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<treatment ID-GBIF-Taxon="154125996" LSID="urn:lsid:plazi:treatment:0B36C0A0BA8AC91A7E3AAD88F4EB5A8A" httpUri="http://treatment.plazi.org/id/0B36C0A0BA8AC91A7E3AAD88F4EB5A8A" lastPageId="141" lastPageNumber="142" pageId="138" pageNumber="139">
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<taxonomicName authority="Dupuis stat. resurr." class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus Dupuis stat. resurr.</taxonomicName>
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<taxonomicName id="DAD5DD4A9DF78B0C370CCD3B5A8A20D4" ID-CoL="76T3W" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
Figs 104A, B, 105
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<normalizedToken id="A2D092A49FA4AD1EB25037477321EB72" originalValue="AD">A-D</normalizedToken>
, 106, 110A
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<subSubSection pageId="138" pageNumber="139" type="reference_group">
<paragraph pageId="138" pageNumber="139">
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
<subSubSection id="126437301C011FDAD3134053B00D7FA7" pageId="138" pageNumber="139" type="reference_group">
<paragraph id="3274E459EC3B39668678389E7A8F88EC" pageId="138" pageNumber="139">
<taxonomicName id="15C2F3033A648BE2AF73825ACF8847FE" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
Dupuis, 1913: 83;
<bibRefCitation pageId="138" pageNumber="139">Csiki 1932</bibRefCitation>
<bibRefCitation id="F198AE032BDA2B8C17728CB7681D5ACC" pageId="138" pageNumber="139">Csiki 1932</bibRefCitation>
: 1369;
<bibRefCitation pageId="138" pageNumber="139">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
<bibRefCitation id="3ED41801FC1353135FDEF2226ED3471A" pageId="138" pageNumber="139">
<normalizedToken id="117ACFDE935F84A718A2FCC9044DA125" originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 379;
<bibRefCitation author="Fedorenko, DN" journalOrPublisher="Russian Entomological Journal" pageId="150" pageNumber="151" pagination="303 - 312" title="A new species of Pericalus s. str. (Coleoptera: Carabidae, Lebiini) from Vietnam, with notes on some Oriental members of the subgenus." url="http://kmkjournals.com/upload/PDF/REJ/26/ent26_4_303_312_Fedorenko_for_Inet.pdf" volume="26" year="2017">Fedorenko 2017</bibRefCitation>
<bibRefCitation id="DCCAF2B99E0D73FA86149840F82531A5" author="Fedorenko, DN" journalOrPublisher="Russian Entomological Journal" pageId="150" pageNumber="151" pagination="303 - 312" title="A new species of Pericalus s. str. (Coleoptera: Carabidae, Lebiini) from Vietnam, with notes on some Oriental members of the subgenus." url="http://kmkjournals.com/upload/PDF/REJ/26/ent26_4_303_312_Fedorenko_for_Inet.pdf" volume="26" year="2017">Fedorenko 2017</bibRefCitation>
: 311.
</paragraph>
<paragraph pageId="138" pageNumber="139">
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus ornatus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="subspecies" species="ornatus" subspecies="formosanus">Pericalus ornatus formosanus</taxonomicName>
<paragraph id="F9B06422E9B824879A94D5850DE63361" pageId="138" pageNumber="139">
<taxonomicName id="040D67077B0A58F10AB06C0E57BD2500" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus ornatus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="subspecies" species="ornatus" subspecies="formosanus">Pericalus ornatus formosanus</taxonomicName>
Dupuis:
<bibRefCitation author="Shi, H" journalOrPublisher="G Haase Soehne, Prague" pageId="153" pageNumber="154" url="https://doi.org/10.3897/zookeys.758.24169" year="2018">Shi and Liang 2018</bibRefCitation>
<bibRefCitation id="515CF3915DF079E3E02ECD4B74064BE9" author="Shi, H" journalOrPublisher="G Haase Soehne, Prague" pageId="153" pageNumber="154" url="https://doi.org/10.3897/zookeys.758.24169" year="2018">Shi and Liang 2018</bibRefCitation>
: 36.
</paragraph>
</subSubSection>
<subSubSection pageId="138" pageNumber="139" type="type material">
<paragraph pageId="138" pageNumber="139">Types and other material examined.</paragraph>
<paragraph pageId="138" pageNumber="139">
<subSubSection id="598ADF3C146FCEE6DEEC38015FB2FB07" pageId="138" pageNumber="139" type="type material">
<paragraph id="5C9899AF8C89534C81A0694FDA66B31A" pageId="138" pageNumber="139">Types and other material examined.</paragraph>
<paragraph id="550CE81CFB796E05C9802E85E9A86903" pageId="138" pageNumber="139">
Holotype (male) labeled &quot;Chip Chip/ II Formosa/Sauter 07-09&quot;;
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[rectangular, red paper]; &quot;
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
<taxonomicName id="D6CC914F6412C87D4F301169A4F80AFA" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
Dupuis/Dupuis det. &quot;;
<normalizedToken originalValue="“Syntypus”">&quot;Syntypus&quot;</normalizedToken>
<normalizedToken id="1A7AD28D907E25068C16E0B83BD2642D" originalValue="“Syntypus”">&quot;Syntypus&quot;</normalizedToken>
; &quot;DEI
<taxonomicName lsidName="" order="Coleoptera" pageId="138" pageNumber="139" rank="order">Coleoptera</taxonomicName>
<taxonomicName id="6EF8E156DDA56124034DA022EB97A75C" class="Insecta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
/# 200415&quot;. Five paratypes and 151 other specimens of
<taxonomicName lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
<taxonomicName id="7EEC71B870317B7D226A3820732A3B6A" lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
: 75 males and 76 females. For further details see EH Strickland Virtual Entomology Museum Database.
</paragraph>
</subSubSection>
<subSubSection pageId="138" pageNumber="139" type="type locality">
<paragraph pageId="138" pageNumber="139">Type locality.</paragraph>
<paragraph pageId="138" pageNumber="139">
<subSubSection id="5D5E445A5A9345910CA997455163884F" pageId="138" pageNumber="139" type="type locality">
<paragraph id="970BFE9497BD552E01556EF21C718CF3" pageId="138" pageNumber="139">Type locality.</paragraph>
<paragraph id="AFBACEE7F251F3066F4E4D17F7169C5F" pageId="138" pageNumber="139">
Taiwan. Kaoshiung County,
<normalizedToken originalValue="“Hoozan”">&quot;Hoozan&quot;</normalizedToken>
<normalizedToken id="196D10AF8AC3241A86E9C1918A443171" originalValue="“Hoozan”">&quot;Hoozan&quot;</normalizedToken>
= Fengshan City.
