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<document id="205F57585CE945777E11F5435822439D" ID-CLB-Dataset="301818" ID-DOI="10.5281/zenodo.13272818" ID-GBIF-Dataset="25561bf2-ef51-4d75-a837-3789f9602786" ID-ISSN="2295-0214" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1723146878667" checkinUser="felipe" docAuthor="Vermeersch, Xavier H. C." docDate="2001" docId="30188791FFD72804FE41FCC885E8E183" docLanguage="en" docName="BelgJEntomol.100.1-18.pdf" docOrigin="Belgian Journal of Entomology 100" docSource="http://dx.doi.org/10.5281/zenodo.13272818" docStyle="DocumentStyle:1C24E9253A34C90F9A25106B2B38F3D3.1:BelgJEntomol.2015-2021.journal_article.type1" docStyleId="1C24E9253A34C90F9A25106B2B38F3D3" docStyleName="BelgJEntomol.2015-2021.journal_article.type1" docStyleVersion="1" docTitle="Titanodula fruhstorferi Vermeersch 2001, comb. nov." docType="treatment" docVersion="2" lastPageNumber="18" masterDocId="CC21FFE9FFDC280BFFB1FF858655E17B" masterDocTitle="Euchomenella adwinae sp. nov., a small stick mantis from southern Central Vietnam (Mantodea: Mantidae: Deroplatyinae: Euchomenellini)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1724918174485" updateUser="ExternalLinkService">
<mods:mods id="930C5F8F702F9B40348EB3D2F3172F43" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE873FADBF2D4FE9D18EBA182B1877D6">
<mods:title id="50AF2DF719F155528C58A00943F5300A">Euchomenella adwinae sp. nov., a small stick mantis from southern Central Vietnam (Mantodea: Mantidae: Deroplatyinae: Euchomenellini)</mods:title>
</mods:titleInfo>
<mods:name id="9923DFF7C9A39A45AF68E891DF763D42" type="personal">
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<mods:namePart id="8C03B3DF24707F3CDED581FACC426908">Vermeersch, Xavier H. C.</mods:namePart>
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<mods:typeOfResource id="C6427FEBFAE949F81985A7F2C45CC12A">text</mods:typeOfResource>
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<mods:title id="50A947855BE3C592681D7468805280F3">Belgian Journal of Entomology</mods:title>
</mods:titleInfo>
<mods:part id="AFC21C77F4BAA5813A6FF752E267ACDC">
<mods:date id="F4EDD7A0BA6DEF0D0A7351A7D2CF57EB">2001</mods:date>
<mods:detail id="4FEE708A3FE3195CE60EB258E448991B" type="volume">
<mods:number id="C6750CC3768C40CCD8FB75776F727E9B">100</mods:number>
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<mods:location id="2B91701530C0838FBF61D8CA3DB21718">
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<mods:classification id="02B666D40C5ACF68405E3D4DE0338F77">journal article</mods:classification>
<mods:identifier id="AA6C9EC3E5D7520B59944E8373203FBC" type="CLB-Dataset">301818</mods:identifier>
<mods:identifier id="3D180C253816778D9E9DB43A57B236D1" type="DOI">10.5281/zenodo.13272818</mods:identifier>
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<mods:identifier id="606C14E8024EEAC7CD0A5DC5FA19A729" type="ISSN">2295-0214</mods:identifier>
</mods:mods>
<treatment id="30188791FFD72804FE41FCC885E8E183" ID-DOI="http://doi.org/10.5281/zenodo.13449993" ID-Zenodo-Dep="13449993" LSID="urn:lsid:plazi:treatment:30188791FFD72804FE41FCC885E8E183" httpUri="http://treatment.plazi.org/id/30188791FFD72804FE41FCC885E8E183" lastPageId="15" pageId="11">
<subSubSection id="F0AB650CFFD72800FE41FCC88208E213" box="[496,1117,845,872]" pageId="11" type="nomenclature">
<paragraph id="B80E3687FFD72800FE41FCC88208E213" blockId="11.[496,1117,845,907]" box="[496,1117,845,872]" pageId="11">
<emphasis id="8AC5EA95FFD72800FE41FCC88208E213" bold="true" box="[496,1117,845,872]" pageId="11">
<taxonomicName id="7FB14D04FFD72800FE41FCC8859FE21C" ID-CoL="8QPYL" authority="(Werner, 1916)" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[496,970,845,872]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="11" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD72800FE41FCC88546E213" bold="true" box="[496,787,845,872]" italics="true" pageId="11">Titanodula fruhstorferi</emphasis>
(Werner, 1916)
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD72800FC63FCCB8208E213" box="[978,1117,846,872]" pageId="11" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD72800FD40FCF4853AE2F0" box="[753,879,881,907]" pageId="11" type="description">
<paragraph id="B80E3687FFD72800FD40FCF4853AE2F0" blockId="11.[496,1117,845,907]" box="[753,879,881,907]" pageId="11">
(
<figureCitation id="208A2A02FFD72800FD48FCF48530E2F0" box="[761,869,881,907]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="11.[247,284,1721,1743]" captionStartId-1="12.[247,284,1798,1820]" captionStartId-2="14.[247,284,1835,1857]" captionTargetBox-0="[251,1383,1100,1695]" captionTargetBox-1="[354,1288,185,1775]" captionTargetBox-2="[298,1335,180,1812]" captionTargetId-0="figure-396@11.[247,1385,1099,1700]" captionTargetId-1="figure-74@12.[330,1303,178,1778]" captionTargetId-2="figure-60@14.[296,1336,178,1814]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Fig. 4. Titanodula fruhstorferi comb. nov., additional specimen ♂ [AS1 – Tam Dao N.P.] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. Titanodula fruhstorferi comb. nov., holotype ♀ (NHMW). E, habitus, dorsal view. F, habitus, ventral view. G, detail of head and prothoracic legs. H, labels. © A−D, Xavier Vermeersch and E−H, Harald Bruckner)" captionText-1="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." captionText-2="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." pageId="11">Figs 4−6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD72800FF46FC2B8350E54A" pageId="11" type="diagnosis">
<paragraph id="B80E3687FFD72800FF46FC2B8350E54A" blockId="11.[247,1386,941,1073]" pageId="11">
<smallCapsWord id="BEE8A05BFFD72800FF46FC2B8729E2BD" baselines="961,961" box="[247,380,942,968]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Diagnosis" pageId="11">DIAGNOSIS</smallCapsWord>
.
<taxonomicName id="7FB14D04FFD72800FE3DFC2884F1E2BC" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[396,676,941,967]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="11" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD72800FE3DFC2884F1E2BC" box="[396,676,941,967]" italics="true" pageId="11">Titanodula fruhstorferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD72800FD1CFC288560E2B3" box="[685,821,941,968]" pageId="11" rank="species">comb. nov.</taxonomicNameLabel>
is very similar to
<taxonomicName id="7FB14D04FFD72800FBA4FC2B82B5E2BC" authorityName="Vermeersch" authorityYear="2001" box="[1045,1248,941,967]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="11" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFD72800FBA4FC2B82B5E2BC" box="[1045,1248,941,967]" italics="true" pageId="11">T. attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD72800FB58FC2B831CE2B3" box="[1257,1353,942,968]" pageId="11" rank="species">sp. nov.</taxonomicNameLabel>
in external appearance although profemoral AvS 1, 2, 10 and 15 and the basal spots of AvS 2, 10 and 15 are noticeably less dark or pronounced compared to
<taxonomicName id="7FB14D04FFD72800FC0EFC7182DDE576" authorityName="Vermeersch" authorityYear="2001" box="[959,1160,1011,1037]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="11" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFD72800FC0EFC7182DDE576" box="[959,1160,1011,1037]" italics="true" pageId="11">T. attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD72800FB3EFC7182B8E575" box="[1167,1261,1012,1038]" pageId="11" rank="species">sp. nov.</taxonomicNameLabel>
However, both species can be readily differentiated by the unique morphology of male genitalia.
</paragraph>
</subSubSection>
<caption id="ECCE660FFFD72800FF46F93C823BE652" pageId="11" startId="11.[247,284,1721,1743]" targetBox="[251,1383,1100,1695]" targetPageId="11" targetType="figure">
<paragraph id="B80E3687FFD72800FF46F93C823BE652" blockId="11.[247,1386,1721,1833]" pageId="11">
Fig. 4.
<taxonomicName id="7FB14D04FFD72800FEF0F93C847EE7B5" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[321,555,1721,1742]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="11" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD72800FEF0F93C847EE7B5" box="[321,555,1721,1742]" italics="true" pageId="11">Titanodula fruhstorferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD72800FD83F93C84F0E7B5" box="[562,677,1721,1743]" pageId="11" rank="species">comb. nov.</taxonomicNameLabel>
, additional specimen ♂ [AS1 – Tam Dao N.P.] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view.
<taxonomicName id="7FB14D04FFD72800FB4AF952873EE671" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="11" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD72800FB4AF952873EE671" italics="true" pageId="11">Titanodula fruhstorferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD72800FEC0F97087B1E670" box="[369,484,1781,1803]" pageId="11" rank="species">comb. nov.</taxonomicNameLabel>
, holotype ♀ (NHMW). E, habitus, dorsal view. F, habitus, ventral view. G, detail of head and prothoracic legs. H, labels. © A−D, Xavier Vermeersch and E−H, Harald Bruckner)
</paragraph>
</caption>
<caption id="ECCE660FFFD02807FF46F88382A6E60D" pageId="12" startId="12.[247,284,1798,1820]" targetBox="[354,1288,185,1775]" targetPageId="12" targetType="figure">
<paragraph id="B80E3687FFD02807FF46F88382A6E60D" blockId="12.[247,1385,1798,1910]" pageId="12">
Fig. 5.
<taxonomicName id="7FB14D04FFD02807FEF1F883847DE660" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[320,552,1798,1819]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="12" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD02807FEF1F883847DE660" box="[320,552,1798,1819]" italics="true" pageId="12">Titanodula fruhstorferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD02807FD9EF88384F6E667" box="[559,675,1798,1820]" pageId="12" rank="species">comb. nov.</taxonomicNameLabel>
, additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view.
<taxonomicName id="7FB14D04FFD02807FC18F8A182F9E642" authorityName="Vermeersch" authorityYear="2001" box="[937,1196,1828,1849]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="12" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFD02807FC18F8A182F9E642" box="[937,1196,1828,1849]" italics="true" pageId="12">Titanodula attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD02807FB05F8A08353E641" box="[1204,1286,1829,1850]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet.
</paragraph>
</caption>
<subSubSection id="F0AB650CFFD12806FF46FF3E8285E18C" pageId="13" type="materials_examined">
<paragraph id="B80E3687FFD12806FF46FF3E8285E18C" blockId="13.[247,1386,183,248]" pageId="13">
<materialsCitation id="08D93CDAFFD12806FF46FF3E8285E18C" collectionCode="MATERIAL, TYPE, NHMW" collectorName="H. Fruhstorfer" country="Vietnam" location="Vietnam" pageId="13" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<collectionCode id="DEA0AE42FFD12806FF46FF3E872DE1A8" box="[247,376,187,213]" pageId="13">MATERIAL</collectionCode>
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.
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<collectionCode id="DEA0AE42FFD12806FDEDFF3B8482E1A8" box="[604,727,190,211]" pageId="13">MATERIAL</collectionCode>
</smallCapsWord>
(examined from detailed photographs):
<typeStatus id="670A8825FFD12806FB67FF3F8313E1AF" box="[1238,1350,186,212]" pageId="13" type="holotype">Holotype</typeStatus>
<emphasis id="8AC5EA95FFD12806FAE2FF32833FE1AF" bold="true" box="[1363,1386,183,212]" pageId="13">
<specimenCount id="AEB7FD0EFFD12806FAE2FF32833FE1AF" box="[1363,1386,183,212]" count="1" pageId="13" type="female">♀</specimenCount>
</emphasis>
(
<figureCitation id="208A2A02FFD12806FEB1FF588712E18C" box="[256,327,221,247]" captionStart="Fig" captionStartId="11.[247,284,1721,1743]" captionTargetBox="[251,1383,1100,1695]" captionTargetId="figure-396@11.[247,1385,1099,1700]" captionTargetPageId="11" captionText="Fig. 4. Titanodula fruhstorferi comb. nov., additional specimen ♂ [AS1 – Tam Dao N.P.] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. Titanodula fruhstorferi comb. nov., holotype ♀ (NHMW). E, habitus, dorsal view. F, habitus, ventral view. G, detail of head and prothoracic legs. H, labels. © A−D, Xavier Vermeersch and E−H, Harald Bruckner)" pageId="13">Fig. 4</figureCitation>
E−F):
<collectingCountry id="C0A67617FFD12806FE2CFF58845DE18C" box="[413,520,221,247]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
, Tonkin, Than Moi, VI−VII, leg.
<collectorName id="15445351FFD12806FC2FFF5B821FE18C" box="[926,1098,221,248]" pageId="13">H. Fruhstorfer</collectorName>
(
<collectionCode id="DEA0AE42FFD12806FBEBFF588291E18C" box="[1114,1220,221,247]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD12806FF46FE888201E05C" box="[247,1108,269,295]" pageId="13" type="discussion">
<paragraph id="B80E3687FFD12806FF46FE888201E05C" blockId="13.[247,1108,269,295]" box="[247,1108,269,295]" pageId="13">
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<collectionCode id="DEA0AE42FFD12806FF46FE88876EE05D" box="[247,315,269,295]" pageId="13">NOTE</collectionCode>
</smallCapsWord>
:
<typeStatus id="670A8825FFD12806FEFAFE8887EEE05C" box="[331,443,269,295]" pageId="13" type="holotype">Holotype</typeStatus>
damaged, abdomen and metathoracic legs are missing.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD12806FF46FEB884C0E314" pageId="13" type="materials_examined">
<paragraph id="B80E3687FFD12806FF46FEB884C0E314" blockId="13.[247,1385,314,624]" pageId="13">
<smallCapsWord id="BEE8A05BFFD12806FF46FEB887C5E02E" baselines="336,336" box="[247,400,317,343]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Additional" pageId="13">ADDITIONAL</smallCapsWord>
<smallCapsWord id="BEE8A05BFFD12806FE2BFEC58440E02E" baselines="336" box="[410,533,320,341]" lowerCaseFontSize="9" mainFontSize="11" normCase="lower" normString="material" pageId="13">MATERIAL</smallCapsWord>
(
<specimenCount id="AEB7FD0EFFD12806FD99FEB88423E02C" box="[552,630,317,343]" count="14" pageId="13" type="male">14♂♂</specimenCount>
,
<specimenCount id="AEB7FD0EFFD12806FD34FEB884FEE02C" box="[645,683,314,343]" count="1" pageId="13" type="female">
1
<emphasis id="8AC5EA95FFD12806FD25FEBF84FEE02C" bold="true" box="[660,683,314,343]" pageId="13">♀</emphasis>
</specimenCount>
): (
<specimenCount id="AEB7FD0EFFD12806FD65FEB884AEE02C" box="[724,763,317,343]" count="1" pageId="13" type="male">1♂</specimenCount>
):
<materialsCitation id="08D93CDAFFD12806FCA6FEB98517E0E6" collectingDate="1833-01" collectionCode="R" collectorName="Tam Dao" country="Vietnam" latitude="22.2" location="Cham Chu" longitude="105.1" pageId="13" specimenCount="1" specimenCount-male="3">
<collectingCountry id="C0A67617FFD12806FCA6FEB985D7E02D" box="[791,898,316,342]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
,
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N.
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.,
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;
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,
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, Night Collecting, leg. J. Constant, J. Bresseel, Genitalia prep. Vermeersch Nr.
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, GTI project, I.G.: 33.092 (
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).
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<materialsCitation id="08D93CDAFFD12806FCFFFE06853DE098" collectingDate="1833-01" collectionCode="R" collectorName="Tam Dao" country="Vietnam" latitude="21.516666" location="Cham Chu" longitude="105.55" pageId="13" specimenCount="1" specimenCount-male="3">
(
<specimenCount id="AEB7FD0EFFD12806FCE9FE0685CCE0E6" box="[856,921,387,413]" count="3" pageId="13" type="male">3♂♂</specimenCount>
):
<collectingCountry id="C0A67617FFD12806FC02FE06824BE0E6" box="[947,1054,387,413]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
,
<collectorName id="15445351FFD12806FB9FFE0682F6E0E6" box="[1070,1187,387,413]" pageId="13">Tam Dao</collectorName>
N.P.,
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;
<geoCoordinate id="DD855040FFD12806FF46FE238738E0BA" box="[247,365,422,449]" degrees="105" direction="east" minutes="33" orientation="longitude" pageId="13" precision="925" value="105.55">105°33’E</geoCoordinate>
,
<date id="CC0F1047FFD12806FE30FE238412E0BA" box="[385,583,422,449]" pageId="13" value="2011-07-25" valueMax="2011-07-30" valueMin="2011-07-25">25−30.VII.2011</date>
, leg. J. Constant, J. Bresseel, Genitalia prep. Vermeersch Nr.
<date id="CC0F1047FFD12806FF46FE4C8721E098" box="[247,372,457,483]" pageId="13" value="1909-01">
<collectingDate id="DC4BE9AFFFD12806FF46FE4C8721E098" box="[247,372,457,483]" pageId="13" value="1909-01">GEN1909</collectingDate>
</date>
, GTI project, I.G.: 31.933 (
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).
</materialsCitation>
<materialsCitation id="08D93CDAFFD12806FCCAFE4C826DE351" collectingDate="1833-01" collectionCode="R" collectorName="Tam Dao" country="Vietnam" latitude="20.316668" location="Cham Chu" longitude="105.60833" pageId="13" specimenCount="1" specimenCount-female="1" specimenCount-male="3">
(
<specimenCount id="AEB7FD0EFFD12806FC34FE4C8583E098" box="[901,982,457,483]" count="10" pageId="13" type="male">10♂♂</specimenCount>
):
<collectingCountry id="C0A67617FFD12806FC44FE4C8235E098" box="[1013,1120,457,483]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
,
<collectorName id="15445351FFD12806FBC9FE4C834DE098" box="[1144,1304,457,483]" pageId="13">Cuc Phuong</collectorName>
N.P.,
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;
<geoCoordinate id="DD855040FFD12806FE2BFE698469E37C" box="[410,572,492,519]" degrees="105" direction="east" minutes="36" orientation="longitude" pageId="13" precision="15" seconds="30" value="105.60833">105°36’30”E</geoCoordinate>
,
<date id="CC0F1047FFD12806FDF6FE698558E37D" box="[583,781,492,518]" pageId="13" value="2011-07-19" valueMax="2011-07-23" valueMin="2011-07-19">19−23.VII.2011</date>
, Light Trap, leg. J. Constant, J. Bresseel, Genitalia prep. Vermeersch Nr.
<date id="CC0F1047FFD12806FDB5FD8A84D4E352" box="[516,641,527,553]" pageId="13" value="1839-01">
<collectingDate id="DC4BE9AFFFD12806FDB5FD8A84D4E352" box="[516,641,527,553]" pageId="13" value="1839-01">GEN1839</collectingDate>
</date>
, GTI project, I.G.: 31.933 (
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).
</materialsCitation>
<materialsCitation id="08D93CDAFFD12806FB8EFD8A84C4E314" collectingDate="1833-01" collectingDateMax="2013-07-11" collectingDateMin="2013-07-07" collectionCode="RBINS" collectorName="J. Bresseel" country="Vietnam" latitude="21.18611" location="Cham Chu" longLatPrecision="21" longitude="106.72389" pageId="13" specimenCount="1" specimenCount-female="1" specimenCount-male="3">
(
<specimenCount id="AEB7FD0EFFD12806FBFAFD8A8225E352" box="[1099,1136,524,553]" count="1" pageId="13" type="female">
1
<emphasis id="8AC5EA95FFD12806FBE8FD898225E352" bold="true" box="[1113,1136,524,553]" pageId="13">♀</emphasis>
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):
<collectingCountry id="C0A67617FFD12806FB38FD8A82A1E352" box="[1161,1268,527,553]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
,
<collectorName id="15445351FFD12806FAB1FD95874DE336" pageId="13">Tay Yen Tu</collectorName>
<taxonomicNameLabel id="91F657EEFFD12806FE9AFDB68737E336" box="[299,354,563,589]" pageId="13" rank="subSpecies">
N.
<collectionCode id="DEA0AE42FFD12806FEFBFDB68737E336" box="[330,354,563,589]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13">R</collectionCode>
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.,
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;
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,
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<collectingDate id="DC4BE9AFFFD12806FD56FDB785C8E337" box="[743,925,562,588]" pageId="13" value="2013-07-07" valueMax="2013-07-11" valueMin="2013-07-07">7−11.VII.2013</collectingDate>
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,
<collectorName id="15445351FFD12806FC04FDB68222E337" box="[949,1143,562,589]" pageId="13">Day Collecting</collectorName>
, leg.
<collectorName id="15445351FFD12806FB61FDB68331E337" box="[1232,1380,562,589]" pageId="13">J. Constant</collectorName>
,
<collectorName id="15445351FFD12806FF46FDD3872EE314" box="[247,379,597,624]" pageId="13">J. Bresseel</collectorName>
, I.G.: 32.454 (
<collectionCode id="DEA0AE42FFD12806FD98FDD084DDE314" box="[553,648,597,623]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="13" type="Museum">RBINS</collectionCode>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD12806FF46FD0387D5E761" pageId="13" type="description">
<paragraph id="B80E3687FFD12806FF46FD038207E209" blockId="13.[247,1386,645,882]" pageId="13">
<smallCapsWord id="BEE8A05BFFD12806FF46FD0387E1E3E5" baselines="665,665" box="[247,436,646,672]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Redescription" pageId="13">REDESCRIPTION</smallCapsWord>
. Measurements (see
<tableCitation id="F533033CFFD12806FD04FD008515E3E4" box="[693,832,645,671]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="7.[247,304,421,443]" captionStartId-1="9.[247,304,316,338]" captionText-0="Table 1. Detailed measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. From left to right: T. attenboroughi holotype ♂, T. attenboroughi paratype ♀, T. fruhstorferi male, additional specimen 1, T. fruhstorferi female, additional specimen 2." captionText-1="Table 2. Measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. to assess intraspecific variability." pageId="13">Tables 1−2</tableCitation>
). Very large and robust praying mantis. Head triangular, antennae filiform. Long but robust pronotum, with smooth dorsal surface. Pronotum very finely denticulate along the margins of the prozone in female, without denticulations in the metazone, entirely smooth edges in males. Spinal formula: F = 4DS/15AvS/4PvS; T = 14AvS/10PvS. Ratios
<specimenCount id="AEB7FD0EFFD12806FDB7FC978475E257" box="[518,544,786,812]" count="1" pageId="13" type="male">♂</specimenCount>
: MzL/PzL: 3.1;
<specimenCount id="AEB7FD0EFFD12806FD57FC9784AAE257" box="[742,767,786,812]" count="1" pageId="13" type="female">♀</specimenCount>
: MzL/PzL: 3. With black spots on the anterior side of the profemora located at the base of the 2
<superScript id="4FC49BCFFFD12806FCBEFCAA8576E239" attach="none" box="[783,803,815,834]" fontSize="8" pageId="13">nd</superScript>
, 10
<superScript id="4FC49BCFFFD12806FCE0FCAA8534E239" attach="left" box="[849,865,815,834]" fontSize="8" pageId="13">th</superScript>
and 15
<superScript id="4FC49BCFFFD12806FC0AFCAA859EE239" attach="left" box="[955,971,815,834]" fontSize="8" pageId="13">th</superScript>
anteroventral spine. Protarsus (all segments) entirely black on the anterior side. Both sexes macropterous.
</paragraph>
<paragraph id="B80E3687FFD12806FF46FC0287D5E761" blockId="13.[247,1386,903,1562]" pageId="13">
<emphasis id="8AC5EA95FFD12806FF46FC0287FCE2DA" box="[247,425,903,929]" italics="true" pageId="13">Male genitalia</emphasis>
(
<figureCitation id="208A2A02FFD12806FE0BFC028471E2D9" box="[442,548,903,930]" captionStart="Fig" captionStartId="12.[247,284,1798,1820]" captionTargetBox="[354,1288,185,1775]" captionTargetId="figure-74@12.[330,1303,178,1778]" captionTargetPageId="12" captionText="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." pageId="13">Fig. 5 A</figureCitation>
): Ventral phallomere of left phallic complex elongated, much longer than broad, with two short and strongly sclerotised distal processes (sdp) of about the same length with acute apex. Median process (sdpm) bend at the middle and pointing sideways, lateral process (sdpl) much more strongly bend and pointing upwards, with much broader base than sdpm. Apical process of left phallomere (paa) complex in shape, with sclerotised apex and sclerotizations near the base, but membranous in the middle. Apex abruptly projecting dorsally and to the left, the apex sharply curved in a 90° hook with narrower but rounded tip. Phalloid apophysis (afa) with two sclerotised processes, anterior process (aafa) small and flat, tubercle-shaped, without additional projections or extensions. Posterior process (pafa) spear-shaped, with very strongly developed base, bulging anteriorly, strongly curving towards an almost straight, very long and heavily sclerotised structure, projecting straight posteriad. Membranous lobe loa rounded and small. Genital lobe goa membranous, sometimes well-developed anteriorly but in some specimens also completely collapsed, and much weaker and wrinkled in appearance posteriorly, running almost parallel to pafa posteriad. Right phallomere with sclerite R1A more or less triangular. Sclerite R3 shovel-shaped, very wide at its anterior end. Ventral process (pia) triangular with many longitudinal grooves on its surface, sclerified ventral process (pva) strongly sclerotized, finger-shaped, ending in a blunt hook. Coxosternite IX (subgenital plate) with small black spikes on the right lateral edge, sometimes also in the middle of the distal edge.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD12806FF46F9B5845DE7EB" pageId="13" type="etymology">
<paragraph id="B80E3687FFD12806FF46F9B5845DE7EB" blockId="13.[247,1385,1583,1680]" pageId="13">
<smallCapsWord id="BEE8A05BFFD12806FF46F9B587C4E733" baselines="1603,1603" box="[247,401,1584,1610]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Etymology" pageId="13">ETYMOLOGY</smallCapsWord>
. This species was described in 1916 by the Austrian taxonomist Franz Werner (1867–1939) in honour of Hans Fruhstorfer (1866−1922), a German explorer, insect trader and
<taxonomicName id="7FB14D04FFD12806FF46F9F387DCE7EB" authorityName="Linnaeus" authorityYear="1758" box="[247,393,1654,1680]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="13" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
specialist.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD12804FF46F92385E8E183" lastPageId="15" pageId="13" type="distribution">
<paragraph id="B80E3687FFD12804FF46F92385E8E183" blockId="13.[247,1385,1701,1763]" lastBlockId="15.[247,1385,186,248]" lastPageId="15" pageId="13">
<smallCapsWord id="BEE8A05BFFD12806FF46F92387F6E7C5" baselines="1721,1721" box="[247,419,1702,1728]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Distribution" pageId="13">DISTRIBUTION</smallCapsWord>
.
