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<document ID-DOI="10.1643/i2021056" approvalRequired="3" approvalRequired_for_document="1" approvalRequired_for_textStreams="1" approvalRequired_for_treatments="1" checkinTime="1678452726402" checkinUser="valdenar" docAuthor="Ng, Heok Hee &amp; Kottelat, Maurice" docDate="2022" docId="067D2B51FFD9FFDF1F68F9FB7BF4F8B8" docLanguage="en" docName="IchthyolHerpetol.110.2.262267.pdf" docOrigin="Ichthyology &amp; Herpetology 110 (2)" docSource="http://dx.doi.org/10.1643/i2021056" docTitle="Glaridoglanis andersonii AMS B." docType="treatment" docVersion="1" lastPageNumber="266" masterDocId="FA445329FFDDFFDB1C5CFFDE784AFFFA" masterDocTitle="A New Glyptosternine Catfish from Myanmar (Actinopterygii: Siluriformes: Sisoridae)" masterLastPageNumber="267" masterPageNumber="262" pageNumber="266" updateTime="1678452727301" updateUser="valdenar">
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<mods:title>A New Glyptosternine Catfish from Myanmar (Actinopterygii: Siluriformes: Sisoridae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ng, Heok Hee</mods:namePart>
<mods:affiliation>c / o Lee Kong Chian Natural History Museum, 2 Conservatory Drive, Singapore 117377</mods:affiliation>
<mods:nameIdentifier type="email">heokhee.ng@gmail.com.</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kottelat, Maurice</mods:namePart>
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<treatment LSID="urn:lsid:plazi:treatment:067D2B51FFD9FFDF1F68F9FB7BF4F8B8" httpUri="http://treatment.plazi.org/id/067D2B51FFD9FFDF1F68F9FB7BF4F8B8" lastPageNumber="266" pageId="4" pageNumber="266">
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<taxonomicName ID-CoL="3G62N" ID-ENA="278783" authority=": AMS B." authorityName="AMS B." box="[820,1094,1573,1595]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="andersonii">
<emphasis box="[820,1087,1573,1595]" italics="true" pageId="4" pageNumber="266">Glaridoglanis andersonii</emphasis>
:
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="266" type="materials_examined">
<paragraph blockId="4.[820,1490,1573,1858]" pageId="4" pageNumber="266">
<materialsCitation pageId="4" pageNumber="266">AMS B.8081, 1, 121.3 mm SL, Myanmar, Bhamo (photograph and radiograph examined);</materialsCitation>
<materialsCitation pageId="4" pageNumber="266">
CMK 26593, 2, 74.1127.6 mm SL, Myanmar, Kachin State, Imawbum Range, Harmaw Chaung, eastern part of Tsaw Law village, northern part of Satt Chaung village and Chaung Lunn village, tributary of NMai Hka, 26813
<emphasis bold="true" box="[1416,1421,1716,1731]" pageId="4" pageNumber="266">
<superScript attach="right" box="[1416,1421,1716,1731]" fontSize="6" pageId="4" pageNumber="266">0</superScript>
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15
<emphasis bold="true" box="[1454,1462,1716,1731]" pageId="4" pageNumber="266">
<superScript attach="right" box="[1454,1462,1716,1731]" fontSize="6" pageId="4" pageNumber="266">00</superScript>
</emphasis>
N, 98820
<emphasis bold="true" box="[884,889,1745,1760]" pageId="4" pageNumber="266">
<superScript attach="none" box="[884,889,1745,1760]" fontSize="6" pageId="4" pageNumber="266">0</superScript>
</emphasis>
07
<emphasis bold="true" box="[919,927,1745,1760]" pageId="4" pageNumber="266">
<superScript attach="right" box="[919,927,1745,1760]" fontSize="6" pageId="4" pageNumber="266">00</superScript>
</emphasis>
E; NRM 10721 (holotype of
<taxonomicName authorityName="Rendahl &amp; Vestergren" authorityYear="1941" box="[1239,1474,1748,1770]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="malaisei">
<emphasis box="[1239,1474,1748,1770]" italics="true" pageId="4" pageNumber="266">Glyptosternon malaisei</emphasis>
</taxonomicName>
), 70.4 mm SL;
</materialsCitation>
<materialsCitation pageId="4" pageNumber="266">
NRM 25108, 3, 98.0129.0 mm SL, Myanmar, Kachin State, Kambawti (¼Kan Paik Ti, 25824
<emphasis bold="true" box="[1416,1421,1804,1819]" pageId="4" pageNumber="266">
<superScript attach="right" box="[1416,1421,1804,1819]" fontSize="6" pageId="4" pageNumber="266">0</superScript>
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24
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<superScript attach="right" box="[1454,1462,1804,1819]" fontSize="6" pageId="4" pageNumber="266">00</superScript>
</emphasis>
N, 98807
<emphasis bold="true" box="[884,889,1833,1848]" pageId="4" pageNumber="266">
<superScript attach="none" box="[884,889,1833,1848]" fontSize="6" pageId="4" pageNumber="266">0</superScript>
</emphasis>
03
<emphasis bold="true" box="[919,927,1833,1848]" pageId="4" pageNumber="266">
<superScript attach="right" box="[919,927,1833,1848]" fontSize="6" pageId="4" pageNumber="266">00</superScript>
</emphasis>
E).
</materialsCitation>
</paragraph>
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</treatment>
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<emphasis bold="true" box="[820,1223,771,794]" pageId="0" pageNumber="262">
<taxonomicName authority="Ng &amp; Kottelat, 2022" authorityName="Ng &amp; Kottelat" authorityYear="2022" box="[820,1063,771,794]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="0" pageNumber="262" phylum="Chordata" rank="species" species="ramosa" status="new species">
<emphasis bold="true" box="[820,1063,771,794]" italics="true" pageId="0" pageNumber="262">Glaridoglanis ramosa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[1079,1223,771,794]" pageId="0" pageNumber="262" rank="species">new species</taxonomicNameLabel>
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<subSubSection pageId="0" pageNumber="262" type="description">
<paragraph blockId="0.[820,1397,771,880]" pageId="0" pageNumber="262">urn:lsid:zoobank.org:act:03C1B59A-D586-45A9-9117- 0726DE94F8F7</paragraph>
<paragraph blockId="0.[820,1397,771,880]" box="[820,908,861,880]" pageId="0" pageNumber="262">
<figureCitation box="[820,908,861,880]" captionStart="FIG" captionStartId="1.[142,170,1647,1667]" captionTargetBox="[142,1530,178,1618]" captionTargetId="figure-174@1.[142,1530,178,1618]" captionTargetPageId="1" captionText="FIG. 1. Glaridoglanis ramosa, holotype, MHNG 2786.064, 104.3 mm SL. Dorsal, lateral, and ventral views (first pectoral-fin element on left damaged)." figureDoi="http://doi.org/10.5281/zenodo.7716355" httpUri="https://zenodo.org/record/7716355/files/figure.png" pageId="0" pageNumber="262">Figure 1</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="262" type="materials_examined">
<paragraph blockId="0.[820,1490,928,1068]" pageId="0" pageNumber="262">
<materialsCitation collectionCode="MHNG" collectorName="Nyein Chan &amp; Aung Lin" location="Shan State" municipality="Myanmar" pageId="0" pageNumber="262" specimenCode="MHNG 2786.064" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[820,945,928,951]" italics="true" pageId="0" pageNumber="262">
<typeStatus box="[820,917,928,951]" pageId="0" pageNumber="262">Holotype</typeStatus>
.—
</emphasis>
<specimenCode box="[945,1138,929,951]" pageId="0" pageNumber="262">MHNG 2786.064</specimenCode>
, 104.3 mm SL,
<collectingMunicipality box="[1314,1423,929,950]" pageId="0" pageNumber="262">Myanmar</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:067D2B51FFDDFFDF1F68FCDD7CEFFA2B:8B0BCC9CFFDDFFDB19C7FC7F7B26FC29" municipality="Myanmar" name="Shan State" pageId="0" pageNumber="262">Shan State</location>
,
<location LSID="urn:lsid:plazi:treatment:067D2B51FFDDFFDF1F68FCDD7CEFFA2B:8B0BCC9CFFDDFFDB1F2BFC617C1BFC2E" box="[887,1105,958,980]" municipality="Myanmar" name="Ywangan Township" pageId="0" pageNumber="262">Ywangan Township</location>
(
<location LSID="urn:lsid:plazi:treatment:067D2B51FFDDFFDF1F68FCDD7CEFFA2B:8B0BCC9CFFDDFFDB1803FC607D3FFC2E" box="[1119,1397,958,980]" municipality="Myanmar" name="Danu Autonomous Zone" pageId="0" pageNumber="262">Danu Autonomous Zone</location>
),
<location LSID="urn:lsid:plazi:treatment:067D2B51FFDDFFDF1F68FCDD7CEFFA2B:8B0BCC9CFFDDFFDB19DBFC607BC5FC0B" municipality="Myanmar" name="Sin Lal Chaung" pageId="0" pageNumber="262">Sin Lal Chaung</location>
,
<locationDeviation pageId="0" pageNumber="262">small stream in limestone area about 14 km E of Ywangan</locationDeviation>
, 21811
<emphasis bold="true" box="[1003,1008,1013,1028]" pageId="0" pageNumber="262">0</emphasis>
23
<emphasis bold="true" box="[1038,1046,1013,1028]" pageId="0" pageNumber="262">00</emphasis>
N, 96833
<emphasis bold="true" box="[1146,1151,1013,1028]" pageId="0" pageNumber="262">0</emphasis>
18
<emphasis bold="true" box="[1181,1189,1013,1028]" pageId="0" pageNumber="262">00</emphasis>
E, about 1190 masl,
<collectorName pageId="0" pageNumber="262">Nyein Chan</collectorName>
and
<collectorName box="[939,1045,1046,1068]" pageId="0" pageNumber="262">Aung Lin</collectorName>
, 5 March 2019.
</materialsCitation>
</paragraph>
<paragraph blockId="0.[820,1490,1099,1150]" pageId="0" pageNumber="262">
<materialsCitation collectionCode="CMK" pageId="0" pageNumber="262" specimenCode="CMK 28568" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[820,954,1099,1122]" italics="true" pageId="0" pageNumber="262">
<typeStatus box="[820,927,1099,1122]" pageId="0" pageNumber="262">Paratypes</typeStatus>
.—
</emphasis>
<specimenCode box="[954,1089,1099,1121]" pageId="0" pageNumber="262">CMK 28568</specimenCode>
, 3, 36.460.6 mm SL, ZRC 62092, 1, 41.1 mm SL, data as for holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="262" type="diagnosis">
<paragraph blockId="0.[820,1490,1181,1468]" pageId="0" pageNumber="262">
<emphasis box="[820,1184,1181,1204]" italics="true" pageId="0" pageNumber="262">
<emphasis bold="true" box="[820,954,1181,1204]" italics="true" pageId="0" pageNumber="262">Diagnosis.—</emphasis>
<taxonomicName authorityName="Ng &amp; Kottelat" authorityYear="2022" box="[954,1184,1181,1203]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="0" pageNumber="262" phylum="Chordata" rank="species" species="ramosa">Glaridoglanis ramosa</taxonomicName>
</emphasis>
is distinguished from
<emphasis box="[1465,1490,1182,1203]" italics="true" pageId="0" pageNumber="262">G.</emphasis>
<taxonomicName box="[820,926,1211,1233]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="0" pageNumber="262" phylum="Chordata" rank="species" species="andersonii">
<emphasis box="[820,926,1211,1233]" italics="true" pageId="0" pageNumber="262">andersonii</emphasis>
</taxonomicName>
in having fewer vertebrae (3940 vs. 4244), more branched pectoral-fin rays (1314 vs. 810), a longer maxillary barbel (reaching beyond proximal half vs. not more than proximal third of first pectoral-fin element; 87101% HL vs. 7579), a shorter pelvic fin (10.816.3% SL vs. 18.022.4) and dorsal-to-adipose distance (7.111.7% SL vs. 17.520.0), absence (vs. presence) of a thin, pale midlateral stripe, and an adipose fin that is strongly incised (vs. without incision or with a weak incision) at the posterior extremity of its base.
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" lastPageNumber="264" pageId="0" pageNumber="262" type="description">
<paragraph blockId="0.[820,1491,1498,1872]" pageId="0" pageNumber="262">
<emphasis bold="true" box="[820,969,1498,1521]" italics="true" pageId="0" pageNumber="262">Description.—</emphasis>
Morphometric data as in
<tableCitation box="[1274,1360,1498,1520]" captionStart="Table 1" captionStartId="3.[142,190,178,198]" captionTargetBox="[142,812,217,1149]" captionTargetPageId="3" captionText="Table 1. Morphometric data for Glaridoglanis ramosa (n ¼ 5)." httpUri="http://table.plazi.org/id/DAABCACFFFDEFFD81CD2FF6C7AAFFF3C" pageId="0" pageNumber="262" tableUuid="DAABCACFFFDEFFD81CD2FF6C7AAFFF3C">Table 1</tableCitation>
. Head and body very strongly depressed, with paired fins greatly enlarged to form elongate ovoid adhesive disc with body. Rostral margin rounded in dorsal view, moderately curved in lateral view. Dorsal profile rising gently and evenly from anterior orbital margin to origin of dorsal fin, then sloping gently ventrally to end of caudal peduncle. Ventral profile flat to anal-fin base, then sloping gently dorsally to end of caudal peduncle. Anus and urogenital openings located just before posterior margin of adpressed pelvic fin.
