Add updates up until 2025-02-21 18:43:30

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(
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)
</paragraph>
</subSubSection>
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<treatmentCitation id="FC7510A4FFFD3119EBD0FE93FE64171A" author="KOTT P." box="[151,474,255,280]" page="61" pageId="34" pageNumber="235" year="1990">
<taxonomicName id="BAD44D36FFFD3119EBD0FE93FE64171A" ID-CoL="VV8P" authority="Kott, 1990: 61" authorityName="Kott" authorityPageNumber="61" authorityYear="1990" box="[151,474,255,280]" class="Ascidiacea" family="Clavelinidae" genus="Clavelina" kingdom="Animalia" order="Aplousobranchia" pageId="34" pageNumber="235" phylum="Chordata" rank="species" species="robusta">
<emphasis id="4FA0EAA7FFFD3119EBD0FE93FE83171A" box="[151,317,256,280]" italics="true" pageId="34" pageNumber="235">Clavelina robusta</emphasis>
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</taxonomicName>
</treatmentCitation>
, fig. 20, pl.
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.
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locality:
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. — Monniot F. &amp; Monniot C. 1996: 210, fig. 37a-c, pl. 6E.
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</paragraph>
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. —
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<collectingCountry id="05C37625FFFD3119EAEEFEC8FD961771" box="[425,552,347,371]" name="Philippines" pageId="34" pageNumber="235">Philippines</collectingCountry>
.
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SW
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, Panglao Island,
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,
<geoCoordinate id="18E05072FFFD3119EA97FEE7FDE3178E" box="[464,605,372,396]" degrees="123" direction="east" minutes="48.56" orientation="longitude" pageId="34" pageNumber="235" precision="9" value="123.80933">123°48.56E</geoCoordinate>
,
<quantity id="BA2C9B50FFFD3119E92BFEE7FD191788" box="[620,679,371,396]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="34" pageNumber="235" unit="m" value="10.0">10 m</quantity>
,
<date id="096A1075FFFD3119EBD0FE1EFEB717A7" box="[151,265,397,421]" pageId="34" pageNumber="235" value="1997-04-20">20.IV.1997</date>
(
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A3
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109).
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</paragraph>
</subSubSection>
<subSubSection id="35CE653EFFFD3119EBD0FE74FDFD1520" pageId="34" pageNumber="235" type="discussion">
<paragraph id="7D6B36B5FFFD3119EBD0FE74FF4517FD" blockId="34.[151,684,487,802]" box="[151,251,487,513]" pageId="34" pageNumber="235">REMARKS</paragraph>
<paragraph id="7D6B36B5FFFD3119EBD0FD9BFDFD1520" blockId="34.[151,684,487,802]" pageId="34" pageNumber="235">
The colony has the typical colour of the species, with yellow rings around the siphons (
<figureCitation id="E5EF2A30FFFD3119EBD8FDDBFEAC1460" box="[159,274,584,610]" captionStart="FIG" captionStartId="157.[151,162,1565,1582]" captionTargetBox="[162,1240,227,1527]" captionTargetId="figure-71@157.[162,1261,227,1550]" captionTargetPageId="157" captionText="FIG. 116. — A, Clavelina robusta Kott, 1990; B, Citorclinum laboutei Monniot F. &amp; Millar, 1988; C, Cystodytes hapu Monniot C. &amp; Monniot F., 1987; D, Eudistoma album Monniot F., 1988; E, Eudistoma fluorescens n. sp.; F, Eudistoma fucatum n. sp." figureDoi="http://doi.org/10.5281/zenodo.5391665" httpUri="https://zenodo.org/record/5391665/files/figure.png" pageId="34" pageNumber="235">Fig. 116A</figureCitation>
). The tunic is transparent. Dark blue and almost black pigment, is concentrated in the body wall, and this makes contracted animals opaque. In formalin, the musculature appears white on a purplish background. This colony contains numerous larvae incubated in the oviduct at the base of the cloacal cavity.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,708 @@
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<subSubSection id="SECID0EWLAI" type="Paratypes">
<paragraph id="90CFE51FA1D1BC4F25562363FE697575">
<heading id="07042C99EE0A7823BF3244032343D53B" reason="title">
<typeStatus id="729017B2EB740FF94D6032635CAD57F3" type="paratype">Paratypes</typeStatus>
.
</heading>
</paragraph>
<paragraph id="ED06EF30025FE809FE0A6A34A9B1811C">
<materialsCitation id="D1342D358C8CE5713620D241A306004E" collectingDate="2023-08-05" collectionCode="MT" collectorName="Chaobo Feng" country="China" county="Leishan County" elevation="1570" latitude="26.3733" location="Leishan County" longLatPrecision="6" longitude="108.1967" specimenCode="LS 20230805003" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou Province" typeStatus="paratype">
<specimenCount id="587FC791ECEC6AAC0C9A4E424C79F2E7" count="1" type="male">One male</specimenCount>
<abbrev id="ABBRID0E3LAI" xlink_title="Moutai Institute">
<collectionCode id="1E0826A9F78178C4AD0C81458242F4F5" country="Russia" name="Mus. Tinro, Vladyvostok">MT</collectionCode>
</abbrev>
<specimenCode id="0EC2657A12D38AE56381896C2BC7558C" collectionCode="LS" country="United Kingdom" name="Linnean Society of London">LS 20230805003</specimenCode>
was collected by
<collectorName id="0B98087B8D7D529B66DD54E6A4659D5E">Chaobo Feng</collectorName>
on
<date id="1DB00338191EDAFD0334B1DAD8B08E58" value="2023-08-05">
<collectingDate id="0634708BCAF28149D73511648AAD1B28" value="2023-08-05">5 August 2023</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="6CE23608F2DEA4AF4C72B8F49EA3C02A" collectingDate="2024-05-18" collectionCode="MT" collectorName="Jing Liu" country="China" county="Leishan County" elevation="1570" latitude="26.3733" location="Leishan County" longLatPrecision="6" longitude="108.1967" specimenCode="LS 20240518009, LS 20240518010" specimenCount="2" specimenCount-female="2" stateProvince="Guizhou Province" typeStatus="paratype">
<specimenCount id="D871CEDAA5EAFE72613C3D318D1D3C34" count="2" type="female">two females</specimenCount>
,
<abbrev id="ABBRID0EAMAI" xlink_title="Moutai Institute">
<collectionCode id="54611A2E9AD77D0E6956F7A11D784C2F" country="Russia" name="Mus. Tinro, Vladyvostok">MT</collectionCode>
