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<document ID-CLB-Dataset="297077" ID-DOI="10.5281/zenodo.7316535" ID-GBIF-Dataset="8712c624-6c5e-4918-b4e3-3f7d3a6b7ed5" ID-ISBN="0-8018-8221-4" checkinTime="1670905144989" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="D11815A16CAB54D2C8C5E7D6530D3C44" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Ctenodactylidae.1536-1537.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press" docTitle="Massoutiera Lataste 1885" docType="treatment" docVersion="4" id="4A4449CCB30C8CFA4278550CD5110DE5" lastPageNumber="1536" masterDocId="4A4449CCB30C8CFA4278550CD5110DE5" masterDocTitle="Order Rodentia - Family Ctenodactylidae" masterLastPageNumber="1537" masterPageNumber="1536" pageNumber="1536" updateTime="1716777739916" updateUser="ExternalLinkService">
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<mods:title id="CC6E1CEB666825F569521479E37E9E3E">Order Rodentia - Family Ctenodactylidae</mods:title>
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<mods:namePart id="DFEE87C3845D8A50A3D9964891BA71C7">Wilson, Don E.</mods:namePart>
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<mods:namePart id="3723AC79FE9228822301F39813E7A299">Reeder, DeeAnn</mods:namePart>
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<mods:title id="58B61F27568EE3AD6A8475E7189CD849">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2</mods:title>
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<taxonomicName id="C1477BE8F53F1748991267001E1AFCFF" ID-CoL="5LTY" authority="Lataste 1885" authorityName="Lataste" authorityYear="1885" class="Mammalia" family="Ctenodactylidae" genus="Massoutiera" kingdom="Animalia" order="Rodentia" pageNumber="1536" phylum="Chordata" rank="genus">
<emphasis id="99A559DD8B7391C916D1A74F8B0C8642" italics="true" pageNumber="1536">Massoutiera</emphasis>
Lataste 1885
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<treatmentCitation id="064B26DC9652E72160E848E0E0C573BF" author="Lataste" authority="Lataste 1885" authorityName="Lataste" authorityYear="1885" class="Mammalia" family="Ctenodactylidae" genus="Massoutiera" issue="3" journal="Le Naturaliste" kingdom="Animalia" order="Rodentia" page="21" pageNumber="1536" phylum="Chordata" rank="genus" title="Massoutiera" volume="7" year="1885">
<taxonomicName id="46A94CA7AD05D36034B7F1838CC2E1BD" ID-CoL="5LTY" authority="Lataste 1885" authorityName="Lataste" authorityYear="1885" class="Mammalia" family="Ctenodactylidae" genus="Massoutiera" kingdom="Animalia" order="Rodentia" pageNumber="1536" phylum="Chordata" rank="genus">
<emphasis id="1016CB2812BD6514E5D39A1210B0D69D" italics="true" pageNumber="1536">Massoutiera</emphasis>
Lataste 1885
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,
<bibCitation id="5D29D5F3BB948EF3A196B372008C7BE9" author="Lataste" issue="3" journal="Le Naturaliste" pageNumber="1536" pagination="21" title="Massoutiera" volume="7" year="1885">Le Naturaliste, 7 (3): 21</bibCitation>
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.
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<paragraph id="A68EB7DA4CD93F26ECBFEDFE8F8B0EC8" pageNumber="1536">
<emphasis id="4FB5B2619BC55237DD67478B5CF2E93A" inLineHeading="true" pageNumber="1536">Type Species:</emphasis>
<taxonomicName id="9A65FCA4E5ED7343FB387FCA6DEC159E" authority="Lataste 1881" authorityName="Lataste" authorityYear="1881" class="Mammalia" family="Ctenodactylidae" genus="Ctenodactylus" kingdom="Animalia" order="Rodentia" pageNumber="1536" phylum="Chordata" rank="species" species="mzabi">
<emphasis id="C78DD7679535090E7D9BF9B4B16341AE" italics="true" pageNumber="1536">Ctenodactylus mzabi</emphasis>
Lataste 1881
</taxonomicName>
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<emphasis id="4C170C0B34525A33471DE29676D0D568" inLineHeading="true" pageNumber="1536">Species and subspecies:</emphasis>
1 species:
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Species
<taxonomicName id="3A30B04CA4DF6820FC848C5B983B2E8B" authority="(Lataste 1881)" baseAuthorityName="Lataste" baseAuthorityYear="1881" class="Mammalia" family="Ctenodactylidae" genus="Massoutiera" kingdom="Animalia" order="Rodentia" pageNumber="1536" phylum="Chordata" rank="species" species="mzabi">
<emphasis id="C9C967D305185CB96CE08084F34677B8" italics="true" pageNumber="1536">Massoutiera mzabi</emphasis>
(Lataste 1881)
</taxonomicName>
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</subSubSection>
</treatment>
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<document id="0452F9C21351431B036041DDAE2933A4" ID-CLB-Dataset="46798" ID-DOI="10.11646/zootaxa.4078.1.22" ID-GBIF-Dataset="f71df488-ad28-4308-aed3-27540ae6ab4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="255899" ID-ZooBank="C45311A2-E7EA-448F-9F38-79E145E53674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460361558807" checkinUser="plazi" docAuthor="Grossi, Paschoal C., Santos, Mariana D. &amp; Almeida, Lúcia M." docDate="2016" docId="D11887F6C451FFB3C6D082D085E0F86E" docLanguage="en" docName="zootaxa.4078.1.22.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclocephala machadoi Grossi, Santos &amp; Almeida, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="246" masterDocId="2D21FF8EC450FFB2C64786128148FF9B" masterDocTitle="Two new species of Cyclocephala (Coleoptera: Scarabaeoidea: Melolonthidae) from Minas Gerais State, Brazil" masterLastPageNumber="251" masterPageNumber="245" pageNumber="246" updateTime="1698294628747" updateUser="ExternalLinkService">
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<mods:title id="39EBCFF36CC0B6F4DA8404DF2E289ECE">Two new species of Cyclocephala (Coleoptera: Scarabaeoidea: Melolonthidae) from Minas Gerais State, Brazil</mods:title>
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<treatment id="D11887F6C451FFB3C6D082D085E0F86E" ID-DOI="http://doi.org/10.5281/zenodo.5659768" ID-GBIF-Taxon="119409862" ID-Zenodo-Dep="5659768" LSID="urn:lsid:plazi:treatment:D11887F6C451FFB3C6D082D085E0F86E" httpUri="http://treatment.plazi.org/id/D11887F6C451FFB3C6D082D085E0F86E" lastPageNumber="246" pageId="1" pageNumber="246">
<subSubSection id="11AB656BC451FFB3C6D082D0800CFB65" pageId="1" pageNumber="246" type="nomenclature">
<paragraph id="590E36E0C451FFB3C6D082D08355FB46" blockId="1.[151,541,1218,1278]" box="[151,541,1218,1245]" pageId="1" pageNumber="246">
<heading id="0246818CC451FFB3C6D082D08355FB46" bold="true" box="[151,541,1218,1245]" fontSize="11" level="1" pageId="1" pageNumber="246" reason="1">
<emphasis id="6BC5EAF2C451FFB3C6D082D08355FB46" bold="true" box="[151,541,1218,1245]" pageId="1" pageNumber="246">
<taxonomicName id="9EB14D63C451FFB3C6D082D080F0FB47" box="[151,440,1218,1244]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C451FFB3C6D082D080F0FB47" bold="true" box="[151,440,1218,1244]" italics="true" pageId="1" pageNumber="246">Cyclocephala machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C451FFB3C7F882D18355FB46" box="[447,541,1219,1245]" pageId="1" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="590E36E0C451FFB3C6D082F7800CFB65" blockId="1.[151,541,1218,1278]" box="[151,324,1253,1278]" pageId="1" pageNumber="246">
(
<figureCitation id="C18A2A65C451FFB3C6D982F7804CFB65" box="[158,260,1253,1278]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Figures 1</figureCitation>
AF)
</paragraph>
</subSubSection>
<subSubSection id="11AB656BC451FFB3C6D0833F818EF837" pageId="1" pageNumber="246" type="materials_examined">
<paragraph id="590E36E0C451FFB3C6D0833F85AEFAF7" blockId="1.[151,1437,1325,2037]" pageId="1" pageNumber="246">
<emphasis id="6BC5EAF2C451FFB3C6D0833F80C9FADD" bold="true" box="[151,385,1325,1350]" pageId="1" pageNumber="246">Material examined.</emphasis>
<typeStatus id="860A8842C451FFB3C7CD833F80BAFADD" box="[394,498,1325,1350]" pageId="1" pageNumber="246" type="holotype">Holotype</typeStatus>
male labeled: “
<collectingCountry id="21A67670C451FFB3C4E3833F83A3FADD" box="[676,747,1325,1350]" name="Brazil" pageId="1" pageNumber="246">Brasil</collectingCountry>
, MG, São Gonçalo/do Rio Preto, P.E. do/Rio Preto, Porteira, Posto (4)/
<date id="2D0F1020C451FFB3C742834680CDFAF6" box="[261,389,1364,1389]" pageId="1" pageNumber="246" value="2011-12-02">
02.xii.