</paragraph>
<paragraph blockId="0.[820,1491,1498,1872]" pageId="0" pageNumber="262">Head broad, approximately as wide as long. Mouth inferior, with gape width approximately half that of head and densely papillate lips. Premaxillary tooth patch with</paragraph>
<paragraph pageId="0" pageNumber="262">
<superScript attach="left" box="[103,112,1895,1907]" fontSize="5" pageId="0" pageNumber="262">1</superScript>
<docAuthorAffiliation box="[119,974,1898,1918]" pageId="0" pageNumber="262">c/o Lee Kong Chian Natural History Museum, 2 Conservatory Drive, Singapore 117377</docAuthorAffiliation>
; Email: (HHN)
<docAuthorEmail box="[1124,1364,1898,1918]" pageId="0" pageNumber="262">heokhee.ng@gmail.com.</docAuthorEmail>
Send reprint requests to HHN.
<superScript attach="left" box="[103,112,1948,1960]" fontSize="5" pageId="0" pageNumber="262">2</superScript>
<docAuthorAffiliation box="[119,616,1952,1972]" pageId="0" pageNumber="262">Rue des Rauraques 6, 2800 Delémont, Switzerland</docAuthorAffiliation>
; Email:
<docAuthorEmail box="[695,916,1952,1971]" pageId="0" pageNumber="262">mkottelat@dplanet.ch.</docAuthorEmail>
Submitted: 13 May 2021. Accepted: 6 October 2021. Associate Editor: R. E. Reis. 2022 by the American Society of Ichthyologists and Herpetologists DOI:
<docID-DOI box="[854,1033,2003,2022]" pageId="0" pageNumber="262">10.1643/i2021056</docID-DOI>
Published online: 10 May 2022
</paragraph>
<paragraph blockId="0.[27,938,2060,2097]" box="[27,765,2060,2078]" pageId="0" pageNumber="262">Downloaded From: https://bioone.org/journals/Ichthyology-&amp;-Herpetology on 08 Mar 2023</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7716355" ID-Zenodo-Dep="7716355" box="[142,1530,1647,1667]" httpUri="https://zenodo.org/record/7716355/files/figure.png" pageId="1" pageNumber="263" startId="1.[142,170,1647,1667]" targetBox="[142,1530,178,1618]" targetPageId="1">
<paragraph blockId="1.[142,194,1647,1667]" box="[142,194,1647,1667]" pageId="1" pageNumber="263">
<heading allCaps="true" bold="true" box="[142,194,1647,1667]" fontSize="8" level="1" pageId="1" pageNumber="263" reason="2">
<emphasis bold="true" box="[142,194,1647,1667]" pageId="1" pageNumber="263">FIG. 1.</emphasis>
</heading>
</paragraph>
<paragraph blockId="1.[216,1530,1647,1667]" box="[216,1530,1647,1667]" pageId="1" pageNumber="263">
<taxonomicName authorityName="Ng &amp; Kottelat" authorityYear="2022" box="[216,416,1647,1667]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="263" phylum="Chordata" rank="species" species="ramosa">
<emphasis box="[216,416,1647,1667]" italics="true" pageId="1" pageNumber="263">Glaridoglanis ramosa</emphasis>
</taxonomicName>
,
<emphasis bold="true" box="[426,1530,1647,1667]" pageId="1" pageNumber="263">holotype, MHNG 2786.064, 104.3 mm SL. Dorsal, lateral, and ventral views (first pectoral-fin element on left damaged).</emphasis>
</paragraph>
</caption>
<paragraph blockId="1.[142,812,1711,2024]" pageId="1" pageNumber="263">anterior margin exposed when mouth is closed. Lower lip with prominent labial fold joined to base of maxillary barbel. Postlabial groove on lower jaw interrupted, ending at base of inner mandibular barbel. Jaw teeth embedded in skin, distally flattened, and oar-shaped. Teeth in single crescentic band on upper jaw and in two well-separated, roughly triangular patches on lower jaw. Palate edentulous. Eye small, dorsolaterally situated, subcutaneous. Gill opening narrow, extending from base of first pectoral-fin ray to level immediately dorsal to base of posteriormost pectoral-fin ray.</paragraph>
<paragraph blockId="1.[859,1529,1711,2024]" pageId="1" pageNumber="263">Barbels in four pairs. Nasal barbel long, reaching beyond posterior orbital margin. Maxillary barbel flattened, with flap of skin fringing posterior margin of distal half; its tip pointed; ventral surface with numerous striae; extending beyond base of first pectoral-fin ray. Inner mandibular barbel slightly flattened, short. Outer mandibular barbel situated lateral to inner mandibular barbel; slightly flattened, reaching to base of first pectoral-fin ray.</paragraph>
<paragraph blockId="1.[859,1529,1711,2024]" pageId="1" pageNumber="263">Dorsal fin without spine, with i,5 (5) rays; surfaces of fin rays covered with small tubercles. Adipose fin with long base;</paragraph>
<paragraph blockId="1.[27,938,2060,2097]" box="[27,765,2060,2078]" pageId="1" pageNumber="263">Downloaded From: https://bioone.org/journals/Ichthyology-&amp;-Herpetology on 08 Mar 2023</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7716357" ID-Zenodo-Dep="7716357" httpUri="https://zenodo.org/record/7716357/files/figure.png" pageId="2" pageNumber="264" startId="2.[1139,1167,1450,1470]" targetBox="[103,1089,181,1518]" targetPageId="2">
<paragraph blockId="2.[1139,1490,1450,1523]" pageId="2" pageNumber="264">
<emphasis bold="true" pageId="2" pageNumber="264">FIG. 2. Irrawaddy River basin showing type locality of</emphasis>
<taxonomicName authorityName="Ng &amp; Kottelat" authorityYear="2022" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="264" phylum="Chordata" rank="species" species="ramosa">
<emphasis italics="true" pageId="2" pageNumber="264">Glaridoglanis ramosa</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="2.[103,773,1562,2024]" pageId="2" pageNumber="264">anterior extremity at approximately midway between bases of pelvic and anal fins, posterior extremity separate from upper procurrent caudal-fin rays and distinctly incised. Anal fin with ii,4 (5) rays, surfaces of fin rays covered with small tubercles. Caudal fin truncate or gently rounded, with i,7,6,i* (1) or i,7,7,i (4) rays; surfaces of fin rays covered with small tubercles. Pelvic fin greatly enlarged, with convex distal margin and i,5 (5) rays; first ray greatly flattened, with numerous striae on ventral surface; dorsal surfaces of fin rays covered with small tubercles. Pectoral fin greatly enlarged, without spine and with i,13,i (1) or i,14,i* (4) rays; first ray greatly flattened, with numerous striae on ventral surface. Dorsal surface of pectoral-fin rays covered with small tubercles. Vertebrae 26þ13¼39* (1), 27þ12¼39 (1), 27þ13¼40 (2), or 29þ11¼40 (1).</paragraph>
<paragraph blockId="2.[820,1490,1561,1762]" pageId="2" pageNumber="264">
<emphasis bold="true" box="[820,963,1561,1584]" italics="true" pageId="2" pageNumber="264">Coloration.—</emphasis>
In 70% ethanol: medium brown on dorsal and lateral surfaces of head and dorsal surface of body, dark yellow on ventral region. Dorsal, anal, and caudal fins brown; dorsal surfaces of pectoral and pelvic fins brown, with ventral surfaces of pectoral and pelvic fins dark yellow. Dorsal surface of nasal and maxillary barbels brown, ventral surface dark yellow. Mandibular barbels dark yellow.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="265" pageId="2" pageNumber="264" type="distribution">
<paragraph blockId="2.[820,1491,1793,2024]" pageId="2" pageNumber="264">
<emphasis box="[820,1325,1793,1816]" italics="true" pageId="2" pageNumber="264">
<emphasis bold="true" box="[820,1106,1793,1816]" italics="true" pageId="2" pageNumber="264">Distribution and habitat.—</emphasis>
<taxonomicName authorityName="Ng &amp; Kottelat" authorityYear="2022" box="[1106,1325,1793,1815]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="264" phylum="Chordata" rank="species" species="ramosa">Glaridoglanis ramosa</taxonomicName>
</emphasis>
is known from a single locality in a cryptorheic basin in Shan State in Myanmar (
<figureCitation box="[952,1025,1853,1876]" captionStart="FIG" captionStartId="2.[1139,1167,1450,1470]" captionTargetBox="[103,1089,181,1518]" captionTargetId="graphics-314@2.[179,1067,211,1508]" captionTargetPageId="2" captionText="FIG. 2. Irrawaddy River basin showing type locality of Glaridoglanis ramosa." figureDoi="http://doi.org/10.5281/zenodo.7716357" httpUri="https://zenodo.org/record/7716357/files/figure.png" pageId="2" pageNumber="264">Fig. 2</figureCitation>
). It was collected in Sin Lal Chaung (21811
<emphasis bold="true" box="[894,899,1879,1894]" pageId="2" pageNumber="264">
<superScript attach="none" box="[894,899,1879,1894]" fontSize="6" pageId="2" pageNumber="264">0</superScript>
</emphasis>
23
<emphasis bold="true" box="[930,938,1879,1894]" pageId="2" pageNumber="264">
<superScript attach="right" box="[930,938,1879,1894]" fontSize="6" pageId="2" pageNumber="264">00</superScript>
</emphasis>
N, 96833
<emphasis bold="true" box="[1047,1052,1879,1894]" pageId="2" pageNumber="264">
<superScript attach="right" box="[1047,1052,1879,1894]" fontSize="6" pageId="2" pageNumber="264">0</superScript>
</emphasis>
18
<emphasis bold="true" box="[1082,1090,1879,1894]" pageId="2" pageNumber="264">
<superScript attach="right" box="[1082,1090,1879,1894]" fontSize="6" pageId="2" pageNumber="264">00</superScript>
</emphasis>
E;
<figureCitation box="[1129,1218,1883,1906]" captionStart="FIG" captionStartId="3.[859,887,1221,1241]" captionTargetBox="[859,1530,178,1192]" captionTargetId="figure-690@3.[859,1530,178,1192]" captionTargetPageId="3" captionText="FIG. 3. Type locality of Glaridoglanis ramosa: (A) Sin Lal Chaung, about 300 m upstream of cave entrance; (B) Sin Lal Chaung at entrance of Yae Win cave. Photographs courtesy of Nyein Chan." figureDoi="http://doi.org/10.5281/zenodo.7716361" httpUri="https://zenodo.org/record/7716361/files/figure.png" pageId="2" pageNumber="264">Fig. 3A</figureCitation>
), a small stream that disappears in Yae Win cave (21811
<emphasis bold="true" box="[1227,1232,1909,1924]" pageId="2" pageNumber="264">
<superScript attach="right" box="[1227,1232,1909,1924]" fontSize="6" pageId="2" pageNumber="264">0</superScript>
</emphasis>
27
<emphasis bold="true" box="[1262,1270,1909,1924]" pageId="2" pageNumber="264">
<superScript attach="right" box="[1262,1270,1909,1924]" fontSize="6" pageId="2" pageNumber="264">00</superScript>
</emphasis>
N, 96833
<emphasis bold="true" box="[1373,1378,1909,1924]" pageId="2" pageNumber="264">
<superScript attach="none" box="[1373,1378,1909,1924]" fontSize="6" pageId="2" pageNumber="264">0</superScript>
</emphasis>
12
<emphasis bold="true" box="[1409,1417,1909,1924]" pageId="2" pageNumber="264">
<superScript attach="right" box="[1409,1417,1909,1924]" fontSize="6" pageId="2" pageNumber="264">00</superScript>
</emphasis>
E;
<figureCitation captionStart="FIG" captionStartId="3.[859,887,1221,1241]" captionTargetBox="[859,1530,178,1192]" captionTargetId="figure-690@3.[859,1530,178,1192]" captionTargetPageId="3" captionText="FIG. 3. Type locality of Glaridoglanis ramosa: (A) Sin Lal Chaung, about 300 m upstream of cave entrance; (B) Sin Lal Chaung at entrance of Yae Win cave. Photographs courtesy of Nyein Chan." figureDoi="http://doi.org/10.5281/zenodo.7716361" httpUri="https://zenodo.org/record/7716361/files/figure.png" pageId="2" pageNumber="264">Fig. 3B</figureCitation>
). The outflow of the Sin Lal Chaung from the cave is not known, although it is locally believed that it flows to Zawgyi Chaung, but no precise area could be suggested. The Zawgyi
</paragraph>
<paragraph blockId="2.[27,938,2060,2097]" pageId="2" pageNumber="264">Downloaded From: https://bioone.org/journals/Ichthyology-&amp;-Herpetology on 08 Mar 2023 Terms of Use: https://bioone.org/terms-of-use Access provided by American Museum of Natural History Library</paragraph>
<caption ID-Table-UUID="DAABCACFFFDEFFD81CD2FF6C7AAFFF3C" box="[142,741,178,198]" httpUri="http://table.plazi.org/id/DAABCACFFFDEFFD81CD2FF6C7AAFFF3C" pageId="3" pageNumber="265" startId="3.[142,190,178,198]" targetBox="[142,812,217,1149]" targetIsTable="true" targetPageId="3">
<paragraph blockId="3.[142,741,178,198]" box="[142,741,178,198]" pageId="3" pageNumber="265">
<tableNote box="[142,213,178,198]" pageId="3" pageNumber="265" targetBox="[142,812,217,1149]" targetPageId="3">Table 1.</tableNote>
<tableNote box="[234,741,178,198]" pageId="3" pageNumber="265" targetBox="[142,812,217,1149]" targetPageId="3">
<emphasis bold="true" box="[234,449,178,198]" pageId="3" pageNumber="265">Morphometric data for</emphasis>
<taxonomicName authorityName="Ng &amp; Kottelat" authorityYear="2022" box="[456,661,178,198]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="265" phylum="Chordata" rank="species" species="ramosa">
<emphasis box="[456,661,178,198]" italics="true" pageId="3" pageNumber="265">Glaridoglanis ramosa</emphasis>
</taxonomicName>
<emphasis bold="true" box="[668,676,178,198]" pageId="3" pageNumber="265">(</emphasis>
<emphasis box="[677,689,178,198]" italics="true" pageId="3" pageNumber="265">n</emphasis>
¼ 5).
