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<document id="D498D5965B6907DFF2D436406F04D54D" ID-DOI="10.1002/tax.12718" ID-ISSN="1996-8175" ID-Zenodo-Dep="14047464" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects" IM.treatments_approvedBy="carolina" checkinTime="1730917809399" checkinUser="felipe" docAuthor="Frisch, Andreas, Ohmura, Yoshihito, Holien, Håkon &amp; Bendiksby, Mika" docDate="2022" docId="03BEB24E4754FFF182DFD76AFCB8D843" docLanguage="en" docName="TAXON.71.5.936-962.pdf" docOrigin="TAXON 71 (5)" docSource="http://dx.doi.org/10.1002/tax.12718" docStyle="DocumentStyle:239A75409313D6488D9D82529296B06F.1:TAXON.2020-.journal_article.open" docStyleId="239A75409313D6488D9D82529296B06F" docStyleName="TAXON.2020-.journal_article.open" docStyleVersion="1" docTitle="Arthrorhaphis" docType="key" docVersion="1" lastPageNumber="957" masterDocId="FF87CA364747FFE4821BD011FFA7DA11" masterDocTitle="A phylogenetic survey of the ascomycete genus Arthrorhaphis (Arthrorhaphidaceae, Lecanoromycetes) including new species in Arthrorhaphis citrinella sensu lato" masterLastPageNumber="962" masterPageNumber="936" pageNumber="955" updateTime="1732226663877" updateUser="carolina" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:titleInfo id="5EEDF6539C6880EBCF624A75B6AE7E22">
<mods:title id="AC56ED18089B997DC0441B709BFA7C39">A phylogenetic survey of the ascomycete genus Arthrorhaphis (Arthrorhaphidaceae, Lecanoromycetes) including new species in Arthrorhaphis citrinella sensu lato</mods:title>
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<mods:name id="C6C7BBCC35EF2C3F33BE02ABD95745C9" type="personal">
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<mods:namePart id="A9119B9AD179C4A410BEFCEF4F2C3643">Frisch, Andreas</mods:namePart>
<mods:affiliation id="466C749464650FA6E8B04D2DEEE4107B">Yoshihito Ohmura, &amp; &amp; Mika Bendiksby &amp; Department of Natural History, NTNU University Museum, Norwegian University of Science and Technology, 7491 Trondheim, &amp;) A. bullata sp. nov., (&amp;) A. bullata sp. nov., (</mods:affiliation>
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<mods:namePart id="6361DC642BE8DDCDCD9C18182549660D">Ohmura, Yoshihito</mods:namePart>
<mods:affiliation id="259CB9907A7B9BF56A5F0E08C9DF4E3B">Håkon Holien &amp; Department of Botany, National Museum of Nature and Science, Tsukuba, 305 - 0005, Japan &amp;) A. catolechioides comb. &amp; stat. nov., (&amp;) A. catolechioides comb. &amp; stat. nov., (</mods:affiliation>
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<mods:namePart id="E5D4FCDE0B32BE8755FCCE9696DC64A3">Holien, Håkon</mods:namePart>
<mods:affiliation id="30E0A08B5BC6E5B57495C50B7DF5DB3D">Yoshihito Ohmura, &amp; &amp; Mika Bendiksby &amp; Department of Natural History, NTNU University Museum, Norwegian University of Science and Technology, 7491 Trondheim, &amp;) A. bullata sp. nov., (&amp;) A. bullata sp. nov., (&amp; &amp; Mika Bendiksby &amp; Faculty of Bioscience and Aquaculture, Nord University, Box 2501, 7729 Steinkjer, Norway &amp;) A. citrinella, (&amp;) A. citrinella, (</mods:affiliation>
</mods:name>
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<mods:namePart id="1C615D3CA18C8F794B1DB7099D7227AA">Bendiksby, Mika</mods:namePart>
<mods:affiliation id="4EB32412CCA3660337C435280F09254F">Yoshihito Ohmura, &amp; &amp; Mika Bendiksby &amp; Department of Natural History, NTNU University Museum, Norwegian University of Science and Technology, 7491 Trondheim, &amp;) A. bullata sp. nov., (&amp;) A. bullata sp. nov., (&amp; Natural History Museum, University of Oslo, 0319 Oslo, Norway &amp;) A. farinosa sp. nov., and (&amp;) A. farinosa A. vulgaris comb. &amp; stat. nov. A sixth phylogenetic clade from the Neotropics remains undescribed herein due Five circumarctic accessions of A. alpina s. l. form a genetically distinct but morphologically poorly understood alpina-vacillans clade, which we preliminarily name “ A. septentrionalis ”. Jointly, our multispecies coalescence single-locus (bGMYC) and multilocus (bPtP, bP &amp; P) datasets, largely support our proposed species hypotheses Keywords integrative taxonomy; lichenicolous fungi; lichens; molecular phylogenetics; Ostropomycetidae;</mods:affiliation>
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<mods:date id="536B6CED24C6FADFBC466E8C4DC9856B">2022</mods:date>
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<mods:number id="642CF8C97653A1BF1544E55138C40E32">2022-10-31</mods:number>
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<mods:number id="869113498B10C8CB2C4F306B711DE895">71</mods:number>
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<mods:identifier id="D54B4D630ABC70489E65D6ECA8BFEC56" type="ISSN">1996-8175</mods:identifier>
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<treatment id="03BEB24E4754FFF182DFD76AFCB8D843" LSID="urn:lsid:plazi:treatment:03BEB24E4754FFF182DFD76AFCB8D843" httpUri="http://treatment.plazi.org/id/03BEB24E4754FFF182DFD76AFCB8D843" lastPageId="21" lastPageNumber="957" pageId="19" pageNumber="955">
<subSubSection id="C30D50D34754FFF782DFD76AFCB8DDA3" pageId="19" pageNumber="955" type="nomenclature">
<paragraph id="8BA803584754FFF782DFD76AFCB8DDA3" blockId="19.[151,799,1915,1970]" pageId="19" pageNumber="955">
<emphasis id="B963DF4A4754FFF782DFD76AFD12DD83" bold="true" box="[196,693,1915,1939]" pageId="19" pageNumber="955">
Artificial key to the genus
<taxonomicName id="4C1778DB4754FFF783EFD76DFD2CDD83" ID-CoL="33DQ" ID-ENA="146289" baseAuthorityName="Obermayer" baseAuthorityYear="1994" box="[500,651,1916,1938]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="genus">
<emphasis id="B963DF4A4754FFF783EFD76DFD2CDD83" bold="true" box="[500,651,1916,1938]" italics="true" pageId="19" pageNumber="955">Arthrorhaphis</emphasis>
</taxonomicName>
.
