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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.88.807" ID-GBIF-Dataset="1858994c-378e-4e6e-bec9-ee565e3b3be1" ID-Pensoft-Pub="1313-2970-88-1" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-88-1" ModsDocOrigin="ZooKeys 88" ModsDocTitle="Family-group names in Coleoptera (Insecta)" checkinTime="1451250740194" checkinUser="pensoft" docAuthor="Bouchard, Patrice, Bousquet, Yves, Davies, Anthony E., Alonso-Zarazaga, Miguel A., Lawrence, John F., Lyal, Chris H. C., Newton, Alfred F., Reid, Chris A. M., Schmitt, Michael, Ślipinski, S. Adam &amp; Smith, Andrew B. T." docDate="2011" docId="4E0D2170BE1D3D28DBC6FB967C191A7F" docLanguage="en" docName="ZooKeys 88: 1-972" docOrigin="ZooKeys 88" docSource="http://dx.doi.org/10.3897/zookeys.88.807" docTitle="Meroscelisini Thomson 1861" docType="treatment" docVersion="6" lastPageNumber="201" masterDocId="FFC1FFDA4E1CE6158E11FFCD0115FF8E" masterDocTitle="Family-group names in Coleoptera (Insecta)" masterLastPageNumber="972" masterPageNumber="1" pageNumber="200" updateTime="1661732154594" updateUser="admin">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Family-group names in Coleoptera (Insecta)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bouchard, Patrice</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bousquet, Yves</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davies, Anthony E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Alonso-Zarazaga, Miguel A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lawrence, John F.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lyal, Chris H. C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Newton, Alfred F.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reid, Chris A. M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schmitt, Michael</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ślipinski, S. Adam</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Andrew B. T.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>88</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>972</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.88.807</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.88.807</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-88-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="159363562" LSID="urn:lsid:plazi:treatment:4E0D2170BE1D3D28DBC6FB967C191A7F" httpUri="http://treatment.plazi.org/id/4E0D2170BE1D3D28DBC6FB967C191A7F" lastPageId="200" lastPageNumber="201" pageId="199" pageNumber="200">
<subSubSection pageId="199" pageNumber="200" type="nomenclature">
<paragraph pageId="199" pageNumber="200">
Tribe
<taxonomicName authority="Thomson, 1861" authorityName="Thomson" authorityYear="1861" lsidName="" pageId="199" pageNumber="200" rank="tribe" tribe="Meroscelisini">Meroscelisini Thomson, 1861</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection lastPageId="200" lastPageNumber="201" pageId="199" pageNumber="200" type="reference_group">
<paragraph pageId="199" pageNumber="200">
<taxonomicName genus="Meroscelisitae" lsidName="Meroscelisitae" pageId="199" pageNumber="200" rank="genus">Meroscelisitae</taxonomicName>
J. Thomson, 1861: 299 [stem: Meroscelis-]. Type genus:
<taxonomicName class="Insecta" family="Cerambycidae" genus="Meroscelisus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Meroscelisus" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="genus">Meroscelisus</taxonomicName>
Audinet-Serville, 1832.
</paragraph>
<paragraph pageId="199" pageNumber="200">
<taxonomicName genus="Tragosomites" lsidName="Tragosomites" pageId="199" pageNumber="200" rank="genus">*Tragosomites</taxonomicName>
Fairmaire, 1864: 119 [stem: Tragosomat-]. Type genus:
<taxonomicName class="Insecta" family="Cerambycidae" genus="Tragosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tragosoma" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="genus">Tragosoma</taxonomicName>
Audinet-Serville, 1832. Comment: original vernacular name unavailable (Art. 11.7.2): subsequently used in latinized form but not generally attributed to Fairmaire (1864); incorrect original stem formation, not in prevailing usage.
</paragraph>
<paragraph pageId="199" pageNumber="200">
<taxonomicName genus="Tragosomitae" lsidName="Tragosomitae" pageId="199" pageNumber="200" rank="genus">Tragosomitae</taxonomicName>
J. Thomson, 1864: 286 [stem: Tragosomat-]. Type genus:
<taxonomicName class="Insecta" family="Cerambycidae" genus="Tragosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tragosoma" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="genus">Tragosoma</taxonomicName>
Audinet-Serville, 1832. Comment: incorrect original stem formation, not in prevailing usage.
