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<document id="CB05796CC5EBB27A08225D87BEEA69C5" ID-DOI="10.11646/phytotaxa.158.1.9" ID-ISSN="1179-3163" ID-Zenodo-Dep="5128216" IM.bibliography_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1627047514536" checkinUser="felipe" docAuthor="Jiménez, Juan A., Toren, David &amp; Shevock, James R." docDate="2014" docId="38491715FFE2CD5CDFBBF80C0709FAE9" docLanguage="en" docName="Phytotaxa.158.1.105-110.pdf" docOrigin="Phytotaxa 158 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.158.1.9" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Didymodon californicus J. A. Jimenez, Toren &amp; Shevock 2014, sp. nov." docType="treatment" docVersion="2" lastPageNumber="108" masterDocId="C4706F6DFFE2CD5FDF29FFA6023AFFBC" masterDocTitle="Didymodon californicus (Pottiaceae), a new species from California, U. S. A." masterLastPageNumber="110" masterPageNumber="105" pageNumber="105" updateTime="1744286783065" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3A7BC7ABCCA1F33C668A6CB6142E82DD">Didymodon californicus (Pottiaceae), a new species from California, U. S. A.</mods:title>
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<mods:namePart id="6CB613617E8E1DB86BABF44CA7C46B4D">Jiménez, Juan A.</mods:namePart>
<mods:affiliation id="36110C9EB43450AAB058D39D247CC247">Departamento de Biología Vegetal (Botánica), Facultad de Biología, Universidad de Murcia, E- 30100 Murcia, Spain.</mods:affiliation>
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<mods:namePart id="A1F2186AE74E5905D301E5E4AD42BBC0">Toren, David</mods:namePart>
<mods:affiliation id="06CFB6993127F4BDE8DA01B5E1DE0E81">Department of Botany, California Academy of Sciences, 55 Music Concourse Dr., Golden Gate Park, San Francisco, CA 94118 U. S. A.</mods:affiliation>
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<mods:namePart id="47D2068D7964895133160920E64366D8">Shevock, James R.</mods:namePart>
<mods:affiliation id="B4DEDC6CCDC267D32AB5DEED228F17E0">Department of Botany, California Academy of Sciences, 55 Music Concourse Dr., Golden Gate Park, San Francisco, CA 94118 U. S. A.</mods:affiliation>
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<taxonomicName id="77E0DD80FFE2CD5FDFBBF80C0120F87F" authority="J. A. Jimenez, Toren &amp; Shevock" authorityName="J. A. Jimenez, Toren &amp; Shevock" authorityYear="2014" box="[146,794,1962,1987]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="0" pageNumber="105" phylum="Bryophyta" rank="species" species="californicus" status="sp. nov.">
<emphasis id="82947A11FFE2CD5FDFBBF80C0393F87F" bold="true" box="[146,425,1962,1987]" italics="true" pageId="0" pageNumber="105">Didymodon californicus</emphasis>
J. A. Jiménez, Toren &amp; Shevock
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<emphasis id="82947A11FFE2CD5FDC08F80D014CF87E" box="[801,886,1963,1986]" italics="true" pageId="0" pageNumber="105">
<taxonomicNameLabel id="99A7C76AFFE2CD5FDC08F80D014CF87E" box="[801,886,1963,1986]" pageId="0" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="28DBBA86FFE2CD5FDCADF80C01E8F87F" box="[900,978,1962,1987]" captionStart="FIGURE 1" captionStartId="1.[151,246,1938,1960]" captionTargetBox="[200,1381,384,1915]" captionTargetId="figure-171@1.[172,1415,349,1951]" captionTargetPageId="1" captionText="FIGURE 1. Didymodon californicus. A. Habit, dry. B, C, D, E. Leaves. F. Ventral surface of leaf apex. G. Basal laminal cells. H. Upper laminal cells. I, J, K. Transverse sections of a leaf from basal to apical part. L. Axillary hair. M. Transverse section of stem. Scale bars: A = 2 mm. BE = 400 µm. F = 60 µm. G = 40 µm. H, L = 30 µm. IK, M = 50 µm. All from holotype." figureDoi="http://doi.org/10.5281/zenodo.5128218" httpUri="https://zenodo.org/record/5128218/files/figure.png" pageId="0" pageNumber="105">Figs. 1</figureCitation>
,
<figureCitation id="28DBBA86FFE2CD5FDCC9F80C01D5F87F" box="[992,1007,1962,1987]" captionStart="FIGURE 2" captionStartId="2.[151,244,1682,1704]" captionTargetBox="[155,1436,155,1648]" captionTargetId="figure-168@2.[155,1440,155,1667]" captionTargetPageId="2" captionText="FIGURE 2. A. Habit, dry. B. Habit, wet. C. Ventral surface of leaf apex. D. Dorsal surface of leaf apex. E. Transverse section of a leaf showing bistratose juxtacostal cells. F. Transverse section of a leaf above midleaf without stereids. A, B from Toren &amp; Dearing 9926; CF from holotype." figureDoi="http://doi.org/10.5281/zenodo.5128220" httpUri="https://zenodo.org/record/5128220/files/figure.png" pageId="0" pageNumber="105">2</figureCitation>
)
</paragraph>
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<paragraph id="B05FA603FFE2CD5EDFBBF869035DFF70" blockId="0.[146,1431,1962,2020]" lastBlockId="1.[151,1436,152,296]" lastPageId="1" lastPageNumber="106" pageId="0" pageNumber="105">
