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<document id="FF96165EB7C5220DB9E6C62F20C568BE" ID-DOI="10.1016/j.rbe.2017.04.004" ID-ISSN="1806-9665" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722635027235" checkinUser="felipe" docAuthor="Ferreira, André da Silva, Almeida, Lúcia M. &amp; Bravo, Freddy" docDate="2017" docId="6B7287A3FFE1FF98F6ACFE5DFD3AFB8B" docLanguage="en" docName="RevBrasEntomol.61.3.208-223.pdf" docOrigin="Revista Brasileira de Entomologia 61 (3)" docSource="http://dx.doi.org/10.1016/j.rbe.2017.04.004" docStyle="DocumentStyle:0303D37C566E4E2A42C6591146119864.3:RevBrasEntomol.2015-.journal_article" docStyleId="0303D37C566E4E2A42C6591146119864" docStyleName="RevBrasEntomol.2015-.journal_article" docStyleVersion="3" docTitle="Pelidnota beckeri Ferreira, Almeida &amp; Bravo, 2017, sp. nov." docType="treatment" docVersion="1" lastPageNumber="211" masterDocId="974BFFDBFFE0FF9BF63EFFF3FF99FFD4" masterDocTitle="Three new species of Pelidnota MacLeay (Coleoptera, Scarabaeidae, Rutelinae) and new distributional records from northeast Brazil" masterLastPageNumber="223" masterPageNumber="208" pageNumber="209" updateTime="1722679709455" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A645701C21827FFFF98BBC7107705EB3">Three new species of Pelidnota MacLeay (Coleoptera, Scarabaeidae, Rutelinae) and new distributional records from northeast Brazil</mods:title>
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<mods:namePart id="97B39345B80D6E851CBB5A5E4852E3A6">Ferreira, André da Silva</mods:namePart>
<mods:affiliation id="3D1E8C98F9AB5326B858DC9ADBBD3A20">Universidade Federal da Bahia, Instituto de Biologia, Programa de Pós-Graduação em Diversidade Animal, Bahia, BA, Brazil</mods:affiliation>
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<mods:namePart id="00F28E4D154C0063F914E0B140330772">Almeida, Lúcia M.</mods:namePart>
<mods:affiliation id="F72F1CB50C05F613363EB7E1FE4FC4E1">Universidade Federal do Paraná, Departamento de Zoologia, Laboratório de Sistemática e Bioecologia de Coleoptera, Curitiba, PR, Brazil</mods:affiliation>
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<mods:namePart id="AB22006B70DC5F2A95D29783EBC07ECB">Bravo, Freddy</mods:namePart>
<mods:affiliation id="6336195E2D961E5BA94371DE620EF783">Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Laboratório de Sistemática de Insetos, Feira de Santana, BA, Brazil</mods:affiliation>
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<subSubSection id="ABC1653EFFE1FF9AF6ACFE5DFE04FE16" box="[146,413,430,450]" pageId="1" pageNumber="209" type="nomenclature">
<paragraph id="E36436B5FFE1FF9AF6ACFE5DFE04FE16" blockId="1.[146,413,430,450]" box="[146,413,430,450]" pageId="1" pageNumber="209">
<heading id="B82C81D9FFE1FF9AF6ACFE5DFE04FE16" bold="true" box="[146,413,430,450]" fontSize="8" level="2" pageId="1" pageNumber="209" reason="2">
<emphasis id="D1AFEAA7FFE1FF9AF6ACFE5DFE04FE16" bold="true" box="[146,413,430,450]" pageId="1" pageNumber="209">
<taxonomicName id="24DB4D36FFE1FF9AF6ACFE5DFEDEFE16" ID-CoL="75YT6" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[146,327,430,450]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="209" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFE1FF9AF6ACFE5DFEDEFE16" bold="true" box="[146,327,430,450]" italics="true" pageId="1" pageNumber="209">Pelidnota beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE1FF9AF772FE5CFE04FE16" box="[332,413,431,450]" pageId="1" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="ABC1653EFFE1FF98F6ACFE38FD3AFB8B" lastPageId="3" lastPageNumber="211" pageId="1" pageNumber="209" type="description">
<paragraph id="E36436B5FFE1FF9AF6ACFE38FF72FE0A" blockId="1.[114,786,459,1985]" box="[146,235,459,478]" pageId="1" pageNumber="209">
<figureCitation id="7BE02A30FFE1FF9AF6ACFE38FF72FE0A" box="[146,235,459,478]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="1" pageNumber="209">Figs. 18</figureCitation>
</paragraph>
<paragraph id="E36436B5FFE1FF9AF6ACFE15FF53FD0D" blockId="1.[114,786,459,1985]" pageId="1" pageNumber="209">
<emphasis id="D1AFEAA7FFE1FF9AF6ACFE15FF69FE2E" box="[146,240,486,506]" italics="true" pageId="1" pageNumber="209">Diagnosis</emphasis>
. Body dark brown; legs brown, lacking reflections. Black lateral spot lacking on pronotum. Elytron with three lateral black spots, one at the humeral knob, one on the elytral half, and one at apex (
<figureCitation id="7BE02A30FFE1FF9AF6F6FDC9FE84FD99" box="[200,285,570,589]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="1" pageNumber="209">Figs. 3, 6</figureCitation>
). Second teeth of maxilla sawed. (
<figureCitation id="7BE02A30FFE1FF9AF44CFDC9FD2DFD99" box="[626,692,570,589]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="1" pageNumber="209">Fig. 25</figureCitation>
). Surface area of pre-mentum excavated and lateral margin of post-mentum rounded (
<figureCitation id="7BE02A30FFE1FF9AF6EAFD81FE8DFD51" box="[212,276,626,645]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="1" pageNumber="209">Fig.29</figureCitation>
). External surface of metatibia densely punctuated, apical margin with two spurs interleaved by two spine-like setae on inner margin and a small spine-like setae on the outer margin (
<figureCitation id="7BE02A30FFE1FF9AF644FD35FF24FD0D" box="[122,189,710,729]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="1" pageNumber="209">Fig. 53</figureCitation>
).
</paragraph>
<paragraph id="E36436B5FFE1FF9AF6ACFD12FC2FFCB0" blockId="1.[114,786,459,1985]" lastBlockId="1.[833,1504,151,1984]" pageId="1" pageNumber="209">
<emphasis id="D1AFEAA7FFE1FF9AF6ACFD12FE05FD21" box="[146,412,737,757]" italics="true" pageId="1" pageNumber="209">
<typeStatus id="3C608817FFE1FF9AF6ACFD12FF73FD21" box="[146,234,737,757]" pageId="1" pageNumber="209" type="holotype">Holotype</typeStatus>
male description
</emphasis>
(
<figureCitation id="7BE02A30FFE1FF9AF791FD11FD97FD21" box="[431,526,738,757]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="1" pageNumber="209">Figs. 13</figureCitation>
). Total length
<quantity id="24239B50FFE1FF9AF48FFD11FC94FD21" box="[689,781,738,757]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="1" pageNumber="209" unit="mm" value="25.9">25.9 mm</quantity>
; width
<quantity id="24239B50FFE1FF9AF68BFD0DFE96FCC5" box="[181,271,766,785]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="209" unit="mm" value="12.6">12.6 mm</quantity>
. Body elongated-oval, slightly convex.
<emphasis id="D1AFEAA7FFE1FF9AF4A5FD0EFD57FCC5" box="[667,718,765,785]" italics="true" pageId="1" pageNumber="209">Color</emphasis>
: Head and pronotum dark brown. Frons brown, slightly darker than pronotum. Scutellum and elytron light brown; elytron slightly lighter in color than head, pronotum, and scutellum. Elytron with three lateral black spots, one at the humeral knob, one on the elytral half, and one at apex. Pygidium dark brown, slightly darker than the rest of the body, with greenish reflections when illuminated with fluorescent light. Legs and ventral surface dark brown, similar in color to the head and pronotum.
<emphasis id="D1AFEAA7FFE1FF9AF47EFC2FFDEBFC24" box="[576,626,988,1008]" italics="true" pageId="1" pageNumber="209">Head</emphasis>
: Antenna with 10 antennomeres, club composed of three antennomeres, scape elongated, longer than antennomeres 27. Clypeus trapezoidal, anterior margin dark and concave (
<figureCitation id="7BE02A30FFE1FF9AF7D6FBC3FDB8FB90" box="[488,545,1072,1092]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="1" pageNumber="209">Fig. 7</figureCitation>
), punctures dense and moderately large. Frontoclypeal suture not evident. Frons width at middle region 23 times larger than clypeus length; punctures dense and moderately large. A set of bristling setae surrounding the eye. Canthus half as long as eye. Interocular distance grather than 3 times the transverse eye diameters. Labrum transversely elongated and not fused to clypeus, with medial compression and small apical tooth, with bristling setae across on all surface (
<figureCitation id="7BE02A30FFE1FF9AF644FAE3FF24FAF7" box="[122,189,1296,1315]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="1" pageNumber="209">Fig. 17</figureCitation>
). Mandibles visible beyond anterior margin of the clypeus, outer margin black and with 2 teeth; apical tooth slightly larger than internal one; molar robust (
<figureCitation id="7BE02A30FFE1FF9AF7EBFAB4FD82FA8F" box="[469,539,1351,1371]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="1" pageNumber="209">Fig. 21</figureCitation>
). Maxilla, galea with 6 teeth, second sawed; palp with 4 palpomeres, segment IV fusiform (
<figureCitation id="7BE02A30FFE1FF9AF644FA8CFF24FA46" box="[122,189,1407,1426]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="1" pageNumber="209">Fig. 25</figureCitation>
). Labium, apex of pre-mentum concave, palp inserted into a lateral and with 3 palpomeres, III elongated; post-mentum with elongated setae dorsally and lateral margin rounded, with elongated moderate setae (
<figureCitation id="7BE02A30FFE1FF9AF75AFA20FE33FA32" box="[356,426,1491,1510]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="1" pageNumber="209">Fig. 29</figureCitation>
).
<emphasis id="D1AFEAA7FFE1FF9AF7FEFA21FDB8FA32" box="[448,545,1490,1510]" italics="true" pageId="1" pageNumber="209">Pronotum</emphasis>
: Convex.
<quantity id="24239B50FFE1FF9AF4B5FA20FD4FFA32" box="[651,726,1491,1510]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="1" pageNumber="209" unit="mm" value="5.1">5.1 mm</quantity>
long, anterior width
<quantity id="24239B50FFE1FF9AF72DFA1CFEC6F9D6" box="[275,351,1519,1538]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="1" pageNumber="209" unit="mm" value="7.6">7.6 mm</quantity>
, median width
<quantity id="24239B50FFE1FF9AF436FA1CFDF8F9D6" box="[520,609,1519,1538]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="1" pageNumber="209" unit="mm" value="11.8">11.8 mm</quantity>
, posterior width
<quantity id="24239B50FFE1FF9AF64CF9F8FF55F9CA" box="[114,204,1547,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="209" unit="mm" value="11.7">11.7 mm</quantity>
. Punctures dense and moderately wide. Anterior margin concave, with anterior angles acute. Lateral margin rounded, with median portion convex. Posterior margins slightly convex. Membranous border surrounding of pronotum.
<emphasis id="D1AFEAA7FFE1FF9AF45FF9AEFD58F9A5" box="[609,705,1629,1649]" italics="true" pageId="1" pageNumber="209">Scutellum</emphasis>
: Width
<quantity id="24239B50FFE1FF9AF64CF989FF27F95A" box="[114,190,1658,1678]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="209" unit="mm" value="2.2">2.2 mm</quantity>
, total length
<quantity id="24239B50FFE1FF9AF768F989FE3AF95A" box="[342,419,1658,1678]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="209" unit="mm" value="1.3">1.3 mm</quantity>
. Punctures dense and moderately large, posterior margin rounded; dark brown, with greenish reflections when exposed to fluorescent light; lacking setae.
<emphasis id="D1AFEAA7FFE1FF9AF4FCF942FC90F911" box="[706,777,1713,1733]" italics="true" pageId="1" pageNumber="209">Elytron</emphasis>
: humeral width
<quantity id="24239B50FFE1FF9AF72DF93DFEF6F935" box="[275,367,1742,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="209" unit="mm" value="12.6">12.6 mm</quantity>
; total length
<quantity id="24239B50FFE1FF9AF7C7F93DFDCAF935" box="[505,595,1742,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="1" pageNumber="209" unit="mm" value="17.8">17.8 mm</quantity>
. Punctures sparse, small to moderate, slightly organized longitudinally, lacking setae. Membranous border present at anterior half. Slightly compressed laterally between humerus and medial portion of elytron. Apex with punctures dense, moderate to large.Elytral suture dark brown. Elytral apex truncated (
<figureCitation id="7BE02A30FFE1FF9AF751F8AAFE2FF8B9" box="[367,438,1881,1901]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="1" pageNumber="209">Fig. 33</figureCitation>
).
<emphasis id="D1AFEAA7FFE1FF9AF7F3F8ABFD94F8B8" box="[461,525,1880,1900]" italics="true" pageId="1" pageNumber="209">Venter</emphasis>
: Punctures dense, moderate to large. Densely covered by brownish setae. Prosternum elongated in the shape of a tubercle but not extending beyond procoxa. Mesoventrite with anterior process, extending beyond mesocoxae.
<emphasis id="D1AFEAA7FFE1FF9AF582FF64FC7CFF7F" box="[956,997,151,171]" italics="true" pageId="1" pageNumber="209">Legs</emphasis>
: Punctures moderate to dense. Tibia with 3 external teeth, apical tooth slightly recurved, larger than the other two; inner apical spur present; dorsal surface with a row of 10 spine-like setae (
<figureCitation id="7BE02A30FFE1FF9AF5C7FF1FFBDBFF2B" box="[1017,1090,236,255]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="1" pageNumber="209">Fig. 41</figureCitation>
); a tooth present on the larger tarsal claw (
<figureCitation id="7BE02A30FFE1FF9AF5BDFEF4FC53FECF" box="[899,970,263,283]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 4752. Anterior tarsal claws of male and left metatibia of female in lateral view.4750, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.4750 = 0.500 mm, Figs.51 and 52 = 1.0mm." pageId="1" pageNumber="209">Fig. 47</figureCitation>
) Metacoxa with posterior projection beyond base of metatrochanter. Metafemur flattened medially. Mesotibia with setae moderate on the inner surface and with sparse spine-like setae on the outer surface; apical margin dark, inner margin with two spurs interleaved with two spine-like setae, and outer margin with 9 spine-like setae. Metatibia slightly enlarged in median region, with slight compression at apex of outer surface; apical margin dark, with two spurs interleaved by two spine-like setae on inner margin and a small spine-like setae on the outer margin; outer surface densely punctuated (
<figureCitation id="7BE02A30FFE1FF9AF33DFDF1FAD1FDC1" box="[1283,1352,514,533]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="1" pageNumber="209">Fig. 53</figureCitation>
). Tarsomere V elongated on all legs, with a ventral tooth. Tarsomeres IIIV with two inner spine-like setae on all legs. Tarsal claws simple on all legs, inner protarsal claw slightly recurved and visibly larger than outer claw. Outer claws of the meso- and metatarsus larger than inner claw.
<emphasis id="D1AFEAA7FFE1FF9AF58BFD7EFB88FD75" box="[949,1041,653,673]" italics="true" pageId="1" pageNumber="209">Abdomen</emphasis>
: Abdominal ventrites convex, with punctures moderate to dense. Setae sparse to moderate.
<emphasis id="D1AFEAA7FFE1FF9AF318FD5AFA1BFD69" box="[1318,1410,681,701]" italics="true" pageId="1" pageNumber="209">Pygidium</emphasis>
: Slightly convex in lateral view. Surface with puncture striae, punctures dense and moderately large. Setae moderate to dense, brownish. Bristling setae present on posterior margin in ventral view.
<emphasis id="D1AFEAA7FFE1FF9AF57FFCEBFC3EFCF8" box="[833,935,792,812]" italics="true" pageId="1" pageNumber="209">Parameres</emphasis>
: (
<figureCitation id="7BE02A30FFE1FF9AF5FFFCEAFBFDFCF8" box="[961,1124,793,812]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="1" pageNumber="209">Figs. 57, 58, 65</figureCitation>
). Symmetrical, fused. Distal margin concave and gradually acute laterally; proximal margin slightly corrugated.
</paragraph>
<paragraph id="E36436B5FFE1FF9AF55EFC9FFA10FBD8" blockId="1.[833,1504,151,1984]" pageId="1" pageNumber="209">
<emphasis id="D1AFEAA7FFE1FF9AF55EFC9FFB47FC54" box="[864,1246,876,896]" italics="true" pageId="1" pageNumber="209">Dimensions in mm (male/female, n</emphasis>
=
<emphasis id="D1AFEAA7FFE1FF9AF2CFFC9FFC00FC48" italics="true" pageId="1" pageNumber="209">
20,
<specimenCount id="F5DDFD3CFFE1FF9AF321FC9FFA18FC54" box="[1311,1409,876,896]" count="10" pageId="1" pageNumber="209" type="male">10 males</specimenCount>
and
<specimenCount id="F5DDFD3CFFE1FF9AF3F9FC9FFC17FC48" count="10" pageId="1" pageNumber="209" type="female">10 females</specimenCount>
).
</emphasis>
Total length 22.926.3/24.828.5; pronotum length 5.16.3/5.67.2; anterior width of pronotum 6.98.7/7.28.2; median width of pronotum 9.811.6/10.812.8; posterior width of pronotum 9.811.6/10.612.7; humerus width 11.012.8/11.513.9; elytron length 15.517.8/15.218.3.
</paragraph>
<paragraph id="E36436B5FFE1FF9AF55EFBE0FC31FA46" blockId="1.[833,1504,151,1984]" pageId="1" pageNumber="209">
<emphasis id="D1AFEAA7FFE1FF9AF55EFBE0FB81FBF3" box="[864,1048,1043,1063]" italics="true" pageId="1" pageNumber="209">
Female (
<typeStatus id="3C608817FFE1FF9AF586FBE0FB89FBF3" box="[952,1040,1043,1063]" pageId="1" pageNumber="209" type="paratype">Paratype</typeStatus>
)
</emphasis>
(
<figureCitation id="7BE02A30FFE1FF9AF212FBE7FB13FBF3" box="[1068,1162,1044,1063]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="1" pageNumber="209">Figs. 46</figureCitation>
). Similar to male, but differing with respect to some characters. Clypeus trapezoidal to triangular, border black, medial section of anterior border straight (
<figureCitation id="7BE02A30FFE1FF9AF3A3FBBFFA4BFB8B" box="[1437,1490,1100,1119]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="1" pageNumber="209">Fig. 8</figureCitation>
). Mandible more developed than in males; pronotum with lateral margin slightly rounded to slightly convex. Pygidium convex, with anterior margin concave. Venter with dense setae, sometimes denser than in males. Tarsal claws simple, first inner tarsal claw larger than outer, and weakly recurved. Outer tarsal claw of meso- and metatarsus slightly larger than inner claw. Spines-like setae on outer surface, and inner setae of inner surface of metatibia moderately dense, sometimes denser than males. The posterior corner of the metacoxa extends beyond of the base of trochanter. Last abdominal ventrite triangular, extending to posterior margin of pygidium.
</paragraph>
<paragraph id="E36436B5FFE1FF9AF55EFA69FC04F9EE" blockId="1.[833,1504,151,1984]" pageId="1" pageNumber="209">
<emphasis id="D1AFEAA7FFE1FF9AF55EFA69FC5FFA7A" box="[864,966,1434,1454]" italics="true" pageId="1" pageNumber="209">
<typeStatus id="3C608817FFE1FF9AF55EFA69FC5BFA7A" box="[864,962,1434,1454]" pageId="1" pageNumber="209" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Similar to
<typeStatus id="3C608817FFE1FF9AF20CFA68FB17FA7A" box="[1074,1166,1435,1454]" pageId="1" pageNumber="209" type="holotype">holotype</typeStatus>
. Dark spots on side of elytron can be faint. Smaller spines-like setae on apical margin of last tibia ranging from 8 to 10. Apical spur on outer edge of last tibia present or not. Spines-like setae on outer surfaces and setae on inner surfaces of meso- and metatibia moderate to dense. Ventral setae moderate to dense.