<quantity id="9E499B05C451FFB3C70D834680CDFAF6" box="[330,389,1364,1389]" metricMagnitude="6" metricUnit="m" metricValue="2.011847" pageId="1" pageNumber="246" unit="m" value="2011847.0">2011</quantity>
</date>
, 847m, black/ligth,
<geoCoordinate id="3C855027C451FFB3C43483468382FAF7" box="[627,714,1360,1388]" direction="south" orientation="latitude" pageId="1" pageNumber="246" precision="555" value="-18.09">18.09o S</geoCoordinate>
,
<geoCoordinate id="3C855027C451FFB3C4A28346820EFAF6" box="[741,838,1360,1389]" direction="west" orientation="longitude" pageId="1" pageNumber="246" precision="555" value="-43.34">43.34o W</geoCoordinate>
/Oliveira &amp; Ferreira legs.” (
<collectionCode id="3FA0AE25C451FFB3C23B83468592FAF7" box="[1148,1242,1364,1388]" pageId="1" pageNumber="246">CERPE</collectionCode>
).
</paragraph>
<paragraph id="590E36E0C451FFB3C680836A818EF837" blockId="1.[151,1437,1325,2037]" pageId="1" pageNumber="246">
<emphasis id="6BC5EAF2C451FFB3C680836A8033FA0A" bold="true" box="[199,379,1400,1425]" pageId="1" pageNumber="246">
<typeStatus id="860A8842C451FFB3C680836A807CFA0A" box="[199,308,1400,1425]" pageId="1" pageNumber="246" type="holotype">Holotype</typeStatus>
male
</emphasis>
.
<emphasis id="6BC5EAF2C451FFB3C7D7836A836EFA0A" box="[400,550,1400,1425]" italics="true" pageId="1" pageNumber="246">Total length:</emphasis>
9.0 mm.
<emphasis id="6BC5EAF2C451FFB3C4E3836A82D7FA0A" box="[676,927,1400,1425]" italics="true" pageId="1" pageNumber="246">Total pronotal width:</emphasis>
<quantity id="9E499B05C451FFB3C5EA836A8553FA0B" box="[941,1051,1400,1425]" metricMagnitude="-3" metricUnit="m" metricValue="3.61" pageId="1" pageNumber="246" unit="mm" value="3.61">3.61 mm</quantity>
.
<emphasis id="6BC5EAF2C451FFB3C26B836A8527FA0A" box="[1068,1135,1400,1425]" italics="true" pageId="1" pageNumber="246">Body:</emphasis>
Dorsal surface glabrous, ventrally moderately setose, setae distinctly elongate. Color almost completely testaceous, but head black except clypeus, antennae light pale whitish yellow, and pronotum and legs slightly darker (
<figureCitation id="C18A2A65C451FFB3C27883D28536FA43" box="[1087,1150,1472,1497]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
A).
<emphasis id="6BC5EAF2C451FFB3C2EB83D285BAFA42" box="[1196,1266,1472,1497]" italics="true" pageId="1" pageNumber="246">Head:</emphasis>
Dorsal surface finely punctate, more punctate on frons; clypeus almost impunctate, sides parallel at base, then weakly convergent to trapezoidal, reflexed apex (
<figureCitation id="C18A2A65C451FFB3C7BA801A830EF9BB" box="[509,582,1544,1569]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Figs 1</figureCitation>
AB). Frontoclypeal suture obsolete. Antennae with lamellae extremely developed, moderately and densely punctate (
<figureCitation id="C18A2A65C451FFB3C4E7803E83A9F9DF" box="[672,737,1580,1605]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
C); first lamella externally with longitudinal row of scattered setae.
<emphasis id="6BC5EAF2C451FFB3C69980428021F9F2" box="[222,361,1616,1641]" italics="true" pageId="1" pageNumber="246">Mouthparts:</emphasis>
Mandibles slender, apices upturned and weakly curved externally; last maxillary palpomere as long as preceding 2 together.
<emphasis id="6BC5EAF2C451FFB3C7B780678320F917" box="[496,616,1653,1676]" italics="true" pageId="1" pageNumber="246">Pronotum:</emphasis>
Surface moderately and uniformly punctate, smooth along center line, posterior margin with a weak bead at middle. Scutellum totally punctate, punctures moderately large.
<emphasis id="6BC5EAF2C451FFB3C30B808A84D4F92A" box="[1356,1436,1688,1713]" italics="true" pageId="1" pageNumber="246">Elytra:</emphasis>
Surface distinctly punctate; striae well defined but weakly marked, punctures fine.
<emphasis id="6BC5EAF2C451FFB3C20380AF85CBF94F" box="[1092,1155,1725,1748]" italics="true" pageId="1" pageNumber="246">Legs:</emphasis>
Protibiae tridentate with basal tooth nearly obsolete (
<figureCitation id="C18A2A65C451FFB3C79580F2835DF963" box="[466,533,1760,1785]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Figs 1</figureCitation>
AB); inner claw widely bifid, forming a right angle (
<figureCitation id="C18A2A65C451FFB3C23080F285FEF963" box="[1143,1206,1760,1785]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
D).
<emphasis id="6BC5EAF2C451FFB3C2A280F3847AF963" box="[1253,1330,1761,1784]" italics="true" pageId="1" pageNumber="246">Venter:</emphasis>
Anterior prosternal margin sinuous and slightly crenulated, distinctly trilobed at middle, with middle lobe more prominent and conspicuous than lateral ones (
<figureCitation id="C18A2A65C451FFB3C47E813A8336F8DB" box="[569,638,1832,1857]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
F). Prosternal process high, but clearly not exceeding coxal height.
<emphasis id="6BC5EAF2C451FFB3C6D0815E8042F8FE" box="[151,266,1868,1893]" italics="true" pageId="1" pageNumber="246">Genitalia:</emphasis>
Parameres symmetrical, moderately elongated, with 2 lateral projections, basal projection larger, middle projection directed laterally; apex not diverging internally, parallel; subapex with small tooth; apex rounded (
<figureCitation id="C18A2A65C451FFB3C328816281EAF837" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
E).
