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<document id="47226E22C201D584AD915377871D0869" ID-DOI="10.3897/zse.100.118522" ID-ZooBank="9D3FFC51-0277-4669-B215-23DA5A1D5483" ID-publisher-id="118522" URI-arpha="2BC007BD-0F6A-5674-86A6-B020044633CA" URI-zoobank="http://zoobank.org/9D3FFC51-0277-4669-B215-23DA5A1D5483" article-type="research-article" checkinTime="1729272491234" checkinUser="pensoft" docAuthor="Meza-Vargas, Vanessa, Ramirez, Jorge L. &amp; Lujan, Nathan K." docDate="2024" docId="A2096C2D90695535B029780B1C461296" docLanguage="en" docName="Zoosystematics and Evolution 100(4): 1387-1400" docOrigin="Zoosystematics and Evolution 100 (4)" docSource="https://zse.pensoft.net/article/118522/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Chaetostoma branickii Steindachner 1881" docType="treatment" docVersion="2" dtd-version="3.0" lastPageNumber="1400" masterDocId="2BC007BD0F6A567486A6B020044633CA" masterDocTitle="The ornate rubbernose pleco (Siluriformes, Loricariidae, Chaetostoma), a new species from the Ucayali River Basin, Peru" masterLastPageNumber="1400" masterPageNumber="1387" pageNumber="1387" updateTime="1729272892402" updateUser="ExternalLinkService">
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<mods:title id="49E4F7AD340F9927D3C8B015E8A6A2E1">The ornate rubbernose pleco (Siluriformes, Loricariidae, Chaetostoma), a new species from the Ucayali River Basin, Peru</mods:title>
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<mods:namePart id="62D6F284BDE0C96598E9437F7625D8D5">Meza-Vargas, Vanessa</mods:namePart>
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<mods:affiliation id="0EAD666CA8202435A65678DF39226485">Departamento de Ictiología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Avenida Arenales 1256, Jesus Maria, Lima 14, Peru</mods:affiliation>
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<mods:namePart id="0422F998645752ABD02CD3674E3AD2D8">Ramirez, Jorge L.</mods:namePart>
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<mods:affiliation id="A18BD6ABE194BE96178775066F8E83BC">Departamento de Ictiología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Avenida Arenales 1256, Jesus Maria, Lima 14, Peru</mods:affiliation>
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<mods:namePart id="A2A9EDF10C1B8CFEF933D1098823340B">Lujan, Nathan K.</mods:namePart>
<mods:affiliation id="5C262FA38531F25262D279B6C46EC7D0">Departamento de Biología Celular y Genética, Facultad de Ciencias Biológicas, Universidad Nacional Mayor de San Marcos, Avenida Germán Amézaga 375, Cercado De Lima, Lima, Peru &amp; Department of Natural History, Royal Ontario Museum, 100 Queens Park, Toronto, ON M 5 S 2 C 6, Canada</mods:affiliation>
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<subSubSection id="818E9C08B200CE436D31B6DD60797EC4" type="nomenclature">
<paragraph id="059EEA66BAD20600385C99641B9663DC">
<taxonomicName id="4FF181802C2CC2847007DAF0A1A4DE5A" ID-CoL="5XQF4" authority="Steindachner 1881" authorityName="Steindachner" authorityYear="1881" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="108D31274E8EEDAC28ACB8288645BC98" italics="true">Chaetostoma branickii</emphasis>
<bibRefCitation id="3C27914B1352C24340ACB4DB8C8EC36B" author="Steindachner F" journalOrPublisher="” Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften in Wien" pagination="103-146" refId="B29" refString="Steindachner F (1881) &quot; Beitrage zur Kenntniss der Flussfische Sudamerika's. II. &quot; Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften in Wien 43: 103 - 146." title="“ Beiträge zur Kenntniss der Flussfische Südamerikas. II." volume="43" year="1881">Steindachner 1881</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="3A461C4F882CBAD4E49DFF30C52EE622" type="description">
<paragraph id="3B97F5810072C400D2B8A5EEE93F650E">
<figureCitation id="6390CD32F762A7363955D39D98A2FCA1" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Original illustrations of Chaetostoma branickii from Steindachner 1881." figureDoi="10.3897/zse.100.118522.figure4" httpUri="https://binary.pensoft.net/fig/1156331">Figs 4</figureCitation>
,
