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<document id="5ACAE8CB841C6512564A72BEDBD95082" ID-DOI="10.11646/zootaxa.5613.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="15216197" ID-ZooBank="FAC715B6-F3A1-470B-880F-56423F69D3A3" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1744667152350" checkinUser="plazi" docAuthor="Syomin, Vitaly, Kolbasova, Glafira, Semenova, Maya & Neretina, Tatiana" docDate="2025" docId="03C2E360FF8A7430FF4676C5AB86FAFE" docLanguage="en" docName="zootaxa.5613.1.3.pdf" docOrigin="Zootaxa 5613 (1)" docSource="https://doi.org/10.11646/zootaxa.5613.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ctenophoricola tzetlini Syomin, Kolbasova, Semenova & Neretina, 2025, sp. nov." docType="treatment" docVersion="3" lastPageNumber="93" masterDocId="FFFB9B18FF8E743BFFD17652AE1AFFAF" masterDocTitle="A new species of an unusual polychaete genus Ctenophoricola (Phyllodocida, Phyllodocidae, Alciopini) from the Indian Ocean" masterLastPageNumber="98" masterPageNumber="82" pageNumber="86" updateTime="1744809039579" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo id="4F326417A846F78BFE180C0B3F1886D2">
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<mods:title id="995CD3F5342141F80A96574D189BA09A">A new species of an unusual polychaete genus Ctenophoricola (Phyllodocida, Phyllodocidae, Alciopini) from the Indian Ocean</mods:title>
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<mods:roleTerm id="B65CE1A3F91C3492BC7E58B2AF5F2D75">Author</mods:roleTerm>
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<mods:namePart id="B8AD96CB1BAE6EAE921F5BF2AE22CF40">Syomin, Vitaly</mods:namePart>
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<mods:affiliation id="B68CD61E41224C7CE235AD3CE43CE10A">Red Sea Research Centre, King Abdullah University of Science and Technology (KAUST), Thuwal, 23955 - 6900, Kingdom of Saudi Arabia & Shirshov Institute of Oceanology of Russian Academy of Sciences, 36 Nahimovskiy Avenue, 117997 Moscow, Russia</mods:affiliation>
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<mods:nameIdentifier id="33E6961E3A20E4221D8D6138189FB837" type="email">svinovod@yandex.ru</mods:nameIdentifier>
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<mods:namePart id="28BA87B75415226E9448F58A4F962EAB">Kolbasova, Glafira</mods:namePart>
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<mods:nameIdentifier id="BC5FAC457B13D81746BF4C69E89CC123" type="ORCID">0000-0001-9252-0473</mods:nameIdentifier>
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<mods:affiliation id="3DC36AEA716B57EAF584437546718EBB">N. A. Pertsov White Sea Biological Station, Lomonosov Moscow State University, 1 Leninskie Gory, building 12, 119234 Moscow, Russia</mods:affiliation>
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<mods:nameIdentifier id="E106267103DEC02354FE5AAEF341F7A5" type="email">utricularia57@yandex.ru</mods:nameIdentifier>
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<mods:namePart id="B859E5330E368CCBBDE394CA9DD0117C">Semenova, Maya</mods:namePart>
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<mods:affiliation id="21798F015AD64FDDA1DF337A8879A638">N. A. Pertsov White Sea Biological Station, Lomonosov Moscow State University, 1 Leninskie Gory, building 12, 119234 Moscow, Russia</mods:affiliation>
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<mods:nameIdentifier id="ED52B3272C58BD9F399858554CC45DA5" type="email">maya.yakuts@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E2298FA854AC573112BA85A357148474">Neretina, Tatiana</mods:namePart>
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<mods:nameIdentifier id="578165C7CF5436110F0CD05E061D7FF0" type="ORCID">0000-0002-0673-1109</mods:nameIdentifier>
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<mods:affiliation id="9C19F4623E0A1BEEBA7AFAB9974043BA">N. A. Pertsov White Sea Biological Station, Lomonosov Moscow State University, 1 Leninskie Gory, building 12, 119234 Moscow, Russia</mods:affiliation>
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<mods:nameIdentifier id="2736E76AA80184F7AFDF88B388A3F7DC" type="email">nertata@wsbs-msu.ru</mods:nameIdentifier>
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<mods:title id="BD4C5732292A66E6E8D9A793E11E4DA7">Zootaxa</mods:title>
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<mods:part id="8F919A04F47EDA609EB7DE83558B97FB">
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<mods:date id="92BC11E6959D179C11E7609A40913EBD">2025</mods:date>
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<mods:number id="1EF929005F071285009A4777B6E48BFE">2025-03-26</mods:number>
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<mods:number id="EF5C0054B4E5A73BE6934E26779C0531">5613</mods:number>
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<mods:url id="55144015A74F06F5C38A817761662636">https://doi.org/10.11646/zootaxa.5613.1.3</mods:url>
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<treatment id="03C2E360FF8A7430FF4676C5AB86FAFE" LSID="urn:lsid:plazi:treatment:03C2E360FF8A7430FF4676C5AB86FAFE" httpUri="http://treatment.plazi.org/id/03C2E360FF8A7430FF4676C5AB86FAFE" lastPageId="11" lastPageNumber="93" pageId="4" pageNumber="86">
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<subSubSection id="C37101FDFF8A743FFF4676C5AD71FF1E" box="[151,875,151,178]" pageId="4" pageNumber="86" type="nomenclature">
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<paragraph id="8BD45276FF8A743FFF4676C5AD71FF1E" blockId="4.[151,968,151,250]" box="[151,875,151,178]" pageId="4" pageNumber="86">
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<emphasis id="B91F8E64FF8A743FFF4676C5AD71FF1E" bold="true" box="[151,875,151,178]" pageId="4" pageNumber="86">
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<taxonomicName id="4C6B29F5FF8A743FFF4676C5AFB6FF1D" ID-CoL="KTTY6" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[151,428,151,178]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="86" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
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<emphasis id="B91F8E64FF8A743FFF4676C5AFB6FF1D" bold="true" box="[151,428,151,178]" italics="true" pageId="4" pageNumber="86">Ctenophoricola tzetlini</emphasis>
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</taxonomicName>
|
||||
<taxonomicNameLabel id="A22C331FFF8A743FFE6276CAAC0AFF1D" box="[435,528,152,178]" pageId="4" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
Syomin and Neretina, 2024
|
||||
</emphasis>
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||||
</heading>
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||||
</paragraph>
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urn:lsid:zoobank.org:act:
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<uuid id="FFCD68A3FF8A743FFE7876E9ADD2FF7A" box="[425,968,187,213]" pageId="4" pageNumber="86">BDF30A6F-6900-492F-82D8-FA9E13BE31A4</uuid>
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<paragraph id="8BD45276FF8A743FFF4676B2AEE3FF55" blockId="4.[151,968,151,250]" box="[151,249,224,250]" pageId="4" pageNumber="86">
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<figureCitation id="13504EF3FF8A743FFF4676B2AEE3FF55" box="[151,249,224,250]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="4.[151,250,1632,1656]" captionStartId-1="5.[151,250,1981,2005]" captionStartId-2="6.[151,250,1160,1184]" captionStartId-3="7.[151,250,1498,1522]" captionStartId-4="8.[151,250,746,770]" captionStartId-5="8.[151,250,1761,1785]" captionStartId-6="11.[151,250,866,890]" captionTargetBox-0="[151,1436,778,1609]" captionTargetBox-1="[437,1111,265,1943]" captionTargetBox-2="[151,1435,525,1137]" captionTargetBox-3="[151,1436,181,1474]" captionTargetBox-4="[151,1435,181,722]" captionTargetBox-5="[151,1435,903,1738]" captionTargetBox-6="[151,1435,237,843]" captionTargetId-0="figure-238@4.[151,1436,778,1609]" captionTargetId-1="figure-30@5.[392,1196,181,1957]" captionTargetId-2="figure-171@6.[151,1435,525,1137]" captionTargetId-3="figure-30@7.[151,1436,181,1474]" captionTargetId-4="figure-20@8.[151,1435,181,722]" captionTargetId-5="figure-76@8.[151,1435,903,1738]" captionTargetId-6="figure-52@11.[151,1435,237,843]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="8" captionTargetPageId-6="11" captionText-0="FIGURE 1. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS10257. General view. A, dorsal view; B, ventral view; both figures to similar scale." captionText-1="FIGURE 2. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Dorsal view. Designations: ar, anterior region of the body; pr, posterior region of the body.Arrowheads point at lateral caeca." captionText-2="FIGURE 3. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, dorsal view; B, ventral view. Designations: co, cornea; el, eye lens; ie, internal eye structures; li, inferior lip; lu, upper lip; m, mouth opening; p, palp; pl, lateral papilla of inferior lip; pr, prostomium; all drawings to similar scale." captionText-3="FIGURE 4. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, lateroventral view; B, frontal view; C, dorsal view. Designations as in FIGURE 3; all drawings to similar scale." captionText-4="FIGURE 5. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of anterior region. A, chaetiger 8, acicular chaeta almost completely retracted; B, chaetiger 14, acicular chaeta protruded; C, chaetiger 18, acicular chaeta retracted. Designations: a, aciculum; ac, acicular chaeta; dl, dorsal lobe; chl, chaetigerous lobe; all drawings to similar scale." captionText-5="FIGURE 6. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of posterior region. A, chaetiger 19; B, chaetiger 20; C, chaetiger 28. Designations: a, aciculum; c, caecum; ca, capillaries; dl, dorsal lobe; chl, chaetigerous lobe; vl, ventral lobe; A and B to similar scale." captionText-6="FIGURE 7. Phylogenetic tree based on concatenated 18S and 28S markers. Numbers in regular font above nodes indicate Bayesian posterior probability; numbers in italic under nodes represent Maximum Likelihood bootstrap values; dash indicates unsupported nodes." figureDoi-0="http://doi.org/10.5281/zenodo.15216199" figureDoi-1="http://doi.org/10.5281/zenodo.15216201" figureDoi-2="http://doi.org/10.5281/zenodo.15216203" figureDoi-3="http://doi.org/10.5281/zenodo.15216206" figureDoi-4="http://doi.org/10.5281/zenodo.15216208" figureDoi-5="http://doi.org/10.5281/zenodo.15216210" figureDoi-6="http://doi.org/10.5281/zenodo.15216212" httpUri-0="https://zenodo.org/record/15216199/files/figure.png" httpUri-1="https://zenodo.org/record/15216201/files/figure.png" httpUri-2="https://zenodo.org/record/15216203/files/figure.png" httpUri-3="https://zenodo.org/record/15216206/files/figure.png" httpUri-4="https://zenodo.org/record/15216208/files/figure.png" httpUri-5="https://zenodo.org/record/15216210/files/figure.png" httpUri-6="https://zenodo.org/record/15216212/files/figure.png" pageId="4" pageNumber="86">Figs 1–7</figureCitation>
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</paragraph>
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||||
</subSubSection>
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<subSubSection id="C37101FDFF8A743FFF46777AAB02FDB5" pageId="4" pageNumber="86" type="materials_examined">
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<paragraph id="8BD45276FF8A743FFF46777AAB02FDB5" blockId="4.[151,1437,295,754]" pageId="4" pageNumber="86">
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<emphasis id="B91F8E64FF8A743FFF46777AAF21FEED" box="[151,315,296,322]" italics="true" pageId="4" pageNumber="86">Type material.</emphasis>
|
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<materialsCitation id="3B03582BFF8A743FFE95777AAFEAFE25" collectingDate="2016-12-12" collectionCode="ZMMU" country="Maldives" elevation="10" latitude="-0.79665" location="Indian Ocean" longLatPrecision="1" longitude="72.484" municipality="South" pageId="4" pageNumber="86" specimenCode="WS10257" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54D0ECD4FF8A743FFE95777AAFB6FEED" box="[324,428,296,322]" pageId="4" pageNumber="86" type="holotype">Holotype</typeStatus>
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<collectionCode id="ED7ACAB3FF8A743FFE67777AAC17FEED" box="[438,525,296,322]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="86" type="Museum">ZMMU</collectionCode>
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||||
<specimenCode id="DBCDFA0DFF8A743FFDC77775AC92FEED" box="[534,648,295,322]" collectionCode="ZMMU" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="86" type="Museum">WS10257</specimenCode>
|
||||
,
|
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,
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<locationDeviation id="9A664B54FF8A743FFCEE777AAD9AFEED" box="[831,896,296,322]" pageId="4" pageNumber="86">South</locationDeviation>
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<collectingCountry id="F37C12E6FF8A743FFC5B777AADEFFEED" box="[906,1013,296,322]" name="Maldives" pageId="4" pageNumber="86">Maldives</collectingCountry>
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||||
,
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<geoCoordinate id="EE5F34B1FF8A743FFBD37775AA61FEED" box="[1026,1147,295,322]" degrees="0.79665" direction="south" orientation="latitude" pageId="4" pageNumber="86" precision="1" value="-0.79665">0.79665°S</geoCoordinate>
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,
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<geoCoordinate id="EE5F34B1FF8A743FFB59777AAB0BFEED" box="[1160,1297,296,322]" degrees="72.48400" direction="east" orientation="longitude" pageId="4" pageNumber="86" precision="1" value="72.484">72.48400°E</geoCoordinate>
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||||
; pelagic,
|
||||
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<quantity id="4C93FF93FF8A743FFA51777AAEB7FEC9" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="86" unit="m" value="10.0">10 m</quantity>
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||||
depth
|
||||
</elevation>
|
||||
;
|
||||
<date id="FFD574B6FF8A743FFED5771EAFFBFECA" box="[260,481,331,358]" pageId="4" pageNumber="86" value="2016-12-12">
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<collectingDate id="EF918D5EFF8A743FFED5771EAFFBFECA" box="[260,481,331,358]" pageId="4" pageNumber="86" value="2016-12-12">12 December 2016</collectingDate>
|
||||
</date>
|
||||
. Originally complete adult female with 32 chaetigers, broke in two otherwise intact fragments during examination.
|
||||
</materialsCitation>
|
||||
Body length
|
||||
<quantity id="4C93FF93FF8A743FFD54773DACF3FE25" box="[645,745,367,394]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="86" unit="mm" value="10.5">10.5 mm</quantity>
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(
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<quantity id="4C93FF93FF8A743FFD297722AD5DFE25" box="[760,839,368,394]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="86" unit="mm" value="12.0">12 mm</quantity>
|
||||
with pygidial cirri). Maximum body width excluding parapodia
|
||||
<quantity id="4C93FF93FF8A743FFEDC77C1AF7EFE01" box="[269,356,403,430]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="86" unit="mm" value="0.6">0.6 mm</quantity>
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||||
in anterior region and
|
||||
<quantity id="4C93FF93FF8A743FFDB977C1ACB2FE01" box="[616,680,403,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="86" unit="in" value="0.5">0.5 in</quantity>
|
||||
posterior region; maximum width with parapodia
|
||||
<quantity id="4C93FF93FF8A743FFB3277C6AB3FFE01" box="[1251,1317,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="86" unit="mm" value="3.0">3 mm</quantity>
|
||||
(chaetiger 28); parapodia length up to
|
||||
<quantity id="4C93FF93FF8A743FFE1E77EAAC32FE7D" box="[463,552,440,466]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="86" unit="mm" value="1.2">1.2 mm</quantity>
|
||||
. Proboscis not everted. Body cavity filled with eggs, especially anterior region and parapodia of posterior region. Fixation: 96% ethanol. Parapodium from chaetiger 26 (right side) taken for DNA extraction. Parapodia of chaetigers 8, 14, 18, 19, 20 and 28 (right side) mounted on microscopic slides.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37101FDFF8A743FFF167476AA3BFD5D" pageId="4" pageNumber="86" type="diagnosis">
|
||||
<paragraph id="8BD45276FF8A743FFF167476AA3BFD5D" blockId="4.[151,1437,295,754]" pageId="4" pageNumber="86">
|
||||
<emphasis id="B91F8E64FF8A743FFF167476AFE6FD91" box="[199,508,548,574]" italics="true" pageId="4" pageNumber="86">
|
||||
Diagnosis.
|
||||
<taxonomicName id="4C6B29F5FF8A743FFE9D7476AFE6FD91" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[332,508,548,574]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="86" phylum="Annelida" rank="genus">Ctenophoricola</taxonomicName>
|
||||
</emphasis>
|
||||
with body relatively large and long. Prostomium conspicuous, non-retractable, nearly as wide as peristomium. Eyes with well-developed hemispherical external cornea directed forward. Two ovoid papillae arising from lip on either side of mouth. Anterior region lacks ciliation. Dorsal lobe of parapodium indistinctly conical. Chaetigerous lobe with aciculum and one long acicular chaeta. Posterior region with scattered bundles of cilia. Gut in posterior region with lateral caeca protruding into parapodia up to half the length of the parapodia. Pygidium large with pair of anal cirri longer than posterior parapodia.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF1402FEFF8A743FFF467032AF9EF933" ID-DOI="http://doi.org/10.5281/zenodo.15216199" ID-Zenodo-Dep="15216199" httpUri="https://zenodo.org/record/15216199/files/figure.png" pageId="4" pageNumber="86" startId="4.[151,250,1632,1656]" targetBox="[151,1436,778,1609]" targetPageId="4" targetType="figure">
|
||||
<paragraph id="8BD45276FF8A743FFF467032AF9EF933" blockId="4.[151,1436,1632,1692]" pageId="4" pageNumber="86">
|
||||
<emphasis id="B91F8E64FF8A743FFF467032AF0EF9D7" bold="true" box="[151,276,1632,1656]" pageId="4" pageNumber="86">FIGURE 1.</emphasis>
|
||||
<taxonomicName id="4C6B29F5FF8A743FFECB7032AC1FF9D7" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[282,517,1632,1656]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="4" pageNumber="86" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF8A743FFECB7032AC1FF9D7" box="[282,517,1632,1656]" italics="true" pageId="4" pageNumber="86">Ctenophoricola tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF8A743FFDDA7032AC46F9D7" bold="true" box="[523,604,1632,1656]" pageId="4" pageNumber="86">
|
||||
<taxonomicNameLabel id="A22C331FFF8A743FFDDA7032AC46F9D7" box="[523,604,1632,1656]" pageId="4" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ZMMU WS10257. General view. A, dorsal view; B, ventral view; both figures to similar scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C37101FDFF8A743CFF1670BDAB1DF8F5" lastPageId="7" lastPageNumber="89" pageId="4" pageNumber="86" type="description">
|
||||
<paragraph id="8BD45276FF8A743FFF1670BDAFD2F813" blockId="4.[151,1437,1774,1980]" pageId="4" pageNumber="86">
|
||||
<emphasis id="B91F8E64FF8A743FFF1670BDAFC2F8A7" box="[199,472,1774,1800]" italics="true" pageId="4" pageNumber="86">
|
||||
Description of
|
||||
<typeStatus id="54D0ECD4FF8A743FFEA170BCAFCEF8A7" box="[368,468,1774,1800]" pageId="4" pageNumber="86" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Body relatively long and slender, divided in two distinct regions (
|
||||
<figureCitation id="13504EF3FF8A743FFB1270BCAB30F8A7" box="[1219,1322,1774,1800]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1632,1656]" captionStartId-1="5.[151,250,1981,2005]" captionTargetBox-0="[151,1436,778,1609]" captionTargetBox-1="[437,1111,265,1943]" captionTargetId-0="figure-238@4.[151,1436,778,1609]" captionTargetId-1="figure-30@5.[392,1196,181,1957]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS10257. General view. A, dorsal view; B, ventral view; both figures to similar scale." captionText-1="FIGURE 2. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Dorsal view. Designations: ar, anterior region of the body; pr, posterior region of the body.Arrowheads point at lateral caeca." figureDoi-0="http://doi.org/10.5281/zenodo.15216199" figureDoi-1="http://doi.org/10.5281/zenodo.15216201" httpUri-0="https://zenodo.org/record/15216199/files/figure.png" httpUri-1="https://zenodo.org/record/15216201/files/figure.png" pageId="4" pageNumber="86">Figs 1–2</figureCitation>
|
||||
). Anterior region cylindrical, with reduced parapodia. Posterior region enlarged and flattened due to enormously expanded parapodia. Trunk cylindrical, its diameter smaller than in anterior region. Anterior region together with prostomium and peristomium with few irregularly distributed single cilia. Numerous bundles of short cilia scattered over posterior region with no apparent regularity. Gut straight in anterior region, with pair of lateral caeca per segment in posterior region (
|
||||
<figureCitation id="13504EF3FF8A743FFEA471F0AFA1F813" box="[373,443,1954,1980]" captionStart="FIGURE 2" captionStartId="5.[151,250,1981,2005]" captionTargetBox="[437,1111,265,1943]" captionTargetId="figure-30@5.[392,1196,181,1957]" captionTargetPageId="5" captionText="FIGURE 2. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Dorsal view. Designations: ar, anterior region of the body; pr, posterior region of the body.Arrowheads point at lateral caeca." figureDoi="http://doi.org/10.5281/zenodo.15216201" httpUri="https://zenodo.org/record/15216201/files/figure.png" pageId="4" pageNumber="86">Fig. 2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF1402FEFF8B743EFF4671EFAD6DF856" ID-DOI="http://doi.org/10.5281/zenodo.15216201" ID-Zenodo-Dep="15216201" httpUri="https://zenodo.org/record/15216201/files/figure.png" pageId="5" pageNumber="87" startId="5.[151,250,1981,2005]" targetBox="[437,1111,265,1943]" targetPageId="5" targetType="figure">
|
||||
<paragraph id="8BD45276FF8B743EFF4671EFAD6DF856" blockId="5.[151,1436,1981,2041]" pageId="5" pageNumber="87">
|
||||
<emphasis id="B91F8E64FF8B743EFF4671EFAF09F87A" bold="true" box="[151,275,1981,2005]" pageId="5" pageNumber="87">FIGURE 2.</emphasis>
|
||||
<taxonomicName id="4C6B29F5FF8B743EFEC871EFAC19F87A" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[281,515,1981,2005]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="5" pageNumber="87" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF8B743EFEC871EFAC19F87A" box="[281,515,1981,2005]" italics="true" pageId="5" pageNumber="87">Ctenophoricola tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF8B743EFDD871ECAC43F879" bold="true" box="[521,601,1982,2006]" pageId="5" pageNumber="87">
|
||||
<taxonomicNameLabel id="A22C331FFF8B743EFDD871ECAC43F879" box="[521,601,1982,2006]" pageId="5" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ZMMU WS 10257. Dorsal view. Designations: ar, anterior region of the body; pr, posterior region of the body. Arrowheads point at lateral caeca.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BD45276FF88743DFF1676C5AF1BFE5A" blockId="6.[151,1437,151,501]" pageId="6" pageNumber="88">
|
||||
Prostomium non-retractable, bears a pair of eyes with well-developed hemispherical cornea directed forward (
|
||||
<figureCitation id="13504EF3FF88743DFF4E76E9AEF2FF7A" box="[159,232,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1160,1184]" captionTargetBox="[151,1435,525,1137]" captionTargetId="figure-171@6.[151,1435,525,1137]" captionTargetPageId="6" captionText="FIGURE 3. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, dorsal view; B, ventral view. Designations: co, cornea; el, eye lens; ie, internal eye structures; li, inferior lip; lu, upper lip; m, mouth opening; p, palp; pl, lateral papilla of inferior lip; pr, prostomium; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216203" httpUri="https://zenodo.org/record/15216203/files/figure.png" pageId="6" pageNumber="88">Figs 3</figureCitation>
|
||||
and
|
||||
<figureCitation id="13504EF3FF88743DFEF376E9AF29FF7A" box="[290,307,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1498,1522]" captionTargetBox="[151,1436,181,1474]" captionTargetId="figure-30@7.[151,1436,181,1474]" captionTargetPageId="7" captionText="FIGURE 4. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, lateroventral view; B, frontal view; C, dorsal view. Designations as in FIGURE 3; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216206" httpUri="https://zenodo.org/record/15216206/files/figure.png" pageId="6" pageNumber="88">4</figureCitation>
|
||||
). Eyes with spherical lenses located directly under cornea. Internal eyes including retina elongated and darkly pigmented, reaching boundary of peristomium and first chaetiger in fixed specimen. Prostomium bears pair of small non-ciliated palps (
|
||||
<figureCitation id="13504EF3FF88743DFDD57751AC7BFEB2" box="[516,609,259,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1160,1184]" captionTargetBox="[151,1435,525,1137]" captionTargetId="figure-171@6.[151,1435,525,1137]" captionTargetPageId="6" captionText="FIGURE 3. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, dorsal view; B, ventral view. Designations: co, cornea; el, eye lens; ie, internal eye structures; li, inferior lip; lu, upper lip; m, mouth opening; p, palp; pl, lateral papilla of inferior lip; pr, prostomium; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216203" httpUri="https://zenodo.org/record/15216203/files/figure.png" pageId="6" pageNumber="88">Figs 3B</figureCitation>
|
||||
;
|
||||
<figureCitation id="13504EF3FF88743DFDBC7751ACA9FEB2" box="[621,691,259,285]" captionStart="FIGURE 4" captionStartId="7.[151,250,1498,1522]" captionTargetBox="[151,1436,181,1474]" captionTargetId="figure-30@7.[151,1436,181,1474]" captionTargetPageId="7" captionText="FIGURE 4. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, lateroventral view; B, frontal view; C, dorsal view. Designations as in FIGURE 3; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216206" httpUri="https://zenodo.org/record/15216206/files/figure.png" pageId="6" pageNumber="88">4A, B</figureCitation>
|
||||
). They are possibly glandulous, with dense granular opaque white content. Peristomium similar in length and width to subsequent segments. Tentacular cirri not found (
|
||||
<figureCitation id="13504EF3FF88743DFAE57775AB8AFEEE" box="[1332,1424,295,321]" captionStart="FIGURE 4" captionStartId="7.[151,250,1498,1522]" captionTargetBox="[151,1436,181,1474]" captionTargetId="figure-30@7.[151,1436,181,1474]" captionTargetPageId="7" captionText="FIGURE 4. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, lateroventral view; B, frontal view; C, dorsal view. Designations as in FIGURE 3; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216206" httpUri="https://zenodo.org/record/15216206/files/figure.png" pageId="6" pageNumber="88">Fig. 4C</figureCitation>
|
||||
). Mouth surrounded by cushion-shaped lips (
|
||||
<figureCitation id="13504EF3FF88743DFD5A7719ACFCFECA" box="[651,742,331,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1160,1184]" captionTargetBox="[151,1435,525,1137]" captionTargetId="figure-171@6.[151,1435,525,1137]" captionTargetPageId="6" captionText="FIGURE 3. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, dorsal view; B, ventral view. Designations: co, cornea; el, eye lens; ie, internal eye structures; li, inferior lip; lu, upper lip; m, mouth opening; p, palp; pl, lateral papilla of inferior lip; pr, prostomium; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216203" httpUri="https://zenodo.org/record/15216203/files/figure.png" pageId="6" pageNumber="88">Fig. 3B</figureCitation>
|
||||
;
|
||||
<figureCitation id="13504EF3FF88743DFD247719AD27FECA" box="[757,829,331,357]" captionStart="FIGURE 4" captionStartId="7.[151,250,1498,1522]" captionTargetBox="[151,1436,181,1474]" captionTargetId="figure-30@7.[151,1436,181,1474]" captionTargetPageId="7" captionText="FIGURE 4. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, lateroventral view; B, frontal view; C, dorsal view. Designations as in FIGURE 3; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216206" httpUri="https://zenodo.org/record/15216206/files/figure.png" pageId="6" pageNumber="88">4A, B</figureCitation>
|
||||
). Upper lip partly fused with bases of palps, opaque white. Inferior lip (or a pair of fused ventro-lateral lips) deep horseshoe-shaped, with dense dark pigmentation. On either side of mouth, one papilla emerges from lateral side of inferior lip. Each papilla consists of swollen base and thinner ovoid distal part, together exceeding corresponding palp two times (
|
||||
<figureCitation id="13504EF3FF88743DFBCE77E5AA6DFE7E" box="[1055,1143,439,465]" captionStart="FIGURE 3" captionStartId="6.[151,250,1160,1184]" captionTargetBox="[151,1435,525,1137]" captionTargetId="figure-171@6.[151,1435,525,1137]" captionTargetPageId="6" captionText="FIGURE 3. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, dorsal view; B, ventral view. Designations: co, cornea; el, eye lens; ie, internal eye structures; li, inferior lip; lu, upper lip; m, mouth opening; p, palp; pl, lateral papilla of inferior lip; pr, prostomium; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216203" httpUri="https://zenodo.org/record/15216203/files/figure.png" pageId="6" pageNumber="88">Fig. 3B</figureCitation>
|
||||
). Proboscis not everted in
|
||||
<typeStatus id="54D0ECD4FF88743DFF467789AEE7FE5A" box="[151,253,475,501]" pageId="6" pageNumber="88" type="holotype">holotype</typeStatus>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF1402FEFF88743DFF4672DAAD90FB47" ID-DOI="http://doi.org/10.5281/zenodo.15216203" ID-Zenodo-Dep="15216203" httpUri="https://zenodo.org/record/15216203/files/figure.png" pageId="6" pageNumber="88" startId="6.[151,250,1160,1184]" targetBox="[151,1435,525,1137]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="8BD45276FF88743DFF4672DAAD90FB47" blockId="6.[151,1437,1160,1256]" pageId="6" pageNumber="88">
|
||||
<emphasis id="B91F8E64FF88743DFF4672DAAF0FFB0F" bold="true" box="[151,277,1160,1184]" pageId="6" pageNumber="88">FIGURE 3.</emphasis>
|
||||
Anterior end of
|
||||
<taxonomicName id="4C6B29F5FF88743DFE1472DAACA8FB0F" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[453,690,1160,1184]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="88" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF88743DFE1472DAACA8FB0F" box="[453,690,1160,1184]" italics="true" pageId="6" pageNumber="88">Ctenophoricola tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF88743DFD6B72DAAD17FB0F" bold="true" box="[698,781,1160,1184]" pageId="6" pageNumber="88">
|
||||
<taxonomicNameLabel id="A22C331FFF88743DFD6B72DAAD17FB0F" box="[698,781,1160,1184]" pageId="6" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ZMMU WS 10257. A, dorsal view; B, ventral view. Designations: co, cornea; el, eye lens; ie, internal eye structures; li, inferior lip; lu, upper lip; m, mouth opening; p, palp; pl, lateral papilla of inferior lip; pr, prostomium; all drawings to similar scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BD45276FF88743DFF167368AEAFFA4A" blockId="6.[151,1437,1338,1940]" pageId="6" pageNumber="88">
|
||||
Anterior body region with 18 chaetigers. Parapodia reduced, consisting of acute chaetigerous lobe and dorsal lobe. Chaetigerous lobe conical, with non-protruding aciculum and one long straight acicular chaeta located below aciculum. Acicular chaeta capable of being completely retracted inside parapodium (
|
||||
<figureCitation id="13504EF3FF88743DFB8A73D0AACDFA33" box="[1115,1239,1410,1437]" captionStart="FIGURE 5" captionStartId="8.[151,250,746,770]" captionTargetBox="[151,1435,181,722]" captionTargetId="figure-20@8.[151,1435,181,722]" captionTargetPageId="8" captionText="FIGURE 5. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of anterior region. A, chaetiger 8, acicular chaeta almost completely retracted; B, chaetiger 14, acicular chaeta protruded; C, chaetiger 18, acicular chaeta retracted. Designations: a, aciculum; ac, acicular chaeta; dl, dorsal lobe; chl, chaetigerous lobe; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216208" httpUri="https://zenodo.org/record/15216208/files/figure.png" pageId="6" pageNumber="88">Fig. 5A, C</figureCitation>
|
||||
) or protruded far outside up to at least three lengths of lobe (
|
||||
