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<document id="B72DE87A34251E506E1451DC56DDEFF0" ID-DOI="10.17111/FragmPalHung.2016.33.3" ID-ISSN="1586-930X" ID-Zenodo-Dep="15627163" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1749503245431" checkinUser="felipe" docAuthor="Szabó, János" docDate="2016" docId="A63287CD6F60FF92CB92E8008A0FFE02" docLanguage="en" docName="FragPalaeontolHung.33.3-30.pdf" docOrigin="Fragmenta Palaeontologica Hungarica 33" docSource="https://doi.org/10.17111/fragmpalhung.2016.33.3" docStyle="DocumentStyle:899F05ADE4C3962F5AA48A6CAFB498A0.1:FragPalaeontolHung.2013-2019.journal_article.type3" docStyleId="899F05ADE4C3962F5AA48A6CAFB498A0" docStyleName="FragPalaeontolHung.2013-2019.journal_article.type3" docStyleVersion="1" docTitle="Cyclostomaria praemonarii Szabó, 2016, n. sp." docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="5A0BFFB56F77FF8BCA67EB4F8B36FFD6" masterDocTitle="Gastropods of the Lower Jurassic Hierlatz Limestone Formation, part 2. Some new archaic type slit-bearing components from the fauna of the Hierlatz Alpe (Hallstatt, Austria) and the Bakony Mts (Hungary)" masterLastPageNumber="30" masterPageNumber="3" pageNumber="26" updateTime="1750156583739" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5BFCE14A2FB70C4B441BF81FF2643733">Gastropods of the Lower Jurassic Hierlatz Limestone Formation, part 2. Some new archaic type slit-bearing components from the fauna of the Hierlatz Alpe (Hallstatt, Austria) and the Bakony Mts (Hungary)</mods:title>
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<mods:namePart id="79A62B17E1AB7A4C81646A4DC4C262D5">Szabó, János</mods:namePart>
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<mods:title id="D3D337C6BD3327CFA7BA168008831021">Fragmenta Palaeontologica Hungarica</mods:title>
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<mods:part id="982BD572593675A9B43D8D2E38BA624E">
<mods:date id="BACD2006233B62109CDEA1E2BC699B82">2016</mods:date>
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<mods:number id="06797B5040A23000BC94A5BA60E4DC53">2016-12-31</mods:number>
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<mods:number id="D04F9B290BD2D7576B34BEC6AF8DC6CF">33</mods:number>
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<mods:url id="D79FB0391772E00E155EA489C54E6D6E">https://doi.org/10.17111/fragmpalhung.2016.33.3</mods:url>
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<mods:classification id="B6B91C655E30ABCFB5BF6BCDC3A92D97">journal article</mods:classification>
<mods:identifier id="C1ED0FAABADDF913B59E116447679939" type="DOI">10.17111/FragmPalHung.2016.33.3</mods:identifier>
<mods:identifier id="8ED888D0A6E0882BA698C61F01E7062D" type="ISSN">1586-930X</mods:identifier>
<mods:identifier id="D36B85E72A6465F804BFCC5DAE3010A5" type="Zenodo-Dep">15627163</mods:identifier>
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<treatment id="A63287CD6F60FF92CB92E8008A0FFE02" LSID="urn:lsid:plazi:treatment:A63287CD6F60FF92CB92E8008A0FFE02" httpUri="http://treatment.plazi.org/id/A63287CD6F60FF92CB92E8008A0FFE02" lastPageId="25" lastPageNumber="28" pageId="23" pageNumber="26">
<subSubSection id="668165506F60FF9CCB92E800885AFCBC" box="[501,876,847,874]" pageId="23" pageNumber="26" type="nomenclature">
<paragraph id="2E2436DB6F60FF9CCB92E800885AFCBC" blockId="23.[501,876,847,909]" box="[501,876,847,874]" pageId="23" pageNumber="26">
<heading id="756C81B76F60FF9CCB92E800885AFCBC" box="[501,876,847,874]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="26" reason="9">
<taxonomicName id="E99B4D586F60FF9CCB92E800881FFCBF" authorityName="Szabó" authorityYear="2016" box="[501,809,847,873]" class="Gastropoda" family="Pleurotomariidae" genus="Cyclostomaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="23" pageNumber="26" phylum="Mollusca" rank="species" species="praemonarii" status="sp. nov.">
<emphasis id="1CEFEAC96F60FF9CCB92E800881FFCBF" box="[501,809,847,873]" italics="true" pageId="23" pageNumber="26">Cyclostomaria praemonarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F60FF9CC948E81F885AFCBC" box="[815,876,848,874]" pageId="23" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="668165506F60FF92C801E83D8A0FFE02" lastPageId="25" lastPageNumber="28" pageId="23" pageNumber="26" type="description">
<paragraph id="2E2436DB6F60FF9CC801E83D89CCFC5B" blockId="23.[501,876,847,909]" box="[614,762,882,909]" pageId="23" pageNumber="26">
(
<figureCitation id="B6A02A5E6F60FF9CC809E83D89C5FC5B" box="[622,755,882,909]" captionStart="Figs 37–40" captionStartId="24.[175,216,1541,1563]" captionTargetBox="[174,1111,1089,1514]" captionTargetId="figure-306@24.[174,1112,1089,1514]" captionTargetPageId="24" captionText="Figs 37–40. Holotype of Cyclostomaria praemonarii n. sp. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria, GBA 2017/001/0001.– 37–38. Magnified apical and lateral views, ×2. – 39–40. The same views without magnification" figureDoi="http://doi.org/10.5281/zenodo.15627171" httpUri="https://zenodo.org/record/15627171/files/figure.png" pageId="23" pageNumber="26">Figs 37–40</figureCitation>
)
</paragraph>
<materialsCitation id="9EF33C866F60FF9CCB71E8F5887FFC21" country="Austria" location="Hierlatz Alpe" municipality="Hallstatt" pageId="23" pageNumber="26" specimenCount="1" typeStatus="holotype">
<paragraph id="2E2436DB6F60FF9CCB71E8F589E3FC05" blockId="23.[219,1157,953,1648]" box="[278,725,953,979]" pageId="23" pageNumber="26">
<emphasis id="1CEFEAC96F60FF9CCB71E8F58A8CFC05" box="[278,442,954,979]" italics="true" pageId="23" pageNumber="26">
<typeStatus id="F12088796F60FF9CCB71E8F58A78FC05" box="[278,334,954,979]" pageId="23" pageNumber="26">Type</typeStatus>
specimen
</emphasis>
– GBA 2017/001/0001.
</paragraph>
<paragraph id="2E2436DB6F60FF9CCB71E8928878FC21" blockId="23.[219,1157,953,1648]" box="[278,846,988,1015]" pageId="23" pageNumber="26">
<emphasis id="1CEFEAC96F60FF9CCB71E8928A91FC20" box="[278,423,988,1014]" italics="true" pageId="23" pageNumber="26">
<typeStatus id="F12088796F60FF9CCB71E8928A78FC20" box="[278,334,989,1014]" pageId="23" pageNumber="26">Type</typeStatus>
locality
</emphasis>
–
<location id="2B4460006F60FF9CCBA4E8938952FC20" LSID="urn:lsid:plazi:treatment:A63287CD6F60FF92CB92E8008A0FFE02:2B4460006F60FF9CCBA4E8938952FC20" box="[451,612,988,1014]" country="Austria" municipality="Hallstatt" name="Hierlatz Alpe" pageId="23" pageNumber="26">Hierlatz Alpe</location>
,
<collectingMunicipality id="CE40ACA16F60FF9CC809E89389E3FC20" box="[622,725,988,1014]" pageId="23" pageNumber="26">Hallstatt</collectingMunicipality>
(
<collectingCountry id="568C764B6F60FF9CC883E8938877FC21" box="[740,833,988,1015]" name="Austria" pageId="23" pageNumber="26">Austria</collectingCountry>
).
</paragraph>
</materialsCitation>
<paragraph id="2E2436DB6F60FF9CCB71EF4F8950FBEA" blockId="23.[219,1157,953,1648]" pageId="23" pageNumber="26">
<materialsCitation id="9EF33C866F60FF9CCB71EF4F8954FBEA" county="Lower Jurassic" location="Hierlatz Limestone" municipality="Pliensbachian" pageId="23" pageNumber="26" specimenCount="1" typeStatus="holotype">
<emphasis id="1CEFEAC96F60FF9CCB71EF4F8AA0FBCF" box="[278,406,1024,1049]" italics="true" pageId="23" pageNumber="26">
<typeStatus id="F12088796F60FF9CCB71EF4F8A79FBCF" box="[278,335,1024,1049]" pageId="23" pageNumber="26">Type</typeStatus>
strata
</emphasis>
–
<location id="2B4460006F60FF9CCBD5E8B089A3FBCF" LSID="urn:lsid:plazi:treatment:A63287CD6F60FF92CB92E8008A0FFE02:2B4460006F60FF9CCBD5E8B089A3FBCF" box="[434,661,1023,1049]" county="Lower Jurassic" municipality="Pliensbachian" name="Hierlatz Limestone" pageId="23" pageNumber="26">Hierlatz Limestone</location>
from the
<location id="2B4460006F60FF9CC960E8B088B9FBCF" LSID="urn:lsid:plazi:treatment:A63287CD6F60FF92CB92E8008A0FFE02:2B4460006F60FF9CC960E8B088B9FBCF" box="[775,911,1023,1049]" county="Lower Jurassic" municipality="Pliensbachian" name="Sinemurian" pageId="23" pageNumber="26">Sinemurian</location>
to lowermost
<collectingMunicipality id="CE40ACA16F60FF9CCE53E8B08A0EFBEA" pageId="23" pageNumber="26">Pliensbachian</collectingMunicipality>
(
<collectingCounty id="C7454E576F60FF9CCB21EF6C8AC3FBEB" box="[326,501,1058,1085]" pageId="23" pageNumber="26">Lower Jurassic</collectingCounty>
) interval
</materialsCitation>
.
</paragraph>
<paragraph id="2E2436DB6F60FF9CCB71EF098889FB55" blockId="23.[219,1157,953,1648]" pageId="23" pageNumber="26">
<emphasis id="1CEFEAC96F60FF9CCB71EF098AC5FB89" box="[278,499,1094,1120]" italics="true" pageId="23" pageNumber="26">Derivation of name</emphasis>
– Referring to the stratigraphically younger [Davoei to Margaritatus Zone (Pliensbachian), Lower Jurassic]
<taxonomicName id="E99B4D586F60FF9CC95AEF26888EFB55" authorityName="Szabo" authorityYear="2008" box="[829,952,1129,1155]" class="Gastropoda" family="Pleurotomariidae" genus="Cyclostomaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="23" pageNumber="26" phylum="Mollusca" rank="species" species="monarii">
<emphasis id="1CEFEAC96F60FF9CC95AEF268866FB55" box="[829,848,1129,1155]" italics="true" pageId="23" pageNumber="26">C</emphasis>
.
<emphasis id="1CEFEAC96F60FF9CC93AEF25888EFB55" box="[861,952,1130,1155]" italics="true" pageId="23" pageNumber="26">monarii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="2E2436DB6F60FF9CCB71EFC28824FA16" blockId="23.[219,1157,953,1648]" pageId="23" pageNumber="26">
<emphasis id="1CEFEAC96F60FF9CCB71EFC28AB5FB70" box="[278,387,1165,1190]" italics="true" pageId="23" pageNumber="26">Diagnosis</emphasis>
– Low spired shell, consisting of whorls with rounded cross-section, two angles at spire and umbilical suture lines, respectively. Top of convexity on whorls in higher position than suture, convex ramp connecting them. Suture running at periphery of former whorl. Base low with convex wall and broad umbilicus. Selenizone between two cords, concave, rather wide and placed at outer face little below top line of whorl. Subregularly repeating, collabral riblets on ramp, crossed by single cord along top of whorl; nodules at intersections. Dense spiral threads between selenizone and periphery; base seemingly smooth. Selenizone without lunulae and spiral ornament.
</paragraph>
<paragraph id="2E2436DB6F60FF9CCB71EE868ADBF9D0" blockId="23.[219,1157,953,1648]" pageId="23" pageNumber="26">
<emphasis id="1CEFEAC96F60FF9CCB71EE868A4DFA35" box="[278,379,1481,1507]" italics="true" pageId="23" pageNumber="26">Material</emphasis>
– A worn, shelly whorl-fragment, surrounding also cross-section of two previous whorls.
</paragraph>
<paragraph id="2E2436DB6F60FF9CCB71ED5F8F16F9FC" blockId="23.[219,1157,953,1648]" box="[278,1056,1551,1578]" pageId="23" pageNumber="26">
<emphasis id="1CEFEAC96F60FF9CCB71ED5F8AAFF9FF" box="[278,409,1552,1577]" italics="true" pageId="23" pageNumber="26">Dimensions</emphasis>
– Height of preserved whorl:
<quantity id="E9639B3E6F60FF9CC89EED408865F9FC" box="[761,851,1551,1578]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="23" pageNumber="26" unit="mm" value="9.5">9.5 mm</quantity>
, width:
<quantity id="E9639B3E6F60FF9CC9D5ED408F2AF9FC" box="[946,1052,1551,1578]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="23" pageNumber="26" unit="mm" value="21.5">21.5 mm</quantity>
.
</paragraph>
<paragraph id="2E2436DB6F60FF93CB71ED7C8ACCFE67" blockId="23.[219,1157,953,1648]" lastBlockId="24.[174,1112,266,1067]" lastPageId="24" lastPageNumber="27" pageId="23" pageNumber="26">
<emphasis id="1CEFEAC96F60FF9CCB71ED7C8AA1F99A" box="[278,407,1587,1612]" italics="true" pageId="23" pageNumber="26">Description</emphasis>
– The remnants indicate a low, almost planispiral shell that has axially round whorl cross-sections; scar of the broken former whorls shows presence of a rather wide umbilicus. A concave selenizone runs well above the midwhorl but somewhat below the most adapical line of the whorl; the belt between the selenizone and the impressed (upper) suture is rather wide and mostly convex but a narrow concave band accompanies the suture. The suture follows the periphery of the former whorl.
</paragraph>
<paragraph id="2E2436DB6F6FFF93CA8EEAF58887FC17" blockId="24.[174,1112,266,1067]" pageId="24" pageNumber="27">The ornament is poorly preserved but the coarser elements are observable. Subregularly repeated, unequally strong riblets are visible in the belt between the selenizone and the adapical suture. Their shape is almost straight in slightly prosocline orientation with a short prosocyrt part along the selenizone. Weaker riblets or thick growth lines seem to be developed also on the outer whorl face occasionally. Two cords, the strongest spiral ornamental elements, limit the selenizone. At the first third of the distance toward the adapical suture from the selenizone, single marked spiral thread is visible and two weak threads are also present between the aforesaid one and the cord along the selenizone. Between the strongest spiral thread on the ramp and the selenizone, the collabral riblets weaken; nodules are at crossings with the riblets. The outer face is covered by dense spiral threads of unequal strength, crossed only by strongly prosocyrt growth-lines, which are visible only in few spots. No lunulae are observable in the selenizone; the probably delicate growth-lines are not observable on the available specimen. Preserved parts of the outermost base zone are smooth.</paragraph>
<paragraph id="2E2436DB6F6FFF92CA8EE88588B5FEBD" blockId="24.[174,1112,266,1067]" lastBlockId="25.[219,1156,266,468]" lastPageId="25" lastPageNumber="28" pageId="24" pageNumber="27">
<emphasis id="1CEFEAC96F6FFF93CA8EE8858A7BFC32" box="[233,333,970,996]" italics="true" pageId="24" pageNumber="27">Remarks</emphasis>
– The available shelly fragment well agrees with the penultimate whorl of the
<typeStatus id="F12088796F6FFF93CB34E8A18AB1FBDE" box="[339,391,1006,1032]" pageId="24" pageNumber="27">type</typeStatus>
specimens of
<taxonomicName id="E99B4D586F6FFF93C85DE8A28875FBD1" authorityName="Szabo" authorityYear="2008" box="[570,835,1005,1031]" class="Gastropoda" family="Pleurotomariidae" genus="Cyclostomaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="24" pageNumber="27" phylum="Mollusca" rank="species" species="monarii">
<emphasis id="1CEFEAC96F6FFF93C85DE8A28875FBD1" box="[570,835,1005,1031]" italics="true" pageId="24" pageNumber="27">Cyclostomaria monarii</emphasis>
</taxonomicName>
in its dimensions and shape. However, the almost full absence of the spiral ornament on the adapical part of the whorl means a significant difference that distinguish this specimen from
<taxonomicName id="E99B4D586F6EFF92CB7CEA618AA3FE91" authorityName="Szabo" authorityYear="2008" box="[283,405,302,328]" class="Gastropoda" family="Pleurotomariidae" genus="Cyclostomaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="25" pageNumber="28" phylum="Mollusca" rank="species" species="monarii">
<emphasis id="1CEFEAC96F6EFF92CB7CEA618A18FE9E" box="[283,302,302,328]" italics="true" pageId="25" pageNumber="28">C</emphasis>
.
<emphasis id="1CEFEAC96F6EFF92CB5DEA618AA3FE91" box="[314,405,302,327]" italics="true" pageId="25" pageNumber="28">monarii</emphasis>
</taxonomicName>
, which has a dense, subregular network ornament adapically and abapically from the selenizone at corresponding shell size.
</paragraph>
<caption id="7AE466536F6FFF93CAC8ED4A883DF9A7" ID-DOI="http://doi.org/10.5281/zenodo.15627171" ID-Zenodo-Dep="15627171" httpUri="https://zenodo.org/record/15627171/files/figure.png" pageId="24" pageNumber="27" startId="24.[175,216,1541,1563]" targetBox="[174,1111,1089,1514]" targetPageId="24" targetType="figure">
<paragraph id="2E2436DB6F6FFF93CAC8ED4A883DF9A7" blockId="24.[174,1112,1541,1649]" pageId="24" pageNumber="27">
<emphasis id="1CEFEAC96F6FFF93CAC8ED4A8A1EF9CC" bold="true" box="[175,296,1541,1563]" pageId="24" pageNumber="27">Figs 37–40.</emphasis>
Holotype of
<taxonomicName id="E99B4D586F6FFF93CBD5ED4A8984F9CC" authorityName="Szabó" authorityYear="2016" box="[434,690,1541,1562]" class="Gastropoda" family="Pleurotomariidae" genus="Cyclostomaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Pleurotomariida" pageId="24" pageNumber="27" phylum="Mollusca" rank="species" species="praemonarii" status="sp. nov.">
<emphasis id="1CEFEAC96F6FFF93CBD5ED4A8984F9CC" box="[434,690,1541,1562]" italics="true" pageId="24" pageNumber="27">Cyclostomaria praemonarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F6FFF93C8DBED4A89C5F9CC" box="[700,755,1541,1562]" pageId="24" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria, GBA 2017/001/0001. –
<emphasis id="1CEFEAC96F6FFF93CB82ED71891AF985" bold="true" box="[485,556,1598,1619]" pageId="24" pageNumber="27">37–38.</emphasis>
Magnified apical and lateral views, ×2. –
<emphasis id="1CEFEAC96F6FFF93C9D4ED7188CFF985" bold="true" box="[947,1017,1598,1619]" pageId="24" pageNumber="27">39–40.</emphasis>
The same views without magnification
</paragraph>
</caption>
<paragraph id="2E2436DB6F6EFF92CB71EA3B8A0FFE02" blockId="25.[219,1156,266,468]" pageId="25" pageNumber="28">
<emphasis id="1CEFEAC96F6EFF92CB71EA3B8A92FE58" box="[278,420,372,398]" italics="true" pageId="25" pageNumber="28">Distribution</emphasis>
– Within the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone Formation in Hierlatz Alpe, Hallstatt,
<collectingCountry id="568C764B6F6EFF92CABCEAF58A02FE02" box="[219,308,442,468]" name="Austria" pageId="25" pageNumber="28">Austria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

74
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Genus
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<typeStatus id="F12088796F60FF9CCB4DE9B78A50FCC4" box="[298,358,760,786]" pageId="23" pageNumber="26">Type</typeStatus>
species –
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<emphasis id="1CEFEAC96F60FF9CCBBCE9B889E9FCC7" box="[475,735,759,785]" italics="true" pageId="23" pageNumber="26">Cyclostomaria monarii</emphasis>
Szabó, 2008
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misidentified as
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<emphasis id="1CEFEAC96F60FF9CCB47E8558927FCE2" box="[288,529,794,820]" italics="true" pageId="23" pageNumber="26">Pleurotomaria suessii</emphasis>
Hörnes,
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in the original genus designation.
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(
<figureCitation id="B6A02A5E6F62FF9EC809EA6289C5FE9E" box="[622,755,301,328]" captionStart="Figs 30–36" captionStartId="20.[174,215,1308,1330]" captionTargetBox="[174,1111,740,1280]" captionTargetId="figure-174@20.[174,1112,740,1280]" captionTargetPageId="20" captionText="Figs 30–36. New species of Foveolaria n. gen.from the Bakony Mts, Hungary localities of the Hierlatz Limestone Formation. – 30–31. Holotype of Foveolaria kocsisi n. sp., GGIH J.08.9.1. from beds with Obtusum to Ibex Zones (Upper Sinemurian to Lower Pliensbachian, Lower Jurassic) mixed fauna in Hierlatz Limestone at Kericser-dűlő, Lókút, Bakony Mts, Hungary. – 30. The only preserved whorl and base parts, ×1. – 31. The same in ×3.5 magnification for the details. – 32–36. Holotype of Foveolaria suemegensis n. sp., GGIH J.08.8.1. from the (?)Upper Sinemurian (Lower Jurassic) Hierlatz Limestone of Mogyorósdomb, Sümeg, Bakony Mts, Hungary. – 32–33. Most informative surface parts of the monotype, ×1. – 32. Nearly parallel view to the inferable peristome plane, showing a short part of the outer lip below the suture. – 33. The shell from the opposite side. – 34. Weakening ornament of the early spire whorls, ×2.5. – 35. Magnified sample view of the obscure ornament on the ramp of the latest whorls, ×5.– 36. Networkornament of the early spire,×4.5" figureDoi="http://doi.org/10.5281/zenodo.15627169" httpUri="https://zenodo.org/record/15627169/files/figure.png" pageId="21" pageNumber="24">Figs 32–36</figureCitation>
)
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1980
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<emphasis id="1CEFEAC96F62FF9ECB68EA2F8A60FEA3" box="[271,342,352,373]" italics="true" pageId="21" pageNumber="24">Sisenna</emphasis>
cf.
<emphasis id="1CEFEAC96F62FF9ECB1FEA2F8A8DFEA3" box="[376,443,352,373]" italics="true" pageId="21" pageNumber="24">procera</emphasis>
(Deslongchamps, 1849)
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– SZABÓ, p. 55, Pl. I,
<figureCitation id="B6A02A5E6F62FF9EC90FEA10888CFEA3" box="[872,954,351,373]" captionStart="Figs 1–3" captionStartId="3.[219,260,1535,1557]" captionTargetBox="[222,1152,1201,1496]" captionTargetId="figure-324@3.[219,1156,1197,1499]" captionTargetPageId="3" captionText="Figs 1–3. The peculiar “quirinii type” ridges, occurring in Wortheniopsis Böhm, 1895, Foveolaria n. gen. and some probably related forms. – 1. Copy of KITTL’s (1899, p. 14, Fig. 2) sketch. – 2. The ridges on the outer face and on the base of Wortheniopsis rakusi n. sp. – 3. The ridges on the base of Foveolaria conoidea n. sp." figureDoi="http://doi.org/10.5281/zenodo.15627165" httpUri="https://zenodo.org/record/15627165/files/figure.png" pageId="21" pageNumber="24">Figs 2–3</figureCitation>
.
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2009
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<emphasis id="1CEFEAC96F62FF9ECB69EA338ABBFE47" box="[270,397,380,401]" italics="true" pageId="21" pageNumber="24">Wortheniopsis</emphasis>
(
<emphasis id="1CEFEAC96F62FF9ECBFDEA328AD7FE44" box="[410,481,381,402]" italics="true" pageId="21" pageNumber="24">Sisenna</emphasis>
) aff.
<emphasis id="1CEFEAC96F62FF9EC875EA33895BFE47" box="[530,621,380,401]" italics="true" pageId="21" pageNumber="24">lokutensis</emphasis>
<bibRefCitation id="4A0A4B2A6F62FF9EC815EA3389D4FE44" author="SZABO J." box="[626,738,380,402]" pageId="21" pageNumber="24" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
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– SZABÓ, p. 34,
<figureCitation id="B6A02A5E6F62FF9EC91CEA33888DFE44" box="[891,955,380,402]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="21" pageNumber="24">Fig. 27</figureCitation>
A-B.
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<typeStatus id="F12088796F62FF9ECB71EAF58A78FE05" box="[278,334,442,467]" pageId="21" pageNumber="24">Type</typeStatus>
specimen
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–
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J.08.8.1.
</paragraph>
<paragraph id="2E2436DB6F62FF9ECB71EA9288C9FE21" blockId="21.[219,1157,441,1664]" box="[278,1023,476,503]" pageId="21" pageNumber="24">
<emphasis id="1CEFEAC96F62FF9ECB71EA928A91FE20" box="[278,423,476,502]" italics="true" pageId="21" pageNumber="24">
<typeStatus id="F12088796F62FF9ECB71EA928A78FE20" box="[278,334,477,502]" pageId="21" pageNumber="24">Type</typeStatus>
locality
</emphasis>
–
<location id="2B4460006F62FF9ECBA4EA9389B7FE20" LSID="urn:lsid:plazi:treatment:A63287CD6F62FF9DC86BEA458924FDCD:2B4460006F62FF9ECBA4EA9389B7FE20" box="[451,641,476,502]" country="Hungary" county="Bakony Mts" municipality="Sumeg" name="Mogyorosdomb" pageId="21" pageNumber="24">Mogyorósdomb</location>
,
<collectingMunicipality id="CE40ACA16F62FF9EC8ECEA9389EBFE20" box="[651,733,476,502]" pageId="21" pageNumber="24">Sümeg</collectingMunicipality>
,
<collectingCounty id="C7454E576F62FF9EC88FEA938842FE21" box="[744,884,476,503]" pageId="21" pageNumber="24">Bakony Mts</collectingCounty>
(
<collectingCountry id="568C764B6F62FF9EC9E4EA9388C5FE21" box="[899,1011,476,503]" name="Hungary" pageId="21" pageNumber="24">Hungary</collectingCountry>
).
</paragraph>
</materialsCitation>
<paragraph id="2E2436DB6F62FF9ECB71E94F8F7BFDCF" blockId="21.[219,1157,441,1664]" box="[278,1101,511,538]" pageId="21" pageNumber="24">
<materialsCitation id="9EF33C866F62FF9ECB71E94F8F7FFDCF" box="[278,1097,511,538]" county="Hierlatz Limestone" location="Upper" municipality="Lower Jurassic" pageId="21" pageNumber="24" specimenCount="1" typeStatus="holotype">
<emphasis id="1CEFEAC96F62FF9ECB71E94F8AA0FDCF" box="[278,406,512,537]" italics="true" pageId="21" pageNumber="24">
<typeStatus id="F12088796F62FF9ECB71E94F8A79FDCF" box="[278,335,512,537]" pageId="21" pageNumber="24">Type</typeStatus>
strata
</emphasis>
–
<location id="2B4460006F62FF9ECBD4EAB08AC8FDCF" LSID="urn:lsid:plazi:treatment:A63287CD6F62FF9DC86BEA458924FDCD:2B4460006F62FF9ECBD4EAB08AC8FDCF" box="[435,510,511,537]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Upper" pageId="21" pageNumber="24">Upper</location>
?
<location id="2B4460006F62FF9EC868EAB089A1FDCF" LSID="urn:lsid:plazi:treatment:A63287CD6F62FF9DC86BEA458924FDCD:2B4460006F62FF9EC868EAB089A1FDCF" box="[527,663,511,537]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Sinemurian" pageId="21" pageNumber="24">Sinemurian</location>
(
<collectingMunicipality id="CE40ACA16F62FF9EC8C2EAB08862FDCC" box="[677,852,511,538]" pageId="21" pageNumber="24">Lower Jurassic</collectingMunicipality>
)
<collectingCounty id="C7454E576F62FF9EC905EAB08F7FFDCF" box="[866,1097,511,537]" pageId="21" pageNumber="24">Hierlatz Limestone</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="2E2436DB6F62FF9ECB71E96C884AFDEB" blockId="21.[219,1157,441,1664]" box="[278,892,546,573]" pageId="21" pageNumber="24">
<emphasis id="1CEFEAC96F62FF9ECB71E96C8AD9FDEA" box="[278,495,546,572]" italics="true" pageId="21" pageNumber="24">Derivation of name</emphasis>
– From the name of town Sümeg.
</paragraph>
<paragraph id="2E2436DB6F62FF9ECB71E9098F49FC81" blockId="21.[219,1157,441,1664]" pageId="21" pageNumber="24">
<emphasis id="1CEFEAC96F62FF9ECB71E9098AB5FD89" box="[278,387,582,607]" italics="true" pageId="21" pageNumber="24">Diagnosis</emphasis>
– Shell having rather high spire and slightly acute early teleoconch. Marked angulation of whorl surface providing slightly pagodiform outline. Ramp feebly convex initially then feebly concave on last whorl. Outer face concave, its basal rim similarly angular to whorl surface. Initially abaxial rim of base then angulation at abapical rim of selenizone corresponding to periphery. Base with moderately convex wall. Weak spiral lines ornament whorls, crossed by thin, collabral threads on early whorls with granules at intersections. Two stronger threads in middle of ramp present on all visible whorls, except last one.
