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<document id="E6D832597EB0F49020686497BFD5BFCD" ENCODING="UTF-8" ID-GBIF-Dataset="ed03b2b0-e845-4a8c-b5e2-3a3740e3f98a" ModsDocAuthor="Smith, F." ModsDocDate="1851" ModsDocID="8200" ModsDocOrigin="http://antbase.org/ants/publications/8200/8200.pdf" ModsDocTitle="List of the specimens of British animals in the collection of the British Museum. Part VI. - Hymenoptera Aculeata." ModsID="8200" checkinTime="1243378439508" checkinUser="christiana" docAuthor="Smith, F." docDate="1851" docId="437D1752A4DF2C5B2F22472E639F665F" docLanguage="en" docName="8200" docOrigin="London: British Museum" docSource="http://antbase.org/ants/publications/8200/8200.pdf" docTitle="Formica fuliginosa" docType="treatment" docVersion="14" lastPageNumber="10" masterDocId="F4C959118C8C94377D908658D7932243" masterDocTitle="List of the specimens of British animals in the collection of the British Museum. Part VI. - Hymenoptera Aculeata." pageNumber="10" updateTime="1701312676551" updateUser="plazi">
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<taxonomicName id="1584037832AA56A5AD737BB178C998A8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139091" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" lsidName="Formica fuliginosa Latreille" lsidName-HNS="Formica fuliginosa Latreille" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="species" species="fuliginosa">FORMICA FULIGINOSA</taxonomicName>
.
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<paragraph id="D039DFD6BC4663E0888C14664B19CDCB" pageNumber="10">
<taxonomicName id="0DD756F38AF62005EB20CF4B8E33F0BF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:139091" authority=", Latr" authorityName="Latr" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" lsidName="Formica fuliginosa Latreille" lsidName-HNS="Formica fuliginosa Latreille" order="Hymenoptera" pageNumber="10" phylum="Arthropoda" rank="species" species="fuliginosa">Formica fuliginosa, Latr</taxonomicName>
. Fourm. 140, t. 5, f. 27, [[ male ]], [[ queen ]], [[? ]]. Nyland. Adno. Mon. Form. Boreal. 915,12. Foerster, Hym. Stud. Form. 28, 11.
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.229.105457" ID-Pensoft-Pub="1314-2003-229-185" ID-Pensoft-UUID="7F9045A5C7F05651BA3D05E555438700" ModsDocID="1314-2003-229-185" checkinTime="1690510977206" checkinUser="pensoft" docAuthor="Kubentayev, Serik A., Efimov, Petr G., Alibekov, Daniyar T., Kupriyanov, Andrey N., Izbastina, Klara S., Khalymbetova, Aizhan E. &amp; Perezhogin, Yuri V." docDate="2023" docId="437D23053B4852A1A05327885585CD24" docLanguage="en" docName="PhytoKeys 229: 185-213" docOrigin="PhytoKeys 229" docPubDate="2023-07-27" docSource="http://dx.doi.org/10.3897/phytokeys.229.105457" docTitle="Dactylorhiza salina Soo 1962" docType="treatment" docVersion="1" id="7F9045A5C7F05651BA3D05E555438700" lastPageNumber="185" masterDocId="7F9045A5C7F05651BA3D05E555438700" masterDocTitle="Review of Orchidaceae of the northern part of Kazakhstan" masterLastPageNumber="213" masterPageNumber="185" pageNumber="185" updateTime="1690510977206" updateUser="pensoft">
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<mods:affiliation id="4765C77BE6C6EC5F1958E2322F3A60C3">L. N. Gumilyov Eurasian National University, 2 Satpayev Str., 010000, Astana, Kazakhstan</mods:affiliation>
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<mods:affiliation id="64450D0DD7FCBC42FCBE133B8DE90042">A. Baitursynov Kostanay Regional University, 47 Baytursynov Str., 110000, Kostanay, Kazakhstan</mods:affiliation>
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<taxonomicName id="2FE8F5B974AE851B0594217ED4B76AE4" ID-CoL="6BZPY" LSID="437D2305-3B48-52A1-A053-27885585CD24" authority="(Turcz. ex Lindl.) Soo" authorityName="Soo" authorityYear="1962" baseAuthorityName="Turcz. ex Lindl." class="Liliopsida" family="Orchidaceae" genus="Dactylorhiza" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dactylorhiza salina" order="Asparagales" pageId="0" pageNumber="185" phylum="Tracheophyta" rank="species" species="salina">
Dactylorhiza salina (Turcz. ex Lindl.)
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<paragraph id="A7C87B402771CCD8F5909D11106BAA88" pageId="0" pageNumber="185">Distribution in adjacent reg.</paragraph>
<paragraph id="D6D27CD315C71F0884B1BB14CC7E2AB0" pageId="0" pageNumber="185">Russia (Siberia), Kazakhstan (Altai, Western Tien Shan, Turanskaya lowlands).</paragraph>
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<paragraph id="AF08E58C987BB3D2B6AC0AC07C249DC6" pageId="0" pageNumber="185">Specimens examined and literature records.</paragraph>
<paragraph id="2B926DDAD74D2EC345E7D971E9E8A8B0" pageId="0" pageNumber="185">
Eastern Upland:
<emphasis id="91330F2091331DE5A996505837431BDB" bold="true" pageId="0" pageNumber="185">Karaganda Region</emphasis>
: Aktogay District: near Aktogay,
<date id="3A3A466814444441D68064576D9AA7BF" value="1917-06-24">24 Jun 1917</date>
,
<emphasis id="467713B632B2E857A9EFEAB9D2155501" italics="true" pageId="0" pageNumber="185">Harin s.n..</emphasis>
(AA!). Western Upland:
<emphasis id="4DB3052BF3B9D9446853C6DC59EF524B" bold="true" pageId="0" pageNumber="185">Karaganda Region</emphasis>
(without detailed locality) (
<bibRefCitation id="8E8F12A5D7EA47F64EA8204FAE362339" DOI="https://doi.org/10.17223/978-5-94621-927-3-2020-23" author="Kupriyanov, AN" journalOrPublisher="Geo, Novosibirsk" pageId="0" pageNumber="185" refId="B28" refString="Kupriyanov, AN, 2020. Synopsis of the Flora of the Kazakh Uplands. Geo, Novosibirsk, DOI: https://doi.org/10.17223/978-5-94621-927-3-2020-23" title="Synopsis of the Flora of the Kazakh Uplands." url="https://doi.org/10.17223/978-5-94621-927-3-2020-23" year="2020">Kupriyanov 2020</bibRefCitation>
). Tobol-Ishim:
<emphasis id="9D0E5B42B15D51A454E82AFA1C94C9FC" bold="true" pageId="0" pageNumber="185">Kostanay Region</emphasis>
: Naurzum District:
<normalizedToken id="4C8A280E50676ADD648627A787EC36D1" originalValue="“Naurzum”">&quot;Naurzum&quot;</normalizedToken>
State Nature Reserve, near Biragach,
<date id="EDB0F94CA6527B5649FB263A571CB477" value="1984-06-08">8 Jun 1984</date>
,
<emphasis id="E209FD56BE9EE2437EFA236EC3AB775B" italics="true" pageId="0" pageNumber="185">
<normalizedToken id="865F7427DBC5DB586356C850ED52E1A6" originalValue="Zaugolnova">Zaugol'nova</normalizedToken>
s.n..
</emphasis>
(MOSP!); near Naurzum-Karagai,
<date id="EC46A2984A9CB37DEDA9C1E00B70D031" value="1909-05-20">20 May 1909</date>
,
<emphasis id="C7634C9805024D25D741BE2D0A3D8259" italics="true" pageId="0" pageNumber="185">Savich and Kucherovskaya s.n..</emphasis>
(LE!); near Karamenda, the shore of Sarymoyin Lake,
<date id="BC90A6A20013F99FDA2B31E17DE8678A" value="1911-07-01">1 Jul 1911</date>
,
<emphasis id="D077FAA8D89C6134947CDCEA1F389C34" italics="true" pageId="0" pageNumber="185">Borodin s.n..</emphasis>
(LE!); Egin-Bulak spring, north of Naurzum Forest,
<date id="CC72C5C4470A71DD0FB526B591A37A0B" value="1934-06-22">22 Jun 1934</date>
,
<emphasis id="55864E741B6FB8028BDD95F3D35E8885" italics="true" pageId="0" pageNumber="185">Pavlov 1396</emphasis>
(MW 0816830!);
<quantity id="72C3C0F252E2D0E278CC1CC3E6C62690" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
south of Aksuat Village,
<date id="B5CF0F5B549917687BB2275B37D788ED" value="1945-06-20">20 Jun 1945</date>
,
<emphasis id="EF1BC06A9DA5FA1B8A74274AC1711084" italics="true" pageId="0" pageNumber="185">Voronov s.n..</emphasis>
(MW 0816882!).
</paragraph>
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<paragraph id="185704777109B4208A398880FD26B7E3" pageId="0" pageNumber="185">Habitat and ecology.</paragraph>
<paragraph id="362835EC7CEF1FF2E3F7250E75375010" pageId="0" pageNumber="185">Lowlands amongst birch trees, damp saline meadows and floodplains of rivers.</paragraph>
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<subSubSection id="A73FA11BD9FB843391F2002E90DA4451" pageId="0" pageNumber="185" type="phenology">
<paragraph id="32BA9A5E8590CE663DCF616CD6306FEF" pageId="0" pageNumber="185">Phenology.</paragraph>
<paragraph id="0053E9D4C7B8836996C63784443C6D82" pageId="0" pageNumber="185">Flowering in Jun-Jul; fruiting in Aug-Sep.</paragraph>
</subSubSection>
<subSubSection id="1CF1A5BCB2935CFFA32DEF6560517E6D" pageId="0" pageNumber="185" type="conservation status">
<paragraph id="F85AED655A61C4DDC3669FB0129CCAD6" pageId="0" pageNumber="185">Conservation status.</paragraph>
<paragraph id="D91CA5CD3A35C4F9B5A0616AB4F52258" pageId="0" pageNumber="185">Not protected.</paragraph>
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<subSubSection id="EA63A240E9A68FF2B3FD4BDD2B72650E" pageId="0" pageNumber="185" type="notes">
<paragraph id="B54FD4A5C01077BE99038ED89A6C63A8" pageId="0" pageNumber="185">Notes.</paragraph>
<paragraph id="8DEB3A3A42320416D08793D451DD54AD" pageId="0" pageNumber="185">
<taxonomicName id="DE1E1A9E52D66AF9268ACC1C642B0213" authorityName="Soo" authorityYear="1962" baseAuthorityName="Turcz. ex Lindl." class="Liliopsida" family="Orchidaceae" genus="Dactylorhiza" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dactylorhiza salina" order="Asparagales" pageId="0" pageNumber="185" phylum="Tracheophyta" rank="species" species="salina">
<emphasis id="B9F93AEC3F46DD3ABD5EE3B5D260F41E" italics="true" pageId="0" pageNumber="185">Dactylorhiza salina</emphasis>
</taxonomicName>
is reported here for the first time for the Tobol-Ishim FR and Kostanay Region.
<taxonomicName id="6F74C1C927E32549CED0DFB81AE39B75" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="D. salina" order="Asparagales" pageId="0" pageNumber="185" phylum="Liliopsida" rank="species" species="salina">
<emphasis id="923BFFB3DFD60A7D39BB934DD83AC08D" italics="true" pageId="0" pageNumber="185">D. salina</emphasis>
</taxonomicName>
was recently erroneously reported for the Turgay FR (
<bibRefCitation id="13208A4C73E349052930A28D4F95DEAA" DOI="https://doi.org/10.3390/d15030369" author="Sumbembayev, AA" journalOrPublisher="Biodiversitas" pageId="0" pageNumber="185" refId="B47" refString="Sumbembayev, AA, Nowak, S, Burzacka-Hinz, A, Kosirog-Ceynowa, A, Szlachetko, DL, 2023. New and noteworthy taxa of the genus Dactylorhiza Necker ex Nevski (Orchidaceae Juss.) in Kazakhstan flora and its response to global warming. Diversity (Basel) 15(3): e369. https://doi.org/10.3390/d15030369" title="New and noteworthy taxa of the genus Dactylorhiza Necker ex Nevski (Orchidaceae Juss.) in Kazakhstan flora and its response to global warming. Diversity (Basel) 15 (3): e 369." url="https://doi.org/10.3390/d15030369" year="2023">Sumbembayev et al. 2023</bibRefCitation>
); in fact, the corresponding localities refer to the Tobol-Ishim FR. There are currently no orchid species known to occur in the Turgay FR.
<taxonomicName id="A6E1DCEE960A0FFFF8F4995FE750AC82" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="D. salina" order="Asparagales" pageId="0" pageNumber="185" phylum="Liliopsida" rank="species" species="salina">
<emphasis id="2E726A2BA093DB5B92CCF04538EA6038" italics="true" pageId="0" pageNumber="185">D. salina</emphasis>
</taxonomicName>
is hardly distinguishable from
<taxonomicName id="5DDE089CCDD7124ECBAECD5A6CFF9AEB" family="Orchidaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="D. umbrosa" order="Asparagales" pageId="0" pageNumber="185" phylum="Liliopsida" rank="species" species="umbrosa">
<emphasis id="2007005C23699E8860F3EFB3670B694E" italics="true" pageId="0" pageNumber="185">D. umbrosa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,144 @@
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.125.34877" ID-GBIF-Dataset="53700efe-1c95-4ea8-ac33-8b9230ca45b4" ID-PMC="PMC6610001" ID-Pensoft-Pub="1314-2003-125-1" ID-Pensoft-UUID="FFF8910AFFD4A824FFC3AF26FFD1FFD5" ID-PubMed="31303810" ID-Zenodo-Dep="3268023" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2003-125-1" ModsDocOrigin="PhytoKeys 125" ModsDocTitle="The Callerya Group redefined and Tribe Wisterieae (Fabaceae) emended based on morphology and data from nuclear and chloroplast DNA sequences" checkinTime="1561619006401" checkinUser="pensoft" docAuthor="Compton, James A., Schrire, Brian D., Koenyves 3, Kalman, Forest, Felix, Malakasi, Panagiota, Sawai Mattapha, &amp; Sirichamorn, Yotsawate" docDate="2019" docId="437D577B0DB73C331D4C45E5765906BD" docLanguage="en" docName="PhytoKeys 125: 1-112" docOrigin="PhytoKeys 125" docSource="http://dx.doi.org/10.3897/phytokeys.125.34877" docTitle="Austrocallerya australis J. Compton &amp; Schrire 2019, comb. nov." docType="treatment" docVersion="7" id="FFF8910AFFD4A824FFC3AF26FFD1FFD5" lastPageNumber="64" masterDocId="FFF8910AFFD4A824FFC3AF26FFD1FFD5" masterDocTitle="The Callerya Group redefined and Tribe Wisterieae (Fabaceae) emended based on morphology and data from nuclear and chloroplast DNA sequences" masterLastPageNumber="112" masterPageNumber="1" pageNumber="63" updateTime="1668139348725" updateUser="ExternalLinkService">
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<mods:title>The Callerya Group redefined and Tribe Wisterieae (Fabaceae) emended based on morphology and data from nuclear and chloroplast DNA sequences</mods:title>
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</mods:role>
<mods:namePart>Schrire, Brian D.</mods:namePart>
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<mods:namePart>Koenyves 3, Kalman</mods:namePart>
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<mods:role>
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</mods:role>
<mods:namePart>Forest, Felix</mods:namePart>
</mods:name>
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<mods:role>
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</mods:role>
<mods:namePart>Malakasi, Panagiota</mods:namePart>
</mods:name>
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<mods:role>
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</mods:role>
<mods:namePart>Sawai Mattapha,</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
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<taxonomicName LSID="5EE7CFC1-FB25-50BB-A2FA-5C9709A2DF37" authority="(Endl.) J. Compton &amp; Schrire" authorityName="J. Compton &amp; Schrire" authorityYear="2019" baseAuthorityName="J. Compton &amp; Schrire" baseAuthorityYear="2019" class="Magnoliopsida" family="Fabaceae" genus="Austrocallerya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Austrocallerya australis" order="Fabales" pageId="62" pageNumber="63" phylum="Tracheophyta" rank="species" species="australis" status="comb. nov.">Austrocallerya australis (Endl.) J.Compton &amp; Schrire</taxonomicName>
<taxonomicNameLabel pageId="62" pageNumber="63">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="reference_group">
<paragraph pageId="62" pageNumber="63">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Pterocarpus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pterocarpus australis" order="Fabales" pageId="62" pageNumber="63" phylum="Tracheophyta" rank="species" species="australis">Pterocarpus australis</taxonomicName>
Endl., Prodr. Fl. Norf.: 94 (1833). Type: Australia, Norfolk Island, &quot;In Insula Norfolk, Ferd. Bauer&quot;, 1804-1805, F.Bauer s.n.., W0046224 (W, lecto.! designated here); W0046223 (W, isolecto.!); W0046225 (W, isolecto.!); &quot;Norfolk Island, Bauer Hb. Brown&quot; K000880984 (K, isolecto.!)
</paragraph>
<paragraph pageId="62" pageNumber="63">
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Millettia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Millettia australis" order="Fabales" pageId="62" pageNumber="63" phylum="Tracheophyta" rank="species" species="australis">Millettia australis</taxonomicName>
(Endl.) Benth. Pl. Jungh. [Miquel] 2: 250 (1852)
</paragraph>
<paragraph pageId="63" pageNumber="64">
<pageBreakToken pageId="63" pageNumber="64" start="start"></pageBreakToken>
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Wisteria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Wisteria australis" order="Fabales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="australis">Wisteria australis</taxonomicName>
(Endl.) F.Muell. Second Systematic Census of Australian Pl.: 68 (1885)
</paragraph>
<paragraph pageId="63" pageNumber="64">
<taxonomicName authorityName="Greene" authorityYear="1891" baseAuthorityName="F.Muell." class="Magnoliopsida" family="Fabaceae" genus="Kraunhia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kraunhia australis" order="Fabales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="australis">Kraunhia australis</taxonomicName>
(Endl.) Greene, Pittonia 2(10): 175 (1891)
</paragraph>
<paragraph pageId="63" pageNumber="64">
<taxonomicName authorityName="Schot" authorityYear="1994" baseAuthorityName="Endl." class="Magnoliopsida" family="Fabaceae" genus="Callerya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Callerya australis" order="Fabales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="australis">Callerya australis</taxonomicName>
(Endl.) Schot, Blumea 39(1): 16 (1994)
</paragraph>
<paragraph pageId="63" pageNumber="64">
=
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Millettia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Millettia maideniana" order="Fabales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="maideniana">Millettia maideniana</taxonomicName>
F.M.Bailey, Queensland Bot. Bull. Agric. 5: 12 (1892). Type: Australia, New South Wales, &quot;Port Macquarie, Bean tree, 16 ins diameter, 25-30 ft high, Nov. 1891,&quot;, J.H.Maiden s.n., BRI-AQ22886 (BRI, holo.!) ≡
<taxonomicName authorityName="F.M.Bailey ex C.Moore" authorityYear="1893" class="Magnoliopsida" family="Fabaceae" genus="Wisteria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Wisteria maideniana" order="Fabales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="maideniana">Wisteria maideniana</taxonomicName>
(F.M.Bailey) C.Moore, Handbook Fl. New South Wales: 517 (1893).