</abbrev>
<specimenCode id="23EC1782B93187B20115471F367D88D7" collectionCode="LS" country="United Kingdom" name="Linnean Society of London">LS 20240518009</specimenCode>
and
<abbrev id="ABBRID0EEMAI" xlink_title="Moutai Institute">
<collectionCode id="8C7410E28939BD4D87D91BBD6789A400" country="Russia" name="Mus. Tinro, Vladyvostok">MT</collectionCode>
</abbrev>
<specimenCode id="A92F42F12163DC25030C31CBB279B534" collectionCode="LS" country="United Kingdom" name="Linnean Society of London">LS 20240518010</specimenCode>
, were collected by
<collectorName id="43E1FF32BD3B4D7183550804E9CCB41C">Jing Liu</collectorName>
on
<date id="7648EAB5A08E298450944A9150B3C240" value="2024-05-18">
<collectingDate id="C57F7A2F1C06776F51FF9BD0DCE4B3D2" value="2024-05-18">18 May 2024</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="2982C7625EFC83A1C52E6E03B65F597F" collectingDate="2024-05-18" collectionCode="MT" collectorName="Caichun Peng" country="China" county="Leishan County" elevation="1570" latitude="26.3733" location="Leishan County" longLatPrecision="6" longitude="108.1967" specimenCode="LS 20240517001, LS 20240517002, LS 20240517003" specimenCount="3" specimenCount-male="3" stateProvince="Guizhou Province" typeStatus="paratype">
and
<specimenCount id="E836E3B7FFDF90220AD410841CBD025F" count="3" type="male">three males</specimenCount>
,
<abbrev id="ABBRID0EIMAI" xlink_title="Moutai Institute">
<collectionCode id="E9EBBB6DD89AEBE4143F2592AF7682D8" country="Russia" name="Mus. Tinro, Vladyvostok">MT</collectionCode>
</abbrev>
<specimenCode id="827A22AB5D1B12DAD8076B1BBCDE6DC7" collectionCode="LS" country="United Kingdom" name="Linnean Society of London">LS 20240517001</specimenCode>
,
<abbrev id="ABBRID0EMMAI" xlink_title="Moutai Institute">
<collectionCode id="A38ACFAFE583EC44E02F7CF4569EC653" country="Russia" name="Mus. Tinro, Vladyvostok">MT</collectionCode>
</abbrev>
<specimenCode id="396F81864FAC30BC75DC60E8564E76B4" collectionCode="LS" country="United Kingdom" name="Linnean Society of London">LS 20240517002</specimenCode>
, and
<abbrev id="ABBRID0EQMAI" xlink_title="Moutai Institute">
<collectionCode id="C97667F569F47E98863B3458B2096FF2" country="Russia" name="Mus. Tinro, Vladyvostok">MT</collectionCode>
</abbrev>
<specimenCode id="869FDA22AD9123E1BEBA9B1572C79F25" collectionCode="LS" country="United Kingdom" name="Linnean Society of London">LS 20240517003</specimenCode>
, were collected by
<collectorName id="074883B9FB7C5AF141CBEF9AD6C27224">Caichun Peng</collectorName>
on
<date id="E925207F16E0E54AC2EBB050C5DE39B0" value="2024-05-18">
<collectingDate id="593E18ED6ECC7C86F96F1C3C81DC9D62" value="2024-05-18">18 May 2024</collectingDate>
</date>
</materialsCitation>
. All of the
<typeStatus id="0223D536D2A4DF222649BE005AEDDDC2" type="paratype">paratypes</typeStatus>
were collected from the same place as the
<typeStatus id="733386F7A5222E6576395818A0B4A919" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUMAI" type="description">
<paragraph id="22F378C4A916FB6E48B1C5947D1F41D4">
<heading id="76EA7E8FEFFBE4C2B9657AC3983C63F1" reason="title">
Description of
<typeStatus id="2B8E7C64762CF79C3ADA18A274316616" type="holotype">holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="96801FB16CF39A665570A8B7D32FE693">
Adult male (Figs
<figureCitation id="C76FCF24B6246C5B9B42295185CD586D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Photos of the holotype MT LS 20240518008 of Gracixalus weii sp. nov. in life. A. Dorsal view; B. Ventral view; C. Dorsal view of hand; D. Ventral view of hand; E. Ventral view of foot. (Photo A E by Shize Li)." figureDoi="10.3897/zse.101.133735.figure4" httpUri="https://binary.pensoft.net/fig/1258205">4</figureCitation>
,
<figureCitation id="C2C0A30A6D053AC2C11FB034BB939515" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of morphological characteristics between the holotype specimen of Gracixalus weii sp. nov. (MT LS 20240518008; A, C, E, G) and Gracixalus jinxiuensis (CIB 58795 (660386); B, D, F, H). A, B. Dorsal view; C, D. Ventral view of hand; E, F. Ventral view; G, H. Ventral view of foot. (Photo A H by Jing Liu)." figureDoi="10.3897/zse.101.133735.figure5" httpUri="https://binary.pensoft.net/fig/1258206">5</figureCitation>
) of moderate body size,
<abbrev id="ABBRID0ECNAI" xlink_title="snout-vent length">SVL</abbrev>
<quantity id="DB4A989DB4956117DFE57BDA7552C935" metricMagnitude="-2" metricUnit="m" metricValue="3.1399999999999997" unit="mm" value="31.4">31.4 mm</quantity>
. The head width was larger than the head length (
<abbrev id="ABBRID0EGNAI" xlink_title="maximum head width">HDW</abbrev>
:
<abbrev id="ABBRID0EKNAI" xlink_title="head length">HDL</abbrev>
= 1.08); the triangular snout showed a pointed appearance in the dorsal view, rounded in the profile, and projecting beyond the margin of the lower jaw; the canthus rostralis was distinct and rounded; the loreal region was oblique and concave; nostrils were oval and significantly protuberant and were closer to the tip of the snout than to the eye; the eyes were large with horizontal pupils; the eye diameter was 0.68 times that of snout length; the distinct tympanum was 0.69 times the eye diameter; the supratympanic fold was distinct, extending from the posterior corner of eye to above the insertion of the arm; no vomerine teeth were observed; the tongue was deeply notched behind.
</paragraph>
<caption id="746C2093B380351B22A8DE216981BA70" ID-DOI="10.3897/zse.101.133735.figure4" ID-arpha="BAC72A0E-C1B5-5E95-86AC-50731E214ADD" httpUri="https://binary.pensoft.net/fig/1258205" startId="F4">
<paragraph id="B69E11B1628B9B67930A0FD78CBB955D">
<label id="10AB1D345A78739E88E6469799C7CAE3">Figure 4.</label>
</paragraph>
<paragraph id="B5A06EAE67DBB90867B21EECE09DF8A6">
Photos of the holotype
<specimenCode id="A49085A6849B1E9CE214AC92C0980CE7">
<abbrev id="ABBRID0EWNAI" xlink_title="Moutai Institute">MT</abbrev>
LS 20240518008
</specimenCode>
of
<taxonomicName id="F463C3D24972B7A2879981CA43A8806A" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="7EA0A3E0FC771B52B709570CC883961D" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6A0579A211AA20711C5C47A61F402962" rank="species">sp. nov.</taxonomicNameLabel>
in life.
