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<document ID-DOI="http://dx.doi.org/10.3897/BDJ.4.e8029" ID-PMC="PMC4867705" ID-Pensoft-Pub="1314-2828-4-8029" ID-PubMed="27226756" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2828-4-e8029" ModsDocOrigin="Biodiversity Data Journal 4" ModsDocTitle="Description of a new species of Acrostichus Rahm 1928 (Nematoda: Diplogastridae) from India with a note on its position and relationship with the congeners" checkinTime="1460370306522" checkinUser="pensoft" docAuthor="Tahseen, Qudsia, Ahlawat, Shikha, Asif, Mohammad &amp; Mustaqim, Malka" docDate="2016" docId="255948EE5432DFECD7DC036D54F03F21" docLanguage="en" docName="BiodivDatJour 4: e8029" docOrigin="Biodiversity Data Journal 4" docSource="http://dx.doi.org/10.3897/BDJ.4.e8029" docTitle="Acrostichus medius Tahseen, Ahlawat, Asif &amp; Mustaqim, 2016, sp. n." docType="treatment" docVersion="3" lastPageNumber="8029" masterDocId="FFCAFF8BA820FFA9FFC75054FF9AFF88" masterDocTitle="Description of a new species of Acrostichus Rahm 1928 (Nematoda: Diplogastridae) from India with a note on its position and relationship with the congeners" masterLastPageNumber="8029" masterPageNumber="8029" pageNumber="8029" updateTime="1668126870976" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Description of a new species of Acrostichus Rahm 1928 (Nematoda: Diplogastridae) from India with a note on its position and relationship with the congeners</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tahseen, Qudsia</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ahlawat, Shikha</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Asif, Mohammad</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mustaqim, Malka</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Biodiversity Data Journal</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>8029</mods:start>
<mods:end>8029</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/BDJ.4.e8029</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.4.e8029</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2828-4-8029</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:255948EE5432DFECD7DC036D54F03F21" httpUri="http://treatment.plazi.org/id/255948EE5432DFECD7DC036D54F03F21" lastPageNumber="8029" pageId="0" pageNumber="8029">
<subSubSection pageId="0" pageNumber="8029" type="nomenclature">
<paragraph pageId="0" pageNumber="8029">
<taxonomicName LSID="urn:lsid:zoobank.org:author:E2679573-3F6B-4221-8817-AFB63266528D" class="Chromadorea" family="Diplogastridae" genus="Acrostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrostichus medius" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="species" species="medius">Acrostichus medius</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="8029">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="materials_examined">
<paragraph pageId="0" pageNumber="8029">Materials</paragraph>
<paragraph pageId="0" pageNumber="8029">
<materialsCitation collectionCode="AMU, NOQ" pageId="0" pageNumber="8029" specimenCount="1" typeStatus="Holotype">
Type status:
<typeStatus pageId="0" pageNumber="8029">Holotype</typeStatus>
. Record Level: type: Acrostichusmedius; modified: NOQ; language: English; rightsHolder: Aligarh Muslim University; bibliographicCitation: Tahseen et al.; institutionID: amu.ac.in; collectionID: urn:lsid:zoobank.org:author:E2679573-3F6B-4221-8817-AFB63266528D; institutionCode:
<collectionCode pageId="0" pageNumber="8029">AMU</collectionCode>
; collectionCode:
<collectionCode pageId="0" pageNumber="8029">NOQ</collectionCode>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="description">
<paragraph pageId="0" pageNumber="8029">Description</paragraph>
<paragraph pageId="0" pageNumber="8029">
Hermaphrodite female (Figs 2, 3, 4, 5, 6, 7 and Table 1): Body slender, medium to large-sized; almost straight after fixation, tapering at both extremities. Cuticle with fine transverse striations; longitudinal ridges delicate, striated (Figs 3e, 4a, e, 5b). Punctations faint, dot-like, running along longitudinal ridges (Fig. 4a). Lateral fields demarcated with two conspicuous ridges about 3-5
<normalizedToken originalValue="µm">µm</normalizedToken>
apart. Lip region continuous with adjoining body. Lips amalgamated; labial sensilla slightly raised. Amphidial apertures elliptical, 2.5-3.5
<normalizedToken originalValue="µm">µm</normalizedToken>
wide and situated about 5-7
<normalizedToken originalValue="µm">µm</normalizedToken>
from anterior end of stoma (Fig. 2c). Cheilostom cuticularised, converging anteriorly with six adradial plates, gymnostom anisomorphic with dorsal wall thickened than sub-ventrals. Stegostom anisotopic, anisomorphic. Dorsal metastegostomal wall with a triangular tooth, each sub-ventral wall provided with relatively smaller tooth (Figs 2c, 3a, b). Pharyngeal corpus muscular, swollen 70-88
<normalizedToken originalValue="µm">µm</normalizedToken>
long; metacorpus, rounded to ovoid strongly developed with thickened lumen, more or less oblong in few specimens; isthmus conspicuously differentiated from metacorpus, 33-45
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Basal bulb small, pyriform, glandular in nature, continuous with isthmus, 16-22
<normalizedToken originalValue="µm">µm</normalizedToken>
x 13-17
<normalizedToken originalValue="µm">µm</normalizedToken>
in dimension (Figs 2d, 3d). Nerve ring encircling isthmus at 67-74% of pharyngeal length from anterior end. Secretory-excretory pore placed posterior to nerve ring or at 81-85% of pharyngeal length from anterior end (Figs 2d, 3e). Body at pharyngeal end 3.5-4.4 times labial diam. wide. Cardia 5-7
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Intestine thin-walled, often with a bacterial pouch present in anterior part. Rectum 1.2-1.5 times anal body diam. long, with three rectal glands. Reproductive system didelphic, amphidelphic; ovaries reflexed, not reaching the level of vulva (Fig. 4b, c); anterior ovary on right side and posterior on left side of intestine. Posterior genital branch usually smaller as compared to anterior branch (Fig. 6b, c, d). In several individuals, the posterior branch represented by a post-uterine sac (Fig. 6e). Oocytes arranged in double row in proliferation zone of each ovary. Oviduct a narrow tube leading to a wider ovoid spermatheca (Fig. 4b). The proximal end of spermatheca showing cluster of sperms (sh) developed within the
<normalizedToken originalValue="hermaphrodite’s">hermaphrodite's</normalizedToken>
gonad. Uteri containing 2-4 eggs of 50-56
<normalizedToken originalValue="µm">µm</normalizedToken>
x 29-32
<normalizedToken originalValue="µm">µm</normalizedToken>
in dimension occasionally in early stage of segmentation. Each uterus with a large ovoid to kidney-shaped dorsal bi- or trilocular pouch connected to vagina mostly filled with larger sperms (sm) transferred as a result of copulation (Fig. 7). Vagina 17-21
<normalizedToken originalValue="µm">µm</normalizedToken>
, cuticularised, forming an ovijector with thick lumen occupying about 1/4th of corresponding body diam.; epiptygma present (Figs 4c, 7b). Vulval lips slightly protruding. Distance between vulva-anus 0.7-0.9 times tail length. Tail long filiform with a fine pointed terminus.
</paragraph>
<paragraph pageId="0" pageNumber="8029">
Male (Figs 2b, f, 5 and Table 1): Similar to female in general appearance but shorter in length and strongly curved in posterior region. Testis single, laterally reflexed, reflexed part 45-52
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Vas deferens a long tube with a tapering ejaculatory duct joining with rectum to form cloaca. Spicules strongly built, heavily cuticularised, strongly arcuate in proximal half, 1.3-1.5 times anal body diam. long with elongated hood-like capitula, a distinct neck and distal part deeply bifurcated with fine extensions and an attenuated ventral arm separated from main body of spicule (Figs 2f, 5a, d, e). Gubernaculum stout, heavily built, 70-82% of spicule length, proximally tapering and curved and distal end with slight protuberances (Figs 2f, 5e). Tail in two parts, an anterior short, conoid part and a posterior long, filamentous part. Genital papillae ten pairs with three precloacal, one adcloacal and six postcloacal pairs. Precloacal pairs GP1 and GP2 closely placed, subventral; GP3 lateroventral. GP4 at level of cloaca. Postcloacal pairs GP5 closely posterior to cloaca; GP6 subventral, more or less one cloacal body diameter posterior to cloaca and nearly at level of phasmids. GP10 subdorsal pair placed slightly posterior to the group of subventrals GP7, GP8 and GP9 (Figs 2f, 5b, c). A membranous rudiment indicating bursa, occasionally confined to posterior most genital papillae including three subventral pairs and one dorsally directed pair (Figs 2f, 5c). Copulatory muscles representing 5-6 pairs of broad bands. Phasmids pore-like, about one anal body diam. posterior to anal opening.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="diagnosis">
<paragraph pageId="0" pageNumber="8029">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="8029">
<taxonomicName class="Chromadorea" family="Diplogastridae" genus="Acrostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrostichus medius" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="species" species="medius">Acrostichus medius</taxonomicName>
n. sp. is characterised by female hermaphrodites having a medium-sized body with cuticle bearing faintly striated longitudinal ridges; lateral fields demarcated with two conspicuous ridges; lips amalgamated, labial sensilla small, papilliform; cheilostom with six adradial plates; dorsal tooth moderately-built slightly larger than subventrals; posterior genital branch relatively smaller; dorsal uterine pouch large, multilocular, oval to rounded occasionally filled with sperms and males with robust, heavily cuticularised spicules with hood-like capitula, appearing deeply forked distally with fine extensions and a ventral attenuated arm; gubernaculum 2/3 of spicule length with curved and tapering proximal end and distal end with slight protuberances; bursa almost absent with rudiments confined to posterior four genital pairs.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="etymology">
<paragraph pageId="0" pageNumber="8029">Etymology</paragraph>
<paragraph pageId="0" pageNumber="8029">
The species name
<normalizedToken originalValue="‘medius’">'medius'</normalizedToken>
is a latin word that indicates the intermediate status of the species showing a blend of characters of
<taxonomicName class="Chromadorea" family="Diplogastridae" genus="Acrostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrostichus" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="genus">Acrostichus</taxonomicName>
and
<taxonomicName class="Chromadorea" family="Diplogastridae" genus="Diplogastrellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplogastrellus" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="genus">Diplogastrellus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="distribution">
<paragraph pageId="0" pageNumber="8029">Distribution</paragraph>
<paragraph pageId="0" pageNumber="8029">
Samples containing
<taxonomicName class="Chromadorea" family="Diplogastridae" genus="Acrostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrostichus medius" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="species" species="medius">Acrostichus medius</taxonomicName>
n. sp. were collected from soil rich in organic matter near State Bank of India at Aligarh, Uttar Pradesh, India at geographical coordinates 27°53'35&quot;N, 78°4'27&quot;E.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="ecology">
<paragraph pageId="0" pageNumber="8029">Ecology</paragraph>
<paragraph pageId="0" pageNumber="8029">
<taxonomicName class="Chromadorea" family="Diplogastridae" genus="Acrostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrostichus" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="genus">Acrostichus</taxonomicName>
is a genus with species reported from all the continents except Australia although biogeography of the genus indicates larger distribution in subtropical to temperate regions (Fig. 8). The individuals have been found to inhabit soil, fresh and polluted water and more specifically reported from aquatic mulm or slime flux or sewage; from soil to rotten decaying matter or from moist husk to rotten plants and from dung to frass of beetles. Most species are bacteriophagous but like other diplogastrids, may also feed on protozoa, fungi and nematodes (
<bibRefCitation author="Bento, G." journalOrPublisher="Nature" pageId="0" pageNumber="8029" pagination="494 - 497" title="Co-option of the hormone signalling module dafachronic acid-DAF- 12 in nematode evolution" volume="466" year="2010">Bento et al. 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="conservation">
<paragraph pageId="0" pageNumber="8029">Conservation</paragraph>
<paragraph pageId="0" pageNumber="8029">Due to inhabiting diverse environment types, the species show extensive diversity reflecting phenotypic plasticity. The variations in the shape of buccal cavity which can be shallow and broad, or narrower and deeper, and the variations in size and shape of dorsal tooth are few such examples.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="biology">
<paragraph pageId="0" pageNumber="8029">Biology</paragraph>
<paragraph pageId="0" pageNumber="8029">
<taxonomicName lsidName="A. medius" pageId="0" pageNumber="8029" rank="species" species="medius">A. medius</taxonomicName>
n. sp. shows hermaphrodite females and males in the population. It demonstrates a gradual reduction in posterior genital branch in several specimens up to the extent of a reminiscent post-uterine sac. Such unique feature indicates towards its transitional status in the evolutionary process showing affinities to both the related yet distinct genera
<taxonomicName class="Chromadorea" family="Diplogastridae" genus="Acrostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrostichus" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="genus">Acrostichus</taxonomicName>
and
<taxonomicName class="Chromadorea" family="Diplogastridae" genus="Diplogastrellus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplogastrellus" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="genus">Diplogastrellus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="taxon discussion">
<paragraph pageId="0" pageNumber="8029">Taxon discussion</paragraph>
<paragraph pageId="0" pageNumber="8029">Relationship with closely related species</paragraph>
<paragraph pageId="0" pageNumber="8029">
<taxonomicName class="Chromadorea" family="Diplogastridae" genus="Acrostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrostichus medius" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="species" species="medius">Acrostichus medius</taxonomicName>
n. sp. most closely resembles
<taxonomicName lsidName="A. consobrinus" pageId="0" pageNumber="8029" rank="species" species="consobrinus">A. consobrinus</taxonomicName>
(
<bibRefCitation author="De Man, J. G." journalOrPublisher="Annales de la Societe Royale Zoologique de, Belgique" pageId="0" pageNumber="8029" pagination="47 - 54" title="Description d'une espèce nouvelle du genre Diplogaster Max Schultze: Diplogasterconsobrinus, nov. sp" volume="51" year="1920">De Man 1920</bibRefCitation>
)
<bibRefCitation author="Massey, C. L." journalOrPublisher="Proceedings of the Helminthological Society of Washington" pageId="0" pageNumber="8029" pagination="67 - 75" title="New species of Diplogasteridae (Nematoda) associated with bark beetles in the United States" volume="29" year="1962">Massey 1962</bibRefCitation>
in most morphometric characteristics but differs in having smaller
<normalizedToken originalValue="‘a’">'a'</normalizedToken>
(13.0-19.6 vs 20.5-24.3) and greater 'c' (9.2-15.8 vs 7-8) values in females and smaller
<normalizedToken originalValue="‘a’">'a'</normalizedToken>
value (15.2-19.5 vs 25.7-29.5) in males; labial sensilla papilliform (vs setose); faint (vs conspicuous) adradial plates; stoma narrow tubular (vs wide tubular); dorsal tooth small (vs robust), slightly larger (vs markedly larger) than subventrals; spicules with prominently demarcated dorsal and ventral arms having fine distal extensions (vs arms not demarcated into arms and genital papillae ten pairs (vs nine pairs) in
<taxonomicName lsidName="A. consobrinus" pageId="0" pageNumber="8029" rank="species" species="consobrinus">A. consobrinus</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="8029">
<taxonomicName class="Chromadorea" family="Diplogastridae" genus="Acrostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acrostichus medius" order="Rhabditida" pageId="0" pageNumber="8029" phylum="Nematoda" rank="species" species="medius">Acrostichus medius</taxonomicName>
n. sp. also resembles
<taxonomicName lsidName="A. superbus" pageId="0" pageNumber="8029" rank="species" species="superbus">A. superbus</taxonomicName>
(
<bibRefCitation author="Paesler, F." journalOrPublisher="Kieler Bucht. Kieler Meeresforschungen" pageId="0" pageNumber="8029" pagination="1 - 48" title="Beitrag zur Kenntnis der im Duenger Bucht" volume="1" year="1946">Paesler 1946</bibRefCitation>
)
<bibRefCitation author="Massey, C. L." journalOrPublisher="Proceedings of the Helminthological Society of Washington" pageId="0" pageNumber="8029" pagination="8 - 13" title="The genus Acrostichus Rahm, 1928, synonym Diplogasteritus Paramanov, 1952 (Nematoda)" volume="33" year="1966">Massey 1966</bibRefCitation>
in most morphometric characteristics but differs in having smaller
<normalizedToken originalValue="‘a’">'a'</normalizedToken>
value in females (13.0-19.6 vs 20.0-24.4) and males (15.2-19.5 vs 25.5-30.2); labial sensilla papilliform (vs setose); faint (vs conspicuous) adradial plates; stoma narrow tubular (vs wide tubular); gymnostom (strongly vs weakly) cuticularised; subventral metastegostomal teeth relatively larger (vs smaller); basal bulb continuous with isthmus (vs distinctly demarcated); males with broad, massive (vs slender, arcuate) spicules having prominent dorsal and ventral arms (vs arms not demarcated) and genital papillae ten pairs (vs nine pairs) in
<taxonomicName lsidName="A. superbus" pageId="0" pageNumber="8029" rank="subspecies" species="superbus" subspecies="apud">A. superbus apud</taxonomicName>
<bibRefCitation author="Weingaertner, I." journalOrPublisher="Zoologische Jahrbuecher (Systematik)" pageId="0" pageNumber="8029" pagination="248 - 317" title="Versuch einer Neuordnung der Gattung Diplogaster Schulze 1857 (Nematoda)." volume="83" year="1955">
<normalizedToken originalValue="Weingärtner">Weingaertner</normalizedToken>
1950
</bibRefCitation>
and
<bibRefCitation author="Gagarin, V. G." journalOrPublisher="Zoosystematica Rossica" pageId="0" pageNumber="8029" pagination="57 - 73" title="A review of the genus Diplogasteritus Paramonov, 1952 with description of a new species from Central Russia (Nematoda, Diplogasterida: Diplogasteridae)" volume="11" year="2002">Gagarin 2002</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="8029">
The new species comes close to
<taxonomicName lsidName="A. dendrophilus" pageId="0" pageNumber="8029" rank="species" species="dendrophilus">A. dendrophilus</taxonomicName>
(
<bibRefCitation author="Weingaertner, I." journalOrPublisher="Zoologische Jahrbuecher (Systematik)" pageId="0" pageNumber="8029" pagination="248 - 317" title="Versuch einer Neuordnung der Gattung Diplogaster Schulze 1857 (Nematoda)." volume="83" year="1955">
<normalizedToken originalValue="Weingärtner">Weingaertner</normalizedToken>
1950
</bibRefCitation>
)
<bibRefCitation author="Massey, C. L." journalOrPublisher="Proceedings of the Helminthological Society of Washington" pageId="0" pageNumber="8029" pagination="8 - 13" title="The genus Acrostichus Rahm, 1928, synonym Diplogasteritus Paramanov, 1952 (Nematoda)" volume="33" year="1966">Massey 1966</bibRefCitation>
in most morphometric characteristics but differs in having smaller
<normalizedToken originalValue="‘a’">'a'</normalizedToken>
value (13.0-19.6 vs 23.3-30.7) in males; smaller
<normalizedToken originalValue="‘b’">'b'</normalizedToken>
value (5.7-6.3 vs 6.8-8.6) in females; greater
<normalizedToken originalValue="‘c’">'c'</normalizedToken>
(3.1-3.6 vs 2.0-3.0) and
<normalizedToken originalValue="‘V’">'V'</normalizedToken>
(43.0-46.9 vs 33.7-36.7) values; labial sensilla papilliform (vs setose); males with broad, massive (vs slender, arcuate) spicules; hood-shaped (vs rounded) capitula; spicules having prominent dorsal and ventral arms (vs arms not demarcated) and genital papillae ten pairs (vs eight pairs) in
<taxonomicName lsidName="A. dendrophilus" pageId="0" pageNumber="8029" rank="subspecies" species="dendrophilus" subspecies="apud">A. dendrophilus apud</taxonomicName>
<bibRefCitation author="Gagarin, V. G." journalOrPublisher="Zoosystematica Rossica" pageId="0" pageNumber="8029" pagination="57 - 73" title="A review of the genus Diplogasteritus Paramonov, 1952 with description of a new species from Central Russia (Nematoda, Diplogasterida: Diplogasteridae)" volume="11" year="2002">Gagarin 2002</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="8029">
The new species also resembles
<taxonomicName lsidName="A. lazarevskajae" pageId="0" pageNumber="8029" rank="species" species="lazarevskajae">A. lazarevskajae</taxonomicName>
(
<bibRefCitation author="Lazarevskaja, S. L." journalOrPublisher="Trudy Geologicheskogo Instituta Akademiya Nauk SSSR (Moskva)" pageId="0" pageNumber="8029" pagination="122 - 127" title="Acrostichusminimus n. sp. (Diplogasteroididae) - a new nematode of the timberman beetle. (In Russian)" volume="14" year="1964">Lazarevskaja 1964</bibRefCitation>
)
<bibRefCitation author="Sudhaus, W." journalOrPublisher="Journal of Nematode Morphology and Systematics" pageId="0" pageNumber="8029" pagination="43 - 90" title="A phylogenetic classification of the Diplogastridae (Secernentea, Nematoda)" volume="6" year="2003">
Sudhaus and
<normalizedToken originalValue="Fürst">Fuerst</normalizedToken>
von Lieven 2003
</bibRefCitation>
in most allometric ratios but differs from it in having larger body size (L= 764-867
<normalizedToken originalValue="µm">µm</normalizedToken>
vs 365-430
<normalizedToken originalValue="µm">µm</normalizedToken>
in females and 611-715
<normalizedToken originalValue="µm">µm</normalizedToken>
vs 290-441
<normalizedToken originalValue="µm">µm</normalizedToken>
in males); greater '
<normalizedToken originalValue="ć'">c'</normalizedToken>
value (9.2-15.8 vs 6.6) and large-sized spicules (36-44
<normalizedToken originalValue="µm">µm</normalizedToken>
vs 23-25
<normalizedToken originalValue="µm">µm</normalizedToken>
) in males; spicules massive (vs slender) with elongated hood-like (vs rounded capitula) and distal part deeply bifurcated (vs bearing very fine spines at tip) and presence [vs absence of an attenuated ventral arm separated from main body of spicules in
<taxonomicName lsidName="A. lazarevskajae" pageId="0" pageNumber="8029" rank="subspecies" species="lazarevskajae" subspecies="apud">A. lazarevskajae apud</taxonomicName>
<bibRefCitation author="Lazarevskaja, S. L." journalOrPublisher="Trudy Geologicheskogo Instituta Akademiya Nauk SSSR (Moskva)" pageId="0" pageNumber="8029" pagination="122 - 127" title="Acrostichusminimus n. sp. (Diplogasteroididae) - a new nematode of the timberman beetle. (In Russian)" volume="14" year="1964">Lazarevskaja 1964</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="8029" type="notes">
<paragraph pageId="0" pageNumber="8029">Notes</paragraph>
<paragraph pageId="0" pageNumber="8029">Most of the species of the genus have been described on very flimsy characteristics hence many of them are likely to be synonymous. We tried to have a comparative assessment of the morphological characteristics of most species. Of the total nominal species, few could not be included largely due to insufficient descriptions available.</paragraph>
</subSubSection>
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Latreille, 1804
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<taxonomicName id="6AF04BB0B0120731708296C8A580F900" ID-CoL="3R86" authority="Latreille, 1804: 134" authorityName="Latreille" authorityPageNumber="134" authorityYear="1804" box="[151,455,1693,1715]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EC21B0120731708296C8A4B2F900" box="[151,245,1693,1715]" italics="true" pageId="3" pageNumber="30">Clubiona</emphasis>
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Clerck, 1757
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<emphasis id="9F84EC21B012073170829751A509F8AD" bold="true" box="[151,334,1796,1822]" italics="true" pageId="3" pageNumber="30">Clubiona blesti</emphasis>
<bibRefCitation id="C9614DC2B012073171409756A5BEF8AE" author="Forster" box="[341,505,1795,1821]" pageId="3" pageNumber="30" refString="Forster, R. R. (1979) The spiders of New Zealand. Part V. Cycloctenidae, Gnaphosidae, Clubionidae. Otago Museum Bulletin, 5, 8 - 95." type="journal article" year="1979">Forster, 1979</bibRefCitation>
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<paragraph id="AD4F3033B012073170829772A5DFF88C" blockId="3.[151,505,1795,1855]" box="[151,408,1831,1855]" pageId="3" pageNumber="30">Figs 1–3, 4a, 5–6, 7c–f</paragraph>
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<paragraph id="AD4F3033B01207317082973AA771F837" blockId="3.[151,822,1902,1925]" box="[151,822,1902,1925]" pageId="3" pageNumber="30">
<treatmentCitationGroup id="8DE0171DB01207317082973AA771F837" box="[151,822,1902,1925]" pageId="3" pageNumber="30">
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<emphasis id="9F84EC21B01207317082973AA575F836" box="[151,306,1903,1925]" italics="true" pageId="3" pageNumber="30">Clubiona blesti</emphasis>
<treatmentCitation id="2C511622B0120731712C973AA5AFF837" author="Forster" box="[313,488,1903,1924]" page="78" pageId="3" pageNumber="30" year="1979">
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: 78
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, fig. 309 (description of female).
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material.
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<collectionCode id="CBE1A8F6B0120731714497E7A5D3F879" box="[337,404,1970,1994]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="3" pageNumber="30">NEW</collectionCode>
<collectionCode id="CBE1A8F6B0120731718B97E4A66CF879" box="[414,555,1969,1994]" pageId="3" pageNumber="30">ZEALAND</collectionCode>
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: Fiordland:
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female (not examined), Lyttles Flat, in wet tussock in bog,
<date id="D94E16F3B012073170DA9780A5DEF85E" box="[207,409,2005,2029]" pageId="3" pageNumber="30" value="1975-02-17">17 February 1975</date>
, A.D. Blest (Otago Museum).
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<emphasis id="9F84EC21B0140737708297E0A554F878" bold="true" box="[151,275,1973,1996]" pageId="5" pageNumber="32">FIGURE 2.</emphasis>
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<emphasis id="9F84EC21B0140737710C97E3A5F3F87F" box="[281,436,1974,1996]" italics="true" pageId="5" pageNumber="32">Clubiona blesti</emphasis>
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. a male habitus (BLCO6), b female habitus (BLCO2).
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<emphasis id="9F84EC21B0170734708296ECA550F97D" bold="true" box="[151,279,1721,1744]" pageId="6" pageNumber="33">FIGURE 3.</emphasis>
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<emphasis id="9F84EC21B0170734713796ECA585F97C" box="[290,450,1721,1743]" italics="true" pageId="6" pageNumber="33">Clubiona blesti</emphasis>
</taxonomicName>
, left pedipalp. c—conductor, e—embolus, rta—retrolateral tibial apophysis, sd—sperm duct, t—tegulum, ta—tegular apophysis. Scale bars = 0.5 mm.
</paragraph>
</caption>
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Other material examined.
<collectingCountry id="D5E770A3B0170734721E9776A6A1F888" box="[523,742,1826,1851]" name="New Zealand" pageId="6" pageNumber="33">
<collectionCode id="CBE1A8F6B0170734721E9776A609F888" box="[523,590,1827,1851]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="33">NEW</collectionCode>
<collectionCode id="CBE1A8F6B0170734724D9777A6A1F888" box="[600,742,1826,1851]" pageId="6" pageNumber="33">ZEALAND</collectionCode>
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: Otago Lakes:
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Temple Peak
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<geoCoordinate id="C8C456F4B017073474C09777A136F889" box="[1237,1393,1826,1850]" direction="east" orientation="longitude" pageId="6" pageNumber="33" precision="1" value="168.4634">168º27.804’E</geoCoordinate>
,
<date id="D94E16F3B017073475959777A578F8ED" pageId="6" pageNumber="33" value="2003-02-27">27 February 2003</date>
, C.J. Vink,
<specimenCount id="BBF6FBBAB017073471D29713A66EF8ED" box="[455,553,1862,1886]" pageId="6" pageNumber="33" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B0170734722F9713A6CFF8ED" box="[570,648,1862,1886]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="6" pageNumber="33">LUNZ</collectionCode>
00012640).
<emphasis id="9F84EC21B0170734731A9713A7A7F8EC" bold="true" box="[783,992,1862,1887]" pageId="6" pageNumber="33">Mid Canterbury:</emphasis>
Ellangowan Scenic Reserve
<geoCoordinate id="C8C456F4B0170734753B9713A1DBF8ED" box="[1326,1436,1862,1886]" direction="south" orientation="latitude" pageId="6" pageNumber="33" precision="92" value="-43.798332">43º47.9’S</geoCoordinate>
<geoCoordinate id="C8C456F4B01707347082973FA55FF831" box="[151,280,1898,1922]" direction="east" orientation="longitude" pageId="6" pageNumber="33" precision="92" value="173.02667">173º01.6’E</geoCoordinate>
,
<date id="D94E16F3B01707347131973FA5BAF831" box="[292,509,1898,1922]" pageId="6" pageNumber="33" value="2006-12-12">12 December 2006</date>
, J. Malumbres-Olarte,
<specimenCount id="BBF6FBBAB01707347317973FA723F831" box="[770,868,1898,1922]" pageId="6" pageNumber="33" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B01707347361973FA784F831" box="[884,963,1898,1922]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="6" pageNumber="33">LUNZ</collectionCode>
00012641); same data except
<date id="D94E16F3B01707347506973FA495F815" pageId="6" pageNumber="33" value="2007-02-20">20 February 2007</date>
,
<specimenCount id="BBF6FBBAB017073470C897DBA578F815" box="[221,319,1934,1958]" pageId="6" pageNumber="33" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B0170734715B97DBA5DBF815" box="[334,412,1934,1958]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="6" pageNumber="33">LUNZ</collectionCode>
00012642).
<emphasis id="9F84EC21B0170734723797DBA690F814" bold="true" box="[546,727,1934,1959]" pageId="6" pageNumber="33">Central Otago:</emphasis>
Te Papanui Conservation Park
<geoCoordinate id="C8C456F4B0170734742E97DBA0EEF815" box="[1083,1193,1934,1958]" direction="south" orientation="latitude" pageId="6" pageNumber="33" precision="92" value="-45.68">45º40.8’S</geoCoordinate>
<geoCoordinate id="C8C456F4B017073474A497DBA175F815" box="[1201,1330,1934,1958]" direction="east" orientation="longitude" pageId="6" pageNumber="33" precision="92" value="169.755">169º45.3’E</geoCoordinate>
,
<date id="D94E16F3B0170734752B97DBA495F879" pageId="6" pageNumber="33" value="2008-03-02">2 March 2008</date>
, J. Malumbres-Olarte,
<specimenCount id="BBF6FBBAB017073471C597E7A65EF879" box="[464,537,1970,1994]" pageId="6" pageNumber="33" type="male">1 male</specimenCount>
(
<collectionCode id="CBE1A8F6B0170734723297E7A632F879" box="[551,629,1970,1994]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="6" pageNumber="33">LUNZ</collectionCode>
00012648); same data,
<specimenCount id="BBF6FBBAB0170734736397E7A791F879" box="[886,982,1970,1994]" pageId="6" pageNumber="33" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B017073473F197E7A075F879" box="[996,1074,1970,1994]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="6" pageNumber="33">LUNZ</collectionCode>
00012646); same data except
<date id="D94E16F3B017073A756F97E7A508FF1C" lastPageId="8" lastPageNumber="35" pageId="6" pageNumber="33" value="2007-12-21">21 December 2007</date>
,
<specimenCount id="BBF6FBBAB019073A714990C2A587FF1C" box="[348,448,151,175]" pageId="8" pageNumber="35" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B019073A71C790C2A667FF1C" box="[466,544,151,175]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="8" pageNumber="35">LUNZ</collectionCode>
00012649); same data except
<geoCoordinate id="C8C456F4B019073A736290C2A7A2FF1C" box="[887,997,151,175]" direction="south" orientation="latitude" pageId="8" pageNumber="35" precision="92" value="-45.665">45º39.9’S</geoCoordinate>
<geoCoordinate id="C8C456F4B019073A73FA90C2A01CFF1C" box="[1007,1115,151,175]" direction="east" orientation="longitude" pageId="8" pageNumber="35" precision="925" value="169.78334">169º47’E</geoCoordinate>
,
<date id="D94E16F3B019073A747C90C2A16DFF1C" box="[1129,1322,151,176]" pageId="8" pageNumber="35" value="2008-01-18">18 January 2008</date>
,
<specimenCount id="BBF6FBBAB019073A752D90C2A1DBFF1C" box="[1336,1436,151,175]" pageId="8" pageNumber="35" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B019073A708B90E9A4ABFF67" box="[158,236,188,212]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="8" pageNumber="35">LUNZ</collectionCode>
00012643); same data except
<date id="D94E16F3B019073A725390E9A751FF67" box="[582,790,188,212]" pageId="8" pageNumber="35" value="2008-02-20">20 February 2008</date>
,
<specimenCount id="BBF6FBBAB019073A733190E9A7CEFF67" box="[804,905,188,212]" pageId="8" pageNumber="35" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B019073A738E90E9A7AEFF67" box="[923,1001,188,212]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="8" pageNumber="35">LUNZ</collectionCode>
00012644); same data except
<geoCoordinate id="C8C456F4B019073A755690E9A1DAFF67" box="[1347,1437,188,212]" direction="south" orientation="latitude" pageId="8" pageNumber="35" precision="925" value="-45.666668">45º40’S</geoCoordinate>
<geoCoordinate id="C8C456F4B019073A7082908AA550FF44" box="[151,279,223,247]" direction="east" orientation="longitude" pageId="8" pageNumber="35" precision="92" value="169.77667">169º46.6’E</geoCoordinate>
,
<date id="D94E16F3B019073A7139908AA599FF44" box="[300,478,223,247]" pageId="8" pageNumber="35" value="2008-03-04">4 March 2008</date>
,
<specimenCount id="BBF6FBBAB019073A71E7908AA61BFF44" box="[498,604,223,247]" pageId="8" pageNumber="35" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B019073A7261908AA685FF44" box="[628,706,223,247]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="8" pageNumber="35">LUNZ</collectionCode>
00012645); same data except
<date id="D94E16F3B019073A7426908AA159FF44" box="[1075,1310,223,247]" pageId="8" pageNumber="35" value="2008-12-14">14 December 2008</date>
,
<specimenCount id="BBF6FBBAB019073A7527908AA1DBFF44" box="[1330,1436,223,247]" pageId="8" pageNumber="35" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B019073A708B9151A4ABFEAF" box="[158,236,260,284]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="8" pageNumber="35">LUNZ</collectionCode>
00012647).
<emphasis id="9F84EC21B019073A71639156A5A1FEAF" bold="true" box="[374,486,259,284]" pageId="8" pageNumber="35">Dunedin:</emphasis>
Swampy Spur
<geoCoordinate id="C8C456F4B019073A728B9151A75BFEAF" box="[670,796,260,284]" direction="south" orientation="latitude" pageId="8" pageNumber="35" precision="9" value="-45.801334">45°48.08’S</geoCoordinate>
<geoCoordinate id="C8C456F4B019073A733D9151A7FEFEAF" box="[808,953,260,284]" direction="east" orientation="longitude" pageId="8" pageNumber="35" precision="9" value="170.49">170°29.40’E</geoCoordinate>
,
<date id="D94E16F3B019073A73DD9151A0CAFEAF" box="[968,1165,260,284]" pageId="8" pageNumber="35" value="2011-10-20">20 October 2011</date>
, M. Wakelin,
<specimenCount id="BBF6FBBAB019073A75229151A1DBFEAF" box="[1335,1436,260,284]" pageId="8" pageNumber="35" type="female">1 female</specimenCount>
(
<collectionCode id="CBE1A8F6B019073A708B9172A4ABFE8C" box="[158,236,295,319]" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="8" pageNumber="35">LUNZ</collectionCode>
00012709).
</paragraph>
<caption id="F98F60BBB0160735708297E0A00FF85C" httpUri="https://zenodo.org/record/212925/files/figure.png" pageId="7" pageNumber="34" targetBox="[359,1225,183,1972]" targetPageId="7">
<paragraph id="AD4F3033B0160735708297E0A00FF85C" blockId="7.[151,1435,1973,2031]" pageId="7" pageNumber="34">
<emphasis id="9F84EC21B0160735708297E0A555F878" bold="true" box="[151,274,1973,1996]" pageId="7" pageNumber="34">FIGURE 4.</emphasis>
Left pedipalps, retrolateral view. a
<taxonomicName id="6AF04BB0B0160735726397E3A756F87F" box="[630,785,1974,1996]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="blesti">
<emphasis id="9F84EC21B0160735726397E3A756F87F" box="[630,785,1974,1996]" italics="true" pageId="7" pageNumber="34">Clubiona blesti</emphasis>
</taxonomicName>
; b
<taxonomicName id="6AF04BB0B0160735732597E3A040F87F" box="[816,1031,1974,1996]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="cambridgei">
<emphasis id="9F84EC21B0160735732597E3A040F87F" box="[816,1031,1974,1996]" italics="true" pageId="7" pageNumber="34">Clubiona cambridgei</emphasis>
</taxonomicName>
; Travis Wetland (CMMC2); c
<taxonomicName id="6AF04BB0B0160735752897E3A5F0F85C" authority="Lake Wairarapa" authorityName="Lake Wairarapa" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="cambridgei">
<emphasis id="9F84EC21B0160735752897E3A54DF85C" italics="true" pageId="7" pageNumber="34">Clubiona cambridgei</emphasis>
, Lake Wairarapa
</taxonomicName>
(CMWA2); d
<taxonomicName id="6AF04BB0B0160735725F978CA789F85C" authority="Lake Wairarapa" authorityName="Lake Wairarapa" box="[586,974,2009,2031]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="cambridgei">
<emphasis id="9F84EC21B0160735725F978CA766F85C" box="[586,801,2009,2031]" italics="true" pageId="7" pageNumber="34">Clubiona cambridgei</emphasis>
, Lake Wairarapa
</taxonomicName>
(CMWA1).
</paragraph>
</caption>
<paragraph id="AD4F3033B019073A70D2911EA6DEFE34" blockId="8.[151,1437,151,968]" pageId="8" pageNumber="35">
<emphasis id="9F84EC21B019073A70D2911EA767FED7" bold="true" box="[199,800,331,356]" pageId="8" pageNumber="35">
Other locality records.
<collectingCountry id="D5E770A3B019073A71CA9119A6FFFED7" box="[479,696,331,356]" name="New Zealand" pageId="8" pageNumber="35">
<collectionCode id="CBE1A8F6B019073A71CA9119A665FED7" box="[479,546,332,356]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="8" pageNumber="35">NEW</collectionCode>
<collectionCode id="CBE1A8F6B019073A723E911EA6FFFED7" box="[555,696,331,356]" pageId="8" pageNumber="35">ZEALAND</collectionCode>
</collectingCountry>
: Nelson:
</emphasis>
Lake Sylvester, Cobb,
<quantity id="6A089DD6B019073A743E9119A0C3FED7" box="[1067,1156,332,356]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="35" unit="m" value="1300.0">1300 m</quantity>
, tussock,
<date id="D94E16F3B019073A74E09119A1DFFED7" box="[1269,1432,332,356]" pageId="8" pageNumber="35" value="1969-04-30">30 April 1969</date>
, J.S. Dugdale (Otago Museum, not examined).
</paragraph>
</subSubSection>
<subSubSection id="E5EA63B8B019073A70D291C6A00AFDEC" pageId="8" pageNumber="35" type="diagnosis">
<paragraph id="AD4F3033B019073A70D291C6A00AFDEC" blockId="8.[151,1437,151,968]" pageId="8" pageNumber="35">
<emphasis id="9F84EC21B019073A70D291C6A506FE1F" bold="true" box="[199,321,403,428]" pageId="8" pageNumber="35">Diagnosis.</emphasis>
<taxonomicName id="6AF04BB0B019073A715291C1A5B4FE1F" box="[327,499,404,428]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="blesti">
<emphasis id="9F84EC21B019073A715291C1A5B4FE1F" box="[327,499,404,428]" italics="true" pageId="8" pageNumber="35">Clubiona blesti</emphasis>
</taxonomicName>
is distinguished from other
<collectingCountry id="D5E770A3B019073A733E91C0A786FE1F" box="[811,961,404,428]" name="New Zealand" pageId="8" pageNumber="35">New Zealand</collectingCountry>
<taxonomicName id="6AF04BB0B019073A73DD91C1A077FE1F" box="[968,1072,404,428]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EC21B019073A73DD91C1A077FE1F" box="[968,1072,404,428]" italics="true" pageId="8" pageNumber="35">Clubiona</emphasis>
</taxonomicName>
species by an abdominal pattern that consists of only a medium band that extends along the entire length of the dorsal surface (
<figureCitation id="35CB2CB6B019073A74AD91E2A0B9FE7C" box="[1208,1278,439,463]" captionStart="FIGURE 2" captionStartId="5.[151,250,1973,1996]" captionTargetBox="[262,1332,157,1949]" captionTargetId="figure@5.[262,1333,157,1949]" captionTargetPageId="5" captionText="FIGURE 2. Clubiona blesti. a male habitus (BLCO 6), b female habitus (BLCO 2)." httpUri="https://zenodo.org/record/212923/files/figure.png" pageId="8" pageNumber="35">Fig. 2</figureCitation>
). This band is either absent, partial or accompanied by lateral stripes in other
<collectingCountry id="D5E770A3B019073A73709188A7BAFE47" box="[869,1021,476,500]" name="New Zealand" pageId="8" pageNumber="35">New Zealand</collectingCountry>
<taxonomicName id="6AF04BB0B019073A74109189A02AFE47" box="[1029,1133,476,500]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EC21B019073A74109189A02AFE47" box="[1029,1133,476,500]" italics="true" pageId="8" pageNumber="35">Clubiona</emphasis>
</taxonomicName>
species (see
<bibRefCitation id="C9614DC2B019073A75109188A1DFFE47" author="Forster" box="[1285,1432,476,500]" pageId="8" pageNumber="35" refString="Forster, R. R. (1979) The spiders of New Zealand. Part V. Cycloctenidae, Gnaphosidae, Clubionidae. Otago Museum Bulletin, 5, 8 - 95." type="journal article" year="1979">Forster 1979</bibRefCitation>
: figs 286-296). Males can be identified by the apically notched retrolateral tibial apophysis, which is broader than in
<taxonomicName id="6AF04BB0B019073A70829270A57FFD8F" box="[151,312,548,572]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="cambridgei">
<emphasis id="9F84EC21B019073A70829270A57FFD8F" box="[151,312,548,572]" italics="true" pageId="8" pageNumber="35">C. cambridgei</emphasis>
</taxonomicName>
(
<figureCitation id="35CB2CB6B019073A71529271A5CDFD8F" box="[327,394,548,572]" captionStart="FIGURE 4" captionStartId="7.[151,250,1973,1996]" captionTargetBox="[359,1225,183,1972]" captionTargetId="figure@7.[357,1230,169,1976]" captionTargetPageId="7" captionText="FIGURE 4. Left pedipalps, retrolateral view. a Clubiona blesti; b Clubiona cambridgei; Travis Wetland (CMMC 2); c Clubiona cambridgei, Lake Wairarapa (CMWA 2); d Clubiona cambridgei, Lake Wairarapa (CMWA 1)." httpUri="https://zenodo.org/record/212925/files/figure.png" pageId="8" pageNumber="35">Fig. 4</figureCitation>
a-d and see
<bibRefCitation id="C9614DC2B019073A72059270A6E4FD8F" author="Forster" box="[528,675,548,572]" pageId="8" pageNumber="35" refString="Forster, R. R. (1979) The spiders of New Zealand. Part V. Cycloctenidae, Gnaphosidae, Clubionidae. Otago Museum Bulletin, 5, 8 - 95." type="journal article" year="1979">Forster 1979</bibRefCitation>
: figs 299–300). Females can be recognised by the oval shape of the copulatory opening, which is constricted somewhat at the anterior end (
<figureCitation id="35CB2CB6B019073A73AB9212A041FDEC" box="[958,1030,583,607]" captionStart="FIGURE 5" captionStartId="8.[151,250,1970,1993]" captionTargetBox="[330,1259,1051,1969]" captionTargetId="figure@8.[320,1266,1041,1975]" captionTargetPageId="8" captionText="FIGURE 5. Clubiona blesti, epigynum, ventral, not cleared. cd—copulatory duct, co—copulatory opening, ea—epigynal atrium. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/212926/files/figure.png" pageId="8" pageNumber="35">Figs 5</figureCitation>
,
<figureCitation id="35CB2CB6B019073A74079212A065FDEC" box="[1042,1058,583,607]" captionStart="FIGURE 6" captionStartId="9.[151,250,1666,1689]" captionTargetBox="[329,1257,664,1666]" captionTargetId="figure@9.[329,1257,661,1670]" captionTargetPageId="9" captionText="FIGURE 6. Clubiona blesti, epigynum, dorsal view, cleared. cd—copulatory duct, fd—fertilisation duct, s—spermatheca. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/212927/files/figure.png" pageId="8" pageNumber="35">6</figureCitation>
,
<figureCitation id="35CB2CB6B019073A743A9212A078FDEC" box="[1071,1087,583,607]" captionStart="FIGURE 7" captionStartId="10.[151,250,1719,1742]" captionTargetBox="[151,1436,155,1700]" captionTargetId="figure@10.[151,1436,155,1700]" captionTargetPageId="10" captionText="FIGURE 7. Epigyna, ventral view. Clubiona cambridgei; a Lake Wairarapa, WA (CMWA 3); b Travis Wetland, MC (CMMC 3). Clubiona blesti; c Ellangowan Scenic Res., MC (BLMC 1); d Ellangowan Scenic Reserve, MC (BLMC 2); e Temple Peak, OL (BLOL 1); f Te Papanui Conservation Park, CO (BLCO 7). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/212928/files/figure.png" pageId="8" pageNumber="35">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E5EA63B8B019073B70D2923EA6CAF94F" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35" type="description">
<paragraph id="AD4F3033B019073A70D2923EA736FD5C" blockId="8.[151,1437,151,968]" pageId="8" pageNumber="35">
<emphasis id="9F84EC21B019073A70D2923EA639FD37" bold="true" box="[199,638,619,644]" pageId="8" pageNumber="35">Description. Male (LUNZ00012648):</emphasis>
Carapace length 3.2, width 2.2, height 1.7; abdomen length 3.7, width 1.8; sternum length 1.7, width 1. AME 0.17, ALE 0.19, PME 0.13, PLE 0.15, AME-AME 0.09, AME-ALE 0.2, ALE-ALE 0.87, PME-PME 0.46, PME-PLE 0.33. Leg I 8.3 (2, 1.5, 2.3, 1.7, 0.8); leg II 8.2 (2.1, 1.5, 2.1, 1.7, 0.8); leg III 6.2(1.7, 1, 1.3, 1.6, 0.6), leg IV 8.7 (2.2, 1.3, 1.9, 2.5, 0.8).
</paragraph>
<paragraph id="AD4F3033B019073A70D292A9A057FCA7" blockId="8.[151,1437,151,968]" box="[199,1040,764,788]" pageId="8" pageNumber="35">Chelicerae with five promarginal teeth in a row and no retromarginal teeth.</paragraph>
<paragraph id="AD4F3033B019073A70D2934AA4B6FCCC" blockId="8.[151,1437,151,968]" pageId="8" pageNumber="35">
Pedipalp as in diagnosis. Embolus arising from tegular apophysis, bent 90 degrees ventrally, extending almost to tegulum. Sperm duct with horizontal U-turn visible through tegulum. Tegular apophysis with nob-like projection (
<figureCitation id="35CB2CB6B019073A708A9332A4A3FCCC" box="[159,228,871,895]" captionStart="FIGURE 3" captionStartId="6.[151,250,1721,1744]" captionTargetBox="[317,1272,182,1682]" captionTargetId="figure@6.[313,1273,161,1700]" captionTargetPageId="6" captionText="FIGURE 3. Clubiona blesti, left pedipalp. c—conductor, e—embolus, rta—retrolateral tibial apophysis, sd—sperm duct, t—tegulum, ta—tegular apophysis. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/212924/files/figure.png" pageId="8" pageNumber="35">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="AD4F3033B019073A70D293D9A046FC74" blockId="8.[151,1437,151,968]" pageId="8" pageNumber="35">
Colour in ethanol (
<figureCitation id="35CB2CB6B019073A71B493D9A5AFFC17" box="[417,488,908,932]" captionStart="FIGURE 2" captionStartId="5.[151,250,1973,1996]" captionTargetBox="[262,1332,157,1949]" captionTargetId="figure@5.[262,1333,157,1949]" captionTargetPageId="5" captionText="FIGURE 2. Clubiona blesti. a male habitus (BLCO 6), b female habitus (BLCO 2)." httpUri="https://zenodo.org/record/212923/files/figure.png" pageId="8" pageNumber="35">Fig. 2</figureCitation>
a). Carapace and sternum yellowish-brown, darker in cephalic area. Abdomen pale grey-brown with dorsal dark-brown medium folium. Legs pale yellow-brown.
</paragraph>
<caption id="F98F60BBB019073A708297E7A5F5F858" httpUri="https://zenodo.org/record/212926/files/figure.png" pageId="8" pageNumber="35" targetBox="[330,1259,1051,1969]" targetPageId="8">
<paragraph id="AD4F3033B019073A708297E7A5F5F858" blockId="8.[151,1436,1970,2027]" pageId="8" pageNumber="35">
<emphasis id="9F84EC21B019073A708297E7A550F87B" bold="true" box="[151,279,1970,1993]" pageId="8" pageNumber="35">FIGURE 5.</emphasis>
<taxonomicName id="6AF04BB0B019073A713797E7A585F87B" box="[290,450,1970,1992]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="blesti">
<emphasis id="9F84EC21B019073A713797E7A585F87B" box="[290,450,1970,1992]" italics="true" pageId="8" pageNumber="35">Clubiona blesti</emphasis>
</taxonomicName>
, epigynum, ventral, not cleared. cd—copulatory duct, co—copulatory opening, ea—epigynal atrium. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="AD4F3033B018073B70D290CDA691FE8C" blockId="9.[151,1436,151,607]" pageId="9" pageNumber="36">
<emphasis id="9F84EC21B018073B70D290CDA5B8FF03" bold="true" box="[199,511,151,176]" pageId="9" pageNumber="36">Female (LUNZ00012640):</emphasis>
Carapace length 3.9, width 2.9, height 1.8; abdomen length 5.8, width 3; sternum length 2.3, width 1.3. AME 0.2, ALE 0.23, PME 0.13, PLE 0.15, AME-AME 0.16, AME-ALE 0.25, ALE-ALE 1.09, PME-PME 0.63, PME-PLE 0.47. Leg I 9.4 (2.5, 1.7, 2.5, 1.8, 0.9); leg II 9.4 (9.4, 2.5, 1.7, 2.5, 1.8, 0.9); leg III 7.3 (2.1, 1.3, 1.6, 1.7, 0.6), leg IV 10.8 (2.9, 1.5, 2.6, 3, 0.8, 10.8). Size range: carapace length 2.2–4.8 (mean 3.5, n=10), body length 6.0–10.2 (mean 8.1, n=10).
</paragraph>
<paragraph id="AD4F3033B018073B70D29119A5F4FE34" blockId="9.[151,1436,151,607]" pageId="9" pageNumber="36">Five promarginal cheliceral teeth, with the second one the largest and the distal one very small, and two or three retromarginal teeth.</paragraph>
<paragraph id="AD4F3033B018073B70D291C1A515FDEC" blockId="9.[151,1436,151,607]" pageId="9" pageNumber="36">
Copulatory opening as in diagnosis. Epigynum strongly sclerotised and laterally flat, with rugose ventral surface; posterior margin with a pair of lobes in all examined specimens except for specimen LUNZ00012642, from Central Canterbury, which had a flat margin. Single copulatory opening with slight septum at posterior edge that leads to anteriorly visible copulatory ducts. Copulatory ducts bent vertically and directed ventrally, and connected to spermathecae. Spermathecae separated into two parts with fertilisation ducts arising from ventral side (
<figureCitation id="35CB2CB6B018073B708A9212A4A0FDEC" box="[159,231,583,607]" captionStart="FIGURE 5" captionStartId="8.[151,250,1970,1993]" captionTargetBox="[330,1259,1051,1969]" captionTargetId="figure@8.[320,1266,1041,1975]" captionTargetPageId="8" captionText="FIGURE 5. Clubiona blesti, epigynum, ventral, not cleared. cd—copulatory duct, co—copulatory opening, ea—epigynal atrium. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/212926/files/figure.png" pageId="9" pageNumber="36">Figs 5</figureCitation>
,
<figureCitation id="35CB2CB6B018073B70E69212A544FDEC" box="[243,259,583,607]" captionStart="FIGURE 6" captionStartId="9.[151,250,1666,1689]" captionTargetBox="[329,1257,664,1666]" captionTargetId="figure@9.[329,1257,661,1670]" captionTargetPageId="9" captionText="FIGURE 6. Clubiona blesti, epigynum, dorsal view, cleared. cd—copulatory duct, fd—fertilisation duct, s—spermatheca. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/212927/files/figure.png" pageId="9" pageNumber="36">6</figureCitation>
,
<figureCitation id="35CB2CB6B018073B711A9212A55AFDEC" box="[271,285,583,607]" captionStart="FIGURE 7" captionStartId="10.[151,250,1719,1742]" captionTargetBox="[151,1436,155,1700]" captionTargetId="figure@10.[151,1436,155,1700]" captionTargetPageId="10" captionText="FIGURE 7. Epigyna, ventral view. Clubiona cambridgei; a Lake Wairarapa, WA (CMWA 3); b Travis Wetland, MC (CMMC 3). Clubiona blesti; c Ellangowan Scenic Res., MC (BLMC 1); d Ellangowan Scenic Reserve, MC (BLMC 2); e Temple Peak, OL (BLOL 1); f Te Papanui Conservation Park, CO (BLCO 7). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/212928/files/figure.png" pageId="9" pageNumber="36">7</figureCitation>
c–f).
</paragraph>
<caption id="F98F60BBB018073B708296D7A523F90B" httpUri="https://zenodo.org/record/212927/files/figure.png" pageId="9" pageNumber="36" targetBox="[329,1257,664,1666]" targetPageId="9">
<paragraph id="AD4F3033B018073B708296D7A523F90B" blockId="9.[151,1437,1666,1720]" pageId="9" pageNumber="36">
<emphasis id="9F84EC21B018073B708296D7A551F92B" bold="true" box="[151,278,1666,1689]" pageId="9" pageNumber="36">FIGURE 6.</emphasis>
<taxonomicName id="6AF04BB0B018073B713596D6A587F92A" box="[288,448,1667,1689]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="blesti">
<emphasis id="9F84EC21B018073B713596D6A587F92A" box="[288,448,1667,1689]" italics="true" pageId="9" pageNumber="36">Clubiona blesti</emphasis>
</taxonomicName>
, epigynum, dorsal view, cleared. cd—copulatory duct, fd—fertilisation duct, s—spermatheca. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="AD4F3033B018073B70D296B1A6CAF94F" blockId="9.[151,1436,1764,1968]" box="[199,653,1764,1789]" pageId="9" pageNumber="36">
Colour in ethanol (
<figureCitation id="35CB2CB6B018073B718F96B1A599F94F" box="[410,478,1764,1788]" captionStart="FIGURE 2" captionStartId="5.[151,250,1973,1996]" captionTargetBox="[262,1332,157,1949]" captionTargetId="figure@5.[262,1333,157,1949]" captionTargetPageId="5" captionText="FIGURE 2. Clubiona blesti. a male habitus (BLCO 6), b female habitus (BLCO 2)." httpUri="https://zenodo.org/record/212923/files/figure.png" pageId="9" pageNumber="36">Fig. 2</figureCitation>
b). As for male.
</paragraph>
</subSubSection>
<subSubSection id="E5EA63B8B018073B70D2975DA4A8F8DB" pageId="9" pageNumber="36" type="biology_ecology">
<paragraph id="AD4F3033B018073B70D2975DA4A8F8DB" blockId="9.[151,1436,1764,1968]" pageId="9" pageNumber="36">
<emphasis id="9F84EC21B018073B70D2975DA560F892" bold="true" box="[199,295,1800,1825]" pageId="9" pageNumber="36">Biology.</emphasis>
<bibRefCitation id="C9614DC2B018073B7125975CA593F893" author="Forster" box="[304,468,1800,1824]" pageId="9" pageNumber="36" refString="Forster, R. R. (1979) The spiders of New Zealand. Part V. Cycloctenidae, Gnaphosidae, Clubionidae. Otago Museum Bulletin, 5, 8 - 95." type="journal article" year="1979">Forster (1979)</bibRefCitation>
suggested that
<taxonomicName id="6AF04BB0B018073B729F975CA6ADF893" box="[650,746,1800,1824]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="blesti">
<emphasis id="9F84EC21B018073B729F975CA6ADF893" box="[650,746,1800,1824]" italics="true" pageId="9" pageNumber="36">C. blesti</emphasis>
</taxonomicName>
is an active hunter associated with tussocks. The examined specimens were collected from the leaves and bases of
<taxonomicName id="6AF04BB0B018073B73129778A0CAF8F7" authority="(Raoul) Zotov" authorityName="(Raoul) Zotov" box="[775,1165,1836,1861]" class="Liliopsida" family="Poaceae" genus="Chionochloa" kingdom="Plantae" order="Poales" pageId="9" pageNumber="36" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis id="9F84EC21B018073B73129778A7A3F8F6" box="[775,996,1837,1861]" italics="true" pageId="9" pageNumber="36">Chionochloa rigida</emphasis>
(Raoul) Zotov
</taxonomicName>
, an alpine tussock grass species.
</paragraph>
</subSubSection>
<subSubSection id="E5EA63B8B018073B70D29721A516F803" pageId="9" pageNumber="36" type="distribution">
<paragraph id="AD4F3033B018073B70D29721A516F803" blockId="9.[151,1436,1764,1968]" pageId="9" pageNumber="36">
<emphasis id="9F84EC21B018073B70D29721A527F83E" bold="true" box="[199,352,1908,1933]" pageId="9" pageNumber="36">Distribution.</emphasis>
<collectingCountry id="D5E770A3B018073B717D9720A647F83F" box="[360,512,1908,1932]" name="New Zealand" pageId="9" pageNumber="36">New Zealand</collectingCountry>
South Island (Fiordland, Dunedin, Central Otago, Otago Lakes, Mid Canterbury, Nelson) (Fig. 1).
</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="9F84EAD4FFB27E13FEA7FE35FDD2FE34" bold="true" box="[264,520,433,459]" italics="true" pageId="5" pageNumber="40">Paraleyrodes perseae</emphasis>
(Quaintance)
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</paragraph>
<paragraph id="AD4F36C6FFB27E13FEA7FE50FE4AFE11" blockId="5.[264,693,433,494]" box="[264,400,468,494]" pageId="5" pageNumber="40">
(
<figureCitation id="35CB2A43FFB27E13FEBFFE50FE5DFE11" box="[272,391,468,494]" captionStart-0="FIGURES 5 – 7" captionStart-1="FIGURES 8 – 16" captionStartId-0="6.[264,382,1707,1731]" captionStartId-1="7.[264,382,1333,1357]" captionTargetBox-0="[359,1267,348,1668]" captionTargetBox-1="[299,1272,320,1308]" captionTargetId-0="figure@6.[305,1282,329,1679]" captionTargetId-1="figure@7.[289,1297,302,1337]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." captionText-1="FIGURES 8 – 16, Paraleyrodes spp, male genitalia. 8, P. p e r s e a e, genital segment, with aedeagus in oblique aspect, Persea americana, Nicaragua (Martin); 9, P. p e r s ea e, aedeagus in lateral aspect, Persea americana, Nicaragua (Martin); 10, P. p e r s e a e, aedeagus in lateral aspect, Citrus limon, Florida (Morrill); 11, P. citricolus Costa Lima, aedeagus in lateral aspect, Citrus sinensis, Madeira (Aguiar); 12, P. a n c o r a Martin, aedeagus of holotype, in dorsal aspect,? Lasianthaea sp., Belize (Martin); 13, P. ancora Martin, aedeagus of paratype, in lateral aspect,? Lasianthaea sp., Belize (Martin); 14, Paraleyrodes sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15, Paraleyrodes sp., aedeagus, in dorsal aspect, Citrus sp., Florida (Dowell); 16, Paraleyrodes sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be P. p ers eae." httpUri-0="https://zenodo.org/record/171848/files/figure.png" httpUri-1="https://zenodo.org/record/171849/files/figure.png" pageId="5" pageNumber="40">Figs 5–10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E5EA654DFFB27E13FEA7FDA0FC2DFA76" pageId="5" pageNumber="40" type="reference_group">
<paragraph id="AD4F36C6FFB27E13FEA7FDA0FCB8FDA4" blockId="5.[264,1323,547,667]" pageId="5" pageNumber="40">
<treatmentCitationGroup id="8DE011E8FFB27E13FEA7FDA0FCB8FDA4" pageId="5" pageNumber="40">
<taxonomicName id="6AF04D45FFB27E13FEA7FDA0FD63FDC4" authority="Quaintance, 1900: 32" authorityName="Quaintance" authorityPageNumber="32" authorityYear="1900" box="[264,697,548,571]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFB27E13FEA7FDA0FE17FDC4" box="[264,461,548,571]" italics="true" pageId="5" pageNumber="40">Aleurodes perseae</emphasis>
<treatmentCitation id="2C5110D7FFB27E13FE7BFDA0FD63FDC4" author="Quaintance" box="[468,697,548,571]" page="32" pageId="5" pageNumber="40" year="1900">
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: 32
</treatmentCitation>
</taxonomicName>
–33.
<typeStatus id="724B8864FFB27E13FD42FDA0FC81FDC4" box="[749,859,548,571]" pageId="5" pageNumber="40" type="lectotype">Lectotype</typeStatus>
,
<collectingCountry id="D5E77656FFB27E13FCCAFDA0FC41FDC4" box="[869,923,548,571]" name="United States of America" pageId="5" pageNumber="40">USA</collectingCountry>
(Florida),
<emphasis id="9F84EAD4FFB27E13FBA2FDA7FB65FDC4" bold="true" box="[1037,1215,547,571]" pageId="5" pageNumber="40">here designated</emphasis>
.
<taxonomicName id="6AF04D45FFB27E13FEA7FDC0FC84FDA4" authority="(Quaintance) Cockerell, 1903: 663" authorityName="(Quaintance) Cockerell" authorityPageNumber="663" authorityYear="1903" box="[264,862,580,603]" class="Insecta" family="Aleyrodidae" genus="Aleurodicus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFB27E13FEA7FDC0FE38FDA4" box="[264,482,580,603]" italics="true" pageId="5" pageNumber="40">Aleurodicus perseae</emphasis>
(Quaintance)
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<bibRefCitation id="C9614B37FFB27E13FDD5FDC0FCF2FDA4" author="Cockerell" box="[634,808,580,603]" pageId="5" pageNumber="46" refString="Cockerell, T. D. A. (1903) The White Fly (Aleyrodes citri) and its allies. Bulletin of the Florida Agricultural Experiment Station 67, 662 - 666." type="journal article" year="1903">Cockerell, 1903</bibRefCitation>
: 663
</treatmentCitation>
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.
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<paragraph id="AD4F36C6FFB27E13FEA7FDE0FE80FD64" blockId="5.[264,1323,547,667]" pageId="5" pageNumber="40">
<treatmentCitationGroup id="8DE011E8FFB27E13FEA7FDE0FE80FD64" pageId="5" pageNumber="40">
<taxonomicName id="6AF04D45FFB27E13FEA7FDE0FC53FD84" ID-CoL="6TPLC" authority="(Quaintance) Quaintance, 1909: 170" authorityName="(Quaintance) Quaintance" authorityPageNumber="170" authorityYear="1909" box="[264,905,612,635]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFB27E13FEA7FDE0FE29FD84" box="[264,499,612,635]" italics="true" pageId="5" pageNumber="40">Paraleyrodes perseae</emphasis>
(
<treatmentCitation id="2C5110D7FFB27E13FDACFDE0FC53FD84" author="Quaintance" box="[515,905,612,635]" page="170" pageId="5" pageNumber="40" year="1909">
<bibRefCitation id="C9614B37FFB27E13FDACFDE0FC89FD84" author="Quaintance" box="[515,851,612,635]" pageId="5" pageNumber="47" refString="Quaintance, A. L. (1909) A new genus of Aleyrodidae, with remarks on Aleyrodes nubifera Berger and Aleyrodes citri Riley and Howard. Technical Series, US Department of Agriculture Bureau of Entomology, 12, 169 - 174." type="journal article" year="1909">Quaintance) Quaintance, 1909</bibRefCitation>
: 170
</treatmentCitation>
</taxonomicName>
–173;
<treatmentCitation id="2C5110D7FFB27E13FC61FDE0FAC7FD84" author="Quaintance" box="[974,1309,612,635]" page="82" pageId="5" pageNumber="40" year="1913">
<bibRefCitation id="C9614B37FFB27E13FC61FDE0FB2CFD84" author="Quaintance" box="[974,1270,612,635]" pageId="5" pageNumber="47" refString="Quaintance, A. L. &amp; Baker, A. C. (1913) Classification of the Aleyrodidae Part I. Technical Series, US Department of Agriculture Bureau of Entomology, 27, 1 - 93." type="journal article" year="1913">
Quaintance &amp;
<collectingCountry id="D5E77656FFB27E13FBC1FDE0FB75FD84" box="[1134,1199,612,635]" name="United States Pacific Island Wildlife Refuges" pageId="5" pageNumber="40">Baker</collectingCountry>
, 1913
</bibRefCitation>
: 82
</treatmentCitation>
– 83.
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F36C6FFB27E13FEA7FD43FC2DFA76" blockId="5.[264,1324,711,1657]" pageId="5" pageNumber="40">
PUPARIUM (n=20).
<quantity id="6A089B23FFB27E13FDACFD43FD77FD1E" box="[515,685,711,737]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" metricValueMax="9.0" metricValueMin="6.2" pageId="5" pageNumber="40" unit="mm" value="0.76" valueMax="0.9" valueMin="0.62">0.62–0.90 mm</quantity>
long, outline oval, margin almost smooth. One pair of large compound pores (
<figureCitation id="35CB2A43FFB27E13FD9DFD6BFDA5FCF6" box="[562,639,751,777]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="5" pageNumber="40">Fig. 7</figureCitation>
) present on cephalic segment and each of abdominal segments V–VIII, all similar in size with overall diameter 20–36 (usually 24–32) m; ring of spinneret cells 10–18 (usually 12–16) m in diameter, with 4–8 cells (
<figureCitation id="35CB2A43FFB27E13FBF3FCBBFB79FCA6" box="[1116,1187,831,857]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="5" pageNumber="40">Fig. 7</figureCitation>
), each with an elongate spline issuing from it (
<figureCitation id="35CB2A43FFB27E13FD31FCE3FD3CFC7E" box="[670,742,871,897]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="5" pageNumber="40">Fig. 5</figureCitation>
); between ring of spinneret cells and boundary of compound pore is a narrow annulus with much finer cells; inner lumen of compound pore (when detail can be seen) divided into irregular segments (
<figureCitation id="35CB2A43FFB27E13FC6AFC33FBD5FC2E" box="[965,1039,951,977]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="5" pageNumber="40">Fig. 7</figureCitation>
). A single pair of small compound pores present, on abdominal segment III,
<quantity id="6A089B23FFB27E13FCC1FC5BFC3FFC06" box="[878,997,991,1017]" metricMagnitude="1" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="5" pageNumber="40" unit="m" value="14.0" valueMax="16.0" valueMin="12.0">12–16 m</quantity>
in diameter, about the same size as the thoracic cicatrices (
<figureCitation id="35CB2A43FFB27E13FDC6FB83FD75FBDE" box="[617,687,1031,1057]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="5" pageNumber="40">Fig. 5</figureCitation>
); the pair of small compound pores usually present on abdominal segment IV of
<taxonomicName id="6AF04D45FFB27E13FDEAFBABFD3BFBB7" ID-CoL="6F9B" box="[581,737,1071,1096]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFB27E13FDEAFBABFD3BFBB7" box="[581,737,1071,1096]" italics="true" pageId="5" pageNumber="40">Paraleyrodes</emphasis>
</taxonomicName>
puparia is absent. Subdorsum sparsely provided with small pore/porette geminate pairs, usually a single pairing to either side of abdominal segments V–VII (but not VIII) (
<figureCitation id="35CB2A43FFB27E13FD2DFBFBFD31FB66" box="[642,747,1151,1177]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="5" pageNumber="40">Figs 5–6</figureCitation>
) and most cephalothoracic segments; abdominal segments III and IV each with a subdorsal cluster of 2–5 slightly larger “bright” pores and (mesad) 1–2 tiny spines (
<figureCitation id="35CB2A43FFB27E13FDEEFB4BFD57FB16" box="[577,653,1231,1257]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="5" pageNumber="40">Fig. 5</figureCitation>
). Outer submargin with a ring of widely­spaced small crater­like glands (
<figureCitation id="35CB2A43FFB27E13FE43FB73FD83FAEE" box="[492,601,1271,1297]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="5" pageNumber="40">Figs 5–6</figureCitation>
) between the ring of submarginal setal bases and puparial margin, each one similar in size to a setal base, about 2–4 per pair of setae. Vasiform orifice (
<figureCitation id="35CB2A43FFB27E13FECAFAC3FE76FA9E" box="[357,428,1351,1377]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="5" pageNumber="40">Fig. 6</figureCitation>
) cordate, smooth; operculum as figured, with a pair of tiny setae near midpoint of posterior edge; lingula head as figured, slightly angular.
