Add updates up until 2024-11-05 16:14:36
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<emphasis id="2FD03229FAB5114A43974A09C948F43C" italics="true">Thomasomys igor</emphasis>
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<emphasis id="A65A146B601D2026D3FF79DF7FA08B14" italics="true">Thomasomys caudivarius</emphasis>
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–
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<bibRefCitation id="79E5FFC3AC204C6FC0FDB35725067881" author="Salazar-Bravo and Yates" editor="Kelt D & Kaspin D" firstAuthor="Salazar-Bravo" journalOrPublisher="Pearson. University of California Publications in Zoology, Berkeley" pagination="747-774" refId="B65" refString="Salazar-Bravo J, Yates TL (2007) A new species of Thomasomys (Cricetidae: Sigmodontinae) from central Bolivia. In: Kelt D, Kaspin D (Eds) The Quintessential Naturalist: Honoring the Life and Legacy of Oliver P. Pearson. University of California Publications in Zoology, Berkeley, 747–774." title="A new species of Thomasomys (Cricetidae: Sigmodontinae) from central Bolivia." volumeTitle="The Quintessential Naturalist: Honoring the Life and Legacy of Oliver P." year="2007">Salazar-Bravo and Yates (2007</bibRefCitation>
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: fig. 2); partim, not
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<emphasis id="6CC860D3C25465EAF6BBE6DFAFC24594" italics="true">Thomasomys caudivarius</emphasis>
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<bibRefCitation id="B5D03429E6D579DB45F006A4FAFC0EA1" author="Anthony HE" journalOrPublisher="American Museum Novitates" pagination="1-14" refId="B4" refString="Anthony HE (1923) Preliminary report on Ecuadorean mammals No. 3. American Museum Novitates 55: 1 - 14." title="Preliminary report on Ecuadorean mammals No. 3." volume="55" year="1923">Anthony, 1923</bibRefCitation>
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.
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<emphasis id="41E363C9BA68941CC9848CB8267E3411" italics="true">Thomasomys caudivarius</emphasis>
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</taxonomicName>
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–
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<bibRefCitation id="B5E5BB95E1179DB7CC19AEC1B4538BC5" DOI="10.1371/journal.pone.0183070" author="Steppan and Schenk" firstAuthor="Steppan" refId="B76" refString="Steppan SJ, Schenk JJ (2017) Muroid rodent phylogenetics: 900 - species tree reveals increasing diversification rates. PloS One 12: e 0183070. https://doi.org/10.1371/journal.pone.0183070" year="2017">Steppan and Schenk (2017</bibRefCitation>
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: fig. 2); partim, not
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<emphasis id="F29EB3C24E407B775B18EA52AAA89EB4" italics="true">Thomasomys caudivarius</emphasis>
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<bibRefCitation id="D5BC017B8610C52C337D6500FA9700D8" author="Anthony HE" journalOrPublisher="American Museum Novitates" pagination="1-14" refId="B4" refString="Anthony HE (1923) Preliminary report on Ecuadorean mammals No. 3. American Museum Novitates 55: 1 - 14." title="Preliminary report on Ecuadorean mammals No. 3." volume="55" year="1923">Anthony, 1923</bibRefCitation>
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<emphasis id="EF80B3294D31C17608905D8EE88FE15C" italics="true">Thomasomys</emphasis>
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sp. 1
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–
|
||||
<bibRefCitation id="6FB176F641D0AF8E055C1A425D752276" DOI="10.31687/saremMN.19.26.2.0.04" author="Brito J & Tinoco N & Curay J & Vargas R & Reyes-Puig C & Romero V & Pardiñas UF" etAl="et al." firstAuthor="Brito" journalOrPublisher="Mastozoología Neotropical" pagination="308-330" refId="B15" refString="Brito J, Tinoco N, Curay J, Vargas R, Reyes-Puig C, Romero V, Pardiñas UF (2019) Diversidad insospechada en los Andes de Ecuador: filogenia del grupo “ cinereus ” de Thomasomys y descripción de una nueva especie (Rodentia, Cricetidae). Mastozoología Neotropical 26: 308–330. https://doi.org/10.31687/saremMN.19.26.2.0.04" title="Diversidad insospechada en los Andes de Ecuador: filogenia del grupo “ cinereus ” de Thomasomys y descripción de una nueva especie (Rodentia, Cricetidae)." volume="26" year="2019">Brito et al. (2019)</bibRefCitation>
|
||||
</treatmentCitation>
|
||||
.
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E6ECI" type="Holotype">
|
||||
<paragraph id="82164F3396B702FEC6116D2A6C7E6962">
|
||||
<heading id="84030FEFAFAB58CC643BEC0D0F9B0D99" reason="title">
|
||||
<typeStatus id="E1CC2A1F8C06D091F60E6B855A2E0703" type="holotype">Holotype</typeStatus>
|
||||
.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="3BA8BA2016D2E4E9372D2EE298EDEC3B">
|
||||
<materialsCitation id="B9E7AAE1F3A52CFA4BEEB76109D1C34E" collectingDate="1994-07-15" collectionCode="MSB" collectorName="Jorge Salazar-Bravo" country="Ecuador" elevation="2875" latitude="-1.716667" location="Cruz de Lizo" longLatPrecision="1" longitude="-78.98333" municipality="Museum of Southwestern Biology" specimenCode="MECN 702, NK 27680" specimenCount="1" specimenCount-female="1" stateProvince="Provincia de Bolivar" typeStatus="holotype">
|
||||
<specimenCode id="C253739BCB4AF79B38FCEA06DB98A876" collectionCode="MECN" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">
|
||||
<named-content id="28A7A40E1A49ECEFBC6C0E2914E71278" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Ciencias Naturales">MECN</named-content>
|
||||
702
|
||||
</specimenCode>
|
||||
, an
|
||||
<specimenCount id="65061F8A1D899E59575AFC4F1138EC4B" count="1" type="female">adult female</specimenCount>
|
||||
collected
|
||||
<date id="6EF1AAABDDBE8B4599C77774F9451EAC" value="1994-07-15">
|
||||
<collectingDate id="2923E2FA371E13A94281DA5D06E6F461" value="1994-07-15">15 July 1994</collectingDate>
|
||||
</date>
|
||||
, by
|
||||
<collectorName id="F24F1916CB6AF9A43504337A6A6795E9">Jorge Salazar-Bravo</collectorName>
|
||||
(field number JSB 716), originally catalogued at Museum of Southwestern Biology
|
||||
<named-content id="0F19A5B12F88BC6F8488F13D2B70213B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/university-new-mexico" xlink_title="Museum of Southwestern Biology at the University of New Mexico">
|
||||
<collectionCode id="F260530113479F7056E99A6FE63A3851" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" type="Museum">MSB</collectionCode>
|
||||
</named-content>
|
||||
: Mamm: 70717 (
|
||||
<specimenCode id="9B069AFE514899B89A67E0F2426FF763" collectionCode="NK">NK 27680</specimenCode>
|
||||
), preserved as dry skin, skull and postcranial skeleton.
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EVFCI" type="description">
|
||||
<paragraph id="9F43569710C8B9431880F6592D9528A0">
|
||||
<heading id="50E410F52F0B6E885FC6D92A65B1B95D" reason="title">
|
||||
Measurements of the
|
||||
<typeStatus id="0A0251A47E90483AD69A9AC20AE9BBA0" type="holotype">holotype</typeStatus>
|
||||
(in mm, mass in g).
