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<document id="93E66C5AA920814F916EA53E474F8589" ID-DOI="10.11646/phytotaxa.677.1.6" ID-ISSN="1179-3163" ID-Zenodo-Dep="14522098" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1734562578332" checkinUser="felipe" docAuthor="Yao, Gang, Luo, Shixiao, Huang, Jiuxiang &amp; Song, Zhuqiu" docDate="2024" docId="03E287DB48700170FF5FF908FA55B00E" docLanguage="en" docName="phytotaxa.677.1.095-101.pdf" docOrigin="Phytotaxa 677 (1)" docSource="https://doi.org/10.11646/phytotaxa.677.1.6" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="(Phyllanthodendron)" docType="key" docVersion="1" lastPageNumber="101" masterDocId="FFDBFFA348750176FFD7FFE1FFFEB126" masterDocTitle="Glochidion liangkwangense (Phyllanthaceae), a new species from Guangdong and Guangxi Provinces, China" masterLastPageNumber="101" masterPageNumber="95" pageNumber="100" updateTime="1741636156536" updateUser="juliana" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4AAEBEA3E46A56937A4C9B283964F16D">Glochidion liangkwangense (Phyllanthaceae), a new species from Guangdong and Guangxi Provinces, China</mods:title>
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<mods:namePart id="9EE420A57476BC868FB5D0BDE4023991">Yao, Gang</mods:namePart>
<mods:nameIdentifier id="90E025EB32029A8A08E2F4ABDF2BB256" type="ORCID">0000-0003-3101-8230</mods:nameIdentifier>
<mods:affiliation id="DD48BDD9DA3204ED2532C4F31261DFC9">College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510642, P. R. China</mods:affiliation>
<mods:nameIdentifier id="711233B741D8D5A2FF5F34F194E54B3B" type="email">gyao@scau.edu.cn</mods:nameIdentifier>
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<mods:namePart id="F97DF8AF141774234E5D963D854C91DD">Luo, Shixiao</mods:namePart>
<mods:nameIdentifier id="53B21E04EF10AA0D5C3090F027355D09" type="ORCID">0000-0002-1813-1944</mods:nameIdentifier>
<mods:affiliation id="CE22DB0F164D7B3469F6307442FE0AE5">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 516650, P. R. China</mods:affiliation>
<mods:nameIdentifier id="9A58DCA53FE0C51F3B49F14B5B21DB0B" type="email">luoshixiao@scbg.ac.cn</mods:nameIdentifier>
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<mods:namePart id="AF61BE584F19C1B11CA531FF35A59BCF">Huang, Jiuxiang</mods:namePart>
<mods:nameIdentifier id="BD7C6CC363CDC3F95B2C65EC0C4C04BC" type="ORCID">0000-0001-8576-5528</mods:nameIdentifier>
<mods:affiliation id="0B176AC2BF54DB5CC78877C1E5D1D8C9">College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou 510642, P. R. China</mods:affiliation>
<mods:nameIdentifier id="BD6221C20A0A8003920E258E6D34A33F" type="email">jxhuang@scau.edu.cn</mods:nameIdentifier>
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<mods:namePart id="A5816546503054F3904FA0D3F842EEB3">Song, Zhuqiu</mods:namePart>
<mods:nameIdentifier id="3A9544298199AAADAF0F7A3AA408183A" type="ORCID">0000-0002-7763-2431</mods:nameIdentifier>
<mods:affiliation id="AAA47D051D0F044ECA3F0C1AA2FE5ED2">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 516650, P. R. China</mods:affiliation>
<mods:nameIdentifier id="3BDAF7134F25DC7FA9FB290393928713" type="email">songzhuqiu@scbg.ac.cn</mods:nameIdentifier>
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<mods:title id="3A1364A45D3D16E49D044CE02718B867">Phytotaxa</mods:title>
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<mods:part id="804504D1E5822C74158C4189847DE3BD">
<mods:date id="FF20B2837FC0394A2A6ACCE1D0C57E94">2024</mods:date>
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<mods:number id="03A2A072133B543CAC277B01DE2EC735">2024-12-12</mods:number>
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<mods:number id="3B53E364B87798EA7100299D86273697">677</mods:number>
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<mods:url id="AC3A67DC1EA7128F24555AA055525802">https://doi.org/10.11646/phytotaxa.677.1.6</mods:url>
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<mods:identifier id="375EF99400D7B89E476AF4F222EED7A3" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="91FCCE271CEF546217A1C8054A1665FB" type="Zenodo-Dep">14522098</mods:identifier>
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<treatment id="03E287DB48700170FF5FF908FA55B00E" LSID="urn:lsid:plazi:treatment:03E287DB48700170FF5FF908FA55B00E" httpUri="http://treatment.plazi.org/id/03E287DB48700170FF5FF908FA55B00E" lastPageId="6" lastPageNumber="101" pageId="5" pageNumber="100">
<subSubSection id="C351654648700173FF5FF908FC90B625" box="[136,878,1768,1795]" pageId="5" pageNumber="100" type="nomenclature">
<paragraph id="8BF436CD48700173FF5FF908FC90B625" blockId="5.[136,878,1768,1795]" box="[136,878,1768,1795]" pageId="5" pageNumber="100">
Key to species of the subgenus
<taxonomicName id="4C4B4D4E48700173FE3EF909FD37B625" box="[489,713,1768,1795]" kingdom="Plantae" pageId="5" pageNumber="100" rank="subGenus" subGenus="Phyllanthodendron">
<emphasis id="B93FEADF48700173FE3EF909FD37B625" bold="true" box="[489,713,1768,1795]" italics="true" pageId="5" pageNumber="100">Phyllanthodendron</emphasis>
</taxonomicName>
in
<taxonomicName id="4C4B4D4E48700173FD3AF909FC90B625" box="[749,878,1768,1795]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="genus">
<emphasis id="B93FEADF48700173FD3AF909FC90B625" bold="true" box="[749,878,1768,1795]" italics="true" pageId="5" pageNumber="100">Glochidion</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
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<keyLead id="30BA95F848700173FF5FF8CCFA55B664" box="[136,1451,1837,1859]" pageId="5" pageNumber="100">
1. Sepals usually aristate at apex, acumens
<quantity id="4CB39B2848700173FD8EF8CCFD56B665" box="[601,680,1837,1859]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="100" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3‒5 mm</quantity>
long; capsules prominently 3-lobed ....................................................................
<emphasis id="B93FEADF48700173FA48F8CCFA55B664" bold="true" box="[1439,1451,1837,1858]" pageId="5" pageNumber="100">2</emphasis>
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</paragraph>
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<emphasis id="B93FEADF48700173FF5FF8ABFF6EB679" bold="true" box="[136,144,1866,1887]" pageId="5" pageNumber="100">-</emphasis>
Sepals usually rounded, acute or acuminate at apex; capsules subglobose or globose-triangular .....................................................
<emphasis id="B93FEADF48700173FA48F8ABFA55B679" bold="true" box="[1439,1451,1866,1887]" pageId="5" pageNumber="100">3</emphasis>
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<paragraph id="8BF436CD48700173FF5FF886FA55B65A" blockId="5.[136,1451,1837,2090]" box="[136,1451,1894,1917]" pageId="5" pageNumber="100">
<keyLead id="30BA95F848700173FF5FF886FA55B65A" box="[136,1451,1894,1917]" pageId="5" pageNumber="100">
2. Tree with succulent and swollen stem base; capsules ca.
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in diam .........................................................................
<taxonomicName id="4C4B4D4E48700173FAE2F886FA55B65A" box="[1333,1451,1894,1916]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="mirabilis">
<emphasis id="B93FEADF48700173FAE2F886FA55B65A" bold="true" box="[1333,1451,1894,1916]" italics="true" pageId="5" pageNumber="100">G. mirabilis</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BF436CD48700173FF5FF865FA55B6BF" blockId="5.[136,1451,1837,2090]" box="[136,1451,1923,1946]" pageId="5" pageNumber="100">
<keyLead id="30BA95F848700173FF5FF865FA55B6BF" box="[136,1451,1923,1946]" pageId="5" pageNumber="100">
<emphasis id="B93FEADF48700173FF5FF865FF6EB6BF" bold="true" box="[136,144,1924,1945]" pageId="5" pageNumber="100">-</emphasis>
Tree with non-succulent and non-swollen stem base; capsules
<quantity id="4CB39B2848700173FCFFF865FC7DB6BF" box="[808,899,1924,1945]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="5" pageNumber="100" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8‒10 mm</quantity>
in diam .......................................................
<taxonomicName id="4C4B4D4E48700173FAF6F865FA55B6BF" box="[1313,1451,1923,1945]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="kaweesakii">
<emphasis id="B93FEADF48700173FAF6F865FA55B6BF" bold="true" box="[1313,1451,1923,1945]" italics="true" pageId="5" pageNumber="100">G. kaweesakii</emphasis>
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<paragraph id="8BF436CD48700173FF5FF840FA55B690" blockId="5.[136,1451,1837,2090]" box="[136,1451,1952,1974]" pageId="5" pageNumber="100">
<keyLead id="30BA95F848700173FF5FF840FA55B690" box="[136,1451,1952,1974]" pageId="5" pageNumber="100">
3. Branchlets conspicuously winged .................................................................................................................................
<taxonomicName id="4C4B4D4E48700173FAF4F841FA55B690" box="[1315,1451,1952,1974]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="dunnianus">
<emphasis id="B93FEADF48700173FAF4F841FA55B690" bold="true" box="[1315,1451,1952,1974]" italics="true" pageId="5" pageNumber="100">G. dunnianus</emphasis>
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<paragraph id="8BF436CD48700173FF5FF85CFA55B6F4" blockId="5.[136,1451,1837,2090]" box="[136,1451,1981,2003]" pageId="5" pageNumber="100">
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<emphasis id="B93FEADF48700173FF5FF85CFF6EB6F4" bold="true" box="[136,144,1981,2002]" pageId="5" pageNumber="100">-</emphasis>
Branchlets terete .................................................................................................................................................................................
<emphasis id="B93FEADF48700173FA48F85CFA55B6F4" bold="true" box="[1439,1451,1981,2002]" pageId="5" pageNumber="100">4</emphasis>
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</paragraph>
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<paragraph id="8BF436CD48700173FF5FF83BFA55B6C9" blockId="5.[136,1451,1837,2090]" box="[136,1451,2010,2032]" pageId="5" pageNumber="100">
<keyLead id="30BA95F848700173FF5FF83BFA55B6C9" box="[136,1451,2010,2032]" pageId="5" pageNumber="100">
4. Leaf blades usually less than
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long .............................................................................................................................................
<emphasis id="B93FEADF48700173FA48F83BFA55B6C9" bold="true" box="[1439,1451,2010,2031]" pageId="5" pageNumber="100">5</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BF436CD48700173FF5FF816FA55B92A" blockId="5.[136,1451,1837,2090]" box="[136,1451,2039,2061]" pageId="5" pageNumber="100">
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- Leaf blade mostly longer than
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...................................................................................................................................................
