Add updates up until 2025-04-15 17:55:54

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ggserver 2025-04-15 18:02:01 +00:00
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@ -1,63 +1,63 @@
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<mods:title id="BA6C88C5A8BB3E4B13C19C3181184763">Hydroporus karabegan sp. nov. from eastern Türkiye (Coleoptera, Dytiscidae, Hydroporini)</mods:title>
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<mods:namePart id="B192308AAF231E4DBC6600076F696527">Fery, Hans</mods:namePart>
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@ -72,11 +72,8 @@
</subSubSection>
<subSubSection id="0E9843F0FFAE29467EFFFC49FA9BD67C" pageId="1" pageNumber="126" type="materials_examined">
<paragraph id="463D107BFFAE29467EFFFC49FD9FD1EC" blockId="1.[151,1436,942,1149]" pageId="1" pageNumber="126">
<materialsCitation id="F6EA1A26FFAE29467EFFFC49FD9BD1EC" country="Turkey" elevation="2180" latitude="38.621666" location="Akdaglar" longLatPrecision="19" longitude="40.135002" municipality="Aricak district" pageId="1" pageNumber="126" specimenCount="1" stateProvince="Elazig" typeStatus="holotype">
<emphasis id="74F6CC69FFAE29467EFFFC49FEBDD1C9" bold="true" box="[151,310,942,969]" pageId="1" pageNumber="126">
<typeStatus id="9939AED9FFAE29467EFFFC49FF59D1C8" box="[151,210,943,969]" pageId="1" pageNumber="126">Type</typeStatus>
locality
</emphasis>
<materialsCitation id="F6EA1A26FFAE29467EFFFC49FD9BD1EC" country="Turkey" elevation="2180" latitude="38.621666" location="Akdaglar plateau, ca. 7 km N Aricak" longLatPrecision="19" longitude="40.135002" municipality="Aricak district" pageId="1" pageNumber="126" specimenCount="1" stateProvince="Elazig province" typeStatus="holotype">
<emphasis id="74F6CC69FFAE29467EFFFC49FEBDD1C9" bold="true" box="[151,310,942,969]" pageId="1" pageNumber="126">Type locality</emphasis>
:
<collectingCountry id="3E9550EBFFAE29467F23FC49FE2CD1C8" box="[331,423,943,969]" name="Turkey" pageId="1" pageNumber="126">Türkiye</collectingCountry>
,
@ -84,12 +81,11 @@ locality
,
<collectingMunicipality id="A6598A01FFAE29467CE9FC49FCA0D1C8" box="[641,811,943,969]" pageId="1" pageNumber="126">Arıcak district</collectingMunicipality>
,
<location id="435D46A0FFAE29467D53FC49FC28D1C8" LSID="urn:lsid:plazi:treatment:CE2BA16DFFAE29427EFFFC80FB0ED3F4:435D46A0FFAE29467D53FC49FC28D1C8" box="[827,931,943,969]" country="Turkey" latitude="38.621666" longLatPrecision="19" longitude="40.135002" municipality="Aricak district" name="Akdaglar" pageId="1" pageNumber="126" stateProvince="Elazig">Akdağlar</location>
plateau, ca.
<locationDeviation id="578F0959FFAE29467A2AFC49FB76D1C8" box="[1090,1277,943,969]" location="Aricak" pageId="1" pageNumber="126">
<location id="435D46A0FFAE29467D53FC49FB76D1C8" LSID="urn:lsid:plazi:treatment:CE2BA16DFFAE29427EFFFC80FB44D730:435D46A0FFAE29467D53FC49FB76D1C8" box="[827,1277,943,969]" country="Turkey" latitude="38.621666" longLatPrecision="19" longitude="40.135002" municipality="Aricak district" name="Akdaglar plateau, ca. 7 km N Aricak" pageId="1" pageNumber="126" stateProvince="Elazig province">
Akdağlar plateau, ca.