</tableNote>
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="265">
<table box="[142,812,217,1149]" gridcols="4" gridrows="32" pageId="3" pageNumber="265">
<tr box="[142,812,217,240]" gridrow="0" pageId="3" pageNumber="265" rowspan-0="1">
<th box="[449,545,217,240]" gridcol="1" gridrow="0" pageId="3" pageNumber="265">
<emphasis bold="true" box="[449,545,217,240]" pageId="3" pageNumber="265">Holotype</emphasis>
</th>
<th box="[561,682,217,240]" gridcol="2" gridrow="0" pageId="3" pageNumber="265">
<emphasis bold="true" box="[588,654,217,240]" pageId="3" pageNumber="265">Range</emphasis>
</th>
<th box="[698,812,217,240]" gridcol="3" gridrow="0" pageId="3" pageNumber="265">
<emphasis bold="true" box="[700,760,217,240]" pageId="3" pageNumber="265">Mean</emphasis>
6
<emphasis bold="true" box="[780,810,217,240]" pageId="3" pageNumber="265">SD</emphasis>
</th>
</tr>
<tr box="[142,812,269,288]" gridrow="1" pageId="3" pageNumber="265" rowspan-3="1">
<th box="[142,433,269,288]" gridcol="0" gridrow="1" pageId="3" pageNumber="265">Standard length (mm)</th>
<td box="[449,545,269,288]" gridcol="1" gridrow="1" pageId="3" pageNumber="265">104.3</td>
<td box="[561,682,269,288]" gridcol="2" gridrow="1" pageId="3" pageNumber="265">36.1104.3</td>
</tr>
<tr box="[142,812,295,318]" gridrow="2" pageId="3" pageNumber="265" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th box="[142,433,295,318]" gridcol="0" gridrow="2" pageId="3" pageNumber="265">
<emphasis bold="true" box="[142,427,295,318]" pageId="3" pageNumber="265">Percent of standard length</emphasis>
</th>
</tr>
<tr box="[142,812,327,346]" gridrow="3" pageId="3" pageNumber="265">
<th box="[142,433,327,346]" gridcol="0" gridrow="3" pageId="3" pageNumber="265">Predorsal length</th>
<td box="[449,545,327,346]" gridcol="1" gridrow="3" pageId="3" pageNumber="265">30.2</td>
<td box="[561,682,327,346]" gridcol="2" gridrow="3" pageId="3" pageNumber="265">30.234.8</td>
<td box="[698,812,327,346]" gridcol="3" gridrow="3" pageId="3" pageNumber="265">32.962.42</td>
</tr>
<tr box="[142,812,355,374]" gridrow="4" pageId="3" pageNumber="265">
<th box="[142,433,355,374]" gridcol="0" gridrow="4" pageId="3" pageNumber="265">Preanal length</th>
<td box="[449,545,355,374]" gridcol="1" gridrow="4" pageId="3" pageNumber="265">72.0</td>
<td box="[561,682,355,374]" gridcol="2" gridrow="4" pageId="3" pageNumber="265">70.074.0</td>
<td box="[698,812,355,374]" gridcol="3" gridrow="4" pageId="3" pageNumber="265">73.361.13</td>
</tr>
<tr box="[142,812,384,403]" gridrow="5" pageId="3" pageNumber="265">
<th box="[142,433,384,403]" gridcol="0" gridrow="5" pageId="3" pageNumber="265">Prepelvic length</th>
<td box="[449,545,384,403]" gridcol="1" gridrow="5" pageId="3" pageNumber="265">37.9</td>
<td box="[561,682,384,403]" gridcol="2" gridrow="5" pageId="3" pageNumber="265">37.943.1</td>
<td box="[698,812,384,403]" gridcol="3" gridrow="5" pageId="3" pageNumber="265">40.562.60</td>
</tr>
<tr box="[142,812,413,432]" gridrow="6" pageId="3" pageNumber="265">
<th box="[142,433,413,432]" gridcol="0" gridrow="6" pageId="3" pageNumber="265">Prepectoral length</th>
<td box="[449,545,413,432]" gridcol="1" gridrow="6" pageId="3" pageNumber="265">13.8</td>
<td box="[561,682,413,432]" gridcol="2" gridrow="6" pageId="3" pageNumber="265">13.816.1</td>
<td box="[698,812,413,432]" gridcol="3" gridrow="6" pageId="3" pageNumber="265">15.061.16</td>
</tr>
<tr box="[142,812,441,461]" gridrow="7" pageId="3" pageNumber="265">
<th box="[142,433,441,461]" gridcol="0" gridrow="7" pageId="3" pageNumber="265">Length of dorsal-fin base</th>
<td box="[449,545,441,461]" gridcol="1" gridrow="7" pageId="3" pageNumber="265">10.0</td>
<td box="[561,682,441,461]" gridcol="2" gridrow="7" pageId="3" pageNumber="265">10.012.7</td>
<td box="[698,812,441,461]" gridcol="3" gridrow="7" pageId="3" pageNumber="265">11.561.39</td>
</tr>
<tr box="[142,812,470,489]" gridrow="8" pageId="3" pageNumber="265">
<th box="[142,433,470,489]" gridcol="0" gridrow="8" pageId="3" pageNumber="265">Length of anal-fin base</th>
<td box="[449,545,470,489]" gridcol="1" gridrow="8" pageId="3" pageNumber="265">7.3</td>
<td box="[561,682,470,489]" gridcol="2" gridrow="8" pageId="3" pageNumber="265">7.39.2</td>
<td box="[698,812,470,489]" gridcol="3" gridrow="8" pageId="3" pageNumber="265">8.360.95</td>
</tr>
<tr box="[142,812,498,518]" gridrow="9" pageId="3" pageNumber="265">
<th box="[142,433,498,518]" gridcol="0" gridrow="9" pageId="3" pageNumber="265">Pelvic-fin length</th>
<td box="[449,545,498,518]" gridcol="1" gridrow="9" pageId="3" pageNumber="265">10.8</td>
<td box="[561,682,498,518]" gridcol="2" gridrow="9" pageId="3" pageNumber="265">10.816.3</td>
<td box="[698,812,498,518]" gridcol="3" gridrow="9" pageId="3" pageNumber="265">13.362.78</td>
</tr>
<tr box="[142,812,527,547]" gridrow="10" pageId="3" pageNumber="265">
<th box="[142,433,527,547]" gridcol="0" gridrow="10" pageId="3" pageNumber="265">Pectoral-fin length</th>
<td box="[449,545,527,547]" gridcol="1" gridrow="10" pageId="3" pageNumber="265">20.1</td>
<td box="[561,682,527,547]" gridcol="2" gridrow="10" pageId="3" pageNumber="265">20.122.4</td>
<td box="[698,812,527,547]" gridcol="3" gridrow="10" pageId="3" pageNumber="265">21.561.21</td>
</tr>
<tr box="[142,812,556,575]" gridrow="11" pageId="3" pageNumber="265">
<th box="[142,433,556,575]" gridcol="0" gridrow="11" pageId="3" pageNumber="265">Caudal-fin length</th>
<td box="[449,545,556,575]" gridcol="1" gridrow="11" pageId="3" pageNumber="265">13.6</td>
<td box="[561,682,556,575]" gridcol="2" gridrow="11" pageId="3" pageNumber="265">13.616.3</td>
<td box="[698,812,556,575]" gridcol="3" gridrow="11" pageId="3" pageNumber="265">14.561.53</td>
</tr>
<tr box="[142,812,585,604]" gridrow="12" pageId="3" pageNumber="265">
<th box="[142,433,585,604]" gridcol="0" gridrow="12" pageId="3" pageNumber="265">Length of adipose-fin base</th>
<td box="[449,545,585,604]" gridcol="1" gridrow="12" pageId="3" pageNumber="265">40.2</td>
<td box="[561,682,585,604]" gridcol="2" gridrow="12" pageId="3" pageNumber="265">38.440.2</td>
<td box="[698,812,585,604]" gridcol="3" gridrow="12" pageId="3" pageNumber="265">39.661.01</td>
</tr>
<tr box="[142,812,613,633]" gridrow="13" pageId="3" pageNumber="265">
<th box="[142,433,613,633]" gridcol="0" gridrow="13" pageId="3" pageNumber="265">Dorsal-to-adipose distance</th>
<td box="[449,545,613,633]" gridcol="1" gridrow="13" pageId="3" pageNumber="265">11.7</td>
<td box="[561,682,613,633]" gridcol="2" gridrow="13" pageId="3" pageNumber="265">7.111.7</td>
<td box="[698,812,613,633]" gridcol="3" gridrow="13" pageId="3" pageNumber="265">9.062.30</td>
</tr>
<tr box="[142,812,642,661]" gridrow="14" pageId="3" pageNumber="265">
<th box="[142,433,642,661]" gridcol="0" gridrow="14" pageId="3" pageNumber="265">Post-adipose distance</th>
<td box="[449,545,642,661]" gridcol="1" gridrow="14" pageId="3" pageNumber="265">11.0</td>
<td box="[561,682,642,661]" gridcol="2" gridrow="14" pageId="3" pageNumber="265">9.911.0</td>
<td box="[698,812,642,661]" gridcol="3" gridrow="14" pageId="3" pageNumber="265">10.460.57</td>
</tr>
<tr box="[142,812,671,690]" gridrow="15" pageId="3" pageNumber="265">
<th box="[142,433,671,690]" gridcol="0" gridrow="15" pageId="3" pageNumber="265">Length of caudal peduncle</th>
<td box="[449,545,671,690]" gridcol="1" gridrow="15" pageId="3" pageNumber="265">17.4</td>
<td box="[561,682,671,690]" gridcol="2" gridrow="15" pageId="3" pageNumber="265">16.017.4</td>
<td box="[698,812,671,690]" gridcol="3" gridrow="15" pageId="3" pageNumber="265">16.760.70</td>
</tr>
<tr box="[142,812,700,719]" gridrow="16" pageId="3" pageNumber="265">
<th box="[142,433,700,719]" gridcol="0" gridrow="16" pageId="3" pageNumber="265">Depth of caudal peduncle</th>
<td box="[449,545,700,719]" gridcol="1" gridrow="16" pageId="3" pageNumber="265">7.9</td>
<td box="[561,682,700,719]" gridcol="2" gridrow="16" pageId="3" pageNumber="265">7.98.9</td>
<td box="[698,812,700,719]" gridcol="3" gridrow="16" pageId="3" pageNumber="265">8.460.50</td>
</tr>
<tr box="[142,812,728,748]" gridrow="17" pageId="3" pageNumber="265">
<th box="[142,433,728,748]" gridcol="0" gridrow="17" pageId="3" pageNumber="265">Body depth at anus</th>
<td box="[449,545,728,748]" gridcol="1" gridrow="17" pageId="3" pageNumber="265">9.9</td>
<td box="[561,682,728,748]" gridcol="2" gridrow="17" pageId="3" pageNumber="265">9.913.1</td>
<td box="[698,812,728,748]" gridcol="3" gridrow="17" pageId="3" pageNumber="265">11.561.60</td>
</tr>
<tr box="[142,812,757,776]" gridrow="18" pageId="3" pageNumber="265">
<th box="[142,433,757,776]" gridcol="0" gridrow="18" pageId="3" pageNumber="265">Pectoralpelvic distance</th>
<td box="[449,545,757,776]" gridcol="1" gridrow="18" pageId="3" pageNumber="265">26.7</td>
<td box="[561,682,757,776]" gridcol="2" gridrow="18" pageId="3" pageNumber="265">26.727.0</td>
<td box="[698,812,757,776]" gridcol="3" gridrow="18" pageId="3" pageNumber="265">26.860.17</td>
</tr>
<tr box="[142,812,786,805]" gridrow="19" pageId="3" pageNumber="265">
<th box="[142,433,786,805]" gridcol="0" gridrow="19" pageId="3" pageNumber="265">Head length</th>
<td box="[449,545,786,805]" gridcol="1" gridrow="19" pageId="3" pageNumber="265">21.0</td>
<td box="[561,682,786,805]" gridcol="2" gridrow="19" pageId="3" pageNumber="265">21.025.6</td>
<td box="[698,812,786,805]" gridcol="3" gridrow="19" pageId="3" pageNumber="265">23.562.33</td>
</tr>
<tr box="[142,812,814,834]" gridrow="20" pageId="3" pageNumber="265">
<th box="[142,433,814,834]" gridcol="0" gridrow="20" pageId="3" pageNumber="265">Head width</th>
<td box="[449,545,814,834]" gridcol="1" gridrow="20" pageId="3" pageNumber="265">18.2</td>
<td box="[561,682,814,834]" gridcol="2" gridrow="20" pageId="3" pageNumber="265">18.219.8</td>
<td box="[698,812,814,834]" gridcol="3" gridrow="20" pageId="3" pageNumber="265">19.160.81</td>
</tr>
<tr box="[142,812,843,862]" gridrow="21" pageId="3" pageNumber="265">
<th box="[142,433,843,862]" gridcol="0" gridrow="21" pageId="3" pageNumber="265">Head depth</th>
<td box="[449,545,843,862]" gridcol="1" gridrow="21" pageId="3" pageNumber="265">8.5</td>
<td box="[561,682,843,862]" gridcol="2" gridrow="21" pageId="3" pageNumber="265">8.510.9</td>
<td box="[698,812,843,862]" gridcol="3" gridrow="21" pageId="3" pageNumber="265">9.761.20</td>
</tr>
<tr box="[142,812,869,892]" gridrow="22" pageId="3" pageNumber="265" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th box="[142,433,869,892]" gridcol="0" gridrow="22" pageId="3" pageNumber="265">
<emphasis bold="true" box="[142,387,869,892]" pageId="3" pageNumber="265">Percent of head length</emphasis>
</th>
</tr>
<tr box="[142,812,900,920]" gridrow="23" pageId="3" pageNumber="265">
<th box="[142,433,900,920]" gridcol="0" gridrow="23" pageId="3" pageNumber="265">Snout length</th>
<td box="[449,545,900,920]" gridcol="1" gridrow="23" pageId="3" pageNumber="265">48</td>
<td box="[561,682,900,920]" gridcol="2" gridrow="23" pageId="3" pageNumber="265">4857</td>