</paragraph>
</subSubSection>
<subSubSection id="E5EA654DFFB27E13FE97FA13FE7BF88F" pageId="5" pageNumber="40" type="description">
<paragraph id="AD4F36C6FFB27E13FE97FA13FDAFF986" blockId="5.[264,1324,711,1657]" pageId="5" pageNumber="40">
ADULT MALE (n=8). Body 0.90–1.00 mm long, genital segment
<quantity id="6A089B23FFB27E13FBFDFA13FAD6FA4E" box="[1106,1292,1431,1457]" metricMagnitude="-4" metricUnit="m" metricValue="7.075" metricValueMax="13.0" metricValueMin="1.15" pageId="5" pageNumber="40" unit="mm" value="0.7075" valueMax="1.3" valueMin="0.115">0.115–1.30 mm</quantity>
in maximum dimension, claspers
<quantity id="6A089B23FFB27E13FD29FA3BFCEDFA26" box="[646,823,1471,1497]" metricMagnitude="-5" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="5" pageNumber="40" unit="mm" value="0.095" valueMax="0.1" valueMin="0.09">0.09–0.10 mm</quantity>
long, antennal flagellum
<quantity id="6A089B23FFB27E13FBD5FA3BFAF1FA26" box="[1146,1323,1471,1497]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.8" metricValueMin="3.7" pageId="5" pageNumber="40" unit="mm" value="0.425" valueMax="0.48" valueMin="0.37">0.37–0.48 mm</quantity>
long. Aedeagal apex as shown in figures 8–10, with 4 ventrally­directed short “thorns” variably visible, depending on precise alignment on slide (lateral or oblique aspect is needed for diagnostic purposes). Vasiform orifice, operculum and lingula as shown, the lingula subtly lobulate (
<figureCitation id="35CB2A43FFB27E13FDB1F9DBFDBEF986" box="[542,612,1631,1657]" captionStart="FIGURES 8 – 16" captionStartId="7.[264,382,1333,1357]" captionTargetBox="[299,1272,320,1308]" captionTargetId="figure@7.[289,1297,302,1337]" captionTargetPageId="7" captionText="FIGURES 8 – 16, Paraleyrodes spp, male genitalia. 8, P. p e r s e a e, genital segment, with aedeagus in oblique aspect, Persea americana, Nicaragua (Martin); 9, P. p e r s ea e, aedeagus in lateral aspect, Persea americana, Nicaragua (Martin); 10, P. p e r s e a e, aedeagus in lateral aspect, Citrus limon, Florida (Morrill); 11, P. citricolus Costa Lima, aedeagus in lateral aspect, Citrus sinensis, Madeira (Aguiar); 12, P. a n c o r a Martin, aedeagus of holotype, in dorsal aspect,? Lasianthaea sp., Belize (Martin); 13, P. ancora Martin, aedeagus of paratype, in lateral aspect,? Lasianthaea sp., Belize (Martin); 14, Paraleyrodes sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15, Paraleyrodes sp., aedeagus, in dorsal aspect, Citrus sp., Florida (Dowell); 16, Paraleyrodes sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be P. p ers eae." httpUri="https://zenodo.org/record/171849/files/figure.png" pageId="5" pageNumber="40">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="AD4F36C6FFB27E13FE97F918FD7EF949" blockId="5.[312,676,1692,1718]" box="[312,676,1692,1718]" pageId="5" pageNumber="40">SPECIMEN DEPOSITORIES.</paragraph>
<paragraph id="AD4F36C6FFB27E13FEA7F95DFC1EF90F" blockId="5.[264,1322,1753,1904]" box="[264,964,1753,1776]" pageId="5" pageNumber="40">
BMNH—The Natural History Museum, London SW7 5BD,
<collectingCountry id="D5E77656FFB27E13FC3FF95DFC1EF90F" box="[912,964,1753,1776]" name="United Kingdom" pageId="5" pageNumber="40">U.K.</collectingCountry>
</paragraph>
<paragraph id="AD4F36C6FFB27E13FEA7F97DFE7BF88F" blockId="5.[264,1322,1753,1904]" pageId="5" pageNumber="40">
CDFA—California Department of Food &amp; Agriculture, Sacramento, California 95832,
<collectingCountry id="D5E77656FFB27E13FB04F97DFB2FF8EF" box="[1195,1269,1785,1808]" name="United States of America" pageId="5" pageNumber="40">U.S.A.</collectingCountry>
USNM—U.S. Department of Agriculture, Beltsville, Maryland 20705,
<collectingCountry id="D5E77656FFB27E13FBACF89DFBEAF8CF" box="[1027,1072,1817,1840]" name="United States of America" pageId="5" pageNumber="40">U.S.</collectingCountry>
A [Sternorrhyncha collections of the
<collectingCountry id="D5E77656FFB27E13FDA4F8BDFD46F8AF" box="[523,668,1849,1872]" name="United States of America" pageId="5" pageNumber="40">United States</collectingCountry>
National Museum of Natural History / Smithsonian Institution].
</paragraph>
</subSubSection>
<caption id="F98F664EFFB17E10FEA7F92FFE6CF88E" httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="6" pageNumber="41" targetBox="[359,1267,348,1668]" targetPageId="6">
<paragraph id="AD4F36C6FFB17E10FEA7F92FFE6CF88E" blockId="6.[264,1323,1706,1905]" pageId="6" pageNumber="41">
<emphasis id="9F84EAD4FFB17E10FEA7F92FFE1AF93C" bold="true" box="[264,448,1706,1732]" pageId="6" pageNumber="41">FIGURES 5–7,</emphasis>
<taxonomicName id="6AF04D45FFB17E10FE7DF928FCBFF93C" authority="Quaintance" authorityName="Quaintance" box="[466,869,1708,1731]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFB17E10FE7DF928FD1CF93C" box="[466,710,1708,1731]" italics="true" pageId="6" pageNumber="41">Paraleyrodes perseae</emphasis>
(Quaintance)
</taxonomicName>
, puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore.
</paragraph>
</caption>
<caption id="F98F664EFFB07E11FEA7FAB1FDACF97A" httpUri="https://zenodo.org/record/171849/files/figure.png" pageId="7" pageNumber="42" targetBox="[299,1272,320,1308]" targetPageId="7">
<paragraph id="AD4F36C6FFB07E11FEA7FAB1FDACF97A" blockId="7.[264,1323,1331,1669]" pageId="7" pageNumber="42">
<emphasis id="9F84EAD4FFB07E11FEA7FAB1FE18FAB2" bold="true" box="[264,450,1331,1357]" pageId="7" pageNumber="42">FIGURES 8–16,</emphasis>
<taxonomicName id="6AF04D45FFB07E11FE67FAB1FD8DFAB3" box="[456,599,1333,1356]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFB07E11FE67FAB1FD8DFAB3" box="[456,599,1333,1356]" italics="true" pageId="7" pageNumber="42">Paraleyrodes</emphasis>
</taxonomicName>
spp, male genitalia. 8,
<emphasis id="9F84EAD4FFB07E11FCE4FAB1FC6EFAB3" box="[843,948,1333,1356]" italics="true" pageId="7" pageNumber="42">P. p e r s e a e</emphasis>
, genital segment, with aedeagus in oblique aspect,
<taxonomicName id="6AF04D45FFB07E11FE1AFADDFDA0FA8F" box="[437,634,1369,1392]" class="Magnoliopsida" family="Lauraceae" genus="Persea" kingdom="Plantae" order="Laurales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="americana">
<emphasis id="9F84EAD4FFB07E11FE1AFADDFDA0FA8F" box="[437,634,1369,1392]" italics="true" pageId="7" pageNumber="42">Persea americana</emphasis>
</taxonomicName>
, Nicaragua (Martin); 9,
<emphasis id="9F84EAD4FFB07E11FC24FADDFC2DFA8F" box="[907,1015,1369,1392]" italics="true" pageId="7" pageNumber="42">P. p e r s ea e</emphasis>
, aedeagus in lateral aspect,
<taxonomicName id="6AF04D45FFB07E11FEA7FAFFFE14FA6D" box="[264,462,1403,1426]" class="Magnoliopsida" family="Lauraceae" genus="Persea" kingdom="Plantae" order="Laurales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="americana">
<emphasis id="9F84EAD4FFB07E11FEA7FAFFFE14FA6D" box="[264,462,1403,1426]" italics="true" pageId="7" pageNumber="42">Persea americana</emphasis>
</taxonomicName>
, Nicaragua (Martin); 10,
<emphasis id="9F84EAD4FFB07E11FD41FAFFFC81FA6D" box="[750,859,1403,1426]" italics="true" pageId="7" pageNumber="42">P. p e r s e a e</emphasis>
, aedeagus in lateral aspect,
<taxonomicName id="6AF04D45FFB07E11FB35FAFFFAFEFA6D" box="[1178,1316,1403,1426]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="limon">
<emphasis id="9F84EAD4FFB07E11FB35FAFFFAFEFA6D" box="[1178,1316,1403,1426]" italics="true" pageId="7" pageNumber="42">Citrus limon</emphasis>
</taxonomicName>
, Florida (Morrill); 11,
<taxonomicName id="6AF04D45FFB07E11FE5DFA1AFD23FA4A" authority="Costa Lima" authorityName="Costa Lima" box="[498,761,1438,1461]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="citricolus">
<emphasis id="9F84EAD4FFB07E11FE5DFA1AFDABFA4A" box="[498,625,1438,1461]" italics="true" pageId="7" pageNumber="42">P. citricolus</emphasis>
Costa Lima
</taxonomicName>
, aedeagus in lateral aspect,
<taxonomicName id="6AF04D45FFB07E11FB8AFA1AFEBDFA27" authority="Madeira (Aguiar)" authorityName="Madeira (Aguiar" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="sinensis">
<emphasis id="9F84EAD4FFB07E11FB8AFA1AFB19FA4A" box="[1061,1219,1438,1461]" italics="true" pageId="7" pageNumber="42">Citrus sinensis</emphasis>
, Madeira (Aguiar)
</taxonomicName>
; 12,
<emphasis id="9F84EAD4FFB07E11FE0FFA45FDDFFA27" box="[416,517,1473,1496]" italics="true" pageId="7" pageNumber="42">P. a n c o r a</emphasis>
Martin, aedeagus of holotype, in dorsal aspect,?
<taxonomicName id="6AF04D45FFB07E11FB84FA45FB75FA27" box="[1067,1199,1473,1496]" class="Magnoliopsida" family="Asteraceae" genus="Lasianthaea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="9F84EAD4FFB07E11FB84FA45FB75FA27" box="[1067,1199,1473,1496]" italics="true" pageId="7" pageNumber="42">Lasianthaea</emphasis>
</taxonomicName>
sp., Belize (Martin); 13,
<taxonomicName id="6AF04D45FFB07E11FE31FA67FD86FA05" authority="Martin" authorityName="Martin" box="[414,604,1507,1530]" class="Magnoliopsida" family="Lauraceae" genus="Persea" kingdom="Plantae" order="Laurales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="ancora">
<emphasis id="9F84EAD4FFB07E11FE31FA67FDD2FA05" box="[414,520,1507,1530]" italics="true" pageId="7" pageNumber="42">P. ancora</emphasis>
Martin
</taxonomicName>
, aedeagus of paratype, in lateral aspect,?
<taxonomicName id="6AF04D45FFB07E11FB85FA67FB74FA05" box="[1066,1198,1507,1530]" class="Magnoliopsida" family="Asteraceae" genus="Lasianthaea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="9F84EAD4FFB07E11FB85FA67FB74FA05" box="[1066,1198,1507,1530]" italics="true" pageId="7" pageNumber="42">Lasianthaea</emphasis>
</taxonomicName>
sp., Belize (Martin); 14,
<taxonomicName id="6AF04D45FFB07E11FE32F982FDF6F9E2" box="[413,556,1542,1565]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFB07E11FE32F982FDF6F9E2" box="[413,556,1542,1565]" italics="true" pageId="7" pageNumber="42">Paraleyrodes</emphasis>
</taxonomicName>
sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15,
<taxonomicName id="6AF04D45FFB07E11FEA7F9ADFE4DF9BF" box="[264,407,1577,1600]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFB07E11FEA7F9ADFE4DF9BF" box="[264,407,1577,1600]" italics="true" pageId="7" pageNumber="42">Paraleyrodes</emphasis>
</taxonomicName>
sp., aedeagus, in dorsal aspect,
<taxonomicName id="6AF04D45FFB07E11FCADF9ADFC9FF9BF" box="[770,837,1577,1600]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="9F84EAD4FFB07E11FCADF9ADFC9FF9BF" box="[770,837,1577,1600]" italics="true" pageId="7" pageNumber="42">Citrus</emphasis>
</taxonomicName>
sp., Florida (Dowell); 16,
<taxonomicName id="6AF04D45FFB07E11FBDCF9ADFAD8F9BF" box="[1139,1282,1577,1600]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFB07E11FBDCF9ADFAD8F9BF" box="[1139,1282,1577,1600]" italics="true" pageId="7" pageNumber="42">Paraleyrodes</emphasis>
</taxonomicName>
sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be
<emphasis id="9F84EAD4FFB07E11FDADF9EAFDB7F97A" box="[514,621,1646,1669]" italics="true" pageId="7" pageNumber="42">P. p ers eae</emphasis>
.
</paragraph>
</caption>
<subSubSection id="E5EA654DFFB07E1EFE97F95FFBEAFA19" lastPageId="8" lastPageNumber="43" pageId="7" pageNumber="42" type="materials_examined">
<paragraph id="AD4F36C6FFB07E11FE97F95FFC62F90A" blockId="7.[264,1325,1755,1901]" box="[312,952,1755,1781]" pageId="7" pageNumber="42">MATERIAL EXAMINED (arranged chronologically).</paragraph>
<paragraph id="AD4F36C6FFB07E1EFE97F887FE48FAE1" blockId="7.[264,1325,1755,1901]" lastBlockId="8.[264,1325,284,1870]" lastPageId="8" lastPageNumber="43" pageId="7" pageNumber="42">
<typeStatus id="724B8864FFB07E11FE97F887FE77F8E2" box="[312,429,1795,1821]" pageId="7" pageNumber="42" type="lectotype">Lectotype</typeStatus>
puparium of
<taxonomicName id="6AF04D45FFB07E11FDE3F880FCF9F8E2" box="[588,803,1796,1821]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFB07E11FDE3F880FCF9F8E2" box="[588,803,1796,1821]" italics="true" pageId="7" pageNumber="42">Aleurodes perseae</emphasis>
</taxonomicName>
, USA—FLORIDA, Fort George, on
<taxonomicName id="6AF04D45FFB07E11FB76F880FE4CF8BA" class="Magnoliopsida" family="Lauraceae" genus="Persea" kingdom="Plantae" order="Laurales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="carolinensis">
<emphasis id="9F84EAD4FFB07E11FB76F880FE4CF8BA" italics="true" pageId="7" pageNumber="42">Persea carolinensis</emphasis>
</taxonomicName>
,
<date id="D94E1006FFB07E11FE07F8AFFDFDF8BA" box="[424,551,1835,1861]" pageId="7" pageNumber="42" value="1880-04-22">22 iv.1880</date>
(Dr R.S.Turner) (
<collectionCode id="CBE1AE03FFB07E11FCADF8AFFC87F8BA" LSID="urn:lsid:biocol.org:col:34871" box="[770,861,1835,1861]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="42">USNM</collectionCode>
);
<typeStatus id="724B8864FFB07E11FCDAF8AFFBF8F8BA" box="[885,1058,1835,1861]" pageId="7" pageNumber="42" type="paralectotype">paralectotypes</typeStatus>
, 1 puparium, 16 firstinstar larvae and egg remains, all on same slide as
<typeStatus id="724B8864FFB07E11FC35F8D7FBDCF892" box="[922,1030,1875,1901]" pageId="7" pageNumber="42" type="lectotype">lectotype</typeStatus>
(
<collectionCode id="CBE1AE03FFB07E11FBB4F8D7FBACF892" LSID="urn:lsid:biocol.org:col:34871" box="[1051,1142,1875,1901]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="42">USNM</collectionCode>
); 1 parasitised puparium,
<specimenCount id="BBF6FD4FFFBF7E1EFE28FE98FE0DFEC9" box="[391,471,284,310]" pageId="8" pageNumber="43" type="adult">2 adult</specimenCount>
males, USA—FLORIDA, Orlando, on
<taxonomicName id="6AF04D45FFBF7E1EFC09FE98FBE3FECA" box="[934,1081,284,309]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="limon">
<emphasis id="9F84EAD4FFBF7E1EFC09FE98FBE3FECA" box="[934,1081,284,309]" italics="true" pageId="8" pageNumber="43">Citrus limon</emphasis>
</taxonomicName>
,
<date id="D94E1006FFBF7E1EFBE7FE98FB64FEC9" box="[1096,1214,284,310]" pageId="8" pageNumber="43" value="1907-01-28">28 i. 1907</date>
(Morrill, Quaintance collection #5204) (
<collectionCode id="CBE1AE03FFBF7E1EFDDCFEC0FD14FEA1" LSID="urn:lsid:biocol.org:col:34871" box="[627,718,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 1 puparium, USA—FLORIDA, St. Augustine, on “bay” [preseumed to be
<taxonomicName id="6AF04D45FFBF7E1EFDEAFEE8FD22FE7A" box="[581,760,364,389]" class="Magnoliopsida" family="Lauraceae" genus="Laurus" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="nobilis">
<emphasis id="9F84EAD4FFBF7E1EFDEAFEE8FD22FE7A" box="[581,760,364,389]" italics="true" pageId="8" pageNumber="43">Laurus nobilis</emphasis>
</taxonomicName>
],
<date id="D94E1006FFBF7E1EFCB6FEE8FC6CFE79" box="[793,950,364,390]" pageId="8" pageNumber="43" value="1910-07-29">29 vii. 1910</date>
(Back, Quaintance collection #20223) (
<collectionCode id="CBE1AE03FFBF7E1EFE2DFE10FE04FE51" LSID="urn:lsid:biocol.org:col:34871" box="[386,478,404,430]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 4 puparia, USA—VIRGINIA, Vienna, no host data,
<date id="D94E1006FFBF7E1EFB33FE10FAF1FE51" box="[1180,1323,404,430]" pageId="8" pageNumber="43" value="1914-10-31">31 x. 1914</date>
(Williams) (
<collectionCode id="CBE1AE03FFBF7E1EFE33FE38FE20FE29" box="[412,506,444,470]" pageId="8" pageNumber="43">BMNH</collectionCode>
); 2 puparia, USA—FLORIDA, Orlando, on
<taxonomicName id="6AF04D45FFBF7E1EFB88FE38FB22FE2A" box="[1063,1272,444,469]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="aurantium">
<emphasis id="9F84EAD4FFBF7E1EFB88FE38FB22FE2A" box="[1063,1272,444,469]" italics="true" pageId="8" pageNumber="43">Citrus aurantium</emphasis>
</taxonomicName>
, no collection date but registered BM 1915­410 (Hargreaves) (
<collectionCode id="CBE1AE03FFBF7E1EFC7BFE60FBE9FE01" box="[980,1075,484,510]" pageId="8" pageNumber="43">BMNH</collectionCode>
); 3 puparia, 3 thirdinstar larvae, USA—GEORGIA, Darien, on
<taxonomicName id="6AF04D45FFBF7E1EFCB2FD88FCBCFDDA" box="[797,870,524,549]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="9F84EAD4FFBF7E1EFCB2FD88FCBCFDDA" box="[797,870,524,549]" italics="true" pageId="8" pageNumber="43">Citrus</emphasis>
</taxonomicName>
[satsuma],
<date id="D94E1006FFBF7E1EFC5CFD88FBA7FDD9" box="[1011,1149,524,550]" pageId="8" pageNumber="43" value="1943-11-24">24 xi. 1943</date>
(Mayer #396) (
<collectionCode id="CBE1AE03FFBF7E1EFEBFFDB0FEB1FDB1" LSID="urn:lsid:biocol.org:col:34871" box="[272,363,564,590]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 5 puparia, 1 third­instar larva, USA—MISSOURI, St. Louis, on
<emphasis id="9F84EAD4FFBF7E1EFB76FDB0FE5DFD8A" italics="true" pageId="8" pageNumber="43">
<taxonomicName id="6AF04D45FFBF7E1EFB76FDB0FE59FD8A" class="Magnoliopsida" family="Lauraceae" genus="Persea" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="pubsecens">Persea pubsecens</taxonomicName>
,
</emphasis>
<date id="D94E1006FFBF7E1EFE3BFDD8FDCCFD89" box="[404,534,604,630]" pageId="8" pageNumber="43" value="1944-01-27">27 i. 1944</date>
(Mayer) (
<collectionCode id="CBE1AE03FFBF7E1EFD38FDD8FD28FD89" LSID="urn:lsid:biocol.org:col:34871" box="[663,754,604,630]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 2 puparia, USA—FLORIDA, Longwood, on
<taxonomicName id="6AF04D45FFBF7E1EFEA7FD00FE76FD62" box="[264,428,644,669]" class="Magnoliopsida" family="Lauraceae" genus="Cinnamomum" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="9F84EAD4FFBF7E1EFEA7FD00FE76FD62" box="[264,428,644,669]" italics="true" pageId="8" pageNumber="43">Cinnamomum</emphasis>
</taxonomicName>
sp.,
<date id="D94E1006FFBF7E1EFDA8FD00FD75FD61" box="[519,687,644,670]" pageId="8" pageNumber="43" value="1960-06-16">16 vi. 1960</date>
(Youtsey, via Weems) (
<collectionCode id="CBE1AE03FFBF7E1EFBA5FD00FBBFFD61" LSID="urn:lsid:biocol.org:col:34871" box="[1034,1125,644,670]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 1 puparium, USA—FLORIDA, Lake Monroe, on
<emphasis id="9F84EAD4FFBF7E1EFD7DFD28FBDDFD3A" box="[722,1031,684,709]" italics="true" pageId="8" pageNumber="43">
<taxonomicName id="6AF04D45FFBF7E1EFD7DFD28FBD9FD3A" box="[722,1027,684,709]" class="Magnoliopsida" family="Adoxaceae" genus="Viburnum" kingdom="Plantae" order="Dipsacales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="odoratissimum">Viburnum odoratissimum</taxonomicName>
,
</emphasis>
<date id="D94E1006FFBF7E1EFBBBFD28FB76FD39" box="[1044,1196,684,710]" pageId="8" pageNumber="43" value="1963-12-30">30 xii. 1963</date>
(Youtsey) (
<collectionCode id="CBE1AE03FFBF7E1EFEBFFD50FEB1FD11" LSID="urn:lsid:biocol.org:col:34871" box="[272,363,724,750]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 4 puparia,
<specimenCount id="BBF6FD4FFFBF7E1EFE55FD50FD93FD11" box="[506,585,724,750]" pageId="8" pageNumber="43" type="adult">1 adult</specimenCount>
male, USA—FLORIDA, Tallahassee, on
<taxonomicName id="6AF04D45FFBF7E1EFB9CFD50FB6AFD12" box="[1075,1200,724,749]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="9F84EAD4FFBF7E1EFB9CFD50FB6AFD12" box="[1075,1200,724,749]" italics="true" pageId="8" pageNumber="43">Anthurium</emphasis>
</taxonomicName>
stems and leaves,
<date id="D94E1006FFBF7E1EFECBFD78FE28FCE9" box="[356,498,764,790]" pageId="8" pageNumber="43" value="1965-03-15">15 iii. 1965</date>
(Weems via Miller) (
<collectionCode id="CBE1AE03FFBF7E1EFD53FD78FC8DFCE9" LSID="urn:lsid:biocol.org:col:34871" box="[764,855,764,790]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 1 puparium, USA—FLORIDA, New Smyrna Beach, on
<emphasis id="9F84EAD4FFBF7E1EFDA2FCA0FD00FCC2" box="[525,730,804,829]" italics="true" pageId="8" pageNumber="43">
<taxonomicName id="6AF04D45FFBF7E1EFDA2FCA0FD0CFCC2" box="[525,726,804,829]" class="Magnoliopsida" family="Myricaceae" genus="Myrica" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="cerifera">Myrica cerifera</taxonomicName>
,
</emphasis>
<date id="D94E1006FFBF7E1EFD41FCA0FC54FCC1" box="[750,910,804,830]" pageId="8" pageNumber="43" value="1972-03-09">09 iii. 1972</date>
(Roberts) (
<collectionCode id="CBE1AE03FFBF7E1EFB83FCA0FB5DFCC1" LSID="urn:lsid:biocol.org:col:34871" box="[1068,1159,804,830]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 4 puparia, USA—HAWAII, intercepted ex­PHILIPPINES, on
<taxonomicName id="6AF04D45FFBF7E1EFC2EFCC8FBD3FC9A" box="[897,1033,844,869]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="betle">
<emphasis id="9F84EAD4FFBF7E1EFC2EFCC8FBD3FC9A" box="[897,1033,844,869]" italics="true" pageId="8" pageNumber="43">Piper betle</emphasis>
</taxonomicName>
,
<date id="D94E1006FFBF7E1EFBB0FCC8FB76FC99" box="[1055,1196,844,870]" pageId="8" pageNumber="43" value="1980-02-29">29 ii. 1980</date>
(intercept #048458) (
<collectionCode id="CBE1AE03FFBF7E1EFE26FCF0FE3EFC71" LSID="urn:lsid:biocol.org:col:34871" box="[393,484,884,910]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 1 puparium, USA—ILLINOIS, intercepted ex­MEXICO, on
<emphasis id="9F84EAD4FFBF7E1EFB65FCF0FEB7FC4A" italics="true" pageId="8" pageNumber="43">
<taxonomicName id="6AF04D45FFBF7E1EFB65FCF0FEB3FC4A" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="43" phylum="Magnoliopsida" rank="species" species="guajava">Psidium guajava</taxonomicName>
,
</emphasis>
<date id="D94E1006FFBF7E1EFED5FC18FE23FC49" box="[378,505,924,950]" pageId="8" pageNumber="43" value="1981-01-20">20 i. 1981</date>
(intercept #3888) (
<collectionCode id="CBE1AE03FFBF7E1EFD47FC18FC9EFC49" LSID="urn:lsid:biocol.org:col:34871" box="[744,836,924,950]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 3 puparia,
<collectingCountry id="D5E77656FFBF7E1EFC4DFC18FB4EFC49" box="[994,1172,924,950]" name="Venezuela" pageId="8" pageNumber="43">VENEZUELA</collectingCountry>
, Aragua, La Candelaria, on
<taxonomicName id="6AF04D45FFBF7E1EFE13FC40FD49FC22" box="[444,659,964,989]" class="Magnoliopsida" family="Lauraceae" genus="Persea" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="americana">
<emphasis id="9F84EAD4FFBF7E1EFE13FC40FD49FC22" box="[444,659,964,989]" italics="true" pageId="8" pageNumber="43">Persea americana</emphasis>
</taxonomicName>
,
<date id="D94E1006FFBF7E1EFD0CFC40FCCDFC21" box="[675,791,964,990]" pageId="8" pageNumber="43" value="1982-03-01">1.iii.1982</date>
, (Arnal) (
<collectionCode id="CBE1AE03FFBF7E1EFC24FC40FC3DFC21" LSID="urn:lsid:biocol.org:col:34871" box="[907,999,964,990]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 1 puparium, 3 third­instar larvae, USA—CALIFORNIA, intercepted ex­PHILIPPINES, on
<emphasis id="9F84EAD4FFBF7E1EFBA2FC68FB01FBFA" box="[1037,1243,1004,1029]" italics="true" pageId="8" pageNumber="43">
<taxonomicName id="6AF04D45FFBF7E1EFBA2FC68FB0DFBFA" box="[1037,1239,1004,1029]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="43" phylum="Magnoliopsida" rank="species" species="guajava">Psidium guajava</taxonomicName>
,
</emphasis>
<date id="D94E1006FFBF7E1EFB4AFC68FE99FBD1" pageId="8" pageNumber="43" value="1982-06-10">10 vi. 1982</date>
(intercept #31335) (
<collectionCode id="CBE1AE03FFBF7E1EFD96FB90FD4EFBD1" LSID="urn:lsid:biocol.org:col:34871" box="[569,660,1044,1070]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
); 7 puparia, 1 third­instar with developing puparium,
<specimenCount id="BBF6FD4FFFBF7E1EFAB4FB90FE98FBA9" pageId="8" pageNumber="43" type="adult">7 adult</specimenCount>
males,
<collectingCountry id="D5E77656FFBF7E1EFE0EFBB8FD8EFBA9" box="[417,596,1084,1110]" name="Nicaragua" pageId="8" pageNumber="43">NICARAGUA</collectingCountry>
, Rio San Juan / Rio Bartola confluence, near El Castillo, on
<taxonomicName id="6AF04D45FFBF7E1EFEA7FBE0FE39FB82" box="[264,483,1124,1149]" class="Magnoliopsida" family="Lauraceae" genus="Persea" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="americana">
<emphasis id="9F84EAD4FFBF7E1EFEA7FBE0FE39FB82" box="[264,483,1124,1149]" italics="true" pageId="8" pageNumber="43">Persea americana</emphasis>
</taxonomicName>
,
<date id="D94E1006FFBF7E1EFE56FBE0FD50FB81" box="[505,650,1124,1150]" pageId="8" pageNumber="43" value="2004-06-23">23 vi. 2004</date>
(Martin #8080) (
<collectionCode id="CBE1AE03FFBF7E1EFCC8FBE0FC1CFB81" box="[871,966,1124,1150]" pageId="8" pageNumber="43">BMNH</collectionCode>
). [Note—as discussed in the Introduction, no characters have been identified that allow adult females of
<taxonomicName id="6AF04D45FFBF7E1EFB21FB08FAF1FB5A" box="[1166,1323,1164,1189]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBF7E1EFB21FB08FAF1FB5A" box="[1166,1323,1164,1189]" italics="true" pageId="8" pageNumber="43">Paraleyrodes</emphasis>
</taxonomicName>
to be determined to species: with the frequency of mixed populations occurring (see Conclusions, below), all females are regarded as only generically determined and are not listed here.]
</paragraph>
<paragraph id="AD4F36C6FFBF7E1EFE97FAA8FBEAFA19" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="43">
The
<typeStatus id="724B8864FFBF7E1EFEC1FAA8FE03FAB9" box="[366,473,1324,1350]" pageId="8" pageNumber="43" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="6AF04D45FFBF7E1EFDABFAA8FD07FABA" box="[516,733,1324,1349]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBF7E1EFDABFAA8FD07FABA" box="[516,733,1324,1349]" italics="true" pageId="8" pageNumber="43">Aleurodes perseae</emphasis>
</taxonomicName>
is one of two puparia on a slide bearing the
<typeStatus id="724B8864FFBF7E1EFB58FAA8FAF3FAB9" box="[1271,1321,1324,1350]" pageId="8" pageNumber="43">type</typeStatus>
data (see above), is clearly indicated on the adjacent red
<collectionCode id="CBE1AE03FFBF7E1EFC67FAD0FBC6FA91" LSID="urn:lsid:biocol.org:col:34871" box="[968,1052,1364,1390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="43">USNM</collectionCode>
<typeStatus id="724B8864FFBF7E1EFB87FAD0FB81FA91" box="[1064,1115,1364,1390]" pageId="8" pageNumber="43">type</typeStatus>
label, and is the specimen with clearly developing adult eyes visible through the cuticle. The
<typeStatus id="724B8864FFBF7E1EFB0DFAF8FAD4FA69" box="[1186,1294,1404,1430]" pageId="8" pageNumber="43" type="lectotype">lectotype</typeStatus>
is the specimen in best condition, and the author considers that improvement through remounting, with proper maceration, would be possible in the future.
</paragraph>
</subSubSection>
<subSubSection id="E5EA654DFFBF7E1EFE97FA70FD60F952" pageId="8" pageNumber="43" type="distribution">
<paragraph id="AD4F36C6FFBF7E1EFE97FA70FE0EF9A1" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="43">
DISTRIBUTION. USA—Florida,
<collectingCountry id="D5E77656FFBF7E1EFCAFFA70FCBBF9F1" box="[768,865,1524,1550]" name="Georgia" pageId="8" pageNumber="43">Georgia</collectingCountry>
, Missouri, Virginia; Neotropical Region—Mexico,
<collectingCountry id="D5E77656FFBF7E1EFE4AF998FDBBF9C9" box="[485,609,1564,1590]" name="Nicaragua" pageId="8" pageNumber="43">Nicaragua</collectingCountry>
,
<collectingCountry id="D5E77656FFBF7E1EFDDEF998FD37F9C9" box="[625,749,1564,1590]" name="Venezuela" pageId="8" pageNumber="43">Venezuela</collectingCountry>
; Austro­oriental Region—[
<collectingCountry id="D5E77656FFBF7E1EFB9BF998FB1BF9C9" box="[1076,1217,1564,1590]" name="Philippines" pageId="8" pageNumber="43">Philippines</collectingCountry>
]; Pacific Region—Hawaii.
</paragraph>
<paragraph id="AD4F36C6FFBF7E1EFE97F9E8FD60F952" blockId="8.[264,1325,284,1870]" pageId="8" pageNumber="43">
HOST PLANT GENERA.
<emphasis id="9F84EAD4FFBF7E1EFDD9F9E8FD60F952" italics="true" pageId="8" pageNumber="43">
<taxonomicName id="6AF04D45FFBF7E1EFDD9F9E8FD2CF97A" box="[630,758,1644,1669]" class="Liliopsida" family="Araceae" genus="Anthurium" kingdom="Plantae" order="Alismatales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Anthurium</taxonomicName>
,
<taxonomicName id="6AF04D45FFBF7E1EFCADF9E8FC70F97A" box="[770,938,1644,1669]" class="Magnoliopsida" family="Lauraceae" genus="Cinnamomum" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Cinnamomum</taxonomicName>
,
<taxonomicName id="6AF04D45FFBF7E1EFC1AF9E8FBDBF97A" box="[949,1025,1644,1669]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Citrus</taxonomicName>
,
<taxonomicName id="6AF04D45FFBF7E1EFBA2F9E8FBB9F97A" box="[1037,1123,1644,1669]" class="Magnoliopsida" family="Lauraceae" genus="Laurus" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Laurus</taxonomicName>
,
<taxonomicName id="6AF04D45FFBF7E1EFBC0F9E8FB1DF97A" box="[1135,1223,1644,1669]" class="Magnoliopsida" family="Myricaceae" genus="Myrica" kingdom="Plantae" order="Fagales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Myrica</taxonomicName>
,
<taxonomicName id="6AF04D45FFBF7E1EFB7DF9E8FAFCF97A" box="[1234,1318,1644,1669]" class="Magnoliopsida" family="Lauraceae" genus="Persea" kingdom="Plantae" order="Laurales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Persea</taxonomicName>
,
<taxonomicName id="6AF04D45FFBF7E1EFEA7F910FE1FF952" authority="Plumeria" authorityName="Plumeria" box="[264,453,1684,1709]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Piper, Plumeria</taxonomicName>
,
<taxonomicName id="6AF04D45FFBF7E1EFE7FF910FDE9F952" box="[464,563,1684,1709]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="43" phylum="Magnoliopsida" rank="genus">Psidium</taxonomicName>
,
<taxonomicName id="6AF04D45FFBF7E1EFD90F910FD6CF952" box="[575,694,1684,1709]" class="Magnoliopsida" family="Adoxaceae" genus="Viburnum" kingdom="Plantae" order="Dipsacales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">Viburnum</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="E5EA654DFFBF7E1DFE97F938FB7BFD61" lastPageId="11" lastPageNumber="46" pageId="8" pageNumber="43" type="description">
<paragraph id="AD4F36C6FFBF7E1FFE97F938FDBBFD89" blockId="8.[264,1325,284,1870]" lastBlockId="9.[264,1323,284,1870]" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43">
DESCRIPTIVE CONFLICT—PUPARIUM. In the original description of
<taxonomicName id="6AF04D45FFBF7E1EFB1AF938FD83F901" authority="Quaintance (1900)" authorityName="Quaintance" authorityYear="1900" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBF7E1EFB1AF938FEB9F902" italics="true" pageId="8" pageNumber="43">Aleurodes perseae</emphasis>
<bibRefCitation id="C9614B37FFBF7E1EFEDBF960FD83F901" author="Quaintance" box="[372,601,1764,1790]" pageId="8" pageNumber="47" refString="Quaintance, A. L. (1900) Contribution towards a monograph of the American Aleurodidae. Technical Series, US Department of Agriculture Bureau of Entomology, 8, 9 - 64." type="journal article" year="1900">Quaintance (1900)</bibRefCitation>
</taxonomicName>
stated that “On the dorsum are 5 pairs of cup­shaped compound pores, 4 pairs on caudal third of case [posterior part of abdomen], and the fifth pair on cephalic segment…..”. However, the puparium was illustrated with five
<emphasis id="9F84EAD4FFBF7E1EFB02F8B0FAF1F8B2" box="[1197,1323,1844,1869]" italics="true" pageId="8" pageNumber="43">abdominal</emphasis>
pairs of compound pores, the anteriormost abdominal pair being significantly smaller than all the others. The original account also described puparial secretions that were much more extensive than is usually seen amongst
<taxonomicName id="6AF04D45FFBE7E1FFD71FEE8FCA1FE7A" box="[734,891,364,389]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBE7E1FFD71FEE8FCA1FE7A" box="[734,891,364,389]" italics="true" pageId="9" pageNumber="44">Paraleyrodes</emphasis>
</taxonomicName>
species: in a later paper
<bibRefCitation id="C9614B37FFBE7E1FFB0AFEE8FE8CFE51" author="Quaintance" pageId="9" pageNumber="47" refString="Quaintance, A. L. (1909) A new genus of Aleyrodidae, with remarks on Aleyrodes nubifera Berger and Aleyrodes citri Riley and Howard. Technical Series, US Department of Agriculture Bureau of Entomology, 12, 169 - 174." type="journal article" year="1909">Quaintance (1909)</bibRefCitation>
quoted the opinion of Dr H.W. Morrill, that puparia with such secretions had proved to be parasitised, and that is confirmed by the present author’s field observations of
<taxonomicName id="6AF04D45FFBE7E1FFEA7FE60FE7FFE02" box="[264,421,484,509]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBE7E1FFEA7FE60FE7FFE02" box="[264,421,484,509]" italics="true" pageId="9" pageNumber="44">Paraleyrodes</emphasis>
</taxonomicName>
puparia (
<figureCitation id="35CB2A43FFBE7E1FFDB2FE60FDB0FE01" box="[541,618,484,510]" captionStart="FIGURES 3 – 4" captionStartId="3.[264,382,1786,1810]" captionTargetBox="[264,1322,326,1762]" captionTargetId="figure@3.[264,1323,326,1778]" captionTargetPageId="3" captionText="FIGURES 3 – 4, Paraleyrodes spp. ex­Belize. 3, feeding puparium, showing characteristic annulus of wax fragments and five pairs of waxy filaments issuing from the large compound pores; 4, characteristic wax curls on parasitised puparium (see discussion on p. 44)." httpUri="https://zenodo.org/record/171847/files/figure.png" pageId="9" pageNumber="44">Fig. 4</figureCitation>
). This then raised the question of whether the originallydescribed form might have been atypical, due to parasitism, with normal specimens (not seen by Quaintance at that time) perhaps having the typical complement of seven pairs of compound pore pairs in total.
</paragraph>
<paragraph id="AD4F36C6FFBE7E1FFE97FD00FB0DFC21" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="44">
<bibRefCitation id="C9614B37FFBE7E1FFE97FD00FDCFFD61" author="Quaintance" box="[312,533,644,670]" pageId="9" pageNumber="47" refString="Quaintance, A. L. (1909) A new genus of Aleyrodidae, with remarks on Aleyrodes nubifera Berger and Aleyrodes citri Riley and Howard. Technical Series, US Department of Agriculture Bureau of Entomology, 12, 169 - 174." type="journal article" year="1909">Quaintance (1909)</bibRefCitation>
redescribed
<taxonomicName id="6AF04D45FFBE7E1FFD1EFD00FC56FD62" box="[689,908,644,669]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBE7E1FFD1EFD00FC56FD62" box="[689,908,644,669]" italics="true" pageId="9" pageNumber="44">Aleurodes perseae</emphasis>
</taxonomicName>
, and formally designated it as the
<typeStatus id="724B8864FFBE7E1FFEA7FD28FEE0FD39" box="[264,314,684,710]" pageId="9" pageNumber="44">type</typeStatus>
species of his new genus
<emphasis id="9F84EAD4FFBE7E1FFDD2FD28FCF8FD3A" box="[637,802,684,709]" italics="true" pageId="9" pageNumber="44">
<taxonomicName id="6AF04D45FFBE7E1FFDD2FD28FCC4FD3A" box="[637,798,684,709]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">Paraleyrodes</taxonomicName>
.
</emphasis>
In his “revised description” the compound pore complement did indeed rise from five pairs (described) and six pairs (illustrated) [in 1900] to seven pairs (both described and illustrated), this being the typical situation throughout the genus as now understood. This revised description was repeated by
<bibRefCitation id="C9614B37FFBE7E1FFEA7FCC8FDBFFC99" author="Quaintance" box="[264,613,844,870]" pageId="9" pageNumber="47" refString="Quaintance, A. L. &amp; Baker, A. C. (1913) Classification of the Aleyrodidae Part I. Technical Series, US Department of Agriculture Bureau of Entomology, 27, 1 - 93." type="journal article" year="1913">
Quaintance &amp;
<collectingCountry id="D5E77656FFBE7E1FFE6FFCC8FDDCFC99" box="[448,518,844,870]" name="United States Pacific Island Wildlife Refuges" pageId="9" pageNumber="44">Baker</collectingCountry>
(1913)
</bibRefCitation>
, but with fresh illustrations provided. Whereas the 1909 drawing was a fairly accurate representation of a typical
<taxonomicName id="6AF04D45FFBE7E1FFC0DFCF0FBE5FC72" box="[930,1087,884,909]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBE7E1FFC0DFCF0FBE5FC72" box="[930,1087,884,909]" italics="true" pageId="9" pageNumber="44">Paraleyrodes</emphasis>
</taxonomicName>
puparium, the 1913 puparial figure showed all six abdominal compound pores to be the same size as each other, a characteristic of
<taxonomicName id="6AF04D45FFBE7E1FFD86FC40FD3AFC22" box="[553,736,964,989]" class="Insecta" family="Aleyrodidae" genus="Metaleurodicus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBE7E1FFD86FC40FD3AFC22" box="[553,736,964,989]" italics="true" pageId="9" pageNumber="44">Metaleurodicus</emphasis>
</taxonomicName>
species that is never seen in
<taxonomicName id="6AF04D45FFBE7E1FFB9BFC40FB0BFC22" box="[1076,1233,964,989]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBE7E1FFB9BFC40FB0BFC22" box="[1076,1233,964,989]" italics="true" pageId="9" pageNumber="44">Paraleyrodes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="AD4F36C6FFBE7E1FFE97FC68FB6BFB31" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="44">
Although puparia with only five pairs of abdominal compound pores were generally (and correctly, see Conclusions) being identified as
<taxonomicName id="6AF04D45FFBE7E1FFCC2FB90FC3CFBD2" box="[877,998,1044,1069]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBE7E1FFCC2FB90FC3CFBD2" box="[877,998,1044,1069]" italics="true" pageId="9" pageNumber="44">P. perseae</emphasis>
</taxonomicName>
for many years,
<bibRefCitation id="C9614B37FFBE7E1FFB00FB90FE39FBA9" author="Paulson" pageId="9" pageNumber="47" refString="Paulson, G. S. &amp; Kumashiro, B. R. (1985) Hawaiian Aleyrodidae. Proceedings, Hawaiian Entomological Society, 25, 103 - 124." type="journal article" year="1985">Paulson &amp; Kumashiro (1985)</bibRefCitation>
stated that puparia of Hawaiian
<taxonomicName id="6AF04D45FFBE7E1FFCDCFBB8FC34FBAA" box="[883,1006,1084,1109]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBE7E1FFCDCFBB8FC34FBAA" box="[883,1006,1084,1109]" italics="true" pageId="9" pageNumber="44">P. perseae</emphasis>
</taxonomicName>
bore six abdominal pairs, thus continuing the confusion over this species’ identity. However examination of material on behalf of the author (Bernarr Kumashiro, personal communication) indicates that the Hawaiian record is correct but that Paulson &amp; Kumashiro’s description was not.
</paragraph>
<paragraph id="AD4F36C6FFBE7E1FFE97FB58FBCBFA69" blockId="9.[264,1323,284,1870]" pageId="9" pageNumber="44">
DESCRIPTIVE CONFLICT—ADULT MALE. In the original description
<bibRefCitation id="C9614B37FFBE7E1FFB0AFB58FE8CFAE1" author="Quaintance" pageId="9" pageNumber="47" refString="Quaintance, A. L. (1900) Contribution towards a monograph of the American Aleurodidae. Technical Series, US Department of Agriculture Bureau of Entomology, 8, 9 - 64." type="journal article" year="1900">Quaintance (1900)</bibRefCitation>
quoted Theodore Pergande’s very basic observations on an adult, but the sex was not noted and Quaintance stated that the single specimen was lost before he could examine it himself. Pergande’s colour observations do suggest that the lost specimen might indeed have belonged to
<taxonomicName id="6AF04D45FFBE7E1FFE77FAF8FDAFFA6A" box="[472,629,1404,1429]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBE7E1FFE77FAF8FDAFFA6A" box="[472,629,1404,1429]" italics="true" pageId="9" pageNumber="44">Paraleyrodes</emphasis>
</taxonomicName>
, but nothing more can be deduced.
</paragraph>
<paragraph id="AD4F36C6FFBE7E1CFE97FA20FCF3FEA1" blockId="9.[264,1323,284,1870]" lastBlockId="10.[264,1325,284,1870]" lastPageId="10" lastPageNumber="45" pageId="9" pageNumber="44">
In his 1909 redescription, Quaintance did describe adults and spoke of Morrill having “been able to obtain the adult in quantity” and having “kindly furnished [Quaintance] with abundant specimens of all stages…..”. However, he described adults generally as having four antennal segments, which is a character of female
<taxonomicName id="6AF04D45FFBE7E1FFC3EF998FBF4F9CA" box="[913,1070,1564,1589]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBE7E1FFC3EF998FBF4F9CA" box="[913,1070,1564,1589]" italics="true" pageId="9" pageNumber="44">Paraleyrodes</emphasis>
</taxonomicName>
only, and the caption of his illustration of such a female antenna simply says “antenna of adult”, suggesting that he did not study any male antennae at all. Quaintance clearly did study at least one male specimen, and the aedeagal apex was illustrated as a two­pronged structure (reproduced here,
<figureCitation id="35CB2A43FFBE7E1FFEE1F938FE72F929" box="[334,424,1724,1750]" captionStart="FIGURES 8 – 16" captionStartId="7.[264,382,1333,1357]" captionTargetBox="[299,1272,320,1308]" captionTargetId="figure@7.[289,1297,302,1337]" captionTargetPageId="7" captionText="FIGURES 8 – 16, Paraleyrodes spp, male genitalia. 8, P. p e r s e a e, genital segment, with aedeagus in oblique aspect, Persea americana, Nicaragua (Martin); 9, P. p e r s ea e, aedeagus in lateral aspect, Persea americana, Nicaragua (Martin); 10, P. p e r s e a e, aedeagus in lateral aspect, Citrus limon, Florida (Morrill); 11, P. citricolus Costa Lima, aedeagus in lateral aspect, Citrus sinensis, Madeira (Aguiar); 12, P. a n c o r a Martin, aedeagus of holotype, in dorsal aspect,? Lasianthaea sp., Belize (Martin); 13, P. ancora Martin, aedeagus of paratype, in lateral aspect,? Lasianthaea sp., Belize (Martin); 14, Paraleyrodes sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15, Paraleyrodes sp., aedeagus, in dorsal aspect, Citrus sp., Florida (Dowell); 16, Paraleyrodes sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be P. p ers eae." httpUri="https://zenodo.org/record/171849/files/figure.png" pageId="9" pageNumber="44">Fig. 16</figureCitation>
), but the lack of comment on the highly unusual and characteristic male antennae is remarkable. In Quaintance &amp; Baker’s (1913) new illustrations, that accompanied the repeated 1909 revised description of “
<taxonomicName id="6AF04D45FFBE7E1FFC66F888FBFEF8DA" box="[969,1060,1804,1829]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBE7E1FFC66F888FBFEF8DA" box="[969,1060,1804,1829]" italics="true" pageId="9" pageNumber="44">perseae</emphasis>
</taxonomicName>
”, the bifurcate male aedeagus seems to have been drawn from the same specimen figured in 1909, but the drawing is not quite identical; also a clearly­female antenna is again figured and captioned as though it could have belonged to either sex.
</paragraph>
<paragraph id="AD4F36C6FFBD7E1CFE97FEE8FC8BFCC1" blockId="10.[264,1325,284,1870]" pageId="10" pageNumber="45">
A single male in BMNH (
<collectingCountry id="D5E77656FFBD7E1CFDC2FEE8FD76FE79" box="[621,684,364,390]" name="United States of America" pageId="10" pageNumber="45">USA</collectingCountry>
, Florida, Orlando, no host data,
<date id="D94E1006FFBD7E1CFB81FEE8FB74FE79" box="[1070,1198,364,390]" pageId="10" pageNumber="45" value="1914-11-12">12.xi.1914</date>
, Williams,
<figureCitation id="35CB2A43FFBD7E1CFEA7FE10FE86FE51" box="[264,348,404,430]" captionStart="FIGURES 8 – 16" captionStartId="7.[264,382,1333,1357]" captionTargetBox="[299,1272,320,1308]" captionTargetId="figure@7.[289,1297,302,1337]" captionTargetPageId="7" captionText="FIGURES 8 – 16, Paraleyrodes spp, male genitalia. 8, P. p e r s e a e, genital segment, with aedeagus in oblique aspect, Persea americana, Nicaragua (Martin); 9, P. p e r s ea e, aedeagus in lateral aspect, Persea americana, Nicaragua (Martin); 10, P. p e r s e a e, aedeagus in lateral aspect, Citrus limon, Florida (Morrill); 11, P. citricolus Costa Lima, aedeagus in lateral aspect, Citrus sinensis, Madeira (Aguiar); 12, P. a n c o r a Martin, aedeagus of holotype, in dorsal aspect,? Lasianthaea sp., Belize (Martin); 13, P. ancora Martin, aedeagus of paratype, in lateral aspect,? Lasianthaea sp., Belize (Martin); 14, Paraleyrodes sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15, Paraleyrodes sp., aedeagus, in dorsal aspect, Citrus sp., Florida (Dowell); 16, Paraleyrodes sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be P. p ers eae." httpUri="https://zenodo.org/record/171849/files/figure.png" pageId="10" pageNumber="45">Fig. 14</figureCitation>
) has a bifurcate aedeagus, but with the prong bases not contiguous as illustrated in the
<date id="D94E1006FFBD7E1CFE94FE38FE21FE29" box="[315,507,444,470]" pageId="10" pageNumber="45" value="1909" valueMax="1913">1909 and 1913</date>
papers. Two further male specimens, in CDFA, (
<collectingCountry id="D5E77656FFBD7E1CFBD2FE38FB66FE29" box="[1149,1212,444,470]" name="United States of America" pageId="10" pageNumber="45">USA</collectingCountry>
, Florida, Broward Co., Davie,
<taxonomicName id="6AF04D45FFBD7E1CFDB8FE60FDBAFE02" box="[535,608,484,509]" class="Magnoliopsida" family="Rutaceae" genus="Citrus" kingdom="Plantae" order="Sapindales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="genus">
<emphasis id="9F84EAD4FFBD7E1CFDB8FE60FDBAFE02" box="[535,608,484,509]" italics="true" pageId="10" pageNumber="45">Citrus</emphasis>
</taxonomicName>
sp.,
<date id="D94E1006FFBD7E1CFD09FE60FCF4FE01" box="[678,814,484,510]" pageId="10" pageNumber="45" value="1985-12-08">08.xii.1985</date>
, Dowell,
<figureCitation id="35CB2A43FFBD7E1CFC02FE60FBD2FE01" box="[941,1032,484,510]" captionStart="FIGURES 8 – 16" captionStartId="7.[264,382,1333,1357]" captionTargetBox="[299,1272,320,1308]" captionTargetId="figure@7.[289,1297,302,1337]" captionTargetPageId="7" captionText="FIGURES 8 – 16, Paraleyrodes spp, male genitalia. 8, P. p e r s e a e, genital segment, with aedeagus in oblique aspect, Persea americana, Nicaragua (Martin); 9, P. p e r s ea e, aedeagus in lateral aspect, Persea americana, Nicaragua (Martin); 10, P. p e r s e a e, aedeagus in lateral aspect, Citrus limon, Florida (Morrill); 11, P. citricolus Costa Lima, aedeagus in lateral aspect, Citrus sinensis, Madeira (Aguiar); 12, P. a n c o r a Martin, aedeagus of holotype, in dorsal aspect,? Lasianthaea sp., Belize (Martin); 13, P. ancora Martin, aedeagus of paratype, in lateral aspect,? Lasianthaea sp., Belize (Martin); 14, Paraleyrodes sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15, Paraleyrodes sp., aedeagus, in dorsal aspect, Citrus sp., Florida (Dowell); 16, Paraleyrodes sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be P. p ers eae." httpUri="https://zenodo.org/record/171849/files/figure.png" pageId="10" pageNumber="45">Fig. 15</figureCitation>
) each have a bifurcate aedegus, but with the prongs almost parallel and directed posteriorly, rather than laterally as in the Williams specimen. Either of these males may belong to the species which Quaintance and Quaintance &amp;
<collectingCountry id="D5E77656FFBD7E1CFDDDFDD8FD62FD89" box="[626,696,604,630]" name="United States Pacific Island Wildlife Refuges" pageId="10" pageNumber="45">Baker</collectingCountry>
thought was
<taxonomicName id="6AF04D45FFBD7E1CFCFAFDD8FC17FD8A" box="[853,973,604,629]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBD7E1CFCFAFDD8FC17FD8A" box="[853,973,604,629]" italics="true" pageId="10" pageNumber="45">P. perseae</emphasis>
</taxonomicName>
, but both remain undescribed and unassociated with puparia. The aedeagus depicted in the
<date id="D94E1006FFBD7E1CFC46FD00FB46FD61" box="[1001,1180,644,670]" pageId="10" pageNumber="45" value="1909" valueMax="1913">1909 and 1913</date>
papers also bears some resemblance to
<emphasis id="9F84EAD4FFBD7E1CFDE5FD28FD6CFD3A" box="[586,694,684,709]" italics="true" pageId="10" pageNumber="45">P. a n c o r a</emphasis>
<bibRefCitation id="C9614B37FFBD7E1CFD6EFD28FCBCFD39" author="Martin" box="[705,870,684,710]" pageId="10" pageNumber="47" refString="Martin, J. H. (2004) Whiteflies of Belize (Hemiptera: Aleyrodidae). Part 1 - introduction and account of the subfamily Aleurodicinae Quaintance &amp; Baker. Zootaxa, 681, 1 - 119." type="journal article" year="2004">Martin (2004)</bibRefCitation>
(
<figureCitation id="35CB2A43FFBD7E1CFCD9FD28FC24FD39" box="[886,1022,684,710]" captionStart="FIGURES 8 – 16" captionStartId="7.[264,382,1333,1357]" captionTargetBox="[299,1272,320,1308]" captionTargetId="figure@7.[289,1297,302,1337]" captionTargetPageId="7" captionText="FIGURES 8 – 16, Paraleyrodes spp, male genitalia. 8, P. p e r s e a e, genital segment, with aedeagus in oblique aspect, Persea americana, Nicaragua (Martin); 9, P. p e r s ea e, aedeagus in lateral aspect, Persea americana, Nicaragua (Martin); 10, P. p e r s e a e, aedeagus in lateral aspect, Citrus limon, Florida (Morrill); 11, P. citricolus Costa Lima, aedeagus in lateral aspect, Citrus sinensis, Madeira (Aguiar); 12, P. a n c o r a Martin, aedeagus of holotype, in dorsal aspect,? Lasianthaea sp., Belize (Martin); 13, P. ancora Martin, aedeagus of paratype, in lateral aspect,? Lasianthaea sp., Belize (Martin); 14, Paraleyrodes sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15, Paraleyrodes sp., aedeagus, in dorsal aspect, Citrus sp., Florida (Dowell); 16, Paraleyrodes sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be P. p ers eae." httpUri="https://zenodo.org/record/171849/files/figure.png" pageId="10" pageNumber="45">Figs 12–13</figureCitation>
), described from material on avocado in
<collectingCountry id="D5E77656FFBD7E1CFE6CFD50FDD4FD11" box="[451,526,724,750]" name="Belize" pageId="10" pageNumber="45">Belize</collectingCountry>
and
<collectingCountry id="D5E77656FFBD7E1CFDFDFD50FD14FD11" box="[594,718,724,750]" name="Nicaragua" pageId="10" pageNumber="45">Nicaragua</collectingCountry>
; however,
<emphasis id="9F84EAD4FFBD7E1CFCF7FD50FC10FD12" box="[856,970,724,749]" italics="true" pageId="10" pageNumber="45">P. a n c o r a</emphasis>
also features a dorso­apical aedeagal process not present in the Quaintance / Quaintance &amp;
<collectingCountry id="D5E77656FFBD7E1CFBABFD78FB90FCE9" box="[1028,1098,764,790]" name="United States Pacific Island Wildlife Refuges" pageId="10" pageNumber="45">Baker</collectingCountry>
drawings, and has not yet been recorded from anywhere in the
<collectingCountry id="D5E77656FFBD7E1CFCA0FCA0FC97FCC1" box="[783,845,804,830]" name="United States of America" pageId="10" pageNumber="45">USA</collectingCountry>
.
</paragraph>
<paragraph id="AD4F36C6FFBD7E1CFE97FCC8FBB0FB31" blockId="10.[264,1325,284,1870]" pageId="10" pageNumber="45">
CONCLUSIONS. After so many years of uncertainty concerning the identity of
<taxonomicName id="6AF04D45FFBD7E1CFABAFCC8FEB9FC72" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBD7E1CFABAFCC8FEB9FC72" italics="true" pageId="10" pageNumber="45">P. perseae</emphasis>
</taxonomicName>
, recent observations have enabled a resolution of the problem, through examination of puparia, associated adult males, and puparia containing fully developed pre­emergence males. The 1900 puparial illustration is correct in its depiction of the compound pores, the “missing” pair of small compound pores is not an artifact of parasitism, and this confirms that
<taxonomicName id="6AF04D45FFBD7E1CFD0BFB90FCC4FBD2" box="[676,798,1044,1069]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBD7E1CFD0BFB90FCC4FBD2" box="[676,798,1044,1069]" italics="true" pageId="10" pageNumber="45">P. perseae</emphasis>
</taxonomicName>
is the only known species of
<taxonomicName id="6AF04D45FFBD7E1CFB21FB90FAF1FBD2" box="[1166,1323,1044,1069]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBD7E1CFB21FB90FAF1FBD2" box="[1166,1323,1044,1069]" italics="true" pageId="10" pageNumber="45">Paraleyrodes</emphasis>
</taxonomicName>
with a pair of very small compound pores on abdominal segment III, but none at all on segment IV. For the future, the distinctive puparia will probably allow accurate identification, even in the absence of confirmatory males. In practice, puparia have hitherto tended to be correctly identified, and it is the males that have not.
</paragraph>
<paragraph id="AD4F36C6FFBD7E1CFE97FB58FE3EF9F1" blockId="10.[264,1325,284,1870]" pageId="10" pageNumber="45">
The aedeagal apex of male specimens recently collected in
<collectingCountry id="D5E77656FFBD7E1CFC59FB58FBB5FB09" box="[1014,1135,1244,1270]" name="Nicaragua" pageId="10" pageNumber="45">Nicaragua</collectingCountry>
(
<figureCitation id="35CB2A43FFBD7E1CFBD0FB58FB30FB09" box="[1151,1258,1244,1270]" captionStart="FIGURES 8 – 16" captionStartId="7.[264,382,1333,1357]" captionTargetBox="[299,1272,320,1308]" captionTargetId="figure@7.[289,1297,302,1337]" captionTargetPageId="7" captionText="FIGURES 8 – 16, Paraleyrodes spp, male genitalia. 8, P. p e r s e a e, genital segment, with aedeagus in oblique aspect, Persea americana, Nicaragua (Martin); 9, P. p e r s ea e, aedeagus in lateral aspect, Persea americana, Nicaragua (Martin); 10, P. p e r s e a e, aedeagus in lateral aspect, Citrus limon, Florida (Morrill); 11, P. citricolus Costa Lima, aedeagus in lateral aspect, Citrus sinensis, Madeira (Aguiar); 12, P. a n c o r a Martin, aedeagus of holotype, in dorsal aspect,? Lasianthaea sp., Belize (Martin); 13, P. ancora Martin, aedeagus of paratype, in lateral aspect,? Lasianthaea sp., Belize (Martin); 14, Paraleyrodes sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15, Paraleyrodes sp., aedeagus, in dorsal aspect, Citrus sp., Florida (Dowell); 16, Paraleyrodes sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be P. p ers eae." httpUri="https://zenodo.org/record/171849/files/figure.png" pageId="10" pageNumber="45">Figs 8–9</figureCitation>
), and reared from puparia that match the
<typeStatus id="724B8864FFBD7E1CFD6AFA80FCEBFAE1" box="[709,817,1284,1310]" pageId="10" pageNumber="45" type="lectotype">lectotype</typeStatus>
puparium of
<taxonomicName id="6AF04D45FFBD7E1CFC71FA80FB66FAE2" box="[990,1212,1284,1309]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBD7E1CFC71FA80FB66FAE2" box="[990,1212,1284,1309]" italics="true" pageId="10" pageNumber="45">Aleurodes perseae</emphasis>
</taxonomicName>
, is quite different in structure from that which
<bibRefCitation id="C9614B37FFBD7E1CFD68FAA8FC7CFAB9" author="Quaintance" box="[711,934,1324,1350]" pageId="10" pageNumber="47" refString="Quaintance, A. L. (1909) A new genus of Aleyrodidae, with remarks on Aleyrodes nubifera Berger and Aleyrodes citri Riley and Howard. Technical Series, US Department of Agriculture Bureau of Entomology, 12, 169 - 174." type="journal article" year="1909">Quaintance (1909)</bibRefCitation>
, and subsequently
<bibRefCitation id="C9614B37FFBD7E1CFB29FAA8FE70FA91" author="Quaintance" pageId="10" pageNumber="47" refString="Quaintance, A. L. &amp; Baker, A. C. (1913) Classification of the Aleyrodidae Part I. Technical Series, US Department of Agriculture Bureau of Entomology, 27, 1 - 93." type="journal article" year="1913">
Quaintance &amp;
<collectingCountry id="D5E77656FFBD7E1CFEA7FAD0FE94FA91" box="[264,334,1364,1390]" name="United States Pacific Island Wildlife Refuges" pageId="10" pageNumber="45">Baker</collectingCountry>
(1913)
</bibRefCitation>
, mistook for
<emphasis id="9F84EAD4FFBD7E1CFDFFFAD0FD0EFA92" box="[592,724,1364,1389]" italics="true" pageId="10" pageNumber="45">
<taxonomicName id="6AF04D45FFBD7E1CFDFFFAD0FD15FA92" box="[592,719,1364,1389]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">P. perseae</taxonomicName>
.
</emphasis>
This finding, combined with the illustration of a puparium with six pairs of abdominal compound pores (
<bibRefCitation id="C9614B37FFBD7E1CFC0CFAF8FBA3FA69" author="Quaintance" box="[931,1145,1404,1430]" pageId="10" pageNumber="47" refString="Quaintance, A. L. (1909) A new genus of Aleyrodidae, with remarks on Aleyrodes nubifera Berger and Aleyrodes citri Riley and Howard. Technical Series, US Department of Agriculture Bureau of Entomology, 12, 169 - 174." type="journal article" year="1909">Quaintance, 1909</bibRefCitation>
), indicates that the redescription of “
<taxonomicName id="6AF04D45FFBD7E1CFDA8FA20FD5BFA42" box="[519,641,1444,1469]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBD7E1CFDA8FA20FD5BFA42" box="[519,641,1444,1469]" italics="true" pageId="10" pageNumber="45">P. perseae</emphasis>
</taxonomicName>
” involved a different species entirely. Unfortunately, the detail of the 1909 description is still insufficient to lead to any conclusion about the species concerned.
</paragraph>
<paragraph id="AD4F36C6FFBD7E1CFE97F998FBFFF8B2" blockId="10.[264,1325,284,1870]" pageId="10" pageNumber="45">
The aedeagus of true
<taxonomicName id="6AF04D45FFBD7E1CFDE7F998FD1FF9CA" box="[584,709,1564,1589]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBD7E1CFDE7F998FD1FF9CA" box="[584,709,1564,1589]" italics="true" pageId="10" pageNumber="45">P. perseae</emphasis>
</taxonomicName>
is illustrated here (probably for the first time) in figures 8–10. This should allow for confident identification of this species when puparia are also present. However, the male aedeagus of
<taxonomicName id="6AF04D45FFBD7E1CFCF2F9E8FC02F97A" box="[861,984,1644,1669]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBD7E1CFCF2F9E8FC02F97A" box="[861,984,1644,1669]" italics="true" pageId="10" pageNumber="45">P. perseae</emphasis>
</taxonomicName>
is very similar to that of
<taxonomicName id="6AF04D45FFBD7E1CFABAF9E8FDDDF951" authority="Costa Lima" authorityName="Costa Lima" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="citricolus">
<emphasis id="9F84EAD4FFBD7E1CFABAF9E8FEADF952" italics="true" pageId="10" pageNumber="45">P. citricolus</emphasis>
Costa Lima
</taxonomicName>
(
<figureCitation id="35CB2A43FFBD7E1CFDB8F910FDB6F951" box="[535,620,1684,1710]" captionStart="FIGURES 8 – 16" captionStartId="7.[264,382,1333,1357]" captionTargetBox="[299,1272,320,1308]" captionTargetId="figure@7.[289,1297,302,1337]" captionTargetPageId="7" captionText="FIGURES 8 – 16, Paraleyrodes spp, male genitalia. 8, P. p e r s e a e, genital segment, with aedeagus in oblique aspect, Persea americana, Nicaragua (Martin); 9, P. p e r s ea e, aedeagus in lateral aspect, Persea americana, Nicaragua (Martin); 10, P. p e r s e a e, aedeagus in lateral aspect, Citrus limon, Florida (Morrill); 11, P. citricolus Costa Lima, aedeagus in lateral aspect, Citrus sinensis, Madeira (Aguiar); 12, P. a n c o r a Martin, aedeagus of holotype, in dorsal aspect,? Lasianthaea sp., Belize (Martin); 13, P. ancora Martin, aedeagus of paratype, in lateral aspect,? Lasianthaea sp., Belize (Martin); 14, Paraleyrodes sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15, Paraleyrodes sp., aedeagus, in dorsal aspect, Citrus sp., Florida (Dowell); 16, Paraleyrodes sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be P. p ers eae." httpUri="https://zenodo.org/record/171849/files/figure.png" pageId="10" pageNumber="45">Fig. 11</figureCitation>
), which typically exhibits six, somewhat digitiform, ventral processes but which otherwise shares the same lateral profile. That
<taxonomicName id="6AF04D45FFBD7E1CFBFBF938FB0BF92A" box="[1108,1233,1724,1749]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">
<emphasis id="9F84EAD4FFBD7E1CFBFBF938FB0BF92A" box="[1108,1233,1724,1749]" italics="true" pageId="10" pageNumber="45">P. perseae</emphasis>
</taxonomicName>
and
<taxonomicName id="6AF04D45FFBD7E1CFABAF938FEADF902" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="citricolus">
<emphasis id="9F84EAD4FFBD7E1CFABAF938FEADF902" italics="true" pageId="10" pageNumber="45">P. citricolus</emphasis>
</taxonomicName>
are particularly closely related is also indicated by the structure of the inner lumen of the large compound pores (
<figureCitation id="35CB2A43FFBD7E1CFD10F888FCDDF8D9" box="[703,775,1804,1830]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="10" pageNumber="45">Fig. 7</figureCitation>
), even though the large compound pores of
<taxonomicName id="6AF04D45FFBD7E1CFABAF888FEACF8B2" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="citricolus">
<emphasis id="9F84EAD4FFBD7E1CFABAF888FEACF8B2" italics="true" pageId="10" pageNumber="45">P. citricolus</emphasis>
</taxonomicName>
are usually over twice the diameter of those of
<emphasis id="9F84EAD4FFBD7E1CFC0AF8B0FBFFF8B2" box="[933,1061,1844,1869]" italics="true" pageId="10" pageNumber="45">
<taxonomicName id="6AF04D45FFBD7E1CFC0AF8B0FBFBF8B2" box="[933,1057,1844,1869]" class="Insecta" family="Aleyrodidae" genus="Aleurodes" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="perseae">P. perseae</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="AD4F36C6FFBC7E1DFE97FE98FB7BFD61" blockId="11.[264,1323,284,670]" pageId="11" pageNumber="46">
A final observation is that this is the second example of a species of
<taxonomicName id="6AF04D45FFBC7E1DFB21FE98FAF1FECA" box="[1166,1323,284,309]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBC7E1DFB21FE98FAF1FECA" box="[1166,1323,284,309]" italics="true" pageId="11" pageNumber="46">Paraleyrodes</emphasis>
</taxonomicName>
being incorrectly identified because described males were not associated with puparia. For many years,
<taxonomicName id="6AF04D45FFBC7E1DFE0AFEE8FCEDFE79" authority="Martin (2001)" authorityName="Martin" authorityYear="2001" box="[421,823,364,390]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="pseudonaranjae">
<emphasis id="9F84EAD4FFBC7E1DFE0AFEE8FD59FE7A" box="[421,643,364,389]" italics="true" pageId="11" pageNumber="46">P. pseudonaranjae</emphasis>
<bibRefCitation id="C9614B37FFBC7E1DFD21FEE8FCEDFE79" author="Martin" box="[654,823,364,390]" pageId="11" pageNumber="46" refString="Martin, J. H. (2001) Description of an invasive new species of Neotropical aleurodicine whitefly (Hemiptera: Aleyrodidae) - a case of complete or partial misidentification? Bulletin of Entomological Research, 91, 101 - 107." type="journal article" year="2001">Martin (2001)</bibRefCitation>
</taxonomicName>
was misidentified as
<taxonomicName id="6AF04D45FFBC7E1DFBEAFEE8FE83FE51" authority="Dozier (1927)" authorityName="Dozier" authorityYear="1927" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="naranjae">
<emphasis id="9F84EAD4FFBC7E1DFBEAFEE8FB0AFE7A" box="[1093,1232,364,389]" italics="true" pageId="11" pageNumber="46">P. naranjae</emphasis>
<bibRefCitation id="C9614B37FFBC7E1DFB74FEE8FE83FE51" author="Dozier" pageId="11" pageNumber="46" refString="Dozier, H. L. (1927) An undescribed white fly attacking citrus in Porto Rico. Journal of Agricultural Research, 34, 853 - 855." type="journal article" year="1927">Dozier (1927)</bibRefCitation>
</taxonomicName>
, based on its puparia. However, the males in invasive populations did not match the two male
<typeStatus id="724B8864FFBC7E1DFE23FE38FE29FE29" box="[396,499,444,470]" pageId="11" pageNumber="46" type="syntype">syntypes</typeStatus>
from Dozier’s original sample of
<taxonomicName id="6AF04D45FFBC7E1DFC1AFE38FB9FFE2A" box="[949,1093,444,469]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="naranjae">
<emphasis id="9F84EAD4FFBC7E1DFC1AFE38FB9FFE2A" box="[949,1093,444,469]" italics="true" pageId="11" pageNumber="46">P. naranjae</emphasis>
</taxonomicName>
, and the invasive populations proved to be a different and undescribed species. The author’s own field experience indicates that, in their native habitat, puparia of
<taxonomicName id="6AF04D45FFBC7E1DFC55FD88FB4DFDDA" box="[1018,1175,524,549]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFBC7E1DFC55FD88FB4DFDDA" box="[1018,1175,524,549]" italics="true" pageId="11" pageNumber="46">Paraleyrodes</emphasis>
</taxonomicName>
species are frequently found with adults of different species feeding on the same individual plant, making the association of adults with puparia difficult. Conversely, invasive populations tend to be taxonomically useful, for they usually comprise only single species.