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N Arıcak
</locationDeviation>
</location>
,
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<geoCoordinate id="23B676BCFFAE29467EFFFC35FEAFD1EC" box="[151,292,979,1005]" degrees="40" direction="east" minutes="08" orientation="longitude" pageId="1" pageNumber="126" precision="15" seconds="06" value="40.135002">40°0806E</geoCoordinate>
@ -103,7 +99,7 @@ N Arıcak
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<emphasis id="74F6CC69FFAE29467EAFFC11FEE6D611" bold="true" box="[199,365,1014,1041]" pageId="1" pageNumber="126">Type material</emphasis>
:
<materialsCitation id="F6EA1A26FFAE29467F12FC11FB25D658" collectingDate="2024-06-04" collectorName="M. Aykut &amp; S. Tusun" determinerName="Fery" elevation="2180" latitude="38.621666" location="Akdaglar" longLatPrecision="19" longitude="40.135002" municipality="Aricak distr." pageId="1" pageNumber="126" specimenCount="1" specimenCount-male="1" stateProvince="Elazig" typeStatus="holotype">
<materialsCitation id="F6EA1A26FFAE29467F12FC11FB25D658" collectingDate="2024-06-04" collectorName="M. Aykut &amp; S. Tusun" country="Turkey" determinerName="Fery" elevation="2180" latitude="38.621666" location="Akdaglar plateau / ca. 7 km N Aricak" longLatPrecision="19" longitude="40.135002" municipality="Aricak distr." pageId="1" pageNumber="126" specimenCount="1" specimenCount-male="1" stateProvince="Elazig prov." typeStatus="holotype">
<typeStatus id="9939AED9FFAE29467F12FC11FE6CD610" box="[378,487,1015,1041]" pageId="1" pageNumber="126" type="holotype">Holotype</typeStatus>
:
<specimenCount id="5084DBF2FFAE29467F98FC11FD8CD610" box="[496,519,1015,1041]" count="1" pageId="1" pageNumber="126" type="male"></specimenCount>
@ -116,12 +112,11 @@ N Arıcak
/
<collectingMunicipality id="A6598A01FFAE29467D32FC11FC6CD610" box="[858,999,1015,1041]" pageId="1" pageNumber="126">Arıcak distr.</collectingMunicipality>
,
<location id="435D46A0FFAE29467D87FC11FBDCD610" LSID="urn:lsid:plazi:treatment:CE2BA16DFFAE29427EFFFC80FB0ED3F4:435D46A0FFAE29467D87FC11FBDCD610" box="[1007,1111,1015,1041]" latitude="38.621666" longLatPrecision="19" longitude="40.135002" municipality="Aricak distr." name="Akdaglar" pageId="1" pageNumber="126" stateProvince="Elazig">Akdağlar</location>
plateau / ca.
<locationDeviation id="578F0959FFAE29467A8FFC11FA05D610" box="[1255,1422,1015,1041]" location="Aricak" pageId="1" pageNumber="126">
<location id="435D46A0FFAE29467D87FC11FA05D610" LSID="urn:lsid:plazi:treatment:CE2BA16DFFAE29427EFFFC80FB44D730:435D46A0FFAE29467D87FC11FA05D610" box="[1007,1422,1015,1041]" latitude="38.621666" longLatPrecision="19" longitude="40.135002" municipality="Aricak distr." name="Akdaglar plateau / ca. 7 km N Aricak" pageId="1" pageNumber="126" stateProvince="Elazig prov.">
Akdağlar plateau / ca.
<quantity id="817ABD9EFFAE29467A8FFC11FA94D610" box="[1255,1311,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="126" unit="km" value="7.0">7 km</quantity>
N Arıcak
</locationDeviation>
</location>
”, “
<geoCoordinate id="23B676BCFFAE29467ECBFBFDFEBAD634" box="[163,305,1051,1077]" degrees="38" direction="north" minutes="37" orientation="latitude" pageId="1" pageNumber="126" precision="15" seconds="18" value="38.621666">38°3718N</geoCoordinate>
<geoCoordinate id="23B676BCFFAE29467F55FBFDFE41D634" box="[317,458,1051,1077]" degrees="40" direction="east" minutes="08" orientation="longitude" pageId="1" pageNumber="126" precision="15" seconds="06" value="40.135002">40°0806E</geoCoordinate>
@ -133,9 +128,7 @@ N Arıcak
<collectorName id="EB7775ADFFAE29467D8CFBFDFBD9D634" box="[996,1106,1051,1077]" pageId="1" pageNumber="126">M. Aykut</collectorName>
&amp;
<collectorName id="EB7775ADFFAE29467A17FBFDFB6DD634" box="[1151,1254,1051,1077]" pageId="1" pageNumber="126">S. Tusun</collectorName>
leg.” [printed], “
<typeStatus id="9939AED9FFAE29467EC9FBD9FE87D658" box="[161,268,1087,1113]" pageId="1" pageNumber="126" type="holotype">Holotype</typeStatus>
/
leg.” [printed], “Holotype /
<taxonomicName id="81826BF8FFAE29467F49FBD9FE2CD658" authorityName="Clairville" authorityYear="1806" box="[289,423,1087,1113]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="74F6CC69FFAE29467F49FBD9FE2CD658" box="[289,423,1087,1113]" italics="true" pageId="1" pageNumber="126">Hydroporus</emphasis>
</taxonomicName>
@ -146,9 +139,7 @@ leg.” [printed], “
<emphasis id="74F6CC69FFAE29467C53FBD9FD1ED658" bold="true" box="[571,661,1087,1113]" pageId="1" pageNumber="126">
<taxonomicNameLabel id="6FC57112FFAE29467C53FBD9FD1ED658" box="[571,661,1087,1113]" pageId="1" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
/ Aykut, Tusun &amp;
<determinerName id="B3E1FCBFFFAE29467D0AFBD9FC1ED658" box="[866,917,1087,1113]" pageId="1" pageNumber="126">Fery</determinerName>
det. 2005” [red, printed]
/ Aykut, Tusun &amp; Fery det. 2005” [red, printed]
</materialsCitation>
.