<figureCitation id="0543E5F07E08BAD23E92B4E486B8CADC" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photo comparison of a nearly 150 - year-old preserved syntype of Chaetostoma branickii (NMW 47270-71, photos by Mark Sabaj &amp; Kyle Luckenbill, copyright NMW) with a live specimen collected in 2018 from the Marañon River, Cajamarca Province, Peru (ROM 109213, photos by NKL). Scale bars: 1 cm." figureDoi="10.3897/zse.100.118522.figure5" httpUri="https://binary.pensoft.net/fig/1156332">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FDF7BAF2ACC57DFCFFB236D374089EB3" type="reference_group">
<paragraph id="9709DE887C1DF5105621B2CBE9D9BBE5">
<treatmentCitationGroup id="467B76CA215CAF03380CD6CD000AF500">
<treatmentCitation id="7A899C08AAA353DEE92112A07A412916">
<taxonomicName id="72D61A4C1246743EC0F969C9FA55645B" ID-CoL="5Y2F4" authority="Regan 1904" authorityName="Regan" authorityYear="1904" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="breve">
<emphasis id="9F1CD480214899E46803B810219552FC" italics="true">Chaetostoma breve</emphasis>
<bibRefCitation id="94EC65EE14865CA245C49B4959F3BA46" DOI="10.1111/j.1096-3642.1904.tb00040.x" author="Regan" firstAuthor="Regan" journalOrPublisher="” Transactions of the Zoological Society of London" pagination="191-350" refId="B21" refString="Regan CT (1904) “ A monograph of the fishes of the family Loricariidae. ” Transactions of the Zoological Society of London 17: 191350. https://doi.org/10.1111/j.1096-3642.1904.tb00040.x" title="“ A monograph of the fishes of the family Loricariidae." volume="17" year="1904">Regan 1904</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[synonym]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6YBK" type="diagnosis">
<paragraph id="8747118B179529E7DD96B7BB77EB7E3D">
<heading id="104810CDB167960745DD8CDF024E4DA6" reason="title">Diagnosis and description.</heading>
</paragraph>
<paragraph id="1266CA9383F7EB59F74EB1CDCADE5547">
As for
<taxonomicName id="04884388A608F6A20EF585FC9F777721" authorityName="Regan" authorityYear="1904" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="breve">
<emphasis id="B113BEB0B961F60A2472D617320C0FBD" italics="true">Chaetostoma breve</emphasis>
</taxonomicName>
in
<bibRefCitation id="ECD7D443A77DA9D3E8D187E18E0F107F" DOI="10.1016/j.ympev.2014.08.020" author="Lujan NK &amp; Armbruster JW &amp; Lovejoy NR &amp; López-Fernández H" journalOrPublisher="Molecular phylogenetics and evolution" pagination="269-288" refId="B11" refString="Lujan NK, Armbruster JW, Lovejoy NR, López-Fernández H (2015 a) Multilocus molecular phylogeny of the suckermouth armored catfishes (Siluriformes: Loricariidae) with a focus on subfamily Hypostominae. Molecular phylogenetics and evolution 82: 269288. https://doi.org/10.1016/j.ympev.2014.08.020" title="Multilocus molecular phylogeny of the suckermouth armored catfishes (Siluriformes: Loricariidae) with a focus on subfamily Hypostominae." volume="82">Lujan et al. (2015 a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUZBK" type="comments">
<paragraph id="DD7FF85AA2F742B2455BB8EDBA21034F">
<heading id="0425C06C0E9469A5F3E7E9D09FC25E27" reason="title">Comments.</heading>
</paragraph>
<paragraph id="00E1DD14ACCA41444F167555DE87B557">
<bibRefCitation id="7D41412E6586BB1898472B96FB8EF3C6" DOI="10.1111/j.1096-3642.1904.tb00040.x" author="Regan CT" firstAuthor="Regan" journalOrPublisher="” Transactions of the Zoological Society of London" pagination="191-350" refId="B21" refString="Regan CT (1904) “ A monograph of the fishes of the family Loricariidae. ” Transactions of the Zoological Society of London 17: 191350. https://doi.org/10.1111/j.1096-3642.1904.tb00040.x" title="“ A monograph of the fishes of the family Loricariidae." volume="17" year="1904">Regan (1904)</bibRefCitation>
based the description of
<taxonomicName id="C33FAE5C2ACA1D658B29CE4283BD9B57" authorityName="Regan" authorityYear="1904" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="breve">
<emphasis id="CA16DB5DBD46F995B114889700D1EA3C" italics="true">Chaetostoma breve</emphasis>
</taxonomicName>
on specimens from the Zamora River in south-eastern
<collectingCountry id="54A9CF4FDEA8D6C2A3DCB80ACA52F279" name="Ecuador">Ecuador</collectingCountry>
.
