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The morphologically most similar species are + +O. lutraria + +, + +O. paralutraria + +sp. nov. +, + +O. sarsii + +and + +O. christiani + +sp. nov. +It differs from these four species by its carapace ornamentation, which features finer reticulations and intermittent, nodulous, anastomosing lirae from about mid-carapace instead of reticulations. The apex of the female rostrum of + +O. lutraria + +and + +O. paralutraria + +is drawn out into an elongate tip. No other species of + +Ozestheria + +features the wing-like flange antero-dorsally on the male rostrum. + + + + + +Type material + + + + +Syntypes + + + + +AUSTRALIA +– + +Western Australia + +• +2 ♀♀ +; Lakeside, Boulder City; + +9 Aug. 1913 + +; +W.B. Alexander +leg.; +WAM 7730 + +. + + + +Other material examined + + + + +AUSTRALIA +– + +Western Australia + +• +3 ♂♂ +; +Lake Carnegie +, +Toonil Pool +; +26°10′27.3″ S +, +122°56′16.9″ E +, + +8 Jun. 2020 + +; +D.J. Cale +leg.; +WAM +C78011 +, +C80198 +, +C80199 + +• + +1 ♂ +; +Lake Carnegie +, +Toonil Pool +; +26°10′27.3″ S +, +122°56′16.9″ E +, + +8 Jun. 2020 + +; +D.J. Cale +leg.; NHMW-ZOO-CR-28494 + +. + + + +Type locality + + + +Western Australia +, Lakeside, Boulder City. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 42a +). Length +10.6–11.3 mm +(HT: +10.6 mm +), height 5.9–6.2 (HT: +5.9 mm +). Coloration reddish-orange, crowded growth bands lighter. 35–46 (HT: 46) growth lines 18–24 (HT: 24) widely spaced and 16–23 (HT: 22) crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, distinct dorso-posterior corner. Posterior margin broadly rounded, suboval, equicurvate (b/H 0.45–0.49, HT: 0.46). Mid-section of ventral margin nearly straight, second growth phase widely rounded. Umbo position submedian (Cr/L 0.29–0.30, HT: 0.30). + + +CARAPACE +ORNAMENTATION +( +Fig. 42f–i +). Larval valve smooth (probably due to abrasion). In dorsal part of carapace, growth bands with strongly reticulating, net-like lirae. In following growth bands and the remainder of non-crowded growth bands, lirae less strongly reticulating, but still anastomosing; lirae intermittent (particularly dorsally within growth bands). Crowded growth bands too narrow to show ornamentation (granular, nodular under SEM). Concentric ridges slightly raised, broad, with moniliform nodules on the dorsal margin. Setae short and thin, rarely preserved (setal pores in one row along all growth lines under SEM). + + +HEAD +( +Fig. 42j +). Condyle short, rounded; occipital notch wide. Condyle lacking anterobasal hump. Margin between condyle and ocular tubercle straight. Ocular tubercle weakly to well developed, forming obtuse angle (~110°) with rostrum. Anterior margin of rostrum strongly convex, antero-dorsally with small, wing-like flange. Apex protruding, pointed, weakly rounded, nearly rectangular. Ventral margin of rostrum weakly concave to straight, with small anterior notch. Naupliar eye small, roundish. Antenna I long with 16–19 lobes (HT: 19), reaching to antenna II flagellomeres VIII–X (HT: IX). Antenna II with 14–15 flagellomeres (HT: 14). + + + +Fig. 42. + +Ozestheria rufa +( +Dakin, 1914 +) + +. +a +. Carapace, male (WAM C78011). +b +. Carapace, female, syntype (WAM 7730). +c +. Carapace, dorsal view (left valve only) female syntype (WAM 7730). +d–e +. Carapace ornamentation (female, syntype, WAM 7730; positions marked in b by rectangles). +d +. Dorsal carapace. +e +. Mid-carapace. +f +. Carapace male (paratype WAM C780119), SEM. +g–i +. Carapace ornamentation (male, WAM C78011; positions marked in f by rectangles), SEM. +g +. Mid-dorsal carapace. +h +. Mid-carapace. +i +. Ventral carapace. +j–k +. Head (antennae not shown). +j +. Male (WAM C78011). +k +. Female, syntype (WAM 7730). +l–m +. Telson. +l +. Male (WAM C78011). +m +. Female, syntype (WAM 7730). +n +. Male, third left thoracopod (WAM C78011). +o +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–c, f= 1 mm; d–e, g–i=0.1 mm; j–n = 0.5 mm. + + + +THORAX +. 25–26 (HT: 26) segments, 23–24 (HT: 24) thoracopod-bearing and two posterior limbless segments not reaching dorsal margin. Only mid-body thoracopod-bearing segments with short spines or setae on dorsal extensions; posterior segments without dorsal spines or setae. + + +THORACOPOD +III (only WAM C78011; +Fig. 42n +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp two-segmented, basal segment shorter than endopod. Exopod ventral extension subequal in extension to endopod, dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( +Fig. 42l +). 6–20 spines (HT: 20). First (anterior) spine enlarged. Spines very small, conical, variable in size and spacing. Posterior-most spine aciculate. Dorsal margin straight to weakly concave. Right terminal claw more strongly curved than left. + + +FURCA +( +Fig. 42l +). Proximally with dorsomedial longitudinal row of 6–15 (HT: 11) setae, row ending distally in a single conical spine. Distal part ⅓–½ of furcal length, with numerous small denticles. + + +Females +( +syntypes +) + + +CARAPACE +( +Fig. 42b–c +). Length +8.4–8.7 mm +, height +4.5–4.6 mm +. Coloration dark brown, nearly black. 18–19, of these 16 widely spaced and 2–3 crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, dorso-posterior corner distinct. Posterior margin oval, greatly extending posteriorly, supracurvate (0.39–0.42). Ventral margin nearly straight. Umbo well developed, extending above dorsal margin, position submedian (0.28–0.29). + + +CARAPACE +ORNAMENTATION +. Larval valve and several following growth bands with irregular, nodulous, medium reticulation (mainly comprising pentagons, hexagons or heptagons). From about mid-carapace ornamentation transitioning to highly anastomosing nodulous lirae; lirae appear intermittent, resulting in granular appearance. Lirae become more pronounced ventrally and posteriorly on carapace. Concentric ridges well developed, raised and broad. No setae visible. + + +HEAD +( +Fig. 42k +). Condyle rounded, short, only weakly protruding; occipital notch wide. Condyle lacking anterobasal hump. Margin between condyle and ocular tubercle straight. Ocular tubercle weakly (?) developed, forming obtuse (~–120°) angle with rostrum. Anterior margin of rostrum dorsally convex, otherwise straight. Apex pointed or slightly rounded (not drawn out), with acute angle (~70°). Ventral margin of rostrum straight. + + +THORAX +. Last few segments without dorsal extensions or spines. + + +TELSON +( + +Fig. +42m + +). 6–9 spines. First (anterior) spine enlarged. All spines rather small. Anterior spines, conical, subequally and widely spaced; posterior spines slightly thinner, more drawn out and aciculate. Dorsal margin straight, last ~¼ concave. Left and right terminal claws equally curved. + + +FURCA +( + +Fig. +42m + +). Distal part ⅓–½ of furcal length, with numerous small denticles (number of setae cannot be determined as individuals were dried out). + + +Distribution +( +Fig. 42o +) + + + +Ozestheria rufa + +is known from central and southern +Western Australia +. + + + + + +Remarks + + + +The +two female +syntypes +are in rather poor condition and appear to have dried out at some point in the past. For this reason, many taxonomically relevant characteristics (e.g., number of body or antennal segments, head shape) could not be assessed. Furthermore, both individuals have been removed from their carapaces, but the bodies and carapace halves were stored together, making it impossible to assign the respective carapace to each body. The latter is not very problematic as they are of similar size and morphologically highly similar. The carapace ornamentation could not be studied in detail due to firmly attached dirt, which could not be removed by strong and continuous sonification. + + +The original description of + +O. rufa + +by +Dakin (1914) +was brief and poorly illustrated. The carapace appeared much narrower and thus more oval. The head lacked the occipital notch (giving it a widely rounded appearance) and the rostrum was probably shown from antero-lateral, giving it a more pointed impression. Because all +syntypes +are female, their morphological features were described in detail and not abbreviated as for the females of other species. + + +We first hesitated to assign the genetically studied males (provisionally termed +O +. sp. X10) to + +O. rufa + +. In the geometric morphometric analyses of carapace shape ( +Fig. 5 +), + +O. rufa + +comb. nov. +appears distinct from all other species and does not overlap with +O +. sp. X10 when individuals are plotted on the PC1– PC2 plane; however, they fully overlap on the PC2–PC3 plane (Supp. file 1_2.2). + +Ozestheria rufa + +was classified as +O +. sp. X10 (probability 100%), but the associated typicality score (0.04) was low. The apparent differences could also be an artifact of the overall low number of specimens in conjunction with sexual dimorphism. As the historic +syntypes +are both poorly preserved females and the freshly collected material all males, a direct comparison of soft body features (e.g., rostrum shape) was not possible. The caparace ornamentation, however, is highly similar. Together with the overall similarity in carapace shape, we decided that the more conservative approach to treat these as a single species is preferrable. More detailed future studies with more specimens and a better representation of males and females might clarify their species status. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FF7BFF0D1749FA37FBD9F831.xml b/data/03/FD/A6/03FDA650FF7BFF0D1749FA37FBD9F831.xml new file mode 100644 index 00000000000..a3129d402d5 --- /dev/null +++ b/data/03/FD/A6/03FDA650FF7BFF0D1749FA37FBD9F831.xml @@ -0,0 +1,540 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria timmsi + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +F9FC30F8-FA27-44BB-9E09-4DB92E842F6E + + + + + +Fig. 47 + + + +Ozestheria +sp. H + +1 – + +Schwentner +et al +. 2015a + +: figs 2, 6. + + + +Ozestheria +sp. H + +– + +Schwentner +et al +. 2020 + +: figs 1–2. + + + + + +Diagnosis + + + + +Ozestheria timmsi + +sp. nov. +is characterized by an elongate, but strongly rounded condyle and a narrow occipital notch; carapace ornamentation dominated by large, pit-like punctae, in later growth bands inconspicuous lirae forming between punctae (lirae mainly posteriorly and ventrally on carapace); male rostrum with strongly convex anterior margin, apex rounded, nearly rectangular (~90–100°), ventral margin convexly curved or straight; female rostrum with weakly convex (nearly straight) anterior margin, apex rectangular and usually rounded, ventral margin slightly convex or s-shaped; 11–14 (male) or 12–14 (female) antenna I lobes reaching to antenna II flagellomeres VII–X (male) or III–VI (female); 16–18 (male) or 15–22 (female) antenna II flagellomeres; 23–24 complete thorax segments; 17–31 telsonic spines, spines small with 3–4 larger spines interspersed, anteriorly broad, conical and posteriorly thin and aciculate; 6–10 furcal setae. + + + +Differential diagnosis + + + + +Ozestheria timmsi + +sp. nov. +can be differentiated from all other species of + +Ozestheria + +by the combination of the carapace ornamentation (dominated by pit-like punctae), the carapace shape, size and coloration (generally dark) as well as the rostrum shape and telson spination. Species with similar carapace ornamentation are + +O. frederikeae + +sp. nov. +and + +O. carnegiensis + +sp. nov. +, which both have shorter carapace lengths (reaching only up to +7 mm +) and fewer complete thorax segments (≤ 21 vs ≥ 23). Furthermore, + +O. carnegiensis + +has fewer antenna flagellomeres (≤ 14 vs ≥ 15) and fewer telsonic spines (11–20 vs 17– 31). Female + +O. frederikeae + +have an undulating anterior rostrum margin and fewer antenna flagellomeres (14–17 vs 15–22). + + + + + +Etymology + + +The species is named in honor of Brian V. Timms. Brian has worked extensively on the ecology and taxonomy of Australia’s large branchiopods and his work has transformed our understanding of their diversity. He also collected the vast majority of the material studied herein; without his relentless efforts and dedication this study would not have been possible. + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +Queensland + +• +♂ +; +Lake Louisa +;, +19°53′46.7″ S +, +144°15′57.4″ E +; + +7 Apr. 2009 + +; +M. Schwentner +and +B.V. Timms +leg.; +GenBank +no: +KJ705772 +( +COI +); +AM +P.91544. + + + + +Paratype + + + + +AUSTRALIA +– + +Queensland + +• +1 ♀ +; same data as for holotype; +GenBank +no: +KJ705761 +( +COI +); +AM +P.91533 + +• + +3 ♂♂ +, +7 ♀♀ +; same data as for holotype; + +12 Apr. 2018 + +; NHMW-ZOO-CR-26637 + +. + + + +Other material examined + + + + +AUSTRALIA +– + +Queensland + +• +2 ♂♂ +, +2 ♀♀ +; +Pelican Lake +; +19°53′46.7″ S +, +144°15′57.4″ E +; + +24 Jun. 2020 + +; +B.V. Timms +leg.; NHMW-ZOO-CR-26638, NHMW-ZOO-CR-26639 + +• + +1 ♂ +; +Salt Bore Lake +; +19°51′22.3″ S +, +144°16′08.5″ E +; + +6 Apr. 2009 + +; +M. Schwentner +and +B.V. Timms +leg.; +AM +P.91532 + +• + +4 ♀♀ +; +Salt Bore Lake +; +19°51′22.3″ S +, +144°16′08.5″ E +; + +24 Jun. 2020 + +; +B.V. Timms +leg.; NHMW-ZOO- CR-26640, NHMW-ZOO-CR-26641 + +. + + + +Type locality + + + +Queensland +. Lake Louisa, +19°53′46.7″ S +, +144°15′57.4″ E +. + + + + + +Description + + + +Males +CARAPACE +( +Fig. 47a, c–e +). Length +8.3–9.6 mm +(HT: +9.6 mm +), height +5.1–5.8 mm +(HT: +5.8 mm +). Coloration light to dark reddish-brown or nearly black, outer margin lighter. 