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<mods:title id="248625C077DE371D814A106A30489BF8">A new species of Dilophochila Bates, 1888 with comments on natural history and distribution of the genus (Coleoptera: Melolonthidae: Rutelinae)</mods:title>
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<mods:title id="87FF9B2E2DB2F0EA2767E9797C8A9326">A new species of Dilophochila Bates, 1888 with comments on natural history and distribution of the genus (Coleoptera: Melolonthidae: Rutelinae)</mods:title>
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<mods:namePart id="D325C04F6A401E77BDF8BDDB4893CB00">Ramírez-Ponce, Andrés</mods:namePart>
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<mods:affiliation id="D3522C318BF31DEB157736DD8D483D53">de Ecología, A. C. (INECOL), Red de Biodiversidad y Sistemática. Xalapa, Veracruz, México.</mods:affiliation>
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<mods:namePart id="848C160E2FE9B2C6B6C27E7C02F4234D">Zaragoza-Caballero, Santiago</mods:namePart>
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<mods:affiliation id="4EF5748AA140958F987692BD5E82D2AB">Universidad Nacional Autónoma de México (UNAM), Instituto de Biología (IB). México, D. F., México.</mods:affiliation>
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<mods:namePart id="4A4AC8F071F6E5A65558E15D2103774B">Curoe, Daniel J.</mods:namePart>
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<subSubSection id="C331657CFFD0C873B199F8D3C22CD996" box="[244,568,1914,1939]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8B9436F7FFD0C873B199F8D3C22CD996" blockId="1.[197,616,1786,1971]" box="[244,568,1914,1939]" pageId="1" pageNumber="2">
<emphasis id="B95FEAE5FFD0C873B199F8D3C22CD996" bold="true" box="[244,568,1914,1939]" pageId="1" pageNumber="2">
@ -67,9 +69,9 @@
<subSubSection id="C331657CFFD0C873B066F833C235D9B6" box="[267,545,1946,1971]" pageId="1" pageNumber="2" type="description">
<paragraph id="8B9436F7FFD0C873B066F833C235D9B6" blockId="1.[197,616,1786,1971]" box="[267,545,1946,1971]" pageId="1" pageNumber="2">
<emphasis id="B95FEAE5FFD0C873B066F833C235D9B6" bold="true" box="[267,545,1946,1971]" pageId="1" pageNumber="2">
<figureCitation id="13102A72FFD0C873B066F833C1CFD9B6" box="[267,475,1946,1971]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." pageId="1" pageNumber="2">Figs. 1A, B, C, E, G</figureCitation>
<figureCitation id="13102A72FFD0C873B066F833C1CFD9B6" box="[267,475,1946,1971]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.13903743" httpUri="https://zenodo.org/record/13903743/files/figure.png" pageId="1" pageNumber="2">Figs. 1A, B, C, E, G</figureCitation>
,
<figureCitation id="13102A72FFD0C873B088F833C235D9B6" box="[485,545,1946,1971]" captionStart="Figure 2" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[99,1412,669,2045]" captionTargetId="figure-200@3.[94,1417,640,2076]" captionTargetPageId="3" captionText="Figure 2. Dilophochilaglabra sp.nov.(A) Lateral habitus;(B)Ventral habitus." pageId="1" pageNumber="2">2A, B</figureCitation>
<figureCitation id="13102A72FFD0C873B088F833C235D9B6" box="[485,545,1946,1971]" captionStart="Figure 2" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[99,1412,669,2045]" captionTargetId="figure-200@3.[94,1417,640,2076]" captionTargetPageId="3" captionText="Figure 2. Dilophochilaglabra sp.nov.(A) Lateral habitus;(B)Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.13903745" httpUri="https://zenodo.org/record/13903745/files/figure.png" pageId="1" pageNumber="2">2A, B</figureCitation>
</emphasis>
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@ -325,11 +327,11 @@ DJCC (
.
<emphasis id="B95FEAE5FFD0C873B54CFC33C472DDB6" bold="true" box="[1057,1126,922,947]" italics="true" pageId="1" pageNumber="2">Color:</emphasis>
Ground color ocher; frons and pronotal discal vittae deep metallic green; elytral intervals IV, VII and IX slightly darker (
<figureCitation id="13102A72FFD0C873B5B1FC73C53BDDF6" box="[1244,1327,986,1011]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." pageId="1" pageNumber="2">Fig. 1A</figureCitation>
<figureCitation id="13102A72FFD0C873B5B1FC73C53BDDF6" box="[1244,1327,986,1011]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.13903743" httpUri="https://zenodo.org/record/13903743/files/figure.png" pageId="1" pageNumber="2">Fig. 1A</figureCitation>
).
