), morphology of the labrum, condition of chitin lines on the forcipular coxosternite (incomplete) (Table
<tableCitation id="1175AC530416F23FE9BCF11C20238ADE" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Taxonomically informative invariant and variable morphological characters for the genera Hovanyx, Mixophilus, and Tuoba based on literature data. Characters in boldface represent putative morphological differences." httpUri="http://table.plazi.org/id/B4073602D8AB8AC245E0D42FF98F554B" tableUuid="B4073602D8AB8AC245E0D42FF98F554B">3</tableCitation>
). The only putative differences between
<taxonomicName id="327B7F85A5A7CAAF4D8176EA69997B2F" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Geophilidae" genus="Hovanyx" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="waterloti">
<emphasis id="7CD42C344A6FA648130171C0DC07414F" italics="true">H. waterloti</emphasis>
</taxonomicName>
and
<taxonomicName id="1F12D74BD2B54C9B6B87F3A66066889D" authorityName="A NHMUK" baseAuthorityName="Pocock" baseAuthorityYear="1891" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="C6CAC73C000777F53AA593DBA54A8DF7" italics="true">T. sydneyensis</emphasis>
</taxonomicName>
are the unipartite labrum and absence of coxal organs in the former (Table
<tableCitation id="52B629D86DD04C6B7216B03A34B93C95" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Taxonomically informative invariant and variable morphological characters for the genera Hovanyx, Mixophilus, and Tuoba based on literature data. Characters in boldface represent putative morphological differences." httpUri="http://table.plazi.org/id/B4073602D8AB8AC245E0D42FF98F554B" tableUuid="B4073602D8AB8AC245E0D42FF98F554B">3</tableCitation>
). These may however be unreliable characters because of inadequate documentation, as published illustrations of
<taxonomicName id="BBCBCF57C1CEC31457B3BFC708BD2D30" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="594038D27063990B5E2785519B1CB9B0" italics="true">Tuoba</emphasis>
</taxonomicName>
specimens and the material here illustrated show great variability in the orientation and shape of the labral pieces, which depending on the degree of sclerotization seen in the specimen and shape and number of denticles on the side pieces, may resemble the labrum of
<taxonomicName id="042BA1600C87BF3236BC61AF09D18C41" class="Chilopoda" family="Dignathodontidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Dignathodontidae</taxonomicName>
under light microscopy. The absence of coxal organs has not been previously reported in any species of
<taxonomicName id="7BA31DD8F9E2C0E04E4A104176F1CD3F" authorityName="Chamberlin" authorityYear="1920" class="Chilopoda" family="Geophilidae" genus="Tuoba" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="33556355D0D9AE457EC4680F8C94590C" italics="true">Tuoba</emphasis>
</taxonomicName>
, however this character shows extensive ontogenetic plasticity within
<taxonomicName id="4012D5330A8321697E8DEC3249460E4F" baseAuthorityName="Bonato" baseAuthorityYear="2011" class="Chilopoda" family="Geophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Geophilidae</taxonomicName>
(
<bibRefCitation id="C7AB27DA56E3AD1562211A6307670D15" author="Horneland and Meidell" firstAuthor="Horneland" issue="3" journalOrPublisher="Soil Organisms" pagination="373-386" refId="B36" refString="Horneland EO, Meidell B (2009) Postembryonic development of Strigamia maritima (Leach, 1817) (Chilopoda: Geophilomorpha, Linotaeniidae) with emphasis on how to separate the different stadia. Soil Organisms 81 (3): 373386. https://soil-organisms.org/index.php/SO/article/view/34" title="Postembryonic development of Strigamia maritima (Leach, 1817) (Chilopoda: Geophilomorpha, Linotaeniidae) with emphasis on how to separate the different stadia." url="https://soil-organisms.org/index.php/SO/article/view/34" volume="81" year="2009">Horneland and Meidell 2009</bibRefCitation>
;
<bibRefCitation id="482C9ACB34111DD589335F4F0FF02C16" author="Gregory and Barber" firstAuthor="Gregory" journalOrPublisher="Bulletin of the British Myriapod and Isopod Group" pagination="2-15" refId="B30" refString="Gregory SJ, Barber AD (2010) Observations of a population, including juveniles, of Geophilus carpophagus Leach, 1815, sensu stricto from Oxfordshire. Bulletin of the British Myriapod and Isopod Group 24: 215." title="Observations of a population, including juveniles, of Geophilus carpophagus Leach, 1815, sensu stricto from Oxfordshire." volume="24" year="2010">Gregory and Barber 2010</bibRefCitation>
;
<bibRefCitation id="8B687D5A4BB74FEB95A0F2B31FBC374A" DOI="10.11646/zootaxa.4058.3.11" author="Tuf and Danyi" firstAuthor="Tuf" issue="3" journalOrPublisher="Zootaxa" pagination="444-450" refId="B103" refString="Tuf IH, Dányi L (2015) True identity of Folkmanovius paralellus Dobroruka (Chilopoda: Geophilomorpha: Geophilidae). Zootaxa 4058 (3): 444450. https://doi.org/10.11646/zootaxa.4058.3.11" title="True identity of Folkmanovius paralellus Dobroruka (Chilopoda: Geophilomorpha: Geophilidae)." volume="4058" year="2015">Tuf and Dányi 2015</bibRefCitation>
;
<bibRefCitation id="6F0223889215EDF7C9A892CE32468EFF" DOI="10.11646/zootaxa.4139.4.3" author="Peretti and Bonato" firstAuthor="Peretti" issue="4" journalOrPublisher="Zootaxa" pagination="499-514" refId="B78" refString="Peretti E, Bonato L (2016) Geophilus pygmaeus (Chilopoda: Geophilidae): clarifying morphology, variation and geographic distribution. Zootaxa 4139 (4): 499514. https://doi.org/10.11646/zootaxa.4139.4.3" title="Geophilus pygmaeus (Chilopoda: Geophilidae): clarifying morphology, variation and geographic distribution." volume="4139" year="2016">Peretti and Bonato 2016</bibRefCitation>
;
<bibRefCitation id="A9BA80F5986F01B5860287C24D82D2BA" DOI="10.1080/07924259.2020.1726514" author="Stojanovic" etAl="et al." firstAuthor="Stojanovic" issue="2" journalOrPublisher="Invertebrate Reproduction &amp; Development" pagination="115-125" refId="B99" refString="Stojanović DZ, Mitić BM, Dudić BD, Gedged AM, Tomić VT, Antić DŽ, Makarov SE (2020) Early development of the centipede Geophilus serbicus (Chilopoda: Geophilomorpha: Geophilidae) from the Balkan Peninsula. Invertebrate Reproduction &amp; Development 64 (2): 115125. https://doi.org/10.1080/07924259.2020.1726514" title="Early development of the centipede Geophilus serbicus (Chilopoda: Geophilomorpha: Geophilidae) from the Balkan Peninsula." volume="64" year="2020">Stojanović et al. 2020</bibRefCitation>
) and the small size of the
<typeStatus id="8AF82088EABBC051E3EAEA6DA25342FF" type="holotype">holotype</typeStatus>
and only known specimen as well as variations in clearing and position of the coxopleuron may render the pit inconspicuous.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,563 @@
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<paragraph id="5568EDF71B7B862A2CEEF729C4D113B9">
<taxonomicName id="A36E88DDB2A12EDBDDB6F66C10390B1C" authority="(Lawrence, 1960)" authorityName="NHMUK" baseAuthorityName="Lawrence" baseAuthorityYear="1960" class="Chilopoda" family="Schendylidae" genus="Ityphilus" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="5B52F87D6E6538208DB393C663790C34" italics="true">Ityphilus</emphasis>
cf.
<emphasis id="74E1976359C0536E02B6CB9E5C75EA2C" italics="true">taeniaformis</emphasis>
(
<bibRefCitation id="FF66B8FD0F560EE9B3AE901D9596A481" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
)
</taxonomicName>
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<paragraph id="08A13DCA35B6126A5FDB54BF9E5B9FA4">
<figureCitation id="27AF263594B608BC59492AAE5BFFF408" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A right antenna, dorsal view B antennal article XIV, ventral view C clusters of type c sensilla on antennal article XIII (top) and IX (bottom), dorsal view." figureDoi="10.3897/zookeys.1228.143007.figure19" httpUri="https://binary.pensoft.net/fig/1257839">Figs 19</figureCitation>
,
<figureCitation id="80CA6E90F361362829C9010604D9994D" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Ityphilus cf. taeniaformis (Lawrence, 1960) NHMUK 015991468 A head and antennae, dorsal view B cephalic shield, dorsal view C right antenna, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure20" httpUri="https://binary.pensoft.net/fig/1257840">20</figureCitation>
,
<figureCitation id="24D9E6DBC59578C85028602F434171F9" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A mandible, lateral view B first and second maxillae, ventral view C anterior margin of second maxillary coxosternite, ventral view D second maxillary pretarsus left (top), right (bottom), ventral view." figureDoi="10.3897/zookeys.1228.143007.figure21" httpUri="https://binary.pensoft.net/fig/1257841">21</figureCitation>
,
<figureCitation id="244E55A8ADDA4C4A65C3311960A41C31" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A forcipular segment, ventral view B left forcipule, ventral view C calyx of venom gland, ventral view D leg-bearing segment 9, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure22" httpUri="https://binary.pensoft.net/fig/1257842">22</figureCitation>
,
<figureCitation id="1234F55B85485EE0E8C6629A780B0F0C" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A ultimate leg-bearing and postpedal segments, ventral view B tip of right ultimate leg telopodite, ventral view C male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure23" httpUri="https://binary.pensoft.net/fig/1257843">23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMECI" type="material">
<paragraph id="5F142442B48ACAB08FD7CD40E9ED0F07">
<heading id="DC217A22B423FC78C5B4FFD4A421540D" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="AE13F95699A980564C262E0DBAF1A10C">
<materialsCitation id="6CB86A9EF4BD66F7DF142BBA4F7A252E" collectingDate="1974-03-21" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Aldabra" specimenCode="NHMUK 015991468" specimenCount="1" specimenCount-male="1">
<specimenCode id="7131E8A75E3DFBC38EFA253CFDDBBF60">NHMUK 015991468</specimenCode>
,
<specimenCount id="A7D798272053D53A86BD0F431002CBF3" count="1" type="male">1 ♂</specimenCount>
,
<location id="15C5551CC77F0B825673D380E8FBD6EA" LSID="urn:lsid:plazi:treatment:6BD3F1C71B37502596C18A3015C2A68D:15C5551CC77F0B825673D380E8FBD6EA" name="Aldabra">Aldabra</location>
,
<date id="9C6DFD08686E20059A2A70E56E04D0C1" value="1974-03-21">
<collectingDate id="AB92A535526277919E22E7E7C4472DA1" value="1974-03-21">21. 03. 1974</collectingDate>
</date>
.,
<collectorName id="4DA869A7E9223F21AEBAADA735E78293">V. W. Spaull</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERECI" type="description">
<paragraph id="C396F17A0AFDC479EFACD659F6E0A7F4">
<heading id="2B3DB64C228344226A041291B8EB7FB3" reason="title">Description.</heading>
</paragraph>
<paragraph id="2129B306C69DB8087AEBDC2CEC8C6C8D">
<emphasis id="25B40EF9ED382DD731FB310EB1873778" bold="true">
<emphasis id="C3D9BF03CE9BAE7582EA3E22F81A6BDB" bold="true" italics="true">Head and antennae</emphasis>
.
</emphasis>
Antennae conspicuously claviform in shape, medially weakly geniculate, with articles IX XIV widened (Figs
<figureCitation id="1806BE3D2E0945A585DC1E29AEE143FA" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A right antenna, dorsal view B antennal article XIV, ventral view C clusters of type c sensilla on antennal article XIII (top) and IX (bottom), dorsal view." figureDoi="10.3897/zookeys.1228.143007.figure19" httpUri="https://binary.pensoft.net/fig/1257839">19 A</figureCitation>
,
<figureCitation id="D672387AA34E40F98AAA985AF33CCDCA" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Ityphilus cf. taeniaformis (Lawrence, 1960) NHMUK 015991468 A head and antennae, dorsal view B cephalic shield, dorsal view C right antenna, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure20" httpUri="https://binary.pensoft.net/fig/1257840">20 A, C</figureCitation>
). Articles IX and XIII with clusters of type c sensilla (
<emphasis id="ED0167DEBEAF0D87A5316D1B94E8F39F" italics="true">sensu</emphasis>
<bibRefCitation id="7B1EAC94D9D40FDF7E2056EED4599A3F" DOI="10.1080/01650521.2017.1412687" author="Pereira" firstAuthor="Pereira" issue="2" journalOrPublisher="Studies on Neotropical Fauna and Environment" pagination="91-106" refId="B75" refString="Pereira LA (2017) A new miniature species of geophilomorph centipede from the Ecuadorian Amazon rainforest (Chilopoda: Geophilomorpha: Ballophilidae). Studies on Neotropical Fauna and Environment 53 (2): 91106. https://doi.org/10.1080/01650521.2017.1412687" title="A new miniature species of geophilomorph centipede from the Ecuadorian Amazon rainforest (Chilopoda: Geophilomorpha: Ballophilidae)." volume="53" year="2017">Pereira 2017</bibRefCitation>
) on the distal edge of the dorsal side (Fig.
<figureCitation id="EE3D4B66FD746D0304CAFAA520A6FCDD" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A right antenna, dorsal view B antennal article XIV, ventral view C clusters of type c sensilla on antennal article XIII (top) and IX (bottom), dorsal view." figureDoi="10.3897/zookeys.1228.143007.figure19" httpUri="https://binary.pensoft.net/fig/1257839">19 C, D</figureCitation>
). Article XIV with two lateral clusters of sensilla basiconica and a small number of spear-shaped sensilla at its apex (Fig.
<figureCitation id="02385CF9BF5F25BC32D46B3C0A7AA7FD" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A right antenna, dorsal view B antennal article XIV, ventral view C clusters of type c sensilla on antennal article XIII (top) and IX (bottom), dorsal view." figureDoi="10.3897/zookeys.1228.143007.figure19" httpUri="https://binary.pensoft.net/fig/1257839">19 B</figureCitation>
). Head approximately as wide as forcipular tergite, 1.14 × broader than long. Curved sulcus near posterior margin. Chaetotaxy of head as in Fig.
<figureCitation id="B8B722D5BF33AB4C7E47DE5616262A15" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Ityphilus cf. taeniaformis (Lawrence, 1960) NHMUK 015991468 A head and antennae, dorsal view B cephalic shield, dorsal view C right antenna, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure20" httpUri="https://binary.pensoft.net/fig/1257840">20 B</figureCitation>
.
</paragraph>
<caption id="D74E5B04A5A96AD8BAA25BBF534F090F" ID-DOI="10.3897/zookeys.1228.143007.figure19" ID-arpha="F3DFF2B9-4B12-5D09-9343-BBBA090AEEE3" httpUri="https://binary.pensoft.net/fig/1257839" startId="F19">
<paragraph id="B792FA3D1E9ED20E063F508EDB2B36CC">
<label id="56FBC671FFE03C67CCA7399903819DE6">Figure 19.</label>
</paragraph>
<paragraph id="AC2B7BCDC3A7B3FE6424DA9408F12B6C">
<taxonomicName id="11DE8F28AE9D4B6426633100630C3541" authority="(Lawrence, 1960)" authorityName="NHMUK" baseAuthorityName="Lawrence" baseAuthorityYear="1960" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="31E31885C9D1F234B46818761CAD4523" italics="true">Ityphilus</emphasis>
cf.
<emphasis id="DE9BC3EA80B0364BD0ADC24E15A064A5" italics="true">taeniaformis</emphasis>
(
<bibRefCitation id="7E16A88D63585B613F167D13C16D7C25" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
)
</taxonomicName>
.
<specimenCode id="CF96244B4D22AD4CFD51F11944F33E79">NHMUK 015991468</specimenCode>
<emphasis id="6F7A83C364B0BDFB2D47C03147412F98" bold="true">A</emphasis>
right antenna, dorsal view
<emphasis id="E37A455CD7AE7887833CD953D98D8471" bold="true">B</emphasis>
antennal article XIV, ventral view
<emphasis id="E5CDAFCD1BA2E2904910002239446D0C" bold="true">C</emphasis>
clusters of type c sensilla on antennal article XIII (top) and IX (bottom), dorsal view.
</paragraph>
</caption>
<caption id="B4EC34AE8934E75C70AB6AEBCC0C43D8" ID-DOI="10.3897/zookeys.1228.143007.figure20" ID-arpha="61267297-F99B-541A-9181-126770724899" httpUri="https://binary.pensoft.net/fig/1257840" startId="F20">
<paragraph id="9CB39E5C75910CDDF3792860B8AAA9C4">
<label id="0C5FD877D1C80EC361AB7AB989A6AACF">Figure 20.</label>
</paragraph>
<paragraph id="9414D90D5EA3AEEFBC4232F202143EB6">
<taxonomicName id="B051EB574A9B2EAAEB52AA7F56A20586" authority="(Lawrence, 1960) NHMUK" authorityName="NHMUK" baseAuthorityName="Lawrence" baseAuthorityYear="1960" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="6B6CDAF1734D8A31AA7044A8C11A5C9D" italics="true">Ityphilus</emphasis>
cf.
<emphasis id="B3A3104C05BB3054C116066B537C9B19" italics="true">taeniaformis</emphasis>
(
<bibRefCitation id="908F786EC728F3CE62E9C9F6F988B554" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
)
<specimenCode id="F3F5F51709E1D51E759229F0E05DDC48">NHMUK 015991468</specimenCode>
</taxonomicName>
<emphasis id="AB03168C92FB57D3951D1E7DF300F77E" bold="true">A</emphasis>
head and antennae, dorsal view
<emphasis id="358941E49E7F13530EC1CBBFA7D739AB" bold="true">B</emphasis>
cephalic shield, dorsal view
<emphasis id="8089D829FCF388BF7FB17E34EE64CA6E" bold="true">C</emphasis>
right antenna, ventral view.
</paragraph>
</caption>
<paragraph id="E07F3E7D31DD7AAFC35B713CF74C6B4C">
<emphasis id="7E6761979292BBB607634E2B5D5C569A" bold="true">
<emphasis id="A807707901A6910F0C634F7BC180DDC9" bold="true" italics="true">Mandibles</emphasis>
.
</emphasis>
Dentate lamella with seven denticles, only six conspicuous in lateral view. Pectinate lamella with approximately 22 hyaline projections (Fig.
<figureCitation id="CEA8F10F249707F9A28B633254A840BA" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A mandible, lateral view B first and second maxillae, ventral view C anterior margin of second maxillary coxosternite, ventral view D second maxillary pretarsus left (top), right (bottom), ventral view." figureDoi="10.3897/zookeys.1228.143007.figure21" httpUri="https://binary.pensoft.net/fig/1257841">21 A</figureCitation>
).
</paragraph>
<caption id="6249FB78F25003FDE6B1DDE703B7EA4B" ID-DOI="10.3897/zookeys.1228.143007.figure21" ID-arpha="44D7461E-CCA9-599C-8B68-F46EBB36FE7C" httpUri="https://binary.pensoft.net/fig/1257841" startId="F21">
<paragraph id="53001402BBD313925A553E472422FAFC">
<label id="1A219C5341856037DEAE4681306D3A40">Figure 21.</label>
</paragraph>
<paragraph id="DF31BACCE01F0F6F24683C57D52ABC4D">
<taxonomicName id="B0927DAFCDD1B5CA422F39047A4020ED" authority="(Lawrence, 1960)" authorityName="NHMUK" baseAuthorityName="Lawrence" baseAuthorityYear="1960" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="7224382A37DB8CA45325E650B2B7E044" italics="true">Ityphilus</emphasis>
cf.
<emphasis id="568CC54D324196F915EE3D432A910BB3" italics="true">taeniaformis</emphasis>
(
<bibRefCitation id="6EDF88B119399A761173724EC8DB1AFD" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
)
</taxonomicName>
.
