Add updates up until 2025-02-07 21:12:41

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<mods:title id="AB49C2DEBE206FEA182ACE948878BE5E">Two new species in the green lacewing genus Santocellus (Neuroptera, Chrysopidae, Leucochrysini)</mods:title>
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<mods:affiliation id="72E57077758D519D7D41BC327C7ED965">Department of Entomology &amp; Nematology, University of California, Davis, CA 95616 and Department of Entomology, Comstock Hall, Cornell University, Ithaca, NY, USA</mods:affiliation>
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<mods:affiliation id="26C5F2566A30671D8C9FA87A180900FA">Department of Entomology &amp; Nematology, University of California, Davis, CA 95616 and Department of Entomology, Comstock Hall, Cornell University, Ithaca, NY, USA</mods:affiliation>
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<paragraph id="9066EBD974A367813818303B35D06AA4">
<taxonomicName id="B03B7132EE5C92FEDC9A5F9EF37AF857" authorityName="Tauber" authorityYear="2025" class="Insecta" family="Chrysopidae" genus="Santocellus" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="legrandi" status="sp. nov.">
@ -74,23 +74,18 @@
</subSubSection>
<subSubSection id="SECID0EFJAC" type="Type specimens">
<paragraph id="8F71C39D187BF228DA140D63F7D53C76">
<heading id="4774B0D64884F867D41F089AAE0B3DA4" reason="title">
<typeStatus id="07B55DA0FE3CE71C0A32FAD248B68693">Type</typeStatus>
specimens.
</heading>
<heading id="4774B0D64884F867D41F089AAE0B3DA4" reason="title">Type specimens.</heading>
</paragraph>
<paragraph id="289E9CC10EF1ADCEB2EF8466E155472E">
<emphasis id="A3230220A5CD63A02EF9113D69E9D724" bold="true">
(sex and figures; verbatim label data [specimen location])
<materialsCitation id="6AF602C1D8389D958C3DADE6B157C12D" collectionCode="CAS" collectorName="P. Duelli &amp; U. &amp; H. Aspoeck" country="Nicaragua" county="Male" elevation="15" latitude="10.915555" location="Bartola" longLatPrecision="21" longitude="-84.299164" municipality="Figs" specimenCount="1" specimenCount-male="1" stateProvince="Rio San Juan" typeStatus="holotype">
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<typeStatus id="D8EE2A034AD35A31051C9E482A99839D" type="holotype">Holotype</typeStatus>
</emphasis>
</emphasis>
<materialsCitation id="EAAE9CE3495037FEC0ABE7561CF5648A" collectionCode="CAS" collectorName="P. Duelli &amp; U. &amp; H. Aspoeck &amp; Allotype" country="Nicaragua" county="Male" elevation="15" latitude="10.915555" location="Bartola" longLatPrecision="21" longitude="-84.299164" municipality="Figs" specimenCount="1" stateProvince="Rio San Juan" typeStatus="holotype">
<collectingCounty id="5DC3B62A932AD70DBE641D127B78B259">Male</collectingCounty>
(
<collectingMunicipality id="47596D85BE9C97CCD5A133582C1E45E7">Figs</collectingMunicipality>
<specimenCount id="2B37C35CCFAFE322A75526D541693292" count="1" type="male">Male</specimenCount>
(Figs
<figureCitation id="9983C4E4C2F80A61B3BE95CB6B388321" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Head of Santocellus legrandi, sp. nov. Holotype, male, Nicaragua, Río San Juan, Bartola (CAS) a frontal b head, prothorax, dorsal c head, frontolateral d ventral. Width across head (dorsal, including eyes), 1.85 mm." figureDoi="10.3897/zookeys.1226.140386.figure1" httpUri="https://binary.pensoft.net/fig/1244337">1</figureCitation>
<figureCitation id="64C936A675CBEB3643C2328C4CFD265F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male genitalia of Santocellus legrandi sp. nov., holotype, Nicaragua, Río San Juan, Bartola (CAS) a gonarcal complex, dorsal b same, lateral c same, frontal d same, posteroventral. Abbreviations: gcr, field of gonocristae on terminal membrane of ninth sternite; gsac gonosaccus; gst gonosetae; g. ap. gonarcal apodeme; mu-d mediuncus, dorsal view; mu-f mediuncus." figureDoi="10.3897/zookeys.1226.140386.figure5" httpUri="https://binary.pensoft.net/fig/1244341">5</figureCitation>
@ -116,54 +111,55 @@ specimens.
