Add updates up until 2025-03-05 16:19:00

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<emphasis id="7DDDB356FFADFFE2FECFFBB3FEC9FBC3" bold="true" box="[269,373,1114,1141]" italics="true" pageId="14" pageNumber="15">Belovius</emphasis>
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<emphasis id="7DDDB356FFADFFE2FE45FBB3FE40FBC2" bold="true" box="[391,508,1114,1140]" italics="true" pageId="14" pageNumber="15">L-nigrum</emphasis>
<bibRefCitation id="2B3812B5FFADFFE2FDC1FBB3FD5DFBC2" author="Matsumura, S." box="[515,737,1114,1140]" pageId="14" pageNumber="15" refId="ref30492" refString="Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Toko-shoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura, 1931</bibRefCitation>
(
<figureCitation id="D79273C1FFADFFE2FD32FBB3FCC4FBC2" box="[752,888,1114,1140]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStart-4="FIGURE 14" captionStartId-0="16.[151,250,1659,1683]" captionStartId-1="17.[151,250,945,969]" captionStartId-2="18.[151,250,1339,1363]" captionStartId-3="19.[151,250,1448,1472]" captionStartId-4="20.[151,250,1084,1108]" captionTargetBox-0="[154,1433,187,1633]" captionTargetBox-1="[154,1433,310,918]" captionTargetBox-2="[182,1410,186,1309]" captionTargetBox-3="[151,1431,345,1425]" captionTargetBox-4="[151,1432,273,1058]" captionTargetId-0="figure-20@16.[151,1435,181,1636]" captionTargetId-1="figure-90@17.[151,1435,309,921]" captionTargetId-2="figure-20@18.[173,1415,181,1315]" captionTargetId-3="figure-106@19.[151,1435,345,1425]" captionTargetId-4="figure-72@20.[151,1435,273,1061]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionText-0="FIGURE 10af. Epeorus L-nigrum, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." captionText-1="FIGURE 11ad. Epeorus L-nigrum, mature nymph. a. Male head, dorsal view; b. Female head, dorsal view; c. Right middle leg, anterior face; d. Gills IVII (from right to left), dorsal view. Scales: ad 1mm." captionText-2="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." captionText-3="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." captionText-4="FIGURE 14ae. Epeorus L-nigrum, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi-0="http://doi.org/10.5281/zenodo.14954007" figureDoi-1="http://doi.org/10.5281/zenodo.14954009" figureDoi-2="http://doi.org/10.5281/zenodo.14954011" figureDoi-3="http://doi.org/10.5281/zenodo.14954013" figureDoi-4="http://doi.org/10.5281/zenodo.14954015" httpUri-0="https://zenodo.org/record/14954007/files/figure.png" httpUri-1="https://zenodo.org/record/14954009/files/figure.png" httpUri-2="https://zenodo.org/record/14954011/files/figure.png" httpUri-3="https://zenodo.org/record/14954013/files/figure.png" httpUri-4="https://zenodo.org/record/14954015/files/figure.png" pageId="14" pageNumber="15">Figs 1014</figureCitation>
)
</emphasis>
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[
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name &quot;Matsumura-hirata-kagerou&quot;]
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L-nigrum
</emphasis>
:
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: p. 1477 (male imago) [
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<emphasis id="7DDDB356FFADFFE2FBF7FB2AFBD5FB6C" box="[1077,1129,1218,1242]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2018: p. 126, pl. 15, figs 14, figs 46-146-4, 47-10 (male imago, male subimago, nymph) [
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: p. 33, pl. 22, fig. 4 (egg) [
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</paragraph>
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<taxonomicName id="88A914C7FFADFFE2FF55FAF6FEFDFA80" ID-CoL="6FNWK" authorityName="Matsumura" authorityYear="1931" box="[151,321,1310,1334]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="l-nigrus">
<emphasis id="7DDDB356FFADFFE2FF55FAF6FEFDFA80" box="[151,321,1310,1334]" italics="true" pageId="14" pageNumber="15">Epeorus l-nigrus</emphasis>
</taxonomicName>
:
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: p. 186, fig. 10FJ (male imago) [
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<collectingRegion id="8D6DA1A6FFADFFE2FC56FAF7FC6AFA80" box="[916,982,1310,1334]" country="Japan" name="Chiba" pageId="14" pageNumber="15">Chiba</collectingRegion>
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<bibRefCitation id="2B3812B5FFADFFE2FC27FAF7FBCBFA81" author="Ishiwata, S." box="[997,1143,1310,1335]" pageId="14" pageNumber="15" pagination="1 - 46" refId="ref29256" refString="Ishiwata, S. (2002) Mayflies of Kanagawa Prefecture, Japan. Kanagawa-Chuho, 138, 1-46. [in Japanese]" type="journal article" year="2002">Ishiwata 2002</bibRefCitation>
: p. 23 (male imago) [
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:
<collectingRegion id="8D6DA1A6FFADFFE2FF05FAD7FE84FAE0" box="[199,312,1342,1366]" country="Japan" name="Kanagawa" pageId="14" pageNumber="15">Kanagawa</collectingRegion>
];
<bibRefCitation id="2B3812B5FFADFFE2FE8BFAD4FDD7FAE3" author="Ishiwata, S. &amp; Takemon, Y." box="[329,619,1341,1365]" pageId="14" pageNumber="15" pagination="31 - 128" refId="ref29453" refString="Ishiwata, S. &amp; Takemon, Y. (2005 b) Ephemeroptera. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic insects of Japan: manual with keys and illustration. Tokai University Press, Kanagawa, pp. 31-128. [in Japanese]" type="book chapter" year="2005">Ishiwata &amp; Takemon 2005b</bibRefCitation>
: p. 105, figs 46-146-4, 47-10 (male imago) [
<collectingCountry id="37BE2FD4FFADFFE2FB86FAD4FB25FAE3" box="[1092,1177,1341,1365]" name="Japan" pageId="14" pageNumber="15">JAPAN</collectingCountry>
];
<bibRefCitation id="2B3812B5FFADFFE2FB69FAD4FA9EFAE3" author="Inada, K." box="[1195,1314,1341,1365]" pageId="14" pageNumber="15" pagination="111 - 112" refId="ref29089" refString="Inada, K. (2007) Distribution of Epeorus l-nigrus and E. latifolium at the Ibokawa River, Hyogo, Japan. Hyogo Freshwater Biology, 59, 111-112. [in Japanese]" type="journal article" year="2007">Inada 2007</bibRefCitation>
: p. 111, fig. 1 (male imago) [
<collectingCountry id="37BE2FD4FFADFFE2FEB3FAB5FE78FAC2" box="[369,452,1372,1396]" name="Japan" pageId="14" pageNumber="15">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFADFFE2FE0CFAB5FDA4FAC2" box="[462,536,1372,1396]" country="Japan" name="Hyogo" pageId="14" pageNumber="15">Hyogo</collectingRegion>
];
<bibRefCitation id="2B3812B5FFADFFE2FDE8FAB5FD6AFAC2" author="Maruyama, H." box="[554,726,1372,1396]" pageId="14" pageNumber="15" pagination="90 - 178" refId="ref30430" refString="Maruyama, H. (2016) Ephemeroptera. In: Maruyama, H. &amp; Hanada, S. (Eds.), A Field Guide to Japanese Aquatic Insects: Adults of Mayflies, Stoneflies and Caddisflies. Zenkoku Noson Kyoiku Kyokai Co., Ltd, Tokyo, pp. 90-178. [in Japanese]" type="book chapter" year="2016">Maruyama 2016</bibRefCitation>
: p. 173, figs
<quantity id="8851C2A1FFADFFE2FC99FAB5FC18FAC2" box="[859,932,1372,1396]" metricMagnitude="-2" metricUnit="kg" metricValue="8.6" metricValueMax="17.1" metricValueMin="0.1" pageId="14" pageNumber="15" unit="g" value="86.0" valueMax="171.0" valueMin="1.0">1-171g</quantity>
, 1-178, 1-275, 1-276, 1-277, 1-278 (male imago, female imago, male subimago, female subimago) [
<collectingCountry id="37BE2FD4FFADFFE2FD05FA92FCA6FA25" box="[711,794,1403,1427]" name="Japan" pageId="14" pageNumber="15">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFADFFE2FCE6FA92FCE6FA24" box="[804,858,1403,1426]" country="Japan" name="Nara" pageId="14" pageNumber="15">Nara</collectingRegion>
], mis-spelling.
</paragraph>
<paragraph id="4F166F44FFADFFE2FF55FA73FCC3FA58" blockId="14.[151,1437,1218,1519]" pageId="14" pageNumber="15">
<taxonomicName id="88A914C7FFADFFE2FF55FA73FEE8FA07" ID-CoL="6FNWP" box="[151,340,1433,1457]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFADFFE2FF55FA73FEE8FA07" box="[151,340,1433,1457]" italics="true" pageId="14" pageNumber="15">Epeorus latifolium</emphasis>
</taxonomicName>
(non
<taxonomicName id="88A914C7FFADFFE2FE4EFA73FD7AFA07" ID-CoL="6FNWP" authority="Ueno, 1928" authorityName="Ueno" authorityYear="1928" box="[396,710,1433,1457]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFADFFE2FE4EFA73FDF4FA07" box="[396,584,1433,1457]" italics="true" pageId="14" pageNumber="15">Epeorus latifolium</emphasis>
<bibRefCitation id="2B3812B5FFADFFE2FD8FFA70FD7AFA07" author="Ueno, M." box="[589,710,1433,1457]" pageId="14" pageNumber="15" pagination="19 - 63" refId="ref31987" refString="Ueno, M. (1928) Some Japanese mayfly nymphs. Memoirs of the college of science, Kyoto Imperial University, Series B, 4 (1), 19-63." type="journal article" year="1928">Uéno, 1928</bibRefCitation>
</taxonomicName>
):
<bibRefCitation id="2B3812B5FFADFFE2FD15FA73FCCBFA07" author="Horasawa, I." box="[727,887,1433,1457]" pageId="14" pageNumber="15" pagination="253 - 258" refId="ref28316" refString="Horasawa, I. (1929) The description of Epeorus latifolium. Kontyu, 3, 253-258. [in Japanese] *" type="journal article" year="1929">Horasawa 1929</bibRefCitation>
: p. 253, figs 13 (male imago, male subimago, nymph) [
<collectingCountry id="37BE2FD4FFADFFE2FF0FFA51FE9CFA66" box="[205,288,1464,1488]" name="Japan" pageId="14" pageNumber="15">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFADFFE2FEEEFA51FE3CFA66" box="[300,384,1464,1488]" country="Japan" name="Nagano" pageId="14" pageNumber="15">Nagano</collectingRegion>
], misidentification (
<bibRefCitation id="2B3812B5FFADFFE2FD8DFA51FD4EFA66" author="Ishiwata, S." box="[591,754,1464,1488]" pageId="14" pageNumber="15" pagination="55 - 84" refId="ref29157" refString="Ishiwata, S. (2001 a) A checklist of Japanese Ephemeroptera. In: The 21 st Century and Aquatic Entomology in East Asia. roceedings of the 1 st. Symposium of Aquatic Entomologists in East Asia (AESEA). Korean Society of Aquatic Entomology Korea, Seoul, pp. 55-84." type="book chapter" year="2001">Ishiwata 2001a</bibRefCitation>
);
<bibRefCitation id="2B3812B5FFADFFE2FCC4FA51FC05FA66" author="Matsumura, S." box="[774,953,1464,1488]" pageId="14" pageNumber="15" refId="ref30550" refString="Matsumura, S. (1933) Illustrated Common Insect of Japan V. Shunyodo, Tokyo, 100 pp. [in Japanese with English description.]" type="book" year="1933">Matsumura 1933</bibRefCitation>
: p. 96, pl. 21, fig. 8 (female imago) [
<collectingCountry id="37BE2FD4FFADFFE2FA84FA51FA25FA66" box="[1350,1433,1464,1488]" name="Japan" pageId="14" pageNumber="15">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFADFFE2FF05FA3FFE8CFA58" box="[199,304,1494,1518]" country="Japan" name="Hokkaido" pageId="14" pageNumber="15">Hokkaido</collectingRegion>
], misidentification;
<bibRefCitation id="2B3812B5FFADFFE2FE3BFA3FFD32FA59" author="Imanishi, K." box="[505,654,1494,1519]" pageId="14" pageNumber="15" pagination="380 - 394" refId="ref28952" refString="Imanishi, K. (1934) Mayflies from Japanese Torrents IV. Notes on the genus Epeorus. Annotationes Zoologicae Japonense, 14, 380-394." type="journal article" year="1934">Imanishi 1934</bibRefCitation>
: p. 389, mis-synonymy.
</paragraph>
</subSubSection>
<subSubSection id="07B33CCFFFADFFE3FF55F9F5FF6BF9C3" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="4F166F44FFADFFE3FF55F9F5FE6EFBA7" blockId="14.[151,1437,1564,2024]" lastBlockId="15.[151,1437,150,1977]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="7DDDB356FFADFFE2FF55F9F5FD69F980" bold="true" box="[151,725,1564,1591]" pageId="14" pageNumber="15">
Materials examined. Type specimens:
<typeStatus id="9012D1E6FFADFFE2FDA2F9F5FD6CF980" box="[608,720,1564,1590]" pageId="14" pageNumber="15" type="syntype">Syntypes</typeStatus>
:
</emphasis>
(an identification label by Imanishi, 1933, &quot;
<taxonomicName id="88A914C7FFADFFE2FB08F9F7FEAFF9ED" authority="Ueno, Syn." authorityName="Ueno, Syn." class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFADFFE2FB08F9F7FA20F981" box="[1226,1436,1565,1591]" italics="true" pageId="14" pageNumber="15">Epeorus latifolium</emphasis>
Uéno, Syn.
</taxonomicName>
<emphasis id="7DDDB356FFADFFE2FEDBF9ABFE5AF9ED" box="[281,486,1602,1627]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="88A914C7FFADFFE2FEDBF9ABFECAF9ED" authorityName="Eaton" authorityYear="1881" box="[281,374,1602,1627]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Epeorus</taxonomicName>
L-nigrum
</emphasis>
Matsumura, Det. K. Imanishi 1933&quot;).
<specimenCount id="59AFA4CDFFADFFE2FC4DF9A8FC64F9ED" box="[911,984,1601,1627]" count="1" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
imago, &quot;
<date id="3B174984FFADFFE2FBF9F9A8FB75F9ED" box="[1083,1225,1601,1627]" pageId="14" pageNumber="15" value="1903-08-09">9 /VIII 1903</date>
, Ishiyama&quot;;
<specimenCount id="59AFA4CDFFADFFE2FA90F9A8FA20F9ED" box="[1362,1436,1601,1627]" count="1" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
imago, &quot;Sapporo, Matsumura;
<date id="3B174984FFADFFE2FE31F98CFDC0F936" box="[499,636,1637,1664]" pageId="14" pageNumber="15" value="1915-07-06">6 /VII 1915</date>
&quot;, &quot;
<emphasis id="7DDDB356FFADFFE2FD5FF98FFCD0F9C9" box="[669,876,1638,1663]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="88A914C7FFADFFE2FD5FF98FFD46F9C9" authorityName="Eaton" authorityYear="1881" box="[669,762,1638,1663]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Epeorus</taxonomicName>
L-nigrum
</emphasis>
M. det. Matsumura&quot;;
<specimenCount id="59AFA4CDFFADFFE2FBAAF98CFB08F9C9" box="[1128,1204,1637,1663]" count="1" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
imago and
<specimenCount id="59AFA4CDFFADFFE2FAF8F98CFA20F9C9" box="[1338,1436,1637,1663]" count="1" pageId="14" pageNumber="15" type="female">1 female</specimenCount>
imago, &quot;Sapporo, Matsumura;
<date id="3B174984FFADFFE2FE30F960FD36F912" box="[498,650,1673,1700]" pageId="14" pageNumber="15" value="1915-07-28">28 /VII 1915</date>
&quot;;
<specimenCount id="59AFA4CDFFADFFE2FD5CF960FD56F915" box="[670,746,1673,1699]" count="1" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
imago (
<figureCitation id="D79273C1FFADFFE2FC8AF960FC20F915" box="[840,924,1673,1699]" captionStart="FIGURE 33" captionStartId="41.[151,250,1657,1681]" captionTargetBox="[154,1434,384,1633]" captionTargetId="figure-126@41.[151,1435,381,1633]" captionTargetPageId="41" captionText="FIGURE 33af. Type specimens of Epeorus L-nigrum. a. Male imago from syntypes of E. L-nigrum, lateral view; b. Ditto, dorsal view; c. Ditto, ventral view; d. Genitalia of the specimen, ventral view; e, f. Labels attached to the specimen. Scales: ac 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14954055" httpUri="https://zenodo.org/record/14954055/files/figure.png" pageId="14" pageNumber="15">Fig. 33</figureCitation>
), &quot;Sapporo, Matsumura;
<date id="3B174984FFADFFE2FB75F960FAF8F915" box="[1207,1348,1673,1699]" pageId="14" pageNumber="15" value="1921-09-28">28 /IX 1921</date>
&quot;, &quot;Type Matsumura;
<emphasis id="7DDDB356FFADFFE2FEE6F947FE55F971" box="[292,489,1709,1735]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="88A914C7FFADFFE2FEE6F947FE3DF971" authorityName="Eaton" authorityYear="1881" box="[292,385,1710,1735]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Epeorus</taxonomicName>
l-nigrum
</emphasis>
M&quot; (red label);
<specimenCount id="59AFA4CDFFADFFE2FD55F944FD5DF971" box="[663,737,1709,1735]" count="1" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
imago, &quot;Sapporo, Matsumura;
<emphasis id="7DDDB356FFADFFE2FBF9F944FBCFF971" bold="true" box="[1083,1139,1709,1735]" pageId="14" pageNumber="15">ĭDz</emphasis>
(means &quot;garden light&quot;),
<date id="3B174984FFADFFE2FA42F944FEB4F95D" pageId="14" pageNumber="15" value="1931-09-10">10 /IX. 1931</date>
&quot;;
<specimenCount id="59AFA4CDFFADFFE2FED9F938FEDAF95D" box="[283,358,1745,1771]" count="2" pageId="14" pageNumber="15" type="male">2 male</specimenCount>
imagines, &quot;Maruyama [in Kanji], 6/7 (year not listed)&quot;;
<specimenCount id="59AFA4CDFFADFFE2FC1AF938FB9EF95D" box="[984,1058,1745,1771]" count="1" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
imago, &quot;Sapporo, Matsumura; 25 /
<collectionCode id="29B8F781FFADFFE2FF62F91CFF01F8B9" box="[160,189,1781,1807]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="14" pageNumber="15">VI</collectionCode>
??8 (year unreadable)&quot;;
<specimenCount id="59AFA4CDFFADFFE2FE04F91CFDB2F8B9" box="[454,526,1781,1807]" count="1" pageId="14" pageNumber="15" type="male">1 male</specimenCount>
imago, &quot;Sapproro, Matsumura; 21 /?? 20 (unreadable)&quot;;
<specimenCount id="59AFA4CDFFADFFE2FBBAF91CFB6BF8B9" box="[1144,1239,1781,1807]" count="1" pageId="14" pageNumber="15" type="female">1 female</specimenCount>
imago, &quot;Sapporo, Matsumura&quot; (collection date not listed).
<emphasis id="7DDDB356FFADFFE2FDAAF8F1FC87F884" bold="true" box="[616,827,1816,1842]" pageId="14" pageNumber="15">Other specimens:</emphasis>
<collectingCountry id="37BE2FD4FFADFFE2FC87F8F0FC21F885" box="[837,925,1817,1843]" name="Japan" pageId="14" pageNumber="15">JAPAN</collectingCountry>
,
<collectingRegion id="8D6DA1A6FFADFFE2FC68F8F0FBF8F885" box="[938,1092,1817,1843]" country="Japan" name="Hokkaido" pageId="14" pageNumber="15">HOKKAIDO</collectingRegion>
, ISHIKARI:
<emphasis id="7DDDB356FFADFFE2FB1FF8F0FAB8F885" bold="true" box="[1245,1284,1817,1843]" pageId="14" pageNumber="15">[3.]</emphasis>
Sapporo-shi, Minami-ku, Jozankei, Toyohira-gawa Riv., upstream of confluence of mainstream and Ekiteisawa-gawa stream,
<quantity id="8851C2A1FFADFFE2FAB0F8D4FF11F8CD" metricMagnitude="2" metricUnit="m" metricValue="5.14" pageId="14" pageNumber="15" unit="m" value="514.0">514 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFADFFE2FF37F888FE2DF8CA" box="[245,401,1889,1916]" degrees="42" direction="north" minutes="51" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="18.1" value="42.855026">
42°51'18.1&quot;
<collectingRegion id="8D6DA1A6FFADFFE2FEB8F888FE2DF8CA" box="[378,401,1889,1916]" country="Japan" name="N" pageId="14" pageNumber="15">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFADFFE2FE5EF888FDFFF8CA" box="[412,579,1889,1916]" degrees="141" direction="east" minutes="08" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="59.0" value="141.14972">141°08'59.0&quot;E</geoCoordinate>
), 1 mature female nymph,
<date id="3B174984FFADFFE2FCA8F888FC5FF8CD" box="[874,995,1889,1915]" pageId="14" pageNumber="15" value="2003-10-18">18.X.2003</date>
, K. Saito;
<emphasis id="7DDDB356FFADFFE2FB94F888FBC1F8CD" bold="true" box="[1110,1149,1889,1915]" pageId="14" pageNumber="15">[4.]</emphasis>
Sapporo-shi, Minami-ku, Jozankei, Usubetsu-gawa Stream (a tributary of Toyohira-gawa Riv.),
<quantity id="8851C2A1FFADFFE2FC61F86CFC56F829" box="[931,1002,1925,1951]" metricMagnitude="2" metricUnit="m" metricValue="4.72" pageId="14" pageNumber="15" unit="m" value="472.0">472 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFADFFE2FBF1F86CFB73F816" box="[1075,1231,1925,1952]" degrees="42" direction="north" minutes="54" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="33.5" value="42.909306">
42°54'33.5&quot;
<collectingRegion id="8D6DA1A6FFADFFE2FB7BF86CFB73F816" box="[1209,1231,1925,1952]" country="Japan" name="N" pageId="14" pageNumber="15">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFADFFE2FB18F86CFA3AF829" box="[1242,1414,1925,1951]" degrees="141" direction="east" minutes="07" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="28.9" value="141.1247">141°07'28.9&quot;E</geoCoordinate>
),1 mature female nymph,
<date id="3B174984FFADFFE2FE65F840FD95F875" box="[423,553,1961,1987]" pageId="14" pageNumber="15" value="2002-09-08">08.IX.2002</date>
, K. Saito;
<emphasis id="7DDDB356FFADFFE2FD69F840FD6EF875" bold="true" box="[683,722,1961,1987]" pageId="14" pageNumber="15">[7.]</emphasis>
Sapporo-shi, Minami-ku, Jozankei, Otarunai-gawa Stream (a tributary of Toyohira-gawa Riv.),
<quantity id="8851C2A1FFADFFE2FDD3F824FDE4F851" box="[529,600,1997,2023]" metricMagnitude="2" metricUnit="m" metricValue="3.94" pageId="14" pageNumber="15" unit="m" value="394.0">394 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFADFFE2FD60F824FC82F85E" box="[674,830,1997,2024]" degrees="43" direction="north" minutes="01" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="56.1" value="43.03225">
43°01'56.1&quot;
<collectingRegion id="8D6DA1A6FFADFFE2FCE5F824FC82F85E" box="[807,830,1997,2024]" country="Japan" name="N" pageId="14" pageNumber="15">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFADFFE2FC8BF824FC4CF851" box="[841,1008,1997,2023]" degrees="141" direction="east" minutes="07" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="42.9" value="141.12859">141°07'42.9&quot;E</geoCoordinate>
), 3 mature male nymphs and 1 mature female nymph,
<date id="3B174984FFACFFE3FE89FF7EFE71FF07" box="[331,461,151,177]" pageId="15" pageNumber="16" value="2007-09-11">11.IX.2007</date>
, K. Saito;
<emphasis id="7DDDB356FFACFFE3FD88FF7FFDC3FF06" bold="true" box="[586,639,150,176]" pageId="15" pageNumber="16">[11.]</emphasis>
Sapporo-shi, Minami-ku, Toyotaki, Toyohira-gawa Riv., mainstream,
<quantity id="8851C2A1FFACFFE3FF55FF52FF5EFF63" box="[151,226,187,213]" metricMagnitude="2" metricUnit="m" metricValue="1.88" pageId="15" pageNumber="16" unit="m" value="188.0">188 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FEF1FF53FE73FF63" box="[307,463,186,213]" degrees="42" direction="north" minutes="57" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="38.4" value="42.960667">
42°57'38.4&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FE7BFF53FE73FF63" box="[441,463,186,213]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FE1CFF53FD3AFF63" box="[478,646,186,213]" degrees="141" direction="east" minutes="13" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="50.9" value="141.2308">141°13'50.9&quot;E</geoCoordinate>
), 1 mature male nymph,
<date id="3B174984FFACFFE3FC6CFF53FB8CFF63" box="[942,1072,186,213]" pageId="15" pageNumber="16" value="2005-06-14">
14.
<collectionCode id="29B8F781FFACFFE3FC0FFF53FC52FF63" box="[973,1006,186,213]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.2005
</date>
, K. Saito; ditto, 1 mature male nymph,
<date id="3B174984FFACFFE3FF36FF37FE35FF4F" box="[244,393,222,249]" pageId="15" pageNumber="16" value="2005-08-13">13.VIII.2005</date>
, K. Saito; ditto,
<specimenCount id="59AFA4CDFFACFFE3FD80FF36FD32FF4F" box="[578,654,223,249]" count="3" pageId="15" pageNumber="16" type="male">3 male</specimenCount>
imagines,
<date id="3B174984FFACFFE3FCCBFF37FC37FF4F" box="[777,907,222,249]" pageId="15" pageNumber="16" value="2020-06-15">
15.
<collectionCode id="29B8F781FFACFFE3FCEAFF37FCF5FF4F" box="[808,841,222,249]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.2020
</date>
,
<collectionCode id="29B8F781FFACFFE3FC55FF36FC15FF4F" box="[919,937,223,249]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi;
<emphasis id="7DDDB356FFACFFE3FB86FF37FBC6FF4E" bold="true" box="[1092,1146,222,248]" pageId="15" pageNumber="16">[12.]</emphasis>
Sapporo-shi, Minami-ku, Makomanai, Toyohira-gawa Riv., mainstream,
<quantity id="8851C2A1FFACFFE3FD64FEEAFD63FEAB" box="[678,735,259,285]" metricMagnitude="1" metricUnit="m" metricValue="8.2" pageId="15" pageNumber="16" unit="m" value="82.0">82 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FCEAFEEBFC79FEAB" box="[808,965,258,285]" degrees="42" direction="north" minutes="59" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="02.5" value="42.984028">
42°59'02.5&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FC6DFEEBFC79FEAB" box="[943,965,258,285]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FC12FEEBFBCBFEAB" box="[976,1143,258,285]" degrees="141" direction="east" minutes="20" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="23.5" value="141.33986">141°20'23.5&quot;E</geoCoordinate>
), 1 mature female nymph,
<date id="3B174984FFACFFE3FF55FECEFEAAFEF7" box="[151,278,295,321]" pageId="15" pageNumber="16" value="2019-04-12">12.IV.2019</date>
,
<collectionCode id="29B8F781FFACFFE3FEDDFECEFE8DFEF7" box="[287,305,295,321]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi:
<emphasis id="7DDDB356FFACFFE3FE0AFECFFE42FEF6" bold="true" box="[456,510,294,320]" pageId="15" pageNumber="16">[13.]</emphasis>
Sapporo-shi, Minami-ku, Makomanai, Toyohira-gawa Riv., mainstream,
<quantity id="8851C2A1FFACFFE3FAE9FECEFADFFEF7" box="[1323,1379,295,321]" metricMagnitude="1" metricUnit="m" metricValue="6.8" pageId="15" pageNumber="16" unit="m" value="68.0">68 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FF5DFEA3FE80FED3" box="[159,316,330,357]" degrees="42" direction="north" minutes="59" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="58.5" value="42.999584">
42°59'58.5&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FEE4FEA3FE80FED3" box="[294,316,330,357]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FE8AFEA2FE4CFED3" box="[328,496,331,357]" degrees="141" direction="east" minutes="20" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="26.0" value="141.34055">141°20'26.0&quot;E</geoCoordinate>
), 5 mature male nymphs and 6 mature female nymphs,
<date id="3B174984FFACFFE3FBA5FEA2FB55FED3" box="[1127,1257,331,357]" pageId="15" pageNumber="16" value="2006-06-26">
26.
<collectionCode id="29B8F781FFACFFE3FB4AFEA2FB14FED3" box="[1160,1192,331,357]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.2006
</date>
, K. Saito; ditto, 2 mature male nymphs and 3 mature female nymphs,
<date id="3B174984FFACFFE3FD2DFE86FCCDFE3F" box="[751,881,367,393]" pageId="15" pageNumber="16" value="2006-09-02">02.IX.2006</date>
, K. Saito;
<emphasis id="7DDDB356FFACFFE3FC24FE87FBA0FE3E" bold="true" box="[998,1052,366,392]" pageId="15" pageNumber="16">[14.]</emphasis>
Sapporo-shi, Minami-ku, Minami 30-jo, Nishi 8-chome, Yamahana-gawa Stream (a tributary of Toyohira-gawa Riv.),
<quantity id="8851C2A1FFACFFE3FBA9FE7BFB15FE1B" box="[1131,1193,402,429]" metricMagnitude="1" metricUnit="m" metricValue="5.1" pageId="15" pageNumber="16" unit="m" value="51.0">51 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FB39FE7AFA24FE1B" box="[1275,1432,403,429]" degrees="43" direction="north" minutes="01" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="09.3" value="43.01925">
43°01'09.3&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FA43FE7AFA24FE1B" box="[1409,1432,403,429]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FF55FE5FFE82FE67" box="[151,318,438,465]" degrees="141" direction="east" minutes="20" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="56.1" value="141.3489">141°20'56.1&quot;E</geoCoordinate>
),
<specimenCount id="59AFA4CDFFACFFE3FE91FE5EFE22FE67" box="[339,414,439,465]" count="1" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
imago,
<date id="3B174984FFACFFE3FE39FE5EFDC1FE67" box="[507,637,439,465]" pageId="15" pageNumber="16" value="2018-06-23">
23.
<collectionCode id="29B8F781FFACFFE3FDD9FE5EFD87FE67" box="[539,571,439,465]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.2018
</date>
, F. Matsui;
<emphasis id="7DDDB356FFACFFE3FCC0FE5FFC8BFE66" bold="true" box="[770,823,438,464]" pageId="15" pageNumber="16">[15.]</emphasis>
Sapporo-shi, Chuo-ku, Minami 13-jo, Nishi 1-chome, Toyohira-gawa Riv., mainstream,
<quantity id="8851C2A1FFACFFE3FDD4FE32FDF3FE43" box="[534,591,475,501]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="16" unit="m" value="28.0">28 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FD59FE33FC8BFE43" box="[667,823,474,501]" degrees="43" direction="north" minutes="02" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="35.3" value="43.043137">
43°02'35.3&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FCE3FE33FC8BFE43" box="[801,823,474,501]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FC81FE32FC56FE43" box="[835,1002,475,501]" degrees="141" direction="east" minutes="21" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="31.7" value="141.35881">141°21'31.7&quot;E</geoCoordinate>
), 4 mature male nymphs and 1 mature female nymph,
<date id="3B174984FFACFFE3FE87FE16FE07FDAF" box="[325,443,511,537]" pageId="15" pageNumber="16" value="2020-05-16">
16.