<emphasis id="F63195A586604852E4E91BC38EA84269" bold="true">A.</emphasis>
Dorsal view;
<emphasis id="119DCF0E064E8661EE4AFB2CC37EBDA3" bold="true">B.</emphasis>
Ventral view;
<emphasis id="2488FF540845B21D4C6CC19FE5AA0DE4" bold="true">C.</emphasis>
Dorsal view of hand;
<emphasis id="509457D830FFEF09E030D0DCDEC5C1D3" bold="true">D.</emphasis>
Ventral view of hand;
<emphasis id="0E011E9767F4F0EF967A1DC828397005" bold="true">E.</emphasis>
Ventral view of foot. (Photo
<emphasis id="73E634E139766AADB92671AA138662E9" bold="true">A E</emphasis>
by Shize Li).
</paragraph>
</caption>
<caption id="B2577FD7948F92887E92A2087A5ACC17" ID-DOI="10.3897/zse.101.133735.figure5" ID-arpha="70309FB3-15ED-51B0-AB0E-77898754A5C5" httpUri="https://binary.pensoft.net/fig/1258206" startId="F5">
<paragraph id="28FE3B350A4E0B586EE805799CDC5865">
<label id="F0A64567ED07EB7701908B346B6A474D">Figure 5.</label>
</paragraph>
<paragraph id="06E6E4C129481E4E70F5727520AC4DD5">
Comparison of morphological characteristics between the holotype specimen of
<taxonomicName id="919AF3BB1B5047C99B74C146AAF45755" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="63B37AE51A18E3BAD5A534FCAD815561" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0A72A653C5FB9E70B26AD2FF3B9E7DA" rank="species">sp. nov.</taxonomicNameLabel>
(
<specimenCode id="751BCF6EB8A7EE0903CF0B90FD35D713">
<abbrev id="ABBRID0EJPAI" xlink_title="Moutai Institute">MT</abbrev>
LS 20240518008
</specimenCode>
;
<emphasis id="C7FBFD0475F2877C8EF1311DA4089E9A" bold="true">A, C, E, G</emphasis>
) and
<taxonomicName id="14B96108BE468A602C6A52CFF0940F80" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jinxiuensis">
<emphasis id="86A63F0AEAA50D500A074EAF5F32BA8F" italics="true">Gracixalus jinxiuensis</emphasis>
</taxonomicName>
(
<specimenCode id="9526EF8F37AC358D07689CEB92F9AD4C">CIB 58795</specimenCode>
(660386);
<emphasis id="A3B8A7249025F2BC0C511C205465E54F" bold="true">B, D, F, H</emphasis>
).
<emphasis id="4053BB0653C9623A3B0C02EF7FAED1DE" bold="true">A, B.</emphasis>
Dorsal view;
<emphasis id="E50FAA588B2906FB0B2AB949FF4E0B56" bold="true">C, D.</emphasis>
Ventral view of hand;
<emphasis id="50ABEE483F0A2BBC93B37E4F646A1B3C" bold="true">E, F.</emphasis>
Ventral view;
<emphasis id="744DEAC3C6EC824F5FF1BAD4F8BC52AB" bold="true">G, H.</emphasis>
Ventral view of foot. (Photo
<emphasis id="C0D67C164F08F27D3872B90AA62CF573" bold="true">A H</emphasis>
by Jing Liu).
</paragraph>
</caption>
<paragraph id="D73799CEDF48B4D323B1DB55ED47E512">
The forelimbs were robust (
<abbrev id="ABBRID0ENQAI" xlink_title="lower arm width">LW</abbrev>
/
<abbrev id="ABBRID0ERQAI" xlink_title="snout-vent length">SVL</abbrev>
= 0.08); the lower arm and hand beyond one-second of body length (
<abbrev id="ABBRID0EVQAI" xlink_title="length of lower arm and hand">LAL</abbrev>
/
<abbrev id="ABBRID0EZQAI" xlink_title="snout-vent length">SVL</abbrev>
= 0.51); fingers were dorsoventrally compressed, with webbing rudimentary but with narrow lateral fringes on the outer edge of all fingers, relative finger lengths were ordered as I &lt;II &lt;IV &lt;III; the tips of all fingers showed well-developed disks with horizontal circummarginal grooves; markedly elevated, prominent, and rounded subarticular tubercles were found on the fingers (one on fingers I and II, and two on fingers III and IV); supernumerary tubercles were present; the prepollex was distinct, oval; a nuptial pad was observed on the dorsolateral surface of the first finger.
</paragraph>
<paragraph id="3B6EDA1C71AFD7CEED0134C73AF000F3">
Hindlimbs were 1.5 times the
<abbrev id="ABBRID0E6QAI" xlink_title="snout-vent length">SVL</abbrev>
; the tibiotarsal articulation reached the mid-eye level when legs were stretched forward; the tibias were longer than the thigh length; heels overlapped when thighs were positioned at right angles to the body; toes were slender, relative lengths I &lt;II &lt;III &lt;V &lt;IV; tips of toes showed expanded discs with distinct transverse circummarginal grooves. Distinct, rounded subarticular tubercles were seen on the fingers (one on fingers I and II, two on fingers III and V, three on finger IV); webbing formula I (2), (2 ½) II (1 ⅔), (3) III (2), (3) IV (3), (2) V; the sole was smooth with small tubercles; the inner metatarsal tubercle was elongated, ellipsoid; the outer metatarsal tubercle and the inner tarsal fold were absent.
</paragraph>
<paragraph id="10A6E13782FDC960A0E6A2D65068440C">The skin on the dorsal surface of the head, dorsum, and dorsal surfaces of the limbs was rough and sparsely scattered with conical tubercles; the ventral surface of the bell and limbs was rough with small tubercles.</paragraph>
</subSubSection>
<subSubSection id="SECID0EERAI" type="Coloration in life">
<paragraph id="7A706D703F362E5C8E4217C433CF95E2">
<heading id="7439492E6DC958A3EDECEF99EA244F16" reason="title">Coloration in life.</heading>
</paragraph>
<paragraph id="E4212018576934F16150A2EEA16438A5">In life, the dorsal surface ranged from brown to beige, with an inversed Y-shaped dark brown marking from the interorbital region, bifurcating into two branches on the shoulder and extending posteriorly; the dorsal surface of the arm and limbs showed dark brown transverse bands; several large dark blotches were observed on the ventrolateral region of flanks; the ventral surface of the throat, chest, and forelimbs was flesh-colored with small dark specks; the belly had large white tubercles; the ventral surface of the hindlimbs was brown; the disks at the tips of the fingers and toes were a pale yellow.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJRAI" type="Color in preservative">
<paragraph id="9C884C42BFEB176FD273453644D3018C">
<heading id="E99C3A99427B502635174B6E3ABF71B8" reason="title">Color in preservative.</heading>
</paragraph>
<paragraph id="74008F381F374A3C2C6641BF5D2BB386">
The dorsal surface of the head, body, and limbs was brown; the Y-shaped marking and transverse bands were black; the ventral surface of the chest and belly faded to white, with a gray throat (Fig.
