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<document id="D583F25C5495989981C420ADC962366C" ID-DOI="10.5252/geodiversitas2024v46a12" ID-ISSN="1638-9395" ID-Zenodo-Dep="13691736" ID-ZooBank="urn:lsid:zoobank.org:pub:02D31D6C-15C2-498A-A6C6-62A0163814C8" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1725524660885" checkinUser="plazi" docAuthor="Bakhouch, Asmaa El, Kerner, Adeline, Azizi, Abdelfattah, Debrenne, Françoise, Jalil, Nour-Eddine, Hafid, Ahmid &amp; Hariri, Khadija El" docDate="2024" docId="AF1FAC31FFE4FF8EFEA4F95E5AE5F929" docLanguage="en" docName="geodiversitas.2024.46.12.pdf" docOrigin="Geodiversitas 46 (12)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a12.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Fouanoucyathus tafraoutiensis El Bakhouch &amp; Kerner 2024, n. gen., n. sp." docType="treatment" docUuid="B5367302-7B81-4539-A744-818C17549559" docUuidSource="ZooBank" docVersion="1" lastPageNumber="449" masterDocId="5326D449FFE2FF88FFB8FFA95905FFC5" masterDocTitle="New archaeocyath genus from the early Cambrian of the western Anti-Atlas, Morocco" masterLastPageNumber="455" masterPageNumber="445" pageNumber="449" updateTime="1726061974691" updateUser="jonas" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="AD54F5A68888A9599BE53C772D1F0345">New archaeocyath genus from the early Cambrian of the western Anti-Atlas, Morocco</mods:title>
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<mods:namePart id="9DDA09ED5E3029525E00F43903BEB053">Bakhouch, Asmaa El</mods:namePart>
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<mods:namePart id="C54708B98F07FC5005A2EECC27240C3C">Kerner, Adeline</mods:namePart>
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<taxonomicName id="E0B666A4FFE4FF8EFEA4F95E58F0F8F5" authority="El Bakhouch &amp; Kerner" authorityName="El Bakhouch &amp; Kerner" authorityYear="2024" family="Carinacyathidae" genus="Fouanoucyathus" kingdom="Animalia" order="Ajacicyathida" pageId="6" pageNumber="449" phylum="Porifera" rank="species" species="tafraoutiensis" status="n. gen., n. sp.">
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El Bakhouch &amp; Kerner
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,
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(
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)
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. — Named after Tafraout, the city from where the specimens were collected.
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. — Tafraout,
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.
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. — Early Cambrian (Cambrian age 4 or Botoman).
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. —
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<collectingCountry id="5FA15DB7FFE4FF8EFC4FFD785D57FD2C" box="[1015,1106,721,745]" name="Morocco" pageId="6" pageNumber="449">Morocco</collectingCountry>
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<materialsCitation id="97DE177AFFE4FF8EFBD4FD785D01FCDB" country="Morocco" county="Issafen Formation" location="western Anti-Atlas" municipality="Tata Group" pageId="6" pageNumber="449" specimenCode="AA-FOU-CI-2" specimenCount="1" typeStatus="holotype">
<specimenCount id="31B0D6AEFFE4FF8EFBD4FD785DE7FD2C" box="[1132,1250,721,745]" count="1" pageId="6" pageNumber="449" type="generic">1 specimen</specimenCount>