<emphasis id="B963DF4A4754FFF78081D76CFD12DD83" bold="true" box="[666,693,1917,1938]" italics="true" pageId="19" pageNumber="955">—</emphasis>
</emphasis>
The morphologically poorly characterised “
<taxonomicName id="4C1778DB4754FFF783F6D78AFCB4DDA3" box="[493,787,1947,1970]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="septentrionalis">
<emphasis id="B963DF4A4754FFF783F6D78AFCB4DDA3" box="[493,787,1947,1970]" italics="true" pageId="19" pageNumber="955">Arthrorhaphis septentrionalis</emphasis>
</taxonomicName>
”
</paragraph>
</subSubSection>
<subSubSection id="C30D50D34754FFF7815ED0AAFAB4DB23" pageId="19" pageNumber="955" type="discussion">
<paragraph id="8BA803584754FFF7815ED0AAFAB4DB23" blockId="19.[837,1485,187,306]" pageId="19" pageNumber="955">
(
<figureCitation id="132C1FDD4754FFF78156D0AAFC3DDAC3" box="[845,922,187,210]" captionStart="Fig" captionStartId="20.[103,135,1874,1895]" captionTargetBox="[221,1318,187,1839]" captionTargetId="figure-0@20.[221,1318,187,1839]" captionTargetPageId="20" captionText="Fig. 8. A, Arthrorhaphis alpina (Schaer., Lichenes Helvetici Exsiccati 532, G, lectotype); B, A. alpina var. jungens (Lichenotheca Graecensis 23, E, isotype); C, A. alpina (Ohmura 10119, TNS); D, A. alpina var. jungens (Kalb, Lichenes Neotropici 577, M); E, A. alpina (Brusse 4515, UPS); F, “A. septentrionalis” (Hansen 026, C). — Scale: A–F, 1 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047480" httpUri="https://zenodo.org/record/14047480/files/figure.png" pageId="19" pageNumber="955">Fig. 8F</figureCitation>
) and
<taxonomicName id="4C1778DB4754FFF781C3D0AAFB00DAC3" box="[984,1191,187,210]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="undefined-1">
<emphasis id="B963DF4A4754FFF781C3D0AAFBC9DAC3" box="[984,1134,187,210]" italics="true" pageId="19" pageNumber="955">Arthrorhaphis</emphasis>
sp. 1
</taxonomicName>
are not included in the key. Characters for the parasitic species have been compiled from
<bibRefCitation id="EF867EA94754FFF7815ED0EAFBA2DB03" author="Obermayer, W." box="[837,1029,251,274]" pageId="19" pageNumber="955" pagination="275 - 333" refId="ref23166" refString="Obermayer, W. 1994. Die Flechtengattung Arthrorhaphis (Arthrorhaphidaceae, Ascomycotina) in Europa und Gronland. Nova Hedwigia 58: 275 - 333." type="journal article" year="1994">Obermayer (1994)</bibRefCitation>
,
<bibRefCitation id="EF867EA94754FFF78615D0EAFAE4DB03" author="Santesson, R. &amp; Tonsberg, T." box="[1038,1347,251,274]" pageId="19" pageNumber="955" pagination="295 - 299" refId="ref24061" refString="Santesson, R. &amp; Tonsberg, T. 1994. Arthrorhaphis aeruginosa and A. olivaceae, two new lichenicolous fungi. Lichenologist 26: 295 - 299. https: // doi. org / 10.1006 / lich. 1994.1021" type="journal article" year="1994">Santesson &amp; Tønsberg (1994)</bibRefCitation>
,
<bibRefCitation id="EF867EA94754FFF78756D0EAFBEADB23" author="Kocourkova, J. &amp; Van den Boom, P. P. G." pageId="19" pageNumber="955" pagination="23 - 35" refId="ref22010" refString="Kocourkova, J. &amp; Van den Boom, P. P. G. 2005. Lichenicolous fungi from the Czech Republic II. Arthrorhaphis arctoparmeliae spec. nov. and some new records for the country. Herzogia 18: 23 - 35." type="journal article" year="2005">Kocourková &amp; Van den Boom (2005)</bibRefCitation>
, and
<bibRefCitation id="EF867EA94754FFF7869ED10AFAA8DB23" author="Etayo, J." box="[1157,1295,283,306]" pageId="19" pageNumber="955" pagination="11 - 535" refId="ref21391" refString="Etayo, J. 2017. Hongos liquenicolas de Ecuador. Opera Lilloana 50: 11 - 535." type="journal article" year="2017">Etayo (2017)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D50D34754FFF1815ED14AFCB8D843" lastPageId="21" lastPageNumber="957" pageId="19" pageNumber="955" type="key">
<key id="FF8654694754FFF1815ED14AFCB8D843" lastPageId="21" lastPageNumber="957" pageId="19" pageNumber="955">
<keyStep id="30E31BFD4754FFF7815ED14AFA6BDBA3" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED14AFA6BDB63" blockId="19.[837,1485,347,1970]" box="[837,1484,347,370]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED14AFA6BDB63" box="[837,1484,347,370]" pageId="19" pageNumber="955">1. Lichenised thallus absent; species strictly lichenicolous .....2</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED16AFA6BDBA3" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED16AFA6BDBA3" pageId="19" pageNumber="955">1. Lichenised thallus present; lichenicolous stages present or absent.........................................................................7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED1AAFA6BDBE3" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED1AAFA6BDBC3" blockId="19.[837,1485,347,1970]" box="[837,1484,443,466]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED1AAFA6BDBC3" box="[837,1484,443,466]" pageId="19" pageNumber="955">2. On the thallus of foliose or squamulose lichens.............3</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED1CAFA6BDBE3" blockId="19.[837,1485,347,1970]" box="[837,1484,475,498]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED1CAFA6BDBE3" box="[837,1484,475,498]" pageId="19" pageNumber="955">2. On the thallus of crustose lichens...................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED1EAFA6AD883" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED1EAFA6AD843" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED1EAFA6AD843" pageId="19" pageNumber="955">
3. Causing aeruginose discolouration of the host thallus; on squamules and podetia of
<taxonomicName id="4C1778DB4754FFF786BED20AFA95D823" box="[1189,1330,539,562]" class="Lecanoromycetes" family="Cladoniaceae" genus="Cladonia" kingdom="Fungi" order="Lecanorales" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="undetermined">
<emphasis id="B963DF4A4754FFF786BED20AFAAFD823" box="[1189,1288,539,562]" italics="true" pageId="19" pageNumber="955">Cladonia</emphasis>
spp
</taxonomicName>
....................... ...................................................