<taxonomicName id="7FB14D04FFD12806FE04F9238433E7C4" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[437,614,1701,1727]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="13" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD12806FE04F9238433E7C4" box="[437,614,1701,1727]" italics="true" pageId="13">T. fruhstorferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD12806FDC5F9208554E7BB" box="[628,769,1701,1728]" pageId="13" rank="species">comb. nov.</taxonomicNameLabel>
inhabits the closed canopy forests of the North Vietnamese lowlands (
<figureCitation id="208A2A02FFD12806FDA3F94D8409E799" box="[530,604,1736,1762]" captionStart="Fig" captionStartId="14.[247,284,1835,1857]" captionTargetBox="[298,1335,180,1812]" captionTargetId="figure-60@14.[296,1336,178,1814]" captionText="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." pageId="13">Fig. 6</figureCitation>
). Forests in this area are highly threatened due to a high rate of deforestation and suitable habitats are strongly fragmented with large gaps between them. More distribution data is needed to assess its conservation status.
</paragraph>
</subSubSection>
</treatment>
</document>

196
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<subSubSection id="F0AB650CFFD8280FFD33FA9A858BE442" box="[642,990,1311,1338]" pageId="4" type="nomenclature">
<paragraph id="B80E3687FFD8280FFD33FA9A858BE442" blockId="4.[445,1188,1311,1408]" box="[642,990,1311,1338]" pageId="4">
<emphasis id="8AC5EA95FFD8280FFD33FA9A858BE442" bold="true" box="[642,990,1311,1338]" pageId="4">
Genus
<taxonomicName id="7FB14D04FFD8280FFD6AFA9A8531E441" ID-CoL="8Q99S" authorityName="Vermeersch" authorityYear="2001" box="[731,868,1311,1338]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8AC5EA95FFD8280FFD6AFA9A8531E441" bold="true" box="[731,868,1311,1338]" italics="true" pageId="4">Titanodula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD8280FFCDDFA9A858BE442" box="[876,990,1311,1337]" pageId="4" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD8280FFE0CFAC385DBE4FB" pageId="4" type="description">
<paragraph id="B80E3687FFD8280FFE0CFAC382F1E427" blockId="4.[445,1188,1311,1408]" box="[445,1188,1346,1372]" pageId="4">
<uri id="CC203A85FFD8280FFE0CFAC382F1E427" box="[445,1188,1346,1372]" pageId="4">
urn:lsid:zoobank.org:act:
<uuid id="CC170C52FFD8280FFD08FAC382F1E427" box="[697,1188,1350,1372]" pageId="4">AFA3EEA0-6982-45D0-B294-3C8B123BD95F</uuid>
</uri>
</paragraph>
<paragraph id="B80E3687FFD8280FFD63FAE085DBE4FB" blockId="4.[445,1188,1311,1408]" box="[722,910,1381,1408]" pageId="4">
(
<figureCitation id="208A2A02FFD8280FFD6BFAE0851DE404" box="[730,840,1381,1407]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[247,284,1261,1283]" captionStartId-1="8.[247,284,1459,1481]" captionStartId-2="10.[247,284,1431,1453]" captionTargetBox-0="[254,1376,536,1218]" captionTargetBox-1="[256,1378,887,1429]" captionTargetBox-2="[247,1385,273,1410]" captionTargetId-0="figure-400@6.[247,1385,529,1227]" captionTargetId-1="figure-458@8.[247,1385,870,1438]" captionTargetId-2="figure-209@10.[247,1385,273,1410]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-1="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-2="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="4">Figs 1−3</figureCitation>
,
<figureCitation id="208A2A02FFD8280FFCE4FAE085D0E4FB" box="[853,901,1381,1408]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[247,284,1798,1820]" captionStartId-1="14.[247,284,1835,1857]" captionTargetBox-0="[354,1288,185,1775]" captionTargetBox-1="[298,1335,180,1812]" captionTargetId-0="figure-74@12.[330,1303,178,1778]" captionTargetId-1="figure-60@14.[296,1336,178,1814]" captionTargetPageId-0="12" captionText-0="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." captionText-1="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." pageId="4">5−6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD8280FFF46FA2785ADE4C7" box="[247,1016,1442,1468]" pageId="4" type="type_taxon">
<paragraph id="B80E3687FFD8280FFF46FA2785ADE4C7" blockId="4.[247,1016,1442,1468]" box="[247,1016,1442,1468]" pageId="4">
Type species:
<taxonomicName id="7FB14D04FFD8280FFE12FA27848FE4C7" authorityName="Vermeersch" authorityYear="2001" box="[419,730,1442,1468]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="species" species="attenboroughi">
<emphasis id="8AC5EA95FFD8280FFE12FA27848FE4C7" box="[419,730,1442,1468]" italics="true" pageId="4">Titanodula attenboroughi</emphasis>
</taxonomicName>
by present designation.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD8280FFF46FA5984ECE622" pageId="4" type="diagnosis">
<paragraph id="B80E3687FFD8280FFF46FA598254E763" blockId="4.[247,1386,1499,1560]" pageId="4">
<smallCapsWord id="BEE8A05BFFD8280FFF46FA5987F1E48F" baselines="1519,1519" box="[247,420,1500,1526]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Differential" pageId="4">DIFFERENTIAL</smallCapsWord>
<smallCapsWord id="BEE8A05BFFD8280FFE1FFA5A847AE48F" baselines="1519" box="[430,559,1503,1524]" lowerCaseFontSize="9" mainFontSize="11" normCase="lower" normString="diagnosis" pageId="4">DIAGNOSIS</smallCapsWord>
.
<taxonomicName id="7FB14D04FFD8280FFDF1FA5E8491E48E" authorityName="Vermeersch" authorityYear="2001" box="[576,708,1499,1525]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8AC5EA95FFD8280FFDF1FA5E8491E48E" box="[576,708,1499,1525]" italics="true" pageId="4">Titanodula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD8280FFD61FA598510E48D" box="[720,837,1500,1526]" pageId="4" rank="genus">gen. nov.</taxonomicNameLabel>
can be distinguished from all other genera within
<taxonomicName id="7FB14D04FFD8280FFEFDFA7B87EBE763" authorityName="Burmeister" authorityYear="1838" box="[332,446,1534,1560]" class="Insecta" family="Mantidae" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="family">Mantidae</taxonomicName>
by the combination of the following characters:
</paragraph>
<paragraph id="B80E3687FFD8280FFF46F9AB84F2E733" blockId="4.[247,679,1582,1608]" box="[247,679,1582,1608]" pageId="4">1) Large and robust praying mantis.</paragraph>
<paragraph id="B80E3687FFD8280FFF46F9DB8574E7E0" blockId="4.[247,1385,1629,1691]" pageId="4">2) Lower frons with two vertical margins within, interrupted in the middle, forming two small but distinct tubercles at its anterior margin.</paragraph>
<paragraph id="B80E3687FFD8280FFF46F935856AE7B1" blockId="4.[247,831,1712,1739]" box="[247,831,1712,1739]" pageId="4">3) Ventral coxal lobe darkened or black ventrad.</paragraph>
<paragraph id="B80E3687FFD8280FFF46F96582CBE781" blockId="4.[247,1182,1760,1787]" box="[247,1182,1760,1787]" pageId="4">4) Darkened or black spots at the base of 2nd, 10th and 15th profemoral AvS.</paragraph>
<paragraph id="B80E3687FFD8280FFF46F89584A1E651" blockId="4.[247,756,1808,1835]" box="[247,756,1808,1835]" pageId="4">5) All protarsal segments black anteriorly.</paragraph>
<paragraph id="B80E3687FFD8280FFF46F8BA84ECE622" blockId="4.[247,697,1855,1882]" box="[247,697,1855,1882]" pageId="4">6) Male genitalia as described below.</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD9280EFF46FF3E87BFE250" pageId="5" type="description">
<paragraph id="B80E3687FFD9280EFF46FF3E82BCE18C" blockId="5.[247,1385,186,248]" pageId="5">
<smallCapsWord id="BEE8A05BFFD9280EFF46FF3E87C2E1A8" baselines="206,206" box="[247,407,187,213]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Description" pageId="5">DESCRIPTION</smallCapsWord>
. Large and very robust praying mantises. Females macropterous, tegmina cover the abdomen completely. Male macropterous, tegmina extend beyond the abdomen.
</paragraph>
<paragraph id="B80E3687FFD9280EFF46FE888341E031" blockId="5.[247,1385,269,330]" pageId="5">
<emphasis id="8AC5EA95FFD9280EFF46FE88876DE05C" box="[247,312,269,295]" italics="true" pageId="5">Head</emphasis>
: triangular, broader than high, with large rounded eyes. Antennae filiform. Long but robust pronotum, with smooth dorsal surface, without projections or lateral expansions.
</paragraph>
<paragraph id="B80E3687FFD9280EFF46FEDA823EE3E8" blockId="5.[247,1385,351,659]" pageId="5">
<emphasis id="8AC5EA95FFD9280EFF46FEDA871BE002" box="[247,334,351,377]" italics="true" pageId="5">Thorax</emphasis>
: pronotum very finely denticulate along the margins of the prozone without denticulations in the metazone in females, entirely smooth edges in males. Dorsal side of the pronotum covered by a very thin whitish waxy layer in adults of both sexes that can easily be removed by direct contact. Profemora with 4 discoidal spines, 15 anteroventral spines and 4 posteroventral spines. Protibiae with 13−14 (rarely 15) anteroventral spines and 10 posteroventral spines. Protarsus (all segments) entirely black anteriorly, concolour with rest of the forelegs (green in all studied specimens) on the posteriorly. Ratios:
<specimenCount id="AEB7FD0EFFD9280EFBDDFDB682D5E336" box="[1132,1152,563,589]" count="1" pageId="5" type="male">♂</specimenCount>
MzL/PzL: 3.2;
<specimenCount id="AEB7FD0EFFD9280EFAFBFDB68334E336" box="[1354,1377,563,589]" count="1" pageId="5" type="female">♀</specimenCount>
: MzL/PzL: 3. All known species have darkened or blackish spots on the anterior side of the profemora located at the base of the 2
<superScript id="4FC49BCFFFD9280EFD0CFDF68484E3FD" attach="none" box="[701,721,627,646]" fontSize="8" pageId="5">nd</superScript>
, 10
<superScript id="4FC49BCFFFD9280EFD4FFDF6855BE3FD" attach="left" box="[766,782,627,646]" fontSize="8" pageId="5">th</superScript>
and 15
<superScript id="4FC49BCFFFD9280EFCD7FDF68523E3FD" attach="left" box="[870,886,627,646]" fontSize="8" pageId="5">th</superScript>
anteroventral spine.
</paragraph>
<paragraph id="B80E3687FFD9280EFF46FD2D87BFE250" blockId="5.[247,1385,680,812]" pageId="5">
<emphasis id="8AC5EA95FFD9280EFF46FD2D87F2E3B9" box="[247,423,680,706]" italics="true" pageId="5">Male genitalia</emphasis>
: with left phallic complex elongated and relatively narrow, lobe L4A projecting anteriorly; afa with small tubercle-shaped aafa and larger pafa with spear shaped projection oriented posteriad; distal processes (sdp) adjacent, relatively short but very strongly curved and about the same size.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD9280EFF46FCC78581E2BE" pageId="5" type="etymology">
<paragraph id="B80E3687FFD9280EFF46FCC78581E2BE" blockId="5.[247,1385,833,965]" pageId="5">
<smallCapsWord id="BEE8A05BFFD9280EFF46FCC787C4E221" baselines="853,853" box="[247,401,834,860]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Etymology" pageId="5">ETYMOLOGY</smallCapsWord>
. The genus name is derived from the ‘Titans’ in Greek mythology, who were a mythological race of giants, thus emphasizing the large size and strength of the species belonging to this genus. The second part of the genus name “-dula” is a reference to the genus
<taxonomicName id="7FB14D04FFD9280EFF46FC2E8726E2BE" authorityName="Burmeister" authorityYear="1838" box="[247,371,939,965]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="5" phylum="Arthropoda" rank="genus">
<emphasis id="8AC5EA95FFD9280EFF46FC2E8726E2BE" box="[247,371,939,965]" italics="true" pageId="5">Hierodula</emphasis>
</taxonomicName>
in which some species were previously described.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD9280EFF46FC5E82ADE525" pageId="5" type="distribution">
<paragraph id="B80E3687FFD9280EFF46FC5E82ADE525" blockId="5.[247,1385,986,1118]" pageId="5">
<smallCapsWord id="BEE8A05BFFD9280EFF46FC5E87F6E288" baselines="1006,1006" box="[247,419,987,1013]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Distribution" pageId="5">DISTRIBUTION</smallCapsWord>
.
<taxonomicName id="7FB14D04FFD9280EFE01FC5F8461E28F" authorityName="Vermeersch" authorityYear="2001" box="[432,564,986,1012]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="8AC5EA95FFD9280EFE01FC5F8461E28F" box="[432,564,986,1012]" italics="true" pageId="5">Titanodula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD9280EFD8DFC5E84FEE28E" box="[572,683,987,1013]" pageId="5" rank="genus">gen. nov.</taxonomicNameLabel>
has a confirmed presence in
<collectingCountry id="C0A67617FFD9280EFBBBFC5F82CCE28F" box="[1034,1177,986,1012]" name="Bangladesh" pageId="5">Bangladesh</collectingCountry>
, Southern
<collectingCountry id="C0A67617FFD9280EFAABFC5F8330E28F" box="[1306,1381,986,1012]" name="China" pageId="5">China</collectingCountry>
,
<collectingCountry id="C0A67617FFD9280EFF46FC788704E56C" box="[247,337,1021,1047]" name="Taiwan" pageId="5">Taiwan</collectingCountry>
and
<collectingCountry id="C0A67617FFD9280EFE38FC7887A0E56C" box="[393,501,1021,1047]" name="Vietnam" pageId="5">Vietnam</collectingCountry>
. It appears that the genus has a wide distribution across the Oriental region but remains limited to suitable forested habitats, making these giant praying mantises vulnerable to habitat loss and deforestation as a result of expanding human activities.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD9280EFF46FBF684C3E5C6" pageId="5" type="taxon_list">
<paragraph id="B80E3687FFD9280EFF46FBF6878AE5F5" blockId="5.[247,479,1139,1166]" box="[247,479,1139,1166]" pageId="5">
<smallCapsWord id="BEE8A05BFFD9280EFF46FBF6870DE5F7" baselines="1158,1159" box="[247,344,1139,1165]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Species" pageId="5">SPECIES</smallCapsWord>
<smallCapsWord id="BEE8A05BFFD9280EFEECFBF28782E5F7" baselines="1159" box="[349,471,1143,1164]" lowerCaseFontSize="9" mainFontSize="11" normCase="lower" normString="included" pageId="5">INCLUDED</smallCapsWord>
:
</paragraph>
<paragraph id="B80E3687FFD9280EFF46FB2684C3E5C6" blockId="5.[247,897,1187,1326]" box="[247,662,1187,1213]" pageId="5">
<taxonomicName id="7FB14D04FFD9280EFF46FB26847BE5C6" authorityName="Vermeersch" authorityYear="2001" box="[247,558,1187,1213]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="5" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFD9280EFF46FB26847BE5C6" box="[247,558,1187,1213]" italics="true" pageId="5">Titanodula attenboroughi</emphasis>
</taxonomicName>
<emphasis id="8AC5EA95FFD9280EFD84FB2684C3E5C6" bold="true" box="[565,662,1187,1213]" pageId="5">
<taxonomicNameLabel id="91F657EEFFD9280EFD84FB2684C3E5C6" box="[565,662,1187,1213]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD9280EFF46FB4D8511E455" pageId="5" type="description">
<paragraph id="B80E3687FFD9280EFF46FB4D85D4E599" blockId="5.[247,897,1187,1326]" box="[247,897,1224,1250]" pageId="5">
<taxonomicName id="7FB14D04FFD9280EFF46FB4D84BBE599" authority="(Giglio-Tos, 1912)" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Giglio-Tos" baseAuthorityYear="1912" box="[247,750,1224,1250]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="5" phylum="Arthropoda" rank="species" species="formosana" status="comb. nov.">
<emphasis id="8AC5EA95FFD9280EFF46FB4D8451E599" box="[247,516,1224,1250]" italics="true" pageId="5">Titanodula formosana</emphasis>
(Giglio-Tos, 1912)
</taxonomicName>
<emphasis id="8AC5EA95FFD9280EFD47FB4D85D4E599" bold="true" box="[758,897,1224,1250]" pageId="5">
<taxonomicNameLabel id="91F657EEFFD9280EFD47FB4D85D4E599" box="[758,897,1224,1250]" pageId="5" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="B80E3687FFD9280EFF46FB6B8534E473" blockId="5.[247,897,1187,1326]" box="[247,865,1262,1288]" pageId="5">
<taxonomicName id="7FB14D04FFD9280EFF46FB6B849BE473" authority="(Werner, 1916)" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[247,718,1262,1288]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="5" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD9280EFF46FB6B845BE473" box="[247,526,1262,1288]" italics="true" pageId="5">Titanodula fruhstorferi</emphasis>
(Werner, 1916)
</taxonomicName>
<emphasis id="8AC5EA95FFD9280EFD67FB6B8534E473" bold="true" box="[726,865,1262,1288]" pageId="5">
<taxonomicNameLabel id="91F657EEFFD9280EFD67FB6B8534E473" box="[726,865,1262,1288]" pageId="5" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="B80E3687FFD9280EFF46FA968511E455" blockId="5.[247,897,1187,1326]" box="[247,836,1299,1326]" pageId="5">
<taxonomicName id="7FB14D04FFD9280EFF46FA9684E5E456" authority="(Saussure, 1870)" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Saussure" baseAuthorityYear="1870" box="[247,688,1299,1325]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="5" phylum="Arthropoda" rank="species" species="grandis" status="comb. nov.">
<emphasis id="8AC5EA95FFD9280EFF46FA96878AE456" box="[247,479,1299,1325]" italics="true" pageId="5">Titanodula grandis</emphasis>
(Saussure, 1870)
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD9280EFD09FA968511E455" box="[696,836,1299,1326]" pageId="5" rank="species">
<emphasis id="8AC5EA95FFD9280EFD09FA968568E456" bold="true" box="[696,829,1299,1325]" pageId="5">comb. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="8ABD3005D3B4BE5762A47813F6B2BAE7">Euchomenella adwinae sp. nov., a small stick mantis from southern Central Vietnam (Mantodea: Mantidae: Deroplatyinae: Euchomenellini)</mods:title>
</mods:titleInfo>
<mods:name id="D21530E94290E562765D4CFCB5B0DAD0" type="personal">
<mods:role id="B9365B60923E8227AF0620161F9E2F25">
<mods:roleTerm id="C7B1C236EB4FF73741841EDC8BFC14F2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BED4A6EA490902B0406950D5C3727C03">Vermeersch, Xavier H. C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="49B50E9B84B6AAA11DC89F68D5891C2E">text</mods:typeOfResource>
<mods:relatedItem id="EDC8BC244B339530E8F58AE0671CA8CA" type="host">
<mods:titleInfo id="641AFE79736A0ED997EE9ADC70BAC8C7">
<mods:title id="AC8DD29997A92B7E296E6C8B4398283F">Belgian Journal of Entomology</mods:title>
</mods:titleInfo>
<mods:part id="3DA6B0AE66F9A10E51DB5D7732C80E6B">
<mods:date id="ACDBFC1BDEDD34FCB28835C76DCD814F">2001</mods:date>
<mods:detail id="0C61769E812DD7E05012E13170D52695" type="volume">
<mods:number id="04EC46CBA4EED4E119C33A04EE1BBF60">100</mods:number>
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<mods:start id="681BADE9854C8EF3CEE9C9141C9FDB4C">1</mods:start>
<mods:end id="B39E88C97D6A445E94D55DB4400C89A9">18</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="FBCE1259DCC8CBA09F185811D7328DF5">
<mods:url id="268936415F629680D5D1D40989B999E4">http://dx.doi.org/10.5281/zenodo.13272818</mods:url>
</mods:location>
<mods:classification id="85BAB3FC5F1BD4785260F34C660B3D87">journal article</mods:classification>
<mods:identifier id="1F3CDAD3C42D37653E74BCAF8F049E05" type="CLB-Dataset">301818</mods:identifier>
<mods:identifier id="F07CE4C85CE52A1330FB42382427F658" type="DOI">10.5281/zenodo.13272818</mods:identifier>
<mods:identifier id="0F675B0728DEE28F2A2B4F7E4916C815" type="GBIF-Dataset">25561bf2-ef51-4d75-a837-3789f9602786</mods:identifier>
<mods:identifier id="6384EE9DBF1954507D11A662405CA071" type="ISSN">2295-0214</mods:identifier>
</mods:mods>
<treatment id="30188791FFD92800FDEAFAD482DCE250" ID-DOI="http://doi.org/10.5281/zenodo.13449985" ID-Zenodo-Dep="13449985" LSID="urn:lsid:plazi:treatment:30188791FFD92800FDEAFAD482DCE250" httpUri="http://treatment.plazi.org/id/30188791FFD92800FDEAFAD482DCE250" lastPageId="11" pageId="5">
<subSubSection id="F0AB650CFFD9280EFDEAFAD48251E417" box="[603,1028,1361,1388]" pageId="5" type="nomenclature">
<paragraph id="B80E3687FFD9280EFDEAFAD48251E417" blockId="5.[445,1187,1361,1459]" box="[603,1028,1361,1388]" pageId="5">
<emphasis id="8AC5EA95FFD9280EFDEAFAD48251E417" bold="true" box="[603,1028,1361,1388]" pageId="5">
<taxonomicName id="7FB14D04FFD9280EFDEAFAD485C9E410" ID-CoL="8QQ4J" authorityName="Vermeersch" authorityYear="2001" box="[603,924,1361,1387]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="5" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFD9280EFDEAFAD485C9E410" bold="true" box="[603,924,1361,1387]" italics="true" pageId="5">Titanodula attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD9280EFC12FAD78251E417" box="[931,1028,1362,1388]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD9280EFE0CFAFD85DBE4C8" pageId="5" type="description">
<paragraph id="B80E3687FFD9280EFE0CFAFD82F6E4F5" blockId="5.[445,1187,1361,1459]" box="[445,1187,1400,1422]" pageId="5">
<uri id="CC203A85FFD9280EFE0CFAFD82F6E4F5" box="[445,1187,1400,1422]" pageId="5">
urn:lsid:zoobank.org:act:
<uuid id="CC170C52FFD9280EFD05FAFD82F6E4F5" box="[692,1187,1400,1422]" pageId="5">4CEFD21C-CB3F-4FB8-A35E-93B0966A5C63</uuid>
</uri>
</paragraph>
<paragraph id="B80E3687FFD9280EFD63FA1D85DBE4C8" blockId="5.[445,1187,1361,1459]" box="[722,910,1432,1459]" pageId="5">
(
<figureCitation id="208A2A02FFD9280EFD6BFA1D851DE4C9" box="[730,840,1432,1458]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[247,284,1261,1283]" captionStartId-1="8.[247,284,1459,1481]" captionStartId-2="10.[247,284,1431,1453]" captionTargetBox-0="[254,1376,536,1218]" captionTargetBox-1="[256,1378,887,1429]" captionTargetBox-2="[247,1385,273,1410]" captionTargetId-0="figure-400@6.[247,1385,529,1227]" captionTargetId-1="figure-458@8.[247,1385,870,1438]" captionTargetId-2="figure-209@10.[247,1385,273,1410]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-1="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-2="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="5">Figs 1−3</figureCitation>
,
<figureCitation id="208A2A02FFD9280EFCE4FA1D85D0E4C8" box="[853,901,1432,1459]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[247,284,1798,1820]" captionStartId-1="14.[247,284,1835,1857]" captionTargetBox-0="[354,1288,185,1775]" captionTargetBox-1="[298,1335,180,1812]" captionTargetId-0="figure-74@12.[330,1303,178,1778]" captionTargetId-1="figure-60@14.[296,1336,178,1814]" captionTargetPageId-0="12" captionText-0="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." captionText-1="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." pageId="5">5−6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD9280EFF46FA50850FE636" pageId="5" type="diagnosis">
<paragraph id="B80E3687FFD9280EFF46FA50850FE636" blockId="5.[247,1386,1492,1870]" pageId="5">
<smallCapsWord id="BEE8A05BFFD9280EFF46FA508729E496" baselines="1512,1512" box="[247,380,1493,1519]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Diagnosis" pageId="5">DIAGNOSIS</smallCapsWord>
. Very large and robust praying mantis. Head triangular, antennae filiform. Long but robust pronotum, with smooth dorsal surface. Pronotum very finely denticulate along the margins of the prozone in female, without denticulations in the metazone, entirely smooth edges in males. Spinal formula: F = 4DS/15AvS/4PvS; T = 13−14AvS/10PvS. Ratios
<specimenCount id="AEB7FD0EFFD9280EFB77F9BB82B5E723" box="[1222,1248,1598,1624]" count="1" pageId="5" type="male">♂</specimenCount>
: MzL/PzL: 3.2;
<specimenCount id="AEB7FD0EFFD9280EFE9DF9E48710E700" box="[300,325,1633,1659]" count="1" pageId="5" type="female">♀</specimenCount>
: MzL/PzL: 2.9. With black spots on the anterior side of the profemora located at the base of the 2
<superScript id="4FC49BCFFFD9280EFEEDF9FB8725E7EA" attach="none" box="[348,368,1662,1681]" fontSize="8" pageId="5">nd</superScript>
, 10
<superScript id="4FC49BCFFFD9280EFE13F9FB87E7E7EA" attach="left" box="[418,434,1662,1681]" fontSize="8" pageId="5">th</superScript>
and 15
<superScript id="4FC49BCFFFD9280EFDA5F9FB8476E7EA" attach="left" box="[532,547,1662,1681]" fontSize="8" pageId="5">th</superScript>
anteroventral spine. Protarsus (all segments) entirely black on the anterior side. Both sexes macropterous. Phalloid apophysis (afa) with two sclerotised processes, anterior process (aafa) small and tubercle-shaped, broad and dome-like at the base with a smaller rounded projection on top of it, located posteriorly from the middle. Posterior process (pafa) spear-shaped, with weakly developed base, almost straight, long and heavily sclerotised, projecting straight or slightly diagonally posteriad.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFDA280DFF46FF3E836BE081" pageId="6" type="materials_examined">
<paragraph id="B80E3687FFDA280DFF46FF3E833CE061" blockId="6.[247,1386,183,283]" pageId="6">
<materialsCitation id="08D93CDAFFDA280DFF46FF3E833CE061" collectingDate="1840-01" collectingDateMax="2018-07-20" collectingDateMin="1840-01" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel" country="Vietnam" latitude="14.474444" location="Gia Lai Province" longLatPrecision="21" longitude="108.54083" pageId="6" specimenCount="1" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="holotype">
<smallCapsWord id="BEE8A05BFFDA280DFF46FF3E8763E1A8" baselines="206,206" box="[247,310,187,213]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Type" pageId="6">
<collectionCode id="DEA0AE42FFDA280DFF46FF3E8763E1A8" box="[247,310,187,213]" pageId="6">TYPE</collectionCode>
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<smallCapsWord id="BEE8A05BFFDA280DFE8EFF3B87EFE1A8" baselines="206" box="[319,442,190,211]" lowerCaseFontSize="9" mainFontSize="11" normCase="lower" normString="material" pageId="6">
<collectionCode id="DEA0AE42FFDA280DFE8EFF3B87EFE1A8" box="[319,442,190,211]" pageId="6">MATERIAL</collectionCode>
</smallCapsWord>
.