</paragraph>
</subSubSection>
<subSubSection pageId="138" pageNumber="139" type="taxonomic notes">
<paragraph pageId="138" pageNumber="139">Taxonomic notes.</paragraph>
<paragraph pageId="138" pageNumber="139">
<subSubSection id="B7EF86F10FF4F254BDCB1DCF8550768D" pageId="138" pageNumber="139" type="taxonomic notes">
<paragraph id="D00648F1112B297EF046AAC946307121" pageId="138" pageNumber="139">Taxonomic notes.</paragraph>
<paragraph id="852730820F80A1EA87D90AC72B99D815" pageId="138" pageNumber="139">
In a recent paper
<bibRefCitation author="Shi, H" journalOrPublisher="G Haase Soehne, Prague" pageId="153" pageNumber="154" url="https://doi.org/10.3897/zookeys.758.24169" year="2018">Shi and Liang (2018)</bibRefCitation>
<bibRefCitation id="9361DF1BAF0DA3D4CA4F295FCDD326FB" author="Shi, H" journalOrPublisher="G Haase Soehne, Prague" pageId="153" pageNumber="154" url="https://doi.org/10.3897/zookeys.758.24169" year="2018">Shi and Liang (2018)</bibRefCitation>
considered
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<taxonomicName id="9DE1654E6467A72E1FD00001FC1043B5" lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
to be a subspecies of
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus ornatus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="ornatus">Pericalus ornatus</taxonomicName>
<taxonomicName id="3EADB9EA27B88716F4BB36E585ADA27C" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus ornatus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="ornatus">Pericalus ornatus</taxonomicName>
Schmidt-Goebel, 1846. While these taxa are similar, we have deduced that there are sufficient differences to maintain the species status of
<taxonomicName lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
<taxonomicName id="EB4EAA29C5A58CFFCE1CA2409EFE971D" lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
. Along with differences in the hind angles of the elytra, and number of discal setae (one in
<taxonomicName lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
<taxonomicName id="4C3173EEFBB09E3D4B40E7E6E5996537" lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
and two in
<taxonomicName lsidName="P. ornatus" pageId="138" pageNumber="139" rank="species" species="ornatus">P. ornatus</taxonomicName>
<taxonomicName id="107F5D279FD086A9009679D72148EF6A" lsidName="P. ornatus" pageId="138" pageNumber="139" rank="species" species="ornatus">P. ornatus</taxonomicName>
), there are slight but consistent differences in the genitalic characteristics. Females of
<taxonomicName lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
<taxonomicName id="62DBE46A17AC4F08B6A3D30F32DBF638" lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
have a spermathecal gland duct that is longer and a spermathecal gland (Fig. 110B) that is larger than seen in specimens of
<taxonomicName lsidName="P. ornatus" pageId="138" pageNumber="139" rank="species" species="ornatus">P. ornatus</taxonomicName>
<taxonomicName id="36A0E7C52CA147CF8DF7CAF7B441B3D7" lsidName="P. ornatus" pageId="138" pageNumber="139" rank="species" species="ornatus">P. ornatus</taxonomicName>
. Males of
<taxonomicName lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
<taxonomicName id="A8A7F4AC27862B49A326927794E397DB" lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
have a phallus with ostium more dorsally situated than in members of
<taxonomicName lsidName="P. ornatus" pageId="138" pageNumber="139" rank="species" species="ornatus">P. ornatus</taxonomicName>
<taxonomicName id="EE87340A5F36FA2B0DC22DB23D4725D0" lsidName="P. ornatus" pageId="138" pageNumber="139" rank="species" species="ornatus">P. ornatus</taxonomicName>
and an endophallus that is obviously narrower in form and has only one prominent endophallic lobe (Fig. 108A), located more basally than the most basal endophallic lobe of specimens of
<taxonomicName lsidName="P. ornatus" pageId="138" pageNumber="139" rank="species" species="ornatus">P. ornatus</taxonomicName>
<taxonomicName id="8F69A579A4E0758155637D6104F239AC" lsidName="P. ornatus" pageId="138" pageNumber="139" rank="species" species="ornatus">P. ornatus</taxonomicName>
.
</paragraph>
<paragraph pageId="138" pageNumber="139">
<paragraph id="E1519C9F39165903B7A62AD3058E352B" pageId="138" pageNumber="139">
Shi and Liang observed that
<bibRefCitation author="Fedorenko, DN" journalOrPublisher="Russian Entomological Journal" pageId="150" pageNumber="151" pagination="303 - 312" title="A new species of Pericalus s. str. (Coleoptera: Carabidae, Lebiini) from Vietnam, with notes on some Oriental members of the subgenus." url="http://kmkjournals.com/upload/PDF/REJ/26/ent26_4_303_312_Fedorenko_for_Inet.pdf" volume="26" year="2017">Fedorenko (2017)</bibRefCitation>
<bibRefCitation id="C7A8924D1209AF00304E3F72C152CF6B" author="Fedorenko, DN" journalOrPublisher="Russian Entomological Journal" pageId="150" pageNumber="151" pagination="303 - 312" title="A new species of Pericalus s. str. (Coleoptera: Carabidae, Lebiini) from Vietnam, with notes on some Oriental members of the subgenus." url="http://kmkjournals.com/upload/PDF/REJ/26/ent26_4_303_312_Fedorenko_for_Inet.pdf" volume="26" year="2017">Fedorenko (2017)</bibRefCitation>
, recorded
<taxonomicName lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
<taxonomicName id="53622A913D00EB26BBB08E6D7060E9D9" lsidName="P. formosanus" pageId="138" pageNumber="139" rank="species" species="formosanus">P. formosanus</taxonomicName>
from Vietnam based on misidentification of
<taxonomicName lsidName="P. acutidens" pageId="138" pageNumber="139" rank="species" species="acutidens">P. acutidens</taxonomicName>
<taxonomicName id="D684DED7263A3A7ACFBE826647EEE4A6" lsidName="P. acutidens" pageId="138" pageNumber="139" rank="species" species="acutidens">P. acutidens</taxonomicName>
. After examining his images of the everted endophallus, it is clear that it was a mistaken identity.
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
<taxonomicName id="07A5F9A4A245BE5CA9F32CE0619CCFC5" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
is restricted to Taiwan and allopatric with all other species of
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="genus">Pericalus</taxonomicName>
<taxonomicName id="75BFE07EF80CCDF5F9F46D8FFFE57421" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="genus">Pericalus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="138" pageNumber="139" type="diagnosis">
<paragraph pageId="138" pageNumber="139">Diagnosis.</paragraph>
<paragraph pageId="138" pageNumber="139">Specimens of this species are easily distinguished from other Taiwanese pericalines by having smooth tarsal claws, two pairs of supraorbital setae, two pairs of latero-marginal setae on the pronotum and a black dorsal coloration with eight maculae on the disc of the elytra.</paragraph>
<subSubSection id="B6906395E967C42AE6D12FA32B2A123B" pageId="138" pageNumber="139" type="diagnosis">
<paragraph id="38EF4F258D706648112E12CB2438D134" pageId="138" pageNumber="139">Diagnosis.</paragraph>
<paragraph id="5C9CA23FFFE2816D5EB1DC7955B49F11" pageId="138" pageNumber="139">Specimens of this species are easily distinguished from other Taiwanese pericalines by having smooth tarsal claws, two pairs of supraorbital setae, two pairs of latero-marginal setae on the pronotum and a black dorsal coloration with eight maculae on the disc of the elytra.</paragraph>
</subSubSection>
<subSubSection lastPageId="140" lastPageNumber="141" pageId="138" pageNumber="139" type="redescription">
<paragraph pageId="138" pageNumber="139">Redescription.</paragraph>
<paragraph pageId="138" pageNumber="139">OBL 7.67 - 11.33 mm. Length (n = ten males, ten females): head 0.92 - 1.24, pronotum 1.28 - 2.12, elytra 4.33 - 6.33, metepisternum 1.04 - 1.40 mm; width: head 2.14 - 3.10, pronotum 3.33 - 4.67, elytra 3.33 - 4.67, metepisternum 0.56 - 0.84 mm.</paragraph>
<paragraph pageId="138" pageNumber="139">Body proportions. HW/HL 2.24 - 2.58; PWM/PL 1.45 - 1.58; EL/EW 1.25 - 1.46; ML/MW 1.40 - 1.94.</paragraph>
<paragraph lastPageId="139" lastPageNumber="140" pageId="138" pageNumber="139">
<subSubSection id="06985D0E39D7D10204644D76253C3120" lastPageId="140" lastPageNumber="141" pageId="138" pageNumber="139" type="redescription">
<paragraph id="DCA8B9B3AC064A15C745F7D8FB4CD56E" pageId="138" pageNumber="139">Redescription.</paragraph>
<paragraph id="3C595249EF7854EFB0B08CC9D6794E48" pageId="138" pageNumber="139">OBL 7.67 - 11.33 mm. Length (n = ten males, ten females): head 0.92 - 1.24, pronotum 1.28 - 2.12, elytra 4.33 - 6.33, metepisternum 1.04 - 1.40 mm; width: head 2.14 - 3.10, pronotum 3.33 - 4.67, elytra 3.33 - 4.67, metepisternum 0.56 - 0.84 mm.</paragraph>
<paragraph id="29180CA63B94E4B9FA74648F6C96C836" pageId="138" pageNumber="139">Body proportions. HW/HL 2.24 - 2.58; PWM/PL 1.45 - 1.58; EL/EW 1.25 - 1.46; ML/MW 1.40 - 1.94.</paragraph>
<paragraph id="427DC4EEC298F61A638759D81ABA9385" lastPageId="139" lastPageNumber="140" pageId="138" pageNumber="139">
Color. Fig. 104A, B. Various. Dorsum of head piceous, clypeus piceous with rufo-brunneous apical edge, antennae and palpi rufo-brunneous, scape sometimes darker; pronotum piceous; elytral disc piceous, with eight testaceous maculae, two in basal 1/3, two near mid-length, and four in apical 1/3, basal macula somewhat dentate, from interval 3 or 4 to interval 6 or 7, always closest to base in interval 5 and closest to apex
<pageBreakToken pageId="139" pageNumber="140" start="start">in</pageBreakToken>
<pageBreakToken id="13B0EFDA556D3DE35E61514A252F8A4F" pageId="139" pageNumber="140" start="start">in</pageBreakToken>
interval 4, mid-length macula from interval 4 to interval 5 or 6, apical macula with small patch from interval 2 to 3 and small patch in interval 7; ventral surface rufo-piceous to piceous; legs rufo-brunneous, tibia darker.