</paragraph>
<paragraph pageId="63" pageNumber="64">
=
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Callerya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Callerya neocaledonica" order="Fabales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="neocaledonica">Callerya neocaledonica</taxonomicName>
I.C.Nielsen &amp; Veillon, Adansonia ser. 3, 27(1): 82 (2005). Type:
<normalizedToken originalValue="Nouvelle-Calédonie">Nouvelle-Caledonie</normalizedToken>
,
<normalizedToken originalValue="Dumbéa">Dumbea</normalizedToken>
, Nakutakoin, western slope of Pic Jacob, c. 150 m, 15 January 1992,
<emphasis italics="true" pageId="63" pageNumber="64">J.M.Veillon</emphasis>
7466, P00625937 (P, holo.!); P00629538 (P, iso.!); P00625939 (P, iso.!); (AAU, iso.), NOU006199 (NOU, iso.).
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="illustration">
<paragraph pageId="63" pageNumber="64">Illustration.</paragraph>
<paragraph pageId="63" pageNumber="64">
Harden in Flora of New South Wales 2: 615 (2000). http://www.floragreatlakes.info/html/rfspecies/callerya.html (
<taxonomicName authorityName="Schot" authorityYear="1994" baseAuthorityName="Endl." class="Magnoliopsida" family="Leguminosae" genus="Callerya" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Callerya australis" order="Fabales" pageId="63" pageNumber="64" phylum="Tracheophyta" rank="species" species="australis">
<emphasis italics="true" pageId="63" pageNumber="64">Callerya australis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="distribution">
<paragraph pageId="63" pageNumber="64">Distribution.</paragraph>
<paragraph pageId="63" pageNumber="64">Australia (Norfolk Island, New South Wales, Queensland); Bougainville Island; Cook Islands; New Britain Island; New Guinea; New Caledonia.</paragraph>
</subSubSection>
<subSubSection pageId="63" pageNumber="64" type="habitat">
<paragraph pageId="63" pageNumber="64">Habitat.</paragraph>
<paragraph pageId="63" pageNumber="64">In rainforest climbing in thickets on forested hillsides at 300 to 1600 m.</paragraph>
</subSubSection>
</treatment>
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<mods:title id="B8DF6A8E0C8A1864C4A57CDD799F2552">Amphicnemis rigiketit sp. nov. from Sarawak, with notes on Amphicnemis remiger Laidlaw, 1912 (Odonata: Zygoptera: Coenagrionidae)</mods:title>
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<emphasis id="F9A0EAB0FFE93248FF6C1C82FD90F91E" bold="true" box="[151,581,1710,1737]" pageId="5" pageNumber="376">
<emphasis id="F9A0EAB0FFE93248FF6C1C82FE42F91E" bold="true" box="[151,407,1710,1737]" italics="true" pageId="5" pageNumber="376">Amphicnemis remiger</emphasis>
<bibRefCitation id="AF454B53FFE93248FE651C82FD90F91E" author="Laidlaw, F. F." box="[414,581,1710,1737]" pageId="5" pageNumber="376" pagination="92 - 99" refId="ref6499" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99., 1 pl." type="journal article" year="1912">Laidlaw, 1912</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="CB6B36A2FFE93248FF6C1CFFFCC4F93A" blockId="5.[151,785,1710,1773]" box="[151,785,1747,1773]" pageId="5" pageNumber="376">Figs (2, 4, 6, 8, 10, 12, 14, 16, 18, 20, 22, 24, 26, 28, 31)</paragraph>
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<emphasis id="F9A0EAB0FFE93248FF6C1D36FE56F8E2" bold="true" box="[151,387,1818,1845]" pageId="5" pageNumber="376">Material examined.</emphasis>
<materialsCitation id="7BBC3CFFFFE93248FE771D30FD0EF88E" ID-GBIF-Occurrence="2460219657" collectingDate="1911-05-11" collectorName="J. C. Moulton &amp; Natural History Museum &amp; London" country="Malaysia" location="Kuala Madalam" municipality="Limbang Division" pageId="5" pageNumber="376" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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.
</emphasis>
<specimenCount id="DDD2FD2BFFE93248FDF21D30FDF5F8E1" box="[521,544,1820,1846]" pageId="5" pageNumber="376" type="male"></specimenCount>
,
<location id="CE0B6079FFE93248FDD51D37FD3DF8E2" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FDD51D37FD3DF8E2" box="[558,744,1819,1845]" country="Malaysia" municipality="Limbang Division" name="Kuala Madalam" pageId="5" pageNumber="376">Kuala Madalam</location>
,
<collectingMunicipality id="2B0FACD8FFE93248FD0E1D37FC13F8E2" box="[757,966,1819,1845]" pageId="5" pageNumber="376">Limbang Division</collectingMunicipality>
,
<collectingCountry id="B3C37632FFE93248FC2F1D37FBE2F8E2" box="[980,1079,1819,1845]" name="Malaysia" pageId="5" pageNumber="376">Sarawak</collectingCountry>
,
<collectingCountry id="B3C37632FFE93248FBBF1D37FB78F8E2" box="[1092,1197,1819,1845]" name="Malaysia" pageId="5" pageNumber="376">Malaysia</collectingCountry>
,
<date id="BF6A1062FFE93248FB401D37FAE5F8E2" box="[1211,1328,1819,1846]" pageId="5" pageNumber="376" value="1911-05-11">
<collectingDate id="AF2EE98AFFE93248FB401D37FAE5F8E2" box="[1211,1328,1819,1846]" pageId="5" pageNumber="376" value="1911-05-11">11 v 1911</collectingDate>
</date>
, leg.
<collectorName id="66215374FFE93248FA8A1D37FF2FF88E" pageId="5" pageNumber="376">J.C. Moulton</collectorName>
, in the
<collectorName id="66215374FFE93248FEA81D13FDA7F88D" box="[339,626,1855,1882]" pageId="5" pageNumber="376">Natural History Museum</collectorName>
,
<collectorName id="66215374FFE93248FD7B1D13FD0EF88E" box="[640,731,1855,1881]" pageId="5" pageNumber="376">London</collectorName>
</materialsCitation>
.
<emphasis id="F9A0EAB0FFE93248FD1C1D12FC71F88E" bold="true" box="[743,932,1854,1881]" pageId="5" pageNumber="376">Other material.</emphasis>
All in authors collection.
<collectingCountry id="B3C37632FFE93248FB2F1D13FAF0F88E" box="[1236,1317,1855,1881]" name="Brunei" pageId="5" pageNumber="376">Brunei</collectingCountry>
:
<materialsCitation id="7BBC3CFFFFE93248FACA1D13FC7FF8A9" ID-GBIF-Occurrence="2460219644" collectingDate="2013-03-02" country="Malaysia" location="Kuala Belai Road" pageId="5" pageNumber="376" specimenCount="1" specimenCount-male="1" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FACA1D13FA8AF88D" box="[1329,1375,1855,1882]" pageId="5" pageNumber="376" type="male">1 ♂</specimenCount>
, peat swamp forest,
<location id="CE0B6079FFE93248FEC71D4FFDD2F8AA" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FEC71D4FFDD2F8AA" box="[316,519,1891,1917]" country="Malaysia" name="Kuala Belai Road" pageId="5" pageNumber="376" stateProvince="Belait">Kuala Belai Road</location>
,
<collectingRegion id="0910F840FFE93248FDE81D4FFD61F8AA" box="[531,692,1891,1917]" country="Brunei" name="Belait" pageId="5" pageNumber="376">Belait District</collectingRegion>
,
<date id="BF6A1062FFE93248FD3B1D4FFCFAF8AA" box="[704,815,1891,1917]" pageId="5" pageNumber="376" value="2013-03-02">
<collectingDate id="AF2EE98AFFE93248FD3B1D4FFCFAF8AA" box="[704,815,1891,1917]" pageId="5" pageNumber="376" value="2013-03-02">2 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE93248FC4D1D4FFA4DF8A9" ID-GBIF-Occurrence="2460219655" box="[950,1432,1891,1918]" collectingDate="2013-03-04" country="Malaysia" location="Kuala Belai Road" pageId="5" pageNumber="376" specimenCount="4" specimenCount-male="4" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FC4D1D4FFC22F8A9" box="[950,1015,1891,1918]" pageId="5" pageNumber="376" type="male">4 ♂♂</specimenCount>
, same location,
<date id="BF6A1062FFE93248FB551D4FFAC8F8AA" box="[1198,1309,1891,1917]" pageId="5" pageNumber="376" value="2013-03-04">
<collectingDate id="AF2EE98AFFE93248FB551D4FFAC8F8AA" box="[1198,1309,1891,1917]" pageId="5" pageNumber="376" value="2013-03-04">4 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE93248FF6C1DABFA89F875" ID-GBIF-Occurrence="2460219656" box="[151,1372,1927,1954]" collectingDate="2013-03-08" country="Malaysia" location="Lumut Pipeline Road East" pageId="5" pageNumber="376" specimenCount="1" specimenCount-female="1" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FF6C1DABFF16F875" box="[151,195,1927,1954]" pageId="5" pageNumber="376" type="female">1 ♀</specimenCount>
, low pH swamp forest accessed from
<location id="CE0B6079FFE93248FD851DA4FC7BF875" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FD851DA4FC7BF875" box="[638,942,1927,1954]" country="Malaysia" name="Lumut Pipeline Road East" pageId="5" pageNumber="376" stateProvince="Belait">Lumut Pipeline Road East</location>
,
<collectingRegion id="0910F840FFE93248FC401DABFB8BF876" box="[955,1118,1927,1953]" country="Brunei" name="Belait" pageId="5" pageNumber="376">Belait District</collectingRegion>
,
<date id="BF6A1062FFE93248FB901DABFB0BF876" box="[1131,1246,1927,1953]" pageId="5" pageNumber="376" value="2013-03-08">
<collectingDate id="AF2EE98AFFE93248FB901DABFB0BF876" box="[1131,1246,1927,1953]" pageId="5" pageNumber="376" value="2013-03-08">8 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE93248FA921DABFB50F811" ID-GBIF-Occurrence="2460219643" collectingDate="2013-03-13" country="Malaysia" location="Lumut Pipeline Road East" pageId="5" pageNumber="376" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FA921DABFA42F875" box="[1385,1431,1927,1954]" pageId="5" pageNumber="376" type="male">1 ♂</specimenCount>
,
<specimenCount id="DDD2FD2BFFE93248FF6C1D87FF16F811" box="[151,195,1963,1990]" pageId="5" pageNumber="376" type="female">1 ♀</specimenCount>
(BRU13_COE103, in tandem with the male), same location,
<date id="BF6A1062FFE93248FC7C1D87FBD2F812" box="[903,1031,1963,1989]" pageId="5" pageNumber="376" value="2013-03-13">
<collectingDate id="AF2EE98AFFE93248FC7C1D87FBD2F812" box="[903,1031,1963,1989]" pageId="5" pageNumber="376" value="2013-03-13">13 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE93248FB691D87FCD8F83D" ID-GBIF-Occurrence="2460219651" collectingDate="2013-03-11" country="Malaysia" location="Labi Road" pageId="5" pageNumber="376" specimenCount="9" specimenCount-male="9" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FB691D87FB00F811" box="[1170,1237,1963,1990]" pageId="5" pageNumber="376" type="male">9 ♂♂</specimenCount>
, peat swamp for- est on
<location id="CE0B6079FFE93248FF191DE3FE8EF83E" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FF191DE3FE8EF83E" box="[226,347,1999,2025]" country="Malaysia" name="Labi Road" pageId="5" pageNumber="376" stateProvince="Belait">Labi Road</location>
,
<collectingRegion id="0910F840FFE93248FE9C1DE3FDDDF83E" box="[359,520,1999,2025]" country="Brunei" name="Belait" pageId="5" pageNumber="376">Belait District</collectingRegion>
,
<date id="BF6A1062FFE93248FDEF1DE3FD44F83E" box="[532,657,1999,2025]" pageId="5" pageNumber="376" value="2013-03-11">
<collectingDate id="AF2EE98AFFE93248FDEF1DE3FD44F83E" box="[532,657,1999,2025]" pageId="5" pageNumber="376" value="2013-03-11">11 iii 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFE9324BFCE31DE3FE32FF64" ID-GBIF-Occurrence="2460219639" collectingDate="2013-05-22" country="Malaysia" lastPageId="6" lastPageNumber="377" location="Seria" pageId="5" pageNumber="376" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Belait">
<specimenCount id="DDD2FD2BFFE93248FCE31DE3FC8FF83D" box="[792,858,1999,2026]" pageId="5" pageNumber="376" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="DDD2FD2BFFE93248FC9C1DE3FC47F83D" box="[871,914,1999,2026]" pageId="5" pageNumber="376" type="female">1 ♀</specimenCount>
, remnant peat swamp forest near
<location id="CE0B6079FFE93248FAEA1DE3FA98F83E" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFE93248FAEA1DE3FA98F83E" box="[1297,1357,1999,2025]" country="Malaysia" name="Seria" pageId="5" pageNumber="376" stateProvince="Belait">Seria</location>
,
<collectingRegion id="0910F840FFE9324BFAA21DE3FF3BFF64" country="Brunei" lastPageId="6" lastPageNumber="377" name="Belait" pageId="5" pageNumber="376">Belait District</collectingRegion>
,
<date id="BF6A1062FFEA324BFF021AB5FEB9FF64" box="[249,364,153,179]" pageId="6" pageNumber="377" value="2013-05-22">
<collectingDate id="AF2EE98AFFEA324BFF021AB5FEB9FF64" box="[249,364,153,179]" pageId="6" pageNumber="377" value="2013-05-22">22 v 2013</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFE091AB5FE75FF00" ID-GBIF-Occurrence="2460219654" collectingDate="2013-05-31" country="Malaysia" location="Rassau area" pageId="6" pageNumber="377" specimenCount="3" specimenCount-male="3" stateProvince="Sabah">
<specimenCount id="DDD2FD2BFFEA324BFE091AB5FDE6FF64" box="[498,563,153,179]" pageId="6" pageNumber="377" type="male">3 ♂♂</specimenCount>
, peat swamp forest near highway,
<location id="CE0B6079FFEA324BFC4D1AB5FBE8FF64" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFC4D1AB5FBE8FF64" box="[950,1085,153,179]" country="Malaysia" name="Rassau area" pageId="6" pageNumber="377" stateProvince="Sabah">Rassau area</location>
, Belait District,
<date id="BF6A1062FFEA324BFB0E1AB5FABDFF64" box="[1269,1384,153,179]" pageId="6" pageNumber="377" value="2013-05-31">
<collectingDate id="AF2EE98AFFEA324BFB0E1AB5FABDFF64" box="[1269,1384,153,179]" pageId="6" pageNumber="377" value="2013-05-31">31 v 2013</collectingDate>
</date>
, leg. RAD.
<collectingCountry id="B3C37632FFEA324BFF1A1A91FE9FFF00" box="[225,330,189,215]" name="Malaysia" pageId="6" pageNumber="377">Malaysia</collectingCountry>
,
<collectingRegion id="0910F840FFEA324BFEAC1A91FE75FF00" box="[343,416,189,215]" country="Malaysia" name="Sabah" pageId="6" pageNumber="377">Sabah</collectingRegion>
</materialsCitation>
:
<materialsCitation id="7BBC3CFFFFEA324BFE501A91FC27FF00" ID-GBIF-Occurrence="2460219647" box="[427,1010,189,215]" collectingDate="2014-05-26" country="Malaysia" location="Klias Forest reserve" pageId="6" pageNumber="377" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="DDD2FD2BFFEA324BFE501A91FE00FF00" box="[427,469,189,215]" pageId="6" pageNumber="377" type="female">1 ♀</specimenCount>
,
<location id="CE0B6079FFEA324BFE1F1A91FD1CFF00" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFE1F1A91FD1CFF00" box="[484,713,189,215]" country="Malaysia" name="Klias Forest reserve" pageId="6" pageNumber="377" stateProvince="Sabah">Klias Forest reserve</location>
,
<date id="BF6A1062FFEA324BFD2F1A91FC9CFF00" box="[724,841,189,215]" pageId="6" pageNumber="377" value="2014-05-26">
<collectingDate id="AF2EE98AFFEA324BFD2F1A91FC9CFF00" box="[724,841,189,215]" pageId="6" pageNumber="377" value="2014-05-26">26 v 2014</collectingDate>
</date>
, leg. unknown
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFC051A91FEA9FF2C" ID-GBIF-Occurrence="2460219653" collectingDate="2014-05-31" country="Malaysia" location="Klias Forest Reserve" pageId="6" pageNumber="377" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sabah">
<specimenCount id="DDD2FD2BFFEA324BFC051A91FBFFFF00" box="[1022,1066,189,215]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
,
<specimenCount id="DDD2FD2BFFEA324BFBCC1A91FBB7FF00" box="[1079,1122,189,215]" pageId="6" pageNumber="377" type="female">1 ♀</specimenCount>
,
<location id="CE0B6079FFEA324BFB8A1A91FA8BFF00" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFB8A1A91FA8BFF00" box="[1137,1374,189,215]" country="Malaysia" name="Klias Forest Reserve" pageId="6" pageNumber="377" stateProvince="Sabah">Klias Forest Reserve</location>
,
<date id="BF6A1062FFEA324BFA911A91FF07FF2C" pageId="6" pageNumber="377" value="2014-05-31">
<collectingDate id="AF2EE98AFFEA324BFA911A91FF07FF2C" pageId="6" pageNumber="377" value="2014-05-31">31 v 2014</collectingDate>
</date>
, leg. unknown
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFE731ACDFA8BFF2C" ID-GBIF-Occurrence="2460219640" box="[392,1374,225,251]" collectingDate="2018-05-28" country="Malaysia" location="Kampung Hiridian Forest Reserve" pageId="6" pageNumber="377" specimenCount="1" specimenCount-male="1">
<specimenCount id="DDD2FD2BFFEA324BFE731ACDFE63FF2C" box="[392,438,225,251]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
,
<location id="CE0B6079FFEA324BFE381ACDFC9EFF2C" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFE381ACDFC9EFF2C" box="[451,843,225,251]" country="Malaysia" name="Kampung Hiridian Forest Reserve" pageId="6" pageNumber="377">Kampung Hiridian Forest Reserve</location>
,
<date id="BF6A1062FFEA324BFCAC1ACDFC18FF2C" box="[855,973,225,251]" pageId="6" pageNumber="377" value="2018-05-28">
<collectingDate id="AF2EE98AFFEA324BFCAC1ACDFC18FF2C" box="[855,973,225,251]" pageId="6" pageNumber="377" value="2018-05-28">28 v 2018</collectingDate>
</date>
, leg. unknown.