<typeStatus id="670A8825FFDA280DFE7DFF3F8469E1AF" box="[460,572,186,212]" pageId="6" type="holotype">Holotype</typeStatus>
<emphasis id="8AC5EA95FFDA280DFDF9FF32840AE1AF" bold="true" box="[584,607,183,212]" pageId="6">
<specimenCount id="AEB7FD0EFFDA280DFDF9FF32840AE1AF" box="[584,607,183,212]" count="1" pageId="6" type="male">♂</specimenCount>
</emphasis>
(
<figureCitation id="208A2A02FFDA280DFDC0FF3F8494E1AF" box="[625,705,186,212]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Figs 1</figureCitation>
,
<figureCitation id="208A2A02FFDA280DFD62FF3F84B7E1AF" box="[723,738,186,212]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="6">3</figureCitation>
A−C):
<collectingCountry id="C0A67617FFDA280DFCF4FF3F85E5E1AF" box="[837,944,186,212]" name="Vietnam" pageId="6">Vietnam</collectingCountry>
,
<collectingRegion id="7A75F865FFDA280DFC0EFF3F82CEE1AF" box="[959,1179,186,212]" country="Vietnam" name="Gia Lai" pageId="6">Gia Lai Province</collectingRegion>
.,
<collectorName id="15445351FFDA280DFB1FFF3E8740E183" pageId="6">
Kon Chu Rang
<taxonomicNameLabel id="91F657EEFFDA280DFF46FF5B8740E183" box="[247,277,222,248]" pageId="6" rank="subSpecies">N.</taxonomicNameLabel>
</collectorName>
<collectionCode id="DEA0AE42FFDA280DFEA7FF5B877BE183" box="[278,302,222,248]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6">R</collectionCode>
.,
<geoCoordinate id="DD855040FFDA280DFEF4FF588789E18C" box="[325,476,221,247]" degrees="14" direction="north" minutes="28" orientation="latitude" pageId="6" precision="15" seconds="28" value="14.474444">14°28’28”N</geoCoordinate>
;
<geoCoordinate id="DD855040FFDA280DFE5EFF5884C4E18C" box="[495,657,221,247]" degrees="108" direction="east" minutes="32" orientation="longitude" pageId="6" precision="15" seconds="27" value="108.54083">108°32’27”E</geoCoordinate>
,
<date id="CC0F1047FFDA280DFD15FF58853CE18C" box="[676,873,221,247]" pageId="6" value="2018-07-13" valueMax="2018-07-20" valueMin="2018-07-13">
<collectingDate id="DC4BE9AFFFDA280DFD15FF58853CE18C" box="[676,873,221,247]" pageId="6" value="2018-07-13" valueMax="2018-07-20" valueMin="2018-07-13">13−20.VII.2018</collectingDate>
</date>
,
<collectorName id="15445351FFDA280DFCCDFF5885B9E18C" box="[892,1004,221,247]" pageId="6">At Light</collectorName>
, leg.
<collectorName id="15445351FFDA280DFB88FF5B829DE18C" box="[1081,1224,221,248]" pageId="6">J. Constant</collectorName>
, J.
<collectorName id="15445351FFDA280DFB4DFF588745E060" pageId="6">Bresseel, X</collectorName>
. Vermeersch,
<collectorName id="15445351FFDA280DFE0AFE85847CE061" box="[443,553,256,282]" pageId="6">Genitalia</collectorName>
prep.
<collectorName id="15445351FFDA280DFDC5FE858566E060" box="[628,819,256,283]" pageId="6">Vermeersch Nr.</collectorName>
<date id="CC0F1047FFDA280DFC88FE8585E3E061" box="[825,950,256,282]" pageId="6" value="1840-01">
<collectingDate id="DC4BE9AFFFDA280DFC88FE8585E3E061" box="[825,950,256,282]" pageId="6" value="1840-01">GEN1840</collectingDate>
</date>
, GTI
<collectorName id="15445351FFDA280DFC4BFE8582C4E061" box="[1018,1169,256,282]" pageId="6">Project, I.G.</collectorName>
: 33.769 (
<collectionCode id="DEA0AE42FFDA280DFB4CFE858309E061" box="[1277,1372,256,282]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="6" type="Museum">RBINS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="B80E3687FFDA280DFF46FEB5836BE081" blockId="6.[247,1385,304,506]" pageId="6">
<typeStatus id="670A8825FFDA280DFF46FEB58739E031" box="[247,364,304,330]" pageId="6" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="AEB7FD0EFFDA280DFE30FEB58794E031" box="[385,449,304,330]" count="3" pageId="6" type="male">3♂♂</specimenCount>
,
<specimenCount id="AEB7FD0EFFDA280DFE63FEB5845AE031" box="[466,527,304,330]" count="6" pageId="6" type="female">6♀♀</specimenCount>
):
<specimenCount id="AEB7FD0EFFDA280DFD9DFEB5843CE031" box="[556,617,304,330]" count="2" pageId="6" type="female">2♀♀</specimenCount>
(
<figureCitation id="208A2A02FFDA280DFDCFFEB58498E031" box="[638,717,304,330]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Fig. 2</figureCitation>
): same data as holotype, Night Collecting.
<materialsCitation id="08D93CDAFFDA280DFB45FEB585B9E0C8" collectingDate="2019-05-27" collectingDateMax="2019-05-28" collectingDateMin="2019-05-27" collectionCode="UniFI" collectorName="L. Bartolozzi &amp; S. Bambi &amp; A. Bandinelli" country="Vietnam" elevation="1110" latitude="14.520278" location="Gia Lai Province" longLatPrecision="21" longitude="108.47" pageId="6" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="paratype">
<typeStatus id="670A8825FFDA280DFB45FEB5833CE031" box="[1268,1385,304,330]" pageId="6" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="AEB7FD0EFFDA280DFEB0FED68717E016" box="[257,322,339,365]" count="2" pageId="6" type="male">2♂♂</specimenCount>
):
<collectingCountry id="C0A67617FFDA280DFEEFFED6879CE016" box="[350,457,339,365]" name="Vietnam" pageId="6">Vietnam</collectingCountry>
,
<collectingRegion id="7A75F865FFDA280DFE6AFED684EDE016" box="[475,696,339,365]" country="Vietnam" name="Gia Lai" pageId="6">Gia Lai Province</collectingRegion>
,
<collectorName id="15445351FFDA280DFD7BFED185E2E015" box="[714,951,339,366]" pageId="6">
Kon Chu Rang
<taxonomicNameLabel id="91F657EEFFDA280DFC28FED185E2E015" box="[921,951,340,366]" pageId="6" rank="subSpecies">N.</taxonomicNameLabel>
</collectorName>
<collectionCode id="DEA0AE42FFDA280DFC09FED18585E015" box="[952,976,340,366]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6">R</collectionCode>
.,
<quantity id="7F499B62FFDA280DFC56FED6826CE016" box="[999,1081,339,366]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="6" unit="km" value="15.0">15 km</quantity>
from HQ,
<elevation id="339CD1B4FFDA280DFB60FED68331E016" box="[1233,1380,339,366]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="6" unit="m" value="1110.0">
<quantity id="7F499B62FFDA280DFB60FED68367E015" box="[1233,1330,339,366]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="6" unit="m" value="1110.0">1110 m</quantity>
asl
</elevation>
,
<geoCoordinate id="DD855040FFDA280DFF46FEF387D8E0EB" box="[247,397,374,400]" degrees="14" direction="north" minutes="31" orientation="latitude" pageId="6" precision="15" seconds="13" value="14.520278">14°31’13”N</geoCoordinate>
;
<geoCoordinate id="DD855040FFDA280DFE1AFEF38419E0EB" box="[427,588,374,400]" degrees="108" direction="east" minutes="28" orientation="longitude" pageId="6" precision="15" seconds="12" value="108.47">108°28’12”E</geoCoordinate>
,
<date id="CC0F1047FFDA280DFDDBFEF3854EE0EB" box="[618,795,374,400]" pageId="6" value="2019-05-27" valueMax="2019-05-28" valueMin="2019-05-27">
<collectingDate id="DC4BE9AFFFDA280DFDDBFEF3854EE0EB" box="[618,795,374,400]" pageId="6" value="2019-05-27" valueMax="2019-05-28" valueMin="2019-05-27">27−28.V.2019</collectingDate>
</date>
,
<collectorName id="15445351FFDA280DFC89FEF385E4E0EB" box="[824,945,374,400]" pageId="6">At Light</collectorName>
, leg.
<collectorName id="15445351FFDA280DFBA2FEF28290E0EB" box="[1043,1221,374,401]" pageId="6">L. Bartolozzi</collectorName>
, S.
<collectorName id="15445351FFDA280DFAA0FEF38745E0C8" pageId="6">Bambi, A</collectorName>
.
<collectorName id="15445351FFDA280DFEADFE1C87EAE0CF" box="[284,447,409,436]" pageId="6">Bandinelli, E</collectorName>
. Orbach. Stored in ethanol (UniFI);
<specimenCount id="AEB7FD0EFFDA280DFCC9FE1C85CAE0C8" box="[888,927,409,435]" count="1" pageId="6" type="male">1♂</specimenCount>
,
<specimenCount id="AEB7FD0EFFDA280DFC1CFE1C85B9E0C8" box="[941,1004,409,435]" count="4" pageId="6" type="female">4♀♀</specimenCount>
</materialsCitation>
:
<materialsCitation id="08D93CDAFFDA280DFC4BFE1C836FE081" collectingDate="1910-01" collectingDateMax="2011-07-10" collectingDateMin="1910-01" collectionCode="R, RBINS" collectorName="J. Constant &amp; J. Bresseel" country="Vietnam" latitude="16.616667" location="Quang Tri Province" longLatPrecision="1281" longitude="106.78333" pageId="6" specimenCount="1" stateProvince="Quang Tri" typeStatus="paratype">
<collectingCountry id="C0A67617FFDA280DFC4BFE1C8230E0C8" box="[1018,1125,409,435]" name="Vietnam" pageId="6">Vietnam</collectingCountry>
,
<collectingRegion id="7A75F865FFDA280DFBC0FE1C8331E0C8" box="[1137,1380,409,435]" country="Vietnam" name="Quang Tri" pageId="6">Quang Tri Province</collectingRegion>
,
<collectorName id="15445351FFDA280DFF46FE3887FEE0AC" box="[247,427,445,471]" pageId="6">
Da Krong
<taxonomicNameLabel id="91F657EEFFDA280DFE3DFE3887FEE0AC" box="[396,427,445,471]" pageId="6" rank="subSpecies">N.</taxonomicNameLabel>
</collectorName>
<collectionCode id="DEA0AE42FFDA280DFE1AFE388796E0AC" box="[427,451,445,471]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6">R</collectionCode>
.,
<geoCoordinate id="DD855040FFDA280DFE6EFE39841EE0AD" box="[479,587,444,470]" degrees="16" direction="north" minutes="37" orientation="latitude" pageId="6" precision="925" value="16.616667">16°37’N</geoCoordinate>
;
<geoCoordinate id="DD855040FFDA280DFDD3FE39848CE0AD" box="[610,729,444,470]" degrees="106" direction="east" minutes="47" orientation="longitude" pageId="6" precision="925" value="106.78333">106°47’E</geoCoordinate>
,
<date id="CC0F1047FFDA280DFD41FE3985CAE0AC" box="[752,927,444,471]" pageId="6" value="2011-07-05" valueMax="2011-07-10" valueMin="2011-07-05">
<collectingDate id="DC4BE9AFFFDA280DFD41FE3985CAE0AC" box="[752,927,444,471]" pageId="6" value="2011-07-05" valueMax="2011-07-10" valueMin="2011-07-05">5-10.VII.2011</collectingDate>
</date>
,
<collectorName id="15445351FFDA280DFC07FE38822DE0AD" box="[950,1144,444,471]" pageId="6">Day Collecting</collectorName>
, leg.
<collectorName id="15445351FFDA280DFB60FE388330E0AC" box="[1233,1381,445,471]" pageId="6">J. Constant</collectorName>
, J. Bresseel,
<collectorName id="15445351FFDA280DFE36FE5A87A0E082" box="[391,501,479,505]" pageId="6">Genitalia</collectorName>
prep.
<collectorName id="15445351FFDA280DFDF0FE5A8557E081" box="[577,770,479,506]" pageId="6">Vermeersch Nr.</collectorName>
<date id="CC0F1047FFDA280DFCB8FE6585D3E081" box="[777,902,480,506]" pageId="6" value="1910-01">
<collectingDate id="DC4BE9AFFFDA280DFCB8FE6585D3E081" box="[777,902,480,506]" pageId="6" value="1910-01">GEN1910</collectingDate>
</date>
, GTI
<collectorName id="15445351FFDA280DFC7DFE5A8236E081" box="[972,1123,479,506]" pageId="6">Project, I.G.</collectorName>
: 31.933 (
<collectionCode id="DEA0AE42FFDA280DFB63FE658364E081" box="[1234,1329,480,506]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="6" type="Museum">RBINS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="ECCE660FFFDA280DFF46FB688254E45A" pageId="6" startId="6.[247,284,1261,1283]" targetBox="[254,1376,536,1218]" targetPageId="6" targetType="figure">
<paragraph id="B80E3687FFDA280DFF46FB688254E45A" blockId="6.[247,1386,1261,1313]" pageId="6">
Fig. 1.
<taxonomicName id="7FB14D04FFDA280DFE8FFB688415E479" authorityName="Vermeersch" authorityYear="2001" box="[318,576,1261,1282]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="6" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFDA280DFE8FFB688415E479" box="[318,576,1261,1282]" italics="true" pageId="6">Titanodula attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFDA280DFDF6FB6884CDE479" box="[583,664,1261,1282]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale.
</paragraph>
</caption>
<subSubSection id="F0AB650CFFDA2800FF46FAC6840FE0AD" lastPageId="11" pageId="6" type="description">
<paragraph id="B80E3687FFDA280DFF46FAC6858DE70D" blockId="6.[247,1386,1346,1654]" pageId="6">
<smallCapsWord id="BEE8A05BFFDA280DFF46FAC687C2E420" baselines="1366,1366" box="[247,407,1347,1373]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Description" pageId="6">DESCRIPTION</smallCapsWord>
.
<smallCapsWord id="BEE8A05BFFDA280DFE19FAC687A4E420" baselines="1366,1366" box="[424,497,1347,1373]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Male" pageId="6">MALE</smallCapsWord>
(
<figureCitation id="208A2A02FFDA280DFDB4FAC78401E427" box="[517,596,1346,1372]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Figs 1</figureCitation>
,
<figureCitation id="208A2A02FFDA280DFDD7FAC78420E427" box="[614,629,1346,1372]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="6">3</figureCitation>
A−C). Measurements (see
<tableCitation id="F533033CFFDA280DFC7DFAC7820DE427" box="[972,1112,1346,1372]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="7.[247,304,421,443]" captionStartId-1="9.[247,304,316,338]" captionText-0="Table 1. Detailed measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. From left to right: T. attenboroughi holotype ♂, T. attenboroughi paratype ♀, T. fruhstorferi male, additional specimen 1, T. fruhstorferi female, additional specimen 2." captionText-1="Table 2. Measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. to assess intraspecific variability." pageId="6">Tables 1−2</tableCitation>
). Colouration (
<figureCitation id="208A2A02FFDA280DFAA3FAC78337E427" box="[1298,1378,1346,1372]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Figs 1</figureCitation>
,
<figureCitation id="208A2A02FFDA280DFF46FAE38751E4FB" box="[247,260,1382,1408]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="6">3</figureCitation>
A−C): All known male specimens of
<taxonomicName id="7FB14D04FFDA280DFD53FAE385E5E404" authorityName="Vermeersch" authorityYear="2001" box="[738,944,1381,1408]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="6" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFDA280DFD53FAE384A6E404" box="[738,755,1382,1407]" italics="true" pageId="6">T</emphasis>
.
<emphasis id="8AC5EA95FFDA280DFCB7FAE085E5E404" box="[774,944,1381,1407]" italics="true" pageId="6">attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFDA280DFC0DFAE3824AE4FB" box="[956,1055,1382,1408]" pageId="6" rank="species">sp. nov.</taxonomicNameLabel>
feature a uniformly green colouration without any variations in darkness or tone between individuals. The dorsal side of pronotum is green and covered in both sexes with a whitish waxy secretion that is easily removed by touch. Underside of pronotum is entirely reddish-brown with a hint of magenta (purplish-red colour, sometimes also referred to as
<taxonomicName id="7FB14D04FFDA280DFCD4FA778593E777" box="[869,966,1522,1548]" class="Insecta" family="Oecophoridae" genus="Fuchsia" kingdom="Animalia" order="Lepidoptera" pageId="6" phylum="Arthropoda" rank="genus">
<emphasis id="8AC5EA95FFDA280DFCD4FA778593E777" box="[869,966,1522,1548]" italics="true" pageId="6">Fuchsia</emphasis>
</taxonomicName>
), but posterior end of pronotum is mostly magenta and is consistently covered with same whitish waxy secretion as on dorsal side. Ventral side of meso and metanotum are also magenta. Meso- and metacoxa are bright green but their trochanters are magenta while rest of legs are green.
</paragraph>
<paragraph id="B80E3687FFDA280DFF46F90E830BE62E" blockId="6.[247,1386,1675,1877]" pageId="6">
<emphasis id="8AC5EA95FFDA280DFF46F90E876DE7DE" box="[247,312,1675,1701]" italics="true" pageId="6">Head</emphasis>
(
<figureCitation id="208A2A02FFDA280DFEFAF90E87E9E7DE" box="[331,444,1675,1701]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Figs 1 C</figureCitation>
;
<figureCitation id="208A2A02FFDA280DFE7AF90E878FE7DE" box="[459,474,1675,1701]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="6">3</figureCitation>
A−C): Wider than long with large, rounded compound eyes. Vertex flat. Ocellar tubercle not visible, in same plane as frons, without protruding ridges. Ocelli small, all about same size. Lower frons transverse with external margins posteriorly and laterally, no margin anteriorly, with two faintly defined vertical ridges internally that run from posterior end towards middle where they are abruptly interrupted. Two more clearly defined tubercules are placed in their extension and border anterior part of lower frons. Clypeus and labrum smooth.
</paragraph>
<paragraph id="B80E3687FFDB280CFF46FF3F8315E0FF" blockId="7.[247,1386,186,388]" pageId="7">
<emphasis id="8AC5EA95FFDB280CFF46FF3F871BE1AF" box="[247,334,186,212]" italics="true" pageId="7">Thorax</emphasis>
(
<figureCitation id="208A2A02FFDB280CFED1FF3F87FAE1AF" box="[352,431,186,212]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="7">Figs 1</figureCitation>
A−B; 3 A−C): Pronotum long, straight and robust. Margins smooth, without denticulations or projections. Dorsal surface also entirely smooth, with a darker spot at each side laterally adjacent to the posterior margin of pronotum. Largest width before one third of pronotum, narrowest width located after middle of pronotum. Cervix with strongly sclerotized lateral and intercervical sclerites, merged in the middle. Postcervical plate and posterior ventral part of pronotum entirely smooth. Ventral lateral margins concolour with rest of pronotum.
</paragraph>
<caption id="ECCE660FFFDB280CFF46FE208568E08C" pageId="7" startId="7.[247,304,421,443]" targetType="table">
<paragraph id="B80E3687FFDB280CFF46FE208568E08C" blockId="7.[247,1386,421,504]" pageId="7">
Table 1. Detailed measurements [mm] of
<taxonomicName id="7FB14D04FFDB280CFD17FE2085FFE0C1" authorityName="Vermeersch" authorityYear="2001" box="[678,938,421,442]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFDB280CFD17FE2085FFE0C1" box="[678,938,421,442]" italics="true" pageId="7">Titanodula attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFDB280CFC02FE238250E0C0" box="[947,1029,422,443]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="7FB14D04FFDB280CFB88FE208376E0C1" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[1081,1315,421,442]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFDB280CFB88FE208376E0C1" box="[1081,1315,421,442]" italics="true" pageId="7">Titanodula fruhstorferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFDB280CFA9AFE208771E0A2" pageId="7" rank="species">comb. nov.</taxonomicNameLabel>
From left to right:
<taxonomicName id="7FB14D04FFDB280CFE53FE4184DFE0A3" authorityName="Vermeersch" authorityYear="2001" box="[482,650,451,473]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="attenboroughi">
<emphasis id="8AC5EA95FFDB280CFE53FE4184DFE0A3" box="[482,650,451,473]" italics="true" pageId="7">T. attenboroughi</emphasis>
</taxonomicName>
holotype ♂,
<taxonomicName id="7FB14D04FFDB280CFCBDFE4185E1E0A3" authorityName="Vermeersch" authorityYear="2001" box="[780,948,451,473]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="attenboroughi">
<emphasis id="8AC5EA95FFDB280CFCBDFE4185E1E0A3" box="[780,948,451,473]" italics="true" pageId="7">T. attenboroughi</emphasis>
</taxonomicName>
paratype ♀,
<taxonomicName id="7FB14D04FFDB280CFB84FE418296E0A3" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[1077,1219,451,473]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="fruhstorferi">
<emphasis id="8AC5EA95FFDB280CFB84FE418296E0A3" box="[1077,1219,451,473]" italics="true" pageId="7">T. fruhstorferi</emphasis>
</taxonomicName>
male, additional specimen 1,
<taxonomicName id="7FB14D04FFDB280CFEC4FE678451E08C" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[373,516,482,503]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="fruhstorferi">
<emphasis id="8AC5EA95FFDB280CFEC4FE678451E08C" box="[373,516,482,503]" italics="true" pageId="7">T. fruhstorferi</emphasis>
</taxonomicName>
female, additional specimen 2.