<figureCitation id="A7170FF30E175761E0F88A9E75644003" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Comparison of morphological characteristics between the holotype specimen of Gracixalus weii sp. nov. (MT LS 20240518008; A, C, E, G) and Gracixalus jinxiuensis (CIB 58795 (660386); B, D, F, H). A, B. Dorsal view; C, D. Ventral view of hand; E, F. Ventral view; G, H. Ventral view of foot. (Photo A H by Jing Liu)." figureDoi="10.3897/zse.101.133735.figure5" httpUri="https://binary.pensoft.net/fig/1258206">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETRAI" type="variation">
<paragraph id="31D8094033097F685CFEF7118A3B7143">
<heading id="896AFE5C2D6F47EEF0CDD436E262E55C" reason="title">Variation.</heading>
</paragraph>
<paragraph id="AC4ADE9AC7ADF43A68376CD7780C6793">
The measurements of all specimens are listed in Table
<tableCitation id="5CF188F529FD6576365A7302EF0745D9" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Measurements of the adult specimens of Gracixalus weii sp. nov. Comparison of male Gracixalus weii sp. nov. (PW) and Gracixalus jinxiuensis (PJ). Units are given in mm. P - values less than 0.05 indicate significant differences. See abbreviations for the morphological characters in the Materials and methods section." httpUri="http://table.plazi.org/id/D72E49A3145B422A40E1746AA6D01A01" tableUuid="D72E49A3145B422A40E1746AA6D01A01">3</tableCitation>
and Suppl. material
<xref id="167121B7ABB2B7C8051D63E109AA8763" ref-type="supplementary-material" rid="S3">3</xref>
. All specimens were very similar in morphology. However, the dorsum of
<specimenCode id="E18A9238F00A477E580974A652C808AF">
<abbrev id="ABBRID0EBSAI" xlink_title="Moutai Institute">
<collectionCode id="998AD4F6BF93A522BA71E2E51A402C46" country="Russia" name="Mus. Tinro, Vladyvostok">MT</collectionCode>
</abbrev>
<collectionCode id="2B73E4A5BD27EEEA7A3E0FB03BD97913" country="United Kingdom" name="Linnean Society of London">LS</collectionCode>
20240517002
</specimenCode>
was brown, with a dark brown inversed Y-shaped marking on the dorsum (Fig.
<figureCitation id="3C10126A803C0F9BE0651515DD04E034" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Color variation in Gracixalus weii sp. nov. A. Dorsolateral view of the male specimen MT LS 20240517002; B. Dorsolateral view of the male specimen MT LS 20240518010. (Photo A, B by Shize Li)." figureDoi="10.3897/zse.101.133735.figure6" httpUri="https://binary.pensoft.net/fig/1258207">6 A</figureCitation>
); in
<specimenCode id="072803E008421F7F68BA1425A06D59E6">
<abbrev id="ABBRID0EJSAI" xlink_title="Moutai Institute">
<collectionCode id="B5B948CEE17B65E25D24F5542311ADA2" country="Russia" name="Mus. Tinro, Vladyvostok">MT</collectionCode>
</abbrev>
<collectionCode id="6FE76606CD74D25DC9EC168B1AA8A369" country="United Kingdom" name="Linnean Society of London">LS</collectionCode>
20240518010
</specimenCode>
, the dorsum and dorsal surfaces of the limbs were chocolate (Fig.
<figureCitation id="4781036FA31E95CA3BE8A63D43C798D1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Color variation in Gracixalus weii sp. nov. A. Dorsolateral view of the male specimen MT LS 20240517002; B. Dorsolateral view of the male specimen MT LS 20240518010. (Photo A, B by Shize Li)." figureDoi="10.3897/zse.101.133735.figure6" httpUri="https://binary.pensoft.net/fig/1258207">6 B</figureCitation>
).
</paragraph>
<caption id="C749C87D77A19FD068224AA6C447C4B2" ID-DOI="10.3897/zse.101.133735.figure6" ID-arpha="43E4F6ED-DE0B-5531-ACDD-874D0CB1A465" httpUri="https://binary.pensoft.net/fig/1258207" startId="F6">
<paragraph id="17D73F73E801C2D295D2576972F7FC4A">
<label id="3C0A0C60012C81A542FB2F1B70D1347D">Figure 6.</label>
</paragraph>
<paragraph id="0F4CC7E39681022581A5C7B4EC272CD8">
Color variation in
<taxonomicName id="4F2080C5B73A99D1006309C731D2B3B0" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="B6B6C3BBFBF5860D1696640EB6C37E2C" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AFDD2A23A595A2B1E72C8B02C57D5107" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="58937C556D16B1C786A41E60DBE9ABA0" bold="true">A.</emphasis>
Dorsolateral view of the male specimen
<specimenCode id="8FB669CD44923253507B6AD813EDBF2A">
<abbrev id="ABBRID0EGTAI" xlink_title="Moutai Institute">MT</abbrev>
LS 20240517002
</specimenCode>
;
<emphasis id="9F1E9A1C5C72EE1F9EFE6B88EA26C02E" bold="true">B.</emphasis>
Dorsolateral view of the male specimen
<specimenCode id="A1DCE9336E6D41B630D1E8F6484FC5ED">
<abbrev id="ABBRID0EMTAI" xlink_title="Moutai Institute">MT</abbrev>
LS 20240518010
</specimenCode>
. (Photo
<emphasis id="CFABAFA64DC8129357EDA806E074A50C" bold="true">A, B</emphasis>
by Shize Li).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EXTAI" type="description">
<paragraph id="5461F0E35F8EFC8F0A0B3DCEDF3F1615">
<heading id="52FE86B7C788A377E56DAD31F09E6B0A" reason="title">Secondary sexual characters.</heading>
</paragraph>
<paragraph id="5EFD582231302916188232472076309B">Adult females were found to be larger than adult males. The adult males showed a single subgular vocal sac. In breeding, the nuptial pad was present on the dorsal surface of the bases of the first finger.</paragraph>
</subSubSection>
<subSubSection id="SECID0E3TAI" type="diagnosis">
<paragraph id="FFB37E79F3B13BBA08A6C407356133AE">
<heading id="A06DBBA3A0C21C3E582217EBB07771D5" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="5FAD4ADE0B62F8005DE47132219E24A1">
<taxonomicName id="950A9A544DAA8F397C158D3746AA742C" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="32110D09EBF2554055FED99614ADB73A" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8F5D35D15FB98C2D7B38F7ABF6E843AC" rank="species">sp. nov.</taxonomicNameLabel>
was assigned to the genus
<taxonomicName id="2ECA7FE3E6D072B3FB30AF20E13AD519" authorityName="Delorme, Dubois, Grosjean &amp; Ohler" authorityYear="2005" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="8A90006219209831B897E1ABD5B8BE6D" italics="true">Gracixalus</emphasis>
</taxonomicName>
based on molecular data and morphological characteristics. Morphologically, the new species presented an intercalary cartilage between the terminal and penultimate phalanges of digits. The tips of the digits expanded into large disks with circummarginal grooves, the vomerine teeth were absent, and the pupil was horizontal (
<bibRefCitation id="4DE958A63759DB9A929B19D23DF14200" DOI="10.3406/linly.2005.13595" author="Delorme" etAl="et al." firstAuthor="Delorme" refId="B7" refString="Delorme M, Dubois A, Grosjean S, Ohler A (2005). Une nouvelle classification générique et subgénérique de la tribu des Philautini (Amphibia, Anura, Rhacophorinae). Bulletin Mensuel de la Société Linnéenne de Lyon 74: 165171. https://doi.org/10.3406/linly.2005.13595" year="2005">Delorme et al. 2005</bibRefCitation>
).
