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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.175.2259" ID-GBIF-Dataset="498b54e4-04bb-4f17-9e6b-4ed915e8538e" ID-PMC="PMC3307346" ID-Pensoft-Pub="1313-2970-175-27" ID-PubMed="22451789" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-175-27" ModsDocOrigin="ZooKeys 175" ModsDocTitle="Loxosceles niedeguidonae (Araneae, Sicariidae) a new species of brown spider from Brazilian semi-arid region" checkinTime="1451249292830" checkinUser="pensoft" docAuthor="Goncalves-de-Andrade, Rute Maria, Bertani, Rogerio, Nagahama, Roberto Hiroaki &amp; Barbosa, Maria Fatima Ribeiro" docDate="2012" docId="4E3C18891CF90C722A060801098AEDCF" docLanguage="en" docName="ZooKeys 175: 27-36" docOrigin="ZooKeys 175" docSource="http://dx.doi.org/10.3897/zookeys.175.2259" docTitle="Loxosceles niedeguidonae Goncalves-de-Andrade, Bertani, Nagahama &amp; Barbosa, 2012, sp. n." docType="treatment" docVersion="3" lastPageNumber="33" masterDocId="9005FF91FF92FF9D9F68FFBDFFDEFFD0" masterDocTitle="Loxosceles niedeguidonae (Araneae, Sicariidae) a new species of brown spider from Brazilian semi-arid region" masterLastPageNumber="36" masterPageNumber="27" pageNumber="28" updateTime="1668153372690" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Loxosceles niedeguidonae (Araneae, Sicariidae) a new species of brown spider from Brazilian semi-arid region</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Goncalves-de-Andrade, Rute Maria</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Bertani, Rogerio</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nagahama, Roberto Hiroaki</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barbosa, Maria Fatima Ribeiro</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>175</mods:number>
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<mods:start>27</mods:start>
<mods:end>36</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.175.2259</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.175.2259</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-175-27</mods:identifier>
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<treatment ID-GBIF-Taxon="152033565" LSID="urn:lsid:plazi:treatment:4E3C18891CF90C722A060801098AEDCF" httpUri="http://treatment.plazi.org/id/4E3C18891CF90C722A060801098AEDCF" lastPageId="6" lastPageNumber="33" pageId="1" pageNumber="28">
<subSubSection pageId="1" pageNumber="28" type="nomenclature">
<paragraph pageId="1" pageNumber="28">
<taxonomicName LSID="urn:lsid:zoobank.org:act:ED9F0A10-B566-4989-AB71-00E1F96BD38E" class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
<taxonomicNameLabel pageId="1" pageNumber="28">sp. n.</taxonomicNameLabel>
Figures 1-13
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="28" type="type material">
<paragraph pageId="1" pageNumber="28">Type material.</paragraph>
<paragraph pageId="1" pageNumber="28">
Holotype: male: Brazil,
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
, Coronel
<normalizedToken originalValue="José">Jose</normalizedToken>
Dias,
<normalizedToken originalValue="Boqueirão">Boqueirao</normalizedToken>
do
<normalizedToken originalValue="Sítio">Sitio</normalizedToken>
da Pedra Furada - Toca,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-8.85">8°51'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-42.55">42°33'W</geoCoordinate>
, 16 March 2009, R. M.
<normalizedToken originalValue="Gonçalves-de-Andrade">Goncalves-de-Andrade</normalizedToken>
&amp; Maria
<normalizedToken originalValue="Fátima">Fatima</normalizedToken>
Ribeiro Barbosa (MNRJ 04359).
</paragraph>
<paragraph pageId="1" pageNumber="28">Paratype female (MNRJ 04360) with same data as for holotype.</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29" type="other material examined">
<paragraph pageId="2" pageNumber="29">
<pageBreakToken pageId="2" pageNumber="29" start="start">Other</pageBreakToken>
material examined.
</paragraph>
<paragraph lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29">
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
. Holotype: Male: Brazil, Bahia, Palmeiras, Chapada Diamantina National Park (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.466666">12°28'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-41.416668">41°25'W</geoCoordinate>
), 15 February 2008, R. Bertani, C. S. Fukushima &amp; R. H. Nagahama, (MNRJ 6047); Paratypes:
<pageBreakToken pageId="3" pageNumber="30" start="start">Brazil</pageBreakToken>
, Bahia: 1 female, with same data as for holotype (MNRJ 6048); 3 females and 1 male,
<normalizedToken originalValue="Lençóis">Lencois</normalizedToken>
, Chapada Diamantina National Park (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-41.383335">41°23'W</geoCoordinate>
), 19 February 2008, same collectors as for holotype, (MNRJ 6049); 1 female, Iraquara, Fazenda Pra- tinha (
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.35">12°21'S</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-41.533333">41°32'W</geoCoordinate>
), 16.II.2008, same collectors as for holotype, (MNRJ 6050).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="30" type="diagnosis">
<paragraph pageId="3" pageNumber="30">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="30">
Males can be readily distinguished from other species, except
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
by the following characters in combination: Palpal tibia more than 1.5 times longer than tarsus, embolus 1.5-1.8 times longer than diameter of tegulum, arising at distal half of tegulum, distally thin but not filiform and hardly curved, particularly not in distal direction (Fig. 2). Males differ from
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
by palpal tarsus almost two times longer than wide (Figs 2-4), embolus less curved (Fig. 4) and faded lateral dentate dark bands on the dorsal side of the carapace (Fig. 9). Females can be recognized by the following characters in combination: A narrow transversal plate in the spermathecae, straight, apically enlarged seminal receptacles and incrassate palpal tarsus, which is broader than palpal tibia (Figs 6-8).
</paragraph>
<caption pageId="3" pageNumber="30">
<paragraph pageId="3" pageNumber="30">
Figures 1-8.
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. 1-4 holotype male 1 carapace 2 left palp 3 bulb, prolateral view 4 bulb, retrolateral view 5-8 paratype female 5 carapace 6 right palp - incrassate tarsus (arrow) 7 spermathecae, dorsal view 8 spermathecae, ventral view. Scale bar: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="30" type="etymology">
<paragraph pageId="3" pageNumber="30">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="30">
The specific name is a patronym in honor of Dr
<normalizedToken originalValue="Niéde">Niede</normalizedToken>
Guidon, one of the most important Brazilian archeologists, internationally acknowledged for her archeological work and her battle for the preservation of archeological sites in Brazil and the conservation of remnant patches of Caatinga vegetation, as well as important social work, especially in the Parque Nacional da Serra da Capivara.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30" type="description">
<paragraph pageId="3" pageNumber="30">Description.</paragraph>
<paragraph pageId="3" pageNumber="30">Male(holotype). Total length (without chelicerae) 6.01. Carapace 3.29 long, 3.1 wide. Eye size: ALE: 0.2, PME: 0.22, PLE: 0.18. Clypeus: 0.32. Interocular distance - PME and PLE: 0.04, PME and ALE: 0.2. Leg formula: II, IV, I, III. Legs and palp length and width in Table 1. Labium 0.75 long, 0.42 wide. Sternum 1.59 long, 1.48 wide. Femur I 2.58 times longer than the carapace. Palpal femur 7.13 times longer than wide, tibia 2.81 times longer than wide (Fig. 2), tarsus almost two times longer than wide (Figs 3-4). Bulb suboval, slightly shorter than tarsus length. Embolus almost straight, approximately two times longer than bulb width, without carina (Fig. 4). Cephalic region of carapace covered by many long setae. Carapace with pars cephalic and chelicerae light brown (Fig. 1). Pars thoracica pale yellow, carapace border mottled, light brown (Figs 1, 9). Legs and palps pale yellow, covered by short grayish setae. Coxae and sternum pale yellow, labium and endites brown. Abdomen covered with grayish setae.</paragraph>
<caption ID-Table-UUID="40EE1F48916D9F6573F544D05309F1AF" httpUri="http://table.plazi.org/id/40EE1F48916D9F6573F544D05309F1AF" pageId="3" pageNumber="30">
<paragraph pageId="3" pageNumber="30">
Table 1.
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. Male holotype. Length/width of right legs and palpal segments.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="30">
<table pageId="3" pageNumber="30">
<tr pageId="3" pageNumber="30">
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">Palp</th>
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">I</th>
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">II</th>
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">III</th>
<th colspan="1" pageId="3" pageNumber="30" rowspan="1">IV</th>
</tr>
</table>
</paragraph>
<caption pageId="3" pageNumber="30">
<paragraph pageId="3" pageNumber="30">
Figures 9-10. Carapaces, males. 9
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n., holotype 10
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
, holotype.
</paragraph>
</caption>
<paragraph lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30">
Female (paratype MNRJ 04360) Total length (without chelicerae) 7.35. Carapace 3.79 long, 3.71 wide. Eye sizes: ALE: 0.22, PME: 0.16, PLE: 0.18. Clypeus: 0.32. Interocular distances - PME and PLE: 0.05, PME and ALE: 0.24. Leg formula: II, I, VI, III. Legs and palp length and width in Table 2. Labium 0.7 long, 0.54 wide.
<pageBreakToken pageId="4" pageNumber="31" start="start">Sternum</pageBreakToken>
1.85 long, 1.78 wide. Femur I 1.58 times longer than carapace. Palpal femur 5.0 times longer than wide, tibia 3.5 longer than wide, tarsus incrassate (Fig. 6). Spermathecae with long, straight, apically enlarged seminal receptacles; transversal plate narrow, weakly sclerotized; atriobursal orifices well visible, ovals, positioned on the internal edge of the central windows; dorsal part of bursa copulatrix weakly sclero- tized (Figs 7-8). Coloration as in male, but darker (Fig. 12). Tarsi and tibiae of palps reddish-brown.
</paragraph>
<caption ID-Table-UUID="A4C00B331346E6B6729856C1B90B7DD9" httpUri="http://table.plazi.org/id/A4C00B331346E6B6729856C1B90B7DD9" pageId="4" pageNumber="31">
<paragraph pageId="4" pageNumber="31">
Table 2.
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. Female paratype (MNRJ 04360). Length/width of right legs and palpal segments.
</paragraph>
</caption>
<paragraph pageId="4" pageNumber="31">
<table pageId="4" pageNumber="31">
<tr pageId="4" pageNumber="31">
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">Palp</th>
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">I</th>
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">II</th>
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">III</th>
<th colspan="1" pageId="4" pageNumber="31" rowspan="1">IV</th>
</tr>
</table>
</paragraph>
<caption pageId="4" pageNumber="31">
<paragraph pageId="4" pageNumber="31">
Figures 11-14.
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. 11 male 12 female 13 male in type locality - Toca do Buraco da Pedra Furada, Parque Nacional Serra da Capivara,
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
, Brazil 14 General view of Parque Nacional Serra da Capivara,
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
, Brazil. Photos 11-12 R. Bertani; 13-14 R. M.
<normalizedToken originalValue="Gonçalves-de-Andrade">Goncalves-de-Andrade</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="32" pageId="4" pageNumber="31" type="discussion">
<paragraph pageId="4" pageNumber="31">Discussion.</paragraph>
<paragraph lastPageId="5" lastPageNumber="32" pageId="4" pageNumber="31">
The female
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. can undoubtedly be included in the gaucho group of
<bibRefCitation pageId="4" pageNumber="31">Gertsch (1967)</bibRefCitation>
due to spermathecae bearing a transverse plate (Figs 7-8), which is characteristic of the group. However, the male resembles species of the laeta group due to its long palpal tibia (Fig. 2). These characteristics are also found in the recently described species
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
with which
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. seems to be closely related. As already discussed by
<bibRefCitation author="Bertani, R" journalOrPublisher="Revista Iberica de Aracnologia" pageId="6" pageNumber="33" title="Loxosceles chapadensis (Araneae: Sicariidae): a new recluse spider species of the gaucho group from Brazil. The Journal of Arachnology 38: 364 - 367. http: // www. americanarachnology. org / JoA _ free / JoA _ v 38 _ n 2 / arac- 38 - 02 - 364. pdf," url="10.1636/A09-92.1" year="2010">Bertani et al. 2010</bibRefCitation>
, the long palpal tibia of males of
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
(and now,
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n.)
<pageBreakToken pageId="5" pageNumber="32" start="start">could</pageBreakToken>
be an homoplasy with species of the laeta and spadicea groups or a plesiomorphy, indicating a more basal position of the species in the gaucho group relative to these other species (
<bibRefCitation author="Bertani, R" journalOrPublisher="Revista Iberica de Aracnologia" pageId="6" pageNumber="33" title="Loxosceles chapadensis (Araneae: Sicariidae): a new recluse spider species of the gaucho group from Brazil. The Journal of Arachnology 38: 364 - 367. http: // www. americanarachnology. org / JoA _ free / JoA _ v 38 _ n 2 / arac- 38 - 02 - 364. pdf," url="10.1636/A09-92.1" year="2010">Bertani et al. 2010</bibRefCitation>
). Despite these shared characteristics they can be separated by genitalic and somatic characteristics. The male has a straighter embolus (Figs 3-4), palpal tarsus almost two times longer than wide and color pattern are not very similar to the typical gaucho pattern, i. e. lateral dentate dark bands on the dorsal side of carapace. The male resembles an individual of the laeta group due to its light-brown coloration and faded dentate dark bands on the carapace (Figs 1-9, 11). The female can be easily separated by the narrow transversal plate in spermathecae (Figs 7-8) and in- crassate palpal tarsus (Fig. 6), which is like the female
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles adelaida" order="Araneae" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="adelaida">Loxosceles adelaida</taxonomicName>
, gaucho group (Gertsch, 1967), but more dilated than the latter.
</paragraph>
<paragraph pageId="5" pageNumber="32">
The distinct, but close geographical distribution of
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles chapadensis" order="Araneae" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="chapadensis">Loxosceles chapadensis</taxonomicName>
and
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. in Northeastern Brazil provides an additional indication that these two species are closely related.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="33" pageId="5" pageNumber="32" type="natural history">
<paragraph pageId="5" pageNumber="32">Natural History.</paragraph>
<paragraph lastPageId="6" lastPageNumber="33" pageId="5" pageNumber="32">
Specimens of
<taxonomicName class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles niedeguidonae" order="Araneae" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="niedeguidonae">Loxosceles niedeguidonae</taxonomicName>
sp. n. were found in the locality of &quot;Buraco da Pedra Furada&quot;. The male was collected over the ground and the female in a rock crevice close to a cave mouth. The arenitic walls in the Parque Nacional Serra da Capivara have innumerous crevices that make typical retreats for brown
<pageBreakToken pageId="6" pageNumber="33" start="start">recluse</pageBreakToken>
spiders (Figs 13-14). However, despite the presence of so many crevices, a low population density was found in the analyzed ecotope.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="33" type="distribution">
<paragraph pageId="6" pageNumber="33">Distribution.</paragraph>
<paragraph pageId="6" pageNumber="33">
Known only from Parque Nacional Serra da Capivara,
<normalizedToken originalValue="Piauí">Piaui</normalizedToken>
State, Brazil.
</paragraph>
</subSubSection>
</treatment>
</document>

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Holmgren, 1857
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1.
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.
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<paragraph id="EBDC1130A3B1DE413D85E294CFCC80DA" pageNumber="79">T. niger, nitidus; capite, thorace antice et abdomine glaberrimis; antennis tarsisque rufo-piceis.</paragraph>
<paragraph id="3597B5BEC3CDCA77105C2D8ED9010DAB" pageNumber="79">Worker. Length 2 lines. Black; the head glassy-smooth and shining; ovate, with the posterior margin of the vertex truncate; in some examples, an indistinct castaneous spot at the sides of the head, in the usual situation of the eyes; the antennae ferruginous. Thorax elongate, compressed at the sides; very smooth and shining anteriorly, with a delicate striation in the middle, the metathorax being finely rugulose; the tarsi rufo-piceous. Abdomen: ovate, very smooth and shining; the nodes subglobose, the basal one being the smallest.</paragraph>
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<paragraph id="5069819111960A8A87375035D6FA69D5" pageNumber="79">Hab. Borneo (Sarawak).</paragraph>
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classification Animalia Hemiptera Miridae
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<taxonomicName LSID="http://zoobank.org/C6849117-A190-43E2-8017-34C5A33D03F4" authority="Wolski &amp; Gorczyca" class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius acutipennis" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="acutipennis">Xenocylapidius acutipennis Wolski &amp; Gorczyca</taxonomicName>
<taxonomicNameLabel pageId="4" pageNumber="77">sp. n.</taxonomicNameLabel>
Figures 1, 9, 16-20, 31
</paragraph>
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<subSubSection pageId="4" pageNumber="77" type="diagnosis">
<paragraph pageId="4" pageNumber="77">Diagnosis.</paragraph>
<paragraph pageId="4" pageNumber="77">Recognized by the dorsum mottled with brownish yellow (Fig. 1); the dark brown antennal segment II; the endosoma with two bundles of spiculi (SP1 and SP2); the medial sclerite (MS) long, weakly curved, tapering toward apex, sharply pointed; the sinistrolateral sclerite (SLS) large, occupying almost half of endosoma, strongly broadened basally, constricted medially; the clublike dextrolateral sclerite (DLS) (Fig. 16); and the right paramere sickle-shaped (Fig. 20).</paragraph>
<paragraph pageId="4" pageNumber="77">
Most similar to
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius gemellus" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="gemellus">Xenocylapidius gemellus</taxonomicName>
in sharing the brownish yellow mottling on dorsum (Figs 1, 5), the rounded extreme apex of apical process of left paramere when viewed dorsally (Figs 19, 35), and the sickle-shaped right paramere. This new species can, however, be distinguished by the dark brownish antennal segment and shape of the endosoma (Figs 16).
</paragraph>
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<subSubSection pageId="4" pageNumber="77" type="description">
<paragraph pageId="4" pageNumber="77">Description.</paragraph>
<paragraph pageId="4" pageNumber="77">Male. COLORATION (Figs 1, 9). Dorsum mostly with mottled, brownish yellow coloration. Head. Vertex and frons mottled with dark brown and yellow; remainder of head dark red with yellow mottling; antennal segment I dirty yellow, with an indistinct, dark yellow tinge basally and with a reddish tinge occupying apical one third of inner surface; segment II dark brown; labium dark brown with indistinct, dirty yellow areas. Thorax.Pronotum. Collar yellow; calli dark brown, with broad, yellowish mottling; anterior margin weakly tinged with red medially; lateral margin and posterior lobe dark brown, tinged with red and dirty yellow; humeral angle and medial portion of posterior margin yellow. Mesoscutum and scutellum. Mostly reddish; mesoscutum weakly tinged with dark brown medially and with dirty yellow area bordering portion being depressed onto lateral margin; scutellum reddish with dirty yellow patch apically. Thoracic pleura. Proepimeron and proepisternum mostly dark brown with reddish areas; remaining pleura reddish, with indistinct yellowish areas. Hemelytron. Corium and clavus dark brown, mottled with yellow; cuneus dark brown, weakly tinged with red, inner angle yellow, apex with a small, dirty yellow patch; membrane fuscous with indistinct, dirty yellow areas. Legs. Procoxa dark brown, dirty yellow apically; meso- and metacoxae yellow; femora dirty yellow brown with reddish areas; tibiae dark brown; tarsi dirty yellow brown. Abdomen. Dark brown with large dirty yellow areas. STRUCTURE, TEXTURE, AND VESTITURE (Figs 1, 9). Head. Antennal segment II weakly broadened toward apex, covered with moderately dense, adpressed and semirecumbent setae, sparse on basal one-fifth of segment II and dense on remainder of segment. Thorax.Pronotum. Lateral margins sharply carinate, somewhat elevated. Mesoscutum and scutellum. Scutellum weakly convex. Hemelytron. Covered with short, relatively dense, adpressed, black setae.</paragraph>
<paragraph pageId="4" pageNumber="77">Male genitalia.Aedeagus (Figs 16-17). Basal sac (BSC) occupying one third of endosoma; sclerotized portion of ductus seminis inside endosoma (DSS) ovoid; secondary gonopore nearly circular, not fully closed; basal fig (BP) irregular in shape; apex of endosoma with two bundles of spiculi (SP1 and SP2); medial sclerite (MS) long, weakly curved, tapering toward apex, sharply pointed; sinistrolateral sclerite (SLS) large, occupying almost half of endosoma, strongly broadened basally, constricted medially, and broadened, nearly cylindrical on apical half; dextrolateral sclerite (DLS) somewhat smaller than SLS, clublike. Left paramere (Figs 18-19). Apical process: lateral view: weakly tapering toward apex, obtuse apically; dorsal view: lateral margins weakly curved, extreme apex rounded; paramere body: dorsal view: weakly broadened toward apex; sensory lobe: massive, just slightly tapering toward apex, obtuse. Right paramere (Fig. 20). Sickle-shaped; apical process: long, thin, arcuate, just slightly narrowed toward apex; paramere body: thin, dorsal margin straight, ventral margin weakly arcuate.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="77" type="measurements">
<paragraph pageId="4" pageNumber="77">Measurements.</paragraph>
<paragraph pageId="4" pageNumber="77">♀/♂ (n=2, holotype measurements second). Body. Length 6.00/4.70, width 2.15/1.76. Head. Length 1.00/0.98, width 0.85/0.79, interocular distance 0.35/0.35. Antenna. Length of segment I 0.74/0.64, II 1.83/1.83 (III and IV missing in both specimens). Labium. Length of segment I 0.98/0.95 (II, III, and IV immeasurable in both specimens). Pronotum. Length 0.85/0.73, width of anterior margin 0.68/0.65, length of lateral margin 0.98/0.80, width of posterior margin 1.60/1.38.</paragraph>
<paragraph pageId="4" pageNumber="77">Female. Similar to male in coloration, structure, texture, and vestiture.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="77" type="biology">
<paragraph pageId="4" pageNumber="77">Biology.</paragraph>
<paragraph pageId="4" pageNumber="77">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="77" type="distribution">
<paragraph pageId="4" pageNumber="77">Distribution.</paragraph>
<paragraph pageId="4" pageNumber="77">Australia (Queensland) (Fig. 31).</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="77" type="etymology">
<paragraph pageId="4" pageNumber="77">Etymology.</paragraph>
<paragraph pageId="4" pageNumber="77">
The specific name is derived from the Latin
<normalizedToken originalValue="“acutus”">&quot;acutus&quot;</normalizedToken>
, meaning sharpened, and is used to denote the sharply pointed mesial process (MS) of the endosoma.
</paragraph>
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<subSubSection pageId="4" pageNumber="77" type="type material">
<paragraph pageId="4" pageNumber="77">Type material.</paragraph>
<paragraph pageId="4" pageNumber="77">
Holotype ♂: Malanda; Queensl[and]
<normalizedToken originalValue="Mjöberg">Mjoeberg</normalizedToken>
; Swedish Museum of Natural History Stockholm NHRS (NHRS); paratype 1 ♀: Glen Lamington Queensl[and]
<normalizedToken originalValue="Mjöberg">Mjoeberg</normalizedToken>
; Swedish Museum of Natural History Stockholm NHRS (NHRS).
</paragraph>
<caption pageId="4" pageNumber="77">
<paragraph pageId="4" pageNumber="77">
Figures 1-8. Dorsal habitus color photographs of
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="genus">Xenocylapidius</taxonomicName>
spp.: 1
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius acutipennis" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="acutipennis">Xenocylapidius acutipennis</taxonomicName>
(holotype) 2
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius ater" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="ater">Xenocylapidius ater</taxonomicName>
(holotype) 3
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius bimaculatus" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="bimaculatus">Xenocylapidius bimaculatus</taxonomicName>
(holotype) 4
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius bioculatus" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="bioculatus">Xenocylapidius bioculatus</taxonomicName>
(♀: Australia N. S. W., Manly nr Sydney, North Head 16-21.2., D. Shcherbakov 1997) 5
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius gemellus" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="gemellus">Xenocylapidius gemellus</taxonomicName>
(holotype) 6
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius gressitti" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="gressitti">Xenocylapidius gressitti</taxonomicName>
(holotype) 7
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius rolandi" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="rolandi">Xenocylapidius rolandi</taxonomicName>
(holotype) 8
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius tamasi" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="tamasi">Xenocylapidius tamasi</taxonomicName>
(♀: New Caledonia, Foret di Thi, 29.X.-1.XI.1967).