<collectingCountry id="B3C37632FFEA324BFB781ACDFB3BFF2C" box="[1155,1262,225,251]" name="Malaysia" pageId="6" pageNumber="377">Malaysia</collectingCountry>
: Sarawak
</materialsCitation>
:
<materialsCitation id="7BBC3CFFFFEA324BFA911ACDFAD0FEC8" ID-GBIF-Occurrence="2460219642" collectingDate="2006-02-11" collectorName="J. Simun" country="Malaysia" county="Headhunters Trail" location="Limbang Division" municipality="Gunung Mulu National Park" pageId="6" pageNumber="377" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="DDD2FD2BFFEA324BFA911ACDFA42FF2C" box="[1386,1431,225,251]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
,
<specimenCount id="DDD2FD2BFFEA324BFF6C1B29FF0DFEC8" box="[151,216,261,287]" pageId="6" pageNumber="377" type="female">2 ♀♀</specimenCount>
,
<collectingCounty id="220A4E2EFFEA324BFF1D1B29FE66FEC8" box="[230,435,261,287]" pageId="6" pageNumber="377">Headhunters Trail</collectingCounty>
,
<collectingMunicipality id="2B0FACD8FFEA324BFE441B29FCD1FEC8" box="[447,772,261,287]" pageId="6" pageNumber="377">Gunung Mulu National Park</collectingMunicipality>
,
<location id="CE0B6079FFEA324BFCEB1B29FC35FEC8" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFCEB1B29FC35FEC8" box="[784,992,261,287]" country="Malaysia" county="Headhunters Trail" municipality="Gunung Mulu National Park" name="Limbang Division" pageId="6" pageNumber="377">Limbang Division</location>
,
<date id="BF6A1062FFEA324BFC171B29FBB4FEC8" box="[1004,1121,261,287]" pageId="6" pageNumber="377" value="2006-02-11">
<collectingDate id="AF2EE98AFFEA324BFC171B29FBB4FEC8" box="[1004,1121,261,287]" pageId="6" pageNumber="377" value="2006-02-11">11 ii 2006</collectingDate>
</date>
, leg.
<collectorName id="66215374FFEA324BFB651B29FAD0FEC8" box="[1182,1285,261,287]" pageId="6" pageNumber="377">J. Simun</collectorName>
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFAEA1B29FAB1FE94" ID-GBIF-Occurrence="2460219649" collectingDate="2006-02-13" collectorName="J. Simun" country="Malaysia" county="Boundary Trail" location="Gunung Mulu National Park" municipality="Mentawei" pageId="6" pageNumber="377" specimenCount="2" specimenCount-male="2">
<specimenCount id="DDD2FD2BFFEA324BFAEA1B29FA86FEC8" box="[1297,1363,261,287]" pageId="6" pageNumber="377" type="male">2 ♂♂</specimenCount>
, Men- tawei
<collectingCounty id="220A4E2EFFEA324BFF271B05FE5EFE94" box="[220,395,297,323]" pageId="6" pageNumber="377">Boundary Trail</collectingCounty>
,
<collectingMunicipality id="2B0FACD8FFEA324BFE6C1B05FDDFFE94" box="[407,522,297,323]" pageId="6" pageNumber="377">Mentawei</collectingMunicipality>
,
<location id="CE0B6079FFEA324BFDEC1B05FC88FE94" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFDEC1B05FC88FE94" box="[535,861,297,323]" country="Malaysia" county="Boundary Trail" municipality="Mentawei" name="Gunung Mulu National Park" pageId="6" pageNumber="377">Gunung Mulu National Park</location>
,
<location id="CE0B6079FFEA324BFC911B05FBEEFE94" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFC911B05FBEEFE94" box="[874,1083,297,323]" country="Malaysia" county="Boundary Trail" municipality="Mentawei" name="Limbang Division" pageId="6" pageNumber="377">Limbang Division</location>
,
<date id="BF6A1062FFEA324BFBB31B05FB6AFE94" box="[1096,1215,297,323]" pageId="6" pageNumber="377" value="2006-02-13">
<collectingDate id="AF2EE98AFFEA324BFBB31B05FB6AFE94" box="[1096,1215,297,323]" pageId="6" pageNumber="377" value="2006-02-13">13 ii 2006</collectingDate>
</date>
, leg.
<collectorName id="66215374FFEA324BFB061B05FAB1FE94" box="[1277,1380,297,323]" pageId="6" pageNumber="377">J. Simun</collectorName>
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFA8A1B05FAB3FEB0" ID-GBIF-Occurrence="2460219648" collectingDate="2011-09-05" country="Malaysia" county="Sungai Dabai" location="Miri Division" municipality="Marudi" pageId="6" pageNumber="377" specimenCount="1" specimenCount-female="1">
<specimenCount id="DDD2FD2BFFEA324BFA8A1B05FA49FE94" box="[1393,1436,297,323]" pageId="6" pageNumber="377" type="female">1 ♀</specimenCount>
(SAR11_12_COE63), dry peat swamp forest near
<collectingCounty id="220A4E2EFFEA324BFD341B61FCBFFEB0" box="[719,874,333,359]" pageId="6" pageNumber="377">Sungai Dabai</collectingCounty>
,
<collectingMunicipality id="2B0FACD8FFEA324BFC8E1B61FC1FFEB0" box="[885,970,333,359]" pageId="6" pageNumber="377">Marudi</collectingMunicipality>
,
<location id="CE0B6079FFEA324BFC2D1B61FBA7FEB0" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFC2D1B61FBA7FEB0" box="[982,1138,333,359]" country="Malaysia" county="Sungai Dabai" municipality="Marudi" name="Miri Division" pageId="6" pageNumber="377">Miri Division</location>
,
<date id="BF6A1062FFEA324BFB851B60FB3EFEB0" box="[1150,1259,332,359]" pageId="6" pageNumber="377" value="2011-09-05">
<collectingDate id="AF2EE98AFFEA324BFB851B60FB3EFEB0" box="[1150,1259,332,359]" pageId="6" pageNumber="377" value="2011-09-05">5 ix 2011</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFA891B61FB69FE5C" ID-GBIF-Occurrence="2460219641" collectingDate="2018-09-06" country="Brunei" location="Trail" pageId="6" pageNumber="377" specimenCount="1" specimenCount-male="1">
<specimenCount id="DDD2FD2BFFEA324BFA891B61FA49FEB0" box="[1394,1436,333,359]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
(SAR11_12_COE332), peat swamp forest on
<collectingCountry id="B3C37632FFEA324BFD641B5DFD3EFE5C" box="[671,747,369,395]" name="Brunei" pageId="6" pageNumber="377">Brunei</collectingCountry>
<location id="CE0B6079FFEA324BFD081B5DFCFEFE5C" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFD081B5DFCFEFE5C" box="[755,811,369,395]" country="Brunei" name="Trail" pageId="6" pageNumber="377">Trail</location>
, Marudi area,
<date id="BF6A1062FFEA324BFC2B1B5DFB95FE5C" box="[976,1088,369,395]" pageId="6" pageNumber="377" value="2018-09-06">
<collectingDate id="AF2EE98AFFEA324BFC2B1B5DFB95FE5C" box="[976,1088,369,395]" pageId="6" pageNumber="377" value="2018-09-06">6 ix 2018</collectingDate>
</date>
, leg. RAD
</materialsCitation>
;
<materialsCitation id="7BBC3CFFFFEA324BFB331B5DFD2CFE78" ID-GBIF-Occurrence="2460219645" collectingDate="2011-09-08" country="Brunei" location="Miri Division" municipality="Marudi" pageId="6" pageNumber="377" specimenCount="1" specimenCount-male="1">
<specimenCount id="DDD2FD2BFFEA324BFB331B5DFB23FE5C" box="[1224,1270,369,395]" pageId="6" pageNumber="377" type="male">1 ♂</specimenCount>
, old rubber on peat, near
<collectingMunicipality id="2B0FACD8FFEA324BFEF01BB9FEB5FE78" box="[267,352,405,431]" pageId="6" pageNumber="377">Marudi</collectingMunicipality>
,
<location id="CE0B6079FFEA324BFE901BB9FDD2FE78" LSID="urn:lsid:plazi:treatment:437D87B4FFE93247FF6C1C82FF09FB10:CE0B6079FFEA324BFE901BB9FDD2FE78" box="[363,519,405,431]" country="Brunei" municipality="Marudi" name="Miri Division" pageId="6" pageNumber="377">Miri Division</location>
,
<date id="BF6A1062FFEA324BFDE81BB9FDABFE78" box="[531,638,405,431]" pageId="6" pageNumber="377" value="2011-09-08">
<collectingDate id="AF2EE98AFFEA324BFDE81BB9FDABFE78" box="[531,638,405,431]" pageId="6" pageNumber="377" value="2011-09-08">8 ix 2011</collectingDate>
</date>
, leg. RAD
</materialsCitation>
.
</paragraph>
<caption id="9FAB662AFFEA324BFF6C1FEAFE6DF9F0" ID-DOI="http://doi.org/10.5281/zenodo.3558211" ID-Zenodo-Dep="3558211" httpUri="https://zenodo.org/record/3558211/files/figure.png" pageId="6" pageNumber="377" startId="6.[151,264,1478,1503]" targetBox="[151,1437,453,1453]" targetPageId="6">
<paragraph id="CB6B36A2FFEA324BFF6C1FEAFE6DF9F0" blockId="6.[151,1437,1478,1575]" pageId="6" pageNumber="377">
<emphasis id="F9A0EAB0FFEA324BFF6C1FEAFE80FA08" bold="true" box="[151,341,1478,1503]" pageId="6" pageNumber="377">FIGURES 1720.</emphasis>
Lateral view of synthorax of
<taxonomicName id="0CD44D21FFEA324BFD721FEBFCC4FA08" authorityName="Selys" authorityYear="1863" box="[649,785,1479,1503]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="genus">
<emphasis id="F9A0EAB0FFEA324BFD721FEBFCC4FA08" box="[649,785,1479,1503]" italics="true" pageId="6" pageNumber="377">Amphicnemis</emphasis>
</taxonomicName>
spp. (17)
<taxonomicName id="0CD44D21FFEA324BFC801FEBFB8EFA08" authorityName="Dow" authorityYear="2019" box="[891,1115,1479,1503]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFEA324BFC801FEBFB8EFA08" box="[891,1115,1479,1503]" italics="true" pageId="6" pageNumber="377">Amphicnemis rigiketit</emphasis>
</taxonomicName>
sp. holotype male: (19)
<taxonomicName id="0CD44D21FFEA324BFAA21FEBFEEEF9D4" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFEA324BFAA21FEBFEEEF9D4" italics="true" pageId="6" pageNumber="377">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFEA324BFEB91FC7FE46F9D5" box="[322,403,1515,1538]" pageId="6" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
, female described; (18) male
<taxonomicName id="0CD44D21FFEA324BFD441FC7FC48F9D4" authorityName="Laidlaw" authorityYear="1912" box="[703,925,1515,1539]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEA324BFD441FC7FC48F9D4" box="[703,925,1515,1539]" italics="true" pageId="6" pageNumber="377">Amphicnemis remiger</emphasis>
</taxonomicName>
(SAR11_12_COE332); (20) female
<taxonomicName id="0CD44D21FFEA324BFAEF1FC7FF33F9F0" authorityName="Laidlaw" authorityYear="1912" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEA324BFAEF1FC7FF33F9F0" italics="true" pageId="6" pageNumber="377">Amphicnemis remiger</emphasis>
</taxonomicName>
(BRU13_COE103).
</paragraph>
</caption>
<paragraph id="CB6B36A2FFEA324BFF3C1C78FCBAF9B8" blockId="6.[151,1437,1620,2044]" box="[199,879,1620,1648]" pageId="6" pageNumber="377">
<emphasis id="F9A0EAB0FFEA324BFF3C1C78FE2CF9B8" bold="true" box="[199,505,1620,1647]" pageId="6" pageNumber="377">Descriptive notes on male</emphasis>
(based on SAR11_12_COE332).
</paragraph>
<paragraph id="CB6B36A2FFEA324BFF3C1C55FA9CF858" blockId="6.[151,1437,1620,2044]" pageId="6" pageNumber="377">
<emphasis id="F9A0EAB0FFEA324BFF3C1C55FED1F944" box="[199,260,1657,1683]" italics="true" pageId="6" pageNumber="377">Head</emphasis>
(
<figureCitation id="53EF2A27FFEA324BFEE81C55FEAFF944" box="[275,378,1657,1683]" captionStart="FIGURES 14" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 14. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="6" pageNumber="377">Figs 2, 4</figureCitation>
). Labium entirely pale. Labrum black, narrowly pale centrally at free margin. Mandible bases black with cream outline except adjacent to genae. Genae pale, this extended narrowly beside compound eye to beyond level of antennae bases. Horizontal surface of clypeus shining black, vertical surface pale with intrusions of black from above (
<figureCitation id="53EF2A27FFEA324BFE771CC9FE07F928" box="[396,466,1765,1791]" captionStart="FIGURES 14" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 14. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="6" pageNumber="377">Fig. 2</figureCitation>
). Vertical face of frons shining black becoming dark metallic green above, but largely occupied by irregular, pale transverse stripe very narrowly divided centrally, just joined to pale colour on genae. Rest of frons, vertex, occiput and underside of head bronzy metallic green. Ocelli whitish. Antenna bases with broad pale stripe on anterior face, this just extending to base of scape, scape black, pale at top, base of pedicel pale, this extended on anterior and posterior faces as stripe running almost to top, remainder brown, flagella missing.
</paragraph>
<paragraph id="CB6B36A2FFEA324AFF3C1DB5FE42FE04" blockId="6.[151,1437,1620,2044]" lastBlockId="7.[151,1437,153,467]" lastPageId="7" lastPageNumber="378" pageId="6" pageNumber="377">
<emphasis id="F9A0EAB0FFEA324BFF3C1DB5FECDF864" box="[199,280,1945,1971]" italics="true" pageId="6" pageNumber="377">Thorax</emphasis>
. Prothorax (
<figureCitation id="53EF2A27FFEA324BFE5A1DB5FDCAF864" box="[417,543,1945,1971]" captionStart="FIGURES 912" captionStartId="4.[151,264,1677,1702]" captionTargetBox="[151,1436,417,1652]" captionTargetId="figure@4.[151,1436,417,1652]" captionTargetPageId="4" captionText="FIGURES 912. Amphicnemis rigiketit sp. nov., prothorax of holotype male: (9), dorsal view; (11) lateral view. Prothorax of male Amphicnemis remiger (SAR11_12_COE332); (10) dorsal view; (12) lateral view." figureDoi="http://doi.org/10.5281/zenodo.3558207" httpUri="https://zenodo.org/record/3558207/files/figure.png" pageId="6" pageNumber="377">Figs 10, 12</figureCitation>
) with pronotum almost entirely dark metallic green apart from pair of small pale marks at rear anterior pronotal lobe, propleuron mostly cream. Posterior pronotal lobe short, collar-like centrally but with rear lateral extremities sharply angulated, directed more to rear than to sides (
<figureCitation id="53EF2A27FFEA324BFBAF1DCDFB73F82C" box="[1108,1190,2017,2043]" captionStart="FIGURES 912" captionStartId="4.[151,264,1677,1702]" captionTargetBox="[151,1436,417,1652]" captionTargetId="figure@4.[151,1436,417,1652]" captionTargetPageId="4" captionText="FIGURES 912. Amphicnemis rigiketit sp. nov., prothorax of holotype male: (9), dorsal view; (11) lateral view. Prothorax of male Amphicnemis remiger (SAR11_12_COE332); (10) dorsal view; (12) lateral view." figureDoi="http://doi.org/10.5281/zenodo.3558207" httpUri="https://zenodo.org/record/3558207/files/figure.png" pageId="6" pageNumber="377">Fig. 12</figureCitation>
). Synthorax (
<figureCitation id="53EF2A27FFEA324BFAC01DCDFA5BF82C" box="[1339,1422,2017,2043]" captionStart="FIGURES 1720" captionStartId="6.[151,264,1478,1503]" captionTargetBox="[151,1437,453,1453]" captionTargetId="figure@6.[151,1437,453,1453]" captionTargetPageId="6" captionText="FIGURES 1720. Lateral view of synthorax of Amphicnemis spp. (17) Amphicnemis rigiketit sp. holotype male: (19) Amphicnemis rigiketit sp. nov., female described; (18) male Amphicnemis remiger (SAR11_12_COE332); (20) female Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558211" httpUri="https://zenodo.org/record/3558211/files/figure.png" pageId="6" pageNumber="377">Fig. 18</figureCitation>
): mesepisternum, mesepimeron, most of mesokatepisternum and metepisternum to top of spiracle, and extending to metepleural suture adjacent to antealar carina, metallic green, remainder of synthorax whitish except dark mark near antealar carina on metepimeron. Legs (right anterior leg absent below trochanter) cream with flexor surface femurs white, poorly defined, incomplete narrow dark stripes on extensor surface femurs, dark brown to black spines, small dark mark lowest part femur and immediately adjacent part flexor surface tibia, black rings between tarsal segments, tips of pale brown claws black. Wings: 12 Px in Fw, 10 (left) or 11 (right) in Hw. Arculus slightly distal to Ax2.
<collectionCode id="ADC5AE67FFEB324AFA7A1B61FA41FEB0" box="[1409,1428,333,359]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="378">R</collectionCode>
<subScript id="575034E7FFEB324AFA6F1B72FA49FEBB" attach="left" box="[1428,1436,350,364]" fontSize="6" pageId="7" pageNumber="378">4</subScript>
arising just distal to subnodus in all wings, IR
<subScript id="575034E7FFEB324AFD681BAEFD4EFE47" attach="left" box="[659,667,386,400]" fontSize="6" pageId="7" pageNumber="378">3</subScript>
joined to it by a short stalk. Pterostigma almost trapezoidal with costal side shorter than anal side, dark brownish grey with whitish border (poorly defined on proximal side), covering ca one underlying cell.