<taxonomicName id="4C1778DB4754FFF786D2D22AFAC5D843" ID-CoL="GZCV" box="[1225,1378,571,594]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="aeruginosa">
<emphasis id="B963DF4A4754FFF786D2D22AFAC5D843" bold="true" box="[1225,1378,571,594]" italics="true" pageId="19" pageNumber="955">A. aeruginosa</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4754FFF7876AD22AFA63D843" box="[1393,1476,571,594]" captionStart="Fig" captionStartId="12.[103,135,1874,1895]" captionTargetBox="[221,1318,187,1839]" captionTargetId="figure-0@12.[221,1318,187,1839]" captionTargetPageId="12" captionText="Fig. 6. A, Arthrorhaphis aeruginosa (TØnsberg 19019, BG, holotype); B, A. arctoparmeliae (Kocourková &amp; Kocourek JK5484 dpl., C); C, A. olivaceae (Santesson 11677, BG, isotype); D, A. muddii (Woods s.n., E, holotype); E, A. grisea (Th.M. Fries s.n., UPS, holotype); F, A. vulgaris (Norrlin s.n., H). — Scale: A–C, 0.5 mm; D–F, 1 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047476" httpUri="https://zenodo.org/record/14047476/files/figure.png" pageId="19" pageNumber="955">Fig. 6A</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED24AFA6AD883" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED24AFA6AD883" pageId="19" pageNumber="955">3. Aeruginose discolouration of the host thallus absent; on other lichens...................................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED28AFA6BD903" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED28AFA6BD8C3" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED28AFA6BD8C3" pageId="19" pageNumber="955">
4. Asci predominantly 4-spored; hymenium 110–150 μm; on
<emphasis id="B963DF4A4754FFF7818ED2AAFB27D8C3" box="[917,1152,699,722]" italics="true" pageId="19" pageNumber="955">
<taxonomicName id="4C1778DB4754FFF7818ED2AAFBB5D8C3" ID-CoL="GZD5" box="[917,1042,699,722]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="phyllobaeis">Phyllobaeis</taxonomicName>
imbricata
</emphasis>
.........................
<taxonomicName id="4C1778DB4754FFF78729D2AAFA6BD8C3" ID-CoL="GZD5" box="[1330,1484,699,722]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="phyllobaeis">
<emphasis id="B963DF4A4754FFF78729D2AAFA6BD8C3" bold="true" box="[1330,1484,699,722]" italics="true" pageId="19" pageNumber="955">A. phyllobaeis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED2CAFA6BD903" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED2CAFA6BD903" pageId="19" pageNumber="955">4. Asci predominantly 6- or 8-spored; hymenium ≥150 μm; on other lichens..............................................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED30AFA6AD983" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED30AFA6AD943" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED30AFA6AD943" pageId="19" pageNumber="955">
5. Asci predominantly 6-spored; hymenium 200–250 μm; on
<taxonomicName id="4C1778DB4754FFF7818ED32AFB20D943" ID-CoL="GB5V" box="[917,1159,827,850]" class="Lecanoromycetes" family="Parmeliaceae" genus="Arctoparmelia" kingdom="Fungi" order="Lecanorales" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="incurva">
<emphasis id="B963DF4A4754FFF7818ED32AFB20D943" box="[917,1159,827,850]" italics="true" pageId="19" pageNumber="955">Arctoparmelia incurva</emphasis>
</taxonomicName>
...
<taxonomicName id="4C1778DB4754FFF786BBD32AFAC3D943" ID-CoL="GZCX" box="[1184,1380,827,850]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="arctoparmeliae">
<emphasis id="B963DF4A4754FFF786BBD32AFAC3D943" bold="true" box="[1184,1380,827,850]" italics="true" pageId="19" pageNumber="955">A. arctoparmeliae</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4754FFF78768D32AFA63D943" box="[1395,1476,827,850]" captionStart="Fig" captionStartId="12.[103,135,1874,1895]" captionTargetBox="[221,1318,187,1839]" captionTargetId="figure-0@12.[221,1318,187,1839]" captionTargetPageId="12" captionText="Fig. 6. A, Arthrorhaphis aeruginosa (TØnsberg 19019, BG, holotype); B, A. arctoparmeliae (Kocourková &amp; Kocourek JK5484 dpl., C); C, A. olivaceae (Santesson 11677, BG, isotype); D, A. muddii (Woods s.n., E, holotype); E, A. grisea (Th.M. Fries s.n., UPS, holotype); F, A. vulgaris (Norrlin s.n., H). — Scale: A–C, 0.5 mm; D–F, 1 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047476" httpUri="https://zenodo.org/record/14047476/files/figure.png" pageId="19" pageNumber="955">Fig. 6B</figureCitation>
)
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</paragraph>
<paragraph id="8BA803584754FFF7815ED34AFA6AD983" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED34AFA6AD983" pageId="19" pageNumber="955">
5. Asci 8-spored; hymenium 150–180 μm; on
<taxonomicName id="4C1778DB4754FFF7875AD34AFC6AD983" ID-CoL="3Z8KW" class="Lecanoromycetes" family="Parmeliaceae" genus="Melanohalea" kingdom="Fungi" order="Lecanorales" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="olivacea">
<emphasis id="B963DF4A4754FFF7875AD34AFC6AD983" italics="true" pageId="19" pageNumber="955">Melanohalea olivacea</emphasis>
</taxonomicName>
........................................