</paragraph>
<caption pageId="139" pageNumber="140">
<paragraph pageId="139" pageNumber="140">
<caption id="4CF5ADCFEFE910B9C9F90659083C4C0C" pageId="139" pageNumber="140">
<paragraph id="B85B07B20E6ECCC3266AF65FDAD8B79D" pageId="139" pageNumber="140">
Figure 104. Dorsal habitus and intrapopulation variation of color pattern of
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
<taxonomicName id="FA944E85EDED4E61763582FA04474CE8" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
Dupuis. A larger elytral macula (common) (OBL 11.02 mm). B smaller elytral macula (OBL 8.02 mm).
</paragraph>
</caption>
<paragraph pageId="139" pageNumber="140">Microsculpture. Head with microsculpture almost isodiametric to isodiametric; pronotum with microsculpture shallow, somewhat transverse to transverse; elytra with sculpticells transverse, single row of isodiametric cells down center of each stria; ventral surface with transverse to almost isodiameteric microsculpture.</paragraph>
<paragraph lastPageId="140" lastPageNumber="141" pageId="139" pageNumber="140">
<paragraph id="D1CEFD69839FC0C45787A549A0B3CCE8" pageId="139" pageNumber="140">Microsculpture. Head with microsculpture almost isodiametric to isodiametric; pronotum with microsculpture shallow, somewhat transverse to transverse; elytra with sculpticells transverse, single row of isodiametric cells down center of each stria; ventral surface with transverse to almost isodiameteric microsculpture.</paragraph>
<paragraph id="8B9D2535AE69221B625D4709BD3E0C89" lastPageId="140" lastPageNumber="141" pageId="139" pageNumber="140">
Macrosculpture and pilosity. Dorsum of head and base of clypeus rugulose, surface with very fine, scattered, setigerous punctures, hardly visible at 50
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="0C9FCE4DFCF739B550FA895301297B32" originalValue="×">x</normalizedToken>
; pronotum disc
<pageBreakToken pageId="140" pageNumber="141" start="start">faintly</pageBreakToken>
<pageBreakToken id="2FEEAA51A973E1BCB0996A90E52EB809" pageId="140" pageNumber="141" start="start">faintly</pageBreakToken>
rugulose centrally, more so along lateral margins, surface with very fine, scattered, setigerous punctures; elytral intervals convex, interval 7 slightly more raised than others in apical 1/3, entire dorsal surface with fine, scattered setigerous punctures, punctures hardly visible but setae easily viewed in lateral view at 50
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="B4E110C3FCB1CD6E07AD7FB869D62D25" originalValue="×">x</normalizedToken>
, striae impunctate; ventral surface of head rugulose to gula suture, remaining ventral surface with randomly scattered setigerous punctures.
</paragraph>
<paragraph pageId="140" pageNumber="141">Fixed setae. Two pairs of supraorbital setae; clypeus with two long, lateral setae; labrum with six setae along apical margin; one pair of suborbital setae; pronotum with two pairs of setae, one at base of lateral margin, one on lateral margin at pronotum max width; 24 - 25 lateral (umbilical) setae in interval 9; elytra with interval 3 with one seta in basal 1/6 (See also, variation); ventral surface with two setae on each of abdominal sterna III to VI; four setae along apical margin of sternum VII.</paragraph>
<paragraph id="5A798EF3F9603AECAFCA018266BFE97F" pageId="140" pageNumber="141">Fixed setae. Two pairs of supraorbital setae; clypeus with two long, lateral setae; labrum with six setae along apical margin; one pair of suborbital setae; pronotum with two pairs of setae, one at base of lateral margin, one on lateral margin at pronotum max width; 24 - 25 lateral (umbilical) setae in interval 9; elytra with interval 3 with one seta in basal 1/6 (See also, variation); ventral surface with two setae on each of abdominal sterna III to VI; four setae along apical margin of sternum VII.</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="141" type="variation">
<paragraph pageId="140" pageNumber="141">Variation.</paragraph>
<paragraph pageId="140" pageNumber="141">of 151 specimens observed only one was observed to have two setae in interval 3, the typical apical setae and one at mid-length; see Fig. 104A (two setae) vs. Fig. 104B (one seta).</paragraph>
<paragraph pageId="140" pageNumber="141">Luster. Dorsal surface moderately glossy; ventral surface glossy.</paragraph>
<paragraph pageId="140" pageNumber="141">Head. Mandibles long, left mandible with distinctive notch on inside and dorsal surface in apical 1/3; labrum deeply bilobed, mentum without tooth; eyes large, convex; palpi cylindrical and elongate and with fine setae.</paragraph>
<paragraph pageId="140" pageNumber="141">Pronotum. Lateral margins explanate, with margins curved slightly upwards, anterior transverse impression moderately shallow, posterior transverse impression deep, median longitudinal impression moderately deep, apico-lateral margins broadly and distinctly lobed, posterio-lateral margins sinuate, right-angled.</paragraph>
<paragraph pageId="140" pageNumber="141">Elytra. Hind angles slightly sinuate, apex of lateral margin with distinctly sharp edge, pointed.</paragraph>
<paragraph pageId="140" pageNumber="141">Hind wings. Macropterous.</paragraph>
<paragraph pageId="140" pageNumber="141">Legs. Tarsal claws smooth, rather long and slender, males with adhesive vestiture ventrally, two rows squamo-setae on tarsomeres 1-3 of fore-leg.</paragraph>
<paragraph pageId="140" pageNumber="141">
<subSubSection id="2F55BC169A79C07EEEC591BCBC8A4D8C" pageId="140" pageNumber="141" type="variation">
<paragraph id="E0E77B3637CE6DA9480595D3FE37850E" pageId="140" pageNumber="141">Variation.</paragraph>
<paragraph id="DAFE13CCB5C4D2DEB1B835D32D1D9F5E" pageId="140" pageNumber="141">of 151 specimens observed only one was observed to have two setae in interval 3, the typical apical setae and one at mid-length; see Fig. 104A (two setae) vs. Fig. 104B (one seta).</paragraph>
<paragraph id="24100594B0FE67A7646DF1451DBED608" pageId="140" pageNumber="141">Luster. Dorsal surface moderately glossy; ventral surface glossy.</paragraph>
<paragraph id="C5949892D2B04A8249641377FF28F312" pageId="140" pageNumber="141">Head. Mandibles long, left mandible with distinctive notch on inside and dorsal surface in apical 1/3; labrum deeply bilobed, mentum without tooth; eyes large, convex; palpi cylindrical and elongate and with fine setae.</paragraph>
<paragraph id="4C8FBBDB1B54F7C7E1890B05C1440ABE" pageId="140" pageNumber="141">Pronotum. Lateral margins explanate, with margins curved slightly upwards, anterior transverse impression moderately shallow, posterior transverse impression deep, median longitudinal impression moderately deep, apico-lateral margins broadly and distinctly lobed, posterio-lateral margins sinuate, right-angled.</paragraph>
<paragraph id="B3346C2784C5C0911607FDB1DAF771D5" pageId="140" pageNumber="141">Elytra. Hind angles slightly sinuate, apex of lateral margin with distinctly sharp edge, pointed.</paragraph>
<paragraph id="44B41B1F9CB60AA6A38D317031391972" pageId="140" pageNumber="141">Hind wings. Macropterous.</paragraph>
<paragraph id="4C2ED07AF899A26A112B23C92005B959" pageId="140" pageNumber="141">Legs. Tarsal claws smooth, rather long and slender, males with adhesive vestiture ventrally, two rows squamo-setae on tarsomeres 1-3 of fore-leg.</paragraph>
<paragraph id="9FC2FDCFA3E3A7CF63DA686DE62E71C7" pageId="140" pageNumber="141">
Male genitalia. Fig. 105
<normalizedToken originalValue="AD">A-D</normalizedToken>
<normalizedToken id="FFAE5F02E4F498B64F16E321F249AC5D" originalValue="AD">A-D</normalizedToken>
. Length 1.80 - 2.06 mm. Ostium catopic, open slightly more on the left side. Phallus cylindrical, widest at mid-length, apex short, rounded at tip; endophallus relatively long and narrow along length, with single distinctive lobe (el) at base.