</paragraph>
<caption id="9FAB662AFFEB324AFF6C19DDFC37FBFA" ID-DOI="http://doi.org/10.5281/zenodo.3558213" ID-Zenodo-Dep="3558213" httpUri="https://zenodo.org/record/3558213/files/figure.png" pageId="7" pageNumber="378" startId="7.[151,264,1009,1034]" targetBox="[158,1430,489,984]" targetPageId="7">
<paragraph id="CB6B36A2FFEB324AFF6C19DDFC37FBFA" blockId="7.[151,1436,1009,1070]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF6C19DDFE83FBDD" bold="true" box="[151,342,1009,1034]" pageId="7" pageNumber="378">FIGURES 2122.</emphasis>
Lateral view of pattern of terminal abdominal segments of female
<taxonomicName id="0CD44D21FFEB324AFBEA19DDFB4CFBDE" authorityName="Selys" authorityYear="1863" box="[1041,1177,1009,1033]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="378" phylum="Arthropoda" rank="genus">
<emphasis id="F9A0EAB0FFEB324AFBEA19DDFB4CFBDE" box="[1041,1177,1009,1033]" italics="true" pageId="7" pageNumber="378">Amphicnemis</emphasis>
</taxonomicName>
spp.: (21),
<taxonomicName id="0CD44D21FFEB324AFAEF19DDFF32FBFA" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFEB324AFAEF19DDFF32FBFA" italics="true" pageId="7" pageNumber="378">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFEB324AFF151E3AFEEAFBFA" box="[238,319,1046,1069]" pageId="7" pageNumber="378" rank="species">sp. nov.</taxonomicNameLabel>
, female described; (22)
<taxonomicName id="0CD44D21FFEB324AFDC91E39FCC5FBFA" authorityName="Laidlaw" authorityYear="1912" box="[562,784,1045,1069]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="378" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEB324AFDC91E39FCC5FBFA" box="[562,784,1045,1069]" italics="true" pageId="7" pageNumber="378">Amphicnemis remiger</emphasis>
</taxonomicName>
(BRU13_COE103).
</paragraph>
</caption>
<paragraph id="CB6B36A2FFEB324AFF3C1E70FD55F9B8" blockId="7.[151,1437,1116,2042]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF3C1E70FEE4FBA1" box="[199,305,1116,1142]" italics="true" pageId="7" pageNumber="378">Abdomen</emphasis>
. S1 cream with apical transverse dark mark extended laterally apically. S2 metallic green above, cream below. S36 largely very dark brown above, cream lower laterally, this becoming darker on successive segments. S7 similar but almost entirely dark brown laterally in apical ca one-third. S8 bronzy black above, dark brown below. S9 entirely bronzy black, S10 white except dorsally where mostly brown. Anal appendages (
<figureCitation id="53EF2A27FFEB324AFAE91EE4FF61FAD1" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Figs 24, 26, 28</figureCitation>
) white. Cerci in lateral view with upper branch narrow at base, directed gently upwards from base, then straight before expanding along lower margin after half-length so that apical part is rounded ventrally and at apex (
<figureCitation id="53EF2A27FFEB324AFAC71F3CFA45FAFD" box="[1340,1424,1296,1322]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Fig. 24</figureCitation>
). Lower branch of cercus tapering from base before expanding slightly subapically where directed gently upwards to narrow but rounded tip, a small dorsal black subapical tooth just visible in this view (
<figureCitation id="53EF2A27FFEB324AFB761F74FB36FAA5" box="[1165,1251,1368,1394]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Fig. 24</figureCitation>
). In dorsal view (
<figureCitation id="53EF2A27FFEB324AFF5B1F50FF27FA40" box="[160,242,1404,1431]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Fig. 26</figureCitation>
) upper branch with a small subbasal subtriangular tooth after which contracted along inner margin before expanding again just after half length, the inner margin with an excised appearance between the subbasal tooth and the subapical expansion.A strong ridge runs to the apex along the outer margin, giving the expanded part the appearance of a heel-like shelf below it. In ventral view lower branch of cercus tapering gently from base, turning gently inwards at ca two-thirds length, a small, poorly defined subapical tooth on inner margin after which running straight or slightly outwards to rounded but marrow tip (
<figureCitation id="53EF2A27FFEB324AFD3F1C1CFCCEF99D" box="[708,795,1584,1610]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="7" pageNumber="378">Fig. 28</figureCitation>
). Paraprocts short, pale with short finger like projection directed to rear in dorsal part (lateral view).
</paragraph>
<paragraph id="CB6B36A2FFEB324AFF3C1C54FB80F945" blockId="7.[151,1437,1116,2042]" box="[199,1109,1656,1682]" pageId="7" pageNumber="378">Measurements (mm). Abdomen without anal appendages 34, cercus ca 1, Hw 20.</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFEB3247FF3C1CB7FF2FFF2C" lastPageId="10" lastPageNumber="381" pageId="7" pageNumber="378" type="description">
<paragraph id="CB6B36A2FFEB324AFF3C1CB7FCC2F961" blockId="7.[151,1437,1116,2042]" box="[199,791,1691,1718]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF3C1CB7FE13F961" bold="true" box="[199,454,1691,1718]" pageId="7" pageNumber="378">Description of female</emphasis>
(based on BRU13_COE103).
</paragraph>
<paragraph id="CB6B36A2FFEB324AFF3C1CECFE0AF90D" blockId="7.[151,1437,1116,2042]" box="[199,479,1728,1754]" pageId="7" pageNumber="378">As male except as noted.</paragraph>
<paragraph id="CB6B36A2FFEB324AFF3C1CC8FA9AF929" blockId="7.[151,1437,1116,2042]" box="[199,1359,1764,1790]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF3C1CC8FED1F929" box="[199,260,1764,1790]" italics="true" pageId="7" pageNumber="378">Head</emphasis>
(
<figureCitation id="53EF2A27FFEB324AFEE81CC8FEA2F929" box="[275,375,1764,1790]" captionStart="FIGURES 58" captionStartId="3.[151,264,1370,1395]" captionTargetBox="[151,1437,345,1345]" captionTargetId="figure@3.[151,1437,345,1345]" captionTargetPageId="3" captionText="FIGURES 58. Amphicnemis rigiketit sp. nov., head of female described: (5), dorsal-frontal view; (7) dorsal view. Head of female Amphicnemis remiger (BRU13_COE103); (6) dorsal-frontal view; (8) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558205" httpUri="https://zenodo.org/record/3558205/files/figure.png" pageId="7" pageNumber="378">Figs 6, 8</figureCitation>
). More extensive pale area on labrum, pale stripe vertical face of frons broadly divided.
</paragraph>
<paragraph id="CB6B36A2FFEB324AFF3C1D24FDEFF82D" blockId="7.[151,1437,1116,2042]" pageId="7" pageNumber="378">
<emphasis id="F9A0EAB0FFEB324AFF3C1D24FECDF8F5" box="[199,280,1800,1826]" italics="true" pageId="7" pageNumber="378">Thorax</emphasis>
. Prothorax (
<figureCitation id="53EF2A27FFEB324AFE5E1D24FDF3F8F4" box="[421,550,1800,1827]" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="7" pageNumber="378">Figs 14, 16</figureCitation>
) almost entirely blue, anterior carina of anterior pronotal lobe black. Posterior pronotal lobe collar-like centrally in dorsal view (
<figureCitation id="53EF2A27FFEB324AFD321D00FCC9F891" box="[713,796,1836,1862]" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="7" pageNumber="378">Fig. 14</figureCitation>
), laterally contracted before slightly folded upwards and then extended downwards to rear as a subtriangular projection, free margin running downwards back to base (
<figureCitation id="53EF2A27FFEB324AFA941D7CFF63F858" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="7" pageNumber="378">Fig. 16</figureCitation>
). Postepimeral strip expanded to rear just below posterior pronotal lobe (this indicated with a white line in
<figureCitation id="53EF2A27FFEB324AFA8B1D58FF63F864" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="7" pageNumber="378">Fig. 16</figureCitation>
). Synthorax (
<figureCitation id="53EF2A27FFEB324AFEA91DB4FE70F865" box="[338,421,1944,1970]" captionStart="FIGURES 1720" captionStartId="6.[151,264,1478,1503]" captionTargetBox="[151,1437,453,1453]" captionTargetId="figure@6.[151,1437,453,1453]" captionTargetPageId="6" captionText="FIGURES 1720. Lateral view of synthorax of Amphicnemis spp. (17) Amphicnemis rigiketit sp. holotype male: (19) Amphicnemis rigiketit sp. nov., female described; (18) male Amphicnemis remiger (SAR11_12_COE332); (20) female Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558211" httpUri="https://zenodo.org/record/3558211/files/figure.png" pageId="7" pageNumber="378">Fig. 20</figureCitation>
) bluish with yellowish cast dorsally (not apparent in life). Legs as male but marking on extensor surface femurs better defined as narrow black line. Wings with 12 Px in Fw, 11 Px in Hw, white border of pterostigma well defined on all sides.
</paragraph>
<caption id="9FAB662AFFE43245FF6C1CAAFDCEF930" ID-DOI="http://doi.org/10.5281/zenodo.3558215" ID-Zenodo-Dep="3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="8" pageNumber="379" startId="8.[151,264,1670,1695]" targetBox="[151,1436,181,1644]" targetPageId="8">
<paragraph id="CB6B36A2FFE43245FF6C1CAAFDCEF930" blockId="8.[151,1436,1670,1767]" pageId="8" pageNumber="379">
<emphasis id="F9A0EAB0FFE43245FF6C1CAAFE83F948" bold="true" box="[151,342,1670,1695]" pageId="8" pageNumber="379">FIGURES 2328.</emphasis>
<taxonomicName id="0CD44D21FFE43245FEA61CAAFDE8F949" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[349,573,1670,1694]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFE43245FEA61CAAFDE8F949" box="[349,573,1670,1694]" italics="true" pageId="8" pageNumber="379">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFE43245FDBE1CABFD4DF949" box="[581,664,1671,1694]" pageId="8" pageNumber="379" rank="species">sp. nov.</taxonomicNameLabel>
, anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male
<taxonomicName id="0CD44D21FFE43245FDA71C86FCEEF915" authorityName="Laidlaw" authorityYear="1912" box="[604,827,1706,1730]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE43245FDA71C86FCEEF915" box="[604,827,1706,1730]" italics="true" pageId="8" pageNumber="379">Amphicnemis remiger</emphasis>
</taxonomicName>
(SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm.
</paragraph>
</caption>
<paragraph id="CB6B36A2FFE43245FF3C1D39FCA1F84C" blockId="8.[151,1436,1813,2019]" pageId="8" pageNumber="379">
<emphasis id="F9A0EAB0FFE43245FF3C1D39FEE4F8F8" box="[199,305,1813,1839]" italics="true" pageId="8" pageNumber="379">Abdomen</emphasis>
(
<figureCitation id="53EF2A27FFE43245FEB81D39FE4FF8F8" box="[323,410,1813,1839]" captionStart="FIGURES 2122" captionStartId="7.[151,264,1009,1034]" captionTargetBox="[158,1430,489,984]" captionTargetId="figure@7.[158,1430,489,984]" captionTargetPageId="7" captionText="FIGURES 2122. Lateral view of pattern of terminal abdominal segments of female Amphicnemis spp.: (21), Amphicnemis rigiketit sp. nov., female described; (22) Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558213" httpUri="https://zenodo.org/record/3558213/files/figure.png" pageId="8" pageNumber="379">Fig. 22</figureCitation>
). Dark marks on dorsum not metallic until apical part S7. S8 yellowish basally laterally, otherwise brown laterally, bronzy black dorsally. S9 brown laterally, bronzy black dorsally with irregular bluish white lateral apical mark. S10 and anal appendages almost entirely bluish white. Ovipositor largely cream, not quite reaching level of tips of cerci apart from styles, which are brown.
</paragraph>
<paragraph id="CB6B36A2FFE43245FF3C1D89FBA3F868" blockId="8.[151,1436,1813,2019]" box="[199,1142,1956,1983]" pageId="8" pageNumber="379">Measurements (mm). Abdomen without anal appendages or ovipositor 36, Hw 22.5.</paragraph>
<paragraph id="CB6B36A2FFE43244FF3C1DE5FD3EFE94" blockId="8.[151,1436,1813,2019]" lastBlockId="9.[151,1436,153,323]" lastPageId="9" lastPageNumber="380" pageId="8" pageNumber="379">
<emphasis id="F9A0EAB0FFE43245FF3C1DE5FE70F834" bold="true" box="[199,421,1992,2019]" pageId="8" pageNumber="379">Notes on variation</emphasis>
. As in
<taxonomicName id="0CD44D21FFE43245FE021DE5FDA6F834" authorityName="Dow" authorityYear="2019" box="[505,627,1993,2019]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFE43245FE021DE5FDA6F834" box="[505,627,1993,2019]" italics="true" pageId="8" pageNumber="379">A. rigiketit</emphasis>
</taxonomicName>
the main variation in markings in both sexes is in those of the anterior part of the head. R
<subScript id="575034E7FFE53244FE981A86FEBEFF6F" attach="left" box="[355,363,170,184]" fontSize="6" pageId="9" pageNumber="380">4</subScript>
occasionally arises at, rather than distal to, the subnodus in one or more wings. A very few males have the central part of the posterior pronotal lobe produced into a very short horn-like structure. Two males (not among those mentioned in the previous sentence) have the rear lateral extremities of the posterior pronotal lobe only slightly angulated. The subbasal spine on the upper branch of the male cercus is sometimes just visible in a typical lateral view, rarely is it clearly visible in such a view.
</paragraph>
<caption id="9FAB662AFFE53244FF6C18ECFB61FD0E" ID-DOI="http://doi.org/10.5281/zenodo.3558217" ID-Zenodo-Dep="3558217" box="[151,1204,704,729]" httpUri="https://zenodo.org/record/3558217/files/figure.png" pageId="9" pageNumber="380" startId="9.[151,264,704,729]" targetBox="[248,1340,345,679]" targetPageId="9">
<paragraph id="CB6B36A2FFE53244FF6C18ECFB61FD0E" blockId="9.[151,1204,704,729]" box="[151,1204,704,729]" pageId="9" pageNumber="380">
<emphasis id="F9A0EAB0FFE53244FF6C18ECFE81FD0E" bold="true" box="[151,340,704,729]" pageId="9" pageNumber="380">FIGURES 2930.</emphasis>
Habitus of
<taxonomicName id="0CD44D21FFE53244FE3018ECFD7FFD0F" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[459,682,704,728]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFE53244FE3018ECFD7FFD0F" box="[459,682,704,728]" italics="true" pageId="9" pageNumber="380">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFE53244FD4B18EDFCD7FD0F" box="[688,770,705,728]" pageId="9" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
: (29) holotype male; (30) female paratype.
</paragraph>
</caption>
<caption id="9FAB662AFFE53244FF6C1DA0FEC1F83A" ID-DOI="http://doi.org/10.5281/zenodo.3558219" ID-Zenodo-Dep="3558219" httpUri="https://zenodo.org/record/3558219/files/figure.png" pageId="9" pageNumber="380" startId="9.[151,250,1932,1957]" targetBox="[261,1329,751,1907]" targetPageId="9">
<paragraph id="CB6B36A2FFE53244FF6C1DA0FEC1F83A" blockId="9.[151,1436,1932,2029]" pageId="9" pageNumber="380">
<emphasis id="F9A0EAB0FFE53244FF6C1DA0FEF5F872" bold="true" box="[151,288,1932,1957]" pageId="9" pageNumber="380">FIGURE 31.</emphasis>
Distributions of
<taxonomicName id="0CD44D21FFE53244FE351DA1FD7BF872" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[462,686,1933,1957]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFE53244FE351DA1FD7BF872" box="[462,686,1933,1957]" italics="true" pageId="9" pageNumber="380">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFE53244FD4E1DA1FCD1F873" box="[693,772,1933,1956]" pageId="9" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
(red circle) and
<taxonomicName id="0CD44D21FFE53244FC561DA1FB5EF872" authorityName="Laidlaw" authorityYear="1912" box="[941,1163,1933,1957]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE53244FC561DA1FB5EF872" box="[941,1163,1933,1957]" italics="true" pageId="9" pageNumber="380">Amphicnemis remiger</emphasis>
</taxonomicName>
(yellow square). Note that the problematic record of
<taxonomicName id="0CD44D21FFE53244FE5E1D9DFDC4F81E" authorityName="Laidlaw" authorityYear="1912" box="[421,529,1969,1993]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE53244FE5E1D9DFDC4F81E" box="[421,529,1969,1993]" italics="true" pageId="9" pageNumber="380">A. remiger</emphasis>
</taxonomicName>
from “Murud” (see the Remarks under
<taxonomicName id="0CD44D21FFE53244FC4A1D9DFBCBF81E" authorityName="Laidlaw" authorityYear="1912" box="[945,1054,1969,1993]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE53244FC4A1D9DFBCBF81E" box="[945,1054,1969,1993]" italics="true" pageId="9" pageNumber="380">A. remiger</emphasis>
</taxonomicName>
) is excluded from the distribution of that species.
</paragraph>
</caption>
<paragraph id="CB6B36A2FFE63247FF3C1AB5FF2FFF2C" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
Measurements (mm). Males: 1114 Px in Fw,
<quantity id="0C2C9B47FFE63247FD191AB5FC92FF64" box="[738,839,153,179]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="10" pageNumber="381" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">1012 in</quantity>
Hw, abdomen without anal appendages 3035, Hw 1821. Females: 1114 Px in Fw,
<quantity id="0C2C9B47FFE63247FDD21A91FD45FF00" box="[553,656,189,215]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.048" metricValueMin="2.54" pageId="10" pageNumber="381" unit="in" value="11.0" valueMax="12.0" valueMin="10.0">1012 in</quantity>
Hw, abdomen without anal appendages or ovipositor 3335.5, Hw 2121.5.