<collectionCode id="29B8F781FFACFFE3FEA6FE16FEC6FDAF" box="[356,378,511,537]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="16" type="Museum">V</collectionCode>
.2020
</date>
,
<collectionCode id="29B8F781FFACFFE3FE07FE16FE6BFDAF" box="[453,471,511,537]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi; ditto, 1 mature male nymph and 1 mature female nymph,
<date id="3B174984FFACFFE3FB30FE16FAC0FDAF" box="[1266,1404,511,537]" pageId="15" pageNumber="16" value="2020-08-19">19.viii.2020</date>
,
<collectionCode id="29B8F781FFACFFE3FA44FE16FA24FDAF" box="[1414,1432,511,537]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi; ditto, 6 mature male nymphs and 1 mature female nymph,
<date id="3B174984FFACFFE3FC7FFDCBFB83FD8B" box="[957,1087,546,573]" pageId="15" pageNumber="16" value="2020-09-25">25.IX.2020</date>
,
<collectionCode id="29B8F781FFACFFE3FB88FDCAFBE0FD8B" box="[1098,1116,547,573]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi; ditto, 1 mature female nymph,
<date id="3B174984FFACFFE3FE8FFDAEFE65FDD7" box="[333,473,583,609]" pageId="15" pageNumber="16" value="2021-07-24">24.VII.2021</date>
,
<collectionCode id="29B8F781FFACFFE3FE25FDAEFE45FDD7" box="[487,505,583,609]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi; ditto, 2 mature female nymphs,
<date id="3B174984FFACFFE3FBD2FDAEFB19FDD7" box="[1040,1189,583,609]" pageId="15" pageNumber="16" value="2021-08-08">08.VIII.2021</date>
,
<collectionCode id="29B8F781FFACFFE3FB76FDAEFB7AFDD7" box="[1204,1222,583,609]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi;
<emphasis id="7DDDB356FFACFFE3FAA5FDAFFA20FDD6" bold="true" box="[1383,1436,582,608]" pageId="15" pageNumber="16">[16.]</emphasis>
Sapporo-shi, Chuo-ku, Kita 1-jo, Higashi 19-chome, Toyohira-gawa Riv., mainstream,
<quantity id="8851C2A1FFACFFE3FBB6FD83FB12FD33" box="[1140,1198,618,645]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="m" value="15.0">15 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FB39FD82FA2BFD33" box="[1275,1431,619,645]" degrees="43" direction="north" minutes="04" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="04.8" value="43.068">
43°04'04.8&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FA43FD82FA2BFD33" box="[1409,1431,619,645]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FF55FD66FE82FD1F" box="[151,318,655,681]" degrees="141" direction="east" minutes="23" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="13.2" value="141.387">141°23'13.2&quot;E</geoCoordinate>
), 11 mature male nymphs and 15 mature female nymphs,
<date id="3B174984FFACFFE3FC06FD66FBFAFD1F" box="[964,1094,655,681]" pageId="15" pageNumber="16" value="2006-06-26">
26.
<collectionCode id="29B8F781FFACFFE3FC24FD66FBBAFD1F" box="[998,1030,655,681]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.2006
</date>
, K. Saito; ditto, 3 mature male nymphs and 9 mature female nymphs,
<date id="3B174984FFACFFE3FD99FD5AFD61FD7B" box="[603,733,691,717]" pageId="15" pageNumber="16" value="2006-09-02">02.IX.2006</date>
, K. Saito; ditto, 1 mature male nymph,
<date id="3B174984FFACFFE3FB6DFD5BFA9BFD7B" box="[1199,1319,690,717]" pageId="15" pageNumber="16" value="2006-10-25">25.X.2006</date>
, K. Saito; ditto,
<specimenCount id="59AFA4CDFFACFFE3FF1AFD3EFE9EFD47" box="[216,290,727,753]" count="1" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
imago,
<date id="3B174984FFACFFE3FEBEFD3EFDB4FD47" box="[380,520,727,753]" pageId="15" pageNumber="16" value="2018-07-26">26.VII.2018</date>
,
<collectionCode id="29B8F781FFACFFE3FDD0FD3EFD98FD47" box="[530,548,727,753]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi; ditto,
<specimenCount id="59AFA4CDFFACFFE3FD3FFD3EFCFBFD47" box="[765,839,727,753]" count="2" pageId="15" pageNumber="16" type="male">2 male</specimenCount>
imagines, 1 mature male nymph and 2 mature female nymphs,
<date id="3B174984FFACFFE3FEC2FD12FE28FCA3" box="[256,404,763,789]" pageId="15" pageNumber="16" value="2021-08-21">21.VIII.2021</date>
,
<collectionCode id="29B8F781FFACFFE3FE63FD12FE0FFCA3" box="[417,435,763,789]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi;
<emphasis id="7DDDB356FFACFFE3FD93FD13FD3AFCA2" bold="true" box="[593,646,762,788]" pageId="15" pageNumber="16">[17.]</emphasis>
Sapporo-shi, Kita-ku, Kita 11-jo, Nishi 1-chome, Sosei-gawa Riv. (a tributary of Fushiko-gawa Riv.),
<quantity id="8851C2A1FFACFFE3FDC9FCF7FDF8FC8F" box="[523,580,798,825]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="m" value="15.0">15 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FD52FCF6FC90FC8F" box="[656,812,799,825]" degrees="43" direction="north" minutes="04" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="30.9" value="43.07525">
43°04'30.9&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FCD4FCF6FC90FC8F" box="[790,812,799,825]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FCFBFCF6FC5CFC8F" box="[825,992,799,825]" degrees="141" direction="east" minutes="21" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="10.9" value="141.35303">141°21'10.9&quot;E</geoCoordinate>
),
<specimenCount id="59AFA4CDFFACFFE3FC37FCF6FBFCFC8F" box="[1013,1088,799,825]" count="2" pageId="15" pageNumber="16" type="male">2 male</specimenCount>
imagines and
<specimenCount id="59AFA4CDFFACFFE3FB24FCF6FAFBFC8F" box="[1254,1351,799,825]" count="1" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
imago,
<date id="3B174984FFACFFE3FF55FCAAFEA5FCEB" box="[151,281,835,861]" pageId="15" pageNumber="16" value="2021-06-23">
23.
<collectionCode id="29B8F781FFACFFE3FF7AFCAAFF65FCEB" box="[184,217,835,861]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="15" pageNumber="16">VI</collectionCode>
.2021
</date>
,
<collectionCode id="29B8F781FFACFFE3FEE6FCAAFE8AFCEB" box="[292,310,835,861]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi; ditto,
<specimenCount id="59AFA4CDFFACFFE3FDD3FCAAFDCEFCEB" box="[529,626,835,861]" count="1" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
imago,
<date id="3B174984FFACFFE3FD0FFCAAFCE5FCEB" box="[717,857,835,861]" pageId="15" pageNumber="16" value="2021-07-02">02.VII.2021</date>
,
<collectionCode id="29B8F781FFACFFE3FCA1FCAAFCC9FCEB" box="[867,885,835,861]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi; ditto,
<specimenCount id="59AFA4CDFFACFFE3FB92FCAAFB0DFCEB" box="[1104,1201,835,861]" count="1" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
imago,
<date id="3B174984FFACFFE3FACEFCAAFA24FCEB" box="[1292,1432,835,861]" pageId="15" pageNumber="16" value="2021-07-12">12.VII.2021</date>
;
<emphasis id="7DDDB356FFACFFE3FF55FC8FFF71FC36" bold="true" box="[151,205,870,896]" pageId="15" pageNumber="16">[21.]</emphasis>
Chitose-shi, Rankoshi, Chitose-gawa Riv., mainstream,
<quantity id="8851C2A1FFACFFE3FC9AFC8EFC29FC37" box="[856,917,871,897]" metricMagnitude="1" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="16" unit="m" value="31.0">31 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FC27FC8EFB3EFC37" box="[997,1154,871,897]" degrees="42" direction="north" minutes="48" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="34.1" value="42.80947">
42°48'34.1&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FBA9FC8EFB3EFC37" box="[1131,1154,871,897]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FB52FC8EFA8BFC37" box="[1168,1335,871,897]" degrees="141" direction="east" minutes="34" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="28.6" value="141.57462">141°34'28.6&quot;E</geoCoordinate>
),
<specimenCount id="59AFA4CDFFACFFE3FA8CFC8EFA20FC37" box="[1358,1436,871,897]" count="1" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
imago,
<date id="3B174984FFACFFE3FF2EFC62FECBFC13" box="[236,375,907,933]" pageId="15" pageNumber="16" value="2020-07-06">06.VII.2020</date>
,
<collectionCode id="29B8F781FFACFFE3FE41FC62FE29FC13" box="[387,405,907,933]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Ito;
<emphasis id="7DDDB356FFACFFE3FE12FC63FDB9FC12" bold="true" box="[464,517,906,932]" pageId="15" pageNumber="16">[22.]</emphasis>
Chitose-shi, Shincho, Chitose-gawa Riv., mainstream,
<quantity id="8851C2A1FFACFFE3FBB4FC62FB0CFC13" box="[1142,1200,907,933]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="16" unit="m" value="13.0">13 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FB39FC62FA24FC13" box="[1275,1432,907,933]" degrees="42" direction="north" minutes="49" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="07.7" value="42.818806">
42°49'07.7&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FA40FC62FA24FC13" box="[1410,1432,907,933]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FF55FC47FE82FC7F" box="[151,318,942,969]" degrees="141" direction="east" minutes="38" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="25.8" value="141.6405">141°38'25.8&quot;E</geoCoordinate>
),
<specimenCount id="59AFA4CDFFACFFE3FE94FC46FE18FC7F" box="[342,420,943,969]" count="1" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
imago and
<specimenCount id="59AFA4CDFFACFFE3FDF0FC46FD2BFC7F" box="[562,663,943,969]" count="3" pageId="15" pageNumber="16" type="female">3 female</specimenCount>
imagines,
<date id="3B174984FFACFFE3FCDAFC47FC26FC7F" box="[792,922,942,969]" pageId="15" pageNumber="16" value="2017-09-05">05.IX.2017</date>
, N. Kuhara; HIYAMA (outside of the study area): Assabu-cho, Shirooka, Sasamodosawa-gawa Riv.,
<quantity id="8851C2A1FFACFFE3FCD7FC3AFCF2FC5B" box="[789,846,979,1005]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="15" pageNumber="16" unit="m" value="76.0">76 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFACFFE3FC5BFC3BFB89FC5B" box="[921,1077,978,1005]" degrees="41" direction="north" minutes="49" orientation="latitude" pageId="15" pageNumber="16" precision="1" seconds="59.0" value="41.833054">
41°49'59.0&quot;
<collectingRegion id="8D6DA1A6FFACFFE3FBDDFC3BFB89FC5B" box="[1055,1077,978,1005]" country="Japan" name="N" pageId="15" pageNumber="16">N</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFACFFE3FB83FC3BFB54FC5B" box="[1089,1256,978,1005]" degrees="140" direction="east" minutes="19" orientation="longitude" pageId="15" pageNumber="16" precision="1" seconds="57.6" value="140.33267">140°19'57.6&quot;E</geoCoordinate>
),
<specimenCount id="59AFA4CDFFACFFE3FB3EFC3AFAF4FC5B" box="[1276,1352,979,1005]" count="1" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
imago,
<date id="3B174984FFACFFE3FF55FC1EFE9FFBA7" box="[151,291,1015,1041]" pageId="15" pageNumber="16" value="2019-07-08">08.VII.2019</date>
,
<collectionCode id="29B8F781FFACFFE3FEECFC1EFEFCFBA7" box="[302,320,1015,1041]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
. Takayanagi.
</paragraph>
<paragraph id="4F166F44FFACFFE3FF05FBF2FD08FB82" blockId="15.[151,1437,150,1977]" box="[199,692,1050,1077]" pageId="15" pageNumber="16">
<emphasis id="7DDDB356FFACFFE3FF05FBF2FD08FB82" bold="true" box="[199,692,1050,1077]" pageId="15" pageNumber="16">
Nymph (mature, in ethanol) (
<figureCitation id="D79273C1FFACFFE3FDE6FBF3FD1BFB82" box="[548,679,1050,1076]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStartId-0="16.[151,250,1659,1683]" captionStartId-1="17.[151,250,945,969]" captionStartId-2="18.[151,250,1339,1363]" captionTargetBox-0="[154,1433,187,1633]" captionTargetBox-1="[154,1433,310,918]" captionTargetBox-2="[182,1410,186,1309]" captionTargetId-0="figure-20@16.[151,1435,181,1636]" captionTargetId-1="figure-90@17.[151,1435,309,921]" captionTargetId-2="figure-20@18.[173,1415,181,1315]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURE 10af. Epeorus L-nigrum, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." captionText-1="FIGURE 11ad. Epeorus L-nigrum, mature nymph. a. Male head, dorsal view; b. Female head, dorsal view; c. Right middle leg, anterior face; d. Gills IVII (from right to left), dorsal view. Scales: ad 1mm." captionText-2="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." figureDoi-0="http://doi.org/10.5281/zenodo.14954007" figureDoi-1="http://doi.org/10.5281/zenodo.14954009" figureDoi-2="http://doi.org/10.5281/zenodo.14954011" httpUri-0="https://zenodo.org/record/14954007/files/figure.png" httpUri-1="https://zenodo.org/record/14954009/files/figure.png" httpUri-2="https://zenodo.org/record/14954011/files/figure.png" pageId="15" pageNumber="16">Figs 1012</figureCitation>
).
</emphasis>
</paragraph>
<paragraph id="4F166F44FFACFFE3FF05FBD6FB64FBEF" blockId="15.[151,1437,150,1977]" box="[199,1240,1087,1113]" pageId="15" pageNumber="16">
Body length 9.811.0 mm (male),
<quantity id="8851C2A1FFACFFE3FD8BFBD6FD6CFBEF" box="[585,720,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" metricValueMax="9.8" metricValueMin="9.4" pageId="15" pageNumber="16" unit="mm" value="9.6" valueMax="9.8" valueMin="9.4">9.49.8 mm</quantity>
(female). Cercus length ca. body length x 1.2.
</paragraph>
<paragraph id="4F166F44FFACFFE3FF05FB8BFF6BF9C3" blockId="15.[151,1437,150,1977]" pageId="15" pageNumber="16">
<emphasis id="7DDDB356FFACFFE3FF05FB8BFEB0FBCB" bold="true" box="[199,268,1122,1149]" italics="true" pageId="15" pageNumber="16">Head.</emphasis>
Shape ellipsoid. Color yellowish-brown to reddish-brown with pale markings. Compound eyes dark-gray to black. Ocelli dark-gray. Antennae brown. Anterior margin densely covered with hair-like setae extending to lateral margins. Dorsal surface of head covered with fine hair-like setae. Sparse longer fine hair-like setae located posterior to eyes (
<figureCitation id="D79273C1FFACFFE3FEA9FB26FE4EFB5F" box="[363,498,1231,1257]" captionStart="FIGURE 11" captionStartId="17.[151,250,945,969]" captionTargetBox="[154,1433,310,918]" captionTargetId="figure-90@17.[151,1435,309,921]" captionTargetPageId="17" captionText="FIGURE 11ad. Epeorus L-nigrum, mature nymph. a. Male head, dorsal view; b. Female head, dorsal view; c. Right middle leg, anterior face; d. Gills IVII (from right to left), dorsal view. Scales: ad 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954009" httpUri="https://zenodo.org/record/14954009/files/figure.png" pageId="15" pageNumber="16">Figs 11a, b</figureCitation>
).
<emphasis id="7DDDB356FFACFFE3FDC9FB26FDD3FB5F" bold="true" box="[523,623,1231,1257]" pageId="15" pageNumber="16">Labrum</emphasis>
: anterior and lateral margins evenly convex, anterior margin concave medially. Ventral surface with long antero-lateral bristles, row of short setae and brush of median fine hair-like setae on each side. Dorsal surface with four long bristles medially and two long bristles near each side of antero-lateral margin, and scattered variable-length setae (
<figureCitation id="D79273C1FFACFFE3FD52FAD2FD4FFAE3" box="[656,755,1339,1365]" captionStart="FIGURE 12" captionStartId="18.[151,250,1339,1363]" captionTargetBox="[182,1410,186,1309]" captionTargetId="figure-20@18.[173,1415,181,1315]" captionTargetPageId="18" captionText="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954011" httpUri="https://zenodo.org/record/14954011/files/figure.png" pageId="15" pageNumber="16">Fig. 12a</figureCitation>
).
<emphasis id="7DDDB356FFACFFE3FCCBFAD3FC3BFAE2" bold="true" box="[777,903,1338,1364]" pageId="15" pageNumber="16">Mandibles</emphasis>
: each outer incisor with three apical teeth and serrated margins, inner incisor with two apical teeth (right mandible) or four apical teeth (left mandible) and outer margin of right inner incisor serrated sharply, right inner incisor slender and almost straight. Tuft of long setae at base of inner incisor of right mandible, tuft of long setae and one brachy-pulmose seta on the base of inner incisor of left mandible (
<figureCitation id="D79273C1FFACFFE3FE94FA22FE6FFA53" box="[342,467,1483,1509]" captionStart="FIGURE 12" captionStartId="18.[151,250,1339,1363]" captionTargetBox="[182,1410,186,1309]" captionTargetId="figure-20@18.[173,1415,181,1315]" captionTargetPageId="18" captionText="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954011" httpUri="https://zenodo.org/record/14954011/files/figure.png" pageId="15" pageNumber="16">Figs 12b, c</figureCitation>
).
<emphasis id="7DDDB356FFACFFE3FE25FA23FDF3FA52" bold="true" box="[487,591,1482,1508]" pageId="15" pageNumber="16">Maxillae</emphasis>
: one long, feather-like seta at base of apical tooth complex, some specimens with two setae (
<figureCitation id="D79273C1FFACFFE3FE84FA06FE15F9BF" box="[326,425,1519,1545]" captionStart="FIGURE 28" captionStartId="36.[151,250,1106,1130]" captionTargetBox="[154,1433,184,1083]" captionTargetId="figure-20@36.[151,1436,181,1083]" captionTargetPageId="36" captionText="FIGURE 28ae. Setae at the base of apical tooth of maxilla (left half right, right half left). a. Epeorus latifolium; b. E. L-nigrum; c. E. curvatulus; d. E. nipponicus; e. E. aesculus." figureDoi="http://doi.org/10.5281/zenodo.14954045" httpUri="https://zenodo.org/record/14954045/files/figure.png" pageId="15" pageNumber="16">Fig. 28b</figureCitation>
).
<emphasis id="7DDDB356FFACFFE3FE79FA06FDDDF9BF" bold="true" box="[443,609,1519,1545]" pageId="15" pageNumber="16">Hypopharynx</emphasis>
: superlinguae distally widened, lingua shape subquadrate.
<emphasis id="7DDDB356FFACFFE3FB33FA07FAECF9BE" bold="true" box="[1265,1360,1518,1544]" pageId="15" pageNumber="16">Labium</emphasis>
: labial palps two segmented, outer margin of proximal one with sparse, thick setae. Distal segment with sparse, hair-like setae on outer margin and dense, brush-like setae on distal 1/3 area. Glossae and paraglossae with long, dense setae.
</paragraph>
</subSubSection>
<subSubSection id="07B33CCFFFACFFFDFF05F997FF61F84C" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="4F166F44FFACFFE3FF05F997FB08F80F" blockId="15.[151,1437,150,1977]" pageId="15" pageNumber="16">
<emphasis id="7DDDB356FFACFFE3FF05F997FE9FF92F" bold="true" box="[199,291,1662,1689]" italics="true" pageId="15" pageNumber="16">Thorax.</emphasis>
Color yellowish-brown to reddish-brown with pale markings dorsally, white ventrally (
<figureCitation id="D79273C1FFACFFE3FB11F996FAA1F92F" box="[1235,1309,1663,1689]" captionStart="FIGURE 10" captionStartId="16.[151,250,1659,1683]" captionTargetBox="[154,1433,187,1633]" captionTargetId="figure-20@16.[151,1435,181,1636]" captionTargetPageId="16" captionText="FIGURE 10af. Epeorus L-nigrum, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." figureDoi="http://doi.org/10.5281/zenodo.14954007" httpUri="https://zenodo.org/record/14954007/files/figure.png" pageId="15" pageNumber="16">Fig 10</figureCitation>
). Pronotum roundly projecting laterally (
<figureCitation id="D79273C1FFACFFE3FE1EF94AFD82F90B" box="[476,574,1699,1725]" captionStart="FIGURE 29" captionStartId="36.[151,250,1557,1581]" captionTargetBox="[152,1428,1237,1525]" captionTargetId="figure-62@36.[151,1436,1227,1533]" captionTargetPageId="36" captionText="FIGURE 29ad. Lateral margin of pronotum, left. a. Epeorus latifolium, and E. L-nigrum; b. E. curvatulus; c. E. nipponicus; d. E. aesculus." figureDoi="http://doi.org/10.5281/zenodo.14954047" httpUri="https://zenodo.org/record/14954047/files/figure.png" pageId="15" pageNumber="16">Fig. 29a</figureCitation>
). Long hair-like setae on medial line to posterior margin of pronotum. Long hair-like setae on medial line to posterior area of mesonotum.
<emphasis id="7DDDB356FFACFFE3FC9BF92EFC24F957" bold="true" box="[857,920,1735,1761]" pageId="15" pageNumber="16">Legs.</emphasis>
Each anterior face of femur with dark-brown transverse markings on its proximal, medial, and distal parts and hypodermal black spots on its base and middle (
<figureCitation id="D79273C1FFACFFE3FF5DF8E6FEBCF89F" box="[159,256,1807,1833]" captionStart="FIGURE 11" captionStartId="17.[151,250,945,969]" captionTargetBox="[154,1433,310,918]" captionTargetId="figure-90@17.[151,1435,309,921]" captionTargetPageId="17" captionText="FIGURE 11ad. Epeorus L-nigrum, mature nymph. a. Male head, dorsal view; b. Female head, dorsal view; c. Right middle leg, anterior face; d. Gills IVII (from right to left), dorsal view. Scales: ad 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954009" httpUri="https://zenodo.org/record/14954009/files/figure.png" pageId="15" pageNumber="16">Fig. 11c</figureCitation>
). Dorsal edge of femur with dense long blade-like setae, ventral edge with sparse short bluntly pointed spines. Femoral surface with diversely shaped setae: oblong, spatulate or semi-circular. Those near ventral edge of each femur tend to be oblong, and those near dorsal edge of each femur tend to be semi-circular. Tibia with two dark-brown markings on base and middle parts and long hair-like setae on dorsal edge. Tarsus darker in proximal area and dorsal edge with long hair-like setae (
<figureCitation id="D79273C1FFACFFE3FD60F876FCBCF80F" box="[674,768,1951,1977]" captionStart="FIGURE 11" captionStartId="17.[151,250,945,969]" captionTargetBox="[154,1433,310,918]" captionTargetId="figure-90@17.[151,1435,309,921]" captionTargetPageId="17" captionText="FIGURE 11ad. Epeorus L-nigrum, mature nymph. a. Male head, dorsal view; b. Female head, dorsal view; c. Right middle leg, anterior face; d. Gills IVII (from right to left), dorsal view. Scales: ad 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954009" httpUri="https://zenodo.org/record/14954009/files/figure.png" pageId="15" pageNumber="16">Fig. 11c</figureCitation>
). Tarsal claw with 45 small denticles.
</paragraph>
<caption id="1BD63FCCFFB3FFFCFF55F992FB9AF90E" ID-DOI="http://doi.org/10.5281/zenodo.14954007" ID-Zenodo-Dep="14954007" httpUri="https://zenodo.org/record/14954007/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1659,1683]" targetBox="[154,1433,187,1633]" targetPageId="16" targetType="figure">
<paragraph id="4F166F44FFB3FFFCFF55F992FB9AF90E" blockId="16.[151,1436,1659,1720]" pageId="16" pageNumber="17">
<emphasis id="7DDDB356FFB3FFFCFF55F992FEFEF925" bold="true" box="[151,322,1659,1683]" pageId="16" pageNumber="17">FIGURE 10af.</emphasis>
<emphasis id="7DDDB356FFB3FFFCFE8AF995FDBEF925" box="[328,514,1660,1683]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="88A914C7FFB3FFFCFE8AF995FE20F925" authorityName="Eaton" authorityYear="1881" box="[328,412,1660,1683]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Epeorus</taxonomicName>
L-nigrum
</emphasis>
, mature nymph, habitus.
<emphasis id="7DDDB356FFB3FFFCFCC6F995FCAAF922" bold="true" box="[772,790,1660,1684]" pageId="16" pageNumber="17">a.</emphasis>
Male, dorsal view;
<emphasis id="7DDDB356FFB3FFFCFC22F995FC48F922" bold="true" box="[992,1012,1660,1684]" pageId="16" pageNumber="17">b.</emphasis>
Male, ventral view;
<emphasis id="7DDDB356FFB3FFFCFB04F995FB64F922" bold="true" box="[1222,1240,1660,1684]" pageId="16" pageNumber="17">c.</emphasis>
Male, lateral view;
<emphasis id="7DDDB356FFB3FFFCFF55F949FF10F90E" bold="true" box="[151,172,1696,1720]" pageId="16" pageNumber="17">d.</emphasis>
Female, dorsal view;
<emphasis id="7DDDB356FFB3FFFCFE4EF949FE21F90E" bold="true" box="[396,413,1696,1720]" pageId="16" pageNumber="17">e.</emphasis>
Female, ventral view;
<emphasis id="7DDDB356FFB3FFFCFD44F976FD2AF901" bold="true" box="[646,662,1695,1719]" pageId="16" pageNumber="17">f.</emphasis>
Female, lateral view.
<emphasis id="7DDDB356FFB3FFFCFCB7F976FC0BF901" bold="true" box="[885,951,1695,1719]" pageId="16" pageNumber="17">Scales</emphasis>
: af 2mm.
</paragraph>
</caption>
<paragraph id="4F166F44FFB3FFFDFF05F8E0FB04FEAB" blockId="16.[151,1437,1801,2008]" lastBlockId="17.[151,1437,151,285]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="7DDDB356FFB3FFFCFF05F8E0FE80F892" bold="true" box="[199,316,1801,1828]" italics="true" pageId="16" pageNumber="17">Abdomen.</emphasis>
Terga yellowish-brown to reddish-brown, each tergum IIIIX with a pair of dark spots. Sterna white. Surface of each tergum with a cluster of long setae located to each side of midline. Posterior margin of each tergum with long setae and long, sharp denticles which are uniform and arranged in an even line (
<figureCitation id="D79273C1FFB3FFFCFB50F8BBFB44F8DA" box="[1170,1272,1874,1900]" captionStart="FIGURE 12" captionStartId="18.[151,250,1339,1363]" captionTargetBox="[182,1410,186,1309]" captionTargetId="figure-20@18.[173,1415,181,1315]" captionTargetPageId="18" captionText="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954011" httpUri="https://zenodo.org/record/14954011/files/figure.png" pageId="16" pageNumber="17">
Figs
<quantity id="8851C2A1FFB3FFFCFB0EF8BBFB44F8DA" box="[1228,1272,1874,1900]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="16" pageNumber="17" unit="g" value="12.0">12g</quantity>
</figureCitation>
,
<figureCitation id="D79273C1FFB3FFFCFAC7F8BBFA8EF8DA" box="[1285,1330,1874,1900]" captionStart="FIGURE 31" captionStartId="38.[151,250,1520,1544]" captionTargetBox="[153,1432,184,1492]" captionTargetId="figure-20@38.[151,1435,181,1495]" captionTargetPageId="38" captionText="FIGURE 31ae. SEM images of abdominal tergum VII (left half). a. Epeorus latifolium; b. E. L-nigrum; c. E. curvatulus; d. E. nipponicus; e. E. aesculus. Scales: ae 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14954051" httpUri="https://zenodo.org/record/14954051/files/figure.png" pageId="16" pageNumber="17">31b</figureCitation>
). Tergum IVII each with three postero-lateral projections, ventral one blunt, lateral one long, pointed and bending slightly upward and dorsal one blunt (
<figureCitation id="D79273C1FFB3FFFCFE2CF873FDF3F802" box="[494,591,1946,1972]" captionStart="FIGURE 30" captionStartId="37.[151,250,1300,1324]" captionTargetBox="[163,1425,193,1260]" captionTargetId="figure-19@37.[151,1435,181,1276]" captionTargetPageId="37" captionText="FIGURE 30af. Postero-lateral projections of abdominal tergum IV, dorsal (left) and lateral (right). a. Epeorus latifolium; b. E. L-nigrum; c. E. curvatulus; d. E. nipponicus; e. E. aesculus; f. Correspondence of projections from a different view (dotted lines)." figureDoi="http://doi.org/10.5281/zenodo.14954049" httpUri="https://zenodo.org/record/14954049/files/figure.png" pageId="16" pageNumber="17">Fig. 30b</figureCitation>
). Posterior margin of Sternum IX with deep medial emargination forming sharp tips and covered in hair-like setae (
<figureCitation id="D79273C1FFB3FFFCFDA3F857FD5DF86E" box="[609,737,1982,2008]" captionStart="FIGURE 12" captionStartId="18.[151,250,1339,1363]" captionTargetBox="[182,1410,186,1309]" captionTargetId="figure-20@18.[173,1415,181,1315]" captionTargetPageId="18" captionText="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954011" httpUri="https://zenodo.org/record/14954011/files/figure.png" pageId="16" pageNumber="17">Figs 12d, e</figureCitation>
). Dorsal surface of cercus with a row of fine setae.
<emphasis id="7DDDB356FFB3FFFCFAE2F857FAE3F86E" bold="true" box="[1312,1375,1982,2008]" pageId="16" pageNumber="17">Gills.</emphasis>
Gills III and IV largest in size, VII smallest. Each gill with variably sized, dark-violet spots on its posterior half, ground color slightly purple (
<figureCitation id="D79273C1FFB2FFFDFE4BFF52FE55FF63" box="[393,489,187,213]" captionStart="FIGURE 11" captionStartId="17.[151,250,945,969]" captionTargetBox="[154,1433,310,918]" captionTargetId="figure-90@17.[151,1435,309,921]" captionTargetPageId="17" captionText="FIGURE 11ad. Epeorus L-nigrum, mature nymph. a. Male head, dorsal view; b. Female head, dorsal view; c. Right middle leg, anterior face; d. Gills IVII (from right to left), dorsal view. Scales: ad 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954009" httpUri="https://zenodo.org/record/14954009/files/figure.png" pageId="17" pageNumber="18">Fig. 11d</figureCitation>
). Gills IIVII with anal-proximal projections, dark colored (
<figureCitation id="D79273C1FFB2FFFDFB49FF52FB57FF63" box="[1163,1259,187,213]" captionStart="FIGURE 11" captionStartId="17.[151,250,945,969]" captionTargetBox="[154,1433,310,918]" captionTargetId="figure-90@17.[151,1435,309,921]" captionTargetPageId="17" captionText="FIGURE 11ad. Epeorus L-nigrum, mature nymph. a. Male head, dorsal view; b. Female head, dorsal view; c. Right middle leg, anterior face; d. Gills IVII (from right to left), dorsal view. Scales: ad 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954009" httpUri="https://zenodo.org/record/14954009/files/figure.png" pageId="17" pageNumber="18">Fig. 11d</figureCitation>
). Costal margin of gill I with fine hair-like setae. IIVII costal margin with rough surface of spines and distal margin with fine setae. Each gill plate with filaments forming a fan-shape together. Gill VII without longitudinal fold.