</paragraph>
</caption>
<caption pageId="4" pageNumber="77">
<paragraph pageId="4" pageNumber="77">
Figures 9-15. Color photographs of
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="genus">Xenocylapidius</taxonomicName>
spp., lateral views: 9
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius acutipennis" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="acutipennis">Xenocylapidius acutipennis</taxonomicName>
(holotype) 10
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius ater" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="ater">Xenocylapidius ater</taxonomicName>
(holotype); 11.
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius bimaculatus" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="bimaculatus">Xenocylapidius bimaculatus</taxonomicName>
(holotype) 12
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius bioculatus" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="bioculatus">Xenocylapidius bioculatus</taxonomicName>
(♀: Australia N. S. W., Manly nr Sydney, North Head 16-21.2., D. Shcherbakov 1997) 13
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius gressitti" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="gressitti">Xenocylapidius gressitti</taxonomicName>
(holotype) 14
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius rolandi" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="rolandi">Xenocylapidius rolandi</taxonomicName>
(holotype) 15
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius tamasi" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="tamasi">Xenocylapidius tamasi</taxonomicName>
(♀: New Caledonia, Foret di Thi, 29.X.-1.XI.1967).
</paragraph>
</caption>
<caption pageId="4" pageNumber="77">
<paragraph pageId="4" pageNumber="77">
Figures 16-25. Male genitalia of
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius acutipennis" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="acutipennis">Xenocylapidius acutipennis</taxonomicName>
(16-20) and
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius ater" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="ater">Xenocylapidius ater</taxonomicName>
(21-25): 16, 21 Endosoma (dorsal view) 17, 22 Basal sac of endosoma (ventral view) 18, 23 Left paramere (left lateral view) 19, 24 Left paramere (dorsal view) 20, 25 Right paramere (right lateral view). APR = apical process of paramere; AR = apical ring of endosomal basal sac; BP = basal fig; BPR = basal process of paramere; BSC = basal sac; DLS = dextrolateral sclerite; DSS = sclerotized portion of ductus seminis inside endosoma; MS = medial sclerite; PB = paramere body; SL = sensory lobe; SLS = sinistrolateral sclerite; SP1 and SP2 = endosomal spiculi.
</paragraph>
</caption>
<caption pageId="4" pageNumber="77">
<paragraph pageId="4" pageNumber="77">
Figures 26-30. Male genitalia of
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius bimaculatus" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="bimaculatus">Xenocylapidius bimaculatus</taxonomicName>
: 26 Endosoma (dorsal view) 27 Basal sac of endosoma (ventral view) 28 Left paramere (left lateral view) 29 Left paramere (dorsal view) 30 Right paramere (right lateral view). APR = apical process of paramere; AR = apical ring of endosomal basal sac; BPR = basal process of paramere; BSC = basal sac; DSS = sclerotized portion of ductus seminis inside endosoma; PB = paramere body; SL = sensory lobe; SLS = sinistrolateral sclerite; SP1 = endosomal spiculi.
</paragraph>
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<caption pageId="4" pageNumber="77">
<paragraph pageId="4" pageNumber="77">
Figure 31. Distribution map of
<taxonomicName class="Insecta" family="Miridae" genus="Xenocylapidius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenocylapidius" order="Hemiptera" pageId="4" pageNumber="77" phylum="Arthropoda" rank="genus">Xenocylapidius</taxonomicName>
spp.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,79 @@
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Genus
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Thorell, 1898
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Thorell, 1898
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<mods:identifier id="86894FAC5774D09CE79483A5CDFD357C" type="DOI">10.11646/zootaxa.4963.1.5</mods:identifier>
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<treatment id="4E3C87F3FFC0FF95C6ECFBFFFC7BFD5C" ID-DOI="http://doi.org/10.5281/zenodo.4696996" ID-GBIF-Taxon="181600554" ID-Zenodo-Dep="4696996" LSID="urn:lsid:plazi:treatment:4E3C87F3FFC0FF95C6ECFBFFFC7BFD5C" httpUri="http://treatment.plazi.org/id/4E3C87F3FFC0FF95C6ECFBFFFC7BFD5C" lastPageId="4" lastPageNumber="119" pageId="1" pageNumber="116">
<subSubSection id="8E8F656EFFC0FF90C6ECFBFFFE78FBD9" box="[151,462,1086,1113]" pageId="1" pageNumber="116" type="nomenclature">
<paragraph id="C62A36E5FFC0FF90C6ECFBFFFE78FBD9" blockId="1.[151,500,1086,1149]" box="[151,462,1086,1113]" pageId="1" pageNumber="116">
<heading id="9D628189FFC0FF90C6ECFBFFFE78FBD9" bold="true" box="[151,462,1086,1113]" fontSize="11" level="1" pageId="1" pageNumber="116" reason="1">
<emphasis id="F4E1EAF7FFC0FF90C6ECFBFFFE78FBD9" bold="true" box="[151,462,1086,1113]" pageId="1" pageNumber="116">
<taxonomicName id="01954D66FFC0FF90C6ECFBFFFEDEFBD9" ID-CoL="98YXW" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[151,360,1086,1113]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="116" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFC0FF90C6ECFBFFFEDEFBD9" bold="true" box="[151,360,1086,1113]" italics="true" pageId="1" pageNumber="116">Belisana bachma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFD2578CFFC0FF90C70BFBFEFE78FBD9" box="[368,462,1087,1113]" pageId="1" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF90C6ECFBA2FE42FBFD" box="[151,500,1122,1149]" pageId="1" pageNumber="116" type="description">
<paragraph id="C62A36E5FFC0FF90C6ECFBA2FE42FBFD" blockId="1.[151,500,1086,1149]" box="[151,500,1122,1149]" pageId="1" pageNumber="116">
<figureCitation id="5EAE2A60FFC0FF90C6ECFBA2FF4CFBFD" box="[151,250,1123,1149]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1860,1884]" captionStartId-1="3.[151,250,1826,1850]" captionTargetBox-0="[231,1357,179,1834]" captionTargetBox-1="[243,1345,181,1802]" captionTargetId-0="figure-19@2.[231,1357,179,1834]" captionTargetId-1="figure-17@3.[243,1345,181,1802]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." captionText-1="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.4719926" figureDoi-1="http://doi.org/10.5281/zenodo.4719928" httpUri-0="https://zenodo.org/record/4719926/files/figure.png" httpUri-1="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 12</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C77DFBA2FEECFBFD" box="[262,346,1123,1149]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="1" pageNumber="116">13AB</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C71EFBA2FE0EFBFD" box="[357,440,1123,1149]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="1" pageNumber="116">14AB</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C7BFFBA3FE42FBFD" box="[452,500,1122,1149]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="1" pageNumber="116">15A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF90C6ECFB6AFCBEFA8D" pageId="1" pageNumber="116" type="materials_examined">
<paragraph id="C62A36E5FFC0FF90C6ECFB6AFCBEFA8D" blockId="1.[151,1437,1194,1977]" pageId="1" pageNumber="116">
<emphasis id="F4E1EAF7FFC0FF90C6ECFB6AFEFFFB44" bold="true" box="[151,329,1194,1221]" pageId="1" pageNumber="116">Type material.</emphasis>
<materialsCitation id="76FD3CB8FFC0FF90C728FB6AFC45FB69" ID-GBIF-Occurrence="3091104307" collectingDate="2015-08-27" collectionCode="IZCAS" collectorName="D. S. Pham" country="Vietnam" elevation="371" latitude="16.229383" location="Male" longLatPrecision="1" longitude="107.84818" municipality="Bach Ma National Park" pageId="1" pageNumber="116" specimenCount="1" typeStatus="holotype">
<emphasis id="F4E1EAF7FFC0FF90C728FB6AFE7FFB45" bold="true" box="[339,457,1195,1221]" pageId="1" pageNumber="116">
<typeStatus id="192E8847FFC0FF90C728FB6AFE72FB45" box="[339,452,1195,1221]" pageId="1" pageNumber="116" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="C34A603EFFC0FF90C7AFFB6AFDB8FB45" LSID="urn:lsid:plazi:treatment:4E3C87F3FFC0FF95C6ECFBFFFC7BFD5C:C34A603EFFC0FF90C7AFFB6AFDB8FB45" box="[468,526,1195,1221]" country="Vietnam" latitude="16.229383" longLatPrecision="1" longitude="107.84818" municipality="Bach Ma National Park" name="Male" pageId="1" pageNumber="116">Male</location>
(
<collectionCode id="A084AE20FFC0FF90C45BFB6AFDC5FB45" box="[544,627,1195,1221]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="116">IZCAS</collectionCode>
Ar41901),
<collectingMunicipality id="264EAC9FFFC0FF90C481FB6AFBBAFB45" box="[762,1036,1195,1221]" pageId="1" pageNumber="116">Bach Ma National Park</collectingMunicipality>
(
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,
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,
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elevation
<quantity id="016D9B00FFC0FF90C77FFB0EFEFBFB69" box="[260,333,1231,1257]" metricMagnitude="2" metricUnit="m" metricValue="3.71" pageId="1" pageNumber="116" unit="m" value="371.0">371 m</quantity>
</elevation>
), Thua Thien Hue,
<collectingCountry id="BE827675FFC0FF90C459FB0EFD32FB69" box="[546,644,1231,1257]" name="Vietnam" pageId="1" pageNumber="116">Vietnam</collectingCountry>
,
<date id="B22B1025FFC0FF90C4F5FB0EFCF7FB69" box="[654,833,1230,1257]" pageId="1" pageNumber="116" value="2015-08-27">
<collectingDate id="A26FE9CDFFC0FF90C4F5FB0EFCF7FB69" box="[654,833,1230,1257]" pageId="1" pageNumber="116" value="2015-08-27">27 August 2015</collectingDate>
</date>
,
<collectorName id="6B605333FFC0FF90C530FB0EFC75FB69" box="[843,963,1231,1257]" pageId="1" pageNumber="116">D.S. Pham</collectorName>
leg.
</materialsCitation>
<materialsCitation id="76FD3CB8FFC0FF90C583FB0EFF09FA8D" ID-GBIF-Occurrence="3091104313" collectionCode="IZCAS" country="Vietnam" pageId="1" pageNumber="116" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="F4E1EAF7FFC0FF90C583FB0EFBCFFB69" bold="true" box="[1016,1145,1231,1257]" pageId="1" pageNumber="116">
<typeStatus id="192E8847FFC0FF90C583FB0EFBC2FB69" box="[1016,1140,1231,1257]" pageId="1" pageNumber="116" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="D093FD6CFFC0FF90C2FBFB0EFB7CFB69" box="[1152,1226,1231,1257]" pageId="1" pageNumber="116" type="male">1 male</specimenCount>
(
<collectionCode id="A084AE20FFC0FF90C2A3FB0EFA9DFB69" box="[1240,1323,1231,1257]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="116">IZCAS</collectionCode>
Ar41902) and
</materialsCitation>
<materialsCitation id="76FD3CB8FFC0FF90C6BCFB32FCB2FA8D" ID-GBIF-Occurrence="3091104303" box="[199,772,1267,1293]" collectingDate="2015-08-27" collectionCode="IZCAS" collectorName="D. S. Pham" country="Vietnam" elevation="371" latitude="16.229383" location="Male" longLatPrecision="1" longitude="107.84818" municipality="Bach Ma National Park" pageId="1" pageNumber="116" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D093FD6CFFC0FF90C6BCFB32FE9EFA8D" box="[199,296,1267,1293]" pageId="1" pageNumber="116" type="female">1 female</specimenCount>
(
<collectionCode id="A084AE20FFC0FF90C74DFB32FE3FFA8D" box="[310,393,1267,1293]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="116">IZCAS</collectionCode>
Ar41903), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF90C6BCFAD6FBCAFAB1" box="[199,1148,1303,1329]" pageId="1" pageNumber="116" type="etymology">
<paragraph id="C62A36E5FFC0FF90C6BCFAD6FBCAFAB1" blockId="1.[151,1437,1194,1977]" box="[199,1148,1303,1329]" pageId="1" pageNumber="116">
<emphasis id="F4E1EAF7FFC0FF90C6BCFAD6FEF8FAB1" bold="true" box="[199,334,1303,1329]" pageId="1" pageNumber="116">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="192E8847FFC0FF90C4CEFAD6FD53FAB1" box="[693,741,1303,1329]" pageId="1" pageNumber="116">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF90C6BCFAFBFA9DF9D1" pageId="1" pageNumber="116" type="diagnosis">
<paragraph id="C62A36E5FFC0FF90C6BCFAFBFA9DF9D1" blockId="1.[151,1437,1194,1977]" pageId="1" pageNumber="116">
<emphasis id="F4E1EAF7FFC0FF90C6BCFAFBFEF4FAD4" bold="true" box="[199,322,1338,1364]" pageId="1" pageNumber="116">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFC0FF90C436FAFAFC7BFAD5" authority="Yao, Pham &amp; Li, 2015" authorityName="Yao, Pham &amp; Li" authorityYear="2015" box="[589,973,1338,1365]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="116" phylum="Arthropoda" rank="species" species="babensis">
<emphasis id="F4E1EAF7FFC0FF90C436FAFAFD78FAD5" box="[589,718,1339,1365]" italics="true" pageId="1" pageNumber="116">B. babensis</emphasis>
<bibRefCitation id="A2044B14FFC0FF90C4A8FAFAFC7BFAD5" author="Yao, Z. &amp; Pham, D. S. &amp; Li, S." box="[723,973,1338,1365]" journalOrPublisher="Zootaxa" pageId="1" pageNumber="116" pagination="1 - 82" part="3909" refId="ref8999" refString="Yao, Z., Pham, D. S. &amp; Li, S. (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa, 3909 (1), 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1.1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species" type="journal article" year="2015">Yao, Pham &amp; Li, 2015</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFC0FF90C27DFAFAFB7AFAD5" author="Yao, Z. &amp; Pham, D. S. &amp; Li, S." box="[1030,1228,1338,1365]" journalOrPublisher="Zootaxa" pageId="1" pageNumber="116" pagination="1 - 82" part="3909" refId="ref8999" refString="Yao, Z., Pham, D. S. &amp; Li, S. (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa, 3909 (1), 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1.1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species" type="journal article" year="2015">
Yao
<emphasis id="F4E1EAF7FFC0FF90C243FAFAFBDFFAD5" box="[1080,1129,1339,1365]" italics="true" pageId="1" pageNumber="116">et al</emphasis>
. 2015: 5
</bibRefCitation>
, figs 13) with the elongated bulbal apophyses (
<figureCitation id="5EAE2A60FFC0FF90C7A5FA9EFD80FAF9" box="[478,566,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2D</figureCitation>
) and vulval anterior arch of a similar dark internal shade (
<figureCitation id="5EAE2A60FFC0FF90C2B2FA9EFA90FAF9" box="[1225,1318,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C349FA9EFAD5FAF9" box="[1330,1379,1375,1401]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="1" pageNumber="116">14B</figureCitation>
), but males can be distinguished by the presence of a strongly conical subdisto-dorsally protruding (arrow in
<figureCitation id="5EAE2A60FFC0FF90C347FA42FA25FA1D" box="[1340,1427,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="1" pageNumber="116">Fig. 1A</figureCitation>
) and bifid disto-prolateral spine (arrow
<quantity id="016D9B00FFC0FF90C430FA66FDC1FA41" box="[587,631,1447,1473]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="116" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFC0FF90C404FA66FD6EFA41" box="[639,728,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="1" pageNumber="116">Figs 1C</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C49DFA66FCA0FA41" box="[742,790,1447,1473]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="1" pageNumber="116">13A</figureCitation>
) on the procursus, by the presence of a frontal apophysis (arrow in
<figureCitation id="5EAE2A60FFC0FF90C77DFA0AFEEFFA65" box="[262,345,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2E</figureCitation>
) and a pair of small ventral apophyses (arrow in
<figureCitation id="5EAE2A60FFC0FF90C504FA0AFC62FA65" box="[895,980,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2C</figureCitation>
) on the clypeus, and the cheliceral distal apophyses are directed laterally (arrow da in
<figureCitation id="5EAE2A60FFC0FF90C4E6FA2EFCB6F989" box="[669,768,1519,1545]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs. 2C</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C575FA2FFCF6F989" box="[782,832,1518,1545]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="1" pageNumber="116">15A</figureCitation>
; distal apophyses directed anteriorly in
<taxonomicName id="01954D66FFC0FF90C372FA2EFA3AF989" authorityName="Yao, Pham &amp; Li" authorityYear="2015" box="[1289,1420,1519,1545]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="116" phylum="Arthropoda" rank="species" species="babensis">
<emphasis id="F4E1EAF7FFC0FF90C372FA2EFA3AF989" box="[1289,1420,1519,1545]" italics="true" pageId="1" pageNumber="116">B. babensis</emphasis>
</taxonomicName>
). Females can be distinguished by having frontal “epigynal” pockets laterally on a sclerotized plate (arrow ep in
<figureCitation id="5EAE2A60FFC0FF90C317F9D2FF6BF9D1" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 2AB</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C693F9F6FE8CF9D1" box="[232,314,1591,1617]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="1" pageNumber="116">14AB</figureCitation>
) and the vulval pore plates are approximately as wide as long (arrow pp in
<figureCitation id="5EAE2A60FFC0FF90C2FDF9F6FB57F9D1" box="[1158,1249,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C297F9F6FAA8F9D1" box="[1260,1310,1591,1617]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="1" pageNumber="116">14B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC0FF95C6BCF99BFABFFD39" lastPageId="4" lastPageNumber="119" pageId="1" pageNumber="116" type="description">
<paragraph id="C62A36E5FFC0FF95C6BCF99BFE75FE01" blockId="1.[151,1437,1194,1977]" lastBlockId="4.[151,1437,151,732]" lastPageId="4" lastPageNumber="119" pageId="1" pageNumber="116">
<emphasis id="F4E1EAF7FFC0FF90C6BCF99BFE29F9F5" bold="true" box="[199,415,1626,1653]" pageId="1" pageNumber="116">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFC0FF90C7C9F99AFDAFF9F5" bold="true" box="[434,537,1627,1653]" pageId="1" pageNumber="116">
<typeStatus id="192E8847FFC0FF90C7C9F99AFDAFF9F5" box="[434,537,1627,1653]" pageId="1" pageNumber="116" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41901): Total length 2.01 (2.16 with clypeus), carapace 0.98 long, 0.65 wide, opisthosoma 1.18 long, 0.80 wide. Leg I: 16.89 (4.55 + 0.19 + 4.42 + 6.67 + 1.15), leg II: 11.08 (3.21 + 0.18 + 2.75 + 4.17 + 0.77), leg III: 7.40 (2.24 + 0.16 + 1.86 + 2.56 + 0.58), leg IV: 10.61 (3.14 + 0.17 + 2.69 + 3.97 + 0.64); tibia I L/d: 55. Distance PME-PME 0.17, diameter PME 0.08, distance PME-ALE 0.11, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFC0FF90C74CF92AFE19F885" box="[311,431,1771,1797]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs 2EF</figureCitation>
. Carapace pale, with brown lateral margins. Ocular area with indistinct dark marks and not elevated. Clypeus brown, with frontal apophysis (arrow in
<figureCitation id="5EAE2A60FFC0FF90C510F8CEFC76F8A9" box="[875,960,1807,1833]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2E</figureCitation>
) and pair of small ventral apophyses (arrow in
<figureCitation id="5EAE2A60FFC0FF90C693F8F2FEF6F8CD" box="[232,320,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Fig. 2C</figureCitation>
). Chelicerae (
<figureCitation id="5EAE2A60FFC0FF90C7A7F8F2FD88F8CD" box="[476,574,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="1" pageNumber="116">Figs. 2C</figureCitation>
,
<figureCitation id="5EAE2A60FFC0FF90C430F8F3FDCAF8CD" box="[587,636,1842,1869]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="1" pageNumber="116">15A</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.38). Sternum pale, approximately as wide as long (0.50). Thoracic furrow absent. Legs pale. Opisthosoma pale, with indistinct large spots. Palp as in
<figureCitation id="5EAE2A60FFC0FF90C7E9F85EFDA6F839" box="[402,528,1951,1977]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="1" pageNumber="116">Figs 1AB</figureCitation>
; coxa unmodified; trochanter with short ventral apophysis with small retrolateral apophysis (arrow
<quantity id="016D9B00FFC5FF95C719FF56FE38FF31" box="[354,398,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="119" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C7EDFF56FE5BFF31" box="[406,493,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Fig. 1B</figureCitation>
); femur with small proximo-dorsal apophysis (arrow
<quantity id="016D9B00FFC5FF95C237FF56FBCEFF31" box="[1100,1144,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="119" unit="in" value="2.0">2 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C2FBFF56FB60FF31" box="[1152,1238,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Fig. 1B</figureCitation>
); procursus (
<figureCitation id="5EAE2A60FFC5FF95C310FF56FF68FF53" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1AD</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C690FF78FE8BFF53" box="[235,317,185,211]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13AB</figureCitation>
) simple proximally but complex distally, strongly protruding subdisto-dorsally (arrow in
<figureCitation id="5EAE2A60FFC5FF95C34EFF78FA38FF53" box="[1333,1422,185,211]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Fig. 1A</figureCitation>
), with bifid disto-prolateral spine (arrow
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<figureCitation id="5EAE2A60FFC5FF95C4EFFF1DFD59FF76" box="[660,751,220,246]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C485FF1DFC87FF76" box="[766,817,220,246]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13A</figureCitation>
), large, curved subdisto-prolateral sclerite (arrow
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<figureCitation id="5EAE2A60FFC5FF95C6ECFF3EFF47FE99" box="[151,241,255,281]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C685FF3EFE87FE99" box="[254,305,255,281]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13A</figureCitation>
), disto-retrolateral membranous lamella (arrow in
<figureCitation id="5EAE2A60FFC5FF95C510FF3EFC70FE99" box="[875,966,255,281]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C5AFFF3EFBB5FE99" box="[980,1027,255,281]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13B</figureCitation>
) and tongue-shaped retrolateral flap (arrow f in
<figureCitation id="5EAE2A60FFC5FF95C76DFEE0FEC7FEBB" box="[278,369,289,315]" captionStart="FIGURE 1" captionStartId="2.[151,250,1860,1884]" captionTargetBox="[231,1357,179,1834]" captionTargetId="figure-19@2.[231,1357,179,1834]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bachma sp. nov., holotype male.AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="4" pageNumber="119">Figs 1D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C706FEE0FE18FEBB" box="[381,430,289,315]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13B</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFC5FF95C47EFEE0FDEAFEBB" box="[517,604,289,315]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Fig. 2D</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia Iat 6% proximally; legs with short, vertical setae on metatarsi, without spines or curved setae; tarsus Iwith 11 distinct pseudopodomeres.