<specimenCode id="119FFA044D3AFA9522184666A1137A71">NHMUK 015991468</specimenCode>
<emphasis id="F5FDFE1E6A51BF0A9A378E83133965E3" bold="true">A</emphasis>
mandible, lateral view
<emphasis id="270254AE25F66B2F3601631A52DB8859" bold="true">B</emphasis>
first and second maxillae, ventral view
<emphasis id="974E9D2ABC6E5CE408FD78FA95023C12" bold="true">C</emphasis>
anterior margin of second maxillary coxosternite, ventral view
<emphasis id="74613636121150B36F69C5EF543E44CC" bold="true">D</emphasis>
second maxillary pretarsus left (top), right (bottom), ventral view.
</paragraph>
</caption>
<paragraph id="BB7809DDDE76F1686F00FEDEAE8BB884">
<emphasis id="7EC9C2BD143D809154BE63C6E78CF821" bold="true">
<emphasis id="7824A57B806DE5A52CA76DE2FF76287E" bold="true" italics="true">Labrum and clypeus</emphasis>
.
</emphasis>
Clypeus with a pair of postantennal setae, a cluster of seven medial setae and one prelabral seta. Lateral pieces of labrum narrow, conspicuously sclerotised, lacking any fringes or projections. Medial piece contiguous with clypeus, poorly sclerotised, membranous and lacking conspicuous hairs or projections.
</paragraph>
<paragraph id="1C91E7C8B4EE6EF5F4B541569E7E7E2D">
<emphasis id="6194C15D466F534C0C60EE4AB78592E8" bold="true">
<emphasis id="BB2BAEC1A3B9C5409D3C46DFBF611C32" bold="true" italics="true">Maxillae</emphasis>
.
</emphasis>
First maxillae with evident, triangular coxal projections, each bearing one sensillum. Telopodites bearing one sensillum each, conspicuously larger than coxal projections and partly covering them (Fig.
<figureCitation id="434CC44740B429962EA920E07B275CCD" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A mandible, lateral view B first and second maxillae, ventral view C anterior margin of second maxillary coxosternite, ventral view D second maxillary pretarsus left (top), right (bottom), ventral view." figureDoi="10.3897/zookeys.1228.143007.figure21" httpUri="https://binary.pensoft.net/fig/1257841">21 B</figureCitation>
). Lappets absent. Second maxillary coxosternite with evident but incomplete medial suture, extending for half of its length (Fig.
<figureCitation id="8A421BAB789890D6409CD731CF6FBBBC" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A mandible, lateral view B first and second maxillae, ventral view C anterior margin of second maxillary coxosternite, ventral view D second maxillary pretarsus left (top), right (bottom), ventral view." figureDoi="10.3897/zookeys.1228.143007.figure21" httpUri="https://binary.pensoft.net/fig/1257841">21 C</figureCitation>
). Each side of the suture bearing one sensillum. Telopodite stout, terminating in large pretarsus. Second maxillary pretarsus spatulate, lateral edges densely fringed (Fig.
<figureCitation id="8E17AB736A05EE2B17695A598D4135C4" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A mandible, lateral view B first and second maxillae, ventral view C anterior margin of second maxillary coxosternite, ventral view D second maxillary pretarsus left (top), right (bottom), ventral view." figureDoi="10.3897/zookeys.1228.143007.figure21" httpUri="https://binary.pensoft.net/fig/1257841">21 D, E</figureCitation>
).
</paragraph>
<paragraph id="1AE4A02A97C3D619406115747581B9AE">
<emphasis id="ECEE2958C4640717571D28CDCE653C92" bold="true">
<emphasis id="817571B5971E707847DE2588F268207A" bold="true" italics="true">Forcipular segment</emphasis>
.
</emphasis>
Exposed face of forcipular coxosternite 2.2 × broader than long (Fig.
<figureCitation id="C7B596C9F5F7F1831B85470BB243BBB2" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A forcipular segment, ventral view B left forcipule, ventral view C calyx of venom gland, ventral view D leg-bearing segment 9, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure22" httpUri="https://binary.pensoft.net/fig/1257842">22 A</figureCitation>
). Chitin lines present, reaching the condyles. Forcipular trochanteroprefemur 1.25 × longer than broad. Calyx of venom gland elongated, ovoid in shape (Fig.
<figureCitation id="F4A9A6B75507CAA182054701228D8923" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A forcipular segment, ventral view B left forcipule, ventral view C calyx of venom gland, ventral view D leg-bearing segment 9, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure22" httpUri="https://binary.pensoft.net/fig/1257842">22 C</figureCitation>
). All forcipular articles without denticles (Fig.
<figureCitation id="0D71F1EC29909C2C26769961E4EFFDF5" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A forcipular segment, ventral view B left forcipule, ventral view C calyx of venom gland, ventral view D leg-bearing segment 9, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure22" httpUri="https://binary.pensoft.net/fig/1257842">22 B</figureCitation>
). Internal margin of tarsungulum smooth. Extended, forcipules do not reach the anterior margin of the head.
</paragraph>
<caption id="7766E282FB83DCB84753FC05E37B01D0" ID-DOI="10.3897/zookeys.1228.143007.figure22" ID-arpha="A9E0FE07-C726-5ACB-B2A7-AD6E40DCF7A2" httpUri="https://binary.pensoft.net/fig/1257842" startId="F22">
<paragraph id="9CBD43839507E5D90B69968E742032EA">
<label id="77741620587B8D679209ABECE8804840">Figure 22.</label>
</paragraph>
<paragraph id="A60B763070BF18EC6D859ABB0680633E">
<taxonomicName id="396A0B2E898415EE2513786C0B16E0C8" authority="(Lawrence, 1960)" authorityName="NHMUK" baseAuthorityName="Lawrence" baseAuthorityYear="1960" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="F48FE184AB34DD06BB43B74210B26743" italics="true">Ityphilus</emphasis>
cf.
<emphasis id="1AA5D214B221F4D0CBC057C3FD7E9FF0" italics="true">taeniaformis</emphasis>
(
<bibRefCitation id="2B393AEAE00E1BB4EA27E3241ECF9E35" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
)
</taxonomicName>
.
<specimenCode id="ED8A503881387993435E311B3AEBDE5B">NHMUK 015991468</specimenCode>
<emphasis id="0EFAF4DD4B578CCC74E98AB5CA6FF452" bold="true">A</emphasis>
forcipular segment, ventral view
<emphasis id="56CADD186C3FAFC6C44D30E7C4DA1367" bold="true">B</emphasis>
left forcipule, ventral view
<emphasis id="B3417DF99616215645C38353B8420286" bold="true">C</emphasis>
calyx of venom gland, ventral view
<emphasis id="1FBD21142495956066EC587687083053" bold="true">D</emphasis>
leg-bearing segment 9, ventral view.
</paragraph>
</caption>
<paragraph id="4A60D0214815288BD551D5FB81C19E77">
<emphasis id="FF6FF1FE39EB09F65CE36E1450070AAC" bold="true">
<emphasis id="0CC54C737E04DD175543FE1E391412AE" bold="true" italics="true">Trunk</emphasis>
.
</emphasis>
75 leg-bearing segments. Pore fields located on raised areas in the middle of all metasternites excluding those of leg-bearing segments 1, 74, and 75. Shape of pore field oval, medially constricted and anteriorly bordered by a line of setae (Fig.
<figureCitation id="F7ED83998157031CEA942F5C0CB7ED62" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A forcipular segment, ventral view B left forcipule, ventral view C calyx of venom gland, ventral view D leg-bearing segment 9, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure22" httpUri="https://binary.pensoft.net/fig/1257842">22 D</figureCitation>
). Colour of pore field bluish grey, conspicuously pigmented relative to surrounding cuticle. Despite the faded colour of the ethanol-preserved specimen, pigmentation of the pore field is conspicuous and the trunk is generally greenish grey in appearance.
</paragraph>
<paragraph id="23DCA5878F95A4885B45654DB7D363B6">
<emphasis id="C79B3C3E218FDA8AEA1EAAC849D59A80" bold="true">
<emphasis id="BF72BF054336FF4AFB1F6A12D324E652" bold="true" italics="true">Ultimate leg</emphasis>
-
<emphasis id="339F713F4253F67703E7B5A8169D15AE" bold="true" italics="true">bearing and postpedal segments</emphasis>
.
</emphasis>
Intercalary pleurites separated from ultimate pretergite by evident sutures. Ultimate metasternite trapezoidal, 1.3 × longer than broad. Coxopleura each with two distinct coxal organs, partially covered by the ultimate metasternite (Fig.
<figureCitation id="139627CB44E7B310B3DC6A096705A2F3" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A ultimate leg-bearing and postpedal segments, ventral view B tip of right ultimate leg telopodite, ventral view C male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure23" httpUri="https://binary.pensoft.net/fig/1257843">23 A</figureCitation>
). Entire ventral side of ultimate leg-bearing segment covered in setae. Ultimate leg telopodite composed of seven articles, all distinctly thickened. Pretarsus absent (Fig.
<figureCitation id="166DAC99BFDC640700263B67E692679F" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A ultimate leg-bearing and postpedal segments, ventral view B tip of right ultimate leg telopodite, ventral view C male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure23" httpUri="https://binary.pensoft.net/fig/1257843">23 B</figureCitation>
). Metatarsus with a small spine subapically. Intermediate sternite indistinct. First genital sternite with straight posterior margin. Gonopods uni-articulate, flanking penis (Fig.
<figureCitation id="48F177FAED85B3F8A3B75B2741717FD8" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A ultimate leg-bearing and postpedal segments, ventral view B tip of right ultimate leg telopodite, ventral view C male gonopods, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure23" httpUri="https://binary.pensoft.net/fig/1257843">23 C</figureCitation>
).
</paragraph>
<caption id="C97FE34434EC0C4713B52312A1B67853" ID-DOI="10.3897/zookeys.1228.143007.figure23" ID-arpha="A9496993-3A2A-5557-9E25-23C070CD7A75" httpUri="https://binary.pensoft.net/fig/1257843" startId="F23">
<paragraph id="557C0DA8F6147AF6D90F94C32FD802CE">
<label id="745D689875F2E06C02BFE1CA60E1EE1C">Figure 23.</label>
</paragraph>
<paragraph id="F41C3A11DBE19997AA2E4BB02EC7F884">
<taxonomicName id="6766CA9F5D8B03A5AEA17D8C88187C34" authority="(Lawrence, 1960)" authorityName="NHMUK" baseAuthorityName="Lawrence" baseAuthorityYear="1960" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="B78E1B22F8F2B18B73C89654AA617A19" italics="true">Ityphilus</emphasis>
cf.
<emphasis id="95C06367D12A2E75D7BFD0B59F74A22D" italics="true">taeniaformis</emphasis>
(
<bibRefCitation id="2D4DDDE18627C886370FEF89A677429E" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
)
</taxonomicName>
.
<specimenCode id="7EA862DFA7582C9448CCC5F1527ABB2C">NHMUK 015991468</specimenCode>
<emphasis id="9FECAC2F80E9886038916633BB277208" bold="true">A</emphasis>
ultimate leg-bearing and postpedal segments, ventral view
<emphasis id="92FD19CDA4A76ACFA50974978DD5DADD" bold="true">B</emphasis>
tip of right ultimate leg telopodite, ventral view
<emphasis id="00143B3130034992AF1AFF4F4FE2165F" bold="true">C</emphasis>
male gonopods, ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ESNCI" type="remarks">
<paragraph id="441CD0D34C89146A59A0EAF8093D6F82">
<heading id="6687183652E9A073A6A8E1CEDB9A56D2" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="DCC2DD37B3BDE2BFC62C68652B93CAAA">
The taxonomy of
<taxonomicName id="48EE1F72255B95FB09B090837CABF61D" authorityName="Cook" authorityYear="1899" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="7BC796A15E208FBBC3A3D419CFE29B02" italics="true">Ityphilus</emphasis>
</taxonomicName>
remains largely unresolved, especially outside of South America, where different authors have disagreed on its relation to
<taxonomicName id="08676A55A9FEBDF326820524EB536382" baseAuthorityName="Verhoeff" baseAuthorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="872DD8936FACDF409F59F85E7CA37BF0" italics="true">Ballophilus</emphasis>
</taxonomicName>
, alternatively considering it a different genus (
<bibRefCitation id="46C3CF95A50CB10014E4E5CEAAC46AA6" author="Attems" firstAuthor="Attems" refId="B4" refString="Attems C (1929) Geophilomorpha. Das Tierreich, 52. W. de Gruyter &amp; Co., Berlin / Leipzig. [XXIII +] 388 pp." year="1929">Attems 1929</bibRefCitation>
) or a synonym of
<taxonomicName id="1081A41FCF581CED0C29B9E64BA56BBC" authority="(Verhoeff 1939)" baseAuthorityName="Verhoeff" baseAuthorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="2742436D725F5FF53BD405D0616BF043" italics="true">Ballophilus</emphasis>
(
<bibRefCitation id="EDC99E59303CED141CC0186268DF740B" author="Verhoeff" firstAuthor="Verhoeff" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere 72: 7198." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff 1939</bibRefCitation>
)
</taxonomicName>
. Recent revision of the genus (
<bibRefCitation id="61EE6B45E9AA5428188C55DEBA840990" DOI="10.11646/zootaxa.1485.1.1" author="Bonato" etAl="et al." firstAuthor="Bonato" journalOrPublisher="Zootaxa" pagination="1-12" refId="B12" refString="Bonato L, Pereira LA, Minelli A (2007) Taxonomic and nomenclatural notes on the centipede genera Chomatobius, Ityphilus, Hapleurytion, Plateurytion, and Steneurytion (Chilopoda: Geophilomorpha). Zootaxa 1485: 112. https://doi.org/10.11646/zootaxa.1485.1.1" title="Taxonomic and nomenclatural notes on the centipede genera Chomatobius, Ityphilus, Hapleurytion, Plateurytion, and Steneurytion (Chilopoda: Geophilomorpha)." volume="1485" year="2007">Bonato et al. 2007</bibRefCitation>
) has maintained the distinction between
<taxonomicName id="F518E0A29C9AB60BFA5C89111E64E25A" authorityName="Cook" authorityYear="1899" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="3D26A259C7D6FC9D1362426A99B917B0" italics="true">Ityphilus</emphasis>
</taxonomicName>
and
<taxonomicName id="07C53FFB8B560AA3E1BB4AE253F20BAC" baseAuthorityName="Verhoeff" baseAuthorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="F7ECEB8E3B7AEFD273E94392AA33BE64" italics="true">Ballophilus</emphasis>
</taxonomicName>
, but remarked on the close relationship between the two, and on cases in which the presence of complete or nearly complete chitin lines is doubtful, such as in
<taxonomicName id="D9926EE0905114175C6230C86DC8ECD4" authority="(Wang, 1955)" baseAuthorityName="Wang" baseAuthorityYear="1955" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="boteltobogensis">
<emphasis id="F5E9DAF869C80E3058A5135BD73C3EDD" italics="true">Ityphilus boteltobogensis</emphasis>
(
<bibRefCitation id="DA79CA3CCDC48214DE09FF1186BFC120" author="Wang YM" journalOrPublisher="Quarterly Journal of the Taiwan Museum" pagination="195-201" refId="B110" refString="Wang YM (1955) Serica 1 b. A preliminary report on the Myriapoda and Arachnida of Lan Yu Islets (Botel Tobago) China. Quarterly Journal of the Taiwan Museum 8: 195-201." title="Serica 1 b. A preliminary report on the Myriapoda and Arachnida of Lan Yu Islets (Botel Tobago) China." volume="8" year="1955">Wang, 1955</bibRefCitation>
)
</taxonomicName>
, despite this character being predominantly used to distinguish the two genera. Similarly, several
<taxonomicName id="C78AAC7660E41AF26DEC4BDA10FB6FB6">
<emphasis id="3939D615DB219C7FA74B7767BE30B3E6" italics="true">Ballophilus</emphasis>
species
</taxonomicName>
are described as bearing a cuticular thickening in the usual position of the chitin line (
<bibRefCitation id="D8E86432A7926AB6F3533BC2E2418516" author="Demange" firstAuthor="Demange" journalOrPublisher="La réserve naturelle integrale du Mont Nimba - Mémoires de lInstitut Française dAfrique Noire" pagination="41-118" refId="B26" refString="Demange JM (1963) Chilopoda. La réserve naturelle integrale du Mont Nimba Mémoires de lInstitut Française dAfrique Noire 5: 41118." title="Chilopoda." volume="5" year="1963">Demange 1963</bibRefCitation>
;
<bibRefCitation id="897D42CA27571CB3148F963F78620B5D" DOI="10.1076/snfe.31.3.170.13335" author="Pereira" etAl="et al." firstAuthor="Pereira" issue="3-4" journalOrPublisher="Studies on Neotropical Fauna and Environment" pagination="170-178" refId="B76" refString="Pereira LA, Foddai D, Minelli A (1997) First Record of a Ballophilid Centipede from Argentina with a Description of Ballophilus ramirezi n. sp. (Chilopoda: Geophilomorpha: Ballophilidae). Studies on Neotropical Fauna and Environment 31 (34): 170178. https://doi.org/10.1076/snfe.31.3.170.13335" title="First Record of a Ballophilid Centipede from Argentina with a Description of Ballophilus ramirezi n. sp. (Chilopoda: Geophilomorpha: Ballophilidae)." volume="31" year="1997">Pereira et al. 1997</bibRefCitation>
), further confounding the utility of this character in taxonomy within the
<taxonomicName id="1D2BC35109411D7D8819240DE682BCA0" class="Chilopoda" family="Ballophilidae" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="family">Ballophilidae</taxonomicName>
. The presence of a median sulcus in the second maxillary coxosternite has been shown to be unreliable in separating
<taxonomicName id="A166D37F1FFCCEFCE3EB873D75F2F560" baseAuthorityName="Verhoeff" baseAuthorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="30278570B9E5248453E6C0D8A3EC66D0" italics="true">Ballophilus</emphasis>
</taxonomicName>
and
<taxonomicName id="855000CFC97E860F178C1E3B235270CD" authorityName="Cook" authorityYear="1899" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="69BD9188486C62C7324C375C9B5CDD50" italics="true">Ityphilus</emphasis>
</taxonomicName>
. This character has been described in both
<taxonomicName id="22BFF1D71210DB53B60254E8B5C99325" baseAuthorityName="Verhoeff" baseAuthorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="2B48A52B3D4C993717F3564E89909277" italics="true">Ballophilus</emphasis>
</taxonomicName>
(
<bibRefCitation id="42FF9ACB896912DE1539C33E1356A333" author="Ribaut" firstAuthor="Ribaut" refId="B90" refString="Ribaut H (1914) Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale (19111912). Résultats Scientifiques. Myriapodes. I. Chilopoda. Libr. A. Schulz, Paris 135." year="1914">Ribaut 1914</bibRefCitation>
;
<bibRefCitation id="C934ED8870A7F48F825090F33877E63B" DOI="10.1076/snfe.31.3.170.13335" author="Pereira" etAl="et al." firstAuthor="Pereira" issue="3-4" journalOrPublisher="Studies on Neotropical Fauna and Environment" pagination="170-178" refId="B76" refString="Pereira LA, Foddai D, Minelli A (1997) First Record of a Ballophilid Centipede from Argentina with a Description of Ballophilus ramirezi n. sp. (Chilopoda: Geophilomorpha: Ballophilidae). Studies on Neotropical Fauna and Environment 31 (34): 170178. https://doi.org/10.1076/snfe.31.3.170.13335" title="First Record of a Ballophilid Centipede from Argentina with a Description of Ballophilus ramirezi n. sp. (Chilopoda: Geophilomorpha: Ballophilidae)." volume="31" year="1997">Pereira et al. 1997</bibRefCitation>
) and
<taxonomicName id="D0FC0FC2E417D9280FB34FF16A9DB661" authority="(Pereira et al. 2000)" baseAuthorityName="Pereira" baseAuthorityYear="2000" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="FF9D3A9FE08259E56834946C99A061F9" italics="true">Ityphilus</emphasis>
(
<bibRefCitation id="7EC1F735CDEF744E01788771B648AA7C" author="Pereira" etAl="et al." firstAuthor="Pereira" journalOrPublisher="Amazoniana" pagination="1-57" refId="B77" refString="Pereira LA, Foddai D, Minelli A (2000) New taxa of Neotropical Geophilomorpha (Chilopoda). Amazoniana 16: 157." title="New taxa of Neotropical Geophilomorpha (Chilopoda)." volume="16" year="2000">Pereira et al. 2000</bibRefCitation>
)
</taxonomicName>
, in some cases as incomplete (
<bibRefCitation id="EC7AFE100188FB9861A61A51248308B3" author="Brolemann" firstAuthor="Brolemann" issue="3" journalOrPublisher="Archives de Zoologie Expérimentale et Générale" pagination="303-340" refId="B15" refString="Brölemann H (1909) À propos dun système des géophilomorphes. Archives de Zoologie Expérimentale et Générale 5 (3): 303340." title="À propos dun système des géophilomorphes." volume="5" year="1909">Brölemann 1909</bibRefCitation>
) (Fig.