;
<location id="22694BFCFFE4FF8EFB55FD7B5CAEFD2C" LSID="urn:lsid:plazi:treatment:AF1FAC31FFE4FF8EFEA4F95E5AE5F929:22694BFCFFE4FF8EFB55FD7B5CAEFD2C" box="[1261,1451,721,745]" country="Morocco" county="Issafen Formation" municipality="Tata Group" name="western Anti-Atlas" pageId="6" pageNumber="449">western Anti-Atlas</location>
,
<collectingMunicipality id="C76D875DFFE4FF8EFC96FD455AA0FCC1" box="[814,933,748,772]" pageId="6" pageNumber="449">Tata Group</collectingMunicipality>
,
<collectingCounty id="CE6865ABFFE4FF8EFC17FD455D6DFCC1" box="[943,1128,748,772]" pageId="6" pageNumber="449">Issafen Formation</collectingCounty>
,
<location id="22694BFCFFE4FF8EFBC9FD455DCFFCC1" LSID="urn:lsid:plazi:treatment:AF1FAC31FFE4FF8EFEA4F95E5AE5F929:22694BFCFFE4FF8EFBC9FD455DCFFCC1" box="[1137,1226,748,772]" county="Issafen Formation" municipality="Tata Group" name="Tafraout" pageId="6" pageNumber="449">Tafraout</location>
, section FO.IS8;
<typeStatus id="F80DA385FFE4FF8EFAC3FD455A58FCDA" pageId="6" pageNumber="449" type="holotype">holotype</typeStatus>
:
<specimenCode id="7710B55CFFE4FF8EFCD1FCAF5D01FCDB" box="[873,1028,774,798]" pageId="6" pageNumber="449">AA-FOU-CI-2</specimenCode>
</materialsCitation>
<materialsCitation id="97DE177AFFE4FF8EFB99FCAF5D81FCAB" country="Morocco" location="western Anti-Atlas" municipality="Tata Group" pageId="6" pageNumber="449" specimenCode="AA-FOU-CI-1, AA-FOU-CI-3, AA- FOU-CI-4" specimenCount="3" stateProvince="Issafen Formation" typeStatus="paratype">
<specimenCount id="31B0D6AEFFE4FF8EFB99FCAF5DA1FCDB" box="[1057,1188,774,798]" count="3" pageId="6" pageNumber="449" type="generic">3 specimens</specimenCount>
;
<location id="22694BFCFFE4FF8EFB09FCAE5C72FCDB" LSID="urn:lsid:plazi:treatment:AF1FAC31FFE4FF8EFEA4F95E5AE5F929:22694BFCFFE4FF8EFB09FCAE5C72FCDB" box="[1201,1399,774,798]" country="Morocco" municipality="Tata Group" name="western Anti-Atlas" pageId="6" pageNumber="449" stateProvince="Issafen Formation">western Anti-Atlas</location>
,
<collectingMunicipality id="C76D875DFFE4FF8EFA3AFCAE5A77FCFC" pageId="6" pageNumber="449">Tata Group</collectingMunicipality>
,
<collectingRegion id="E572D3C5FFE4FF8EFCC6FC885D3CFCFC" box="[894,1081,801,825]" pageId="6" pageNumber="449">Issafen Formation</collectingRegion>
,
<location id="22694BFCFFE4FF8EFBFCFC885D98FCFC" LSID="urn:lsid:plazi:treatment:AF1FAC31FFE4FF8EFEA4F95E5AE5F929:22694BFCFFE4FF8EFBFCFC885D98FCFC" box="[1092,1181,801,825]" municipality="Tata Group" name="Tafraout" pageId="6" pageNumber="449" stateProvince="Issafen Formation">Tafraout</location>
;
<typeStatus id="F80DA385FFE4FF8EFB11FC885C09FCFC" box="[1193,1292,801,825]" pageId="6" pageNumber="449" type="paratype">paratypes</typeStatus>
:
<specimenCode id="7710B55CFFE4FF8EFAA0FC885CAAFCFC" box="[1304,1455,801,825]" pageId="6" pageNumber="449">AA-FOU-CI-1</specimenCode>
from section FO.IS4;
<specimenCode id="7710B55CFFE4FF8EFBBFFC925D9BFC96" box="[1031,1182,827,851]" pageId="6" pageNumber="449">AA-FOU-CI-3</specimenCode>
from section FO.IS10;
<specimenCode id="7710B55CFFE4FF8EFA31FC925A9EFCAB" pageId="6" pageNumber="449">AA-FOU-CI-4</specimenCode>
from section FO.IS24
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6FAC4EACFFE4FF8EFC95FC2F5D70FC58" box="[813,1141,901,925]" pageId="6" pageNumber="449" type="description">