</paragraph>
<paragraph id="2E2436DB6F62FF9ECB71E8108A1FFC4B" blockId="21.[219,1157,441,1664]" pageId="21" pageNumber="24">
<emphasis id="1CEFEAC96F62FF9ECB71E8108A4DFCAF" box="[278,379,863,889]" italics="true" pageId="21" pageNumber="24">Material</emphasis>
– Single, fragmentary specimen without earliest whorls and peristome.
</paragraph>
<paragraph id="2E2436DB6F62FF9ECB71E8E9890AFC35" blockId="21.[219,1157,441,1664]" pageId="21" pageNumber="24">
<emphasis id="1CEFEAC96F62FF9ECB71E8E98AAFFC69" box="[278,409,934,959]" italics="true" pageId="21" pageNumber="24">Dimensions</emphasis>
– Preserved width:
<quantity id="E9639B3E6F62FF9EC8E7E8E989E3FC16" box="[640,725,934,960]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="21" pageNumber="24" unit="mm" value="33.0">33 mm</quantity>
; earliest measurable spiral angle: 55°; spiral angle on last whorl: 63°.
</paragraph>
<paragraph id="2E2436DB6F62FF9ECB71E8A28981FAF6" blockId="21.[219,1157,441,1664]" pageId="21" pageNumber="24">
<emphasis id="1CEFEAC96F62FF9ECB71E8A28AA1FBD0" box="[278,407,1005,1030]" italics="true" pageId="21" pageNumber="24">Description</emphasis>
– A clearly gradate shell which has a relatively high spire with slightly acute early teleoconch. No information is available from the protoconch. A marked angulation is present on the whorl surface already from the early shell that causes a slightly pagodiform outline. The ramp is feebly convex on the early whorls but becomes feebly concave for the last whorl. The outer face is concave, its basal rim similarly angular to that on the whorl surface. Initially, the abaxial rim of the base then the angulation at the abapical rim of selenizone corresponds to the periphery. The base seems fairly convex from the available fragment as whole and has a moderately convex wall.
</paragraph>
<paragraph id="2E2436DB6F62FF9ECB71EE6688E9FA7A" blockId="21.[219,1157,441,1664]" pageId="21" pageNumber="24">Weak spiral lines ornament the whorls, crossed by thin collabral threads on the early whorls with granules at the intersections. Two marked threads in the middle of the ramp are present on all visible whorls, except the last one. The outer face is ornamented by several equally spaced, obscure spiral striae.</paragraph>
<paragraph id="2E2436DB6F62FF9DCB71EEF98A70FE02" blockId="21.[219,1157,441,1664]" lastBlockId="22.[174,1112,266,539]" lastPageId="22" lastPageNumber="25" pageId="21" pageNumber="24">
<emphasis id="1CEFEAC96F62FF9ECB71EEF98A4DFA06" box="[278,379,1462,1488]" italics="true" pageId="21" pageNumber="24">Remarks</emphasis>
– The slightly acute spire and the marked, keel-like angulation of the whorls suggested affinity of the species to
<taxonomicName id="E99B4D586F62FF9EC951EE9689E6F9C0" authority="J. A. Eudes-Deslongchamps, 1849" authorityName="J. A. Eudes-Deslongchamps" authorityYear="1849" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="21" pageNumber="24" phylum="Mollusca" rank="variety" species="faveolata" variety="procera">
<emphasis id="1CEFEAC96F62FF9EC951EE968F66FA25" box="[822,1104,1497,1523]" italics="true" pageId="21" pageNumber="24">Pleurotomaria faveolata</emphasis>
var.
<emphasis id="1CEFEAC96F62FF9ECABCEEB28A18F9C0" box="[219,302,1533,1558]" italics="true" pageId="21" pageNumber="24">procera</emphasis>
J. A.
<bibRefCitation id="4A0A4B2A6F62FF9ECB1FEEB389E6F9C0" author="EUDES-DESLONGCHAMPS J. A." box="[376,720,1532,1558]" pageId="21" pageNumber="24" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">Eudes-Deslongchamps, 1849</bibRefCitation>
</taxonomicName>
drawings (
<bibRefCitation id="4A0A4B2A6F62FF9EC931EEB388C6F9C1" author="SZABO J." box="[854,1008,1532,1559]" pageId="21" pageNumber="24" pagination="49 - 71" refId="ref12779" refString="SZABO J. 1980: Lower and Middle Jurassic Gastropods from the Bakony Mountains (Hungary). Part II. Pleurotomariacea and Fissurellacea (Archaeogastropoda). - Annales historico-naturales Musei nationalis hungarici 72: 49-71." type="journal article" year="1980">SZABÓ 1980</bibRefCitation>
) but photos in
<bibRefCitation id="4A0A4B2A6F62FF9ECA9BED508970F9EC" author="FISCHER J. - C. &amp; WEBER C." box="[252,582,1567,1594]" pageId="21" pageNumber="24" pagination="451" refId="ref11914" refString="FISCHER J. - C. &amp; WEBER C. 1997: Volume II, Gasteropodes Jurassiques. - In: FISCHER J. C. (ed.): Revision critique de la Paleontologie Francaise, incluant la reedition de la original. - pp. 300 + 622, Pls 451; Paris, Masson &amp; Museum national d'Histoire naturelle." type="book chapter" year="1997">FISCHER &amp; WEBER (1997)</bibRefCitation>
display a specimen as “morphe
<taxonomicName id="E99B4D586F62FF9EC9AFED508F19F9EF" baseAuthorityName="Deslongchamps" baseAuthorityYear="1849" box="[968,1071,1567,1593]" class="Gastropoda" family="Pleurotomariidae" genus="Sisenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="21" pageNumber="24" phylum="Mollusca" rank="species" species="procera">
”
<emphasis id="1CEFEAC96F62FF9EC9BBED6F8F19F9EF" box="[988,1071,1568,1593]" italics="true" pageId="21" pageNumber="24">procera</emphasis>
</taxonomicName>
that is strongly different from the earliest figures and not similar to the Sümeg specimen. The shell is somewhat similar to that of
<taxonomicName id="E99B4D586F62FF9EC88AED298FB3F956" authority="(Szabo, 2009)" baseAuthorityName="Szabo" baseAuthorityYear="2009" box="[749,1157,1637,1664]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="24" phylum="Bryozoa" rank="species" species="lokutensis">
<emphasis id="1CEFEAC96F62FF9EC88AED2988E8F956" box="[749,990,1638,1664]" italics="true" pageId="21" pageNumber="24">Foveolaria lokutensis</emphasis>
(
<bibRefCitation id="4A0A4B2A6F62FF9EC98BED298F4BF956" author="SZABO J." box="[1004,1149,1637,1664]" pageId="21" pageNumber="24" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
)
</taxonomicName>
therefore the name
<taxonomicName id="E99B4D586F61FF9DCBFFEA458879FEF2" authorityName="Szabo" authorityYear="2009" box="[408,847,266,293]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" isUncertain="true" kingdom="Animalia" order="Pleurotomariida" pageId="22" pageNumber="25" phylum="Mollusca" rank="species" species="lokutensis" subGenus="Sisenna">
<emphasis id="1CEFEAC96F61FF9DCBFFEA45890EFEF2" box="[408,568,266,292]" italics="true" pageId="22" pageNumber="25">Wortheniopsis</emphasis>
(
<emphasis id="1CEFEAC96F61FF9DC82DEA448994FEF3" box="[586,674,267,293]" italics="true" pageId="22" pageNumber="25">Sisenna</emphasis>
) aff.
<emphasis id="1CEFEAC96F61FF9DC8B8EA458879FEF2" box="[735,847,266,292]" italics="true" pageId="22" pageNumber="25">lokutensis</emphasis>
</taxonomicName>
has been given to this species subsequently. Since the earliest parts are slightly acute and the angulations of the whorls are more distinct, keel like, while the shell outline is simply conoidal in
<taxonomicName id="E99B4D586F61FF9DCB5BEA3A8AFCFE58" authorityName="Szabo" authorityYear="2009" box="[316,458,372,398]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="22" pageNumber="25" phylum="Mollusca" rank="species" species="lokutensis">
<emphasis id="1CEFEAC96F61FF9DCB5BEA3A8A7AFE58" box="[316,332,373,398]" italics="true" pageId="22" pageNumber="25">F</emphasis>
.
<emphasis id="1CEFEAC96F61FF9DCB3DEA3B8AFCFE58" box="[346,458,372,398]" italics="true" pageId="22" pageNumber="25">lokutensis</emphasis>
</taxonomicName>
, the ramp is narrower and the outer face is wider than in
<taxonomicName id="E99B4D586F61FF9DCAABEAD78A6CFE67" authorityName="Szabo" authorityYear="2009" box="[204,346,407,434]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="22" pageNumber="25" phylum="Mollusca" rank="species" species="lokutensis">
<emphasis id="1CEFEAC96F61FF9DCAABEAD78BEAFE67" box="[204,220,408,433]" italics="true" pageId="22" pageNumber="25">F</emphasis>
.
<emphasis id="1CEFEAC96F61FF9DCA8DEAD88A6CFE67" box="[234,346,407,433]" italics="true" pageId="22" pageNumber="25">lokutensis</emphasis>
</taxonomicName>
, establishing a new species seems satisfactorily supported by the morphology.
</paragraph>
<paragraph id="2E2436DB6F61FF9DCA8EEA928924FDCD" blockId="22.[174,1112,266,539]" pageId="22" pageNumber="25">
<emphasis id="1CEFEAC96F61FF9DCA8EEA928A41FE21" box="[233,375,477,503]" italics="true" pageId="22" pageNumber="25">Distribution</emphasis>
– Upper(?) Sinemurian Hierlatz Limestone, Mogyorósdomb, Sümeg, Bakony Mts,
<collectingCountry id="568C764B6F61FF9DCBC3E94E8938FDCD" box="[420,526,513,539]" name="Hungary" pageId="22" pageNumber="25">Hungary</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

212
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<paragraph id="2E2436DB6F65FF99CB9AEE3E883EFA5D" blockId="18.[509,776,1393,1455]" box="[509,776,1393,1419]" pageId="18" pageNumber="21">
<heading id="756C81B76F65FF99CB9AEE3E883EFA5D" box="[509,776,1393,1419]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="21" reason="9">
<taxonomicName id="E99B4D586F65FF99CB9AEE3E89F3FA5D" authorityName="Szabó" authorityYear="2016" box="[509,709,1393,1419]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="21" phylum="Bryozoa" rank="species" species="kocsisi" status="sp. nov.">
<emphasis id="1CEFEAC96F65FF99CB9AEE3E89F3FA5D" box="[509,709,1393,1419]" italics="true" pageId="18" pageNumber="21">Foveolaria kocsisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F65FF99C8ACEE3E883EFA5D" box="[715,776,1393,1419]" pageId="18" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="2E2436DB6F65FF99C85FEEDB89FBFA79" blockId="18.[509,776,1393,1455]" box="[568,717,1428,1455]" pageId="18" pageNumber="21">
(
<figureCitation id="B6A02A5E6F65FF99C827EEDB89F3FA79" box="[576,709,1428,1455]" captionStart="Figs 30–36" captionStartId="20.[174,215,1308,1330]" captionTargetBox="[174,1111,740,1280]" captionTargetId="figure-174@20.[174,1112,740,1280]" captionTargetPageId="20" captionText="Figs 30–36. New species of Foveolaria n. gen.from the Bakony Mts, Hungary localities of the Hierlatz Limestone Formation. – 30–31. Holotype of Foveolaria kocsisi n. sp., GGIH J.08.9.1. from beds with Obtusum to Ibex Zones (Upper Sinemurian to Lower Pliensbachian, Lower Jurassic) mixed fauna in Hierlatz Limestone at Kericser-dűlő, Lókút, Bakony Mts, Hungary. – 30. The only preserved whorl and base parts, ×1. – 31. The same in ×3.5 magnification for the details. – 32–36. Holotype of Foveolaria suemegensis n. sp., GGIH J.08.8.1. from the (?)Upper Sinemurian (Lower Jurassic) Hierlatz Limestone of Mogyorósdomb, Sümeg, Bakony Mts, Hungary. – 32–33. Most informative surface parts of the monotype, ×1. – 32. Nearly parallel view to the inferable peristome plane, showing a short part of the outer lip below the suture. – 33. The shell from the opposite side. – 34. Weakening ornament of the early spire whorls, ×2.5. – 35. Magnified sample view of the obscure ornament on the ramp of the latest whorls, ×5.– 36. Networkornament of the early spire,×4.5" figureDoi="http://doi.org/10.5281/zenodo.15627169" httpUri="https://zenodo.org/record/15627169/files/figure.png" pageId="18" pageNumber="21">Figs 30–31</figureCitation>
)
</paragraph>
<paragraph id="2E2436DB6F65FF99CAC9EE898857FA0A" blockId="18.[174,989,1478,1529]" box="[174,865,1478,1500]" pageId="18" pageNumber="21">
1980
<taxonomicName id="E99B4D586F65FF99CA85EE89896FFA0A" authority="(Deslongchamps, 1849)" baseAuthorityName="Deslongchamps" baseAuthorityYear="1849" box="[226,601,1478,1500]" class="Gastropoda" family="Pleurotomariidae" genus="Sisenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="21" phylum="Mollusca" rank="species" species="pinguis">
<emphasis id="1CEFEAC96F65FF99CA85EE898A46FA0A" box="[226,368,1478,1500]" italics="true" pageId="18" pageNumber="21">Sisenna pinguis</emphasis>
(Deslongchamps, 1849)
</taxonomicName>
– SZABÓ, p. 53, Pl. I,
<figureCitation id="B6A02A5E6F65FF99C941EE89886AFA0A" box="[806,860,1478,1500]" captionStart="Figs 1–3" captionStartId="3.[219,260,1535,1557]" captionTargetBox="[222,1152,1201,1496]" captionTargetId="figure-324@3.[219,1156,1197,1499]" captionTargetPageId="3" captionText="Figs 1–3. The peculiar “quirinii type” ridges, occurring in Wortheniopsis Böhm, 1895, Foveolaria n. gen. and some probably related forms. – 1. Copy of KITTL’s (1899, p. 14, Fig. 2) sketch. – 2. The ridges on the outer face and on the base of Wortheniopsis rakusi n. sp. – 3. The ridges on the base of Foveolaria conoidea n. sp." figureDoi="http://doi.org/10.5281/zenodo.15627165" httpUri="https://zenodo.org/record/15627165/files/figure.png" pageId="18" pageNumber="21">Fig. 1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="668165506F65FF99CAC9EEAC88EBFA2E" box="[174,989,1507,1529]" pageId="18" pageNumber="21" type="reference_group">
<paragraph id="2E2436DB6F65FF99CAC9EEAC88EBFA2E" blockId="18.[174,989,1478,1529]" box="[174,989,1507,1529]" pageId="18" pageNumber="21">
2009
<taxonomicName id="E99B4D586F65FF99CA85EEAC89F9FA2F" authority="Szabo, 2009" authorityName="Szabo" authorityYear="2009" box="[226,719,1507,1529]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" isUncertain="true" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="21" phylum="Mollusca" rank="species" species="hierlatzensis" subGenus="Sisenna">
<emphasis id="1CEFEAC96F65FF99CA85EEAC8A8DFA2E" box="[226,443,1507,1528]" italics="true" pageId="18" pageNumber="21">Wortheniopsis (Sisenna)</emphasis>
aff.
<emphasis id="1CEFEAC96F65FF99CB85EEAC8961FA2E" box="[482,599,1507,1528]" italics="true" pageId="18" pageNumber="21">hierlatzensis</emphasis>
<bibRefCitation id="4A0A4B2A6F65FF99C83AEEAC89F9FA2F" author="SZABO J." box="[605,719,1507,1529]" pageId="18" pageNumber="21" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
</taxonomicName>
– SZABÓ, p. 34,
<figureCitation id="B6A02A5E6F65FF99C90DEEAC88EFFA2E" box="[874,985,1507,1529]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="18" pageNumber="21">Fig. 27C, D</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="668165506F65FF9FCA8EED6F8A73FDCD" lastPageId="20" lastPageNumber="23" pageId="18" pageNumber="21" type="materials_examined">
<materialsCitation id="9EF33C866F65FF99CA8EED6F889FF98B" collectionCode="GGIH" country="Hungary" county="Bakony Mts" location="Kericser-dulo" municipality="Lokut" pageId="18" pageNumber="21" specimenCount="1" typeStatus="holotype">
<paragraph id="2E2436DB6F65FF99CA8EED6F896DF9EF" blockId="18.[233,942,1567,1629]" box="[233,603,1567,1594]" pageId="18" pageNumber="21">
<emphasis id="1CEFEAC96F65FF99CA8EED6F8ABBF9EF" box="[233,397,1568,1593]" italics="true" pageId="18" pageNumber="21">
<typeStatus id="F12088796F65FF99CA8EED6F8A17F9EF" box="[233,289,1568,1593]" pageId="18" pageNumber="21">Type</typeStatus>
specimen
</emphasis>
–
<collectionCode id="488AAE1E6F65FF99CBCEED508AC1F9EF" box="[425,503,1567,1593]" pageId="18" pageNumber="21">GGIH</collectionCode>
J.08.9.1.
</paragraph>
<paragraph id="2E2436DB6F65FF99CA8EED0C8898F98B" blockId="18.[233,942,1567,1629]" box="[233,942,1602,1629]" pageId="18" pageNumber="21">
<emphasis id="1CEFEAC96F65FF99CA8EED0C8A4CF98A" box="[233,378,1602,1628]" italics="true" pageId="18" pageNumber="21">
<typeStatus id="F12088796F65FF99CA8EED0C8A17F98A" box="[233,289,1603,1628]" pageId="18" pageNumber="21">Type</typeStatus>
locality
</emphasis>
–
<location id="2B4460006F65FF99CBF1ED0D890EF98A" LSID="urn:lsid:plazi:treatment:A63287CD6F65FF9FCB9AEE3E8F6EFD7E:2B4460006F65FF99CBF1ED0D890EF98A" box="[406,568,1602,1628]" country="Hungary" county="Bakony Mts" municipality="Lokut" name="Kericser-dulo" pageId="18" pageNumber="21">Kericser-dűlő</location>
,
<collectingMunicipality id="CE40ACA16F65FF99C825ED0D89BAF98A" box="[578,652,1602,1628]" pageId="18" pageNumber="21">Lókút</collectingMunicipality>
,
<collectingCounty id="C7454E576F65FF99C8F1ED0D8815F98B" box="[662,803,1602,1629]" pageId="18" pageNumber="21">Bakony Mts</collectingCounty>
(
<collectingCountry id="568C764B6F65FF99C955ED0D8894F98B" box="[818,930,1602,1629]" name="Hungary" pageId="18" pageNumber="21">Hungary</collectingCountry>
).
</paragraph>
</materialsCitation>
<paragraph id="2E2436DB6F64FF98CB71EA4488D3FE91" blockId="19.[219,1157,266,1630]" pageId="19" pageNumber="22">
<emphasis id="1CEFEAC96F64FF98CB71EA448AADFEF2" box="[278,411,267,292]" italics="true" pageId="19" pageNumber="22">
<typeStatus id="F12088796F64FF98CB71EA448A79FEF2" box="[278,335,267,292]" pageId="19" pageNumber="22">Type</typeStatus>
strata
</emphasis>
– Beds with Obtusum to Ibex Zones (Upper Sinemurian to Lower Pliensbachian, Jurassic) mixed fauna in Hierlatz Limestone.
</paragraph>
<paragraph id="2E2436DB6F64FF98CB71EA1D89ACFE58" blockId="19.[219,1157,266,1630]" pageId="19" pageNumber="22">
<emphasis id="1CEFEAC96F64FF98CB71EA1D8AC6FEBD" box="[278,496,337,363]" italics="true" pageId="19" pageNumber="22">Derivation of name</emphasis>
– Dedicated to Lajos KOCSIS (Sümeg), leader worker of the collecting staff at the
<typeStatus id="F12088796F64FF98C864EA3B8901FE58" box="[515,567,372,398]" pageId="19" pageNumber="22">type</typeStatus>
locality.
</paragraph>
<paragraph id="2E2436DB6F64FF98CB71EAD78F11FD52" blockId="19.[219,1157,266,1630]" pageId="19" pageNumber="22">
<emphasis id="1CEFEAC96F64FF98CB71EAD78AB5FE67" box="[278,387,408,433]" italics="true" pageId="19" pageNumber="22">Diagnosis</emphasis>
– Gradate shell with whorls of flattish concave ramp and concave outer face. Suture impressed, running slightly below periphery. Base subglobular with convex wall and no or only narrow umbilicus. Concave selenizone between sharp threads. Ornament consisting of network from spiral and collabral threads with granules at intersection points on ramp and adapical part of outer face. Quirinii
<typeStatus id="F12088796F64FF98CB25E9078A40FDB4" box="[322,374,584,610]" pageId="19" pageNumber="22">type</typeStatus>
spiral ridges on abapical belt of outer face and on base with short, riblet-like granules, aligned into seemingly continuous collabral threads.
</paragraph>
<paragraph id="2E2436DB6F64FF98CB71E9C2895FFD1D" blockId="19.[219,1157,266,1630]" pageId="19" pageNumber="22">
<emphasis id="1CEFEAC96F64FF98CB71E9C28A4DFD71" box="[278,379,653,679]" italics="true" pageId="19" pageNumber="22">Material</emphasis>
– Two fragmentary specimens, one is shelly, the other is an inner mould with small shell fragments.
</paragraph>
<paragraph id="2E2436DB6F64FF98CB71E99A8F38FD38" blockId="19.[219,1157,266,1630]" box="[278,1038,724,750]" pageId="19" pageNumber="22">
<emphasis id="1CEFEAC96F64FF98CB71E99A8AAFFD38" box="[278,409,725,750]" italics="true" pageId="19" pageNumber="22">Dimensions</emphasis>
– Preserved height:
<quantity id="E9639B3E6F64FF98C8EEE99B89E9FD38" box="[649,735,724,750]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="22" unit="mm" value="11.0">11 mm</quantity>
; preserved width:
<quantity id="E9639B3E6F64FF98C9D1E99B8F3CFD38" box="[950,1034,724,750]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="22" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
<paragraph id="2E2436DB6F64FF98CB71E9B78F00FBA7" blockId="19.[219,1157,266,1630]" pageId="19" pageNumber="22">
<emphasis id="1CEFEAC96F64FF98CB71E9B78AA0FCC7" box="[278,406,760,785]" italics="true" pageId="19" pageNumber="22">Description</emphasis>
– The shells are gradate with clearly angulate whorls. The ramp is flattish but slightly convex just below the suture; its abapical rim is concave along the selenizone. The selenizone itself is moderately wide, concave between two sharp spiral cords; the abapical one just coincides with angulation of the whorl surface. The outer face is rather wide and its surface is nearly parallel to the axis but having also a feebly concave median belt. Its lower edge is slightly swollen next the rounded-angular periphery. The suture follows the abapically first spiral ridge from the peripheral one. The base is subglobular with convex wall, and it is seemingly anomphalous or possibly narrowly phaneromphalous. No peristome part can be observed in the available specimens; the growth lines indicate a prosocline outer lip with a rather deep sinus at the height of the inferred exhalant slit.
</paragraph>
<paragraph id="2E2436DB6F64FF98CB71EF358AB7FA78" blockId="19.[219,1157,266,1630]" pageId="19" pageNumber="22">
Collabral riblets and spiral threads intersect each other with granules on the ramp and the riblets weaken towards the selenizone. The shape of the spiral threads gradually changes into quirinii
<typeStatus id="F12088796F64FF98C8D7EF8E89D2FB0D" box="[688,740,1217,1243]" pageId="19" pageNumber="22">type</typeStatus>
ridges on the outer face in abaxial direction; only the latter ones are observed on the base. Instead of riblets, collabral threads are present on the outer face. Seemingly, they are continued also on the base but there the threads are actually composed of short riblets between pairs of the quirinii
<typeStatus id="F12088796F64FF98CBB6EE018933FABE" box="[465,517,1358,1384]" pageId="19" pageNumber="22">type</typeStatus>
ridges and are aligned as apparently continuous collabral threads; they are most conspicuous at the periphery. The selenizone bears dense lunulae.
</paragraph>
<paragraph id="2E2436DB6F64FF98CB71EEF88845F988" blockId="19.[219,1157,266,1630]" pageId="19" pageNumber="22">
<emphasis id="1CEFEAC96F64FF98CB71EEF88A4CFA07" box="[278,378,1463,1489]" italics="true" pageId="19" pageNumber="22">Remarks</emphasis>
– Both specimens are fragments of gradate shells of similar size; the inner mould one shows the conoidal-gradate outline. Though their size suggests early ontogenetic stage, the sharp angulation of the whorls and the dense lunulae seem to indicate nearly full-grown shells. The available information about the shell is satisfactory for the distinction of this species.
</paragraph>
<paragraph id="2E2436DB6F63FF9FCA8EEA458A73FDCD" blockId="20.[174,1114,266,680]" pageId="20" pageNumber="23">
The shell fragments are most similar to
<taxonomicName id="E99B4D586F63FF9FC8D3EA458F6EFEF3" authority="(Szabo, 2009)" baseAuthorityName="Szabo" baseAuthorityYear="2009" box="[692,1112,266,293]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="23" phylum="Bryozoa" rank="species" species="hierlatzensis">
<emphasis id="1CEFEAC96F63FF9FC8D3EA45888FFEF2" box="[692,953,266,292]" italics="true" pageId="20" pageNumber="23">Foveolaria hierlatzensis</emphasis>
(
<bibRefCitation id="4A0A4B2A6F63FF9FC9A1EA458F66FEF3" author="SZABO J." box="[966,1104,266,293]" pageId="20" pageNumber="23" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
)
</taxonomicName>
but its larger shells are without angulation in the growth stage where the size is similar to that of
<taxonomicName id="E99B4D586F63FF9FCB2FEA1E8927FEBD" authorityName="Szabó" authorityYear="2016" box="[328,529,337,363]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="23" phylum="Bryozoa" rank="species" species="kocsisi" status="sp. nov.">
<emphasis id="1CEFEAC96F63FF9FCB2FEA1E8927FEBD" box="[328,529,337,363]" italics="true" pageId="20" pageNumber="23">Foveolaria kocsisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F63FF9FC87DEA1E896DFEBD" box="[538,603,337,363]" pageId="20" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
; the position and morphology of the selenizone, and the ramp are also different. The selenizone lies clearly above the angulation in
<taxonomicName id="E99B4D586F63FF9FCB63EAD78A5EFE67" authorityName="Szabó" authorityYear="2016" box="[260,360,407,433]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="23" phylum="Bryozoa" rank="species" species="kocsisi" status="n. sp.">
<emphasis id="1CEFEAC96F63FF9FCB63EAD78A5EFE67" box="[260,360,407,433]" italics="true" pageId="20" pageNumber="23">F. kocsisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F63FF9FCB09EAD78A9AFE64" box="[366,428,408,434]" pageId="20" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
in the level of the ramp, however, it blunts the angulation in
<taxonomicName id="E99B4D586F63FF9FCAACEAF48A42FE02" baseAuthorityName="Szabo" baseAuthorityYear="2009" box="[203,372,442,468]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="23" phylum="Bryozoa" rank="species" species="hierlatzensis">
<emphasis id="1CEFEAC96F63FF9FCAACEAF48A42FE02" box="[203,372,442,468]" italics="true" pageId="20" pageNumber="23">F. hierlatzensis</emphasis>
</taxonomicName>
, forms a third surface between the ramp and the outer face, and angled to both. The ramp is flattish concave in
<taxonomicName id="E99B4D586F63FF9FC883EA91887FFE21" authorityName="Szabó" authorityYear="2016" box="[740,841,477,503]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="23" phylum="Bryozoa" rank="species" species="kocsisi" status="n. sp.">
<emphasis id="1CEFEAC96F63FF9FC883EA91887FFE21" box="[740,841,477,503]" italics="true" pageId="20" pageNumber="23">F. kocsisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F63FF9FC936EA9188A7FE2E" box="[849,913,478,504]" pageId="20" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
but convex in
<emphasis id="1CEFEAC96F63FF9FCE24EA918A73FDCD" italics="true" pageId="20" pageNumber="23">
<taxonomicName id="E99B4D586F63FF9FCE24EA918A77FDCD" baseAuthorityName="Szabo" baseAuthorityYear="2009" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="23" phylum="Bryozoa" rank="species" species="hierlatzensis">F. hierlatzensis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="668165506F63FF9FCA8EE96B8855FDB4" pageId="20" pageNumber="23" type="reference_group">
<paragraph id="2E2436DB6F63FF9FCA8EE96B8855FDB4" blockId="20.[174,1114,266,680]" pageId="20" pageNumber="23">
<taxonomicName id="E99B4D586F63FF9FCA8EE96B897EFDE9" authority="(Szabo, 2009)" baseAuthorityName="Szabo" baseAuthorityYear="2009" box="[233,584,548,575]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="23" phylum="Bryozoa" rank="species" species="jancsii">
<emphasis id="1CEFEAC96F63FF9FCA8EE96B8A9FFDE8" box="[233,425,548,574]" italics="true" pageId="20" pageNumber="23">Foveolaria jancsii</emphasis>
(
<bibRefCitation id="4A0A4B2A6F63FF9FCBD1E96B8977FDE9" author="SZABO J." box="[438,577,548,575]" pageId="20" pageNumber="23" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
)
</taxonomicName>
has convex ramp, and coeloconoidal shell with less markedly gradate outline, differently from
<taxonomicName id="E99B4D586F63FF9FC8DAE9078816FDB7" authorityName="Szabó" authorityYear="2016" box="[701,800,583,609]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="23" phylum="Bryozoa" rank="species" species="kocsisi" status="n. sp.">
<emphasis id="1CEFEAC96F63FF9FC8DAE9078816FDB7" box="[701,800,583,609]" italics="true" pageId="20" pageNumber="23">F. kocsisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F63FF9FC941E9078855FDB4" box="[806,867,584,610]" pageId="20" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="668165506F63FF9FCA8EE9258F6EFD7E" pageId="20" pageNumber="23" type="distribution">
<paragraph id="2E2436DB6F63FF9FCA8EE9258F6EFD7E" blockId="20.[174,1114,266,680]" pageId="20" pageNumber="23">
<emphasis id="1CEFEAC96F63FF9FCA8EE9258A4EFD52" box="[233,376,618,644]" italics="true" pageId="20" pageNumber="23">Distribution</emphasis>
– Beds with mixed fauna of Obtusum to Ibex Zones (Upper Sinemurian to lowermost Pliensbachian), Kericser, Lókút, Bakony Mts,
<collectingCountry id="568C764B6F63FF9FC98FE9C18F65FD7E" box="[1000,1107,654,680]" name="Hungary" pageId="20" pageNumber="23">Hungary</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="1482AA7173402D68CE5A9E67BF0320DA">Gastropods of the Lower Jurassic Hierlatz Limestone Formation, part 2. Some new archaic type slit-bearing components from the fauna of the Hierlatz Alpe (Hallstatt, Austria) and the Bakony Mts (Hungary)</mods:title>
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<paragraph id="2E2436DB6F66FF9AC841EF5E880CFBFD" blockId="17.[550,826,1041,1103]" box="[550,826,1041,1067]" pageId="17" pageNumber="20">
<heading id="756C81B76F66FF9AC841EF5E880CFBFD" box="[550,826,1041,1067]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="20" reason="9">
<taxonomicName id="E99B4D586F66FF9AC841EF5E89C1FBFD" authorityName="Szabó" authorityYear="2016" box="[550,759,1041,1067]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="species" species="globosa" status="sp. nov.">
<emphasis id="1CEFEAC96F66FF9AC841EF5E89C1FBFD" box="[550,759,1041,1067]" italics="true" pageId="17" pageNumber="20">Foveolaria globosa</emphasis>
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<taxonomicNameLabel id="07DC57B26F66FF9AC89AEF5E880CFBFD" box="[765,826,1041,1067]" pageId="17" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="B6A02A5E6F66FF9AC809EF7B89C4FB99" box="[622,754,1076,1103]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="17" pageNumber="20">Figs 23–26</figureCitation>
)
</paragraph>
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<typeStatus id="F12088796F66FF9ACB71EF258A78FB55" box="[278,334,1130,1155]" pageId="17" pageNumber="20">Type</typeStatus>
specimen
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– GBA 2017/001/0005.