<emphasis id="B93FEADF48700173FA48F816FA55B92A" bold="true" box="[1439,1451,2039,2060]" pageId="5" pageNumber="100">6</emphasis>
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<paragraph id="8BF436CD48700173FF5FF7F2FA55B90F" blockId="5.[136,1451,1837,2090]" box="[136,1451,2067,2090]" pageId="5" pageNumber="100">
<keyLead id="30BA95F848700173FF5FF7F2FA55B90F" box="[136,1451,2067,2090]" pageId="5" pageNumber="100">
5. Leaf blades 1.6‒2.0 cm long, and
<quantity id="4CB39B2848700173FDCEF7F5FD7AB90F" box="[537,644,2068,2089]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="5" pageNumber="100" unit="cm" value="0.8" valueMax="0.9" valueMin="0.7">0.7‒0.9 cm</quantity>
wide; capsules ca.
<quantity id="4CB39B2848700173FCE9F7F2FC79B90F" box="[830,903,2067,2089]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="100" unit="mm" value="4.5">4.5 mm</quantity>
in diam....................................................
<taxonomicName id="4C4B4D4E48700173FAC6F7F5FA55B90F" box="[1297,1451,2067,2089]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="dongmoense">
<emphasis id="B93FEADF48700173FAC6F7F5FA55B90F" bold="true" box="[1297,1451,2067,2089]" italics="true" pageId="5" pageNumber="100">G. dongmoense</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF436CD48730170FF5FFF7EFA55B193" blockId="6.[136,1451,159,296]" box="[136,1451,159,181]" pageId="6" pageNumber="101">
<keyLead id="30BA95F848730170FF5FFF7EFA55B193" box="[136,1451,159,181]" pageId="6" pageNumber="101">
<emphasis id="B93FEADF48730170FF5FFF7EFF6EB192" bold="true" box="[136,144,159,180]" pageId="6" pageNumber="101">-</emphasis>
Leaf blades usually 1.0‒
<quantity id="4CB39B2848730170FE6AFF7EFE00B193" box="[445,510,159,181]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="101" unit="cm" value="3.5">3.5 cm</quantity>
long, and
<quantity id="4CB39B2848730170FDB3FF7EFD31B193" box="[612,719,159,181]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="6" pageNumber="101" unit="cm" value="1.0" valueMax="1.5" valueMin="0.5">0.5‒1.5 cm</quantity>
wide; capsules
<quantity id="4CB39B2848730170FCBFFF7EFC49B193" box="[872,951,159,181]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="6" pageNumber="101" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5‒8 mm</quantity>
in diam.....................................
<taxonomicName id="4C4B4D4E48730170FB32FF7EFA55B193" authorityName="Croizat" authorityYear="1942" baseAuthorityName="Handel-Mazzetti" baseAuthorityYear="1931" box="[1253,1451,159,181]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="101" phylum="Tracheophyta" rank="species" species="anthopotamicum">
<emphasis id="B93FEADF48730170FB32FF7EFA55B193" bold="true" box="[1253,1451,159,181]" italics="true" pageId="6" pageNumber="101">G. anthopotamicum</emphasis>
</taxonomicName>
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</paragraph>
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<keyLead id="30BA95F848730170FF5FFF5DFA55B1C8" pageId="6" pageNumber="101">
6. Sepals caudate or narrowly acuminate at apex; lateral veins absent in sepals; capsules
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in diam., triangular-globose.......... ..............................................................................................................................................................................
<taxonomicName id="4C4B4D4E48730170FB3AFF38FA55B1C8" box="[1261,1451,216,238]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="101" phylum="Tracheophyta" rank="species" species="liangkwangense">
<emphasis id="B93FEADF48730170FB3AFF38FA55B1C8" bold="true" box="[1261,1451,216,238]" italics="true" pageId="6" pageNumber="101">G. liangkwangense</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF436CD48730170FF5FFF17FA55B00E" blockId="6.[136,1451,159,296]" pageId="6" pageNumber="101">
<keyLead id="30BA95F848730170FF5FFF17FA55B00E" pageId="6" pageNumber="101">
- Sepals acute to shortly acuminate at apex; lateral veins present in sepals; capsules
<quantity id="4CB39B2848730170FC3EFF17FBBAB02D" box="[1001,1092,246,267]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="6" pageNumber="101" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
in diam., subglobose ......................... .................................................................................................................................................................................
<taxonomicName id="4C4B4D4E48730170FB28FEF3FA55B00E" box="[1279,1451,274,296]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="6" pageNumber="101" phylum="Tracheophyta" rank="species" species="yangchunense">
<emphasis id="B93FEADF48730170FB28FEF3FA55B00E" bold="true" box="[1279,1451,274,296]" italics="true" pageId="6" pageNumber="101">G. yangchunense</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>

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<mods:title id="42CADE7BDFC0864CA181E5D304DF98C8">Glochidion liangkwangense (Phyllanthaceae), a new species from Guangdong and Guangxi Provinces, China</mods:title>
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<mods:namePart id="59EA2F1385FF36F2BB795B9A372132C3">Yao, Gang</mods:namePart>
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<mods:namePart id="C39AA1BCD4B30D8AEE2A319DC2BF69F3">Luo, Shixiao</mods:namePart>
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<mods:affiliation id="EE41CDCF452CA5D59B1068F8B556F85A">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 516650, P. R. China</mods:affiliation>
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<mods:namePart id="DCF90A04BDC6C18501E7E66FDFB3AD81">Huang, Jiuxiang</mods:namePart>
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<mods:namePart id="F9BB204DEB010C21E4C77CACCBD5FA6A">Song, Zhuqiu</mods:namePart>
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<mods:affiliation id="BFFA0385C446CD28273E5266E0E89773">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 516650, P. R. China</mods:affiliation>
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<treatment id="03E287DB48740173FF5FFD3EFAEEB79D" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D" httpUri="http://treatment.plazi.org/id/03E287DB48740173FF5FFD3EFAEEB79D" lastPageId="5" lastPageNumber="100" pageId="1" pageNumber="96">
<subSubSection id="C351654648740177FF5FFD3EFD27B3D3" box="[136,729,735,757]" pageId="1" pageNumber="96" type="nomenclature">
<paragraph id="8BF436CD48740177FF5FFD3EFD27B3D3" blockId="1.[136,1452,735,1082]" box="[136,729,735,757]" pageId="1" pageNumber="96">
<heading id="D0BC81A148740177FF5FFD3EFD27B3D3" box="[136,729,735,757]" pageId="1" pageNumber="96">
<taxonomicName id="4C4B4D4E48740177FF5FFD3EFD74B3D3" authority="Gang Yao" authorityName="Gang Yao" box="[136,650,735,757]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="96" phylum="Tracheophyta" rank="species" species="liangkwangense">
<emphasis id="B93FEADF48740177FF5FFD3EFE63B3D3" bold="true" box="[136,413,735,757]" italics="true" pageId="1" pageNumber="96">Glochidion liangkwangense</emphasis>
Gang Yao &amp; Z.Q. Song,
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<taxonomicNameLabel id="A20C57A448740177FD58FD01FD27B3D3" box="[655,729,736,757]" pageId="1" pageNumber="96">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C351654648740177FD09FD01FC9EB3D3" box="[734,864,736,758]" pageId="1" pageNumber="96" type="description">
<paragraph id="8BF436CD48740177FD09FD01FC9EB3D3" blockId="1.[136,1452,735,1082]" box="[734,864,736,758]" pageId="1" pageNumber="96">
(
<figureCitation id="13702A4848740177FD33FD01FCBEB3D3" box="[740,832,736,758]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="1" pageNumber="96">Figures 1</figureCitation>
,
<figureCitation id="13702A4848740177FC9BFD01FCA6B3D3" box="[844,856,736,757]" captionStart="FIGURE 3" captionStartId="4.[136,229,1188,1210]" captionTargetBox="[153,1430,206,1160]" captionTargetId="figure-29@4.[151,1436,206,1160]" captionTargetPageId="4" captionText="FIGURE 3. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A. Holotype (Gang Yao &amp; Zhu-Qiu Song 2024010, IBSC1025532). B. Isotype (IBSC1025533)." figureDoi="http://doi.org/10.5281/zenodo.14522104" httpUri="https://zenodo.org/record/14522104/files/figure.png" pageId="1" pageNumber="96">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C351654648740177FF5FFCE5FD3EB2E8" pageId="1" pageNumber="96" type="diagnosis">
<paragraph id="8BF436CD48740177FF5FFCE5FD3EB2E8" blockId="1.[136,1452,735,1082]" pageId="1" pageNumber="96">
<emphasis id="B93FEADF48740177FF5FFCE5FF15B23F" bold="true" box="[136,235,772,793]" italics="true" pageId="1" pageNumber="96">Diagnosis</emphasis>
:—
<taxonomicName id="4C4B4D4E48740177FEDEFCE5FDE6B23F" box="[265,536,772,793]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="96" phylum="Tracheophyta" rank="species" species="liangkwangense">
<emphasis id="B93FEADF48740177FEDEFCE5FDE6B23F" box="[265,536,772,793]" italics="true" pageId="1" pageNumber="96">Glochidion liangkwangense</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4B4D4E48740177FD07FCE5FC6EB23F" authorityName="Croizat" authorityYear="1942" baseAuthorityName="Handel-Mazzetti" baseAuthorityYear="1931" box="[720,912,772,793]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="96" phylum="Tracheophyta" rank="species" species="anthopotamicum">
<emphasis id="B93FEADF48740177FD07FCE5FC6EB23F" box="[720,912,772,793]" italics="true" pageId="1" pageNumber="96">G. anthopotamicum</emphasis>
</taxonomicName>
in morphology, but differs from the latter by its lateral veins in leaf blades elevated abaxially (especially for leaf blades grow at the lower part of branchlets) (
<emphasis id="B93FEADF48740177FB7FFCC9FB42B21B" box="[1192,1212,808,829]" italics="true" pageId="1" pageNumber="96">vs</emphasis>
. not elevated abaxially); sepals erect and usually extend outward (
<emphasis id="B93FEADF48740177FD9DFCADFDA0B247" box="[586,606,844,865]" italics="true" pageId="1" pageNumber="96">vs</emphasis>
. spreaded or even reflexed at the upper part, and jointly formed a urceolate shape at the lower part), glabrous or subglabrous along the midrib (vs. densely pubescent); sepals of female flowers in 2 series, but different in shape and size (
<emphasis id="B93FEADF48740177FE83FC75FE96B28F" box="[340,360,916,937]" italics="true" pageId="1" pageNumber="96">vs</emphasis>
. in 2 series but subequal in both of shape and size); disc segments short and spatulate to slightly T-shaped (
<emphasis id="B93FEADF48740177FAB6FC75FA8BB28F" box="[1377,1397,916,937]" italics="true" pageId="1" pageNumber="96">vs</emphasis>
. long and linear); capsules triangular-globose (
<emphasis id="B93FEADF48740177FD91FC59FDA4B2EB" box="[582,602,952,973]" italics="true" pageId="1" pageNumber="96">vs</emphasis>
. globose).