<paragraph id="27091D27FFE4FF8EFC95FC2F5D70FC58" blockId="6.[813,1141,901,925]" box="[813,1141,901,925]" pageId="6" pageNumber="449">
<smallCapsWord id="21EF8BFBFFE4FF8EFC95FC2F5ACAFC59" baselines="919,920" box="[813,975,902,925]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Measurements" pageId="6" pageNumber="449">MEASUREMENTS</smallCapsWord>
. — See
<tableCitation id="6A34289CFFE4FF8EFB9CFC2C5D70FC58" box="[1060,1141,901,925]" captionStart="TABLE" captionStartId="8.[388,399,1677,1694]" captionTargetPageId="8" captionText="TABLE 2. — Measurements of Fouanoucyathus tafraoutiensis El Bakhouch &amp; Kerner, n. gen., n. sp." pageId="6" pageNumber="449">Table 2.</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="6FAC4EACFFE4FF8EFC95FC1C5D91FBD9" pageId="6" pageNumber="449" type="diagnosis">
<paragraph id="27091D27FFE4FF8EFC95FC1C5D91FBD9" blockId="6.[813,1456,949,1052]" pageId="6" pageNumber="449">
<smallCapsWord id="21EF8BFBFFE4FF8EFC95FC1C5A9EFC0E" baselines="966,967" box="[813,923,949,972]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="6" pageNumber="449">DIAGNOSIS</smallCapsWord>
. — Intervallum
<quantity id="E04EB0C2FFE4FF8EFBF2FC1C5D99FC09" box="[1098,1180,949,973]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="6" pageNumber="449" unit="mm" value="6.5" valueMax="9.0" valueMin="4.0">4-9 mm</quantity>
wide; outer wall with 3 to 5 vertical rows of canals per intersept; septa completely porous; some synapticulae grouped into synapticular tabulae (
<figureCitation id="BF8D01A2FFE4FF8EFA86FC435C83FBC4" box="[1342,1414,1002,1026]" captionStart="FIG" captionStartId="7.[132,142,1940,1957]" captionTargetBox="[133,1454,216,1896]" captionTargetId="figure-16@7.[132,1455,215,1897]" captionTargetPageId="7" captionText="FIG. 2. — Fouanoucyathus tafraoutiensis El Bakhouch &amp; Kerner, n. gen., n. sp., holotype AA-FOU-CI-2 from section FO.IS8: A, transverse view of specimen prior to preparation;B, transverse section showing wall and septal porosity (inner wall at right);C, detail of longitudinal section showing the outer and inner walls with canalsand septal porosity;D, detail of longitudinal section showing inner wall porosity (central cavity at right);E, detailof longitudinal section showing outer wall porosity (intervallum at right); F, longitudinal section showing wall,septal porosity and a synapticular tabulae (inner wall at right).Scale bars:A,9 mm;B, 0.5 mm;C, E, F, 1 mm; D, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13691740" httpUri="https://zenodo.org/record/13691740/files/figure.png" pageId="6" pageNumber="449">Fig. 2F</figureCitation>
,
<figureCitation id="BF8D01A2FFE4FF8EFA34FC435A46FBD9" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[133,1454,216,1473]" captionTargetId="figure-235@8.[132,1455,215,1474]" captionTargetPageId="8" captionText="FIG. 3. — Fouanoucyathus tafraoutiensis El Bakhouch &amp; Kerner, n. gen., n. sp., paratype AA-FOU-CI-3 from FO.IS10: A, transverse view of specimen prior to preparation; B, detail of longitudinal section showing walls, septal porosity and synapticular tabulae (inner wall at left); C, detail of longitudinal section showing inner wall canals (central cavity at left); D, detail of longitudinal section showing outer wall canals and synapticular tabulae (intervallum at right). Scale bars: A, 8.5 mm; B, 2 mm; C, 1 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13691742" httpUri="https://zenodo.org/record/13691742/files/figure.png" pageId="6" pageNumber="449">3B, D</figureCitation>