</paragraph>
</subSubSection>
<subSubSection id="11AB656BC451FFB3C68081AA85E0F86E" pageId="1" pageNumber="246" type="etymology">
<paragraph id="590E36E0C451FFB3C68081AA85E0F86E" blockId="1.[151,1437,1325,2037]" pageId="1" pageNumber="246">
<emphasis id="6BC5EAF2C451FFB3C68081AA800EF84A" bold="true" box="[199,326,1976,2001]" pageId="1" pageNumber="246">Etymology</emphasis>
. This species is named in honor of the entomologist and friend, Dr. Angelo Barbosa Monteiro Machado, expert in the taxonomy of dragonflies (Odonata), on occasion of his 80th birthday.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,282 @@
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<mods:title id="73EE91F1803E3103CC33F1A8A4636A42">Two new species of Cyclocephala (Coleoptera: Scarabaeoidea: Melolonthidae) from Minas Gerais State, Brazil</mods:title>
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<mods:namePart id="5F58235C02AE63832188E62986BA2B35">Grossi, Paschoal C.</mods:namePart>
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<mods:namePart id="61CCA4009FB72DDAAFD235EB304BD13A">Almeida, Lúcia M.</mods:namePart>
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<treatment id="D11887F6C454FFB7C6D0832480C9F9A3" ID-DOI="http://doi.org/10.5281/zenodo.5659770" ID-GBIF-Taxon="119409861" ID-Zenodo-Dep="5659770" LSID="urn:lsid:plazi:treatment:D11887F6C454FFB7C6D0832480C9F9A3" httpUri="http://treatment.plazi.org/id/D11887F6C454FFB7C6D0832480C9F9A3" lastPageId="5" lastPageNumber="250" pageId="4" pageNumber="249">
<subSubSection id="11AB656BC454FFB6C6D08324803BFAEA" pageId="4" pageNumber="249" type="nomenclature">
<paragraph id="590E36E0C454FFB6C6D08324835FFACB" blockId="4.[151,535,1334,1394]" box="[151,535,1334,1360]" pageId="4" pageNumber="249">
<heading id="0246818CC454FFB6C6D08324835FFACB" bold="true" box="[151,535,1334,1360]" fontSize="11" level="1" pageId="4" pageNumber="249" reason="1">
<emphasis id="6BC5EAF2C454FFB6C6D08324835FFACB" bold="true" box="[151,535,1334,1360]" pageId="4" pageNumber="249">
<taxonomicName id="9EB14D63C454FFB6C6D0832480FBFACB" box="[151,435,1334,1360]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C454FFB6C6D0832480FBFACB" bold="true" box="[151,435,1334,1360]" italics="true" pageId="4" pageNumber="249">Cyclocephala everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C454FFB6C7FD8324835FFACB" box="[442,535,1334,1360]" pageId="4" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="590E36E0C454FFB6C6D0834B803BFAEA" blockId="4.[151,535,1334,1394]" box="[151,371,1369,1394]" pageId="4" pageNumber="249">
(
<figureCitation id="C18A2A65C454FFB6C6D9834B804DFAEA" box="[158,261,1369,1394]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="4" pageNumber="249">Figures 1</figureCitation>
G, 2AE)
</paragraph>
</subSubSection>
<subSubSection id="11AB656BC454FFB7C6D083B6818FFE13" lastPageId="5" lastPageNumber="250" pageId="4" pageNumber="249" type="materials_examined">
<paragraph id="590E36E0C454FFB6C6D083B682E2FA7B" blockId="4.[151,1436,1440,2008]" pageId="4" pageNumber="249">
<emphasis id="6BC5EAF2C454FFB6C6D083B68030FA26" bold="true" box="[151,376,1444,1469]" pageId="4" pageNumber="249">Material examined</emphasis>
.
<typeStatus id="860A8842C454FFB6C7C183B680A6FA26" box="[390,494,1444,1469]" pageId="4" pageNumber="249" type="holotype">Holotype</typeStatus>
male labeled: “
<collectingCountry id="21A67670C454FFB6C4DC83B683AAFA26" box="[667,738,1444,1469]" name="Brazil" pageId="4" pageNumber="249">Brasil</collectingCountry>
, MG,
<quantity id="9E499B05C454FFB6C56283B68215FA26" box="[805,861,1444,1469]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="249" unit="km" value="8.0">8 km</quantity>
ao N de/Cristália,
<date id="2D0F1020C454FFB6C26A83B685EAFA26" box="[1069,1186,1444,1469]" pageId="4" pageNumber="249" value="2010-02-16">16.ii.2010</date>
, luz/
<geoCoordinate id="3C855027C454FFB6C29F83B68465FA27" box="[1240,1325,1440,1469]" direction="south" orientation="latitude" pageId="4" pageNumber="249" precision="925" value="-16.65">16o39'S</geoCoordinate>
,
<geoCoordinate id="3C855027C454FFB6C37F83B684D0FA26" box="[1336,1432,1440,1469]" direction="west" orientation="longitude" pageId="4" pageNumber="249" precision="925" value="-42.883335">42o53'W</geoCoordinate>
,
<quantity id="9E499B05C454FFB6C6D083D58191FA7B" box="[151,217,1479,1504]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="4" pageNumber="249" unit="m" value="720.0">720m</quantity>
,/Grossi, Parizotto &amp; Melo/ao amanhecer (56: a.m.)” (
<collectionCode id="3FA0AE25C454FFB6C50683DA82D7FA7B" box="[833,927,1480,1504]" pageId="4" pageNumber="249">CERPE</collectionCode>
).
</paragraph>
<paragraph id="590E36E0C454FFB7C68083FE818FFE13" blockId="4.[151,1436,1440,2008]" lastBlockId="5.[151,1436,151,464]" lastPageId="5" lastPageNumber="250" pageId="4" pageNumber="249">
<emphasis id="6BC5EAF2C454FFB6C68083FE8032F99E" bold="true" box="[199,378,1516,1541]" pageId="4" pageNumber="249">
<typeStatus id="860A8842C454FFB6C68083FE807CF99E" box="[199,308,1516,1541]" pageId="4" pageNumber="249" type="holotype">Holotype</typeStatus>
male
</emphasis>
.
<emphasis id="6BC5EAF2C454FFB6C7C983FE836BF99E" box="[398,547,1516,1541]" italics="true" pageId="4" pageNumber="249">Total length:</emphasis>
<quantity id="9E499B05C454FFB6C46883FE83D4F99F" box="[559,668,1516,1540]" metricMagnitude="-3" metricUnit="m" metricValue="9.01" pageId="4" pageNumber="249" unit="mm" value="9.01">9.01 mm</quantity>
.
<emphasis id="6BC5EAF2C454FFB6C4EB83FE82EDF99E" box="[684,933,1516,1541]" italics="true" pageId="4" pageNumber="249">Total pronotal width:</emphasis>
<quantity id="9E499B05C454FFB6C5F483FE8557F99F" box="[947,1055,1516,1541]" metricMagnitude="-3" metricUnit="m" metricValue="3.59" pageId="4" pageNumber="249" unit="mm" value="3.59">3.59 mm</quantity>
.
<emphasis id="6BC5EAF2C454FFB6C27783FE853AF99E" box="[1072,1138,1516,1541]" italics="true" pageId="4" pageNumber="249">Body:</emphasis>
Dorsal surface glabrous, ventrally densely setose, setae distinctly elongate. Color brownish red, with lighter areas on clypeal apex, pronotum, scutellum (
<figureCitation id="C18A2A65C454FFB6C7DD802680ACF9D7" box="[410,484,1588,1613]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="4" pageNumber="249">Fig. 2</figureCitation>
A), legs, and venter.
<emphasis id="6BC5EAF2C454FFB6C4A380268262F9D6" box="[740,810,1588,1613]" italics="true" pageId="4" pageNumber="249">Head:</emphasis>
Shape transverse with abrupt narrowing to clypeus, sparsely punctate; punctures larger on clypeus and on sides; clypeus narrowly parabolic, ⅓ of head width, with elongated punctures, anteriorly concave, and with Cshaped punctures, apex reflexed and bordered (
<figureCitation id="C18A2A65C454FFB6C375806E8433F90F" box="[1330,1403,1660,1685]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="4" pageNumber="249">Fig. 2</figureCitation>
B). Frontoclypeal suture obsolete at middle, with 2 lateral, transverse impressions. Antennae with lamellae extremely developed, almost
<date id="2D0F1020C454FFB6C72C80D680CFF947" box="[363,391,1732,1756]" pageId="4" pageNumber="249">3x</date>
longer than antennomeres 37 together, and strongly and densely punctate with a spongiose aspect, and with scattered setae (
<figureCitation id="C18A2A65C454FFB6C44F80F58303F89B" box="[520,587,1767,1792]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="4" pageNumber="249">Fig. 2</figureCitation>
C); first lamella externally with longitudinal row of dense setae; eyes large, wider than clypeal width (
<figureCitation id="C18A2A65C454FFB6C7FE811E80B3F8BF" box="[441,507,1804,1829]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="4" pageNumber="249">Figs 1</figureCitation>
G2B).
<emphasis id="6BC5EAF2C454FFB6C41E811E83ADF8BE" box="[601,741,1804,1829]" italics="true" pageId="4" pageNumber="249">Mouthparts:</emphasis>
Mandibles slender, apices upturned and externally curved and with elongated setae, basal half laterally crenulated; last maxillar palpomere slightly longer than the preceding 2 together.