</paragraph>
<caption pageId="140" pageNumber="141">
<paragraph pageId="140" pageNumber="141">
<caption id="58C93E3CE9B4DF50FFA7A818F51ECDCF" pageId="140" pageNumber="141">
<paragraph id="4A327AC31AE26FEB678058B8E5734F35" pageId="140" pageNumber="141">
Figure 105. Digital images of male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
<taxonomicName id="06022C74316E571FCCA507D41182ECD3" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
Dupuis. A left lateral aspect, endophallus everted B right lateral aspect C ventral aspect D left lateral aspect. Legend: el endophallic lobe.
</paragraph>
</caption>
<paragraph pageId="140" pageNumber="141">Female genitalia. Fig. 110A. Width 1.15 - 1.30 mm. Gonocoxite 2 (gc2) distinctively long and narrow, relatively uniform in width along length; three lateral ensiform setae spaced widely apart (les), one dorsal ensiform seta. Sensory furrow, furrow pegs, and associated nematiform setae not observed. One spermatheca present (sp1), elongate and cylindrical, expanding slightly in apical half; one spermathecal accessory gland (sg), associated spermathecal gland duct (sgd), with attachment site just before widening of spermatheca.</paragraph>
<paragraph id="9DC21E6605EE7D6FAD90D35312C6D80D" pageId="140" pageNumber="141">Female genitalia. Fig. 110A. Width 1.15 - 1.30 mm. Gonocoxite 2 (gc2) distinctively long and narrow, relatively uniform in width along length; three lateral ensiform setae spaced widely apart (les), one dorsal ensiform seta. Sensory furrow, furrow pegs, and associated nematiform setae not observed. One spermatheca present (sp1), elongate and cylindrical, expanding slightly in apical half; one spermathecal accessory gland (sg), associated spermathecal gland duct (sgd), with attachment site just before widening of spermatheca.</paragraph>
</subSubSection>
<subSubSection lastPageId="141" lastPageNumber="142" pageId="140" pageNumber="141" type="habitat">
<paragraph pageId="140" pageNumber="141">Habitat, habits, and seasonal occurrence.</paragraph>
<paragraph lastPageId="141" lastPageNumber="142" pageId="140" pageNumber="141">
<subSubSection id="C970E26F47880917AF653AC7952978C6" lastPageId="141" lastPageNumber="142" pageId="140" pageNumber="141" type="habitat">
<paragraph id="0E3BA68459BB7F8C5D1A3F37EAEFA98D" pageId="140" pageNumber="141">Habitat, habits, and seasonal occurrence.</paragraph>
<paragraph id="1A0A4207C1A62835CC1DE3356D5A3893" lastPageId="141" lastPageNumber="142" pageId="140" pageNumber="141">
The known elevational range of
<taxonomicName lsidName="P. formosanus" pageId="140" pageNumber="141" rank="species" species="formosanus">P. formosanus</taxonomicName>
<taxonomicName id="3C4B60E7B099A96AF63FFD3869E65838" lsidName="P. formosanus" pageId="140" pageNumber="141" rank="species" species="formosanus">P. formosanus</taxonomicName>
is from 500 to 2095 meters with the majority of adults being collected between 1800 and 2000 meters. Adults of this species are crepuscular and are found in mixed primary and secondary forest of montane areas, typically in moist areas.
<pageBreakToken pageId="141" pageNumber="142" start="start">They</pageBreakToken>
<pageBreakToken id="BC00973C5F33546DE4B6457D1A69ABA1" pageId="141" pageNumber="142" start="start">They</pageBreakToken>
can be found inside and under deadwood during the day and on deadwood at night. Specimens are easily captured as they do not run or fly when illuminated. Specimens have been collected throughout the year with most collected from May to July. Methods of collecting include light trap on ground, u.v. light sheet, pitfall trap, and hand collecting.
</paragraph>
</subSubSection>
<subSubSection pageId="141" pageNumber="142" type="distribution">
<paragraph pageId="141" pageNumber="142">Geographical distribution.</paragraph>
<paragraph pageId="141" pageNumber="142">
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
<subSubSection id="AEA4505E2232309005867C5DF82375BA" pageId="141" pageNumber="142" type="distribution">
<paragraph id="9988640EF822A3D8BBA4C5F99E79F6D6" pageId="141" pageNumber="142">Geographical distribution.</paragraph>
<paragraph id="FC4B430CA406495C789108A61EE3891B" pageId="141" pageNumber="142">
<taxonomicName id="486871A6804142538EA301F0FA3C0D3D" authority="Dupuis, 1913" authorityName="Dupuis" authorityYear="1913" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus formosanus" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="formosanus">Pericalus formosanus</taxonomicName>
is known only from Taiwan. See Figure 106.
</paragraph>
<caption pageId="141" pageNumber="142">
<paragraph pageId="141" pageNumber="142">
<caption id="97300A12E642D005351B6E151C078737" pageId="141" pageNumber="142">
<paragraph id="D6D2B76A459B64FEBF0681B0CEEAF8CC" pageId="141" pageNumber="142">
Figure 106. Map showing known localities for species of the genus
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="genus">Pericalus</taxonomicName>
<taxonomicName id="7F3E9C2127CF291B1A8661C497B45F2B" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="genus">Pericalus</taxonomicName>
MacLeay, in Taiwan.
</paragraph>
</caption>

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Subgenus
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<taxonomicName id="24C3A731066A786F03C5180D76629282" ID-CoL="8MH5X" authority="s. str." class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="(Pericalus)" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="subGenus" subGenus="Pericalus">Pericalus s. str.</taxonomicName>
</paragraph>
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<paragraph pageId="138" pageNumber="139">
<taxonomicName class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="genus">Pericalus</taxonomicName>
<subSubSection id="4F75DA403BFBD1CA501459D35DC87B7F" pageId="138" pageNumber="139" type="reference_group">
<paragraph id="33DFCDA9371094811A2F7BF355387954" pageId="138" pageNumber="139">
<taxonomicName id="A3C1A5997C9E9D457BC14DB81F977DBA" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="genus">Pericalus</taxonomicName>
MacLeay, 1825: 15; Schmidt-Goebel 1846: 85;
<bibRefCitation pageId="138" pageNumber="139">Lacordaire 1854</bibRefCitation>
<bibRefCitation id="8A080D89ED244A1B5693D77BBD0A0849" pageId="138" pageNumber="139">Lacordaire 1854</bibRefCitation>
: 147;
<bibRefCitation pageId="138" pageNumber="139">Schaum 1860</bibRefCitation>
<bibRefCitation id="5765514CC48DF35318248B07270389E9" pageId="138" pageNumber="139">Schaum 1860</bibRefCitation>
: 190;
<bibRefCitation author="Chaudoir, M de" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="149" pageNumber="150" pagination="116 - 131" title="Beitrag zur Kenntnis einiger Carabicinen-Gattungen." volume="5" year="1861">Chaudoir 1861</bibRefCitation>
<bibRefCitation id="079FEB0117E79FB899A59E3A5AEAD71E" author="Chaudoir, M de" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="149" pageNumber="150" pagination="116 - 131" title="Beitrag zur Kenntnis einiger Carabicinen-Gattungen." volume="5" year="1861">Chaudoir 1861</bibRefCitation>