</paragraph>
</caption>
<paragraph id="B80E3687FFDB280CFEFDFD988357E707" pageId="7">
<table id="CAB1C427FFDBD7F4FEFDFD888375E706" box="[332,1312,525,1661]" gridcols="5" gridrows="25" pageId="7">
<tr id="068134C5FFDBD7F4FEFDFD888375E33A" box="[332,1312,525,577]" gridrow="0" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFD8884C5E33A" box="[332,656,525,577]" gridcol="0" gridrow="0" pageId="7">
<emphasis id="8AC5EA95FFDB280CFEFDFD9887BCE349" bold="true" box="[332,489,541,562]" pageId="7">Measurements</emphasis>
</th>
<th id="45505DB9FFDBD7F4FD53FD888564E33A" box="[738,817,525,577]" gridcol="1" gridrow="0" pageId="7">
<emphasis id="8AC5EA95FFDB280CFD52FD8B8564E33A" bold="true" pageId="7">
<taxonomicName id="7FB14D04FFDB280CFD52FD8B8552E33A" authority="HT" authorityName="HT" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="att">
<emphasis id="8AC5EA95FFDB280CFD52FD8B854DE358" bold="true" box="[739,792,526,547]" italics="true" pageId="7">T. att</emphasis>
HT
</taxonomicName>
(♂)
</emphasis>
</th>
<th id="45505DB9FFDBD7F4FC31FD88859EE33A" box="[896,971,525,577]" gridcol="2" gridrow="0" pageId="7">
<emphasis id="8AC5EA95FFDB280CFC30FD8B859EE33A" bold="true" pageId="7">
<taxonomicName id="7FB14D04FFDB280CFC30FD8B85F4E33A" authority="PT" authorityName="PT" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="att">
<emphasis id="8AC5EA95FFDB280CFC30FD8B85E3E358" bold="true" box="[897,950,526,547]" italics="true" pageId="7">T. att</emphasis>
PT
</taxonomicName>
(♀)
</emphasis>
</th>
<th id="45505DB9FFDBD7F4FBADFD888226E33A" box="[1052,1139,525,577]" gridcol="3" gridrow="0" pageId="7">
<emphasis id="8AC5EA95FFDB280CFBADFD8B8226E33A" bold="true" pageId="7">
<taxonomicName id="7FB14D04FFDB280CFBADFD8B826BE33A" authority="AS" authorityName="AS" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="fru">
<emphasis id="8AC5EA95FFDB280CFBADFD8B820DE358" bold="true" box="[1052,1112,525,547]" italics="true" pageId="7">T. fru</emphasis>
AS
</taxonomicName>
1 (♂)
</emphasis>
</th>
<th id="45505DB9FFDBD7F4FB78FD888375E33A" box="[1225,1312,525,577]" gridcol="4" gridrow="0" pageId="7">
<emphasis id="8AC5EA95FFDB280CFB78FD8B8375E33A" bold="true" pageId="7">
<taxonomicName id="7FB14D04FFDB280CFB78FD8B82BCE33A" authority="AS" authorityName="AS" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="fru">
<emphasis id="8AC5EA95FFDB280CFB78FD8B8350E358" bold="true" box="[1225,1285,525,547]" italics="true" pageId="7">T. fru</emphasis>
AS
</taxonomicName>
2 (♀)
</emphasis>
</th>
</tr>
<tr id="068134C5FFDBD7F4FEFDFDD28375E316" box="[332,1312,599,621]" gridrow="1" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFDD284C5E316" box="[332,656,599,621]" gridcol="0" gridrow="1" pageId="7">Total length</th>
<td id="45505DB9FFDBD7F4FD53FDD28564E316" box="[738,817,599,621]" gridcol="1" gridrow="1" pageId="7">99.6</td>
<td id="45505DB9FFDBD7F4FC31FDD2859EE316" box="[896,971,599,621]" gridcol="2" gridrow="1" pageId="7">102.2</td>
<td id="45505DB9FFDBD7F4FBADFDD28226E316" box="[1052,1139,599,621]" gridcol="3" gridrow="1" pageId="7">107</td>
<td id="45505DB9FFDBD7F4FB78FDD28375E316" box="[1225,1312,599,621]" gridcol="4" gridrow="1" pageId="7">93.7</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFD068375E3E2" box="[332,1312,643,665]" gridrow="2" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFD0684C5E3E2" box="[332,656,643,665]" gridcol="0" gridrow="2" pageId="7">Body length</th>
<td id="45505DB9FFDBD7F4FD53FD068564E3E2" box="[738,817,643,665]" gridcol="1" gridrow="2" pageId="7">-</td>
<td id="45505DB9FFDBD7F4FC31FD06859EE3E2" box="[896,971,643,665]" gridcol="2" gridrow="2" pageId="7">97.3</td>
<td id="45505DB9FFDBD7F4FBADFD068226E3E2" box="[1052,1139,643,665]" gridcol="3" gridrow="2" pageId="7">88.4</td>
<td id="45505DB9FFDBD7F4FB78FD068375E3E2" box="[1225,1312,643,665]" gridcol="4" gridrow="2" pageId="7">91.9</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFD2A8375E3BE" box="[332,1312,687,709]" gridrow="3" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFD2A84C5E3BE" box="[332,656,687,709]" gridcol="0" gridrow="3" pageId="7">Head width</th>
<td id="45505DB9FFDBD7F4FD53FD2A8564E3BE" box="[738,817,687,709]" gridcol="1" gridrow="3" pageId="7">10.3</td>
<td id="45505DB9FFDBD7F4FC31FD2A859EE3BE" box="[896,971,687,709]" gridcol="2" gridrow="3" pageId="7">12.5</td>
<td id="45505DB9FFDBD7F4FBADFD2A8226E3BE" box="[1052,1139,687,709]" gridcol="3" gridrow="3" pageId="7">10.5</td>
<td id="45505DB9FFDBD7F4FB78FD2A8375E3BE" box="[1225,1312,687,709]" gridcol="4" gridrow="3" pageId="7">11.9</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFD5E8375E38A" box="[332,1312,731,753]" gridrow="4" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFD5E84C5E38A" box="[332,656,731,753]" gridcol="0" gridrow="4" pageId="7">Head height</th>
<td id="45505DB9FFDBD7F4FD53FD5E8564E38A" box="[738,817,731,753]" gridcol="1" gridrow="4" pageId="7">8.6</td>
<td id="45505DB9FFDBD7F4FC31FD5E859EE38A" box="[896,971,731,753]" gridcol="2" gridrow="4" pageId="7">11.2</td>
<td id="45505DB9FFDBD7F4FBADFD5E8226E38A" box="[1052,1139,731,753]" gridcol="3" gridrow="4" pageId="7">8.2</td>
<td id="45505DB9FFDBD7F4FB78FD5E8375E38A" box="[1225,1312,731,753]" gridcol="4" gridrow="4" pageId="7">10.6</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFC828375E266" box="[332,1312,775,797]" gridrow="5" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFC8284C5E266" box="[332,656,775,797]" gridcol="0" gridrow="5" pageId="7">Pronotum length</th>
<td id="45505DB9FFDBD7F4FD53FC828564E266" box="[738,817,775,797]" gridcol="1" gridrow="5" pageId="7">29.3</td>
<td id="45505DB9FFDBD7F4FC31FC82859EE266" box="[896,971,775,797]" gridcol="2" gridrow="5" pageId="7">35</td>
<td id="45505DB9FFDBD7F4FBADFC828226E266" box="[1052,1139,775,797]" gridcol="3" gridrow="5" pageId="7">30.2</td>
<td id="45505DB9FFDBD7F4FB78FC828375E266" box="[1225,1312,775,797]" gridcol="4" gridrow="5" pageId="7">33.2</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFCB68375E232" box="[332,1312,819,841]" gridrow="6" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFCB684C5E232" box="[332,656,819,841]" gridcol="0" gridrow="6" pageId="7">Pronotum width</th>
<td id="45505DB9FFDBD7F4FD53FCB68564E232" box="[738,817,819,841]" gridcol="1" gridrow="6" pageId="7">8.2</td>
<td id="45505DB9FFDBD7F4FC31FCB6859EE232" box="[896,971,819,841]" gridcol="2" gridrow="6" pageId="7">10.4</td>
<td id="45505DB9FFDBD7F4FBADFCB68226E232" box="[1052,1139,819,841]" gridcol="3" gridrow="6" pageId="7">8.9</td>
<td id="45505DB9FFDBD7F4FB78FCB68375E232" box="[1225,1312,819,841]" gridcol="4" gridrow="6" pageId="7">10.2</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFCDA8375E20E" box="[332,1312,863,885]" gridrow="7" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFCDA84C5E20E" box="[332,656,863,885]" gridcol="0" gridrow="7" pageId="7">Pronotum narrow width</th>
<td id="45505DB9FFDBD7F4FD53FCDA8564E20E" box="[738,817,863,885]" gridcol="1" gridrow="7" pageId="7">4.7</td>
<td id="45505DB9FFDBD7F4FC31FCDA859EE20E" box="[896,971,863,885]" gridcol="2" gridrow="7" pageId="7">6</td>
<td id="45505DB9FFDBD7F4FBADFCDA8226E20E" box="[1052,1139,863,885]" gridcol="3" gridrow="7" pageId="7">4.9</td>
<td id="45505DB9FFDBD7F4FB78FCDA8375E20E" box="[1225,1312,863,885]" gridcol="4" gridrow="7" pageId="7">5</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFC0E8375E2DA" box="[332,1312,907,929]" gridrow="8" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFC0E84C5E2DA" box="[332,656,907,929]" gridcol="0" gridrow="8" pageId="7">Prozone length</th>
<td id="45505DB9FFDBD7F4FD53FC0E8564E2DA" box="[738,817,907,929]" gridcol="1" gridrow="8" pageId="7">7</td>
<td id="45505DB9FFDBD7F4FC31FC0E859EE2DA" box="[896,971,907,929]" gridcol="2" gridrow="8" pageId="7">8.9</td>
<td id="45505DB9FFDBD7F4FBADFC0E8226E2DA" box="[1052,1139,907,929]" gridcol="3" gridrow="8" pageId="7">7.3</td>
<td id="45505DB9FFDBD7F4FB78FC0E8375E2DA" box="[1225,1312,907,929]" gridcol="4" gridrow="8" pageId="7">8.3</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFC328375E2B6" box="[332,1312,951,973]" gridrow="9" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFC3284C5E2B6" box="[332,656,951,973]" gridcol="0" gridrow="9" pageId="7">Metazone length</th>
<td id="45505DB9FFDBD7F4FD53FC328564E2B6" box="[738,817,951,973]" gridcol="1" gridrow="9" pageId="7">22.3</td>
<td id="45505DB9FFDBD7F4FC31FC32859EE2B6" box="[896,971,951,973]" gridcol="2" gridrow="9" pageId="7">26.1</td>
<td id="45505DB9FFDBD7F4FBADFC328226E2B6" box="[1052,1139,951,973]" gridcol="3" gridrow="9" pageId="7">22.9</td>
<td id="45505DB9FFDBD7F4FB78FC328375E2B6" box="[1225,1312,951,973]" gridcol="4" gridrow="9" pageId="7">24.9</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFC668375E282" box="[332,1312,995,1017]" gridrow="10" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFC6684C5E282" box="[332,656,995,1017]" gridcol="0" gridrow="10" pageId="7">Tegmen length</th>
<td id="45505DB9FFDBD7F4FD53FC668564E282" box="[738,817,995,1017]" gridcol="1" gridrow="10" pageId="7">69</td>
<td id="45505DB9FFDBD7F4FC31FC66859EE282" box="[896,971,995,1017]" gridcol="2" gridrow="10" pageId="7">61.2</td>
<td id="45505DB9FFDBD7F4FBADFC668226E282" box="[1052,1139,995,1017]" gridcol="3" gridrow="10" pageId="7">73.8</td>
<td id="45505DB9FFDBD7F4FB78FC668375E282" box="[1225,1312,995,1017]" gridcol="4" gridrow="10" pageId="7">59.2</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFB8B8375E55E" box="[332,1312,1038,1061]" gridrow="11" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFB8B84C5E55E" box="[332,656,1038,1061]" gridcol="0" gridrow="11" pageId="7">Ala lenght</th>
<td id="45505DB9FFDBD7F4FD53FB8B8564E55E" box="[738,817,1038,1061]" gridcol="1" gridrow="11" pageId="7">62.2</td>
<td id="45505DB9FFDBD7F4FC31FB8B859EE55E" box="[896,971,1038,1061]" gridcol="2" gridrow="11" pageId="7">55.5</td>
<td id="45505DB9FFDBD7F4FBADFB8B8226E55E" box="[1052,1139,1038,1061]" gridcol="3" gridrow="11" pageId="7">-</td>
<td id="45505DB9FFDBD7F4FB78FB8B8375E55E" box="[1225,1312,1038,1061]" gridcol="4" gridrow="11" pageId="7">-</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFBBF8375E52B" box="[332,1312,1082,1104]" gridrow="12" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFBBF84C5E52B" box="[332,656,1082,1104]" gridcol="0" gridrow="12" pageId="7">Procoxa length</th>
<td id="45505DB9FFDBD7F4FD53FBBF8564E52B" box="[738,817,1082,1104]" gridcol="1" gridrow="12" pageId="7">16.8</td>
<td id="45505DB9FFDBD7F4FC31FBBF859EE52B" box="[896,971,1082,1104]" gridcol="2" gridrow="12" pageId="7">22.5</td>
<td id="45505DB9FFDBD7F4FBADFBBF8226E52B" box="[1052,1139,1082,1104]" gridcol="3" gridrow="12" pageId="7">18.5</td>
<td id="45505DB9FFDBD7F4FB78FBBF8375E52B" box="[1225,1312,1082,1104]" gridcol="4" gridrow="12" pageId="7">21.4</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFBE38375E507" box="[332,1312,1126,1148]" gridrow="13" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFBE384C5E507" box="[332,656,1126,1148]" gridcol="0" gridrow="13" pageId="7">Profemur length</th>
<td id="45505DB9FFDBD7F4FD53FBE38564E507" box="[738,817,1126,1148]" gridcol="1" gridrow="13" pageId="7">20.4</td>
<td id="45505DB9FFDBD7F4FC31FBE3859EE507" box="[896,971,1126,1148]" gridcol="2" gridrow="13" pageId="7">25.6</td>
<td id="45505DB9FFDBD7F4FBADFBE38226E507" box="[1052,1139,1126,1148]" gridcol="3" gridrow="13" pageId="7">21.2</td>
<td id="45505DB9FFDBD7F4FB78FBE38375E507" box="[1225,1312,1126,1148]" gridcol="4" gridrow="13" pageId="7">23.1</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFB178375E5D3" box="[332,1312,1170,1192]" gridrow="14" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFB1784C5E5D3" box="[332,656,1170,1192]" gridcol="0" gridrow="14" pageId="7">Protibia length</th>
<td id="45505DB9FFDBD7F4FD53FB178564E5D3" box="[738,817,1170,1192]" gridcol="1" gridrow="14" pageId="7">12.9</td>
<td id="45505DB9FFDBD7F4FC31FB17859EE5D3" box="[896,971,1170,1192]" gridcol="2" gridrow="14" pageId="7">17.3</td>
<td id="45505DB9FFDBD7F4FBADFB178226E5D3" box="[1052,1139,1170,1192]" gridcol="3" gridrow="14" pageId="7">13.8</td>
<td id="45505DB9FFDBD7F4FB78FB178375E5D3" box="[1225,1312,1170,1192]" gridcol="4" gridrow="14" pageId="7">16.1</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFB3B8375E5AF" box="[332,1312,1214,1236]" gridrow="15" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFB3B84C5E5AF" box="[332,656,1214,1236]" gridcol="0" gridrow="15" pageId="7">Protarsus length</th>
<td id="45505DB9FFDBD7F4FD53FB3B8564E5AF" box="[738,817,1214,1236]" gridcol="1" gridrow="15" pageId="7">14</td>
<td id="45505DB9FFDBD7F4FC31FB3B859EE5AF" box="[896,971,1214,1236]" gridcol="2" gridrow="15" pageId="7">13.3</td>
<td id="45505DB9FFDBD7F4FBADFB3B8226E5AF" box="[1052,1139,1214,1236]" gridcol="3" gridrow="15" pageId="7">13.6</td>
<td id="45505DB9FFDBD7F4FB78FB3B8375E5AF" box="[1225,1312,1214,1236]" gridcol="4" gridrow="15" pageId="7">15.1</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFB6F8375E47B" box="[332,1312,1258,1280]" gridrow="16" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFB6F84C5E47B" box="[332,656,1258,1280]" gridcol="0" gridrow="16" pageId="7">Mesofemur length</th>
<td id="45505DB9FFDBD7F4FD53FB6F8564E47B" box="[738,817,1258,1280]" gridcol="1" gridrow="16" pageId="7">19.2</td>
<td id="45505DB9FFDBD7F4FC31FB6F859EE47B" box="[896,971,1258,1280]" gridcol="2" gridrow="16" pageId="7">24.2</td>
<td id="45505DB9FFDBD7F4FBADFB6F8226E47B" box="[1052,1139,1258,1280]" gridcol="3" gridrow="16" pageId="7">21.2</td>
<td id="45505DB9FFDBD7F4FB78FB6F8375E47B" box="[1225,1312,1258,1280]" gridcol="4" gridrow="16" pageId="7">21.9</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFA938375E457" box="[332,1312,1302,1324]" gridrow="17" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFA9384C5E457" box="[332,656,1302,1324]" gridcol="0" gridrow="17" pageId="7">Mesotibia length</th>
<td id="45505DB9FFDBD7F4FD53FA938564E457" box="[738,817,1302,1324]" gridcol="1" gridrow="17" pageId="7">16</td>
<td id="45505DB9FFDBD7F4FC31FA93859EE457" box="[896,971,1302,1324]" gridcol="2" gridrow="17" pageId="7">20.6</td>
<td id="45505DB9FFDBD7F4FBADFA938226E457" box="[1052,1139,1302,1324]" gridcol="3" gridrow="17" pageId="7">17.3</td>
<td id="45505DB9FFDBD7F4FB78FA938375E457" box="[1225,1312,1302,1324]" gridcol="4" gridrow="17" pageId="7">17.8</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFAC78375E423" box="[332,1312,1346,1368]" gridrow="18" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFAC784C5E423" box="[332,656,1346,1368]" gridcol="0" gridrow="18" pageId="7">Mesotarsus length</th>
<td id="45505DB9FFDBD7F4FD53FAC78564E423" box="[738,817,1346,1368]" gridcol="1" gridrow="18" pageId="7">10.7</td>
<td id="45505DB9FFDBD7F4FC31FAC7859EE423" box="[896,971,1346,1368]" gridcol="2" gridrow="18" pageId="7">10.8</td>
<td id="45505DB9FFDBD7F4FBADFAC78226E423" box="[1052,1139,1346,1368]" gridcol="3" gridrow="18" pageId="7">11.4</td>
<td id="45505DB9FFDBD7F4FB78FAC78375E423" box="[1225,1312,1346,1368]" gridcol="4" gridrow="18" pageId="7">11.3</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFAEB8375E4FF" box="[332,1312,1390,1412]" gridrow="19" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFAEB84C5E4FF" box="[332,656,1390,1412]" gridcol="0" gridrow="19" pageId="7">Metafemur length</th>
<td id="45505DB9FFDBD7F4FD53FAEB8564E4FF" box="[738,817,1390,1412]" gridcol="1" gridrow="19" pageId="7">22.3</td>
<td id="45505DB9FFDBD7F4FC31FAEB859EE4FF" box="[896,971,1390,1412]" gridcol="2" gridrow="19" pageId="7">28</td>
<td id="45505DB9FFDBD7F4FBADFAEB8226E4FF" box="[1052,1139,1390,1412]" gridcol="3" gridrow="19" pageId="7">25.8</td>
<td id="45505DB9FFDBD7F4FB78FAEB8375E4FF" box="[1225,1312,1390,1412]" gridcol="4" gridrow="19" pageId="7">25.2</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFA1F8375E4CB" box="[332,1312,1434,1456]" gridrow="20" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFA1F84C5E4CB" box="[332,656,1434,1456]" gridcol="0" gridrow="20" pageId="7">Metatibia length</th>
<td id="45505DB9FFDBD7F4FD53FA1F8564E4CB" box="[738,817,1434,1456]" gridcol="1" gridrow="20" pageId="7">21.8</td>
<td id="45505DB9FFDBD7F4FC31FA1F859EE4CB" box="[896,971,1434,1456]" gridcol="2" gridrow="20" pageId="7">28.6</td>
<td id="45505DB9FFDBD7F4FBADFA1F8226E4CB" box="[1052,1139,1434,1456]" gridcol="3" gridrow="20" pageId="7">24.4</td>
<td id="45505DB9FFDBD7F4FB78FA1F8375E4CB" box="[1225,1312,1434,1456]" gridcol="4" gridrow="20" pageId="7">25.5</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFA438375E4A7" box="[332,1312,1478,1500]" gridrow="21" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFA4384C5E4A7" box="[332,656,1478,1500]" gridcol="0" gridrow="21" pageId="7">Metatarsus length</th>
<td id="45505DB9FFDBD7F4FD53FA438564E4A7" box="[738,817,1478,1500]" gridcol="1" gridrow="21" pageId="7">12.9</td>
<td id="45505DB9FFDBD7F4FC31FA43859EE4A7" box="[896,971,1478,1500]" gridcol="2" gridrow="21" pageId="7">14.6</td>
<td id="45505DB9FFDBD7F4FBADFA438226E4A7" box="[1052,1139,1478,1500]" gridcol="3" gridrow="21" pageId="7">14.6</td>
<td id="45505DB9FFDBD7F4FB78FA438375E4A7" box="[1225,1312,1478,1500]" gridcol="4" gridrow="21" pageId="7">13.9</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDFA778375E75E" box="[332,1312,1522,1573]" gridrow="22" pageId="7">
<th id="45505DB9FFDBD7F4FEFDFA7784C5E75E" box="[332,656,1522,1573]" gridcol="0" gridrow="22" pageId="7">Anteroventral femoral spine count</th>
<td id="45505DB9FFDBD7F4FD53FA778564E75E" box="[738,817,1522,1573]" gridcol="1" gridrow="22" pageId="7">15/15</td>
<td id="45505DB9FFDBD7F4FC31FA77859EE75E" box="[896,971,1522,1573]" gridcol="2" gridrow="22" pageId="7">15/15</td>
<td id="45505DB9FFDBD7F4FBADFA778226E75E" box="[1052,1139,1522,1573]" gridcol="3" gridrow="22" pageId="7">15/15</td>
<td id="45505DB9FFDBD7F4FB78FA778375E75E" box="[1225,1312,1522,1573]" gridcol="4" gridrow="22" pageId="7">15/15</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDF9BE8375E72A" box="[332,1312,1595,1617]" gridrow="23" pageId="7">
<th id="45505DB9FFDBD7F4FEFDF9BE84C5E72A" box="[332,656,1595,1617]" gridcol="0" gridrow="23" pageId="7">Anteroventral tibial spine count</th>
<td id="45505DB9FFDBD7F4FD53F9BE8564E72A" box="[738,817,1595,1617]" gridcol="1" gridrow="23" pageId="7">13/13</td>
<td id="45505DB9FFDBD7F4FC31F9BE859EE72A" box="[896,971,1595,1617]" gridcol="2" gridrow="23" pageId="7">14/14</td>
<td id="45505DB9FFDBD7F4FBADF9BE8226E72A" box="[1052,1139,1595,1617]" gridcol="3" gridrow="23" pageId="7">14/14</td>
<td id="45505DB9FFDBD7F4FB78F9BE8375E72A" box="[1225,1312,1595,1617]" gridcol="4" gridrow="23" pageId="7">14/14</td>
</tr>
<tr id="068134C5FFDBD7F4FEFDF9E28375E706" box="[332,1312,1639,1661]" gridrow="24" pageId="7">
<th id="45505DB9FFDBD7F4FEFDF9E284C5E706" box="[332,656,1639,1661]" gridcol="0" gridrow="24" pageId="7">Posteroventral tibial spine count</th>
<td id="45505DB9FFDBD7F4FD53F9E28564E706" box="[738,817,1639,1661]" gridcol="1" gridrow="24" pageId="7">10/9</td>
<td id="45505DB9FFDBD7F4FC31F9E2859EE706" box="[896,971,1639,1661]" gridcol="2" gridrow="24" pageId="7">10/10</td>
<td id="45505DB9FFDBD7F4FBADF9E28226E706" box="[1052,1139,1639,1661]" gridcol="3" gridrow="24" pageId="7">10/10</td>
<td id="45505DB9FFDBD7F4FB78F9E28375E706" box="[1225,1312,1639,1661]" gridcol="4" gridrow="24" pageId="7">10/10</td>
</tr>
</table>
</paragraph>
<paragraph id="B80E3687FFDB2803FF46F946845EE237" blockId="7.[247,1385,1731,1934]" lastBlockId="8.[247,1386,180,845]" lastPageId="8" pageId="7">
<emphasis id="8AC5EA95FFDB280CFF46F9468793E7A6" box="[247,454,1731,1757]" italics="true" pageId="7">Prothoracic legs</emphasis>
(
<figureCitation id="208A2A02FFDB280CFE69F9468472E7A5" box="[472,551,1731,1758]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="7">Figs 1</figureCitation>
A−B, D; 3 A−C): Coxa with 8−10 very small more or less blunt spinules of more or less equal size with same colour as coxa. Dorsal and ventral coxal lobes rounded, equal in length and adjacent, both lobes equally as broad. Black spot at proximal edge of ventral coxal lobe. Trochanter with black spot at anterior tip, adjacent matching black spot of same size on anterior side of femur. Femur with gently convex but straight dorsal margin, surface entirely smooth. Femoral brush ellipse-shaped, starting from 12
<superScript id="4FC49BCFFFDB280CFBCFF8E882DBE6FB" attach="left" box="[1150,1166,1901,1920]" fontSize="8" pageId="7">th</superScript>
or 13
<superScript id="4FC49BCFFFDB280CFB6EF8E882BAE6FB" attach="left" box="[1247,1263,1901,1920]" fontSize="8" pageId="7">th</superScript>
AvS and ending just a little distally of 15
<superScript id="4FC49BCFFFD42803FDC5FF3184D6E1BC" attach="left" box="[628,643,180,199]" fontSize="8" pageId="8">th</superScript>
AvS. Genicular lobes large and rounded, with minute, almost invisible genicular spur. Colouration of prothoracic legs green posteriorly, yellow anteriorly. Tibial spine groove proximally placed from middle of femur. AvS 1, 2, 10 and 15 black, black only along the distal and proximal edges or strongly apically infuscate (variable colouration between specimens) with black spot at base of AvS 2, 10 and 15 that slightly extends on anterior surface of femur; other AvS concolour with femur, only apically infuscate; AvS 10 and 12 large, but with 2 much smaller spines of equal size between them. Large and medium spines alternate between AvS 1 and 12, first spine being a medium one; medium spines subtly inclined towards anterior side of femur. PvS 4 smaller than others, PvS 1, 2 and 3 about equal in size; all concolour with femur and slightly apically infuscate. DS 1 and 4 more or less equal in size, DS 2 slightly longer than DS1, DS 3 longest and about 2x size of DS 2; DS 1 and 3 entirely black on anterior side (note: in some specimens only DS1 entirely black and DS3 apically infuscate), apically infuscate on posterior side, others apically infuscate on both sides. DS1 with black spot at base on anterior side. Protibiae smooth with longitudinal groove across entire length of posterior side. Tibial AvS gradually elongated towards tibial spur, all concolour to tibia but apically infuscate. PvS slightly more procumbent, spines gradually longer towards distal end. Tibial spur apically infuscate. Protarsi apically darkened posteriorly, entirely black anteriorly; first tarsomere longer than others combined. Spinal formula: F = 4DS/15AvS/4PvS; T = 13−14AvS/10PvS.
</paragraph>
<caption id="ECCE660FFFD42803FF46FA36826AE49C" pageId="8" startId="8.[247,284,1459,1481]" targetBox="[256,1378,887,1429]" targetPageId="8" targetType="figure">
<paragraph id="B80E3687FFD42803FF46FA36826AE49C" blockId="8.[247,1385,1459,1511]" pageId="8">
Fig. 2.
<taxonomicName id="7FB14D04FFD42803FE8CFA36846AE4B3" authorityName="Vermeersch" authorityYear="2001" box="[317,575,1459,1480]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="8" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFD42803FE8CFA36846AE4B3" box="[317,575,1459,1480]" italics="true" pageId="8">Titanodula attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD42803FDF4FA3684C3E4B3" box="[581,662,1459,1480]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale.
</paragraph>
</caption>
<paragraph id="B80E3687FFD42803FF46F98C8592E7D4" blockId="8.[247,1385,1545,1712]" pageId="8">
<emphasis id="8AC5EA95FFD42803FF46F98C8409E758" box="[247,604,1545,1571]" italics="true" pageId="8">Meso- and metathoracic legs</emphasis>
(
<figureCitation id="208A2A02FFD42803FDDDF98C84EFE758" box="[620,698,1545,1571]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="8">Figs 1</figureCitation>
A−B; 3 A−C): Long and slender, without dilatations or projections. Femora with rounded genicular lobes and a short genicular spur. Tibiae with two apical spurs. Tarsi 5-segmented, green and slightly darkened apically. First tarsomere of mesotarsus shorter than remaining segments combined, first tarsomere of metatarsus about equal in length compared to remaining segments combined.
</paragraph>
<paragraph id="B80E3687FFD42803FF46F94385D9E633" blockId="8.[247,1386,1733,1864]" pageId="8">
<emphasis id="8AC5EA95FFD42803FF46F9438781E7A4" box="[247,468,1733,1759]" italics="true" pageId="8">Tegmina and alae</emphasis>
(
<figureCitation id="208A2A02FFD42803FE55F9408467E7A4" box="[484,562,1733,1759]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="8">Figs 1</figureCitation>
A−B; 3 A−C): Tegmina uniformly green on costal area, hyaline in discoidal area, longer than tip of abdomen when folded in rest. Veins green, stigma white with strong hue of yellowish-green. Stigma located anteriorly from middle of tegmen. Alae fully developed, hyaline with only greenish anterior margin.