</paragraph>
<paragraph id="E36436B5FFE1FF9AF55EF9B2FC08F814" blockId="1.[833,1504,151,1984]" pageId="1" pageNumber="209">
<emphasis id="D1AFEAA7FFE1FF9AF55EF9B2FC68F981" box="[864,1009,1601,1621]" italics="true" pageId="1" pageNumber="209">Type material.</emphasis>
<materialsCitation id="53B33CE8FFE1FF9AF5C4F9B1FB8AF959" collectingDate="1984-11" collectionCode="MZFS" collectorName="G. V. dos Santos" country="Brazil" location="Bahia" pageId="1" pageNumber="209" specimenCode="MZFS#55573" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<typeStatus id="3C608817FFE1FF9AF5C4F9B1FBCEF981" box="[1018,1111,1602,1621]" pageId="1" pageNumber="209" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="9BCC7625FFE1FF9AF29DF9B0FB74F982" box="[1187,1261,1603,1622]" name="Brazil" pageId="1" pageNumber="209">BRAZIL</collectingCountry>
,
<emphasis id="D1AFEAA7FFE1FF9AF2C4F9B2FAA8F981" box="[1274,1329,1601,1621]" italics="true" pageId="1" pageNumber="209">
<collectingRegion id="211FF857FFE1FF9AF2C4F9B2FAA8F981" box="[1274,1329,1601,1621]" country="Brazil" name="Bahia" pageId="1" pageNumber="209">Bahia</collectingRegion>
</emphasis>
: (“Una, Fazenda Reunidas Unacau”);
<date id="97651075FFE1FF9AF22BF9ADFBFDF9A5" box="[1045,1124,1630,1649]" pageId="1" pageNumber="209" value="1984-11">
<collectingDate id="8721E99DFFE1FF9AF22BF9ADFBFDF9A5" box="[1045,1124,1630,1649]" pageId="1" pageNumber="209" value="1984-11">XI.1984</collectingDate>
</date>
,
<collectorName id="4E2E5363FFE1FF9AF24FF9ADFA8AF9A5" box="[1137,1299,1630,1649]" pageId="1" pageNumber="209">G.V. dos Santos</collectorName>
col. (
<specimenCode id="B37D9ECEFFE1FF9AF370F9ADFA41F9A5" box="[1358,1496,1630,1649]" collectionCode="MZFS" pageId="1" pageNumber="209">MZFS#55573</specimenCode>
) (
<collectionCode id="85CAAE70FFE1FF9AF579F989FC1AF959" box="[839,899,1658,1677]" pageId="1" pageNumber="209">MZFS</collectionCode>
) (
<figureCitation id="7BE02A30FFE1FF9AF5A9F989FB93F959" box="[919,1034,1658,1677]" captionStart="Figs" captionStartId="13.[114,149,1725,1739]" captionTargetBox="[179,720,149,1695]" captionTargetId="figure-13@13.[179,720,149,1695]" captionTargetPageId="13" captionText="Figs.6977. Labels attachedtothe hotypes.6971,Pelidnota beckerisp.nov.;7274, P. nordestina sp. nov.; 7577, P.pernambucana sp. nov." pageId="1" pageNumber="209">Figs. 6971</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="53B33CE8FFE1FF9AF223F98AFBB9F911" collectingDate="2001-05-10" collectionCode="MZFS" collectorName="E. Neto" country="Brazil" elevation="430" location="Santa Terezinha" municipality="Pedra Branca" pageId="1" pageNumber="209" specimenCode="MZFS#2308" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
<emphasis id="D1AFEAA7FFE1FF9AF223F98AFB1AF959" box="[1053,1155,1657,1677]" italics="true" pageId="1" pageNumber="209">
<typeStatus id="3C608817FFE1FF9AF223F98AFBE6F959" box="[1053,1151,1657,1677]" pageId="1" pageNumber="209" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="F5DDFD3CFFE1FF9AF2B6F989FB79F959" box="[1160,1248,1658,1677]" count="1" pageId="1" pageNumber="209" type="female">1 female</specimenCount>
(“
<location id="E604606EFFE1FF9AF2C8F989FA05F959" LSID="urn:lsid:plazi:treatment:6B7287A3FFE1FF98F6ACFE5DFD3AFB8B:E604606EFFE1FF9AF2C8F989FA05F959" box="[1270,1436,1658,1677]" country="Brazil" municipality="Pedra Branca" name="Santa Terezinha" pageId="1" pageNumber="209" stateProvince="Bahia">Santa Terezinha</location>
,
<collectingMunicipality id="0300ACCFFFE1FF9AF398F989FC12F97D" pageId="1" pageNumber="209">Pedra Branca</collectingMunicipality>
”; “
<quantity id="24239B50FFE1FF9AF591F965FC76F97D" box="[943,1007,1686,1705]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="209" unit="m" value="430.0">
<elevation id="68F6D186FFE1FF9AF591F965FC76F97D" box="[943,1007,1686,1705]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="209" unit="m" value="430.0">430 m</elevation>
</quantity>
”) (12
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50
<emphasis id="D1AFEAA7FFE1FF9AF26DF961FBCEF974" bold="true" box="[1107,1111,1682,1696]" pageId="1" pageNumber="209">
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</emphasis>
S/39
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</emphasis>
30
<emphasis id="D1AFEAA7FFE1FF9AF297F961FB34F974" bold="true" box="[1193,1197,1682,1696]" pageId="1" pageNumber="209">
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</emphasis>
W),
<date id="97651075FFE1FF9AF2DCF965FAD0F97D" box="[1250,1353,1686,1705]" pageId="1" pageNumber="209" value="2001-05-10">
<collectingDate id="8721E99DFFE1FF9AF2DCF965FAD0F97D" box="[1250,1353,1686,1705]" pageId="1" pageNumber="209" value="2001-05-10">10.V.2001</collectingDate>
</date>
.
<collectorName id="4E2E5363FFE1FF9AF362F964FA37F97E" box="[1372,1454,1687,1706]" pageId="1" pageNumber="209">E. Neto</collectorName>
col. (
<specimenCode id="B37D9ECEFFE1FF9AF579F941FC5CF911" box="[839,965,1714,1733]" collectionCode="MZFS" pageId="1" pageNumber="209">MZFS#2308</specimenCode>
) (
<collectionCode id="85CAAE70FFE1FF9AF5E5F941FB82F911" box="[987,1051,1714,1733]" pageId="1" pageNumber="209">MZFS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="53B33CE8FFE1FF9AF213F941FC47F929" collectingDate="2007-11-17" collectionCode="MZFS, DZUP" collectorName="Joao &amp; Zafira &amp; Daniela" country="Brazil" location="Itubera" municipality="Michelin" pageId="1" pageNumber="209" specimenCode="MZFS#33533" specimenCount="5" specimenCount-male="5" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="F5DDFD3CFFE1FF9AF213F941FBE6F911" box="[1069,1151,1714,1733]" count="5" pageId="1" pageNumber="209" type="male">5 males</specimenCount>
(“
<location id="E604606EFFE1FF9AF2A6F941FB7DF911" LSID="urn:lsid:plazi:treatment:6B7287A3FFE1FF98F6ACFE5DFD3AFB8B:E604606EFFE1FF9AF2A6F941FB7DF911" box="[1176,1252,1714,1733]" country="Brazil" municipality="Michelin" name="Itubera" pageId="1" pageNumber="209" stateProvince="Bahia">Ituberá</location>
,
<collectingMunicipality id="0300ACCFFFE1FF9AF2CCF941FAC8F911" box="[1266,1361,1714,1733]" pageId="1" pageNumber="209">Michelin</collectingMunicipality>
”);
<date id="97651075FFE1FF9AF350F941FA45F911" box="[1390,1500,1714,1733]" pageId="1" pageNumber="209" value="2007-11-17">
<collectingDate id="8721E99DFFE1FF9AF350F941FA45F911" box="[1390,1500,1714,1733]" pageId="1" pageNumber="209" value="2007-11-17">17.XI.2007</collectingDate>
</date>
,
<collectorName id="4E2E5363FFE1FF9AF57FF93DFCF7F935" box="[833,878,1742,1761]" pageId="1" pageNumber="209">João</collectorName>
,
<collectorName id="4E2E5363FFE1FF9AF544F93DFC2DF935" box="[890,948,1742,1761]" pageId="1" pageNumber="209">Zafira</collectorName>
&amp;
<collectorName id="4E2E5363FFE1FF9AF5EBF93DFBBBF935" box="[981,1058,1742,1761]" pageId="1" pageNumber="209">Daniela</collectorName>
col. (
<specimenCode id="B37D9ECEFFE1FF9AF263F93DFB73F935" box="[1117,1258,1742,1761]" collectionCode="MZFS" pageId="1" pageNumber="209">MZFS#33533</specimenCode>
; 33862 (
<collectionCode id="85CAAE70FFE1FF9AF37BF93DFA1CF935" box="[1349,1413,1742,1761]" pageId="1" pageNumber="209">MZFS</collectionCode>
); 33863; 33864 (
<collectionCode id="85CAAE70FFE1FF9AF5AFF919FC49F929" box="[913,976,1770,1789]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="1" pageNumber="209" type="University or college">DZUP</collectionCode>
))
</materialsCitation>
.
<materialsCitation id="53B33CE8FFE1FF9AF5D2F919FBA8F8E1" collectingDate="2007-12-07" collectionCode="MZFS" country="Brazil" elevation="493" location="Sao Felix do Coribe" municipality="Coribe - Km" pageId="1" pageNumber="209" specimenCode="MZFS#41178" specimenCount="1" specimenCount-male="1" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="F5DDFD3CFFE1FF9AF5D2F919FBACF929" box="[1004,1077,1770,1789]" count="1" pageId="1" pageNumber="209" type="male">1 male</specimenCount>
(“
<location id="E604606EFFE1FF9AF271F919FABBF929" LSID="urn:lsid:plazi:treatment:6B7287A3FFE1FF98F6ACFE5DFD3AFB8B:E604606EFFE1FF9AF271F919FABBF929" box="[1103,1314,1770,1789]" country="Brazil" municipality="Coribe - Km" name="Sao Felix do Coribe" pageId="1" pageNumber="209" stateProvince="Bahia">São Félix do Coribe</location>
,
<collectingMunicipality id="0300ACCFFFE1FF9AF30EF919FA2FF929" box="[1328,1462,1770,1789]" pageId="1" pageNumber="209">Coribe Km</collectingMunicipality>
24, “
<quantity id="24239B50FFE1FF9AF574F8F5FC10F8CD" box="[842,905,1798,1817]" metricMagnitude="2" metricUnit="m" metricValue="4.93" pageId="1" pageNumber="209" unit="m" value="493.0">
<elevation id="68F6D186FFE1FF9AF574F8F5FC10F8CD" box="[842,905,1798,1817]" metricMagnitude="2" metricUnit="m" metricValue="4.93" pageId="1" pageNumber="209" unit="m" value="493.0">493 m</elevation>
</quantity>
”) (13
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33
<emphasis id="D1AFEAA7FFE1FF9AF5D4F8F1FC77F8C4" bold="true" box="[1002,1006,1794,1808]" pageId="1" pageNumber="209">
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</emphasis>
S/44
<emphasis id="D1AFEAA7FFE1FF9AF222F8F1FBBDF8C4" bold="true" box="[1052,1060,1794,1808]" pageId="1" pageNumber="209">
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15
<emphasis id="D1AFEAA7FFE1FF9AF201F8F1FBDAF8C4" bold="true" box="[1087,1091,1794,1808]" pageId="1" pageNumber="209">
<superScript id="14AE9BFDFFE1FF9AF201F8F1FBDAF8C4" attach="left" box="[1087,1091,1794,1808]" fontSize="6" pageId="1" pageNumber="209">l</superScript>
</emphasis>
W),
<date id="97651075FFE1FF9AF24AF8F5FB70F8CD" box="[1140,1257,1798,1817]" pageId="1" pageNumber="209" value="2007-12-07">
<collectingDate id="8721E99DFFE1FF9AF24AF8F5FB70F8CD" box="[1140,1257,1798,1817]" pageId="1" pageNumber="209" value="2007-12-07">07.XII.2007</collectingDate>
</date>
, FB, TZ, NA &amp; JRA col. (
<specimenCode id="B37D9ECEFFE1FF9AF579F8D1FC4BF8E1" box="[839,978,1826,1845]" collectionCode="MZFS" pageId="1" pageNumber="209">MZFS#41178</specimenCode>
) (
<collectionCode id="85CAAE70FFE1FF9AF5D3F8D1FBB2F8E1" box="[1005,1067,1826,1845]" pageId="1" pageNumber="209">MZFS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="53B33CE8FFE1FF9AF27CF8D1FB4AF8B9" collectingDate="1985-10-28" collectionCode="MZFS" collectorName="J. Becker" country="Brazil" location="Itamaraju" pageId="1" pageNumber="209" specimenCode="MZFS#55574" specimenCount="8" specimenCount-female="2" specimenCount-male="6" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="F5DDFD3CFFE1FF9AF27CF8D1FB00F8E1" box="[1090,1177,1826,1845]" count="6" pageId="1" pageNumber="209" type="male">6 males</specimenCount>
and
<specimenCount id="F5DDFD3CFFE1FF9AF2E2F8D1FADFF8E1" box="[1244,1350,1826,1845]" count="2" pageId="1" pageNumber="209" type="female">2 females</specimenCount>
(“
<location id="E604606EFFE1FF9AF35AF8D1FA55F8E1" LSID="urn:lsid:plazi:treatment:6B7287A3FFE1FF98F6ACFE5DFD3AFB8B:E604606EFFE1FF9AF35AF8D1FA55F8E1" box="[1380,1484,1826,1845]" country="Brazil" name="Itamaraju" pageId="1" pageNumber="209" stateProvince="Bahia">Itamaraju</location>
”),
<date id="97651075FFE1FF9AF57FF8CEFC3EF884" box="[833,935,1853,1872]" pageId="1" pageNumber="209" value="1985-10-28">
<collectingDate id="8721E99DFFE1FF9AF57FF8CEFC3EF884" box="[833,935,1853,1872]" pageId="1" pageNumber="209" value="1985-10-28">28.X.1985</collectingDate>
</date>
,
<collectorName id="4E2E5363FFE1FF9AF589F8CDFB8DF884" box="[951,1044,1853,1873]" pageId="1" pageNumber="209">J. Becker</collectorName>
col. (
<specimenCode id="B37D9ECEFFE1FF9AF26BF8CEFB78F884" box="[1109,1249,1853,1872]" collectionCode="MZFS" pageId="1" pageNumber="209">MZFS#55574</specimenCode>
; 55575; 55576; 55577; 55578; 55579; 55580; 55581) (
<collectionCode id="85CAAE70FFE1FF9AF2B0F8A9FB55F8B9" box="[1166,1228,1882,1901]" pageId="1" pageNumber="209">MZFS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="53B33CE8FFE1FF9AF2DEF8A9FAE8F85C" collectingDate="1985-08" collectionCode="MNRJ" collectorName="Roppa &amp; J. Becker" country="Brazil" location="Itamaraju" pageId="1" pageNumber="209" specimenCount="14" specimenCount-female="11" specimenCount-male="3" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="F5DDFD3CFFE1FF9AF2DEF8A9FAAAF8B8" box="[1248,1331,1881,1901]" count="3" pageId="1" pageNumber="209" type="male">3 males</specimenCount>
and
<specimenCount id="F5DDFD3CFFE1FF9AF353F8A9FA46F8B8" box="[1389,1503,1881,1901]" count="11" pageId="1" pageNumber="209" type="female">11 females</specimenCount>
(“
<location id="E604606EFFE1FF9AF56EF886FC21F85C" LSID="urn:lsid:plazi:treatment:6B7287A3FFE1FF98F6ACFE5DFD3AFB8B:E604606EFFE1FF9AF56EF886FC21F85C" box="[848,952,1909,1928]" country="Brazil" name="Itamaraju" pageId="1" pageNumber="209" stateProvince="Bahia">Itamaraju</location>
”),
<date id="97651075FFE1FF9AF5EAF886FBA8F85C" box="[980,1073,1909,1928]" pageId="1" pageNumber="209" value="1985-08">
<collectingDate id="8721E99DFFE1FF9AF5EAF886FBA8F85C" box="[980,1073,1909,1928]" pageId="1" pageNumber="209" value="1985-08">VIII.1985</collectingDate>
</date>
,
<collectorName id="4E2E5363FFE1FF9AF202F885FBE5F85D" box="[1084,1148,1910,1929]" pageId="1" pageNumber="209">Roppa</collectorName>
&amp;
<collectorName id="4E2E5363FFE1FF9AF2A5F885FB6DF85C" box="[1179,1268,1909,1929]" pageId="1" pageNumber="209">J. Becker</collectorName>
col. (
<collectionCode id="85CAAE70FFE1FF9AF315F886FAF2F85C" box="[1323,1387,1909,1928]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="1" pageNumber="209">MNRJ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="53B33CE8FFE1FF9AF342F886FC14F814" collectingDate="2012-12-28" collectingDateMax="2012-12-29" collectingDateMin="2012-12-28" collectionCode="MNRJ" collectorName="M. L. Monne" country="Brazil" location="Camacan" municipality="Serra Bonita" pageId="1" pageNumber="209" specimenCount="3" specimenCount-female="3" stateProvince="Bahia" typeStatus="paratype">
<specimenCount id="F5DDFD3CFFE1FF9AF342F886FA46F85C" box="[1404,1503,1909,1928]" count="3" pageId="1" pageNumber="209" type="female">3 females</specimenCount>
(“
<location id="E604606EFFE1FF9AF56EF862FC29F870" LSID="urn:lsid:plazi:treatment:6B7287A3FFE1FF98F6ACFE5DFD3AFB8B:E604606EFFE1FF9AF56EF862FC29F870" box="[848,944,1937,1956]" country="Brazil" municipality="Serra Bonita" name="Camacan" pageId="1" pageNumber="209" stateProvince="Bahia">Camacan</location>
, RPPN
<collectingMunicipality id="0300ACCFFFE1FF9AF5C9F862FBE1F870" box="[1015,1144,1937,1956]" pageId="1" pageNumber="209">Serra Bonita</collectingMunicipality>
”),
<date id="97651075FFE1FF9AF2ACF862FAB4F870" box="[1170,1325,1937,1956]" pageId="1" pageNumber="209" value="2012-12-28" valueMax="2012-12-29" valueMin="2012-12-28">
<collectingDate id="8721E99DFFE1FF9AF2ACF862FAB4F870" box="[1170,1325,1937,1956]" pageId="1" pageNumber="209" value="2012-12-28" valueMax="2012-12-29" valueMin="2012-12-28">2829.XII.2012</collectingDate>
</date>
,
<collectorName id="4E2E5363FFE1FF9AF309F861FA2FF870" box="[1335,1462,1937,1957]" pageId="1" pageNumber="209">M. L. Monné</collectorName>
col. (
<collectionCode id="85CAAE70FFE1FF9AF579F85EFC1EF814" box="[839,903,1965,1984]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="1" pageNumber="209">MNRJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="B7A4663DFFE2FF99F66BF873FB11F871" pageId="2" pageNumber="210" startId="2.[85,120,1920,1934]" targetBox="[253,1300,156,1885]" targetPageId="2" targetType="figure">
<paragraph id="E36436B5FFE2FF99F66BF873FB11F871" blockId="2.[85,1474,1919,1957]" pageId="2" pageNumber="210">
<emphasis id="D1AFEAA7FFE2FF99F66BF873FF37F85A" bold="true" box="[85,174,1920,1934]" pageId="2" pageNumber="210">Figs. 110.</emphasis>
<taxonomicName id="24DB4D36FFE2FF99F689F88CFE99F85A" authorityName="MacLeay" authorityYear="1819" box="[183,256,1919,1934]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="D1AFEAA7FFE2FF99F689F88CFE99F85A" box="[183,256,1919,1934]" italics="true" pageId="2" pageNumber="210">Pelidnota</emphasis>
</taxonomicName>
. 18,
<taxonomicName id="24DB4D36FFE2FF99F70EF88CFE2CF85A" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[304,437,1919,1934]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="210" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFE2FF99F70EF88CFE2CF85A" box="[304,437,1919,1934]" italics="true" pageId="2" pageNumber="210">Pelidnota beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE2FF99F784F873FE65F85A" box="[442,508,1920,1934]" pageId="2" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910,
<taxonomicName id="24DB4D36FFE2FF99F738F865FE52F871" authority="(Drury, 1778)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[262,459,1942,1957]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="210" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFE2FF99F738F865FEC1F871" box="[262,344,1942,1957]" italics="true" pageId="2" pageNumber="210">P. unicolor</emphasis>
(Drury, 1778)
</taxonomicName>
(dorsal, lateral). Scale bars: Figs. 16 = 9.0 mm, Figs. 78 = 2.5 mm, Figs. 910 = 8.2 mm.