<bibRefCitation id="D2169AD24A072C4F15A71F07E5E94754" DOI="10.1016/j.ympev.2014.08.020" author="Lujan NK &amp; Armbruster JW &amp; Lovejoy NR &amp; López-Fernández H" journalOrPublisher="Molecular phylogenetics and evolution" pagination="269-288" refId="B11" refString="Lujan NK, Armbruster JW, Lovejoy NR, López-Fernández H (2015 a) Multilocus molecular phylogeny of the suckermouth armored catfishes (Siluriformes: Loricariidae) with a focus on subfamily Hypostominae. Molecular phylogenetics and evolution 82: 269288. https://doi.org/10.1016/j.ympev.2014.08.020" title="Multilocus molecular phylogeny of the suckermouth armored catfishes (Siluriformes: Loricariidae) with a focus on subfamily Hypostominae." volume="82">Lujan et al. (2015 a</bibRefCitation>
) re-described
<taxonomicName id="E68042EBFC74AB0D49A8F668988C1E11" authorityName="Regan" authorityYear="1904" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="breve">
<emphasis id="C1E8D0186ADE0661BE030837DFA13F4A" italics="true">Chaetostoma breve</emphasis>
</taxonomicName>
based on specimens from throughout most of the species range, from the
<collectingRegion id="FE149993342B8A6942189A5094639F7A" country="Ecuador" name="Napo">Napo</collectingRegion>
River in the north (although the species is also now known from more northern
<collectingRegion id="7704D3D0258B8170137EF77CD1151B81" country="Colombia" name="Caqueta">Caqueta</collectingRegion>
River headwaters in
<collectingCountry id="625DB4DB482B168FE7A837BB7D619D93" name="Colombia">Colombia</collectingCountry>
;
<collectionCode id="F9DC888070F25162AB29A6CBA648784D" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" type="Museum">ROM</collectionCode>
107831, 107845) to the Marañon River in the south.
<bibRefCitation id="B2C478E687937464C41AF3E2E0B9BF01" DOI="10.1016/j.ympev.2014.08.020" author="Lujan NK &amp; Armbruster JW &amp; Lovejoy NR &amp; López-Fernández H" journalOrPublisher="Molecular phylogenetics and evolution" pagination="269-288" refId="B11" refString="Lujan NK, Armbruster JW, Lovejoy NR, López-Fernández H (2015 a) Multilocus molecular phylogeny of the suckermouth armored catfishes (Siluriformes: Loricariidae) with a focus on subfamily Hypostominae. Molecular phylogenetics and evolution 82: 269288. https://doi.org/10.1016/j.ympev.2014.08.020" title="Multilocus molecular phylogeny of the suckermouth armored catfishes (Siluriformes: Loricariidae) with a focus on subfamily Hypostominae." volume="82">Lujan et al. (2015 a</bibRefCitation>
) also generated a multi-locus molecular phylogenetic hypothesis spanning over 24 valid and undescribed species, with
<taxonomicName id="1427027D72B771CEB0CB1CCF4427501D" authorityName="Regan" authorityYear="1904" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="breve">
<emphasis id="C3A2DD10440A2246C1FC94EC32BEFE18" italics="true">Chaetostoma breve</emphasis>
</taxonomicName>
represented by samples from the Napo, Pastaza, Marañon, Santiago and Huallaga River Basins. Bayesian and Maximum Likelihood analyses found uniformly strong support for monophyly of the entire
<taxonomicName id="F6FC9574F223ACC2C4C022BF73B502E7" authorityName="Regan" authorityYear="1904" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="breve">
<emphasis id="5D06A1EA6DDB3D3E155DA2D81B1424C5" italics="true">Chaetostoma breve</emphasis>
</taxonomicName>
clade (Bayesian posterior probability: 1.00, Maximum Likelihood bootstrap: 100).
