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<mods:title id="DB17DB07CC3B846E18196E036C92B4FE">Stenus attenboroughi nov. sp. and records of Stenus LATREILLE, 1797 from New Guinea (Coleoptera, Staphylinidae)</mods:title>
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<mods:namePart id="1D368DE50F86F1C900322604CAF0B17A">M, Cyclops</mods:namePart>
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<heading id="D0B481A57551FFD27BA16074FDB8BA28" box="[163,528,1076,1098]" fontSize="8" level="2" pageId="5" pageNumber="1010" reason="2">
<taxonomicName id="4C434D4A7551FFD27BA16074FDB8BA28" ID-CoL="529D7" authority="PUTHZ, 1971" authorityName="PUTHZ" authorityYear="1971" box="[163,528,1076,1098]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1010" phylum="Arthropoda" rank="species" species="hypostenoides">
<emphasis id="B937EADB7551FFD27BA16074FE67BA28" bold="true" box="[163,463,1076,1098]" pageId="5" pageNumber="1010">
<emphasis id="B937EADB7551FFD27BA16074FE29BA28" bold="true" box="[163,385,1076,1098]" italics="true" pageId="5" pageNumber="1010">Stenus hypostenoides</emphasis>
<bibRefCitation id="EFD24B387551FFD27A8A6074FE67BA28" author="PUTHZ V" box="[392,463,1077,1098]" pageId="5" pageNumber="1010" pagination="447 - 469" refId="ref3136" refString="PUTHZ V. (1971): Revision of the Stenus species of New Guinea. Part I. (Coleoptera: Staphylinidae). - Pacific Insects 13 (3 - 4): 447 - 469." type="journal article" year="1971">PUTHZ</bibRefCitation>
</emphasis>
,
<emphasis id="B937EADB7551FFD27ADF6074FDB8BA28" bold="true" box="[477,528,1077,1098]" pageId="5" pageNumber="1010">1971</emphasis>
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</paragraph>
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<materialsCitation id="3B2B3C947551FFD27BA1601AFCE7BAEA" collectingDate="2019-05-18" elevation="440" latitude="-2.5362556" location="West-Papua" longLatPrecision="1" longitude="140.51297" municipality="Cyclops Mountains" pageId="5" pageNumber="1010" specimenCount="1" specimenCount-male="1">
M a t e r i a l e x a m i n e d:
<specimenCount id="9D45FD407551FFD27912601AFD87BA12" box="[528,559,1115,1136]" pageId="5" pageNumber="1010" type="male">1♂</specimenCount>
:
<location id="8E9C60127551FFD2794F601AFD11BA12" LSID="urn:lsid:plazi:treatment:03EA87DF7551FFD27BA16074FE27BB3A:8E9C60127551FFD2794F601AFD11BA12" box="[589,697,1115,1136]" latitude="-2.5362556" longLatPrecision="1" longitude="140.51297" municipality="Cyclops Mountains" name="West-Papua" pageId="5" pageNumber="1010">West-Papua</location>
,
<collectingMunicipality id="6B98ACB37551FFD279D5601AFC31BA13" box="[727,921,1115,1137]" pageId="5" pageNumber="1010">Cyclops Mountains</collectingMunicipality>
,
<geoCoordinate id="EE77500E7551FFD278B5601BFB86BA12" box="[951,1070,1114,1136]" degrees="2" direction="south" minutes="32" orientation="latitude" pageId="5" pageNumber="1010" precision="1" seconds="10.52" value="-2.5362556">2°32’10.52&quot;S</geoCoordinate>
<geoCoordinate id="EE77500E7551FFD27BCD6033FEF7BAEA" box="[207,351,1138,1160]" degrees="140" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="1010" precision="1" seconds="46.71" value="140.51297">140°30’46.71&quot;E</geoCoordinate>
,
<quantity id="4CBB9B2C7551FFD27A6C6032FE05BAEA" box="[366,429,1139,1160]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="1010" unit="m" value="440.0">
<elevation id="006ED1FA7551FFD27A6C6032FE05BAEA" box="[366,429,1139,1160]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="1010" unit="m" value="440.0">440 m</elevation>
</quantity>
, loamy and mossy wall, stream,
<date id="FFFD10097551FFD279EE6033FCE7BAEA" box="[748,847,1138,1160]" pageId="5" pageNumber="1010" value="2019-05-18">