</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFE63247FF3C1B29FF09FB10" pageId="10" pageNumber="381" type="discussion">
<paragraph id="CB6B36A2FFE63247FF3C1B29FB30FE20" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
<emphasis id="F9A0EAB0FFE63247FF3C1B29FEE1FEC8" bold="true" box="[199,308,261,287]" pageId="10" pageNumber="381">Remarks</emphasis>
. The
<typeStatus id="146F8800FFE63247FE8C1B29FE0CFEC8" box="[375,473,261,287]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0CD44D21FFE63247FDFB1B29FDACFEC8" authorityName="Laidlaw" authorityYear="1912" box="[512,633,261,287]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FDFB1B29FDACFEC8" box="[512,633,261,287]" italics="true" pageId="10" pageNumber="381">A. remiger</emphasis>
</taxonomicName>
is from somewhere in the vicinity of the mouth of the Madalam River (Kuala Madalam), a tributary of the Limbang River, in Sarawaks Limbang Division (
<bibRefCitation id="AF454B53FFE63247FBA51B05FAD5FE94" author="Laidlaw, F. F." box="[1118,1280,297,323]" pageId="10" pageNumber="381" pagination="92 - 99" refId="ref6499" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99., 1 pl." type="journal article" year="1912">Laidlaw 1912</bibRefCitation>
), collected on the
<collectingCountry id="B3C37632FFE63247FF3B1B61FEF5FEB0" box="[192,288,333,359]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
Museums 1911 expedition to Mount Batu Lawi (
<bibRefCitation id="AF454B53FFE63247FCB01B61FC25FEB0" author="Moulton, J. C." box="[843,1008,333,359]" pageId="10" pageNumber="381" pagination="1 - 58" refId="ref6783" refString="Moulton, J. C. (1912) An expedition to Mount Batu Lawi. Journal of the Straits Branch of the Royal Asiatic Society, 63, 1 - 58." type="journal article" year="1912">Moulton 1912</bibRefCitation>
). Later
<bibRefCitation id="AF454B53FFE63247FBBF1B61FACEFEB0" author="Laidlaw, F. F." box="[1092,1307,333,359]" pageId="10" pageNumber="381" pagination="63 - 80" refId="ref6564" refString="Laidlaw, F. F. (1913) Contributions to a study of the dragonfly fauna of Borneo. Part I. The Corduliinae: The genus Amphicnemis: The legion Protoneura. Proceedings of the Zoological Society of London, 1913, 63 - 80, pl. 4 excl. https: // doi. org / 10.1111 / j. 1096 - 3642.1913. tb 01984. x" type="journal article" year="1913">Laidlaw (1913: 72)</bibRefCitation>
mistakenly gave the type locality as Batu Lawi, possibly believing that Kuala Madalam is actually in the vicinity of the mountain (actually it is more than
<quantity id="0C2C9B47FFE63247FE181BB9FDF9FE78" box="[483,556,405,431]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="381" unit="km" value="70.0">70 km</quantity>
distant from it). I have compared recently collected specimens of
<taxonomicName id="0CD44D21FFE63247FADF1BB9FA48FE78" authorityName="Laidlaw" authorityYear="1912" box="[1316,1437,405,431]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FADF1BB9FA48FE78" box="[1316,1437,405,431]" italics="true" pageId="10" pageNumber="381">A. remiger</emphasis>
</taxonomicName>
directly against the
<typeStatus id="146F8800FFE63247FE8E1B95FE02FE04" box="[373,471,441,467]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
(in the Natural History Museum, London) to confirm their identity. The
<typeStatus id="146F8800FFE63247FAF71B95FABBFE04" box="[1292,1390,441,467]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
is a slightly immature individual, so a fresh mature male specimen was used for the description above.
</paragraph>
<paragraph id="CB6B36A2FFE63247FF3C182DFE11FD50" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
The type locality is at low altitude and it is likely that the
<typeStatus id="146F8800FFE63247FCBC182DFC7CFDCC" box="[839,937,513,539]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
was collected in swamp forest of some type. Most subsequent records, where the habitat is recorded, are also from swamp forest, often peat swamp forest, or streams in swamp or alluvial forest; my own specimens are from such habitats.
<bibRefCitation id="AF454B53FFE63247FBE81865FB11FDB4" author="Orr, A. G." box="[1043,1220,585,611]" pageId="10" pageNumber="381" pagination="167 - 220" refId="ref6844" refString="Orr, A. G. (2001) An annotated checklist of the Odonata of Brunei with ecological notes and descriptions of hitherto unknown males and larvae. International Journal of Odonatology, 4 (2), 167 - 220. https: // doi. org / 10.1080 / 13887890.2001.9748168" type="journal article" year="2001">Orr (2001: 183)</bibRefCitation>
states “common in swampy areas” for
<collectingCountry id="B3C37632FFE63247FE8B1841FE15FD50" box="[368,448,621,647]" name="Brunei" pageId="10" pageNumber="381">Brunei</collectingCountry>
.
</paragraph>
<paragraph id="CB6B36A2FFE63247FF3C18BDFDC9FC7F" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
There is a rather puzzling record of this species in
<bibRefCitation id="AF454B53FFE63247FD0B18BDFC01FD7B" author="Laidlaw, F. F." box="[752,980,657,684]" pageId="10" pageNumber="381" pagination="311 - 342" refId="ref6636" refString="Laidlaw, F. F. (1920) Contributions to a study of the dragonfly fauna of Borneo. Part IV. A list of species known to occur in the island. Proceedings of the Zoological Society of London, 1920, 311 - 342. https: // doi. org / 10.1111 / j. 1469 - 7998.1920. tb 07074. x" type="journal article" year="1920">Laidlaw (1920: 336)</bibRefCitation>
where as well as again incorrectly giving the
<typeStatus id="146F8800FFE63247FF3A1899FF24FD18" box="[193,241,693,719]" pageId="10" pageNumber="381">type</typeStatus>
locality as Batu Lawi, he lists two males, stated to be “imperfect”, from “Murud”, collected on the 20
<superScript id="3CA19BEAFFE63247FA8A1898FAABFD15" attach="left" box="[1393,1406,692,706]" fontSize="6" pageId="10" pageNumber="381">th</superScript>
of
<date id="BF6A1062FFE63247FF6C18F5FE9EFD24" box="[151,331,729,755]" pageId="10" pageNumber="381" value="1914-12">December 1914</date>
. Murud presumably refers to Gunung Murud, the highest mountain in
<collectingCountry id="B3C37632FFE63247FBAC18F5FB6EFD24" box="[1111,1211,729,755]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
. Collectors from the
<collectingCountry id="B3C37632FFE63247FF6C18D1FF22FCC0" box="[151,247,765,791]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
Museum went to Gunung Murud, in late 1914 (
<bibRefCitation id="AF454B53FFE63247FCF418D1FC66FCC0" author="Moulton, J. C." box="[783,947,764,791]" pageId="10" pageNumber="381" refId="ref6816" refString="Moulton, J. C. (1915) Twelfth report on the Sarawak Museum, 1914. Sarawak Government Printing Office, Kuching, 48 pp." type="book" year="1915">Moulton 1915</bibRefCitation>
), but it is not clear if they were still there in December and these specimens may have been collected somewhere on their route back. If these specimens were actually collected in the highlands it seems doubtful that they were actually
<taxonomicName id="0CD44D21FFE63247FC001969FBA6FC88" authorityName="Laidlaw" authorityYear="1912" box="[1019,1139,837,863]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FC001969FBA6FC88" box="[1019,1139,837,863]" italics="true" pageId="10" pageNumber="381">A. remiger</emphasis>
</taxonomicName>
and might actually be the unnamed
<taxonomicName id="0CD44D21FFE63247FEFE1945FE49FC54" authorityName="Selys" authorityYear="1863" box="[261,412,873,899]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="genus">
<emphasis id="F9A0EAB0FFE63247FEFE1945FE49FC54" box="[261,412,873,899]" italics="true" pageId="10" pageNumber="381">Amphicnemis</emphasis>
</taxonomicName>
species recorded in
<bibRefCitation id="AF454B53FFE63247FD841945FC90FC54" author="Dow, R. A. &amp; Reels, G. T. &amp; Ngiam, R. W. J." box="[639,837,872,899]" pageId="10" pageNumber="381" pagination="1 - 17" refId="ref6386" refString="Dow, R. A., Reels, G. T. &amp; Ngiam, R. W. J. (2015) Odonata collected at Usun Apau National Park, Miri Division, Sarawak, Malaysia in April and May 2012. International Dragonfly Fund Report, 79, 1 - 17." type="journal article" year="2015">
Dow
<emphasis id="F9A0EAB0FFE63247FD471946FD38FC54" box="[700,749,873,899]" italics="true" pageId="10" pageNumber="381">et al</emphasis>
. (2015)
</bibRefCitation>
from Usun Apau and the Tama Abu Range (the range to which Gunung Murud belongs).
</paragraph>
<paragraph id="CB6B36A2FFE63247FF3C199DFC7EFB8C" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
Laidlaws description of the male anal appendages of
<taxonomicName id="0CD44D21FFE63247FCD3199DFC75FC1C" authorityName="Laidlaw" authorityYear="1912" box="[808,928,945,971]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FCD3199DFC75FC1C" box="[808,928,945,971]" italics="true" pageId="10" pageNumber="381">A. remiger</emphasis>
</taxonomicName>
is somewhat misleading. He states, referring to the upper branch of the cercus as the upper pair of anal appendages, “The shaft a little bowed with a small tooth at its middle” (
<bibRefCitation id="AF454B53FFE63247FEC619D5FDDDFBC3" author="Laidlaw, F. F." box="[317,520,1017,1044]" pageId="10" pageNumber="381" pagination="92 - 99" refId="ref6499" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99., 1 pl." type="journal article" year="1912">Laidlaw 1912: 96</bibRefCitation>
). However, the illustration of the anal appendages of the
<typeStatus id="146F8800FFE63247FB7C19D5FB3CFBC4" box="[1159,1257,1017,1043]" pageId="10" pageNumber="381" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="AF454B53FFE63247FB0319D5FA4DFBC4" author="Laidlaw, F. F." box="[1272,1432,1017,1043]" pageId="10" pageNumber="381" pagination="92 - 99" refId="ref6499" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99., 1 pl." type="journal article" year="1912">Laidlaw 1912</bibRefCitation>
:
<figureCitation id="53EF2A27FFE63247FF6C1E31FF0FFBE0" box="[151,218,1053,1079]" captionStart="FIGURES 14" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 14. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="10" pageNumber="381">Fig. 4</figureCitation>
) correctly shows the tooth in a subbasal position, as in all specimens that I have examined (including that illustrated here), although the tooth is only rarely visible in lateral view.
</paragraph>
<paragraph id="CB6B36A2FFE63247FF3C1E49FF09FB10" blockId="10.[151,1437,152,1223]" pageId="10" pageNumber="381">
<taxonomicName id="0CD44D21FFE63247FF3C1E49FE6BFBA8" authorityName="Laidlaw" authorityYear="1912" box="[199,446,1125,1151]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE63247FF3C1E49FE6BFBA8" box="[199,446,1125,1151]" italics="true" pageId="10" pageNumber="381">Amphicnemis remiger</emphasis>
</taxonomicName>
appears to have a rather restricted distribution in
<collectingCountry id="B3C37632FFE63247FC091E49FB94FBA8" box="[1010,1089,1125,1151]" name="Brunei" pageId="10" pageNumber="381">Brunei</collectingCountry>
, immediately adjacent parts of
<collectingCountry id="B3C37632FFE63247FF6C1EA5FF22FB74" box="[151,247,1161,1187]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
and extending to the south-west of
<collectingRegion id="0910F840FFE63247FD741EA5FD06FB74" box="[655,723,1161,1187]" country="Malaysia" name="Sabah" pageId="10" pageNumber="381">Sabah</collectingRegion>
(
<figureCitation id="53EF2A27FFE63247FD1F1EA5FCECFB74" box="[740,825,1161,1187]" captionStart="FIGURE 31" captionStartId="9.[151,250,1932,1957]" captionTargetBox="[261,1329,751,1907]" captionTargetId="figure@9.[259,1329,750,1907]" captionTargetPageId="9" captionText="FIGURE 31. Distributions of Amphicnemis rigiketit sp. nov. (red circle) and Amphicnemis remiger (yellow square). Note that the problematic record of A. remiger from “Murud” (see the Remarks under A. remiger) is excluded from the distribution of that species." figureDoi="http://doi.org/10.5281/zenodo.3558219" httpUri="https://zenodo.org/record/3558219/files/figure.png" pageId="10" pageNumber="381">Fig. 31</figureCitation>
). In
<collectingCountry id="B3C37632FFE63247FC961EA5FC18FB74" box="[877,973,1161,1187]" name="Malaysia" pageId="10" pageNumber="381">Sarawak</collectingCountry>
it has not been found west of the Baram River.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,464 @@
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<mods:title id="B5569A9D2EB018CFE90668171C596DEA">Amphicnemis rigiketit sp. nov. from Sarawak, with notes on Amphicnemis remiger Laidlaw, 1912 (Odonata: Zygoptera: Coenagrionidae)</mods:title>
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<treatment id="437D87B4FFED3248FF6C1B05FEBBFF2C" ID-DOI="http://doi.org/10.5281/zenodo.5933226" ID-GBIF-Taxon="160605720" ID-Zenodo-Dep="5933226" LSID="urn:lsid:plazi:treatment:437D87B4FFED3248FF6C1B05FEBBFF2C" httpUri="http://treatment.plazi.org/id/437D87B4FFED3248FF6C1B05FEBBFF2C" lastPageId="5" lastPageNumber="376" pageId="1" pageNumber="372">
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<paragraph id="CB6B36A2FFED324CFF6C1B05FDDDFE94" blockId="1.[151,861,297,359]" box="[151,520,297,324]" pageId="1" pageNumber="372">
<heading id="902381CEFFED324CFF6C1B05FDDDFE94" bold="true" box="[151,520,297,324]" fontSize="11" level="1" pageId="1" pageNumber="372" reason="1">
<emphasis id="F9A0EAB0FFED324CFF6C1B05FDDDFE94" bold="true" box="[151,520,297,324]" pageId="1" pageNumber="372">
<taxonomicName id="0CD44D21FFED324CFF6C1B05FE71FE93" ID-CoL="9WZYF" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[151,420,297,324]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFED324CFF6C1B05FE71FE93" bold="true" box="[151,420,297,324]" italics="true" pageId="1" pageNumber="372">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFED324CFE501B05FDDDFE94" box="[427,520,297,323]" pageId="1" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFED324CFF6C1B61FC88FEB0" box="[151,861,332,359]" pageId="1" pageNumber="372" type="description">
<paragraph id="CB6B36A2FFED324CFF6C1B61FC88FEB0" blockId="1.[151,861,297,359]" box="[151,861,332,359]" pageId="1" pageNumber="372">
(
<figureCitation id="53EF2A27FFED324CFF641B61FEDFFEB0" box="[159,266,333,359]" captionStart="FIGURES 14" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 14. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="1" pageNumber="372">Figs. 1, 3</figureCitation>
,
<figureCitation id="53EF2A27FFED324CFEEC1B60FE96FEB0" box="[279,323,332,359]" captionStart="FIGURES 58" captionStartId="3.[151,264,1370,1395]" captionTargetBox="[151,1437,345,1345]" captionTargetId="figure@3.[151,1437,345,1345]" captionTargetPageId="3" captionText="FIGURES 58. Amphicnemis rigiketit sp. nov., head of female described: (5), dorsal-frontal view; (7) dorsal view. Head of female Amphicnemis remiger (BRU13_COE103); (6) dorsal-frontal view; (8) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558205" httpUri="https://zenodo.org/record/3558205/files/figure.png" pageId="1" pageNumber="372">5, 7</figureCitation>
,
<figureCitation id="53EF2A27FFED324CFEB41B61FE5DFEB0" box="[335,392,333,359]" captionStart="FIGURES 912" captionStartId="4.[151,264,1677,1702]" captionTargetBox="[151,1436,417,1652]" captionTargetId="figure@4.[151,1436,417,1652]" captionTargetPageId="4" captionText="FIGURES 912. Amphicnemis rigiketit sp. nov., prothorax of holotype male: (9), dorsal view; (11) lateral view. Prothorax of male Amphicnemis remiger (SAR11_12_COE332); (10) dorsal view; (12) lateral view." figureDoi="http://doi.org/10.5281/zenodo.3558207" httpUri="https://zenodo.org/record/3558207/files/figure.png" pageId="1" pageNumber="372">9, 11</figureCitation>
,
<figureCitation id="53EF2A27FFED324CFE6F1B61FE09FEB0" box="[404,476,332,359]" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="1" pageNumber="372">13, 15</figureCitation>
,
<figureCitation id="53EF2A27FFED324CFE131B61FDE5FEB0" box="[488,560,333,359]" captionStart="FIGURES 1720" captionStartId="6.[151,264,1478,1503]" captionTargetBox="[151,1437,453,1453]" captionTargetId="figure@6.[151,1437,453,1453]" captionTargetPageId="6" captionText="FIGURES 1720. Lateral view of synthorax of Amphicnemis spp. (17) Amphicnemis rigiketit sp. holotype male: (19) Amphicnemis rigiketit sp. nov., female described; (18) male Amphicnemis remiger (SAR11_12_COE332); (20) female Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558211" httpUri="https://zenodo.org/record/3558211/files/figure.png" pageId="1" pageNumber="372">17, 19</figureCitation>
,
<figureCitation id="53EF2A27FFED324CFDC71B61FD8FFEB0" box="[572,602,333,359]" captionStart="FIGURES 2122" captionStartId="7.[151,264,1009,1034]" captionTargetBox="[158,1430,489,984]" captionTargetId="figure@7.[158,1430,489,984]" captionTargetPageId="7" captionText="FIGURES 2122. Lateral view of pattern of terminal abdominal segments of female Amphicnemis spp.: (21), Amphicnemis rigiketit sp. nov., female described; (22) Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558213" httpUri="https://zenodo.org/record/3558213/files/figure.png" pageId="1" pageNumber="372">21</figureCitation>
,
<figureCitation id="53EF2A27FFED324CFD9D1B61FD0DFEB0" box="[614,728,332,359]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="1" pageNumber="372">23, 25, 27</figureCitation>
,
<figureCitation id="53EF2A27FFED324CFD1F1B61FCF9FEB0" box="[740,812,333,359]" captionStart="FIGURES 2930" captionStartId="9.[151,264,704,729]" captionTargetBox="[248,1340,345,679]" captionTargetId="figure@9.[248,1340,345,679]" captionTargetPageId="9" captionText="FIGURES 2930. Habitus of Amphicnemis rigiketit sp. nov.: (29) holotype male; (30) female paratype." figureDoi="http://doi.org/10.5281/zenodo.3558217" httpUri="https://zenodo.org/record/3558217/files/figure.png" pageId="1" pageNumber="372">29, 30</figureCitation>
,
<figureCitation id="53EF2A27FFED324CFCC31B61FC80FEB0" box="[824,853,333,359]" captionStart="FIGURE 31" captionStartId="9.[151,250,1932,1957]" captionTargetBox="[261,1329,751,1907]" captionTargetId="figure@9.[259,1329,750,1907]" captionTargetPageId="9" captionText="FIGURE 31. Distributions of Amphicnemis rigiketit sp. nov. (red circle) and Amphicnemis remiger (yellow square). Note that the problematic record of A. remiger from “Murud” (see the Remarks under A. remiger) is excluded from the distribution of that species." figureDoi="http://doi.org/10.5281/zenodo.3558219" httpUri="https://zenodo.org/record/3558219/files/figure.png" pageId="1" pageNumber="372">31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFED324CFF6C1BB9FC60FDE8" pageId="1" pageNumber="372" type="materials_examined">
<paragraph id="CB6B36A2FFED324CFF6C1BB9FD90FE20" blockId="1.[151,1437,404,2015]" pageId="1" pageNumber="372">
<materialsCitation id="7BBC3CFFFFED324CFF6C1BB9FD90FE20" ID-GBIF-Occurrence="2460219650" collectingDate="2018-02-21" collectionCode="R" collectorName="A. Dow &amp; Natural History Museum &amp; London." country="Malaysia" county="Malaysian Borneo" elevation="13" latitude="1.4096" location="Kota Samarahan" longLatPrecision="7" longitude="110.4046" municipality="Samarahan Division" pageId="1" pageNumber="372" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="F9A0EAB0FFED324CFF6C1BB9FEDEFE78" bold="true" box="[151,267,405,431]" pageId="1" pageNumber="372">
<typeStatus id="146F8800FFED324CFF6C1BB9FED3FE78" box="[151,262,405,431]" pageId="1" pageNumber="372" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="DDD2FD2BFFED324CFEE81BB9FEE8FE78" box="[275,317,405,431]" pageId="1" pageNumber="372" type="male">1 ♂</specimenCount>
(SAR18_COE58), in forest around stream, near road from
<location id="CE0B6079FFED324CFC201BB9FB40FE78" LSID="urn:lsid:plazi:treatment:437D87B4FFED3248FF6C1B05FEBBFF2C:CE0B6079FFED324CFC201BB9FB40FE78" box="[987,1173,405,431]" country="Malaysia" county="Malaysian Borneo" latitude="1.4096" longLatPrecision="7" longitude="110.4046" municipality="Samarahan Division" name="Kota Samarahan" pageId="1" pageNumber="372">Kota Samarahan</location>
to
<location id="CE0B6079FFED324CFB411BB9FAC1FE78" LSID="urn:lsid:plazi:treatment:437D87B4FFED3248FF6C1B05FEBBFF2C:CE0B6079FFED324CFB411BB9FAC1FE78" box="[1210,1300,405,431]" country="Malaysia" county="Malaysian Borneo" latitude="1.4096" longLatPrecision="7" longitude="110.4046" municipality="Samarahan Division" name="Siburan" pageId="1" pageNumber="372">Siburan</location>
,
<collectingMunicipality id="2B0FACD8FFED324CFADB1BB9FF2FFE04" pageId="1" pageNumber="372">Samarahan Division</collectingMunicipality>
,
<collectingCountry id="B3C37632FFED324CFEFD1B95FEBCFE04" box="[262,361,441,467]" name="Malaysia" pageId="1" pageNumber="372">Sarawak</collectingCountry>
,
<collectingCounty id="220A4E2EFFED324CFE8E1B95FD93FE04" box="[373,582,441,467]" pageId="1" pageNumber="372">Malaysian Borneo</collectingCounty>
,
<geoCoordinate id="AEE05065FFED324CFDAA1B95FD60FE04" box="[593,693,441,467]" degrees="1.4096" direction="north" orientation="latitude" pageId="1" pageNumber="372" precision="5" value="1.4096">1.4096N</geoCoordinate>
,
<geoCoordinate id="AEE05065FFED324CFD3A1B95FCE9FE04" box="[705,828,441,467]" degrees="110.4046" direction="east" orientation="longitude" pageId="1" pageNumber="372" precision="5" value="110.4046">110.4046E</geoCoordinate>
, ca
<elevation id="40F9D191FFED324CFC931B95FC02FE04" box="[872,983,441,467]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="372" unit="m" value="13.0">
<quantity id="0C2C9B47FFED324CFC931B95FC4FFE04" box="[872,922,441,467]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="372" unit="m" value="13.0">13m</quantity>
a.s.l.