</paragraph>
</subSubSection>
</treatment>
</document>

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Quaintance
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<taxonomicName id="6AF04D45FFB37E12FEA7FAADFD48FAC0" ID-CoL="6F9B" authority="Quaintance, 1909: 169" authorityName="Quaintance" authorityPageNumber="169" authorityYear="1909" box="[264,658,1320,1344]" class="Insecta" family="Aleyrodidae" genus="Paraleyrodes" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="9F84EAD4FFB37E12FEA7FAADFE4DFABF" box="[264,407,1321,1344]" italics="true" pageId="4" pageNumber="39">Paraleyrodes</emphasis>
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<bibRefCitation id="C9614B37FFB37E12FE31FAACFD85FAC0" author="Quaintance" box="[414,607,1320,1343]" pageId="4" pageNumber="47" refString="Quaintance, A. L. (1909) A new genus of Aleyrodidae, with remarks on Aleyrodes nubifera Berger and Aleyrodes citri Riley and Howard. Technical Series, US Department of Agriculture Bureau of Entomology, 12, 169 - 174." type="journal article" year="1909">Quaintance, 1909</bibRefCitation>
: 169
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–170.
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species
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<emphasis id="9F84EAD4FFB37E12FCC7FAADFBF4FABF" box="[872,1070,1321,1344]" italics="true" pageId="4" pageNumber="39">Aleurodes perseae</emphasis>
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: 32
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– 33, by original designation and monotypy.
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GENERIC DIAGNOSIS.
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comprises species that are physically much smaller than most other members of the
<taxonomicName id="6AF04D45FFB37E12FD72FA3DFC5CFA2C" box="[733,902,1465,1491]" class="Insecta" family="Aleyrodidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="subFamily" subFamily="Aleurodicinae">Aleurodicinae</taxonomicName>
, adults and puparia often less than
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in length.
<emphasis id="9F84EAD4FFB37E12FE6EFA65FDFCFA04" bold="true" box="[449,550,1505,1531]" pageId="4" pageNumber="39">Puparia</emphasis>
: abdomen with 5 or 6 pairs of compound pores, of which the anteriormost 1 or 2 pairs are much smaller than the remaining 4 abdominal pairs and the cephalic pair (
<figureCitation id="35CB2A43FFB37E12FE02F9B5FE28F9B4" box="[429,498,1585,1611]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="4" pageNumber="39">Fig. 5</figureCitation>
); two pairs of cicatrices present on thorax (scars of compound pores in third­instar, one shown in
<figureCitation id="35CB2A43FFB37E12FDF2F9DDFD71F98C" box="[605,683,1625,1651]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="4" pageNumber="39">Fig. 5</figureCitation>
); a pair of submedian cephalic setae present, but corresponding thoracic setal pairs absent; outer submargin with a row of 14 pairs of hairlike setae, including the nominal caudal pair; lingula extending beyond boundary of vasiform orifice, bearing 4 stout setae (
<figureCitation id="35CB2A43FFB37E12FD60F955FCCCF914" box="[719,790,1745,1771]" captionStart="FIGURES 5 – 7" captionStartId="6.[264,382,1707,1731]" captionTargetBox="[359,1267,348,1668]" captionTargetId="figure@6.[305,1282,329,1679]" captionTargetPageId="6" captionText="FIGURES 5 – 7, Paraleyrodes perseae (Quaintance), puparium. 5, puparial margin, outersubmarginal glands, submarginal setae, large abdominal compound pores, small compound pore on abdominal segment III, bright pores and adjacent spines, and posterior thoracic cicatrice; 6, posterodorsal detail, with vasiform orifice, operculum, lingula, eighth abdominal setae, outersubmarginal glands and submarginal &amp; posterior marginal setae; 7, stylised drawing of large compound pore." httpUri="https://zenodo.org/record/171848/files/figure.png" pageId="4" pageNumber="39">Fig. 6</figureCitation>
).
<emphasis id="9F84EAD4FFB37E12FC82F955FC5AF914" bold="true" box="[813,896,1745,1771]" pageId="4" pageNumber="39">Adults</emphasis>
: females have 4 articulated antennal segments, the distal 2 segments sparsely provided with small, rounded, sensoria; males have only 3 articulated antennal segments, the very long distal segment (
<figureCitation id="35CB2A43FFB37E12FBD6F8A5FB1BF8C4" box="[1145,1217,1825,1851]" captionStart="FIGURES 1 – 2" captionStartId="2.[264,382,1770,1794]" captionTargetBox="[264,1322,329,1741]" captionTargetId="figure@2.[264,1323,329,1761]" captionTargetPageId="2" captionText="FIGURES 1 – 2, Paraleyrodes spp. ex­Belize. 1, oviposition “ nest ” with eggs visible inside; 2, adult male, showing characteristic long single­segment antennal flagellum." httpUri="https://zenodo.org/record/171846/files/figure.png" pageId="4" pageNumber="39">Fig. 2</figureCitation>
) heavily sensoriate and often appearing reddish in life; males have complex aedeagal apices (e.g.
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<figureCitation id="35CB2A43FFB27E13FEA7FE98FE5AFEC9" box="[264,384,284,310]" captionStart="FIGURES 8 – 16" captionStartId="7.[264,382,1333,1357]" captionTargetBox="[299,1272,320,1308]" captionTargetId="figure@7.[289,1297,302,1337]" captionTargetPageId="7" captionText="FIGURES 8 – 16, Paraleyrodes spp, male genitalia. 8, P. p e r s e a e, genital segment, with aedeagus in oblique aspect, Persea americana, Nicaragua (Martin); 9, P. p e r s ea e, aedeagus in lateral aspect, Persea americana, Nicaragua (Martin); 10, P. p e r s e a e, aedeagus in lateral aspect, Citrus limon, Florida (Morrill); 11, P. citricolus Costa Lima, aedeagus in lateral aspect, Citrus sinensis, Madeira (Aguiar); 12, P. a n c o r a Martin, aedeagus of holotype, in dorsal aspect,? Lasianthaea sp., Belize (Martin); 13, P. ancora Martin, aedeagus of paratype, in lateral aspect,? Lasianthaea sp., Belize (Martin); 14, Paraleyrodes sp., aedeagus, in dorsal aspect, (no host data) Florida (Williams); 15, Paraleyrodes sp., aedeagus, in dorsal aspect, Citrus sp., Florida (Dowell); 16, Paraleyrodes sp, aedeagus and claspers in dorsal aspect, Florida, reproduced from Quaintance (1909) where this was incorrectly stated to be P. p ers eae." httpUri="https://zenodo.org/record/171849/files/figure.png" pageId="5" pageNumber="40">Figs 8–16</figureCitation>
) which are species­diagnostic (almost all other known whiteflies have simple aedeagal apices).
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<paragraph id="AD4F195BA841FFE6D6FFFBD68C57FBC4" blockId="18.[159,723,1101,1349]" box="[159,274,1101,1125]" pageId="18" pageNumber="19">Catalogue</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A841FFE6D6FFFBE98F81FAE4" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="AD4F195BA841FFE6D6FFFBE98F13FB2B" blockId="18.[159,723,1101,1349]" box="[159,598,1138,1162]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="8DE03E75A841FFE6D6FFFBE98F13FB2B" box="[159,598,1138,1162]" pageId="18" pageNumber="19">
<taxonomicName id="6AF062D8A841FFE6D6FFFBE98CCCFB2B" ID-CoL="8MFK7" authorityName="BATES Compared" authorityYear="1892" box="[159,393,1138,1162]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A841FFE6D6FFFBE98CCCFB2B" box="[159,393,1138,1162]" italics="true" pageId="18" pageNumber="19">Clivina costulipennis</emphasis>
</taxonomicName>
;
<treatmentCitation id="2C513F4AA841FFE6D7F8FBE98F0BFB2B" author="Bates, H. W." box="[408,590,1138,1162]" page="281" pageId="18" pageNumber="19" year="1892">
<bibRefCitation id="C96164AAA841FFE6D7F8FBE98F0BFB2B" author="Bates, H. W." box="[408,590,1138,1162]" pageId="18" pageNumber="19" pagination="267 - 428" refId="ref27786" refString="Bates, H. W. (1892): Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. - Annali del Museo Civico di Storia Naturale di Genova 32: 267 - 428." type="journal article" year="1892">Bates 1892: 281</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA841FFE6D6FFFB038F97FB11" blockId="18.[159,723,1101,1349]" box="[159,722,1176,1200]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="8DE03E75A841FFE6D6FFFB038F97FB11" box="[159,722,1176,1200]" pageId="18" pageNumber="19">
<taxonomicName id="6AF062D8A841FFE6D6FFFB038C91FB11" ID-CoL="8MFK7" authority="BATES" authorityName="BATES Compared" authorityYear="1892" box="[159,468,1176,1200]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A841FFE6D6FFFB038CCCFB11" box="[159,393,1176,1200]" italics="true" pageId="18" pageNumber="19">Clivina costulipennis</emphasis>
BATES
</taxonomicName>
;
<treatmentCitation id="2C513F4AA841FFE6D782FB038F8FFB11" author="Andrewes, H. E." box="[482,714,1176,1200]" page="373" pageId="18" pageNumber="19" year="1926">
<bibRefCitation id="C96164AAA841FFE6D782FB038F8FFB11" author="Andrewes, H. E." box="[482,714,1176,1200]" pageId="18" pageNumber="19" pagination="371 - 381" refId="ref27411" refString="Andrewes, H. E. (1926): Papers on Oriental Carabidae. XVII. - The Annals and Magazine of Natural History Series 9 17: 371 - 381." type="journal article" year="1926">Andrewes 1926: 373</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA841FFE6D6FFFB268F96FB74" blockId="18.[159,723,1101,1349]" box="[159,723,1213,1237]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="8DE03E75A841FFE6D6FFFB268F96FB74" box="[159,723,1213,1237]" pageId="18" pageNumber="19">
<taxonomicName id="6AF062D8A841FFE6D6FFFB268C91FB74" ID-CoL="8MFK7" authority="BATES" authorityName="BATES Compared" authorityYear="1892" box="[159,468,1213,1237]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A841FFE6D6FFFB268CCCFB74" box="[159,393,1213,1237]" italics="true" pageId="18" pageNumber="19">Clivina costulipennis</emphasis>
BATES
</taxonomicName>
;
<treatmentCitation id="2C513F4AA841FFE6D782FB268F89FB74" author="Andrewes, H. E." box="[482,716,1213,1237]" page="366" pageId="18" pageNumber="19" year="1929">
<bibRefCitation id="C96164AAA841FFE6D782FB268F89FB74" author="Andrewes, H. E." box="[482,716,1213,1237]" pageId="18" pageNumber="19" refId="ref27444" refString="Andrewes, H. E. (1929): The fauna of British India, including Ceylon and Burma. Coleoptera, Carabidae, Vol. I., Carabinae. - Taylor &amp; Francis, London, 431 pp." type="book" year="1929">Andrewes 1929: 366</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA841FFE6D6FFFB798F95FB5B" blockId="18.[159,723,1101,1349]" box="[159,720,1250,1274]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="8DE03E75A841FFE6D6FFFB798F95FB5B" box="[159,720,1250,1274]" pageId="18" pageNumber="19">
<taxonomicName id="6AF062D8A841FFE6D6FFFB798C91FB5B" ID-CoL="8MFK7" authority="BATES" authorityName="BATES Compared" authorityYear="1892" box="[159,468,1250,1274]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A841FFE6D6FFFB798CCCFB5B" box="[159,393,1250,1274]" italics="true" pageId="18" pageNumber="19">Clivina costulipennis</emphasis>
BATES
</taxonomicName>
;
<treatmentCitation id="2C513F4AA841FFE6D782FB798F8CFB5B" author="Andrewes, H. E." box="[482,713,1250,1274]" page="113" pageId="18" pageNumber="19" year="1930">
<bibRefCitation id="C96164AAA841FFE6D782FB798F8CFB5B" author="Andrewes, H. E." box="[482,713,1250,1274]" pageId="18" pageNumber="19" refId="ref27486" refString="Andrewes, H. E. (1930): Catalogue of Indian Insects. Part 18 - Carabidae. - 389 pp. Government of India Central Publication Branch, Calcutta." type="book" year="1930">Andrewes 1930: 113</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA841FFE6D6FFFA938FE8FA81" blockId="18.[159,723,1101,1349]" box="[159,685,1288,1312]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="8DE03E75A841FFE6D6FFFA938FE8FA81" box="[159,685,1288,1312]" pageId="18" pageNumber="19">
<taxonomicName id="6AF062D8A841FFE6D6FFFA938C91FA81" ID-CoL="8MFK7" authority="BATES" authorityName="BATES Compared" authorityYear="1892" box="[159,468,1288,1312]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A841FFE6D6FFFA938CCCFA81" box="[159,393,1288,1312]" italics="true" pageId="18" pageNumber="19">Clivina costulipennis</emphasis>
BATES
</taxonomicName>
;
<treatmentCitation id="2C513F4AA841FFE6D782FA938FE1FA81" author="Lorenz, W." box="[482,676,1288,1312]" page="131" pageId="18" pageNumber="19" year="1998">
<bibRefCitation id="C96164AAA841FFE6D782FA938FE1FA81" author="Lorenz, W." box="[482,676,1288,1312]" pageId="18" pageNumber="19" refId="ref28004" refString="Lorenz, W. (1998): Systematic list of extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). - Tutzing, printed by the author. 502 pp." type="book" year="1998">Lorenz 1998: 131</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA841FFE6D6FFFAB68F81FAE4" blockId="18.[159,723,1101,1349]" box="[159,708,1325,1349]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="8DE03E75A841FFE6D6FFFAB68F81FAE4" box="[159,708,1325,1349]" pageId="18" pageNumber="19">
<taxonomicName id="6AF062D8A841FFE6D6FFFAB68C91FAE4" ID-CoL="8MFK7" authority="BATES" authorityName="BATES Compared" authorityYear="1892" box="[159,468,1325,1349]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A841FFE6D6FFFAB68CCCFAE4" box="[159,393,1325,1349]" italics="true" pageId="18" pageNumber="19">Clivina costulipennis</emphasis>
BATES
</taxonomicName>
;
<treatmentCitation id="2C513F4AA841FFE6D782FAB68FF9FAE4" author="Balkenohl, M." box="[482,700,1325,1349]" page="14" pageId="18" pageNumber="19" year="2001">
<bibRefCitation id="C96164AAA841FFE6D782FAB68FF9FAE4" author="Balkenohl, M." box="[482,700,1325,1349]" pageId="18" pageNumber="19" refId="ref27660" refString="Balkenohl, M. (2001): Key and Catalogue of the tribe Clivinini from the Oriental realm, with revisions of the genera Thliboclivina KULT, and Trilophidius JEANNEL (Insecta, Coleoptera, Carabidae, Scarititae, Clivinini). - 83 pp., Sofia, Moscow (Pensoft Series Faunistica 21)." type="book" year="2001">Balkenohl 2001: 14</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A841FFE7D6FFFAE38C0AFC24" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="AD4F195BA841FFE7D6FFFAE38C66FF44" blockId="18.[159,1043,1397,1573]" lastBlockId="19.[121,1007,93,901]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Type material:
<materialsCitation id="1D981306A841FFE7D735FAE38C5AFF44" ID-GBIF-Occurrence="3497007326" collectionCode="MCSNG" collectorName="L. Fea." lastPageId="19" lastPageNumber="20" location="Palon" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Bago" typeStatus="holotype">
<typeStatus id="724BA7F9A841FFE6D735FAE38C86FA31" box="[341,451,1400,1424]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A841FFE6D7B4FAEE8CADFA31" box="[468,488,1397,1424]" pageId="18" pageNumber="19" type="male">♂</specimenCount>
, labels: brown with black frame, black printed: “
<location id="A82F4F80A841FFE6D6B7FA068C59FA14" LSID="urn:lsid:plazi:treatment:2559A84DA841FFE2D6FFFB998E71FC0B:A82F4F80A841FFE6D6B7FA068C59FA14" box="[215,284,1437,1461]" name="Palon" pageId="18" pageNumber="19" stateProvince="Bago">Palon</location>
(
<location id="A82F4F80A841FFE6D74DFA068C29FA14" LSID="urn:lsid:plazi:treatment:2559A84DA841FFE2D6FFFB998E71FC0B:A82F4F80A841FFE6D74DFA068C29FA14" box="[301,364,1437,1461]" name="Pegu" pageId="18" pageNumber="19" stateProvince="Bago">Pegú</location>
) L.Fea.VIII.
<date id="D94E3F9BA841FFE6D78FFA068F6CFA14" box="[495,553,1437,1461]" pageId="18" pageNumber="19" value="1987-09">IX.87</date>
” / brown, handwritten in black: “
<taxonomicName id="6AF062D8A841FFE6D5A2FA068CDEFA7B" authority="Bates" authorityName="BATES Compared" authorityYear="1892" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costulipennis">Clivina costulipennis Bates</taxonomicName>
” / yellowish-brown, handwritten in black: “
<taxonomicName id="6AF062D8A841FFE6D5BCFA598C73F9A1" authorityName="BATES Compared" authorityYear="1892" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costulipennis">Cliv. costulipennis</taxonomicName>
(typus!) Bates” / white, red-printed with red frame: “TYPUS” / brown, printed in black: “Mus. Civ. Genova” / white with black frame, handwritten in black: “
<taxonomicName id="6AF062D8A840FFE7D727FFC68F6AFFD4" authority="Bates" authorityName="BATES Compared" authorityYear="1892" box="[327,559,93,117]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="costulipennis">costulipennis Bates</taxonomicName>
” / red with black frame, black printed “HOLOTYPUS” and handwritten “
<taxonomicName id="6AF062D8A840FFE7D792FF198E34FF3B" authority="Bates, 1892" authorityName="BATES Compared" authorityYear="1892" box="[498,881,130,154]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="costulipennis">
Clivina costulipennis
<bibRefCitation id="C96164AAA840FFE7D489FF198E34FF3B" author="Bates, H. W." box="[745,881,130,154]" pageId="19" pageNumber="20" pagination="267 - 428" refId="ref27786" refString="Bates, H. W. (1892): Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. - Annali del Museo Civico di Storia Naturale di Genova 32: 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
</taxonomicName>
” Additional label: white, black printed: “Genitalpräparat wasserlÖslich M. Balkenohl
<date id="D94E3F9BA840FFE7D619FF568DE7FF44" box="[121,162,205,229]" pageId="19" pageNumber="20" value="2007-01">I 07</date>
” (
<collectionCode id="CBE1819EA840FFE7D6DBFF568C5FFF44" box="[187,282,205,229]" collectionName="MCSNG" pageId="19" pageNumber="20">MCSNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="AD4F195BA840FFE7D619FF698C0AFC24" blockId="19.[121,1007,93,901]" pageId="19" pageNumber="20">
Additional material:
<materialsCitation id="1D981306A840FFE7D71CFF698E56FEF4" ID-GBIF-Occurrence="3497007307" collectingDate="1990-05-04" collectingDateMax="1990-06-19" collectingDateMin="1990-05-04" collectionCode="CBM-ZSM, CBB" collectorName="Malicky / Clivina" country="Thailand" determinerName="M. Baehr'" location="Doi Inthanon" municipality="Doi Inthanon" pageId="19" pageNumber="20" specimenCount="6" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="BBF6D2D2A840FFE7D71CFF698CEAFEAB" box="[380,431,239,266]" pageId="19" pageNumber="20" type="male">2 ♂</specimenCount>
,
<specimenCount id="BBF6D2D2A840FFE7D7A7FF698CBFFEAB" box="[455,506,239,266]" pageId="19" pageNumber="20" type="female">1 ♀</specimenCount>
,
<specimenCount id="BBF6D2D2A840FFE7D473FF698FF6FEAB" box="[531,691,242,266]" pageId="19" pageNumber="20" type="generic">3 specimens</specimenCount>
:
<collectingCountry id="D5E759CBA840FFE7D4A8FF698E74FEAB" box="[712,817,242,266]" name="Thailand" pageId="19" pageNumber="20">Thailand</collectingCountry>
,
<collectingMunicipality id="4D2B8321A840FFE7D525FF698EACFEAB" box="[837,1001,242,266]" country="Thailand" name="Doi Inthanon" pageId="19" pageNumber="20">Doi Inthanon</collectingMunicipality>
,
<date id="D94E3F9BA840FFE7D619FE838C47FE91" box="[121,258,280,304]" pageId="19" pageNumber="20" value="1990-05-04" valueMax="1990-05-06" valueMin="1990-05-04">
<collectingDate id="C90AC673A840FFE7D619FE838C47FE91" box="[121,258,280,304]" pageId="19" pageNumber="20" value="1990-05-04" valueMax="1990-05-06" valueMin="1990-05-04">4.–6.5.1990</collectingDate>
</date>
,
<date id="D94E3F9BA840FFE7D772FE838CDDFE91" box="[274,408,280,304]" pageId="19" pageNumber="20" value="1990-05-08" valueMax="1990-05-09" valueMin="1990-05-08">
<collectingDate id="C90AC673A840FFE7D772FE838CDDFE91" box="[274,408,280,304]" pageId="19" pageNumber="20" value="1990-05-08" valueMax="1990-05-09" valueMin="1990-05-08">8.–9.5.1990</collectingDate>
</date>
,
<date id="D94E3F9BA840FFE7D7C9FE838F0FFE91" box="[425,586,280,304]" pageId="19" pageNumber="20" value="1990-05-23" valueMax="1990-05-25" valueMin="1990-05-23">
<collectingDate id="C90AC673A840FFE7D7C9FE838F0FFE91" box="[425,586,280,304]" pageId="19" pageNumber="20" value="1990-05-23" valueMax="1990-05-25" valueMin="1990-05-23">23.–25.5.1990</collectingDate>
</date>
,
<date id="D94E3F9BA840FFE7D43BFE838FB9FE91" box="[603,764,280,304]" pageId="19" pageNumber="20" value="1990-06-12" valueMax="1990-06-19" valueMin="1990-06-12">
<collectingDate id="C90AC673A840FFE7D43BFE838FB9FE91" box="[603,764,280,304]" pageId="19" pageNumber="20" value="1990-06-12" valueMax="1990-06-19" valueMin="1990-06-12">12.–19.6.1990</collectingDate>
</date>
, leg.
<collectorName id="00057C8DA840FFE7D520FE838DD0FEF4" pageId="19" pageNumber="20">
Malicky /
<taxonomicName id="6AF062D8A840FFE7D5D5FE838DD0FEF4" authority="BAT." authorityName="BATES Compared" authorityYear="1892" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A840FFE7D5D5FE838DD0FEF4" italics="true" pageId="19" pageNumber="20">Clivina</emphasis>
</taxonomicName>
</collectorName>
costulipennis BAT. det.
<determinerName id="5893F59FA840FFE7D7D2FEA68F58FEF4" box="[434,541,317,341]" pageId="19" pageNumber="20">M.Baehr’</determinerName>
94 (
<collectionCode id="CBE1819EA840FFE7D42CFEA68F81FEF4" box="[588,708,317,341]" pageId="19" pageNumber="20">CBM-ZSM</collectionCode>
,
<collectionCode id="CBE1819EA840FFE7D4B3FEA68E4CFEF4" box="[723,777,317,341]" collectionName="CBB" pageId="19" pageNumber="20">CBB</collectionCode>
);
</materialsCitation>
<materialsCitation id="1D981306A840FFE7D57EFEA68F9FFE01" ID-GBIF-Occurrence="3497007328" collectingDate="2013-11-13" collectionCode="BMNH" collectorName="H. E. Andrewes" location="Penang" municipality="Penang" pageId="19" pageNumber="20" specimenCount="1">
<specimenCount id="BBF6D2D2A840FFE7D57EFEA68EEEFEF4" box="[798,939,317,341]" pageId="19" pageNumber="20" type="generic">1 specimen</specimenCount>
:
<collectingMunicipality id="4D2B8321A840FFE7D5DAFEA68DEDFEDB" pageId="19" pageNumber="20">Penang</collectingMunicipality>
,G.E.Bryant.
<date id="D94E3F9BA840FFE7D755FEF98CD7FEDB" box="[309,402,354,378]" pageId="19" pageNumber="20" value="2013-11-13">
<collectingDate id="C90AC673A840FFE7D755FEF98CD7FEDB" box="[309,402,354,378]" pageId="19" pageNumber="20" value="2013-11-13">13.XI.13</collectingDate>
</date>
/
<collectorName id="00057C8DA840FFE7D7D1FEF98F15FEDB" box="[433,592,354,378]" pageId="19" pageNumber="20">
H.E.
<taxonomicName id="6AF062D8A840FFE7D7B5FEF98F15FEDB" box="[469,592,354,378]" pageId="19" pageNumber="1" rank="subGenus" status="subgen. nov." subGenus="Andrewes">Andrewes</taxonomicName>
</collectorName>
Coll. B.M.1945–97. /
<taxonomicName id="6AF062D8A840FFE7D531FEF98CE6FE01" authority="BATES Compared" authorityName="BATES Compared" authorityYear="1892" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A840FFE7D531FEF98D91FE01" italics="true" pageId="19" pageNumber="20">Clivina costulipennis</emphasis>
BATES Compared
</taxonomicName>
with type H.E.A. (
<collectionCode id="CBE1819EA840FFE7D41EFE138F95FE01" box="[638,720,392,416]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="19" pageNumber="20">BMNH</collectionCode>
);
</materialsCitation>
<materialsCitation id="1D981306A840FFE7D483FE138C76FD94" ID-GBIF-Occurrence="3497007335" collectingDate="1996-05-09" collectingDateMax="1996-05-17" collectingDateMin="1996-05-09" collectionCode="NHMW" collectorName="Jendek &amp; Sausa" elevation="1100" latitude="25.426666" location="Meghalaya" longLatPrecision="124" longitude="90.325" pageId="19" pageNumber="20" specimenCount="2">
<specimenCount id="BBF6D2D2A840FFE7D483FE138E28FE01" box="[739,877,392,416]" pageId="19" pageNumber="20" type="generic">1 specimen</specimenCount>
: Malay Pa. Perak / 58J81 / Doherty / Fry Coll 1905.100. /
<taxonomicName id="6AF062D8A840FFE7D4CCFE368DABFE4B" authority="BATES Compared" authorityName="BATES Compared" authorityYear="1892" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A840FFE7D4CCFE368EDBFE64" box="[684,926,429,453]" italics="true" pageId="19" pageNumber="20">Clivina costulipennis</emphasis>
BATES Compared
</taxonomicName>
with type H.E.A.;
<specimenCount id="BBF6D2D2A840FFE7D7A7FE498F15FE4B" box="[455,592,466,490]" pageId="19" pageNumber="20" type="generic">1 specimen</specimenCount>
: NE-INDIA:
<location id="A82F4F80A840FFE7D4B6FE498E16FE4B" LSID="urn:lsid:plazi:treatment:2559A84DA841FFE2D6FFFB998E71FC0B:A82F4F80A840FFE7D4B6FE498E16FE4B" box="[726,851,466,490]" latitude="25.426666" longLatPrecision="124" longitude="90.325" name="Meghalaya" pageId="19" pageNumber="20">Meghalaya</location>
W
<location id="A82F4F80A840FFE7D51CFE498DF8FDB1" LSID="urn:lsid:plazi:treatment:2559A84DA841FFE2D6FFFB998E71FC0B:A82F4F80A840FFE7D51CFE498DF8FDB1" latitude="25.426666" longLatPrecision="124" longitude="90.325" name="Garo Hills Norek" pageId="19" pageNumber="20">Garo Hills Norek</location>
NP; ca.
<quantity id="6A08B4BEA840FFE7D74EFE638CC2FDB1" box="[302,391,504,528]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="20" unit="m" value="1100.0">
<elevation id="26DDFE68A840FFE7D74EFE638CC2FDB1" box="[302,391,504,528]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="20" unit="m" value="1100.0">1100 m</elevation>
</quantity>
/
<geoCoordinate id="C8C47F9CA840FFE7D7CDFE638F65FDB1" box="[429,544,504,528]" degrees="25" direction="north" minutes="25. 6" orientation="latitude" pageId="19" pageNumber="20" precision="92" value="25.426666">25°25.6´N</geoCoordinate>
<geoCoordinate id="C8C47F9CA840FFE7D44DFE638FDBFDB1" box="[557,670,504,528]" degrees="90" direction="east" minutes="19. 5" orientation="longitude" pageId="19" pageNumber="20" precision="92" value="90.325">90°19.5´E</geoCoordinate>
<date id="D94E3F9BA840FFE7D4CBFE638E78FDB1" box="[683,829,504,528]" pageId="19" pageNumber="20" value="1996-05-09" valueMax="1996-05-17" valueMin="1996-05-09">
<collectingDate id="C90AC673A840FFE7D4CBFE638E78FDB1" box="[683,829,504,528]" pageId="19" pageNumber="20" value="1996-05-09" valueMax="1996-05-17" valueMin="1996-05-09">9.–17.5.1996</collectingDate>
</date>
leg.
<collectorName id="00057C8DA840FFE7D5E0FE638D85FD94" pageId="19" pageNumber="20">Jendek &amp; Sausa</collectorName>
(
<collectionCode id="CBE1819EA840FFE7D6B0FD868C6CFD94" box="[208,297,541,565]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="19" pageNumber="20" type="Museum">NHMW</collectionCode>
);
</materialsCitation>
<materialsCitation id="1D981306A840FFE7D75DFD868DFFFD04" ID-GBIF-Occurrence="3497007311" collectingDate="1988-04-18" collectingDateMax="1988-04-27" collectingDateMin="1988-04-18" collectionCode="BMNH, CBB" country="Thailand" elevation="300" latitude="15.466666" location="Umphang District Song Bae Stream" longLatPrecision="1285" longitude="98.8" pageId="19" pageNumber="20" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Tak Province">
<specimenCount id="BBF6D2D2A840FFE7D75DFD868C2CFD94" box="[317,361,538,565]" pageId="19" pageNumber="20" type="male">2 ♂</specimenCount>
,
<specimenCount id="BBF6D2D2A840FFE7D71AFD868CE3FD94" box="[378,422,538,565]" pageId="19" pageNumber="20" type="female">3 ♀</specimenCount>
, W.
<collectingCountry id="D5E759CBA840FFE7D7B1FD868F0BFD94" box="[465,590,541,565]" name="Thailand" pageId="19" pageNumber="20">THAILAND</collectingCountry>
:
<quantity id="6A08B4BEA840FFE7D43BFD868FE1FD94" box="[603,676,541,565]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="20" unit="m" value="300.0">
<elevation id="26DDFE68A840FFE7D43BFD868FE1FD94" box="[603,676,541,565]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="20" unit="m" value="300.0">300m</elevation>
</quantity>
., Thung Yai Wildlife Sanctuary.
<geoCoordinate id="C8C47F9CA840FFE7D6CDFDD98C4EFDFB" box="[173,267,578,602]" degrees="15" direction="north" minutes="28" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="15.466666">15°28´N</geoCoordinate>
–
<geoCoordinate id="C8C47F9CA840FFE7D74BFDD98CCEFDFB" box="[299,395,578,602]" degrees="98" direction="east" minutes="48" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="98.8">98°48´E</geoCoordinate>
. /
<collectingRegion id="6F34D7B9A840FFE7D7CDFDD98F7AFDFB" box="[429,575,578,602]" country="Thailand" name="Tak" pageId="19" pageNumber="20">Tak Province</collectingRegion>
<location id="A82F4F80A840FFE7D429FDD98EAEFDFB" LSID="urn:lsid:plazi:treatment:2559A84DA841FFE2D6FFFB998E71FC0B:A82F4F80A840FFE7D429FDD98EAEFDFB" box="[585,1003,578,602]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Umphang District Song Bae Stream" pageId="19" pageNumber="20" stateProvince="Tak Province">Umphang District Song Bae Stream</location>
.
<date id="D94E3F9BA840FFE7D619FDF38C53FD21" box="[121,278,616,640]" pageId="19" pageNumber="20" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">
<collectingDate id="C90AC673A840FFE7D619FDF38C53FD21" box="[121,278,616,640]" pageId="19" pageNumber="20" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">18–27.iv.1988</collectingDate>
</date>
. / Evergreen rain forest M.J.D.Brendell. B.M.1988–183. (
<collectionCode id="CBE1819EA840FFE7D5FCFDF38EAEFD21" box="[924,1003,616,640]" pageId="19" pageNumber="20">BMNH</collectionCode>
,
<collectionCode id="CBE1819EA840FFE7D619FD168DEAFD04" box="[121,175,653,677]" pageId="19" pageNumber="20">CBB</collectionCode>
);
</materialsCitation>
<materialsCitation id="1D981306A840FFE7D6A1FD168CCBFD6B" ID-GBIF-Occurrence="3497007316" collectingDate="1996-02-23" collectingDateMax="1996-02-25" collectingDateMin="1996-02-23" collectionCode="CBP" collectorName="K. Majer" country="Thailand" location="Hat Som Paen env" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Ranong prov.">
<specimenCount id="BBF6D2D2A840FFE7D6A1FD168DA8FD04" box="[193,237,650,677]" pageId="19" pageNumber="20" type="male">1 ♂</specimenCount>
,
<collectingCountry id="D5E759CBA840FFE7D69CFD168C26FD04" box="[252,355,653,677]" name="Thailand" pageId="19" pageNumber="20">Thailand</collectingCountry>
,
<date id="D94E3F9BA840FFE7D70FFD168F54FD04" box="[367,529,653,677]" pageId="19" pageNumber="20" value="1996-02-23" valueMax="1996-02-25" valueMin="1996-02-23">
<collectingDate id="C90AC673A840FFE7D70FFD168F54FD04" box="[367,529,653,677]" pageId="19" pageNumber="20" value="1996-02-23" valueMax="1996-02-25" valueMin="1996-02-23">23.–25.ii.1996</collectingDate>
</date>
<collectingRegion id="6F34D7B9A840FFE7D47AFD168FF6FD04" box="[538,691,653,677]" country="Thailand" name="Ranong" pageId="19" pageNumber="20">Ranong prov.</collectingRegion>
9°57´98°45´
<location id="A82F4F80A840FFE7D528FD168DE1FD6B" LSID="urn:lsid:plazi:treatment:2559A84DA841FFE2D6FFFB998E71FC0B:A82F4F80A840FFE7D528FD168DE1FD6B" country="Thailand" name="Hat Som Paen env" pageId="19" pageNumber="20" stateProvince="Ranong prov.">Hat Som Paen env</location>
.
<collectorName id="00057C8DA840FFE7D6CFFD298C4BFD6B" box="[175,270,690,714]" pageId="19" pageNumber="20">K. Majer</collectorName>
leg. (
<collectionCode id="CBE1819EA840FFE7D72EFD298CC1FD6B" box="[334,388,690,714]" pageId="19" pageNumber="20">CBP</collectionCode>
);
</materialsCitation>
<materialsCitation id="1D981306A840FFE7D7F5FD298ED7FD51" ID-GBIF-Occurrence="3497007312" collectingDate="1999-04-26" collectionCode="CBB" collectorName="Dembicky &amp; Pacholatko" country="India" latitude="25.45" location="Nokrek" longLatPrecision="1246" longitude="90.316666" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Meghalaya">
<specimenCount id="BBF6D2D2A840FFE7D7F5FD298C85FD6B" box="[405,448,687,714]" pageId="19" pageNumber="20" type="female">1 ♀</specimenCount>
, NE
<collectingCountry id="D5E759CBA840FFE7D795FD298F70FD6B" box="[501,565,690,714]" name="India" pageId="19" pageNumber="20">INDIA</collectingCountry>
<collectingRegion id="6F34D7B9A840FFE7D45DFD298FFCFD6B" box="[573,697,690,714]" country="India" name="Meghalaya" pageId="19" pageNumber="20">Meghalaya</collectingRegion>
<location id="A82F4F80A840FFE7D4A1FD298E57FD6B" LSID="urn:lsid:plazi:treatment:2559A84DA841FFE2D6FFFB998E71FC0B:A82F4F80A840FFE7D4A1FD298E57FD6B" box="[705,786,690,714]" country="India" latitude="25.45" longLatPrecision="1246" longitude="90.316666" name="Nokrek" pageId="19" pageNumber="20" stateProvince="Meghalaya">Nokrek</location>
n.p.,3kmSDaribokgirl
<geoCoordinate id="C8C47F9CA840FFE7D6C7FD438C4CFD51" box="[167,265,728,752]" degrees="25" direction="north" minutes="27" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="25.45">25°27´N</geoCoordinate>
,
<geoCoordinate id="C8C47F9CA840FFE7D775FD438C30FD51" box="[277,373,728,752]" degrees="90" direction="east" minutes="19" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="90.316666">90°19´E</geoCoordinate>
,
<date id="D94E3F9BA840FFE7D7E2FD438CB0FD51" box="[386,501,728,752]" pageId="19" pageNumber="20" value="1999-04-26">
<collectingDate id="C90AC673A840FFE7D7E2FD438CB0FD51" box="[386,501,728,752]" pageId="19" pageNumber="20" value="1999-04-26">26.4.1999</collectingDate>
</date>
<collectorName id="00057C8DA840FFE7D79EFD4F8F35FD51" box="[510,624,724,752]" pageId="19" pageNumber="20">Dembický</collectorName>
&amp;
<collectorName id="00057C8DA840FFE7D4F4FD438E57FD51" box="[660,786,728,752]" pageId="19" pageNumber="20">Pacholátko</collectorName>
lgt. (
<collectionCode id="CBE1819EA840FFE7D531FD438ECDFD51" box="[849,904,728,752]" pageId="19" pageNumber="20">CBB</collectionCode>
);
</materialsCitation>
<materialsCitation id="1D981306A840FFE7D5FAFD438F0AFC9B" ID-GBIF-Occurrence="3497007320" collectingDate="1998-05-01" collectingDateMax="1998-05-15" collectingDateMin="1998-05-01" collectionCode="CBP" collectorName="Michal Bednarik" country="Thailand" elevation="1400" latitude="19.316668" location="San Pakia" longLatPrecision="1272" longitude="98.833336" municipality="San Pakia" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Chiang Mai prov.">
<specimenCount id="BBF6D2D2A840FFE7D5FAFD438DF0FCB4" pageId="19" pageNumber="20" type="generic">1 specimen</specimenCount>
: N.
<collectingCountry id="D5E759CBA840FFE7D681FD668C1FFCB4" box="[225,346,765,789]" name="Thailand" pageId="19" pageNumber="20">THAILAND</collectingCountry>
,
<collectingRegion id="6F34D7B9A840FFE7D705FD668F62FCB4" box="[357,551,765,789]" country="Thailand" name="Chiang Mai" pageId="19" pageNumber="20">Chiang Mai prov.</collectingRegion>
<collectingMunicipality id="4D2B8321A840FFE7D44EFD668FE1FCB4" box="[558,676,765,789]" pageId="19" pageNumber="20">San Pakia</collectingMunicipality>
,
<geoCoordinate id="C8C47F9CA840FFE7D4CFFD668E54FCB4" box="[687,785,765,789]" degrees="19" direction="north" minutes="19" orientation="latitude" pageId="19" pageNumber="20" precision="925" value="19.316668">19°19’N</geoCoordinate>
,
<geoCoordinate id="C8C47F9CA840FFE7D57DFD668EC6FCB4" box="[797,899,765,789]" degrees="98" direction="east" minutes="50" orientation="longitude" pageId="19" pageNumber="20" precision="925" value="98.833336">98°50’ E</geoCoordinate>
,
<quantity id="6A08B4BEA840FFE7D5EFFD668EAFFCB4" box="[911,1002,765,789]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="19" pageNumber="20" unit="m" value="1400.0">
<elevation id="26DDFE68A840FFE7D5EFFD668EAFFCB4" box="[911,1002,765,789]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="19" pageNumber="20" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="D94E3F9BA840FFE7D619FCB98C4EFC9B" box="[121,267,802,826]" pageId="19" pageNumber="20" value="1998-05-01" valueMax="1998-05-15" valueMin="1998-05-01">
<collectingDate id="C90AC673A840FFE7D619FCB98C4EFC9B" box="[121,267,802,826]" pageId="19" pageNumber="20" value="1998-05-01" valueMax="1998-05-15" valueMin="1998-05-01">1.–15.5.1998</collectingDate>
</date>
,
<collectorName id="00057C8DA840FFE7D779FCB98C97FC9A" box="[281,466,801,827]" pageId="19" pageNumber="20">Michal Bednařik</collectorName>
leg. (
<collectionCode id="CBE1819EA840FFE7D46EFCB98F01FC9B" box="[526,580,802,826]" pageId="19" pageNumber="20">CBP</collectionCode>
);
</materialsCitation>
<materialsCitation id="1D981306A840FFE7D435FCB98C0AFC24" ID-GBIF-Occurrence="3497007322" collectingDate="2004-05-11" collectingDateMax="2004-05-12" collectingDateMin="2004-05-11" collectionCode="NHMB" collectorName="Dembicky" country="India" elevation="900" location="SW of CHERRAPUNJEE" pageId="19" pageNumber="20" specimenCount="1" stateProvince="MEGHALAYA">
<specimenCount id="BBF6D2D2A840FFE7D435FCB98F9FFC9B" box="[597,730,802,826]" pageId="19" pageNumber="20" type="generic">1 specimen</specimenCount>
: NE
<collectingCountry id="D5E759CBA840FFE7D569FCB98E0BFC9B" box="[777,846,802,826]" name="India" pageId="19" pageNumber="20">INDIA</collectingCountry>
,
<collectingRegion id="6F34D7B9A840FFE7D538FCB98EAEFC9B" box="[856,1003,802,826]" country="India" name="Meghalaya" pageId="19" pageNumber="20">MEGHALAYA</collectingRegion>
,
<location id="A82F4F80A840FFE7D619FCD38C3AFCC1" LSID="urn:lsid:plazi:treatment:2559A84DA841FFE2D6FFFB998E71FC0B:A82F4F80A840FFE7D619FCD38C3AFCC1" box="[121,383,840,864]" country="India" name="SW of CHERRAPUNJEE" pageId="19" pageNumber="20" stateProvince="MEGHALAYA">SW of CHERRAPUNJEE</location>
25°13´–15´N 91°40´E;
<quantity id="6A08B4BEA840FFE7D4F4FCD38F92FCC1" box="[660,727,840,864]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="19" pageNumber="20" unit="m" value="900.0">
<elevation id="26DDFE68A840FFE7D4F4FCD38F92FCC1" box="[660,727,840,864]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="19" pageNumber="20" unit="m" value="900.0">900m</elevation>
</quantity>
L.
<collectorName id="00057C8DA840FFE7D562FCD38E30FCC1" box="[770,885,836,864]" pageId="19" pageNumber="20">Dembický</collectorName>
leg.;
<date id="D94E3F9BA840FFE7D5DEFCD38DACFC24" pageId="19" pageNumber="20" value="2004-05-11" valueMax="2004-05-12" valueMin="2004-05-11">
<collectingDate id="C90AC673A840FFE7D5DEFCD38DACFC24" pageId="19" pageNumber="20" value="2004-05-11" valueMax="2004-05-12" valueMin="2004-05-11">11.– 12.v.2004</collectingDate>
</date>
(
<collectionCode id="CBE1819EA840FFE7D695FCF68C00FC24" box="[245,325,877,901]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="19" pageNumber="20">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A840FFE2D619FC468DAEFE01" lastPageId="22" lastPageNumber="23" pageId="19" pageNumber="20" type="description">
<paragraph id="AD4F195BA840FFE7D619FC468E47FC54" blockId="19.[121,770,989,1014]" box="[121,770,989,1014]" pageId="19" pageNumber="20">
<emphasis id="9F84C549A840FFE7D619FC468C5DFC57" bold="true" box="[121,280,989,1014]" pageId="19" pageNumber="20">Redescription</emphasis>
(
<figureCitation id="35CB05DEA840FFE7D744FC468C2DFC54" box="[292,360,989,1013]" captionStart="Figs 1–4" captionStartId="7.[121,158,1494,1512]" captionTargetBox="[145,1002,80,1436]" captionTargetPageId="7" captionText="Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284320" httpUri="https://zenodo.org/record/6284320/files/figure.png" pageId="19" pageNumber="20">Figs 2</figureCitation>
,
<figureCitation id="35CB05DEA840FFE7D716FC468CD7FC54" box="[374,402,989,1013]" captionStart="Figs 15–16" captionStartId="14.[159,196,1489,1507]" captionTargetBox="[159,1043,104,1429]" captionTargetId="figure-49@14.[159,1043,104,761]" captionTargetPageId="14" captionText="Figs 15–16: Orictites ssp., head and pronotum, lateral view; 15: Orictites (subgenus Orictites) minotaur ANDREWES, 1931,; 16: Orictites (subgenus Semictites) costulipennis BATES, 1892." figureDoi="http://doi.org/10.5281/zenodo.6284338" httpUri="https://zenodo.org/record/6284338/files/figure.png" pageId="19" pageNumber="20">16</figureCitation>
,
<figureCitation id="35CB05DEA840FFE7D7FFFC468CFEFC54" box="[415,443,989,1013]" captionStart="Figs 17–18" captionStartId="18.[159,196,780,798]" captionTargetBox="[159,1043,91,754]" captionText="Figs 17–18: Orictites spp., Maxilla, maxillary and labial palpomere. 17: O. minotaur ANDREWES, 1931 (subgenus Orictites); 18: O. costulipennis BATES, 1892 (subgenus Semictites subg. nov.)." figureDoi="http://doi.org/10.5281/zenodo.6284342" httpUri="https://zenodo.org/record/6284342/files/figure.png" pageId="19" pageNumber="20">18</figureCitation>
,
<figureCitation id="35CB05DEA840FFE7D7A8FC468CA0FC54" box="[456,485,989,1013]" captionStart="Figs 19–20" captionStartId="22.[159,196,1496,1514]" captionTargetBox="[159,1043,1049,1463]" captionText="Figs 19–20: Orictites spp., aedeagus, ventral view. 19: O. minotaur ANDREWES, 1931, lectotype, scale bar 1.5 mm; 20: O. costulipennis BATES, 1892, holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284346" httpUri="https://zenodo.org/record/6284346/files/figure.png" pageId="19" pageNumber="20">20</figureCitation>
,
<figureCitation id="35CB05DEA840FFE7D792FC468F56FC54" box="[498,531,989,1013]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="19" pageNumber="20">32</figureCitation>
; pp. 8, 15, 19, 23, 35)
</paragraph>
<paragraph id="AD4F195BA840FFE7D619FBB38E44FBC4" blockId="19.[121,1006,1064,1573]" pageId="19" pageNumber="20">
Measurements:
<typeStatus id="724BA7F9A840FFE7D752FBB38CDBFBE1" box="[306,414,1064,1088]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
: Body length
<quantity id="6A08B4BEA840FFE7D45DFBB38FD0FBE1" box="[573,661,1064,1088]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="19" pageNumber="20" unit="mm" value="5.1">5.1 mm</quantity>
, width
<quantity id="6A08B4BEA840FFE7D489FBB38E15FBE1" box="[745,848,1064,1088]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="19" pageNumber="20" unit="mm" value="1.45">1.45 mm</quantity>
; ratio length/ width of pronotum 0.98; ratio length/width of elytra 1.91.
</paragraph>
<paragraph id="AD4F195BA840FFE7D6FEFBE98FA3FB74" blockId="19.[121,1006,1064,1573]" pageId="19" pageNumber="20">
Additional material (n = 10): Body length
<quantity id="6A08B4BEA840FFE7D41AFBE98E47FB2B" box="[634,770,1138,1162]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="19" pageNumber="20" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5.5–6.5 mm</quantity>
(
<emphasis id="9F84C549A840FFE7D572FBEA8E67FB2B" bold="true" box="[786,802,1137,1162]" italics="true" pageId="19" pageNumber="20"></emphasis>
=
<quantity id="6A08B4BEA840FFE7D523FBE98ED8FB2B" box="[835,925,1138,1162]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="19" pageNumber="20" unit="mm" value="5.6">5.6 mm</quantity>
), width
<quantity id="6A08B4BEA840FFE7D619FB038C5FFB11" box="[121,282,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" metricValueMax="1.75" metricValueMin="1.49" pageId="19" pageNumber="20" unit="mm" value="1.62" valueMax="1.75" valueMin="1.49">1.49–1.75 mm</quantity>
(
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=
<quantity id="6A08B4BEA840FFE7D739FB038C85FB11" box="[345,448,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="19" pageNumber="20" unit="mm" value="1.62">1.62 mm</quantity>
), ratio length/width of pronotum 0.93 – 1.02 (
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= 0.96), ratio length/width of elytra 1.8 – 1.95 (
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= 1.87).
</paragraph>
<paragraph id="AD4F195BA840FFE7D6FEFB798CC5FAE4" blockId="19.[121,1006,1064,1573]" pageId="19" pageNumber="20">Colour: Shiny. Black to dark-brown. Wings of clypeus and supraantennal plates more or less transparent red-brown, antennae, labrum and palpi yellow brown, legs red-brown.</paragraph>
<paragraph id="AD4F195BA840FFE0D6FEFAC98E20FAE4" blockId="19.[121,1006,1064,1573]" lastBlockId="20.[159,1044,93,1573]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
Head: A fifth narrower than pronotum. Clypeus with small rounded tooth laterally, anterior margin slightly excised, with raised knob between the two clypeal setae, posterior part of knob covered with big and medium sized punctures; margin of wings convex, hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge. Clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, smooth, with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by flat transverse depression and group of big punctures, with costate, nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, with flat punctured transverse furrow, sharp furrow between frons and frontal carina. Frons moderately convex, with very minute scattered punctures. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes of moderate size, flattened but still convex, genae enlarged, regularly rounded, not as high as and shorter than eyes, enclosing eyes posteriorly, forming distinct angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as distinct step, with band of bigger punctures almost doubled, neck laterally covered with reticulation and densely scattered medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, nearly as long as head, flattened, stout, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A847FFE0D412FCDF8FCCFCF6" attach="left" box="[626,649,836,855]" fontSize="8" pageId="20" pageNumber="21">nd</superScript>
segment, 2
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segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with bigger sized punctures and transverse reticulation, Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with broad and rounded tubercle, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming a broad obtuse triangle, bisetose, not projecting as far as lateral lobes. Paragenae sharp, carina-like, with distinct tooth anteriorly, rounded at base. Antennae short, reaching up to middle of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 moniliform, antennomeres somewhat flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with few additional long setae.
</paragraph>
<paragraph id="AD4F195BA847FFE1D6A4FAC98C83FE64" blockId="20.[159,1044,93,1573]" lastBlockId="21.[121,1006,93,1573]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">Pronotum: Square, as long as wide, sides slightly convex at middle, distinctly rounded in anterior quarter to anterior angles. Anterior margin straight. Reflexed lateral margin smooth, from posterior angle to base running as a straight line. Lateral channel conspicuously broad between the two lateral setigerous punctures, with isodiametric reticulation and very few irregular punctures. Anterior setigerous puncture located at the end of anterior quarter approximated to the convexity of the disc of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle projecting, rounded, formed by reflexed lateral margin, posterior angle developed as sharp distinct tooth, projecting laterally. Anterior transverse line formed by distinct, partly connected punctures. Median line deep and conspicuously broad, running up to level of anterior transverse line but not joining, smaller at base, adjoining base. Surface scattered with some fine to medium sized punctures and with isodiametric reticulation laterally and at base, basal impression indistinct, basal channel broad, deep. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA846FFE1D6FEFE498F69FC24" blockId="21.[121,1006,93,1573]" pageId="21" pageNumber="22">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles. Reflexed margin with fine crenulation in anterior quarter, margin slightly thickened at humerus, bending with channel over rounded humerus up to 5
<superScript id="5A85B413A846FFE1D759FDFF8C09FDD6" attach="left" box="[313,332,612,631]" fontSize="8" pageId="21" pageNumber="22">th</superScript>
stria. No humeral tooth. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1
<superScript id="5A85B413A846FFE1D43EFD118F2AFD3C" attach="left" box="[606,623,650,669]" fontSize="8" pageId="21" pageNumber="22">st</superScript>
stria. Striae 1–4 free at base, all striae deep, broad, more or less distinctly punctuate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura costiform, raised. All intervals conspicuously convex, convexity increasing laterally, thought costate, 3
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and 4
<superScript id="5A85B413A846FFE1D524FC848E12FC93" attach="left" box="[836,855,799,818]" fontSize="8" pageId="21" pageNumber="22">th</superScript>
broader and raised at base, both with tubercle at base, 8
<superScript id="5A85B413A846FFE1D4E5FCDF8FDDFCF6" attach="left" box="[645,664,836,855]" fontSize="8" pageId="21" pageNumber="22">th</superScript>
carinate. Interval 3 with four setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA846FFE1D6FEFC098CA6FC0B" blockId="21.[121,1006,93,1573]" box="[158,483,914,938]" pageId="21" pageNumber="22">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA846FFE1D6FEFC238C4DFB74" blockId="21.[121,1006,93,1573]" pageId="21" pageNumber="22">
Ventral surface: Epipleuron shiny, covered with medium sized scattered punctures and indistinct isodiametric reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle like an inverted V, mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with big punctures at base laterally, 3
<superScript id="5A85B413A846FFE1D6DEFBF48D94FB23" attach="left" box="[190,209,1135,1154]" fontSize="8" pageId="21" pageNumber="22">rd</superScript>
to 5
<superScript id="5A85B413A846FFE1D766FBF48C5CFB23" attach="left" box="[262,281,1135,1154]" fontSize="8" pageId="21" pageNumber="22">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite six smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA846FFE1D6FEFB798CDDFA31" blockId="21.[121,1006,93,1573]" pageId="21" pageNumber="22">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, basal tarsomere elongated, nearly as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA846FFE1D6FEFA068C83F984" blockId="21.[121,1006,93,1573]" pageId="21" pageNumber="22">
Male genitalia (
<figureCitation id="35CB05DEA846FFE1D739FA068CEBFA14" box="[345,430,1437,1461]" captionStart="Figs 19–20" captionStartId="22.[159,196,1496,1514]" captionTargetBox="[159,1043,1049,1463]" captionText="Figs 19–20: Orictites spp., aedeagus, ventral view. 19: O. minotaur ANDREWES, 1931, lectotype, scale bar 1.5 mm; 20: O. costulipennis BATES, 1892, holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284346" httpUri="https://zenodo.org/record/6284346/files/figure.png" pageId="21" pageNumber="22">Fig. 20</figureCitation>
): Relatively long, moderately sclerotized. Median lobe slender, moderately arcuate, slightly twisted apically. Endophallus strongly folded, teeth not visible. Parameres of about same length, both slender, slightly twisted, asetose.
</paragraph>
<paragraph id="AD4F195BA845FFE2D6A4FFC68E68FF61" blockId="22.[159,1043,93,938]" pageId="22" pageNumber="23">
Female genitalia (
<figureCitation id="35CB05DEA845FFE2D7FAFFC68CA8FFD4" box="[410,493,93,117]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="22" pageNumber="23">Fig. 32</figureCitation>
): Coxostylus slightly sigmoid, dorso-ventrally moderately depressed, gently curved to apex, with one seta at apex, with one longer ensiform and four nematiform setae in middle part.
</paragraph>
<paragraph id="AD4F195BA845FFE2D6A4FF568DAEFE01" blockId="22.[159,1043,93,938]" pageId="22" pageNumber="23">
Variation: The anterior transverse line of the pronotum is more or less deeply engraved. The isodiametric reticulation in the lateral channel of the pronotum is more or less distinctly developed. Specimens from
<collectingRegion id="6F34D7B9A845FFE2D5F6FE838E9BFE91" box="[918,990,280,304]" country="Malaysia" name="Sabah" pageId="22" pageNumber="23">Sabah</collectingRegion>
and
<collectingCountry id="D5E759CBA845FFE2D6FFFEA68C47FEF4" box="[159,258,317,341]" name="Malaysia" pageId="22" pageNumber="23">Sarawak</collectingCountry>
(
<collectingCountry id="D5E759CBA845FFE2D772FEA68C39FEF4" box="[274,380,317,341]" name="Malaysia" pageId="22" pageNumber="23">Malaysia</collectingCountry>
) have the tubercles on the 3
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and 4
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interval at base indistinctly developed. The number of punctures on the disc of the pronotum varies.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A845FFE2D6A4FE368E31FD04" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="AD4F195BA845FFE2D6A4FE368E31FD04" blockId="22.[159,1043,93,938]" pageId="22" pageNumber="23">Diagnosis: A medium-sized almost black species with distinct knob on the clypeus and distinct flat transverse punctured depression behind, without humeral tooth, tubercles at the base of intervals 3 and 4 of the elytra, and four setigerous punctures on interval 3. Distinguished from all other species by the combination of the following characters: The frons of the head is covered by minute scattered punctures, the reflexed lateral margin of the pronotum is smooth, and all intervals of the elytra are conspicuously convex.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A845FFE2D6A4FD298C81FCB4" pageId="22" pageNumber="23" type="distribution">
<paragraph id="AD4F195BA845FFE2D6A4FD298C81FCB4" blockId="22.[159,1043,93,938]" pageId="22" pageNumber="23">
Distribution: Known from North, middle and Western parts of
<collectingCountry id="D5E759CBA845FFE2D5D0FD298956FD6B" box="[944,1043,690,714]" name="Thailand" pageId="22" pageNumber="23">Thailand</collectingCountry>
and from
<collectingRegion id="6F34D7B9A845FFE2D76CFD438C3FFD51" box="[268,378,728,752]" country="India" name="Meghalaya" pageId="22" pageNumber="23">Megalaya</collectingRegion>
(NE
<collectingCountry id="D5E759CBA845FFE2D7CFFD438CA9FD51" box="[431,492,728,752]" name="India" pageId="22" pageNumber="23">India</collectingCountry>
). Some specimens were collected at
<quantity id="6A08B4BEA845FFE2D5ECFD438E97FD51" box="[908,978,728,752]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="22" pageNumber="23" unit="m" value="450.0">450 m</quantity>
up to
<quantity id="6A08B4BEA845FFE2D6FFFD668DB1FCB4" box="[159,244,765,789]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="23" unit="m" value="1400.0">1400 m</quantity>
in litter of forests.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A845FFE2D6A4FCB98E71FC0B" pageId="22" pageNumber="23" type="discussion">
<paragraph id="AD4F195BA845FFE2D6A4FCB98E71FC0B" blockId="22.[159,1043,93,938]" pageId="22" pageNumber="23">
Remarks: Bates based the description on
<specimenCount id="BBF6D2D2A845FFE2D4FBFCB98E47FC9B" box="[667,770,802,826]" pageId="22" pageNumber="23" type="male">one male</specimenCount>
specimens deposited in MCSNG and he described the species as “curious” (
<bibRefCitation id="C96164AAA845FFE2D487FCD38EE4FCC1" author="Bates, H. W." box="[743,929,840,864]" pageId="22" pageNumber="23" pagination="267 - 428" refId="ref27786" refString="Bates, H. W. (1892): Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. - Annali del Museo Civico di Storia Naturale di Genova 32: 267 - 428." type="journal article" year="1892">Bates 1892: 282</bibRefCitation>
) obviously due to the striking differences to the
<taxonomicName id="6AF062D8A845FFE2D405FCF68E4BFC24" box="[613,782,877,901]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9F84C549A845FFE2D405FCF68FF7FC24" box="[613,690,877,901]" italics="true" pageId="22" pageNumber="23">Clivina</emphasis>
species
</taxonomicName>
described in the same contribution.
<bibRefCitation id="C96164AAA845FFE2D757FC098CF4FC0B" author="Kult, K." box="[311,433,914,938]" pageId="22" pageNumber="23" pagination="16 - 32" refId="ref27914" refString="Kult, K. (1951): Revision of the genus Clivina, Latr., from Oriental region. Revise rodu Clivina Latr. z orientalni oblasti. (Col. Carabidae). - Casopis Ceskoslovenske Spolecnosti Entomologicke 48: 16 - 32." type="journal article" year="1951">Kult (1951)</bibRefCitation>
did not mention the species at all.
</paragraph>
</subSubSection>
</treatment>
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319
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<paragraph id="AD4F195BA84BFFECD6FFFBD68F09FBC4" blockId="24.[159,588,1101,1126]" box="[159,588,1101,1126]" pageId="24" pageNumber="25">
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<taxonomicName id="6AF062D8A84BFFECD6FFFBD68F09FBC4" authority="LouWERENS, 1964" authorityName="LouWERENS" authorityYear="1964" box="[159,588,1101,1126]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A84BFFECD6FFFBD68C3CFBC7" bold="true" box="[159,377,1101,1126]" italics="true" pageId="24" pageNumber="25">Orictites mjoebergi</emphasis>
<bibRefCitation id="C96164AAA84BFFECD71FFBD68F09FBC4" author="Louwerens, C. J." box="[383,588,1101,1125]" pageId="24" pageNumber="25" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">LouWERENS, 1964</bibRefCitation>
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<paragraph id="AD4F195BA84BFFECD6FFFB038C57FB11" blockId="24.[159,755,1176,1312]" box="[159,274,1176,1200]" pageId="24" pageNumber="25">Catalogue</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84BFFECD6FFFB268FB6FA81" pageId="24" pageNumber="25" type="reference_group">
<paragraph id="AD4F195BA84BFFECD6FFFB268FCCFB74" blockId="24.[159,755,1176,1312]" box="[159,649,1213,1237]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="8DE03E75A84BFFECD6FFFB268FCCFB74" box="[159,649,1213,1237]" pageId="24" pageNumber="25">
<emphasis id="9F84C549A84BFFECD6FFFB268CC5FB74" box="[159,384,1213,1237]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="6AF062D8A84BFFECD6FFFB268C3CFB74" authorityName="LouWERENS" authorityYear="1964" box="[159,377,1213,1237]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mjoebergi">Orictites mjoebergi</taxonomicName>
;
</emphasis>
<treatmentCitation id="2C513F4AA84BFFECD7E6FB268FC4FB74" author="Louwerens, C. J." box="[390,641,1213,1237]" page="172" pageId="24" pageNumber="25" year="1964">
<bibRefCitation id="C96164AAA84BFFECD7E6FB268FC4FB74" author="Louwerens, C. J." box="[390,641,1213,1237]" pageId="24" pageNumber="25" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens, 1964: 172</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA84BFFECD6FFFB798F99FB5B" blockId="24.[159,755,1176,1312]" box="[159,732,1250,1274]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="8DE03E75A84BFFECD6FFFB798F99FB5B" box="[159,732,1250,1274]" pageId="24" pageNumber="25">
<taxonomicName id="6AF062D8A84BFFECD6FFFB798F46FB5B" authority="LouWERENS" authorityName="LouWERENS" authorityYear="1964" box="[159,515,1250,1274]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A84BFFECD6FFFB798C3CFB5B" box="[159,377,1250,1274]" italics="true" pageId="24" pageNumber="25">Orictites mjoebergi</emphasis>
LouWERENS
</taxonomicName>
;
<treatmentCitation id="2C513F4AA84BFFECD471FB798F91FB5B" author="Lorenz, W." box="[529,724,1250,1274]" page="135" pageId="24" pageNumber="25" year="1998">
<bibRefCitation id="C96164AAA84BFFECD471FB798F91FB5B" author="Lorenz, W." box="[529,724,1250,1274]" pageId="24" pageNumber="25" refId="ref28004" refString="Lorenz, W. (1998): Systematic list of extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). - Tutzing, printed by the author. 502 pp." type="book" year="1998">Lorenz 1998: 135</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA84BFFECD6FFFA938FB6FA81" blockId="24.[159,755,1176,1312]" box="[159,755,1288,1312]" pageId="24" pageNumber="25">
<treatmentCitationGroup id="8DE03E75A84BFFECD6FFFA938FB6FA81" box="[159,755,1288,1312]" pageId="24" pageNumber="25">
<taxonomicName id="6AF062D8A84BFFECD6FFFA938F46FA81" authority="LouWERENS" authorityName="LouWERENS" authorityYear="1964" box="[159,515,1288,1312]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A84BFFECD6FFFA938C3CFA81" box="[159,377,1288,1312]" italics="true" pageId="24" pageNumber="25">Orictites mjoebergi</emphasis>
LouWERENS
</taxonomicName>
;
<treatmentCitation id="2C513F4AA84BFFECD471FA938FAEFA81" author="Balkenohl, M." box="[529,747,1288,1312]" page="18" pageId="24" pageNumber="25" year="2001">
<bibRefCitation id="C96164AAA84BFFECD471FA938FAEFA81" author="Balkenohl, M." box="[529,747,1288,1312]" pageId="24" pageNumber="25" refId="ref27660" refString="Balkenohl, M. (2001): Key and Catalogue of the tribe Clivinini from the Oriental realm, with revisions of the genera Thliboclivina KULT, and Trilophidius JEANNEL (Insecta, Coleoptera, Carabidae, Scarititae, Clivinini). - 83 pp., Sofia, Moscow (Pensoft Series Faunistica 21)." type="book" year="2001">Balkenohl 2001: 18</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84BFFECD6FFFAC98C09F984" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="AD4F195BA84BFFECD6FFFAC98C09F984" blockId="24.[159,1043,1362,1573]" pageId="24" pageNumber="25">
Material examined:
<materialsCitation id="1D981306A84BFFECD7EEFAC98C09F984" ID-GBIF-Occurrence="3497007340" collectionCode="NHRS" collectorName="O. Borneo MjOberg" determinerName="C. J. Louwerens" location="Pajau River" municipality="Pajau River" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
Type material:
<typeStatus id="724BA7F9A84BFFECD423FAC98FF0FACB" box="[579,693,1362,1386]" pageId="24" pageNumber="25" type="lectotype">Lectotype</typeStatus>
(by present designation).
<specimenCount id="BBF6D2D2A84BFFECD599FAC88948FACB" box="[1017,1037,1363,1386]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
, with labels and data: white, handwritten: “Type
<taxonomicName id="6AF062D8A84BFFECD4AAFAE38E6EFA31" authorityName="ANDREWES" authorityYear="1931" box="[714,811,1400,1424]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Orictites</taxonomicName>
mjÖbergi sp.n.” and printed “det.
<determinerName id="5893F59FA84BFFECD749FA068C91FA14" box="[297,468,1437,1461]" pageId="24" pageNumber="25">C.J.Louwerens</determinerName>
” / blue: “3491 E91 +” / 2
<superScript id="5A85B413A84BFFECD499FA018E55FA0C" attach="left" box="[761,784,1434,1453]" fontSize="8" pageId="24" pageNumber="25">nd</superScript>
pin with original small paper card: small, white, “
<location id="A82F4F80A84BFFECD7ACFA598F10FA7B" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84BFFECD7ACFA598F10FA7B" box="[460,597,1474,1498]" municipality="Pajau River" name="Pajau River" pageId="24" pageNumber="25">Pajau River</location>
” / white, “O. Borneo MjÖberg” / small blue: “3492 E91 +” / big, white, “
<taxonomicName id="6AF062D8A84BFFECD452FA738FB1F9A1" authority="Andr." authorityName="ANDREWES" authorityYear="1931" box="[562,756,1512,1536]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">ORICTITES Andr.</taxonomicName>
” / big, white: “MjÖbergi Louw.” (
<collectionCode id="CBE1819EA84BFFECD698F9968C07F984" box="[248,322,1549,1573]" collectionName="Sweden, Stockholm, Naturhistoriska riksmuseet" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="24" pageNumber="25">NHRS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84AFFEDD6FEFFC68F00FD04" pageId="25" pageNumber="26" type="discussion">
<paragraph id="AD4F195BA84AFFEDD6FEFFC68F00FD04" blockId="25.[121,1006,93,938]" pageId="25" pageNumber="26">
Remarks: C.J.