<figureCitation id="13504EF3FF88743DFD5773F4ACFAFA6E" box="[646,736,1446,1473]" captionStart="FIGURE 5" captionStartId="8.[151,250,746,770]" captionTargetBox="[151,1435,181,722]" captionTargetId="figure-20@8.[151,1435,181,722]" captionTargetPageId="8" captionText="FIGURE 5. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of anterior region. A, chaetiger 8, acicular chaeta almost completely retracted; B, chaetiger 14, acicular chaeta protruded; C, chaetiger 18, acicular chaeta retracted. Designations: a, aciculum; ac, acicular chaeta; dl, dorsal lobe; chl, chaetigerous lobe; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216208" httpUri="https://zenodo.org/record/15216208/files/figure.png" pageId="6" pageNumber="88">Fig. 5B</figureCitation>
|
||||
). Dorsal lobe of indistinct conical shape with wide base (
|
||||
<figureCitation id="13504EF3FF88743DFABE73F4AEBDFA4A" captionStart="FIGURE 5" captionStartId="8.[151,250,746,770]" captionTargetBox="[151,1435,181,722]" captionTargetId="figure-20@8.[151,1435,181,722]" captionTargetPageId="8" captionText="FIGURE 5. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of anterior region. A, chaetiger 8, acicular chaeta almost completely retracted; B, chaetiger 14, acicular chaeta protruded; C, chaetiger 18, acicular chaeta retracted. Designations: a, aciculum; ac, acicular chaeta; dl, dorsal lobe; chl, chaetigerous lobe; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216208" httpUri="https://zenodo.org/record/15216208/files/figure.png" pageId="6" pageNumber="88">Fig. 5</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF88743DFF1673BCACB7F83B" blockId="6.[151,1437,1338,1940]" pageId="6" pageNumber="88">
|
||||
Posterior region with 14 chaetigers. Parapodia are an order of magnitude longer than those of anterior region and exceed corresponding body width up to 2.5 times. Dorsal and ventral lobes fused to chaetigerous lobe and overgrow it forming an expanded oval paddle-like structure with trilobed free end (
|
||||
<figureCitation id="13504EF3FF88743DFC377064AA37F9FF" box="[998,1069,1590,1616]" captionStart="FIGURE 6" captionStartId="8.[151,250,1761,1785]" captionTargetBox="[151,1435,903,1738]" captionTargetId="figure-76@8.[151,1435,903,1738]" captionTargetPageId="8" captionText="FIGURE 6. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of posterior region. A, chaetiger 19; B, chaetiger 20; C, chaetiger 28. Designations: a, aciculum; c, caecum; ca, capillaries; dl, dorsal lobe; chl, chaetigerous lobe; vl, ventral lobe; A and B to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216210" httpUri="https://zenodo.org/record/15216210/files/figure.png" pageId="6" pageNumber="88">Fig. 6</figureCitation>
|
||||
). Longest and widest parapodia located in chaetiger 28 (
|
||||
<figureCitation id="13504EF3FF88743DFE7C7008AC12F9DB" box="[429,520,1626,1652]" captionStart="FIGURE 6" captionStartId="8.[151,250,1761,1785]" captionTargetBox="[151,1435,903,1738]" captionTargetId="figure-76@8.[151,1435,903,1738]" captionTargetPageId="8" captionText="FIGURE 6. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of posterior region. A, chaetiger 19; B, chaetiger 20; C, chaetiger 28. Designations: a, aciculum; c, caecum; ca, capillaries; dl, dorsal lobe; chl, chaetigerous lobe; vl, ventral lobe; A and B to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216210" httpUri="https://zenodo.org/record/15216210/files/figure.png" pageId="6" pageNumber="88">Fig. 6C</figureCitation>
|
||||
). Parapodia decrease in length and become narrower in shape both forward and backward. Lateral caeca follow same pattern: they are short and just enter bases of anterior parapodia of posterior region (
|
||||
<figureCitation id="13504EF3FF88743DFF3C70F0AF5EF913" box="[237,324,1698,1724]" captionStart="FIGURE 6" captionStartId="8.[151,250,1761,1785]" captionTargetBox="[151,1435,903,1738]" captionTargetId="figure-76@8.[151,1435,903,1738]" captionTargetPageId="8" captionText="FIGURE 6. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of posterior region. A, chaetiger 19; B, chaetiger 20; C, chaetiger 28. Designations: a, aciculum; c, caecum; ca, capillaries; dl, dorsal lobe; chl, chaetigerous lobe; vl, ventral lobe; A and B to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216210" httpUri="https://zenodo.org/record/15216210/files/figure.png" pageId="6" pageNumber="88">Fig. 6B</figureCitation>
|
||||
), protrude into parapodia to up to half their length in middle of region (
|
||||
<figureCitation id="13504EF3FF88743DFB8970F0AAB5F913" box="[1112,1199,1698,1724]" captionStart="FIGURE 6" captionStartId="8.[151,250,1761,1785]" captionTargetBox="[151,1435,903,1738]" captionTargetId="figure-76@8.[151,1435,903,1738]" captionTargetPageId="8" captionText="FIGURE 6. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of posterior region. A, chaetiger 19; B, chaetiger 20; C, chaetiger 28. Designations: a, aciculum; c, caecum; ca, capillaries; dl, dorsal lobe; chl, chaetigerous lobe; vl, ventral lobe; A and B to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216210" httpUri="https://zenodo.org/record/15216210/files/figure.png" pageId="6" pageNumber="88">Fig. 6C</figureCitation>
|
||||
), and become shorter again in posteriormost segments. Acicula long and slender, not protruding. Chaetae very fine, short capillaries; they only slightly protrude forming barely visible brush around tip of parapodium. In first and last parapodia of posterior body region, bases of chaetae attached within basal part of parapodium (
|
||||
<figureCitation id="13504EF3FF88743DFC6A715CAA2DF887" box="[955,1079,1806,1832]" captionStart="FIGURE 6" captionStartId="8.[151,250,1761,1785]" captionTargetBox="[151,1435,903,1738]" captionTargetId="figure-76@8.[151,1435,903,1738]" captionTargetPageId="8" captionText="FIGURE 6. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of posterior region. A, chaetiger 19; B, chaetiger 20; C, chaetiger 28. Designations: a, aciculum; c, caecum; ca, capillaries; dl, dorsal lobe; chl, chaetigerous lobe; vl, ventral lobe; A and B to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216210" httpUri="https://zenodo.org/record/15216210/files/figure.png" pageId="6" pageNumber="88">Fig. 6A–B</figureCitation>
|
||||
). In biggest parapodia in middle of region, bases of chaetae moved towards end of parapodium, to around boundary of second and last thirds of its length; main part of internal space of parapodium is occupied by caecum (
|
||||
<figureCitation id="13504EF3FF88743DFC0F7104AA22F8DF" box="[990,1080,1878,1904]" captionStart="FIGURE 6" captionStartId="8.[151,250,1761,1785]" captionTargetBox="[151,1435,903,1738]" captionTargetId="figure-76@8.[151,1435,903,1738]" captionTargetPageId="8" captionText="FIGURE 6. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Parapodia of posterior region. A, chaetiger 19; B, chaetiger 20; C, chaetiger 28. Designations: a, aciculum; c, caecum; ca, capillaries; dl, dorsal lobe; chl, chaetigerous lobe; vl, ventral lobe; A and B to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216210" httpUri="https://zenodo.org/record/15216210/files/figure.png" pageId="6" pageNumber="88">Fig. 6C</figureCitation>
|
||||
). Pygidium large, with tapering cirri longer than posterior parapodia (
|
||||
<figureCitation id="13504EF3FF88743DFDEB7128ACBAF83B" box="[570,672,1914,1940]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,250,1632,1656]" captionStartId-1="5.[151,250,1981,2005]" captionTargetBox-0="[151,1436,778,1609]" captionTargetBox-1="[437,1111,265,1943]" captionTargetId-0="figure-238@4.[151,1436,778,1609]" captionTargetId-1="figure-30@5.[392,1196,181,1957]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS10257. General view. A, dorsal view; B, ventral view; both figures to similar scale." captionText-1="FIGURE 2. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. Dorsal view. Designations: ar, anterior region of the body; pr, posterior region of the body.Arrowheads point at lateral caeca." figureDoi-0="http://doi.org/10.5281/zenodo.15216199" figureDoi-1="http://doi.org/10.5281/zenodo.15216201" httpUri-0="https://zenodo.org/record/15216199/files/figure.png" httpUri-1="https://zenodo.org/record/15216201/files/figure.png" pageId="6" pageNumber="88">Figs 1–2</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<caption id="DF1402FEFF89743CFF467388ADC9F9B9" ID-DOI="http://doi.org/10.5281/zenodo.15216206" ID-Zenodo-Dep="15216206" httpUri="https://zenodo.org/record/15216206/files/figure.png" pageId="7" pageNumber="89" startId="7.[151,250,1498,1522]" targetBox="[151,1436,181,1474]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="8BD45276FF89743CFF467388ADC9F9B9" blockId="7.[151,1437,1498,1558]" pageId="7" pageNumber="89">
|
||||
<emphasis id="B91F8E64FF89743CFF467388AF0FFA5D" bold="true" box="[151,277,1498,1522]" pageId="7" pageNumber="89">FIGURE 4.</emphasis>
|
||||
Anterior end of
|
||||
<taxonomicName id="4C6B29F5FF89743CFE147388ACABFA5D" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[453,689,1498,1522]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="89" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF89743CFE147388ACABFA5D" box="[453,689,1498,1522]" italics="true" pageId="7" pageNumber="89">Ctenophoricola tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF89743CFD6B7388AD16FA5D" bold="true" box="[698,780,1498,1522]" pageId="7" pageNumber="89">
|
||||
<taxonomicNameLabel id="A22C331FFF89743CFD6B7388AD16FA5D" box="[698,780,1498,1522]" pageId="7" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ZMMU WS 10257. A, lateroventral view; B, frontal view; C, dorsal view. Designations as in FIGURE 3; all drawings to similar scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BD45276FF89743CFF16703AAB1DF8F5" blockId="7.[151,1437,1640,1919]" pageId="7" pageNumber="89">
|
||||
Coloration: fixed specimen transparent whitish with opaque mass of eggs filling body (
|
||||
<figureCitation id="13504EF3FF89743CFB40703AAAC8F92D" box="[1169,1234,1640,1666]" captionStart="FIGURE 1" captionStartId="4.[151,250,1632,1656]" captionTargetBox="[151,1436,778,1609]" captionTargetId="figure-238@4.[151,1436,778,1609]" captionTargetPageId="4" captionText="FIGURE 1. Ctenophoricola tzetlini sp. nov., holotype ZMMU WS10257. General view. A, dorsal view; B, ventral view; both figures to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216199" httpUri="https://zenodo.org/record/15216199/files/figure.png" pageId="7" pageNumber="89">Fig. 1</figureCitation>
|
||||
). Palps and upper lip opaque white due to dense granular content. Inferior lip with papillae dark brown. Anterior region of body with little pigmentation: brown longitudinal spot dorsally on prostomium and first three segments (
|
||||
<figureCitation id="13504EF3FF89743CFB7470E2AAE6F965" box="[1189,1276,1712,1738]" captionStart="FIGURE 4" captionStartId="7.[151,250,1498,1522]" captionTargetBox="[151,1436,181,1474]" captionTargetId="figure-30@7.[151,1436,181,1474]" captionTargetPageId="7" captionText="FIGURE 4. Anterior end of Ctenophoricola tzetlini sp. nov., holotype ZMMU WS 10257. A, lateroventral view; B, frontal view; C, dorsal view. Designations as in FIGURE 3; all drawings to similar scale." figureDoi="http://doi.org/10.5281/zenodo.15216206" httpUri="https://zenodo.org/record/15216206/files/figure.png" pageId="7" pageNumber="89">Fig. 4C</figureCitation>
|
||||
), and traces of brownish pigmentation at parapodial bases posteriorly. Posterior region of body with multiple reddish brown spots formed by branching chromatophores. Dorsally, scattered spots cover most of trunk; ventrally, spots form single transverse row in each segment running from one parapodial base to another. Parapodia with bold scattered bands along upper margin of dorsal parapodial lobes and indistinct bands on anterior and posterior surfaces.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37101FDFF897432FF167137AA1DF85A" lastPageId="9" lastPageNumber="91" pageId="7" pageNumber="89" type="biology_ecology">
|
||||
<paragraph id="8BD45276FF89743CFF167137AFF1F8D1" blockId="7.[151,1437,1640,1919]" box="[199,491,1892,1919]" pageId="7" pageNumber="89">Hosts absent or unknown.</paragraph>
|
||||
<caption id="DF1402FEFF867433FF4674B8AB05FCE5" ID-DOI="http://doi.org/10.5281/zenodo.15216208" ID-Zenodo-Dep="15216208" httpUri="https://zenodo.org/record/15216208/files/figure.png" pageId="8" pageNumber="90" startId="8.[151,250,746,770]" targetBox="[151,1435,181,722]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8BD45276FF867433FF4674B8AB05FCE5" blockId="8.[151,1437,746,842]" pageId="8" pageNumber="90">
|
||||
<emphasis id="B91F8E64FF867433FF4674B8AF08FCAC" bold="true" box="[151,274,746,771]" pageId="8" pageNumber="90">FIGURE 5.</emphasis>
|
||||
<taxonomicName id="4C6B29F5FF867433FEC974B8AC18FCAD" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[280,514,746,770]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="90" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF867433FEC974B8AC18FCAD" box="[280,514,746,770]" italics="true" pageId="8" pageNumber="90">Ctenophoricola tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF867433FDD674B9AC4DFCAC" bold="true" box="[519,599,747,771]" pageId="8" pageNumber="90">
|
||||
<taxonomicNameLabel id="A22C331FFF867433FDD674B9AC4DFCAC" box="[519,599,747,771]" pageId="8" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ZMMU WS 10257. Parapodia of anterior region. A, chaetiger 8, acicular chaeta almost completely retracted; B, chaetiger 14, acicular chaeta protruded; C, chaetiger 18, acicular chaeta retracted. Designations: a, aciculum; ac, acicular chaeta; dl, dorsal lobe; chl, chaetigerous lobe; all drawings to similar scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="DF1402FEFF867433FF4670B3AC0FF8EE" ID-DOI="http://doi.org/10.5281/zenodo.15216210" ID-Zenodo-Dep="15216210" httpUri="https://zenodo.org/record/15216210/files/figure.png" pageId="8" pageNumber="90" startId="8.[151,250,1761,1785]" targetBox="[151,1435,903,1738]" targetPageId="8" targetType="figure">
|
||||
<paragraph id="8BD45276FF867433FF4670B3AC0FF8EE" blockId="8.[151,1437,1761,1858]" pageId="8" pageNumber="90">
|
||||
<emphasis id="B91F8E64FF867433FF4670B3AF0EF956" bold="true" box="[151,276,1761,1785]" pageId="8" pageNumber="90">FIGURE 6.</emphasis>
|
||||
<taxonomicName id="4C6B29F5FF867433FECD70B3AC1DF956" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[284,519,1761,1785]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="90" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF867433FECD70B3AC1DF956" box="[284,519,1761,1785]" italics="true" pageId="8" pageNumber="90">Ctenophoricola tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF867433FDDE70B3AC7BF956" bold="true" box="[527,609,1761,1785]" pageId="8" pageNumber="90">
|
||||
<taxonomicNameLabel id="A22C331FFF867433FDDE70B3AC7BF956" box="[527,609,1761,1785]" pageId="8" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, holotype ZMMU WS 10257. Parapodia of posterior region. A, chaetiger 19; B, chaetiger 20; C, chaetiger 28. Designations: a, aciculum; c, caecum; ca, capillaries; dl, dorsal lobe; chl, chaetigerous lobe; vl, ventral lobe; A and B to similar scale.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BD45276FF877432FF1676C5AC40FD95" blockId="9.[151,1437,151,2037]" pageId="9" pageNumber="91">
|
||||
<emphasis id="B91F8E64FF877432FF1676C5AF33FF1E" box="[199,297,151,177]" italics="true" pageId="9" pageNumber="91">Remarks</emphasis>
|
||||
. The original diagnosis of the genus (
|
||||
<bibRefCitation id="EFFA2F87FF877432FD1F76C5ADD0FF1E" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[718,970,151,177]" pageId="9" pageNumber="91" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF877432FC8276C5AD91FF1E" box="[851,907,151,177]" italics="true" pageId="9" pageNumber="91">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
) was amended to include the new species which differs in many points from previously described species, and stress the difference between
|
||||
<taxonomicName id="4C6B29F5FF877432FB3D76E9AB86FF7A" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[1260,1436,187,213]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B91F8E64FF877432FB3D76E9AB86FF7A" box="[1260,1436,187,213]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
</taxonomicName>
|
||||
and free-living genera of
|
||||
<taxonomicName id="4C6B29F5FF877432FE79768DAC0CFF56" box="[424,534,223,249]" class="Polychaeta" family="Phyllodocidae" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="tribe" tribe="Alciopini">Alciopini</taxonomicName>
|
||||
.These amendments were as follows: 1) Body
|
||||
<emphasis id="B91F8E64FF877432FBDB768CAA74FF57" bold="true" box="[1034,1134,222,248]" pageId="9" pageNumber="91">variably</emphasis>
|
||||
covered by cilia (“densely” in the original diagnosis; however, even the description of
|
||||
<taxonomicName id="4C6B29F5FF877432FC957751ADB6FEB3" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[836,940,259,285]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="rousei">
|
||||
<emphasis id="B91F8E64FF877432FC957751ADB6FEB3" box="[836,940,259,285]" italics="true" pageId="9" pageNumber="91">C. rousei</emphasis>
|
||||
</taxonomicName>
|
||||
from the original paper mentions “sparse” rather than “dense” ciliation; 2)
|
||||
<emphasis id="B91F8E64FF877432FDDB7774AB67FEEF" bold="true" box="[522,1405,294,321]" pageId="9" pageNumber="91">Chaetae absent or represented by single acicular chaeta per parapodium</emphasis>
|
||||
in anterior body region (anterior region without chaetae in original diagnoses); 3) Posterior region enlarged due to larger segments
|
||||
<emphasis id="B91F8E64FF877432FE9F773CAC80FE27" bold="true" box="[334,666,366,392]" pageId="9" pageNumber="91">and/or expanded parapodia</emphasis>
|
||||
: added to include the formally undescribed
|
||||
<taxonomicName id="4C6B29F5FF877432FB59773CAB45FE27" box="[1160,1375,366,392]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undetermined">
|
||||
<emphasis id="B91F8E64FF877432FB59773CAB22FE27" box="[1160,1336,366,392]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
sp.
|
||||
</taxonomicName>
|
||||
from the Gulf of
|
||||
<collectingRegion id="49AF9C94FF877432FEF477C0AF8DFE03" box="[293,407,402,428]" country="United States of America" name="California" pageId="9" pageNumber="91">California</collectingRegion>
|
||||
(hereinafter referred to as
|
||||
<taxonomicName id="4C6B29F5FF877432FD0477C0ADDDFE03" box="[725,967,402,428]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF877432FD0477C0AD9FFE03" box="[725,901,402,428]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
) and
|
||||
<taxonomicName id="4C6B29F5FF877432FBDC77C0AB09FE03" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[1037,1299,402,428]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF877432FBDC77C0AB09FE03" box="[1037,1299,402,428]" italics="true" pageId="9" pageNumber="91">Ctenophoricola tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF877432FACF77C0AB66FE03" bold="true" box="[1310,1404,402,428]" pageId="9" pageNumber="91">
|
||||
<taxonomicNameLabel id="A22C331FFF877432FACF77C0AB66FE03" box="[1310,1404,402,428]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
in which segments of the posterior region are not enlarged; 4) Lips surrounding mouth: added because this character is common to all representatives of
|
||||
<taxonomicName id="4C6B29F5FF877432FDFF778BACC4FE5C" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[558,734,473,499]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B91F8E64FF877432FDFF778BACC4FE5C" box="[558,734,473,499]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
</taxonomicName>
|
||||
known so far and differs notably from typical morphology of mouth organs in
|
||||
<taxonomicName id="4C6B29F5FF877432FE8277AFAFDAFDB8" box="[339,448,509,535]" class="Polychaeta" family="Phyllodocidae" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="tribe" tribe="Alciopini">Alciopini</taxonomicName>
|
||||
; 5) Eyes extending through
|
||||
<emphasis id="B91F8E64FF877432FD2C77AFAD3DFDB8" bold="true" box="[765,807,509,535]" pageId="9" pageNumber="91">one</emphasis>
|
||||
to several anterior segments (extending through several anterior segments in original diagnosis).
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF877432FF167416AD82FC0F" blockId="9.[151,1437,151,2037]" pageId="9" pageNumber="91">
|
||||
Body of our specimen is considerably longer and comparatively more slender than in other
|
||||
<taxonomicName id="4C6B29F5FF877432FB3D7416AEF1FD2D" pageId="9" pageNumber="91">
|
||||
<emphasis id="B91F8E64FF877432FB3D7416AB86FDF1" box="[1260,1436,580,606]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
, which is illustrated by the following measurements. Its overall length (
|
||||
<quantity id="4C93FF93FF877432FBD4743AAA71FD2C" box="[1029,1131,616,643]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="9" pageNumber="91" unit="mm" value="10.5">10.5 mm</quantity>
|
||||
,
|
||||
<quantity id="4C93FF93FF877432FBA7743AAADEFD2C" box="[1142,1220,616,643]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="91" unit="mm" value="12.0">12 mm</quantity>
|
||||
with pygidial cirri) exceeds 3.5 times the maximum lengths of valid species, and 2.1 times the maximum length of
|
||||
<taxonomicName id="4C6B29F5FF877432FB1474DEAEB0FD65" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF877432FB1474DEAB6FFD09" box="[1221,1397,652,678]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
(
|
||||
<tableCitation id="C6E967CDFF877432FF6874FDAF0AFD65" box="[185,272,687,714]" captionStart="TABLE 3" captionStartId="10.[152,245,151,175]" captionTargetPageId="10" captionText="TABLE 3. Comparison of morphological differences between known Ctenophoricola species. Previously described species characterized according to (San Martín et al. 2021). Designations: na, not available; ~, character is not specified in the description, but an approximate estimate is taken from the figure." pageId="9" pageNumber="91">Table 3</tableCitation>
|
||||
). Length to maximum width ratios in anterior and posterior body regions of
|
||||
<taxonomicName id="4C6B29F5FF877432FBBD74E2AAC3FD66" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[1132,1241,687,714]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF877432FBBD74E2AAC3FD66" box="[1132,1241,687,714]" italics="true" pageId="9" pageNumber="91">C. tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF877432FB3074E2AB20FD65" bold="true" box="[1249,1338,688,714]" pageId="9" pageNumber="91">
|
||||
<taxonomicNameLabel id="A22C331FFF877432FB3074E2AB20FD65" box="[1249,1338,688,714]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
are 17.5 and 21, respectively. Meanwhile, the same ratios are around 3 and 4 times smaller, respectively, in both
|
||||
<taxonomicName id="4C6B29F5FF877432FADA7486AB87FD42" box="[1291,1437,724,750]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="masanorii">
|
||||
<emphasis id="B91F8E64FF877432FADA7486AB87FD42" box="[1291,1437,724,750]" italics="true" pageId="9" pageNumber="91">C. masanorii</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C6B29F5FF877432FF1674A5AF36FCBE" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[199,300,759,785]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="rousei">
|
||||
<emphasis id="B91F8E64FF877432FF1674A5AF36FCBE" box="[199,300,759,785]" italics="true" pageId="9" pageNumber="91">C. rousei</emphasis>
|
||||
</taxonomicName>
|
||||
. Width excluding parapodia is not provided for
|
||||
<taxonomicName id="4C6B29F5FF877432FC9674A5AA21FCBE" box="[839,1083,759,785]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF877432FC9674A5ADEDFCBE" box="[839,1015,759,785]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
, but measurements of its image from the original paper produce corresponding ratios ~ 9 and ~ 8, which is 2 and 2.5 times smaller, respectively. Body width to maximum parapodia length ratio in posterior region equals
|
||||
<quantity id="4C93FF93FF877432FC0A756DAA01FCF7" box="[987,1051,830,857]" metricMagnitude="-3" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="91" unit="in" value="0.3">0.3 in</quantity>
|
||||
<taxonomicName id="4C6B29F5FF877432FBF3756DAA95FCF7" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[1058,1167,830,857]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF877432FBF3756DAA95FCF7" box="[1058,1167,830,857]" italics="true" pageId="9" pageNumber="91">C. tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22C331FFF877432FB47756DAAF7FCF6" box="[1174,1261,831,857]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
This parameter could only be measured from figures in other species; it was only slightly bigger in
|
||||
<taxonomicName id="4C6B29F5FF877432FB817530AB5BFCD3" box="[1104,1345,866,893]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF877432FB817530AB1AFCD3" box="[1104,1280,866,892]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
(~ 0.5), but exceeded
|
||||
<taxonomicName id="4C6B29F5FF877432FEE075D4AF84FC0F" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[305,414,902,928]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF877432FEE075D4AF84FC0F" box="[305,414,902,928]" italics="true" pageId="9" pageNumber="91">C. tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22C331FFF877432FE7475D4AFE6FC0F" box="[421,508,902,928]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
3–6 times in other species (
|
||||
<tableCitation id="C6E967CDFF877432FCE275D4AD90FC0F" box="[819,906,902,928]" captionStart="TABLE 3" captionStartId="10.[152,245,151,175]" captionTargetPageId="10" captionText="TABLE 3. Comparison of morphological differences between known Ctenophoricola species. Previously described species characterized according to (San Martín et al. 2021). Designations: na, not available; ~, character is not specified in the description, but an approximate estimate is taken from the figure." pageId="9" pageNumber="91">Table 3</tableCitation>
|
||||
).
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF877432FF1675F8AA4DFB80" blockId="9.[151,1437,151,2037]" pageId="9" pageNumber="91">
|
||||
Ciliary cover varies considerably between different
|
||||
<taxonomicName id="4C6B29F5FF877432FC9575F8AA46FC6B" box="[836,1116,938,964]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undetermined">
|
||||
<emphasis id="B91F8E64FF877432FC9575F8ADEEFC6B" box="[836,1012,938,964]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
species.
|
||||
</taxonomicName>
|
||||
Whereas
|
||||
<taxonomicName id="4C6B29F5FF877432FB0F75F8AB60FC6C" box="[1246,1402,938,964]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="masanorii">
|
||||
<emphasis id="B91F8E64FF877432FB0F75F8AB60FC6C" box="[1246,1402,938,964]" italics="true" pageId="9" pageNumber="91">C. masanorii</emphasis>
|
||||
</taxonomicName>
|
||||
is “densely covered by short cilia”, body of
|
||||
<taxonomicName id="4C6B29F5FF877432FDB8759CACD5FC48" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[617,719,974,1000]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="rousei">
|
||||
<emphasis id="B91F8E64FF877432FDB8759CACD5FC48" box="[617,719,974,1000]" italics="true" pageId="9" pageNumber="91">C. rousei</emphasis>
|
||||
</taxonomicName>
|
||||
is “sparsely ciliate” (
|
||||
<bibRefCitation id="EFFA2F87FF877432FC6F759FAAA4FC48" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[958,1214,973,999]" pageId="9" pageNumber="91" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF877432FB92759CAA66FC48" box="[1091,1148,973,999]" italics="true" pageId="9" pageNumber="91">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
). Information about
|
||||
<taxonomicName id="4C6B29F5FF877432FF4675A3AF93FBA4" box="[151,393,1009,1036]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF877432FF4675A3AF5DFBA4" box="[151,327,1009,1035]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
is unavailable. In
|
||||
<taxonomicName id="4C6B29F5FF877432FD8D75A3ACD0FBA4" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[604,714,1009,1035]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF877432FD8D75A3ACD0FBA4" box="[604,714,1009,1035]" italics="true" pageId="9" pageNumber="91">C. tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF877432FD0375A3AD37FBA4" bold="true" box="[722,813,1009,1035]" pageId="9" pageNumber="91">
|
||||
<taxonomicNameLabel id="A22C331FFF877432FD0375A3AD37FBA4" box="[722,813,1009,1035]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
, ciliation differs between body regions. While anterior region has quite a few isolated cilia, posterior region bears numerous bundles of cilia.
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF877432FF16726BAF48FB35" blockId="9.[151,1437,151,2037]" pageId="9" pageNumber="91">
|
||||
Lower lips surrounding mouth are described in all
|
||||
<taxonomicName id="4C6B29F5FF877432FD28726AAA19FBFD" box="[761,1027,1080,1106]" pageId="9" pageNumber="91">
|
||||
<emphasis id="B91F8E64FF877432FD28726AADB3FBFD" box="[761,937,1080,1106]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
; however, no papillae or appendages on lips are mentioned by (
|
||||
<bibRefCitation id="EFFA2F87FF877432FE6C720EACDAFBD9" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[445,704,1116,1142]" pageId="9" pageNumber="91" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF877432FD95720FAC66FBD9" box="[580,636,1116,1142]" italics="true" pageId="9" pageNumber="91">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
). Contrary to that, two papillae exceeding palps in size are found in our specimen.
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF877432FF1672F6AD34FA86" blockId="9.[151,1437,151,2037]" pageId="9" pageNumber="91">
|
||||
Lateral caeca are absent in
|
||||
<emphasis id="B91F8E64FF877432FE2B72F6AC8EFB11" box="[506,660,1188,1214]" italics="true" pageId="9" pageNumber="91">
|
||||
<taxonomicName id="4C6B29F5FF877432FE2B72F6AC8AFB11" box="[506,656,1188,1214]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="masanorii">C. masanorii</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
“sometimes protrude into parapodia” by at least the width of the gut in
|
||||
<emphasis id="B91F8E64FF877432FF65729AAF3BFB4E" box="[180,289,1224,1250]" italics="true" pageId="9" pageNumber="91">
|
||||
<taxonomicName id="4C6B29F5FF877432FF65729AAF07FB4E" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[180,285,1224,1250]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="rousei">C. rousei</taxonomicName>
|
||||
,
|
||||
</emphasis>
|
||||
and not described in
|
||||
<taxonomicName id="4C6B29F5FF877432FDC37295AD1BFB4D" box="[530,769,1223,1250]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF877432FDC37295ACD8FB4E" box="[530,706,1223,1249]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
(
|
||||
<bibRefCitation id="EFFA2F87FF877432FCC1729AAA0BFB4D" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[784,1041,1223,1250]" pageId="9" pageNumber="91" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF877432FC47729AADD4FB4E" box="[918,974,1223,1249]" italics="true" pageId="9" pageNumber="91">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
). In our specimen, the lateral caeca consistently extend into the parapodia: they protrude into almost all parapodia of the posterior region, extending to half the length of the parapodia in the middle of the region.
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF877432FF167361AD39F98C" blockId="9.[151,1437,151,2037]" pageId="9" pageNumber="91">
|
||||
All previously described
|
||||
<taxonomicName id="4C6B29F5FF877432FE347361ACF4FAE2" box="[485,750,1331,1357]" pageId="9" pageNumber="91">
|
||||
<emphasis id="B91F8E64FF877432FE347361AC8FFAE2" box="[485,661,1331,1357]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
do not possess chaetae in parapodia of anterior region; only acicula are present (
|
||||
<bibRefCitation id="EFFA2F87FF877432FE507305AC8AFADE" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[385,656,1366,1393]" pageId="9" pageNumber="91" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF877432FDDF7305AC53FADF" box="[526,585,1366,1392]" italics="true" pageId="9" pageNumber="91">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
). On the contrary, our specimen possesses long retractable acicular chaetae located below acicula. Capillary chaetae of the posterior region of our specimen are extremely short and fine; they just show up their tips at the end of a parapodial lobe. Contrary to that, in all three species described and depicted by
|
||||
<bibRefCitation id="EFFA2F87FF877432FEF37390AC2FFA73" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[290,565,1474,1500]" pageId="9" pageNumber="91" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF877432FE767390AFFAFA73" box="[423,480,1474,1500]" italics="true" pageId="9" pageNumber="91">et al.</emphasis>
|
||||
(2021)
|
||||
</bibRefCitation>
|
||||
, capillaries form a well-developed fan or fascicle. However, we are not quite confident with this character in our specimen, since this species is capable of retracting chaetae to a great extent (observed for acicular chaetae in its anterior body region).