<taxonomicName id="4C1778DB4754FFF786FBD36AFAC3D983" box="[1248,1380,891,914]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="olivaceae">
<emphasis id="B963DF4A4754FFF786FBD36AFAC3D983" bold="true" box="[1248,1380,891,914]" italics="true" pageId="19" pageNumber="955">A. olivaceae</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4754FFF78768D36AFA63D983" box="[1395,1476,891,914]" captionStart="Fig" captionStartId="12.[103,135,1874,1895]" captionTargetBox="[221,1318,187,1839]" captionTargetId="figure-0@12.[221,1318,187,1839]" captionTargetPageId="12" captionText="Fig. 6. A, Arthrorhaphis aeruginosa (TØnsberg 19019, BG, holotype); B, A. arctoparmeliae (Kocourková &amp; Kocourek JK5484 dpl., C); C, A. olivaceae (Santesson 11677, BG, isotype); D, A. muddii (Woods s.n., E, holotype); E, A. grisea (Th.M. Fries s.n., UPS, holotype); F, A. vulgaris (Norrlin s.n., H). — Scale: A–C, 0.5 mm; D–F, 1 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047476" httpUri="https://zenodo.org/record/14047476/files/figure.png" pageId="19" pageNumber="955">Fig. 6C</figureCitation>
)
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</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED38AFA6ADE43" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED38AFA6AD9E3" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED38AFA6AD9E3" pageId="19" pageNumber="955">
6. Hymenium (except epihymenium) ± clear; ascospores 3.5–4.5(–5) μm wide, 10–15-septate; on
<taxonomicName id="4C1778DB4754FFF7872FD3AAFC12D9E3" ID-CoL="9XM6S" class="Lecanoromycetes" family="Icmadophilaceae" genus="Dibaeis" kingdom="Fungi" order="Pertusariales" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="baeomyces">
<emphasis id="B963DF4A4754FFF7872FD3AAFC12D9E3" italics="true" pageId="19" pageNumber="955">Dibaeis baeomyces</emphasis>
</taxonomicName>
................................................
<taxonomicName id="4C1778DB4754FFF786ECD3CAFAC4D9E3" ID-CoL="GZD4" box="[1271,1379,987,1010]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="muddii">
<emphasis id="B963DF4A4754FFF786ECD3CAFAC4D9E3" bold="true" box="[1271,1379,987,1010]" italics="true" pageId="19" pageNumber="955">A. muddii</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4754FFF7876AD3CAFA63D9E3" box="[1393,1476,987,1010]" captionStart="Fig" captionStartId="12.[103,135,1874,1895]" captionTargetBox="[221,1318,187,1839]" captionTargetId="figure-0@12.[221,1318,187,1839]" captionTargetPageId="12" captionText="Fig. 6. A, Arthrorhaphis aeruginosa (TØnsberg 19019, BG, holotype); B, A. arctoparmeliae (Kocourková &amp; Kocourek JK5484 dpl., C); C, A. olivaceae (Santesson 11677, BG, isotype); D, A. muddii (Woods s.n., E, holotype); E, A. grisea (Th.M. Fries s.n., UPS, holotype); F, A. vulgaris (Norrlin s.n., H). — Scale: A–C, 0.5 mm; D–F, 1 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047476" httpUri="https://zenodo.org/record/14047476/files/figure.png" pageId="19" pageNumber="955">Fig. 6D</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED3EAFA6ADE43" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED3EAFA6ADE43" pageId="19" pageNumber="955">
6. Hymenium strongly inspersed; ascospores 2.5–3.5 (–4.5) μm wide, 7–9(13)-septate; on
<taxonomicName id="4C1778DB4754FFF786E0D40AFA17DE23" ID-CoL="KCNR" box="[1275,1456,1051,1074]" class="Lecanoromycetes" family="Baeomycetaceae" genus="Baeomyces" kingdom="Fungi" order="Baeomycetales" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="rufus">
<emphasis id="B963DF4A4754FFF786E0D40AFA17DE23" box="[1275,1456,1051,1074]" italics="true" pageId="19" pageNumber="955">Baeomyces rufus</emphasis>
</taxonomicName>
.... ............................................................