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Diagnosis:
<taxonomicName id="77E0DD80FFE2CD5FDE21F86803C5F858" authorityName="J. A. Jimenez, Toren &amp; Shevock" authorityYear="2014" box="[264,511,1998,2020]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="0" pageNumber="105" phylum="Bryophyta" rank="species" species="californicus">Didymodon californicus</taxonomicName>
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differs from the otherwise similar
<taxonomicName id="77E0DD80FFE2CD5FDC40F86807ADF858" authority="Bridel (1827: 830) Zander (1978: 25)" authorityName="Bridel (1827: 830) Zander" authorityYear="1978" box="[873,1431,1998,2020]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="0" pageNumber="105" phylum="Bryophyta" rank="species" species="vinealis">
<emphasis id="82947A11FFE2CD5FDC40F868060CF858" box="[873,1078,1998,2020]" italics="true" pageId="0" pageNumber="105">Didymodon vinealis</emphasis>
<bibRefCitation id="D471DBF2FFE2CD5FDB17F86806C7F858" author="Bridel" box="[1086,1277,1998,2020]" firstAuthor="Bridel" page="830" pageId="0" pageNumber="105" refId="ref2540" refString="Bridel, S. E. (1827) Bryologia universa seu systematica ad novam methodum dispositio, historia et descriptio omnium muscorum frondosorum huscusque cognitorum cum synonymia ex auctoribus probatissimis, vol. 2. J. A. Barth, Leipzig. http: // dx. doi. org / 10.5962 / bhl. title. 60260" type="book" year="1827">Bridel (1827: 830)</bibRefCitation>
<bibRefCitation id="D471DBF2FFE2CD5EDA2CF86802D2FF12" author="Zander, R. H." firstAuthor="Zander" lastPageId="1" lastPageNumber="106" page="25" pageId="0" pageNumber="105" pagination="11 - 32" refId="ref3011" refString="Zander, R. H. (1978) New combinations in Didymodon (Musci) and a key to the taxa in North America North of Mexico. Phytologia 41: 11 - 32." type="journal article" year="1978">Zander (1978: 25)</bibRefCitation>
</taxonomicName>
in its obtuse or widely acute leaf apex not ending in a conical cell, plane leaf margins throughout and costa ending below the apex.
</paragraph>
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<subSubSection id="F8FAF588FFE3CD5EDFBEFF700063FE94" pageId="1" pageNumber="106" type="materials_examined">
<paragraph id="B05FA603FFE3CD5EDFBEFF700063FE94" blockId="1.[151,1436,152,296]" pageId="1" pageNumber="106">
TYPE:—
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<collectingCountry id="C8F7E693FFE3CD5EDFC4FF73030BFF57" box="[237,305,213,235]" name="United States of America" pageId="1" pageNumber="106">U.S.A.</collectingCountry>
<collectingRegion id="722468E1FFE3CD5EDE11FF730399FF57" box="[312,419,213,235]" country="United States of America" name="California" pageId="1" pageNumber="106">California</collectingRegion>
:
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,
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Sierra National Forest along road 8S05 above the San Joaquin River Canyon,
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NE of Stevenson Creek
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. T8S, R24E, section 33,
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&amp;
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20632
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(
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;
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).
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<caption id="E49FF68BFFE3CD5EDFBEF83406D8F85E" ID-DOI="http://doi.org/10.5281/zenodo.5128218" ID-Zenodo-Dep="5128218" httpUri="https://zenodo.org/record/5128218/files/figure.png" pageId="1" pageNumber="106" startId="1.[151,246,1938,1960]" targetBox="[200,1381,384,1915]" targetPageId="1">
<paragraph id="B05FA603FFE3CD5EDFBEF83406D8F85E" blockId="1.[151,1437,1938,2018]" pageId="1" pageNumber="106">
<emphasis id="82947A11FFE3CD5EDFBEF8340335F814" bold="true" box="[151,271,1938,1960]" pageId="1" pageNumber="106">FIGURE 1.</emphasis>
<taxonomicName id="77E0DD80FFE3CD5EDE3FF834003EF81B" authorityName="J. A. Jimenez, Toren &amp; Shevock" authorityYear="2014" box="[278,516,1938,1959]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="1" pageNumber="106" phylum="Bryophyta" rank="species" species="californicus">
<emphasis id="82947A11FFE3CD5EDE3FF834003EF81B" box="[278,516,1938,1959]" italics="true" pageId="1" pageNumber="106">Didymodon californicus</emphasis>
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. A. Habit, dry. B, C, D, E. Leaves. F. Ventral surface of leaf apex. G. Basal laminal cells. H. Upper laminal cells. I, J, K. Transverse sections of a leaf from basal to apical part. L. Axillary hair. M. Transverse section of stem. Scale bars: A = 2 mm. BE = 400 µm. F = 60 µm. G = 40 µm. H, L = 30 µm. IK, M = 50 µm. All from holotype.