<emphasis id="6BC5EAF2C454FFB6C74181478037F8F7" box="[262,383,1877,1900]" italics="true" pageId="4" pageNumber="249">Pronotum:</emphasis>
Surface weakly and sparsely punctate, punctures stronger posteriorly; posterior margin slightly projected backward, rounded, with no bead; sides with long setae, weakly crenulated. Scutellum punctate; punctures small to large, some elongate.
<emphasis id="6BC5EAF2C454FFB6C418818E83E6F82E" box="[607,686,1948,1973]" italics="true" pageId="4" pageNumber="249">Elytra:</emphasis>
Surface distinctly and irregularly punctate; striae not well defined, slightly wrinkled near suture.
<emphasis id="6BC5EAF2C454FFB6C7A481D3836AF843" box="[483,546,1985,2008]" italics="true" pageId="4" pageNumber="249">Legs:</emphasis>
Protibiae with 3 distinct teeth, teeth decreasing in size from apex to base; apical tooth twice as long as middle tooth, middle tooth more than
<date id="2D0F1020C455FFB7C516868A8225FF2B" box="[849,877,152,176]" pageId="5" pageNumber="250">3x</date>
longer than basal tooth (
<figureCitation id="C18A2A65C455FFB7C2CA8685859EFF2B" box="[1165,1238,151,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="5" pageNumber="250">Figs 2</figureCitation>
AB); inner claw widely bifid, forming a right angle (
<figureCitation id="C18A2A65C455FFB7C47586AE833FFF4F" box="[562,631,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="5" pageNumber="250">Fig. 2</figureCitation>
D).
<emphasis id="6BC5EAF2C455FFB7C4E486AF83B8FF4F" box="[675,752,189,212]" italics="true" pageId="5" pageNumber="250">Venter:</emphasis>
Anterior prosternal margin sinuous and crenulated by setae insertion, distinctly trilobed at middle, with lateral lobes more prominent than middle lobe, conspicuous (
<figureCitation id="C18A2A65C455FFB7C37286CD843DFF63" box="[1333,1397,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="5" pageNumber="250">Fig. 1</figureCitation>
G). Prosternal process high, exceeding coxal height.
<emphasis id="6BC5EAF2C455FFB7C4FF87168262FE86" box="[696,810,260,285]" italics="true" pageId="5" pageNumber="250">Genitalia:</emphasis>
Parameres symmetrical with elongated parameres, with 2 lateral projections, one basal and one at middle; basal larger, middle one projecting backwards; apices diverging internally, externally with small serrations; subapex with a slightly diverging, rounded tooth at external angle (
<figureCitation id="C18A2A65C455FFB7C328875E81EEFE13" captionStart="FIGURE 2" captionStartId="3.[151,250,1966,1988]" captionTargetBox="[215,1381,209,1933]" captionTargetId="figure@3.[187,1400,193,1945]" captionTargetPageId="3" captionText="FIGURE 2. Cyclocephala everardoi sp. nov. A. holotype male, dorsal habitus; B. head pronotum, and protibiae in detail; C. antennal lamellae; D. anterior protarsus; E. parameres in caudal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/255901/files/figure.png" pageId="5" pageNumber="250">Fig. 2</figureCitation>
E).
</paragraph>
</subSubSection>
<subSubSection id="11AB656BC455FFB7C680878680C9F9A3" pageId="5" pageNumber="250" type="etymology">
<paragraph id="590E36E0C455FFB7C6808786857AFE4B" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="250">
<emphasis id="6BC5EAF2C455FFB7C6808786800EFE36" bold="true" box="[199,326,404,429]" pageId="5" pageNumber="250">Etymology</emphasis>
. This species is named in honor of the dedicated entomologist, Everardo José Grossi, father of the first author, a doctor with a passion for beetles and an expert in rhinoceros beetles.
</paragraph>
<caption id="0DCE6668C455FFB7C6D084128583FD8C" ID-Table-UUID="0DCE6668C455FFB7C6D084128583FD8C" box="[151,1227,511,535]" httpUri="http://table.plazi.org/id/0DCE6668C455FFB7C6D084128583FD8C" pageId="5" pageNumber="250" targetBox="[151,1408,562,1199]" targetIsTable="true" targetPageId="5">
<paragraph id="590E36E0C455FFB7C6D084128583FD8C" blockId="5.[151,1227,511,535]" box="[151,1227,511,535]" pageId="5" pageNumber="250">
TABLE 1. Main characters to differentiate the two new species of
<taxonomicName id="9EB14D63C455FFB7C5C487ED855FFD8C" box="[899,1047,511,535]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="6BC5EAF2C455FFB7C5C487ED855FFD8C" bold="true" box="[899,1047,511,535]" italics="true" pageId="5" pageNumber="250">Cyclocephala</emphasis>
</taxonomicName>
described here.
</paragraph>
</caption>
<paragraph id="590E36E0C455FFB7C6D884208053FB34" pageId="5" pageNumber="250">
<table id="2BB1C440C455004DC6D0842084C8FB34" box="[151,1408,562,1199]" gridcols="3" gridrows="9" pageId="5" pageNumber="250">
<tr id="E78134A2C455004DC6D0842084C8FDD3" box="[151,1408,562,584]" gridrow="0" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08420802DFDD3" box="[151,357,562,584]" gridcol="0" gridrow="0" pageId="5" pageNumber="250">
<emphasis id="6BC5EAF2C455FFB7C6D884208040FDD3" bold="true" box="[159,264,562,584]" pageId="5" pageNumber="250">Character</emphasis>
</th>
<th id="A4505DDEC455004DC7B1842082E5FDD3" box="[502,941,562,584]" gridcol="1" gridrow="0" pageId="5" pageNumber="250">
<emphasis id="6BC5EAF2C455FFB7C7B18420838EFDD3" bold="true" box="[502,710,562,584]" pageId="5" pageNumber="250">
<taxonomicName id="9EB14D63C455FFB7C7B18420833CFDD3" box="[502,628,562,584]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C7B18420833CFDD3" bold="true" box="[502,628,562,584]" italics="true" pageId="5" pageNumber="250">C. machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C43D8420838EFDD3" box="[634,710,562,584]" pageId="5" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="A4505DDEC455004DC581842084C8FDD3" box="[966,1408,562,584]" gridcol="2" gridrow="0" pageId="5" pageNumber="250">
<emphasis id="6BC5EAF2C455FFB7C581842085D9FDD3" bold="true" box="[966,1169,562,584]" pageId="5" pageNumber="250">
<taxonomicName id="9EB14D63C455FFB7C58184208577FDD3" box="[966,1087,562,584]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C58184208577FDD3" bold="true" box="[966,1087,562,584]" italics="true" pageId="5" pageNumber="250">C. everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C202842085D9FDD3" box="[1093,1169,562,584]" pageId="5" pageNumber="250" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="E78134A2C455004DC6D0844884C8FDF4" box="[151,1408,602,623]" gridrow="1" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08448802DFDF4" box="[151,357,602,623]" gridcol="0" gridrow="1" pageId="5" pageNumber="250">Color</th>
<td id="A4505DDEC455004DC7B1844882E5FDF4" box="[502,941,602,623]" gridcol="1" gridrow="1" pageId="5" pageNumber="250">Testaceous with black head</td>
<td id="A4505DDEC455004DC581844884C8FDF4" box="[966,1408,602,623]" gridcol="2" gridrow="1" pageId="5" pageNumber="250">Brownish red</td>
</tr>
<tr id="E78134A2C455004DC6D0849084C8FD2F" box="[151,1408,642,692]" gridrow="2" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08490802DFD2F" box="[151,357,642,692]" gridcol="0" gridrow="2" pageId="5" pageNumber="250">Clypeus</th>
<td id="A4505DDEC455004DC7B1849082E5FD2F" box="[502,941,642,692]" gridcol="1" gridrow="2" pageId="5" pageNumber="250">Almost impunctate, strongly reflexed.</td>