36 –50 (HT: 50) growth lines, of these 28–35 widely spaced (HT: 35) and 4–15 (HT: 15) crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, dorso-posterior corner rounded. Posterior margin widely oval, equicurvate (b/H 0.47–0.50, HT: 0.48). Ventral margin rounded, strongly and uniformly curved. Umbo position anterior to submedian (Cr/L 0.22–0.27, HT: 0.26). + + +CARAPACE +ORNAMENTATION +( +Fig. 47f–i +). All growth bands densely punctate (larval valve and first few growth bands appear to be smooth in several individuals, but that is probably due to abrasion), punctae large and pit-like or net-like. From about mid-carapace, inconspicuous lirae forming between punctae ventrally on growth bands, lirae stronger defined ventrally and posteriorly on the carapace. Crowded growth bands with less conspicuous punctae, space between punctae slightly raised, lirae-like (especially posteriorly). Concentric ridges slightly raised and smooth. Setae filiform, preserved usually only on outer growth lines (under SEM all growth lines with single row of setal pores). + + + +Fig. 47. + +Ozestheria timmsi + +sp. nov. +a–e +. Carapace. +a +. Male, holotype (P.91544). +b +. Female, paratype (P.91533). +c +. Male, holotype, dorsal view (only left valve shown; P.91544). +d +. Male, antero-dorsal part of carapace (P.91532), SEM. +e +. Male, postero-ventral part of carapace (P.91532), SEM. +f–h +. Carapace ornamentation of male (P.91532; positions marked in d, e by rectangles), SEM. +f +. Mid-carapace. +g +. Posterior carapace. +h +. Ventral carapace. +i +. Mid-dorsal carapace ornamentation of male, holotype (P.91544; position marked in a by rectangle). +j–k +. Head (antennae not shown). +j +. Male, holotype (AM P:91544). +k +. Female, paratype (P.91533). +l–m +. Telson. +l +. Male (P.91532; posteriorly damaged), SEM. +m +. Male, holotype (P.91544). +n +. Male, holotype third right thoracopod (P.91544). +o +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–e, j–k, m–n =0.5 mm; f–i =0.1 mm; l =0.2 mm. + + + +HEAD +( +Fig. 47j +). Condyle long, distally rounded; occipital notch narrow. Condyle with weakly developed anterobasal hump. Margin between condyle and ocular tubercle straight. Ocular tubercle weakly developed, forming obtuse to straight angle (~100–180°) angle with rostrum. Anterior margin of rostrum strongly convex. Ventral margin of rostrum convexly curved or straight, no anterior notch; apex rounded, nearly rectangular (~90–100°). Naupliar eye elongated, subrectangular to subtriangular with rounded edges. Antenna I long with 11–14 lobes (HT: 11), reaching to antenna II flagellomeres VII–IX (HT: VIII). Antenna II with 16–18 flagellomeres (HT: 18). + + +THORAX +. 24–25 (HT: 25) segments, 23–24 (HT: 23) thoracopod-bearing and 1–2 (HT: one) posterior limbless segment not reaching dorsal margin. Dorsal armature increasing in size posteriorly over successive segments, central spines long and thin, laterally row of smaller spines. In posteriormost segments central spines stouter and shorter. + + +THORACOPOD +III (only P.91533; +Fig. 47n +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp one-segmented. Exopod ventral extension subequal in extension to endopod, dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( +Fig. 47l–m +). 17–29 spines (HT: 29). First (anterior) spine enlarged. Spines subequal in length, small and thin.Anterior spines conical, posterior spines becoming thinner and more drawn out (aciculate) and more densely spaced. 3–4 larger spines (~2 × in size) interspersed, one about half-length of telson. Dorsal margin straight to slightly concave. Right terminal claw more strongly curved than left. + + +FURCA +( +Fig. 47l–m +). Proximally with dorsomedial longitudinal row of 8–10 (HT: damaged) setae, row ending distally in a single conical spine. Distal part ⅔ of furcal length, with numerous small denticles. + + +Females + + +Overall appearance as in males. Carapace ( +Fig. 47b +) length +7.9–11.5 mm +(mean: +8.7 mm +), height 5.0– +7.6 mm +(mean: +5.5 mm +); 36–45 (41) growth lines, 23–25 (mean: 29) widely spaced and 5–20 (mean: 13) crowded; Cr/L 0.21–0.26 (mean: 0.24) and b/H 0.46–0.53 (mean: 0.50). Anterior margin of rostrum only weakly convex (nearly straight), apex more rectangular than in males but usually rounded. Ventral margin slightly convex or s-shaped. Angle between eye bulge and rostrum widely obtuse, nearly straight ( +Fig. 47k +). Antenna I short with 12–14 (mean: 13) small, well-defined lobes, lobes smaller than in males; reaching to antenna II flagellomeres III–VI (mean: IV). Antenna II with 15–22 flagellomeres (mean: 18). 24–25 (mean: 25) segments, 23–24 (mean: 24) thoracopod-bearing and one to two posterior limbless segments not reaching dorsal margin. Telson with 24–31 (mean: 27) dorsal spines; up to 3–5 larger spines interspersed. Furca with 6–10 setae. + + +Distribution +( +Fig. 47o +) + + + +Ozestheria timmsi + +sp. nov. +is known only from three localities in northern +Queensland +. It is the only species of + +Ozestheria + +recorded so close to the tropical climate zone. + + + + + +Remarks + + + +The carapace shape of + +Ozestheria timmsi + +sp. nov. +( +Fig. 6 +) is distinct from that of most other species and overlaps with those of + +O. frederikeae + +sp. nov. +(only marginally), + +O. sivesae + +sp. nov. +, + +O. setifera + +sp. nov. +, + +O. mariae + +, + +O. gemina + +sp. nov. +, + +O. jonnae + +sp. nov. +(only marginally), + +O. marthae + +sp. nov. +, + +O. weeksi + +sp. nov. +, + +O. cancellata + +comb. nov. +, + +O. quinlanae + +sp. nov. +, + +O. pilbarensis + +sp. nov. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FF7CFF00174EFD3DFDD2FA4D.xml b/data/03/FD/A6/03FDA650FF7CFF00174EFD3DFDD2FA4D.xml new file mode 100644 index 00000000000..1223d5ba87c --- /dev/null +++ b/data/03/FD/A6/03FDA650FF7CFF00174EFD3DFDD2FA4D.xml @@ -0,0 +1,510 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria sivesae + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +E86EC450-37BD-41D3-B398-2AC8A2F38177 + + + + + +Fig. 46 + + + +Ozestheria +sp. I + +– + +Schwentner +et al +. 2015a + +: figs 2, 6; 2020: figs 1–2. + + + + + +Diagnosis + + + + +Ozestheria sivesae + +sp. nov. +is characterized by a long condyle and a narrow occipital notch; a broadly rounded ventral carapace margin; carapace ornamentation punctate dorsally on carapace, from mid-dorsal carapace smooth band dorsally within growth bands followed by punctae and shallow lirae (smooth bands become more pronounced in following growth bands); male rostrum with straight anterior margin, apex rounded with acute angle (close to rectangular), ventral margin concave to nearly straight; female rostrum anterior margin weakly concave, apex weakly drawn out (not into acutely pointed tip) with acute angle (nearly rectangular), ventral margin weakly convex to straight; 12–16 (male) or 11–14 (female) antenna I lobes reaching to antenna II flagellomeres V–VI (male) or IV–V (female); 12–14 (male) or 10–13 (female) antenna II flagellomeres; 19–21 complete thorax segments; 19–27 telsonic spines of varying size, larger and smaller spines almost alternating, anterior spines conical, posterior spines thinner, elongate, aciculate; 8–12 furcal setae. + + + +Differential diagnosis + + + + +Ozestheria sivesae + +sp. nov. +can be easily differentiated from most other Australian species of + +Ozestheria + +by the characteristic carapace ornamentation with the smooth band dorsally within growth bands. Three other species have (partially) smooth growth bands: + +Ozestheria jiangi + +sp. nov. +, + +Ozestheria setifera + +sp. nov. +and + +Ozestheria echidna + +sp. nov. +Neither + +O. jiangi + +nor + +O. echidna + +have punctae ventrally of the smooth carapace ornamentation. + +Ozestheria setifera + +has nodular, more irregular lirae. Furthermore, + +O. setifera + +and + +O. echidna + +have dense, conspicuous setation along all growth bands (setae mostly broken off in + +O. sivesae + +). In + +O. setifera + +setal pores on concentric ridges are arranged in 2–3 rows. In + +O. echidna + +only short lirae are present and these only mid-dorsally on the carapace (ventrally on carapace growth bands completely smooth) and in + +O. jiangi + +dorsal growth bands are largely smooth (only short ventral lirae, which become longer in later growth bands, from about mid-dorsal carapace). + +Ozestheria sivesae + +differs from these and most other species also in the telsonic spination with the very variably sized spines. + + + + + +Etymology + + + +The species is named in honor of Claire Sives, who has worked on the ecology of the large branchiopod fauna on and around Bloodwood Station, +New South Wales +, and who took part in collecting some of the specimens studied here, including the species named after her. + + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +New South Wales + +• +♂ +; +Bloodwood Station +, +Freshwater Lake +, +29°29′14.7″ S +, +144°49′59.0″ E +; + +19 Feb. 2010 + +, +M. Schwentner +, +C. Sieves +and +B.V. Timms +leg.; +GenBank +no: +KJ705775 +( +COI +); +AM +P.91547. + + + + +Paratypes + + + + +AUSTRALIA +– + +New South Wales + +• +2 ♀♀ +; same data as for holotype; +GenBank +nos: +KJ705776, KJ705777 +( +COI +); +AM +P.91548, P.91549 + +• + +1 ♂ +; same data as for holotype; GenBank no: +KJ705774 +; NHMW-ZOO-CR-28482 + +. + + +Additional material +(not examined) + + + +AUSTRALIA +– + +Queensland + +• +1 juv. +; +Rockwell Station +, +Coolibah +swamp; +28°54′03.2″ S +, +144°59′22.6″ E +; + +1 Apr. 2009 + +; +M. Schwentner +and +B.V. Timms +leg.; +AM +P. 91545 + +. + + + +Type locality + + + +New South Wales +, Bloodwood Station, Freshwater Lake, +29°29′14.7″ S +, +144°49′59.0″ E +. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 46a, c–d +). Length 3.9–4.0 mm (HT: 4.0 mm), height +2.4–2.5 mm +(HT: +2.5 mm +). Coloration lightly yellowish-brown. 20–22 (HT: 22) growth lines, 16–20 evenly spaced, and 0–6 crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, rounded dorso-posterior corner. Posterior margin broadly rounded, suboval, equicurvate (b/H 0.47–0.48, HT: 0.47). Ventral margin broadly rounded. Umbo position anterior (Cr/L 0.23–0.24, HT: 0.23). + + +CARAPACE +ORNAMENTATION +( +Fig. 46e–h +). Larval valve and following growth bands finely reticulate to punctate. From about mid-dorsal carapace, unordered, shallow and short lirae appearing between punctae ventrally within growth bands, dorsal part of growth bands smooth; the dorsally smooth band becomes more pronounced in subsequent growth bands. Crowded growth bands too closely set to see ornamentation. Concentric ridges slightly raised, broad, with dorsal row of minute nodules in moniliform row ventrally on the carapace (visible under SEM). Setae filiform, preferentially preserved along the outer margin of the carapace. Setal pores in single row along all growth lines. + + +HEAD +( +Fig. 46i +). Condyle long, distally acute; occipital notch narrow. Condyle with anterobasal hump. Margin between condyle and ocular tubercle straight or weakly concave. Ocular tubercle weakly developed, forming widely obtuse angle with rostrum (close to 180°). Anterior margin of rostrum straight. Ventral margin of rostrum concave to nearly straight; apex extending well ventrally, broadly rounded, acute (close to 90°). Naupliar eye elongated, sub-rectangular. Antenna I with 12–16 (HT: 16) lobes, reaching to antenna II flagellomeres V–VI (HT: VI). Antenna II with 12–14 flagellomeres (HT: 14). + + +THORAX +. 20–21(HT:21)segments, 19–20 (HT: 20) thoracopod-bearing and one posterior limbless segment not reaching dorsal margin. Last thirteen thoracopod-bearing segments with dorsal extensions bearing spines. Dorsal extensions increasing in size posteriorly over successive segments, well developed in last nine segments. Spines thin and elongated, central spines stronger and broader in posterior segments. The +holotype +(P.91547) with a single spine situated between the last clearly demarcated segment and the telson on the left side of the body (this appears to be a deformation, maybe due to a not properly developed segment; not present in the other individuals). + + + +Fig. 46. + +Ozestheria sivesae + +sp. nov. +a–d +. Carapace. +a +. Male, holotype (P.91547). +b +. Female, paratype (NHMW-ZOO-CR-28482). +c +. Male, holotype, dorsal view (left valve only; P.91547). +d +. Male, paratype (P. 91548), SEM. +e–h +. Carapace ornamentation of male paratype (P.91548, positions marked in d by rectangles), SEM. +e +. Larval valve. +f +. Posterior carapace. +g +. Ventral carapace. +h +. Mid-carapace. +i–j +. Heads, antennae not shown. +i +. Male, holotype (P.91547). +j +. Female, paratype (P.9456). +k +. Male, holotype third right thoracopod (P.91547). +l–m +. Telson. +l +. Male, holotype (AM P:91547). +m +. Male, paratype (P.91548), SEM. +n +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–d =0.5 mm; e–h =0.05 mm; i–m =0.1 mm. + + + +THORACOPOD +III (only P.91547; +Fig. 46k +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp two-segmented, basal segment shorter than endopod. Exopod ventral extension shorter in extension to endopod, dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( +Fig. 46l–m +). 