); in cups of diameter
<quantity id="E04EB0C2FFE4FF8EFB99FBAD5D95FBD9" box="[1057,1168,1028,1052]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="5.0" metricValueMin="2.2" pageId="6" pageNumber="449" unit="mm" value="36.0" valueMax="50.0" valueMin="22.0">22-50 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="6FAC4EACFFE4FF8EFC95FBFC5C1CFA28" pageId="6" pageNumber="449" type="description">
<paragraph id="27091D27FFE4FF8EFC95FBFC5AC7FBA8" blockId="6.[811,1457,1109,1517]" box="[813,962,1109,1135]" pageId="6" pageNumber="449">
<smallCapsWord id="21EF8BFBFFE4FF8EFC95FBFC5AC7FBA8" baselines="1128,1128" box="[813,962,1109,1135]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="6" pageNumber="449">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="27091D27FFE4FF8EFC95FBDD5AC9FA88" blockId="6.[811,1457,1109,1517]" pageId="6" pageNumber="449">
Cup conical, diameter
<quantity id="E04EB0C2FFE4FF8EFB9BFBDD5D93FB4A" box="[1059,1174,1140,1167]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="5.0" metricValueMin="2.2" pageId="6" pageNumber="449" unit="mm" value="36.0" valueMax="50.0" valueMin="22.0">22-50 mm</quantity>
with intervallum
<quantity id="E04EB0C2FFE4FF8EFAE0FBDD5CAAFB4A" box="[1368,1455,1140,1167]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="6" pageNumber="449" unit="mm" value="6.5" valueMax="9.0" valueMin="4.0">4-9 mm</quantity>
wide, outer wall with 3-5 rows of canals (diameter
<quantity id="E04EB0C2FFE4FF8EFAF6FB3D5A5EFB0B" metricMagnitude="-4" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.8999999999999999" pageId="6" pageNumber="449" unit="mm" value="0.145" valueMax="0.2" valueMin="0.09">0.09-0.2 mm</quantity>
, lintels
<quantity id="E04EB0C2FFE4FF8EFC0DFB1D5D58FB0B" box="[949,1117,1204,1230]" metricMagnitude="-5" metricUnit="m" metricValue="6.5" metricValueMax="7" metricValueMin="6.0" pageId="6" pageNumber="449" unit="mm" value="0.065" valueMax="0.07" valueMin="0.06">0.06-0.07 mm</quantity>
, porosity coefficient 1.3-3.3), wall thickness (
<quantity id="E04EB0C2FFE4FF8EFC65FB7D5D6FFB2B" box="[989,1130,1236,1262]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="449" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2-0.3 mm</quantity>
). Inner wall with one row of straight canal per intersept (diameter
<quantity id="E04EB0C2FFE4FF8EFB6FFB5D5C64FACB" box="[1239,1377,1268,1294]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="6" pageNumber="449" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4-0.7 mm</quantity>
, lintels
<quantity id="E04EB0C2FFE4FF8EFC96FABD5AD5FAEB" box="[814,976,1300,1326]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" metricValueMax="2.3" metricValueMin="1.1" pageId="6" pageNumber="449" unit="mm" value="0.17" valueMax="0.23" valueMin="0.11">0.11-0.23 mm</quantity>
, porosity coefficient 1.9-6.4), wall thickness (
<quantity id="E04EB0C2FFE4FF8EFC8BFA9A5ABAFA88" box="[819,959,1331,1357]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="6" pageNumber="449" unit="mm" value="0.8999999999999999" valueMax="1.2" valueMin="0.6">0.6-1.2 mm</quantity>
).