</paragraph>
</caption>
<caption id="E49FF68BFFE0CD5DDFBEF934036CF95E" ID-DOI="http://doi.org/10.5281/zenodo.5128220" ID-Zenodo-Dep="5128220" httpUri="https://zenodo.org/record/5128220/files/figure.png" pageId="2" pageNumber="107" startId="2.[151,244,1682,1704]" targetBox="[155,1436,155,1648]" targetPageId="2">
<paragraph id="B05FA603FFE0CD5DDFBEF934036CF95E" blockId="2.[150,1436,1682,1762]" pageId="2" pageNumber="107">
<emphasis id="82947A11FFE0CD5DDFBEF9340336F914" bold="true" box="[151,268,1682,1704]" pageId="2" pageNumber="107">FIGURE 2.</emphasis>
A. Habit, dry. B. Habit, wet. C. Ventral surface of leaf apex. D. Dorsal surface of leaf apex. E. Transverse section of a leaf showing bistratose juxtacostal cells. F. Transverse section of a leaf above midleaf without stereids. A, B from
<emphasis id="82947A11FFE0CD5DDB93F91607AFF979" box="[1210,1429,1712,1733]" italics="true" pageId="2" pageNumber="107">Toren &amp; Dearing 9926</emphasis>
; CF from holotype.
</paragraph>
</caption>
<subSubSection id="F8FAF588FFE0CD5CDFBEF8800750FD81" lastPageId="3" lastPageNumber="108" pageId="2" pageNumber="107" type="description">
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Plants
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high, growing in dense turfs, green or green-brown to red-brown; stems erect, irregularly branched, not radiculose, without hyalodermis, sclerodermis differentiated, central strand weakly developed; axillary hairs filiform, 34 cells long, with 1(2) brown basal cell and hyaline upper cells. Rhizoidal tubers absent. Leaves appressed to strongly incurved, seldom slightly twisted when dry, erect-patent or more rarely spreading when moist, often rather distant, lanceolate to widely-lanceolate, base sometimes ovate, 1.73.1 ×
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, channelled ventrally in the upper part, lamina unistratose, sometimes juxtacostal lamina cells bistratose in several rows, red to reddish orange with KOH; apex obtuse or widely acute, not cucullate; margins plane, unistratose, very seldom irregularly bistratose in 1(2) rows of cells in distal part, often decurrent on stem; costa strong,
<quantity id="77180BE6FFE1CD5CDA32FF1A07A6FF68" box="[1307,1436,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="3" pageNumber="108" unit="mm" value="110.0" valueMax="150.0" valueMin="70.0">70150 µm</quantity>
wide at base, ending 25 cells below the apex, sometimes sinuose above midleaf, not spurred; ventral surface cells of the costa subquadrate to oblate, smooth, vanishing near the apex, and replaced by a hyaline area of widely rectangular cells, dorsal surface cells of the costa subquadrate to oblate, smooth, in cross-section below midleaf semicircular, with 23 layers of guide cells, with 46 cells in each layer, without ventral stereids, 12 layers of dorsal stereids present, lunulate in shape, without hydroids, ventral surface cells differentiated, smooth, dorsal surface cells differentiated, smooth; upper and middle lamina cells subquadrate, rounded or oblate, 3.28(9.6) × 3.28(9.6) µm, essentially smooth, sometimes with low papillae, not bulging, thick-walled; juxtacostal basal cells quadrate to shortly rectangular, 7.530(35) ×
<quantity id="77180BE6FFE1CD5CDD88FE7A0108FE48" box="[673,818,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="12.5" metricValueMin="0.5" pageId="3" pageNumber="108" unit="mm" value="6.5" valueMax="12.5" valueMin="0.5">0.512.5 µm</quantity>
, smooth, not bulging, slightly and evenly thick-walled; marginal basal cells mostly oblate, some subquadrate, 3.29.6 ×
<quantity id="77180BE6FFE1CD5CDCA3FE590630FDA4" box="[906,1034,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" metricValueMax="1.6" metricValueMin="0.64" pageId="3" pageNumber="108" unit="mm" value="11.2" valueMax="16.0" valueMin="6.4">6.416 µm</quantity>
, smooth, not bulging, transversely thick-walled, forming a differentiated area. Gemmae absent. Perigonia, perichaetia and sporophytes unknown.