<figureCitation id="A380FA11C324B1B870E7AD1BBC0D3610" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Ityphilus cf. taeniaformis (Lawrence, 1960). NHMUK 015991468 A mandible, lateral view B first and second maxillae, ventral view C anterior margin of second maxillary coxosternite, ventral view D second maxillary pretarsus left (top), right (bottom), ventral view." figureDoi="10.3897/zookeys.1228.143007.figure21" httpUri="https://binary.pensoft.net/fig/1257841">21 B, C</figureCitation>
), and is not included in the most recent diagnosis of the latter genus (
<bibRefCitation id="2D63F768CDE8952AAB2F1B74F0F400E2" DOI="10.11646/zootaxa.1485.1.1" author="Bonato" etAl="et al." firstAuthor="Bonato" journalOrPublisher="Zootaxa" pagination="1-12" refId="B12" refString="Bonato L, Pereira LA, Minelli A (2007) Taxonomic and nomenclatural notes on the centipede genera Chomatobius, Ityphilus, Hapleurytion, Plateurytion, and Steneurytion (Chilopoda: Geophilomorpha). Zootaxa 1485: 112. https://doi.org/10.11646/zootaxa.1485.1.1" title="Taxonomic and nomenclatural notes on the centipede genera Chomatobius, Ityphilus, Hapleurytion, Plateurytion, and Steneurytion (Chilopoda: Geophilomorpha)." volume="1485" year="2007">Bonato et al. 2007</bibRefCitation>
).
</paragraph>
<paragraph id="CE114C0A4BA9A9602F54592468AD19EF">
<bibRefCitation id="2DA9ADC3691FC28F67D145AD8F2A8450" author="Verhoeff KW" firstAuthor="Verhoeff" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere 72: 7198." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff (1939)</bibRefCitation>
described two
<taxonomicName id="F0519219A78689C79654FE5FE6D1AF9F">
<emphasis id="BA50EE2C7BDB1CFC03F6150E70064556" italics="true">Ballophilus</emphasis>
species
</taxonomicName>
from
<collectingCountry id="43B6B8E0F50107EBBE09B886DC29C654" name="Mauritius">Mauritius</collectingCountry>
,
<taxonomicName id="5652B9FD53740E19FE11C687F58AF027" authority="Verhoeff, 1939" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="5FD9B50B15B693F76AE6D1DF5C8BF7F4" italics="true">B. lawrencei</emphasis>
<bibRefCitation id="B1A6322918E2E70CC3A80EB3F4D47092" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 72: 71-98." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff, 1939</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="EF5B6952243086992250253B4413FC0A" authority="Verhoeff, 1939" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="mauritianus">
<emphasis id="0BCA57C0A4835862A22851C49C15FFC5" italics="true">B. mauritianus</emphasis>
<bibRefCitation id="E839C40568F0BA25CC8177BD4AF56AB5" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 72: 71-98." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff, 1939</bibRefCitation>
</taxonomicName>
. Both are known from single specimens but, according to their original description, compare closely with
<taxonomicName id="86A198C807E2FF347B9C195F3A6632A0" authority="Ribaut, 1914" authorityName="Ribaut" authorityYear="1914" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="allauadi">
<emphasis id="8F08E463FC50005095AE26C764477E6C" italics="true">Ballophilus allauadi</emphasis>
<bibRefCitation id="739B4EAC17F9FAEA708659F5F9D49A70" author="Ribaut H" refId="B90" refString="Ribaut H (1914) Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale (1911 - 1912). Resultats Scientifiques. Myriapodes. I. Chilopoda. Libr. A. Schulz, Paris 1 - 35." year="1914">Ribaut, 1914</bibRefCitation>
</taxonomicName>
described from Eastern Africa. The Aldabra specimen differs from these species in the presence of pore fields on all but the first and last two leg-bearing segment metasternites (contrasting with the absence of the pore field on the first and the last four metasternites). Additionally, the distal end of the antenna of
<taxonomicName id="61484347ECC8CD1BC6F14ACFF2A5D09D" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="311AC4B82ECE2E662AA69D70573C06E0" italics="true">B. lawrencei</emphasis>
</taxonomicName>
is illustrated as markedly less clavate than observed for the Aldabra specimen. Despite this, the
<taxonomicName id="CC6C4FC4ECD1A8CC3A63867549ED41B0" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="3D4C1A9762C24FE439F9F13C6B4D8C21" italics="true">Ityphilus</emphasis>
</taxonomicName>
specimen collected in Aldabra overlaps in the shape of the metasternal pore field, its position on a raised area and in the relative elongation of the ultimate leg-bearing segment telopodite and the number of leg-bearing segments with
<taxonomicName id="C8A9D80AE2962B48AC614D6BE6C0911E" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lawrencei">
<emphasis id="4C9260AD45727BA4D923D30EF1E084F5" italics="true">B. lawrencei</emphasis>
</taxonomicName>
. As all Mauritian
<taxonomicName id="E3E24B999ED75EA12B053904B121F2C5">
<emphasis id="48C16E22CC71214BFFB93633C967B853" italics="true">Ballophilus</emphasis>
species
</taxonomicName>
are known solely from their
<typeStatus id="0CAC2BAB6123CD1A7F61D14707E54E3E" type="holotype">holotypes</typeStatus>
, it is not possible to comment on intraspecific variability that may account for the overlap in these traits.
</paragraph>
<paragraph id="1EBB86AE06F6A6A7B9C1AF83E55121C1">
<taxonomicName id="AE68BC907F699B93452AE10F7C98129F" authority="Pocock, 1906" authorityName="Pocock" authorityYear="1906" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="maldivensis">
<emphasis id="8E26D7B2AF15D696358700EB2F08BCDC" italics="true">Ballophilus maldivensis</emphasis>
<bibRefCitation id="DFE1A00B31472B3336FC0847E75462F0" author="Pocock RI" refId="B85" refString="Pocock RI (1906) Chilopoda and Diplopoda. In: Gardiner JS (E.) The Fauna and Geography of the Maldive and Laccadive Archipelagoes 2, Supplement 2. Cambridge University Press, 1042 - 1045." year="1906">Pocock, 1906</bibRefCitation>
</taxonomicName>
, described from the
<collectingCountry id="DE9D84B26CE70D5F086CE0A61B55E909" name="Maldives">Maldives</collectingCountry>
, similarly resembles the Aldabra specimen in the shape of the metasternal pore fields and their pigmentation. The incomplete original description did not allow for comparison of any other putative diagnostic characters beside the number of leg-bearing segments (
<quantity id="4575D344D33156BB116B985E54EC596B" metricMagnitude="0" metricUnit="m" metricValue="1.7018" unit="in" value="67.0">67 in</quantity>
the female
<typeStatus id="5B05546D39B91313D8DB90CE3EA5C373" type="holotype">holotype</typeStatus>
), which is lower than that of the Aldabra specimen (
<quantity id="B23C71A0CDE48D3EA5FDE5C624B5C2D0" metricMagnitude="0" metricUnit="m" metricValue="1.905" unit="in" value="75.0">75 in</quantity>
a male). Re-examination of the
<typeStatus id="C014E279D1837EFB0DBF71A9998908D3" type="holotype">holotype</typeStatus>
(Fig.
<figureCitation id="CE5095A1400C337203FE27429170E483" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Ballophilus maldivensis Pocock, 1906. BMNH # 200555 A head and forcipular segment, ventral view B magnified view of forcipular coxosternite showing lateral pleurite and condyle, right side, ventral view C metasternite of leg-bearing segment 7, ventral view D ultimate leg-bearing and postpedal segments, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure24" httpUri="https://binary.pensoft.net/fig/1257844">24</figureCitation>
), the sole known specimen, revealed several features that further distance it from the Aldabra specimen and bring it closer to the currently accepted diagnosis of
<taxonomicName id="942B7F67A6A31F1B934B5D4B34640CBD" baseAuthorityName="Verhoeff" baseAuthorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="39B2F56F7BAF950B8DE264DAF0B7E2B4" italics="true">Ballophilus</emphasis>
</taxonomicName>
.
<taxonomicName id="BCD6E90227D792772D5803517FAAD67F" authorityName="Pocock" authorityYear="1906" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="maldivensis">
<emphasis id="B5FD42D135FFC267C1C91CC38D2A9C54" italics="true">Ballophilus maldivensis</emphasis>
</taxonomicName>
lacks chitin lines or any cuticular thickenings near their position (Fig.
<figureCitation id="155922A1C1A4C2A0738F03823349FF29" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Ballophilus maldivensis Pocock, 1906. BMNH # 200555 A head and forcipular segment, ventral view B magnified view of forcipular coxosternite showing lateral pleurite and condyle, right side, ventral view C metasternite of leg-bearing segment 7, ventral view D ultimate leg-bearing and postpedal segments, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure24" httpUri="https://binary.pensoft.net/fig/1257844">24 A, B</figureCitation>
). Other important differences from
<taxonomicName id="DEE0E218287EEDAC49E167D478935BC9" authorityName="NHMUK" baseAuthorityName="Lawrence" baseAuthorityYear="1960" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="4791A97E25EF2E008C29ED7255B248EC" italics="true">I.</emphasis>
cf.
<emphasis id="CC13D5E0DB7BC983ABDFBD62A829467F" italics="true">taeniaformis</emphasis>
</taxonomicName>
from Aldabra include the absence of the metasternal pore field from the last four leg-bearing segments and a more strongly enlarged ultimate leg telopodite (Fig.
<figureCitation id="1947191C88C3A2EA29CF51551726574D" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Ballophilus maldivensis Pocock, 1906. BMNH # 200555 A head and forcipular segment, ventral view B magnified view of forcipular coxosternite showing lateral pleurite and condyle, right side, ventral view C metasternite of leg-bearing segment 7, ventral view D ultimate leg-bearing and postpedal segments, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure24" httpUri="https://binary.pensoft.net/fig/1257844">24 D</figureCitation>
).
</paragraph>
<caption id="F1F673DA17B304E020A0A6FA33D8A104" ID-DOI="10.3897/zookeys.1228.143007.figure24" ID-arpha="902CDC77-479C-5AF7-8A05-F5B52BE3EF58" httpUri="https://binary.pensoft.net/fig/1257844" startId="F24">
<paragraph id="832732932B374FCC0CF7ACA2A7576B0C">
<label id="F60EEE52E3D84786EF257FB7ECAA3037">Figure 24.</label>
</paragraph>
<paragraph id="A67E10773A4C9910618FF90AFB11F32E">
<taxonomicName id="BF94778FC4230B90EB3008BF31ECDBA1" authority="Pocock, 1906" authorityName="Pocock" authorityYear="1906" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="maldivensis">
<emphasis id="929AE3EE69EA1A8A0739DA26FFC35AEE" italics="true">Ballophilus maldivensis</emphasis>
<bibRefCitation id="A8AB29D6CB05E176FD7121CB528DDC1E" author="Pocock RI" refId="B85" refString="Pocock RI (1906) Chilopoda and Diplopoda. In: Gardiner JS (E.) The Fauna and Geography of the Maldive and Laccadive Archipelagoes 2, Supplement 2. Cambridge University Press, 1042 - 1045." year="1906">Pocock, 1906</bibRefCitation>
</taxonomicName>
. BMNH # 200555
<emphasis id="C0522FCA51FCCCFF6882CEDD121107E2" bold="true">A</emphasis>
head and forcipular segment, ventral view
<emphasis id="A8BD34177CD76E22F06681A6AAD5448E" bold="true">B</emphasis>
magnified view of forcipular coxosternite showing lateral pleurite and condyle, right side, ventral view
<emphasis id="FA5B33435387F16C80B07465388D6E2F" bold="true">C</emphasis>
metasternite of leg-bearing segment 7, ventral view
<emphasis id="F7E078BF1B2875EB353843427AC32821" bold="true">D</emphasis>
ultimate leg-bearing and postpedal segments, ventral view.
</paragraph>
</caption>
<paragraph id="313E092F0FF7BC73C0EC87AD75E3D52E">
<bibRefCitation id="7B2A6B19EE055D0F89401D6ED81451B7" author="Lawrence RF" firstAuthor="Lawrence" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes Chilopodes. Faune de Madagascar 12: 1123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence (1960)</bibRefCitation>
described three species assigned to
<taxonomicName id="2E8EBE16FA2EC0DC51FC082D45A388BA" baseAuthorityName="Verhoeff" baseAuthorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="9AB5A094BB3145CFDBAFA28555B24753" italics="true">Ballophilus</emphasis>
</taxonomicName>
from
<collectingCountry id="D482D995AC59F4B8FB02AA30C5F9B1DE" name="Madagascar">Madagascar</collectingCountry>
, of which
<taxonomicName id="7B4A189F3230F4860F2018562F0F3182" authority="Lawrence, 1960" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="C989F89EB2AA61C47691117850D2741E" italics="true">Ballophilus taeniaformis</emphasis>
<bibRefCitation id="EAEECCEA092CDED5DA2ECB94A098421B" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
</taxonomicName>
overlaps in nearly all diagnostic traits with the Aldabra material examined, differing only in the number of teeth on the dentate lamella of the mandible (
<quantity id="5AC04B20B51EF6DC49BFB270024B8747" metricMagnitude="-1" metricUnit="m" metricValue="1.778" unit="in" value="7.0">7 in</quantity>
the Aldabra specimen compared to eight or nine in
<taxonomicName id="641B066A6AC311390927684A756516ED" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="CE2CC084CE1DA497BC097736A6AAADDE" italics="true">B. taeniaformis</emphasis>
</taxonomicName>
). Lawrence made no comment on the presence or absence of the chitin line on the forcipular coxosternite for the species he assigned to
<taxonomicName id="09A462BFBFFEF354D208FCFD4F71975B" baseAuthorityName="Verhoeff" baseAuthorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="6B46FB12B38BA79023FCD2DA72A24AC0" italics="true">Ballophilus</emphasis>
</taxonomicName>
. As noted above, this character has been used to differentiate between the genera
<taxonomicName id="842FCFA09E70808368A138A9558430CD" baseAuthorityName="Verhoeff" baseAuthorityYear="1939" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="D5BFBE771B6518A2F2341C4B5E3CAB9A" italics="true">Ballophilus</emphasis>
</taxonomicName>
and
<taxonomicName id="5F9D0574A0A3DC6A8D71C732A26010BB" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="744FC1650007652A7EBD8FD11F10AB7E" italics="true">Ityphilus</emphasis>
</taxonomicName>
, although its variability as discussed above and the lack of consensus on the status of
<taxonomicName id="5237A0CC71C024BF8AE5E5B9F33BD600" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="F56686B683385750785933BEA1CC27BE" italics="true">Ityphilus</emphasis>
</taxonomicName>
at the time of Lawrences original description of
<taxonomicName id="FE6DED1855624833A31E5D1385F113FC" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Ballophilidae" genus="Ballophilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="taeniaformis">
<emphasis id="C82E9EBEC60106688EB478E6344DF6EE" italics="true">B. taeniaformis</emphasis>
</taxonomicName>
prompt us to refer the examined specimen from Aldabra to
<taxonomicName id="731F3E2E697D807B791331153E0C9393" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="B26FED17343F39C122DDFD032D5A96C7" italics="true">Ityphilus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CCB346204C69035E86077D606769240E">
In nearly all characters examined, the singular specimen from Aldabra agrees with the description of
<taxonomicName id="7F9307F6CDA10E434E428E232B51BA7C" authority="(Attems 1900)" baseAuthorityName="Attems" baseAuthorityYear="1900" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="melanostigma">
<emphasis id="A2C7FFDAEB460C905988061DE58F3E18" italics="true">Ityphilus melanostigma</emphasis>
(Attems 1900)
</taxonomicName>
and the subsequent redescription of this species from specimens collected in the
<collectingCountry id="6E98F7917445187B07CFB81E38305DEB" name="Seychelles">Seychelles</collectingCountry>
(
<bibRefCitation id="1D6ACC7493866F2A7D22E4D31D60EC1F" author="Bonato and Minelli" firstAuthor="Bonato" journalOrPublisher="Phelsuma" pagination="9-38" refId="B11" refString="Bonato L, Minelli A (2010) The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha). Phelsuma 18: 938." title="The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha)." volume="18" year="2010">Bonato and Minelli 2010</bibRefCitation>
). However, the greatly reduced number of leg-bearing segments (75) relative to the presently known range within
<taxonomicName id="E6AE852CDBE8E297CF6610147C4C9675" baseAuthorityName="Attems" baseAuthorityYear="1900" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="melanostigma">
<emphasis id="F2076BBF0AE9B8B8E6CFFB55B5243BD8" italics="true">I. melanostigma</emphasis>
</taxonomicName>
(95101) suggest specific distinction, as extensive variation in leg-bearing segment number is not generally known from other species of
<taxonomicName id="91A44601A3847D69804B3D25121BA281" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="3D06AD0857932FA34901C8B0AC481D07" italics="true">Ityphilus</emphasis>
</taxonomicName>
. Other putative differences to the original description of
<taxonomicName id="FC955B74F7621121E7A9DB04DA3C9B4D" baseAuthorityName="Attems" baseAuthorityYear="1900" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="melanostigma">
<emphasis id="D5DFD8752742A52D3DB21F7EB2A95A94" italics="true">I. melanostigma</emphasis>
</taxonomicName>
include the greater relative enlargement of the ultimate leg pair telopodites in the Aldabra specimen, relative to
<taxonomicName id="714D166921AB9BF8E6C2306434DEAB10" class="Chilopoda" family="Ballophilidae" genus="Ityphilus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="45760F5AF332A665E1D7A60CCD958CF6" italics="true">Ityphilus</emphasis>
</taxonomicName>
specimens illustrated from the
<collectingCountry id="1B66807CF3BB500FB420B5C2D6C97789" name="Seychelles">Seychelles</collectingCountry>
(
<bibRefCitation id="2ECC2AEDC6379C52193C52270D74D302" author="Bonato and Minelli" firstAuthor="Bonato" journalOrPublisher="Phelsuma" pagination="9-38" refId="B11" refString="Bonato L, Minelli A (2010) The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha). Phelsuma 18: 938." title="The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha)." volume="18" year="2010">Bonato and Minelli 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,213 @@
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<treatment id="7547F35C65035FE18A9BA80624AE48CC" ID-arpha="7547F35C-6503-5FE1-8A9B-A80624AE48CC" LSID="urn:lsid:plazi:treatment:7547F35C65035FE18A9BA80624AE48CC" httpUri="http://treatment.plazi.org/id/7547F35C65035FE18A9BA80624AE48CC" scope_class="Chilopoda" scope_family="Mecistocephalidae" scope_order="Geophilomorpha">
<subSubSection id="1FA51EA7F8DC8135C0506F5EDEBAD6CB" type="nomenclature">
<paragraph id="D79B73602ED962194B7240EE0600E74A">
<taxonomicName id="92B70AE692A7ED444AD3ED7A51B798D8" ID-CoL="3YKCJ" authority="Verhoeff, 1939" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lohmanderi">
<emphasis id="824FD4B0B4D025E0CDD68E06FD88CA50" italics="true">Mecistocephalus lohmanderi</emphasis>
<bibRefCitation id="2DB470C74740EDBEC6945385A9D7A084" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 72: 71-98." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff, 1939</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="2A85D4E290FC30CBBCCCE44D0BC9ED89" type="description">
<paragraph id="AF9D6E227A81125BF3D8EC5C5D3FCA42">
<figureCitation id="2BC4B81946FFCA944C19A58D282CDA37" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Mecistocephalus lohmanderi Verhoeff, 1939, NHMUK 015991460, forcipular segment, dorsal view B Mecistocephalus angusticeps (Ribaut, 1914), NHMUK 015991466, forcipular segment, dorsal view." figureDoi="10.3897/zookeys.1228.143007.figure11" httpUri="https://binary.pensoft.net/fig/1257830">Fig. 11 A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXEBI" type="material">
<paragraph id="E466503C0921A5841940FEA9D29B6532">
<heading id="A215050E3F52DAAC7E30DE467BA6B826" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="94967F9371C8107714C417EB5B4587B4">
<specimenCount id="64D9F4995F8AE1F6CFC40C1CAD4FD323" count="2" type="generic">2 specimens</specimenCount>
:
<materialsCitation id="4EF780995EFDC2333BD0F7DF249CD969" collectionCode="NHMUK" collectorName="V. W. Spaull" county="Aldabra" location="Black Path" municipality="Picard" specimenCode="NHMUK 015991459" specimenCount="1" specimenCount-female="1">
<specimenCode id="A145F5CD64BA4742031E53379B5F7499">NHMUK 015991459</specimenCode>
,
<specimenCount id="D270DB9595B2B081B29045F3F6A8FD85" count="1" type="female">1 ♀</specimenCount>
,
<location id="48C4FFF95E870589861CADCD96A9ABCF" LSID="urn:lsid:plazi:treatment:7547F35C65035FE18A9BA80624AE48CC:48C4FFF95E870589861CADCD96A9ABCF" county="Aldabra" municipality="Picard" name="Black Path">Black Path</location>
,
<collectingMunicipality id="7DA02EA8A3AFDAD537BC0E238920612B">Picard</collectingMunicipality>
,
<collectingCounty id="7684FC473EB8855E72CE44482C5A7E1C">Aldabra</collectingCounty>
, Summer 1975, leg.