</paragraph>
<paragraph id="27091D27FFE4FF8EFCFBFAFA5C39FA08" blockId="6.[811,1457,1109,1517]" pageId="6" pageNumber="449">
Septa are perced by numerous vertical pore rows, with the number varying from 6 to 21 rows; (perforation diameter
<quantity id="E04EB0C2FFE4FF8EFC95FA3A5AD4FA6B" box="[813,977,1427,1454]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="2.2" metricValueMin="1.4" pageId="6" pageNumber="449" unit="mm" value="0.18" valueMax="0.22" valueMin="0.14">0.14-0.22 mm</quantity>
, lintels
<quantity id="E04EB0C2FFE4FF8EFB9CFA3A5DC3FA68" box="[1060,1222,1427,1453]" metricMagnitude="-4" metricUnit="m" metricValue="2.55" metricValueMax="3.4" metricValueMin="1.7" pageId="6" pageNumber="449" unit="mm" value="0.255" valueMax="0.34" valueMin="0.17">0.17-0.34 mm</quantity>
) (
<figureCitation id="BF8D01A2FFE4FF8EFB65FA3A5C34FA68" box="[1245,1329,1427,1453]" captionStart="FIG" captionStartId="7.[132,142,1940,1957]" captionTargetBox="[133,1454,216,1896]" captionTargetId="figure-16@7.[132,1455,215,1897]" captionTargetPageId="7" captionText="FIG. 2. — Fouanoucyathus tafraoutiensis El Bakhouch &amp; Kerner, n. gen., n. sp., holotype AA-FOU-CI-2 from section FO.IS8: A, transverse view of specimen prior to preparation;B, transverse section showing wall and septal porosity (inner wall at right);C, detail of longitudinal section showing the outer and inner walls with canalsand septal porosity;D, detail of longitudinal section showing inner wall porosity (central cavity at right);E, detailof longitudinal section showing outer wall porosity (intervallum at right); F, longitudinal section showing wall,septal porosity and a synapticular tabulae (inner wall at right).Scale bars:A,9 mm;B, 0.5 mm;C, E, F, 1 mm; D, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13691740" httpUri="https://zenodo.org/record/13691740/files/figure.png" pageId="6" pageNumber="449">Fig. 2C</figureCitation>
,
<figureCitation id="BF8D01A2FFE4FF8EFA84FA3A5C5AFA68" box="[1340,1375,1427,1453]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[133,1454,216,1473]" captionTargetId="figure-235@8.[132,1455,215,1474]" captionTargetPageId="8" captionText="FIG. 3. — Fouanoucyathus tafraoutiensis El Bakhouch &amp; Kerner, n. gen., n. sp., paratype AA-FOU-CI-3 from FO.IS10: A, transverse view of specimen prior to preparation; B, detail of longitudinal section showing walls, septal porosity and synapticular tabulae (inner wall at left); C, detail of longitudinal section showing inner wall canals (central cavity at left); D, detail of longitudinal section showing outer wall canals and synapticular tabulae (intervallum at right). Scale bars: A, 8.5 mm; B, 2 mm; C, 1 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13691742" httpUri="https://zenodo.org/record/13691742/files/figure.png" pageId="6" pageNumber="449">3B</figureCitation>
). Some synapticulae grouped into synapticular tabulae.
</paragraph>
<paragraph id="27091D27FFE4FF8EFCFBFA7A5C1CFA28" blockId="6.[811,1457,1109,1517]" box="[835,1305,1491,1517]" pageId="6" pageNumber="449">RK 1.4-2.3. IC 1/4,0-1/9,0. GPC 1.1-1.9</paragraph>
</subSubSection>
<subSubSection id="6FAC4EACFFE4FF8EFC95F9BA5AE5F929" pageId="6" pageNumber="449" type="discussion">
<paragraph id="27091D27FFE4FF8EFC95F9BA5A91F9EE" blockId="6.[813,1457,1555,1772]" box="[813,916,1555,1581]" pageId="6" pageNumber="449">
<smallCapsWord id="21EF8BFBFFE4FF8EFC95F9BA5A91F9EE" baselines="1574,1574" box="[813,916,1555,1581]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="6" pageNumber="449">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="27091D27FFE4FF8EFC95F99B5AE5F929" blockId="6.[813,1457,1555,1772]" pageId="6" pageNumber="449">
The canals on the inner wall for this new taxon are quite regular in size and shape (