</paragraph>
</subSubSection>
<subSubSection id="F8FAF588FFE1CD5CDFEFFDE10739FC84" pageId="3" pageNumber="108" type="materials_examined">
<paragraph id="B05FA603FFE1CD5CDFEFFDE10739FC84" blockId="3.[151,1436,151,1365]" pageId="3" pageNumber="108">
<emphasis id="82947A11FFE1CD5CDFEFFDE1006BFDDC" bold="true" box="[198,593,583,608]" pageId="3" pageNumber="108">Additional specimens examined:</emphasis>
<materialsCitation id="0088AC5EFFE1CD5CDD5BFDE1064BFD14" collectingDate="1997-07-02" collectionCode="CAS, MO, MUB, NY" collectorName="Shevock" country="United States of America" county="Glenn County" elevation="1509" latitude="39.525" location="Mendocino National Forest, along forest highway M- 3, 4.9 miles southeast of Ivory Mill Saddle and 3.9 miles northeast of Sheetiron Mountain" longLatPrecision="19" longitude="-122.683334" pageId="3" pageNumber="108" specimenCount="1" stateProvince="CALIFORNIA">
<collectingCountry id="C8F7E693FFE1CD5CDD5BFDE10080FDDC" box="[626,698,583,608]" name="United States of America" pageId="3" pageNumber="108">U.S.A.</collectingCountry>
<collectingRegion id="722468E1FFE1CD5CDDECFDEA0177FDE3" box="[709,845,588,607]" country="United States of America" name="California" pageId="3" pageNumber="108">CALIFORNIA</collectingRegion>
:
<collectingCounty id="593EDE8FFFE1CD5CDC76FDE10638FDDC" box="[863,1026,583,608]" pageId="3" pageNumber="108">Glenn County</collectingCounty>
,
<location id="B53FF0D8FFE1CD5CDB39FDE107A2FD39" LSID="urn:lsid:plazi:treatment:38491715FFE2CD5CDFBBF80C0709FAE9:B53FF0D8FFE1CD5CDB39FDE107A2FD39" country="United States of America" county="Glenn County" latitude="39.525" longLatPrecision="19" longitude="-122.683334" name="Mendocino National Forest, along forest highway M- 3, 4.9 miles southeast of Ivory Mill Saddle and 3.9 miles northeast of Sheetiron Mountain" pageId="3" pageNumber="108" stateProvince="CALIFORNIA">
Mendocino National Forest, along forest highway M-3,
<quantity id="77180BE6FFE1CD5CDEA6FDCA03C1FD39" box="[399,507,620,645]" metricMagnitude="3" metricUnit="m" metricValue="7.8857856" pageId="3" pageNumber="108" unit="mi" value="4.9">4.9 miles</quantity>
southeast of Ivory Mill Saddle and
<quantity id="77180BE6FFE1CD5CDC8DFDCA062AFD39" box="[932,1040,620,645]" metricMagnitude="3" metricUnit="m" metricValue="6.2764416" pageId="3" pageNumber="108" unit="mi" value="3.9">3.9 miles</quantity>
northeast of Sheetiron Mountain
</location>
,
<geoCoordinate id="D5D4C0C4FFE1CD5CDFBEFD290320FD1B" box="[151,282,655,680]" degrees="39" direction="north" minutes="31" orientation="latitude" pageId="3" pageNumber="108" precision="15" seconds="30" value="39.525">39º31'30''N</geoCoordinate>
,
<geoCoordinate id="D5D4C0C4FFE1CD5CDE0FFD290381FD1B" box="[294,443,655,680]" degrees="122" direction="west" minutes="41" orientation="longitude" pageId="3" pageNumber="108" precision="15" seconds="00" value="-122.683334">122º41'00''W</geoCoordinate>
,
<quantity id="77180BE6FFE1CD5CDEEEFD29001AFD1B" box="[455,544,655,680]" metricMagnitude="3" metricUnit="m" metricValue="1.509" pageId="3" pageNumber="108" unit="m" value="1509.0">
<elevation id="3BCD4130FFE1CD5CDEEEFD29001AFD1B" box="[455,544,655,680]" metricMagnitude="3" metricUnit="m" metricValue="1.509" pageId="3" pageNumber="108" unit="m" value="1509.0">1509 m</elevation>
</quantity>
,
<date id="C45E80C3FFE1CD5CDD05FD29009FFD14" box="[556,677,655,680]" pageId="3" pageNumber="108" value="1997-07-02">
<collectingDate id="D41A792BFFE1CD5CDD05FD29009FFD14" box="[556,677,655,680]" pageId="3" pageNumber="108" value="1997-07-02">2 Jul 1997</collectingDate>
</date>
,
<emphasis id="82947A11FFE1CD5CDD98FD290160FD14" box="[689,858,655,680]" italics="true" pageId="3" pageNumber="108">
<collectorName id="1D15C3D5FFE1CD5CDD98FD290137FD14" box="[689,781,655,680]" pageId="3" pageNumber="108">Shevock</collectorName>
15818
</emphasis>
(