<taxonomicName id="4C1778DB4754FFF7871FD42AFAC1DE43" ID-CoL="GZD3" box="[1284,1382,1083,1106]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="grisea">
<emphasis id="B963DF4A4754FFF7871FD42AFAC1DE43" bold="true" box="[1284,1382,1083,1106]" italics="true" pageId="19" pageNumber="955">A. grisea</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4754FFF7876ED42AFA63DE43" box="[1397,1476,1083,1106]" captionStart="Fig" captionStartId="12.[103,135,1874,1895]" captionTargetBox="[221,1318,187,1839]" captionTargetId="figure-0@12.[221,1318,187,1839]" captionTargetPageId="12" captionText="Fig. 6. A, Arthrorhaphis aeruginosa (TØnsberg 19019, BG, holotype); B, A. arctoparmeliae (Kocourková &amp; Kocourek JK5484 dpl., C); C, A. olivaceae (Santesson 11677, BG, isotype); D, A. muddii (Woods s.n., E, holotype); E, A. grisea (Th.M. Fries s.n., UPS, holotype); F, A. vulgaris (Norrlin s.n., H). — Scale: A–C, 0.5 mm; D–F, 1 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047476" httpUri="https://zenodo.org/record/14047476/files/figure.png" pageId="19" pageNumber="955">Fig. 6E</figureCitation>
)
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</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED44AFA6ADEE3" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED44AFA6BDE83" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED44AFA6BDE83" pageId="19" pageNumber="955">
7. Ascospores of the alpina, jungens or vacillans
<typeStatus id="54ACBDFA4754FFF78772D44AFA3DDE63" box="[1385,1434,1115,1138]" pageId="19" pageNumber="955">type</typeStatus>
; Ca-oxalate crystals in the medulla usually present..............8
</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED48AFA6ADEE3" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED48AFA6ADEE3" pageId="19" pageNumber="955">
7. Ascospores of the citrinella
<typeStatus id="54ACBDFA4754FFF78685D48AFB68DEA3" box="[1182,1231,1179,1202]" pageId="19" pageNumber="955">type</typeStatus>
; Ca-oxalate crystals absent in the medulla or beneath the soredia (
<taxonomicName id="4C1778DB4754FFF78757D4AAFA6BDEC3" ID-CoL="GZCY" box="[1356,1484,1211,1234]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="citrinella">
<emphasis id="B963DF4A4754FFF78757D4AAFA6BDEC3" box="[1356,1484,1211,1234]" italics="true" pageId="19" pageNumber="955">A. citrinella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25062314754FFF7816FD4CAFC2EDEE3" box="[884,905,1243,1266]" pageId="19" pageNumber="955" sensu="lato">s.l.</taxonomicNameLabel>
)................................................................................ 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED4EAFA6BDF43" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED4EAFA6BDF03" blockId="19.[837,1485,347,1970]" box="[837,1484,1275,1298]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED4EAFA6BDF03" box="[837,1484,1275,1298]" pageId="19" pageNumber="955">
8. Ascospores (vacillans
<typeStatus id="54ACBDFA4754FFF78642D4EAFB2EDF03" box="[1113,1161,1275,1298]" pageId="19" pageNumber="955">type</typeStatus>
); soredia absent.....
<taxonomicName id="4C1778DB4754FFF7874BD4EAFA6BDF03" ID-CoL="GZD6" baseAuthorityName="Obermayer" baseAuthorityYear="1995" box="[1360,1484,1275,1298]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="vacillans">
<emphasis id="B963DF4A4754FFF7874BD4EAFA6BDF03" bold="true" box="[1360,1484,1275,1298]" italics="true" pageId="19" pageNumber="955">A. vacillans</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED50AFA6BDF43" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED50AFA6BDF43" pageId="19" pageNumber="955">
8. Ascospores of the alpina or jungens
<typeStatus id="54ACBDFA4754FFF786E8D50AFA83DF23" box="[1267,1316,1307,1330]" pageId="19" pageNumber="955">type</typeStatus>
; soredia present or absent.........................................................................9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED54AFA6ADFC3" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED54AFA6ADF83" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED54AFA6ADF83" pageId="19" pageNumber="955">
9. Ascospores alpina
<typeStatus id="54ACBDFA4754FFF78627D54AFBCEDF63" box="[1084,1129,1371,1394]" pageId="19" pageNumber="955">type</typeStatus>
..................................................... ...........................
<emphasis id="B963DF4A4754FFF78636D56AFA93DF83" bold="true" box="[1069,1332,1403,1426]" pageId="19" pageNumber="955">
<taxonomicName id="4C1778DB4754FFF78636D56AFAAADF83" ID-CoL="GZCW" authority="var. alpina" box="[1069,1293,1403,1426]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="variety" sensu="lato" species="alpina" variety="alpina">
<emphasis id="B963DF4A4754FFF78636D56AFB36DF83" bold="true" box="[1069,1169,1403,1426]" italics="true" pageId="19" pageNumber="955">A. alpina</emphasis>
var.
<emphasis id="B963DF4A4754FFF786D3D56AFAAADF83" bold="true" box="[1224,1293,1403,1426]" italics="true" pageId="19" pageNumber="955">alpina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25062314754FFF7870ED56AFA93DF83" box="[1301,1332,1403,1426]" pageId="19" pageNumber="955" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132C1FDD4754FFF78759D56AFA64DF83" box="[1346,1475,1403,1426]" captionStart="Fig" captionStartId="20.[103,135,1874,1895]" captionTargetBox="[221,1318,187,1839]" captionTargetId="figure-0@20.[221,1318,187,1839]" captionTargetPageId="20" captionText="Fig. 8. A, Arthrorhaphis alpina (Schaer., Lichenes Helvetici Exsiccati 532, G, lectotype); B, A. alpina var. jungens (Lichenotheca Graecensis 23, E, isotype); C, A. alpina (Ohmura 10119, TNS); D, A. alpina var. jungens (Kalb, Lichenes Neotropici 577, M); E, A. alpina (Brusse 4515, UPS); F, “A. septentrionalis” (Hansen 026, C). — Scale: A–F, 1 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047480" httpUri="https://zenodo.org/record/14047480/files/figure.png" pageId="19" pageNumber="955">Fig. 8A,C,E</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED58AFA6ADFC3" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED58AFA6ADFC3" pageId="19" pageNumber="955">
9. Ascospores jungens
<typeStatus id="54ACBDFA4754FFF78655D58AFBDBDFA3" box="[1102,1148,1435,1458]" pageId="19" pageNumber="955">type</typeStatus>
.................................................. ............................
<emphasis id="B963DF4A4754FFF78629D5AAFAECDFC3" bold="true" box="[1074,1355,1467,1490]" pageId="19" pageNumber="955">
<taxonomicName id="4C1778DB4754FFF78629D5AAFA82DFC3" box="[1074,1317,1467,1490]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="variety" sensu="lato" species="alpina" variety="jungens">
<emphasis id="B963DF4A4754FFF78629D5AAFB30DFC3" bold="true" box="[1074,1175,1467,1490]" italics="true" pageId="19" pageNumber="955">A. alpina</emphasis>
var.