<named-content id="45CE3629DD88011A5ACA59E3CB32B381" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">
<collectionCode id="E58EA14E5A3FF3AF1FD17D11010A0710" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
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<typeStatus id="E0977EFAC19ACEB873A86517043360C0" type="allotype">
<collectorName id="C88DAF44D75CA1EF23011CF8E64482FC">Allotype</collectorName>
</typeStatus>
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]
</materialsCitation>
</emphasis>
.
<materialsCitation id="1E4A9B80D384C08606896078FA8F43D3" collectionCode="SDMC" collectorName="J. T. Doyen &amp; P. A. Opler" country="Costa Rica" location="N. San Ysidro" municipality="Female" specimenCount="1" specimenCount-female="1" stateProvince="San Jose Prov." typeStatus="allotype">
<emphasis id="1AA4124C21F8F83429A55E9648A1422A" bold="true" italics="true">
<typeStatus id="E0977EFAC19ACEB873A86517043360C0" type="allotype">Allotype</typeStatus>
</emphasis>
<materialsCitation id="C7D31BE0D1CFE8DF4E3BC97251F64F58" collectionCode="T, SDMC" collectorName="N. San Ysidro &amp; P. A. Opler" country="Costa Rica" location="Fig." municipality="Female" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
<collectingMunicipality id="99D83850FE975A26F85FCB9D0DCE798E">Female</collectingMunicipality>
(
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<specimenCount id="4998255EA7A3E922205D84969063522B" count="1" type="female">Female</specimenCount>
(Fig.
<figureCitation id="8473DD95BED88A4A1AA2CFF3E5480D81" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cleared abdominal terminus of female Santocellus legrandi sp. nov., allotype, Costa Rica, San José (SDMC) a segments A 7 to terminus, lateral b genitalia, lateral c subgenitale, distal view d spermathecal complex below bursa copulatrix, lateral view. Abbreviations: b. c. bursa copulatrix; c. c. callus cerci; g. l. gonapophysis lateralis; k-sg distal knob of subgenitale; sg-b base of subgenitale; sp spermatheca; sp. d. spermathecal duct; sp. i. spermathecal invagination; S 7 seventh sternite; T 8 eighth tergite; vel, spermathecal velum." figureDoi="10.3897/zookeys.1226.140386.figure6" httpUri="https://binary.pensoft.net/fig/1244342">6</figureCitation>
). (1)
<collectingCountry id="6C4BA9832F55FD5543E3EB05DDF7FC79" name="Costa Rica">Costa Rica</collectingCountry>
,
<quantity id="020A47D8D798282363A19365A8ED493C" metricMagnitude="4" metricUnit="m" metricValue="2.2" unit="km" value="22.0">22 km</quantity>
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<locationDeviation id="0B2ACB1DD4A067C4512ABAD836528610" metricMagnitude="4" metricUnit="m" metricValue="2.2" unit="km" value="22.0">22 km</locationDeviation>
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<collectorName id="6084DA9A3037B723CD20BB6E1FFE1A85">N. San Ysidro</collectorName>
<location id="475E5985EC7ECD6AAC3CBF656C0995B9" LSID="urn:lsid:plazi:treatment:E596F489F95457B7BC3DD5E91CE47B2D:475E5985EC7ECD6AAC3CBF656C0995B9" country="Costa Rica" municipality="Female" name="N. San Ysidro" stateProvince="San Jose Prov.">N. San Ysidro</location>
, /
<collectingRegion id="2EEA835DCE6573E1BDBE9381EE5790B6" country="Costa Rica" name="San Jose">San Jose Prov.</collectingRegion>
/ V- 24-85 (2) J.