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="7C6F271F2332F9A5C082F2A6E36C8ECF">
|
||||
<abbrev id="ABBRID0E2FCI" xlink_title="head and body length">HB</abbrev>
|
||||
= 118;
|
||||
<abbrev id="ABBRID0E6FCI" xlink_title="tail length">TL</abbrev>
|
||||
= 160;
|
||||
<abbrev id="ABBRID0EDGCI" xlink_title="hind foot length">HF</abbrev>
|
||||
= 27, E = 19; W = 30;
|
||||
<abbrev id="ABBRID0EHGCI" xlink_title="Condylo-incisive length">CIL</abbrev>
|
||||
= 27.44;
|
||||
<abbrev id="ABBRID0ELGCI" xlink_title="length of upper diastema">LD</abbrev>
|
||||
= 8.49;
|
||||
<abbrev id="ABBRID0EPGCI" xlink_title="crown length of maxillary toothrow">LM</abbrev>
|
||||
= 4.74;
|
||||
<abbrev id="ABBRID0ETGCI" xlink_title="breadth of the first maxillary molar">
|
||||
<collectionCode id="6D90125350B46EEA36D8A39EF457DD61" country="United Kingdom" name="Bristol Museum">BM</collectionCode>
|
||||
1
|
||||
</abbrev>
|
||||
= 1.6;
|
||||
<abbrev id="ABBRID0EXGCI" xlink_title="length of incisive foramina">LIF</abbrev>
|
||||
= 5.77;
|
||||
<abbrev id="ABBRID0E2GCI" xlink_title="breadth of incisive foramina">BIF</abbrev>
|
||||
= 1.75;
|
||||
<abbrev id="ABBRID0E6GCI" xlink_title="breadth of the palatal bridge">BPB</abbrev>
|
||||
= 3.53;
|
||||
<abbrev id="ABBRID0EDHCI" xlink_title="breadth of zygomatic plate">BZP</abbrev>
|
||||
= 2.14;
|
||||
<abbrev id="ABBRID0EHHCI" xlink_title="length of incisive foramina">LIF</abbrev>
|
||||
= 5.19;
|
||||
<abbrev id="ABBRID0ELHCI" xlink_title="zygomatic breadth">ZB</abbrev>
|
||||
= 15.15;
|
||||
<abbrev id="ABBRID0EPHCI" xlink_title="depth of upper incisor">DI</abbrev>
|
||||
= 1.45;
|
||||
<abbrev id="ABBRID0ETHCI" xlink_title="breadth across both upper incisor tips">BIT</abbrev>
|
||||
= 1.81;
|
||||
<abbrev id="ABBRID0EXHCI" xlink_title="length of rostrum">LR</abbrev>
|
||||
= 9.51;
|
||||
<abbrev id="ABBRID0E2HCI" xlink_title="length of nasals">LN</abbrev>
|
||||
= 11.61;
|
||||
<abbrev id="ABBRID0E6HCI" xlink_title="breadth of rostrum">
|
||||
<collectionCode id="F39643BB91C3F2D557A9001C0C3E1B31" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||||
</abbrev>
|
||||
= 5.18;
|
||||
<abbrev id="ABBRID0EDICI" xlink_title="orbital fossa length">OL</abbrev>
|
||||
= 7.73;
|
||||
<abbrev id="ABBRID0EHICI" xlink_title="breadth of mesopterygoid foramen">BMF</abbrev>
|
||||
= 2.43;
|
||||
<abbrev id="ABBRID0ELICI" xlink_title="breadth between occipital condyles">BCO</abbrev>
|
||||
= 7, 13;
|
||||
<abbrev id="ABBRID0EPICI" xlink_title="braincase breadth">BCB</abbrev>
|
||||
= 13.31;
|
||||
<abbrev id="ABBRID0ETICI" xlink_title="length of mandible">LMN</abbrev>
|
||||
= 15.07;
|
||||
<abbrev id="ABBRID0EXICI" xlink_title="depth of mandibular ramus">DR</abbrev>
|
||||
= 3.25. External and craniodental of additional specimens are listed in Table
|
||||
<tableCitation id="1A8D4688390415E9A7381209A5B476E5" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of morphometric measurements of all specimens in mm. Species names are accompanied by the number of analyzed individuals between parentheses. Minimum and maximum values are accompanied by mean and standard deviation between parentheses. Abbreviations of characters are detailed in the text." httpUri="http://table.plazi.org/id/B5A75659D53B773ADE5A518BE89C41B5" tableUuid="B5A75659D53B773ADE5A518BE89C41B5">4</tableCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E6ICI" type="type locality">
|
||||
<paragraph id="4216A88D280D4BE85FC45DE7F07DC571">
|
||||
<heading id="295218F0B6FDC2FBB2CA6D44FA18E5C4" reason="title">Type locality.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="A1F87ABE6F59459814215D331733991A">
|
||||
<collectingCountry id="43FFC8468656E2D3415295306717BDC0" name="Ecuador">Ecuador</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="5298EE1710DDF06140CC7EEFD2B32699" country="Ecuador" name="Bolivar">Provincia de Bolívar</collectingRegion>
|
||||
,
|
||||
<collectingMunicipality id="81666D8534399085622A47F1686114AF">Río Tatahuazo</collectingMunicipality>
|
||||
,
|
||||
<locationDeviation id="04E18F021E4C4AA2E37F7349373FC3BD">
|
||||
<quantity id="DD732D0FD504803A5E9F3EF0A260CBAB" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="km" value="2.5">2.5 km</quantity>
|
||||
NE de
|
||||
</locationDeviation>
|
||||
<location id="607920871F5A514B488BE33DAEAAA5C2" LSID="urn:lsid:plazi:treatment:8D7253C2E25D570D9C60CC83C0F61931:607920871F5A514B488BE33DAEAAA5C2" country="Ecuador" latitude="-1.716667" longLatPrecision="1" longitude="-78.98333" municipality="Rio Tatahuazo" name="Cruz de Lizo" stateProvince="Provincia de Bolivar">Cruz de Lizo</location>
|
||||
(
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EIJCI" specific-use="{"type":"Point","coordinates":[-78.983330,-1.716667]}">
|
||||
<geoCoordinate id="57A39E045A71E4C8BBBC45EE95185585" degrees="1.716667" direction="south" orientation="latitude" precision="1" value="-1.716667">– 1.716667</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="1085BE848B690CAEBE923B671E78B90A" degrees="78.98333" direction="west" orientation="longitude" precision="1" value="-78.98333">– 78.98333</geoCoordinate>
|
||||
</named-content>
|
||||
, WGS 84 coordinates taken by GPS at the collection site, elevation
|
||||
<quantity id="918D5FF1CC1F179D5D1ABA943D142928" metricMagnitude="3" metricUnit="m" metricValue="2.875" unit="m" value="2875.0">
|
||||
<elevation id="8F9FAB438DD1FDD2CF0FDF1CE2CD6066" metricMagnitude="3" metricUnit="m" metricValue="2.875" unit="m" value="2875.0">2,875 m</elevation>
|
||||
</quantity>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ENJCI" type="Paratypes">
|
||||
<paragraph id="196A38C8DB43EDA37BD4CB7FAC1ABB0A">
|
||||
<heading id="D80EB735FEFCB8E3030C2FEE76DC885B" reason="title">
|
||||
<typeStatus id="F50C28F2E963DD438B4B3597066A62EE" type="paratype">Paratypes</typeStatus>
|
||||
.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="68DDD3D0BC716096CAFE769FAC7100E6">
|
||||
<named-content id="5D8AEA6DFF3C06D3A74B99F3B98723EC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Ciencias Naturales">
|
||||
<collectionCode id="85204E9884BB6AFA742BD1376C16F64B" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
|
||||
</named-content>
|
||||
700, juvenile male,
|
||||
<named-content id="C099A79BB35726B0CE316CD98463610E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Ciencias Naturales">
|
||||
<collectionCode id="90834950243217200E05BF7013416FC5" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
|
||||
</named-content>
|
||||
701, adult male, preserved as dry skins and cleaned skulls, by Jorge Salazar-Bravo on
|
||||
<date id="FC8A3C1DE508E69E06B023E89F964D7F" value="1994-07-15">15 July 1994</date>
|
||||
.
|
||||
<named-content id="491ED7BFB0FA8B537B915E108BFF1347" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Ciencias Naturales">
|
||||
<collectionCode id="FA931EAF3256797209B7ABFEBD803B57" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
|
||||
</named-content>
|
||||
703, male adult, preserved as dry skins and cleaned skulls, by Nelson Monar.