<bibRefCitation id="C96164AAA84AFFEDD720FFC68F4AFFD4" author="Louwerens, C. J." box="[320,527,93,117]" pageId="25" pageNumber="26" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens (1964)</bibRefCitation>
based the description on
<specimenCount id="BBF6D2D2A84AFFEDD521FFC68EA8FFD4" box="[833,1005,93,117]" pageId="25" pageNumber="26" type="generic">two specimens</specimenCount>
from Pajan River. One of them is located in the NHRS (see material examined), the other one could not be located and is obviously lost. The available specimen isdesignated here as
<typeStatus id="724BA7F9A84AFFEDD7C7FF568F57FF44" box="[423,530,205,229]" pageId="25" pageNumber="26" type="lectotype">lectotype</typeStatus>
in the interst of the stability. In the specimen the right wing of the clypeus is slightly more angled (dorsal view) and more fused with the median part. I recognise this character as intraindividual variation. Exactly this asymmetric character is described by
<bibRefCitation id="C96164AAA84AFFEDD511FEA68D80FEDB" author="Louwerens, C. J." pageId="25" pageNumber="26" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens (1964)</bibRefCitation>
. However, careful investigation revealed that this specimen is a male and not a female as indicated by Louwerens. The inner structures were slightly damaged, obviously by a former mounting attempt. The left antennae misses segments 8–11. It should be mentioned that the sketch provided by Louwerens does not display a realistic picture of the species. In addition,
<bibRefCitation id="C96164AAA84AFFEDD512FE638C45FD94" author="Louwerens, C. J." pageId="25" pageNumber="26" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens (1964: 173)</bibRefCitation>
describes the presence of five setigerous punctures on interval 3 of the elytra. However, the type specimen possesses four, respectively five setigerous punctures (right/left side). The additional specimens all exhibit four setigerous punctures on either side.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84AFFEDD6FEFD298C3EFC0B" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="AD4F195BA84AFFEDD6FEFD298C3EFC0B" blockId="25.[121,1006,93,938]" pageId="25" pageNumber="26">
Additional material:
<materialsCitation id="1D981306A84AFFEDD7FAFD298EF4FD51" ID-GBIF-Occurrence="3497007317" collectingDate="1991-08-05" collectionCode="SMNS" country="NORD-SUMATRA" elevation="1500" location="Kiefernwald" municipality="Umg. Siulakosa" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1">
<specimenCount id="BBF6D2D2A84AFFEDD7FAFD298C8FFD6B" box="[410,458,690,714]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
,
<collectingCountry id="D5E759CBA84AFFEDD7BFFD298FE3FD6B" box="[479,678,690,714]" pageId="25" pageNumber="26">NORD-SUMATRA</collectingCountry>
:
<collectingMunicipality id="4D2B8321A84AFFEDD4D7FD298E37FD6B" box="[695,882,690,714]" name="Umg. Siulakosa" pageId="25" pageNumber="26">Umg. Siulakosa</collectingMunicipality>
<date id="D94E3F9BA84AFFEDD5E0FD298EA3FD6B" box="[896,998,690,714]" pageId="25" pageNumber="26" value="1991-08-05">
<collectingDate id="C90AC673A84AFFEDD5E0FD298EA3FD6B" box="[896,998,690,714]" pageId="25" pageNumber="26" value="1991-08-05">5.8.1991</collectingDate>
</date>
, ERBER / Samosir, ob. Tomok
<quantity id="6A08B4BEA84AFFEDD7ABFD438F63FD51" box="[459,550,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="m" value="1500.0">
<elevation id="26DDFE68A84AFFEDD7ABFD438F63FD51" box="[459,550,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="25" pageNumber="26" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<location id="A82F4F80A84AFFEDD452FD438FFBFD51" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD452FD438FFBFD51" box="[562,702,728,752]" country="NORD-SUMATRA" municipality="Umg. Siulakosa" name="Kiefernwald" pageId="25" pageNumber="26">Kiefernwald</location>
,
<location id="A82F4F80A84AFFEDD4A9FD438E0BFD51" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD4A9FD438E0BFD51" box="[713,846,728,752]" country="NORD-SUMATRA" municipality="Umg. Siulakosa" name="unter Rinde" pageId="25" pageNumber="26">unter Rinde</location>
(
<collectionCode id="CBE1819EA84AFFEDD53DFD438EEEFD51" box="[861,939,728,752]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="25" pageNumber="26">SMNS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A84AFFEDD5DDFD438E37FCB4" ID-GBIF-Occurrence="3497007332" collectingDate="1995-03" collectionCode="CBP" collectorName="M. Malindang North Sambuanga" country="Philippines" location="M. Malindang North Sambuanga" municipality="Mindanao" pageId="25" pageNumber="26" specimenCount="1" specimenCount-male="1">
<specimenCount id="BBF6D2D2A84AFFEDD5DDFD438EADFD4E" box="[957,1000,728,752]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
,
<collectingCountry id="D5E759CBA84AFFEDD619FD668D9FFCB4" box="[121,218,765,789]" name="Philippines" pageId="25" pageNumber="26">Filippine</collectingCountry>
<collectingMunicipality id="4D2B8321A84AFFEDD681FD668C17FCB4" box="[225,338,765,789]" country="Philippines" name="Mindanao" pageId="25" pageNumber="26">Mindanao</collectingMunicipality>
<date id="D94E3F9BA84AFFEDD739FD668CEBFCB4" box="[345,430,765,789]" pageId="25" pageNumber="26" value="1995-03">
<collectingDate id="C90AC673A84AFFEDD739FD668CEBFCB4" box="[345,430,765,789]" pageId="25" pageNumber="26" value="1995-03">III.1995</collectingDate>
</date>
<location id="A82F4F80A84AFFEDD7D6FD668E6DFCB4" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD7D6FD668E6DFCB4" box="[438,808,765,789]" country="Philippines" municipality="Mindanao" name="M. Malindang North Sambuanga" pageId="25" pageNumber="26">M. Malindang North Sambuanga</location>
(
<collectionCode id="CBE1819EA84AFFEDD556FD668E29FCB4" box="[822,876,765,789]" pageId="25" pageNumber="26">CBP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A84AFFEDD51DFD668ECCFC9B" ID-GBIF-Occurrence="3497007341" collectingDate="2014-11" collectionCode="CBB" collectorName="N Luzon" country="PHIL- IPPINES" location="N Luzon" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" stateProvince="Ifugao">
<specimenCount id="BBF6D2D2A84AFFEDD51DFD668EE3FCB4" box="[893,934,765,789]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
,
<collectingCountry id="D5E759CBA84AFFEDD5D4FD668D9DFC9B" pageId="25" pageNumber="26">PHILIPPINES</collectingCountry>
,
<location id="A82F4F80A84AFFEDD682FCB98C06FC9B" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD682FCB98C06FC9B" box="[226,323,802,826]" country="PHIL- IPPINES" name="N Luzon" pageId="25" pageNumber="26" stateProvince="Ifugao">N Luzon</location>
,
<location id="A82F4F80A84AFFEDD72EFCB98CDEFC9B" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD72EFCB98CDEFC9B" box="[334,411,802,826]" country="PHIL- IPPINES" name="Ifugao" pageId="25" pageNumber="26" stateProvince="Ifugao">Ifugao</location>
,
<location id="A82F4F80A84AFFEDD7C5FCB98CBAFC9B" LSID="urn:lsid:plazi:treatment:2559A84DA84BFFE8D6FFFBD68E7EFC24:A82F4F80A84AFFEDD7C5FCB98CBAFC9B" box="[421,511,802,826]" country="PHIL- IPPINES" name="Banaue" pageId="25" pageNumber="26" stateProvince="Ifugao">Banaue</location>
,
<date id="D94E3F9BA84AFFEDD469FCB98F25FC9B" box="[521,608,802,826]" pageId="25" pageNumber="26" value="2014-11">
<collectingDate id="C90AC673A84AFFEDD469FCB98F25FC9B" box="[521,608,802,826]" pageId="25" pageNumber="26" value="2014-11">XI 2014</collectingDate>
</date>
. local collector leg. (
<collectionCode id="CBE1819EA84AFFEDD52CFCB98EC6FC9B" box="[844,899,802,826]" collectionName="CBB" pageId="25" pageNumber="26">CBB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A84AFFEDD5F4FCB98C32FC0B" ID-GBIF-Occurrence="3497007306" collectionCode="BMNH" collectorName="Banninger &amp; H. E. Andrewes" country="Philippines" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Ifugao">
<specimenCount id="BBF6D2D2A84AFFEDD5F4FCB98DEBFCC1" pageId="25" pageNumber="26" type="generic">1 specimen</specimenCount>
: head and pronotum lost, F. C. DRESCHER G. Tangkoeban Prahoe 4000 – 5000 Voet Praeanger, Java
<date id="D94E3F9BA84AFFEDD7D0FCF68F4AFC24" box="[432,527,877,901]" pageId="25" pageNumber="26" value="1933-06">VI. 1933</date>
/ Ex coll.
<collectorName id="00057C8DA84AFFEDD4E5FCF68FBDFC24" box="[645,760,877,901]" pageId="25" pageNumber="26">Bänninger</collectorName>
/
<collectorName id="00057C8DA84AFFEDD574FCF68EF6FC24" box="[788,947,877,901]" pageId="25" pageNumber="26">
H.E.
<taxonomicName id="6AF062D8A84AFFEDD559FCF68EF6FC24" box="[825,947,877,901]" pageId="25" pageNumber="1" rank="subGenus" status="subgen. nov." subGenus="Andrewes">Andrewes</taxonomicName>
</collectorName>
Coll. B.M.1945–97. (
<collectionCode id="CBE1819EA84AFFEDD741FC098C34FC0B" box="[289,369,914,938]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="25" pageNumber="26">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84AFFE8D619FB998F6FFDFB" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26" type="description">
<paragraph id="AD4F195BA84AFFEDD619FB998F19FBBB" blockId="25.[121,604,1026,1051]" box="[121,604,1026,1051]" pageId="25" pageNumber="26">
<emphasis id="9F84C549A84AFFEDD619FB998C5DFBBA" bold="true" box="[121,280,1026,1051]" pageId="25" pageNumber="26">Redescription</emphasis>
(
<figureCitation id="35CB05DEA84AFFEDD744FB998C2DFBBB" box="[292,360,1026,1050]" captionStart="Figs 1–4" captionStartId="7.[121,158,1494,1512]" captionTargetBox="[145,1002,80,1436]" captionTargetPageId="7" captionText="Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284320" httpUri="https://zenodo.org/record/6284320/files/figure.png" pageId="25" pageNumber="26">Figs 3</figureCitation>
,
<figureCitation id="35CB05DEA84AFFEDD715FB998CD4FBBB" box="[373,401,1026,1050]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="25" pageNumber="26">21</figureCitation>
,
<figureCitation id="35CB05DEA84AFFEDD7FEFB998CFAFBBB" box="[414,447,1026,1050]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="25" pageNumber="26">33</figureCitation>
; pp. 8, 24, 35)
</paragraph>
<paragraph id="AD4F195BA84AFFEDD619FBD68FBBFB2B" blockId="25.[121,1006,1101,1573]" pageId="25" pageNumber="26">
Measurements:
<typeStatus id="724BA7F9A84AFFEDD751FBD68CE3FBC4" box="[305,422,1101,1125]" pageId="25" pageNumber="26" type="lectotype">Lectotype</typeStatus>
: Body length
<quantity id="6A08B4BEA84AFFEDD422FBD68FDEFBC4" box="[578,667,1101,1125]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="25" pageNumber="26" unit="mm" value="5.4">5.4 mm</quantity>
, width
<quantity id="6A08B4BEA84AFFEDD48CFBD68E17FBC4" box="[748,850,1101,1125]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="25" pageNumber="26" unit="mm" value="1.52">1.52 mm</quantity>
; ratio length/ width of pronotum 1.07; ratio length/width of elytra 1.95.
</paragraph>
<paragraph id="AD4F195BA84AFFEDD6FEFB038F9CFB5B" blockId="25.[121,1006,1101,1573]" pageId="25" pageNumber="26">
Additional material (n = 3): Body length
<quantity id="6A08B4BEA84AFFEDD40BFB038E47FB11" box="[619,770,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="5.415" metricValueMax="5.9" metricValueMin="4.93" pageId="25" pageNumber="26" unit="mm" value="5.415" valueMax="5.9" valueMin="4.93">4.93–5.9 mm</quantity>
(
<emphasis id="9F84C549A84AFFEDD572FB0C8E67FB11" bold="true" box="[786,802,1175,1200]" italics="true" pageId="25" pageNumber="26"></emphasis>
=
<quantity id="6A08B4BEA84AFFEDD523FB038ED8FB11" box="[835,925,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="25" pageNumber="26" unit="mm" value="5.4">5.4 mm</quantity>
), width
<quantity id="6A08B4BEA84AFFEDD619FB268C5CFB74" box="[121,281,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.51" metricValueMax="1.65" metricValueMin="1.37" pageId="25" pageNumber="26" unit="mm" value="1.51" valueMax="1.65" valueMin="1.37">1.37–1.65 mm</quantity>
(
<emphasis id="9F84C549A84AFFEDD74AFB278C7FFB74" bold="true" box="[298,314,1212,1237]" italics="true" pageId="25" pageNumber="26"></emphasis>
=
<quantity id="6A08B4BEA84AFFEDD73BFB268C87FB74" box="[347,450,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="25" pageNumber="26" unit="mm" value="1.53">1.53 mm</quantity>
); ratio length/width of pronotum 0.96–1.06 (
<emphasis id="9F84C549A84AFFEDD5A4FB278E91FB74" bold="true" box="[964,980,1212,1237]" italics="true" pageId="25" pageNumber="26"></emphasis>
= 1.0); ratio length/width of elytra 1.78–1.97 (
<emphasis id="9F84C549A84AFFEDD40AFB7A8F3FFB5B" bold="true" box="[618,634,1249,1274]" italics="true" pageId="25" pageNumber="26"></emphasis>
= 1.89).
</paragraph>
<paragraph id="AD4F195BA84AFFEDD6FEFA938E69FACB" blockId="25.[121,1006,1101,1573]" pageId="25" pageNumber="26">Colour: Shiny. Black. Elytra red-brown, pronotum darker. Wings of clypeus and supraantennal plates transparent red-brown. Antennae, mouthparts, middle and hind legs yellowish brown, front legs dark red-brown.</paragraph>
<paragraph id="AD4F195BA84AFFEED6FEFAE38C59FA81" blockId="25.[121,1006,1101,1573]" lastBlockId="26.[159,1044,93,1573]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
Head: A quarter narrower than pronotum. Clypeus limited laterally by small obtuse angled tooth, anterior margin straight, with blunt raised knob between the two clypeal setae, posterior part of knob covered with medium sized punctures, anterior part smooth; wings obtuse-angular, rounded at tip, hollowed out, divided from middle part of clypeus by slight incisions and carinae, divided from supraantennal plates by distinct obtusely angled incisions and indistinct carina, clypeus and wings reflexed margined, supraantennal plates vaulted, elongated as supraorbital carina, smooth, with reflexed rounded margin, frons separated from clypeus by flat transverse depression and a group of punctures formed like a letter V, with nearly parallel running costate frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina. Frons moderately convex, nearly smooth, with small but distinct longitudinal impression at middle. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from a tubercle. Eyes of moderate size, moderately convex, genae enlarged, regularly rounded, not as high as and a third shorter than eyes, enclosing eyes posteriorly. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as distinct step, with band of big punctures, neck laterally densely covered with medium sized punctures. Labrum indistinctly trilobed, appearing straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles half as long as head, flattened, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled and somewhat hollowed out dorsally. Maxilla nearly straight, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, longer than 2
<superScript id="5A85B413A849FFEED4CBFCDF8F87FCF6" attach="left" box="[683,706,836,855]" fontSize="8" pageId="26" pageNumber="27">nd</superScript>
segment, 2
<superScript id="5A85B413A849FFEED52EFCDF8E20FCF6" attach="left" box="[846,869,836,855]" fontSize="8" pageId="26" pageNumber="27">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with medium sized punctures and irregular reticulation. Gula smooth. Submentum and mentum distinctly separated; submentum with two pairs of setae; mentum keeled at middle, nearly smooth, with fine and complete reflexed margin, lateral lobes projecting, thought obtusely angled at tips, with two setae near base of each lobe, median tooth forming broad obtuse triangle, bisetose, less projecting than lateral lobes. Paragenae sharp, carina-like, without tooth. Antennae of medium length, reaching over middle of pronotum, scapus carinate, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 transverse (L/
<geoCoordinate id="C8C47F9CA849FFEED4F7FB268FB4FB74" box="[663,753,1213,1237]" degrees="0. 80" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="555" value="-0.8">W 0.80</geoCoordinate>
), antennomeres subglobose, nearly dull, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA849FFEFD6A4FAB68DF9FDFB" blockId="26.[159,1044,93,1573]" lastBlockId="27.[121,1006,93,1573]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Pronotum: Square (L/
<geoCoordinate id="C8C47F9CA849FFEED7A6FAB68F65FAE4" box="[454,544,1325,1349]" degrees="1. 07" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="555" value="-1.07">W 1.07</geoCoordinate>
), sides straight but slightly diverging to base, gently rounded before posterior angle. Anterior margin slightly bisinuate. Reflexed lateral margin nearly smooth, with indistinct reticulation (100x), from posterior angle to base running as smooth, slightly convex line. Lateral channel distinctly broadened between the two lateral setigerous punctures, with fine irregular reticulation, with a few medium-sized punctures situated in broadened part of channel, basal channel broad, deep, foveolate. Anterior setigerous puncture located at end of anterior quarter, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle slightly but distinctly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as large rounded tooth, projecting laterally. Anterior transverse line sharp, punctured, not adjoining lateral channel. Median line deep, broad, thought with indistinct blunt carina-like vault bilaterally, surpassing level of anterior transverse line but not joining, smaller at base, joining base. Surface with two longitudinal groups of paramedian and paralateral big punctures and with a few fine, irregularly situated punctures, with irregular reticulation laterally, shiny on disc; basal impression developed as small flat furrow; base of disc with indistinct reticulation. Basal part of disc elongated posteriorly and bilaterally at declivity, as tooth-like vault slightly hanging over basal channel (lateral view). Flange raised keel-like (lateral view).
</paragraph>
<paragraph id="AD4F195BA848FFEFD6FEFDF38EA0FBC4" blockId="27.[121,1006,93,1573]" pageId="27" pageNumber="28">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, not completely visible from above due to convexity of intervals 7 and 8, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of punctures laterally. Reflexed margin with fine crenulation in anterior quarter, channel broadened at humerus, bending with margin over rounded toothless humerus up to 5
<superScript id="5A85B413A848FFEFD469FC848F59FC93" attach="left" box="[521,540,799,818]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
stria. Basal tubercle big, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, broad, distinctly punctate-striate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura costiform, raised, basal area around striole moderately impressed. Intervals shiny, distinctly convex, 3
<superScript id="5A85B413A848FFEFD4E8FC418FDEFC4C" attach="left" box="[648,667,986,1005]" fontSize="8" pageId="27" pageNumber="28">rd</superScript>
broader, strikingly raised and with large tubercle at base, 4
<superScript id="5A85B413A848FFEFD7A5FC648C9DFBB3" attach="left" box="[453,472,1023,1042]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
broader and raised at base but not as strongly as 3
<superScript id="5A85B413A848FFEFD6CBFBBF8DFBFB96" attach="none" box="[171,190,1060,1079]" fontSize="8" pageId="27" pageNumber="28">rd</superScript>
, 7
<superScript id="5A85B413A848FFEFD6BDFBBF8DB5FB96" attach="left" box="[221,240,1060,1079]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
running to apex as obtuse carina. 8
<superScript id="5A85B413A848FFEFD4FBFBBF8FEBFB96" attach="left" box="[667,686,1060,1079]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
small, completely carinate. Interval 3 with four/five setigerous punctures adjoining stria 3 (see variation).
</paragraph>
<paragraph id="AD4F195BA848FFEFD6FEFBE98CA6FB2B" blockId="27.[121,1006,93,1573]" box="[158,483,1138,1162]" pageId="27" pageNumber="28">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA848FFEFD6FEFB038CFEFA7B" blockId="27.[121,1006,93,1573]" pageId="27" pageNumber="28">
Ventral surface: Proepisternum with complete submarginal ridge, with some punctures, with irregular reticulation laterally. Proepipleuron small, with punctures and indistinct reticulation. Epipleuron broadened and with big punctures in basal quarter. Proepisternum somewhat swollen at level of posterior setigerous puncture, prosternite smooth, double keeled at middle, mesosternum smooth. Sternites smooth at middle, each with big punctures at base laterally, 3
<superScript id="5A85B413A848FFEFD736FAEF8C2CFA26" attach="left" box="[342,361,1396,1415]" fontSize="8" pageId="27" pageNumber="28">rd</superScript>
to 5
<superScript id="5A85B413A848FFEFD7C2FAEF8CF0FA26" attach="left" box="[418,437,1396,1415]" fontSize="8" pageId="27" pageNumber="28">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA848FFE8D6FEFA738CFBFF61" blockId="27.[121,1006,93,1573]" lastBlockId="28.[159,1044,93,901]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral and lateral, no dorsal furrow, basal tarsomere elongated, longer than the following four together. Mesotibia with apical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA84FFFE8D6A4FF568F90FE91" blockId="28.[159,1044,93,901]" pageId="28" pageNumber="29">
Male genitalia (
<figureCitation id="35CB05DEA84FFFE8D71EFF568C94FF44" box="[382,465,205,229]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="28" pageNumber="29">Fig. 21</figureCitation>
): Relatively short, median lobe broad, regularly arcuate at middle, apex twisted apically. Endophallus with bristles laterally. Both parameres slender, slightly twisted, asetose.
</paragraph>
<paragraph id="AD4F195BA84FFFE8D6A4FEA68EC8FE64" blockId="28.[159,1044,93,901]" pageId="28" pageNumber="29">
Female genitalia (
<figureCitation id="35CB05DEA84FFFE8D7EDFEA68C9FFEF4" box="[397,474,317,341]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="28" pageNumber="29">Fig. 33</figureCitation>
): Coxostylus of moderate width, distinctly curved to apex, with two preapical setae close together, dorsally with some minute scars in apical half, with obtuse knob-like tooth dorsally at base, with two short ensiform setae at middle, with four longer nematiform setae.
</paragraph>
<paragraph id="AD4F195BA84FFFE8D6A4FE498F6FFDFB" blockId="28.[159,1044,93,901]" pageId="28" pageNumber="29">
Variation: Bilateral variation is observed in the right clypeal wing, mentioned under remarks. In addition, in the
<typeStatus id="724BA7F9A84FFFE8D4E3FE638FABFDB1" box="[643,750,504,528]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
the left elytra bears five setigerous punctures and the right one four. The other specimens all have four setigerous punctures on each side.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84FFFE8D6A4FDF38E7EFC24" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="AD4F195BA84FFFE8D6A4FDF38E7EFC24" blockId="28.[159,1044,93,901]" pageId="28" pageNumber="29">
Diagnosis: A medium-sized almost black species with red-brownish elytra, with distinct knob on the clypeus, humerus without tooth, and four setigerous punctures on interval 3 (but compare remarks). Only in this species and in
<taxonomicName id="6AF062D8A84FFFE8D6FFFD438C17FD51" authorityName="Balkenohl" authorityYear="2017" box="[159,338,728,752]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="anteriorlatus">
<emphasis id="9F84C549A84FFFE8D6FFFD438C17FD51" box="[159,338,728,752]" italics="true" pageId="28" pageNumber="29">O. anteriorlatus</emphasis>
</taxonomicName>
is the basal part of the disc of the pronotum posteriorly elongated bilaterally at the declivity as a tooth-like vault slightly hanging over the basal channel. By contrast to
<taxonomicName id="6AF062D8A84FFFE8D797FCB98FEEFC9B" authorityName="Balkenohl" authorityYear="2017" box="[503,683,802,826]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="anteriorlatus">
<emphasis id="9F84C549A84FFFE8D797FCB98FEEFC9B" box="[503,683,802,826]" italics="true" pageId="28" pageNumber="29">O. anteriorlatus</emphasis>
</taxonomicName>
,
<taxonomicName id="6AF062D8A84FFFE8D4DAFCB98E0AFC9B" authorityName="LouWERENS" authorityYear="1964" box="[698,847,802,826]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A84FFFE8D4DAFCB98E0AFC9B" box="[698,847,802,826]" italics="true" pageId="28" pageNumber="29">O. mjoebergi</emphasis>
</taxonomicName>
exhibits a tubercle at the base of interval 3 of the elytra, only, and the furrow between clypeus and frons consists of punctures in form of the letter V.
</paragraph>
</subSubSection>
</treatment>
</document>

150
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<taxonomicName id="6AF062D8A84CFFEBD619FE498C28FE4A" ID-CoL="8MBN7" authorityName="Balkenohl" authorityYear="2017" box="[121,365,466,491]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="plurisetosus" status="sp. nov.">
<emphasis id="9F84C549A84CFFEBD619FE498C28FE4A" bold="true" box="[121,365,466,491]" italics="true" pageId="31" pageNumber="32">Orictites plurisetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A84CFFEBD714FE498C89FE4B" box="[372,460,466,490]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="AD4F195BA84CFFEBD619FD868DA4FD21" blockId="31.[121,1006,541,640]" pageId="31" pageNumber="32">
Type material:
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<typeStatus id="724BA7F9A84CFFEBD750FD868CDBFD94" box="[304,414,541,565]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
:
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, labels: white, black printed: “
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–
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S Calapan Balete
</locationDeviation>
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</date>
leg.
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(13)” (
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).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84CFFD5D619FD4C8F66FBE1" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="description">
<paragraph id="AD4F195BA84CFFEBD619FD4C8CABFD51" blockId="31.[121,494,727,752]" box="[121,494,727,752]" pageId="31" pageNumber="32">
<emphasis id="9F84C549A84CFFEBD619FD4C8DB9FD51" bold="true" box="[121,252,727,752]" pageId="31" pageNumber="32">Description</emphasis>
(
<figureCitation id="35CB05DEA84CFFEBD768FD438C08FD51" box="[264,333,728,752]" captionStart="Figs 5–8" captionStartId="8.[159,196,1494,1512]" captionTargetBox="[159,1021,146,1417]" captionTargetId="figure-66@8.[159,590,846,1416]" captionTargetPageId="8" captionText="Figs 5–8: Orictites spp., habitus. 5: O. plurisetosus sp. nov., holotype, male; 6: O. anteriortenuis sp. nov., holotype, male; 7: O. omnipunctatus sp. nov., holotype, male; 8: O. ampliosetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284326" httpUri="https://zenodo.org/record/6284326/files/figure.png" pageId="31" pageNumber="32">Figs 5</figureCitation>
,
<figureCitation id="35CB05DEA84CFFEBD73AFD438C3EFD51" box="[346,379,728,752]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="31" pageNumber="32">23</figureCitation>
; pp. 9, 24)
</paragraph>
<paragraph id="AD4F195BA84CFFEBD619FCB98FE5FCC1" blockId="31.[121,1006,802,1573]" pageId="31" pageNumber="32">
Measurements: Body length
<quantity id="6A08B4BEA84CFFEBD7A8FCB98F77FC9B" box="[456,562,802,826]" metricMagnitude="-3" metricUnit="m" metricValue="3.84" pageId="31" pageNumber="32" unit="mm" value="3.84">3.84 mm</quantity>
, width
<quantity id="6A08B4BEA84CFFEBD4E7FCB98FB5FC9B" box="[647,752,802,826]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="31" pageNumber="32" unit="mm" value="1.02">1.02 mm</quantity>
; ratio length/width of pronotum 1.06; ratio length/width of elytra 2.06.
</paragraph>
<paragraph id="AD4F195BA84CFFEBD6FEFCF68EEAFC0B" blockId="31.[121,1006,802,1573]" pageId="31" pageNumber="32">Colour: Shiny. Brown. Pronotum, all reflexed margins, lines, striae and punctures darker. Labrum and palpi yellowish. Antennae and legs brown.</paragraph>
<paragraph id="AD4F195BA84CFFD4D6FEFC238DA0FC9B" blockId="31.[121,1006,802,1573]" lastBlockId="32.[159,1044,93,1573]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
Head: Slightly narrower than pronotum. Clypeus with small rounded tooth laterally, anterior margin slightly excised, with blunt raised elevation between the two clypeal setae, with punctures reaching up to frons, margin of wings convex, flattened, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to posterior eye level, minutely punctured (100x), with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by indistinct transverse depression, with costate nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina, frons moderately convex, punctured. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes and genae prominent, nearly as wide as anterior angles of pronotum, moderately convex, genae enlarged, rounded, not as high and shorter than eyes, enclosing eyes posteriorly, forming obtuse angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as line of punctures, neck laterally covered with reticulation and punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A873FFD4D412FEC48FCCFED3" attach="left" box="[626,649,351,370]" fontSize="8" pageId="32" pageNumber="33">nd</superScript>
segment, 2
<superScript id="5A85B413A873FFD4D579FEC48E75FED3" attach="left" box="[793,816,351,370]" fontSize="8" pageId="32" pageNumber="33">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with medium sized punctures and some reticulation, Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae of moderate length, reaching up to posterior third of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 moniliform (L/
<geoCoordinate id="C8C47F9CA873FFD4D42EFD438FD0FD51" box="[590,661,728,752]" degrees="1. 0" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="5555" value="-1.0">W 1.0</geoCoordinate>
), antennomeres cylindrical, with shiny areas, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA873FFD4D6A4FCD38F97FAE4" blockId="32.[159,1044,93,1573]" pageId="32" pageNumber="33">Pronotum: Square, longer than wide, sides straight, rounded at anterior angles. Anterior margin straight. Reflexed lateral margin smooth, slightly convex for a short distance before posterior angles. Margin from posterior angle to base running as slightly sigmoid line. Lateral channel distinctly broadened between the two lateral setigerous punctures, smooth. Anterior setigerous puncture located at the end of anterior quarter adjoining convexity of disc of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles indistinctly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as sharp distinct tooth, projecting antero-laterally. Anterior transverse line sharp, deep. Median line deep and moderately broad, surpassing anterior transverse line without joining, fine at base, not joining base. Surface irregularly covered with big punctures, with distinct punctured basal impression. Basal channel broad, deep. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA873FFD5D6A4FAC98E75FEF4" blockId="32.[159,1044,93,1573]" lastBlockId="33.[121,1006,93,1386]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
Elytra: Subcylindrical, sides indistinctly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of fine punctures laterally. Reflexed margin with one or two scars posterior humerus, margin slightly thickened at humerus, bending with channel over rounded humerus up to 5
<superScript id="5A85B413A873FFD4D5F5FA7F8EEDFA56" attach="left" box="[917,936,1508,1527]" fontSize="8" pageId="32" pageNumber="33">th</superScript>
stria. No tooth at humerus. Basal tubercle distinct, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, distinctly punctate, striae 1 and 2 ending free at apex, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura raised. All intervals distinctly convex, 3
<superScript id="5A85B413A872FFD5D4E3FF518FD3FF7C" attach="left" box="[643,662,202,221]" fontSize="8" pageId="33" pageNumber="34">rd</superScript>
and 4
<superScript id="5A85B413A872FFD5D4BCFF518FAAFF7C" attach="left" box="[732,751,202,221]" fontSize="8" pageId="33" pageNumber="34">th</superScript>
broader and distinctly raised at base, both with tubercle at base, 7
<superScript id="5A85B413A872FFD5D41FFF748FD7FEA3" attach="left" box="[639,658,239,258]" fontSize="8" pageId="33" pageNumber="34">th</superScript>
and 8
<superScript id="5A85B413A872FFD5D4BDFF748FB5FEA3" attach="left" box="[733,752,239,258]" fontSize="8" pageId="33" pageNumber="34">th</superScript>
carinate. All intervals shiny on disc. Interval 3 with 16/18, 5 with 19, and 7 with 15 setigerous punctures, somewhat irregularly placed on the respective interval.
</paragraph>
<paragraph id="AD4F195BA872FFD5D6FEFEF98CA6FEDB" blockId="33.[121,1006,93,1386]" box="[158,483,354,378]" pageId="33" pageNumber="34">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA872FFD5D6FEFE138F62FD21" blockId="33.[121,1006,93,1386]" pageId="33" pageNumber="34">
Ventral surface: Epipleuron shiny, with punctures and indistinct reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle, mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with big punctures at base, slightly denser laterally, 3
<superScript id="5A85B413A872FFD5D788FD818CBEFD8C" attach="left" box="[488,507,538,557]" fontSize="8" pageId="33" pageNumber="34">rd</superScript>
to 5
<superScript id="5A85B413A872FFD5D450FD818F06FD8C" attach="left" box="[560,579,538,557]" fontSize="8" pageId="33" pageNumber="34">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA872FFD5D6FEFD168CDDFC9B" blockId="33.[121,1006,93,1386]" pageId="33" pageNumber="34">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, smooth, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA872FFD5D6FEFCD38E96FC71" blockId="33.[121,1006,93,1386]" pageId="33" pageNumber="34">
Male genitalia (
<figureCitation id="35CB05DEA872FFD5D735FCD38CE3FCC1" box="[341,422,840,864]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="33" pageNumber="34">Fig. 23</figureCitation>
): Moderately long, moderately sclerotized. Median lobe slender, moderately arcuate, slightly twisted basally, apex stick-like. Endophallus basally strongly folded, with some bristles towards apex. Parameres of about same length, dorsal one broader, slightly distorted, asetose.
</paragraph>
<paragraph id="AD4F195BA872FFD5D6FEFC468C9DFC54" blockId="33.[121,1006,93,1386]" box="[158,472,989,1013]" pageId="33" pageNumber="34">Female genitalia: Unknown.</paragraph>
<paragraph id="AD4F195BA872FFD5D6FEFB998F66FBE1" blockId="33.[121,1006,93,1386]" pageId="33" pageNumber="34">
Variation: The number of setigerous punctures on the 3
<superScript id="5A85B413A872FFD5D54CFC648E7AFBB3" attach="left" box="[812,831,1023,1042]" fontSize="8" pageId="33" pageNumber="34">rd</superScript>
interval of the elytra seems to vary intraindividually.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A872FFD5D6FEFBD68E85FB5B" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="AD4F195BA872FFD5D6FEFBD68E85FB5B" blockId="33.[121,1006,93,1386]" pageId="33" pageNumber="34">Diagnosis: A small medium-brown species with the knob on the clypeus developed as elevation and indistinct transverse depression behind, no humeral tooth, and a tubercle at the base of interval 3 and 4 of the elytra. Distinguished from all other species by the series of setigerous punctures on intervals 3, 5, and 7, and the eyes and genae which are projecting laterally.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A872FFD5D6FEFA938FB4FA81" box="[158,753,1288,1312]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="AD4F195BA872FFD5D6FEFA938FB4FA81" blockId="33.[121,1006,93,1386]" box="[158,753,1288,1312]" pageId="33" pageNumber="34">
Distribution: Known from Mindoro (
<collectingCountry id="D5E759CBA872FFD5D44CFA938FEBFA81" box="[556,686,1288,1312]" name="Philippines" pageId="33" pageNumber="34">Philippines</collectingCountry>
) only.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A872FFD5D6FEFAB68C0AFACB" pageId="33" pageNumber="34" type="etymology">
<paragraph id="AD4F195BA872FFD5D6FEFAB68C0AFACB" blockId="33.[121,1006,93,1386]" pageId="33" pageNumber="34">Etymology: The name refers to the numerous setae on the intervals 3, 5 and 7 of the elytra.</paragraph>
</subSubSection>
</treatment>
</document>

178
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<taxonomicName id="6AF062D8A84FFFE8D6FFFC468CE8FC57" ID-CoL="8MCCG" authorityName="Balkenohl" authorityYear="2017" box="[159,429,989,1014]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="desuntsetosus" status="sp. nov.">
<emphasis id="9F84C549A84FFFE8D6FFFC468CE8FC57" bold="true" box="[159,429,989,1014]" italics="true" pageId="28" pageNumber="29">Orictites desuntsetosus</emphasis>
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<paragraph id="AD4F195BA84FFFE8D6FFFBB38FBFFB2B" blockId="28.[159,1044,1064,1162]" pageId="28" pageNumber="29">
Type material:
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<typeStatus id="724BA7F9A84FFFE8D737FBB38C83FBE1" box="[343,454,1064,1088]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
:
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, Labels: white, black printed: “leg.
<collectorName id="00057C8DA84FFFE8D5CAFBB38E9FFBE1" box="[938,986,1064,1088]" pageId="28" pageNumber="29">Jäch</collectorName>
(11)
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–
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Luzon
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E Lucena City Quezon NP
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<date id="D94E3F9BA84FFFE8D578FBD68ED3FBC4" box="[792,918,1101,1125]" pageId="28" pageNumber="29" value="1992-11-23">
<collectingDate id="C90AC673A84FFFE8D578FBD68ED3FBC4" box="[792,918,1101,1125]" pageId="28" pageNumber="29" value="1992-11-23">23.11.1992</collectingDate>
</date>
” (
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)
</materialsCitation>
.
<materialsCitation id="1D981306A84FFFE8D6FFFBE98FB3FB2B" ID-GBIF-Occurrence="3497007301" box="[159,758,1138,1162]" collectingDate="1992-11-23" collectionCode="NHMW, CBB" collectorName="Jach" country="Philippines" location="30 km E Lucena City" pageId="28" pageNumber="29" specimenCount="3" specimenCount-female="3" stateProvince="Lucena" typeStatus="paratype">
<typeStatus id="724BA7F9A84FFFE8D6FFFBE98C52FB2B" box="[159,279,1138,1162]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="BBF6D2D2A84FFFE8D740FBE98C0DFB2B" box="[288,328,1138,1162]" pageId="28" pageNumber="29" type="female">3 ♀</specimenCount>
, same data as
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(
<collectionCode id="CBE1819EA84FFFE8D401FBE98FF1FB2B" box="[609,692,1138,1162]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="29" type="Museum">NHMW</collectionCode>
/
<collectionCode id="CBE1819EA84FFFE8D4DCFBE98FB5FB2B" box="[700,752,1138,1162]" collectionName="CBB" pageId="28" pageNumber="29">CBB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84FFFEBD6FFFB798ED9FFD4" lastPageId="31" lastPageNumber="32" pageId="28" pageNumber="29" type="description">
<paragraph id="AD4F195BA84FFFE8D6FFFB798F2CFB5B" blockId="28.[159,617,1250,1275]" box="[159,617,1250,1275]" pageId="28" pageNumber="29">
<emphasis id="9F84C549A84FFFE8D6FFFB798C67FB5A" bold="true" box="[159,290,1250,1275]" pageId="28" pageNumber="29">Description</emphasis>
(
<figureCitation id="35CB05DEA84FFFE8D74EFB798C31FB5B" box="[302,372,1250,1274]" captionStart="Figs 1–4" captionStartId="7.[121,158,1494,1512]" captionTargetBox="[145,1002,80,1436]" captionTargetPageId="7" captionText="Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284320" httpUri="https://zenodo.org/record/6284320/files/figure.png" pageId="28" pageNumber="29">Figs 4</figureCitation>
,
<figureCitation id="35CB05DEA84FFFE8D7E1FB798CD8FB5B" box="[385,413,1250,1274]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="28" pageNumber="29">22</figureCitation>
,
<figureCitation id="35CB05DEA84FFFE8D7CAFB798C89FB5B" box="[426,460,1250,1274]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="28" pageNumber="29">34</figureCitation>
; pp. 8, 24, 35)
</paragraph>
<paragraph id="AD4F195BA84FFFE8D6FFFAB68F80FA31" blockId="28.[159,1044,1324,1573]" pageId="28" pageNumber="29">
Measurements (n = 4): Body length
<quantity id="6A08B4BEA84FFFE8D455FAB68F9FFAE4" box="[565,730,1325,1349]" metricMagnitude="-3" metricUnit="m" metricValue="4.515" metricValueMax="4.65" metricValueMin="4.38" pageId="28" pageNumber="29" unit="mm" value="4.515" valueMax="4.65" valueMin="4.38">4.38–4.65 mm</quantity>
(
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=
<quantity id="6A08B4BEA84FFFE8D577FAB68E3BFAE4" box="[791,894,1325,1349]" metricMagnitude="-3" metricUnit="m" metricValue="4.49" pageId="28" pageNumber="29" unit="mm" value="4.49">4.49 mm</quantity>
), width
<quantity id="6A08B4BEA84FFFE8D5B4FAB68C44FACB" metricMagnitude="-3" metricUnit="m" metricValue="1.28" metricValueMax="1.31" metricValueMin="1.25" pageId="28" pageNumber="29" unit="mm" value="1.28" valueMax="1.31" valueMin="1.25">1.25– 1.31 mm</quantity>
(
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=
<quantity id="6A08B4BEA84FFFE8D75FFAC98CE3FACB" box="[319,422,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="28" pageNumber="29" unit="mm" value="1.29">1.29 mm</quantity>
), ratio length/width of pronotum 0.98–0.99 (
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= 0.98), ratio length/width of elytra 2.25–2.31 (
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= 2.29).
</paragraph>
<paragraph id="AD4F195BA84FFFE8D6A4FA068E6FFA7B" blockId="28.[159,1044,1324,1573]" pageId="28" pageNumber="29">Colour: Shiny. Brown. Reflexed margins, lines, striae, and punctures darker. Labrum and palpi yellowish. Antennae and legs brown.</paragraph>
<paragraph id="AD4F195BA84FFFE9D6A4FA738F8CFAE4" blockId="28.[159,1044,1324,1573]" lastBlockId="29.[121,1006,93,1573]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Head: A quarter narrower than pronotum. Clypeus deeply excised, with small rounded tooth laterally nearly fused with clypeal wings, with blunt raised elevation between the two clypeal setae, anteriorly smooth, posterior part of elevation covered with big punctures, margin of wings convex, flattened, divided from middle part of clypeus by slight carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to posterior eye level, smooth, with reflexed rounded margin, as supraorbital carina, frons separated from clypeus by flat transverse depression with punctures, with costate posteriorly diverging frontal carinae at each side, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina, frons moderately convex, smooth. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes relatively big, convex, genae distinct, enclosing eyes posteriorly, forming obtuse angle at neck. Grooves for reception of antennae short, length 0.3 of eye diameter. Neck constriction marked by punctures with wider distance, neck laterally covered with reticulation and densely scattered set medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, broadened at base, acutely curved at apex, carinae of right scrobe interrupted at middle, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A84EFFE9D74DFC148C01FC03" attach="left" box="[301,324,911,930]" fontSize="8" pageId="29" pageNumber="30">nd</superScript>
segment, 2
<superScript id="5A85B413A84EFFE9D7ACFC148CA6FC03" attach="left" box="[460,483,911,930]" fontSize="8" pageId="29" pageNumber="30">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with punctures. Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae relatively long, reaching up to posterior setigerous puncture of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 submoniliform (L/
<geoCoordinate id="C8C47F9CA84EFFE9D5B2FB798DE9FA81" degrees="0. 89" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="-0.89">W 0.89</geoCoordinate>
), antennomeres somewhat flattened, with shiny areas, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA84EFFEAD6FEFAC98EF7FEF4" blockId="29.[121,1006,93,1573]" lastBlockId="30.[159,1044,93,1573]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">Pronotum: Square, as long as wide, sides straight, slightly diverging posteriorly. Anterior margin straight. Reflexed lateral margin smooth. Margin from posterior angle to base running as sigmoid line. Lateral channel conspicuously broad between the two lateral setigerous punctures, nearly smooth. Anterior setigerous puncture located at the end of anterior quarter approximated to the convexity of the disc of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle indistinctly projecting, angled, formed by reflexed lateral margin, posterior angle developed as distinct tooth, projecting antero-laterally. Anterior transverse line formed by some distant punctures. Median line deep, broad, surpassing anterior transverse line without joining, not joining base. Surface with some irregularly situated medium sized punctures, with basal impression. Basal channel broad, deep. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA84DFFEAD6A4FEF98C45FCB4" blockId="30.[159,1044,93,1573]" pageId="30" pageNumber="31">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of small punctures laterally. Reflexed margin with two indistinct scars (120x) in anterior third, bending with channel over rounded humerus up to 5
<superScript id="5A85B413A84DFFEAD40CFE6F8F3AFDA6" attach="left" box="[620,639,500,519]" fontSize="8" pageId="30" pageNumber="31">th</superScript>
stria. No tooth at humerus. Basal tubercle distinct, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, indistinctly punctate, striae 1 and 2 ending free at apex, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura raised. All intervals distinctly convex, 3
<superScript id="5A85B413A84DFFEAD7D0FD348C86FD63" attach="left" box="[432,451,687,706]" fontSize="8" pageId="30" pageNumber="31">rd</superScript>
broader and distinctly raised at base, with tubercle at base, 8
<superScript id="5A85B413A84DFFEAD76EFD4F8C64FD46" attach="left" box="[270,289,724,743]" fontSize="8" pageId="30" pageNumber="31">th</superScript>
carinate. All intervals shiny on disc. Interval 3 or any other interval asetose.
</paragraph>
<paragraph id="AD4F195BA84DFFEAD6A4FCB98F4DFC9B" blockId="30.[159,1044,93,1573]" box="[196,520,802,826]" pageId="30" pageNumber="31">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA84DFFEAD6A4FCD38F4FFBE1" blockId="30.[159,1044,93,1573]" pageId="30" pageNumber="31">
Ventral surface: Epipleuron shiny, with scattered punctures and indistinct reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle. Mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with big punctures at base, denser laterally, 3
<superScript id="5A85B413A84DFFEAD7B1FC418CA1FC4C" attach="left" box="[465,484,986,1005]" fontSize="8" pageId="30" pageNumber="31">rd</superScript>
to 5
<superScript id="5A85B413A84DFFEAD479FC418F69FC4C" attach="left" box="[537,556,986,1005]" fontSize="8" pageId="30" pageNumber="31">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA84DFFEAD6A4FBD68CFBFB5B" blockId="30.[159,1044,93,1573]" pageId="30" pageNumber="31">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, smooth, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA84DFFEAD6A4FA938CA1FA31" blockId="30.[159,1044,93,1573]" pageId="30" pageNumber="31">
Male genitalia (
<figureCitation id="35CB05DEA84DFFEAD711FA938CFBFA81" box="[369,446,1288,1312]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="30" pageNumber="31">Fig. 22</figureCitation>
): Moderately sclerotized. Median lobe slender, moderately arcuate, not twisted, apex stick-like. Endophallus without visible teeth. Parameres of about same length, both slender, each distorted anterior apex, with minute seta preapically.
</paragraph>
<paragraph id="AD4F195BA84DFFEAD6A4FA068E6AF984" blockId="30.[159,1044,93,1573]" pageId="30" pageNumber="31">
Female genitalia. (
<figureCitation id="35CB05DEA84DFFEAD7FFFA068CB7FA14" box="[415,498,1437,1461]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="30" pageNumber="31">Fig. 34</figureCitation>
): Coxostylus slender, nearly straight in basal part, gently curved to apex, with short obtuse knob-like tooth dorsally at base, with one seta preapically, with one blunt and one acute short ensiform seta, with five nematiform setae. Ramus of moderate size.
</paragraph>
<paragraph id="AD4F195BA84CFFEBD6FEFFC68ED9FFD4" blockId="31.[121,1006,93,378]" box="[158,924,93,117]" pageId="31" pageNumber="32">
Variation: In one of the
<typeStatus id="724BA7F9A84CFFEBD7C5FFC68F5DFFD4" box="[421,536,93,117]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
there are single punctures visible.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84CFFEBD6FEFF198E79FE91" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="AD4F195BA84CFFEBD6FEFF198E79FE91" blockId="31.[121,1006,93,378]" pageId="31" pageNumber="32">Diagnosis: A small medium-brown species with the knob on the clypeus developed as elevation and indistinct transverse depression behind, no humeral tooth, and a tubercle only at the base of interval 3 of the elytra. Distinguished from all other species by the completely missing setigerous punctures on intervals of the elytra and the deeply excised clypeus.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84CFFEBD6FEFEA68E53FEF4" box="[158,790,317,341]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="AD4F195BA84CFFEBD6FEFEA68E53FEF4" blockId="31.[121,1006,93,378]" box="[158,790,317,341]" pageId="31" pageNumber="32">
Distribution: Known from the Luzon Island (
<collectingCountry id="D5E759CBA84CFFEBD4E7FEA68E49FEF4" box="[647,780,317,341]" name="Philippines" pageId="31" pageNumber="32">Philippines</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A84CFFEBD6FEFEF98E24FEDB" box="[158,865,354,378]" pageId="31" pageNumber="32" type="etymology">
<paragraph id="AD4F195BA84CFFEBD6FEFEF98E24FEDB" blockId="31.[121,1006,93,378]" box="[158,865,354,378]" pageId="31" pageNumber="32">Etymology: The name refers to the missing setae on the elytra.</paragraph>
</subSubSection>
</treatment>
</document>

238
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<emphasis id="9F84C549A850FFF7D6A7FFC68C6CFFD7" bold="true" box="[199,297,93,118]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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<paragraph id="AD4F195BA850FFF7D619FF338DA9FF61" blockId="3.[121,616,168,453]" box="[121,236,168,192]" pageId="3" pageNumber="4">Catalogue</paragraph>
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<emphasis id="9F84C549A850FFF7D619FF568D9CFF44" box="[121,217,205,229]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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<emphasis id="9F84C549A850FFF7D619FF698D9CFEAB" box="[121,217,242,266]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
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;
<treatmentCitation id="2C513F4AA850FFF7D706FF698F51FEAB" author="Csiki E." box="[358,532,242,266]" page="638" pageId="3" pageNumber="4" year="1933">
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<emphasis id="9F84C549A850FFF7D619FE838D9CFE91" box="[121,217,280,304]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
ANDREWES
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;
<treatmentCitation id="2C513F4AA850FFF7D706FE838F25FE91" author="Louwerens, C. J." box="[358,608,280,304]" page="172" pageId="3" pageNumber="4" year="1964">
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<emphasis id="9F84C549A850FFF7D619FEA68D9CFEF4" box="[121,217,317,341]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
ANDREWES
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<emphasis id="9F84C549A850FFF7D619FEF98D9CFEDB" box="[121,217,354,378]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
ANDREWES
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<emphasis id="9F84C549A850FFF7D619FE138D9CFE01" box="[121,217,392,416]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
ANDREWES
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;
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<emphasis id="9F84C549A850FFF7D619FE368D9CFE64" box="[121,217,429,453]" italics="true" pageId="3" pageNumber="4">Orictites</emphasis>
ANDREWES
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;
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.
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<subSubSection id="E5EA4AD0A850FFF7D619FD868E84FD94" box="[121,961,541,565]" pageId="3" pageNumber="4" type="type_taxon">
<paragraph id="AD4F195BA850FFF7D619FD868E84FD94" blockId="3.[121,961,541,565]" box="[121,961,541,565]" pageId="3" pageNumber="4">
<heading id="F607AE37A850FFF7D619FD868E84FD94" box="[121,961,541,565]" centered="true" fontSize="12" level="2" pageId="3" pageNumber="4" reason="2">
<typeStatus id="724BA7F9A850FFF7D619FD868DEBFD94" box="[121,174,541,565]" pageId="3" pageNumber="4">Type</typeStatus>
species:
<taxonomicName id="6AF062D8A850FFF7D777FD868FEFFD94" ID-CoL="8M924" authority="ANDREWES, 1931" authorityName="ANDREWES" authorityYear="1931" box="[279,682,541,565]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="minotaur">
<emphasis id="9F84C549A850FFF7D777FD868CA1FD94" box="[279,484,541,565]" italics="true" pageId="3" pageNumber="4">Orictites minotaur</emphasis>
<bibRefCitation id="C96164AAA850FFF7D78BFD868FEFFD94" author="Andrewes, H. E." box="[491,682,541,565]" pageId="3" pageNumber="4" pagination="431 - 485" refId="ref27519" refString="Andrewes, H. E. (1931): On the Carabidae of the Mount Kinabalu. - Journal of the FMS Museum, Kuala Lumpur 16: 431 - 485." type="journal article" year="1931">ANDREWES, 1931</bibRefCitation>
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, by original designation.
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<paragraph id="AD4F195BA850FFF0D619FDF38FE1FF61" blockId="3.[121,1007,616,1573]" lastBlockId="4.[159,1044,93,192]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Diagnosis: General appearance more or less like a
<taxonomicName id="6AF062D8A850FFF7D4AAFDF38E52FD21" authorityName="LATREILLE" authorityYear="1802" box="[714,791,616,640]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="9F84C549A850FFF7D4AAFDF38E52FD21" box="[714,791,616,640]" italics="true" pageId="3" pageNumber="4">Clivina</emphasis>
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. Clypeus variable, straight anteriorly or excised or with teeth, with raised knob between the two clypeal setae or with blunt elevation, wings small or well-developed, more or less projecting, divided from supraantennal plates by distinct notches and from middle part of clypeus by distinct or indistinct notches. Frons separated from clypeus by band of big punctures and/or by broad flat furrow. Longitudinal frontal carinae distinct. Supraorbital carinae distinct. Eyes oval, flattened, enclosed posteriorly by distinctly vaulted genae of different length, temporae carinate; antennae relatively short, with some long setae among the usual pubescence. Labrum 7-setose. Mandibles strong, right or left one with tooth at middle or basally. Pronotum square, almost flattened with tooth at posterior angle, anterior angles formed by reflexed lateral margin, rounded and projecting anteriorly or angled. Reflexed lateral margin more or less crenulated. Lateral channel distinctly broad, separated from disc by longitudinal vault, median line conspicuously deep and broad, surface in most of the species covered with deep and broad punctures. Basal channel always broad and deep. Elytra subcylindrical, stria 1–4 free at base, intervals more or less convex, asetose or with setigerous punctures on interval 3, or 3, 5, and 7. Third interval of elytra broadened and raised at base with distinct tubercle, in most species the same is true for interval 4. Middle tibia with spur near apex furnished with seta. Terminal abdominal sternite with two widely separated apical setae at each side. Ventral surface almost covered with punctures. Distinguished from all other
<taxonomicName id="6AF062D8A850FFF7D740FA068C28FA14" authorityName="LATREILLE" authorityYear="1802" box="[288,365,1437,1461]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="9F84C549A850FFF7D740FA068C28FA14" box="[288,365,1437,1461]" italics="true" pageId="3" pageNumber="4">Clivina</emphasis>
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-related genera by the following combination of characters: the flattened eyes with distinctly enlarged genae, the distinct carinae on the head, the transverse square and flattened pronotum with unusually broad lateral channel, the big and usually numerous punctures on the head, pronotum, and ventral surface, the broad and deep base of the pronotum, the distinctly convex intervals of the elytra, and the distinctly broadened, raised and tubercle possessing intervals 3 and 4 at base.
</paragraph>
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<paragraph id="AD4F195BA857FFF0D6FFFE8C8C2BFE91" blockId="4.[159,366,279,304]" box="[159,366,279,304]" pageId="4" pageNumber="5">
<emphasis id="9F84C549A857FFF0D6FFFE8C8C2BFE91" bold="true" box="[159,366,279,304]" pageId="4" pageNumber="5">Genus description</emphasis>
</paragraph>
<paragraph id="AD4F195BA857FFF0D6FFFEF98F11FEDB" blockId="4.[159,1044,354,1573]" box="[159,596,354,378]" pageId="4" pageNumber="5">Size: Small to medium-sized, elongate.</paragraph>
<paragraph id="AD4F195BA857FFF0D6FFFE138FB9FE01" blockId="4.[159,1044,354,1573]" box="[159,764,392,416]" pageId="4" pageNumber="5">Colour: Shiny, without metallic lustre, black or brown.</paragraph>
<paragraph id="AD4F195BA857FFF0D6A4FE368C26FA14" blockId="4.[159,1044,354,1573]" pageId="4" pageNumber="5">
Head: Wide. Clypeus straight, excised, or with teeth anteriorly, limited laterally by small rounded or sharp tooth, with raised knob in between the two clypeal setae or with blunt elevation; wings always well-developed, rounded or angled, hollowed out, divided from supraantennal plates and from middle part of clypeus by notches. Supraantennal plates vaulted, with reflexed rounded margin, frons separated from clypeus by transverse furrow or depression, with or without big punctures, with nearly parallel running carinae at each side posteriorly, each carina with furrow mesially, carinae enclosing the convex frons; frons smooth or with punctures. Supraorbital setae located in broad furrow between eye and frontal carina, the posterior one arising from a small tubercle. Eyes oval, not big, orbit projecting laterally, tempora with distinct supraorbital carina, subtemporae and genae distinctly vaulted, rounded, nearly as high but slightly shorter than eyes, distinctly enclosing eyes posteriorly. Grooves for reception of antennae short. Neck constriction formed by punctures or indistinct. Labrum 7-setose, ciliate laterally, almost with isodiametric reticulation. Mandibles shorter than head, distinctly broadened at base, acute at apex, upper margin of scrobe of both mandibles sharply carinate, mandibular teeth differently developed in the two subgenera. Maxilla bent slightly sigmoidal. Apical maxillary palpomere slender, fusiform bottle-like. Apical labial palpomere slightly arcuate, fusiform bottle-like or securiform, second segment bisetose. Ligula somewhat truncated at apex, with one seta. Paraglossae slender, membranous.
<collectingRegion id="6F34D7B9A857FFF0D7B4FB268F77FB74" box="[468,562,1213,1237]" country="Myanmar" name="Chin" pageId="4" pageNumber="5">Mentum</collectingRegion>
and submentum divided by sharp furrow, with two pairs of setae each, median tooth broad, obtuse at apex, bisetose, lobes smooth or with wrinkles, longer than tooth, acute at apex, margins with carinae. Antennae short, not reaching over two thirds of pronotum, scapus with one seta dorsoapically, scapus and pedicellus in most of the cases convered with fine reticulation, segments 4–11 densely pubescent, with few additional long setae.
</paragraph>
<paragraph id="AD4F195BA857FFF1D6A4FA598C51FEF4" blockId="4.[159,1044,354,1573]" lastBlockId="5.[121,1006,93,1573]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">Pronotum: Square, as long as wide or broader than long, almost flattened. Anterior margin straight or slightly bisinuate. Reflexed lateral margin smooth or crenulated. Lateral channel distinctly broadened between the two lateral setigerous punctures, basal channel broad and deep. Anterior setigerous puncture located backwards, the posterior one located at level of posterior angle, removed from lateral channel. Anterior angles more or less projecting anteriorly, posterior angles well developed. Anterior transverse line sharp, punctured, or indistinct. Median line deep and conspicuously broad, often with blunt carina bilaterally. Surface nearly always covered with punctures of different size.</paragraph>
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Elytra: Subcylindrical. Marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, with second row of smaller punctures. Reflexed margin with more or less developed crenulation, more distinct at humerus. Reflexed lateral margin bending parallel to channel over rounded humerus up to stria 5, with or without humeral tooth. Basal tubercle big, with setigerous puncture, situated at declivity of 1
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stria. Striae 1–4 free at base, all striae deep, with more or less distinct punctures, striae 1–2 either ending free at apex or joining apically, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura raised. Intervals conspicuously convex, with increasing distinctness laterally, 3
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interval and in some cases the4
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as well broader and raised at base, with big tubercle at base, 7
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running to apex as more or less obtuse carina, 8
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small, carinate. Intervals asetose or with setigerous punctures on interval 3, or 3, 5, and 7.
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<paragraph id="AD4F195BA856FFF1D6FEFCF68C9AFC24" blockId="5.[121,1006,93,1573]" box="[158,479,877,901]" pageId="5" pageNumber="6">Hind wings: Well developed.</paragraph>
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Ventral surface: Proepisternum with big scattered punctures and indistinct reticulation, in a few species with some small punctures, submarginal furrow engraved. Metasternum smooth at middle, punctured laterally. Epipleura somewhat broadened basally, with row of big punctures. Sternites 3
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to 5
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with paralateral ambulatory setae at each side, covered densely and irregularly with big punctures or punctures limited to lateral margin, ventral strigae distinct, with transverse reticulation; terminal sternite with two widely separated apical setae at each side.
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<paragraph id="AD4F195BA856FFF1D6FEFB268E54FAE4" blockId="5.[121,1006,93,1573]" pageId="5" pageNumber="6">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral and lateral. Basal tarsomere distinctly elongated. Mesotibia with long apical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
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Male genitalia: In general of
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-
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, with spatulate or stick-like apex.
</paragraph>
<paragraph id="AD4F195BA856FFF1D6FEFAE38C2EFA7B" blockId="5.[121,1006,93,1573]" pageId="5" pageNumber="6">Female genitalia: Stylomere more or less sigmoidal, with or without preapical setae, with one or two ensiform seta at middle and some nematiform setae laterally in basal half.</paragraph>
<paragraph id="AD4F195BA856FFF1D6FEFA738E1EF9A1" blockId="5.[121,1006,93,1573]" box="[158,859,1512,1536]" pageId="5" pageNumber="6">Sexual dimorphism: External sexual differences not observed.</paragraph>
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Distribution. South
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including
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, Sumatra, Borneo, and Sulawesi. Specimens were also found in North East
<collectingCountry id="D5E759CBA855FFF2D48CFFC68E61FFD4" box="[748,804,93,117]" name="India" pageId="6" pageNumber="7">India</collectingCountry>
(
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Hills) and in the West of Papua (
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; p. 63).
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<paragraph id="AD4F195BA855FFF2D6A4FF338CB0FEAB" blockId="6.[159,1043,93,266]" pageId="6" pageNumber="7">Habitat: Not well known. Specific labels of some specimens indicate finds in mountainous areas. Some species were found under bark of trees and rotten wood, e.g. in a pine forest.</paragraph>
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<emphasis id="9F84C549A855FFF2D6FFFEF98C54FEDA" bold="true" box="[159,273,354,379]" pageId="6" pageNumber="7">Subgenus</emphasis>
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<emphasis id="9F84C549A855FFF2D777FEF98C3CFEDA" bold="true" box="[279,377,354,379]" italics="true" pageId="6" pageNumber="7">Orictites</emphasis>
<bibRefCitation id="C96164AAA855FFF2D71FFEF98F7BFEDB" author="Andrewes, H. E." box="[383,574,354,378]" pageId="6" pageNumber="7" pagination="431 - 485" refId="ref27519" refString="Andrewes, H. E. (1931): On the Carabidae of the Mount Kinabalu. - Journal of the FMS Museum, Kuala Lumpur 16: 431 - 485." type="journal article" year="1931">ANDREWES, 1931</bibRefCitation>
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(
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,
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,
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; pp. 8, 15, 19)
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species
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, by original designation.
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Diagnosis: Black. General appearance like small-sized elongate
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Anterior margin of clypeus with four distinctly projecting teeth, clypeal wings sharply angled, tooth-like projecting. Mandibles as long as head; dorsal margin of scrobe of both mandibles at base with carina-like tooth, right mandible with distinct tooth at middle, left mandible with indistinct knob like tooth basally. Apical maxillary palpi elongate, bottle-like fusiform, labial palpi slender, slightly arcuate bottle-like fusiform. Pronotum distinctly broader than long, convex at middle of disc, anterior angles conspicuously projecting. Spatula of aedeagus broad. Parameres with two apical and one subapical setae. Coxostylus with one subapical seta. Body length
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<mods:identifier id="DCDDF1E913149EAB630D556BE547C3E3" type="DOI">10.5169/seals-787048</mods:identifier>
<mods:identifier id="B792B8833FE6FA15B55AD3A9A0C62390" type="GBIF-Dataset">8c468ca9-0ee3-4200-b5a3-6adfd98caaea</mods:identifier>
<mods:identifier id="32594CBD082EEA61B8EBD10DBB11102E" type="ISSN">2624-9170</mods:identifier>
<mods:identifier id="EBE421B65B09F430DF2D7D9CB8CB7B7A" type="Zenodo-Dep">6284316</mods:identifier>
</mods:mods>
<treatment id="2559A84DA855FFF8D6FFFC2C8956FAE4" ID-DOI="http://doi.org/10.5281/zenodo.6315602" ID-GBIF-Taxon="193586548" ID-Zenodo-Dep="6315602" LSID="urn:lsid:plazi:treatment:2559A84DA855FFF8D6FFFC2C8956FAE4" httpUri="http://treatment.plazi.org/id/2559A84DA855FFF8D6FFFC2C8956FAE4" lastPageId="12" lastPageNumber="13" pageId="6" pageNumber="7">
<subSection id="C47F00B0A855FFF2D6FFFC2C8E2FFC71" box="[159,874,951,976]" pageId="6" pageNumber="7" type="multiple">
<subSubSection id="E5EA4AD0A855FFF2D6FFFC2C8E2FFC71" box="[159,874,951,976]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="AD4F195BA855FFF2D6FFFC2C8E2FFC71" blockId="6.[159,874,951,976]" box="[159,874,951,976]" pageId="6" pageNumber="7">
<heading id="F607AE37A855FFF2D6FFFC2C8E2FFC71" box="[159,874,951,976]" centered="true" fontSize="12" level="2" pageId="6" pageNumber="7" reason="2">
<emphasis id="9F84C549A855FFF2D6FFFC2C8CCAFC71" bold="true" box="[159,399,951,976]" pageId="6" pageNumber="7">
Subgenus
<taxonomicName id="6AF062D8A855FFF2D777FC2C8CCAFC71" authorityName="Balkenoh" box="[279,399,951,976]" class="Insecta" family="Carabidae" genus="Semictites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="subgen. nov.">
<emphasis id="9F84C549A855FFF2D777FC2C8CCAFC71" bold="true" box="[279,399,951,976]" italics="true" pageId="6" pageNumber="7">Semictites</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="84B77832A855FFF2D7F5FC238F63FC71" box="[405,550,952,976]" pageId="6" pageNumber="7" rank="subGenus">subgen. nov.</taxonomicNameLabel>
(
<figureCitation id="35CB05DEA855FFF2D451FC238F30FC71" box="[561,629,952,976]" captionStart="Figs 1–4" captionStartId="7.[121,158,1494,1512]" captionTargetBox="[145,1002,80,1436]" captionTargetPageId="7" captionText="Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284320" httpUri="https://zenodo.org/record/6284320/files/figure.png" pageId="6" pageNumber="7">Figs 2</figureCitation>
,
<figureCitation id="35CB05DEA855FFF2D4E3FC238FDAFC71" box="[643,671,952,976]" captionStart="Figs 15–16" captionStartId="14.[159,196,1489,1507]" captionTargetBox="[159,1043,104,1429]" captionTargetId="figure-49@14.[159,1043,104,761]" captionTargetPageId="14" captionText="Figs 15–16: Orictites ssp., head and pronotum, lateral view; 15: Orictites (subgenus Orictites) minotaur ANDREWES, 1931,; 16: Orictites (subgenus Semictites) costulipennis BATES, 1892." figureDoi="http://doi.org/10.5281/zenodo.6284338" httpUri="https://zenodo.org/record/6284338/files/figure.png" pageId="6" pageNumber="7">16</figureCitation>
,
<figureCitation id="35CB05DEA855FFF2D4CCFC238F88FC71" box="[684,717,952,976]" captionStart="Figs 17–18" captionStartId="18.[159,196,780,798]" captionTargetBox="[159,1043,91,754]" captionText="Figs 17–18: Orictites spp., Maxilla, maxillary and labial palpomere. 17: O. minotaur ANDREWES, 1931 (subgenus Orictites); 18: O. costulipennis BATES, 1892 (subgenus Semictites subg. nov.)." figureDoi="http://doi.org/10.5281/zenodo.6284342" httpUri="https://zenodo.org/record/6284342/files/figure.png" pageId="6" pageNumber="7">18</figureCitation>
; pp. 8, 15, 19)
</heading>
</paragraph>
</subSubSection>
</subSection>
<subSubSection id="E5EA4AD0A855FFF2D6FFFB998944FBBB" box="[159,1025,1026,1050]" pageId="6" pageNumber="7" type="type_taxon">
<paragraph id="AD4F195BA855FFF2D6FFFB998944FBBB" blockId="6.[159,1043,1026,1536]" box="[159,1025,1026,1050]" pageId="6" pageNumber="7">
<heading id="F607AE37A855FFF2D6FFFB998944FBBB" box="[159,1025,1026,1050]" centered="true" fontSize="12" level="2" pageId="6" pageNumber="7" reason="2">
<typeStatus id="724BA7F9A855FFF2D6FFFB998D91FBBB" box="[159,212,1026,1050]" pageId="6" pageNumber="7">Type</typeStatus>
species
<taxonomicName id="6AF062D8A855FFF2D755FB998FACFBBB" ID-CoL="8MFK7" authority="BATES, 1892: 281" authorityName="BATES Compared" authorityPageNumber="281" authorityYear="1892" box="[309,745,1026,1050]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A855FFF2D755FB998F65FBBB" box="[309,544,1026,1050]" italics="true" pageId="6" pageNumber="7">Clivina costulipennis</emphasis>
<bibRefCitation id="C96164AAA855FFF2D446FB998FACFBBB" author="Bates, H. W." box="[550,745,1026,1050]" pageId="6" pageNumber="7" pagination="267 - 428" refId="ref27786" refString="Bates, H. W. (1892): Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. - Annali del Museo Civico di Storia Naturale di Genova 32: 267 - 428." type="journal article" year="1892">BATES, 1892: 281</bibRefCitation>
</taxonomicName>
, by present designation.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A855FFF2D6FFFBB38C74FA14" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="AD4F195BA855FFF2D6FFFBB38C74FA14" blockId="6.[159,1043,1026,1536]" pageId="6" pageNumber="7">
Diagnosis: Black or brown. General appearance like a
<taxonomicName id="6AF062D8A855FFF2D57CFBB38E2CFBE1" authorityName="LATREILLE" authorityYear="1802" box="[796,873,1064,1088]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="9F84C549A855FFF2D57CFBB38E2CFBE1" box="[796,873,1064,1088]" italics="true" pageId="6" pageNumber="7">Clivina</emphasis>
</taxonomicName>
. Anterior margin of clypeus straight or excised, without teeth, clypeal wings small, obtuse angled or rounded. Mandibles distinctly shorter than head; dorsal margin of scrobe of both mandibles at base without elevated tooth, right mandible with indistinct tooth at base, left mandible with small but distinct tooth slightly anterior to base. Apical maxillary palpomere securiform, apical labial palpomere nearly straight, fusiform, slightly bottle-like. Pronotum about as long as wide, almost flattened on disc, anterior angles angulate, more or less projecting. Aedeagus with spatula or stick-like. Parameres asetose or with up to two apical setae. Coxostylus with one or two subapical setae. Body length smaller than
<quantity id="6A08B4BEA855FFF2D6B8FA068C68FA14" box="[216,301,1437,1461]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="7" unit="mm" value="7.4">7.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A855FFF2D6FFFA598C14F9A1" pageId="6" pageNumber="7" type="etymology">
<paragraph id="AD4F195BA855FFF2D6FFFA598C14F9A1" blockId="6.[159,1043,1026,1536]" pageId="6" pageNumber="7">
Etymology: The name refers to the numerous similarities shared with the subgenus
<emphasis id="9F84C549A855FFF2D68AFA738C14F9A1" box="[234,337,1512,1536]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="6AF062D8A855FFF2D68AFA738C08F9A1" authorityName="ANDREWES" authorityYear="1931" box="[234,333,1512,1536]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Orictites</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<caption id="F98F49D3A854FFF3D619FA4D8C0AF9BC" ID-DOI="http://doi.org/10.5281/zenodo.6284320" ID-Zenodo-Dep="6284320" httpUri="https://zenodo.org/record/6284320/files/figure.png" pageId="7" pageNumber="8" startId="7.[121,158,1494,1512]" targetBox="[145,1002,80,1436]" targetPageId="7">
<paragraph id="AD4F195BA854FFF3D619FA4D8C0AF9BC" blockId="7.[121,1005,1494,1565]" pageId="7" pageNumber="8">
Figs 1–4:
<taxonomicName id="6AF062D8A854FFF3D6AEFA4D8C03FA49" box="[206,326,1494,1512]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9F84C549A854FFF3D6AEFA4D8C5CFA49" bold="true" box="[206,281,1494,1512]" italics="true" pageId="7" pageNumber="8">Orictites</emphasis>
spp.
</taxonomicName>
, habitus. 1:
<taxonomicName id="6AF062D8A854FFF3D7CFFA4D8FE9FA49" authority="ANDREWES, 1931" authorityName="ANDREWES" authorityYear="1931" box="[431,684,1494,1512]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="minotaur">
<emphasis id="9F84C549A854FFF3D7CFFA4D8CF9FA49" bold="true" box="[431,444,1494,1512]" italics="true" pageId="7" pageNumber="8">O</emphasis>
.
<emphasis id="9F84C549A854FFF3D7A7FA4D8F50FA49" bold="true" box="[455,533,1494,1512]" italics="true" pageId="7" pageNumber="8">minotaur</emphasis>
<bibRefCitation id="C96164AAA854FFF3D47BFA4D8FE9FA49" author="Andrewes, H. E." box="[539,684,1494,1512]" pageId="7" pageNumber="8" pagination="431 - 485" refId="ref27519" refString="Andrewes, H. E. (1931): On the Carabidae of the Mount Kinabalu. - Journal of the FMS Museum, Kuala Lumpur 16: 431 - 485." type="journal article" year="1931">ANDREWES, 1931</bibRefCitation>
</taxonomicName>
, lectotype, male; 2:
<taxonomicName id="6AF062D8A854FFF3D501FA4D8DA6F9A3" authority="BATES, 1892" authorityName="BATES Compared" authorityYear="1892" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A854FFF3D501FA4D8E2BFA49" bold="true" box="[865,878,1494,1512]" italics="true" pageId="7" pageNumber="8">O</emphasis>
.
<emphasis id="9F84C549A854FFF3D518FA4D8EA8FA49" bold="true" box="[888,1005,1494,1512]" italics="true" pageId="7" pageNumber="8">costulipennis</emphasis>
<bibRefCitation id="C96164AAA854FFF3D619FA6B8DA6F9A3" author="Bates, H. W." box="[121,227,1520,1538]" pageId="7" pageNumber="8" pagination="267 - 428" refId="ref27786" refString="Bates, H. W. (1892): Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. - Annali del Museo Civico di Storia Naturale di Genova 32: 267 - 428." type="journal article" year="1892">BATES, 1892</bibRefCitation>
</taxonomicName>
, holotype, male; 3:
<taxonomicName id="6AF062D8A854FFF3D7F2FA6B8FE6F9A3" authority="LouWERENS, 1964" authorityName="LouWERENS" authorityYear="1964" box="[402,675,1520,1538]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A854FFF3D7F2FA6B8CDAF9A3" bold="true" box="[402,415,1520,1538]" italics="true" pageId="7" pageNumber="8">O</emphasis>
.