</paragraph>
<paragraph id="AF021D18F57C2160B2F94E52658B23EA">
<taxonomicName id="E152035D1E7EF034E130C9052D6ADF0D" authorityName="Steindachner" authorityYear="1881" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="9D7E36C727266B7C35294AE4F40203A7" italics="true">Chaetostoma branickii</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="E33C299EB24550E7EB9578C2D36B7AF0" author="Steindachner F" firstAuthor="Steindachner" journalOrPublisher="” Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften in Wien" pagination="103-146" refId="B29" refString="Steindachner F (1881) “ Beiträge zur Kenntniss der Flussfische Südamerikas. II. ” Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften in Wien 43: 103146." title="“ Beiträge zur Kenntniss der Flussfische Südamerikas. II." volume="43" year="1881">Steindachner (1881)</bibRefCitation>
, based on specimens from the Chota River near
<collectingRegion id="041DD8D5F12F844E50990FB687908C35" country="Peru" name="Cajamarca">Cajamarca</collectingRegion>
in northern
<collectingCountry id="98900A4AB0A36CFD68FA378383FD138C" name="Peru">Peru</collectingCountry>
. That portion of the Marañon River Basin was poorly sampled at the time of
<bibRefCitation id="C9EA99FB6C32D49FBF1928A93F2EAB7C" DOI="10.1643/CI-15-246" author="Lujan NK &amp; Meza-Vargas V &amp; Barriga-Salazar R" issue="3" journalOrPublisher="Copeia" pagination="651-663" refId="B13" refString="Lujan NK, Meza-Vargas V, Barriga-Salazar R (2015 c) Two new Chaetostoma group (Loricariidae: Hypostominae) sister genera from opposite sides of the Andes Mountains in Ecuador, with the description of one new species. Copeia 103 (3): 651663. https://doi.org/10.1643/CI-15-246" title="Two new Chaetostoma group (Loricariidae: Hypostominae) sister genera from opposite sides of the Andes Mountains in Ecuador, with the description of one new species." volume="103">Lujan et al. (2015 c</bibRefCitation>
), thus few specimens and no tissues from that specific region were available, making it reasonable to assume that a distinct, endemic species might exist there. Moreover, the
<typeStatus id="8354BA850B67C30C5BF66463BFBE7D83" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="AEA1D92D166057DFE848C753768055AC" authorityName="Steindachner" authorityYear="1881" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="AB4C88453EB7FCC45C30E274E47C28EB" italics="true">Chaetostoma branickii</emphasis>
</taxonomicName>
were in Vienna and not directly accessible to the authors.
<bibRefCitation id="921FA59F1D2F4A5D9FF643A020AA8371" DOI="10.1643/CI-15-246" author="Lujan NK &amp; Meza-Vargas V &amp; Barriga-Salazar R" issue="3" journalOrPublisher="Copeia" pagination="651-663" refId="B13" refString="Lujan NK, Meza-Vargas V, Barriga-Salazar R (2015 c) Two new Chaetostoma group (Loricariidae: Hypostominae) sister genera from opposite sides of the Andes Mountains in Ecuador, with the description of one new species. Copeia 103 (3): 651663. https://doi.org/10.1643/CI-15-246" title="Two new Chaetostoma group (Loricariidae: Hypostominae) sister genera from opposite sides of the Andes Mountains in Ecuador, with the description of one new species." volume="103">Lujan et al. (2015 c</bibRefCitation>
) concluded that
<taxonomicName id="45BACCD48AB153796F124A28E36936E0" authorityName="Steindachner" authorityYear="1881" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="D5486D2DCA8FDA1740388588D9DCC026" italics="true">Chaetostoma branickii</emphasis>
</taxonomicName>
might be more closely allied with
<taxonomicName id="57A4ED2B7DAD905D5ED2CA8B059E23C0" authorityName="Steindachner" authorityYear="1882" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="taczanowskii">
<emphasis id="D5EACF44D794FC566537CC2FC9F73FCF" italics="true">Chaetostoma taczanowskii</emphasis>
</taxonomicName>
, which was described by
<bibRefCitation id="26DF8B313989777B35B1C830CF713D2D" author="Steindachner F" firstAuthor="Steindachner" journalOrPublisher="” Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften in Wien" pagination="103-146" refId="B29" refString="Steindachner F (1881) “ Beiträge zur Kenntniss der Flussfische Südamerikas. II. ” Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften in Wien 43: 103146." title="“ Beiträge zur Kenntniss der Flussfische Südamerikas. II." volume="43" year="1881">Steindachner (1881)</bibRefCitation>
, based on specimens from the Huallaga River drainage to the south. To further investigate the validity of