|
||||
<abbrev content-type="institution" id="ABBRID0ELKCI" xlink_title="Museo de la Escuela Politécnica Nacional">
|
||||
<collectionCode id="18D579F5370034A628123FA46EC4CC2D">MEPN</collectionCode>
|
||||
</abbrev>
|
||||
6203, adult female preserved as dry skin and cleaned skull, by Igor Castro on
|
||||
<date id="1D40308AF6D722ABE6C2757FFCF06F0C" value="1994-07-17" valueMax="1994-07-18" valueMin="1994-07-17">17 and 18 July 1994</date>
|
||||
, respectively, collected at
|
||||
<quantity id="39217186A0BF05818C90122329FDCCB4" metricMagnitude="3" metricUnit="m" metricValue="2.5" unit="km" value="2.5">2.5 km</quantity>
|
||||
NE de Cruz de Lizo the same locality of the
|
||||
<typeStatus id="22AAB5CF4A661EF2078C5F5B58254A74" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
<named-content id="7A80A66B5C89EC2223717C02DB60AA4E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/university-new-mexico" xlink_title="Museum of Southwestern Biology at the University of New Mexico">
|
||||
<collectionCode id="9647BBC77ED8019836AD30C8E25A1798" country="USA" httpUri="http://grbio.org/cool/km06-gtbn" name="Museum of Southwestern Biology" type="Museum">MSB</collectionCode>
|
||||
</named-content>
|
||||
70712, juvenile female, collected at Cruz de Lizo (
|
||||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2KCI" specific-use="{"type":"Point","coordinates":[-78.950000,-1.716667]}">– 1.716667, – 78.95000</named-content>
|
||||
,
|
||||
<quantity id="9A454A50A45BC93E418813288369A521" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="m" value="3000.0">3,000 m</quantity>
|
||||
) preserved as dry skins and cleaned skulls, by I. Castro on
|
||||
<date id="2D466E09799E43E686FCAC7D8316DCEB" value="1994-07-18">18 July 1994</date>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EALCI" type="etymology">
|
||||
<paragraph id="581CA454A6E3EB6AA6EB5C7FE52E5B23">
|
||||
<heading id="B07F51C9563F571AF0E1CA80C9D41CDC" reason="title">Etymology.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="57736C1FB171072D54598711697DCF4C">
|
||||
This species is named in honor of Igor A. Castro Revelo (1971–2022), Ecuadorian, prominent collector of rodents and curator of the mammal collection at the Museo Ecuatoriano de Ciencias Naturales (
|
||||
<named-content id="30B822AF87F092944AEA6AD328F011CD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Ciencias Naturales">
|
||||
<collectionCode id="FC053E4F2988507B1A24AA001EBBF4C0" country="Ecuador" httpUri="http://grbio.org/cool/rm44-by7m" name="Museo Ecuadoriano de Ciencias Naturales" type="">MECN</collectionCode>
|
||||
</named-content>
|
||||
) during the period 1994–2001. The species epithet is formed from the name “ Igor ” taken as a noun in apposition.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EOLCI" type="diagnosis">
|
||||
<paragraph id="40FE2E7D1DB3522610CEC8FA941D4E09">
|
||||
<heading id="285CC66D3F539EC33CF301368CC50350" reason="title">Diagnosis.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="96FE5843656C323F146D5E6C8DED97B1">
|
||||
Species of
|
||||
<taxonomicName id="2C203AF5C3FE23E0B2E9626274A84BDB" authorityName="Coues" authorityYear="1884" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="genus">
|
||||
<emphasis id="8C8251B75263128E6FE482C9CDD8D3DD" italics="true">Thomasomys</emphasis>
|
||||
</taxonomicName>
|
||||
with a unique combination of characters, as follows: Head-body length
|
||||
<quantity id="4C01BFAA640E2E05C39E77F87F65B2F6" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" metricValueMax="1.22" metricValueMin="1.02" unit="mm" value="112.0" valueMax="122.0" valueMin="102.0">102–122 mm</quantity>
|
||||
, with long tail (~ 130 % of head-body length); interorbital region narrow with rounded supraorbital margins; zygomatic arches converging anteriorly; long incisive foramina covering approximately ~ 72 % of the diastema, but not extending posteriorly between the molar series; M 1 without reaching the posterior edge of the zygomatic plate; subsquamosal fenestra larger than the postglenoid foramen; upper incisors opisthodont; M 3 with paraflexus and mesoflexus long and fused; M 3 comparatively large; procingulum of m 1 divided.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E2LCI" type="description">
|
||||
<paragraph id="6F19D85E1BEC5AFFDACA282B2A9D7472">
|
||||
<heading id="90D64F3B610B97F06DA97A5591245117" reason="title">
|
||||
Morphological description of the
|
||||
<typeStatus id="39A8C359F974529CA59DB3C40D3014B9" type="holotype">holotype</typeStatus>
|
||||
and variation.
|
||||
</heading>
|
||||
</paragraph>
|
||||
<paragraph id="4C75F13CFB929F5BEE768626849D6AF5">
|
||||
Fine, dense, and soft coat, about
|
||||
<quantity id="94CFED8239593FB61FAB421FDC75B4EA" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" unit="mm" value="12.0" valueMax="13.0" valueMin="11.0">11–13 mm</quantity>
|
||||
long on the back and rump. Dorsal coloration Prouts Brown (Color 47), along the flanks changing to Drab (color 19). Ventral coat Medium Neutral Gray (Color 298) basally, with superficial Raw Umber (Color 24); not clearly separated from the dorsal coloration (Fig.
|
||||
<figureCitation id="27EF1F329CB79C3952CD901DA88480B4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dry skin in dorsal (A, B), lateral (C – E), and ventral (E, F) views of Thomasomys otavalo, sp. nov. (A, C, E; MECN 6912, holotype), and T. igor sp. nov. (B, D, F; MECN 702, holotype). Scale bar: 20 mm." figureDoi="10.3897/vz.74.e128528.figure4" httpUri="https://binary.pensoft.net/fig/1168227">4 B, D, F</figureCitation>
|
||||
). Mystacial vibrissae long, extending beyond the pinnae when placed backward along the head. Ears covered with short, blackish hairs not contrasting with the color of the head. Hairs on metapodials and fingers and toes white and dark giving a gray-haired appearance; ungual tufts white, abundant and extending beyond the edge of the claws (Fig.
|
||||
<figureCitation id="519CEFEF027D415E38CC58B1754137F7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Selected external and soft anatomical traits of Thomasomys: A, C dorsal and (B, D) plantar surface of right pes (A, B: T. otavalo sp. nov, MECN 5604, paratype; C, D T. igor sp. nov, MECN 708, paratype); E soft palate (T. otavalo sp. nov, MECN 6921, paratype); F, G gross morphology of the stomach (T. otavalo sp. nov, MECN 5700, paratype). Abbreviations: 1–5, digits; b, bordering fold; co, cornified epithelium; d, duodenum; d 1 – d 3, diastemal rugae; ge, glandular epithelium; i, incisura angularis; i 1–15, interdental rugae. Scale bars: 5 mm." figureDoi="10.3897/vz.74.e128528.figure5" httpUri="https://binary.pensoft.net/fig/1168228">5 C, D</figureCitation>
|
||||
). Tail longer than the combined length of head and body (about 130 % of
|
||||
<abbrev id="ABBRID0EJMCI" xlink_title="head and body length">HB</abbrev>
|
||||
), uniformly dark, but with white tips (<
|
||||
<quantity id="145D92194D6BAB7E4F80FDAE53ACE2D6" metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
|
||||
, Fig.