<collectorName id="4CF5B79E74A4850CA3F30E3756CC6EF7">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="31556680A5A2317A23EE8DFEFFCB5FCF" collectedFrom="Casuarina litter" collectingDate="1974-12-10" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Picard" specimenCode="NHMUK 015991460" specimenCount="1" specimenCount-male="1">
<specimenCode id="E63FC354E82BD71D36E5DE3D777C29E6">NHMUK 015991460</specimenCode>
,
<specimenCount id="92A41401BD0F6609A1DFEE430D760FCA" count="1" type="male">1 ♂</specimenCount>
,
<collectedFrom id="E7147C1D99C16FAED96571014FC23040">
<taxonomicName id="1584AD287BEADB76149CED241B50443C" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="genus">
<emphasis id="518B0438F13C477909136A3D8EF4976F" italics="true">Casuarina</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="484E67D6E91FAFBD1C6150D49B637503" LSID="urn:lsid:plazi:treatment:7547F35C65035FE18A9BA80624AE48CC:484E67D6E91FAFBD1C6150D49B637503" name="Picard">Picard</location>
,
<date id="445A2285D2AECA175BF5A3B2A3B25C70" value="1974-12-10">
<collectingDate id="A3F41F0DD25B629864F521D55AE975BB" value="1974-12-10">10. 12. 1974</collectingDate>
</date>
, leg.
<collectorName id="A3651759BE7383972D9C3E87F562F54F">V. W. Spaull</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEFBI" type="remarks">
<paragraph id="31B0EC4A4558C872E6E41616507003AC">
<heading id="29A46599511B42EB9212DEC53C2C19F0" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="B84752E5247FC435EC81989D8BA1B1A5">
Originally described from
<collectingCountry id="E2CB47741AEFD187748C305027E74A6B" name="Mauritius">Mauritius</collectingCountry>
(
<bibRefCitation id="6F9B837575E563607340741DACC58F75" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 72: 71-98." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff, 1939</bibRefCitation>
),
<taxonomicName id="DA74993D2FCB84F339C91631BB716AAF" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lohmanderi">
<emphasis id="A910496B0C857D711A53C5659E0B0A60" italics="true">M. lohmanderi</emphasis>
</taxonomicName>
has been found on other Western Indian Ocean Islands (
<bibRefCitation id="949D1950EA807CA5011053BE272101E4" author="Bonato and Minelli" firstAuthor="Bonato" journalOrPublisher="Phelsuma" pagination="9-38" refId="B11" refString="Bonato L, Minelli A (2010) The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha). Phelsuma 18: 938." title="The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha)." volume="18" year="2010">Bonato and Minelli 2010</bibRefCitation>
;
<bibRefCitation id="1173B4ADA6242085792E70DAA5E4B7D7" DOI="10.1080/00222933.2024.2395903" author="Popovici" etAl="et al." firstAuthor="Popovici" issue="41-44" journalOrPublisher="Journal of Natural History" pagination="1885-1915" refId="B87" refString="Popovici G, Edgecombe GD, Hall DW (2024) New Chilopoda from the Chagos Archipelago. Journal of Natural History 58 (4144): 18851915. https://doi.org/10.1080/00222933.2024.2395903" title="New Chilopoda from the Chagos Archipelago." volume="58" year="2024">Popovici et al. 2024</bibRefCitation>
). Similarly to
<taxonomicName id="8DB92C9C2A3F4A7C1D2071C4CEEAAE58" baseAuthorityName="Ribaut" baseAuthorityYear="1914" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="66BDB469383AF7205E36E8F0502DD68C" italics="true">M. angusticeps</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="8F8615010BC5C65ED571A82D0D8B4274" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Mecistocephalus lohmanderi Verhoeff, 1939, NHMUK 015991460, forcipular segment, dorsal view B Mecistocephalus angusticeps (Ribaut, 1914), NHMUK 015991466, forcipular segment, dorsal view." figureDoi="10.3897/zookeys.1228.143007.figure11" httpUri="https://binary.pensoft.net/fig/1257830">11 B</figureCitation>
), examined specimens lack forcipular cerri (Fig.
<figureCitation id="D637E32B1292392CF28D7BB6ECB9A081" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A Mecistocephalus lohmanderi Verhoeff, 1939, NHMUK 015991460, forcipular segment, dorsal view B Mecistocephalus angusticeps (Ribaut, 1914), NHMUK 015991466, forcipular segment, dorsal view." figureDoi="10.3897/zookeys.1228.143007.figure11" httpUri="https://binary.pensoft.net/fig/1257830">11 A</figureCitation>
). Presently examined specimens are most similar to
<taxonomicName id="A25160E8DE09D52E6234A2FF7BCC5C38" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lohmanderi">
<emphasis id="D1F3B1D0BAC5398A4323A8A8815AD3EE" italics="true">M. lohmanderi</emphasis>
</taxonomicName>
specimens collected from the Diego Garcia atoll (
<bibRefCitation id="1F0DCA4CEBC2CB670D2CF576EA336C9E" DOI="10.1080/00222933.2024.2395903" author="Popovici" etAl="et al." firstAuthor="Popovici" issue="41-44" journalOrPublisher="Journal of Natural History" pagination="1885-1915" refId="B87" refString="Popovici G, Edgecombe GD, Hall DW (2024) New Chilopoda from the Chagos Archipelago. Journal of Natural History 58 (4144): 18851915. https://doi.org/10.1080/00222933.2024.2395903" title="New Chilopoda from the Chagos Archipelago." volume="58" year="2024">Popovici et al. 2024</bibRefCitation>
). Adults (female
<quantity id="AA380EBA9F3ACB72468813B4FDDEF76A" metricMagnitude="-2" metricUnit="m" metricValue="3.4" unit="mm" value="34.0">34 mm</quantity>
body length, male
<quantity id="1DC5FCB258D7B29CCA81B5DF53E6AFA5" metricMagnitude="-2" metricUnit="m" metricValue="3.4" unit="mm" value="34.0">34 mm</quantity>
body length) in the Aldabra sample lack dark subcutaneous pigment patches and maintain the characteristic clypeal setation pattern described in
<taxonomicName id="9AF772424EAC53E9A889354B1249FDB9" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lohmanderi">
<emphasis id="FDEB2F98064968D861F7E855D090C6E4" italics="true">M. lohmanderi</emphasis>
</taxonomicName>
from other localities (
<bibRefCitation id="B39342F1C22BB66BC97565D3FCAB1E4D" author="Bonato and Minelli" firstAuthor="Bonato" journalOrPublisher="Phelsuma" pagination="9-38" refId="B11" refString="Bonato L, Minelli A (2010) The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha). Phelsuma 18: 938." title="The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha)." volume="18" year="2010">Bonato and Minelli 2010</bibRefCitation>
;
<bibRefCitation id="5356DBD4AD02C5D490FF5E0ADA80FD02" DOI="10.1080/00222933.2024.2395903" author="Popovici" etAl="et al." firstAuthor="Popovici" issue="41-44" journalOrPublisher="Journal of Natural History" pagination="1885-1915" refId="B87" refString="Popovici G, Edgecombe GD, Hall DW (2024) New Chilopoda from the Chagos Archipelago. Journal of Natural History 58 (4144): 18851915. https://doi.org/10.1080/00222933.2024.2395903" title="New Chilopoda from the Chagos Archipelago." volume="58" year="2024">Popovici et al. 2024</bibRefCitation>
). Similarly, specimens assigned to
<taxonomicName id="D4195A7B0E0729A81BE9165B41DBC5F5" authorityName="Lucas" authorityYear="1863" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="120444CB343D3FC34013A673F9DFB08B" italics="true">M. insularis</emphasis>
</taxonomicName>
described from
<collectingCountry id="4A1E7E53B9817F5C7577A4C058717BC0" name="Yemen">Socotra</collectingCountry>
(
<bibRefCitation id="7D3E972D83601FBD23428085D7B50C81" DOI="10.1080/09397140.1990.10637589" author="Lewis and Wranik" firstAuthor="Lewis" journalOrPublisher="Zoology in The Middle East" pagination="61-70" refId="B64" refString="Lewis JGE, Wranik W (1990) On the centipedes of Yemen. Zoology in The Middle East 4: 6170. https://doi.org/10.1080/09397140.1990.10637589" title="On the centipedes of Yemen." volume="4" year="1990">Lewis and Wranik 1990</bibRefCitation>
) match all diagnostic characters outlined for
<taxonomicName id="C72AB9CB831BE70CF04D81A852D1E727" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lohmanderi">
<emphasis id="6579F3DF0708C2379B9A29E4842AB57E" italics="true">M. lohmanderi</emphasis>
</taxonomicName>
, and can be referred to this taxon, thus extending its range to island localities in the Northwestern Indian Ocean.
</paragraph>
<paragraph id="9AB99CFEF757293DF0D8B9C56E8A9900">
Records of large adult specimens (
<quantity id="87D1C20F097FCB18D035F1472AE7C777" metricMagnitude="-2" metricUnit="m" metricValue="8.05" metricValueMax="9.1" metricValueMin="7.0" unit="mm" value="80.5" valueMax="91.0" valueMin="70.0">7091 mm</quantity>
) assigned to
<taxonomicName id="A742A07E37206DAAA48D66D952A80286" authorityName="Lucas" authorityYear="1863" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="0321FC1B3A7F4B77F7FA74C26055826F" italics="true">Mecistocephalus insularis</emphasis>
</taxonomicName>
from the Arabian Peninsula (
<bibRefCitation id="457BF634DC20E2B93ACDDAB2CE07E410" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Fauna of Saudi Arabia" pagination="137-156" refId="B54" refString="Lewis JGE (1996) Further records of scolopendromorph and geophilomorph centipedes from the Arabian Peninsula with a note by Dr. E. H. Eason on Lithobius erythrocephalus cronebergii Sseliwanoff. Fauna of Saudi Arabia 15: 137156." title="Further records of scolopendromorph and geophilomorph centipedes from the Arabian Peninsula with a note by Dr. E. H. Eason on Lithobius erythrocephalus cronebergii Sseliwanoff." volume="15" year="1996">Lewis 1996</bibRefCitation>
) are fully consistent with the revised description of
<taxonomicName id="13475EDBA14B503B22537929C6F85D86" authority="Attems, 1900" authorityName="Attems" authorityYear="1900" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="glabridorsalis">
<emphasis id="200BBBBDFEE69AD32D24AFE692099360" italics="true">Mecistocephalus glabridorsalis</emphasis>
Attems, 1900
</taxonomicName>
from the
<collectingCountry id="6CA7A4B857208579251CBC6642907646" name="Seychelles">Seychelles</collectingCountry>
(
<bibRefCitation id="016511E542354ED6589BE1076AB481E7" author="Bonato and Minelli" firstAuthor="Bonato" journalOrPublisher="Phelsuma" pagination="9-38" refId="B11" refString="Bonato L, Minelli A (2010) The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha). Phelsuma 18: 938." title="The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha)." volume="18" year="2010">Bonato and Minelli 2010</bibRefCitation>
) and are almost certainly misidentified
<taxonomicName id="1F4BB8B0E674B8AC7648797EC77D0163" authorityName="Attems" authorityYear="1900" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="glabridorsalis">
<emphasis id="F31A66C1E803BF352BF953C2CCE401B6" italics="true">M. glabridorsalis</emphasis>
</taxonomicName>
. In particular, the clypeal morphology illustrated for specimens from
<collectingCountry id="55DBB19BB2E846C71E755031DE3E5F31" name="Saudi Arabia">Saudi Arabia</collectingCountry>
shares the presence of a small non-areolate insula anterior to the plagulae with specimens from the
<collectingCountry id="B5EA1F7B4AC0F9C9BA2C6EC9A7A24E08" name="Seychelles">Seychelles</collectingCountry>
and the arrangement of setae in a transverse line on the areolate part of the clypeus. This morphology has hitherto only been recorded in
<taxonomicName id="60521986A19B685775BFB94B25B854EB" authorityName="Attems" authorityYear="1900" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="glabridorsalis">
<emphasis id="0043AC8379B3B2CCE5765DAB80398D02" italics="true">M. glabridorsalis</emphasis>
</taxonomicName>
and
<taxonomicName id="B3925F030252DF27AECEAA4937002D8E" authorityName="Newport" authorityYear="1843" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="punctifrons">
<emphasis id="20644DC711331A100DC54F78F3B7DCF5" italics="true">M. punctifrons</emphasis>
</taxonomicName>
<bibRefCitation id="A1126E14CD18811D5E37EFBF7864F809" DOI="10.1080/03745484409442576" author="Newport G" journalOrPublisher="Annals and Magazine of Natural History" pagination="94-101" refId="B71" refString="Newport G (1843) On some new genera of the class Myriapoda. Proceedings of the Zoological Society of London, 10 (1842): 177 - 181. Newport G. 1844. A list of the species of Myriapoda, order Chilopoda, contained in the cabinets of the British Museum, with a synoptic description of forty-seven species. Annals and Magazine of Natural History 13: 94-101. https://doi.org/10.1080/03745484409442576" title="On some new genera of the class Myriapoda. Proceedings of the Zoological Society of London, 10 (1842): 177-181. Newport G. 1844. A list of the species of Myriapoda, order Chilopoda, contained in the cabinets of the British Museum, with a synoptic description of forty-seven species." volume="13" year="1843">Newport, 1843</bibRefCitation>
(
<bibRefCitation id="E988AF01A44901A98D10E17A88984113" DOI="10.1080/03946975.2004.10531198" author="Bonato and Minelli" firstAuthor="Bonato" issue="1" journalOrPublisher="Tropical Zoology" pagination="15-63" refId="B10" refString="Bonato L, Minelli A (2004) The centipede genus Mecistocephalus Newport 1843 in the Indian Peninsula (Chilopoda Geophilomorpha Mecistocephalidae). Tropical Zoology 17 (1): 1563. https://doi.org/10.1080/03946975.2004.10531198" title="The centipede genus Mecistocephalus Newport 1843 in the Indian Peninsula (Chilopoda Geophilomorpha Mecistocephalidae)." volume="17" year="2004">Bonato and Minelli 2004</bibRefCitation>
), casting further doubt on the true identity and distribution of
<taxonomicName id="23DAD959A414EFE263B7DE9D8986F209" authorityName="Lucas" authorityYear="1863" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="D871A0D5F36C97FEADC0C29F5BF0EFB1" italics="true">M. insularis</emphasis>
</taxonomicName>
. Insufficient data on morphological variability in
<taxonomicName id="8B351C58C0C43D67576E8D827F36D1A4" authorityName="Verhoeff" authorityYear="1939" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="lohmanderi">
<emphasis id="0A3E9EF569E0C9C0672E01E4ADCB8365" italics="true">M. lohmanderi</emphasis>
</taxonomicName>
and the uncertain status of
<taxonomicName id="3B92A3863342E04F0EC58E54B495CD75" authorityName="Lucas" authorityYear="1863" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="C94615056F956CBFA554CB67CE42A4E4" italics="true">M. insularis</emphasis>
</taxonomicName>
records from past literature prevent further inferences on the taxonomic validity and relationships between these species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,190 @@
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<subSubSection id="6A2443AE2DA4EF1C66A9073FC1081A83" type="nomenclature">
<paragraph id="1BC767E7E6419D647A0C872601E2E3AD">
<taxonomicName id="AAB058C53BC6DB4DCD6B3D70F449728E" ID-CoL="4SY3P" authority="Attems, 1952" authorityName="Attems" authorityYear="1952" class="Chilopoda" family="Geophilidae" genus="Ribautia" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="paucipes">
<emphasis id="C87F4360537C82DAC5C3FD8EB24EC5DE" italics="true">Ribautia</emphasis>
cf.
<emphasis id="955666816B1BE4011470CDC3ADDAC2AF" italics="true">paucipes</emphasis>
<bibRefCitation id="87E9CC822A074116C0A18677B1F16858" author="Attems C" journalOrPublisher="Annales du Musée Royal du Congo Belge, Tervuren (Belgique)" pagination="1-139" refId="B5" refString="Attems C (1952) Neue Myriopoden des Belgischen Congo. Annales du Musee Royal du Congo Belge, Tervuren (Belgique) 18: 1-139." title="Neue Myriopoden des Belgischen Congo." volume="18" year="1952">Attems, 1952</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="153D3712593EE65436A0E16FA965A387" type="description">
<paragraph id="334E6502AB4DB6E8951D1451D120487C">
<figureCitation id="E8A66D718FEAB47E230C6CE2E8DDC1C7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ribautia cf. paucipes Attems, 1952. NHMUK 015991467 A head and forcipular segment, ventral view B anterior margin of second maxillary coxosternite, ventral view C calyx of venom gland, lateral view D ultimate leg-bearing and postpedal segments, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure5" httpUri="https://binary.pensoft.net/fig/1257824">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3KBG" type="material">
<paragraph id="9FFDF65097EB53F1FD571589071E1E52">
<heading id="7A24834E9CD5AB9A0B5ED9FF7B19AA20" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="D3D432F49754461BC79BF588C1ECE475">
<materialsCitation id="A0A5D85999C36E485D4C7B91C6F29891" collectedFrom="Casuarina litter" collectingDate="1974-12-10" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Picard" municipality="Aldabra" specimenCode="NHMUK 015991467" specimenCount="1" specimenCount-juvenile="1">
<specimenCode id="E61FE208960D715BC060576E51986F79">NHMUK 015991467</specimenCode>
,
<specimenCount id="C754919474C4BE976EF0145032C4B7B9" count="1" type="juvenile">1 juvenile</specimenCount>
,
<collectedFrom id="80B3F8616825D178ED87169A355C63A0">
<taxonomicName id="5FBC2A6CEEC2880094E2345C0BEC7295" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="genus">
<emphasis id="82D0D07878D4C5FC6ADAE2506DFD5376" italics="true">Casuarina</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="8221E809BFBDC25BFA65A7747DCD9CFF" LSID="urn:lsid:plazi:treatment:8558B2970A0059EF8D8D9517C5F0B6D5:8221E809BFBDC25BFA65A7747DCD9CFF" municipality="Aldabra" name="Picard">Picard</location>
,
<collectingMunicipality id="6CFEE46B6F8D7CA187EC728945870FBB">Aldabra</collectingMunicipality>
,
<date id="6D8016B328B55B1325793950B5B77A00" value="1974-12-10">
<collectingDate id="20D9A2EB3C7EDD6A4D2C789FF09A445B" value="1974-12-10">10. 12. 1974</collectingDate>
</date>
, leg.