</paragraph>
<paragraph pageId="199" pageNumber="200">
<taxonomicName genus="Closterides" lsidName="Closterides" pageId="199" pageNumber="200" rank="genus">
<normalizedToken originalValue="Clostérides">Closterides</normalizedToken>
</taxonomicName>
Lacordaire, 1868: 149 [stem: Closter-]. Type genus:
<taxonomicName class="Insecta" family="Cerambycidae" genus="Closterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Closterus" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="genus">Closterus</taxonomicName>
Audinet-Serville, 1832. Comment: original vernacular name available (Art. 11.7.2): first used in latinized form by Pascoe (1869b: 676, as
<taxonomicName lsidName="" pageId="199" pageNumber="200" rank="subFamily" subFamily="Closterinae">Closterinae</taxonomicName>
), generally accepted as in Lameere (1913: 81, as
<taxonomicName genus="Closteri" lsidName="Closteri" pageId="199" pageNumber="200" rank="genus">Closteri</taxonomicName>
).
</paragraph>
<paragraph lastPageId="200" lastPageNumber="201" pageId="199" pageNumber="200">
<taxonomicName genus="Monodesmides" lsidName="Monodesmides" pageId="199" pageNumber="200" rank="genus">*Monodesmides</taxonomicName>
Lacordaire, 1868: 157 [stem: Monodesm-]. Type genus:
<taxonomicName class="Insecta" family="Cerambycidae" genus="Monodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monodesmus" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="genus">Monodesmus</taxonomicName>
Audinet-Serville, 1832. Comment: original vernacular name unavail
<pageBreakToken pageId="200" pageNumber="201" start="start">able</pageBreakToken>
(Art. 11.7.2): subsequently used in latinized form but not generally attributed to Lacordaire (1868).
</paragraph>
<paragraph pageId="200" pageNumber="201">
<taxonomicName lsidName="" pageId="200" pageNumber="201" rank="subFamily" subFamily="Monodesminae">Monodesminae</taxonomicName>
Gahan, 1890: 299 [stem: Monodesm-]. Type genus:
<taxonomicName class="Insecta" family="Cerambycidae" genus="Monodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monodesmus" order="Coleoptera" pageId="200" pageNumber="201" phylum="Arthropoda" rank="genus">Monodesmus</taxonomicName>
Audinet-Serville, 1832.
</paragraph>
<paragraph pageId="200" pageNumber="201">
<taxonomicName genus="Luluina" lsidName="Luluina" pageId="200" pageNumber="201" rank="genus">Luluina</taxonomicName>
Gilmour, 1956: 222 [stem: Lulu-]. Type genus:
<taxonomicName class="Insecta" family="Cerambycidae" genus="Lulua" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lulua" order="Coleoptera" pageId="200" pageNumber="201" phylum="Arthropoda" rank="genus">Lulua</taxonomicName>
Burgeon, 1931.
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ENCODING="UTF-8" ID-DOI="https://doi.org/10.5962/bhl.title.542" ID-GBIF-Dataset="57ea6fdb-2533-44e5-b60b-736758a88d19" ID-Zenodo-Dep="3922206" ID-ZooBank="2C6327E1-5560-4DB4-B9CA-76A0FA03D975" checkinTime="1593218702860" checkinUser="admin" docAuthor="Linnaeus, Carolus" docDate="1758" docId="4E0D24815A67D5AD0CDA882204DDB49C" docLanguage="de" docName="Linnaeus1758Treatments.xml" docOrigin="Stockholm: Laurentius Salvius" docSource="https://archive.org/download/mobot31753000798865/mobot31753000798865.pdf" docTitle="Cimex striatus Linnaeus, 1758, spec. nov." docType="treatment" docUuid="CB940F04-030A-441F-A29F-5ED8AF155005" docUuidSource="ZooBank" docVersion="13" lastPageNumber="449" masterDocId="3F896AFC21C459639D0D71D9EDB8875C" masterDocTitle="Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis" pageNumber="449" updateTime="1643600433100" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systema Naturae per regna tria naturae: secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Linnaeus, Carolus</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:originInfo>
<mods:dateIssued>1758</mods:dateIssued>
<mods:publisher>Laurentius Salvius</mods:publisher>
<mods:place>
<mods:placeTerm>Stockholm</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:location>
<mods:url>https://archive.org/download/mobot31753000798865/mobot31753000798865.pdf</mods:url>
</mods:location>
<mods:classification>book</mods:classification>
<mods:identifier type="ZooBank">2C6327E1-5560-4DB4-B9CA-76A0FA03D975</mods:identifier>
<mods:identifier type="DOI">https://doi.org/10.5962/bhl.title.542</mods:identifier>
<mods:identifier type="Zenodo-Dep">3922206</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3919143" ID-GBIF-Taxon="164811961" ID-Zenodo-Dep="3919143" LSID="urn:lsid:zoobank.org:act:CB940F04-030A-441F-A29F-5ED8AF155005" httpUri="http://treatment.plazi.org/id/4E0D24815A67D5AD0CDA882204DDB49C" lastPageNumber="449" pageNumber="449" sourcePageUrl="https://www.biodiversitylibrary.org/page/727360">
<subSubSection type="nomenclature">
<paragraph pageNumber="449">
<taxonomicName LSID="urn:lsid:zoobank.org:act:CB940F04-030A-441F-A29F-5ED8AF155005" authorityName="Linnaeus" authorityYear="1758" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="striatus" status="spec. nov." zbkClass="Insecta" zbkKingdom="Animalia" zbkOrder="Hemiptera">Cimex striatus</taxonomicName>
[
<taxonomicNameLabel rank="species">spec. nov.</taxonomicNameLabel>
]