</paragraph>
</caption>
<paragraph id="E36436B5FFE3FF98F6ACFF64FD8CFF13" blockId="3.[114,785,151,1120]" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F6ACFF64FE9EFF7F" box="[146,263,151,171]" italics="true" pageId="3" pageNumber="211">Distribution</emphasis>
.
<collectingCountry id="9BCC7625FFE3FF98F72FFF6BFEC7FF7F" box="[273,350,152,171]" name="Brazil" pageId="3" pageNumber="211">BRAZIL</collectingCountry>
:
<collectingRegion id="211FF857FFE3FF98F758FF6BFE4CFF7F" box="[358,469,152,171]" country="Brazil" name="Bahia" pageId="3" pageNumber="211">Bahia state</collectingRegion>
(Una, Santa Terezinha, Ituberá, São Félix do Coribe, Itamaraju, Camacan).
</paragraph>
<paragraph id="E36436B5FFE3FF98F6ACFF3CFD73FF2B" blockId="3.[114,785,151,1120]" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F6ACFF3CFF60FF37" box="[146,249,207,227]" italics="true" pageId="3" pageNumber="211">Etymology</emphasis>
. This species was named in honor of Johann Becker who collected a large number of specimens used as
<typeStatus id="3C608817FFE3FF98F441FF1FFD7CFF2B" box="[639,741,236,255]" pageId="3" pageNumber="211" type="paratype">paratypes</typeStatus>
.
</paragraph>
<paragraph id="E36436B5FFE3FF98F6ACFEF5FEBCFCB0" blockId="3.[114,785,151,1120]" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F6ACFEF5FF7FFECE" box="[146,230,262,282]" italics="true" pageId="3" pageNumber="211">Remarks</emphasis>
.
<taxonomicName id="24DB4D36FFE3FF98F6CAFEF5FE07FECE" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[244,414,262,282]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="D1AFEAA7FFE3FF98F6CAFEF5FE07FECE" box="[244,414,262,282]" italics="true" pageId="3" pageNumber="211">Pelidnota beckeri</emphasis>
</taxonomicName>
is distinguished as follows (
<taxonomicName id="24DB4D36FFE3FF98F4F0FEF5FEAAFEE2" authority="(Drury, 1778)" baseAuthorityName="Drury" baseAuthorityYear="1778" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFE3FF98F4F0FEF5FF3BFEE2" italics="true" pageId="3" pageNumber="211">P. unicolor</emphasis>
(Drury, 1778)
</taxonomicName>
characters in brackets): elytra with dark spots (
<figureCitation id="7BE02A30FFE3FF98F644FECCFF40FE86" box="[122,217,319,338]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="3" pageNumber="211">Figs. 13</figureCitation>
) (without dark spots (
<figureCitation id="7BE02A30FFE3FF98F7ECFECCFDD9FE86" box="[466,576,319,338]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="3" pageNumber="211">Figs. 9, 10</figureCitation>
)); second tooth on the inner margin of maxila small and less acute (
<figureCitation id="7BE02A30FFE3FF98F441FEA8FD5CFEBA" box="[639,709,347,366]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 21</figureCitation>
) (large and more acute (
<figureCitation id="7BE02A30FFE3FF98F71FFE84FEFCFE5E" box="[289,357,375,394]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 22</figureCitation>
)); second teeth of maxillary galea sawed, and IV palpomere fusiform (
<figureCitation id="7BE02A30FFE3FF98F7A4FE60FE46FE72" box="[410,479,403,422]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="3" pageNumber="211">Fig. 25</figureCitation>
) (teeth sawed absent, and IV palpomere cylindrical (
<figureCitation id="7BE02A30FFE3FF98F75FFE5CFE3CFE16" box="[353,421,431,450]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="3" pageNumber="211">Fig. 26</figureCitation>
)); surface area of pre-mentum and insertion area of labial palp less excavated, and lateral margin of post-mentum more rounded (
<figureCitation id="7BE02A30FFE3FF98F797FE14FE74FE2E" box="[425,493,487,506]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="3" pageNumber="211">Fig. 29</figureCitation>
) (surface area and insertion palp more excavated, and lateral margin less rounded (
<figureCitation id="7BE02A30FFE3FF98F48AFDF1FD60FDC2" box="[692,761,514,534]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="3" pageNumber="211">Fig. 30</figureCitation>
)); elytral apex truncated (
<figureCitation id="7BE02A30FFE3FF98F752FDEDFE2BFDE6" box="[364,434,542,562]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="3" pageNumber="211">Fig. 33</figureCitation>
) (rounded (
<figureCitation id="7BE02A30FFE3FF98F40AFDEDFDE3FDE6" box="[564,634,542,562]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="3" pageNumber="211">Fig. 34</figureCitation>
)); externodistal tooth of protibia large and strongly curved (
<figureCitation id="7BE02A30FFE3FF98F449FDC9FD26FD99" box="[631,703,570,589]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="3" pageNumber="211">Fig. 41</figureCitation>
) (small and slightly curved (
<figureCitation id="7BE02A30FFE3FF98F772FDA5FE0BFDBD" box="[332,402,598,617]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="3" pageNumber="211">Fig. 42</figureCitation>
)); anteroexternal angle of metacoxa acute (
<figureCitation id="7BE02A30FFE3FF98F688FD81FF61FD51" box="[182,248,626,645]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="3" pageNumber="211">Fig. 37</figureCitation>
) (rounded (
<figureCitation id="7BE02A30FFE3FF98F74EFD81FE2BFD51" box="[368,434,626,645]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="3" pageNumber="211">Fig. 38</figureCitation>
)); apical tubercle of protarsal claw small and rounded (
<figureCitation id="7BE02A30FFE3FF98F702FD7DFEE7FD75" box="[316,382,654,673]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 4752. Anterior tarsal claws of male and left metatibia of female in lateral view.4750, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.4750 = 0.500 mm, Figs.51 and 52 = 1.0mm." pageId="3" pageNumber="211">Fig. 47</figureCitation>
) (large and acute (
<figureCitation id="7BE02A30FFE3FF98F407FD7DFDE2FD75" box="[569,635,654,673]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 4752. Anterior tarsal claws of male and left metatibia of female in lateral view.4750, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.4750 = 0.500 mm, Figs.51 and 52 = 1.0mm." pageId="3" pageNumber="211">Fig. 48</figureCitation>
)); female with metatibial apex straight and with spine-like setae (
<figureCitation id="7BE02A30FFE3FF98F4BAFD59FD50FD69" box="[644,713,682,701]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 4752. Anterior tarsal claws of male and left metatibia of female in lateral view.4750, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.4750 = 0.500 mm, Figs.51 and 52 = 1.0mm." pageId="3" pageNumber="211">Fig. 51</figureCitation>
) (apex with a acute angle and without spine-like setae (
<figureCitation id="7BE02A30FFE3FF98F451FD35FD2DFD0D" box="[623,692,710,729]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 4752. Anterior tarsal claws of male and left metatibia of female in lateral view.4750, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.4750 = 0.500 mm, Figs.51 and 52 = 1.0mm." pageId="3" pageNumber="211">Fig. 52</figureCitation>
)); external surface of metatibia densely punctuated (
<figureCitation id="7BE02A30FFE3FF98F47CFD11FD1CFD21" box="[578,645,738,757]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="3" pageNumber="211">Fig. 53</figureCitation>
) (moderately punctuated (
<figureCitation id="7BE02A30FFE3FF98F6CAFD0DFEAFFCC4" box="[244,310,765,785]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="3" pageNumber="211">Fig. 54</figureCitation>
)); parameres distal margin acute laterally, and with proximal margin slightly corrugated (
<figureCitation id="7BE02A30FFE3FF98F414FCEAFD5BFCF8" box="[554,706,793,812]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="3" pageNumber="211">Figs. 57, 58, 65</figureCitation>
) (distal margin rounded laterally, and proximal margin strongly corrugated (
<figureCitation id="7BE02A30FFE3FF98F644FCA2FE96FCB0" box="[122,271,849,868]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="3" pageNumber="211">Figs. 59, 60, 66</figureCitation>
)).
</paragraph>
<paragraph id="E36436B5FFE3FF98F6ACFC9FFD3AFB8B" blockId="3.[114,785,151,1120]" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F6ACFC9FFE2CFC54" box="[146,437,876,896]" italics="true" pageId="3" pageNumber="211">
Comments.
<taxonomicName id="24DB4D36FFE3FF98F737FC9FFE2CFC54" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[265,437,876,896]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="beckeri">Pelidnota beckeri</taxonomicName>
</emphasis>
is found in different ecoregions (
<bibRefCitation id="874A4B44FFE3FF98F644FC7AFED1FC48" author="Instituto LIFE" box="[122,328,905,924]" pageId="3" pageNumber="211" pagination="35 - 52" refId="ref10592" refString="Instituto LIFE, 2015. Bahia, in: Ecorregioes do Brasil - Prioridades Terrestres e Marinhas. Serie cadernos tecnicos, Volume 3, Brasil., pp. 35 - 52." type="book chapter" year="2015">Instituto LIFE, 2015</bibRefCitation>
) in
<collectingRegion id="211FF857FFE3FF98F747FC7AFE76FC48" box="[377,495,905,924]" country="Brazil" name="Bahia" pageId="3" pageNumber="211">Bahia state</collectingRegion>
.
<materialsCitation id="53B33CE8FFE3FF98F7C3FC7AFD39FB8B" county="Atlantic Rain Forest" location="Sao Felix do Coribe" municipality="Caatinga" pageId="3" pageNumber="211" specimenCount="1" stateProvince="Bahia" typeStatus="paratype">
The new species was collected in rainy coastal forest sites (
<collectingCounty id="0A054E39FFE3FF98F7D3FC56FD59FC6C" box="[493,704,933,952]" pageId="3" pageNumber="211">Atlantic Rain Forest</collectingCounty>
) in the municipalities of Ituberá, Una, Itamaraju, and Camacan; one of the
<typeStatus id="3C608817FFE3FF98F6A3FC2EFE98FC24" box="[157,257,989,1008]" pageId="3" pageNumber="211" type="paratype">paratypes</typeStatus>
was collected in a transition zone between rainy and semi-deciduous forests,
<quantity id="24239B50FFE3FF98F7A7FC0AFE7DFBD8" box="[409,484,1017,1036]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="211" unit="km" value="100.0">100 km</quantity>
inland from the coast (Pedra Branca, municipality of Santa Terezinha);
<specimenCount id="F5DDFD3CFFE3FF98F416FBE6FD29FBFC" box="[552,688,1045,1064]" count="1" pageId="3" pageNumber="211" type="generic" typeStatus="paratype">one paratype</specimenCount>
was collected in a dry forest site (
<collectingMunicipality id="0300ACCFFFE3FF98F74AFBC3FE56FB97" box="[372,463,1072,1091]" pageId="3" pageNumber="211">Caatinga</collectingMunicipality>
) in the municipality of
<location id="E604606EFFE3FF98F486FBC2FF4AFB8B" LSID="urn:lsid:plazi:treatment:6B7287A3FFE1FF98F6ACFE5DFD3AFB8B:E604606EFFE3FF98F486FBC2FF4AFB8B" county="Atlantic Rain Forest" municipality="Caatinga" name="Sao Felix do Coribe" pageId="3" pageNumber="211" stateProvince="Bahia">São Félix do Coribe</location>
near
<location id="E604606EFFE3FF98F732FBBFFE74FB8B" LSID="urn:lsid:plazi:treatment:6B7287A3FFE1FF98F6ACFE5DFD3AFB8B:E604606EFFE3FF98F732FBBFFE74FB8B" box="[268,493,1100,1119]" county="Atlantic Rain Forest" municipality="Caatinga" name="Santa Maria da Vitoria" pageId="3" pageNumber="211" stateProvince="Bahia">Santa Maria da Vitoria</location>
in western
<collectingRegion id="211FF857FFE3FF98F45BFBBFFD39FB8B" box="[613,672,1100,1119]" country="Brazil" name="Bahia" pageId="3" pageNumber="211">Bahia</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,308 @@
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<mods:namePart id="D90F7DBEDE8106E9B7187AAB5E9C86DD">Ferreira, André da Silva</mods:namePart>
<mods:affiliation id="AEC0858B2FA5330D51ADFC33AA6227BB">Universidade Federal da Bahia, Instituto de Biologia, Programa de Pós-Graduação em Diversidade Animal, Bahia, BA, Brazil</mods:affiliation>
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<mods:affiliation id="4F088010B55176E5A5E2A3F57A2B27F9">Universidade Federal do Paraná, Departamento de Zoologia, Laboratório de Sistemática e Bioecologia de Coleoptera, Curitiba, PR, Brazil</mods:affiliation>
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<mods:namePart id="663622BD86F46EA3BB9204411FE9E733">Bravo, Freddy</mods:namePart>
<mods:affiliation id="4D3215451EF9E54964C5B947E3F9BB48">Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Laboratório de Sistemática de Insetos, Feira de Santana, BA, Brazil</mods:affiliation>
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<subSubSection id="ABC1653EFFE3FF98F6ACFB70FE5AFB43" box="[146,451,1155,1175]" pageId="3" pageNumber="211" type="nomenclature">
<paragraph id="E36436B5FFE3FF98F6ACFB70FE5AFB43" blockId="3.[146,451,1155,1175]" box="[146,451,1155,1175]" pageId="3" pageNumber="211">
<heading id="B82C81D9FFE3FF98F6ACFB70FE5AFB43" bold="true" box="[146,451,1155,1175]" fontSize="8" level="2" pageId="3" pageNumber="211" reason="2">
<emphasis id="D1AFEAA7FFE3FF98F6ACFB70FE5AFB43" bold="true" box="[146,451,1155,1175]" pageId="3" pageNumber="211">
<taxonomicName id="24DB4D36FFE3FF98F6ACFB70FEF4FB43" ID-CoL="75ZWV" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[146,365,1155,1175]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE3FF98F6ACFB70FEF4FB43" bold="true" box="[146,365,1155,1175]" italics="true" pageId="3" pageNumber="211">Pelidnota nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE3FF98F74CFB77FE5AFB43" box="[370,451,1156,1175]" pageId="3" pageNumber="211" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ABC1653EFFE3FF9FF6ACFB53FD10F8E7" lastPageId="4" lastPageNumber="212" pageId="3" pageNumber="211" type="description">
<paragraph id="E36436B5FFE3FF98F6ACFB53FE8DFB67" blockId="3.[114,786,1184,1985]" box="[146,276,1184,1203]" pageId="3" pageNumber="211">
(
<figureCitation id="7BE02A30FFE3FF98F6A4FB53FE95FB67" box="[154,268,1184,1203]" captionStart="Figs" captionStartId="4.[85,120,1355,1369]" captionTargetBox="[250,1305,180,1309]" captionTargetId="figure-13@4.[249,1309,149,1324]" captionTargetPageId="4" captionText="Figs. 1116. Pelidnota. 1113, Pelidnota nordestina sp. nov., male holotype (dorsal, ventral, lateral); 1416, P. pernambucana sp. nov., male holotype (dorsal, ventral, lateral). Scale bars: Figs. 1113 =5.5 mm, Figs.1416= 5.0mm." pageId="3" pageNumber="211">Figs. 1113</figureCitation>
)
</paragraph>
<paragraph id="E36436B5FFE3FF98F6ACFB48FED3FA1E" blockId="3.[114,786,1184,1985]" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F6ACFB48FF69FB1B" box="[146,240,1211,1231]" italics="true" pageId="3" pageNumber="211">Diagnosis</emphasis>
. Labrum triangular (
<figureCitation id="7BE02A30FFE3FF98F7EDFB4FFD85FB1B" box="[467,540,1212,1231]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 19</figureCitation>
). Second teeth on the inner margin of mandible larger than first and third (
<figureCitation id="7BE02A30FFE3FF98F482FB2BFC9DFB3F" box="[700,772,1240,1259]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 23</figureCitation>
). Second teeth of maxillary galea sawed (
<figureCitation id="7BE02A30FFE3FF98F427FB07FDC6FAD3" box="[537,607,1268,1287]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="3" pageNumber="211">Fig. 27</figureCitation>
). Surface area of pre-mentum excavated and lateral margin of post-mentum slight rounded (
<figureCitation id="7BE02A30FFE3FF98F6EBFAD8FE8FFAEB" box="[213,278,1323,1343]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="3" pageNumber="211">Fig. 31</figureCitation>
). Anteroexternal coxal angle of metacoxa rounded and little acute (
<figureCitation id="7BE02A30FFE3FF98F72AFAB4FECFFA8F" box="[276,342,1351,1371]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="3" pageNumber="211">Fig. 39</figureCitation>
). Pygidium dark brown, with reddish reflections on disk, with slight metallic green reflections only on lateral portion of basal margin (
<figureCitation id="7BE02A30FFE3FF98F75BFA8CFE3FFA46" box="[357,422,1407,1426]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="3" pageNumber="211">Fig.45</figureCitation>
). Six spine-like setae on the external longitudinal carinae and six setae on the inner longitudinal carinae of metatibia (
<figureCitation id="7BE02A30FFE3FF98F6C5FA44FEA4FA1E" box="[251,317,1463,1482]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="3" pageNumber="211">Fig. 55</figureCitation>
).
</paragraph>
<paragraph id="E36436B5FFE3FF98F6ACFA21FABAF9CA" blockId="3.[114,786,1184,1985]" lastBlockId="3.[833,1504,152,1984]" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F6ACFA21FE14FA32" box="[146,397,1490,1510]" italics="true" pageId="3" pageNumber="211">
<typeStatus id="3C608817FFE3FF98F6ACFA21FF73FA32" box="[146,234,1490,1510]" pageId="3" pageNumber="211" type="holotype">Holotype</typeStatus>
male description
</emphasis>
. Total length
<quantity id="24239B50FFE3FF98F42CFA20FDF2FA32" box="[530,619,1491,1510]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="3" pageNumber="211" unit="mm" value="25.7">25.7 mm</quantity>
, width
<quantity id="24239B50FFE3FF98F48DFA20FC97FA32" box="[691,782,1491,1510]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="3" pageNumber="211" unit="mm" value="13.1">13.1 mm</quantity>
. Body elongated-oval, slightly convex.
<emphasis id="D1AFEAA7FFE3FF98F7C5FA1DFDB7F9D6" box="[507,558,1518,1538]" italics="true" pageId="3" pageNumber="211">Color</emphasis>
: Head golden-brown, with black margin. Pronotum, scutellum, and elytra golden-brown and with metallic green reflections. Anterior margin of pronotum and scutellum dark. Pronotum with one and elytra with two lateral black spots. Pygidium dark brown, with reddish reflections in middle, with slight metallic green reflections only on lateral portion of basal margin. Ventral surface and legs of same color as pygidium, and with metallic green reflections.
<emphasis id="D1AFEAA7FFE3FF98F479F942FDE0F911" box="[583,633,1713,1733]" italics="true" pageId="3" pageNumber="211">Head</emphasis>
: Antenna with 10 antennomeres; club composed by three antennomeres; scape elongated, shorter than antennomeres IIVII. Clypeus trapezoidal, puncture dense and large to moderately large; anterior margin concave. Frontoclypeal suture not evident. Frons width at middle region 23 times length of clypeus; punctures moderate to dense and moderately wide. Ocular canthus not reaching the middle of the eye. Interocular distance 3 times the transverse eye diameters. Labrum triangular and transversely elongated (
<figureCitation id="7BE02A30FFE3FF98F466F862FD05F870" box="[600,668,1937,1956]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 19</figureCitation>
), not fused to clypeus, with medial compression; acuminate anterior margin and posterior convex. Mandibles partially visible in dorsal view, outer margin black, with 2 exposed, recurved, outer teeth; 3 smaller teeth on inner margin, II larger than others; molar robust (
<figureCitation id="7BE02A30FFE3FF98F3AEFF23FA4BFF37" box="[1424,1490,208,227]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 23</figureCitation>
). Maxilla with galea bearing 6 teeth, first and second sawed; palp with 4 palpomeres, IV elongated and fusiform (
<figureCitation id="7BE02A30FFE3FF98F30AFEF4FAE0FECF" box="[1332,1401,263,283]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="3" pageNumber="211">Fig. 27</figureCitation>
). Labium, mentum with apex concave; post-mentum with elongated dense setae laterally; palp inserted into a lateral cavity in pre-mentum, with 3 palpomeres, III elongate (
<figureCitation id="7BE02A30FFE3FF98F2BBFEA8FB5FFEBA" box="[1157,1222,347,366]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="3" pageNumber="211">Fig. 31</figureCitation>
).