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF877432FF16707FAEC2F921" blockId="9.[151,1437,151,2037]" pageId="9" pageNumber="91">
|
||||
Pygidium is “small with two rounded, short anal cirri” both in
|
||||
<taxonomicName id="4C6B29F5FF877432FCB8707FADE3F9E8" box="[873,1017,1581,1607]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="masanorii">
|
||||
<emphasis id="B91F8E64FF877432FCB8707FADE3F9E8" box="[873,1017,1581,1607]" italics="true" pageId="9" pageNumber="91">C. masanorii</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C6B29F5FF877432FBFB707FAB8FF9E8" authority="(San Martin et al. 2021)" baseAuthorityName="San Martin" baseAuthorityYear="2021" box="[1066,1429,1581,1607]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="rousei">
|
||||
<emphasis id="B91F8E64FF877432FBFB707FAA97F9E8" box="[1066,1165,1581,1607]" italics="true" pageId="9" pageNumber="91">C. rousei</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFFA2F87FF877432FB48707FAB97F9E8" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[1177,1421,1581,1607]" pageId="9" pageNumber="91" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF877432FACB707CAB4AF9E8" box="[1306,1360,1581,1607]" italics="true" pageId="9" pageNumber="91">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. Our specimen has enlarged pygidium with a pair of long anal cirri, a feature which it shares with
|
||||
<taxonomicName id="4C6B29F5FF877432FB3D7002AECEF921" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF877432FB3D7002AB86F9C5" box="[1260,1436,1616,1642]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF877432FF1670CAAA1DF85A" blockId="9.[151,1437,151,2037]" pageId="9" pageNumber="91">
|
||||
The most striking difference between
|
||||
<taxonomicName id="4C6B29F5FF877432FDA270CAACFAF91D" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[627,736,1688,1714]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF877432FDA270CAACFAF91D" box="[627,736,1688,1714]" italics="true" pageId="9" pageNumber="91">C. tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF877432FD3970CAAD58F91D" bold="true" box="[744,834,1688,1714]" pageId="9" pageNumber="91">
|
||||
<taxonomicNameLabel id="A22C331FFF877432FD3970CAAD58F91D" box="[744,834,1688,1714]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
and two described
|
||||
<taxonomicName id="4C6B29F5FF877432FBCF70CAAB3DF91D" box="[1054,1319,1688,1714]" pageId="9" pageNumber="91">
|
||||
<emphasis id="B91F8E64FF877432FBCF70CAAAD4F91D" box="[1054,1230,1688,1714]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
is that the eyes of the new species possess well-developed rigid cornea occupying the anterodorsal side of the prostomium, making the latter wide and presumably non-retractable. Contrary to that, prostomium is “small, semicircular, without external eyes” (
|
||||
<bibRefCitation id="EFFA2F87FF877432FE9B7151AC48F8B2" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[330,594,1795,1821]" pageId="9" pageNumber="91" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF877432FE047156AC0AF8B2" box="[469,528,1795,1821]" italics="true" pageId="9" pageNumber="91">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
) in both
|
||||
<taxonomicName id="4C6B29F5FF877432FD6F7156AD4EF8B2" box="[702,852,1796,1822]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="masanorii">
|
||||
<emphasis id="B91F8E64FF877432FD6F7156AD4EF8B2" box="[702,852,1796,1822]" italics="true" pageId="9" pageNumber="91">C. masanorii</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C6B29F5FF877432FC5E7156ADEDF8B2" box="[911,1015,1796,1822]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="rousei">
|
||||
<emphasis id="B91F8E64FF877432FC5E7156ADEDF8B2" box="[911,1015,1796,1822]" italics="true" pageId="9" pageNumber="91">C. rousei</emphasis>
|
||||
</taxonomicName>
|
||||
. Internal eyes in these species reach posteriorly chaetigers
|
||||
<quantity id="4C93FF93FF877432FE5F7175AFFDF8EE" box="[398,487,1831,1858]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" metricValueMax="1.27" metricValueMin="1.016" pageId="9" pageNumber="91" unit="in" value="4.5" valueMax="5.0" valueMin="4.0">4 to 5 in</quantity>
|
||||
fixed specimens (
|
||||
<quantity id="4C93FF93FF877432FD7E7175AD10F8EE" box="[687,778,1831,1857]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="9" pageNumber="91" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">
|
||||
2 to 4
|
||||
<emphasis id="B91F8E64FF877432FD25717AAD10F8EE" box="[756,778,1832,1857]" italics="true" pageId="9" pageNumber="91">in</emphasis>
|
||||
</quantity>
|
||||
vivo) (
|
||||
<bibRefCitation id="EFFA2F87FF877432FC847175AA4BF8EE" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[853,1105,1831,1857]" pageId="9" pageNumber="91" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF877432FC08717AAA0AF8EE" box="[985,1040,1831,1857]" italics="true" pageId="9" pageNumber="91">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
). The supporting video shows that the eyes move forward and backward inside the worm’s body. Apparently, this is impossible for
|
||||
<taxonomicName id="4C6B29F5FF877432FAFD7119AB86F8CA" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[1324,1436,1867,1893]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF877432FAFD7119AB86F8CA" box="[1324,1436,1867,1893]" italics="true" pageId="9" pageNumber="91">C. tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF877432FF46713DAEE9F826" bold="true" box="[151,243,1903,1929]" pageId="9" pageNumber="91">
|
||||
<taxonomicNameLabel id="A22C331FFF877432FF46713DAEE9F826" box="[151,243,1903,1929]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
in which internal portions of eyes are attached to rigid external cornea and extend only to the boundary between peristomium and the first parapodial segment. The description of
|
||||
<taxonomicName id="4C6B29F5FF877432FC0771C1AADFF802" box="[982,1221,1939,1965]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF877432FC0771C1AA9CF802" box="[982,1158,1939,1965]" italics="true" pageId="9" pageNumber="91">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
lacks these details; but judging from the figure, its internal eyes reach the middle of the first chaetiger posteriorly and adjoin to some external elevations anteriorly. This makes it more similar to
|
||||
<taxonomicName id="4C6B29F5FF877432FCE87189ADBCF85A" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[825,934,2011,2037]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="91" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF877432FCE87189ADBCF85A" box="[825,934,2011,2037]" italics="true" pageId="9" pageNumber="91">C. tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF877432FC7C7189AA1DF85A" bold="true" box="[941,1031,2011,2037]" pageId="9" pageNumber="91">
|
||||
<taxonomicNameLabel id="A22C331FFF877432FC7C7189AA1DF85A" box="[941,1031,2011,2037]" pageId="9" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF1402FEFF847431FF4976C5AD8EFF58" pageId="10" pageNumber="92" startId="10.[152,245,151,175]" targetBox="[163,1424,263,1579]" targetIsTable="true" targetPageId="10" targetType="table">
|
||||
<paragraph id="8BD45276FF847431FF4976C5AD8EFF58" blockId="10.[151,1436,151,247]" pageId="10" pageNumber="92">
|
||||
<emphasis id="B91F8E64FF847431FF4976C5AF09FF00" bold="true" box="[152,275,151,175]" pageId="10" pageNumber="92">TABLE 3.</emphasis>
|
||||
Comparison of morphological differences between known
|
||||
<taxonomicName id="4C6B29F5FF847431FC7876C5AAB5FF00" box="[937,1199,151,175]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="undetermined">
|
||||
<emphasis id="B91F8E64FF847431FC7876C5AA4AFF00" box="[937,1104,151,175]" italics="true" pageId="10" pageNumber="92">Ctenophoricola</emphasis>
|
||||
species.
|
||||
</taxonomicName>
|
||||
Previously described species characterized according to (
|
||||
<bibRefCitation id="EFFA2F87FF847431FDC576EEAD11FF7B" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[532,779,187,212]" pageId="10" pageNumber="92" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF847431FD4576EEACD0FF7C" box="[660,714,187,211]" italics="true" pageId="10" pageNumber="92">et al.</emphasis>
|
||||
2021
|
||||
</bibRefCitation>
|
||||
). Designations: na, not available; ~, character is not specified in the description, but an approximate estimate is taken from the figure.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="8BD45276FF847431FF727755AB59F984" pageId="10" pageNumber="92">
|
||||
<table id="F96BA0D6FF848BC4FF727755AB8AF984" box="[163,1424,263,1579]" gridcols="5" gridrows="17" pageId="10" pageNumber="92">
|
||||
<tr id="355B5034FF848BC4FF727755AB8AFEB0" box="[163,1424,263,287]" gridrow="0" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF727755AF9AFEB0" box="[163,384,263,287]" gridcol="0" gridrow="0" pageId="10" pageNumber="92">
|
||||
<emphasis id="B91F8E64FF847431FF727755AF08FEB0" bold="true" box="[163,274,263,287]" pageId="10" pageNumber="92">Character</emphasis>
|
||||
</th>
|
||||
<th id="768A3948FF848BC4FE487755AC5FFEB0" box="[409,581,263,287]" gridcol="1" gridrow="0" pageId="10" pageNumber="92">
|
||||
<taxonomicName id="4C6B29F5FF847431FE7D7755AC29FEB0" box="[428,563,263,287]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="masanorii">
|
||||
<emphasis id="B91F8E64FF847431FE7D7755AC29FEB0" bold="true" box="[428,563,263,287]" italics="true" pageId="10" pageNumber="92">C. masanorii</emphasis>
|
||||
</taxonomicName>
|
||||
</th>
|
||||
<th id="768A3948FF848BC4FD8D7755AD2CFEB0" box="[604,822,263,287]" gridcol="2" gridrow="0" pageId="10" pageNumber="92">
|
||||
<taxonomicName id="4C6B29F5FF847431FD4B7755ACE2FEB0" box="[666,760,263,287]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="rousei">
|
||||
<emphasis id="B91F8E64FF847431FD4B7755ACE2FEB0" bold="true" box="[666,760,263,287]" italics="true" pageId="10" pageNumber="92">C. rousei</emphasis>
|
||||
</taxonomicName>
|
||||
</th>
|
||||
<th id="768A3948FF848BC4FC9E7755AA2BFEB0" box="[847,1073,263,287]" gridcol="3" gridrow="0" pageId="10" pageNumber="92">
|
||||
<taxonomicName id="4C6B29F5FF847431FC837755AA35FEB0" box="[850,1071,263,287]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF847431FC837755AA35FEB0" bold="true" box="[850,1071,263,287]" pageId="10" pageNumber="92">
|
||||
<emphasis id="B91F8E64FF847431FC837755ADEEFEB0" bold="true" box="[850,1012,263,287]" italics="true" pageId="10" pageNumber="92">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</th>
|
||||
<th id="768A3948FF848BC4FB9B7755AB8AFEB0" box="[1098,1424,263,287]" gridcol="4" gridrow="0" pageId="10" pageNumber="92">
|
||||
<emphasis id="B91F8E64FF847431FB9B7755AB8AFEB0" bold="true" box="[1098,1424,263,287]" pageId="10" pageNumber="92">
|
||||
<taxonomicName id="4C6B29F5FF847431FB9B7755AB22FEB0" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[1098,1336,263,287]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF847431FB9B7755AB22FEB0" bold="true" box="[1098,1336,263,287]" italics="true" pageId="10" pageNumber="92">Ctenophoricola tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="A22C331FFF847431FAEE7755AB8AFEB0" box="[1343,1424,263,287]" pageId="10" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF727762AB8AFEC3" box="[163,1424,304,364]" gridrow="1" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF727762AF9AFEC3" box="[163,384,304,364]" gridcol="0" gridrow="1" pageId="10" pageNumber="92">maximum body length, mm</th>
|
||||
<td id="768A3948FF848BC4FE487762AC5FFEC3" box="[409,581,304,364]" gridcol="1" gridrow="1" pageId="10" pageNumber="92">3</td>
|
||||
<td id="768A3948FF848BC4FD8D7762AD2CFEC3" box="[604,822,304,364]" gridcol="2" gridrow="1" pageId="10" pageNumber="92">3</td>
|
||||
<td id="768A3948FF848BC4FC9E7762AA2BFEC3" box="[847,1073,304,364]" gridcol="3" gridrow="1" pageId="10" pageNumber="92">5</td>
|
||||
<td id="768A3948FF848BC4FB9B7762AB8AFEC3" box="[1098,1424,304,364]" gridcol="4" gridrow="1" pageId="10" pageNumber="92">10.5</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF72772EAB8AFE17" box="[163,1424,380,440]" gridrow="2" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF72772EAB8AFE17" box="[163,1424,380,440]" colspan="5" colspanRight="4" gridcol="0" gridrow="2" pageId="10" pageNumber="92">length to maximum width ratio:</th>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF72779AAB8AFDAB" box="[163,1424,456,516]" gridrow="3" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF72779AAF9AFDAB" box="[163,384,456,516]" gridcol="0" gridrow="3" pageId="10" pageNumber="92">anterior region</th>
|
||||
<td id="768A3948FF848BC4FE48779AAC5FFDAB" box="[409,581,456,516]" gridcol="1" gridrow="3" pageId="10" pageNumber="92">7.5</td>
|
||||
<td id="768A3948FF848BC4FD8D779AAD2CFDAB" box="[604,822,456,516]" gridcol="2" gridrow="3" pageId="10" pageNumber="92">6</td>
|
||||
<td id="768A3948FF848BC4FC9E779AAA2BFDAB" box="[847,1073,456,516]" gridcol="3" gridrow="3" pageId="10" pageNumber="92">~9</td>
|
||||
<td id="768A3948FF848BC4FB9B779AAB8AFDAB" box="[1098,1424,456,516]" gridcol="4" gridrow="3" pageId="10" pageNumber="92">17.5</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF727446AB8AFDFF" box="[163,1424,532,592]" gridrow="4" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF727446AF9AFDFF" box="[163,384,532,592]" gridcol="0" gridrow="4" pageId="10" pageNumber="92">posterior region</th>
|
||||
<td id="768A3948FF848BC4FE487446AC5FFDFF" box="[409,581,532,592]" gridcol="1" gridrow="4" pageId="10" pageNumber="92">5</td>
|
||||
<td id="768A3948FF848BC4FD8D7446AD2CFDFF" box="[604,822,532,592]" gridcol="2" gridrow="4" pageId="10" pageNumber="92">4.6</td>
|
||||
<td id="768A3948FF848BC4FC9E7446AA2BFDFF" box="[847,1073,532,592]" gridcol="3" gridrow="4" pageId="10" pageNumber="92">~8</td>
|
||||
<td id="768A3948FF848BC4FB9B7446AB8AFDFF" box="[1098,1424,532,592]" gridcol="4" gridrow="4" pageId="10" pageNumber="92">21</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF727433AB8AFD6E" box="[163,1424,609,705]" gridrow="5" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF727433AF9AFD6E" box="[163,384,609,705]" gridcol="0" gridrow="5" pageId="10" pageNumber="92">width to parapodial length ratio (posterior region)</th>
|
||||
<td id="768A3948FF848BC4FE487433AC5FFD6E" box="[409,581,609,705]" gridcol="1" gridrow="5" pageId="10" pageNumber="92">~2</td>
|
||||
<td id="768A3948FF848BC4FD8D7433AD2CFD6E" box="[604,822,609,705]" gridcol="2" gridrow="5" pageId="10" pageNumber="92">~1</td>
|
||||
<td id="768A3948FF848BC4FC9E7433AA2BFD6E" box="[847,1073,609,705]" gridcol="3" gridrow="5" pageId="10" pageNumber="92">~0.5</td>
|
||||
<td id="768A3948FF848BC4FB9B7433AB8AFD6E" box="[1098,1424,609,705]" gridcol="4" gridrow="5" pageId="10" pageNumber="92">0.3</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF727483AB8AFCA2" box="[163,1424,721,781]" gridrow="6" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF727483AB8AFCA2" box="[163,1424,721,781]" colspan="5" colspanRight="4" gridcol="0" gridrow="6" pageId="10" pageNumber="92">expanded posterior region:</th>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF72754FAB8AFC99" box="[163,1424,797,822]" gridrow="7" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF72754FAF9AFC99" box="[163,384,797,822]" gridcol="0" gridrow="7" pageId="10" pageNumber="92">segments</th>
|
||||
<td id="768A3948FF848BC4FE48754FAC5FFC99" box="[409,581,797,822]" gridcol="1" gridrow="7" pageId="10" pageNumber="92">enlarged</td>
|
||||
<td id="768A3948FF848BC4FD8D754FAD2CFC99" box="[604,822,797,822]" gridcol="2" gridrow="7" pageId="10" pageNumber="92">slightly enlarged</td>
|
||||
<td id="768A3948FF848BC4FC9E754FAA2BFC99" box="[847,1073,797,822]" gridcol="3" gridrow="7" pageId="10" pageNumber="92">~not enlarged</td>
|
||||
<td id="768A3948FF848BC4FB9B754FAB8AFC99" box="[1098,1424,797,822]" gridcol="4" gridrow="7" pageId="10" pageNumber="92">not enlarged</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF727517AB8AFCF2" box="[163,1424,837,861]" gridrow="8" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF727517AF9AFCF2" box="[163,384,837,861]" gridcol="0" gridrow="8" pageId="10" pageNumber="92">parapodia</th>
|
||||
<td id="768A3948FF848BC4FE487517AC5FFCF2" box="[409,581,837,861]" gridcol="1" gridrow="8" pageId="10" pageNumber="92">enlarged</td>
|
||||
<td id="768A3948FF848BC4FD8D7517AD2CFCF2" box="[604,822,837,861]" gridcol="2" gridrow="8" pageId="10" pageNumber="92">enlarged</td>
|
||||
<td id="768A3948FF848BC4FC9E7517AA2BFCF2" box="[847,1073,837,861]" gridcol="3" gridrow="8" pageId="10" pageNumber="92">~greatly enlarged</td>
|
||||
<td id="768A3948FF848BC4FB9B7517AB8AFCF2" box="[1098,1424,837,861]" gridcol="4" gridrow="8" pageId="10" pageNumber="92">greatly enlarged</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF72753FAB8AFC06" box="[163,1424,877,937]" gridrow="9" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF72753FAF9AFC06" box="[163,384,877,937]" gridcol="0" gridrow="9" pageId="10" pageNumber="92">ciliary cover</th>
|
||||
<td id="768A3948FF848BC4FE48753FAC5FFC06" box="[409,581,877,937]" gridcol="1" gridrow="9" pageId="10" pageNumber="92">dense</td>
|
||||
<td id="768A3948FF848BC4FD8D753FAD2CFC06" box="[604,822,877,937]" gridcol="2" gridrow="9" pageId="10" pageNumber="92">sparse</td>
|
||||
<td id="768A3948FF848BC4FC9E753FAA2BFC06" box="[847,1073,877,937]" gridcol="3" gridrow="9" pageId="10" pageNumber="92">na</td>
|
||||
<td id="768A3948FF848BC4FB9B753FAB8AFC06" box="[1098,1424,877,937]" gridcol="4" gridrow="9" pageId="10" pageNumber="92">sparse anteriorly, bundles of cilia posteriorly</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF7275E8AB8AFC7D" box="[163,1424,954,978]" gridrow="10" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF7275E8AF9AFC7D" box="[163,384,954,978]" gridcol="0" gridrow="10" pageId="10" pageNumber="92">lip papillae</th>
|
||||
<td id="768A3948FF848BC4FE4875E8AC5FFC7D" box="[409,581,954,978]" gridcol="1" gridrow="10" pageId="10" pageNumber="92">absent</td>
|
||||
<td id="768A3948FF848BC4FD8D75E8AD2CFC7D" box="[604,822,954,978]" gridcol="2" gridrow="10" pageId="10" pageNumber="92">absent</td>
|
||||
<td id="768A3948FF848BC4FC9E75E8AA2BFC7D" box="[847,1073,954,978]" gridcol="3" gridrow="10" pageId="10" pageNumber="92">absent</td>
|
||||
<td id="768A3948FF848BC4FB9B75E8AB8AFC7D" box="[1098,1424,954,978]" gridcol="4" gridrow="10" pageId="10" pageNumber="92">present</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF7275B0AB8AFBED" box="[163,1424,994,1090]" gridrow="11" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF7275B0AF9AFBED" box="[163,384,994,1090]" gridcol="0" gridrow="11" pageId="10" pageNumber="92">lateral caeca in posterior region</th>
|
||||
<td id="768A3948FF848BC4FE4875B0AC5FFBED" box="[409,581,994,1090]" gridcol="1" gridrow="11" pageId="10" pageNumber="92">absent</td>
|
||||
<td id="768A3948FF848BC4FD8D75B0AD2CFBED" box="[604,822,994,1090]" gridcol="2" gridrow="11" pageId="10" pageNumber="92">present, sometimes protrude into parapodia</td>
|
||||
<td id="768A3948FF848BC4FC9E75B0AA2BFBED" box="[847,1073,994,1090]" gridcol="3" gridrow="11" pageId="10" pageNumber="92">na</td>
|
||||
<td id="768A3948FF848BC4FB9B75B0AB8AFBED" box="[1098,1424,994,1090]" gridcol="4" gridrow="11" pageId="10" pageNumber="92">present, protrude into parapodia to up to half of their length</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF727200AB8AFB21" box="[163,1424,1106,1166]" gridrow="12" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF727200AF9AFB21" box="[163,384,1106,1166]" gridcol="0" gridrow="12" pageId="10" pageNumber="92">acicular chaetae in anterior region</th>
|
||||
<td id="768A3948FF848BC4FE487200AC5FFB21" box="[409,581,1106,1166]" gridcol="1" gridrow="12" pageId="10" pageNumber="92">absent</td>
|
||||
<td id="768A3948FF848BC4FD8D7200AD2CFB21" box="[604,822,1106,1166]" gridcol="2" gridrow="12" pageId="10" pageNumber="92">absent</td>
|
||||
<td id="768A3948FF848BC4FC9E7200AA2BFB21" box="[847,1073,1106,1166]" gridcol="3" gridrow="12" pageId="10" pageNumber="92">absent</td>
|
||||
<td id="768A3948FF848BC4FB9B7200AB8AFB21" box="[1098,1424,1106,1166]" gridcol="4" gridrow="12" pageId="10" pageNumber="92">present</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF7272CCAB8AFB75" box="[163,1424,1182,1242]" gridrow="13" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF7272CCAF9AFB75" box="[163,384,1182,1242]" gridcol="0" gridrow="13" pageId="10" pageNumber="92">pygidium</th>
|
||||
<td id="768A3948FF848BC4FE4872CCAC5FFB75" box="[409,581,1182,1242]" gridcol="1" gridrow="13" pageId="10" pageNumber="92">small with short rounded cirri</td>
|
||||
<td id="768A3948FF848BC4FD8D72CCAD2CFB75" box="[604,822,1182,1242]" gridcol="2" gridrow="13" pageId="10" pageNumber="92">small with short rounded cirri</td>
|
||||
<td id="768A3948FF848BC4FC9E72CCAA2BFB75" box="[847,1073,1182,1242]" gridcol="3" gridrow="13" pageId="10" pageNumber="92">enlarged with long digitiform cirri</td>
|
||||
<td id="768A3948FF848BC4FB9B72CCAB8AFB75" box="[1098,1424,1182,1242]" gridcol="4" gridrow="13" pageId="10" pageNumber="92">enlarged with long digitiform cirri</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF7272B9AB8AFAAC" box="[163,1424,1259,1283]" gridrow="14" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF7272B9AF9AFAAC" box="[163,384,1259,1283]" gridcol="0" gridrow="14" pageId="10" pageNumber="92">external eyes</th>
|
||||
<td id="768A3948FF848BC4FE4872B9AC5FFAAC" box="[409,581,1259,1283]" gridcol="1" gridrow="14" pageId="10" pageNumber="92">absent</td>
|
||||
<td id="768A3948FF848BC4FD8D72B9AD2CFAAC" box="[604,822,1259,1283]" gridcol="2" gridrow="14" pageId="10" pageNumber="92">absent</td>
|
||||
<td id="768A3948FF848BC4FC9E72B9AA2BFAAC" box="[847,1073,1259,1283]" gridcol="3" gridrow="14" pageId="10" pageNumber="92">na</td>
|
||||
<td id="768A3948FF848BC4FB9B72B9AB8AFAAC" box="[1098,1424,1259,1283]" gridcol="4" gridrow="14" pageId="10" pageNumber="92">present</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF727341AB8AFADC" box="[163,1424,1299,1395]" gridrow="15" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF727341AF9AFADC" box="[163,384,1299,1395]" gridcol="0" gridrow="15" pageId="10" pageNumber="92">eyes in fixed specimens reach posteriorly</th>
|
||||
<td id="768A3948FF848BC4FE487341AC5FFADC" box="[409,581,1299,1395]" gridcol="1" gridrow="15" pageId="10" pageNumber="92">chaetigers 4–5</td>
|
||||
<td id="768A3948FF848BC4FD8D7341AD2CFADC" box="[604,822,1299,1395]" gridcol="2" gridrow="15" pageId="10" pageNumber="92">chaetigers 4–5</td>
|
||||
<td id="768A3948FF848BC4FC9E7341AA2BFADC" box="[847,1073,1299,1395]" gridcol="3" gridrow="15" pageId="10" pageNumber="92">~middle of chaetiger 1</td>
|
||||
<td id="768A3948FF848BC4FB9B7341AB8AFADC" box="[1098,1424,1299,1395]" gridcol="4" gridrow="15" pageId="10" pageNumber="92">peristomium / chaetiger 1 boundary</td>
|
||||
</tr>
|
||||
<tr id="355B5034FF848BC4FF7273D1AB8AF984" box="[163,1424,1411,1579]" gridrow="16" pageId="10" pageNumber="92">
|
||||
<th id="768A3948FF848BC4FF7273D1AF9AF984" box="[163,384,1411,1579]" gridcol="0" gridrow="16" pageId="10" pageNumber="92">coloration pattern</th>
|
||||
<td id="768A3948FF848BC4FE4873D1AC5FF984" box="[409,581,1411,1579]" gridcol="1" gridrow="16" pageId="10" pageNumber="92">transverse pigmented bands on dorsum</td>
|
||||
<td id="768A3948FF848BC4FD8D73D1AD2CF984" box="[604,822,1411,1579]" gridcol="2" gridrow="16" pageId="10" pageNumber="92">scattered red spots on dorsum</td>
|
||||
<td id="768A3948FF848BC4FC9E73D1AA2BF984" box="[847,1073,1411,1579]" gridcol="3" gridrow="16" pageId="10" pageNumber="92">small pigmented spots in rows anteriorly on dorsum and ventrum, and on all parapodia of posterior region</td>
|
||||
<td id="768A3948FF848BC4FB9B73D1AB8AF984" box="[1098,1424,1411,1579]" gridcol="4" gridrow="16" pageId="10" pageNumber="92">almost unpigmented anteriorly, posterior with scattered reddish brown spots on dorsum and parapodia, spots in transverse rows on ventrum</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<subSubSection id="C37101FDFF847431FF1670D3AFF8F94B" pageId="10" pageNumber="92" type="etymology">
|
||||
<paragraph id="8BD45276FF847431FF1670D3AFF8F94B" blockId="10.[151,1437,1665,2016]" pageId="10" pageNumber="92">
|
||||
<emphasis id="B91F8E64FF847431FF1670D3AF5FF934" box="[199,325,1665,1691]" italics="true" pageId="10" pageNumber="92">Etymology.</emphasis>
|
||||
The species is named after our teacher, colleague and friend, Professor Alexander Tzetlin, to honor his contribution to the study of annelid morphology and evolution, and for his incredible ability to share his wormloving attitude through years.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37101FDFF847430FF1670BFAC7CFF7A" lastPageId="11" lastPageNumber="93" pageId="10" pageNumber="92" type="discussion">
|
||||
<paragraph id="8BD45276FF847430FF1670BFAC7CFF7A" blockId="10.[151,1437,1665,2016]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="93" pageId="10" pageNumber="92">
|
||||
<emphasis id="B91F8E64FF847431FF1670BFAFDEF8A8" box="[199,452,1773,1799]" italics="true" pageId="10" pageNumber="92">Phylogenetic analysis.</emphasis>
|
||||
The tree based on concatenated
|
||||
<emphasis id="B91F8E64FF847431FCE370BFADB9F8A7" box="[818,931,1773,1800]" italics="true" pageId="10" pageNumber="92">18S rRNA</emphasis>
|
||||
and
|
||||
<emphasis id="B91F8E64FF847431FC0B70BFAA51F8A7" box="[986,1099,1773,1800]" italics="true" pageId="10" pageNumber="92">28S rRNA</emphasis>
|
||||
sequences (
|
||||
<figureCitation id="13504EF3FF847431FB0270BFAB0DF8A8" box="[1235,1303,1773,1799]" captionStart="FIGURE 7" captionStartId="11.[151,250,866,890]" captionTargetBox="[151,1435,237,843]" captionTargetId="figure-52@11.[151,1435,237,843]" captionTargetPageId="11" captionText="FIGURE 7. Phylogenetic tree based on concatenated 18S and 28S markers. Numbers in regular font above nodes indicate Bayesian posterior probability; numbers in italic under nodes represent Maximum Likelihood bootstrap values; dash indicates unsupported nodes." figureDoi="http://doi.org/10.5281/zenodo.15216212" httpUri="https://zenodo.org/record/15216212/files/figure.png" pageId="10" pageNumber="92">Fig. 7</figureCitation>
|
||||
) placed our specimen as a sister lineage to the only
|
||||
<taxonomicName id="4C6B29F5FF847431FD867143AD1DF884" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[599,775,1809,1835]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B91F8E64FF847431FD867143AD1DF884" box="[599,775,1809,1835]" italics="true" pageId="10" pageNumber="92">Ctenophoricola</emphasis>
|
||||
</taxonomicName>
|
||||
sequences available so far,
|
||||
<taxonomicName id="4C6B29F5FF847431FB937143AF71F8E0" authority="San Martin et al., 2020" authorityName="San Martin" authorityYear="2020" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="masanorii">
|
||||
<emphasis id="B91F8E64FF847431FB937143AB71F884" box="[1090,1387,1809,1835]" italics="true" pageId="10" pageNumber="92">Ctenophoricola masanorii</emphasis>
|
||||
San Martín
|
||||
<emphasis id="B91F8E64FF847431FF3A7164AF3FF8E0" box="[235,293,1845,1871]" italics="true" pageId="10" pageNumber="92">et al.</emphasis>
|
||||
, 2020
|
||||
</taxonomicName>
|
||||
from the original study. Distance between
|
||||
<taxonomicName id="4C6B29F5FF847431FC857167AA1EF8E0" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[852,1028,1845,1871]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B91F8E64FF847431FC857167AA1EF8E0" box="[852,1028,1845,1871]" italics="true" pageId="10" pageNumber="92">Ctenophoricola</emphasis>
|
||||
</taxonomicName>
|
||||
lineages (0.012) is within the range of between-species distances in the analyzed dataset. Thus, distances between considered
|
||||
<taxonomicName id="4C6B29F5FF847431FB7D710BAB7AF8DC" box="[1196,1376,1881,1907]" pageId="10" pageNumber="92">
|
||||
<emphasis id="B91F8E64FF847431FB7D710BAB1FF8DC" box="[1196,1285,1881,1907]" italics="true" pageId="10" pageNumber="92">Vanadis</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
vary from
|
||||
<quantity id="4C93FF93FF847431FF07712FAF2EF838" box="[214,308,1917,1943]" metricMagnitude="-4" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="92" unit="in" value="0.006">0.006 in</quantity>
|
||||
close species group
|
||||
<taxonomicName id="4C6B29F5FF847431FDF2712CAD9BF837" authority="(McIntosh, 1885)" authorityName="ZMMU WS" baseAuthorityName="McIntosh" baseAuthorityYear="1885" box="[547,897,1917,1944]" class="Polychaeta" family="Phyllodocidae" genus="Vanadis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="antarctica">
|
||||
<emphasis id="B91F8E64FF847431FDF2712CACA8F838" box="[547,690,1918,1943]" italics="true" pageId="10" pageNumber="92">V. antarctica</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFFA2F87FF847431FD12712FAD63F837" author="McIntosh, W. C." box="[707,889,1917,1944]" pageId="10" pageNumber="92" pagination="1 - 509" refId="ref9637" refString="McIntosh, W. C. [as M'Intosh] (1885) Report on the Annelida Polychaeta collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76, Zoology, 12 (34), i - xxxvi + 1-509." type="journal article" year="1885">McIntosh, 1885</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
—
|
||||
<taxonomicName id="4C6B29F5FF847431FC7E712CAB15F837" authority="(Levinsen, 1885)" authorityName="ZMMU WS" baseAuthorityName="Levinsen" baseAuthorityYear="1885" box="[943,1295,1917,1944]" class="Polychaeta" family="Phyllodocidae" genus="Vanadis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="longissima">
|
||||
<emphasis id="B91F8E64FF847431FC7E712CAA5FF838" box="[943,1093,1917,1943]" italics="true" pageId="10" pageNumber="92">V. longissima</emphasis>
|
||||
(
|
||||
<bibRefCitation id="EFFA2F87FF847431FB87712FAB1DF837" author="Levinsen, G. M. R." box="[1110,1287,1917,1944]" pageId="10" pageNumber="92" pagination="321 - 44" refId="ref9360" refString="Levinsen, G. M. R. (1885) Spolia atlantica. Om nogle pelagiske Annulata. Det Kongelige Danske videnskabernes selskabs skrifter. Naturvidenskabelig og mathematisk afdeling, Series 6, 3 (2), 321-44." type="journal article" year="1885">Levinsen, 1885</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