</paragraph>
<paragraph id="B80E3687FFD52802FF46FF3F845EE061" blockId="9.[247,1386,186,283]" pageId="9">
<emphasis id="8AC5EA95FFD52802FF46FF3F873CE1AF" box="[247,361,186,212]" italics="true" pageId="9">Abdomen</emphasis>
(
<figureCitation id="208A2A02FFD52802FEC8FF3F8794E1AF" box="[377,449,186,212]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="9">Fig. 1</figureCitation>
A−B): Fusiform, uniformly green ventrally, yellowish-green dorsally. Cerci setose, not flattened, with 22−23 segments, proximal segments being merged together and difficult to distinguish.
</paragraph>
<caption id="ECCE660FFFD52802FF46FEB9847EE00B" pageId="9" startId="9.[247,304,316,338]" targetType="table">
<paragraph id="B80E3687FFD52802FF46FEB9847EE00B" blockId="9.[247,1385,316,368]" pageId="9">
Table 2. Measurements [mm] of
<taxonomicName id="7FB14D04FFD52802FDFEFEB98500E02A" authorityName="Vermeersch" authorityYear="2001" box="[591,853,316,337]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFD52802FDFEFEB98500E02A" box="[591,853,316,337]" italics="true" pageId="9">Titanodula attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD52802FCEFFEB985E5E02A" box="[862,944,316,337]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="7FB14D04FFD52802FC57FEB98284E02A" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[998,1233,316,337]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD52802FC57FEB98284E02A" box="[998,1233,316,337]" italics="true" pageId="9">Titanodula fruhstorferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD52802FB6BFEB98318E02A" box="[1242,1357,316,338]" pageId="9" rank="species">comb. nov.</taxonomicNameLabel>
to assess intraspecific variability.
</paragraph>
</caption>
<paragraph id="B80E3687FFD52802FE25FE0082B8E471" pageId="9">
<table id="CAB1C427FFD5D7F4FE25FE0182BBE471" box="[404,1262,388,1290]" gridcols="5" gridrows="21" pageId="9">
<tr id="068134C5FFD5D7F4FE25FE0182BBE0E7" box="[404,1262,388,412]" gridrow="0" pageId="9">
<th id="45505DB9FFD5D7F4FE25FE0182BBE0E7" box="[404,1262,388,412]" colspan="5" colspanRight="4" gridcol="0" gridrow="0" pageId="9">
<emphasis id="8AC5EA95FFD52802FE25FE0085A4E0E7" bold="true" box="[404,1009,388,412]" pageId="9">
<taxonomicName id="7FB14D04FFD52802FE25FE0084F5E0E0" authorityName="Vermeersch" authorityYear="2001" box="[404,672,389,411]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFD52802FE25FE0084F5E0E0" bold="true" box="[404,672,389,411]" italics="true" pageId="9">Titanodula attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD52802FD17FE0384ADE0E0" box="[678,760,390,411]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
(♂) - from 2 specimens
</emphasis>
</th>
</tr>
<tr id="068134C5FFD5D7F4FE25FE3782BBE0BC" box="[404,1262,434,455]" gridrow="1" pageId="9" rowspan-0="1">
<td id="45505DB9FFD5D7F4FD4FFE378563E0BC" box="[766,822,434,455]" gridcol="1" gridrow="1" pageId="9">
<emphasis id="8AC5EA95FFD52802FD4FFE378575E0BC" bold="true" box="[766,800,434,455]" pageId="9">TL</emphasis>
</td>
<td id="45505DB9FFD5D7F4FCC4FE3785F8E0BC" box="[885,941,434,455]" gridcol="2" gridrow="1" pageId="9">
<emphasis id="8AC5EA95FFD52802FCC4FE3785C2E0BC" bold="true" box="[885,919,434,455]" pageId="9">BL</emphasis>
</td>
<td id="45505DB9FFD5D7F4FC5DFE3782D6E0BC" box="[1004,1155,434,455]" gridcol="3" gridrow="1" pageId="9">
<emphasis id="8AC5EA95FFD52802FC5DFE378225E0BC" bold="true" box="[1004,1136,434,455]" pageId="9">PL PW</emphasis>
</td>
<td id="45505DB9FFD5D7F4FB03FE3782BBE0BC" box="[1202,1262,434,455]" gridcol="4" gridrow="1" pageId="9">
<emphasis id="8AC5EA95FFD52802FB03FE3782B5E0BC" bold="true" box="[1202,1248,434,455]" pageId="9">TgL</emphasis>
</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FE5B82BBE08F" box="[404,1262,478,500]" gridrow="2" pageId="9">
<th id="45505DB9FFD5D7F4FE25FE5B8402E08F" box="[404,599,478,500]" gridcol="0" gridrow="2" pageId="9">Average</th>
<td id="45505DB9FFD5D7F4FD4FFE5B8563E08F" box="[766,822,478,500]" gridcol="1" gridrow="2" pageId="9">95.6</td>
<td id="45505DB9FFD5D7F4FCC4FE5B85F8E08F" box="[885,941,478,500]" gridcol="2" gridrow="2" pageId="9">-</td>
<td id="45505DB9FFD5D7F4FC5DFE5B82D6E08F" box="[1004,1155,478,500]" gridcol="3" gridrow="2" pageId="9">28.9 8.2</td>
<td id="45505DB9FFD5D7F4FB03FE5B82BBE08F" box="[1202,1262,478,500]" gridcol="4" gridrow="2" pageId="9">65.4</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FD8F82BBE35B" box="[404,1262,522,544]" gridrow="3" pageId="9">
<th id="45505DB9FFD5D7F4FE25FD8F8402E35B" box="[404,599,522,544]" gridcol="0" gridrow="3" pageId="9">Standard Deviation</th>
<td id="45505DB9FFD5D7F4FD4FFD8F8563E35B" box="[766,822,522,544]" gridcol="1" gridrow="3" pageId="9">4</td>
<td id="45505DB9FFD5D7F4FCC4FD8F85F8E35B" box="[885,941,522,544]" gridcol="2" gridrow="3" pageId="9">-</td>
<td id="45505DB9FFD5D7F4FC5DFD8F82D6E35B" box="[1004,1155,522,544]" gridcol="3" gridrow="3" pageId="9">0.45 0</td>
<td id="45505DB9FFD5D7F4FB03FD8F82BBE35B" box="[1202,1262,522,544]" gridcol="4" gridrow="3" pageId="9">3.6</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FDB382BBE337" box="[404,1262,566,588]" gridrow="4" pageId="9">
<th id="45505DB9FFD5D7F4FE25FDB38402E337" box="[404,599,566,588]" gridcol="0" gridrow="4" pageId="9">Max value</th>
<td id="45505DB9FFD5D7F4FD4FFDB38563E337" box="[766,822,566,588]" gridcol="1" gridrow="4" pageId="9">99.6</td>
<td id="45505DB9FFD5D7F4FCC4FDB385F8E337" box="[885,941,566,588]" gridcol="2" gridrow="4" pageId="9">-</td>
<td id="45505DB9FFD5D7F4FC5DFDB382D6E337" box="[1004,1155,566,588]" gridcol="3" gridrow="4" pageId="9">29.3 8.2</td>
<td id="45505DB9FFD5D7F4FB03FDB382BBE337" box="[1202,1262,566,588]" gridcol="4" gridrow="4" pageId="9">69</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FDE782BBE303" box="[404,1262,610,632]" gridrow="5" pageId="9">
<th id="45505DB9FFD5D7F4FE25FDE78402E303" box="[404,599,610,632]" gridcol="0" gridrow="5" pageId="9">Min value</th>
<td id="45505DB9FFD5D7F4FD4FFDE78563E303" box="[766,822,610,632]" gridcol="1" gridrow="5" pageId="9">91.6</td>
<td id="45505DB9FFD5D7F4FCC4FDE785F8E303" box="[885,941,610,632]" gridcol="2" gridrow="5" pageId="9">-</td>
<td id="45505DB9FFD5D7F4FC5DFDE782D6E303" box="[1004,1155,610,632]" gridcol="3" gridrow="5" pageId="9">28.4 8.2</td>
<td id="45505DB9FFD5D7F4FB03FDE782BBE303" box="[1202,1262,610,632]" gridcol="4" gridrow="5" pageId="9">61.8</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FD0882BBE3D8" box="[404,1262,653,675]" gridrow="6" pageId="9">
<th id="45505DB9FFD5D7F4FE25FD0882BBE3D8" box="[404,1262,653,675]" colspan="5" colspanRight="4" gridcol="0" gridrow="6" pageId="9">
<emphasis id="8AC5EA95FFD52802FE24FD088553E3D8" bold="true" box="[405,774,653,675]" pageId="9">
<taxonomicName id="7FB14D04FFD52802FE24FD0884F5E3D8" authorityName="Vermeersch" authorityYear="2001" box="[405,672,653,675]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="8AC5EA95FFD52802FE24FD0884F5E3D8" bold="true" box="[405,672,653,675]" italics="true" pageId="9">Titanodula attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD52802FD17FD0B84ADE3D8" box="[678,760,654,675]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
(
</emphasis>
<emphasis id="8AC5EA95FFD52802FCB7FD088542E3D9" bold="true" box="[774,791,653,674]" pageId="9">♀</emphasis>
<emphasis id="8AC5EA95FFD52802FCA6FD0885BAE3D8" bold="true" box="[791,1007,653,675]" pageId="9">) - from 6 specimens</emphasis>
</th>
</tr>
<tr id="068134C5FFD5D7F4FE25FD3F82BBE3B4" box="[404,1262,698,719]" gridrow="7" pageId="9" rowspan-0="1">
<td id="45505DB9FFD5D7F4FD4FFD3F8563E3B4" box="[766,822,698,719]" gridcol="1" gridrow="7" pageId="9">
<emphasis id="8AC5EA95FFD52802FD4FFD3F8575E3B4" bold="true" box="[766,800,698,719]" pageId="9">TL</emphasis>
</td>
<td id="45505DB9FFD5D7F4FCC4FD3F85F8E3B4" box="[885,941,698,719]" gridcol="2" gridrow="7" pageId="9">
<emphasis id="8AC5EA95FFD52802FCC4FD3F85C2E3B4" bold="true" box="[885,919,698,719]" pageId="9">BL</emphasis>
</td>
<td id="45505DB9FFD5D7F4FC5DFD3F82D6E3B4" box="[1004,1155,698,719]" gridcol="3" gridrow="7" pageId="9">
<emphasis id="8AC5EA95FFD52802FC5DFD3F8225E3B4" bold="true" box="[1004,1136,698,719]" pageId="9">PL PW</emphasis>
</td>
<td id="45505DB9FFD5D7F4FB03FD3F82BBE3B4" box="[1202,1262,698,719]" gridcol="4" gridrow="7" pageId="9">
<emphasis id="8AC5EA95FFD52802FB03FD3F82B5E3B4" bold="true" box="[1202,1248,698,719]" pageId="9">TgL</emphasis>
</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FD6082BBE380" box="[404,1262,741,763]" gridrow="8" pageId="9">
<th id="45505DB9FFD5D7F4FE25FD608402E380" box="[404,599,741,763]" gridcol="0" gridrow="8" pageId="9">Average</th>
<td id="45505DB9FFD5D7F4FD4FFD608563E380" box="[766,822,741,763]" gridcol="1" gridrow="8" pageId="9">105.1</td>
<td id="45505DB9FFD5D7F4FCC4FD6085F8E380" box="[885,941,741,763]" gridcol="2" gridrow="8" pageId="9">99.8</td>
<td id="45505DB9FFD5D7F4FC5DFD6082D6E380" box="[1004,1155,741,763]" gridcol="3" gridrow="8" pageId="9">36.3 10.8</td>
<td id="45505DB9FFD5D7F4FB03FD6082BBE380" box="[1202,1262,741,763]" gridcol="4" gridrow="8" pageId="9">64</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FC9482BBE25C" box="[404,1262,785,807]" gridrow="9" pageId="9">
<th id="45505DB9FFD5D7F4FE25FC948402E25C" box="[404,599,785,807]" gridcol="0" gridrow="9" pageId="9">Standard Deviation</th>
<td id="45505DB9FFD5D7F4FD4FFC948563E25C" box="[766,822,785,807]" gridcol="1" gridrow="9" pageId="9">7</td>
<td id="45505DB9FFD5D7F4FCC4FC9485F8E25C" box="[885,941,785,807]" gridcol="2" gridrow="9" pageId="9">9.2</td>
<td id="45505DB9FFD5D7F4FC5DFC9482D6E25C" box="[1004,1155,785,807]" gridcol="3" gridrow="9" pageId="9">2.6 0.7</td>
<td id="45505DB9FFD5D7F4FB03FC9482BBE25C" box="[1202,1262,785,807]" gridcol="4" gridrow="9" pageId="9">4.8</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FCB882BBE228" box="[404,1262,829,851]" gridrow="10" pageId="9">
<th id="45505DB9FFD5D7F4FE25FCB88402E228" box="[404,599,829,851]" gridcol="0" gridrow="10" pageId="9">Max value</th>
<td id="45505DB9FFD5D7F4FD4FFCB88563E228" box="[766,822,829,851]" gridcol="1" gridrow="10" pageId="9">115</td>
<td id="45505DB9FFD5D7F4FCC4FCB885F8E228" box="[885,941,829,851]" gridcol="2" gridrow="10" pageId="9">115</td>
<td id="45505DB9FFD5D7F4FC5DFCB882D6E228" box="[1004,1155,829,851]" gridcol="3" gridrow="10" pageId="9">39.6 12</td>
<td id="45505DB9FFD5D7F4FB03FCB882BBE228" box="[1202,1262,829,851]" gridcol="4" gridrow="10" pageId="9">71.3</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FCEC82BBE204" box="[404,1262,873,895]" gridrow="11" pageId="9">
<th id="45505DB9FFD5D7F4FE25FCEC8402E204" box="[404,599,873,895]" gridcol="0" gridrow="11" pageId="9">Min value</th>
<td id="45505DB9FFD5D7F4FD4FFCEC8563E204" box="[766,822,873,895]" gridcol="1" gridrow="11" pageId="9">97.7</td>
<td id="45505DB9FFD5D7F4FCC4FCEC85F8E204" box="[885,941,873,895]" gridcol="2" gridrow="11" pageId="9">88.4</td>
<td id="45505DB9FFD5D7F4FC5DFCEC82D6E204" box="[1004,1155,873,895]" gridcol="3" gridrow="11" pageId="9">33.8 9.9</td>
<td id="45505DB9FFD5D7F4FB03FCEC82BBE204" box="[1202,1262,873,895]" gridcol="4" gridrow="11" pageId="9">59.2</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FC1182BBE2D0" box="[404,1262,916,939]" gridrow="12" pageId="9">
<th id="45505DB9FFD5D7F4FE25FC1182BBE2D0" box="[404,1262,916,939]" colspan="5" colspanRight="4" gridcol="0" gridrow="12" pageId="9">
<emphasis id="8AC5EA95FFD52802FE24FC11855AE2D1" bold="true" box="[405,783,916,938]" pageId="9">
<taxonomicName id="7FB14D04FFD52802FE24FC1184D3E2D1" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[405,646,916,938]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD52802FE24FC1184D3E2D1" bold="true" box="[405,646,916,938]" italics="true" pageId="9">Titanodula fruhstorferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD52802FD3CFC108554E2D1" box="[653,769,917,938]" pageId="9" rank="species">comb. nov.</taxonomicNameLabel>
(
</emphasis>
<emphasis id="8AC5EA95FFD52802FCA1FC108574E2D1" bold="true" box="[784,801,917,938]" pageId="9">♂</emphasis>
<emphasis id="8AC5EA95FFD52802FC90FC118250E2D0" bold="true" box="[801,1029,916,939]" pageId="9">) - from 14 specimens</emphasis>
</th>
</tr>
<tr id="068134C5FFD5D7F4FE25FC4482BBE2AD" box="[404,1262,961,982]" gridrow="13" pageId="9" rowspan-0="1">
<td id="45505DB9FFD5D7F4FD4FFC448563E2AD" box="[766,822,961,982]" gridcol="1" gridrow="13" pageId="9">
<emphasis id="8AC5EA95FFD52802FD4FFC448575E2AD" bold="true" box="[766,800,961,982]" pageId="9">TL</emphasis>
</td>
<td id="45505DB9FFD5D7F4FCC4FC4485F8E2AD" box="[885,941,961,982]" gridcol="2" gridrow="13" pageId="9">
<emphasis id="8AC5EA95FFD52802FCC4FC4485C2E2AD" bold="true" box="[885,919,961,982]" pageId="9">BL</emphasis>
</td>
<td id="45505DB9FFD5D7F4FC5DFC4482D6E2AD" box="[1004,1155,961,982]" gridcol="3" gridrow="13" pageId="9">
<emphasis id="8AC5EA95FFD52802FC5DFC448225E2AD" bold="true" box="[1004,1136,961,982]" pageId="9">PL PW</emphasis>
</td>
<td id="45505DB9FFD5D7F4FB03FC4482BBE2AD" box="[1202,1262,961,982]" gridcol="4" gridrow="13" pageId="9">
<emphasis id="8AC5EA95FFD52802FB03FC4482B5E2AD" bold="true" box="[1202,1248,961,982]" pageId="9">TgL</emphasis>
</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FC6882BBE578" box="[404,1262,1005,1027]" gridrow="14" pageId="9">
<th id="45505DB9FFD5D7F4FE25FC688402E578" box="[404,599,1005,1027]" gridcol="0" gridrow="14" pageId="9">Average</th>
<td id="45505DB9FFD5D7F4FD4FFC688563E578" box="[766,822,1005,1027]" gridcol="1" gridrow="14" pageId="9">105.4</td>
<td id="45505DB9FFD5D7F4FCC4FC6885F8E578" box="[885,941,1005,1027]" gridcol="2" gridrow="14" pageId="9">90.46</td>
<td id="45505DB9FFD5D7F4FC5DFC6882D6E578" box="[1004,1155,1005,1027]" gridcol="3" gridrow="14" pageId="9">30.7 8.8</td>
<td id="45505DB9FFD5D7F4FB03FC6882BBE578" box="[1202,1262,1005,1027]" gridcol="4" gridrow="14" pageId="9">71.9</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FB9D82BBE555" box="[404,1262,1048,1070]" gridrow="15" pageId="9">
<th id="45505DB9FFD5D7F4FE25FB9D8402E555" box="[404,599,1048,1070]" gridcol="0" gridrow="15" pageId="9">Standard Deviation</th>
<td id="45505DB9FFD5D7F4FD4FFB9D8563E555" box="[766,822,1048,1070]" gridcol="1" gridrow="15" pageId="9">4.5</td>
<td id="45505DB9FFD5D7F4FCC4FB9D85F8E555" box="[885,941,1048,1070]" gridcol="2" gridrow="15" pageId="9">3.8</td>
<td id="45505DB9FFD5D7F4FC5DFB9D82D6E555" box="[1004,1155,1048,1070]" gridcol="3" gridrow="15" pageId="9">1.5 0.4</td>
<td id="45505DB9FFD5D7F4FB03FB9D82BBE555" box="[1202,1262,1048,1070]" gridcol="4" gridrow="15" pageId="9">3.3</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FBC182BBE521" box="[404,1262,1092,1114]" gridrow="16" pageId="9">
<th id="45505DB9FFD5D7F4FE25FBC18402E521" box="[404,599,1092,1114]" gridcol="0" gridrow="16" pageId="9">Max value</th>
<td id="45505DB9FFD5D7F4FD4FFBC18563E521" box="[766,822,1092,1114]" gridcol="1" gridrow="16" pageId="9">110.3</td>
<td id="45505DB9FFD5D7F4FCC4FBC185F8E521" box="[885,941,1092,1114]" gridcol="2" gridrow="16" pageId="9">96.8</td>
<td id="45505DB9FFD5D7F4FC5DFBC182D6E521" box="[1004,1155,1092,1114]" gridcol="3" gridrow="16" pageId="9">33.5 9.3</td>
<td id="45505DB9FFD5D7F4FB03FBC182BBE521" box="[1202,1262,1092,1114]" gridcol="4" gridrow="16" pageId="9">75.5</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FBF582BBE5FD" box="[404,1262,1136,1158]" gridrow="17" pageId="9">
<th id="45505DB9FFD5D7F4FE25FBF58402E5FD" box="[404,599,1136,1158]" gridcol="0" gridrow="17" pageId="9">Min value</th>
<td id="45505DB9FFD5D7F4FD4FFBF58563E5FD" box="[766,822,1136,1158]" gridcol="1" gridrow="17" pageId="9">95</td>
<td id="45505DB9FFD5D7F4FCC4FBF585F8E5FD" box="[885,941,1136,1158]" gridcol="2" gridrow="17" pageId="9">81.7</td>
<td id="45505DB9FFD5D7F4FC5DFBF582D6E5FD" box="[1004,1155,1136,1158]" gridcol="3" gridrow="17" pageId="9">27.8 8.1</td>
<td id="45505DB9FFD5D7F4FB03FBF582BBE5FD" box="[1202,1262,1136,1158]" gridcol="4" gridrow="17" pageId="9">64.5</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FB1982BBE5C9" box="[404,1262,1180,1202]" gridrow="18" pageId="9">
<th id="45505DB9FFD5D7F4FE25FB1982BBE5C9" box="[404,1262,1180,1202]" colspan="5" colspanRight="4" gridcol="0" gridrow="18" pageId="9">
<emphasis id="8AC5EA95FFD52802FE24FB19855AE5C9" bold="true" box="[405,783,1180,1202]" pageId="9">
<taxonomicName id="7FB14D04FFD52802FE24FB1984D3E5C9" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[405,646,1180,1202]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
<emphasis id="8AC5EA95FFD52802FE24FB1984D3E5C9" bold="true" box="[405,646,1180,1202]" italics="true" pageId="9">Titanodula fruhstorferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD52802FD3CFB198557E5CA" box="[653,770,1180,1201]" pageId="9" rank="species">comb. nov.</taxonomicNameLabel>
(
</emphasis>
<emphasis id="8AC5EA95FFD52802FCA1FB198574E5CA" bold="true" box="[784,801,1180,1201]" pageId="9">♀</emphasis>
<emphasis id="8AC5EA95FFD52802FC90FB198266E5C9" bold="true" box="[801,1075,1180,1202]" pageId="9">) - from 1 specimen [AS2]</emphasis>
</th>
</tr>
<tr id="068134C5FFD5D7F4FE25FB4D82BBE5A6" box="[404,1262,1224,1245]" gridrow="19" pageId="9" rowspan-0="1">
<td id="45505DB9FFD5D7F4FD4FFB4D8563E5A6" box="[766,822,1224,1245]" gridcol="1" gridrow="19" pageId="9">
<emphasis id="8AC5EA95FFD52802FD4FFB4D8575E5A6" bold="true" box="[766,800,1224,1245]" pageId="9">TL</emphasis>
</td>
<td id="45505DB9FFD5D7F4FCC4FB4D85F8E5A6" box="[885,941,1224,1245]" gridcol="2" gridrow="19" pageId="9">
<emphasis id="8AC5EA95FFD52802FCC4FB4D85C2E5A6" bold="true" box="[885,919,1224,1245]" pageId="9">BL</emphasis>
</td>
<td id="45505DB9FFD5D7F4FC5DFB4D82D6E5A6" box="[1004,1155,1224,1245]" gridcol="3" gridrow="19" pageId="9">
<emphasis id="8AC5EA95FFD52802FC5DFB4D8225E5A6" bold="true" box="[1004,1136,1224,1245]" pageId="9">PL PW</emphasis>
</td>
<td id="45505DB9FFD5D7F4FB03FB4D82BBE5A6" box="[1202,1262,1224,1245]" gridcol="4" gridrow="19" pageId="9">
<emphasis id="8AC5EA95FFD52802FB03FB4D82B5E5A6" bold="true" box="[1202,1248,1224,1245]" pageId="9">TgL</emphasis>
</td>
</tr>
<tr id="068134C5FFD5D7F4FE25FB7182BBE471" box="[404,1262,1268,1290]" gridrow="20" pageId="9">
<th id="45505DB9FFD5D7F4FE25FB718402E471" box="[404,599,1268,1290]" gridcol="0" gridrow="20" pageId="9">Value</th>
<td id="45505DB9FFD5D7F4FD4FFB718563E471" box="[766,822,1268,1290]" gridcol="1" gridrow="20" pageId="9">93.7</td>
<td id="45505DB9FFD5D7F4FCC4FB7185F8E471" box="[885,941,1268,1290]" gridcol="2" gridrow="20" pageId="9">91.9</td>
<td id="45505DB9FFD5D7F4FC5DFB7182D6E471" box="[1004,1155,1268,1290]" gridcol="3" gridrow="20" pageId="9">33.2 10.2</td>
<td id="45505DB9FFD5D7F4FB03FB7182BBE471" box="[1202,1262,1268,1290]" gridcol="4" gridrow="20" pageId="9">59.2</td>
</tr>
</table>
</paragraph>
<paragraph id="B80E3687FFD52801FF46FAD582E1E18C" blockId="9.[247,1386,1360,1878]" lastBlockId="10.[247,1385,186,248]" lastPageId="10" pageId="9">
<emphasis id="8AC5EA95FFD52802FF46FAD587F3E411" box="[247,422,1360,1386]" italics="true" pageId="9">Male genitalia</emphasis>
(
<figureCitation id="208A2A02FFD52802FE04FAD587A9E410" box="[437,508,1360,1387]" captionStart="Fig" captionStartId="12.[247,284,1798,1820]" captionTargetBox="[354,1288,185,1775]" captionTargetId="figure-74@12.[330,1303,178,1778]" captionTargetPageId="12" captionText="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." pageId="9">Fig. 5</figureCitation>
B−D): Ventral phallomere of left phallic complex elongated, much longer than broad, with two short and strongly sclerotised distal processes (sdp) of about the same length with acute apex. Median process (sdpm) bend at the middle and pointing sideways, lateral process (sdpl) much more strongly bend and pointing upwards, with broader base than sdpm. Apical process of left phallomere (paa) complex in shape, with sclerotised apex and sclerotizations near the base, but membranous in the middle. Apex abruptly projecting dorsally and to the left, the apex sharply curved in a 90° hook with narrower but rounded tip. Phalloid apophysis (afa) with two sclerotised processes, anterior process (aafa) small and tubercle-shaped, broad and dome-like at the base with a smaller rounded projection on top of it, located posteriorly from the middle. Posterior process (pafa) spear-shaped, almost straight from the base onwards, long and heavily sclerotised, projecting straight or slightly diagonally posteriad. Membranous lobe loa rounded and small. Genital lobe goa strongly developed, extending laterally over its whole length. Right phallomere with sclerite R1A more or less triangular. Sclerite R3 shovel-shaped, very wide at its anterior end. Ventral process (pia) triangular with many longitudinal grooves on its surface, sclerified ventral process (pva) strongly sclerotized, finger-shaped, ending in a blunt hook. Coxosternite IX (subgenital plate) with small black spikes on the right lateral edge, sometimes also in the middle of the distal edge.