</paragraph>
<caption id="1BD63FCCFFB2FFFDFF55FC58FC62FC5B" ID-DOI="http://doi.org/10.5281/zenodo.14954009" ID-Zenodo-Dep="14954009" httpUri="https://zenodo.org/record/14954009/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,945,969]" targetBox="[154,1433,310,918]" targetPageId="17" targetType="figure">
<paragraph id="4F166F44FFB2FFFDFF55FC58FC62FC5B" blockId="17.[151,1436,945,1005]" pageId="17" pageNumber="18">
<emphasis id="7DDDB356FFB2FFFDFF55FC58FEFCFC7F" bold="true" box="[151,320,945,969]" pageId="17" pageNumber="18">FIGURE 11ad</emphasis>
.
<emphasis id="7DDDB356FFB2FFFDFE8CFC5BFDB5FC7F" box="[334,521,946,969]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="88A914C7FFB2FFFDFE8CFC5BFE1EFC7F" authorityName="Eaton" authorityYear="1881" box="[334,418,946,969]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">Epeorus</taxonomicName>
L-nigrum
</emphasis>
, mature nymph.
<emphasis id="7DDDB356FFB2FFFDFD7AFC58FD77FC7F" bold="true" box="[696,715,945,969]" pageId="17" pageNumber="18">a.</emphasis>
Male head, dorsal view;
<emphasis id="7DDDB356FFB2FFFDFC0DFC58FC5FFC7F" bold="true" box="[975,995,945,969]" pageId="17" pageNumber="18">b.</emphasis>
Female head, dorsal view;
<emphasis id="7DDDB356FFB2FFFDFB3CFC58FAB3FC7F" bold="true" box="[1278,1295,945,969]" pageId="17" pageNumber="18">c.</emphasis>
Right middle leg, anterior face;
<emphasis id="7DDDB356FFB2FFFDFE8DFC3CFED8FC5B" bold="true" box="[335,356,981,1005]" pageId="17" pageNumber="18">d.</emphasis>
Gills IVII (from right to left), dorsal view.
<emphasis id="7DDDB356FFB2FFFDFCEAFC3CFCD6FC5B" bold="true" box="[808,874,981,1005]" pageId="17" pageNumber="18">Scales</emphasis>
: ad 1mm.
</paragraph>
</caption>
<paragraph id="4F166F44FFB2FFFDFF05FBD0FD0FFBE4" blockId="17.[151,1437,1080,2042]" box="[199,691,1080,1107]" pageId="17" pageNumber="18">
<emphasis id="7DDDB356FFB2FFFDFF05FBD0FD0FFBE4" bold="true" box="[199,691,1080,1107]" pageId="17" pageNumber="18">
Male (in ethanol), redescription (
<figureCitation id="D79273C1FFB2FFFDFD92FBD1FD18FBE4" box="[592,676,1080,1106]" captionStart="FIGURE 13" captionStartId="19.[151,250,1448,1472]" captionTargetBox="[151,1431,345,1425]" captionTargetId="figure-106@19.[151,1435,345,1425]" captionTargetPageId="19" captionText="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="17" pageNumber="18">Fig. 13</figureCitation>
).
</emphasis>
</paragraph>
<paragraph id="4F166F44FFB2FFFDFF05FBB5FB4DFBC1" blockId="17.[151,1437,1080,2042]" box="[199,1265,1116,1143]" pageId="17" pageNumber="18">
Body length
<quantity id="8851C2A1FFB2FFFDFE9AFBB5FE4CFBC1" box="[344,496,1116,1143]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" metricValueMax="1.29" metricValueMin="0.96" pageId="17" pageNumber="18" unit="mm" value="11.25" valueMax="12.9" valueMin="9.6">9.612.9 mm</quantity>
. Cercus length ca. body length x 2.8. Forewing length
<quantity id="8851C2A1FFB2FFFDFB96FBB4FB50FBC1" box="[1108,1260,1117,1143]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" metricValueMax="1.33" metricValueMin="0.93" pageId="17" pageNumber="18" unit="mm" value="11.3" valueMax="13.3" valueMin="9.3">9.313.3 mm</quantity>
.
</paragraph>
<paragraph id="4F166F44FFB2FFFDFF05FB69FD4BFB09" blockId="17.[151,1437,1080,2042]" pageId="17" pageNumber="18">
<emphasis id="7DDDB356FFB2FFFDFF05FB69FEB9FB2D" bold="true" box="[199,261,1152,1179]" italics="true" pageId="17" pageNumber="18">Head</emphasis>
. Color white. Frons shape triangular. Compound eyes dark-gray to black, light-blueish-gray when alive. Ocelli grayish-white with dark-gray border (
<figureCitation id="D79273C1FFB2FFFDFD4BFB4DFD56FB09" box="[649,746,1188,1215]" captionStart="FIGURE 13" captionStartId="19.[151,250,1448,1472]" captionTargetBox="[151,1431,345,1425]" captionTargetId="figure-106@19.[151,1435,345,1425]" captionTargetPageId="19" captionText="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="17" pageNumber="18">
Fig.
<quantity id="8851C2A1FFB2FFFDFD7FFB4CFD56FB09" box="[701,746,1189,1215]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="17" pageNumber="18" unit="g" value="13.0">13g</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="4F166F44FFB2FFFDFF05FB21FD94F9B4" blockId="17.[151,1437,1080,2042]" pageId="17" pageNumber="18">
<emphasis id="7DDDB356FFB2FFFDFF05FB21FE9FFB55" bold="true" box="[199,291,1224,1251]" italics="true" pageId="17" pageNumber="18">Thorax.</emphasis>
Ground color brownish-white to translucent, internal tissues visible. Pronotum with a pair of black line. Mesonotum with thick black markings on adjacent parts of sutures (
<figureCitation id="D79273C1FFB2FFFDFC19FB05FB83FAB1" box="[987,1087,1260,1287]" captionStart="FIGURE 13" captionStartId="19.[151,250,1448,1472]" captionTargetBox="[151,1431,345,1425]" captionTargetId="figure-106@19.[151,1435,345,1425]" captionTargetPageId="19" captionText="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="17" pageNumber="18">
Fig.
<quantity id="8851C2A1FFB2FFFDFBD3FB04FB83FAB1" box="[1041,1087,1261,1287]" metricMagnitude="-2" metricUnit="kg" metricValue="1.3" pageId="17" pageNumber="18" unit="g" value="13.0">13g</quantity>
</figureCitation>
). Meso- and metapleuron with black markings (
<figureCitation id="D79273C1FFB2FFFDFE96FAF9FE09FA9D" box="[340,437,1296,1323]" captionStart="FIGURE 13" captionStartId="19.[151,250,1448,1472]" captionTargetBox="[151,1431,345,1425]" captionTargetId="figure-106@19.[151,1435,345,1425]" captionTargetPageId="19" captionText="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="17" pageNumber="18">Fig. 13a</figureCitation>
).
<emphasis id="7DDDB356FFB2FFFDFE08FAF9FDA1FA9C" bold="true" box="[458,541,1296,1322]" pageId="17" pageNumber="18">Wings.</emphasis>
Hyaline, veins white to translucent. Pterostigmatic area cloudy. Forewing with black markings on the base of radius and cubitus to costal brace, junction of costal brace and costa and base of costa (
<figureCitation id="D79273C1FFB2FFFDFF22FAB1FE83FAC5" box="[224,319,1368,1395]" captionStart="FIGURE 13" captionStartId="19.[151,250,1448,1472]" captionTargetBox="[151,1431,345,1425]" captionTargetId="figure-106@19.[151,1435,345,1425]" captionTargetPageId="19" captionText="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="17" pageNumber="18">Fig. 13e</figureCitation>
). Forewing axillary cord color black. Hindwing with black spot on the base of cubitus to the base of radius.
<emphasis id="7DDDB356FFB2FFFDFF28FA94FE95FA21" bold="true" box="[234,297,1405,1431]" pageId="17" pageNumber="18">Legs.</emphasis>
Anterior face of fore femur with black line on its proximal half. Fore tibia with dark spot on its distal tip. Fore tarsus longest with each segment becoming shorter distally and each joint dark. Mid and hind femur with discontinuous black lines. Fourth segment of mid and hind tarsus shortest (
<figureCitation id="D79273C1FFB2FFFDFC17FA2DFBEBFA68" box="[981,1111,1476,1502]" captionStart="FIGURE 13" captionStartId="19.[151,250,1448,1472]" captionTargetBox="[151,1431,345,1425]" captionTargetId="figure-106@19.[151,1435,345,1425]" captionTargetPageId="19" captionText="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="17" pageNumber="18">Figs 13bd</figureCitation>
). Tarsal claws dissimilar, one hooked-shape and the other oblong.
</paragraph>
<paragraph id="4F166F44FFB2FFFDFF05F9E5FB9BF900" blockId="17.[151,1437,1080,2042]" pageId="17" pageNumber="18">
<emphasis id="7DDDB356FFB2FFFDFF05F9E5FE80F991" bold="true" box="[199,316,1548,1575]" italics="true" pageId="17" pageNumber="18">Abdomen.</emphasis>
Ground color white to translucent. Posterior margin of each tergum color dark. Segments VIII X color white, internal organs and tissues visible (
<figureCitation id="D79273C1FFB2FFFDFD1CF9D9FCFCF9FD" box="[734,832,1584,1611]" captionStart="FIGURE 13" captionStartId="19.[151,250,1448,1472]" captionTargetBox="[151,1431,345,1425]" captionTargetId="figure-106@19.[151,1435,345,1425]" captionTargetPageId="19" captionText="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="17" pageNumber="18">Fig. 13a</figureCitation>
). Posterior margin of styliger plate shape edged and triangular, forceps base with developed projection which has sharp spines. Forceps four-segmented, with short comb-like spines on each inner surface, distal two segments dark in color, fourth segment with sharp setae on its surface. Penis lobes without spines, titillators developed (
<figureCitation id="D79273C1FFB2FFFDFCD9F975FCC5F900" box="[795,889,1692,1718]" captionStart="FIGURE 13" captionStartId="19.[151,250,1448,1472]" captionTargetBox="[151,1431,345,1425]" captionTargetId="figure-106@19.[151,1435,345,1425]" captionTargetPageId="19" captionText="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="17" pageNumber="18">Fig. 13f</figureCitation>
). Cercus white.
</paragraph>
<paragraph id="4F166F44FFB2FFFDFF05F928FD9DF96C" blockId="17.[151,1437,1080,2042]" box="[199,545,1728,1755]" pageId="17" pageNumber="18">
<emphasis id="7DDDB356FFB2FFFDFF05F928FD9DF96C" bold="true" box="[199,545,1728,1755]" pageId="17" pageNumber="18">
Female (in ethanol) (
<figureCitation id="D79273C1FFB2FFFDFE7CF929FDAEF96C" box="[446,530,1728,1754]" captionStart="FIGURE 14" captionStartId="20.[151,250,1084,1108]" captionTargetBox="[151,1432,273,1058]" captionTargetId="figure-72@20.[151,1435,273,1061]" captionTargetPageId="20" captionText="FIGURE 14ae. Epeorus L-nigrum, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954015" httpUri="https://zenodo.org/record/14954015/files/figure.png" pageId="17" pageNumber="18">Fig. 14</figureCitation>
).
</emphasis>
</paragraph>
<paragraph id="4F166F44FFB2FFFDFF05F90DFAB1F949" blockId="17.[151,1437,1080,2042]" box="[199,1293,1764,1791]" pageId="17" pageNumber="18">
Body length
<quantity id="8851C2A1FFB2FFFDFE9AF90DFE42F949" box="[344,510,1764,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" metricValueMax="1.31" metricValueMin="1.05" pageId="17" pageNumber="18" unit="mm" value="11.8" valueMax="13.1" valueMin="10.5">10.513.1 mm</quantity>
. Cercus length ca. body length x 2.1. Forewing length
<quantity id="8851C2A1FFB2FFFDFBA0F90DFAB4F949" box="[1122,1288,1764,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.3649999999999998" metricValueMax="1.51" metricValueMin="1.22" pageId="17" pageNumber="18" unit="mm" value="13.649999999999999" valueMax="15.1" valueMin="12.2">12.215.1 mm</quantity>
.
</paragraph>
<paragraph id="4F166F44FFB2FFFDFF05F8E1FA28F894" blockId="17.[151,1437,1080,2042]" box="[199,1428,1800,1827]" pageId="17" pageNumber="18">
<emphasis id="7DDDB356FFB2FFFDFF05F8E1FEB0F895" bold="true" box="[199,268,1800,1827]" italics="true" pageId="17" pageNumber="18">Head.</emphasis>
Color white. Frons shape triangular. Compound eyes dark-gray to black, light-blueish gray when alive.
</paragraph>
<paragraph id="4F166F44FFB2FFFDFF05F8C5FAE1F838" blockId="17.[151,1437,1080,2042]" pageId="17" pageNumber="18">
<emphasis id="7DDDB356FFB2FFFDFF05F8C5FE9FF8F1" bold="true" box="[199,291,1836,1863]" italics="true" pageId="17" pageNumber="18">Thorax.</emphasis>
Coloration as in male imago (
<figureCitation id="D79273C1FFB2FFFDFDB3F8C5FD6DF8F1" box="[625,721,1836,1863]" captionStart="FIGURE 14" captionStartId="20.[151,250,1084,1108]" captionTargetBox="[151,1432,273,1058]" captionTargetId="figure-72@20.[151,1435,273,1061]" captionTargetPageId="20" captionText="FIGURE 14ae. Epeorus L-nigrum, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954015" httpUri="https://zenodo.org/record/14954015/files/figure.png" pageId="17" pageNumber="18">Fig. 14a</figureCitation>
).
<emphasis id="7DDDB356FFB2FFFDFD21F8C5FC8AF8F0" bold="true" box="[739,822,1836,1862]" pageId="17" pageNumber="18">Wings.</emphasis>
Veins slightly brown. Basal markings as in male imago (
<figureCitation id="D79273C1FFB2FFFDFF5DF8B9FEBEF8DD" box="[159,258,1872,1899]" captionStart="FIGURE 14" captionStartId="20.[151,250,1084,1108]" captionTargetBox="[151,1432,273,1058]" captionTargetId="figure-72@20.[151,1435,273,1061]" captionTargetPageId="20" captionText="FIGURE 14ae. Epeorus L-nigrum, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954015" httpUri="https://zenodo.org/record/14954015/files/figure.png" pageId="17" pageNumber="18">Fig. 14a</figureCitation>
).
<emphasis id="7DDDB356FFB2FFFDFED5F8B8FEEAF8DD" bold="true" box="[279,342,1873,1899]" pageId="17" pageNumber="18">Legs.</emphasis>
Coloration and markings as in male imago. Fore tarsus not elongated. Fourth tarsal segment of each leg shortest, other segments almost equal length (
<figureCitation id="D79273C1FFB2FFFDFD38F89DFCC1F838" box="[762,893,1908,1934]" captionStart="FIGURE 14" captionStartId="20.[151,250,1084,1108]" captionTargetBox="[151,1432,273,1058]" captionTargetId="figure-72@20.[151,1435,273,1061]" captionTargetPageId="20" captionText="FIGURE 14ae. Epeorus L-nigrum, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954015" httpUri="https://zenodo.org/record/14954015/files/figure.png" pageId="17" pageNumber="18">Figs 14bd</figureCitation>
). Tarsal claws dissimilar as in male imago.
</paragraph>
<paragraph id="4F166F44FFB2FFFDFF05F871FF61F84C" blockId="17.[151,1437,1080,2042]" pageId="17" pageNumber="18">
<emphasis id="7DDDB356FFB2FFFDFF05F871FE80F805" bold="true" box="[199,316,1944,1971]" italics="true" pageId="17" pageNumber="18">Abdomen.</emphasis>
White to translucent, greenish eggs visible when alive. Posterior margin of subgenital plate almost straight. Posterior margin of sternum IX with shallow emargination and both tips relatively sharp (
<figureCitation id="D79273C1FFB2FFFDFB1EF855FA87F861" box="[1244,1339,1980,2007]" captionStart="FIGURE 14" captionStartId="20.[151,250,1084,1108]" captionTargetBox="[151,1432,273,1058]" captionTargetId="figure-72@20.[151,1435,273,1061]" captionTargetPageId="20" captionText="FIGURE 14ae. Epeorus L-nigrum, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954015" httpUri="https://zenodo.org/record/14954015/files/figure.png" pageId="17" pageNumber="18">Fig. 14e</figureCitation>
). Cercus white.
</paragraph>
</subSubSection>
<caption id="1BD63FCCFFB1FFFEFF55FAD2FB53FA2D" ID-DOI="http://doi.org/10.5281/zenodo.14954011" ID-Zenodo-Dep="14954011" httpUri="https://zenodo.org/record/14954011/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1339,1363]" targetBox="[182,1410,186,1309]" targetPageId="18" targetType="figure">
<paragraph id="4F166F44FFB1FFFEFF55FAD2FB53FA2D" blockId="18.[151,1436,1339,1436]" pageId="18" pageNumber="19">
<emphasis id="7DDDB356FFB1FFFEFF55FAD2FE83FAE5" bold="true" box="[151,319,1339,1363]" pageId="18" pageNumber="19">FIGURE 12ag</emphasis>
.
<emphasis id="7DDDB356FFB1FFFEFE8EFAD5FDBBFAE5" box="[332,519,1340,1363]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="88A914C7FFB1FFFEFE8EFAD5FE1CFAE5" authorityName="Eaton" authorityYear="1881" box="[332,416,1340,1363]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Epeorus</taxonomicName>
L-nigrum
</emphasis>
, nymph.
<emphasis id="7DDDB356FFB1FFFEFDA4FAD2FDC5FAE5" bold="true" box="[614,633,1339,1363]" pageId="18" pageNumber="19">a.</emphasis>
Labrum (left half in ventral view, right half in dorsal view);
<emphasis id="7DDDB356FFB1FFFEFB22FAD2FB48FAE5" bold="true" box="[1248,1268,1339,1363]" pageId="18" pageNumber="19">b.</emphasis>
Incisors of right mandible (ventral view);
<emphasis id="7DDDB356FFB1FFFEFE59FAB6FE11FAC1" bold="true" box="[411,429,1375,1399]" pageId="18" pageNumber="19">c.</emphasis>
Incisors of left mandible (ventral view);
<emphasis id="7DDDB356FFB1FFFEFC94FAB6FCD6FAC1" bold="true" box="[854,874,1375,1399]" pageId="18" pageNumber="19">d.</emphasis>
Sternum IX of male;
<emphasis id="7DDDB356FFB1FFFEFB8CFAB6FBE3FAC1" bold="true" box="[1102,1119,1375,1399]" pageId="18" pageNumber="19">e.</emphasis>
Sternum IX of female;
<emphasis id="7DDDB356FFB1FFFEFA94FAB6FAD9FAC1" bold="true" box="[1366,1381,1375,1399]" pageId="18" pageNumber="19">f.</emphasis>
New exoskeleton of incisors of left mandible, through the old one, ventral view;
<emphasis id="7DDDB356FFB1FFFEFC51FA6AFC1AFA2D" bold="true" box="[915,934,1411,1435]" pageId="18" pageNumber="19">g.</emphasis>
Posterior margin of tergum VII.
</paragraph>
</caption>
<subSubSection id="07B33CCFFFB1FFFFFF05FA04FADEFEF7" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="4F166F44FFB1FFFFFF05FA04FADEFEF7" blockId="18.[151,1437,1517,2011]" lastBlockId="19.[151,1437,150,321]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="7DDDB356FFB1FFFEFF05FA04FDDBF9BE" bold="true" box="[199,615,1517,1544]" pageId="18" pageNumber="19">
Diagnosis and comparison.
<emphasis id="7DDDB356FFB1FFFEFDCFFA04FDDBF9BE" bold="true" box="[525,615,1517,1544]" italics="true" pageId="18" pageNumber="19">Nymph.</emphasis>
</emphasis>
The nymph of
<emphasis id="7DDDB356FFB1FFFEFCD2FA07FC24F9B1" box="[784,920,1518,1543]" italics="true" pageId="18" pageNumber="19">E. L-nigrum</emphasis>
is quite similar to that of
<taxonomicName id="88A914C7FFB1FFFEFB70FA07FA81F9B1" box="[1202,1341,1517,1543]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB1FFFEFB70FA07FA81F9B1" box="[1202,1341,1517,1543]" italics="true" pageId="18" pageNumber="19">E. latifolium</emphasis>
</taxonomicName>
. Among our samples, the body size of
<taxonomicName id="88A914C7FFB1FFFEFE25F9FBFDCFF99D" box="[487,627,1553,1579]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB1FFFEFE25F9FBFDCFF99D" box="[487,627,1553,1579]" italics="true" pageId="18" pageNumber="19">E. latifolium</emphasis>
</taxonomicName>
is larger than
<emphasis id="7DDDB356FFB1FFFEFCD6F9FBFC21F99D" box="[788,925,1554,1579]" italics="true" pageId="18" pageNumber="19">E. L-nigrum</emphasis>
from May to June (reaching up to
<quantity id="8851C2A1FFB1FFFEFAE4F9F8FAC9F99A" box="[1318,1397,1553,1580]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="mm" value="15.0">15 mm</quantity>
vs.
<quantity id="8851C2A1FFB1FFFEFF55F9DCFF54F9E6" box="[151,232,1589,1616]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="19" unit="mm" value="11.0">11 mm</quantity>
, respectively). However, in samples collected later in the season,
<taxonomicName id="88A914C7FFB1FFFEFC0FF9DFFBE6F9F9" box="[973,1114,1589,1615]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB1FFFEFC0FF9DFFBE6F9F9" box="[973,1114,1589,1615]" italics="true" pageId="18" pageNumber="19">E. latifolium</emphasis>
</taxonomicName>
individuals become smaller. The following morphological characteristics can be used to differentiate the nymphs of the two species: 1) the shape of the marginal denticles on the abdominal terga differs, with
<taxonomicName id="88A914C7FFB1FFFEFC54F997FB9FF921" box="[918,1059,1661,1687]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB1FFFEFC54F997FB9FF921" box="[918,1059,1661,1687]" italics="true" pageId="18" pageNumber="19">E. latifolium</emphasis>
</taxonomicName>
having short, fine denticles (
<figureCitation id="D79273C1FFB1FFFEFAA8F994FF09F90D" captionStart="FIGURE 7" captionStartId="12.[151,250,1746,1770]" captionTargetBox="[169,1421,317,1715]" captionTargetId="figure-76@12.[157,1431,309,1723]" captionTargetPageId="12" captionText="FIGURE 7ak. Epeorus latifolium, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill IV; h. Gill VII; i. Sternum IX of male; j. Sternum IX of female; k. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954001" httpUri="https://zenodo.org/record/14954001/files/figure.png" pageId="18" pageNumber="19">Figs 7k</figureCitation>
,
<figureCitation id="D79273C1FFB1FFFEFF01F948FF50F90D" box="[195,236,1697,1723]" captionStart="FIGURE 31" captionStartId="38.[151,250,1520,1544]" captionTargetBox="[153,1432,184,1492]" captionTargetId="figure-20@38.[151,1435,181,1495]" captionTargetPageId="38" captionText="FIGURE 31ae. SEM images of abdominal tergum VII (left half). a. Epeorus latifolium; b. E. L-nigrum; c. E. curvatulus; d. E. nipponicus; e. E. aesculus. Scales: ae 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14954051" httpUri="https://zenodo.org/record/14954051/files/figure.png" pageId="18" pageNumber="19">31a</figureCitation>
) and
<emphasis id="7DDDB356FFB1FFFEFEECF94BFE04F90D" box="[302,440,1698,1723]" italics="true" pageId="18" pageNumber="19">E. L-nigrum</emphasis>
having long, sharp denticles (
<figureCitation id="D79273C1FFB1FFFEFCD2F948FCCAF90D" box="[784,886,1697,1723]" captionStart="FIGURE 12" captionStartId="18.[151,250,1339,1363]" captionTargetBox="[182,1410,186,1309]" captionTargetId="figure-20@18.[173,1415,181,1315]" captionTargetPageId="18" captionText="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954011" httpUri="https://zenodo.org/record/14954011/files/figure.png" pageId="18" pageNumber="19">
Figs
<quantity id="8851C2A1FFB1FFFEFC88F948FCCAF90D" box="[842,886,1697,1723]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="18" pageNumber="19" unit="g" value="12.0">12g</quantity>
</figureCitation>
,
<figureCitation id="D79273C1FFB1FFFEFC41F948FC0DF90D" box="[899,945,1697,1723]" captionStart="FIGURE 31" captionStartId="38.[151,250,1520,1544]" captionTargetBox="[153,1432,184,1492]" captionTargetId="figure-20@38.[151,1435,181,1495]" captionTargetPageId="38" captionText="FIGURE 31ae. SEM images of abdominal tergum VII (left half). a. Epeorus latifolium; b. E. L-nigrum; c. E. curvatulus; d. E. nipponicus; e. E. aesculus. Scales: ae 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14954051" httpUri="https://zenodo.org/record/14954051/files/figure.png" pageId="18" pageNumber="19">31b</figureCitation>
); 2) the posterior tip of abdominal sternum IX is rounded in
<taxonomicName id="88A914C7FFB1FFFEFEA3F92FFE4CF969" box="[353,496,1733,1759]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB1FFFEFEA3F92FFE4CF969" box="[353,496,1733,1759]" italics="true" pageId="18" pageNumber="19">E. latifolium</emphasis>
</taxonomicName>
(
<figureCitation id="D79273C1FFB1FFFEFDC0F92CFDD2F969" box="[514,622,1733,1759]" captionStart="FIGURE 7" captionStartId="12.[151,250,1746,1770]" captionTargetBox="[169,1421,317,1715]" captionTargetId="figure-76@12.[157,1431,309,1723]" captionTargetPageId="12" captionText="FIGURE 7ak. Epeorus latifolium, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill IV; h. Gill VII; i. Sternum IX of male; j. Sternum IX of female; k. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954001" httpUri="https://zenodo.org/record/14954001/files/figure.png" pageId="18" pageNumber="19">Figs 7i, j</figureCitation>
) but pointed in
<emphasis id="7DDDB356FFB1FFFEFCEEF92FFC04F969" box="[812,952,1734,1759]" italics="true" pageId="18" pageNumber="19">E. L-nigrum</emphasis>
(
<figureCitation id="D79273C1FFB1FFFEFC08F92CFBECF969" box="[970,1104,1733,1759]" captionStart="FIGURE 12" captionStartId="18.[151,250,1339,1363]" captionTargetBox="[182,1410,186,1309]" captionTargetId="figure-20@18.[173,1415,181,1315]" captionTargetPageId="18" captionText="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954011" httpUri="https://zenodo.org/record/14954011/files/figure.png" pageId="18" pageNumber="19">Figs 12d, e</figureCitation>
); and 3) the outer margin of the left mandibles inner incisor lacks sharp serrations in
<emphasis id="7DDDB356FFB1FFFEFCE5F903FC0EF8B5" box="[807,946,1770,1795]" italics="true" pageId="18" pageNumber="19">E. L-nigrum</emphasis>
(
<figureCitation id="D79273C1FFB1FFFEFC00F900FBFCF8B5" box="[962,1088,1769,1795]" captionStart="FIGURE 12" captionStartId="18.[151,250,1339,1363]" captionTargetBox="[182,1410,186,1309]" captionTargetId="figure-20@18.[173,1415,181,1315]" captionTargetPageId="18" captionText="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954011" httpUri="https://zenodo.org/record/14954011/files/figure.png" pageId="18" pageNumber="19">Figs 12c, f</figureCitation>
) but has sharp serrations in
<taxonomicName id="88A914C7FFB1FFFEFA46F903FEB8F891" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB1FFFEFA46F903FEB8F891" italics="true" pageId="18" pageNumber="19">E. latifolium</emphasis>
</taxonomicName>
(
<figureCitation id="D79273C1FFB1FFFEFED6F8E4FEDBF891" box="[276,359,1805,1831]" captionStart="FIGURE 7" captionStartId="12.[151,250,1746,1770]" captionTargetBox="[169,1421,317,1715]" captionTargetId="figure-76@12.[157,1431,309,1723]" captionTargetPageId="12" captionText="FIGURE 7ak. Epeorus latifolium, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill IV; h. Gill VII; i. Sternum IX of male; j. Sternum IX of female; k. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954001" httpUri="https://zenodo.org/record/14954001/files/figure.png" pageId="18" pageNumber="19">Fig. 7c</figureCitation>
). However, it appears that characteristics 1) and 2) are reliable diagnostic features earlier in the summer, but become less distinct later in the season (after September). In this study, specimens were differentiated by a combination of these two features. The serration on the mandibles may vary depending on the time elapsed since molting, but observations of the inner cuticle of the incisor suggest that serration is consistently absent in
<emphasis id="7DDDB356FFB1FFFEFF55F877FE9FF801" box="[151,291,1950,1975]" italics="true" pageId="18" pageNumber="19">E. L-nigrum</emphasis>
(
<figureCitation id="D79273C1FFB1FFFEFEF7F874FE2AF801" box="[309,406,1949,1975]" captionStart="FIGURE 12" captionStartId="18.[151,250,1339,1363]" captionTargetBox="[182,1410,186,1309]" captionTargetId="figure-20@18.[173,1415,181,1315]" captionTargetPageId="18" captionText="FIGURE 12ag. Epeorus L-nigrum, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Sternum IX of male; e. Sternum IX of female; f. New exoskeleton of incisors of left mandible, through the old one, ventral view; g. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954011" httpUri="https://zenodo.org/record/14954011/files/figure.png" pageId="18" pageNumber="19">Fig. 12f</figureCitation>
). This morphological trait may be useful for identifying this species, although it requires dissection and high-magnification observation. The proposed morphological description of
<emphasis id="7DDDB356FFB1FFFEFB5BF82BFA9FF86D" box="[1177,1315,1986,2011]" italics="true" pageId="18" pageNumber="19">E. L-nigrum</emphasis>
nymphs is based on specimens identified through molecular analysis.