<collectorName id="8557AA61DDCEC6AFA5DDAACB1428091D">V. W. Spaull</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJLBG" type="remarks">
<paragraph id="2C6C09736783E0F2D14F3F79EC495A74">
<heading id="F95BCFB1A49DFA26FBA9C151CC9DF684" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="215D76D1978104A7B94A0A79121705B0">
The sexually immature specimen found in the present sample displays all diagnostic characters that support its assignment to
<taxonomicName id="0755C14BA19547FCBB62F685F8D1E00D" authorityName="Brolemann" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Ribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="66DC4C1854D7C743C8DB03A2F59FE52C" italics="true">Ribautia</emphasis>
</taxonomicName>
, comprising an elongate cephalic shield, lack of lappets on the first maxillae, the two halves of the second maxillary coxosternite being united by a sclerotised isthmus and the pleural sutures of the forcipular coxosternite being parallel to its lateral edge distally. A potentially novel ontogenetic observation is the incomplete separation of the two halves of the second maxillary coxosternite by an isthmus (Fig.
<figureCitation id="48A6F16C1880456594C2CE8DDCC06F96" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Ribautia cf. paucipes Attems, 1952. NHMUK 015991467 A head and forcipular segment, ventral view B anterior margin of second maxillary coxosternite, ventral view C calyx of venom gland, lateral view D ultimate leg-bearing and postpedal segments, ventral view." figureDoi="10.3897/zookeys.1228.143007.figure5" httpUri="https://binary.pensoft.net/fig/1257824">5 B</figureCitation>
), as is a characteristic of
<taxonomicName id="C2137EE257EB6B134F2273EDAF1A63FD" authorityName="Brolemann" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Ribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="E8B3BD941ABE2FB5B704FB32F8635D0E" italics="true">Ribautia</emphasis>
</taxonomicName>
. Taking into account the very small size (length
<quantity id="A5165A0FF93A465BE6BE1DE33BC63EFA" metricMagnitude="-3" metricUnit="m" metricValue="7.0" unit="mm" value="7.0">7 mm</quantity>
) and sexual immaturity of the specimen, this may be a character state that becomes more conspicuous in older individuals.
</paragraph>
<caption id="97FEF1699A22A841EB2F119E01AF3361" ID-DOI="10.3897/zookeys.1228.143007.figure5" ID-arpha="3935CE69-3843-5004-8BB1-B31725204CD5" httpUri="https://binary.pensoft.net/fig/1257824" startId="F5">
<paragraph id="49A4D40D612DB88D4162F1067E7F298A">
<label id="1FC110605406F290B2E82AAC956DFAD3">Figure 5.</label>
</paragraph>
<paragraph id="C902F887A2BFF60990E00E3339688AB3">
<taxonomicName id="CE5F7D9553866B731E9E4003E58BFD40" authority="Attems, 1952" authorityName="Attems" authorityYear="1952" class="Chilopoda" family="Geophilidae" genus="Ribautia" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="paucipes">
<emphasis id="A2667096CE8E9638D9642C9E3AFD9537" italics="true">Ribautia</emphasis>
cf.
<emphasis id="C8F05B48C88F545787E071EC71674652" italics="true">paucipes</emphasis>
<bibRefCitation id="6D20229FB1F23FA16AE0D4B345E1AFD3" author="Attems C" journalOrPublisher="Annales du Musée Royal du Congo Belge, Tervuren (Belgique)" pagination="1-139" refId="B5" refString="Attems C (1952) Neue Myriopoden des Belgischen Congo. Annales du Musee Royal du Congo Belge, Tervuren (Belgique) 18: 1-139." title="Neue Myriopoden des Belgischen Congo." volume="18" year="1952">Attems, 1952</bibRefCitation>
</taxonomicName>
.
<specimenCode id="333CED98A9F01DACF4FC9B1D1651DC73">NHMUK 015991467</specimenCode>
<emphasis id="575307B3CB19EA918B233D5CBC123DB2" bold="true">A</emphasis>
head and forcipular segment, ventral view
<emphasis id="9AC77624E9BCFCD68C209A92A940B877" bold="true">B</emphasis>
anterior margin of second maxillary coxosternite, ventral view
<emphasis id="5562D2E7D6BC6F37EB6CB3914174E7DE" bold="true">C</emphasis>
calyx of venom gland, lateral view
<emphasis id="F03BDECAF8441FF39F967F01B065CB7B" bold="true">D</emphasis>
ultimate leg-bearing and postpedal segments, ventral view.
</paragraph>
</caption>
<paragraph id="167F6DF18D0D8C918E149E61DC7ED4C8">
Beside the very low number of leg-bearing segments (37), which is shared with
<taxonomicName id="3E5D49F29169DF7E65288DD62020F4BE" authorityName="Attems" authorityYear="1952" class="Chilopoda" family="Geophilidae" genus="Ribautia" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="paucipes">
<emphasis id="A0F46D31EC6DFE3B8B6FA62B1CEABEBD" italics="true">R.</emphasis>
cf.
<emphasis id="26AA8315BC6AAF8248C45FEB4F166D74" italics="true">paucipes</emphasis>
</taxonomicName>
reported from the
<collectingCountry id="89193FB39310485D1A677B1BFCC35426" name="Seychelles">Seychelles</collectingCountry>
(
<bibRefCitation id="E9F243E3D04FDBC013BF5BB5316A1539" author="Bonato and Minelli" firstAuthor="Bonato" journalOrPublisher="Phelsuma" pagination="9-38" refId="B11" refString="Bonato L, Minelli A (2010) The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha). Phelsuma 18: 938." title="The geophilomorph centipedes of the Seychelles (Chilopoda: Geophilomorpha)." volume="18" year="2010">Bonato and Minelli 2010</bibRefCitation>
), the developmental stage of the present specimen does not allow for satisfactory evaluation of potential morphological differences between
<taxonomicName id="F40EBB48155265DF2130324566924347" authorityName="Brolemann" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Ribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="B3AA81E3A6C2F1C08E3C9A60B976C670" italics="true">Ribautia</emphasis>
</taxonomicName>
specimens described from Western Indian Ocean localities and type material of
<taxonomicName id="F7B69BB4EC32F4D2098522E291263251" authorityName="Attems" authorityYear="1952" class="Chilopoda" family="Geophilidae" genus="Ribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="paucipes">
<emphasis id="CF846A3B57341ABFC17D4EB9B3648B95" italics="true">Ribautia paucipes</emphasis>
</taxonomicName>
described from the environs of Lake Kivu in Central Africa. Notably, the Aldabra specimen lacks evident denticles on the anterior margin of the forcipular coxosternite and the interior margin of the forcipular trochanteroprefemur, although these are clearly illustrated in the original description of
<taxonomicName id="C269B544AF8CCC7A3997014D6BADFE6D" authorityName="Attems" authorityYear="1952" class="Chilopoda" family="Geophilidae" genus="Ribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="paucipes">
<emphasis id="3ACA5FE467F323947C0F63BF16E1DB92" italics="true">R. paucipes</emphasis>
</taxonomicName>
(
<bibRefCitation id="CB4B4D307A8AA0DF3DDA4DD564D0BC17" author="Attems" firstAuthor="Attems" journalOrPublisher="Annales du Musée Royal du Congo Belge, Tervuren (Belgique)" pagination="1-139" refId="B5" refString="Attems C (1952) Neue Myriopoden des Belgischen Congo. Annales du Musée Royal du Congo Belge, Tervuren (Belgique) 18: 1139." title="Neue Myriopoden des Belgischen Congo." volume="18" year="1952">Attems 1952</bibRefCitation>
: fig. 203). The specimen also lacks conspicuous coxal organs or coxal pores, a character state not noted in
<taxonomicName id="1448C5B3E858C4F75E9FA7C0ACBDB264" authorityName="Attems" authorityYear="1952" class="Chilopoda" family="Geophilidae" genus="Ribautia" isUncertain="true" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="paucipes">
<emphasis id="D824382739807C5250A7710C5F952205" italics="true">R.</emphasis>
cf.
<emphasis id="278C5F3D5CB3E5AD8395D70C30608CAC" italics="true">paucipes</emphasis>
</taxonomicName>
recorded from the
<collectingCountry id="AED52E95247B13043E1444288FB27151" name="Seychelles">Seychelles</collectingCountry>
. Ontogenetic variation in the number of coxal pores has been well-documented for other geophilid centipedes (
<bibRefCitation id="33A3171EAB0FAC59D7CB09064DA5E06F" author="Horneland and Meidell" firstAuthor="Horneland" issue="3" journalOrPublisher="Soil Organisms" pagination="373-386" refId="B36" refString="Horneland EO, Meidell B (2009) Postembryonic development of Strigamia maritima (Leach, 1817) (Chilopoda: Geophilomorpha, Linotaeniidae) with emphasis on how to separate the different stadia. Soil Organisms 81 (3): 373386. https://soil-organisms.org/index.php/SO/article/view/34" title="Postembryonic development of Strigamia maritima (Leach, 1817) (Chilopoda: Geophilomorpha, Linotaeniidae) with emphasis on how to separate the different stadia." url="https://soil-organisms.org/index.php/SO/article/view/34" volume="81" year="2009">Horneland and Meidell 2009</bibRefCitation>
;
<bibRefCitation id="B5712C0CDE9202DB13C71675B872BA52" author="Gregory and Barber" firstAuthor="Gregory" journalOrPublisher="Bulletin of the British Myriapod and Isopod Group" pagination="2-15" refId="B30" refString="Gregory SJ, Barber AD (2010) Observations of a population, including juveniles, of Geophilus carpophagus Leach, 1815, sensu stricto from Oxfordshire. Bulletin of the British Myriapod and Isopod Group 24: 215." title="Observations of a population, including juveniles, of Geophilus carpophagus Leach, 1815, sensu stricto from Oxfordshire." volume="24" year="2010">Gregory and Barber 2010</bibRefCitation>
;
<bibRefCitation id="E4C0FD8D51166AB39BFD2F9D5AED904E" DOI="10.1186/s12983-014-0058-9" author="Brena" firstAuthor="Brena" refId="B14" refString="Brena C (2014) The embryoid development of Strigamia maritima and its bearing on post-embryonic segmentation of geophilomorph centipedes. Frontiers in Zoology 11: 58. https://doi.org/10.1186/s12983-014-0058-9" year="2014">Brena 2014</bibRefCitation>
;
<bibRefCitation id="60677AE9EBD93DF1DBB6CE38D69D8BA6" DOI="10.1080/07924259.2020.1726514" author="Stojanovic" etAl="et al." firstAuthor="Stojanovic" issue="2" journalOrPublisher="Invertebrate Reproduction &amp; Development" pagination="115-125" refId="B99" refString="Stojanović DZ, Mitić BM, Dudić BD, Gedged AM, Tomić VT, Antić DŽ, Makarov SE (2020) Early development of the centipede Geophilus serbicus (Chilopoda: Geophilomorpha: Geophilidae) from the Balkan Peninsula. Invertebrate Reproduction &amp; Development 64 (2): 115125. https://doi.org/10.1080/07924259.2020.1726514" title="Early development of the centipede Geophilus serbicus (Chilopoda: Geophilomorpha: Geophilidae) from the Balkan Peninsula." volume="64" year="2020">Stojanović et al. 2020</bibRefCitation>
), increasing with body size at each postembryonic stage, and being absent in comparably sized adolescens stages of some species (
<bibRefCitation id="EFCC956E7999FA44A967EF04209E537B" author="Gregory and Barber" firstAuthor="Gregory" journalOrPublisher="Bulletin of the British Myriapod and Isopod Group" pagination="2-15" refId="B30" refString="Gregory SJ, Barber AD (2010) Observations of a population, including juveniles, of Geophilus carpophagus Leach, 1815, sensu stricto from Oxfordshire. Bulletin of the British Myriapod and Isopod Group 24: 215." title="Observations of a population, including juveniles, of Geophilus carpophagus Leach, 1815, sensu stricto from Oxfordshire." volume="24" year="2010">Gregory and Barber 2010</bibRefCitation>
). Additionally, the second maxillary pretarsus of the present specimen is markedly shorter and less acuminate than illustrated for
<taxonomicName id="E0C799B8DF550B9D0111069A3CBB82AA" authorityName="Attems" authorityYear="1952" class="Chilopoda" family="Geophilidae" genus="Ribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="paucipes">
<emphasis id="CBD70F767B6FF47F4D3A8F6F8F4CCBDC" italics="true">R. paucipes</emphasis>
</taxonomicName>
from continental Africa. As the ontogenetic variation of the morphology of the second maxillae in
<taxonomicName id="D5A5BDBB90BA17A59E5A09767B747D0C" authorityName="Brolemann" authorityYear="1909" class="Chilopoda" family="Geophilidae" genus="Ribautia" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="genus">
<emphasis id="22ACCD49F7AEA0992BC98369380FEFCE" italics="true">Ribautia</emphasis>
</taxonomicName>
is not presently known and in other characters the specimen strongly resembles individuals described from the
<collectingCountry id="8D5C60CC95B6CAC9F67BF37EE2026C13" name="Seychelles">Seychelles</collectingCountry>
, we maintain its presently assigned identity.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,48 +1,48 @@
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<treatment id="8692BA1F9EAF5459A438AB09F9D650CD" ID-DOI="http://doi.org/10.5281/zenodo.14907696" ID-Zenodo-Dep="14907696" ID-arpha="8692BA1F-9EAF-5459-A438-AB09F9D650CD" LSID="urn:lsid:plazi:treatment:8692BA1F9EAF5459A438AB09F9D650CD" httpUri="http://treatment.plazi.org/id/8692BA1F9EAF5459A438AB09F9D650CD" scope_class="Chilopoda" scope_family="Mecistocephalidae" scope_order="Geophilomorpha">
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<paragraph id="D7555F5663B14BA8A417E7987FDAFD66">
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@ -65,73 +65,123 @@
<paragraph id="FB0131777E0F45A33429970889EBD0AE">
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:
<collectionCode id="7E9C7026429789956ADC2276DDBF4002" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991461,
<materialsCitation id="890EDDD516CE0E8F129A6DE8DF0E4398" collectedFrom="Pandanus tectorius soil and litter" collectingDate="1974-03-24" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Cinq Cases" specimenCode="NHMUK 015991461" specimenCount="1" specimenCount-male="1">
<specimenCode id="7A00B923249A93552D69CC142DA22B2D">NHMUK 015991461</specimenCode>
,
<specimenCount id="44EAEDA379A48804E92BA3336E779D3A" count="1" type="male">1 ♂</specimenCount>
,
<collectedFrom id="321A1888656A857309FD5E3521C4C322">
<taxonomicName id="4CBF3C3D59DBEF7050CCF2B48FB034BA" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" phylum="Tracheophyta" rank="species" species="tectorius">
<emphasis id="CAA24BA0D13C1AEE0F935E1CB92C55B9" italics="true">Pandanus tectorius</emphasis>
</taxonomicName>
soil and litter, Cinq Cases,
<date id="F40FB19066D0D19FA989CDBB0AB184F7" value="1974-03-24">24. 03. 1974</date>
soil and litter
</collectedFrom>
,
<location id="14CD85C5AC980719981751E3EF59155B" LSID="urn:lsid:plazi:treatment:8692BA1F9EAF5459A438AB09F9D650CD:14CD85C5AC980719981751E3EF59155B" name="Cinq Cases">Cinq Cases</location>
,
<date id="F40FB19066D0D19FA989CDBB0AB184F7" value="1974-03-24">
<collectingDate id="CD8BD51ECCCA58CF84BC68FF68795AF8" value="1974-03-24">24. 03. 1974</collectingDate>
</date>
, leg.
<collectionCode id="4E392863EEC2D6FF537E5BDF8124B1D5" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. W. Spaull;
<collectionCode id="C3E41689380C1E6820CC68A812705B7D" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991462,
<collectorName id="A29181ADBA60C553BA2AB0E0ED732748">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="6FE1E7A445761C18562762D4771D7E19" collectingDate="1974-03-11" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Cinq Cases" specimenCode="NHMUK 015991462" specimenCount="1" specimenCount-female="1">
<specimenCode id="9C66D2FA65C683EDC1BDD76999971D7D">NHMUK 015991462</specimenCode>
,
<specimenCount id="07D673F22EABCD125BC1710F98360CF9" count="1" type="female">1 ♀</specimenCount>
,
<locationDeviation id="5C7D64531A94439694273BA418D2A489" bearing="0.0" distance="2000.0">
<quantity id="A89AD89F056841E070A3FD6741735B3D" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
N of Cinq Cases,
<date id="C39F4B6C60F1F6D3BDE096419A3FA232" value="1974-03-11">11. 03. 1974</date>
N of
</locationDeviation>
<location id="2883650BEB74998D67D54BF86C478158" LSID="urn:lsid:plazi:treatment:8692BA1F9EAF5459A438AB09F9D650CD:2883650BEB74998D67D54BF86C478158" name="Cinq Cases">Cinq Cases</location>
,
<date id="C39F4B6C60F1F6D3BDE096419A3FA232" value="1974-03-11">
<collectingDate id="D255BCFFA2073B49B6245BC2E9590EBF" value="1974-03-11">11. 03. 1974</collectingDate>
</date>
, leg.
<collectionCode id="37722756E9989C63863B54D848E9AC5E" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. W. Spaull;
<collectionCode id="F06809DE3DAD15E6331D86087AA7CECE" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991463,
<collectorName id="BADF84D3A29EB8078FB742D4764B7D21">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="87B1FF2B57A948E84AE8C66EFCA2BFCE" collectedFrom="Sideroxylon litter" collectingDate="1974-03-10" collectingDateMax="1974-03-17" collectingDateMin="1974-03-10" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Cinq Cases" specimenCode="NHMUK 015991463" specimenCount="6" specimenCount-juvenile="6">
<specimenCode id="9130BCE70505B58E86C25D96E7E01CFE">NHMUK 015991463</specimenCode>
,
<specimenCount id="9356BDF2D2F6674B062E73FC03B01DBF" count="6" type="juvenile">6 juveniles</specimenCount>
,
<collectedFrom id="F276C2237BF6CB531841F3F839CFCDA9">
<taxonomicName id="1448D3E0B91ACFA122A616447FBF7F84" class="Magnoliopsida" family="Sapotaceae" genus="Sideroxylon" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="genus">
<emphasis id="CC632BE5389336D19331B18DD19D9145" italics="true">Sideroxylon</emphasis>
</taxonomicName>
litter, Cinq Cases,
<date id="57CE28ABFEC4606AFAC5586E817A6AE7" value="1974-03-10" valueMax="1974-03-17" valueMin="1974-03-10">1017.03. 1974</date>
litter
</collectedFrom>
,
<location id="F0FED46F7C5E2AB7AB31CF7BF9FF9849" LSID="urn:lsid:plazi:treatment:8692BA1F9EAF5459A438AB09F9D650CD:F0FED46F7C5E2AB7AB31CF7BF9FF9849" name="Cinq Cases">Cinq Cases</location>
,
<date id="57CE28ABFEC4606AFAC5586E817A6AE7" value="1974-03-10" valueMax="1974-03-17" valueMin="1974-03-10">
<collectingDate id="DB6A8F24FE52D2D4079FEFCD84A45C23" value="1974-03-10" valueMax="1974-03-17" valueMin="1974-03-10">1017.03. 1974</collectingDate>
</date>
, leg.