|
||||
<figureCitation id="8C282354B7EFB3E9C3D4D11DB12197DA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distal portion of the tail of some species of Thomasomys: A T. otavalo sp. nov. (MECN 5604, paratype); B T. ucucha (MECN 2691); C T. igor sp. nov. (MECN 708, paratype); D T. silvestris (MECN 5054). Scale bar: 10 mm." figureDoi="10.3897/vz.74.e128528.figure6" httpUri="https://binary.pensoft.net/fig/1168229">6 C</figureCitation>
|
||||
); with short sparse hairs, giving a naked appearance at least up to the proximal half, while the distal half is somewhat more hairy; tail end with some larger hairs (<
|
||||
<quantity id="4E1ACD142FF38D4F3E5A7A62711DC81B" metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
|
||||
); tail covered with rectangular scales (14 or 15 rows / cm near base), with three dark brown hairs emerging from base of each scale, no longer than 1.5–2 scale rows, ventral hairs with whitish tips. Mammae six in inguinal, abdominal, and postaxial pairs.
|
||||
</paragraph>
|
||||
<paragraph id="4EDE3B43D23CBFCB0FA629F96474E4F9">
|
||||
Skull medium for the genus (
|
||||
<quantity id="44324FAA260A274F1DE1F97EC48BD838" metricMagnitude="-2" metricUnit="m" metricValue="2.665" metricValueMax="2.7399999999999998" metricValueMin="2.59" unit="mm" value="26.65" valueMax="27.4" valueMin="25.9">25.9–27.4 mm</quantity>
|
||||
<abbrev id="ABBRID0ETMCI" xlink_title="Condylo-incisive length">
|
||||
<collectionCode id="EBBE3E982E5FB8D9AD2D099819C97214">CIL</collectionCode>
|
||||
</abbrev>
|
||||
). Rostrum long and narrow (Fig.
|
||||
<figureCitation id="5BBC31DE1F89DFDF2E91434C734467A1" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Thomasomys igor sp. nov. (Río Tatahuazo, Bolívar, Ecuador): cranium in (A) dorsal, (B) ventral, and (C) lateral views, and mandible in (D) labial view (MECN 702, holotype). Scale bar: 1 mm." figureDoi="10.3897/vz.74.e128528.figure13" httpUri="https://binary.pensoft.net/fig/1168236">13 A</figureCitation>
|
||||
), with nasal and premaxillary bones extending beyond anterior face of incisors; gnathic process poorly developed. Posterior margin of nasal bone reaches and exceeds the plane of the lacrimal bone. Shallow zygomatic notches. Enlarged lacrimal bones, triangular in appearance. Narrow interorbital region with smooth external borders, leaving the maxillary alveolar processes and labial part of the molars exposed in dorsal view. Zygomatic arches narrow anteriorly. Supraorbital region with divergent posterior borders (sensu
|
||||
<bibRefCitation id="E4EEA5009364CA57A0C950C0D49584A5" author="Steppan" firstAuthor="Steppan" journalOrPublisher="Fieldiana Zoology" pagination="1-112" refId="B75" refString="Steppan S (1995) Revision of the tribe Phyllotini (Rodentia: Sigmondontinae), with a phylogenetic hypothesis for the Sigmodontinae. Revisión de la tribu Phyllotini (Rodentia: Sigmondontinae), con una hipótesis filogenética para los Sigmodontinae. Fieldiana Zoology 1464: 1–112." title="Revision of the tribe Phyllotini (Rodentia: Sigmondontinae), with a phylogenetic hypothesis for the Sigmodontinae. Revisión de la tribu Phyllotini (Rodentia: Sigmondontinae), con una hipótesis filogenética para los Sigmodontinae." volume="1464" year="1995">Steppan 1995</bibRefCitation>
|
||||
). Frontoparietal suture “
|
||||
<collectionCode id="223DC6430AB6B85E20DCBCB863D867CE" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
|
||||
” shaped. Broad and rounded braincase, slightly flattened at the outer edges. Broad and concave exoccipital. Dorsal profile (in lateral view) distinctly flattened from nasal tips to middle frontal region; anterior margin of zygomatic plate slightly sloping backward. No further development of the ethmoturbinals. Narrow infraorbital foramen. Not distinguishable lambdoidal crest. M 1 without reaching the posterior edge of the zygomatic plate (Fig.
|
||||
<figureCitation id="74BAABECEC6A6866BB13989D169AE6AE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Morphological comparisons. Comparison of diastemal and palate region in four species of Thomasomys. A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928). Arrow in (A) indicates that the anterior border of the M 1 at the same level of the posterior end of the zygomatic plate; the lower dotted line indicates the penetration of the mesopterygoid fossa in relation to the hypoflexus of the M 3; abbreviations: hf, hypoflexid; if, incisive foramina; mpf, mesopterygoid fossa; pal, palate; ppp, posterolateral palatal pit; zp, zygomatic plate. Scale bar: 5 mm." figureDoi="10.3897/vz.74.e128528.figure8" httpUri="https://binary.pensoft.net/fig/1168231">8 C</figureCitation>
|
||||
). Slender zygomatic arches, with long jugals spanning a similar segment in the zygomatic process of the maxillary and squamosal bone. Alisphenoid strut wide and short. Carotid circulation primitive (Pattern 1, sensu
|
||||
<bibRefCitation id="D606BB2EBBF16422BFAEFCCB6CC42A25" author="Voss" firstAuthor="Voss" journalOrPublisher="Bulletin of the American Museum of Natural History" pagination="259-493" refId="B85" refString="Voss RS (1988) Systematics and ecology of ichthyomyine rodents (Muroidea): Patterns of morphological evolution in a small adaptive radiation. Bulletin of the American Museum of Natural History 188: 259–493." title="Systematics and ecology of ichthyomyine rodents (Muroidea): Patterns of morphological evolution in a small adaptive radiation." volume="188" year="1988">Voss 1988</bibRefCitation>
|
||||
), as indicated by large stapedial foramen, prominent squamosal-alisphenoid groove, and sphenofrontal foramen. Postglenoid foramen approximately twice as large as the subsquamosal fenestra; hamular process of squamosal thin, long, slightly curved and applied distally over the mastoid auditory capsule; tegmen tympani broadly overlapping the posterior suspensory process of squamosal. Upper edge of the mastoid does not exceed the edge of the subsquamosal fenestra (Fig.
|
||||
<figureCitation id="1CC9D4ADBD95BD8026ED1A155D912011" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Alisphenoid-mastoid region comparisons in lateral view in four species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928). Arrow in (B) illustrated to upper edge of the mastoid that exceed the edge of the subsquamosal fenestra; while the oblique dotted line indicates the suture between the squamosal and parietal. Abbreviations: hp, hamular process of squamosal; exo, exoccipital; ip, interparietal; pa, parietal; lpp, lateral parietal portion; mas, mastoid; mf, mastoid fenestra; pgf, postglenoid foramen; sq, squamosal; ssf, subsquamosal fenestra; zps, zygomatic process of the squamosal bone. For ease of comparison are not included to scale" figureDoi="10.3897/vz.74.e128528.figure9" httpUri="https://binary.pensoft.net/fig/1168232">9 C</figureCitation>
|
||||
). Bullae small; pars flaccida of tympanic membrane present, large; orbicular apophysis of malleus well developed. Paraoccipital process small. Hill foramen very small; long and narrow incisive foramina (averaging 72 % of diastemal length), not approaching first molar alveoli. Narrow premaxillary process, maxillary septum of the incisive foramen very thin and long. Short and broad palate (sensu
|
||||
<bibRefCitation id="1E8B843B51938E36626C7B0BA4AD4215" author="Hershkovitz" firstAuthor="Hershkovitz" journalOrPublisher="Fieldiana Zoology" pagination="1-524" refId="B30" refString="Hershkovitz P (1962) Evolution of Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group. Fieldiana Zoology 46: 1–524." title="Evolution of Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group." volume="46" year="1962">Hershkovitz 1962</bibRefCitation>
|
||||
), with the mesopterygoid fossa entering between the molars and reaching the protocone of M 3. Posterolateral palatal pit small and inconspicuous. Wide, slightly divergent-sided mesopterygoid fossa, extending anteriorly between third molars; bony roof perforated by wide, slit-like sphenopalatine openings flanking the basisphenoid. Basisphenoid wide with slightly flat edges. Small foramen ovale. Parapterygoid fossae narrow, approximately triangular, with shallow (unexcavated) anterior limits. Middle lacerate foramen narrow. Lateral expressions of the parietals present, small (Fig.