<emphasis id="9F84C549A854FFF3D7CAFA6B8F47F9A3" bold="true" box="[426,514,1520,1538]" italics="true" pageId="7" pageNumber="8">mjoebergi</emphasis>
<bibRefCitation id="C96164AAA854FFF3D466FA6B8FE6F9A3" author="Louwerens, C. J." box="[518,675,1520,1538]" pageId="7" pageNumber="8" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">LouWERENS, 1964</bibRefCitation>
</taxonomicName>
, lectotype, male; 4:
<taxonomicName id="6AF062D8A854FFF3D537FA6B8EA8F9A3" authorityName="Balkenohl" authorityYear="2017" box="[855,1005,1520,1538]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="desuntsetosus" status="sp. nov.">
<emphasis id="9F84C549A854FFF3D537FA6B8E21F9A3" bold="true" box="[855,868,1520,1538]" italics="true" pageId="7" pageNumber="8">O</emphasis>
.
<emphasis id="9F84C549A854FFF3D50EFA6B8EA8F9A3" bold="true" box="[878,1005,1520,1538]" italics="true" pageId="7" pageNumber="8">desuntsetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A854FFF3D619F9908DFAF9BC" box="[121,191,1547,1565]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male.
</paragraph>
</caption>
<caption id="F98F49D3A85BFFFCD6FFFA4D8C44F9BC" ID-DOI="http://doi.org/10.5281/zenodo.6284326" ID-Zenodo-Dep="6284326" httpUri="https://zenodo.org/record/6284326/files/figure.png" pageId="8" pageNumber="9" startId="8.[159,196,1494,1512]" targetBox="[159,1021,146,1417]" targetPageId="8">
<paragraph id="AD4F195BA85BFFFCD6FFFA4D8C44F9BC" blockId="8.[159,1044,1494,1565]" pageId="8" pageNumber="9">
Figs 5–8:
<taxonomicName id="6AF062D8A85BFFFCD697FA4D8C35FA49" box="[247,368,1494,1512]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9F84C549A85BFFFCD697FA4D8C07FA49" bold="true" box="[247,322,1494,1512]" italics="true" pageId="8" pageNumber="9">Orictites</emphasis>
spp.
</taxonomicName>
, habitus. 5:
<taxonomicName id="6AF062D8A85BFFFCD7BCFA4D8F24FA49" authorityName="Balkenohl" authorityYear="2017" box="[476,609,1494,1512]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="plurisetosus" status="sp. nov.">
<emphasis id="9F84C549A85BFFFCD7BCFA4D8CACFA49" bold="true" box="[476,489,1494,1512]" italics="true" pageId="8" pageNumber="9">O</emphasis>
.
<emphasis id="9F84C549A85BFFFCD795FA4D8F24FA49" bold="true" box="[501,609,1494,1512]" italics="true" pageId="8" pageNumber="9">plurisetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A85BFFFCD407FA4D8FE9FA49" box="[615,684,1494,1512]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male; 6:
<taxonomicName id="6AF062D8A85BFFFCD53FFA4D8EB1FA49" authorityName="Balkenohl" authorityYear="2017" box="[863,1012,1494,1512]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="anteriortenuis" status="sp. nov.">
<emphasis id="9F84C549A85BFFFCD53FFA4D8E29FA49" bold="true" box="[863,876,1494,1512]" italics="true" pageId="8" pageNumber="9">O</emphasis>
.
<emphasis id="9F84C549A85BFFFCD518FA4D8EB1FA49" bold="true" box="[888,1012,1494,1512]" italics="true" pageId="8" pageNumber="9">anteriortenuis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A85BFFFCD599FA4D8D81F9A3" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male; 7:
<taxonomicName id="6AF062D8A85BFFFCD70FFA6B8F4CF9A3" authorityName="Balkenohl" authorityYear="2017" box="[367,521,1520,1538]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="omnipunctatus" status="sp. nov.">
<emphasis id="9F84C549A85BFFFCD70FFA6B8C39F9A3" bold="true" box="[367,380,1520,1538]" italics="true" pageId="8" pageNumber="9">O</emphasis>
.
<emphasis id="9F84C549A85BFFFCD7E7FA6B8F4CF9A3" bold="true" box="[391,521,1520,1538]" italics="true" pageId="8" pageNumber="9">omnipunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A85BFFFCD46DFA6B8F15F9A3" box="[525,592,1520,1538]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male; 8:
<taxonomicName id="6AF062D8A85BFFFCD49DFA6B8ED6F9A3" authorityName="Balkenohl" authorityYear="2017" box="[765,915,1520,1538]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ampliosetosus" status="sp. nov.">
<emphasis id="9F84C549A85BFFFCD49DFA6B8E4FF9A3" bold="true" box="[765,778,1520,1538]" italics="true" pageId="8" pageNumber="9">O</emphasis>
.
<emphasis id="9F84C549A85BFFFCD574FA6B8ED6F9A3" bold="true" box="[788,915,1520,1538]" italics="true" pageId="8" pageNumber="9">ampliosetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A85BFFFCD5F8FA6B8E9BF9A3" box="[920,990,1520,1538]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male.
</paragraph>
</caption>
<caption id="F98F49D3A85AFFFDD619FA4D8C46F9BC" ID-DOI="http://doi.org/10.5281/zenodo.6284330" ID-Zenodo-Dep="6284330" httpUri="https://zenodo.org/record/6284330/files/figure.png" pageId="9" pageNumber="10" startId="9.[121,158,1494,1512]" targetBox="[145,998,148,1407]" targetPageId="9">
<paragraph id="AD4F195BA85AFFFDD619FA4D8C46F9BC" blockId="9.[121,1007,1494,1565]" pageId="9" pageNumber="10">
Figs 9–12:
<taxonomicName id="6AF062D8A85AFFFDD6B8FA4D8C14FA49" box="[216,337,1494,1512]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9F84C549A85AFFFDD6B8FA4D8C66FA49" bold="true" box="[216,291,1494,1512]" italics="true" pageId="9" pageNumber="10">Orictites</emphasis>
spp.
</taxonomicName>
, habitus. 9:
<taxonomicName id="6AF062D8A85AFFFDD7DCFA4D8F5FFA49" authorityName="Balkenohl" authorityYear="2017" box="[444,538,1494,1512]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="9F84C549A85AFFFDD7DCFA4D8C8CFA49" bold="true" box="[444,457,1494,1512]" italics="true" pageId="9" pageNumber="10">O</emphasis>
.
<emphasis id="9F84C549A85AFFFDD7B4FA4D8F5FFA49" bold="true" box="[468,538,1494,1512]" italics="true" pageId="9" pageNumber="10">barclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A85AFFFDD47FFA4D8F21FA49" box="[543,612,1494,1512]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female; 10:
<taxonomicName id="6AF062D8A85AFFFDD54FFA4D8EDAFA49" authorityName="Balkenohl" authorityYear="2017" box="[815,927,1494,1512]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brancuccii" status="sp. nov.">
<emphasis id="9F84C549A85AFFFDD54FFA4D8E79FA49" bold="true" box="[815,828,1494,1512]" italics="true" pageId="9" pageNumber="10">O</emphasis>
.
<emphasis id="9F84C549A85AFFFDD527FA4D8EDAFA49" bold="true" box="[839,927,1494,1512]" italics="true" pageId="9" pageNumber="10">brancuccii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A85AFFFDD5C4FA4D8EACFA49" box="[932,1001,1494,1512]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male; 11:
<taxonomicName id="6AF062D8A85AFFFDD74EFA6B8CBDF9A3" authorityName="Balkenohl" authorityYear="2017" box="[302,504,1520,1538]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="barclayi" subSpecies="minorfrater">
<emphasis id="9F84C549A85AFFFDD74EFA6B8C7EF9A3" bold="true" box="[302,315,1520,1538]" italics="true" pageId="9" pageNumber="10">O</emphasis>
.
<emphasis id="9F84C549A85AFFFDD727FA6B8CBDF9A3" bold="true" box="[327,504,1520,1538]" italics="true" pageId="9" pageNumber="10">barclayi minorfrater</emphasis>
</taxonomicName>
ssp. nov., holotype, male; 12:
<taxonomicName id="6AF062D8A85AFFFDD571FA6B8EDEF9A3" authorityName="Balkenohl" authorityYear="2017" box="[785,923,1520,1538]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="tubercucollis" status="sp. nov.">
<emphasis id="9F84C549A85AFFFDD571FA6B8E5BF9A3" bold="true" box="[785,798,1520,1538]" italics="true" pageId="9" pageNumber="10">O</emphasis>
.
<emphasis id="9F84C549A85AFFFDD54AFA6B8EDEF9A3" bold="true" box="[810,923,1520,1538]" italics="true" pageId="9" pageNumber="10">tubercucollis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A85AFFFDD5C2FA6B8EAEF9A3" box="[930,1003,1520,1538]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male.
</paragraph>
</caption>
<caption id="F98F49D3A859FFFED6FFFCBA8CD0FCEF" ID-DOI="http://doi.org/10.5281/zenodo.6284334" ID-Zenodo-Dep="6284334" httpUri="https://zenodo.org/record/6284334/files/figure.png" pageId="10" pageNumber="11" startId="10.[159,196,801,819]" targetBox="[159,1036,143,748]" targetPageId="10">
<paragraph id="AD4F195BA859FFFED6FFFCBA8CD0FCEF" blockId="10.[159,1043,801,846]" pageId="10" pageNumber="11">
Figs 13–14:
<taxonomicName id="6AF062D8A859FFFED769FCBA8CC7FC92" box="[265,386,801,819]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9F84C549A859FFFED769FCBA8C11FC92" bold="true" box="[265,340,801,819]" italics="true" pageId="10" pageNumber="11">Orictites</emphasis>
spp.
</taxonomicName>
, habitus. 13:
<taxonomicName id="6AF062D8A859FFFED798FCBA8FCDFC92" authorityName="Balkenohl" authorityYear="2017" box="[504,648,801,819]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="charleshuberi" status="sp. nov.">
<emphasis id="9F84C549A859FFFED798FCBA8F40FC92" bold="true" box="[504,517,801,819]" italics="true" pageId="10" pageNumber="11">O</emphasis>
.
<emphasis id="9F84C549A859FFFED470FCBA8FCDFC92" bold="true" box="[528,648,801,819]" italics="true" pageId="10" pageNumber="11">charleshuberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A859FFFED4EEFCBA8F97FC92" box="[654,722,801,819]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, female; 14:
<taxonomicName id="6AF062D8A859FFFED5FEFCBA8DFFFCEF" authorityName="Balkenohl" authorityYear="2017" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="anteriorlatus" status="sp. nov.">
<emphasis id="9F84C549A859FFFED5FEFCBA8EEEFC92" bold="true" box="[926,939,801,819]" italics="true" pageId="10" pageNumber="11">O</emphasis>
.
<emphasis id="9F84C549A859FFFED5D6FCBA8DFFFCEF" bold="true" italics="true" pageId="10" pageNumber="11">anteriorlatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A859FFFED6DEFCA78C47FCEF" box="[190,258,828,846]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male.
</paragraph>
</caption>
<subSubSection id="E5EA4AD0A859FFF8D6FFFBBE8956FAE4" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="key">
<paragraph id="AD4F195BA859FFFED6FFFBBE8CCEFBE0" blockId="10.[159,395,1061,1089]" box="[159,395,1061,1089]" pageId="10" pageNumber="11">
<heading id="F607AE37A859FFFED6FFFBBE8CCEFBE0" box="[159,395,1061,1089]" centered="true" fontSize="12" level="2" pageId="10" pageNumber="11" reason="3">
<emphasis id="9F84C549A859FFFED6FFFBBE8CCEFBE0" bold="true" box="[159,395,1061,1089]" pageId="10" pageNumber="11">Key to the species</emphasis>
</heading>
</paragraph>
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<keyStep id="160401FEA859FFFED6FFFBE98951F984" pageId="10" pageNumber="11">
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<emphasis id="9F84C549A859FFFED6FFFBE98DE8FB2B" bold="true" box="[159,173,1138,1162]" pageId="10" pageNumber="11">1</emphasis>
Frontal margin of clypeus with four distinctly projecting teeth; clypeal wings tooth-like projecting anteriorly, sharply angled (distinctly less than 90°); mandibles as long as head; dorsal margin of scrobe of both mandibles at base with carina-like tooth; pronotum distinctly broader than long; length
<quantity id="6A08B4BEA859FFFED773FA938CDBFA81" box="[275,414,1288,1312]" metricMagnitude="-2" metricUnit="m" metricValue="1.015" metricValueMax="1.03" metricValueMin="1.0" pageId="10" pageNumber="11" unit="mm" value="10.15" valueMax="10.3" valueMin="10.0">10–10.3 mm</quantity>
............................ Subgenus
<taxonomicName id="6AF062D8A859FFFED48CFA9C8C08FAE4" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9F84C549A859FFFED48CFA9C8E0BFA81" bold="true" box="[748,846,1287,1312]" italics="true" pageId="10" pageNumber="11">Orictites</emphasis>
<bibRefCitation id="C96164AAA859FFFED534FA938956FA81" author="Andrewes, H. E." box="[852,1043,1288,1312]" pageId="10" pageNumber="11" pagination="431 - 485" refId="ref27519" refString="Andrewes, H. E. (1931): On the Carabidae of the Mount Kinabalu. - Journal of the FMS Museum, Kuala Lumpur 16: 431 - 485." type="journal article" year="1931">ANDREWES, 1931</bibRefCitation>
One species
</taxonomicName>
.......................................................
<taxonomicName id="6AF062D8A859FFFED487FAB68956FAE4" ID-CoL="8M924" authority="ANDREWES, 1931" authorityName="ANDREWES" authorityYear="1931" box="[743,1043,1325,1350]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="minotaur">
<emphasis id="9F84C549A859FFFED487FAB68E0BFAE7" bold="true" box="[743,846,1325,1350]" italics="true" pageId="10" pageNumber="11">minotaur</emphasis>
<bibRefCitation id="C96164AAA859FFFED534FAB68956FAE4" author="Andrewes, H. E." box="[852,1043,1325,1349]" pageId="10" pageNumber="11" pagination="431 - 485" refId="ref27519" refString="Andrewes, H. E. (1931): On the Carabidae of the Mount Kinabalu. - Journal of the FMS Museum, Kuala Lumpur 16: 431 - 485." type="journal article" year="1931">ANDREWES, 1931</bibRefCitation>
</taxonomicName>
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</paragraph>
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<emphasis id="9F84C549A859FFFED6FFFAE28DE9FA31" box="[159,172,1401,1424]" italics="true" pageId="10" pageNumber="11">–</emphasis>
Frontal margin of clypeus without teeth; clypeal wings small, obtusely angled or rounded; mandibles distinctly shorter than head; dorsal margin of scrobe of both mandibles at base without elevated tooth; pronotum about as long as wide (ratio length/width 0.9–1.1); length shorter than
<quantity id="6A08B4BEA859FFFED6A5F9968C53F984" box="[197,278,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="10" pageNumber="11" unit="mm" value="7.4">7.4 mm</quantity>
............................................... 2: Subgenus
<taxonomicName id="6AF062D8A859FFFED565F9968E38F987" box="[773,893,1549,1574]" pageId="10" pageNumber="11" rank="subGenus" status="subgen. nov." subGenus="Semictites">
<emphasis id="9F84C549A859FFFED565F9968E38F987" bold="true" box="[773,893,1549,1574]" italics="true" pageId="10" pageNumber="11">Semictites</emphasis>
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<taxonomicNameLabel id="84B77832A859FFFED5E3F9968951F984" box="[899,1044,1549,1573]" pageId="10" pageNumber="11" rank="subGenus">subgen. nov.</taxonomicNameLabel>
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</paragraph>
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<keyStep id="160401FEA858FFFFD619FFC68EA8FF61" pageId="11" pageNumber="12">
<paragraph id="AD4F195BA858FFFFD619FFC68EABFF3B" blockId="11.[121,1006,93,192]" pageId="11" pageNumber="12">
<keyLead id="1601BA6EA858FFFFD619FFC68EABFF3B" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FFC68DC2FFD4" bold="true" box="[121,135,93,117]" pageId="11" pageNumber="12">2</emphasis>
Intervals of elytra without any setigerous punctures; length
<quantity id="6A08B4BEA858FFFFD524FFC68ED9FFD4" box="[836,924,93,117]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="12" unit="mm" value="4.5">4.5 mm</quantity>
. .............................................................................
<taxonomicName id="6AF062D8A858FFFFD48AFF198ED4FF3A" ID-CoL="8MCCG" authorityName="Balkenohl" authorityYear="2017" box="[746,913,130,155]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="desuntsetosus" status="sp. nov.">
<emphasis id="9F84C549A858FFFFD48AFF198ED4FF3A" bold="true" box="[746,913,130,155]" italics="true" pageId="11" pageNumber="12">desuntsetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A858FFFFD5F6FF198EABFF3B" box="[918,1006,130,154]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="AD4F195BA858FFFFD619FF328EA8FF61" blockId="11.[121,1006,93,192]" box="[121,1005,168,192]" pageId="11" pageNumber="12">
<keyLead id="1601BA6EA858FFFFD619FF328EA8FF61" box="[121,1005,168,192]" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FF328DC3FF61" box="[121,134,169,192]" italics="true" pageId="11" pageNumber="12">–</emphasis>
Interval 3, or 3, 5 and 7 of elytra with setigerous punctures ..................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="160401FEA858FFFFD619FF698EABFEDB" pageId="11" pageNumber="12">
<paragraph id="AD4F195BA858FFFFD619FF698EABFE91" blockId="11.[121,1006,242,378]" pageId="11" pageNumber="12">
<keyLead id="1601BA6EA858FFFFD619FF698EABFE91" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FF698DC2FEAB" bold="true" box="[121,135,242,266]" pageId="11" pageNumber="12">3</emphasis>
Intervals 3, 5, and 7 of elytra with series of setigerous punctures; clypeus deeply excised anteriorly; length
<quantity id="6A08B4BEA858FFFFD476FE838F2FFE91" box="[534,618,280,304]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="12" unit="mm" value="3.8">3.8 mm</quantity>
...................
<taxonomicName id="6AF062D8A858FFFFD563FE8C8ED5FE91" ID-CoL="8MBN7" authorityName="Balkenohl" authorityYear="2017" box="[771,912,279,304]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="plurisetosus" status="sp. nov.">
<emphasis id="9F84C549A858FFFFD563FE8C8ED5FE91" bold="true" box="[771,912,279,304]" italics="true" pageId="11" pageNumber="12">plurisetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A858FFFFD5F6FE838EABFE91" box="[918,1006,280,304]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="AD4F195BA858FFFFD619FEA58EABFEDB" blockId="11.[121,1006,242,378]" pageId="11" pageNumber="12">
<keyLead id="1601BA6EA858FFFFD619FEA58EABFEDB" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FEA58DC3FEF4" box="[121,134,318,341]" italics="true" pageId="11" pageNumber="12">–</emphasis>
Setigerous punctures on interval 3 of elytra, only; clypeus straight or slightly excised anteriorly .......................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="160401FEA858FFFFD619FE368EA8FE4B" pageId="11" pageNumber="12">
<paragraph id="AD4F195BA858FFFFD619FE368EA8FE64" blockId="11.[121,1005,429,490]" box="[121,1005,429,453]" pageId="11" pageNumber="12">
<keyLead id="1601BA6EA858FFFFD619FE368EA8FE64" box="[121,1005,429,453]" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FE368DC2FE64" bold="true" box="[121,135,429,453]" pageId="11" pageNumber="12">4</emphasis>
With up to seven setigerous punctures on interval 3 of elytra .................. 5
</keyLead>
</paragraph>
<paragraph id="AD4F195BA858FFFFD619FE488EA8FE4B" blockId="11.[121,1005,429,490]" box="[121,1005,466,490]" pageId="11" pageNumber="12">
<keyLead id="1601BA6EA858FFFFD619FE488EA8FE4B" box="[121,1005,466,490]" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FE488DC3FE4B" box="[121,134,467,490]" italics="true" pageId="11" pageNumber="12">–</emphasis>
With four setigerous punctures on interval 3 of elytra .............................. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="160401FEA858FFFFD619FD868EA8FD6B" pageId="11" pageNumber="12">
<paragraph id="AD4F195BA858FFFFD619FD868EABFD21" blockId="11.[121,1006,541,714]" pageId="11" pageNumber="12">
<keyLead id="1601BA6EA858FFFFD619FD868EABFD21" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FD868DC2FD94" bold="true" box="[121,135,541,565]" pageId="11" pageNumber="12">5</emphasis>
Black; with five setigerous punctures on interval 3 of elytra; knob on clypeus indistinct, developed as elevation; interval 3 and 4 of elytra at base with a tubercle each; length
<quantity id="6A08B4BEA858FFFFD7BEFDF38F76FD21" box="[478,563,616,640]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="11" pageNumber="12" unit="mm" value="4.8">4.8 mm</quantity>
<emphasis id="9F84C549A858FFFFD45FFDF38ED4FD21" box="[575,913,615,640]" italics="true" pageId="11" pageNumber="12">
........................
<taxonomicName id="6AF062D8A858FFFFD48FFDFC8ED4FD21" ID-CoL="8MLMV" authorityName="Balkenohl" authorityYear="2017" box="[751,913,615,640]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="anteriortenuis" status="sp. nov.">
<emphasis id="9F84C549A858FFFFD48FFDFC8ED4FD21" bold="true" box="[751,913,615,640]" italics="true" pageId="11" pageNumber="12">anteriortenuis</emphasis>
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</emphasis>
<taxonomicNameLabel id="84B77832A858FFFFD5F6FDF38EABFD21" box="[918,1006,616,640]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<keyLead id="1601BA6EA858FFFFD619FD158EA8FD6B" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FD158DC3FD04" box="[121,134,654,677]" italics="true" pageId="11" pageNumber="12">–</emphasis>
With more than five setigerous punctures on interval 3 of elytra; brown or black species ............................................................................................. 6
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</paragraph>
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<keyStep id="160401FEA858FFFFD619FD668EA8FBE1" pageId="11" pageNumber="12">
<paragraph id="AD4F195BA858FFFFD619FD668EABFC71" blockId="11.[121,1006,765,1088]" pageId="11" pageNumber="12">
<keyLead id="1601BA6EA858FFFFD619FD668EABFC71" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FD668DC2FCB4" bold="true" box="[121,135,765,789]" pageId="11" pageNumber="12">6</emphasis>
Brown; with up to seven setigerous punctures on interval 3 of elytra (four to seven punctures, variable, see tab. under description); interval 3 and 4 at base with tubercle; knob on clypeus indistinct, developed as elevation; punctures of neck constriction of head visible laterally, no constriction at middle; lateral margin of pronotum straight, anterior angles not projecting; length 3.4–4.0 mm .................................................
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<emphasis id="9F84C549A858FFFFD486FC2C8ED5FC71" bold="true" box="[742,912,951,976]" italics="true" pageId="11" pageNumber="12">omnipunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A858FFFFD5F6FC238EABFC71" box="[918,1006,952,976]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
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<keyLead id="1601BA6EA858FFFFD619FC458EA8FBE1" pageId="11" pageNumber="12">
<emphasis id="9F84C549A858FFFFD619FC458DC3FC54" box="[121,134,990,1013]" italics="true" pageId="11" pageNumber="12">–</emphasis>
Black; with six or seven setigerous punctures on interval 3 of elytra; only interval 3 at base with tubercle; knob on clypeus distinct, neck constriction of head with big punctures; length
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........................................ 7
</keyLead>
</paragraph>
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Lateral margin of pronotum straight, anterior angles distinctly projecting; median line of pronotum conspicuously broad, interval 3 of elytra with six to seven setigerous punctures; length
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<emphasis id="9F84C549A858FFFFD4D0FB268ED4FB77" box="[688,913,1213,1238]" italics="true" pageId="11" pageNumber="12">
.......
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<emphasis id="9F84C549A858FFFFD48AFB268ED4FB77" bold="true" box="[746,913,1213,1238]" italics="true" pageId="11" pageNumber="12">ampliosetosus</emphasis>
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</emphasis>
<taxonomicNameLabel id="84B77832A858FFFFD5F6FB268EABFB74" box="[918,1006,1213,1237]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
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<emphasis id="9F84C549A858FFFFD619FB788DC3FB5B" box="[121,134,1251,1274]" italics="true" pageId="11" pageNumber="12">–</emphasis>
Lateral margin of pronotum convex, anterior angles indistinctly projecting; median line of pronotum moderately narrow ............................................ 8
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</paragraph>
</keyStep>
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<paragraph id="AD4F195BA858FFFFD619FAC98EABFA14" blockId="11.[121,1006,1362,1574]" pageId="11" pageNumber="12">
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<emphasis id="9F84C549A858FFFFD619FAC98DC2FACB" bold="true" box="[121,135,1362,1386]" pageId="11" pageNumber="12">8</emphasis>
Median line of pronotum finely adjoining basal constriction; without central puncture on frons; interval 3 of elytra with seven setigerous punctures; length
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<emphasis id="9F84C549A858FFFFD7F5FA068ED5FA17" box="[405,912,1437,1462]" italics="true" pageId="11" pageNumber="12">
.........................................................
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<emphasis id="9F84C549A858FFFFD555FA068ED5FA17" bold="true" box="[821,912,1437,1462]" italics="true" pageId="11" pageNumber="12">barclayi</emphasis>
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</emphasis>
<taxonomicNameLabel id="84B77832A858FFFFD5F6FA068EABFA14" box="[918,1006,1437,1461]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="9F84C549A858FFFFD619FA588DC3FA7B" box="[121,134,1475,1498]" italics="true" pageId="11" pageNumber="12">–</emphasis>
Median line of pronotum not joining basal constriction; with central puncture on frons; interval 3 of elytra with six setigerous punctures; length
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<emphasis id="9F84C549A858FFFFD767F9968EC1F987" box="[263,900,1549,1574]" italics="true" pageId="11" pageNumber="12">
........................................................
<emphasis id="9F84C549A858FFFFD4C1F9968EC1F987" bold="true" box="[673,900,1549,1574]" italics="true" pageId="11" pageNumber="12">
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minorfrater
</emphasis>
</emphasis>
ssp. nov.
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<emphasis id="9F84C549A85FFFF8D6FFFFC68DE8FFD4" bold="true" box="[159,173,93,117]" pageId="12" pageNumber="13">9</emphasis>
Elytra without tooth at humerus .............................................................. 12
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<emphasis id="9F84C549A85FFFF8D6FFFF188DE9FF3B" box="[159,172,131,154]" italics="true" pageId="12" pageNumber="13">–</emphasis>
Elytra with distinct tooth at humerus ...................................................... 10
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<emphasis id="9F84C549A85FFFF8D6FFFF568DFEFF44" bold="true" box="[159,187,205,229]" pageId="12" pageNumber="13">10</emphasis>
Bigger and more massive species; pronotum distinctly transverse, disc distinctly convex, surface sparsely covered with small punctures; anterior transverse line of pronotum indistinct; preapical spine of mesotibia nearly at apex of tibia, length
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.......................................
<taxonomicName id="6AF062D8A85FFFF8D523FEA68EF3FEF7" ID-CoL="8M9MH" authorityName="Balkenohl" authorityYear="2017" box="[835,950,317,342]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="brancuccii" status="sp. nov.">
<emphasis id="9F84C549A85FFFF8D523FEA68EF3FEF7" bold="true" box="[835,950,317,342]" italics="true" pageId="12" pageNumber="13">brancuccii</emphasis>
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<taxonomicNameLabel id="84B77832A85FFFF8D5DCFEA68951FEF4" box="[956,1044,317,341]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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<emphasis id="9F84C549A85FFFF8D6FFFEF88DE9FEDB" box="[159,172,355,378]" italics="true" pageId="12" pageNumber="13">–</emphasis>
Smaller and slender species; pronotum as long as wide or slightly broader than long, disc somewhat flattened, surface densely covered with big punctures; anterior transverse line of pronotum distinct; preapical spine of mesotibia located distinctly preapically, length up to
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................. 11
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Elytra with tooth at humerus of moderate size, tooth rounded; reflexed margin of pronotum at base tooth-like raised bilaterally; lateral margin of pronotum with few fine scars anteriorly; length
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..............................
<emphasis id="9F84C549A85FFFF8D68DFD168EF3FD07" box="[237,950,653,678]" italics="true" pageId="12" pageNumber="13">
...............................................................................
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<emphasis id="9F84C549A85FFFF8D6FFFD288DE9FD6B" box="[159,172,691,714]" italics="true" pageId="12" pageNumber="13">–</emphasis>
Elytra with tooth at humerus big, sharp; reflexed margin of pronotum at base in total of about same appearance; lateral margin of pronotum crenulated; length
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<emphasis id="9F84C549A85FFFF8D7DAFD668EF3FCB7" box="[442,950,765,790]" italics="true" pageId="12" pageNumber="13">
................................................
<taxonomicName id="6AF062D8A85FFFF8D579FD668EF3FCB7" ID-CoL="8MJW8" authorityName="Balkenohl" authorityYear="2017" box="[793,950,765,790]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="charleshuberi" status="sp. nov.">
<emphasis id="9F84C549A85FFFF8D579FD668EF3FCB7" bold="true" box="[793,950,765,790]" italics="true" pageId="12" pageNumber="13">charleshuberi</emphasis>
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Disc of pronotum with basal part bilaterally elongated posteriorly as toothlike vault slightly hanging over basal channel; lateral margin of pronotum with traces of scars .................................................................................. 13
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<paragraph id="AD4F195BA85FFFF8D6FFFC228956FBBB" blockId="12.[159,1044,839,1051]" pageId="12" pageNumber="13">
<keyLead id="1601BA6EA85FFFF8D6FFFC228956FBBB" pageId="12" pageNumber="13">
<emphasis id="9F84C549A85FFFF8D6FFFC228DE9FC71" box="[159,172,953,976]" italics="true" pageId="12" pageNumber="13">–</emphasis>
Disc of pronotum in basal part regularly developed; lateral margin of pronotum smooth; pronotum as long as wide; length
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.......................... ......................................................................
<taxonomicName id="6AF062D8A85FFFF8D489FB998956FBBB" ID-CoL="8MFK7" authority="BATES, 1892" authorityName="BATES Compared" authorityYear="1892" box="[745,1043,1026,1051]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A85FFFF8D489FB998EC7FBBA" bold="true" box="[745,898,1026,1051]" italics="true" pageId="12" pageNumber="13">costulipennis</emphasis>
<bibRefCitation id="C96164AAA85FFFF8D5E8FB998956FBBB" author="Bates, H. W." box="[904,1043,1026,1050]" pageId="12" pageNumber="13" pagination="267 - 428" refId="ref27786" refString="Bates, H. W. (1892): Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. - Annali del Museo Civico di Storia Naturale di Genova 32: 267 - 428." type="journal article" year="1892">BATES, 1892</bibRefCitation>
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<emphasis id="9F84C549A85FFFF8D6FFFBD68DFEFBC4" bold="true" box="[159,187,1101,1125]" pageId="12" pageNumber="13">13</emphasis>
Clypeus without distinct knob, developed as broad elevation without punctures; intervals 3 and 4 at base of elytra with tubercles; interval 3 of elytra with four setigerous punctures; length
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.................................
<emphasis id="9F84C549A85FFFF8D68DFB268EF3FB77" box="[237,950,1213,1238]" italics="true" pageId="12" pageNumber="13">
..............................................................................
<taxonomicName id="6AF062D8A85FFFF8D542FB268EF3FB77" ID-CoL="8MJVP" authorityName="Balkenohl" authorityYear="2017" box="[802,950,1213,1238]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anteriorlatus" status="sp. nov.">
<emphasis id="9F84C549A85FFFF8D542FB268EF3FB77" bold="true" box="[802,950,1213,1238]" italics="true" pageId="12" pageNumber="13">anteriorlatus</emphasis>
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</emphasis>
<taxonomicNameLabel id="84B77832A85FFFF8D5DCFB268951FB74" box="[956,1044,1213,1237]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="AD4F195BA85FFFF8D6FFFB788956FAE4" blockId="12.[159,1044,1101,1350]" pageId="12" pageNumber="13">
<keyLead id="1601BA6EA85FFFF8D6FFFB788956FAE4" pageId="12" pageNumber="13">
<emphasis id="9F84C549A85FFFF8D6FFFB788DE9FB5B" box="[159,172,1251,1274]" italics="true" pageId="12" pageNumber="13">–</emphasis>
Clypeus with distinct knob, with punctures posterior to knob in form of a V; only interval 3 at base of elytra with tubercle; interval 3 of elytra with four/ five setigerous punctures; length
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<emphasis id="9F84C549A85FFFF8D4FDFAB68F85FAE4" box="[669,704,1325,1349]" italics="true" pageId="12" pageNumber="13">.....</emphasis>
<taxonomicName id="6AF062D8A85FFFF8D4ADFAB68956FAE4" authority="LouWERENS, 1964" authorityName="LouWERENS" authorityYear="1964" box="[717,1043,1325,1350]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A85FFFF8D4ADFAB68E05FAE7" bold="true" box="[717,832,1325,1350]" italics="true" pageId="12" pageNumber="13">mjoebergi</emphasis>
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</taxonomicName>
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321
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<emphasis id="9F84C549A85EFFF9D619FF568C02FF47" bold="true" box="[121,327,205,230]" italics="true" pageId="13" pageNumber="14">Orictites minotaur</emphasis>
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<paragraph id="AD4F195BA85EFFF9D619FE838DA9FE91" blockId="13.[121,716,280,490]" box="[121,236,280,304]" pageId="13" pageNumber="14">Catalogue</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A85EFFF9D619FEA68FF1FE4B" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="AD4F195BA85EFFF9D619FEA68F03FEF4" blockId="13.[121,716,280,490]" box="[121,582,317,341]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="8DE03E75A85EFFF9D619FEA68F03FEF4" box="[121,582,317,341]" pageId="13" pageNumber="14">
<taxonomicName id="6AF062D8A85EFFF9D619FEA68C03FEF4" ID-CoL="8M924" authorityName="ANDREWES" authorityYear="1931" box="[121,326,317,341]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="minotaur">
<emphasis id="9F84C549A85EFFF9D619FEA68C03FEF4" box="[121,326,317,341]" italics="true" pageId="13" pageNumber="14">Orictites minotaur</emphasis>
</taxonomicName>
;
<treatmentCitation id="2C513F4AA85EFFF9D735FEA68F78FEF4" author="Andrewes, H. E." box="[341,573,317,341]" page="440" pageId="13" pageNumber="14" year="1931">
<bibRefCitation id="C96164AAA85EFFF9D735FEA68F78FEF4" author="Andrewes, H. E." box="[341,573,317,341]" pageId="13" pageNumber="14" pagination="431 - 485" refId="ref27519" refString="Andrewes, H. E. (1931): On the Carabidae of the Mount Kinabalu. - Journal of the FMS Museum, Kuala Lumpur 16: 431 - 485." type="journal article" year="1931">Andrewes 1931: 440</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA85EFFF9D619FEF98FCCFEDB" blockId="13.[121,716,280,490]" box="[121,649,354,378]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="8DE03E75A85EFFF9D619FEF98FCCFEDB" box="[121,649,354,378]" pageId="13" pageNumber="14">
<taxonomicName id="6AF062D8A85EFFF9D619FEF98C80FEDB" ID-CoL="8M924" authority="ANDREWES" authorityName="ANDREWES" authorityYear="1931" box="[121,453,354,378]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="minotaur">
<emphasis id="9F84C549A85EFFF9D619FEF98C03FEDB" box="[121,326,354,378]" italics="true" pageId="13" pageNumber="14">Orictites minotaur</emphasis>
ANDREWES
</taxonomicName>
;
<treatmentCitation id="2C513F4AA85EFFF9D7B3FEF98FC4FEDB" author="Csiki E." box="[467,641,354,378]" page="638" pageId="13" pageNumber="14" year="1933">
<bibRefCitation id="C96164AAA85EFFF9D7B3FEF98FC4FEDB" author="Csiki E." box="[467,641,354,378]" pageId="13" pageNumber="14" pagination="623 - 641" refId="ref27828" refString="Csiki E. (1933): Coleopterorum Catalogus auspiciis et auxilio. W. Junk editus a S. Schenkling. 127: Carabidae: Carabinae III, Corrigenda et Addenda. - W. Junk, Berlin: 623 - 641." type="book chapter" year="1933">Csiki 1933: 638</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA85EFFF9D619FE138F89FE01" blockId="13.[121,716,280,490]" box="[121,716,392,416]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="8DE03E75A85EFFF9D619FE138F89FE01" box="[121,716,392,416]" pageId="13" pageNumber="14">
<taxonomicName id="6AF062D8A85EFFF9D619FE138C80FE01" ID-CoL="8M924" authority="ANDREWES" authorityName="ANDREWES" authorityYear="1931" box="[121,453,392,416]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="minotaur">
<emphasis id="9F84C549A85EFFF9D619FE138C03FE01" box="[121,326,392,416]" italics="true" pageId="13" pageNumber="14">Orictites minotaur</emphasis>
ANDREWES
</taxonomicName>
;
<treatmentCitation id="2C513F4AA85EFFF9D7B3FE138F81FE01" author="Louwerens, C. J." box="[467,708,392,416]" page="174" pageId="13" pageNumber="14" year="1964">
<bibRefCitation id="C96164AAA85EFFF9D7B3FE138F81FE01" author="Louwerens, C. J." box="[467,708,392,416]" pageId="13" pageNumber="14" pagination="171 - 189" refId="ref27961" refString="Louwerens, C. J. (1964): An annotated list of the Carabidae, chiefly collected in East Borneo by Dr Eric MjOberg with descriptions of new species. - Entomologisk Tidskrift, Stockholm 85: 171 - 189." type="journal article" year="1964">Louwerens 1964: 174</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA85EFFF9D619FE368FD8FE64" blockId="13.[121,716,280,490]" box="[121,669,429,453]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="8DE03E75A85EFFF9D619FE368FD8FE64" box="[121,669,429,453]" pageId="13" pageNumber="14">
<taxonomicName id="6AF062D8A85EFFF9D619FE368C80FE64" ID-CoL="8M924" authority="ANDREWES" authorityName="ANDREWES" authorityYear="1931" box="[121,453,429,453]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="minotaur">
<emphasis id="9F84C549A85EFFF9D619FE368C03FE64" box="[121,326,429,453]" italics="true" pageId="13" pageNumber="14">Orictites minotaur</emphasis>
ANDREWES
</taxonomicName>
;
<treatmentCitation id="2C513F4AA85EFFF9D7B3FE368FD0FE64" author="Lorenz, W." box="[467,661,429,453]" page="135" pageId="13" pageNumber="14" year="1998">
<bibRefCitation id="C96164AAA85EFFF9D7B3FE368FD0FE64" author="Lorenz, W." box="[467,661,429,453]" pageId="13" pageNumber="14" refId="ref28004" refString="Lorenz, W. (1998): Systematic list of extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodidae). - Tutzing, printed by the author. 502 pp." type="book" year="1998">Lorenz 1998: 135</bibRefCitation>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="AD4F195BA85EFFF9D619FE498FF1FE4B" blockId="13.[121,716,280,490]" box="[121,692,466,490]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="8DE03E75A85EFFF9D619FE498FF1FE4B" box="[121,692,466,490]" pageId="13" pageNumber="14">
<taxonomicName id="6AF062D8A85EFFF9D619FE498C80FE4B" ID-CoL="8M924" authority="ANDREWES" authorityName="ANDREWES" authorityYear="1931" box="[121,453,466,490]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="minotaur">
<emphasis id="9F84C549A85EFFF9D619FE498C03FE4B" box="[121,326,466,490]" italics="true" pageId="13" pageNumber="14">Orictites minotaur</emphasis>
ANDREWES
</taxonomicName>
;
<treatmentCitation id="2C513F4AA85EFFF9D7B3FE498FE8FE4B" author="Balkenohl, M." box="[467,685,466,490]" page="18" pageId="13" pageNumber="14" year="2001">
<bibRefCitation id="C96164AAA85EFFF9D7B3FE498FE8FE4B" author="Balkenohl, M." box="[467,685,466,490]" pageId="13" pageNumber="14" refId="ref27660" refString="Balkenohl, M. (2001): Key and Catalogue of the tribe Clivinini from the Oriental realm, with revisions of the genera Thliboclivina KULT, and Trilophidius JEANNEL (Insecta, Coleoptera, Carabidae, Scarititae, Clivinini). - 83 pp., Sofia, Moscow (Pensoft Series Faunistica 21)." type="book" year="2001">Balkenohl 2001: 18</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A85EFFF9D619FD868CBCFC9B" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="AD4F195BA85EFFF9D619FD868F2DFD51" blockId="13.[121,1007,540,1088]" pageId="13" pageNumber="14">
Type material:
<materialsCitation id="1D981306A85EFFF9D748FD868F2DFD51" ID-GBIF-Occurrence="3497007338" collectingDate="1929-04-12" collectionCode="BMNH" collectorName="H. M. Pendlebury" county="Lumu Lumu" elevation="1701" location="Mt. Kinabalu" municipality="Mt. Kinabalu" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="724BA7F9A85EFFF9D748FD868CDCFD94" box="[296,409,541,565]" pageId="13" pageNumber="14" type="lectotype">Lectotype</typeStatus>
(by present designation):
<specimenCount id="BBF6D2D2A85EFFF9D4B2FD878FA6FD95" box="[722,739,540,564]" pageId="13" pageNumber="14" type="male">♂</specimenCount>
, with labels and data: round, white, with red border, “Type” / white and yellow: “Mt. Kinabalu, Brit.N.Borneo. B.M.1931–386.” / white, handwritten: “
<taxonomicName id="6AF062D8A85EFFF9D570FDF38EA8FD21" authorityName="ANDREWES" authorityYear="1931" box="[784,1005,616,640]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="minotaur">Orictites minotaur</taxonomicName>
Type Andr.” and printed “H.E.
<taxonomicName id="6AF062D8A85EFFF9D7B6FD168F13FD04" box="[470,598,653,677]" pageId="13" pageNumber="1" rank="subGenus" status="subgen. nov." subGenus="Andrewes">Andrewes</taxonomicName>
det.” / light red: “B.N. BORNEO.
<collectingMunicipality id="4D2B8321A85EFFF9D619FD298C54FD6B" box="[121,273,690,714]" pageId="13" pageNumber="14">Mt. Kinabalu</collectingMunicipality>
,
<collectingCounty id="442E61D7A85EFFF9D77DFD298CECFD6B" box="[285,425,690,714]" pageId="13" pageNumber="14">Lumu Lumu</collectingCounty>
,
<quantity id="6A08B4BEA85EFFF9D7D5FD298F54FD6B" box="[437,529,690,714]" metricMagnitude="3" metricUnit="m" metricValue="1.700784" pageId="13" pageNumber="14" unit="ft" value="5580.0">
<elevation id="26DDFE68A85EFFF9D7D5FD298F54FD6B" box="[437,529,690,714]" metricMagnitude="3" metricUnit="m" metricValue="1.700784" pageId="13" pageNumber="14" unit="ft" value="5580.0">5580 ft.</elevation>
</quantity>
<date id="D94E3F9BA85EFFF9D47AFD298F91FD6B" box="[538,724,687,714]" pageId="13" pageNumber="14" value="1929-04-12">
<collectingDate id="C90AC673A85EFFF9D47AFD298F91FD6B" box="[538,724,687,714]" pageId="13" pageNumber="14" value="1929-04-12">
April 12
<superScript id="5A85B413A85EFFF9D416FD348FCCFD63" attach="right" box="[630,649,687,706]" fontSize="8" pageId="13" pageNumber="14">th</superScript>
, 1929
</collectingDate>
</date>
”; with backside: “
<collectorName id="00057C8DA85EFFF9D5CDFD298DBEFD51" pageId="13" pageNumber="14">H. M. Pendlebury</collectorName>
coll. F.M.S. Museums.” (
<collectionCode id="CBE1819EA85EFFF9D46EFD438F1BFD51" box="[526,606,728,752]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="13" pageNumber="14">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="AD4F195BA85EFFF9D619FD668CBCFC9B" blockId="13.[121,1007,540,1088]" pageId="13" pageNumber="14">
Additional material:
<materialsCitation id="1D981306A85EFFF9D709FD668CB0FC9B" ID-GBIF-Occurrence="3497007325" collectingDate="1996-04-03" collectingDateMax="1996-04-20" collectingDateMin="1996-04-03" collectionCode="CBB" collectorName="J. Kadleg" elevation="1700" location="Malaysia-Borneo" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="BBF6D2D2A85EFFF9D709FD668CD0FCB4" box="[361,405,762,789]" pageId="13" pageNumber="14" type="female">1 ♀</specimenCount>
,
<location id="A82F4F80A85EFFF9D7C6FD668F2DFCB4" LSID="urn:lsid:plazi:treatment:2559A84DA85EFFE5D619FF568EC0FB2B:A82F4F80A85EFFF9D7C6FD668F2DFCB4" box="[422,616,765,789]" name="Malaysia-Borneo" pageId="13" pageNumber="14" stateProvince="Sabah">Malaysia-Borneo</location>
<location id="A82F4F80A85EFFF9D412FD668FFFFCB4" LSID="urn:lsid:plazi:treatment:2559A84DA85EFFE5D619FF568EC0FB2B:A82F4F80A85EFFF9D412FD668FFFFCB4" box="[626,698,765,789]" name="Sabah" pageId="13" pageNumber="14" stateProvince="Sabah">Sabah</location>
<collectingDate id="C90AC673A85EFFF9D4A5FD668E39FCB4" box="[709,892,765,789]" pageId="13" pageNumber="14">
<date id="D94E3F9BA85EFFF9D4A5FD668E63FCB4" box="[709,806,765,789]" pageId="13" pageNumber="14" value="2020-03-21">21.3.–20</date>
.4.1996
</collectingDate>
GUNUNG EMAS
<quantity id="6A08B4BEA85EFFF9D6A2FCB98C54FC9B" box="[194,273,802,826]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="m" value="1700.0">
<elevation id="26DDFE68A85EFFF9D6A2FCB98C54FC9B" box="[194,273,802,826]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="m" value="1700.0">1700m</elevation>
</quantity>
lgt.
<collectorName id="00057C8DA85EFFF9D723FCB98CEFFC9B" box="[323,426,802,826]" pageId="13" pageNumber="14">J. Kadleg</collectorName>
, (
<collectionCode id="CBE1819EA85EFFF9D7D8FCB98CAAFC9B" box="[440,495,802,826]" collectionName="CBB" pageId="13" pageNumber="14">CBB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A85EFFF9D6FEFCD38F4DFBE1" pageId="13" pageNumber="14" type="discussion">
<paragraph id="AD4F195BA85EFFF9D6FEFCD38F4DFBE1" blockId="13.[121,1007,540,1088]" pageId="13" pageNumber="14">
Remarks:
<taxonomicName id="6AF062D8A85EFFF9D772FCD38C91FCC1" box="[274,468,840,864]" pageId="13" pageNumber="1" rank="subGenus" status="subgen. nov." subGenus="Andrewes">
<bibRefCitation id="C96164AAA85EFFF9D772FCD38C91FCC1" author="Andrewes, H. E." box="[274,468,840,864]" pageId="13" pageNumber="14" pagination="431 - 485" refId="ref27519" refString="Andrewes, H. E. (1931): On the Carabidae of the Mount Kinabalu. - Journal of the FMS Museum, Kuala Lumpur 16: 431 - 485." type="journal article" year="1931">Andrewes (1931)</bibRefCitation>
</taxonomicName>
based his description on
<specimenCount id="BBF6D2D2A85EFFF9D563FCD38EEAFCC1" box="[771,943,840,864]" pageId="13" pageNumber="14" type="generic">two specimens</specimenCount>
from Lumu Lumu. One of them is deposited in the BMNH and is labelled as “Type”. The specimen was pinned in the middle between the two elytra so that the elytra are opened. It has been mounted on a paper card and the original pin has been used for that card. The second specimen is not anymore in the BMNH, could not be located, and is obviously lost. In the interst of the stability the
<typeStatus id="724BA7F9A85EFFF9D6A0FBB38C6EFBE1" box="[192,299,1064,1088]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
is designated here.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A85EFFE5D619FB0C8C77FD51" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14" type="description">
<paragraph id="AD4F195BA85EFFF9D619FB0C8FB9FB11" blockId="13.[121,764,1175,1200]" box="[121,764,1175,1200]" pageId="13" pageNumber="14">
<emphasis id="9F84C549A85EFFF9D619FB0C8C5DFB11" bold="true" box="[121,280,1175,1200]" pageId="13" pageNumber="14">Redescription</emphasis>
(
<figureCitation id="35CB05DEA85EFFF9D744FB038C2DFB11" box="[292,360,1176,1200]" captionStart="Figs 1–4" captionStartId="7.[121,158,1494,1512]" captionTargetBox="[145,1002,80,1436]" captionTargetPageId="7" captionText="Figs 1–4: Orictites spp., habitus. 1: O. minotaur ANDREWES, 1931, lectotype, male; 2: O. costulipennis BATES, 1892, holotype, male; 3: O. mjoebergi LouWERENS, 1964, lectotype, male; 4: O. desuntsetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284320" httpUri="https://zenodo.org/record/6284320/files/figure.png" pageId="13" pageNumber="14">Figs 1</figureCitation>
,
<figureCitation id="35CB05DEA85EFFF9D715FB038CD4FB11" box="[373,401,1176,1200]" captionStart="Figs 15–16" captionStartId="14.[159,196,1489,1507]" captionTargetBox="[159,1043,104,1429]" captionTargetId="figure-49@14.[159,1043,104,761]" captionTargetPageId="14" captionText="Figs 15–16: Orictites ssp., head and pronotum, lateral view; 15: Orictites (subgenus Orictites) minotaur ANDREWES, 1931,; 16: Orictites (subgenus Semictites) costulipennis BATES, 1892." figureDoi="http://doi.org/10.5281/zenodo.6284338" httpUri="https://zenodo.org/record/6284338/files/figure.png" pageId="13" pageNumber="14">15</figureCitation>
,
<figureCitation id="35CB05DEA85EFFF9D7FEFB038CF2FB11" box="[414,439,1176,1200]" captionStart="Figs 17–18" captionStartId="18.[159,196,780,798]" captionTargetBox="[159,1043,91,754]" captionText="Figs 17–18: Orictites spp., Maxilla, maxillary and labial palpomere. 17: O. minotaur ANDREWES, 1931 (subgenus Orictites); 18: O. costulipennis BATES, 1892 (subgenus Semictites subg. nov.)." figureDoi="http://doi.org/10.5281/zenodo.6284342" httpUri="https://zenodo.org/record/6284342/files/figure.png" pageId="13" pageNumber="14">17</figureCitation>
,
<figureCitation id="35CB05DEA85EFFF9D7A4FB038C9AFB11" box="[452,479,1176,1200]" captionStart="Figs 19–20" captionStartId="22.[159,196,1496,1514]" captionTargetBox="[159,1043,1049,1463]" captionText="Figs 19–20: Orictites spp., aedeagus, ventral view. 19: O. minotaur ANDREWES, 1931, lectotype, scale bar 1.5 mm; 20: O. costulipennis BATES, 1892, holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284346" httpUri="https://zenodo.org/record/6284346/files/figure.png" pageId="13" pageNumber="14">19</figureCitation>
,
<figureCitation id="35CB05DEA85EFFF9D78DFB038F48FB11" box="[493,525,1176,1200]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="13" pageNumber="14">31</figureCitation>
; pp. 8, 15, 19, 23, 35)
</paragraph>
<paragraph id="AD4F195BA85EFFF9D619FB798FBBFA81" blockId="13.[121,1006,1250,1573]" pageId="13" pageNumber="14">
Measurements:
<typeStatus id="724BA7F9A85EFFF9D751FB798CE3FB5B" box="[305,422,1250,1274]" pageId="13" pageNumber="14" type="lectotype">Lectotype</typeStatus>
. Body length
<quantity id="6A08B4BEA85EFFF9D422FB798FE2FB5B" box="[578,679,1250,1274]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="13" pageNumber="14" unit="mm" value="10.1">10.1 mm</quantity>
, width
<quantity id="6A08B4BEA85EFFF9D499FB798E17FB5B" box="[761,850,1250,1274]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="14" unit="mm" value="2.7">2.7 mm</quantity>
; ratio length/ width of pronotum 0.74; ratio length/width of elytra 1.85.
</paragraph>
<paragraph id="AD4F195BA85EFFF9D6FEFAB68E45FACB" blockId="13.[121,1006,1250,1573]" pageId="13" pageNumber="14">
Additional material: Body length
<quantity id="6A08B4BEA85EFFF9D472FAB68F32FAE4" box="[530,631,1325,1349]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="13" pageNumber="14" unit="mm" value="10.3">10.3 mm</quantity>
, width
<quantity id="6A08B4BEA85EFFF9D4A6FAB68E5BFAE4" box="[710,798,1325,1349]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="14" unit="mm" value="2.8">2.8 mm</quantity>
; ratio length/ width of pronotum 0.76; ratio length/width of elytra 1.82.
</paragraph>
<paragraph id="AD4F195BA85EFFF9D6FEFAE38D8DFA7B" blockId="13.[121,1006,1250,1573]" pageId="13" pageNumber="14">Colour: Shiny. Black. Wings of clypeus and supraantennal plates transparent red-brown, antennae, mouthparts, tarsalia medium brown, legs dark redbrown.</paragraph>
<paragraph id="AD4F195BA85EFFFBD6FEFA738E2EFA14" blockId="13.[121,1006,1250,1573]" lastBlockId="15.[121,1006,93,1573]" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
Head: Relatively wide, 15% narrower than pronotum. Clypeus with four anteriorly projecting teeth, with blunt raised knob between the two clypeal setae, transversally hollowed out between anterior margin and knob, wings of clypeus sharply angled (&lt;90°), projecting, divided from supraantennal plates by distinct obtusely angled incision and from middle part of clypeus by broad rounded incision. Supraantennal plates vaulted, with reflexed rounded margin, hollowed out anteriorly; frons separated from clypeus by band of big punctures, with obtuse nearly parallel running carinae at each side posteriorly, carinae with furrows medially and laterally, carinae enclosing the slightly convex frons, frons with irregular big and small punctures, with longitudinal impression at middle. Supraorbital carina flat, with two setae, posterior one arising from big tubercle. Eyes not big, projecting, tempora carinate, genae distinctly vaulted, nearly as high and as long as eyes, enclosing eyes posteriorly, forming distinct angle at neck. Eyes and tempora separated from middle part of head by distinct supraorbital carinae and deep broad furrows. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction formed by doubled irregular row of big punctures, neck laterally densely covered with medium sized punctures. Labrum formed by two broad lobes, with slight emargination at middle, 7-setose (in the
<typeStatus id="724BA7F9A85CFFFBD4F0FD298FBEFD6B" box="[656,763,690,714]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
one seta is missing), ciliate laterally, with isodiametric reticulation. Mandibles robust, nearly as long as head, relatively slender, acute at apex, upper margin of scrobe of both mandibles at base with carina-like tooth, right one with distinct tooth at middle, left one with knob-like tooth at base visible in opened position, both mandibles hollowed out at base; scrobe rounded dorsally and ventrally in apical half, with no carinae, appearing rounded at sides. Maxilla bent sigmoidally, acutely hooked at apex. Apical maxillary palpomere slender, straight. Apical labial palpomere slender, straight, more than twice as long as 2
<superScript id="5A85B413A85CFFFBD503FC418E3FFC4C" attach="left" box="[867,890,986,1005]" fontSize="8" pageId="15" pageNumber="16">nd</superScript>
segment, 2
<superScript id="5A85B413A85CFFFBD6E7FC648DDBFBB3" attach="left" box="[135,158,1023,1042]" fontSize="8" pageId="15" pageNumber="16">nd</superScript>
segment bisetose. Ligula bilobed at apex, with long seta; paraglossae slender. Ventral surface of neck covered with big punctures, with transverse wrinkles near gula. Gula smooth. Submentum and mentum distinctly separated; submentum with two pairs of setae; mentum keeled at middle, nearly smooth, with fine and complete reflexed margin, lateral lobes projecting, thought obtusely angled at tips, with two setae near base of each lobe, median tooth forming broad obtuse triangle, bisetose, projecting as far as lateral lobes. Paragena sharp, carina-like, without tooth. Antennae short, reaching up to middle of pronotum, scapus carinate, with one seta dorsoapically, scapus and pedicellus longitudinally reticulated, segments 5–10 broader than long (L/
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), antennomeres flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with some additional longer setae.
</paragraph>
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<paragraph id="AD4F195BA85DFFFAD6FFFA4A8EFDFA5C" blockId="14.[159,1006,1489,1533]" pageId="14" pageNumber="15">
Figs 15–16:
<taxonomicName id="6AF062D8A85DFFFAD76DFA4A8C1DFA42" authorityName="ANDREWES" authorityYear="1931" box="[269,344,1489,1507]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="9F84C549A85DFFFAD76DFA4A8C1DFA42" bold="true" box="[269,344,1489,1507]" italics="true" pageId="14" pageNumber="15">Orictites</emphasis>
</taxonomicName>
ssp., head and pronotum, lateral view; 15:
<taxonomicName id="6AF062D8A85DFFFAD486FA4A8E74FA42" authorityName="ANDREWES" authorityYear="1931" box="[742,817,1489,1507]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="9F84C549A85DFFFAD486FA4A8E74FA42" bold="true" box="[742,817,1489,1507]" italics="true" pageId="14" pageNumber="15">Orictites</emphasis>
</taxonomicName>
(subgenus
<taxonomicName id="6AF062D8A85DFFFAD5FCFA4A8EA2FA42" authorityName="ANDREWES" authorityYear="1931" box="[924,999,1489,1507]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="9F84C549A85DFFFAD5FCFA4A8EA2FA42" bold="true" box="[924,999,1489,1507]" italics="true" pageId="14" pageNumber="15">Orictites</emphasis>
</taxonomicName>
)
<taxonomicName id="6AF062D8A85DFFFAD6FFFA708CC6FA5C" authority="ANDREWES, 1931" authorityName="ANDREWES" authorityYear="1931" box="[159,387,1515,1533]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="minotaur">
<emphasis id="9F84C549A85DFFFAD6FFFA708DA8FA5C" bold="true" box="[159,237,1515,1533]" italics="true" pageId="14" pageNumber="15">minotaur</emphasis>
<bibRefCitation id="C96164AAA85DFFFAD692FA708CC6FA5C" author="Andrewes, H. E." box="[242,387,1515,1533]" pageId="14" pageNumber="15" pagination="431 - 485" refId="ref27519" refString="Andrewes, H. E. (1931): On the Carabidae of the Mount Kinabalu. - Journal of the FMS Museum, Kuala Lumpur 16: 431 - 485." type="journal article" year="1931">ANDREWES, 1931</bibRefCitation>
</taxonomicName>
,; 16:
<taxonomicName id="6AF062D8A85DFFFAD7D8FA708F46FA5C" authorityName="ANDREWES" authorityYear="1931" box="[440,515,1515,1533]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="9F84C549A85DFFFAD7D8FA708F46FA5C" bold="true" box="[440,515,1515,1533]" italics="true" pageId="14" pageNumber="15">Orictites</emphasis>
</taxonomicName>
(subgenus
<taxonomicName id="6AF062D8A85DFFFAD407FA708F87FA5C" box="[615,706,1515,1533]" pageId="14" pageNumber="15" rank="subGenus" subGenus="Semictites">
<emphasis id="9F84C549A85DFFFAD407FA708F87FA5C" bold="true" box="[615,706,1515,1533]" italics="true" pageId="14" pageNumber="15">Semictites</emphasis>
</taxonomicName>
)
<taxonomicName id="6AF062D8A85DFFFAD4AEFA708EF7FA5C" authority="BATES, 1892" authorityName="BATES Compared" authorityYear="1892" box="[718,946,1515,1533]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="costulipennis">
<emphasis id="9F84C549A85DFFFAD4AEFA708E06FA5C" bold="true" box="[718,835,1515,1533]" italics="true" pageId="14" pageNumber="15">costulipennis</emphasis>
<bibRefCitation id="C96164AAA85DFFFAD528FA708EF7FA5C" author="Bates, H. W." box="[840,946,1515,1533]" pageId="14" pageNumber="15" pagination="267 - 428" refId="ref27786" refString="Bates, H. W. (1892): Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. - Annali del Museo Civico di Storia Naturale di Genova 32: 267 - 428." type="journal article" year="1892">BATES, 1892</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="AD4F195BA85CFFE4D6FEFA598F97FD21" blockId="15.[121,1006,93,1573]" lastBlockId="16.[159,1044,93,1573]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">Pronotum: Subquadratic, broader than long. Sides parallel at middle but convex slightly before posterior angles. Anterior margin slightly bisinuate. Reflexed lateral margin distinctly crenulated up to posterior angle, smooth up to base, running from posterior angle to base as sigmoid curve. Lateral channel broad, subfoveolate between the two lateral setigerous punctures, paralateral part of disc slightly vaulted, basal channel broad and deep. Anterior setigerous puncture at level of anterior transverse line, the posterior one located before level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles distinctly projecting, rounded, formed by reflexed lateral margin, posterior angles sharp, not projecting laterally due to convexity of lateral margin in basal part. Anterior transverse line sharp, not reaching lateral channel. Median line deep and conspicuously broad, with blunt carinae bilaterally, surpassing level of anterior transverse line but not joining, running nearly up to channel of base, not adjoining base. Surface with scattered big punctures and some irregular big wrinkles, with fine isodiametric reticulation towards base, with longitudinal vault parallel to lateral channel. Basal part of disc elongated posteriorly and bilaterally, hanging over basal channel (lateral view). Flange at base sharply raised (lateral view).</paragraph>
<paragraph id="AD4F195BA843FFE4D6A4FD168E4CFBC4" blockId="16.[159,1044,93,1573]" pageId="16" pageNumber="17">
Elytra: Elongate, sides parallel at middle. Marginal channel broad, completely visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, no fold-like apical carina visible. Reflexed margin with fine crenulation up to apex, distinct at humerus, margin and channel bending over rounded humerus up to 5
<superScript id="5A85B413A843FFE4D408FC848F3EFC93" attach="left" box="[616,635,799,818]" fontSize="8" pageId="16" pageNumber="17">th</superScript>
stria, with rounded humeral tooth. Basal tubercle big, with setigerous puncture, situated at declivity of 1
<superScript id="5A85B413A843FFE4D5A7FCDF8E9DFCF6" attach="left" box="[967,984,836,855]" fontSize="8" pageId="16" pageNumber="17">st</superScript>
stria 3. Striae 1–4 free at base, 7
<superScript id="5A85B413A843FFE4D786FCF18CBCFCDC" attach="left" box="[486,505,874,893]" fontSize="8" pageId="16" pageNumber="17">th</superScript>
shortened at humerus and nearly joining with stria 6, all striae deep, with indistinct row of punctures, striae 1 and 2 ending free at apex, 3–4 and 5–6 joined at apex. Striole distinct, deepened at base, with small sharp tubercle at base. Intervals 1–4 moderately convex, others distinctly convex, all flattened towards apex, 3
<superScript id="5A85B413A843FFE4D4D6FC648F8CFBB3" attach="left" box="[694,713,1023,1042]" fontSize="8" pageId="16" pageNumber="17">rd</superScript>
at base broader, raised, and with longitudinal tubercle, 8
<superScript id="5A85B413A843FFE4D784FBBF8CB2FB96" attach="left" box="[484,503,1060,1079]" fontSize="8" pageId="16" pageNumber="17">th</superScript>
small, carinate at base and apically. Interval 3 with five robust setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA843FFE4D6A4FBE98D93FA14" blockId="16.[159,1044,93,1573]" pageId="16" pageNumber="17">
Hind wings: Fully developed. Ventral surface: Proepisternum with submarginal ridge. Proepipleuron small, nearly smooth, submarginal furrow distinct. Epipleuron broadened in basal third, with row of partly connected big punctures in basal quarter. Proepisternum, prosternite, episternum, and lateral parts of mesosternum covered densely and irregularly with big punctures, mesosternum smooth at middle. Sternites smooth, 3
<superScript id="5A85B413A843FFE4D49CFAB18E4AFA9C" attach="left" box="[764,783,1322,1341]" fontSize="8" pageId="16" pageNumber="17">rd</superScript>
to 5
<superScript id="5A85B413A843FFE4D524FAB18E12FA9C" attach="left" box="[836,855,1322,1341]" fontSize="8" pageId="16" pageNumber="17">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite six with flat transverse depression apically, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA843FFE5D6A4FA598CFDFF44" blockId="16.[159,1044,93,1573]" lastBlockId="17.[121,1006,93,1162]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">Legs: Protibia with slight dorsal furrow at base, digitation distinct, with two big and one small lateral preapical denticles, all furnished with seta close to apex, apical spine curved hook-like ventrally and laterally, movable spur moderately curved, inner denticle with corkscrew-like seta; basal tarsomere elongated, longer than the following three together. Mesotibia with big long apical spine, furnished with one seta apically and another one at middle. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA842FFE5D6FEFF698F50FE4B" blockId="17.[121,1006,93,1162]" pageId="17" pageNumber="18">
Male genitalia (
<figureCitation id="35CB05DEA842FFE5D72DFF698CDFFEAB" box="[333,410,242,266]" captionStart="Figs 19–20" captionStartId="22.[159,196,1496,1514]" captionTargetBox="[159,1043,1049,1463]" captionText="Figs 19–20: Orictites spp., aedeagus, ventral view. 19: O. minotaur ANDREWES, 1931, lectotype, scale bar 1.5 mm; 20: O. costulipennis BATES, 1892, holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284346" httpUri="https://zenodo.org/record/6284346/files/figure.png" pageId="17" pageNumber="18">Fig. 19</figureCitation>
): Median lobe sclerotized, regularly and moderately arcuate in middle part, distinctly bent at apex, apex formed as slightly asymmetric rounded big spatula. Oroficium large, closing lips less sclerotized. No spines or teeth visible on endophallus (at 500x by optimized condenser and different filters). Parameres asymmetrical, slender, somewhat twisted, ventral one with four medium sized setae at apex, dorsal one with three medium sized and one very fine seta at apex.
</paragraph>
<paragraph id="AD4F195BA842FFE5D6FEFE638FCEFDFB" blockId="17.[121,1006,93,1162]" pageId="17" pageNumber="18">
Female genitalia (
<figureCitation id="35CB05DEA842FFE5D707FE638CF1FDB1" box="[359,436,504,528]" captionStart="Figs 31–34" captionStartId="34.[159,196,1291,1309]" captionTargetBox="[159,1043,80,1253]" captionText="Figs 31–34: Orictites spp., coxostylus. 31: O. minotaur ANDREWES, 1931, scale bar 0.5 mm; 32: O. costulipennis BATES, 1892, scale bar 0.1 mm. 33: O. mjoebergi LouWERENS, 1964, scale bar 0.1 mm; 34: O. desuntsetosus sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284358" httpUri="https://zenodo.org/record/6284358/files/figure.png" pageId="17" pageNumber="18">Fig. 31</figureCitation>
): Coxostylus sigmoid, dorso-ventrally moderately depressed, gently curved to apex, with one seta at apex, with one longer ensiform and four nematiform setae in middle part.
</paragraph>
<paragraph id="AD4F195BA842FFE5D6FEFDF38C77FD51" blockId="17.[121,1006,93,1162]" pageId="17" pageNumber="18">
Variation: The labrum is 6-setose in the
<typeStatus id="724BA7F9A842FFE5D403FDF38F8BFD21" box="[611,718,616,640]" pageId="17" pageNumber="18" type="lectotype">lectotype</typeStatus>
and 7-setose in the additional specimen. In the additional specimen one of the ambulatory setae of the sternite is missing. The anterior transverse line of the pronotum is sharper in the
<typeStatus id="724BA7F9A842FFE5D6DFFD438C6BFD51" box="[191,302,728,752]" pageId="17" pageNumber="18" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A842FFE5D6FEFD668DA8FBE1" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="AD4F195BA842FFE5D6FEFD668DA8FBE1" blockId="17.[121,1006,93,1162]" pageId="17" pageNumber="18">Diagnosis: A big black species with distinct knob on the clypeus and flat broad transverse impression with band of big punctures behind knob, with rounded humeral tooth, tubercle at the base of interval 3 of the elytra, and five setigerous punctures on interval 3. Moreover, the pronotum is distinctly broader than long, and the margin of pronotum and elytra is crenulated. Distinguished from all other species by the long and slender mandibles with a scrobe that is not carinate in the apical half, and they each exhibit an elevated tooth near the base. In addition, the clypeus possesses four projecting teeth anteriorly.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A842FFE5D6FEFBD68EC0FB2B" pageId="17" pageNumber="18" type="distribution">
<paragraph id="AD4F195BA842FFE5D6FEFBD68EC0FB2B" blockId="17.[121,1006,93,1162]" pageId="17" pageNumber="18">
Distribution: Known from the North of Borneo only. The
<typeStatus id="724BA7F9A842FFE5D579FBD68E09FBC4" box="[793,844,1101,1125]" pageId="17" pageNumber="18">type</typeStatus>
was collected at
<quantity id="6A08B4BEA842FFE5D6F5FBE98DA3FB2B" box="[149,230,1138,1162]" metricMagnitude="3" metricUnit="m" metricValue="1.700784" pageId="17" pageNumber="18" unit="ft" value="5580.0">5580 ft</quantity>
(
<quantity id="6A08B4BEA842FFE5D694FBE98C0CFB2B" box="[244,329,1138,1162]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="18" unit="m" value="1700.0">1700 m</quantity>
) and the additional specimen at the same altitude.