<td box="[698,812,900,920]" gridcol="3" gridrow="23" pageId="3" pageNumber="265">5165.2</td>
</tr>
<tr box="[142,812,929,948]" gridrow="24" pageId="3" pageNumber="265">
<th box="[142,433,929,948]" gridcol="0" gridrow="24" pageId="3" pageNumber="265">Interorbital distance</th>
<td box="[449,545,929,948]" gridcol="1" gridrow="24" pageId="3" pageNumber="265">23</td>
<td box="[561,682,929,948]" gridcol="2" gridrow="24" pageId="3" pageNumber="265">2331</td>
<td box="[698,812,929,948]" gridcol="3" gridrow="24" pageId="3" pageNumber="265">2864.2</td>
</tr>
<tr box="[142,812,958,977]" gridrow="25" pageId="3" pageNumber="265">
<th box="[142,433,958,977]" gridcol="0" gridrow="25" pageId="3" pageNumber="265">Eye diameter</th>
<td box="[449,545,958,977]" gridcol="1" gridrow="25" pageId="3" pageNumber="265">7</td>
<td box="[561,682,958,977]" gridcol="2" gridrow="25" pageId="3" pageNumber="265">711</td>
<td box="[698,812,958,977]" gridcol="3" gridrow="25" pageId="3" pageNumber="265">962.1</td>
</tr>
<tr box="[142,812,986,1006]" gridrow="26" pageId="3" pageNumber="265">
<th box="[142,433,986,1006]" gridcol="0" gridrow="26" pageId="3" pageNumber="265">Nasal barbel length</th>
<td box="[449,545,986,1006]" gridcol="1" gridrow="26" pageId="3" pageNumber="265">52</td>
<td box="[561,682,986,1006]" gridcol="2" gridrow="26" pageId="3" pageNumber="265">3652</td>
<td box="[698,812,986,1006]" gridcol="3" gridrow="26" pageId="3" pageNumber="265">4568.3</td>
</tr>
<tr box="[142,812,1015,1035]" gridrow="27" pageId="3" pageNumber="265">
<th box="[142,433,1015,1035]" gridcol="0" gridrow="27" pageId="3" pageNumber="265">Maxillary barbel length</th>
<td box="[449,545,1015,1035]" gridcol="1" gridrow="27" pageId="3" pageNumber="265">101</td>
<td box="[561,682,1015,1035]" gridcol="2" gridrow="27" pageId="3" pageNumber="265">87101</td>
<td box="[698,812,1015,1035]" gridcol="3" gridrow="27" pageId="3" pageNumber="265">9367.1</td>
</tr>
<tr box="[142,812,1044,1063]" gridrow="28" pageId="3" pageNumber="265">
<th box="[142,433,1044,1063]" gridcol="0" gridrow="28" pageId="3" pageNumber="265">Inner mandibular barbel</th>
<td box="[449,545,1044,1063]" gridcol="1" gridrow="28" pageId="3" pageNumber="265">26</td>
<td box="[561,682,1044,1063]" gridcol="2" gridrow="28" pageId="3" pageNumber="265">2126</td>
<td box="[698,812,1044,1063]" gridcol="3" gridrow="28" pageId="3" pageNumber="265">2362.5</td>
</tr>
<tr box="[142,812,1073,1092]" gridrow="29" pageId="3" pageNumber="265" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th box="[142,433,1073,1092]" gridcol="0" gridrow="29" pageId="3" pageNumber="265">length</th>
</tr>
<tr box="[142,812,1101,1121]" gridrow="30" pageId="3" pageNumber="265">
<th box="[142,433,1101,1121]" gridcol="0" gridrow="30" pageId="3" pageNumber="265">Outer mandibular barbel</th>
<td box="[449,545,1101,1121]" gridcol="1" gridrow="30" pageId="3" pageNumber="265">49</td>
<td box="[561,682,1101,1121]" gridcol="2" gridrow="30" pageId="3" pageNumber="265">3249</td>
<td box="[698,812,1101,1121]" gridcol="3" gridrow="30" pageId="3" pageNumber="265">4068.5</td>
</tr>
<tr box="[142,812,1130,1149]" gridrow="31" pageId="3" pageNumber="265" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th box="[142,433,1130,1149]" gridcol="0" gridrow="31" pageId="3" pageNumber="265">length</th>
</tr>
</table>
</paragraph>
<paragraph blockId="3.[142,812,1213,1563]" pageId="3" pageNumber="265">Chaung occupies the watershed north of the karst massive; it is a tributary of Myitnge Chaung, which enters the Ayeyarwaddy (Irrawaddy) River near Mandalay.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7716361" ID-Zenodo-Dep="7716361" httpUri="https://zenodo.org/record/7716361/files/figure.png" pageId="3" pageNumber="265" startId="3.[859,887,1221,1241]" targetBox="[859,1530,178,1192]" targetPageId="3">
<paragraph blockId="3.[859,1529,1221,1294]" pageId="3" pageNumber="265">
<emphasis bold="true" box="[859,1071,1221,1241]" pageId="3" pageNumber="265">FIG. 3. Type locality of</emphasis>
<taxonomicName authorityName="Ng &amp; Kottelat" authorityYear="2022" box="[1076,1280,1221,1241]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="265" phylum="Chordata" rank="species" species="ramosa">
<emphasis box="[1076,1280,1221,1241]" italics="true" pageId="3" pageNumber="265">Glaridoglanis ramosa</emphasis>
</taxonomicName>
: (A) Sin Lal Chaung, about 300 m upstream of cave entrance; (B) Sin Lal Chaung at entrance of Yae Win cave. Photographs courtesy of Nyein Chan.
</paragraph>
</caption>
<paragraph blockId="3.[142,812,1213,1563]" pageId="3" pageNumber="265">
At the collecting site and time (dry season), the Sin Lal Chaung was a clear water stream, about 5 m wide and 0.31.0 m deep, flowing over a substrate of sand, gravel, and stones, and occasional stretches with quieter water and silty sand. The vegetation of the surrounding area comprises scattered forest and home gardens. The fish were observed upstream of the entrance of Yae Win cave and inside the cave including in total darkness.
<taxonomicName baseAuthorityName="Blyth" baseAuthorityYear="1860" box="[314,630,1511,1533]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalichthys" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="265" phylum="Chordata" rank="species" species="berdmorei">
<emphasis box="[314,630,1511,1533]" italics="true" pageId="3" pageNumber="265">Lepidocephalichthys berdmorei</emphasis>
</taxonomicName>
and
<taxonomicName box="[692,812,1512,1533]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="265" phylum="Chordata" rank="species" species="undetermined">
<emphasis box="[692,771,1512,1533]" italics="true" pageId="3" pageNumber="265">Devario</emphasis>
sp.
</taxonomicName>
were the only other fish species collected at this locality.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="265" type="etymology">
<paragraph blockId="3.[142,812,1595,1767]" pageId="3" pageNumber="265">
<emphasis bold="true" box="[142,285,1595,1618]" italics="true" pageId="3" pageNumber="265">Etymology.—</emphasis>
The specific epithet comes from the Latin adjective
<emphasis box="[258,345,1626,1647]" italics="true" pageId="3" pageNumber="265">ramosus</emphasis>
,
<emphasis box="[369,390,1626,1647]" italics="true" pageId="3" pageNumber="265">-a</emphasis>
,
<emphasis box="[413,455,1626,1647]" italics="true" pageId="3" pageNumber="265">-um</emphasis>
, which means having many branches. This is used in allusion to the higher number of branched pectoral-fin rays in this species compared to its sole congener (
<taxonomicName box="[265,408,1714,1736]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="265" phylum="Chordata" rank="species" species="andersonii">
<emphasis box="[265,408,1714,1736]" italics="true" pageId="3" pageNumber="265">G. andersonii</emphasis>
</taxonomicName>
). As
<taxonomicName authorityName="Norman" authorityYear="1925" box="[474,612,1714,1736]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="265" phylum="Chordata" rank="genus">
<emphasis box="[474,612,1714,1736]" italics="true" pageId="3" pageNumber="265">Glaridoglanis</emphasis>
</taxonomicName>
is feminine (
<emphasis box="[774,811,1714,1736]" italics="true" pageId="3" pageNumber="265">fide</emphasis>
<bibRefCitation author="Kottelat, M." box="[142,302,1744,1767]" pageId="3" pageNumber="265" pagination="1 - 663" refId="ref4097" refString="Kottelat, M. 2013. The fishes of the inland waters of Southeast Asia: a catalogue and core bibiography of the fishes known to occur in freshwaters, mangroves and estuaries. Raffles Bulletin of Zoology Supplement 27: 1 - 663." type="journal article" year="2013">Kottelat, 2013</bibRefCitation>
), the specific epithet is declined accordingly.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="266" pageId="3" pageNumber="265" type="discussion">
<paragraph blockId="3.[142,285,1806,1829]" box="[142,285,1806,1829]" pageId="3" pageNumber="265">
<heading bold="true" box="[142,285,1806,1829]" fontSize="9" level="2" pageId="3" pageNumber="265" reason="2">
<emphasis bold="true" box="[142,285,1806,1829]" pageId="3" pageNumber="265">DISCUSSION</emphasis>
</heading>
</paragraph>
<paragraph blockId="3.[142,812,1852,2024]" lastBlockId="3.[859,1529,1339,2024]" pageId="3" pageNumber="265">
<bibRefCitation author="Day, F." box="[142,269,1852,1875]" pageId="3" pageNumber="265" pagination="511 - 527" refId="ref3859" refString="Day, F. 1870. Remarks on some of the fishes in the Calcutta Museum. - Part I. Proceedings of the Zoological Society of London 1869: 511 - 527." type="journal article" year="1870">Day (1870)</bibRefCitation>
described
<taxonomicName authorityName="Day" authorityYear="1870" box="[396,614,1852,1874]" class="Actinopterygii" family="Sisoridae" genus="Exostoma" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="265" phylum="Chordata" rank="species" species="andersonii">
<emphasis box="[396,614,1852,1874]" italics="true" pageId="3" pageNumber="265">Exostoma andersonii</emphasis>
</taxonomicName>
from four specimens collected from Yunnan, China: two from Hotham (¼Husaxiang, 24828
<emphasis bold="true" box="[359,364,1908,1923]" pageId="3" pageNumber="265">
<superScript attach="right" box="[359,364,1908,1923]" fontSize="6" pageId="3" pageNumber="265">0</superScript>
</emphasis>
09
<emphasis bold="true" box="[394,402,1908,1923]" pageId="3" pageNumber="265">
<superScript attach="right" box="[394,402,1908,1923]" fontSize="6" pageId="3" pageNumber="265">00</superScript>
</emphasis>
N, 97853
<emphasis bold="true" box="[501,506,1908,1923]" pageId="3" pageNumber="265">
<superScript attach="right" box="[501,506,1908,1923]" fontSize="6" pageId="3" pageNumber="265">0</superScript>
</emphasis>
54
<emphasis bold="true" box="[536,544,1908,1923]" pageId="3" pageNumber="265">
<superScript attach="right" box="[536,544,1908,1923]" fontSize="6" pageId="3" pageNumber="265">00</superScript>
</emphasis>
E) and two from Ponsee (approx. 24828
<emphasis bold="true" box="[322,327,1938,1953]" pageId="3" pageNumber="265">
<superScript attach="right" box="[322,327,1938,1953]" fontSize="6" pageId="3" pageNumber="265">0</superScript>
</emphasis>
N, 97842
<emphasis bold="true" box="[438,443,1938,1953]" pageId="3" pageNumber="265">
<superScript attach="right" box="[438,443,1938,1953]" fontSize="6" pageId="3" pageNumber="265">0</superScript>
</emphasis>
E). Both localities are in the Irrawaddy drainage; see
<bibRefCitation author="Anderson, J." box="[413,599,1971,1994]" journalOrPublisher="Macmillan, London" pageId="3" pageNumber="265" refId="ref3678" refString="Anderson, J. 1876. Mandalay to Momien: A Narrative of the Two Expeditions to Western China of 1868 and 1875, under Colonel Edward B. Sladen and Colonel Horace Browne. Macmillan, London." title="Mandalay to Momien: A Narrative of the Two Expeditions to Western China of 1868 and 1875, under Colonel Edward B. Sladen and Colonel Horace Browne" type="book" year="1876">Anderson (1876)</bibRefCitation>
for travel itinerary and maps.