</paragraph>
<paragraph id="29EB70CF78F91F61FACC911B504A1FCE">
<taxonomicName id="3A0BB2B071D850592DAD3AAB2DFD579E" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="92EEAE8BE47862B1DBEE6A4D00BBA57D" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="495FF6478009A65B2586A5B0D0F94539" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from other congeners by the following characteristics: (1) moderate body size (
<abbrev id="ABBRID0EFVAI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm in males and 35.236.0 mm in females); (2) head width larger than head length; (3) distinct tympanum; (4) brown to beige dorsum, with an inversed Y-shaped dark brown marking; (5) rough skin of the dorsal and lateral surfaces of the head, body, and limbs, sparsely scattered with tubercles; (6) toes with moderately developed webbing, while finger webbing was rudimentary; (7) nuptial pads on base of finger I; (8) males had a single subgular vocal sac; (9) tibiotarsal articulation reaching the mid-eye when leg stretched forward; (10) heels overlapping when legs at right angle to body.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJVAI" type="Morphological comparisons">
<paragraph id="D60CBB5DCD9BFEBABE326D0E8F267AB8">
<heading id="4297DB965E82827BF70E6CE8878260F2" reason="title">Morphological comparisons.</heading>
</paragraph>
<paragraph id="E29C263C786FE5ACE5F1A5681A78A438">
A summary of morphological comparisons is present in Suppl. material
<xref id="E9D49FD441EA02063558C7EFC14FAF49" ref-type="supplementary-material" rid="S1">1</xref>
.
<taxonomicName id="A8F5D1527D633AD8ECE4BFFC3B817D8A" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="68FD11547C8D66630290085F00484F86" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2520E50605FB8949328CF4B6A94B0930" rank="species">sp. nov.</taxonomicNameLabel>
appears to be most closely related to
<taxonomicName id="D1FD7795C8C9805A753FD2EE5C042BD8" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jinxiuensis">
<emphasis id="78EFCC087427BAF2E5DD96C078E9A5F8" italics="true">G. jinxiuensis</emphasis>
</taxonomicName>
but differs from it by having a relatively larger size,
<abbrev id="ABBRID0EJWAI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm in adult males and 35.236.0 mm in adult females (vs.
<quantity id="E94AC832167F764843D0263F95EF8CF3" metricMagnitude="-2" metricUnit="m" metricValue="2.525" metricValueMax="2.63" metricValueMin="2.42" unit="mm" value="25.25" valueMax="26.3" valueMin="24.2">24.226.3 mm</quantity>
in males, 28.0
<quantity id="6B94635B2315EC9675E2D55FFDA23501" metricMagnitude="-2" metricUnit="m" metricValue="2.92" unit="mm" value="29.2">29.2 mm</quantity>
in females); head width larger than head length (vs. the length and width of the head are almost equal); sole of foot and palmar smooth with sparse small tubercles (vs. rough with dense large tubercles); heels overlapped when thighs are positioned at right angles to the body (vs. heels just meeting); tibiotarsal articulation reaching the level of mid-eye when leg stretched forward (vs. posterior angle of eye);
<taxonomicName id="B594BF266EA8AB0F5606732E4B2A9511" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="9135DC77B511EAA43951FE83169E1B47" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2941D8F0151863CF24A629C917FDD4BD" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="83518CB569EB02CC21D7606280124225" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jinxiuensis">
<emphasis id="E4C50E294BF70413B396B4798D035EAE" italics="true">G. jinxiuensis</emphasis>
</taxonomicName>
by having significantly lower ratios of
<abbrev id="ABBRID0EDXAI" xlink_title="eye diameter">ED</abbrev>
,
<abbrev id="ABBRID0EHXAI" xlink_title="interorbital distance">IOD</abbrev>
, and
<abbrev id="ABBRID0ELXAI" xlink_title="snout length">SL</abbrev>
to
<abbrev id="ABBRID0EPXAI" xlink_title="snout-vent length">SVL</abbrev>
in males and having significantly higher ratios of
<abbrev id="ABBRID0ETXAI" xlink_title="head length">HDL</abbrev>
,
<abbrev id="ABBRID0EXXAI" xlink_title="internasal distance">IND</abbrev>
,
<abbrev id="ABBRID0E2XAI" xlink_title="maximal tympanum diameter">TYD</abbrev>
, and
<abbrev id="ABBRID0E6XAI" xlink_title="length of foot and tarsus">TFL</abbrev>
to
<abbrev id="ABBRID0EDYAI" xlink_title="snout-vent length">SVL</abbrev>
in males (all p-values &lt;0.05; Table
<tableCitation id="DFB96ECE7F502874341C75A3E8914B66" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Measurements of the adult specimens of Gracixalus weii sp. nov. Comparison of male Gracixalus weii sp. nov. (PW) and Gracixalus jinxiuensis (PJ). Units are given in mm. P - values less than 0.05 indicate significant differences. See abbreviations for the morphological characters in the Materials and methods section." httpUri="http://table.plazi.org/id/D72E49A3145B422A40E1746AA6D01A01" tableUuid="D72E49A3145B422A40E1746AA6D01A01">3</tableCitation>
).
</paragraph>
<paragraph id="ABBF81A2D74B11E0C7ED8A560A697BFC">
<taxonomicName id="03410D3972037FF41340F1838E5A4506" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="6A05D9CCCC33A693191174C5C4DECCC6" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3F13174602914124C6357FD136EED506" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4674181B11E5B0CE958594E79637AECE" baseAuthorityName="Matsui &amp; Orlov" baseAuthorityYear="2004" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ananjevae">
<emphasis id="FE17949AD271FF580D1EFBCB24FE6759" italics="true">G. ananjevae</emphasis>
</taxonomicName>
by having a smaller body size in females
<abbrev id="ABBRID0EDZAI" xlink_title="snout-vent length">SVL</abbrev>
35.236.0 mm in adult females (vs.