<collectingDate id="EFB9E9E17551FFD279EE6033FCE7BAEA" box="[748,847,1138,1160]" pageId="5" pageNumber="1010" value="2019-05-18">18.05.2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2B3C947551FFD278626032FB82BAC2" collectingDate="2019-05-27" county="Timua river" latitude="-2.441075" location="West-Papua" municipality="Foja Mountains" pageId="5" pageNumber="1010" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D45FD407551FFD278626032FCD6BAEA" box="[864,894,1139,1160]" pageId="5" pageNumber="1010" type="male">1♂</specimenCount>
:
<location id="8E9C60127551FFD2788F6032FC51BAEA" LSID="urn:lsid:plazi:treatment:03EA87DF7551FFD27BA16074FE27BB3A:8E9C60127551FFD2788F6032FC51BAEA" box="[909,1017,1139,1160]" county="Timua river" latitude="-2.441075" municipality="Foja Mountains" name="West-Papua" pageId="5" pageNumber="1010">West-Papua</location>
,
<collectingMunicipality id="6B98ACB37551FFD27F056032FE87BAC2" pageId="5" pageNumber="1010">Foja Mountains</collectingMunicipality>
,
<geoCoordinate id="EE77500E7551FFD27A3A60CBFE07BAC2" box="[312,431,1162,1184]" degrees="2" direction="south" minutes="26" orientation="latitude" pageId="5" pageNumber="1010" precision="1" seconds="27.87" value="-2.441075">2°26’27.87&quot;S</geoCoordinate>
138°46’27.283, 280 m,
<collectingCounty id="629D4E457551FFD2798460CBFD59BAC2" box="[646,753,1162,1184]" pageId="5" pageNumber="1010">Timua river</collectingCounty>
, small loamy waterfall,
<date id="FFFD10097551FFD278C760CBFB82BAC2" box="[965,1066,1162,1184]" pageId="5" pageNumber="1010" value="2019-05-27">
<collectingDate id="EFB9E9E17551FFD278C760CBFB82BAC2" box="[965,1066,1162,1184]" pageId="5" pageNumber="1010" value="2019-05-27">27.05.2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2B3C947551FFD27BCD60E2FE63BAB2" collectingDate="2019-05-29" county="Timua river" elevation="350" latitude="-2.4717057" location="West-Papua" longLatPrecision="1" longitude="138.77443" municipality="Foja Mountains" pageId="5" pageNumber="1010" specimenCount="3" specimenCount-female="3">
<specimenCount id="9D45FD407551FFD27BCD60E2FF56BADA" box="[207,254,1187,1208]" pageId="5" pageNumber="1010" type="female">3♀♀</specimenCount>
:
<location id="8E9C60127551FFD27A0860E2FEDEBADA" LSID="urn:lsid:plazi:treatment:03EA87DF7551FFD27BA16074FE27BB3A:8E9C60127551FFD27A0860E2FEDEBADA" box="[266,374,1187,1208]" county="Timua river" latitude="-2.4717057" longLatPrecision="1" longitude="138.77443" municipality="Foja Mountains" name="West-Papua" pageId="5" pageNumber="1010">West-Papua</location>
,
<collectingMunicipality id="6B98ACB37551FFD27A8360E3FDB8BADA" box="[385,528,1186,1208]" pageId="5" pageNumber="1010">Foja Mountains</collectingMunicipality>
,
<geoCoordinate id="EE77500E7551FFD2791960E2FD3BBADA" box="[539,659,1187,1208]" degrees="2" direction="south" minutes="28" orientation="latitude" pageId="5" pageNumber="1010" precision="1" seconds="18.14" value="-2.4717057">2°28’18.14&quot;S</geoCoordinate>
<geoCoordinate id="EE77500E7551FFD2799860E3FC83BADA" box="[666,811,1186,1208]" degrees="138" direction="east" minutes="46" orientation="longitude" pageId="5" pageNumber="1010" precision="1" seconds="27.98" value="138.77443">138°46’27.98&quot;E</geoCoordinate>
,
<quantity id="4CBB9B2C7551FFD2783460E3FCDBBADA" box="[822,883,1186,1208]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="1010" unit="m" value="350.0">
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,
<collectingCounty id="629D4E457551FFD2787D60E3FC43BADA" box="[895,1003,1186,1208]" pageId="5" pageNumber="1010">Timua river</collectingCounty>
, loamy bluish grey wall,
<date id="FFFD10097551FFD27A6760FBFE63BAB2" box="[357,459,1210,1232]" pageId="5" pageNumber="1010" value="2019-05-29">
<collectingDate id="EFB9E9E17551FFD27A6760FBFE63BAB2" box="[357,459,1210,1232]" pageId="5" pageNumber="1010" value="2019-05-29">29.05.2019</collectingDate>
</date>
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.