</paragraph>
</subSubSection>
<subSubSection id="C351654648740177FF5FFC3DFEE2B51F" pageId="1" pageNumber="96" type="materials_examined">
<paragraph id="8BF436CD48740177FF5FFC3DFEE2B51F" blockId="1.[136,1452,735,1082]" pageId="1" pageNumber="96">
<materialsCitation id="3B233C9048740177FF5FFC3DFBD0B530" collectingDate="2024-04-02" collectionCode="IBSC" collectorName="Gang Yao &amp; Zhu-Qiu Song" country="China" county="Type" elevation="66" latitude="22.185535" location="Kongtongyan" longLatPrecision="1" longitude="111.74618" municipality="Yangchun Hsien" pageId="1" pageNumber="96" specimenCode="IBSC1025532" specimenCount="1" stateProvince="Guangdong Province" typeStatus="holotype">
Type:—
<collectingCountry id="F35C765D48740177FF00FC3DFED8B2D7" box="[215,294,988,1009]" name="China" pageId="1" pageNumber="96">CHINA</collectingCountry>
.
<collectingRegion id="498FF82F48740177FEE1FC3DFDF5B2D4" box="[310,523,988,1010]" country="China" name="Guangdong" pageId="1" pageNumber="96">Guangdong Province</collectingRegion>
,
<collectingMunicipality id="6B90ACB748740177FDCCFC3DFD3DB2D4" box="[539,707,988,1010]" pageId="1" pageNumber="96">Yangchun Hsien</collectingMunicipality>
,
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, in limestone hills, elevation ca.
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,
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,
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,
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<collectingDate id="EFB1E9E548740177FEB0FBE1FE23B533" box="[359,477,1024,1046]" pageId="1" pageNumber="96" value="2024-04-02">2April 2024</collectingDate>
</date>
,
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<collectorName id="26BE531B48740177FE33FBE1FDBEB533" box="[484,576,1024,1045]" pageId="1" pageNumber="96">Gang Yao</collectorName>
&amp;
<collectorName id="26BE531B48740177FD8EFBE1FD23B533" box="[601,733,1024,1045]" pageId="1" pageNumber="96">Zhu-Qiu Song</collectorName>
2024010
</emphasis>
(
<typeStatus id="54F0886F48740177FCE9FBE1FC69B530" box="[830,919,1024,1046]" pageId="1" pageNumber="96" type="holotype">holotype</typeStatus>
:
<specimenCode id="DBED9EB648740177FC49FBE1FBD4B530" box="[926,1066,1024,1046]" pageId="1" pageNumber="96">IBSC1025532</specimenCode>
;
</materialsCitation>
<materialsCitation id="3B233C9048740177FBE6FBE1FEE6B51F" collectingDate="2024-04-02" collectionCode="IBSC" collectorName="Gang Yao &amp; Zhu-Qiu Song" country="China" county="Type" elevation="66" latitude="22.185535" location="Kongtongyan" longLatPrecision="1" longitude="111.74618" municipality="Yangchun Hsien" pageId="1" pageNumber="96" specimenCode="IBSC1025533, IBSC1025534" specimenCount="1" stateProvince="Guangdong" typeStatus="isotype">
<typeStatus id="54F0886F48740177FBE6FBE1FB7CB530" box="[1073,1154,1024,1046]" pageId="1" pageNumber="96" type="isotype">isotypes</typeStatus>
,
<specimenCode id="DBED9EB648740177FB5EFBE1FAEAB530" box="[1161,1300,1024,1046]" pageId="1" pageNumber="96">IBSC1025533</specimenCode>
,
<specimenCode id="DBED9EB648740177FACCFBE1FA56B530" box="[1307,1448,1024,1046]" pageId="1" pageNumber="96">IBSC1025534</specimenCode>
;
<figureCitation id="13702A4848740177FF6AFBC5FEF1B51F" box="[189,271,1060,1082]" captionStart="FIGURE 3" captionStartId="4.[136,229,1188,1210]" captionTargetBox="[153,1430,206,1160]" captionTargetId="figure-29@4.[151,1436,206,1160]" captionTargetPageId="4" captionText="FIGURE 3. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A. Holotype (Gang Yao &amp; Zhu-Qiu Song 2024010, IBSC1025532). B. Isotype (IBSC1025533)." figureDoi="http://doi.org/10.5281/zenodo.14522104" httpUri="https://zenodo.org/record/14522104/files/figure.png" pageId="1" pageNumber="96">Figure 3</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C351654648740177FF5FFB89FE37B609" pageId="1" pageNumber="96" type="description">
<paragraph id="8BF436CD48740177FF5FFB89FE37B609" blockId="1.[136,1452,1128,2091]" pageId="1" pageNumber="96">
Description:—Shrubs,
<quantity id="4CB39B2848740177FE5DFB89FE04B5A5" box="[394,506,1128,1155]" metricMagnitude="0" metricUnit="m" metricValue="1.65" metricValueMax="2.5" metricValueMin="0.8" pageId="1" pageNumber="96" unit="m" value="1.65" valueMax="2.5" valueMin="0.8">0.8–2.5 m</quantity>
tall, erect, monoecious; stem gray-brown; branches terete, subglabrous to pubescent, green when young. Stipules short,
<quantity id="4CB39B2848740177FDD0FB6DFD9AB581" box="[519,612,1164,1191]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="96" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
long, usually triangular at lower part of branches and narrowly triangular at upper part of branches. Petiole
<quantity id="4CB39B2848740177FE30FB51FDBBB5ED" box="[487,581,1200,1227]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="1" pageNumber="96" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long, subglabrous. Leaves simple, alternate, distichous; leaf blades 2.0–7.0 × 1.0–
<quantity id="4CB39B2848740177FF6EFB35FEF6B5C9" box="[185,264,1236,1263]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="96" unit="cm" value="3.5">3.5 cm</quantity>
, usually coriaceous, broadly ovate or elliptic, usually glabrous, or puberulent on the vines abaxially when young; leaf blades at upper part of branches usually smaller, lateral veins in 5–7 pairs; leaf blades at lower part of branches usually larger, 3.0–7.5 × 2.0–5.0 cm, lateral veins in 5–9 pairs; leaf blades margin entire, slightly revolute, apex acute to acuminate, base broadly truncate to sub-rounded, or rarely cordiform, usually glaucous abaxially in individuals habituated at lower elevations; midrib and lateral veins flattened above, slightly elevated below (especially the midrib), usually anastomosing near margin. Male flowers clustered axillary, usually inserted at the lower part of branches; pedicles 3.0–
<quantity id="4CB39B2848740177FE63FA4DFDF2B4E1" box="[436,524,1452,1479]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="96" unit="mm" value="4.5">4.5 mm</quantity>
long, glabrous or sparsely pubescent; sepals 5, 3.0–3.5 × 1.0–
<quantity id="4CB39B2848740177FB0BFA4DFACBB4E1" box="[1244,1333,1452,1479]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="96" unit="mm" value="1.5">1.5 mm</quantity>
, erect and usually extend outward, imbricate, yellowish-white, midrib elevated on abaxial surface, narrowly triangular or ovatetriangular and usually caudate at apex, glabrous or rarely sparsely pubescent; petals absent; disc segments 5, free, Tshaped, or sub-spatulate but rounded or truncate at apex, slightly greenish yellow; stamens 3, filaments connate into a terete column, anthers erect, dehiscing longitudinally, connectives apiculate. Female flowers usually solidary axillary and inserted at the upper part of branches; sepals 6, erect and usually extend outward, imbricate, in 2 series, inner ones ovate and with a caudate at apex, outer ones narrowly triangular, usually glabrous; petals absent; disc segments 6, free, extended outside, T-shaped or slightly expand at apex, slightly greenish yellow; ovary 3-locular, pubescent at least at the upper part; styles 3; stigmas evidently bifid. Capsules
<quantity id="4CB39B2848740177FC49F92DFC00B7C1" box="[926,1022,1740,1767]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="96" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
in diam., globose-triangular, smooth, brownish when mature, fruiting pedicels
<quantity id="4CB39B2848740177FD8BF911FD2EB62D" box="[604,720,1776,1803]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="1" pageNumber="96" unit="mm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 mm</quantity>
long. Seeds 2–2.6 ×
<quantity id="4CB39B2848740177FC13F911FBC7B62D" box="[964,1081,1776,1803]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="1" pageNumber="96" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 mm</quantity>
, obscurely laterally compressed, dark brownish when mature.