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<paragraph id="2E2436DB6F66FF9ACB71EFC28878FB71" blockId="17.[219,1157,1129,1648]" box="[278,846,1164,1191]" pageId="17" pageNumber="20">
<emphasis id="1CEFEAC96F66FF9ACB71EFC28A91FB70" box="[278,423,1164,1190]" italics="true" pageId="17" pageNumber="20">
<typeStatus id="F12088796F66FF9ACB71EFC28A78FB70" box="[278,334,1165,1190]" pageId="17" pageNumber="20">Type</typeStatus>
locality
</emphasis>
–
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,
<collectingMunicipality id="CE40ACA16F66FF9AC809EFC389E3FB70" box="[622,725,1164,1190]" pageId="17" pageNumber="20">Hallstatt</collectingMunicipality>
(
<collectingCountry id="568C764B6F66FF9AC883EFC38877FB71" box="[740,833,1164,1191]" name="Austria" pageId="17" pageNumber="20">Austria</collectingCountry>
).
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<paragraph id="2E2436DB6F66FF9ACB71EFFF8AF0FB3A" blockId="17.[219,1157,1129,1648]" pageId="17" pageNumber="20">
<materialsCitation id="9EF33C866F66FF9ACB71EFFF8AF4FB3A" county="Hierlatz Limestone" location="Sinemurian" municipality="Lower Jurassic" pageId="17" pageNumber="20" specimenCount="1" typeStatus="holotype">
<emphasis id="1CEFEAC96F66FF9ACB71EFFF8AABFB1F" box="[278,413,1200,1225]" italics="true" pageId="17" pageNumber="20">
<typeStatus id="F12088796F66FF9ACB71EFFF8A79FB1F" box="[278,335,1200,1225]" pageId="17" pageNumber="20">Type</typeStatus>
strata
</emphasis>
–
<location id="2B4460006F66FF9ACBAEEFE08967FB1F" LSID="urn:lsid:plazi:treatment:A63287CD6F66FF99C841EF5E8A3DFA92:2B4460006F66FF9ACBAEEFE08967FB1F" box="[457,593,1199,1225]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Sinemurian" pageId="17" pageNumber="20">Sinemurian</location>
to lowermost
<location id="2B4460006F66FF9AC96AEFE08887FB1F" LSID="urn:lsid:plazi:treatment:A63287CD6F66FF99C841EF5E8A3DFA92:2B4460006F66FF9AC96AEFE08887FB1F" box="[781,945,1199,1225]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Pliensbachian" pageId="17" pageNumber="20">Pliensbachian</location>
(
<collectingMunicipality id="CE40ACA16F66FF9AC9A1EFE08F4BFB1C" box="[966,1149,1199,1226]" pageId="17" pageNumber="20">Lower Jurassic</collectingMunicipality>
)
<collectingCounty id="C7454E576F66FF9ACABCEF9D8AF4FB3A" box="[219,450,1234,1260]" pageId="17" pageNumber="20">Hierlatz Limestone</collectingCounty>
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.
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<emphasis id="1CEFEAC96F66FF9ACB71EFB98AD8FAD9" box="[278,494,1270,1296]" italics="true" pageId="17" pageNumber="20">Derivation of name</emphasis>
– Globosa (Lat.) = globular; referring to the last whorl of the shell, providing the shell with subglobular shape.
</paragraph>
<paragraph id="2E2436DB6F66FF9ACB71EE7289FBF9A6" blockId="17.[219,1157,1129,1648]" pageId="17" pageNumber="20">
<emphasis id="1CEFEAC96F66FF9ACB71EE728AB5FA80" box="[278,387,1341,1366]" italics="true" pageId="17" pageNumber="20">Diagnosis</emphasis>
– Subglobular shell with rounded whorls; both usual angulations of
<taxonomicName id="E99B4D586F66FF9ACA9BEE108A40FAAF" authorityName="Szabó" authorityYear="2016" box="[252,374,1375,1401]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="genus">
<emphasis id="1CEFEAC96F66FF9ACA9BEE108A40FAAF" box="[252,374,1375,1401]" italics="true" pageId="17" pageNumber="20">Foveolaria</emphasis>
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obscure. Selenizone blunting angulation at abapical rim of ramp, flat-convex outer face hiding other one at outer rim of base. Base subglobular with strongly convex wall and rather broad umbilicus. Ornament of collabral riblets and thin spiral threads on ramp, lunulae between strong limiting threads of selenizone, strong spiral threads on outer face, crossed by thin collabral threads, vanishing in abaxial belt of base. Base covered by quirinii
<typeStatus id="F12088796F66FF9AC9F2ED5F88FFF9FC" box="[917,969,1552,1578]" pageId="17" pageNumber="20">type</typeStatus>
ridges. Growth lines prosocline-prosocyrt on ramp, prosocline-feebly prosocyrt on outer face and prosocline-feebly opisthocyrt on base.
</paragraph>
<paragraph id="2E2436DB6F65FF99CA8EEA45887BFEF2" blockId="18.[174,1112,266,1348]" box="[233,845,266,293]" pageId="18" pageNumber="21">
<emphasis id="1CEFEAC96F65FF99CA8EEA458A78FEF2" box="[233,334,266,292]" italics="true" pageId="18" pageNumber="21">Material</emphasis>
– Single, damaged specimen of two whorls.
</paragraph>
<paragraph id="2E2436DB6F65FF99CA8EEA618919FE9E" blockId="18.[174,1112,266,1348]" box="[233,559,301,328]" pageId="18" pageNumber="21">
<emphasis id="1CEFEAC96F65FF99CA8EEA618A5AFE91" box="[233,364,302,327]" italics="true" pageId="18" pageNumber="21">Dimensions</emphasis>
– Width:
<quantity id="E9639B3E6F65FF99CB82EA61891DFE9E" box="[485,555,302,328]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="21" unit="mm" value="9.0">9 mm</quantity>
.
</paragraph>
<paragraph id="2E2436DB6F65FF99CA8EEA1D8F15FDB7" blockId="18.[174,1112,266,1348]" pageId="18" pageNumber="21">
<emphasis id="1CEFEAC96F65FF99CA8EEA1D8A5FFEBD" box="[233,361,338,363]" italics="true" pageId="18" pageNumber="21">Description</emphasis>
– Both angulations of the latest whorls, common in
<taxonomicName id="E99B4D586F65FF99C9BFEA1E8F66FEBD" authorityName="Szabó" authorityYear="2016" box="[984,1104,337,363]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="21" phylum="Bryozoa" rank="genus">
<emphasis id="1CEFEAC96F65FF99C9BFEA1E8F66FEBD" box="[984,1104,337,363]" italics="true" pageId="18" pageNumber="21">Foveolaria</emphasis>
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, are obscure on the available specimen, therefore the whorls have almost evenly arched surface. A rather wide selenizone is blunting the adapical angulation at the abapical rim of the ramp, and the flat-convex outer face almost fully hides also the second angulation at the outer rim of the base but the strongest spiral cord somewhat highlights it and gives the periphery. On the earlier whorl, this angulation is still clearly visible above the suture, which is impressed. The base itself is subglobular with a strongly convex wall and a rather broad phaneromphalus.
</paragraph>
<paragraph id="2E2436DB6F65FF99CA8EE9258F39FC32" blockId="18.[174,1112,266,1348]" pageId="18" pageNumber="21">
The ornament of the ramp on the last whorl in the available specimen consists of marked collabral riblets, which are equally strong along their full length and thin, dense spiral threads. Regularly repeating lunulae are present in the selenizone between strong boundary threads. Few (4) marked spiral threads are visible on the outer face; their strength increases and their interspaces widen in abapical direction. The strongest thread provides the periphery and the outer rim of the base; the suture is positioned slightly adaxially from this thread. The outer face is crossed by thin collabral threads, vanishing within the abaxial belt of the base; tiny granules are in the intersections. Quirinii
<typeStatus id="F12088796F65FF99C907E8CB88A2FC48" box="[864,916,900,926]" pageId="18" pageNumber="21">type</typeStatus>
ridges cover the base. The growth lines are prosocline-prosocyrt on the ramp, prosocline-feebly prosocyrt on the outer face and prosocline-feebly opisthocyrt on the base.
</paragraph>
<paragraph id="2E2436DB6F65FF99CA8EE8A28F6EFB43" blockId="18.[174,1112,266,1348]" pageId="18" pageNumber="21">
<emphasis id="1CEFEAC96F65FF99CA8EE8A28A7DFBD1" box="[233,331,1005,1031]" italics="true" pageId="18" pageNumber="21">Remarks</emphasis>
– Two rounded angulations do not occur on late whorls of the shell in other species. The most similar, rather globose species is
<taxonomicName id="E99B4D586F65FF99C92DEF5E8934FB99" authority="(J. A. Eudes-Deslongchamps, 1849)" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="21" phylum="Bryozoa" rank="species" species="pinguis">
<emphasis id="1CEFEAC96F65FF99C92DEF5E8F23FBFD" box="[842,1045,1041,1067]" italics="true" pageId="18" pageNumber="21">Foveolaria pinguis</emphasis>
(J. A.
<bibRefCitation id="4A0A4B2A6F65FF99CAC9EF7B8ACCFB99" author="EUDES-DESLONGCHAMPS J. A." box="[174,506,1076,1103]" pageId="18" pageNumber="21" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">Eudes-Deslongchamps, 1849</bibRefCitation>
)
</taxonomicName>
as figured by
<bibRefCitation id="4A0A4B2A6F65FF99C8C5EF7B88EAFB99" author="FISCHER J. - C. &amp; WEBER C." box="[674,988,1076,1103]" pageId="18" pageNumber="21" pagination="451" refId="ref11914" refString="FISCHER J. - C. &amp; WEBER C. 1997: Volume II, Gasteropodes Jurassiques. - In: FISCHER J. C. (ed.): Revision critique de la Paleontologie Francaise, incluant la reedition de la original. - pp. 300 + 622, Pls 451; Paris, Masson &amp; Museum national d'Histoire naturelle." type="book chapter" year="1997">FISCHER &amp; WEBER (1997)</bibRefCitation>
but its angulations and a subvertical outer face are present also on the latest whorl, furthermore, it has an ornament of granulate riblets, denser than in
<taxonomicName id="E99B4D586F65FF99C929EF358F2FFB42" authorityName="Szabó" authorityYear="2016" box="[846,1049,1146,1172]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="21" phylum="Bryozoa" rank="species" species="globosa" status="sp. nov.">
<emphasis id="1CEFEAC96F65FF99C929EF358F2FFB42" box="[846,1049,1146,1172]" italics="true" pageId="18" pageNumber="21">Foveolaria globosa</emphasis>
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<taxonomicNameLabel id="07DC57B26F65FF99CE7AEF348F6EFB43" box="[1053,1112,1147,1173]" pageId="18" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="2E2436DB6F65FF99CA8EEFD28F6EFB0D" blockId="18.[174,1112,266,1348]" pageId="18" pageNumber="21">
The lack of globosity in the species, discussed in this paper, well distinguishes them from
<taxonomicName id="E99B4D586F65FF99CB57EF8D8AACFB0D" authorityName="Szabó" authorityYear="2016" box="[304,410,1217,1243]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="21" phylum="Bryozoa" rank="species" species="globosa">
<emphasis id="1CEFEAC96F65FF99CB57EF8D8AACFB0D" box="[304,410,1217,1243]" italics="true" pageId="18" pageNumber="21">F. globosa</emphasis>
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, however, further details are added also in their descriptions.
</paragraph>
<paragraph id="2E2436DB6F65FF99CA8EEFAB8A3DFA92" blockId="18.[174,1112,266,1348]" pageId="18" pageNumber="21">
<emphasis id="1CEFEAC96F65FF99CA8EEFAB8A41FB28" box="[233,375,1252,1278]" italics="true" pageId="18" pageNumber="21">Distribution</emphasis>
– Within the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone Formation in Hierlatz Alpe, Hallstatt,
<collectingCountry id="568C764B6F65FF99CAC9EE658A31FA92" box="[174,263,1322,1348]" name="Austria" pageId="18" pageNumber="21">Austria</collectingCountry>
.
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<mods:url id="3FF18FD20699148DA3811CEFD5BCADCE">https://doi.org/10.17111/fragmpalhung.2016.33.3</mods:url>
</mods:location>
<mods:classification id="24C07297CC01B5C5E0B44923949AC56F">journal article</mods:classification>
<mods:identifier id="D86E6CD3B11735826906AF9F536AEEC2" type="DOI">10.17111/FragmPalHung.2016.33.3</mods:identifier>
<mods:identifier id="F25FBF20B2D7C2774368965A2CCE6C39" type="ISSN">1586-930X</mods:identifier>
<mods:identifier id="C9EFA734B5CB6AE2784BA9775CDDCD27" type="Zenodo-Dep">15627163</mods:identifier>
</mods:mods>
<treatment id="A63287CD6F71FF82CB8BE8A28A0FFDE8" LSID="urn:lsid:plazi:treatment:A63287CD6F71FF82CB8BE8A28A0FFDE8" httpUri="http://treatment.plazi.org/id/A63287CD6F71FF82CB8BE8A28A0FFDE8" lastPageId="9" lastPageNumber="12" pageId="6" pageNumber="9">
<subSubSection id="668165506F71FF8DCB8BE8A2882CFBDE" box="[492,794,1005,1032]" pageId="6" pageNumber="9" type="nomenclature">
<paragraph id="2E2436DB6F71FF8DCB8BE8A2882CFBDE" blockId="6.[492,794,1005,1067]" box="[492,794,1005,1032]" pageId="6" pageNumber="9">
<heading id="756C81B76F71FF8DCB8BE8A2882CFBDE" box="[492,794,1005,1032]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="9" reason="9">
<taxonomicName id="E99B4D586F71FF8DCB8BE8A289E1FBD1" authorityName="Szabó" authorityYear="2016" box="[492,727,1005,1031]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="6" pageNumber="9" phylum="Mollusca" rank="species" species="rakusi" status="sp. nov.">
<emphasis id="1CEFEAC96F71FF8DCB8BE8A289E1FBD1" box="[492,727,1005,1031]" italics="true" pageId="6" pageNumber="9">Wortheniopsis rakusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F71FF8DC8BAE8A1882CFBDE" box="[733,794,1006,1032]" pageId="6" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="668165506F71FF8DC858EF5E89F0FBFD" box="[575,710,1040,1067]" pageId="6" pageNumber="9" type="description">
<paragraph id="2E2436DB6F71FF8DC858EF5E89F0FBFD" blockId="6.[492,794,1005,1067]" box="[575,710,1040,1067]" pageId="6" pageNumber="9">
(
<figureCitation id="B6A02A5E6F71FF8DC820EF5E8988FBFD" box="[583,702,1040,1067]" captionStart="Figs 4–14" captionStartId="8.[174,214,1437,1459]" captionTargetBox="[174,1111,265,1412]" captionTargetId="figure-18@8.[174,1111,265,1412]" captionTargetPageId="8" captionText="Figs 4–14.Wortheniopsisspecies fromtheHierlatz LimestoneFormation.– 4–10.Wortheniopsisrakusi n.sp.from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe,Hallstatt, Austria,GBA 2017/001/0002,all ×4.– 4. Dorsal view.– 5. Peristomal view.– 6. Basal view. – 7–9. Same views as 4–6 but in natural (×1) size. – 10. Details of the ornament on thespire whorls,×7.– 11–14.Wortheniopsis bakonyensis n. sp.,?Lower Sinemurian (Lower Jurassic) Hierlatz Limestone variety with rich micrite content, Csárdahegy, Úrkút, Bakony Mts, Hungary, GGIH J.08.3.1. – 11. Peristomal view of the holotype, ×4. – 12. Lateral view to show the best preserved outline part, ×4.– 13–14. The same views as in figures 11–12 but in natural (×1) size" figureDoi="http://doi.org/10.5281/zenodo.15627167" httpUri="https://zenodo.org/record/15627167/files/figure.png" pageId="6" pageNumber="9">Figs 4–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="668165506F71FF8DCA8EEF098A54FB1F" pageId="6" pageNumber="9" type="materials_examined">
<materialsCitation id="9EF33C866F71FF8DCA8EEF09882AFB55" country="Austria" location="Hierlatz Alpe" municipality="Hallstatt" pageId="6" pageNumber="9" specimenCount="1" typeStatus="holotype">
<paragraph id="2E2436DB6F71FF8DCA8EEF09899EFBB6" blockId="6.[174,1112,1094,1613]" box="[233,680,1094,1120]" pageId="6" pageNumber="9">
<emphasis id="1CEFEAC96F71FF8DCA8EEF098ABBFB89" box="[233,397,1094,1119]" italics="true" pageId="6" pageNumber="9">
<typeStatus id="F12088796F71FF8DCA8EEF098A17FB89" box="[233,289,1094,1119]" pageId="6" pageNumber="9">Type</typeStatus>
specimen
</emphasis>
– GBA 2017/001/0002.
</paragraph>
<paragraph id="2E2436DB6F71FF8DCA8EEF258816FB55" blockId="6.[174,1112,1094,1613]" box="[233,800,1128,1155]" pageId="6" pageNumber="9">
<emphasis id="1CEFEAC96F71FF8DCA8EEF258A4CFB55" box="[233,378,1129,1155]" italics="true" pageId="6" pageNumber="9">
<typeStatus id="F12088796F71FF8DCA8EEF258A17FB55" box="[233,289,1130,1155]" pageId="6" pageNumber="9">Type</typeStatus>
locality
</emphasis>
–
<location id="2B4460006F71FF8DCBF1EF268901FB55" LSID="urn:lsid:plazi:treatment:A63287CD6F71FF82CB8BE8A28A0FFDE8:2B4460006F71FF8DCBF1EF268901FB55" box="[406,567,1129,1155]" country="Austria" municipality="Hallstatt" name="Hierlatz Alpe" pageId="6" pageNumber="9">Hierlatz Alpe</location>
,
<collectingMunicipality id="CE40ACA16F71FF8DC826EF26899EFB55" box="[577,680,1129,1155]" pageId="6" pageNumber="9">Hallstatt</collectingMunicipality>
(
<collectingCountry id="568C764B6F71FF8DC8D0EF278822FB55" box="[695,788,1128,1155]" name="Austria" pageId="6" pageNumber="9">Austria</collectingCountry>
).
</paragraph>
</materialsCitation>
<paragraph id="2E2436DB6F71FF8DCA8EEFC28A54FB1F" blockId="6.[174,1112,1094,1613]" pageId="6" pageNumber="9">
<materialsCitation id="9EF33C866F71FF8DCA8EEFC28A68FB1F" county="Hierlatz Limestone" location="Sinemurian" municipality="Lower Jurassic" pageId="6" pageNumber="9" specimenCount="1" typeStatus="holotype">
<emphasis id="1CEFEAC96F71FF8DCA8EEFC28A51FB70" box="[233,359,1165,1190]" italics="true" pageId="6" pageNumber="9">
<typeStatus id="F12088796F71FF8DCA8EEFC28A14FB70" box="[233,290,1165,1190]" pageId="6" pageNumber="9">Type</typeStatus>
strata
</emphasis>
–
<location id="2B4460006F71FF8DCBE5EFC3893CFB70" LSID="urn:lsid:plazi:treatment:A63287CD6F71FF82CB8BE8A28A0FFDE8:2B4460006F71FF8DCBE5EFC3893CFB70" box="[386,522,1164,1190]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Sinemurian" pageId="6" pageNumber="9">Sinemurian</location>
to lowermost
<location id="2B4460006F71FF8DC8CBEFC38866FB70" LSID="urn:lsid:plazi:treatment:A63287CD6F71FF82CB8BE8A28A0FFDE8:2B4460006F71FF8DC8CBEFC38866FB70" box="[684,848,1164,1190]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Pliensbachian" pageId="6" pageNumber="9">Pliensbachian</location>
(
<collectingMunicipality id="CE40ACA16F71FF8DC93BEFC38F3DFB71" box="[860,1035,1164,1191]" pageId="6" pageNumber="9">Lower Jurassic</collectingMunicipality>
)
<collectingCounty id="C7454E576F71FF8DCE70EFC38A68FB1F" pageId="6" pageNumber="9">Hierlatz Limestone</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="668165506F71FF8DCA8EEF9C8F0DFB3A" box="[233,1083,1234,1261]" pageId="6" pageNumber="9" type="etymology">
<paragraph id="2E2436DB6F71FF8DCA8EEF9C8F0DFB3A" blockId="6.[174,1112,1094,1613]" box="[233,1083,1234,1261]" pageId="6" pageNumber="9">
<emphasis id="1CEFEAC96F71FF8DCA8EEF9C8AF4FB3A" box="[233,450,1234,1260]" italics="true" pageId="6" pageNumber="9">Derivation of name</emphasis>
– Dedicated to Milos Rakús, late Slovakian colleague.
</paragraph>
</subSubSection>
<subSubSection id="668165506F71FF8DCA8EEFB98904FA80" pageId="6" pageNumber="9" type="materials_examined">
<paragraph id="2E2436DB6F71FF8DCA8EEFB98904FA80" blockId="6.[174,1112,1094,1613]" pageId="6" pageNumber="9">
<emphasis id="1CEFEAC96F71FF8DCA8EEFB98A78FAC6" box="[233,334,1270,1296]" italics="true" pageId="6" pageNumber="9">Material</emphasis>
– Single specimen that consists of four fragmentary whorls; protoconch and the earliest teleoconch whorls are lacking, just like the latest, near peristome parts of the last whorl.
</paragraph>
</subSubSection>
<subSubSection id="668165506F71FF8CCA8EEE2F892DFA5D" lastPageId="7" lastPageNumber="10" pageId="6" pageNumber="9" type="description">
<paragraph id="2E2436DB6F71FF8DCA8EEE2F882DFAAC" blockId="6.[174,1112,1094,1613]" box="[233,795,1375,1402]" pageId="6" pageNumber="9">
<emphasis id="1CEFEAC96F71FF8DCA8EEE2F8A5AFAAF" box="[233,364,1376,1401]" italics="true" pageId="6" pageNumber="9">Dimensions</emphasis>
– Width:
<quantity id="E9639B3E6F71FF8DCB82EE108979FAAC" box="[485,591,1375,1402]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="6" pageNumber="9" unit="mm" value="11.5">11.5 mm</quantity>
; spiral angle: 45°.
</paragraph>
<paragraph id="2E2436DB6F71FF8CCA8EEECC8815FE02" blockId="6.[174,1112,1094,1613]" lastBlockId="7.[219,1157,266,1666]" lastPageId="7" lastPageNumber="10" pageId="6" pageNumber="9">
<emphasis id="1CEFEAC96F71FF8DCA8EEECC8A60FA4A" box="[233,342,1411,1436]" italics="true" pageId="6" pageNumber="9">Diagnosis</emphasis>
– High turbiniform, thin walled shell; slightly convex whorls, shallow impressed suture, last whorl somewhat inflated. Early whorls rounded-angular at both sides of suture, last whorl turning into base along wide, even arch. Last whorl with obscure angulation and slightly concave ramp below adapical suture. Base as whole subconoidal, its wall convex;?narrowly phaneromphalous. Selenizone rather narrow, concave and bordered by two threads above midwhorl on latest whorls. Ornament of spiral threads above selenizone, crossed by collabral threads with granules at intersecting points. Sparse, asymmetrical, weak spiral ridges below selenizone and on base. Growth lines fine, prosocyrt and just gently prosocline above and below selenizone on whorls, feebly opisthocline on base with shallow opisthocyrt part abaxially. Selenizone with sparse lunulae on early whorls, with growth lines only on last whorl.
</paragraph>
<paragraph id="2E2436DB6F70FF8CCB71EA918F23FBD1" blockId="7.[219,1157,266,1666]" pageId="7" pageNumber="10">
<emphasis id="1CEFEAC96F70FF8CCB71EA918AA1FE21" box="[278,407,478,503]" italics="true" pageId="7" pageNumber="10">Description</emphasis>
– A high turbiniform, very thin walled shell represents the species. It is composed of slightly convex whorls, which are separated by a slightly impressed, almost flush suture. The earliest preserved whorls are rounded-angular at both sides of the suture but the last whorl turns into the base without angulation along a wide, even arch. The spire whorls are slightly convex and there is no angulation on their surface but the belt between the selenizone and the adapical suture becomes feebly concave on the last whorl; actually a concave ramp develops. Along the abapical boundary of the ramp, which coincides with the abapical rim of the selenizone, an angulation with the outer face is just observable. The selenizone itself seems flat on the early whorls but concave on the last one; its width is rather narrow, about 12% of the sutures distance on the penultimate whorl. The outer face itself is wider than the ramp and has flat surface that bends into the base along a wide, unbroken arch. The base is convex, subconoidal as a whole and it has convex wall that evenly turns into an axial deepening. Its matrix infilling seems to hide a narrow umbilicus. The selenizone is narrow, concave and two threads border it well above the midwhorl on all preserved whorls.
</paragraph>
<paragraph id="2E2436DB6F70FF8CCB71EF5E892DFA5D" blockId="7.[219,1157,266,1666]" pageId="7" pageNumber="10">
Several equally spaced spiral threads ornament the whole surface adapically from the selenizone that are crossed by collabral threads and granules are at their intersecting points. Towards the last whorl, the number of the growth lines increases. Sparse, asymmetrical, weak spiral ridges, belonging to the
<typeStatus id="F12088796F70FF8CCE67EF348F02FB43" box="[1024,1076,1147,1173]" pageId="7" pageNumber="10">type</typeStatus>
which has been first observed in
<taxonomicName id="E99B4D586F70FF8CC875EFD288E8FB6E" authority="(Stoppani, 1858)" baseAuthorityName="Stoppani" baseAuthorityYear="1858" box="[530,990,1181,1208]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="7" pageNumber="10" phylum="Mollusca" rank="species" species="quirinii">
<emphasis id="1CEFEAC96F70FF8CC875EFD28826FB61" box="[530,784,1181,1207]" italics="true" pageId="7" pageNumber="10">Wortheniopsis quirinii</emphasis>
(Stoppani, 1858)
</taxonomicName>
, see in
<bibRefCitation id="4A0A4B2A6F70FF8CCE51EFD18A17FB0D" author="KITTL E." pageId="7" pageNumber="10" pagination="1 - 237" refId="ref12368" refString="KITTL E. 1899: Die Gastropoden der Esinokalke, nebst einer Revision der Gastropoden der Marmolatakalke. - Annalen des kaiserlich-koniglichen naturhistorishen Hofmuseums 14: 1-237, Pls 1 - 18." type="journal article" year="1899">KITTL (1899</bibRefCitation>
, p. 14,
<figureCitation id="B6A02A5E6F70FF8CCB16EF8E8A81FB0D" box="[369,439,1216,1243]" captionStart="Figs 1–3" captionStartId="3.[219,260,1535,1557]" captionTargetBox="[222,1152,1201,1496]" captionTargetId="figure-324@3.[219,1156,1197,1499]" captionTargetPageId="3" captionText="Figs 1–3. The peculiar “quirinii type” ridges, occurring in Wortheniopsis Böhm, 1895, Foveolaria n. gen. and some probably related forms. – 1. Copy of KITTL’s (1899, p. 14, Fig. 2) sketch. – 2. The ridges on the outer face and on the base of Wortheniopsis rakusi n. sp. – 3. The ridges on the base of Foveolaria conoidea n. sp." figureDoi="http://doi.org/10.5281/zenodo.15627165" httpUri="https://zenodo.org/record/15627165/files/figure.png" pageId="7" pageNumber="10">Fig. 2</figureCitation>
.), ornament the whorls below the selenizone and on the base. Lunulae are present on the selenizone of the early whorls but they are missing on the last whorl. The growth lines are very fine and have a peculiar arrangement; they are prosocyrt and just gently prosocline above and below the selenizone on the whorls and feebly opisthocline on the base with a shallow opisthocyrt part on the abaxial half of the base.