</elevation>
,
<date id="BF6A1062FFED324CFC1F1B95FB8CFE04" box="[996,1113,441,467]" pageId="1" pageNumber="372" value="2018-02-21">
<collectingDate id="AF2EE98AFFED324CFC1F1B95FB8CFE04" box="[996,1113,441,467]" pageId="1" pageNumber="372" value="2018-02-21">21 ii 2018</collectingDate>
</date>
, leg.
<collectionCode id="ADC5AE67FFED324CFB6E1B95FB7EFE04" box="[1173,1195,441,467]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="372">R</collectionCode>
.
<collectorName id="66215374FFED324CFB551B95FADDFE04" box="[1198,1288,441,467]" pageId="1" pageNumber="372">A. Dow</collectorName>
, deposited in the
<collectorName id="66215374FFED324CFF3B1BF1FE0EFE20" box="[192,475,477,503]" pageId="1" pageNumber="372">Natural History Museum</collectorName>
,
<collectorName id="66215374FFED324CFE1D1BF1FD90FE20" box="[486,581,477,503]" pageId="1" pageNumber="372">London.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="CB6B36A2FFED324CFF3C182DFC60FDE8" blockId="1.[151,1437,404,2015]" pageId="1" pageNumber="372">
<emphasis id="F9A0EAB0FFED324CFF3C182DFE93FDCC" bold="true" box="[199,326,513,539]" pageId="1" pageNumber="372">
<typeStatus id="146F8800FFED324CFF3C182DFE94FDCC" box="[199,321,513,539]" pageId="1" pageNumber="372" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="DDD2FD2BFFED324CFEAB182DFE47FDCC" box="[336,402,513,539]" pageId="1" pageNumber="372" type="male">6 ♂♂</specimenCount>
(SAR18_COE53-57, 59),
<materialsCitation id="7BBC3CFFFFED324CFD3D182CFEDEFDE8" ID-GBIF-Occurrence="2460219646" collectionCode="R" collectorName="A. Dow" pageId="1" pageNumber="372" specimenCount="5" specimenCount-female="5" typeStatus="holotype">
<specimenCount id="DDD2FD2BFFED324CFD3D182CFCDDFDCC" box="[710,776,512,539]" pageId="1" pageNumber="372" type="female">5 ♀♀</specimenCount>
(SAR18_COE60-64), location and date as
<typeStatus id="146F8800FFED324CFAFB182DFAB0FDCC" box="[1280,1381,513,539]" pageId="1" pageNumber="372" type="holotype">holotype</typeStatus>
, leg.
<collectionCode id="ADC5AE67FFED324CFF6C1809FF78FDE8" box="[151,173,549,575]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="372">R</collectionCode>
.
<collectorName id="66215374FFED324CFF4A1809FEDEFDE8" box="[177,267,549,575]" pageId="1" pageNumber="372">A. Dow</collectorName>
</materialsCitation>
,
<materialsCitation id="7BBC3CFFFFED324CFEED1809FC64FDE8" ID-GBIF-Occurrence="2460219652" box="[278,945,549,575]" country="Malaysia" location="Sarawak" pageId="1" pageNumber="372" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="DDD2FD2BFFED324CFEED1809FE97FDE8" box="[278,322,549,575]" pageId="1" pageNumber="372" type="male">1 ♂</specimenCount>
,
<specimenCount id="DDD2FD2BFFED324CFEB51809FEADFDE8" box="[334,376,549,575]" pageId="1" pageNumber="372" type="female">1 ♀</specimenCount>
to be deposited in the
<collectingCountry id="B3C37632FFED324CFD8E1809FD00FDE8" box="[629,725,549,575]" name="Malaysia" pageId="1" pageNumber="372">Sarawak</collectingCountry>
Museum, Kuching
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFED324CFF3C1865FB20FC88" pageId="1" pageNumber="372" type="etymology">
<paragraph id="CB6B36A2FFED324CFF3C1865FB20FC88" blockId="1.[151,1437,404,2015]" pageId="1" pageNumber="372">
<emphasis id="F9A0EAB0FFED324CFF3C1865FE9BFDB4" bold="true" box="[199,334,585,611]" pageId="1" pageNumber="372">Etymology.</emphasis>
The species epithet,
<taxonomicName id="0CD44D21FFED324CFDC31865FD44FDB4" authorityName="Dow" authorityYear="2019" box="[568,657,585,611]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFED324CFDC31865FD44FDB4" box="[568,657,585,611]" italics="true" pageId="1" pageNumber="372">rigiketit</emphasis>
</taxonomicName>
, is a noun in apposition, formed from the names of two highly skilled Iban (the largest indigenous group in
<collectingCountry id="B3C37632FFED324CFDBC1841FD7CFD5F" box="[583,681,621,648]" name="Malaysia" pageId="1" pageNumber="372">Sarawak</collectingCountry>
) collectors, Ketit and Rigi, employed by the
<collectingCountry id="B3C37632FFED324CFB491841FAC7FD50" box="[1202,1298,621,647]" name="Malaysia" pageId="1" pageNumber="372">Sarawak</collectingCountry>
Museum in the early Twentieth Century. Both collectors took part in the expedition during which the
<typeStatus id="146F8800FFED324CFB6818BDFB20FD7C" box="[1171,1269,657,683]" pageId="1" pageNumber="372" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0CD44D21FFED324CFAE618BDFA43FD7C" authorityName="Laidlaw" authorityYear="1912" box="[1309,1430,657,683]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFED324CFAE618BDFA43FD7C" box="[1309,1430,657,683]" italics="true" pageId="1" pageNumber="372">A. remiger</emphasis>
</taxonomicName>
, the closest species to
<taxonomicName id="0CD44D21FFED324CFE771899FDD0FD18" authorityName="Dow" authorityYear="2019" box="[396,517,693,719]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFED324CFE771899FDD0FD18" box="[396,517,693,719]" italics="true" pageId="1" pageNumber="372">A. rigiketit</emphasis>
</taxonomicName>
, was collected. Although the
<typeStatus id="146F8800FFED324CFCAE1899FC62FD18" box="[853,951,693,719]" pageId="1" pageNumber="372" type="holotype">holotype</typeStatus>
of
<taxonomicName id="0CD44D21FFED324CFC251899FB83FD18" authorityName="Laidlaw" authorityYear="1912" box="[990,1110,693,719]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFED324CFC251899FB83FD18" box="[990,1110,693,719]" italics="true" pageId="1" pageNumber="372">A. remiger</emphasis>
</taxonomicName>
is attributed to J.C. Moulton (likely merely because it was Moulton who sent the specimen to F.F. Laidlaw and it was not customary at the time to explicitly name native collectors) it is entirely possible that it was actually collected by one of these two collectors. In any case the contributions of the
<collectingCountry id="B3C37632FFED324CFDA5190DFD6BFCEC" box="[606,702,801,827]" name="Malaysia" pageId="1" pageNumber="372">Sarawak</collectingCountry>
Museums Iban collectors to our knowledge of Sarawaks natural history are largely undocumented but significant, see
<bibRefCitation id="AF454B53FFED324CFD141969FC5AFC88" author="Morris, J. R." box="[751,911,837,863]" pageId="1" pageNumber="372" pagination="6 - 8" refId="ref6731" refString="Morris, J. R. (2018) The contributions of Sarawakian collecting networks to the history of science in Southeast Asia c. 1890 - 1940. Paper presented at the Borneo Research Council Conference, Kuching, Sarawak, 6 - 8 August 2018. [unkown pagination]" type="proceedings paper" year="2018">Morris (2018)</bibRefCitation>
, and deserve wider recognition.
</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFED324EFF3C1944FEDAF9F0" lastPageId="3" lastPageNumber="374" pageId="1" pageNumber="372" type="description">
<paragraph id="CB6B36A2FFED324CFF3C1944FA9CFBA8" blockId="1.[151,1437,404,2015]" pageId="1" pageNumber="372">
<emphasis id="F9A0EAB0FFED324CFF3C1944FE33FC54" bold="true" box="[199,486,872,899]" pageId="1" pageNumber="372">
Description of
<typeStatus id="146F8800FFED324CFE821945FE35FC54" box="[377,480,873,899]" pageId="1" pageNumber="372" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="F9A0EAB0FFED324CFE151945FDFEFC54" box="[494,555,873,899]" italics="true" pageId="1" pageNumber="372">Head</emphasis>
(
<figureCitation id="53EF2A27FFED324CFDC01945FD74FC54" box="[571,673,873,899]" captionStart="FIGURES 14" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 14. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="1" pageNumber="372">Figs 1, 3</figureCitation>
). Labium entirely pale. Labrum black, cream in lower ca one third. Mandible bases cream with dark central patch. Genae cream, this extended narrowly beside compound eye to beyond level of antennae bases. Horizontal surface of clypeus shining black, vertical surface pale brown with intrusions of black from above (
<figureCitation id="53EF2A27FFED324CFE2919F9FDC3FC38" box="[466,534,981,1007]" captionStart="FIGURES 14" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 14. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="1" pageNumber="372">Fig. 1</figureCitation>
). Vertical face of frons shining black becoming dark metallic green above, but largely occupied by irregular, complete cream transverse stripe joined to pale colour on genae. Rest of frons, vertex, occiput and underside of head bronzy metallic green. Ocelli whitish. Antenna bases with broad yellowish stripe on anterior face, this just extending to base of scape, scape black, yellowish at top, base of pedicel yellowish, this extended on anterior and posterior faces as stripe running almost to top, remainder brown, flagella missing.
</paragraph>
<paragraph id="CB6B36A2FFED324CFF3C1EA5FA97FA30" blockId="1.[151,1437,404,2015]" pageId="1" pageNumber="372">
<emphasis id="F9A0EAB0FFED324CFF3C1EA5FECDFB74" box="[199,280,1161,1187]" italics="true" pageId="1" pageNumber="372">Thorax</emphasis>
. Prothorax (
<figureCitation id="53EF2A27FFED324CFE5F1EA5FDC1FB74" box="[420,532,1161,1187]" captionStart="FIGURES 912" captionStartId="4.[151,264,1677,1702]" captionTargetBox="[151,1436,417,1652]" captionTargetId="figure@4.[151,1436,417,1652]" captionTargetPageId="4" captionText="FIGURES 912. Amphicnemis rigiketit sp. nov., prothorax of holotype male: (9), dorsal view; (11) lateral view. Prothorax of male Amphicnemis remiger (SAR11_12_COE332); (10) dorsal view; (12) lateral view." figureDoi="http://doi.org/10.5281/zenodo.3558207" httpUri="https://zenodo.org/record/3558207/files/figure.png" pageId="1" pageNumber="372">Figs 9, 11</figureCitation>
) with anterior pronotal lobe cream except anterior carina black, middle and posterior pronotal lobes metallic green, propleuron mostly cream. Posterior pronotal lobe short, collar-like centrally but with rear lateral extremities produced as outward directed, finger-like projections. Synthorax (
<figureCitation id="53EF2A27FFED324CFB481EFDFAD2FB3C" box="[1203,1287,1233,1259]" captionStart="FIGURES 1720" captionStartId="6.[151,264,1478,1503]" captionTargetBox="[151,1437,453,1453]" captionTargetId="figure@6.[151,1437,453,1453]" captionTargetPageId="6" captionText="FIGURES 1720. Lateral view of synthorax of Amphicnemis spp. (17) Amphicnemis rigiketit sp. holotype male: (19) Amphicnemis rigiketit sp. nov., female described; (18) male Amphicnemis remiger (SAR11_12_COE332); (20) female Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558211" httpUri="https://zenodo.org/record/3558211/files/figure.png" pageId="1" pageNumber="372">Fig. 17</figureCitation>
): mesepisternum, most of mesepimeron, upper part of mesokatepisternum and small area of metepisternum adjacent to antealar carina metallic green, with small pale area in upper part mesepimeron immediately adjacent to antealar carina, remainder of synthorax cream. Legs (left anterior leg absent below trochanter) cream except dark brown to black spines, small dark mark lowest part femur and immediately adjacent part flexor surface tibia, black rings between tarsal segments, tips of claws black. Wings: 12 Px in Fw, 11 (left) or 12 (right) in Hw. Arculus slightly distal to Ax2. R
<subScript id="575034E7FFED324CFF511F96FF67FA1F" attach="left" box="[170,178,1466,1480]" fontSize="6" pageId="1" pageNumber="372">4</subScript>
arising well distal to subnodus in all wings, IR
<subScript id="575034E7FFED324CFD3A1F96FD1CFA1F" attach="left" box="[705,713,1466,1480]" fontSize="6" pageId="1" pageNumber="372">3</subScript>
joined to it by a short stalk. Pterostigma almost trapezoidal with costal side shorter than anal side, dark brownish grey with whitish border, covering ca one underlying cell.
</paragraph>
<paragraph id="CB6B36A2FFED324CFF3C1FDDFCFCF86B" blockId="1.[151,1437,404,2015]" pageId="1" pageNumber="372">
<emphasis id="F9A0EAB0FFED324CFF3C1FDDFEE4F9DC" box="[199,305,1521,1547]" italics="true" pageId="1" pageNumber="372">Abdomen</emphasis>
. S1 cream with apical transverse dark mark. S2 metallic green above, cream below. S36 largely very dark brown above, cream lower laterally. S7 similar but dark markings extending lower laterally in apical ca onethird. S8 bronzy black above, brownish below. S9 almost entirely bronzy black, S10 dark brown and black above, mostly brownish white laterally, pale brown to rear. Anal appendages (
<figureCitation id="53EF2A27FFED324CFC4B1C71FB8EF9A0" box="[944,1115,1628,1655]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="1" pageNumber="372">Figs 23, 25, 27</figureCitation>
) white. Cerci in lateral view (
<figureCitation id="53EF2A27FFED324CFF5B1CADFF21F94C" box="[160,244,1664,1691]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="1" pageNumber="372">Fig. 25</figureCitation>
) with upper branch narrow at base, directed gently upwards, narrowing further to ca two-thirds length where abruptly expanded and turned down at slightly more than a right angle, from downturn upper margin running straight then slightly out to rounded apex, lower margin curved. Lower branch tapering from base, directed gently down, abruptly expanding along upper margin at apex with black dorsal interior tooth just before apex. In dorsal view (
<figureCitation id="53EF2A27FFED324CFF271D3DFEF8F8FC" box="[220,301,1809,1835]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="1" pageNumber="372">Fig. 23</figureCitation>
) upper branch of cercus of approximately even width from base to shortly before ca half-length, lacking any well-defined projection although with a slight swelling just before contraction, expanded apical part appearing as a rounded shelf below an outer ridge running to apex. In ventral view (
<figureCitation id="53EF2A27FFED324CFBFF1D75FB82F8A4" box="[1028,1111,1881,1907]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="1" pageNumber="372">Fig. 27</figureCitation>
) lower branch tapering from base, slightly expanded at apex, where very weakly bifurcated. Paraprocts short, pale and brown with short finger like projection directed to rear in dorsal part (lateral view).