<figureCitation id="BF8D01A2FFE4FF8EFBF4F9FB5D98F9A9" box="[1100,1181,1618,1645]" captionStart="FIG" captionStartId="10.[132,143,1430,1447]" captionTargetBox="[133,1454,215,1386]" captionTargetId="figure-335@10.[132,1455,215,1387]" captionTargetPageId="10" captionText="FIG. 5. — Fouanoucyathus tafraoutiensis El Bakhouch &amp; Kerner,n. gen., n. sp., paratype AA-FOU-CI-1 from section FO.IS4:A, longitudinal view of specimen prior to preparation; B, oblique transverse section showing details of walls and septa porosity; C, longitudinal section (inner wall at right); D, detail of C showing inner wall porosity (central cavity at right); E, detail of C showing outer wall porosity. Scale bars: A, 4 mm; B, 0.4 mm; C, 1.5 mm; D, 1 mm; E, 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.13691746" httpUri="https://zenodo.org/record/13691746/files/figure.png" pageId="6" pageNumber="449">Fig. 5B</figureCitation>
) and their direction varies from horizontal to upward with all the intermediate directions. Septa are completely porous (6-21 rows) and their pores have round to slightly longitudinally elongate shape (
<figureCitation id="BF8D01A2FFE4FF8EFCC4F97B5AD7F929" box="[892,978,1746,1772]" captionStart="FIG" captionStartId="8.[132,143,1516,1533]" captionTargetBox="[133,1454,216,1473]" captionTargetId="figure-235@8.[132,1455,215,1474]" captionTargetPageId="8" captionText="FIG. 3. — Fouanoucyathus tafraoutiensis El Bakhouch &amp; Kerner, n. gen., n. sp., paratype AA-FOU-CI-3 from FO.IS10: A, transverse view of specimen prior to preparation; B, detail of longitudinal section showing walls, septal porosity and synapticular tabulae (inner wall at left); C, detail of longitudinal section showing inner wall canals (central cavity at left); D, detail of longitudinal section showing outer wall canals and synapticular tabulae (intervallum at right). Scale bars: A, 8.5 mm; B, 2 mm; C, 1 mm; D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13691742" httpUri="https://zenodo.org/record/13691742/files/figure.png" pageId="6" pageNumber="449">Fig. 3C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,178 @@
<document id="F9F80B8632B5CAF9A453717F25B1EC79" ID-DOI="10.5252/geodiversitas2024v46a12" ID-ISSN="1638-9395" ID-Zenodo-Dep="13691736" ID-ZooBank="urn:lsid:zoobank.org:pub:02D31D6C-15C2-498A-A6C6-62A0163814C8" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1725524660885" checkinUser="plazi" docAuthor="Bakhouch, Asmaa El, Kerner, Adeline, Azizi, Abdelfattah, Debrenne, Françoise, Jalil, Nour-Eddine, Hafid, Ahmid &amp; Hariri, Khadija El" docDate="2024" docId="AF1FAC31FFE4FF8EFF2EFCBF5B91F96A" docLanguage="en" docName="geodiversitas.2024.46.12.pdf" docOrigin="Geodiversitas 46 (12)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a12.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Fouanoucyathus El Bakhouch &amp; Kerner 2024, n. gen." docType="treatment" docUuid="EFFD9C78-3BA5-4216-8931-4D2694F7F3F2" docUuidSource="ZooBank" docVersion="3" lastPageNumber="449" masterDocId="5326D449FFE2FF88FFB8FFA95905FFC5" masterDocTitle="New archaeocyath genus from the early Cambrian of the western Anti-Atlas, Morocco" masterLastPageNumber="455" masterPageNumber="445" pageNumber="449" updateTime="1726061974691" updateUser="jonas" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
<mods:mods id="43BE60BF2832983A2960404D1CF3409C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AEA47536759D19AECDA65B552984DA3B">
<mods:title id="C790B76BD9BC89F41E1A42E553902139">New archaeocyath genus from the early Cambrian of the western Anti-Atlas, Morocco</mods:title>
</mods:titleInfo>
<mods:name id="9CE9A52FC9C29532A19A8D0F13ADB435" type="personal">
<mods:role id="88BAC367FBD6725FD09CB1501E2B27B0">
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</mods:role>
<mods:namePart id="6AA8E76E66A71D0D42F824DF4A965793">Bakhouch, Asmaa El</mods:namePart>
<mods:affiliation id="0C5120907B11FB8FAF3F902AD33952EC">Laboratoire de Géo-ressources, Géoenvironnement et Génie civil, Département des Sciences de la Terre, Faculté des Sciences et Techniques, Université Cadi-Ayyad, boîte postale 549, 40000 Marrakech (Morocco)</mods:affiliation>
<mods:nameIdentifier id="EBBAB6CDF69DEE4CBB4D0D7B74A20DA7" type="email">asmaa.bakhouche@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="CE66C1CAE123BC82369017A06B92D2CC" type="personal">