</paragraph>
<caption id="92EA666DFFC3FF93C6ECF885FB23F86C" ID-DOI="http://doi.org/10.5281/zenodo.4719926" ID-Zenodo-Dep="4719926" httpUri="https://zenodo.org/record/4719926/files/figure.png" pageId="2" pageNumber="117" startId="2.[151,250,1860,1884]" targetBox="[231,1357,179,1834]" targetPageId="2">
<paragraph id="C62A36E5FFC3FF93C6ECF885FB23F86C" blockId="2.[151,1437,1860,2028]" pageId="2" pageNumber="117">
<emphasis id="F4E1EAF7FFC3FF93C6ECF885FEA5F8DC" bold="true" box="[151,275,1860,1884]" pageId="2" pageNumber="117">FIGURE 1.</emphasis>
<taxonomicName id="01954D66FFC3FF93C763F885FE71F8DC" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[280,455,1860,1884]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="117" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFC3FF93C763F885FE71F8DC" box="[280,455,1860,1884]" italics="true" pageId="2" pageNumber="117">Belisana bachma</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFC3FF93C7B7F885FDAAF8DC" bold="true" box="[460,540,1860,1884]" pageId="2" pageNumber="117">
<taxonomicNameLabel id="EFD2578CFFC3FF93C7B7F885FDAAF8DC" box="[460,540,1860,1884]" pageId="2" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at subdistodorsal protruding part of procursus; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid disto-prolateral spine, arrow 2 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD).
</paragraph>
</caption>
<caption id="92EA666DFFC2FF92C6ECF8E3FEA6F86E" ID-DOI="http://doi.org/10.5281/zenodo.4719928" ID-Zenodo-Dep="4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="3" pageNumber="118" startId="3.[151,250,1826,1850]" targetBox="[243,1345,181,1802]" targetPageId="3">
<paragraph id="C62A36E5FFC2FF92C6ECF8E3FEA6F86E" blockId="3.[151,1436,1826,2030]" pageId="3" pageNumber="118">
<emphasis id="F4E1EAF7FFC2FF92C6ECF8E3FEA2F8BA" bold="true" box="[151,276,1826,1850]" pageId="3" pageNumber="118">FIGURE 2.</emphasis>
<taxonomicName id="01954D66FFC2FF92C767F8E3FE7AF8BA" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[284,460,1826,1850]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFC2FF92C767F8E3FE7AF8BA" box="[284,460,1826,1850]" italics="true" pageId="3" pageNumber="118">Belisana bachma</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFC2FF92C7AFF8E3FD90F8BA" bold="true" box="[468,550,1826,1850]" pageId="3" pageNumber="118">
<taxonomicNameLabel id="EFD2578CFFC2FF92C7AFF8E3FD90F8BA" box="[468,550,1826,1850]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyseson clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH).
</paragraph>
</caption>
<paragraph id="C62A36E5FFC5FF95C6BCFE48FD9DFDF3" blockId="4.[151,1437,151,732]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFE48FEA8FE23" bold="true" box="[199,286,393,419]" pageId="4" pageNumber="119">Female</emphasis>
(IZCAS Ar41903): Similar to male, habitus as in
<figureCitation id="5EAE2A60FFC5FF95C51BFE48FC57FE23" box="[864,993,393,419]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2GH</figureCitation>
. Total length 2.19 (2.32 with clypeus), carapace 0.75 long, 0.63 wide, opisthosoma 1.16 long, 0.81 wide. Tibia I: 3.70; tibia I L/d: 46. Distance PME-PME 0.13, diameter PME 0.08, distance PME-ALE 0.09, AME absent. Sternum approximately as wide as long (0.56). Ocular area without marks. External female genitalia (
<figureCitation id="5EAE2A60FFC5FF95C489FE30FCF8FD8B" box="[754,846,497,523]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2A</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C522FE30FC3CFD8B" box="[857,906,497,523]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14A</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets (arrow ep in
<figureCitation id="5EAE2A60FFC5FF95C7F3FDD5FDB0FDAE" box="[392,518,532,558]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2AB</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C469FDD5FDD1FDAE" box="[530,615,532,558]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14AB</figureCitation>
), 0.48 apart. Vulva (
<figureCitation id="5EAE2A60FFC5FF95C535FDD5FC1DFDAE" box="[846,939,532,558]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C5CCFDD5FC51FDAE" box="[951,999,532,558]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14B</figureCitation>
) with W-shaped anterior arch (median ridge strongly protruding, arrow in
<figureCitation id="5EAE2A60FFC5FF95C459FDF6FDCBFDD1" box="[546,637,567,593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C4F3FDF6FD01FDD1" box="[648,695,567,593]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14B</figureCitation>
) and pair of large, nearly trapezoidal pore plates (arrow pp in
<figureCitation id="5EAE2A60FFC5FF95C317FDF6FF0CFDF3" captionStart="FIGURE 2" captionStartId="3.[151,250,1826,1850]" captionTargetBox="[243,1345,181,1802]" captionTargetId="figure-17@3.[243,1345,181,1802]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bachma sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Prosoma, frontal view, arrow points at pair of small ventral apophyses on clypeus; D. Bulb, prolateral view; EH. Habitus (E. Dorsal view, arrow points at frontal apophysis on clypeus; F. Lateral view; G. Dorsal view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719928" httpUri="https://zenodo.org/record/4719928/files/figure.png" pageId="4" pageNumber="119">Figs 2B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C6BDFD98FF41FDF4" box="[198,247,601,628]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="4" pageNumber="119">16B</figureCitation>
), without serrated sclerites.
</paragraph>
<paragraph id="C62A36E5FFC5FF95C6BCFDBDFCD4FD16" blockId="4.[151,1437,151,732]" box="[199,866,636,663]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFDBDFEF7FD16" bold="true" box="[199,321,636,662]" pageId="4" pageNumber="119">Variation:</emphasis>
Tibia Iin another male (IZCAS Ar41902): 4.47.
</paragraph>
<paragraph id="C62A36E5FFC5FF95C6BCFD5EFABFFD39" blockId="4.[151,1437,151,732]" box="[199,1289,670,697]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFD5EFE33FD38" bold="true" box="[199,389,670,697]" pageId="4" pageNumber="119">Natural history.</emphasis>
The species was found in leaf litter and collected by small plot direct searching.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF95C6BCFD00FC7BFD5C" box="[199,973,705,732]" pageId="4" pageNumber="119" type="distribution">
<paragraph id="C62A36E5FFC5FF95C6BCFD00FC7BFD5C" blockId="4.[151,1437,151,732]" box="[199,973,705,732]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFD00FED7FD5B" bold="true" box="[199,353,705,731]" pageId="4" pageNumber="119">Distribution.</emphasis>
<collectingCountry id="BE827675FFC5FF95C71CFD00FE70FD5B" box="[359,454,705,731]" name="Vietnam" pageId="4" pageNumber="119">Vietnam</collectingCountry>
(Thua Thien Hue,
<typeStatus id="192E8847FFC5FF95C4E0FD03FD7DFD5C" box="[667,715,706,732]" pageId="4" pageNumber="119">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFC5FF95C515FD00FC76FD5C" box="[878,960,705,732]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="4" pageNumber="119">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,300 @@
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<mods:title id="2FBC85BEC0D55BCEFE030FEFDCD7EF90">Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia</mods:title>
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<mods:namePart id="E2325D5E663144A6A385E2D8D7A6B423">Li, Shuqiang</mods:namePart>
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<treatment id="4E3C87F3FFC5FF96C6ECFCE8FC3AFF7A" ID-DOI="http://doi.org/10.5281/zenodo.4696998" ID-GBIF-Taxon="181600553" ID-Zenodo-Dep="4696998" LSID="urn:lsid:plazi:treatment:4E3C87F3FFC5FF96C6ECFCE8FC3AFF7A" httpUri="http://treatment.plazi.org/id/4E3C87F3FFC5FF96C6ECFCE8FC3AFF7A" lastPageId="7" lastPageNumber="122" pageId="4" pageNumber="119">
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<taxonomicName id="01954D66FFC5FF95C6ECFCE8FE39FCC4" ID-CoL="98YXV" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[151,399,809,836]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFC5FF95C6ECFCE8FE39FCC4" bold="true" box="[151,399,809,836]" italics="true" pageId="4" pageNumber="119">Belisana cucphuong</emphasis>
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<paragraph id="C62A36E5FFC5FF95C6ECFC8DFE45FCE7" blockId="4.[151,501,809,871]" box="[151,499,844,871]" pageId="4" pageNumber="119">
<figureCitation id="5EAE2A60FFC5FF95C6ECFC8DFF4DFCE6" box="[151,251,844,870]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,1806,1830]" captionStartId-1="6.[151,250,1875,1899]" captionTargetBox-0="[249,1339,180,1783]" captionTargetBox-1="[226,1361,181,1851]" captionTargetId-0="figure-17@5.[249,1339,180,1783]" captionTargetId-1="figure-19@6.[226,1361,181,1851]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." captionText-1="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.4719932" figureDoi-1="http://doi.org/10.5281/zenodo.4719936" httpUri-0="https://zenodo.org/record/4719932/files/figure.png" httpUri-1="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 34</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C77DFC8DFEECFCE6" box="[262,346,844,870]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13CD</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C71EFC8DFE0FFCE6" box="[357,441,844,870]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14CD</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C7BFFC8DFE45FCE7" box="[452,499,844,871]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="4" pageNumber="119">15B</figureCitation>
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<emphasis id="F4E1EAF7FFC5FF95C6ECFC50FEF3FC2B" bold="true" box="[151,325,913,939]" pageId="4" pageNumber="119">Type material.</emphasis>
<materialsCitation id="76FD3CB8FFC5FF95C736FC50FC73FC4E" ID-GBIF-Occurrence="3091104310" collectingDate="2015-08-20" collectionCode="IZCAS" collectorName="D. S. Pham" country="Vietnam" elevation="437" latitude="20.349533" location="Male" longLatPrecision="1" longitude="105.59334" municipality="Cuc Phuong National Park" pageId="4" pageNumber="119" specimenCount="1" stateProvince="Ninh Binh" typeStatus="holotype">
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<typeStatus id="192E8847FFC5FF95C736FC50FE08FC2B" box="[333,446,913,939]" pageId="4" pageNumber="119" type="holotype">Holotype</typeStatus>
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(
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Ar41904),
<collectingMunicipality id="264EAC9FFFC5FF95C49DFC50FBA5FC2B" box="[742,1043,913,939]" pageId="4" pageNumber="119">Cuc Phuong National Park</collectingMunicipality>
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,
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,
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elevation
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</elevation>
),
<collectingRegion id="0451F807FFC5FF95C71DFC75FE68FC4E" box="[358,478,948,974]" country="Vietnam" name="Ninh Binh" pageId="4" pageNumber="119">Ninh Binh</collectingRegion>
,
<collectingCountry id="BE827675FFC5FF95C791FC75FDFAFC4E" box="[490,588,948,974]" name="Vietnam" pageId="4" pageNumber="119">Vietnam</collectingCountry>
,
<date id="B22B1025FFC5FF95C423FC75FCB8FC4F" box="[600,782,948,975]" pageId="4" pageNumber="119" value="2015-08-20">
<collectingDate id="A26FE9CDFFC5FF95C423FC75FCB8FC4F" box="[600,782,948,975]" pageId="4" pageNumber="119" value="2015-08-20">20 August 2015</collectingDate>
</date>
,
<collectorName id="6B605333FFC5FF95C561FC75FC22FC4E" box="[794,916,948,974]" pageId="4" pageNumber="119">D.S. Pham</collectorName>
leg.
</materialsCitation>
<materialsCitation id="76FD3CB8FFC5FF95C5B7FC75FE59FC71" ID-GBIF-Occurrence="3091104301" collectingDate="2015-08-20" collectionCode="IZCAS" collectorName="D. S. Pham" country="Vietnam" elevation="437" latitude="20.349533" location="Male" longLatPrecision="1" longitude="105.59334" municipality="Cuc Phuong National Park" pageId="4" pageNumber="119" specimenCount="2" specimenCount-female="2" stateProvince="Ninh Binh" typeStatus="paratype">
<emphasis id="F4E1EAF7FFC5FF95C5B7FC75FBFBFC4E" bold="true" box="[972,1101,948,974]" pageId="4" pageNumber="119">
<typeStatus id="192E8847FFC5FF95C5B7FC75FBFEFC4E" box="[972,1096,948,974]" pageId="4" pageNumber="119" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="D093FD6CFFC5FF95C22EFC75FB74FC4E" box="[1109,1218,948,974]" pageId="4" pageNumber="119" type="female">2 females</specimenCount>
(
<collectionCode id="A084AE20FFC5FF95C2A9FC75FA93FC4E" box="[1234,1317,948,974]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="119">IZCAS</collectionCode>
Ar41905 41906), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF95C6BCFC38FBCAFB93" box="[199,1148,1017,1044]" pageId="4" pageNumber="119" type="etymology">
<paragraph id="C62A36E5FFC5FF95C6BCFC38FBCAFB93" blockId="4.[151,1437,913,2015]" box="[199,1148,1017,1044]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFC38FEF8FB93" bold="true" box="[199,334,1017,1043]" pageId="4" pageNumber="119">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="192E8847FFC5FF95C4CEFC3BFD53FB94" box="[693,741,1018,1044]" pageId="4" pageNumber="119">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF95C6BCFBDDFE8EFA87" pageId="4" pageNumber="119" type="diagnosis">
<paragraph id="C62A36E5FFC5FF95C6BCFBDDFE8EFA87" blockId="4.[151,1437,913,2015]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFBDDFEF4FBB6" bold="true" box="[199,322,1052,1078]" pageId="4" pageNumber="119">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFC5FF95C434FBDCFBB6FBB7" authority="Yao, Pham &amp; Li, 2015" authorityName="Yao, Pham &amp; Li" authorityYear="2015" box="[591,1024,1052,1079]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="F4E1EAF7FFC5FF95C434FBDCFD49FBB6" box="[591,767,1053,1078]" italics="true" pageId="4" pageNumber="119">B. vietnamensis</emphasis>
<bibRefCitation id="A2044B14FFC5FF95C57FFBDCFBB6FBB7" author="Yao, Z. &amp; Pham, D. S. &amp; Li, S." box="[772,1024,1052,1079]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="119" pagination="1 - 82" part="3909" refId="ref8999" refString="Yao, Z., Pham, D. S. &amp; Li, S. (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa, 3909 (1), 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1.1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species" type="journal article" year="2015">Yao, Pham &amp; Li, 2015</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFC5FF95C241FBDCFAA6FBB6" author="Yao, Z. &amp; Pham, D. S. &amp; Li, S." box="[1082,1296,1052,1079]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="119" pagination="1 - 82" part="3909" refId="ref8999" refString="Yao, Z., Pham, D. S. &amp; Li, S. (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa, 3909 (1), 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1.1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species" type="journal article" year="2015">
Yao
<emphasis id="F4E1EAF7FFC5FF95C216FBDCFB28FBB6" box="[1133,1182,1052,1078]" italics="true" pageId="4" pageNumber="119">et al</emphasis>
. 2015: 10
</bibRefCitation>
, figs 2931) with the elongated bulbal apophyses (
<figureCitation id="5EAE2A60FFC5FF95C433FBFEFD17FBD9" box="[584,673,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Fig. 4D</figureCitation>
) and the male chelicerae (
<figureCitation id="5EAE2A60FFC5FF95C5AAFBFEFB83FBD9" box="[977,1077,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs. 4C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C238FBFFFBC2FBD9" box="[1091,1140,1086,1113]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="4" pageNumber="119">15B</figureCitation>
), but males can be distinguished by the long distal spine, 7 times as long as wide (arrow
<quantity id="016D9B00FFC5FF95C511FBA0FC20FBFB" box="[874,918,1121,1147]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="119" unit="in" value="2.0">2 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C5E6FBA0FC4EFBFB" box="[925,1016,1121,1147]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C27FFBA0FB83FBFB" box="[1028,1077,1121,1147]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C23AFBA0FBF9FBFB" box="[1089,1103,1121,1147]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">4</figureCitation>
times in
<taxonomicName id="01954D66FFC5FF95C2C3FBA3FADFFBFB" authorityName="Yao, Pham &amp; Li" authorityYear="2015" box="[1208,1385,1122,1147]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="F4E1EAF7FFC5FF95C2C3FBA3FADFFBFB" box="[1208,1385,1122,1147]" italics="true" pageId="4" pageNumber="119">B. vietnamensis</emphasis>
</taxonomicName>
), on the procursus, and by the presence of a long ventral apophysis on the palpal trochanter (arrow
<quantity id="016D9B00FFC5FF95C2ADFB45FAB5FB1E" box="[1238,1283,1156,1182]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="119" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C377FB45FAD4FB1E" box="[1292,1378,1156,1182]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
) and distinct ventral apophysis on palpal femur (arrow
<quantity id="016D9B00FFC5FF95C4A9FB66FD49FB41" box="[722,767,1191,1217]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="4" pageNumber="119" unit="in" value="3.0">3 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C57CFB66FCE9FB41" box="[775,863,1191,1217]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
). Females can be distinguished by the vulval pore plates (arrow pp in
<figureCitation id="5EAE2A60FFC5FF95C70EFB08FE66FB63" box="[373,464,1225,1251]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C7A6FB08FDB9FB63" box="[477,527,1225,1251]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14D</figureCitation>
) being longer than wide (vulval pore plates approximately as wide as long in
<taxonomicName id="01954D66FFC5FF95C3FFFB0BFE9EFA86" authorityName="Yao, Pham &amp; Li" authorityYear="2015" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="119" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="F4E1EAF7FFC5FF95C3FFFB0BFE9EFA86" italics="true" pageId="4" pageNumber="119">B. vietnamensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC5FF96C6BCFACFFB67FF55" lastPageId="7" lastPageNumber="122" pageId="4" pageNumber="119" type="description">
<paragraph id="C62A36E5FFC5FF95C6BCFACFFE23F96C" blockId="4.[151,1437,913,2015]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCFACFFE29FAA9" bold="true" box="[199,415,1294,1321]" pageId="4" pageNumber="119">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFC5FF95C7C9FACEFDAFFAA9" bold="true" box="[434,537,1295,1321]" pageId="4" pageNumber="119">
<typeStatus id="192E8847FFC5FF95C7C9FACEFDAFFAA9" box="[434,537,1295,1321]" pageId="4" pageNumber="119" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41904): Total length 2.22 (2.35 with clypeus), carapace 0.94 long, 0.78 wide, opisthosoma 1.41 long, 0.78 wide. Leg I: (5.70 + 0.38 + 5.77 + + ), leg II: 13.00 (3.97 + 0.32 + 3.33 + 4.10 + 1.28), leg III: 8.28 (2.50 + 0.26 + 1.99 + 2.82 + 0.71), leg IV: 14.15 (6.03 + 0.30 + 2.88 + 3.85 + 1.09); tibia I L/d: 38. Distance PME-PME 0.09, diameter PME 0.09, distance PME-ALE 0.09, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFC5FF95C6ECFA58FEB8FA33" box="[151,270,1433,1459]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4EF</figureCitation>
. Carapace pale, with brown lateral margins. Ocular area not elevated. Clypeus unmodified. Chelicerae (
<figureCitation id="5EAE2A60FFC5FF95C6E4FA7DFEB5FA56" box="[159,259,1468,1494]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs. 4C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C774FA7DFE88FA57" box="[271,318,1468,1495]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="4" pageNumber="119">15B</figureCitation>
) with pair of small proximo-lateral apophyses and pair of short distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.04). Sternum pale, approximately as wide as long (0.50). Thoracic furrow absent. Legs pale. Opisthosoma pale, with large, dark spots. Palps as in
<figureCitation id="5EAE2A60FFC5FF95C362F9C0FA2EF99B" box="[1305,1432,1537,1563]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3AB</figureCitation>
; coxa unmodified; trochanter with long ventral apophysis (arrow
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<figureCitation id="5EAE2A60FFC5FF95C5C9F9E5FBBDF9BE" box="[946,1035,1572,1598]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
); femur with small proximo-dorsal apophysis (arrow
<quantity id="016D9B00FFC5FF95C71BF986FE3DF9E1" box="[352,395,1607,1633]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="119" unit="in" value="2.0">2 in</quantity>
<figureCitation id="5EAE2A60FFC5FF95C7E9F986FE51F9E1" box="[402,487,1607,1633]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
) and distinct ventral apophysis (arrow
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<figureCitation id="5EAE2A60FFC5FF95C5B4F986FB93F9E1" box="[975,1061,1607,1633]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Fig. 3B</figureCitation>
); procursus (
<figureCitation id="5EAE2A60FFC5FF95C2CCF986FA80F9E1" box="[1207,1334,1607,1633]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3AD</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C339F986FA22F9E1" box="[1346,1428,1607,1633]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13CD</figureCitation>
) simple proximally but complex distally, with large subdisto-prolateral sclerite (arrow
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<figureCitation id="5EAE2A60FFC5FF95C2E5F9A8FB4CF903" box="[1182,1274,1641,1667]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C373F9A8FA8FF903" box="[1288,1337,1641,1667]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13C</figureCitation>
), curved distal spine (arrow
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<figureCitation id="5EAE2A60FFC5FF95C7E4F94DFE4FF926" box="[415,505,1676,1702]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3C</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C47EF94DFD82F926" box="[517,564,1676,1702]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13C</figureCitation>
) and subdisto-retrolateral membranous process (arrow in
<figureCitation id="5EAE2A60FFC5FF95C2C0F94DFAA1F926" box="[1211,1303,1676,1702]" captionStart="FIGURE 3" captionStartId="5.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-17@5.[249,1339,180,1783]" captionTargetPageId="5" captionText="FIGURE 3. Belisana cucphuong sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="4" pageNumber="119">Figs 3D</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C359F94DFAE0F926" box="[1314,1366,1676,1702]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="4" pageNumber="119">13D</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFC5FF95C6DBF96EFF40F949" box="[160,246,1711,1737]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Fig. 4D</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia Iat 5% proximally; without spines or curved setae.
</paragraph>
<paragraph id="C62A36E5FFC5FF95C6BCF935FC88F85E" blockId="4.[151,1437,913,2015]" pageId="4" pageNumber="119">
<emphasis id="F4E1EAF7FFC5FF95C6BCF935FEA8F88E" bold="true" box="[199,286,1780,1806]" pageId="4" pageNumber="119">Female</emphasis>
(IZCAS Ar41906): Similar to male, habitus as in
<figureCitation id="5EAE2A60FFC5FF95C51BF935FC57F88E" box="[864,993,1780,1806]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4GH</figureCitation>
. Total length 1.90 (2.00 with clypeus), carapace 0.72 long, 0.66 wide, opisthosoma 1.28 long, 0.66 wide. Tibia I: 4.04; tibia I L/d: 51. Distance PME-PME 0.13, diameter PME 0.06, distance PME-ALE 0.09, AME absent. Sternum slightly wider than long (0.50/0.40). Opisthosoma with indistinct dark spots. External female genitalia (
<figureCitation id="5EAE2A60FFC5FF95C5DBF89DFBB7F8F6" box="[928,1025,1884,1910]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4A</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C26BF89DFBF6F8F6" box="[1040,1088,1884,1910]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14C</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets on sclerotized plate (arrow ep in
<figureCitation id="5EAE2A60FFC5FF95C542F8BEFC00F819" box="[825,950,1919,1945]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4AB</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C5B9F8BEFBA0F819" box="[962,1046,1919,1945]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14CD</figureCitation>
), 0.03 apart. Vulva (
<figureCitation id="5EAE2A60FFC5FF95C280F8BEFAE0F819" box="[1275,1366,1919,1945]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C318F8BEFA22F819" box="[1379,1428,1919,1945]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14D</figureCitation>
) with wavy anterior arch (median ridge strongly protruding, arrow in
<figureCitation id="5EAE2A60FFC5FF95C5E9F860FC5AF83B" box="[914,1004,1953,1979]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C58CF860FB9FF83B" box="[1015,1065,1953,1979]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14D</figureCitation>
) and pair of nearly elliptical pore plates (arrow pp in
<figureCitation id="5EAE2A60FFC5FF95C70AF805FE7DF85E" box="[369,459,1988,2014]" captionStart="FIGURE 4" captionStartId="6.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@6.[226,1361,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Belisana cucphuong sp. nov., holotype male (CF) and paratype female (AB, GH).A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="4" pageNumber="119">Figs 4B</figureCitation>
,
<figureCitation id="5EAE2A60FFC5FF95C7ACF805FDBCF85E" box="[471,522,1988,2014]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="4" pageNumber="119">14D</figureCitation>
), without serrated sclerites.