<collectionCode id="D6F13EC6FFE1CD5CDC42FD29019FFD14" box="[875,933,655,680]" pageId="3" pageNumber="108">CAS</collectionCode>
,
<collectionCode id="D6F13EC6FFE1CD5CDC98FD2901DBFD14" box="[945,993,655,680]" pageId="3" pageNumber="108">MO</collectionCode>
,
<collectionCode id="D6F13EC6FFE1CD5CDCC4FD36060AFD1B" box="[1005,1072,656,679]" pageId="3" pageNumber="108">MUB</collectionCode>
,
<collectionCode id="D6F13EC6FFE1CD5CDB15FD290653FD14" box="[1084,1129,655,680]" pageId="3" pageNumber="108">NY</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0088AC5EFFE1CD5CDB55FD2907A2FD4C" collectingDate="2012-07-12" collectionCode="CAS, MUB" collectorName="Toren &amp; Dearing" county="Lake County" elevation="1646" latitude="39.433334" location="Mendocino National Forest, east of Crockett Peak along old un-numbered logging road south of Forest Road M 3 and southwest of Waters Camp" longLatPrecision="19" longitude="-122.75694" pageId="3" pageNumber="108" specimenCount="1">
<collectingCounty id="593EDE8FFFE1CD5CDB55FD29072AFD14" box="[1148,1296,655,680]" pageId="3" pageNumber="108">Lake County</collectingCounty>
,
<location id="B53FF0D8FFE1CD5CDA35FD2903FDFD4C" LSID="urn:lsid:plazi:treatment:38491715FFE2CD5CDFBBF80C0709FAE9:B53FF0D8FFE1CD5CDA35FD2903FDFD4C" county="Lake County" latitude="39.433334" longLatPrecision="19" longitude="-122.75694" name="Mendocino National Forest, east of Crockett Peak along old un-numbered logging road south of Forest Road M 3 and southwest of Waters Camp" pageId="3" pageNumber="108">Mendocino National Forest, east of Crockett Peak along old un-numbered logging road south of Forest Road M3 and southwest of Waters Camp</location>
,
<geoCoordinate id="D5D4C0C4FFE1CD5CDEFAFD71006FFD53" box="[467,597,727,752]" degrees="39" direction="north" minutes="26" orientation="latitude" pageId="3" pageNumber="108" precision="15" seconds="00" value="39.433334">39º26'00''N</geoCoordinate>
,
<geoCoordinate id="D5D4C0C4FFE1CD5CDD48FD7100CCFD53" box="[609,758,727,752]" degrees="122" direction="west" minutes="45" orientation="longitude" pageId="3" pageNumber="108" precision="15" seconds="25" value="-122.75694">122º45'25''W</geoCoordinate>
,
<quantity id="77180BE6FFE1CD5CDC28FD710162FD53" box="[769,856,727,752]" metricMagnitude="3" metricUnit="m" metricValue="1.646" pageId="3" pageNumber="108" unit="m" value="1646.0">
<elevation id="3BCD4130FFE1CD5CDC28FD710162FD53" box="[769,856,727,752]" metricMagnitude="3" metricUnit="m" metricValue="1.646" pageId="3" pageNumber="108" unit="m" value="1646.0">1646 m</elevation>
</quantity>
,
<date id="C45E80C3FFE1CD5CDC4DFD7101D1FD4C" box="[868,1003,727,752]" pageId="3" pageNumber="108" value="2012-07-12">
<collectingDate id="D41A792BFFE1CD5CDC4DFD7101D1FD4C" box="[868,1003,727,752]" pageId="3" pageNumber="108" value="2012-07-12">12 Jul 2012</collectingDate>
</date>
,
<emphasis id="82947A11FFE1CD5CDCDFFD7F06CCFD4C" box="[1014,1270,727,752]" italics="true" pageId="3" pageNumber="108">
<collectorName id="1D15C3D5FFE1CD5CDCDFFD7F060FFD4C" box="[1014,1077,729,752]" pageId="3" pageNumber="108">Toren</collectorName>
&amp;
<collectorName id="1D15C3D5FFE1CD5CDB73FD7E068DFD53" box="[1114,1207,728,751]" pageId="3" pageNumber="108">Dearing</collectorName>
9926
</emphasis>
(
<collectionCode id="D6F13EC6FFE1CD5CDA2FFD71077AFD4C" box="[1286,1344,727,752]" pageId="3" pageNumber="108">CAS</collectionCode>
,
<collectionCode id="D6F13EC6FFE1CD5CDA65FD7107AAFD4C" box="[1356,1424,727,752]" pageId="3" pageNumber="108">MUB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0088AC5EFFE1CD5CDFBEFD5A06C5FC84" collectingDate="2000-07-29" collectionCode="CAS, MUB" collectorName="Shevock &amp; Norris" county="Madera County" elevation="1372" latitude="37.275" location="Sierra National Forest along Minarets Road, 1.8 miles north of Fish Creek and 1.5 miles south of Slide Creek" longLatPrecision="19" longitude="-119.35833" pageId="3" pageNumber="108" specimenCount="1">