21–27 spines (HT: 21). First (anterior) spine enlarged. Spines of varying size, smaller and slightly enlarged spines more-or-less alternating; anterior spines conical, posterior spines thinner, elongate, aciculate. Dorsal margin nearly straight. Right terminal claw more strongly curved than left. + + +FURCA +( +Fig. 46l–m +). Proximally with dorsomedial longitudinal row of 12 setae, row ending distally in a single conical spine. Distal part ⅔ of furca length, with numerous small denticles. + + +Females + + +Overall appearance as in males. Carapace ( +Fig. 46b +) length +3.6–3.9 mm +, height +2.2–2.5 mm +; 17–18 growth lines, all of these widely spaced, Cr/L 0.25 and b/H 0.46–0.49. Anterior margin of rostrum weakly concave; apex with acute angle (close to 90°), weakly drawn out; ventral margin weakly convex to straight; overall rostrum shape trapezoidal ( +Fig. 46j +). Naupliar eye elongated, sub-rectangular to sub-triangular. Antenna I with 11–14 small lobes, lobes smaller than in males; reaching to antenna II flagellomeres IV–V. Antenna II with 10–13 flagellomeres. 21 segments, 20–21 thoracopod-bearing and none to one posterior limbless segment not reaching dorsal margin. Telson with 19–21 dorsal spines; left and right terminal claws equally curved. Furca with 8 setae. + + +Distribution +( +Fig. 46n +) + + +Currently known only from two localities in the catchment of the Paroo River in eastern +Australia +at the border between +New South Wales +and +Queensland +. + + + + + +Remarks + + + +Because only few specimens were available, the morphological variability of the species is not well characterized. The carapace shape of + +Ozestheria sivesae + +sp. nov. +( +Fig. 6 +) is distinct from that of most other species and overlaps fully with those of + +O. cancellata + +comb. nov. +, + +O. weeksi + +sp. nov. +, + +O. pilbarensis + +sp. nov. +and partly with + +O. timmsi + +sp. nov. +, + +O. setifera + +sp. nov. +, + +O. gemina + +sp. nov. +, + +O. marthae + +sp. nov. +, and + +O +. +selmae + +sp. nov. +(marginally). + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FF86FFFE176EFE64FDD2FE48.xml b/data/03/FD/A6/03FDA650FF86FFFE176EFE64FDD2FE48.xml new file mode 100644 index 00000000000..831b5b4dd7f --- /dev/null +++ b/data/03/FD/A6/03FDA650FF86FFFE176EFE64FDD2FE48.xml @@ -0,0 +1,423 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria paralutraria + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +4DECD2EF-56DA-495A-B637-8BCA981B6BB6 + + + + + +Fig. 33 + + + + + +Diagnosis + + + + +Ozestheria paralutraria + +sp. nov. +is characterized by a short condyle and wide occipital notch; a straight ventral carapace margin; a supracurvate posterior margin (b/H 0.32–0.38); carapace ornamentation with large, well-developed polygonal reticulations, center of each polygon smooth or with granular secondary ornamentation (best seen under SEM); male rostrum with strongly convex anterior margin, apex rounded with right angle, ventral margin strongly concave, pointing apex downwards; female rostrum anterior margin weakly convex, apex pointed and drawn out into acute tip, ventral margin weakly concave; 15 (male) or 13–18 (female) antenna I lobes reaching to antenna II flagellomeres VI–VIII (male) or V–VI (female); 14–15 (male and female) antenna II flagellomeres; 23–24 complete thorax segments; 16–27 very small and irregularly spaced, conical spines; 12 furcal setae. + + + +Differential diagnosis + + + + +Ozestheria paralutraria + +sp. nov. +is most similar to + +O. lutraria + +from which it can be differentiated by it having fewer thoracic segments (24 vs 25–27 complete segments), more telsonic spines ( +16–18 in + +O. paralutraria + +, whereas + +O. lutraria + +rarely has more than 15), fewer and longer setae on the carapace (in a single row vs in two rows along each concentric ridge in + +O. lutraria + +; best seen under SEM) and length of the carapace (up to +9 mm +vs> +10 mm +in + +O. lutraria + +; n= 4 vs n= 73). + +Ozestheria paralutraria + +has a straight ventral carapace margin, which in combination with a short condyle is present only in + +O. lutraria + +, + +O. fuersichi + +sp. nov. +and + +O. rufa + +. The carapace ornamentation of + +O. rufa + +features distinctly smaller reticulations in early growth bands and nodulous lirae in mid-carapace rather than the well-defined reticulations of + +O. paralutraria + +. + +Ozestheria fuersichi + +lacks the well-defined reticulation in mid-carapace. + +Ozestheria paralutraria + +can be further differentiated from most other species by the strongly supracurvate posterior carapace margin (b/H 0.32–0.38). + + + + + +Etymology + + + +The name is based on another morphologically similar species – + +O. lutraria + +. The Greek prefix ‘ +para +’ (meaning ‘altered’ or ‘irregular’) hints at the great morphological similarity of the two species. + + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +Western Australia + +• +♀ +; K.P.W floodout, +22 km +N of +Wittenoom +; +22°7′46.7″ S +, +118°24′4.2″ E +; + +4 Aug. 2015 + +; +B.V. Timms +leg.; +GenBank +no: PQ427013 ( +COI +); +WAM +C78013 +. + + + + +Paratypes + + + + +AUSTRALIA +– + +Western Australia + +• +1 ♂ +, +1 ♀ +; same data as for holotype; +WAM +C80216 +, +C80217 + +• + +1 ♂ +; same data as for holotype; NHMW-ZOO-CR-28498 + +. + + + +Type locality + + + +Western Australia +, K.P.W floodout, +22 km +N of Wittenoom, +22°7′46.7″ S +, +118°24′4.2″ E +. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 33a, c +). Length +8.9–9.9 mm +, height 4.4–5.0 mm. Coloration light yellow-orange, crowded growth bands lighter. 24–27 growth lines, 15–20 widely spaced and 9–17 crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, distinct dorso-posterior corner. Posterior margin rounded, greatly extending posteriorly, supracurvate (b/H 0.36–0.38). Ventral margin straight. Umbo position anterior (Cr/L 0.21–0.24). + + +CARAPACE +ORNAMENTATION +(see also +Fig. 33e–f +). Larval valve and first few growth bands appear smooth (might be due to abrasion). All other non-crowded growth bands with medium to large reticulations. Reticulations form polygonal mesh across each growth band with each polygon usually being a pentagon, hexagon or heptagon. Under SEM (based on SEM of female carapace), polygon centers smooth or granular in late juvenile growth bands, without secondary reticulations. Ornamentation uniform across all non-crowded growth bands; reticulations become irregular and transition to broken lines on growth bands of incipient carapace crowding; crowded growth bands very narrow, without obvious ornamentation (under SEM, crowded growth bands with irregular, granular, lirae-like ornamentation). Concentric ridges raised. Setae short, thin and inconspicuous, in many individuals none visible; under SEM one row of setae and corresponding setal pores along all growth lines. + + +HEAD +( + +Fig. +33g + +). Condyle short, rounded; occipital notch wide. Condyle with anterobasal hump. Margin between condyle and ocular tubercle straight. Ocular tubercle well developed, forming obtuse (~120°) angle with rostrum. Anterior margin of rostrum strongly convex. Apex rounded with nearly rectangular angle. Ventral margin of rostrum deeply concave, pointing apex downwards. Naupliar eye small, roundish. Antenna I long with 15 lobes, reaching to antenna II flagellomeres VI–VIII. Antenna II with 14–15 flagellomeres. + + +THORAX +. 25 segments, 24 thoracopod-bearing and one posterior limbless segment not reaching dorsal margin. Only few segments with spine-bearing dorsal extensions, posterior segments lacking spines. + + +THORACOPOD +III (only NHMW-CR-28498; +Fig. 33k +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp two-segmented, basal segment shorter than endopod. Exopod ventral extension slightly shorter in extension to endopod, dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( +Fig. 33i +). 16–18 spines. First (anterior) spine enlarged. All spines conical; spines small or tiny, irregular in size and spacing. Dorsal margin weakly concave. Right terminal claw more strongly curved than left. + + +FURCA +( +Fig. 33i +). Broken off in all studied males. + + +Females + + +Overall appearance as in males. Carapace ( +Fig. 33b, d +) length 8.0– +8.9 mm +(HT: +8.9 mm +), height +4.1– 4.2 mm +(HT: +4.2 mm +); 17–18 (HT: 17) growth lines, 14–15 (HT: 14) widely spaced and 2–4 (HT: 3) crowded; Cr/L0.21–0.24 (HT: 0.24) and b/H 0.32–0.35 (HT: 0.35). Angle between ocular tubercle and rostrum obtuse (~160°) to straight ( +Fig. 33h +). Anterior margin of rostrum weakly convex, apex drawn out into acute, pointed tip; ventral margin weakly concave. Antenna I with 13–18 small lobes (HT: 18), lobes smaller than in males; reaching to antenna II flagellomeres V–VI (HT: VI). Antenna II with 14–15 flagellomeres (HT: 14). 24–25 (HT: 24) segments, 23 thoracopod-bearing and one to two (HT: 1) posterior limbless segments not reaching dorsal margin. Telson with 16–27 (HT: 27) dorsal spines (HT: posteriormost spines tiny); left and right terminal claws equally curved ( +Fig. 33j +). Furca with 12 setae, followed by single elongate spine; distal part ⅓–½ (HT: 1/3) of furcal length, with numerous small denticles. + + +Distribution +( +Fig. 33l +) + + + +Ozestheria paralutraria + +sp. nov. +is known only from a single locality in northwestern +Western Australia +. + + + + +Fig. 33. + +Ozestheria paralutraria + +sp. nov. +a–d +. Carapace. +a +. Male, paratype (NHMW-ZOO-CR-28498). +b +. Female, holotype (WAM C78013). +c +. Dorsal view male paratype (left valve only; NHMW-ZOO- CR-28498). +d +. Female, paratype (WAM C80217), SEM. +e–f +. Carapace ornamentation (positions marked in b and d by rectangles). +e +. Posterior carapace, female paratype (WAM C80217), SEM. +f +. Posteroventral carapace, female, holotype (WAM C78013). +g–h +. Head (antennae not shown). +g +. Male, paratype (NHMW-ZOO-CR-28498). +h +. Female, holotype (WAM C78013). +i–j +. Telson. +i +. Male, paratype, furca broken off (WAM C80216). +j +. Female, holotype (WAM C78013). +k +. Male, third left thoracopod (paratype, NHMW-ZOO-CR-28498). +l +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–d =1 mm; e–f, i–k = 0.2 mm; g–h=0.5 mm. + + + + + +Remarks + + + +A female was selected as the +holotype +, because the telson and furca of each male were damaged. The carapace shape of + +Ozestheria paralutraria + +sp. nov. +( +Fig. 5 +) is distinct from that of most other species and overlaps partly with + +O. lutraria + +. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FF96FFEE174DFEECFAB8FE3C.xml b/data/03/FD/A6/03FDA650FF96FFEE174DFEECFAB8FE3C.xml new file mode 100644 index 00000000000..79e78fa2d0b --- /dev/null +++ b/data/03/FD/A6/03FDA650FF96FFEE174DFEECFAB8FE3C.xml @@ -0,0 +1,424 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria richteri + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +35804CD8-1540-44C0-BB7D-F55A92D44692 + + + + + +Fig. 38 + + + +Ozestheria +sp. G + +– + +Schwentner +et al +. 2015 a + +: figs 2, 6; 2020: figs 1–2. + + + + + +Diagnosis + + + + +Ozestheria richteri + +sp. nov. +is characterized by a short condyle with a hump at its base and a mediumwide occipital notch; a rounded ventral carapace margin; carapace ornamentation dorsally on carapace smooth or with irregular depression, from mid-carapace with granular or nodulous intermittent lirae; male rostrum with convex anterior margin, apex strongly rounded with right angle, ventral margin straight or with slightly convex hump; female rostrum anterior margin convex, apex strongly rounded, ventral margin convex with anterior notch; 9–11 (male) or 8–12 (female) antenna I lobes reaching to antenna II flagellomeres IV–V (male) or IV–V (female); 8–9 (male) or 9 (female) antenna II flagellomeres; 19 complete thorax segments; 10–17 large spines, varying in shape between conical and elongate and aciculate, unequal in size; 0–1 furcal setae. + + + +Differential diagnosis + + + + +Ozestheria richteri + +sp. nov. +can be differentiated from all other species by the shape of the condyle, rostrum and carapace ornamentation. It is the only species with a short condyle and wide occipital notch, which has a well-defined hump at the condyle base (this is usually only present in species with a longer condyle) and has fewer furcal setae (0–1) than most other species. The condyle is uniquely shaped by being short, but distally pointed. The broad male rostrum and nodular carapace ornamentation (present on growth bands of later ontogenetic stages) are shared only with + +O. pellucida + +and to a lesser degree + +O. rufa + +and no other short-condyled species. + +Ozestheria pellucida + +differs from + +O. richteri + +by the condyle shape, the lighter and pellucid carapace and the number of furcal setae (14 vs 0–1), and by the lack of liral ornamentation (well visible posteriorly on the carapace of + +O. richteri + +). + +Ozestheria rufa + +differs from + +O. richteri + +by the pointed, drawn-out rostral apex, the number of antenna I lobes (16–19 vs 8–12), the number of complete thorax segments (23–24 vs 19) and the number of furcal setae (6–15 vs 0–1). + + + + + +Etymology + + + +The species is named in honor of the German zoologist Stefan Richter. Without Stefan’s contribution and support all the newly described species of + +Ozestheria + +would not have been discovered. He supervised and planned MS’ PhD thesis, which lay the foundation for this publication, participated in collecting specimens and started MS’ interest in these fascinating animals. + + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +Western Australia + +• +♂ +; +Paynes Find +, +Bullamanya Rock +, pool 5; +29°09′50.9″ S +, +117°39′40.4″ E +; + +20 Aug. 2011 + +, +B.V. Timms +leg.; +GenBank +no: +KJ705759 +( +COI +); +AM +P.91531. + + + + +Paratypes + + + + +AUSTRALIA +– + +Western Australia + +• +2 ♂♂ +, +1 ♀ +; same data as for holotype; +GenBank +nos: +KJ705756 +to +KJ705758 +( +COI +); +AM +P.91528 to P.91530 + +• + +1 ♀ +; same data as for holotype; +GenBank +no: +KJ705755 +( +COI +); NHMW-ZOO-CR-28480 + +. + + + +Type locality + + + +Western Australia +, Paynes Find, Bullamanya Rock, pool 5, +29°09′50.9″ S +, +117°39′40.4″ E +. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 38a, c–d +). Length +4.8–5.1 mm +(HT: +5.1 mm +), height +2.9–3.1 mm +(HT: +3.1 mm +). Coloration dorsally dark brown to nearly black (~⅓–⅔ of carapace) with roundish dark area below umbo, fading into yellowish-brown ventrally. 15–17 growth lines, 13 widely spaced and 2–4 crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, dorso-posterior corner rounded. Posterior margin widely rounded, supracurvate (b/H 0.39–0.44, HT: 0.44). Ventral margin widely rounded. Umbo position anterior to submedian (Cr/L 0.23–0.26, HT: 0.26). + + +CARAPACE +ORNAMENTATION +( +Fig. 38e–g +). Larval valve and following growth bands smooth, following growth bands appear smooth, partly with irregular depressions or very anastomosing lirae. From about mid carapace, with shallow, inconspicuous, subparallel, anastomosing lirae forming ventrally on growth bands (nodular under SEM, highly nodular with incipient crowding in the ventral and anterior part of the carapace). Concentric ridges shallow. Setae spiniform, preserved along carapace margin (under SEM setal pores along all growth lines, dorsally and medially on carapace with two alternating rows of setal pores on early concentric ridges, ventrally a single row). + + +HEAD +( +Fig. 38h +). Condyle short, slightly elongate and pointed distally; occipital notch medium wide. Condyle with well-developed anterobasal hump. Margin between condyle and ocular tubercle weakly concave. Ocular tubercle well developed, forming obtuse angle with rostrum (close to 90°). Anterior margin of rostrum convex, protruding anteriorly. Ventral margin of rostrum nearly straight or with slightly convex hump mid-length (HT: with hump) with notch close to apex, apex broadly rounded, acute. Naupliar eye triangular. Antenna I with 9–11 (HT: 11) lobes, reaching to antenna II flagellomeres IV–V (HT: IV). Antenna II with 8–9 (HT: 9) flagellomeres. + + +THORAX +. 20 segments, 19 thoracopod-bearing and one posterior limbless segment not reaching dorsal margin. Dorsal armature well developed in last eight segments, spines thin and elongated, central spines stronger and broader in posterior segments. + + +THORACOPOD +III (only P.91531; +Fig. 38l +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp two-segmented, basal segment shorter than endopod. Exopod ventral extension shorter in extension than endopod; dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( +Fig. 38j–k +). 8–12 (HT: 12) spines. First (anterior) spine enlarged. Spines of highly variable size and spacing (mostly widely spaced, space increases posteriorly), varying in shape between broad, conical and elongate, slender, aciculate. Several larger spines interspersed among much smaller spines. Dorsal margin nearly straight, posteriorly slightly concavely curved. Right terminal claw more strongly curved than left. + + +FURCA +( +Fig. 38j–k +). Proximally with dorsomedial longitudinal row of 0–1 seta (HT: 1) setae, row ending distally in a single conical spine. Distal part ½ of furca length, with numerous small denticles. + + +Females + + +Very similar to males. Carapace ( +Fig. 38b +) length 3.9–4.0 mm, height +2.5–2.6 mm +. 16–17 growth lines, of these 12–14 evenly spaced and 3–4 crowded; Cr/L 0.23–0.24 and b/H 0.47–0.48. Ocular tubercle and rostrum form acute angle (~20–80°); rostrum anterior margin convex; apex widely rounded, rectangular (not drawn out), ventral margin convex with anterior notch ( +Fig. 38i +). Antenna I with 8–12 indistinct lobes, reaching to antenna II flagellomeres IV–V. Antenna II with nine flagellomeres. Telson 16–17 spines, their shape, size and spacing as in males. Furca bearing 0–1 seta. + + + +Fig. 38. + +Ozestheria richteri + +sp. nov. +a–d +. Carapace. +a +. Male, holotype (P.91531). +b +. Female, paratype (P.91528). +c +. Male, holotype, dorsal view (only left valve shown, P.91531). +d +. Male, paratype (P.91529), SEM. +e–g +. Carapace ornamentation of male paratype (P.91529; positions marked in d by rectangles). +e +. Mid-dorsal carapace. +f +. Mid-carapace. +g +. Ventral carapace. +h–i +. Head (antennae not shown). +h +. Male, holotype (P.91531). +i +. Female, paratype (P.91528). +j–k +. Telson. +j +. Male, holotype (P.91531). +k +. Male, paratype (P.91529), SEM. +l +. Male, holotype third right thoracopod (P.91531). +m +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–d =0.5 mm; e–l=0.1 mm. + + + +Distribution +( + +Fig. +38m + +) + + +Currently known only from its +type +locality in western +Western Australia +. + + + + + +Remarks + + + +The carapace shape of + +Ozestheria richteri + +sp. nov. +( +Fig. 5 +) is distinct from that of most other species and overlaps partly with those of + +O. rubra + +, + +O. henryae + +sp. nov. +, + +O. berneyi + +and + +O. gemina + +sp. nov. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FFADFFD31741FEECFE79FE14.xml b/data/03/FD/A6/03FDA650FFADFFD31741FEECFE79FE14.xml new file mode 100644 index 00000000000..84964e83e12 --- /dev/null +++ b/data/03/FD/A6/03FDA650FFADFFD31741FEECFE79FE14.xml @@ -0,0 +1,454 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria henryae + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +26EFE2F6-F832-45F9-AD47-50E142A3EDA8 + + + + + +Fig. 22 + + + +Ozestheria +sp. D + +3 – + +Schwentner +et al +. 2015a + +: figs 2, 6. + + + +Ozestheria rubra + +– + +Schwentner +et al +. 2020 + +: figs 1–2. + + + + + +Diagnosis + + + + +Ozestheria henryae + +sp. nov. +is characterized by a short condyle and wide occipital notch; a rounded ventral carapace margin; carapace ornamentation with medium to large, well-developed polygonal reticulations, each polygon with polygonal secondary ornamentation (best seen under SEM); male rostrum with strongly convex anterior margin, apex strongly rounded with acute (nearly rectangular), ventral margin concave, pointing apex downwards; female rostrum anterior margin slightly s-shaped, apex pointed (not elongated or drawn out), ventral margin slightly concave; 14–18 (male) or 14 (female) antenna I lobes reaching to antenna II flagellomeres V–VII (male) or III (female); 14–16 (male) or 13 (female) antenna II flagellomeres; 22 complete thorax segments; 21–30 small, unequally sized and spaced conical telsonic spines, spines in the central part of the telson enlarged; 5–6 furcal setae. + + + +Differential diagnosis + + + +See differential diagnosis of + +O. elliptica + +. + + + + + +Etymology + + + +The species is named in honor of Marguerite Henry (1895–1982), who was one of the few female Australian zoologists and taxonomists in the early 20 +th +century. She described one species of + +Ozestheria +– + + +Ozestheria rubra +( +Henry, 1924 +) + +. + +Ozestheria henryae + +sp. nov. +is probably the sister species to + +O. rubra + +. For details on her scientific life and achievements see +Damkaer (2014) +. + + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +Queensland + +• +♂ +; +Lake Galilee +; +22°25′37.3″ S +, +145°42′13.4″ E +; + +15 Feb. 2010 + +; +M. Schwentner +, +C. Sieves +and +B.V. Timms +leg.; +GenBank +no: +KJ705637 +( +COI +); +AM +P.91408. + + + + +Paratypes + + + + +AUSTRALIA +– + +Queensland + +• +2 ♂♂ +, +1 ♀ +; same data as for holotype; +GenBank +nos: +KJ705610, KJ705610, KJ705636 +( +COI +); +AM +P.91381, P.91382, P.91407 + +• + +1 ♂ +; same data as for holotype; GenBank no: +KJ705638 +; NHWM-CR-28476 + +. + + + +Type locality + + + +Australia +, Lake Galilee, +22°25′37.3″ S +, +145°42′13.4″ E +. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 22a, c–d +). Length +5.4–7.3 mm +(HT: +6.9 mm +), height +3.3–4.3 mm +(HT: +4.3 mm +). Coloration light. 17–23 (HT: 23) growth lines, 17–20 (HT: 20) widely spaced and 0–3 crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, distinct dorso-posterior corner. Posterior margin broadly rounded, supracurvate (b/H 0.40–0.41, HT: 0.40). Ventral margin widely rounded. Umbo position anterior (Cr/L 0.19–0.22, HT: 0.22). + + +CARAPACE +ORNAMENTATION +( +Fig. 22e–h +). Larval valve with shallow reticulations. Each growth band with medium to large, well-developed, strongly raised reticulations. Reticulations form polygonal mesh across each growth band with each polygon usually being a pentagon, hexagon or heptagon. Polygon-size increasing during ontogeny, largest in the dorsal to median part of each growth band; under SEM secondary mesh or polygonal reticulation within each primary polygon (less strongly developed or absent ventrally within growth bands and on carapace). Ornamentation uniform across all non-crowded growth bands, crowded growth bands usually a single row of polygonal reticulations resulting in radial appearance. Concentric ridges raised. Setae mostly long and thick; preferentially preserved ventral and posterior parts of the carapace. Setal pores in single, irregular row along all growth lines. + + + +Fig. 22. + +Ozestheria henryae + +sp. nov. +a +. Carapace (male, holotype, P.91408). +b +. Carapace (female paratype, P.91381). +c +. Carapace, dorsal view (right valve only, male, holotype, P.91408). +d–g +. Male, paratype (P.91407), SEM. +d +. Carapace. +e +. Mid-carapace ornamentation (position marked in d by rectangle). +f +. Posterior carapace ornamentation (position marked in d by rectangle). +g +. Ventral carapace ornamentation, including crowded growth lines (position marked in d by rectangle). +h +. Mid-carapace ornamentation (male, holotype, P.91408). +i–j +. Head (antennae not shown). +i +. Male, holotype (P.91408). +j +. Female, paratype (P.91381). +k +. Third left thoracopod (male, holotype, P.91408). +l–m +. Telson. +l +. Male, paratype (P.91407), SEM. +m +. Male, holotype (P.91408). +n +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–d = 1 mm; e–h, l =0.1 mm; i–k =0.5 mm; m =0.2 mm. + + + +HEAD +( +Fig. 22i +). Condyle rounded, short, only weakly protruding; occipital notch wide. Condyle lacking anterobasal hump. Margin between condyle and ocular tubercle straight. Ocular tubercle well developed, forming obtuse (~90°–120°) angle with rostrum. Anterior margin of rostrum strongly convex. Apex strongly rounded, acute (nearly rectangular). Ventral margin of rostrum weakly concave, pointing apex slightly downwards; small notch anteriorly. Naupliar eye elongated, sub-triangular to sub-rectangular. Antenna I long with 14–18 (HT: 14) lobes, reaching to antenna II flagellomeres V–VII (HT: VII). Antenna II with 14–16 (HT: 14) flagellomeres. + + +THORAX +. 22–23 (HT: 22) segments, 22 thoracopod-bearing and none to one posterior limbless segment not reaching dorsal margin. Mid to posterior thoracopod-bearing segments with spine bearing dorsal extensions. Dorsal extensions increasing in size posteriorly over successive segments; spines mostly short, in posterior segments with fewer spines and central spines stouter but shorter. + + +THORACOPOD +III (only P.91408; +Fig. 22k +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp one-segmented. Exopod ventral extension slightly overreaching endopod, dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( +Fig. 22l–m +). 21–30 (HT: 30) spines. First (anterior) spine enlarged. Spines conical, subequally spaced, anterior spines smaller, followed by several larger spines close to the central part of the telson (with few interspersed smaller spines); posteriorly spines slightly thinner and more drawn out and increasing in length (last ~¼ of telson). Dorsal margin straight, posteriorly weakly concavely curved. Right terminal claw more strongly curved than left, in some individuals equally curved. + + +FURCA +( +Fig. 22l–m +). Proximally with dorsomedial longitudinal row of 5–6 (HT: 5) setae, row ending distally in a single conical spine. Distal part ⅔ of furcal length, with numerous small denticles. + + +Female + + +Overall appearance as in males. Carapace ( +Fig. 22b +) length +6.8 mm +, height +4.2 mm +; 18 growth lines, these 16 widely spaced and 2 crowded; Cr/L 0.22 and b/H 0.4. Ocular tubercle forming nearly straight angle with rostrum ( +Fig. 22j +). Anterior margin of rostrum slightly s-shaped (dorsally slightly convex, ventrally slightly concave); apex pointed (~45°), weakly drawn out; ventral margin weakly concavely curved. Antenna I with 14 small lobes, lobes smaller than in males; reaching to antenna II flagellomere III. Antenna II with 13 flagellomeres. 