<collectionCode id="2019E518663D9A8815E3FE26F437D20A" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. W. Spaull;
<collectionCode id="68DAAC1CA1EE98644729F44154DFE85B" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991464,
<collectorName id="D6B5CCD243C8FE898BBEDA98C348D5ED">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="F0BDB980B9442EAF9DA094DA2F2167D6" collectedFrom="Pandanus tectorius soil and litter" collectingDate="1974-03-27" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Cinq Cases / Point Hodoul" specimenCode="NHMUK 015991464" specimenCount="1" specimenCount-female="1">
<specimenCode id="EEBACB836659608B54F3F929289D8299">NHMUK 015991464</specimenCode>
,
<specimenCount id="514F4EB64F26CD6C9779548B41050F86" count="1" type="female">1 ♀</specimenCount>
,
<collectedFrom id="D6C1071232D676F29C3ABC5D2887E465">
<taxonomicName id="707F0188DE7C03CDF259CF0FCD35296B" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" phylum="Tracheophyta" rank="species" species="tectorius">
<emphasis id="799F2C7B922AB6084E66CF34C91946A7" italics="true">Pandanus tectorius</emphasis>
</taxonomicName>
soil and litter, Cinq Cases / Point Hodoul,
<date id="740D5E47F40B1FAA80EB1EF86F1B4E9A" value="1974-03-27">27. 03. 1974</date>
soil and litter
</collectedFrom>
,
<location id="9B284CDA7C34853772F6137EAFBF781B" LSID="urn:lsid:plazi:treatment:8692BA1F9EAF5459A438AB09F9D650CD:9B284CDA7C34853772F6137EAFBF781B" name="Cinq Cases / Point Hodoul">Cinq Cases / Point Hodoul</location>
,
<date id="740D5E47F40B1FAA80EB1EF86F1B4E9A" value="1974-03-27">
<collectingDate id="45BCB0B5BD295CB2E89591F0A2E69C88" value="1974-03-27">27. 03. 1974</collectingDate>
</date>
, leg.
<collectionCode id="3696C3D2278DCBD1929774B08BBC1377" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. W. Spaull;
<collectionCode id="50B6A7CAED963873E78EFE861A65F2D9" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991465,
<collectorName id="7E7721F1D86BB1B3A3013017DF6A8B50">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="1DAA372925A779DB30D93B20319D54C3" collectingDate="1974-12-03" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Gionnet" municipality="Aldabra" specimenCode="NHMUK 015991465" specimenCount="1" specimenCount-male="1">
<specimenCode id="67F96BEF9E1FF718656A6DE54548A5B3">NHMUK 015991465</specimenCode>
,
<specimenCount id="02F9A232CA95BE249EE820BEF35722F8" count="1" type="male">1 ♂</specimenCount>
, Gionnet, Aldabra,
<date id="C16B78BA2F0ED6F041E49EE8B145543F" value="1974-12-03">03. 12. 1974</date>
,
<location id="35E4CCA12248D69517094EB2C3A857C6" LSID="urn:lsid:plazi:treatment:8692BA1F9EAF5459A438AB09F9D650CD:35E4CCA12248D69517094EB2C3A857C6" municipality="Aldabra" name="Gionnet">Gionnet</location>
,
<collectingMunicipality id="52143B85F2C7F39FCFA5100E5A5BAF3C">Aldabra</collectingMunicipality>
,
<date id="C16B78BA2F0ED6F041E49EE8B145543F" value="1974-12-03">
<collectingDate id="BCAFF390CCB819623B1B4F4BB16C5652" value="1974-12-03">03. 12. 1974</collectingDate>
</date>
, leg.
<collectionCode id="6A3E317AE1EB33BE10593F7A7CEEBC9C" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. W. Spaull;
<collectionCode id="EEBAA57E47CB381104BC926EB4C72066" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991466,
<collectorName id="793D7AE705772263C25102886AE8C91A">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="111A342C44B0010AA005E3E351D84CB0" collectedFrom="Casuarina" collectingDate="1974-02-04" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Aldabra" specimenCode="NHMUK 015991466" specimenCount="1" specimenCount-male="1">
<specimenCode id="83C0C59E192D31AEEA53FD4FEFD81C04">NHMUK 015991466</specimenCode>
,
<specimenCount id="3BD88B9EE0E4FD4FE41385EA0F45DE2F" count="1" type="male">1 ♂</specimenCount>
,
<taxonomicName id="196C0931B53807873F47DBAC206DAED9" authority=", Aldabra" authorityName="Aldabra" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="genus">
<emphasis id="102B601F20B9E0855B4551155F9FFCC0" italics="true">Casuarina</emphasis>
, Aldabra
<taxonomicName id="096E7AA68AC1E06CF36DFE1D4969B176" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="genus">
<emphasis id="102B601F20B9E0855B4551155F9FFCC0" italics="true">
<collectedFrom id="C515952F07C9317F77DCDCBAF07DA94D">Casuarina</collectedFrom>
</emphasis>
</taxonomicName>
,
<date id="A96181896B63BDEC7BD00543BC6119F4" value="1974-02-04">04. 02. 1974</date>
<location id="16D88655F1A592C867A88D03CC60B9FA" LSID="urn:lsid:plazi:treatment:8692BA1F9EAF5459A438AB09F9D650CD:16D88655F1A592C867A88D03CC60B9FA" name="Aldabra">Aldabra</location>
,
<date id="A96181896B63BDEC7BD00543BC6119F4" value="1974-02-04">
<collectingDate id="4D3551D41DEBA13D0507CCCF7837E4FC" value="1974-02-04">04. 02. 1974</collectingDate>
</date>
, leg.
<collectionCode id="1835DE506CA5BDCA456B312E2592AC3C" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. W. Spaull.
<collectorName id="CDBD5D5CDF0343D803FA7D8688051FCF">V. W. Spaull</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6ABI" type="remarks">
@ -170,7 +220,9 @@ within its range.
<emphasis id="F842A72489FBF77D1AF43DA55D135123" italics="true">Mecistocephalus lohmanderi</emphasis>
<bibRefCitation id="DD2262579C26BA2A8CB4756AD4781D68" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 72: 71-98." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff, 1939</bibRefCitation>
</taxonomicName>
, NHMUK 015991460, forcipular segment, dorsal view
,
<specimenCode id="5D7573615BAA7E8F205F94B4A9C6946E">NHMUK 015991460</specimenCode>
, forcipular segment, dorsal view
<emphasis id="CF03E74FE047AB514D7FB9F61D1F1707" bold="true">B</emphasis>
<taxonomicName id="86D20B3FFD13456959EBBF77D190E5BF" authority="(Ribaut, 1914)" baseAuthorityName="Ribaut" baseAuthorityYear="1914" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B4D98A7DF4F0C75EC793551D32B30FB1" italics="true">Mecistocephalus angusticeps</emphasis>
@ -178,7 +230,9 @@ within its range.
<bibRefCitation id="A38E3A382958DFC33B63ED07F3B1FE13" author="Ribaut H" refId="B90" refString="Ribaut H (1914) Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale (1911 - 1912). Resultats Scientifiques. Myriapodes. I. Chilopoda. Libr. A. Schulz, Paris 1 - 35." year="1914">Ribaut, 1914</bibRefCitation>
)
</taxonomicName>
, NHMUK 015991466, forcipular segment, dorsal view.
,
<specimenCode id="670F3A4E4BA680275C9D2C6853DDAB3C">NHMUK 015991466</specimenCode>
, forcipular segment, dorsal view.
</paragraph>
</caption>
</subSubSection>

View file

@ -1,48 +1,48 @@
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@ -94,9 +105,7 @@ species
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(
<bibRefCitation id="E734EF8B73265FA9F65608E176804F0F" DOI="10.1080/00222930110098508" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Journal of Natural History" pagination="79-106" refId="B55" refString="Lewis JGE (2002) The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha). Journal of Natural History 36: 79106. https://doi.org/10.1080/00222930110098508" title="The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha)." volume="36" year="2002">Lewis 2002</bibRefCitation>
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locality. In the singular Aldabra specimen examined, the presence of minute accessory spurs on the pretarsus of leg pairs 120 (Fig.
) completed the summary original description and provided additional illustrations of material from near the type locality. In the singular Aldabra specimen examined, the presence of minute accessory spurs on the pretarsus of leg pairs 120 (Fig.
<figureCitation id="C1DE4214965833F7FB2DA42F75FE4A4F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Cryptops mauritianus Verhoeff, 1939. NHMUK 015991434 A anterior margin of forcipular coxosternite, ventral view B calyx of venom gland. Left forcipule (top), right forcipule (bottom) C coxopleuron of ultimate leg-bearing segment, lateral view D Ultimate leg telopodite, lateral view E tibia and tarsal articles of ultimate leg, lateral view F tibial comb (top), tarsal comb of left ultimate leg (middle), tarsal comb of right ultimate leg (bottom) G pretarsus of leg pair 18, lateral view." figureDoi="10.3897/zookeys.1228.143007.figure3" httpUri="https://binary.pensoft.net/fig/1257822">3 G</figureCitation>
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<bibRefCitation id="E4168991606C8612AD49458F0FE28AFE" author="Verhoeff KW" journalOrPublisher="Zoologische Jahrbucher, Abteilung für Systematik, Ökologie und Geographie der Tiere" pagination="71-98" refId="B108" refString="Verhoeff KW (1939) Chilopoden der Insel Mauritius. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere 72: 71-98." title="Chilopoden der Insel Mauritius." volume="72" year="1939">Verhoeff, 1939</bibRefCitation>
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anterior margin of forcipular coxosternite, ventral view
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<taxonomicName id="791AE201ACD4702836C3812212E80009" ID-CoL="ZZ7F" authority="Takakuwa, 1936" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="6DF98C2E0C2CA05426D9EB4F260F9D93" italics="true">Cryptops nigropictus</emphasis>
<bibRefCitation id="1E112F018DEE41F962B242FE6E609F68" author="Takakuwa Y" journalOrPublisher="Transactions of the Sapporo Natural History Society" pagination="236-241" refId="B101" refString="Takakuwa Y (1936) Uber die japanischen Cryptops - Arten. Transactions of the Sapporo Natural History Society 14: 236-241." title="Über die japanischen Cryptops - Arten." volume="14" year="1936">Takakuwa, 1936</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0B0A15FAF8AF8AA120D78C14E36983DE" type="description">
<paragraph id="5810B3EF69484D19D264F9F8BCAA9731">
<figureCitation id="6769F217B3710B6778F22AEDB72DDD4D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cryptops nigropictus Takakuwa, 1936 A, F, K, I, B NHMUK 015991426 H, J NHMUK 015991446 G NHMUK 015991424 C “ Cryptops daszaki ” holotype, Île aux Aigrettes, 19. 10. 1995 D, E paratype, Île aux Aigrettes, 18. 10. 1995 A, B, C, D, E anterior margin of forcipular coxosternite, ventral view F calyx of venom gland, lateral view G, H coxopleuron of ultimate leg-bearing segment, lateral view I, J tibia and tarsus of ultimate leg telopodite, lateral view K pretarsus of leg pair 8, lateral view." figureDoi="10.3897/zookeys.1228.143007.figure4" httpUri="https://binary.pensoft.net/fig/1257823">Fig. 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXIAG" type="material">
<paragraph id="393699E974FBB74575B7DC0500F890B3">
<heading id="C6A19E43248E6D1D38F6C0B461E09F1A" reason="title">Examined material.</heading>
</paragraph>
<paragraph id="4545E68C4FA2156CED951AD9126D4413">
<specimenCount id="55AC539C2CAF75DE02FF443A3284593F" count="12" type="generic">12 specimens</specimenCount>
:
<materialsCitation id="1E4707D8DA0C6F18BB6ED337463814BC" collectionCode="NHMUK" county="Aldabra" location="Black Path" municipality="Picard" specimenCode="NHMUK 015991424" specimenCount="3">
<specimenCode id="2E576C16B52C5D31990FE4F93E88E61E">NHMUK 015991424</specimenCode>
,
<specimenCount id="06ACD07D933E02072DA3E2C5045996CE" count="3" type="generic">3 specimens</specimenCount>
,
<location id="5DB31C3AE36A755AE177C8539913540C" LSID="urn:lsid:plazi:treatment:ADBE62175EF05AEA9D94A0DE96CB9C55:5DB31C3AE36A755AE177C8539913540C" county="Aldabra" municipality="Picard" name="Black Path">Black Path</location>
,
<collectingMunicipality id="F4FB20C27653ACC4803DBD092303BDA6">Picard</collectingMunicipality>
,
<collectingCounty id="E46C40283CAC3C67938A485D76D6CF1F">Aldabra</collectingCounty>
, Summer 1975
</materialsCitation>
;
<materialsCitation id="552373116A01865C17927D00E7CDCF16" collectedFrom="Calophyllum litter" collectingDate="1975-01-14" collectionCode="NHMUK" location="Takamaka" municipality="Grande Terre" specimenCode="NHMUK 015991426" specimenCount="3">
<specimenCode id="A640D5844E35A4BCDBA6E261AC798750">NHMUK 015991426</specimenCode>
,
<specimenCount id="6B9DB00FB12C476C1415DCABC080C512" count="3" type="generic">3 specimens</specimenCount>
,
<collectedFrom id="351A44A4515623FDD9DF04081B8418DD">
<taxonomicName id="5A9D363D22906B535924BE0EA285E3EF" class="Magnoliopsida" family="Calophyllaceae" genus="Calophyllum" kingdom="Plantae" order="Malpighiales" phylum="Tracheophyta" rank="genus">
<emphasis id="72992736FF7FE3BF3985668AA9FB8200" italics="true">Calophyllum</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="BEC3960CC63CC42DC8FCF794F87AC540" LSID="urn:lsid:plazi:treatment:ADBE62175EF05AEA9D94A0DE96CB9C55:BEC3960CC63CC42DC8FCF794F87AC540" municipality="Grande Terre" name="Takamaka">Takamaka</location>
,
<collectingMunicipality id="126AF5F0B880534928525032A83A9C5D">Grande Terre</collectingMunicipality>
,
<date id="881E095945694554AA20A58EC83AE6A1" value="1975-01-14">
<collectingDate id="E8AC4D4EB3360D1F6E47EEA9F6E9DCF6" value="1975-01-14">14. 01. 1975</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="282D103B86C81F8404D899C30F83ACE6" collectedFrom="Sideroxylon litter" collectingDate="1975-02-14" collectionCode="NHMUK" collectorName="V. W. Spaull" location="Au Parc" municipality="Aldabra" specimenCode="NHMUK 015991427" specimenCount="2">
<specimenCode id="A6020A58B056EB1FF59F9EF62933357E">NHMUK 015991427</specimenCode>
,
<specimenCount id="B0F591979640CDA339D945CE969FD425" count="2" type="generic">2 specimens</specimenCount>
,
<collectedFrom id="67EA5AEE1A0B7E2E7B2213112A937B9C">
<taxonomicName id="5B8828C1BA9153205E4A0C9D71F07C02" class="Magnoliopsida" family="Sapotaceae" genus="Sideroxylon" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="genus">
<emphasis id="9CC56CC585DA84794147A89E980045B9" italics="true">Sideroxylon</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="69BA02920496AC1F772C9248E8938426" LSID="urn:lsid:plazi:treatment:ADBE62175EF05AEA9D94A0DE96CB9C55:69BA02920496AC1F772C9248E8938426" municipality="Aldabra" name="Au Parc">Au Parc</location>
,
<collectingMunicipality id="0B3458863A0C13B209BA0DA08341E4ED">Aldabra</collectingMunicipality>
,
<date id="1D76A0A13EF48F8432AA66A4FC41EB86" value="1975-02-14">
<collectingDate id="A2EDD94E81C476B160E9FE8E3A3D9CB4" value="1975-02-14">14. 02. 1975</collectingDate>
</date>
.