<collectingCounty id="593EDE8FFFE1CD5CDFBEFD5A0372FCA9" box="[151,328,764,789]" pageId="3" pageNumber="108">Madera County</collectingCounty>
,
<location id="B53FF0D8FFE1CD5CDE7AFD5A0327FC84" LSID="urn:lsid:plazi:treatment:38491715FFE2CD5CDFBBF80C0709FAE9:B53FF0D8FFE1CD5CDE7AFD5A0327FC84" county="Madera County" latitude="37.275" longLatPrecision="19" longitude="-119.35833" name="Sierra National Forest along Minarets Road, 1.8 miles north of Fish Creek and 1.5 miles south of Slide Creek" pageId="3" pageNumber="108">
Sierra National Forest along Minarets Road,
<quantity id="77180BE6FFE1CD5CDC67FD5A018FFCA9" box="[846,949,764,789]" metricMagnitude="3" metricUnit="m" metricValue="2.8968192" pageId="3" pageNumber="108" unit="mi" value="1.8">1.8 miles</quantity>
north of Fish Creek and
<quantity id="77180BE6FFE1CD5CDBF8FD5A0703FCA9" box="[1233,1337,764,789]" metricMagnitude="3" metricUnit="m" metricValue="2.414016" pageId="3" pageNumber="108" unit="mi" value="1.5">1.5 miles</quantity>
south of Slide Creek
</location>
,
<geoCoordinate id="D5D4C0C4FFE1CD5CDE01FCB90390FC84" box="[296,426,799,824]" degrees="37" direction="north" minutes="16" orientation="latitude" pageId="3" pageNumber="108" precision="15" seconds="30" value="37.275">37º16'30''N</geoCoordinate>
,
<geoCoordinate id="D5D4C0C4FFE1CD5CDE9FFCB90070FC84" box="[438,586,799,824]" degrees="119" direction="west" minutes="21" orientation="longitude" pageId="3" pageNumber="108" precision="15" seconds="30" value="-119.35833">119º21'30''W</geoCoordinate>
,
<quantity id="77180BE6FFE1CD5CDD7CFCB90096FC8B" box="[597,684,799,824]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="3" pageNumber="108" unit="m" value="1372.0">
<elevation id="3BCD4130FFE1CD5CDD7CFCB90096FC8B" box="[597,684,799,824]" metricMagnitude="3" metricUnit="m" metricValue="1.3719999999999999" pageId="3" pageNumber="108" unit="m" value="1372.0">1372 m</elevation>
</quantity>
,
<date id="C45E80C3FFE1CD5CDD91FCB90107FC84" box="[696,829,799,824]" pageId="3" pageNumber="108" value="2000-07-29">
<collectingDate id="D41A792BFFE1CD5CDD91FCB90107FC84" box="[696,829,799,824]" pageId="3" pageNumber="108" value="2000-07-29">29 Jul 2000</collectingDate>
</date>
,
<emphasis id="82947A11FFE1CD5CDC61FCB90664FC84" box="[840,1118,799,824]" italics="true" pageId="3" pageNumber="108">
<collectorName id="1D15C3D5FFE1CD5CDC61FCB9019EFC84" box="[840,932,799,824]" pageId="3" pageNumber="108">Shevock</collectorName>
&amp;
<collectorName id="1D15C3D5FFE1CD5CDCE1FC86062BFC8B" box="[968,1041,800,823]" pageId="3" pageNumber="108">Norris</collectorName>
19786
</emphasis>
(
<collectionCode id="D6F13EC6FFE1CD5CDB44FCB9069DFC84" box="[1133,1191,799,824]" pageId="3" pageNumber="108">CAS</collectionCode>
,
<collectionCode id="D6F13EC6FFE1CD5CDB9AFCB906CDFC84" box="[1203,1271,799,824]" pageId="3" pageNumber="108">MUB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F8FAF588FFE1CD5CDFEFFCE2035AFC3C" pageId="3" pageNumber="108" type="etymology">
<paragraph id="B05FA603FFE1CD5CDFEFFCE2035AFC3C" blockId="3.[151,1436,151,1365]" pageId="3" pageNumber="108">
<emphasis id="82947A11FFE1CD5CDFEFFCE2036BFCE1" bold="true" box="[198,337,836,861]" pageId="3" pageNumber="108">Etymology:</emphasis>
—The epithet
<taxonomicName id="77E0DD80FFE1CD5CDED2FCE200BBFCE1" authorityName="J. A. Jimenez, Toren &amp; Shevock" authorityYear="2014" box="[507,641,836,861]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="108" phylum="Bryophyta" rank="species" species="californicus">
<emphasis id="82947A11FFE1CD5CDED2FCE200BBFCE1" box="[507,641,836,861]" italics="true" pageId="3" pageNumber="108">californicus</emphasis>
</taxonomicName>
refers to the State of
<collectingRegion id="722468E1FFE1CD5CDCA2FCE201C7FCE1" box="[907,1021,836,861]" country="United States of America" name="California" pageId="3" pageNumber="108">California</collectingRegion>
where this species appears to be a regional endemic.