<quantity id="41677F4A38F52D9450C22207A531B369" metricMagnitude="-2" metricUnit="m" metricValue="4.34" unit="mm" value="43.4">43.4 mm</quantity>
), with distinctive conical tubercles on dorsum (vs. absent) and skin of throat granular (vs. plain); from
<taxonomicName id="2C28AF31DEC031F57EC01D8061E19F5B" baseAuthorityName="Boulenger" baseAuthorityYear="1893" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="carinensis">
<emphasis id="6B7EDA9B352678CE2F0EF1A51265DC27" italics="true">G. carinensis</emphasis>
</taxonomicName>
by having distinctive conical tubercles on dorsum (vs. absent); from
<taxonomicName id="20CAC4028767B6B189594773B2C127E7" baseAuthorityName="Bourret" baseAuthorityYear="1937" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="gracilipes">
<emphasis id="5FF6BA7C8BA6461A348B876926C930BE" italics="true">G. gracilipes</emphasis>
</taxonomicName>
by having bigger body size both in males and females (
<abbrev id="ABBRID0E4ZAI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm in males and 35.236.0 mm in females vs. 20.024.0 mm in males and
<quantity id="7A1DA34241BF4D4BADE9B6E019334E49" metricMagnitude="-2" metricUnit="m" metricValue="2.76" metricValueMax="2.88" metricValueMin="2.6399999999999997" unit="mm" value="27.6" valueMax="28.8" valueMin="26.4">26.428.8 mm</quantity>
in females), distinctive conical tubercles present on dorsum (versus absent), dorsal surface brown (vs. greenish), tibiotarsal articulation reaching to mid-eye (vs. reaching to between eye and nostril) and throat granular (vs. smooth); from
<taxonomicName id="CDAD3624AC3AB47158CE6943E49EC33D" baseAuthorityName="Wang, Zeng, Liu &amp; Wang" baseAuthorityYear="2018" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="guangdongensis">
<emphasis id="C019AD001EFBB5D5C858F3489379B3DA" italics="true">G. guangdongensis</emphasis>
</taxonomicName>
by having distinctive conical tubercles on dorsum (vs. absent), finger webbing rudimentary (vs. absent) and linea masculine absent (vs. present); from
<taxonomicName id="6DA28982D950EAEE56A9AC5F5FDF0CF2" baseAuthorityName="Zeng, Zhao, Chen, Chen, Zhang &amp; Wang" baseAuthorityYear="2017" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="jinggangensis">
<emphasis id="4484719B066F1BEF2F17F851F61E62A1" italics="true">G. jinggangensis</emphasis>
</taxonomicName>
by having distinctive conical tubercles on dorsum (vs. absent) and nuptial pads present only on finger I (vs. nuptial pads present on fingers I and II); from
<taxonomicName id="E08E93EE3DCA313896D48B9E4A2EB6C3" baseAuthorityName="Rowley, Le, Dau, Hoang &amp; Cao" baseAuthorityYear="2014" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="lumarius">
<emphasis id="4CB59A909A6A26BE5B901CE2DB74F88D" italics="true">G. lumarius</emphasis>
</taxonomicName>
by having a smaller body size in males (
<abbrev id="ABBRID0EC2AI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm vs.
<quantity id="9F67860D8121967DEC9F4E65B74EAE76" metricMagnitude="-2" metricUnit="m" metricValue="4.025" metricValueMax="4.16" metricValueMin="3.8899999999999997" unit="mm" value="40.25" valueMax="41.6" valueMin="38.9">38.941.6 mm</quantity>
) and males with an internal vocal sac (vs. an external vocal sac); from
<taxonomicName id="C7C19FC000D37589495FBCD007E48188" baseAuthorityName="Ye &amp; Hu" baseAuthorityYear="1984" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="medogensis">
<emphasis id="4E65EB280FCCEBE6D67F85344E5864BC" italics="true">G. medogensis</emphasis>
</taxonomicName>
by having a bigger body size in males (
<abbrev id="ABBRID0ER2AI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm vs.
<quantity id="7548E0875AD0E6E33515060CC9887C01" metricMagnitude="-2" metricUnit="m" metricValue="2.65" unit="mm" value="26.5">26.5 mm</quantity>
), having distinctive conical tubercles on dorsum (vs. absent) and linea masculine absent (vs. present); from
<taxonomicName id="70A59ABFE2FF808C33CEC48588689D79" baseAuthorityName="Mo, Zhang, Luo, Zhou &amp; Chen" baseAuthorityYear="2013" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="nonggangensis">
<emphasis id="AB77D41534D38513300A201E1B48101B" italics="true">G. nonggangensis</emphasis>
</taxonomicName>
by having a bigger body size in females (
<abbrev id="ABBRID0EA3AI" xlink_title="snout-vent length">SVL</abbrev>
35.236.0 mm vs.
<quantity id="B6E67D31FC62F65AB045E6D22FFB35E9" metricMagnitude="-2" metricUnit="m" metricValue="2.705" metricValueMax="2.73" metricValueMin="2.68" unit="mm" value="27.05" valueMax="27.3" valueMin="26.8">26.827.3 mm</quantity>
), having distinctive conical tubercles on dorsum (vs. absent), finger webbing rudimentary (vs. absent), nuptial pads present on finger I (vs. nuptial pads absent) and tibiotarsal articulation reaching to mid-eye (vs. reaching to tip of snout); from
<taxonomicName id="39DAB0FEC2F0FCF9D559934ED22FC620" baseAuthorityName="Boruah, Deepak, Patel, Jithin, Yomcha &amp; Das" baseAuthorityYear="2023" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="patkaiensis">
<emphasis id="8EB189BA07A835954FAD6D16CF4CC6F3" italics="true">G. patkaiensis</emphasis>
</taxonomicName>
by having bigger body size in males (
<abbrev id="ABBRID0EP3AI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm vs.
<quantity id="61169D851986E28F93EFA0FFC8F7F45B" metricMagnitude="-2" metricUnit="m" metricValue="2.505" metricValueMax="2.65" metricValueMin="2.36" unit="mm" value="25.05" valueMax="26.5" valueMin="23.6">23.626.5 mm</quantity>
), finger webbing rudimentary (vs. absent), males with an internal vocal sac (vs. an external vocal sac) and tibiotarsal articulation reaching to mid-eye (vs. reaching to tip of snout); from
<taxonomicName id="11986F9A10D3B5B71E3DC56B97D491E1" baseAuthorityName="Rowley, Dau, Nguyen, Cao &amp; Nguyen" baseAuthorityYear="2011" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="quangi">
<emphasis id="7A46D92D6D96333792CDF1EB06317E0B" italics="true">G. quangi</emphasis>
</taxonomicName>
by having bigger body size both in males and females (
<abbrev id="ABBRID0E53AI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm in adult males and 35.236.0 mm in adult females vs. 21.024.0 mm in males and
<quantity id="62BA90D38D63A490646B33C0662FA959" metricMagnitude="-2" metricUnit="m" metricValue="2.705" metricValueMax="2.73" metricValueMin="2.68" unit="mm" value="27.05" valueMax="27.3" valueMin="26.8">26.827.3 mm</quantity>
in females), finger webbing rudimentary (vs. absent), males with an internal vocal sac (vs. an external vocal sac) and skin of throat granular (vs. absent); from
<taxonomicName id="F845C7C9C0C3277B15151FA4A193C638" baseAuthorityName="Rowley, Dau, Nguyen, Cao &amp; Nguyen" baseAuthorityYear="2011" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="quangi">
<emphasis id="7EA05D285728FEAAEF1DBBF165EC0D0D" italics="true">G. quangi</emphasis>
</taxonomicName>
by having bigger body size in males (
<abbrev id="ABBRID0EN4AI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm vs.