<collectorName id="AE073966EFE6CB4C8F2F14435ABF07F0">V. W. Spaull</collectorName>
leg
</materialsCitation>
;
<materialsCitation id="ADED9F39F48AAFF09B610A9AA9B694A6" collectingDate="1975-02-28" collectionCode="NHMUK" location="Picard" municipality="Aldabra" specimenCode="NHMUK 015991428" specimenCount="1">
<specimenCode id="B627E93127F9559AC16EE22A58018D36">NHMUK 015991428</specimenCode>
,
<specimenCount id="06F3133847CAB1E273A662690FC0BEEF" count="1" type="generic">1 specimen</specimenCount>
,
<location id="F5E485B26E481A6CFB9BABF1DE3DD6CA" LSID="urn:lsid:plazi:treatment:ADBE62175EF05AEA9D94A0DE96CB9C55:F5E485B26E481A6CFB9BABF1DE3DD6CA" municipality="Aldabra" name="Picard">Picard</location>
,
<collectingMunicipality id="71A688867198DD72D4E8810324EFFCB6">Aldabra</collectingMunicipality>
,
<date id="DA937CFDBB2A132ADD8B0F4B87AB9E51" value="1975-02-28">
<collectingDate id="D608B4636E613B1FA32F6DAACE9BF1C2" value="1975-02-28">28. 02. 1975</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="61183DF8E490ED7C09EEC68C40797EEC" collectedFrom="Sideroxylon litter" collectingDate="1975-03-28" collectionCode="NHMUK" location="Ile. Michel" municipality="Aldabra" specimenCode="NHMUK 015991445" specimenCount="1">
<specimenCode id="C599BE56F85CEC38D052EE7A6A0DF969">NHMUK 015991445</specimenCode>
,
<specimenCount id="59E195FA03D4308657887AEFA8F364AD" count="1" type="generic">1 specimen</specimenCount>
,
<collectedFrom id="33D3FEDB58FA34A8BAAAD78FF29B9846">
<taxonomicName id="7484CB90FDFF76B842AAD310C5AB8B40" class="Magnoliopsida" family="Sapotaceae" genus="Sideroxylon" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="genus">
<emphasis id="41FC3F6C5BE63A1AAE576A839EE04883" italics="true">Sideroxylon</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="5D50D2BA31856789868A0417D56979DA" LSID="urn:lsid:plazi:treatment:ADBE62175EF05AEA9D94A0DE96CB9C55:5D50D2BA31856789868A0417D56979DA" municipality="Aldabra" name="Ile. Michel">Ile. Michel</location>
,
<collectingMunicipality id="DA44A7658F9633DD9B7C21178B55DCB7">Aldabra</collectingMunicipality>
,
<date id="AF2BBBB5FC9AE45D7874CCF395137DFC" value="1975-03-28">
<collectingDate id="D8E40A84D7C0B57C4F76B08982EF342A" value="1975-03-28">28. 03. 1975</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="EF40082F7E8731B872ED5FDA5A504B03" collectedFrom="Casuarina litter" collectingDate="1974-05-08" collectionCode="NHMUK" location="Picard" municipality="Aldabra" specimenCode="NHMUK 015991446" specimenCount="2">
<specimenCode id="CA9758376F71B07D52846211B82A41B1">NHMUK 015991446</specimenCode>
,
<specimenCount id="CAD90764DB03D862DC6D5401EAC809A3" count="2" type="generic">2 specimens</specimenCount>
,
<collectedFrom id="CE88D028BE901ED22155A8650FC7D66E">
<taxonomicName id="6623C012455A6A6AF53CB1E7641F81A3" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="genus">
<emphasis id="C3F7EC173B5DB4E6A776259293B0046D" italics="true">Casuarina</emphasis>
</taxonomicName>
litter
</collectedFrom>
,
<location id="6C967DE1BEC530EF5D4DB0BBC8DE7CB9" LSID="urn:lsid:plazi:treatment:ADBE62175EF05AEA9D94A0DE96CB9C55:6C967DE1BEC530EF5D4DB0BBC8DE7CB9" municipality="Aldabra" name="Picard">Picard</location>
,
<collectingMunicipality id="54A0F7BB45B89C485170AD16DB512F5B">Aldabra</collectingMunicipality>
,
<date id="8EC9269BB491646F66DD8EB903DFFE3F" value="1974-05-08">
<collectingDate id="F453E92CE1BF5C5DC3BF11460E7A5ABE" value="1974-05-08">08. 05. 1974</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZJAG" type="remarks">
<paragraph id="44308315097D7EF6A73FF64BD52D9837">
<heading id="9D0053139D5748DE32521A76F6D1EB61" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="A751EABC61DB66DFCED9FC978141F55C">
Taxonomic revision of
<taxonomicName id="EFDDA10D648EB2381C68CA399B098343">
<emphasis id="E8F33342AC1ECCBAAC4B119B742512C5" italics="true">Cryptops</emphasis>
species
</taxonomicName>
belonging to the
<taxonomicName id="38C7E7EBC74819E5CF69082F81C7078E" authorityName="Donovan" authorityYear="1810" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="hortensis">
<emphasis id="148527A0E40DDB7885B736115450CE6B" italics="true">hortensis</emphasis>
</taxonomicName>
group ” identified
<taxonomicName id="3769CB148F32AE26189B03BBF6B5D69C" authority="Lawrence, 1960" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="decoratus">
<emphasis id="5DE32E8188F85D5EAF7AD669942AC9F1" italics="true">C. decoratus</emphasis>
<bibRefCitation id="719D87E688D7419E5E126EFD71C84D88" author="Lawrence RF" journalOrPublisher="Faune de Madagascar" pagination="1-123" refId="B46" refString="Lawrence RF (1960) Myriapodes - Chilopodes. Faune de Madagascar 12: 1-123." title="Myriapodes - Chilopodes." volume="12" year="1960">Lawrence, 1960</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="885B888F1E1D863107F5B64091C3D5E5" authority="Chamberlin, 1941" authorityName="Chamberlin" authorityYear="1941" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="melanotypus">
<emphasis id="DE2592647CD200517AF935E4B484B48A" italics="true">C. melanotypus</emphasis>
Chamberlin, 1941
</taxonomicName>
, and
<taxonomicName id="DBBD481D20534916EF4EC7EDE3B5817D" authority="Takakuwa, 1936" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="CBE255B413757D8E553C4572F01E0C55" italics="true">C. nigropictus</emphasis>
<bibRefCitation id="3E714107B5DB216B285D7A36ED3BDEF2" author="Takakuwa Y" journalOrPublisher="Transactions of the Sapporo Natural History Society" pagination="236-241" refId="B101" refString="Takakuwa Y (1936) Uber die japanischen Cryptops - Arten. Transactions of the Sapporo Natural History Society 14: 236-241." title="Über die japanischen Cryptops - Arten." volume="14" year="1936">Takakuwa, 1936</bibRefCitation>
</taxonomicName>
as a potential species complex, raising doubts on the taxonomic validity of many morphologically similar species (
<bibRefCitation id="1EA692F9D3999CEB7CA252D11A63FB03" DOI="10.3897/ijm.4.1116" author="Lewis" firstAuthor="Lewis" journalOrPublisher="International Journal of Myriapodology" pagination="11-50" refId="B61" refString="Lewis JGE (2011) A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha). International Journal of Myriapodology 4: 1150. https://doi.org/10.3897/ijm.4.1116" title="A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha)." volume="4" year="2011">Lewis 2011</bibRefCitation>
). A reliable diagnostic trait mentioned by
<bibRefCitation id="408FFC5A6BE6D8C07D01CD45C86B7DC7" DOI="10.3897/ijm.4.1116" author="Lewis JGE" firstAuthor="Lewis" journalOrPublisher="International Journal of Myriapodology" pagination="11-50" refId="B61" refString="Lewis JGE (2011) A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha). International Journal of Myriapodology 4: 1150. https://doi.org/10.3897/ijm.4.1116" title="A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha)." volume="4" year="2011">Lewis (2011)</bibRefCitation>
is the presence of a single long pretarsal accessory spur in
<taxonomicName id="A40F47E56801249821D4CDD83BBCE352" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="F9E91D420498EFF3F02DCD1E528D2C15" italics="true">C. nigropictus</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="FCAC40238C3978CC34F65812401BE759" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cryptops nigropictus Takakuwa, 1936 A, F, K, I, B NHMUK 015991426 H, J NHMUK 015991446 G NHMUK 015991424 C “ Cryptops daszaki ” holotype, Île aux Aigrettes, 19. 10. 1995 D, E paratype, Île aux Aigrettes, 18. 10. 1995 A, B, C, D, E anterior margin of forcipular coxosternite, ventral view F calyx of venom gland, lateral view G, H coxopleuron of ultimate leg-bearing segment, lateral view I, J tibia and tarsus of ultimate leg telopodite, lateral view K pretarsus of leg pair 8, lateral view." figureDoi="10.3897/zookeys.1228.143007.figure4" httpUri="https://binary.pensoft.net/fig/1257823">4 K</figureCitation>
), differentiating it from
<taxonomicName id="C2F539125DD106FDEE965821EA95C772" authorityName="Lawrence" authorityYear="1960" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="decoratus">
<emphasis id="72230ECE05EEC6DC2F1BAABB95605993" italics="true">C. decoratus</emphasis>
</taxonomicName>
and
<taxonomicName id="D957257F69A947CC0374E1569D995D84" authorityName="Chamberlin" authorityYear="1941" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="melanotypus">
<emphasis id="F2B2813EE77E74E816914F35F0F07235" italics="true">C. melanotypus</emphasis>
</taxonomicName>
which bear two small pretarsal accessory spurs on leg pairs 120. The Aldabra specimens key out to
<taxonomicName id="3220462142D5FB90549F27D9C8F975DF" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="F8C6F60514C511E130B5B370C10122A7" italics="true">C. nigropictus</emphasis>
</taxonomicName>
in the key provided within the same article and agree with its revised description. Although originally described from
<collectingCountry id="6FED8574CAFAB972DF8C2204F235869F" name="China">East Asia</collectingCountry>
, various nominal taxa recorded from East Africa and islands of the Indian Ocean (including
<collectingCountry id="D85BA6DE84012D235B9C3645FF010B82" name="Mauritius">Mauritius</collectingCountry>
and
<collectingRegion id="A9CB9850E466E34345E3194C28577B74" country="Mauritius" name="Rodrigues">Rodrigues</collectingRegion>
) have been placed in synonymy with
<taxonomicName id="01B082DB31AEDEDCDF484AFF8A5C7615" authority="(Lewis 2011)" baseAuthorityName="Lewis" baseAuthorityYear="2011" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="E4478E1D108FE1E212E411A31ECCC26D" italics="true">C. nigropictus</emphasis>
(
<bibRefCitation id="0B6D3A68903AA8B57A71CEE7D60F13BA" DOI="10.3897/ijm.4.1116" author="Lewis" firstAuthor="Lewis" journalOrPublisher="International Journal of Myriapodology" pagination="11-50" refId="B61" refString="Lewis JGE (2011) A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha). International Journal of Myriapodology 4: 1150. https://doi.org/10.3897/ijm.4.1116" title="A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha)." volume="4" year="2011">Lewis 2011</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="39C052DFF9E49F41C004AF0B558BB2E7" ID-DOI="10.3897/zookeys.1228.143007.figure4" ID-arpha="0C5E785E-4D27-544B-8FE3-AF89980C0F1C" httpUri="https://binary.pensoft.net/fig/1257823" startId="F4">
<paragraph id="AE367D50CF7CB20B8D8FD7062597AA91">
<label id="5E6DCFD6DBF9A98792ECD1AEA5E16EF0">Figure 4.</label>
</paragraph>
<paragraph id="6D0D88C3214B5F4DDBF7040D20EEE8BB">
<taxonomicName id="7E1E45E5DB55B42D181DB228E54EAEBD" authority="Takakuwa, 1936" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="6CDDAE9B3AE115B68A9681ED8B50AA6C" italics="true">Cryptops nigropictus</emphasis>
<bibRefCitation id="7E5566700C008F4355DDB353B5BB76F7" author="Takakuwa Y" journalOrPublisher="Transactions of the Sapporo Natural History Society" pagination="236-241" refId="B101" refString="Takakuwa Y (1936) Uber die japanischen Cryptops - Arten. Transactions of the Sapporo Natural History Society 14: 236-241." title="Über die japanischen Cryptops - Arten." volume="14" year="1936">Takakuwa, 1936</bibRefCitation>
</taxonomicName>
<emphasis id="F30871B0AEC36B6F2F5A4CDD50B0E731" bold="true">A, F, K, I, B</emphasis>
<specimenCode id="8D72A27A56AFD6717758B42F2806B10B">NHMUK 015991426</specimenCode>
<emphasis id="262E5AC0527E05D29694F94A75D821F9" bold="true">H, J</emphasis>
<specimenCode id="5387D0B19598F7D98692D07AD43CB9B8">NHMUK 015991446</specimenCode>
<emphasis id="A1F8E106EA0D67257AC26DEAE219CF98" bold="true">G</emphasis>
<specimenCode id="7A0F0FAD87780CE0459AB36F291A6670">NHMUK 015991424</specimenCode>
<emphasis id="080621F98B4CD6820679CB9E9E32A81B" bold="true">C</emphasis>
<taxonomicName id="B51D1DDE91F3326245E23B3DDF4FB70E" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="FE3851059F0E3BC6B960D3B0D3C6A5CA" italics="true">Cryptops daszaki</emphasis>
</taxonomicName>
” holotype, Île aux Aigrettes, 19. 10. 1995
<emphasis id="B0034FFAFF45B360C3DFD07BD408E767" bold="true">D, E</emphasis>
paratype, Île aux Aigrettes, 18. 10. 1995
<emphasis id="49041883446AC69E4260E9642112C5E4" bold="true">A, B, C, D, E</emphasis>
anterior margin of forcipular coxosternite, ventral view
<emphasis id="07DD818C7B6E53682980C0F2E76E8AA2" bold="true">F</emphasis>
calyx of venom gland, lateral view
<emphasis id="56ACEC1209833E09F48EC6DB66FC587F" bold="true">G, H</emphasis>
coxopleuron of ultimate leg-bearing segment, lateral view
<emphasis id="D3F678E5B5C13787334A34F8C5CAE630" bold="true">I, J</emphasis>
tibia and tarsus of ultimate leg telopodite, lateral view
<emphasis id="EB945061D2E937CA1A4F83778E10E3BC" bold="true">K</emphasis>
pretarsus of leg pair 8, lateral view.
</paragraph>
</caption>
<paragraph id="E28A28350763B95156D2F8BDACC0AEF1">
<taxonomicName id="B45021372A9328CA1377AEF01F03C285" authority="Lewis, 2002" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="46B25158681CF8ACEEC7EE1B930B297B" italics="true">Cryptops daszaki</emphasis>
<bibRefCitation id="BFD302CD7D4C3B565186C6A204C05947" DOI="10.1080/00222930110098508" author="Lewis JGE" journalOrPublisher="Journal of Natural History" pagination="79-106" refId="B55" refString="Lewis JGE (2002) The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha). Journal of Natural History 36: 79-106. https://doi.org/10.1080/00222930110098508" title="The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha)." volume="36" year="2002">Lewis, 2002</bibRefCitation>
</taxonomicName>
was described from several localities in
<collectingCountry id="AE3AF1F6B1FE1A94043241E0F0D7BA89" name="Mauritius">Mauritius</collectingCountry>
.
<bibRefCitation id="1E6808CD90AFF865AB9FA1503DD174F5" DOI="10.1080/00222930110098508" author="Lewis JGE" firstAuthor="Lewis" journalOrPublisher="Journal of Natural History" pagination="79-106" refId="B55" refString="Lewis JGE (2002) The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha). Journal of Natural History 36: 79106. https://doi.org/10.1080/00222930110098508" title="The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha)." volume="36" year="2002">Lewis (2002)</bibRefCitation>
remarked on the small body size (
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) of sexually mature specimens and their apparent juvenile characteristics (low number of coxal pores, reduced setation). The specimens collected from Aldabra with a similar body length to specimens assigned by Lewis to
<taxonomicName id="39B4042F3AA6EF960D8734349B4FA2CE" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="979C40F74938B3A04ACB6EC9871A0944" italics="true">C. daszaki</emphasis>
</taxonomicName>
overlap in morphology with respect to the usual lack of subcuticular dark pigment, the number of setae on the anterior margin of the forcipular coxosternite (2 or 3 on each side in
<taxonomicName id="D74A82A227929141237B9DC0BA86A352" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="D14053DEFDF91703B6ABE3901296F326" italics="true">C. daszaki</emphasis>
</taxonomicName>
, 3 or 4 on each side in
<taxonomicName id="AA5CAED69AB153CC84424992F0C62171" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="DA3C03809477BBD158271583670DFE5E" italics="true">C. nigropictus</emphasis>
</taxonomicName>
; Fig.
<figureCitation id="A9643BB0625D50878BC4033AC28812EA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cryptops nigropictus Takakuwa, 1936 A, F, K, I, B NHMUK 015991426 H, J NHMUK 015991446 G NHMUK 015991424 C “ Cryptops daszaki ” holotype, Île aux Aigrettes, 19. 10. 1995 D, E paratype, Île aux Aigrettes, 18. 10. 1995 A, B, C, D, E anterior margin of forcipular coxosternite, ventral view F calyx of venom gland, lateral view G, H coxopleuron of ultimate leg-bearing segment, lateral view I, J tibia and tarsus of ultimate leg telopodite, lateral view K pretarsus of leg pair 8, lateral view." figureDoi="10.3897/zookeys.1228.143007.figure4" httpUri="https://binary.pensoft.net/fig/1257823">4 C</figureCitation>
versus Fig.
<figureCitation id="4386A29006E60F0388E2074BC20CD2DC" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cryptops nigropictus Takakuwa, 1936 A, F, K, I, B NHMUK 015991426 H, J NHMUK 015991446 G NHMUK 015991424 C “ Cryptops daszaki ” holotype, Île aux Aigrettes, 19. 10. 1995 D, E paratype, Île aux Aigrettes, 18. 10. 1995 A, B, C, D, E anterior margin of forcipular coxosternite, ventral view F calyx of venom gland, lateral view G, H coxopleuron of ultimate leg-bearing segment, lateral view I, J tibia and tarsus of ultimate leg telopodite, lateral view K pretarsus of leg pair 8, lateral view." figureDoi="10.3897/zookeys.1228.143007.figure4" httpUri="https://binary.pensoft.net/fig/1257823">4 A, B, D</figureCitation>
, respectively), the relative length of the single pretarsal accessory spur on legs 120 (&gt; 1 / 2 the length of the pretarsus) and the number of coxal pores (5 or
<quantity id="5E1771152CDC9E28AA3C4BC016FA1F34" metricMagnitude="-1" metricUnit="m" metricValue="1.524" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="5E9A92A666086EB0C5722C17ACAE87A3" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="DE182F14F66FE63EA6F397149D29A0AC" italics="true">C. daszaki</emphasis>
</taxonomicName>
,
<quantity id="CE40E70B962E994172C894AADD44B9D8" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.286" metricValueMin="1.016" unit="in" value="6.5" valueMax="9.0" valueMin="4.0">49 in</quantity>
<taxonomicName id="3B830F63D7AD4BC22EF74E02EEC44D63" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="9796647C8DF69801AB7D3D34D769ABB3" italics="true">C. nigropictus</emphasis>
</taxonomicName>
) but show greater variation than
<taxonomicName id="95FC0143704D5F3912AB6AE759062565" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="C2323C9000937F09A8F4E2D211CD5660" italics="true">C. daszaki</emphasis>
</taxonomicName>
in the number of saw teeth on the tibia (35) and tarsus (2 or 3). The number of tibial and tarsal saw teeth is known to be intraspecifically variable in
<taxonomicName id="74137FA2CD56C6B8B071EB1ABD70874B" authorityName="Leach" authorityYear="1815" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="genus">
<emphasis id="DFC5858B869032A5B047BA944E0028DE" italics="true">Cryptops</emphasis>
</taxonomicName>
(
<bibRefCitation id="939B978BE3097D9243F85E5DC772BC58" author="Iorio and Geoffroy" firstAuthor="Iorio" refId="B39" refString="Iorio E, Geoffroy J-J (2003) Etude des Scolopendromorphes français du genre Cryptops Leach 1814 Première partie: Cryptops parisi Brolemann, 1920 et C. parisi var. cristata Ribaut, 1925 (Chilopoda, Scolopendromorpha, Cryptopidae). Bulletin de Phyllie n ° 184 e trimestre: 2837." year="2003">Iorio and Geoffroy 2003</bibRefCitation>
;
<bibRefCitation id="AA505DB1F06F690D49B7944B4205474D" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Soil Organisms" pagination="505-518" refId="B58" refString="Lewis JGE (2009) A review of some characters used in the taxonomy of Cryptops (subgenus Cryptops) (Chilopoda: Scolopendromorpha: Cryptopidae). Soil Organisms 81: 505518." title="A review of some characters used in the taxonomy of Cryptops (subgenus Cryptops) (Chilopoda: Scolopendromorpha: Cryptopidae)." volume="81" year="2009">Lewis 2009</bibRefCitation>
) and scales allometrically with body size, as exemplified by the largest
<taxonomicName id="663DDB050C54469CDC37B4F10F3DFD0F" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="01C7FBCDD27F524E06DCCCB4A1D05A83" italics="true">C. nigropictus</emphasis>
</taxonomicName>
specimen in our sample (
<quantity id="45C0726AC3FF9AFC15BD8BAF2CAACC1A" metricMagnitude="-2" metricUnit="m" metricValue="1.2" unit="mm" value="12.0">12 mm</quantity>
,
<specimenCode id="00EF2400836FBBC18B165E78FB71A108">
<collectionCode id="128BEA55DC1286C772F9489E6CB72EFD" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991426
</specimenCode>
; Fig.
<figureCitation id="57EDB5E3502994D364D79D2CFB35745D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cryptops nigropictus Takakuwa, 1936 A, F, K, I, B NHMUK 015991426 H, J NHMUK 015991446 G NHMUK 015991424 C “ Cryptops daszaki ” holotype, Île aux Aigrettes, 19. 10. 1995 D, E paratype, Île aux Aigrettes, 18. 10. 1995 A, B, C, D, E anterior margin of forcipular coxosternite, ventral view F calyx of venom gland, lateral view G, H coxopleuron of ultimate leg-bearing segment, lateral view I, J tibia and tarsus of ultimate leg telopodite, lateral view K pretarsus of leg pair 8, lateral view." figureDoi="10.3897/zookeys.1228.143007.figure4" httpUri="https://binary.pensoft.net/fig/1257823">4 I</figureCitation>
).
</paragraph>
<paragraph id="2AED7731E8815D4A8F586AB74819F36F">
The only other diagnostic trait given by
<bibRefCitation id="DB679D5C8DA784D027D0BF5B61A84466" DOI="10.3897/ijm.4.1116" author="Lewis JGE" firstAuthor="Lewis" journalOrPublisher="International Journal of Myriapodology" pagination="11-50" refId="B61" refString="Lewis JGE (2011) A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha). International Journal of Myriapodology 4: 1150. https://doi.org/10.3897/ijm.4.1116" title="A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha)." volume="4" year="2011">Lewis (2011)</bibRefCitation>
separating
<taxonomicName id="6661F0BC0EADF5007019074E861A4913" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="77F4BDBFD909A77F29B502D852CC719E" italics="true">C. daszaki</emphasis>
</taxonomicName>
and
<taxonomicName id="F91844FFF3C14E9FDD5F6AA260D23664" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="BEDAD18559A1BEFDBF5B8BDB103AD7A6" italics="true">C. nigropictus</emphasis>
</taxonomicName>
is the position of the setae near the anterior margin of the forcipular coxosternite, being described as “ marginal ” in
<taxonomicName id="6EEBF93BC8F0F7E3EB312082635C5648" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="049BB08B282B853CD2CEB5DABDC70C2E" italics="true">C. daszaki</emphasis>
</taxonomicName>
and “ submarginal ” in
<taxonomicName id="144A71002CEC02A1C765311BEC72A162" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="B3D32C454DE8F5FBC1329E73437C6DBE" italics="true">C. nigropictus</emphasis>
</taxonomicName>
. Re-examination of the type specimens of
<taxonomicName id="F715969219469DB02D5DCB2B15B4580E" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="ADA9BDC7D00AF1E97D07E44665222EB8" italics="true">C. daszaki</emphasis>
</taxonomicName>
revealed that these setae occupy a submarginal position (Fig.