</paragraph>
<paragraph id="CB6B36A2FFED324CFF3C1DE9FBBCF808" blockId="1.[151,1437,404,2015]" box="[199,1129,1988,2015]" pageId="1" pageNumber="372">Measurements (mm). Abdomen without anal appendages 33.5, cercus ca 1, Hw 21.</paragraph>
<caption id="9FAB662AFFEE324FFF6C1E9BFBD4FB23" ID-DOI="http://doi.org/10.5281/zenodo.3558203" ID-Zenodo-Dep="3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="2" pageNumber="373" startId="2.[151,264,1207,1232]" targetBox="[151,1437,179,1179]" targetPageId="2">
<paragraph id="CB6B36A2FFEE324FFF6C1E9BFBD4FB23" blockId="2.[151,1436,1207,1268]" pageId="2" pageNumber="373">
<emphasis id="F9A0EAB0FFEE324FFF6C1E9BFEEEFB07" bold="true" box="[151,315,1207,1232]" pageId="2" pageNumber="373">FIGURES 14.</emphasis>
<taxonomicName id="0CD44D21FFEE324FFEB91E9BFDF4FB18" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[322,545,1207,1231]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFEE324FFEB91E9BFDF4FB18" box="[322,545,1207,1231]" italics="true" pageId="2" pageNumber="373">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFEE324FFDD31E94FDACFB18" box="[552,633,1208,1231]" pageId="2" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
, head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male
<taxonomicName id="0CD44D21FFEE324FFF6C1EF7FEA0FB24" authorityName="Laidlaw" authorityYear="1912" box="[151,373,1243,1267]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="373" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEE324FFF6C1EF7FEA0FB24" box="[151,373,1243,1267]" italics="true" pageId="2" pageNumber="373">Amphicnemis remiger</emphasis>
</taxonomicName>
(SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view.
</paragraph>
</caption>
<paragraph id="CB6B36A2FFEE324FFF3C1F0EFEB7FA53" blockId="2.[151,1437,1314,2024]" pageId="2" pageNumber="373">
<emphasis id="F9A0EAB0FFEE324FFF3C1F0EFECBFAEB" bold="true" box="[199,286,1314,1340]" pageId="2" pageNumber="373">Female</emphasis>
(based on SAR18_COE64). Overall similar to male except in the thorax and as noted below.
<emphasis id="F9A0EAB0FFEE324FFADB1F0EFA88FAEB" box="[1312,1373,1314,1340]" italics="true" pageId="2" pageNumber="373">Head</emphasis>
(
<figureCitation id="53EF2A27FFEE324FFA911F0EFF16FAB7" captionStart="FIGURES 58" captionStartId="3.[151,264,1370,1395]" captionTargetBox="[151,1437,345,1345]" captionTargetId="figure@3.[151,1437,345,1345]" captionTargetPageId="3" captionText="FIGURES 58. Amphicnemis rigiketit sp. nov., head of female described: (5), dorsal-frontal view; (7) dorsal view. Head of female Amphicnemis remiger (BRU13_COE103); (6) dorsal-frontal view; (8) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558205" httpUri="https://zenodo.org/record/3558205/files/figure.png" pageId="2" pageNumber="373">Figs 5, 7</figureCitation>
). Only basal ca one-third of labrum black. Dark patches on mandible bases adjacent to base of labrum. Median ocellus yellowish.
</paragraph>
<paragraph id="CB6B36A2FFEE324FFF3C1FA2FC52F973" blockId="2.[151,1437,1314,2024]" pageId="2" pageNumber="373">
<emphasis id="F9A0EAB0FFEE324FFF3C1FA2FECDFA7F" box="[199,280,1422,1448]" italics="true" pageId="2" pageNumber="373">Thorax</emphasis>
. Prothorax (
<figureCitation id="53EF2A27FFEE324FFE501FA2FDE6FA7F" box="[427,563,1421,1448]" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="2" pageNumber="373">Figs 13, 15</figureCitation>
) with posterior pronotal lobe collar-like centrally in dorsal view (
<figureCitation id="53EF2A27FFEE324FFAC31FA2FA5BFA7F" box="[1336,1422,1422,1448]" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="2" pageNumber="373">Fig. 13</figureCitation>
), slightly folded upwards laterally then extended downwards to rear as rounded flap, free margin arching upwards back to base (
<figureCitation id="53EF2A27FFEE324FFED51FFAFEAEFA27" box="[302,379,1493,1520]" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="2" pageNumber="373">Fig 15</figureCitation>
). Almost entire prothorax bluish grey with indistinct darker and lighter areas, lateral parts anterior carina of anterior pronotal lobe black, centrally pale. Postepimeral strip slightly expanded just below posterior pronotal lobe (this is difficult to see because it is in the shadow of the lateral parts of the posterior pronotal lobe, indicated with a white line in
<figureCitation id="53EF2A27FFEE324FFE171C6EFD95F98B" box="[492,576,1601,1628]" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="2" pageNumber="373">Fig. 15</figureCitation>
). Synthorax (
<figureCitation id="53EF2A27FFEE324FFD221C6EFCF9F98B" box="[729,812,1602,1628]" captionStart="FIGURES 1720" captionStartId="6.[151,264,1478,1503]" captionTargetBox="[151,1437,453,1453]" captionTargetId="figure@6.[151,1437,453,1453]" captionTargetPageId="6" captionText="FIGURES 1720. Lateral view of synthorax of Amphicnemis spp. (17) Amphicnemis rigiketit sp. holotype male: (19) Amphicnemis rigiketit sp. nov., female described; (18) male Amphicnemis remiger (SAR11_12_COE332); (20) female Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558211" httpUri="https://zenodo.org/record/3558211/files/figure.png" pageId="2" pageNumber="373">Fig. 19</figureCitation>
) dorsum bluish with a greenish cast, remainder bluish, become paler lower laterally, venter pale. Legs with broad black stripe on extensor surface of femur. Wings with arculus only very slightly distal to Ax 2, 14 Px in Fw, 13 Px in Hw.
</paragraph>
<paragraph id="CB6B36A2FFEE324FFF3C1C82FEF8F8C7" blockId="2.[151,1437,1314,2024]" pageId="2" pageNumber="373">
<emphasis id="F9A0EAB0FFEE324FFF3C1C82FEE4F91F" box="[199,305,1710,1736]" italics="true" pageId="2" pageNumber="373">Abdomen</emphasis>
(
<figureCitation id="53EF2A27FFEE324FFEBA1C82FE4EF91F" box="[321,411,1710,1736]" captionStart="FIGURES 2122" captionStartId="7.[151,264,1009,1034]" captionTargetBox="[158,1430,489,984]" captionTargetId="figure@7.[158,1430,489,984]" captionTargetPageId="7" captionText="FIGURES 2122. Lateral view of pattern of terminal abdominal segments of female Amphicnemis spp.: (21), Amphicnemis rigiketit sp. nov., female described; (22) Amphicnemis remiger (BRU13_COE103)." figureDoi="http://doi.org/10.5281/zenodo.3558213" httpUri="https://zenodo.org/record/3558213/files/figure.png" pageId="2" pageNumber="373">Figs 21</figureCitation>
). S9 blackish with poorly defined pale lateral mark. S10 black below, whitish blue above. Cerci whitish, shorter than S10. Ovipositor pale, not extending further than tips of cerci except for mostly dark brown styles.
</paragraph>
<paragraph id="CB6B36A2FFEE324FFF3C1D36FB5EF8E3" blockId="2.[151,1437,1314,2024]" box="[199,1163,1817,1844]" pageId="2" pageNumber="373">Measurements (mm). Abdomen without anal appendages or ovipositor 37.5, Hw 23.5.</paragraph>
<paragraph id="CB6B36A2FFEE324EFF3C1D11FB3BFE94" blockId="2.[151,1437,1314,2024]" lastBlockId="3.[151,1436,153,323]" lastPageId="3" lastPageNumber="374" pageId="2" pageNumber="373">
<emphasis id="F9A0EAB0FFEE324FFF3C1D11FDE5F88F" bold="true" box="[199,560,1853,1880]" pageId="2" pageNumber="373">
Variation in
<typeStatus id="146F8800FFEE324FFEA61D12FE00F88F" box="[349,469,1854,1880]" pageId="2" pageNumber="373" type="paratype">paratypes</typeStatus>
. Males.
</emphasis>
There is little variation in markings except for small differences in the pattern of parts of the front of the head, none significant. There is some variation in the shape of the pterostigma with the costal side less short relative to the anal side in some, and R
<subScript id="575034E7FFEE324FFCDC1DBBFCFAF872" attach="left" box="[807,815,1943,1957]" fontSize="6" pageId="2" pageNumber="373">4</subScript>
is less distal to the subnodus in some. Many of the
<typeStatus id="146F8800FFEE324FFA981DAAFF12F813" pageId="2" pageNumber="373" type="paratype">paratype</typeStatus>
males are semi-teneral with distorted anal appendages.
<emphasis id="F9A0EAB0FFEE324FFCBC1D86FC65F813" bold="true" box="[839,944,1962,1988]" pageId="2" pageNumber="373">Females.</emphasis>
Two of the female
<typeStatus id="146F8800FFEE324FFB691D86FB28F813" box="[1170,1277,1962,1988]" pageId="2" pageNumber="373" type="paratype">paratypes</typeStatus>
are immature with the synthorax pale red to below the interpleural suture and most of the prothorax red. The legs are also pale red and lack the black stripe on the extensor surface of the femur, the brown marks on the abdomen are paler and the pale mark on S9 faint. On one of the mature females the posterior femurs are almost white on the flexor surface, in one of the immature females the femurs are white on middle and posterior legs, and on the other immature female the femurs are partially white on the posterior legs only. One of the mature females has the dark stripes on the extensor surface of the femur reduced. Other variation is minor and similar to that in the male
<typeStatus id="146F8800FFEF324EFB801B05FB3FFE94" box="[1147,1258,297,323]" pageId="3" pageNumber="374" type="paratype">paratypes</typeStatus>
.
</paragraph>
<caption id="9FAB662AFFEF324EFF6C1F76FBFFFA40" ID-DOI="http://doi.org/10.5281/zenodo.3558205" ID-Zenodo-Dep="3558205" httpUri="https://zenodo.org/record/3558205/files/figure.png" pageId="3" pageNumber="374" startId="3.[151,264,1370,1395]" targetBox="[151,1437,345,1345]" targetPageId="3">
<paragraph id="CB6B36A2FFEF324EFF6C1F76FBFFFA40" blockId="3.[151,1436,1370,1431]" pageId="3" pageNumber="374">
<emphasis id="F9A0EAB0FFEF324EFF6C1F76FEE8FAA4" bold="true" box="[151,317,1370,1395]" pageId="3" pageNumber="374">FIGURES 58.</emphasis>
<taxonomicName id="0CD44D21FFEF324EFEBE1F77FDF3FAA4" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[325,550,1371,1395]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFEF324EFEBE1F77FDF3FAA4" box="[325,550,1371,1395]" italics="true" pageId="3" pageNumber="374">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFEF324EFDD51F77FD54FAA5" box="[558,641,1371,1394]" pageId="3" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
, head of female described: (5), dorsal-frontal view; (7) dorsal view. Head of female
<taxonomicName id="0CD44D21FFEF324EFF191F53FE15FA40" authorityName="Laidlaw" authorityYear="1912" box="[226,448,1407,1431]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEF324EFF191F53FE15FA40" box="[226,448,1407,1431]" italics="true" pageId="3" pageNumber="374">Amphicnemis remiger</emphasis>
</taxonomicName>
(BRU13_COE103); (6) dorsal-frontal view; (8) dorsal view.
</paragraph>
</caption>
<paragraph id="CB6B36A2FFEF324EFF3C1FEAFEDAF9F0" blockId="3.[151,1436,1476,2044]" pageId="3" pageNumber="374">
Measurements (mm). Males: 1214 Px in Fw,
<quantity id="0C2C9B47FFEF324EFD1F1FE9FC9CFA08" box="[740,841,1477,1503]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="3" pageNumber="374" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">1112 in</quantity>
Hw, Abdomen without anal appendages 33.536.5, Hw 2021. Female: 1214 Px in Fw,
<quantity id="0C2C9B47FFEF324EFDC21FC5FD4EF9D4" box="[569,667,1513,1539]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="3" pageNumber="374" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">1112 in</quantity>
Hw, abdomen without anal appendages or ovipositor 36.538.5, Hw 22.523.5.
</paragraph>
</subSubSection>
<subSubSection id="83CE6529FFEF3249FF3C1C1CFC89FE5C" lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374" type="diagnosis">
<paragraph id="CB6B36A2FFEF3249FF3C1C1CFC89FE5C" blockId="3.[151,1436,1476,2044]" lastBlockId="4.[151,1436,152,395]" lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374">
<emphasis id="F9A0EAB0FFEF324EFF3C1C1CFE97F99C" bold="true" box="[199,322,1584,1611]" pageId="3" pageNumber="374">Diagnosis.</emphasis>
The male is easily separated from its known, named, congeners in Borneo except
<taxonomicName id="0CD44D21FFEF324EFB1D1C1EFEDBF9B8" authority="Lieftinck, 1940" authorityName="Lieftinck" authorityYear="1940" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="F9A0EAB0FFEF324EFB1D1C1EFA81F99C" box="[1254,1364,1586,1612]" italics="true" pageId="3" pageNumber="374">A. mariae</emphasis>
<bibRefCitation id="AF454B53FFEF324EFAA71C1DFEDBF9B8" author="Lieftinck, M. A." pageId="3" pageNumber="374" pagination="337 - 390" refId="ref6703" refString="Lieftinck, M. A. (1940) Descriptions and records of South-East Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article" year="1940">Lieftinck, 1940</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="0CD44D21FFEF324EFEBC1C7AFE6AF9B8" authorityName="Laidlaw" authorityYear="1912" box="[327,447,1622,1648]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEF324EFEBC1C7AFE6AF9B8" box="[327,447,1622,1648]" italics="true" pageId="3" pageNumber="374">A. remiger</emphasis>
</taxonomicName>
, by the combination of posterior pronotal lobe lacking a long central horn, pterostigma the same colour in both pairs of wings and not bicoloured, except for a narrow pale border, and the upper branch of the cerci lacking any well-developed projections. Males of
<taxonomicName id="0CD44D21FFEF324EFCD01CB2FC4CF960" authorityName="Lieftinck" authorityYear="1940" box="[811,921,1694,1720]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="F9A0EAB0FFEF324EFCD01CB2FC4CF960" box="[811,921,1694,1720]" italics="true" pageId="3" pageNumber="374">A. mariae</emphasis>
</taxonomicName>
and
<taxonomicName id="0CD44D21FFEF324EFC341CB2FB93F960" authorityName="Laidlaw" authorityYear="1912" box="[975,1094,1694,1720]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEF324EFC341CB2FB93F960" box="[975,1094,1694,1720]" italics="true" pageId="3" pageNumber="374">A. remiger</emphasis>
</taxonomicName>
have a better defined subbasal projection on the cerci (visible in dorsal view, see
<figureCitation id="53EF2A27FFEF324EFD3D1CEDFCC0F90C" box="[710,789,1729,1755]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="3" pageNumber="374">Fig. 24</figureCitation>
here and Plate 15,
<figureCitation id="53EF2A27FFEF324EFC111CEDFBFEF90C" box="[1002,1067,1729,1755]" captionStart="FIGURES 14" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 14. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="3" pageNumber="374">
Fig.
<quantity id="0C2C9B47FFEF324EFBE61CEDFBFEF90C" box="[1053,1067,1729,1755]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="374" unit="in" value="1.0">1</quantity>
</figureCitation>
in
<bibRefCitation id="AF454B53FFEF324EFBB51CEDFADAF90C" author="Lieftinck, M. A." box="[1102,1295,1729,1755]" pageId="3" pageNumber="374" pagination="337 - 390" refId="ref6703" refString="Lieftinck, M. A. (1940) Descriptions and records of South-East Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article" year="1940">Lieftinck (1940))</bibRefCitation>
and lack the abrupt downturn present in the cerci of
<taxonomicName id="0CD44D21FFEF324EFDB61CCAFD10F928" authorityName="Dow" authorityYear="2019" box="[589,709,1765,1792]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFEF324EFDB61CCAFD10F928" box="[589,709,1765,1792]" italics="true" pageId="3" pageNumber="374">A. rigiketit</emphasis>
</taxonomicName>
; the upper branch of the cercus in
<taxonomicName id="0CD44D21FFEF324EFBBF1CCAFB67F928" authorityName="Lieftinck" authorityYear="1940" box="[1092,1202,1766,1792]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="F9A0EAB0FFEF324EFBBF1CCAFB67F928" box="[1092,1202,1766,1792]" italics="true" pageId="3" pageNumber="374">A. mariae</emphasis>
</taxonomicName>
forms a smooth arch (Plate 15,
<figureCitation id="53EF2A27FFEF324EFEF01D25FE9BF8F4" box="[267,334,1801,1827]" captionStart="FIGURES 14" captionStartId="2.[151,264,1207,1232]" captionTargetBox="[151,1437,179,1179]" captionTargetId="figure@2.[151,1437,179,1179]" captionTargetPageId="2" captionText="FIGURES 14. Amphicnemis rigiketit sp. nov., head of holotype male: (1), dorsal-frontal view; (3) dorsal view. Head of male Amphicnemis remiger (SAR11_12_COE332); (2) dorsal-frontal view; (4) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3558203" httpUri="https://zenodo.org/record/3558203/files/figure.png" pageId="3" pageNumber="374">
Fig.
