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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.175.2259" ID-GBIF-Dataset="498b54e4-04bb-4f17-9e6b-4ed915e8538e" ID-PMC="PMC3307346" ID-Pensoft-Pub="1313-2970-175-27" ID-PubMed="22451789" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-175-27" ModsDocOrigin="ZooKeys 175" ModsDocTitle="Loxosceles niedeguidonae (Araneae, Sicariidae) a new species of brown spider from Brazilian semi-arid region" checkinTime="1451249292830" checkinUser="pensoft" docAuthor="Goncalves-de-Andrade, Rute Maria, Bertani, Rogerio, Nagahama, Roberto Hiroaki &amp; Barbosa, Maria Fatima Ribeiro" docDate="2012" docId="4E3C18891CF90C722A060801098AEDCF" docLanguage="en" docName="ZooKeys 175: 27-36" docOrigin="ZooKeys 175" docSource="http://dx.doi.org/10.3897/zookeys.175.2259" docTitle="Loxosceles niedeguidonae Goncalves-de-Andrade, Bertani, Nagahama &amp; Barbosa, 2012, sp. n." docType="treatment" docVersion="3" lastPageNumber="33" masterDocId="9005FF91FF92FF9D9F68FFBDFFDEFFD0" masterDocTitle="Loxosceles niedeguidonae (Araneae, Sicariidae) a new species of brown spider from Brazilian semi-arid region" masterLastPageNumber="36" masterPageNumber="27" pageNumber="28" updateTime="1668153372690" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Loxosceles niedeguidonae (Araneae, Sicariidae) a new species of brown spider from Brazilian semi-arid region</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Goncalves-de-Andrade, Rute Maria</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bertani, Rogerio</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nagahama, Roberto Hiroaki</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barbosa, Maria Fatima Ribeiro</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>175</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>27</mods:start>
<mods:end>36</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.175.2259</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.175.2259</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-175-27</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152033565" LSID="urn:lsid:plazi:treatment:4E3C18891CF90C722A060801098AEDCF" httpUri="http://treatment.plazi.org/id/4E3C18891CF90C722A060801098AEDCF" lastPageId="6" lastPageNumber="33" pageId="1" pageNumber="28">
<subSubSection pageId="1" pageNumber="28" type="nomenclature">
<paragraph pageId="1" pageNumber="28">
<taxonomicName LSID="urn:lsid:zoobank.org:act:ED9F0A10-B566-4989-AB71-00E1F96BD38E" class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="28">sp. n.</taxonomicNameLabel>
Figures 1-13
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="28" type="type material">
<paragraph pageId="1" pageNumber="28">Type material.</paragraph>
<paragraph pageId="1" pageNumber="28">
Holotype: male: Brazil,
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
, Coronel
<normalizedToken originalValue="José">Jose</normalizedToken>
Dias,
<normalizedToken originalValue="Boqueirão">Boqueirao</normalizedToken>
do
<normalizedToken originalValue="Sítio">Sitio</normalizedToken>
da Pedra Furada - Toca,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-8.85">8°51'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-42.55">42°33'W</geoCoordinate>
, 16 March 2009, R. M.
<normalizedToken originalValue="Gonçalves-de-Andrade">Goncalves-de-Andrade</normalizedToken>
&amp; Maria
<normalizedToken originalValue="Fátima">Fatima</normalizedToken>
Ribeiro Barbosa (MNRJ 04359).
</paragraph>
<paragraph pageId="1" pageNumber="28">Paratype female (MNRJ 04360) with same data as for holotype.</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29" type="other material examined">
<paragraph pageId="2" pageNumber="29">
<pageBreakToken pageId="2" pageNumber="29" start="start">Other</pageBreakToken>
material examined.
</paragraph>
<paragraph lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29">
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
. Holotype: Male: Brazil, Bahia, Palmeiras, Chapada Diamantina National Park (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.466666">12°28'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-41.416668">41°25'W</geoCoordinate>
), 15 February 2008, R. Bertani, C. S. Fukushima &amp; R. H. Nagahama, (MNRJ 6047); Paratypes:
<pageBreakToken pageId="3" pageNumber="30" start="start">Brazil</pageBreakToken>
, Bahia: 1 female, with same data as for holotype (MNRJ 6048); 3 females and 1 male,
<normalizedToken originalValue="Lençóis">Lencois</normalizedToken>
, Chapada Diamantina National Park (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-41.383335">41°23'W</geoCoordinate>
), 19 February 2008, same collectors as for holotype, (MNRJ 6049); 1 female, Iraquara, Fazenda Pra- tinha (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.35">12°21'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-41.533333">41°32'W</geoCoordinate>
), 16.II.2008, same collectors as for holotype, (MNRJ 6050).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="30" type="diagnosis">
<paragraph pageId="3" pageNumber="30">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="30">
Males can be readily distinguished from other species, except
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
by the following characters in combination: Palpal tibia more than 1.5 times longer than tarsus, embolus 1.5-1.8 times longer than diameter of tegulum, arising at distal half of tegulum, distally thin but not filiform and hardly curved, particularly not in distal direction (Fig. 2). Males differ from
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
by palpal tarsus almost two times longer than wide (Figs 2-4), embolus less curved (Fig. 4) and faded lateral dentate dark bands on the dorsal side of the carapace (Fig. 9). Females can be recognized by the following characters in combination: A narrow transversal plate in the spermathecae, straight, apically enlarged seminal receptacles and incrassate palpal tarsus, which is broader than palpal tibia (Figs 6-8).