</paragraph>
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<subSubSection id="C35965427551FFD27BA060ABFE27BB3A" pageId="5" pageNumber="1010" type="description">
<paragraph id="8BFC36C97551FFD27BA060ABFE27BB3A" blockId="5.[162,1070,1258,1368]" pageId="5" pageNumber="1010">
This species was collected together with
<taxonomicName id="4C434D4A7551FFD2797160ABFC8CBA9D" authority="Mainda &amp; M &amp; M, 2021" authorityName="Mainda &amp; M &amp; M" authorityYear="2021" box="[627,804,1258,1279]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1010" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="B937EADB7551FFD2797160ABFC8CBA9D" box="[627,804,1258,1279]" italics="true" pageId="5" pageNumber="1010">S. attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457A07551FFD2783160AAFCD5BB62" box="[819,893,1259,1280]" pageId="5" pageNumber="1010" rank="species">nov.sp.</taxonomicNameLabel>
in the Cyclops Mountains, where it lives in the same habitat. In the Foja Mountains it was only collected on bluish grey loamy walls in the valley of the Timua river, where it lives together with
<taxonomicName id="4C434D4A7551FFD27F186164FF49BB35" authorityName="PUTHZ" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1010" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B937EADB7551FFD27F186164FF49BB35" italics="true" pageId="5" pageNumber="1010">S. balkei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C434D4A7551FFD27A106103FE21BB35" authorityName="FAUVEL" authorityYear="1878" box="[274,393,1346,1367]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1010" phylum="Arthropoda" rank="species" species="prismalis">
<emphasis id="B937EADB7551FFD27A106103FE21BB35" box="[274,393,1346,1367]" italics="true" pageId="5" pageNumber="1010">S. prismalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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data/03/EA/87/03EA87DF7551FFD27BA166DFFC0ABD5D.xml
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@ -1,87 +0,0 @@
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<emphasis id="B937EADB7551FFD27BA166DFFEEDBCD6" bold="true" box="[163,325,670,692]" italics="true" pageId="5" pageNumber="1010">Stenus agricola</emphasis>
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M a t e r i a l e x a m i n e d:
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,
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: West-Papua, Sentani Airport, Lake Sentani,
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, overgrown shore (grass),
<date id="FFFD10097551FFD27947669DFD02BC90" box="[581,682,732,754]" pageId="5" pageNumber="1010" value="2019-09-06">09.06.2019</date>
.
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This species was found at the overgrown shore of Lake Sentani between grass, which was submerged under water. It was collected together with
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<taxonomicName id="4C434D4A7551FFD279CD6768FC94BD5C" authorityName="Motschulsky" authorityYear="1858" box="[719,828,809,830]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1010" phylum="Arthropoda" rank="species" species="piliferus">S. piliferus</taxonomicName>
obesulus
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82
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M a t e r i a l e x a m i n e d:
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, West-Papua, Foja Mountains,
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,
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, Timua river, loamy wall,
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.
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<paragraph id="8BFC36C97551FFD27BA06796FE52BA68" blockId="5.[162,1069,983,1034]" pageId="5" pageNumber="1010">The species was collected on the steep wall of the Timua river. There the species lives on loamy ground and between rocks.</paragraph>
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C35965427555FFD47BA166B4FC5EBB90" lastPageId="3" lastPageNumber="1008" pageId="1" pageNumber="1006" type="description">
<paragraph id="8BFC36C97555FFD67BA166B4FE3FBD30" blockId="1.[163,1071,724,850]" pageId="1" pageNumber="1006">
T y p e m a t e r i a l e x a m i n e d:
<typeStatus id="54F8886B7555FFD6794266B4FD31BD69" box="[576,665,757,779]" pageId="1" pageNumber="1006" type="holotype">Holotype</typeStatus>
<specimenCount id="9D45FD407555FFD6799B66B4FD00BD69" box="[665,680,757,779]" pageId="1" pageNumber="1006" type="male">♂</specimenCount>
:&quot;West-Papua:Sentani,Cyclops Mountains, loamy and mossy wall, stream,
<quantity id="4CBB9B2C7555FFD67ADD674CFDB0BD40" box="[479,536,781,802]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="1" pageNumber="1006" unit="m" value="440.0">440 m</quantity>
,
<geoCoordinate id="EE77500E7555FFD67922674CFD3FBD41" box="[544,663,781,803]" degrees="2" direction="south" minutes="32" orientation="latitude" pageId="1" pageNumber="1006" precision="1" seconds="10.52" value="-2.5362556">2°32’10.52&quot;S</geoCoordinate>
<geoCoordinate id="EE77500E7555FFD6799E674CFC84BD41" box="[668,812,781,803]" degrees="140" direction="east" minutes="30" orientation="longitude" pageId="1" pageNumber="1006" precision="1" seconds="46.71" value="140.51297">140°30’46.71&quot;E</geoCoordinate>
,
<date id="FFFD10097555FFD67831674CFC3FBD41" box="[819,919,781,803]" pageId="1" pageNumber="1006" value="2019-05-18">18.05.2019</date>
&quot; [cM].