</paragraph>
</subSubSection>
<subSubSection id="F8FAF588FFE1CD5CDFEFFC2A0709FAE9" pageId="3" pageNumber="108" type="distribution">
<paragraph id="B05FA603FFE1CD5CDFEFFC2A0151FAB1" blockId="3.[151,1436,151,1365]" pageId="3" pageNumber="108">
<emphasis id="82947A11FFE1CD5CDFEFFC2A03CDFC19" bold="true" box="[198,503,908,933]" pageId="3" pageNumber="108">Habitat and distribution:</emphasis>
<taxonomicName id="77E0DD80FFE1CD5CDD3AFC2A011EFC19" authorityName="J. A. Jimenez, Toren &amp; Shevock" authorityYear="2014" box="[531,804,908,933]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="108" phylum="Bryophyta" rank="species" species="californicus">
<emphasis id="82947A11FFE1CD5CDD3AFC2A011EFC19" box="[531,804,908,933]" italics="true" pageId="3" pageNumber="108">Didymodon californicus</emphasis>
</taxonomicName>
, based on the known occurrences, is viewed as a semirheophytic species where it is seasonally submerged or inundated during snowmelt and peak rain events along perennial and intermittent streams. However, during most of the year the plants are dry. This ecological partition of habitat along the high water zone of stream and river banks is also similar to that displayed by
<taxonomicName id="77E0DD80FFE1CD5CDA3AFC5100F4FB89" authority="De Notaris (1838: 320) Hill (1981: 599)" authorityName="De Notaris (1838: 320) Hill" authorityYear="1981" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="108" phylum="Bryophyta" rank="species" species="insulanus">
<emphasis id="82947A11FFE1CD5CDA3AFC510339FB89" italics="true" pageId="3" pageNumber="108">Didymodon insulanus</emphasis>
<bibRefCitation id="D471DBF2FFE1CD5CDE20FBBA002AFB89" author="De Notaris" box="[265,528,1052,1077]" firstAuthor="De Notaris" page="320" pageId="3" pageNumber="108" pagination="281 - 326" refId="ref2623" refString="De Notaris, G. (1838) Specimen de Tortulis italicis. Memorie della Reale Accademia delle Scienze di Torino 40: 281 - 326." type="journal article" year="1838">De Notaris (1838: 320)</bibRefCitation>
<bibRefCitation id="D471DBF2FFE1CD5CDD3EFBBA00F4FB89" author="Hill" box="[535,718,1052,1077]" firstAuthor="Hill" page="599" pageId="3" pageNumber="108" pagination="599 - 602" refId="ref2779" refString="Hill, M. O. (1981) [1982] New combinations in European mosses. I. Pottiaceae. Journal of Bryology 11: 599 - 602. http: // dx. doi. org / 10.1179 / jbr. 1981.11.4.599" type="journal article" year="1981">Hill (1981: 599)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="77E0DD80FFE1CD5CDC2FFBBB0698FB89" authority="Culmann (1907: 100)" authorityName="Culmann" authorityPageNumber="100" authorityYear="1907" box="[774,1186,1052,1077]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="108" phylum="Bryophyta" rank="species" species="nicholsonii">
<emphasis id="82947A11FFE1CD5CDC2FFBBB019FFB89" box="[774,933,1052,1077]" italics="true" pageId="3" pageNumber="108">D. nicholsonii</emphasis>
<bibRefCitation id="D471DBF2FFE1CD5CDC85FBBA0698FB89" author="Culmann" box="[940,1186,1052,1077]" firstAuthor="Culmann" page="100" pageId="3" pageNumber="108" pagination="100 - 102" refId="ref2600" refString="Culmann, P. (1907) Barbula aut Didymodon nicholsonii spec. nov. Revue Bryologique 34: 100 - 102." type="journal article" year="1907">Culmann (1907: 100)</bibRefCitation>
</taxonomicName>
. Both of these species are frequently encountered along streams and rivers where they too are seasonally submerged. Even some
<collectingRegion id="722468E1FFE1CD5CDFBEFBC20331FBC1" box="[151,267,1124,1149]" country="United States of America" name="California" pageId="3" pageNumber="108">California</collectingRegion>
populations of
<taxonomicName id="77E0DD80FFE1CD5CDEEBFBC20092FBC1" authorityName="Bridel (1827: 830) Zander" authorityYear="1978" box="[450,680,1124,1149]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="108" phylum="Bryophyta" rank="species" species="vinealis">