<emphasis id="B963DF4A4754FFF786D5D5AAFA82DFC3" bold="true" box="[1230,1317,1467,1490]" italics="true" pageId="19" pageNumber="955">jungens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25062314754FFF78730D5AAFAECDFC3" box="[1323,1355,1467,1490]" pageId="19" pageNumber="955" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132C1FDD4754FFF78742D5AAFA64DFC3" box="[1369,1475,1467,1490]" captionStart="Fig" captionStartId="20.[103,135,1874,1895]" captionTargetBox="[221,1318,187,1839]" captionTargetId="figure-0@20.[221,1318,187,1839]" captionTargetPageId="20" captionText="Fig. 8. A, Arthrorhaphis alpina (Schaer., Lichenes Helvetici Exsiccati 532, G, lectotype); B, A. alpina var. jungens (Lichenotheca Graecensis 23, E, isotype); C, A. alpina (Ohmura 10119, TNS); D, A. alpina var. jungens (Kalb, Lichenes Neotropici 577, M); E, A. alpina (Brusse 4515, UPS); F, “A. septentrionalis” (Hansen 026, C). — Scale: A–F, 1 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047480" httpUri="https://zenodo.org/record/14047480/files/figure.png" pageId="19" pageNumber="955">Fig. 8B,D</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED5CAFA6ADC63" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED5CAFA6BDC03" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED5CAFA6BDC03" pageId="19" pageNumber="955">10. Soredia or granules absent; thallus distinctly bullate-areolate; parasitic stages absent...........................................11</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED60AFA6ADC63" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED60AFA6ADC63" pageId="19" pageNumber="955">10. Soredia or granules present (occasionally indistinct); thallus areolae present or absent; parasitic stages present or absent...........................................................................12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF7815ED66AFA6ADD43" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF7815ED66AFA6ADCE3" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED66AFA6ADCE3" pageId="19" pageNumber="955">
11. Thallus surface ± smooth and shiny; areolae folded in ridges and with central cavity or resting in umbrella-like fashion on black hyphal strands; Australasia ................... ..............................................
<taxonomicName id="4C1778DB4754FFF786B2D6CAFAC4DCE3" ID-CoL="7KZGP" box="[1193,1379,1755,1778]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="catolechioides">
<emphasis id="B963DF4A4754FFF786B2D6CAFAC4DCE3" bold="true" box="[1193,1379,1755,1778]" italics="true" pageId="19" pageNumber="955">A. catolechioides</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4754FFF7876AD6CAFA63DCE3" box="[1393,1476,1755,1778]" captionStart="Fig" captionStartId="15.[151,183,1874,1895]" captionTargetBox="[269,1366,187,1839]" captionTargetId="figure-0@15.[269,1366,187,1839]" captionTargetPageId="15" captionText="Fig. 7. A, Arthrorhaphis catolechioides (Moberg &amp; Owe-Larsson NZ2:5, UPS); B, A. citrinella (Odelvik 10598, S); C, A. citrinella (Frisch 15/No100, TRH); D, A. farinosa (Ohmura 9312, TNS); E, A. farinosa (Thor 33718, UPS); F, A. bullata (Ohmura 10730, TNS). — Scale: A, B, E &amp; F, 1 mm; C, 0.5 mm; D, 5 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047478" httpUri="https://zenodo.org/record/14047478/files/figure.png" pageId="19" pageNumber="955">Fig. 7A</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BA803584754FFF7815ED6EAFA6ADD43" blockId="19.[837,1485,347,1970]" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF7815ED6EAFA6ADD43" pageId="19" pageNumber="955">
11. Thallus surface ± verrucose, matt to slightly shiny; areolae convex to distinctly bullate, not folded in ridges; pale yellow medulla present;
<collectingCountry id="F30043C84754FFF78663D72AFB7BDD43" box="[1144,1244,1851,1874]" name="China" pageId="19" pageNumber="955">East Asia</collectingCountry>
...
<taxonomicName id="4C1778DB4754FFF786E0D72AFAC0DD43" ID-CoL="C92QZ" authorityName="Frisch &amp; Y. Ohmura" authorityYear="2022" box="[1275,1383,1851,1874]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="19" pageNumber="955" phylum="Ascomycota" rank="species" species="bullata">
<emphasis id="B963DF4A4754FFF786E0D72AFAC0DD43" bold="true" box="[1275,1383,1851,1874]" italics="true" pageId="19" pageNumber="955">A. bullata</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4754FFF7876DD72AFA63DD43" box="[1398,1476,1851,1874]" captionStart="Fig" captionStartId="15.[151,183,1874,1895]" captionTargetBox="[269,1366,187,1839]" captionTargetId="figure-0@15.[269,1366,187,1839]" captionTargetPageId="15" captionText="Fig. 7. A, Arthrorhaphis catolechioides (Moberg &amp; Owe-Larsson NZ2:5, UPS); B, A. citrinella (Odelvik 10598, S); C, A. citrinella (Frisch 15/No100, TRH); D, A. farinosa (Ohmura 9312, TNS); E, A. farinosa (Thor 33718, UPS); F, A. bullata (Ohmura 10730, TNS). — Scale: A, B, E &amp; F, 1 mm; C, 0.5 mm; D, 5 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047478" httpUri="https://zenodo.org/record/14047478/files/figure.png" pageId="19" pageNumber="955">Fig. 7F</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4754FFF1815ED74AFCB8DB03" lastPageId="21" lastPageNumber="957" pageId="19" pageNumber="955">
<paragraph id="8BA803584754FFF1815ED74AFCB8DAE3" blockId="19.[837,1485,347,1970]" lastBlockId="21.[151,799,187,594]" lastPageId="21" lastPageNumber="957" pageId="19" pageNumber="955">
<keyLead id="30E6A06D4754FFF1815ED74AFCB8DAE3" lastPageId="21" lastPageNumber="957" pageId="19" pageNumber="955">
12. Thallus forming small compact colonies on saxicolous bryophytes and cyanobacteria; thallus surface entirely disintegrated into finely granular to farinose soredia; parasitic stages absent;
<collectingCountry id="F30043C84752FFF183A4D0AAFD84DAC3" box="[447,547,187,210]" name="China" pageId="21" pageNumber="957">East Asia</collectingCountry>
and Scandinavia .......... ....................................................