<typeStatus id="54F8886B7555FFD678DB674CFB86BD40" box="[985,1070,781,802]" pageId="1" pageNumber="1006" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D45FD407555FFD67BD46764FEB8BD58" box="[214,272,805,826]" pageId="1" pageNumber="1006" type="male">11♂♂</specimenCount>
,
<specimenCount id="9D45FD407555FFD67A196764FEFCBD59" box="[283,340,805,827]" pageId="1" pageNumber="1006" type="female">17♀♀</specimenCount>
): Same collecting data as the HT [cM,
<specimenCount id="9D45FD407555FFD679C36764FD76BD58" box="[705,734,805,826]" pageId="1" pageNumber="1006" type="male">1♂</specimenCount>
/
<specimenCount id="9D45FD407555FFD679E56764FD57BD58" box="[743,767,805,826]" pageId="1" pageNumber="1006" type="female">1♀</specimenCount>
NHMW,
<specimenCount id="9D45FD407555FFD6785F6764FCD2BD58" box="[861,890,805,826]" pageId="1" pageNumber="1006" type="male">1♂</specimenCount>
/
<specimenCount id="9D45FD407555FFD678816764FC3FBD58" box="[899,919,805,826]" pageId="1" pageNumber="1006" type="female">1♀</specimenCount>
SMNK,
<specimenCount id="9D45FD407555FFD678F26764FBA4BD58" box="[1008,1036,805,826]" pageId="1" pageNumber="1006" type="male">1♂</specimenCount>
/
<specimenCount id="9D45FD407555FFD67F176764FB83BD58" box="[1045,1067,805,826]" pageId="1" pageNumber="1006" type="female">1♀</specimenCount>
SMNS,
<specimenCount id="9D45FD407555FFD67A1B677CFE9EBD30" box="[281,310,829,850]" pageId="1" pageNumber="1006" type="male">1♂</specimenCount>
/
<specimenCount id="9D45FD407555FFD67A42677CFEFFBD30" box="[320,343,829,850]" pageId="1" pageNumber="1006" type="female">1♀</specimenCount>
ZSM].
</paragraph>
<paragraph id="8BFC36C97555FFD67BA06723FC2BBA68" blockId="1.[162,1070,865,1034]" pageId="1" pageNumber="1006">
D e s c r i p t i o n: Measurements of the HT (in mm): BL: 4.70, DE: 0.48, FBL: 2.20, EL: 0.90, EW: 0.75, HW: 0.78, PL: 0.63, PW: 0.55, SL: 0.68. Measurements of the
<specimenCount id="9D45FD407555FFD678EB673EFB86BDF6" box="[1001,1070,895,916]" pageId="1" pageNumber="1006" type="male">10 ♂♂</specimenCount>
PTT (in mm): BL: 4.55-4.70, DE: 0.47-0.50, FBL: 2.20-2.36, EL: 0.88-0.95, EW: 0.80- 0.83, HW: 0.78-0.83, PL: 0.60-0.65, PW: 0.55-0.60, SL: 0.55-0.63. Measurements of the
<specimenCount id="9D45FD407555FFD67BA16796FF40BD8E" box="[163,232,983,1004]" pageId="1" pageNumber="1006" type="female">17 ♀♀</specimenCount>
PTT (in mm): BL: 4.50-4.70, DE: 0.48-0.53, FBL: 2.30-2.35, EL: 0.85-0.975, EW: 0.73-0.83, HW: 0.78-0.83, PL: 0.58-0.68, PW: 0.55-0.58, SL: 0.63-0.68.
</paragraph>
<paragraph id="8BFC36C97555FFD67BA16058FE1ABA0B" blockId="1.[163,1070,1049,1130]" pageId="1" pageNumber="1006">
Habitus as in
<figureCitation id="13782A4C7555FFD67A2F6058FEC1BA4C" box="[301,361,1049,1071]" captionStart="Fig" captionStartId="2.[163,196,1185,1207]" captionText="Fig. 1: Habitus of S. attenboroughi nov.sp., male paratype." pageId="1" pageNumber="1006">Fig. 1</figureCitation>
. Macropterous, metallic blue-green-violet, shining, with metallic hue, without microsculpture; maxillary palpi, legs and antennae orange, last segments infuscate; labrum aeneous.