</paragraph>
</subSubSection>
<subSubSection id="C351654648740174FF6AF8D9FA6BB002" lastPageId="2" lastPageNumber="97" pageId="1" pageNumber="96" type="materials_examined">
<paragraph id="8BF436CD48740174FF6AF8D9FA6BB002" blockId="1.[136,1452,1128,2091]" lastBlockId="2.[136,1452,158,293]" lastPageId="2" lastPageNumber="97" pageId="1" pageNumber="96">
<materialsCitation id="3B233C9048740177FF6AF8D9FD74B651" collectionCode="IBSC" collectorName="Hua-Shing Chiu" country="China" location="Chuanbu Town" municipality="Luoding Hsien" pageId="1" pageNumber="96" specimenCode="IBSC0325784" specimenCount="1" stateProvince="Guangdong Province" typeStatus="paratype">
<emphasis id="B93FEADF48740177FF6AF8D9FD09B674" box="[189,759,1848,1875]" italics="true" pageId="1" pageNumber="96">
<emphasis id="B93FEADF48740177FF6AF8D9FD20B674" bold="true" box="[189,734,1848,1875]" italics="true" pageId="1" pageNumber="96">
Additional specimens examined (
<typeStatus id="54F0886F48740177FD8BF8D9FD33B674" box="[604,717,1848,1874]" pageId="1" pageNumber="96" type="paratype">paratypes</typeStatus>
):
</emphasis>
—
</emphasis>
<collectingCountry id="F35C765D48740177FD20F8D9FCAAB674" box="[759,852,1848,1874]" name="China" pageId="1" pageNumber="96">CHINA</collectingCountry>
:
<collectingRegion id="498FF82F48740177FCBEF8D9FB9AB674" box="[873,1124,1848,1874]" country="China" name="Guangdong" pageId="1" pageNumber="96">Guangdong Province</collectingRegion>
,
<collectingMunicipality id="6B90ACB748740177FBACF8D9FACEB674" box="[1147,1328,1848,1874]" pageId="1" pageNumber="96">Luoding Hsien</collectingMunicipality>
,
<location id="8E94601648740177FA91F8D9FF34B651" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648740177FA91F8D9FF34B651" country="China" municipality="Luoding Hsien" name="Chuanbu Town" pageId="1" pageNumber="96" stateProvince="Guangdong Province">Chuanbu Town</location>
,
<emphasis id="B93FEADF48740177FF0EF8BDFE37B650" box="[217,457,1884,1910]" italics="true" pageId="1" pageNumber="96">
<collectorName id="26BE531B48740177FF0EF8BDFE6DB650" box="[217,403,1884,1910]" pageId="1" pageNumber="96">Hua-Shing Chiu</collectorName>
561
</emphasis>
(
<specimenCode id="DBED9EB648740177FE0CF8BDFD7DB651" box="[475,643,1884,1911]" pageId="1" pageNumber="96">IBSC0325784</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B233C9048740177FD4FF8BDFCEDB6BD" collectingDate="1991-11-24" collectionCode="IIBSC" collectorName="Nian Liu" country="China" county="Yangchun Hsien" elevation="300" location="Kongdongyan" municipality="Kongtongyan" pageId="1" pageNumber="96" specimenCode="IIBSC0325785" specimenCount="1" stateProvince="Guangdong Province">
<collectingCounty id="62954E4148740177FD4FF8BDFCA9B650" box="[664,855,1884,1910]" pageId="1" pageNumber="96">Yangchun Hsien</collectingCounty>
,
<collectingMunicipality id="6B90ACB748740177FCB0F8BCFBFCB651" box="[871,1026,1885,1911]" pageId="1" pageNumber="96">Kongtongyan</collectingMunicipality>
,
<location id="8E94601648740177FBC6F8BDFB4DB650" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648740177FBC6F8BDFB4DB650" box="[1041,1203,1884,1910]" country="China" county="Yangchun Hsien" municipality="Kongtongyan" name="Kongdongyan" pageId="1" pageNumber="96">Kongdongyan</location>
,
<elevation id="0066D1FE48740177FB16F8BDFA7EB651" box="[1217,1408,1884,1911]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="96" unit="m" value="300.0">
elevation
<quantity id="4CB39B2848740177FAE4F8BDFA7EB651" box="[1331,1408,1884,1911]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="96" unit="m" value="300.0">300 m</quantity>
</elevation>
,
<date id="FFF5100D48740177FA58F8BDFEBAB6BC" pageId="1" pageNumber="96" value="1991-11-24">
<collectingDate id="EFB1E9E548740177FA58F8BDFEBAB6BC" pageId="1" pageNumber="96" value="1991-11-24">24 November 1991</collectingDate>
</date>
,
<emphasis id="B93FEADF48740177FE83F860FDB6B6BC" box="[340,584,1920,1946]" italics="true" pageId="1" pageNumber="96">
<collectorName id="26BE531B48740177FE83F860FE45B6BC" box="[340,443,1921,1946]" pageId="1" pageNumber="96">Nian Liu</collectorName>
et al. 3052
</emphasis>
(
<specimenCode id="DBED9EB648740177FD8CF861FCF5B6BD" box="[603,779,1920,1947]" pageId="1" pageNumber="96">IIBSC0325785</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B233C9048740177FCF5F861FE11B699" collectingDate="1956-09-10" collectionCode="IBK, IBSC" collectorName="Chi Wang" country="China" county="Yangchun Hsien" elevation="300" location="Yangchun Hsien" municipality="Kongtongyan" pageId="1" pageNumber="96" specimenCode="IBK00168390, IBSC0325786" specimenCount="1" stateProvince="Guangdong Province">
<location id="8E94601648740177FCF5F861FC1CB6BC" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648740177FCF5F861FC1CB6BC" box="[802,994,1920,1946]" country="China" name="Yangchun Hsien" pageId="1" pageNumber="96">Yangchun Hsien</location>
,
<date id="FFF5100D48740177FC26F861FB29B6BD" box="[1009,1239,1920,1947]" pageId="1" pageNumber="96" value="1956-09-10">
<collectingDate id="EFB1E9E548740177FC26F861FB29B6BD" box="[1009,1239,1920,1947]" pageId="1" pageNumber="96" value="1956-09-10">10 September 1956</collectingDate>
</date>
,
<emphasis id="B93FEADF48740177FB30F861FA55B6BC" box="[1255,1451,1920,1946]" italics="true" pageId="1" pageNumber="96">
<collectorName id="26BE531B48740177FB30F861FAA4B6BC" box="[1255,1370,1920,1946]" pageId="1" pageNumber="96">Chi Wang</collectorName>
42121
</emphasis>
(
<specimenCode id="DBED9EB648740177FF47F845FECAB698" box="[144,308,1956,1982]" pageId="1" pageNumber="96">IBK00168390</specimenCode>
,
<specimenCode id="DBED9EB648740177FE96F845FE19B699" box="[321,487,1956,1983]" pageId="1" pageNumber="96">IBSC0325786</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B233C9048740177FE2DF845FCA8B6C5" collectingDate="1935-10-16" collectingDateMax="1957-07-03" collectingDateMin="1935-10-16" collectionCode="IBK, IBSC" collectorName="Chi Wang" country="China" county="Yangchun Hsien" elevation="300" location="Kongtongyan" municipality="Kongtongyan" pageId="1" pageNumber="96" specimenCode="IBK00168392, IBSC0325782, IBSC0325780, IBSC0325781, IBSC0325783" specimenCount="1" stateProvince="Guangxi">
<collectingCounty id="62954E4148740177FE2DF845FD49B698" box="[506,695,1956,1982]" pageId="1" pageNumber="96">Yangchun Hsien</collectingCounty>
,
<collectingMunicipality id="6B90ACB748740177FD13F844FCA1B699" box="[708,863,1957,1983]" pageId="1" pageNumber="96">Kongtongyan</collectingMunicipality>
,
<date id="FFF5100D48740177FCBBF845FBD5B699" box="[876,1067,1956,1983]" pageId="1" pageNumber="96" value="1935-10-16">
<collectingDate id="EFB1E9E548740177FCBBF845FBD5B699" box="[876,1067,1956,1983]" pageId="1" pageNumber="96" value="1935-10-16">16 October 1935</collectingDate>
</date>
<emphasis id="B93FEADF48740177FBE3F845FB0CB698" box="[1076,1266,1956,1982]" italics="true" pageId="1" pageNumber="96">
<collectorName id="26BE531B48740177FBE3F845FB5AB698" box="[1076,1188,1956,1982]" pageId="1" pageNumber="96">Chi Wang</collectorName>
38481
</emphasis>
(
<specimenCode id="DBED9EB648740177FAD5F845FA59B698" box="[1282,1447,1956,1982]" pageId="1" pageNumber="96">IBK00168392</specimenCode>
,
<specimenCode id="DBED9EB648740177FF5FF829FED5B6C5" box="[136,299,1992,2019]" pageId="1" pageNumber="96">IBSC0325782</specimenCode>
,
<specimenCode id="DBED9EB648740177FEEBF829FE21B6C5" box="[316,479,1992,2019]" pageId="1" pageNumber="96">IBSC0325780</specimenCode>
,
<specimenCode id="DBED9EB648740177FE26F829FD6AB6C5" box="[497,660,1992,2019]" pageId="1" pageNumber="96">IBSC0325781</specimenCode>
,
<specimenCode id="DBED9EB648740177FD72F829FCB4B6C5" box="[677,842,1992,2019]" pageId="1" pageNumber="96">IBSC0325783</specimenCode>
).
</materialsCitation>
<materialsCitation id="3B233C9048740177FCB4F829FCEFB920" collectingDate="1957-07-03" collectingDateMax="1957-07-03" collectingDateMin="1935-10-16" collectionCode="IBSC, KUN" collectorName="Yin-Kun Li" country="China" county="Yangchun Hsien" elevation="300" location="Du'an Hsien" municipality="Kongtongyan" pageId="1" pageNumber="96" specimenCode="IBSC0325794, KUN1294369" specimenCount="1" stateProvince="Guangxi Province">
<collectingRegion id="498FF82F48740177FCB4F829FBC8B6C4" box="[867,1078,1992,2018]" country="China" name="Guangxi" pageId="1" pageNumber="96">Guangxi Province</collectingRegion>
,
<location id="8E94601648740177FB9FF829FB21B6C4" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648740177FB9FF829FB21B6C4" box="[1096,1247,1992,2018]" country="China" county="Yangchun Hsien" municipality="Kongtongyan" name="Du'an Hsien" pageId="1" pageNumber="96" stateProvince="Guangxi Province">Du’an Hsien</location>
,
<location id="8E94601648740177FB27F829FAAEB6C4" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648740177FB27F829FAAEB6C4" box="[1264,1360,1992,2018]" country="China" county="Yangchun Hsien" municipality="Kongtongyan" name="Shangfu" pageId="1" pageNumber="96" stateProvince="Guangxi Province">Shangfu</location>
,
<date id="FFF5100D48740177FAB5F829FF3DB921" pageId="1" pageNumber="96" value="1957-07-03">
<collectingDate id="EFB1E9E548740177FAB5F829FF3DB921" pageId="1" pageNumber="96" value="1957-07-03">3 July 1957</collectingDate>
</date>
,
<emphasis id="B93FEADF48740177FF19F80CFE59B920" box="[206,423,2028,2054]" italics="true" pageId="1" pageNumber="96">
<collectorName id="26BE531B48740177FF19F80CFEB6B920" box="[206,328,2029,2054]" pageId="1" pageNumber="96">Yin-Kun Li</collectorName>
P01570
</emphasis>
(
<specimenCode id="DBED9EB648740177FE62F80DFDA4B921" box="[437,602,2028,2055]" pageId="1" pageNumber="96">IBSC0325794</specimenCode>
,
<specimenCode id="DBED9EB648740177FDB2F80DFCF7B920" box="[613,777,2028,2054]" pageId="1" pageNumber="96">KUN1294369</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B233C9048740177FCCBF80DFD11B90D" collectingDate="1935-08-26" collectionCode="IBK, IBSC" collectorName="Sik-Pang Ko" country="China" county="Jingxi Hsien" elevation="300" location="Longyangshan" municipality="Longyangshan" pageId="1" pageNumber="96" specimenCode="IBK00168381, IBK00168387, IBSC0325793" specimenCount="1" stateProvince="Guangxi Province">
<collectingCounty id="62954E4148740177FCCBF80DFC52B920" box="[796,940,2028,2054]" pageId="1" pageNumber="96">Jingxi Hsien</collectingCounty>