<emphasis id="B95FEAE5FFD0C873B42BFC73C59EDDF6" bold="true" box="[1350,1418,986,1011]" italics="true" pageId="1" pageNumber="2">Head:</emphasis>
Clypeal margin with v-shaped notch; noticeably reflexed anteriorly; clypeal surface densely rugose, with scattered, moderately short, slightly curved setae; frontoclypeal suture complete (
<figureCitation id="13102A72FFD0C873B559FBF3C494DA76" box="[1076,1152,1114,1139]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." pageId="1" pageNumber="2">Fig. 1B</figureCitation>
<figureCitation id="13102A72FFD0C873B559FBF3C494DA76" box="[1076,1152,1114,1139]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.13903743" httpUri="https://zenodo.org/record/13903743/files/figure.png" pageId="1" pageNumber="2">Fig. 1B</figureCitation>
). Frons with depression widening anteriorly; surface of depression as on clypeus except without setae; rest of surface of frons punctate; punctures dense,large, contiguous anteriorly, moderately dense and shallow posterior to eyes; supraocular area with 6-7 long, decumbent setae.
<emphasis id="B95FEAE5FFD0C873B5E9FB53C4EBDB16" bold="true" box="[1156,1279,1274,1299]" italics="true" pageId="1" pageNumber="2">Pronotum:</emphasis>
Anterior angles acute and projected anteriorly, lateral margins sinuate, posterior angles slightly obtuse; surface irregularly, not densely punctate; punctures large; each side with 2 long, decumbent setae; anterior and lateral margins with long, irregularly spaced setae; lateral and posterior margins beaded.
@ -337,13 +339,13 @@ Anterior angles acute and projected anteriorly, lateral margins sinuate, posteri
Surface irregularly, densely punctate; punctures moderate in size.
<emphasis id="B95FEAE5FFD0C873B5ADFA73C50FDBF6" bold="true" box="[1216,1307,1498,1523]" italics="true" pageId="1" pageNumber="2">Elytron:</emphasis>
Glabrous, with 10 striae between suture and humeral umbone; striae slightly impressed by rows of large, round punctures separated approximately by their own diameter; intervals IV, VII and IX wider, clearly more elevated and darker than adjacent intervals (
<figureCitation id="13102A72FFD0C873B54EF9D3C465D896" box="[1059,1137,1658,1683]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." pageId="1" pageNumber="2">Fig. 1A</figureCitation>
<figureCitation id="13102A72FFD0C873B54EF9D3C465D896" box="[1059,1137,1658,1683]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.13903743" httpUri="https://zenodo.org/record/13903743/files/figure.png" pageId="1" pageNumber="2">Fig. 1A</figureCitation>
); epipleuron not expanded in basal half (
<figureCitation id="13102A72FFD0C873B2A7F933C40DD8B6" box="[970,1049,1690,1715]" captionStart="Figure 2" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[99,1412,669,2045]" captionTargetId="figure-200@3.[94,1417,640,2076]" captionTargetPageId="3" captionText="Figure 2. Dilophochilaglabra sp.nov.(A) Lateral habitus;(B)Ventral habitus." pageId="1" pageNumber="2">Fig. 2A</figureCitation>
<figureCitation id="13102A72FFD0C873B2A7F933C40DD8B6" box="[970,1049,1690,1715]" captionStart="Figure 2" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[99,1412,669,2045]" captionTargetId="figure-200@3.[94,1417,640,2076]" captionTargetPageId="3" captionText="Figure 2. Dilophochilaglabra sp.nov.(A) Lateral habitus;(B)Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.13903745" httpUri="https://zenodo.org/record/13903745/files/figure.png" pageId="1" pageNumber="2">Fig. 2A</figureCitation>
).