<emphasis id="7DDDB356FFB0FFFFFCF7FF7FFC7AFF07" bold="true" box="[821,966,150,177]" italics="true" pageId="19" pageNumber="20">Male imago.</emphasis>
Distinguishing between
<emphasis id="7DDDB356FFB0FFFFFB23FF7EFAD0FF06" box="[1249,1388,151,176]" italics="true" pageId="19" pageNumber="20">E. L-nigrum</emphasis>
and
<taxonomicName id="88A914C7FFB0FFFFFF55FF52FE9FFF63" box="[151,291,187,213]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB0FFFFFF55FF52FE9FFF63" box="[151,291,187,213]" italics="true" pageId="19" pageNumber="20">E. latifolium</emphasis>
</taxonomicName>
in male imago is straightforward based on their penial structures:
<emphasis id="7DDDB356FFB0FFFFFBC0FF52FB37FF62" box="[1026,1163,187,212]" italics="true" pageId="19" pageNumber="20">E. L-nigrum</emphasis>
has titillators, which are absent in
<taxonomicName id="88A914C7FFB0FFFFFEC4FF36FE28FF4F" box="[262,404,223,249]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB0FFFFFEC4FF36FE28FF4F" box="[262,404,223,249]" italics="true" pageId="19" pageNumber="20">E. latifolium</emphasis>
</taxonomicName>
. Additionally,
<emphasis id="7DDDB356FFB0FFFFFDFFFF36FD74FF4E" box="[573,712,223,248]" italics="true" pageId="19" pageNumber="20">E. L-nigrum</emphasis>
lacks lateral projections, whereas
<taxonomicName id="88A914C7FFB0FFFFFB91FF36FB5CFF4F" box="[1107,1248,223,249]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB0FFFFFB91FF36FB5CFF4F" box="[1107,1248,223,249]" italics="true" pageId="19" pageNumber="20">E. latifolium</emphasis>
</taxonomicName>
possesses them. The black markings at the base of the forewing also differ:
<emphasis id="7DDDB356FFB0FFFFFC84FEEAFC6EFEAA" box="[838,978,259,284]" italics="true" pageId="19" pageNumber="20">E. L-nigrum</emphasis>
has &quot;L&quot; or &quot;Z&quot;-shaped markings on the costal brace and the base of the radius, while
<taxonomicName id="88A914C7FFB0FFFFFD56FECEFC9CFEF7" box="[660,800,295,321]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB0FFFFFD56FECEFC9CFEF7" box="[660,800,295,321]" italics="true" pageId="19" pageNumber="20">E. latifolium</emphasis>
</taxonomicName>
displays spotted markings only on the costal brace.
</paragraph>
</subSubSection>
<caption id="1BD63FCCFFB0FFFFFF55FA41FD9DF9BE" ID-DOI="http://doi.org/10.5281/zenodo.14954013" ID-Zenodo-Dep="14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1448,1472]" targetBox="[151,1431,345,1425]" targetPageId="19" targetType="figure">
<paragraph id="4F166F44FFB0FFFFFF55FA41FD9DF9BE" blockId="19.[151,1436,1448,1544]" pageId="19" pageNumber="20">
<emphasis id="7DDDB356FFB0FFFFFF55FA41FE83FA76" bold="true" box="[151,319,1448,1472]" pageId="19" pageNumber="20">FIGURE 13ag</emphasis>
.
<emphasis id="7DDDB356FFB0FFFFFE8EFA40FDBBFA76" box="[332,519,1449,1472]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="88A914C7FFB0FFFFFE8EFA40FE1CFA76" authorityName="Eaton" authorityYear="1881" box="[332,416,1449,1472]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Epeorus</taxonomicName>
L-nigrum
</emphasis>
, male imago.
<emphasis id="7DDDB356FFB0FFFFFD54FA41FD15FA76" bold="true" box="[662,681,1448,1472]" pageId="19" pageNumber="20">a.</emphasis>
Habitus, lateral view;
<emphasis id="7DDDB356FFB0FFFFFC4DFA41FC1FFA76" bold="true" box="[911,931,1448,1472]" pageId="19" pageNumber="20">b.</emphasis>
Right hind leg, anterior face;
<emphasis id="7DDDB356FFB0FFFFFB16FA41FB59FA76" bold="true" box="[1236,1253,1448,1472]" pageId="19" pageNumber="20">c.</emphasis>
Right middle leg, anterior face;
<emphasis id="7DDDB356FFB0FFFFFEE7FA25FE86FA52" bold="true" box="[293,314,1484,1508]" pageId="19" pageNumber="20">d.</emphasis>
Right fore leg, anterior face;
<emphasis id="7DDDB356FFB0FFFFFDA9FA25FDC0FA52" bold="true" box="[619,636,1484,1508]" pageId="19" pageNumber="20">e.</emphasis>
Base of forewing, dorsal view;
<emphasis id="7DDDB356FFB0FFFFFC04FA25FC69FA52" bold="true" box="[966,981,1484,1508]" pageId="19" pageNumber="20">f.</emphasis>
Genitalia, ventral view;
<emphasis id="7DDDB356FFB0FFFFFB11FA25FB5AFA52" bold="true" box="[1235,1254,1484,1508]" pageId="19" pageNumber="20">g.</emphasis>
Head and thorax, dorsal view.
<emphasis id="7DDDB356FFB0FFFFFEDAFA19FEE6F9BE" bold="true" box="[280,346,1520,1544]" pageId="19" pageNumber="20">Scales</emphasis>
: a 5mm, bd 1mm.
</paragraph>
</caption>
<subSubSection id="07B33CCFFFB0FFF8FF05F9B3FE4EFA2F" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="discussion">
<paragraph id="4F166F44FFB0FFFFFF05F9B3FA30F89E" blockId="19.[151,1437,1626,2013]" pageId="19" pageNumber="20">
<emphasis id="7DDDB356FFB0FFFFFF05F9B3FE21F9C3" bold="true" box="[199,413,1626,1653]" pageId="19" pageNumber="20">
Remarks.
<emphasis id="7DDDB356FFB0FFFFFE80F9B3FE21F9C3" bold="true" box="[322,413,1626,1653]" italics="true" pageId="19" pageNumber="20">Nymph.</emphasis>
</emphasis>
The nymph was described for the first time.
<emphasis id="7DDDB356FFB0FFFFFC57F9B3FB9AF9C2" bold="true" box="[917,1062,1626,1653]" italics="true" pageId="19" pageNumber="20">Male imago.</emphasis>
The specimens here identified as
<emphasis id="7DDDB356FFB0FFFFFF55F996FE9CF92E" box="[151,288,1663,1688]" italics="true" pageId="19" pageNumber="20">E. L-nigrum</emphasis>
agree with the original description (
<bibRefCitation id="2B3812B5FFB0FFFFFD76F997FCCAF92E" author="Matsumura, S." box="[692,886,1662,1688]" pageId="19" pageNumber="20" refId="ref30492" refString="Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Toko-shoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura 1931</bibRefCitation>
) and the
<typeStatus id="9012D1E6FFB0FFFFFC1FF996FB88F92F" box="[989,1076,1663,1689]" pageId="19" pageNumber="20" type="syntype">syntype</typeStatus>
specimens.
<emphasis id="7DDDB356FFB0FFFFFB7FF997FAD5F92E" bold="true" box="[1213,1385,1662,1689]" italics="true" pageId="19" pageNumber="20">Female imago.</emphasis>
The female imago was described for the first time.
<emphasis id="7DDDB356FFB0FFFFFD5CF94BFC24F90B" bold="true" box="[670,920,1698,1725]" italics="true" pageId="19" pageNumber="20">Subgenus attribution.</emphasis>
This species can be attributed to the subgenus
<taxonomicName id="88A914C7FFB0FFFFFF55F92FFF4BF956" authorityName="Tshernova" authorityYear="1981" box="[151,247,1734,1760]" pageId="19" pageNumber="20" rank="subGenus" subGenus="Belovius">
<emphasis id="7DDDB356FFB0FFFFFF55F92FFF4BF956" box="[151,247,1734,1760]" italics="true" pageId="19" pageNumber="20">Belovius</emphasis>
</taxonomicName>
by the following features: 1) wide-rounded anal-proximal projection on gills IIVI (
<figureCitation id="D79273C1FFB0FFFFFB0AF92FFA96F956" box="[1224,1322,1734,1760]" captionStart="FIGURE 11" captionStartId="17.[151,250,945,969]" captionTargetBox="[154,1433,310,918]" captionTargetId="figure-90@17.[151,1435,309,921]" captionTargetPageId="17" captionText="FIGURE 11ad. Epeorus L-nigrum, mature nymph. a. Male head, dorsal view; b. Female head, dorsal view; c. Right middle leg, anterior face; d. Gills IVII (from right to left), dorsal view. Scales: ad 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954009" httpUri="https://zenodo.org/record/14954009/files/figure.png" pageId="19" pageNumber="20">Fig. 11d</figureCitation>
); 2) Male imago with titillators on its penis (a valuable character in
<taxonomicName id="88A914C7FFB0FFFFFCFFF903FC21F8B2" authorityName="Tshernova" authorityYear="1981" box="[829,925,1770,1796]" pageId="19" pageNumber="42" rank="subGenus" subGenus="Belovius">
<emphasis id="7DDDB356FFB0FFFFFCFFF903FC21F8B2" box="[829,925,1770,1796]" italics="true" pageId="19" pageNumber="20">Belovius</emphasis>
</taxonomicName>
) (
<figureCitation id="D79273C1FFB0FFFFFC7BF903FBABF8B2" box="[953,1047,1770,1796]" captionStart="FIGURE 13" captionStartId="19.[151,250,1448,1472]" captionTargetBox="[151,1431,345,1425]" captionTargetId="figure-106@19.[151,1435,345,1425]" captionTargetPageId="19" captionText="FIGURE 13ag. Epeorus L-nigrum, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view; g. Head and thorax, dorsal view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954013" httpUri="https://zenodo.org/record/14954013/files/figure.png" pageId="19" pageNumber="20">Fig. 13f</figureCitation>
) (
<bibRefCitation id="2B3812B5FFB0FFFFFBF3F903FB02F8B2" author="Kluge, N. J." box="[1073,1214,1770,1796]" pageId="19" pageNumber="20" refId="ref29818" refString="Kluge, N. J. (2004) The Phylogenetic System of Ephemeroptera. Kluwer Academic Publishers, Dordrecht, 442 pp. https://doi.org/10.1007/978-94-007-0872-3" type="book" year="2004">Kluge 2004</bibRefCitation>
).
<bibRefCitation id="2B3812B5FFB0FFFFFB11F903FA20F8B2" author="Tshernova, O. A." box="[1235,1436,1770,1796]" pageId="19" pageNumber="20" pagination="323 - 336" refId="ref31871" refString="Tshernova, O. A. (1981) On the systematics of adult mayflies of the genus Epeorus Eaton, 1881 (Ephemeroptera, Heptageniidae). Entomologicheskoe Obozrenie, 60 (2), 323-336. [in Russian]" type="journal article" year="1981">Tshernova (1981)</bibRefCitation>
probably observed this species and established the subgenus
<taxonomicName id="88A914C7FFB0FFFFFC82F8E7FC1CF89E" authorityName="Tshernova" authorityYear="1981" box="[832,928,1806,1832]" pageId="19" pageNumber="20" rank="subGenus" subGenus="Belovius">
<emphasis id="7DDDB356FFB0FFFFFC82F8E7FC1CF89E" box="[832,928,1806,1832]" italics="true" pageId="19" pageNumber="20">Belovius</emphasis>
</taxonomicName>
. Further notes are in the discussion section.
</paragraph>
<paragraph id="4F166F44FFB0FFF8FF05F8DBFAB4FF4F" blockId="19.[151,1437,1626,2013]" lastBlockId="20.[151,1437,151,249]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
This species was originally described based on male imaginal specimens collected in &quot;Sapporo&quot;. The description appeared in
<emphasis id="7DDDB356FFB0FFFFFEE0F8BFFD4EF8C6" box="[290,754,1878,1904]" italics="true" pageId="19" pageNumber="20">6000 Illustrated Insects of Japan-Empire</emphasis>
(
<bibRefCitation id="2B3812B5FFB0FFFFFCC0F8BFFC7AF8C6" author="Matsumura, S." box="[770,966,1878,1904]" pageId="19" pageNumber="20" refId="ref30492" refString="Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Toko-shoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura 1931</bibRefCitation>
;
<bibRefCitation id="2B3812B5FFB0FFFFFC11F8BFFBB3F8C6" author="Matsumura, S." box="[979,1039,1878,1904]" pageId="19" pageNumber="20" pagination="199 - 200" refId="ref30516" refString="Matsumura, S. (1932) New Genera and New Species Described in &quot; 6000 Illustrated Insects of Japan-Empire &quot;. Insecta matsumurana, 6 (4), 199-200." type="journal article" year="1932">1932</bibRefCitation>
), a publication noted for numerous taxonomic and nomenclatural issues that have complicated species concepts for many
<taxonomicName id="88A914C7FFB0FFFFFBB3F892FB69F823" box="[1137,1237,1915,1941]" class="Flasuviricetes" family="Flaviviridae" genus="Japanese" higherTaxonomySource="CoL" kingdom="Orthornavirae" order="Amarillovirales" pageId="19" pageNumber="47" phylum="Kitrinoviricota" rank="genus">Japanese</taxonomicName>
insects due to its insufficient descriptions (see
<bibRefCitation id="2B3812B5FFB0FFFFFE27F877FDD3F80E" author="Ueno, M." box="[485,623,1950,1976]" pageId="19" pageNumber="20" pagination="104 - 113" refId="ref32080" refString="Ueno, M. (1931 c) About the Dr. Matsumura's &quot; 6000 Illustrated Insects of Japan-Empire &quot;. The Transactions of the Kansai Entomological Society, 2, 104-113. [in Japanese] *" type="journal article" year="1931">Uéno 1931c</bibRefCitation>
regarding mayflies). Fortunately, we were able to examine
<typeStatus id="9012D1E6FFB0FFFFFAD8F876FAC0F80F" box="[1306,1404,1951,1977]" pageId="19" pageNumber="20" type="syntype">syntypes</typeStatus>
of
<emphasis id="7DDDB356FFB0FFFFFF55F82AFE98F86A" box="[151,292,1987,2012]" italics="true" pageId="19" pageNumber="20">E. L-nigrum</emphasis>
and found no contradictions in morphological characters among the materials examined (
<figureCitation id="D79273C1FFB0FFFFFAFAF82BFA33F86A" box="[1336,1423,1986,2012]" captionStart="FIGURE 33" captionStartId="41.[151,250,1657,1681]" captionTargetBox="[154,1434,384,1633]" captionTargetId="figure-126@41.[151,1435,381,1633]" captionTargetPageId="41" captionText="FIGURE 33af. Type specimens of Epeorus L-nigrum. a. Male imago from syntypes of E. L-nigrum, lateral view; b. Ditto, dorsal view; c. Ditto, ventral view; d. Genitalia of the specimen, ventral view; e, f. Labels attached to the specimen. Scales: ac 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14954055" httpUri="https://zenodo.org/record/14954055/files/figure.png" pageId="19" pageNumber="20">Fig. 33</figureCitation>
). This species was considered a synonym of
<taxonomicName id="88A914C7FFB7FFF8FDBCFF7EFCB7FF07" box="[638,779,151,177]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB7FFF8FDBCFF7EFCB7FF07" box="[638,779,151,177]" italics="true" pageId="20" pageNumber="21">E. latifolium</emphasis>
</taxonomicName>
from
<bibRefCitation id="2B3812B5FFB7FFF8FC92FF7EFBB8FF07" author="Imanishi, K." box="[848,1028,151,177]" pageId="20" pageNumber="21" pagination="380 - 394" refId="ref28952" refString="Imanishi, K. (1934) Mayflies from Japanese Torrents IV. Notes on the genus Epeorus. Annotationes Zoologicae Japonense, 14, 380-394." type="journal article" year="1934">Imanishi (1934)</bibRefCitation>
onward; however,
<bibRefCitation id="2B3812B5FFB7FFF8FB1FFF7EFA20FF07" author="Ishiwata, S." box="[1245,1436,151,177]" pageId="20" pageNumber="21" pagination="163 - 200" refId="ref29216" refString="Ishiwata, S. (2001 b) Mayflies of Chiba Prefecture, Checklist, Diagnoses and Keys. Journal of the Natural History Museum and Institute, Chiba, 7, 163-200. [in Japanese]" type="journal article" year="2001">Ishiwata (2001b)</bibRefCitation>
suggested that
<emphasis id="7DDDB356FFB7FFF8FEFDFF52FE74FF62" box="[319,456,187,212]" italics="true" pageId="20" pageNumber="21">E. L-nigrum</emphasis>
and
<taxonomicName id="88A914C7FFB7FFF8FDC2FF52FD30FF63" box="[512,652,187,213]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="latifolium">
<emphasis id="7DDDB356FFB7FFF8FDC2FF52FD30FF63" box="[512,652,187,213]" italics="true" pageId="20" pageNumber="21">E. latifolium</emphasis>
</taxonomicName>
could be distinguished by the shape of male genitalia, recognizing
<emphasis id="7DDDB356FFB7FFF8FA46FF52FEBDFF4E" italics="true" pageId="20" pageNumber="21">E. L-nigrum</emphasis>
(spelled as &quot;
<taxonomicName id="88A914C7FFB7FFF8FE53FF36FDB5FF4F" authorityName="Matsumura" authorityYear="1931" box="[401,521,223,249]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="l-nigrus">
<emphasis id="7DDDB356FFB7FFF8FE53FF36FDB5FF4F" box="[401,521,223,249]" italics="true" pageId="20" pageNumber="21">E. l-nigrus</emphasis>
</taxonomicName>
&quot;) as a valid species (
<bibRefCitation id="2B3812B5FFB7FFF8FD37FF36FC1AFF4F" author="Ishiwata, S." box="[757,934,223,249]" pageId="20" pageNumber="21" pagination="55 - 84" refId="ref29157" refString="Ishiwata, S. (2001 a) A checklist of Japanese Ephemeroptera. In: The 21 st Century and Aquatic Entomology in East Asia. roceedings of the 1 st. Symposium of Aquatic Entomologists in East Asia (AESEA). Korean Society of Aquatic Entomology Korea, Seoul, pp. 55-84." type="book chapter" year="2001">Ishiwata 2001a</bibRefCitation>
). This study supports that view.
</paragraph>
<caption id="1BD63FCCFFB7FFF8FF55FBD5FE1CFB2A" ID-DOI="http://doi.org/10.5281/zenodo.14954015" ID-Zenodo-Dep="14954015" httpUri="https://zenodo.org/record/14954015/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1084,1108]" targetBox="[151,1432,273,1058]" targetPageId="20" targetType="figure">
<paragraph id="4F166F44FFB7FFF8FF55FBD5FE1CFB2A" blockId="20.[151,1437,1084,1180]" pageId="20" pageNumber="21">
<emphasis id="7DDDB356FFB7FFF8FF55FBD5FEFAFBE2" bold="true" box="[151,326,1084,1108]" pageId="20" pageNumber="21">FIGURE 14ae.</emphasis>
<emphasis id="7DDDB356FFB7FFF8FE8FFBD4FDB5FBE2" box="[333,521,1085,1108]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="88A914C7FFB7FFF8FE8FFBD4FE1DFBE2" authorityName="Eaton" authorityYear="1881" box="[333,417,1085,1108]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Epeorus</taxonomicName>
L-nigrum
</emphasis>
, female imago.
<emphasis id="7DDDB356FFB7FFF8FD73FBD5FD78FBE2" bold="true" box="[689,708,1084,1108]" pageId="20" pageNumber="21">a.</emphasis>
Habitus, lateral view;
<emphasis id="7DDDB356FFB7FFF8FC72FBD5FC78FBE2" bold="true" box="[944,964,1084,1108]" pageId="20" pageNumber="21">b.</emphasis>
Right hind leg, anterior face;
<emphasis id="7DDDB356FFB7FFF8FB3FFBD5FAB2FBE2" bold="true" box="[1277,1294,1084,1108]" pageId="20" pageNumber="21">c.</emphasis>
Right middle leg, anterior face;
<emphasis id="7DDDB356FFB7FFF8FE8CFB89FEDEFBCE" bold="true" box="[334,354,1120,1144]" pageId="20" pageNumber="21">d.</emphasis>
Right fore leg, anterior face;
<emphasis id="7DDDB356FFB7FFF8FD4EFB89FD21FBCE" bold="true" box="[652,669,1120,1144]" pageId="20" pageNumber="21">e.</emphasis>
Subgenital plate and sternum IX (with ventral part of tergum), ventral view.
<emphasis id="7DDDB356FFB7FFF8FF55FB6DFF65FB2A" bold="true" box="[151,217,1156,1180]" pageId="20" pageNumber="21">Scales</emphasis>
: a 5mm, bd 1mm.
</paragraph>
</caption>
<paragraph id="4F166F44FFB7FFF8FF05FB06FE4EFA2F" blockId="20.[151,1436,1262,1469]" pageId="20" pageNumber="21">
<emphasis id="7DDDB356FFB7FFF8FF05FB06FDF5FABE" bold="true" box="[199,585,1262,1289]" pageId="20" pageNumber="21">Emergence periods and habitat.</emphasis>
Fully mature nymphs and imagines were continuously found from June to October, suggesting that this species is multivoltine. Mature nymphs collected in May and June were relatively large (ca.,
<quantity id="8851C2A1FFB7FFF8FF0CFADEFE9CFAE7" box="[206,288,1335,1361]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="20" pageNumber="21" unit="mm" value="11.0">11 mm</quantity>
), while those obtained from August to October were smaller (ca.,
<quantity id="8851C2A1FFB7FFF8FC3FFADEFBFEFAE7" box="[1021,1090,1335,1361]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="21" unit="mm" value="8.0">8 mm</quantity>
). This species is most common in the midstream riffles of the Toyohira-gawa River (localities [11.][17.]) but is absent from smaller mountain streams (localities [10.], [19.]).
</paragraph>
</subSubSection>
<subSubSection id="07B33CCFFFB7FFF8FF05FA4BFC20FA0A" box="[199,924,1442,1469]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="4F166F44FFB7FFF8FF05FA4BFC20FA0A" blockId="20.[151,1436,1262,1469]" box="[199,924,1442,1469]" pageId="20" pageNumber="21">
<emphasis id="7DDDB356FFB7FFF8FF05FA4BFEDDFA0A" bold="true" box="[199,353,1442,1468]" pageId="20" pageNumber="21">Distribution.</emphasis>
<collectingCountry id="37BE2FD4FFB7FFF8FEA5FA4AFE1BFA0B" box="[359,423,1443,1469]" name="Japan" pageId="20" pageNumber="21">Japan</collectingCountry>
(
<collectingRegion id="8D6DA1A6FFB7FFF8FE74FA4BFD95FA0A" box="[438,553,1442,1468]" country="Japan" name="Hokkaido" pageId="20" pageNumber="21">Hokkaido</collectingRegion>
, Honshu, Shikoku, and Kyushu).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,890 @@
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<mods:title id="7A0102AED6D6FF1EDE5D514718743515">Taxonomic review of Epeorus s. l. Eaton, 1881 (Ephemeroptera: Heptageniidae) in Sapporo City and adjacent areas, Hokkaido, Japan</mods:title>
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<paragraph id="4F166F44FFB7FFF8FF55F9BEFC0CF9C6" blockId="20.[151,944,1622,1685]" box="[151,944,1622,1649]" pageId="20" pageNumber="21">
<heading id="145ED828FFB7FFF8FF55F9BEFC0CF9C6" bold="true" box="[151,944,1622,1649]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="7DDDB356FFB7FFF8FF55F9BEFC0CF9C6" bold="true" box="[151,944,1622,1649]" pageId="20" pageNumber="21">
<taxonomicName id="88A914C7FFB7FFF8FF55F9BEFCADF9C6" ID-CoL="6FNVL" authority="Matsumura, 1931" authorityName="Matsumura" authorityYear="1931" box="[151,785,1622,1649]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="curvatulus" subGenus="Caucasiron">
<emphasis id="7DDDB356FFB7FFF8FF55F9BEFF40F9C7" bold="true" box="[151,252,1623,1649]" italics="true" pageId="20" pageNumber="21">Epeorus</emphasis>
(
<emphasis id="7DDDB356FFB7FFF8FECFF9BFFE25F9C6" bold="true" box="[269,409,1622,1648]" italics="true" pageId="20" pageNumber="21">Caucasiron</emphasis>
)
<emphasis id="7DDDB356FFB7FFF8FE69F9BFFD90F9C7" bold="true" box="[427,556,1622,1649]" italics="true" pageId="20" pageNumber="21">curvatulus</emphasis>
<bibRefCitation id="2B3812B5FFB7FFF8FDF1F9BEFCADF9C6" author="Matsumura, S." box="[563,785,1622,1649]" pageId="20" pageNumber="21" refId="ref30492" refString="Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Toko-shoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura, 1931</bibRefCitation>
</taxonomicName>
(
<figureCitation id="D79273C1FFB7FFF8FCE2F9BFFC1BF9C6" box="[800,935,1622,1648]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStart-2="FIGURE 17" captionStart-3="FIGURE 18" captionStart-4="FIGURE 19" captionStartId-0="21.[151,250,1829,1853]" captionStartId-1="22.[151,250,968,992]" captionStartId-2="23.[151,250,1882,1906]" captionStartId-3="24.[151,250,1048,1072]" captionStartId-4="24.[151,250,1855,1879]" captionTargetBox-0="[153,1431,419,1803]" captionTargetBox-1="[154,1433,183,940]" captionTargetBox-2="[159,1435,464,1849]" captionTargetBox-3="[151,1433,198,1018]" captionTargetBox-4="[151,1432,1205,1825]" captionTargetId-0="figure-111@21.[151,1435,415,1806]" captionTargetId-1="figure-20@22.[151,1436,181,944]" captionTargetId-2="figure-165@23.[151,1435,453,1858]" captionTargetId-3="figure-20@24.[151,1436,198,1024]" captionTargetId-4="figure-70@24.[151,1435,1205,1831]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionTargetPageId-3="24" captionTargetPageId-4="24" captionText-0="FIGURE 15af. Epeorus curvatulus, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." captionText-1="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." captionText-2="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." captionText-3="FIGURE 18af. Epeorus curvatulus, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view. Scales: a 5 mm, bd 1mm." captionText-4="FIGURE 19ae. Epeorus curvatulus, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi-0="http://doi.org/10.5281/zenodo.14954017" figureDoi-1="http://doi.org/10.5281/zenodo.14954019" figureDoi-2="http://doi.org/10.5281/zenodo.14954021" figureDoi-3="http://doi.org/10.5281/zenodo.14954023" figureDoi-4="http://doi.org/10.5281/zenodo.14954027" httpUri-0="https://zenodo.org/record/14954017/files/figure.png" httpUri-1="https://zenodo.org/record/14954019/files/figure.png" httpUri-2="https://zenodo.org/record/14954021/files/figure.png" httpUri-3="https://zenodo.org/record/14954023/files/figure.png" httpUri-4="https://zenodo.org/record/14954027/files/figure.png" pageId="20" pageNumber="21">Figs 1519</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="4F166F44FFB7FFF8FF55F992FDF7F922" blockId="20.[151,944,1622,1685]" box="[151,587,1658,1685]" pageId="20" pageNumber="21">[Japanse name &quot;Uéno-hirata-kagerou&quot;]</paragraph>
</subSubSection>
<subSubSection id="07B33CCFFFB7FFF9FF55F929FADDFF65" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="reference_group">
<paragraph id="4F166F44FFB7FFF8FF55F929FE90F8C6" blockId="20.[151,1437,1727,2027]" pageId="20" pageNumber="21">
<taxonomicName id="88A914C7FFB7FFF8FF55F929FEE3F961" ID-CoL="6FNVL" authorityName="Matsumura" authorityYear="1931" box="[151,351,1727,1751]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFB7FFF8FF55F929FEE3F961" box="[151,351,1727,1751]" italics="true" pageId="20" pageNumber="21">Epeorus curvatulus</emphasis>
</taxonomicName>
:
<bibRefCitation id="2B3812B5FFB7FFF8FEACF956FDA3F961" author="Matsumura, S." box="[366,543,1727,1751]" pageId="20" pageNumber="21" refId="ref30492" refString="Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Toko-shoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura 1931</bibRefCitation>
: p. 1477 (male imago) [
<collectingCountry id="37BE2FD4FFB7FFF8FCDAF956FCD7F961" box="[792,875,1727,1751]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFB7FFF8FCB7F956FC07F961" box="[885,955,1727,1751]" country="Japan" name="Tokyo" pageId="20" pageNumber="21">Tokyo</collectingRegion>
];
<bibRefCitation id="2B3812B5FFB7FFF8FC0FF956FBC3F961" author="Matsumura, S." box="[973,1151,1727,1751]" pageId="20" pageNumber="21" refId="ref30550" refString="Matsumura, S. (1933) Illustrated Common Insect of Japan V. Shunyodo, Tokyo, 100 pp. [in Japanese with English description.]" type="book" year="1933">Matsumura 1933</bibRefCitation>
: p. 97, pl. 21, fig. 13 (male imago) [
<collectingCountry id="37BE2FD4FFB7FFF8FEDFF937FECCF940" box="[285,368,1758,1782]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
: Honshu];
<bibRefCitation id="2B3812B5FFB7FFF8FE26F937FDC5F940" author="Ishiwata, S." box="[484,633,1758,1782]" pageId="20" pageNumber="21" pagination="1 - 46" refId="ref29256" refString="Ishiwata, S. (2002) Mayflies of Kanagawa Prefecture, Japan. Kanagawa-Chuho, 138, 1-46. [in Japanese]" type="journal article" year="2002">Ishiwata 2002</bibRefCitation>
: p. 21, figs 167170 (nymph, male imago) [
<collectingCountry id="37BE2FD4FFB7FFF8FB8AF937FB27F940" box="[1096,1179,1758,1782]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFB7FFF8FB65F937FAA4F940" box="[1191,1304,1758,1782]" country="Japan" name="Kanagawa" pageId="20" pageNumber="21">Kanagawa</collectingRegion>
];
<bibRefCitation id="2B3812B5FFB7FFF8FAE8F937FED2F8A2" author="Ishiwata, S. &amp; Takemon, Y." pageId="20" pageNumber="21" pagination="31 - 128" refId="ref29453" refString="Ishiwata, S. &amp; Takemon, Y. (2005 b) Ephemeroptera. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic insects of Japan: manual with keys and illustration. Tokai University Press, Kanagawa, pp. 31-128. [in Japanese]" type="book chapter" year="2005">Ishiwata &amp; Takemon 2005b</bibRefCitation>
: p. 105, figs 45-3, 47-4, 47-5 (male imago, nymph) [
<collectingCountry id="37BE2FD4FFB7FFF8FCB9F915FC6CF8A2" box="[891,976,1788,1812]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
];
<bibRefCitation id="2B3812B5FFB7FFF8FC1DF914FB36F8A2" author="Maruyama, H." box="[991,1162,1788,1813]" pageId="20" pageNumber="21" pagination="90 - 178" refId="ref30430" refString="Maruyama, H. (2016) Ephemeroptera. In: Maruyama, H. &amp; Hanada, S. (Eds.), A Field Guide to Japanese Aquatic Insects: Adults of Mayflies, Stoneflies and Caddisflies. Zenkoku Noson Kyoiku Kyokai Co., Ltd, Tokyo, pp. 90-178. [in Japanese]" type="book chapter" year="2016">Maruyama 2016</bibRefCitation>
: p. 171, figs 1-171c, 1-174, 1-261, 1-262, 1-263, 1-264 (male imago, female imago, male subimago, female subimago) [
<collectingCountry id="37BE2FD4FFB7FFF8FBB6F8F2FB7BF885" box="[1140,1223,1819,1843]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFB7FFF8FB10F8F2FAA0F885" box="[1234,1308,1819,1843]" country="Japan" name="Hyogo" pageId="20" pageNumber="21">Hyogo</collectingRegion>
]; Ishiwata
<emphasis id="7DDDB356FFB7FFF8FA48F8F5FF5EF8E4" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2018: p. 126, figs 45-3, 47-4, 47-5 (male imago, nymph) [
<collectingCountry id="37BE2FD4FFB7FFF8FCF5F8D3FC30F8E4" box="[823,908,1850,1874]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
];
<bibRefCitation id="2B3812B5FFB7FFF8FC5EF8D3FB1CF8E4" author="Ishiwata, S. &amp; Fujitani, T." box="[924,1184,1850,1874]" pageId="20" pageNumber="21" pagination="1 - 63" refId="ref29324" refString="Ishiwata, S. &amp; Fujitani, T. (2018) Eggs of Japanese Ephemeroptera. Kanagawa Institute of Technology, Division for Environmental Chemistry Research Report, 7 (Supplement 2), March 2018, 1-63." type="journal article" year="2018">Ishiwata &amp; Fujitani 2018</bibRefCitation>
: p. 32, pl.21, fig. 4 (egg) [
<collectingCountry id="37BE2FD4FFB7FFF8FF0FF8B1FE9EF8C6" box="[205,290,1880,1904]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
].