</paragraph>
</subSubSection>
<subSubSection type="description">
<paragraph pageNumber="449">
C. oblongus niger, elytris luteis fusco striatis apice pedibusque rubris.
<emphasis italics="true">Fn. svec.</emphasis>
680.
</paragraph>
</subSubSection>
<subSubSection type="distribution">
<paragraph pageNumber="449">
<emphasis italics="true">Habitat in</emphasis>
Ulmi
<emphasis italics="true">campestris foliis.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.700.11784" ID-GBIF-Dataset="30d1c76a-41e1-4f66-9fb5-460bf381f752" ID-PMC="PMC5711039" ID-Pensoft-Pub="1313-2970-700-1" ID-PubMed="29358897" ID-ZBK="EBA4322720AD4CFFA04E8D2542DDA3D6" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-700-1" ModsDocOrigin="ZooKeys 700" ModsDocTitle="Revision of the ant genus Melophorus (Hymenoptera, Formicidae)" checkinTime="1506009303549" checkinUser="pensoft" docAuthor="Heterick, Brian E., Castalanelli, Mark &amp; Shattuck, Steve O." docDate="2017" docId="4E0D8DD3535037838DDA6E9DE40BBC5B" docLanguage="en" docName="ZooKeys 700: 1-420" docOrigin="ZooKeys 700" docSource="http://dx.doi.org/10.3897/zookeys.700.11784" docTitle="Melophorus griseus Heterick, Castalanelli &amp; Shattuck, sp. n." docType="treatment" docUuid="5B0258CA-A311-4AC4-BBD7-B1DA09D0CBCB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="98" masterDocId="FF93E60CA711CB7BFF95D521FFFDFF9A" masterDocTitle="Revision of the ant genus Melophorus (Hymenoptera, Formicidae)" masterLastPageNumber="420" masterPageNumber="1" pageNumber="96" updateTime="1668164819442" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the ant genus Melophorus (Hymenoptera, Formicidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Heterick, Brian E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Castalanelli, Mark</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shattuck, Steve O.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>700</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>420</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.700.11784</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.700.11784</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-700-1</mods:identifier>
<mods:identifier type="ZBK">EBA4322720AD4CFFA04E8D2542DDA3D6</mods:identifier>
<mods:identifier type="ZooBank">EBA4322720AD4CFFA04E8D2542DDA3D6</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="150814512" LSID="urn:lsid:zoobank.org:act:5B0258CA-A311-4AC4-BBD7-B1DA09D0CBCB" httpUri="http://treatment.plazi.org/id/4E0D8DD3535037838DDA6E9DE40BBC5B" lastPageId="97" lastPageNumber="98" pageId="95" pageNumber="96">
<subSubSection pageId="95" pageNumber="96" type="nomenclature">
<paragraph pageId="95" pageNumber="96">
<taxonomicName LSID="http://zoobank.org/5B0258CA-A311-4AC4-BBD7-B1DA09D0CBCB" authority="Heterick, Castalanelli &amp; Shattuck" class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus griseus" order="Hymenoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="griseus">Melophorus griseus Heterick, Castalanelli &amp; Shattuck</taxonomicName>
<taxonomicNameLabel pageId="95" pageNumber="96">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="95" pageNumber="96" type="types">
<paragraph pageId="95" pageNumber="96">Types.</paragraph>
<paragraph pageId="95" pageNumber="96">
Holotype minor worker from Argyle Diamonds via Kununurra, Western Australia, September 1989, A. T. Postle, Site 15
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus" order="Hymenoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="genus">Melophorus</taxonomicName>
sp. [JDM32-004555] (WAM). Paratypes: 2 minor workers from Gibb River Road turnoff
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-15.834933">15°50.096'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="128.31107">128°18.664'E</geoCoordinate>
, Western Australia, 9 February 2013 (3.40pm), Heterick, B.E., Sample 35: parking area with tall grass nearby, ABRS M21 (WAM).