<collectionCode id="94AD5D8D0B49040F77E479C4C9060BAE" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
. Doyen, /
/ V- 24-85 (2)
<collectorName id="7188DE365B60F4E2BFD9616F30A08F1F">J. T. Doyen</collectorName>
, /
<collectorName id="50B09D97E59D2A24190A14681E46691F">P. A. Opler</collectorName>
/ Collectors [
<named-content id="8BD90292507B20E17840F430C06AE4EB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/san-diego-natural-history-museum" xlink_title="San Diego Natural History Museum">
<collectionCode id="ADDEAD6E24EE3EB1DCFF9D613596A908">SDMC</collectionCode>
</named-content>
].
]
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.
<materialsCitation id="77DED1E57D075B0A9EA0DE9D81D4976E" collectingDate="2001-07-31" collectionCode="OUMNH" collectorName="G. C. McGavin" country="Belize" county="Chiquibul Forest" location="Nature Trail" municipality="Las Cuevas" specimenCode="OUM-2002-006" specimenCount="1" specimenCount-female="1" stateProvince="Cayo" typeStatus="paratype">
<emphasis id="8766D9AB62206AD799BAF5036759F4CF" bold="true" italics="true">
<typeStatus id="4828B410212DACC7CAEDD226E8F8FA6C" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
<materialsCitation id="9AE95CCA23F694E544D8A8C78A825616" collectingDate="2001-07-31" collectionCode="OUMNH" collectorName="G. C. McGavin Coll. &amp; Pres. G. C. McGavin" country="Belize" county="Female." location="Chiquibul Forest" municipality="Chiquibul Forest. Las Cuevas" specimenCount="1" stateProvince="Cayo" typeStatus="holotype">
<collectingCounty id="7F55955886F1A0A0FBAFEC1823E4D303">Female.</collectingCounty>
(1)
<specimenCount id="08CA287B50668CAFDBDBB6546BA583B4" count="1" type="female">Female</specimenCount>
. (1)
<collectingCountry id="06116495582DB57123422B843AFCB22A" name="Belize">Belize</collectingCountry>
:
<collectingRegion id="5B8945D137F69CA63113671F4AF12513" country="Belize" name="Cayo">Cayo</collectingRegion>
/
<collectingMunicipality id="DF77FC246B94737D672D74DD5F85A621">
<location id="B5DAFF8A1EAFD8C94A947CA04589FA0F" LSID="urn:lsid:plazi:treatment:E596F489F95457B7BC3DD5E91CE47B2D:B5DAFF8A1EAFD8C94A947CA04589FA0F" country="Belize" county="Female." municipality="Chiquibul Forest. Las Cuevas" name="Chiquibul Forest" stateProvince="Cayo">Chiquibul Forest</location>
. Las Cuevas
</collectingMunicipality>
<collectingCounty id="7FD3317EF090D85EB35BD3E067761771">Chiquibul Forest</collectingCounty>
.