</paragraph>
<caption id="92EA666DFFC4FF94C6ECF8CFFC26F837" ID-DOI="http://doi.org/10.5281/zenodo.4719932" ID-Zenodo-Dep="4719932" httpUri="https://zenodo.org/record/4719932/files/figure.png" pageId="5" pageNumber="120" startId="5.[151,250,1806,1830]" targetBox="[249,1339,180,1783]" targetPageId="5">
<paragraph id="C62A36E5FFC4FF94C6ECF8CFFC26F837" blockId="5.[151,1437,1806,1975]" pageId="5" pageNumber="120">
<emphasis id="F4E1EAF7FFC4FF94C6ECF8CFFEA5F8A7" bold="true" box="[151,275,1806,1831]" pageId="5" pageNumber="120">FIGURE 3.</emphasis>
<taxonomicName id="01954D66FFC4FF94C762F8CEFE51F8A7" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[281,487,1807,1831]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="120" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFC4FF94C762F8CEFE51F8A7" box="[281,487,1807,1831]" italics="true" pageId="5" pageNumber="120">Belisana cucphuong</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFC4FF94C796F8CEFD88F8A7" bold="true" box="[493,574,1807,1831]" pageId="5" pageNumber="120">
<taxonomicNameLabel id="EFD2578CFFC4FF94C796F8CEFD88F8A7" box="[493,574,1807,1831]" pageId="5" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at long ventral apophysis on palpal trochanter, arrow 2 points at small proximo-dorsal apophysis on palpal femur, arrow 3 points at ventral apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at large subdisto-prolateral sclerite, arrow 2 points at curved distal spine; D. Retrolateral view, arrow points at subdisto-retrolateral membranous process). pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD).
</paragraph>
</caption>
<caption id="92EA666DFFC7FF97C6ECF892FE7FF87B" ID-DOI="http://doi.org/10.5281/zenodo.4719936" ID-Zenodo-Dep="4719936" httpUri="https://zenodo.org/record/4719936/files/figure.png" pageId="6" pageNumber="121" startId="6.[151,250,1875,1899]" targetBox="[226,1361,181,1851]" targetPageId="6">
<paragraph id="C62A36E5FFC7FF97C6ECF892FE7FF87B" blockId="6.[151,1437,1875,2043]" pageId="6" pageNumber="121">
<emphasis id="F4E1EAF7FFC7FF97C6ECF892FEA4F8EB" bold="true" box="[151,274,1875,1899]" pageId="6" pageNumber="121">FIGURE 4.</emphasis>
<taxonomicName id="01954D66FFC7FF97C763F892FE50F8EB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[280,486,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="121" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFC7FF97C763F892FE50F8EB" box="[280,486,1875,1899]" italics="true" pageId="6" pageNumber="121">Belisana cucphuong</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFC7FF97C790F892FD8DF8EB" bold="true" box="[491,571,1875,1899]" pageId="6" pageNumber="121">
<taxonomicNameLabel id="EFD2578CFFC7FF97C790F892FD8DF8EB" box="[491,571,1875,1899]" pageId="6" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at strongly protruding median ridge of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH).
</paragraph>
</caption>
<paragraph id="C62A36E5FFC6FF96C6BCFF57FCCEFF31" blockId="7.[199,1233,150,250]" box="[199,888,150,177]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFF57FEF7FF30" bold="true" box="[199,321,150,176]" pageId="7" pageNumber="122">Variation:</emphasis>
Tibia Iin another female (IZCAS Ar39905): 3.99.
</paragraph>
<paragraph id="C62A36E5FFC6FF96C6BCFF7AFB67FF55" blockId="7.[199,1233,150,250]" box="[199,1233,186,213]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFF7AFE33FF54" bold="true" box="[199,389,186,213]" pageId="7" pageNumber="122">Natural history.</emphasis>
The species was found on domed webs attached to the underside of leaves.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC6FF96C6BCFF1FFC3AFF7A" box="[199,908,222,250]" pageId="7" pageNumber="122" type="distribution">
<paragraph id="C62A36E5FFC6FF96C6BCFF1FFC3AFF7A" blockId="7.[199,1233,150,250]" box="[199,908,222,250]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFF1FFED7FF78" bold="true" box="[199,353,222,248]" pageId="7" pageNumber="122">Distribution.</emphasis>
<collectingCountry id="BE827675FFC6FF96C71CFF1EFE70FF79" box="[359,454,223,249]" name="Vietnam" pageId="7" pageNumber="122">Vietnam</collectingCountry>
(
<collectingRegion id="0451F807FFC6FF96C7AFFF1EFDF8FF79" box="[468,590,223,249]" country="Vietnam" name="Ninh Binh" pageId="7" pageNumber="122">Ninh Binh</collectingRegion>
,
<typeStatus id="192E8847FFC6FF96C422FF1EFD3FFF79" box="[601,649,223,249]" pageId="7" pageNumber="122">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFC6FF96C557FF1EFCC9FF7A" box="[812,895,223,250]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="7" pageNumber="122">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,290 @@
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<emphasis id="F4E1EAF7FFC6FF96C6ECFEFEFEEDFEDA" bold="true" box="[151,347,319,346]" italics="true" pageId="7" pageNumber="122">Belisana jaegeri</emphasis>
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<taxonomicNameLabel id="EFD2578CFFC6FF96C718FE81FE76FEDA" box="[355,448,320,346]" pageId="7" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="C62A36E5FFC6FF96C6ECFEA5FE54FEFF" blockId="7.[151,482,319,383]" box="[151,482,356,383]" pageId="7" pageNumber="122">
<figureCitation id="5EAE2A60FFC6FF96C6ECFEA5FF4CFEFF" box="[151,250,356,383]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="8.[151,250,1875,1899]" captionStartId-1="9.[151,250,1892,1916]" captionTargetBox-0="[226,1361,181,1851]" captionTargetBox-1="[226,1361,199,1869]" captionTargetId-0="figure-19@8.[226,1361,181,1851]" captionTargetId-1="figure-17@9.[226,1361,199,1869]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." captionText-1="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.4719940" figureDoi-1="http://doi.org/10.5281/zenodo.4719942" httpUri-0="https://zenodo.org/record/4719940/files/figure.png" httpUri-1="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">
Figs 5
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<figureCitation id="5EAE2A60FFC6FF96C77DFEA5FEE7FEFE" box="[262,337,356,382]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="7" pageNumber="122">13EF</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C726FEA5FE1EFEFE" box="[349,424,356,382]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="7" pageNumber="122">14EF</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C7C8FEA5FE54FEFF" box="[435,482,356,383]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="7" pageNumber="122">15C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC6FF96C6ECFE6DFD2CFD8E" pageId="7" pageNumber="122" type="materials_examined">
<paragraph id="C62A36E5FFC6FF96C6ECFE6DFD2CFD8E" blockId="7.[151,1437,428,1751]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6ECFE6DFEFFFE46" bold="true" box="[151,329,428,454]" pageId="7" pageNumber="122">Type material.</emphasis>
<materialsCitation id="76FD3CB8FFC6FF96C72FFE6DFAE5FE6A" ID-GBIF-Occurrence="3091104311" collectingDate="2015-02-16" collectionCode="SMF" collectorName="P. Jager &amp; T. Laufs" country="Malaysia" county="Malay Peninsula" elevation="1300" latitude="3.7184167" location="Telecom" longLatPrecision="1" longitude="101.75274" municipality="Fraser's Hill" pageId="7" pageNumber="122" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<emphasis id="F4E1EAF7FFC6FF96C72FFE6DFE7CFE46" bold="true" box="[340,458,428,454]" pageId="7" pageNumber="122">
<typeStatus id="192E8847FFC6FF96C72FFE6DFE73FE46" box="[340,453,428,454]" pageId="7" pageNumber="122" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male (
<collectionCode id="A084AE20FFC6FF96C459FE6DFDE9FE46" box="[546,607,428,454]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="122">SMF</collectionCode>
), secondary forest (
<geoCoordinate id="A3A15022FFC6FF96C53CFE6DFC5FFE46" box="[839,1001,428,454]" degrees="3" direction="north" minutes="43" orientation="latitude" pageId="7" pageNumber="122" precision="1" seconds="06.30" value="3.7184167">3°4306.30”N</geoCoordinate>
,
<geoCoordinate id="A3A15022FFC6FF96C583FE6DFB04FE47" box="[1016,1202,428,455]" degrees="101" direction="east" minutes="45" orientation="longitude" pageId="7" pageNumber="122" precision="1" seconds="09.86" value="101.75274">101°4509.86”E</geoCoordinate>
,
<elevation id="4DB8D1D6FFC6FF96C2BAFE6DFA39FE47" box="[1217,1423,428,455]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="122" unit="m" value="1300.0">
elevation
<quantity id="016D9B00FFC6FF96C348FE6DFA39FE47" box="[1331,1423,428,455]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="122" unit="m" value="1300.0">1300 m</quantity>
</elevation>
),
<location id="C34A603EFFC6FF96C6ECFE11FF41FE6A" LSID="urn:lsid:plazi:treatment:4E3C87F3FFC6FF96C6ECFEFEFC6CF957:C34A603EFFC6FF96C6ECFE11FF41FE6A" box="[151,247,464,490]" country="Malaysia" county="Malay Peninsula" latitude="3.7184167" longLatPrecision="1" longitude="101.75274" municipality="Fraser's Hill" name="Telecom" pageId="7" pageNumber="122" stateProvince="Pahang">Telecom</location>
loop,
<collectingMunicipality id="264EAC9FFFC6FF96C745FE11FE7BFE6A" box="[318,461,464,490]" pageId="7" pageNumber="122">Frasers Hill</collectingMunicipality>
,
<collectingRegion id="0451F807FFC6FF96C7A3FE11FD9BFE6A" box="[472,557,464,490]" country="Malaysia" name="Pahang" pageId="7" pageNumber="122">Pahang</collectingRegion>
,
<collectingCounty id="2F4B4E69FFC6FF96C442FE11FD41FE6A" box="[569,759,464,490]" pageId="7" pageNumber="122">Malay Peninsula</collectingCounty>
,
<collectingCountry id="BE827675FFC6FF96C579FE11FCDDFE6A" box="[770,875,464,490]" name="Malaysia" pageId="7" pageNumber="122">Malaysia</collectingCountry>
,
<date id="B22B1025FFC6FF96C50CFE11FBF7FE6B" box="[887,1089,464,491]" pageId="7" pageNumber="122" value="2015-02-16">
<collectingDate id="A26FE9CDFFC6FF96C50CFE11FBF7FE6B" box="[887,1089,464,491]" pageId="7" pageNumber="122" value="2015-02-16">16 February 2015</collectingDate>
</date>
,
<collectorName id="6B605333FFC6FF96C237FE10FB14FE6B" box="[1100,1186,465,491]" pageId="7" pageNumber="122">P. Jäger</collectorName>
&amp;
<collectorName id="6B605333FFC6FF96C2BEFE10FA94FE6A" box="[1221,1314,464,491]" pageId="7" pageNumber="122">T. Laufs</collectorName>
leg.
</materialsCitation>
<materialsCitation id="76FD3CB8FFC6FF96C322FE11FD20FD8E" ID-GBIF-Occurrence="3091104302" collectingDate="2015-02-16" collectionCode="SMF" collectorName="P. Jager &amp; T. Laufs" country="Malaysia" county="Malay Peninsula" elevation="1300" latitude="3.7184167" location="Telecom" longLatPrecision="1" longitude="101.75274" municipality="Fraser's Hill" pageId="7" pageNumber="122" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="paratype">
<emphasis id="F4E1EAF7FFC6FF96C322FE11FF62FD8E" bold="true" pageId="7" pageNumber="122">
<typeStatus id="192E8847FFC6FF96C322FE11FF78FD8E" pageId="7" pageNumber="122" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="D093FD6CFFC6FF96C6A0FE35FE8AFD8E" box="[219,316,500,526]" pageId="7" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode id="A084AE20FFC6FF96C730FE35FE3EFD8E" box="[331,392,500,526]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="7" pageNumber="122">SMF</collectionCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC6FF96C6BCFDD9FF79FDD7" pageId="7" pageNumber="122" type="etymology">
<paragraph id="C62A36E5FFC6FF96C6BCFDD9FF79FDD7" blockId="7.[151,1437,428,1751]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFDD9FEF8FDB2" bold="true" box="[199,334,536,562]" pageId="7" pageNumber="122">Etymology.</emphasis>
The specific name is a patronym in honor of the collector Peter Jäger; noun (name) in genitive case.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC6FF96C6BCFDA1FC8EFCF7" pageId="7" pageNumber="122" type="diagnosis">
<paragraph id="C62A36E5FFC6FF96C6BCFDA1FC8EFCF7" blockId="7.[151,1437,428,1751]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFDA1FEF4FDFA" bold="true" box="[199,322,608,634]" pageId="7" pageNumber="122">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFC6FF96C419FDA0FCD2FDFB" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[610,868,608,635]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="fraser">
<emphasis id="F4E1EAF7FFC6FF96C419FDA0FD7FFDFA" box="[610,713,608,634]" italics="true" pageId="7" pageNumber="122">B. fraser</emphasis>
<bibRefCitation id="A2044B14FFC6FF96C4AEFDA1FCD2FDFB" author="Huber, B. A." box="[725,868,608,635]" journalOrPublisher="Zoologica" pageId="7" pageNumber="122" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber, 2005</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFC6FF96C5D2FDA1FBD2FDFB" author="Huber, B. A." box="[937,1124,608,635]" journalOrPublisher="Zoologica" pageId="7" pageNumber="122" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber 2005: 51</bibRefCitation>
, figs 69, 320324) and
<taxonomicName id="01954D66FFC6FF96C3FFFDA0FE64FD1E" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="xiaolongha">
<emphasis id="F4E1EAF7FFC6FF96C3FFFDA0FEA3FD1E" italics="true" pageId="7" pageNumber="122">B. xiaolongha</emphasis>
<bibRefCitation id="A2044B14FFC6FF96C765FD45FE64FD1E" author="Zhu, W. &amp; Li, S." box="[286,466,644,670]" journalOrPublisher="Zootaxa" pageId="7" pageNumber="122" pagination="557 - 566" part="4933" refId="ref9059" refString="Zhu, W. &amp; Li, S. (2021) Two new spiders of the genus Belisana (Araneae: Pholcidae) from Xishuangbanna Rainforest, China. Zootaxa, 4933 (4), 557 - 566. https: // doi. org / 10.11646 / zootaxa. 4933.4.6" title="Two new spiders of the genus Belisana (Araneae: Pholcidae) from Xishuangbanna Rainforest, China" type="journal article" year="2021">Zhu &amp; Li, 2021</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFC6FF96C46BFD45FD4FFD1F" author="Zhu, W. &amp; Li, S." box="[528,761,644,671]" journalOrPublisher="Zootaxa" pageId="7" pageNumber="122" pagination="557 - 566" part="4933" refId="ref9059" refString="Zhu, W. &amp; Li, S. (2021) Two new spiders of the genus Belisana (Araneae: Pholcidae) from Xishuangbanna Rainforest, China. Zootaxa, 4933 (4), 557 - 566. https: // doi. org / 10.11646 / zootaxa. 4933.4.6" title="Two new spiders of the genus Belisana (Araneae: Pholcidae) from Xishuangbanna Rainforest, China" type="journal article" year="2021">Zhu &amp; Li 2021: 561</bibRefCitation>
, figs 3
<geoCoordinate id="A3A15022FFC6FF96C52DFD45FC27FD1F" box="[854,913,644,671]" degrees="4, 5" direction="east" orientation="longitude" pageId="7" pageNumber="122" precision="5555" value="4.5">4, 5E</geoCoordinate>
F) with the elongated bulbal apophyses (
<figureCitation id="5EAE2A60FFC6FF96C314FD45FF0BFD43" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Fig. 6D</figureCitation>
), but males can be distinguished by brown marks on the clypeus (
<figureCitation id="5EAE2A60FFC6FF96C5DAFD69FC46FD43" box="[929,1008,680,707]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Fig. 6I</figureCitation>
). They also can be distinguished from
<taxonomicName id="01954D66FFC6FF96C6ECFD0CFF4CFD66" authorityName="Huber" authorityYear="2005" box="[151,250,716,742]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="fraser">
<emphasis id="F4E1EAF7FFC6FF96C6ECFD0CFF4CFD66" box="[151,250,716,742]" italics="true" pageId="7" pageNumber="122">B. fraser</emphasis>
</taxonomicName>
by having the cheliceral distal apophyses directed laterally (arrow da in
<figureCitation id="5EAE2A60FFC6FF96C24EFD0DFB21FD66" box="[1077,1175,716,742]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs. 6C</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C2DFFD0DFB60FD67" box="[1188,1238,716,743]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="7" pageNumber="122">15C</figureCitation>
; distal apophyses directed anteriorly in
<taxonomicName id="01954D66FFC6FF96C7F4FD30FE42FC8A" authorityName="Huber" authorityYear="2005" box="[399,500,752,778]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="fraser">
<emphasis id="F4E1EAF7FFC6FF96C7F4FD30FE42FC8A" box="[399,500,752,778]" italics="true" pageId="7" pageNumber="122">B. fraser</emphasis>
</taxonomicName>
), and females can be distinguished by having frontal “epigynal” pockets laterally on the sclerotized plate (arrow ep in
<figureCitation id="5EAE2A60FFC6FF96C449FCD5FD19FCAE" box="[562,687,788,814]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs 6AB</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C4C0FCD5FCB1FCAE" box="[699,775,788,814]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="7" pageNumber="122">14EF</figureCitation>
) and curved vulval pore plates (arrow pp in
<figureCitation id="5EAE2A60FFC6FF96C281FCD5FAE2FCAE" box="[1274,1364,788,814]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs 6B</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C31BFCD5FA26FCAE" box="[1376,1424,788,814]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="7" pageNumber="122">14F</figureCitation>
). Females differ from
<taxonomicName id="01954D66FFC6FF96C7FFFCF8FD95FCD2" authorityName="Zhu &amp; Li" authorityYear="2021" box="[388,547,824,850]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="xiaolongha">
<emphasis id="F4E1EAF7FFC6FF96C7FFFCF8FD95FCD2" box="[388,547,824,850]" italics="true" pageId="7" pageNumber="122">B. xiaolongha</emphasis>
</taxonomicName>
by the semicircular flap and by a distal spine seen in retrolateral view of the procursus (
<figureCitation id="5EAE2A60FFC6FF96C768FC9DFE29FCF7" box="[275,415,860,887]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Figs 5B, 5D</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C7D0FC9DFE6FFCF6" box="[427,473,860,886]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="7" pageNumber="122">13F</figureCitation>
; narrow flap in
<taxonomicName id="01954D66FFC6FF96C4F1FC9CFC9EFCF6" authorityName="Zhu &amp; Li" authorityYear="2021" box="[650,808,860,886]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="122" phylum="Arthropoda" rank="species" species="xiaolongha">
<emphasis id="F4E1EAF7FFC6FF96C4F1FC9CFC9EFCF6" box="[650,808,860,886]" italics="true" pageId="7" pageNumber="122">B. xiaolongha</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC6FF96C6BCFC41FEF4F932" pageId="7" pageNumber="122" type="description">
<paragraph id="C62A36E5FFC6FF96C6BCFC41FD6CFA13" blockId="7.[151,1437,428,1751]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFC41FE2BFC1A" bold="true" box="[199,413,896,922]" pageId="7" pageNumber="122">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFC6FF96C7D5FC41FDA3FC1A" bold="true" box="[430,533,896,922]" pageId="7" pageNumber="122">
<typeStatus id="192E8847FFC6FF96C7D5FC41FDA3FC1A" box="[430,533,896,922]" pageId="7" pageNumber="122" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 2.03 (2.19 with clypeus), carapace 0.88 long, 0.78 wide, opisthosoma 1.31 long, 0.88 wide. Leg I missing, leg II: 14.71 (3.85 + 0.40 + 3.53 + 5.58 + 1.35), leg III: 9.02 (2.56 + 0.30 + 2.12 + 3.21 + 0.83), leg IV: 12.53 (3.53 + 0.34 + 3.21 + 4.49 + 0.96). Distance PME-PME 0.11, diameter PME 0.06, distance PME-ALE 0.09, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFC6FF96C491FC2DFCE9FB86" box="[746,863,1004,1030]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs 6EF</figureCitation>
. Carapace pale, with brown lateral margins. Ocular area not elevated. Clypeus (
<figureCitation id="5EAE2A60FFC6FF96C7A9FBD1FDA9FBAB" box="[466,543,1040,1067]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Fig. 6I</figureCitation>
) with brown marks, unmodified. Chelicerae (
<figureCitation id="5EAE2A60FFC6FF96C25FFBD1FB30FBAA" box="[1060,1158,1040,1066]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs. 6C</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C2E8FBD1FB75FBAB" box="[1171,1219,1040,1067]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="7" pageNumber="122">15C</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.62). Sternum pale, slightly wider than long (0.63/0.56). Thoracic furrow absent. Legs pale, but dark brown on femur, patellae and tibia-metatarsus joints, without darker rings. Opisthosoma pale, without spots. Palps as in
<figureCitation id="5EAE2A60FFC6FF96C7C6FB61FD8AFB3B" box="[445,572,1184,1211]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Figs 5AB</figureCitation>
; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur with small ventral hump (arrow
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<figureCitation id="5EAE2A60FFC6FF96C44CFB05FD3BFB5F" box="[567,653,1220,1247]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Fig. 5B</figureCitation>
) and small proximo-dorsal apophysis (arrow
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<figureCitation id="5EAE2A60FFC6FF96C2B9FB05FAACFB5F" box="[1218,1306,1220,1247]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Fig. 5B</figureCitation>
); procursus (
<figureCitation id="5EAE2A60FFC6FF96C6E4FB29FEA9FA83" box="[159,287,1256,1283]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Figs 5AD</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C750FB29FECEFA82" box="[299,376,1256,1282]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="7" pageNumber="122">13EF</figureCitation>
) simple proximally but complex distally, with curved subdisto-dorsal sclerite (arrow
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<figureCitation id="5EAE2A60FFC6FF96C317FB29FF0CFAA7" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Figs 5C</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C6BCFACDFF41FAA6" box="[199,247,1292,1318]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="7" pageNumber="122">13E</figureCitation>
), long disto-prolateral spine (arrow
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<figureCitation id="5EAE2A60FFC6FF96C4BCFACDFC95FAA7" box="[711,803,1292,1319]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Figs 5C</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C54BFACDFCD6FAA6" box="[816,864,1292,1318]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="7" pageNumber="122">13E</figureCitation>
), long subdisto-prolateral sclerite (arrow
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<figureCitation id="5EAE2A60FFC6FF96C317FACDFF0CFACB" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Figs 5C</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C6BEFAF1FF43FACA" box="[197,245,1328,1354]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="7" pageNumber="122">13E</figureCitation>
), disto-retrolateral membranous lamella (arrow in
<figureCitation id="5EAE2A60FFC6FF96C553FAF1FC32FACB" box="[808,900,1328,1355]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Figs 5D</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C5F4FAF1FC0AFACA" box="[911,956,1328,1354]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="7" pageNumber="122">13F</figureCitation>
) and semicircular retrolateral membranous flap (arrow f in
<figureCitation id="5EAE2A60FFC6FF96C732FA95FE13FAEF" box="[329,421,1364,1391]" captionStart="FIGURE 5" captionStartId="8.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@8.[226,1361,181,1851]" captionTargetPageId="8" captionText="FIGURE 5. Belisana jaegeri sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small ventral hump on palpal femur, arrow 2 points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdisto-dorsal sclerite, arrow 2 points at long disto-prolateral spine, arrow 3 points at long subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719940" httpUri="https://zenodo.org/record/4719940/files/figure.png" pageId="7" pageNumber="122">Figs 5D</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C7CBFA95FE68FAEE" box="[432,478,1364,1390]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="7" pageNumber="122">13F</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFC6FF96C44EFA95FD3AFAEF" box="[565,652,1364,1391]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Fig. 6D</figureCitation>
) with hooked apophysis and simple embolus. Legs with short vertical setae on metatarsi, without spines and curved setae.