<emphasis id="82947A11FFE1CD5CDEEBFBC20092FBC1" box="[450,680,1124,1149]" italics="true" pageId="3" pageNumber="108">Didymodon vinealis</emphasis>
</taxonomicName>
can be viewed ecologically as being rheophytic where the plants
<taxonomicName id="77E0DD80FFE1CD5CDFBEFB2102C6FB1C" box="[151,252,1159,1184]" form="flat" pageId="3" pageNumber="108" rank="form">form flat</taxonomicName>
mat-like colonies, over rock slabs with seasonal water flow, more reminiscent of the growth pattern of
<taxonomicName id="77E0DD80FFE1CD5CDFBEFB0A03D9FB79" authorityName="B. Bremer" authorityYear="1980" baseAuthorityName="C.Mull." box="[151,483,1196,1221]" class="Bryopsida" family="Grimmiaceae" genus="Schistidium" kingdom="Plantae" order="Grimmiales" pageId="3" pageNumber="108" phylum="Bryophyta" rank="species" species="cinclidodonteum">
<emphasis id="82947A11FFE1CD5CDFBEFB0A03D9FB79" box="[151,483,1196,1221]" italics="true" pageId="3" pageNumber="108">Schistidium cinclidodonteum</emphasis>
</taxonomicName>
(Müll. Hal. in
<bibRefCitation id="D471DBF2FFE1CD5CDDBAFB0A0178FB79" author="Roll" box="[659,834,1196,1221]" firstAuthor="Roll" page="387" pageId="3" pageNumber="108" pagination="417 - 424" refId="ref2870" refString="Roll, J. (1890) Vorlaufige Mittheilungen uber die von in Jahr 1888 in Nord-Amerika Gesammelten Neuen Arten und Variataten der Laubmoose. Botanisches Centralblatt 44: 385 - 391, 417 - 424." type="journal article" year="1890">Röll 1890: 387</bibRefCitation>
) Bremer (108: 1980) populations. This may explain why this new species of
<taxonomicName id="77E0DD80FFE1CD5CDE86FB690009FB54" authorityName="Hedwig" authorityYear="1801" box="[431,563,1231,1256]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="108" phylum="Bryophyta" rank="genus">
<emphasis id="82947A11FFE1CD5CDE86FB690009FB54" box="[431,563,1231,1256]" italics="true" pageId="3" pageNumber="108">Didymodon</emphasis>
</taxonomicName>
has been overlooked and under-collected. Many bryologists defer collecting these common species, especially when sporophytes are lacking.
</paragraph>
<paragraph id="B05FA603FFE1CD5CDFEFFAB10709FAE9" blockId="3.[151,1436,151,1365]" pageId="3" pageNumber="108">
<taxonomicName id="77E0DD80FFE1CD5CDFEFFAB103ECFA8C" authorityName="J. A. Jimenez, Toren &amp; Shevock" authorityYear="2014" box="[198,470,1303,1328]" class="Bryopsida" family="Pottiaceae" genus="Didymodon" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="108" phylum="Bryophyta" rank="species" species="californicus">
<emphasis id="82947A11FFE1CD5CDFEFFAB103ECFA8C" box="[198,470,1303,1328]" italics="true" pageId="3" pageNumber="108">Didymodon californicus</emphasis>
</taxonomicName>
occurs in the central Sierra
<collectingRegion id="722468E1FFE1CD5CDC3BFAB1015DFA8C" box="[786,871,1303,1328]" country="United States of America" name="Nevada" pageId="3" pageNumber="108">Nevada</collectingRegion>
and the northern Coast Ranges of
<collectingRegion id="722468E1FFE1CD5CDBC4FAB10765FA8C" box="[1261,1375,1303,1328]" country="United States of America" name="California" pageId="3" pageNumber="108">California</collectingRegion>
(
<figureCitation id="28DBBA86FFE1CD5CDA47FAB1029DFAE9" captionStart="FIGURE 3" captionStartId="4.[151,244,1549,1571]" captionTargetBox="[157,1436,159,1538]" captionTargetId="figure-238@4.[156,1441,157,1539]" captionTargetPageId="4" captionText="FIGURE 3. Map of California showing the known distribution of Didymodon californicus." figureDoi="http://doi.org/10.5281/zenodo.5128222" httpUri="https://zenodo.org/record/5128222/files/figure.png" pageId="3" pageNumber="108">Fig. 3</figureCitation>
). All known occurrences are on public land providing various opportunities for long-term conservation.
</paragraph>
</subSubSection>
</treatment>
</document>