<bibRefCitation author="Anderson, J." box="[269,514,2001,2024]" journalOrPublisher="Quaritch, London" pageId="3" pageNumber="265" refId="ref3717" refString="Anderson, J. 1879. Anatomical and Zoological Researches: Comprising an Account of the Zoological Results of the Two Expeditions to Western Yunnan in 1868 and 1875; and a Monograph of the Two Cetacean Genera, Platanista and Orcella. Quaritch, London." title="Anatomical and Zoological Researches: Comprising an Account of the Zoological Results of the Two Expeditions to Western Yunnan in 1868 and 1875; and a Monograph of the Two Cetacean Genera, Platanista and Orcella" type="book" year="1879">Anderson (1879: 866)</bibRefCitation>
explicitly mentioned that he collected only four specimens, on which Day based his description. The four syntypes were deposited in the Zoological Survey of India in Kolkata, India (ZSI), where
<bibRefCitation author="Hora, S. L." box="[859,1039,1429,1452]" pageId="3" pageNumber="265" pagination="1 - 44" refId="ref3993" refString="Hora, S. L. 1923. Notes on fishes in the Indian Museum. V. On the composite genus Glyptosternon McClelland. Records of the Indian Museum 25: 1 - 44, Pls. 1 - 4." type="journal article" year="1923">Hora (1923: 37)</bibRefCitation>
examined them. He reported that two of them are desiccated and have fallen into pieces; the remaining two are in a better condition. In a revision of glyptosternoid fishes,
<bibRefCitation author="Hora, S. L. &amp; E. G. Silas" box="[1133,1424,1519,1542]" pageId="3" pageNumber="265" pagination="5 - 29" refId="ref4035" refString="Hora, S. L., and E. G. Silas. 1952 [1951]. Notes on fishes in the Indian Museum. XLVII. Revision of the glyptosternoid fishes of the family Sisoridae, with descriptions of new genera and species. Records of the Indian Museum 49: 5 - 29, Pl. 1." type="journal article" year="1952">Hora and Silas (1952: 22)</bibRefCitation>
mention ZSI F9173/1 (1) and 9174/1 (1), both desiccated and one with head separated from body, and comment that the two remaining specimens (ZSI 596) have been lost, apparently in the 1943 flood that destroyed part of the museums collection (
<bibRefCitation author="Chopra, B." box="[984,1140,1670,1693]" journalOrPublisher="Records of the Indian Museum" pageId="3" pageNumber="265" pagination="347 - 355" part="44" refId="ref3764" refString="Chopra, B. 1946. Zoological Survey of India, 1942 - 1945. Records of the Indian Museum 44: 347 - 355." title="Zoological Survey of India" type="journal article" year="1946">Chopra, 1946</bibRefCitation>
).
</paragraph>
<paragraph blockId="3.[859,1529,1339,2024]" pageId="3" pageNumber="265">
In a catalog of fish type-specimens in ZSI,
<bibRefCitation author="Menon, A. G. K. &amp; G. M. Yazdani" pageId="3" pageNumber="265" pagination="91 - 190" refId="ref4218" refString="Menon, A. G. K., and G. M. Yazdani. 1968 [1963]. Catalogue of type specimens in the Zoological Survey of India. Part 2. Fishes. Records of the Zoological Survey of India 61: 91 - 190." type="journal article" year="1968">Menon and Yazdani (1968: 127)</bibRefCitation>
mention two specimens and give three catalog numbers F 9173/1; F 9174/1 &amp; A.S.B. Cat. No. 596 (ASB refers to the Asiatic Society of Bengal, whose collection formed the nucleus of ZSI collection). In a catalog of the Day fish collections,
<bibRefCitation author="Whitehead, P. J. P. &amp; P. K. Talwar" box="[1041,1426,1851,1874]" pageId="3" pageNumber="265" pagination="1 - 4" refId="ref4555" refString="Whitehead, P. J. P., and P. K. Talwar. 1976. Francis Day (1829 - 1889) and his collections of Indian fishes. Bulletin of the British Museum (Natural History) Historical Series 5: 1 - 189, Pls. 1 - 4." type="book chapter" year="1976">Whitehead and Talwar (1976: 157)</bibRefCitation>
mention that two specimens (A.595) were present in ZSI and two specimens (A.596) were lost or destroyed. Some of the specimens in the Day collection had been given to AMS (Australian Museum, Sydney) in 1884 by Day himself; among them, AMS B.8081, 127 mm SL, labeled from
</paragraph>
<paragraph blockId="3.[27,938,2060,2097]" box="[27,765,2060,2078]" pageId="3" pageNumber="265">Downloaded From: https://bioone.org/journals/Ichthyology-&amp;-Herpetology on 08 Mar 2023</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7716364" ID-Zenodo-Dep="7716364" httpUri="https://zenodo.org/record/7716364/files/figure.png" pageId="4" pageNumber="266" startId="4.[103,131,995,1015]" targetBox="[103,1489,178,965]" targetPageId="4">
<paragraph blockId="4.[103,1491,994,1041]" pageId="4" pageNumber="266">
<emphasis bold="true" box="[103,323,995,1015]" pageId="4" pageNumber="266">FIG. 4. Lateral views of</emphasis>
<taxonomicName box="[329,561,994,1015]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="andersonii">
<emphasis box="[329,561,994,1015]" italics="true" pageId="4" pageNumber="266">Glaridoglanis andersonii</emphasis>
</taxonomicName>
:
<emphasis bold="true" box="[573,841,995,1015]" pageId="4" pageNumber="266">(A) NRM 10721, holotype of</emphasis>
<taxonomicName authorityName="Rendahl &amp; Vestergren" authorityYear="1941" box="[846,1064,994,1015]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="malaisei">
<emphasis box="[846,1064,994,1015]" italics="true" pageId="4" pageNumber="266">Glyptosternon malaisei</emphasis>
</taxonomicName>
, 70.4 mm SL; (B) CMK 26593, 127.6 mm SL. Images not to scale. Figure 4A courtesy of NRM.
</paragraph>
</caption>
<paragraph blockId="4.[103,773,1086,2024]" pageId="4" pageNumber="266">
Bhamo, China was treated as a possible syntype by
<bibRefCitation author="Ferraris, C. J., Jr. &amp; M. A. McGrouther &amp; K. L. Parkinson" pageId="4" pageNumber="266" pagination="289 - 306" refId="ref3931" refString="Ferraris, C. J., Jr., M. A. McGrouther, and K. L. Parkinson. 2000. A critical review of the types and putative types of southern Asian marine and freshwater fish species in the Australian Museum named by Francis Day. Records of the Australian Museum 52: 289 - 306." type="journal article" year="2000">Ferraris et al. (2000: 294)</bibRefCitation>
. However, since there were only four syntypes and Hora had examined all in 1923 in ZSI, the AMS specimen cannot be a syntype.
</paragraph>
<paragraph blockId="4.[103,773,1086,2024]" pageId="4" pageNumber="266">
We were unable to examine the ZSI syntypes, but were able to examine photographs and radiographs of the AMS specimen. A comparison with the data given in
<bibRefCitation author="Day, F." box="[649,773,1263,1286]" pageId="4" pageNumber="266" pagination="511 - 527" refId="ref3859" refString="Day, F. 1870. Remarks on some of the fishes in the Calcutta Museum. - Part I. Proceedings of the Zoological Society of London 1869: 511 - 527." type="journal article" year="1870">Day (1870)</bibRefCitation>
and
<bibRefCitation author="Hora, S. L." box="[152,285,1292,1315]" pageId="4" pageNumber="266" pagination="1 - 44" refId="ref3993" refString="Hora, S. L. 1923. Notes on fishes in the Indian Museum. V. On the composite genus Glyptosternon McClelland. Records of the Indian Museum 25: 1 - 44, Pls. 1 - 4." type="journal article" year="1923">Hora (1923)</bibRefCitation>
indicates that the syntypes of
<taxonomicName box="[628,765,1292,1314]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="andersonii">
<emphasis box="[628,765,1292,1314]" italics="true" pageId="4" pageNumber="266">G. andersonii</emphasis>
</taxonomicName>
, the AMS specimen, and specimens from northern Myanmar (all collected within 200 km of both type localities of
<taxonomicName class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="andersonii">
<emphasis italics="true" pageId="4" pageNumber="266">G. andersonii</emphasis>
</taxonomicName>
) we have examined all share 910 branched pectoral-fin rays and an adipose fin with at most a weak incision at the posterior extremity of its base. Based on this information, we conclude that all of these specimens are conspecific.
</paragraph>
<paragraph blockId="4.[103,773,1086,2024]" lastBlockId="4.[820,1490,1085,1489]" pageId="4" pageNumber="266">
<taxonomicName authorityName="Rendahl &amp; Vestergren" authorityYear="1941" box="[127,365,1529,1551]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="malaisei">
<emphasis box="[127,365,1529,1551]" italics="true" pageId="4" pageNumber="266">Glyptosternon malaisei</emphasis>
</taxonomicName>
has been considered a valid species of
<taxonomicName authorityName="McClelland" authorityYear="1842" box="[133,275,1558,1580]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[133,275,1558,1580]" italics="true" pageId="4" pageNumber="266">Glyptosternon</emphasis>
</taxonomicName>
in recent checklists (
<bibRefCitation author="Ferraris, C. J., Jr." box="[513,666,1559,1581]" pageId="4" pageNumber="266" pagination="1 - 628" refId="ref3893" refString="Ferraris, C. J., Jr. 2007. Checklist of catfishes, recent and fossil (Osteichthyes: Siluriformes), and catalogue of siluriform primary types. Zootaxa 1418: 1 - 628." type="journal article" year="2007">Ferraris, 2007</bibRefCitation>
;
<bibRefCitation author="Kottelat, M." pageId="4" pageNumber="266" pagination="1 - 663" refId="ref4097" refString="Kottelat, M. 2013. The fishes of the inland waters of Southeast Asia: a catalogue and core bibiography of the fishes known to occur in freshwaters, mangroves and estuaries. Raffles Bulletin of Zoology Supplement 27: 1 - 663." type="journal article" year="2013">Kottelat, 2013</bibRefCitation>
), and its taxonomic status has not been discussed in recent studies on
<taxonomicName authorityName="McClelland" authorityYear="1842" box="[302,444,1617,1639]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[302,444,1617,1639]" italics="true" pageId="4" pageNumber="266">Glyptosternon</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation author="Thoni, R. J. &amp; E. Simonov &amp; O. Artaev &amp; S. Asylbaeva &amp; S. U. Aibek &amp; B. A. Levin" box="[514,717,1617,1640]" pageId="4" pageNumber="266" pagination="435 - 442" refId="ref4488" refString="Thoni, R. J., E. Simonov, O. Artaev, S. Asylbaeva, S. U. Aibek, and B. A. Levin. 2017. A century in synonymy: molecular and morphological evidence for the revalidation of Glyptosternon osshanini (Herzenstein, 1889) (Actinopterygii: Sisoridae). Zootaxa 4277: 435 - 442." type="journal article" year="2017">Thoni et al., 2017</bibRefCitation>
). We have examined the holotype and reidentify it as a species of
<taxonomicName authorityName="Norman" authorityYear="1925" box="[103,244,1676,1698]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[103,244,1676,1698]" italics="true" pageId="4" pageNumber="266">Glaridoglanis</emphasis>
</taxonomicName>
, on the basis of the spatulate, homodont dentition (vs. conical, homodont dentition in
<taxonomicName authorityName="McClelland" authorityYear="1842" box="[622,764,1706,1728]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[622,764,1706,1728]" italics="true" pageId="4" pageNumber="266">Glyptosternon</emphasis>
</taxonomicName>
) on both jaws and the premaxillary tooth patch not extending (vs. extending in
<taxonomicName authorityName="McClelland" authorityYear="1842" box="[435,577,1765,1787]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[435,577,1765,1787]" italics="true" pageId="4" pageNumber="266">Glyptosternon</emphasis>
</taxonomicName>
) posterolaterally.