22 segments, all of these thoracopod-bearing. Telson with 26 dorsal spines; left and right terminal claws equally curved. Furca with 5 setae. + + +Distribution +( +Fig. 22n +) + + + +Ozestheria henryae + +sp. nov. +is known only from the northern regions of the Cooper Creek catchment in central +Queensland +. + + + + + +Remarks + + + + +Schwentner +et al +. (2020) + +wrongly identified this species as + +O. rubra + +. A comparison with the respective +type +material showed that + +O. henryae + +sp. nov. +is not conspecific with + +O. rubra +. + +So far only a single female of + +O. henryae + +could be studied, so the intraspecific variability cannot be assessed for female characters (e.g., rostrum shape). + + +The carapace shape of + +Ozestheria henryae + +sp. nov. +( +Fig. 5 +) is distinct from that of most other species and overlaps fully with that of + +O. rubra + +and partly with those of + +O. gemina + +sp. nov. +, + +O. matuwa + +sp. nov. +and + +O. richteri + +sp. nov. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FFB5FFCB1740FB0DFB8EFA11.xml b/data/03/FD/A6/03FDA650FFB5FFCB1740FB0DFB8EFA11.xml new file mode 100644 index 00000000000..a1a8513342b --- /dev/null +++ b/data/03/FD/A6/03FDA650FFB5FFCB1740FB0DFB8EFA11.xml @@ -0,0 +1,483 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria matuwa + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +4F91152F-FCDD-4306-9467-B580768A9FA0 + + + + + +Fig. 29 + + + + + +Diagnosis + + + + +Ozestheria matuwa + +sp. nov. +is characterized by a short condyle and a wide occipital notch; a rounded ventral carapace margin; carapace ornamentation with large, well-developed polygonal reticulations, each polygon with polygonal secondary ornamentation (best seen under SEM); male rostrum with strongly convex anterior margin, apex rounded with acute to right angle, ventral margin concave, pointing apex downwards; female rostrum anterior margin straight or strongly convex, apex pointed (elongated, drawn out), ventral margin slightly concave; 12–19 (male) or 13–14 (female) antenna I lobes reaching to antenna II flagellomeres VII–IX (male) or III–IV (female); 11–14 (male) or 11–13 (female) antenna II flagellomeres; 22–24 complete thorax segments; 18–27 small, unequally sized and spaced conical telsonic spines, some spines in the central part of the telson enlarged; 3–14 furcal setae. + + + +Differential diagnosis + + + +See differential diagnosis of + +O. elliptica + +. + + + + + +Etymology + + +The name is derived from Matuwa, the traditional owner’s name for the Lake Carnegie area in the Martu language; noun in apposition. + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +Western Australia + +• +♂ +; +Gidgi Lake +, + +20 km +N of Kalgoorlie + +; +30°36′11″ S +, +121°24′51.7″ E +; + +18 Feb. 2014 + +; +K. Quinlan +leg.; +GenBank +no: PQ427000 ( +COI +); +WAM +C78010 +. + + + + +Paratypes + + + + +AUSTRALIA +– + +Western Australia + +• +2 ♂♂ +; same data as for holotype; +GenBank +nos: PQ426999, PQ427003 ( +COI +); +WAM +C80207 +, +C80208 + +• + +1 ♂ +; same data as for holotype; GenBank no: PQ427001 ( +COI +); NHMW-ZOO-CR-28496 + +. + + + +Other material examined + + + + +AUSTRALIA +– + +Western Australia + +• +3 ♂♂ +, +1 ♀ +; +Wetland + +2.4 km +S of Boondi Rock + +; +31°11′40.4″ S +, +120°23′55.7″ E +; + +24 Feb. 2017 + +; +K. Quinlan +leg.; +WAM +C78006 +, +C80204 +to +C80206 + +• + +1 ♂ +, +2 ♀♀ +; +Lindsay Gordon Lagoon +, +SW of Lorna Glen Homestead +; +26°15′45.2″ S +, +121°29′51.2″ E +; + +18 Mar. 2017 + +; +K. Quinlan +leg.; +WAM +C80202 +, +C80203 +, +C80201 + +• + +4 ♂♂ +, +1 ♀ +; +Wetland + +50 km +SW of Bullabulling + +(Goldfields Woodlands Conservation Park); +31°16′44.5″ S +, +120°32′36″ E +; + +22 Feb. 2017 + +; +K. Quinlan +leg.; +WAM +C78007 +, +C80209 +to +C80212 + +. + + + +Type locality + + + +Western Australia +, Gidgi Lake, +20 km +N of Kalgoorlie, +30°36′11″ S +, +121°24′51.7″ E +. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 29a, c–d +). Length +6.4–9.4 mm +(HT: +9.4 mm +, mean: +7.7 mm +), height 3.5–5.3 (HT: +5.2 mm +, mean: +4.3 mm +). Coloration whitish to yellow-brownish or brown, crowded growth bands lighter, often whitish. 19–23 (HT: 22, mean: 21) growth lines, 15–20 (HT: 17, mean: 17) widely spaced and 2–7 (HT: 5, mean: 4) crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, distinct dorso-posterior corner. Posterior margin broadly rounded, suboval, supracurvate to infracurvate (b/H 0.41–0.48, HT: 0.45, mean: 0.43). Ventral margin widely rounded. Umbo position anterior (Cr/L 0.21–0.24, HT: 0.24, mean: 0.22). + + +CARAPACE +ORNAMENTATION +( +Fig. 29e–i +). Larval valve with shallow reticulations. Each growth band with large, well-developed, strongly raised reticulations. Reticulations form polygonal mesh across each growth band with each polygon usually being a pentagon, hexagon or heptagon. Polygon-size increasing during ontogeny, largest in the median part of each growth band; under SEM secondary mesh or polygonal reticulation within each primary polygon (less strongly developed or absent ventrally within growth bands). Ornamentation uniform across all non-crowded growth bands, crowded growth bands either too narrow to show reticulation or with a single row of polygonal reticulations, sometimes resulting in radial appearance. Concentric ridges raised. Setae of varying length; preferentially preserved in ventral and posterior parts of the carapace. Setal pores in single, irregular row along all growth lines. + + +HEAD +( +Fig. 29j +). Condyle short, distally rounded; occipital notch wide. Condyle lacking anterobasal hump. Margin between condyle and ocular tubercle straight to slightly concave. Ocular tubercle well developed, forming obtuse, nearly rectangular (~90–120°) angle with rostrum. Anterior margin of rostrum strongly convex. Apex weakly rounded, acute to nearly rectangular (~90°), pointing slightly downwards. Ventral margin of rostrum weakly to strongly concave (HT: strongly concave), in some individuals with notch anteriorly (HT: weakly developed). Naupliar eye elongated, subtriangular to subrectangular.Antenna I long with 12–19 lobes (HT: 16; mean: 15), reaching to antenna II flagellomeres VII–IX (HT: IX; mean: VIII). Antenna II with 11–14 flagellomeres (HT: 14; mean: 13). + + + +Fig. 29. + +Ozestheria matuwa + +sp. nov. +a–d +. Carapace. +a +. Male, holotype (WAM C78010). +b +. Female (WAM C80202). +c +. Dorsal view, male, holotype (left valve only; WAM C78010). +d +. Male, paratype (WAM C80208), SEM. +e–h +. Carapace ornamentation (male paratype WAM C80208; positions marked in a, d and f by rectangles), SEM. +e +. Dorsal carapace. +f +. Mid-carapace. +g +. Ventral carapace. +h +. Midcarapace, detail. +i +. Carapace ornamentation (male, holotype WAM C78010), ventral. +j–l +. Head (antennae not shown). +j +. Male, holotype (WAM C78010). +k +. Female (WAM C80202). +l +. Female (WAM C80212). +m–n +. Telson. +m +. Male, holotype (WAM C78010). +n +. Male (WAM C80209). +o +. Third left thoracopod (holotype, WAM C78010). +p +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–d =1 mm; e–g, i =0.2 mm; h= 0.05 mm; j–o =0.5 mm. + + + +THORAX +. 23–25 (HT: 24; mean: 24) segments, 22–24 (HT: 23; mean: 23) thoracopod-bearing and one (HT: one) posterior limbless segment not reaching dorsal margin. In most thoracopod-bearing segments dorsal extensions bearing several spines and setae, posterior segments with only few spines. + + +THORACOPOD +III (only WAM C78010; +Fig. 29o +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp two-segmented, basal segment shorter than endopod. Exopod ventral extension subequal to slightly shorter in extension to endopod, dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( + +Fig. +29m +–n + +). 18–27 spines (HT: 24; mean: 22). First (anterior) spine enlarged. Spines on anterior ½ to ¾ of telson short, conical, varying in size; posterior spines thinner, elongate, aciculate, increasing in size; aciculate spines preceded by a few (usually 2) particularly small spines; 1–4 larger spines interspersed. Dorsal margin straight or anteriorly convex, then concavely curved. Right terminal claw more strongly curved than left. + + +FURCA +( + +Fig. +29m +–n + +). Proximally with dorsomedial longitudinal row of 8–14 (HT: 9, mean: 10) setae, row ending distally in a single conical spine. Distal part ⅓–⅔ of furcal length, with numerous small denticles. + + +Females + + +Overall appearance as in males. Carapace ( +Fig. 29b +) length +5.6–7.4 mm +, height +3.1–4.2 mm +; 19–25 growth lines, 14–18 and 2–10; Cr/L 0.21–0.23 and b/H 0.42–0.47. Angle between ocular tubercle and rostrum obtuse (~120–150°) ( +Fig. 29k–l +). Anterior margin of rostrum strongly convex to nearly straight (slightly undulating); apex with acute angle (~50°), pointed, drawn out into acute tip; ventral margin weakly concave. Antenna I with 13–14 small lobes, lobes smaller than in males; reaching to antenna II flagellomeres III–IV. Antenna II with 11–13 flagellomeres. 24 segments, of these 23 thoracopod-bearing and one posterior limb-less segment not reaching dorsal margin. Telson with 19–23 dorsal spines; right terminal claws stronger curved. Furca with 3–8 setae. + + +Distribution +( +Fig. 29p +) + + + +Ozestheria matuwa + +sp. nov. +has been recorded in central and southern +Western Australia +. + + + + + +Remarks + + + +The carapace shape of + +Ozestheria matuwa + +sp. nov. +( +Fig. 5 +) is distinct from that of most other species and partly overlaps with those of + +O. rubra + +, + +O. henryae + +and + +O. christiani + +sp. nov. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FFC0FFA41740FCFEFE24FA12.xml b/data/03/FD/A6/03FDA650FFC0FFA41740FCFEFE24FA12.xml new file mode 100644 index 00000000000..7797053cc30 --- /dev/null +++ b/data/03/FD/A6/03FDA650FFC0FFA41740FCFEFE24FA12.xml @@ -0,0 +1,518 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria echidna + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +09E54F6C-67ED-45FE-8DE6-6167AF1C7BA6 + + + + + +Fig. 16 + + + + + +Diagnosis + + + + +Ozestheria echidna + +sp. nov. +is characterized by a long condyle and a narrow occipital notch; a broadly rounded ventral carapace margin; carapace ornamentation dorsally on carapace punctate, following growth bands smooth with short lirae ventrally within growth bands, lirae decrease in size in following growth bands (lirae absent in growth bands of later ontogenetic stages); carapace setae very long and thick, very densely arranged along most concentric ridges, giving the whole carapace a furry or pelt like appearance (setal pores of broken off setae clearly visible under stereo microscope); male rostrum with weakly convex anterior margin, apex rounded with right angle, ventral margin strongly concave, pointing apex downwards; female rostrum anterior margin straight or undulating, apex right angle (not rounded or drawn out into acute tip), ventral margin weakly concave or weakly convex; 9–14 (male) or 8–15 (female) antenna I lobes reaching to antenna II flagellomeres IV–IX (male) or IV–V (female); 11– 16 (male) or 12–16 (female) antenna II flagellomeres; 19–21 complete thorax segments; 31–44 telsonic spines, most spines aciculate, densely spaced, usually no larger spines interspersed, slightly increasing in length posteriorly; 4–11 furcal setae. + + + +Differential diagnosis + + + + +Ozestheria echidna + +sp. nov. +can be easily differentiated from most other Australian species of + +Ozestheria + +by the dense setation of the carapace (in most other species the majority of setae are broken off) and the characteristic carapace ornamentation with the extensive smooth band dorsally within growth bands and only short ventral lirae (lirae completely missing ventrally on carapace). + +Ozestheria setifera + +sp. nov. +also has a dense carapace setation, but it has shorter and thinner setae, more pronounced and nodular lirae on the growth bands (which do not decrease in size ventrally on carapace) and fewer telsonic spines (18–31 vs 28–44). + +Ozestheria jiangi + +sp. nov. +and + +O. sivesae + +sp. nov. +also have smooth bands in their carapace ornamentation, but their growth bands contain prominent lirae (separated by punctae in + +O. sivesae + +) ventrally on the carapace; they also lack the dense setation and differ in the shape of the male and female rostrum. + + + + + +Etymology + + +The species is named after an iconic Australian animal, the echidna, due to the thick, dense and long spines on the species’ carapace. Echidna is a junior homonym that has become the common word for this species group; noun in apposition. + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +Western Australia + +• +♂ +; +Chaddolinna Pool +, + +5 km +E of Mulga Downs + +; +22°18′0.1″ S +, +118°49′32.5″ E +; + +13 Mar. 2007 + +; +A.M. Pinder +leg.; +WAM +C77994 +. + + + + +Paratypes + + + + +AUSTRALIA +– + +Western Australia + +• +1 ♂ +, +2 ♀♀ +; same data as for holotype; +WAM +C80241 +to +C80243 + +• + +1 ♂ +; same data as for holotype; NHMW-ZOO-CR-29004 + +. + + + +Other material examined + + + + +AUSTRALIA +– + +Western Australia + +• +1 ♂ +, +1 ♀ +; 14 +Mile Pool +, + +114 km +N of Newman + +; +22°33′13″ S +, +119°51′49.1″ E +; + +15 Mar. 2016 + +; +A.M. Pinder +leg.; +WAM +C80233 +, +C80232 + +• + +4 ♂♂ +, +1 ♀ +; +Paradise Pool +, + +61 km +SW of Pardoo + +; +20°22′35″ S +, +119°25′21″ E +; + +13 Sep. 2014 + +; +B.V. Timms +leg.; +WAM +C77992 +, +C80237 +to +C80240 + +• + +4 ♂♂ +; +Moorimoordinina Native Well +, + +118 km +S of Nullagine + +; +22°30′48″ S +, +119°46′15.4″ E +; + +10 Apr. 2008 + +; +A.M. Pinder +leg.; +WAM +C77991 +, +C80244 +to +C80246 + +. + + + +Type locality + + + +Western Australia +, Chaddolinna Pool, +5 km +E of Mulga Downs, +22°18′0.1″ S +, +118°49′32.5″ E +. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 16a, c +). Length +5.3–7.3 mm +(HT: +6.7 mm +, mean: +6.3 mm +), height +3.1–4.5 mm +(HT: 4.0 mm, mean: +3.8 mm +). Coloration varying from light orange-brownish to light reddish-brown; outer margin lighter. 34–65 (HT: 43; mean: 48) growth lines, 12–58 (HT: 34; mean: 35) widely spaced and 3–37 (HT: 9; mean: 14) crowded; also, wide growth bands rather densely spaced. + + +CARAPACE +SHAPE +. Dorsal margin straight, dorso-posterior corner rounded. Posterior margin broadly rounded, suboval, equicurvate (b/H 0.48–0.52; HT: 0.49, mean: 0.51). Ventral margin broadly rounded. Umbo position submedian (Cr/L 0.25–0.30; HT: 0.26, mean: 0.27). + + +CARAPACE +ORNAMENTATION +(compare +Fig. 16e–f +). Larval valve and first few growth bands granular or punctate (puntate under SEM), in some individuals inconspicuous lirae forming between punctae. Following growth bands, dorsally within growth bands smooth, ventrally with short lirae; lirae decrease in extension in progressing growth bands (lirae become so reduced that they are only visible when terminating in the concentric ridge). Ornamentation usually concealed by setae. Crowded growth bands without apparent ornamentation. Concentric ridges shallow; under SEM smooth, appears serrated dorsally where the short lirae terminate. Setae very long and thick, very densely arranged along most concentric ridges, giving the whole carapace a furry or pelt-like appearance; setal pores of broken off setae clearly visible under stereo microscope; setal pores in irregular row along all growth lines. + + +HEAD +( +Fig. 16i +). Condyle long, distally rounded; occipital notch narrow. Condyle with anterobasal hump. Margin between condyle and ocular tubercle straight to weakly concave. Ocular tubercle weakly developed, forming obtuse (~150°) angle with rostrum. Anterior margin of rostrum weakly convex. Apex rounded with right angle. Ventral margin of rostrum strongly concave, pointing apex downwards. Naupliar eye subtriangular, varying in size. Antenna I long with 9–14 lobes (HT: 13; mean: 12), reaching to antenna II flagellomeres IV–IX (HT: IX; mean: VII). Antenna II with 11–16 flagellomeres (HT: 15; mean: 14). + + + +Fig. 16. + +Ozestheria echidna + +sp. nov. +a–d +. Carapace. +a +. Male, holotype (WAM C77994). +b +. Female, paratype (WAM C80243). +c +. Dorsal view, male, holotype (right valve only; WAM C77994). +d +. Female, paratype (WAM C80242), SEM. +e–g +. Carapace ornamentation (female paratype WAM C80242; positions marked in d by rectangles), SEM. +e +. Dorsal carapace. +f +. Mid-carapace. +g +. Mid-ventral carapace. +h +. Carapace setation (male, holotype, WAM C77994). +i–j +. Head (antennae not shown). +i +. Male, holotype (WAM C77994). +j +. Female, paratype (WAM C80243). +k +. Telson (male, holotype, WAM C77994). +l +. Male, third left thoracopod (holotype, WAM C77994). +m +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–d =1 mm; e–g, j–k =0.1 mm; h–i, l =0.5 mm. + + + +THORAX +. 20–21 (HT: 20, mean: 21) segments, 19–21 (HT: 19; mean: 20) thoracopod-bearing and one to no posterior limbless segment, not reaching dorsal margin. Most thoracopod-bearing segments with spine bearing dorsal extensions. + + +THORACOPOD +III (only WAM C77994; +Fig. 16l +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp two-segmented, basal segment subequal to endopod. Exopod ventral extension subequal in extension to endopod, dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( +Fig. 16k +). 28–44 spines (HT: 35; mean: 35). First (anterior) spine enlarged. Following spines slender, elongate; most anterior spines slightly conical, posterior spines aciculate; densely and regularly spaced, usually no larger spines interspersed; increasing in length posteriorly. Dorsal margin concave or undulating. Right terminal claw more strongly curved. + + +FURCA +( +Fig. 16k +). Proximally with dorsomedial longitudinal row of 4–11 (HT: 8, mean: 9) setae, row ending distally in a single conical spine. Distal part ~⅔ of furcal length, with numerous small denticles. + + +Females + + +Overall appearance as in males. Carapace ( +Fig. 16b, d +) length +5.6–6.7 mm +, height +3.4–4.4 mm +; 34–63 growth lines, 26–55 widely spaced and 3–8 crowded; Cr/L 0.26–0.29; b/H 0.48. Angle between head and rostrum nearly straight to obtuse (~150°) ( +Fig. 16j +). Rostrum frontal margin straight or slightly undulating. Apex with right angle, not rounded; ventral margin varying from weakly concave to weakly convex. Antenna I with 8–15 small lobes, lobes smaller than in males; reaching to antenna II flagellomeres IV–V. Antenna II with 12–16 flagellomeres. Telson with 31–38 dorsal spines; left and right terminal claws usually equally curved, sometimes right stronger curved. Furca with 8–14 setae. + + +Distribution +( + +Fig. +16m + +) + + + +Ozestheria echidna + +sp. nov. +is known from four geographically relatively close localities in northwestern +Western Australia +. + + + + + +Remarks + + + +The carapace shape of + +Ozestheria echidna + +sp. nov. +( +Fig. 6 +) is distinct from that of most other species and overlaps partly with those of + +O. jiangi + +sp. nov. +, + +O. minor + +comb. nov. +, + +O. jonnae + +sp. nov. +, + +O. marthae + +sp. nov. +, + +O. selmae + +sp. nov. +, + +O. typica + +comb. nov. +, + +O. cancellata + +comb. nov. +, + +O. weeksi + +sp. nov. +, and + +O. glabra + +sp. nov. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FFC5FFB8175EFD32FDEAFDAA.xml b/data/03/FD/A6/03FDA650FFC5FFB8175EFD32FDEAFDAA.xml new file mode 100644 index 00000000000..9ba95b03d68 --- /dev/null +++ b/data/03/FD/A6/03FDA650FFC5FFB8175EFD32FDEAFDAA.xml @@ -0,0 +1,402 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria christiani + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +ED04772A-3C1F-4DCA-8725-76752888C3E3 + + + + + +Fig. 15 + + + +Ozestheria +sp. F + +– + +Schwentner +et al +. 2015a + +: figs 2, 6. + + + +Ozestheria sarsii + +– + +Schwentner +et al +. 2020 + +: figs 1–2. + + + + + +Diagnosis + + + + +Ozestheria christiani + +sp. nov. +is characterized by a short condyle and wide occipital notch; a rounded ventral carapace margin with well-defined most ventral point; carapace ornamentation with large, well-developed polygonal reticulations, many polygons intermittent and with small projections towards the polygon’s center, each polygon without secondary ornamentation (best seen under SEM); male rostrum with convex anterior margin, apex rounded, ventral margin with anterior hump then strongly concave, pointing apex downwards; 20–23 (male) antenna I lobes reaching to antenna II flagellomeres VI–VII (male); 10–11 (male) antenna II flagellomeres; 22–23 complete thorax segments; 15–18 small spines, anterior spines conical spines, posterior spines elongate and aciculate; 7–12 furcal setae. + + + +Differential diagnosis + + + + +Ozestheria christiani + +sp. nov. +can be easily differentiated from most other species of + +Ozestheria + +by the shape and ornamentation of the carapace as well as the telson spination. The morphologically most similar species are + +O. sarsii + +, + +O +. +rufa + +, + +O. paralutraria + +sp. nov. +and + +O. lutraria +. + +These can be easily differentiated by their ornamentation, as the polygonal reticulations of + +O. christiani + +are partly intermittent and with small projections towards the polygon’s center. + + + + + +Etymology + + +The species is named after Christian Pott, the partner of MH. + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +South Australia + +• +♂ +; dugout on +Wentworth Road +, about + +15 km +W of Lake + +Victoria +; +33°53′03.4″ S +, +140°58′39.1″ E +; + +13 Mar. 2011 + +; +M. Schwentner +and +B.V. Timms +leg.; +GenBank +no: +KJ705663 +( +COI +); +AM +P.91434. + + + + +Paratypes + + + + +AUSTRALIA +– + +South Australia + +• +3 ♂♂ +; same data as for holotype; +GenBank +nos: +KJ705660 +to +KJ705662 +( +COI +); +AM +P.91431 to P.91433 + +• + +1 ♂ +; same data as for holotype; +GenBank +no: +KJ705664 +( +COI +); NHMW-ZOO-CR-28479 + +. + + + +Type locality + + + +South Australia +, dugout Wentworth Road, about +15 km +W of Lake +Victoria +, +33°53′03.4″S +, +140°58′39.1″ E +. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 15a–c +). Length +8.9–10.5 mm +(HT: +8.9 mm +), height 5.0– +6.1 mm +(HT: 5.0 mm). Coloration dark brown, with lighter ventral margin and lighter (yellowish) spot on mid-frontal carapace. 21–23 (HT: 22) growth lines, 13–15 (HT: 14) widely spaced and 6–9 (HT: 8) crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, dorso-posterior corner rounded. Posterior margin broadly rounded, suboval, supracurvate to equicurvate (b/H 0.39–0.46, HT: 0.40). Ventral margin strongly curved, with clearly defined point of greatest extension. Umbo position anterior (Cr/L 0.18–0.21, HT: 0.21). + + +CARAPACE +ORNAMENTATION +( +Fig. 15d–e +). Larval valves appear smooth (probably artifact due to abrasion). Each growth band with medium to large polygonal reticulations with each polygon usually being a pentagon, hexagon or heptagon. Many polygons not fully closed and with additional lirae projecting into the polygon’s center. No secondary reticulation visible under SEM. Crowded growth bands without apparent ornamentation, but with uneven texture. Concentric ridges raised. No setae visible, under SEM setal pores in single row along posterior growth lines. + + +HEAD +( +Fig. 15f–g +). Condyle short, rounded; occipital notch wide. Condyle lacking anterobasal hump. Margin between condyle and ocular tubercle concave. Ocular tubercle well developed, forming obtuse, close to 90° angle with rostrum. Small tubercle ventrally below eye in most specimens (HT: present). Anterior margin of rostrum convex. Apex strongly rounded with obtuse angle, followed by ventral indentation (lacking in one individual). Ventral margin of rostrum with hump following indentation, then deeply concave with obtuse angle about half-length, pointing apex slightly downwards. Naupliar eye small, triangular. Antenna I long with 20–23 lobes (HT: 22), reaching to antenna II flagellomeres VI–VII (HT: VI). Antenna II with 10–11 flagellomeres (HT: 11). + + +THORAX +. 23–24 (HT: 24) segments, 22–23 (HT: 22) thoracopod-bearing and 1–2 (HT: two) posterior limbless segments not reaching dorsal margin. Dorsal extensions with numerous short spines, decreasing in number posteriorly. In posterior segments central spines stouter. + + +THORACOPOD +III (only P.91434; +Fig. 15j +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp two-segmented, basal segment shorter than endopod. Exopod ventral extension subequal or slightly longer in extension to endopod, dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( +Fig. 15h–i +). 15–18 spines (HT: 15). First (anterior) spine enlarged. Spines conical, posteriorly slightly thinner, drawn out and aciculate. Most spines small (compared to telson size), few (usually 1–3) slightly larger spines interspersed in anterior half of telson. Anterior most spines not arranged along dorsal margin but slightly lateral; spacing of spines irregular, usually widely spaced. Anterior half to two-thirds of dorsal margin nearly straight, posteriorly concavely curved. Right terminal claw more strongly curved than left. + + +FURCA +( +Fig. 15h–i +). Proximally with dorsomedial longitudinal row of 7–12 (HT: nine) setae, row ending distally in a single conical spine. Distal part ⅓ of furcal length, with numerous small denticles. + + +Female + +Unknown. + +Distribution +( +Fig. 15k +) + + +The species is known only from its +type +locality, an artificial pool in south-eastern +South Australia +. + + + + + +Remarks + + + +Currently, only males are known. + +Schwentner +et al +. (2020) + +wrongly identified this species as + +O. sarsii + +. A comparison with the respective +type +material showed that + +O. christiani + +sp. nov. +is not conspecific with + +O. sarsii +. + +This was further corroborated by the classification based on carapace shape, where the probability that + +O. sarsii + +can be assigned to + +O. christiani + +was 0% with a typicality of 0.0. The carapace shape of + +O. christiani + +( +Fig. 5 +) is distinct from that of most other species and overlaps partly with those of + +O. rubra + +and + +O. matuwa + +sp. nov. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FFC6FFBE176CFDC3FAC2FD43.xml b/data/03/FD/A6/03FDA650FFC6FFBE176CFDC3FAC2FD43.xml new file mode 100644 index 00000000000..6d80d5760c3 --- /dev/null +++ b/data/03/FD/A6/03FDA650FFC6FFBE176CFDC3FAC2FD43.xml @@ -0,0 +1,376 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria carnegiensis + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +16235DFE-FAC3-4158-B253-6F1FF8B4D8B9 + + + + + +Fig. 14 + + + + + +Diagnosis + + + + +Ozestheria carnegiensis + +sp. nov. +is characterized by an elongate, but strongly rounded condyle with a narrow occipital notch (which is wider than in most species with a long condyle); carapace ornamentation dominated by punctae, in later growth bands nodular and very inconspicuous lirae (best seen under SEM) forming between dominant punctae, concentric ridges weakly developed and punctate; male rostrum with strongly convex (sometimes undulating) anterior margin, apex rounded with nearly rectangular angle, ventral margin weakly concave; 11–15 (male) antenna I lobes reaching to antenna II flagellomeres VIII–X (male); 12–14 (male) antenna II flagellomeres; 19–20 complete thorax segments; 11–20 telsonic spines, widely and irregularly spaced, varying in length with several larger spines interspersed, anteriorly broad, conical and posteriorly thinner and elongated, increasing in size posteriorly, most spines rather large; 7–8 furcal setae. + + + +Differential diagnosis + + + + +Ozestheria carnegiensis + +sp. nov. +can be differentiated from all other species of + +Ozestheria + +by the length of the condyle (which is somewhat intermediate between the typically short or long condyle) and the carapace ornamentation, which is dominated by punctae with a very inconspicuous transition to a liraedominated ornamentation on growth bands of the secondary growth phase. Other species with such a punctae-dominated carapace ornamentation are + +O. timmsi + +sp. nov. +and + +O. frederikeae + +sp. nov. +, which both have longer condyles and a larger number of complete thorax segments (≥21 vs 19–20), antenna II flagellomeres (≥ 14 vs ≤ 14) and telsonic spines (17–31 vs 11–20), and distinct carapace shapes with more convexly curved ventral margins. + +Ozestheria gemina + +sp. nov. +has a similar condyle length but differs in carapace ornamentation (well-developed lirae, fewer and less dominant punctae), rostrum shape and telson spination (spines smaller, usually only 2–3 larger interspersed). Moreover, + +Ozestheria carnegiensis + +sp. nov. +appears to be the only species with a short, nodular epipod in most thoracopods. + + + + + +Etymology + + +The species is named after Lake Carnegie, the only locality where the species has been recorded so far. + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +Western Australia + +• +♂ +; +Lake Carnegie +(Windidda 2); +26°15′52.7″ S +, +122°16′29.23″ E +; + +4 Jun. 2020 + +; +D.J. Cale +leg.; +GenBank +no: PQ427040 ( +COI +); +WAM +C78003 +. + + + + +Paratypes + + + + +AUSTRALIA +– + +Western Australia + +• +3 ♂♂ +; same data as for holotype; +WAM +C80213 +, +WAM +C80214 +, +WAM +C80215 + +• + +1 ♂ +; same data as for holotype; NHMW-ZOO-CR-28497 + +. + + + +Type locality + + + +Western Australia +, Lake Carnegie (Windidda 2), +26°15′52.7″ S +, +122°16′29.23″ E +. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 14a–d +). Length +5.7–6.4 mm +(HT: +6.1 mm +), height 3.5–4.0 mm (HT: +3.8 mm +). Coloration light brownish, crowded growth bands lighter. ~70 (in most individuals the carapace was covered in thick layer of dirt and algae, preventing detailed study of carapace features) growth lines, ~16 widely spaced and ~55 crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, distinct dorso-posterior corner. Posterior margin broadly rounded, suboval, equicurvate to infracurvate (b/H 0.54–0.61, HT: 0.61). Ventral margin widely rounded. Umbo position submedian (Cr/L 0.31–0.35, HT: 0.33). + + +CARAPACE +ORNAMENTATION +(due to very thick layer of dirt and fungi, difficult to study; +Fig. 14e–g +). Larval valve probably punctate. In the dorsal and median part of the carapace, growth bands punctate; in later growth bands very short, inconspicuous nodular lirae appearing between punctae (best visible under SEM), lirae not reaching across full growth band width. Crowded growth bands and growth bands of the secondary growth phase with short, inconspicuous, subparallel lirae. Concentric ridges shallow, punctate, with smooth dorsal and smooth to serrate ventral margins. No obvious setae preserved (setal pores in single row along all growth lines under SEM). + + +HEAD +( +Fig. 14h +). Condyle long, distally rounded; occipital notch narrow (but wider than in most species with a long condyle). Condyle lacking anterobasal hump. Margin between condyle and ocular tubercle concave. Ocular tubercle well developed, forming obtuse angle (~100–110°) with rostrum. Anterior margin of rostrum strongly convex, sometimes undulating. Apex rounded with nearly rectangular angle. Ventral margin of rostrum with slight notch anteriorly and weakly concave mid-length. Naupliar eye large and subtriangular. Antenna I long with 11–15 lobes (HT: 11), reaching to antenna II flagellomeres VIII–X (HT: X). Antenna II with 12–14 flagellomeres (HT: 14). + + +THORAX +. 21–22 (HT: 21) segments, 19–20 (HT: 20) thoracopod-bearing and none to two (HT: one) posterior limbless segments not reaching dorsal margin. Most segments with dorsal extensions bearing several short spines, especially on last segments spines very short and inconspicuous. + + +THORACOPOD +III (only WAM C78003; +Fig. 14j +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp two-segmented, basal segment shorter than endopod. Exopod ventral extension subequal in extension to endopod, dorsal extension wide, narrowing distally, greatly overreaching epipod. Epipod very short, rounded, nodular. + + +TELSON +( +Fig. 14i +). 11–20 spines (HT: 20). First (anterior) spine enlarged. Spines on anterior ⅔ of telson broad, conical; following spines becoming longer and slightly thinner. Spines vary in size, overall large-sized, several particularly large spines interspersed. Dorsal margin anterior ⅔ straight to slightly concave, then concave. Right terminal claw more strongly curved than left. + + +FURCA +( +Fig. 14i +). Proximally with dorsomedial longitudinal row of 7–8 (HT: 7) setae, row ending distally in a single conical spine. Distal part ½ of furcal length, with numerous small denticles. + + +Female + +Unknown. + + + + +Distribution + + + + +Ozestheria carnegiensis + +sp. nov. +is known from a single locality in central +Western Australia +. + + + + +Fig. 14. + +Ozestheria carnegiensis + +sp. nov. +a–d +. Carapace. +a +. Male, holotype (WAM C78003). +b +. Male, paratype (WAM C80213). +c +. Dorsal view, male, holotype (left valve only; WAM C78003). +d +. Male, paratype (WAM C80213), SEM. +e–g +. Carapace ornamentation (male paratype WAM C80213; positions marked in d by rectangles), SEM. +e +. Mid-carapace. +f +. Posterodorsal carapace. +g +. Ventral carapace. +h–j +. Male, holotype (WAM C78003). +h +. Head (antennae not shown). +i +. Telson. +j +. Third left thoracopod. +k +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–d= 1 mm; e–g =0.05 mm; h, j =0.5 mm; i =0.1 mm. + + + + + +Remarks + + + +Currently, only males are known. In most individuals the carapace was covered by a thick layer of dirt and algae, preventing detailed study of carapace features. The extremely short epipod observed at the third thoracopod of the +holotype +appears like a damage or growth defect; however, also most other thoracopods of this and all other specimens have similarly short epipods. In some specimens these are slightly longer (about twice the length seen in the +holotype +), but still much smaller than in all other species. The claspers and a few more posterior thoracopods have the long and elongated epipods observed in all other species. + + +Because only few specimens were available, the morphological variability of the species is not well characterized. The carapace shape of + +O. carnegiensis + +sp. nov. +( +Fig. 6 +) is distinct from that of most other species and overlaps with those of + +O. beleriandensis + +sp. nov. +, + +O. typica + +comb. nov. +, and + +O. selmae + +sp. nov. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FFD9FFAE175AF922FD1DFE6C.xml b/data/03/FD/A6/03FDA650FFD9FFAE175AF922FD1DFE6C.xml new file mode 100644 index 00000000000..250532b7cba --- /dev/null +++ b/data/03/FD/A6/03FDA650FFD9FFAE175AF922FD1DFE6C.xml @@ -0,0 +1,561 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria fuersichi + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +862B6BE4-936F-4F8A-A2F0-B8BB4581CE5B + + + + + +Fig. 19 + + + +Ozestheria +sp. E + +– + +Schwentner +et al +. 2015a + +: figs 2, 6; 2020: figs 1–2. + + + + + +Diagnosis + + + + +Ozestheria fuersichi + +sp. nov. +is characterized by a long condyle and a narrow occipital notch; carapace ornamentation on larval valve and first growth bands tiny polygonal reticulation, following growth bands with widely spaced subparallel lirae, punctae (hardly visible) between lirae; male rostrum with weakly convex to straight anterior margin, apex pointed with acute angle (~60°) rarely rounded, ventral margin concavely curved; female rostrum with concave anterior margin, apex weakly drawn out and pointed with acute angle (~70–80°), ventral margin weakly convex to straight; 13–14 (male) or 10–16 (female) antenna I lobes reaching to antenna II flagellomeres VI–VIII (male) or IV–V (female); 10–12 (male) or 11–14 (female) antenna II flagellomeres; 22 complete thorax segments; 19–25 telsonic spines, spines short with 1–4 larger spines interspersed, anterior spines conical, posterior spines increase slightly in size and are longer, drawn out, aciculate and more closely spaced; 6–10 furcal setae. + + + +Differential diagnosis + + + + +Ozestheria fuersichi + +sp. nov. +can be differentiated from many other species of + +Ozestheria + +by the narrow occipital notch and long condyle in combination with the carapace ornamentation (dominated by punctate ornamentation dorsally on the carapace, transitioning to distinct, subparallel lirae during ontogeny), except from + +O. cancellata + +comb. nov. +, + +O. minor + +comb. nov. +, + +O +. +typica + +comb. nov. +, + +O. jonnae + +sp. nov. +, + +O. marthae + +sp. nov. +, + +O. selmae + +sp. nov. +, + +O. radiata + +sp. nov. +, + +O. bourkensis + +sp. nov. +, + +O. rincewindi + +sp. nov. +, + +O. barcaldinensis + +sp. nov. +, + +O. ngamurru + +sp. nov. +, + +O. beleriandensis + +sp. nov. +, + +O. quinlanae + +sp. nov. +, + +O. glabra + +sp. nov. +, + +O. pilbarensis + +sp. nov. +and + +O. weeksi + +sp. nov. +, and differentiating these species can be difficult. + +Ozestheria fuersichi + +differs from these species in having polygonal ornamentations on the ontogenetically earliest growth bands (shared only with + +O. pilbarensis + +), widely spaced lirae that are not continuous but a row of nodular structures (only seen under SEM), in the shape of the male rostrum (which is more elongated and slender than in the other species), the shape of the female rostrum (with a concave anterior margin which curves the apex anteriorly) and the spination of the telson with several distinctly larger spines interspersed (though this is also present in some of the other species). + + + + + +Etymology + + +The species is named after German paleontologist Franz T. Fürsich, honoring his significant contributions to invertebrate paleontology, which sparked MH’s curiosity in fossil clam shrimp. + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +Queensland + +• +♂ +; +Barcaldine Region +, pool next to +Lake Dunn +; +22°36′16.4″ S +, +145°40′21.8″ E +; + +14 Feb. 2010 + +; +M. Schwentner +, +C. Sieves +and and +B.V. Timms +leg.; +GenBank +no: +KJ705655 +( +COI +); +AM +P.91426. + + + + +Paratypes + + + + +AUSTRALIA +– + +Queensland + +• +1 ♂ +, +1 ♀ +; same data as for holotype; +GenBank +nos: +KJ705653, KJ705656 +( +COI +); +AM +P.91424, P.91427 + +• + +1 ♀ +; same data as for holotype; +GenBank +no: +KJ705654 +( +COI +); +NHMW- +ZOO-CR-28478 + +. + + + +Other material examined + + + + +AUSTRALIA +– + +Queensland + +• + +1 +♂ + +; old dugout close to +Lake Dunn +; +22°36′12.9″ S +, +145°40′26.6″ E +; + +14 Feb. 2010 + +; +M. Schwentner +, +C. Sieves +and +B.V. Timms +leg.; +AM +P.91420 + +• + +3 ♀♀ +; old borrow pit, +Monklands Road +; +23°37′34.8″ S +, +146°21′11.7″ E +; + +14 Feb. 2010 + +; +M. Schwentner +, +C. Sieves +and +B.V. Timms +leg.; +AM +P.91421 to P.91423 + +. + + + +Type locality + + + +Australia +, +Queensland +, Barcaldine Region, pool next to Lake Dunn, +22°36′16.4″ S +, +145°40′21.8″ E +. + + + + + +Description + + + +Males + + +CARAPACE +( +Fig. 19a–c +). Length +3.7–4.3 mm +(HT: +3.7 mm +), height +2.3–2.7 mm +(HT: +2.3 mm +). Coloration light brown, outer margin lighter. 