</paragraph>
<paragraph id="8BFC36C97555FFD67BA16038FC3BBB66" blockId="1.[163,1070,1145,1284]" pageId="1" pageNumber="1006">Head 1.04 times broader than elytra, median portion of frons about half as wide as lateral portions, elevated, not reaching the high of the inner eye margin. Median portion separated from lateral portions by two narrow lateral furrows. Punctation dense and coarse, diameter of punctures larger than basal cross-section of antennomere III. Antennae moderately slender, when reflexed not extending to the posterior margin of pronotum.</paragraph>
<paragraph id="8BFC36C97555FFD67BA16155FE78BBDE" blockId="1.[163,1070,1299,1468]" pageId="1" pageNumber="1006">Pronotum 1.15 times as long as broad, broadest in the posterior third, sides towards anterior margin straight, towards posterior margin concavely narrowed. Punctation coarse and very dense, partly coalescent, largest punctures in dorsal middle as large as basal diameter of antennomere III, interstices much narrower than diameter of punctures. Legs slender; metatarsomere I as long as the combined length of metatarsomeres II–V and longer than metatarsomere V.</paragraph>
<caption id="DF3C66417556FFD57BA160E0FD12BAD4" box="[163,698,1185,1207]" pageId="2" pageNumber="1007" startId="2.[163,196,1185,1207]">
<paragraph id="8BFC36C97556FFD57BA160E0FD12BAD4" blockId="2.[163,698,1185,1207]" box="[163,698,1185,1207]" pageId="2" pageNumber="1007">
<emphasis id="B937EADB7556FFD57BA160E0FF70BAD4" bold="true" box="[163,216,1185,1207]" pageId="2" pageNumber="1007">Fig. 1</emphasis>
: Habitus of
<taxonomicName id="4C434D4A7556FFD57A4B60E0FE48BAD4" authority="Mainda &amp; M &amp; M, 2021" authorityName="Mainda &amp; M &amp; M" authorityYear="2021" box="[329,480,1185,1206]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1007" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="B937EADB7556FFD57A4B60E0FE48BAD4" box="[329,480,1185,1206]" italics="true" pageId="2" pageNumber="1007">S. attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457A07556FFD57AE460E0FD82BAD4" box="[486,554,1185,1206]" pageId="2" pageNumber="1007" rank="species">nov.sp.</taxonomicNameLabel>
, male paratype.
</paragraph>
</caption>
<paragraph id="8BFC36C97556FFD57BA16084FCBEBB77" blockId="2.[163,1069,1221,1302]" pageId="2" pageNumber="1007">Elytra 1.2 times as long as broad, humeral angles prominent, sides nearly parallel, suture unmodified. Punctation coarse and dense, not coalescent, punctures about as large as those on pronotum, interstices narrower than diameter of punctures.</paragraph>
<paragraph id="8BFC36C97556FFD57BA16164FCA1BB3A" blockId="2.[163,1069,1317,1368]" pageId="2" pageNumber="1007">Abdomen cylindrical; basal impressions of tergites I-III deep; punctation very fine, nearly invisible, interstices much wider than diameter of punctures.</paragraph>
<paragraph id="8BFC36C97556FFD57BA16129FD06B84C" blockId="2.[163,1070,1384,1582]" pageId="2" pageNumber="1007">
M a l e: Femora slightly dilated, tibiae simple. Sternites III–V unmodified, sternite VI broadly impressed in posterior half, with long erect setae; sternite VII broadly impressed, sides of impression extending posteriorly, densely set with long erected setae; sternite VIII with a narrowed notch in posterior fivth (
<figureCitation id="13782A4C7556FFD579446181FD2CBBB4" box="[582,644,1472,1494]" captionStart="Figs 2-5" captionStartId="3.[163,203,1342,1364]" captionTargetBox="[163,1073,579,1331]" captionTargetId="figure-291@3.[163,1073,579,1331]" captionTargetPageId="3" captionText="Figs 2-5: Aedeagus of S. attenboroughi nov.sp. (2) and S. anteros (5); Sternite VIII of S. attenboroughi nov.sp. (3) and S. anteros (4)." figureDoi="http://doi.org/10.5281/zenodo.5277537" httpUri="https://zenodo.org/record/5277537/files/figure.png" pageId="2" pageNumber="1007">Fig. 3</figureCitation>
); sternite IX serrate apicolaterally; tergite X with rounded posterior margin. Apical portion of aedeagus (
<figureCitation id="13782A4C7556FFD5785C619CFC0ABB91" box="[862,930,1501,1523]" captionStart="Figs 2-5" captionStartId="3.[163,203,1342,1364]" captionTargetBox="[163,1073,579,1331]" captionTargetId="figure-291@3.[163,1073,579,1331]" captionTargetPageId="3" captionText="Figs 2-5: Aedeagus of S. attenboroughi nov.sp. (2) and S. anteros (5); Sternite VIII of S. attenboroughi nov.sp. (3) and S. anteros (4)." figureDoi="http://doi.org/10.5281/zenodo.5277537" httpUri="https://zenodo.org/record/5277537/files/figure.png" pageId="2" pageNumber="1007">Fig. 2</figureCitation>
) spatula-like enlarged, median lobe with two long expulsion-bands; internal sac asymmetric; parameres shorter than median lobe, with seven setae apically.