</paragraph>
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</treatment>
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271
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<taxonomicName id="6AF062D8A862FFC5D619FA068C16FA17" ID-CoL="8M9MH" authorityName="Balkenohl" authorityYear="2017" box="[121,339,1437,1462]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="brancuccii" status="sp. nov.">
<emphasis id="9F84C549A862FFC5D619FA068C16FA17" bold="true" box="[121,339,1437,1462]" italics="true" pageId="49" pageNumber="50">Orictites brancuccii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A862FFC5D73AFA068CF7FA14" box="[346,434,1437,1461]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A862FFC6D619FA738FCDFE01" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="AD4F195BA862FFC6D619FA738F15FF61" blockId="49.[121,1005,1512,1573]" lastBlockId="50.[159,1044,93,490]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
Type material:
<materialsCitation id="1D981306A862FFC6D74CFA738F15FF61" ID-GBIF-Occurrence="3497007330" collectingDate="2009-05-10" collectingDateMax="2009-05-30" collectingDateMin="2009-05-10" collectionCode="NHMB" collectorName="Z. Kraus &amp; M. Brancucci &amp; M. Geiser &amp; D. Hauck &amp; V. Kuban" country="&quot; LAOS-NE" elevation="1450" lastPageId="50" lastPageNumber="51" latitude="103.333336" location="Phonsavan" municipality="Xieng Khouang" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Xiangkhouang" typeStatus="holotype">
<typeStatus id="724BA7F9A862FFC5D74CFA738CDFF9A1" box="[300,410,1512,1536]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A862FFC5D7C8FA738CF9FA5E" box="[424,444,1512,1535]" pageId="49" pageNumber="50" type="male">♂</specimenCount>
, labels: white, black printed:
<collectingCountry id="D5E759CBA862FFC5D547FA738EDAF9A1" box="[807,927,1512,1536]" pageId="49" pageNumber="50">“LAOS-NE</collectingCountry>
,
<collectingMunicipality id="4D2B8321A862FFC5D5CDFA738D9BF984" country="&quot; LAOS-NE" latitude="103.333336" municipality="Phou Sane Mt." name="Xieng Khouang" pageId="49" pageNumber="50" stateProvince="Xiangkhouang">Xieng Khouang</collectingMunicipality>
prov.,~ 19°37–8’N 103°20´E,
<location id="A82F4F80A862FFC5D42AF9968F8DF984" LSID="urn:lsid:plazi:treatment:2559A84DA862FFC0D619FA068CCAFB11:A82F4F80A862FFC5D42AF9968F8DF984" box="[586,712,1549,1573]" country="&quot; LAOS-NE" latitude="103.333336" municipality="Xieng Khouang" name="Phonsavan" pageId="49" pageNumber="50" stateProvince="Xiangkhouang">Phonsavan</location>
(
<locationDeviation id="BCFD0079A862FFC5D4BCF9968E1DF984" bearing="45.0" box="[732,856,1549,1573]" distance="30000.0" pageId="49" pageNumber="50">
<quantity id="6A08B4BEA862FFC5D4BCF9968E62F984" box="[732,807,1549,1573]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="49" pageNumber="50" unit="km" value="30.0">30 km</quantity>
NE
</locationDeviation>
):
<location id="A82F4F80A862FFC6D50EF9968D8DFFD4" LSID="urn:lsid:plazi:treatment:2559A84DA862FFC0D619FA068CCAFB11:A82F4F80A862FFC6D50EF9968D8DFFD4" country="&quot; LAOS-NE" lastPageId="50" lastPageNumber="51" latitude="103.333336" municipality="Xieng Khouang" name="Phou Sane Mt." pageId="49" pageNumber="50" stateProvince="Xiangkhouang">Phou Sane Mt.</location>
,~
<quantity id="6A08B4BEA861FFC6D6BBFFC68CC0FFD4" box="[219,389,93,117]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="50" pageNumber="51" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">
<elevation id="26DDFE68A861FFC6D6BBFFC68CC0FFD4" box="[219,389,93,117]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="50" pageNumber="51" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400–1500 m</elevation>
</quantity>
,
<date id="D94E3F9BA861FFC6D7F5FFC68F72FFD4" box="[405,567,93,117]" pageId="50" pageNumber="51" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<collectingDate id="C90AC673A861FFC6D7F5FFC68F72FFD4" box="[405,567,93,117]" pageId="50" pageNumber="51" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">10.–30.v.2009</collectingDate>
</date>
,
<collectorName id="00057C8DA861FFC6D42BFFC68FEAFFD4" box="[587,687,93,117]" pageId="50" pageNumber="51">Z. Kraus</collectorName>
leg.” / “NHMB
<location id="A82F4F80A861FFC6D519FFC68EFBFFD4" LSID="urn:lsid:plazi:treatment:2559A84DA862FFC0D619FA068CCAFB11:A82F4F80A861FFC6D519FFC68EFBFFD4" box="[889,958,93,117]" country="&quot; LAOS-NE" latitude="103.333336" municipality="Xieng Khouang" name="Basel" pageId="50" pageNumber="51" stateProvince="Xiangkhouang">Basel</location>
, NMPC
<location id="A82F4F80A861FFC6D6FFFF198C69FF3B" LSID="urn:lsid:plazi:treatment:2559A84DA862FFC0D619FA068CCAFB11:A82F4F80A861FFC6D6FFFF198C69FF3B" box="[159,300,130,154]" country="&quot; LAOS-NE" latitude="103.333336" municipality="Xieng Khouang" name="Prague Laos" pageId="50" pageNumber="51" stateProvince="Xiangkhouang">Prague Laos</location>
2009
<location id="A82F4F80A861FFC6D716FF198CB0FF3B" LSID="urn:lsid:plazi:treatment:2559A84DA862FFC0D619FA068CCAFB11:A82F4F80A861FFC6D716FF198CB0FF3B" box="[374,501,130,154]" country="&quot; LAOS-NE" latitude="103.333336" municipality="Xieng Khouang" name="Expedition" pageId="50" pageNumber="51" stateProvince="Xiangkhouang">Expedition</location>
:
<collectorName id="00057C8DA861FFC6D79FFF198FD3FF3B" box="[511,662,130,154]" pageId="50" pageNumber="51">M. Brancucci</collectorName>
,
<collectorName id="00057C8DA861FFC6D4C1FF198E54FF3B" box="[673,785,130,154]" pageId="50" pageNumber="51">M. Geiser</collectorName>
,
<collectorName id="00057C8DA861FFC6D57CFF198E3AFF3B" box="[796,895,130,154]" pageId="50" pageNumber="51">Z. Kraus</collectorName>
,
<collectorName id="00057C8DA861FFC6D5E9FF198EB6FF3B" box="[905,1011,130,154]" pageId="50" pageNumber="51">D. Hauck</collectorName>
,
<collectorName id="00057C8DA861FFC6D59EFF198DACFF60" pageId="50" pageNumber="51">V. Kubáň</collectorName>
” / “COLL: NHM Basel” (
<collectionCode id="CBE1819EA861FFC6D796FF338F03FF61" box="[502,582,168,192]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="50" pageNumber="51">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="AD4F195BA861FFC6D6FFFF568FCDFE01" blockId="50.[159,1044,93,490]" pageId="50" pageNumber="51">
<materialsCitation id="1D981306A861FFC6D6FFFF568F09FEF4" ID-GBIF-Occurrence="3497007339" collectingDate="2012-05-25" collectingDateMax="2012-05-31" collectingDateMin="2012-05-25" collectionCode="NHMB, COLL, CBB" collectorName="M. Brancucci &amp; M. Geiser &amp; K. Phanthavong &amp; S. Xayalath" country="Laos" elevation="700" latitude="16.133333" location="Xe Xap NPA" pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="Salavan prov." typeStatus="paratype">
<typeStatus id="724BA7F9A861FFC6D6FFFF568C5CFF44" box="[159,281,205,229]" pageId="50" pageNumber="51" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="BBF6D2D2A861FFC6D747FF568C11FF44" box="[295,340,205,229]" pageId="50" pageNumber="51" type="male">1 ♂</specimenCount>
: “
<collectingCountry id="D5E759CBA861FFC6D70FFF568CF3FF44" box="[367,438,205,229]" name="Laos" pageId="50" pageNumber="51">LAOS</collectingCountry>
,
<collectingRegion id="6F34D7B9A861FFC6D7A5FF568F2FFF44" box="[453,618,205,229]" country="Laos" name="Salavan" pageId="50" pageNumber="51">Salavan prov.</collectingRegion>
,
<location id="A82F4F80A861FFC6D418FF568E4EFF44" LSID="urn:lsid:plazi:treatment:2559A84DA862FFC0D619FA068CCAFB11:A82F4F80A861FFC6D418FF568E4EFF44" box="[632,779,205,229]" country="Laos" latitude="16.133333" name="Xe Xap NPA" pageId="50" pageNumber="51" stateProvince="Salavan prov.">Xe Xap NPA</location>
,
<locationDeviation id="BCFD0079A861FFC6D579FF568E91FF44" box="[793,980,205,229]" pageId="50" pageNumber="51">
<quantity id="6A08B4BEA861FFC6D579FF568E1DFF44" box="[793,856,205,229]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="50" pageNumber="51" unit="km" value="15.0">15km</quantity>
NE Ta-Oy
</locationDeviation>
,
<location id="A82F4F80A861FFC6D582FF568C67FEAB" LSID="urn:lsid:plazi:treatment:2559A84DA862FFC0D619FA068CCAFB11:A82F4F80A861FFC6D582FF568C67FEAB" country="Laos" latitude="16.133333" name="BAN DOUB env." pageId="50" pageNumber="51" stateProvince="Salavan prov.">BAN DOUB env.</location>
,
<quantity id="6A08B4BEA861FFC6D750FF698C8CFEAB" box="[304,457,242,266]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="50" pageNumber="51" unit="m" value="700.0" valueMax="1000.0" valueMin="400.0">
<elevation id="26DDFE68A861FFC6D750FF698C8CFEAB" box="[304,457,242,266]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="50" pageNumber="51" unit="m" value="700.0" valueMax="1000.0" valueMin="400.0">400–1000 m</elevation>
</quantity>
, 16°8´N / 106°40–43´E,
<date id="D94E3F9BA861FFC6D491FF698EECFEAB" box="[753,937,242,266]" pageId="50" pageNumber="51" value="2012-05-25" valueMax="2012-05-31" valueMin="2012-05-25">
<collectingDate id="C90AC673A861FFC6D491FF698EECFEAB" box="[753,937,242,266]" pageId="50" pageNumber="51" value="2012-05-25" valueMax="2012-05-31" valueMin="2012-05-25">25.–31. V. 2012</collectingDate>
</date>
/
<collectionCode id="CBE1819EA861FFC6D5ABFF698956FEAB" box="[971,1043,242,266]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="50" pageNumber="51">NHMB</collectionCode>
Basel Laos 2012 Expedition,
<collectorName id="00057C8DA861FFC6D798FE838FD6FE91" box="[504,659,280,304]" pageId="50" pageNumber="51">M. Brancucci</collectorName>
,
<collectorName id="00057C8DA861FFC6D4FFFE838E57FE91" box="[671,786,280,304]" pageId="50" pageNumber="51">M. Geiser</collectorName>
,
<collectorName id="00057C8DA861FFC6D540FE838E93FE91" box="[800,982,280,304]" pageId="50" pageNumber="51">K. Phanthavong</collectorName>
&amp;
<collectorName id="00057C8DA861FFC6D59DFE838C46FEF4" pageId="50" pageNumber="51">S. Xayalath</collectorName>
/
<collectionCode id="CBE1819EA861FFC6D743FEA68C26FEF4" box="[291,355,317,341]" pageId="50" pageNumber="51">COLL</collectionCode>
: NHM Basel“ (
<collectionCode id="CBE1819EA861FFC6D46EFEA68F03FEF4" box="[526,582,317,341]" pageId="50" pageNumber="51">CBB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A861FFC6D43AFEA68FC1FE01" ID-GBIF-Occurrence="3497007302" collectingDate="2000-11-25" collectingDateMax="2000-11-30" collectingDateMin="2000-11-25" collectionCode="CBP" collectorName="E. Jendek &amp; P. Pacholatko" country="Laos" elevation="250" latitude="18.116667" location="Khammouan prov." longLatPrecision="0" longitude="104.0" pageId="50" pageNumber="51" specimenCount="1" specimenCount-male="1" stateProvince="Khammouan prov." typeStatus="paratype">
<specimenCount id="BBF6D2D2A861FFC6D43AFEA68FC2FEF4" box="[602,647,317,341]" pageId="50" pageNumber="51" type="male">1 ♂</specimenCount>
: “
<collectingCountry id="D5E759CBA861FFC6D4C1FEA68FA1FEF4" box="[673,740,317,341]" name="Laos" pageId="50" pageNumber="51">LAOS</collectingCountry>
centr.,
<collectingRegion id="6F34D7B9A861FFC6D520FEA68951FEF4" box="[832,1044,317,341]" country="Laos" name="Khammouan" pageId="50" pageNumber="51">Khammouan prov.</collectingRegion>
4–16.XI.,
<date id="D94E3F9BA861FFC6D76FFEF98C87FEDB" box="[271,450,354,378]" pageId="50" pageNumber="51" value="2000-11-25" valueMax="2000-11-30" valueMin="2000-11-25">
<collectingDate id="C90AC673A861FFC6D76FFEF98C87FEDB" box="[271,450,354,378]" pageId="50" pageNumber="51" value="2000-11-25" valueMax="2000-11-30" valueMin="2000-11-25">25–30.XI. 2000</collectingDate>
</date>
BAN KHOUN NGEUN env. N18°07´, E 104°29´alt.
<quantity id="6A08B4BEA861FFC6D6FFFE138D9AFE01" box="[159,223,392,416]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="50" pageNumber="51" unit="m" value="250.0">
<elevation id="26DDFE68A861FFC6D6FFFE138D9AFE01" box="[159,223,392,416]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="50" pageNumber="51" unit="m" value="250.0">250m</elevation>
</quantity>
<collectorName id="00057C8DA861FFC6D685FE138C08FE01" box="[229,333,392,416]" pageId="50" pageNumber="51">E. Jendek</collectorName>
&amp;
<collectorName id="00057C8DA861FFC6D70BFE138F47FE01" box="[363,514,392,416]" pageId="50" pageNumber="51">P. Pacholátko</collectorName>
leg.“ (
<collectionCode id="CBE1819EA861FFC6D429FE138F3AFE01" box="[585,639,392,416]" pageId="50" pageNumber="51">CBP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A861FFC6D6A4FE368F4CFE4B" pageId="50" pageNumber="51" type="discussion">
<paragraph id="AD4F195BA861FFC6D6A4FE368F4CFE4B" blockId="50.[159,1044,93,490]" pageId="50" pageNumber="51">
Remarks: In the
<typeStatus id="724BA7F9A861FFC6D7E2FE368CADFE64" box="[386,488,429,453]" pageId="50" pageNumber="51" type="paratype">paratype</typeStatus>
some antennomeres are missing as well as two tarsomeres of the right hind leg.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A861FFC0D6FFFDD98F30FCB4" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51" type="description">
<paragraph id="AD4F195BA861FFC6D6FFFDD98F6BFDFB" blockId="50.[159,558,578,603]" box="[159,558,578,603]" pageId="50" pageNumber="51">
<emphasis id="9F84C549A861FFC6D6FFFDD98C67FDFA" bold="true" box="[159,290,578,603]" pageId="50" pageNumber="51">Description</emphasis>
(
<figureCitation id="35CB05DEA861FFC6D74EFDD98CC5FDFB" box="[302,384,578,602]" captionStart="Figs 9–12" captionStartId="9.[121,158,1494,1512]" captionTargetBox="[145,998,148,1407]" captionTargetId="figure-60@9.[567,998,842,1405]" captionTargetPageId="9" captionText="Figs 9–12: Orictites spp., habitus. 9: O. barclayi sp. nov., holotype, female; 10: O. brancuccii sp. nov., holotype, male; 11: O. barclayi minorfrater ssp. nov., holotype, male; 12: O. tubercucollis sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284330" httpUri="https://zenodo.org/record/6284330/files/figure.png" pageId="50" pageNumber="51">Figs 10</figureCitation>
,
<figureCitation id="35CB05DEA861FFC6D7EEFDD98CEAFDFB" box="[398,431,578,602]" captionStart="Figs 27–30" captionStartId="24.[159,196,982,1000]" captionTargetBox="[159,1043,130,960]" captionText="Figs 27–30: Orictites spp., aedeagus, ventral view. 27: O. brancuccii sp. nov., holotype, scale bar 0.2 mm; 28: O. barclayi minorfrater ssp. nov., holotype, scale bar 0.2 mm; 29: O. tubercucollis sp. nov., holotype, scale bar 0.2 mm; 30: O. anteriorlatus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284354" httpUri="https://zenodo.org/record/6284354/files/figure.png" pageId="50" pageNumber="51">27</figureCitation>
; pp. 10, 25)
</paragraph>
<paragraph id="AD4F195BA861FFC6D6FFFD168F3CFD51" blockId="50.[159,1044,652,1573]" pageId="50" pageNumber="51">
Measurements (n = 3): Body length
<quantity id="6A08B4BEA861FFC6D45BFD168FEDFD04" box="[571,680,653,677]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" metricValueMax="7.4" metricValueMin="7.0" pageId="50" pageNumber="51" unit="mm" value="7.2" valueMax="7.4" valueMin="7.0">7–7.4 mm</quantity>
(
<emphasis id="9F84C549A861FFC6D4D8FD178F8DFD04" bold="true" box="[696,712,652,677]" italics="true" pageId="50" pageNumber="51"></emphasis>
=
<quantity id="6A08B4BEA861FFC6D488FD168E08FD04" box="[744,845,653,677]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" pageId="50" pageNumber="51" unit="mm" value="7.25">7.25 mm</quantity>
), width
<quantity id="6A08B4BEA861FFC6D5C5FD168D8CFD6B" metricMagnitude="-3" metricUnit="m" metricValue="2.185" metricValueMax="2.19" metricValueMin="2.18" pageId="50" pageNumber="51" unit="mm" value="2.185" valueMax="2.19" valueMin="2.18">2.18–2.19 mm</quantity>
(
<emphasis id="9F84C549A861FFC6D6B8FD2A8DADFD6B" bold="true" box="[216,232,689,714]" italics="true" pageId="50" pageNumber="51"></emphasis>
=
<quantity id="6A08B4BEA861FFC6D766FD298C29FD6B" box="[262,364,690,714]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="50" pageNumber="51" unit="mm" value="2.18">2.18 mm</quantity>
); ratio length/width of pronotum 0.88–0.90 (
<emphasis id="9F84C549A861FFC6D508FD2A8E3DFD6B" bold="true" box="[872,888,689,714]" italics="true" pageId="50" pageNumber="51"></emphasis>
= 0.89); ratio length/width of elytra 1.7–1.75 (
<emphasis id="9F84C549A861FFC6D46BFD4C8F5EFD51" bold="true" box="[523,539,727,752]" italics="true" pageId="50" pageNumber="51"></emphasis>
= 1.73).
</paragraph>
<paragraph id="AD4F195BA861FFC6D6A4FD668E2EFCB4" blockId="50.[159,1044,652,1573]" box="[196,875,765,789]" pageId="50" pageNumber="51">Colour: Shiny. Black. Antennae, mouthparts and legs brown.</paragraph>
<paragraph id="AD4F195BA861FFC7D6A4FCB98CD8FDFB" blockId="50.[159,1044,652,1573]" lastBlockId="51.[121,1006,93,1573]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
Head: A quarter narrower than pronotum. Clypeus fused with wings, clypeal wings indistinctly projecting, anterior margin straight, with raised broad transverse knob between the two clypeal setae, margin of wings obtusely angled, hollowed out, divided from middle part of clypeus by carinae, divided from supraantennal plates by distinct nearly right-angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, smooth, with reflexed rounded margin, interrupted from supraorbital carina. Frons separated from clypeus by transverse broad furrow, with median impression on frons (indistinct in
<specimenCount id="BBF6D2D2A861FFC6D4A7FBD68E2DFBC4" box="[711,872,1101,1125]" pageId="50" pageNumber="51" type="generic" typeStatus="paratype">one paratype</specimenCount>
), broad furrow between frons and frontal carina, frons moderately convex, smooth. Clypeus with transverse rugae and a few small punctures. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle at posterior genae level. Eyes flattened but still convex. Genae distinct, gently rounded, as long as eyes, not as high as eyes, forming obtuse angle at neck. Grooves for reception of antennae of moderate length, 0.5 of eye diameter. Neck constriction distinct, with band of punctures. Neck laterally covered with small punctures and rugae. Labrum advanced at middle, 7-setose, ciliate laterally, nearly smooth in anterior part, with isodiametric reticulation in posterior part. Mandibles robust, length half of head, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles rounded towards base. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, indistinctly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A860FFC7D4AFFFE48FA3FF33" attach="left" box="[719,742,127,146]" fontSize="8" pageId="51" pageNumber="52">nd</superScript>
segment, 2
<superScript id="5A85B413A860FFC7D50DFFE48EC1FF33" attach="left" box="[877,900,127,146]" fontSize="8" pageId="51" pageNumber="52">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with rugae and a few punctures. Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with broad and rounded tubercle, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming broad obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae relatively long, reaching up to posterior third of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine longitudinal reticulation, segments 5–10 elongate (L/
<geoCoordinate id="C8C47F9CA860FFC7D5B2FE638DECFD94" degrees="1. 55" direction="west" orientation="longitude" pageId="51" pageNumber="52" precision="555" value="-1.55">W 1.55</geoCoordinate>
), antennomeres cylindriform, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA860FFC7D6FEFDF38ED0FB2B" blockId="51.[121,1006,93,1573]" pageId="51" pageNumber="52">Pronotum: Square, distinctly wider than long, outline with transverse appearance, disc distinctly convex, reflexed margin convex from anterior to posterior angles. Disc distinctly convex. Anterior margin bisinuate. Reflexed lateral margin with traces of scars (120x). Margin from posterior angle to base running as slightly convex line. Lateral channel distinctly broadened between the two lateral setigerous punctures, with isodiametric reticulation. Anterior setigerous puncture located in anterior quarter adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle distinctly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as tooth, projecting antero-laterally. Anterior transverse line indistinct, formed by minute punctures. Median line deep, moderately broad, rugose, reaching up to anterior transverse line but not joining, not adjoining base. Surface with small punctures and basally with transverse rugae, basal impression indistinct. Basal channel short, moderately broad. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA860FFC7D6FEFB038E06F984" blockId="51.[121,1006,93,1573]" pageId="51" pageNumber="52">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles and a second row of minute punctures. Reflexed margin slightly crenulated posterior to humerus. Margin bending with channel over rounded humerus up to 5
<superScript id="5A85B413A860FFC7D7C6FAB18CFCFA9C" attach="left" box="[422,441,1322,1341]" fontSize="8" pageId="51" pageNumber="52">th</superScript>
stria. Humerus with distinct tooth. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1
<superScript id="5A85B413A860FFC7D52EFAD48E1AFAC3" attach="left" box="[846,863,1359,1378]" fontSize="8" pageId="51" pageNumber="52">st</superScript>
stria. Striae 1–4 free at base, all striae deep, nearly impunctate, striae 1 and 2 ending free at apex, 3–4, and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct. Intervals moderately convex, 3
<superScript id="5A85B413A860FFC7D4F1FA248FE1FA73" attach="left" box="[657,676,1471,1490]" fontSize="8" pageId="51" pageNumber="52">rd</superScript>
and4
<superScript id="5A85B413A860FFC7D486FA248FBCFA73" attach="left" box="[742,761,1471,1490]" fontSize="8" pageId="51" pageNumber="52">th</superScript>
moderately raised at base, both with tubercle at base, 7
<superScript id="5A85B413A860FFC7D473FA7F8F63FA56" attach="left" box="[531,550,1508,1527]" fontSize="8" pageId="51" pageNumber="52">th</superScript>
and 8
<superScript id="5A85B413A860FFC7D411FA7F8FC1FA56" attach="left" box="[625,644,1508,1527]" fontSize="8" pageId="51" pageNumber="52">th</superScript>
carinate. All intervals shiny on disc. Interval 3 with four setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA867FFC0D6A4FFC68F4DFFD4" blockId="52.[159,1045,93,1200]" box="[196,520,93,117]" pageId="52" pageNumber="53">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA867FFC0D6A4FF198CDDFEF4" blockId="52.[159,1045,93,1200]" pageId="52" pageNumber="53">
Ventral surface: Epipleuron shiny, with big punctures, submarginal furrow distinct. Proepisternum and episternum with punctures, keeled at middle like an inverted V, mesosternum smooth. Sternites with punctures, 3
<superScript id="5A85B413A867FFC0D5E1FF518ED1FF7C" attach="left" box="[897,916,202,221]" fontSize="8" pageId="52" pageNumber="53">rd</superScript>
to 5
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with paralateral ambulatory setae at each side, ventral strigae distinct, with transverse reticulation, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA867FFC0D6A4FEF98EA9FDB1" blockId="52.[159,1045,93,1200]" pageId="52" pageNumber="53">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, with indistinct dorsal furrow, with minute punctures, basal tarsomere elongated, longer than the following four together. Mesotibia with preapical spine nearly at apex of tibia, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA867FFC0D6A4FD868E30FD04" blockId="52.[159,1045,93,1200]" pageId="52" pageNumber="53">
Male genitalia (
<figureCitation id="35CB05DEA867FFC0D71CFD868C88FD94" box="[380,461,541,565]" captionStart="Figs 27–30" captionStartId="24.[159,196,982,1000]" captionTargetBox="[159,1043,130,960]" captionText="Figs 27–30: Orictites spp., aedeagus, ventral view. 27: O. brancuccii sp. nov., holotype, scale bar 0.2 mm; 28: O. barclayi minorfrater ssp. nov., holotype, scale bar 0.2 mm; 29: O. tubercucollis sp. nov., holotype, scale bar 0.2 mm; 30: O. anteriorlatus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284354" httpUri="https://zenodo.org/record/6284354/files/figure.png" pageId="52" pageNumber="53">Fig. 27</figureCitation>
): Moderately long, distinctly sclerotized. Median lobe moderately arcuate, apex not distorted, without spatula. Endophallus with numerous blunt teeth. Parameres of about same length, both slender, longer than median lobe, ventral one distinctly twisted, asetose.
</paragraph>
<paragraph id="AD4F195BA867FFC0D6A4FD298CBBFD6B" blockId="52.[159,1045,93,1200]" box="[196,510,690,714]" pageId="52" pageNumber="53">Female genitalia: Unknown.</paragraph>
<paragraph id="AD4F195BA867FFC0D6A4FD438F30FCB4" blockId="52.[159,1045,93,1200]" pageId="52" pageNumber="53">
Variation: In the
<typeStatus id="724BA7F9A867FFC0D7E3FD438CADFD51" box="[387,488,728,752]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
the median impression on the clypeus is deeply impressed, in the
<typeStatus id="724BA7F9A867FFC0D708FD668C8BFCB4" box="[360,462,765,789]" pageId="52" pageNumber="53" type="paratype">paratype</typeStatus>
it is indistinct.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A867FFC0D6A4FCB98F57FC54" pageId="52" pageNumber="53" type="diagnosis">
<paragraph id="AD4F195BA867FFC0D6A4FCB98F57FC54" blockId="52.[159,1045,93,1200]" pageId="52" pageNumber="53">Diagnosis: A big massive black species with transverse knob on the clypeus and transverse broad furrow, with distinct, moderately sharp humeral tooth, tubercles at the base of intervals 3 and 4 of the elytra, and four setigerous punctures on interval 3. Moreover, the anterior transverse line is indistinctly developed. Distinguished from all other species by the transverse and conspicuously convex pronotum.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A867FFC0D6A4FB998DB3FBE1" pageId="52" pageNumber="53" type="distribution">
<paragraph id="AD4F195BA867FFC0D6A4FB998DB3FBE1" blockId="52.[159,1045,93,1200]" pageId="52" pageNumber="53">
Distribution: Known from
<collectingCountry id="D5E759CBA867FFC0D463FB998F7BFBBB" box="[515,574,1026,1050]" name="Laos" pageId="52" pageNumber="53">Laos</collectingCountry>
, collected at an altitude of 400–1500 metres.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A867FFC0D6A4FBD68CCAFB11" pageId="52" pageNumber="53" type="etymology">
<paragraph id="AD4F195BA867FFC0D6A4FBD68CCAFB11" blockId="52.[159,1045,93,1200]" pageId="52" pageNumber="53">
Etymology: The species is dedicated to the memory of Dr. Michel Brancucci, Curator
<taxonomicName id="6AF062D8A867FFC0D69BFBE98C33FB2B" authorityName="Linnaeus" authorityYear="1758" box="[251,374,1138,1162]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
and Head of the Department of Entomology (NHMB) who passed away in 2012.
</paragraph>
</subSubSection>
</treatment>
</document>

138
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<taxonomicName id="6AF062D8A863FFC4D6FFFB798CAFFB5A" ID-CoL="8MMPJ" authorityName="Balkenohl" authorityYear="2017" box="[159,490,1250,1275]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="barclayi" status="subsp. nov." subSpecies="minorfrater">
<emphasis id="9F84C549A863FFC4D6FFFB798CAFFB5A" bold="true" box="[159,490,1250,1275]" italics="true" pageId="48" pageNumber="49">Orictites barclayi minorfrater</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A863FFC4D790FB798F11FB5B" box="[496,596,1250,1274]" pageId="48" pageNumber="49" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
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<subSubSection id="E5EA4AD0A863FFC4D6FFFAB68CFDFA14" pageId="48" pageNumber="49" type="materials_examined">
<paragraph id="AD4F195BA863FFC4D6FFFAB68CFDFA14" blockId="48.[159,1044,1325,1461]" pageId="48" pageNumber="49">
Type material:
<materialsCitation id="1D981306A863FFC4D72AFAB68CFDFA14" ID-GBIF-Occurrence="3497007308" collectingDate="2003-03-04" collectingDateMax="2003-03-18" collectingDateMin="2003-03-04" collectionCode="NHMB" collectorName="Dembicky" country="SUMATRA" elevation="475" latitude="-2.0833333" location="SUMATRA" longLatPrecision="1307" longitude="101.25" pageId="48" pageNumber="49" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<typeStatus id="724BA7F9A863FFC4D72AFAB68CF3FAE4" box="[330,438,1325,1349]" pageId="48" pageNumber="49" type="holotype">Holotype</typeStatus>
:
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, labels: white, black printed: “W
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prov.: Kerinci Seblat N.P.:
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NE Tapan
</locationDeviation>
: MUARA SAKO
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E env.:
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<geoCoordinate id="C8C47F9CA863FFC4D5C5FAC9894AFACB" box="[933,1039,1362,1386]" degrees="101" direction="east" minutes="15" orientation="longitude" pageId="48" pageNumber="49" precision="925" value="101.25">101°15´E</geoCoordinate>
:
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:
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leg.:
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</date>
” / “Collection Naturhistorisches Museum Basel” (
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).
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</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A862FFC5D619FFC68C9DFC0B" pageId="49" pageNumber="50" type="description">
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<emphasis id="9F84C549A862FFC5D619FFC68DB9FFD7" bold="true" box="[121,252,93,118]" pageId="49" pageNumber="50">Description</emphasis>
(
<figureCitation id="35CB05DEA862FFC5D768FFC68C1CFFD4" box="[264,345,93,117]" captionStart="Figs 9–12" captionStartId="9.[121,158,1494,1512]" captionTargetBox="[145,998,148,1407]" captionTargetId="figure-60@9.[567,998,842,1405]" captionTargetPageId="9" captionText="Figs 9–12: Orictites spp., habitus. 9: O. barclayi sp. nov., holotype, female; 10: O. brancuccii sp. nov., holotype, male; 11: O. barclayi minorfrater ssp. nov., holotype, male; 12: O. tubercucollis sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284330" httpUri="https://zenodo.org/record/6284330/files/figure.png" pageId="49" pageNumber="50">Figs 11</figureCitation>
,
<figureCitation id="35CB05DEA862FFC5D707FFC68CCDFFD4" box="[359,392,93,117]" captionStart="Figs 27–30" captionStartId="24.[159,196,982,1000]" captionTargetBox="[159,1043,130,960]" captionText="Figs 27–30: Orictites spp., aedeagus, ventral view. 27: O. brancuccii sp. nov., holotype, scale bar 0.2 mm; 28: O. barclayi minorfrater ssp. nov., holotype, scale bar 0.2 mm; 29: O. tubercucollis sp. nov., holotype, scale bar 0.2 mm; 30: O. anteriorlatus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284354" httpUri="https://zenodo.org/record/6284354/files/figure.png" pageId="49" pageNumber="50">28</figureCitation>
; pp. 10, 25)
</paragraph>
<paragraph id="AD4F195BA862FFC5D619FF338FDAFF44" blockId="49.[121,1006,168,1349]" pageId="49" pageNumber="50">
Measurements: Body length
<quantity id="6A08B4BEA862FFC5D7A8FF338F74FF61" box="[456,561,168,192]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" pageId="49" pageNumber="50" unit="mm" value="4.25">4.25 mm</quantity>
, width
<quantity id="6A08B4BEA862FFC5D4E7FF338FB5FF61" box="[647,752,168,192]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="49" pageNumber="50" unit="mm" value="1.09">1.09 mm</quantity>
; ratio length/width of pronotum 1.04; ratio length/width of elytra 1.93.
</paragraph>
<paragraph id="AD4F195BA862FFC5D619FF698DBCFE91" blockId="49.[121,1006,168,1349]" pageId="49" pageNumber="50">The subspecies differs from the nominotypical subspecies in the following characters:</paragraph>
<paragraph id="AD4F195BA862FFC5D6FEFEA68E29FEF4" blockId="49.[121,1006,168,1349]" box="[158,876,317,341]" pageId="49" pageNumber="50">Colour: Intervals of elytra with reddish-brown stripes at middle.</paragraph>
<paragraph id="AD4F195BA862FFC5D6FEFEF98C31FE4B" blockId="49.[121,1006,168,1349]" pageId="49" pageNumber="50">
Head: Frontal carinae convex, with medium sized puncture on the frons. Labrum anteriorly slightly advanced. Punctures of neck constriction bigger than in the nominotypical subspecies. Antennae with segments 5–10 slightly transverse (L/
<geoCoordinate id="C8C47F9CA862FFC5D774FE498C20FE4B" box="[276,357,466,490]" degrees="0. 76" direction="west" orientation="longitude" pageId="49" pageNumber="50" precision="555" value="-0.76">W 0.76</geoCoordinate>
).
</paragraph>
<paragraph id="AD4F195BA862FFC5D6FEFE638D93FDFB" blockId="49.[121,1006,168,1349]" pageId="49" pageNumber="50">Pronotum: Median line not quite reaching level of anterior transverse line, indistinctly joining base. Surface with irregularly scattered punctures of medium size.</paragraph>
<paragraph id="AD4F195BA862FFC5D6FEFDF38C66FD04" blockId="49.[121,1006,168,1349]" pageId="49" pageNumber="50">Elytra: Intervals 3 and 4 at base with tubercle. Interval 3 with six setigerous punctures.</paragraph>
<paragraph id="AD4F195BA862FFC5D6FEFD298CFBFD6B" blockId="49.[121,1006,168,1349]" box="[158,446,690,714]" pageId="49" pageNumber="50">Legs: Anterior tibia shiny.</paragraph>
<paragraph id="AD4F195BA862FFC5D6FEFD438CACFC24" blockId="49.[121,1006,168,1349]" pageId="49" pageNumber="50">
Male genitalia (
<figureCitation id="35CB05DEA862FFC5D72EFD438CD9FD51" box="[334,412,728,752]" captionStart="Figs 27–30" captionStartId="24.[159,196,982,1000]" captionTargetBox="[159,1043,130,960]" captionText="Figs 27–30: Orictites spp., aedeagus, ventral view. 27: O. brancuccii sp. nov., holotype, scale bar 0.2 mm; 28: O. barclayi minorfrater ssp. nov., holotype, scale bar 0.2 mm; 29: O. tubercucollis sp. nov., holotype, scale bar 0.2 mm; 30: O. anteriorlatus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284354" httpUri="https://zenodo.org/record/6284354/files/figure.png" pageId="49" pageNumber="50">Fig. 28</figureCitation>
): Moderately long, slightly sclerotized. Median lobe slender, moderately and regularly arcuate, spatula stick-like at apex. Endophallus with some minute teeth. Parameres of about same length, both slender, dorsal one distinctly twisted, both of them with two setae apically (in the dorsal one, best visible at 400x).
</paragraph>
<paragraph id="AD4F195BA862FFC5D6FEFC098C9DFC0B" blockId="49.[121,1006,168,1349]" box="[158,472,914,938]" pageId="49" pageNumber="50">Female genitalia: Unknown.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A862FFC5D6FEFC238DF9FBE1" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="AD4F195BA862FFC5D6FEFC238DF9FBE1" blockId="49.[121,1006,168,1349]" pageId="49" pageNumber="50">
Diagnosis: Smaller body than the nominotypical subspecies and distinguished by the six setigerous punctures on the 3
<superScript id="5A85B413A862FFC5D4D2FC418F80FC4C" attach="left" box="[690,709,986,1005]" fontSize="8" pageId="49" pageNumber="50">rd</superScript>
interval of the elytra and tubercles at base of intervals 3 and 4, and the puncture in the middle of the frons.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A862FFC5D6FEFBD68EA2FBC4" box="[158,999,1101,1125]" pageId="49" pageNumber="50" type="distribution">
<paragraph id="AD4F195BA862FFC5D6FEFBD68EA2FBC4" blockId="49.[121,1006,168,1349]" box="[158,999,1101,1125]" pageId="49" pageNumber="50">
Distribution: Known from the Kerinci Seblat Natonal Park in
<collectingRegion id="6F34D7B9A862FFC5D520FBD68EA6FBC4" box="[832,995,1101,1125]" country="Indonesia" name="Sumatera Barat" pageId="49" pageNumber="50">West Sumatra</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A862FFC5D6FEFBE98CE6FB74" pageId="49" pageNumber="50" type="discussion">
<paragraph id="AD4F195BA862FFC5D6FEFBE98CE6FB74" blockId="49.[121,1006,168,1349]" pageId="49" pageNumber="50">Remark: Whether this subspecies deserves species status should be revaluated when male and female genitalia of both subspecies are at hand and more material is available.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A862FFC5D6FEFB798C32FAE4" pageId="49" pageNumber="50" type="etymology">
<paragraph id="AD4F195BA862FFC5D6FEFB798C32FAE4" blockId="49.[121,1006,168,1349]" pageId="49" pageNumber="50">Etymology. By contrast to the nominotypical subspecies, the specimen has a smaller body size and is a male serving for the derivation of the name, which means “little brother”.</paragraph>
</subSubSection>
</treatment>
</document>

401
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<heading id="F607AE37A867FFC0D6FFFAB68CB2FAE4" box="[159,503,1325,1350]" centered="true" fontSize="12" level="2" pageId="52" pageNumber="53" reason="2">
<taxonomicName id="6AF062D8A867FFC0D6FFFAB68CDCFAE7" ID-CoL="8MDDT" authorityName="Balkenohl" authorityYear="2017" box="[159,409,1325,1350]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="tubercucollis" status="sp. nov.">
<emphasis id="9F84C549A867FFC0D6FFFAB68CDCFAE7" bold="true" box="[159,409,1325,1350]" italics="true" pageId="52" pageNumber="53">Orictites tubercucollis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A867FFC0D7FFFAB68CB2FAE4" box="[415,503,1325,1349]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A867FFC0D6FFFAE38E42FA14" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="AD4F195BA867FFC0D6FFFAE38E42FA14" blockId="52.[159,1043,1400,1573]" pageId="52" pageNumber="53">
Type material:
<materialsCitation id="1D981306A867FFC0D729FAE38E42FA14" ID-GBIF-Occurrence="3497007346" collectingDate="1995-08-14" collectionCode="CBM-ZSM" country="Thailand" location="Doi Suthep Pui NP" pageId="52" pageNumber="53" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="holotype">
<typeStatus id="724BA7F9A867FFC0D729FAE38CF0FA31" box="[329,437,1400,1424]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A867FFC0D7A1FAE38C90FA2E" box="[449,469,1400,1423]" pageId="52" pageNumber="53" type="male">♂</specimenCount>
, labels: white, black printed: “
<collectingCountry id="D5E759CBA867FFC0D553FAE38EDCFA31" box="[819,921,1400,1424]" name="Thailand" pageId="52" pageNumber="53">Thailand</collectingCountry>
<date id="D94E3F9BA867FFC0D5C1FAE38956FA31" box="[929,1043,1400,1424]" pageId="52" pageNumber="53" value="1995-08-14">
<collectingDate id="C90AC673A867FFC0D5C1FAE38956FA31" box="[929,1043,1400,1424]" pageId="52" pageNumber="53" value="1995-08-14">14.8.1995</collectingDate>
</date>
(
<collectingRegion id="6F34D7B9A867FFC0D6C5FA068C62FA14" box="[165,295,1437,1461]" country="Thailand" name="Chiang Mai" pageId="52" pageNumber="53">Chiang Mai</collectingRegion>
)
<location id="A82F4F80A867FFC0D753FA068F44FA14" LSID="urn:lsid:plazi:treatment:2559A84DA867FFCCD6FFFAB68CDFFD21:A82F4F80A867FFC0D753FA068F44FA14" box="[307,513,1437,1461]" country="Thailand" name="Doi Suthep Pui NP" pageId="52" pageNumber="53" stateProvince="Chiang Mai">Doi Suthep Pui NP</location>
R. Grimm” (
<collectionCode id="CBE1819EA867FFC0D4E6FA068FBBFA14" box="[646,766,1437,1461]" pageId="52" pageNumber="53">CBM-ZSM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A867FFC1D6A4FA598C24FD51" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="discussion">
<paragraph id="AD4F195BA867FFC0D6A4FA598E95FA7B" blockId="52.[159,1043,1400,1573]" box="[196,976,1474,1498]" pageId="52" pageNumber="53">
Remark: In the
<typeStatus id="724BA7F9A867FFC0D70FFA598C91FA7B" box="[367,468,1474,1498]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
some antennomeres are missing at one side.
</paragraph>
<paragraph id="AD4F195BA867FFC1D6FFFA738C24FD51" blockId="52.[159,1043,1400,1573]" lastBlockId="53.[121,1007,93,752]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<materialsCitation id="1D981306A867FFC0D6FFFA738CB9F984" ID-GBIF-Occurrence="3497007323" collectingDate="2004-05-07" collectingDateMax="2004-05-10" collectingDateMin="2004-05-07" collectionCode="CBB" collectorName="R. Grimm" country="Thailand" elevation="1200" location="Doi Suthep" municipality="Doi Suthep" pageId="52" pageNumber="53" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="paratype">
<typeStatus id="724BA7F9A867FFC0D6FFFA738C5EF9A1" box="[159,283,1512,1536]" pageId="52" pageNumber="53" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="BBF6D2D2A867FFC0D750FA738C20FA5E" box="[304,357,1512,1536]" pageId="52" pageNumber="53" type="male">1 ♂</specimenCount>
, “
<collectingCountry id="D5E759CBA867FFC0D7E6FA738CB1F9A1" box="[390,500,1512,1536]" name="Thailand" pageId="52" pageNumber="53">Thailand</collectingCountry>
, NWW
<collectingRegion id="6F34D7B9A867FFC0D43BFA738FA8F9A1" box="[603,749,1512,1536]" country="Thailand" name="Chiang Mai" pageId="52" pageNumber="53">Chiang Mai</collectingRegion>
,
<collectingMunicipality id="4D2B8321A867FFC0D562FA738ED1F9A1" box="[770,916,1512,1536]" pageId="52" pageNumber="53">Doi Suthep</collectingMunicipality>
.
<quantity id="6A08B4BEA867FFC0D5C9FA73894AF9A1" box="[937,1039,1512,1536]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="52" pageNumber="53" unit="m" value="1200.0">
<elevation id="26DDFE68A867FFC0D5C9FA73894AF9A1" box="[937,1039,1512,1536]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="52" pageNumber="53" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<date id="D94E3F9BA867FFC0D6FFF9968C74F984" box="[159,305,1549,1573]" pageId="52" pageNumber="53" value="2004-05-07" valueMax="2004-05-10" valueMin="2004-05-07">
<collectingDate id="C90AC673A867FFC0D6FFF9968C74F984" box="[159,305,1549,1573]" pageId="52" pageNumber="53" value="2004-05-07" valueMax="2004-05-10" valueMin="2004-05-07">7.–10.5.2004</collectingDate>
</date>
,
<collectorName id="00057C8DA867FFC0D75EF9968CEFF984" box="[318,426,1549,1573]" pageId="52" pageNumber="53">R. Grimm</collectorName>
” (
<collectionCode id="CBE1819EA867FFC0D7DEF9968CB0F984" box="[446,501,1549,1573]" collectionName="CBB" pageId="52" pageNumber="53">CBB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A867FFC1D465F9968CECFFD4" ID-GBIF-Occurrence="3497007327" collectingDate="1990-03-27" collectingDateMax="1990-03-29" collectingDateMin="1990-03-27" collectionCode="CBM-, ZSM" collectorName="A. Riedel" lastPageId="53" lastPageNumber="54" location="Sarawak Kuching. Bako NP" pageId="52" pageNumber="53" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="BBF6D2D2A867FFC0D465F9968F6BF984" box="[517,558,1549,1573]" pageId="52" pageNumber="53" type="female">1 ♀</specimenCount>
, “
<location id="A82F4F80A867FFC0D422F9968E2AF984" LSID="urn:lsid:plazi:treatment:2559A84DA867FFCCD6FFFAB68CDFFD21:A82F4F80A867FFC0D422F9968E2AF984" box="[578,879,1549,1573]" name="Sarawak Kuching. Bako NP" pageId="52" pageNumber="53">Sarawak Kuching.Bako NP</location>
<date id="D94E3F9BA867FFC0D515F9968956F984" box="[885,1043,1549,1573]" pageId="52" pageNumber="53" value="1990-03-27" valueMax="1990-03-29" valueMin="1990-03-27">
<collectingDate id="C90AC673A867FFC0D515F9968956F984" box="[885,1043,1549,1573]" pageId="52" pageNumber="53" value="1990-03-27" valueMax="1990-03-29" valueMin="1990-03-27">27.–29.3.1990</collectingDate>
</date>
leg.
<collectorName id="00057C8DA866FFC1D6C1FFC68C4CFFD4" box="[161,265,93,117]" pageId="53" pageNumber="54">A.Riedel</collectorName>
” (
<collectionCode id="CBE1819EA866FFC1D747FFC68C2DFFD4" box="[295,360,93,117]" pageId="53" pageNumber="54">CBM-</collectionCode>
<collectionCode id="CBE1819EA866FFC1D708FFC68CE6FFD4" box="[360,419,93,117]" collectionName="Germany, Muenchen [= Munich], Zoologische Staatssammlung" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="53" pageNumber="54" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A866FFC1D7DBFFC68EC7FF61" ID-GBIF-Occurrence="3497007336" collectingDate="2001-08" collectionCode="BMNH" country="Indonesia" county="Borneo" latitude="-0.05" location="Borneo Kalimantan Tengah Busang" longLatPrecision="1308" longitude="113.98333" municipality="Busang" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Kalimantan Tengah" typeStatus="paratype">
<specimenCount id="BBF6D2D2A866FFC1D7DBFFC68F09FFD4" box="[443,588,93,117]" pageId="53" pageNumber="54" type="generic">1 specimen</specimenCount>
: “
<collectingCountry id="D5E759CBA866FFC1D406FFC68FB1FFD4" box="[614,756,93,117]" name="Indonesia" pageId="53" pageNumber="54">INDONESIA</collectingCountry>
;
<location id="A82F4F80A866FFC1D567FFC68C6EFF3B" LSID="urn:lsid:plazi:treatment:2559A84DA867FFCCD6FFFAB68CDFFD21:A82F4F80A866FFC1D567FFC68C6EFF3B" country="Indonesia" county="Borneo" latitude="-0.05" longLatPrecision="1308" longitude="113.98333" municipality="Busang" name="Borneo Kalimantan Tengah Busang" pageId="53" pageNumber="54" stateProvince="Kalimantan Tengah">Borneo Kalimantan Tengah Busang</location>
/
<location id="A82F4F80A866FFC1D728FF198CCCFF3B" LSID="urn:lsid:plazi:treatment:2559A84DA867FFCCD6FFFAB68CDFFD21:A82F4F80A866FFC1D728FF198CCCFF3B" box="[328,393,130,154]" country="Indonesia" county="Borneo" latitude="-0.05" longLatPrecision="1308" longitude="113.98333" municipality="Busang" name="Rekut" pageId="53" pageNumber="54" stateProvince="Kalimantan Tengah">Rekut</location>
confl.
<geoCoordinate id="C8C47F9CA866FFC1D7BCFF198F6AFF3B" box="[476,559,130,154]" degrees="0" direction="south" minutes="03" orientation="latitude" pageId="53" pageNumber="54" precision="925" value="-0.05">0°03´S</geoCoordinate>
,
<geoCoordinate id="C8C47F9CA866FFC1D45CFF198FE6FF3B" box="[572,675,130,154]" degrees="113" direction="east" minutes="59" orientation="longitude" pageId="53" pageNumber="54" precision="925" value="113.98333">113°59´E</geoCoordinate>
/ Brendell / Mendel
<date id="D94E3F9BA866FFC1D5FDFF198DF6FF61" pageId="53" pageNumber="54" value="2001-08">
<collectingDate id="C90AC673A866FFC1D5FDFF198DF6FF61" pageId="53" pageNumber="54" value="2001-08">August 2001</collectingDate>
</date>
General collecting / 2001–191´Barito Ulu 2001´
<collectionCode id="CBE1819EA866FFC1D569FF338E19FF61" box="[777,860,168,192]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E)”
</materialsCitation>
;
<materialsCitation id="1D981306A866FFC1D5F0FF338F4FFEF4" ID-GBIF-Occurrence="3497007344" collectingDate="2009-05-10" collectingDateMax="2009-05-30" collectingDateMin="2009-05-10" collectionCode="NHMB, CBB" collectorName="M. Brancucci &amp; M. Geiser &amp; Z. Krauss &amp; D. Hauck &amp; V. Kuban" country="Laos" elevation="1450" latitude="103.333336" location="Phonsavan" municipality="Phonsavan" pageId="53" pageNumber="54" specimenCount="3" stateProvince="Xieng Khouang prov." typeStatus="paratype">
<specimenCount id="BBF6D2D2A866FFC1D5F0FF338DF9FF44" pageId="53" pageNumber="54" type="generic">3 specimens</specimenCount>
: “
<collectingCountry id="D5E759CBA866FFC1D6B3FF568C53FF44" box="[211,278,205,229]" name="Laos" pageId="53" pageNumber="54">LAOS</collectingCountry>
–NE,
<collectingRegion id="6F34D7B9A866FFC1D739FF568F12FF44" box="[345,599,205,229]" country="Laos" name="Xiangkhouang" pageId="53" pageNumber="54">Xieng Khouang prov.</collectingRegion>
,~19°37–8´N 103°20´E,
<location id="A82F4F80A866FFC1D50FFF568EA8FF44" LSID="urn:lsid:plazi:treatment:2559A84DA867FFCCD6FFFAB68CDFFD21:A82F4F80A866FFC1D50FFF568EA8FF44" box="[879,1005,205,229]" country="Laos" latitude="103.333336" municipality="Phonsavan" name="Phonsavan" pageId="53" pageNumber="54" stateProvince="Xieng Khouang prov.">Phonsavan</location>
(
<locationDeviation id="BCFD0079A866FFC1D6E2FF698DB1FEAB" box="[130,244,242,266]" pageId="53" pageNumber="54">
<quantity id="6A08B4BEA866FFC1D6E2FF698D87FEAB" box="[130,194,242,266]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="53" pageNumber="54" unit="km" value="30.0">30km</quantity>
NE
</locationDeviation>
):
<location id="A82F4F80A866FFC1D76AFF698CFBFEAB" LSID="urn:lsid:plazi:treatment:2559A84DA867FFCCD6FFFAB68CDFFD21:A82F4F80A866FFC1D76AFF698CFBFEAB" box="[266,446,242,266]" country="Laos" latitude="103.333336" municipality="Phonsavan" name="Phou Sane Mt." pageId="53" pageNumber="54" stateProvince="Xieng Khouang prov.">Phou Sane Mt.</location>
,~
<quantity id="6A08B4BEA866FFC1D7B2FF698F2BFEAB" box="[466,622,242,266]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="53" pageNumber="54" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">
<elevation id="26DDFE68A866FFC1D7B2FF698F2BFEAB" box="[466,622,242,266]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="53" pageNumber="54" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400–1500m</elevation>
</quantity>
,
<date id="D94E3F9BA866FFC1D41DFF698E5AFEAB" box="[637,799,242,266]" pageId="53" pageNumber="54" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<collectingDate id="C90AC673A866FFC1D41DFF698E5AFEAB" box="[637,799,242,266]" pageId="53" pageNumber="54" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">10.–30.v.2009</collectingDate>
</date>
, Z. Krauss leg. / NHMB Basel Prague Laos 2009 Expedition:
<collectorName id="00057C8DA866FFC1D407FE838FB9FE91" box="[615,764,280,304]" pageId="53" pageNumber="54">M. Brancucci</collectorName>
,
<collectorName id="00057C8DA866FFC1D566FE838E31FE91" box="[774,884,280,304]" pageId="53" pageNumber="54">M. Geiser</collectorName>
,
<collectorName id="00057C8DA866FFC1D51DFE838EAEFE91" box="[893,1003,280,304]" pageId="53" pageNumber="54">Z. Krauss</collectorName>
,
<collectorName id="00057C8DA866FFC1D619FEA68DA1FEF4" box="[121,228,317,341]" pageId="53" pageNumber="54">D. Hauck</collectorName>
,
<collectorName id="00057C8DA866FFC1D691FEA68C1CFEF6" box="[241,345,314,343]" pageId="53" pageNumber="54">V. Kubáň</collectorName>
(
<collectionCode id="CBE1819EA866FFC1D708FEA68CF2FEF4" box="[360,439,317,341]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="53" pageNumber="54">NHMB</collectionCode>
,
<collectionCode id="CBE1819EA866FFC1D7A3FEA68CBEFEF4" box="[451,507,317,341]" pageId="53" pageNumber="54">CBB</collectionCode>
)“
</materialsCitation>
;
<materialsCitation id="1D981306A866FFC1D477FEA68D86FE4B" ID-GBIF-Occurrence="3497007334" collectingDate="2009-05-30" collectingDateMax="2009-05-31" collectingDateMin="2009-05-30" collectionCode="NHMB, CBB" collectorName="M. Brancucci &amp; M. Geiser &amp; Z. Krauss &amp; D. Hauck &amp; V. Kuban" country="Laos" elevation="1150" latitude="19.433332" location="Phonsavan town to Phou Padaeng" longLatPrecision="1271" longitude="103.21667" municipality="Phonsavan town" pageId="53" pageNumber="54" specimenCount="2" stateProvince="Xieng Khouang prov." typeStatus="paratype">
<specimenCount id="BBF6D2D2A866FFC1D477FEA68FE9FEF4" box="[535,684,317,341]" pageId="53" pageNumber="54" type="generic">2 specimens</specimenCount>
: “
<collectingCountry id="D5E759CBA866FFC1D4DFFEA68E47FEF4" box="[703,770,317,341]" name="Laos" pageId="53" pageNumber="54">LAOS</collectingCountry>
–NE,
<collectingRegion id="6F34D7B9A866FFC1D521FEA68DFDFEDB" country="Laos" name="Xiangkhouang" pageId="53" pageNumber="54">Xieng Khouang prov.</collectingRegion>
,~
<geoCoordinate id="C8C47F9CA866FFC1D6A8FEF98C6DFEDB" box="[200,296,354,378]" degrees="19" direction="north" minutes="26" orientation="latitude" pageId="53" pageNumber="54" precision="925" value="19.433332">19°26´N</geoCoordinate>
/
<geoCoordinate id="C8C47F9CA866FFC1D752FEF98CDBFEDB" box="[306,414,354,378]" degrees="103" direction="east" minutes="13" orientation="longitude" pageId="53" pageNumber="54" precision="925" value="103.21667">103°13´E</geoCoordinate>
,
<location id="A82F4F80A866FFC1D7CCFEF98E06FEDB" LSID="urn:lsid:plazi:treatment:2559A84DA867FFCCD6FFFAB68CDFFD21:A82F4F80A866FFC1D7CCFEF98E06FEDB" box="[428,835,354,378]" country="Laos" latitude="19.433332" longLatPrecision="1271" longitude="103.21667" municipality="Phonsavan town" name="Phonsavan town to Phou Padaeng" pageId="53" pageNumber="54" stateProvince="Xieng Khouang prov.">Phonsavan town to Phou Padaeng</location>
,
<quantity id="6A08B4BEA866FFC1D530FEF98EAFFEDB" box="[848,1002,354,378]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="53" pageNumber="54" unit="m" value="1150.0" valueMax="1200.0" valueMin="1100.0">
<elevation id="26DDFE68A866FFC1D530FEF98EAFFEDB" box="[848,1002,354,378]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="53" pageNumber="54" unit="m" value="1150.0" valueMax="1200.0" valueMin="1100.0">1100–1200m</elevation>
</quantity>
,
<date id="D94E3F9BA866FFC1D619FE138C59FE01" box="[121,284,392,416]" pageId="53" pageNumber="54" value="2009-05-30" valueMax="2009-05-31" valueMin="2009-05-30">
<collectingDate id="C90AC673A866FFC1D619FE138C59FE01" box="[121,284,392,416]" pageId="53" pageNumber="54" value="2009-05-30" valueMax="2009-05-31" valueMin="2009-05-30">30.–31.v.2009</collectingDate>
</date>
, M. Geiser leg. / NHMB Basel, NMPC Prague Laos 2009 Expedition:
<collectorName id="00057C8DA866FFC1D765FE368CE5FE64" box="[261,416,429,453]" pageId="53" pageNumber="54">M. Brancucci</collectorName>
,
<collectorName id="00057C8DA866FFC1D7CCFE368F5AFE64" box="[428,543,429,453]" pageId="53" pageNumber="54">M. Geiser</collectorName>
,
<collectorName id="00057C8DA866FFC1D44BFE368FD8FE64" box="[555,669,429,453]" pageId="53" pageNumber="54">Z. Krauss</collectorName>
,
<collectorName id="00057C8DA866FFC1D4C9FE368E53FE64" box="[681,790,429,453]" pageId="53" pageNumber="54">D. Hauck</collectorName>
,
<collectorName id="00057C8DA866FFC1D543FE368EC9FE67" box="[803,908,428,454]" pageId="53" pageNumber="54">V. Kubáň</collectorName>
(
<collectionCode id="CBE1819EA866FFC1D5FBFE368EAFFE64" box="[923,1002,429,453]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="53" pageNumber="54">NHMB</collectionCode>
,
<collectionCode id="CBE1819EA866FFC1D619FE498DF6FE4B" box="[121,179,466,490]" pageId="53" pageNumber="54">CBB</collectionCode>
)“
</materialsCitation>
;
<materialsCitation id="1D981306A866FFC1D6B5FE498E93FDFB" ID-GBIF-Occurrence="3497007309" collectingDate="2009-05-10" collectingDateMax="2009-05-30" collectingDateMin="2009-05-10" collectionCode="NHMB" collectorName="M. Brancucci &amp; M. Geiser &amp; Z. Krauss &amp; D. Hauck &amp; V. Kuban" country="Laos" elevation="1400" latitude="103.333336" location="Phou Sane Mt." municipality="Ban Na Lam" pageId="53" pageNumber="54" specimenCount="2" specimenCount-male="1" stateProvince="Xieng Khouang prov." typeStatus="paratype">
<specimenCount id="BBF6D2D2A866FFC1D6B5FE498C4DFE4B" box="[213,264,466,490]" pageId="53" pageNumber="54" type="male">1 ♂</specimenCount>
,
<specimenCount id="BBF6D2D2A866FFC1D77EFE498CF4FE4B" box="[286,433,466,490]" pageId="53" pageNumber="54" type="generic">1 specimen</specimenCount>
, “
<collectingCountry id="D5E759CBA866FFC1D7ABFE498F4AFE4B" box="[459,527,466,490]" name="Laos" pageId="53" pageNumber="54">LAOS</collectingCountry>
–NE,
<collectingRegion id="6F34D7B9A866FFC1D437FE498E1AFE4B" box="[599,863,466,490]" country="Laos" name="Xiangkhouang" pageId="53" pageNumber="54">Xieng Khouang prov.</collectingRegion>
,~19°37–8´N 103°20´E,
<locationDeviation id="BCFD0079A866FFC1D690FE638C9FFDB1" box="[240,474,504,528]" location="Phonsavan" pageId="53" pageNumber="54">
<quantity id="6A08B4BEA866FFC1D690FE638C75FDB1" box="[240,304,504,528]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="53" pageNumber="54" unit="km" value="30.0">30km</quantity>
NE Phonsavan
</locationDeviation>
:
<collectingMunicipality id="4D2B8321A866FFC1D791FE638F3FFDB1" box="[497,634,504,528]" pageId="53" pageNumber="54">Ban Na Lam</collectingMunicipality>
<emphasis id="9F84C549A866FFC1D4E0FE6D8FDFFDB1" bold="true" box="[640,666,502,528]" italics="true" pageId="53" pageNumber="54"></emphasis>
<location id="A82F4F80A866FFC1D4C2FE638E09FDB1" LSID="urn:lsid:plazi:treatment:2559A84DA867FFCCD6FFFAB68CDFFD21:A82F4F80A866FFC1D4C2FE638E09FDB1" box="[674,844,504,528]" country="Laos" latitude="103.333336" municipality="Ban Na Lam" name="Phou Sane Mt." pageId="53" pageNumber="54" stateProvince="Xieng Khouang prov.">Phou Sane Mt.</location>
,
<quantity id="6A08B4BEA866FFC1D52FFE638EAFFDB1" box="[847,1002,504,528]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="53" pageNumber="54" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">
<elevation id="26DDFE68A866FFC1D52FFE638EAFFDB1" box="[847,1002,504,528]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="53" pageNumber="54" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300–1500m</elevation>
</quantity>
,
<date id="D94E3F9BA866FFC1D619FD868C5FFD94" box="[121,282,541,565]" pageId="53" pageNumber="54" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<collectingDate id="C90AC673A866FFC1D619FD868C5FFD94" box="[121,282,541,565]" pageId="53" pageNumber="54" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">10.–30.v.2009</collectingDate>
</date>
, M. Brancucci leg. / NHMB Basel NMPC Prague Laos 2009 Expedition:
<collectorName id="00057C8DA866FFC1D761FDD98CD3FDFB" box="[257,406,578,602]" pageId="53" pageNumber="54">M. Brancucci</collectorName>
,
<collectorName id="00057C8DA866FFC1D7C0FDD98F4BFDFB" box="[416,526,578,602]" pageId="53" pageNumber="54">M. Geiser</collectorName>
,
<collectorName id="00057C8DA866FFC1D478FDD98FC0FDFB" box="[536,645,578,602]" pageId="53" pageNumber="54">Z. Krauss</collectorName>
,
<collectorName id="00057C8DA866FFC1D4EEFDD98FB2FDFB" box="[654,759,578,602]" pageId="53" pageNumber="54">D. Hauck</collectorName>
,
<collectorName id="00057C8DA866FFC1D561FDD98E22FDFD" box="[769,871,575,604]" pageId="53" pageNumber="54">V. Kubáň</collectorName>
(
<collectionCode id="CBE1819EA866FFC1D513FDD98E83FDFB" box="[883,966,578,602]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="53" pageNumber="54">NHMB</collectionCode>
)“
</materialsCitation>
;
<materialsCitation id="1D981306A866FFC1D5BFFDD98DA4FD04" ID-GBIF-Occurrence="3497007319" collectingDate="1917-02" collectionCode="BMNH" country="Laos" location="Tonkin Hanoi" pageId="53" pageNumber="54" specimenCount="1" specimenCount-female="1" stateProvince="Xiangkhouang" typeStatus="paratype">
<specimenCount id="BBF6D2D2A866FFC1D5BFFDD98DC8FDDE" pageId="53" pageNumber="54" type="female">1 ♀</specimenCount>
, “
<location id="A82F4F80A866FFC1D6C3FDF38C79FD21" LSID="urn:lsid:plazi:treatment:2559A84DA867FFCCD6FFFAB68CDFFD21:A82F4F80A866FFC1D6C3FDF38C79FD21" box="[163,316,616,640]" country="Laos" name="Tonkin Hanoi" pageId="53" pageNumber="54" stateProvince="Xiangkhouang">Tonkin Hanoi</location>
<date id="D94E3F9BA866FFC1D723FDF38CF7FD21" box="[323,434,616,640]" pageId="53" pageNumber="54" value="1917-02">
<collectingDate id="C90AC673A866FFC1D723FDF38CF7FD21" box="[323,434,616,640]" pageId="53" pageNumber="54" value="1917-02">Feb. 1917</collectingDate>
</date>
R.V.de Salvaua. /
<taxonomicName id="6AF062D8A866FFC1D4E3FDF38FBCFD21" box="[643,761,616,640]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="9F84C549A866FFC1D4E3FDF38F95FD21" box="[643,720,616,640]" italics="true" pageId="53" pageNumber="54">Clivina</emphasis>
sp.
</taxonomicName>
/ Brit. Mus. 1921–89 (
<collectionCode id="CBE1819EA866FFC1D61FFD168D97FD04" box="[127,210,653,677]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="53" pageNumber="54">BMNH</collectionCode>
)“
</materialsCitation>
.
<materialsCitation id="1D981306A866FFC1D68EFD168C18FD51" ID-GBIF-Occurrence="3497007329" collectingDate="2008-12-23" collectingDateMax="2008-12-28" collectingDateMin="2008-12-23" collectionCode="IRSNB" collectorName="A. Napolov" country="Laos" latitude="1.1333333" location="Putussibau" longLatPrecision="1308" longitude="113.0" municipality="Ca." pageId="53" pageNumber="54" specimenCount="1" stateProvince="Xiangkhouang" typeStatus="paratype">
<specimenCount id="BBF6D2D2A866FFC1D68EFD168C32FD04" box="[238,375,653,677]" pageId="53" pageNumber="54" type="generic">1 specimen</specimenCount>
: “Coll. I.R.Sc.N.B. BORNEO:ca
<locationDeviation id="BCFD0079A866FFC1D485FD168EAFFD04" box="[741,1002,653,677]" location="Pontianak" pageId="53" pageNumber="54">
<quantity id="6A08B4BEA866FFC1D485FD168E71FD04" box="[741,820,653,677]" metricMagnitude="5" metricUnit="m" metricValue="4.2" pageId="53" pageNumber="54" unit="km" value="420.0">420km</quantity>
NEE Pontianak
</locationDeviation>
, Ca.
<locationDeviation id="BCFD0079A866FFC1D6C4FD298CE3FD6B" box="[164,422,690,714]" pageId="53" pageNumber="54">
<quantity id="6A08B4BEA866FFC1D6C4FD298DA1FD6B" box="[164,228,690,714]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="53" pageNumber="54" unit="km" value="30.0">30km</quantity>
NNE Putussibau
</locationDeviation>
,
<geoCoordinate id="C8C47F9CA866FFC1D7CFFD298F46FD6B" box="[431,515,690,714]" degrees="1" direction="north" minutes="08" orientation="latitude" pageId="53" pageNumber="54" precision="925" value="1.1333333">1°08´N</geoCoordinate>
,
<geoCoordinate id="C8C47F9CA866FFC1D46EFD298F3FFD6B" box="[526,634,690,714]" degrees="113" direction="east" minutes="00" orientation="longitude" pageId="53" pageNumber="54" precision="925" value="113.0">113°00´E</geoCoordinate>
,
<date id="D94E3F9BA866FFC1D4E5FD298E7BFD6B" box="[645,830,690,714]" pageId="53" pageNumber="54" value="2008-12-23" valueMax="2008-12-28" valueMin="2008-12-23">
<collectingDate id="C90AC673A866FFC1D4E5FD298E7BFD6B" box="[645,830,690,714]" pageId="53" pageNumber="54" value="2008-12-23" valueMax="2008-12-28" valueMin="2008-12-23">23–28–XII–2008</collectingDate>
</date>
I.G.31.412. Leg
<collectorName id="00057C8DA866FFC1D619FD438DB6FD51" box="[121,243,728,752]" pageId="53" pageNumber="54">A.Napolov</collectorName>
“ (
<collectionCode id="CBE1819EA866FFC1D767FD438C12FD51" box="[263,343,728,752]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="53" pageNumber="54">IRSNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A866FFC3D619FCDC8F00FA81" lastPageId="55" lastPageNumber="56" pageId="53" pageNumber="54" type="description">
<paragraph id="AD4F195BA866FFC1D619FCDC8F18FCC1" blockId="53.[121,605,839,864]" box="[121,605,839,864]" pageId="53" pageNumber="54">
<emphasis id="9F84C549A866FFC1D619FCDC8DB9FCC1" bold="true" box="[121,252,839,864]" pageId="53" pageNumber="54">Description</emphasis>
(
<figureCitation id="35CB05DEA866FFC1D768FCD38C1EFCC1" box="[264,347,840,864]" captionStart="Figs 9–12" captionStartId="9.[121,158,1494,1512]" captionTargetBox="[145,998,148,1407]" captionTargetId="figure-60@9.[567,998,842,1405]" captionTargetPageId="9" captionText="Figs 9–12: Orictites spp., habitus. 9: O. barclayi sp. nov., holotype, female; 10: O. brancuccii sp. nov., holotype, male; 11: O. barclayi minorfrater ssp. nov., holotype, male; 12: O. tubercucollis sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284330" httpUri="https://zenodo.org/record/6284330/files/figure.png" pageId="53" pageNumber="54">Figs 12</figureCitation>
,
<figureCitation id="35CB05DEA866FFC1D708FCD38CC6FCC1" box="[360,387,840,864]" captionStart="Figs 27–30" captionStartId="24.[159,196,982,1000]" captionTargetBox="[159,1043,130,960]" captionText="Figs 27–30: Orictites spp., aedeagus, ventral view. 27: O. brancuccii sp. nov., holotype, scale bar 0.2 mm; 28: O. barclayi minorfrater ssp. nov., holotype, scale bar 0.2 mm; 29: O. tubercucollis sp. nov., holotype, scale bar 0.2 mm; 30: O. anteriorlatus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284354" httpUri="https://zenodo.org/record/6284354/files/figure.png" pageId="53" pageNumber="54">29</figureCitation>
, 39; pp. 10, 25, 37)
</paragraph>
<paragraph id="AD4F195BA866FFC1D619FC098FDBFC54" blockId="53.[121,1006,913,1573]" pageId="53" pageNumber="54">
Measurements (n = 9): Body length
<quantity id="6A08B4BEA866FFC1D474FC098FE8FC0B" box="[532,685,914,938]" metricMagnitude="-3" metricUnit="m" metricValue="5.91" metricValueMax="6.52" metricValueMin="5.3" pageId="53" pageNumber="54" unit="mm" value="5.91" valueMax="6.52" valueMin="5.3">5.3–6.52 mm</quantity>
, (
<emphasis id="9F84C549A866FFC1D4A1FC0A8F94FC0B" bold="true" box="[705,721,913,938]" italics="true" pageId="53" pageNumber="54"></emphasis>
=
<quantity id="6A08B4BEA866FFC1D490FC098E1DFC0B" box="[752,856,914,938]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" pageId="53" pageNumber="54" unit="mm" value="5.85">5.85 mm</quantity>
), width
<quantity id="6A08B4BEA866FFC1D5CEFC098D8AFC71" metricMagnitude="-3" metricUnit="m" metricValue="1.66" metricValueMax="1.9" metricValueMin="1.42" pageId="53" pageNumber="54" unit="mm" value="1.66" valueMax="1.9" valueMin="1.42">1.42– 1.9 mm</quantity>
(
<emphasis id="9F84C549A866FFC1D6BFFC2C8DAAFC71" bold="true" box="[223,239,951,976]" italics="true" pageId="53" pageNumber="54"></emphasis>
=
<quantity id="6A08B4BEA866FFC1D771FC238C3CFC71" box="[273,377,952,976]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="53" pageNumber="54" unit="mm" value="1.67">1.67 mm</quantity>
), ratio length/width of pronotum 0.93–1.01 (
<emphasis id="9F84C549A866FFC1D51AFC2C8ECFFC71" bold="true" box="[890,906,951,976]" italics="true" pageId="53" pageNumber="54"></emphasis>
= 0.97), ratio length/width of elytra 1.74–1.93 (
<emphasis id="9F84C549A866FFC1D44FFC478F7AFC54" bold="true" box="[559,575,988,1013]" italics="true" pageId="53" pageNumber="54"></emphasis>
= 1.88).