</paragraph>
</subSubSection>
<subSubSection id="668165506F70FF82CB71EEDB8FB3FE02" lastPageId="9" lastPageNumber="12" pageId="7" pageNumber="10" type="discussion">
<paragraph id="2E2436DB6F70FF8CCB71EEDB89A9F954" blockId="7.[219,1157,266,1666]" pageId="7" pageNumber="10">
<emphasis id="1CEFEAC96F70FF8CCB71EEDB8A4CFA78" box="[278,378,1428,1454]" italics="true" pageId="7" pageNumber="10">Remarks</emphasis>
– The only specimen of
<taxonomicName id="E99B4D586F70FF8CC8FEEEDB88B3FA78" authorityName="Szabó" authorityYear="2016" box="[665,901,1428,1454]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="7" pageNumber="10" phylum="Mollusca" rank="species" species="rakusi" status="sp. nov.">
<emphasis id="1CEFEAC96F70FF8CC8FEEEDB88B3FA78" box="[665,901,1428,1454]" italics="true" pageId="7" pageNumber="10">Wortheniopsis rakusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F70FF8CC9EBEEDB88FFFA78" box="[908,969,1428,1454]" pageId="7" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
is exceptionally well preserved compared to the
<typeStatus id="F12088796F70FF8CC831EEF7898AFA04" box="[598,700,1464,1490]" pageId="7" pageNumber="10" type="syntype">syntypes</typeStatus>
of the Middle Triassic type species. In spite of the considerable age difference, their comparison shows only slight differences in the shell shape and the details of the ornament.
<taxonomicName id="E99B4D586F70FF8CC9C6EEB18F26F9C1" authorityName="Szabó" authorityYear="2016" box="[929,1040,1533,1560]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="7" pageNumber="10" phylum="Mollusca" rank="species" species="rakusi" status="n. sp.">
<emphasis id="1CEFEAC96F70FF8CC9C6EEB1888DF9C1" box="[929,955,1534,1559]" italics="true" pageId="7" pageNumber="10">W</emphasis>
.
<emphasis id="1CEFEAC96F70FF8CC9AEEEB28F26F9C1" box="[969,1040,1533,1559]" italics="true" pageId="7" pageNumber="10">rakusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F70FF8CCE70EEB18F60F9CE" box="[1047,1110,1534,1560]" pageId="7" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
has about 10° wider spiral angle, less impressed suture, less convex whorls, so more conoidal outline, and a network ornament with granules at the crossings between the selenizone and the adapical suture.
</paragraph>
<caption id="7AE466536F7FFF83CAC9EED28F0DF9AB" ID-DOI="http://doi.org/10.5281/zenodo.15627167" ID-Zenodo-Dep="15627167" httpUri="https://zenodo.org/record/15627167/files/figure.png" pageId="8" pageNumber="11" startId="8.[174,214,1437,1459]" targetBox="[174,1111,265,1412]" targetPageId="8" targetType="figure">
<paragraph id="2E2436DB6F7FFF83CAC9EED28F0DF9AB" blockId="8.[174,1112,1437,1661]" pageId="8" pageNumber="11">
<emphasis id="1CEFEAC96F7FFF83CAC9EED28A24FA64" bold="true" box="[174,274,1437,1459]" pageId="8" pageNumber="11">Figs 4–14.</emphasis>
<taxonomicName id="E99B4D586F7FFF83CB72EED28AE1FA65" box="[277,471,1437,1459]" pageId="8" pageNumber="11">
<emphasis id="1CEFEAC96F7FFF83CB72EED28AA5FA64" box="[277,403,1437,1458]" italics="true" pageId="8" pageNumber="11">Wortheniopsis</emphasis>
species
</taxonomicName>
from the Hierlatz Limestone Formation.–
<emphasis id="1CEFEAC96F7FFF83C904EED288ADFA64" bold="true" box="[867,923,1437,1458]" pageId="8" pageNumber="11">4–10.</emphasis>
<taxonomicName id="E99B4D586F7FFF83C9F9EED28F61FA64" authorityName="Szabó" authorityYear="2016" box="[926,1111,1437,1458]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="8" pageNumber="11" phylum="Mollusca" rank="species" species="rakusi" status="sp. nov.">
<emphasis id="1CEFEAC96F7FFF83C9F9EED28F61FA64" box="[926,1111,1437,1458]" italics="true" pageId="8" pageNumber="11">Wortheniopsis rakusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7FFF83CAC9EEF48BE9FA06" box="[174,223,1467,1488]" pageId="8" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria, GBA 2017/001/0002, all ×4. –
<emphasis id="1CEFEAC96F7FFF83C90BEE988849FA3A" bold="true" box="[876,895,1495,1516]" pageId="8" pageNumber="11">4.</emphasis>
Dorsal view. –
<emphasis id="1CEFEAC96F7FFF83CE6CEE988F28FA3A" bold="true" box="[1035,1054,1495,1516]" pageId="8" pageNumber="11">5.</emphasis>
Peristomal view. –
<emphasis id="1CEFEAC96F7FFF83CB57EEBB8A75F9DF" bold="true" box="[304,323,1524,1545]" pageId="8" pageNumber="11">6.</emphasis>
Basal view. –
<emphasis id="1CEFEAC96F7FFF83CBA2EEBB8AC7F9DF" bold="true" box="[453,497,1524,1545]" pageId="8" pageNumber="11">7–9.</emphasis>
Same views as 4–6 but in natural (×1) size. –
<emphasis id="1CEFEAC96F7FFF83C9F1EEBB8883F9DF" bold="true" box="[918,949,1524,1545]" pageId="8" pageNumber="11">10.</emphasis>
Details of the ornament on the spire whorls, ×7. –
<emphasis id="1CEFEAC96F7FFF83CB8EED5E8918F9F0" bold="true" box="[489,558,1553,1574]" pageId="8" pageNumber="11">11–14.</emphasis>
<taxonomicName id="E99B4D586F7FFF83C856ED5E882BF9F0" authorityName="Szabó" authorityYear="2016" box="[561,797,1553,1574]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="8" pageNumber="11" phylum="Mollusca" rank="species" species="bakonyensis" status="sp. nov.">
<emphasis id="1CEFEAC96F7FFF83C856ED5E882BF9F0" box="[561,797,1553,1574]" italics="true" pageId="8" pageNumber="11">Wortheniopsis bakonyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7FFF83C945ED5E8862F9F0" box="[802,852,1553,1574]" pageId="8" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
,?Lower Sinemurian (Lower Jurassic) Hierlatz Limestone
<taxonomicName id="E99B4D586F7FFF83CBDCED62891DF995" box="[443,555,1581,1603]" pageId="8" pageNumber="11" rank="variety" variety="with">variety with</taxonomicName>
rich micrite content, Csárdahegy, Úrkút, Bakony Mts, Hungary, GGIH J.08.3.1. –
<emphasis id="1CEFEAC96F7FFF83CBEFED058A91F989" bold="true" box="[392,423,1610,1631]" pageId="8" pageNumber="11">11.</emphasis>
Peristomal view of the holotype, ×4. –
<emphasis id="1CEFEAC96F7FFF83C979ED05880BF989" bold="true" box="[798,829,1610,1631]" pageId="8" pageNumber="11">12.</emphasis>
Lateral view to show the best preserved outline part, ×4. –
<emphasis id="1CEFEAC96F7FFF83CBBFED28892BF9AA" bold="true" box="[472,541,1639,1660]" pageId="8" pageNumber="11">13–14.</emphasis>
The same views as in figures 11–12 but in natural (×1) size
</paragraph>
</caption>
<paragraph id="2E2436DB6F7EFF82CB71EA448FB3FE67" blockId="9.[219,1157,266,574]" pageId="9" pageNumber="12">
<taxonomicName id="E99B4D586F7EFF82CB71EA448AB3FEF2" authorityName="Szabó" authorityYear="2016" box="[278,389,266,293]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="rakusi" status="n. sp.">
<emphasis id="1CEFEAC96F7EFF82CB71EA448A06FEF2" box="[278,304,267,292]" italics="true" pageId="9" pageNumber="12">W</emphasis>
.
<emphasis id="1CEFEAC96F7EFF82CB59EA458AB3FEF2" box="[318,389,266,292]" italics="true" pageId="9" pageNumber="12">rakusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7EFF82CBEBEA448AFDFEF3" box="[396,459,267,293]" pageId="9" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
clearly shows that the growth lines and the collabral ornamental elements terminate at the threads, delimiting the anal fasciole, which indicate a true selenizone, not a sinuszone. Lunulae on the penultimate whorl and lunuliform growth lines on the last whorl endorse this assumption. Both species have quirinii
<typeStatus id="F12088796F7EFF82CB12EAD78A9FFE64" box="[373,425,408,434]" pageId="9" pageNumber="12">type</typeStatus>
ridges between the selenizone and the axial region of the base.
</paragraph>
<paragraph id="2E2436DB6F7EFF82CB71EAF48FB3FE02" blockId="9.[219,1157,266,574]" box="[278,1157,442,469]" pageId="9" pageNumber="12">
For distinction from
<taxonomicName id="E99B4D586F7EFF82C861EAF5881CFE02" authorityName="Szabó" authorityYear="2016" box="[518,810,442,468]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="bakonyensis" status="sp. nov.">
<emphasis id="1CEFEAC96F7EFF82C861EAF5881CFE02" box="[518,810,442,468]" italics="true" pageId="9" pageNumber="12">Wortheniopsis bakonyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7EFF82C948EAF4885DFE03" box="[815,875,443,469]" pageId="9" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
see the next description.
</paragraph>
</subSubSection>
<subSubSection id="668165506F7EFF82CB71EA928A0FFDE8" pageId="9" pageNumber="12" type="distribution">
<paragraph id="2E2436DB6F7EFF82CB71EA928A0FFDE8" blockId="9.[219,1157,266,574]" pageId="9" pageNumber="12">
<emphasis id="1CEFEAC96F7EFF82CB71EA928A92FE21" box="[278,420,477,503]" italics="true" pageId="9" pageNumber="12">Distribution</emphasis>
– Within Sinemurian to lowermost Pliensbachian (Lower Jurassic) part of the Hierlatz Limestone Formation in Hierlatz Alpe, Hallstatt,
<collectingCountry id="568C764B6F7EFF82CABCE96B8A02FDE8" box="[219,308,548,574]" name="Austria" pageId="9" pageNumber="12">Austria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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<mods:title id="C8CA08505C0142649C8F30CD4CD67FA9">Gastropods of the Lower Jurassic Hierlatz Limestone Formation, part 2. Some new archaic type slit-bearing components from the fauna of the Hierlatz Alpe (Hallstatt, Austria) and the Bakony Mts (Hungary)</mods:title>
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<subSubSection id="668165506F73FF8FCBD1E82E8879FCAA" box="[438,847,865,892]" pageId="4" pageNumber="7" type="nomenclature">
<paragraph id="2E2436DB6F73FF8FCBD1E82E8879FCAA" blockId="4.[187,1098,689,962]" box="[438,847,865,892]" pageId="4" pageNumber="7">
<heading id="756C81B76F73FF8FCBD1E82E8879FCAA" box="[438,847,865,892]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="7" reason="2">
Genus
<taxonomicName id="E99B4D586F73FF8FC860E82E8879FCAA" ID-CoL="7PL4R" authority="J. Bohm, 1895" authorityName="J. Bohm" authorityYear="1895" box="[519,847,865,892]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="4" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F73FF8FC860E82E8993FCAD" box="[519,677,865,891]" italics="true" pageId="4" pageNumber="7">Wortheniopsis</emphasis>
J.
<bibRefCitation id="4A0A4B2A6F73FF8FC8A5E82E8879FCAA" author="BOHM J." box="[706,847,865,892]" pageId="4" pageNumber="7" pagination="211 - 308" refId="ref11860" refString="BOHM J. 1895: Die Gastropoden des Marmolatakalkes. - Palaeontographica 42: 211-308." type="journal article" year="1895">Böhm, 1895</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="668165506F73FF8DCADCE8CB8876FC14" lastPageId="6" lastPageNumber="9" pageId="4" pageNumber="7" type="materials_examined">
<paragraph id="2E2436DB6F73FF8FCADCE8CB8817FC14" blockId="4.[187,1098,689,962]" pageId="4" pageNumber="7">
<typeStatus id="F12088796F73FF8FCADCE8CB8BC1FC48" box="[187,247,900,926]" pageId="4" pageNumber="7">Type</typeStatus>
species –
<taxonomicName id="E99B4D586F73FF8FCB0BE8CB881BFC48" authority="Kittl, 1894" authorityName="Kittl" authorityYear="1894" box="[364,813,900,926]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="4" pageNumber="7" phylum="Mollusca" rank="species" species="margarethae">
<emphasis id="1CEFEAC96F73FF8FCB0BE8CB8991FC48" box="[364,679,900,926]" italics="true" pageId="4" pageNumber="7">Pleurotomaria margarethae</emphasis>
<bibRefCitation id="4A0A4B2A6F73FF8FC8CAE8CB881BFC48" author="KITTL E." box="[685,813,900,926]" pageId="4" pageNumber="7" pagination="99 - 182" refId="ref12329" refString="KITTL E. 1894: Die Triadischen Gastropoden der Marmolata und verwandter Fundstellen in der weissen Riffkalken Sudtirols. - Jahrbuch der kaiserlich-koniglichen Geologischen Reichsanstalt 44 (1): 99-182, Pls 1 - 6." type="journal article" year="1894">Kittl, 1894</bibRefCitation>
</taxonomicName>
, Marmolada Limestone, Ladinian (Middle Triassic)
</paragraph>
<paragraph id="2E2436DB6F73FF8FCA8EE8938AADFA35" blockId="4.[174,1112,988,1648]" pageId="4" pageNumber="7">
<emphasis id="1CEFEAC96F73FF8FCA8EE8938AF0FC20" box="[233,454,988,1014]" italics="true" pageId="4" pageNumber="7">Emended diagnosis</emphasis>
– Moderately high turbiniform to subconoidal, thin walled shells of convex whorls. Earliest shell parts unknown. Suture variably impressed. Surface of early whorls evenly arched, later whorls having obscure spiral angulation adapically from midwhorl. Flat to concave ramp between upper suture and angulation. Flat or feebly concave selenizone between two spiral threads, abapical one just overlapping ridge of angulation. Feebly concave to feebly convex outer face, being wider than ramp. Transition from outer face to base widely and evenly arched. Base subglobose as whole with convex wall, anomphalous or narrowly phaneromphalous. Aperture rounded with angle at terminus of suture. Thin, smooth shell layer representing parietal lip; columellar/umbilical lip vertical, thin or weakly thickened. Ornament of spiral and collabral threads and granules on whorls, threadlike spiral ridges of asymmetric triangle cross-section on base and on outer face (quirinii
<typeStatus id="F12088796F73FF8FC844EECC8961FA4B" box="[547,599,1411,1437]" pageId="4" pageNumber="7">type</typeStatus>
ridges). Growth lines feebly prosocline and fairly prosocyrt on both sides of selenizone, respectively, feebly opisthocline and opisthocyrt on base.
</paragraph>
<paragraph id="2E2436DB6F73FF8ECA8EEEA3898FFEBD" blockId="4.[174,1112,988,1648]" lastBlockId="5.[219,1157,266,1630]" lastPageId="5" lastPageNumber="8" pageId="4" pageNumber="7">
<emphasis id="1CEFEAC96F73FF8FCA8EEEA38A7BF9D0" box="[233,333,1516,1542]" italics="true" pageId="4" pageNumber="7">Remarks</emphasis>
– All
<typeStatus id="F12088796F73FF8FCBF3EEA28ACCF9D1" box="[404,506,1517,1543]" pageId="4" pageNumber="7" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="E99B4D586F73FF8FC878EEA38862F9D0" box="[543,852,1516,1542]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="4" pageNumber="7" phylum="Mollusca" rank="species" species="margarethae">
<emphasis id="1CEFEAC96F73FF8FC878EEA38862F9D0" box="[543,852,1516,1542]" italics="true" pageId="4" pageNumber="7">Wortheniopsis margarethae</emphasis>
</taxonomicName>
are of very poor preservation, therefore the nature of the exhalant system, critical for classification, has remained debatable for the recently active researchers of the Triassic gastropods. The original descriptions and drawings do not provide satisfactory help to see doubtlessly if the type species has a slit and selenizone or only peristomal sinus and sinuszone on the whorls. Therefore a re-study of the specimens has become necessary for further discussion.
</paragraph>
<paragraph id="2E2436DB6F72FF8ECB71EA3B8AFBFC48" blockId="5.[219,1157,266,1630]" pageId="5" pageNumber="8">
The restudy of the
<typeStatus id="F12088796F72FF8ECB9AEA3B8952FE58" box="[509,612,372,398]" pageId="5" pageNumber="8" type="syntype">syntypes</typeStatus>
(
<specimenCode id="7E3D9EA06F72FF8EC813EA3B882FFE59" box="[628,793,372,399]" collectionCode="NHMV" pageId="5" pageNumber="8">NHMV 1969</specimenCode>
/1103) has confirmed, as their figures (
<bibRefCitation id="4A0A4B2A6F72FF8ECB5CEAD88AE4FE67" author="KITTL E." box="[315,466,407,433]" pageId="5" pageNumber="8" pagination="99 - 182" refId="ref12329" refString="KITTL E. 1894: Die Triadischen Gastropoden der Marmolata und verwandter Fundstellen in der weissen Riffkalken Sudtirols. - Jahrbuch der kaiserlich-koniglichen Geologischen Reichsanstalt 44 (1): 99-182, Pls 1 - 6." type="journal article" year="1894">KITTL 1894</bibRefCitation>
, Pl.
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,
<figureCitation id="B6A02A5E6F72FF8EC853EAD889ADFE64" box="[564,667,407,434]" captionStart="Figs 1–3" captionStartId="3.[219,260,1535,1557]" captionTargetBox="[222,1152,1201,1496]" captionTargetId="figure-324@3.[219,1156,1197,1499]" captionTargetPageId="3" captionText="Figs 1–3. The peculiar “quirinii type” ridges, occurring in Wortheniopsis Böhm, 1895, Foveolaria n. gen. and some probably related forms. – 1. Copy of KITTL’s (1899, p. 14, Fig. 2) sketch. – 2. The ridges on the outer face and on the base of Wortheniopsis rakusi n. sp. – 3. The ridges on the base of Foveolaria conoidea n. sp." figureDoi="http://doi.org/10.5281/zenodo.15627165" httpUri="https://zenodo.org/record/15627165/files/figure.png" pageId="5" pageNumber="8">Figs 1–3</figureCitation>
) also show, that even the best preserved
<typeStatus id="F12088796F72FF8ECABCEAF48A01FE03" box="[219,311,443,469]" pageId="5" pageNumber="8" type="syntype">syntype</typeStatus>
of the type species bears only very small pieces of badly worn shell fragments on inner mould and their outermost layer is lacking. Actually, the lines of the ornament are much more obscure than the drawings show them. A narrow, smooth and flattish ribbon between two eroded spiral threads represents the anal fasciole just adapically from an obscure angulation of the whorl surface, aligned to the abapical spiral thread. Some distinct lines of KITTL’s drawing (1894, Pl.
<collectionCode id="488AAE1E6F72FF8ECB37E9C18A47FD7E" box="[336,369,654,680]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="8">VI</collectionCode>
,
<figureCitation id="B6A02A5E6F72FF8ECB1CE9C28A8BFD7E" box="[379,445,653,680]" captionStart="Figs 1–3" captionStartId="3.[219,260,1535,1557]" captionTargetBox="[222,1152,1201,1496]" captionTargetId="figure-324@3.[219,1156,1197,1499]" captionTargetPageId="3" captionText="Figs 1–3. The peculiar “quirinii type” ridges, occurring in Wortheniopsis Böhm, 1895, Foveolaria n. gen. and some probably related forms. – 1. Copy of KITTL’s (1899, p. 14, Fig. 2) sketch. – 2. The ridges on the outer face and on the base of Wortheniopsis rakusi n. sp. – 3. The ridges on the base of Foveolaria conoidea n. sp." figureDoi="http://doi.org/10.5281/zenodo.15627165" httpUri="https://zenodo.org/record/15627165/files/figure.png" pageId="5" pageNumber="8">Fig. 2</figureCitation>
) show that the growth lines of the subsutural ramp and those of the outer face form a sharp angle at the ridge of the obscure angulation i.e. at the abapical spiral thread. Actually, those few lines that cross the belt between the two spiral threads are straight linear damages and are not aligned to the growth-lines. Waste majority of the collabral lines terminate at or before reaching the two spiral threads, limiting the smooth belt that must be therefore a selenizone. The selenizone is smooth like in the latest growth phases of the two new Jurassic species.
</paragraph>
<paragraph id="2E2436DB6F72FF8ECB71E8E889DBFB42" blockId="5.[219,1157,266,1630]" pageId="5" pageNumber="8">
Similarly to
<taxonomicName id="E99B4D586F72FF8ECBD1E8E889CDFC17" authorityName="Kittl" authorityYear="1894" box="[438,763,935,961]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="5" pageNumber="8" phylum="Mollusca" rank="species" species="margarethae">
<emphasis id="1CEFEAC96F72FF8ECBD1E8E889CDFC17" box="[438,763,935,961]" italics="true" pageId="5" pageNumber="8">Pleurotomaria margarethae</emphasis>
</taxonomicName>
,
<taxonomicName id="E99B4D586F72FF8EC977E8E88FB2FC14" authority="Stoppani, 1858" authorityName="Stoppani" authorityYear="1858" box="[784,1156,935,962]" class="Gastropoda" family="Turbinidae" genus="Turbo" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="8" phylum="Mollusca" rank="species" species="quirinii">
<emphasis id="1CEFEAC96F72FF8EC977E8E88888FC17" box="[784,958,935,961]" italics="true" pageId="5" pageNumber="8">Turbo quirinii</emphasis>
Stoppani, 1858
</taxonomicName>
and
<taxonomicName id="E99B4D586F72FF8ECB72E8858998FC33" authority="Stoppani, 1858" authorityName="Stoppani" authorityYear="1858" box="[277,686,970,997]" class="Gastropoda" family="Turbinidae" genus="Turbo" kingdom="Animalia" order="Trochida" pageId="5" pageNumber="8" phylum="Mollusca" rank="species" species="funiculatus">
<emphasis id="1CEFEAC96F72FF8ECB72E8858ADDFC32" box="[277,491,970,996]" italics="true" pageId="5" pageNumber="8">Turbo funiculatus</emphasis>
Stoppani, 1858
</taxonomicName>
occurred in the same formation, the Marmolada Limestone (Ladinian, Middle Triassic). First
<bibRefCitation id="4A0A4B2A6F72FF8EC918E8A18F71FBDE" author="MARIANI E." box="[895,1095,1005,1032]" pageId="5" pageNumber="8" pagination="113 - 137" refId="ref12569" refString="MARIANI E. 1896: Appunti di paleontologia Lombarda. - Atti della Societa italiana di scienze naturali e del Museo civico di storia naturale di Milano 36: 113-137, 3 Pls." type="book chapter" year="1896">MARIANI (1896)</bibRefCitation>
then
<bibRefCitation id="4A0A4B2A6F72FF8ECABCEF5E8A48FBFD" author="KITTL E." box="[219,382,1040,1067]" pageId="5" pageNumber="8" pagination="1 - 237" refId="ref12368" refString="KITTL E. 1899: Die Gastropoden der Esinokalke, nebst einer Revision der Gastropoden der Marmolatakalke. - Annalen des kaiserlich-koniglichen naturhistorishen Hofmuseums 14: 1-237, Pls 1 - 18." type="journal article" year="1899">KITTL (1899)</bibRefCitation>
regarded these species also as
<taxonomicName id="E99B4D586F72FF8EC883EF5E88B4FBFD" authorityName="J. Bohm" authorityYear="1895" box="[740,898,1041,1067]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="5" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F72FF8EC883EF5E88B4FBFD" box="[740,898,1041,1067]" italics="true" pageId="5" pageNumber="8">Wortheniopsis</emphasis>
</taxonomicName>
. The latter author has also unified the two latter species, however, as
<bibRefCitation id="4A0A4B2A6F72FF8EC96DEF7B8890FB98" author="KITTL E." box="[778,934,1076,1102]" pageId="5" pageNumber="8" pagination="1 - 237" refId="ref12368" refString="KITTL E. 1899: Die Gastropoden der Esinokalke, nebst einer Revision der Gastropoden der Marmolatakalke. - Annalen des kaiserlich-koniglichen naturhistorishen Hofmuseums 14: 1-237, Pls 1 - 18." type="journal article" year="1899">KITTL (1899</bibRefCitation>
, Pl. 14–15) figures show it, the interpretation of this species involves so significantly different morphologies that it might belong to two species.
</paragraph>
<paragraph id="2E2436DB6F72FF8ECB71EFD28A0BFAF4" blockId="5.[219,1157,266,1630]" pageId="5" pageNumber="8">
<collectorName id="836E530D6F72FF8ECB71EFD28AA7FB61" box="[278,401,1181,1207]" pageId="5" pageNumber="8">According</collectorName>
to unpublished data of
<collectorName id="836E530D6F72FF8EC8DCEFD18804FB61" box="[699,818,1181,1208]" pageId="5" pageNumber="8">S. Monari</collectorName>
(Padua University), slitless, sinuszone bearing gastropod with morphology similar to the
<typeStatus id="F12088796F72FF8EC9B7EF8E8F32FB0D" box="[976,1028,1217,1243]" pageId="5" pageNumber="8">type</typeStatus>
species of
<taxonomicName id="E99B4D586F72FF8ECABCEFAB8A4FFB28" authorityName="J. Bohm" authorityYear="1895" box="[219,377,1252,1278]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="5" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F72FF8ECABCEFAB8A4FFB28" box="[219,377,1252,1278]" italics="true" pageId="5" pageNumber="8">Wortheniopsis</emphasis>
</taxonomicName>
occurs in the fauna of the Marmolada Limestone (
<collectionCode id="488AAE1E6F72FF8EC9B5EFAB8F74FB28" box="[978,1090,1252,1278]" pageId="5" pageNumber="8">MONARI</collectionCode>
, pers. comm.).
</paragraph>
<paragraph id="2E2436DB6F72FF8ECB71EE658A88FA78" blockId="5.[219,1157,266,1630]" pageId="5" pageNumber="8">
The Late Triassic
<taxonomicName id="E99B4D586F72FF8ECB84EE6588A4FA92" authority="Kutassy, 1927" authorityName="Kutassy" authorityYear="1927" box="[483,914,1322,1349]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="5" pageNumber="8" phylum="Mollusca" rank="species" species="budensis">
<emphasis id="1CEFEAC96F72FF8ECB84EE6589DCFA92" box="[483,746,1322,1348]" italics="true" pageId="5" pageNumber="8">Wortheniopsis budensis</emphasis>
Kutassy, 1927
</taxonomicName>
, in lack of any traces of the exhalant system, is excluded from the comparisons. The name
<taxonomicName id="E99B4D586F72FF8EC981EE028FB2FAB1" authorityName="J. Bohm" authorityYear="1895" box="[998,1156,1357,1383]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="5" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F72FF8EC981EE028FB2FAB1" box="[998,1156,1357,1383]" italics="true" pageId="5" pageNumber="8">Wortheniopsis</emphasis>
</taxonomicName>
has also been applied to some Palaeozoic forms but their revision is out of the scope of this paper.