<td id="A4505DDEC455004DC581849084C8FD2F" box="[966,1408,642,692]" gridcol="2" gridrow="2" pageId="5" pageNumber="250">With elongated and Cshaped punctures, slightly reflexed</td>
</tr>
<tr id="E78134A2C455004DC6D084D584C8FD61" box="[151,1408,711,762]" gridrow="3" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D084D5802DFD61" box="[151,357,711,762]" gridcol="0" gridrow="3" pageId="5" pageNumber="250">Frontoclypeal suture</th>
<td id="A4505DDEC455004DC7B184D582E5FD61" box="[502,941,711,762]" gridcol="1" gridrow="3" pageId="5" pageNumber="250">Obsolete, lacking impression</td>
<td id="A4505DDEC455004DC58184D584C8FD61" box="[966,1408,711,762]" gridcol="2" gridrow="3" pageId="5" pageNumber="250">Obsolete at middle and with two transverse impressions</td>
</tr>
<tr id="E78134A2C455004DC6D0851F84C8FCA4" box="[151,1408,781,831]" gridrow="4" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D0851F802DFCA4" box="[151,357,781,831]" gridcol="0" gridrow="4" pageId="5" pageNumber="250">Pronotum</th>
<td id="A4505DDEC455004DC7B1851F82E5FCA4" box="[502,941,781,831]" gridcol="1" gridrow="4" pageId="5" pageNumber="250">Surface moderately and uniformly punctate, smooth at center line</td>
<td id="A4505DDEC455004DC581851F84C8FCA4" box="[966,1408,781,831]" gridcol="2" gridrow="4" pageId="5" pageNumber="250">Surface weakly and sparsely punctate, punctures posteriorly stronger</td>
</tr>
<tr id="E78134A2C455004DC6D0854084C8FC1F" box="[151,1408,850,900]" gridrow="5" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08540802DFC1F" box="[151,357,850,900]" gridcol="0" gridrow="5" pageId="5" pageNumber="250">Elytra</th>
<td id="A4505DDEC455004DC7B1854082E5FC1F" box="[502,941,850,900]" gridcol="1" gridrow="5" pageId="5" pageNumber="250">Striae distinct, marked by lines of punctures; interstriae almost impunctate</td>
<td id="A4505DDEC455004DC581854084C8FC1F" box="[966,1408,850,900]" gridcol="2" gridrow="5" pageId="5" pageNumber="250">Striae indistinct and surface nearly totally, uniformly punctate</td>
</tr>
<tr id="E78134A2C455004DC6D0858584C8FC7C" box="[151,1408,919,999]" gridrow="6" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D08585802DFC7C" box="[151,357,919,999]" gridcol="0" gridrow="6" pageId="5" pageNumber="250">Protibiae</th>
<td id="A4505DDEC455004DC7B1858582E5FC7C" box="[502,941,919,999]" gridcol="1" gridrow="6" pageId="5" pageNumber="250">Basal tooth weak, and more than 3x smaller than middle tooth</td>
<td id="A4505DDEC455004DC581858584C8FC7C" box="[966,1408,919,999]" gridcol="2" gridrow="6" pageId="5" pageNumber="250">Basal tooth more developed, and distinctly projected, three times smaller than middle tooth</td>
</tr>
<tr id="E78134A2C455004DC6D085E884C8FBB7" box="[151,1408,1018,1068]" gridrow="7" pageId="5" pageNumber="250">
<th id="A4505DDEC455004DC6D085E8802DFBB7" box="[151,357,1018,1068]" gridcol="0" gridrow="7" pageId="5" pageNumber="250">Aedeagi</th>
<td id="A4505DDEC455004DC7B185E882E5FBB7" box="[502,941,1018,1068]" gridcol="1" gridrow="7" pageId="5" pageNumber="250">Parameres with apices parallel; middle tooth small, projecting laterally</td>
<td id="A4505DDEC455004DC58185E884C8FBB7" box="[966,1408,1018,1068]" gridcol="2" gridrow="7" pageId="5" pageNumber="250">Parameres with apices diverging; middle tooth more developed, projecting backwards</td>
</tr>
<tr id="E78134A2C455004DC6D0828784C8FB34" box="[151,1408,1173,1199]" gridrow="8" pageId="5" pageNumber="250" rowspan-1="1" rowspan-2="1">
<th id="A4505DDEC455004DC6D08287802DFB34" box="[151,357,1173,1199]" gridcol="0" gridrow="8" pageId="5" pageNumber="250">Discussion</th>
</tr>
</table>
</paragraph>
<paragraph id="590E36E0C455FFB7C6D082CE80C9F9A3" blockId="5.[151,1436,1244,1592]" pageId="5" pageNumber="250">
The specimens were collected with UV light.
<taxonomicName id="9EB14D63C455FFB7C4F682CE8289FB6E" box="[689,961,1244,1269]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C4F682CE8289FB6E" box="[689,961,1244,1269]" italics="true" pageId="5" pageNumber="250">Cyclocephala everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C58B82CE8561FB6E" box="[972,1065,1244,1269]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C58B82CE8561FB6E" bold="true" box="[972,1065,1244,1269]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
was taken at dawn in
<date id="2D0F1020C455FFB7C37082CE819AFA83" pageId="5" pageNumber="250" value="2010-02">February 2010</date>
, between 5:00 and 6:00 A.M., in an area of sandy and rupestrian Cerrado fields (
<figureCitation id="C18A2A65C455FFB7C2C982ED859BFA83" box="[1166,1235,1279,1304]" captionStart="FIGURE 3" captionStartId="4.[151,250,1212,1234]" captionTargetBox="[177,1408,243,1176]" captionTargetId="figure@4.[171,1416,193,1190]" captionTargetPageId="4" captionText="FIGURE 3. General aspect of the region of Cristália municipality, Minas Gerais State, Brazil, where the holotype of C. everardoi sp. nov. was collected." httpUri="https://zenodo.org/record/255902/files/figure.png" pageId="5" pageNumber="250">Fig. 3</figureCitation>
A).
<taxonomicName id="9EB14D63C455FFB7C34582ED804DFAA6" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C34582ED804DFAA6" italics="true" pageId="5" pageNumber="250">Cyclocephala machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C74A8336802FFAA6" box="[269,359,1316,1341]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C74A8336802FFAA6" bold="true" box="[269,359,1316,1341]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
lacks other information. Both
<typeStatus id="860A8842C455FFB7C48483368278FAA6" box="[707,816,1316,1341]" pageId="5" pageNumber="250" type="holotype">holotypes</typeStatus>
are pinned; in
<taxonomicName id="9EB14D63C455FFB7C59C83378520FAA6" box="[987,1128,1316,1341]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C59C83378520FAA6" box="[987,1128,1316,1341]" italics="true" pageId="5" pageNumber="250">C. everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C23783368582FAA6" box="[1136,1226,1316,1341]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C23783368582FAA6" bold="true" box="[1136,1226,1316,1341]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
part of the middle tooth of left protibia is missing, while in
<taxonomicName id="9EB14D63C455FFB7C42F835A83BFFAFB" box="[616,759,1351,1376]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C42F835A83BFFAFB" box="[616,759,1351,1376]" italics="true" pageId="5" pageNumber="250">C. machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C54783558212FAFB" box="[768,858,1351,1376]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C54783558212FAFB" bold="true" box="[768,858,1351,1376]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
part of the right first and second antennal lamellae are missing. They were initially confused with
<taxonomicName id="9EB14D63C455FFB7C4E0837E85C9FA1F" authority="Erichson 1847" authorityName="Erichson" authorityYear="1847" box="[679,1153,1388,1413]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="macrophylla">
<emphasis id="6BC5EAF2C455FFB7C4E0837E829DFA1E" box="[679,981,1388,1413]" italics="true" pageId="5" pageNumber="250">Cyclocephala macrophylla</emphasis>
Erichson 1847
</taxonomicName>
.