<materialsCitation id="F6EA1A26FFAE29467AD0FBD9FA87D67C" collectingDate="2024-08-28" collectorName="M. Aykut &amp; S. Tusun" determinerName="Fery" elevation="2180" latitude="38.621666" location="Akdaglar" longLatPrecision="19" longitude="40.135002" municipality="Aricak distr." pageId="1" pageNumber="126" specimenCount="9" specimenCount-female="2" specimenCount-male="7" stateProvince="Elazig" typeStatus="paratype">
@ -157,15 +148,11 @@ det. 2005” [red, printed]
<specimenCount id="5084DBF2FFAE29467B5CFBD9FAEBD658" box="[1332,1376,1087,1113]" count="7" pageId="1" pageNumber="126" type="male">7 ♂</specimenCount>
,
<specimenCount id="5084DBF2FFAE29467B03FBD9FA1ED658" box="[1387,1429,1087,1113]" count="2" pageId="1" pageNumber="126" type="female">2 ♀</specimenCount>
, same label texts as
<typeStatus id="9939AED9FFAE29467F07FB85FE5ED67C" box="[367,469,1123,1149]" pageId="1" pageNumber="126" type="holotype">holotype</typeStatus>
; 8 exs, idem, but date
, same label texts as holotype; 8 exs, idem, but date
<date id="323C36BBFFAE29467CA7FB85FCC8D67C" box="[719,835,1123,1149]" pageId="1" pageNumber="126" value="2024-08-28">
<collectingDate id="2278CF53FFAE29467CA7FB85FCC8D67C" box="[719,835,1123,1149]" pageId="1" pageNumber="126" value="2024-08-28">28.8.2024</collectingDate>
</date>
; all
<typeStatus id="9939AED9FFAE29467D19FB85FC57D67C" box="[881,988,1123,1149]" pageId="1" pageNumber="126" type="paratype">paratypes</typeStatus>
with a respective red label
; all paratypes with a respective red label
</materialsCitation>
.
</paragraph>
@ -344,5 +331,76 @@ Fery, 2009
. The specific name of the new species relates to “Karabegan”, a historical name of the Arıcak district. It is a noun in the nominative singular standing in apposition to the generic name.
</paragraph>
</subSubSection>
<subSubSection id="0E9843F0FFAA29427EFFFDA0FC1DD634" pageId="5" pageNumber="130" type="diagnosis">
<paragraph id="463D107BFFAA29427EFFFDA0FE28D061" blockId="5.[151,419,582,608]" box="[151,419,582,608]" pageId="5" pageNumber="130">
<heading id="1D75A717FFAA29427EFFFDA0FE28D061" bold="true" box="[151,419,582,608]" fontSize="11" level="1" pageId="5" pageNumber="130" reason="1">
<emphasis id="74F6CC69FFAA29427EFFFDA0FE28D061" bold="true" box="[151,419,582,608]" pageId="5" pageNumber="130">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="463D107BFFAA29427EFFFD69FB01D1A4" blockId="5.[151,1437,655,1329]" pageId="5" pageNumber="130">
The new species is similar to
<taxonomicName id="81826BF8FFAA29427F84FD69FD79D0A8" authorityName="Fery" authorityYear="2009" box="[492,754,655,681]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="holzschuhi">
<emphasis id="74F6CC69FFAA29427F84FD69FD79D0A8" box="[492,754,655,681]" italics="true" pageId="5" pageNumber="130">Hydroporus holzschuhi</emphasis>
</taxonomicName>
and
<taxonomicName id="81826BF8FFAA29427D44FD69FC68D0A9" authorityName="Erman &amp; Fery" authorityYear="2000" box="[812,995,655,680]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="erzerumensis">
<emphasis id="74F6CC69FFAA29427D44FD69FC68D0A9" box="[812,995,655,680]" italics="true" pageId="5" pageNumber="130">H. erzerumensis</emphasis>
</taxonomicName>
, both also from
<collectingCountry id="3E9550EBFFAA29427AC9FD69FB76D0A8" box="[1185,1277,655,681]" name="Turkey" pageId="5" pageNumber="130">Türkiye</collectingCountry>
. Compared to the former it is somewhat more elongate because of the slightly more parallel sides of the body. The two species are best distinguished by the shape of the median lobe (cf.