<emphasis id="D1AFEAA7FFE3FF98F2E6FEA9FAA0FEBA" box="[1240,1337,346,366]" italics="true" pageId="3" pageNumber="211">Pronotum</emphasis>
: Convex. Length
<quantity id="24239B50FFE3FF98F57FFE84FC14FE5E" box="[833,909,375,394]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="211" unit="mm" value="6.3">6.3 mm</quantity>
, anterior width
<quantity id="24239B50FFE3FF98F205FE84FB1EFE5E" box="[1083,1159,375,394]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="3" pageNumber="211" unit="mm" value="7.6">7.6 mm</quantity>
, mean width
<quantity id="24239B50FFE3FF98F322FE84FAECFE5E" box="[1308,1397,375,394]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="211" unit="mm" value="11.5">11.5 mm</quantity>
, posterior width
<quantity id="24239B50FFE3FF98F5B8FE60FC79FE72" box="[902,992,403,422]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="211" unit="mm" value="11.5">11.5 mm</quantity>
. Punctures dense and moderately wide. Anterior margin concave, with anterior angles acute. Posterior margin convex.Lateral margin rounded. Membranous border produced weakly anterior to the anterior marginal bead.
<emphasis id="D1AFEAA7FFE3FF98F2E6FE15FAA1FE2E" box="[1240,1336,486,506]" italics="true" pageId="3" pageNumber="211">Scutellum</emphasis>
: Width
<quantity id="24239B50FFE3FF98F3B1FE14FA42FE2E" box="[1423,1499,487,506]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="211" unit="mm" value="0.3">0.3 mm</quantity>
, total length 1.0 mm. Punctures moderate and moderately large. Posterior margin rounded. Lacking setae.
<emphasis id="D1AFEAA7FFE3FF98F2D3FDEEFAADFDE5" box="[1261,1332,541,561]" italics="true" pageId="3" pageNumber="211">Elytron</emphasis>
: Humeral width
<quantity id="24239B50FFE3FF98F57FFDC9FC03FD9A" box="[833,922,570,590]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="3" pageNumber="211" unit="mm" value="12.9">12.9 mm</quantity>
, total length
<quantity id="24239B50FFE3FF98F21FFDC9FBE2FD9A" box="[1057,1147,570,590]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="3" pageNumber="211" unit="mm" value="17.2">17.2 mm</quantity>
. Surface smooth (punctures sparse to moderate, and small to moderate, slightly organized longitudinally), lacking setae. Membranous border present at the anterior half. Two dark spots present, one in humerus and other in lateral margin. Apex with punctures moderate and moderately wide. Elytra with dark brown suture and truncate apex (
<figureCitation id="7BE02A30FFE3FF98F369FD35FA05FD0D" box="[1367,1436,710,729]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="3" pageNumber="211">Fig. 35</figureCitation>
).
<emphasis id="D1AFEAA7FFE3FF98F38CFD36FCC5FD21" italics="true" pageId="3" pageNumber="211">Venter</emphasis>
: Punctures dense and moderately wide. Moderately covered by brown setae. Prosternum elongated in the shape of a tubercle but not extending beyond procoxae. Mesoventrite with anterior process between mesocoxae. Mesoventrite medial area with sparse setae.
<emphasis id="D1AFEAA7FFE3FF98F541FCA3FC31FCB0" box="[895,936,848,868]" italics="true" pageId="3" pageNumber="211">Legs</emphasis>
: Punctures dense. Protibia with dark outer margin and 3 teeth, apical tooth slightly recurved and larger than the other two; inner apical spur present; dorsal longitudinal ridge present and with a row of 8 brown setae (
<figureCitation id="7BE02A30FFE3FF98F255FC56FB34FC6C" box="[1131,1197,933,952]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="3" pageNumber="211">Fig. 43</figureCitation>
); inner protarsal claw slightly recurved and larger than outer claw (
<figureCitation id="7BE02A30FFE3FF98F280FC32FA98FC00" box="[1214,1281,961,980]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 4752. Anterior tarsal claws of male and left metatibia of female in lateral view.4750, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.4750 = 0.500 mm, Figs.51 and 52 = 1.0mm." pageId="3" pageNumber="211">Fig. 49</figureCitation>
). Metacoxa with posterior projection beyond base of metatrochanter; anteroexternal angle rounded and slightly sharp (
<figureCitation id="7BE02A30FFE3FF98F293FC0AFB6AFBD8" box="[1197,1267,1017,1036]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="3" pageNumber="211">Fig. 39</figureCitation>
). Metafemur flattened medially. Mesotibia with setae moderate on inner surface, spine-like setae sparse on outer surface; apical margin dark, inner margin with two larger spurs interleaved with two smaller spine-like setae, outer margin with 10 smaller spine-like setae. Metatibia with apical margin dark, inner margin with two larger spurs interleaved with two smaller spine-like setae, outer margin with one small spine-like setae; external longitudinal carinae with six spinae-like setae and inner longitudinal carinae with six setae present (
<figureCitation id="7BE02A30FFE3FF98F3AFFB2BFA4BFB3F" box="[1425,1490,1240,1259]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="3" pageNumber="211">Fig.55</figureCitation>
). Tarsomeres with small ventral tooth, most evident on meso- and metatarsomeres V.Tarsomeres 24, with two inner spine-like setae on all legs. Tarsal claws simple. Outer claws of meso- and metatarsus larger than inner claws.
<emphasis id="D1AFEAA7FFE3FF98F25EFAB5FB25FA8E" box="[1120,1212,1350,1370]" italics="true" pageId="3" pageNumber="211">Abdomen</emphasis>
: Abdominal ventrite convex and punctures moderate to dense, and with sparse setae. Ventrite VI ending before posterior margin of pygidium.
<emphasis id="D1AFEAA7FFE3FF98F31AFA8DFA19FA46" box="[1316,1408,1406,1426]" italics="true" pageId="3" pageNumber="211">Pygidium</emphasis>
: (
<figureCitation id="7BE02A30FFE3FF98F3ABFA8CFA41FA46" box="[1429,1496,1407,1426]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="3" pageNumber="211">Fig. 45</figureCitation>
) slightly convex in lateral view. Surface densely striate and with moderately dense setae. Setae whitish, moderately dense. A gradual concavity present on posterior margin.
<emphasis id="D1AFEAA7FFE3FF98F2F5FA21FAA8FA32" box="[1227,1329,1490,1510]" italics="true" pageId="3" pageNumber="211">Parameres</emphasis>
: (
<figureCitation id="7BE02A30FFE3FF98F378FA20FA41FA32" box="[1350,1496,1491,1510]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="3" pageNumber="211">Figs. 61, 62, 67</figureCitation>
) symmetrical, fused; distal margin V-shaped, gradually acute laterally; proximal margin gradually semi-quadrate.
</paragraph>
<paragraph id="E36436B5FFE3FF98F55EF9D6FA46F9A5" blockId="3.[833,1504,152,1984]" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F55EF9D6FC72F9ED" box="[864,1003,1573,1593]" italics="true" pageId="3" pageNumber="211">Type material</emphasis>
.
<materialsCitation id="53B33CE8FFE3FF98F5C4F9D4FA46F9A5" collectingDate="2015-03-25" collectionCode="MZFS" collectorName="E. M. Menezes" country="Brazil" elevation="510" location="Cachoeira Veu da Noiva" municipality="Bonito" pageId="3" pageNumber="211" specimenCode="MZFS#55726" specimenCount="1" stateProvince="Pernambuco" typeStatus="holotype">
<typeStatus id="3C608817FFE3FF98F5C4F9D4FBCEF9EE" box="[1018,1111,1575,1594]" pageId="3" pageNumber="211" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="9BCC7625FFE3FF98F29DF9D4FB74F9EE" box="[1187,1261,1575,1594]" name="Brazil" pageId="3" pageNumber="211">BRAZIL</collectingCountry>
,
<emphasis id="D1AFEAA7FFE3FF98F2C4F9D6FAEAF9ED" box="[1274,1395,1573,1593]" italics="true" pageId="3" pageNumber="211">
<collectingRegion id="211FF857FFE3FF98F2C4F9D6FAEAF9ED" box="[1274,1395,1573,1593]" country="Brazil" name="Pernambuco" pageId="3" pageNumber="211">Pernambuco</collectingRegion>
</emphasis>
: (“
<collectingMunicipality id="0300ACCFFFE3FF98F3A8F9D4FA45F9EE" box="[1430,1500,1575,1594]" pageId="3" pageNumber="211">Bonito</collectingMunicipality>
,
<location id="E604606EFFE3FF98F57FF9B1FBDBF981" LSID="urn:lsid:plazi:treatment:6B7287A3FFE3FF9FF6ACFB70FD10F8E7:E604606EFFE3FF98F57FF9B1FBDBF981" box="[833,1090,1602,1622]" country="Brazil" municipality="Bonito" name="Cachoeira Veu da Noiva" pageId="3" pageNumber="211" stateProvince="Pernambuco">Cachoeira Véu da Noiva</location>
”, “
<quantity id="24239B50FFE3FF98F25CF9B1FB38F982" box="[1122,1185,1602,1622]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="211" unit="m" value="510.0">
<elevation id="68F6D186FFE3FF98F25CF9B1FB38F982" box="[1122,1185,1602,1622]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="211" unit="m" value="510.0">510 m</elevation>
</quantity>
”) (8
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32
<emphasis id="D1AFEAA7FFE3FF98F2CBF9CDFB60F998" bold="true" box="[1269,1273,1598,1612]" pageId="3" pageNumber="211">
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</emphasis>
32.4
<emphasis id="D1AFEAA7FFE3FF98F318F9CDFAB6F998" bold="true" box="[1318,1327,1598,1612]" pageId="3" pageNumber="211">
<superScript id="14AE9BFDFFE3FF98F318F9CDFAB6F998" attach="right" box="[1318,1327,1598,1612]" fontSize="6" pageId="3" pageNumber="211">ll</superScript>
</emphasis>
S/35
<emphasis id="D1AFEAA7FFE3FF98F363F9CDFAFCF998" bold="true" box="[1373,1381,1598,1612]" pageId="3" pageNumber="211">
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</emphasis>
42
<emphasis id="D1AFEAA7FFE3FF98F3BEF9CDFA1DF998" bold="true" box="[1408,1412,1598,1612]" pageId="3" pageNumber="211">
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</emphasis>
53.9
<emphasis id="D1AFEAA7FFE3FF98F38FF9CDFA23F998" bold="true" box="[1457,1466,1598,1612]" pageId="3" pageNumber="211">
<superScript id="14AE9BFDFFE3FF98F38FF9CDFA23F998" attach="right" box="[1457,1466,1598,1612]" fontSize="6" pageId="3" pageNumber="211">ll</superScript>
</emphasis>
W),
<date id="97651075FFE3FF98F57FF9ADFC37F9A5" box="[833,942,1630,1649]" pageId="3" pageNumber="211" value="2015-03-25">
<collectingDate id="8721E99DFFE3FF98F57FF9ADFC37F9A5" box="[833,942,1630,1649]" pageId="3" pageNumber="211" value="2015-03-25">25.III.2015</collectingDate>
</date>
.
<collectorName id="4E2E5363FFE3FF98F588F9ACFBD8F9A6" box="[950,1089,1631,1650]" pageId="3" pageNumber="211">E.M. Menezes</collectorName>
col. (
<specimenCode id="B37D9ECEFFE3FF98F24AF9ADFB66F9A5" box="[1140,1279,1630,1649]" collectionCode="MZFS" pageId="3" pageNumber="211">MZFS#55726</specimenCode>
) (
<collectionCode id="85CAAE70FFE3FF98F32FF9ADFAD4F9A5" box="[1297,1357,1630,1649]" pageId="3" pageNumber="211">MZFS</collectionCode>
) (
<figureCitation id="7BE02A30FFE3FF98F35EF9ADFA4BF9A5" box="[1376,1490,1630,1649]" captionStart="Figs" captionStartId="13.[114,149,1725,1739]" captionTargetBox="[179,720,149,1695]" captionTargetId="figure-13@13.[179,720,149,1695]" captionTargetPageId="13" captionText="Figs.6977. Labels attachedtothe hotypes.6971,Pelidnota beckerisp.nov.;7274, P. nordestina sp. nov.; 7577, P.pernambucana sp. nov." pageId="3" pageNumber="211">Figs. 7274</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="E36436B5FFE3FF98F55EF98AFA82F959" blockId="3.[833,1504,152,1984]" box="[864,1307,1657,1678]" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F55EF98AFC45F959" box="[864,988,1657,1677]" italics="true" pageId="3" pageNumber="211">Distribution.</emphasis>
Known only from
<typeStatus id="3C608817FFE3FF98F2A9F988FB5DF95A" box="[1175,1220,1659,1678]" pageId="3" pageNumber="211">type</typeStatus>
locality.
</paragraph>
<paragraph id="E36436B5FFE3FF98F55EF966FBD8F911" blockId="3.[833,1504,152,1984]" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F55EF966FC5EF97D" box="[864,967,1685,1705]" italics="true" pageId="3" pageNumber="211">Etymology</emphasis>
. The specific epithet
<taxonomicName id="24DB4D36FFE3FF98F293F966FA8CF97D" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1197,1301,1685,1705]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="nordestina">
<emphasis id="D1AFEAA7FFE3FF98F293F966FA8CF97D" box="[1197,1301,1685,1705]" italics="true" pageId="3" pageNumber="211">nordestina</emphasis>
</taxonomicName>
refers to its origin from northeastern
<collectingCountry id="9BCC7625FFE3FF98F23FF941FBA4F911" box="[1025,1085,1714,1733]" name="Brazil" pageId="3" pageNumber="211">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="E36436B5FFE3FF9FF55EF93EFECAF928" blockId="3.[833,1504,152,1984]" lastBlockId="4.[85,756,1434,1844]" lastPageId="4" lastPageNumber="212" pageId="3" pageNumber="211">
<emphasis id="D1AFEAA7FFE3FF98F55EF93EFB11F935" box="[864,1160,1741,1761]" italics="true" pageId="3" pageNumber="211">
Remarks.
<taxonomicName id="24DB4D36FFE3FF98F581F93EFB11F935" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[959,1160,1741,1761]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="nordestina">Pelidnota nordestina</taxonomicName>
</emphasis>
is distinguished as follows (
<taxonomicName id="24DB4D36FFE3FF98F399F93EFC22F929" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="211" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="D1AFEAA7FFE3FF98F399F93EFC22F929" italics="true" pageId="3" pageNumber="211">Pelidnota beckeri</emphasis>
</taxonomicName>
characters in brackets): golden-brown with metallic green reflections, especially on the pronotum and ventral surface, and a black spot is present on both sides of pronotum, with two black spots on the elytron (
<figureCitation id="7BE02A30FFE3FF98F272F8CEFB20F885" box="[1100,1209,1853,1873]" captionStart="Figs" captionStartId="4.[85,120,1355,1369]" captionTargetBox="[250,1305,180,1309]" captionTargetId="figure-13@4.[249,1309,149,1324]" captionTargetPageId="4" captionText="Figs. 1116. Pelidnota. 1113, Pelidnota nordestina sp. nov., male holotype (dorsal, ventral, lateral); 1416, P. pernambucana sp. nov., male holotype (dorsal, ventral, lateral). Scale bars: Figs. 1113 =5.5 mm, Figs.1416= 5.0mm." pageId="3" pageNumber="211">Figs.11, 13</figureCitation>
) (pronotum and ventral body surface dark brown, with no spots on pronotum but three black spots on elytron (
<figureCitation id="7BE02A30FFE3FF98F5CDF886FBD3F85C" box="[1011,1098,1909,1928]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="3" pageNumber="211">Figs. 1, 3</figureCitation>
)); labrum triangular (
<figureCitation id="7BE02A30FFE3FF98F317F886FAF5F85C" box="[1321,1388,1909,1928]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 19</figureCitation>
) (transversal (
<figureCitation id="7BE02A30FFE3FF98F552F862FC36F870" box="[876,943,1937,1956]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 17</figureCitation>
)); inner teeth of mandible more acuminate and larger (
<figureCitation id="7BE02A30FFE3FF98F577F85EFC15F814" box="[841,908,1965,1984]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 23</figureCitation>
) (small (
<figureCitation id="7BE02A30FFE3FF98F5D9F85EFBB3F814" box="[999,1066,1965,1984]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="3" pageNumber="211">Fig. 21</figureCitation>
)); first and second teeth of maxillary galea sawed, and IV palpomere more elongated (
<figureCitation id="7BE02A30FFE4FF9FF424FA69FDC6FA79" box="[538,607,1434,1453]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="4" pageNumber="212">Fig. 27</figureCitation>
) (only second teeth sawed, and IV palpomere less longer (
<figureCitation id="7BE02A30FFE4FF9FF419FA45FDF4FA1D" box="[551,621,1462,1481]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="4" pageNumber="212">Fig. 25</figureCitation>
)); excavated area on the surface of pre-mentum larger, and lateral margin of post-mentum less rounded and with elongated dense setae (
<figureCitation id="7BE02A30FFE4FF9FF492FA1EFD75F9D5" box="[684,748,1517,1537]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="4" pageNumber="212">Fig.31</figureCitation>
) (excavated area small, lateral margin more rounded, and with moderate setae (
<figureCitation id="7BE02A30FFE4FF9FF6E8F9D6FE82F9EC" box="[214,283,1573,1592]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="4" pageNumber="212">Fig. 29</figureCitation>
)); anteroexternal angle of metacoxa rounded and slightly sharp (
<figureCitation id="7BE02A30FFE4FF9FF723F9B2FEFBF980" box="[285,354,1601,1620]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="4" pageNumber="212">Fig. 39</figureCitation>
) (acute (
<figureCitation id="7BE02A30FFE4FF9FF781F9B2FD9AF980" box="[447,515,1601,1620]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="4" pageNumber="212">Fig. 37</figureCitation>
)); protarsal tubercle of male located more on the claw apex (
<figureCitation id="7BE02A30FFE4FF9FF7DCF9AEFDBEF9A4" box="[482,551,1629,1648]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 4752. Anterior tarsal claws of male and left metatibia of female in lateral view.4750, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.4750 = 0.500 mm, Figs.51 and 52 = 1.0mm." pageId="4" pageNumber="212">Fig. 49</figureCitation>
) (located not so on the claw apex (
<figureCitation id="7BE02A30FFE4FF9FF6D1F98AFEABF958" box="[239,306,1657,1676]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 4752. Anterior tarsal claws of male and left metatibia of female in lateral view.4750, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.4750 = 0.500 mm, Figs.51 and 52 = 1.0mm." pageId="4" pageNumber="212">Fig. 47</figureCitation>
)); external surface of metatibia less punctuated, and inner longitudinal carinae with six setae (
<figureCitation id="7BE02A30FFE4FF9FF45BF966FD31F97C" box="[613,680,1685,1704]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="4" pageNumber="212">Fig. 55</figureCitation>
) (more punctuated and with more than six setae (
<figureCitation id="7BE02A30FFE4FF9FF434F942FDD7F910" box="[522,590,1713,1732]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="4" pageNumber="212">Fig. 53</figureCitation>
)); shapes of the parameres distal margin, any less acute laterally (
<figureCitation id="7BE02A30FFE4FF9FF47CF93EFD36F934" box="[578,687,1741,1760]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="4" pageNumber="212">Figs. 61, 67</figureCitation>
) (apex more acute (
<figureCitation id="7BE02A30FFE4FF9FF6EBF91BFEDFF92F" box="[213,326,1768,1787]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="4" pageNumber="212">Figs. 57, 65</figureCitation>
).
</paragraph>
<caption id="B7A4663DFFE4FF9FF66BFAB8FD94FAA4" pageId="4" pageNumber="212" startId="4.[85,120,1355,1369]" targetBox="[250,1305,180,1309]" targetPageId="4" targetType="figure">
<paragraph id="E36436B5FFE4FF9FF66BFAB8FD94FAA4" blockId="4.[85,1476,1354,1392]" pageId="4" pageNumber="212">
<emphasis id="D1AFEAA7FFE4FF9FF66BFAB8FF21FA8D" bold="true" box="[85,184,1355,1369]" pageId="4" pageNumber="212">Figs. 1116.</emphasis>
<taxonomicName id="24DB4D36FFE4FF9FF6FFFAB9FE93FA8D" authorityName="MacLeay" authorityYear="1819" box="[193,266,1354,1369]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="212" phylum="Arthropoda" rank="genus">
<emphasis id="D1AFEAA7FFE4FF9FF6FFFAB9FE93FA8D" box="[193,266,1354,1369]" italics="true" pageId="4" pageNumber="212">Pelidnota</emphasis>
</taxonomicName>
. 1113,
<taxonomicName id="24DB4D36FFE4FF9FF770FAB9FE77FA8D" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[334,494,1354,1369]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="212" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE4FF9FF770FAB9FE77FA8D" box="[334,494,1354,1369]" italics="true" pageId="4" pageNumber="212">Pelidnota nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE4FF9FF7CDFAB8FDAAFA8D" box="[499,563,1355,1369]" pageId="4" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (dorsal, ventral, lateral); 1416,
<taxonomicName id="24DB4D36FFE4FF9FF588FAB9FBA4FA8D" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[950,1085,1354,1369]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="212" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE4FF9FF588FAB9FBA4FA8D" box="[950,1085,1354,1369]" italics="true" pageId="4" pageNumber="212">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE4FF9FF27FFAB8FB18FA8D" box="[1089,1153,1355,1369]" pageId="4" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
, male holotype (dorsal, ventral, lateral). Scale bars: Figs. 1113 = 5.5 mm, Figs. 1416 = 5.0 mm.