<taxonomicName id="4C1778DB4752FFF1803FD0CAFD39DAE3" ID-CoL="C92R3" authorityName="Frisch &amp; Y. Ohmura" authorityYear="2022" box="[548,670,219,242]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="21" pageNumber="957" phylum="Ascomycota" rank="species" species="farinosa">
<emphasis id="B963DF4A4752FFF1803FD0CAFD39DAE3" bold="true" box="[548,670,219,242]" italics="true" pageId="21" pageNumber="957">A. farinosa</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4752FFF180B6D0CAFCB1DAE3" box="[685,790,219,242]" captionStart="Fig" captionStartId="15.[151,183,1874,1895]" captionTargetBox="[269,1366,187,1839]" captionTargetId="figure-0@15.[269,1366,187,1839]" captionTargetPageId="15" captionText="Fig. 7. A, Arthrorhaphis catolechioides (Moberg &amp; Owe-Larsson NZ2:5, UPS); B, A. citrinella (Odelvik 10598, S); C, A. citrinella (Frisch 15/No100, TRH); D, A. farinosa (Ohmura 9312, TNS); E, A. farinosa (Thor 33718, UPS); F, A. bullata (Ohmura 10730, TNS). — Scale: A, B, E &amp; F, 1 mm; C, 0.5 mm; D, 5 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047478" httpUri="https://zenodo.org/record/14047478/files/figure.png" pageId="21" pageNumber="957">Fig. 7D,E</figureCitation>
)
</keyLead>
</paragraph>
<caption id="DF6853D04753FFF0827CD743FCFDDD8F" ID-DOI="http://doi.org/10.5281/zenodo.14047480" ID-Zenodo-Dep="14047480" httpUri="https://zenodo.org/record/14047480/files/figure.png" pageId="20" pageNumber="956" startId="20.[103,135,1874,1895]" targetBox="[221,1318,187,1839]" targetPageId="20" targetType="figure">
<paragraph id="8BA803584753FFF0827CD743FCFDDD8F" blockId="20.[103,1437,1874,1950]" pageId="20" pageNumber="956">
<emphasis id="B963DF4A4753FFF0827CD743FF11DD77" bold="true" box="[103,182,1874,1895]" pageId="20" pageNumber="956">Fig. 8. A</emphasis>
,
<taxonomicName id="4C1778DB4753FFF082DBD743FE24DD77" box="[192,387,1874,1894]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="20" pageNumber="956" phylum="Ascomycota" rank="species" species="alpina">
<emphasis id="B963DF4A4753FFF082DBD743FE24DD77" box="[192,387,1874,1894]" italics="true" pageId="20" pageNumber="956">Arthrorhaphis alpina</emphasis>
</taxonomicName>
(Schaer., Lichenes Helvetici Exsiccati 532, G, lectotype);
<emphasis id="B963DF4A4753FFF08188D743FC05DD77" bold="true" box="[915,930,1874,1894]" pageId="20" pageNumber="956">B</emphasis>
,
<taxonomicName id="4C1778DB4753FFF081B6D743FBD5DD77" box="[941,1138,1874,1894]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="20" pageNumber="956" phylum="Ascomycota" rank="variety" species="alpina" variety="jungens">
<emphasis id="B963DF4A4753FFF081B6D743FBA7DD77" box="[941,1024,1874,1894]" italics="true" pageId="20" pageNumber="956">A. alpina</emphasis>
var.
<emphasis id="B963DF4A4753FFF08631D743FBD5DD77" box="[1066,1138,1874,1894]" italics="true" pageId="20" pageNumber="956">jungens</emphasis>
</taxonomicName>
(Lichenotheca Graecensis 23, E, isotype);
<emphasis id="B963DF4A4753FFF082A5D77FFF68DD93" bold="true" box="[190,207,1902,1922]" pageId="20" pageNumber="956">C</emphasis>
,
<taxonomicName id="4C1778DB4753FFF082C0D77FFE96DD93" box="[219,305,1902,1922]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="20" pageNumber="956" phylum="Ascomycota" rank="species" species="alpina">
<emphasis id="B963DF4A4753FFF082C0D77FFE96DD93" box="[219,305,1902,1922]" italics="true" pageId="20" pageNumber="956">A. alpina</emphasis>
</taxonomicName>
(
<emphasis id="B963DF4A4753FFF08324D77FFE6BDD93" box="[319,460,1902,1922]" italics="true" pageId="20" pageNumber="956">Ohmura 10119</emphasis>
, TNS);
<emphasis id="B963DF4A4753FFF08002D77FFD8DDD93" bold="true" box="[537,554,1902,1922]" pageId="20" pageNumber="956">D</emphasis>
,
<taxonomicName id="4C1778DB4753FFF0802DD77FFCA5DD93" box="[566,770,1902,1922]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="20" pageNumber="956" phylum="Ascomycota" rank="variety" species="alpina" variety="jungens">
<emphasis id="B963DF4A4753FFF0802DD77FFD2BDD93" box="[566,652,1902,1922]" italics="true" pageId="20" pageNumber="956">A. alpina</emphasis>
var.
<emphasis id="B963DF4A4753FFF080A1D77FFCA5DD93" box="[698,770,1902,1922]" italics="true" pageId="20" pageNumber="956">jungens</emphasis>
</taxonomicName>
(Kalb, Lichenes Neotropici 577, M);
<emphasis id="B963DF4A4753FFF0867FD77FFBD4DD93" bold="true" box="[1124,1139,1902,1922]" pageId="20" pageNumber="956">E</emphasis>
,
<taxonomicName id="4C1778DB4753FFF0869BD77FFB71DD93" box="[1152,1238,1902,1922]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="20" pageNumber="956" phylum="Ascomycota" rank="species" species="alpina">
<emphasis id="B963DF4A4753FFF0869BD77FFB71DD93" box="[1152,1238,1902,1922]" italics="true" pageId="20" pageNumber="956">A. alpina</emphasis>
</taxonomicName>
(
<emphasis id="B963DF4A4753FFF086FFD77FFAF0DD93" box="[1252,1367,1902,1922]" italics="true" pageId="20" pageNumber="956">Brusse 4515</emphasis>
, UPS);
<emphasis id="B963DF4A4753FFF0827CD79BFFD2DD8F" bold="true" box="[103,117,1930,1950]" pageId="20" pageNumber="956">F</emphasis>
, “
<taxonomicName id="4C1778DB4753FFF08292D79BFE8CDD8F" box="[137,299,1930,1950]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="20" pageNumber="956" phylum="Ascomycota" rank="species" species="septentrionalis">
<emphasis id="B963DF4A4753FFF08292D79BFE8CDD8F" box="[137,299,1930,1950]" italics="true" pageId="20" pageNumber="956">A. septentrionalis</emphasis>
</taxonomicName>
” (
<emphasis id="B963DF4A4753FFF08359D79BFE15DD8F" box="[322,434,1930,1950]" italics="true" pageId="20" pageNumber="956">Hansen 026</emphasis>
, C). — Scale: A–F, 1 mm. Photos: A. Frisch.