<mods:role id="5413DF14C12D9A67B855347FE1034C98">
<mods:roleTerm id="61D19B92E5FE0A30E918401A1C36A7CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="71CE1D572F936497659BF06B1A7AD899">Kerner, Adeline</mods:namePart>
<mods:affiliation id="BBACF125305E7FC74BE52EFD31A6DA54">UMR 7207 - Center for Research on Palaeontology - Paris, CNRS - Sorbonne Université - Muséum national dHistoire naturelle, Bâtiment de géologie, case postale 48, 57 rue Cuvier, 75231 Paris cedex 05 (France)</mods:affiliation>
<mods:nameIdentifier id="038B4C0988943AA1C04BE6656A3EC702" type="email">adeline.kerner@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="C01BE6D3A75DA9AA7F1FCAD70E9CCE3B" type="personal">
<mods:role id="326FD665B2434235DDB2769784B366DB">
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<mods:namePart id="BC7D1E7134D3ACF7045730590359A760">Azizi, Abdelfattah</mods:namePart>
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<mods:namePart id="2A246D15D1041BB88D617C3A2175F384">Jalil, Nour-Eddine</mods:namePart>
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<mods:namePart id="FAE4978F8A486BFDFEBD2E4EE0D90D7B">Hariri, Khadija El</mods:namePart>
<mods:affiliation id="B57B760C098EC3B417D64C3E5A3F433C">Laboratoire de Géo-ressources, Géoenvironnement et Génie civil, Département des Sciences de la Terre, Faculté des Sciences et Techniques, Université Cadi-Ayyad, boîte postale 549, 40000 Marrakech (Morocco)</mods:affiliation>
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<mods:title id="5F64AA3A36D9CED5E4463817B3C8B091">Geodiversitas</mods:title>
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<mods:part id="7EDBD0F51C76552C0F26F7E56E5A4134">
<mods:date id="1F899164E874BFB0B6E33A4C69E9913B">2024</mods:date>
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<mods:number id="5E7660159365852BF877AB610843C6AC">2024-09-05</mods:number>
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<mods:number id="9C5F1C6772A4DAECC02D9EEC469B6391">46</mods:number>
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<mods:number id="2F78A9125B5A190DDB5FB05DC9762B7D">12</mods:number>
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<mods:url id="FE16221DE2C1D6929E7F0B48292F0EB7">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a12.pdf</mods:url>
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<mods:identifier id="C9A5525223E7C3C7B280E5841142A5A5" type="DOI">10.5252/geodiversitas2024v46a12</mods:identifier>
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Genus
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<emphasis id="15C2C135FFE4FF8EFF5DFCBF5895FCF5" box="[229,400,790,816]" italics="true" pageId="6" pageNumber="449">Fouanoucyathus</emphasis>
El Bakhouch &amp; Kerner
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,
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</heading>
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urn:lsid:zoobank.org:act:
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. — Named after the Fouanou Syncline where the specimen was discovered.
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<paragraph id="27091D27FFE4FF8EFF3CFC4C585EFB89" blockId="6.[132,777,996,1100]" pageId="6" pageNumber="449">
<smallCapsWord id="21EF8BFBFFE4FF8EFF3CFC4C59B3FC3F" baselines="1014,1014" box="[132,182,997,1020]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="6" pageNumber="449">
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. —
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<emphasis id="15C2C135FFE4FF8EFE84FC4D5B58FC39" box="[316,605,996,1020]" italics="true" pageId="6" pageNumber="449">Fouanoucyathus tafraoutiensis</emphasis>
El Bakhouch &amp; Kerner
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,
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, Early Cambrian (Cambrian age 4 or Botoman), Issafen Formation, Fouanou Syncline, Ait Abdellah, western Anti-Atlas,
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.