,
<location id="8E94601648740177FC6FF80DFB9EB920" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648740177FC6FF80DFB9EB920" box="[952,1120,2028,2054]" country="China" county="Jingxi Hsien" municipality="Longyangshan" name="Longyangshan" pageId="1" pageNumber="96">Longyangshan</location>
, 26 Augst 1935,
<emphasis id="B93FEADF48740177FACAF80DFF30B90C" italics="true" pageId="1" pageNumber="96">
<collectorName id="26BE531B48740177FACAF80DFA52B920" box="[1309,1452,2028,2054]" pageId="1" pageNumber="96">Sik-Pang Ko</collectorName>
55629
</emphasis>
(
<specimenCode id="DBED9EB648740177FF08F7F1FE7AB90C" box="[223,388,2064,2090]" pageId="1" pageNumber="96">IBK00168381</specimenCode>
,
<specimenCode id="DBED9EB648740177FE46F7F1FDCAB90C" box="[401,564,2064,2090]" pageId="1" pageNumber="96">IBK00168387</specimenCode>
,
<specimenCode id="DBED9EB648740177FD96F7F1FD19B90D" box="[577,743,2064,2091]" pageId="1" pageNumber="96">IBSC0325793</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B233C9048740174FD2BF7F1FC9BB19E" collectingDate="2013-10-15" collectingDateMax="1957-07-03" collectingDateMin="1935-10-16" collectionCode="GXMG, GXMI" collectorName="Lingyun Expedition" country="China" county="Jingxi Hsien" elevation="956" lastPageId="2" lastPageNumber="97" location="Lingyun Hsien" municipality="Longyangshan" pageId="1" pageNumber="96" specimenCode="GXMG0117289, GXMI063810" specimenCount="1" stateProvince="Guangxi Province">
<location id="8E94601648740177FD2BF7F1FC56B90C" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648740177FD2BF7F1FC56B90C" box="[764,936,2064,2090]" country="China" name="Lingyun Hsien" pageId="1" pageNumber="96">Lingyun Hsien</location>
,
<elevation id="0066D1FE48740177FC61F7F1FB8FB90D" box="[950,1137,2064,2091]" metricMagnitude="2" metricUnit="m" metricValue="9.559999999999999" pageId="1" pageNumber="96" unit="m" value="956.0">
elevation
<quantity id="4CB39B2848740177FBF2F7F1FB8FB90D" box="[1061,1137,2064,2091]" metricMagnitude="2" metricUnit="m" metricValue="9.559999999999999" pageId="1" pageNumber="96" unit="m" value="956.0">956 m</quantity>
</elevation>
,
<date id="FFF5100D48740177FBA8F7F1FABCB90C" box="[1151,1346,2064,2091]" pageId="1" pageNumber="96" value="2013-10-15">
<collectingDate id="EFB1E9E548740177FBA8F7F1FABCB90C" box="[1151,1346,2064,2091]" pageId="1" pageNumber="96" value="2013-10-15">15 October 2013</collectingDate>
</date>
,
<emphasis id="B93FEADF48740174FA87F7F0FE20B19E" italics="true" lastPageId="2" lastPageNumber="97" pageId="1" pageNumber="96">
<collectorName id="26BE531B48740174FA87F7F0FEFFB19E" lastPageId="2" lastPageNumber="97" pageId="1" pageNumber="96">Lingyun Expedition</collectorName>
451027131015013
</emphasis>
(
<specimenCode id="DBED9EB648770174FE38FF7FFD54B19E" box="[495,682,158,184]" pageId="2" pageNumber="97">GXMG0117289</specimenCode>
,
<specimenCode id="DBED9EB648770174FD6EFF7FFCA3B19E" box="[697,861,158,184]" pageId="2" pageNumber="97">GXMI063810</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B233C9048770174FCA4FF7FFCE6B1FB" collectingDate="2018-08-10" collectionCode="IBK" collectorName="Liujiang Expedition" country="China" county="Jingxi Hsien" elevation="126" location="Liujiang Hsien" municipality="Liujiang Hsien" pageId="2" pageNumber="97" specimenCode="IBK00425489" specimenCount="1" stateProvince="Guangxi Province">
<collectingMunicipality id="6B90ACB748770174FCA4FF7FFBDEB19E" box="[883,1056,158,184]" pageId="2" pageNumber="97">Liujiang Hsien</collectingMunicipality>
,
<elevation id="0066D1FE48770174FBF9FF7FFB12B19F" box="[1070,1260,158,185]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="2" pageNumber="97" unit="m" value="126.0">
elevation
<quantity id="4CB39B2848770174FB48FF7FFB12B19F" box="[1183,1260,158,185]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="2" pageNumber="97" unit="m" value="126.0">126 m</quantity>
</elevation>
, 10 Augst 2018,
<emphasis id="B93FEADF48770174FF5FFF22FDA2B1FA" box="[136,604,194,220]" italics="true" pageId="2" pageNumber="97">
<collectorName id="26BE531B48770174FF5FFF22FE98B1FA" box="[136,358,194,220]" pageId="2" pageNumber="97">Liujiang Expedition</collectorName>
450221180810025LY
</emphasis>
(
<specimenCode id="DBED9EB648770174FDBEFF23FCEFB1FB" box="[617,785,194,221]" pageId="2" pageNumber="97">IBK00425489</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B233C9048770174FCF4FF23FD2EB026" collectingDate="2011-09-01" collectingDateMax="1957-07-03" collectingDateMin="1935-10-16" collectionCode="IBK" collectorName="Jin-Cai Yang" country="China" county="Jingxi Hsien" elevation="220" location="Longhushan" municipality="Long'an Hsien" pageId="2" pageNumber="97" specimenCode="IBK00306774" specimenCount="1" stateProvince="Guangxi Province">
<collectingMunicipality id="6B90ACB748770174FCF4FF23FC32B1FA" box="[803,972,194,220]" pageId="2" pageNumber="97">Long’an Hsien</collectingMunicipality>
,
<location id="8E94601648770174FC01FF23FB9DB1FA" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648770174FC01FF23FB9DB1FA" box="[982,1123,194,220]" country="China" municipality="Long'an Hsien" name="Longhushan" pageId="2" pageNumber="97">Longhushan</location>
,
<date id="FFF5100D48770174FBB9FF23FAC4B1FA" box="[1134,1338,194,220]" pageId="2" pageNumber="97" value="2011-09-01">
<collectingDate id="EFB1E9E548770174FBB9FF23FAC4B1FA" box="[1134,1338,194,220]" pageId="2" pageNumber="97" value="2011-09-01">1 September 2011</collectingDate>
</date>
,
<elevation id="0066D1FE48770174FA92FF23FF2FB027" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="97" unit="m" value="220.0">
elevation
<quantity id="4CB39B2848770174FF5FFF07FF2FB027" box="[136,209,230,257]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="97" unit="m" value="220.0">220 m</quantity>
</elevation>
,
<emphasis id="B93FEADF48770174FF0AFF06FDECB026" box="[221,530,230,257]" italics="true" pageId="2" pageNumber="97">
<collectorName id="26BE531B48770174FF0AFF06FE8EB026" box="[221,368,231,257]" pageId="2" pageNumber="97">Jin-Cai Yang</collectorName>
et al. LH0459
</emphasis>
(
<specimenCode id="DBED9EB648770174FDF6FF07FD36B026" box="[545,712,230,256]" pageId="2" pageNumber="97">IBK00306774</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B233C9048770174FD0CFF07FDC7B002" collectingDate="1958-10-04" collectingDateMax="1957-07-03" collectingDateMin="1935-10-16" collectionCode="IBK, KUN" collectorName="Huan-Qi Li" country="China" county="Jingxi Hsien" elevation="220" location="Paotai Mountain" municipality="Longzhou Hsien" pageId="2" pageNumber="97" specimenCode="IBK00168394, KUN1294370" specimenCount="1" stateProvince="Guangxi Province">
<collectingMunicipality id="6B90ACB748770174FD0CFF07FC66B026" box="[731,920,230,256]" pageId="2" pageNumber="97">Longzhou Hsien</collectingMunicipality>
,
<location id="8E94601648770174FC73FF07FB9FB026" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648770174FC73FF07FB9FB026" box="[932,1121,230,256]" country="China" municipality="Longzhou Hsien" name="Paotai Mountain" pageId="2" pageNumber="97">Paotai Mountain</location>
,
<date id="FFF5100D48770174FBBBFF07FAE0B027" box="[1132,1310,230,257]" pageId="2" pageNumber="97" value="1958-10-04">
<collectingDate id="EFB1E9E548770174FBBBFF07FAE0B027" box="[1132,1310,230,257]" pageId="2" pageNumber="97" value="1958-10-04">4 October 1958</collectingDate>
</date>
,
<emphasis id="B93FEADF48770174FAFEFF06FF30B002" italics="true" pageId="2" pageNumber="97">
<collectorName id="26BE531B48770174FAFEFF06FA55B026" box="[1321,1451,231,257]" pageId="2" pageNumber="97">Huan-Qi Li</collectorName>
40064
</emphasis>
(
<specimenCode id="DBED9EB648770174FF0AFEEBFE7FB002" box="[221,385,266,292]" pageId="2" pageNumber="97">IBK00168394</specimenCode>
,
<specimenCode id="DBED9EB648770174FE5AFEEBFDCFB002" box="[397,561,266,292]" pageId="2" pageNumber="97">KUN1294370</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B233C9048770174FD94FEEBFA6FB002" box="[579,1425,266,293]" collectingDate="2024-09-07" collectingDateMax="1957-07-03" collectingDateMin="1935-10-16" collectionCode="IBSC" collectorName="Gang Yao" country="China" county="Jingxi Hsien" elevation="220" location="Nandan Hsien" municipality="Longzhou Hsien" pageId="2" pageNumber="97" specimenCount="1" stateProvince="Guangxi Province">
<location id="8E94601648770174FD94FEEBFD1BB002" LSID="urn:lsid:plazi:treatment:03E287DB48740173FF5FFD3EFAEEB79D:8E94601648770174FD94FEEBFD1BB002" box="[579,741,266,292]" country="China" name="Nandan Hsien" pageId="2" pageNumber="97">Nandan Hsien</location>
,
<date id="FFF5100D48770174FD26FEEAFC3EB002" box="[753,960,266,293]" pageId="2" pageNumber="97" value="2024-09-07">
<collectingDate id="EFB1E9E548770174FD26FEEAFC3EB002" box="[753,960,266,293]" pageId="2" pageNumber="97" value="2024-09-07">7 September 2024</collectingDate>
</date>
,
<emphasis id="B93FEADF48770174FC1CFEEAFAC8B002" box="[971,1334,266,293]" italics="true" pageId="2" pageNumber="97">
<collectorName id="26BE531B48770174FC1CFEEAFBC7B002" box="[971,1081,267,293]" pageId="2" pageNumber="97">Gang Yao</collectorName>
YGGXTE2024090701
</emphasis>
(
<collectionCode id="ED5AAE0848770174FA92FEEBFA77B002" box="[1349,1417,266,292]" pageId="2" pageNumber="97">IBSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF34664548770174FF5FF898FD2FB938" ID-DOI="http://doi.org/10.5281/zenodo.14522100" ID-Zenodo-Dep="14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="2" pageNumber="97" startId="2.[136,229,1913,1935]" targetBox="[201,1384,314,1882]" targetPageId="2" targetType="figure">
<paragraph id="8BF436CD48770174FF5FF898FD2FB938" blockId="2.[136,1452,1913,2079]" pageId="2" pageNumber="97">
<emphasis id="B93FEADF48770174FF5FF898FF01B6A8" bold="true" box="[136,255,1913,1935]" pageId="2" pageNumber="97">FIGURE 1.</emphasis>
<taxonomicName id="4C4B4D4E48770174FED2F898FD87B6A9" authority="Gang Yao" authorityName="Gang Yao" box="[261,633,1913,1935]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="97" phylum="Tracheophyta" rank="species" species="liangkwangense">
<emphasis id="B93FEADF48770174FED2F898FDEDB6A8" box="[261,531,1913,1934]" italics="true" pageId="2" pageNumber="97">Glochidion liangkwangense</emphasis>
Gang Yao
</taxonomicName>
&amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song.