</paragraph>
<paragraph id="2E2436DB6F72FF8ECB71EEF88ABFF988" blockId="5.[219,1157,266,1630]" pageId="5" pageNumber="8">
The emended diagnosis above is based on the study of the
<typeStatus id="F12088796F72FF8EC9B6EEF78F01FA04" box="[977,1079,1464,1490]" pageId="5" pageNumber="8" type="syntype">syntypes</typeStatus>
of the type species,
<taxonomicName id="E99B4D586F72FF8ECB10EE95880DFA22" authority="Kittl, 1894" authorityName="Kittl" authorityYear="1894" box="[375,827,1498,1524]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="5" pageNumber="8" phylum="Mollusca" rank="species" species="margarethae">
<emphasis id="1CEFEAC96F72FF8ECB10EE958982FA22" box="[375,692,1498,1524]" italics="true" pageId="5" pageNumber="8">Pleurotomaria margarethae</emphasis>
<bibRefCitation id="4A0A4B2A6F72FF8EC8DCEE95880DFA22" author="KITTL E." box="[699,827,1498,1524]" pageId="5" pageNumber="8" pagination="99 - 182" refId="ref12329" refString="KITTL E. 1894: Die Triadischen Gastropoden der Marmolata und verwandter Fundstellen in der weissen Riffkalken Sudtirols. - Jahrbuch der kaiserlich-koniglichen Geologischen Reichsanstalt 44 (1): 99-182, Pls 1 - 6." type="journal article" year="1894">Kittl, 1894</bibRefCitation>
</taxonomicName>
and monotypes of two new Jurassic species,
<taxonomicName id="E99B4D586F72FF8ECBF9EEB289BDF9C1" authorityName="Szabó" authorityYear="2016" box="[414,651,1533,1559]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="5" pageNumber="8" phylum="Mollusca" rank="species" species="rakusi" status="sp. nov.">
<emphasis id="1CEFEAC96F72FF8ECBF9EEB289BDF9C1" box="[414,651,1533,1559]" italics="true" pageId="5" pageNumber="8">Wortheniopsis rakusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F72FF8EC8F3EEB189E5F9CE" box="[660,723,1534,1560]" pageId="5" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="E99B4D586F72FF8EC977EEB28F0BF9C1" authorityName="Szabó" authorityYear="2016" box="[784,1085,1533,1559]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="5" pageNumber="8" phylum="Mollusca" rank="species" species="bakonyensis" status="sp. nov.">
<emphasis id="1CEFEAC96F72FF8EC977EEB28F0BF9C1" box="[784,1085,1533,1559]" italics="true" pageId="5" pageNumber="8">Wortheniopsis bakonyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F72FF8ECE21EEB18FB3F9CE" box="[1094,1157,1534,1560]" pageId="5" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
The morphology of these three species is similar to each other so they seem to be closely related.
</paragraph>
<paragraph id="2E2436DB6F71FF8DCA8EEA45886AFCAD" blockId="6.[174,1112,266,962]" pageId="6" pageNumber="9">
The morphology of the growth lines in the new specimens permits to reconstruct elements of the outer lip that has not yet been found in preserved form. It is just very feebly prosocline, almost orthocline as a whole and composed of prosocyrt parts at both sides of the selenizone, respectively, and feebly opisthocyrt on the base. The growth lines meet the selenizone in 50–60° angles at its both sides. It probably means that the rim of the outer lip is angular at both sides of a peristomal fissure, like in
<taxonomicName id="E99B4D586F71FF8DCB3EEA928ACCFE21" authorityName="Gray" authorityYear="1847" box="[345,506,477,503]" class="Gastropoda" family="Scissurellidae" kingdom="Animalia" order="Lepetellida" pageId="6" pageNumber="9" phylum="Mollusca" rank="family">Scissurellidae</taxonomicName>
. This morphology is different in Pleurotomarioidea where a wide, strongly prosocline-prosocyrt growth line belt is characteristic adapically from the selenizone, prosocyrt and narrow belt between the base and the selenizone, and markedly parasigmoidal on the base. Furthermore, these growth lines asymptotically approach the selenizone, i.e. the rim of the peristome is widely arched at both sides of the slit. These differences support the non-pleurotomarioidean systematic position for
<emphasis id="1CEFEAC96F71FF8DC8EEE9FE8818FD1D" box="[649,814,689,715]" italics="true" pageId="6" pageNumber="9">
<taxonomicName id="E99B4D586F71FF8DC8EEE9FE881FFD1D" authorityName="J. Bohm" authorityYear="1895" box="[649,809,689,715]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="6" pageNumber="9" phylum="Mollusca" rank="genus">Wortheniopsis</taxonomicName>
.
</emphasis>
The sinus-bearing counterpart (“
<taxonomicName id="E99B4D586F71FF8DCB7BE99B8908FD39" authority="Stoppani" authorityName="Stoppani" authorityYear="1858" box="[284,574,724,751]" class="Gastropoda" family="Turbinidae" genus="Turbo" kingdom="Animalia" order="Trochida" pageId="6" pageNumber="9" phylum="Mollusca" rank="species" species="quirinii">
<emphasis id="1CEFEAC96F71FF8DCB7BE99B8A54FD38" box="[284,354,724,750]" italics="true" pageId="6" pageNumber="9">Turbo</emphasis>
”
<emphasis id="1CEFEAC96F71FF8DCB12E99A8AFAFD38" box="[373,460,725,750]" italics="true" pageId="6" pageNumber="9">quirinii</emphasis>
Stoppani
</taxonomicName>
) in the Middle Triassic seems to represent an earlier evolutionary stage, and suggests belonging to one of the archaic higher taxa. Although similar situation occurs also in some other family rank taxa, e.g. in
<taxonomicName id="E99B4D586F71FF8DCAC9E8728A4EFC81" authorityName="Koken" authorityYear="1896" box="[174,376,829,855]" class="Gastropoda" family="Raphistomatidae" kingdom="Animalia" pageId="6" pageNumber="9" phylum="Mollusca" rank="family">Raphistomatidae</taxonomicName>
, containing also genera of similar shell morphology, tentatively
<taxonomicName id="E99B4D586F71FF8DCAC9E82E8A0AFCAD" authorityName="Wenz" authorityYear="1938" box="[174,316,865,891]" class="Gastropoda" family="Pithodeidae" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="6" pageNumber="9" phylum="Mollusca" rank="family">Pithodeidae</taxonomicName>
is selected as family position for
<taxonomicName id="E99B4D586F71FF8DC8DFE82E8862FCAD" authorityName="J. Bohm" authorityYear="1895" box="[696,852,865,891]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="6" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F71FF8DC8DFE82E8862FCAD" box="[696,852,865,891]" italics="true" pageId="6" pageNumber="9">Wortheniopsis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="2E2436DB6F71FF8DCA8EE8CB8876FC14" blockId="6.[174,1112,266,962]" pageId="6" pageNumber="9">
<emphasis id="1CEFEAC96F71FF8DCA8EE8CB8A41FC48" box="[233,375,900,926]" italics="true" pageId="6" pageNumber="9">Distribution</emphasis>
– Beside the Alpine Middle Triassic (Marmolada Limestone) here verified in the Lower Jurassic (Hierlatz Limestone).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="A1A6E281E9863BA4D4151BA95DCA5567">Gastropods of the Lower Jurassic Hierlatz Limestone Formation, part 2. Some new archaic type slit-bearing components from the fauna of the Hierlatz Alpe (Hallstatt, Austria) and the Bakony Mts (Hungary)</mods:title>
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<paragraph id="2E2436DB6F79FF85CB97EE958823FA23" blockId="14.[496,789,1498,1560]" box="[496,789,1498,1525]" pageId="14" pageNumber="17">
<heading id="756C81B76F79FF85CB97EE958823FA23" box="[496,789,1498,1525]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="17" reason="9">
<taxonomicName id="E99B4D586F79FF85CB97EE9589E4FA22" authorityName="Szabó" authorityYear="2016" box="[496,722,1498,1524]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="17" phylum="Bryozoa" rank="species" species="conoidea" status="sp. nov.">
<emphasis id="1CEFEAC96F79FF85CB97EE9589E4FA22" box="[496,722,1498,1524]" italics="true" pageId="14" pageNumber="17">Foveolaria conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F79FF85C8BFEE948823FA23" box="[728,789,1499,1525]" pageId="14" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="668165506F79FF9AC85FEEB28A0FFC32" lastPageId="17" lastPageNumber="20" pageId="14" pageNumber="17" type="description">
<paragraph id="2E2436DB6F79FF85C85FEEB289FBF9CE" blockId="14.[496,789,1498,1560]" box="[568,717,1533,1560]" pageId="14" pageNumber="17">
(
<figureCitation id="B6A02A5E6F79FF85C827EEB289F3F9CE" box="[576,709,1533,1560]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="14" pageNumber="17">Figs 19–22</figureCitation>
)
</paragraph>
<materialsCitation id="9EF33C866F79FF85CA8EED7C882AF9A6" country="Austria" location="Hierlatz Alpe" municipality="Hallstatt" pageId="14" pageNumber="17" specimenCount="1" typeStatus="holotype">
<paragraph id="2E2436DB6F79FF85CA8EED7C899EF99B" blockId="14.[233,800,1587,1648]" box="[233,680,1587,1613]" pageId="14" pageNumber="17">
<emphasis id="1CEFEAC96F79FF85CA8EED7C8ABBF99A" box="[233,397,1587,1612]" italics="true" pageId="14" pageNumber="17">
<typeStatus id="F12088796F79FF85CA8EED7C8A17F99A" box="[233,289,1587,1612]" pageId="14" pageNumber="17">Type</typeStatus>
specimen
</emphasis>
– GBA 2017/001/0004.
</paragraph>
<paragraph id="2E2436DB6F79FF85CA8EED198816F9A6" blockId="14.[233,800,1587,1648]" box="[233,800,1621,1648]" pageId="14" pageNumber="17">
<emphasis id="1CEFEAC96F79FF85CA8EED198A4CF9A6" box="[233,378,1622,1648]" italics="true" pageId="14" pageNumber="17">
<typeStatus id="F12088796F79FF85CA8EED198A17F9B9" box="[233,289,1622,1647]" pageId="14" pageNumber="17">Type</typeStatus>
locality
</emphasis>
–
<location id="2B4460006F79FF85CBF1ED198901F9A6" LSID="urn:lsid:plazi:treatment:A63287CD6F79FF9ACB97EE958A0FFC32:2B4460006F79FF85CBF1ED198901F9A6" box="[406,567,1622,1648]" country="Austria" municipality="Hallstatt" name="Hierlatz Alpe" pageId="14" pageNumber="17">Hierlatz Alpe</location>
,
<collectingMunicipality id="CE40ACA16F79FF85C826ED19899EF9A6" box="[577,680,1622,1648]" pageId="14" pageNumber="17">Hallstatt</collectingMunicipality>
(
<collectingCountry id="568C764B6F79FF85C8D0ED1A8822F9A6" box="[695,788,1621,1648]" name="Austria" pageId="14" pageNumber="17">Austria</collectingCountry>
).
</paragraph>
</materialsCitation>
<paragraph id="2E2436DB6F78FF84CB71EA448AB9FE91" blockId="15.[219,1157,266,1383]" pageId="15" pageNumber="18">
<materialsCitation id="9EF33C866F78FF84CB71EA448ABAFE91" county="Hierlatz Limestone" location="Sinemurian" municipality="Lower Jurassic" pageId="15" pageNumber="18" specimenCount="1" typeStatus="holotype">
<emphasis id="1CEFEAC96F78FF84CB71EA448AA3FEF2" box="[278,405,267,292]" italics="true" pageId="15" pageNumber="18">
<typeStatus id="F12088796F78FF84CB71EA448A79FEF2" box="[278,335,267,292]" pageId="15" pageNumber="18">Type</typeStatus>
strata
</emphasis>
–
<location id="2B4460006F78FF84CBC8EA458901FEF2" LSID="urn:lsid:plazi:treatment:A63287CD6F79FF9ACB97EE958A0FFC32:2B4460006F78FF84CBC8EA458901FEF2" box="[431,567,266,292]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Sinemurian" pageId="15" pageNumber="18">Sinemurian</location>
to lowermost
<location id="2B4460006F78FF84C8BEEA45884BFEF2" LSID="urn:lsid:plazi:treatment:A63287CD6F79FF9ACB97EE958A0FFC32:2B4460006F78FF84C8BEEA45884BFEF2" box="[729,893,266,292]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Pliensbachian" pageId="15" pageNumber="18">Pliensbachian</location>
(
<collectingMunicipality id="CE40ACA16F78FF84C9EDEA458F0EFEF3" box="[906,1080,266,293]" pageId="15" pageNumber="18">Lower Jurassic</collectingMunicipality>
)
<collectingCounty id="C7454E576F78FF84CE22EA458ABAFE91" pageId="15" pageNumber="18">Hierlatz Limestone</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="2E2436DB6F78FF84CB71EA1D88E4FEBD" blockId="15.[219,1157,266,1383]" box="[278,978,337,363]" pageId="15" pageNumber="18">
<emphasis id="1CEFEAC96F78FF84CB71EA1D8AD9FEBD" box="[278,495,337,363]" italics="true" pageId="15" pageNumber="18">Derivation of name</emphasis>
– Referring to the conoidal shell outline.
</paragraph>
<paragraph id="2E2436DB6F78FF84CB71EA3A895EFD71" blockId="15.[219,1157,266,1383]" pageId="15" pageNumber="18">
<emphasis id="1CEFEAC96F78FF84CB71EA3A8AB5FE58" box="[278,387,373,398]" italics="true" pageId="15" pageNumber="18">Diagnosis</emphasis>
– Conoidal shell outline with slightly convex whorls with obscure angulation feebly below midwhorl. Ramp flattish with narrow convex belt subsuturally and concavity along selenizone. Selenizone concave and angled to ramp and outer face so blunting angulation of whorl. Outer face concave, limited by markedly swollen abapical belt, providing periphery of shell and rim of base. Base low conoidal as whole and having slightly convex wall; anomphalous or minutely phaneromphalous. Spiral threads and collabral riblets on whorls except in selenizone, spiral ones vanishing from later whorl. Base covered with almost evenly spaced quirinii
<typeStatus id="F12088796F78FF84CB87E9C18922FD7E" box="[480,532,654,680]" pageId="15" pageNumber="18">type</typeStatus>
ridges.
</paragraph>
<paragraph id="2E2436DB6F78FF84CB71E9FE8F68FD1D" blockId="15.[219,1157,266,1383]" box="[278,1118,689,715]" pageId="15" pageNumber="18">
<emphasis id="1CEFEAC96F78FF84CB71E9FE8A4DFD1D" box="[278,379,689,715]" italics="true" pageId="15" pageNumber="18">Material</emphasis>
– Single specimen, composed of remains of three latest whorls.
</paragraph>
<paragraph id="2E2436DB6F78FF84CB71E99A8947FD38" blockId="15.[219,1157,266,1383]" box="[278,625,724,750]" pageId="15" pageNumber="18">
<emphasis id="1CEFEAC96F78FF84CB71E99A8AAFFD38" box="[278,409,725,750]" italics="true" pageId="15" pageNumber="18">Dimensions</emphasis>
– Width:
<quantity id="E9639B3E6F78FF84C875E99B895BFD38" box="[530,621,724,750]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="15" pageNumber="18" unit="mm" value="9.4">9.4 mm</quantity>
.
</paragraph>
<paragraph id="2E2436DB6F78FF84CB71E9B78876FB98" blockId="15.[219,1157,266,1383]" pageId="15" pageNumber="18">
<emphasis id="1CEFEAC96F78FF84CB71E9B78AA1FCC7" box="[278,407,760,785]" italics="true" pageId="15" pageNumber="18">Description</emphasis>
– The only shell has conoidal, just slightly gradate outline with an obscure angulation feebly below the midwhorl. The rather steep ramp is flattish with a narrow convex belt subsuturally and some concavity along the selenizone. The selenizone itself is concave and angled both to the ramp and the outer face obscuring the angulation of the whorls. The outer face is concave; a markedly swollen belt limits it abapically, providing also the periphery of the shell and outer rim of the base that joins with angulation to the whorl surface. The slightly impressed suture does not follow the periphery but a line slightly adaxially from it on the base. As a whole, the base is low conoidal with slightly convex wall; the shell is anomphalous or minutely phaneromphalous.
</paragraph>
<paragraph id="2E2436DB6F78FF9ACB71EF18894DFE02" blockId="15.[219,1157,266,1383]" lastBlockId="17.[219,1157,266,996]" lastPageId="17" lastPageNumber="20" pageId="15" pageNumber="18">
Spiral threads and collabral ribs/riblets ornament the whorls apart from the selenizone; granules at the intersections are frequently preserved. The spiral threads are sparse and much weaker than the ribs on the ramp and gradually vanish for the last growth phase. The ribs of this area start from the suture and almost vanish before reaching the selenizone; their subregularly repeated interspaces gradually increase on the last half whorl. On the outer face, spiral and collabral threads form a dense network; first the collabral ones change into short riblets, crossing the spiral threads, then they fully disappear for the last whorl. The spiral threads also vanish from the adapical belt of the outer face but remain in the swollen peripheral belt. On the selenizone of the preserved shell parts, no lunulae are found. The base is covered with almost evenly spaced moderately dense quirinii
<typeStatus id="F12088796F66FF9ACBB1EA3B893CFE58" box="[470,522,372,398]" pageId="17" pageNumber="20">type</typeStatus>
ridges. The growth lines are rather strongly prosocline and prosocyrt on the ramp, prosocyrt-orthocline on the outer face and slightly prosocline-opisthocyrt on the base.
</paragraph>
<caption id="7AE466536F78FF84CABCEED48F7DF9AD" pageId="15" pageNumber="18" startId="15.[219,260,1435,1457]" targetBox="[174,1111,289,1639]" targetPageId="16" targetType="figure">
<paragraph id="2E2436DB6F78FF84CABCEED48F7DF9AD" blockId="15.[219,1157,1435,1659]" pageId="15" pageNumber="18">
<emphasis id="1CEFEAC96F78FF84CABCEED48A66FA66" bold="true" box="[219,336,1435,1457]" pageId="15" pageNumber="18">Figs 15–29.</emphasis>
New species of
<taxonomicName id="E99B4D586F78FF84CB82EED3897FFA67" authorityName="Szabó" authorityYear="2016" box="[485,585,1436,1457]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="18" phylum="Bryozoa" rank="genus" status="gen. nov.">
<emphasis id="1CEFEAC96F78FF84CB82EED3897FFA67" box="[485,585,1436,1457]" italics="true" pageId="15" pageNumber="18">Foveolaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F78FF84C829EED389BBFA67" box="[590,653,1436,1457]" pageId="15" pageNumber="18" rank="genus">n. gen.</taxonomicNameLabel>
from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. –
<emphasis id="1CEFEAC96F78FF84CE58EEF78FB3FA1B" bold="true" box="[1087,1157,1464,1485]" pageId="15" pageNumber="18">15–18.</emphasis>
Holotype of
<taxonomicName id="E99B4D586F78FF84CB3FEE9A8912FA3C" authorityName="Szabó" authorityYear="2016" box="[344,548,1493,1514]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="18" phylum="Bryozoa" rank="species" species="bicarinata" status="sp. nov.">
<emphasis id="1CEFEAC96F78FF84CB3FEE9A8912FA3C" box="[344,548,1493,1514]" italics="true" pageId="15" pageNumber="18">Foveolaria bicarinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F78FF84C84DEE998956FA3D" box="[554,608,1494,1515]" pageId="15" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
, GBA 2017/001/0003, all ×3.8. –
<emphasis id="1CEFEAC96F78FF84C9CBEE9A88FDFA3C" bold="true" box="[940,971,1493,1514]" pageId="15" pageNumber="18">15.</emphasis>
Dorsal view. –
<emphasis id="1CEFEAC96F78FF84CE02EE9A8FB3FA3C" bold="true" box="[1125,1157,1493,1514]" pageId="15" pageNumber="18">16.</emphasis>
Apical view. –
<emphasis id="1CEFEAC96F78FF84CB09EEBD8AB8F9D1" bold="true" box="[366,398,1522,1543]" pageId="15" pageNumber="18">17.</emphasis>
Basal view. –
<emphasis id="1CEFEAC96F78FF84C87BEEBD890AF9D1" bold="true" box="[540,572,1522,1543]" pageId="15" pageNumber="18">18.</emphasis>
Peristomal view. –
<emphasis id="1CEFEAC96F78FF84C898EEBD8873F9D1" bold="true" box="[767,837,1522,1543]" pageId="15" pageNumber="18">19–22.</emphasis>
Holotype of
<taxonomicName id="E99B4D586F78FF84C9ADEEBD8FB3F9D1" authorityName="Szabó" authorityYear="2016" box="[970,1157,1522,1543]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="18" phylum="Bryozoa" rank="species" species="conoidea" status="sp. nov.">
<emphasis id="1CEFEAC96F78FF84C9ADEEBD8FB3F9D1" box="[970,1157,1522,1543]" italics="true" pageId="15" pageNumber="18">Foveolaria conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F78FF84CABCED408A26F9F2" box="[219,272,1551,1572]" pageId="15" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
, GBA 2017/001/0004, all ×4. –
<emphasis id="1CEFEAC96F78FF84C82CED40895DF9F2" bold="true" box="[587,619,1551,1572]" pageId="15" pageNumber="18">19.</emphasis>
Dorsal view. –
<emphasis id="1CEFEAC96F78FF84C963ED408812F9F2" bold="true" box="[772,804,1551,1572]" pageId="15" pageNumber="18">20.</emphasis>
Apical view. –
<emphasis id="1CEFEAC96F78FF84C9DDED4088ECF9F2" bold="true" box="[954,986,1551,1572]" pageId="15" pageNumber="18">21.</emphasis>
Basal view. –
<emphasis id="1CEFEAC96F78FF84CE02ED408FB3F9F2" bold="true" box="[1125,1157,1551,1572]" pageId="15" pageNumber="18">22.</emphasis>
Peristomal view. –
<emphasis id="1CEFEAC96F78FF84CBF7ED648AE0F996" bold="true" box="[400,470,1579,1600]" pageId="15" pageNumber="18">23–26.</emphasis>
Holotype of
<taxonomicName id="E99B4D586F78FF84C834ED6389C8F997" authorityName="Szabó" authorityYear="2016" box="[595,766,1580,1601]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="18" phylum="Bryozoa" rank="species" species="globosa" status="sp. nov.">
<emphasis id="1CEFEAC96F78FF84C834ED6389C8F997" box="[595,766,1580,1601]" italics="true" pageId="15" pageNumber="18">Foveolaria globosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F78FF84C965ED638803F997" box="[770,821,1580,1601]" pageId="15" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
, GBA 2017/001/0005, all ×4. –
<emphasis id="1CEFEAC96F78FF84CE02ED648FB3F996" bold="true" box="[1125,1157,1579,1600]" pageId="15" pageNumber="18">23.</emphasis>
Dorsal view. –
<emphasis id="1CEFEAC96F78FF84CB09ED078AB8F98B" bold="true" box="[366,398,1608,1629]" pageId="15" pageNumber="18">24.</emphasis>
Apical view. –
<emphasis id="1CEFEAC96F78FF84C843ED078972F98B" bold="true" box="[548,580,1608,1629]" pageId="15" pageNumber="18">25.</emphasis>
Basal view. –
<emphasis id="1CEFEAC96F78FF84C8A8ED0789D9F98B" bold="true" box="[719,751,1608,1629]" pageId="15" pageNumber="18">26.</emphasis>
Peristomal view. –
<emphasis id="1CEFEAC96F78FF84C9C9ED0788C3F98B" bold="true" box="[942,1013,1608,1629]" pageId="15" pageNumber="18">27–29.</emphasis>
<taxonomicName id="E99B4D586F78FF84C99CED078FB7F988" box="[1019,1153,1608,1630]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="18" phylum="Bryozoa" rank="species" species="undetermined">
<emphasis id="1CEFEAC96F78FF84C99CED078F69F98B" box="[1019,1119,1608,1629]" italics="true" pageId="15" pageNumber="18">Foveolaria</emphasis>
sp.
</taxonomicName>
, GBA 2017/001/0006, all ×4. –
<emphasis id="1CEFEAC96F78FF84C858ED2A8969F9AC" bold="true" box="[575,607,1637,1658]" pageId="15" pageNumber="18">27.</emphasis>
Apertural view. –
<emphasis id="1CEFEAC96F78FF84C968ED2A8819F9AC" bold="true" box="[783,815,1637,1658]" pageId="15" pageNumber="18">28.</emphasis>
Basal view. –
<emphasis id="1CEFEAC96F78FF84C9D3ED2A88E2F9AC" bold="true" box="[948,980,1637,1658]" pageId="15" pageNumber="18">29.</emphasis>
Dorsal view
</paragraph>
</caption>
<paragraph id="2E2436DB6F66FF9ACB71EA928812FD71" blockId="17.[219,1157,266,996]" pageId="17" pageNumber="20">
<emphasis id="1CEFEAC96F66FF9ACB71EA928A4CFE21" box="[278,378,477,503]" italics="true" pageId="17" pageNumber="20">Remarks</emphasis>
– The vanishing ornament of the latest whorl resembles the last whorl of
<taxonomicName id="E99B4D586F66FF9ACB21E94E8903FDCD" authorityName="Szabó" authorityYear="2016" box="[326,565,513,539]" class="Gastropoda" family="Pithodeidae" genus="Foveolata" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="17" pageNumber="20" phylum="Mollusca" rank="species" species="bicarinata" status="sp. nov.">
<emphasis id="1CEFEAC96F66FF9ACB21E94E8903FDCD" box="[326,565,513,539]" italics="true" pageId="17" pageNumber="20">Foveolata bicarinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F66FF9AC85BE94E894CFDCD" box="[572,634,513,539]" pageId="17" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
The steeper ramp, the less impressed suture, consequently less gradate outline, and the selenizone, blunting the angulation of the whorl surface distinguish
<taxonomicName id="E99B4D586F66FF9AC80EE9088879FDB7" authorityName="Szabó" authorityYear="2016" box="[617,847,583,609]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="species" species="conoidea" status="sp. nov.">
<emphasis id="1CEFEAC96F66FF9AC80EE9088879FDB7" box="[617,847,583,609]" italics="true" pageId="17" pageNumber="20">Foveolaria conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F66FF9AC93EE90788AFFDB4" box="[857,921,584,610]" pageId="17" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
from
<taxonomicName id="E99B4D586F66FF9AC98FE9078F48FDB7" authorityName="Szabó" authorityYear="2016" box="[1000,1150,583,609]" class="Gastropoda" family="Pithodeidae" genus="Foveolata" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="17" pageNumber="20" phylum="Mollusca" rank="species" species="bicarinata">
<emphasis id="1CEFEAC96F66FF9AC98FE9078F48FDB7" box="[1000,1150,583,609]" italics="true" pageId="17" pageNumber="20">F. bicarinata</emphasis>
</taxonomicName>
. Besides, the latter species has a rather broad umbilicus but
<taxonomicName id="E99B4D586F66FF9AC9D5E9248F03FD52" authorityName="Szabó" authorityYear="2016" box="[946,1077,618,645]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="species" species="conoidea">
<emphasis id="1CEFEAC96F66FF9AC9D5E92488F4FD52" box="[946,962,619,644]" italics="true" pageId="17" pageNumber="20">F</emphasis>
.
<emphasis id="1CEFEAC96F66FF9AC9B4E9258F03FD52" box="[979,1077,618,644]" italics="true" pageId="17" pageNumber="20">conoidea</emphasis>
</taxonomicName>
seems anomphalous or very narrowly phaneromphalous.
</paragraph>
<paragraph id="2E2436DB6F66FF9ACB71E9FE8929FCC7" blockId="17.[219,1157,266,996]" pageId="17" pageNumber="20">
The subglobular shell, the more convex surfaces (ramp, outer face and base wall) and the presence of a quite wide umbilicus in
<taxonomicName id="E99B4D586F66FF9AC95CE99B8F3BFD38" authorityName="Szabó" authorityYear="2016" box="[827,1037,724,750]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="species" species="globosa" status="sp. nov.">
<emphasis id="1CEFEAC96F66FF9AC95CE99B8F3BFD38" box="[827,1037,724,750]" italics="true" pageId="17" pageNumber="20">Foveolaria globosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F66FF9ACE73E99B8F64FD38" box="[1044,1106,724,750]" pageId="17" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
distinguish it from
<emphasis id="1CEFEAC96F66FF9ACBFDE9B78929FCC7" box="[410,543,759,785]" italics="true" pageId="17" pageNumber="20">
<taxonomicName id="E99B4D586F66FF9ACBFDE9B7892CFCC7" authorityName="Szabó" authorityYear="2016" box="[410,538,759,785]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="species" species="conoidea">F. conoidea</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="2E2436DB6F66FF9ACB71E8558F20FCAD" blockId="17.[219,1157,266,996]" pageId="17" pageNumber="20">
<taxonomicName id="E99B4D586F66FF9ACB71E8558928FCE2" authorityName="Szabó" authorityYear="2016" box="[278,542,794,820]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="species" species="suemegensis" status="sp. nov.">
<emphasis id="1CEFEAC96F66FF9ACB71E8558928FCE2" box="[278,542,794,820]" italics="true" pageId="17" pageNumber="20">Foveolaria suemegensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F66FF9AC84FE854895FFCE3" box="[552,617,795,821]" pageId="17" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
has similarly steep ramp like
<taxonomicName id="E99B4D586F66FF9AC9B3E8548F62FCE2" authorityName="Szabó" authorityYear="2016" box="[980,1108,794,820]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="species" species="conoidea">
<emphasis id="1CEFEAC96F66FF9AC9B3E8548F62FCE2" box="[980,1108,794,820]" italics="true" pageId="17" pageNumber="20">F. conoidea</emphasis>
</taxonomicName>
but their ornaments at identical size are strongly different; marked in
<taxonomicName id="E99B4D586F66FF9AC9BCE8718F6EFC81" authorityName="Szabó" authorityYear="2016" box="[987,1112,829,855]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="species" species="conoidea">
<emphasis id="1CEFEAC96F66FF9AC9BCE8718F6EFC81" box="[987,1112,829,855]" italics="true" pageId="17" pageNumber="20">F. conoidea</emphasis>
</taxonomicName>
but scarce in
<emphasis id="1CEFEAC96F66FF9ACB21E82D8AD8FCAD" box="[326,494,866,891]" italics="true" pageId="17" pageNumber="20">
<taxonomicName id="E99B4D586F66FF9ACB21E82D8ADCFCAD" authorityName="Szabó" authorityYear="2016" box="[326,490,866,891]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="17" pageNumber="20" phylum="Bryozoa" rank="species" species="suemegensis">F. suemegensis</taxonomicName>
.