<emphasis id="B95FEAE5FFD0C873B545F933C46FD8B6" bold="true" box="[1064,1147,1690,1715]" italics="true" pageId="1" pageNumber="2">Venter:</emphasis>
Surface completely covered by long, scattered setae (
<figureCitation id="13102A72FFD0C873B500F913C4A8D8D6" box="[1133,1212,1722,1747]" captionStart="Figure 2" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[99,1412,669,2045]" captionTargetId="figure-200@3.[94,1417,640,2076]" captionTargetPageId="3" captionText="Figure 2. Dilophochilaglabra sp.nov.(A) Lateral habitus;(B)Ventral habitus." pageId="1" pageNumber="2">Fig. 2B</figureCitation>
<figureCitation id="13102A72FFD0C873B500F913C4A8D8D6" box="[1133,1212,1722,1747]" captionStart="Figure 2" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[99,1412,669,2045]" captionTargetId="figure-200@3.[94,1417,640,2076]" captionTargetPageId="3" captionText="Figure 2. Dilophochilaglabra sp.nov.(A) Lateral habitus;(B)Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.13903745" httpUri="https://zenodo.org/record/13903745/files/figure.png" pageId="1" pageNumber="2">Fig. 2B</figureCitation>
).
<taxonomicName id="4C2B4D74FFD0C873B5A0F913C549D8D6" authority=": In" authorityName="In" box="[1229,1373,1722,1747]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="2" phylum="Chordata" rank="genus">
<emphasis id="B95FEAE5FFD0C873B5A0F913C554D8D6" bold="true" box="[1229,1344,1722,1747]" italics="true" pageId="1" pageNumber="2">Pygidium:</emphasis>
@ -352,14 +354,14 @@ In
lateral view, uniformly convex; surface moderately densely setose.
<emphasis id="B95FEAE5FFD0C873B2E3F953C3DDD916" bold="true" box="[910,969,1786,1811]" italics="true" pageId="1" pageNumber="2">Legs:</emphasis>
Protibia (
<figureCitation id="13102A72FFD0C873B556F953C49FD916" box="[1083,1163,1786,1811]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." pageId="1" pageNumber="2">Fig. 1C</figureCitation>
<figureCitation id="13102A72FFD0C873B556F953C49FD916" box="[1083,1163,1786,1811]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.13903743" httpUri="https://zenodo.org/record/13903743/files/figure.png" pageId="1" pageNumber="2">Fig. 1C</figureCitation>
) with margins mostly subparallel; external margin with 2 distinct teeth; subapical projection small, obtuse, weakly broadened; protarsomeres 1-4 moderately wide; protarsomere 5 with 2 basal, acute denticles; inner protarsal claw angulate and narrowly cleft.
<emphasis id="B95FEAE5FFD0C873B2D1F833C47FD9B6" bold="true" box="[956,1131,1946,1971]" italics="true" pageId="1" pageNumber="2">Male Genitalia:</emphasis>
Base of parameres broadly and evenly curved; in caudal view, parameres elongate subtriangular, external margins weakly sinuate before preapical constriction; apices markedly broadened, rounded; in lateral view, ventral margin not folded (arrow) (
<figureCitation id="13102A72FFD0C873B235F793C3B2D656" box="[856,934,2106,2131]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." pageId="1" pageNumber="2">Fig. 1G</figureCitation>
<figureCitation id="13102A72FFD0C873B235F793C3B2D656" box="[856,934,2106,2131]" captionStart="Figure 1" captionStartId="2.[170,229,2074,2099]" captionTargetBox="[185,1482,206,2029]" captionTargetId="figure-1@2.[180,1484,204,2044]" captionTargetPageId="2" captionText="Figure 1. Diagnostic characters of Dilophochila glabra sp.nov. (A,B,C,E, G) and D.miahuatleca Morón &amp; Howden (D, F,H). (A) Dorsal habitus;(B) Head,pronotum and anterior legs;(C, D) Protibia;(E,F) Parameres (caudal view);(G,H) Aedeagus (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.13903743" httpUri="https://zenodo.org/record/13903743/files/figure.png" pageId="1" pageNumber="2">Fig. 1G</figureCitation>
).