<bibRefCitation author="Rendahl, H. &amp; G. Vestergren" box="[103,457,1795,1818]" pageId="4" pageNumber="266" pagination="213 - 214" refId="ref4388" refString="Rendahl, H., and G. Vestergren. 1941. Eine neue Art der Gattung Glyptosternon s. str. aus dem nordostlichen Birma. Zoologischer Anzeiger 133: 213 - 214." type="journal article" year="1941">Rendahl and Vestergren (1941)</bibRefCitation>
only compared
<taxonomicName authorityName="Rendahl &amp; Vestergren" authorityYear="1941" box="[650,772,1795,1817]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="malaisei">
<emphasis box="[650,772,1795,1817]" italics="true" pageId="4" pageNumber="266">G. malaisei</emphasis>
</taxonomicName>
with
<taxonomicName baseAuthorityName="Regan" baseAuthorityYear="1905" box="[169,325,1824,1846]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="maculatum">
<emphasis box="[169,325,1824,1846]" italics="true" pageId="4" pageNumber="266">G. maculatum</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="McClelland" authorityYear="1842" box="[397,553,1824,1846]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="reticulatum">
<emphasis box="[397,553,1824,1846]" italics="true" pageId="4" pageNumber="266">G. reticulatum</emphasis>
</taxonomicName>
, distinguishing
<taxonomicName authorityName="Rendahl &amp; Vestergren" authorityYear="1941" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="malaisei">
<emphasis italics="true" pageId="4" pageNumber="266">G. malaisei</emphasis>
</taxonomicName>
from them by the morphology of the adipose fin, coloration, and differences in the relative sizes of the eye and pectoral fin. Our studies fail to find any significant differences in morphology and morphometry between the holotype of
<taxonomicName authorityName="Rendahl &amp; Vestergren" authorityYear="1941" box="[238,354,1972,1994]" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="malaisei">
<emphasis box="[238,354,1972,1994]" italics="true" pageId="4" pageNumber="266">G. malaisei</emphasis>
</taxonomicName>
(
<figureCitation box="[369,447,1972,1995]" captionStart="FIG" captionStartId="4.[103,131,995,1015]" captionTargetBox="[103,1489,178,965]" captionTargetId="figure-539@4.[103,1489,178,965]" captionTargetPageId="4" captionText="FIG. 4. Lateral views of Glaridoglanis andersonii: (A) NRM 10721, holotype of Glyptosternon malaisei, 70.4 mm SL; (B) CMK 26593, 127.6 mm SL. Images not to scale. Figure 4A courtesy of NRM." figureDoi="http://doi.org/10.5281/zenodo.7716364" httpUri="https://zenodo.org/record/7716364/files/figure.png" pageId="4" pageNumber="266">Fig. 4A</figureCitation>
) and fresh material identified as
<taxonomicName box="[132,384,2001,2023]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="andersonii">
<emphasis box="[132,384,2001,2023]" italics="true" pageId="4" pageNumber="266">Glaridoglanis andersonii</emphasis>
</taxonomicName>
collected from northern Myanmar (CMK 26593;
<figureCitation box="[982,1064,1085,1108]" captionStart="FIG" captionStartId="4.[103,131,995,1015]" captionTargetBox="[103,1489,178,965]" captionTargetId="figure-539@4.[103,1489,178,965]" captionTargetPageId="4" captionText="FIG. 4. Lateral views of Glaridoglanis andersonii: (A) NRM 10721, holotype of Glyptosternon malaisei, 70.4 mm SL; (B) CMK 26593, 127.6 mm SL. Images not to scale. Figure 4A courtesy of NRM." figureDoi="http://doi.org/10.5281/zenodo.7716364" httpUri="https://zenodo.org/record/7716364/files/figure.png" pageId="4" pageNumber="266">Fig. 4B</figureCitation>
). We therefore consider
<taxonomicName authorityName="Rendahl &amp; Vestergren" authorityYear="1941" class="Actinopterygii" family="Sisoridae" genus="Glyptosternon" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="malaisei">
<emphasis italics="true" pageId="4" pageNumber="266">Glyptosternon malaisei</emphasis>
</taxonomicName>
to be a junior subjective synonym of
<taxonomicName class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="andersonii">
<emphasis italics="true" pageId="4" pageNumber="266">Glaridoglanis andersonii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="4.[820,1490,1085,1489]" pageId="4" pageNumber="266">
One of the characters previously used to diagnose
<taxonomicName authorityName="Norman" authorityYear="1925" box="[820,958,1203,1225]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[820,958,1203,1225]" italics="true" pageId="4" pageNumber="266">Glaridoglanis</emphasis>
</taxonomicName>
from other glyptosternines is the presence of 810 branched pectoral-fin rays (
<bibRefCitation author="Ng, H. H." box="[1195,1302,1232,1255]" pageId="4" pageNumber="266" pagination="97 - 157" refId="ref4268" refString="Ng, H. H. 2015. Phylogenetic systematics of the Asian catfish family Sisoridae (Actinopterygii: Siluriformes). Ichthyological Exploration of Freshwaters 26: 97 - 157." type="journal article" year="2015">Ng, 2015</bibRefCitation>
), which is lower than that reported in many other genera (1116 in
<taxonomicName authorityName="Sauvage" authorityYear="1874" box="[820,980,1290,1312]" class="Actinopterygii" family="Sisoridae" genus="Chimarrichthys" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[820,980,1290,1312]" italics="true" pageId="4" pageNumber="266">Chimarrichthys</emphasis>
</taxonomicName>
, 1416 in
<taxonomicName authorityName="Zhou, Li &amp; Thomson" authorityYear="2011" box="[1104,1280,1290,1312]" class="Actinopterygii" family="Sisoridae" genus="Creteuchiloglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[1104,1280,1290,1312]" italics="true" pageId="4" pageNumber="266">Creteuchiloglanis</emphasis>
</taxonomicName>
, 1720 in
<taxonomicName authorityName="Smith" authorityYear="1933" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="266">Oreoglanis</emphasis>
</taxonomicName>
, 1617 in
<taxonomicName authorityName="Wu, He &amp; Chu" authorityYear="1981" box="[978,1135,1320,1342]" class="Actinopterygii" family="Sisoridae" genus="Parachiloglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[978,1135,1320,1342]" italics="true" pageId="4" pageNumber="266">Parachiloglanis</emphasis>
</taxonomicName>
, 1316 in
<taxonomicName authorityName="Pellegrin" authorityYear="1936" box="[1263,1431,1320,1342]" class="Actinopterygii" family="Sisoridae" genus="Pareuchiloglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[1263,1431,1320,1342]" italics="true" pageId="4" pageNumber="266">Pareuchiloglanis</emphasis>
</taxonomicName>
, and 1618 in
<taxonomicName authorityName="Chu" authorityYear="1979" box="[925,1080,1349,1371]" class="Actinopterygii" family="Sisoridae" genus="Pseudexostoma" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[925,1080,1349,1371]" italics="true" pageId="4" pageNumber="266">Pseudexostoma</emphasis>
</taxonomicName>
). The discovery of
<taxonomicName authorityName="Ng &amp; Kottelat" authorityYear="2022" box="[1294,1401,1350,1371]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="species" species="ramosa">
<emphasis box="[1294,1401,1350,1371]" italics="true" pageId="4" pageNumber="266">G. ramosa</emphasis>
</taxonomicName>
renders the number of branched pectoral-fin rays (now 814) less useful as a diagnostic character for the genus, now distinguishing
<taxonomicName authorityName="Norman" authorityYear="1925" box="[926,1064,1437,1459]" class="Actinopterygii" family="Sisoridae" genus="Glaridoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[926,1064,1437,1459]" italics="true" pageId="4" pageNumber="266">Glaridoglanis</emphasis>
</taxonomicName>
only from
<taxonomicName authorityName="Smith" authorityYear="1933" box="[1200,1308,1437,1459]" class="Actinopterygii" family="Sisoridae" genus="Oreoglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[1200,1308,1437,1459]" italics="true" pageId="4" pageNumber="266">Oreoglanis</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Wu, He &amp; Chu" authorityYear="1981" box="[1326,1483,1437,1459]" class="Actinopterygii" family="Sisoridae" genus="Parachiloglanis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[1326,1483,1437,1459]" italics="true" pageId="4" pageNumber="266">Parachiloglanis</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Chu" authorityYear="1979" box="[870,1025,1466,1488]" class="Actinopterygii" family="Sisoridae" genus="Pseudexostoma" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="266" phylum="Chordata" rank="genus">
<emphasis box="[870,1025,1466,1488]" italics="true" pageId="4" pageNumber="266">Pseudexostoma</emphasis>
</taxonomicName>
in this regard.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,106 @@
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Key to species of
<taxonomicName id="49D44D345931FF935421408EFDE908C5" ID-CoL="5CX9J" authorityName="WOLLASTON" authorityYear="1854" box="[364,541,1222,1244]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="BCA0EAA55931FF935421408EFDE908C5" bold="true" box="[364,541,1222,1244]" italics="true" pageId="4" pageNumber="329">Zargus schaumii</emphasis>
</taxonomicName>
-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C6CE653C5931FF9255FF40BDFBCA0DD9" lastPageId="5" lastPageNumber="330" pageId="4" pageNumber="329" type="key">
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<paragraph id="8E6B36B75931FF9355FF40BDFBC9097D" blockId="4.[178,1085,1268,1601]" pageId="4" pageNumber="329">
<keyLead id="352595825931FF9355FF40BDFBC9097D" pageId="4" pageNumber="329">
1 Elytral striae interrupted at about regular intervals, consisting of a series of elongate impressions strongly deepened. Pronotum cordiform with lateral sides towards posterior angles more or less rectilinearly or even weakly sinuately narrowed, on disc usually strongly wrinkled. Antennae and legs long and slender (ratio BL/MTL about 3.12).
<collectingRegion id="4C10F8555931FF93545B4106FE92097D" box="[278,358,1359,1380]" country="Portugal" name="Madeira" pageId="4" pageNumber="329">Madeira</collectingRegion>
main isle.........................................................................
<taxonomicName id="49D44D345931FF9356CB4106FBC9097D" ID-CoL="5CX9J" authority="WOLL." authorityName="WOLLASTON" authorityYear="1854" box="[902,1085,1359,1380]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="BCA0EAA55931FF9356CB4106FC03097D" box="[902,1015,1359,1380]" italics="true" pageId="4" pageNumber="329">Z. schaumii</emphasis>
WOLL.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8E6B36B75931FF9355FF4127FBC909DE" blockId="4.[178,1085,1268,1601]" pageId="4" pageNumber="329">
<keyLead id="352595825931FF9355FF4127FBC909DE" pageId="4" pageNumber="329">- Elytral striae interrupted less regularly, the elongate impressions flat or only suggested. Pronotum subquadrate with lateral sides towards posterior angles more or less weakly rounded, or almost rectangularly narrowed, on disc usually less strongly wrinkled. Antennae and legs shorter and rather more robust (ratio BL/MTL about 3.21-3.30)..........2</keyLead>
</paragraph>
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<keyLead id="352595825931FF9255FF4198FBC90D1B" lastPageId="5" lastPageNumber="330" pageId="4" pageNumber="329">
2 Elytral striae with a weak tendency to be interrupted with shallow, not very noticeable elongate impressions, disc weakly convex, subapical lateral sinuation weak, each elytron broadly rounded apically (
<figureCitation id="16EF2A325931FF93570841B7FD730A0A" box="[581,647,1534,1555]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="4" pageNumber="329">Fig. 8</figureCitation>
). Pronotum with lateral sides towards posterior angles more or less weakly rounded. Labrum with lobes narrow, acuminate, only weakly rounded at tip (
<figureCitation id="16EF2A325931FF9354B54265FDC20A58" box="[504,566,1580,1601]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="4" pageNumber="329">Fig. 6</figureCitation>
). Legs shorter (ratio BL/MTL about 3.30). Body piceous black with margins of pronotum and elytra narrowly lightened, legs more or less darkened. Deserta Grande. ...............................................................
<taxonomicName id="49D44D345930FF9256C644A4FBC90D1B" ID-CoL="5CX9D" authority="WOLL." authorityName="WOLLASTON" authorityYear="1854" box="[907,1085,236,258]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="330" phylum="Arthropoda" rank="species" species="desertae">
<emphasis id="BCA0EAA55930FF9256C644A4FC020D18" box="[907,1014,236,258]" italics="true" pageId="5" pageNumber="330">Z. desertae</emphasis>
WOLL.
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<keyLead id="352595825930FF9255FF4545FBCA0DD9" pageId="5" pageNumber="330">
- Elytral striae with stronger and deeper elongate impressions (but not as strong as in
<taxonomicName id="49D44D345930FF92515D4545FEDB0D2E" ID-CoL="5CX9J" authorityName="WOLLASTON" authorityYear="1854" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="330" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="BCA0EAA55930FF92515D4545FEDB0D2E" italics="true" pageId="5" pageNumber="330">Z. schaumii</emphasis>
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), disc flattened, subapical lateral sinuation stronger, sutural angles somewhat angulate (
<figureCitation id="16EF2A325930FF92540D4570FE710D57" box="[320,389,313,334]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="5" pageNumber="330">Fig. 7</figureCitation>
). Pronotum with lateral sides towards posterior angles almost rectangularly narrowed. Labrum with lobes not acuminate, wider, widely rounded at tip (
<figureCitation id="16EF2A325930FF9255B2452FFECF0D62" box="[255,315,358,379]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="5" pageNumber="330">Fig. 5</figureCitation>
). Legs somewhat longer (ratio BL/MTL 3.21). Body reddish piceous, with a very faint brassy tinge, margins of pronotum and elytra somewhat wider lightened, appendages yellowish with tarsi somewhat infuscated.
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main isle........................... .................................................................................................................
<taxonomicName id="49D44D345930FF9256C245E2FC180DD9" ID-CoL="5CX9H" authority="Wrase, 2010" authorityName="Wrase" authorityYear="2010" box="[911,1004,427,448]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="330" phylum="Arthropoda" rank="species" species="putzeri" status="sp. nov.">
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</treatment>
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<emphasis id="BCA0EAA55937FF9555FF40FFFEBF08D2" bold="true" box="[178,331,1206,1227]" italics="true" pageId="2" pageNumber="327">Zargus putzeri</emphasis>
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y p e m a t e r i a l:
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: &quot;
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&gt; /
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, below / nesting colony of
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/
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(MATHEWS 1934) / (Genista-Erica arborea zone, moist litter, sifted) /
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&quot; (cWR).
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D i a g n o s i s: A typical
<taxonomicName id="49D44D345937FF9554934171FDD20954" box="[478,550,1336,1357]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="327" phylum="Arthropoda" rank="genus">
<emphasis id="BCA0EAA55937FF9554934171FDD20954" box="[478,550,1336,1357]" italics="true" pageId="2" pageNumber="327">Zargus</emphasis>
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species of large size (with all characters given by
<bibRefCitation id="EA454B465937FF9555FF411CFE610972" author="MACHADO A" box="[178,405,1365,1387]" pageId="2" pageNumber="327" refId="ref4138" refString="MACHADO A. (1992): Monografia de los carabidos de las Islas Canarias (Insecta, Coleoptera). - La Laguna: Instituto de Estudios Canarios, 734 pp." type="book" year="1992">MACHADO 1992: 445</bibRefCitation>
: head elongate, two supraorbital setae, clypeus widely excavate, labrum bilobed, V-shaped, with base widely membranous, mentum without tooth, mandibles elongate, bidentate, ligula membranous, bisetose, paraglossa weakly protruding, labial palpomere 2 bisetose, antennae fine and long, pubescent from antennomere 4, pronotum distincly flattened along margin, elytra with scutellar pore puncture in interval 1 and with two pore punctures in or at stria 2, hindwings reduced to small relicts). Body reddish piceous, with a very faint brassy tinge, appendages yellowish with tarsi somewhat infuscated. Pronotum almost quadratic, elytra ovate with striae somewhat irregu- larly interrupted, and consisting of a series of shallow, elongate impressions. Habitus see
<figureCitation id="16EF2A325936FF9455FF44B1FF070D17" box="[178,243,248,270]" captionStart="Figs 1-3" captionStartId="8.[178,218,820,840]" captionTargetBox="[197,1072,231,792]" captionTargetId="figure-43@8.[178,1085,219,806]" captionTargetPageId="8" captionText="Figs 1-3: Zargus, habitus: (1) Z. putzeri nov.sp.; (2) Z. schaumii WOLL.; (3) Z. desertae WOLL." figureDoi="http://doi.org/10.5281/zenodo.4507201" httpUri="https://zenodo.org/record/4507201/files/figure.png" pageId="3" pageNumber="328">Fig. 1.</figureCitation>
</paragraph>
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D e s c r i p t i o n: Body length
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; width
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.