: 123;
<bibRefCitation pageId="138" pageNumber="139">
<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
<bibRefCitation id="0F21D3A541E04D92D430ACFB12713C6B" pageId="138" pageNumber="139">
<normalizedToken id="9ED90DA4D3E5930C3A546238B01995FD" originalValue="Jedlička">Jedlicka</normalizedToken>
1963
</bibRefCitation>
: 373;
<bibRefCitation author="Baehr, M" journalOrPublisher="Spixiana" pageId="148" pageNumber="149" pagination="33 - 39" title="Review of the Pericalusguttatus-complex (Insecta, Coleoptera, Carabidae, Lebiinae)." url="https://www.biodiversitylibrary.org/part/66549" volume="23" year="2000">Baehr 2000</bibRefCitation>
<bibRefCitation id="DA4FCFD157A013F452C04A19459BA309" author="Baehr, M" journalOrPublisher="Spixiana" pageId="148" pageNumber="149" pagination="33 - 39" title="Review of the Pericalusguttatus-complex (Insecta, Coleoptera, Carabidae, Lebiinae)." url="https://www.biodiversitylibrary.org/part/66549" volume="23" year="2000">Baehr 2000</bibRefCitation>
: 33;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). Second ed. Germany" pageId="152" pageNumber="153" title="Systematic List of Extant Ground Beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="B3C20D59006F31AC01589C244879113C" author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). Second ed. Germany" pageId="152" pageNumber="153" title="Systematic List of Extant Ground Beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
: 455;
<bibRefCitation author="Fedorenko, DN" journalOrPublisher="Russian Entomological Journal" pageId="150" pageNumber="151" pagination="303 - 312" title="A new species of Pericalus s. str. (Coleoptera: Carabidae, Lebiini) from Vietnam, with notes on some Oriental members of the subgenus." url="http://kmkjournals.com/upload/PDF/REJ/26/ent26_4_303_312_Fedorenko_for_Inet.pdf" volume="26" year="2017">Fedorenko 2017</bibRefCitation>
<bibRefCitation id="E89BA5B0C9C909485AD933960DC73433" author="Fedorenko, DN" journalOrPublisher="Russian Entomological Journal" pageId="150" pageNumber="151" pagination="303 - 312" title="A new species of Pericalus s. str. (Coleoptera: Carabidae, Lebiini) from Vietnam, with notes on some Oriental members of the subgenus." url="http://kmkjournals.com/upload/PDF/REJ/26/ent26_4_303_312_Fedorenko_for_Inet.pdf" volume="26" year="2017">Fedorenko 2017</bibRefCitation>
: 303;
<bibRefCitation author="Hongliang, S" journalOrPublisher="Annales de la Societe entomologique de Belgique" pageId="151" pageNumber="152" url="https://doi.org/10.3897/zookeys.758.24169" year="2018">Hongliang and Hongbin 2018</bibRefCitation>
<bibRefCitation id="FC0DBE0DD8C9530DE66BF3398F715C68" author="Hongliang, S" journalOrPublisher="Annales de la Societe entomologique de Belgique" pageId="151" pageNumber="152" url="https://doi.org/10.3897/zookeys.758.24169" year="2018">Hongliang and Hongbin 2018</bibRefCitation>
: 19.
</paragraph>
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<paragraph pageId="138" pageNumber="139">Type species.</paragraph>
<paragraph pageId="138" pageNumber="139">
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<subSubSection id="FAFC3E551A000E00359A23D59DF89588" pageId="138" pageNumber="139" type="type species">
<paragraph id="40C357CC50B1F8EBD40B0C2397C2C2D9" pageId="138" pageNumber="139">Type species.</paragraph>
<paragraph id="3E275485A6689C6AD9DFA5FAC941E92D" pageId="138" pageNumber="139">
<taxonomicName id="135F689A9C012EFA11AF49D1770D0F4D" class="Insecta" family="Carabidae" genus="Pericalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pericalus cicindeloides" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="cicindeloides">Pericalus cicindeloides</taxonomicName>
MacLeay, 1825 (monobasic).
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@ -1,121 +1,120 @@
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Subgenus
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Kirby, 1825: 94-97;
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<normalizedToken id="64896459EA657B74F2A8FAF50C160198" originalValue="Schmidt-Göbel">Schmidt-Goebel</normalizedToken>
1846
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: 80;
<bibRefCitation pageId="32" pageNumber="33">Lacordaire 1854</bibRefCitation>
<bibRefCitation id="73ECCEEADBE297FCA08195240EB851BF" pageId="32" pageNumber="33">Lacordaire 1854</bibRefCitation>
: 145;
<bibRefCitation author="Chaudoir, M de" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="149" pageNumber="150" pagination="116 - 131" title="Beitrag zur Kenntnis einiger Carabicinen-Gattungen." volume="5" year="1861">Chaudoir 1861</bibRefCitation>
<bibRefCitation id="322D20C575CCA077982BDAC055811C82" author="Chaudoir, M de" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="149" pageNumber="150" pagination="116 - 131" title="Beitrag zur Kenntnis einiger Carabicinen-Gattungen." volume="5" year="1861">Chaudoir 1861</bibRefCitation>
: 116;
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</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="33" type="type species">
<paragraph pageId="32" pageNumber="33">Type species.</paragraph>
<paragraph pageId="32" pageNumber="33">
<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus hardwickii" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hardwickii">Catascopus hardwickii</taxonomicName>
<subSubSection id="382D9954107EF1D9B7BF54C9E8F949BB" pageId="32" pageNumber="33" type="type species">
<paragraph id="3A67D9A78620FD437D2B52E138D93AC5" pageId="32" pageNumber="33">Type species.</paragraph>
<paragraph id="C161AF19EA398FC6527632952BE6A14B" pageId="32" pageNumber="33">
<taxonomicName id="ECF35A64A9DBB9A04F987C52D0737E45" class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus hardwickii" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hardwickii">Catascopus hardwickii</taxonomicName>
Kirby, 1825 (monotypic).
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="33" type="type locality">
<paragraph pageId="32" pageNumber="33">Type locality.</paragraph>
<paragraph pageId="32" pageNumber="33">India.</paragraph>
<subSubSection id="BDCB44B551684C12DCFCE134D4204F45" pageId="32" pageNumber="33" type="type locality">
<paragraph id="B0A82331F004F62BA68A7185DBE5C795" pageId="32" pageNumber="33">Type locality.</paragraph>
<paragraph id="D7694B0662FE7E70F55AD19E6B3C7478" pageId="32" pageNumber="33">India.</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="33" type="recognition of taiwanese species of catascopus">
<paragraph pageId="32" pageNumber="33">
<subSubSection id="20E6E1EFA4127B4808D3360DFB9BB964" pageId="32" pageNumber="33" type="recognition of taiwanese species of catascopus">
<paragraph id="AE989236924A6537A7B64281076DC42E" pageId="32" pageNumber="33">
Recognition of Taiwanese species of
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<taxonomicName id="BA7940EB2344A87742EB334A156CC3FA" class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Catascopus</taxonomicName>
s. str.