</paragraph>
</caption>
<paragraph id="E36436B5FFE4FF9FF64BF8F0FD10F8E7" blockId="4.[85,756,1434,1844]" pageId="4" pageNumber="212">
<emphasis id="D1AFEAA7FFE4FF9FF64BF8F0FF7BF8C3" box="[117,226,1795,1815]" italics="true" pageId="4" pageNumber="212">Comments.</emphasis>
The specimen was collected in a tropical forest in an enclave in the Caatinga ecoregion (
<bibRefCitation id="874A4B44FFE4FF9FF789F8D3FDE2F8E7" author="Instituto LIFE" box="[439,635,1824,1844]" pageId="4" pageNumber="212" pagination="35 - 52" refId="ref10592" refString="Instituto LIFE, 2015. Bahia, in: Ecorregioes do Brasil - Prioridades Terrestres e Marinhas. Serie cadernos tecnicos, Volume 3, Brasil., pp. 35 - 52." type="book chapter" year="2015">Instituto LIFE, 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,760 @@
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<mods:title id="074F624813A247612C62C50C9C759735">Three new species of Pelidnota MacLeay (Coleoptera, Scarabaeidae, Rutelinae) and new distributional records from northeast Brazil</mods:title>
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<mods:namePart id="9798BB2055A77246ECF5A2DC10CE3211">Ferreira, André da Silva</mods:namePart>
<mods:affiliation id="43C38909E676A40C666D7F6517FFEC1F">Universidade Federal da Bahia, Instituto de Biologia, Programa de Pós-Graduação em Diversidade Animal, Bahia, BA, Brazil</mods:affiliation>
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<mods:namePart id="4BD77991DF4B1FC9D9D5AA35609578C0">Almeida, Lúcia M.</mods:namePart>
<mods:affiliation id="A924F340507ED53ED4383FB511A70F17">Universidade Federal do Paraná, Departamento de Zoologia, Laboratório de Sistemática e Bioecologia de Coleoptera, Curitiba, PR, Brazil</mods:affiliation>
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<mods:namePart id="14D736D656A8F507FDD985478A0C6877">Bravo, Freddy</mods:namePart>
<mods:affiliation id="3FD5FA502FE4D2D7E50D98E7713FF9A2">Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Laboratório de Sistemática de Insetos, Feira de Santana, BA, Brazil</mods:affiliation>
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<treatment id="6B7287A3FFE4FF96F64BF8A4FA79F814" LSID="urn:lsid:plazi:treatment:6B7287A3FFE4FF96F64BF8A4FA79F814" httpUri="http://treatment.plazi.org/id/6B7287A3FFE4FF96F64BF8A4FA79F814" lastPageId="13" lastPageNumber="221" pageId="4" pageNumber="212">
<subSubSection id="ABC1653EFFE4FF9FF64BF8A4FE4AF8BF" box="[117,467,1879,1899]" pageId="4" pageNumber="212" type="nomenclature">
<paragraph id="E36436B5FFE4FF9FF64BF8A4FE4AF8BF" blockId="4.[117,467,1879,1899]" box="[117,467,1879,1899]" pageId="4" pageNumber="212">
<heading id="B82C81D9FFE4FF9FF64BF8A4FE4AF8BF" bold="true" box="[117,467,1879,1899]" fontSize="8" level="2" pageId="4" pageNumber="212" reason="2">
<emphasis id="D1AFEAA7FFE4FF9FF64BF8A4FE4AF8BF" bold="true" box="[117,467,1879,1899]" pageId="4" pageNumber="212">
<taxonomicName id="24DB4D36FFE4FF9FF64BF8A4FEE5F8BF" ID-CoL="6TZWZ" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[117,380,1879,1899]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="212" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE4FF9FF64BF8A4FEE5F8BF" bold="true" box="[117,380,1879,1899]" italics="true" pageId="4" pageNumber="212">Pelidnota pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE4FF9FF7BCF8ABFE4AF8BF" box="[386,467,1880,1899]" pageId="4" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ABC1653EFFE4FF96F64BF887FA79F814" lastPageId="13" lastPageNumber="221" pageId="4" pageNumber="212" type="description">
<paragraph id="E36436B5FFE4FF9FF64BF887FF61F853" blockId="4.[85,756,1908,1983]" box="[117,248,1908,1927]" pageId="4" pageNumber="212">
(
<figureCitation id="7BE02A30FFE4FF9FF643F887FF69F853" box="[125,240,1908,1927]" captionStart="Figs" captionStartId="4.[85,120,1355,1369]" captionTargetBox="[250,1305,180,1309]" captionTargetId="figure-13@4.[249,1309,149,1324]" captionTargetPageId="4" captionText="Figs. 1116. Pelidnota. 1113, Pelidnota nordestina sp. nov., male holotype (dorsal, ventral, lateral); 1416, P. pernambucana sp. nov., male holotype (dorsal, ventral, lateral). Scale bars: Figs. 1113 =5.5 mm, Figs.1416= 5.0mm." pageId="4" pageNumber="212">Figs. 1416</figureCitation>
)
</paragraph>
<paragraph id="E36436B5FFE4FF9FF64BF87CFD6DF86B" blockId="4.[85,756,1908,1983]" pageId="4" pageNumber="212">
<emphasis id="D1AFEAA7FFE4FF9FF64BF87CFF4AF877" box="[117,211,1935,1955]" italics="true" pageId="4" pageNumber="212">Diagnosis</emphasis>
. Second teeth on the external margin of mandible four times than first; second teeth of inner margin bigger than others
</paragraph>
<paragraph id="E36436B5FFE4FF9FF51AFA69FCE2F9EC" blockId="4.[804,1475,1434,1983]" pageId="4" pageNumber="212">
(
<figureCitation id="7BE02A30FFE4FF9FF512FA69FCEBFA79" box="[812,882,1434,1453]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="4" pageNumber="212">Fig. 24</figureCitation>
). First teeth of maxillary galea sawed and IV palpomere elongated (
<figureCitation id="7BE02A30FFE4FF9FF5A4FA45FC78FA1D" box="[922,993,1462,1481]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="4" pageNumber="212">Fig. 28</figureCitation>
). Surface area of pre-mentum excavated and lateral margin of post-mentum rounded (
<figureCitation id="7BE02A30FFE4FF9FF2FEFA21FA98FA31" box="[1216,1281,1490,1509]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="4" pageNumber="212">Fig.32</figureCitation>
). Elytral apex truncated and slightly acute (
<figureCitation id="7BE02A30FFE4FF9FF222FA1EFBC7F9D4" box="[1052,1118,1517,1536]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="4" pageNumber="212">Fig. 36</figureCitation>
). Anteroexternal angle of metacoxa strongly acute (
<figureCitation id="7BE02A30FFE4FF9FF5F8F9FAFB93F9C8" box="[966,1034,1545,1564]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="4" pageNumber="212">Fig. 40</figureCitation>
). Pygidium green with metallic reflections (
<figureCitation id="7BE02A30FFE4FF9FF512F9D6FCF6F9EC" box="[812,879,1573,1592]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="4" pageNumber="212">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="E36436B5FFE4FF9CF57AF9B3FC8BF9F2" blockId="4.[804,1475,1434,1983]" lastBlockId="7.[114,786,1332,1965]" lastPageId="7" lastPageNumber="215" pageId="4" pageNumber="212">
<emphasis id="D1AFEAA7FFE4FF9FF57AF9B3FBDCF980" box="[836,1093,1600,1620]" italics="true" pageId="4" pageNumber="212">
<typeStatus id="3C608817FFE4FF9FF57AF9B3FC05F980" box="[836,924,1600,1620]" pageId="4" pageNumber="212" type="holotype">Holotype</typeStatus>
male description.
</emphasis>
Total length
<quantity id="24239B50FFE4FF9FF2FAF9B2FA84F981" box="[1220,1309,1601,1621]" metricMagnitude="-2" metricUnit="m" metricValue="2.33" pageId="4" pageNumber="212" unit="mm" value="23.3">23.3 mm</quantity>
, width
<quantity id="24239B50FFE4FF9FF35BF9B2FA26F981" box="[1381,1471,1601,1621]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="4" pageNumber="212" unit="mm" value="11.2">11.2 mm</quantity>
. Body elongated-oval, slightly convex.
<emphasis id="D1AFEAA7FFE4FF9FF2F8F9AFFB60F9A4" box="[1222,1273,1628,1648]" italics="true" pageId="4" pageNumber="212">Color</emphasis>
: Head, pronotum, scutellum, and elytron golden-brown. Posterior margin of vertex black. Anterior margin of pronotum dark brown. Medial portion of posterior margin of pronotum, posterior margin of scutellum with metallic green reflections. Pronotum with one and elytra with two lateral black spots. Pygidium dark brown. Ventral surface and legs same color as pygidium. Pygidium, ventral surface and legs with metallic green reflections.
<emphasis id="D1AFEAA7FFE4FF9FF207F8ECFBF2F8E7" box="[1081,1131,1823,1843]" italics="true" pageId="4" pageNumber="212">Head</emphasis>
: Antenna with 10 antennomers; club composed by three antennomeres; scape elongated, shorter than IIVII. Clypeus trapezoidal, punctures dense and large to moderately large; anterior margin concave. Frontoclypeal suture not evident. Frons width at meddle region 23 times length of clypeus; punctures moderate to dense and moderately wide. Ocular canthus not reaching the middle of the eye; interocular distance 4 times the transverse eye diameters. Labrum transversal (
<figureCitation id="7BE02A30FFE5FF9EF474F8C3FD15F897" box="[586,652,1840,1859]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="5" pageNumber="213">Fig. 20</figureCitation>
) not fused to clypeus, with medial compression,setae on the entire surface; anterior and posterior margin straight. Mandibles extend beyond apical margin of clypeus, outer margin black, with 2 exposed, recurved, outer teeth, and 3 teeth on internal, the first worn and second acute (
<figureCitation id="7BE02A30FFE5FF9EF577F8E7FC16F8FC" box="[841,911,1812,1832]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="5" pageNumber="213">Fig. 24</figureCitation>
); molar robust. Maxilla with galea bearing 6 teeth, first sawed; palp with 4 palpomeres, IV elongated (
<figureCitation id="7BE02A30FFE5FF9EF30CF8C3FAE1F897" box="[1330,1400,1840,1859]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="5" pageNumber="213">Fig. 28</figureCitation>
). Labium, post-mentum with elongate moderate setae; mentum with apex concave; palp inserted into a lateral cavity in the pre-mentum, with 3 palpomeres, III elongate, equal III together (
<figureCitation id="7BE02A30FFE5FF9EF374F877FA15F843" box="[1354,1420,1924,1943]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="5" pageNumber="213">Fig. 32</figureCitation>
).
<emphasis id="D1AFEAA7FFE5FF9EF3A3F870FCF0F867" italics="true" pageId="5" pageNumber="213">Pronotum</emphasis>
: Convex. Length
<quantity id="24239B50FFE5FF9EF224F853FBFFF867" box="[1050,1126,1952,1971]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="213" unit="mm" value="4.8">4.8 mm</quantity>
, anterior width
<quantity id="24239B50FFE5FF9EF332F853FAC0F867" box="[1292,1369,1952,1971]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="5" pageNumber="213" unit="mm" value="5.9">5.9 mm</quantity>
, mean width
<quantity id="24239B50FFE6FF9DF66BF9F6FF37F9CC" box="[85,174,1541,1560]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="6" pageNumber="214" unit="mm" value="10.3">10.3 mm</quantity>
, posterior width
<quantity id="24239B50FFE6FF9DF760F9F6FE5CF9CC" box="[350,453,1541,1560]" metricMagnitude="-2" metricUnit="m" metricValue="1.022" pageId="6" pageNumber="214" unit="mm" value="10.22">10.22 mm</quantity>
. Punctures dense and moderately wide. Anterior margin concave, with anterior angles acute. Posterior margin convex. Lateral margin rounded. Membranous border surrounding pronotal disk.
<emphasis id="D1AFEAA7FFE6FF9DF786F9A4FD81F9BF" box="[440,536,1623,1643]" italics="true" pageId="6" pageNumber="214">Scutellum</emphasis>
: Width
<quantity id="24239B50FFE6FF9DF450F9ABFD23F9B8" box="[622,698,1624,1644]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="214" unit="mm" value="0.3">0.3 mm</quantity>
, total length
<quantity id="24239B50FFE6FF9DF6A4F987FF7EF953" box="[154,231,1652,1671]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="214" unit="mm" value="1.2">1.2 mm</quantity>
.Punctures moderate and moderately wide.Posterior margin rounded. Lacking setae.
<emphasis id="D1AFEAA7FFE6FF9DF7A3F97CFE7DF977" box="[413,484,1679,1699]" italics="true" pageId="6" pageNumber="214">Elytron</emphasis>
: Humeral width
<quantity id="24239B50FFE6FF9DF4A9F963FD69F977" box="[663,752,1680,1699]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="6" pageNumber="214" unit="mm" value="11.2">11.2 mm</quantity>
, elytral length
<quantity id="24239B50FFE6FF9DF6DDF95FFEA4F96B" box="[227,317,1708,1727]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="6" pageNumber="214" unit="mm" value="16.4">16.4 mm</quantity>
. Surface smooth (punctures sparse to moderate, small to moderate, slightly organized longitudinally), lacking setae. Membranous border present at the anterior half. Two dark spots present, one in humerus and other in lateral margin. Apex with punctures sparse and moderately wide. Elytra with suture dark brown and truncated, and slightly acute apex (
<figureCitation id="7BE02A30FFE6FF9DF450F8C4FD2BF89E" box="[622,690,1847,1866]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="6" pageNumber="214">Fig. 36</figureCitation>
).
<emphasis id="D1AFEAA7FFE6FF9DF4F8F8C5FFE9F8B2" italics="true" pageId="6" pageNumber="214">Venter</emphasis>
: Punctures dense and moderately wide. Moderately covered by yellow setae. Prosternum with anterior process, but not extending beyond procoxa. Mesoventrite with anterior process between mesocoxae. Mesoventrite lateral area with moderate setae.
<emphasis id="D1AFEAA7FFE6FF9DF4FDF855FD75F86E" box="[707,748,1958,1978]" italics="true" pageId="6" pageNumber="214">Legs</emphasis>
: Punctures dense. Protibia with dark outer margin and 3 teeth, apical tooth slightly recurved and larger than the other two; apical spur present; dorsal longitudinal ridge present and with a row of 8 brown setae (
<figureCitation id="7BE02A30FFE6FF9DF591F9ABFC6BF9BF" box="[943,1010,1624,1643]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="6" pageNumber="214">Fig. 44</figureCitation>
). Metacoxa with posterior projection extending beyond base of metatrochanter; anteroexternal angle strongly acute (
<figureCitation id="7BE02A30FFE6FF9DF559F963FC31F977" box="[871,936,1680,1699]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="6" pageNumber="214">Fig. 40</figureCitation>
). Metafemur flattened medially. Mesotibia with moderate setae on inner surface, spines and sparse spine-like setae on outer surface; apical margin dark, inner margin with two larger spurs interleaved with two small spine-like setae, outer margin with 10 spine-like setae. Metatibia with apical margin dark, inner margin with two larger spines interleaved with two smaller spines, outer margin with one small spine-like setae; four spine-like setae on external longitudinal carinae (
<figureCitation id="7BE02A30FFE6FF9DF240F8A0FB5AF8B2" box="[1150,1219,1875,1894]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="6" pageNumber="214">Fig. 56</figureCitation>
). Tarsomeres with small ventral tooth, most evident on meso- and metatarsomeres V. Tarsomeres 24, with two inner spine-like setae on all legs. Tarsal claws simple, inner protarsal claw slightly recurved and larger than outer claw (
<figureCitation id="7BE02A30FFE7FF9CF6CEFAC7FEADFA93" box="[240,308,1332,1351]" captionStart="Figs" captionStartId="10.[85,120,1787,1801]" captionTargetBox="[197,1362,153,1753]" captionTargetId="figure-13@10.[197,1362,150,1756]" captionTargetPageId="10" captionText="Figs. 4752. Anterior tarsal claws of male and left metatibia of female in lateral view.4750, anterior tarsal claws of male;47,Pelidnota beckeri sp.nov.; 48,P.unicolor (Drury); 49, P. nordestina sp. nov.; 50, P. pernambucana sp. nov.; 51 and 52, metatibia of female; 51, P. beckeri sp. nov., a spine-like seta; 52, P. unicolor (Drury), a spine (cuticular projection). Scale bars: Figs.4750 = 0.500 mm, Figs.51 and 52 = 1.0mm." pageId="7" pageNumber="215">Fig. 50</figureCitation>
). Outer claws of meso- and metatarsus larger than inner claws.
<emphasis id="D1AFEAA7FFE7FF9CF71AFABCFE19FAB7" box="[292,384,1359,1379]" italics="true" pageId="7" pageNumber="215">Abdomen</emphasis>
: Abdominal ventrite convex, punctures moderate to dense, and with moderate setae. Sternite VI ending before posterior margin of pygidium. Margins of abdominal ventrites darker than the rest of the ventrites.
<emphasis id="D1AFEAA7FFE7FF9CF475FA51FD3EFA62" box="[587,679,1442,1462]" italics="true" pageId="7" pageNumber="215">Pygidium</emphasis>
: (
<figureCitation id="7BE02A30FFE7FF9CF4FFFA50FC90FA62" box="[705,777,1443,1462]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="7" pageNumber="215">Fig. 46</figureCitation>
) convex in lateral view. Surface densely striate and with moderately to dense brown setae. Posterior margin slightly concave.
<emphasis id="D1AFEAA7FFE7FF9CF49DFA29FC90FA3A" box="[675,777,1498,1518]" italics="true" pageId="7" pageNumber="215">Parameres</emphasis>
: (
<figureCitation id="7BE02A30FFE7FF9CF644FA04FE8AF9DE" box="[122,275,1527,1546]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="7" pageNumber="215">Figs. 63, 64, 68</figureCitation>
) symmetrical, fused; apex concave, distal margin slightly rounded laterally; proximal margin straight and truncated.