<figureCitation id="DAF6BCC6B514AF1D6381DC1091F19968" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Cryptops nigropictus Takakuwa, 1936 A, F, K, I, B NHMUK 015991426 H, J NHMUK 015991446 G NHMUK 015991424 C “ Cryptops daszaki ” holotype, Île aux Aigrettes, 19. 10. 1995 D, E paratype, Île aux Aigrettes, 18. 10. 1995 A, B, C, D, E anterior margin of forcipular coxosternite, ventral view F calyx of venom gland, lateral view G, H coxopleuron of ultimate leg-bearing segment, lateral view I, J tibia and tarsus of ultimate leg telopodite, lateral view K pretarsus of leg pair 8, lateral view." figureDoi="10.3897/zookeys.1228.143007.figure4" httpUri="https://binary.pensoft.net/fig/1257823">4 C</figureCitation>
), which may be misinterpreted as marginal depending on the orientation of the specimens. As both of the two putative diagnostic traits separating
<taxonomicName id="13B71FB6E690BA42DAEA83C6C0A0FFAF" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="F292534831736B08006E449367F46257" italics="true">C. daszaki</emphasis>
</taxonomicName>
and
<taxonomicName id="109E10114283E60350A04E20CE7491E7" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="4FE2D4C82488E75183BCC7CB024BEFCB" italics="true">C. nigropictus</emphasis>
</taxonomicName>
are fully encompassed by intraspecific variation within
<taxonomicName id="27A2B5DD635409CCD64928EDACC2407B" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="746FABCB3041916A5DE094B12B1CB69B" italics="true">C. nigropictus</emphasis>
</taxonomicName>
and the latter species has been recorded from nearby localities (
<bibRefCitation id="36B0E8CA1FAC03699D7433CB11434150" DOI="10.1080/00222930110098508" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Journal of Natural History" pagination="79-106" refId="B55" refString="Lewis JGE (2002) The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha). Journal of Natural History 36: 79106. https://doi.org/10.1080/00222930110098508" title="The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha)." volume="36" year="2002">Lewis 2002</bibRefCitation>
), we consider
<taxonomicName id="EA30C5B4DA6220C6563615DDB70925ED" authorityName="Lewis" authorityYear="2002" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="daszaki">
<emphasis id="880C0639ED825E0C221F28C84B61AA57" italics="true">C. daszaki</emphasis>
</taxonomicName>
a likely junior subjective synonym of
<taxonomicName id="A492EA091FCB2DF8C758B78D44789605" authorityName="Takakuwa" authorityYear="1936" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="80318572CF203B7599CC03208CD35D63" italics="true">C. nigropictus</emphasis>
</taxonomicName>
, from which it cannot be reliably distinguished by morphology alone. Similarly, the presence of subcuticular dark pigment cannot be reliably used to separate putative
<taxonomicName id="E14D20F21C88C55368162A945B626B51">
<emphasis id="BFB3B80C51C7038C45F3E775AA543EE7" italics="true">Cryptops</emphasis>
species
</taxonomicName>
in Western Indian Ocean and Eastern African localities. This character is variable within our sample, with all specimens with the exception of
<specimenCode id="0A2B51BCA78760650DD7E1A65EA60301">
<collectionCode id="FB957D2EB56F20B8F6BFADC6CC4EE7E5" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991428
</specimenCode>
lacking dark subcuticular pigment. Dark pigmentation is also variable in the type specimens of
<taxonomicName id="215C4D1919137BB64EAA736EC9D48BA8" authority="Lewis, 1967" authorityName="Lewis" authorityYear="1967" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="A5C0CB1D3F5B769FD7A836E68DFA5D28" italics="true">Cryptops niloticus</emphasis>
<bibRefCitation id="9C389F31F048DC3CDD224D03F351CC42" DOI="10.1111/j.1095-8312.1967.tb00973.x" author="Lewis JGE" journalOrPublisher="Proceedings of the Linnean Society of London" pagination="185-207" refId="B51" refString="Lewis JGE (1967) The scolopendromorph centipedes of the Sudan with remarks on taxonomic characters in the Scolopendridae. Proceedings of the Linnean Society of London 178: 185-207. https://doi.org/10.1111/j.1095-8312.1967.tb00973.x" title="The scolopendromorph centipedes of the Sudan with remarks on taxonomic characters in the Scolopendridae." volume="178" year="1967">Lewis, 1967</bibRefCitation>
</taxonomicName>
(present in male
<typeStatus id="8DE72FE6D939BA19E10A9C6E9592DFA6" type="allotype">allotype</typeStatus>
but absent in female
<typeStatus id="1AFDCF5897186596D0DB34A17EFE1CEB" type="holotype">holotype</typeStatus>
and in juvenile
<typeStatus id="EBCBEFA5573179F2A7690D5FF1CF50DA" type="paratype">paratype</typeStatus>
), a species previously recorded from the Western Indian Ocean (
<bibRefCitation id="AAA57202BDDF7720EB5C8F3FF9F3262F" DOI="10.1080/00222930110098508" author="Lewis" firstAuthor="Lewis" journalOrPublisher="Journal of Natural History" pagination="79-106" refId="B55" refString="Lewis JGE (2002) The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha). Journal of Natural History 36: 79106. https://doi.org/10.1080/00222930110098508" title="The scolopendromorph centipedes of Mauritius and Rodrigues and their adjacent islets (Chilopoda: Scolopendromorpha)." volume="36" year="2002">Lewis 2002</bibRefCitation>
) but subsequently synonymised with
<taxonomicName id="E2E3A7123E4EF0ED93DDCE5B50E3A065" authority="(Lewis 2011)" baseAuthorityName="Lewis" baseAuthorityYear="2011" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="nigropictus">
<emphasis id="27243C524A829A10EF6E33A554F38F26" italics="true">C. nigropictus</emphasis>
(
<bibRefCitation id="38F3138B5E421297794AD18AE005B935" DOI="10.3897/ijm.4.1116" author="Lewis" firstAuthor="Lewis" journalOrPublisher="International Journal of Myriapodology" pagination="11-50" refId="B61" refString="Lewis JGE (2011) A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha). International Journal of Myriapodology 4: 1150. https://doi.org/10.3897/ijm.4.1116" title="A review of the species in the genus Cryptops Leach, 1815 from the Old World related to Cryptops (Cryptops) hortensis (Donovan, 1810) (Chilopoda, Scolopendromorpha)." volume="4" year="2011">Lewis 2011</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,48 +1,48 @@
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<mods:title id="C864616813FBB8DBFB5F25BCBCAC24AC">Centipedes (Myriapoda, Chilopoda) of Aldabra Atoll (Seychelles)</mods:title>
<document id="1FF0D830C8C5C993D300C3B54DA8FA7D" ID-DOI="10.3897/zookeys.1228.143007" ID-ZooBank="3A5C895C-ACF6-4E44-BD53-6296E1F9EAB1" ID-publisher-id="143007" URI-arpha="C30A29E7-DD94-5DA1-B8D1-F45BB72750DE" URI-zoobank="http://zoobank.org/3A5C895C-ACF6-4E44-BD53-6296E1F9EAB1" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1740162329141" checkinUser="pensoft" docAuthor="Popovici, George &amp; Edgecombe, Gregory D." docDate="2025" docId="DC1DCBCACB155101A2370EB887D8E379" docLanguage="en" docName="ZooKeys 1228: 225-273" docOrigin="ZooKeys 1228" docSource="https://zookeys.pensoft.net/article/143007/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Cryptops japonicus Takakuwa 1934" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="273" masterDocId="C30A29E7DD945DA1B8D1F45BB72750DE" masterDocTitle="Centipedes (Myriapoda, Chilopoda) of Aldabra Atoll (Seychelles)" masterLastPageNumber="273" masterPageNumber="225" pageNumber="225" updateTime="1740173442479" updateUser="admin">
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<mods:title id="658FD9B84BE159FBF442844049DAA4BA">Centipedes (Myriapoda, Chilopoda) of Aldabra Atoll (Seychelles)</mods:title>
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<mods:namePart id="70188B204677A5F66AFE324FC58B0D92">Popovici, George</mods:namePart>
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<mods:affiliation id="FBC59AFC14CD44C4BFB02377E0BC142C">Department of Life Sciences, Imperial College London, London SW 7 2 AZ, UK &amp; &quot; Grigore Antipa &quot; National Museum of Natural History, Pavel Dimitrievici Kiseleff St. 1, 011341, Bucharest, Romania</mods:affiliation>
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<treatment id="DC1DCBCACB155101A2370EB887D8E379" ID-DOI="http://doi.org/10.5281/zenodo.14907689" ID-Zenodo-Dep="14907689" ID-arpha="DC1DCBCA-CB15-5101-A237-0EB887D8E379" LSID="urn:lsid:plazi:treatment:DC1DCBCACB155101A2370EB887D8E379" httpUri="http://treatment.plazi.org/id/DC1DCBCACB155101A2370EB887D8E379">
<treatment id="DC1DCBCACB155101A2370EB887D8E379" ID-DOI="http://doi.org/10.5281/zenodo.14907689" ID-Zenodo-Dep="14907689" ID-arpha="DC1DCBCA-CB15-5101-A237-0EB887D8E379" LSID="urn:lsid:plazi:treatment:DC1DCBCACB155101A2370EB887D8E379" httpUri="http://treatment.plazi.org/id/DC1DCBCACB155101A2370EB887D8E379" scope_class="Chilopoda" scope_family="Cryptopidae" scope_order="Scolopendromorpha">
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<paragraph id="D37BF1B7D0045F50BA9D54DF4DAA8F61">
<taxonomicName id="822EFC591D16C1E974F6124FCF3C2BD6" ID-CoL="6BLXC" authority="Takakuwa, 1934" authorityName="Takakuwa" authorityYear="1934" class="Chilopoda" family="Cryptopidae" genus="Cryptops" isUncertain="true" kingdom="Animalia" order="Scolopendromorpha" phylum="Arthropoda" rank="species" species="japonicus">
@ -65,34 +65,62 @@ cf.
<paragraph id="07A09430B271B3EC65CAA543F50CEAF9">
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:
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015991425,
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<specimenCode id="32D37E9A41E2806A9508F856C0AA3F65">NHMUK 015991425</specimenCode>
,
<specimenCount id="BB2DDD67162D0E1630D802E18EEAEF7C" count="4" type="generic">4 specimens</specimenCount>
, Black Path, Picard, Summer 1975, leg.
<collectionCode id="704D31574509D5B7230A80E44DE49F97" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
. W. Spaull;
<collectionCode id="18AB70FE85555FE57F8B2389424E9F70" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991429,
,
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,
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, Summer 1975, leg.
<collectorName id="D05701A79CBEB340F87E927C069CDE4E">V. W. Spaull</collectorName>
</materialsCitation>
;
<materialsCitation id="B0F9278518F741730270CFA3227C7F71" collectingDate="1973-11-10" collectionCode="NHMUK" location="Aldabra" specimenCode="NHMUK 015991429" specimenCount="1">
<specimenCode id="D02CD0954C2B24CD0AFD78097399ADCB">NHMUK 015991429</specimenCode>
,
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, Aldabra,
<date id="70E31A703A7A18FEB5A043794AE28940" value="1973-11-10">10. 11. 1973</date>
,
<location id="97D4E676B5D5EBD4D851A45F1BC711F0" LSID="urn:lsid:plazi:treatment:DC1DCBCACB155101A2370EB887D8E379:97D4E676B5D5EBD4D851A45F1BC711F0" name="Aldabra">Aldabra</location>
,
<date id="70E31A703A7A18FEB5A043794AE28940" value="1973-11-10">
<collectingDate id="C1C57460349E7CBE0B30F523D13150CF" value="1973-11-10">10. 11. 1973</collectingDate>
</date>
</materialsCitation>
;
<collectionCode id="C08066C88252278754102CF02DBC5469" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991430,
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,
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, Point Hodoul, Grande Terre,
<date id="4884811786AA9437D8D9C9168820CA49" value="1974-03-22">22. 03. 1974</date>
,
<location id="57B63FBE3E40FE22F2E5BD69A4BDC976" LSID="urn:lsid:plazi:treatment:DC1DCBCACB155101A2370EB887D8E379:57B63FBE3E40FE22F2E5BD69A4BDC976" municipality="Grande Terre" name="Point Hodoul">Point Hodoul</location>
,
<collectingMunicipality id="10D80389D651735C47430D98293E38B5">Grande Terre</collectingMunicipality>
,
<date id="4884811786AA9437D8D9C9168820CA49" value="1974-03-22">
<collectingDate id="4EC5B3A4E3B72D78342F0FC1EB4D2293" value="1974-03-22">22. 03. 1974</collectingDate>
</date>
</materialsCitation>
;
<collectionCode id="E25977E9D4FC9438309F24153D34747D" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991431,
<materialsCitation id="FF72CEF0C4950CD89A09E7CDB7005FC9" collectingDate="1974-06-08" collectionCode="NHMUK" location="Ile. Malabar" specimenCode="NHMUK 015991431" specimenCount="2">
<specimenCode id="A6DA8A346F1B2685236E8BD385632223">NHMUK 015991431</specimenCode>
,
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, Ile. Malabar,
<date id="05468718727284E29438BC8D538554D2" value="1974-06-08">08. 06. 1974</date>
,
<location id="E979664F0764EC9A828DDE9E8167D371" LSID="urn:lsid:plazi:treatment:DC1DCBCACB155101A2370EB887D8E379:E979664F0764EC9A828DDE9E8167D371" name="Ile. Malabar">Ile. Malabar</location>
,
<date id="05468718727284E29438BC8D538554D2" value="1974-06-08">
<collectingDate id="7ABBF7094BAAA6058D663BA50940386B" value="1974-06-08">08. 06. 1974</collectingDate>
</date>
</materialsCitation>
;
<collectionCode id="D6502A668C4681D039C676E1997D5213" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991432,
<materialsCitation id="AC12E520B0D295FB2A31E68B57539AC4" collectedFrom="Casuarina and Sideroxylon litter" collectingDate="1974-02-12" collectionCode="NHMUK" location="Anse Cedres" specimenCode="NHMUK 015991432" specimenCount="18">
<specimenCode id="20B5220E49124AEAFA27A1BCEA8F616A">NHMUK 015991432</specimenCode>
,
<specimenCount id="14EFA1AA8C89F75628EA48FC9A27D68E" count="18" type="generic">18 specimens</specimenCount>
,
<collectedFrom id="7D9C4F45C4077027C4233C0F3093EBC5">
<taxonomicName id="981B9EF0B8B11B3DFD8DCAD66B613B90" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" phylum="Tracheophyta" rank="genus">
<emphasis id="2E7BAB716F30A0BD08D98745C65AB1D6" italics="true">Casuarina</emphasis>
</taxonomicName>
@ -100,14 +128,27 @@ and
<taxonomicName id="95FF36A2F6158F8EDAD0F61A647E305B" class="Magnoliopsida" family="Sapotaceae" genus="Sideroxylon" kingdom="Plantae" order="Ericales" phylum="Tracheophyta" rank="genus">
<emphasis id="6D53794B94F5462CF137C1E08EA1FEB2" italics="true">Sideroxylon</emphasis>
</taxonomicName>
litter, Anse Cedres,
<date id="BA707682AC3B9C47A2F49710456DBF57" value="1974-02-12">12. 02. 1974</date>
litter
</collectedFrom>
,
<location id="62B5EE288E69E3B6D663CA744C995CDB" LSID="urn:lsid:plazi:treatment:DC1DCBCACB155101A2370EB887D8E379:62B5EE288E69E3B6D663CA744C995CDB" name="Anse Cedres">Anse Cedres</location>
,
<date id="BA707682AC3B9C47A2F49710456DBF57" value="1974-02-12">
<collectingDate id="CDE2BDB2CA9153D08D52154CD8EA5C3E" value="1974-02-12">12. 02. 1974</collectingDate>
</date>
</materialsCitation>
;
<collectionCode id="C05F52F9285F1BA2EF8B4DB7328AC884" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
015991433,
<materialsCitation id="7F2730604C341659C86FD26B5830D13F" collectingDate="1974-12-14" collectionCode="NHMUK" location="Esprit" specimenCode="NHMUK 015991433" specimenCount="4">
<specimenCode id="735A0A11CA330375CF8B856DB246D3D5">NHMUK 015991433</specimenCode>
,
<specimenCount id="632D9DAE434F70A49D00F8AD307548A9" count="4" type="generic">4 specimens</specimenCount>
, Esprit,
<date id="B5BB82EC452EE135784362AAC27DD709" value="1974-12-14">14. 12. 1974</date>
,
<location id="E03D91CC307E9E7FA3CB25BE7A4E57FA" LSID="urn:lsid:plazi:treatment:DC1DCBCACB155101A2370EB887D8E379:E03D91CC307E9E7FA3CB25BE7A4E57FA" name="Esprit">Esprit</location>
,
<date id="B5BB82EC452EE135784362AAC27DD709" value="1974-12-14">
<collectingDate id="10F70C594406170823F896CFC706B71B" value="1974-12-14">14. 12. 1974</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
@ -172,17 +213,20 @@ cf.
<bibRefCitation id="15261A89CB257BDBF90C5259033673F7" author="Takakuwa Y" journalOrPublisher="Transactions of the Sapporo Natural History Society" pagination="398-406" refId="B100" refString="Takakuwa Y (1934) Sechs neue Arten der Chilopoden aus Japan. Transactions of the Sapporo Natural History Society 13: 398-406." title="Sechs neue Arten der Chilopoden aus Japan." volume="13" year="1934">Takakuwa, 1934</bibRefCitation>
</taxonomicName>
<emphasis id="3DB04C087A9FA52246A7B96EFCD706AA" bold="true">A, B, D</emphasis>
NHMUK 015991433
<specimenCode id="76D0C81437F091450ACDB6CE8B229049">NHMUK 015991433</specimenCode>
<emphasis id="D6573F4F183950A460C52DB082E88FE4" bold="true">A</emphasis>
anterior margin of forcipular coxosternite, ventral view
<emphasis id="B35D6D6A3F2B5AB41F78DD7532F69A5A" bold="true">B</emphasis>
Calyx of venom gland, lateral view
<emphasis id="8A66A79CF7287780CD40467D23AB06B2" bold="true">C</emphasis>
NHMUK 015991430, coxopleuron of ultimate leg-bearing segment, lateral view
<specimenCode id="D06B403D6C11B6FF9B2633A936B066D8">NHMUK 015991430</specimenCode>
, coxopleuron of ultimate leg-bearing segment, lateral view
<emphasis id="FFF2B79C68504860094650F39ED3428A" bold="true">D</emphasis>
femur, tibia, and tarsus 1 of ultimate leg telopodite, lateral view
<emphasis id="AAB011829A99BCDFA07777A23E0AA5FD" bold="true">E</emphasis>
. NHMUK 015991429, pretarsus of leg pair 8, lateral view.
.
<specimenCode id="80E3AFD361AC3948609C1E522020A184">NHMUK 015991429</specimenCode>
, pretarsus of leg pair 8, lateral view.
</paragraph>
</caption>
<paragraph id="3B5EEBF3A337B1A7FCC1DD3539758F03">