<figureCitation id="DEB90CFEFFAA29427D5AFD31FC4BD0F0" box="[818,960,727,753]" captionStart="FIGURE 49" captionStartId="3.[151,250,1767,1791]" captionTargetBox="[151,1433,184,1737]" captionTargetId="figure-17@3.[151,1436,181,1743]" captionTargetPageId="3" captionText="FIGURE 49. (47) Hydroporus karabegan sp. nov.: (4) median lobe of aedeagus (a: ventral view, b: lateral view); (5) left paramere; (6) gonocoxosternum; (7) gonocoxae. (89) median lobe of aedeagus of (8) Hydroporus holzschuhi Fery, 2009 (redrawn from Fery 2000: 536); (9), idem of H. erzurumensis Erman &amp; Fery, 2000 (redrawn from Erman &amp; Fery 2000: 174)." figureDoi="http://doi.org/10.5281/zenodo.15214877" httpUri="https://zenodo.org/record/15214877/files/figure.png" pageId="5" pageNumber="130">Figs 4 and 8</figureCitation>
). Both have the distal part in ventral view evenly rounded; however, the lobe of the new species is distinctly broader. In lateral view, the lobe of
<taxonomicName id="81826BF8FFAA29427B6BFD1DFA17D114" authorityName="Fery" authorityYear="2009" box="[1283,1436,763,789]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="holzschuhi">
<emphasis id="74F6CC69FFAA29427B6BFD1DFA17D114" box="[1283,1436,763,789]" italics="true" pageId="5" pageNumber="130">H. holzschuhi</emphasis>
</taxonomicName>
is basally much more bent and in distal third more or less straight. That of the new species is basally less bent and in distal third still slightly curved. From the more northern
<taxonomicName id="81826BF8FFAA29427D11FCA5FBB8D15D" authorityName="Erman &amp; Fery" authorityYear="2000" box="[889,1075,835,860]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="erzurumensis">
<emphasis id="74F6CC69FFAA29427D11FCA5FBB8D15D" box="[889,1075,835,860]" italics="true" pageId="5" pageNumber="130">H. erzurumensis</emphasis>
</taxonomicName>
the new species can be clearly distinguished by the shape of the distal part of the median lobe in ventral view (
<figureCitation id="DEB90CFEFFAA29427A7FFC81FBD7D180" box="[1047,1116,871,897]" captionStart="FIGURE 49" captionStartId="3.[151,250,1767,1791]" captionTargetBox="[151,1433,184,1737]" captionTargetId="figure-17@3.[151,1436,181,1743]" captionTargetPageId="3" captionText="FIGURE 49. (47) Hydroporus karabegan sp. nov.: (4) median lobe of aedeagus (a: ventral view, b: lateral view); (5) left paramere; (6) gonocoxosternum; (7) gonocoxae. (89) median lobe of aedeagus of (8) Hydroporus holzschuhi Fery, 2009 (redrawn from Fery 2000: 536); (9), idem of H. erzurumensis Erman &amp; Fery, 2000 (redrawn from Erman &amp; Fery 2000: 174)." figureDoi="http://doi.org/10.5281/zenodo.15214877" httpUri="https://zenodo.org/record/15214877/files/figure.png" pageId="5" pageNumber="130">Fig. 9</figureCitation>
). For comparison with other species from the Eastern Mediterranean region see Fery (2009) and
<bibRefCitation id="22136D8AFFAA29427DF8FC6DFB0DD1A4" author="Fery, H. &amp; Erman, O. K." box="[912,1158,907,933]" pageId="5" pageNumber="130" pagination="1 - 12" refId="ref3562" refString="Fery, H. &amp; Erman, O. K. (2009) Five new species of the longulus-group of Hydroporus Clairville, 1806 from north-eastern Turkey (Coleoptera: Dytiscidae). Zootaxa, 2033 (1), 1-12. https://doi.org/10.11646/zootaxa.2033.1.1" type="journal article" year="2009">Fery &amp; Erman (2009)</bibRefCitation>
.