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</figureCitation>
in
<bibRefCitation id="AF454B53FFEF324EFE881D25FDECF8F4" author="Lieftinck, M. A." box="[371,569,1801,1827]" pageId="3" pageNumber="374" pagination="337 - 390" refId="ref6703" refString="Lieftinck, M. A. (1940) Descriptions and records of South-East Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article" year="1940">Lieftinck (1940))</bibRefCitation>
. The shape of the apical part of the lower branch of the cercus is different in
<taxonomicName id="0CD44D21FFEF324EFF6C1D02FEDAF890" authorityName="Dow" authorityYear="2019" box="[151,271,1837,1864]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFEF324EFF6C1D02FEDAF890" box="[151,271,1837,1864]" italics="true" pageId="3" pageNumber="374">A. rigiketit</emphasis>
</taxonomicName>
and
<taxonomicName id="0CD44D21FFEF324EFEBE1D02FE6EF890" authorityName="Laidlaw" authorityYear="1912" box="[325,443,1838,1864]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEF324EFEBE1D02FE6EF890" box="[325,443,1838,1864]" italics="true" pageId="3" pageNumber="374">A. remiger</emphasis>
</taxonomicName>
(compare
<figureCitation id="53EF2A27FFEF324EFDC91D01FD7AF890" box="[562,687,1836,1863]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="3" pageNumber="374">Figs 25, 27</figureCitation>
with
<figureCitation id="53EF2A27FFEF324EFD151D01FCBAF890" box="[750,879,1837,1863]" captionStart="FIGURES 2328" captionStartId="8.[151,264,1670,1695]" captionTargetBox="[151,1436,181,1644]" captionTargetId="figure@8.[151,1436,181,1644]" captionTargetPageId="8" captionText="FIGURES 2328. Amphicnemis rigiketit sp. nov., anal appendages of of holotype male: (23), dorsal view; (25) lateral view; (27) ventral view. Anal appendages of male Amphicnemis remiger (SAR11_12_COE332); (24) dorsal view; (26) lateral view; (28) ventral view. All scale bars 5mm." figureDoi="http://doi.org/10.5281/zenodo.3558215" httpUri="https://zenodo.org/record/3558215/files/figure.png" pageId="3" pageNumber="374">Figs 26, 28</figureCitation>
). The male of
<taxonomicName id="0CD44D21FFEF324EFBF41D02FB53F890" authorityName="Laidlaw" authorityYear="1912" box="[1039,1158,1838,1864]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEF324EFBF41D02FB53F890" box="[1039,1158,1838,1864]" italics="true" pageId="3" pageNumber="374">A. remiger</emphasis>
</taxonomicName>
has the anterior pronotal lobe almost entirely dark coloured but it is mostly pale in
<taxonomicName id="0CD44D21FFEF324EFCE61D7EFC40F8BC" authorityName="Dow" authorityYear="2019" box="[797,917,1873,1900]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFEF324EFCE61D7EFC40F8BC" box="[797,917,1873,1900]" italics="true" pageId="3" pageNumber="374">A. rigiketit</emphasis>
</taxonomicName>
(and
<taxonomicName id="0CD44D21FFEF324EFC2F1D7EFB97F8BC" authorityName="Lieftinck" authorityYear="1940" box="[980,1090,1874,1900]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="F9A0EAB0FFEF324EFC2F1D7EFB97F8BC" box="[980,1090,1874,1900]" italics="true" pageId="3" pageNumber="374">A. mariae</emphasis>
</taxonomicName>
). The lateral extremities of the posterior pronotal lobe are produced as finger-like projections in
<taxonomicName id="0CD44D21FFEF324EFC881D5AFC3EF858" authorityName="Dow" authorityYear="2019" box="[883,1003,1909,1936]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFEF324EFC881D5AFC3EF858" box="[883,1003,1909,1936]" italics="true" pageId="3" pageNumber="374">A. rigiketit</emphasis>
</taxonomicName>
but these are lacking in
<taxonomicName id="0CD44D21FFEF324EFB051D5AFAB9F858" authorityName="Lieftinck" authorityYear="1940" box="[1278,1388,1910,1936]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="F9A0EAB0FFEF324EFB051D5AFAB9F858" box="[1278,1388,1910,1936]" italics="true" pageId="3" pageNumber="374">A. mariae</emphasis>
</taxonomicName>
and
<taxonomicName id="0CD44D21FFEF324EFF6C1DB6FEC5F864" authorityName="Laidlaw" authorityYear="1912" box="[151,272,1946,1972]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEF324EFF6C1DB6FEC5F864" box="[151,272,1946,1972]" italics="true" pageId="3" pageNumber="374">A. remiger</emphasis>
</taxonomicName>
. The female can be separated from those of most species by the combination of the same pterostigma characteristics as the male and posterior prothoracic lobe lacking a strong central horn and projected to rear laterally. It can be separated from that of
<taxonomicName id="0CD44D21FFEF324EFDD71DCEFD77F82C" authorityName="Laidlaw" authorityYear="1912" box="[556,674,2018,2044]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="374" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFEF324EFDD71DCEFD77F82C" box="[556,674,2018,2044]" italics="true" pageId="3" pageNumber="374">A. remiger</emphasis>
</taxonomicName>
by the shape of the lateral parts of the posterior prothoracic lobe and the expansion of the postepimeral strip (compare
<figureCitation id="53EF2A27FFE83249FD3A1AB5FCBFFF63" box="[705,874,152,180]" captionStart="FIGURES 1316" captionStartId="5.[151,264,1532,1557]" captionTargetBox="[151,1436,272,1507]" captionTargetId="figure@5.[151,1436,272,1507]" captionTargetPageId="5" captionText="FIGURES 1316. Amphicnemis rigiketit sp. nov., prothorax of female described: (13), dorsal view; (15) lateral view. Prothorax of female Amphicnemis remiger (BRU13_COE103); (14) dorsal view; (16) lateral view. The expanded part of the postepimeral strip is indicated with a white line in Figs 15 and 16." figureDoi="http://doi.org/10.5281/zenodo.3558209" httpUri="https://zenodo.org/record/3558209/files/figure.png" pageId="4" pageNumber="375">Figs 15 and 16</figureCitation>
). The female of
<taxonomicName id="0CD44D21FFE83249FBD91AB5FB45FF64" authorityName="Lieftinck" authorityYear="1940" box="[1058,1168,153,179]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="F9A0EAB0FFE83249FBD91AB5FB45FF64" box="[1058,1168,153,179]" italics="true" pageId="4" pageNumber="375">A. mariae</emphasis>
</taxonomicName>
is unknown but may be similar. Using unpublished information supplied by myself,
<bibRefCitation id="AF454B53FFE83249FCBE1A91FB56FF00" author="Kosterin, O. E. &amp; Kompier, T." box="[837,1155,189,215]" pageId="4" pageNumber="375" pagination="281 - 294" refId="ref6442" refString="Kosterin, O. E. &amp; Kompier, T. (2018) Amphicnemis valentini sp. nov. from the Cardamom ecoregion in Cambodia and Vietnam (Odonata: Coenagrionidae). Zootaxa, 4429 (2), 281 - 294. https: // doi. org / 10.11646 / zootaxa. 4429.2.4" type="journal article" year="2018">Kosterin &amp; Kompier (2018)</bibRefCitation>
noted that the female of
<taxonomicName id="0CD44D21FFE83249FF6C1ACDFE67FF2C" authority="Lieftinck, 1940" authorityName="Lieftinck" authorityYear="1940" box="[151,434,225,251]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="annae">
<emphasis id="F9A0EAB0FFE83249FF6C1ACDFF29FF2C" box="[151,252,225,251]" italics="true" pageId="4" pageNumber="375">A. annae</emphasis>
<bibRefCitation id="AF454B53FFE83249FEFF1ACDFE67FF2C" author="Lieftinck, M. A." box="[260,434,225,251]" pageId="4" pageNumber="375" pagination="337 - 390" refId="ref6703" refString="Lieftinck, M. A. (1940) Descriptions and records of South-East Asiatic Odonata (II). Treubia, 17, 337 - 390." type="journal article" year="1940">Lieftinck, 1940</bibRefCitation>
</taxonomicName>
is apparently polymorphic in the structure of the posterior prothoracic lobe, with a long central horn present in some individuals in a small number of populations but not in most. The form without the central horn is the only one recorded in
<collectingCountry id="B3C37632FFE83249FDA51B05FD6BFE94" box="[606,702,297,323]" name="Malaysia" pageId="4" pageNumber="375">Sarawak</collectingCountry>
but this hornless form is easily distinguished from the female of
<taxonomicName id="0CD44D21FFE83249FF6C1B61FEDAFEB0" authorityName="Dow" authorityYear="2019" box="[151,271,333,359]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFE83249FF6C1B61FEDAFEB0" box="[151,271,333,359]" italics="true" pageId="4" pageNumber="375">A. rigiketit</emphasis>
</taxonomicName>
because in dorsal view the central part of the posterior pronotal lobe appears almost rectangular with the lateral extremities projecting more to the rear in the same view.
</paragraph>
</subSubSection>
<caption id="9FAB662AFFE83249FF6C1CA1FBD2F91D" ID-DOI="http://doi.org/10.5281/zenodo.3558207" ID-Zenodo-Dep="3558207" httpUri="https://zenodo.org/record/3558207/files/figure.png" pageId="4" pageNumber="375" startId="4.[151,264,1677,1702]" targetBox="[151,1436,417,1652]" targetPageId="4">
<paragraph id="CB6B36A2FFE83249FF6C1CA1FBD2F91D" blockId="4.[151,1436,1677,1738]" pageId="4" pageNumber="375">
<emphasis id="F9A0EAB0FFE83249FF6C1CA1FE97F971" bold="true" box="[151,322,1677,1702]" pageId="4" pageNumber="375">FIGURES 912</emphasis>
.
<taxonomicName id="0CD44D21FFE83249FEB41CA1FDFAF972" authority="Dow, 2019" authorityName="Dow" authorityYear="2019" box="[335,559,1677,1701]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="rigiketit" status="sp. nov.">
<emphasis id="F9A0EAB0FFE83249FEB41CA1FDFAF972" box="[335,559,1677,1701]" italics="true" pageId="4" pageNumber="375">Amphicnemis rigiketit</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E29357CBFFE83249FDCD1CA2FD5DF972" box="[566,648,1678,1701]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
, prothorax of holotype male: (9), dorsal view; (11) lateral view. Prothorax of male
<taxonomicName id="0CD44D21FFE83249FF341C9DFE78F91E" authorityName="Laidlaw" authorityYear="1912" box="[207,429,1713,1737]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE83249FF341C9DFE78F91E" box="[207,429,1713,1737]" italics="true" pageId="4" pageNumber="375">Amphicnemis remiger</emphasis>
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(SAR11_12_COE332); (10) dorsal view; (12) lateral view.
</paragraph>
</caption>
<subSubSection id="83CE6529FFE83248FF3C1CD4FEBBFF2C" lastPageId="5" lastPageNumber="376" pageId="4" pageNumber="375" type="discussion">
<paragraph id="CB6B36A2FFE83249FF3C1CD4FA9DF83D" blockId="4.[151,1437,1784,2026]" pageId="4" pageNumber="375">
<emphasis id="F9A0EAB0FFE83249FF3C1CD4FEEEF8C5" bold="true" box="[199,315,1784,1810]" pageId="4" pageNumber="375">Remarks.</emphasis>
Although none were taken in tandem, the female is associated with the male by reasonable supposition, males of no other species of
<taxonomicName id="0CD44D21FFE83249FDBA1D30FD0DF8E1" authorityName="Selys" authorityYear="1863" box="[577,728,1820,1846]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="F9A0EAB0FFE83249FDBA1D30FD0DF8E1" box="[577,728,1820,1846]" italics="true" pageId="4" pageNumber="375">Amphicnemis</emphasis>
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were found at the site. The male is clearly closely related to
<taxonomicName id="0CD44D21FFE83249FA7F1D30FF3AF88D" authorityName="Laidlaw" authorityYear="1912" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE83249FA7F1D30FF3AF88D" italics="true" pageId="4" pageNumber="375">A. remiger</emphasis>
</taxonomicName>
and the female is similar to that of
<taxonomicName id="0CD44D21FFE83249FD7C1D6CFCD5F88D" authorityName="Laidlaw" authorityYear="1912" box="[647,768,1856,1882]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE83249FD7C1D6CFCD5F88D" box="[647,768,1856,1882]" italics="true" pageId="4" pageNumber="375">A. remiger</emphasis>
</taxonomicName>
, further supporting the association of the sexes. It is likely that some characters of colour and pattern, not mentioned in the diagnosis, also consistently serve to distinguish
<taxonomicName id="0CD44D21FFE83249FA7F1D48FF3AF875" authorityName="Laidlaw" authorityYear="1912" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="remiger">
<emphasis id="F9A0EAB0FFE83249FA7F1D48FF3AF875" italics="true" pageId="4" pageNumber="375">A. remiger</emphasis>
</taxonomicName>
from
<taxonomicName id="0CD44D21FFE83249FECD1DA4FE65F875" authorityName="Dow" authorityYear="2019" box="[310,432,1928,1954]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFE83249FECD1DA4FE65F875" box="[310,432,1928,1954]" italics="true" pageId="4" pageNumber="375">A. rigiketit</emphasis>
</taxonomicName>
, for instance the extent of metallic colouration on the synthorax of males of the and the extent of pale colouration on abdominal segment
<quantity id="0C2C9B47FFE83249FD351D80FCDCF811" box="[718,777,1964,1990]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="375" unit="in" value="10.0">10 in</quantity>
females. However, the pattern of the anterior parts of the head is sufficiently variable in both species (and many other
<taxonomicName id="0CD44D21FFE83249FCBB1DFCFC02F83D" authorityName="Selys" authorityYear="1863" box="[832,983,2000,2026]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="F9A0EAB0FFE83249FCBB1DFCFC02F83D" box="[832,983,2000,2026]" italics="true" pageId="4" pageNumber="375">Amphicnemis</emphasis>
</taxonomicName>
) that it is of no diagnostic value.
</paragraph>
<paragraph id="CB6B36A2FFE93248FF3C1AB4FEBBFF2C" blockId="5.[151,1436,152,251]" pageId="5" pageNumber="376">
All specimens of
<taxonomicName id="0CD44D21FFE93248FE771AB5FDD1FF65" authorityName="Dow" authorityYear="2019" box="[396,516,152,179]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="rigiketit">
<emphasis id="F9A0EAB0FFE93248FE771AB5FDD1FF65" box="[396,516,152,179]" italics="true" pageId="5" pageNumber="376">A. rigiketit</emphasis>
</taxonomicName>
were collected either hanging over a steep sided, turbid stream or around pools or trailside, perched on foliage, in the adjacent forest, which becomes flooded at least occasionally during the wettest periods of the year.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,121 @@
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</paragraph>
</subSubSection>
<subSubSection id="FE31C3BAAEA2983115896E6A18D74ECC" pageId="0" pageNumber="980" type="materials_examined">
<paragraph id="EA30738F7E7E9325AE65360823419D55" pageId="0" pageNumber="980">Materials</paragraph>
<paragraph id="90F6A48D7ED4909F0170A46F9269C68D" pageId="0" pageNumber="980">
<materialsCitation id="8AF568015F50083C5E01319082AE1B6D" collectingDate="2011-07-14" collectorName="Kuntner, Gregoric, Candek" country="Switzerland" latitude="46.5166" location="Grison Alps, Alp Flix, Salategnas" longitude="9.6516" pageId="0" pageNumber="980" specimenCount="4" specimenCount-female="4">
Occurrence: recordedBy:
<collectorName id="92C27718BF8F701406E0F6166DC8CA7D" pageId="0" pageNumber="980">
Kuntner,
<normalizedToken id="6ED841D4BDA105679B09B2ADAA25A903" originalValue="Gregorič">Gregoric</normalizedToken>
,
<normalizedToken id="74CD7731E45BB0718A3A48C33CA35A91" originalValue="Čandek">Candek</normalizedToken>
</collectorName>
; sex:
<specimenCount id="0BD706D1E31F921057B81A3371B75318" type="female">4 females</specimenCount>
; Location: locationID: CH26; country:
<collectingCountry id="39BC55035AE7E4CFE01991D73070BBED" pageId="0" pageNumber="980">Switzerland</collectingCountry>
; locality:
<location id="6B26CCF91E2F1156BC423286A99D57EB" LSID="urn:lsid:plazi:treatment:437DB729FF4F3B9E8073D8BCDCAF5A60:6B26CCF91E2F1156BC423286A99D57EB" country="Switzerland" latitude="46.5166" longitude="9.6516" name="Grison Alps, Alp Flix, Salategnas" pageId="0" pageNumber="980">Grison Alps, Alp Flix, Salategnas</location>
; minimumElevationInMeters: 1987; maximumElevationInMeters: 1987; decimalLatitude:
<geoCoordinate id="E0699E2DAA66837F95FAB107CEFB3EE7" orientation="latitude" pageId="0" pageNumber="980" value="46.5166">46.5166</geoCoordinate>
; decimalLongitude:
<geoCoordinate id="1FD8F497AC72D1389D2F42C46CC33EB6" orientation="longitude" pageId="0" pageNumber="980" value="9.6516">9.6516</geoCoordinate>
; Event: eventDate:
<collectingDate id="EE8EFE15D38A794AEA1F207D5EDC6376" pageId="0" pageNumber="980" value="2011-07-14">2011-07-14</collectingDate>
; habitat: grassland
</materialsCitation>
<materialsCitation id="421CCA6FBED8002509D8C727BBF268D2" collectingDate="2011-07-16" collectorName="Kuntner, Gregoric, Candek" country="Switzerland" latitude="46.5203" location="Grison Alps, Alp Flix, Salategnas" longitude="9.6458" pageId="0" pageNumber="980" specimenCount="2" specimenCount-female="2">
Occurrence: recordedBy:
<collectorName id="37056E48E93C0AD7D5F8663C2D193BF2" pageId="0" pageNumber="980">
Kuntner,
<normalizedToken id="F9A921ED12268BE218D2FD84149A6E7C" originalValue="Gregorič">Gregoric</normalizedToken>
,
<normalizedToken id="B1ECD00529E1240CE9C1FA999DF115E8" originalValue="Čandek">Candek</normalizedToken>
</collectorName>
; sex:
<specimenCount id="08F7B6FD225017977EC3151D5A37114A" type="female">2 females</specimenCount>
; Location: locationID: CH32; country:
<collectingCountry id="4A99B5C3524B605C9EE3041FBA03604A" pageId="0" pageNumber="980">Switzerland</collectingCountry>
; locality:
<location id="573522F84400560906905F924D0562C1" LSID="urn:lsid:plazi:treatment:437DB729FF4F3B9E8073D8BCDCAF5A60:573522F84400560906905F924D0562C1" country="Switzerland" latitude="46.5203" longitude="9.6458" name="Grison Alps, Alp Flix, Salategnas" pageId="0" pageNumber="980">Grison Alps, Alp Flix, Salategnas</location>
; minimumElevationInMeters: 1955; maximumElevationInMeters: 1955; decimalLatitude:
<geoCoordinate id="3FDFE565F4E71EA1FF49578319F2DE4E" orientation="latitude" pageId="0" pageNumber="980" value="46.5203">46.5203</geoCoordinate>
; decimalLongitude:
<geoCoordinate id="6C6C7BBE0540561500F5237DEF075B84" orientation="longitude" pageId="0" pageNumber="980" value="9.6458">9.6458</geoCoordinate>
; Event: eventDate:
<collectingDate id="70115155A7174D12E4B495EE3AFE16F6" pageId="0" pageNumber="980" value="2011-07-16">2011-07-16</collectingDate>
; habitat: timberline forest, moss
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>