</paragraph>
<paragraph id="4F166F44FFB7FFF8FF55F891FD87F85D" blockId="20.[151,1437,1727,2027]" pageId="20" pageNumber="21">
<treatmentCitationGroup id="6FB9486AFFB7FFF8FF55F891FD87F85D" pageId="20" pageNumber="21">
<taxonomicName id="88A914C7FFB7FFF8FF55F891FE96F839" ID-CoL="6FNHR" box="[151,298,1912,1935]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="7DDDB356FFB7FFF8FF55F891FE96F839" box="[151,298,1912,1935]" italics="true" pageId="20" pageNumber="21">Epeorus uenoi</emphasis>
</taxonomicName>
auct., non
<bibRefCitation id="2B3812B5FFB7FFF8FE5EF89EFDEEF839" author="Matsumura, S." box="[412,594,1911,1935]" pageId="20" pageNumber="21" refId="ref30550" refString="Matsumura, S. (1933) Illustrated Common Insect of Japan V. Shunyodo, Tokyo, 100 pp. [in Japanese with English description.]" type="book" year="1933">Matsumura, 1933</bibRefCitation>
:
<treatmentCitation id="CE084955FFB7FFF8FD9EF89EFC80F839" author="Imanishi, K." box="[604,828,1911,1935]" page="383" pageId="20" pageNumber="21" year="1934">
<bibRefCitation id="2B3812B5FFB7FFF8FD9EF89EFD4EF839" author="Imanishi, K." box="[604,754,1911,1935]" pageId="20" pageNumber="21" pagination="380 - 394" refId="ref28952" refString="Imanishi, K. (1934) Mayflies from Japanese Torrents IV. Notes on the genus Epeorus. Annotationes Zoologicae Japonense, 14, 380-394." type="journal article" year="1934">Imanishi 1934</bibRefCitation>
: p. 383
</treatmentCitation>
pl. 18, fig. 2 (nymph, male imago, female imago) [
<collectingCountry id="37BE2FD4FFB7FFF8FA84F89EFA25F839" box="[1350,1433,1911,1935]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFB7FFF8FF05F87FFEB0F818" box="[199,268,1942,1966]" country="Japan" name="Kyoto" pageId="20" pageNumber="21">Kyoto</collectingRegion>
];
<treatmentCitation id="CE084955FFB7FFF8FEDFF87FFDBDF81B" author="Imanishi, K." box="[285,513,1941,1966]" page="252" pageId="20" pageNumber="21" year="1940">
<bibRefCitation id="2B3812B5FFB7FFF8FEDFF87FFE08F818" author="Imanishi, K." box="[285,436,1942,1966]" pageId="20" pageNumber="21" pagination="169 - 263" refId="ref28981" refString="Imanishi, K. (1940) Ephemeroptera of Manchuria, Inner Mongolia and Korea. In: Kawamura, T. (Ed.), Report of the liminological survey of Kwantung and Manchoukuo, Kyoto, Japan, 1940, pp. 169-263. [in Japanese]" type="journal article" year="1940">Imanishi 1940</bibRefCitation>
: p. 252
</treatmentCitation>
, fig. 33 (nymph) [
<collectingCountry id="37BE2FD4FFB7FFF8FD7EF87FFCB3F818" box="[700,783,1942,1966]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFB7FFF8FCD8F87FFCE3F818" box="[794,863,1942,1966]" country="Japan" name="Kyoto" pageId="20" pageNumber="21">Kyoto</collectingRegion>
];
<treatmentCitation id="CE084955FFB7FFF8FCB3F87FFB98F818" author="Gose, K." box="[881,1060,1941,1966]" page="20" pageId="20" pageNumber="21" year="1962">
<bibRefCitation id="2B3812B5FFB7FFF8FCB3F87FFC5EF81B" author="Gose, K." box="[881,994,1941,1966]" pageId="20" pageNumber="21" pagination="6 - 24" refId="ref28133" refString="Gose, K. (1962) Ephemeroptera. In: Tsuda, M. (Ed.), Aquatic Entomology. Hokuryukan, Tokyo, pp. 6-24. [in Japanese]" type="book chapter" year="1962">Gose 1962</bibRefCitation>
: p. 20
</treatmentCitation>
, figs 1-11-1a, 1b (nymph) [
<collectingCountry id="37BE2FD4FFB7FFF8FAFCF87FFA2FF818" box="[1342,1427,1942,1966]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
];
<treatmentCitation id="CE084955FFB7FFF8FF05F85DFECBF87A" author="Gose, K." box="[199,375,1972,1996]" page="44" pageId="20" pageNumber="21" year="1979">
<bibRefCitation id="2B3812B5FFB7FFF8FF05F85DFE8BF87A" author="Gose, K." box="[199,311,1972,1996]" pageId="20" pageNumber="21" pagination="44 - 46" refId="ref28170" refString="Gose, K. (1979) The mayflies of Japanese (4). Aquabiology, 1 (4), 44-46. [in Japanese]" type="journal article" year="1979">Gose 1979</bibRefCitation>
, p. 44
</treatmentCitation>
, figs 8, 10, 22 (male imago, nymph) [
<collectingCountry id="37BE2FD4FFB7FFF8FD34F85DFCF7F87A" box="[758,843,1972,1996]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
];
<treatmentCitation id="CE084955FFB7FFF8FC9EF85DFB91F87A" author="Okazaki, H." box="[860,1069,1972,1996]" page="22" pageId="20" pageNumber="21" year="1984">
<bibRefCitation id="2B3812B5FFB7FFF8FC9EF85DFC51F87A" author="Okazaki, H." box="[860,1005,1972,1996]" pageId="20" pageNumber="21" pagination="19 - 27" refId="ref30763" refString="Okazaki, H. (1984) Observation of the eggs of Japanese mayflies by scanning electron microscope. Biology of Inland Waters, 3, 19-27. [in Japanese]" type="journal article" year="1984">Okazaki 1984</bibRefCitation>
: p. 22
</treatmentCitation>
, fig. 46 (egg) [
<collectingCountry id="37BE2FD4FFB7FFF8FB00F85DFAABF87A" box="[1218,1303,1972,1996]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
];
<treatmentCitation id="CE084955FFB7FFF8FAEAF85DFF40F85D" author="Gose, K." page="16" pageId="20" pageNumber="21" year="1985">
<bibRefCitation id="2B3812B5FFB7FFF8FAEAF85DFA24F87A" author="Gose, K." box="[1320,1432,1972,1996]" pageId="20" pageNumber="21" pagination="7 - 32" refId="ref28200" refString="Gose, K. (1985) Ephemeroptera. In: Kawai, T. (Ed.), An Illustrated Book of Aquatic Insects of Japan. Univ. Tokai Press, Tokyo, pp. 7-32. [in Japanese]" type="book chapter" year="1985">Gose 1985</bibRefCitation>
: p. 16
</treatmentCitation>
, figs 34, 35 (nymph) [
<collectingCountry id="37BE2FD4FFB7FFF8FE1EF83AFD8DF85D" box="[476,561,2003,2027]" name="Japan" pageId="20" pageNumber="21">JAPAN</collectingCountry>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="4F166F44FFB6FFF9FF55FF7EFC67FF19" blockId="21.[151,1377,151,212]" box="[151,987,151,175]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="6FB9486AFFB6FFF9FF55FF7EFC67FF19" box="[151,987,151,175]" pageId="21" pageNumber="22">
<taxonomicName id="88A914C7FFB6FFF9FF55FF7EFE24FF18" ID-CoL="6FNHR" box="[151,408,151,175]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="uenoi" subGenus="Belovius">
<emphasis id="7DDDB356FFB6FFF9FF55FF7EFF57FF18" box="[151,235,151,174]" italics="true" pageId="21" pageNumber="22">Epeorus</emphasis>
(
<emphasis id="7DDDB356FFB6FFF9FF38FF7EFEEDFF19" box="[250,337,151,175]" italics="true" pageId="21" pageNumber="22">Belovius</emphasis>
)
<emphasis id="7DDDB356FFB6FFF9FEA2FF7EFE24FF18" box="[352,408,151,174]" italics="true" pageId="21" pageNumber="22">uenoi</emphasis>
</taxonomicName>
:
<treatmentCitation id="CE084955FFB6FFF9FE64FF7EFD2AFF19" author="Tshernova, O. A." box="[422,662,151,175]" page="334" pageId="21" pageNumber="22" year="1981">
<bibRefCitation id="2B3812B5FFB6FFF9FE64FF7EFDF7FF19" author="Tshernova, O. A." box="[422,587,151,175]" pageId="21" pageNumber="22" pagination="323 - 336" refId="ref31871" refString="Tshernova, O. A. (1981) On the systematics of adult mayflies of the genus Epeorus Eaton, 1881 (Ephemeroptera, Heptageniidae). Entomologicheskoe Obozrenie, 60 (2), 323-336. [in Russian]" type="journal article" year="1981">Tshernova 1981</bibRefCitation>
: p. 334
</treatmentCitation>
, fig. 11 (male imago) [
<collectingCountry id="37BE2FD4FFB6FFF9FCBFFF7EFC6EFF19" box="[893,978,151,175]" name="Japan" pageId="21" pageNumber="22">JAPAN</collectingCountry>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="4F166F44FFB6FFF9FF55FF53FADDFF65" blockId="21.[151,1377,151,212]" box="[151,1377,185,212]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="6FB9486AFFB6FFF9FF55FF53FADDFF65" box="[151,1377,185,212]" pageId="21" pageNumber="22">
<taxonomicName id="88A914C7FFB6FFF9FF55FF53FF4CFF65" box="[151,240,185,211]" class="Insecta" family="Heptageniidae" genus="Iron" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="7DDDB356FFB6FFF9FF55FF53FF7AFF65" box="[151,198,186,211]" italics="true" pageId="21" pageNumber="22">Iron</emphasis>
<materialsCitation id="FFC16519FFB6FFF9FF0FFF50FF4CFF65" box="[205,240,185,211]" country="Japan" location="Kyoto" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Kyoto">sp.</materialsCitation>
</taxonomicName>
?:
<treatmentCitation id="CE084955FFB6FFF9FECAFF50FE71FF65" author="Ueno, M." box="[264,461,185,211]" page="28" pageId="21" pageNumber="22" year="1928">
<bibRefCitation id="2B3812B5FFB6FFF9FECAFF50FE34FF65" author="Ueno, M." box="[264,392,185,211]" pageId="21" pageNumber="22" pagination="19 - 63" refId="ref31987" refString="Ueno, M. (1928) Some Japanese mayfly nymphs. Memoirs of the college of science, Kyoto Imperial University, Series B, 4 (1), 19-63." type="journal article" year="1928">Uéno 1928</bibRefCitation>
: p. 28
</treatmentCitation>
, pl. 5, figs 4, 4a4i, pl. 6, figs 4j4n (nymph) [
<collectingCountry id="37BE2FD4FFB6FFF9FC15FF50FB8FFF65" box="[983,1075,185,211]" name="Japan" pageId="21" pageNumber="22">JAPAN</collectingCountry>
:
<collectingRegion id="8D6DA1A6FFB6FFF9FBFCFF50FB3BFF65" box="[1086,1159,185,211]" country="Japan" name="Kyoto" pageId="21" pageNumber="22">Kyoto</collectingRegion>
,
<collectingRegion id="8D6DA1A6FFB6FFF9FB50FF50FB50FF65" box="[1170,1260,185,211]" country="Japan" name="Nagano" pageId="21" pageNumber="22">Nagano</collectingRegion>
,
<collectingRegion id="8D6DA1A6FFB6FFF9FB34FF50FAEAFF65" box="[1270,1366,185,211]" country="Japan" name="Toyama" pageId="21" pageNumber="22">Toyama</collectingRegion>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="07B33CCFFFB6FFF5FF55FEE8FF6CF9F1" lastPageId="25" lastPageNumber="26" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="4F166F44FFB6FFF9FF55FEE8FD1EFE31" blockId="21.[151,1436,256,391]" pageId="21" pageNumber="22">
<emphasis id="7DDDB356FFB6FFF9FF55FEE8FD5CFEAC" bold="true" box="[151,736,256,283]" pageId="21" pageNumber="22">
Materials examined. Type specimen (
<typeStatus id="9012D1E6FFB6FFF9FD9DFEE9FD6FFEAC" box="[607,723,256,282]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
):
</emphasis>
(identification label by Imanishi, 1933 &quot;
<taxonomicName id="88A914C7FFB6FFF9FB7CFEEBFE53FE89" authority="Matsumura, K. Imanishi, 1933" authorityName="Matsumura, K. Imanishi" authorityYear="1933" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFB6FFF9FB7CFEEBFA20FEAD" box="[1214,1436,257,283]" italics="true" pageId="21" pageNumber="22">Epeorus curvatulus</emphasis>
Matsumura, K. Imanishi, 1933
</taxonomicName>
&quot;).
<materialsCitation id="FFC16519FFB6FFF9FDC8FECCFD27FE30" determinerName="Matsumura" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1">
<specimenCount id="59AFA4CDFFB6FFF9FDC8FECCFDEFFE89" box="[522,595,293,319]" count="1" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
imago, &quot;Tamagawa [
<emphasis id="7DDDB356FFB6FFF9FCFCFECFFC12FEF6" bold="true" box="[830,942,294,320]" pageId="21" pageNumber="22">タマガハ</emphasis>
] 7/25 (year not listed)&quot;, &quot;
<taxonomicName id="88A914C7FFB6FFF9FB06FECFFA20FE89" authorityName="Matsumura, K. Imanishi" authorityYear="1933" box="[1220,1436,293,319]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFB6FFF9FB06FECFFA20FE89" box="[1220,1436,293,319]" italics="true" pageId="21" pageNumber="22">Epeorus curvatulus</emphasis>
</taxonomicName>
det.
<determinerName id="BACA8380FFB6FFF9FF0AFEA0FEF2FED5" box="[200,334,329,355]" pageId="21" pageNumber="22">Matsumura</determinerName>
&quot;, &quot;Two tails color white [
<emphasis id="7DDDB356FFB6FFF9FDAFFEA3FC8DFED2" bold="true" box="[621,817,330,356]" pageId="21" pageNumber="22">二*の¹は̊e</emphasis>
]&quot;, &quot;Type Matsumura&quot; (red label), &quot;
<taxonomicName id="88A914C7FFB6FFF9FB03FEA3FF54FE31" authority="Matsu." authorityName="Matsumura" authorityYear="1931" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFB6FFF9FB03FEA3FA20FED5" box="[1217,1436,329,355]" italics="true" pageId="21" pageNumber="22">Epeorus curvatulus</emphasis>
Matsu.
</taxonomicName>
&quot; (probably attached recently)
<emphasis id="7DDDB356FFB6FFF9FDFBFE85FDFEFE30" bold="true" box="[569,578,364,390]" pageId="21" pageNumber="22">(</emphasis>
<figureCitation id="D79273C1FFB6FFF9FD81FE84FD2EFE31" box="[579,658,365,391]" captionStart="FIGURE 34" captionStartId="42.[151,250,1297,1321]" captionTargetBox="[153,1432,240,1270]" captionTargetId="figure-43@42.[151,1435,237,1273]" captionTargetPageId="42" captionText="FIGURE 34af. Type specimen of Epeorus curvatulus. a. Holotype of E. curvatulus, dorsal view; b. Ditto, ventral view; c. Ditto, lateral view; d. Abdomen of the specimen (forceps lost), lateral view; e. Genitalia of the specimen, ventral view; f. Labels attached to the specimen. Scales: ac 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14954057" httpUri="https://zenodo.org/record/14954057/files/figure.png" pageId="21" pageNumber="22">Fig. 34</figureCitation>
<emphasis id="7DDDB356FFB6FFF9FD50FE85FD27FE30" bold="true" box="[658,667,364,390]" pageId="21" pageNumber="22">)</emphasis>
</materialsCitation>
.
</paragraph>
<caption id="1BD63FCCFFB6FFF9FF55F8CCFBC4F8D7" ID-DOI="http://doi.org/10.5281/zenodo.14954017" ID-Zenodo-Dep="14954017" httpUri="https://zenodo.org/record/14954017/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1829,1853]" targetBox="[153,1431,419,1803]" targetPageId="21" targetType="figure">
<paragraph id="4F166F44FFB6FFF9FF55F8CCFBC4F8D7" blockId="21.[151,1436,1829,1889]" pageId="21" pageNumber="22">
<emphasis id="7DDDB356FFB6FFF9FF55F8CCFEF8F88B" bold="true" box="[151,324,1829,1853]" pageId="21" pageNumber="22">FIGURE 15af.</emphasis>
<taxonomicName id="88A914C7FFB6FFF9FE8EF8CFFDA8F88B" authorityName="Matsumura" authorityYear="1931" box="[332,532,1829,1853]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFB6FFF9FE8EF8CFFDA8F88B" box="[332,532,1829,1853]" italics="true" pageId="21" pageNumber="22">Epeorus curvatulus</emphasis>
</taxonomicName>
, mature nymph, habitus.
<emphasis id="7DDDB356FFB6FFF9FCDDF8CCFC8EF88B" bold="true" box="[799,818,1829,1853]" pageId="21" pageNumber="22">a.</emphasis>
Male, dorsal view;
<emphasis id="7DDDB356FFB6FFF9FBC8F8CCFBA3F88B" bold="true" box="[1034,1055,1829,1853]" pageId="21" pageNumber="22">b.</emphasis>
Male, ventral view;
<emphasis id="7DDDB356FFB6FFF9FAC2F8CCFAADF88B" bold="true" box="[1280,1297,1829,1853]" pageId="21" pageNumber="22">c.</emphasis>
Male, lateral view;
<emphasis id="7DDDB356FFB6FFF9FF1EF8A0FF4CF8D7" bold="true" box="[220,240,1865,1889]" pageId="21" pageNumber="22">d.</emphasis>
Female, dorsal view;
<emphasis id="7DDDB356FFB6FFF9FE15F8A0FE55F8D7" bold="true" box="[471,489,1865,1889]" pageId="21" pageNumber="22">e.</emphasis>
Female, ventral view;
<emphasis id="7DDDB356FFB6FFF9FD1AF8A0FD5BF8D7" bold="true" box="[728,743,1865,1889]" pageId="21" pageNumber="22">f.</emphasis>
Female, lateral view.
<emphasis id="7DDDB356FFB6FFF9FC04F8A0FBB4F8D7" bold="true" box="[966,1032,1865,1889]" pageId="21" pageNumber="22">Scales</emphasis>
: af 2mm.
</paragraph>
</caption>
<caption id="1BD63FCCFFB5FFFAFF55FC21FAEBFBB3" ID-DOI="http://doi.org/10.5281/zenodo.14954019" ID-Zenodo-Dep="14954019" httpUri="https://zenodo.org/record/14954019/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,968,992]" targetBox="[154,1433,183,940]" targetPageId="22" targetType="figure">
<paragraph id="4F166F44FFB5FFFAFF55FC21FAEBFBB3" blockId="22.[151,1437,968,1029]" pageId="22" pageNumber="23">
<emphasis id="7DDDB356FFB5FFFAFF55FC21FEFCFC56" bold="true" box="[151,320,968,992]" pageId="22" pageNumber="23">FIGURE 16ae</emphasis>
.
<taxonomicName id="88A914C7FFB5FFFAFE8CFC20FDAAFC56" authorityName="Matsumura" authorityYear="1931" box="[334,534,968,992]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFB5FFFAFE8CFC20FDAAFC56" box="[334,534,968,992]" italics="true" pageId="22" pageNumber="23">Epeorus curvatulus</emphasis>
</taxonomicName>
, mature nymph.
<emphasis id="7DDDB356FFB5FFFAFD0AFC20FD67FC57" bold="true" box="[712,731,969,993]" pageId="22" pageNumber="23">a.</emphasis>
Male head, dorsal view;
<emphasis id="7DDDB356FFB5FFFAFC26FC20FC45FC57" bold="true" box="[996,1017,969,993]" pageId="22" pageNumber="23">b.</emphasis>
Frontal margin of male head;
<emphasis id="7DDDB356FFB5FFFAFAFAFC20FAF5FC57" bold="true" box="[1336,1353,969,993]" pageId="22" pageNumber="23">c.</emphasis>
Female head, dorsal view;
<emphasis id="7DDDB356FFB5FFFAFE95FC04FED7FBB3" bold="true" box="[343,363,1005,1029]" pageId="22" pageNumber="23">d.</emphasis>
Right middle leg, anterior face;
<emphasis id="7DDDB356FFB5FFFAFD77FC04FD7AFBB3" bold="true" box="[693,710,1005,1029]" pageId="22" pageNumber="23">e.</emphasis>
Gills IVII (from right to left), dorsal view.
<emphasis id="7DDDB356FFB5FFFAFB49FC05FB71FBB2" bold="true" box="[1163,1229,1004,1028]" pageId="22" pageNumber="23">Scales</emphasis>
: a, ce 1mm.
</paragraph>
</caption>
<paragraph id="4F166F44FFB5FFFAFF05FBBFFD9AF94F" blockId="22.[151,1437,1110,2037]" pageId="22" pageNumber="23">
<emphasis id="7DDDB356FFB5FFFAFF05FBBFFE23FBC6" bold="true" box="[199,415,1110,1136]" pageId="22" pageNumber="23">Other specimens:</emphasis>
<materialsCitation id="FFC16519FFB5FFFAFE6FFBBEFC04FB23" collectingDate="2002-09-09" collectingDateMax="2021-08-21" collectingDateMin="2002-09-09" collectionCode="T" collectorName="K. Saito" country="Japan" county="Sapporo-shi" elevation="678" latitude="41.833054" location="Jozankei" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" pageId="22" pageNumber="23" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido">
<collectingCountry id="37BE2FD4FFB5FFFAFE6FFBBEFDB9FBC7" box="[429,517,1111,1137]" name="Japan" pageId="22" pageNumber="23">JAPAN</collectingCountry>
,
<collectingRegion id="8D6DA1A6FFB5FFFAFDD4FBBEFD0CFBC7" box="[534,688,1111,1137]" country="Japan" name="Hokkaido" pageId="22" pageNumber="23">HOKKAIDO</collectingRegion>
, ISHIKARI:
<emphasis id="7DDDB356FFB5FFFAFC91FBBFFCC6FBC6" bold="true" box="[851,890,1110,1136]" pageId="22" pageNumber="23">[1.]</emphasis>
<collectingCounty id="A67717C8FFB5FFFAFC4BFBBEFBAFFBC7" box="[905,1043,1111,1137]" pageId="22" pageNumber="23">Sapporo-shi</collectingCounty>
,
<collectingMunicipality id="AF72F53EFFB5FFFAFBE7FBBEFB1AFBC7" box="[1061,1190,1111,1137]" pageId="22" pageNumber="23">Minami-ku</collectingMunicipality>
,
<location id="4A76399FFFB5FFFAFB7AFBBEFAA1FBC7" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFB7AFBBEFAA1FBC7" box="[1208,1309,1111,1137]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Jozankei" pageId="22" pageNumber="23" stateProvince="Hokkaido">Jozankei</location>
, Toyohira-gawa Riv., mainstream,
<quantity id="8851C2A1FFB5FFFAFE77FB92FDBDFB23" box="[437,513,1147,1173]" metricMagnitude="2" metricUnit="m" metricValue="6.779999999999999" pageId="22" pageNumber="23" unit="m" value="678.0">678 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFB5FFFAFD94FB93FD4EFB23" box="[598,754,1146,1173]" degrees="42" direction="north" minutes="47" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="57.0" value="42.799168">42°47'57.0&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB5FFFAFCC1FB92FC16FB23" box="[771,938,1147,1173]" degrees="141" direction="east" minutes="10" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="14.3" value="141.17064">141°10'14.3&quot;E</geoCoordinate>
),
</materialsCitation>
<materialsCitation id="FFC16519FFB5FFFAFC06FB92FD87FB6B" collectingDate="2002-09-09" collectingDateMax="2021-08-21" collectingDateMin="2002-09-09" collectionCode="T" collectorName="K. Saito" country="Japan" county="Sapporo-shi" elevation="678" latitude="41.833054" location="Jozankei" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" pageId="22" pageNumber="23" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido">
1 mature male nymph,
<date id="3B174984FFB5FFFAFB1EFB93FACCFB23" box="[1244,1392,1146,1173]" pageId="22" pageNumber="23" value="2003-08-15">
<collectingDate id="2B53B06CFFB5FFFAFB1EFB93FACCFB23" box="[1244,1392,1146,1173]" pageId="22" pageNumber="23" value="2003-08-15">15.VIII.2003</collectingDate>
</date>
,
<collectorName id="E25C0A92FFB5FFFAFA43FB92FF68FB0F" pageId="22" pageNumber="23">K. Saito</collectorName>
;
<emphasis id="7DDDB356FFB5FFFAFF23FB77FEB4FB0E" bold="true" box="[225,264,1182,1208]" pageId="22" pageNumber="23">[4.]</emphasis>
<location id="4A76399FFFB5FFFAFED3FB76FE27FB0F" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFED3FB76FE27FB0F" box="[273,411,1183,1209]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Sapporo-shi" pageId="22" pageNumber="23" stateProvince="Hokkaido">Sapporo-shi</location>
,
<location id="4A76399FFFB5FFFAFE6AFB76FD95FB0F" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFE6AFB76FD95FB0F" box="[424,553,1183,1209]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Minami-ku" pageId="22" pageNumber="23" stateProvince="Hokkaido">Minami-ku</location>
,
<location id="4A76399FFFB5FFFAFDF4FB76FD27FB0F" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFDF4FB76FD27FB0F" box="[566,667,1183,1209]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Jozankei" pageId="22" pageNumber="23" stateProvince="Hokkaido">Jozankei</location>
,
<location id="4A76399FFFB5FFFAFD6AFB76FC10FB0F" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFD6AFB76FC10FB0F" box="[680,940,1183,1209]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Usubetsu-gawa Stream" pageId="22" pageNumber="23" stateProvince="Hokkaido">Usubetsu-gawa Stream</location>
(a tributary of Toyohira-gawa Riv.),
<quantity id="8851C2A1FFB5FFFAFA96FB76FA20FB0F" box="[1364,1436,1183,1209]" metricMagnitude="2" metricUnit="m" metricValue="4.72" pageId="22" pageNumber="23" unit="m" value="472.0">472 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFB5FFFAFF1FFB2BFEC5FB6B" box="[221,377,1218,1245]" degrees="42" direction="north" minutes="54" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="33.5" value="42.909306">42°54'33.5&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB5FFFAFE45FB2AFD92FB6B" box="[391,558,1219,1245]" degrees="141" direction="east" minutes="07" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="28.9" value="141.1247">141°07'28.9&quot;E</geoCoordinate>
),
</materialsCitation>
<materialsCitation id="FFC16519FFB5FFFAFD86FB2AFA9EFA93" collectingDate="2002-09-09" collectingDateMax="2021-08-21" collectingDateMin="2002-09-09" collectionCode="T" collectorName="K. Saito" country="Japan" county="Sapporo-shi" elevation="678" latitude="41.833054" location="Jozankei" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" pageId="22" pageNumber="23" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido">
1 mature male nymph and 2 mature female nymphs,
<date id="3B174984FFB5FFFAFB65FB2AFA94FB6B" box="[1191,1320,1219,1245]" pageId="22" pageNumber="23" value="2002-09-09">
<collectingDate id="2B53B06CFFB5FFFAFB65FB2AFA94FB6B" box="[1191,1320,1219,1245]" pageId="22" pageNumber="23" value="2002-09-09">09.IX.2002</collectingDate>
</date>
,
<collectorName id="E25C0A92FFB5FFFAFAF4FB2AFA24FB6B" box="[1334,1432,1219,1245]" pageId="22" pageNumber="23">K. Saito</collectorName>
;
<emphasis id="7DDDB356FFB5FFFAFF55FB0EFF03FAB7" bold="true" box="[151,191,1255,1281]" pageId="22" pageNumber="23">[5.]</emphasis>
<location id="4A76399FFFB5FFFAFF0BFB0EFEEFFAB7" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFF0BFB0EFEEFFAB7" box="[201,339,1255,1281]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Sapporo-shi" pageId="22" pageNumber="23" stateProvince="Hokkaido">Sapporo-shi</location>
,
<location id="4A76399FFFB5FFFAFEA3FB0EFE5EFAB7" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFEA3FB0EFE5EFAB7" box="[353,482,1255,1281]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Minami-ku" pageId="22" pageNumber="23" stateProvince="Hokkaido">Minami-ku</location>
,
<location id="4A76399FFFB5FFFAFE33FB0EFDEAFAB7" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFE33FB0EFDEAFAB7" box="[497,598,1255,1281]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Jozankei" pageId="22" pageNumber="23" stateProvince="Hokkaido">Jozankei</location>
,
<location id="4A76399FFFB5FFFAFDA6FB0EFCF8FAB7" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFDA6FB0EFCF8FAB7" box="[612,836,1255,1281]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Shirai-gawa Stream" pageId="22" pageNumber="23" stateProvince="Hokkaido">Shirai-gawa Stream</location>
(a tributary of Toyohira-gawa Riv.), the confluence of
<location id="4A76399FFFB5FFFAFF7CFAE2FE18FA93" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFF7CFAE2FE18FA93" box="[190,420,1291,1317]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Shirai-gawa Stream" pageId="22" pageNumber="23" stateProvince="Hokkaido">Shirai-gawa Stream</location>
and
<location id="4A76399FFFB5FFFAFE2EFAE2FD42FA93" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFE2EFAE2FD42FA93" box="[492,766,1291,1317]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Migimata-gawa Stream" pageId="22" pageNumber="23" stateProvince="Hokkaido">Migimata-gawa Stream</location>
,
<quantity id="8851C2A1FFB5FFFAFCD0FAE3FCDEFA93" box="[786,866,1290,1317]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="22" pageNumber="23" unit="m" value="450.0">450 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFB5FFFAFC7FFAE3FBE5FA93" box="[957,1113,1290,1317]" degrees="42" direction="north" minutes="59" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="39.6" value="42.994335">42°59'39.6&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB5FFFAFBAFFAE2FAA9FA93" box="[1133,1301,1291,1317]" degrees="141" direction="east" minutes="04" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="42.9" value="141.07858">141°04'42.9&quot;E</geoCoordinate>
),
</materialsCitation>
<materialsCitation id="FFC16519FFB5FFFAFAF3FAE2FEF0FA27" collectingDate="2002-09-09" collectingDateMax="2021-08-21" collectingDateMin="2002-09-09" collectionCode="T" collectorName="K. Saito" country="Japan" county="Sapporo-shi" elevation="678" latitude="41.833054" location="Jozankei" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" pageId="22" pageNumber="23" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido">
3 mature female nymphs,
<date id="3B174984FFB5FFFAFE97FAC6FE56FAFF" box="[341,490,1327,1353]" pageId="22" pageNumber="23" value="2004-08-07">
<collectingDate id="2B53B06CFFB5FFFAFE97FAC6FE56FAFF" box="[341,490,1327,1353]" pageId="22" pageNumber="23" value="2004-08-07">07.VIII.2004</collectingDate>
</date>
,
<collectorName id="E25C0A92FFB5FFFAFE35FAC6FDE4FAFF" box="[503,600,1327,1353]" pageId="22" pageNumber="23">K. Saito</collectorName>
; ditto, 1 mature female nymph,
<date id="3B174984FFB5FFFAFC08FAC6FBF4FAFF" box="[970,1096,1327,1353]" pageId="22" pageNumber="23" value="2019-04-07">
<collectingDate id="2B53B06CFFB5FFFAFC08FAC6FBF4FAFF" box="[970,1096,1327,1353]" pageId="22" pageNumber="23" value="2019-04-07">07.IV.2019</collectingDate>
</date>
,
<collectionCode id="29B8F781FFB5FFFAFB97FAC6FBDBFAFF" box="[1109,1127,1327,1353]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T</collectionCode>
.