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<paragraph id="8E6B36B75936FF9455FF450BFDA80D8A" blockId="3.[178,1085,322,403]" pageId="3" pageNumber="328">Colour: Body reddish piceous, with a very faint brassy tinge, elytral interval 7 and 8 somewhat infuscated, margins of pronotum and elytra somewhat lightened, appendages yellowish with tarsi somewhat infuscated.</paragraph>
<paragraph id="8E6B36B75936FF9455FF45EBFEBA0E34" blockId="3.[178,1086,418,557]" pageId="3" pageNumber="328">
Head (
<figureCitation id="16EF2A325936FF9455BB45EBFEC60DA1" box="[246,306,418,440]" captionStart="Figs 1-3" captionStartId="8.[178,218,820,840]" captionTargetBox="[197,1072,231,792]" captionTargetId="figure-43@8.[178,1085,219,806]" captionTargetPageId="8" captionText="Figs 1-3: Zargus, habitus: (1) Z. putzeri nov.sp.; (2) Z. schaumii WOLL.; (3) Z. desertae WOLL." figureDoi="http://doi.org/10.5281/zenodo.4507201" httpUri="https://zenodo.org/record/4507201/files/figure.png" pageId="3" pageNumber="328">Fig. 1</figureCitation>
) elongate, much narrower than pronotum (ratio PW/HW: 1.67). Eyes small and fairly flat, eye diameter about 3.64 times as long as almost rectilinearly narrowed tempora (seen in dorsal view). Lobes of labrum wide (
<figureCitation id="16EF2A325936FF9457B74594FCCF0DEA" box="[762,827,477,499]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="3" pageNumber="328">Fig. 5</figureCitation>
), widely rounded at tip. Antennae long, slender, reaching scarcely to middle of elytra, pubescent from antennomere 4.
</paragraph>
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Pronotum (
<figureCitation id="16EF2A325936FF94546A4674FE910E4A" box="[295,357,573,595]" captionStart="Figs 1-3" captionStartId="8.[178,218,820,840]" captionTargetBox="[197,1072,231,792]" captionTargetId="figure-43@8.[178,1085,219,806]" captionTargetPageId="8" captionText="Figs 1-3: Zargus, habitus: (1) Z. putzeri nov.sp.; (2) Z. schaumii WOLL.; (3) Z. desertae WOLL." figureDoi="http://doi.org/10.5281/zenodo.4507201" httpUri="https://zenodo.org/record/4507201/files/figure.png" pageId="3" pageNumber="328">Fig. 1</figureCitation>
) almost square (ratio PW/PL: 1.25), widest at about end of anterior third, shortly behind insertion of lateral seta. Anterior margin strongly emarginate, anterior angles somewhat acutely projecting forward, rounded at tip, from there slightly curved laterally till insertion of lateral seta, from there basad almost rectangularly narrowed to posterior angles which are rounded and angled strongly forward, at insertion of seta very weakly angulate. Base medially slightly emarginate, laterally to insertion of seta strongly curved (ratio PW/PBaW: 1.16), distinctly wider than anterior margin (ratio PBaW/PAW: 1.27). Distincly flattened along lateral margin, the latter somewhat reflexed anterior to posterior angles, disc flat. Medial longitudinal impression fine, not deeply impressed, terminated at anterior transverse impression and, becoming indistinct, terminated somewhat anterior to basal margin, posterior transverse impression indistinct, basal impressions round and shallow, whole pronotum impunctate, around termination of longitudinal impression at base somewhat weakly longitudinally wrinkled.
</paragraph>
<paragraph id="8E6B36B75936FF9455FF4788FDA408E5" blockId="3.[178,1086,961,1276]" pageId="3" pageNumber="328">
Elytra (
<figureCitation id="16EF2A325936FF9455B34788FECD0FCE" box="[254,313,961,983]" captionStart="Figs 1-3" captionStartId="8.[178,218,820,840]" captionTargetBox="[197,1072,231,792]" captionTargetId="figure-43@8.[178,1085,219,806]" captionTargetPageId="8" captionText="Figs 1-3: Zargus, habitus: (1) Z. putzeri nov.sp.; (2) Z. schaumii WOLL.; (3) Z. desertae WOLL." figureDoi="http://doi.org/10.5281/zenodo.4507201" httpUri="https://zenodo.org/record/4507201/files/figure.png" pageId="3" pageNumber="328">Fig. 1</figureCitation>
) ovate (ratio EL/EW: 1.37), with rounded humeri, widest somewhat behind middle, flat on disc. Basal bead strongly sinuate, strongly curved inside humerus, weakly curved towards lateral margin, subapical lateral sinuation weak, sutural angle somewhat angulate (
<figureCitation id="16EF2A325936FF9454544050FEAF0836" box="[281,347,1049,1071]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="3" pageNumber="328">Fig. 7</figureCitation>
). Scutellar pore puncture present in interval 1, two pore punctures in stria 2, the first one shortly anterior to middle, the second one about at beginning of apical fourth. Striae fine, somewhat irregularly interrupted, and consisting of a series of shallow, elongate impressions. Umbellicate series of left elytron with 13 setigerous punctures (forming three groups: 5+1+7), at right side consisting of 15 punctures in a row with punctures of somewhat greater distance from each other at middle. Elytra obviously fused together at suture, relicts of hindwings after removing of abdomen not visible, in all probability extremely reduced.
</paragraph>
<paragraph id="8E6B36B75936FF9455FF4142FC340945" blockId="3.[178,1085,1291,1372]" pageId="3" pageNumber="328">Sterna: Impunctate, abdominal sternites laterally somewhat wrinkled. Metepisternum short, ventral margin about as long as anterior margin. Last sternite with about 14 seta subapically and with two setae inserted at middle somewhat anterior to them.</paragraph>
<paragraph id="8E6B36B75936FF9455FF4125FEF90987" blockId="3.[178,1087,1387,1438]" pageId="3" pageNumber="328">
Legs shorter and rather more robust than in
<taxonomicName id="49D44D345936FF9457314125FD000999" authorityName="WOLLASTON" authorityYear="1854" box="[636,756,1387,1409]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="BCA0EAA55936FF9457314125FD000999" box="[636,756,1387,1409]" italics="true" pageId="3" pageNumber="328">Z. schaumii</emphasis>
</taxonomicName>
but somewhat longer than in
<taxonomicName id="49D44D345936FF9451674125FEF30987" authorityName="WOLLASTON" authorityYear="1854" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="328" phylum="Arthropoda" rank="species" species="desertae">
<emphasis id="BCA0EAA55936FF9451674125FEF30987" italics="true" pageId="3" pageNumber="328">Z. desertae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8E6B36B75936FF9455FF41E7FD6009F8" blockId="3.[178,1085,1454,1505]" pageId="3" pageNumber="328">Microsculpture mesh pattern on upper surface consisting of strongly engraved isodiametric meshes, causing a surface fairly matt.</paragraph>
<paragraph id="8E6B36B75936FF9455FF41B9FDDC0A1F" blockId="3.[178,552,1520,1542]" box="[178,552,1520,1542]" pageId="3" pageNumber="328">Median lobe of aedeagus: Unknown.</paragraph>
<paragraph id="8E6B36B75936FF9355FF425FFE520D32" blockId="3.[178,1085,1558,1609]" lastBlockId="4.[178,1085,219,299]" lastPageId="4" lastPageNumber="329" pageId="3" pageNumber="328">
Female genitalia (
<figureCitation id="16EF2A325936FF945439425FFE4D0A35" box="[372,441,1558,1580]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="3" pageNumber="328">Fig. 9</figureCitation>
): Hemisternite asymmetrical, with some longer and shorter bristles apically. Basal stylomere without setae, apical stylomere moderately curved, with a fine double-sensilla in a furrow at external margin shortly anterior to apex, with one peg-like seta at external margin about at middle, and with two peg-like setae at internal margin anterior to apex.
</paragraph>
<paragraph id="8E6B36B75931FF9355FF4573FC760DB1" blockId="4.[178,1085,314,424]" pageId="4" pageNumber="329">
C o m p a r i s o n s: The new species belongs to the group of species with strongly interrupted elytral striae, consisting of a series of elongate impressions strongly deepened (
<taxonomicName id="49D44D345931FF9355F7453FFE8A0D92" authority="WOLL." authorityName="WOLLASTON" authorityYear="1854" box="[186,382,373,395]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="BCA0EAA55931FF9355F7453FFEC60D93" box="[186,306,373,395]" italics="true" pageId="4" pageNumber="329">Z. schaumii</emphasis>
WOLL.
</taxonomicName>
), or with light striae having a weak tendency to be interrupted with shallow, not very noticeable elongate impressions (
<taxonomicName id="49D44D345931FF9357F445DAFC860DB1" authority="WOLL." authorityName="WOLLASTON" authorityYear="1854" box="[697,882,402,424]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="desertae">
<emphasis id="BCA0EAA55931FF9357F445DAFCDE0DBE" box="[697,810,402,424]" italics="true" pageId="4" pageNumber="329">Z. desertae</emphasis>
WOLL.
</taxonomicName>
).
</paragraph>
<paragraph id="8E6B36B75931FF9355FF45F1FCA50ECC" blockId="4.[178,1085,439,725]" pageId="4" pageNumber="329">
<taxonomicName id="49D44D345931FF9355FF45F1FEED0DD4" authority="Wrase, 2010" authorityName="Wrase" authorityYear="2010" box="[178,281,440,461]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="putzeri" status="sp. nov.">
<emphasis id="BCA0EAA55931FF9355FF45F1FEED0DD4" box="[178,281,440,461]" italics="true" pageId="4" pageNumber="329">Z. putzeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A79357DE5931FF93546845F1FE720DD4" box="[293,390,440,461]" pageId="4" pageNumber="329" rank="species">nov.spec.</taxonomicNameLabel>
has a somewhat intermediate position between these species, it shares with
<taxonomicName id="49D44D345931FF93547F459FFE580DF3" authorityName="WOLLASTON" authorityYear="1854" box="[306,428,469,491]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="BCA0EAA55931FF93547F459FFE580DF3" box="[306,428,469,491]" italics="true" pageId="4" pageNumber="329">Z. schaumii</emphasis>
</taxonomicName>
the flattened elytra with subapical lateral sinuation weak and with sutural angle somewhat angulate (
<figureCitation id="16EF2A325931FF93570A45BBFD730E11" box="[583,647,498,520]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="4" pageNumber="329">Fig. 7</figureCitation>
), the labrum with lobes wider, not acuminate, widely rounded at tip, and the reddish piceous coloration of the body, often with a very faint brassy tinge (in
<taxonomicName id="49D44D345931FF93548A4664FDCD0E5B" authorityName="WOLLASTON" authorityYear="1854" box="[455,569,557,578]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="desertae">
<emphasis id="BCA0EAA55931FF93548A4664FDCD0E5B" box="[455,569,557,578]" italics="true" pageId="4" pageNumber="329">Z. desertae</emphasis>
</taxonomicName>
elytral disc weakly convex, with subapical lateral sinuation missing or only indicated (
<figureCitation id="16EF2A325931FF9357614603FD980E79" box="[556,620,586,608]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="4" pageNumber="329">Fig. 8</figureCitation>
), labrum with lobes narrow, acuminate, only weakly rounded at tip, compare
<figureCitation id="16EF2A325931FF9354B4462EFD8C0E64" box="[505,632,615,637]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="4" pageNumber="329">Figs 5 and 6</figureCitation>
, and body piceous black). In construction of the pronotum and with its legs shorter and rather more robust it is closer to
<taxonomicName id="49D44D345931FF93568B46CCFBCD0E83" authorityName="WOLLASTON" authorityYear="1854" box="[966,1081,645,666]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="desertae">
<emphasis id="BCA0EAA55931FF93568B46CCFBCD0E83" box="[966,1081,645,666]" italics="true" pageId="4" pageNumber="329">Z. desertae</emphasis>
</taxonomicName>
. Its elytral elongate impressions are distincly more shallow than in
<taxonomicName id="49D44D345931FF9356DE46EAFBFB0EAE" authorityName="WOLLASTON" authorityYear="1854" box="[915,1039,674,696]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="schaumii">
<emphasis id="BCA0EAA55931FF9356DE46EAFBFB0EAE" box="[915,1039,674,696]" italics="true" pageId="4" pageNumber="329">Z. schaumii</emphasis>
</taxonomicName>
but stronger than in
<taxonomicName id="49D44D345931FF93541A4689FE3C0ECD" authorityName="WOLLASTON" authorityYear="1854" box="[343,456,703,725]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="desertae">
<emphasis id="BCA0EAA55931FF93541A4689FE3C0ECD" box="[343,456,703,725]" italics="true" pageId="4" pageNumber="329">Z. desertae</emphasis>
</taxonomicName>
which has only indicated impressions.