</paragraph>
<paragraph id="8BFC36C97557FFD47BA16495FD97BF64" blockId="3.[163,1070,211,262]" pageId="3" pageNumber="1008">F e m a l e: Sternite VIII with rounded posterior margin; valvifera serrate apicolaterally; tergite X with rounded posterior margin.</paragraph>
<paragraph id="8BFC36C97557FFD47BA16557FD46BC56" blockId="3.[163,1071,278,564]" pageId="3" pageNumber="1008">
D i f f e r e n t i a l d i a g n o s i s: This species belongs to the group of
<taxonomicName id="4C434D4A7557FFD478EE6557FE36BF2A" authority="FAUVEL, 1878" authorityName="FAUVEL" authorityYear="1878" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="prismalis">
<emphasis id="B937EADB7557FFD478EE6557FEA9BF2A" italics="true" pageId="3" pageNumber="1008">Stenus prismalis</emphasis>
FAUVEL, 1878
</taxonomicName>
. In this group
<taxonomicName id="4C434D4A7557FFD479366575FCA5BF2A" authority="Mainda &amp; M &amp; M, 2021" authorityName="Mainda &amp; M &amp; M" authorityYear="2021" box="[564,781,307,329]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="B937EADB7557FFD479366575FCA5BF2A" box="[564,781,307,329]" italics="true" pageId="3" pageNumber="1008">Stenus attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457A07557FFD478146575FCC8BF2B" box="[790,864,308,329]" pageId="3" pageNumber="1008" rank="species">nov.sp.</taxonomicNameLabel>
is closely related to
<taxonomicName id="4C434D4A7557FFD47BA16510FE6ABF04" authority="PUTHZ, 2016" authorityName="PUTHZ" authorityYear="2016" box="[163,450,336,358]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="anteros">
<emphasis id="B937EADB7557FFD47BA16510FE90BF04" box="[163,312,337,358]" italics="true" pageId="3" pageNumber="1008">Stenus anteros</emphasis>
<bibRefCitation id="EFD24B387557FFD47A3D6510FE6ABF04" author="PUTHZ V" box="[319,450,336,358]" pageId="3" pageNumber="1008" pagination="779 - 834" refId="ref3243" refString="PUTHZ V. (2016): Ubersicht uber die Steninen der melanesischen Subregion (Coleoptera, Staphylinidae). - Linzer biologische Beitrage 48 (1): 779 - 834." type="journal article" year="2016">PUTHZ, 2016</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F35476597557FFD47AD26510FD32BF04" box="[464,666,336,358]" name="Papua New Guinea" pageId="3" pageNumber="1008">Papua New Guinea</collectingCountry>
: Haia), from which it is distinguishable by the male sexual characters (
<figureCitation id="13782A4C7557FFD47ADD652FFD93BFE6" box="[479,571,366,388]" captionStart="Figs 2-5" captionStartId="3.[163,203,1342,1364]" captionTargetBox="[163,1073,579,1331]" captionTargetId="figure-291@3.[163,1073,579,1331]" captionTargetPageId="3" captionText="Figs 2-5: Aedeagus of S. attenboroughi nov.sp. (2) and S. anteros (5); Sternite VIII of S. attenboroughi nov.sp. (3) and S. anteros (4)." figureDoi="http://doi.org/10.5281/zenodo.5277537" httpUri="https://zenodo.org/record/5277537/files/figure.png" pageId="3" pageNumber="1008">Figs 4, 5</figureCitation>
) and by sparser punctation of tergite V. From
<taxonomicName id="4C434D4A7557FFD47F19652FFEC5BFC3" authority="PUTHZ, 1969" authorityName="PUTHZ" authorityYear="1969" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="aglaia">
<emphasis id="B937EADB7557FFD47F19652FFF4CBFC2" italics="true" pageId="3" pageNumber="1008">S. aglaia</emphasis>
<bibRefCitation id="EFD24B387557FFD47BE865CDFEC5BFC3" author="PUTHZ V" box="[234,365,395,417]" pageId="3" pageNumber="1008" pagination="525 - 528" refId="ref3102" refString="PUTHZ V. (1969): The first Stenus species from the Solomons (Coleoptera: Staphylinidae). - Pacific Insects 11 (3 - 4): 525 - 528." type="journal article" year="1969">PUTHZ, 1969</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F35476597557FFD47A7B65CAFD81BFC3" box="[377,553,395,417]" name="Solomon Islands" pageId="3" pageNumber="1008">Solomon Islands</collectingCountry>
:
<collectingRegion id="4987F82B7557FFD4793065CAFD19BFC3" box="[562,689,395,417]" country="Solomon Islands" name="Guadalcanal" pageId="3" pageNumber="1008">Guadalcanal</collectingRegion>