</paragraph>
<paragraph pageId="32" pageNumber="33">Color. Various.</paragraph>
<paragraph pageId="32" pageNumber="33">Fixed setae. Two pairs of supraorbital setae; clypeus with two lateral setae; labrum with six setae along apical margin; one pair of suborbital setae.; 16-17 lateral (umbilical) setae in interval 9; two setae on each of abdominal sterna III to VI, two setae along apical margin of sternum VII in males, four setae along apical margin of sternum VII in females.</paragraph>
<paragraph pageId="32" pageNumber="33">Head. Mandibles curved, left mandible slightly more curved and pointed at apex then right mandible, right mandible with additional, small tooth on inside cutting surface near mid-length of mandible; labrum bilobed; mentum with single broad tooth; eyes convex; palpi cylindrical, elongate.</paragraph>
<paragraph pageId="32" pageNumber="33">Elytra. Humeri broadly rounded; elytral margin shallowly impressed in basal 1/3.</paragraph>
<paragraph pageId="32" pageNumber="33">Hind Wings. Macropterus.</paragraph>
<paragraph pageId="32" pageNumber="33">Legs. Tarsal claws smooth. Males with adhesive vestiture ventrally, two rows of squamo-setae on tarsomeres 1-3 of fore-leg.</paragraph>
<paragraph pageId="32" pageNumber="33">Male genitalia. Ostium left pleuropic. Phallus cylindrical.</paragraph>
<paragraph pageId="32" pageNumber="33">Female genitalia. Gonocoxite 2 (gc2) wide at base, narrowing sharply toward apex, moderately curved. Sensory furrow, furrow pegs and associated nematiform setae not observed.</paragraph>
<paragraph id="AE186BB17C56340EF4DFE9316FC3B347" pageId="32" pageNumber="33">Color. Various.</paragraph>
<paragraph id="54C208C0DFA151F5BC3ABF5CE7CAE1B2" pageId="32" pageNumber="33">Fixed setae. Two pairs of supraorbital setae; clypeus with two lateral setae; labrum with six setae along apical margin; one pair of suborbital setae.; 16-17 lateral (umbilical) setae in interval 9; two setae on each of abdominal sterna III to VI, two setae along apical margin of sternum VII in males, four setae along apical margin of sternum VII in females.</paragraph>
<paragraph id="271B2D18A6633B978E3238D4B34B7C82" pageId="32" pageNumber="33">Head. Mandibles curved, left mandible slightly more curved and pointed at apex then right mandible, right mandible with additional, small tooth on inside cutting surface near mid-length of mandible; labrum bilobed; mentum with single broad tooth; eyes convex; palpi cylindrical, elongate.</paragraph>
<paragraph id="1B1D9B311718822AD3ACF2ECA711922A" pageId="32" pageNumber="33">Elytra. Humeri broadly rounded; elytral margin shallowly impressed in basal 1/3.</paragraph>
<paragraph id="563FECDB35C2487D2EE6C2319A0D671B" pageId="32" pageNumber="33">Hind Wings. Macropterus.</paragraph>
<paragraph id="C34E4FD228E2E4A8D1B5D1C6DFEBA460" pageId="32" pageNumber="33">Legs. Tarsal claws smooth. Males with adhesive vestiture ventrally, two rows of squamo-setae on tarsomeres 1-3 of fore-leg.</paragraph>
<paragraph id="0564DE2429EB95C6317A9306D79296AF" pageId="32" pageNumber="33">Male genitalia. Ostium left pleuropic. Phallus cylindrical.</paragraph>
<paragraph id="59030B4FA20A56B0C9A416664E7EA710" pageId="32" pageNumber="33">Female genitalia. Gonocoxite 2 (gc2) wide at base, narrowing sharply toward apex, moderately curved. Sensory furrow, furrow pegs and associated nematiform setae not observed.</paragraph>
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@ -1,96 +1,95 @@
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<taxonomicName id="56C0EA68627BF05F56C13232C4016612" class="Insecta" family="Carabidae" genus="Catascopoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopoides" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Catascopoides</taxonomicName>
Habu, 1967: 78;
<bibRefCitation pageId="44" pageNumber="45">Shpeley and Ball 2000</bibRefCitation>
<bibRefCitation id="0A70A77C68B8BB0DAD6483AE1C8F8A5C" pageId="44" pageNumber="45">Shpeley and Ball 2000</bibRefCitation>
: 112;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). Second ed. Germany" pageId="152" pageNumber="153" title="Systematic List of Extant Ground Beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="A5D14F63443369DAFD9B09A236C06FB2" author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). Second ed. Germany" pageId="152" pageNumber="153" title="Systematic List of Extant Ground Beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
: 454.
</paragraph>
<paragraph pageId="44" pageNumber="45">
<taxonomicName class="Insecta" family="Carabidae" genus="Dentiscopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dentiscopus" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Dentiscopus</taxonomicName>
<paragraph id="3AE8B8D8D4ED89E847C732501A115F55" pageId="44" pageNumber="45">
<taxonomicName id="285BAEE69744498B374C1F720A7CB162" class="Insecta" family="Carabidae" genus="Dentiscopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dentiscopus" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">Dentiscopus</taxonomicName>
Straneo, 1994: 148.
</paragraph>
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<subSubSection pageId="44" pageNumber="45" type="type species">
<paragraph pageId="44" pageNumber="45">Type species.</paragraph>
<paragraph pageId="44" pageNumber="45">
<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus mirabilis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="mirabilis">Catascopus mirabilis</taxonomicName>
<bibRefCitation pageId="44" pageNumber="45">Bates 1892</bibRefCitation>
<subSubSection id="ABB636D51B4BE4D404D29A2830809AB6" pageId="44" pageNumber="45" type="type species">
<paragraph id="8F40BF14B8E35FBC69607805F8DCA70F" pageId="44" pageNumber="45">Type species.</paragraph>
<paragraph id="8AC843AD59E94F390822E9F6C3787EB7" pageId="44" pageNumber="45">
<taxonomicName id="DC5E2B0FADE3B9A975809DB09005CCDE" class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus mirabilis" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="mirabilis">Catascopus mirabilis</taxonomicName>
<bibRefCitation id="B39E276520B143DE9B615D40F2D086FB" pageId="44" pageNumber="45">Bates 1892</bibRefCitation>
(by original designation).
</paragraph>
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<subSubSection pageId="44" pageNumber="45" type="type locality">
<paragraph pageId="44" pageNumber="45">Type locality.</paragraph>
<paragraph pageId="44" pageNumber="45">Southeast Asia (Myanmar and the Malay Peninsula).</paragraph>
<subSubSection id="2FE8DD91C2D004F46AE9F3BC2D06BAAD" pageId="44" pageNumber="45" type="type locality">
<paragraph id="44690653A2FCC4B49AAD3B369EC8CD43" pageId="44" pageNumber="45">Type locality.</paragraph>
<paragraph id="87181B2759A21E52B935A4F3B12676C3" pageId="44" pageNumber="45">Southeast Asia (Myanmar and the Malay Peninsula).</paragraph>
</subSubSection>
<subSubSection pageId="45" pageNumber="46" type="taxonomic notes">
<paragraph pageId="45" pageNumber="46">
<pageBreakToken pageId="45" pageNumber="46" start="start">Taxonomic</pageBreakToken>
<subSubSection id="977B706C30E8B634938379DA882AAF24" pageId="45" pageNumber="46" type="taxonomic notes">
<paragraph id="55ADEEAC0DE8F351228DDC7A579A3A94" pageId="45" pageNumber="46">
<pageBreakToken id="2282F3AE5141189D0256A761B7AC1244" pageId="45" pageNumber="46" start="start">Taxonomic</pageBreakToken>
notes.
</paragraph>
<paragraph pageId="45" pageNumber="46">
<paragraph id="B9D6D446501AC77223ACC0ABBA8DD2B1" pageId="45" pageNumber="46">
In a short footnote,
<bibRefCitation author="Habu, A" journalOrPublisher="Tokyo Electrical Engineering College Press, Tokyo" pageId="151" pageNumber="152" title="Carabidae. Truncatipennes group. (Insecta: Coleoptera). Fauna Japonica." year="1967">Habu (1967)</bibRefCitation>
<bibRefCitation id="77FCB7EB46803400A988BBDDC3D79877" author="Habu, A" journalOrPublisher="Tokyo Electrical Engineering College Press, Tokyo" pageId="151" pageNumber="152" title="Carabidae. Truncatipennes group. (Insecta: Coleoptera). Fauna Japonica." year="1967">Habu (1967)</bibRefCitation>
introduced the subgenus
<taxonomicName lsidName="(Catascopoides)" pageId="45" pageNumber="46" rank="subGenus" subGenus="Catascopoides">Catascopoides</taxonomicName>
<taxonomicName id="1B54C714E42D94C42319E30362E9F4A9" lsidName="(Catascopoides)" pageId="45" pageNumber="46" rank="subGenus" subGenus="Catascopoides">Catascopoides</taxonomicName>
. His rationale for this subgenus included both unique mandibular form (Fig. 23B) and distinctive pubescence on the ventral surface of the fore and mid femora. Later, and apparently unaware of
<normalizedToken originalValue="Habus">Habu's</normalizedToken>
<normalizedToken id="11BB1D1D90B2B0D168FEEA3AD436B570" originalValue="Habus">Habu's</normalizedToken>
short treatment,
<bibRefCitation author="Straneo, SL" journalOrPublisher="Elytron" pageId="154" pageNumber="155" pagination="141 - 172" title="Sulle species Orientali del genere Catascopus Kirby, 1825 (Coleoptera: Carabidae)." volume="8" year="1994">Straneo (1994)</bibRefCitation>
<bibRefCitation id="ABC17041E3F8DA17F7AD7F78F4FAE183" author="Straneo, SL" journalOrPublisher="Elytron" pageId="154" pageNumber="155" pagination="141 - 172" title="Sulle species Orientali del genere Catascopus Kirby, 1825 (Coleoptera: Carabidae)." volume="8" year="1994">Straneo (1994)</bibRefCitation>
, erected the now synonymized subgenus
<taxonomicName lsidName="(Dentiscopus)" pageId="45" pageNumber="46" rank="subGenus" subGenus="Dentiscopus">Dentiscopus</taxonomicName>
<taxonomicName id="ABFABC12A04BD0553D37C5338B13AED3" lsidName="(Dentiscopus)" pageId="45" pageNumber="46" rank="subGenus" subGenus="Dentiscopus">Dentiscopus</taxonomicName>
, that was based largely on the same characters. These characteristics are indeed unique within the
<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Catascopus</taxonomicName>
<taxonomicName id="5674DCD2A62278D66B276DE3366D65EE" class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">Catascopus</taxonomicName>
s. l. and the three currently known species appear to be quite closely related.