</paragraph>
<caption id="B7A4663DFFE5FF9EF64CF936FA42F93E" pageId="5" pageNumber="213" startId="5.[114,149,1733,1747]" targetBox="[209,1398,152,1698]" targetPageId="5" targetType="figure">
<paragraph id="E36436B5FFE5FF9EF64CF936FA42F93E" blockId="5.[114,1503,1733,1770]" pageId="5" pageNumber="213">
<emphasis id="D1AFEAA7FFE5FF9EF64CF936FF4FF907" bold="true" box="[114,214,1733,1747]" pageId="5" pageNumber="213">Figs. 1724.</emphasis>
Mouthparts. 1720, labrum in frontal view; 17,
<taxonomicName id="24DB4D36FFE5FF9EF459F936FD2BF900" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[615,690,1733,1748]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="213" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFE5FF9EF459F936FD2BF900" box="[615,690,1733,1748]" italics="true" pageId="5" pageNumber="213">P. beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE5FF9EF489F935FD63F900" box="[695,762,1734,1748]" pageId="5" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
; 18,
<taxonomicName id="24DB4D36FFE5FF9EF521F936FC2EF907" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[799,951,1733,1748]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="213" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFE5FF9EF521F936FCEBF900" box="[799,882,1733,1748]" italics="true" pageId="5" pageNumber="213">P. unicolor</emphasis>
(Drury)
</taxonomicName>
; 19,
<taxonomicName id="24DB4D36FFE5FF9EF5E3F936FBDAF900" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[989,1091,1733,1748]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="213" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE5FF9EF5E3F936FBDAF900" box="[989,1091,1733,1748]" italics="true" pageId="5" pageNumber="213">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE5FF9EF277F935FB12F900" box="[1097,1163,1734,1748]" pageId="5" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
; 20,
<taxonomicName id="24DB4D36FFE5FF9EF28EF936FAA1F900" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1200,1336,1733,1748]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="213" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE5FF9EF28EF936FAA1F900" box="[1200,1336,1733,1748]" italics="true" pageId="5" pageNumber="213">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE5FF9EF300F935FA19F900" box="[1342,1408,1734,1748]" pageId="5" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
; 2124, left mandible in dorsolateral view; 21,
<taxonomicName id="24DB4D36FFE5FF9EF7B3F928FD8BF93E" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[397,530,1755,1770]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="213" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFE5FF9EF7B3F928FD8BF93E" box="[397,530,1755,1770]" italics="true" pageId="5" pageNumber="213">Pelidnota beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE5FF9EF429F92FFDC1F93E" box="[535,600,1756,1770]" pageId="5" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
; 22,
<taxonomicName id="24DB4D36FFE5FF9EF442F928FC8BF93E" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[636,786,1755,1770]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="213" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFE5FF9EF442F928FD57F93E" box="[636,718,1755,1770]" italics="true" pageId="5" pageNumber="213">P. unicolor</emphasis>
(Drury)
</taxonomicName>
; 23,
<taxonomicName id="24DB4D36FFE5FF9EF509F928FC05F93E" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[823,924,1755,1770]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="213" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE5FF9EF509F928FC05F93E" box="[823,924,1755,1770]" italics="true" pageId="5" pageNumber="213">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE5FF9EF59EF92FFC7BF93E" box="[928,994,1756,1770]" pageId="5" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
; 24,
<taxonomicName id="24DB4D36FFE5FF9EF23BF928FB15F93E" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1029,1164,1755,1770]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="213" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE5FF9EF23BF928FB15F93E" box="[1029,1164,1755,1770]" italics="true" pageId="5" pageNumber="213">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE5FF9EF2AFF92FFB49F93E" box="[1169,1232,1756,1770]" pageId="5" pageNumber="213" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: Figs. 1724 = 2.0 mm.
</paragraph>
</caption>
<caption id="B7A4663DFFE6FF9DF66BFA46FDECFA0E" pageId="6" pageNumber="214" startId="6.[85,120,1461,1475]" targetBox="[246,1311,154,1428]" targetPageId="6" targetType="figure">
<paragraph id="E36436B5FFE6FF9DF66BFA46FDECFA0E" blockId="6.[85,1476,1461,1498]" pageId="6" pageNumber="214">
<emphasis id="D1AFEAA7FFE6FF9DF66BFA46FF21FA17" bold="true" box="[85,184,1461,1475]" pageId="6" pageNumber="214">Figs. 2528.</emphasis>
Mouthparts. 25 and 26, left maxilla in dorsal view; 25,
<taxonomicName id="24DB4D36FFE6FF9DF440FA46FD51FA10" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[638,712,1461,1476]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="214" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFE6FF9DF440FA46FD51FA10" box="[638,712,1461,1476]" italics="true" pageId="6" pageNumber="214">P. beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE6FF9DF4F2FA45FC97FA10" box="[716,782,1462,1476]" pageId="6" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
; 26,
<taxonomicName id="24DB4D36FFE6FF9DF50CFA46FC51FA17" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[818,968,1461,1476]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="214" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFE6FF9DF50CFA46FC1DFA10" box="[818,900,1461,1476]" italics="true" pageId="6" pageNumber="214">P. unicolor</emphasis>
(Drury)
</taxonomicName>
; 2728, left maxilla in dorsal view; 27,
<taxonomicName id="24DB4D36FFE6FF9DF339FA46FA3EFA10" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1287,1447,1461,1476]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="214" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE6FF9DF339FA46FA3EFA10" box="[1287,1447,1461,1476]" italics="true" pageId="6" pageNumber="214">Pelidnota nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE6FF9DF392FA45FFE5FA0E" pageId="6" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
; 28,
<taxonomicName id="24DB4D36FFE6FF9DF6A1FA38FEBFFA0E" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[159,294,1483,1498]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="214" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE6FF9DF6A1FA38FEBFFA0E" box="[159,294,1483,1498]" italics="true" pageId="6" pageNumber="214">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE6FF9DF715FA3FFEF3FA0E" box="[299,362,1484,1498]" pageId="6" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: Figs. 2528 = 2.0 mm.
</paragraph>
</caption>
<caption id="B7A4663DFFE7FF9CF64CFB3BFF6FFB39" pageId="7" pageNumber="215" startId="7.[114,149,1224,1238]" targetBox="[224,1373,161,1190]" targetPageId="7" targetType="figure">
<paragraph id="E36436B5FFE7FF9CF64CFB3BFF6FFB39" blockId="7.[114,1505,1223,1261]" pageId="7" pageNumber="215">
<emphasis id="D1AFEAA7FFE7FF9CF64CFB3BFF4CFB02" bold="true" box="[114,213,1224,1238]" pageId="7" pageNumber="215">Figs. 2932.</emphasis>
Mouthparts. 2932, labium in dorsal view; 29,
<taxonomicName id="24DB4D36FFE7FF9CF466FB34FD3BFB02" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[600,674,1223,1238]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFE7FF9CF466FB34FD3BFB02" box="[600,674,1223,1238]" italics="true" pageId="7" pageNumber="215">P. beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE7FF9CF498FB3BFD71FB02" box="[678,744,1224,1238]" pageId="7" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
; 30,
<taxonomicName id="24DB4D36FFE7FF9CF535FB34FC38FB02" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[779,929,1223,1238]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFE7FF9CF535FB34FCC4FB02" box="[779,861,1223,1238]" italics="true" pageId="7" pageNumber="215">P. unicolor</emphasis>
(Drury)
</taxonomicName>
; 31,
<taxonomicName id="24DB4D36FFE7FF9CF5FBFB34FBB3FB02" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[965,1066,1223,1238]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE7FF9CF5FBFB34FBB3FB02" box="[965,1066,1223,1238]" italics="true" pageId="7" pageNumber="215">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE7FF9CF211FB3BFBE9FB02" box="[1071,1136,1224,1238]" pageId="7" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
; 32,
<taxonomicName id="24DB4D36FFE7FF9CF2AAFB34FA83FB02" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1172,1306,1223,1238]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE7FF9CF2AAFB34FA83FB02" box="[1172,1306,1223,1238]" italics="true" pageId="7" pageNumber="215">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE7FF9CF321FB3BFAC7FB02" box="[1311,1374,1224,1238]" pageId="7" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: Figs. 2932 = 2.0 mm.
</paragraph>
</caption>
<paragraph id="E36436B5FFE7FF9CF6ACF9DDFC88F9AE" blockId="7.[114,786,1332,1965]" pageId="7" pageNumber="215">
<emphasis id="D1AFEAA7FFE7FF9CF6ACF9DDFEBBF996" box="[146,290,1582,1602]" italics="true" pageId="7" pageNumber="215">Type material.</emphasis>
<materialsCitation id="53B33CE8FFE7FF9CF715F9DCFC88F9AE" collectingDate="2015-03-25" collectionCode="MZFS" collectorName="E. M. Menezes" country="Brazil" elevation="510" location="Cachoeira Veu da Noiva" municipality="Bonito" pageId="7" pageNumber="215" specimenCode="MZFS#55727" specimenCount="1" stateProvince="Pernambuco" typeStatus="holotype">
<typeStatus id="3C608817FFE7FF9CF715F9DCFE11F996" box="[299,392,1583,1602]" pageId="7" pageNumber="215" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="9BCC7625FFE7FF9CF7EBF9DCFD86F996" box="[469,543,1583,1602]" name="Brazil" pageId="7" pageNumber="215">BRAZIL</collectingCountry>
,
<emphasis id="D1AFEAA7FFE7FF9CF412F9DDFD3CF996" box="[556,677,1582,1602]" italics="true" pageId="7" pageNumber="215">
<collectingRegion id="211FF857FFE7FF9CF412F9DDFD3CF996" box="[556,677,1582,1602]" country="Brazil" name="Pernambuco" pageId="7" pageNumber="215">Pernambuco</collectingRegion>
</emphasis>
: (“
<collectingMunicipality id="0300ACCFFFE7FF9CF4F9F9DCFC94F996" box="[711,781,1583,1602]" pageId="7" pageNumber="215">Bonito</collectingMunicipality>
,
<location id="E604606EFFE7FF9CF64CF9B8FEEAF98A" LSID="urn:lsid:plazi:treatment:6B7287A3FFE4FF96F64BF8A4FA79F814:E604606EFFE7FF9CF64CF9B8FEEAF98A" box="[114,371,1611,1630]" country="Brazil" municipality="Bonito" name="Cachoeira Veu da Noiva" pageId="7" pageNumber="215" stateProvince="Pernambuco">Cachoeira Véu da Noiva</location>
”, “
<quantity id="24239B50FFE7FF9CF7ADF9B8FE4AF98A" box="[403,467,1611,1630]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="215" unit="m" value="510.0">
<elevation id="68F6D186FFE7FF9CF7ADF9B8FE4AF98A" box="[403,467,1611,1630]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="215" unit="m" value="510.0">510 m</elevation>
</quantity>
”) (8
<emphasis id="D1AFEAA7FFE7FF9CF43DF9B4FD92F981" bold="true" box="[515,523,1607,1621]" pageId="7" pageNumber="215">
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32
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</emphasis>
32.4
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<superScript id="14AE9BFDFFE7FF9CF469F9B4FDF9F981" attach="right" box="[599,608,1607,1621]" fontSize="6" pageId="7" pageNumber="215">ll</superScript>
</emphasis>
S/35
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42
<emphasis id="D1AFEAA7FFE7FF9CF48FF9B4FD2CF981" bold="true" box="[689,693,1607,1621]" pageId="7" pageNumber="215">
<superScript id="14AE9BFDFFE7FF9CF48FF9B4FD2CF981" attach="left" box="[689,693,1607,1621]" fontSize="6" pageId="7" pageNumber="215">l</superScript>
</emphasis>
53.9
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</emphasis>
W),
<date id="97651075FFE7FF9CF64CF995FF46F9AD" box="[114,223,1638,1657]" pageId="7" pageNumber="215" value="2015-03-25">
<collectingDate id="8721E99DFFE7FF9CF64CF995FF46F9AD" box="[114,223,1638,1657]" pageId="7" pageNumber="215" value="2015-03-25">25.III.2015</collectingDate>
</date>
.
<collectorName id="4E2E5363FFE7FF9CF6D6F994FEEBF9AE" box="[232,370,1639,1658]" pageId="7" pageNumber="215">E.M. Menezes</collectorName>
col. (
<specimenCode id="B37D9ECEFFE7FF9CF798F995FDA8F9AD" box="[422,561,1638,1657]" collectionCode="MZFS" pageId="7" pageNumber="215">MZFS#55727</specimenCode>
) (
<collectionCode id="85CAAE70FFE7FF9CF47CF994FDE7F9AE" box="[578,638,1639,1658]" pageId="7" pageNumber="215">MZFS</collectionCode>
) (
<figureCitation id="7BE02A30FFE7FF9CF4AFF994FC9AF9AD" box="[657,771,1638,1658]" captionStart="Figs" captionStartId="13.[114,149,1725,1739]" captionTargetBox="[179,720,149,1695]" captionTargetId="figure-13@13.[179,720,149,1695]" captionTargetPageId="13" captionText="Figs.6977. Labels attachedtothe hotypes.6971,Pelidnota beckerisp.nov.;7274, P. nordestina sp. nov.; 7577, P.pernambucana sp. nov." pageId="7" pageNumber="215">Figs. 7577</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="E36436B5FFE7FF9CF6ACF972FDD5F941" blockId="7.[114,786,1332,1965]" box="[146,588,1665,1686]" pageId="7" pageNumber="215">
<emphasis id="D1AFEAA7FFE7FF9CF6ACF972FE94F941" box="[146,269,1665,1685]" italics="true" pageId="7" pageNumber="215">Distribution.</emphasis>
Known only from
<typeStatus id="3C608817FFE7FF9CF7F7F970FE6FF942" box="[457,502,1667,1686]" pageId="7" pageNumber="215">type</typeStatus>
locality.
</paragraph>
<paragraph id="E36436B5FFE7FF9CF6ACF96EFD12F919" blockId="7.[114,786,1332,1965]" pageId="7" pageNumber="215">
<emphasis id="D1AFEAA7FFE7FF9CF6ACF96EFF66F965" box="[146,255,1693,1713]" italics="true" pageId="7" pageNumber="215">Etymology.</emphasis>
The specific epithet
<taxonomicName id="24DB4D36FFE7FF9CF436F96EFD03F965" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[520,666,1693,1713]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="pernambucana">
<emphasis id="D1AFEAA7FFE7FF9CF436F96EFD03F965" box="[520,666,1693,1713]" italics="true" pageId="7" pageNumber="215">pernambucana</emphasis>
</taxonomicName>
refers to
<collectingRegion id="211FF857FFE7FF9CF64CF949FEB1F91A" box="[114,296,1722,1742]" country="Brazil" name="Pernambuco" pageId="7" pageNumber="215">Pernambuco state</collectingRegion>
where the
<typeStatus id="3C608817FFE7FF9CF7A5F949FE6CF919" box="[411,501,1722,1741]" pageId="7" pageNumber="215" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
<paragraph id="E36436B5FFE7FF9CF6ACF926FADBF8E9" blockId="7.[114,786,1332,1965]" lastBlockId="7.[833,1504,1332,1965]" pageId="7" pageNumber="215">
<emphasis id="D1AFEAA7FFE7FF9CF6ACF926FE74F93D" box="[146,493,1749,1769]" italics="true" pageId="7" pageNumber="215">
Remarks.
<taxonomicName id="24DB4D36FFE7FF9CF6CBF926FE74F93D" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[245,493,1749,1769]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="pernambucana">Pelidnota pernambucana</taxonomicName>
</emphasis>
is distinguished as follows (
<taxonomicName id="24DB4D36FFE7FF9CF644F902FEDCF8D1" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[122,325,1777,1797]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="nordestina">
<emphasis id="D1AFEAA7FFE7FF9CF644F902FEDCF8D1" box="[122,325,1777,1797]" italics="true" pageId="7" pageNumber="215">Pelidnota nordestina</emphasis>
</taxonomicName>
characters in brackets): posterior margin of head less black (black dark); mandibles small, not exceeding the lateral margin of the clypeus (exceeding the lateral margin of the clypeus; labrum transversal (
<figureCitation id="7BE02A30FFE7FF9CF79CF8B5FE71F88D" box="[418,488,1862,1881]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="7" pageNumber="215">Fig. 20</figureCitation>
) (triangular (
<figureCitation id="7BE02A30FFE7FF9CF44BF8B5FD22F88D" box="[629,699,1862,1881]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="7" pageNumber="215">Fig. 19</figureCitation>
)); inner teeth of mandible larger (
<figureCitation id="7BE02A30FFE7FF9CF7BCF891FE5EF8A1" box="[386,455,1890,1909]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="7" pageNumber="215">Fig. 24</figureCitation>
) (small (
<figureCitation id="7BE02A30FFE7FF9CF416F891FDF4F8A1" box="[552,621,1890,1909]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="7" pageNumber="215">Fig. 23</figureCitation>
)); first teeth of maxillary galea sawed, IV palpomere of maxilla, more elongated and insertion area of palp more excavated (
<figureCitation id="7BE02A30FFE7FF9CF420F86AFDC6F879" box="[542,607,1945,1965]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="7" pageNumber="215">Fig.28</figureCitation>
) (first and second teeth sawed and IV palpomere less elongated (
<figureCitation id="7BE02A30FFE7FF9CF335FAC7FAD5FA93" box="[1291,1356,1332,1351]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="7" pageNumber="215">Fig.27</figureCitation>
)); surface area of pre-mentum and insertion area of palp more excavated, lateral margin of post-mentum less rounded, and with elongated moderate setae (
<figureCitation id="7BE02A30FFE7FF9CF59BFA74FC7CFA4F" box="[933,997,1415,1435]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="7" pageNumber="215">Fig.32</figureCitation>
) (less excavated, more rounded, and with elongate dense setae (
<figureCitation id="7BE02A30FFE7FF9CF5FBFA50FB91FA62" box="[965,1032,1443,1462]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="7" pageNumber="215">Fig. 31</figureCitation>
)); external teeth of protibia small and slightly curved (
<figureCitation id="7BE02A30FFE7FF9CF5A8FA4CFC43FA06" box="[918,986,1471,1490]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="7" pageNumber="215">Fig. 44</figureCitation>
) (larger and sharply curved (
<figureCitation id="7BE02A30FFE7FF9CF337FA4CFAD7FA06" box="[1289,1358,1471,1490]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="7" pageNumber="215">Fig. 43</figureCitation>
)); elytra apex truncated (
<figureCitation id="7BE02A30FFE7FF9CF58DFA28FC61FA3A" box="[947,1016,1499,1518]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="7" pageNumber="215">Fig. 36</figureCitation>
) (slightly more acute (
<figureCitation id="7BE02A30FFE7FF9CF2DBFA28FAB3FA3A" box="[1253,1322,1499,1518]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="7" pageNumber="215">Fig. 35</figureCitation>
)); anteroexternal angle of metacoxa strongly acute (
<figureCitation id="7BE02A30FFE7FF9CF294FA04FB76F9DE" box="[1194,1263,1527,1546]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="7" pageNumber="215">Fig. 40</figureCitation>
) (rounded and slightly sharp (
<figureCitation id="7BE02A30FFE7FF9CF5B6F9E0FC55F9F2" box="[904,972,1555,1574]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="7" pageNumber="215">Fig. 39</figureCitation>
)); pygidium dark brown with strong metallic green reflections around pygidium (
<figureCitation id="7BE02A30FFE7FF9CF247F9DCFB26F996" box="[1145,1215,1583,1602]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="7" pageNumber="215">Fig. 46</figureCitation>
) (dark brown with reddish reflections around pygidium, and slight metallic green reflections only on the sides of the posterior margin of pygidium (
<figureCitation id="7BE02A30FFE7FF9CF3BDF994FA51F9AD" box="[1411,1480,1638,1658]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="7" pageNumber="215">Fig. 45</figureCitation>
)); small apical spurs and cuticle projection-shaped spur of metatibia; six spine-like setae on external longitudinal carinae, and six setae on inner longitudinal carinae of metatibia (
<figureCitation id="7BE02A30FFE7FF9CF301F949FA1DF919" box="[1343,1412,1722,1741]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="7" pageNumber="215">Fig. 56</figureCitation>
) (larger; more than six (
<figureCitation id="7BE02A30FFE7FF9CF5E2F925FB86F93D" box="[988,1055,1750,1769]" captionStart="Figs" captionStartId="11.[114,149,1821,1835]" captionTargetBox="[281,1346,152,1789]" captionTargetId="figure-13@11.[267,1349,149,1790]" captionTargetPageId="11" captionText="Figs. 5356. Right metatibia of male in lateral view. 53, Pelidnota beckeri sp. nov.; 54, P. unicolor (Drury); 55, P. nordestina sp. nov.; 56, P. pernambucana sp. nov. Scale bars: Figs.5356= 2.0mm." pageId="7" pageNumber="215">Fig. 55</figureCitation>
); parameres distal margin slightly rounded, and proximal margin straight and truncated (
<figureCitation id="7BE02A30FFE7FF9CF330F901FA3AF8D1" box="[1294,1443,1778,1797]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="7" pageNumber="215">Figs. 63, 64, 68</figureCitation>
) (distal margin V-shaped and gradually acute laterally, and proximal margin gradually semi-quadrate (
<figureCitation id="7BE02A30FFE7FF9CF2A9F8D9FAB5F8E9" box="[1175,1324,1834,1853]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="7" pageNumber="215">Figs. 61, 62, 67</figureCitation>
)).
</paragraph>
<paragraph id="E36436B5FFE7FF9CF55EF8B6FAEDF8A1" blockId="7.[833,1504,1332,1965]" pageId="7" pageNumber="215">
<emphasis id="D1AFEAA7FFE7FF9CF55EF8B6FC57F88D" box="[864,974,1861,1881]" italics="true" pageId="7" pageNumber="215">Comments.</emphasis>
The specimen was collected in a tropical forest in an enclave in the Caatinga ecoregion (
<bibRefCitation id="874A4B44FFE7FF9CF29DF891FAFEF8A1" author="Instituto LIFE" box="[1187,1383,1890,1909]" pageId="7" pageNumber="215" pagination="35 - 52" refId="ref10592" refString="Instituto LIFE, 2015. Bahia, in: Ecorregioes do Brasil - Prioridades Terrestres e Marinhas. Serie cadernos tecnicos, Volume 3, Brasil., pp. 35 - 52." type="book chapter" year="2015">Instituto LIFE, 2015</bibRefCitation>
).