</paragraph>
<paragraph id="463D107BFFAA29427EAFFC49FC1DD634" blockId="5.[151,1437,655,1329]" pageId="5" pageNumber="130">
Differences in the shape of the parameres and female genitalia between the three species have also been recognised but, considering the small amount of specimens of
<taxonomicName id="81826BF8FFAA29427D37FC35FC70D1EC" authorityName="Fery" authorityYear="2009" box="[863,1019,979,1005]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="holzschuhi">
<emphasis id="74F6CC69FFAA29427D37FC35FC70D1EC" box="[863,1019,979,1005]" italics="true" pageId="5" pageNumber="130">H. holzschuhi</emphasis>
</taxonomicName>
and
<taxonomicName id="81826BF8FFAA29427A5BFC35FB67D1ED" authorityName="Erman &amp; Fery" authorityYear="2000" box="[1075,1260,979,1004]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="erzurumensis">
<emphasis id="74F6CC69FFAA29427A5BFC35FB67D1ED" box="[1075,1260,979,1004]" italics="true" pageId="5" pageNumber="130">H. erzurumensis</emphasis>
</taxonomicName>
to our disposal we refrain from giving details since we believe that distinguishing the species based solely on female characters is not at all trustworthy (as it is in most species of the
<taxonomicName id="81826BF8FFAA29427CB2FBFDFCB1D634" authorityName="Mulsant &amp; Rey" authorityYear="1861" box="[730,826,1051,1077]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="74F6CC69FFAA29427CB2FBFDFCB1D634" box="[730,826,1051,1077]" italics="true" pageId="5" pageNumber="130">longulus</emphasis>
</taxonomicName>
-group).
</paragraph>
</subSubSection>
<subSubSection id="0E9843F0FFAA29427EAFFBD9FB44D730" pageId="5" pageNumber="130" type="discussion">
<paragraph id="463D107BFFAA29427EAFFBD9FB44D730" blockId="5.[151,1437,655,1329]" pageId="5" pageNumber="130">
<emphasis id="74F6CC69FFAA29427EAFFBD9FE81D658" bold="true" box="[199,266,1087,1113]" pageId="5" pageNumber="130">Notes</emphasis>
: The distance between the collecting site of the new species and that of
<taxonomicName id="81826BF8FFAA29427A2AFBD9FB56D658" authorityName="Fery" authorityYear="2009" box="[1090,1245,1087,1113]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="holzschuhi">
<emphasis id="74F6CC69FFAA29427A2AFBD9FB56D658" box="[1090,1245,1087,1113]" italics="true" pageId="5" pageNumber="130">H. holzschuhi</emphasis>
</taxonomicName>
is only about
<quantity id="817ABD9EFFAA29427BE8FBD8FF35D67C" metricMagnitude="4" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="130" unit="km" value="95.0">95 km</quantity>
. Therefore, at first glance, it is somewhat surprising that two different species were found in areas so close to each other. However, it must be considered that the Akdağlar plateau in the Akdağ mountains and the Şerafettin mountains (area of
<taxonomicName id="81826BF8FFAA29427F14FB4DFD91D6C4" authorityName="Fery" authorityYear="2009" box="[380,538,1195,1221]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="130" phylum="Arthropoda" rank="species" species="holzschuhi">
<emphasis id="74F6CC69FFAA29427F14FB4DFD91D6C4" box="[380,538,1195,1221]" italics="true" pageId="5" pageNumber="130">H. holzschuhi</emphasis>
</taxonomicName>
) are separated by the Murat River basin. Additionally, the Akdağ mountains belongs to the Maden mountain range, a part of the eastern Taurus mountains group, and consists of a wide variety of sedimentary rocks, primarily volcanic rocks, which is called the Maden Complex in terms of lithology. The Şerafettin mountains, on the other hand, consist of basalt, andesite and tufa (
<bibRefCitation id="22136D8AFFAA29427D85FAF1FB48D730" author="Nehme, C. &amp; Al, A. &amp; Karadogan, S. &amp; Kuzucuoglu, C. &amp; Pons-Branchu, E. &amp; Mouralis, D." box="[1005,1219,1303,1329]" pageId="5" pageNumber="130" pagination="147 - 159" refId="ref3992" refString="Nehme, C., Al, A., Karadogan, S., Kuzucuoglu, C., Pons-Branchu, E., &amp; Mouralis, D. (2023). Karst landscapes in south-eastern Turkey: potential of cave speleothems to record the last glacial maximum and Holocene climate. Quaternaire. Revue de l'Association Francaise pour l'Etude du Quaternaire, 34 (3), 147-159. https://doi.org/10.4000/quaternaire.18304" type="journal article" year="2023">
Nehme
<emphasis id="74F6CC69FFAA29427A2FFAF1FBF4D730" box="[1095,1151,1303,1329]" italics="true" pageId="5" pageNumber="130">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>