and Savo Isl.) it is distinguished by a finer abdominal punctation and the male sexual characters, from
<taxonomicName id="4C434D4A7557FFD4784F65E8FB86BFDC" authority="PUTHZ, 1971" authorityName="PUTHZ" authorityYear="1971" box="[845,1070,425,446]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="amor">
<emphasis id="B937EADB7557FFD4784F65E8FC37BFDC" box="[845,927,425,446]" italics="true" pageId="3" pageNumber="1008">S. amor</emphasis>
<bibRefCitation id="EFD24B387557FFD478AA65E8FB86BFDC" author="PUTHZ V" box="[936,1070,425,446]" pageId="3" pageNumber="1008" pagination="447 - 469" refId="ref3136" refString="PUTHZ V. (1971): Revision of the Stenus species of New Guinea. Part I. (Coleoptera: Staphylinidae). - Pacific Insects 13 (3 - 4): 447 - 469." type="journal article" year="1971">PUTHZ, 1971</bibRefCitation>
</taxonomicName>
(Biak Island) by a shorter metatarsomere I. It is distinguished from
<taxonomicName id="4C434D4A7557FFD4785E6587FB86BFBE" authority="PUTHZ, 2016" authorityName="PUTHZ" authorityYear="2016" box="[860,1070,454,476]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="eros">
<emphasis id="B937EADB7557FFD4785E6587FC0ABFBE" box="[860,930,454,476]" italics="true" pageId="3" pageNumber="1008">S. eros</emphasis>
<bibRefCitation id="EFD24B387557FFD478AB6587FB86BFBE" author="PUTHZ V" box="[937,1070,454,476]" pageId="3" pageNumber="1008" pagination="779 - 834" refId="ref3243" refString="PUTHZ V. (2016): Ubersicht uber die Steninen der melanesischen Subregion (Coleoptera, Staphylinidae). - Linzer biologische Beitrage 48 (1): 779 - 834." type="journal article" year="2016">PUTHZ, 2016</bibRefCitation>
</taxonomicName>
(West-Papua: Manokwari Prov.) by the male sexual characters and by a shorter metatarsomere I, from
<taxonomicName id="4C434D4A7557FFD47A956640FD02BC74" authority="PUTHZ, 1969" authorityName="PUTHZ" authorityYear="1969" box="[407,682,512,534]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="aphrodite">
<emphasis id="B937EADB7557FFD47A956640FDBEBC77" box="[407,534,512,534]" italics="true" pageId="3" pageNumber="1008">S. aphrodite</emphasis>
<bibRefCitation id="EFD24B387557FFD479236640FD02BC74" author="PUTHZ V" box="[545,682,512,534]" pageId="3" pageNumber="1008" pagination="525 - 528" refId="ref3102" refString="PUTHZ V. (1969): The first Stenus species from the Solomons (Coleoptera: Staphylinidae). - Pacific Insects 11 (3 - 4): 525 - 528." type="journal article" year="1969">PUTHZ, 1969</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C434D4A7557FFD479BA6640FC3BBC74" authority="LAST, 1970" authorityName="LAST" authorityYear="1970" box="[696,915,513,534]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="fretus">
<emphasis id="B937EADB7557FFD479BA6640FCB8BC74" box="[696,784,513,534]" italics="true" pageId="3" pageNumber="1008">S. fretus</emphasis>
<bibRefCitation id="EFD24B387557FFD4781B6640FC3BBC74" author="LAST H" box="[793,915,513,534]" pageId="3" pageNumber="1008" pagination="347 - 362" refId="ref2965" refString="LAST H. R. (1970): A record of the species of the genus Stenus (Coleoptera, Staphylinidae) with descriptions of new species collected by Dr. R. Hornabrook in New Guinea. - Journal of Natural History 4 / 3: 347 - 362. doi: 10.1080 / 00222937000770321." type="journal article" year="1970">LAST, 1970</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C434D4A7557FFD478C96640FE8DBC51" authority="PUTHZ, 1971" authorityName="PUTHZ" authorityYear="1971" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="cupido">
<emphasis id="B937EADB7557FFD478C96640FB86BC77" box="[971,1070,512,534]" italics="true" pageId="3" pageNumber="1008">S. cupido</emphasis>
<bibRefCitation id="EFD24B387557FFD47BA1665FFE8DBC51" author="PUTHZ V" box="[163,293,542,563]" pageId="3" pageNumber="1008" pagination="447 - 469" refId="ref3136" refString="PUTHZ V. (1971): Revision of the Stenus species of New Guinea. Part I. (Coleoptera: Staphylinidae). - Pacific Insects 13 (3 - 4): 447 - 469." type="journal article" year="1971">PUTHZ, 1971</bibRefCitation>
</taxonomicName>
only by the different male sexual characters.