</paragraph>
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<treatment ID-GBIF-Taxon="154125948" LSID="urn:lsid:plazi:treatment:F56214254B59A9573E05F58F8BA3CB78" httpUri="http://treatment.plazi.org/id/F56214254B59A9573E05F58F8BA3CB78" lastPageNumber="52" pageId="51" pageNumber="52">
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<paragraph pageId="51" pageNumber="52">
<pageBreakToken pageId="51" pageNumber="52" start="start">Subgenus</pageBreakToken>
<taxonomicName authority="Jeannel" family="Carabidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="(Coptoderina)" order="Coleoptera" pageId="51" pageNumber="52" rank="subGenus" subGenus="Coptoderina">Coptoderina Jeannel</taxonomicName>
<treatment id="F56214254B59A9573E05F58F8BA3CB78" ID-GBIF-Taxon="154125948" LSID="urn:lsid:plazi:treatment:F56214254B59A9573E05F58F8BA3CB78" httpUri="http://treatment.plazi.org/id/F56214254B59A9573E05F58F8BA3CB78" lastPageNumber="52" pageId="51" pageNumber="52">
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<paragraph id="46C6F21996EE8B344B9F64FEE57BD27C" pageId="51" pageNumber="52">
<pageBreakToken id="3BBA8EC80FC788D9DF931E43BC62DCE0" pageId="51" pageNumber="52" start="start">Subgenus</pageBreakToken>
<taxonomicName id="B39110D017B69A988F73B1F9C470E896" ID-CoL="8MH5X" authority="Jeannel" class="Insecta" family="Carabidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="(Coptoderina)" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="subGenus" subGenus="Coptoderina">Coptoderina Jeannel</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="reference_group">
<paragraph pageId="51" pageNumber="52">
<taxonomicName class="Insecta" family="Carabidae" genus="Coptoderina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptoderina" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">Coptoderina</taxonomicName>
<subSubSection id="B7001A14D31143A760A0DCDF574DEBCB" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="406AAC44C3EE1B4AD1B2CB08D729936A" pageId="51" pageNumber="52">
<taxonomicName id="2DF25D8A9FE67D051FF73266EE88B8EB" class="Insecta" family="Carabidae" genus="Coptoderina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptoderina" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">Coptoderina</taxonomicName>
Jeannel, 1949: 1924;
<bibRefCitation author="Nakane, T" journalOrPublisher="The Entomological review of Japan" pageId="153" pageNumber="154" pagination="45 - 50" title="On the species of the genus Coptoderina in Japan and Formosa (Coleoptera, Caraboidea, Coptoderinae)." volume="7" year="1956">Nakane and Ohkura 1956</bibRefCitation>
<bibRefCitation id="83BB544DBAB7BB342E2210238EAEEDB1" author="Nakane, T" journalOrPublisher="The Entomological review of Japan" pageId="153" pageNumber="154" pagination="45 - 50" title="On the species of the genus Coptoderina in Japan and Formosa (Coleoptera, Caraboidea, Coptoderinae)." volume="7" year="1956">Nakane and Ohkura 1956</bibRefCitation>
: 45;
<bibRefCitation author="Habu, A" journalOrPublisher="Kontyu" pageId="151" pageNumber="152" pagination="114 - 117" title="A new species of Coptodera from Japan (Coleoptera, Carabidae)." volume="25" year="1957">Habu 1957</bibRefCitation>
<bibRefCitation id="6B5993FFA3301499065BC151E5DB9CDE" author="Habu, A" journalOrPublisher="Kontyu" pageId="151" pageNumber="152" pagination="114 - 117" title="A new species of Coptodera from Japan (Coleoptera, Carabidae)." volume="25" year="1957">Habu 1957</bibRefCitation>
: 114;
<bibRefCitation author="Shibata, T" journalOrPublisher="The Entomological Review of Japan" pageId="153" pageNumber="154" pagination="39 - 47" title="Notes on the Carabid-beetles of Japan. I. (Coleoptera)." volume="17" year="1964">Shibata 1964</bibRefCitation>
<bibRefCitation id="3DE580B0CB3D8E6B96ECFDD6D7335570" author="Shibata, T" journalOrPublisher="The Entomological Review of Japan" pageId="153" pageNumber="154" pagination="39 - 47" title="Notes on the Carabid-beetles of Japan. I. (Coleoptera)." volume="17" year="1964">Shibata 1964</bibRefCitation>
: 40;
<bibRefCitation author="Ball, GE" journalOrPublisher="Quaestiones Entomologicae" pageId="149" pageNumber="150" pagination="143 - 242" title="Pericaline Lebiini: notes on classification, a synopsis of the New World genera, and a revision of the genus Phloeoxena Chaudoir (Coleoptera: Carabidae)." volume="11" year="1975">Ball 1975</bibRefCitation>
<bibRefCitation id="D72A0CED71F1B1A2817E1D99FD9AA733" author="Ball, GE" journalOrPublisher="Quaestiones Entomologicae" pageId="149" pageNumber="150" pagination="143 - 242" title="Pericaline Lebiini: notes on classification, a synopsis of the New World genera, and a revision of the genus Phloeoxena Chaudoir (Coleoptera: Carabidae)." volume="11" year="1975">Ball 1975</bibRefCitation>
: 177;
<bibRefCitation author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). Second ed. Germany" pageId="152" pageNumber="153" title="Systematic List of Extant Ground Beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
<bibRefCitation id="E42560AFFA8751630703CD9AEA9DC2EE" author="Lorenz, W" journalOrPublisher="Paussinae, Cicindelinae, Rhysodinae). Second ed. Germany" pageId="152" pageNumber="153" title="Systematic List of Extant Ground Beetles of the World. (Insecta, Coleoptera, Adephaga: Trachypachidae &amp; Carabidae incl." year="2005">Lorenz 2005</bibRefCitation>
: 457;
<bibRefCitation author="Baehr, M" journalOrPublisher="Spixiana" pageId="149" pageNumber="150" pagination="33 - 35" title="A new subspecies of Coptoderapapuella Darlington from New Britain (Insecta, Coleoptera, Carabidae, Lebiinae)." volume="28" year="2005">Baehr 2005</bibRefCitation>
<bibRefCitation id="8E9384C68E5EB4BFF629A4A108819F91" author="Baehr, M" journalOrPublisher="Spixiana" pageId="149" pageNumber="150" pagination="33 - 35" title="A new subspecies of Coptoderapapuella Darlington from New Britain (Insecta, Coleoptera, Carabidae, Lebiinae)." volume="28" year="2005">Baehr 2005</bibRefCitation>
: 33;
<bibRefCitation author="Baehr, M" journalOrPublisher="Coleoptera" pageId="149" pageNumber="150" pagination="175 - 199" title="The Australian species of the genus Coptodera Dejean. (Coleoptera, Carabidae, Lebiinae)." volume="11" year="2007">2007</bibRefCitation>
<bibRefCitation id="5A98070B6F5401886C7A8535552AFCDC" author="Baehr, M" journalOrPublisher="Coleoptera" pageId="149" pageNumber="150" pagination="175 - 199" title="The Australian species of the genus Coptodera Dejean. (Coleoptera, Carabidae, Lebiinae)." volume="11" year="2007">2007</bibRefCitation>
: 175;
<bibRefCitation pageId="51" pageNumber="52">2014</bibRefCitation>
<bibRefCitation id="0792505CDB8931195B1FA5D4F6F6AC97" pageId="51" pageNumber="52">2014</bibRefCitation>
: 249;
<bibRefCitation author="Park, J K" journalOrPublisher="The Entomological review of Japan" pageId="153" pageNumber="154" url="https://doi.org/10.1111/j.1748-5967.2011.00324.x" year="2011">Park et al. 2011</bibRefCitation>
<bibRefCitation id="850BA551A380182E0046586DEC6ECB44" author="Park, J K" journalOrPublisher="The Entomological review of Japan" pageId="153" pageNumber="154" url="https://doi.org/10.1111/j.1748-5967.2011.00324.x" year="2011">Park et al. 2011</bibRefCitation>
: 103;
<bibRefCitation author="Gamboa, S" journalOrPublisher="Zootaxa" pageId="150" pageNumber="151" pagination="592 - 596" title="A new fossil species of the genus Coptodera Dejean, 1825 (Coleoptera: Carabidae: Lebiinae) from Baltic amber." url="https://doi.org/10.11646/zootaxa.3981.4.9" volume="3981" year="2015">Gamboa and Ortuno 2015</bibRefCitation>
<bibRefCitation id="4CD10ABFD3AC4AB9C62BEA2457E5D157" author="Gamboa, S" journalOrPublisher="Zootaxa" pageId="150" pageNumber="151" pagination="592 - 596" title="A new fossil species of the genus Coptodera Dejean, 1825 (Coleoptera: Carabidae: Lebiinae) from Baltic amber." url="https://doi.org/10.11646/zootaxa.3981.4.9" volume="3981" year="2015">Gamboa and Ortuno 2015</bibRefCitation>
: 593.