to 0.016 (
|
||||
<taxonomicName id="4C6B29F5FF847431FA59712CAFA8F814" authority="Claparede, 1870" authorityName="Claparede" authorityYear="1870" class="Polychaeta" family="Phyllodocidae" genus="Vanadis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="formosa">
|
||||
<emphasis id="B91F8E64FF847431FA59712CAEE9F814" italics="true" pageId="10" pageNumber="92">V. formosa</emphasis>
|
||||
<bibRefCitation id="EFFA2F87FF847431FF2B71F3AFA8F814" author="Claparede, E." box="[250,434,1953,1979]" pageId="10" pageNumber="92" pagination="365 - 542" refId="ref8073" refString="Claparede, E. (1870) Les Annelides Chetopodes du Golfe de Naples. Supplement. Memoires de la Societe de physique et d'histoire naturelle de Geneve, 20 (2), 365-542. https://doi.org/10.5962/bhl.title.2142" type="journal article" year="1870">Claparède, 1870</bibRefCitation>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C6B29F5FF847431FE3871F0AD0EF814" authority="Treadwell, 1906" authorityName="Treadwell" authorityYear="1906" box="[489,788,1953,1979]" class="Polychaeta" family="Phyllodocidae" genus="Vanadis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="species" species="minuta">
|
||||
<emphasis id="B91F8E64FF847431FE3871F0AC48F814" box="[489,594,1954,1979]" italics="true" pageId="10" pageNumber="92">V. minuta</emphasis>
|
||||
<bibRefCitation id="EFFA2F87FF847431FD8871F3AD0EF814" author="Treadwell, A. L." box="[601,788,1953,1979]" pageId="10" pageNumber="92" pagination="1145 - 1181" refId="ref10907" refString="Treadwell, A. L. (1906) Polychaetous annelids of the Hawaiian Islands collected by the steamer Albatross in 1902. Bulletin of the United States Fish Commission, 23 (3), 1145-1181." type="journal article" year="1906">Treadwell, 1906</bibRefCitation>
|
||||
</taxonomicName>
|
||||
); distance between two
|
||||
<taxonomicName id="4C6B29F5FF847431FBF071F3AB2FF814" box="[1057,1333,1953,1979]" pageId="10" pageNumber="92">
|
||||
<emphasis id="B91F8E64FF847431FBF071F3AAC4F814" box="[1057,1246,1953,1979]" italics="true" pageId="10" pageNumber="92">Rhynchonereella</emphasis>
|
||||
species
|
||||
</taxonomicName>
|
||||
is 0.012. The common node of
|
||||
<taxonomicName id="4C6B29F5FF847431FE5A7197AC21F870" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[395,571,1989,2015]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B91F8E64FF847431FE5A7197AC21F870" box="[395,571,1989,2015]" italics="true" pageId="10" pageNumber="92">Ctenophoricola</emphasis>
|
||||
</taxonomicName>
|
||||
lineages is well-supported (
|
||||
<figureCitation id="13504EF3FF847431FCA37197ADACF870" box="[882,950,1989,2015]" captionStart="FIGURE 7" captionStartId="11.[151,250,866,890]" captionTargetBox="[151,1435,237,843]" captionTargetId="figure-52@11.[151,1435,237,843]" captionTargetPageId="11" captionText="FIGURE 7. Phylogenetic tree based on concatenated 18S and 28S markers. Numbers in regular font above nodes indicate Bayesian posterior probability; numbers in italic under nodes represent Maximum Likelihood bootstrap values; dash indicates unsupported nodes." figureDoi="http://doi.org/10.5281/zenodo.15216212" httpUri="https://zenodo.org/record/15216212/files/figure.png" pageId="10" pageNumber="92">Fig. 7</figureCitation>
|
||||
). However, all basal nodes within
|
||||
<taxonomicName id="4C6B29F5FF847431FAE37197AB86F870" box="[1330,1436,1989,2015]" class="Polychaeta" family="Phyllodocidae" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="92" phylum="Annelida" rank="tribe" tribe="Alciopini">Alciopini</taxonomicName>
|
||||
including the one nesting
|
||||
<taxonomicName id="4C6B29F5FF857430FE1676C5AC6DFF1E" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[455,631,151,177]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="93" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B91F8E64FF857430FE1676C5AC6DFF1E" box="[455,631,151,177]" italics="true" pageId="11" pageNumber="93">Ctenophoricola</emphasis>
|
||||
</taxonomicName>
|
||||
within
|
||||
<taxonomicName id="4C6B29F5FF857430FD0376C5AD26FF1E" box="[722,828,151,177]" class="Polychaeta" family="Phyllodocidae" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="93" phylum="Annelida" rank="tribe" tribe="Alciopini">Alciopini</taxonomicName>
|
||||
lack support, thus leaving the position of the genus within
|
||||
<taxonomicName id="4C6B29F5FF857430FF3576E9AF54FF7A" box="[228,334,187,213]" class="Polychaeta" family="Phyllodocidae" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="93" phylum="Annelida" rank="tribe" tribe="Alciopini">Alciopini</taxonomicName>
|
||||
unresolved in this study.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="DF1402FEFF857430FF467530AF47FC6D" ID-DOI="http://doi.org/10.5281/zenodo.15216212" ID-Zenodo-Dep="15216212" httpUri="https://zenodo.org/record/15216212/files/figure.png" pageId="11" pageNumber="93" startId="11.[151,250,866,890]" targetBox="[151,1435,237,843]" targetPageId="11" targetType="figure">
|
||||
<paragraph id="8BD45276FF857430FF467530AF47FC6D" blockId="11.[151,1436,866,962]" pageId="11" pageNumber="93">
|
||||
<emphasis id="B91F8E64FF857430FF467530AF0CFCD4" bold="true" box="[151,278,866,891]" pageId="11" pageNumber="93">FIGURE 7.</emphasis>
|
||||
Phylogenetic tree based on concatenated 18S and 28S markers. Numbers in regular font above nodes indicate Bayesian posterior probability; numbers in italic under nodes represent Maximum Likelihood bootstrap values; dash indicates unsupported nodes.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="C37101FDFF857430FF467244AAD9FB34" pageId="11" pageNumber="93" type="multiple">
|
||||
<paragraph id="8BD45276FF857430FF467244ACF9FB80" blockId="11.[151,739,1045,1071]" box="[151,739,1045,1071]" pageId="11" pageNumber="93">
|
||||
<emphasis id="B91F8E64FF857430FF467244ACF9FB80" box="[151,739,1045,1071]" italics="true" pageId="11" pageNumber="93">
|
||||
Key to formally described
|
||||
<taxonomicName id="4C6B29F5FF857430FE1F7247AC9CFB80" box="[462,646,1045,1071]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="93" phylum="Annelida" rank="genus">Ctenophoricola</taxonomicName>
|
||||
species
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF857430FF46720FAAD9FB34" blockId="11.[151,1436,1117,1179]" pageId="11" pageNumber="93">
|
||||
Below, we present a short key including
|
||||
<taxonomicName id="4C6B29F5FF857430FDBB720FACC2FBD8" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[618,728,1117,1143]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="93" phylum="Annelida" rank="species" species="tzetlini" status="sp. nov.">
|
||||
<emphasis id="B91F8E64FF857430FDBB720FACC2FBD8" box="[618,728,1117,1143]" italics="true" pageId="11" pageNumber="93">C. tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B91F8E64FF857430FD30720FAD26FBD8" bold="true" box="[737,828,1117,1143]" pageId="11" pageNumber="93">
|
||||
<taxonomicNameLabel id="A22C331FFF857430FD30720FAD26FBD8" box="[737,828,1117,1143]" pageId="11" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
and two other formally described species. We do not consider undescribed
|
||||
<taxonomicName id="4C6B29F5FF857430FE5A72D3AC60FB34" box="[395,634,1153,1179]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="93" phylum="Annelida" rank="species" species="undefined-A">
|
||||
<emphasis id="B91F8E64FF857430FE5A72D3AC21FB34" box="[395,571,1153,1179]" italics="true" pageId="11" pageNumber="93">Ctenophoricola</emphasis>
|
||||
sp. A
|
||||
</taxonomicName>
|
||||
from the original paper by
|
||||
<bibRefCitation id="EFFA2F87FF857430FC7F72D3AAA5FB34" author="San Martin, G. & Alvarez-Campos, P. & Kondo, Y. & Nunez, J. & Fernandez-Alamo, M. A. & Pleijel, F. & Goetz, F. E. & Nygren, A. & Osborn, K." box="[942,1215,1153,1179]" pageId="11" pageNumber="93" pagination="672 - 694" refId="ref10510" refString="San Martin, G., Alvarez-Campos, P., Kondo, Y., Nunez, J., Fernandez-Alamo, M. A., Pleijel, F., Goetz, F. E., Nygren, A. & Osborn, K. (2021 (online 2020 )) New symbiotic association in marine annelids: ectoparasites of comb jellies. Zoological Journal of the Linnean Society, 191 (3), 672-694. https://doi.org/10.1093/zoolinnean/zlaa034" type="journal article" year="2021">
|
||||
San Martin
|
||||
<emphasis id="B91F8E64FF857430FBE372D0AA79FB34" box="[1074,1123,1153,1179]" italics="true" pageId="11" pageNumber="93">et al</emphasis>
|
||||
. (2021)
|
||||
</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37101FDFF857430FF467294AB86FAFE" pageId="11" pageNumber="93" type="key">
|
||||
<key id="FFFA0547FF857430FF467294AB86FAFE" pageId="11" pageNumber="93">
|
||||
<keyStep id="309F4AD3FF857430FF467294AB86FABA" pageId="11" pageNumber="93">
|
||||
<paragraph id="8BD45276FF857430FF467294AB86FB74" blockId="11.[151,1436,1222,1361]" box="[151,1436,1222,1244]" pageId="11" pageNumber="93">
|
||||
<keyLead id="309AF143FF857430FF467294AB86FB74" box="[151,1436,1222,1244]" pageId="11" pageNumber="93">1 Prostomium without external eyes; parapodia of anterior region only with acicula; body relatively short and stout........ 2</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF857430FF4672B1AB86FABA" blockId="11.[151,1436,1222,1361]" pageId="11" pageNumber="93">
|
||||
<keyLead id="309AF143FF857430FF4672B1AB86FABA" pageId="11" pageNumber="93">
|
||||
- Prostomium with external eyes; parapodia of anterior region with acicular chaeta below aciculum; body relatively slender..............................................................................................
|
||||
<taxonomicName id="4C6B29F5FF857430FAEE7353AB86FABA" ID-CoL="KTTY6" authorityName="Syomin & Kolbasova & Semenova & Neretina" authorityYear="2025" box="[1343,1436,1280,1302]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="93" phylum="Annelida" rank="species" species="tzetlini">
|
||||
<emphasis id="B91F8E64FF857430FAEE7353AB86FABA" box="[1343,1436,1280,1302]" italics="true" pageId="11" pageNumber="93">C. tzetlini</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="309F4AD3FF857430FF46734CAB86FAFE" pageId="11" pageNumber="93">
|
||||
<paragraph id="8BD45276FF857430FF46734CAB86FA9C" blockId="11.[151,1436,1222,1361]" box="[151,1436,1310,1332]" pageId="11" pageNumber="93">
|
||||
<keyLead id="309AF143FF857430FF46734CAB86FA9C" box="[151,1436,1310,1332]" pageId="11" pageNumber="93">
|
||||
2 Gut without lateral caeca; transverse pigmented bands on dorsum....................................
|
||||
<taxonomicName id="4C6B29F5FF857430FACE734CAB86FA9C" ID-CoL="85RBT" box="[1311,1436,1310,1331]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="93" phylum="Annelida" rank="species" species="masanorii">
|
||||
<emphasis id="B91F8E64FF857430FACE734CAB86FA9C" box="[1311,1436,1310,1331]" italics="true" pageId="11" pageNumber="93">C. masanorii</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="8BD45276FF857430FF467369AB86FAFE" blockId="11.[151,1436,1222,1361]" box="[151,1436,1339,1361]" pageId="11" pageNumber="93">
|
||||
<keyLead id="309AF143FF857430FF467369AB86FAFE" box="[151,1436,1339,1361]" pageId="11" pageNumber="93">
|
||||
- Gut with lateral caeca in posterior region; scattered red spots on dorsum..................................
|
||||
<taxonomicName id="4C6B29F5FF857430FA947369AB86FAFE" ID-CoL="85RBV" box="[1349,1436,1339,1361]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="11" pageNumber="93" phylum="Annelida" rank="species" species="rousei">
|
||||
<emphasis id="B91F8E64FF857430FA947369AB86FAFE" box="[1349,1436,1339,1361]" italics="true" pageId="11" pageNumber="93">C. rousei</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</key>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
105
data/03/C2/E3/03C2E360FF8D7438FF467044AF7FF83C.xml
Normal file
105
data/03/C2/E3/03C2E360FF8D7438FF467044AF7FF83C.xml
Normal file
|
|
@ -0,0 +1,105 @@
|
|||
<document id="3DC07BC22868AD1B2672E7FC5D3A2C17" ID-DOI="10.11646/zootaxa.5613.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="15216197" ID-ZooBank="FAC715B6-F3A1-470B-880F-56423F69D3A3" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1744667152350" checkinUser="plazi" docAuthor="Syomin, Vitaly, Kolbasova, Glafira, Semenova, Maya & Neretina, Tatiana" docDate="2025" docId="03C2E360FF8D7438FF467044AF7FF83C" docLanguage="en" docName="zootaxa.5613.1.3.pdf" docOrigin="Zootaxa 5613 (1)" docSource="https://doi.org/10.11646/zootaxa.5613.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alciopa;Alciopina;Ctenophoricola;Krohnia;Mauita;Naiades;Plotohelmis;Pseudalciopa;Rhynchonereella;Torrea;Vanadis;Watelio Ehlers 1864" docType="treatment" docVersion="3" lastPageNumber="85" masterDocId="FFFB9B18FF8E743BFFD17652AE1AFFAF" masterDocTitle="A new species of an unusual polychaete genus Ctenophoricola (Phyllodocida, Phyllodocidae, Alciopini) from the Indian Ocean" masterLastPageNumber="98" masterPageNumber="82" pageNumber="85" updateTime="1744809039579" updateUser="juliana" zenodo-license-document="CLOSED">
|
||||
<mods:mods id="838DF7BEB0D6BA2A6B80D8132A7AC4C8" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="0651F81A20292EB4F65A2544A51BCE61">
|
||||
<mods:title id="CA20B5F4EDD9414B2B3E3C025F25CB37">A new species of an unusual polychaete genus Ctenophoricola (Phyllodocida, Phyllodocidae, Alciopini) from the Indian Ocean</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="A54258606E80D3DD28C6B50C7DFA6116" type="personal">
|
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<mods:role id="B4E4E3420BB34143FFF9A504F80277B8">
|
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<mods:roleTerm id="525CCB880E3085AA9EFC5B971ABCE3A4">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="E861A6399A61146C2954CAACCDCE381B">Syomin, Vitaly</mods:namePart>
|
||||
<mods:nameIdentifier id="D8CBFFD4D245A03F1DFAAA474ED33131" type="ORCID">0000-0003-1257-3298</mods:nameIdentifier>
|
||||
<mods:affiliation id="5CB0C9DA8B3E14DFB8B0921F10E5853C">Red Sea Research Centre, King Abdullah University of Science and Technology (KAUST), Thuwal, 23955 - 6900, Kingdom of Saudi Arabia & Shirshov Institute of Oceanology of Russian Academy of Sciences, 36 Nahimovskiy Avenue, 117997 Moscow, Russia</mods:affiliation>
|
||||
<mods:nameIdentifier id="EA0BF490A95B03877163AE07024ED32B" type="email">svinovod@yandex.ru</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name id="332FBB50FD1DCDF5597E66A8B9C3D182" type="personal">
|
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<mods:role id="45AEE20073CDEDBE5C7B543FF410D7FB">
|
||||
<mods:roleTerm id="A790A26985DBD02B162415465A365C0B">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="92E950A511E5E0F006392F48BF3DFEDE">Kolbasova, Glafira</mods:namePart>
|
||||
<mods:nameIdentifier id="9934CDD33E5F886D667FF447E44D5971" type="ORCID">0000-0001-9252-0473</mods:nameIdentifier>
|
||||
<mods:affiliation id="0902B8BB8EF31450CD6D836958DDC9E3">N. A. Pertsov White Sea Biological Station, Lomonosov Moscow State University, 1 Leninskie Gory, building 12, 119234 Moscow, Russia</mods:affiliation>
|
||||
<mods:nameIdentifier id="69E1EC231615BBC014EE5FE33BED2338" type="email">utricularia57@yandex.ru</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name id="438C42FA8E643275B1FAC54E7AB34707" type="personal">
|
||||
<mods:role id="2EC7CA9732D58855C6F7D4DAA8265EE4">
|
||||
<mods:roleTerm id="DF7783A65FC16584C023F311B048A17C">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="70DA92A362470CFF367F321B79334DAC">Semenova, Maya</mods:namePart>
|
||||
<mods:nameIdentifier id="BDF448D45EAFF22F2AEF70B9FB28CC11" type="ORCID">0009-0000-9994-1879</mods:nameIdentifier>
|
||||
<mods:affiliation id="F985BF3FE1B3180A6896EA657839B5D5">N. A. Pertsov White Sea Biological Station, Lomonosov Moscow State University, 1 Leninskie Gory, building 12, 119234 Moscow, Russia</mods:affiliation>
|
||||
<mods:nameIdentifier id="0A31D4FF83D8E2A8029C51494352D5BB" type="email">maya.yakuts@gmail.com</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name id="68A642FA01497E786ED82EFCF5AB16AE" type="personal">
|
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<mods:role id="5EE11EDC4BCE8268B429354908E5B94A">
|
||||
<mods:roleTerm id="90EB97D067DDFCA6805C5C518B721836">Author</mods:roleTerm>
|
||||
</mods:role>
|
||||
<mods:namePart id="A8A3EF6F58DFABE8BEF627DC0BF4B93A">Neretina, Tatiana</mods:namePart>
|
||||
<mods:nameIdentifier id="D77AB1C4FC17C14A774331B6E7BC6760" type="ORCID">0000-0002-0673-1109</mods:nameIdentifier>
|
||||
<mods:affiliation id="A230F76F74DF29364D772351DF9798A9">N. A. Pertsov White Sea Biological Station, Lomonosov Moscow State University, 1 Leninskie Gory, building 12, 119234 Moscow, Russia</mods:affiliation>
|
||||
<mods:nameIdentifier id="13F10D41B81303E4FFCF9D1C35876FFC" type="email">nertata@wsbs-msu.ru</mods:nameIdentifier>
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<mods:title id="F3A2450DB8162B022F35310E6CC24A18">Zootaxa</mods:title>
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<mods:date id="EAF8FE3BB8BEE78B6B50D429CCDCE9C2">2025</mods:date>
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<treatment id="03C2E360FF8D7438FF467044AF7FF83C" LSID="urn:lsid:plazi:treatment:03C2E360FF8D7438FF467044AF7FF83C" httpUri="http://treatment.plazi.org/id/03C2E360FF8D7438FF467044AF7FF83C" lastPageNumber="85" pageId="3" pageNumber="85">
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<subSubSection id="C37101FDFF8D7438FF467044AFEFF99F" box="[151,501,1558,1584]" pageId="3" pageNumber="85" type="nomenclature">
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<paragraph id="8BD45276FF8D7438FF467044AFEFF99F" blockId="3.[151,501,1558,1584]" box="[151,501,1558,1584]" pageId="3" pageNumber="85">
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|
||||
<emphasis id="B91F8E64FF8D7438FF467044AFEFF99F" bold="true" box="[151,501,1558,1584]" pageId="3" pageNumber="85">
|
||||
Tribe
|
||||
<taxonomicName id="4C6B29F5FF8D7438FF307044AFEFF99F" ID-CoL="85R7B" authority="Ehlers, 1864" authorityName="Ehlers" authorityYear="1864" box="[225,501,1558,1584]" class="Polychaeta" family="Phyllodocidae" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="85" phylum="Annelida" rank="tribe" tribe="Alciopini">
|
||||
Alciopini
|
||||
<bibRefCitation id="EFFA2F87FF8D7438FE8B7044AFEFF99F" author="Ehlers, E. H." box="[346,501,1558,1584]" pageId="3" pageNumber="85" refId="ref8352" refString="Ehlers, E. H. (1864) Die Borstenwurmer (Annelida Chaetopoda) nach systematischen und anatomischen Untersuchungen dargestellt. Vol. 1. W. Engelmann, Leipzig, 268 pp. https://doi.org/10.5962/bhl.title.2081" type="book" year="1864">Ehlers, 1864</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37101FDFF8D7438FF467009AB8DF9DC" box="[151,1431,1627,1653]" pageId="3" pageNumber="85" type="type_taxon">
|
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<paragraph id="8BD45276FF8D7438FF467009AB8DF9DC" blockId="3.[151,1431,1627,1653]" box="[151,1431,1627,1653]" pageId="3" pageNumber="85">
|
||||
Genus
|
||||
<taxonomicName id="4C6B29F5FF8D7438FF0F7009AB8DF9DC" authority="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn, 2020" authorityName="San Martin, Alvarez-Campos, Kondo, Nunez, Fernandez-Alamo, Pleijel, Goetz, Nygren, Osborn" authorityYear="2020" box="[222,1431,1627,1653]" class="Polychaeta" family="Phyllodocidae" genus="Ctenophoricola" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="85" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B91F8E64FF8D7438FF0F7009AF67F9DC" box="[222,381,1627,1651]" italics="true" pageId="3" pageNumber="85">Ctenophoricola</emphasis>
|
||||
San Martín, Álvarez-Campos, Kondo, Núñez, Fernández-Álamo, Pleijel, Goetz, Nygren, Osborn, 2020
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C37101FDFF8D7438FF4670F0AF7FF83C" pageId="3" pageNumber="85" type="diagnosis">
|
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<paragraph id="8BD45276FF8D7438FF4670F0AF7FF83C" blockId="3.[151,1437,1697,1939]" pageId="3" pageNumber="85">
|
||||
<emphasis id="B91F8E64FF8D7438FF4670F0AF90F914" box="[151,394,1697,1723]" italics="true" pageId="3" pageNumber="85">Diagnosis (amended)</emphasis>
|
||||
:
|
||||
<taxonomicName id="4C6B29F5FF8D7438FE4B70F3AC1EF914" box="[410,516,1697,1723]" class="Polychaeta" family="Phyllodocidae" kingdom="Animalia" order="Phyllodocida" pageId="3" pageNumber="85" phylum="Annelida" rank="tribe" tribe="Alciopini">Alciopini</taxonomicName>
|
||||
with body divided in two distinctly different regions, variably covered by cilia. Anterior region highly contractile; parapodia consist of small acute chaetigerous lobe with aciculum, and indistinct dorsal lobe. Posterior region enlarged due to larger segments and/or expanded parapodia. Posterior parapodia trilobed, chaetigerous lobe with thin single aciculum and multiple capillary chaetae. All parapodia uniramous. Prostomium with two tiny retractile palps, without antennae. Pharynx eversible, unarmed, without papillae. Mouth surrounded by swollen lips. Two distinctly pigmented, elongate, lensed eyes, extending through one to several anterior segments.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
238
data/03/F1/15/03F115033B151057FF67FF0AFB55FFDE.xml
Normal file
238
data/03/F1/15/03F115033B151057FF67FF0AFB55FFDE.xml
Normal file
|
|
@ -0,0 +1,238 @@
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<mods:title id="0C35035D8F2B11DAC7B6C149ABEF52F2">Melanophylla angustior (Melanophyllaceae), a new species from southeastern Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="9BC6532EE6380C1F18B9F8A215AA2D0A" type="personal">
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<mods:namePart id="EA3EEA548E18C4043064C2608C59D554">Rabenantoandro, Johny</mods:namePart>
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<mods:title id="5532C1C9FB86EF08C580693C1BEAED73">Adansonia</mods:title>
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<mods:part id="B4763822AB69063C95A71B3B22689C27">
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<mods:date id="2050EEF5F350E6DF3BE2A4A0037491F1">2002</mods:date>
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<mods:title id="A36B3B065A9D9AC9F31E375858C6FAA6">3</mods:title>
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<subSubSection id="C342F79E3B151057FF67FF0AFE3DF9B8" pageId="1" pageNumber="264" type="nomenclature">
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<taxonomicName id="4C58DF963B151057FF67FF0AFE9EF9BF" ID-CoL="3Z9MG" authority="McPherson & Rabenantoandro" authorityName="McPherson & Rabenantoandro" authorityYear="2002" class="Magnoliopsida" family="Torricelliaceae" genus="Melanophylla" kingdom="Plantae" order="Apiales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="angustior" status="sp. nov.">
|
||||
<emphasis id="B92C78073B151057FF67FF0AFE16F99E" bold="true" box="[151,480,170,197]" pageId="1" pageNumber="264">Melanophylla angustior</emphasis>
|
||||
McPherson & Rabenantoandro
|
||||
</taxonomicName>
|
||||
,
|
||||
<emphasis id="B92C78073B151057FE84FF69FE3DF9B8" bold="true" box="[372,459,201,227]" pageId="1" pageNumber="264">
|
||||
<taxonomicNameLabel id="A21FC57C3B151057FE84FF69FE3DF9B8" box="[372,459,201,227]" pageId="1" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C342F79E3B151057FF41FEA3FEADF8DA" pageId="1" pageNumber="264" type="description">
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<paragraph id="8BE7A4153B151057FF41FEA3FEADF8DA" blockId="1.[151,684,259,385]" pageId="1" pageNumber="264">
|
||||
<emphasis id="B92C78073B151057FF41FEA3FEA7F816" italics="true" pageId="1" pageNumber="264">
|
||||
A speciebus cognitibus combinatione formae (oblongae, anguste ellipticae vel anguste obovatae) cum amplitudine foliorum (
|
||||
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|
||||
</emphasis>
|
||||
×
|
||||
<emphasis id="B92C78073B151057FE9CFE95FEADF8DA" italics="true" pageId="1" pageNumber="264">
|
||||
<quantity id="4CA009F03B151057FE9CFE95FE33F817" box="[364,453,309,332]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="1" pageNumber="264" unit="cm" value="6.25" valueMax="7.5" valueMin="5.0">5-7.5 cm</quantity>
|
||||
), longitudine petiolorum (
|
||||
<quantity id="4CA009F03B151057FF6EFEEEFF0EF83D" box="[158,248,334,358]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.7" metricValueMin="3.0" pageId="1" pageNumber="264" unit="cm" value="3.85" valueMax="4.7" valueMin="3.0">3-4.7 cm</quantity>
|
||||
), longitudine pedicellorum (
|
||||
<quantity id="4CA009F03B151057FDF0FEEEFDB9F83D" box="[512,591,334,358]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="1" pageNumber="264" unit="mm" value="0.5" valueMax="1.0" valueMin="0.0">0-1 mm</quantity>
|
||||
), et colore florum (albo) differt.
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C342F79E3B151057FF41FE00FE1EFB45" pageId="1" pageNumber="264" type="materials_examined">
|
||||
<paragraph id="8BE7A4153B151057FF41FE00FE1EFB45" blockId="1.[151,684,416,542]" pageId="1" pageNumber="264">
|
||||
<typeStatus id="54E31AB73B151057FF41FE00FF02F8ED" box="[177,244,416,439]" pageId="1" pageNumber="264">TYPUS</typeStatus>
|
||||
. —
|
||||
<materialsCitation id="3B30AE483B151057FED9FE01FEB9FB45" collectingDate="2001-11-14" collectionCode="MO" collectorName="Rabenantoandro & McPherson" country="Madagascar" elevation="30" latitude="-23.021944" location="Reserve Speciale de Manombo" longLatPrecision="21" longitude="47.732224" municipality="Prov. Fianarantosoa" pageId="1" pageNumber="264" specimenCount="1" typeStatus="holotype">
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<emphasis id="B92C78073B151057FED9FE01FD5DF8E3" box="[297,683,416,441]" italics="true" pageId="1" pageNumber="264">
|
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<collectorName id="26ADC1C33B151057FED9FE01FE21F8E2" box="[297,471,417,441]" pageId="1" pageNumber="264">Rabenantoandro</collectorName>
|
||||
&
|
||||
<collectorName id="26ADC1C33B151057FE0EFE01FD87F8E2" box="[510,625,417,441]" pageId="1" pageNumber="264">McPherson</collectorName>
|
||||
774,
|
||||
</emphasis>
|
||||
<collectingCountry id="F34FE4853B151057FF67FE1AFEFBF889" box="[151,269,442,466]" name="Madagascar" pageId="1" pageNumber="264">Madagascar</collectingCountry>
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||||
,
|
||||
<collectingMunicipality id="6B833E6F3B151057FEE8FE1AFE16F889" box="[280,480,442,466]" pageId="1" pageNumber="264">Prov. Fianarantosoa</collectingMunicipality>
|
||||
,
|
||||
<location id="8E87F2CE3B151057FE1CFE1AFEF2F8B7" LSID="urn:lsid:plazi:treatment:03F115033B151057FF67FF0AFB55FFDE:8E87F2CE3B151057FE1CFE1AFEF2F8B7" country="Madagascar" latitude="-23.021944" longLatPrecision="21" longitude="47.732224" municipality="Prov. Fianarantosoa" name="Reserve Speciale de Manombo" pageId="1" pageNumber="264">Réserve Spéciale de Manombo</location>
|
||||
,
|
||||
<location id="8E87F2CE3B151057FEE0FE74FF12FB5E" LSID="urn:lsid:plazi:treatment:03F115033B151057FF67FF0AFB55FFDE:8E87F2CE3B151057FEE0FE74FF12FB5E" country="Madagascar" latitude="-23.021944" longLatPrecision="21" longitude="47.732224" municipality="Prov. Fianarantosoa" name="foret dense humide de basse altitude sur laterite" pageId="1" pageNumber="264">forêt dense humide de basse altitude sur latérite</location>
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||||
,
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||||
<geoCoordinate id="EE6CC2D23B151057FF01FE4DFE86FB5E" box="[241,368,493,517]" degrees="23" direction="south" minutes="01" orientation="latitude" pageId="1" pageNumber="264" precision="15" seconds="19" value="-23.021944">23º01’19”S</geoCoordinate>
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,
|
||||
<geoCoordinate id="EE6CC2D23B151057FE8DFE4DFE08FB5E" box="[381,510,493,517]" degrees="47" direction="east" minutes="43" orientation="longitude" pageId="1" pageNumber="264" precision="15" seconds="56" value="47.732224">47º43’56”E</geoCoordinate>
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,
|
||||
<quantity id="4CA009F03B151057FDFCFE4DFDB0FB58" box="[524,582,492,517]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="264" unit="m" value="30.0">
|
||||
<elevation id="007543263B151057FDFCFE4DFDB0FB58" box="[524,582,492,517]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="264" unit="m" value="30.0">30 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<date id="FFE682D53B151057FDA4FE4DFF3CFB45" pageId="1" pageNumber="264" value="2001-11-14">
|
||||
<collectingDate id="EFA27B3D3B151057FDA4FE4DFF3CFB45" pageId="1" pageNumber="264" value="2001-11-14">14 Nov. 2001</collectingDate>
|
||||
</date>
|
||||
(holo-,
|
||||
<collectionCode id="ED493CD03B151057FEE8FDA6FEBEFB45" box="[280,328,518,542]" pageId="1" pageNumber="264">MO</collectionCode>
|
||||
!;
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B30AE483B151057FEA6FDA6FE56FB45" box="[342,416,518,542]" collectingDate="2001-11-14" collectionCode="P" collectorName="Rabenantoandro & McPherson" country="Madagascar" elevation="30" latitude="-23.021944" location="Reserve Speciale de Manombo" longLatPrecision="21" longitude="47.732224" municipality="Prov. Fianarantosoa" pageId="1" pageNumber="264" specimenCount="1" typeStatus="isotype">
|
||||
iso-,
|
||||
<collectionCode id="ED493CD03B151057FE76FDA6FE6EFB45" box="[390,408,518,542]" pageId="1" pageNumber="264">P</collectionCode>
|
||||
!,
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B30AE483B151057FE56FDA6FE12FB45" box="[422,484,518,542]" collectingDate="2001-11-14" collectionCode=" TEF" collectorName="Rabenantoandro & McPherson" country="Madagascar" elevation="30" latitude="-23.021944" location="Reserve Speciale de Manombo" longLatPrecision="21" longitude="47.732224" municipality="Prov. Fianarantosoa" pageId="1" pageNumber="264" specimenCount="1" typeStatus="isotype">
|
||||
<collectionCode id="ED493CD03B151057FE56FDA6FE2FFB45" box="[422,473,518,542]" pageId="1" pageNumber="264">TEF</collectionCode>
|
||||
!)