</emphasis>
Their selenizones are also in different position.
</paragraph>
<paragraph id="2E2436DB6F66FF9ACB71E8CB8A0FFC32" blockId="17.[219,1157,266,996]" pageId="17" pageNumber="20">
<emphasis id="1CEFEAC96F66FF9ACB71E8CB8A92FC48" box="[278,420,900,926]" italics="true" pageId="17" pageNumber="20">Distribution</emphasis>
– Within the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone Formation in Hierlatz Alpe, Hallstatt,
<collectingCountry id="568C764B6F66FF9ACABCE8858A02FC32" box="[219,308,970,996]" name="Austria" pageId="17" pageNumber="20">Austria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<subSubSection id="668165506F7AFF86C874E8D78878FC65" box="[531,846,920,947]" pageId="13" pageNumber="16" type="nomenclature">
<paragraph id="2E2436DB6F7AFF86C874E8D78878FC65" blockId="13.[531,846,920,982]" box="[531,846,920,947]" pageId="13" pageNumber="16">
<heading id="756C81B76F7AFF86C874E8D78878FC65" box="[531,846,920,947]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="16" reason="9">
<taxonomicName id="E99B4D586F7AFF86C874E8D7883DFC64" authorityName="Szabó" authorityYear="2016" box="[531,779,920,946]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="bicarinata" status="sp. nov.">
<emphasis id="1CEFEAC96F7AFF86C874E8D7883DFC64" box="[531,779,920,946]" italics="true" pageId="13" pageNumber="16">Foveolaria bicarinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7AFF86C976E8D68878FC65" box="[785,846,921,947]" pageId="13" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="668165506F7AFF85C801E8F38A3DFA78" lastPageId="14" lastPageNumber="17" pageId="13" pageNumber="16" type="description">
<paragraph id="2E2436DB6F7AFF86C801E8F389CDFC00" blockId="13.[531,846,920,982]" box="[614,763,955,982]" pageId="13" pageNumber="16">
(
<figureCitation id="B6A02A5E6F7AFF86C809E8F389C4FC00" box="[622,754,955,982]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="13" pageNumber="16">Figs 15–18</figureCitation>
)
</paragraph>
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<paragraph id="2E2436DB6F7AFF86CB71E8BE89E3FBDD" blockId="13.[219,1157,1009,1633]" box="[278,725,1009,1035]" pageId="13" pageNumber="16">
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<typeStatus id="F12088796F7AFF86CB71E8BE8A78FBDC" box="[278,334,1009,1034]" pageId="13" pageNumber="16">Type</typeStatus>
specimen
</emphasis>
– GBA 2017/001/0003.
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB71EF5B8878FBF8" blockId="13.[219,1157,1009,1633]" box="[278,846,1043,1070]" pageId="13" pageNumber="16">
<emphasis id="1CEFEAC96F7AFF86CB71EF5B8A91FBF8" box="[278,423,1044,1070]" italics="true" pageId="13" pageNumber="16">
<typeStatus id="F12088796F7AFF86CB71EF5B8A78FBFB" box="[278,334,1044,1069]" pageId="13" pageNumber="16">Type</typeStatus>
locality
</emphasis>
–
<location id="2B4460006F7AFF86CBA4EF5B8952FBF8" LSID="urn:lsid:plazi:treatment:A63287CD6F7AFF85C874E8D78A3DFA78:2B4460006F7AFF86CBA4EF5B8952FBF8" box="[451,612,1044,1070]" country="Austria" municipality="Hallstatt" name="Hierlatz Alpe" pageId="13" pageNumber="16">Hierlatz Alpe</location>
,
<collectingMunicipality id="CE40ACA16F7AFF86C809EF5B89E3FBF8" box="[622,725,1044,1070]" pageId="13" pageNumber="16">Hallstatt</collectingMunicipality>
(
<collectingCountry id="568C764B6F7AFF86C883EF5C8877FBF8" box="[740,833,1043,1070]" name="Austria" pageId="13" pageNumber="16">Austria</collectingCountry>
).
</paragraph>
</materialsCitation>
<paragraph id="2E2436DB6F7AFF86CB71EF778AF0FBA2" blockId="13.[219,1157,1009,1633]" pageId="13" pageNumber="16">
<materialsCitation id="9EF33C866F7AFF86CB71EF778AF4FBA2" county="Hierlatz Limestone" location="Within Sinemurian" municipality="Lower Jurassic" pageId="13" pageNumber="16" specimenCount="1" typeStatus="holotype">
<emphasis id="1CEFEAC96F7AFF86CB71EF778AA4FB87" box="[278,402,1080,1105]" italics="true" pageId="13" pageNumber="16">
<typeStatus id="F12088796F7AFF86CB71EF778A78FB87" box="[278,334,1080,1105]" pageId="13" pageNumber="16">Type</typeStatus>
strata
</emphasis>
–
<location id="2B4460006F7AFF86CBCCEF7889BEFB87" LSID="urn:lsid:plazi:treatment:A63287CD6F7AFF85C874E8D78A3DFA78:2B4460006F7AFF86CBCCEF7889BEFB87" box="[427,648,1079,1105]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Within Sinemurian" pageId="13" pageNumber="16">Within Sinemurian</location>
to lowermost
<location id="2B4460006F7AFF86C941EF7888F1FB87" LSID="urn:lsid:plazi:treatment:A63287CD6F7AFF85C874E8D78A3DFA78:2B4460006F7AFF86C941EF7888F1FB87" box="[806,967,1079,1105]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Pliensbachian" pageId="13" pageNumber="16">Pliensbachian</location>
(
<collectingMunicipality id="CE40ACA16F7AFF86C9B4EF788F4BFB84" box="[979,1149,1079,1106]" pageId="13" pageNumber="16">Lower Jurassic</collectingMunicipality>
)
<collectingCounty id="C7454E576F7AFF86CABCEF158AF4FBA2" box="[219,450,1114,1140]" pageId="13" pageNumber="16">Hierlatz Limestone</collectingCounty>
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.
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB71EF318F26FB6C" blockId="13.[219,1157,1009,1633]" pageId="13" pageNumber="16">
<emphasis id="1CEFEAC96F7AFF86CB71EF318AC7FB41" box="[278,497,1149,1175]" italics="true" pageId="13" pageNumber="16">Derivation of name</emphasis>
– Bicarinata (Lat.) = having two ridges; referring to the sharp spiral angulations, limiting the concave outer face of the whorls.
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB71EF8B8812FAD7" blockId="13.[219,1157,1009,1633]" pageId="13" pageNumber="16">
<emphasis id="1CEFEAC96F7AFF86CB71EF8B8A4DFB08" box="[278,379,1220,1246]" italics="true" pageId="13" pageNumber="16">Material</emphasis>
– Single fragmentary specimen with remnants of three whorls that are characteristic enough to outline a new species.
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB71EE4489B6FAF3" blockId="13.[219,1157,1009,1633]" box="[278,640,1290,1317]" pageId="13" pageNumber="16">
<emphasis id="1CEFEAC96F7AFF86CB71EE448AAFFAF2" box="[278,409,1291,1316]" italics="true" pageId="13" pageNumber="16">Dimensions</emphasis>
– Width:
<quantity id="E9639B3E6F7AFF86C875EE45894AFAF3" box="[530,636,1290,1317]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="13" pageNumber="16" unit="mm" value="12.2">12.2 mm</quantity>
.
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB71EE6188DDF9B7" blockId="13.[219,1157,1009,1633]" pageId="13" pageNumber="16">
<emphasis id="1CEFEAC96F7AFF86CB71EE618AB4FA91" box="[278,386,1326,1351]" italics="true" pageId="13" pageNumber="16">Diagnosis</emphasis>
– Gradate, medium high shell of whorls with rather steep ramp and concave outer face with sharply angular edges, abapical one providing periphery. Base low, subglobose as whole with moderately convex wall; phaneromphalous, umbilical rim rounded angular. Suture rather deeply impressed, running somewhat adaxially from periphery of former whorl. Ramp ornamented by regularly repeating collabral riblets; outer face with single thin thread above midline and some obscure striae above it. Below peripheral sharpness, another but weaker keel visible, remaining parts of base covered by quirinii
<typeStatus id="F12088796F7AFF86C955ED6B8853F9E8" box="[818,869,1572,1598]" pageId="13" pageNumber="16">type</typeStatus>
spiral ridges. Selenizone bearing lunulae on earlier whorls, gradually vanishing on last whorl.
</paragraph>
<paragraph id="2E2436DB6F79FF85CA8EEA448F70FD52" blockId="14.[174,1112,266,1454]" pageId="14" pageNumber="17">
<emphasis id="1CEFEAC96F79FF85CA8EEA448A5FFEF2" box="[233,361,267,292]" italics="true" pageId="14" pageNumber="17">Description</emphasis>
– A gradate, medium high shell is available; its reconstructed height is about
<quantity id="E9639B3E6F79FF85CB05EA618A81FE9E" box="[354,439,302,328]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="17" unit="mm" value="20.0">20 mm</quantity>
. The whorls have rather steep ramp that is slightly convex on the earlier whorls but flat on the last whorl. At its abapical edge, the spiral angulation is marked, sharp keel-like on the last whorl but less prominent, slightly rounded on the earlier shell parts. A strong spiral thread gives the adapical boundary for the selenizone that is concave. The outer face is markedly concave with an abapical edge that is also sharply angular. This ridge corresponds to the periphery. The base is low, subglobose as a whole and has moderately convex wall; it is rather broadly phaneromphalous and has a rounded angular umbilical rim. Near the peripheral angulation, a weaker keel also develops on the base; the suture is seamed about halfway between these two keels, so the suture is rather deeply impressed.
</paragraph>
<paragraph id="2E2436DB6F79FF85CA8EE9C289EFFB98" blockId="14.[174,1112,266,1454]" pageId="14" pageNumber="17">
Regularly repeating collabral riblets ornament the ramp, they start at the adapical suture, and with gradual weakening, terminate at the rim of the selenizone. On the ramp, no spiral ornament is observable on the preserved shell fragment. The outer face bears a single thin thread above the midline and some obscure striae are also visible above it. Between the weak keel of the base at the peripheral belt and the umbilicus, quirinii
<typeStatus id="F12088796F79FF85C8C3E87189EEFC8E" box="[676,728,830,856]" pageId="14" pageNumber="17">type</typeStatus>
spiral ridges represent the ornament. They are denser in the abaxial half than in the remaining part of the base. The selenizone bears lunulae on the earlier whorls that gradually vanish on the last whorl. The growth lines are strongly prosocline on the ramp with markedly prosocyrt subsutural and feebly prosocyrt remaining parts; on the outer face, the growth lines are feebly prosocline and clearly prosocyrt. The growth lines of the base are more prosocline (~45°) than it is common in
<taxonomicName id="E99B4D586F79FF85C97DEF5E88A2FBFD" authorityName="Szabó" authorityYear="2016" box="[794,916,1041,1067]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="17" phylum="Bryozoa" rank="genus">
<emphasis id="1CEFEAC96F79FF85C97DEF5E88A2FBFD" box="[794,916,1041,1067]" italics="true" pageId="14" pageNumber="17">Foveolaria</emphasis>
</taxonomicName>
(~30°), and they are somewhat more opisthocyrt than it is usual.
</paragraph>
<paragraph id="2E2436DB6F79FF85CA8EEF1889A9FA92" blockId="14.[174,1112,266,1454]" pageId="14" pageNumber="17">
<emphasis id="1CEFEAC96F79FF85CA8EEF18895FFBA7" box="[233,617,1111,1138]" italics="true" pageId="14" pageNumber="17">
Remarks –
<taxonomicName id="E99B4D586F79FF85CB08EF18895FFBA7" authorityName="Szabó" authorityYear="2016" box="[367,617,1111,1137]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="17" phylum="Bryozoa" rank="species" species="bicarinata" status="sp. nov.">Foveolaria bicarinata</taxonomicName>
</emphasis>
<taxonomicNameLabel id="07DC57B26F79FF85C815EF178984FBA4" box="[626,690,1112,1138]" pageId="14" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
resembles
<taxonomicName id="E99B4D586F79FF85C951EF1788EEFBA7" authorityName="Szabó" authorityYear="2016" box="[822,984,1112,1137]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="17" phylum="Bryozoa" rank="species" species="suemegensis" status="n. sp.">
<emphasis id="1CEFEAC96F79FF85C951EF1788EEFBA7" box="[822,984,1112,1137]" italics="true" pageId="14" pageNumber="17">F. suemegensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F79FF85C986EF178F17FBA4" box="[993,1057,1112,1138]" pageId="14" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
(see below) in the keel like angulations on the whorl surface and at the periphery, limiting a markedly concave outer face. The latter species has higher, somewhat acute spire, much widely developed and steeper ramp. Moreover,
<taxonomicName id="E99B4D586F79FF85C9D3EF8D8F61FB0D" authorityName="Szabó" authorityYear="2016" box="[948,1111,1217,1243]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="17" phylum="Bryozoa" rank="species" species="suemegensis">
<emphasis id="1CEFEAC96F79FF85C9D3EF8D88F2FB0D" box="[948,964,1218,1243]" italics="true" pageId="14" pageNumber="17">F</emphasis>
.
<emphasis id="1CEFEAC96F79FF85C9B5EF8D8F61FB0D" box="[978,1111,1218,1243]" italics="true" pageId="14" pageNumber="17">suemegensis</emphasis>
</taxonomicName>
has only barely developed or obscure spiral ornament on the latest whorls, while
<taxonomicName id="E99B4D586F79FF85CAC9EE478A74FAF7" authorityName="Szabó" authorityYear="2016" box="[174,322,1287,1313]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="17" phylum="Bryozoa" rank="species" species="bicarinata">
<emphasis id="1CEFEAC96F79FF85CAC9EE478A74FAF7" box="[174,322,1287,1313]" italics="true" pageId="14" pageNumber="17">F. bicarinata</emphasis>
</taxonomicName>
bears few but clear spiral threads and sharp collabral riblets. Their dimensions are also significantly different.
</paragraph>
<paragraph id="2E2436DB6F79FF85CA8EEE028A3DFA78" blockId="14.[174,1112,266,1454]" pageId="14" pageNumber="17">
<emphasis id="1CEFEAC96F79FF85CA8EEE028A41FAB1" box="[233,375,1357,1383]" italics="true" pageId="14" pageNumber="17">Distribution</emphasis>
– Within the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone Formation in Hierlatz Alpe, Hallstatt,
<collectingCountry id="568C764B6F79FF85CAC9EEDB8A31FA78" box="[174,263,1428,1454]" name="Austria" pageId="14" pageNumber="17">Austria</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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444
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Genus
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<emphasis id="1CEFEAC96F7DFF81C822EE488989FAF7" box="[581,703,1287,1313]" italics="true" pageId="10" pageNumber="13">Foveolaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7DFF81C8A2EE478824FAF4" box="[709,786,1288,1314]" pageId="10" pageNumber="13" rank="genus">n. gen.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="668165506F7DFF81CA8EEE72889CFA80" box="[233,938,1340,1367]" pageId="10" pageNumber="13" type="type_taxon">
<paragraph id="2E2436DB6F7DFF81CA8EEE72889CFA80" blockId="10.[174,1112,1340,1648]" box="[233,938,1340,1367]" pageId="10" pageNumber="13">
<emphasis id="1CEFEAC96F7DFF81CA8EEE728AB0FA81" box="[233,390,1341,1367]" italics="true" pageId="10" pageNumber="13">
<typeStatus id="F12088796F7DFF81CA8EEE728A17FA80" box="[233,289,1341,1366]" pageId="10" pageNumber="13">Type</typeStatus>
species –
</emphasis>
<taxonomicName id="E99B4D586F7DFF81CBEAEE738890FA80" authority="Szabo, 2009" authorityName="Szabo" authorityYear="2009" box="[397,934,1340,1367]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="lokutensis" subGenus="Sisenna">
<emphasis id="1CEFEAC96F7DFF81CBEAEE73891DFA80" box="[397,555,1340,1366]" italics="true" pageId="10" pageNumber="13">Wortheniopsis</emphasis>
(
<emphasis id="1CEFEAC96F7DFF81C85CEE7389A4FA80" box="[571,658,1340,1366]" italics="true" pageId="10" pageNumber="13">Sisenna</emphasis>
)
<emphasis id="1CEFEAC96F7DFF81C8C4EE738824FA80" box="[675,786,1340,1366]" italics="true" pageId="10" pageNumber="13">lokutensis</emphasis>
<bibRefCitation id="4A0A4B2A6F7DFF81C97FEE738890FA80" author="SZABO J." box="[792,934,1340,1366]" pageId="10" pageNumber="13" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="668165506F7DFF81CA8EEE2F898BFA16" pageId="10" pageNumber="13" type="etymology">
<paragraph id="2E2436DB6F7DFF81CA8EEE2F898BFA16" blockId="10.[174,1112,1340,1648]" pageId="10" pageNumber="13">
<emphasis id="1CEFEAC96F7DFF81CA8EEE2F8AF5FAAF" box="[233,451,1375,1401]" italics="true" pageId="10" pageNumber="13">Derivation of name</emphasis>
– Foveola (Lat.) = small hole, alluding to
<taxonomicName id="E99B4D586F7DFF81C9D5EE108996FA4B" authority="J. A. Eudes-Deslongchamp, 1849" authorityName=", J. A. EUDES-DESLONGCHAMPS" authorityYear="1849" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="foveolata">
<emphasis id="1CEFEAC96F7DFF81C9D5EE108A21FA4A" italics="true" pageId="10" pageNumber="13">Pleurotomaria foveolata</emphasis>
J. A. Eudes-Deslongchamp, 1849
</taxonomicName>
, the earliest species name for a group of species, included here to
<taxonomicName id="E99B4D586F7DFF81CB97EEE9895CFA16" authorityName="Szabó" authorityYear="2016" box="[496,618,1446,1472]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="10" pageNumber="13" phylum="Bryozoa" rank="genus" status="gen. nov.">
<emphasis id="1CEFEAC96F7DFF81CB97EEE9895CFA16" box="[496,618,1446,1472]" italics="true" pageId="10" pageNumber="13">Foveolaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7DFF81C817EEE9898BFA16" box="[624,701,1446,1472]" pageId="10" pageNumber="13" rank="genus">n. gen.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="668165506F7DFF80CA8EEE8589AEFE21" lastPageId="11" lastPageNumber="14" pageId="10" pageNumber="13" type="diagnosis">
<paragraph id="2E2436DB6F7DFF80CA8EEE8589AEFE21" blockId="10.[174,1112,1340,1648]" lastBlockId="11.[219,1157,266,1665]" lastPageId="11" lastPageNumber="14" pageId="10" pageNumber="13">
<emphasis id="1CEFEAC96F7DFF81CA8EEE858A60FA35" box="[233,342,1482,1507]" italics="true" pageId="10" pageNumber="13">Diagnosis</emphasis>
– Turbiniform to conoidal or coeloconoidal, thin-walled shells of slightly convex whorls, being obscurely to clearly angular in latest growth stages. Base convex, narrowly phaneromphalous, some possibly anomphalous. Slit rather long, selenizone concave or flat between two threads at or below midwhorl as abapical belt of ramp or blunting angulation of whorl surface. Outer whorl face more or less concave. Abaxial rim of base angular, reinforced by cord or swollen belt, providing periphery. Apex blunt, protoconch low with evenly convex whorls. Ornament consisting of collabral riblets and spiral threads in high variability of patterns; granules frequently occurring at crossing points. Lunulae mostly on early selenizone. Growth lines delicate, more prosocline and prosocyrt adapically from selenizone, feebly prosocline to opisthocline abapically; prosocline and slightly opisthocyrt on base.
</paragraph>
</subSubSection>
<subSubSection id="668165506F7CFF80CB71E94D8A72F957" pageId="11" pageNumber="14" type="description">
<paragraph id="2E2436DB6F7CFF80CB71E94D8903FB28" blockId="11.[219,1157,266,1665]" pageId="11" pageNumber="14">
<emphasis id="1CEFEAC96F7CFF80CB71E94D8AA1FDCD" box="[278,407,514,539]" italics="true" pageId="11" pageNumber="14">Description</emphasis>
– Shells of
<taxonomicName id="E99B4D586F7CFF80C841E94E8996FDCD" authorityName="Szabó" authorityYear="2016" box="[550,672,513,539]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="11" pageNumber="14" phylum="Bryozoa" rank="genus">
<emphasis id="1CEFEAC96F7CFF80C841E94E8996FDCD" box="[550,672,513,539]" italics="true" pageId="11" pageNumber="14">Foveolaria</emphasis>
</taxonomicName>
approach turbiniform to conoidal shape with more or less gradate outline and variable spire height. They are very thin-walled, the early teleoconch whorls are simply convex but the latest whorls become obscurely to markedly angular at or below the midwhorl. The protoconch is poorly known; it has depressed turbiniform shape in the
<typeStatus id="F12088796F7CFF80C90AE9C18897FD7E" box="[877,929,654,680]" pageId="11" pageNumber="14">type</typeStatus>
species with evenly convex whorls and rather deeply impressed suture. The suture remains more or less impressed also along the subsequent whorls. The last whorl meets the base in a more or less rounded angulation that is reinforced by a swollen spiral belt or cord. The base as a whole has subconoidal to subglobular shape with feebly to strongly convex wall. In most species, a narrow phaneromphalus is observed; in some cases absence of the umbilicus seems probable. The slit is rather long as it is demonstrated by the only photo available (
<bibRefCitation id="4A0A4B2A6F7CFF80C966E8CB8F07FC48" author="FISCHER J. - C. &amp; WEBER C." box="[769,1073,900,926]" pageId="11" pageNumber="14" pagination="451" refId="ref11914" refString="FISCHER J. - C. &amp; WEBER C. 1997: Volume II, Gasteropodes Jurassiques. - In: FISCHER J. C. (ed.): Revision critique de la Paleontologie Francaise, incluant la reedition de la original. - pp. 300 + 622, Pls 451; Paris, Masson &amp; Museum national d'Histoire naturelle." type="book chapter" year="1997">FISCHER &amp; WEBER 1997</bibRefCitation>
, Pl. 25,
<figureCitation id="B6A02A5E6F7CFF80CABCE8E88A18FC14" box="[219,302,935,962]" captionStart="Figs 4–14" captionStartId="8.[174,214,1437,1459]" captionTargetBox="[174,1111,265,1412]" captionTargetId="figure-18@8.[174,1111,265,1412]" captionTargetPageId="8" captionText="Figs 4–14.Wortheniopsisspecies fromtheHierlatz LimestoneFormation.– 4–10.Wortheniopsisrakusi n.sp.from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe,Hallstatt, Austria,GBA 2017/001/0002,all ×4.– 4. Dorsal view.– 5. Peristomal view.– 6. Basal view. – 7–9. Same views as 4–6 but in natural (×1) size. – 10. Details of the ornament on thespire whorls,×7.– 11–14.Wortheniopsis bakonyensis n. sp.,?Lower Sinemurian (Lower Jurassic) Hierlatz Limestone variety with rich micrite content, Csárdahegy, Úrkút, Bakony Mts, Hungary, GGIH J.08.3.1. – 11. Peristomal view of the holotype, ×4. – 12. Lateral view to show the best preserved outline part, ×4.– 13–14. The same views as in figures 11–12 but in natural (×1) size" figureDoi="http://doi.org/10.5281/zenodo.15627167" httpUri="https://zenodo.org/record/15627167/files/figure.png" pageId="11" pageNumber="14">Fig. 12</figureCitation>
), it extends about 60° abaperturally. The selenizone is mostly concave or rarely flat between two threads at or, more commonly, somewhat below the midwhorl. Usually, it lies as abapical belt within the plane of the ramp and its abapical thread coincides with the ridge of angulation of the whorls but less frequently, it forms a third surface of the whorls, which blunts the angulation between the ramp and the outer face. Width of the selenizone is around 15% of the distance between two neighbouring sutures. In some species, the limiting threads are lacking or substituted by shallow striae on the last whorl. Outer face of the whorls is flat to concave and nearly parallel to the coiling axis; usually, its abapical rim corresponds to the periphery.
</paragraph>
<paragraph id="2E2436DB6F7CFF80CB71EE488A72F957" blockId="11.[219,1157,266,1665]" pageId="11" pageNumber="14">
The ornament is composed of collabral riblets and spiral threads in high variability of patterns, frequently combined with granules and/or short riblets between pairs of spiral ridges at the crossing points. The short riblets are usually aligned collabrally forming false threads. Quirinii
<typeStatus id="F12088796F7CFF80C940EE3E886DFA5D" box="[807,859,1393,1419]" pageId="11" pageNumber="14">type</typeStatus>
ridges are also developed usually only on the base and mainly in the stratigraphically older (Sinemurian) species; rarely they occur also on the abapical part of the outer face. Lunulae occur mostly on the early selenizone parts, rarely present also on latest whorls. The growth lines are delicate, clearly prosocline and prosocyrt adapically from the selenizone but slightly prosocline to opisthocline and prosocyrt between the selenizone and the periphery, slightly prosocline and feebly opisthocyrt on the base.
</paragraph>
</subSubSection>
<subSubSection id="668165506F7BFF86CA8EEA458852FE34" lastPageId="13" lastPageNumber="16" pageId="12" pageNumber="15" type="discussion">
<paragraph id="2E2436DB6F7BFF87CA8EEA4588F1FCC7" blockId="12.[174,1114,266,1666]" pageId="12" pageNumber="15">
<emphasis id="1CEFEAC96F7BFF87CA8EEA458A7BFEF2" box="[233,333,266,292]" italics="true" pageId="12" pageNumber="15">Remarks</emphasis>
– The shell form and many details are similar in
<taxonomicName id="E99B4D586F7BFF87C9E1EA458F12FEF2" authorityName="J. Bohm" authorityYear="1895" box="[902,1060,266,292]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F7BFF87C9E1EA458F12FEF2" box="[902,1060,266,292]" italics="true" pageId="12" pageNumber="15">Wortheniopsis</emphasis>
</taxonomicName>
and
<taxonomicName id="E99B4D586F7BFF87CAC9EA628A1EFE91" authorityName="Szabó" authorityYear="2016" box="[174,296,301,327]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="genus">
<emphasis id="1CEFEAC96F7BFF87CAC9EA628A1EFE91" box="[174,296,301,327]" italics="true" pageId="12" pageNumber="15">Foveolaria</emphasis>
</taxonomicName>
. Most reliable distinctive characters are the position of the selenizone, the orientation and shape of the growth lines (outer lip), the width of the outer face and the distribution of the quirinii
<typeStatus id="F12088796F7BFF87C812EA3B899FFE58" box="[629,681,372,398]" pageId="12" pageNumber="15">type</typeStatus>
ridges on the shells. The selenizone is above the midwhorl in
<taxonomicName id="E99B4D586F7BFF87CBDEEAD88961FE67" authorityName="J. Bohm" authorityYear="1895" box="[441,599,407,433]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F7BFF87CBDEEAD88961FE67" box="[441,599,407,433]" italics="true" pageId="12" pageNumber="15">Wortheniopsis</emphasis>
</taxonomicName>
but at/or below the midwhorl in
<emphasis id="1CEFEAC96F7BFF87C9B1EAD88F6EFE67" box="[982,1112,407,433]" italics="true" pageId="12" pageNumber="15">
<taxonomicName id="E99B4D586F7BFF87C9B1EAD88F62FE67" authorityName="Szabó" authorityYear="2016" box="[982,1108,407,433]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="genus">Foveolaria</taxonomicName>
.
</emphasis>
Consequently, the width of the outer face is significantly narrower in
<taxonomicName id="E99B4D586F7BFF87C9BAEAF58F61FE02" authorityName="Szabó" authorityYear="2016" box="[989,1111,442,468]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="genus">
<emphasis id="1CEFEAC96F7BFF87C9BAEAF58F61FE02" box="[989,1111,442,468]" italics="true" pageId="12" pageNumber="15">Foveolaria</emphasis>
</taxonomicName>
than in
<emphasis id="1CEFEAC96F7BFF87CB77EA928A80FE21" box="[272,438,477,503]" italics="true" pageId="12" pageNumber="15">
<taxonomicName id="E99B4D586F7BFF87CB77EA928A84FE21" authorityName="J. Bohm" authorityYear="1895" box="[272,434,477,503]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="genus">Wortheniopsis</taxonomicName>
.