19–21 growth lines (HT: 19), 15–18 (HT: 16) widely spaced and 3–6 (HT: 3) crowded. + + +CARAPACE +SHAPE +. Dorsal margin straight, distinct dorso-posterior corner. Posterior margin broadly rounded, weakly extending posteriorly, supracurvate to equicurvate (b/H 0.46–0.49, HT: 0.46). Ventral margin nearly straight. Umbo position submedian (Cr/L 0.26–0.27, HT: 0.26). + + +CARAPACE +ORNAMENTATION +( +Fig. 19d–e +). Larval valve and following few growth bands covered in mesh of shallow, poorly visible polygonal reticulations, mostly tetragons, pentagons and hexagons. Following growth bands with widely spaced, subparallel lirae; lirae dorsally often anastomosing or confluent, with shorter lirae intercalating on the ventral part of each growth band (predominantly in the posterior region of the carapace), under SEM lirae not continuous but formed by a series of nodular structures; punctae interspersed between lirae (but poorly visible under incident light). Crowded growth bands lacking apparent ornamentation (punctate under SEM). Concentric ridges raised. Setae filiform, under SEM a single row of setal pores along all growth lines. + + +HEAD +( + +Fig. +19g + +). Condyle long, distally rounded; occipital notch narrow. Condyle with weakly developed or absent anterobasal hump. Margin between condyle and ocular tubercle straight to weakly concave. Ocular tubercle weakly developed, forming obtuse angle with rostrum (~120°). Anterior margin of rostrum weakly convex to straight (HT: weakly convex). Ventral margin of rostrum concavely curved with obtuse angle about half-length, pointing apex slightly downwards; apex pointed, acute (~60°), rarely rounded. Naupliar eye large and elongated, subrectangular with irregular margins to subtriangular or suboval. Antenna I with 13–14 lobes (HT: 14), reaching to antenna II flagellomeres VI–VIII (HT: VIII). Antenna II with 10–12 flagellomeres (HT: 11). + + +THORAX +. 22–23 (HT: 22) segments, 22 thoracopod-bearing and none to one (HT: none) posterior limbless segment not reaching dorsal margin. Dorsal armature with thin and elongated spines, becoming shorter and stouter in posterior segments. + + +THORACOPOD +III (only P.91426; +Fig. 19f +). Endite I short and curved dorsally. Endites II–V broad, decreasing in size. Endite V palp two-segmented, basal segment sub-equal to endopod. Exopod ventral extension shorter in extension to endopod, dorsal extension wide, narrowing distally, overreaching epipod. Epipod long, cylindric. + + +TELSON +( +Fig. 19i +). 19–23 spines (HT: 20). First (anterior) spine enlarged. Spines short, conical, subequal in size and spacing, slightly increasing in size posteriorly, with 1–4 (HT: 4) larger spines interspersed; posterior most spines slightly larger, drawn out and aciculate. Dorsal margin nearly straight. Terminal claws subequally curved. + + +FURCA +( +Fig. 19i +). Proximally with dorsomedial longitudinal row of 8–10 (HT: eight) setae, row ending distally in a single conical spine. Distal part of ~⅔ of furca length, widely curved, with numerous small denticles. + + +Females + + +Very similar to males. Carapace length +4.4– 5.1 mm +, height +2.7 – 3.2 mm +. 17–19 growth lines, of these 15–18 widely spaced and 0–3 crowded; Cr/L 0.24–0.28 and b/H 0.44–0.49. Ocular tubercle and rostrum form nearly straight angle ( +Fig. 19h +). Anterior margin of rostrum concave, apex weakly drawn out, pointed, with acute angle (~70–80°), ventral margin weakly convex to straight. Antenna I with 10–16 indistinct lobes, reaching to antenna II flagellomeres IV–V. Antenna II with 11–14 flagellomeres. Telson 19–25 spines ( +Fig. 19j +), shape and arrangement as in males. Furca bearing 6–10 setae. + + +Distribution +( +Fig. 19k +) + + +The species occurs in central +Queensland +, in the northern regions of the Cooper Creek catchment. + + + + +Fig. 19. + +Ozestheria fuersichi + +sp. nov. +a +. Carapace (male, holotype, P.91426). +b +. Carapace (male paratype, P.91427), SEM. +c +. Carapace, dorsal view (right valve only, male, holotype, P.91426). +d–e +. Carapace ornamentation (male paratype, P.91427; positions marked in b by rectangles), SEM. +d +. Dorsal carapace. +e +. Mid-carapace. +f +. Third left thoracopod (male, holotype, P.91426). +g–h +. Head (antennae not shown). +g +. Male, holotype (P.91426). +h +. Female, paratype (P.91424). +i–j +. Telson. +i +. Male, paratype (P.91427). +j +. Female, paratype (P.91424). +k +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–c, g= 0.5 mm; d–f, i–j =0.1 mm; h =0.2 mm. + + + + + +Remarks + + + +The carapace shape of + +Ozestheria fuersichi + +sp. nov. +( +Fig. 6 +) is distinct from all that of other species in the analysis of short-condyled species ( +Fig. 5 +). In the analysis of long-condyled species, + +O. fuersichi + +overlaps partly with + +O. minor + +comb. nov. +, + +O +. +selmae + +sp. nov. +, + +O. cancellata + +comb. nov. +and + +O. weeksi + +sp. nov. +and fully with + +O. setifera + +sp. nov. + + + + \ No newline at end of file diff --git a/data/03/FD/A6/03FDA650FFDBFFA21755FEECFAC2F954.xml b/data/03/FD/A6/03FDA650FFDBFFA21755FEECFAC2F954.xml new file mode 100644 index 00000000000..4705144546e --- /dev/null +++ b/data/03/FD/A6/03FDA650FFDBFFA21755FEECFAC2F954.xml @@ -0,0 +1,436 @@ + + + +Revision of the Australian Ozestheria Schwentner & Richter, 2015 (Crustacea: Branchiopoda: Spinicaudata) fauna, with the descriptions of 27 new species + + + +Author + +Schwentner, Martin +720F61F1-5EAC-42D2-A66C-B91B439DB2A1 +Naturhistorisches Museum Wien, 1010 Vienna, Austria. +martin.schwentner@nhm.at + + + +Author + +Hethke, Manja +288F3F34-AD0D-4DF7-AD5D-434FACC207E5 +LWL-Museum für Naturkunde, Westfälisches Landesmuseum mit Planetarium, Referat Paläontologie, D- 48161 Münster, Germany and Fachrichtung Paläontologie, Department Geowissenschaften, Freie Universität Berlin, D- 12249 Berlin, Germany. +manja.hethke@lwl.org + +text + + +European Journal of Taxonomy + + +2025 + +2025-05-23 + + +992 + + +1 +172 + + + + +https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2905/13169 + +journal article +10.5852/ejt.2025.992.2905 +2118-9773 +24F7D1C9-A2DA-4F31-B6FE-7A7DDF54D202 + + + + + + +Ozestheria frederikeae + +sp. nov. + + + + +urn:lsid:zoobank.org:act: +115FEB5B-8E8B-4979-B524-087E7023A7C5 + + + + + +Fig. 18 + + + +Ozestheria +sp. H + +2 – + +Schwentner +et al +. 2015a + +: figs 2, 6. + + + + + +Diagnosis + + + + +Ozestheria frederikeae + +sp. nov. +is characterized by a long condyle and a narrow occipital notch; carapace ornamentation dominated by pit-like punctae, in later growth bands lirae forming between punctae; female rostrum with undulating anterior margin, apex rounded, nearly rectangular (in some individuals protruding), ventral margin straight to convex (sometimes concave mid-length); 12–14 (female) antenna I lobes reaching to antenna II flagellomeres III–IV (female); 14–17 (female) antenna II flagellomeres; 21–22 complete thorax segments; 22–31 telsonic spines, spines small with ~2 larger spines interspersed, anteriorly conical and posteriorly thin and aciculate; 6 furcal setae. + + + +Differential diagnosis + + + + +Ozestheria frederikeae + +sp. nov. +can be differentiated from all other species of + +Ozestheria + +by the combination of the carapace ornamentation (dominated by pit-like punctae), carapace shape, rostrum shape and telson spination. Species with similar carapace ornamentation are + +O. timmsi + +sp. nov. +and + +O. carnegiensis + +sp. nov. + +Ozestheria timmsi + +differs by having a larger carapace ( +7.9–11.5 mm +vs 6.1– 7.0 mm), more antenna flagellomeres (15–22 vs 14–17) and more complete thorax segments (23–24 vs 21). + +Ozestheria carnegiensis + +differs by having a slightly shorter and more rounded condyle, less distinct lirae on the carapace, fewer antenna flagellomeres (≤ 14 vs ≥ 14) and larger and fewer (11–20 vs 22–31) telsonic spines. + + + + + +Etymology + + +The species is named after Frederike Korth, the wife of MS – for everything. + + + + +Type material + + + + +Holotype + + + + +AUSTRALIA +– + +Queensland + +• +♀ +; turbid pool +80 km +S of +Charleville +; +27°04′59.0″ S +, +146°01′42.2″ E +; + +17 Feb. 2010 + +; +M. Schwentner +, +C. Sieves +and +B.V. Timms +leg.; +GenBank +no: +KJ705769 +( +COI +); +AM +P.91541. + + + + +Paratypes + + + + +AUSTRALIA +– + +Queensland + +• +3 ♀♀ +; same data as for holotype; +GenBank +nos: +KJ705767, KJ705770, KJ705771 +( +COI +); +AM +P. 91539, P.91542, P.91543 + +• + +1 ♀ +; same data as for holotype; +GenBank +no: +KJ705768 +( +COI +); NHMW-ZOO-CR-28481 + +. + + +Additional material +(not examined) + + + +AUSTRALIA +– + +Queensland + +• 5 juvs; big pool on +Meandarra Road +; +27°22′43.9″ S +, +150°01′18.1″ E +; + +12 Feb. 2010 + +; +M. Schwentner +, +C. Sieves +and +B.V. Timms +leg.; +AM +P.91534 to P.91538 + +. + + + +Type locality + + + +Australia +, +Queensland +, turbid pool +80 km +S of Charleville, +27°04′59.0″ S +, +146°01′42.2″ E +. + + + + + +Description + + + +Females + + +CARAPACE +( +Fig. 18a–c +). Length 6.1–7.0 mm (HT: 7.0 mm), height 4.3–4.7 (HT: +4.6 mm +). Coloration reddish- or orange-brown, crowded growth bands lighter. 46–52 (HT: 46) growth lines, 16–22 (HT: 20) widely spaced and 26–32 (HT: 26) crowded; in some individuals ~2–3 widely spaced growth bands recurrently interspersed between crowded growth bands (secondary growth phase). + + +CARAPACE +SHAPE +. Dorsal margin straight, dorso-posterior corner rounded. Posterior margin widely rounded, suboval, equicurvate to infracurvate (b/H 0.50–0.57, HT: 0.50). Ventral margin widely and equally rounded. Umbo anterior to submedian (Cr/L 0.23–0.26, HT: 0.24). + + + +Fig. 18. + +Ozestheria frederikeae + +sp. nov. +a–c +. Carapace. +a +. Female, holotype (P.91541). +b +. Female, paratype (P.91542), SEM. +c +. Female, holotype, dorsal view (only left valve shown; AM P:91541). +d–g +. Carapace ornamentation of female paratype (P.91542; positions marked in b by rectangles), SEM. +d +. Dorsal carapace. +e +. Mid-carapace. +f +. Posterodorsal carapace. +g +. Ventral carapace. +h–i +. Head (antennae not shown). +h +. Female, paratype (P.91542), SEM. +i +. Female, holotype (P.91541). +j–k +. Telson. +j +. Female, paratype (P. 91542), SEM. +k +. Female, holotype (P.91541). +m +. Distribution map (produced in ArcMap 10.7; see caption Fig. 7 for sources). Scale bars: a–c=0.5 mm; d–g =0.05 mm; h–k=0.2 mm. + + + +CARAPACE +ORNAMENTATION +( +Fig. 18d–g +). Larval valve and first few growth bands smooth to finely granulated, under SEM finely punctate. Subsequent growth bands with irregular, poorly defined pits (punctae-like). From about mid-carapace, space between punctae raised to lirae, increasing in size on subsequent growth bands (giving the punctae an elongate, oval appearance under SEM). Crowded growth bands often too closely spaced for apparent ornamentation, else poorly defined lirae. Concentric ridges slightly raised (dorsally serrated under SEM and with small nodules in moniliform row). Setae filiform, preferentially preserved close to ventral margin; under SEM a single row of setal pores along all growth lines. + + +HEAD +( +Fig. 18h–i +). Condyle long, distally rounded; occipital notch narrow. Condyle with weak anterobasal hump. Margin between condyle and ocular tubercle straight. Ocular tubercle weakly developed, forming obtuse (~120°) to nearly straight angle with rostrum. Anterior margin of rostrum undulating: dorsally weakly convex, ventrally weakly concave. Apex rounded, nearly rectangular (in some individuals protruding). Ventral margin of rostrum straight (HT) to convex (sometimes concave mid-length). Naupliar eye elongated, subtriangular to subrectangular (rounded edges).Antenna I long with 12–14 lobes (HT: 14), reaching to antenna II flagellomeres III–IV (HT: IV). Antenna II with 14–17 flagellomeres (HT: 14). + + +THORAX +. 21–22 (HT: 21) segments, 21 thoracopod-bearing and none to one (HT: none) posterior limbless segments not reaching dorsal margin. Mid and posterior thoracopod-bearing segments with spine bearing dorsal extensions. Dorsal extensions increase in size posteriorly over successive segments. Spines short and stout, in posterior segments central spines increase in size and the total number of spines decreases. + + +TELSON +( +Fig. 18j–k +). 22–31 spines (HT: 29). First (anterior) spine enlarged. Spines mostly tiny, conical; posterior ⅓ with slightly thinner, larger and more drawn-out spines. Spines subequal in length and spacing, few (usually two) slightly larger spines interspersed in anterior ⅔ of spines. One or a few anterior spines not arranged along dorsal margin but slightly lateral. Dorsal margin nearly straight, not concavely curved. Right terminal claw more strongly curved than left, claws notably slender. + + +FURCA +( +Fig. 18j–k +). Proximally with dorsomedial longitudinal row of 6 setae, row ending distally in a single conical spine. Distal part ½–⅔ of furcal length, with numerous small denticles. + + +Males + +Unknown. + +Distribution +( + +Fig. +18m + +) + + +The species is known only from two localities in southern +Queensland +close to the border to +New South Wales +. These populations are separated by ~ +400 km +; thus, further populations can be assumed in this region. + + + + + +Remarks + + + +So far, only females have been collected. The second population identified by + +Schwentner +et al +. (2015a) + +yielded only juveniles. + + +Because only few specimens were available, the morphological variability of the species is not well characterized. The carapace shape of + +O. frederikeae + +sp. nov. +( +Fig. 6 +) is distinct from that of most other species and partly overlaps with + +O. timmsi + +sp. nov. +(marginally), + +O. jonnae + +sp. nov. +, + +O. cancellata + +comb. nov. + + + + \ No newline at end of file