</paragraph>
<caption id="ECCE660FFFD62801FF46FA128482E4B1" pageId="10" startId="10.[247,284,1431,1453]" targetBox="[247,1385,273,1410]" targetPageId="10" targetType="figure">
<paragraph id="B80E3687FFD62801FF46FA128482E4B1" blockId="10.[247,1385,1431,1483]" pageId="10">
Fig. 3.
<taxonomicName id="7FB14D04FFD62801FEF1FA128416E4D7" authorityName="Vermeersch" authorityYear="2001" box="[320,579,1431,1452]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="10" phylum="Arthropoda" rank="species" species="attenboroughi" status="gen. nov. et sp. nov.">
<emphasis id="8AC5EA95FFD62801FEF1FA128416E4D7" box="[320,579,1431,1452]" italics="true" pageId="10">Titanodula attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="91F657EEFFD62801FDFBFA12854CE4D7" box="[586,793,1431,1452]" pageId="10" rank="species">gen. nov. et sp. nov.</taxonomicNameLabel>
, live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang
<taxonomicNameLabel id="91F657EEFFD62801FD83FA308431E4B1" box="[562,612,1461,1482]" pageId="10" rank="subSpecies">N.R.</taxonomicNameLabel>
, VII. 2018.
</paragraph>
</caption>
<paragraph id="B80E3687FFD62801FF46FA6887D3E752" blockId="10.[247,1385,1516,1577]" pageId="10">
<smallCapsWord id="BEE8A05BFFD62801FF46FA68870FE77E" baselines="1536,1536" box="[247,346,1517,1543]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Female" pageId="10">FEMALE</smallCapsWord>
(
<figureCitation id="208A2A02FFD62801FEDFFA6987EAE77D" box="[366,447,1516,1542]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="10">Figs 2</figureCitation>
,
<figureCitation id="208A2A02FFD62801FE61FA69878AE77D" box="[464,479,1516,1542]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="10">3</figureCitation>
D−E). Measurements (see
<tableCitation id="F533033CFFD62801FC86FA698590E77D" box="[823,965,1516,1542]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="7.[247,304,421,443]" captionStartId-1="9.[247,304,316,338]" captionText-0="Table 1. Detailed measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. From left to right: T. attenboroughi holotype ♂, T. attenboroughi paratype ♀, T. fruhstorferi male, additional specimen 1, T. fruhstorferi female, additional specimen 2." captionText-1="Table 2. Measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. to assess intraspecific variability." pageId="10">Tables 1−2</tableCitation>
). Similar to male, with following differences:
</paragraph>
<paragraph id="B80E3687FFD62801FF46F9BA8593E707" blockId="10.[247,1385,1599,1660]" pageId="10">
<emphasis id="8AC5EA95FFD62801FF46F9BA871BE722" box="[247,334,1599,1625]" italics="true" pageId="10">Thorax</emphasis>
(
<figureCitation id="208A2A02FFD62801FEEEF9BA87F9E721" box="[351,428,1599,1626]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="10">Figs 2</figureCitation>
A−B, 3 D−E): Margins of pronotum finely denticulated anteriorly, fading out after pronotal supracoxal dilatation, absent in posterior end.
</paragraph>
<paragraph id="B80E3687FFD62801FF46F91784CEE789" blockId="10.[247,1385,1682,1779]" pageId="10">
<emphasis id="8AC5EA95FFD62801FF46F917879CE7D7" box="[247,457,1682,1708]" italics="true" pageId="10">Prothoracic legs</emphasis>
(
<figureCitation id="208A2A02FFD62801FE6CF9178465E7D7" box="[477,560,1682,1708]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="10">Figs 2</figureCitation>
A−B, D, 3 D−E): Coxae with 8−10 small sharp and regularly interspaced spinules of more or less equal size with same colour as coxa, spinules larger and more pronounced than in the male.
</paragraph>
<paragraph id="B80E3687FFD62801FF46F88C84A2E6F0" blockId="10.[247,1385,1800,1932]" pageId="10">
<emphasis id="8AC5EA95FFD62801FF46F88C879AE659" box="[247,463,1800,1826]" italics="true" pageId="10">Tegmina and alae</emphasis>
(
<figureCitation id="208A2A02FFD62801FE6DF88D8472E659" box="[476,551,1800,1826]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="10">Figs 2</figureCitation>
A−B, 3 D−E): Tegmina uniformly green in costal area, partly hyaline in discoidal area, being green anteriorly, then quickly becoming hyaline in a gradient posteriorly. Tegmina reaching tip of abdomen when folded in rest. Alae fully developed, hyaline with faintly green anterior margin.
</paragraph>
<paragraph id="B80E3687FFD72800FF46FF3F8218E18C" blockId="11.[247,1385,186,248]" pageId="11">
<emphasis id="8AC5EA95FFD72800FF46FF3F873CE1AF" box="[247,361,186,212]" italics="true" pageId="11">Abdomen</emphasis>
(
<figureCitation id="208A2A02FFD72800FEC8FF3F87B0E1AF" box="[377,485,186,212]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="11">Figs 2 A</figureCitation>
,
<figureCitation id="208A2A02FFD72800FE43FF3F8476E1AF" box="[498,547,186,212]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="11">3 D</figureCitation>
): Elongated ovaliform, much broader than in male, uniformly green ventrally, bright yellowish dorsally with a hint of green near the edges.
</paragraph>
<paragraph id="B80E3687FFD72800FF46FE88840FE0AD" blockId="11.[247,1385,269,470]" pageId="11">
<smallCapsWord id="BEE8A05BFFD72800FF46FE88876EE05D" baselines="288,289" box="[247,315,269,295]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Note" pageId="11">NOTE</smallCapsWord>
: The shape of the pronotum in females appears to be somewhat variable between different locations. Females from Da Krong
<taxonomicNameLabel id="91F657EEFFD72800FD74FEB584ABE031" box="[709,766,304,330]" pageId="11" rank="subSpecies">N.R.</taxonomicNameLabel>
have a noticeably more pronounced supracoxal dilatation and a metazona that appears thinner, while females from Kon Chu Rang
<taxonomicNameLabel id="91F657EEFFD72800FB6AFED18341E015" box="[1243,1300,340,366]" pageId="11" rank="subSpecies">N.R.</taxonomicNameLabel>
have a more subtle sinusoidal transition of the pronotal edge between these parts. Male genitalia are the same between different locations and provide the most stable and reliable characters to correctly identify the species.
</paragraph>
</subSubSection>
<subSubSection id="F0AB650CFFD72800FF46FE6884D9E314" pageId="11" type="etymology">
<paragraph id="B80E3687FFD72800FF46FE6884D9E314" blockId="11.[247,1385,492,623]" pageId="11">
<smallCapsWord id="BEE8A05BFFD72800FF46FE6887C4E37E" baselines="512,512" box="[247,401,493,519]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Etymology" pageId="11">ETYMOLOGY</smallCapsWord>
. The species epithet is a patronym dedicated to Sir David Attenborough, one of the world's most beloved naturalists, in acknowledgment for his life-long endeavours to disseminate knowledge on all the beings that are part of the natural world and to advocate for their protection and conservation.
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appears to be an endemic of undisturbed forests in the Vietnamese Annamite mountains. However, forests in the known distribution area are highly threatened and continue to disappear at an alarming rate. More distribution data are needed to evaluate the conservation status according to the IUCN Red Lists assessment guidelines, in the meanwhile the species is considered DD (Data Deficient).
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(
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<emphasis id="3DAFEACDAA0F16558F11F6C3581CFD2D" box="[565,789,698,724]" italics="true" pageId="4">Hierodula confusa</emphasis>
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<taxonomicNameLabel id="269C57B6AA0F16558E3FF6C35871FD2D" box="[795,888,698,724]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from the closest species
<taxonomicName id="C8DB4D5CAA0F16558DD3F6A75ACEFD0E" authorityName="Burmeister" authorityYear="1838" box="[247,455,733,759]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="species" species="membranacea">
<emphasis id="3DAFEACDAA0F16558DD3F6A75ACEFD0E" box="[247,455,733,759]" italics="true" pageId="4">H. membranacea</emphasis>
</taxonomicName>
by:
</paragraph>
<paragraph id="0F6436DFAA0F16558DD3F774583AFC2F" blockId="4.[247,1386,780,983]" pageId="4">
(1) The punctuation of the inner side of the procoxae:
<taxonomicName id="C8DB4D5CAA0F16558EA1F7745F01FCDE" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[901,1032,781,807]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA0F16558EA1F7745F01FCDE" box="[901,1032,781,807]" italics="true" pageId="4">H. confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA0F16558934F7745F67FCDE" box="[1040,1134,781,807]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="97E02A5AAA0F1655895AF7755FC2FCDE" box="[1150,1227,780,807]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="4">Figs 1</figureCitation>
A−C; 2 E) − small whitish tubercles are covering the entire anterior surface of the procoxae at irregular intervals, these also form a more or less straight row running parallel with the dorsal coxal margin;
<taxonomicName id="C8DB4D5CAA0F16558C7FF70E5921FC69" authorityName="Burmeister" authorityYear="1838" box="[347,552,886,912]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="species" species="membranacea">
<emphasis id="3DAFEACDAA0F16558C7FF70E5921FC69" box="[347,552,886,912]" italics="true" pageId="4">H. membranacea</emphasis>
</taxonomicName>
− only one single row of small whitish tubercles arranged in a straight row running parallel with the dorsal coxal margin (only faintly visible in some specimens), remaining anterior surface of procoxae smooth.
</paragraph>
<paragraph id="0F6436DFAA0F16558DD3F7955AADFB20" blockId="4.[247,1386,1004,1241]" pageId="4">
(2) Morphology of the male genitalia:
<taxonomicName id="C8DB4D5CAA0F16558FE7F794584FFBFF" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[707,838,1004,1030]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA0F16558FE7F794584FFBFF" box="[707,838,1004,1030]" italics="true" pageId="4">H. confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA0F16558E68F79558A2FBFF" box="[844,939,1004,1030]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="97E02A5AAA0F16558E9EF7955F01FBFF" box="[954,1032,1004,1030]" captionStart="Fig" captionStartId="8.[247,284,1876,1898]" captionTargetBox="[266,1367,1421,1845]" captionTargetId="figure-230@8.[247,1385,1411,1854]" captionTargetPageId="8" captionText="Fig. 4. Hierodula confusa sp. nov., genitalia prep. Vermeersch GEN1804. A, left phallic complex. B, detail of anterior process of left phallomere. C, right phallic complex. D, coxosternite IX (subgenital plate)." figureDoi="http://doi.org/10.5281/zenodo.13271608" httpUri="https://zenodo.org/record/13271608/files/figure.png" pageId="4">Figs 4</figureCitation>
;
<figureCitation id="97E02A5AAA0F16558931F7955F2DFBFE" box="[1045,1060,1004,1031]" captionStart="Fig" captionStartId="9.[247,284,1757,1779]" captionTargetBox="[270,1355,573,1708]" captionTargetId="figure-263@9.[247,1385,559,1737]" captionTargetPageId="9" captionText="Fig. 5. Hierodula membranacea, genitalia ♂: A, anterior process of left phallomere, dorsal view. B, distal part of left phallic complex, dorsal view. C, distal part of right phallic complex, ventral view. Hierodula confusa sp. nov., genitalia ♂: D, anterior process of left phallomere, dorsal view. E, distal part of left phallic complex, dorsal view. F, distal part of right phallic complex, ventral view. Rhombodera extensicollis, genitalia ♂: G, anterior process of left phallomere, dorsal view. H, distal part of left phallic complex, dorsal view. I, distal part of right phallic complex, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13271610" httpUri="https://zenodo.org/record/13271610/files/figure.png" pageId="4">5</figureCitation>
D−F) – afa with a strongly sclerotised and acute spine-shaped anterior process aafa, posterior process pafa irregularly shaped, wrinkled and membranous, completely and densely covered with very small and short spine-like setae, distal process with long and slender sdpl, bifurcating around the middle with very small sdpm (see also description below);
<taxonomicName id="C8DB4D5CAA0F16558E0AF00058F6FB6B" authorityName="Burmeister" authorityYear="1838" box="[814,1023,1144,1170]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="species" species="membranacea">
<emphasis id="3DAFEACDAA0F16558E0AF00058F6FB6B" box="[814,1023,1144,1170]" italics="true" pageId="4">H. membranacea</emphasis>
</taxonomicName>
(
<figureCitation id="97E02A5AAA0F16558934F0015F50FB6B" box="[1040,1113,1144,1170]" captionStart="Fig" captionStartId="10.[247,284,1894,1916]" captionText="Fig. 6. Distribution map of Hierodula confusa sp. nov." pageId="4">Fig. 6</figureCitation>
A−C) – aafa and pafa both small and sclerified, pafa longer than aafa, distal process sdpl very long and slender, without sdpm.
</paragraph>
<paragraph id="0F6436DFAA0F16558DD3F0975A4CFA8B" blockId="4.[247,1385,1262,1394]" pageId="4">
<taxonomicName id="C8DB4D5CAA0F16558DD3F0975AD0FAF1" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[247,473,1262,1288]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA0F16558DD3F0975AD0FAF1" box="[247,473,1262,1288]" italics="true" pageId="4">Hierodula confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA0F16558CC5F0965948FAF0" box="[481,577,1263,1289]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="C8DB4D5CAA0F16558FC6F0965893FAF1" authorityName="Serville" authorityYear="1839" box="[738,922,1262,1288]" class="Insecta" family="Mantidae" genus="Rhombodera" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="species" species="extensicollis">
<emphasis id="3DAFEACDAA0F16558FC6F0965893FAF1" box="[738,922,1262,1288]" italics="true" pageId="4">R. extensicollis</emphasis>
</taxonomicName>
by the shape of the pronotum without large lateral expansion, the missing expansion of the dorsal procoxal edge, smaller dark spots on the ventral side of meso- and metathorax, and the differences in male genitalia illustrated in
<figureCitation id="97E02A5AAA0F16558DD3F1215A4CFA8B" box="[247,325,1368,1394]" captionStart="Fig" captionStartId="10.[247,284,1894,1916]" captionText="Fig. 6. Distribution map of Hierodula confusa sp. nov." pageId="4">Fig. 6.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="47C16554AA0F16558DD3F1F15A58F9F2" pageId="4" type="etymology">
<paragraph id="0F6436DFAA0F16558DD3F1F15A58F9F2" blockId="4.[247,1386,1415,1547]" pageId="4">
<smallCapsWord id="0982A003AA0F16558DD3F1F15A98FA59" baselines="1435,1435" box="[247,401,1416,1442]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Etymology" pageId="4">ETYMOLOGY</smallCapsWord>
. The species epithet, from the Latin ‘confusa’ means vague/indefinite or obscure. The name is referring to the great similarity in external morphology with its close relative
<taxonomicName id="C8DB4D5CAA0F16558DD3F1B45920FA1E" authorityName="Burmeister" authorityYear="1838" box="[247,553,1485,1511]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="4" phylum="Arthropoda" rank="species" species="membranacea">
<emphasis id="3DAFEACDAA0F16558DD3F1B45920FA1E" box="[247,553,1485,1511]" italics="true" pageId="4">Hierodula membranacea</emphasis>
</taxonomicName>
and the subsequent confusions and misidentifications that resulted from it.
</paragraph>
</subSubSection>
<subSubSection id="47C16554AA0F16548DD3F2585911FE88" lastPageId="5" pageId="4" type="materials_examined">
<paragraph id="0F6436DFAA0F16558DD3F2585E60F8AA" blockId="4.[247,1385,1568,1876]" pageId="4">
<materialsCitation id="BFB33C82AA0F16558DD3F2585965F9C2" box="[247,620,1568,1595]" collectionCode="TYPE, MATERIAL" pageId="4" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<smallCapsWord id="0982A003AA0F16558DD3F2585A3FF9C0" baselines="1588,1588" box="[247,310,1569,1595]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Type" pageId="4">
<collectionCode id="69CAAE1AAA0F16558DD3F2585A3FF9C0" box="[247,310,1569,1595]" pageId="4">TYPE</collectionCode>
</smallCapsWord>
<smallCapsWord id="0982A003AA0F16558C67F25D5AB7F9C0" baselines="1588" box="[323,446,1572,1593]" lowerCaseFontSize="9" mainFontSize="11" normCase="lower" normString="material" pageId="4">
<collectionCode id="69CAAE1AAA0F16558C67F25D5AB7F9C0" box="[323,446,1572,1593]" pageId="4">MATERIAL</collectionCode>
</smallCapsWord>
.
<typeStatus id="D060887DAA0F16558CF1F259594CF9C3" box="[469,581,1568,1594]" pageId="4" type="holotype">Holotype</typeStatus>
<specimenCount id="19DDFD56AA0F16558F71F2585965F9C2" box="[597,620,1569,1595]" count="1" pageId="4" type="male">♂</specimenCount>
</materialsCitation>
and
<typeStatus id="D060887DAA0F16558F91F2585821F9C2" box="[693,808,1569,1595]" pageId="4" type="paratype">paratypes</typeStatus>
(
<specimenCount id="19DDFD56AA0F16558E66F2595860F9C3" box="[834,873,1568,1594]" count="3" pageId="4" type="male">3♂</specimenCount>
,
<specimenCount id="19DDFD56AA0F16558E5AF25958AAF9C3" box="[894,931,1568,1594]" count="4" pageId="4" type="female">4♀</specimenCount>
):
<materialsCitation id="BFB33C82AA0F16558EE2F2595F56F9C2" box="[966,1119,1568,1595]" collectionCode="TYPE, MATERIAL" isTailEnumeration="true" pageId="4" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="D060887DAA0F16558EE2F2595F3FF9C3" box="[966,1078,1568,1594]" pageId="4" type="holotype">Holotype</typeStatus>
<specimenCount id="19DDFD56AA0F16558961F2585F56F9C2" box="[1093,1119,1569,1595]" count="1" pageId="4" type="male">♂</specimenCount>
</materialsCitation>
:
<materialsCitation id="BFB33C82AA0F16558957F2595A74F978" collectingDate="2019-04-25" collectionCode="T, THNHM" collectorName="Unnahachote" country="Thailand" location="Ramintra" municipality="Kannayao dist." pageId="4" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="holotype">
<collectingCountry id="77CC764FAA0F16558957F2595FE8F9C3" box="[1139,1249,1568,1594]" name="Thailand" pageId="4">Thailand</collectingCountry>
,
<collectingRegion id="CD1FF83DAA0F165589D1F2595E6CF9C3" box="[1269,1381,1568,1594]" country="Thailand" name="Krung Thep Mahanakhon" pageId="4">Bangkok</collectingRegion>
,
<collectingMunicipality id="EF00ACA5AA0F16558DD3F23D5AB0F9A4" box="[247,441,1603,1630]" pageId="4">Kannayao dist.</collectingMunicipality>
,
<location id="0A046004AA0F16558CF5F23A5948F9A4" LSID="urn:lsid:plazi:treatment:877287C9AA0F165B8C82F64E59CAFCC9:0A046004AA0F16558CF5F23A5948F9A4" box="[465,577,1603,1629]" country="Thailand" municipality="Kannayao dist." name="Ramintra" pageId="4" stateProvince="Krung Thep Mahanakhon">Ramintra</location>
subdist.,
<date id="7B65101FAA0F16558FEEF23A585EF9A7" box="[714,855,1603,1630]" pageId="4" value="2019-04-25">
<collectingDate id="6B21E9F7AA0F16558FEEF23A585EF9A7" box="[714,855,1603,1630]" pageId="4" value="2019-04-25">25.IV.2019</collectingDate>
</date>
, leg.
<collectionCode id="69CAAE1AAA0F16558E88F23D58CBF9A7" box="[940,962,1604,1630]" name="Tavera, Department of Geology and Geophysics" pageId="4">T</collectionCode>
.
<collectorName id="A22E5309AA0F16558EFDF23A5F72F9A4" box="[985,1147,1603,1629]" pageId="4">Unnahachote</collectorName>
, THNHM-I-19528 (
<collectionCode id="69CAAE1AAA0F16558DDBF21E5A7CF978" box="[255,373,1639,1665]" pageId="4">THNHM</collectionCode>
)
</materialsCitation>
.
<typeStatus id="D060887DAA0F16558CACF21E5AFBF978" box="[392,498,1639,1665]" pageId="4" type="paratype">Paratype</typeStatus>
(
<specimenCount id="19DDFD56AA0F16558F20F21E592EF978" box="[516,551,1639,1665]" count="1" pageId="4" type="female">1♀</specimenCount>
):
<materialsCitation id="BFB33C82AA0F16558F65F21E5883F95D" collectingDate="2020-04-15" collectionCode="T, THNHM" collectorName="Unnahachote" country="Thailand" location="Ramintra" municipality="Kannayao dist." pageId="4" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="paratype">
<collectingCountry id="77CC764FAA0F16558F65F21E59A6F978" box="[577,687,1639,1665]" name="Thailand" pageId="4">Thailand</collectingCountry>
,
<collectingRegion id="CD1FF83DAA0F16558F9EF21E5823F978" box="[698,810,1639,1665]" country="Thailand" name="Krung Thep Mahanakhon" pageId="4">Bangkok</collectingRegion>
,
<collectingMunicipality id="EF00ACA5AA0F16558E11F21E58E2F978" box="[821,1003,1639,1665]" pageId="4">Kannayao dist.</collectingMunicipality>
,
<location id="0A046004AA0F16558ED3F21E5F6EF978" LSID="urn:lsid:plazi:treatment:877287C9AA0F165B8C82F64E59CAFCC9:0A046004AA0F16558ED3F21E5F6EF978" box="[1015,1127,1639,1665]" country="Thailand" municipality="Kannayao dist." name="Ramintra" pageId="4" stateProvince="Krung Thep Mahanakhon">Ramintra</location>
subdist.,
<date id="7B65101FAA0F165589FCF21F5E6DF978" box="[1240,1380,1638,1665]" pageId="4" value="2020-04-15">
<collectingDate id="6B21E9F7AA0F165589FCF21F5E6DF978" box="[1240,1380,1638,1665]" pageId="4" value="2020-04-15">15.IV.2020</collectingDate>
</date>
, leg.
<collectionCode id="69CAAE1AAA0F16558C12F2F35A45F95D" box="[310,332,1674,1700]" name="Tavera, Department of Geology and Geophysics" pageId="4">T</collectionCode>
.
<collectorName id="A22E5309AA0F16558C46F2F3590DF95D" box="[354,516,1674,1700]" pageId="4">Unnahachote</collectorName>
, THNHM-I-19529 (
<collectionCode id="69CAAE1AAA0F16558E29F2F3588AF95D" box="[781,899,1674,1700]" pageId="4">THNHM</collectionCode>
)
</materialsCitation>
.
<typeStatus id="D060887DAA0F16558E84F2F35F03F95D" box="[928,1034,1674,1700]" pageId="4" type="paratype">Paratype</typeStatus>
(
<specimenCount id="19DDFD56AA0F1655890DF2F35F45F95D" box="[1065,1100,1674,1700]" count="1" pageId="4" type="female">1♀</specimenCount>
):
<materialsCitation id="BFB33C82AA0F16558954F2F35A74F913" collectingDate="2019-12-15" collectionCode="T, THNHM" collectorName="Unnahachote" country="Thailand" location="Ramintra" municipality="Kannayao dist." pageId="4" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="paratype">
<collectingCountry id="77CC764FAA0F16558954F2F35FD7F95D" box="[1136,1246,1674,1700]" name="Thailand" pageId="4">Thailand</collectingCountry>
,
<collectingRegion id="CD1FF83DAA0F165589D1F2F35E6CF95D" box="[1269,1381,1674,1700]" country="Thailand" name="Krung Thep Mahanakhon" pageId="4">Bangkok</collectingRegion>
,
<collectingMunicipality id="EF00ACA5AA0F16558DD3F2D75ABEF93E" box="[247,439,1709,1736]" pageId="4">Kannayao dist.</collectingMunicipality>
,
<location id="0A046004AA0F16558CE8F2D45935F93E" LSID="urn:lsid:plazi:treatment:877287C9AA0F165B8C82F64E59CAFCC9:0A046004AA0F16558CE8F2D45935F93E" box="[460,572,1709,1735]" country="Thailand" municipality="Kannayao dist." name="Ramintra" pageId="4" stateProvince="Krung Thep Mahanakhon">Ramintra</location>
subdist.,
<date id="7B65101FAA0F16558FE5F2D45856F93E" box="[705,863,1709,1736]" pageId="4" value="2019-12-15">
<collectingDate id="6B21E9F7AA0F16558FE5F2D45856F93E" box="[705,863,1709,1736]" pageId="4" value="2019-12-15">15 XII.2019</collectingDate>
</date>
, leg.
<collectionCode id="69CAAE1AAA0F16558E95F2D758CEF931" box="[945,967,1710,1736]" name="Tavera, Department of Geology and Geophysics" pageId="4">T</collectionCode>
.
<collectorName id="A22E5309AA0F16558EFFF2D45F74F93E" box="[987,1149,1709,1735]" pageId="4">Unnahachote</collectorName>
, THNHM-I-19530 (
<collectionCode id="69CAAE1AAA0F16558DDBF2A95A7CF913" box="[255,373,1744,1770]" pageId="4">THNHM</collectionCode>
)
</materialsCitation>
.