</paragraph>
<paragraph id="C62A36E5FFC6FF96C6BCFA5DFCC4F9EA" blockId="7.[151,1437,428,1751]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCFA5DFEA8FA36" bold="true" box="[199,286,1436,1462]" pageId="7" pageNumber="122">Female</emphasis>
: Similar to male, habitus as in
<figureCitation id="5EAE2A60FFC6FF96C4F9FA5DFCB4FA36" box="[642,770,1436,1462]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs 6GH</figureCitation>
. Total length 2.22 (2.31 with clypeus), carapace 0.84 long, 0.88 wide, opisthosoma 1.47 long, 0.91 wide. Tibia I: 4.17; tibia I L/d: 42. Distance PME-PME 0.16, diameter PME 0.09, distance PME-ALE 0.09, AME absent. Sternum slightly wider than long (0.59/0.50). Clypeus pale, without marks. External female genitalia (
<figureCitation id="5EAE2A60FFC6FF96C475F9C9FDDDF9A2" box="[526,619,1544,1570]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs 6A</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C40EF9C9FD12F9A2" box="[629,676,1544,1570]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="7" pageNumber="122">14E</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets (arrow ep in
<figureCitation id="5EAE2A60FFC6FF96C6ECF9EDFEA2F9C6" box="[151,276,1580,1606]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs 6AB</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C75BF9EDFED9F9C6" box="[288,367,1580,1606]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="7" pageNumber="122">14EF</figureCitation>
), 0.54 apart. Vulva (
<figureCitation id="5EAE2A60FFC6FF96C428F9EDFD19F9C6" box="[595,687,1580,1606]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs 6B</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C4C0F9EDFD51F9C6" box="[699,743,1580,1606]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="7" pageNumber="122">14F</figureCitation>
) with pointed anterior arch and pair of large, curved and oval pore plates (arrow pp in
<figureCitation id="5EAE2A60FFC6FF96C7D1F991FDB2F9EA" box="[426,516,1616,1642]" captionStart="FIGURE 6" captionStartId="9.[151,250,1892,1916]" captionTargetBox="[226,1361,199,1869]" captionTargetId="figure-17@9.[226,1361,199,1869]" captionTargetPageId="9" captionText="FIGURE 6. Belisana jaegeri sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view); I. Clypeus, frontal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AD, I), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719942" httpUri="https://zenodo.org/record/4719942/files/figure.png" pageId="7" pageNumber="122">Figs 6B</figureCitation>
,
<figureCitation id="5EAE2A60FFC6FF96C46BF991FD88F9EA" box="[528,574,1616,1642]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="7" pageNumber="122">14F</figureCitation>
), without serrated sclerites.
</paragraph>
<paragraph id="C62A36E5FFC6FF96C6BCF9B5FEF4F932" blockId="7.[151,1437,428,1751]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCF9B5FE33F90E" bold="true" box="[199,389,1652,1678]" pageId="7" pageNumber="122">Natural history.</emphasis>
The species was found in secondary forest, on embankments along the road at night and collected by hand.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFC6FF96C6BCF97DFC6CF957" box="[199,986,1724,1751]" pageId="7" pageNumber="122" type="distribution">
<paragraph id="C62A36E5FFC6FF96C6BCF97DFC6CF957" blockId="7.[151,1437,428,1751]" box="[199,986,1724,1751]" pageId="7" pageNumber="122">
<emphasis id="F4E1EAF7FFC6FF96C6BCF97DFED7F956" bold="true" box="[199,353,1724,1750]" pageId="7" pageNumber="122">Distribution.</emphasis>
<collectingCountry id="BE827675FFC6FF96C71CF97DFE78F956" box="[359,462,1724,1750]" name="Malaysia" pageId="7" pageNumber="122">Malaysia</collectingCountry>
(Malay Peninsula,
<typeStatus id="192E8847FFC6FF96C4D3F97CFD6EF957" box="[680,728,1725,1751]" pageId="7" pageNumber="122">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFC6FF96C500F97DFC7BF957" box="[891,973,1724,1751]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="7" pageNumber="122">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,270 @@
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<mods:title id="AB39FDD08F6516AC8BDD3EF4136E449D">Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia</mods:title>
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<emphasis id="F4E1EAF7FFCBFF9BC6ECFF56FEEDFF32" bold="true" box="[151,347,151,178]" italics="true" pageId="10" pageNumber="125">Belisana kachin</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFD2578CFFCBFF9BC718FF56FE76FF31" box="[355,448,151,177]" pageId="10" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="8E8F656EFFCBFF9BC6ECFF7AFE4EFF56" box="[151,504,187,214]" pageId="10" pageNumber="125" type="description">
<paragraph id="C62A36E5FFCBFF9BC6ECFF7AFE4EFF56" blockId="10.[151,504,151,214]" box="[151,504,187,214]" pageId="10" pageNumber="125">
<figureCitation id="5EAE2A60FFCBFF9BC6ECFF7AFF4DFF55" box="[151,251,187,213]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="11.[151,250,1875,1899]" captionStartId-1="12.[151,250,1875,1899]" captionTargetBox-0="[226,1361,181,1851]" captionTargetBox-1="[226,1361,181,1851]" captionTargetId-0="figure-17@11.[226,1361,181,1851]" captionTargetId-1="figure-19@12.[226,1361,181,1851]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 7. Belisana kachin sp. nov., holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." captionText-1="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.4719944" figureDoi-1="http://doi.org/10.5281/zenodo.4719947" httpUri-0="https://zenodo.org/record/4719944/files/figure.png" httpUri-1="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 78</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC77DFF7AFEEAFF55" box="[262,348,187,213]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13GH</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC71CFF7AFE0AFF55" box="[359,444,187,213]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14GH</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC7BCFF7AFE4EFF56" box="[455,504,187,214]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6ECFEC2FB23FEE5" pageId="10" pageNumber="125" type="materials_examined">
<paragraph id="C62A36E5FFCBFF9BC6ECFEC2FB23FEE5" blockId="10.[151,1437,259,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6ECFEC2FEFDFE9D" bold="true" box="[151,331,259,285]" pageId="10" pageNumber="125">Type material.</emphasis>
<materialsCitation id="76FD3CB8FFCBFF9BC72CFEC2FED0FEE5" ID-GBIF-Occurrence="3091104304" collectingDate="2017-05-12" collectionCode="IZCAS" collectorName="Z. Chen &amp; J. Wu" country="Myanmar" county="Putao" elevation="1171" latitude="27.577435" location="Roadside" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" pageId="10" pageNumber="125" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="F4E1EAF7FFCBFF9BC72CFEC2FE78FE9D" bold="true" box="[343,462,259,285]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC72CFEC2FE7EFE9D" box="[343,456,259,285]" pageId="10" pageNumber="125" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male (
<collectionCode id="A084AE20FFCBFF9BC453FEC2FDCDFE9D" box="[552,635,259,285]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="10" pageNumber="125">IZCAS</collectionCode>
Ar41907),
<location id="C34A603EFFCBFF9BC57CFEC2FCD9FE9D" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9BC6ECFF56FCC0FA42:C34A603EFFCBFF9BC57CFEC2FCD9FE9D" box="[775,879,259,285]" country="Myanmar" county="Putao" latitude="27.577435" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" name="Roadside" pageId="10" pageNumber="125" stateProvince="Kachin">Roadside</location>
between
<location id="C34A603EFFCBFF9BC59DFEC2FB9FFE9D" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9BC6ECFF56FCC0FA42:C34A603EFFCBFF9BC59DFEC2FB9FFE9D" box="[998,1065,259,285]" country="Myanmar" county="Putao" latitude="27.577435" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" name="Camp" pageId="10" pageNumber="125" stateProvince="Kachin">Camp</location>
2 to
<location id="C34A603EFFCBFF9BC208FEC2FB65FE9D" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9BC6ECFF56FCC0FA42:C34A603EFFCBFF9BC208FEC2FB65FE9D" box="[1139,1235,259,285]" country="Myanmar" county="Putao" latitude="27.577435" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" name="Ziradum" pageId="10" pageNumber="125" stateProvince="Kachin">Ziradum</location>
(
<geoCoordinate id="A3A15022FFCBFF9BC293FEC2FA21FE9D" box="[1256,1431,259,285]" degrees="27" direction="north" minutes="34" orientation="latitude" pageId="10" pageNumber="125" precision="1" seconds="38.76" value="27.577435">27°3438.76”N</geoCoordinate>
,
<geoCoordinate id="A3A15022FFCBFF9BC6ECFEE6FEF5FEC1" box="[151,323,295,321]" degrees="97" direction="east" minutes="04" orientation="longitude" pageId="10" pageNumber="125" precision="1" seconds="32.64" value="97.07573">97°0432.64”E</geoCoordinate>
,
<elevation id="4DB8D1D6FFCBFF9BC72BFEE6FDACFEC2" box="[336,538,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.1709999999999998" pageId="10" pageNumber="125" unit="m" value="1171.0">
elevation
<quantity id="016D9B00FFCBFF9BC7BBFEE6FDACFEC2" box="[448,538,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.1709999999999998" pageId="10" pageNumber="125" unit="m" value="1171.0">1171 m</quantity>
</elevation>
),
<collectingMunicipality id="264EAC9FFFCBFF9BC44BFEE6FC16FEC1" box="[560,928,295,321]" pageId="10" pageNumber="125">Hponkanrazi Wildlife Sanctuary</collectingMunicipality>
,
<collectingCounty id="2F4B4E69FFCBFF9BC5D6FEE9FC46FEC2" box="[941,1008,296,322]" pageId="10" pageNumber="125">Putao</collectingCounty>
,
<collectingRegion id="0451F807FFCBFF9BC586FEE6FBE7FEC1" box="[1021,1105,295,321]" country="Myanmar" name="Kachin" pageId="10" pageNumber="125">Kachin</collectingRegion>
,
<collectingCountry id="BE827675FFCBFF9BC225FEE9FB7BFEC2" box="[1118,1229,296,322]" name="Myanmar" pageId="10" pageNumber="125">Myanmar</collectingCountry>
,
<date id="B22B1025FFCBFF9BC2A1FEE6FAC1FEC1" box="[1242,1399,295,322]" pageId="10" pageNumber="125" value="2017-05-12">
<collectingDate id="A26FE9CDFFCBFF9BC2A1FEE6FAC1FEC1" box="[1242,1399,295,322]" pageId="10" pageNumber="125" value="2017-05-12">12 May 2017</collectingDate>
</date>
,
<collectorName id="6B605333FFCBFF9BC3FFFEE9FF64FEE5" pageId="10" pageNumber="125">Z. Chen</collectorName>
&amp;
<collectorName id="6B605333FFCBFF9BC68DFE8DFE83FEE6" box="[246,309,332,358]" pageId="10" pageNumber="125">J. Wu</collectorName>
leg.
</materialsCitation>
<materialsCitation id="76FD3CB8FFCBFF9BC716FE8AFB27FEE5" ID-GBIF-Occurrence="3091104305" box="[365,1169,331,358]" collectingDate="2017-05-12" collectionCode="IZCAS" collectorName="Z. Chen &amp; J. Wu" country="Myanmar" county="Putao" elevation="1171" latitude="27.577435" location="Roadside" longLatPrecision="1" longitude="97.07573" municipality="Hponkanrazi Wildlife Sanctuary" pageId="10" pageNumber="125" specimenCount="2" specimenCount-female="2" stateProvince="Kachin" typeStatus="paratype">
<emphasis id="F4E1EAF7FFCBFF9BC716FE8AFE58FEE5" bold="true" box="[365,494,331,357]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC716FE8AFE5FFEE5" box="[365,489,331,357]" pageId="10" pageNumber="125" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="D093FD6CFFCBFF9BC78EFE8AFDD7FEE5" box="[501,609,331,357]" pageId="10" pageNumber="125" type="female">2 females</specimenCount>
(
<collectionCode id="A084AE20FFCBFF9BC40BFE8AFD75FEE5" box="[624,707,331,357]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="10" pageNumber="125">IZCAS</collectionCode>
Ar4190841909), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCFEAEFBCAFE09" box="[199,1148,367,394]" pageId="10" pageNumber="125" type="etymology">
<paragraph id="C62A36E5FFCBFF9BC6BCFEAEFBCAFE09" blockId="10.[151,1437,259,1474]" box="[199,1148,367,394]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFEAEFEF8FE09" bold="true" box="[199,334,367,393]" pageId="10" pageNumber="125">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="192E8847FFCBFF9BC4CEFEB1FD53FE0A" box="[693,741,368,394]" pageId="10" pageNumber="125">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCFE52FD2EFDE1" pageId="10" pageNumber="125" type="diagnosis">
<paragraph id="C62A36E5FFCBFF9BC6BCFE52FD2EFDE1" blockId="10.[151,1437,259,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFE52FEF4FE2D" bold="true" box="[199,322,403,429]" pageId="10" pageNumber="125">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFCBFF9BC429FE55FCEDFE2E" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[594,859,403,430]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="nujiang">
<emphasis id="F4E1EAF7FFCBFF9BC429FE55FD7EFE2D" box="[594,712,404,429]" italics="true" pageId="10" pageNumber="125">B. nujiang</emphasis>
<bibRefCitation id="A2044B14FFCBFF9BC4B4FE52FCEDFE2E" author="Huber, B. A." box="[719,859,403,430]" journalOrPublisher="Zoologica" pageId="10" pageNumber="125" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber, 2005</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFCBFF9BC5ECFE52FBFCFE2D" author="Huber, B. A." box="[919,1098,403,430]" journalOrPublisher="Zoologica" pageId="10" pageNumber="125" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber 2005: 14</bibRefCitation>
, figs 7, 8, 57, 8387, 93111) with the elongated distal apophyses on the male chelicerae (
<figureCitation id="5EAE2A60FFCBFF9BC531FE76FC18FE51" box="[842,942,439,465]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs. 8C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC5C6FE76FC58FE52" box="[957,1006,439,466]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15D</figureCitation>
) and bulbal apophyses (
<figureCitation id="5EAE2A60FFCBFF9BC37DFE76FAD7FE51" box="[1286,1377,439,465]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Fig. 8D</figureCitation>
), but males can be distinguished by the absence of a distal spine and retrolateral flap on the procursus (
<figureCitation id="5EAE2A60FFCBFF9BC28FFE1AFAF8FE76" box="[1268,1358,475,502]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Fig. 7D</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC327FE1AFA26FE75" box="[1372,1424,475,501]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13H</figureCitation>
). Females can be distinguished from other species by the external genitalia with an anterior arch extending to the posterior part (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC700FDE2FE63FDBD" box="[379,469,547,573]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC79AFDE2FDA3FDBD" box="[481,533,547,573]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
; anterior arch extending to the median part in
<taxonomicName id="01954D66FFCBFF9BC263FDE5FB3BFDBD" authorityName="Huber" authorityYear="2005" box="[1048,1165,548,573]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="nujiang">
<emphasis id="F4E1EAF7FFCBFF9BC263FDE5FB3BFDBD" box="[1048,1165,548,573]" italics="true" pageId="10" pageNumber="125">B. nujiang</emphasis>
</taxonomicName>
) and the different shape of the pore plates (arrow pp in
<figureCitation id="5EAE2A60FFCBFF9BC78AFD86FDFDFDE1" box="[497,587,583,609]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC42CFD86FD3DFDE1" box="[599,651,583,609]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCFDAAFB67FA1D" pageId="10" pageNumber="125" type="description">
<paragraph id="C62A36E5FFCBFF9BC6BCFDAAFC7DFBD9" blockId="10.[151,1437,259,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFDAAFE29FD05" bold="true" box="[199,415,619,645]" pageId="10" pageNumber="125">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFCBFF9BC7C9FDAAFDAFFD05" bold="true" box="[434,537,619,645]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC7C9FDAAFDAFFD05" box="[434,537,619,645]" pageId="10" pageNumber="125" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41907): Total length 1.70 (1.76 with clypeus), carapace 0.64 long, 0.62 wide, opisthosoma 1.12 long, 0.60 wide. Leg I: 16.34 (4.49 + 0.25 + 4.10 + 6.35 + 1.15), leg II: 11.13 (3.40 + 0.23 + 2.82 + 3.85 + 0.83), leg III: 7.44 (2.18 + 0.20 + 1.92 + 2.56 + 0.58), leg IV: 9.70 (2.88 + 0.21 + 2.69 + 3.21 + 0.71); tibia I L/d: 43. Distance PME-PME 0.19, diameter PME 0.06, distance PME-ALE 0.08, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFCBFF9BC6AAFD3AFEF3FC95" box="[209,325,763,789]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8EF</figureCitation>
. Carapace pale. Ocular area not elevated. Clypeus unmodified. Chelicerae (
<figureCitation id="5EAE2A60FFCBFF9BC2F7FD3AFB5BFC95" box="[1164,1261,763,789]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs. 8C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC282FD3AFA9CFC96" box="[1273,1322,763,790]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15D</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.44). Sternum pale, slightly wider than long (0.53/0.52). Thoracic furrow absent. Legs pale. Opisthosoma pale, without spots. Palps as in
<figureCitation id="5EAE2A60FFCBFF9BC5E1FCA6FBAEFC01" box="[922,1048,871,897]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7AB</figureCitation>
; coxa unmodified; trochanter with ventral apophysis (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC7A7FC4AFD80FC26" box="[476,566,907,934]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Fig. 7B</figureCitation>
); femur without proximo-dorsal apophysis; procursus (
<figureCitation id="5EAE2A60FFCBFF9BC2CBFC4AFA85FC25" box="[1200,1331,907,933]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7AD</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC33BFC4AFA22FC25" box="[1344,1428,907,933]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13GH</figureCitation>
) simple proximally but complex distally, with curved subdisto-dorsal sclerite (arrow
<quantity id="016D9B00FFCBFF9BC23BFC6EFBDDFC49" box="[1088,1131,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="125" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFCBFF9BC209FC6EFB7DFC49" box="[1138,1227,943,969]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC2ACFC6EFABCFC49" box="[1239,1290,943,969]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13G</figureCitation>
), large distoprolateral sclerite (arrow
<quantity id="016D9B00FFCBFF9BC7C1FC12FE51FC6D" box="[442,487,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="125" unit="in" value="2.0">2 in</quantity>
<figureCitation id="5EAE2A60FFCBFF9BC78BFC12FDFAFC6D" box="[496,588,979,1005]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC420FC12FD38FC6D" box="[603,654,979,1005]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13G</figureCitation>
), inverted L-shaped subdisto-prolateral sclerite (arrow
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<figureCitation id="5EAE2A60FFCBFF9BC340FC12FA21FC6D" box="[1339,1431,979,1005]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC6ECFC36FF7EFB91" box="[151,200,1015,1041]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13G</figureCitation>
) and disto-retrolateral membranous lamella (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC564FC36FCCFFB92" box="[799,889,1015,1042]" captionStart="FIGURE 7" captionStartId="11.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@11.[226,1361,181,1851]" captionTargetPageId="11" captionText="FIGURE 7. Belisana kachin sp. nov., holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="10" pageNumber="125">Figs 7D</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC5FFFC36FC01FB91" box="[900,951,1015,1041]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13H</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFCBFF9BC271FC36FBD6FB91" box="[1034,1120,1015,1041]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Fig. 8D</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia Iat 5% proximally; legs with short vertical setae on metatarsi, without spines or curved setae; tarsus Iwith 9 distinct pseudopodomeres.
</paragraph>
<paragraph id="C62A36E5FFCBFF9BC6BCFBA2FE01FAD5" blockId="10.[151,1437,259,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFBA2FEA8FBFD" bold="true" box="[199,286,1123,1149]" pageId="10" pageNumber="125">Female</emphasis>
(IZCAS Ar41909): Similar to male, habitus as in
<figureCitation id="5EAE2A60FFCBFF9BC51BFBA2FC57FBFD" box="[864,993,1123,1149]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8GH</figureCitation>
. Total length 1.60 (1.66 with clypeus), carapace 0.60 long, 0.52 wide, opisthosoma 1.06 long, 0.60 wide. Tibia I: 1.75; tibia I L/d: 21. Distance PME-PME 0.13, diameter PME 0.05, distance PME-ALE 0.04, AME absent. Sternum slightly wider than long (0.44/0.42). External female genitalia (
<figureCitation id="5EAE2A60FFCBFF9BC7C6FB0EFDACFB69" box="[445,538,1231,1257]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8A</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC45EFB0EFDE1FB69" box="[549,599,1231,1257]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14G</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets laterally on sclerotized plate (arrow ep in
<figureCitation id="5EAE2A60FFCBFF9BC7DBFB32FDABFA8D" box="[416,541,1267,1293]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8AB</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC453FB32FDC8FA8D" box="[552,638,1267,1293]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14GH</figureCitation>
), 0.42 apart. Vulva (
<figureCitation id="5EAE2A60FFCBFF9BC524FB32FC0DFA8D" box="[863,955,1267,1293]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC5BDFB32FC41FA8D" box="[966,1015,1267,1293]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
) with pointed anterior arch extending to posterior part, with scaly edge (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC406FAD6FD61FAB1" box="[637,727,1303,1329]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC498FAD6FCA2FAB1" box="[739,788,1303,1329]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
) and pair of large pore plates (arrow pp in
<figureCitation id="5EAE2A60FFCBFF9BC28DFAD6FAE6FAB1" box="[1270,1360,1303,1329]" captionStart="FIGURE 8" captionStartId="12.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-19@12.[226,1361,181,1851]" captionTargetPageId="12" captionText="FIGURE 8. Belisana kachin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="10" pageNumber="125">Figs 8B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC327FAD6FA39FAB1" box="[1372,1423,1303,1329]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="10" pageNumber="125">14H</figureCitation>
), without serrated sclerites.
</paragraph>
<paragraph id="C62A36E5FFCBFF9BC6BCFA9EFCD5FAF9" blockId="10.[151,1437,259,1474]" box="[199,867,1375,1402]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFA9EFEF7FAF9" bold="true" box="[199,321,1375,1401]" pageId="10" pageNumber="125">Variation:</emphasis>
Tibia Iin another male (IZCAS Ar41908): 4.04.