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<smallCapsWord id="21EF8BFBFFE4FF8EFF3CFBCD59F7FBBF" baselines="1141,1142" box="[132,242,1124,1147]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="6" pageNumber="449">DIAGNOSIS</smallCapsWord>
. — Outer wall with several vertical rows of horizontal to upwardly projecting slightly S-shaped canals; inner wall with one vertical row of horizontal to upwardly S-shaped canals per intersept, septa completely porous, linked by synapticulae.
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<subSubSection id="6FAC4EACFFE4FF8EFF3DFB5F5B91F96A" pageId="6" pageNumber="449" type="discussion">
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<smallCapsWord id="21EF8BFBFFE4FF8EFF3DFB5F580EFACB" baselines="1289,1289" box="[133,267,1270,1296]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Discussion" pageId="6" pageNumber="449">DISCUSSION</smallCapsWord>
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<paragraph id="27091D27FFE4FF8EFF3CFABF5B91F96A" blockId="6.[131,776,1270,1712]" pageId="6" pageNumber="449">
<taxonomicName id="E0B666A4FFE4FF8EFF3CFABF5B32FAF5" authority="El Bakhouch &amp; Kerner" authorityName="El Bakhouch &amp; Kerner" authorityYear="2024" box="[132,567,1302,1328]" family="Carinacyathidae" genus="Fouanoucyathus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="449" phylum="Porifera" rank="genus" status="gen. nov.">
<emphasis id="15C2C135FFE4FF8EFF3CFABF582AFAF5" box="[132,303,1302,1328]" italics="true" pageId="6" pageNumber="449">Fouanoucyathus</emphasis>
El Bakhouch &amp; Kerner
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,
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is the only carinacyathid genus with synapticulae, it differs to
<taxonomicName id="E0B666A4FFE4FF8EFD08FA9F59D4FAB5" authorityName="Vologdin" authorityYear="1932" genus="Carinacyathus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="449" phylum="Porifera" rank="genus">
<emphasis id="15C2C135FFE4FF8EFD08FA9F59D4FAB5" italics="true" pageId="6" pageNumber="449">Carinacyathus</emphasis>
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in the shape of canals on the outer wall (S-shaped but not straight), absence of supplementary bracts externally and the appearance of the inner wall (S-shaped but not straight, horizontal to upwardly but not downwardly plus no additional bracts on central cavity side).
<emphasis id="15C2C135FFE4FF8EFD1DFA7F59CBF9CA" italics="true" pageId="6" pageNumber="449">Hupercyathelus</emphasis>
differs in possessing an outer wall with downwardly projecting canals and bearing supplementary independent microporous sheath externally and an inner wall with several rows of canals.
<taxonomicName id="E0B666A4FFE4FF8EFE8FF9FC58C8F9AA" box="[311,461,1621,1647]" genus="Porocyathellus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="449" phylum="Porifera" rank="genus">
<emphasis id="15C2C135FFE4FF8EFE8FF9FC58C8F9AA" box="[311,461,1621,1647]" italics="true" pageId="6" pageNumber="449">Porocyathellus</emphasis>
</taxonomicName>
differs in possessing several rows of canals on the inner wall.
<taxonomicName id="E0B666A4FFE4FF8EFE54F9DC5BBEF94A" box="[492,699,1653,1679]" genus="Vologdinocyathellus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="449" phylum="Porifera" rank="genus">
<emphasis id="15C2C135FFE4FF8EFE54F9DC5BBEF94A" box="[492,699,1653,1679]" italics="true" pageId="6" pageNumber="449">Vologdinocyathellus</emphasis>
</taxonomicName>
differs in possessing an inner wall with straight canals.
</paragraph>
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