<typeStatus id="D060887DAA0F16558CABF2A85AF1F912" box="[399,504,1745,1771]" pageId="4" type="paratype">Paratype</typeStatus>
(
<specimenCount id="19DDFD56AA0F16558F35F2A95930F913" box="[529,569,1744,1770]" count="1" pageId="4" type="male">1♂</specimenCount>
):
<materialsCitation id="BFB33C82AA0F16558F72F2A9590EF8F4" country="Thailand" elevation="200" latitude="13.584389" location="Suan Phueng dist." longLatPrecision="1" longitude="99.23222" pageId="4" specimenCount="200" stateProvince="Ratchaburi" typeStatus="paratype">
<collectingCountry id="77CC764FAA0F16558F72F2A959CDF913" box="[598,708,1744,1770]" name="Thailand" pageId="4">Thailand</collectingCountry>
,
<collectingRegion id="CD1FF83DAA0F16558FF3F2A958A3F912" box="[727,938,1744,1771]" country="Thailand" name="Ratchaburi" pageId="4">Ratchaburi prov.</collectingRegion>
,
<location id="0A046004AA0F16558E99F2A95FADF913" LSID="urn:lsid:plazi:treatment:877287C9AA0F165B8C82F64E59CAFCC9:0A046004AA0F16558E99F2A95FADF913" box="[957,1188,1744,1770]" country="Thailand" latitude="13.584389" longLatPrecision="1" longitude="99.23222" name="Suan Phueng dist." pageId="4" stateProvince="Ratchaburi">Suan Phueng dist.</location>
,
<geoCoordinate id="6AEF5018AA0F16558993F2A95E6DF912" box="[1207,1380,1744,1771]" degrees="13" direction="north" minutes="35" orientation="latitude" pageId="4" precision="1" seconds="03.8" value="13.584389">13°35´03.8”N</geoCoordinate>
;
<geoCoordinate id="6AEF5018AA0F16558DD3F28A5AB9F8F7" box="[247,432,1779,1806]" degrees="099" direction="east" minutes="13" orientation="longitude" pageId="4" precision="1" seconds="56.0" value="99.23222">099°13´56.0”E</geoCoordinate>
,
<specimenCount id="19DDFD56AA0F16558C9EF28A590EF8F4" box="[442,519,1779,1805]" count="200" pageId="4" type="generic">
<quantity id="C8239B3AAA0F16558C9EF28A590AF8F4" box="[442,515,1779,1805]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" unit="m" value="200.0">
<elevation id="84F6D1ECAA0F16558C9EF28A590AF8F4" box="[442,515,1779,1805]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="4" unit="m" value="200.0">200m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
<materialsCitation id="BFB33C82AA0F16558F28F28A5FA7F8F4" box="[524,1198,1779,1806]" collectingDate="2020-04-23" collectionCode="TUPC" collectorName="K. Jiaranaisakul" country="Thailand" pageId="4" specimenCount="1" stateProvince="Ratchaburi" typeStatus="paratype">
asl.,
<collectingMethod id="D69A4EC8AA0F16558F66F28A59B8F8F7" box="[578,689,1779,1806]" pageId="4">light trap</collectingMethod>
,
<date id="7B65101FAA0F16558F9FF28A584EF8F4" box="[699,839,1779,1805]" pageId="4" value="2020-04-23">
<collectingDate id="6B21E9F7AA0F16558F9FF28A584EF8F4" box="[699,839,1779,1805]" pageId="4" value="2020-04-23">23.IV.2020</collectingDate>
</date>
,
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leg. (
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)
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.
<typeStatus id="D060887DAA0F1655899CF28D5E28F8F7" box="[1208,1313,1780,1806]" pageId="4" type="paratype">Paratype</typeStatus>
(
<specimenCount id="19DDFD56AA0F16558810F28A5E5EF8F4" box="[1332,1367,1779,1805]" count="1" pageId="4" type="female">1♀</specimenCount>
):
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<collectingCountry id="77CC764FAA0F16558DD3F36F5A6CF8C9" box="[247,357,1814,1840]" name="Thailand" pageId="4">Thailand</collectingCountry>
,
<collectingRegion id="CD1FF83DAA0F16558C56F36F594CF8C8" box="[370,581,1814,1841]" country="Thailand" name="Nonthaburi" pageId="4">Nonthaburi prov.</collectingRegion>
,
<location id="0A046004AA0F16558F76F36F59F2F8C9" LSID="urn:lsid:plazi:treatment:877287C9AA0F165B8C82F64E59CAFCC9:0A046004AA0F16558F76F36F59F2F8C9" box="[594,763,1814,1841]" country="Thailand" name="Pak Kret dist." pageId="4" stateProvince="Nonthaburi">Pak Kret dist.</location>
,
<date id="7B65101FAA0F16558E2DF36F5883F8C9" box="[777,906,1814,1840]" pageId="4" value="2019-10-30">
<collectingDate id="6B21E9F7AA0F16558E2DF36F5883F8C9" box="[777,906,1814,1840]" pageId="4" value="2019-10-30">30.X.2019</collectingDate>
</date>
,
<collectorName id="A22E5309AA0F16558EBCF36E5F4FF8C9" box="[920,1094,1814,1841]" pageId="4">W. Chanaparn</collectorName>
leg. (
<collectionCode id="69CAAE1AAA0F165589A8F36F5FE9F8C9" box="[1164,1248,1814,1840]" pageId="4">TUPC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="D060887DAA0F165589D0F36E5E60F8C8" box="[1268,1385,1815,1841]" pageId="4" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="19DDFD56AA0F16558C26F3405A21F8AA" box="[258,296,1849,1875]" count="1" pageId="4" type="male">1♂</specimenCount>
,
<specimenCount id="19DDFD56AA0F16558C61F3405A63F8AA" box="[325,362,1849,1875]" count="1" pageId="4" type="female">1♀</specimenCount>
):
<materialsCitation id="BFB33C82AA0F16558CB0F3405E6FF8AA" box="[404,1382,1849,1876]" collectingDate="2019-07-20" country="Thailand" location="Ramintra" municipality="Kannayao dist." pageId="4" specimenCount="1" stateProvince="Krung Thep Mahanakhon" typeStatus="paratype">
<collectingCountry id="77CC764FAA0F16558CB0F340590BF8AA" box="[404,514,1849,1875]" name="Thailand" pageId="4">Thailand</collectingCountry>
,
<collectingRegion id="CD1FF83DAA0F16558F39F3405984F8AA" box="[541,653,1849,1875]" country="Thailand" name="Krung Thep Mahanakhon" pageId="4">Bangkok</collectingRegion>
,
<collectingMunicipality id="EF00ACA5AA0F16558F8CF3435867F8AA" box="[680,878,1849,1876]" pageId="4">Kannayao dist.</collectingMunicipality>
,
<location id="0A046004AA0F16558EADF34058F0F8AA" LSID="urn:lsid:plazi:treatment:877287C9AA0F165B8C82F64E59CAFCC9:0A046004AA0F16558EADF34058F0F8AA" box="[905,1017,1849,1875]" country="Thailand" municipality="Kannayao dist." name="Ramintra" pageId="4" stateProvince="Krung Thep Mahanakhon">Ramintra</location>
subdist.,
<date id="7B65101FAA0F165589AFF3405E28F8AA" box="[1163,1313,1849,1875]" pageId="4" value="2019-07-20">
<collectingDate id="6B21E9F7AA0F165589AFF3405E28F8AA" box="[1163,1313,1849,1875]" pageId="4" value="2019-07-20">20.VII.2019</collectingDate>
</date>
, leg
</materialsCitation>
.
</paragraph>
<paragraph id="0F6436DFAA0E16548DD3F4C6586BFF05" blockId="5.[247,1386,190,252]" pageId="5">
<collectionCode id="69CAAE1AAA0E16548DD3F4C65A04FF20" box="[247,269,191,217]" name="Tavera, Department of Geology and Geophysics" pageId="5">T</collectionCode>
.
<materialsCitation id="BFB33C82AA0E16548C3DF4C75856FF05" collectingDate="1804-01" collectionCode="RBINS" collectorName="Unnahachote. I. G. &amp; Vermeersch Nr. &amp; P. Rolin." country="Thailand" location="Genitalia" pageId="5" specimenCount="1" specimenCount-male="1" stateProvince="Krung Thep Mahanakhon" typeStatus="paratype">
<collectorName id="A22E5309AA0E16548C3DF4C75AFEFF20" box="[281,503,190,217]" pageId="5">Unnahachote. I.G.</collectorName>
:34.197 (
<collectionCode id="69CAAE1AAA0E16548F49F4C659C5FF20" box="[621,716,191,217]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="5" type="Museum">RBINS</collectionCode>
).
<typeStatus id="D060887DAA0E16548FC3F4C65859FF20" box="[743,848,191,217]" pageId="5" type="paratype">Paratype</typeStatus>
(
<specimenCount id="19DDFD56AA0E16548E4DF4C65899FF20" box="[873,912,191,217]" count="1" pageId="5" type="male">1♂</specimenCount>
):
<collectingCountry id="77CC764FAA0E16548E8AF4C75F15FF21" box="[942,1052,190,216]" name="Thailand" pageId="5">Thailand</collectingCountry>
,
<collectingRegion id="CD1FF83DAA0E1654890AF4C75F97FF21" box="[1070,1182,190,216]" country="Thailand" name="Krung Thep Mahanakhon" pageId="5">Bangkok</collectingRegion>
,
<location id="0A046004AA0E16548994F4C75E17FF21" LSID="urn:lsid:plazi:treatment:877287C9AA0F165B8C82F64E59CAFCC9:0A046004AA0E16548994F4C75E17FF21" box="[1200,1310,190,216]" country="Thailand" name="Genitalia" pageId="5" stateProvince="Krung Thep Mahanakhon">Genitalia</location>
prep.
<collectorName id="A22E5309AA0E16548DD3F49B5ABEFF05" box="[247,439,226,252]" pageId="5">Vermeersch Nr.</collectorName>
<date id="7B65101FAA0E16548C9BF49B5935FF05" box="[447,572,226,252]" pageId="5" value="1804-01">
<collectingDate id="6B21E9F7AA0E16548C9BF49B5935FF05" box="[447,572,226,252]" pageId="5" value="1804-01">GEN1804</collectingDate>
</date>
, leg.
<collectorName id="A22E5309AA0E16548F58F49B59EEFF05" box="[636,743,226,252]" pageId="5">P. Rolin.</collectorName>
(
<collectionCode id="69CAAE1AAA0E16548FD2F49B585CFF05" box="[758,853,226,252]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="5" type="Museum">RBINS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="0F6436DFAA0E16548DD3F56B5911FE88" blockId="5.[247,1385,273,370]" pageId="5">
<materialsCitation id="BFB33C82AA0E16548DD3F56B591DFE88" collectingDate="2018-04-15" collectionCode="MATERIAL, EXAMINED, FROM, PHOTOGRAPHS" collectorName="I. Dugdale &amp; P. Phetsri" country="Thailand" county="Baan Maka Nature Resort" latitude="12.843861" location="Kaeng Krachan" longLatPrecision="1" longitude="99.59089" municipality="Song Pe Nong" pageId="5" specimenCount="1" specimenCount-male="1" stateProvince="Phetchaburi">
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<collectionCode id="69CAAE1AAA0E16548F6EF56C59FEFED3" box="[586,759,277,298]" pageId="5">PHOTOGRAPHS</collectionCode>
</smallCapsWord>
.
<collectingCountry id="77CC764FAA0E16548E20F568587DFED2" box="[772,884,273,299]" name="Thailand" pageId="5">Thailand</collectingCountry>
:
<specimenCount id="19DDFD56AA0E16548E58F56858AEFED5" box="[892,935,273,300]" count="1" pageId="5" type="male">1 ♂</specimenCount>
(
<figureCitation id="97E02A5AAA0E16548E91F5685F10FED2" box="[949,1049,273,299]" captionStart="Fig" captionStartId="8.[247,284,1140,1162]" captionTargetBox="[247,1385,409,1120]" captionTargetId="figure-226@8.[247,1385,409,1120]" captionTargetPageId="8" captionText="Fig. 3. Hierodula confusa sp. nov., live photographs: A−B, paratype ♀ [PT1]. C, sub-adult female nymph. D, adult male at light trap, Baan Maka Nature Resort, Thailand, © I. Dugdale &amp; P. Phetsri." figureDoi="http://doi.org/10.5281/zenodo.13271606" httpUri="https://zenodo.org/record/13271606/files/figure.png" pageId="5">Fig. 3 D</figureCitation>
):
<collectingCounty id="E6054E53AA0E16548908F56B5E6CFED5" box="[1068,1381,273,300]" pageId="5">Baan Maka Nature Resort</collectingCounty>
,
<collectingMunicipality id="EF00ACA5AA0E16548DD3F54D5AB6FEB6" box="[247,447,308,335]" pageId="5">Song Pe Nong</collectingMunicipality>
,
<location id="0A046004AA0E16548CF3F54C59ABFEB7" LSID="urn:lsid:plazi:treatment:877287C9AA0F165B8C82F64E59CAFCC9:0A046004AA0E16548CF3F54C59ABFEB7" box="[471,674,308,335]" country="Thailand" county="Baan Maka Nature Resort" latitude="12.843861" longLatPrecision="1" longitude="99.59089" municipality="Song Pe Nong" name="Kaeng Krachan" pageId="5" stateProvince="Phetchaburi">Kaeng Krachan</location>
,
<collectingRegion id="CD1FF83DAA0E16548F9DF54D5845FEB7" box="[697,844,308,334]" country="Thailand" name="Phetchaburi" pageId="5">Phetchaburi</collectingRegion>
,
<geoCoordinate id="6AEF5018AA0E16548E45F54D5F00FEB6" box="[865,1033,308,335]" degrees="12" direction="north" minutes="50" orientation="latitude" pageId="5" precision="1" seconds="37.9" value="12.843861">12°50'37.9&quot;N</geoCoordinate>
;
<geoCoordinate id="6AEF5018AA0E1654893BF54D5FCAFEB6" box="[1055,1219,308,335]" degrees="99" direction="east" minutes="35" orientation="longitude" pageId="5" precision="1" seconds="27.2" value="99.59089">99°35'27.2&quot;E</geoCoordinate>
,
<date id="7B65101FAA0E165489FCF54D5E6DFEB6" box="[1240,1380,308,335]" pageId="5" value="2018-04-15">
<collectingDate id="6B21E9F7AA0E165489FCF54D5E6DFEB6" box="[1240,1380,308,335]" pageId="5" value="2018-04-15">15.IV.2018</collectingDate>
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,
<collectorName id="A22E5309AA0E16548DD3F5215A7FFE88" box="[247,374,343,370]" pageId="5">I. Dugdale</collectorName>
&amp;
<collectorName id="A22E5309AA0E16548CB9F521591DFE88" box="[413,532,343,370]" pageId="5">P. Phetsri</collectorName>
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.
</paragraph>
</subSubSection>
<subSubSection id="47C16554AA0E165B8DD3F5F15997FD27" lastPageId="10" pageId="5" type="description">
<paragraph id="0F6436DFAA0E16548DD3F5F15885FE58" blockId="5.[247,908,391,418]" box="[247,908,391,418]" pageId="5">
<smallCapsWord id="0982A003AA0E16548DD3F5F15A9EFE59" baselines="411,411" box="[247,407,392,418]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Description" pageId="5">DESCRIPTION</smallCapsWord>
.
<smallCapsWord id="0982A003AA0E16548C80F5F15AE4FE59" baselines="411,411" box="[420,493,392,418]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Male" pageId="5">MALE</smallCapsWord>
(
<typeStatus id="D060887DAA0E16548CD9F5FE5967FE58" box="[509,622,391,417]" pageId="5" type="holotype">holotype</typeStatus>
): (
<figureCitation id="97E02A5AAA0E16548FAEF5FE59DFFE58" box="[650,726,391,417]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="5">Figs 1</figureCitation>
A−B; 2 C−D).
</paragraph>
<caption id="5BA46657AA0E16548DD3F2A35F04F8F7" ID-DOI="http://doi.org/10.5281/zenodo.13271602" ID-Zenodo-Dep="13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="5" startId="5.[247,284,1754,1776]" targetBox="[254,1380,448,1725]" targetPageId="5" targetType="figure">
<paragraph id="0F6436DFAA0E16548DD3F2A35F04F8F7" blockId="5.[247,1386,1754,1806]" pageId="5">
Fig. 1.
<taxonomicName id="C8DB4D5CAA0E16548C61F2A3590AF916" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[325,515,1754,1775]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="5" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA0E16548C61F2A3590AF916" box="[325,515,1754,1775]" italics="true" pageId="5">Hierodula confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA0E16548F29F2A25954F909" box="[525,605,1755,1776]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view.
</paragraph>
</caption>
<paragraph id="0F6436DFAA0D16578DD3F4C25910FF2D" blockId="6.[247,537,186,213]" box="[247,537,186,213]" pageId="6">
<emphasis id="3DAFEACDAA0D16578DD3F4C25AAEFF2D" box="[247,423,187,212]" italics="true" pageId="6">Measurements</emphasis>
:
<tableCitation id="42590364AA0D16578C93F4C3591AFF2D" box="[439,531,186,212]" captionStart="Table 1" captionStartId="6.[247,304,246,268]" captionText="Table 1. Measurements of the genus diagnostic morphological parts." pageId="6">Table 1</tableCitation>
.
</paragraph>
<caption id="5BA46657AA0D16578DD3F48F58AEFEF2" box="[247,935,246,268]" pageId="6" startId="6.[247,304,246,268]" targetType="table">
<paragraph id="0F6436DFAA0D16578DD3F48F58AEFEF2" blockId="6.[247,935,246,268]" box="[247,935,246,268]" pageId="6">Table 1. Measurements of the genus diagnostic morphological parts.</paragraph>
</caption>
<paragraph id="0F6436DFAA0D16578C0DF55B58EBFB28" pageId="6">
<table id="7DDBC47FAA0DE9AE8C0DF55B58F6FB2B" box="[297,1023,290,1234]" gridcols="3" gridrows="22" pageId="6">
<tr id="B1EB349DAA0DE9AE8C0DF55B58F6FECE" box="[297,1023,290,311]" gridrow="0" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF55B5988FECE" box="[297,641,290,311]" gridcol="0" gridrow="0" pageId="6">
<emphasis id="3DAFEACDAA0D16578C0DF55B5ACFFECE" bold="true" box="[297,454,290,311]" pageId="6">Measurements</emphasis>
</th>
<th id="F23A5DE1AA0DE9AE8E21F55B585DFECE" box="[773,852,290,311]" gridcol="1" gridrow="0" pageId="6">
<emphasis id="3DAFEACDAA0D16578E22F55B5823FECE" bold="true" box="[774,810,290,311]" pageId="6">HT</emphasis>
(♂)
</th>
<th id="F23A5DE1AA0DE9AE8E8CF55B58F6FECE" box="[936,1023,290,311]" gridcol="2" gridrow="0" pageId="6">
<emphasis id="3DAFEACDAA0D16578E8CF55B58DCFECE" bold="true" box="[936,981,290,311]" pageId="6">PT1</emphasis>
(♀)
</th>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF53758F6FE9D" box="[297,1023,334,356]" gridrow="1" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF5375988FE9D" box="[297,641,334,356]" gridcol="0" gridrow="1" pageId="6">Total length</th>
<td id="F23A5DE1AA0DE9AE8E21F537585DFE9D" box="[773,852,334,356]" gridcol="1" gridrow="1" pageId="6">79.4</td>
<td id="F23A5DE1AA0DE9AE8E8CF53758F6FE9D" box="[936,1023,334,356]" gridcol="2" gridrow="1" pageId="6">91.9</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF50058F6FE76" box="[297,1023,377,399]" gridrow="2" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF5005988FE76" box="[297,641,377,399]" gridcol="0" gridrow="2" pageId="6">Head width</th>
<td id="F23A5DE1AA0DE9AE8E21F500585DFE76" box="[773,852,377,399]" gridcol="1" gridrow="2" pageId="6">8.6</td>
<td id="F23A5DE1AA0DE9AE8E8CF50058F6FE76" box="[936,1023,377,399]" gridcol="2" gridrow="2" pageId="6">11.4</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF5DC58F6FE42" box="[297,1023,421,443]" gridrow="3" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF5DC5988FE42" box="[297,641,421,443]" gridcol="0" gridrow="3" pageId="6">Head height</th>
<td id="F23A5DE1AA0DE9AE8E21F5DC585DFE42" box="[773,852,421,443]" gridcol="1" gridrow="3" pageId="6">6.2</td>
<td id="F23A5DE1AA0DE9AE8E8CF5DC58F6FE42" box="[936,1023,421,443]" gridcol="2" gridrow="3" pageId="6">8.8</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF5A858F6FE1E" box="[297,1023,465,487]" gridrow="4" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF5A85988FE1E" box="[297,641,465,487]" gridcol="0" gridrow="4" pageId="6">Pronotum length</th>
<td id="F23A5DE1AA0DE9AE8E21F5A8585DFE1E" box="[773,852,465,487]" gridcol="1" gridrow="4" pageId="6">24.1</td>
<td id="F23A5DE1AA0DE9AE8E8CF5A858F6FE1E" box="[936,1023,465,487]" gridcol="2" gridrow="4" pageId="6">30.5</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF58458F6FDEA" box="[297,1023,509,531]" gridrow="5" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF5845988FDEA" box="[297,641,509,531]" gridcol="0" gridrow="5" pageId="6">Pronotum width</th>
<td id="F23A5DE1AA0DE9AE8E21F584585DFDEA" box="[773,852,509,531]" gridcol="1" gridrow="5" pageId="6">5.9</td>
<td id="F23A5DE1AA0DE9AE8E8CF58458F6FDEA" box="[936,1023,509,531]" gridcol="2" gridrow="5" pageId="6">7.9</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF65058F6FDC6" box="[297,1023,553,575]" gridrow="6" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF6505988FDC6" box="[297,641,553,575]" gridcol="0" gridrow="6" pageId="6">Pronotum narrow width</th>
<td id="F23A5DE1AA0DE9AE8E21F650585DFDC6" box="[773,852,553,575]" gridcol="1" gridrow="6" pageId="6">3.6</td>
<td id="F23A5DE1AA0DE9AE8E8CF65058F6FDC6" box="[936,1023,553,575]" gridcol="2" gridrow="6" pageId="6">4.8</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF62C58F6FD92" box="[297,1023,597,619]" gridrow="7" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF62C5988FD92" box="[297,641,597,619]" gridcol="0" gridrow="7" pageId="6">Prozona length</th>
<td id="F23A5DE1AA0DE9AE8E21F62C585DFD92" box="[773,852,597,619]" gridcol="1" gridrow="7" pageId="6">6.0</td>
<td id="F23A5DE1AA0DE9AE8E8CF62C58F6FD92" box="[936,1023,597,619]" gridcol="2" gridrow="7" pageId="6">7.5</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF6F858F6FD6E" box="[297,1023,641,663]" gridrow="8" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF6F85988FD6E" box="[297,641,641,663]" gridcol="0" gridrow="8" pageId="6">Metazona length</th>
<td id="F23A5DE1AA0DE9AE8E21F6F8585DFD6E" box="[773,852,641,663]" gridcol="1" gridrow="8" pageId="6">18.1</td>
<td id="F23A5DE1AA0DE9AE8E8CF6F858F6FD6E" box="[936,1023,641,663]" gridcol="2" gridrow="8" pageId="6">23.0</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF6D458F6FD3A" box="[297,1023,685,707]" gridrow="9" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF6D45988FD3A" box="[297,641,685,707]" gridcol="0" gridrow="9" pageId="6">Tegmen length</th>
<td id="F23A5DE1AA0DE9AE8E21F6D4585DFD3A" box="[773,852,685,707]" gridcol="1" gridrow="9" pageId="6">52.2</td>
<td id="F23A5DE1AA0DE9AE8E8CF6D458F6FD3A" box="[936,1023,685,707]" gridcol="2" gridrow="9" pageId="6">55.4</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF6A158F6FD16" box="[297,1023,728,751]" gridrow="10" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF6A15988FD16" box="[297,641,728,751]" gridcol="0" gridrow="10" pageId="6">Ala length</th>
<td id="F23A5DE1AA0DE9AE8E21F6A1585DFD16" box="[773,852,728,751]" gridcol="1" gridrow="10" pageId="6">45.5</td>
<td id="F23A5DE1AA0DE9AE8E8CF6A158F6FD16" box="[936,1023,728,751]" gridcol="2" gridrow="10" pageId="6">47.6</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF77C58F6FCE2" box="[297,1023,773,795]" gridrow="11" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF77C5988FCE2" box="[297,641,773,795]" gridcol="0" gridrow="11" pageId="6">Procoxa length</th>
<td id="F23A5DE1AA0DE9AE8E21F77C585DFCE2" box="[773,852,773,795]" gridcol="1" gridrow="11" pageId="6">14.7</td>
<td id="F23A5DE1AA0DE9AE8E8CF77C58F6FCE2" box="[936,1023,773,795]" gridcol="2" gridrow="11" pageId="6">19.6</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF74858F6FCBE" box="[297,1023,817,839]" gridrow="12" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF7485988FCBE" box="[297,641,817,839]" gridcol="0" gridrow="12" pageId="6">Profemur length</th>
<td id="F23A5DE1AA0DE9AE8E21F748585DFCBE" box="[773,852,817,839]" gridcol="1" gridrow="12" pageId="6">17.0</td>
<td id="F23A5DE1AA0DE9AE8E8CF74858F6FCBE" box="[936,1023,817,839]" gridcol="2" gridrow="12" pageId="6">22.7</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF72558F6FC8B" box="[297,1023,860,882]" gridrow="13" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF7255988FC8B" box="[297,641,860,882]" gridcol="0" gridrow="13" pageId="6">Protibia length</th>
<td id="F23A5DE1AA0DE9AE8E21F725585DFC8B" box="[773,852,860,882]" gridcol="1" gridrow="13" pageId="6">10.6</td>
<td id="F23A5DE1AA0DE9AE8E8CF72558F6FC8B" box="[936,1023,860,882]" gridcol="2" gridrow="13" pageId="6">13.9</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF7F158F6FC67" box="[297,1023,904,926]" gridrow="14" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF7F15988FC67" box="[297,641,904,926]" gridcol="0" gridrow="14" pageId="6">Mesofemur length</th>
<td id="F23A5DE1AA0DE9AE8E21F7F1585DFC67" box="[773,852,904,926]" gridcol="1" gridrow="14" pageId="6">18.1</td>
<td id="F23A5DE1AA0DE9AE8E8CF7F158F6FC67" box="[936,1023,904,926]" gridcol="2" gridrow="14" pageId="6">22.5</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF7CD58F6FC33" box="[297,1023,948,970]" gridrow="15" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF7CD5988FC33" box="[297,641,948,970]" gridcol="0" gridrow="15" pageId="6">Mesotibia length</th>
<td id="F23A5DE1AA0DE9AE8E21F7CD585DFC33" box="[773,852,948,970]" gridcol="1" gridrow="15" pageId="6">14.4</td>
<td id="F23A5DE1AA0DE9AE8E8CF7CD58F6FC33" box="[936,1023,948,970]" gridcol="2" gridrow="15" pageId="6">19.7</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF79958F6FC0F" box="[297,1023,992,1014]" gridrow="16" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF7995988FC0F" box="[297,641,992,1014]" gridcol="0" gridrow="16" pageId="6">Metafemur length</th>
<td id="F23A5DE1AA0DE9AE8E21F799585DFC0F" box="[773,852,992,1014]" gridcol="1" gridrow="16" pageId="6">21.9</td>
<td id="F23A5DE1AA0DE9AE8E8CF79958F6FC0F" box="[936,1023,992,1014]" gridcol="2" gridrow="16" pageId="6">26.7</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF07558F6FBDB" box="[297,1023,1036,1058]" gridrow="17" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF0755988FBDB" box="[297,641,1036,1058]" gridcol="0" gridrow="17" pageId="6">Metatibia length</th>
<td id="F23A5DE1AA0DE9AE8E21F075585DFBDB" box="[773,852,1036,1058]" gridcol="1" gridrow="17" pageId="6">22.4</td>
<td id="F23A5DE1AA0DE9AE8E8CF07558F6FBDB" box="[936,1023,1036,1058]" gridcol="2" gridrow="17" pageId="6">30.9</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF04158F6FBB7" box="[297,1023,1080,1102]" gridrow="18" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF0415988FBB7" box="[297,641,1080,1102]" gridcol="0" gridrow="18" pageId="6">Metatarsus length</th>
<td id="F23A5DE1AA0DE9AE8E21F041585DFBB7" box="[773,852,1080,1102]" gridcol="1" gridrow="18" pageId="6">6.0</td>
<td id="F23A5DE1AA0DE9AE8E8CF04158F6FBB7" box="[936,1023,1080,1102]" gridcol="2" gridrow="18" pageId="6">7.2</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF01D58F6FB83" box="[297,1023,1124,1146]" gridrow="19" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF01D5988FB83" box="[297,641,1124,1146]" gridcol="0" gridrow="19" pageId="6">Anteroventral femoral spine count</th>
<td id="F23A5DE1AA0DE9AE8E21F01D585DFB83" box="[773,852,1124,1146]" gridcol="1" gridrow="19" pageId="6">15</td>
<td id="F23A5DE1AA0DE9AE8E8CF01D58F6FB83" box="[936,1023,1124,1146]" gridcol="2" gridrow="19" pageId="6">15</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF0E958F6FB5F" box="[297,1023,1168,1190]" gridrow="20" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF0E95988FB5F" box="[297,641,1168,1190]" gridcol="0" gridrow="20" pageId="6">Anteroventral tibial spine count</th>
<td id="F23A5DE1AA0DE9AE8E21F0E9585DFB5F" box="[773,852,1168,1190]" gridcol="1" gridrow="20" pageId="6">14/15</td>
<td id="F23A5DE1AA0DE9AE8E8CF0E958F6FB5F" box="[936,1023,1168,1190]" gridcol="2" gridrow="20" pageId="6">14</td>
</tr>
<tr id="B1EB349DAA0DE9AE8C0DF0C558F6FB2B" box="[297,1023,1212,1234]" gridrow="21" pageId="6">
<th id="F23A5DE1AA0DE9AE8C0DF0C55988FB2B" box="[297,641,1212,1234]" gridcol="0" gridrow="21" pageId="6">Posteroventral tibial spine count</th>
<td id="F23A5DE1AA0DE9AE8E21F0C5585DFB2B" box="[773,852,1212,1234]" gridcol="1" gridrow="21" pageId="6">9/11</td>
<td id="F23A5DE1AA0DE9AE8E8CF0C558F6FB2B" box="[936,1023,1212,1234]" gridcol="2" gridrow="21" pageId="6">10/11</td>
</tr>
</table>
</paragraph>
<paragraph id="0F6436DFAA0D16578DD3F1755905FADC" blockId="6.[247,524,1291,1318]" box="[247,524,1291,1318]" pageId="6">
<emphasis id="3DAFEACDAA0D16578DD3F1755A4DFADC" box="[247,324,1292,1317]" italics="true" pageId="6">Ratios</emphasis>
<specimenCount id="19DDFD56AA0D16578C68F1755A6FFADF" box="[332,358,1292,1318]" count="1" pageId="6" type="male">♂</specimenCount>
: MzL/PzL: 3.