</paragraph>
<caption id="DF3C66417557FFD47BA1617FFD9FBB08" ID-DOI="http://doi.org/10.5281/zenodo.5277537" ID-Zenodo-Dep="5277537" httpUri="https://zenodo.org/record/5277537/files/figure.png" pageId="3" pageNumber="1008" startId="3.[163,203,1342,1364]" targetBox="[163,1073,579,1331]" targetPageId="3">
<paragraph id="8BFC36C97557FFD47BA1617FFD9FBB08" blockId="3.[163,1071,1342,1387]" pageId="3" pageNumber="1008">
<emphasis id="B937EADB7557FFD47BA1617FFF5EBB31" bold="true" box="[163,246,1342,1364]" pageId="3" pageNumber="1008">Figs 2-5</emphasis>
: Aedeagus of
<taxonomicName id="4C434D4A7557FFD47A8C617EFD84BB31" authority="Mainda &amp; M &amp; M, 2021" authorityName="Mainda &amp; M &amp; M" authorityYear="2021" box="[398,556,1342,1364]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="B937EADB7557FFD47A8C617EFD84BB31" box="[398,556,1342,1364]" italics="true" pageId="3" pageNumber="1008">S. attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457A07557FFD4793B617EFDD3BB36" box="[569,635,1343,1364]" pageId="3" pageNumber="1008" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="B937EADB7557FFD4798C617FFD31BB31" bold="true" box="[654,665,1342,1363]" pageId="3" pageNumber="1008">2</emphasis>
) and
<taxonomicName id="4C434D4A7557FFD479D9617EFC95BB36" authorityName="PUTHZ" authorityYear="2016" box="[731,829,1343,1364]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="anteros">
<emphasis id="B937EADB7557FFD479D9617EFC95BB36" box="[731,829,1343,1364]" italics="true" pageId="3" pageNumber="1008">S. anteros</emphasis>
</taxonomicName>
(
<emphasis id="B937EADB7557FFD47853617FFCF4BB31" bold="true" box="[849,860,1342,1363]" pageId="3" pageNumber="1008">5</emphasis>
); Sternite VIII of
<taxonomicName id="4C434D4A7557FFD47F1F617EFE8CBB08" authority="Mainda &amp; M &amp; M, 2021" authorityName="Mainda &amp; M &amp; M" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="B937EADB7557FFD47F1F617EFE8CBB08" italics="true" pageId="3" pageNumber="1008">S. attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20457A07557FFD47A2B6114FEC3BB08" box="[297,363,1365,1386]" pageId="3" pageNumber="1008" rank="species">nov.sp.</taxonomicNameLabel>
(
<emphasis id="B937EADB7557FFD47A7A6114FE2BBB08" bold="true" box="[376,387,1365,1386]" pageId="3" pageNumber="1008">3</emphasis>
) and
<taxonomicName id="4C434D4A7557FFD47AB56114FDB9BB09" authorityName="PUTHZ" authorityYear="2016" box="[439,529,1365,1387]" class="Insecta" family="Staphylinidae" genus="Stenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1008" phylum="Arthropoda" rank="species" species="anteros">
<emphasis id="B937EADB7557FFD47AB56114FDB9BB09" box="[439,529,1365,1387]" italics="true" pageId="3" pageNumber="1008">S. anteros</emphasis>
</taxonomicName>
(
<emphasis id="B937EADB7557FFD4791C6114FD81BB08" bold="true" box="[542,553,1365,1386]" pageId="3" pageNumber="1008">4</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BFC36C97557FFD47BA161C4FC5EBB90" blockId="3.[163,1070,1412,1522]" pageId="3" pageNumber="1008">E t y m o l o g y: The new species is named in honor of Sir David Attenborough, whose outstanding documentaries have made the beauty of the natural world accessible to millions of people. But they have also shown how vulnerable our planet is and that its biodiversity is critically endangered, if humanity does not change its current course.</paragraph>
</subSubSection>
</treatment>
</document>