</paragraph>
<paragraph id="AD4F195BA866FFC1D6FEFB998FB3FBC4" blockId="53.[121,1006,913,1573]" pageId="53" pageNumber="54">
Colour: Shiny. Black to dark-brown. Wings of clypeus and supraantennal plates dark-brown, slightly transparent in
<specimenCount id="BBF6D2D2A866FFC1D435FBB38FB7FBE1" box="[597,754,1064,1088]" pageId="53" pageNumber="54" type="generic" typeStatus="paratype">one paratype</specimenCount>
, antennae and labrum dark-brown, palpi yellowish brown, legs dark red-brown.
</paragraph>
<paragraph id="AD4F195BA866FFC2D6FEFBE98EAFFBBB" blockId="53.[121,1006,913,1573]" lastBlockId="54.[159,1044,93,1573]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
Head: A fifth narrower than pronotum. Clypeus with small rounded tooth laterally, nearly fused with wings, anterior margin slightly excised, with blunt raised knob between the two clypeal setae, anteriorly with some rugae, posterior part of knob covered with big punctures and rugae, margin of wings convex, slightly hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, smooth, with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by flat transverse depression and transverse rugae, with costate, nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina, frons moderately convex, with very minute and scattered punctures. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes of moderate size, flattened but still convex, genae enlarged, regularly rounded, not as high as and shorter than eyes, enclosing eyes posteriorly, forming obtuse angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as distinct step, with bigger punctures connected line-like, neck laterally covered with reticulation and densely scattered medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), and both somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A865FFC2D44CFDFF8F06FDD6" attach="left" box="[556,579,612,631]" fontSize="8" pageId="54" pageNumber="55">nd</superScript>
segment, 2
<superScript id="5A85B413A865FFC2D4AAFDFF8FA4FDD6" attach="left" box="[714,737,612,631]" fontSize="8" pageId="54" pageNumber="55">nd</superScript>
segment bisetose. Ventral surface of neck covered with punctures and some reticulation. Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming broad obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae of moderate length, reaching distinctly over middle of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 subelongate (L/
<geoCoordinate id="C8C47F9CA865FFC2D694FC468C7CFC54" box="[244,313,989,1013]" degrees="1. 4" direction="west" orientation="longitude" pageId="54" pageNumber="55" precision="5555" value="-1.4">W 1.4</geoCoordinate>
), antennomeres slightly flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA865FFC3D6A4FBB38F52FF61" blockId="54.[159,1044,93,1573]" lastBlockId="55.[121,1006,93,1573]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">Pronotum: Square, thought slightly transverse, sides slightly convex at middle, distinctly rounded in anterior quarter to anterior angles. Anterior margin slightly bisinuate. Reflexed lateral margin with some fine scars in anterior part. Margin from posterior angle to base running as slightly convex line, with a distinct tubercle bilaterally at base. Lateral channel conspicuously broad between the two lateral setigerous punctures, with isodiametric reticulation and very few irregular punctures. Anterior setigerous puncture located at the end of anterior fifth adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles projecting, rounded, formed by reflexed lateral margin, posterior angle developed as sharp distinct tooth, projecting antero-laterally. Anterior transverse line formed by distinct, partly connected punctures. Median line deep and conspicuously broad, rugose, running up to level of anterior transverse line but not joining, smaller at base, adjoining base. Surface irregularly covered with big punctures and with irregular reticulation laterally and at base, basal impression short but well visible, basal channel broad, deep. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA864FFC3D6FEFF568FD2FD21" blockId="55.[121,1006,93,1573]" pageId="55" pageNumber="56">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles. Reflexed margin with fine crenulation in anterior third, margin slightly thickened at humerus, bending with channel over rounded humerus up to 5
<superScript id="5A85B413A864FFC3D757FEC48C0FFED3" attach="left" box="[311,330,351,370]" fontSize="8" pageId="55" pageNumber="56">th</superScript>
stria. Humerus with distinct rounded tooth. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1
<superScript id="5A85B413A864FFC3D57EFE1F8E6AFE36" attach="left" box="[798,815,388,407]" fontSize="8" pageId="55" pageNumber="56">st</superScript>
stria. Striae 1–4 free at base, all striae deep, distinctly punctate, striae 1 and 2 ending free at apex, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura costiform, raised. All intervals conspicuously convex, thought costate, 3
<superScript id="5A85B413A864FFC3D42CFD818F1AFD8C" attach="left" box="[588,607,538,557]" fontSize="8" pageId="55" pageNumber="56">rd</superScript>
and4
<superScript id="5A85B413A864FFC3D4FEFD818FF4FD8C" attach="left" box="[670,689,538,557]" fontSize="8" pageId="55" pageNumber="56">th</superScript>
broader and raised at base, both with tubercle at base, 8
<superScript id="5A85B413A864FFC3D7A6FDA48C9CFDF3" attach="left" box="[454,473,575,594]" fontSize="8" pageId="55" pageNumber="56">th</superScript>
carinate. All intervals shiny on disc. Interval 3 with four setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA864FFC3D6FEFD168CA6FD04" blockId="55.[121,1006,93,1573]" box="[158,483,653,677]" pageId="55" pageNumber="56">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA864FFC3D6FEFD298ED3FC0B" blockId="55.[121,1006,93,1573]" pageId="55" pageNumber="56">
Ventral surface: Epipleuron shiny, covered with big scattered punctures and indistinct reticulation, submarginal furrow distinct. Proepisternum and episternum with punctures, prosternite slightly punctured, keeled at middle, mesosternum nearly smooth. Sternites with transverse to isodiametric reticulation, each with big punctures at base, denser laterally, 3
<superScript id="5A85B413A864FFC3D539FCDF8E29FCF6" attach="left" box="[857,876,836,855]" fontSize="8" pageId="55" pageNumber="56">rd</superScript>
to 5
<superScript id="5A85B413A864FFC3D5C1FCDF8EF1FCF6" attach="left" box="[929,948,836,855]" fontSize="8" pageId="55" pageNumber="56">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA864FFC3D6FEFC238C51FBC4" blockId="55.[121,1006,93,1573]" pageId="55" pageNumber="56">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA864FFC3D6FEFBE98F00FA81" blockId="55.[121,1006,93,1573]" pageId="55" pageNumber="56">
Male genitalia (
<figureCitation id="35CB05DEA864FFC3D735FBE98CE2FB2B" box="[341,423,1138,1162]" captionStart="Figs 27–30" captionStartId="24.[159,196,982,1000]" captionTargetBox="[159,1043,130,960]" captionText="Figs 27–30: Orictites spp., aedeagus, ventral view. 27: O. brancuccii sp. nov., holotype, scale bar 0.2 mm; 28: O. barclayi minorfrater ssp. nov., holotype, scale bar 0.2 mm; 29: O. tubercucollis sp. nov., holotype, scale bar 0.2 mm; 30: O. anteriorlatus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284354" httpUri="https://zenodo.org/record/6284354/files/figure.png" pageId="55" pageNumber="56">Fig. 29</figureCitation>
): Relatively long, moderately sclerotized. Median lobe slender, moderately arcuate, spatula slightly distorted and broadened. Endophallus strongly folded, teeth not visible. Parameres of about same length, both slender, longer than median lobe (in natural position), slightly twisted, with minute pili at the apical tip.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A864FFCCD6FEFAB68942FF44" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="discussion">
<paragraph id="AD4F195BA864FFC3D6FEFAB68CB9FA31" blockId="55.[121,1006,93,1573]" pageId="55" pageNumber="56">Remarks: The minute setae on the apical tip of the parameres are sensitive and tend to break during mounting. The setigerous punctures are minute so that high magnification is needed.</paragraph>
<paragraph id="AD4F195BA864FFCCD6FEFA068C7AFF3B" blockId="55.[121,1006,93,1573]" lastBlockId="56.[159,1044,93,640]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
Female genitalia (
<figureCitation id="35CB05DEA864FFC3D707FA068CF1FA14" box="[359,436,1437,1461]" captionStart="Figs 39–40" captionStartId="36.[159,196,705,723]" captionTargetBox="[159,1032,80,662]" captionTargetId="figure-227@36.[159,590,115,627]" captionTargetPageId="36" captionText="Figs 39–40: Orictites spp., coxostylus. 39: O. tubercucollis sp. nov., scale bar 0.1 mm; 40: O. charleshuberi sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284374" httpUri="https://zenodo.org/record/6284374/files/figure.png" pageId="55" pageNumber="56">Fig. 39</figureCitation>
): Coxostylus slender, slightly sigmoid, dorso-ventrally moderately depressed, gently curved to apex, with obtuse knob-like tooth laterally at base, with two setae apically close together, with one short and blunt robust ensiform seta, and one acute short ensiform seta, both of them in middle part, with four nematiform setae in the middle part and towards base.
</paragraph>
<paragraph id="AD4F195BA86BFFCCD6A4FF338942FF44" blockId="56.[159,1044,93,640]" pageId="56" pageNumber="57">Variation: Besides the total length, slight differences in coloration, and the degree in punctation on the ventral surface, there was no variation observed.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A86BFFCCD6A4FF698CA4FE4B" pageId="56" pageNumber="57" type="diagnosis">
<paragraph id="AD4F195BA86BFFCCD6A4FF698CA4FE4B" blockId="56.[159,1044,93,640]" pageId="56" pageNumber="57">Diagnosis: A medium-sized black species with distinct knob on the clypeus and distinct flat transverse depression with rugae behind, with distinct but rounded humeral tooth, tubercles at the base of intervals 3 and 4 of the elytra, and four setigerous punctures on interval 3. In addition, the female coxostylus exhibits two ensiform setae in the middle part. Distinguished from all other species by the presence of a bilateral tubercle on the reflexed margin at the base of the pronotum.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A86BFFCCD6A4FE638E9EFDB1" box="[196,987,504,528]" pageId="56" pageNumber="57" type="distribution">
<paragraph id="AD4F195BA86BFFCCD6A4FE638E9EFDB1" blockId="56.[159,1044,93,640]" box="[196,987,504,528]" pageId="56" pageNumber="57">
Distribution. The species was collected in
<collectingCountry id="D5E759CBA86BFFCCD4FEFE638E41FDB1" box="[670,772,504,528]" name="Thailand" pageId="56" pageNumber="57">Thailand</collectingCountry>
,
<collectingCountry id="D5E759CBA86BFFCCD56DFE638E02FDB1" box="[781,839,504,528]" name="Laos" pageId="56" pageNumber="57">Laos</collectingCountry>
, and Borneo.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A86BFFCCD6A4FD868E15FD94" box="[196,848,541,565]" pageId="56" pageNumber="57" type="biology_ecology">
<paragraph id="AD4F195BA86BFFCCD6A4FD868E15FD94" blockId="56.[159,1044,93,640]" box="[196,848,541,565]" pageId="56" pageNumber="57">Collecting altitudes were between 1000 and 1500 metres.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A86BFFCCD6A4FDD98CDFFD21" pageId="56" pageNumber="57" type="etymology">
<paragraph id="AD4F195BA86BFFCCD6A4FDD98CDFFD21" blockId="56.[159,1044,93,640]" pageId="56" pageNumber="57">Etymology: The name refers to the tubercles on the reflexed margin at the base of the pronotum.</paragraph>
</subSubSection>
</treatment>
</document>

159
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<taxonomicName id="6AF062D8A868FFCFD619FD168C31FD07" ID-CoL="8MJVP" authorityName="Balkenohl" authorityYear="2017" box="[121,372,653,678]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="anteriorlatus" status="sp. nov.">
<emphasis id="9F84C549A868FFCFD619FD168C31FD07" bold="true" box="[121,372,653,678]" italics="true" pageId="59" pageNumber="60">Orictites anteriorlatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A868FFCFD71AFD168C97FD04" box="[378,466,653,677]" pageId="59" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="E5EA4AD0A868FFCFD619FD438CC6FC9B" pageId="59" pageNumber="60" type="materials_examined">
<paragraph id="AD4F195BA868FFCFD619FD438CC6FC9B" blockId="59.[121,1005,728,864]" pageId="59" pageNumber="60">
Type material:
<materialsCitation id="1D981306A868FFCFD74CFD438CC6FC9B" ID-GBIF-Occurrence="3497007314" collectingDate="1999-12-25" collectionCode="CBM-ZSM" location="Mt. Lompo Battang" pageId="59" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<typeStatus id="724BA7F9A868FFCFD74CFD438CDFFD51" box="[300,410,728,752]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A868FFCFD7C9FD438CF8FD4E" box="[425,445,728,751]" pageId="59" pageNumber="60" type="male">♂</specimenCount>
, labels: white, black printed: “Mt. Lompo Battang S.C. Sulawesi
<date id="D94E3F9BA868FFCFD73CFD668CA9FCB4" box="[348,492,765,789]" pageId="59" pageNumber="60" value="1999-12-25">
<collectingDate id="C90AC673A868FFCFD73CFD668CA9FCB4" box="[348,492,765,789]" pageId="59" pageNumber="60" value="1999-12-25">25–XII–1999</collectingDate>
</date>
M. ANDO” / blue, black printed: “K. ANDO Collection” (
<collectionCode id="CBE1819EA868FFCFD762FCB98C3FFC9B" box="[258,378,802,826]" pageId="59" pageNumber="60">CBM-ZSM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
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<paragraph id="AD4F195BA868FFCFD619FCD38F8BFCC1" blockId="59.[121,1005,728,864]" box="[121,718,840,864]" pageId="59" pageNumber="60">
Remark: In the
<typeStatus id="724BA7F9A868FFCFD743FCD38CCDFCC1" box="[291,392,840,864]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
the left hind tibia is missing.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A868FFC9D619FC2C8F3DFB2B" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60" type="description">
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<emphasis id="9F84C549A868FFCFD619FC2C8DB9FC71" bold="true" box="[121,252,951,976]" pageId="59" pageNumber="60">Description</emphasis>
(
<figureCitation id="35CB05DEA868FFCFD768FC238C1EFC71" box="[264,347,952,976]" captionStart="Figs 13–14" captionStartId="10.[159,196,801,819]" captionTargetBox="[159,1036,143,748]" captionTargetPageId="10" captionText="Figs 13–14: Orictites spp., habitus. 13: O. charleshuberi sp. nov., holotype, female; 14: O. anteriorlatus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284334" httpUri="https://zenodo.org/record/6284334/files/figure.png" pageId="59" pageNumber="60">Figs 14</figureCitation>
,
<figureCitation id="35CB05DEA868FFCFD708FC238CCCFC71" box="[360,393,952,976]" captionStart="Figs 27–30" captionStartId="24.[159,196,982,1000]" captionTargetBox="[159,1043,130,960]" captionText="Figs 27–30: Orictites spp., aedeagus, ventral view. 27: O. brancuccii sp. nov., holotype, scale bar 0.2 mm; 28: O. barclayi minorfrater ssp. nov., holotype, scale bar 0.2 mm; 29: O. tubercucollis sp. nov., holotype, scale bar 0.2 mm; 30: O. anteriorlatus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284354" httpUri="https://zenodo.org/record/6284354/files/figure.png" pageId="59" pageNumber="60">30</figureCitation>
; pp. 11, 25)
</paragraph>
<paragraph id="AD4F195BA868FFCFD619FB998F3FFBE1" blockId="59.[121,1006,1026,1573]" pageId="59" pageNumber="60">
Measurements: Body length
<quantity id="6A08B4BEA868FFCFD7A1FB998F5CFBBB" box="[449,537,1026,1050]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="59" pageNumber="60" unit="mm" value="5.8">5.8 mm</quantity>
, width
<quantity id="6A08B4BEA868FFCFD40AFB998F87FBBB" box="[618,706,1026,1050]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="59" pageNumber="60" unit="mm" value="1.6">1.6 mm</quantity>
; ratio length/width of pronotum 0.99; ratio length/width of elytra 1.94.
</paragraph>
<paragraph id="AD4F195BA868FFCFD6FEFBD68F73FB11" blockId="59.[121,1006,1026,1573]" pageId="59" pageNumber="60">Colour: Shiny. Black to dark-brown. Wings of clypeus and supraantennal plates brown and slightly transparent, antennae, labrum and mandibles brown, palpi yellowish, legs red-brown.</paragraph>
<paragraph id="AD4F195BA868FFC8D6FEFB268FD4FC71" blockId="59.[121,1006,1026,1573]" lastBlockId="60.[159,1044,93,1573]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
Head: Wide, a sixth narrower than pronotum. Clypeus with small rounded indistinctly projecting tooth laterally, separated from wings, anterior margin excised, with blunt raised broad knob between the two clypeal setae, margin of wings convex, hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, smooth, with reflexed rounded margin, elongated as supraorbital carina. Frons separated from clypeus by transverse furrow interrupted at middle, sharp furrow between frons and frontal carina, frons moderately convex. Frons and clypeus smooth. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle behind genae level. Eyes well developed, convex, genae distinct, regularly rounded, not as high as eyes, enclosing eyes posteriorly by one third, forming nearly right angle at neck. Grooves for reception of antennae of moderate length, 0.5 of eye diameter. Neck constriction distinct, with band of big punctures. Neck laterally covered with reticulation and some medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with fine isodiametric reticulation. Mandibles robust, length three quarters of head, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles rounded towards base. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A86FFFC8D539FE6F8E35FDA6" attach="left" box="[857,880,500,519]" fontSize="8" pageId="60" pageNumber="61">nd</superScript>
segment, 2
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segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with punctures and reticulation. Submentum and mentum separated; submentum with four, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae of moderate length, reaching up to middle of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine isodiametric reticulation, segments 5–10 transverse (L/
<geoCoordinate id="C8C47F9CA86FFFC8D496FCF68E14FC24" box="[758,849,877,901]" degrees="0. 73" direction="west" orientation="longitude" pageId="60" pageNumber="61" precision="555" value="-0.73">W 0.73</geoCoordinate>
), antennomeres slightly flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA86FFFC8D6A4FC468EEEF984" blockId="60.[159,1044,93,1573]" pageId="60" pageNumber="61">Pronotum: Square, wider than long, outline reminescent of the inverted frustum of a pyramid, sides straight, convex in anterior quarter for a short distance to anterior angles. Anterior margin distinctly bisinuate. Reflexed lateral margin with traces of scars (120x). Margin from posterior angle to base running as a straight line. Lateral channel disctinctly broadened between the two lateral setigerous punctures, with isodiametric reticulation and few punctures. Anterior setigerous puncture located at the end of anterior fifth adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles distinctly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as blunt tooth. Anterior transverse line narrow, sharp, formed by punctures. Median line deep, relatively narrow, rugose, surpassing anterior transverse line but not joining, joining base. Surface with a few punctures and transverse rugae, basal impression indistinct. Basal channel moderately broad, deep. Basal part of disc elongated posteriorly and bilaterally at declivity as tooth-like vault slightly hanging over basal channel (lateral view). Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA86EFFC9D6FEFFC68F69FE4B" blockId="61.[121,1006,93,1536]" pageId="61" pageNumber="62">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles and with a second row of punctures. Reflexed margin with a few indistinct scars posterior to humerus (120x). Margin bending with channel over rounded humerus up to 5
<superScript id="5A85B413A86EFFC9D458FF748F0EFEA3" attach="left" box="[568,587,239,258]" fontSize="8" pageId="61" pageNumber="62">th</superScript>
stria. Humerus without tooth. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1
<superScript id="5A85B413A86EFFC9D5F8FE8F8EECFE86" attach="left" box="[920,937,276,295]" fontSize="8" pageId="61" pageNumber="62">st</superScript>
stria. Striae 1–4 free at base, all striae deep, indistinctly punctate, striae 1–2, 3–4, and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct. Intervals distinctly convex, 3
<superScript id="5A85B413A86EFFC9D7B1FE1F8CA1FE36" attach="left" box="[465,484,388,407]" fontSize="8" pageId="61" pageNumber="62">rd</superScript>
and4
<superScript id="5A85B413A86EFFC9D449FE1F8F79FE36" attach="left" box="[553,572,388,407]" fontSize="8" pageId="61" pageNumber="62">th</superScript>
distinctly raised at base, both with tubercle at base, 8
<superScript id="5A85B413A86EFFC9D732FE318C20FE1C" attach="left" box="[338,357,426,445]" fontSize="8" pageId="61" pageNumber="62">th</superScript>
carinate. All intervals shiny on disc. Interval 3 with four setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA86EFFC9D6FEFE638CA6FDB1" blockId="61.[121,1006,93,1536]" box="[158,483,504,528]" pageId="61" pageNumber="62">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA86EFFC9D6FEFD868DAEFCB4" blockId="61.[121,1006,93,1536]" pageId="61" pageNumber="62">
Ventral surface: Epipleuron shiny, with some punctures, submarginal furrow distinct, with transverse rugae. Proepisternum and episternum with a few punctures, prosternite nearly smooth, keeled at middle, mesosternum smooth. Sternites with few punctures laterally, 3
<superScript id="5A85B413A86EFFC9D4A5FD118F9DFD3C" attach="left" box="[709,728,650,669]" fontSize="8" pageId="61" pageNumber="62">rd</superScript>
to 5
<superScript id="5A85B413A86EFFC9D574FD118E62FD3C" attach="left" box="[788,807,650,669]" fontSize="8" pageId="61" pageNumber="62">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, with transverse reticulation, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA86EFFC9D6FEFCB98E8EFC0B" blockId="61.[121,1006,93,1536]" pageId="61" pageNumber="62">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA86EFFC9D6FEFC238CD1FBE1" blockId="61.[121,1006,93,1536]" pageId="61" pageNumber="62">
Male genitalia (
<figureCitation id="35CB05DEA86EFFC9D72DFC238CDEFC71" box="[333,411,952,976]" captionStart="Figs 27–30" captionStartId="24.[159,196,982,1000]" captionTargetBox="[159,1043,130,960]" captionText="Figs 27–30: Orictites spp., aedeagus, ventral view. 27: O. brancuccii sp. nov., holotype, scale bar 0.2 mm; 28: O. barclayi minorfrater ssp. nov., holotype, scale bar 0.2 mm; 29: O. tubercucollis sp. nov., holotype, scale bar 0.2 mm; 30: O. anteriorlatus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284354" httpUri="https://zenodo.org/record/6284354/files/figure.png" pageId="61" pageNumber="62">Fig. 30</figureCitation>
): Relatively long, distinctly sclerotized. Median lobe slender, moderately arcuate, slightly twisted apically. Endophallus strongly folded, teeth not visible. Parameres of about equal length, both slender, slightly twisted, asetose.
</paragraph>
<paragraph id="AD4F195BA86EFFC9D6FEFBD68C9DFBC4" blockId="61.[121,1006,93,1536]" box="[158,472,1101,1125]" pageId="61" pageNumber="62">Female genitalia: Unknown.</paragraph>
<paragraph id="AD4F195BA86EFFC9D6FEFBE98F3DFB2B" blockId="61.[121,1006,93,1536]" box="[158,632,1138,1162]" pageId="61" pageNumber="62">Variation: No bilateral variation observed.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A86EFFC9D6FEFB038FD0FA14" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="AD4F195BA86EFFC9D6FEFB038FD0FA14" blockId="61.[121,1006,93,1536]" pageId="61" pageNumber="62">
Diagnosis: A medium-sized almost black species, with blunt broad knob on the clypeus, humerus without tooth, and four setigerous punctures on interval 3 of the elytra. Only in this species and in
<taxonomicName id="6AF062D8A86EFFC9D4EFFB798E61FB5B" authorityName="LouWERENS" authorityYear="1964" box="[655,804,1250,1274]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A86EFFC9D4EFFB798E61FB5B" box="[655,804,1250,1274]" italics="true" pageId="61" pageNumber="62">O. mjoebergi</emphasis>
</taxonomicName>
the basal part of the disc of the pronotum is posteriorly elongated bilaterally at the declivity as a tooth-like vault slightly hanging over the basal channel. In contrast to
<taxonomicName id="6AF062D8A86EFFC9D5B5FAB68DA9FACB" authorityName="LouWERENS" authorityYear="1964" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="9F84C549A86EFFC9D5B5FAB68DA9FACB" italics="true" pageId="61" pageNumber="62">O. mjoebergi</emphasis>
</taxonomicName>
,
<taxonomicName id="6AF062D8A86EFFC9D69BFAC98CE8FACB" authorityName="Balkenohl" authorityYear="2017" box="[251,429,1362,1386]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="anteriorlatus" status="sp. nov.">
<emphasis id="9F84C549A86EFFC9D69BFAC98CE8FACB" box="[251,429,1362,1386]" italics="true" pageId="61" pageNumber="62">O. anteriorlatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A86EFFC9D7D4FAC98F48FACB" box="[436,525,1362,1386]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
exhibits tubercles at the base of intervals 3 and 4 of the elytra, the furrow between clypeus and frons is interrupted at middle, and clypeus and frons are impunctured.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A86EFFC9D6FEFA598FFAFA7B" box="[158,703,1474,1498]" pageId="61" pageNumber="62" type="distribution">
<paragraph id="AD4F195BA86EFFC9D6FEFA598FFAFA7B" blockId="61.[121,1006,93,1536]" box="[158,703,1474,1498]" pageId="61" pageNumber="62">
Distribution: Known from the South of
<collectingRegion id="6F34D7B9A86EFFC9D432FA598FFEFA7B" box="[594,699,1474,1498]" country="Indonesia" name="Sulawesi Utara" pageId="61" pageNumber="62">Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A86EFFC9D6FEFA738E3CF9A1" box="[158,889,1512,1536]" pageId="61" pageNumber="62" type="etymology">
<paragraph id="AD4F195BA86EFFC9D6FEFA738E3CF9A1" blockId="61.[121,1006,93,1536]" box="[158,889,1512,1536]" pageId="61" pageNumber="62">Etymology: The name refers to the anteriorly widened pronotum.</paragraph>
</subSubSection>
</treatment>
</document>

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<emphasis id="9F84C549A86BFFCCD6FFFD4C8CE6FD51" bold="true" box="[159,419,727,752]" italics="true" pageId="56" pageNumber="57">Orictites charleshuberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A86BFFCCD7CAFD438F44FD51" box="[426,513,728,752]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="E5EA4AD0A86BFFCCD6FFFCB98FA1FC54" pageId="56" pageNumber="57" type="materials_examined">
<paragraph id="AD4F195BA86BFFCCD6FFFCB98EA0FC0B" blockId="56.[159,1044,802,1050]" pageId="56" pageNumber="57">
Type material:
<materialsCitation id="1D981306A86BFFCCD72DFCB98EA0FC0B" ID-GBIF-Occurrence="3497007345" collectingDate="1938-05-10" collectionCode="BMNH" collectorName="H. E. Andrewes" country="Taiwan" elevation="950" location="Karapin Near Ari Formosa" pageId="56" pageNumber="57" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="724BA7F9A86BFFCCD72DFCB98CFFFC9B" box="[333,442,802,826]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A86BFFCCD7A7FCB98CB6FC9B" box="[455,499,802,826]" pageId="56" pageNumber="57" type="female">1 ♀</specimenCount>
, labels: beige, black printed and handwritten: “
<location id="A82F4F80A86BFFCCD6C7FCD38C95FCC1" LSID="urn:lsid:plazi:treatment:2559A84DA86BFFCFD6FFFD4C8DEEFD94:A82F4F80A86BFFCCD6C7FCD38C95FCC1" box="[167,464,840,864]" country="Taiwan" name="Karapin Near Ari Formosa" pageId="56" pageNumber="57">Karapin Near Ari Formosa</location>
.
<date id="D94E3F9BA86BFFCCD7B9FCD38F17FCC1" box="[473,594,840,864]" pageId="56" pageNumber="57" value="1938-05-10">
<collectingDate id="C90AC673A86BFFCCD7B9FCD38F17FCC1" box="[473,594,840,864]" pageId="56" pageNumber="57" value="1938-05-10">10–V 1938</collectingDate>
</date>
Coll. Yoshio Yano” / beige, handwritten in black ink: “Karapin Mt. Ari (
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.)” / beige, black printed: “
<collectorName id="00057C8DA86BFFCCD510FCF68956FC24" box="[880,1043,877,901]" pageId="56" pageNumber="57">
H.E.
<taxonomicName id="6AF062D8A86BFFCCD5F6FCF68956FC24" box="[918,1043,877,901]" pageId="56" pageNumber="1" rank="subGenus" status="subgen. nov." subGenus="Andrewes">Andrewes</taxonomicName>
</collectorName>
Coll. B.M.1945—97.” / backside, handwritten in black ink: “2541” (
<collectionCode id="CBE1819EA86BFFCCD5EBFC098E9EFC0B" box="[907,987,914,938]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="56" pageNumber="57">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="AD4F195BA86BFFCCD6FFFC238FA1FC54" blockId="56.[159,1044,802,1050]" pageId="56" pageNumber="57">
<materialsCitation id="1D981306A86BFFCCD6FFFC238FA5FC54" ID-GBIF-Occurrence="3497007342" collectingDate="1993-06-06" collectingDateMax="1993-06-26" collectingDateMin="1993-06-06" collectionCode="CBP" collectorName="J. DALIHOD" country="Taiwan" location="YUCHIH SUN MOON LAIE" pageId="56" pageNumber="57" specimenCode="LAIE 16" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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:
<specimenCount id="BBF6D2D2A86BFFCCD775FC238C05FC6E" box="[277,320,952,976]" pageId="56" pageNumber="57" type="female">1 ♀</specimenCount>
, labels: white, printed “TAIWAN P.o.CHINA PULI –
<location id="A82F4F80A86BFFCCD5E2FC238C64FC54" LSID="urn:lsid:plazi:treatment:2559A84DA86BFFCFD6FFFD4C8DEEFD94:A82F4F80A86BFFCCD5E2FC238C64FC54" country="Taiwan" name="YUCHIH SUN MOON LAIE" pageId="56" pageNumber="57">YÜCHIH SUN MOON LAIE</location>
<collectingDate id="C90AC673A86BFFCCD747FC468C9AFC54" box="[295,479,989,1013]" pageId="56" pageNumber="57">
<date id="D94E3F9BA86BFFCCD747FC468CCCFC54" box="[295,393,989,1013]" pageId="56" pageNumber="57" value="1926-06-16">16.6.–26</date>
.6.1993
</collectingDate>
<collectorName id="00057C8DA86BFFCCD785FC468F24FC54" box="[485,609,989,1013]" pageId="56" pageNumber="57">J. DALIHOD</collectorName>
lgt.“ (
<collectionCode id="CBE1819EA86BFFCCD4C4FC468F9FFC54" box="[676,730,989,1013]" pageId="56" pageNumber="57">CBP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A86BFFCCD6FFFB99894AFBBB" box="[159,1039,1026,1050]" pageId="56" pageNumber="57" type="discussion">
<paragraph id="AD4F195BA86BFFCCD6FFFB99894AFBBB" blockId="56.[159,1044,802,1050]" box="[159,1039,1026,1050]" pageId="56" pageNumber="57">
Remark: In the
<typeStatus id="724BA7F9A86BFFCCD729FB998CEAFBBB" box="[329,431,1026,1050]" pageId="56" pageNumber="57" type="paratype">paratype</typeStatus>
the right antennae is missing from joint two onwards.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A86BFFCED6FFFBE98FB1F984" lastPageId="58" lastPageNumber="59" pageId="56" pageNumber="57" type="description">
<paragraph id="AD4F195BA86BFFCCD6FFFBE98F74FB2B" blockId="56.[159,561,1138,1163]" box="[159,561,1138,1163]" pageId="56" pageNumber="57">
<emphasis id="9F84C549A86BFFCCD6FFFBE98C67FB2A" bold="true" box="[159,290,1138,1163]" pageId="56" pageNumber="57">Description</emphasis>
(
<figureCitation id="35CB05DEA86BFFCCD74EFBE98CC4FB2B" box="[302,385,1138,1162]" captionStart="Figs 13–14" captionStartId="10.[159,196,801,819]" captionTargetBox="[159,1036,143,748]" captionTargetPageId="10" captionText="Figs 13–14: Orictites spp., habitus. 13: O. charleshuberi sp. nov., holotype, female; 14: O. anteriorlatus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284334" httpUri="https://zenodo.org/record/6284334/files/figure.png" pageId="56" pageNumber="57">Figs 13</figureCitation>
,
<figureCitation id="35CB05DEA86BFFCCD7EEFBE98CF5FB2B" box="[398,432,1138,1162]" captionStart="Figs 39–40" captionStartId="36.[159,196,705,723]" captionTargetBox="[159,1032,80,662]" captionTargetId="figure-227@36.[159,590,115,627]" captionTargetPageId="36" captionText="Figs 39–40: Orictites spp., coxostylus. 39: O. tubercucollis sp. nov., scale bar 0.1 mm; 40: O. charleshuberi sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284374" httpUri="https://zenodo.org/record/6284374/files/figure.png" pageId="56" pageNumber="57">40</figureCitation>
; pp. 11, 37)
</paragraph>
<paragraph id="AD4F195BA86BFFCCD6FFFB268EB0FB5B" blockId="56.[159,1044,1213,1573]" pageId="56" pageNumber="57">
Measurements (n = 2): Body length 5.8/
<quantity id="6A08B4BEA86BFFCCD419FB268F91FB74" box="[633,724,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="56" pageNumber="57" unit="mm" value="5.8">5.8 mm</quantity>
, width 1.65/
<quantity id="6A08B4BEA86BFFCCD506FB268E94FB74" box="[870,977,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="56" pageNumber="57" unit="mm" value="1.68">1.68 mm</quantity>
; ratio length/width of pronotum 0.98/0.95; ratio length/width of elytra 1.84/1.86.
</paragraph>
<paragraph id="AD4F195BA86BFFCCD6A4FA938F11FACB" blockId="56.[159,1044,1213,1573]" pageId="56" pageNumber="57">Colour: Shiny. Black to dark-brown. Wings of clypeus, supraantennal plates and labrum dark-brown, slightly transparent at margins, antennae brown, palpi yellowish brown, legs red-brown.</paragraph>
<paragraph id="AD4F195BA86BFFCDD6A4FAE38E43FA31" blockId="56.[159,1044,1213,1573]" lastBlockId="57.[121,1006,93,1573]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
Head: A quarter narrower than pronotum. Clypeus nearly fused with wings, with small rounded tooth laterally, anterior margin slightly excised, with blunt raised knob between the two clypeal seatae, margin of wings convex, slightly hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, shiny but with indistinct irregular reticulation, with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by flat transverse depression, depression with broad band of big punctures, elongated V-like on frons, with costate, nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina. Frons moderately convex, with very few punctures. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes of moderate size, flattened but still convex, genae enlarged, regularly rounded, not as high as and shorter than eyes, enclosing eyes posteriorly, forming rectangular angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as distinct step, with broader band of big punctures, interrupted at middle. Neck laterally covered with reticulation and densely scattered with punctures of medium size. Labrum indistinctly trilobed, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, irregularly reticulated, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A86AFFCDD74DFC148C01FC03" attach="left" box="[301,324,911,930]" fontSize="8" pageId="57" pageNumber="58">nd</superScript>
segment, 2
<superScript id="5A85B413A86AFFCDD7ACFC148CA6FC03" attach="left" box="[460,483,911,930]" fontSize="8" pageId="57" pageNumber="58">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck covered with bigger sized punctures and transverse reticulation, Submentum and mentum distinctly separated; submentum with four bigger, about equally spaced setigerous punctures; mentum at base bilaterally with broad and rounded tubercle, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming broad obtuse triangle, bisetose, not projecting as far as lateral lobes. Paragenae sharp, carina-like, with distinct tooth anteriorly, rounded at base. Antennae relatively long, reaching up to beginning of posterior third of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with isodiametric reticulation, segments 5–10 moniliform, antennomeres somewhat flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA86AFFCED6FEFA068F78FDB1" blockId="57.[121,1006,93,1573]" lastBlockId="58.[159,1044,93,1573]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">Pronotum: Square, nearly as long as wide, sides slightly convex at middle, rounded in anterior tenth to anterior angles. Anterior margin nearly straight. Reflexed lateral margin distinctly crenulated. Margin from posterior angle to base running as straight line, with small bilateral tubercle at base. Lateral channel broad between the two lateral setigerous punctures, with isodiametric reticulation and some punctures. Anterior setigerous puncture located at the end of anterior quarter adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles slightly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as distinct tooth, projecting antero-laterally. Anterior transverse line relatively fine, formed by partly connected punctures. Median line moderately deep, reticulated, surpassing anterior transverse line without joining, smaller at base, joining base. Surface with smaller and bigger punctures, with isodiametric reticulation laterally and at base, basal impression indistinct, basal channel distinct, broad and deep. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA869FFCED6A4FD868FC3FC71" blockId="58.[159,1044,93,1573]" pageId="58" pageNumber="59">
Elytra: Subcylindrical. Sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles. Reflexed margin with distinct crenulation in anterior half, margin thickened at humerus, bending with channel over rounded humerus up to 5
<superScript id="5A85B413A869FFCED738FD348C2EFD63" attach="left" box="[344,363,687,706]" fontSize="8" pageId="58" pageNumber="59">th</superScript>
stria. Humeral tooth big, sharp. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1
<superScript id="5A85B413A869FFCED4CAFD4F8FFEFD46" attach="left" box="[682,699,724,743]" fontSize="8" pageId="58" pageNumber="59">st</superScript>
stria. Striae 1–4 free at base, striae deep, punctate-striate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura somewhat raised, reticulated. Intervals moderately convex, 6
<superScript id="5A85B413A869FFCED5D6FCDF8E8CFCF6" attach="left" box="[950,969,836,855]" fontSize="8" pageId="58" pageNumber="59">th</superScript>
to 8
<superScript id="5A85B413A869FFCED260FCDF8956FCF6" attach="none" box="[1024,1043,836,855]" fontSize="8" pageId="58" pageNumber="59">th</superScript>
carinate, 3
<superScript id="5A85B413A869FFCED779FCF18C69FCDC" attach="left" box="[281,300,874,893]" fontSize="8" pageId="58" pageNumber="59">rd</superScript>
and4
<superScript id="5A85B413A869FFCED70DFCF18CC5FCDC" attach="left" box="[365,384,874,893]" fontSize="8" pageId="58" pageNumber="59">th</superScript>
slightly raised at base, both with tubercle at base. Intervals shiny, with isodiametric reticulation laterally and at base. Interval 3 with four setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA869FFCED6A4FC468F4DFC54" blockId="58.[159,1044,93,1573]" box="[196,520,989,1013]" pageId="58" pageNumber="59">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA869FFCED6A4FB998EE4FB74" blockId="58.[159,1044,93,1573]" pageId="58" pageNumber="59">
Ventral surface: Epipleuron shiny, covered with big, scattered punctures. submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle like an inverted V, mesosternum smooth. Sternites with transverse to irregular reticulation, each with bigger punctures laterally, 3
<superScript id="5A85B413A869FFCED585FBF48EB2FB23" attach="left" box="[997,1015,1135,1154]" fontSize="8" pageId="58" pageNumber="59">rd</superScript>
to 5
<superScript id="5A85B413A869FFCED6CCFB0F8DFBFB06" attach="left" box="[172,190,1172,1191]" fontSize="8" pageId="58" pageNumber="59">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA869FFCED6A4FB798F70FA31" blockId="58.[159,1044,93,1573]" pageId="58" pageNumber="59">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, with indistinct reticulation, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA869FFCED6A4FA068CA0FA14" blockId="58.[159,1044,93,1573]" box="[196,485,1437,1461]" pageId="58" pageNumber="59">Male genitalia: Unknown.</paragraph>
<paragraph id="AD4F195BA869FFCED6A4FA598FB1F984" blockId="58.[159,1044,93,1573]" pageId="58" pageNumber="59">
Female genitalia (
<figureCitation id="35CB05DEA869FFCED7FCFA598CB6FA7B" box="[412,499,1474,1498]" captionStart="Figs 39–40" captionStartId="36.[159,196,705,723]" captionTargetBox="[159,1032,80,662]" captionTargetId="figure-227@36.[159,590,115,627]" captionTargetPageId="36" captionText="Figs 39–40: Orictites spp., coxostylus. 39: O. tubercucollis sp. nov., scale bar 0.1 mm; 40: O. charleshuberi sp. nov., scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284374" httpUri="https://zenodo.org/record/6284374/files/figure.png" pageId="58" pageNumber="59">Fig. 40</figureCitation>
): Coxostylus slightly sigmoid, gently curved to apex, apex broad and with a pair of setae close together, with five longer nematiform and one slender ensiform seta at middle.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A868FFCFD6FEFFC68EFCFFD4" box="[158,953,93,117]" pageId="59" pageNumber="60" type="discussion">
<paragraph id="AD4F195BA868FFCFD6FEFFC68EFCFFD4" blockId="59.[121,1006,93,565]" box="[158,953,93,117]" pageId="59" pageNumber="60">
Variation: In the
<typeStatus id="724BA7F9A868FFCFD738FFC68C86FFD4" box="[344,451,93,117]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
, striae 1 and 2 adjoining indistinctly at apex.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A868FFCFD6FEFF198FF2FEF4" pageId="59" pageNumber="60" type="diagnosis">
<paragraph id="AD4F195BA868FFCFD6FEFF198FF2FEF4" blockId="59.[121,1006,93,565]" pageId="59" pageNumber="60">Diagnosis: A medium-sized nearly black species with distinct knob on the clypeus and distinct transverse depression with punctures behind in form of the letter V, with distinct and sharp humeral tooth, with tubercles at the base of intervals 3 and 4 of the elytra, and four setigerous punctures on interval 3. Distinguished from all other species by the crenulation of the lateral margin of the pronotum, and of the elytra in the anterior half.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A868FFCFD6FEFEF98C05FE01" pageId="59" pageNumber="60" type="distribution">
<paragraph id="AD4F195BA868FFCFD6FEFEF98C05FE01" blockId="59.[121,1006,93,565]" pageId="59" pageNumber="60">
Distribution: Known from
<collectingCountry id="D5E759CBA868FFCFD7ADFEF98F59FEDB" box="[461,540,354,378]" name="Taiwan" pageId="59" pageNumber="60">Taiwan</collectingCountry>
only. The
<typeStatus id="724BA7F9A868FFCFD4FAFEF98E44FEDB" box="[666,769,354,378]" pageId="59" pageNumber="60" type="paratype">paratype</typeStatus>
was collected at an altitude of
<specimenCount id="BBF6D2D2A868FFCFD693FE138C05FE01" box="[243,320,392,416]" pageId="59" pageNumber="60" type="generic">
<quantity id="6A08B4BEA868FFCFD693FE138C78FE01" box="[243,317,392,416]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="59" pageNumber="60" unit="m" value="950.0">950 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A868FFCFD6FEFE368DEEFD94" pageId="59" pageNumber="60" type="etymology">
<paragraph id="AD4F195BA868FFCFD6FEFE368DEEFD94" blockId="59.[121,1006,93,565]" pageId="59" pageNumber="60">
Etymology: The name is a patronym of Dr. Charles Huber, former curator at the Naturhistorisches Museum der Burgergemeinde
<collectingRegion id="6F34D7B9A868FFCFD488FE498E5BFE4B" box="[744,798,466,490]" country="Switzerland" name="Bern" pageId="59" pageNumber="60">Bern</collectingRegion>
(
<collectingCountry id="D5E759CBA868FFCFD54BFE498EF2FE4B" box="[811,951,466,490]" name="Switzerland" pageId="59" pageNumber="60">Switzerland</collectingCountry>
) and specialist in
<taxonomicName id="6AF062D8A868FFCFD767FE638C30FDB1" box="[263,373,504,528]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="subFamily" subFamily="Nebriinae">Nebriinae</taxonomicName>
who supported my work significantly over the last decade.
</paragraph>
</subSubSection>
</treatment>
</document>

267
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<subSubSection id="E5EA4AD0A874FFD3D619FCF68CADFC24" box="[121,488,877,902]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="AD4F195BA874FFD3D619FCF68CADFC24" blockId="39.[121,488,877,902]" box="[121,488,877,902]" pageId="39" pageNumber="40">
<heading id="F607AE37A874FFD3D619FCF68CADFC24" box="[121,488,877,902]" centered="true" fontSize="12" level="2" pageId="39" pageNumber="40" reason="2">
<taxonomicName id="6AF062D8A874FFD3D619FCF68CCFFC27" ID-CoL="8M6VK" authorityName="Balkenohl" authorityYear="2017" box="[121,394,877,902]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="omnipunctatus" status="sp. nov.">
<emphasis id="9F84C549A874FFD3D619FCF68CCFFC27" bold="true" box="[121,394,877,902]" italics="true" pageId="39" pageNumber="40">Orictites omnipunctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A874FFD3D7F0FCF68CADFC24" box="[400,488,877,901]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="E5EA4AD0A874FFD3D619FC238EE0FB11" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="AD4F195BA874FFD3D619FC238E77FC54" blockId="39.[121,1006,952,1200]" pageId="39" pageNumber="40">
Type material:
<materialsCitation id="1D981306A874FFD3D742FC238E77FC54" ID-GBIF-Occurrence="3497007310" collectingDate="1990-12-14" collectingDateMax="1990-12-18" collectingDateMin="1990-12-14" collectionCode="NHMB" collectorName="Bolm" country="Philippines" elevation="150" location="PORT BARTON" pageId="39" pageNumber="40" specimenCount="1" specimenCount-male="1" stateProvince="Palawan" typeStatus="holotype">
<typeStatus id="724BA7F9A874FFD3D742FC238CCBFC71" box="[290,398,952,976]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A874FFD3D7F8FC238CE9FC6E" box="[408,428,952,975]" pageId="39" pageNumber="40" type="male">♂</specimenCount>
, labels: white, black printed: “
<collectingCountry id="D5E759CBA874FFD3D565FC238ED8FC71" box="[773,925,952,976]" name="Philippines" pageId="39" pageNumber="40">PHILIPPINES</collectingCountry>
,
<quantity id="6A08B4BEA874FFD3D5C7FC238EA8FC71" box="[935,1005,952,976]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="39" pageNumber="40" unit="m" value="150.0">
<elevation id="26DDFE68A874FFD3D5C7FC238EA8FC71" box="[935,1005,952,976]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="39" pageNumber="40" unit="m" value="150.0">150 m</elevation>
</quantity>
<collectingRegion id="6F34D7B9A874FFD3D619FC468D98FC54" box="[121,221,989,1013]" country="Philippines" name="Palawan" pageId="39" pageNumber="40">Palawan</collectingRegion>
,
<location id="A82F4F80A874FFD3D686FC468CCCFC54" LSID="urn:lsid:plazi:treatment:2559A84DA874FFDED619FCF68C4FFC71:A82F4F80A874FFD3D686FC468CCCFC54" box="[230,393,989,1013]" country="Philippines" name="PORT BARTON" pageId="39" pageNumber="40" stateProvince="Palawan">PORT BARTON</location>
<date id="D94E3F9BA874FFD3D7EFFC468F08FC54" box="[399,589,989,1013]" pageId="39" pageNumber="40" value="1990-12-14" valueMax="1990-12-18" valueMin="1990-12-14">
<collectingDate id="C90AC673A874FFD3D7EFFC468F08FC54" box="[399,589,989,1013]" pageId="39" pageNumber="40" value="1990-12-14" valueMax="1990-12-18" valueMin="1990-12-14">14.–18.Dec 1990</collectingDate>
</date>
,
<collectorName id="00057C8DA874FFD3D43BFC468FD0FC54" box="[603,661,989,1013]" pageId="39" pageNumber="40">Bolm</collectorName>
lgt.” (
<collectionCode id="CBE1819EA874FFD3D4B8FC468E6DFC54" box="[728,808,989,1013]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="39" pageNumber="40">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="AD4F195BA874FFD3D619FB998EE0FB11" blockId="39.[121,1006,952,1200]" pageId="39" pageNumber="40">
<materialsCitation id="1D981306A874FFD3D619FB998E12FBBB" ID-GBIF-Occurrence="3497007331" box="[121,855,1026,1050]" collectingDate="1990-12-14" collectingDateMax="1990-12-18" collectingDateMin="1990-12-14" collectionCode="NHMB, CBB" country="Philippines" elevation="150" location="Bolm" pageId="39" pageNumber="40" specimenCount="4" stateProvince="Palawan" typeStatus="paratype">
<typeStatus id="724BA7F9A874FFD3D619FB998DB7FBBB" box="[121,242,1026,1050]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="BBF6D2D2A874FFD3D760FB998CDDFBBB" box="[256,408,1026,1050]" pageId="39" pageNumber="40" type="generic">4 specimens</specimenCount>
, same data as holotype (
<collectionCode id="CBE1819EA874FFD3D4A5FB998E56FBBB" box="[709,787,1026,1050]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="39" pageNumber="40">NHMB</collectionCode>
/
<collectionCode id="CBE1819EA874FFD3D57BFB998E14FBBB" box="[795,849,1026,1050]" collectionName="CBB" pageId="39" pageNumber="40">CBB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A874FFD3D504FB998EAFFBC4" ID-GBIF-Occurrence="3497007343" collectingDate="1999-12-06" collectingDateMax="1999-12-10" collectingDateMin="1999-12-06" collectionCode="SMNS, CBB" collectorName="A. Riedel" country="Philippines" elevation="700" location="Torosik" municipality="Kotamobagu Metalibaru" pageId="39" pageNumber="40" specimenCount="5" specimenCount-female="1" specimenCount-male="1" stateProvince="SULAWESI" typeStatus="paratype">
<specimenCount id="BBF6D2D2A874FFD3D504FB998ED4FBBB" box="[868,913,1026,1050]" pageId="39" pageNumber="40" type="male">1 ♂</specimenCount>
,
<specimenCount id="BBF6D2D2A874FFD3D5C2FB998E8BFBBB" box="[930,974,1026,1050]" pageId="39" pageNumber="40" type="female">1 ♀</specimenCount>
,
<specimenCount id="BBF6D2D2A874FFD3D5BFFB998DBCFBE1" pageId="39" pageNumber="40" type="generic">3 specimens</specimenCount>
. “
<collectingRegion id="6F34D7B9A874FFD3D772FBB38CD3FBE1" box="[274,406,1064,1088]" country="Indonesia" name="Sulawesi Utara" pageId="39" pageNumber="40">SULAWESI</collectingRegion>
:
<collectingMunicipality id="4D2B8321A874FFD3D7C8FBB38F89FBE1" box="[424,716,1064,1088]" pageId="39" pageNumber="40">Kotamobagu Metalibaru</collectingMunicipality>
,
<emphasis id="9F84C549A874FFD3D4BEFBBD8FB2FBE1" bold="true" box="[734,759,1062,1088]" italics="true" pageId="39" pageNumber="40"></emphasis>
<location id="A82F4F80A874FFD3D497FBB38E17FBE1" LSID="urn:lsid:plazi:treatment:2559A84DA874FFDED619FCF68C4FFC71:A82F4F80A874FFD3D497FBB38E17FBE1" box="[759,850,1064,1088]" country="Philippines" municipality="Kotamobagu Metalibaru" name="Torosik" pageId="39" pageNumber="40" stateProvince="SULAWESI">Torosik</location>
,
<location id="A82F4F80A874FFD3D535FBB38EAFFBE1" LSID="urn:lsid:plazi:treatment:2559A84DA874FFDED619FCF68C4FFC71:A82F4F80A874FFD3D535FBB38EAFFBE1" box="[853,1002,1064,1088]" country="Philippines" municipality="Kotamobagu Metalibaru" name="Gn. Tongara" pageId="39" pageNumber="40" stateProvince="SULAWESI">Gn. Tongara</location>
,
<date id="D94E3F9BA874FFD3D619FBD68DBEFBC4" box="[121,251,1101,1125]" pageId="39" pageNumber="40" value="1999-12-10">
<collectingDate id="C90AC673A874FFD3D619FBD68DBEFBC4" box="[121,251,1101,1125]" pageId="39" pageNumber="40" value="1999-12-10">10.XII.1999</collectingDate>
</date>
<quantity id="6A08B4BEA874FFD3D765FBD68C0FFBC4" box="[261,330,1101,1125]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="39" pageNumber="40" unit="m" value="700.0">
<elevation id="26DDFE68A874FFD3D765FBD68C0FFBC4" box="[261,330,1101,1125]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="39" pageNumber="40" unit="m" value="700.0">700m</elevation>
</quantity>
, and
<date id="D94E3F9BA874FFD3D7EBFBD68F45FBC4" box="[395,512,1101,1125]" pageId="39" pageNumber="40" value="1999-12-06">
<collectingDate id="C90AC673A874FFD3D7EBFBD68F45FBC4" box="[395,512,1101,1125]" pageId="39" pageNumber="40" value="1999-12-06">6.XII.1999</collectingDate>
</date>
,
<quantity id="6A08B4BEA874FFD3D470FBD68FECFBC4" box="[528,681,1101,1125]" metricMagnitude="3" metricUnit="m" metricValue="1.275" metricValueMax="1.45" metricValueMin="1.1" pageId="39" pageNumber="40" unit="m" value="1275.0" valueMax="1450.0" valueMin="1100.0">
<elevation id="26DDFE68A874FFD3D470FBD68FECFBC4" box="[528,681,1101,1125]" metricMagnitude="3" metricUnit="m" metricValue="1.275" metricValueMax="1.45" metricValueMin="1.1" pageId="39" pageNumber="40" unit="m" value="1275.0" valueMax="1450.0" valueMin="1100.0">1100–1450m</elevation>
</quantity>
, leg.
<collectorName id="00057C8DA874FFD3D4BDFBD68E01FBC4" box="[733,836,1101,1125]" pageId="39" pageNumber="40">A.Riedel</collectorName>
” (
<collectionCode id="CBE1819EA874FFD3D53CFBD68EEDFBC4" box="[860,936,1101,1125]" collectionName="Germany, Stuttgart, Staatliches Museum fuer Naturkunde" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="39" pageNumber="40">SMNS</collectionCode>
/
<collectionCode id="CBE1819EA874FFD3D5D0FBD68EA0FBC4" box="[944,997,1101,1125]" collectionName="CBB" pageId="39" pageNumber="40">CBB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="1D981306A874FFD3D619FBE98EE4FB11" ID-GBIF-Occurrence="3497007321" collectedFrom="Under bark of fallen tree" collectingDate="1980-01-27" collectingDateMax="1980-04-20" collectingDateMin="1980-01-27" collectionCode="BMNH, CBB" collectorName="M. J. D. Brendell" country="Philippines" location="TENGAH" municipality="Ranu River Area" pageId="39" pageNumber="40" specimenCount="3" specimenCount-female="3" stateProvince="SULAWESI" typeStatus="paratype">
<specimenCount id="BBF6D2D2A874FFD3D619FBE98DE0FB2B" box="[121,165,1138,1162]" pageId="39" pageNumber="40" type="female">3 ♀</specimenCount>
, “
<collectingRegion id="6F34D7B9A874FFD3D6DDFBE98C7AFB2B" box="[189,319,1138,1162]" country="Indonesia" name="Sulawesi Utara" pageId="39" pageNumber="40">SULAWESI</collectingRegion>
,
<location id="A82F4F80A874FFD3D72BFBE98CF5FB2B" LSID="urn:lsid:plazi:treatment:2559A84DA874FFDED619FCF68C4FFC71:A82F4F80A874FFD3D72BFBE98CF5FB2B" box="[331,432,1138,1162]" country="Philippines" municipality="Ranu River Area" name="TENGAH" pageId="39" pageNumber="40" stateProvince="SULAWESI">TENGAH</location>
:
<location id="A82F4F80A874FFD3D7DCFBE98F09FB2B" LSID="urn:lsid:plazi:treatment:2559A84DA874FFDED619FCF68C4FFC71:A82F4F80A874FFD3D7DCFBE98F09FB2B" box="[444,588,1138,1162]" country="Philippines" municipality="Ranu River Area" name="Nr. Morowali" pageId="39" pageNumber="40" stateProvince="SULAWESI">Nr.Morowali</location>
,
<location id="A82F4F80A874FFD3D438FBE98E5DFB2B" LSID="urn:lsid:plazi:treatment:2559A84DA874FFDED619FCF68C4FFC71:A82F4F80A874FFD3D438FBE98E5DFB2B" box="[600,792,1138,1162]" country="Philippines" municipality="Ranu River Area" name="Ranu River Area" pageId="39" pageNumber="40" stateProvince="SULAWESI">Ranu River Area</location>
.
<date id="D94E3F9BA874FFD3D545FBE98E92FB2B" box="[805,983,1138,1162]" pageId="39" pageNumber="40" value="1980-01-27" valueMax="1980-04-20" valueMin="1980-01-27">
<collectingDate id="C90AC673A874FFD3D545FBE98E92FB2B" box="[805,983,1138,1162]" pageId="39" pageNumber="40" value="1980-01-27" valueMax="1980-04-20" valueMin="1980-01-27">27.i.–20.iv.1980</collectingDate>
</date>
/
<collectorName id="00057C8DA874FFD3D619FB038C5AFB11" box="[121,287,1176,1200]" pageId="39" pageNumber="40">M.J.D.Brendell</collectorName>
, B.M.1980–280 /
<collectedFrom id="1EEC5AA6A874FFD3D788FB038FB8FB11" box="[488,765,1176,1200]" pageId="39" pageNumber="40">Under bark of fallen tree</collectedFrom>
” (
<collectionCode id="CBE1819EA874FFD3D572FB038E1BFB11" box="[786,862,1176,1200]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="39" pageNumber="40">BMNH</collectionCode>
/
<collectionCode id="CBE1819EA874FFD3D506FB038EDEFB11" box="[870,923,1176,1200]" collectionName="CBB" pageId="39" pageNumber="40">CBB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A874FFD3D619FA9C8CA2F984" pageId="39" pageNumber="40" type="description">
<paragraph id="AD4F195BA874FFD3D619FA9C8F7AFA81" blockId="39.[121,575,1287,1312]" box="[121,575,1287,1312]" pageId="39" pageNumber="40">
<emphasis id="9F84C549A874FFD3D619FA9C8DB9FA81" bold="true" box="[121,252,1287,1312]" pageId="39" pageNumber="40">Description</emphasis>
(
<figureCitation id="35CB05DEA874FFD3D769FA938C0FFA81" box="[265,330,1288,1312]" captionStart="Figs 5–8" captionStartId="8.[159,196,1494,1512]" captionTargetBox="[159,1021,146,1417]" captionTargetId="figure-66@8.[159,590,846,1416]" captionTargetPageId="8" captionText="Figs 5–8: Orictites spp., habitus. 5: O. plurisetosus sp. nov., holotype, male; 6: O. anteriortenuis sp. nov., holotype, male; 7: O. omnipunctatus sp. nov., holotype, male; 8: O. ampliosetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284326" httpUri="https://zenodo.org/record/6284326/files/figure.png" pageId="39" pageNumber="40">Figs 7</figureCitation>
,
<figureCitation id="35CB05DEA874FFD3D738FA938C31FA81" box="[344,372,1288,1312]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="39" pageNumber="40">25</figureCitation>
,
<figureCitation id="35CB05DEA874FFD3D7E0FA938CE4FA81" box="[384,417,1288,1312]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="39" pageNumber="40">36</figureCitation>
; pp. 9, 24, 36)
</paragraph>
<paragraph id="AD4F195BA874FFD3D619FAC98FD4FA14" blockId="39.[121,1006,1361,1573]" pageId="39" pageNumber="40">
Measurements (n = 10): Body length
<quantity id="6A08B4BEA874FFD3D44CFAC98F8AFACB" box="[556,719,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.95" metricValueMin="3.45" pageId="39" pageNumber="40" unit="mm" value="3.7" valueMax="3.95" valueMin="3.45">3.45–3.95 mm</quantity>
(
<emphasis id="9F84C549A874FFD3D481FACA8FB5FACB" bold="true" box="[737,752,1361,1386]" italics="true" pageId="39" pageNumber="40"></emphasis>
=
<quantity id="6A08B4BEA874FFD3D56AFAC98E20FACB" box="[778,869,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="39" pageNumber="40" unit="mm" value="3.7">3.7 mm</quantity>
); width 0.9- 1.03 (
<emphasis id="9F84C549A874FFD3D6DDFAEC8D88FA31" bold="true" box="[189,205,1399,1424]" italics="true" pageId="39" pageNumber="40"></emphasis>
=
<quantity id="6A08B4BEA874FFD3D692FAE38C18FA31" box="[242,349,1400,1424]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="39" pageNumber="40" unit="mm" value="0.95">0.95 mm</quantity>
); ration length/width of pronotum 1.03–1.11 (
<emphasis id="9F84C549A874FFD3D516FAEC8EC3FA31" bold="true" box="[886,902,1399,1424]" italics="true" pageId="39" pageNumber="40"></emphasis>
= 1.07); ratio length/width of elytra 2.0–2.09 (
<emphasis id="9F84C549A874FFD3D443FA078F76FA14" bold="true" box="[547,563,1436,1461]" italics="true" pageId="39" pageNumber="40"></emphasis>
= 2.05).
</paragraph>
<paragraph id="AD4F195BA874FFD3D6FEFA598CA2F984" blockId="39.[121,1006,1361,1573]" pageId="39" pageNumber="40">Colour: Shiny. Red-brown. Elytra red-brown, pronotum darker. Wings of clypeus and supraantennal plates transparent red-brown, antennae, mouthparts, and legs yellowish brown.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87BFFDCD6A4FFC68CD1F9A1" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="AD4F195BA87BFFDCD6A4FFC68CD1F9A1" blockId="40.[159,1044,93,1536]" pageId="40" pageNumber="41">
Head: A fifth narrower than pronotum. Clypeus limited laterally by small rounded tooth, anterior margin excised, convex between the two clypeal setae, smooth but covered with medium sized punctures, wings obtuse-angular, margin of wings convex, slightly hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches, clypeus and wings reflexed margined, supraantennal plates distinctly vaulted up to mid-eye level, smooth, reflexed margin rounded, elongated as supraorbital carina, frons not separated from clypeus, thought indistinct flat depression visible laterally, supraorbital plates elongated posteriorly as obtuse diverging carina up to neck constriction, separated from supraorbital carinae by broad furrow and from frons by sharp diverging furrow. Frons distinctly convex without median impression, covered irregularly with punctures of large and medium size. Supraorbital setae located in broad supraorbital furrows, the posterior one arising from tubercle. Eyes of moderate size, flattened but still convex, genae distinct, regularly rounded, as high as and shorter than eyes, enclosing eyes posteriorly by one sixth, forming obtuse angle at neck. Grooves for reception of antennae short, length about 0.4 of eye diameter. Neck constriction nearly invisible at middle, laterally with densely scattered medium sized punctures. Labrum slightly advanced at middle, 7-setose, ciliate laterally, with fine isodiametric reticulation (120x). Mandibles half as long as head, flattened, stout, broadened at base, gently curved at apex, carinae of scrobe complete, both mandibles rounded towards base, right one ventrally with small tooth at middle. Both mandibles somewhat hollowed out dorsally, both of them with small but distinct tooth near base (visible in completely opened position). Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A87BFFDCD539FBBF8E35FB96" attach="left" box="[857,880,1060,1079]" fontSize="8" pageId="40" pageNumber="41">nd</superScript>
segment, 2
<superScript id="5A85B413A87BFFDCD59CFBBF8956FB96" attach="none" box="[1020,1043,1060,1079]" fontSize="8" pageId="40" pageNumber="41">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with few punctures and reticulation, Submentum and mentum distinctly separated; submentum with four, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae short, just reaching over middle of pronotum, scapus knee-like angled, with one seta dorso-apically, scapus and pedicellus nearly smooth, segments 5–10 transverse (L/
<geoCoordinate id="C8C47F9CA87BFFDCD79AFA068F06FA14" box="[506,579,1437,1461]" degrees="0. 8" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="5555" value="-0.8">W 0.8</geoCoordinate>
), antennomeres slightly flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87AFFDED6FEFFC68CCCFD94" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="AD4F195BA87AFFDDD6FEFFC68E4EFDFB" blockId="41.[121,1006,93,1536]" pageId="41" pageNumber="42">Pronotum: Square, distinctly longer than wide, sides straight, gently rounded before posterior angle. Anterior margin straight. Reflexed lateral margin smooth, from posterior angle to base running as straight line. Lateral channel broadened between the two lateral setigerous punctures, smooth, with very few fine punctures. Anterior setigerous puncture located at the end of anterior quarter adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by twice diameter of pore. Anterior angles not projecting, nearly right-angled in dorsal view, formed by reflexed lateral margin, posterior angle developed as distinct tooth. Anterior transverse line formed by distinct partly connected punctures. Median line deep, broad, surpassing level of anterior transverse line without joining, smaller at base, joining base. Whole surface irregularly covered with medium and big sized punctures. No basal impression, no reticulation at base. Basal channel broad, deep. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA87AFFDDD6FEFDF38E9AFBC4" blockId="41.[121,1006,93,1536]" pageId="41" pageNumber="42">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, completely visible from above in anterior two thirds, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of punctures laterally. Reflexed margin with a few fine, indistinct scars posterior to humerus, channel slightly thickened at humerus, margin bending with channel over rounded humerus up to 5
<superScript id="5A85B413A87AFFDDD4E9FC848FD9FC93" attach="left" box="[649,668,799,818]" fontSize="8" pageId="41" pageNumber="42">th</superScript>
stria. No tooth at humerus. Basal tubercle distinct, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, broad, punctate, striae 1 and 7 joining at apex, 2 ending free, 3 and 4, 5 and 6 joined apically; striae 6 and 7 shortened at humerus. Striole short but distinct, interval between striole and sutura costiform, raised. All intervals conspicuously convex, thought costate, 2
<superScript id="5A85B413A87AFFDDD6E7FC648DDBFBB3" attach="left" box="[135,158,1023,1042]" fontSize="8" pageId="41" pageNumber="42">nd</superScript>
to4
<superScript id="5A85B413A87AFFDDD6ADFC648DA5FBB3" attach="left" box="[205,224,1023,1042]" fontSize="8" pageId="41" pageNumber="42">th</superScript>
slightly broader at base, 3
<superScript id="5A85B413A87AFFDDD476FC648F6CFBB3" attach="left" box="[534,553,1023,1042]" fontSize="8" pageId="41" pageNumber="42">rd</superScript>
and4
<superScript id="5A85B413A87AFFDDD40AFC648F38FBB3" attach="left" box="[618,637,1023,1042]" fontSize="8" pageId="41" pageNumber="42">th</superScript>
with small tubercle at base, 6
<superScript id="5A85B413A87AFFDDD5BBFC648EABFBB3" attach="right" box="[987,1006,1023,1042]" fontSize="8" pageId="41" pageNumber="42">th</superScript>
costate, 7
<superScript id="5A85B413A87AFFDDD68DFBBF8C45FB96" attach="left" box="[237,256,1060,1079]" fontSize="8" pageId="41" pageNumber="42">th</superScript>
and 8
<superScript id="5A85B413A87AFFDDD72DFBBF8C25FB96" attach="left" box="[333,352,1060,1079]" fontSize="8" pageId="41" pageNumber="42">th</superScript>
completely carinae in total. All intervals shiny on disc. Interval 3 with four to seven setigerous punctures (see
<tableCitation id="E0722CE0A87AFFDDD4B7FBD68E5DFBC4" box="[727,792,1101,1125]" captionStart="Tab" captionStartId="42.[159,193,1105,1123]" captionTargetPageId="42" captionText="Tab. 1: Orictites omnipunctatus sp. nov.: Variation of the number of setigerous punctures on the 3rd interval of the right/left elytra." httpUri="http://table.plazi.org/id/F98F49D3A879FFDED6FFFBCA8CEAFBDF" pageId="41" pageNumber="42" tableUuid="F98F49D3A879FFDED6FFFBCA8CEAFBDF">Tab. 1</tableCitation>
) adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA87AFFDDD6FEFBE98CA6FB2B" blockId="41.[121,1006,93,1536]" box="[158,483,1138,1162]" pageId="41" pageNumber="42">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA87AFFDDD6FEFB038F69F9A1" blockId="41.[121,1006,93,1536]" pageId="41" pageNumber="42">
Ventral surface: Proepipleuron small, submarginal furrow distinct, engraved. Epipleuron broadened and with big punctures in basal quarter. Proepisternum somewhat swollen at level of posterior setigerous puncture, proepisternum and episternum covered densely and irregularly with big punctures, prosternite smooth at middle, with confluent double keel at middle, mesosternum smooth at middle. Sternites with big and medium-sized punctures, 3
<superScript id="5A85B413A87AFFDDD6B3FAEF8DA3FA26" attach="left" box="[211,230,1396,1415]" fontSize="8" pageId="41" pageNumber="42">rd</superScript>
to 5
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with paralateral ambulatory setae at each side, ventral strigae distinct; sternite 6 with big punctures at base, becoming finer at middle, minute towards apex, with two widely separated apical setae at each side, the inner one arising from tooth of margin.
</paragraph>
<paragraph id="AD4F195BA879FFDED6A4FFC68DACFEAB" blockId="42.[159,1044,93,976]" pageId="42" pageNumber="43">Legs: Anterior tibia with strong digitation, with two big lateral preapical denticles, third one small, apical spine distinctly curved ventral and lateral, no dorsal furrow, basal tarsomere as long as the following three together. Mesotibia with apical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA879FFDED6A4FE838E45FEDB" blockId="42.[159,1044,93,976]" pageId="42" pageNumber="43">
Male genitalia (
<figureCitation id="35CB05DEA879FFDED71AFE838C8EFE91" box="[378,459,280,304]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="42" pageNumber="43">Fig. 25</figureCitation>
): Median lobe slender, more distinctly arcuate at middle, oroficium large, endophallus with some small teeth, apex spatulate. Both parameres slender, dorsal one with seta at apex.
</paragraph>
<paragraph id="AD4F195BA879FFDED6A4FE138F72FE4B" blockId="42.[159,1044,93,976]" pageId="42" pageNumber="43">
Female genitalia (
<figureCitation id="35CB05DEA879FFDED7EEFE138C9BFE01" box="[398,478,392,416]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="42" pageNumber="43">Fig. 36</figureCitation>
): Coxostylus broad, slightly curved at apex, with one apical seta. Ensiform seta at middle short, broad. With five longer nematiform setae in middle and basal part.