</paragraph>
<paragraph id="E36436B5FFE7FF96F55EF88FFA79FF13" blockId="7.[833,1504,1332,1965]" lastBlockId="13.[833,1504,152,311]" lastPageId="13" lastPageNumber="221" pageId="7" pageNumber="215">
<emphasis id="D1AFEAA7FFE7FF9CF55EF88FFB22F844" box="[864,1211,1916,1936]" italics="true" pageId="7" pageNumber="215">
Identification key.
<taxonomicName id="24DB4D36FFE7FF9CF22AF88FFB22F844" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1044,1211,1916,1936]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="beckeri">Pelidnota beckeri</taxonomicName>
</emphasis>
,
<taxonomicName id="24DB4D36FFE7FF9CF2FBF88FFADDF844" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1221,1348,1916,1936]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="nordestina">
<emphasis id="D1AFEAA7FFE7FF9CF2FBF88FFADDF844" box="[1221,1348,1916,1936]" italics="true" pageId="7" pageNumber="215">P. nordestina</emphasis>
</taxonomicName>
and
<taxonomicName id="24DB4D36FFE7FF9CF34BF88FFC11F878" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="215" phylum="Arthropoda" rank="species" species="pernambucana">
<emphasis id="D1AFEAA7FFE7FF9CF34BF88FFC11F878" italics="true" pageId="7" pageNumber="215">P. pernambucana</emphasis>
</taxonomicName>
were included in couplet 48 of the identification key for the uniformly brown and robust; pronotum densely covered by ventral bristles.
<collectingCountry id="9BCC7625FFEDFF96F5EEFF47FB8EFF13" box="[976,1047,180,199]" name="Bolivia" pageId="13" pageNumber="221">Bolivia</collectingCountry>
,
<collectingCountry id="9BCC7625FFEDFF96F21EFF47FB1FFF13" box="[1056,1158,180,199]" name="Argentina" pageId="13" pageNumber="221">Argentina</collectingCountry>
, and southern
<collectingCountry id="9BCC7625FFEDFF96F325FF47FACCFF13" box="[1307,1365,180,199]" name="Brazil" pageId="13" pageNumber="221">Brazil</collectingCountry>
.. ... ...... 45
</paragraph>
<caption id="B7A4663DFFE8FF93F66BF8BEFC32F85D" pageId="8" pageNumber="216" startId="8.[85,120,1869,1883]" targetBox="[271,1287,149,1839]" targetPageId="8" targetType="figure">
<paragraph id="E36436B5FFE8FF93F66BF8BEFC32F85D" blockId="8.[85,1476,1869,1929]" pageId="8" pageNumber="216">
<emphasis id="D1AFEAA7FFE8FF93F66BF8BEFF21F88F" bold="true" box="[85,184,1869,1883]" pageId="8" pageNumber="216">Figs. 3340.</emphasis>
Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33,
<taxonomicName id="24DB4D36FFE8FF93F241F8BEFA9CF888" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1151,1285,1869,1884]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="216" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFE8FF93F241F8BEFA9CF888" box="[1151,1285,1869,1884]" italics="true" pageId="8" pageNumber="216">Pelidnota beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE8FF93F337F8BDFAD2F888" box="[1289,1355,1870,1884]" pageId="8" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
; 34,
<taxonomicName id="24DB4D36FFE8FF93F34EF8BEFF0CF8A6" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="216" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFE8FF93F34EF8BEFA5BF888" box="[1392,1474,1869,1884]" italics="true" pageId="8" pageNumber="216">P. unicolor</emphasis>
(Drury)
</taxonomicName>
; 35,
<taxonomicName id="24DB4D36FFE8FF93F685F890FEB8F8A6" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[187,289,1891,1906]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="216" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE8FF93F685F890FEB8F8A6" box="[187,289,1891,1906]" italics="true" pageId="8" pageNumber="216">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE8FF93F719F897FEF3F8A6" box="[295,362,1892,1906]" pageId="8" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
; 36,
<taxonomicName id="24DB4D36FFE8FF93F7B1F890FD8EF8A6" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[399,535,1891,1906]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="216" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE8FF93F7B1F890FD8EF8A6" box="[399,535,1891,1906]" italics="true" pageId="8" pageNumber="216">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE8FF93F423F897FDC6F8A6" box="[541,607,1892,1906]" pageId="8" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
; 3740, anteroexternal coxal angle of right metacoxa; 37,
<taxonomicName id="24DB4D36FFE8FF93F200F890FB10F8A6" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1086,1161,1891,1906]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="216" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFE8FF93F200F890FB10F8A6" box="[1086,1161,1891,1906]" italics="true" pageId="8" pageNumber="216">P. beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE8FF93F2B0F897FB48F8A6" box="[1166,1233,1892,1906]" pageId="8" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
; 38,
<taxonomicName id="24DB4D36FFE8FF93F2C8F890FA16F8A6" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[1270,1423,1891,1906]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="216" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFE8FF93F2C8F890FAD0F8A6" box="[1270,1353,1891,1906]" italics="true" pageId="8" pageNumber="216">P. unicolor</emphasis>
(Drury)
</taxonomicName>
; 39,
<taxonomicName id="24DB4D36FFE8FF93F38BF890FF31F85D" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="216" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE8FF93F38BF890FF31F85D" italics="true" pageId="8" pageNumber="216">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE8FF93F693F888FF77F85D" box="[173,238,1915,1929]" pageId="8" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
; 40,
<taxonomicName id="24DB4D36FFE8FF93F72CF889FE00F85D" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[274,409,1914,1929]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="216" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE8FF93F72CF889FE00F85D" box="[274,409,1914,1929]" italics="true" pageId="8" pageNumber="216">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE8FF93F7A3F888FE45F85D" box="[413,476,1915,1929]" pageId="8" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: Figs. 3336 = 0.500 mm, Figs. 3740 = 1.0 mm.
</paragraph>
</caption>
<caption id="B7A4663DFFE9FF92F64CF94EFB26F936" pageId="9" pageNumber="217" startId="9.[114,149,1725,1739]" targetBox="[348,1282,149,1690]" targetPageId="9" targetType="figure">
<paragraph id="E36436B5FFE9FF92F64CF94EFB26F936" blockId="9.[114,1503,1725,1762]" pageId="9" pageNumber="217">
<emphasis id="D1AFEAA7FFE9FF92F64CF94EFF4CF91F" bold="true" box="[114,213,1725,1739]" pageId="9" pageNumber="217">Figs. 4146.</emphasis>
Protibia and pygidium in dorsal view. 4144, left protíbia; 41,
<taxonomicName id="24DB4D36FFE9FF92F4E4F94EFCF9F918" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[730,864,1725,1740]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="217" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFE9FF92F4E4F94EFCF9F918" box="[730,864,1725,1740]" italics="true" pageId="9" pageNumber="217">Pelidnota beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE9FF92F55AF94DFC3FF918" box="[868,934,1726,1740]" pageId="9" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
; 42,
<taxonomicName id="24DB4D36FFE9FF92F5F5F94EFBFBF91F" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[971,1122,1725,1740]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="217" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFE9FF92F5F5F94EFB84F918" box="[971,1053,1725,1740]" italics="true" pageId="9" pageNumber="217">P. unicolor</emphasis>
(Drury)
</taxonomicName>
; 43,
<taxonomicName id="24DB4D36FFE9FF92F2B9F94EFB75F918" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1159,1260,1725,1740]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="217" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE9FF92F2B9F94EFB75F918" box="[1159,1260,1725,1740]" italics="true" pageId="9" pageNumber="217">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE9FF92F2CFF94DFAADF918" box="[1265,1332,1726,1740]" pageId="9" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
; 44,
<taxonomicName id="24DB4D36FFE9FF92F366F94EFA46F918" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1368,1503,1725,1740]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="217" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE9FF92F366F94EFA46F918" box="[1368,1503,1725,1740]" italics="true" pageId="9" pageNumber="217">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE9FF92F64CF927FF2DF936" box="[114,180,1748,1762]" pageId="9" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
; 4546, pygidium; 45,
<taxonomicName id="24DB4D36FFE9FF92F752F920FE48F936" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[364,465,1747,1762]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="217" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFE9FF92F752F920FE48F936" box="[364,465,1747,1762]" italics="true" pageId="9" pageNumber="217">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE9FF92F7EBF927FD8EF936" box="[469,535,1748,1762]" pageId="9" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
; 46,
<taxonomicName id="24DB4D36FFE9FF92F404F920FD58F936" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[570,705,1747,1762]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="217" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFE9FF92F404F920FD58F936" box="[570,705,1747,1762]" italics="true" pageId="9" pageNumber="217">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFE9FF92F4F8F927FC9CF936" box="[710,773,1748,1762]" pageId="9" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: Figs. 4142 = 1.0 mm, Figs. 4576 = 2.7 mm.
</paragraph>
</caption>
<caption id="B7A4663DFFEAFF91F66BF908FD38F8E3" pageId="10" pageNumber="218" startId="10.[85,120,1787,1801]" targetBox="[197,1362,153,1753]" targetPageId="10" targetType="figure">
<paragraph id="E36436B5FFEAFF91F66BF908FD38F8E3" blockId="10.[85,1475,1786,1847]" pageId="10" pageNumber="218">
<emphasis id="D1AFEAA7FFEAFF91F66BF908FF2EF8DD" bold="true" box="[85,183,1787,1801]" pageId="10" pageNumber="218">Figs. 4752.</emphasis>
Anterior tarsal claws of male and left metatibia of female in lateral view. 4750, anterior tarsal claws of male; 47,
<taxonomicName id="24DB4D36FFEAFF91F27CF909FB5EF8DD" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1090,1223,1786,1801]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="218" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFEAFF91F27CF909FB5EF8DD" box="[1090,1223,1786,1801]" italics="true" pageId="10" pageNumber="218">Pelidnota beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEAFF91F2F4F908FA93F8DD" box="[1226,1290,1787,1801]" pageId="10" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
; 48,
<taxonomicName id="24DB4D36FFEAFF91F315F909FA59F8DD" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[1323,1472,1786,1801]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="218" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFEAFF91F315F909FAE5F8DD" box="[1323,1404,1786,1801]" italics="true" pageId="10" pageNumber="218">P.unicolor</emphasis>
(Drury)
</taxonomicName>
; 49,
<taxonomicName id="24DB4D36FFEAFF91F64DF8E2FF43F8F4" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[115,218,1809,1824]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="218" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFEAFF91F64DF8E2FF43F8F4" box="[115,218,1809,1824]" italics="true" pageId="10" pageNumber="218">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEAFF91F6DEF8E1FEBAF8F4" box="[224,291,1810,1824]" pageId="10" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
; 50,
<taxonomicName id="24DB4D36FFEAFF91F777F8E2FE48F8F4" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[329,465,1809,1824]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="218" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFEAFF91F777F8E2FE48F8F4" box="[329,465,1809,1824]" italics="true" pageId="10" pageNumber="218">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEAFF91F7E9F8E1FD83F8F4" box="[471,538,1810,1824]" pageId="10" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
; 51 and 52, metatibia of female; 51,
<taxonomicName id="24DB4D36FFEAFF91F574F8E2FC0CF8F4" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[842,917,1809,1824]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="218" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFEAFF91F574F8E2FC0CF8F4" box="[842,917,1809,1824]" italics="true" pageId="10" pageNumber="218">P. beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEAFF91F5A5F8E1FC45F8F4" box="[923,988,1810,1824]" pageId="10" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
, a spine-like seta; 52,
<taxonomicName id="24DB4D36FFEAFF91F2AAF8E2FAB2F8F4" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[1172,1323,1809,1824]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="218" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFEAFF91F2AAF8E2FB7EF8F4" box="[1172,1255,1809,1824]" italics="true" pageId="10" pageNumber="218">P. unicolor</emphasis>
(Drury)
</taxonomicName>
, a spine (cuticular projection). Scale bars: Figs. 4750 = 0.500 mm, Figs. 51 and 52 = 1.0 mm.
</paragraph>
</caption>
<caption id="B7A4663DFFEBFF90F64CF8EEFE86F896" pageId="11" pageNumber="219" startId="11.[114,149,1821,1835]" targetBox="[281,1346,152,1789]" targetPageId="11" targetType="figure">
<paragraph id="E36436B5FFEBFF90F64CF8EEFE86F896" blockId="11.[114,1505,1821,1858]" pageId="11" pageNumber="219">
<emphasis id="D1AFEAA7FFEBFF90F64CF8EEFF4CF8FF" bold="true" box="[114,213,1821,1835]" pageId="11" pageNumber="219">Figs. 5356.</emphasis>
Right metatibia of male in lateral view. 53,
<taxonomicName id="24DB4D36FFEBFF90F403F8EEFD5AF8F8" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[573,707,1821,1836]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="219" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFEBFF90F403F8EEFD5AF8F8" box="[573,707,1821,1836]" italics="true" pageId="11" pageNumber="219">Pelidnota beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEBFF90F4F9F8EDFC90F8F8" box="[711,777,1822,1836]" pageId="11" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
; 54,
<taxonomicName id="24DB4D36FFEBFF90F513F8EEFC5CF8FF" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[813,965,1821,1836]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="219" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFEBFF90F513F8EEFC19F8F8" box="[813,896,1821,1836]" italics="true" pageId="11" pageNumber="219">P. unicolor</emphasis>
(Drury)
</taxonomicName>
; 55,
<taxonomicName id="24DB4D36FFEBFF90F5D7F8EEFBD6F8F8" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1001,1103,1821,1836]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="219" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFEBFF90F5D7F8EEFBD6F8F8" box="[1001,1103,1821,1836]" italics="true" pageId="11" pageNumber="219">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEBFF90F26AF8EDFB0FF8F8" box="[1108,1174,1822,1836]" pageId="11" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
; 56,
<taxonomicName id="24DB4D36FFEBFF90F284F8EEFAD8F8F8" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1210,1345,1821,1836]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="219" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFEBFF90F284F8EEFAD8F8F8" box="[1210,1345,1821,1836]" italics="true" pageId="11" pageNumber="219">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEBFF90F378F8EDFA1CF8F8" box="[1350,1413,1822,1836]" pageId="11" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: Figs. 5356 = 2.0 mm.
</paragraph>
</caption>
<caption id="B7A4663DFFECFF97F66BF90EFC3BF8ED" pageId="12" pageNumber="220" startId="12.[85,120,1789,1803]" targetBox="[219,1340,153,1758]" targetPageId="12" targetType="figure">
<paragraph id="E36436B5FFECFF97F66BF90EFC3BF8ED" blockId="12.[85,1476,1789,1849]" pageId="12" pageNumber="220">
<emphasis id="D1AFEAA7FFECFF97F66BF90EFF21F8DF" bold="true" box="[85,184,1789,1803]" pageId="12" pageNumber="220">Figs. 5768.</emphasis>
Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58,
<taxonomicName id="24DB4D36FFECFF97F578F90EFC55F8D8" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[838,972,1789,1804]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="220" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFECFF97F578F90EFC55F8D8" box="[838,972,1789,1804]" italics="true" pageId="12" pageNumber="220">Pelidnota beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFECFF97F5EEF90DFB97F8D8" box="[976,1038,1790,1804]" pageId="12" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
(dorsal, lateral); 59 and 60,
<taxonomicName id="24DB4D36FFECFF97F2D1F90EFA18F8DF" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[1263,1409,1789,1804]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="220" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFECFF97F2D1F90EFAD8F8D8" box="[1263,1345,1789,1804]" italics="true" pageId="12" pageNumber="220">P. unicolor</emphasis>
(Drury)
</taxonomicName>
(dorsal, lateral); 61 and 62,
<taxonomicName id="24DB4D36FFECFF97F6CCF8E0FECEF8F6" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[242,343,1811,1826]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="220" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFECFF97F6CCF8E0FECEF8F6" box="[242,343,1811,1826]" italics="true" pageId="12" pageNumber="220">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFECFF97F765F8E7FE03F8F6" box="[347,410,1812,1826]" pageId="12" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
(dorsal, lateral); 63 and 64,
<taxonomicName id="24DB4D36FFECFF97F444F8E0FC98F8F6" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[634,769,1811,1826]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="220" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFECFF97F444F8E0FC98F8F6" box="[634,769,1811,1826]" italics="true" pageId="12" pageNumber="220">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFECFF97F53BF8E7FCDDF8F6" box="[773,836,1812,1826]" pageId="12" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
(dorsal, lateral); 6568, paramers margin drawing; 65,
<taxonomicName id="24DB4D36FFECFF97F33DF8E0FAD4F8F6" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1283,1357,1811,1826]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="220" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFECFF97F33DF8E0FAD4F8F6" box="[1283,1357,1811,1826]" italics="true" pageId="12" pageNumber="220">P. beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFECFF97F36FF8E7FA0BF8F6" box="[1361,1426,1812,1826]" pageId="12" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
; 66,
<taxonomicName id="24DB4D36FFECFF97F38BF8E0FF43F8ED" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="220" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFECFF97F38BF8E0FF0CF8ED" italics="true" pageId="12" pageNumber="220">P. unicolor</emphasis>
(Drury)
</taxonomicName>
; 67,
<taxonomicName id="24DB4D36FFECFF97F6C0F8D9FEFAF8ED" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[254,355,1834,1849]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="220" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFECFF97F6C0F8D9FEFAF8ED" box="[254,355,1834,1849]" italics="true" pageId="12" pageNumber="220">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFECFF97F759F8D8FE30F8ED" box="[359,425,1835,1849]" pageId="12" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
; 68,
<taxonomicName id="24DB4D36FFECFF97F7F3F8D9FDCDF8ED" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[461,596,1834,1849]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="220" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFECFF97F7F3F8D9FDCDF8ED" box="[461,596,1834,1849]" italics="true" pageId="12" pageNumber="220">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFECFF97F466F8D8FD0EF8ED" box="[600,663,1835,1849]" pageId="12" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: Figs. 5764 = 2.5 mm.
</paragraph>
</caption>
<paragraph id="E36436B5FFEDFF96F57FFF23FA43FEE2" blockId="13.[833,1504,152,311]" pageId="13" pageNumber="221">
- Species of the Atlantic Forest and Caatinga ecoregions, Northeastern
<collectingCountry id="9BCC7625FFEDFF96F59BFF1FFC78FF2B" box="[933,993,236,255]" name="Brazil" pageId="13" pageNumber="221">Brazil</collectingCountry>
. Body uniformly brown or dark brown with dark spots present or not on pronotum, humeral angle and lateral elytra.. ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ......
<emphasis id="D1AFEAA7FFEDFF96F3FEFED0FA43FEE2" bold="true" box="[1472,1498,291,310]" pageId="13" pageNumber="221">48</emphasis>
</paragraph>
<paragraph id="E36436B5FFEDFF96F56CFE84FA43FE5E" blockId="13.[833,1504,375,506]" box="[850,1498,375,394]" pageId="13" pageNumber="221">48(43) Dorsal surface uniformly brown.. ... ... ... ... ... ... .... A</paragraph>
<paragraph id="E36436B5FFEDFF96F57FFE60FA46FE2E" blockId="13.[833,1504,375,506]" pageId="13" pageNumber="221">
-
<materialsCitation id="53B33CE8FFEDFF96F56CFE60FA46FE2E" location="Brazilian Atlantic Forest" pageId="13" pageNumber="221" specimenCount="1" stateProvince="Bahia, Espirito Santo, Minas Gerais, Rio de Janeiro">
Head, pronotum and scutellum dark brown; elytra brown. Distribution:
<location id="E604606EFFEDFF96F59CFE5CFB0CFE16" LSID="urn:lsid:plazi:treatment:6B7287A3FFE4FF96F64BF8A4FA79F814:E604606EFFEDFF96F59CFE5CFB0CFE16" box="[930,1173,431,450]" name="Brazilian Atlantic Forest" pageId="13" pageNumber="221" stateProvince="Bahia, Espirito Santo, Minas Gerais, Rio de Janeiro">Brazilian Atlantic Forest</location>
and deciduous forest (Caatinga), states of
<collectingRegion id="211FF857FFEDFF96F584FE38FA79FE0A" box="[954,1504,459,478]" isEnumeration="true" pageId="13" pageNumber="221">Bahia, Espírito Santo, Minas Gerais, Rio de Janeiro</collectingRegion>
.. ... ... ... ... ... ... ... ... ... ... ... ....