<location id="4A76399FFFB5FFFAFBB6FAC6FB44FAFF" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFBB6FAC6FB44FAFF" box="[1140,1272,1327,1353]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Takayanagi" pageId="22" pageNumber="23" stateProvince="Hokkaido">Takayanagi</location>
;
<emphasis id="7DDDB356FFB5FFFAFAC7FAC7FA91FAFE" bold="true" box="[1285,1325,1326,1352]" pageId="22" pageNumber="23">[8.]</emphasis>
<location id="4A76399FFFB5FFFAFAF4FAC6FF06FADB" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFAF4FAC6FF06FADB" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Sapporo-shi" pageId="22" pageNumber="23" stateProvince="Hokkaido">Sapporo-shi</location>
,
<location id="4A76399FFFB5FFFAFF0BFABAFEF6FADB" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFF0BFABAFEF6FADB" box="[201,330,1363,1389]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Minami-ku" pageId="22" pageNumber="23" stateProvince="Hokkaido">Minami-ku</location>
,
<location id="4A76399FFFB5FFFAFE9BFABAFE02FADB" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFE9BFABAFE02FADB" box="[345,446,1363,1389]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Jozankei" pageId="22" pageNumber="23" stateProvince="Hokkaido">Jozankei</location>
,
<location id="4A76399FFFB5FFFAFE0EFABAFD11FADB" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFE0EFABAFD11FADB" box="[460,685,1363,1389]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Shirai-gawa Stream" pageId="22" pageNumber="23" stateProvince="Hokkaido">Shirai-gawa Stream</location>
(a tributary of Toyohira-gawa Riv.),
<quantity id="8851C2A1FFB5FFFAFBA2FABAFB16FADB" box="[1120,1194,1363,1389]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="23" unit="m" value="300.0">300 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFB5FFFAFB39FABBFA2BFADB" box="[1275,1431,1362,1389]" degrees="42" direction="north" minutes="58" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="37.7" value="42.97714">42°58'37.7&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB5FFFAFF55FA9EFE82FA27" box="[151,318,1399,1425]" degrees="141" direction="east" minutes="08" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="16.4" value="141.1379">141°08'16.4&quot;E</geoCoordinate>
),
</materialsCitation>
<materialsCitation id="FFC16519FFB5FFFAFE9AFA9EFB7CFA03" collectingDate="2002-09-09" collectingDateMax="2021-08-21" collectingDateMin="2002-09-09" collectionCode="T" collectorName="K. Saito" country="Japan" county="Sapporo-shi" elevation="678" latitude="41.833054" location="Jozankei" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" pageId="22" pageNumber="23" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido">
1 mature female nymph,
<date id="3B174984FFB5FFFAFD45FA9FFCA0FA27" box="[647,796,1398,1425]" pageId="22" pageNumber="23" value="2005-08-13">
<collectingDate id="2B53B06CFFB5FFFAFD45FA9FFCA0FA27" box="[647,796,1398,1425]" pageId="22" pageNumber="23" value="2005-08-13">13.VIII.2005</collectingDate>
</date>
,
<collectorName id="E25C0A92FFB5FFFAFCEFFA9EFC2DFA27" box="[813,913,1399,1425]" pageId="22" pageNumber="23">K. Saito</collectorName>
;
<emphasis id="7DDDB356FFB5FFFAFC60FA9FFC64FA26" bold="true" box="[930,984,1398,1424]" pageId="22" pageNumber="23">[15.]</emphasis>
<location id="4A76399FFFB5FFFAFC26FA9EFBD2FA27" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFC26FA9EFBD2FA27" box="[996,1134,1399,1425]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Sapporo-shi" pageId="22" pageNumber="23" stateProvince="Hokkaido">Sapporo-shi</location>
,
<location id="4A76399FFFB5FFFAFBBDFA9EFB58FA27" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFBBDFA9EFB58FA27" box="[1151,1252,1399,1425]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Chuo-ku" pageId="22" pageNumber="23" stateProvince="Hokkaido">Chuo-ku</location>
,
<location id="4A76399FFFB5FFFAFB37FA9EFAF2FA27" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFB37FA9EFAF2FA27" box="[1269,1358,1399,1425]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Minami" pageId="22" pageNumber="23" stateProvince="Hokkaido">Minami</location>
13-jo,
<location id="4A76399FFFB5FFFAFF55FA72FF68FA03" LSID="urn:lsid:plazi:treatment:C700DE52FFB7FFF6FF55F9BEFC20FDAF:4A76399FFFB5FFFAFF55FA72FF68FA03" box="[151,212,1435,1461]" country="Japan" county="Sapporo-shi" latitude="41.833054" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" name="Nishi" pageId="22" pageNumber="23" stateProvince="Hokkaido">Nishi</location>
1-chome, Toyohira-gawa Riv., mainstream,
<quantity id="8851C2A1FFB5FFFAFD17FA72FCADFA03" box="[725,785,1435,1461]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="22" pageNumber="23" unit="m" value="28.0">28 m</quantity>
a.s.l. (
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,
<geoCoordinate id="2A9D0983FFB5FFFAFBCEFA72FB0EFA03" box="[1036,1202,1435,1461]" degrees="141" direction="east" minutes="21" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="31.7" value="141.35881">141°21'31.7&quot;E</geoCoordinate>
),
</materialsCitation>
<materialsCitation id="FFC16519FFB5FFFAFB0BFA72FC47F997" collectingDate="2002-09-09" collectingDateMax="2021-08-21" collectingDateMin="2002-09-09" collectionCode="T" collectorName="K. Saito" country="Japan" county="Sapporo-shi" elevation="678" latitude="41.833054" location="Jozankei" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" pageId="22" pageNumber="23" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido">
<specimenCount id="59AFA4CDFFB5FFFAFB0BFA72FA91FA03" box="[1225,1325,1435,1461]" count="1" pageId="22" pageNumber="23" type="female">1 female</specimenCount>
imago, 1 mature male nymph and 3 mature female nymphs,
<date id="3B174984FFB5FFFAFD1CFA56FCCFFA6F" box="[734,883,1471,1497]" pageId="22" pageNumber="23" value="2020-08-19">
<collectingDate id="2B53B06CFFB5FFFAFD1CFA56FCCFFA6F" box="[734,883,1471,1497]" pageId="22" pageNumber="23" value="2020-08-19">19.VIII.2020</collectingDate>
</date>
,
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. Takayanagi; ditto, 1 mature male nymph and 2 mature female nymph,
<date id="3B174984FFB5FFFAFE7FFA0BFD83FA4B" box="[445,575,1506,1533]" pageId="22" pageNumber="23" value="2020-09-25">
<collectingDate id="2B53B06CFFB5FFFAFE7FFA0BFD83FA4B" box="[445,575,1506,1533]" pageId="22" pageNumber="23" value="2020-09-25">25.IX.2020</collectingDate>
</date>
,
<collectionCode id="29B8F781FFB5FFFAFD8FFA0AFDE3FA4B" box="[589,607,1507,1533]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T</collectionCode>
. Takayanagi;
<emphasis id="7DDDB356FFB5FFFAFD3DFA0BFC89FA4A" bold="true" box="[767,821,1506,1532]" pageId="22" pageNumber="23">[16.]</emphasis>
Sapporo-shi, Chuo-ku, Kita 1-jo, Higashi 19-chome, Toyohira-gawa Riv., mainstream,
<quantity id="8851C2A1FFB5FFFAFDD5F9EFFDEDF997" box="[535,593,1542,1569]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="23" unit="m" value="15.0">15 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFB5FFFAFD5CF9EEFC86F997" box="[670,826,1543,1569]" degrees="43" direction="north" minutes="04" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="04.8" value="43.068">43°04'04.8&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB5FFFAFC85F9EEFC51F997" box="[839,1005,1543,1569]" degrees="141" direction="east" minutes="23" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="13.2" value="141.387">141°23'13.2&quot;E</geoCoordinate>
),
</materialsCitation>
<materialsCitation id="FFC16519FFB5FFFAFBC1F9EEFE43F93B" collectingDate="2002-09-09" collectingDateMax="2021-08-21" collectingDateMin="2002-09-09" collectionCode="T" collectorName="K. Saito" country="Japan" county="Sapporo-shi" elevation="678" latitude="41.833054" location="Jozankei" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" pageId="22" pageNumber="23" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido">
2 mature male nymphs,
<date id="3B174984FFB5FFFAFAD4F9EEFA24F997" box="[1302,1432,1543,1569]" pageId="22" pageNumber="23" value="2006-09-02">
<collectingDate id="2B53B06CFFB5FFFAFAD4F9EEFA24F997" box="[1302,1432,1543,1569]" pageId="22" pageNumber="23" value="2006-09-02">02.IX.2006</collectingDate>
</date>
,
<collectorName id="E25C0A92FFB5FFFAFF55F9C2FF4BF9F3" box="[151,247,1579,1605]" pageId="22" pageNumber="23">K. Saito</collectorName>
; ditto,
<specimenCount id="59AFA4CDFFB5FFFAFE86F9C2FE33F9F3" box="[324,399,1579,1605]" count="1" pageId="22" pageNumber="23" type="male">1 male</specimenCount>
imago,
<specimenCount id="59AFA4CDFFB5FFFAFE29F9C2FDF0F9F3" box="[491,588,1579,1605]" count="1" pageId="22" pageNumber="23" type="female">1 female</specimenCount>
imago, 9 mature male nymphs and 7 mature female nymphs,
<date id="3B174984FFB5FFFAFAC1F9C2FA2BF9F3" box="[1283,1431,1579,1605]" pageId="22" pageNumber="23" value="2021-08-21">
<collectingDate id="2B53B06CFFB5FFFAFAC1F9C2FA2BF9F3" box="[1283,1431,1579,1605]" pageId="22" pageNumber="23" value="2021-08-21">21.VIII.2021</collectingDate>
</date>
,
<collectionCode id="29B8F781FFB5FFFAFF55F9A6FF15F9DF" box="[151,169,1615,1641]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T</collectionCode>
. Takayanagi; KAMIKAWA (outside of the study area): Kamikawa-cho, Souunkyo, Ishikari-gawa Riv.,
<quantity id="8851C2A1FFB5FFFAFADCF9A6FADFF9DF" box="[1310,1379,1615,1641]" metricMagnitude="2" metricUnit="m" metricValue="6.11" pageId="22" pageNumber="23" unit="m" value="611.0">611 m</quantity>
a.s.l. (
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,
<geoCoordinate id="2A9D0983FFB5FFFAFE89F99BFE4EF93B" box="[331,498,1650,1677]" degrees="142" direction="east" minutes="56" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="34.5" value="142.94292">142°56'34.5&quot;E</geoCoordinate>
),
</materialsCitation>
<materialsCitation id="FFC16519FFB5FFFAFDCBF99AFD46F907" collectingDate="2002-09-09" collectingDateMax="2021-08-21" collectingDateMin="2002-09-09" collectionCode="T" collectorName="K. Saito" country="Japan" county="Sapporo-shi" elevation="678" latitude="41.833054" location="Jozankei" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" pageId="22" pageNumber="23" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido">
<specimenCount id="59AFA4CDFFB5FFFAFDCBF99AFDE4F93B" box="[521,600,1651,1677]" count="1" pageId="22" pageNumber="23" type="male">1 male</specimenCount>
imago,
<date id="3B174984FFB5FFFAFD78F99AFC86F93B" box="[698,826,1651,1677]" pageId="22" pageNumber="23" value="2020-09-11">
<collectingDate id="2B53B06CFFB5FFFAFD78F99AFC86F93B" box="[698,826,1651,1677]" pageId="22" pageNumber="23" value="2020-09-11">11.IX.2020</collectingDate>
</date>
,
<collectionCode id="29B8F781FFB5FFFAFC8BF99AFCE7F93B" box="[841,859,1651,1677]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T</collectionCode>
. Hirose; Horokanai-cho, Shumarinai, Shumarinaigawa Riv.,
<quantity id="8851C2A1FFB5FFFAFED0F97EFEE4F907" box="[274,344,1687,1713]" metricMagnitude="2" metricUnit="m" metricValue="2.68" pageId="22" pageNumber="23" unit="m" value="268.0">268 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFB5FFFAFE62F97FFD80F907" box="[416,572,1686,1713]" degrees="44" direction="north" minutes="17" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="56.0" value="44.29889">44°17'56.0&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB5FFFAFD84F97EFD52F907" box="[582,750,1687,1713]" degrees="142" direction="east" minutes="09" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="31.0" value="142.1586">142°09'31.0&quot;E</geoCoordinate>
),
</materialsCitation>
<materialsCitation id="FFC16519FFB5FFFAFCC2F97EFD9EF94F" collectingDate="2002-09-09" collectingDateMax="2021-08-21" collectingDateMin="2002-09-09" collectionCode="T" collectorName="K. Saito" country="Japan" county="Sapporo-shi" elevation="678" latitude="41.833054" location="Jozankei" longLatPrecision="1" longitude="140.33267" municipality="Minami-ku" pageId="22" pageNumber="23" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Hokkaido">
<specimenCount id="59AFA4CDFFB5FFFAFCC2F97EFCF6F907" box="[768,842,1687,1713]" count="1" pageId="22" pageNumber="23" type="male">1 male</specimenCount>
imago,
<date id="3B174984FFB5FFFAFC60F97EFB84F907" box="[930,1080,1687,1713]" pageId="22" pageNumber="23" value="2020-08-02">
<collectingDate id="2B53B06CFFB5FFFAFC60F97EFB84F907" box="[930,1080,1687,1713]" pageId="22" pageNumber="23" value="2020-08-02">02.VIII.2020</collectingDate>
</date>
,
<collectionCode id="29B8F781FFB5FFFAFB83F97EFBEFF907" box="[1089,1107,1687,1713]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T</collectionCode>
. Ito; HIYAMA (outside of the study area): Assabu-cho, Shirooka, Sasamodosawa-gawa Riv.,
<quantity id="8851C2A1FFB5FFFAFC94F952FC33F963" box="[854,911,1723,1749]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="22" pageNumber="23" unit="m" value="76.0">76 m</quantity>
a.s.l. (
<geoCoordinate id="2A9D0983FFB5FFFAFC1BF953FBC9F963" box="[985,1141,1722,1749]" degrees="41" direction="north" minutes="49" orientation="latitude" pageId="22" pageNumber="23" precision="1" seconds="59.0" value="41.833054">41°49'59.0&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB5FFFAFB42F953FA9BF963" box="[1152,1319,1722,1749]" degrees="140" direction="east" minutes="19" orientation="longitude" pageId="22" pageNumber="23" precision="1" seconds="57.6" value="140.33267">140°19'57.6&quot;E</geoCoordinate>
),
<specimenCount id="59AFA4CDFFB5FFFAFAF9F952FA20F963" box="[1339,1436,1723,1749]" count="1" pageId="22" pageNumber="23" type="female">1 female</specimenCount>
imago,
<date id="3B174984FFB5FFFAFF29F936FECBF94F" box="[235,375,1759,1785]" pageId="22" pageNumber="23" value="2019-07-08">
<collectingDate id="2B53B06CFFB5FFFAFF29F936FECBF94F" box="[235,375,1759,1785]" pageId="22" pageNumber="23" value="2019-07-08">08.VII.2019</collectingDate>
</date>
,
<collectionCode id="29B8F781FFB5FFFAFE40F936FE28F94F" box="[386,404,1759,1785]" name="Tavera, Department of Geology and Geophysics" pageId="22" pageNumber="23">T</collectionCode>
. Takayanagi
</materialsCitation>
.
</paragraph>
<paragraph id="4F166F44FFB5FFFAFF05F8EAFCDCF8AA" blockId="22.[151,1437,1110,2037]" box="[199,864,1794,1821]" pageId="22" pageNumber="23">
<emphasis id="7DDDB356FFB5FFFAFF05F8EAFCDCF8AA" bold="true" box="[199,864,1794,1821]" pageId="22" pageNumber="23">
Nymph (mature, in ethanol), redescription (
<figureCitation id="D79273C1FFB5FFFAFD12F8EBFCE8F8AA" box="[720,852,1794,1820]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStart-2="FIGURE 17" captionStartId-0="21.[151,250,1829,1853]" captionStartId-1="22.[151,250,968,992]" captionStartId-2="23.[151,250,1882,1906]" captionTargetBox-0="[153,1431,419,1803]" captionTargetBox-1="[154,1433,183,940]" captionTargetBox-2="[159,1435,464,1849]" captionTargetId-0="figure-111@21.[151,1435,415,1806]" captionTargetId-1="figure-20@22.[151,1436,181,944]" captionTargetId-2="figure-165@23.[151,1435,453,1858]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIGURE 15af. Epeorus curvatulus, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." captionText-1="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." captionText-2="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi-0="http://doi.org/10.5281/zenodo.14954017" figureDoi-1="http://doi.org/10.5281/zenodo.14954019" figureDoi-2="http://doi.org/10.5281/zenodo.14954021" httpUri-0="https://zenodo.org/record/14954017/files/figure.png" httpUri-1="https://zenodo.org/record/14954019/files/figure.png" httpUri-2="https://zenodo.org/record/14954021/files/figure.png" pageId="22" pageNumber="23">Figs 1517</figureCitation>
).
</emphasis>
</paragraph>
<paragraph id="4F166F44FFB5FFFAFF05F8CEFABEF8F7" blockId="22.[151,1437,1110,2037]" box="[199,1282,1830,1857]" pageId="22" pageNumber="23">
Body length 10.011.0 mm (male), 10.0
<quantity id="8851C2A1FFB5FFFAFD5AF8CFFD46F8F7" box="[664,762,1830,1857]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="22" pageNumber="23" unit="mm" value="12.5">12.5 mm</quantity>
(female). Cercus length ca. body length x 1.2.
</paragraph>
<paragraph id="4F166F44FFB5FFFBFF05F8A3FAF6FE1B" blockId="22.[151,1437,1110,2037]" lastBlockId="23.[151,1437,150,429]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis id="7DDDB356FFB5FFFAFF05F8A3FEB0F8D3" bold="true" box="[199,268,1866,1893]" italics="true" pageId="22" pageNumber="23">Head.</emphasis>
Shape trapezoidal. Color dark-brown, with white spots. Compound eyes dark-gray. Ocelli dark gray. Antennae brown (
<figureCitation id="D79273C1FFB5FFFAFEA6F886FE5BF83F" box="[356,487,1903,1929]" captionStart="FIGURE 16" captionStartId="22.[151,250,968,992]" captionTargetBox="[154,1433,183,940]" captionTargetId="figure-20@22.[151,1436,181,944]" captionTargetPageId="22" captionText="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954019" httpUri="https://zenodo.org/record/14954019/files/figure.png" pageId="22" pageNumber="23">Figs 16a, c</figureCitation>
). Anterior margin very densely covered with fine medio-dorsally directed hair-like setae extending to lateral margins (
<figureCitation id="D79273C1FFB5FFFAFDECF87AFD28F81B" box="[558,660,1939,1965]" captionStart="FIGURE 16" captionStartId="22.[151,250,968,992]" captionTargetBox="[154,1433,183,940]" captionTargetId="figure-20@22.[151,1436,181,944]" captionTargetPageId="22" captionText="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954019" httpUri="https://zenodo.org/record/14954019/files/figure.png" pageId="22" pageNumber="23">Fig. 16b</figureCitation>
). Dorsal surface covered with fine hair-like setae.
<emphasis id="7DDDB356FFB5FFFAFB23F87AFAF9F81B" bold="true" box="[1249,1349,1939,1965]" pageId="22" pageNumber="23">Labrum</emphasis>
: lateral margins straight and widened anteriorly, anterior margin gently concave, dorsal surface with 810 bristles anteromedially and 2 bristles near each side of anterolateral margin. Ventral surface with a row of bristles along lateral margins extending to anterior margins and fine, long hair-like setae medially (
<figureCitation id="D79273C1FFB4FFFBFBDFFF7EFB3CFF07" box="[1053,1152,151,177]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="23" pageNumber="24">Fig. 17a</figureCitation>
).
<emphasis id="7DDDB356FFB4FFFBFB54FF7FFAA8FF06" bold="true" box="[1174,1300,150,176]" pageId="23" pageNumber="24">Mandibles</emphasis>
: each outer incisor with three apical teeth and serrated margins, inner incisor with two apical teeth (right mandible) or three apical teeth (left mandible) and serrated sharply on outer margin of both incisors, right inner incisor slender and almost straight. Tuft of long setae on the base of inner incisor of right mandible, tuft of long setae and one comb-like seta on the base of inner incisor of left mandible (
<figureCitation id="D79273C1FFB4FFFBFD78FECEFC84FEF7" box="[698,824,295,321]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="23" pageNumber="24">Figs 17b, c</figureCitation>
).
<emphasis id="7DDDB356FFB4FFFBFC8EFECFFC08FEF6" bold="true" box="[844,948,294,320]" pageId="23" pageNumber="24">Maxillae</emphasis>
: one penicillate setae at base of apical tooth complex (
<figureCitation id="D79273C1FFB4FFFBFECBFEA2FED7FED3" box="[265,363,331,357]" captionStart="FIGURE 28" captionStartId="36.[151,250,1106,1130]" captionTargetBox="[154,1433,184,1083]" captionTargetId="figure-20@36.[151,1436,181,1083]" captionTargetPageId="36" captionText="FIGURE 28ae. Setae at the base of apical tooth of maxilla (left half right, right half left). a. Epeorus latifolium; b. E. L-nigrum; c. E. curvatulus; d. E. nipponicus; e. E. aesculus." figureDoi="http://doi.org/10.5281/zenodo.14954045" httpUri="https://zenodo.org/record/14954045/files/figure.png" pageId="23" pageNumber="24">Fig. 28c</figureCitation>
).
<emphasis id="7DDDB356FFB4FFFBFE43FEA2FD9BFED3" bold="true" box="[385,551,331,357]" pageId="23" pageNumber="24">Hypopharynx</emphasis>
: superlinguae distally widened. Lingua shape quadrate.
<emphasis id="7DDDB356FFB4FFFBFB69FEA3FAB6FED2" bold="true" box="[1195,1290,330,356]" pageId="23" pageNumber="24">Labium</emphasis>
: labial palps two-segmented, outer margin of proximal one with sparse-thick setae. Distal segment with sparse, hair-like setae on outer surface and dense, brush-like setae on distal 1/3 area. Glossae and paraglossae with long, dense setae.
</paragraph>
<caption id="1BD63FCCFFB4FFFBFF55F8B3FB81F80F" ID-DOI="http://doi.org/10.5281/zenodo.14954021" ID-Zenodo-Dep="14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1882,1906]" targetBox="[159,1435,464,1849]" targetPageId="23" targetType="figure">
<paragraph id="4F166F44FFB4FFFBFF55F8B3FB81F80F" blockId="23.[151,1436,1882,1978]" pageId="23" pageNumber="24">
<emphasis id="7DDDB356FFB4FFFBFF55F8B3FEFDF8C4" bold="true" box="[151,321,1882,1906]" pageId="23" pageNumber="24">FIGURE 17aj.</emphasis>
<taxonomicName id="88A914C7FFB4FFFBFE85F8B2FDB0F8C4" authorityName="Matsumura" authorityYear="1931" box="[327,524,1882,1906]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFB4FFFBFE85F8B2FDB0F8C4" box="[327,524,1882,1906]" italics="true" pageId="23" pageNumber="24">Epeorus curvatulus</emphasis>
</taxonomicName>
, nymph.
<emphasis id="7DDDB356FFB4FFFBFDA9F8B3FDC2F8C4" bold="true" box="[619,638,1882,1906]" pageId="23" pageNumber="24">a.</emphasis>
Labrum (left half in ventral view, right half in dorsal view);
<emphasis id="7DDDB356FFB4FFFBFB23F8B3FB49F8C4" bold="true" box="[1249,1269,1882,1906]" pageId="23" pageNumber="24">b.</emphasis>
Incisors of right mandible (ventral view);
<emphasis id="7DDDB356FFB4FFFBFE5BF897FE16F820" bold="true" box="[409,426,1918,1942]" pageId="23" pageNumber="24">c.</emphasis>
Incisors of left mandible (ventral view);
<emphasis id="7DDDB356FFB4FFFBFC8FF897FCDEF820" bold="true" box="[845,866,1918,1942]" pageId="23" pageNumber="24">d.</emphasis>
Setae on dorsal surface of femora;
<emphasis id="7DDDB356FFB4FFFBFB09F897FB60F820" bold="true" box="[1227,1244,1918,1942]" pageId="23" pageNumber="24">e.</emphasis>
Gill I;
<emphasis id="7DDDB356FFB4FFFBFAE5F897FA8BF820" bold="true" box="[1319,1335,1918,1942]" pageId="23" pageNumber="24">f.</emphasis>
Gill II;
<emphasis id="7DDDB356FFB4FFFBFA4BF897FA20F820" bold="true" box="[1417,1436,1918,1942]" pageId="23" pageNumber="24">g.</emphasis>
Gill VII;
<emphasis id="7DDDB356FFB4FFFBFF37F84BFEB5F80C" bold="true" box="[245,265,1954,1978]" pageId="23" pageNumber="24">h.</emphasis>
Sternum IX of male;
<emphasis id="7DDDB356FFB4FFFBFE25F84BFE48F80C" bold="true" box="[487,500,1954,1978]" pageId="23" pageNumber="24">i.</emphasis>
Sternum IX of female;
<emphasis id="7DDDB356FFB4FFFBFD24F84BFD48F80C" bold="true" box="[742,756,1954,1978]" pageId="23" pageNumber="24">j.</emphasis>
Posterior margin of tergum VII.
</paragraph>
</caption>
<caption id="1BD63FCCFFBBFFF4FF55FBF1FF6EFBCE" ID-DOI="http://doi.org/10.5281/zenodo.14954023" ID-Zenodo-Dep="14954023" httpUri="https://zenodo.org/record/14954023/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1048,1072]" targetBox="[151,1433,198,1018]" targetPageId="24" targetType="figure">
<paragraph id="4F166F44FFBBFFF4FF55FBF1FF6EFBCE" blockId="24.[151,1436,1048,1144]" pageId="24" pageNumber="25">
<emphasis id="7DDDB356FFBBFFF4FF55FBF1FE87FB86" bold="true" box="[151,315,1048,1072]" pageId="24" pageNumber="25">FIGURE 18af</emphasis>
.
<taxonomicName id="88A914C7FFBBFFF4FE85FBF0FDB0FB86" authorityName="Matsumura" authorityYear="1931" box="[327,524,1048,1072]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFBBFFF4FE85FBF0FDB0FB86" box="[327,524,1048,1072]" italics="true" pageId="24" pageNumber="25">Epeorus curvatulus</emphasis>
</taxonomicName>
, male imago.
<emphasis id="7DDDB356FFBBFFF4FD59FBF1FD12FB86" bold="true" box="[667,686,1048,1072]" pageId="24" pageNumber="25">a.</emphasis>
Habitus, lateral view;
<emphasis id="7DDDB356FFBBFFF4FC51FBF1FC1BFB86" bold="true" box="[915,935,1048,1072]" pageId="24" pageNumber="25">b.</emphasis>
Right hind leg, anterior face;
<emphasis id="7DDDB356FFBBFFF4FB17FBF1FB5AFB86" bold="true" box="[1237,1254,1048,1072]" pageId="24" pageNumber="25">c.</emphasis>
Right middle leg, anterior face;
<emphasis id="7DDDB356FFBBFFF4FEE7FBD5FE85FBE2" bold="true" box="[293,313,1084,1108]" pageId="24" pageNumber="25">d.</emphasis>
Right fore leg, anterior face;
<emphasis id="7DDDB356FFBBFFF4FDABFBD5FDC6FBE2" bold="true" box="[617,634,1084,1108]" pageId="24" pageNumber="25">e.</emphasis>
Base of forewing, dorsal view;
<emphasis id="7DDDB356FFBBFFF4FC00FBD5FC6DFBE2" bold="true" box="[962,977,1084,1108]" pageId="24" pageNumber="25">f.</emphasis>
Genitalia, ventral view.
<emphasis id="7DDDB356FFBBFFF4FB0EFBD5FAB2FBE2" bold="true" box="[1228,1294,1084,1108]" pageId="24" pageNumber="25">Scales</emphasis>
: a 5 mm, bd 1mm.
</paragraph>
</caption>
<caption id="1BD63FCCFFBBFFF4FF55F8D6FE1CF829" ID-DOI="http://doi.org/10.5281/zenodo.14954027" ID-Zenodo-Dep="14954027" httpUri="https://zenodo.org/record/14954027/files/figure.png" pageId="24" pageNumber="25" startId="24.[151,250,1855,1879]" targetBox="[151,1432,1205,1825]" targetPageId="24" targetType="figure">
<paragraph id="4F166F44FFBBFFF4FF55F8D6FE1CF829" blockId="24.[151,1437,1855,1951]" pageId="24" pageNumber="25">
<emphasis id="7DDDB356FFBBFFF4FF55F8D6FEF9F8E1" bold="true" box="[151,325,1855,1879]" pageId="24" pageNumber="25">FIGURE 19ae.</emphasis>
<taxonomicName id="88A914C7FFBBFFF4FE8EF8A9FDAFF8E1" authorityName="Matsumura" authorityYear="1931" box="[332,531,1855,1879]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFBBFFF4FE8EF8A9FDAFF8E1" box="[332,531,1855,1879]" italics="true" pageId="24" pageNumber="25">Epeorus curvatulus</emphasis>
</taxonomicName>
, female imago.