</paragraph>
<paragraph id="C62A36E5FFCBFF9BC6BCFA42FB67FA1D" blockId="10.[151,1437,259,1474]" box="[199,1233,1411,1438]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFA42FE33FA1D" bold="true" box="[199,389,1411,1437]" pageId="10" pageNumber="125">Natural history.</emphasis>
The species was found on domed webs attached to the underside of leaves.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCFA66FCC0FA42" box="[199,886,1447,1474]" pageId="10" pageNumber="125" type="distribution">
<paragraph id="C62A36E5FFCBFF9BC6BCFA66FCC0FA42" blockId="10.[151,1437,259,1474]" box="[199,886,1447,1474]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCFA66FED7FA41" bold="true" box="[199,353,1447,1473]" pageId="10" pageNumber="125">Distribution.</emphasis>
<collectingCountry id="BE827675FFCBFF9BC71CFA69FE62FA42" box="[359,468,1448,1474]" name="Myanmar" pageId="10" pageNumber="125">Myanmar</collectingCountry>
(
<collectingRegion id="0451F807FFCBFF9BC798FA66FD8EFA41" box="[483,568,1447,1473]" country="Myanmar" name="Kachin" pageId="10" pageNumber="125">Kachin</collectingRegion>
,
<typeStatus id="192E8847FFCBFF9BC438FA69FDC5FA42" box="[579,627,1448,1474]" pageId="10" pageNumber="125">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFCBFF9BC56DFA66FCDFFA42" box="[790,873,1447,1474]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="10" pageNumber="125">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,269 @@
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<taxonomicName id="01954D66FFCBFF9BC6ECF9D2FEFAF9AD" ID-CoL="98YXX" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[151,332,1555,1582]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="putao" status="sp. nov.">
<emphasis id="F4E1EAF7FFCBFF9BC6ECF9D2FEFAF9AD" bold="true" box="[151,332,1555,1582]" italics="true" pageId="10" pageNumber="125">Belisana putao</emphasis>
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<paragraph id="C62A36E5FFCBFF9BC6ECF9F6FE38F9D2" blockId="10.[151,434,1555,1618]" box="[151,398,1591,1618]" pageId="10" pageNumber="125">
<figureCitation id="5EAE2A60FFCBFF9BC6ECF9F6FEBFF9D1" box="[151,265,1591,1617]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="13.[151,250,1875,1899]" captionStartId-1="14.[151,250,1910,1934]" captionTargetBox-0="[226,1361,181,1851]" captionTargetBox-1="[214,1374,181,1886]" captionTargetId-0="figure-17@13.[226,1361,181,1851]" captionTargetId-1="figure-19@14.[214,1374,181,1886]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." captionText-1="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (AB), 0.20 (CE)." figureDoi-0="http://doi.org/10.5281/zenodo.4719949" figureDoi-1="http://doi.org/10.5281/zenodo.4719951" httpUri-0="https://zenodo.org/record/4719949/files/figure.png" httpUri-1="https://zenodo.org/record/4719951/files/figure.png" pageId="10" pageNumber="125">Figs 910</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC76FF9F6FEE3F9D1" box="[276,341,1591,1617]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13IJ</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC71AF9F6FE38F9D2" box="[353,398,1591,1618]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6ECF9BEFB64F93D" pageId="10" pageNumber="125" type="materials_examined">
<paragraph id="C62A36E5FFCBFF9BC6ECF9BEFB64F93D" blockId="10.[151,1437,1663,2014]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6ECF9BEFEFEF919" bold="true" box="[151,328,1663,1689]" pageId="10" pageNumber="125">Type material.</emphasis>
<materialsCitation id="76FD3CB8FFCBFF9BC728F9BEFB64F93D" ID-GBIF-Occurrence="3091104312" collectingDate="2017-05-07" collectionCode="IZCAS" collectorName="Z. Chen &amp; J. Wu" country="Myanmar" elevation="874" latitude="27.496883" location="Male" longLatPrecision="1" longitude="97.17108" pageId="10" pageNumber="125" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="F4E1EAF7FFCBFF9BC728F9BEFE7FF919" bold="true" box="[339,457,1663,1689]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC728F9BEFE72F919" box="[339,452,1663,1689]" pageId="10" pageNumber="125" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="C34A603EFFCBFF9BC7AFF9BEFDB8F919" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCBFF9EC6ECF9D2FCC0FD3A:C34A603EFFCBFF9BC7AFF9BEFDB8F919" box="[468,526,1663,1689]" country="Myanmar" latitude="27.496883" longLatPrecision="1" longitude="97.17108" name="Male" pageId="10" pageNumber="125" stateProvince="Kachin">Male</location>
(
<collectionCode id="A084AE20FFCBFF9BC464F9BEFDC4F919" box="[543,626,1663,1689]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="10" pageNumber="125">IZCAS</collectionCode>
Ar41910), roadside
<locationDeviation id="D7982FC7FFCBFF9BC519F9BEFB63F919" box="[866,1237,1663,1689]" firstLocation="Wasadum" pageId="10" pageNumber="125" secondLocation="Ziradum">between Wasadum and Ziradum</locationDeviation>
(
<geoCoordinate id="A3A15022FFCBFF9BC293F9BEFA21F919" box="[1256,1431,1663,1689]" degrees="27" direction="north" minutes="29" orientation="latitude" pageId="10" pageNumber="125" precision="1" seconds="48.78" value="27.496883">27°2948.78”N</geoCoordinate>
,
<geoCoordinate id="A3A15022FFCBFF9BC6ECF962FEF5F93E" box="[151,323,1699,1726]" degrees="97" direction="east" minutes="10" orientation="longitude" pageId="10" pageNumber="125" precision="1" seconds="15.90" value="97.17108">97°1015.90”E</geoCoordinate>
,
<elevation id="4DB8D1D6FFCBFF9BC735F962FDB0F93E" box="[334,518,1699,1726]" metricMagnitude="2" metricUnit="m" metricValue="8.74" pageId="10" pageNumber="125" unit="m" value="874.0">
elevation
<quantity id="016D9B00FFCBFF9BC7C7F962FDB0F93E" box="[444,518,1699,1726]" metricMagnitude="2" metricUnit="m" metricValue="8.74" pageId="10" pageNumber="125" unit="m" value="874.0">874 m</quantity>
</elevation>
), Putao,
<collectingRegion id="0451F807FFCBFF9BC413F962FD0AF93D" box="[616,700,1699,1725]" country="Myanmar" name="Kachin" pageId="10" pageNumber="125">Kachin</collectingRegion>
,
<collectingCountry id="BE827675FFCBFF9BC4BCF965FC80F93E" box="[711,822,1700,1726]" name="Myanmar" pageId="10" pageNumber="125">Myanmar</collectingCountry>
,
<date id="B22B1025FFCBFF9BC53BF962FC6FF93D" box="[832,985,1699,1726]" pageId="10" pageNumber="125" value="2017-05-07">
<collectingDate id="A26FE9CDFFCBFF9BC53BF962FC6FF93D" box="[832,985,1699,1726]" pageId="10" pageNumber="125" value="2017-05-07">07 May 2017</collectingDate>
</date>
,
<collectorName id="6B605333FFCBFF9BC59FF965FB88F93D" box="[996,1086,1699,1726]" pageId="10" pageNumber="125">Z. Chen</collectorName>
&amp;
<collectorName id="6B605333FFCBFF9BC219F965FB17F93E" box="[1122,1185,1700,1726]" pageId="10" pageNumber="125">J. Wu</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCF906FBCBF961" box="[199,1149,1735,1762]" pageId="10" pageNumber="125" type="etymology">
<paragraph id="C62A36E5FFCBFF9BC6BCF906FBCBF961" blockId="10.[151,1437,1663,2014]" box="[199,1149,1735,1762]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCF906FEF8F961" bold="true" box="[199,334,1735,1761]" pageId="10" pageNumber="125">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="192E8847FFCBFF9BC4CDF909FD50F962" box="[694,742,1736,1762]" pageId="10" pageNumber="125">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9BC6BCF92AFEF7F815" pageId="10" pageNumber="125" type="diagnosis">
<paragraph id="C62A36E5FFCBFF9BC6BCF92AFEF7F815" blockId="10.[151,1437,1663,2014]" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCF92AFEF4F885" bold="true" box="[199,322,1771,1797]" pageId="10" pageNumber="125">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFCBFF9BC434F92DFCE5F886" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[591,851,1771,1798]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="125" phylum="Arthropoda" rank="species" species="pianma">
<emphasis id="F4E1EAF7FFCBFF9BC434F92DFD74F885" box="[591,706,1772,1797]" italics="true" pageId="10" pageNumber="125">B. pianma</emphasis>
<bibRefCitation id="A2044B14FFCBFF9BC4B2F92AFCE5F886" author="Huber, B. A." box="[713,851,1771,1798]" journalOrPublisher="Zoologica" pageId="10" pageNumber="125" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber, 2005</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFCBFF9BC5F5F92AFBF6F885" author="Huber, B. A." box="[910,1088,1771,1798]" journalOrPublisher="Zoologica" pageId="10" pageNumber="125" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber 2005: 14</bibRefCitation>
, figs 5, 6, 56, 8892, 112130) with the elongated distal apophyses on the male chelicerae (
<figureCitation id="5EAE2A60FFCBFF9BC551F8CEFC2CF8A9" box="[810,922,1807,1833]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="10" pageNumber="125">Figs. 10B</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC5DFF8CEFC64F8AA" box="[932,978,1807,1834]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="10" pageNumber="125">15E</figureCitation>
) but can be distinguished by the presence of a subdisto-ventral membranous lamella (arrow
<quantity id="016D9B00FFCBFF9BC4BBF8F2FD5CF8CD" box="[704,746,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="125" unit="in" value="3.0">3 in</quantity>
<figureCitation id="5EAE2A60FFCBFF9BC494F8F2FCFEF8CD" box="[751,840,1843,1869]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="10" pageNumber="125">Figs 9C</figureCitation>
,
<figureCitation id="5EAE2A60FFCBFF9BC528F8F2FCCFF8CD" box="[851,889,1843,1869]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="10" pageNumber="125">13I</figureCitation>
) on the procursus, a pair of angular apophyses on the bulbal apophyses (arrow in
<figureCitation id="5EAE2A60FFCBFF9BC78AF896FDEDF8F1" box="[497,603,1879,1905]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="10" pageNumber="125">Figs 10A</figureCitation>
), the absence of a ventral hump on the palpal femur and a pair of apophyses on the clypeus.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCBFF9EC6BCF85EFB67FD16" lastPageId="15" lastPageNumber="130" pageId="10" pageNumber="125" type="description">
<paragraph id="C62A36E5FFCBFF9EC6BCF85EFAEDFDD1" blockId="10.[151,1437,1663,2014]" lastBlockId="15.[151,1437,150,698]" lastPageId="15" lastPageNumber="130" pageId="10" pageNumber="125">
<emphasis id="F4E1EAF7FFCBFF9BC6BCF85EFE29F839" bold="true" box="[199,415,1951,1977]" pageId="10" pageNumber="125">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFCBFF9BC7C9F85EFDAFF839" bold="true" box="[434,537,1951,1977]" pageId="10" pageNumber="125">
<typeStatus id="192E8847FFCBFF9BC7C9F85EFDAFF839" box="[434,537,1951,1977]" pageId="10" pageNumber="125" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41910): Total length 1.60 (1.66 with clypeus), carapace 0.66 long, 0.66 wide, opisthosoma 1.00 long, 0.53 wide. Leg I: 11.39 (3.00 + 0.17 + 2.97 + 4.09 + 1.16), leg II: 7.65 (2.03 + 0.15 + 1.91 + 2.72 + 0.84), leg III: 4.85 (1.41 + 0.09 + 1.13 + 1.59 + 0.63), leg IV: 8.04 (2.28 + 0.16 + 1.94 + 2.75 + 0.91); tibia I L/d: 50. Distance PME-PME 0.14, diameter PME 0.06, distance PME-ALE 0.09, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFCEFF9EC6A8FF1DFEEBFF76" box="[211,349,220,246]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="15" pageNumber="130">Figs 10CE</figureCitation>
. Carapace pale. Ocular area not elevated. Clypeus unmodified. Chelicerae (
<figureCitation id="5EAE2A60FFCEFF9EC2D4FF1DFA97FF76" box="[1199,1313,220,246]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="15" pageNumber="130">Figs. 10B</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC357FF1DFAECFF77" box="[1324,1370,220,247]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="15" pageNumber="130">15E</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.39). Sternum pale, approximately as wide as long (0.41). Thoracic furrow absent. Legs pale. Opisthosoma pale, without spots. Palps as in
<figureCitation id="5EAE2A60FFCEFF9EC5B6FE85FBFAFEDE" box="[973,1100,324,350]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9AB</figureCitation>
; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur with retrolatero-dorsal apophysis (arrow in
<figureCitation id="5EAE2A60FFCEFF9EC2F0FEA6FB6CFE01" box="[1163,1242,359,385]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Fig 9B</figureCitation>
); procursus (
<figureCitation id="5EAE2A60FFCEFF9EC311FEA6FF6BFE23" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9AD</figureCitation>
) curved (arrow in
<figureCitation id="5EAE2A60FFCEFF9EC7CFFE48FDB8FE23" box="[436,526,393,419]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Fig. 9A</figureCitation>
), complex distally, with droplet-shaped subdisto-dorsal sclerite (arrow
<quantity id="016D9B00FFCEFF9EC34CFE48FAD2FE23" box="[1335,1380,393,419]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="130" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFCEFF9EC317FE48FF0CFE46" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC6BDFE6DFF58FE46" box="[198,238,428,454]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13I</figureCitation>
), small subdisto-prolateral sclerite (arrow
<quantity id="016D9B00FFCEFF9EC4B0FE6DFD40FE46" box="[715,758,428,454]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="130" unit="in" value="2.0">2 in</quantity>
<figureCitation id="5EAE2A60FFCEFF9EC486FE6DFCE1FE46" box="[765,855,428,454]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC51FFE6DFC3AFE46" box="[868,908,428,454]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13I</figureCitation>
), semicircular subdisto-prolateral membranous lamella (arrow
<quantity id="016D9B00FFCEFF9EC738FE0EFED9FE69" box="[323,367,463,489]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="15" pageNumber="130" unit="in" value="3.0">3 in</quantity>
<figureCitation id="5EAE2A60FFCEFF9EC70DFE0EFE67FE69" box="[374,465,463,489]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC7A6FE0EFDB3FE69" box="[477,517,463,489]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13I</figureCitation>
), subdisto-ventral membranous lamella (arrow in
<figureCitation id="5EAE2A60FFCEFF9EC243FE0EFB22FE69" box="[1080,1172,463,489]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9D</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC2DAFE0EFB7EFE69" box="[1185,1224,463,489]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13J</figureCitation>
) and with retrolateral membranous flap (arrow f in
<figureCitation id="5EAE2A60FFCEFF9EC46BFE30FDDDFD8B" box="[528,619,497,523]" captionStart="FIGURE 9" captionStartId="13.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@13.[226,1361,181,1851]" captionTargetPageId="13" captionText="FIGURE 9. Belisana putao sp. nov., holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="15" pageNumber="130">Figs 9D</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC40CFE30FD17FD8B" box="[631,673,497,523]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13J</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFCEFF9EC48DFE30FCEAFD8B" box="[758,860,497,523]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="15" pageNumber="130">Fig. 10A</figureCitation>
) with hooked apophysis (pair of angular apophyses subdistally, arrow in
<figureCitation id="5EAE2A60FFCEFF9EC7FBFDD5FE5EFDAE" box="[384,488,532,558]" captionStart="FIGURE 10" captionStartId="14.[151,250,1910,1934]" captionTargetBox="[214,1374,181,1886]" captionTargetId="figure-19@14.[214,1374,181,1886]" captionTargetPageId="14" captionText="FIGURE 10. Belisana putao sp. nov., holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (AB), 0.20 (CE)." figureDoi="http://doi.org/10.5281/zenodo.4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="15" pageNumber="130">Figs 10A</figureCitation>
) and simple embolus. Retrolateral trichobothria of tibia Iat 6% proximally; legs with short vertical setae on metatarsi, without spines and curved setae; tarsus Iwith 15 distinct pseudopodomeres.
</paragraph>
<caption id="92EA666DFFCAFF9AC6ECF892FDC0F87B" ID-DOI="http://doi.org/10.5281/zenodo.4719944" ID-Zenodo-Dep="4719944" httpUri="https://zenodo.org/record/4719944/files/figure.png" pageId="11" pageNumber="126" startId="11.[151,250,1875,1899]" targetBox="[226,1361,181,1851]" targetPageId="11">
<paragraph id="C62A36E5FFCAFF9AC6ECF892FDC0F87B" blockId="11.[151,1436,1875,2043]" pageId="11" pageNumber="126">
<emphasis id="F4E1EAF7FFCAFF9AC6ECF892FEA6F8EB" bold="true" box="[151,272,1875,1899]" pageId="11" pageNumber="126">FIGURE 7</emphasis>
.
<taxonomicName id="01954D66FFCAFF9AC764F892FE73F8EB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[287,453,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="126" phylum="Arthropoda" rank="species" species="kachin" status="sp. nov.">
<emphasis id="F4E1EAF7FFCAFF9AC764F892FE73F8EB" box="[287,453,1875,1899]" italics="true" pageId="11" pageNumber="126">Belisana kachin</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFCAFF9AC7B6F892FD97F8EB" bold="true" box="[461,545,1875,1899]" pageId="11" pageNumber="126">
<taxonomicNameLabel id="EFD2578CFFCAFF9AC7B6F892FD97F8EB" box="[461,545,1875,1899]" pageId="11" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. AB. Palp (A. Prolateral view; B. Retrolateral view, arrow points at ventral apophysis on palpal trochanter); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at curved subdistodorsal sclerite, arrow 2 points at large disto-prolateral sclerite, arrow 3 points at inverted L-shaped subdisto-prolateral sclerite; D. Retrolateral view, arrow points at disto-retrolateral membranous process). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD).
</paragraph>
</caption>
<caption id="92EA666DFFCDFF9DC6ECF892FEA6F87B" ID-DOI="http://doi.org/10.5281/zenodo.4719947" ID-Zenodo-Dep="4719947" httpUri="https://zenodo.org/record/4719947/files/figure.png" pageId="12" pageNumber="127" startId="12.[151,250,1875,1899]" targetBox="[226,1361,181,1851]" targetPageId="12">
<paragraph id="C62A36E5FFCDFF9DC6ECF892FEA6F87B" blockId="12.[151,1437,1875,2043]" pageId="12" pageNumber="127">
<emphasis id="F4E1EAF7FFCDFF9DC6ECF892FEA3F8EB" bold="true" box="[151,277,1875,1899]" pageId="12" pageNumber="127">FIGURE 8.</emphasis>
<taxonomicName id="01954D66FFCDFF9DC766F892FE74F8EB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[285,450,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="127" phylum="Arthropoda" rank="species" species="kachin" status="sp. nov.">
<emphasis id="F4E1EAF7FFCDFF9DC766F892FE74F8EB" box="[285,450,1875,1899]" italics="true" pageId="12" pageNumber="127">Belisana kachin</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFCDFF9DC7B1F892FDABF8EB" bold="true" box="[458,541,1875,1899]" pageId="12" pageNumber="127">
<taxonomicNameLabel id="EFD2578CFFCDFF9DC7B1F892FDABF8EB" box="[458,541,1875,1899]" pageId="12" pageNumber="127" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view, arrow points at extending part of anterior arch; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH).
</paragraph>
</caption>
<caption id="92EA666DFFCCFF9CC6ECF892FC50F87B" ID-DOI="http://doi.org/10.5281/zenodo.4719949" ID-Zenodo-Dep="4719949" httpUri="https://zenodo.org/record/4719949/files/figure.png" pageId="13" pageNumber="128" startId="13.[151,250,1875,1899]" targetBox="[226,1361,181,1851]" targetPageId="13">
<paragraph id="C62A36E5FFCCFF9CC6ECF892FC50F87B" blockId="13.[151,1436,1875,2043]" pageId="13" pageNumber="128">
<emphasis id="F4E1EAF7FFCCFF9CC6ECF892FEB9F8EB" bold="true" box="[151,271,1875,1899]" pageId="13" pageNumber="128">FIGURE 9</emphasis>
.
<taxonomicName id="01954D66FFCCFF9CC766F892FE0EF8EB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[285,440,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="128" phylum="Arthropoda" rank="species" species="putao" status="sp. nov.">
<emphasis id="F4E1EAF7FFCCFF9CC766F892FE0EF8EB" box="[285,440,1875,1899]" italics="true" pageId="13" pageNumber="128">Belisana putao</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFCCFF9CC7BBF892FDA5F8EB" bold="true" box="[448,531,1875,1899]" pageId="13" pageNumber="128">
<taxonomicNameLabel id="EFD2578CFFCCFF9CC7BBF892FDA5F8EB" box="[448,531,1875,1899]" pageId="13" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. AB. Palp (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at retrolatero-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at drop-shaped subdisto-dorsal sclerite, arrow 2 points at small subdisto-prolateral sclerite, arrow 3 points at semicircular subdisto-prolateral membranous lamella; D. Retrolateral view, arrow points at subdisto-ventral membranous lamella). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD).
</paragraph>
</caption>
<caption id="92EA666DFFCFFF9FC6ECF8B7FAD9F856" ID-DOI="http://doi.org/10.5281/zenodo.4719951" ID-Zenodo-Dep="4719951" httpUri="https://zenodo.org/record/4719951/files/figure.png" pageId="14" pageNumber="129" startId="14.[151,250,1910,1934]" targetBox="[214,1374,181,1886]" targetPageId="14">
<paragraph id="C62A36E5FFCFFF9FC6ECF8B7FAD9F856" blockId="14.[151,1436,1910,2006]" pageId="14" pageNumber="129">
<emphasis id="F4E1EAF7FFCFFF9FC6ECF8B7FE97F80E" bold="true" box="[151,289,1910,1934]" pageId="14" pageNumber="129">FIGURE 10.</emphasis>
<taxonomicName id="01954D66FFCFFF9FC752F8B7FE75F80E" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[297,451,1910,1934]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="129" phylum="Arthropoda" rank="species" species="putao" status="sp. nov.">
<emphasis id="F4E1EAF7FFCFFF9FC752F8B7FE75F80E" box="[297,451,1910,1934]" italics="true" pageId="14" pageNumber="129">Belisana putao</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFCFFF9FC7B0F8B7FDABF80E" bold="true" box="[459,541,1910,1934]" pageId="14" pageNumber="129">
<taxonomicNameLabel id="EFD2578CFFCFFF9FC7B0F8B7FDABF80E" box="[459,541,1910,1934]" pageId="14" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A. Bulb, prolateral view, arrow points at pair of angular apophyses on bulbal apophysis; B. Chelicerae, frontal view; CE. Habitus (C. Dorsal view; D. Ventral view; E. Lateral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.05 (AB), 0.20 (CE).
</paragraph>
</caption>
<paragraph id="C62A36E5FFCEFF9EC6BCFD98FE13FDF3" blockId="15.[151,1437,150,698]" box="[199,421,601,627]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCFD98FE9EFDF3" bold="true" box="[199,296,601,627]" pageId="15" pageNumber="130">Female:</emphasis>
Unknown.