<collectingMunicipality id="82A9DF79B84DB1ABB52DA1AD6C415509">Las Cuevas</collectingMunicipality>
/
<location id="753DE13829E0861B7B4709E57BCC42D4" LSID="urn:lsid:plazi:treatment:E596F489F95457B7BC3DD5E91CE47B2D:753DE13829E0861B7B4709E57BCC42D4" country="Belize" county="Female." municipality="Chiquibul Forest. Las Cuevas" name="Nature Trail" stateProvince="Cayo">Nature Trail</location>
<location id="753DE13829E0861B7B4709E57BCC42D4" LSID="urn:lsid:plazi:treatment:E596F489F95457B7BC3DD5E91CE47B2D:753DE13829E0861B7B4709E57BCC42D4" country="Belize" county="Chiquibul Forest" municipality="Las Cuevas" name="Nature Trail" stateProvince="Cayo">Nature Trail</location>
/
<date id="653CEB501668E6F78482B3C20C760260" value="2001-07-31">
<collectingDate id="D8C5140F1A74C5838113AFE664E12AA3" value="2001-07-31">
@ -173,48 +169,48 @@ July 2001
</collectingDate>
</date>
/
<collectorName id="F54BA1653C9964E9D273CB93C58A131D">G. C. McGavin Coll.</collectorName>
(2)
<collectorName id="0D2259B13100D01A6BCC92BEB3D58903">G. C. McGavin Coll.</collectorName>
/
<collectorName id="E7D37F747699DCF3D638D99F9B54EB90">Pres. G. C. McGavin</collectorName>
/ OUM- 2002-006 [
<collectorName id="0B05080BF525D39974C801BF7C3538E9">G. C. McGavin</collectorName>
Coll. (2)
<collectorName id="A2AED53456C3C39F7857CA142D67403F">G. C. McGavin</collectorName>
Coll. / Pres. G. C. McGavin /
<specimenCode id="40D59811D528D682D2C6C8AC0D1DC453">OUM-2002-006</specimenCode>
[
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].
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.
<materialsCitation id="94EE4680E89BD56649418C39959BD715" collectionCode="CAS" collectorName="T. H. Haldell &amp; Phillip A. Adams" country="Honduras" county="Male." location="Barraneo" municipality="Rio Paulaya" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="98641DEE3EE8F2514FD76BEAD79AD8C5" bold="true" italics="true">
<typeStatus id="61B248FEC841C5AA3966EA35687DD9C9" type="paratype">Paratype</typeStatus>
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<materialsCitation id="9561ECEA64CDE18537ACF031C6487207" collectionCode="T, CAS" collectorName="H. Haldell &amp; Phillip A. Adams" country="Honduras" county="Male." location="Barraneo" municipality="Rio Paulaya" specimenCount="1" typeStatus="holotype">
<collectingCounty id="3B35528829C3292740E293BDF2FD8ECC">Male.</collectingCounty>
(1)
<specimenCount id="EE9667B919A0DBEDA58740904BD2ABE6" count="1" type="male">Male</specimenCount>
. (1)
<collectingCountry id="B37BD4296B6A611D56770C05A7DCAE2E" name="Honduras">Honduras</collectingCountry>
1923 /
<collectingMunicipality id="DF104567DF1E1C9F7F045A44F5133E98">Rio Paulaya</collectingMunicipality>
: /
<location id="351A18DC94037E93D4225C15E597C3FD" LSID="urn:lsid:plazi:treatment:E596F489F95457B7BC3DD5E91CE47B2D:351A18DC94037E93D4225C15E597C3FD" country="Honduras" county="Male." municipality="Rio Paulaya" name="Barraneo">Barraneo</location>
, iv- 17 / 303
<collectionCode id="29DD89F32240FB424954D43980AB4BEA" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
.
<collectorName id="2849DCBF177AD3F72541CF9AC8132146">H. Haldell</collectorName>
<collectorName id="AA7BCE9D3E57E7BD1F373493128480DD">T. H. Haldell</collectorName>
(2)
<collectorName id="87B9CCD25CF3CDE5E80D56EF267EC382">Phillip A. Adams</collectorName>
/ Collection / 1998 bequest to / Calif. Acad. Sci. [
<named-content id="164A7D4E5B884FDFABAD14A03E1715CD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/california-academy-sciences" xlink_title="California Academy of Sciences">
<collectionCode id="7A7A798F0EBF0D688ADE9A2093B100AC" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" type="Museum">CAS</collectionCode>
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].
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.