<quantity id="7E8563758F0A9DE8B590BF75EAB7E464" metricMagnitude="-2" metricUnit="m" metricValue="2.85" unit="mm" value="28.5">28.5 mm</quantity>
), skin of throat granular (vs. absent) and tibiotarsal articulation reaching to mid-eye (vs. reaching to snout); from
<taxonomicName id="9433EFD74AC5B4B8174F21B783EE5BD7" baseAuthorityName="Matsui, Ohler, Eto &amp; Nguyen" baseAuthorityYear="2017" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="sapaensis">
<emphasis id="7649512154C7590A81A99B636C10337F" italics="true">G. sapaensis</emphasis>
</taxonomicName>
by having bigger body size in males (
<abbrev id="ABBRID0E34AI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm vs.
<quantity id="924F4CCFE2D44C48A27E9B586D26D980" metricMagnitude="-2" metricUnit="m" metricValue="2.52" metricValueMax="2.96" metricValueMin="2.08" unit="mm" value="25.2" valueMax="29.6" valueMin="20.8">20.829.6 mm</quantity>
) and with distinctive conical tubercles on dorsum (vs. absent); from
<taxonomicName id="0B163411B5A3CBDB257CCEC9149B83F5" baseAuthorityName="Matsui, Khonsue, Panha &amp; Eto" baseAuthorityYear="2015" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="seesom">
<emphasis id="978C43A93A659DB43D532E3232F89F09" italics="true">G. seesom</emphasis>
</taxonomicName>
by having bigger body size both in males and females (
<abbrev id="ABBRID0EL5AI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm in males and 35.236.0 mm in females vs. 21.623.0 mm in males and
<quantity id="880AA1CCBC7D80CDBC7EB41A7F8E05C9" metricMagnitude="-2" metricUnit="m" metricValue="2.4299999999999997" metricValueMax="2.54" metricValueMin="2.32" unit="mm" value="24.299999999999997" valueMax="25.4" valueMin="23.2">23.225.4 mm</quantity>
in females), with distinctive conical tubercles on dorsum (vs. absent), males with an internal vocal sac (vs. an external vocal sac) and tibiotarsal articulation reaching to mid-eye (vs. reaching between eye and nostril); from
<taxonomicName id="20615B13B89D2694D78005E5F770264E" baseAuthorityName="Orlov, Ho &amp; Nguyen" baseAuthorityYear="2004" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="supercornutus">
<emphasis id="7E0FFDE40FE04887315E2D6DD90FB7EA" italics="true">G. supercornutus</emphasis>
</taxonomicName>
by having bigger body size in males (
<abbrev id="ABBRID0E15AI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm vs. 22.0
<quantity id="750F2111F4BAF2F5EF4F7D03AE8141DF" metricMagnitude="-2" metricUnit="m" metricValue="2.41" unit="mm" value="24.1">24.1 mm</quantity>
); from
<taxonomicName id="B5877FC09B22B1593D04BE5BC8A1BDF2" baseAuthorityName="Chen, Bei, Liao, Zhou &amp; Mo" baseAuthorityYear="2018" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="tianlinensis">
<emphasis id="7D64EAE18DD7475FCE26C4AC4CB4D363" italics="true">G. tianlinensis</emphasis>
</taxonomicName>
by having distinctive conical tubercles on dorsum (vs. absent), males with an internal vocal sac (vs. an external vocal sac) and nuptial pads present only on finger I (vs. nuptial pads present on fingers I and II); from
<taxonomicName id="80FB0518452FFA9B8EB964E28CB7C5AA" baseAuthorityName="Rowley, Le, Hoang, Cao &amp; Dau" baseAuthorityYear="2020" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="trieng">
<emphasis id="CDA082728A5D6A07E9E8783AE42C7C9E" italics="true">G. trieng</emphasis>
</taxonomicName>
by having smaller body size in males (
<abbrev id="ABBRID0EU6AI" xlink_title="snout-vent length">SVL</abbrev>
30.134.0 mm vs.
<quantity id="FA678E204B057F14931D71C1A78DBDC8" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" metricValueMax="4.14" metricValueMin="3.72" unit="mm" value="39.3" valueMax="41.4" valueMin="37.2">37.241.4 mm</quantity>
), finger webbing rudimentary (vs. absent) and males with an internal vocal sac (vs. an external vocal sac); from
<taxonomicName id="F29CB5C546C068C5DE72DAED37C93498" baseAuthorityName="Tran, Pham, Le, Nguyen, Ziegler &amp; Pham" baseAuthorityYear="2023" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="truongi">
<emphasis id="751FBFA6236D2DAF49087445A51D7F7A" italics="true">G. truongi</emphasis>
</taxonomicName>
by having smaller body size in females (
<abbrev id="ABBRID0EDABI" xlink_title="snout-vent length">SVL</abbrev>
35.236.0 mm vs.
<quantity id="F6B431E509FC8804E4513977A76819A0" metricMagnitude="-2" metricUnit="m" metricValue="3.845" metricValueMax="3.9299999999999997" metricValueMin="3.76" unit="mm" value="38.45" valueMax="39.3" valueMin="37.6">37.639.3 mm</quantity>
), finger webbing rudimentary (vs. absent), having distinctive conical tubercles on dorsum (vs. absent) and tibiotarsal articulation reaching to mid-eye (vs. reaching between eye and nostril); from
<taxonomicName id="261DD0C6104DD5B9736A270F995ABE8F" baseAuthorityName="Yu, Li, Wang, Rao, Wu &amp; Yang" baseAuthorityYear="2019" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="5346E35A152C0573E9CD0B826C06482F" italics="true">G. yunnanensis</emphasis>
</taxonomicName>
by males with an internal vocal sac (vs. an external vocal sac); freom
<taxonomicName id="C54D2EF724D93F2C457A060A32FC72AF" baseAuthorityName="Le, Do, Tran, Nguyen, Orlov, Ninh &amp; Nguyen" baseAuthorityYear="2021" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="0E8B283B4C78AA8BF24340345060991B" italics="true">G. ziegleri</emphasis>
</taxonomicName>
by tibiotarsal articulation reaching to mid-eye (vs. reaching tip of snout).