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="type species">
<paragraph pageId="51" pageNumber="52">Type species.</paragraph>
<paragraph pageId="51" pageNumber="52">
<taxonomicName class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus equestris" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="equestris">Catascopus equestris</taxonomicName>
<subSubSection id="B33686F6125CE4BF38C7345527F6A5FA" pageId="51" pageNumber="52" type="type species">
<paragraph id="FE6A78CA930B81370320B7DC1DE61C48" pageId="51" pageNumber="52">Type species.</paragraph>
<paragraph id="5605FD61C016E7EF25425BA472ED8415" pageId="51" pageNumber="52">
<taxonomicName id="DBAA9DB8FBD53B70D3D927749D5E3738" class="Insecta" family="Carabidae" genus="Catascopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catascopus equestris" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="equestris">Catascopus equestris</taxonomicName>
Boheman, 1848.
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="type locality">
<paragraph pageId="51" pageNumber="52">Type locality.</paragraph>
<paragraph pageId="51" pageNumber="52">South Africa.</paragraph>
<subSubSection id="A1BB2B64B06DB59A57E5646B5EA266AC" pageId="51" pageNumber="52" type="type locality">
<paragraph id="ABD22369B65467FB25519CDF870A93B0" pageId="51" pageNumber="52">Type locality.</paragraph>
<paragraph id="2FBF1E3B481FBAB1AEBCE45C146FEE8A" pageId="51" pageNumber="52">South Africa.</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="recognition of taiwanese species of coptoderina">
<paragraph pageId="51" pageNumber="52">
<subSubSection id="0F1F299A4AF6A395760E57D890398F0B" pageId="51" pageNumber="52" type="recognition of taiwanese species of coptoderina">
<paragraph id="66248B79C69C290270F39C4B760E0676" pageId="51" pageNumber="52">
Recognition of Taiwanese species of
<taxonomicName class="Insecta" family="Carabidae" genus="Coptoderina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptoderina" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">Coptoderina</taxonomicName>
<taxonomicName id="515EB1AE05A5D3DC1995FFA01C1594EF" class="Insecta" family="Carabidae" genus="Coptoderina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coptoderina" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">Coptoderina</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="">
<paragraph pageId="51" pageNumber="52">.</paragraph>
<paragraph pageId="51" pageNumber="52">Color. Various.</paragraph>
<paragraph pageId="51" pageNumber="52">Fixed setae. Two pairs of supraorbital setae; clypeus with two lateral setae; labrum with six setae along apical margin; one pair of suborbital setae; pronotum with two pairs of setae set in raised punctures, one at base of lateral margin and one on lateral margin at pronotum max width; 17 lateral (umbilical) setae in interval 9; ventrally, two setae on each of abdominal sterna III to VI, four setae along apical margin of sternum VII.</paragraph>
<paragraph pageId="51" pageNumber="52">Head. Mentum with no tooth; eyes large, convex; palpi cylindrical, elongate, and setose.</paragraph>
<paragraph pageId="51" pageNumber="52">Elytra. Humeri broadly rounded; lateral margin explanate, slightly more so in basal 1/3.</paragraph>
<paragraph pageId="51" pageNumber="52">Hind wings. Macropterous.</paragraph>
<paragraph pageId="51" pageNumber="52">Legs. Tarsal claws denticulate, three to four denticles per claw, males with adhesive vestiture ventrally, two rows of squamo-setae on tarsomeres 1-3 of fore-leg.</paragraph>
<paragraph pageId="51" pageNumber="52">Female genitalia. Gonocoxite 2 (gc2) long, narrow, slightly curved; three lateral ensiform setae (les) and one dorsal ensiform seta (des) present. Sensory furrow, furrow pegs and associated nematiform setae not observed.</paragraph>
<subSubSection id="9B33DE472B6AF6697B14D2D217000A0D" pageId="51" pageNumber="52" type="">
<paragraph id="ECC3059822EC7312A046C36E89ED6F43" pageId="51" pageNumber="52">.</paragraph>
<paragraph id="ABFE90D0AB898B4258D0842370F34319" pageId="51" pageNumber="52">Color. Various.</paragraph>
<paragraph id="6BF0FE3391FCD18CDFAD48FC3D6AE54A" pageId="51" pageNumber="52">Fixed setae. Two pairs of supraorbital setae; clypeus with two lateral setae; labrum with six setae along apical margin; one pair of suborbital setae; pronotum with two pairs of setae set in raised punctures, one at base of lateral margin and one on lateral margin at pronotum max width; 17 lateral (umbilical) setae in interval 9; ventrally, two setae on each of abdominal sterna III to VI, four setae along apical margin of sternum VII.</paragraph>
<paragraph id="22A3D510F25EE78EA327C62CAAA4BBBD" pageId="51" pageNumber="52">Head. Mentum with no tooth; eyes large, convex; palpi cylindrical, elongate, and setose.</paragraph>
<paragraph id="A2EA3FCFD01864160D0E6ADFD975C38B" pageId="51" pageNumber="52">Elytra. Humeri broadly rounded; lateral margin explanate, slightly more so in basal 1/3.</paragraph>
<paragraph id="4F447DBDD541C947049228459BA3AE89" pageId="51" pageNumber="52">Hind wings. Macropterous.</paragraph>
<paragraph id="96ECA5A0DA5A93C7DAC05DEBB02BB654" pageId="51" pageNumber="52">Legs. Tarsal claws denticulate, three to four denticles per claw, males with adhesive vestiture ventrally, two rows of squamo-setae on tarsomeres 1-3 of fore-leg.</paragraph>
<paragraph id="0E08720E176BABEC92A5C452F1808DE6" pageId="51" pageNumber="52">Female genitalia. Gonocoxite 2 (gc2) long, narrow, slightly curved; three lateral ensiform setae (les) and one dorsal ensiform seta (des) present. Sensory furrow, furrow pegs and associated nematiform setae not observed.</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="taxonomic notes">
<paragraph pageId="51" pageNumber="52">Taxonomic notes.</paragraph>
<paragraph pageId="51" pageNumber="52">
<subSubSection id="31FA10851F8DC1F2B6146EC9B6981AA6" pageId="51" pageNumber="52" type="taxonomic notes">
<paragraph id="2ADC2C4D07D25E3228123D94E7ECDE09" pageId="51" pageNumber="52">Taxonomic notes.</paragraph>
<paragraph id="1B1BAE165A8FAE5CA5DF5949895838A8" pageId="51" pageNumber="52">
All species from the Oriental Region belong to the subgenus
<taxonomicName lsidName="(Coptoderina)" pageId="51" pageNumber="52" rank="subGenus" subGenus="Coptoderina">Coptoderina</taxonomicName>
<taxonomicName id="CE35E966B81058B8E7AD9080B9DBEFE5" lsidName="(Coptoderina)" pageId="51" pageNumber="52" rank="subGenus" subGenus="Coptoderina">Coptoderina</taxonomicName>
Jeannel, 1949. They are distinguished by the median lobe of males with ostium catopic.
</paragraph>
</subSubSection>