<materialsCitation id="FE4D26AB40E3F4982DB9FE0A501C9B15" collectionCode="NHMUK" country="Panama" determinerName="P. Adams" location="Chiriqui" specimenCount="1" specimenCount-male="1" stateProvince="Chiriqui" typeStatus="paratype">
<emphasis id="2634896BD6FA1D2169536DCBA0F89835" bold="true" italics="true">
<typeStatus id="A31DBCDA871159901B751803CE129CF8" type="paratype">Paratype</typeStatus>
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<materialsCitation id="8605F33FABDF7200E03B972BA7A26F87" collectionCode="NHMUK" country="Panama" determinerName="P. Adams" location="Male." specimenCount="1" stateProvince="Chiriqui" typeStatus="holotype">
<location id="9C0AA9CAEBE17FF15DFEAF2388991DA2" LSID="urn:lsid:plazi:treatment:E596F489F95457B7BC3DD5E91CE47B2D:9C0AA9CAEBE17FF15DFEAF2388991DA2" country="Panama" name="Male." stateProvince="Chiriqui">Male.</location>
(1)
<specimenCount id="BF9C6A524333E98D7059280DE4CCB40E" count="1" type="male">Male</specimenCount>
. (1)
<collectingRegion id="8425BE03ED463F176036238B702E74AB" country="Panama" name="Chiriqui">Chiriqui</collectingRegion>
(2) prob.
<taxonomicName id="B788781130F5E676D0CE361CCCE85DD3" authority="Esb-Pet" authorityName="Esben-Petersen" authorityYear="1933" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" phylum="Arthropoda" rank="species" species="risi">
@ -486,26 +482,16 @@ diameter), with 3335 long trichobothria; cupuliform bases of variable size [t
<quantity id="05885583441D5A279ACE82A12688E4AE" metricMagnitude="-5" metricUnit="m" metricValue="2.6" unit="mm" value="0.026">0.026 mm</quantity>
) than those on periphery (~
<quantity id="826DBE146FADCA94A0A175826A99C797" metricMagnitude="-5" metricUnit="m" metricValue="1.4" unit="mm" value="0.014">0.014 mm</quantity>
)]. Dorsum of
<collectionCode id="A3AE5728F438E7FA85A462838FA7848A" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct rounded distally, fused mesally, with very small to no cleft; no midline suture visible; with long setae, densest along distal margin; dorsal apodeme simple, straight, short, extending around proximal side of callus cerci. S 8 + 9 fused without suture, tapering to acute apex (lateral view); S 9 slightly darker than S 8; ventral apodeme extending across dorsal surface of S 8 + 9; distal margin of S 8 + 9 spatulate, with terminal membrane prominent; membrane bearing moderately heavy gonocristae in two broad (somewhat irregular) lateral patches, single, small mesal patch of several (~ 3) gonocristae.
)]. Dorsum of T 9 + ectoproct rounded distally, fused mesally, with very small to no cleft; no midline suture visible; with long setae, densest along distal margin; dorsal apodeme simple, straight, short, extending around proximal side of callus cerci. S 8 + 9 fused without suture, tapering to acute apex (lateral view); S 9 slightly darker than S 8; ventral apodeme extending across dorsal surface of S 8 + 9; distal margin of S 8 + 9 spatulate, with terminal membrane prominent; membrane bearing moderately heavy gonocristae in two broad (somewhat irregular) lateral patches, single, small mesal patch of several (~ 3) gonocristae.
<emphasis id="33BFF024980E9FE3BE057463F7416161" bold="true" italics="true">Coloration</emphasis>
: Dorsum of each segment pale with large black central mark; marks on
<collectionCode id="C5B365F7E9E9EC717DE97BA9C5862801" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
2
<collectionCode id="CB0E7DC28C4C97FB0903973234EAFD15" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
8 with prominent white spot in center of black area; mark on
<collectionCode id="D5440B1E1D64F2EB6B1FFC3848AEB8F2" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
1 without white spot; venter, pleuron white to cream. Setae golden, long; dorsal half of callus cerci light cream colored; ventral half black.
: Dorsum of each segment pale with large black central mark; marks on T 2 T 8 with prominent white spot in center of black area; mark on T 1 without white spot; venter, pleuron white to cream. Setae golden, long; dorsal half of callus cerci light cream colored; ventral half black.