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C342F79E3B151057FF41FDF8FB88FA57" pageId="1" pageNumber="264" type="description">
|
||||
<paragraph id="8BE7A4153B151057FF41FDF8FB88FA57" blockId="1.[151,684,600,1669]" lastBlockId="1.[714,1248,170,780]" pageId="1" pageNumber="264">
|
||||
Tree c.
|
||||
<quantity id="4CA009F03B151057FEF2FDF9FED9FB29" box="[258,303,600,627]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="264" unit="m" value="8.0">8 m</quantity>
|
||||
tall; branches
|
||||
<quantity id="4CA009F03B151057FE3EFDF9FDD1FB29" box="[462,551,600,627]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="1" pageNumber="264" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7-9 mm</quantity>
|
||||
in diameter just below their leafy portions, light-coloured, glabrous, marked by closely spaced leafscars, each of which half-encircles the stem and displays c. 10 protruding leaf traces. Leaves somewhat clustered near the branch ends, the leafy portion of the stem c.
|
||||
<quantity id="4CA009F03B151057FF42FCB1FEF7FA71" box="[178,257,784,811]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="1" pageNumber="264" unit="cm" value="6.5" valueMax="8.0" valueMin="5.0">5-8 cm</quantity>
|
||||
long. Petioles
|
||||
<quantity id="4CA009F03B151057FE54FCB1FDFFFA71" box="[420,521,784,811]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.7" metricValueMin="3.0" pageId="1" pageNumber="264" unit="cm" value="3.85" valueMax="4.7" valueMin="3.0">3-4.7 cm</quantity>
|
||||
long,
|
||||
<quantity id="4CA009F03B151057FDA2FCB1FD5AFA71" box="[594,684,784,811]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="1" pageNumber="264" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2-4 mm</quantity>
|
||||
wide except at the amplexicaul base (
|
||||
<quantity id="4CA009F03B151057FDBDFC8FFD5AFA13" box="[589,684,814,841]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="1" pageNumber="264" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5-9 mm</quantity>
|
||||
wide), the width near the apex c.
|
||||
<quantity id="4CA009F03B151057FDE5FCEEFDAEFA3C" box="[533,600,845,872]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="264" unit="mm" value="2.0">2 mm</quantity>
|
||||
in side view, caniculate, the rims of the channel often thin and wing-like the entire length of the petiole, not suberose at maturity, glabrous except for the margins of the wings, which are often minutely and obscurely pubescent; petiole length/blade length ratio c. 1/5-1/7. Blades oblong, narrowly elliptic, or narrowly obovate;
|
||||
<quantity id="4CA009F03B151057FE7BFB85FE0CFD65" box="[395,506,1060,1087]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="264" unit="cm" value="25.0" valueMax="30.0" valueMin="20.0">20-30 cm</quantity>
|
||||
long,
|
||||
<quantity id="4CA009F03B151057FDB5FB85FD5AFD65" box="[581,684,1060,1087]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="1" pageNumber="264" unit="cm" value="6.25" valueMax="7.5" valueMin="5.0">5-7.5 cm</quantity>
|
||||
wide (excepting the reduced leaves immediately below the inflorescence); length/width ratio 4/1- 5/1; fibrous on being torn (as are other species in the genus); base rather broadly acute; apex narrowly obtuse or briefly acuminate-obtuse; margin subentire to shallowly dentate, especially distally, somewhat revolute; venation slightly raised adaxially, more prominent abaxially, the midrib pinkish in vivo, the secondary veins 10-12 on each side of the midrib; surfaces glabrous; texture chartaceous to subcoriaceous on drying. Inflorescence terminal, paniculate, c.
|
||||
<quantity id="4CA009F03B151057FEAEFA35FE39FCF5" box="[350,463,1428,1455]" metricMagnitude="-1" metricUnit="m" metricValue="1.175" metricValueMax="1.4" metricValueMin="0.95" pageId="1" pageNumber="264" unit="cm" value="11.75" valueMax="14.0" valueMin="9.5">9.5-14 cm</quantity>
|
||||
long, c.
|
||||
<quantity id="4CA009F03B151057FDC1FA35FD7BFCF5" box="[561,653,1428,1455]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="1" pageNumber="264" unit="cm" value="9.0" valueMax="10.0" valueMin="8.0">8-10 cm</quantity>
|
||||
in diameter, of c. 8 major branches, the lower ones in the axils of reduced leaves and c.
|
||||
<quantity id="4CA009F03B151057FE09FA72FD99FCB0" box="[505,623,1489,1516]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="8.5" metricValueMin="6.5" pageId="1" pageNumber="264" unit="cm" value="7.5" valueMax="8.5" valueMin="6.5">6.5-8.5 cm</quantity>
|
||||
long; axes, bracts, and bracteoles minutely pubescent; each flower subtended by a roughly triangular bract
|
||||
<quantity id="4CA009F03B151057FF26F98EFEB0FF1C" box="[214,326,1581,1608]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="264" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5-2 mm</quantity>
|
||||
long and two smaller bracteoles that are attached immediately beneath the ovary; pedicels 0-0.5 (-1) mm long (and thus the flowers sessile or nearly so). Flowers apparently protandrous, the stamens opening quickly and soon falling, the styles remaining and apparently receptive for a longer period. Buds up to
|
||||
<quantity id="4CA009F03B151057FBBCFEA7FB54F87B" box="[1100,1186,262,289]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="264" unit="mm" value="5.5">5.5 mm</quantity>
|
||||
long. Calyx epigynous, c.
|
||||
<quantity id="4CA009F03B151057FC6BFE86FC2DF864" box="[923,987,293,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="264" unit="mm" value="1.0">1 mm</quantity>
|
||||
long,
|
||||
<quantity id="4CA009F03B151057FBECFE86FBAAF864" box="[1052,1116,293,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="264" unit="mm" value="2.0">2 mm</quantity>
|
||||
in diameter, 5-lobed (the lobes c.
|
||||
<quantity id="4CA009F03B151057FC4EFEE5FBEFF805" box="[958,1049,324,351]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="264" unit="mm" value="0.5">0.5 mm</quantity>
|
||||
long), somewhat pubescent abaxially and more so on the margin. Corolla in bud ellipsoidal, up to
|
||||
<quantity id="4CA009F03B151057FBB3FE22FB6BF8C0" box="[1091,1181,385,412]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="264" unit="mm" value="3.5">3.5 mm</quantity>
|
||||
long,
|
||||
<quantity id="4CA009F03B151057FD3AFE01FCE9F8E1" box="[714,799,416,443]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="264" unit="mm" value="2.5">2.5 mm</quantity>
|
||||
in diameter, the 5 petals slightly imbricate in bud, reflexed at anthesis and
|
||||
<quantity id="4CA009F03B151057FBE9FE1FFB87F883" box="[1049,1137,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="264" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4-5 mm</quantity>
|
||||
long,
|
||||
<quantity id="4CA009F03B151057FB44FE1FFCF8F8AC" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="264" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5- 2 mm</quantity>
|
||||
wide, yellowish white, the margins and adaxial surface minutely papillose. Stamens 5,
|
||||
<quantity id="4CA009F03B151057FB44FE5DFCFCFB6F" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="1" pageNumber="264" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5- 3 mm</quantity>
|
||||
long, the filaments
|
||||
<quantity id="4CA009F03B151057FC2AFDBBFBB3FB6F" box="[986,1093,538,565]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="264" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5-2 mm</quantity>
|
||||
long, attached basally/abaxially between the lobes of the sagittate anthers, these
|
||||
<quantity id="4CA009F03B151057FC95FDF9FC51FB29" box="[869,935,600,627]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="264" unit="mm" value="2.0">2 mm</quantity>
|
||||
long and laterally dehiscent. Ovary
|
||||
<quantity id="4CA009F03B151057FCEAFDD7FC7BFBCB" box="[794,909,630,657]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="264" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2-2.5 mm</quantity>
|
||||
long,
|
||||
<quantity id="4CA009F03B151057FC29FDD7FBBAFBCB" box="[985,1100,630,657]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="1" pageNumber="264" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 mm</quantity>
|
||||
in diameter, glabrous or minutely pubescent; styles 3,
|
||||
<quantity id="4CA009F03B151057FB40FD36FCD2FB95" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="1" pageNumber="264" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5- 3.5 mm</quantity>
|
||||
long at anthesis, the distal half flaring somewhat into the papillose-glandular adaxial stigmatic surfaces. Fruit unknown. —
|
||||
<figureCitation id="1363B8903B151057FBCBFD52FB8EFA57" box="[1083,1144,754,780]" captionStart="Fig" captionStartId="2.[280,308,1108,1125]" captionTargetBox="[174,1195,161,1074]" captionTargetId="figure-218@2.[174,1195,161,1074]" captionTargetPageId="2" captionText="Fig. 1. — Melanophylla angustior: flowering shoot and flower. (Rabenantoandro & McPherson 774)." figureDoi="http://doi.org/10.5281/zenodo.5180849" httpUri="https://zenodo.org/record/5180849/files/figure.png" pageId="1" pageNumber="264">Fig. 1</figureCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C342F79E3B151057FD14FC8FFB42FCB7" pageId="1" pageNumber="264" type="discussion">
|
||||
<paragraph id="8BE7A4153B151057FD14FC8FFB42FCB7" blockId="1.[714,1247,814,1669]" pageId="1" pageNumber="264">
|
||||
The new species is unique within its genus in its combination of narrow leaf blades (
|
||||
<quantity id="4CA009F03B151057FB89FCEEFB28FA3C" box="[1145,1246,845,872]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="1" pageNumber="264" unit="cm" value="6.25" valueMax="7.5" valueMin="5.0">5-7.5 cm</quantity>
|
||||
wide, the length/width ratio 4/1-5/1), relatively short petioles (
|
||||
<quantity id="4CA009F03B151057FC83FC2BFC2DFAFF" box="[883,987,906,933]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.7" metricValueMin="3.0" pageId="1" pageNumber="264" unit="cm" value="3.85" valueMax="4.7" valueMin="3.0">3-4.7 cm</quantity>
|
||||
long, the length/blade length ratio c. 1/5-1/7), and sessile or subsessile, yellowish white flowers. In
|
||||
<bibRefCitation id="EFC9D9E43B151057FC04FC69FB2CFAB8" author="SCHATZ G. E. & LOWRY II P. P. & WOLF A. - E." box="[1012,1242,968,995]" pageId="1" pageNumber="264" pagination="233 - 242" refId="ref1541" refString="SCHATZ G. E., LOWRY II P. P. & WOLF A. - E. 1998. - Endemic families of Madagascar. I. A synoptic revision of Melanophylla Baker (Melanophyllaceae). Adansonia, ser. 3, 20: 233 - 242." type="book chapter" year="1998">SCHATZ et al. (1998)</bibRefCitation>
|
||||
, the new species would key either to the couplet separating
|
||||
<taxonomicName id="4C58DF963B151057FCCEFBA5FBE4FD7B" box="[830,1042,1029,1056]" class="Magnoliopsida" family="Torricelliaceae" genus="Melanophylla" kingdom="Plantae" order="Apiales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="madagascariensis">
|
||||
<emphasis id="B92C78073B151057FCCEFBA5FBE4FD7B" box="[830,1042,1029,1056]" italics="true" pageId="1" pageNumber="264">M. madagascariensis</emphasis>
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="4C58DF963B151057FBBBFBA5FB48FD45" box="[1099,1214,1029,1054]" class="Magnoliopsida" family="Torricelliaceae" genus="Melanophylla" kingdom="Plantae" order="Apiales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="perrieri">
|
||||
<emphasis id="B92C78073B151057FBBBFBA5FB48FD45" box="[1099,1214,1029,1054]" italics="true" pageId="1" pageNumber="264">M. perrieri</emphasis>
|
||||
</taxonomicName>
|
||||
(if the leaves were considered coriaceous), or else to
|
||||
<taxonomicName id="4C58DF963B151057FD3AFBE3FC95FD06" box="[714,867,1091,1117]" class="Magnoliopsida" family="Torricelliaceae" genus="Melanophylla" kingdom="Plantae" order="Apiales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="aucubifolia">
|
||||
<emphasis id="B92C78073B151057FD3AFBE3FC95FD06" box="[714,867,1091,1117]" italics="true" pageId="1" pageNumber="264">M. aucubifolia</emphasis>
|
||||
</taxonomicName>
|
||||
(if the leaves were considered chartaceous). However,
|
||||
<taxonomicName id="4C58DF963B151057FC50FBC1FB83FD27" box="[928,1141,1121,1148]" class="Magnoliopsida" family="Torricelliaceae" genus="Melanophylla" kingdom="Plantae" order="Apiales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="madagascariensis">
|
||||
<emphasis id="B92C78073B151057FC50FBC1FB83FD27" box="[928,1141,1121,1148]" italics="true" pageId="1" pageNumber="264">M. madagascariensis</emphasis>
|
||||
</taxonomicName>
|
||||
differs in having wider leaf blades (length/width ratio c. 2/1), thicker and longer petioles (petiole/blade ratio 1/2-1/3), evident pedicels (
|
||||
<quantity id="4CA009F03B151057FBCAFB1EFB62FD8C" box="[1082,1172,1213,1240]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="264" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1-3 mm</quantity>
|
||||
long), and smaller flowers;
|
||||
<taxonomicName id="4C58DF963B151057FC5EFB7DFB64FDAE" box="[942,1170,1244,1271]" class="Magnoliopsida" family="Torricelliaceae" genus="Melanophylla" kingdom="Plantae" order="Apiales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="perrieri">
|
||||
<emphasis id="B92C78073B151057FC5EFB7DFB64FDAE" box="[942,1170,1244,1271]" italics="true" pageId="1" pageNumber="264">Melanophylla perrieri</emphasis>
|
||||
</taxonomicName>
|
||||
differs in having wider leaf blades (length/width ratio c. 3/1), pedicels
|
||||
<quantity id="4CA009F03B151057FC9EFABAFC3DFC68" box="[878,971,1305,1332]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="1" pageNumber="264" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1-3 mm</quantity>
|
||||
long, and pink flowers;
|
||||
<taxonomicName id="4C58DF963B151057FD3AFA99FC2FFC08" box="[714,985,1337,1363]" class="Magnoliopsida" family="Torricelliaceae" genus="Melanophylla" kingdom="Plantae" order="Apiales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="aucubifolia">
|
||||
<emphasis id="B92C78073B151057FD3AFA99FC2FFC08" box="[714,985,1337,1363]" italics="true" pageId="1" pageNumber="264">Melanophylla aucubifolia</emphasis>
|
||||
</taxonomicName>
|
||||
differs in having wider leaf blades (length/width ratio c. 3/1), relatively longer petioles (petiole/blade ratio c. 1/3), and pedicels
|
||||
<quantity id="4CA009F03B151057FCB3FA35FC46FCF5" box="[835,944,1428,1455]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="264" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3-5 mm</quantity>
|
||||
long. Geographically,
|
||||
<taxonomicName id="4C58DF963B151057FD3AFA13FC3DFC97" authorityName="McPherson & Rabenantoandro" authorityYear="2002" box="[714,971,1459,1485]" class="Magnoliopsida" family="Torricelliaceae" genus="Melanophylla" kingdom="Plantae" order="Apiales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="angustior">
|
||||
<emphasis id="B92C78073B151057FD3AFA13FC3DFC97" box="[714,971,1459,1485]" italics="true" pageId="1" pageNumber="264">Melanophylla angustior</emphasis>
|
||||
</taxonomicName>
|
||||
occurs well south of all known populations of the above three species.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C342F79E3B151057FD14FA51FB55FFDE" pageId="1" pageNumber="264" type="conservation">
|
||||
<paragraph id="8BE7A4153B151057FD14FA51FB55FFDE" blockId="1.[714,1247,814,1669]" pageId="1" pageNumber="264">
|
||||
CONSERVATION STATUS. — Given that the new species appears to be infrequent in the single, but stable, small forest from which it is known, we consider it Endangered, as it satisfies the IUCN’s criterion D for that category (
|
||||
<bibRefCitation id="EFC9D9E43B151057FBFBF9CBFB61FFDE" author="IUCN" box="[1035,1175,1643,1669]" pageId="1" pageNumber="264" refId="ref1510" refString="IUCN 2001. - IUCN Red List Categories and Criteria: Version 3. 1. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, UK." type="book" year="2001">IUCN 2001</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
467
data/0B/66/87/0B66879F91700510BCF7D311619EFE01.xml
Normal file
467
data/0B/66/87/0B66879F91700510BCF7D311619EFE01.xml
Normal file
|
|
@ -0,0 +1,467 @@
|
|||
<document id="1C9B8370FF4581F771C5F6A98C17153C" ID-DOI="10.11646/zootaxa.5609.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="15215020" ID-ZooBank="5F98331A-A9B1-4825-A477-5628EA14B6CA" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1744663275477" checkinUser="plazi" docAuthor="Ahmed, Shakoor, Banerjee, Dhriti & Marimuthu, Nithyanandam" docDate="2025" docId="0B66879F91700510BCF7D311619EFE01" docLanguage="en" docName="zootaxa.5609.2.5.pdf" docOrigin="Zootaxa 5609 (2)" docSource="https://doi.org/10.11646/zootaxa.5609.2.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Priodoscolex maduraiensis Ahmed & Marimuthu 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="245" masterDocId="F75FFFE791760519BC60D27E6514FFAC" masterDocTitle="Two new earthworm species of the genera Eutyphoeus Michaelsen, 1900 and Priodoscolex Gates, 1940 (Crassiclitellata: Acanthodrilidae) from India" masterLastPageNumber="248" masterPageNumber="236" pageNumber="242" updateTime="1744809129855" updateUser="jonas" zenodo-license-document="CLOSED">
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<mods:mods id="6F670E1ED7AC670B3E42C3E96E6F3B4B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B116891B46BFAC7268E0A33F240A9A71">Two new earthworm species of the genera Eutyphoeus Michaelsen, 1900 and Priodoscolex Gates, 1940 (Crassiclitellata: Acanthodrilidae) from India</mods:title>
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<mods:namePart id="81138087361859B99149352343DD19C8">Ahmed, Shakoor</mods:namePart>
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<mods:affiliation id="70B4C4DDE014E899BD2CE2D19F78C3BB">Zoological Survey of India (Ministry of Environment, Forest and Climate Change, Government of India), FPS Building, Indian Museum complex, Kolkata 700016, India.</mods:affiliation>
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</mods:role>
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<mods:namePart id="BA61DFF55533675EC55D0F9F635CC3E6">Banerjee, Dhriti</mods:namePart>
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<mods:affiliation id="3EA226E8A6E4D538F86128E07C4565A6">Zoological Survey of India (Ministry of Environment, Forest and Climate Change, Government of India), M Block, New Alipore, Kolkata 700053, India.</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="D0C6806BC5A3D97DC3751A9D334C8377">Marimuthu, Nithyanandam</mods:namePart>
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<mods:affiliation id="6734BFE0B5609CC198F6BB8BB7CA6B44">Zoological Survey of India (Ministry of Environment, Forest and Climate Change, Government of India), FPS Building, Indian Museum complex, Kolkata 700016, India.</mods:affiliation>
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<mods:title id="62D66F10BC15E46D006078A560F1FD5B">Zootaxa</mods:title>
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<mods:part id="1D974D60109DCFEC6B309C0598321219">
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<mods:date id="F4800A83775EAB2E8E7E507208856D00">2025</mods:date>
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<mods:number id="5B21722C7D28085C9C0AB3778DCFD205">2025-03-20</mods:number>
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<mods:number id="7AD8886AB53B3991CA1AFC0CEB9C38E2">5609</mods:number>
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<mods:url id="E99F97EFF8BDF419B8B8AF43B91BC929">https://doi.org/10.11646/zootaxa.5609.2.5</mods:url>
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<treatment id="0B66879F91700510BCF7D311619EFE01" LSID="urn:lsid:plazi:treatment:0B66879F91700510BCF7D311619EFE01" httpUri="http://treatment.plazi.org/id/0B66879F91700510BCF7D311619EFE01" lastPageId="9" lastPageNumber="245" pageId="6" pageNumber="242">
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<subSubSection id="CBD565029170051FBCF7D311662BFE26" box="[151,831,367,394]" pageId="6" pageNumber="242" type="nomenclature">
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<paragraph id="837036899170051FBCF7D311662BFE26" blockId="6.[151,1163,367,466]" box="[151,831,367,394]" pageId="6" pageNumber="242">
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<taxonomicName id="44CF4D0A9170051FBCF7D31167C2FE25" ID-CoL="KTTX5" authority="Ahmed & Marimuthu" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[151,726,367,394]" class="Clitellata" family="Acanthodrilidae" genus="Priodoscolex" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="242" phylum="Annelida" rank="species" species="maduraiensis" status="sp. nov.">
|
||||
<emphasis id="B1BBEA9B9170051FBCF7D31164CAFE26" bold="true" box="[151,478,367,394]" italics="true" pageId="6" pageNumber="242">Priodoscolex maduraiensis</emphasis>
|
||||
<emphasis id="B1BBEA9B9170051FBD85D30F67C2FE25" bold="true" box="[485,726,369,393]" pageId="6" pageNumber="242">Ahmed & Marimuthu</emphasis>
|
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</taxonomicName>
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<emphasis id="B1BBEA9B9170051FBEB6D30F662BFE26" bold="true" box="[726,831,368,394]" pageId="6" pageNumber="242">
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,
|
||||
<taxonomicNameLabel id="AA8857E09170051FBE81D30E662BFE26" box="[737,831,368,394]" pageId="6" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD565029170051FBCF7D3ED641FFE7E" pageId="6" pageNumber="242" type="description">
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<paragraph id="837036899170051FBCF7D3ED619FFE02" blockId="6.[151,1163,367,466]" box="[151,1163,403,430]" pageId="6" pageNumber="242">
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<uuid id="F7690C5C9170051FBEE4D3ED619FFE02" box="[644,1163,403,430]" pageId="6" pageNumber="242">2F7495D8-B95E-4F48-B796-54C989FB3065</uuid>
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</uri>
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</paragraph>
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<paragraph id="837036899170051FBCF7D3C6641FFE7E" blockId="6.[151,1163,367,466]" box="[151,267,439,466]" pageId="6" pageNumber="242">
|
||||
(
|
||||
<figureCitation id="1BF42A0C9170051FBCFFD3C66417FE7E" box="[159,259,439,466]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="6.[151,250,1791,1815]" captionStartId-1="7.[151,250,1899,1923]" captionTargetBox-0="[177,1411,886,1768]" captionTargetBox-1="[164,1423,489,1876]" captionTargetId-0="figure-196@6.[177,1411,886,1768]" captionTargetId-1="figure-187@7.[164,1423,489,1876]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 4. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view C. prostomium, D. showing male pores and female pores. Abbreviations: pr, prostomium; fp, female pores, cl, clitellum; mp, male pores; spp, spermathecal pores." captionText-1="FIGURE 5. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. internal anatomy (anterior), B. Spermathecae. C. Calciferous glands and prostates glands. Abbreviations: gz, gizzard; di, diverticulum; am, ampulla; sv–xi, seminal vesicles of segment 11, sv–xii, seminal vesicle of segment 12, ts–x, male funnels of segment 10; ts–xi, male funnels of segment 11; cg, calciferous glands; pd, prostatic duct; pg, prostate glands." figureDoi-0="http://doi.org/10.5281/zenodo.15215030" figureDoi-1="http://doi.org/10.5281/zenodo.15215032" httpUri-0="https://zenodo.org/record/15215030/files/figure.png" httpUri-1="https://zenodo.org/record/15215032/files/figure.png" pageId="6" pageNumber="242">Figs 4–5</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD565029170051FBCF7D07E61DCFD5E" pageId="6" pageNumber="242" type="materials_examined">
|
||||
<paragraph id="837036899170051FBCF7D07E6009FDCE" blockId="6.[151,1436,511,862]" pageId="6" pageNumber="242">
|
||||
<emphasis id="B1BBEA9B9170051FBCF7D07E6428FDB5" bold="true" box="[151,316,511,538]" pageId="6" pageNumber="242">Type material</emphasis>
|
||||
.
|
||||
<materialsCitation id="33A73CD49170051FBD29D07E61EDFD92" collectingDate="2004-11-08" collectionCode="ZSI -GNC" collectorName="C. K. Mandal & S. Dhani" country="India" county="Madurai" elevation="133" latitude="9.923514" location="Madurai" longLatPrecision="1" longitude="78.10858" pageId="6" pageNumber="242" specimenCode="An8036/2" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
|
||||
<emphasis id="B1BBEA9B9170051FBD29D07E64A4FDB6" box="[329,432,512,538]" italics="true" pageId="6" pageNumber="242">
|
||||
<typeStatus id="5C74882B9170051FBD29D07E64A4FDB6" box="[329,432,512,538]" pageId="6" pageNumber="242" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
. Clitellate (
|
||||
<collectionCode id="E5DEAE4C9170051FBE4FD38167B2FDB5" box="[559,678,511,537]" pageId="6" pageNumber="242">ZSI–GNC</collectionCode>
|
||||
–
|
||||
<specimenCode id="D3699EF29170051FBED5D3816632FDB5" box="[693,806,511,537]" pageId="6" pageNumber="242">An8036/2</specimenCode>
|
||||
),
|
||||
<location id="861060529170051FBF58D07E6683FDB6" LSID="urn:lsid:plazi:treatment:0B66879F91700510BCF7D311619EFE01:861060529170051FBF58D07E6683FDB6" box="[824,919,512,538]" country="India" county="Madurai" latitude="9.923514" longLatPrecision="1" longitude="78.10858" name="Madurai" pageId="6" pageNumber="242" stateProvince="Tamil Nadu">Madurai</location>
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(
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<geoCoordinate id="E6FB504E9170051FBFC4D3816125FDB6" box="[932,1073,511,538]" degrees="9.923514" direction="north" orientation="latitude" pageId="6" pageNumber="242" precision="1" value="9.923514">9.923514°N</geoCoordinate>
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||||
,
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||||
<geoCoordinate id="E6FB504E9170051FB85AD38161C6FDB6" box="[1082,1234,511,538]" degrees="78.108583" direction="east" orientation="longitude" pageId="6" pageNumber="242" precision="1" value="78.10858">78.108583°E</geoCoordinate>
|
||||
),
|
||||
<collectingCounty id="6A114E059170051FB883D07E6056FDB6" box="[1251,1346,512,538]" pageId="6" pageNumber="242">Madurai</collectingCounty>
|
||||
district,
|
||||
<collectingRegion id="410BF86B9170051FBCF7D05A640BFD92" box="[151,287,548,574]" country="India" name="Tamil Nadu" pageId="6" pageNumber="242">Tamil Nadu</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="FBD876199170051FBD4BD05A647CFD92" box="[299,360,548,574]" name="India" pageId="6" pageNumber="242">India</collectingCountry>
|
||||
;
|
||||
<quantity id="44379B6C9170051FBD14D05A64A7FD92" box="[372,435,548,574]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="6" pageNumber="242" unit="m" value="133.0">
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<elevation id="08E2D1BA9170051FBD14D05A64A7FD92" box="[372,435,548,574]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="6" pageNumber="242" unit="m" value="133.0">133m</elevation>
|
||||
</quantity>
|
||||
above mean sea level;
|
||||
<date id="F77110499170051FBEDAD05A669EFD92" box="[698,906,548,574]" pageId="6" pageNumber="242" value="2004-11-08">
|
||||
<collectingDate id="E735E9A19170051FBEDAD05A669EFD92" box="[698,906,548,574]" pageId="6" pageNumber="242" value="2004-11-08">8 November 2004</collectingDate>
|
||||
</date>
|
||||
; leg.
|
||||
<collectorName id="2E3A535F9170051FBFA6D05A614CFD92" box="[966,1112,548,574]" pageId="6" pageNumber="242">C.K. Mandal</collectorName>
|
||||
and
|
||||
<collectorName id="2E3A535F9170051FB8EFD05A61E1FD92" box="[1167,1269,548,574]" pageId="6" pageNumber="242">S. Dhani</collectorName>
|
||||
.
|
||||
</materialsCitation>
|
||||
<materialsCitation id="33A73CD49170051FB960D05A600DFDCE" collectingDate="2004-11-08" collectionCode="ZSI -GNC" collectorName="C. K. Mandal & S. Dhani" country="India" county="Madurai" elevation="133" latitude="9.923514" location="Madurai" longLatPrecision="1" longitude="78.10858" pageId="6" pageNumber="242" specimenCode="An8037/2" specimenCount="16" stateProvince="Tamil Nadu" typeStatus="paratype">
|
||||
<emphasis id="B1BBEA9B9170051FB960D05A6066FD91" box="[1280,1394,548,573]" italics="true" pageId="6" pageNumber="242">
|
||||
<typeStatus id="5C74882B9170051FB960D05A6066FD91" box="[1280,1394,548,573]" pageId="6" pageNumber="242" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
.
|
||||
<specimenCount id="95C9FD009170051FB9E0D05A6088FD92" box="[1408,1436,548,574]" pageId="6" pageNumber="242">14</specimenCount>
|
||||
clitellates,
|
||||
<specimenCount id="95C9FD009170051FBD71D036640BFDCE" box="[273,287,584,610]" pageId="6" pageNumber="242">2</specimenCount>
|
||||
aclitellates and 16 tail parts (
|
||||
<collectionCode id="E5DEAE4C9170051FBE07D03667C9FDCE" box="[615,733,584,610]" pageId="6" pageNumber="242">ZSI–GNC</collectionCode>
|
||||
–
|
||||
<specimenCode id="D3699EF29170051FBE8CD036664AFDCE" box="[748,862,584,610]" pageId="6" pageNumber="242">An8037/2</specimenCode>
|
||||
), collection details as same as holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
<paragraph id="837036899170051FBCA7D01561DCFD5E" blockId="6.[151,1436,511,862]" pageId="6" pageNumber="242">
|
||||
<emphasis id="B1BBEA9B9170051FBCA7D0156468FD2A" bold="true" box="[199,380,619,646]" pageId="6" pageNumber="242">
|
||||
<emphasis id="B1BBEA9B9170051FBCA7D0156461FD2A" bold="true" box="[199,373,619,646]" italics="true" pageId="6" pageNumber="242">Other material</emphasis>
|
||||
.
|
||||
</emphasis>
|
||||
<materialsCitation id="33A73CD49170051FBDE7D01565C3FD62" collectingDate="2004-11-30" collectionCode="ZSI -GNC" collectorName="A. Mishra" country="India" county="Salem" elevation="1438" latitude="11.796355" location="Yercaud" longLatPrecision="1" longitude="78.21641" municipality="Nagalur village" pageId="6" pageNumber="242" specimenCode="An3038/2" specimenCount="5" stateProvince="Tamil Nadu">
|
||||
<specimenCount id="95C9FD009170051FBDE7D0156481FD2A" box="[391,405,619,646]" pageId="6" pageNumber="242">5</specimenCount>
|
||||
aclitellates (
|
||||
<collectionCode id="E5DEAE4C9170051FBE4AD01267B5FD2A" box="[554,673,620,646]" pageId="6" pageNumber="242">ZSI–GNC</collectionCode>
|
||||
–
|
||||
<specimenCode id="D3699EF29170051FBED0D0126636FD2A" box="[688,802,620,646]" pageId="6" pageNumber="242">An3038/2</specimenCode>
|
||||
),
|
||||
<location id="861060529170051FBF57D0126687FD2A" LSID="urn:lsid:plazi:treatment:0B66879F91700510BCF7D311619EFE01:861060529170051FBF57D0126687FD2A" box="[823,915,620,646]" country="India" county="Salem" latitude="11.796355" longLatPrecision="1" longitude="78.21641" municipality="Nagalur village" name="Yercaud" pageId="6" pageNumber="242" stateProvince="Tamil Nadu">Yercaud</location>
|
||||
(
|
||||
<geoCoordinate id="E6FB504E9170051FBFC6D0126154FD2A" box="[934,1088,620,646]" degrees="11.774038" direction="north" orientation="latitude" pageId="6" pageNumber="242" precision="1" value="11.774038">11.774038°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="E6FB504E9170051FB82FD01561F0FD29" box="[1103,1252,619,645]" degrees="78.216717" direction="east" orientation="longitude" pageId="6" pageNumber="242" precision="1" value="78.21672">78.216717°E</geoCoordinate>
|
||||
) (habitat coffee plantation),
|
||||
<collectingCounty id="6A114E059170051FBD7DD0EE6477FD06" box="[285,355,656,682]" pageId="6" pageNumber="242">Salem</collectingCounty>
|
||||
district,
|
||||
<collectingRegion id="410BF86B9170051FBDA3D0EE675DFD06" box="[451,585,656,682]" country="India" name="Tamil Nadu" pageId="6" pageNumber="242">Tamil Nadu</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="FBD876199170051FBE33D0EE6784FD06" box="[595,656,656,682]" name="India" pageId="6" pageNumber="242">India</collectingCountry>
|
||||
;
|
||||
<quantity id="44379B6C9170051FBEFAD0EE67FAFD06" box="[666,750,656,682]" metricMagnitude="3" metricUnit="m" metricValue="1.438" pageId="6" pageNumber="242" unit="m" value="1438.0">
|
||||
<elevation id="08E2D1BA9170051FBEFAD0EE67FAFD06" box="[666,750,656,682]" metricMagnitude="3" metricUnit="m" metricValue="1.438" pageId="6" pageNumber="242" unit="m" value="1438.0">1438 m</elevation>
|
||||
</quantity>
|
||||
above mean sea level;
|
||||
<date id="F77110499170051FBF8DD0EE61D3FD06" box="[1005,1223,656,682]" pageId="6" pageNumber="242" value="2004-11-30">
|
||||
<collectingDate id="E735E9A19170051FBF8DD0EE61D3FD06" box="[1005,1223,656,682]" pageId="6" pageNumber="242" value="2004-11-30">30 November 2004</collectingDate>
|
||||
</date>
|
||||
; leg.