</paragraph>
</subSubSection>
<subSubSection id="SECID0E4ABI" type="distribution">
<paragraph id="1419DDCB41631427FB7688D5EB93C904">
<heading id="8E3029114C5EB280B80F0320C87F05DA" reason="title">Distribution and habitats.</heading>
</paragraph>
<paragraph id="7CB973C6B20B6427110C6594BB30D6F0">
At present,
<taxonomicName id="91D9D5AC19C87D7C96BC075749EDE87B" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="AC4B03B5EBBE532C8626C796AB60D6B8" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65083D0FE2996378FBE68E988BB5A8E2" rank="species">sp. nov.</taxonomicNameLabel>
is only found in Leigongshan Nature Reserve, Leishan County,
<collectingRegion id="C6A24EA87D74649076B51B4B0DD88DFE" country="China" name="Guizhou">Guizhou Province</collectingRegion>
,
<collectingCountry id="A226956003B3673B7E4BE916D423A9E0" name="China">China</collectingCountry>
. The population inhabits the mountain forests at elevations between 1600 and
<quantity id="CC7CF70FFCE8E46F60B7893F6760DFD6" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</quantity>
and is often found on bamboo near streams (Fig.
<figureCitation id="33778B73DF276693ECD8073A9B9F2FA9" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitat of Gracixalus weii sp. nov. in the type locality, Leishan County, Guizhou Province, China. (Photo by Jing Liu)." figureDoi="10.3897/zse.101.133735.figure7" httpUri="https://binary.pensoft.net/fig/1258208">7</figureCitation>
).
<taxonomicName id="0F1A6C088496DB3A45745589C5E8A1CD" authority="Li, Xu, Liu, Jiang, Wei &amp; Wang, 2018" authorityName="Li, Xu, Liu, Jiang, Wei &amp; Wang" authorityYear="2018" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="leishanensis">
<emphasis id="09AF459D34FF7A867A60FD72AC687E5D" italics="true">Boulenophrys leishanensis</emphasis>
<bibRefCitation id="4CAED62B89AAC73FB0EC63EB942B36A5" DOI="10.16373/j.cnki.ahr.180072" author="Li SZ &amp; Xu N &amp; Liu J &amp; Jiang JP &amp; Wei G &amp; Wang B" issue="4" journalOrPublisher="Asian Herpetological Research" pagination="224-239" refId="B21" refString="Li SZ, Xu N, Liu J, Jiang JP, Wei G, Wang B (2018) A new species of the Asian toad genus Megophrys sensu lato (Amphibia, Anura, Megophryidae) from Guizhou Province, China. Asian Herpetological Research 9 (4): 224-239. https://doi.org/10.16373/j.cnki.ahr.180072" title="A new species of the Asian toad genus Megophrys sensu lato (Amphibia, Anura, Megophryidae) from Guizhou Province, China." volume="9" year="2018">Li, Xu, Liu, Jiang, Wei &amp; Wang, 2018</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="11B7DE09303B99D26E9D6357F09C3AE8" authority="Liu &amp; Hu, 1973" authorityName="Liu &amp; Hu" authorityYear="1973" class="Amphibia" family="Megophryidae" genus="Boulenophrys" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="spinata">
<emphasis id="FE4E964D40CDF3943FCAEA7816E008E2" italics="true">B. spinata</emphasis>
Liu &amp; Hu, 1973
</taxonomicName>
;
<taxonomicName id="9A391F886C689B13851624BE2DDE17B6" authority="Qian, Xia, Cao, Xiao &amp; Yang, 2020" authorityName="Qian, Xia, Cao, Xiao &amp; Yang" authorityYear="2020" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="wulingensis">
<emphasis id="318EF0D38B6CEF65D3FC9B5C75507D57" italics="true">Leptobrachella wulingensis</emphasis>
Qian, Xia, Cao, Xiao &amp; Yang, 2020
</taxonomicName>
; and
<taxonomicName id="7BFA0AE0303611B764A5B3D3710CA19F" authority="Liu &amp; Hu, 1973" authorityName="Liu &amp; Hu" authorityYear="1973" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="leishanensis">
<emphasis id="4ACAD23A35675EC5410A2608DF7F33C9" italics="true">Leptobrachium leishanensis</emphasis>
Liu &amp; Hu, 1973
</taxonomicName>
were also found in the type locality of the new species. After investigation, we only heard the calls of
<taxonomicName id="407BAA75D6317C34E42490D1414B2B05" authorityName="Delorme, Dubois, Grosjean &amp; Ohler" authorityYear="2005" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="genus">
<emphasis id="AA39EFB44D8F84009BDB98613287037B" italics="true">Gracixalus</emphasis>
</taxonomicName>
in mid to late May, and no calls were heard in other seasons, so we speculate that the breeding season may be in mid to late May.
</paragraph>
<caption id="07C90BB394FE2B65FADE3DBE12E3C6E6" ID-DOI="10.3897/zse.101.133735.figure7" ID-arpha="C5D8A96C-9DCA-51A0-BC60-F52E4C1C639D" httpUri="https://binary.pensoft.net/fig/1258208" startId="F7">
<paragraph id="EAA173F940358480397902F176DACD1C">
<label id="50E7DE028F487E9B957E80220E23D1E4">Figure 7.</label>
</paragraph>
<paragraph id="50A18D12206C7727DE9AE57B32BB9B49">
Habitat of
<taxonomicName id="7785386E1DDC5D0EF37112540D9E10F7" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii" status="sp. nov.">
<emphasis id="6D4485A0732B25B0CDAEBFF0CFE814FB" italics="true">Gracixalus weii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B548EF008FAF58EF5E830FFBE5D43144" rank="species">sp. nov.</taxonomicNameLabel>
in the type locality, Leishan County, Guizhou Province, China. (Photo by Jing Liu).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E3DBI" type="etymology">
<paragraph id="C6D1B282E94C6768CBF942E1A888A456">
<heading id="84C888E991FE9FD6CB37983F9738E323" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="E08480DFA84995BA3966C42465C6D945">
The specific epithet
<taxonomicName id="C29C3E3A7DD0747BAD23B8DF6F7054B5" authorityName="Liu &amp; Peng &amp; Wang &amp; Feng &amp; Shen &amp; Li &amp; Chen &amp; Su &amp; Tang" authorityYear="2025" class="Amphibia" family="Rhacophoridae" genus="Gracixalus" kingdom="Animalia" order="Anura" phylum="Chordata" rank="species" species="weii">
<emphasis id="E923F3F78D4A436AA7567927DAB2A247" italics="true">weii</emphasis>
</taxonomicName>
” was given in honor of Professor Gang Wei from Guiyang University, in recognition of his contributions to the systematic studies and biodiversity conservation of amphibians in
<collectingCountry id="FD12319E343EB42292D395A2BEB76AFD" name="China">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>