</emphasis>
The growth lines (outer lip) are almost orthocline, just feebly prosocline in
<taxonomicName id="E99B4D586F7BFF87CBCDE94E897EFDCD" authorityName="J. Bohm" authorityYear="1895" box="[426,584,513,539]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F7BFF87CBCDE94E897EFDCD" box="[426,584,513,539]" italics="true" pageId="12" pageNumber="15">Wortheniopsis</emphasis>
</taxonomicName>
but clearly prosocline in
<taxonomicName id="E99B4D586F7BFF87C9EEE94E8F35FDCD" authorityName="Szabó" authorityYear="2016" box="[905,1027,513,539]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="genus">
<emphasis id="1CEFEAC96F7BFF87C9EEE94E8F35FDCD" box="[905,1027,513,539]" italics="true" pageId="12" pageNumber="15">Foveolaria</emphasis>
</taxonomicName>
in the belt adapically from the selenizone. The quirinii
<typeStatus id="F12088796F7BFF87C88FE96B882AFDE8" box="[744,796,548,574]" pageId="12" pageNumber="15">type</typeStatus>
ridges appear between the abapical edge of the selenizone and the axial region of the base in
<taxonomicName id="E99B4D586F7BFF87C9D5E9088F66FDB7" authorityName="J. Bohm" authorityYear="1895" box="[946,1104,583,609]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F7BFF87C9D5E9088F66FDB7" box="[946,1104,583,609]" italics="true" pageId="12" pageNumber="15">Wortheniopsis</emphasis>
</taxonomicName>
, however, they are present mostly on the base or, mainly in stratigraphically younger species, substituted by simple spiral threads in
<taxonomicName id="E99B4D586F7BFF87C95BE9C28880FD71" authorityName="Szabó" authorityYear="2016" box="[828,950,653,679]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="genus">
<emphasis id="1CEFEAC96F7BFF87C95BE9C28880FD71" box="[828,950,653,679]" italics="true" pageId="12" pageNumber="15">Foveolaria</emphasis>
</taxonomicName>
. In few cases, they are recognisable also on the outer face in strongly modified form, e.g. in
<taxonomicName id="E99B4D586F7BFF87CE24E9FD8ADEFD39" authority="(Szabo, 2009)" baseAuthorityName="Szabo" baseAuthorityYear="2009" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="species" species="hierlatzensis">
<emphasis id="1CEFEAC96F7BFF87CE24E9FD8A0BFD38" italics="true" pageId="12" pageNumber="15">F. hierlatzensis</emphasis>
(
<bibRefCitation id="4A0A4B2A6F7BFF87CB29E99B8AE9FD39" author="SZABO J." box="[334,479,724,751]" pageId="12" pageNumber="15" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
)
</taxonomicName>
or in
<taxonomicName id="E99B4D586F7BFF87C854E99A89AEFD38" authorityName="Szabó" authorityYear="2016" box="[563,664,724,750]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="species" species="kocsisi" status="n. sp.">
<emphasis id="1CEFEAC96F7BFF87C854E99A89AEFD38" box="[563,664,724,750]" italics="true" pageId="12" pageNumber="15">F. kocsisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7BFF87C8C7E99B89D7FD38" box="[672,737,724,750]" pageId="12" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
, with short riblet-like granules, rapidly diminishing abapically before reaching the next spiral ridge.
</paragraph>
<paragraph id="2E2436DB6F7BFF87CA8EE8548873FBFD" blockId="12.[174,1114,266,1666]" pageId="12" pageNumber="15">
In common cases, the selenizone is the abapical belt of the ramp in
<taxonomicName id="E99B4D586F7BFF87C9B0E8558F67FCE2" authorityName="Szabó" authorityYear="2016" box="[983,1105,794,820]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="genus">
<emphasis id="1CEFEAC96F7BFF87C9B0E8558F67FCE2" box="[983,1105,794,820]" italics="true" pageId="12" pageNumber="15">Foveolaria</emphasis>
</taxonomicName>
, however, in some species, the selenizone does not lie in the plane of the ramp but blunts the angulation of the whorl surface like e.g. in
<taxonomicName id="E99B4D586F7BFF87C94AE82E8F27FCAD" authorityName="Szabó" authorityYear="2016" box="[813,1041,865,891]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="species" species="conoidea" status="sp. nov.">
<emphasis id="1CEFEAC96F7BFF87C94AE82E8F27FCAD" box="[813,1041,865,891]" italics="true" pageId="12" pageNumber="15">Foveolaria conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7BFF87CE7EE82E8F61FCAD" box="[1049,1111,865,891]" pageId="12" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
or
<taxonomicName id="E99B4D586F7BFF87CAB7E8CB8A95FC48" authorityName="Szabó" authorityYear="2016" box="[208,419,900,926]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="species" species="globosa" status="sp. nov.">
<emphasis id="1CEFEAC96F7BFF87CAB7E8CB8A95FC48" box="[208,419,900,926]" italics="true" pageId="12" pageNumber="15">Foveolaria globosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7BFF87CBCCE8CB8ADCFC48" box="[427,490,900,926]" pageId="12" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
(see below). If the selenizone is not clearly concave, like in
<taxonomicName id="E99B4D586F7BFF87CB67E8E8899CFC14" authority="(Szabo, 2009)" baseAuthorityName="Szabo" baseAuthorityYear="2009" box="[256,682,935,962]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="species" species="hierlatzensis">
<emphasis id="1CEFEAC96F7BFF87CB67E8E88931FC17" box="[256,519,935,961]" italics="true" pageId="12" pageNumber="15">Foveolaria hierlatzensis</emphasis>
(
<bibRefCitation id="4A0A4B2A6F7BFF87C872E8E88997FC14" author="SZABO J." box="[533,673,935,962]" pageId="12" pageNumber="15" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
)
</taxonomicName>
, these forms may recall the shape of
<taxonomicName id="E99B4D586F7BFF87CAC9E8858AECFC33" authority="Cox, 1956" authorityName="Cox" authorityYear="1956" box="[174,474,970,997]" class="Gastropoda" family="Pleurotomariidae" genus="Bathrotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="genus">
<emphasis id="1CEFEAC96F7BFF87CAC9E8858A6EFC32" box="[174,344,970,996]" italics="true" pageId="12" pageNumber="15">Bathrotomaria</emphasis>
Cox, 1956
</taxonomicName>
. However, in the latter genus, the selenizone is clearly convex, its adapical belt is part of the ramp and the abapical part belongs to the outer face, i.e. the selenizone itself is also mostly angular.
</paragraph>
<paragraph id="2E2436DB6F7BFF87CA8EEF7B8AF2F957" blockId="12.[174,1114,266,1666]" pageId="12" pageNumber="15">
<emphasis id="1CEFEAC96F7BFF87CA8EEF7B8A00FB98" box="[233,310,1076,1102]" italics="true" pageId="12" pageNumber="15">Species</emphasis>
– With the name
<taxonomicName id="E99B4D586F7BFF87C87CEF7B8A5EFBA4" authority=", J. A. EUDES-DESLONGCHAMPS (1849)" authorityName=", J. A. EUDES-DESLONGCHAMPS" authorityYear="1849" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="foveolata">
<emphasis id="1CEFEAC96F7BFF87C87CEF7B8819FB98" box="[539,815,1076,1102]" italics="true" pageId="12" pageNumber="15">Pleurotomaria foveolata</emphasis>
, J. A.
<bibRefCitation id="4A0A4B2A6F7BFF87C918EF7B8A5EFBA4" author="EUDES-DESLONGCHAMPS J. A." pageId="12" pageNumber="15" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">EUDES-DESLONGCHAMPS (1849)</bibRefCitation>
</taxonomicName>
covered six different but doubtlessly related shell forms that he regarded as varieties (
<taxonomicName id="E99B4D586F7BFF87CBD6EF358910FB42" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[433,550,1146,1172]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="species" species="trochoidea">
<emphasis id="1CEFEAC96F7BFF87CBD6EF358910FB42" box="[433,550,1146,1172]" italics="true" pageId="12" pageNumber="15">trochoidea</emphasis>
</taxonomicName>
,
<taxonomicName id="E99B4D586F7BFF87C852EF35899FFB42" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[565,681,1146,1172]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="species" species="subturrita">
<emphasis id="1CEFEAC96F7BFF87C852EF35899FFB42" box="[565,681,1146,1172]" italics="true" pageId="12" pageNumber="15">subturrita</emphasis>
</taxonomicName>
,
<taxonomicName id="E99B4D586F7BFF87C8DFEF348830FB42" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[696,774,1147,1172]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="species" species="turrita">
<emphasis id="1CEFEAC96F7BFF87C8DFEF348830FB42" box="[696,774,1147,1172]" italics="true" pageId="12" pageNumber="15">turrita</emphasis>
</taxonomicName>
,
<taxonomicName id="E99B4D586F7BFF87C972EF348850FB42" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[789,870,1147,1172]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="procera">
<emphasis id="1CEFEAC96F7BFF87C972EF348850FB42" box="[789,870,1147,1172]" italics="true" pageId="12" pageNumber="15">procera</emphasis>
</taxonomicName>
,
<taxonomicName id="E99B4D586F7BFF87C912EF3488F3FB42" baseAuthorityName="Deslongchamps" baseAuthorityYear="1849" box="[885,965,1147,1172]" class="Gastropoda" family="Pleurotomariidae" genus="Sisenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="pinguis">
<emphasis id="1CEFEAC96F7BFF87C912EF3488F3FB42" box="[885,965,1147,1172]" italics="true" pageId="12" pageNumber="15">pinguis</emphasis>
</taxonomicName>
and
<taxonomicName id="E99B4D586F7BFF87CE65EF358BE0FB61" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="12" pageNumber="15" phylum="Bryozoa" rank="species" species="ellipsoidea">
<emphasis id="1CEFEAC96F7BFF87CE65EF358BE0FB61" italics="true" pageId="12" pageNumber="15">ellipsoidea</emphasis>
</taxonomicName>
). D’ORBIGNY (1854) raised these varieties, with some modifications, to species rank with indication of own authorship:
<taxonomicName id="E99B4D586F7BFF87C8DBEF8E88EFFB0D" baseAuthorityName="d'Orbigny" baseAuthorityYear="1853" box="[700,985,1217,1243]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="subturrita">
<emphasis id="1CEFEAC96F7BFF87C8DBEF8E88EFFB0D" box="[700,985,1217,1243]" italics="true" pageId="12" pageNumber="15">Pleurotomaria subturrita</emphasis>
</taxonomicName>
,
<taxonomicName id="E99B4D586F7BFF87C981EF8D8F67FB0D" box="[998,1105,1218,1243]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="pinguis">
<emphasis id="1CEFEAC96F7BFF87C981EF8D8F67FB0D" box="[998,1105,1218,1243]" italics="true" pageId="12" pageNumber="15">P. pinguis</emphasis>
</taxonomicName>
,
<taxonomicName id="E99B4D586F7BFF87CAC9EFAA8A0AFB28" box="[174,316,1252,1278]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="ellipsoidea">
<emphasis id="1CEFEAC96F7BFF87CAC9EFAA8A0AFB28" box="[174,316,1252,1278]" italics="true" pageId="12" pageNumber="15">P. ellipsoidea</emphasis>
</taxonomicName>
,
<taxonomicName id="E99B4D586F7BFF87CB2EEFAA8AC6FB28" box="[329,496,1252,1278]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="subfaveolata">
<emphasis id="1CEFEAC96F7BFF87CB2EEFAA8AC6FB28" box="[329,496,1252,1278]" italics="true" pageId="12" pageNumber="15">P. subfaveolata</emphasis>
</taxonomicName>
,
<taxonomicName id="E99B4D586F7BFF87CB9AEFAA895FFB28" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[509,617,1253,1278]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="procera">
<emphasis id="1CEFEAC96F7BFF87CB9AEFAA895FFB28" box="[509,617,1253,1278]" italics="true" pageId="12" pageNumber="15">P. procera</emphasis>
</taxonomicName>
. The name
<taxonomicName id="E99B4D586F7BFF87C897EFAB8852FB28" baseAuthorityName="d'Orbigny" baseAuthorityYear="1853" box="[752,868,1252,1278]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="subturrita">
<emphasis id="1CEFEAC96F7BFF87C897EFAB8852FB28" box="[752,868,1252,1278]" italics="true" pageId="12" pageNumber="15">subturrita</emphasis>
</taxonomicName>
in d’Orbigny’s interpretation contains also
<taxonomicName id="E99B4D586F7BFF87CBDDEE4788B7FAF7" authority="J. A. Eudes-Deslongchamps" authorityName="J. A. Eudes-Deslongchamps" box="[442,897,1287,1314]" pageId="12" pageNumber="15" rank="variety" variety="turrita">
var.
<emphasis id="1CEFEAC96F7BFF87CB8DEE47890EFAF7" box="[490,568,1288,1313]" italics="true" pageId="12" pageNumber="15">turrita</emphasis>
J. A. Eudes-Deslongchamps
</taxonomicName>
; the name
<taxonomicName id="E99B4D586F7BFF87C99CEE488BD1FA92" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="subfaveolata">
<emphasis id="1CEFEAC96F7BFF87C99CEE488BD1FA92" italics="true" pageId="12" pageNumber="15">subfaveolata</emphasis>
</taxonomicName>
is given instead of
<taxonomicName id="E99B4D586F7BFF87CBA2EE6488EBFA93" authority="J. A. Eudes-Deslongchamps. By" authorityName="J. A. Eudes-Deslongchamps. By" box="[453,989,1322,1349]" pageId="12" pageNumber="15" rank="variety" variety="ellipsoidea">
var.
<emphasis id="1CEFEAC96F7BFF87CB91EE65895CFA92" box="[502,618,1322,1348]" italics="true" pageId="12" pageNumber="15">ellipsoidea</emphasis>
J. A. Eudes-Deslongchamps. By
</taxonomicName>
the
<collectionCode id="488AAE1E6F7BFF87CE77EE658F61FA92" box="[1040,1111,1322,1348]" pageId="12" pageNumber="15">ICZN</collectionCode>
(1999) rules, the true author of these species names is J. A. Eudes-Deslongchamps, except
<taxonomicName id="E99B4D586F7BFF87CB67EE3E8ABAFA5D" box="[256,396,1393,1419]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="subfaveolata">
<emphasis id="1CEFEAC96F7BFF87CB67EE3E8ABAFA5D" box="[256,396,1393,1419]" italics="true" pageId="12" pageNumber="15">subfaveolata</emphasis>
</taxonomicName>
, and the original species name (
<taxonomicName id="E99B4D586F7BFF87C96DEE3E8844FA5D" authorityName=", J. A. EUDES-DESLONGCHAMPS" authorityYear="1849" box="[778,882,1393,1419]" class="Gastropoda" family="Pleurotomariidae" genus="Pleurotomaria" kingdom="Animalia" order="Pleurotomariida" pageId="12" pageNumber="15" phylum="Mollusca" rank="species" species="foveolata">
<emphasis id="1CEFEAC96F7BFF87C96DEE3E8844FA5D" box="[778,882,1393,1419]" italics="true" pageId="12" pageNumber="15">foveolata</emphasis>
</taxonomicName>
) becomes a nomen nudum. This name remains applicable only if a reviser (1) rejects any subdivision, or (2) reduces the rank of the varieties as
<bibRefCitation id="4A0A4B2A6F7BFF87C8FDEEF888EEFA04" author="FISCHER J. - C. &amp; WEBER C." box="[666,984,1463,1490]" pageId="12" pageNumber="15" pagination="451" refId="ref11914" refString="FISCHER J. - C. &amp; WEBER C. 1997: Volume II, Gasteropodes Jurassiques. - In: FISCHER J. C. (ed.): Revision critique de la Paleontologie Francaise, incluant la reedition de la original. - pp. 300 + 622, Pls 451; Paris, Masson &amp; Museum national d'Histoire naturelle." type="book chapter" year="1997">FISCHER &amp; WEBER (1997)</bibRefCitation>
have done it; in their species interpretation the former varieties are treated as “morphes”. In either cases
<typeStatus id="F12088796F7BFF87CB00EEB28AE4F9C1" box="[359,466,1533,1559]" pageId="12" pageNumber="15" type="lectotype">lectotype</typeStatus>
or
<typeStatus id="F12088796F7BFF87C866EEB18969F9CE" box="[513,607,1534,1560]" pageId="12" pageNumber="15" type="neotype">neotype</typeStatus>
selection is necessary like in
<bibRefCitation id="4A0A4B2A6F7BFF87C9ADEEB18A56F9ED" author="FISCHER J. - C. &amp; WEBER C." pageId="12" pageNumber="15" pagination="451" refId="ref11914" refString="FISCHER J. - C. &amp; WEBER C. 1997: Volume II, Gasteropodes Jurassiques. - In: FISCHER J. C. (ed.): Revision critique de la Paleontologie Francaise, incluant la reedition de la original. - pp. 300 + 622, Pls 451; Paris, Masson &amp; Museum national d'Histoire naturelle." type="book chapter" year="1997">FISCHER &amp; WEBER (1997)</bibRefCitation>
. Following the arguments in
<bibRefCitation id="4A0A4B2A6F7BFF87C8D8ED6F886BF9ED" author="SZABO J." box="[703,861,1568,1595]" pageId="12" pageNumber="15" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">SZABÓ (2009</bibRefCitation>
, p. 28), J. A.
<bibRefCitation id="4A0A4B2A6F7BFF87C999ED6E8AE9F989" author="EUDES-DESLONGCHAMPS J. A." pageId="12" pageNumber="15" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">EUDES-DESLONGCHAMPS (1849)</bibRefCitation>
varieties are treated as species also in this paper but without a deep revision.
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB5DEA458852FE34" blockId="13.[314,1157,266,770]" pageId="13" pageNumber="16">
<heading id="756C81B76F7AFF86CB5DEA458852FE34" fontSize="11" level="2" pageId="13" pageNumber="16" reason="4">
<taxonomicName id="E99B4D586F7AFF86CB5DEA4588BAFEF4" authority="(J. A. Eudes-Deslongchamps, 1849)" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[314,908,266,291]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="trochoidea">
<emphasis id="1CEFEAC96F7AFF86CB5DEA458922FEF4" box="[314,532,266,290]" italics="true" pageId="13" pageNumber="16">Foveolaria trochoidea</emphasis>
(J. A.
<bibRefCitation id="4A0A4B2A6F7AFF86C832EA4588B5FEF4" author="EUDES-DESLONGCHAMPS J. A." box="[597,899,266,291]" pageId="13" pageNumber="16" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">Eudes-Deslongchamps, 1849</bibRefCitation>
)
</taxonomicName>
, p. 73, Pl. XV,
<figureCitation id="B6A02A5E6F7AFF86CE7AEA458FB7FEF4" box="[1053,1153,266,291]" captionStart="Figs 1–3" captionStartId="3.[219,260,1535,1557]" captionTargetBox="[222,1152,1201,1496]" captionTargetId="figure-324@3.[219,1156,1197,1499]" captionTargetPageId="3" captionText="Figs 1–3. The peculiar “quirinii type” ridges, occurring in Wortheniopsis Böhm, 1895, Foveolaria n. gen. and some probably related forms. – 1. Copy of KITTL’s (1899, p. 14, Fig. 2) sketch. – 2. The ridges on the outer face and on the base of Wortheniopsis rakusi n. sp. – 3. The ridges on the base of Foveolaria conoidea n. sp." figureDoi="http://doi.org/10.5281/zenodo.15627165" httpUri="https://zenodo.org/record/15627165/files/figure.png" pageId="13" pageNumber="16">Figs 2a, b</figureCitation>
.
<taxonomicName id="E99B4D586F7AFF86CB5DEA6588BDFE94" authority="(J. A. Eudes-Deslongchamps, 1849)" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[314,907,298,323]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="subturrita">
<emphasis id="1CEFEAC96F7AFF86CB5DEA658925FE94" box="[314,531,298,322]" italics="true" pageId="13" pageNumber="16">Foveolaria subturrita</emphasis>
(J. A.
<bibRefCitation id="4A0A4B2A6F7AFF86C832EA6588B5FE94" author="EUDES-DESLONGCHAMPS J. A." box="[597,899,298,323]" pageId="13" pageNumber="16" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">Eudes-Deslongchamps, 1849</bibRefCitation>
)
</taxonomicName>
, p. 73, Pl. XV,
<figureCitation id="B6A02A5E6F7AFF86CE7AEA658FB7FE94" box="[1053,1153,298,323]" captionStart="Figs 1–3" captionStartId="3.[219,260,1535,1557]" captionTargetBox="[222,1152,1201,1496]" captionTargetId="figure-324@3.[219,1156,1197,1499]" captionTargetPageId="3" captionText="Figs 1–3. The peculiar “quirinii type” ridges, occurring in Wortheniopsis Böhm, 1895, Foveolaria n. gen. and some probably related forms. – 1. Copy of KITTL’s (1899, p. 14, Fig. 2) sketch. – 2. The ridges on the outer face and on the base of Wortheniopsis rakusi n. sp. – 3. The ridges on the base of Foveolaria conoidea n. sp." figureDoi="http://doi.org/10.5281/zenodo.15627165" httpUri="https://zenodo.org/record/15627165/files/figure.png" pageId="13" pageNumber="16">Figs 3a, b</figureCitation>
.
<taxonomicName id="E99B4D586F7AFF86CB5DEA05884EFEB4" authority="(J. A. Eudes-Deslongchamps, 1849)" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[314,888,330,355]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="turrita">
<emphasis id="1CEFEAC96F7AFF86CB5DEA058AC0FEB4" box="[314,502,330,354]" italics="true" pageId="13" pageNumber="16">Foveolaria turrita</emphasis>
(J. A.
<bibRefCitation id="4A0A4B2A6F7AFF86C85CEA058846FEB4" author="EUDES-DESLONGCHAMPS J. A." box="[571,880,330,355]" pageId="13" pageNumber="16" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">Eudes-Deslongchamps, 1849</bibRefCitation>
)
</taxonomicName>
, p. 74, Pl. XV,
<figureCitation id="B6A02A5E6F7AFF86CE77EA058F41FEB4" box="[1040,1143,330,355]" captionStart="Figs 4–14" captionStartId="8.[174,214,1437,1459]" captionTargetBox="[174,1111,265,1412]" captionTargetId="figure-18@8.[174,1111,265,1412]" captionTargetPageId="8" captionText="Figs 4–14.Wortheniopsisspecies fromtheHierlatz LimestoneFormation.– 4–10.Wortheniopsisrakusi n.sp.from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe,Hallstatt, Austria,GBA 2017/001/0002,all ×4.– 4. Dorsal view.– 5. Peristomal view.– 6. Basal view. – 7–9. Same views as 4–6 but in natural (×1) size. – 10. Details of the ornament on thespire whorls,×7.– 11–14.Wortheniopsis bakonyensis n. sp.,?Lower Sinemurian (Lower Jurassic) Hierlatz Limestone variety with rich micrite content, Csárdahegy, Úrkút, Bakony Mts, Hungary, GGIH J.08.3.1. – 11. Peristomal view of the holotype, ×4. – 12. Lateral view to show the best preserved outline part, ×4.– 13–14. The same views as in figures 11–12 but in natural (×1) size" figureDoi="http://doi.org/10.5281/zenodo.15627167" httpUri="https://zenodo.org/record/15627167/files/figure.png" pageId="13" pageNumber="16">Figs 4a, b</figureCitation>
.
<taxonomicName id="E99B4D586F7AFF86CB5DEA25884DFE54" authority="(J. A. Eudes-Deslongchamps, 1849)" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[314,891,362,387]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="procera">
<emphasis id="1CEFEAC96F7AFF86CB5DEA258ACFFE54" box="[314,505,362,386]" italics="true" pageId="13" pageNumber="16">Foveolaria procera</emphasis>
(J. A.
<bibRefCitation id="4A0A4B2A6F7AFF86C859EA258845FE54" author="EUDES-DESLONGCHAMPS J. A." box="[574,883,362,387]" pageId="13" pageNumber="16" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">Eudes-Deslongchamps, 1849</bibRefCitation>
)
</taxonomicName>
, p. 74, Pl. XV,
<figureCitation id="B6A02A5E6F7AFF86CE74EA258F4CFE54" box="[1043,1146,362,387]" captionStart="Figs 4–14" captionStartId="8.[174,214,1437,1459]" captionTargetBox="[174,1111,265,1412]" captionTargetId="figure-18@8.[174,1111,265,1412]" captionTargetPageId="8" captionText="Figs 4–14.Wortheniopsisspecies fromtheHierlatz LimestoneFormation.– 4–10.Wortheniopsisrakusi n.sp.from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe,Hallstatt, Austria,GBA 2017/001/0002,all ×4.– 4. Dorsal view.– 5. Peristomal view.– 6. Basal view. – 7–9. Same views as 4–6 but in natural (×1) size. – 10. Details of the ornament on thespire whorls,×7.– 11–14.Wortheniopsis bakonyensis n. sp.,?Lower Sinemurian (Lower Jurassic) Hierlatz Limestone variety with rich micrite content, Csárdahegy, Úrkút, Bakony Mts, Hungary, GGIH J.08.3.1. – 11. Peristomal view of the holotype, ×4. – 12. Lateral view to show the best preserved outline part, ×4.– 13–14. The same views as in figures 11–12 but in natural (×1) size" figureDoi="http://doi.org/10.5281/zenodo.15627167" httpUri="https://zenodo.org/record/15627167/files/figure.png" pageId="13" pageNumber="16">Figs 5a, b</figureCitation>
.
<taxonomicName id="E99B4D586F7AFF86CB5DEAC58842FE74" authority="(J. A. Eudes-Deslongchamps, 1849)" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[314,884,394,419]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="pinguis">
<emphasis id="1CEFEAC96F7AFF86CB5DEAC58AC1FE74" box="[314,503,394,418]" italics="true" pageId="13" pageNumber="16">Foveolaria pinguis</emphasis>
(J. A.
<bibRefCitation id="4A0A4B2A6F7AFF86C85FEAC5885AFE74" author="EUDES-DESLONGCHAMPS J. A." box="[568,876,394,419]" pageId="13" pageNumber="16" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">Eudes-Deslongchamps, 1849</bibRefCitation>
)
</taxonomicName>
, p. 75, Pl. XV,
<figureCitation id="B6A02A5E6F7AFF86CE60EAC58FB7FE74" box="[1031,1153,394,419]" captionStart="Figs 4–14" captionStartId="8.[174,214,1437,1459]" captionTargetBox="[174,1111,265,1412]" captionTargetId="figure-18@8.[174,1111,265,1412]" captionTargetPageId="8" captionText="Figs 4–14.Wortheniopsisspecies fromtheHierlatz LimestoneFormation.– 4–10.Wortheniopsisrakusi n.sp.from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe,Hallstatt, Austria,GBA 2017/001/0002,all ×4.– 4. Dorsal view.– 5. Peristomal view.– 6. Basal view. – 7–9. Same views as 4–6 but in natural (×1) size. – 10. Details of the ornament on thespire whorls,×7.– 11–14.Wortheniopsis bakonyensis n. sp.,?Lower Sinemurian (Lower Jurassic) Hierlatz Limestone variety with rich micrite content, Csárdahegy, Úrkút, Bakony Mts, Hungary, GGIH J.08.3.1. – 11. Peristomal view of the holotype, ×4. – 12. Lateral view to show the best preserved outline part, ×4.– 13–14. The same views as in figures 11–12 but in natural (×1) size" figureDoi="http://doi.org/10.5281/zenodo.15627167" httpUri="https://zenodo.org/record/15627167/files/figure.png" pageId="13" pageNumber="16">Figs 6a, b, c</figureCitation>
.
<taxonomicName id="E99B4D586F7AFF86CB5DEAE588BDFE14" authority="(J. A. Eudes-Deslongchamps, 1849)" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[314,907,426,451]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="ellipsoidea">
<emphasis id="1CEFEAC96F7AFF86CB5DEAE58925FE14" box="[314,531,426,450]" italics="true" pageId="13" pageNumber="16">Foveolaria ellipsoidea</emphasis>
(J. A.
<bibRefCitation id="4A0A4B2A6F7AFF86C832EAE588B5FE14" author="EUDES-DESLONGCHAMPS J. A." box="[597,899,426,451]" pageId="13" pageNumber="16" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">Eudes-Deslongchamps, 1849</bibRefCitation>
)
</taxonomicName>
, p. 75, Pl. XV,
<figureCitation id="B6A02A5E6F7AFF86CE7AEAE58FB7FE14" box="[1053,1153,426,451]" captionStart="Figs 4–14" captionStartId="8.[174,214,1437,1459]" captionTargetBox="[174,1111,265,1412]" captionTargetId="figure-18@8.[174,1111,265,1412]" captionTargetPageId="8" captionText="Figs 4–14.Wortheniopsisspecies fromtheHierlatz LimestoneFormation.– 4–10.Wortheniopsisrakusi n.sp.from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe,Hallstatt, Austria,GBA 2017/001/0002,all ×4.– 4. Dorsal view.– 5. Peristomal view.– 6. Basal view. – 7–9. Same views as 4–6 but in natural (×1) size. – 10. Details of the ornament on thespire whorls,×7.– 11–14.Wortheniopsis bakonyensis n. sp.,?Lower Sinemurian (Lower Jurassic) Hierlatz Limestone variety with rich micrite content, Csárdahegy, Úrkút, Bakony Mts, Hungary, GGIH J.08.3.1. – 11. Peristomal view of the holotype, ×4. – 12. Lateral view to show the best preserved outline part, ×4.– 13–14. The same views as in figures 11–12 but in natural (×1) size" figureDoi="http://doi.org/10.5281/zenodo.15627167" httpUri="https://zenodo.org/record/15627167/files/figure.png" pageId="13" pageNumber="16">Figs 7a, b</figureCitation>
.
<taxonomicName id="E99B4D586F7AFF86CB5DEA858991FE34" authority="(Szabo, 2009)" baseAuthorityName="Szabo" baseAuthorityYear="2009" box="[314,679,458,482]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="urkutensis">
<emphasis id="1CEFEAC96F7AFF86CB5DEA858923FE34" box="[314,533,458,482]" italics="true" pageId="13" pageNumber="16">Foveolaria urkutensis</emphasis>
(
<bibRefCitation id="4A0A4B2A6F7AFF86C846EA858996FE34" author="SZABO J." box="[545,672,458,482]" pageId="13" pageNumber="16" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
)
</taxonomicName>
, p. 29,
<figureCitation id="B6A02A5E6F7AFF86C88BEA858805FE34" box="[748,819,458,483]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="13" pageNumber="16">Figs 23</figureCitation>
A-E.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="668165506F7AFF86CB5DEAA58AFAFCD4" pageId="13" pageNumber="16" type="taxon_list">
<paragraph id="2E2436DB6F7AFF86CB5DEAA58848FDD4" blockId="13.[314,1157,266,770]" box="[314,894,490,515]" pageId="13" pageNumber="16">
<taxonomicName id="E99B4D586F7AFF86CB5DEAA589F5FDD4" authority="(Szabo, 2009)" baseAuthorityName="Szabo" baseAuthorityYear="2009" box="[314,707,490,514]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="hierlatzensis">
<emphasis id="1CEFEAC96F7AFF86CB5DEAA58906FDD4" box="[314,560,490,514]" italics="true" pageId="13" pageNumber="16">Foveolaria hierlatzensis</emphasis>
(
<bibRefCitation id="4A0A4B2A6F7AFF86C85AEAA5898AFDD4" author="SZABO J." box="[573,700,490,514]" pageId="13" pageNumber="16" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
)
</taxonomicName>
, p. 30,
<figureCitation id="B6A02A5E6F7AFF86C96FEAA58866FDD4" box="[776,848,490,515]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="13" pageNumber="16">Figs 24</figureCitation>
A-F.