|
||||
<figureCitation id="1D194DCA77BBB422A1D438A37ECEDF78" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Alisphenoid-mastoid region comparisons in lateral view in four species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928). Arrow in (B) illustrated to upper edge of the mastoid that exceed the edge of the subsquamosal fenestra; while the oblique dotted line indicates the suture between the squamosal and parietal. Abbreviations: hp, hamular process of squamosal; exo, exoccipital; ip, interparietal; pa, parietal; lpp, lateral parietal portion; mas, mastoid; mf, mastoid fenestra; pgf, postglenoid foramen; sq, squamosal; ssf, subsquamosal fenestra; zps, zygomatic process of the squamosal bone. For ease of comparison are not included to scale" figureDoi="10.3897/vz.74.e128528.figure9" httpUri="https://binary.pensoft.net/fig/1168232">9 C</figureCitation>
|
||||
). Auditory bullae small and uninflated with short and wide Eustachian tubes. Dentary moderately long, with long and narrow coronoid process (extending beyond upper edge of condylar process); postcondylar and mental processes poorly developed; deep sigmoid notch. Semilunar recess symmetrical, with lower edge ends pointed. Capsular projection of the root of the incisor inconspicuous.
|
||||
</paragraph>
|
||||
<caption id="485811B0087515A057D3F1434889AB01" ID-DOI="10.3897/vz.74.e128528.figure13" ID-arpha="AEA40C65-97E2-5C04-8867-917C56080B13" httpUri="https://binary.pensoft.net/fig/1168236" startId="F13">
|
||||
<paragraph id="DA2DD5E8EE065C10AA204B4068EE9586">
|
||||
<label id="F36450DE4C067841898C47C9D5687D5C">Figure 13.</label>
|
||||
</paragraph>
|
||||
<paragraph id="8E8446A8D818235FBE3D62B244563CDF">
|
||||
<taxonomicName id="A0B9C8A81A2EFE59D15DBB9C1B4EA6C2" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="78AA7228663B6D7D1DB3813ECD1C4DE8" italics="true">Thomasomys igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="8FFC77442DC5E61D26581DEAB20FE4E3" bold="true">
|
||||
<taxonomicNameLabel id="5485A3EC6C656D5D27B0A3444B097788" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
(Río Tatahuazo, Bolívar, Ecuador): cranium in (
|
||||
<emphasis id="5EA0E4FFEE99ED9D67E533354CAB3EC8" bold="true">A</emphasis>
|
||||
) dorsal, (
|
||||
<emphasis id="08F46A93E35928BDAEC5AB1E1BEC8D9D" bold="true">B</emphasis>
|
||||
) ventral, and (
|
||||
<emphasis id="019DFD3FAAF256DFB7F74CCC67142BF5" bold="true">C</emphasis>
|
||||
) lateral views, and mandible in (
|
||||
<emphasis id="EC41CD6F0153B6249369E41B20257420" bold="true">D</emphasis>
|
||||
) labial view (
|
||||
<named-content id="BD105BB3E517FBBA799CDF883773358A" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-ecuadoriano-de-ciencias-naturales" xlink_title="Museo Ecuatoriano de Ciencias Naturales">MECN</named-content>
|
||||
702, holotype). Scale bar: 1 mm.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="BF1DDB1D853F67A9966CABE914B3FA89">
|
||||
Upper incisors small, slender, and opisthodont (Thomas’s angle of 80 °,
|
||||
<bibRefCitation id="76EFEE472DF7BCD19F2914E1DB8DF8F7" DOI="10.1080/00222931908673892" author="Thomas" firstAuthor="Thomas" journalOrPublisher="Annals and Magazine of Natural History" pagination="289-290" refId="B79" refString="Thomas O (1919) The method of taking the incisive index in rodents. Annals and Magazine of Natural History 4: 289–290. https://doi.org/10.1080/00222931908673892" title="The method of taking the incisive index in rodents." volume="4" year="1919">Thomas 1919</bibRefCitation>
|
||||
; see Fig.
|
||||
<figureCitation id="ED70D74D6CC1B2FF3915A3EE33E420D1" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Comparison of the left anterior portion of the rostrum, viewed from left side, in four species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928). Thomas’ angles according to incisive and molar planes are indicated as well as the extension of the molar series. Arrow in (A) point to the anterior border of the nasal bone extending beyond the anterior face of the incisors relative to (B); abbreviations: n, nasal; nc, nasolacrimal capsule; p, premaxillary; sf, supraorbital foramen; zp, zygomatic plate. Scale bar: 5 mm" figureDoi="10.3897/vz.74.e128528.figure10" httpUri="https://binary.pensoft.net/fig/1168233">10 C</figureCitation>
|
||||
), with front enamel Light Chrome Orange (color 76); brachydont and pentalophodont molars (sensu
|
||||
<bibRefCitation id="BE0340F4CAA5FCD2890B24B1EAB19697" author="Hershkovitz" firstAuthor="Hershkovitz" journalOrPublisher="Fieldiana Zoology" pagination="1-524" refId="B30" refString="Hershkovitz P (1962) Evolution of Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group. Fieldiana Zoology 46: 1–524." title="Evolution of Neotropical cricetine rodents (Muridae) with special reference to the phyllotine group." volume="46" year="1962">Hershkovitz 1962</bibRefCitation>
|
||||
). Upper molars in left and right parallel series, small and pentalodont; hypsodont and cingulate in juveniles; coronal surfaces crested when unworn; main cusps slightly opposite and inclined backward when viewed from the side. M 1 rectangular in outline with procingulum divided by the anteromedian flexus into subequal anterolabial and anterolingual conules; anteroloph large; mesoloph short and / or segmented. M 2 squared in outline; mesoloph showing the same condition as in M 1. M 3 enlarged relative to M 2, rounded in outline with conspicuous anteroloph; paraflexus and metaflexus long and fused (Fig.
|
||||
<figureCitation id="DDBF795C75FC7C6EB4A81A74880B5595" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Comparison of the upper right molar series in occlusal view among some species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928); abbreviations: af, anteromedian flexus; al, anteroloph; ml, mesoloph; mfl, metaflexus; pf, paraflexus; prf, protoflexus. Scale bar: 1 mm." figureDoi="10.3897/vz.74.e128528.figure11" httpUri="https://binary.pensoft.net/fig/1168234">11 C</figureCitation>
|
||||
); mesoloph present in juvenile specimens; hypoflexid conspicuous. Lower molars with alternate, posteriorly inclined main cusps viewed from the side. First lower molar (m 1) with anteromedian flexid dividing procingulum into subequal anterolabial and anterolingual conules (Fig.
|
||||
<figureCitation id="72C3B5856187BD808B701C57B9D64F34" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Comparison of the lower right molar series in occlusal view among some species of Thomasomys: A T. otavalo sp. nov. (MECN 6912, holotype); B T. ucucha (MECN 7937); C T. igor sp. nov. (MECN 702, holotype); D T. silvestris (MECN 4928); abbreviations: ml, mesolophid; prf, protoflexid. Scale bar: 1 mm." figureDoi="10.3897/vz.74.e128528.figure12" httpUri="https://binary.pensoft.net/fig/1168235">12 C</figureCitation>
|
||||
); mesolophid short; mesolophid of m 2 short, hypoflexid deep; m 3 slightly shorter than m 2.