</paragraph>
<paragraph id="8E6B36B75931FF9355FF46ADFBE80806" blockId="4.[178,1086,740,1055]" pageId="4" pageNumber="329">
The gonocoxites and hemisternites (
<figureCitation id="16EF2A325931FF93576246ADFD9B0EE3" box="[559,623,740,762]" captionStart="Figs 5-9" captionStartId="9.[178,218,1315,1335]" captionTargetBox="[202,1061,217,1298]" captionTargetId="figure-0@9.[178,1085,212,1304]" captionTargetPageId="9" captionText="Figs 5-9: Zargus: 5, 6: Apical part of labrum, dorsal aspect: (5) Z. putzeri nov.sp.; (6) Z. desertae WOLL.; 7, 8: Outline of apical portion of left elytron, dorsal aspect; (7) Z. putzeri nov.sp.; (8) Z. desertae WOLL.; (9) Z. putzeri nov.sp., female genitalia, left hemisternite with gonocoxites, dorsal aspect; scale bar: 0.39 mm (5, 6); 1.6 mm (7, 8); 0.47 mm (9)." figureDoi="http://doi.org/10.5281/zenodo.4507207" httpUri="https://zenodo.org/record/4507207/files/figure.png" pageId="4" pageNumber="329">Fig. 9</figureCitation>
) are similar to each other in the species in question, but more material is wanted to realize if small differences are constant. Ratios (see table 1) can show only tendencies as the number of available specimens was limited. Many values are fairly variable (the fairly strong intraspecific variability probably can be explained by the circumstance that specimens of different populations were investigated which have no contact to each other due to aptery), values between species overlap partly. The ratio PW/PBAW shows in
<taxonomicName id="49D44D345931FF93577147DCFD6A0FB3" authority="Wrase, 2010" authorityName="Wrase" authorityYear="2010" box="[572,670,917,938]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="putzeri" status="sp. nov.">
<emphasis id="BCA0EAA55931FF93577147DCFD6A0FB3" box="[572,670,917,938]" italics="true" pageId="4" pageNumber="329">Z. putzeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A79357DE5931FF9357E847DCFCF10FB3" box="[677,773,917,938]" pageId="4" pageNumber="329" rank="species">nov.spec.</taxonomicNameLabel>
the lowest value, and the ratio PBAW/PAW is the highest, not overlapping with the values of the remaining two species, but more material is needed to confirm consistence. But other external characters revealed clear differences between these three species which can be differentiated in the following key. For data on variation in some values among
<taxonomicName id="49D44D345931FF9356434043FCA20806" box="[782,854,1034,1055]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="genus">
<emphasis id="BCA0EAA55931FF9356434043FCA20806" box="[782,854,1034,1055]" italics="true" pageId="4" pageNumber="329">Zargus</emphasis>
</taxonomicName>
species see table 1.
</paragraph>
<paragraph id="8E6B36B75931FF9355FF4067FCC90885" blockId="4.[178,1086,1070,1180]" pageId="4" pageNumber="329">
The other congeners of
<taxonomicName id="49D44D345931FF9354F04066FDF1085D" box="[445,517,1071,1092]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="genus">
<emphasis id="BCA0EAA55931FF9354F04066FDF1085D" box="[445,517,1071,1092]" italics="true" pageId="4" pageNumber="329">Zargus</emphasis>
</taxonomicName>
from
<collectingRegion id="4C10F8555931FF9357024067FD57085D" box="[591,675,1070,1092]" country="Portugal" name="Madeira" pageId="4" pageNumber="329">Madeira</collectingRegion>
(
<taxonomicName id="49D44D345931FF9357F44066FC60085D" authority="WOLL." authorityName="WOLLASTON" authorityYear="1854" box="[697,916,1071,1092]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="pellucidus">
<emphasis id="BCA0EAA55931FF9357F44066FCB6085D" box="[697,834,1071,1092]" italics="true" pageId="4" pageNumber="329">Z. pellucidus</emphasis>
WOLL.
</taxonomicName>
and
<taxonomicName id="49D44D345931FF93569C4066FF010878" authority="WOLL." authorityName="WOLLASTON" authorityYear="1860" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="monizii">
<emphasis id="BCA0EAA55931FF93569C4066FBC8085D" box="[977,1084,1071,1092]" italics="true" pageId="4" pageNumber="329">Z. monizii</emphasis>
WOLL.
</taxonomicName>
) have the elytra regularly and fairly deeply striate with striae having no tendency to be interrupted, and are additionally of smaller body size, therefore
<taxonomicName id="49D44D345931FF9356384023FC230866" authority="Wrase, 2010" authorityName="Wrase" authorityYear="2010" box="[885,983,1130,1151]" class="Insecta" family="Carabidae" genus="Zargus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="329" phylum="Arthropoda" rank="species" species="putzeri" status="sp. nov.">
<emphasis id="BCA0EAA55931FF9356384023FC230866" box="[885,983,1130,1151]" italics="true" pageId="4" pageNumber="329">Z. putzeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A79357DE5931FF9356934023FBCA0866" box="[990,1086,1130,1151]" pageId="4" pageNumber="329" rank="species">nov.spec.</taxonomicNameLabel>
on these characters alone cannot be confused with these species.
</paragraph>
</subSubSection>
</treatment>
</document>

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<taxonomicName authority="Barr, 1952 a" class="Insecta" family="Cleridae" genus="Araeodontia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araeodontia marginalis" order="Coleoptera" pageId="8" pageNumber="83" phylum="Arthropoda" rank="species" species="marginalis">Araeodontia marginalis Barr, 1952a</taxonomicName>
Figs 1B, 18B
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<subSubSection pageId="9" pageNumber="84" type="paratype">
<paragraph pageId="9" pageNumber="84">
<pageBreakToken pageId="9" pageNumber="84" start="start">Paratype</pageBreakToken>
.
</paragraph>
<paragraph pageId="9" pageNumber="84">One male examined.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="84" type="type locality">
<paragraph pageId="9" pageNumber="84">Type locality.</paragraph>
<paragraph pageId="9" pageNumber="84">Mexico, Samalayuca, Chihuahua. Type depository: American Museum of Natural History (AMNH).</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="84" type="distribution">
<paragraph pageId="9" pageNumber="84">Distribution.</paragraph>
<paragraph pageId="9" pageNumber="84">USA: TX; Mexico: Chihuahua, Coahuila, Sonora.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="84" type="differential diagnosis">
<paragraph pageId="9" pageNumber="84">Differential diagnosis.</paragraph>
<paragraph pageId="9" pageNumber="84">
<taxonomicName class="Insecta" family="Cleridae" genus="Araeodontia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araeodontia marginalis" order="Coleoptera" pageId="9" pageNumber="84" phylum="Arthropoda" rank="species" species="marginalis">Araeodontia marginalis</taxonomicName>
is most similar to
<taxonomicName lsidName="A. isabellae" pageId="9" pageNumber="84" rank="species" species="isabellae">A. isabellae</taxonomicName>
. The fascia pattern on the elytral disc can be used to separate these species.
<taxonomicName class="Insecta" family="Cleridae" genus="Araeodontia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araeodontia marginalis" order="Coleoptera" pageId="9" pageNumber="84" phylum="Arthropoda" rank="species" species="marginalis">Araeodontia marginalis</taxonomicName>
has two longitudinal fasciae that extend from the elytral base to the apex, the first band is located adjacent to the elytral suture and the second runs along the epipleural fold; for some specimens, the latter band can be absent on the anterior half of the elytral disc; both fasciae are interconnected at the apex (Fig. 1B).
<taxonomicName class="Insecta" family="Cleridae" genus="Araeodontia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Araeodontia isabellae" order="Coleoptera" pageId="9" pageNumber="84" phylum="Arthropoda" rank="species" species="isabellae">Araeodontia isabellae</taxonomicName>
has the elytral disc uniformly light testaceous and each elytron has one brown macula (Fig. 1A).
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="86" pageId="9" pageNumber="84" type="redescription">
<paragraph pageId="9" pageNumber="84">Redescription.</paragraph>
<paragraph pageId="9" pageNumber="84">Male. Form: Body relatively stout, elongate. Color: Head, anterior margin of pronotum and mouthparts brown to dark brown; pronotum, thorax, elytra, abdomen and legs testaceous to light brown; two brown, longitudinal fasciae on each elytron, the first located on the elytral suture, and extends from anterior margin of elytra and reaches apex, this fascia abruptly reduced in width on second and last fourth, the second adjacent to epipleural fold, and also extends from anterior margin of elytron and reaches apex, this fascia may be reduced to absent on the anterior half of elytral length, both fasciae may be interconnected at the elytral apex (Fig. 1B).</paragraph>
<paragraph pageId="10" pageNumber="85">
<pageBreakToken pageId="10" pageNumber="85" start="start">Head</pageBreakToken>
: Feebly vested by light, recumbent setae; surface weakly punctate; frons bi-impressed; eyes enlarged, bulging laterally, coarsely faceted; antennae extending to anterior third of elytra; third antennomere about twice the length of second antennomere; antennomeres 3-10 about the same length; antennomeres 4-10 somewhat robust, slightly serrate; eleventh antennomere robust, subacuminate.
</paragraph>
<paragraph pageId="10" pageNumber="85">Thorax: Pronotum scarcely punctate; faintly rugose laterally, smooth; vested by semierect seta interspersed with fine, recumbent setae; broadest at middle; disc flat, very feebly impressed in front of middle, more strongly constricted behind middle, subbasal tumescence absent. Mesoventrite very finely vested, smooth. Metaventrite smooth, convex, puncticulate; covered with fine, semi-recumbent setae. Scutellum subquadrate, notched posteriorly.</paragraph>
<paragraph pageId="10" pageNumber="85">Legs: Vested with short, recumbent setae intermixed with long, erect and semi-erect setae. Femora rugulose; finely punctate. Tibiae longitudinally rugose, punctate, vested with short, recumbent setae intermixed with semi-erect setae.</paragraph>
<paragraph pageId="10" pageNumber="85">
Elytra: Humeri rounded, indicated; sides subparallel, widest behind middle; base wider than pronotum; disc flattened apically; apices subtriangular, slightly dehiscent; disc convex; vestiture composed of stiff, erect and semi-erect setae intermixed with stiff, semirecumbent setae; sculpturing consisting of small, shallow punctations arranged in striae that gradually reduce in size on middle third and do not reach elytral apex; interstices smooth, 4.0
<normalizedToken originalValue="×">x</normalizedToken>
the width of punctuation at anterior margin.
</paragraph>
<paragraph pageId="10" pageNumber="85">Abdomen: Ventrites 1-4 rugulose, feebly vested with short, recumbent setae, indistinctly, finely punctate. First visible ventrite about the same length of second ventrite, ventrite 3-4 subquadrate, smooth, feebly vested with fine recumbent setae. Fifth visible ventrite reduced, convex, lateral margins subparallel, posterior margin broadly, slightly emarginate. Sixth visible ventrite subquadrate, surface somewhat concave medially, convex laterally, feebly punctate; lateral margins oblique; posterior margin broadly, moderately deeply emarginate, emargination V-shaped, posterolateral angles rounded. Fifth tergite convex; finely punctate; rugulose; lateral margin subparallel; posterior margin broadly, shallowly, slightly, emarginate. Sixth tergite subtriangular; surface convex; longer than broad; finely punctate; scarcely covered with short, recumbent setae; lateral margins oblique; posterior margin narrowly, very shallowly emarginate; hind angles rounded. Posterior margin of sixth tergite produced ventrally, fully covering sixth visible ventrite.</paragraph>
<paragraph pageId="10" pageNumber="85">Aedeagus: Phallobasic apodeme present; phallus with copulatory piece expanded at apex; phallic plate without denticles; intraspicular plate present, elongate; phallobasic apodeme long, expanded distally; phallobase trigonal; parameres free; tegmen complete, covering phallus; parameres pointed anteriorly; endophallic struts long, the length of tegmen; endophallic struts slender distally (Fig. 18B).</paragraph>
<paragraph lastPageId="11" lastPageNumber="86" pageId="10" pageNumber="85">
Sexual dimorphism: The female of
<taxonomicName lsidName="A. marginalis" pageId="10" pageNumber="85" rank="species" species="marginalis">A. marginalis</taxonomicName>
can be separated from males based on the structure of the last abdominal segment. In females, the lateral and posterior margins of the sixth tergite and the sixth visible ventrite are broadly rounded, making a single semicircular margin; males have the sixth tergite and the sixth visible ventrite subquadrate in shape, and the posterior margin narrowly, shallowly emarginate, the
<pageBreakToken pageId="11" pageNumber="86" start="start">emargination</pageBreakToken>
seen in the sixth visible ventrite is slightly deeper than that observed in the sixth tergite. Remaining characters are similar.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="86" type="materials_examined">
<paragraph pageId="11" pageNumber="86">Material examined.</paragraph>
<paragraph pageId="11" pageNumber="86">PARATYPE: 1 male: Pine Springs, TX, VII-12-16-1928, W. Benedict.</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="86" type="materials_examined">
<paragraph pageId="11" pageNumber="86">Additional material examined.</paragraph>
<paragraph pageId="11" pageNumber="86">USA: 1 male: Hudspeth Co., TX, 9 mi SW Dell City, VII-31-1950, R. F. Smith; 2 males: Valentine, TX, VI-25-1947, R. H. Beawer. MEXICO: 1 male, 1 female: Sonora, Mexico, near San Jose beach, Ciudad Obregon, 40 mi SW of V-16-23-1961, Howden and Martin; 1 female: Coahuila, Mexico, sand dunes, near Bilbao, 8 mi N of Viesca, V-30-31-1981, J. Doyen and J. Liebherr.</paragraph>
</subSubSection>
</treatment>
</document>

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