</paragraph>
<caption id="DF54667FFFD3C870B1C7F7B3C3BBD656" pageId="2" pageNumber="3" startId="2.[170,229,2074,2099]" targetBox="[185,1482,206,2029]" targetPageId="2" targetType="figure">
<caption id="DF54667FFFD3C870B1C7F7B3C3BBD656" ID-DOI="http://doi.org/10.5281/zenodo.13903743" ID-Zenodo-Dep="13903743" httpUri="https://zenodo.org/record/13903743/files/figure.png" pageId="2" pageNumber="3" startId="2.[170,229,2074,2099]" targetBox="[185,1482,206,2029]" targetPageId="2" targetType="figure">
<paragraph id="8B9436F7FFD3C870B1C7F7B3C3BBD656" blockId="2.[170,1493,2074,2131]" pageId="2" pageNumber="3">
Figure 1. Diagnostic characters of
<taxonomicName id="4C2B4D74FFD3C870B0ABF7B3C275D636" authority="Ramírez-Ponce &amp; Zaragoza-Caballero &amp; Curoe, 2023" authorityName="Ramírez-Ponce &amp; Zaragoza-Caballero &amp; Curoe" authorityYear="2023" box="[454,609,2074,2099]" class="Insecta" family="Scarabaeidae" genus="Dilophochila" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="glabra" status="sp. nov.">
@ -430,11 +432,11 @@ which are remarkably acute and expanded externally, and the base of parameres no
<emphasis id="B95FEAE5FFD5C876B370FB93C2D7DA56" box="[541,707,1082,1107]" italics="true" pageId="4" pageNumber="5">D. miahuatleca</emphasis>
</taxonomicName>
). In addition, they are distributed in two mountainous systems separated by the central valleys of the state of Oaxaca (
<figureCitation id="13102A72FFD5C876B1DCFB33C0F9DAB6" box="[177,237,1178,1203]" captionStart="Figure 3" captionStartId="4.[170,229,707,732]" captionTargetBox="[172,793,206,675]" captionTargetId="figure-848@4.[170,794,204,676]" captionTargetPageId="4" captionText="Figure 3. Distribution of the species of the genus Dilophochila Bates." pageId="4" pageNumber="5">Fig. 3</figureCitation>
<figureCitation id="13102A72FFD5C876B1DCFB33C0F9DAB6" box="[177,237,1178,1203]" captionStart="Figure 3" captionStartId="4.[170,229,707,732]" captionTargetBox="[172,793,206,675]" captionTargetId="figure-848@4.[170,794,204,676]" captionTargetPageId="4" captionText="Figure 3. Distribution of the species of the genus Dilophochila Bates." figureDoi="http://doi.org/10.5281/zenodo.13903747" httpUri="https://zenodo.org/record/13903747/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF54667FFFD2C871B133F793C2CED656" box="[94,730,2106,2131]" pageId="3" pageNumber="4" startId="3.[94,153,2106,2131]" targetBox="[99,1412,669,2045]" targetPageId="3" targetType="figure">
<caption id="DF54667FFFD2C871B133F793C2CED656" ID-DOI="http://doi.org/10.5281/zenodo.13903745" ID-Zenodo-Dep="13903745" box="[94,730,2106,2131]" httpUri="https://zenodo.org/record/13903745/files/figure.png" pageId="3" pageNumber="4" startId="3.[94,153,2106,2131]" targetBox="[99,1412,669,2045]" targetPageId="3" targetType="figure">
<paragraph id="8B9436F7FFD2C871B133F793C2CED656" blockId="3.[94,730,2106,2131]" box="[94,730,2106,2131]" pageId="3" pageNumber="4">
Figure 2.
<taxonomicName id="4C2B4D74FFD2C871B1D9F793C15AD656" authority="Ramírez-Ponce &amp; Zaragoza-Caballero &amp; Curoe, 2023" authorityName="Ramírez-Ponce &amp; Zaragoza-Caballero &amp; Curoe" authorityYear="2023" box="[180,334,2106,2131]" class="Insecta" family="Scarabaeidae" genus="Dilophochila" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="glabra" status="sp. nov.">
@ -444,7 +446,7 @@ Figure 2.
(A) Lateral habitus;(B)Ventral habitus.
</paragraph>
</caption>
<caption id="DF54667FFFD5C876B1C7FD6AC2CFDCD9" box="[170,731,707,732]" pageId="4" pageNumber="5" startId="4.[170,229,707,732]" targetBox="[172,793,206,675]" targetPageId="4" targetType="figure">
<caption id="DF54667FFFD5C876B1C7FD6AC2CFDCD9" ID-DOI="http://doi.org/10.5281/zenodo.13903747" ID-Zenodo-Dep="13903747" box="[170,731,707,732]" httpUri="https://zenodo.org/record/13903747/files/figure.png" pageId="4" pageNumber="5" startId="4.[170,229,707,732]" targetBox="[172,793,206,675]" targetPageId="4" targetType="figure">
<paragraph id="8B9436F7FFD5C876B1C7FD6AC2CFDCD9" blockId="4.[170,731,707,732]" box="[170,731,707,732]" pageId="4" pageNumber="5">
Figure 3. Distribution of the species of the genus
<taxonomicName id="4C2B4D74FFD5C876B32DFD6AC2CFDCD9" authority="Bates." authorityName="Bates." box="[576,731,707,732]" class="Insecta" family="Scarabaeidae" genus="Dilophochila" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">