<emphasis id="7DDDB356FFBBFFF4FD7BF8D6FD77F8E1" bold="true" box="[697,715,1855,1879]" pageId="24" pageNumber="25">a.</emphasis>
Habitus, lateral view;
<emphasis id="7DDDB356FFBBFFF4FC77F8D6FC75F8E1" bold="true" box="[949,969,1855,1879]" pageId="24" pageNumber="25">b.</emphasis>
Right hind leg, anterior face;
<emphasis id="7DDDB356FFBBFFF4FB3CF8D6FAB3F8E1" bold="true" box="[1278,1295,1855,1879]" pageId="24" pageNumber="25">c.</emphasis>
Right middle leg, anterior face;
<emphasis id="7DDDB356FFBBFFF4FE8CF88AFEDEF8CD" bold="true" box="[334,354,1891,1915]" pageId="24" pageNumber="25">d.</emphasis>
Right fore leg, anterior face;
<emphasis id="7DDDB356FFBBFFF4FD4EF88AFD21F8CD" bold="true" box="[652,669,1891,1915]" pageId="24" pageNumber="25">e.</emphasis>
Subgenital plate and sternum IX (with ventral part of tergum), ventral view.
<emphasis id="7DDDB356FFBBFFF4FF55F86EFF65F829" bold="true" box="[151,217,1927,1951]" pageId="24" pageNumber="25">Scales</emphasis>
: a 5mm, bd 1mm.
</paragraph>
</caption>
<paragraph id="4F166F44FFBAFFF5FF05FF7FFB3EFED2" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FF7FFE9FFF07" bold="true" box="[199,291,150,177]" italics="true" pageId="25" pageNumber="26">Thorax.</emphasis>
Color dark-brown, with pale markings dorsally (
<figureCitation id="D79273C1FFBAFFF5FCFBFF7EFC08FF07" box="[825,948,150,177]" captionStart="FIGURE 15" captionStartId="21.[151,250,1829,1853]" captionTargetBox="[153,1431,419,1803]" captionTargetId="figure-111@21.[151,1435,415,1806]" captionTargetPageId="21" captionText="FIGURE 15af. Epeorus curvatulus, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." figureDoi="http://doi.org/10.5281/zenodo.14954017" httpUri="https://zenodo.org/record/14954017/files/figure.png" pageId="25" pageNumber="26">Figs 15a, d</figureCitation>
). Ventrally white to grayish-white (
<figureCitation id="D79273C1FFBAFFF5FAF7FF7EFF1AFF63" captionStart="FIGURE 15" captionStartId="21.[151,250,1829,1853]" captionTargetBox="[153,1431,419,1803]" captionTargetId="figure-111@21.[151,1435,415,1806]" captionTargetPageId="21" captionText="FIGURE 15af. Epeorus curvatulus, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." figureDoi="http://doi.org/10.5281/zenodo.14954017" httpUri="https://zenodo.org/record/14954017/files/figure.png" pageId="25" pageNumber="26">Figs 15b, e</figureCitation>
). Pronotum slightly projecting laterally (
<figureCitation id="D79273C1FFBAFFF5FDB2FF52FD6FFF63" box="[624,723,187,213]" captionStart="FIGURE 29" captionStartId="36.[151,250,1557,1581]" captionTargetBox="[152,1428,1237,1525]" captionTargetId="figure-62@36.[151,1436,1227,1533]" captionTargetPageId="36" captionText="FIGURE 29ad. Lateral margin of pronotum, left. a. Epeorus latifolium, and E. L-nigrum; b. E. curvatulus; c. E. nipponicus; d. E. aesculus." figureDoi="http://doi.org/10.5281/zenodo.14954047" httpUri="https://zenodo.org/record/14954047/files/figure.png" pageId="25" pageNumber="26">Fig. 29b</figureCitation>
).
<emphasis id="7DDDB356FFBAFFF5FD25FF52FC9AFF63" bold="true" box="[743,806,187,213]" pageId="25" pageNumber="26">Legs.</emphasis>
Each femur with dark brown markings and hypodermal black spot on its middle part of anterior face (
<figureCitation id="D79273C1FFBAFFF5FD49FF36FD57FF4F" box="[651,747,223,249]" captionStart="FIGURE 16" captionStartId="22.[151,250,968,992]" captionTargetBox="[154,1433,183,940]" captionTargetId="figure-20@22.[151,1436,181,944]" captionTargetPageId="22" captionText="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954019" httpUri="https://zenodo.org/record/14954019/files/figure.png" pageId="25" pageNumber="26">Fig. 16d</figureCitation>
), blade-like long setae on its dorsal edge, short bluntly pointed spines on its ventral edge. Setae on femoral surface variably shaped, most oval to semi-circular (
<figureCitation id="D79273C1FFBAFFF5FB2BFEEBFAF1FEAA" box="[1257,1357,258,284]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="25" pageNumber="26">Fig. 17d</figureCitation>
). Tibia with two dark-brown markings and long hair-like setae on dorsal edge (
<figureCitation id="D79273C1FFBAFFF5FC0BFECFFB90FEF6" box="[969,1068,294,320]" captionStart="FIGURE 16" captionStartId="22.[151,250,968,992]" captionTargetBox="[154,1433,183,940]" captionTargetId="figure-20@22.[151,1436,181,944]" captionTargetPageId="22" captionText="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954019" httpUri="https://zenodo.org/record/14954019/files/figure.png" pageId="25" pageNumber="26">Fig. 16d</figureCitation>
). Tarsus darker in proximal area and with long hair-like setae on its dorsal edge (
<figureCitation id="D79273C1FFBAFFF5FD73FEA3FCAEFED2" box="[689,786,330,356]" captionStart="FIGURE 16" captionStartId="22.[151,250,968,992]" captionTargetBox="[154,1433,183,940]" captionTargetId="figure-20@22.[151,1436,181,944]" captionTargetPageId="22" captionText="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954019" httpUri="https://zenodo.org/record/14954019/files/figure.png" pageId="25" pageNumber="26">Fig. 16d</figureCitation>
). Tarsal claw with 34 denticles.
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FE84FA21FD8D" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FE84FE80FE3E" bold="true" box="[199,316,365,392]" italics="true" pageId="25" pageNumber="26">Abdomen.</emphasis>
Color dark-brown, each tergum with a pair of dark spots (
<figureCitation id="D79273C1FFBAFFF5FC03FE84FBFCFE31" box="[961,1088,365,392]" captionStart="FIGURE 15" captionStartId="21.[151,250,1829,1853]" captionTargetBox="[153,1431,419,1803]" captionTargetId="figure-111@21.[151,1435,415,1806]" captionTargetPageId="21" captionText="FIGURE 15af. Epeorus curvatulus, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." figureDoi="http://doi.org/10.5281/zenodo.14954017" httpUri="https://zenodo.org/record/14954017/files/figure.png" pageId="25" pageNumber="26">Figs 15a, d</figureCitation>
). Sterna white to grayish-white (
<figureCitation id="D79273C1FFBAFFF5FF5DFE78FE9DFE1D" box="[159,289,401,428]" captionStart="FIGURE 15" captionStartId="21.[151,250,1829,1853]" captionTargetBox="[153,1431,419,1803]" captionTargetId="figure-111@21.[151,1435,415,1806]" captionTargetPageId="21" captionText="FIGURE 15af. Epeorus curvatulus, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." figureDoi="http://doi.org/10.5281/zenodo.14954017" httpUri="https://zenodo.org/record/14954017/files/figure.png" pageId="25" pageNumber="26">Figs 15b, e</figureCitation>
). Each tergum with long hair-like setae densely arranged along the midline. Posterior margin of each tergum without long setae and with spines of variable lengths which overlap, forming 23 levels (
<figureCitation id="D79273C1FFBAFFF5FB34FE5CFAEAFE79" box="[1270,1366,437,463]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="25" pageNumber="26">Figs 17j</figureCitation>
,
<figureCitation id="D79273C1FFBAFFF5FAA1FE5CFA2CFE79" box="[1379,1424,437,463]" captionStart="FIGURE 31" captionStartId="38.[151,250,1520,1544]" captionTargetBox="[153,1432,184,1492]" captionTargetId="figure-20@38.[151,1435,181,1495]" captionTargetPageId="38" captionText="FIGURE 31ae. SEM images of abdominal tergum VII (left half). a. Epeorus latifolium; b. E. L-nigrum; c. E. curvatulus; d. E. nipponicus; e. E. aesculus. Scales: ae 50 µm." figureDoi="http://doi.org/10.5281/zenodo.14954051" httpUri="https://zenodo.org/record/14954051/files/figure.png" pageId="25" pageNumber="26">31c</figureCitation>
). Tergum IVII each with three short postero-lateral projections, ventral and lateral ones blunt and dorsal one pointed and bending upward (
<figureCitation id="D79273C1FFBAFFF5FE52FE15FE4EFDA1" box="[400,498,508,535]" captionStart="FIGURE 30" captionStartId="37.[151,250,1300,1324]" captionTargetBox="[163,1425,193,1260]" captionTargetId="figure-19@37.[151,1435,181,1276]" captionTargetPageId="37" captionText="FIGURE 30af. Postero-lateral projections of abdominal tergum IV, dorsal (left) and lateral (right). a. Epeorus latifolium; b. E. L-nigrum; c. E. curvatulus; d. E. nipponicus; e. E. aesculus; f. Correspondence of projections from a different view (dotted lines)." figureDoi="http://doi.org/10.5281/zenodo.14954049" httpUri="https://zenodo.org/record/14954049/files/figure.png" pageId="25" pageNumber="26">Fig. 30c</figureCitation>
). Posterior margin of sternum IX triangular, with a sharp medial emargination and with long hair-like setae from middle to lateral edges (
<figureCitation id="D79273C1FFBAFFF5FD33FDC9FCD5FD8C" box="[753,873,544,570]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="25" pageNumber="26">Figs 17h, i</figureCitation>
). Dorsal surface of cercus with a row of fine setae.
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF55FDADFB95FD7F" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF55FDADFF6AFDE8" bold="true" box="[151,214,580,606]" pageId="25" pageNumber="26">Gills.</emphasis>
Dorsally greyish to brownish, ventrally same (
<figureCitation id="D79273C1FFBAFFF5FD39FDADFCE2FDE8" box="[763,862,580,606]" captionStart="FIGURE 16" captionStartId="22.[151,250,968,992]" captionTargetBox="[154,1433,183,940]" captionTargetId="figure-20@22.[151,1436,181,944]" captionTargetPageId="22" captionText="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954019" httpUri="https://zenodo.org/record/14954019/files/figure.png" pageId="25" pageNumber="26">Fig. 16e</figureCitation>
). Gill I extending anteriorly and meeting beneath mesothorax (
<figureCitation id="D79273C1FFBAFFF5FEE9FD81FE11FD34" box="[299,429,615,642]" captionStart="FIGURE 15" captionStartId="21.[151,250,1829,1853]" captionTargetBox="[153,1431,419,1803]" captionTargetId="figure-111@21.[151,1435,415,1806]" captionTargetPageId="21" captionText="FIGURE 15af. Epeorus curvatulus, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." figureDoi="http://doi.org/10.5281/zenodo.14954017" httpUri="https://zenodo.org/record/14954017/files/figure.png" pageId="25" pageNumber="26">Figs 15b, e</figureCitation>
,
<figureCitation id="D79273C1FFBAFFF5FE7EFD81FE5BFD34" box="[444,487,616,642]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="25" pageNumber="26">17e</figureCitation>
). Posterior tip of Gill VII slightly expanded beneath sternum. Rounded projections at costal-proximal margin of gill plates IIVII (
<figureCitation id="D79273C1FFBAFFF5FD68FD62FC9BFD10" box="[682,807,651,678]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="25" pageNumber="26">Figs 17f, g</figureCitation>
). Costal margin with rough surface. Each gill plate with filaments forming a fan-shape together. Gill VII with longitudinal fold (
<figureCitation id="D79273C1FFBAFFF5FC79FD46FBA1FD7F" box="[955,1053,687,713]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="25" pageNumber="26">
Fig.
<quantity id="8851C2A1FFBAFFF5FC2DFD46FBA1FD7F" box="[1007,1053,687,713]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" pageId="25" pageNumber="26" unit="g" value="17.0">17g</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FD3AFCBFFD5A" blockId="25.[151,1437,150,2037]" box="[199,771,722,749]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FD3AFCBFFD5A" bold="true" box="[199,771,722,749]" pageId="25" pageNumber="26">
Male imago (in ethanol), redescription (
<figureCitation id="D79273C1FFBAFFF5FD62FD3BFD48FD5A" box="[672,756,722,748]" captionStart="FIGURE 18" captionStartId="24.[151,250,1048,1072]" captionTargetBox="[151,1433,198,1018]" captionTargetId="figure-20@24.[151,1436,198,1024]" captionTargetPageId="24" captionText="FIGURE 18af. Epeorus curvatulus, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view. Scales: a 5 mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954023" httpUri="https://zenodo.org/record/14954023/files/figure.png" pageId="25" pageNumber="26">Fig. 18</figureCitation>
).
</emphasis>
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FD1EFB33FCA7" blockId="25.[151,1437,150,2037]" box="[199,1167,758,785]" pageId="25" pageNumber="26">
Body length
<quantity id="8851C2A1FFBAFFF5FE9AFD1EFE03FCA7" box="[344,447,759,785]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="25" pageNumber="26" unit="mm" value="12.6">12.6 mm</quantity>
. Cercus length ca. body length x 2.5. Forewing length
<quantity id="8851C2A1FFBAFFF5FBE1FD1FFB36FCA7" box="[1059,1162,758,785]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="25" pageNumber="26" unit="mm" value="14.5">14.5 mm</quantity>
.
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FCF3FC9CFCEE" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FCF3FEB0FC83" bold="true" box="[199,268,794,821]" italics="true" pageId="25" pageNumber="26">Head.</emphasis>
Yellowish-white. Frons margin with brown border. Frons shape relatively round. Compound eyes dark-gray to black. Ocelli grayish-white with dark-gray border.
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FC88FA20FC16" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FC88FE9FFCCA" bold="true" box="[199,291,865,892]" italics="true" pageId="25" pageNumber="26">Thorax.</emphasis>
Ground color orange to brick-color, internal tissues visible. Pronotum without markings. Mesonotum without markings, darker in scutellum. Mesopleuron with dark markings. Metanotum dark on scutellum (
<figureCitation id="D79273C1FFBAFFF5FAEDFC6CFA33FC16" box="[1327,1423,901,928]" captionStart="FIGURE 18" captionStartId="24.[151,250,1048,1072]" captionTargetBox="[151,1433,198,1018]" captionTargetId="figure-20@24.[151,1436,198,1024]" captionTargetPageId="24" captionText="FIGURE 18af. Epeorus curvatulus, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view. Scales: a 5 mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954023" httpUri="https://zenodo.org/record/14954023/files/figure.png" pageId="25" pageNumber="26">Fig. 18a</figureCitation>
).
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF55FC40FCA6FBE4" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF55FC40FF56FC75" bold="true" box="[151,234,937,963]" pageId="25" pageNumber="26">Wings.</emphasis>
Hyaline, veins white. Pterostigmatic area cloudy. Forewing with black markings on costal brace to the base of radius and media (
<figureCitation id="D79273C1FFBAFFF5FE43FC24FE5CFC51" box="[385,480,973,999]" captionStart="FIGURE 18" captionStartId="24.[151,250,1048,1072]" captionTargetBox="[151,1433,198,1018]" captionTargetId="figure-20@24.[151,1436,198,1024]" captionTargetPageId="24" captionText="FIGURE 18af. Epeorus curvatulus, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view. Scales: a 5 mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954023" httpUri="https://zenodo.org/record/14954023/files/figure.png" pageId="25" pageNumber="26">Fig. 18e</figureCitation>
). Hindwing with black markings on the same part as forewing.
<emphasis id="7DDDB356FFBAFFF5FB5CFC24FB61FC51" bold="true" box="[1182,1245,973,999]" pageId="25" pageNumber="26">Legs.</emphasis>
Each femur with a black spot on the middle part. Fore tibia dark at the tip. Each segment of fore tarsus becoming shorter distally and each joint color dark. Fourth segment shortest on mid and hind tarsus, each joint color dark (
<figureCitation id="D79273C1FFBAFFF5FB7DFBFDFAF8FB98" box="[1215,1348,1044,1070]" captionStart="FIGURE 18" captionStartId="24.[151,250,1048,1072]" captionTargetBox="[151,1433,198,1018]" captionTargetId="figure-20@24.[151,1436,198,1024]" captionTargetPageId="24" captionText="FIGURE 18af. Epeorus curvatulus, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view. Scales: a 5 mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954023" httpUri="https://zenodo.org/record/14954023/files/figure.png" pageId="25" pageNumber="26">Figs 18bd</figureCitation>
). Tarsal claws dissimilar, one hooked-shape and the other oblong.
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FBB2FE81FAB3" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FBB2FE80FBC0" bold="true" box="[199,316,1115,1142]" italics="true" pageId="25" pageNumber="26">Abdomen.</emphasis>
Ground color brownish-white to transparent, posterior margins of segments IIX with dark-brown streaks. Segment VIIIX yellowish white, internal organs and tissues visible (
<figureCitation id="D79273C1FFBAFFF5FC35FB69FBEAFB2C" box="[1015,1110,1152,1178]" captionStart="FIGURE 18" captionStartId="24.[151,250,1048,1072]" captionTargetBox="[151,1433,198,1018]" captionTargetId="figure-20@24.[151,1436,198,1024]" captionTargetPageId="24" captionText="FIGURE 18af. Epeorus curvatulus, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view. Scales: a 5 mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954023" httpUri="https://zenodo.org/record/14954023/files/figure.png" pageId="25" pageNumber="26">Fig. 18a</figureCitation>
). Posterior margin of styliger plate bulging, tip rounded. Projection on forceps base not developed. Forceps four segmented, distal two segments color dark. Penis lobes without spines, with complex concavity, titillators developed (
<figureCitation id="D79273C1FFBAFFF5FB46FB2EFB5AFB57" box="[1156,1254,1223,1249]" captionStart="FIGURE 18" captionStartId="24.[151,250,1048,1072]" captionTargetBox="[151,1433,198,1018]" captionTargetId="figure-20@24.[151,1436,198,1024]" captionTargetPageId="24" captionText="FIGURE 18af. Epeorus curvatulus, male imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Base of forewing, dorsal view; f. Genitalia, ventral view. Scales: a 5 mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954023" httpUri="https://zenodo.org/record/14954023/files/figure.png" pageId="25" pageNumber="26">Fig. 18f</figureCitation>
). Cercus white, basally brown.
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FAE6FD71FA9E" blockId="25.[151,1437,150,2037]" box="[199,717,1294,1321]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FAE6FD71FA9E" bold="true" box="[199,717,1294,1321]" pageId="25" pageNumber="26">
Female (in ethanol), redescription (
<figureCitation id="D79273C1FFBAFFF5FDA8FAE7FD02FA9E" box="[618,702,1294,1320]" captionStart="FIGURE 19" captionStartId="24.[151,250,1855,1879]" captionTargetBox="[151,1432,1205,1825]" captionTargetId="figure-70@24.[151,1435,1205,1831]" captionTargetPageId="24" captionText="FIGURE 19ae. Epeorus curvatulus, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954027" httpUri="https://zenodo.org/record/14954027/files/figure.png" pageId="25" pageNumber="26">Fig. 19</figureCitation>
).
</emphasis>
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FADBFB31FAFB" blockId="25.[151,1437,150,2037]" box="[199,1165,1330,1357]" pageId="25" pageNumber="26">
Body length
<quantity id="8851C2A1FFBAFFF5FE9AFADBFE02FAFB" box="[344,446,1330,1357]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="25" pageNumber="26" unit="mm" value="11.9">11.9 mm</quantity>
. Cercus length ca. body length x 2.4. Forewing length
<quantity id="8851C2A1FFBAFFF5FBE0FADBFB35FAFB" box="[1058,1161,1330,1357]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="25" pageNumber="26" unit="mm" value="13.5">13.5 mm</quantity>
.
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FABFFAEEFAC6" blockId="25.[151,1437,150,2037]" box="[199,1362,1366,1393]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FABFFEB0FAC7" bold="true" box="[199,268,1366,1393]" italics="true" pageId="25" pageNumber="26">Head.</emphasis>
Yellowish-white. Frons triangular. Compound eyes dark-gray to black, grayish-green when alive.
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FA90FC58FA6D" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FA90FE9FFA22" bold="true" box="[199,291,1401,1428]" italics="true" pageId="25" pageNumber="26">Thorax.</emphasis>
Coloration as in male imago (
<figureCitation id="D79273C1FFBAFFF5FDABFA93FD7AFA22" box="[617,710,1402,1428]" captionStart="FIGURE 19" captionStartId="24.[151,250,1855,1879]" captionTargetBox="[151,1432,1205,1825]" captionTargetId="figure-70@24.[151,1435,1205,1831]" captionTargetPageId="24" captionText="FIGURE 19ae. Epeorus curvatulus, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954027" httpUri="https://zenodo.org/record/14954027/files/figure.png" pageId="25" pageNumber="26">Fig. 19a</figureCitation>
).
<emphasis id="7DDDB356FFBAFFF5FD1AFA93FC97FA22" bold="true" box="[728,811,1402,1428]" pageId="25" pageNumber="26">Wings.</emphasis>
Veins dark.Markings as in male imago.
<emphasis id="7DDDB356FFBAFFF5FB23FA93FA9CFA22" bold="true" box="[1249,1312,1402,1428]" pageId="25" pageNumber="26">Legs.</emphasis>
Coloration and markings as in male imago. Fore tarsus not elongated. Fourth segment shortest on each tarsus, other segments almost equal length (
<figureCitation id="D79273C1FFBAFFF5FE43FA28FDB8FA6D" box="[385,516,1473,1499]" captionStart="FIGURE 19" captionStartId="24.[151,250,1855,1879]" captionTargetBox="[151,1432,1205,1825]" captionTargetId="figure-70@24.[151,1435,1205,1831]" captionTargetPageId="24" captionText="FIGURE 19ae. Epeorus curvatulus, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954027" httpUri="https://zenodo.org/record/14954027/files/figure.png" pageId="25" pageNumber="26">Figs 19bd</figureCitation>
). Tarsal claws dissimilar as in male imago.
</paragraph>
<paragraph id="4F166F44FFBAFFF5FF05FA0DFF6CF9F1" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05FA0DFE80FA49" bold="true" box="[199,316,1508,1535]" italics="true" pageId="25" pageNumber="26">Abdomen.</emphasis>
Coloration and markings as in male imago. Reddish eggs visible when alive (
<figureCitation id="D79273C1FFBAFFF5FB7DFA0CFA9DFA49" box="[1215,1313,1509,1535]" captionStart="FIGURE 19" captionStartId="24.[151,250,1855,1879]" captionTargetBox="[151,1432,1205,1825]" captionTargetId="figure-70@24.[151,1435,1205,1831]" captionTargetPageId="24" captionText="FIGURE 19ae. Epeorus curvatulus, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954027" httpUri="https://zenodo.org/record/14954027/files/figure.png" pageId="25" pageNumber="26">Fig. 19a</figureCitation>
). Posterior margin of subgenital plate quadrate, apices rounded. Posterior margin of sternum IX with medial emargination (
<figureCitation id="D79273C1FFBAFFF5FAADF9E0FF7FF9F1" captionStart="FIGURE 19" captionStartId="24.[151,250,1855,1879]" captionTargetBox="[151,1432,1205,1825]" captionTargetId="figure-70@24.[151,1435,1205,1831]" captionTargetPageId="24" captionText="FIGURE 19ae. Epeorus curvatulus, female imago. a. Habitus, lateral view; b. Right hind leg, anterior face; c. Right middle leg, anterior face; d. Right fore leg, anterior face; e. Subgenital plate and sternum IX (with ventral part of tergum), ventral view. Scales: a 5mm, bd 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954027" httpUri="https://zenodo.org/record/14954027/files/figure.png" pageId="25" pageNumber="26">Fig. 19e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07B33CCFFFBAFFF5FF05F9B9FC1FF94C" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="4F166F44FFBAFFF5FF05F9B9FC1FF94C" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05F9B9FDDAF9DD" bold="true" box="[199,614,1616,1643]" pageId="25" pageNumber="26">
Diagnosis and comparison.
<emphasis id="7DDDB356FFBAFFF5FDC9F9B9FDDAF9DD" bold="true" box="[523,614,1616,1643]" italics="true" pageId="25" pageNumber="26">Nymph.</emphasis>
</emphasis>
This species is easily distinguished from other species in the study area by 1) a strongly extended gill I (
<figureCitation id="D79273C1FFBAFFF5FE15F99DFDE8F939" box="[471,596,1652,1679]" captionStart="FIGURE 15" captionStartId="21.[151,250,1829,1853]" captionTargetBox="[153,1431,419,1803]" captionTargetId="figure-111@21.[151,1435,415,1806]" captionTargetPageId="21" captionText="FIGURE 15af. Epeorus curvatulus, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." figureDoi="http://doi.org/10.5281/zenodo.14954017" httpUri="https://zenodo.org/record/14954017/files/figure.png" pageId="25" pageNumber="26">Figs 15b, e</figureCitation>
,
<figureCitation id="D79273C1FFBAFFF5FDA2F99DFD35F938" box="[608,649,1652,1678]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="25" pageNumber="26">17e</figureCitation>
) and 2) a prominent dorsal projection of the postero-lateral projections located on each abdominal tergum (
<figureCitation id="D79273C1FFBAFFF5FDFAF971FD20F904" box="[568,668,1688,1714]" captionStart="FIGURE 30" captionStartId="37.[151,250,1300,1324]" captionTargetBox="[163,1425,193,1260]" captionTargetId="figure-19@37.[151,1435,181,1276]" captionTargetPageId="37" captionText="FIGURE 30af. Postero-lateral projections of abdominal tergum IV, dorsal (left) and lateral (right). a. Epeorus latifolium; b. E. L-nigrum; c. E. curvatulus; d. E. nipponicus; e. E. aesculus; f. Correspondence of projections from a different view (dotted lines)." figureDoi="http://doi.org/10.5281/zenodo.14954049" httpUri="https://zenodo.org/record/14954049/files/figure.png" pageId="25" pageNumber="26">Fig. 30c</figureCitation>
).
<emphasis id="7DDDB356FFBAFFF5FD71F97EFCFAF904" bold="true" box="[691,838,1687,1714]" italics="true" pageId="25" pageNumber="26">Male imago.</emphasis>
This species has black markings at the base of the forewing and has titillators on its penis, similar to
<emphasis id="7DDDB356FFBAFFF5FD0CF955FCE4F963" box="[718,856,1724,1749]" italics="true" pageId="25" pageNumber="26">E. L-nigrum</emphasis>
, but its styliger plate is rounded, and the projection at the base of the forceps is less developed in contrast to
<emphasis id="7DDDB356FFBAFFF5FCD1F909FC20F94F" box="[787,924,1760,1785]" italics="true" pageId="25" pageNumber="26">E. L-nigrum</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="07B33CCFFFBAFFF6FF05F8EAFBAFFE43" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="discussion">
<paragraph id="4F166F44FFBAFFF5FF05F8EAFB24F843" blockId="25.[151,1437,150,2037]" pageId="25" pageNumber="26">
<emphasis id="7DDDB356FFBAFFF5FF05F8EAFE27F8AB" bold="true" box="[199,411,1794,1821]" pageId="25" pageNumber="26">
Remarks.
<emphasis id="7DDDB356FFBAFFF5FE83F8EBFE27F8AB" bold="true" box="[321,411,1794,1821]" italics="true" pageId="25" pageNumber="26">Nymph.</emphasis>
</emphasis>
The specimens identified here as
<taxonomicName id="88A914C7FFBAFFF5FCD2F8EDFC18F8AB" authorityName="Matsumura" authorityYear="1931" box="[784,932,1795,1821]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFBAFFF5FCD2F8EDFC18F8AB" box="[784,932,1795,1821]" italics="true" pageId="25" pageNumber="26">E. curvatulus</emphasis>
</taxonomicName>
agree with the description of
<taxonomicName id="88A914C7FFBAFFF5FB28F8EDFA83F8AB" box="[1258,1343,1795,1821]" class="Insecta" family="Heptageniidae" genus="Iron" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="7DDDB356FFBAFFF5FB28F8EDFAA5F8AB" box="[1258,1305,1796,1821]" italics="true" pageId="25" pageNumber="26">Iron</emphasis>
sp.
</taxonomicName>
? (
<bibRefCitation id="2B3812B5FFBAFFF5FA9CF8EAFF6CF8F7" author="Ueno, M." pageId="25" pageNumber="26" pagination="19 - 63" refId="ref31987" refString="Ueno, M. (1928) Some Japanese mayfly nymphs. Memoirs of the college of science, Kyoto Imperial University, Series B, 4 (1), 19-63." type="journal article" year="1928">Uéno 1928</bibRefCitation>
) which supports
<bibRefCitation id="2B3812B5FFBAFFF5FE53F8CEFDF4F8F7" author="Imanishi, K." box="[401,584,1831,1857]" pageId="25" pageNumber="26" pagination="380 - 394" refId="ref28952" refString="Imanishi, K. (1934) Mayflies from Japanese Torrents IV. Notes on the genus Epeorus. Annotationes Zoologicae Japonense, 14, 380-394." type="journal article" year="1934">Imanishi (1934)</bibRefCitation>
.
<emphasis id="7DDDB356FFBAFFF5FD91F8CFFD5FF8F7" bold="true" box="[595,739,1830,1857]" italics="true" pageId="25" pageNumber="26">Male imago.</emphasis>
The specimens identified here as
<taxonomicName id="88A914C7FFBAFFF5FBA3F8C1FB4AF8F7" authorityName="Matsumura" authorityYear="1931" box="[1121,1270,1831,1857]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFBAFFF5FBA3F8C1FB4AF8F7" box="[1121,1270,1831,1857]" italics="true" pageId="25" pageNumber="26">E. curvatulus</emphasis>
</taxonomicName>
agree with the original description and
<typeStatus id="9012D1E6FFBAFFF5FE6DF8A2FDADF8D3" box="[431,529,1867,1893]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="2B3812B5FFBAFFF5FDE3F8A2FD54F8D3" author="Matsumura, S." box="[545,744,1867,1893]" pageId="25" pageNumber="26" refId="ref30492" refString="Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Toko-shoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura 1931</bibRefCitation>
).