</paragraph>
</caption>
<paragraph id="8BA803584752FFF1828CD0EAFCB8DB03" blockId="21.[151,799,187,594]" box="[151,799,251,274]" pageId="21" pageNumber="957">
<keyLead id="30E6A06D4752FFF1828CD0EAFCB8DB03" box="[151,799,251,274]" pageId="21" pageNumber="957">12. Thallus otherwise; parasitic stages present or absent........13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E31BFD4752FFF1828CD10AFCB8D843" pageId="21" pageNumber="957">
<paragraph id="8BA803584752FFF1828CD10AFCB8DB83" blockId="21.[151,799,187,594]" pageId="21" pageNumber="957">
<keyLead id="30E6A06D4752FFF1828CD10AFCB8DB83" pageId="21" pageNumber="957">
13. Thallus entirely of small loose to compact aggregations of granular soredia on saxicolous bryophytes; parasitic stages absent; Europe and
<collectingCountry id="F30043C84752FFF183FBD14AFD89DB63" box="[480,558,347,370]" name="Iceland" pageId="21" pageNumber="957">Iceland</collectingCountry>
.................................... ...................................................
<taxonomicName id="4C1778DB4752FFF18007D16AFD39DB83" ID-CoL="GZCY" box="[540,670,379,402]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="21" pageNumber="957" phylum="Ascomycota" rank="species" species="citrinella">
<emphasis id="B963DF4A4752FFF18007D16AFD39DB83" bold="true" box="[540,670,379,402]" italics="true" pageId="21" pageNumber="957">A. citrinella</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4752FFF180B7D16AFCB2DB83" box="[684,789,379,402]" captionStart="Fig" captionStartId="15.[151,183,1874,1895]" captionTargetBox="[269,1366,187,1839]" captionTargetId="figure-0@15.[269,1366,187,1839]" captionTargetPageId="15" captionText="Fig. 7. A, Arthrorhaphis catolechioides (Moberg &amp; Owe-Larsson NZ2:5, UPS); B, A. citrinella (Odelvik 10598, S); C, A. citrinella (Frisch 15/No100, TRH); D, A. farinosa (Ohmura 9312, TNS); E, A. farinosa (Thor 33718, UPS); F, A. bullata (Ohmura 10730, TNS). — Scale: A, B, E &amp; F, 1 mm; C, 0.5 mm; D, 5 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047478" httpUri="https://zenodo.org/record/14047478/files/figure.png" pageId="21" pageNumber="957">Fig. 7B,C</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8BA803584752FFF1828CD18AFCB8D843" blockId="21.[151,799,187,594]" pageId="21" pageNumber="957">
<keyLead id="30E6A06D4752FFF1828CD18AFCB8D843" pageId="21" pageNumber="957">
13. Thallus of discrete to confluent areolae on soil, terricolous bryophytes and plant remains, or parasitic on
<taxonomicName id="4C1778DB4752FFF180C4D1AAFE9CDBE3" class="Lecanoromycetes" family="Baeomycetaceae" genus="Baeomyces" kingdom="Fungi" order="Baeomycetales" pageId="21" pageNumber="957" phylum="Ascomycota" rank="species" species="undetermined">
<emphasis id="B963DF4A4752FFF180C4D1AAFEA1DBE3" italics="true" pageId="21" pageNumber="957">Baeomyces</emphasis>
spp.
</taxonomicName>
(rarely on other terricolous lichens); areolae breaking into soredia, rarely completely disintegrated or esorediate; widespread in the Northern Hemisphere........ ........................................................
<taxonomicName id="4C1778DB4752FFF18059D22AFD1ED843" box="[578,697,571,594]" class="Lecanoromycetes" family="Arthrorhaphidaceae" genus="Arthrorhaphis" kingdom="Fungi" pageId="21" pageNumber="957" phylum="Ascomycota" rank="species" species="vulgaris">
<emphasis id="B963DF4A4752FFF18059D22AFD1ED843" bold="true" box="[578,697,571,594]" italics="true" pageId="21" pageNumber="957">A. vulgaris</emphasis>
</taxonomicName>
(
<figureCitation id="132C1FDD4752FFF180D3D22AFCB2D843" box="[712,789,571,594]" captionStart="Fig" captionStartId="12.[103,135,1874,1895]" captionTargetBox="[221,1318,187,1839]" captionTargetId="figure-0@12.[221,1318,187,1839]" captionTargetPageId="12" captionText="Fig. 6. A, Arthrorhaphis aeruginosa (TØnsberg 19019, BG, holotype); B, A. arctoparmeliae (Kocourková &amp; Kocourek JK5484 dpl., C); C, A. olivaceae (Santesson 11677, BG, isotype); D, A. muddii (Woods s.n., E, holotype); E, A. grisea (Th.M. Fries s.n., UPS, holotype); F, A. vulgaris (Norrlin s.n., H). — Scale: A–C, 0.5 mm; D–F, 1 mm. Photos: A. Frisch." figureDoi="http://doi.org/10.5281/zenodo.14047476" httpUri="https://zenodo.org/record/14047476/files/figure.png" pageId="21" pageNumber="957">Fig. 6F</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>