|
||||
<collectorName id="2E3A535F9170051FB89ED0EE607AFD06" box="[1278,1390,656,682]" pageId="6" pageNumber="242">A. Mishra</collectorName>
|
||||
and party.
|
||||
</materialsCitation>
|
||||
<materialsCitation id="33A73CD49170051FBCBFD0CA61D0FD5E" collectingDate="2004-11-30" collectionCode="ZSI -GNC" collectorName="A. Mishra" country="India" county="Salem" elevation="1412" latitude="11.796355" location="Nagalur village" longLatPrecision="1" longitude="78.21641" municipality="Yercaud" pageId="6" pageNumber="242" specimenCode="An3039/2" specimenCount="1" stateProvince="Tamil Nadu">
|
||||
<specimenCount id="95C9FD009170051FBCBFD0CA65F9FD62" box="[223,237,692,718]" pageId="6" pageNumber="242">1</specimenCount>
|
||||
aclitellate (
|
||||
<collectionCode id="E5DEAE4C9170051FBD14D0CA64FDFD62" box="[372,489,692,718]" pageId="6" pageNumber="242">ZSI–GNC</collectionCode>
|
||||
–
|
||||
<specimenCode id="D3699EF29170051FBD98D0CA677CFD62" box="[504,616,692,718]" pageId="6" pageNumber="242">An3039/2</specimenCode>
|
||||
)
|
||||
<location id="861060529170051FBE19D0CA663CFD62" LSID="urn:lsid:plazi:treatment:0B66879F91700510BCF7D311619EFE01:861060529170051FBE19D0CA663CFD62" box="[633,808,692,718]" country="India" county="Salem" latitude="11.796355" longLatPrecision="1" longitude="78.21641" municipality="Yercaud" name="Nagalur village" pageId="6" pageNumber="242" stateProvince="Tamil Nadu">Nagalur village</location>
|
||||
(
|
||||
<geoCoordinate id="E6FB504E9170051FBF59D0CD66C7FD62" box="[825,979,691,718]" degrees="11.796355" direction="north" orientation="latitude" pageId="6" pageNumber="242" precision="1" value="11.796355">11.796355°N</geoCoordinate>
|
||||
,
|
||||
<geoCoordinate id="E6FB504E9170051FBF80D0CD616CFD61" box="[992,1144,691,717]" degrees="78.216404" direction="east" orientation="longitude" pageId="6" pageNumber="242" precision="1" value="78.21641">78.216404°E</geoCoordinate>
|
||||
),
|
||||
<collectingMunicipality id="6314ACF39170051FB8ECD0CA61FFFD62" box="[1164,1259,692,718]" pageId="6" pageNumber="242">Yercaud</collectingMunicipality>
|
||||
,
|
||||
<collectingCounty id="6A114E059170051FB898D0CA602AFD62" box="[1272,1342,692,718]" pageId="6" pageNumber="242">Salem</collectingCounty>
|
||||
district,
|
||||
<collectingRegion id="410BF86B9170051FBCF7D0A6640AFD5E" box="[151,286,728,754]" country="India" name="Tamil Nadu" pageId="6" pageNumber="242">Tamil Nadu</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="FBD876199170051FBD4AD0A66473FD5E" box="[298,359,728,754]" name="India" pageId="6" pageNumber="242">India</collectingCountry>
|
||||
;
|
||||
<quantity id="44379B6C9170051FBD12D0A664D3FD5E" box="[370,455,728,754]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="6" pageNumber="242" unit="m" value="1412.0">
|
||||
<elevation id="08E2D1BA9170051FBD12D0A664D3FD5E" box="[370,455,728,754]" metricMagnitude="3" metricUnit="m" metricValue="1.412" pageId="6" pageNumber="242" unit="m" value="1412.0">1412 m</elevation>
|
||||
</quantity>
|
||||
above mean sea level;
|
||||
<date id="F77110499170051FBEABD0A666BCFD5E" box="[715,936,728,754]" pageId="6" pageNumber="242" value="2004-11-30">
|
||||
<collectingDate id="E735E9A19170051FBEABD0A666BCFD5E" box="[715,936,728,754]" pageId="6" pageNumber="242" value="2004-11-30">30 November 2004</collectingDate>
|
||||
</date>
|
||||
; leg.
|
||||
<collectorName id="2E3A535F9170051FBF81D0A66146FD5E" box="[993,1106,728,754]" pageId="6" pageNumber="242">A. Mishra</collectorName>
|
||||
and party
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD565029170051FBCA7D08261A5FCF2" pageId="6" pageNumber="242" type="etymology">
|
||||
<paragraph id="837036899170051FBCA7D08261A5FCF2" blockId="6.[151,1436,511,862]" pageId="6" pageNumber="242">
|
||||
<emphasis id="B1BBEA9B9170051FBCA7D082645AFCBA" bold="true" box="[199,334,764,790]" pageId="6" pageNumber="242">Etymology.</emphasis>
|
||||
The species epithet
|
||||
<taxonomicName id="44CF4D0A9170051FBE23D08267F9FCBA" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[579,749,764,790]" class="Clitellata" family="Acanthodrilidae" genus="Priodoscolex" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="242" phylum="Annelida" rank="species" species="maduraiensis">
|
||||
“
|
||||
<emphasis id="B1BBEA9B9170051FBE30D08267F9FCBA" box="[592,749,764,790]" italics="true" pageId="6" pageNumber="242">maduraiensis</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B1BBEA9B9170051FBE8CD08267E2FCBA" box="[748,758,764,790]" italics="true" pageId="6" pageNumber="242">”</emphasis>
|
||||
is derived from the name of the city, Madurai where the specimens have been collected. The rationale behind the species’ name is the history of the city, Madurai had been recorded by the Greek envoy, Megasthenes as early as the third century BCE (
|
||||
<bibRefCitation id="E75E4B789170051FB862D13A61B0FCF2" author="Zvelebil, K. V." box="[1026,1188,836,862]" pageId="6" pageNumber="242" refId="ref8251" refString="Zvelebil, K. V. (1992) Companion studies to the history of Tamil literature. Leiden, Brill, 277 pp." type="book" year="1992">Zvelebil 1992</bibRefCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="D7B066019170051FBCF7D4816475F8F3" ID-DOI="http://doi.org/10.5281/zenodo.15215030" ID-Zenodo-Dep="15215030" httpUri="https://zenodo.org/record/15215030/files/figure.png" pageId="6" pageNumber="242" startId="6.[151,250,1791,1815]" targetBox="[177,1411,886,1768]" targetPageId="6" targetType="figure">
|
||||
<paragraph id="837036899170051FBCF7D4816475F8F3" blockId="6.[151,1436,1791,1887]" pageId="6" pageNumber="242">
|
||||
<emphasis id="B1BBEA9B9170051FBCF7D4816401F8BB" bold="true" box="[151,277,1791,1815]" pageId="6" pageNumber="242">FIGURE 4.</emphasis>
|
||||
<taxonomicName id="44CF4D0A9170051FBD7ED4816636F8BB" authority="Ahmed & Marimuthu" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[286,802,1791,1816]" class="Clitellata" family="Acanthodrilidae" genus="Priodoscolex" kingdom="Animalia" order="Crassiclitellata" pageId="6" pageNumber="242" phylum="Annelida" rank="species" species="maduraiensis" status="sp. nov.">
|
||||
<emphasis id="B1BBEA9B9170051FBD7ED4816722F8BB" box="[286,566,1791,1815]" italics="true" pageId="6" pageNumber="242">Priodoscolex maduraiensis</emphasis>
|
||||
Ahmed & Marimuthu
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="AA8857E09170051FBF4ED57E666BF8BB" box="[814,895,1792,1815]" pageId="6" pageNumber="242" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="B1BBEA9B9170051FBFE7D481668DF8BB" bold="true" box="[903,921,1791,1815]" pageId="6" pageNumber="242">A</emphasis>
|
||||
. lateral view,
|
||||
<emphasis id="B1BBEA9B9170051FB84FD57E6154F8B4" bold="true" box="[1071,1088,1792,1816]" pageId="6" pageNumber="242">B</emphasis>
|
||||
. ventral view
|
||||
<emphasis id="B1BBEA9B9170051FB8B9D48161FFF8BB" bold="true" box="[1241,1259,1791,1815]" pageId="6" pageNumber="242">C</emphasis>
|
||||
. prostomium,
|
||||
<emphasis id="B1BBEA9B9170051FB9E4D57E6082F8B4" bold="true" box="[1412,1430,1792,1816]" pageId="6" pageNumber="242">D</emphasis>
|
||||
. showing male pores and female pores. Abbreviations:
|
||||
<emphasis id="B1BBEA9B9170051FBEB0D55A67F3F897" box="[720,743,1828,1851]" italics="true" pageId="6" pageNumber="242">pr</emphasis>
|
||||
, prostomium;
|
||||
<emphasis id="B1BBEA9B9170051FBFE0D55D6680F897" box="[896,916,1827,1851]" italics="true" pageId="6" pageNumber="242">fp</emphasis>
|
||||
, female pores,
|
||||
<emphasis id="B1BBEA9B9170051FB858D55D615EF897" box="[1080,1098,1827,1851]" italics="true" pageId="6" pageNumber="242">cl</emphasis>
|
||||
, clitellum;
|
||||
<emphasis id="B1BBEA9B9170051FB8A3D55A61F6F897" box="[1219,1250,1828,1851]" italics="true" pageId="6" pageNumber="242">mp</emphasis>
|
||||
, male pores;
|
||||
<emphasis id="B1BBEA9B9170051FB913D55A6082F897" box="[1395,1430,1828,1851]" italics="true" pageId="6" pageNumber="242">spp</emphasis>
|
||||
, spermathecal pores.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="CBD5650291700511BCA7D5CF6023FF55" lastPageId="8" lastPageNumber="244" pageId="6" pageNumber="242" type="diagnosis">
|
||||
<paragraph id="8370368991700511BCA7D5CF6023FF55" blockId="6.[151,1436,1969,2032]" lastBlockId="8.[151,1437,151,717]" lastPageId="8" lastPageNumber="244" pageId="6" pageNumber="242">
|
||||
<emphasis id="B1BBEA9B9170051FBCA7D5CF642EF867" bold="true" box="[199,314,1969,1995]" pageId="6" pageNumber="242">Diagnosis</emphasis>
|
||||
. Large sized; length
|
||||
<quantity id="44379B6C9170051FBE7FD5CC67B7F860" box="[543,675,1970,1996]" metricMagnitude="-1" metricUnit="m" metricValue="1.86" metricValueMax="2.8" metricValueMin="0.9199999999999999" pageId="6" pageNumber="242" unit="mm" value="186.0" valueMax="280.0" valueMin="92.0">92–280mm</quantity>
|
||||
; width
|
||||
<quantity id="44379B6C9170051FBE91D5CF665DF867" box="[753,841,1969,1995]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="6" pageNumber="242" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6mm</quantity>
|
||||
, number of segments 211–389. Segment 7 biannulate, 8–12 with light annuli (two anterior and two posteriors of setal arc and one with setal arc). Colour, light brownish with reddish clitellum (preserved specimens). Prostomium prolobous. First dorsal pore in intersegmental furrow 10/11. Setae perichaetine; on the pre-clitellar region there is no dorsal or ventral setal break, post clitellum dorsal
|
||||
<emphasis id="B1BBEA9B9171051EB917D2C56099FF78" box="[1399,1421,187,212]" italics="true" pageId="7" pageNumber="243">zz</emphasis>
|
||||
= 2–2.5
|
||||
<emphasis id="B1BBEA9B9171051EBCB6D2A165F9FF54" box="[214,237,223,248]" italics="true" pageId="7" pageNumber="243">yz</emphasis>
|
||||
; except segment 18, the ventral most setae on segments 12–21 some time on segment 11 and 22 displaced anterior and posterior to the setal line; ventral most setae on segment 18 absent. Clitellum prominent, reddish colour, saddle shaped on segments 13–18, dorsal pores absent, 5–6 setae on each side visible ventral part of clitellum. Female pores paired in a small bulge oval area on segment 14 anterior to setal arc. Male pores closely paired on segment 18, no setae between male pores. Spermathecal pores 3 pairs, closely paired about the level of
|
||||
<emphasis id="B1BBEA9B9171051EB970D3116038FE25" box="[1296,1324,367,393]" italics="true" pageId="7" pageNumber="243">ab</emphasis>
|
||||
setal line, in segments 7,8 and 9, just posterior to the intersegmental furrow 6/7/8/9. Genital markings absent, except light bulging of ventral area on segments, 14,15,16, at the place of displaced setae. Typhlosole begin in segments 26–28. Spermathecae, three pairs in segments 7,8, and 9 sometime anterior diverted in segments 6, 7, and 8; ampulla club shaped with digitiform ectal diverticulum. Calciferous glands 4 pairs in segments 14, 15, 16, and 17, the glands in segments 15–17 are bilobed, whereas glands of segment 14 are small and single lobe. Penial setae absent.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="D7B066019171051EBCF7D51567CBF843" ID-DOI="http://doi.org/10.5281/zenodo.15215032" ID-Zenodo-Dep="15215032" httpUri="https://zenodo.org/record/15215032/files/figure.png" pageId="7" pageNumber="243" startId="7.[151,250,1899,1923]" targetBox="[164,1423,489,1876]" targetPageId="7" targetType="figure">
|
||||
<paragraph id="837036899171051EBCF7D51567CBF843" blockId="7.[151,1436,1899,2031]" pageId="7" pageNumber="243">
|
||||
<emphasis id="B1BBEA9B9171051EBCF7D5156401F828" bold="true" box="[151,277,1899,1924]" pageId="7" pageNumber="243">FIGURE 5.</emphasis>
|
||||
<taxonomicName id="44CF4D0A9171051EBD7DD5156634F82F" authority="Ahmed & Marimuthu" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[285,800,1899,1924]" class="Clitellata" family="Acanthodrilidae" genus="Priodoscolex" kingdom="Animalia" order="Crassiclitellata" pageId="7" pageNumber="243" phylum="Annelida" rank="species" species="maduraiensis" status="sp. nov.">
|
||||
<emphasis id="B1BBEA9B9171051EBD7DD5156720F82F" box="[285,564,1899,1923]" italics="true" pageId="7" pageNumber="243">Priodoscolex maduraiensis</emphasis>
|
||||
Ahmed & Marimuthu
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="AA8857E09171051EBF4CD5126669F82F" box="[812,893,1900,1923]" pageId="7" pageNumber="243" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="B1BBEA9B9171051EBFE3D5156681F82F" bold="true" box="[899,917,1899,1923]" pageId="7" pageNumber="243">A</emphasis>
|
||||
. internal anatomy (anterior),
|
||||
<emphasis id="B1BBEA9B9171051EB8A7D51261CCF828" bold="true" box="[1223,1240,1900,1924]" pageId="7" pageNumber="243">B</emphasis>
|
||||
. Spermathecae.
|
||||
<emphasis id="B1BBEA9B9171051EB9E4D5156082F82F" bold="true" box="[1412,1430,1899,1923]" pageId="7" pageNumber="243">C</emphasis>
|
||||
. Calciferous glands and prostates glands. Abbreviations:
|
||||
<emphasis id="B1BBEA9B9171051EBEB9D5EE67E4F80B" box="[729,752,1936,1959]" italics="true" pageId="7" pageNumber="243">gz</emphasis>
|
||||
, gizzard;
|
||||
<emphasis id="B1BBEA9B9171051EBF37D5F1667FF80B" box="[855,875,1935,1959]" italics="true" pageId="7" pageNumber="243">di</emphasis>
|
||||
, diverticulum;
|
||||
<emphasis id="B1BBEA9B9171051EB869D5EE613CF80B" box="[1033,1064,1936,1959]" italics="true" pageId="7" pageNumber="243">am</emphasis>
|
||||
, ampulla;
|
||||
<emphasis id="B1BBEA9B9171051EB8F6D5EE61DEF80B" box="[1174,1226,1936,1959]" italics="true" pageId="7" pageNumber="243">sv–xi</emphasis>
|
||||
, seminal vesicles of segment 11,
|
||||
<emphasis id="B1BBEA9B9171051EBD7BD5CA6442F867" box="[283,342,1972,1995]" italics="true" pageId="7" pageNumber="243">sv–xii</emphasis>
|
||||
, seminal vesicle of segment 12,
|
||||
<emphasis id="B1BBEA9B9171051EBECCD5CA67C0F867" box="[684,724,1972,1995]" italics="true" pageId="7" pageNumber="243">ts–x</emphasis>
|
||||
, male funnels of segment 10;
|
||||
<emphasis id="B1BBEA9B9171051EB872D5CA6155F867" box="[1042,1089,1972,1995]" italics="true" pageId="7" pageNumber="243">ts–xi</emphasis>
|
||||
, male funnels of segment 11;
|
||||
<emphasis id="B1BBEA9B9171051EB91ED5CA6082F867" box="[1406,1430,1972,1995]" italics="true" pageId="7" pageNumber="243">cg</emphasis>
|
||||
, calciferous glands;
|
||||
<emphasis id="B1BBEA9B9171051EBD3CD5A96461F843" box="[348,373,2007,2031]" italics="true" pageId="7" pageNumber="243">pd</emphasis>
|
||||
, prostatic duct;
|
||||
<emphasis id="B1BBEA9B9171051EBE79D5A66726F843" box="[537,562,2008,2031]" italics="true" pageId="7" pageNumber="243">pg</emphasis>
|
||||
, prostate glands.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="CBD56502917E0510BCA7D37C6191FEB1" lastPageId="9" lastPageNumber="245" pageId="8" pageNumber="244" type="description">
|
||||
<paragraph id="83703689917E0511BCA7D37C6098FD61" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="244">
|
||||
<emphasis id="B1BBEA9B917E0511BCA7D37C644DFEB0" bold="true" box="[199,345,258,284]" pageId="8" pageNumber="244">Description.</emphasis>
|
||||
<emphasis id="B1BBEA9B917E0511BD07D37D64C4FEB1" box="[359,464,259,285]" italics="true" pageId="8" pageNumber="244">External:</emphasis>
|
||||
Colour light brownish with reddish clitellum (preserved specimen). Dimensions:
|
||||
<typeStatus id="5C74882B917E0511BCF7D3596410FEED" box="[151,260,295,321]" pageId="8" pageNumber="244" type="holotype">Holotype</typeStatus>
|
||||
: length
|
||||
<quantity id="44379B6C917E0511BD3DD35864AAFEED" box="[349,446,294,321]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" pageId="8" pageNumber="244" unit="mm" value="275.0">275 mm</quantity>
|
||||
; width
|
||||
<quantity id="44379B6C917E0511BE73D3586741FEED" box="[531,597,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="244" unit="mm" value="5.0">5 mm</quantity>
|
||||
at segment 8, 4.7 at clitellum and 4.2 at segment 20; number of segments 308;
|
||||
<typeStatus id="5C74882B917E0511BCB1D3356455FEC9" box="[209,321,331,357]" pageId="8" pageNumber="244" type="paratype">paratypes</typeStatus>
|
||||
: length
|
||||
<quantity id="44379B6C917E0511BDFBD3356709FEC9" box="[411,541,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="1.86" metricValueMax="2.8" metricValueMin="0.9199999999999999" pageId="8" pageNumber="244" unit="mm" value="186.0" valueMax="280.0" valueMin="92.0">92–280mm</quantity>
|
||||
, width
|
||||
<quantity id="44379B6C917E0511BE12D33567DEFEC9" box="[626,714,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="8" pageNumber="244" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6mm</quantity>
|
||||
, segment 211–389; other material: length
|
||||
<quantity id="44379B6C917E0511B8C9D3356038FEC9" box="[1193,1324,331,357]" metricMagnitude="-1" metricUnit="m" metricValue="1.525" metricValueMax="2.08" metricValueMin="0.97" pageId="8" pageNumber="244" unit="mm" value="152.5" valueMax="208.0" valueMin="97.0">97–208mm</quantity>
|
||||
, width
|
||||
<quantity id="44379B6C917E0511B9E0D33565FDFE25" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="8" pageNumber="244" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3– 3.5mm</quantity>
|
||||
, segments 316–376. Prostomium prolobous (
|
||||
<figureCitation id="1BF42A0C917E0511BE94D3116644FE25" box="[756,848,367,393]" captionStart="FIGURE 4" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[177,1411,886,1768]" captionTargetId="figure-196@6.[177,1411,886,1768]" captionTargetPageId="6" captionText="FIGURE 4. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view C. prostomium, D. showing male pores and female pores. Abbreviations: pr, prostomium; fp, female pores, cl, clitellum; mp, male pores; spp, spermathecal pores." figureDoi="http://doi.org/10.5281/zenodo.15215030" httpUri="https://zenodo.org/record/15215030/files/figure.png" pageId="8" pageNumber="244">Fig. 4C</figureCitation>
|
||||
). First dorsal pore in intersegmental furrow 10/11. Setae Perichaetine, 32 on segment 2, 74 on segment 6, 86 on segment 12, 96 on segment 19. On the pre-clitellar region there is no dorsal or ventral setal break, post clitellum dorsal
|
||||
<emphasis id="B1BBEA9B917E0511BFF4D3C966BEFE7C" box="[916,938,439,464]" italics="true" pageId="8" pageNumber="244">zz</emphasis>
|
||||
= 2
|
||||
<emphasis id="B1BBEA9B917E0511BFAFD3C966F2FE7C" box="[975,998,439,464]" italics="true" pageId="8" pageNumber="244">yz</emphasis>
|
||||
. Further, the ventral setae on segments 11–20, except segment 18 displaced anterior and posterior to the setal line. Ventral most setae on segment 18 absent. Clitellum prominent, reddish colour, saddle shaped on segments 13–18, dorsal pores absent, 5–6 setae on each side visible ventral on the clitellum thickening (
|
||||
<figureCitation id="1BF42A0C917E0511BE9CD05D66B5FD91" box="[764,929,547,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[177,1411,886,1768]" captionTargetId="figure-196@6.[177,1411,886,1768]" captionTargetPageId="6" captionText="FIGURE 4. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view C. prostomium, D. showing male pores and female pores. Abbreviations: pr, prostomium; fp, female pores, cl, clitellum; mp, male pores; spp, spermathecal pores." figureDoi="http://doi.org/10.5281/zenodo.15215030" httpUri="https://zenodo.org/record/15215030/files/figure.png" pageId="8" pageNumber="244">Fig. 4A, B, D</figureCitation>
|
||||
). Female pores paired, pre-setal on segment 14, nearly equi-distance between 13/14 and setal arc of segment 14. Male pores closely paired on segment 18, no setae between male pores, seminal grooves absent (
|
||||
<figureCitation id="1BF42A0C917E0511BEB5D0156644FD29" box="[725,848,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[177,1411,886,1768]" captionTargetId="figure-196@6.[177,1411,886,1768]" captionTargetPageId="6" captionText="FIGURE 4. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view C. prostomium, D. showing male pores and female pores. Abbreviations: pr, prostomium; fp, female pores, cl, clitellum; mp, male pores; spp, spermathecal pores." figureDoi="http://doi.org/10.5281/zenodo.15215030" httpUri="https://zenodo.org/record/15215030/files/figure.png" pageId="8" pageNumber="244">Fig. 4B, D</figureCitation>
|
||||
). Spermathecal pores 3 pairs, closely paired at
|
||||
<emphasis id="B1BBEA9B917E0511B93DD015606DFD29" box="[1373,1401,619,645]" italics="true" pageId="8" pageNumber="244">ab</emphasis>
|
||||
, in segments 7,8 and 9, just posterior to the intersegmental furrow 6/7/8/9 (
|
||||
<figureCitation id="1BF42A0C917E0511BFA4D0F16109FD05" box="[964,1053,655,681]" captionStart="FIGURE 4" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[177,1411,886,1768]" captionTargetId="figure-196@6.[177,1411,886,1768]" captionTargetPageId="6" captionText="FIGURE 4. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view C. prostomium, D. showing male pores and female pores. Abbreviations: pr, prostomium; fp, female pores, cl, clitellum; mp, male pores; spp, spermathecal pores." figureDoi="http://doi.org/10.5281/zenodo.15215030" httpUri="https://zenodo.org/record/15215030/files/figure.png" pageId="8" pageNumber="244">Fig. 4B</figureCitation>
|
||||
). Genital markings absent, except epithelium bulging at ventral areas on segments, 14,15,16 and 17, at place of displaced setae. Penial setae absent.