|
||||
</paragraph>
|
||||
<paragraph id="F20AF013F152344A98B88A3787E34651">Tuberculum of first rib articulates with transverse processes of seventh cervical and first thoracic vertebrae; second thoracic vertebra with differentially elongated neural spine; vertebral column composed of 19 thoracolumbar, 4 sacral (fused), and 30–39 caudal vertebrae; usually the second and third caudal vertebrae with small but complete hemal arches; 12 ribs. Details of soft anatomy and genitalia unknown.</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0EUPCI" type="comparisons">
|
||||
<paragraph id="37EF3E4A98D468021FCF8EA8B2FEADF5">
|
||||
<heading id="B6D9DE93F9A0F2A87835BD379E8BF0DD" reason="title">Comparisons.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="79CC77EA6F00B2D08B9E35B21377AB6F">
|
||||
<taxonomicName id="AFB4E4FF7EC0597788FAF9B142296F2B" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="DC45F72608046CA5085B10953EADB07D" italics="true">Thomasomys igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="4EC195BB8BB05B651961F291353DA5C1" bold="true">
|
||||
<taxonomicNameLabel id="2A6D0F8E47B5FC7A9E94CF4C98664EEC" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is retrieved as the sister species to
|
||||
<taxonomicName id="E15D2C819057287DFD384B3725AEBAB5" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
|
||||
<emphasis id="38622B20C5039366A1A9FA4EFFD99762" italics="true">T. silvestris</emphasis>
|
||||
</taxonomicName>
|
||||
(Fig.
|
||||
<figureCitation id="A5B2BC7F8486D1AC3FF31C1AB583815E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Cyt b gene tree of the genus Thomasomys and close outgroups. The tree was inferred by maximum likelihood (ML) using a nucleotide substitution model (GTR + F + G 4); it contains 222 terminals belonging to 36 named species of Thomasomys, including the two new species here described (indicated with black arrows), it also contains at least 20 putative new species of Thomasomys, and 20 outgroups of the genera Rhipidomys, Chilomys and Rhagomys. Nodal support statistics for the analyses in IQ-Tree are from: ultrafast bootstrap approximation (UF), SH-like approximate likelihood ratio test (aL), and the aBayes test (aB). White sections of the circles indicate UF and aL bootstrap frequencies <75 % and aB posterior probabilities <0.95, whereas black sections of the circles indicate UF and aL bootstrap frequencies ~ 75 % and aB posterior probabilities ~ 0.95. Insets represent differences in topology and branch lengths of three ML approaches. Inset A: Collapsed gene tree of the main figure. Inset B: Collapsed gene tree inferred with a codon substitution model. Inset C: Collapsed gene tree estimated with the RAxML tree with the CAT approximation; black circles represent bootstrap values ~ 75 %. Terminals are labeled with GenBank accession number, museum voucher number and scientific name — in case that the species have not been described, the sequences are indicated with ‘ sp’ and a geographical proxy — ‘ ONT ’ indicates that the sequence was obtained with MinION Oxford Nanopore Technologies." figureDoi="10.3897/vz.74.e128528.figure1" httpUri="https://binary.pensoft.net/fig/1168224">1</figureCitation>
|
||||
: Part A). To construct this section, we used specimens referred to
|
||||
<taxonomicName id="D9DCF1179B6B5AEA14CF92C1D3B1D8BB" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
|
||||
<emphasis id="AA32319E87D6F8991EB230EE2804CE91" italics="true">Thomasomys silvestris</emphasis>
|
||||
</taxonomicName>
|
||||
(sensu stricto) from western
|
||||
<collectingRegion id="AEEDC7886FC415A479E4B0A2D803C15C" country="Ecuador" name="Cotopaxi">Provincia de Cotopaxi</collectingRegion>
|
||||
and
|
||||
<collectingRegion id="E4DC2C10289AB4715182FFA278FBE63D" country="Ecuador" name="Pichincha">Pichincha</collectingRegion>
|
||||
(
|
||||
<bibRefCitation id="ECD7966890B844509D642A17D72CCF7C" author="Anthony HE" refId="B5" refString="Anthony HE (1924 a) Preliminary report on Ecuadorean mammals. No. 4. American Museum Novitates 114: 6.">Anthony 1924 a</bibRefCitation>
|
||||
,
|
||||
<bibRefCitation id="66612B730E2F30B2769897D37FE63DE5" author="Anthony HE" refId="B6" refString="Anthony HE (1924 b) Preliminary report on Ecuadorean mammals. No. 6. American Museum Novitates 139: 9.">1924 b</bibRefCitation>
|
||||
;
|
||||
<bibRefCitation id="132B19C5ED4C10389CDD3090743A7B8E" DOI="10.31687/saremMN.19.26.2.0.04" author="Brito" etAl="et al." firstAuthor="Brito" journalOrPublisher="Mastozoología Neotropical" pagination="308-330" refId="B15" refString="Brito J, Tinoco N, Curay J, Vargas R, Reyes-Puig C, Romero V, Pardiñas UF (2019) Diversidad insospechada en los Andes de Ecuador: filogenia del grupo “ cinereus ” de Thomasomys y descripción de una nueva especie (Rodentia, Cricetidae). Mastozoología Neotropical 26: 308–330. https://doi.org/10.31687/saremMN.19.26.2.0.04" title="Diversidad insospechada en los Andes de Ecuador: filogenia del grupo “ cinereus ” de Thomasomys y descripción de una nueva especie (Rodentia, Cricetidae)." volume="26" year="2019">Brito et al. 2019</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="11C9914B408C1233E0EB5932F1C698F6" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="05B91FBA321034203D32A6C5309C0E75" italics="true">Thomasomys igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="D93141E8B0D19A8CDABD88087B97F0C0" bold="true">
|
||||
<taxonomicNameLabel id="B20B1E153BF2C6E96E8F64EE22ED46D5" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is a medium-sized species (Table
|
||||
<tableCitation id="EB40D5F2E41274D0988653950EC50E7F" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Summary of morphometric measurements of all specimens in mm. Species names are accompanied by the number of analyzed individuals between parentheses. Minimum and maximum values are accompanied by mean and standard deviation between parentheses. Abbreviations of characters are detailed in the text." httpUri="http://table.plazi.org/id/B5A75659D53B773ADE5A518BE89C41B5" tableUuid="B5A75659D53B773ADE5A518BE89C41B5">4</tableCitation>
|
||||
), which differs from
|
||||
<taxonomicName id="BC674F3F2E78B937A56E27733EBDC737" authorityName="Anthony" authorityYear="1924" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="silvestris">
|
||||
<emphasis id="D0AF51E8D71A35D01F919B3761632781" italics="true">T. silvestris</emphasis>
|
||||
</taxonomicName>
|
||||
(characteristics in parentheses) by having slightly shorter tail ~ 130 % of
|
||||
<abbrev id="ABBRID0EJSCI" xlink_title="head and body length">HB</abbrev>
|
||||
(~ 140 %). Craniodentally, qualitative differences between the two species are conspicuous: M 1 without reaching the posterior edge of the zygomatic plate (beyond); M 3 with paraflexus and metaflexus long and fused (short); procingulum of m 1 divided (undivided or indistinct).