<taxonomicName id="24DB4D36FFEDFF96F2F8FE15FA46FE2E" authority="Perty" authorityName="Perty" authorityYear="1830" box="[1222,1503,486,506]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="221" phylum="Arthropoda" rank="species" species="chalcothorax">
<emphasis id="D1AFEAA7FFEDFF96F2F8FE15FA3AFE2E" box="[1222,1443,486,506]" italics="true" pageId="13" pageNumber="221">Pelidnota chalcothorax</emphasis>
Perty
</taxonomicName>
</materialsCitation>
</paragraph>
<paragraph id="E36436B5FFEDFF96F56CFDC9FA46FD9A" blockId="13.[833,1504,570,701]" box="[850,1503,570,590]" pageId="13" pageNumber="221">A(48) Black spots present on elytra or pronotum.. ... ... ... .... B</paragraph>
<paragraph id="E36436B5FFEDFF96F57FFDA4FA79FD69" blockId="13.[833,1504,570,701]" pageId="13" pageNumber="221">
- Black spots absent on elytra and pronotum absent (
<figureCitation id="7BE02A30FFEDFF96F355FDA5FA4AFDBD" box="[1387,1491,598,617]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="13" pageNumber="221">Figs. 9, 10</figureCitation>
), legs more or less brown, body darker than head, pronotum, and elytra. Distribution: Argentinian to
<collectingRegion id="211FF857FFEDFF96F31CFD7DFA45FD75" box="[1314,1500,654,673]" country="Brazil" name="Pernambuco" pageId="13" pageNumber="221">Pernambuco state</collectingRegion>
... ... ... ... ... ... ... ... ... ... ... ... ... ....
<taxonomicName id="24DB4D36FFEDFF96F2DEFD5AFA79FD69" authority="(Drury)" baseAuthorityName="Drury" baseAuthorityYear="1778" box="[1248,1504,681,701]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="221" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="D1AFEAA7FFEDFF96F2DEFD5AFA09FD69" box="[1248,1424,681,701]" italics="true" pageId="13" pageNumber="221">Pelidnota unicolor</emphasis>
(Drury)
</taxonomicName>
</paragraph>
<paragraph id="E36436B5FFEDFF96F56CFD0DFA40FCB0" blockId="13.[833,1504,765,1036]" pageId="13" pageNumber="221">B(A) Body brown; legs dark brown with metallic green reflections; one black lateral spot on pronotum and two on elytron (one at humerus of elytral and one on lateral half); elytral apex truncated, slightly acute or not.. ... ... ... ... ... ... ... ... ... .... C</paragraph>
<paragraph id="E36436B5FFEDFF96F57FFC9EFEDFF8F8" blockId="13.[833,1504,765,1036]" lastBlockId="13.[114,785,1788,1892]" pageId="13" pageNumber="221">
- Body dark brown; legs brown, lacking metallic reflections; pronotum without spots; three black spots laterally on elytron (one at humerus and two on lateral half) (
<figureCitation id="7BE02A30FFEDFF96F2D7FC56FAD0FC6C" box="[1257,1353,933,952]" captionStart="Figs" captionStartId="2.[85,120,1920,1934]" captionTargetBox="[253,1300,156,1885]" captionTargetId="figure-13@2.[252,1307,149,1889]" captionTargetPageId="2" captionText="Figs. 110. Pelidnota. 18, Pelidnota beckeri sp. nov.; 13, male holotype (dorsal, ventral, lateral); 46, female paratype (dorsal, ventral, lateral); 78, clypeus in dorsal view (male, female); 910, P.unicolor (Drury, 1778) (dorsal, lateral). Scale bars: Figs. 16 =9.0 mm, Figs.78 =2.5 mm, Figs. 910= 8.2mm." pageId="13" pageNumber="221">Figs. 1, 3</figureCitation>
); elytral apex truncated (
<figureCitation id="7BE02A30FFEDFF96F5F4FC32FB8DFC00" box="[970,1044,961,980]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="13" pageNumber="221">Fig. 33</figureCitation>
); parameres distal margin apex gradually acute laterally (
<figureCitation id="7BE02A30FFEDFF96F5CCFC2EFB10FC3B" box="[1010,1161,988,1008]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="13" pageNumber="221">Figs. 57, 58, 65</figureCitation>
). Distribution:
<collectingCountry id="9BCC7625FFEDFF96F31BFC2EFAF8FC24" box="[1317,1377,989,1008]" name="Brazil" pageId="13" pageNumber="221">Brazil</collectingCountry>
,
<collectingRegion id="211FF857FFEDFF96F354FC2EFA45FC24" box="[1386,1500,989,1008]" country="Brazil" name="Bahia" pageId="13" pageNumber="221">Bahia state</collectingRegion>
... ... ... ... ... ... ... ... ... ... ... ... ... ....
<taxonomicName id="24DB4D36FFEDFF96F2DCFC04FA10FBDF" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1250,1417,1015,1035]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="221" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFEDFF96F2DCFC04FA10FBDF" box="[1250,1417,1015,1035]" italics="true" pageId="13" pageNumber="221">Pelidnota beckeri</emphasis>
</taxonomicName>
<emphasis id="D1AFEAA7FFEDFF96F3B0FC0AFA46FBD8" bold="true" box="[1422,1503,1017,1036]" pageId="13" pageNumber="221">
<taxonomicNameLabel id="CA9C57DCFFEDFF96F3B0FC0AFA46FBD8" box="[1422,1503,1017,1036]" pageId="13" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
species of
<taxonomicName id="24DB4D36FFEDFF96F6E9F90FFE23F8C4" authority="(Soula, 2009)" baseAuthorityName="Soula" baseAuthorityYear="2009" box="[215,442,1788,1808]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="D1AFEAA7FFEDFF96F6E9F90FFEABF8C4" box="[215,306,1788,1808]" italics="true" pageId="13" pageNumber="221">Pelidnota</emphasis>
(Soula, 2009)
</taxonomicName>
. Key modified and translated from Soula (2009: 3034).
</paragraph>
<caption id="B7A4663DFFEDFF96F64CF94EFDB3F936" pageId="13" pageNumber="221" startId="13.[114,149,1725,1739]" targetBox="[179,720,149,1695]" targetPageId="13" targetType="figure">
<paragraph id="E36436B5FFEDFF96F64CF94EFDB3F936" blockId="13.[114,786,1725,1762]" pageId="13" pageNumber="221">
<emphasis id="D1AFEAA7FFEDFF96F64CF94EFF4AF91F" bold="true" box="[114,211,1725,1739]" pageId="13" pageNumber="221">Figs. 6977.</emphasis>
Labels attached to the hotypes. 6971,
<taxonomicName id="24DB4D36FFEDFF96F430F94EFD0BF918" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[526,658,1725,1740]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="221" phylum="Arthropoda" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="D1AFEAA7FFEDFF96F430F94EFD0BF918" box="[526,658,1725,1740]" italics="true" pageId="13" pageNumber="221">Pelidnota beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEDFF96F4AAF94DFD4CF918" box="[660,725,1726,1740]" pageId="13" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
; 7274,
<taxonomicName id="24DB4D36FFEDFF96F64CF920FF4EF936" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[114,215,1747,1762]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="221" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFEDFF96F64CF920FF4EF936" box="[114,215,1747,1762]" italics="true" pageId="13" pageNumber="221">P. nordestina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEDFF96F6E5F927FE84F936" box="[219,285,1748,1762]" pageId="13" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
; 7577,
<taxonomicName id="24DB4D36FFEDFF96F761F920FE7FF936" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[351,486,1747,1762]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="221" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFEDFF96F761F920FE7FF936" box="[351,486,1747,1762]" italics="true" pageId="13" pageNumber="221">P. pernambucana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA9C57DCFFEDFF96F7D5F927FDB3F936" box="[491,554,1748,1762]" pageId="13" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="E36436B5FFEDFF96F6ACF8C6FC88F8B0" blockId="13.[114,785,1788,1892]" pageId="13" pageNumber="221">
43(38) Andean species; light brown, fairly large and elongated body (length
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, width
<quantity id="24239B50FFEDFF96F7D2F8A2FDC3F8B0" box="[492,602,1873,1892]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="13" pageNumber="221" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">1214 mm</quantity>
).. ... ... ... .... 44
</paragraph>
<paragraph id="E36436B5FFEDFF96F64CF861FA79FA7A" blockId="13.[114,785,1937,1985]" lastBlockId="13.[833,1504,1100,1789]" pageId="13" pageNumber="221">
- Sub-Andean species, southernmost; usually smaller individuals (length
<quantity id="24239B50FFEDFF96F6ECF85EFED9F815" box="[210,320,1965,1985]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.4" metricValueMin="1.5" pageId="13" pageNumber="221" unit="mm" value="19.5" valueMax="24.0" valueMin="15.0">1524 mm</quantity>
, width
<quantity id="24239B50FFEDFF96F7B4F85EFE73F814" box="[394,490,1965,1984]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="13" pageNumber="221" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">913 mm</quantity>
) bodies, brown dorsally; legs C(B) Labrum triangular (
<figureCitation id="7BE02A30FFEDFF96F25FFBBFFB32FB8B" box="[1121,1195,1100,1119]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="13" pageNumber="221">Fig. 19</figureCitation>
); teeth on inner margin of mandible small (
<figureCitation id="7BE02A30FFEDFF96F5C7FB9BFBA3FBAF" box="[1017,1082,1128,1147]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="13" pageNumber="221">Fig. 23</figureCitation>
)); first and second teeth of maxillary galea sawed and IV palpomere less elongated (
<figureCitation id="7BE02A30FFEDFF96F33AFB77FAD3FB43" box="[1284,1354,1156,1175]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="13" pageNumber="221">Fig. 27</figureCitation>
); surface area of pre-mentum and insertion area of palp less excavated, lateral margin of post-mentum more rounded, and with elongated dense setae (
<figureCitation id="7BE02A30FFEDFF96F5AAFB2BFC4FFB3F" box="[916,982,1240,1259]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="13" pageNumber="221">Fig. 31</figureCitation>
)); elytra apex truncated and slightly acute (
<figureCitation id="7BE02A30FFEDFF96F3B0FB2BFA49FB3E" box="[1422,1488,1239,1259]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="13" pageNumber="221">Fig. 35</figureCitation>
); metacoxa with anteroexternal angle rounded and slightly acute (
<figureCitation id="7BE02A30FFEDFF96F565FAFCFC39FAF6" box="[859,928,1295,1314]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="13" pageNumber="221">Fig. 39</figureCitation>
); protibia with distal external tooth large and strongly recurved (
<figureCitation id="7BE02A30FFEDFF96F580FAD8FB9DFAEA" box="[958,1028,1323,1342]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="13" pageNumber="221">Fig. 43</figureCitation>
); pygidium dark brown, with reddish reflections on disk, with slight metallic green reflections on lateral area of posterior margin (
<figureCitation id="7BE02A30FFEDFF96F219FA90FBF3FAA2" box="[1063,1130,1379,1398]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="13" pageNumber="221">Fig. 45</figureCitation>
); parameres distal margin acute laterally (
<figureCitation id="7BE02A30FFEDFF96F5A2FA8CFBA0FA46" box="[924,1081,1407,1426]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="13" pageNumber="221">Figs. 61, 62, 67</figureCitation>
). Distribution:
<collectingCountry id="9BCC7625FFEDFF96F2E7FA8CFA8CFA46" box="[1241,1301,1407,1426]" name="Brazil" pageId="13" pageNumber="221">Brazil</collectingCountry>
,
<collectingRegion id="211FF857FFEDFF96F31FFA8CFA45FA46" box="[1313,1500,1407,1426]" country="Brazil" name="Pernambuco" pageId="13" pageNumber="221">Pernambuco state</collectingRegion>
... ... ... ... ... ... ... ... ... ... ... ... ....
<taxonomicName id="24DB4D36FFEDFF96F2FAFA69FA15FA7A" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1220,1420,1434,1454]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="221" phylum="Arthropoda" rank="species" species="nordestina" status="sp. nov.">
<emphasis id="D1AFEAA7FFEDFF96F2FAFA69FA15FA7A" box="[1220,1420,1434,1454]" italics="true" pageId="13" pageNumber="221">Pelidnota nordestina</emphasis>
</taxonomicName>
<emphasis id="D1AFEAA7FFEDFF96F3AEFA68FA79FA7A" bold="true" box="[1424,1504,1435,1454]" pageId="13" pageNumber="221">
<taxonomicNameLabel id="CA9C57DCFFEDFF96F3AEFA68FA79FA7A" box="[1424,1504,1435,1454]" pageId="13" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="E36436B5FFEDFF96F57FFA44FA4EF929" blockId="13.[833,1504,1100,1789]" pageId="13" pageNumber="221">
- Labrum transversal (
<figureCitation id="7BE02A30FFEDFF96F218FA44FBF0FA1E" box="[1062,1129,1463,1482]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="13" pageNumber="221">Fig. 20</figureCitation>
); teeth on inner margin of mandible larger (
<figureCitation id="7BE02A30FFEDFF96F59FFA20FC70FA32" box="[929,1001,1491,1510]" captionStart="Figs" captionStartId="5.[114,149,1733,1747]" captionTargetBox="[209,1398,152,1698]" captionTargetId="figure-13@5.[205,1411,148,1703]" captionTargetPageId="5" captionText="Figs. 1724. Mouthparts. 1720, labrum in frontal view; 17, P. beckeri sp. nov.; 18, P. unicolor (Drury); 19, P. nordestina sp. nov.; 20, P. pernambucana sp. nov.; 2124, left mandible in dorsolateral view; 21, Pelidnota beckeri sp. nov.; 22, P. unicolor (Drury); 23, P. nordestina sp. nov.; 24, P.pernambucana sp. nov. Scale bars: Figs.1724 = 2.0mm." pageId="13" pageNumber="221">Fig. 24</figureCitation>
); only first teeth of maxillary galea sawed, IV palpomere of maxilla, more elongated (
<figureCitation id="7BE02A30FFEDFF96F2C0FA1DFADCF9D6" box="[1278,1349,1518,1538]" captionStart="Figs" captionStartId="6.[85,120,1461,1475]" captionTargetBox="[246,1311,154,1428]" captionTargetId="figure-13@6.[245,1314,148,1431]" captionTargetPageId="6" captionText="Figs. 2528. Mouthparts. 25 and 26, left maxilla in dorsal view; 25, P. beckeri sp. nov.; 26, P. unicolor (Drury); 2728, left maxilla in dorsal view; 27, Pelidnota nordestina sp. nov.; 28, P.pernambucana sp. nov. Scale bars: Figs. 2528 = 2.0mm." pageId="13" pageNumber="221">Fig. 28</figureCitation>
); surface area of pre-mentum and insertion area of palp more excavated, lateral margin of post-mentum less rounded, and with elongate moderate setae (
<figureCitation id="7BE02A30FFEDFF96F236F9B1FBC8F981" box="[1032,1105,1602,1621]" captionStart="Figs" captionStartId="7.[114,149,1224,1238]" captionTargetBox="[224,1373,161,1190]" captionTargetId="figure-13@7.[223,1392,148,1193]" captionTargetPageId="7" captionText="Figs. 2932. Mouthparts. 2932, labium in dorsal view; 29, P.beckeri sp. nov.; 30, P. unicolor (Drury); 31, P.nordestina sp. nov.; 32, P.pernambucana sp. nov. Scale bars: Figs. 2932 = 2.0mm." pageId="13" pageNumber="221">Fig. 32</figureCitation>
); elytral apex truncated and slightly acute (
<figureCitation id="7BE02A30FFEDFF96F5A6F9ADFC42F9A5" box="[920,987,1630,1649]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="13" pageNumber="221">Fig. 36</figureCitation>
); anteroexternal angle of metacoxa strongly acute (
<figureCitation id="7BE02A30FFEDFF96F565F989FC3CF959" box="[859,933,1658,1677]" captionStart="Figs" captionStartId="8.[85,120,1869,1883]" captionTargetBox="[271,1287,149,1839]" captionTargetId="figure-13@8.[271,1287,149,1839]" captionTargetPageId="8" captionText="Figs. 3340. Elytral apex in frontal view and anteroexternal coxal angle of right metacoxa in lateral view. 3336, Elytral apex; 33, Pelidnota beckeri sp. nov.; 34, P. unicolor (Drury); 35, P. nordestina sp. nov.; 36, P. pernambucana sp. nov.; 3740, anteroexternal coxal angle of right metacoxa; 37, P. beckeri sp. nov.; 38, P. unicolor (Drury); 39, P. nordestina sp. nov.; 40, P. pernambucana sp. nov. Scale bars: Figs. 3336 =0.500 mm, Figs. 3740 =1.0 mm." pageId="13" pageNumber="221">Fig. 40</figureCitation>
); distal external tooth of protibia slightly recurved (
<figureCitation id="7BE02A30FFEDFF96F565F965FC07F97D" box="[859,926,1686,1705]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="13" pageNumber="221">Fig. 44</figureCitation>
); pygidium green and with metallic reflections (
<figureCitation id="7BE02A30FFEDFF96F3B3F965FA49F97D" box="[1421,1488,1686,1705]" captionStart="Figs" captionStartId="9.[114,149,1725,1739]" captionTargetBox="[348,1282,149,1690]" captionTargetId="figure-13@9.[331,1286,148,1695]" captionTargetPageId="9" captionText="Figs. 4146. Protibia and pygidium in dorsal view. 4144, left protíbia; 41, Pelidnota beckeri sp. nov.; 42, P. unicolor (Drury); 43, P. nordestina sp. nov.; 44, P. pernambucana sp. nov.; 4546, pygidium; 45, P. nordestina sp. nov.; 46, P.pernambucana sp. nov. Scale bars: Figs. 4142 = 1.0mm, Figs. 4576 =2.7 mm." pageId="13" pageNumber="221">Fig. 46</figureCitation>
); parameres distal margin rounded and slightly acute laterally (
<figureCitation id="7BE02A30FFEDFF96F5B4F93DFBB6F935" box="[906,1071,1742,1761]" captionStart="Figs" captionStartId="12.[85,120,1789,1803]" captionTargetBox="[219,1340,153,1758]" captionTargetId="figure-13@12.[219,1341,150,1758]" captionTargetPageId="12" captionText="Figs. 5768. Male genitalia and paramers margin drawing. 5764, male genitalia; 57 and 58, Pelidnota beckeri sp. nov. (dorsal, lateral); 59 and 60, P. unicolor (Drury) (dorsal, lateral); 61 and 62, P. nordestina sp. nov. (dorsal, lateral); 63 and 64, P.pernambucana sp. nov. (dorsal, lateral); 6568, paramers margin drawing; 65, P. beckeri sp. nov.; 66, P. unicolor (Drury); 67, P. nordestina sp. nov.; 68, P. pernambucana sp. nov. Scale bars: Figs. 5764 =2.5 mm." pageId="13" pageNumber="221">Figs. 63, 64, 68</figureCitation>
). Distribution:
<collectingCountry id="9BCC7625FFEDFF96F2EAF93DFA89F935" box="[1236,1296,1742,1761]" name="Brazil" pageId="13" pageNumber="221">Brazil</collectingCountry>
,
<collectingRegion id="211FF857FFEDFF96F320F93DFA45F935" box="[1310,1500,1742,1761]" country="Brazil" name="Pernambuco" pageId="13" pageNumber="221">Pernambuco state</collectingRegion>
... ... ... ... ... ... ... ... ... ... ....
<taxonomicName id="24DB4D36FFEDFF96F2B3F91AFA19F929" authorityName="Ferreira &amp; Almeida &amp; Bravo" authorityYear="2017" box="[1165,1408,1769,1789]" class="Insecta" family="Scarabaeidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="221" phylum="Arthropoda" rank="species" species="pernambucana" status="sp. nov.">
<emphasis id="D1AFEAA7FFEDFF96F2B3F91AFA19F929" box="[1165,1408,1769,1789]" italics="true" pageId="13" pageNumber="221">Pelidnota pernambucana</emphasis>
</taxonomicName>
<emphasis id="D1AFEAA7FFEDFF96F3B8F919FA4EF929" bold="true" box="[1414,1495,1770,1789]" pageId="13" pageNumber="221">
<taxonomicNameLabel id="CA9C57DCFFEDFF96F3B8F919FA4EF929" box="[1414,1495,1770,1789]" pageId="13" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="E36436B5FFEDFF96F56CF8CEFA46F85C" blockId="13.[833,1504,1853,1984]" pageId="13" pageNumber="221">52(37) Andean or Sub-Andean species; more or less uniformly green, elytron green and body ochre; punctures of the pronotum small to large.. ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... .... 53</paragraph>
<paragraph id="E36436B5FFEDFF96F57FF862FA79F814" blockId="13.[833,1504,1853,1984]" pageId="13" pageNumber="221">- Amazonian or Atlantic Forest species; color distinctly more ochre.. ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... .... 58</paragraph>
</subSubSection>
</treatment>
</document>