</paragraph>
<caption pageId="3" pageNumber="30">
<paragraph pageId="3" pageNumber="30">
Figures 1-8.
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. 1-4 holotype male 1 carapace 2 left palp 3 bulb, prolateral view 4 bulb, retrolateral view 5-8 paratype female 5 carapace 6 right palp - incrassate tarsus (arrow) 7 spermathecae, dorsal view 8 spermathecae, ventral view. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="30" type="etymology">
<paragraph pageId="3" pageNumber="30">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="30">
The specific name is a patronym in honor of Dr
<normalizedToken originalValue="Niéde">Niede</normalizedToken>
Guidon, one of the most important Brazilian archeologists, internationally acknowledged for her archeological work and her battle for the preservation of archeological sites in Brazil and the conservation of remnant patches of Caatinga vegetation, as well as important social work, especially in the Parque Nacional da Serra da Capivara.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30" type="description">
<paragraph pageId="3" pageNumber="30">Description.</paragraph>
<paragraph pageId="3" pageNumber="30">Male(holotype). Total length (without chelicerae) 6.01. Carapace 3.29 long, 3.1 wide. Eye size: ALE: 0.2, PME: 0.22, PLE: 0.18. Clypeus: 0.32. Interocular distance - PME and PLE: 0.04, PME and ALE: 0.2. Leg formula: II, IV, I, III. Legs and palp length and width in Table 1. Labium 0.75 long, 0.42 wide. Sternum 1.59 long, 1.48 wide. Femur I 2.58 times longer than the carapace. Palpal femur 7.13 times longer than wide, tibia 2.81 times longer than wide (Fig. 2), tarsus almost two times longer than wide (Figs 3-4). Bulb suboval, slightly shorter than tarsus length. Embolus almost straight, approximately two times longer than bulb width, without carina (Fig. 4). Cephalic region of carapace covered by many long setae. Carapace with pars cephalic and chelicerae light brown (Fig. 1). Pars thoracica pale yellow, carapace border mottled, light brown (Figs 1, 9). Legs and palps pale yellow, covered by short grayish setae. Coxae and sternum pale yellow, labium and endites brown. Abdomen covered with grayish setae.</paragraph>
<caption ID-Table-UUID="40EE1F48916D9F6573F544D05309F1AF" httpUri="http://table.plazi.org/id/40EE1F48916D9F6573F544D05309F1AF" pageId="3" pageNumber="30">
<paragraph pageId="3" pageNumber="30">
Table 1.
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. Male holotype. Length/width of right legs and palpal segments.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="30">
<table pageId="3" pageNumber="30">
<tr pageId="3" pageNumber="30">
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">Palp</th>
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">I</th>
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">II</th>
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">III</th>
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">IV</th>
</tr>
</table>
</paragraph>
<caption pageId="3" pageNumber="30">
<paragraph pageId="3" pageNumber="30">
Figures 9-10. Carapaces, males. 9
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n., holotype 10
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
, holotype.
</paragraph>
</caption>
<paragraph lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30">
Female (paratype MNRJ 04360) Total length (without chelicerae) 7.35. Carapace 3.79 long, 3.71 wide. Eye sizes: ALE: 0.22, PME: 0.16, PLE: 0.18. Clypeus: 0.32. Interocular distances - PME and PLE: 0.05, PME and ALE: 0.24. Leg formula: II, I, VI, III. Legs and palp length and width in Table 2. Labium 0.7 long, 0.54 wide.
<pageBreakToken pageId="4" pageNumber="31" start="start">Sternum</pageBreakToken>
1.85 long, 1.78 wide. Femur I 1.58 times longer than carapace. Palpal femur 5.0 times longer than wide, tibia 3.5 longer than wide, tarsus incrassate (Fig. 6). Spermathecae with long, straight, apically enlarged seminal receptacles; transversal plate narrow, weakly sclerotized; atriobursal orifices well visible, ovals, positioned on the internal edge of the central windows; dorsal part of bursa copulatrix weakly sclero- tized (Figs 7-8). Coloration as in male, but darker (Fig. 12). Tarsi and tibiae of palps reddish-brown.