</paragraph>
<paragraph id="1C81746651BA58F8327099E28725F055">
<emphasis id="72F774EA6B01371ABDE698DDC7B08092" bold="true">
<emphasis id="1A8641650A14F842F945ECAE822436A7" bold="true" italics="true">Male genitalia</emphasis>
.
</emphasis>
Tignum, gonapsis, parameres, quadrate hood, pseudopenis, spinellae absent; S 8 + 9 fused, elongate, not extending beyond
<collectionCode id="0A4BB43DD13EC3CDDB6BEE76A070122A" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct. Gonarcus broad, with very small gonocornua on frontolateral margin of gonarcal bridge, with relatively large, rounded gonarcal arms expanded forward slightly; mediuncus in two sections; basal section extending from below gonarcal bridge, with pair of internal apodemes dorsomesally, with dorsal surface membranous, broad basally, tapering distally; distal section of mediuncus extending perpendicularly from tip of basal section, distally with flange-like lateral arms partially surrounding mesal beak; gonosaccus beneath mediuncus, stiff, flat, laterally with a few gonosetae on chalazae.
Tignum, gonapsis, parameres, quadrate hood, pseudopenis, spinellae absent; S 8 + 9 fused, elongate, not extending beyond T 9 + ectoproct. Gonarcus broad, with very small gonocornua on frontolateral margin of gonarcal bridge, with relatively large, rounded gonarcal arms expanded forward slightly; mediuncus in two sections; basal section extending from below gonarcal bridge, with pair of internal apodemes dorsomesally, with dorsal surface membranous, broad basally, tapering distally; distal section of mediuncus extending perpendicularly from tip of basal section, distally with flange-like lateral arms partially surrounding mesal beak; gonosaccus beneath mediuncus, stiff, flat, laterally with a few gonosetae on chalazae.
</paragraph>
<paragraph id="CFC8D9F243F0F1AC9EA424043D7029A4">
<emphasis id="FDECD8E62DADDF433C2C5697EE78EB00" bold="true" italics="true">Female abdomen</emphasis>
@ -521,11 +507,7 @@ in diameter, with 3637 trichobothria of variable length; cupuliform bases als
<geoCoordinate id="BA8D89DAD9FF1E332EB0C6E3BDE15420" degrees="0.029" direction="west" orientation="longitude" precision="55" value="-0.029">0.029</geoCoordinate>
mm
</quantity>
). Dorsum of
<collectionCode id="F681F6FD6A4E180131AD2E3D276BA1E9" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct rounded posteriorly, tall (lateral view), extending approximately to lateral margin of gonapophysis lateralis. Sternite 7 with dorsal margin relatively straight; distal margin blunt, not particularly heavy, with setae mostly missing. Gonapophyses laterales slender, occupying approximately half of abdominal posterior margin; interior membranous area not greatly expanded or inflatable.
<collectionCode id="02C4B14EC64728C0760288ED6C03D50E" name="Tavera, Department of Geology and Geophysics">T</collectionCode>
9 + ectoproct truncate posteriorly, extending slightly below gonapophyses laterales (lateral view). Gonapophyses laterales occupying ~ 6070 % of posterior margin of abdomen.
). Dorsum of T 9 + ectoproct rounded posteriorly, tall (lateral view), extending approximately to lateral margin of gonapophysis lateralis. Sternite 7 with dorsal margin relatively straight; distal margin blunt, not particularly heavy, with setae mostly missing. Gonapophyses laterales slender, occupying approximately half of abdominal posterior margin; interior membranous area not greatly expanded or inflatable. T 9 + ectoproct truncate posteriorly, extending slightly below gonapophyses laterales (lateral view). Gonapophyses laterales occupying ~ 6070 % of posterior margin of abdomen.
</paragraph>
<paragraph id="5AD597955ADCA869F03B44DD9B67E11B">
<emphasis id="E90FB19B6CB2620B0B08E21B4A459064" bold="true">