</paragraph>
<paragraph id="0F6436DFAA0D16578DD3F142595BFA18" blockId="6.[247,1386,1339,1505]" pageId="6">
<emphasis id="3DAFEACDAA0D16578DD3F1425A31FAAC" box="[247,312,1339,1365]" italics="true" pageId="6">Head</emphasis>
(
<figureCitation id="97E02A5AAA0D16578C6FF1425A9EFAAC" box="[331,407,1339,1365]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="6">Figs 1</figureCitation>
A−B; 2 C): Wider than long with large, rounded compound eyes. Vertex flat. Ocellar tubercle not elevated, in same plane as frons, without protruding ridges. Ocelli relatively large. Lower frons transverse with external margins posteriorly and laterally, no margin anteriorly, posterior margin almost flat, with two faintly defined vertical ridges internally. Clypeus and labrum smooth.
</paragraph>
<paragraph id="0F6436DFAA0D16578DD3F18E5FC6F91D" blockId="6.[247,1385,1527,1764]" pageId="6">
<emphasis id="3DAFEACDAA0D16578DD3F18E5A47F9E8" box="[247,334,1527,1553]" italics="true" pageId="6">Thorax</emphasis>
(
<figureCitation id="97E02A5AAA0D16578C44F18E5AA7F9E8" box="[352,430,1527,1553]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="6">Figs 1</figureCitation>
A−B; 2 D): Pronotum long, broadest point at level of procoxal insertion site. Lateral margins gently sinusoidal in prozona and anterior half of metazona, parallel sided in posterior half of metazona. Margins smooth, without denticulations or projections, flattened along the edges, in particular posteriorly from the widest point until half the metazona, yet not wide enough to be considered as a distinct lateral expansion of pronotum. Dorsal surface entirely smooth. Cervix with strongly sclerotized lateral and intercervical sclerites, touching in middle. Postcervical plate and posterior ventral part of pronotum entirely smooth.
</paragraph>
<paragraph id="0F6436DFAA0D16578DD3F2805E60F884" blockId="6.[247,1386,1785,1917]" pageId="6">
<emphasis id="3DAFEACDAA0D16578DD3F2805ACBF8EA" box="[247,450,1785,1811]" italics="true" pageId="6">Prothoracic legs</emphasis>
(
<figureCitation id="97E02A5AAA0D16578CF4F2805914F8EA" box="[464,541,1785,1811]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="6">Figs 1</figureCitation>
A−B; 2 E): Coxa with 10−16 small and very weakly developed blunt spinules of more or less equal size with same colour as coxa (amount and size of spinules can strongly vary between individuals). Coxal anterior surface completely covered with irregularly interspaced small whitish tubercles of roughly same size as spinules on dorsal edge. Dorsal and
</paragraph>
<paragraph id="0F6436DFAA0C16568DD3F4C75FE9FDA2" blockId="7.[247,1386,190,603]" pageId="7">
ventral coxal lobes rounded, equal in shape, length and adjacent, both lobes equally as broad. Femur with straight dorsal margin, surface entirely smooth. Femoral brush ellipse-shaped, starting from 12
<superScript id="F8AE9B97AA0C16568C9FF4865AC2FEEB" attach="left" box="[443,459,255,274]" fontSize="8" pageId="7">th</superScript>
AvS and ending at 15
<superScript id="F8AE9B97AA0C16568FFFF48659E2FEEB" attach="left" box="[731,747,255,274]" fontSize="8" pageId="7">th</superScript>
AvS. Genicular lobes large and rounded, with small genicular spur. Colouration of prothoracic legs entirely uniform to body colour. Tibial spine groove placed slightly proximally from middle of femur. AvS unequal in size and arranged as follows:
<emphasis id="3DAFEACDAA0C16568C40F5175AF0FE71" bold="true" box="[356,505,366,392]" pageId="7">iIiIiIiIiIiIiiI</emphasis>
. AvS 12 and 15 larger but with 2 smaller spines of equal size between them. All spines apically infuscate only. PvS 4 smaller than others, others about equal in size; all slightly apically infuscate. DS 2 and 4 more or less equal in size, DS 2 longest and DS1 smallest. Tibial AvS gradually elongated towards tibial spur, all concolour to tibia and slightly apically infuscate. PvS slightly more procumbent, spines gradually longer towards distal end. Tibial spur apically infuscate. Protarsi with dark spot at distal end; first tarsomere longer than others combined. Spinal formula: F = 4DS/15AvS/4PvS; T = 14−15AvS/9−11PvS.
</paragraph>
<caption id="5BA46657AA0C16568DD3F2175A4EF926" ID-DOI="http://doi.org/10.5281/zenodo.13271604" ID-Zenodo-Dep="13271604" httpUri="https://zenodo.org/record/13271604/files/figure.png" pageId="7" startId="7.[247,284,1646,1668]" targetBox="[279,1385,629,1625]" targetPageId="7" targetType="figure">
<paragraph id="0F6436DFAA0C16568DD3F2175A4EF926" blockId="7.[247,1386,1646,1759]" pageId="7">
Fig. 2.
<taxonomicName id="C8DB4D5CAA0C16568C1AF2175AF0F97A" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[318,505,1646,1667]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA0C16568C1AF2175AF0F97A" box="[318,505,1646,1667]" italics="true" pageId="7">Hierodula confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA0C16568CDBF2165946F97D" box="[511,591,1647,1668]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
Paratype ♀ [PT1] (THNHM). A, head, frontal view. B, pronotum, dorsal view. Holotype ♂ (THNHM). C, head, frontal view. D, pronotum, dorsal view. Live specimens. E, detail of prothoracic legs. F, dentral view of nymph abdomen with visible intersegmental colouration, typically shown during deimatic display.
</paragraph>
</caption>
<paragraph id="0F6436DFAA0316598DD3F4C25F90FE7D" blockId="8.[247,1386,186,388]" pageId="8">
<emphasis id="3DAFEACDAA0316598DD3F4C2595BFF2D" box="[247,594,186,212]" italics="true" pageId="8">Meso- and metathoracic legs</emphasis>
(
<figureCitation id="97E02A5AAA0316598F44F4C35996FF2D" box="[608,671,186,212]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="8">Fig 1</figureCitation>
A−B): Long and slender, without dilatations or projections. Femora with rounded genicular lobes and a short genicular spur. Tibiae with two apical spurs. Tarsi 5-segmented, concolour and slightly darkened apically on ventral surface; 3
<superScript id="F8AE9B97AA03165989D3F4835E01FEF4" attach="left" box="[1271,1288,250,269]" fontSize="8" pageId="8">rd</superScript>
and 4
<superScript id="F8AE9B97AA031659887EF4835E63FEF4" attach="none" box="[1370,1386,250,269]" fontSize="8" pageId="8">th</superScript>
tarsomere with small black spot proximally near the joint with previous tarsomere. First tarsomere of mesotarsus slightly shorter than remaining segments combined, first tarsomere of metatarsus about equal in length compared to remaining segments combined.
</paragraph>
<caption id="5BA46657AA0316598DD3F00D5F38FB51" ID-DOI="http://doi.org/10.5281/zenodo.13271606" ID-Zenodo-Dep="13271606" httpUri="https://zenodo.org/record/13271606/files/figure.png" pageId="8" startId="8.[247,284,1140,1162]" targetBox="[247,1385,409,1120]" targetPageId="8" targetType="figure">
<paragraph id="0F6436DFAA0316598DD3F00D5F38FB51" blockId="8.[247,1385,1140,1192]" pageId="8">
Fig. 3.
<taxonomicName id="C8DB4D5CAA0316598C18F00D5AFEFB70" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[316,503,1140,1161]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="8" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA0316598C18F00D5AFEFB70" box="[316,503,1140,1161]" italics="true" pageId="8">Hierodula confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA0316598CD8F00C5944FB73" box="[508,589,1141,1162]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
, live photographs: A−B, paratype ♀ [PT1]. C, sub-adult female nymph. D, adult male at light trap, Baan Maka Nature Resort, Thailand, © I. Dugdale &amp; P. Phetsri.
</paragraph>
</caption>
<paragraph id="0F6436DFAA0316598DD3F0BF59E5FABE" blockId="8.[247,1385,1221,1351]" pageId="8">
<emphasis id="3DAFEACDAA0316598DD3F0BF5ADAFB26" box="[247,467,1221,1247]" italics="true" pageId="8">Tegmina and alae</emphasis>
(
<figureCitation id="97E02A5AAA0316598CC6F0BC592DFB26" box="[482,548,1221,1247]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="8">Fig 1</figureCitation>
A−B): Tegmina uniformly concolour to the body (usually green) on costal area, hyaline in discoidal area, longer than tip of abdomen when folded in rest. Veins green, stigma white with strong hue of yellowish-green. Stigma located proximally from middle of tegmen. Alae fully developed, hyaline.
</paragraph>
<paragraph id="0F6436DFAA0316598DD3F12558D1FA8F" blockId="8.[247,984,1372,1398]" box="[247,984,1372,1398]" pageId="8">
<emphasis id="3DAFEACDAA0316598DD3F1255A60FA8F" box="[247,361,1372,1398]" italics="true" pageId="8">Abdomen</emphasis>
(
<figureCitation id="97E02A5AAA0316598C5CF1255AB7FA8F" box="[376,446,1372,1398]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="8">Fig. 1</figureCitation>
A−B): Fusiform. Cerci setose, not flattened.
</paragraph>
<caption id="5BA46657AA0316598DD3F32D5FDDF871" ID-DOI="http://doi.org/10.5281/zenodo.13271608" ID-Zenodo-Dep="13271608" httpUri="https://zenodo.org/record/13271608/files/figure.png" pageId="8" startId="8.[247,284,1876,1898]" targetBox="[266,1367,1421,1845]" targetPageId="8" targetType="figure">
<paragraph id="0F6436DFAA0316598DD3F32D5FDDF871" blockId="8.[247,1385,1876,1928]" pageId="8">
Fig. 4.
<taxonomicName id="C8DB4D5CAA0316598C67F32D5909F890" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[323,512,1876,1897]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="8" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA0316598C67F32D5909F890" box="[323,512,1876,1897]" italics="true" pageId="8">Hierodula confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA0316598F2DF32D5955F890" box="[521,604,1876,1897]" pageId="8" rank="species">sp. nov.</taxonomicNameLabel>
, genitalia prep. Vermeersch GEN1804. A, left phallic complex. B, detail of anterior process of left phallomere. C, right phallic complex. D, coxosternite IX (subgenital plate).
</paragraph>
</caption>
<paragraph id="0F6436DFAA031658887EF3B45801FDEC" blockId="8.[1370,1385,1997,2023]" lastBlockId="9.[247,1386,190,533]" lastPageId="9" pageId="8">
9
<emphasis id="3DAFEACDAA0216588DD3F4C75A60FF21" box="[247,361,190,216]" italics="true" pageId="9">Genitalia</emphasis>
(
<figureCitation id="97E02A5AAA0216588C5EF4C75AC2FF20" box="[378,459,190,217]" captionStart="Fig" captionStartId="8.[247,284,1876,1898]" captionTargetBox="[266,1367,1421,1845]" captionTargetId="figure-230@8.[247,1385,1411,1854]" captionTargetPageId="8" captionText="Fig. 4. Hierodula confusa sp. nov., genitalia prep. Vermeersch GEN1804. A, left phallic complex. B, detail of anterior process of left phallomere. C, right phallic complex. D, coxosternite IX (subgenital plate)." figureDoi="http://doi.org/10.5281/zenodo.13271608" httpUri="https://zenodo.org/record/13271608/files/figure.png" pageId="9">Figs 4</figureCitation>
;
<figureCitation id="97E02A5AAA0216588CFEF4C75AE0FF20" box="[474,489,190,217]" captionStart="Fig" captionStartId="9.[247,284,1757,1779]" captionTargetBox="[270,1355,573,1708]" captionTargetId="figure-263@9.[247,1385,559,1737]" captionTargetPageId="9" captionText="Fig. 5. Hierodula membranacea, genitalia ♂: A, anterior process of left phallomere, dorsal view. B, distal part of left phallic complex, dorsal view. C, distal part of right phallic complex, ventral view. Hierodula confusa sp. nov., genitalia ♂: D, anterior process of left phallomere, dorsal view. E, distal part of left phallic complex, dorsal view. F, distal part of right phallic complex, ventral view. Rhombodera extensicollis, genitalia ♂: G, anterior process of left phallomere, dorsal view. H, distal part of left phallic complex, dorsal view. I, distal part of right phallic complex, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13271610" httpUri="https://zenodo.org/record/13271610/files/figure.png" pageId="9">5</figureCitation>
D−F): Ventral phallomere of left phallic complex elongated, longer than broad. At about half its length the distal process bifurcates into a very small medial process (sdpm) and a long more or less straight lateral process (sdpl) with acute apex. Apical process of left phallomere (paa) broad and flattened. Phalloid apophysis (afa) with a strongly sclerotised and spine-shaped anterior process (aafa). Posterior process (pafa) irregularly shaped, wrinkled and membranous, completely and densely covered with very small and short spine-like setae. Right phallomere with main posterior lobe fda triangular but rather elongated (longer than broad), sclerite R3 shovel-shaped; very wide at its anterior end, pia and pva both strongly sclerified. Coxosternite IX (subgenital plate) smooth, with small hair-like setae near the edges, without spikes or other sclerified structures.
</paragraph>
<caption id="5BA46657AA0216588DD3F2A45AD7F873" ID-DOI="http://doi.org/10.5281/zenodo.13271610" ID-Zenodo-Dep="13271610" httpUri="https://zenodo.org/record/13271610/files/figure.png" pageId="9" startId="9.[247,284,1757,1779]" targetBox="[270,1355,573,1708]" targetPageId="9" targetType="figure">
<paragraph id="0F6436DFAA0216588DD3F2A45AD7F873" blockId="9.[247,1386,1757,1930]" pageId="9">
Fig. 5.
<taxonomicName id="C8DB4D5CAA0216588C1BF2A45932F90B" authorityName="Burmeister" authorityYear="1838" box="[319,571,1757,1778]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="membranacea">
<emphasis id="3DAFEACDAA0216588C1BF2A45932F90B" box="[319,571,1757,1778]" italics="true" pageId="9">Hierodula membranacea</emphasis>
</taxonomicName>
, genitalia ♂: A, anterior process of left phallomere, dorsal view. B, distal part of left phallic complex, dorsal view. C, distal part of right phallic complex, ventral view.
<taxonomicName id="C8DB4D5CAA0216588970F2825E06F8E9" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[1108,1295,1787,1808]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA0216588970F2825E06F8E9" box="[1108,1295,1787,1808]" italics="true" pageId="9">Hierodula confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA0216588830F2855E6FF8E8" box="[1300,1382,1788,1809]" pageId="9" rank="species">sp. nov.</taxonomicNameLabel>
, genitalia ♂: D, anterior process of left phallomere, dorsal view. E, distal part of left phallic complex, dorsal view. F, distal part of right phallic complex, ventral view.
<taxonomicName id="C8DB4D5CAA0216588E27F34E5F0EF8B5" authorityName="Serville" authorityYear="1839" box="[771,1031,1847,1868]" class="Insecta" family="Mantidae" genus="Rhombodera" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="extensicollis">
<emphasis id="3DAFEACDAA0216588E27F34E5F0EF8B5" box="[771,1031,1847,1868]" italics="true" pageId="9">Rhombodera extensicollis</emphasis>
</taxonomicName>
, genitalia ♂: G, anterior process of left phallomere, dorsal view. H, distal part of left phallic complex, dorsal view. I, distal part of right phallic complex, ventral view.
</paragraph>
</caption>
<paragraph id="0F6436DFAA01165B8DD3F4C35809FEE2" blockId="10.[247,1385,186,283]" pageId="10">
<emphasis id="3DAFEACDAA01165B8DD3F4C35A83FF2D" box="[247,394,186,212]" italics="true" pageId="10">Colouration</emphasis>
(
<figureCitation id="97E02A5AAA01165B8CBFF4C35AE5FF2D" box="[411,492,186,212]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="10">Figs 1</figureCitation>
;
<figureCitation id="97E02A5AAA01165B8CDAF4C35902FF2D" box="[510,523,186,212]" captionStart="Fig" captionStartId="8.[247,284,1140,1162]" captionTargetBox="[247,1385,409,1120]" captionTargetId="figure-226@8.[247,1385,409,1120]" captionTargetPageId="8" captionText="Fig. 3. Hierodula confusa sp. nov., live photographs: A−B, paratype ♀ [PT1]. C, sub-adult female nymph. D, adult male at light trap, Baan Maka Nature Resort, Thailand, © I. Dugdale &amp; P. Phetsri." figureDoi="http://doi.org/10.5281/zenodo.13271606" httpUri="https://zenodo.org/record/13271606/files/figure.png" pageId="10">3</figureCitation>
A−C): All currently known specimens of
<taxonomicName id="C8DB4D5CAA01165B893CF4C35FF5FF2D" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[1048,1276,186,212]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="10" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA01165B893CF4C35FF5FF2D" box="[1048,1276,186,212]" italics="true" pageId="10">Hierodula confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA01165B8823F4C25E60FF2C" box="[1287,1385,187,213]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
feature a uniformly green colouration, although it is likely that brown individuals exist as is occasionally observed in
<taxonomicName id="C8DB4D5CAA01165B8F0CF57859F1FEE3" authorityName="Burmeister" authorityYear="1838" box="[552,760,256,282]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="10" phylum="Arthropoda" rank="species" species="membranacea">
<emphasis id="3DAFEACDAA01165B8F0CF57859F1FEE3" box="[552,760,256,282]" italics="true" pageId="10">H. membranacea</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="0F6436DFAA01165B8DD3F54958B3FEB3" blockId="10.[247,954,304,330]" box="[247,954,304,330]" pageId="10">
<smallCapsWord id="0982A003AA01165B8DD3F5495A53FEB0" baselines="323,324" box="[247,346,304,330]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Female" pageId="10">FEMALE</smallCapsWord>
(
<typeStatus id="D060887DAA01165B8C4CF5495AD1FEB3" box="[360,472,304,330]" pageId="10" type="paratype">Paratype</typeStatus>
: PT1): (
<figureCitation id="97E02A5AAA01165B8F1CF549598DFEB3" box="[568,644,304,330]" captionStart="Fig" captionStartId="5.[247,284,1754,1776]" captionTargetBox="[254,1380,448,1725]" captionTargetId="figure-109@5.[247,1385,441,1733]" captionTargetPageId="5" captionText="Fig. 1. Hierodula confusa sp. nov. Holotype ♂ (THNHM). A, habitus, ventral view. B, habitus, dorsal view. Paratype ♀ [PT1] (THNHM) C, habitus, ventral view. D, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13271602" httpUri="https://zenodo.org/record/13271602/files/figure.png" pageId="10">Figs 1</figureCitation>
C−D; 2 A−B, F; 3 A−C).
</paragraph>
<paragraph id="0F6436DFAA01165B8DD3F519583FFE83" blockId="10.[247,822,351,378]" box="[247,822,351,378]" pageId="10">
<emphasis id="3DAFEACDAA01165B8DD3F5195AAEFE80" box="[247,423,352,377]" italics="true" pageId="10">Measurements</emphasis>
:
<tableCitation id="42590364AA01165B8C93F526591AFE83" box="[439,531,351,378]" captionStart="Table 1" captionStartId="6.[247,304,246,268]" captionText="Table 1. Measurements of the genus diagnostic morphological parts." pageId="10">Table 1</tableCitation>
.
<emphasis id="3DAFEACDAA01165B8F05F5195967FE80" box="[545,622,352,377]" italics="true" pageId="10">Ratios</emphasis>
<specimenCount id="19DDFD56AA01165B8F51F5195987FE83" box="[629,654,352,378]" count="1" pageId="10" type="female">♀</specimenCount>
: MzL/PzL: 3.
</paragraph>
<paragraph id="0F6436DFAA01165B8DD3F5E95F77FE50" blockId="10.[247,1150,399,426]" box="[247,1150,399,426]" pageId="10">Very similar to male but more robust, different in the following characters:</paragraph>
<paragraph id="0F6436DFAA01165B8DD3F5C65A6AFE05" blockId="10.[247,1385,447,508]" pageId="10">
(1) More acute spinules on dorsal margin of procoxae (approximately
<quantity id="C8239B3AAA01165B8910F5C65F94FE20" box="[1076,1181,447,473]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.588" metricValueMin="5.08" pageId="10" unit="in" value="21.0" valueMax="22.0" valueMin="20.0">20−22 in</quantity>
female vs
<quantity id="C8239B3AAA01165B8838F5C65A07FE05" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="4.064" metricValueMin="2.54" pageId="10" unit="in" value="13.0" valueMax="16.0" valueMin="10.0">10−16 in</quantity>
male);
</paragraph>
<paragraph id="0F6436DFAA01165B8DD3F66B5F98FDB6" blockId="10.[247,1385,530,591]" pageId="10">(2) Pronotum more robust, finely denticulated in prozona and anterior part of metazona, denticulations fading away distally, smooth edges in distal part of metazona;</paragraph>
<paragraph id="0F6436DFAA01165B8DD3F61D58BCFD87" blockId="10.[247,949,612,639]" box="[247,949,612,639]" pageId="10">(3) Tegmina completely opaque, reaching tip of abdomen;</paragraph>
<paragraph id="0F6436DFAA01165B8DD3F6ED59FBFD57" blockId="10.[247,754,660,687]" box="[247,754,660,687]" pageId="10">(4) Costal area of forewing wider distally;</paragraph>
<paragraph id="0F6436DFAA01165B8DD3F6BD5997FD27" blockId="10.[247,670,708,735]" box="[247,670,708,735]" pageId="10">(5) Abdomen broad and ovaliform.</paragraph>
</subSubSection>
<subSubSection id="47C16554AA01165B8DD3F68D59CAFCC9" pageId="10" type="distribution">
<paragraph id="0F6436DFAA01165B8DD3F68D59CAFCC9" blockId="10.[247,1380,755,817]" pageId="10">
<smallCapsWord id="0982A003AA01165B8DD3F68D5AAAFCF5" baselines="775,775" box="[247,419,756,782]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Distribution" pageId="10">DISTRIBUTION</smallCapsWord>
. The distribution of
<taxonomicName id="C8DB4D5CAA01165B8FBDF68A5872FCF4" authorityName="Vermeersch &amp; Unnahachote" authorityYear="1877" box="[665,891,755,781]" class="Insecta" family="Mantidae" genus="Hierodula" kingdom="Animalia" order="Mantodea" pageId="10" phylum="Arthropoda" rank="species" species="confusa" status="sp. nov.">
<emphasis id="3DAFEACDAA01165B8FBDF68A5872FCF4" box="[665,891,755,781]" italics="true" pageId="10">Hierodula confusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="269C57B6AA01165B8EA6F68D58E9FCF7" box="[898,992,756,782]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
as currently confirmed includes
<collectingCountry id="77CC764FAA01165B8DD3F76E5A6BFCC8" box="[247,354,791,817]" name="Thailand" pageId="10">Thailand</collectingCountry>
and Southern
<collectingCountry id="77CC764FAA01165B8F35F76E5950FCC8" box="[529,601,791,817]" name="China" pageId="10">China</collectingCountry>
(
<figureCitation id="97E02A5AAA01165B8F4DF76E59BDFCC9" box="[617,692,790,817]" captionStart="Fig" captionStartId="10.[247,284,1894,1916]" captionText="Fig. 6. Distribution map of Hierodula confusa sp. nov." pageId="10">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>