</paragraph>
</subSubSection>
<subSubSection pageId="95" pageNumber="96" type="materials_examined">
<paragraph pageId="95" pageNumber="96">Other material examined.</paragraph>
<paragraph pageId="95" pageNumber="96">Western Australia: Gibb River Rd turnoff (Heterick, B.E., B.E. [M217/M218]).</paragraph>
</subSubSection>
<subSubSection pageId="95" pageNumber="96" type="diagnosis">
<paragraph pageId="95" pageNumber="96">Diagnosis.</paragraph>
<paragraph pageId="95" pageNumber="96">
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus griseus" order="Hymenoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="griseus">Melophorus griseus</taxonomicName>
is a member of the
<taxonomicName lsidName="M. aeneovirens" pageId="95" pageNumber="96" rank="species" species="aeneovirens">M. aeneovirens</taxonomicName>
species-group (in full-face view, the anterior clypeal margin convex, apron-like and covering whole or part of the retracted mandible, except in
<taxonomicName lsidName="M. nemophilus" pageId="95" pageNumber="96" rank="species" species="nemophilus">M. nemophilus</taxonomicName>
, the medial clypeal sector often produced so that it is protrusive when seen in profile; the psammophore frequently with coarse and well-separated ammochaetae, these always placed on or just above anterior margin; in profile, the propodeum elongate and oblique or broadly rounded), and the
<taxonomicName lsidName="M. aeneovirens" pageId="95" pageNumber="96" rank="species" species="aeneovirens">M. aeneovirens</taxonomicName>
species-complex (in full-face view, psammophore ranged along or just above anterior margin of clypeus and following the curve of the margin; anterior margin of clypeus broadly medially produced, and often with central notch that may be deeply impressed, but is never acuminate at its midpoint; metatibia with maximum of two rows of preapical spines). In
<taxonomicName lsidName="M. griseus" pageId="95" pageNumber="96" rank="species" species="griseus">M. griseus</taxonomicName>
the tibiae possess fine, appressed pubescence in addition to stout, socketed, appressed to subdecumbent setae
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus griseus" order="Hymenoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="griseus">Melophorus griseus</taxonomicName>
differs from
<taxonomicName lsidName="M. curtus" pageId="95" pageNumber="96" rank="species" species="curtus">M. curtus</taxonomicName>
by the features mentioned above. The Frontal carinae of the minor worker is raised and laminate at the edges, the frontal triangle is narrowly semi-oval, the sides of head are concave below eyes, giving it a bell-shaped appearance and, in profile, the in profile, pronotum and mesonotum form a gentle curve. These features separate it from its likely nearest relative,
<taxonomicName lsidName="M. gibbosus" pageId="95" pageNumber="96" rank="species" species="gibbosus">M. gibbosus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="97" lastPageNumber="98" pageId="95" pageNumber="96" type="description">
<paragraph pageId="95" pageNumber="96">Minor worker description.</paragraph>
<paragraph lastPageId="97" lastPageNumber="98" pageId="95" pageNumber="96">
Head. Head approximately oval, sides of head divergent towards mandibular articulations; posterior margin of head planar or weakly convex; frons matt or with weak sheen, microreticulate or microreticulate-shagreenate;
<pageBreakToken pageId="96" pageNumber="97" start="start">frons</pageBreakToken>
consisting of appressed pubescence, with many short, unmodified, erect setae. Eye moderate (eye length 0.20-0.49 length of side of head capsule); in full-face view, eyes set above midpoint of head capsule; in profile, eye set anteriad of midline of head capsule; eyes elliptical or slightly reniform. In full-face view, frontal carinae straight, divergent posteriad; frontal lobes straight, elevated. Anteromedial clypeal margin narrowly convex and protruding, clypeal midpoint distinctly notched; clypeal psammophore set at or just above anterior clypeal margin; palp formula 6,4. Five mandibular teeth in minor worker; mandibles triangular, weakly incurved; third mandibular tooth distinctly shorter than apical tooth and teeth numbers two and four; masticatory margin of mandibles approximately vertical or weakly oblique. Mesosoma. Integument of pronotum, mesonotum and mesopleuron matt with indistinct shagreenate sculpture throughout; anterior mesosoma in profile broadly convex; appearance of erect pronotal setae short, (i.e., longest erect setae shorter than length of eye) and unmodified; in profile, metanotal groove shallow, broadly V or U-shaped; propodeum matt or with a weak sheen and indistinctly shagreenate; propodeum smoothly rounded or with indistinct angle; propodeal dorsum and declivity confluent; erect propodeal setae present and abundant (greater than 12); appressed propodeal setulae long and closely aligned, creating pubescence; propodeal spiracle situated at least twice its width from