|
||||
</paragraph>
|
||||
<caption id="D7B06601917E0511BCF8D08266E3FCB8" box="[152,1015,764,788]" pageId="8" pageNumber="244" startId="8.[152,244,764,788]" targetBox="[163,1422,805,1519]" targetIsTable="true" targetPageId="8" targetType="table">
|
||||
<paragraph id="83703689917E0511BCF8D08266E3FCB8" blockId="8.[152,1015,764,788]" box="[152,1015,764,788]" pageId="8" pageNumber="244">
|
||||
<emphasis id="B1BBEA9B917E0511BCF8D082641BFCB8" bold="true" box="[152,271,764,788]" pageId="8" pageNumber="244">TABLE 1.</emphasis>
|
||||
Comparative analysis between the new species and described species
|
||||
</paragraph>
|
||||
</caption>
|
||||
<paragraph id="83703689917E0511BCC3D15B600AFA67" pageId="8" pageNumber="244">
|
||||
<table id="F1CFC429917EFAE6BCC3D15B609AFA43" box="[163,1422,805,1519]" gridcols="3" gridrows="11" pageId="8" pageNumber="244">
|
||||
<tr id="3DFF34CB917EFAE6BCC3D15B609AFC91" box="[163,1422,805,829]" gridrow="0" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D15B647EFC91" box="[163,362,805,829]" gridcol="0" gridrow="0" pageId="8" pageNumber="244">Characters/species</th>
|
||||
<th id="7E2E5DB7917EFAE6BDE4D15B66CEFC91" box="[388,986,805,829]" gridcol="1" gridrow="0" pageId="8" pageNumber="244">
|
||||
<taxonomicName id="44CF4D0A917E0511BDE4D15B6669FC91" authority="Ahmed & Marimuthu" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[388,893,805,829]" class="Clitellata" family="Acanthodrilidae" genus="Priodoscolex" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="244" phylum="Annelida" rank="species" species="maduraiensis" status="sp. nov.">
|
||||
<emphasis id="B1BBEA9B917E0511BDE4D15B678DFC91" box="[388,665,805,829]" italics="true" pageId="8" pageNumber="244">Priodoscolex maduraiensis</emphasis>
|
||||
Ahmed & Marimuthu
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="AA8857E0917E0511BFE7D15B66C2FC90" box="[903,982,805,828]" pageId="8" pageNumber="244" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</th>
|
||||
<th id="7E2E5DB7917EFAE6BF94D15B609AFC91" box="[1012,1422,805,829]" gridcol="2" gridrow="0" pageId="8" pageNumber="244">
|
||||
<taxonomicName id="44CF4D0A917E0511BF94D15B6076FC91" authority="Gates, 1940 Length" authorityName="Gates" authorityYear="1940" box="[1012,1378,805,829]" class="Clitellata" family="Acanthodrilidae" genus="Priodoscolex" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="244" phylum="Annelida" rank="species" species="montanus">
|
||||
<emphasis id="B1BBEA9B917E0511BF94D15B61F7FC90" box="[1012,1251,805,829]" italics="true" pageId="8" pageNumber="244">Priodoscolex montanus</emphasis>
|
||||
<bibRefCitation id="E75E4B78917E0511B889D15B6076FC91" author="Gates, G. E." box="[1257,1378,805,829]" pageId="8" pageNumber="244" pagination="115 - 143" refId="ref6481" refString="Gates, G. E. (1940) Indian earthworms. VIII - XI. Records of the Indian Museum, 42 (1), 115-143. https://doi.org/10.26515/rzsi/v42/i1/1940/162455" type="journal article" year="1940">Gates, 1940</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</th>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D133609AFCC9" box="[163,1422,845,869]" gridrow="1" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D133647EFCC9" box="[163,362,845,869]" gridcol="0" gridrow="1" pageId="8" pageNumber="244">Length</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D13366CEFCC9" box="[388,986,845,869]" gridcol="1" gridrow="1" pageId="8" pageNumber="244">92–280 mm</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D133609AFCC9" box="[1012,1422,845,869]" gridcol="2" gridrow="1" pageId="8" pageNumber="244">230–313 mm</td>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D10B609AFC21" box="[163,1422,885,909]" gridrow="2" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D10B647EFC21" box="[163,362,885,909]" gridcol="0" gridrow="2" pageId="8" pageNumber="244">Width</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D10B66CEFC21" box="[388,986,885,909]" gridcol="1" gridrow="2" pageId="8" pageNumber="244">3–6 mm</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D10B609AFC21" box="[1012,1422,885,909]" gridcol="2" gridrow="2" pageId="8" pageNumber="244">3–4 mm</td>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D1E3609AFC1A" box="[163,1422,925,950]" gridrow="3" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D1E3647EFC1A" box="[163,362,925,950]" gridcol="0" gridrow="3" pageId="8" pageNumber="244">Segments</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D1E366CEFC1A" box="[388,986,925,950]" gridcol="1" gridrow="3" pageId="8" pageNumber="244">211–389</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D1E3609AFC1A" box="[1012,1422,925,950]" gridcol="2" gridrow="3" pageId="8" pageNumber="244">306–384</td>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D1B8609AFBE6" box="[163,1422,966,1098]" gridrow="4" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D1B8647EFBE6" box="[163,362,966,1098]" gridcol="0" gridrow="4" pageId="8" pageNumber="244">Setae</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D1B866CEFBE6" box="[388,986,966,1098]" gridcol="1" gridrow="4" pageId="8" pageNumber="244">Perichaetine, except segment 18, ventral setae on segments 12–21 sometimes 11 and 22 displaced anteriorly or posteriorly of the setal line</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D1B8609AFBE6" box="[1012,1422,966,1098]" gridcol="2" gridrow="4" pageId="8" pageNumber="244">Perichaetine, ventral-most setae may be displaced anteriorly or posteriorly to produce a zigzagged appearance of the transverse setal line</td>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D624609AFBD8" box="[163,1422,1114,1140]" gridrow="5" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D624647EFBD8" box="[163,362,1114,1140]" gridcol="0" gridrow="5" pageId="8" pageNumber="244">Clitellum</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D62466CEFBD8" box="[388,986,1114,1140]" gridcol="1" gridrow="5" pageId="8" pageNumber="244">Saddle shaped on segments 13–18</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D624609AFBD8" box="[1012,1422,1114,1140]" gridcol="2" gridrow="5" pageId="8" pageNumber="244">Saddle shaped on segments 13, ½13– 19</td>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D6FC609AFB12" box="[163,1422,1154,1214]" gridrow="6" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D6FC647EFB12" box="[163,362,1154,1214]" gridcol="0" gridrow="6" pageId="8" pageNumber="244">Spermathecal pores</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D6FC66CEFB12" box="[388,986,1154,1214]" gridcol="1" gridrow="6" pageId="8" pageNumber="244">Three pairs 7,8 and 9 (just posterior to 6/7,7/8 and 8/9</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D6FC609AFB12" box="[1012,1422,1154,1214]" gridcol="2" gridrow="6" pageId="8" pageNumber="244">Two pairs in 8 and 9 (slightly posterior to7/8 and 8/9)</td>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D6B0609AFAA6" box="[163,1422,1230,1290]" gridrow="7" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D6B0647EFAA6" box="[163,362,1230,1290]" gridcol="0" gridrow="7" pageId="8" pageNumber="244">Male pores</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D6B066CEFAA6" box="[388,986,1230,1290]" gridcol="1" gridrow="7" pageId="8" pageNumber="244">Closely paired in segment 18 in a square-shaped depression</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D6B0609AFAA6" box="[1012,1422,1230,1290]" gridcol="2" gridrow="7" pageId="8" pageNumber="244">Closely paired in segment 18 in an oval, transverse genital shield</td>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D765609AFAFB" box="[163,1422,1307,1367]" gridrow="8" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D765647EFAFB" box="[163,362,1307,1367]" gridcol="0" gridrow="8" pageId="8" pageNumber="244">Typhlosole</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D76566CEFAFB" box="[388,986,1307,1367]" gridcol="1" gridrow="8" pageId="8" pageNumber="244">Simple, lamelliform begin in segments 26–28</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D765609AFAFB" box="[1012,1422,1307,1367]" gridcol="2" gridrow="8" pageId="8" pageNumber="244">Simple, lamelliform begin in segments 27–31</td>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D719609AFAD3" box="[163,1422,1383,1407]" gridrow="9" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D719647EFAD3" box="[163,362,1383,1407]" gridcol="0" gridrow="9" pageId="8" pageNumber="244">Seminal vesicles</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D71966CEFAD3" box="[388,986,1383,1407]" gridcol="1" gridrow="9" pageId="8" pageNumber="244">In segments 11 and 12</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D719609AFAD3" box="[1012,1422,1383,1407]" gridcol="2" gridrow="9" pageId="8" pageNumber="244">In segment 11 and 12</td>
|
||||
</tr>
|
||||
<tr id="3DFF34CB917EFAE6BCC3D7F1609AFA43" box="[163,1422,1423,1519]" gridrow="10" pageId="8" pageNumber="244">
|
||||
<th id="7E2E5DB7917EFAE6BCC3D7F1647EFA43" box="[163,362,1423,1519]" gridcol="0" gridrow="10" pageId="8" pageNumber="244">Spermathecae</th>
|
||||
<td id="7E2E5DB7917EFAE6BDE4D7F166CEFA43" box="[388,986,1423,1519]" gridcol="1" gridrow="10" pageId="8" pageNumber="244">Three pairs in segments 7, 8 and 9 sometimes anterior directed in segments 6, 7 and 8, with digitiform, ectal diverticulum</td>
|
||||
<td id="7E2E5DB7917EFAE6BF94D7F1609AFA43" box="[1012,1422,1423,1519]" gridcol="2" gridrow="10" pageId="8" pageNumber="244">Two pairs in segments 8 and 9, with digitiform, ectal diverticulum</td>
|
||||
</tr>
|
||||
</table>
|
||||
</paragraph>
|
||||
<paragraph id="83703689917E0510BCA7D43D6191FEB1" blockId="8.[151,1437,1602,2025]" lastBlockId="9.[151,1436,150,429]" lastPageId="9" lastPageNumber="245" pageId="8" pageNumber="244">
|
||||
<emphasis id="B1BBEA9B917E0511BCA7D43D643EF9F1" box="[199,298,1603,1629]" italics="true" pageId="8" pageNumber="244">Internal:</emphasis>
|
||||
Septa 6/7–11/12 strongly thickened, septa 5/6 and 12/13 thickened. Gizzard large cylindrical in segment 5. Intestine begin in segment 19. Typhlosole simple, lamelliform, begin in segment 28 ending in segment 204. Calciferous gland extramural, 4 pairs in segments 14–17 (
|
||||
<figureCitation id="1BF42A0C917E0511BF02D4F566CBF909" box="[866,991,1674,1701]" captionStart="FIGURE 5" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1423,489,1876]" captionTargetId="figure-187@7.[164,1423,489,1876]" captionTargetPageId="7" captionText="FIGURE 5. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. internal anatomy (anterior), B. Spermathecae. C. Calciferous glands and prostates glands. Abbreviations: gz, gizzard; di, diverticulum; am, ampulla; sv–xi, seminal vesicles of segment 11, sv–xii, seminal vesicle of segment 12, ts–x, male funnels of segment 10; ts–xi, male funnels of segment 11; cg, calciferous glands; pd, prostatic duct; pg, prostate glands." figureDoi="http://doi.org/10.5281/zenodo.15215032" httpUri="https://zenodo.org/record/15215032/files/figure.png" pageId="8" pageNumber="244">Fig. 5A, C</figureCitation>
|
||||
). The calciferous glands of segment 14 are small in seize each with a single lobe, whereas, the glands of segments 15,16,17 are bilobed and well developed communicating dorsally, open into the intestine laterally. Nephridia meroic. Large cluster of enteronephric paired tufts in segments 4 and 5; exonephric paired tufts in segments 3 and 6–13 located at the anterior side of septa 3/4 and 6/7–13/14, open ventrally through long duct; several exonephric nephridia on the body wall of segment 14 to the last segment. Megameronephridia absent. Last pair of hearts in segment 12. Dorsal blood vessel complete, extend to the segment 3. Subneural trunk present, turning lateral in segment 15, join the latero-parietal vessel in segment 13. At segment 14 the subneural vessel branched, leave two branches on parities of segments 15 and 16 and one branch diminish posterior in segment 17. Ovaries and funnels in segment 13, ovaries grapes like, attached with posterior face of septa 12/13. Holandric, testis (testes free) and funnels in segments 10 and 11 (
|
||||
<figureCitation id="1BF42A0C917E0511B836D5B161A4F845" box="[1110,1200,1998,2025]" captionStart="FIGURE 5" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1423,489,1876]" captionTargetId="figure-187@7.[164,1423,489,1876]" captionTargetPageId="7" captionText="FIGURE 5. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. internal anatomy (anterior), B. Spermathecae. C. Calciferous glands and prostates glands. Abbreviations: gz, gizzard; di, diverticulum; am, ampulla; sv–xi, seminal vesicles of segment 11, sv–xii, seminal vesicle of segment 12, ts–x, male funnels of segment 10; ts–xi, male funnels of segment 11; cg, calciferous glands; pd, prostatic duct; pg, prostate glands." figureDoi="http://doi.org/10.5281/zenodo.15215032" httpUri="https://zenodo.org/record/15215032/files/figure.png" pageId="8" pageNumber="244">Fig. 5A</figureCitation>
|
||||
). Seminal vesicles in segments 11 and 12 (5A, C). Prostates tubulo-racemose, flattened, extend in segments 18–20; duct
|
||||
<emphasis id="B1BBEA9B917F0510B8BCD2E961FCFF1C" box="[1244,1256,151,176]" italics="true" pageId="9" pageNumber="245">c</emphasis>
|
||||
shaped, curved, about
|
||||
<quantity id="44379B6C917F0510BCBBD2C5640EFF79" box="[219,282,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="245" unit="mm" value="3.0">3 mm</quantity>
|
||||
long; vas deferens join the prostatic duct ectally in segment 18 (
|
||||
<figureCitation id="1BF42A0C917F0510BFBAD2C56125FF79" box="[986,1073,186,213]" captionStart="FIGURE 5" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1423,489,1876]" captionTargetId="figure-187@7.[164,1423,489,1876]" captionTargetPageId="7" captionText="FIGURE 5. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. internal anatomy (anterior), B. Spermathecae. C. Calciferous glands and prostates glands. Abbreviations: gz, gizzard; di, diverticulum; am, ampulla; sv–xi, seminal vesicles of segment 11, sv–xii, seminal vesicle of segment 12, ts–x, male funnels of segment 10; ts–xi, male funnels of segment 11; cg, calciferous glands; pd, prostatic duct; pg, prostate glands." figureDoi="http://doi.org/10.5281/zenodo.15215032" httpUri="https://zenodo.org/record/15215032/files/figure.png" pageId="9" pageNumber="245">Fig. 5C</figureCitation>
|
||||
). Spermathecae unidiverticulate, anterior directed in segments 6,7 and 8; spermathecal duct short, ampulla sacular, nearly club-shaped; digitiform, ectal diverticulum, about 2/3 length of duct and ampulla (
|
||||
<figureCitation id="1BF42A0C917F0510BF7CD37D6681FEB1" box="[796,917,258,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1899,1923]" captionTargetBox="[164,1423,489,1876]" captionTargetId="figure-187@7.[164,1423,489,1876]" captionTargetPageId="7" captionText="FIGURE 5. Priodoscolex maduraiensis Ahmed & Marimuthu, sp. nov. A. internal anatomy (anterior), B. Spermathecae. C. Calciferous glands and prostates glands. Abbreviations: gz, gizzard; di, diverticulum; am, ampulla; sv–xi, seminal vesicles of segment 11, sv–xii, seminal vesicle of segment 12, ts–x, male funnels of segment 10; ts–xi, male funnels of segment 11; cg, calciferous glands; pd, prostatic duct; pg, prostate glands." figureDoi="http://doi.org/10.5281/zenodo.15215032" httpUri="https://zenodo.org/record/15215032/files/figure.png" pageId="9" pageNumber="245">Fig. 5A, B</figureCitation>
|
||||
). Penial setae absent.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD56502917F0510BCA7D35861DCFEED" box="[199,1224,294,321]" pageId="9" pageNumber="245" type="distribution">
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<paragraph id="83703689917F0510BCA7D35861DCFEED" blockId="9.[151,1436,150,429]" box="[199,1224,294,321]" pageId="9" pageNumber="245">
|
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<emphasis id="B1BBEA9B917F0510BCA7D358644DFEEC" bold="true" box="[199,345,294,320]" pageId="9" pageNumber="245">Distribution</emphasis>
|
||||
.
|
||||
<collectingCountry id="FBD87619917F0510BD07D35964B1FEED" box="[359,421,295,321]" name="India" pageId="9" pageNumber="245">India</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="410BF86B917F0510BDCFD3596763FEED" box="[431,631,295,321]" country="India" name="Tamil Nadu" pageId="9" pageNumber="245">Tamil Nadu State</collectingRegion>
|
||||
: Madurai District: Madurai; Salam District: Nagalur.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD56502917F0510BCA7D335619EFE01" pageId="9" pageNumber="245" type="discussion">
|
||||
<paragraph id="83703689917F0510BCA7D335619EFE01" blockId="9.[151,1436,150,429]" pageId="9" pageNumber="245">
|
||||
<emphasis id="B1BBEA9B917F0510BCA7D335642FFEC9" bold="true" box="[199,315,331,357]" pageId="9" pageNumber="245">Remarks.</emphasis>
|
||||
The new species is differentiated (
|
||||
<tableCitation id="CE4D0332917F0510BEA7D335660FFEC9" box="[711,795,331,357]" captionStart="TABLE 1" captionStartId="8.[152,244,764,788]" captionTargetPageId="8" captionText="TABLE 1. Comparative analysis between the new species and described species" pageId="9" pageNumber="245">Table 1</tableCitation>
|
||||
) from the known species
|
||||
<taxonomicName id="44CF4D0A917F0510B825D33565DBFE25" authority="Gates, 1940" authorityName="Gates" authorityYear="1940" class="Clitellata" family="Acanthodrilidae" genus="Priodoscolex" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="245" phylum="Annelida" rank="species" species="montanus">
|
||||
<emphasis id="B1BBEA9B917F0510B825D3356059FEC8" box="[1093,1357,331,357]" italics="true" pageId="9" pageNumber="245">Priodoscolex montanus</emphasis>
|
||||
<bibRefCitation id="E75E4B78917F0510B935D33565DBFE25" author="Gates, G. E." pageId="9" pageNumber="245" pagination="115 - 143" refId="ref6481" refString="Gates, G. E. (1940) Indian earthworms. VIII - XI. Records of the Indian Museum, 42 (1), 115-143. https://doi.org/10.26515/rzsi/v42/i1/1940/162455" type="journal article" year="1940">Gates, 1940</bibRefCitation>
|
||||
</taxonomicName>
|
||||
by the number of spermathecal pores (3 pairs
|
||||
<emphasis id="B1BBEA9B917F0510BE80D31167E3FE24" box="[736,759,367,392]" italics="true" pageId="9" pageNumber="245">vs</emphasis>
|
||||
2 pairs), origin of typhlosole 26–28
|
||||
<emphasis id="B1BBEA9B917F0510B8F9D31161A4FE24" box="[1177,1200,367,392]" italics="true" pageId="9" pageNumber="245">vs</emphasis>
|
||||
27–31. Extension of clitellum 13–18
|
||||
<emphasis id="B1BBEA9B917F0510BD2FD3ED6472FE00" box="[335,358,403,428]" italics="true" pageId="9" pageNumber="245">vs</emphasis>
|
||||
13–19. Posterior dorsal break
|
||||
<emphasis id="B1BBEA9B917F0510BEDED3ED67C1FE00" box="[702,725,403,428]" italics="true" pageId="9" pageNumber="245">vs</emphasis>
|
||||
anterior dorsal and ventral setal break.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
90
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90
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.
|
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species.
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@ -78,11 +79,19 @@ species.
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Beddard, 1883
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=
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(Beddard, 1883)
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.
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<collectingCountry id="FBD876199174051BBD07D3ED64F9FE01" box="[359,493,403,429]" name="Bangladesh" pageId="2" pageNumber="238">Bangladesh</collectingCountry>
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371
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||||
<mods:detail id="5752036185208AE2A7FE6878EEC07479" type="issue">
|
||||
<mods:number id="8636C3B76863FB73A5FFDB0D115880DB">2</mods:number>
|
||||
</mods:detail>
|
||||
<mods:extent id="7990EFC66810A1CADEBC6A4108C80834" unit="page">
|
||||
<mods:start id="7CD548E405B2015AAE9010325E643199">236</mods:start>
|
||||
<mods:end id="6AFF9C8CED240CEE81DD89464D193396">248</mods:end>
|
||||
</mods:extent>
|
||||
</mods:part>
|
||||
</mods:relatedItem>
|
||||
<mods:location id="6AD7AD56876914E672F47F43766CCE18">
|
||||
<mods:url id="E7B09937B047C962FA2766FBCCE939F9">https://doi.org/10.11646/zootaxa.5609.2.5</mods:url>
|
||||
</mods:location>
|
||||
<mods:classification id="07299A8E11E3E5442016F51AFFD04F25">journal article</mods:classification>
|
||||
<mods:identifier id="95D661E824A3112FAFD03335A310009E" type="DOI">10.11646/zootaxa.5609.2.5</mods:identifier>
|
||||
<mods:identifier id="A400BB949D3B4F5BA34ECB37E90667AC" type="ISSN">1175-5326</mods:identifier>
|
||||
<mods:identifier id="9870D1E74FB00D5D94A5BDDAAF2E9338" type="Zenodo-Dep">15215020</mods:identifier>
|
||||
<mods:identifier id="A29ECAFBF301564822C2FBC12C9C27D6" type="ZooBank">5F98331A-A9B1-4825-A477-5628EA14B6CA</mods:identifier>
|
||||
</mods:mods>
|
||||
<treatment id="0B66879F9174051CBCF7D38166CBFEC9" LSID="urn:lsid:plazi:treatment:0B66879F9174051CBCF7D38166CBFEC9" httpUri="http://treatment.plazi.org/id/0B66879F9174051CBCF7D38166CBFEC9" lastPageId="5" lastPageNumber="240" pageId="2" pageNumber="238">
|
||||
<subSubSection id="CBD565029174051BBCF7D3816638FDB5" box="[151,812,511,538]" pageId="2" pageNumber="238" type="nomenclature">
|
||||
<paragraph id="837036899174051BBCF7D3816638FDB5" blockId="2.[151,1182,511,609]" box="[151,812,511,538]" pageId="2" pageNumber="238">
|
||||
<heading id="D83881E59174051BBCF7D3816638FDB5" bold="true" box="[151,812,511,538]" fontSize="11" level="1" pageId="2" pageNumber="238" reason="1">
|
||||
<emphasis id="B1BBEA9B9174051BBCF7D3816638FDB5" bold="true" box="[151,812,511,538]" pageId="2" pageNumber="238">
|
||||
<taxonomicName id="44CF4D0A9174051BBCF7D38167D5FDB5" authority="Ahmed & Marimuthu" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[151,705,511,538]" class="Clitellata" family="Acanthodrilidae" genus="Eutyphoeus" kingdom="Animalia" order="Crassiclitellata" pageId="2" pageNumber="238" phylum="Annelida" rank="species" species="unithecus" status="sp. nov.">
|
||||
<emphasis id="B1BBEA9B9174051BBCF7D38164B1FDB6" bold="true" box="[151,421,511,538]" italics="true" pageId="2" pageNumber="238">Eutyphoeus unithecus</emphasis>
|
||||
Ahmed & Marimuthu
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="AA8857E09174051BBEAED3816638FDB5" box="[718,812,511,537]" pageId="2" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD565029174051BBCF7D05D641FFDCD" pageId="2" pageNumber="238" type="description">
|
||||
<paragraph id="837036899174051BBCF7D05D618AFD92" blockId="2.[151,1182,511,609]" box="[151,1182,547,574]" pageId="2" pageNumber="238">
|
||||
http://zoobank.org/
|
||||
<uri id="F75E3A8B9174051BBD13D05D618AFD92" box="[371,1182,547,574]" pageId="2" pageNumber="238">
|
||||
urn:lsid:zoobank.org:act:
|
||||
<uuid id="F7690C5C9174051BBEE4D05D618AFD92" box="[644,1182,547,574]" pageId="2" pageNumber="238">825A4379-2F56-435E-AEA3-2C6BF59D2EC1</uuid>
|
||||
</uri>
|
||||
</paragraph>
|
||||
<paragraph id="837036899174051BBCF7D039641FFDCD" blockId="2.[151,1182,511,609]" box="[151,267,583,609]" pageId="2" pageNumber="238">
|
||||
(
|
||||
<figureCitation id="1BF42A0C9174051BBCFFD0396416FDCD" box="[159,258,583,609]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1857,1881]" captionStartId-1="3.[151,250,1841,1865]" captionTargetBox-0="[151,1436,883,1831]" captionTargetBox-1="[151,1436,181,1817]" captionTargetId-0="figure-145@2.[151,1436,883,1831]" captionTargetId-1="figure-21@3.[151,1436,181,1817]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view: male pore, genital markings and spermathecal pore, C. prostomium, D. showing spermathecal pore. Abbreviations: pr, prostomium; gm, genital markings; cl, clitellum; spp, spermathecal pore; mp, male pores; ps, penial setae." captionText-1="FIGURE 2. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. internal anatomy, B. spermathecae (dorsal side), C. penial setae 10x, D. penial setae 100x, E. Calciferous glands, F. Spermathecae (ventral side). Abbreviations: di, diverticulum; am, ampulla; gz, gizzard; ts, testes; sv, seminal vesicles; in, intestine; vd, vas deferens; pd, prostatic duct; pg, prostate glands; vdo, vas deferens opening; dv, dorsal blood vessel; vn, ventral nerve cord; cg, calciferous glands; he, hearts." figureDoi-0="http://doi.org/10.5281/zenodo.15215024" figureDoi-1="http://doi.org/10.5281/zenodo.15215026" httpUri-0="https://zenodo.org/record/15215024/files/figure.png" httpUri-1="https://zenodo.org/record/15215026/files/figure.png" pageId="2" pageNumber="238">Figs 1–2</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD565029174051BBCF7D0F16026FCB9" pageId="2" pageNumber="238" type="materials_examined">
|
||||
<paragraph id="837036899174051BBCF7D0F16026FCB9" blockId="2.[151,1437,655,862]" pageId="2" pageNumber="238">
|
||||
<emphasis id="B1BBEA9B9174051BBCF7D0F1642FFD05" bold="true" box="[151,315,655,681]" pageId="2" pageNumber="238">Type material</emphasis>
|
||||
.
|
||||
<materialsCitation id="33A73CD49174051BBD27D0F16401FCB9" collectingDate="1999-10-26" collectionCode="ZSI -GNC" collectorName="P. S. Chaudhuri." country="India" county="Khowai" elevation="52" latitude="24.063906" location="Khowai" longLatPrecision="1" longitude="91.6007" pageId="2" pageNumber="238" specimenCode="An8034/2" specimenCount="1" stateProvince="Tripura" typeStatus="holotype">
|
||||
<emphasis id="B1BBEA9B9174051BBD27D0F164BAFD05" box="[327,430,655,681]" italics="true" pageId="2" pageNumber="238">
|
||||
<typeStatus id="5C74882B9174051BBD27D0F164BAFD05" box="[327,430,655,681]" pageId="2" pageNumber="238" type="holotype">Holotype</typeStatus>
|
||||
</emphasis>
|
||||
: Clitellate (
|
||||
<collectionCode id="E5DEAE4C9174051BBE4DD0F167B7FD05" box="[557,675,655,681]" pageId="2" pageNumber="238">ZSI–GNC</collectionCode>
|
||||
–
|
||||
<specimenCode id="D3699EF29174051BBED2D0F16630FD05" box="[690,804,655,681]" pageId="2" pageNumber="238">An8034/2</specimenCode>
|
||||
),
|
||||
<location id="861060529174051BBF54D0F16699FD05" LSID="urn:lsid:plazi:treatment:0B66879F9174051CBCF7D38166CBFEC9:861060529174051BBF54D0F16699FD05" box="[820,909,655,681]" country="India" county="Khowai" latitude="24.063906" longLatPrecision="1" longitude="91.6007" name="Khowai" pageId="2" pageNumber="238" stateProvince="Tripura">Khowai</location>
|
||||
(
|
||||
<geoCoordinate id="E6FB504E9174051BBFF9D0F1612DFD06" box="[921,1081,655,682]" degrees="24.063905" direction="north" orientation="latitude" pageId="2" pageNumber="238" precision="1" value="24.063906">24.063905° N</geoCoordinate>
|
||||
;
|
||||
<geoCoordinate id="E6FB504E9174051BB822D0F161CEFD05" box="[1090,1242,655,681]" degrees="91.600703" direction="east" orientation="longitude" pageId="2" pageNumber="238" precision="1" value="91.6007">91.600703°E</geoCoordinate>
|
||||
),
|
||||
<collectingCounty id="6A114E059174051BB88AD0F16057FD05" box="[1258,1347,655,681]" pageId="2" pageNumber="238">Khowai</collectingCounty>
|
||||
district,
|
||||
<collectingRegion id="410BF86B9174051BBCF7D0CD65F9FD61" box="[151,237,691,717]" country="India" name="Tripura" pageId="2" pageNumber="238">Tripura</collectingRegion>
|
||||
,
|
||||
<collectingCountry id="FBD876199174051BBC9AD0CD6423FD61" box="[250,311,691,717]" name="India" pageId="2" pageNumber="238">India</collectingCountry>
|
||||
;
|
||||
<quantity id="44379B6C9174051BBD25D0CD6494FD62" box="[325,384,691,718]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="238" unit="m" value="52.0">
|
||||
<elevation id="08E2D1BA9174051BBD25D0CD6494FD62" box="[325,384,691,718]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="238" unit="m" value="52.0">52 m</elevation>
|
||||
</quantity>
|
||||
above mean sea level;
|
||||
<date id="F77110499174051BBEEFD0CD6644FD61" box="[655,848,691,717]" pageId="2" pageNumber="238" value="1999-10-26">
|
||||
<collectingDate id="E735E9A19174051BBEEFD0CD6644FD61" box="[655,848,691,717]" pageId="2" pageNumber="238" value="1999-10-26">26 October 1999</collectingDate>
|
||||
</date>
|
||||
(date here is based on the deposition of specimens in the Zoological Survey of India, Kolkata, whereas specimens were collected during monsoon season); leg.
|
||||
<collectorName id="2E3A535F9174051BB912D0A96401FCB9" pageId="2" pageNumber="238">P.S. Chaudhuri.</collectorName>
|
||||
</materialsCitation>
|
||||
<materialsCitation id="33A73CD49174051BBD7BD082603AFCB9" box="[283,1326,763,790]" collectingDate="1999-10-26" collectionCode="ZSI -GNC" collectorName="P. S. Chaudhuri." country="India" county="Khowai" elevation="52" latitude="24.063906" location="Khowai" longLatPrecision="1" longitude="91.6007" pageId="2" pageNumber="238" specimenCode="An8035/2" specimenCount="1" stateProvince="Tripura" typeStatus="paratype">
|
||||
<emphasis id="B1BBEA9B9174051BBD7BD0826499FCB9" box="[283,397,764,789]" italics="true" pageId="2" pageNumber="238">
|
||||
<typeStatus id="5C74882B9174051BBD7BD0826499FCB9" box="[283,397,764,789]" pageId="2" pageNumber="238" type="paratype">Paratypes</typeStatus>
|
||||
</emphasis>
|
||||
: 5 clitellates (
|
||||
<collectionCode id="E5DEAE4C9174051BBE4BD08567B6FCBA" box="[555,674,763,790]" pageId="2" pageNumber="238">ZSI–GNC</collectionCode>
|
||||
–
|
||||
<specimenCode id="D3699EF29174051BBED1D0856636FCBA" box="[689,802,763,790]" pageId="2" pageNumber="238">An8035/2</specimenCode>
|
||||
), collection details same as for as the holotype
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD565029174051BBCA7D161612BFCF1" pageId="2" pageNumber="238" type="etymology">
|
||||
<paragraph id="837036899174051BBCA7D161612BFCF1" blockId="2.[151,1437,655,862]" pageId="2" pageNumber="238">
|
||||
<emphasis id="B1BBEA9B9174051BBCA7D161645AFC95" bold="true" box="[199,334,799,825]" pageId="2" pageNumber="238">Etymology.</emphasis>
|
||||
The specific name
|
||||
<taxonomicName id="44CF4D0A9174051BBE4BD16167BDFC95" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[555,681,799,825]" class="Clitellata" family="Acanthodrilidae" genus="Eutyphoeus" kingdom="Animalia" order="Crassiclitellata" pageId="2" pageNumber="238" phylum="Annelida" rank="species" species="unithecus">
|
||||
“
|
||||
<emphasis id="B1BBEA9B9174051BBE58D16167BDFC95" box="[568,681,799,825]" italics="true" pageId="2" pageNumber="238">unithecus</emphasis>
|
||||
</taxonomicName>
|
||||
<emphasis id="B1BBEA9B9174051BBEC8D16167A6FC95" box="[680,690,799,825]" italics="true" pageId="2" pageNumber="238">”</emphasis>
|
||||
is derived from the Latin words “uni” meaning one or single, and “thecae” mean spermathecae which appropriately describes the single spermatheca.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<caption id="D7B066019174051BBCF7D53F6648F80C" ID-DOI="http://doi.org/10.5281/zenodo.15215024" ID-Zenodo-Dep="15215024" httpUri="https://zenodo.org/record/15215024/files/figure.png" pageId="2" pageNumber="238" startId="2.[151,250,1857,1881]" targetBox="[151,1436,883,1831]" targetPageId="2" targetType="figure">
|
||||
<paragraph id="837036899174051BBCF7D53F6648F80C" blockId="2.[151,1437,1857,1953]" pageId="2" pageNumber="238">
|
||||
<emphasis id="B1BBEA9B9174051BBCF7D53F6407F8F5" bold="true" box="[151,275,1857,1881]" pageId="2" pageNumber="238">FIGURE 1.</emphasis>
|
||||
<taxonomicName id="44CF4D0A9174051BBD79D53F67CFF8F5" authority="Ahmed & Marimuthu" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[281,731,1857,1881]" class="Clitellata" family="Acanthodrilidae" genus="Eutyphoeus" kingdom="Animalia" order="Crassiclitellata" pageId="2" pageNumber="238" phylum="Annelida" rank="species" species="unithecus" status="sp. nov.">
|
||||
<emphasis id="B1BBEA9B9174051BBD79D53F64E3F8F5" box="[281,503,1857,1881]" italics="true" pageId="2" pageNumber="238">Eutyphoeus unithecus</emphasis>
|
||||
Ahmed & Marimuthu
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="AA8857E09174051BBE85D53F6627F8F4" box="[741,819,1857,1880]" pageId="2" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="B1BBEA9B9174051BBF59D53F6645F8F5" bold="true" box="[825,849,1857,1881]" pageId="2" pageNumber="238">A.</emphasis>
|
||||
lateral view,
|
||||
<emphasis id="B1BBEA9B9174051BBFB9D53F66E4F8F5" bold="true" box="[985,1008,1857,1881]" pageId="2" pageNumber="238">B.</emphasis>
|
||||
ventral view: male pore, genital markings and spermathecal pore,
|
||||
<emphasis id="B1BBEA9B9174051BBDEAD51B64B7F8D1" bold="true" box="[394,419,1893,1917]" pageId="2" pageNumber="238">C.</emphasis>
|
||||
prostomium,
|
||||
<emphasis id="B1BBEA9B9174051BBE52D51B675EF8D1" bold="true" box="[562,586,1893,1917]" pageId="2" pageNumber="238">D.</emphasis>
|
||||
showing spermathecal pore. Abbreviations:
|
||||
<emphasis id="B1BBEA9B9174051BB872D518613DF8D1" box="[1042,1065,1894,1917]" italics="true" pageId="2" pageNumber="238">pr</emphasis>
|
||||
, prostomium;
|
||||
<emphasis id="B1BBEA9B9174051BB8DED51861C9F8D1" box="[1214,1245,1894,1917]" italics="true" pageId="2" pageNumber="238">gm</emphasis>
|
||||
, genital markings;
|
||||
<emphasis id="B1BBEA9B9174051BBCF7D5F765BDF80D" box="[151,169,1929,1953]" italics="true" pageId="2" pageNumber="238">cl</emphasis>
|
||||
, clitellum;
|
||||
<emphasis id="B1BBEA9B9174051BBD7ED5F46455F80D" box="[286,321,1930,1953]" italics="true" pageId="2" pageNumber="238">spp</emphasis>
|
||||
, spermathecal pore;
|
||||
<emphasis id="B1BBEA9B9174051BBE74D5F46727F80D" box="[532,563,1930,1953]" italics="true" pageId="2" pageNumber="238">mp</emphasis>
|
||||
, male pores;
|
||||
<emphasis id="B1BBEA9B9174051BBEDBD5F467C6F80D" box="[699,722,1930,1953]" italics="true" pageId="2" pageNumber="238">ps</emphasis>
|
||||
, penial setae.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<caption id="D7B066019175051ABCF7D54F645BF875" ID-DOI="http://doi.org/10.5281/zenodo.15215026" ID-Zenodo-Dep="15215026" httpUri="https://zenodo.org/record/15215026/files/figure.png" pageId="3" pageNumber="239" startId="3.[151,250,1841,1865]" targetBox="[151,1436,181,1817]" targetPageId="3" targetType="figure">
|
||||
<paragraph id="837036899175051ABCF7D54F6110F8C1" blockId="3.[151,1436,1841,2009]" pageId="3" pageNumber="239">
|
||||
<emphasis id="B1BBEA9B9175051ABCF7D54F6406F8E5" bold="true" box="[151,274,1841,1865]" pageId="3" pageNumber="239">FIGURE 2.</emphasis>
|
||||
<taxonomicName id="44CF4D0A9175051ABD78D54F67CDF8E5" authority="Ahmed & Marimuthu" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[280,729,1841,1865]" class="Clitellata" family="Acanthodrilidae" genus="Eutyphoeus" kingdom="Animalia" order="Crassiclitellata" pageId="3" pageNumber="239" phylum="Annelida" rank="species" species="unithecus" status="sp. nov.">
|
||||
<emphasis id="B1BBEA9B9175051ABD78D54F64E2F8E5" box="[280,502,1841,1865]" italics="true" pageId="3" pageNumber="239">Eutyphoeus unithecus</emphasis>
|
||||
Ahmed & Marimuthu
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="AA8857E09175051ABE82D54C6625F8E5" box="[738,817,1842,1865]" pageId="3" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
<emphasis id="B1BBEA9B9175051ABF56D54F665CF8E5" bold="true" box="[822,840,1841,1865]" pageId="3" pageNumber="239">A</emphasis>
|
||||
. internal anatomy,
|
||||
<emphasis id="B1BBEA9B9175051AB869D54F610EF8E5" bold="true" box="[1033,1050,1841,1865]" pageId="3" pageNumber="239">B</emphasis>
|
||||
. spermathecae (dorsal side),
|
||||
<emphasis id="B1BBEA9B9175051AB920D54F6046F8E5" bold="true" box="[1344,1362,1841,1865]" pageId="3" pageNumber="239">C</emphasis>
|
||||
. penial setae 10x,
|
||||
<emphasis id="B1BBEA9B9175051ABD63D52B6401F8C1" bold="true" box="[259,277,1877,1901]" pageId="3" pageNumber="239">D</emphasis>
|
||||
. penial setae 100x,
|
||||
<emphasis id="B1BBEA9B9175051ABDBED52B64FBF8C1" bold="true" box="[478,495,1877,1901]" pageId="3" pageNumber="239">E</emphasis>
|
||||
. Calciferous glands,
|
||||
<emphasis id="B1BBEA9B9175051ABEA6D52B67C1F8C1" bold="true" box="[710,725,1877,1901]" pageId="3" pageNumber="239">F</emphasis>
|
||||
. Spermathecae (ventral side).