</paragraph>
</caption>
<caption id="DF34664548760175FF5FF89CFD13B6D9" ID-DOI="http://doi.org/10.5281/zenodo.14522102" ID-Zenodo-Dep="14522102" httpUri="https://zenodo.org/record/14522102/files/figure.png" pageId="3" pageNumber="98" startId="3.[136,229,1917,1939]" targetBox="[157,1429,187,1886]" targetPageId="3" targetType="figure">
<paragraph id="8BF436CD48760175FF5FF89CFD13B6D9" blockId="3.[136,1451,1917,2047]" pageId="3" pageNumber="98">
<emphasis id="B93FEADF48760175FF5FF89CFF01B6B4" bold="true" box="[136,255,1917,1939]" pageId="3" pageNumber="98">FIGURE 2.</emphasis>
<taxonomicName id="4C4B4D4E48760175FED1F89CFDE5B6B4" authorityName="Croizat" authorityYear="1942" baseAuthorityName="Handel-Mazzetti" baseAuthorityYear="1931" box="[262,539,1917,1938]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="98" phylum="Tracheophyta" rank="species" species="anthopotamicum">
<emphasis id="B93FEADF48760175FED1F89CFDE5B6B4" box="[262,539,1917,1938]" italics="true" pageId="3" pageNumber="98">Glochidion anthopotamicum</emphasis>
</taxonomicName>
(Hand.-Mazz) R.W. Bouman. A. Holotype (
<emphasis id="B93FEADF48760175FC19F89CFEFDB691" italics="true" pageId="3" pageNumber="98">Handel-Mazzetti, H.R.E. von, Iter sinense (1914- 1918) 10377</emphasis>
, WU0046581). B. Habit. C. Male flowers from the holotype (lateral view). D. Male flowers, disc segments (white arrows indicated). E. Male flowers. F. Female flower (lateral view). G. Female flower (top view), disc segments (white arrows indicated), stigma (yellow arrows indicated). Photographs (B, D–G) by Gang Yao.
</paragraph>
</caption>
<caption id="DF34664548710172FF5FFB45FE8BB5F8" ID-DOI="http://doi.org/10.5281/zenodo.14522104" ID-Zenodo-Dep="14522104" httpUri="https://zenodo.org/record/14522104/files/figure.png" pageId="4" pageNumber="99" startId="4.[136,229,1188,1210]" targetBox="[153,1430,206,1160]" targetPageId="4" targetType="figure">
<paragraph id="8BF436CD48710172FF5FFB45FE8BB5F8" blockId="4.[136,1452,1188,1246]" pageId="4" pageNumber="99">
<emphasis id="B93FEADF48710172FF5FFB45FF00B59F" bold="true" box="[136,254,1188,1210]" pageId="4" pageNumber="99">FIGURE 3.</emphasis>
<taxonomicName id="4C4B4D4E48710172FED3FB45FD89B59F" authority="Gang Yao" authorityName="Gang Yao" box="[260,631,1188,1209]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="99" phylum="Tracheophyta" rank="species" species="liangkwangense">
<emphasis id="B93FEADF48710172FED3FB45FDECB59F" box="[260,530,1188,1209]" italics="true" pageId="4" pageNumber="99">Glochidion liangkwangense</emphasis>
Gang Yao
</taxonomicName>
&amp; Z.Q. Song. A. Holotype (
<emphasis id="B93FEADF48710172FC5CFB45FB15B59F" box="[907,1259,1188,1209]" italics="true" pageId="4" pageNumber="99">Gang Yao &amp; Zhu-Qiu Song 2024010</emphasis>
, IBSC1025532). B. Isotype (IBSC1025533).
</paragraph>
</caption>
<subSubSection id="C351654648710172FF6AFAEDFC8BB448" pageId="4" pageNumber="99" type="biology_ecology">
<paragraph id="8BF436CD48710172FF6AFAEDFB8AB46D" blockId="4.[136,1452,1292,2075]" pageId="4" pageNumber="99">
<emphasis id="B93FEADF48710172FF6AFAEDFE2FB401" bold="true" box="[189,465,1292,1319]" italics="true" pageId="4" pageNumber="99">Habitat and distribution</emphasis>
:—
<taxonomicName id="4C4B4D4E48710172FE22FAEDFCD2B400" box="[501,812,1292,1318]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="99" phylum="Tracheophyta" rank="species" species="liangkwangense">
<emphasis id="B93FEADF48710172FE22FAEDFCD2B400" box="[501,812,1292,1318]" italics="true" pageId="4" pageNumber="99">Glochidion liangkwangense</emphasis>
</taxonomicName>
is found in multiple localities in
<collectingRegion id="498FF82F48710172FB42FAEDFAE6B400" box="[1173,1304,1292,1318]" country="China" name="Guangdong" pageId="4" pageNumber="99">Guangdong</collectingRegion>
and
<collectingRegion id="498FF82F48710172FA9CFAEDFA55B400" box="[1355,1451,1292,1318]" country="China" name="Guangxi" pageId="4" pageNumber="99">Guangxi</collectingRegion>
Provinces,
<collectingCountry id="F35C765D48710172FED3FAD1FEB9B46C" box="[260,327,1328,1354]" name="China" pageId="4" pageNumber="99">China</collectingCountry>
(
<figureCitation id="13702A4848710172FE81FAD1FE47B46C" box="[342,441,1328,1354]" captionStart="FIGURE 4" captionStartId="5.[136,229,1592,1614]" captionTargetBox="[161,1426,612,1555]" captionTargetId="figure-248@5.[151,1436,602,1565]" captionTargetPageId="5" captionText="FIGURE 4. Distribution map of Glochidion liangkwangense Gang Yao &amp; Z.Q. Song (red stars indicated)." figureDoi="http://doi.org/10.5281/zenodo.14522106" httpUri="https://zenodo.org/record/14522106/files/figure.png" pageId="4" pageNumber="99">Figure 4</figureCitation>
). It grows in limestone hills at elevations from
<quantity id="4CB39B2848710172FC1DFAD1FBFDB46D" box="[970,1027,1328,1355]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="99" unit="m" value="50.0">50 m</quantity>
to
<specimenCount id="9D4DFD4448710172FBF1FAD1FB8AB46D" box="[1062,1140,1328,1355]" count="960" pageId="4" pageNumber="99" type="generic">
<quantity id="4CB39B2848710172FBF1FAD1FB8EB46D" box="[1062,1136,1328,1355]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="4" pageNumber="99" unit="m" value="960.0">960 m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="8BF436CD48710172FF6AFAB5FC8BB448" blockId="4.[136,1452,1292,2075]" box="[189,885,1364,1391]" pageId="4" pageNumber="99">
<emphasis id="B93FEADF48710172FF6AFAB5FEC6B449" bold="true" box="[189,312,1364,1391]" italics="true" pageId="4" pageNumber="99">Phenology</emphasis>
:—Flowering and fruiting from April to December.
</paragraph>
</subSubSection>
<subSubSection id="C351654648710172FF6AFA99FEB1B490" pageId="4" pageNumber="99" type="etymology">
<paragraph id="8BF436CD48710172FF6AFA99FEB1B490" blockId="4.[136,1452,1292,2075]" pageId="4" pageNumber="99">
<emphasis id="B93FEADF48710172FF6AFA99FEC6B4B5" bold="true" box="[189,312,1400,1427]" italics="true" pageId="4" pageNumber="99">Etymology</emphasis>
:—
<taxonomicName id="4C4B4D4E48710172FE8BFA99FD67B4B4" box="[348,665,1400,1426]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="99" phylum="Tracheophyta" rank="species" species="liangkwangense">
<emphasis id="B93FEADF48710172FE8BFA99FD67B4B4" box="[348,665,1400,1426]" italics="true" pageId="4" pageNumber="99">Glochidion liangkwangense</emphasis>
</taxonomicName>
is named after its distribution regions, in
<collectingRegion id="498FF82F48710172FB5DFA99FAF3B4B4" box="[1162,1293,1400,1426]" country="China" name="Guangdong" pageId="4" pageNumber="99">Guangdong</collectingRegion>
and
<collectingRegion id="498FF82F48710172FA9CFA99FA55B4B4" box="[1355,1451,1400,1426]" country="China" name="Guangxi" pageId="4" pageNumber="99">Guangxi</collectingRegion>
Provinces,
<collectingCountry id="F35C765D48710172FED3FA7DFEB5B490" box="[260,331,1436,1462]" name="China" pageId="4" pageNumber="99">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C351654648710172FF6AFA21FCC0B4FC" box="[189,830,1472,1499]" pageId="4" pageNumber="99" type="vernacular_names">
<paragraph id="8BF436CD48710172FF6AFA21FCC0B4FC" blockId="4.[136,1452,1292,2075]" box="[189,830,1472,1499]" pageId="4" pageNumber="99">
<emphasis id="B93FEADF48710172FF6AFA21FE7AB4FC" box="[189,388,1472,1499]" italics="true" pageId="4" pageNumber="99">
<emphasis id="B93FEADF48710172FF6AFA21FE95B4FC" bold="true" box="[189,363,1472,1499]" italics="true" pageId="4" pageNumber="99">
<taxonomicName id="4C4B4D4E48710172FF6AFA21FE9AB4FC" box="[189,356,1472,1499]" class="Aves" family="Anatidae" genus="Chinese" kingdom="Animalia" order="Anseriformes" pageId="4" pageNumber="99" phylum="Chordata" rank="species" species="name">Chinese name</taxonomicName>
:
</emphasis>
—
</emphasis>
Liang Guang Zhu Zi Mu (两ḞȕƗ†)
</paragraph>
</subSubSection>
<subSubSection id="C351654648710172FF6AFA05FE3BB760" pageId="4" pageNumber="99" type="conservation">
<paragraph id="8BF436CD48710172FF6AFA05FE3BB760" blockId="4.[136,1452,1292,2075]" pageId="4" pageNumber="99">
<emphasis id="B93FEADF48710172FF6AFA05FDCEB4D8" bold="true" box="[189,560,1508,1535]" italics="true" pageId="4" pageNumber="99">Preliminary conservation status</emphasis>
:—It is considered here as LC according to current information and the IUCN Red List criteria (
<bibRefCitation id="EFDA4B3C48710172FE9AF9E9FC90B704" author="IUCN Standards &amp; Petitions Committee" box="[333,878,1544,1570]" pageId="4" pageNumber="99" pagination="1 - 60" publicationUrl="http://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf" refId="ref3445" refString="IUCN Standards and Petitions Committee. (2019) IUCN Standards and Petitions Committee. Standards and Petitions Committee, 1 (August), 1-60. [http://intranet.iucn.org/webfiles/doc/SSC/RedList/RedListGuidelines.pdf]" title="IUCN Standards and Petitions Committee. Standards and Petitions Committee, 1 (August)" type="url" year="2019">IUCN Standards and Petitions Committee, 2019</bibRefCitation>
) for its wide distribution regions and multiple wild populations and individuals.