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB5DE945880BFDF4" blockId="13.[314,1157,266,770]" box="[314,829,522,547]" pageId="13" pageNumber="16">
<taxonomicName id="E99B4D586F7AFF86CB5DE94589B5FDF4" authority="(Szabo, 2009)" baseAuthorityName="Szabo" baseAuthorityYear="2009" box="[314,643,522,546]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="jancsii">
<emphasis id="1CEFEAC96F7AFF86CB5DE9458AC6FDF4" box="[314,496,522,546]" italics="true" pageId="13" pageNumber="16">Foveolaria jancsii</emphasis>
(
<bibRefCitation id="4A0A4B2A6F7AFF86CB9AE945894AFDF4" author="SZABO J." box="[509,636,522,546]" pageId="13" pageNumber="16" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
)
</taxonomicName>
, p. 31,
<figureCitation id="B6A02A5E6F7AFF86C8AFE9458839FDF4" box="[712,783,522,547]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="13" pageNumber="16">Figs 25</figureCitation>
A-P.
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB5DE9658855FD94" blockId="13.[314,1157,266,770]" box="[314,867,554,579]" pageId="13" pageNumber="16">
<taxonomicName id="E99B4D586F7AFF86CB5DE9658990FD94" authority="(Szabo, 2009)" baseAuthorityName="Szabo" baseAuthorityYear="2009" box="[314,678,554,578]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="lokutensis">
<emphasis id="1CEFEAC96F7AFF86CB5DE9658925FD94" box="[314,531,554,578]" italics="true" pageId="13" pageNumber="16">Foveolaria lokutensis</emphasis>
(
<bibRefCitation id="4A0A4B2A6F7AFF86C847E96589A9FD94" author="SZABO J." box="[544,671,554,578]" pageId="13" pageNumber="16" pagination="1 - 108" refId="ref12819" refString="SZABO J. 2009: Gastropods of the Early Jurassic Hierlatz Limestone Formation; part 1: a revision of type collections from Austrian and Hungarian localities. - Fragmenta Palaeontologica Hungarica 26: 1-108." type="journal article" year="2009">Szabó, 2009</bibRefCitation>
)
</taxonomicName>
, p. 32,
<figureCitation id="B6A02A5E6F7AFF86C88DE9658805FD94" box="[746,819,554,579]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="13" pageNumber="16">Figs 26</figureCitation>
A-E.
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB5DE905896FFDB4" blockId="13.[314,1157,266,770]" box="[314,601,586,610]" pageId="13" pageNumber="16">
<taxonomicName id="E99B4D586F7AFF86CB5DE905892DFDB4" authorityName="Szabó" authorityYear="2016" box="[314,539,586,610]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="bicarinata" status="sp. nov.">
<emphasis id="1CEFEAC96F7AFF86CB5DE905892DFDB4" box="[314,539,586,610]" italics="true" pageId="13" pageNumber="16">Foveolaria bicarinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7AFF86C846E904896FFDB4" box="[545,601,587,610]" pageId="13" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB5DE9258972FD54" blockId="13.[314,1157,266,770]" box="[314,580,618,642]" pageId="13" pageNumber="16">
<taxonomicName id="E99B4D586F7AFF86CB5DE9258931FD54" authorityName="Szabó" authorityYear="2016" box="[314,519,618,642]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="conoidea" status="sp. nov.">
<emphasis id="1CEFEAC96F7AFF86CB5DE9258931FD54" box="[314,519,618,642]" italics="true" pageId="13" pageNumber="16">Foveolaria conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7AFF86C86AE9248972FD54" box="[525,580,619,642]" pageId="13" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB5DE9C58903FD74" blockId="13.[314,1157,266,770]" box="[314,565,650,674]" pageId="13" pageNumber="16">
<taxonomicName id="E99B4D586F7AFF86CB5DE9C58ACEFD74" authorityName="Szabó" authorityYear="2016" box="[314,504,650,674]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="globosa" status="sp. nov.">
<emphasis id="1CEFEAC96F7AFF86CB5DE9C58ACEFD74" box="[314,504,650,674]" italics="true" pageId="13" pageNumber="16">Foveolaria globosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7AFF86CB99E9C48903FD74" box="[510,565,651,674]" pageId="13" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB5DE9E58953FD14" blockId="13.[314,1157,266,770]" box="[314,613,682,706]" pageId="13" pageNumber="16">
<taxonomicName id="E99B4D586F7AFF86CB5DE9E58911FD14" authorityName="Szabó" authorityYear="2016" box="[314,551,682,706]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="suemegensis" status="sp. nov.">
<emphasis id="1CEFEAC96F7AFF86CB5DE9E58911FD14" box="[314,551,682,706]" italics="true" pageId="13" pageNumber="16">Foveolaria suemegensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7AFF86C84AE9E48953FD14" box="[557,613,683,706]" pageId="13" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB5DE985891BFD34" blockId="13.[314,1157,266,770]" box="[314,557,714,738]" pageId="13" pageNumber="16">
<taxonomicName id="E99B4D586F7AFF86CB5DE9858AD9FD34" authorityName="Szabó" authorityYear="2016" box="[314,495,714,738]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="kocsisi" status="sp. nov.">
<emphasis id="1CEFEAC96F7AFF86CB5DE9858AD9FD34" box="[314,495,714,738]" italics="true" pageId="13" pageNumber="16">Foveolaria kocsisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7AFF86CB92E984891BFD34" box="[501,557,715,738]" pageId="13" pageNumber="16" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="2E2436DB6F7AFF86CB5DE9A58AFAFCD4" blockId="13.[314,1157,266,770]" box="[314,460,746,770]" pageId="13" pageNumber="16">
<taxonomicName id="E99B4D586F7AFF86CB5DE9A58AFAFCD4" box="[314,460,746,770]" class="Gymnolaemata" family="Foveolariidae" genus="Foveolaria" kingdom="Animalia" order="Cheilostomatida" pageId="13" pageNumber="16" phylum="Bryozoa" rank="species" species="undetermined">
<emphasis id="1CEFEAC96F7AFF86CB5DE9A58A9FFCD4" box="[314,425,746,770]" italics="true" pageId="13" pageNumber="16">Foveolaria</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="668165506F7AFF86CB71E860888DFCBB" pageId="13" pageNumber="16" type="distribution">
<paragraph id="2E2436DB6F7AFF86CB71E860888DFCBB" blockId="13.[219,1156,814,877]" pageId="13" pageNumber="16">
<emphasis id="1CEFEAC96F7AFF86CB71E8608A92FC9F" box="[278,420,815,841]" italics="true" pageId="13" pageNumber="16">Distribution</emphasis>
– Sinemurian to Pliensbachian (Jurassic) in deposits of the Western Tethys and the Western European epicontinental seas.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="5A0E4E56F455B05F53192177D84D1686">Gastropods of the Lower Jurassic Hierlatz Limestone Formation, part 2. Some new archaic type slit-bearing components from the fauna of the Hierlatz Alpe (Hallstatt, Austria) and the Bakony Mts (Hungary)</mods:title>
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<mods:namePart id="CC409C183E4F3C2654434E5877598642">Szabó, János</mods:namePart>
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<subSubSection id="668165506F7EFF82CB9EE9258851FD53" box="[505,871,618,645]" pageId="9" pageNumber="12" type="nomenclature">
<paragraph id="2E2436DB6F7EFF82CB9EE9258851FD53" blockId="9.[505,871,618,680]" box="[505,871,618,645]" pageId="9" pageNumber="12">
<heading id="756C81B76F7EFF82CB9EE9258851FD53" box="[505,871,618,645]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="12" reason="9">
<taxonomicName id="E99B4D586F7EFF82CB9EE9258812FD52" authorityName="Szabó" authorityYear="2016" box="[505,804,618,644]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="bakonyensis" status="sp. nov.">
<emphasis id="1CEFEAC96F7EFF82CB9EE9258812FD52" box="[505,804,618,644]" italics="true" pageId="9" pageNumber="12">Wortheniopsis bakonyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7EFF82C94DE9248851FD53" box="[810,871,619,645]" pageId="9" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="668165506F7EFF82C801E9C289CCFD7E" box="[614,762,653,680]" pageId="9" pageNumber="12" type="description">
<paragraph id="2E2436DB6F7EFF82C801E9C289CCFD7E" blockId="9.[505,871,618,680]" box="[614,762,653,680]" pageId="9" pageNumber="12">
(
<figureCitation id="B6A02A5E6F7EFF82C809E9C289C7FD7E" box="[622,753,653,680]" captionStart="Figs 4–14" captionStartId="8.[174,214,1437,1459]" captionTargetBox="[174,1111,265,1412]" captionTargetId="figure-18@8.[174,1111,265,1412]" captionTargetPageId="8" captionText="Figs 4–14.Wortheniopsisspecies fromtheHierlatz LimestoneFormation.– 4–10.Wortheniopsisrakusi n.sp.from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe,Hallstatt, Austria,GBA 2017/001/0002,all ×4.– 4. Dorsal view.– 5. Peristomal view.– 6. Basal view. – 7–9. Same views as 4–6 but in natural (×1) size. – 10. Details of the ornament on thespire whorls,×7.– 11–14.Wortheniopsis bakonyensis n. sp.,?Lower Sinemurian (Lower Jurassic) Hierlatz Limestone variety with rich micrite content, Csárdahegy, Úrkút, Bakony Mts, Hungary, GGIH J.08.3.1. – 11. Peristomal view of the holotype, ×4. – 12. Lateral view to show the best preserved outline part, ×4.– 13–14. The same views as in figures 11–12 but in natural (×1) size" figureDoi="http://doi.org/10.5281/zenodo.15627167" httpUri="https://zenodo.org/record/15627167/files/figure.png" pageId="9" pageNumber="12">Figs 11–14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="668165506F7EFF82CABCE9F08A6BFD24" pageId="9" pageNumber="12" type="reference_group">
<paragraph id="2E2436DB6F7EFF82CABCE9F08A6BFD24" blockId="9.[219,1157,703,754]" pageId="9" pageNumber="12">
2009
<taxonomicName id="E99B4D586F7EFF82CB68E98F88D8FD03" authority="(J. A. Eudes-Deslongchamps, 1849)" baseAuthorityName="J. A. Eudes-Deslongchamps" baseAuthorityYear="1849" box="[271,1006,703,725]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" isUncertain="true" kingdom="Animalia" order="Pleurotomariida" pageId="9" pageNumber="12" phylum="Mollusca" rank="species" species="procera" subGenus="Wortheniopsis">
<emphasis id="1CEFEAC96F7EFF82CB68E98F8AA7FD03" box="[271,401,704,725]" italics="true" pageId="9" pageNumber="12">Wortheniopsis</emphasis>
(
<emphasis id="1CEFEAC96F7EFF82CBF9E98F8916FD03" box="[414,544,704,725]" italics="true" pageId="9" pageNumber="12">Wortheniopsis</emphasis>
) aff.
<emphasis id="1CEFEAC96F7EFF82C837E98F89A5FD03" box="[592,659,704,725]" italics="true" pageId="9" pageNumber="12">procera</emphasis>
(J. A.
<bibRefCitation id="4A0A4B2A6F7EFF82C8B5E9F088D0FD03" author="EUDES-DESLONGCHAMPS J. A." box="[722,998,703,725]" pageId="9" pageNumber="12" pagination="1 - 157" refId="ref11878" refString="EUDES-DESLONGCHAMPS J. A. 1849: Memoires sur les Pleurotomaries des terrains secondaires du Calvados. - Memoires de la Societe Linneenne de Normandie 8: 1-157, Pls 1 - 18." type="journal article" year="1849">Eudes-Deslongchamps, 1849</bibRefCitation>
)
</taxonomicName>
– SZABÓ, p. 29,
<figureCitation id="B6A02A5E6F7EFF82CB71E9938A6CFD24" box="[278,346,732,754]" captionStart="Figs 15–29" captionStartId="15.[219,260,1435,1457]" captionTargetId="figure-18@16.[174,1112,289,1639]" captionTargetPageId="16" captionText="Figs 15–29. New species of Foveolaria n. gen. from the Sinemurian to lowermost Pliensbachian interval (Lower Jurassic) of the Hierlatz Limestone in the Hierlatz Alpe, Hallstatt, Austria. – 15–18. Holotype of Foveolaria bicarinata n. sp., GBA 2017/001/0003, all ×3.8. – 15. Dorsal view. – 16. Apical view. – 17. Basal view. – 18. Peristomal view. – 19–22. Holotype of Foveolaria conoidea n. sp., GBA 2017/001/0004, all ×4. – 19. Dorsal view. – 20. Apical view. – 21. Basal view. – 22. Peristomal view.– 23–26. Holotype of Foveolaria globosa n. sp., GBA 2017/001/0005, all ×4. – 23. Dorsal view. – 24. Apical view. – 25. Basal view. – 26. Peristomal view. – 27–29. Foveolaria sp., GBA 2017/001/0006, all ×4. – 27. Apertural view. – 28. Basal view. – 29. Dorsal view" pageId="9" pageNumber="12">Fig. 22</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="668165506F7EFF81CB71E8558A80FB0D" lastPageId="10" lastPageNumber="13" pageId="9" pageNumber="12" type="materials_examined">
<materialsCitation id="9EF33C866F7EFF82CB71E85588F6FC81" collectionCode="GGIH" country="Hungary" county="Bakony Mts" location="Csardahegy" municipality="Urkut" pageId="9" pageNumber="12" specimenCount="1" typeStatus="holotype">
<paragraph id="2E2436DB6F7EFF82CB71E85589BEFCE5" blockId="9.[219,1157,793,1664]" box="[278,648,793,819]" pageId="9" pageNumber="12">
<emphasis id="1CEFEAC96F7EFF82CB71E8558A8CFCE5" box="[278,442,794,819]" italics="true" pageId="9" pageNumber="12">
<typeStatus id="F12088796F7EFF82CB71E8558A78FCE5" box="[278,334,794,819]" pageId="9" pageNumber="12">Type</typeStatus>
specimen
</emphasis>
–
<collectionCode id="488AAE1E6F7EFF82CBB1E8568912FCE5" box="[470,548,793,819]" pageId="9" pageNumber="12">GGIH</collectionCode>
J.08.3.1.
</paragraph>
<paragraph id="2E2436DB6F7EFF82CB71E87288F2FC81" blockId="9.[219,1157,793,1664]" box="[278,964,828,855]" pageId="9" pageNumber="12">
<emphasis id="1CEFEAC96F7EFF82CB71E8728A91FC80" box="[278,423,828,854]" italics="true" pageId="9" pageNumber="12">
<typeStatus id="F12088796F7EFF82CB71E8728A78FC80" box="[278,334,829,854]" pageId="9" pageNumber="12">Type</typeStatus>
locality
</emphasis>
–
<location id="2B4460006F7EFF82CBA4E8738979FC80" LSID="urn:lsid:plazi:treatment:A63287CD6F7EFF81CB9EE9258A80FB0D:2B4460006F7EFF82CBA4E8738979FC80" box="[451,591,828,854]" country="Hungary" county="Bakony Mts" municipality="Urkut" name="Csardahegy" pageId="9" pageNumber="12">Csárdahegy</location>
,
<collectingMunicipality id="CE40ACA16F7EFF82C83EE8738994FC81" box="[601,674,828,855]" pageId="9" pageNumber="12">Úrkút</collectingMunicipality>
,
<collectingCounty id="C7454E576F7EFF82C8CAE873880FFC81" box="[685,825,828,855]" pageId="9" pageNumber="12">Bakony Mts</collectingCounty>
(
<collectingCountry id="568C764B6F7EFF82C92FE873888EFC81" box="[840,952,828,855]" name="Hungary" pageId="9" pageNumber="12">Hungary</collectingCountry>
).
</paragraph>
</materialsCitation>
<paragraph id="2E2436DB6F7EFF82CB71E82F8995FC4B" blockId="9.[219,1157,793,1664]" pageId="9" pageNumber="12">
<materialsCitation id="9EF33C866F7EFF82CB71E82F89A9FC4B" county="Hierlatz Limestone" location="Lower" municipality="Lower Jurassic" pageId="9" pageNumber="12" specimenCount="1" typeStatus="holotype">
<emphasis id="1CEFEAC96F7EFF82CB71E82F8AA0FCAF" box="[278,406,864,889]" italics="true" pageId="9" pageNumber="12">
<typeStatus id="F12088796F7EFF82CB71E82F8A79FCAF" box="[278,335,864,889]" pageId="9" pageNumber="12">Type</typeStatus>
strata
</emphasis>
–
<location id="2B4460006F7EFF82CBD3E8108AC9FCAF" LSID="urn:lsid:plazi:treatment:A63287CD6F7EFF81CB9EE9258A80FB0D:2B4460006F7EFF82CBD3E8108AC9FCAF" box="[436,511,863,889]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Lower" pageId="9" pageNumber="12">Lower</location>
?
<location id="2B4460006F7EFF82C868E81089A1FCAF" LSID="urn:lsid:plazi:treatment:A63287CD6F7EFF81CB9EE9258A80FB0D:2B4460006F7EFF82C868E81089A1FCAF" box="[527,663,863,889]" county="Hierlatz Limestone" municipality="Lower Jurassic" name="Sinemurian" pageId="9" pageNumber="12">Sinemurian</location>
(
<collectingMunicipality id="CE40ACA16F7EFF82C8C1E8108863FCAC" box="[678,853,863,890]" pageId="9" pageNumber="12">Lower Jurassic</collectingMunicipality>
)
<collectingCounty id="C7454E576F7EFF82C903E8108F71FCAF" box="[868,1095,863,889]" pageId="9" pageNumber="12">Hierlatz Limestone</collectingCounty>
<taxonomicName id="E99B4D586F7EFF82CE29E8108A0AFC4A" pageId="9" pageNumber="12" rank="variety" variety="with">variety with</taxonomicName>
relatively rich micrite content
</materialsCitation>
.
</paragraph>
<paragraph id="2E2436DB6F7EFF82CB71E8E98A59FC35" blockId="9.[219,1157,793,1664]" pageId="9" pageNumber="12">
<emphasis id="1CEFEAC96F7EFF82CB71E8E98AD8FC69" box="[278,494,934,960]" italics="true" pageId="9" pageNumber="12">Derivation of name</emphasis>
– Referring to the wider surroundings of the locality, the Bakony Mts.
</paragraph>
<paragraph id="2E2436DB6F7EFF82CB71E8A38878FBFF" blockId="9.[219,1157,793,1664]" pageId="9" pageNumber="12">
<emphasis id="1CEFEAC96F7EFF82CB71E8A38A4DFBD0" box="[278,379,1004,1030]" italics="true" pageId="9" pageNumber="12">Material</emphasis>
– Single specimen that consists of three partial whorls; the earliest whorls and the near-peristome parts are fully lacking.
</paragraph>
<paragraph id="2E2436DB6F7EFF82CB71EF7C8802FB9B" blockId="9.[219,1157,793,1664]" box="[278,820,1074,1101]" pageId="9" pageNumber="12">
<emphasis id="1CEFEAC96F7EFF82CB71EF7C8AAFFB9A" box="[278,409,1075,1100]" italics="true" pageId="9" pageNumber="12">Dimensions</emphasis>
– Width:
<quantity id="E9639B3E6F7EFF82C875EF7C895EFB9B" box="[530,616,1075,1101]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="12" unit="mm" value="14.0">14 mm</quantity>
; spiral angle: 50°.
</paragraph>
<paragraph id="2E2436DB6F7EFF82CB71EF1989ACFAB1" blockId="9.[219,1157,793,1664]" pageId="9" pageNumber="12">
<emphasis id="1CEFEAC96F7EFF82CB71EF198AB5FBB9" box="[278,387,1110,1135]" italics="true" pageId="9" pageNumber="12">Diagnosis</emphasis>
– Ramp flat and steep, outer face concave; earlier whorls rounded-angular above suture, last whorl evenly rounded toward base. Concave selenizone between two sharp ridges, abapical one just overlapping angulation of whorl surface. Base subconoidal as a whole with slightly convex wall; moderately broad umbilicus with rounded-angular rim. Umbilical lip seeming subvertical. Spiral ornament on full shell surface, excepting selenizone, threads on ramp, quirinii
<typeStatus id="F12088796F7EFF82CABCEE668A39FA95" box="[219,271,1321,1347]" pageId="9" pageNumber="12">type</typeStatus>
ridges on outer face and base, all finely granulated even within umbilicus. Collabral threads observable on ramp.
</paragraph>
<paragraph id="2E2436DB6F7EFF81CB71EE3F8986FE67" blockId="9.[219,1157,793,1664]" lastBlockId="10.[174,1114,266,1243]" lastPageId="10" lastPageNumber="13" pageId="9" pageNumber="12">
<emphasis id="1CEFEAC96F7EFF82CB71EE3F8AA1FA5F" box="[278,407,1392,1417]" italics="true" pageId="9" pageNumber="12">Description</emphasis>
– The shell is high turbiniform, extremely thin walled with slightly convex whorls that have an angulation above the midwhorl. The ramp is feebly concave on the earlier whorls but flat on the last whorl. The selenizone is slightly protruding from its environs and it is fairly concave on all whorls; its position is on the ramp just along the angulation. Abapically, a concave outer whorl face limits the selenizone, which is wider than the ramp and rounded-angular above the suture; this angulation is more marked on the earlier whorls and widely rounded, practically absent on the last whorl. The suture appears deeply impressed since it is built somewhat adaxially from the periphery of the whorls. The base is subconoidal with slightly convex wall and a moderately wide umbilicus, having a rounded-angular rim. Remnants suggest subaxial orientation of the umbilical lip and no trace of the parietal lip has been found; no more information, concerning the peristome is preserved.
</paragraph>
<paragraph id="2E2436DB6F7DFF81CA8EEAF5889DFCC7" blockId="10.[174,1114,266,1243]" pageId="10" pageNumber="13">
All whorls are covered by spiral ornament, except the selenizone. It consists of spiral threads on the ramp and quirinii
<typeStatus id="F12088796F7DFF81C8C5EA9189E0FE2E" box="[674,726,478,504]" pageId="10" pageNumber="13">type</typeStatus>
ridges on the outer face and the base. Weak collabral threads are observable on the ramp and the outer face of the earliest whorl with granules at the crossings with the spiral threads. The collabral threads of the ramp are vanishing only on the last whorl but lacking already from the outer face of the penultimate whorl. However, the granules remain present also on the spiral threads and ridges of the latest shell parts, even on the base and within the umbilicus. The growth lines and the collabral ornament are feebly prosocline and clearly prosocyrt on both sides along the selenizone. The growth lines of the base are nearly orthocline and somewhat opisthocyrt.
</paragraph>
<paragraph id="2E2436DB6F7DFF81CA8EE8558AFCFCAD" blockId="10.[174,1114,266,1243]" pageId="10" pageNumber="13">
<emphasis id="1CEFEAC96F7DFF81CA8EE8558A7BFCE2" box="[233,333,794,820]" italics="true" pageId="10" pageNumber="13">Remarks</emphasis>
– Granules of the spiral ridges on the base are almost fully removed by erosion, however some remnants indicate in strong magnification that they are present on all ridges.
</paragraph>
<paragraph id="2E2436DB6F7DFF81CA8EE8CB8F3DFB42" blockId="10.[174,1114,266,1243]" pageId="10" pageNumber="13">
<taxonomicName id="E99B4D586F7DFF81CA8EE8CB8920FC48" authorityName="Szabó" authorityYear="2016" box="[233,534,900,926]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="bakonyensis" status="sp. nov.">
<emphasis id="1CEFEAC96F7DFF81CA8EE8CB8920FC48" box="[233,534,900,926]" italics="true" pageId="10" pageNumber="13">Wortheniopsis bakonyensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7DFF81C879E8CB8968FC48" box="[542,606,900,926]" pageId="10" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
differs from
<taxonomicName id="E99B4D586F7DFF81C89AE8CA885CFC48" authorityName="Szabó" authorityYear="2016" box="[765,874,900,926]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="rakusi" status="n. sp.">
<emphasis id="1CEFEAC96F7DFF81C89AE8CA885CFC48" box="[765,874,900,926]" italics="true" pageId="10" pageNumber="13">W. rakusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07DC57B26F7DFF81C915E8CB8887FC48" box="[882,945,900,926]" pageId="10" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
in having the angulation of the whorl surface from a much earlier growth stage; i. e. all whorls (3) are angulated in
<taxonomicName id="E99B4D586F7DFF81CBC7E8848979FC32" authorityName="Szabó" authorityYear="2016" box="[416,591,970,997]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="bakonyensis">
<emphasis id="1CEFEAC96F7DFF81CBC7E8848A8CFC32" box="[416,442,971,996]" italics="true" pageId="10" pageNumber="13">W</emphasis>
.
<emphasis id="1CEFEAC96F7DFF81CBAEE8858979FC32" box="[457,591,970,996]" italics="true" pageId="10" pageNumber="13">bakonyensis</emphasis>
</taxonomicName>
but only the last one in
<taxonomicName id="E99B4D586F7DFF81C911E88488D4FC32" authorityName="Szabó" authorityYear="2016" box="[886,994,970,996]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="rakusi">
<emphasis id="1CEFEAC96F7DFF81C911E88488D4FC32" box="[886,994,970,996]" italics="true" pageId="10" pageNumber="13">W. rakusi</emphasis>
</taxonomicName>
. It means also that a ramp has developed only on the last whorl of
<taxonomicName id="E99B4D586F7DFF81C93DE8A188F1FBD1" authorityName="Szabó" authorityYear="2016" box="[858,967,1005,1031]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="rakusi">
<emphasis id="1CEFEAC96F7DFF81C93DE8A188F1FBD1" box="[858,967,1005,1031]" italics="true" pageId="10" pageNumber="13">W. rakusi</emphasis>
</taxonomicName>
while in
<taxonomicName id="E99B4D586F7DFF81CE50E8A18A02FBFD" authorityName="Szabó" authorityYear="2016" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="bakonyensis">
<emphasis id="1CEFEAC96F7DFF81CE50E8A18F67FBD1" box="[1079,1105,1006,1031]" italics="true" pageId="10" pageNumber="13">W</emphasis>
.
<emphasis id="1CEFEAC96F7DFF81CAC9EF5E8A02FBFD" box="[174,308,1041,1067]" italics="true" pageId="10" pageNumber="13">bakonyensis</emphasis>
</taxonomicName>
it is present (at least) two whorls earlier.
<taxonomicName id="E99B4D586F7DFF81C94EEF5D88EEFBFD" authorityName="Szabó" authorityYear="2016" box="[809,984,1041,1067]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="bakonyensis">
<emphasis id="1CEFEAC96F7DFF81C94EEF5D8875FBFD" box="[809,835,1042,1067]" italics="true" pageId="10" pageNumber="13">W</emphasis>
.
<emphasis id="1CEFEAC96F7DFF81C935EF5E88EEFBFD" box="[850,984,1041,1067]" italics="true" pageId="10" pageNumber="13">bakonyensis</emphasis>
</taxonomicName>
has feebly concave outer face on all preserved whorls but
<taxonomicName id="E99B4D586F7DFF81C883EF7A8867FB98" authorityName="Szabó" authorityYear="2016" box="[740,849,1076,1102]" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="rakusi">
<emphasis id="1CEFEAC96F7DFF81C883EF7A8867FB98" box="[740,849,1076,1102]" italics="true" pageId="10" pageNumber="13">W. rakusi</emphasis>
</taxonomicName>
has flat outer face on the last whorl and feebly convex corresponding belt on the former whorls. In
<taxonomicName id="E99B4D586F7DFF81CE5DEF178BC3FB42" authorityName="Szabó" authorityYear="2016" class="Gastropoda" family="Plethospiridae" genus="Wortheniopsis" kingdom="Animalia" order="Pleurotomariida" pageId="10" pageNumber="13" phylum="Mollusca" rank="species" species="rakusi">
<emphasis id="1CEFEAC96F7DFF81CE5DEF178BC3FB42" italics="true" pageId="10" pageNumber="13">W. rakusi</emphasis>
</taxonomicName>
, no granulation is present neither on the outer face nor on the base.
</paragraph>
<paragraph id="2E2436DB6F7DFF81CA8EEFD28A80FB0D" blockId="10.[174,1114,266,1243]" pageId="10" pageNumber="13">
<emphasis id="1CEFEAC96F7DFF81CA8EEFD28A41FB61" box="[233,375,1181,1207]" italics="true" pageId="10" pageNumber="13">Distribution</emphasis>
– Lower? Sinemurian Hierlatz Limestone, Csárdahegy, Úrkút, Bakony Mts,
<collectingCountry id="568C764B6F7DFF81CB20EF8E8A87FB0D" box="[327,433,1217,1243]" name="Hungary" pageId="10" pageNumber="13">Hungary</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>