<pageBreakToken pageId="97" pageNumber="98" start="start">the</pageBreakToken>
declivitous face of propodeum, and shorter (length &lt;0.50
<normalizedToken originalValue="×">x</normalizedToken>
height of propodeum). Petiole. In profile, petiolar node squamiform; in full-face view, shape of petiolar node generally rounded with median indentation or hollow; node matt with indistinct microsculpture. Gaster weakly shining with indistinct shagreenation; pilosity of first gastral tergite consisting of thick, appressed setae that form pubescence, interspersed with numerous short, bristly, erect setae. General characters. Colour brown, gaster slightly darker than foreparts.
</paragraph>
</subSubSection>
<subSubSection pageId="97" pageNumber="98" type="description">
<paragraph pageId="97" pageNumber="98">Measurements.</paragraph>
<paragraph pageId="97" pageNumber="98">Worker (n =1): CI 91; EI 23; EL 0.23; HL 1.10; HW 1.01; ML 1.58; MTL 1; PpH 0.166; PpL 0.62; SI 136; SL 1.40.</paragraph>
</subSubSection>
<subSubSection pageId="97" pageNumber="98" type="comments">
<paragraph pageId="97" pageNumber="98">Comments.</paragraph>
<paragraph pageId="97" pageNumber="98">
This ant is differentiated from
<taxonomicName lsidName="M. canus" pageId="97" pageNumber="98" rank="species" species="canus">M. canus</taxonomicName>
in lacking the long, silky setae and the dentiform mandible, the mandible in this case being five-toothed. The lack of a thoracic hump also helps to separate the minor worker from the minor worker of
<taxonomicName lsidName="M. gibbosus" pageId="97" pageNumber="98" rank="species" species="gibbosus">M. gibbosus</taxonomicName>
. Most workers have bristly, erect setae on the antennae and tibiae. Workers collected in long grass near Dunham River, Western Australia, were extremely timid solitary foragers among (much more numerous) workers of
<taxonomicName class="Insecta" family="Formicidae" genus="Iridomyrmex" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Iridomyrmex minor" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="minor">Iridomyrmex minor</taxonomicName>
, which they resemble in the field. The species is both genetically and morphologically closely related to
<taxonomicName lsidName="M. canus" pageId="97" pageNumber="98" rank="species" species="canus">M. canus</taxonomicName>
but is more restricted in its distribution, being apparently confined to the top end, with collections made in NT, QLD and WA (TERC = 'Group
<normalizedToken originalValue="L’">L'</normalizedToken>
) and the Kimberley region in Western Australia (WAM).
</paragraph>
</subSubSection>
<subSubSection pageId="97" pageNumber="98" type="etymology">
<paragraph pageId="97" pageNumber="98">Etymology.</paragraph>
<paragraph pageId="97" pageNumber="98">
Latin
<taxonomicName lsidName="griseus" pageId="97" pageNumber="98" rank="species" species="griseus">griseus</taxonomicName>
(
<normalizedToken originalValue="‘gray’">'gray'</normalizedToken>
); adjective in the nominative singular.
</paragraph>
<caption pageId="97" pageNumber="98">
<paragraph pageId="97" pageNumber="98">
Figure 14.
<taxonomicName class="Insecta" family="Formicidae" genus="Melophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melophorus griseus" order="Hymenoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="griseus">Melophorus griseus</taxonomicName>
sp. n.: minor worker holotype (JDM32-004555) frons (a), profile (b) and dorsum (c); distribution map for the species (d). Low resolution scale bars: 0.5 mm (
<normalizedToken originalValue="a–c">a-c</normalizedToken>
).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>

65
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<taxonomicName id="0810BE611585E8843541FC0B9DDD5D2C" ID-CoL="5JJQF" authority="Thomas 1899" authorityName="Thomas" authorityYear="1899" class="Mammalia" family="Didelphidae" genus="Marmosa" kingdom="Animalia" order="Didelphimorphia" pageNumber="9" phylum="Chordata" rank="subSpecies" species="robinsoni" subSpecies="simonsi">
<emphasis id="5F5CAA0D2A85FFF897914FAE574BD9E2" italics="true" pageNumber="9">Marmosa robinsoni</emphasis>
subsp.