</paragraph>
</subSubSection>
<subSubSection id="C351654648710173FF6AF9B1FAEEB79D" lastPageId="5" lastPageNumber="100" pageId="4" pageNumber="99" type="discussion">
<paragraph id="8BF436CD48710173FF6AF9B1FB5DB363" blockId="4.[136,1452,1292,2075]" lastBlockId="5.[136,1452,158,581]" lastPageId="5" lastPageNumber="100" pageId="4" pageNumber="99">
<emphasis id="B93FEADF48710172FF6AF9B1FDF8B74D" bold="true" box="[189,518,1616,1643]" italics="true" pageId="4" pageNumber="99">Morphological relationships</emphasis>
:—Morphologically, the new species is characterized by its sepals that are caudate at apex and have elevated midrib abaxially (
<figureCitation id="13702A4848710172FD57F995FD0CB7A8" box="[640,754,1652,1678]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="4" pageNumber="99">Figure 1F</figureCitation>
), spatulate to slightly T-shaped glands in flowers (
<figureCitation id="13702A4848710172FAE5F995FF48B794" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="4" pageNumber="99">Figure 1E, G, I</figureCitation>
), as well as extended connectives of anthers. All these characters indicate that it is a member of the traditionally circumscribed
<taxonomicName id="4C4B4D4E48710172FEFCF95DFDFCB7F0" box="[299,514,1724,1750]" pageId="4" pageNumber="100" rank="subGenus" subGenus="Phyllanthodendron">
<emphasis id="B93FEADF48710172FEFCF95DFDFCB7F0" box="[299,514,1724,1750]" italics="true" pageId="4" pageNumber="99">Phyllanthodendron</emphasis>
</taxonomicName>
. Furthermore, the lack of a pedicelled clustered fascicle indicates that the new species belongs to the subgenus
<taxonomicName id="4C4B4D4E48710172FE45F901FD97B7DC" box="[402,617,1760,1786]" pageId="4" pageNumber="99" rank="subGenus" subGenus="Phyllanthodendron">
<emphasis id="B93FEADF48710172FE45F901FD97B7DC" box="[402,617,1760,1786]" italics="true" pageId="4" pageNumber="99">Phyllanthodendron</emphasis>
</taxonomicName>
in the newly circumscribed
<taxonomicName id="4C4B4D4E48710172FC4BF901FBE4B7DC" box="[924,1050,1760,1786]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="99" phylum="Tracheophyta" rank="genus">
<emphasis id="B93FEADF48710172FC4BF901FBE4B7DC" box="[924,1050,1760,1786]" italics="true" pageId="4" pageNumber="99">Glochidion</emphasis>
</taxonomicName>
, as defined by
<bibRefCitation id="EFDA4B3C48710172FB6BF900FA56B7DC" author="Bouman, R. W. &amp; Kebler, P. J. &amp; Telford, I. R. &amp; Bruhl, J. J. &amp; Strijk, J. S. &amp; Saunders, R. M. &amp; Esser, H. J. &amp; Falcon-Hi-dalgo, B. &amp; van Welzen, P. C." box="[1212,1448,1760,1787]" journalOrPublisher="Phytotaxa" pageId="4" pageNumber="99" pagination="1 - 100" part="540" publicationUrl="https://doi.org/10.11646/phytotaxa.540.1.1" refId="ref3379" refString="Bouman, R. W., Kebler, P. J., Telford, I. R., Bruhl, J. J., Strijk, J. S., Saunders, R. M., Esser, H. J., Falcon-Hi-dalgo, B. &amp; van Welzen, P. C. (2022) A revised phylogenetic classification of tribe Phyllantheae (Phyllanthaceae). Phytotaxa 540 (1): 1-100. https://doi.org/10.11646/phytotaxa.540.1.1" title="A revised phylogenetic classification of tribe Phyllantheae (Phyllanthaceae)" type="book chapter" year="2022">
Bouman
<emphasis id="B93FEADF48710172FAC8F900FAABB7DC" box="[1311,1365,1760,1786]" italics="true" pageId="4" pageNumber="99">et al.</emphasis>
(2022)
</bibRefCitation>
. In
<bibRefCitation id="EFDA4B3C48710172FF70F8E4FE62B638" author="Bouman, R. W. &amp; Kebler, P. J. &amp; Telford, I. R. &amp; Bruhl, J. J. &amp; Strijk, J. S. &amp; Saunders, R. M. &amp; Esser, H. J. &amp; Falcon-Hi-dalgo, B. &amp; van Welzen, P. C." box="[167,412,1796,1823]" journalOrPublisher="Phytotaxa" pageId="4" pageNumber="99" pagination="1 - 100" part="540" publicationUrl="https://doi.org/10.11646/phytotaxa.540.1.1" refId="ref3379" refString="Bouman, R. W., Kebler, P. J., Telford, I. R., Bruhl, J. J., Strijk, J. S., Saunders, R. M., Esser, H. J., Falcon-Hi-dalgo, B. &amp; van Welzen, P. C. (2022) A revised phylogenetic classification of tribe Phyllantheae (Phyllanthaceae). Phytotaxa 540 (1): 1-100. https://doi.org/10.11646/phytotaxa.540.1.1" title="A revised phylogenetic classification of tribe Phyllantheae (Phyllanthaceae)" type="book chapter" year="2022">
Bouman
<emphasis id="B93FEADF48710172FED9F8E4FEB9B638" box="[270,327,1796,1822]" italics="true" pageId="4" pageNumber="99">et al.</emphasis>
(2022)
</bibRefCitation>
, five species were accepted in the subgenus
<taxonomicName id="4C4B4D4E48710172FC44F8E5FB94B638" box="[915,1130,1796,1822]" pageId="4" pageNumber="99" rank="subGenus" subGenus="Phyllanthodendron">
<emphasis id="B93FEADF48710172FC44F8E5FB94B638" box="[915,1130,1796,1822]" italics="true" pageId="4" pageNumber="99">Phyllanthodendron</emphasis>
</taxonomicName>
, and the sixth species of the subgenus, viz.
<taxonomicName id="4C4B4D4E48710172FEE6F8C8FE0AB664" box="[305,500,1832,1859]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="99" phylum="Tracheophyta" rank="species" species="yangchunense">
<emphasis id="B93FEADF48710172FEE6F8C8FE0AB664" box="[305,500,1832,1859]" italics="true" pageId="4" pageNumber="99">G. yangchunense</emphasis>
</taxonomicName>
Z.Q. Song &amp; Gang Yao (2024: 230), was described recently from Yangchun Hsien,
<collectingRegion id="498FF82F48710172FF5FF8ADFE86B640" box="[136,376,1868,1894]" country="China" name="Guangdong" pageId="4" pageNumber="99">Guangdong Province</collectingRegion>
,
<collectingCountry id="F35C765D48710172FE52F8ADFE32B640" box="[389,460,1868,1894]" name="China" pageId="4" pageNumber="99">China</collectingCountry>
. The new species most resembles
<taxonomicName id="4C4B4D4E48710172FC99F8ACFBD2B640" authorityName="Croizat" authorityYear="1942" baseAuthorityName="Handel-Mazzetti" baseAuthorityYear="1931" box="[846,1068,1868,1895]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="99" phylum="Tracheophyta" rank="species" species="anthopotamicum">
<emphasis id="B93FEADF48710172FC99F8ACFBD2B640" box="[846,1068,1868,1895]" italics="true" pageId="4" pageNumber="99">G. anthopotamicum</emphasis>
</taxonomicName>
in morphology, and specimens of the new species are deposited mainly under the latter name in most herbariums in
<collectingCountry id="F35C765D48710172FBE5F891FB87B6AC" box="[1074,1145,1904,1930]" name="China" pageId="4" pageNumber="99">China</collectingCountry>
. However, results from the present study show that the two species could be easily distinguished by characters of flowers and capsules. The new species has erect sepals that are usually extend outward (
<figureCitation id="13702A4848710172FCD7F859FC72B6F4" box="[768,908,1976,2002]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="4" pageNumber="99">Figure 1E‒J</figureCitation>
), glabrous or subglabrous along midrib of sepals (
<figureCitation id="13702A4848710172FF47F83DFEDFB6D0" box="[144,289,2012,2038]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="4" pageNumber="99">Figure 1H, J</figureCitation>
), 2 series of sepals of female flowers different in shape and size (
<figureCitation id="13702A4848710172FBDCF83DFB69B6D0" box="[1035,1175,2012,2038]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="4" pageNumber="99">Figure 1E, I</figureCitation>
), spatulate to slightly Tshaped disc segments in flowers (
<figureCitation id="13702A4848710172FDD6F7E1FD71B93C" box="[513,655,2048,2074]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="4" pageNumber="99">Figure 1G, I</figureCitation>
), styles usually swollen at apex, capsules that are triangular-globose in shape (
<figureCitation id="13702A4848700173FF00FF7FFE91B19E" box="[215,367,158,184]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="5" pageNumber="100">Figure 1K, L</figureCitation>
). In contrast,
<taxonomicName id="4C4B4D4E48700173FDDFFF7EFD19B19E" authorityName="Croizat" authorityYear="1942" baseAuthorityName="Handel-Mazzetti" baseAuthorityYear="1931" box="[520,743,158,185]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="anthopotamicum">
<emphasis id="B93FEADF48700173FDDFFF7EFD19B19E" box="[520,743,158,185]" italics="true" pageId="5" pageNumber="100">G. anthopotamicum</emphasis>
</taxonomicName>
has sepals that are spreading or even reflexed at the upper part but jointly formed an urceolate shape at the lower part (
<figureCitation id="13702A4848700173FCDBFF23FC58B1FA" box="[780,934,194,220]" captionStart="FIGURE 2" captionStartId="3.[136,229,1917,1939]" captionTargetBox="[157,1429,187,1886]" captionTargetId="figure-18@3.[157,1429,187,1890]" captionTargetPageId="3" captionText="FIGURE 2. Glochidion anthopotamicum (Hand.-Mazz) R.W. Bouman. A. Holotype (Handel-Mazzetti, H.R.E. von, Iter sinense (1914- 1918) 10377, WU0046581). B. Habit. C. Male flowers from the holotype (lateral view). D. Male flowers, disc segments (white arrows indicated). E. Male flowers. F. Female flower (lateral view). G. Female flower (top view), disc segments (white arrows indicated), stigma (yellow arrows indicated). Photographs (B, D–G) by Gang Yao." figureDoi="http://doi.org/10.5281/zenodo.14522102" httpUri="https://zenodo.org/record/14522102/files/figure.png" pageId="5" pageNumber="100">Figure 2C‒G</figureCitation>
), midrib of sepals that are densely pubescent (