|
||||
</paragraph>
|
||||
<paragraph id="A1A74D71179D3881089E54D1F4372E6B">
|
||||
Other
|
||||
<taxonomicName id="00F401792F63C9FC95ECFA62C76D8F9E">
|
||||
<emphasis id="F01B2372601F40DEA5FC801C47A50A7A" italics="true">Thomasomys</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
who could be confused with
|
||||
<taxonomicName id="7E883C9ECB8784CB5885C82AC715F40F" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="8DF056B664499AD73362E10CA86DDE88" italics="true">T. igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="4716E63F48FB3AEAF33774963983CA5D" bold="true">
|
||||
<taxonomicNameLabel id="7BE031303DBE13655993C0C5072F56E9" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
are
|
||||
<taxonomicName id="41FF08862FDA70886C3CF8664CC4A604" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo">
|
||||
<emphasis id="20ED500DF2E4B6B9FFF86E8A096276F7" italics="true">T. otavalo</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="2867D880D877B420D0C556DC35C15E71" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
|
||||
<emphasis id="4FB22458C9BCE86DF5C167B1EC9D7224" italics="true">T. ucucha</emphasis>
|
||||
</taxonomicName>
|
||||
. However,
|
||||
<taxonomicName id="B9B3840BE3F7B947354630B298AF2DF4" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="4BB1F953305CA9CD0CB49F4AE52AA19B" italics="true">T. igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="706308F526447D6F31189A0CA4D6A2F4" bold="true">
|
||||
<taxonomicNameLabel id="EF446C7AB27522A418A8AE113822A0D0" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
can be differentiated from these species by the white tail tip <
|
||||
<quantity id="94129DDC95408D13B829759357170997" metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
|
||||
(
|
||||
<quantity id="D7F47A51DA86295A0136197992C47E38" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" unit="mm" value="25.0" valueMax="35.0" valueMin="15.0">15–35 mm</quantity>
|
||||
in
|
||||
<taxonomicName id="205FC704CA1437A06D0852C6463E98A9" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo">
|
||||
<emphasis id="1ABCAE8B4777B25A45D7DC0E0D494258" italics="true">T. otavalo</emphasis>
|
||||
</taxonomicName>
|
||||
and <5
|
||||
<taxonomicName id="F696EB45E6CB893D9C2A24AF69286404" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
|
||||
<emphasis id="932CFD05884ED428504807947659F96D" italics="true">T. ucucha</emphasis>
|
||||
</taxonomicName>
|
||||
). Regarding the skull,
|
||||
<taxonomicName id="BEA1583DE09E299EFD1EB7A00048EA1E" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="DAFF7A6651110D2ED60BB79656323E9D" italics="true">T. igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="7BFF24C0C4B24BED9B11D838CF10D05F" bold="true">
|
||||
<taxonomicNameLabel id="C674EE8FA1CCB486946BB2DD55E8146B" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
can be distinguished by the incisive foramina (72 % of
|
||||
<abbrev id="ABBRID0EJVCI" xlink_title="length of upper diastema">LD</abbrev>
|
||||
), while
|
||||
<taxonomicName id="C27B163C7B80320917BF3D5635D3B93D" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo">
|
||||
<emphasis id="A563E7446DA84E67FAEC26E8BFEE604A" italics="true">T. otavalo</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="93FB3BC7C94EFF29A7FEA01F473CA479" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
|
||||
<emphasis id="1E96B8B313D79DF9B9BE066F3D5E6642" italics="true">T. ucucha</emphasis>
|
||||
</taxonomicName>
|
||||
are shorter (63 and 57 % of
|
||||
<abbrev id="ABBRID0EDWCI" xlink_title="length of upper diastema">LD</abbrev>
|
||||
, respectively). As for the molars, in
|
||||
<taxonomicName id="2E2A51D5E796F61F2A1F9BE50FA0D875" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="FBAF5D875A6FF7A753A29EEE50286C54" italics="true">T. igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="5075CA133C182228DDC3C9DAFC2F225F" bold="true">
|
||||
<taxonomicNameLabel id="6475ECD421B946AABC036AFDD061DB76" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
in M 3 the paraflexus and metaflexus long and fused, while in
|
||||
<taxonomicName id="4FA8BA995D25614EA97D8EC65A855C0E" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="otavalo">
|
||||
<emphasis id="CBA976FC8C6D4591F53F0BF9DA0B2ED6" italics="true">T. otavalo</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="CA254C316B7B85EF772BD3CF1F4B4027" baseAuthorityName="MECN" baseAuthorityYear="2691" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="ucucha">
|
||||
<emphasis id="2270F3A62CF60B17E5E2DD71F737102C" italics="true">T. ucucha</emphasis>
|
||||
</taxonomicName>
|
||||
they are short.
|
||||
<taxonomicName id="B5BBD2F80B7C32B57A3164F682D195E6" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="0F74791DD99028CA9FE054AE462C52FD" italics="true">Thomasomys igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="66C71F51219B8EA532D0E77EE6F9553C" bold="true">
|
||||
<taxonomicNameLabel id="5927CD6434FAA4D14D8C2C7A6CAB08F8" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
was previously referred to as
|
||||
<taxonomicName id="D9091BF7E690EA327C4932B7323E569E" authorityName="Anthony" authorityYear="1923" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="caudivarius">
|
||||
<emphasis id="8026143472F55D6BB8DEB14551AA8D93" italics="true">T. caudivarius</emphasis>
|
||||
</taxonomicName>
|
||||
by Salazar and Yates (2007),
|
||||
<bibRefCitation id="06DA1CDA519E0E7E65980A5A1F4BECC2" author="Pacheco V" editor="Patton JL & Pardiñas UFJ & D’Elia G" firstAuthor="Pacheco" journalOrPublisher="University of Chicago Press, Chicago, IL" pagination="617-682" refId="B50" refString="Pacheco V (2015) Genus Thomasomys Coues, 1884. In: Patton JL, Pardiñas UFJ, D’Elia G (Eds) Mammals of South America, Volume 2: Rodents. University of Chicago Press, Chicago, IL, 617–682." title="Genus Thomasomys Coues, 1884." volumeTitle="Mammals of South America, Volume 2: Rodents." year="2015">Pacheco (2015)</bibRefCitation>
|
||||
, and Steppan and Schenck (2017), however,
|
||||
<taxonomicName id="21FA482E50368FDEFB5304677ADE748D" authorityName="Anthony" authorityYear="1923" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="caudivarius">
|
||||
<emphasis id="65779E57C55F594ABD8B5017A3F7EFC6" italics="true">T. caudivarius</emphasis>
|
||||
</taxonomicName>
|
||||
is larger in size (see
|
||||
<bibRefCitation id="B2EA1C54449DF7B5EC6B6BE35B2257F5" DOI="10.31687/saremMN.19.26.2.0.04" author="Brito" etAl="et al." firstAuthor="Brito" journalOrPublisher="Mastozoología Neotropical" pagination="308-330" refId="B15" refString="Brito J, Tinoco N, Curay J, Vargas R, Reyes-Puig C, Romero V, Pardiñas UF (2019) Diversidad insospechada en los Andes de Ecuador: filogenia del grupo “ cinereus ” de Thomasomys y descripción de una nueva especie (Rodentia, Cricetidae). Mastozoología Neotropical 26: 308–330. https://doi.org/10.31687/saremMN.19.26.2.0.04" title="Diversidad insospechada en los Andes de Ecuador: filogenia del grupo “ cinereus ” de Thomasomys y descripción de una nueva especie (Rodentia, Cricetidae)." volume="26" year="2019">Brito et al. 2019</bibRefCitation>
|
||||
: table 2) and has a disjunct distribution on the southwestern slope of the Andes. Another species with which
|
||||
<taxonomicName id="AF2FC924BD6ECE90E96A9CABD35F6BFC" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="D672F8BFD76C15CA7C1C155117845CBC" italics="true">Thomasomys igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="A5753DA0EAF2B532B11F990C2625981A" bold="true">
|
||||
<taxonomicNameLabel id="3A91EC6CB93FFC37733DFB0C9EEB5F5A" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
could be confused is
|
||||
<taxonomicName id="013DAB7B216B2B6BB349BA1B2C659447" baseAuthorityName="Tomes" baseAuthorityYear="1860" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="aureus">
|
||||
<emphasis id="A9D48CBFBB83F0A7D9FEA89FEE311181" italics="true">T. aureus</emphasis>
|
||||
</taxonomicName>
|
||||
(sympatric species), however they are easily distinguished because the new species is notoriously smaller ~ 113 % of
|
||||
<abbrev id="ABBRID0ENZCI" xlink_title="head and body length">HB</abbrev>
|
||||
(~ 336).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0ERZCI" type="distribution">
|
||||
<paragraph id="B235523FF21214EEFBE413178B04F345">
|
||||
<heading id="531781DCF03344AAE9128CB9964CC255" reason="title">Distribution.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="DF33D0BFFD00208824483F7135A59716">
|
||||
<taxonomicName id="80D2B7AF8130F08289E0F2A45DB7C6A0" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="6142CF86DE7534EC111A8C804D7B1164" italics="true">Thomasomys igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="BDBEE4ECE9EB7C4F61BB42D96A178091" bold="true">
|
||||
<taxonomicNameLabel id="B0B3F7F88C0F8961D06B879AF39DF07E" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is known only from one locality, near to Cruz de Lizo,
|
||||
<collectingRegion id="C2D7D3D36B7A0D68C7DA9D61DA070E74" country="Ecuador" name="Bolivar">Provincia de Bolívar</collectingRegion>
|
||||
, in the intersections of the Bosque Protector Cashca Totoras, at elevations of
|
||||
<quantity id="665FFEBC13788E9B26F8178090CDBD68" metricMagnitude="3" metricUnit="m" metricValue="2.9375" metricValueMax="3.0" metricValueMin="2.875" unit="m" value="2937.5" valueMax="3000.0" valueMin="2875.0">
|
||||
<geoCoordinate id="1560CAE484D07D1BE68941C223CBD8FA" degrees="2,875" direction="north" orientation="latitude" precision="55" value="2.875">2,875</geoCoordinate>
|
||||
<geoCoordinate id="A8E2CA5B0C199577E60A4820CE0C852D" degrees="3,000" direction="west" orientation="longitude" precision="55" value="-3.0">–3,000</geoCoordinate>
|
||||
m
|
||||
</quantity>
|
||||
. The new species is geographically delimited by the basins of the rivers Angamarca (north), Chanchán (south), to the east by the inter-Andean valley and to the west by the tropical rainforest (see Fig.