<emphasis id="7DDDB356FFBAFFF5FD3CF8A3FC46F8D3" bold="true" box="[766,1018,1866,1893]" italics="true" pageId="25" pageNumber="26">Subgenus attribution.</emphasis>
<bibRefCitation id="2B3812B5FFBAFFF5FBC6F8A2FB20F8D3" author="Kluge, N. J." box="[1028,1180,1867,1893]" pageId="25" pageNumber="26" refId="ref29818" refString="Kluge, N. J. (2004) The Phylogenetic System of Ephemeroptera. Kluwer Academic Publishers, Dordrecht, 442 pp. https://doi.org/10.1007/978-94-007-0872-3" type="book" year="2004">Kluge (2004)</bibRefCitation>
attributed this species to the subgenus
<taxonomicName id="88A914C7FFBAFFF5FE90F899FE3DF83F" authorityName="Eaton" authorityYear="1883" box="[338,385,1904,1929]" pageId="25" pageNumber="26" rank="subGenus" subGenus="Iron">
<emphasis id="7DDDB356FFBAFFF5FE90F899FE3DF83F" box="[338,385,1904,1929]" italics="true" pageId="25" pageNumber="26">Iron</emphasis>
</taxonomicName>
, but it should be placed in the subgenus
<taxonomicName id="88A914C7FFBAFFF5FC9BF886FC67F83F" authorityName="Kluge" authorityYear="1997" box="[857,987,1903,1929]" pageId="25" pageNumber="26" rank="subGenus" subGenus="Caucasiron">
<emphasis id="7DDDB356FFBAFFF5FC9BF886FC67F83F" box="[857,987,1903,1929]" italics="true" pageId="25" pageNumber="26">Caucasiron</emphasis>
</taxonomicName>
due to the following characteristics: 1) hairs on the anterior margin of the head are directed dorsomedially (nymphs,
<figureCitation id="D79273C1FFBAFFF5FBC3F87AFBDEF81B" box="[1025,1122,1939,1965]" captionStart="FIGURE 16" captionStartId="22.[151,250,968,992]" captionTargetBox="[154,1433,183,940]" captionTargetId="figure-20@22.[151,1436,181,944]" captionTargetPageId="22" captionText="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954019" httpUri="https://zenodo.org/record/14954019/files/figure.png" pageId="25" pageNumber="26">Fig. 16b</figureCitation>
); 2) each of gills IIVII has a well-developed projection (nymphs,
<figureCitation id="D79273C1FFBAFFF5FD84F85EFD15F867" box="[582,681,1975,2001]" captionStart="FIGURE 16" captionStartId="22.[151,250,968,992]" captionTargetBox="[154,1433,183,940]" captionTargetId="figure-20@22.[151,1436,181,944]" captionTargetPageId="22" captionText="FIGURE 16ae. Epeorus curvatulus, mature nymph. a. Male head, dorsal view; b. Frontal margin of male head; c. Female head, dorsal view; d. Right middle leg, anterior face; e. Gills IVII (from right to left), dorsal view. Scales: a, ce 1mm." figureDoi="http://doi.org/10.5281/zenodo.14954019" httpUri="https://zenodo.org/record/14954019/files/figure.png" pageId="25" pageNumber="26">Figs 16e</figureCitation>
,
<figureCitation id="D79273C1FFBAFFF5FD76F85EFD4BF867" box="[692,759,1975,2001]" captionStart="FIGURE 17" captionStartId="23.[151,250,1882,1906]" captionTargetBox="[159,1435,464,1849]" captionTargetId="figure-165@23.[151,1435,453,1858]" captionTargetPageId="23" captionText="FIGURE 17aj. Epeorus curvatulus, nymph. a. Labrum (left half in ventral view, right half in dorsal view); b. Incisors of right mandible (ventral view); c. Incisors of left mandible (ventral view); d. Setae on dorsal surface of femora; e. Gill I; f. Gill II; g. Gill VII; h. Sternum IX of male; i. Sternum IX of female; j. Posterior margin of tergum VII." figureDoi="http://doi.org/10.5281/zenodo.14954021" httpUri="https://zenodo.org/record/14954021/files/figure.png" pageId="25" pageNumber="26">17f, g</figureCitation>
); and 3) gill plates form a “suction disc” (nymphs,
<figureCitation id="D79273C1FFBAFFF5FAF6F85EFF1AF843" captionStart="FIGURE 15" captionStartId="21.[151,250,1829,1853]" captionTargetBox="[153,1431,419,1803]" captionTargetId="figure-111@21.[151,1435,415,1806]" captionTargetPageId="21" captionText="FIGURE 15af. Epeorus curvatulus, mature nymph, habitus. a. Male, dorsal view; b. Male, ventral view; c. Male, lateral view; d. Female, dorsal view; e. Female, ventral view; f. Female, lateral view. Scales: af 2mm." figureDoi="http://doi.org/10.5281/zenodo.14954017" httpUri="https://zenodo.org/record/14954017/files/figure.png" pageId="25" pageNumber="26">Figs 15b, e</figureCitation>
). Additionally, the male imago of this species lacks latero-dorsal spines on its penis lobes.
</paragraph>
<paragraph id="4F166F44FFB9FFF6FF05FF7EFEBEFE1B" blockId="26.[151,1436,151,537]" pageId="26" pageNumber="27">
This species was originally described based on male imaginal specimens collected from the Tama-gawa River in
<collectingRegion id="8D6DA1A6FFB9FFF6FF39FF52FEF9FF63" box="[251,325,187,213]" country="Japan" name="Tokyo" pageId="26" pageNumber="27">Tokyo</collectingRegion>
, Honshu (
<bibRefCitation id="2B3812B5FFB9FFF6FE75FF52FDC7FF63" author="Matsumura, S." box="[439,635,187,213]" pageId="26" pageNumber="27" refId="ref30492" refString="Matsumura, S. (1931) 6000 Illustrated Insects of Japan-Empire. Toko-shoin, Tokyo, 1497 pp. [in Japanese]" type="book" year="1931">Matsumura 1931</bibRefCitation>
;
<bibRefCitation id="2B3812B5FFB9FFF6FD4AFF52FD78FF63" author="Matsumura, S." box="[648,708,187,213]" pageId="26" pageNumber="27" pagination="199 - 200" refId="ref30516" refString="Matsumura, S. (1932) New Genera and New Species Described in &quot; 6000 Illustrated Insects of Japan-Empire &quot;. Insecta matsumurana, 6 (4), 199-200." type="journal article" year="1932">1932</bibRefCitation>
). The
<typeStatus id="9012D1E6FFB9FFF6FCC9FF52FCD1FF63" box="[779,877,187,213]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
is preserved at SEHU, and male imagines of this species can be confidently identified by comparison with the type specimen (
<figureCitation id="D79273C1FFB9FFF6FC36FF36FBF4FF4F" box="[1012,1096,223,249]" captionStart="FIGURE 34" captionStartId="42.[151,250,1297,1321]" captionTargetBox="[153,1432,240,1270]" captionTargetId="figure-43@42.[151,1435,237,1273]" captionTargetPageId="42" captionText="FIGURE 34af. Type specimen of Epeorus curvatulus. a. Holotype of E. curvatulus, dorsal view; b. Ditto, ventral view; c. Ditto, lateral view; d. Abdomen of the specimen (forceps lost), lateral view; e. Genitalia of the specimen, ventral view; f. Labels attached to the specimen. Scales: ac 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14954057" httpUri="https://zenodo.org/record/14954057/files/figure.png" pageId="26" pageNumber="27">Fig. 34</figureCitation>
). Nymphs of this species were identified using DNA barcoding, and their morphological characteristics matched those of Uéno's
<taxonomicName id="88A914C7FFB9FFF6FB27FEEAFA87FEAB" box="[1253,1339,259,285]" class="Insecta" family="Heptageniidae" genus="Iron" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="7DDDB356FFB9FFF6FB27FEEAFAA8FEAA" box="[1253,1300,259,284]" italics="true" pageId="26" pageNumber="27">Iron</emphasis>
sp.
</taxonomicName>
? (
<bibRefCitation id="2B3812B5FFB9FFF6FA9CFEEAFF6FFEF7" author="Ueno, M." pageId="26" pageNumber="27" pagination="19 - 63" refId="ref31987" refString="Ueno, M. (1928) Some Japanese mayfly nymphs. Memoirs of the college of science, Kyoto Imperial University, Series B, 4 (1), 19-63." type="journal article" year="1928">Uéno 1928</bibRefCitation>
). Although
<bibRefCitation id="2B3812B5FFB9FFF6FE94FECEFDB6FEF7" author="Imanishi, K." box="[342,522,295,321]" pageId="26" pageNumber="27" pagination="380 - 394" refId="ref28952" refString="Imanishi, K. (1934) Mayflies from Japanese Torrents IV. Notes on the genus Epeorus. Annotationes Zoologicae Japonense, 14, 380-394." type="journal article" year="1934">Imanishi (1934)</bibRefCitation>
associated Uéno's
<taxonomicName id="88A914C7FFB9FFF6FD1DFECEFC8AFEF7" box="[735,822,295,321]" class="Insecta" family="Heptageniidae" genus="Iron" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="7DDDB356FFB9FFF6FD1DFECEFCB2FEF6" box="[735,782,295,320]" italics="true" pageId="26" pageNumber="27">Iron</emphasis>
sp.
</taxonomicName>
? with
<taxonomicName id="88A914C7FFB9FFF6FC48FECEFC5BFEF6" box="[906,999,295,320]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="7DDDB356FFB9FFF6FC48FECEFC5BFEF6" box="[906,999,295,320]" italics="true" pageId="26" pageNumber="27">E. uenoi</emphasis>
</taxonomicName>
, the male genital structures of the type specimens of
<taxonomicName id="88A914C7FFB9FFF6FEF4FEA2FE72FED3" authorityName="Matsumura" authorityYear="1931" box="[310,462,331,357]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="curvatulus">
<emphasis id="7DDDB356FFB9FFF6FEF4FEA2FE72FED3" box="[310,462,331,357]" italics="true" pageId="26" pageNumber="27">E. curvatulus</emphasis>
</taxonomicName>
(
<figureCitation id="D79273C1FFB9FFF6FE22FEA2FDFDFED3" box="[480,577,331,357]" captionStart="FIGURE 34" captionStartId="42.[151,250,1297,1321]" captionTargetBox="[153,1432,240,1270]" captionTargetId="figure-43@42.[151,1435,237,1273]" captionTargetPageId="42" captionText="FIGURE 34af. Type specimen of Epeorus curvatulus. a. Holotype of E. curvatulus, dorsal view; b. Ditto, ventral view; c. Ditto, lateral view; d. Abdomen of the specimen (forceps lost), lateral view; e. Genitalia of the specimen, ventral view; f. Labels attached to the specimen. Scales: ac 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14954057" httpUri="https://zenodo.org/record/14954057/files/figure.png" pageId="26" pageNumber="27">Fig. 34e</figureCitation>
) and
<taxonomicName id="88A914C7FFB9FFF6FD41FEA2FD5FFED2" box="[643,739,331,356]" class="Insecta" family="Heptageniidae" genus="Epeorus" kingdom="Animalia" order="Ephemeroptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="uenoi">
<emphasis id="7DDDB356FFB9FFF6FD41FEA2FD5FFED2" box="[643,739,331,356]" italics="true" pageId="26" pageNumber="27">E. uenoi</emphasis>
</taxonomicName>
(
<figureCitation id="D79273C1FFB9FFF6FD37FEA2FCEBFED3" box="[757,855,331,357]" captionStart="FIGURE 32" captionStartId="40.[151,249,1073,1097]" captionTargetBox="[154,1434,184,1049]" captionTargetId="figure-20@40.[151,1435,181,1049]" captionTargetPageId="40" captionText="FIGURE 32af. Type specimens of Epeorus uenoi. a. Male imago from the syntypes of E. uenoi, dorsal view; b. Ditto, ventral view; c. Ditto, lateral view; d. Genitalia of the specimen, ventral view; e, f. Labels attached to the specimen. Scales: ac 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14954053" httpUri="https://zenodo.org/record/14954053/files/figure.png" pageId="26" pageNumber="27">Fig. 32d</figureCitation>
) are markedly different, making this identification highly questionable.
<bibRefCitation id="2B3812B5FFB9FFF6FE41FE86FD86FE3F" author="Ishiwata, S." box="[387,570,367,393]" pageId="26" pageNumber="27" pagination="55 - 84" refId="ref29157" refString="Ishiwata, S. (2001 a) A checklist of Japanese Ephemeroptera. In: The 21 st Century and Aquatic Entomology in East Asia. roceedings of the 1 st. Symposium of Aquatic Entomologists in East Asia (AESEA). Korean Society of Aquatic Entomology Korea, Seoul, pp. 55-84." type="book chapter" year="2001">Ishiwata (2001a</bibRefCitation>
;
<bibRefCitation id="2B3812B5FFB9FFF6FD84FE86FD2CFE3F" author="Ishiwata, S." box="[582,656,367,393]" pageId="26" pageNumber="27" pagination="163 - 200" refId="ref29216" refString="Ishiwata, S. (2001 b) Mayflies of Chiba Prefecture, Checklist, Diagnoses and Keys. Journal of the Natural History Museum and Institute, Chiba, 7, 163-200. [in Japanese]" type="journal article" year="2001">2001b</bibRefCitation>
; 2018) suggested there may be misidentifications in Imanishi's series of works.
</paragraph>
<paragraph id="4F166F44FFB9FFF6FF05FE5EFBAFFE43" blockId="26.[151,1436,151,537]" pageId="26" pageNumber="27">
<emphasis id="7DDDB356FFB9FFF6FF05FE5EFD85FE66" bold="true" box="[199,569,438,465]" pageId="26" pageNumber="27">Emergence period and habitat.</emphasis>
Fully mature nymphs and imagines were obtained from July to October. This species was collected from a wide area within the Toyohira-gawa River system.
</paragraph>
</subSubSection>
<subSubSection id="07B33CCFFFB9FFF6FF05FE17FC20FDAF" box="[199,924,510,537]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="4F166F44FFB9FFF6FF05FE17FC20FDAF" blockId="26.[151,1436,151,537]" box="[199,924,510,537]" pageId="26" pageNumber="27">
<emphasis id="7DDDB356FFB9FFF6FF05FE17FEDDFDAE" bold="true" box="[199,353,510,536]" pageId="26" pageNumber="27">Distribution.</emphasis>
<collectingCountry id="37BE2FD4FFB9FFF6FEA5FE16FE1BFDAF" box="[359,423,511,537]" name="Japan" pageId="26" pageNumber="27">Japan</collectingCountry>
(
<collectingRegion id="8D6DA1A6FFB9FFF6FE74FE16FD95FDAF" box="[438,553,511,537]" country="Japan" name="Hokkaido" pageId="26" pageNumber="27">Hokkaido</collectingRegion>
, Honshu, Shikoku, and Kyushu).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -1,54 +1,54 @@
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<mods:number id="AF183223F80434F5BE6DEE5E10D954EC">5594</mods:number>
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<mods:number id="B2C1CFC979A0D41B593F19B627B0BE1C">5594</mods:number>
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<mods:url id="38392A69A956383B069536DC7208B0DA">https://doi.org/10.11646/zootaxa.5594.1.1</mods:url>
<mods:location id="B3FDE9D62D9EDB266E6D289C72793FE1">
<mods:url id="EB1F45DFD11BA5F73837F2BADCF54407">https://doi.org/10.11646/zootaxa.5594.1.1</mods:url>
</mods:location>
<mods:classification id="8A90ED8DE8D0609596F8FF391124F865">journal article</mods:classification>
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<mods:identifier id="1C6FBCBDCB87D065689070194D356F78" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="651546FF736C30AF6F32D7F05EBE5E8B" type="Zenodo-Dep">14953985</mods:identifier>
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<mods:classification id="C4F6AA0DCE12623070B01DAF2FC59DF7">journal article</mods:classification>
<mods:identifier id="849EF064D69FD5D9B85D4248088A41E5" type="DOI">10.11646/zootaxa.5594.1.1</mods:identifier>
<mods:identifier id="79C60CAF8EE6F37A133FA6F5864DB54D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8BA9AEA87932DF89D15C46199B76CF38" type="Zenodo-Dep">14953985</mods:identifier>
<mods:identifier id="C9F873F2ABF305CD9BB8E04C10129261" type="ZooBank">343CB495-974B-44E6-BE81-1D5A552BD777</mods:identifier>
</mods:mods>
<treatment id="C700DE52FFB9FFF2FF55FD5AFF64FD1F" LSID="urn:lsid:plazi:treatment:C700DE52FFB9FFF2FF55FD5AFF64FD1F" httpUri="http://treatment.plazi.org/id/C700DE52FFB9FFF2FF55FD5AFF64FD1F" lastPageId="30" lastPageNumber="31" pageId="26" pageNumber="27">
<treatment id="C700DE52FFB9FFF2FF55FD5AFF64FD1F" ID-DOI="http://doi.org/10.5281/zenodo.14975376" ID-Zenodo-Dep="14975376" LSID="urn:lsid:plazi:treatment:C700DE52FFB9FFF2FF55FD5AFF64FD1F" httpUri="http://treatment.plazi.org/id/C700DE52FFB9FFF2FF55FD5AFF64FD1F" lastPageId="30" lastPageNumber="31" pageId="26" pageNumber="27">
<subSubSection id="07B33CCFFFB9FFF6FF55FD5AFCC8FD7A" box="[151,884,690,717]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="4F166F44FFB9FFF6FF55FD5AFCC8FD7A" blockId="26.[151,884,690,753]" box="[151,884,690,717]" pageId="26" pageNumber="27">
<heading id="145ED828FFB9FFF6FF55FD5AFCC8FD7A" bold="true" box="[151,884,690,717]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
@ -319,18 +319,17 @@ Quan
<subSubSection id="07B33CCFFFB9FFF0FF55FA60FEE1FEAB" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="4F166F44FFB9FFF6FF55FA60FC0AF8BD" blockId="26.[151,1437,1417,1876]" pageId="26" pageNumber="27">
<emphasis id="7DDDB356FFB9FFF6FF55FA60FE31FA15" bold="true" box="[151,397,1417,1443]" pageId="26" pageNumber="27">Materials examined.</emphasis>
<materialsCitation id="FFC16519FFB9FFF6FE57FA60FF4DFA71" location="Kyoto" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Kyoto">
<materialsCitation id="FFC16519FFB9FFF6FE57FA60FDECFA71" location="Kyoto" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Kyoto">
<emphasis id="7DDDB356FFB9FFF6FE57FA60FDE7FA15" bold="true" box="[405,603,1417,1443]" pageId="26" pageNumber="27">
<typeStatus id="9012D1E6FFB9FFF6FE57FA60FE6CFA15" box="[405,464,1417,1443]" pageId="26" pageNumber="27">Type</typeStatus>
specimens:
</emphasis>
We investigated The
<collectingRegion id="8D6DA1A6FFB9FFF6FC97FA63FC27FA12" box="[853,923,1418,1444]" country="Japan" name="Kyoto" pageId="26" pageNumber="27">Kyoto</collectingRegion>
University Museum collection but could not find the
University Museum collection but could not find the type specimen. Probably lost
</materialsCitation>
<typeStatus id="9012D1E6FFB9FFF6FF3FFA47FE91FA7E" box="[253,301,1454,1480]" pageId="26" pageNumber="27">type</typeStatus>
specimen. Probably lost.
<materialsCitation id="FFC16519FFB9FFF6FD9CFA44FC0EF8BD" collectingDate="2002-09-08" collectingDateMax="2021-08-21" collectingDateMin="2002-09-08" collectionCode="T" collectorName="K. Saito &amp; Sapporo-shi &amp; Minami-ku &amp; Jozankei &amp; Shirai-gawa Stream &amp; Migimata-gawa Stream &amp; Chuo-ku &amp; Minami &amp; Nishi &amp; Toyohira-gawa Riv. &amp; Takayanagi" country="Japan" county="Sapporo-shi" elevation="472" latitude="43.068" location="Jozankei" longLatPrecision="1" longitude="141.387" municipality="Minami-ku" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
.
<materialsCitation id="FFC16519FFB9FFF6FD9CFA44FEF0F9B9" collectingDate="2002-09-08" collectingDateMax="2021-08-21" collectingDateMin="2002-09-08" collectionCode="T" collectorName="K. Saito &amp; Sapporo-shi &amp; Minami-ku &amp; Jozankei &amp; Shirai-gawa Stream &amp; Migimata-gawa Stream &amp; Chuo-ku &amp; Minami &amp; Nishi &amp; Toyohira-gawa Riv. &amp; Takayanagi" country="Japan" county="Sapporo-shi" elevation="472" latitude="43.068" location="Jozankei" longLatPrecision="1" longitude="141.387" municipality="Minami-ku" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
<emphasis id="7DDDB356FFB9FFF6FD9CFA44FC8FFA71" bold="true" box="[606,819,1453,1479]" pageId="26" pageNumber="27">
<typeStatus id="9012D1E6FFB9FFF6FD9CFA44FC92FA71" box="[606,814,1453,1479]" isOtherTypeStatus="true" pageId="26" pageNumber="27">Other specimens</typeStatus>
:
@ -348,15 +347,15 @@ specimen. Probably lost.
,
<location id="4A76399FFFB9FFF6FE5EFA38FD1FFA5D" LSID="urn:lsid:plazi:treatment:C700DE52FFB9FFF2FF55FD5AFF64FD1F:4A76399FFFB9FFF6FE5EFA38FD1FFA5D" box="[412,675,1489,1515]" country="Japan" county="Sapporo-shi" latitude="43.068" longLatPrecision="1" longitude="141.387" municipality="Minami-ku" name="Usubetsu-gawa Stream" pageId="26" pageNumber="27" stateProvince="Hokkaido">Usubetsu-gawa Stream</location>
(a tributary of Toyohira-gawa Riv.),
<elevation id="C4848877FFB9FFF6FB9FFA38FB54FA5D" box="[1117,1256,1489,1516]" metricMagnitude="2" metricUnit="m" metricValue="4.72" pageId="26" pageNumber="27" unit="m" value="472.0">
<quantity id="8851C2A1FFB9FFF6FB9FFA38FB14FA5A" box="[1117,1192,1489,1516]" metricMagnitude="2" metricUnit="m" metricValue="4.72" pageId="26" pageNumber="27" unit="m" value="472.0">472 m</quantity>
a.s.l.
</elevation>
(
a.s.l. (
<geoCoordinate id="2A9D0983FFB9FFF6FB39FA38FA2BFA5A" box="[1275,1431,1489,1516]" degrees="42" direction="north" minutes="54" orientation="latitude" pageId="26" pageNumber="27" precision="1" seconds="33.5" value="42.909306">42°54'33.5&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB9FFF6FF55FA1CFE82F9B9" box="[151,318,1525,1551]" degrees="141" direction="east" minutes="07" orientation="longitude" pageId="26" pageNumber="27" precision="1" seconds="28.9" value="141.1247">141°07'28.9&quot;E</geoCoordinate>
), 2 mature male nymphs,
),
</materialsCitation>
<materialsCitation id="FFC16519FFB9FFF6FE91FA1CFD35F9E1" collectingDate="2002-09-08" collectingDateMax="2021-08-21" collectingDateMin="2002-09-08" collectionCode="T" collectorName="K. Saito &amp; Sapporo-shi &amp; Minami-ku &amp; Jozankei &amp; Shirai-gawa Stream &amp; Migimata-gawa Stream &amp; Chuo-ku &amp; Minami &amp; Nishi &amp; Toyohira-gawa Riv. &amp; Takayanagi" country="Japan" county="Sapporo-shi" elevation="472" latitude="43.068" location="Jozankei" longLatPrecision="1" longitude="141.387" municipality="Minami-ku" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
2 mature male nymphs,
<date id="3B174984FFB9FFF6FDA7FA1CFD64F9B9" box="[613,728,1525,1551]" pageId="26" pageNumber="27" value="2002-09-08">
<collectingDate id="2B53B06CFFB9FFF6FDA7FA1CFD64F9B9" box="[613,728,1525,1551]" pageId="26" pageNumber="27" value="2002-09-08">8.IX.2002</collectingDate>
</date>
@ -376,15 +375,15 @@ a.s.l.
and
<collectorName id="E25C0A92FFB9FFF6FB48F9F0FA24F985" box="[1162,1432,1561,1587]" pageId="26" pageNumber="27">Migimata-gawa Stream</collectorName>
,
<elevation id="C4848877FFB9FFF6FF55F9D4FEA0F9E1" box="[151,284,1597,1624]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="26" pageNumber="27" unit="m" value="450.0">
<quantity id="8851C2A1FFB9FFF6FF55F9D4FF63F9EE" box="[151,223,1597,1624]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="26" pageNumber="27" unit="m" value="450.0">450 m</quantity>
a.s.l.
</elevation>
(
a.s.l. (
<geoCoordinate id="2A9D0983FFB9FFF6FEEEF9D4FE74F9EE" box="[300,456,1597,1624]" degrees="42" direction="north" minutes="59" orientation="latitude" pageId="26" pageNumber="27" precision="1" seconds="39.6" value="42.994335">42°59'39.6&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB9FFF6FE17F9D4FDC0F9E1" box="[469,636,1597,1623]" degrees="141" direction="east" minutes="04" orientation="longitude" pageId="26" pageNumber="27" precision="1" seconds="42.9" value="141.07858">141°04'42.9&quot;E</geoCoordinate>
), 1 mature male nymph and 1 mature female nymph,
),
</materialsCitation>
<materialsCitation id="FFC16519FFB9FFF6FD53F9D4FE47F929" collectingDate="2002-09-08" collectingDateMax="2021-08-21" collectingDateMin="2002-09-08" collectionCode="T" collectorName="K. Saito &amp; Sapporo-shi &amp; Minami-ku &amp; Jozankei &amp; Shirai-gawa Stream &amp; Migimata-gawa Stream &amp; Chuo-ku &amp; Minami &amp; Nishi &amp; Toyohira-gawa Riv. &amp; Takayanagi" country="Japan" county="Sapporo-shi" elevation="472" latitude="43.068" location="Jozankei" longLatPrecision="1" longitude="141.387" municipality="Minami-ku" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
1 mature male nymph and 1 mature female nymph,
<date id="3B174984FFB9FFF6FB22F9D4FAC9F9E1" box="[1248,1397,1597,1623]" pageId="26" pageNumber="27" value="2004-08-07">
<collectingDate id="2B53B06CFFB9FFF6FB22F9D4FAC9F9E1" box="[1248,1397,1597,1623]" pageId="26" pageNumber="27" value="2004-08-07">07.VIII.2004</collectingDate>
</date>
@ -402,15 +401,15 @@ a.s.l.
1-chome,
<collectorName id="E25C0A92FFB9FFF6FC58F988FBC2F9CD" box="[922,1150,1633,1659]" pageId="26" pageNumber="27">Toyohira-gawa Riv.</collectorName>
, mainstream,
<elevation id="C4848877FFB9FFF6FAE3F988FA21F9CD" box="[1313,1437,1633,1660]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="26" pageNumber="27" unit="m" value="28.0">
<quantity id="8851C2A1FFB9FFF6FAE3F988FAE2F9CA" box="[1313,1374,1633,1660]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="26" pageNumber="27" unit="m" value="28.0">28 m</quantity>
a.s.l.
</elevation>
(
a.s.l. (
<geoCoordinate id="2A9D0983FFB9FFF6FF5DF96CFE87F916" box="[159,315,1669,1696]" degrees="43" direction="north" minutes="02" orientation="latitude" pageId="26" pageNumber="27" precision="1" seconds="35.3" value="43.043137">43°02'35.3&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB9FFF6FE85F96CFE52F929" box="[327,494,1669,1695]" degrees="141" direction="east" minutes="21" orientation="longitude" pageId="26" pageNumber="27" precision="1" seconds="31.7" value="141.35881">141°21'31.7&quot;E</geoCoordinate>
), 2 mature male nymphs and 4 mature female nymphs,
),
</materialsCitation>
<materialsCitation id="FFC16519FFB9FFF6FDC0F96CFA89F951" collectingDate="2002-09-08" collectingDateMax="2021-08-21" collectingDateMin="2002-09-08" collectionCode="T" collectorName="K. Saito &amp; Sapporo-shi &amp; Minami-ku &amp; Jozankei &amp; Shirai-gawa Stream &amp; Migimata-gawa Stream &amp; Chuo-ku &amp; Minami &amp; Nishi &amp; Toyohira-gawa Riv. &amp; Takayanagi" country="Japan" county="Sapporo-shi" elevation="472" latitude="43.068" location="Jozankei" longLatPrecision="1" longitude="141.387" municipality="Minami-ku" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
2 mature male nymphs and 4 mature female nymphs,
<date id="3B174984FFB9FFF6FB9AF96CFB51F929" box="[1112,1261,1669,1695]" pageId="26" pageNumber="27" value="2020-08-19">
<collectingDate id="2B53B06CFFB9FFF6FB9AF96CFB51F929" box="[1112,1261,1669,1695]" pageId="26" pageNumber="27" value="2020-08-19">19.VIII.2020</collectingDate>
</date>
@ -440,15 +439,15 @@ mature nymph,
1-jo,
<location id="4A76399FFFB9FFF6FEC5F924FEE2F951" LSID="urn:lsid:plazi:treatment:C700DE52FFB9FFF2FF55FD5AFF64FD1F:4A76399FFFB9FFF6FEC5F924FEE2F951" box="[263,350,1741,1767]" country="Japan" county="Sapporo-shi" latitude="43.068" longLatPrecision="1" longitude="141.387" municipality="Minami-ku" name="Higashi" pageId="26" pageNumber="27" stateProvince="Hokkaido">Higashi</location>
19-chome, Toyohira-gawa Riv., mainstream,
<elevation id="C4848877FFB9FFF6FC98F924FC70F951" box="[858,972,1741,1768]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="15.0">
<quantity id="8851C2A1FFB9FFF6FC98F924FC2EF95E" box="[858,914,1741,1768]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="m" value="15.0">15 m</quantity>
a.s.l.
</elevation>
(
a.s.l. (
<geoCoordinate id="2A9D0983FFB9FFF6FC18F924FBCAF951" box="[986,1142,1741,1767]" degrees="43" direction="north" minutes="04" orientation="latitude" pageId="26" pageNumber="27" precision="1" seconds="04.8" value="43.068">43°04'04.8&quot;N</geoCoordinate>
,
<geoCoordinate id="2A9D0983FFB9FFF6FB43F924FA9BF951" box="[1153,1319,1741,1767]" degrees="141" direction="east" minutes="23" orientation="longitude" pageId="26" pageNumber="27" precision="1" seconds="13.2" value="141.387">141°23'13.2&quot;E</geoCoordinate>
), 1 mature male nymph and 1 mature female nymph,
),
</materialsCitation>
<materialsCitation id="FFC16519FFB9FFF6FAF9F924FC0EF8BD" collectingDate="2002-09-08" collectingDateMax="2021-08-21" collectingDateMin="2002-09-08" collectionCode="T" collectorName="K. Saito &amp; Sapporo-shi &amp; Minami-ku &amp; Jozankei &amp; Shirai-gawa Stream &amp; Migimata-gawa Stream &amp; Chuo-ku &amp; Minami &amp; Nishi &amp; Toyohira-gawa Riv. &amp; Takayanagi" country="Japan" county="Sapporo-shi" elevation="472" latitude="43.068" location="Jozankei" longLatPrecision="1" longitude="141.387" municipality="Minami-ku" pageId="26" pageNumber="27" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hokkaido">
1 mature male nymph and 1 mature female nymph,
<date id="3B174984FFB9FFF6FDB0F918FCBAF8BD" box="[626,774,1777,1803]" pageId="26" pageNumber="27" value="2021-08-21">
<collectingDate id="2B53B06CFFB9FFF6FDB0F918FCBAF8BD" box="[626,774,1777,1803]" pageId="26" pageNumber="27" value="2021-08-21">21.VIII.2021</collectingDate>
</date>

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