<emphasis id="ACD512EFE056F1F091D111F461FCBF82" italics="true" pageNumber="9">simonsi</emphasis>
Thomas 1899
</taxonomicName>
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<emphasis id="1E4E84B6194FB667249F1E576F012F9C" inLineHeading="true" pageNumber="9">Synonyms:</emphasis>
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<emphasis id="52ACDC16F306B21F47D48F41213AF8A4" italics="true" pageNumber="9">Marmosa robinsoni</emphasis>
subsp.
<emphasis id="57BBC579FAE883AF5C29D797B9065944" italics="true" pageNumber="9">mimetra</emphasis>
Thomas 1921
</taxonomicName>
.
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214
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<mods:title>A survey of the spider genus Dysdera Latreille, 1804 (Araneae, Dysderidae) in Iran, with fourteen new species and notes on two fossil genera</mods:title>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Turku, Finland</mods:affiliation>
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<mods:namePart>Marusik, Yuri M.</mods:namePart>
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<mods:affiliation>Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Szűts, Tamas</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8954-0641</mods:nameIdentifier>
<mods:affiliation>Department of Ecology, University of Veterinary Medicine Budapest, Rottenbiller u. 50, Budapest, 1077, Hungary</mods:affiliation>
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<paragraph pageId="0" pageNumber="43">
<taxonomicName LSID="https://zoobank.org/5CCD54FC-78A6-4AA9-A521-C53C8C70F140" authority="Zamani &amp; Marusik &amp; Szűts, 2023" authorityName="Zamani &amp; Marusik &amp; Szűts" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera persica" order="Araneae" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="persica" status="sp. nov.">Dysdera persica</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="43">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection pageId="0" pageNumber="43" type="description">
<paragraph pageId="0" pageNumber="43">
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Female (A-C) and male (D-F) of Dysdera persica sp. nov., habitus A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1146.97517.figure15" httpUri="https://binary.pensoft.net/fig/806626" pageId="0" pageNumber="43">Figs 15A-F</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Male of Dysdera persica sp. nov., bulb A retrolateral view B prolateral view C anterior view D posterior view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1146.97517.figure16" httpUri="https://binary.pensoft.net/fig/806627" pageId="0" pageNumber="43">, 16A-D</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Female of Dysdera persica sp. nov., endogyne A ventral view B dorsal view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1146.97517.figure17" httpUri="https://binary.pensoft.net/fig/806628" pageId="0" pageNumber="43">, 17A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="type material">
<paragraph pageId="0" pageNumber="43">Type material.</paragraph>
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" italics="true" pageId="0" pageNumber="43">Holotype</emphasis>
♂ (ZMUT), Iran: Golestan Province: Shast Kalateh,
<geoCoordinate degrees="36" direction="north" minutes="45" orientation="latitude" precision="925" value="36.75">36°45'N</geoCoordinate>
,
<geoCoordinate degrees="54" direction="east" minutes="21" orientation="longitude" precision="925" value="54.35">54°21'E</geoCoordinate>
, 2017 (R. Rafiei-Jahed).