<taxonomicName id="9EB14D63C455FFB7C2CB837E84D3FA1E" box="[1164,1435,1388,1413]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C2CB837E84D3FA1E" box="[1164,1435,1388,1413]" italics="true" pageId="5" pageNumber="250">Cyclocephala machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C6D0839D81B9FA33" box="[151,241,1423,1448]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C6D0839D81B9FA33" bold="true" box="[151,241,1423,1448]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
is similar to
<taxonomicName id="9EB14D63C455FFB7C7C383828359FA33" box="[388,529,1423,1448]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="species" species="everardoi" status="sp. nov.">
<emphasis id="6BC5EAF2C455FFB7C7C383828359FA33" box="[388,529,1423,1448]" italics="true" pageId="5" pageNumber="250">C. everardoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C455FFB7C45F839D833AFA33" box="[536,626,1423,1448]" pageId="5" pageNumber="250" rank="species">
<emphasis id="6BC5EAF2C455FFB7C45F839D833AFA33" bold="true" box="[536,626,1423,1448]" pageId="5" pageNumber="250">sp. nov.</emphasis>
</taxonomicNameLabel>
and was also confused with this. In comparing them, we concluded that they were similar but distinct species and they are diagnosed in the
<tableCitation id="1433035BC455FFB7C5DD83A682A7FA57" box="[922,1007,1460,1485]" captionStart="TABLE 1" captionStartId="5.[151,239,512,535]" captionTargetBox="[151,1408,562,1199]" captionTargetPageId="5" captionText="TABLE 1. Main characters to differentiate the two new species of Cyclocephala described here." httpUri="http://table.plazi.org/id/0DCE6668C455FFB7C6D084128583FD8C" pageId="5" pageNumber="250" tableUuid="0DCE6668C455FFB7C6D084128583FD8C">Table 1</tableCitation>
. We also believe that they compose a new species group within
<emphasis id="6BC5EAF2C455FFB7C78383C5832DFA6B" box="[452,613,1495,1520]" italics="true" pageId="5" pageNumber="250">
<taxonomicName id="9EB14D63C455FFB7C78383C58328FA6B" box="[452,608,1495,1520]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="250" phylum="Arthropoda" rank="genus">Cyclocephala</taxonomicName>
,
</emphasis>
being diagnosed by the short body length, extremely elongate antennal lamellae, eyes very developed, clypeus elongate and truncate, somewhat quadrate and inner claw of anterior tarsi with a wide incision.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,271 @@
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<mods:title id="22914D401770C86D66F20D8530149FD8">The Buprestidae (Coleoptera, Buprestoidea) of the Tuscan Archipelago (Italy)</mods:title>
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<paragraph id="3499861A670BDB31D264CAB8EA800D7C" pageId="0" pageNumber="117362">Materials</paragraph>
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<emphasis id="123185A51A3CE2328D1750D4CFEDCB2A" bold="true" pageId="0" pageNumber="117362">Type status:</emphasis>
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:
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:
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scientificName:
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(Anthaxia) mendizabali
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:
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Via
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<collectorName id="E49923F3232057FDAE7E2009BDC18346">Identification</collectorName>
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identifiedBy:
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eventDate:
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<collectorName id="6BA86658B760C6D77EC872BF9C70E20B">Record Level</collectorName>
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collectionCode: LFPC
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status:
</emphasis>
<materialsCitation id="AC4C467859ACF1D5EF071307B9CC895F" collectingDate="2014-01-06" collectorName="Leonardo Forbicioni, Identification, E. Paggetti, Event, Record Level" country="Italy" county="Sant'Ilario" determinerName="E. Paggetti" latitude="42.763233" location="Occurrence" longitude="10.221358" municipality="Campo" specimenCount="1" specimenCount-adult="1" typeStatus="Other material">
<typeStatus id="410213AC2D36E832853EDAE8FAFDDB7D" pageId="0" pageNumber="117362">Other material</typeStatus>
.
<emphasis id="2BF0AA0792C4880EB8A636467D833EC1" bold="true" pageId="0" pageNumber="117362">
<location id="571CC3A3E2667F16D6D15F4344D74E18" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:571CC3A3E2667F16D6D15F4344D74E18" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Occurrence">Occurrence</location>
:
</emphasis>
recordedBy:
<collectorName id="E6F7A2F702F087F8A3AB74DD978D1510" pageId="0" pageNumber="117362">
<location id="F66483713106723578EF76C886837AB4" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:F66483713106723578EF76C886837AB4" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Leonardo Forbicioni">Leonardo Forbicioni</location>
</collectorName>
; individualCount:
<specimenCount id="7D6C603DCB69AEE7EAEEEB34E1B8AE3D" count="1" pageId="0" pageNumber="117362" type="generic">1</specimenCount>
; lifeStage:
<specimenType id="43C29204C9DEB0832E1AD682C26791E6" pageId="0" pageNumber="117362">adult</specimenType>
; occurrenceID:
<uuid id="8DFBA6826146D7FEB865E4B17E35BE96">2BBABC28-A652-52DD-A340-4C167163B7E5</uuid>
;
<emphasis id="6B48AE895483F62EB68D9A018C63E1A2" bold="true" pageId="0" pageNumber="117362">
<location id="344A91732EA86F058AF2DEA1890D917E" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:344A91732EA86F058AF2DEA1890D917E" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Taxon">Taxon</location>
:
</emphasis>
scientificName:
<location id="4B29F5B54B246218EFDDA82C8A36DE1B" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:4B29F5B54B246218EFDDA82C8A36DE1B" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Anthaxia">Anthaxia</location>
(Anthaxia) mendizabali
<location id="2A72997DAE8564604A6A25E56EC25D28" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:2A72997DAE8564604A6A25E56EC25D28" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Cobos">Cobos</location>
, 1965; order:
<location id="BDBEE31B9B3AB924BA90F5A6018D52EE" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:BDBEE31B9B3AB924BA90F5A6018D52EE" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Coleoptera">Coleoptera</location>
; family:
<location id="A4C7D651836E11EBF0DA9019368A8A6B" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:A4C7D651836E11EBF0DA9019368A8A6B" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Buprestidae">Buprestidae</location>
; genus:
<location id="E44C075B07AC7E826CBA420A8A17AA84" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:E44C075B07AC7E826CBA420A8A17AA84" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Anthaxia">Anthaxia</location>
; subgenus:
<location id="6403A5E0DB6658698B1B9F7D7F414C42" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:6403A5E0DB6658698B1B9F7D7F414C42" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Anthaxia">Anthaxia</location>
; specificEpithet: mendizabali; scientificNameAuthorship:
<location id="A216845EDA134A50A2D1D3B931E0225D" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:A216845EDA134A50A2D1D3B931E0225D" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Cobos">Cobos</location>
, 1965;
<emphasis id="FAAD3766B786B3DBA253189786E4B475" bold="true" pageId="0" pageNumber="117362">
<location id="E0D347CC6DED4D78CCABACACEBAA8F96" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:E0D347CC6DED4D78CCABACACEBAA8F96" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Location">Location</location>
:
</emphasis>
islandGroup:
<location id="378ADA8DC796180A1D47E35AAD9C7E8A" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:378ADA8DC796180A1D47E35AAD9C7E8A" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Tuscan Archipelago">Tuscan Archipelago</location>
; island: Isola
<location id="5BC6D4EF4C1100CD06EAEE37A96A2B0B" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:5BC6D4EF4C1100CD06EAEE37A96A2B0B" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="d'Elba">d'Elba</location>
; country:
<collectingCountry id="A26178B41EF6D70B03BA11C36CA9459B" name="Italy" pageId="0" pageNumber="117362">Italy</collectingCountry>
; countryCode: IT; stateProvince:
<location id="AC50CCCC0CF2CDCB52029D99B8BAD229" LSID="urn:lsid:plazi:treatment:D118DDA3EAE2529E9923956405846178:AC50CCCC0CF2CDCB52029D99B8BAD229" country="Italy" county="Sant'Ilario" latitude="42.763233" longitude="10.221358" municipality="Campo" name="Livorno">Livorno</location>
; county:
<collectingMunicipality id="03BC651114A23CBD9B10DDB6FA72EB6D">Campo</collectingMunicipality>
nell'Elba; municipality:
<collectingCounty id="1C5B8533CF1D55648D183CC8E05BA2FD">Sant'Ilario</collectingCounty>
; decimalLatitude:
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; decimalLongitude:
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; geodeticDatum: WGS84; coordinatePrecision: 0.0002;
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<collectorName id="4EC1CFBEAE412D475BC27858F601FEB3">Identification</collectorName>
:
</emphasis>
identifiedBy:
<determinerName id="985902B041BCF417811F20CF9197859F" pageId="0" pageNumber="117362">
<collectorName id="DD1D5889046BDB3A741FB2C2AE3DBBA7">E. Paggetti</collectorName>
</determinerName>
;
<emphasis id="A2B6EBB4542DDEBA4C9E25BE806F9FF0" bold="true" pageId="0" pageNumber="117362">
<collectorName id="9F5D1EBABA1ACA39D6EB58431A97B5A4">Event</collectorName>
:
</emphasis>
eventDate:
<collectingDate id="7B228F7146F4B16897728304C3EB2B27" pageId="0" pageNumber="117362" value="2014-01-06">2014-06-01</collectingDate>
;
<emphasis id="5DEDD4AB402D7C768074543B43CDCB5F" bold="true" pageId="0" pageNumber="117362">
<collectorName id="77601531E3F6B8AA4CEB6161BC0EEA35">Record Level</collectorName>
:
</emphasis>
collectionCode: LFPC
</materialsCitation>
</materialsCitation>
</materialsCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="69883266A67707EDC905FDAF9424792B" pageId="0" pageNumber="117362" type="conservation status">
<paragraph id="E884EFE0C83C85A77EC719B8B92F0BA3" pageId="0" pageNumber="117362">Conservation status</paragraph>
<paragraph id="3E881A2C29FC6A9EAC6FB070D9FE6762" pageId="0" pageNumber="117362">LC</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,82 @@
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.602.8136" ID-PMC="PMC4978033" ID-Pensoft-Pub="1313-2970-602-1" ID-Pensoft-UUID="FFA86D39FFE2FFF3FF8AFFEBC209FFDE" ID-PubMed="27551193" ID-Zenodo-Dep="126863" ID-ZooBank="65EFA27673454AC69B78DBE7E98D6103" ModsDocID="1313-2970-602-1" checkinTime="1467777339758" checkinUser="pensoft" docAuthor="Neubauer, Thomas A." docDate="2016" docId="D118DF7543775FC1B550B982B7759AF1" docLanguage="en" docName="ZooKeys 602: 1-358" docOrigin="ZooKeys 602" docPubDate="2016-07-05" docSource="http://dx.doi.org/10.3897/zookeys.602.8136" docTitle="Melanopsis (Mesopotamia) aroussiana subsp. var. var. semilaevigata Pallary 1939, [invalid]" docType="treatment" docVersion="4" id="FFA86D39FFE2FFF3FF8AFFEBC209FFDE" lastPageNumber="247" masterDocId="FFA86D39FFE2FFF3FF8AFFEBC209FFDE" masterDocTitle="A nomenclator of extant and fossil taxa of the Melanopsidae (Gastropoda, Cerithioidea)" masterLastPageNumber="358" masterPageNumber="1" pageNumber="247" updateTime="1668163284802" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A nomenclator of extant and fossil taxa of the Melanopsidae (Gastropoda, Cerithioidea)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Neubauer, Thomas A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1398-9941</mods:nameIdentifier>
<mods:affiliation>Geological-Paleontological Department, Natural History Museum Vienna, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">thomas.neubauer@nhm-wien.ac.at</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-07-05</mods:number>
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<mods:detail type="volume">
<mods:number>602</mods:number>
</mods:detail>
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<mods:start>1</mods:start>
<mods:end>358</mods:end>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="Zenodo-Dep">126863</mods:identifier>
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<treatment ID-GBIF-Taxon="182232938" LSID="urn:lsid:plazi:treatment:D118DF7543775FC1B550B982B7759AF1" httpUri="http://treatment.plazi.org/id/D118DF7543775FC1B550B982B7759AF1" lastPageNumber="247" pageId="246" pageNumber="247">
<subSubSection pageId="246" pageNumber="247" type="nomenclature">
<paragraph pageId="246" pageNumber="247">
<taxonomicName LSID="D118DF75-4377-5FC1-B550-B982B7759AF1" authority="Pallary, 1939" authorityName="Pallary" authorityYear="1939" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Melanopsis (Mesopotamia) aroussiana subsp. var. var. semilaevigata" order="Sorbeoconcha" pageId="246" pageNumber="247" phylum="Mollusca" rank="variety" species="aroussiana" status="[invalid]" subGenus="Mesopotamia" subSpecies="var." variety="semilaevigata">Melanopsis (Mesopotamia) aroussiana var. semilaevigata Pallary, 1939</taxonomicName>
<taxonomicNameLabel pageId="246" pageNumber="247">[invalid]</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="246" pageNumber="247" type="original source">
<paragraph pageId="246" pageNumber="247">Original source.</paragraph>
<paragraph pageId="246" pageNumber="247">
<bibRefCitation author="Pallary, P" journalOrPublisher="Memoires de l'Institut d'Egypte" pageId="325" pageNumber="326" pagination="1 - 141" refId="B395" refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141" title="Deuxieme addition a la faune malacologique de la Syrie." volume="39" year="1939">Pallary 1939</bibRefCitation>
: 101.
</paragraph>
</subSubSection>
<subSubSection pageId="246" pageNumber="247" type="type locality">
<paragraph pageId="246" pageNumber="247">Type locality.</paragraph>
<paragraph pageId="246" pageNumber="247">
Not explicitly stated but probably the same as for the species (
<normalizedToken originalValue="“‘">&quot;'</normalizedToken>
Ain Arouss&quot; ['Ayn al 'Arūs, near Tall Abyaḑ], Syria).
</paragraph>
</subSubSection>
<subSubSection pageId="246" pageNumber="247" type="remarks">
<paragraph pageId="246" pageNumber="247">Remarks.</paragraph>
<paragraph pageId="246" pageNumber="247">
<bibRefCitation author="Pallary, P" journalOrPublisher="Memoires de l'Institut d'Egypte" pageId="325" pageNumber="326" pagination="1 - 141" refId="B395" refString="Pallary, P, 1939. Deuxieme addition a la faune malacologique de la Syrie. Memoires de l'Institut d'Egypte 39: 1 - 141" title="Deuxieme addition a la faune malacologique de la Syrie." volume="39" year="1939">Pallary (1939)</bibRefCitation>
indicated an illustration (&quot;Fig. 4&quot;) of this taxon, which, however, does not appear as such in the plate captions. Junior homonym of
<taxonomicName authorityName="Pallary" authorityYear="1928" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" lsidName="Melanopsis seurati subsp. semilaevigata" order="Pteropoda" pageId="246" pageNumber="247" phylum="Mollusca" rank="subSpecies" species="seurati" subSpecies="semilaevigata">
<emphasis italics="true" pageId="246" pageNumber="247">Melanopsis seurati semilaevigata</emphasis>
</taxonomicName>
Pallary, 1928 and
<taxonomicName authorityName="Pallary" authorityYear="1928" class="Gastropoda" family="Melanopsidae" genus="Melanopsis" kingdom="Animalia" lsidName="Melanopsis eximia subsp. semilaevigata" order="Pteropoda" pageId="246" pageNumber="247" phylum="Mollusca" rank="subSpecies" species="eximia" subSpecies="semilaevigata">
<emphasis italics="true" pageId="246" pageNumber="247">Melanopsis eximia semilaevigata</emphasis>
</taxonomicName>
Pallary, 1928 (simultaneously published; no priority fixed yet; see Note 1).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,136 @@
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<mods:title>Description of a Cretaceous amber fossil putatively of the tribe Coprophilini (Coleoptera, Staphylinidae, Oxytelinae)</mods:title>
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<mods:name type="personal">
<mods:role>
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<mods:namePart>Makranczy, Gyoergy</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yamamoto, Shuhei</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Engel, Michael S.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:date>2018</mods:date>
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<mods:identifier type="ZooBank">789CE1ADE7784C898DBFAD7A01C7ECDE</mods:identifier>
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<treatment ID-GBIF-Taxon="147292387" LSID="urn:lsid:zoobank.org:act:910F415D-CEA9-4E31-BF75-801F8A73BE80" httpUri="http://treatment.plazi.org/id/D118E07F0BC3361F11AF8DDC95559D80" lastPageId="3" lastPageNumber="84" pageId="1" pageNumber="82">
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<taxonomicName LSID="http://zoobank.org/910F415D-CEA9-4E31-BF75-801F8A73BE80" class="Insecta" family="Staphylinidae" genus="Gollandia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gollandia" order="Coleoptera" pageId="1" pageNumber="82" phylum="Arthropoda" rank="genus">Gollandia</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="82">gen. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="82" type="type species">
<paragraph pageId="1" pageNumber="82">Type species.</paragraph>
<paragraph pageId="1" pageNumber="82">
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gollandia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gollandia planata" order="Coleoptera" pageId="1" pageNumber="82" phylum="Arthropoda" rank="species" species="planata">Gollandia planata</taxonomicName>
sp. n., (described below).