</paragraph>
<paragraph id="C62A36E5FFCEFF9EC6BCFDBDFB67FD16" blockId="15.[151,1437,150,698]" box="[199,1233,636,663]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCFDBDFE33FD16" bold="true" box="[199,389,636,662]" pageId="15" pageNumber="130">Natural history.</emphasis>
The species was found on domed webs attached to the underside of leaves.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCEFF9EC6BCFD5FFCC0FD3A" box="[199,886,670,698]" pageId="15" pageNumber="130" type="distribution">
<paragraph id="C62A36E5FFCEFF9EC6BCFD5FFCC0FD3A" blockId="15.[151,1437,150,698]" box="[199,886,670,698]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCFD5FFED7FD38" bold="true" box="[199,353,670,696]" pageId="15" pageNumber="130">Distribution.</emphasis>
<collectingCountry id="BE827675FFCEFF9EC71CFD5EFE62FD39" box="[359,468,671,697]" name="Myanmar" pageId="15" pageNumber="130">Myanmar</collectingCountry>
(
<collectingRegion id="0451F807FFCEFF9EC798FD5EFD8EFD39" box="[483,568,671,697]" country="Myanmar" name="Kachin" pageId="15" pageNumber="130">Kachin</collectingRegion>
,
<typeStatus id="192E8847FFCEFF9EC438FD5EFDC5FD39" box="[579,627,671,697]" pageId="15" pageNumber="130">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFCEFF9EC56DFD5EFCDFFD3A" box="[790,873,671,698]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="15" pageNumber="130">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="0C4D9CA2420559D5C481F3460B59FEE0" ID-CLB-Dataset="7182" ID-DOI="10.11646/zootaxa.4963.1.5" ID-GBIF-Dataset="ee9bfd7b-3b05-4403-9af2-e54beaae5e69" ID-ISSN="1175-5326" ID-Zenodo-Dep="4719924" ID-ZooBank="6B4B8F71-7B08-4027-A76B-7EE8C987C9BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618562743172" checkinUser="plazi" docAuthor="Zhu, Wenhui &amp; Li, Shuqiang" docDate="2021" docId="4E3C87F3FFCEFF84C6ECFCC7FC34F871" docLanguage="en" docName="zootaxa.4963.1.5.pdf" docOrigin="Zootaxa 4963 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana tarang Zhu &amp; Li 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="136" masterDocId="B205FF8BFFC1FF91C67BFFC1FFB6FF80" masterDocTitle="Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia" masterLastPageNumber="137" masterPageNumber="115" pageNumber="130" updateTime="1698937141523" updateUser="ExternalLinkService">
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<mods:title id="ED63F497FEF1696FF81E4A7269DDCD86">Six new species of the spider genus Belisana (Araneae: Pholcidae) from Southeast Asia</mods:title>
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<mods:namePart id="59FAEBC7E198A4A519AD88816B3E67F7">Zhu, Wenhui</mods:namePart>
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<mods:affiliation id="758CD24B85B44AA6B664BA1B9D7861F9">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
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<mods:namePart id="EE1619BC6743FD65A484F1DFA9BCED17">Li, Shuqiang</mods:namePart>
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<mods:affiliation id="27A313D30D8DA717E796D313FB4BABB1">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
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<treatment id="4E3C87F3FFCEFF84C6ECFCC7FC34F871" ID-DOI="http://doi.org/10.5281/zenodo.4697008" ID-GBIF-Taxon="181600549" ID-Zenodo-Dep="4697008" LSID="urn:lsid:plazi:treatment:4E3C87F3FFCEFF84C6ECFCC7FC34F871" httpUri="http://treatment.plazi.org/id/4E3C87F3FFCEFF84C6ECFCC7FC34F871" lastPageId="21" lastPageNumber="136" pageId="15" pageNumber="130">
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<emphasis id="F4E1EAF7FFCEFF9EC6ECFCC7FE0BFCA1" bold="true" box="[151,445,774,801]" pageId="15" pageNumber="130">
<taxonomicName id="01954D66FFCEFF9EC6ECFCC7FEEEFCA1" ID-CoL="98YXT" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[151,344,774,801]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="tarang" status="sp. nov.">
<emphasis id="F4E1EAF7FFCEFF9EC6ECFCC7FEEEFCA1" bold="true" box="[151,344,774,801]" italics="true" pageId="15" pageNumber="130">Belisana tarang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFD2578CFFCEFF9EC71BFCC6FE0BFCA1" box="[352,445,775,801]" pageId="15" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="C62A36E5FFCEFF9EC6ECFCE8FE40FCC4" blockId="15.[151,502,774,836]" box="[151,502,809,836]" pageId="15" pageNumber="130">
<figureCitation id="5EAE2A60FFCEFF9EC6ECFCE8FEA0FCC3" box="[151,278,809,835]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="16.[151,250,1806,1830]" captionStartId-1="17.[151,250,1875,1899]" captionTargetBox-0="[249,1339,180,1783]" captionTargetBox-1="[226,1361,181,1851]" captionTargetId-0="figure-19@16.[249,1339,180,1783]" captionTargetId-1="figure-17@17.[226,1361,181,1851]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." captionText-1="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.4719953" figureDoi-1="http://doi.org/10.5281/zenodo.4719955" httpUri-0="https://zenodo.org/record/4719953/files/figure.png" httpUri-1="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs 1112</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC75AFCE8FEC5FCC3" box="[289,371,809,835]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13KL</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC704FCE8FE76FCC3" box="[383,448,809,835]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="15" pageNumber="130">14IJ</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC7B0FCE8FE40FCC4" box="[459,502,809,836]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="15" pageNumber="130">15F</figureCitation>
</paragraph>
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<paragraph id="C62A36E5FFCEFF9EC6ECFCAEFB8CFC4E" blockId="15.[151,1437,878,2015]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6ECFCAEFEF2FC08" bold="true" box="[151,324,878,905]" pageId="15" pageNumber="130">Type material.</emphasis>
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<emphasis id="F4E1EAF7FFCEFF9EC731FCAEFE76FC09" bold="true" box="[330,448,879,905]" pageId="15" pageNumber="130">
<typeStatus id="192E8847FFCEFF9EC731FCAEFE0DFC09" box="[330,443,879,905]" pageId="15" pageNumber="130" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male (
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Ar41916), near entrance of
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(
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,
<geoCoordinate id="A3A15022FFCEFF9EC2A5FCAFFA2EFC09" box="[1246,1432,878,905]" degrees="100" direction="east" minutes="24" orientation="longitude" pageId="15" pageNumber="130" precision="1" seconds="58.62" value="100.41628">100°2458.62”E</geoCoordinate>
,
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elevation
<quantity id="016D9B00FFCEFF9EC77CFC50FEE5FC2C" box="[263,339,913,940]" metricMagnitude="2" metricUnit="m" metricValue="8.77" pageId="15" pageNumber="130" unit="m" value="877.0">877 m</quantity>
</elevation>
),
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,
<collectingRegion id="0451F807FFCEFF9EC782FC53FD2CFC2B" box="[505,666,913,940]" country="Indonesia" name="Sumatera Barat" pageId="15" pageNumber="130">West Sumatra</collectingRegion>
,
<collectingRegion id="0451F807FFCEFF9EC4DCFC50FCB1FC2B" box="[679,775,913,939]" country="Indonesia" name="Sumatera Utara" pageId="15" pageNumber="130">Sumatra</collectingRegion>
,
<collectingCountry id="BE827675FFCEFF9EC56EFC50FC33FC2B" box="[789,901,913,939]" name="Indonesia" pageId="15" pageNumber="130">Indonesia</collectingCountry>
,
<date id="B22B1025FFCEFF9EC5E9FC50FB99FC2B" box="[914,1071,913,940]" pageId="15" pageNumber="130" value="2014-05-13">
<collectingDate id="A26FE9CDFFCEFF9EC5E9FC50FB99FC2B" box="[914,1071,913,940]" pageId="15" pageNumber="130" value="2014-05-13">13 May 2014</collectingDate>
</date>
,
<collectorName id="6B605333FFCEFF9EC247FC53FB3FFC2C" box="[1084,1161,914,940]" pageId="15" pageNumber="130">Z. Yao</collectorName>
leg.
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<emphasis id="F4E1EAF7FFCEFF9EC2BFFC50FAF3FC2B" bold="true" box="[1220,1349,913,939]" pageId="15" pageNumber="130">
<typeStatus id="192E8847FFCEFF9EC2BFFC50FAF6FC2B" box="[1220,1344,913,939]" pageId="15" pageNumber="130" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="D093FD6CFFCEFF9EC334FC50FA2AFC2B" box="[1359,1436,913,939]" pageId="15" pageNumber="130" type="male">1 male</specimenCount>
(
<collectionCode id="A084AE20FFCEFF9EC6E4FC75FF44FC4E" box="[159,242,948,974]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="15" pageNumber="130">IZCAS</collectionCode>
Ar41917) and
</materialsCitation>
<materialsCitation id="76FD3CB8FFCEFF9EC7E1FC75FB80FC4E" ID-GBIF-Occurrence="3091104309" box="[410,1078,948,975]" collectingDate="2014-05-13" collectionCode="IZCAS" collectorName="Z. Yao" country="Indonesia" elevation="877" latitude="0.21088333" location="Tarang Cave" longLatPrecision="1" longitude="100.41628" municipality="Bukittinggi" pageId="15" pageNumber="130" specimenCount="2" specimenCount-female="2" stateProvince="Sumatera Barat" typeStatus="paratype">
<specimenCount id="D093FD6CFFCEFF9EC7E1FC75FDB0FC4E" box="[410,518,948,974]" pageId="15" pageNumber="130" type="female">2 females</specimenCount>
(
<collectionCode id="A084AE20FFCEFF9EC46EFC75FDDEFC4E" box="[533,616,948,974]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="15" pageNumber="130">IZCAS</collectionCode>
Ar4191841919), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCEFF9EC6BCFC16FBCAFC71" box="[199,1148,983,1009]" pageId="15" pageNumber="130" type="etymology">
<paragraph id="C62A36E5FFCEFF9EC6BCFC16FBCAFC71" blockId="15.[151,1437,878,2015]" box="[199,1148,983,1009]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCFC16FEF8FC71" bold="true" box="[199,334,983,1009]" pageId="15" pageNumber="130">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="192E8847FFCEFF9EC4CEFC16FD53FC71" box="[693,741,983,1009]" pageId="15" pageNumber="130">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCEFF9EC6BCFC38FF4AFAA9" pageId="15" pageNumber="130" type="diagnosis">
<paragraph id="C62A36E5FFCEFF9EC6BCFC38FF4AFAA9" blockId="15.[151,1437,878,2015]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCFC38FEF4FB93" bold="true" box="[199,322,1017,1043]" pageId="15" pageNumber="130">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="01954D66FFCEFF9EC435FC3BFCDFFB94" authority="Huber, 2005" authorityName=", Huber" authorityYear="2005" box="[590,873,1017,1044]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="sandakan">
<emphasis id="F4E1EAF7FFCEFF9EC435FC3BFD61FB93" box="[590,727,1017,1043]" italics="true" pageId="15" pageNumber="130">B. sandakan</emphasis>
<bibRefCitation id="A2044B14FFCEFF9EC4A5FC38FCDFFB94" author="Huber, B. A." box="[734,873,1017,1044]" journalOrPublisher="Zoologica" pageId="15" pageNumber="130" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber, 2005</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="A2044B14FFCEFF9EC5D8FC38FBE2FB94" author="Huber, B. A." box="[931,1108,1017,1044]" journalOrPublisher="Zoologica" pageId="15" pageNumber="130" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber 2005: 59</bibRefCitation>
, figs 360375) with the same shape of the bulbal apophyses (
<figureCitation id="5EAE2A60FFCEFF9EC479FBDDFDDAFBB6" box="[514,620,1052,1078]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Fig. 12D</figureCitation>
), but males can be distinguished by the presence of a nearly triangular subdisto-prolateral sclerite (arrow
<quantity id="016D9B00FFCEFF9EC462FBFEFDF2FBD9" box="[537,580,1087,1113]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="15" pageNumber="130" unit="in" value="1.0">1 in</quantity>
<figureCitation id="5EAE2A60FFCEFF9EC430FBFEFD04FBD9" box="[587,690,1087,1113]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Figs 11C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC4C5FBFEFD59FBD9" box="[702,751,1087,1113]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13K</figureCitation>
) and triangular retrolateral membranous flap (arrow f in
<figureCitation id="5EAE2A60FFCEFF9EC317FBFEFF7FFBFB" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Figs 11D</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC6AEFBA0FEB5FBFB" box="[213,259,1121,1147]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13L</figureCitation>
) on the procursus (
<figureCitation id="5EAE2A60FFCEFF9EC7A0FBA0FDDEFBFB" box="[475,616,1121,1147]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Figs 11AD</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC40FFBA0FD71FBFB" box="[628,711,1121,1147]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13KL</figureCitation>
), and by having the cheliceral distal apophyses directed laterally (arrow da in
<figureCitation id="5EAE2A60FFCEFF9EC75DFB45FE22FB1E" box="[294,404,1156,1182]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs. 12C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC7DBFB45FE78FB1F" box="[416,462,1156,1183]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="15" pageNumber="130">15F</figureCitation>
; distal apophyses directed downward in
<taxonomicName id="01954D66FFCEFF9EC5EAFB44FBADFB1E" authorityName=", Huber" authorityYear="2005" box="[913,1051,1156,1182]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="sandakan">
<emphasis id="F4E1EAF7FFCEFF9EC5EAFB44FBADFB1E" box="[913,1051,1156,1182]" italics="true" pageId="15" pageNumber="130">B. sandakan</emphasis>
</taxonomicName>
). Females can be distinguished by having the frontal “epigynal” pockets laterally on a sclerotized plate (arrow ep in
<figureCitation id="5EAE2A60FFCEFF9EC247FB66FB7EFB41" box="[1084,1224,1191,1217]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs 12AB</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC2AEFB66FAA2FB41" box="[1237,1300,1191,1217]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="15" pageNumber="130">14IJ</figureCitation>
) and a long vulval anterior arch, 2/3 of the vulva length (
<figureCitation id="5EAE2A60FFCEFF9EC4F0FB08FD43FB63" box="[651,757,1225,1251]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs 12B</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC57BFB08FC9FFB63" box="[768,809,1225,1251]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="15" pageNumber="130">14J</figureCitation>
) (vs. short vulval anterior arch, 1/2 of the vulva length). It can also distinguished by a longer total body length (total body length &lt;1.4, “tiny” size in
<taxonomicName id="01954D66FFCEFF9EC2C7FB2CFF58FAA9" authority=", Huber 2005: 3" authorityName=", Huber" authorityPageNumber="3" authorityYear="2005" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="130" phylum="Arthropoda" rank="species" species="sandakan">
<emphasis id="F4E1EAF7FFCEFF9EC2C7FB2CFAF1FA86" box="[1212,1351,1260,1286]" italics="true" pageId="15" pageNumber="130">B. sandakan</emphasis>
,
<bibRefCitation id="A2044B14FFCEFF9EC32DFB2DFF58FAA9" author="Huber, B. A." journalOrPublisher="Zoologica" pageId="15" pageNumber="130" pagination="1 - 126" part="155" refId="ref8453" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." title="High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae)" type="journal article" year="2005">Huber 2005: 3</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFCEFF84C6BCFAF0FABFF84C" lastPageId="21" lastPageNumber="136" pageId="15" pageNumber="130" type="description">
<paragraph id="C62A36E5FFCEFF9EC6BCFAF0FD2DF88F" blockId="15.[151,1437,878,2015]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCFAF0FE29FACB" bold="true" box="[199,415,1329,1355]" pageId="15" pageNumber="130">Description. Male</emphasis>
(
<emphasis id="F4E1EAF7FFCEFF9EC7C9FAF0FDAFFACB" bold="true" box="[434,537,1329,1355]" pageId="15" pageNumber="130">
<typeStatus id="192E8847FFCEFF9EC7C9FAF0FDAFFACB" box="[434,537,1329,1355]" pageId="15" pageNumber="130" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41916): Total length 1.46 (1.54 with clypeus), carapace 0.63 long, 0.63 wide, opisthosoma 0.91 long, 0.69 wide. Leg I missing (see variation), leg II: 7.45 (2.03 + 0.25 + 1.88 + 2.63 + 0.66), leg III: 3.23 (0.88 + 0.13 + 0.88 + 1.09 + 0.25), leg IV: 5.07 (1.47 + 0.19 + 1.19 + 1.63 + 0.59). Distance PME-PME 0.06, diameter PME 0.08, distance PME-ALE 0.08, AME absent. Habitus as in
<figureCitation id="5EAE2A60FFCEFF9EC2DFFA58FA9EFA33" box="[1188,1320,1433,1459]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs 12EF</figureCitation>
. Carapace pale, with yellow median marks. Ocular area not elevated. Clypeus unmodified. Chelicerae (
<figureCitation id="5EAE2A60FFCEFF9EC2CBFA7DFA97FA56" box="[1200,1313,1468,1494]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs. 12C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC355FA7DFAECFA57" box="[1326,1370,1468,1495]" captionStart="FIGURE 15" captionStartId="20.[151,250,1726,1750]" captionTargetBox="[151,1435,199,1702]" captionTargetId="figure-19@20.[151,1435,199,1702]" captionTargetPageId="20" captionText="FIGURE 15. Chelicerae in lateral views.A. B. bachma sp. nov.; B. B. cucphuong sp. nov.; C. B. jaegeri sp. nov.; D. B. kachin sp. nov.; E. B. putao sp. nov.; F. B. tarang sp. nov. ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="15" pageNumber="130">15F</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with sclerotized and pointed tip (distance between tips of distal apophyses: 0.23). Sternum pale, approximately as wide as long (0.38). Thoracic furrow absent. Legs pale, without darker rings. Opisthosoma pale, without spots. Palps as in
<figureCitation id="5EAE2A60FFCEFF9EC2DEF9E5FA86F9BE" box="[1189,1328,1572,1598]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Figs 11AB</figureCitation>
; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur with small proximo-dorsal apophysis (arrow in
<figureCitation id="5EAE2A60FFCEFF9EC6ECF9A8FF4DF903" box="[151,251,1641,1667]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Fig. 11B</figureCitation>
); procursus (
<figureCitation id="5EAE2A60FFCEFF9EC7F1F9A8FDA3F903" box="[394,533,1641,1667]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Figs 11AD</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC45BF9A8FDC7F903" box="[544,625,1641,1667]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13KL</figureCitation>
) curved (arrow in
<figureCitation id="5EAE2A60FFCEFF9EC545F9A8FC14F903" box="[830,930,1641,1667]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Fig. 11A</figureCitation>
), complex distally, with large, nearly triangular subdisto-prolateral sclerite (arrow
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<figureCitation id="5EAE2A60FFCEFF9EC412F94DFD79F926" box="[617,719,1676,1702]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Figs 11C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC4A1F94DFCBBF926" box="[730,781,1676,1702]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13K</figureCitation>
), small subdisto-dorsal sclerite (arrow
<quantity id="016D9B00FFCEFF9EC2C0F94DFB53F926" box="[1211,1253,1676,1702]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="130" unit="in" value="2.0">2 in</quantity>
<figureCitation id="5EAE2A60FFCEFF9EC290F94DFAE7F926" box="[1259,1361,1676,1702]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Figs 11C</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC327F94DFA38F926" box="[1372,1422,1676,1702]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13K</figureCitation>
), large distal membranous process (arrow in
<figureCitation id="5EAE2A60FFCEFF9EC405F96EFD51F949" box="[638,743,1711,1737]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Figs 11D</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC488F96EFC97F949" box="[755,801,1711,1737]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13L</figureCitation>
) and triangular retrolateral membranous flap (arrow f in
<figureCitation id="5EAE2A60FFCEFF9EC6ECF910FEB7F96B" box="[151,257,1745,1771]" captionStart="FIGURE 11" captionStartId="16.[151,250,1806,1830]" captionTargetBox="[249,1339,180,1783]" captionTargetId="figure-19@16.[249,1339,180,1783]" captionTargetPageId="16" captionText="FIGURE 11. Belisana tarang sp. nov., holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD)." figureDoi="http://doi.org/10.5281/zenodo.4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="15" pageNumber="130">Figs 11D</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC775F910FE88F96B" box="[270,318,1745,1771]" captionStart="FIGURE 13" captionStartId="18.[151,250,1875,1899]" captionTargetBox="[239,1334,203,1844]" captionTargetId="figure-19@18.[226,1361,181,1851]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively).AB. B. bachma sp. nov.; CD. B. cucphuong sp. nov.; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov.; IJ. B. putao sp. nov.; KL. B. tarang sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="15" pageNumber="130">13L</figureCitation>
); bulb (
<figureCitation id="5EAE2A60FFCEFF9EC7EDF910FE4BF96B" box="[406,509,1745,1771]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Fig. 12D</figureCitation>
) with short, hooked apophysis and simple embolus. Legs with short vertical setae on metatarsi, without spines and curved setae.
</paragraph>
<paragraph id="C62A36E5FFCEFF9EC6BCF8D6FD4CF85E" blockId="15.[151,1437,878,2015]" pageId="15" pageNumber="130">
<emphasis id="F4E1EAF7FFCEFF9EC6BCF8D6FEA8F8B1" bold="true" box="[199,286,1815,1841]" pageId="15" pageNumber="130">Female</emphasis>
(IZCAS Ar41918): Similar to male, habitus as in
<figureCitation id="5EAE2A60FFCEFF9EC523F8D6FC51F8B1" box="[856,999,1815,1841]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs 12GH</figureCitation>
. Total length 1.39 (1.47 with clypeus), carapace 0.47 long, 0.47 wide, opisthosoma 1.00 long, 0.94 wide. Tibia I: 1.75; tibia I L/d: 29. Distance PME-PME 0.07, diameter PME 0.06, distance PME-ALE 0.07, AME absent. Sternum slightly wider than long (0.38/0.34). External female genitalia (
<figureCitation id="5EAE2A60FFCEFF9EC7BBF8BEFD9AF819" box="[448,556,1919,1945]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs 12A</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC44CF8BEFDE8F819" box="[567,606,1919,1945]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="15" pageNumber="130">14I</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets (arrow ep in
<figureCitation id="5EAE2A60FFCEFF9EC317F8BEFF5CF83B" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs 12AB</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC68DF860FE8EF83B" box="[246,312,1953,1979]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="15" pageNumber="130">14IJ</figureCitation>
), 0.34 apart. Vulva (
<figureCitation id="5EAE2A60FFCEFF9EC460F860FD33F83B" box="[539,645,1953,1979]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs 12B</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC4EBF860FD0EF83B" box="[656,696,1953,1979]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="15" pageNumber="130">14J</figureCitation>
) with W-shaped anterior arch and pair of long, narrow pore plates (arrow pp in
<figureCitation id="5EAE2A60FFCEFF9EC752F805FE27F85E" box="[297,401,1988,2014]" captionStart="FIGURE 12" captionStartId="17.[151,250,1875,1899]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@17.[226,1361,181,1851]" captionTargetPageId="17" captionText="FIGURE 12. Belisana tarang sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="15" pageNumber="130">Figs 12B</figureCitation>
,
<figureCitation id="5EAE2A60FFCEFF9EC7E6F805FE70F85E" box="[413,454,1988,2014]" captionStart="FIGURE 14" captionStartId="19.[151,250,1910,1934]" captionTargetBox="[231,1358,188,1876]" captionTargetId="figure-17@19.[214,1374,181,1886]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. bachma sp. nov., arrow points at strongly protruding median ridge of anterior arch; CD. B. cucphuong sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. jaegeri sp. nov.; GH. B. kachin sp. nov., arrow points at extending part of anterior arch; IJ. B. tarang sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="15" pageNumber="130">14J</figureCitation>
), without serrated sclerites.
</paragraph>
<caption id="92EA666DFFD1FF81C6ECF8CFFEA4F837" ID-DOI="http://doi.org/10.5281/zenodo.4719953" ID-Zenodo-Dep="4719953" httpUri="https://zenodo.org/record/4719953/files/figure.png" pageId="16" pageNumber="131" startId="16.[151,250,1806,1830]" targetBox="[249,1339,180,1783]" targetPageId="16">
<paragraph id="C62A36E5FFD1FF81C6ECF8CFFEA4F837" blockId="16.[151,1436,1806,1975]" pageId="16" pageNumber="131">
<emphasis id="F4E1EAF7FFD1FF81C6ECF8CFFEAEF8A7" bold="true" box="[151,280,1806,1831]" pageId="16" pageNumber="131">FIGURE 11</emphasis>
.