|
||||
</paragraph>
|
||||
<paragraph id="837036899175051ABCF7D507645BF875" blockId="3.[151,1436,1841,2009]" pageId="3" pageNumber="239">
|
||||
Abbreviations:
|
||||
<emphasis id="B1BBEA9B9175051ABD56D507645EF83D" box="[310,330,1913,1937]" italics="true" pageId="3" pageNumber="239">di</emphasis>
|
||||
, diverticulum;
|
||||
<emphasis id="B1BBEA9B9175051ABD8AD504671DF83D" box="[490,521,1914,1937]" italics="true" pageId="3" pageNumber="239">am</emphasis>
|
||||
, ampulla;
|
||||
<emphasis id="B1BBEA9B9175051ABE1AD5046785F83D" box="[634,657,1914,1937]" italics="true" pageId="3" pageNumber="239">gz</emphasis>
|
||||
, gizzard;
|
||||
<emphasis id="B1BBEA9B9175051ABE9AD504661FF83D" box="[762,779,1914,1937]" italics="true" pageId="3" pageNumber="239">ts</emphasis>
|
||||
, testes;
|
||||
<emphasis id="B1BBEA9B9175051ABF02D5046663F83D" box="[866,887,1914,1937]" italics="true" pageId="3" pageNumber="239">sv</emphasis>
|
||||
, seminal vesicles;
|
||||
<emphasis id="B1BBEA9B9175051AB85DD5046145F83D" box="[1085,1105,1914,1937]" italics="true" pageId="3" pageNumber="239">in</emphasis>
|
||||
, intestine;
|
||||
<emphasis id="B1BBEA9B9175051AB8A6D50761CAF83D" box="[1222,1246,1913,1937]" italics="true" pageId="3" pageNumber="239">vd</emphasis>
|
||||
, vas deferens;
|
||||
<emphasis id="B1BBEA9B9175051AB91DD5076082F83D" box="[1405,1430,1913,1937]" italics="true" pageId="3" pageNumber="239">pd</emphasis>
|
||||
, prostatic duct;
|
||||
<emphasis id="B1BBEA9B9175051ABD51D5E0645EF819" box="[305,330,1950,1973]" italics="true" pageId="3" pageNumber="239">pg</emphasis>
|
||||
, prostate glands;
|
||||
<emphasis id="B1BBEA9B9175051ABE62D5E36733F819" box="[514,551,1949,1973]" italics="true" pageId="3" pageNumber="239">vdo</emphasis>
|
||||
, vas deferens opening;
|
||||
<emphasis id="B1BBEA9B9175051ABF7CD5E36620F819" box="[796,820,1949,1973]" italics="true" pageId="3" pageNumber="239">dv</emphasis>
|
||||
, dorsal blood vessel;
|
||||
<emphasis id="B1BBEA9B9175051AB877D5E0613BF819" box="[1047,1071,1950,1973]" italics="true" pageId="3" pageNumber="239">vn</emphasis>
|
||||
, ventral nerve cord;
|
||||
<emphasis id="B1BBEA9B9175051AB967D5E0600BF819" box="[1287,1311,1950,1973]" italics="true" pageId="3" pageNumber="239">cg</emphasis>
|
||||
, calciferous glands;
|
||||
<emphasis id="B1BBEA9B9175051ABC87D5BF65EBF875" box="[231,255,1985,2009]" italics="true" pageId="3" pageNumber="239">he</emphasis>
|
||||
, hearts.
|
||||
</paragraph>
|
||||
</caption>
|
||||
<subSubSection id="CBD565029172051DBCA7D2E8644CFDCD" pageId="4" pageNumber="240" type="diagnosis">
|
||||
<paragraph id="837036899172051DBCA7D2E8644CFDCD" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="240">
|
||||
<emphasis id="B1BBEA9B9172051DBCA7D2E8642EFF1C" bold="true" box="[199,314,150,176]" pageId="4" pageNumber="240">Diagnosis</emphasis>
|
||||
. Length
|
||||
<quantity id="44379B6C9172051DBDC3D2E9673BFF1D" box="[419,559,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" metricValueMax="1.3199999999999998" metricValueMin="0.86" pageId="4" pageNumber="240" unit="mm" value="109.0" valueMax="132.0" valueMin="86.0">86–132 mm</quantity>
|
||||
, width
|
||||
<quantity id="44379B6C9172051DBEE5D2E967CAFF1D" box="[645,734,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="240" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4mm</quantity>
|
||||
, segments 184–243. Colour brownish (preserved specimens). Segment 5 bi-annulate, further 6–12 tri-annulate. Prostomium tanylobous. Setae lumbricine. First dorsal pore in intersegmental furrow 11/12. Male pores open into a small, oval porophore, median to
|
||||
<emphasis id="B1BBEA9B9172051DB819D2A16193FF54" box="[1145,1159,223,248]" italics="true" pageId="4" pageNumber="240">a</emphasis>
|
||||
setal line, penes absent. Spermathecal pore single, midventral in intersegmental furrow 7/8. Female pores paired, presetal at or slight median to
|
||||
<emphasis id="B1BBEA9B9172051DBCD5D35965D7FEEC" box="[181,195,295,320]" italics="true" pageId="4" pageNumber="240">a</emphasis>
|
||||
setal line. Genital markings, unpaired, disc shaped, post-setal, mid ventral on segments 12 (lateral to
|
||||
<emphasis id="B1BBEA9B9172051DB92BD3596073FEED" box="[1355,1383,295,321]" italics="true" pageId="4" pageNumber="240">bb</emphasis>
|
||||
), 13 (
|
||||
<emphasis id="B1BBEA9B9172051DBCC1D33565A9FEC9" box="[161,189,331,357]" italics="true" pageId="4" pageNumber="240">bb</emphasis>
|
||||
) and 16 (with in
|
||||
<emphasis id="B1BBEA9B9172051DBDEDD33564BDFEC8" box="[397,425,331,356]" italics="true" pageId="4" pageNumber="240">aa</emphasis>
|
||||
), sometime post setal on segments 14 and 15. Typhlosole simple, lamelliform begin in segment 26 ending with the beginning of supra-intestinal glands. Lateral typhlosole, low ridges in segments 27 and 28. Spermatheca with bilobed irregular or dumbbell shaped ampulla, with 2–3 digitiform diverticulum (two lateral and sometime one median). Supra-intestinal glands 4–5 pairs in segments 76, 84–80, 87. Ventral and median intestinal ceaca
|
||||
<quantity id="44379B6C9172051DBD32D3A56494FE59" box="[338,384,475,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="240" unit="in" value="4.0">4 in</quantity>
|
||||
segments 31–37. Seminal vesicles in segment 12 extending to segments 13, 14. Prostates tubular, extending from segment 17 to 18 with long wavy duct. Penial setae
|
||||
<quantity id="44379B6C9172051DBF98D381612BFDB5" box="[1016,1087,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="4" pageNumber="240" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4–6 in</quantity>
|
||||
each battery, length
|
||||
<quantity id="44379B6C9172051DB94BD380608CFDB5" box="[1323,1432,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="240" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2mm</quantity>
|
||||
, tip slightly curved and pointed, ornamented with triangular teeth. Genital markings glands sessile in segments 12, 13, sometime 14.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD565029172051DBCA7D0146695F935" pageId="4" pageNumber="240" type="description">
|
||||
<paragraph id="837036899172051DBCA7D01461F9FC65" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="240">
|
||||
<emphasis id="B1BBEA9B9172051DBCA7D014644DFD28" bold="true" box="[199,345,618,644]" pageId="4" pageNumber="240">Description.</emphasis>
|
||||
<emphasis id="B1BBEA9B9172051DBD02D01564D6FD29" box="[354,450,619,645]" italics="true" pageId="4" pageNumber="240">External</emphasis>
|
||||
: Medium size worm, colour brownish (preserved specimens). Dimensions:
|
||||
<typeStatus id="5C74882B9172051DB94CD015608DFD29" box="[1324,1433,619,645]" pageId="4" pageNumber="240" type="holotype">Holotype</typeStatus>
|
||||
: length
|
||||
<quantity id="44379B6C9172051DBC83D0F1642FFD05" box="[227,315,655,681]" metricMagnitude="-1" metricUnit="m" metricValue="1.22" pageId="4" pageNumber="240" unit="mm" value="122.0">122mm</quantity>
|
||||
, width
|
||||
<quantity id="44379B6C9172051DBDEAD0F164D0FD05" box="[394,452,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="240" unit="mm" value="4.0">4mm</quantity>
|
||||
at clitellum, segments 243;
|
||||
<typeStatus id="5C74882B9172051DBE99D0F1667DFD05" box="[761,873,655,681]" pageId="4" pageNumber="240" type="paratype">paratypes</typeStatus>
|
||||
: length
|
||||
<quantity id="44379B6C9172051DBFDDD0F16151FD05" box="[957,1093,655,681]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" metricValueMax="1.3199999999999998" metricValueMin="0.86" pageId="4" pageNumber="240" unit="mm" value="109.0" valueMax="132.0" valueMin="86.0">86–132 mm</quantity>
|
||||
, width
|
||||
<quantity id="44379B6C9172051DB8F4D0F161F8FD05" box="[1172,1260,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="240" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4mm</quantity>
|
||||
, segments 184– 232. Body cylindrical, segment 5 bi-annulate, 6–12 tri-annulate. Prostomium tanylobous (
|
||||
<figureCitation id="1BF42A0C9172051DB8ECD0CD601CFD61" box="[1164,1288,691,717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1857,1881]" captionTargetBox="[151,1436,883,1831]" captionTargetId="figure-145@2.[151,1436,883,1831]" captionTargetPageId="2" captionText="FIGURE 1. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view: male pore, genital markings and spermathecal pore, C. prostomium, D. showing spermathecal pore. Abbreviations: pr, prostomium; gm, genital markings; cl, clitellum; spp, spermathecal pore; mp, male pores; ps, penial setae." figureDoi="http://doi.org/10.5281/zenodo.15215024" httpUri="https://zenodo.org/record/15215024/files/figure.png" pageId="4" pageNumber="240">Fig. 1A, C</figureCitation>
|
||||
). First dorsal pore in intersegmental furrow 11/12. Setae lumbricine, present from segment 2,
|
||||
<emphasis id="B1BBEA9B9172051DB87ED0A961D9FD5D" box="[1054,1229,727,753]" italics="true" pageId="4" pageNumber="240">aa:ab:bc:cd:dd</emphasis>
|
||||
=1.9:1.2:2:1.3:7 at 5;
|
||||
<emphasis id="B1BBEA9B9172051DBCD3D0856475FCB9" box="[179,353,763,789]" italics="true" pageId="4" pageNumber="240">aa:ab:bc:cd:dd</emphasis>
|
||||
=1.6:1.2:3:1.6:8 at 8;
|
||||
<emphasis id="B1BBEA9B9172051DBE32D0856614FCB9" box="[594,768,763,789]" italics="true" pageId="4" pageNumber="240">aa:ab:bc:cd:dd</emphasis>
|
||||
= 2:1:2:1.5:8 at 20. Clitellum annular, on segments ½13–17, setae present and dorsal pores (visible in 13/14). Spermathecal pore single, midventral in intersegmental furrow 7/8 between to
|
||||
<emphasis id="B1BBEA9B9172051DBD78D13D6420FCF0" box="[280,308,835,860]" italics="true" pageId="4" pageNumber="240">aa</emphasis>
|
||||
setal line (
|
||||
<figureCitation id="1BF42A0C9172051DBDCDD13D673CFCF1" box="[429,552,835,861]" captionStart="FIGURE 1" captionStartId="2.[151,250,1857,1881]" captionTargetBox="[151,1436,883,1831]" captionTargetId="figure-145@2.[151,1436,883,1831]" captionTargetPageId="2" captionText="FIGURE 1. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view: male pore, genital markings and spermathecal pore, C. prostomium, D. showing spermathecal pore. Abbreviations: pr, prostomium; gm, genital markings; cl, clitellum; spp, spermathecal pore; mp, male pores; ps, penial setae." figureDoi="http://doi.org/10.5281/zenodo.15215024" httpUri="https://zenodo.org/record/15215024/files/figure.png" pageId="4" pageNumber="240">Fig. 1B, D</figureCitation>
|
||||
). Avestibulate, male pores median to
|
||||
<emphasis id="B1BBEA9B9172051DBFA9D13D66C3FCF0" box="[969,983,835,860]" italics="true" pageId="4" pageNumber="240">a</emphasis>
|
||||
open in to a small rounded porophore in segment 17, penes absent (
|
||||
<figureCitation id="1BF42A0C9172051DBDA3D119670DFC2D" box="[451,537,871,897]" captionStart="FIGURE 1" captionStartId="2.[151,250,1857,1881]" captionTargetBox="[151,1436,883,1831]" captionTargetId="figure-145@2.[151,1436,883,1831]" captionTargetPageId="2" captionText="FIGURE 1. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view: male pore, genital markings and spermathecal pore, C. prostomium, D. showing spermathecal pore. Abbreviations: pr, prostomium; gm, genital markings; cl, clitellum; spp, spermathecal pore; mp, male pores; ps, penial setae." figureDoi="http://doi.org/10.5281/zenodo.15215024" httpUri="https://zenodo.org/record/15215024/files/figure.png" pageId="4" pageNumber="240">Fig. 1B</figureCitation>
|
||||
). Female pores paired, pre-setal, slight median to
|
||||
<emphasis id="B1BBEA9B9172051DB829D1196143FC2C" box="[1097,1111,871,896]" italics="true" pageId="4" pageNumber="240">a</emphasis>
|
||||
setal lines. Genital markings unpaired, disc shaped, median, post-setal on segments 12 (lateral to
|
||||
<emphasis id="B1BBEA9B9172051DBFE9D1F566B1FC09" box="[905,933,907,933]" italics="true" pageId="4" pageNumber="240">bb</emphasis>
|
||||
), 13 (
|
||||
<emphasis id="B1BBEA9B9172051DBF87D1F56117FC09" box="[999,1027,907,933]" italics="true" pageId="4" pageNumber="240">bb</emphasis>
|
||||
), 16 (
|
||||
<emphasis id="B1BBEA9B9172051DB824D1F56174FC08" box="[1092,1120,907,932]" italics="true" pageId="4" pageNumber="240">aa</emphasis>
|
||||
), slight developed post setal on segment 14 and 15 median to
|
||||
<emphasis id="B1BBEA9B9172051DBE6AD1D16732FC64" box="[522,550,943,968]" italics="true" pageId="4" pageNumber="240">aa</emphasis>
|
||||
(
|
||||
<figureCitation id="1BF42A0C9172051DBE55D1D16798FC65" box="[565,652,943,969]" captionStart="FIGURE 1" captionStartId="2.[151,250,1857,1881]" captionTargetBox="[151,1436,883,1831]" captionTargetId="figure-145@2.[151,1436,883,1831]" captionTargetPageId="2" captionText="FIGURE 1. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. lateral view, B. ventral view: male pore, genital markings and spermathecal pore, C. prostomium, D. showing spermathecal pore. Abbreviations: pr, prostomium; gm, genital markings; cl, clitellum; spp, spermathecal pore; mp, male pores; ps, penial setae." figureDoi="http://doi.org/10.5281/zenodo.15215024" httpUri="https://zenodo.org/record/15215024/files/figure.png" pageId="4" pageNumber="240">Fig. 1B</figureCitation>
|
||||
). Penial setae projecting from the male genital region.
|
||||
</paragraph>
|
||||
<paragraph id="837036899172051DBCA7D1AD61BDF9A5" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="240">
|
||||
<emphasis id="B1BBEA9B9172051DBCA7D1AD6435FC41" box="[199,289,979,1005]" italics="true" pageId="4" pageNumber="240">Internal</emphasis>
|
||||
: Septa 4/5–5/6 and 9/10–11/12 strongly thickened, 8/9 thickened, 6/7 and 7/8 absent. Gizzard single, large, in space between septa 5/6 and 8/9; intestine begin in segment 15 (
|
||||
<figureCitation id="1BF42A0C9172051DBFDBD1896125FBBD" box="[955,1073,1015,1041]" captionStart="FIGURE 2" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-21@3.[151,1436,181,1817]" captionTargetPageId="3" captionText="FIGURE 2. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. internal anatomy, B. spermathecae (dorsal side), C. penial setae 10x, D. penial setae 100x, E. Calciferous glands, F. Spermathecae (ventral side). Abbreviations: di, diverticulum; am, ampulla; gz, gizzard; ts, testes; sv, seminal vesicles; in, intestine; vd, vas deferens; pd, prostatic duct; pg, prostate glands; vdo, vas deferens opening; dv, dorsal blood vessel; vn, ventral nerve cord; cg, calciferous glands; he, hearts." figureDoi="http://doi.org/10.5281/zenodo.15215026" httpUri="https://zenodo.org/record/15215026/files/figure.png" pageId="4" pageNumber="240">Fig. 2A, E</figureCitation>
|
||||
); typhlosole simple, lamelliform begin in segment 26 ending posteriorly in segment 84 with the beginning of supra-intestinal glands; lateral typhlosole, low ridges in segments 27 and 28. Calciferous glands, paired, intramural in segment 12 (
|
||||
<figureCitation id="1BF42A0C9172051DB815D64161D8FBF5" box="[1141,1228,1087,1113]" captionStart="FIGURE 2" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-21@3.[151,1436,181,1817]" captionTargetPageId="3" captionText="FIGURE 2. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. internal anatomy, B. spermathecae (dorsal side), C. penial setae 10x, D. penial setae 100x, E. Calciferous glands, F. Spermathecae (ventral side). Abbreviations: di, diverticulum; am, ampulla; gz, gizzard; ts, testes; sv, seminal vesicles; in, intestine; vd, vas deferens; pd, prostatic duct; pg, prostate glands; vdo, vas deferens opening; dv, dorsal blood vessel; vn, ventral nerve cord; cg, calciferous glands; he, hearts." figureDoi="http://doi.org/10.5281/zenodo.15215026" httpUri="https://zenodo.org/record/15215026/files/figure.png" pageId="4" pageNumber="240">Fig. 2E</figureCitation>
|
||||
). Lateral intestinal caeca absent, median ventral intestinal caeca
|
||||
<quantity id="44379B6C9172051DBEC8D61D67C2FBD1" box="[680,726,1123,1149]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="240" unit="in" value="4.0">4 in</quantity>
|
||||
segments 34–37. Supra-intestinal glands 4 pairs in segments 84–87. Nephridia, meroic; enteronephric tuft nephridia in segment 3, numerous exonephric micro-nephridia on parities in segment 5 and posterior segments arranged in transverse rows from segment 18; paired exonephric megameronephridia in each segment posterior to the supra-intestinal glands. Dorsal vessel complete extending to segment 3, with heart in segment 5 and 6; last pairs of heart in segment 13. Metandric, testes and funnels in segment 11, funnels absent in segment 10. Seminal vesicles in segment 12 extending to segment 13 (
|
||||
<figureCitation id="1BF42A0C9172051DB952D7696084FA9D" box="[1330,1424,1303,1329]" captionStart="FIGURE 2" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-21@3.[151,1436,181,1817]" captionTargetPageId="3" captionText="FIGURE 2. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. internal anatomy, B. spermathecae (dorsal side), C. penial setae 10x, D. penial setae 100x, E. Calciferous glands, F. Spermathecae (ventral side). Abbreviations: di, diverticulum; am, ampulla; gz, gizzard; ts, testes; sv, seminal vesicles; in, intestine; vd, vas deferens; pd, prostatic duct; pg, prostate glands; vdo, vas deferens opening; dv, dorsal blood vessel; vn, ventral nerve cord; cg, calciferous glands; he, hearts." figureDoi="http://doi.org/10.5281/zenodo.15215026" httpUri="https://zenodo.org/record/15215026/files/figure.png" pageId="4" pageNumber="240">Fig. 2A</figureCitation>
|
||||
). Ovaries and funnels in segment 13, ovaries fan shaped attached on the posterior face septa 12/13. The Vas deferens enlarged before passing into the parietes just behind the prostatic duct (
|
||||
<figureCitation id="1BF42A0C9172051DBFD8D7216107FAD5" box="[952,1043,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-21@3.[151,1436,181,1817]" captionTargetPageId="3" captionText="FIGURE 2. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. internal anatomy, B. spermathecae (dorsal side), C. penial setae 10x, D. penial setae 100x, E. Calciferous glands, F. Spermathecae (ventral side). Abbreviations: di, diverticulum; am, ampulla; gz, gizzard; ts, testes; sv, seminal vesicles; in, intestine; vd, vas deferens; pd, prostatic duct; pg, prostate glands; vdo, vas deferens opening; dv, dorsal blood vessel; vn, ventral nerve cord; cg, calciferous glands; he, hearts." figureDoi="http://doi.org/10.5281/zenodo.15215026" httpUri="https://zenodo.org/record/15215026/files/figure.png" pageId="4" pageNumber="240">Fig. 2A</figureCitation>
|
||||
). Prostates tubular, extending from segment 17 to 18, with long wavy duct. Spermatheca single in segment 8, ampulla large, bilobed, with short duct. Bi-diverticulate, digitiform, ental diverticulum, on lateral sides of the duct (
|
||||
<figureCitation id="1BF42A0C9172051DBFBAD7D9617BFA6D" box="[986,1135,1447,1473]" captionStart="FIGURE 2" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-21@3.[151,1436,181,1817]" captionTargetPageId="3" captionText="FIGURE 2. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. internal anatomy, B. spermathecae (dorsal side), C. penial setae 10x, D. penial setae 100x, E. Calciferous glands, F. Spermathecae (ventral side). Abbreviations: di, diverticulum; am, ampulla; gz, gizzard; ts, testes; sv, seminal vesicles; in, intestine; vd, vas deferens; pd, prostatic duct; pg, prostate glands; vdo, vas deferens opening; dv, dorsal blood vessel; vn, ventral nerve cord; cg, calciferous glands; he, hearts." figureDoi="http://doi.org/10.5281/zenodo.15215026" httpUri="https://zenodo.org/record/15215026/files/figure.png" pageId="4" pageNumber="240">Fig. 2A, B, F</figureCitation>
|
||||
). Penial setae 6 per battery, length
|
||||
<quantity id="44379B6C9172051DBC87D7B46440FA49" box="[231,340,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="4" pageNumber="240" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2mm</quantity>
|
||||
, 24–26µm width, tip spoon shaped and pointed, ornamented with triangular teeth (
|
||||
<figureCitation id="1BF42A0C9172051DB970D7B56084FA49" box="[1296,1424,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-21@3.[151,1436,181,1817]" captionTargetPageId="3" captionText="FIGURE 2. Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. A. internal anatomy, B. spermathecae (dorsal side), C. penial setae 10x, D. penial setae 100x, E. Calciferous glands, F. Spermathecae (ventral side). Abbreviations: di, diverticulum; am, ampulla; gz, gizzard; ts, testes; sv, seminal vesicles; in, intestine; vd, vas deferens; pd, prostatic duct; pg, prostate glands; vdo, vas deferens opening; dv, dorsal blood vessel; vn, ventral nerve cord; cg, calciferous glands; he, hearts." figureDoi="http://doi.org/10.5281/zenodo.15215026" httpUri="https://zenodo.org/record/15215026/files/figure.png" pageId="4" pageNumber="240">Fig. 2C, D</figureCitation>
|
||||
). Genital markings glands sessile, associated with genital markings of segments 12, 13 and 14.
|
||||
</paragraph>
|
||||
<paragraph id="837036899172051DBCA7D46C6695F935" blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="240">
|
||||
<emphasis id="B1BBEA9B9172051DBCA7D46C645EF980" bold="true" box="[199,330,1554,1580]" pageId="4" pageNumber="240">Variations.</emphasis>
|
||||
Genital markings on segments 13, 14, and 16 are well-developed, whereas the markings on segments 14 and 15 exhibit a slightly whitish color (
|
||||
<specimenCount id="95C9FD009172051DBE16D44967E9F9FD" box="[630,765,1591,1617]" count="4" pageId="4" pageNumber="240" type="generic" typeStatus="paratype">4 paratypes</specimenCount>
|
||||
). Most of the dissected specimens bear one pair of the ental diverticulum, whereas one dissected
|
||||
<typeStatus id="5C74882B9172051DBE54D4256780F9D9" box="[564,660,1627,1653]" pageId="4" pageNumber="240" type="paratype">paratype</typeStatus>
|
||||
has three diverticulums. Sometimes the ampulla is constricted by the ventral cord, which forms dumbbell-shaped or irregular two lobes.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD565029172051DBCA7D4DC6667F911" box="[199,883,1698,1725]" pageId="4" pageNumber="240" type="distribution">
|
||||
<paragraph id="837036899172051DBCA7D4DC6667F911" blockId="4.[151,1437,150,2013]" box="[199,883,1698,1725]" pageId="4" pageNumber="240">
|
||||
<emphasis id="B1BBEA9B9172051DBCA7D4DC644DF910" bold="true" box="[199,345,1698,1724]" pageId="4" pageNumber="240">Distribution</emphasis>
|
||||
.
|
||||
<collectingCountry id="FBD876199172051DBD07D4DD64B1F911" box="[359,421,1699,1725]" name="India" pageId="4" pageNumber="240">India</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="410BF86B9172051DBDCFD4DD6751F911" box="[431,581,1699,1725]" country="India" name="Tripura" pageId="4" pageNumber="240">Tripura State</collectingRegion>
|
||||
: Khowai District: Khowai.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="CBD565029172051CBCA7D4B866CBFEC9" lastPageId="5" lastPageNumber="241" pageId="4" pageNumber="240" type="discussion">
|
||||
<paragraph id="837036899172051CBCA7D4B866CBFEC9" blockId="4.[151,1437,150,2013]" lastBlockId="5.[151,1437,151,357]" lastPageId="5" lastPageNumber="241" pageId="4" pageNumber="240">
|
||||
<emphasis id="B1BBEA9B9172051DBCA7D4B86421F94D" bold="true" box="[199,309,1734,1761]" italics="true" pageId="4" pageNumber="240">Remarks.</emphasis>
|
||||
The proposed species
|
||||
<taxonomicName id="44CF4D0A9172051DBE56D4B96107F94C" authority="Ahmed & Marimuthu" authorityName="Ahmed & Marimuthu" authorityYear="2025" box="[566,1043,1735,1761]" class="Clitellata" family="Acanthodrilidae" genus="Eutyphoeus" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="240" phylum="Annelida" rank="species" species="unithecus" status="sp. nov.">
|
||||
<emphasis id="B1BBEA9B9172051DBE56D4B9663FF94D" box="[566,811,1735,1761]" italics="true" pageId="4" pageNumber="240">Eutyphoeus unithecus</emphasis>
|
||||
Ahmed & Marimuthu
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="AA8857E09172051DB87DD4B96160F94D" box="[1053,1140,1735,1761]" pageId="4" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
|
||||
differs from its congeners in terms of the number of spermathecae, spermathecal pore position and male pore position. All known species of
|
||||
<taxonomicName id="44CF4D0A9172051DBCF7D571640EF885" authorityName="Michaelsen" authorityYear="1900" box="[151,282,1807,1833]" class="Clitellata" family="Acanthodrilidae" genus="Eutyphoeus" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="240" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B1BBEA9B9172051DBCF7D571640EF885" box="[151,282,1807,1833]" italics="true" pageId="4" pageNumber="240">Eutyphoeus</emphasis>
|
||||
</taxonomicName>
|
||||
have one pair of spermathecal pores in intersegmental furrow 7/8 and one pair of spermathecae, whereas the new species has a single spermathecal pore at the midventral line between
|
||||
<emphasis id="B1BBEA9B9172051DB850D54D6158F8E0" box="[1072,1100,1843,1868]" italics="true" pageId="4" pageNumber="240">aa</emphasis>
|
||||
and with a single associated spermatheca. Principal component analysis (
|
||||
<figureCitation id="1BF42A0C9172051DBEFCD52967EDF8DD" box="[668,761,1879,1905]" captionStart="FIGURE 3" captionStartId="5.[151,250,1770,1794]" captionTargetBox="[152,1434,385,1745]" captionTargetId="figure-121@5.[151,1435,381,1747]" captionTargetPageId="5" captionText="FIGURE 3. Multivariate analyses. Principal component analysis (A) of the proposed new species Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. based on the taxonomic characters, Canonical Analysis of Principal coordinates (CAP) of the proposed new species Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. based on number of spermathecae (B) and male pores position (C). Species code 1–52 and sp. nov. (Refer species list in supplementary data). Abbreviations: Ln, length; Wd, width; Sg, segments; Spm no., number of spermathecae; Spmp., spermathecal pore position with respect to setal line; Mpp, male pore position with respect to setal line; Gm, genital markings; Mvic, mid–ventral intestinal caeca; Sig, supra–intestinal glands; To, typhlosole origin; Pspb, penial setae per battery; Ps ln, penial setae length." figureDoi="http://doi.org/10.5281/zenodo.15215028" httpUri="https://zenodo.org/record/15215028/files/figure.png" pageId="4" pageNumber="240">Fig. 3A</figureCitation>
|
||||
) of species coming under the genus,
|
||||
<taxonomicName id="44CF4D0A9172051DB8D8D529602FF8DD" authorityName="Michaelsen" authorityYear="1900" box="[1208,1339,1879,1905]" class="Clitellata" family="Acanthodrilidae" genus="Eutyphoeus" kingdom="Animalia" order="Crassiclitellata" pageId="4" pageNumber="240" phylum="Annelida" rank="genus">
|
||||
<emphasis id="B1BBEA9B9172051DB8D8D529602FF8DD" box="[1208,1339,1879,1905]" italics="true" pageId="4" pageNumber="240">Eutyphoeus</emphasis>
|
||||
</taxonomicName>
|
||||
showed that there is a strong variability (PC1: 29.5% variance) between the species based on the observed taxonomical characters and the most influenced taxonomical characters observed were genital markings. There was also a variability (PC2: 14.2% variance) between the species and the influenced taxonomical characters observed were the number of spermathecae followed by the position of spermathecal pore and male pore. It was also observed that the proposed species became an outlier (distinguished) from all the species of the similar genus due to the distinct taxonomical characters such as spermathecae pores and male pore position followed by the number of spermathecae. Corresponding to the single spermathecal pore, the male openings are very close together in the proposed species. Based on these characteristics, CAP analysis used male pores position and number of spermathecae as factors to distinguish the proposed one from the other described species (
|
||||
<figureCitation id="1BF42A0C9173051CBF39D33566C6FEC9" box="[857,978,331,357]" captionStart="FIGURE 3" captionStartId="5.[151,250,1770,1794]" captionTargetBox="[152,1434,385,1745]" captionTargetId="figure-121@5.[151,1435,381,1747]" captionTargetPageId="5" captionText="FIGURE 3. Multivariate analyses. Principal component analysis (A) of the proposed new species Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. based on the taxonomic characters, Canonical Analysis of Principal coordinates (CAP) of the proposed new species Eutyphoeus unithecus Ahmed & Marimuthu, sp. nov. based on number of spermathecae (B) and male pores position (C). Species code 1–52 and sp. nov. (Refer species list in supplementary data). Abbreviations: Ln, length; Wd, width; Sg, segments; Spm no., number of spermathecae; Spmp., spermathecal pore position with respect to setal line; Mpp, male pore position with respect to setal line; Gm, genital markings; Mvic, mid–ventral intestinal caeca; Sig, supra–intestinal glands; To, typhlosole origin; Pspb, penial setae per battery; Ps ln, penial setae length." figureDoi="http://doi.org/10.5281/zenodo.15215028" httpUri="https://zenodo.org/record/15215028/files/figure.png" pageId="5" pageNumber="241">Fig. 3B, C</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
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Reference in a new issue