<emphasis bold="true" italics="true" pageId="0" pageNumber="43">Paratypes</emphasis>
: 7♂5♀ (ZMUT), same data as the holotype; 1♂ (ZMUT), Mazandaran Province: Polur,
<geoCoordinate degrees="35" direction="north" minutes="50" orientation="latitude" precision="925" value="35.833332">35°50'N</geoCoordinate>
,
<geoCoordinate degrees="52" direction="east" minutes="03" orientation="longitude" precision="925" value="52.05">52°03'E</geoCoordinate>
, 10.2015 (A. Zamani); 1♂ (MHNG), Kiyasar,
<geoCoordinate degrees="36" direction="north" minutes="14" orientation="latitude" precision="925" value="36.233334">36°14'N</geoCoordinate>
,
<geoCoordinate degrees="53" direction="east" minutes="33" orientation="longitude" precision="925" value="53.55">53°33'E</geoCoordinate>
, 1500 m, 11.07.1975 (A. Senglet).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="etymology">
<paragraph pageId="0" pageNumber="43">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="43">The specific epithet is an adjective, referring to the historical region of the Middle East, located in eastern Mesopotamia, which is now Iran.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="diagnosis">
<paragraph pageId="0" pageNumber="43">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="43">
The male of this species differs from those of the other species of the
<taxonomicName authorityName="Kroneberg" authorityYear="1875" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera aculeata" order="Araneae" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="aculeata">
<emphasis italics="true" pageId="0" pageNumber="43">Dysdera aculeata</emphasis>
</taxonomicName>
group occurring in Iran by the extremely long bulb (especially psembolus, i.e., twice longer than tegulum), and by the tegulum with posterior margin 1.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than anterior margin (vs. equal or shorter in length). The female of
<taxonomicName lsidName="D. persica" pageId="0" pageNumber="43" rank="species" species="persica">
<emphasis italics="true" pageId="0" pageNumber="43">D. persica</emphasis>
</taxonomicName>
sp. nov. differs from those of its congeners by having the broadest dorsal arch, bearing almost angled anterior corners (vs. rounded).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="description">
<paragraph pageId="0" pageNumber="43">Description.</paragraph>
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Male (Holotype).</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Female (A-C) and male (D-F) of Dysdera persica sp. nov., habitus A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1146.97517.figure15" httpUri="https://binary.pensoft.net/fig/806626" pageId="0" pageNumber="43">15D-F</figureCitation>
. Total length 9.06. Carapace 4.19 long, 3.20 wide. Eye diameters: AME 0.14, PME 0.16, PLE 0.17. Carapace, sternum, chelicerae, labium, and maxillae reddish brown. Legs orange. Abdomen greyish, without any pattern. Spinnerets uniformly dark yellowish. Measurements of legs: I: 13.98 (3.96, 2.36, 3.43, 3.52, 0.71), II: 12.87 (3.60, 2.11, 3.07, 3.27, 0.82), III: 9.38 (2.74, 1.51, 1.84, 2.70, 0.59), IV: 12.37 (3.54, 1.95, 2.66, 3.40, 0.82). Spination: I: Fe: 2pl. II: Fe: 1pl. III: Fe: 1pl; Ti: 5pl, 3rl, 4v; Mt: 5d, 9pl, 5rl, 3v. IV: Fe: 4d; Ti: 4pl, 6rl, 1v; Mt: 5d, 6pl, 7rl, 6v. Ti III-IV with a row of thin rigid ventral setae.
</paragraph>
<caption doi="10.3897/zookeys.1146.97517.figure15" httpUri="https://binary.pensoft.net/fig/806626" pageId="0" pageNumber="43" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Figure 15.</emphasis>
Female (
<emphasis bold="true" pageId="0" pageNumber="43">A-C</emphasis>
) and male (
<emphasis bold="true" pageId="0" pageNumber="43">D-F</emphasis>
) of
<taxonomicName authorityName="Zamani &amp; Marusik &amp; Szűts" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera persica" order="Araneae" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="persica">
<emphasis italics="true" pageId="0" pageNumber="43">Dysdera persica</emphasis>
</taxonomicName>
sp. nov., habitus
<emphasis bold="true" pageId="0" pageNumber="43">A, D</emphasis>
dorsal view
<emphasis bold="true" pageId="0" pageNumber="43">B, E</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="43">C, F</emphasis>
lateral view. Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="43">
Palp as in Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Male of Dysdera persica sp. nov., bulb A retrolateral view B prolateral view C anterior view D posterior view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1146.97517.figure16" httpUri="https://binary.pensoft.net/fig/806627" pageId="0" pageNumber="43">16A-D</figureCitation>
; bulb ca. 2.3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide; tegulum bell-shaped, almost as long as wide; psembolus 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than tegulum; median crest triangular, ca. 2
<normalizedToken originalValue="×">x</normalizedToken>
shorter than length of psembolus, ca. 2.4
<normalizedToken originalValue="×">x</normalizedToken>
wider than high; posterior apophysis broad; incision between tegulum and psembolus present; retrolateral crest gradually rounded.