</paragraph>
<paragraph id="AD4F195BA879FFDED6A4FE638CCCFD94" blockId="42.[159,1044,93,976]" pageId="42" pageNumber="43">
Variation: For variation of the setigerous punctures on the 3
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interval of the elytra see
<tableCitation id="E0722CE0A879FFDED75FFD868CC7FD94" box="[319,386,541,565]" captionStart="Tab" captionStartId="42.[159,193,1105,1123]" captionTargetPageId="42" captionText="Tab. 1: Orictites omnipunctatus sp. nov.: Variation of the number of setigerous punctures on the 3rd interval of the right/left elytra." httpUri="http://table.plazi.org/id/F98F49D3A879FFDED6FFFBCA8CEAFBDF" pageId="42" pageNumber="43" tableUuid="F98F49D3A879FFDED6FFFBCA8CEAFBDF">Tab. 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A879FFDED6A4FDD98F96FCB4" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="AD4F195BA879FFDED6A4FDD98F96FCB4" blockId="42.[159,1044,93,976]" pageId="42" pageNumber="43">Diagnosis: A small pale-brown species with the knob on the clypeus developed as elevation and indistinct flat transverse depression laterally, no humeral tooth, and with tubercles at the base of interval 3 and 4 of the elytra. Distinguished from all other species by the series of setigerous punctures on interval 3 of the elytra, the numerous and dense punctures on many parts of the body, and the coxostylus with one apical seta.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A879FFDED6A4FCB98CA9FC24" pageId="42" pageNumber="43" type="distribution">
<paragraph id="AD4F195BA879FFDED6A4FCB98CA9FC24" blockId="42.[159,1044,93,976]" pageId="42" pageNumber="43">
Distribution: Known from
<collectingRegion id="6F34D7B9A879FFDED791FCB98F13FC9B" box="[497,598,802,826]" country="Indonesia" name="Sulawesi Utara" pageId="42" pageNumber="43">Sulawesi</collectingRegion>
and
<collectingRegion id="6F34D7B9A879FFDED4F3FCB98FB2FC9B" box="[659,759,802,826]" country="Philippines" name="Palawan" pageId="42" pageNumber="43">Palawan</collectingRegion>
. The altitude of one collection spot is
<quantity id="6A08B4BEA879FFDED723FCD38CC8FCC1" box="[323,397,840,864]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="42" pageNumber="43" unit="m" value="150.0">150 m</quantity>
, of another one
<specimenCount id="BBF6D2D2A879FFDED422FCD38FD5FCC1" box="[578,656,840,864]" pageId="42" pageNumber="43" type="generic">
<quantity id="6A08B4BEA879FFDED422FCD38FC9FCC1" box="[578,652,840,864]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="42" pageNumber="43" unit="m" value="700.0">700 m</quantity>
.
</specimenCount>
One set of specimens was collected under bark of a fallen tree.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A879FFDED6A4FC098C4FFC71" pageId="42" pageNumber="43" type="etymology">
<paragraph id="AD4F195BA879FFDED6A4FC098C4FFC71" blockId="42.[159,1044,93,976]" pageId="42" pageNumber="43">Etymology: The name refers to the numerous punctures on many parts of the body.</paragraph>
</subSubSection>
</treatment>
</document>

224
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<taxonomicName id="6AF062D8A877FFD0D6FFFCDC8CEDFCC1" ID-CoL="8MLMV" authorityName="Balkenohl" authorityYear="2017" box="[159,424,839,864]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="anteriortenuis" status="sp. nov.">
<emphasis id="9F84C549A877FFD0D6FFFCDC8CEDFCC1" bold="true" box="[159,424,839,864]" italics="true" pageId="36" pageNumber="37">Orictites anteriortenuis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A877FFD0D7CEFCD38F43FCC1" box="[430,518,840,864]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="E5EA4AD0A877FFD0D6FFFC098FD0FC54" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="AD4F195BA877FFD0D6FFFC098FD0FC54" blockId="36.[159,1044,914,1237]" pageId="36" pageNumber="37">
Type material:
<materialsCitation id="1D981306A877FFD0D73AFC098FD0FC54" ID-GBIF-Occurrence="3497007304" collectingDate="1996-07-23" collectionCode="CBM-ZSM" elevation="700" location="Strasse von Nabire nach Mapia km 54 Sek. wald" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="West Papua" typeStatus="holotype">
<typeStatus id="724BA7F9A877FFD0D73AFC098C8CFC0B" box="[346,457,914,938]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A877FFD0D7BAFC088CABFC0B" box="[474,494,915,938]" pageId="36" pageNumber="37" type="male">♂</specimenCount>
Labels: white, black printed, black framed: “
<date id="D94E3F9BA877FFD0D6CBFC238C5EFC71" box="[171,283,952,976]" pageId="36" pageNumber="37" value="1996-07-23">
<collectingDate id="C90AC673A877FFD0D6CBFC238C5EFC71" box="[171,283,952,976]" pageId="36" pageNumber="37" value="1996-07-23">23.7.1996</collectingDate>
</date>
29 Schüle/ Stüben
<collectingRegion id="6F34D7B9A877FFD0D474FC238FDAFC71" box="[532,671,952,976]" country="Indonesia" name="Papua Barat" pageId="36" pageNumber="37">West Papua</collectingRegion>
<quantity id="6A08B4BEA877FFD0D4C9FC238FAFFC71" box="[681,746,952,976]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="37" unit="m" value="700.0">
<elevation id="26DDFE68A877FFD0D4C9FC238FAFFC71" box="[681,746,952,976]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="36" pageNumber="37" unit="m" value="700.0">700m</elevation>
</quantity>
<location id="A82F4F80A877FFD0D494FC238CD8FC54" LSID="urn:lsid:plazi:treatment:2559A84DA877FFD3D6FFFCDC8EA8FCB4:A82F4F80A877FFD0D494FC238CD8FC54" name="Strasse von Nabire nach Mapia km 54 Sek. wald" pageId="36" pageNumber="37" stateProvince="West Papua">Strasse von Nabire nach Mapia km 54 Sek.wald</location>
/ Busch” (
<collectionCode id="CBE1819EA877FFD0D473FC468FCEFC54" box="[531,651,989,1013]" pageId="36" pageNumber="37">CBM-ZSM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A877FFD0D6A4FB998F59FB74" pageId="36" pageNumber="37" type="discussion">
<paragraph id="AD4F195BA877FFD0D6A4FB998C4BFBE1" blockId="36.[159,1044,914,1237]" pageId="36" pageNumber="37">
Remarks: In the
<typeStatus id="724BA7F9A877FFD0D7E0FB998CA0FBBB" box="[384,485,1026,1050]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
the digitation of the left front tibia is missing its apical tip.
</paragraph>
<paragraph id="AD4F195BA877FFD0D6FFFBD68F59FB74" blockId="36.[159,1044,914,1237]" pageId="36" pageNumber="37">
<materialsCitation id="1D981306A877FFD0D6FFFBD68F20FB2B" ID-GBIF-Occurrence="3497007305" collectingDate="1997-10" collectionCode="CBB" collectorName="M. Balke" country="Iran" elevation="0" location="Jaya" municipality="Nabire-Ilaga" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="724BA7F9A877FFD0D6FFFBD68C5FFBC4" box="[159,282,1101,1125]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="BBF6D2D2A877FFD0D74BFBD68C1EFBC4" box="[299,347,1101,1125]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
, labels: white, black printed: “
<collectingCountry id="D5E759CBA877FFD0D481FBD68E57FBC4" box="[737,786,1101,1125]" name="Iran" pageId="36" pageNumber="37">Iran</collectingCountry>
<location id="A82F4F80A877FFD0D540FBD68E1CFBC4" LSID="urn:lsid:plazi:treatment:2559A84DA877FFD3D6FFFCDC8EA8FCB4:A82F4F80A877FFD0D540FBD68E1CFBC4" box="[800,857,1101,1125]" country="Iran" municipality="Nabire-Ilaga" name="Jaya" pageId="36" pageNumber="37">Jaya</location>
,
<location id="A82F4F80A877FFD0D53CFBD68EAAFBC4" LSID="urn:lsid:plazi:treatment:2559A84DA877FFD3D6FFFCDC8EA8FCB4:A82F4F80A877FFD0D53CFBD68EAAFBC4" box="[860,1007,1101,1125]" country="Iran" municipality="Nabire-Ilaga" name="Nabire-Ilaga" pageId="36" pageNumber="37">Nabire-Ilaga</location>
,
<location id="A82F4F80A877FFD0D591FBD68DF9FB2B" LSID="urn:lsid:plazi:treatment:2559A84DA877FFD3D6FFFCDC8EA8FCB4:A82F4F80A877FFD0D591FBD68DF9FB2B" country="Iran" municipality="Nabire-Ilaga" name="km 54" pageId="36" pageNumber="37">
km
<quantity id="6A08B4BEA877FFD0D6FFFBE98DF9FB2B" box="[159,188,1138,1162]" metricMagnitude="4" metricUnit="m" metricValue="5.475" pageId="36" pageNumber="37" unit="m" value="54750.0">54</quantity>
</location>
,
<elevation id="26DDFE68A877FFD0D6A4FBE98C43FB2B" box="[196,262,1138,1162]" pageId="36" pageNumber="37">750m</elevation>
<date id="D94E3F9BA877FFD0D776FBE98C2DFB2B" box="[278,360,1138,1162]" pageId="36" pageNumber="37" value="1997-10">
<collectingDate id="C90AC673A877FFD0D776FBE98C2DFB2B" box="[278,360,1138,1162]" pageId="36" pageNumber="37" value="1997-10">X.1997</collectingDate>
</date>
leg.
<collectorName id="00057C8DA877FFD0D7C1FBE98F40FB2B" box="[417,517,1138,1162]" pageId="36" pageNumber="37">M.Balke</collectorName>
” (
<collectionCode id="CBE1819EA877FFD0D446FBE98F1AFB2B" box="[550,607,1138,1162]" collectionName="CBB" pageId="36" pageNumber="37">CBB</collectionCode>
)
</materialsCitation>
;
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, “
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/
<collectingCountry id="D5E759CBA877FFD0D5C8FBE98955FB2B" box="[936,1040,1138,1162]" pageId="36" pageNumber="37">Sumatra</collectingCountry>
: MjÖberg /
<taxonomicName id="6AF062D8A877FFD0D77CFB038CBAFB11" authority="Bates" authorityName="BATES Compared" authorityYear="1892" box="[284,511,1176,1200]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="costulipennis">costulipennis Bates</taxonomicName>
(petit ex.)
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H.E.
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</determinerName>
det. / 4122 E91 + /
<taxonomicName id="6AF062D8A877FFD0D6FFFB268CF1FB74" authority="H. W. Bates" authorityName="H. W. Bates" box="[159,436,1213,1237]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="costulipennis">costulipennis H.W.Bates</taxonomicName>
” (
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A877FFD3D6FFFAB68F10FEDB" lastPageId="39" lastPageNumber="40" pageId="36" pageNumber="37" type="description">
<paragraph id="AD4F195BA877FFD0D6FFFAB68F22FAE4" blockId="36.[159,615,1325,1350]" box="[159,615,1325,1350]" pageId="36" pageNumber="37">
<emphasis id="9F84C549A877FFD0D6FFFAB68C67FAE7" bold="true" box="[159,290,1325,1350]" pageId="36" pageNumber="37">Description</emphasis>
(
<figureCitation id="35CB05DEA877FFD0D751FAB68C32FAE4" box="[305,375,1325,1349]" captionStart="Figs 5–8" captionStartId="8.[159,196,1494,1512]" captionTargetBox="[159,1021,146,1417]" captionTargetId="figure-66@8.[159,590,846,1416]" captionTargetPageId="8" captionText="Figs 5–8: Orictites spp., habitus. 5: O. plurisetosus sp. nov., holotype, male; 6: O. anteriortenuis sp. nov., holotype, male; 7: O. omnipunctatus sp. nov., holotype, male; 8: O. ampliosetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284326" httpUri="https://zenodo.org/record/6284326/files/figure.png" pageId="36" pageNumber="37">Figs 6</figureCitation>
,
<figureCitation id="35CB05DEA877FFD0D7E1FAB68CE7FAE4" box="[385,418,1325,1349]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="36" pageNumber="37">24</figureCitation>
,
<figureCitation id="35CB05DEA877FFD0D7CBFAB68C8EFAE4" box="[427,459,1325,1349]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="36" pageNumber="37">35</figureCitation>
; pp. 9, 24, 36)
</paragraph>
<paragraph id="AD4F195BA877FFD0D6FFFAE38F82FA7B" blockId="36.[159,1044,1399,1498]" pageId="36" pageNumber="37">
Measurements (n = 3): Body length
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=
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), width
<quantity id="6A08B4BEA877FFD0D5B5FAE38C47FA14" metricMagnitude="-3" metricUnit="m" metricValue="1.245" metricValueMax="1.28" metricValueMin="1.21" pageId="36" pageNumber="37" unit="mm" value="1.245" valueMax="1.28" valueMin="1.21">1.21– 1.28 mm</quantity>
(
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=
<quantity id="6A08B4BEA877FFD0D722FA068CEFFA14" box="[322,426,1437,1461]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="36" pageNumber="37" unit="mm" value="1.24">1.24 mm</quantity>
), ratio length/width of pronotum 1.01–1.05 (
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= 1.03), ratio length/width of elytra 1.99–2.09 (
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= 2.04).
</paragraph>
<paragraph id="AD4F195BA876FFD1D6FEFFC68CC5FF61" blockId="37.[121,1006,93,1536]" pageId="37" pageNumber="38">Colour: Shiny. Black. Wings of clypeus and supraantennal plates brown and slightly transparent, antennae, labrum and mandibles brown, palpi yellowish brown, legs red-brown.</paragraph>
<paragraph id="AD4F195BA876FFD1D6FEFF568DFAF9A1" blockId="37.[121,1006,93,1536]" pageId="37" pageNumber="38">
Head: A third narrower than pronotum. Clypeus with small rounded slightly projecting tooth laterally, distinctly separated from wings, anterior margin excised, convex between the two clypeal setae, margin of wings convex, slightly hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted over mid-eye level, widened laterally, smooth, with reflexed rounded margin, elongated as supraorbital carina. Frons separated from clypeus by indistinct transverse depression, sharp furrow between frons and frontal carina, frons moderately convex. Frons and clypeus with big punctures. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes well developed, convex. Genae distinct, regularly rounded, not as high as eyes, enclosing eyes posteriorly by one fifth, forming obtuse angle at neck. Grooves for reception of antennae short, length 0.25 of eye diameter. Neck constriction marked as slightly impressed line, impunctate. Neck laterally covered with reticulation and densely scattered medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with fine isodiametric reticulation. Mandibles robust, shorter than head, broadened at base, apical half elongated, acutely curved at apex, carinae of scrobe complete, angled, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A876FFD1D74DFBBF8C01FB96" attach="left" box="[301,324,1060,1079]" fontSize="8" pageId="37" pageNumber="38">nd</superScript>
segment, 2
<superScript id="5A85B413A876FFD1D7ACFBBF8CA6FB96" attach="left" box="[460,483,1060,1079]" fontSize="8" pageId="37" pageNumber="38">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with punctures and some transverse reticulation. Submentum and mentum distinctly separated; submentum with four bigger setigerous punctures; mentum at base bilaterally with rounded elevation, with a few longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth obtuse, triangle-like, bisetose, not projecting as far as lateral lobes. Antennae relatively long, reaching distinctly over two thirds of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine isodiametric reticulation, segments 5–10 transverse (L/
<geoCoordinate id="C8C47F9CA876FFD1D77AFA068C26FA14" box="[282,355,1437,1461]" degrees="0. 8" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="5555" value="-0.8">W 0.8</geoCoordinate>
), antennomeres slightly flattened, with shiny areas on flattened parts, segments 4–11 densely pubescent, with few additional long setae.
</paragraph>
<paragraph id="AD4F195BA875FFD2D6A4FFC68FC7FD04" blockId="38.[159,1045,93,1573]" pageId="38" pageNumber="39">Pronotum: Square, longer than wide, outline reminiscent of the frustum of a pyramid, sides straight, convex in anterior quarter for a short distance to anterior angles. Anterior margin indistinctly bisinuate. Reflexed lateral margin without any scars. Margin from posterior angle to base running as slightly sigmoid line. Lateral channel distinctly broadened between the two lateral setigerous punctures, with isodiametric reticulation and very few indistinct punctures. Anterior setigerous puncture located at the end of anterior quarter adjoining disc of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by twice the diameter of pore. Anterior angle slightly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as tooth, projecting laterally not as far as widest part of pronotum. Anterior transverse line narrow, sharp. Median line deep, broad, rugose, running up to level of anterior transverse line but not joining, smaller at base, just adjoining base. Surface irregularly covered with punctures of different size, basal impression indistinct. Basal channel moderately broad, deep. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA875FFD2D6A4FD298FE9FBE1" blockId="38.[159,1045,93,1573]" pageId="38" pageNumber="39">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles and second row of punctures. Reflexed margin with a few indistinct scars in anterior third, margin bending with channel over rounded humerus up to 5
<superScript id="5A85B413A875FFD2D68FFCDF8C47FCF6" attach="left" box="[239,258,836,855]" fontSize="8" pageId="38" pageNumber="39">th</superScript>
stria. Humerus without tooth. Basal tubercle distinct, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, indistinctly punctate, striae 1–2, 3–4, and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura slightly raised. Intervals moderately convex, 3
<superScript id="5A85B413A875FFD2D4C6FC418FFCFC4C" attach="left" box="[678,697,986,1005]" fontSize="8" pageId="38" pageNumber="39">rd</superScript>
and4
<superScript id="5A85B413A875FFD2D498FC418E4EFC4C" attach="left" box="[760,779,986,1005]" fontSize="8" pageId="38" pageNumber="39">th</superScript>
slightly raised at base, both with tubercle at base, 8
<superScript id="5A85B413A875FFD2D78CFC648CBAFBB3" attach="left" box="[492,511,1023,1042]" fontSize="8" pageId="38" pageNumber="39">th</superScript>
carinate. All intervals shiny on disc. Interval 3 with five setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA875FFD2D6A4FBD68F44FBC4" blockId="38.[159,1045,93,1573]" box="[196,513,1101,1125]" pageId="38" pageNumber="39">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA875FFD2D6A4FBE98C28FACB" blockId="38.[159,1045,93,1573]" pageId="38" pageNumber="39">
Ventral surface: Epipleuron shiny, covered with indistinct reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle, mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with punctures at base, slightly denser laterally, 3
<superScript id="5A85B413A875FFD2D756FA9F8C0CFAB6" attach="left" box="[310,329,1284,1303]" fontSize="8" pageId="38" pageNumber="39">rd</superScript>
to 5
<superScript id="5A85B413A875FFD2D7E0FA9F8CD6FAB6" attach="left" box="[384,403,1284,1303]" fontSize="8" pageId="38" pageNumber="39">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA875FFD2D6A4FAE38C6CF984" blockId="38.[159,1045,93,1573]" pageId="38" pageNumber="39">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, with flat dorsal furrow, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA874FFD3D6FEFFC68F85FF44" blockId="39.[121,1006,93,789]" pageId="39" pageNumber="40">
Male genitalia (
<figureCitation id="35CB05DEA874FFD3D738FFC68CEEFFD4" box="[344,427,93,117]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="39" pageNumber="40">Fig. 24</figureCitation>
): Moderately long and sclerotized. Median lobe slender, moderately arcuate, not twisted, nearly straight preapically, apex stick-like. Endophallus with group of teeth apically. Parameres of about same length, both slender, distorted preapically, asetose.
</paragraph>
<paragraph id="AD4F195BA874FFD3D6FEFF698F10FEDB" blockId="39.[121,1006,93,789]" pageId="39" pageNumber="40">
Female genitalia. (
<figureCitation id="35CB05DEA874FFD3D70DFF698CFFFEAB" box="[365,442,242,266]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="39" pageNumber="40">Fig. 35</figureCitation>
): Coxostylus stout, of moderate length, distinctly curved to apex, with a pair of setae arising closely together preapically, with three longer nematiform setae basally, with two bigger acute ensiform setae at middle and a smaller one towards base.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A874FFD3D6FEFE138E27FE64" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="AD4F195BA874FFD3D6FEFE138E27FE64" blockId="39.[121,1006,93,789]" pageId="39" pageNumber="40">
Variation: The basal impression of the pronotum varies in distinctness. Striae 1 and 2 indistinctly joined at apex at one side in a
<typeStatus id="724BA7F9A874FFD3D493FE368E18FE64" box="[755,861,429,453]" pageId="39" pageNumber="40" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A874FFD3D6FEFE498CE8FD04" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="AD4F195BA874FFD3D6FEFE498CE8FD04" blockId="39.[121,1006,93,789]" pageId="39" pageNumber="40">Diagnosis: A medium-sized black species with the knob on the clypeus developed as elevation and indistinct transverse depression behind, no humeral tooth, and tubercles at the base of intervals 3 and 4 of the elytra. Distinguished from all other species by the presence of five setigerous punctures on interval 3 of the elytra, and the neck constriction, developed as slightly impressed impunctate line.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A874FFD3D6FEFD298D83FD51" pageId="39" pageNumber="40" type="distribution">
<paragraph id="AD4F195BA874FFD3D6FEFD298D83FD51" blockId="39.[121,1006,93,789]" pageId="39" pageNumber="40">
Distribution. Known from West of
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collected at an altitude of 700 and
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<quantity id="6A08B4BEA874FFD3D619FD438D86FD51" box="[121,195,728,752]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="39" pageNumber="40" unit="m" value="750.0">750 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A874FFD3D6FEFD668EA8FCB4" box="[158,1005,765,789]" pageId="39" pageNumber="40" type="etymology">
<paragraph id="AD4F195BA874FFD3D6FEFD668EA8FCB4" blockId="39.[121,1006,93,789]" box="[158,1005,765,789]" pageId="39" pageNumber="40">Etymology: The name refers to the width of the pronotum decreasing anteriorly.</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:classification id="735B9F26099D73C35A80D9FFBAAB51DD">journal article</mods:classification>
<mods:identifier id="446C52578DBA9BD5160226EF4CA6474D" type="CLB-Dataset">20497</mods:identifier>
<mods:identifier id="8B477C96BA1575194671DC459D29C174" type="DOI">10.5169/seals-787048</mods:identifier>
<mods:identifier id="6487A8076006A9B815FDD321D4D39DA5" type="GBIF-Dataset">8c468ca9-0ee3-4200-b5a3-6adfd98caaea</mods:identifier>
<mods:identifier id="F1CA1C2CE7FBA895AD545729976E306B" type="ISSN">2624-9170</mods:identifier>
<mods:identifier id="291A5ACD967A4F7C13DDACE278327BC1" type="Zenodo-Dep">6284316</mods:identifier>
</mods:mods>
<treatment id="2559A84DA878FFD9D619FFC68E60FA14" ID-DOI="http://doi.org/10.5281/zenodo.6315583" ID-GBIF-Taxon="193586553" ID-Zenodo-Dep="6315583" LSID="urn:lsid:plazi:treatment:2559A84DA878FFD9D619FFC68E60FA14" httpUri="http://treatment.plazi.org/id/2559A84DA878FFD9D619FFC68E60FA14" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44">
<subSubSection id="E5EA4AD0A878FFDFD619FFC68CA0FFD4" box="[121,485,93,118]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="AD4F195BA878FFDFD619FFC68CA0FFD4" blockId="43.[121,485,93,118]" box="[121,485,93,118]" pageId="43" pageNumber="44">
<heading id="F607AE37A878FFDFD619FFC68CA0FFD4" box="[121,485,93,118]" centered="true" fontSize="12" level="2" pageId="43" pageNumber="44" reason="2">
<taxonomicName id="6AF062D8A878FFDFD619FFC68CC2FFD7" ID-CoL="8MH6D" authorityName="Balkenohl" authorityYear="2017" box="[121,391,93,118]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ampliosetosus" status="sp. nov.">
<emphasis id="9F84C549A878FFDFD619FFC68CC2FFD7" bold="true" box="[121,391,93,118]" italics="true" pageId="43" pageNumber="44">Orictites ampliosetosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A878FFDFD7EDFFC68CA0FFD4" box="[397,485,93,117]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A878FFDFD619FF338D9FFE01" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="AD4F195BA878FFDFD619FF338DA5FEAB" blockId="43.[121,1006,168,453]" pageId="43" pageNumber="44">
Type material:
<materialsCitation id="1D981306A878FFDFD744FF338DA5FEAB" ID-GBIF-Occurrence="3497007315" collectingDate="1993-07-27" collectingDateMax="1993-07-31" collectingDateMin="1993-07-27" collectionCode="NHMW" collectorName="Schuh" country="Malaysia" elevation="1600" location="Pahang Cameron Highlands Umg. Tanah Rata" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="holotype">
<typeStatus id="724BA7F9A878FFDFD744FF338CD5FF61" box="[292,400,168,192]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A878FFDFD7FBFF338CEAFF1E" box="[411,431,168,191]" pageId="43" pageNumber="44" type="male">♂</specimenCount>
, labels: white, black printed: “
<collectingCountry id="D5E759CBA878FFDFD56EFF338ECBFF61" box="[782,910,168,192]" name="Malaysia" pageId="43" pageNumber="44">MALAYSIA</collectingCountry>
:
<location id="A82F4F80A878FFDFD5F9FF338F00FF44" LSID="urn:lsid:plazi:treatment:2559A84DA878FFD9D619FFC68E60FA14:A82F4F80A878FFDFD5F9FF338F00FF44" country="Malaysia" name="Pahang Cameron Highlands Umg. Tanah Rata" pageId="43" pageNumber="44" stateProvince="Pahang">Pahang Cameron Highlands Umg. Tanah Rata</location>
,
<quantity id="6A08B4BEA878FFDFD436FF568FF1FF44" box="[598,692,205,229]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="43" pageNumber="44" unit="m" value="1600.0">
<elevation id="26DDFE68A878FFDFD436FF568FF1FF44" box="[598,692,205,229]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="43" pageNumber="44" unit="m" value="1600.0">1600 m</elevation>
</quantity>
<date id="D94E3F9BA878FFDFD4A2FF568E24FF44" box="[706,865,205,229]" pageId="43" pageNumber="44" value="1993-07-27" valueMax="1993-07-31" valueMin="1993-07-27">
<collectingDate id="C90AC673A878FFDFD4A2FF568E24FF44" box="[706,865,205,229]" pageId="43" pageNumber="44" value="1993-07-27" valueMax="1993-07-31" valueMin="1993-07-27">27.–31.7.1993</collectingDate>
</date>
lg.
<collectorName id="00057C8DA878FFDFD5F9FF568EA0FF44" box="[921,997,205,229]" pageId="43" pageNumber="44">Schuh</collectorName>
” (
<collectionCode id="CBE1819EA878FFDFD61FFF698D93FEAB" box="[127,214,242,266]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="43" pageNumber="44" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="AD4F195BA878FFDFD619FE838D9FFE01" blockId="43.[121,1006,168,453]" pageId="43" pageNumber="44">
<materialsCitation id="1D981306A878FFDFD619FE838FFDFEF4" ID-GBIF-Occurrence="3497007303" collectingDate="1990-04-01" collectingDateMax="1990-04-03" collectingDateMin="1990-04-01" collectionCode="CBM-ZSM" collectorName="Riedel" country="Malaysia" location="Cameron Highl. Gn. Beremban" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="724BA7F9A878FFDFD619FE838DB1FE91" box="[121,244,280,304]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="BBF6D2D2A878FFDFD765FE838C73FE8E" box="[261,310,280,304]" pageId="43" pageNumber="44" type="male">1 ♂</specimenCount>
, Labels: white, black printed: “
<collectingCountry id="D5E759CBA878FFDFD4A1FE838E6EFE91" box="[705,811,280,304]" name="Malaysia" pageId="43" pageNumber="44">Malaysia</collectingCountry>
,
<location id="A82F4F80A878FFDFD54EFE838C56FEF4" LSID="urn:lsid:plazi:treatment:2559A84DA878FFD9D619FFC68E60FA14:A82F4F80A878FFDFD54EFE838C56FEF4" country="Malaysia" name="Cameron Highl. Gn. Beremban" pageId="43" pageNumber="44">Cameron Highl. Gn.Beremban</location>
<date id="D94E3F9BA878FFDFD77BFEA68CE4FEF4" box="[283,417,317,341]" pageId="43" pageNumber="44" value="1990-04-01" valueMax="1990-04-03" valueMin="1990-04-01">
<collectingDate id="C90AC673A878FFDFD77BFEA68CE4FEF4" box="[283,417,317,341]" pageId="43" pageNumber="44" value="1990-04-01" valueMax="1990-04-03" valueMin="1990-04-01">1.–3.4.1990</collectingDate>
</date>
leg.
<collectorName id="00057C8DA878FFDFD7BAFEA68F60FEF4" box="[474,549,317,341]" pageId="43" pageNumber="44">Riedel</collectorName>
” (
<collectionCode id="CBE1819EA878FFDFD45AFEA68FF7FEF4" box="[570,690,317,341]" pageId="43" pageNumber="44">CBM-ZSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="1D981306A878FFDFD4A2FEA68D90FE01" ID-GBIF-Occurrence="3497007337" collectingDate="1993-04-22" collectingDateMax="1993-04-28" collectingDateMin="1993-04-22" collectingMethod="light trap" collectionCode="MHNG" collectorName="LObl &amp; Calama" country="Malaysia" elevation="1500" location="23 Cameron Highls" pageId="43" pageNumber="44" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="paratype">
<specimenCount id="BBF6D2D2A878FFDFD4A2FEA68FA9FEF4" box="[706,748,317,341]" pageId="43" pageNumber="44" type="female">1 ♀</specimenCount>
: “W.
<collectingCountry id="D5E759CBA878FFDFD541FEA68ECAFEF4" box="[801,911,317,341]" name="Malaysia" pageId="43" pageNumber="44">Malaysia</collectingCountry>
:
<collectingRegion id="6F34D7B9A878FFDFD5F9FEA68EABFEF4" box="[921,1006,317,341]" country="Malaysia" name="Pahang" pageId="43" pageNumber="44">Pahang</collectingRegion>
#
<location id="A82F4F80A878FFDFD6EEFEF98C22FEDB" LSID="urn:lsid:plazi:treatment:2559A84DA878FFD9D619FFC68E60FA14:A82F4F80A878FFDFD6EEFEF98C22FEDB" box="[142,359,354,378]" country="Malaysia" name="23 Cameron Highls" pageId="43" pageNumber="44" stateProvince="Pahang">23 Cameron Highls</location>
.
<quantity id="6A08B4BEA878FFDFD711FEF98CFAFEDB" box="[369,447,354,378]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="m" value="1500.0">
<elevation id="26DDFE68A878FFDFD711FEF98CFAFEDB" box="[369,447,354,378]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="43" pageNumber="44" unit="m" value="1500.0">1500m</elevation>
</quantity>
<location id="A82F4F80A878FFDFD7A5FEF98F02FEDB" LSID="urn:lsid:plazi:treatment:2559A84DA878FFD9D619FFC68E60FA14:A82F4F80A878FFDFD7A5FEF98F02FEDB" box="[453,583,354,378]" country="Malaysia" name="Tanah Rata" pageId="43" pageNumber="44" stateProvince="Pahang">Tanah Rata</location>
,
<date id="D94E3F9BA878FFDFD430FEF98F93FEDB" box="[592,726,354,378]" pageId="43" pageNumber="44" value="1993-04-22" valueMax="1993-04-28" valueMin="1993-04-22">
<collectingDate id="C90AC673A878FFDFD430FEF98F93FEDB" box="[592,726,354,378]" pageId="43" pageNumber="44" value="1993-04-22" valueMax="1993-04-28" valueMin="1993-04-22">22–28.IV.93</collectingDate>
</date>
<collectorName id="00057C8DA878FFDFD4BCFEF98E54FEDB" box="[732,785,354,378]" pageId="43" pageNumber="44">LÖbl</collectorName>
&amp;
<collectorName id="00057C8DA878FFDFD542FEF98E39FEDB" box="[802,892,354,378]" pageId="43" pageNumber="44">Calama</collectorName>
,
<collectingMethod id="74B1614CA878FFDFD51EFEF98EA0FEDB" box="[894,997,354,378]" pageId="43" pageNumber="44">light trap</collectingMethod>
” (
<collectionCode id="CBE1819EA878FFDFD61FFE138D95FE01" box="[127,208,392,416]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="43" pageNumber="44" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A878FFD9D619FE368F20FBC4" lastPageId="45" lastPageNumber="46" pageId="43" pageNumber="44" type="description">
<paragraph id="AD4F195BA878FFDFD619FE368E34FE64" blockId="43.[121,1006,168,453]" box="[121,881,429,453]" pageId="43" pageNumber="44">
Remark: In one of the
<typeStatus id="724BA7F9A878FFDFD710FE368CA2FE64" box="[368,487,429,453]" pageId="43" pageNumber="44" type="paratype">paratypes</typeStatus>
, parts of the antennae are missing.
</paragraph>
<paragraph id="AD4F195BA878FFDFD619FD868F05FD94" blockId="43.[121,576,541,566]" box="[121,576,541,566]" pageId="43" pageNumber="44">
<emphasis id="9F84C549A878FFDFD619FD868DB9FD97" bold="true" box="[121,252,541,566]" pageId="43" pageNumber="44">Description</emphasis>
(
<figureCitation id="35CB05DEA878FFDFD768FD868C08FD94" box="[264,333,541,565]" captionStart="Figs 5–8" captionStartId="8.[159,196,1494,1512]" captionTargetBox="[159,1021,146,1417]" captionTargetId="figure-66@8.[159,590,846,1416]" captionTargetPageId="8" captionText="Figs 5–8: Orictites spp., habitus. 5: O. plurisetosus sp. nov., holotype, male; 6: O. anteriortenuis sp. nov., holotype, male; 7: O. omnipunctatus sp. nov., holotype, male; 8: O. ampliosetosus sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284326" httpUri="https://zenodo.org/record/6284326/files/figure.png" pageId="43" pageNumber="44">Figs 8</figureCitation>
,
<figureCitation id="35CB05DEA878FFDFD73BFD868C32FD94" box="[347,375,541,565]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="43" pageNumber="44">26</figureCitation>
,
<figureCitation id="35CB05DEA878FFDFD7E4FD868CE0FD94" box="[388,421,541,565]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="43" pageNumber="44">37</figureCitation>
; pp. 9, 24, 36)
</paragraph>
<paragraph id="AD4F195BA878FFDFD619FDF38F21FD6B" blockId="43.[121,1006,615,1573]" pageId="43" pageNumber="44">
Measurements (n = 3): Body length
<quantity id="6A08B4BEA878FFDFD46FFDF38FD5FD21" box="[527,656,616,640]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.1" metricValueMin="5.9" pageId="43" pageNumber="44" unit="mm" value="6.5" valueMax="7.1" valueMin="5.9">5.9–7.1 mm</quantity>
(
<emphasis id="9F84C549A878FFDFD4FFFDFC8FEAFD21" bold="true" box="[671,687,615,640]" italics="true" pageId="43" pageNumber="44"></emphasis>
=
<quantity id="6A08B4BEA878FFDFD4ADFDF38E70FD21" box="[717,821,616,640]" metricMagnitude="-3" metricUnit="m" metricValue="6.64" pageId="43" pageNumber="44" unit="mm" value="6.64">6.64 mm</quantity>
), width
<quantity id="6A08B4BEA878FFDFD5EBFDF38DE6FD04" metricMagnitude="-3" metricUnit="m" metricValue="1.725" metricValueMax="1.85" metricValueMin="1.6" pageId="43" pageNumber="44" unit="mm" value="1.725" valueMax="1.85" valueMin="1.6">1.6–1.85 mm</quantity>
(
<emphasis id="9F84C549A878FFDFD6D3FD178D86FD04" bold="true" box="[179,195,652,677]" italics="true" pageId="43" pageNumber="44"></emphasis>
=
<quantity id="6A08B4BEA878FFDFD682FD168C0CFD04" box="[226,329,653,677]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="43" pageNumber="44" unit="mm" value="1.75">1.75 mm</quantity>
), ratio length/width of pronotum 0.87–1.01 (
<emphasis id="9F84C549A878FFDFD521FD178E14FD04" bold="true" box="[833,849,652,677]" italics="true" pageId="43" pageNumber="44"></emphasis>
= 0.96), ratio length/width of elytra 1.95–2.07 (
<emphasis id="9F84C549A878FFDFD795FD2A8F40FD6B" bold="true" box="[501,517,689,714]" italics="true" pageId="43" pageNumber="44"></emphasis>
= 1.99).
</paragraph>
<paragraph id="AD4F195BA878FFDFD6FEFD438D8DFC9B" blockId="43.[121,1006,615,1573]" pageId="43" pageNumber="44">Colour: Shiny. Black. Wings of clypeus and supraantennal plates slightly transparent brown. Labrum black. Antennae and legs dark-brown, palpi red brown.</paragraph>
<paragraph id="AD4F195BA878FFD8D6FEFCD38C86FD6B" blockId="43.[121,1006,615,1573]" lastBlockId="44.[159,1044,93,1573]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
Head: A sixth narrower than pronotum. Clypeus with small rounded tooth laterally, anterior margin slightly excised, with blunt raised knob in between the two clypeal setae, anteriorly smooth, posterior part of knob covered with big punctures, margin of wings convex, slightly hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, smooth, with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by flat transverse depression with rugae, with costate, nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina, frons moderately convex, smooth. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes relatively small, moderately convex, genae enlarged, rounded, not as high as and shorter than eyes, enclosing eyes posteriorly, forming obtuse angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked as distinct step, with bigger punctures partly connected, neck laterally covered with reticulation and densely scattered medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A87FFFD8D733FE8F8C2FFE86" attach="left" box="[339,362,276,295]" fontSize="8" pageId="44" pageNumber="45">nd</superScript>
segment, 2
<superScript id="5A85B413A87FFFD8D792FE8F8F4CFE86" attach="left" box="[498,521,276,295]" fontSize="8" pageId="44" pageNumber="45">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with a few punctures and reticulation. Submentum and mentum distinctly separated; submentum with four, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae relatively long, reaching up to posterior third of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 elongate (L/
<geoCoordinate id="C8C47F9CA87FFFD8D532FDF38ED2FD21" box="[850,919,616,640]" degrees="1. 5" direction="west" orientation="longitude" pageId="44" pageNumber="45" precision="5555" value="-1.5">W 1.5</geoCoordinate>
), antennomeres cylindrical, with shiny areas, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA87FFFD8D6A4FD438C2DFA81" blockId="44.[159,1044,93,1573]" pageId="44" pageNumber="45">Pronotum: Square, slightly wider than long, sides straight, rounded at anterior angles. Anterior margin indistinctly bisinuate. Reflexed lateral margin smooth, convex for a short distance before posterior angles. Margin from posterior angle to base running as straight line. Lateral channel conspicuously broad between the two lateral setigerous punctures, with isodiametric reticulation and very few irregular punctures. Anterior setigerous puncture located at the end of anterior quarter adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angles distinctly projecting, rounded, formed by reflexed lateral margin, posterior angle developed as sharp distinct tooth, projecting laterally. Anterior transverse line narrow, formed by distinct, partly connected punctures. Median line deep and conspicuously broad, nearly smooth, surpassing anterior transverse line without joining, fine at base, not adjoining base. Surface bilaterally with two irregularly longitudinal groups of punctures with slight basal impression. Basal channel broad, deep. Flange raised keellike (lateral view).</paragraph>
<paragraph id="AD4F195BA87FFFD9D6A4FAB68F19FEAB" blockId="44.[159,1044,93,1573]" lastBlockId="45.[121,1006,93,1461]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of punctures laterally. Reflexed margin with some scars in anterior third, margin slightly thickened at humerus, bending with channel over rounded humerus up to 5
<superScript id="5A85B413A87FFFD8D54FFA248E07FA73" attach="left" box="[815,834,1471,1490]" fontSize="8" pageId="44" pageNumber="45">th</superScript>
stria. No tooth at humerus. Basal tubercle distinct, with setigerous puncture, situated at declivity of first stria. Striae 1–4 free at base, all striae deep, distinctly punctate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura raised. All intervals conspicuously convex, thought costate, 3
<superScript id="5A85B413A87EFFD9D448FF3F8F7EFF16" attach="left" box="[552,571,164,183]" fontSize="8" pageId="45" pageNumber="46">rd</superScript>
broader and distinctly raised at base, with tubercle at base, 8
<superScript id="5A85B413A87EFFD9D7EBFF518CDBFF7C" attach="left" box="[395,414,202,221]" fontSize="8" pageId="45" pageNumber="46">th</superScript>
carinate. All intervals shiny on disc. Interval 3 with 6–7 setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA87EFFD9D6FEFE838CA6FE91" blockId="45.[121,1006,93,1461]" box="[158,483,280,304]" pageId="45" pageNumber="46">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA87EFFD9D6FEFEA68FC8FD94" blockId="45.[121,1006,93,1461]" pageId="45" pageNumber="46">
Ventral surface: Epipleuron shiny, covered with big scattered punctures and indistinct reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle, mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with punctures at base, denser laterally, 3
<superScript id="5A85B413A87EFFD9D459FE548F09FE43" attach="left" box="[569,588,463,482]" fontSize="8" pageId="45" pageNumber="46">rd</superScript>
to 5
<superScript id="5A85B413A87EFFD9D4E8FE548FDEFE43" attach="left" box="[648,667,463,482]" fontSize="8" pageId="45" pageNumber="46">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA87EFFD9D6FEFDD98CDDFD51" blockId="45.[121,1006,93,1461]" pageId="45" pageNumber="46">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, smooth, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA87EFFD9D6FEFD668CD0FC24" blockId="45.[121,1006,93,1461]" pageId="45" pageNumber="46">
Male genitalia (
<figureCitation id="35CB05DEA87EFFD9D732FD668CE7FCB4" box="[338,418,765,789]" captionStart="Figs 21–26" captionStartId="23.[121,158,1467,1485]" captionTargetBox="[120,1005,145,1417]" captionTargetPageId="23" captionText="Figs 21–26: Orictites spp., aedeagus, ventral view. 21: O. mjoebergi LouWERENS, 1964, lectotype, scale bar 0.2 mm; 22: O. desuntsetosus sp. nov., holotype, scale bar 0.15 mm; 23: O. plurisetosus sp. nov., holotype, scale bar 0.2 mm; 24: O. anteriortenuis sp. nov., holotype, scale bar 0.15 mm; 25: O. omnipunctatus sp. nov., holotype, scale bar 0.2 mm; 26: O. ampliosetosus sp. nov., holotype, scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6284350" httpUri="https://zenodo.org/record/6284350/files/figure.png" pageId="45" pageNumber="46">Fig. 26</figureCitation>
): Relatively short, moderately sclerotized. Median lobe broad, moderately arcuate, moderately distorted. Endophallus strongly folded, teeth not visible. Parameres of about same length, both slender, slightly twisted, asetose.
</paragraph>
<paragraph id="AD4F195BA87EFFD9D6FEFC098E2AFC54" blockId="45.[121,1006,93,1461]" pageId="45" pageNumber="46">
Female genitalia (
<figureCitation id="35CB05DEA87EFFD9D713FC098C82FC0B" box="[371,455,914,938]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="45" pageNumber="46">Fig. 37</figureCitation>
): Coxostylus slightly sigmoid, dorso-ventrally moderately depressed, gently curved to apex, with one seta at apex, with one short but robust ensiform seta, with six nematiform setae medially.
</paragraph>
<paragraph id="AD4F195BA87EFFD9D6FEFB998F20FBC4" blockId="45.[121,1006,93,1461]" pageId="45" pageNumber="46">
Variation: Of the
<specimenCount id="BBF6D2D2A87EFFD9D703FB998F5BFBBB" box="[355,542,1026,1050]" pageId="45" pageNumber="46" type="generic">three specimens</specimenCount>
investigated one exhibits six setigerous punctures on the 3
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interval of each elytra, one seven on each elytra, and one six on one side and seven on the other side.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87EFFD9D6FEFBE98E67FA81" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="AD4F195BA87EFFD9D6FEFBE98E67FA81" blockId="45.[121,1006,93,1461]" pageId="45" pageNumber="46">Diagnosis: A medium-sized black species with distinct knob on the clypeus and distinct flat transverse depression behind, no humeral tooth, and a tubercle at the base of interval 3 of the elytra. Distinguished from all other species by the presence of 6–7 setigerous punctures on interval 3, and the conspicuously broad median line of the pronotum not adjoining base.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87EFFD9D6FEFAB68FAEFACB" pageId="45" pageNumber="46" type="distribution">
<paragraph id="AD4F195BA87EFFD9D6FEFAB68FAEFACB" blockId="45.[121,1006,93,1461]" pageId="45" pageNumber="46">
Distribution: The species is found in the Cameron Highlands of
<collectingCountry id="D5E759CBA87EFFD9D5E8FAB68EA8FAE4" box="[904,1005,1325,1349]" name="Malaysia" pageId="45" pageNumber="46">Malaysia</collectingCountry>
and the material was collected at an altitude of
<quantity id="6A08B4BEA87EFFD9D4EFFAC98FA2FACB" box="[655,743,1362,1386]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="45" pageNumber="46" unit="m" value="1200.0">1200 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87EFFD9D6FEFAE38E60FA14" pageId="45" pageNumber="46" type="etymology">
<paragraph id="AD4F195BA87EFFD9D6FEFAE38E60FA14" blockId="45.[121,1006,93,1461]" pageId="45" pageNumber="46">
Etymology: The name refers to the additional setae on the 3
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interval of the elytra, which differs from the usual pattern of four setae.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="3B70C8F7D9840B6EFFA984597B2EFE52">Revision of the genus Orictites ANDREWES, 1931 (Coleoptera, Carabidae, Clivinini)</mods:title>
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<subSubSection id="E5EA4AD0A87DFFDAD6FFFFC68CFAFFD4" box="[159,447,93,118]" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="AD4F195BA87DFFDAD6FFFFC68CFAFFD4" blockId="46.[159,447,93,118]" box="[159,447,93,118]" pageId="46" pageNumber="47">
<heading id="F607AE37A87DFFDAD6FFFFC68CFAFFD4" box="[159,447,93,118]" centered="true" fontSize="12" level="2" pageId="46" pageNumber="47" reason="2">
<taxonomicName id="6AF062D8A87DFFDAD6FFFFC68C24FFD7" ID-CoL="8MG4G" authorityName="Balkenohl" authorityYear="2017" box="[159,353,93,118]" class="Insecta" family="Carabidae" genus="Orictites" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="barclayi" status="sp. nov.">
<emphasis id="9F84C549A87DFFDAD6FFFFC68C24FFD7" bold="true" box="[159,353,93,118]" italics="true" pageId="46" pageNumber="47">Orictites barclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84B77832A87DFFDAD708FFC68CFAFFD4" box="[360,447,93,117]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87DFFDAD6FFFF338CF9FEDB" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="AD4F195BA87DFFDAD6FFFF338CF9FEDB" blockId="46.[159,1044,168,453]" pageId="46" pageNumber="47">
Type material:
<materialsCitation id="1D981306A87DFFDAD734FF338CF9FEDB" ID-GBIF-Occurrence="3497007333" collectingDate="1921-11-15" collectionCode="BMNH" collectorName="H. M. Pendlebury" location="MALAYA SELANGOR" municipality="Gombak Valley" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="MALAYA SELANGOR" typeStatus="holotype">
<typeStatus id="724BA7F9A87DFFDAD734FF338C87FF61" box="[340,450,168,192]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
:
<specimenCount id="BBF6D2D2A87DFFDAD7B2FF338CA3FF1E" box="[466,486,168,191]" pageId="46" pageNumber="47" type="female">♀</specimenCount>
, labels: beige, black printed and handwritten in black ink: “
<collectingRegion id="6F34D7B9A87DFFDAD75EFF568F6BFF44" box="[318,558,205,229]" pageId="46" pageNumber="47">MALAYA SELANGOR</collectingRegion>
, F.M.S. Gombak Valley Gd.
<date id="D94E3F9BA87DFFDAD510FF568EB4FF44" box="[880,1009,205,229]" pageId="46" pageNumber="47" value="1921-11-15">
<collectingDate id="C90AC673A87DFFDAD510FF568EB4FF44" box="[880,1009,205,229]" pageId="46" pageNumber="47" value="1921-11-15">11 15 1921</collectingDate>
</date>
” / backside printed: “
<collectorName id="00057C8DA87DFFDAD717FF698F77FEAB" box="[375,562,242,266]" pageId="46" pageNumber="47">H.M.Pendlebury</collectorName>
Coll: F.M.S.Museums.” / beige, handwritten in black ink and printed: “
<taxonomicName id="6AF062D8A87DFFDAD794FE838E96FE91" authority="Bates H. E. Andrewes" box="[500,979,280,304]" class="Insecta" family="Carabidae" genus="Clivina" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="costulipennis">Clivina costulipennis Bates H.E.Andrewes</taxonomicName>
det.” / small, beige, black printed: “346” / white, black printed: “Ex F.M.S.Museum: B.M.1955–354.” (
<collectionCode id="CBE1819EA87DFFDAD701FEF98CF4FEDB" box="[353,433,354,378]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="46" pageNumber="47">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87DFFDAD6FFFE138E5CFE64" pageId="46" pageNumber="47" type="discussion">
<paragraph id="AD4F195BA87DFFDAD6FFFE138E5CFE64" blockId="46.[159,1044,168,453]" pageId="46" pageNumber="47">Remark: The specimen was pinned centrally in between the two elytra, so the elytra are opened. It has been mounted on a paper card.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A87DFFC4D6FFFD868FDBFCC1" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
<paragraph id="AD4F195BA87DFFDAD6FFFD868F66FD94" blockId="46.[159,547,541,566]" box="[159,547,541,566]" pageId="46" pageNumber="47">
<emphasis id="9F84C549A87DFFDAD6FFFD868C67FD97" bold="true" box="[159,290,541,566]" pageId="46" pageNumber="47">Description</emphasis>
(
<figureCitation id="35CB05DEA87DFFDAD74EFD868C37FD94" box="[302,370,541,565]" captionStart="Figs 9–12" captionStartId="9.[121,158,1494,1512]" captionTargetBox="[145,998,148,1407]" captionTargetId="figure-60@9.[567,998,842,1405]" captionTargetPageId="9" captionText="Figs 9–12: Orictites spp., habitus. 9: O. barclayi sp. nov., holotype, female; 10: O. brancuccii sp. nov., holotype, male; 11: O. barclayi minorfrater ssp. nov., holotype, male; 12: O. tubercucollis sp. nov., holotype, male." figureDoi="http://doi.org/10.5281/zenodo.6284330" httpUri="https://zenodo.org/record/6284330/files/figure.png" pageId="46" pageNumber="47">Figs 9</figureCitation>
,
<figureCitation id="35CB05DEA87DFFDAD71FFD868CE5FD94" box="[383,416,541,565]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="46" pageNumber="47">38</figureCitation>
; pp. 10, 36)
</paragraph>
<paragraph id="AD4F195BA87DFFDAD6FFFDF38FDBFD04" blockId="46.[159,1044,616,1573]" pageId="46" pageNumber="47">
Measurements: Body length
<quantity id="6A08B4BEA87DFFDAD786FDF38F7AFD21" box="[486,575,616,640]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="46" pageNumber="47" unit="mm" value="4.8">4.8 mm</quantity>
, width 1,4 mm; ratio length/width of pronotum 1.03; ratio length/width of elytra 1.93.
</paragraph>
<paragraph id="AD4F195BA87DFFDAD6A4FD298DABFCB4" blockId="46.[159,1044,616,1573]" pageId="46" pageNumber="47">Colour: Shiny. Black. Wings of clypeus and supraantennal plates slightly transparent brown. Labrum black. Antennae and legs dark-brown, palpi red brown.</paragraph>
<paragraph id="AD4F195BA87DFFDBD6A4FCB98E43FD21" blockId="46.[159,1044,616,1573]" lastBlockId="47.[121,1006,93,1573]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
Head: A fifth narrower than pronotum. Clypeus with small rounded tooth laterally, anterior margin excised, with blunt raised knob in between the two clypeal setae, anteriorly smooth, posterior part of knob covered with big punctures, margin of wings convex, slightly hollowed out, divided from middle part of clypeus by slight notches and carinae, divided from supraantennal plates by distinct obtusely angled notches and ridge, clypeus and wings reflexed margined, supraantennal plates vaulted up to mid-eye level, smooth, with reflexed rounded margin, elongated as supraorbital carina, frons separated from clypeus by flat transverse depression with punctures, with costate, nearly parallel running frontal carinae at each side posteriorly, with broad furrow between frontal carina and supraorbital carina, sharp furrow between frons and frontal carina, frons moderately convex, smooth. Supraorbital setae located in broad furrows between supraorbital carina and frontal carina, the posterior one arising from tubercle. Eyes subglobose, genae indistinct, enclosing eyes posteriorly, separated from eye by small ridge, forming obtuse angle at neck. Grooves for reception of antennae short, length 0.4 of eye diameter. Neck constriction marked by band of big punctures, neck laterally covered with reticulation and densely scattered medium sized punctures. Labrum straight, 7-setose, ciliate laterally, with isodiametric reticulation. Mandibles robust, shorter than head, flattened, stout, broadened at base, acutely curved at apex, carinae of scrobe complete, both mandibles obtusely angled towards base (opened position), both mandibles somewhat hollowed out dorsally. Maxilla distinctly curved, acutely hooked at apex. Apical maxillary palpomere slender, slightly securiform. Apical labial palpomere slender, straight, slightly longer than 2
<superScript id="5A85B413A87CFFDBD74DFF518C01FF7C" attach="left" box="[301,324,202,221]" fontSize="8" pageId="47" pageNumber="48">nd</superScript>
segment, 2
<superScript id="5A85B413A87CFFDBD7ACFF518CA6FF7C" attach="left" box="[460,483,202,221]" fontSize="8" pageId="47" pageNumber="48">nd</superScript>
segment bisetose. Ligula with one long seta; paraglossae slender. Ventral surface of neck with punctures and reticulation. Submentum and mentum distinctly separated; submentum with four, about equally spaced setigerous punctures; mentum at base bilaterally with rounded elevation, with some longitudinal carinae, with fine and complete reflexed margin, lateral lobes projecting, nearly right-angled at tips, with a seta near base of each lobe, median tooth forming broad obtuse triangle, bisetose, not projecting as far as lateral lobes. Antennae of moderate length, reaching up to middle of pronotum, scapus knee-like angled, with one seta dorsoapically, scapus and pedicellus with fine reticulation, segments 5–10 submoniliform (L/
<geoCoordinate id="C8C47F9CA87CFFDBD6F6FDD98DA5FDFB" box="[150,224,578,602]" degrees="0. 9" direction="west" orientation="longitude" pageId="47" pageNumber="48" precision="5555" value="-0.9">W 0.9</geoCoordinate>
), antennomeres somewhat flattened, with shiny areas, segments 4–11 densely pubescent, with a few additional long setae.
</paragraph>
<paragraph id="AD4F195BA87CFFDBD6FEFD168C51FB11" blockId="47.[121,1006,93,1573]" pageId="47" pageNumber="48">Pronotum: Square, slightly longer than wide, sides slightly convex. Anterior margin straight. Reflexed lateral margin smooth. Margin from posterior angle to base running as straight line. Lateral channel conspicuously broad between the two lateral setigerous punctures, with isodiametric reticulation and very few irregular punctures. Anterior setigerous puncture located at the end of anterior quarter adjoining convexity of pronotum, the posterior one located at level of posterior angle, removed from lateral channel by diameter of pore. Anterior angle indistinctly projecting, formed by reflexed lateral margin, posterior angle developed as sharp distinct tooth, projecting laterally. Anterior transverse line narrow, formed by distinct, partly connected punctures. Median line deep and moderately narrow, with some punctures, surpassing anterior transverse line without joining, fine at base, adjoining base. Surface bilaterally with two irregularly longitudinal groups of punctures with distinct basal impression. Basal channel broad, deep. Flange raised keel-like (lateral view).</paragraph>
<paragraph id="AD4F195BA87CFFC4D6FEFB268F17FF3B" blockId="47.[121,1006,93,1573]" lastBlockId="48.[159,1044,90,1162]" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48">
Elytra: Subcylindrical, sides slightly diverging, marginal channel broad, visible from above, with uninterrupted row of big setigerous punctures arising from broad tubercles, with additional row of punctures laterally. Reflexed margin with some scars in anterior third, margin slightly thickened at humerus, bending with channel over rounded humerus up to 5
<superScript id="5A85B413A87CFFDBD56AFAD48E58FAC3" attach="left" box="[778,797,1359,1378]" fontSize="8" pageId="47" pageNumber="48">th</superScript>
stria. No tooth at humerus. Basal tubercle distinct, with setigerous puncture, situated at declivity of 1
<superScript id="5A85B413A87CFFDBD6A7FA018D9DFA0C" attach="left" box="[199,216,1434,1453]" fontSize="8" pageId="47" pageNumber="48">st</superScript>
stria. Striae 1–4 free at base, all striae deep, distinctly punctate, striae 1–2, 3–4 and 5–6 joined apically; striae 6 and 7 shortened at humerus. Striole distinct, interval between striole and sutura raised. All intervals conspicuously convex, thought costate, 3
<superScript id="5A85B413A87CFFDBD78EF9918F44F9BC" attach="left" box="[494,513,1546,1565]" fontSize="8" pageId="47" pageNumber="48">rd</superScript>
broader and distinctly raised at base, with tubercle at base, 8
<superScript id="5A85B413A863FFC4D714FFC18CC2FFCC" attach="left" box="[372,391,90,109]" fontSize="8" pageId="48" pageNumber="49">th</superScript>
carinate. All intervals shiny on disc. Interval 3 with seven setigerous punctures adjoining stria 3.
</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FF338F4DFF61" blockId="48.[159,1044,90,1162]" box="[196,520,168,192]" pageId="48" pageNumber="49">Hind wings: Fully developed.</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FF568FF6FE64" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">
Ventral surface: Epipleuron shiny, covered with big scattered punctures and indistinct reticulation, submarginal furrow distinct. Proepisternum and episternum nearly smooth, prosternite smooth, keeled at middle, mesosternum smooth. Sternites with transverse to isodiametric reticulation, each with big punctures at base, denser laterally, 3
<superScript id="5A85B413A863FFC4D414FEC48FC2FED3" attach="left" box="[628,647,351,370]" fontSize="8" pageId="48" pageNumber="49">rd</superScript>
to 5
<superScript id="5A85B413A863FFC4D4DBFEC48F8BFED3" attach="left" box="[699,718,351,370]" fontSize="8" pageId="48" pageNumber="49">th</superScript>
with paralateral ambulatory setae at each side, ventral strigae distinct, sternite 6 smooth at middle, with two widely separated apical setae at each side.
</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FE498CFBFD21" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">Legs: Anterior tibia with strong digitation, with two big and one small lateral preapical denticles, apical spine curved distinctly towards ventral, no dorsal furrow, reticulated, basal tarsomere elongated, as long as the following four together. Mesotibia with preapical spine, furnished with strong seta. All tarsomeres relatively broad.</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FD168CA0FD04" blockId="48.[159,1044,90,1162]" box="[196,485,653,677]" pageId="48" pageNumber="49">Male genitalia: Unknown.</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FD298F5DFC9B" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">
Female genitalia (
<figureCitation id="35CB05DEA863FFC4D7F9FD298CABFD6B" box="[409,494,690,714]" captionStart="Figs 35–38" captionStartId="35.[121,158,1306,1324]" captionTargetBox="[121,1005,147,1260]" captionTargetId="figure-64@35.[121,552,811,1130]" captionTargetPageId="35" captionText="Figs 35–38: Orictites spp., coxostylus. 35: O. anteriortenuis sp. nov., scale bar 0.1 mm; 36: O. omnipunctatus sp. nov., scale bar 0.1 mm; 37: O. ampliosetosus sp. nov., scale bar 0.1 mm; 38: O. barclayi sp. nov., holotype, scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6284368" httpUri="https://zenodo.org/record/6284368/files/figure.png" pageId="48" pageNumber="49">Fig. 38</figureCitation>
): Coxostylus slender, slightly sigmoid, gently curved to apex, with short obtuse knob-like tooth dorsally at base, with two setae apically tight together, with one short robust ensiform seta, with five nematiforme setae. Ramus short.
</paragraph>
<paragraph id="AD4F195BA863FFC4D6A4FCD38FDBFCC1" blockId="48.[159,1044,90,1162]" box="[196,670,840,864]" pageId="48" pageNumber="49">Variation: No bilateral variation observed.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A863FFC4D6A4FCF68E4DFBBB" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="AD4F195BA863FFC4D6A4FCF68E4DFBBB" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">Diagnosis: A medium-sized black species with distinct knob on the clypeus and distinct flat transverse depression behind, no humeral tooth, and a tubercle at the base of interval 3 of the elytra. Distinguished from all other species by the presence of seven setigerous punctures on interval 3, and the moderately narrow median line of the pronotum joining base.</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A863FFC4D6A4FBB38FEAFBE1" box="[196,687,1064,1088]" pageId="48" pageNumber="49" type="distribution">
<paragraph id="AD4F195BA863FFC4D6A4FBB38FEAFBE1" blockId="48.[159,1044,90,1162]" box="[196,687,1064,1088]" pageId="48" pageNumber="49">
Distribution: Known from West of
<collectingCountry id="D5E759CBA863FFC4D422FBB38FEEFBE1" box="[578,683,1064,1088]" name="Malaysia" pageId="48" pageNumber="49">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E5EA4AD0A863FFC4D6A4FBD68F9EFB2B" pageId="48" pageNumber="49" type="etymology">
<paragraph id="AD4F195BA863FFC4D6A4FBD68F9EFB2B" blockId="48.[159,1044,90,1162]" pageId="48" pageNumber="49">Etymology. The name is a patronym of Maxwell V. L. Barclay (BMNH) who supported my work over the last time significantly.</paragraph>
</subSubSection>
</treatment>
</document>

121
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<mods:title>Extension of the phasmid genus Presbistus to Cambodia with a new species and notes on genitalia and captive breeding (Phasmida, Aschiphasmatidae, Aschiphasmatinae)</mods:title>
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<mods:namePart>Bresseel, Joachim</mods:namePart>
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Genus
<taxonomicName LSID="2559AD6F-21BC-53ED-B2EB-FA48CAE73308" authority="Kirby, 1896" authorityName="Kirby" authorityYear="1896" class="Insecta" family="Aschiphasmatidae" genus="Presbistus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Presbistus" order="Phasmida" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">Presbistus Kirby, 1896</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="105" type="reference_group">
<paragraph pageId="0" pageNumber="105">
<taxonomicName authorityName="Kirby" authorityYear="1896" class="Insecta" family="Aschiphasmatidae" genus="Presbistus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Presbistus" order="Phasmida" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">Presbistus</taxonomicName>
Kirby, 1896: 475. -
<bibRefCitation author="Kirby, WF" journalOrPublisher="British Museum, London" pageId="0" pageNumber="105" refId="B25" refString="Kirby, WF, 1904. A Synonymic Catalogue of Orthoptera. 1. OrthopteraEuplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). British Museum, London" title="A Synonymic Catalogue of Orthoptera. 1. OrthopteraEuplexoptera, Cursoria et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae)." year="1904">Kirby 1904</bibRefCitation>
: 475 [in
<taxonomicName family="Aschiphasmatidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Phasmida" pageId="0" pageNumber="105" rank="subFamily" subFamily="Aschiphasminae">Aschiphasminae</taxonomicName>
]. -
<bibRefCitation author="Redtenbacher, J" journalOrPublisher="British Museum, London" pageId="0" pageNumber="105" refId="B30" refString="Redtenbacher, J, 1906. Die Insektenfamilie der Phasmiden. I. Phasmidae Areolatae. Leipzig, 180 pp." title="Die Insektenfamilie der Phasmiden. I. Phasmidae Areolatae. Leipzig, 180 pp." year="1906">Redtenbacher 1906</bibRefCitation>
: 78 [redescribed]. -
<bibRefCitation author="Ragge, DR" journalOrPublisher="British Museum, London" pageId="0" pageNumber="105" refId="B29" refString="Ragge, DR, 1955. The wing-venation of the OrthopteraSaltatoria with notes on dictyopteran wing-venation. British Museum, London" title="The wing-venation of the OrthopteraSaltatoria with notes on dictyopteran wing-venation." year="1955">Ragge 1955</bibRefCitation>
: 377 [wing venation]. -
<bibRefCitation author="Bradley, JC" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="105" pagination="176 - 208" refId="B1" refString="Bradley, JC, Galil, BS, 1977. The taxonomic arrangement of the Phasmatodea with keys to the subfamilies and tribes. Proceedings of the Entomological Society of Washington 79: 176 - 208" title="The taxonomic arrangement of the Phasmatodea with keys to the subfamilies and tribes." volume="79" year="1977">Bradley and Galil 1977</bibRefCitation>
: 200 [in
<taxonomicName authorityName="Brunner von Wattenwyl" authorityYear="1893" family="Aschiphasmatidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Phasmida" pageId="0" pageNumber="105" rank="subFamily" subFamily="Aschiphasmatinae">Aschiphasmatinae</taxonomicName>
]. -
<bibRefCitation author="Brock, PD" journalOrPublisher="Malaysian Nature Society, Kuala Lumpur" pageId="0" pageNumber="105" refId="B9" refString="Brock, PD, 1999. Stick and Leaf Insects of Peninsular Malaysia and Singapore. Malaysian Nature Society, Kuala Lumpur" title="Stick and Leaf Insects of Peninsular Malaysia and Singapore." year="1999">Brock 1999</bibRefCitation>
: 147, 154, 184 [species from Peninsular Malaysia]. -
<bibRefCitation author="Bragg, PE" journalOrPublisher="Natural History Publications (Borneo), Kota Kinabalu" pageId="0" pageNumber="105" refId="B6" refString="Bragg, PE, 2001. Phasmids of Borneo. Natural History Publications (Borneo), Kota Kinabalu" title="Phasmids of Borneo." year="2001">Bragg 2001</bibRefCitation>
: 323, 642 [species from Borneo]. - Zompro, 2004: 318 [in superfamily
<taxonomicName authorityName="Kirby" authorityYear="1896" class="Insecta" family="Aschiphasmatidae" genus="Presbistus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Phasmida" pageId="0" pageNumber="105" phylum="Arthropoda" rank="superFamily" superFamily="Aschiphasmatoidea">Aschiphasmatoidea</taxonomicName>
]. -
<bibRefCitation author="Otte, D" journalOrPublisher="The Insect Diversity Association and the Academy of Natural Sciences, Philadelphia" pageId="0" pageNumber="105" refId="B26" refString="Otte, D, Brock, PD, 2005. Phasmida Species File. Catalog of Stick and Leaf Insects of the world, 2nd edn. The Insect Diversity Association and the Academy of Natural Sciences, Philadelphia" title="Phasmida Species File. Catalog of Stick and Leaf Insects of the world, 2 nd edn." year="2005">Otte and Brock 2005</bibRefCitation>
: 280 [Catalogued]. -
<bibRefCitation author="Seow-Choen, F" journalOrPublisher="Natural History Publications (Borneo), Kota Kinabalu" pageId="0" pageNumber="105" refId="B33" refString="Seow-Choen, F, 2016. A taxonomic guide to the stick insects of Borneo including new genera and species. Natural History Publications (Borneo), Kota Kinabalu" title="A taxonomic guide to the stick insects of Borneo including new genera and species." year="2016">Seow-Choen 2016</bibRefCitation>
: 359 [species recorded and figured from Borneo]. -
<bibRefCitation author="Seow-Choen, F" journalOrPublisher="Natural History Publications (Borneo), Kota Kinabalu" pageId="0" pageNumber="105" refId="B34" refString="Seow-Choen, F, 2017. A taxonomic guide to the stick insects of Singapore. Natural History Publications (Borneo), Kota Kinabalu" title="A taxonomic guide to the stick insects of Singapore." year="2017">Seow-Choen 2017</bibRefCitation>
: 128 [species recorded and figured from Singapore]. -
<bibRefCitation author="Seow-Choen, F" journalOrPublisher="Natural History Publications (Borneo), Kota Kinabalu" pageId="0" pageNumber="105" refId="B35" refString="Seow-Choen, F, 2018. A taxonomic guide to the stick insects of Sumatra. Volume 1. Natural History Publications (Borneo), Kota Kinabalu" title="A taxonomic guide to the stick insects of Sumatra. Volume 1." year="2018">Seow-Choen 2018</bibRefCitation>
: 577 [species recorded and figured from Sumatra].
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="105" type="type species">
<paragraph pageId="0" pageNumber="105">
<emphasis italics="true" pageId="0" pageNumber="105">Type species.</emphasis>
-
</paragraph>
<paragraph pageId="0" pageNumber="105">
<taxonomicName authorityName="G. R. Gray" authorityYear="1835" class="Insecta" family="Aschiphasmatidae" genus="Perlamorpha" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Perlamorpha peleus" order="Phasmatodea" pageId="0" pageNumber="105" phylum="Arthropoda" rank="species" species="peleus">
<emphasis italics="true" pageId="0" pageNumber="105">Perlamorpha peleus</emphasis>
</taxonomicName>
Gray, 1835 by original designation.
</paragraph>
<paragraph pageId="0" pageNumber="105">
The genus
<taxonomicName authorityName="Kirby" authorityYear="1896" class="Insecta" family="Vitaceae" genus="Presbistus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Presbistus" order="Phasmatodea" pageId="0" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="105">Presbistus</emphasis>
</taxonomicName>
can be distinguished from other members of the
<taxonomicName authorityName="Brunner von Wattenwyl" authorityYear="1893" family="Aschiphasmatidae" kingdom="Animalia" lsidName="" order="Phasmida" pageId="0" pageNumber="105" rank="subFamily" subFamily="Aschiphasmatinae">Aschiphasmatinae</taxonomicName>
by the combination of the following set of characters (adapted from
<bibRefCitation author="Bragg, PE" journalOrPublisher="Natural History Publications (Borneo), Kota Kinabalu" pageId="0" pageNumber="105" refId="B6" refString="Bragg, PE, 2001. Phasmids of Borneo. Natural History Publications (Borneo), Kota Kinabalu" title="Phasmids of Borneo." year="2001">Bragg 2001</bibRefCitation>
):
</paragraph>
<paragraph pageId="0" pageNumber="105">Body and legs brown with some black markings.</paragraph>
<paragraph pageId="0" pageNumber="105">Mesonotum unarmed.</paragraph>
<paragraph pageId="0" pageNumber="105">Profemora incurving at base; all femora with distinct medioventral carina, armed with at least a few minute spines.</paragraph>
<paragraph pageId="0" pageNumber="105">Tegmina triangular and spine-like; anal region of hind wing translucent brown.</paragraph>
<paragraph pageId="0" pageNumber="105">Apex of male abdomen distinctly swollen.</paragraph>
<paragraph pageId="0" pageNumber="105">Apex of female abdomen almost arrowhead shaped in dorsal view. Subgenital plate with anterior portion globose, posterior half medially keeled and tapering.</paragraph>
<paragraph pageId="0" pageNumber="105">Cerci cylindrical.</paragraph>
<paragraph pageId="0" pageNumber="105">Eggs longer than high, lacking setae.</paragraph>
</subSubSection>
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