<figureCitation id="13702A4848700173FF47FF07FEDAB026" box="[144,292,230,256]" captionStart="FIGURE 2" captionStartId="3.[136,229,1917,1939]" captionTargetBox="[157,1429,187,1886]" captionTargetId="figure-18@3.[157,1429,187,1890]" captionTargetPageId="3" captionText="FIGURE 2. Glochidion anthopotamicum (Hand.-Mazz) R.W. Bouman. A. Holotype (Handel-Mazzetti, H.R.E. von, Iter sinense (1914- 1918) 10377, WU0046581). B. Habit. C. Male flowers from the holotype (lateral view). D. Male flowers, disc segments (white arrows indicated). E. Male flowers. F. Female flower (lateral view). G. Female flower (top view), disc segments (white arrows indicated), stigma (yellow arrows indicated). Photographs (B, D–G) by Gang Yao." figureDoi="http://doi.org/10.5281/zenodo.14522102" httpUri="https://zenodo.org/record/14522102/files/figure.png" pageId="5" pageNumber="100">Figure 2E, F</figureCitation>
), 2 series of sepals of female flowers subequivalent in both of shape and size (
<figureCitation id="13702A4848700173FB70FF07FADFB026" box="[1191,1313,230,256]" captionStart="FIGURE 2" captionStartId="3.[136,229,1917,1939]" captionTargetBox="[157,1429,187,1886]" captionTargetId="figure-18@3.[157,1429,187,1890]" captionTargetPageId="3" captionText="FIGURE 2. Glochidion anthopotamicum (Hand.-Mazz) R.W. Bouman. A. Holotype (Handel-Mazzetti, H.R.E. von, Iter sinense (1914- 1918) 10377, WU0046581). B. Habit. C. Male flowers from the holotype (lateral view). D. Male flowers, disc segments (white arrows indicated). E. Male flowers. F. Female flower (lateral view). G. Female flower (top view), disc segments (white arrows indicated), stigma (yellow arrows indicated). Photographs (B, D–G) by Gang Yao." figureDoi="http://doi.org/10.5281/zenodo.14522102" httpUri="https://zenodo.org/record/14522102/files/figure.png" pageId="5" pageNumber="100">Figure 2G</figureCitation>
), linear disc segments in flowers (
<figureCitation id="13702A4848700173FEA1FEEBFDF0B002" box="[374,526,266,292]" captionStart="FIGURE 2" captionStartId="3.[136,229,1917,1939]" captionTargetBox="[157,1429,187,1886]" captionTargetId="figure-18@3.[157,1429,187,1890]" captionTargetPageId="3" captionText="FIGURE 2. Glochidion anthopotamicum (Hand.-Mazz) R.W. Bouman. A. Holotype (Handel-Mazzetti, H.R.E. von, Iter sinense (1914- 1918) 10377, WU0046581). B. Habit. C. Male flowers from the holotype (lateral view). D. Male flowers, disc segments (white arrows indicated). E. Male flowers. F. Female flower (lateral view). G. Female flower (top view), disc segments (white arrows indicated), stigma (yellow arrows indicated). Photographs (B, D–G) by Gang Yao." figureDoi="http://doi.org/10.5281/zenodo.14522102" httpUri="https://zenodo.org/record/14522102/files/figure.png" pageId="5" pageNumber="100">Figure 2D, G</figureCitation>
), styles not swollen at apex (
<figureCitation id="13702A4848700173FC98FEEBFC39B002" box="[847,967,266,292]" captionStart="FIGURE 2" captionStartId="3.[136,229,1917,1939]" captionTargetBox="[157,1429,187,1886]" captionTargetId="figure-18@3.[157,1429,187,1890]" captionTargetPageId="3" captionText="FIGURE 2. Glochidion anthopotamicum (Hand.-Mazz) R.W. Bouman. A. Holotype (Handel-Mazzetti, H.R.E. von, Iter sinense (1914- 1918) 10377, WU0046581). B. Habit. C. Male flowers from the holotype (lateral view). D. Male flowers, disc segments (white arrows indicated). E. Male flowers. F. Female flower (lateral view). G. Female flower (top view), disc segments (white arrows indicated), stigma (yellow arrows indicated). Photographs (B, D–G) by Gang Yao." figureDoi="http://doi.org/10.5281/zenodo.14522102" httpUri="https://zenodo.org/record/14522102/files/figure.png" pageId="5" pageNumber="100">Figure 2G</figureCitation>
), and capsules that are globose in shape (
<bibRefCitation id="EFDA4B3C48700173FA46FEEBFF3AB06E" author="Li" firstAuthor="Li" pageId="5" pageNumber="100" pagination="1 - 60" refId="ref3598" refString="(August), 1 - 60. [http: // intranet. iucn. org / webfiles / doc / SSC / RedList / RedListGuidelines. pdf] Li, P. T. (1994) Phyllanthodendron. In: Li, P. T. (ed.) Flora Repubulicae Popularis Sinicae, Tomus 44 (1). Science Press, Beijing, 116 -" type="book chapter" year="1994">Li 1994</bibRefCitation>
). Additionally, the new species is also similar to
<taxonomicName id="4C4B4D4E48700173FD38FECEFC4FB06E" box="[751,945,302,329]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="yangchunense">
<emphasis id="B93FEADF48700173FD38FECEFC4FB06E" box="[751,945,302,329]" italics="true" pageId="5" pageNumber="100">G. yangchunense</emphasis>
</taxonomicName>
morphologically in having sepals of flowers that are extend outward and T-shaped disc segments in flowers (Song and Yao 2024). However, the new species differs from the latter by its smaller leaves at the upper part of branchlets and larger leaves at the lower part of branchlets (
<figureCitation id="13702A4848700173FF47FE7BFEFCB092" box="[144,258,410,436]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="5" pageNumber="100">Figure 1C</figureCitation>
) (vs. larger leaves at the upper part of branchlets and smaller leaves at the lower part of branchlets), sepals caudate or narrowly acuminate at apex (
<figureCitation id="13702A4848700173FD99FE5FFD24B0FE" box="[590,730,446,472]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="5" pageNumber="100">Figure 1E‒J</figureCitation>
) (vs. sepals acute to shortly acuminate at apex), lateral veins in sepals are absent (
<figureCitation id="13702A4848700173FE84FE03FE00B0DA" box="[339,510,482,508]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="5" pageNumber="100">Figure 1E, G, I</figureCitation>
) (vs. present), smaller capsules (
<quantity id="4CB39B2848700173FCBAFE03FC35B0DB" box="[877,971,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="5" pageNumber="100" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
in diameter) that are triangular-globose in polar view (
<figureCitation id="13702A4848700173FEDBFDE7FE60B306" box="[268,414,518,544]" captionStart="FIGURE 1" captionStartId="2.[136,229,1913,1935]" captionTargetBox="[201,1384,314,1882]" captionTargetId="figure-85@2.[201,1385,314,1885]" captionTargetPageId="2" captionText="FIGURE 1. Glochidion liangkwangense Gang Yao &amp; Z.Q. Song. A, Habit. B. Detail of the apex of branches. C. Branches. D. Stipules. E. Pistillate flower, top view. F. Inflorescence, female flowers (red arrows indicated) and male flowers (white arrows indicated). G. Male flower (top view), disc segments (red arrow indicated). H. Male flower (lateral view), extended connective of anthers (white arrow indicated). I. Female flower (top view), disc segments (red arrow indicated), stigma (white arrow indicated). J. Female flower (lateral view). K, L. fruits and seeds. Photographs by Zhu-Qiu Song." figureDoi="http://doi.org/10.5281/zenodo.14522100" httpUri="https://zenodo.org/record/14522100/files/figure.png" pageId="5" pageNumber="100">Figure 1K, L</figureCitation>
) (vs.
<quantity id="4CB39B2848700173FE0BFDE7FDB8B307" box="[476,582,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="5" pageNumber="100" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
in diameter and subglobose in polar view). Additionally, the new species also much differs from other members accepted currently within the subgenus
<taxonomicName id="4C4B4D4E48700173FC13FDCBFB65B362" box="[964,1179,554,580]" pageId="5" pageNumber="100" rank="subGenus" subGenus="Phyllanthodendron">
<emphasis id="B93FEADF48700173FC13FDCBFB65B362" box="[964,1179,554,580]" italics="true" pageId="5" pageNumber="100">Phyllanthodendron</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF34664548700173FF5FF9D9FB78B768" ID-DOI="http://doi.org/10.5281/zenodo.14522106" ID-Zenodo-Dep="14522106" box="[136,1158,1592,1614]" httpUri="https://zenodo.org/record/14522106/files/figure.png" pageId="5" pageNumber="100" startId="5.[136,229,1592,1614]" targetBox="[161,1426,612,1555]" targetPageId="5" targetType="figure">
<paragraph id="8BF436CD48700173FF5FF9D9FB78B768" blockId="5.[136,1158,1592,1614]" box="[136,1158,1592,1614]" pageId="5" pageNumber="100">
<emphasis id="B93FEADF48700173FF5FF9D9FF00B76B" bold="true" box="[136,254,1592,1614]" pageId="5" pageNumber="100">FIGURE 4.</emphasis>
Distribution map of
<taxonomicName id="4C4B4D4E48700173FE10F9D9FCC6B768" authority="Gang Yao" authorityName="Gang Yao" box="[455,824,1592,1614]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="species" species="liangkwangense">
<emphasis id="B93FEADF48700173FE10F9D9FD2AB76B" box="[455,724,1592,1613]" italics="true" pageId="5" pageNumber="100">Glochidion liangkwangense</emphasis>
Gang Yao
</taxonomicName>
&amp; Z.Q. Song (red stars indicated).
</paragraph>
</caption>
<paragraph id="8BF436CD48700173FF6AF99CFAEEB79D" blockId="5.[136,1451,1661,1723]" pageId="5" pageNumber="100">
Due to the lack of specialized literature for the identification of
<taxonomicName id="4C4B4D4E48700173FC7CF99CFAAEB7B1" box="[939,1360,1661,1687]" class="Magnoliopsida" family="Phyllanthaceae" genus="Glochidion" kingdom="Plantae" order="Malpighiales" pageId="5" pageNumber="100" phylum="Tracheophyta" rank="subGenus" subGenus="Phyllanthodendron">
<emphasis id="B93FEADF48700173FC7CF99CFBD7B7B1" box="[939,1065,1661,1687]" italics="true" pageId="5" pageNumber="100">Glochidion</emphasis>
subg.
<emphasis id="B93FEADF48700173FBAEF99CFAAEB7B1" box="[1145,1360,1661,1687]" italics="true" pageId="5" pageNumber="100">Phyllanthodendron</emphasis>
</taxonomicName>
species occurring in Asia, and considering the great similarity between them, it provides a key to identify them.
</paragraph>
</subSubSection>
</treatment>
</document>