<taxonomicName id="01954D66FFD1FF81C75FF8CEFE71F8A6" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[292,455,1807,1831]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="131" phylum="Arthropoda" rank="species" species="tarang" status="sp. nov.">
<emphasis id="F4E1EAF7FFD1FF81C75FF8CEFE71F8A6" box="[292,455,1807,1831]" italics="true" pageId="16" pageNumber="131">Belisana tarang</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD1FF81C7B6F8CEFDABF8A7" bold="true" box="[461,541,1807,1831]" pageId="16" pageNumber="131">
<taxonomicNameLabel id="EFD2578CFFD1FF81C7B6F8CEFDABF8A7" box="[461,541,1807,1831]" pageId="16" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. AB. Palpal (Bulb removed) (A. Prolateral view, arrow points at curved procursus; B. Retrolateral view, arrow points at small proximo-dorsal apophysis on palpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly triangular subdisto-prolateral sclerite, arrow 2 points at small subdisto-dorsal sclerite; D. Retrolateral view, arrow points at large distal membranous process). f = flap, pr = procursus. Scale bars: 0.10 (AB), 0.05 (CD).
</paragraph>
</caption>
<caption id="92EA666DFFD0FF80C6ECF892FB72F857" ID-DOI="http://doi.org/10.5281/zenodo.4719955" ID-Zenodo-Dep="4719955" httpUri="https://zenodo.org/record/4719955/files/figure.png" pageId="17" pageNumber="132" startId="17.[151,250,1875,1899]" targetBox="[226,1361,181,1851]" targetPageId="17">
<paragraph id="C62A36E5FFD0FF80C6ECF892FB72F857" blockId="17.[151,1436,1875,2007]" pageId="17" pageNumber="132">
<emphasis id="F4E1EAF7FFD0FF80C6ECF892FE97F8EB" bold="true" box="[151,289,1875,1899]" pageId="17" pageNumber="132">FIGURE 12.</emphasis>
<taxonomicName id="01954D66FFD0FF80C753F892FE7BF8EB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[296,461,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="132" phylum="Arthropoda" rank="species" species="tarang" status="sp. nov.">
<emphasis id="F4E1EAF7FFD0FF80C753F892FE7BF8EB" box="[296,461,1875,1899]" italics="true" pageId="17" pageNumber="132">Belisana tarang</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD0FF80C7AFF892FD90F8EB" bold="true" box="[468,550,1875,1899]" pageId="17" pageNumber="132">
<taxonomicNameLabel id="EFD2578CFFD0FF80C7AFF892FD90F8EB" box="[468,550,1875,1899]" pageId="17" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Chelicerae, frontal view; D. Bulb, prolateral view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.30 (EH).
</paragraph>
</caption>
<caption id="92EA666DFFD3FF83C6ECF892FC19F833" ID-DOI="http://doi.org/10.5281/zenodo.4719957" ID-Zenodo-Dep="4719957" httpUri="https://zenodo.org/record/4719957/files/figure.png" pageId="18" pageNumber="133" startId="18.[151,250,1875,1899]" targetBox="[239,1334,203,1844]" targetPageId="18">
<paragraph id="C62A36E5FFD3FF83C6ECF892FC19F833" blockId="18.[151,1436,1875,1971]" pageId="18" pageNumber="133">
<emphasis id="F4E1EAF7FFD3FF83C6ECF892FE96F8EB" bold="true" box="[151,288,1875,1899]" pageId="18" pageNumber="133">FIGURE 13.</emphasis>
Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB.
<taxonomicName id="01954D66FFD3FF83C791F8B9FDE3F80F" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[490,597,1911,1935]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="133" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFD3FF83C791F8B9FDE3F80F" box="[490,597,1911,1935]" italics="true" pageId="18" pageNumber="133">B. bachma</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD3FF83C420F8B6FD1DF80F" bold="true" box="[603,683,1911,1935]" pageId="18" pageNumber="133">
<taxonomicNameLabel id="EFD2578CFFD3FF83C420F8B6FD1DF80F" box="[603,683,1911,1935]" pageId="18" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; CD.
<taxonomicName id="01954D66FFD3FF83C48FF8B9FCC9F80F" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[756,895,1911,1935]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="133" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFD3FF83C48FF8B9FCC9F80F" box="[756,895,1911,1935]" italics="true" pageId="18" pageNumber="133">B. cucphuong</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD3FF83C5FEF8B6FC62F80F" bold="true" box="[901,980,1911,1935]" pageId="18" pageNumber="133">
<taxonomicNameLabel id="EFD2578CFFD3FF83C5FEF8B6FC62F80F" box="[901,980,1911,1935]" pageId="18" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; EF.
<taxonomicName id="01954D66FFD3FF83C26DF8B9FBCFF80F" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[1046,1145,1912,1935]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="133" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
<emphasis id="F4E1EAF7FFD3FF83C26DF8B9FBCFF80F" box="[1046,1145,1912,1935]" italics="true" pageId="18" pageNumber="133">B. jaegeri</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD3FF83C204F8B6FB79F80F" bold="true" box="[1151,1231,1911,1935]" pageId="18" pageNumber="133">
<taxonomicNameLabel id="EFD2578CFFD3FF83C204F8B6FB79F80F" box="[1151,1231,1911,1935]" pageId="18" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; GH.
<taxonomicName id="01954D66FFD3FF83C362F8B9FACFF80F" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[1305,1401,1911,1935]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="133" phylum="Arthropoda" rank="species" species="kachin" status="sp. nov.">
<emphasis id="F4E1EAF7FFD3FF83C362F8B9FACFF80F" box="[1305,1401,1911,1935]" italics="true" pageId="18" pageNumber="133">B. kachin</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD3FF83C305F8B6FF75F833" bold="true" pageId="18" pageNumber="133">
<taxonomicNameLabel id="EFD2578CFFD3FF83C305F8B6FF75F833" pageId="18" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; IJ.
<taxonomicName id="01954D66FFD3FF83C686F85DFEE5F833" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[253,339,1948,1971]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="133" phylum="Arthropoda" rank="species" species="putao" status="sp. nov.">
<emphasis id="F4E1EAF7FFD3FF83C686F85DFEE5F833" box="[253,339,1948,1971]" italics="true" pageId="18" pageNumber="133">B. putao</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD3FF83C722F85AFE1FF833" bold="true" box="[345,425,1947,1971]" pageId="18" pageNumber="133">
<taxonomicNameLabel id="EFD2578CFFD3FF83C722F85AFE1FF833" box="[345,425,1947,1971]" pageId="18" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; KL.
<taxonomicName id="01954D66FFD3FF83C789F85DFDE4F833" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[498,594,1948,1971]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="133" phylum="Arthropoda" rank="species" species="tarang" status="sp. nov.">
<emphasis id="F4E1EAF7FFD3FF83C789F85DFDE4F833" box="[498,594,1948,1971]" italics="true" pageId="18" pageNumber="133">B. tarang</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD3FF83C423F85AFD1FF833" bold="true" box="[600,681,1947,1971]" pageId="18" pageNumber="133">
<taxonomicNameLabel id="EFD2578CFFD3FF83C423F85AFD1FF833" box="[600,681,1947,1971]" pageId="18" pageNumber="133" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
f = flap. Scale bars: 0.05.
</paragraph>
</caption>
<caption id="92EA666DFFD2FF82C6ECF8B7FC90F87A" ID-DOI="http://doi.org/10.5281/zenodo.4719959" ID-Zenodo-Dep="4719959" httpUri="https://zenodo.org/record/4719959/files/figure.png" pageId="19" pageNumber="134" startId="19.[151,250,1910,1934]" targetBox="[231,1358,188,1876]" targetPageId="19">
<paragraph id="C62A36E5FFD2FF82C6ECF8B7FC90F87A" blockId="19.[151,1436,1910,2042]" pageId="19" pageNumber="134">
<emphasis id="F4E1EAF7FFD2FF82C6ECF8B7FE96F80E" bold="true" box="[151,288,1910,1934]" pageId="19" pageNumber="134">FIGURE 14.</emphasis>
Drawings of female genitalia in ventral and dorsal views. AB.
<taxonomicName id="01954D66FFD2FF82C5D0F8B6FBA1F80E" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[939,1047,1910,1934]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="134" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFD2FF82C5D0F8B6FBA1F80E" box="[939,1047,1910,1934]" italics="true" pageId="19" pageNumber="134">B. bachma</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD2FF82C266F8B7FBD8F80E" bold="true" box="[1053,1134,1910,1934]" pageId="19" pageNumber="134">
<taxonomicNameLabel id="EFD2578CFFD2FF82C266F8B7FBD8F80E" box="[1053,1134,1910,1934]" pageId="19" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, arrow points at strongly protruding median ridge of anterior arch; CD.
<taxonomicName id="01954D66FFD2FF82C42EF85AFD69F832" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[597,735,1946,1970]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="134" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFD2FF82C42EF85AFD69F832" box="[597,735,1946,1970]" italics="true" pageId="19" pageNumber="134">B. cucphuong</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD2FF82C49EF85BFC83F832" bold="true" box="[741,821,1946,1970]" pageId="19" pageNumber="134">
<taxonomicNameLabel id="EFD2578CFFD2FF82C49EF85BFC83F832" box="[741,821,1946,1970]" pageId="19" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, arrow points at strongly protruding median ridge of anterior arch; EF.
<taxonomicName id="01954D66FFD2FF82C77DF87EFEDDF856" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[262,363,1983,2006]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="134" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
<emphasis id="F4E1EAF7FFD2FF82C77DF87EFEDDF856" box="[262,363,1983,2006]" italics="true" pageId="19" pageNumber="134">B. jaegeri</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD2FF82C70AF87FFE74F856" bold="true" box="[369,450,1982,2006]" pageId="19" pageNumber="134">
<taxonomicNameLabel id="EFD2578CFFD2FF82C70AF87FFE74F856" box="[369,450,1982,2006]" pageId="19" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; GH.
<taxonomicName id="01954D66FFD2FF82C474F87EFDD9F856" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[527,623,1982,2006]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="134" phylum="Arthropoda" rank="species" species="kachin" status="sp. nov.">
<emphasis id="F4E1EAF7FFD2FF82C474F87EFDD9F856" box="[527,623,1982,2006]" italics="true" pageId="19" pageNumber="134">B. kachin</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD2FF82C40DF87FFD71F856" bold="true" box="[630,711,1982,2006]" pageId="19" pageNumber="134">
<taxonomicNameLabel id="EFD2578CFFD2FF82C40DF87FFD71F856" box="[630,711,1982,2006]" pageId="19" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, arrow points at extending part of anterior arch; IJ.
<taxonomicName id="01954D66FFD2FF82C29FF87EFAF2F856" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[1252,1348,1983,2006]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="134" phylum="Arthropoda" rank="species" species="tarang" status="sp. nov.">
<emphasis id="F4E1EAF7FFD2FF82C29FF87EFAF2F856" box="[1252,1348,1983,2006]" italics="true" pageId="19" pageNumber="134">B. tarang</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD2FF82C330F87FFA2AF856" bold="true" box="[1355,1436,1982,2006]" pageId="19" pageNumber="134">
<taxonomicNameLabel id="EFD2578CFFD2FF82C330F87FFA2AF856" box="[1355,1436,1982,2006]" pageId="19" pageNumber="134" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10.
</paragraph>
</caption>
<caption id="92EA666DFFD5FF85C6ECF97FFAA1F97A" ID-DOI="http://doi.org/10.5281/zenodo.4719961" ID-Zenodo-Dep="4719961" httpUri="https://zenodo.org/record/4719961/files/figure.png" pageId="20" pageNumber="135" startId="20.[151,250,1726,1750]" targetBox="[151,1435,199,1702]" targetPageId="20">
<paragraph id="C62A36E5FFD5FF85C6ECF97FFAA1F97A" blockId="20.[151,1437,1726,1786]" pageId="20" pageNumber="135">
<emphasis id="F4E1EAF7FFD5FF85C6ECF97FFE96F956" bold="true" box="[151,288,1726,1750]" pageId="20" pageNumber="135">FIGURE 15.</emphasis>
Chelicerae in lateral views. A.
<taxonomicName id="01954D66FFD5FF85C427F97EFD7EF956" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[604,712,1726,1750]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="135" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFD5FF85C427F97EFD7EF956" box="[604,712,1726,1750]" italics="true" pageId="20" pageNumber="135">B. bachma</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD5FF85C4B5F97FFCA9F956" bold="true" box="[718,799,1726,1750]" pageId="20" pageNumber="135">
<taxonomicNameLabel id="EFD2578CFFD5FF85C4B5F97FFCA9F956" box="[718,799,1726,1750]" pageId="20" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; B.
<taxonomicName id="01954D66FFD5FF85C531F97EFC63F956" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[842,981,1726,1750]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="135" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFD5FF85C531F97EFC63F956" box="[842,981,1726,1750]" italics="true" pageId="20" pageNumber="135">B. cucphuong</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD5FF85C5A0F97FFB9DF956" bold="true" box="[987,1067,1726,1750]" pageId="20" pageNumber="135">
<taxonomicNameLabel id="EFD2578CFFD5FF85C5A0F97FFB9DF956" box="[987,1067,1726,1750]" pageId="20" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; C.
<taxonomicName id="01954D66FFD5FF85C22DF97EFB0CF956" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[1110,1210,1727,1750]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="135" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
<emphasis id="F4E1EAF7FFD5FF85C22DF97EFB0CF956" box="[1110,1210,1727,1750]" italics="true" pageId="20" pageNumber="135">B. jaegeri</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD5FF85C2BBF97FFAA6F956" bold="true" box="[1216,1296,1726,1750]" pageId="20" pageNumber="135">
<taxonomicNameLabel id="EFD2578CFFD5FF85C2BBF97FFAA6F956" box="[1216,1296,1726,1750]" pageId="20" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; D.
<taxonomicName id="01954D66FFD5FF85C346F97EFA2BF956" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[1341,1437,1726,1750]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="135" phylum="Arthropoda" rank="species" species="kachin" status="sp. nov.">
<emphasis id="F4E1EAF7FFD5FF85C346F97EFA2BF956" box="[1341,1437,1726,1750]" italics="true" pageId="20" pageNumber="135">B. kachin</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD5FF85C6ECF923FF5EF97A" bold="true" box="[151,232,1762,1786]" pageId="20" pageNumber="135">
<taxonomicNameLabel id="EFD2578CFFD5FF85C6ECF923FF5EF97A" box="[151,232,1762,1786]" pageId="20" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; E.
<taxonomicName id="01954D66FFD5FF85C769F922FEDEF97A" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[274,360,1763,1786]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="135" phylum="Arthropoda" rank="species" species="putao" status="sp. nov.">
<emphasis id="F4E1EAF7FFD5FF85C769F922FEDEF97A" box="[274,360,1763,1786]" italics="true" pageId="20" pageNumber="135">B. putao</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD5FF85C715F923FE08F97A" bold="true" box="[366,446,1762,1786]" pageId="20" pageNumber="135">
<taxonomicNameLabel id="EFD2578CFFD5FF85C715F923FE08F97A" box="[366,446,1762,1786]" pageId="20" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; F.
<taxonomicName id="01954D66FFD5FF85C79EF922FDF3F97A" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[485,581,1763,1786]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="135" phylum="Arthropoda" rank="species" species="tarang" status="sp. nov.">
<emphasis id="F4E1EAF7FFD5FF85C79EF922FDF3F97A" box="[485,581,1763,1786]" italics="true" pageId="20" pageNumber="135">B. tarang</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD5FF85C430F923FD2DF97A" bold="true" box="[587,667,1762,1786]" pageId="20" pageNumber="135">
<taxonomicNameLabel id="EFD2578CFFD5FF85C430F923FD2DF97A" box="[587,667,1762,1786]" pageId="20" pageNumber="135" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
ba = bulbal apophysis, da = distal apophysis. Scale bars: 0.05.
</paragraph>
</caption>
<caption id="92EA666DFFD4FF84C6ECF93EFB78F8BC" ID-DOI="http://doi.org/10.5281/zenodo.4719963" ID-Zenodo-Dep="4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="21" pageNumber="136" startId="21.[151,250,1791,1815]" targetBox="[151,1435,181,1768]" targetPageId="21">
<paragraph id="C62A36E5FFD4FF84C6ECF93EFB78F8BC" blockId="21.[151,1437,1791,1852]" pageId="21" pageNumber="136">
<emphasis id="F4E1EAF7FFD4FF84C6ECF93EFE97F897" bold="true" box="[151,289,1791,1815]" pageId="21" pageNumber="136">FIGURE 16.</emphasis>
Distribution records of six new species of the genus
<taxonomicName id="01954D66FFD4FF84C53EF8C1FC28F898" authorityName="Thorell" authorityYear="1898" box="[837,926,1792,1816]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="F4E1EAF7FFD4FF84C53EF8C1FC28F898" box="[837,926,1792,1816]" italics="true" pageId="21" pageNumber="136">Belisana</emphasis>
</taxonomicName>
from Southern Asia. 1.
<taxonomicName id="01954D66FFD4FF84C2EDF8C1FAB2F898" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[1174,1284,1792,1816]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="136" phylum="Arthropoda" rank="species" species="bachma" status="sp. nov.">
<emphasis id="F4E1EAF7FFD4FF84C2EDF8C1FAB2F898" box="[1174,1284,1792,1816]" italics="true" pageId="21" pageNumber="136">B. bachma</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD4FF84C370F8C1FAEBF898" bold="true" box="[1291,1373,1792,1816]" pageId="21" pageNumber="136">
<taxonomicNameLabel id="EFD2578CFFD4FF84C370F8C1FAEBF898" box="[1291,1373,1792,1816]" pageId="21" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 2.
<taxonomicName id="01954D66FFD4FF84C3FDF8C1FEB0F8BC" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="136" phylum="Arthropoda" rank="species" species="cucphuong" status="sp. nov.">
<emphasis id="F4E1EAF7FFD4FF84C3FDF8C1FEB0F8BC" italics="true" pageId="21" pageNumber="136">B. cucphuong</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD4FF84C776F8E5FEEBF8BC" bold="true" box="[269,349,1828,1852]" pageId="21" pageNumber="136">
<taxonomicNameLabel id="EFD2578CFFD4FF84C776F8E5FEEBF8BC" box="[269,349,1828,1852]" pageId="21" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 3.
<taxonomicName id="01954D66FFD4FF84C7FFF8E5FE5EF8BB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[388,488,1828,1851]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="136" phylum="Arthropoda" rank="species" species="jaegeri" status="sp. nov.">
<emphasis id="F4E1EAF7FFD4FF84C7FFF8E5FE5EF8BB" box="[388,488,1828,1851]" italics="true" pageId="21" pageNumber="136">B. jaegeri</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD4FF84C794F8E5FD89F8BC" bold="true" box="[495,575,1828,1852]" pageId="21" pageNumber="136">
<taxonomicNameLabel id="EFD2578CFFD4FF84C794F8E5FD89F8BC" box="[495,575,1828,1852]" pageId="21" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 4.
<taxonomicName id="01954D66FFD4FF84C41DF8E5FD70F8BC" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[614,710,1828,1852]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="136" phylum="Arthropoda" rank="species" species="kachin" status="sp. nov.">
<emphasis id="F4E1EAF7FFD4FF84C41DF8E5FD70F8BC" box="[614,710,1828,1852]" italics="true" pageId="21" pageNumber="136">B. kachin</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD4FF84C4B7F8E5FCABF8BC" bold="true" box="[716,797,1828,1852]" pageId="21" pageNumber="136">
<taxonomicNameLabel id="EFD2578CFFD4FF84C4B7F8E5FCABF8BC" box="[716,797,1828,1852]" pageId="21" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 5.
<taxonomicName id="01954D66FFD4FF84C53FF8E5FC2CF8BB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[836,922,1828,1851]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="136" phylum="Arthropoda" rank="species" species="putao" status="sp. nov.">
<emphasis id="F4E1EAF7FFD4FF84C53FF8E5FC2CF8BB" box="[836,922,1828,1851]" italics="true" pageId="21" pageNumber="136">B. putao</emphasis>
</taxonomicName>
<emphasis id="F4E1EAF7FFD4FF84C5DBF8E5FC47F8BC" bold="true" box="[928,1009,1828,1852]" pageId="21" pageNumber="136">
<taxonomicNameLabel id="EFD2578CFFD4FF84C5DBF8E5FC47F8BC" box="[928,1009,1828,1852]" pageId="21" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 6.
<taxonomicName id="01954D66FFD4FF84C263F8E5FBCEF8BB" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[1048,1144,1828,1851]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="136" phylum="Arthropoda" rank="species" species="tarang" status="sp. nov.">
<emphasis id="F4E1EAF7FFD4FF84C263F8E5FBCEF8BB" box="[1048,1144,1828,1851]" italics="true" pageId="21" pageNumber="136">B. tarang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFD2578CFFD4FF84C205F8E5FB78F8BC" box="[1150,1230,1828,1852]" pageId="21" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="C62A36E5FFD4FF84C6BCF8A8FB65F828" blockId="21.[151,1436,1897,2033]" pageId="21" pageNumber="136">
<emphasis id="F4E1EAF7FFD4FF84C6BCF8A8FEF7F803" bold="true" box="[199,321,1897,1923]" pageId="21" pageNumber="136">Variation:</emphasis>
In male
<typeStatus id="192E8847FFD4FF84C7D9F8ABFDB4F804" box="[418,514,1898,1924]" pageId="21" pageNumber="136" type="paratype">paratype</typeStatus>
(IZCAS Ar41917): Leg I: 10.57 (2.66 + 0.28 + 2.72 + 3.75 + 1.16); tibia I L/d: 45. Retrolateral trichobothria of tibia Iat 7% proximally; tarsus Iwith 16 distinct pseudopodomeres.
</paragraph>
<paragraph id="C62A36E5FFD4FF84C6BCF873FABFF84C" blockId="21.[151,1436,1897,2033]" box="[199,1289,1969,1996]" pageId="21" pageNumber="136">
<emphasis id="F4E1EAF7FFD4FF84C6BCF873FE33F84B" bold="true" box="[199,389,1969,1996]" pageId="21" pageNumber="136">Natural history.</emphasis>
The species was found in leaf litter and collected by small plot direct searching.
</paragraph>
</subSubSection>
<subSubSection id="8E8F656EFFD4FF84C6BCF814FC34F871" box="[199,898,2005,2033]" pageId="21" pageNumber="136" type="distribution">
<paragraph id="C62A36E5FFD4FF84C6BCF814FC34F871" blockId="21.[151,1436,1897,2033]" box="[199,898,2005,2033]" pageId="21" pageNumber="136">
<emphasis id="F4E1EAF7FFD4FF84C6BCF814FED7F86F" bold="true" box="[199,353,2005,2031]" pageId="21" pageNumber="136">Distribution.</emphasis>
<collectingCountry id="BE827675FFD4FF84C71CF817FE62F870" box="[359,468,2006,2032]" name="Indonesia" pageId="21" pageNumber="136">Indonesia</collectingCountry>
(
<collectingRegion id="0451F807FFD4FF84C798F817FDF3F870" box="[483,581,2006,2032]" country="Indonesia" name="Sumatera Utara" pageId="21" pageNumber="136">Sumatra</collectingRegion>
,
<typeStatus id="192E8847FFD4FF84C42BF817FD36F870" box="[592,640,2006,2032]" pageId="21" pageNumber="136">type</typeStatus>
locality only;
<figureCitation id="5EAE2A60FFD4FF84C558F817FCC3F871" box="[803,885,2006,2033]" captionStart="FIGURE 16" captionStartId="21.[151,250,1791,1815]" captionTargetBox="[151,1435,181,1768]" captionTargetId="figure-17@21.[151,1435,181,1768]" captionTargetPageId="21" captionText="FIGURE 16. Distribution records of six new species of the genus Belisana from Southern Asia. 1. B. bachma sp. nov.; 2. B. cucphuong sp. nov.; 3. B. jaegeri sp. nov.; 4. B. kachin sp. nov.; 5. B. putao sp. nov.; 6. B. tarang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4719963" httpUri="https://zenodo.org/record/4719963/files/figure.png" pageId="21" pageNumber="136">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>