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="83" pageId="1" pageNumber="82" type="diagnosis">
<paragraph pageId="1" pageNumber="82">Diagnosis.</paragraph>
<paragraph lastPageId="2" lastPageNumber="83" pageId="1" pageNumber="82">
Head. Head somewhat retracted under large pronotum; head capsule rather short. Epistomal sulcus not well visible, but presence suggested by a tranvserse
<normalizedToken originalValue="run">'run'</normalizedToken>
of air between amber and cuticle. Supraantennal prominences weak. Antennomeres with long tactile setae near apices (prominent on articles 3-11). Labial palp trimerous, basal two palpomeres rather stout, last palpomere thin. Labrum with two thick, forward-directed setae. Mandibles not prominent, apices acute. Maxillary palp tetramerous, basal three palpomeres moderately elongate, last palpomere much wider
<pageBreakToken pageId="2" pageNumber="83" start="start">and</pageBreakToken>
long, not reduced, apex pointed. Gular sulci seemingly widely separated at base but confluent anteriorly (this area is not well visible as preserved). Neck separated by gentle constriction and (at least laterally) a groove. Thorax. Pronotum strongly explanate, margin slightly reflexed, marginal bead present, lateral edge finely serrate/sinuous. Laterally with a strong seta at each of
<normalizedToken originalValue="anteroangularis">'anteroangularis'</normalizedToken>
and
<normalizedToken originalValue="lateralis">'lateralis'</normalizedToken>
positions, plus strong seta on both sides well inside lateral margin at about 1/3 length, posterior edge appearing slightly concave (might be artefact of preservation). Pronotal disc with shallow impressions, with fine and dense punctation and setation. Procoxae contiguous, projecting; procoxal fissure present and open (Figure 7). Mesoscutellum (Figure 8) with apex exposed and somewhat impressed without distinct pattern. Elytra finely and randomly punctate. Mesocoxae narrowly separated by mesosternal process (Figure 9). Legs slender (metatibia especially elongate), with regular rows of tibial spines (more slender than strong), and a conspicuous mesotibial spur (and a second spur half size at half-length towards femoral joint). Tarsal formula 4-4-4, no tarsal lobes (Figure 10), but empodial setae strong (Figure 11). Elytra with epipleural ridge, seemingly with a fine and shallow dorsal groove following it from inside, epipleura strongly deflexed and rather wide but epipleural fold thin to inconspicuous. Post-scutellar area with a pair of elongate impressions along suture. Shoulders prominently developed, narrowly rounded, even slightly projecting forward in relation to anterior edge at mesoscutellar area, posterior margin slightly oblique but straight from suture to outer 3/4, slightly incurved (concave) in outer 1/4 thereby producing a somewhat sharp outer corner in dorsal view. Abdomen. Abdomen with only six visible segments (not counting segments
<normalizedToken originalValue="IXX">IX-X</normalizedToken>
, often retracted under VIII), second abdominal segment not developed. With two pairs of laterosclerites. Apex of tergite VII seemingly without well-developed palisade fringe (difficult to judge; an air bubble under this structure obscures almost its entire width), apex of segment conspicuously widening (not narrowing to base of next segment), surface somewhat concave. Tergite VIII with apical edge truncate medially or slightly concave. Apex of sternite VIII without modification.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="84" pageId="2" pageNumber="83" type="differential diagnosis">
<paragraph pageId="2" pageNumber="83">Differential diagnosis.</paragraph>
<paragraph lastPageId="3" lastPageNumber="84" pageId="2" pageNumber="83">
All extant
<taxonomicName lsidName="" pageId="2" pageNumber="83" rank="tribe" tribe="Coprophilini">Coprophilini</taxonomicName>
have a 5-5-5 tarsal formula, and even the fossil genus
<taxonomicName class="Insecta" family="Staphylinidae" genus="Mesocoprophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesocoprophilus" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">Mesocoprophilus</taxonomicName>
has five tarsomeres, so the 4-4-4 condition in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gollandia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gollandia" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">Gollandia</taxonomicName>
is significant. The new genus differs greatly from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Mesocoprophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesocoprophilus" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">Mesocoprophilus</taxonomicName>
in the antennal structure, stout and short in
<taxonomicName class="Insecta" family="Staphylinidae" genus="Mesocoprophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesocoprophilus" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">Mesocoprophilus</taxonomicName>
, slender and elongate with well-developed tactile setae on all antennomeres in the present fossil. The neck (lateral constriction, postoccipital groove) also differentiates this genus from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Mesocoprophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesocoprophilus" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">Mesocoprophilus</taxonomicName>
where these features are absent. The lack of striae or puncture rows on the elytra makes this genus distinct from all extant
<taxonomicName lsidName="" pageId="2" pageNumber="83" rank="tribe" tribe="Coprophilini">Coprophilini</taxonomicName>
, while a distinction from
<taxonomicName class="Insecta" family="Staphylinidae" genus="Mesocoprophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesocoprophilus" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="genus">Mesocoprophilus</taxonomicName>
cannot be made as that fossil lacks its dorsal portion (
<bibRefCitation author="Cai, C-Y" journalOrPublisher="Insect Systematics &amp; Evolution" pageId="6" pageNumber="87" pagination="213 - 220" title="Mesocoprophilusclavatus, a new oxyteline rove beetle (Coleoptera: Staphylinidae) from the Early Cretaceous of China." url="https://doi.org/10.1163/1876312X-44022100" volume="44" year="2013">Cai and Huang 2013</bibRefCitation>
). The present fossil is also peculiar in the slender and elongate appendages. The present-day representatives of
<taxonomicName lsidName="" pageId="2" pageNumber="83" rank="tribe" tribe="Coprophilini">Coprophilini</taxonomicName>
lack such strongly formed, almost forward-projecting shoulders and the new genus has more slender antennae and palp, more slender tibiae, a procoxal cavity far removed from the pronotal margin, a prominently explanate pronotum, and the mesosternal process extending much more posteriorly. Two unusual traits for this subfamily are the posteriorly slightly incised elytral corners and the cylindrical, wide
<pageBreakToken pageId="3" pageNumber="84" start="start">apex</pageBreakToken>
of segment VII (not narrowing to the base of segment VIII), both features otherwise characteristic of the subfamily
<taxonomicName lsidName="" pageId="3" pageNumber="84" rank="subfamily" subfamily="Aleocharinae">Aleocharinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="84" type="systematic placement">
<paragraph pageId="3" pageNumber="84">Systematic placement.</paragraph>
<paragraph pageId="3" pageNumber="84">
The only feature that clearly unites the fossil with extant
<taxonomicName lsidName="" pageId="3" pageNumber="84" rank="tribe" tribe="Coprophilini">Coprophilini</taxonomicName>
is the lack of the well-developed second sternite. Beyond that, the head shape is reminiscent of
<taxonomicName class="Insecta" family="Staphylinidae" genus="Homalotrichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalotrichus" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">Homalotrichus</taxonomicName>
, while the pronotum bears some similarity to that of some
<taxonomicName class="Insecta" family="Staphylinidae" genus="Coprophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophilus" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">Coprophilus</taxonomicName>
(e.g.,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Coprophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophilus striatulus" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="striatulus">Coprophilus striatulus</taxonomicName>
(Fabricius, 1793) plus its close relatives) and to a lesser extent some
<taxonomicName class="Insecta" family="Staphylinidae" genus="Homalotrichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalotrichus" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">Homalotrichus</taxonomicName>
(e.g.,
<taxonomicName class="Insecta" family="Staphylinidae" genus="Homalotrichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalotrichus impressicollis" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="impressicollis">Homalotrichus impressicollis</taxonomicName>
Solier, 1849), but none of these are as explanate as in the fossil.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="84" type="etymology">
<paragraph pageId="3" pageNumber="84">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="84">
The new genus is named after Susan Golland, exhibition developer at the Field Museum of Natural History, Chicago, whom the first author met at 10:32am on 14 March 2018 in front of Crystal
<normalizedToken originalValue="Maiers">Maier's</normalizedToken>
office. The fossil specimen described here was shown to him by the second author later on the same day. The gender of the name is considered feminine.
</paragraph>
</subSubSection>
</treatment>
</document>