</paragraph>
<caption ID-Table-UUID="A4C00B331346E6B6729856C1B90B7DD9" httpUri="http://table.plazi.org/id/A4C00B331346E6B6729856C1B90B7DD9" pageId="4" pageNumber="31">
<paragraph pageId="4" pageNumber="31">
Table 2.
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. Female paratype (MNRJ 04360). Length/width of right legs and palpal segments.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="31">
<table pageId="4" pageNumber="31">
<tr pageId="4" pageNumber="31">
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">Palp</th>
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">I</th>
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">II</th>
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">III</th>
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">IV</th>
</tr>
</table>
</paragraph>
<caption pageId="4" pageNumber="31">
<paragraph pageId="4" pageNumber="31">
Figures 11-14.
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. 11 male 12 female 13 male in type locality - Toca do Buraco da Pedra Furada, Parque Nacional Serra da Capivara,
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
, Brazil 14 General view of Parque Nacional Serra da Capivara,
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
, Brazil. Photos 11-12 R. Bertani; 13-14 R. M.
<normalizedToken originalValue="Gonçalves-de-Andrade">Goncalves-de-Andrade</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="32" pageId="4" pageNumber="31" type="discussion">
<paragraph pageId="4" pageNumber="31">Discussion.</paragraph>
<paragraph lastPageId="5" lastPageNumber="32" pageId="4" pageNumber="31">
The female
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. can undoubtedly be included in the gaucho group of
<bibRefCitation pageId="4" pageNumber="31">Gertsch (1967)</bibRefCitation>
due to spermathecae bearing a transverse plate (Figs 7-8), which is characteristic of the group. However, the male resembles species of the laeta group due to its long palpal tibia (Fig. 2). These characteristics are also found in the recently described species
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
with which
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. seems to be closely related. As already discussed by
<bibRefCitation author="Bertani, R" journalOrPublisher="Revista Iberica de Aracnologia" pageId="6" pageNumber="33" title="Loxosceles chapadensis (Araneae: Sicariidae): a new recluse spider species of the gaucho group from Brazil. The Journal of Arachnology 38: 364 - 367. http: // www. americanarachnology. org / JoA _ free / JoA _ v 38 _ n 2 / arac- 38 - 02 - 364. pdf," url="10.1636/A09-92.1" year="2010">Bertani et al. 2010</bibRefCitation>
, the long palpal tibia of males of
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
(and now,
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n.)
<pageBreakToken pageId="5" pageNumber="32" start="start">could</pageBreakToken>
be an homoplasy with species of the laeta and spadicea groups or a plesiomorphy, indicating a more basal position of the species in the gaucho group relative to these other species (
<bibRefCitation author="Bertani, R" journalOrPublisher="Revista Iberica de Aracnologia" pageId="6" pageNumber="33" title="Loxosceles chapadensis (Araneae: Sicariidae): a new recluse spider species of the gaucho group from Brazil. The Journal of Arachnology 38: 364 - 367. http: // www. americanarachnology. org / JoA _ free / JoA _ v 38 _ n 2 / arac- 38 - 02 - 364. pdf," url="10.1636/A09-92.1" year="2010">Bertani et al. 2010</bibRefCitation>
). Despite these shared characteristics they can be separated by genitalic and somatic characteristics. The male has a straighter embolus (Figs 3-4), palpal tarsus almost two times longer than wide and color pattern are not very similar to the typical gaucho pattern, i. e. lateral dentate dark bands on the dorsal side of carapace. The male resembles an individual of the laeta group due to its light-brown coloration and faded dentate dark bands on the carapace (Figs 1-9, 11). The female can be easily separated by the narrow transversal plate in spermathecae (Figs 7-8) and in- crassate palpal tarsus (Fig. 6), which is like the female
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles adelaida" order="Araneae" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="adelaida">Loxosceles adelaida</taxonomicName>
, gaucho group (Gertsch, 1967), but more dilated than the latter.
</paragraph>
<paragraph pageId="5" pageNumber="32">
The distinct, but close geographical distribution of
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
and
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. in Northeastern Brazil provides an additional indication that these two species are closely related.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="33" pageId="5" pageNumber="32" type="natural history">
<paragraph pageId="5" pageNumber="32">Natural History.</paragraph>
<paragraph lastPageId="6" lastPageNumber="33" pageId="5" pageNumber="32">
Specimens of
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. were found in the locality of &quot;Buraco da Pedra Furada&quot;. The male was collected over the ground and the female in a rock crevice close to a cave mouth. The arenitic walls in the Parque Nacional Serra da Capivara have innumerous crevices that make typical retreats for brown
<pageBreakToken pageId="6" pageNumber="33" start="start">recluse</pageBreakToken>
spiders (Figs 13-14). However, despite the presence of so many crevices, a low population density was found in the analyzed ecotope.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="33" type="distribution">
<paragraph pageId="6" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="33">
Known only from Parque Nacional Serra da Capivara,
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
State, Brazil.
</paragraph>
</subSubSection>
</treatment>
</document>