|
||||
<figureCitation id="1AA36FFBBCC14AD41A7B4EBBC8AB64FE" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A relief map of Ecuador is depicted with hypsometric tints that show the elevation variations across the region. The sampling localities of five Thomasomys species are marked on the map." figureDoi="10.3897/vz.74.e128528.figure14" httpUri="https://binary.pensoft.net/fig/1168237">14</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="1398158674C08C9F410B9CFE6C8357F5" ID-DOI="10.3897/vz.74.e128528.figure14" ID-arpha="9DFA8C98-8922-5CD1-8909-D6A788EC4A91" httpUri="https://binary.pensoft.net/fig/1168237" startId="F14">
|
||||
<paragraph id="384FBF65B82F878039722B7A578C0E44">
|
||||
<label id="0B7C82C78755E2B13E07F98D1E37BE3D">Figure 14.</label>
|
||||
</paragraph>
|
||||
<paragraph id="3586F6926F27F73C463BE0B1C1743909">
|
||||
A relief map of Ecuador is depicted with hypsometric tints that show the elevation variations across the region. The sampling localities of five
|
||||
<taxonomicName id="DEF4C5369C6B58CF0265CF93D46056CD">
|
||||
<emphasis id="69CD375F5D0A4B73936EE8BFEA1C5B4B" italics="true">Thomasomys</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
are marked on the map.
|
||||
</paragraph>
|
||||
</caption>
|
||||
</subSubSection>
|
||||
<subSubSection id="SECID0E31CI" type="natural history">
|
||||
<paragraph id="62F5E1C7096A721298E66A2AA1BB738B">
|
||||
<heading id="8A156F899D5585383CD97B1BD1A808EF" reason="title">Natural history.</heading>
|
||||
</paragraph>
|
||||
<paragraph id="F201E59F292D566B9BD5DD336EFCE886">
|
||||
<taxonomicName id="16943AA1C0814F84B697E87FC78C9250" authorityName="Brito & García & Castellanos & Gavilanes & Curay & Carrión-Olmedo & Reyes-Barriga & Guayasamin & Salazar-Bravo & Pinto" authorityYear="2024" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="igor" status="sp. nov.">
|
||||
<emphasis id="F479D5EEA547CB40C76E44ABAD1FC956" italics="true">Thomasomys igor</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="CAAE9D5091D423C4941AEE95A10A496D" bold="true">
|
||||
<taxonomicNameLabel id="9D2A0593DB9E14FF9F3115FFCC731DFB" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
is found within the temperate and high Andean zoogeographic areas (
|
||||
<bibRefCitation id="2D3F4287221161A4DE5DFA6C08673B26" author="Albuja" etAl="et al." firstAuthor="Albuja" journalOrPublisher="Escuela Politécnica Nacional, Quito" refId="B1" refString="Albuja L, Almendáriz A, Barriga R, Montalvo LD, Cáceres F, Román JL (2012) Fauna de vertebrados del Ecuador. Escuela Politécnica Nacional, Quito, 490 pp. http://bibdigital.epn.edu.ec/handle/15000/6686" title="Fauna de vertebrados del Ecuador." url="http://bibdigital.epn.edu.ec/handle/15000/6686" year="2012">Albuja et al. 2012</bibRefCitation>
|
||||
). The ecosystem corresponds to montane forest (
|
||||
<bibRefCitation id="08897CCE1C13B4FC99948E3E01F12548" journalOrPublisher="Subsecretaría de Patrimonio Natural, Quito" refId="B42" refString="Ministerio del Ambiente del Ecuador (2013) Sistema de clasificación de los ecosistemas del Ecuador continental. Subsecretaría de Patrimonio Natural, Quito, 143 pp." title="Sistema de clasificación de los ecosistemas del Ecuador continental." year="2013">
|
||||
Ministerio del Ambiente del
|
||||
<collectingCountry id="60416ED652F2985571D50DA76845EB1E" name="Ecuador">Ecuador</collectingCountry>
|
||||
2013
|
||||
</bibRefCitation>
|
||||
), which is characterized by trees with abundant orchids, ferns and bromeliads. The species was collected in sympatry with the shrew-opossum,
|
||||
<taxonomicName id="9DC2753034C50AE8F00BD9F691327CD3" baseAuthorityName="Tomes" baseAuthorityYear="1863" class="Mammalia" family="Caenolestidae" genus="Caenolestes" kingdom="Animalia" order="Paucituberculata" phylum="Chordata" rank="species" species="fuliginosus">
|
||||
<emphasis id="3F0F3433328F7B915CF42BE823D1822A" italics="true">Caenolestes fuliginosus</emphasis>
|
||||
</taxonomicName>
|
||||
, the shrew
|
||||
<taxonomicName id="81049CAA23655F6DCA43899DACC551C6" baseAuthorityName="Thomas" baseAuthorityYear="1912" class="Mammalia" family="Soricidae" genus="Cryptotis" kingdom="Animalia" order="Soricomorpha" phylum="Chordata" rank="species" species="equatoris">
|
||||
<emphasis id="28ABE52E5E586DD37FADD1FB035A84A6" italics="true">Cryptotis equatoris</emphasis>
|
||||
</taxonomicName>
|
||||
and the rodents
|
||||
<taxonomicName id="8E33F89CF8694E27A71E7E92D351195B" authorityName="Thomas" authorityYear="1894" class="Mammalia" family="Cricetidae" genus="Akodon" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="mollis">
|
||||
<emphasis id="7E7D5D59E035FE355E10CA505B367171" italics="true">Akodon mollis</emphasis>
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicName id="E4894273D68519D3B92B981FCF92814C" authority="(Tomes, 1860)" baseAuthorityName="Tomes" baseAuthorityYear="1860" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="albigularis">
|
||||
<emphasis id="8007C4F3E0D740D224CA44BDECE8CBFA" italics="true">Nephelomys albigularis</emphasis>
|
||||
(Tomes, 1860)
|
||||
</taxonomicName>
|
||||
, and
|
||||
<taxonomicName id="2FD8506A7F6DFC155B5B4AA848D1BEDB" authority="(Tomes, 1860)" baseAuthorityName="Tomes" baseAuthorityYear="1860" class="Mammalia" family="Cricetidae" genus="Thomasomys" kingdom="Animalia" order="Rodentia" phylum="Chordata" rank="species" species="aureus">
|
||||
<emphasis id="C4CA3C149997CCE3E35D79BDC5F22677" italics="true">Thomasomys aureus</emphasis>
|
||||
(Tomes, 1860)
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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