</paragraph>
<caption doi="10.3897/zookeys.1146.97517.figure16" httpUri="https://binary.pensoft.net/fig/806627" pageId="0" pageNumber="43" start="Figure 16" startId="F16">
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Figure 16.</emphasis>
Male of
<taxonomicName authorityName="Zamani &amp; Marusik &amp; Szűts" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera persica" order="Araneae" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="persica">
<emphasis italics="true" pageId="0" pageNumber="43">Dysdera persica</emphasis>
</taxonomicName>
sp. nov., bulb
<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
retrolateral view
<emphasis bold="true" pageId="0" pageNumber="43">B</emphasis>
prolateral view
<emphasis bold="true" pageId="0" pageNumber="43">C</emphasis>
anterior view
<emphasis bold="true" pageId="0" pageNumber="43">D</emphasis>
posterior view. Scale bars: 0.25 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Female.</emphasis>
Habitus as in Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Female (A-C) and male (D-F) of Dysdera persica sp. nov., habitus A, D dorsal view B, E ventral view C, F lateral view. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1146.97517.figure15" httpUri="https://binary.pensoft.net/fig/806626" pageId="0" pageNumber="43">15A-C</figureCitation>
. Total length 11.0. Carapace 4.25 long, 3.16 wide. Eye diameters: AME 0.17, PME 0.18, PLE 0.15. Carapace and chelicerae dark reddish violet; sternum, labium, and maxillae reddish brown. Leg reddish brown. Abdomen greyish, without any pattern. Spinnerets uniformly greyish. Measurements of legs: I: 11.02 (3.15, 1.98, 2.59, 2.51, 0.79), II: 9.61 (2.72, 1.76, 2.32, 2.14, 0.67), III: 8.07 (2.27, 1.33, 1.62, 2.14, 0.71), IV: 10.54 (3.02, 1.66, 2.15, 2.89, 0.82). Spination: I: Fe: 3pl. II: Fe: 1pl. III: Fe: 1d; Ti: 4pl, 4rl; Mt: 12pl, 5rl, 4v. IV: Fe: 4d; Ti: 5pl, 4rl, 1v; Mt: 14pl, 9rl, 3v. Ti III-IV with a row of thin rigid ventral setae.
</paragraph>
<paragraph pageId="0" pageNumber="43">
Endogyne as in Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Female of Dysdera persica sp. nov., endogyne A ventral view B dorsal view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1146.97517.figure17" httpUri="https://binary.pensoft.net/fig/806628" pageId="0" pageNumber="43">17A, B</figureCitation>
; length/width ratio ca. 1.9; receptacle 3.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, 1.2
<normalizedToken originalValue="×">x</normalizedToken>
wider than transverse bar; anterior angles rounded; dorsal arch trapezoidal; transverse bar straight, ca. 1.8
<normalizedToken originalValue="×">x</normalizedToken>
longer than receptacle; lateral edges with almost horizontal anterior margins, as long as width of receptacle; posterior diverticulum rectangular.
</paragraph>
<caption doi="10.3897/zookeys.1146.97517.figure17" httpUri="https://binary.pensoft.net/fig/806628" pageId="0" pageNumber="43" start="Figure 17" startId="F17">
<paragraph pageId="0" pageNumber="43">
<emphasis bold="true" pageId="0" pageNumber="43">Figure 17.</emphasis>
Female of
<taxonomicName authorityName="Zamani &amp; Marusik &amp; Szűts" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera persica" order="Araneae" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="persica">
<emphasis italics="true" pageId="0" pageNumber="43">Dysdera persica</emphasis>
</taxonomicName>
sp. nov., endogyne
<emphasis bold="true" pageId="0" pageNumber="43">A</emphasis>
ventral view
<emphasis bold="true" pageId="0" pageNumber="43">B</emphasis>
dorsal view. Scale bars: 0.25 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="43" type="distribution">
<paragraph pageId="0" pageNumber="43">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="43">
Known only from listed localities in Golestan and Mazandaran provinces, northern Iran (Fig.
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Distribution records of Dysdera spp. in Iran 1 D. achaemenes sp. nov. 2 D. bakhtiari sp. nov. 3 D. hormuzensis sp. nov. 4 D. iranica sp. nov. 5 D. isfahanica sp. nov. 6 D. mazeruni sp. nov. 7 D. persica sp. nov. 8 D. pococki 9 D. sagartia sp. nov. and D. medes sp. nov. 10 D. verkana sp. nov. 11 D. xerxesi sp. nov. 12 D. damavandica sp. nov. 13 D. tapuria sp. nov. 14 D. genoensis sp. nov." figureDoi="10.3897/zookeys.1146.97517.figure35" httpUri="https://binary.pensoft.net/fig/806646" pageId="0" pageNumber="43">35</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>