<taxonomicName id="E1FA25F70ECC7ED02E49239FE0AE90CF" authority=", Lujan" authorityName="Lujan" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="F5E50EE441184ADAA10EFC0133754383" italics="true">Chaetostoma branickii</emphasis>
, Lujan
</taxonomicName>
and colleagues sampled the upper middle Marañon River in 2018, collecting over 540 fresh specimens and 60 tissues of
<taxonomicName id="D282DABBAB38C526AFB24E8B7003CD8A" authorityName="Meza-Vargas &amp; Ramirez &amp; Lujan" authorityYear="2024" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="genus">
<emphasis id="7EBB2AE3995F1AA70FD08F81D5143E84" italics="true">Chaetostoma</emphasis>
</taxonomicName>
. This new material allowed a more detailed comparison of fresh, nearly topotypic specimens with the specimens on which
<bibRefCitation id="65F4DB0DE0529F8ABA4869C4CCA9F37B" DOI="10.1016/j.ympev.2014.08.020" author="Lujan NK &amp; Armbruster JW &amp; Lovejoy NR &amp; López-Fernández H" journalOrPublisher="Molecular phylogenetics and evolution" pagination="269-288" refId="B11" refString="Lujan NK, Armbruster JW, Lovejoy NR, López-Fernández H (2015 a) Multilocus molecular phylogeny of the suckermouth armored catfishes (Siluriformes: Loricariidae) with a focus on subfamily Hypostominae. Molecular phylogenetics and evolution 82: 269288. https://doi.org/10.1016/j.ympev.2014.08.020" title="Multilocus molecular phylogeny of the suckermouth armored catfishes (Siluriformes: Loricariidae) with a focus on subfamily Hypostominae." volume="82">Lujan et al. (2015 a</bibRefCitation>
) based their re-description of
<taxonomicName id="CFCE3494A3FAF7B7B369E345C9D47707" authorityName="Regan" authorityYear="1904" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="breve">
<emphasis id="8195D6BB1D0E476C3135B608A08B1CF2" italics="true">Chaetostoma breve</emphasis>
</taxonomicName>
and type images of
<taxonomicName id="4D41EBF511D330A4E9F48E8C478BA508" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="2837F985588774D3A298A50F6B129A79" italics="true">Chaetostoma branickii</emphasis>
</taxonomicName>
(Figs
<figureCitation id="82F164E9EDC47AC848BAB30D7333BB0F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Original illustrations of Chaetostoma branickii from Steindachner 1881." figureDoi="10.3897/zse.100.118522.figure4" httpUri="https://binary.pensoft.net/fig/1156331">4</figureCitation>
,
<figureCitation id="1CBB72F857F61538B3F503D0DCA04DD5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Photo comparison of a nearly 150 - year-old preserved syntype of Chaetostoma branickii (NMW 47270-71, photos by Mark Sabaj &amp; Kyle Luckenbill, copyright NMW) with a live specimen collected in 2018 from the Marañon River, Cajamarca Province, Peru (ROM 109213, photos by NKL). Scale bars: 1 cm." figureDoi="10.3897/zse.100.118522.figure5" httpUri="https://binary.pensoft.net/fig/1156332">5</figureCitation>
). Based on this comparison, it has become clear that these are the same species and that
<taxonomicName id="AB1FAEA453C322C58ADFF65134F722EA" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="22A24A748FD566F09D5923241F0A375A" italics="true">Chaetostoma branickii</emphasis>
</taxonomicName>
should thus be recognised as the senior synonym of
<taxonomicName id="DD7674E2BB24435A1AB05B1A281F695D" authorityName="Regan" authorityYear="1904" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="breve">
<emphasis id="55F06ABB9120021CA522674A763406CE" italics="true">Chaetostoma breve</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="BBAC97376BCA438D77C7D526509DDC7E" ID-DOI="10.3897/zse.100.118522.figure4" ID-arpha="4993E651-771D-578E-8AED-76F1538F121C" httpUri="https://binary.pensoft.net/fig/1156331" startId="F4">
<paragraph id="E07D998D7002F8611FBFAB670EE4FEA6">
<label id="5E5EB46C3DA28A5DB4F10AAE86DC75B2">Figure 4.</label>
</paragraph>
<paragraph id="25DBF3734BA0541D7607C86ABAD85FED">
Original illustrations of
<taxonomicName id="1EBA3F5B5DDF9B2D2135326ED9BA4170" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="338B42994193E79D1047A95CEA724F9D" italics="true">Chaetostoma branickii</emphasis>
</taxonomicName>
from
<bibRefCitation id="0194158068E15D8E7F82E17B0D719E0F" author="Steindachner" firstAuthor="Steindachner" journalOrPublisher="” Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften in Wien" pagination="103-146" refId="B29" refString="Steindachner F (1881) “ Beiträge zur Kenntniss der Flussfische Südamerikas. II. ” Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften in Wien 43: 103146." title="“ Beiträge zur Kenntniss der Flussfische Südamerikas. II." volume="43" year="1881">Steindachner 1881</bibRefCitation>
.
</paragraph>
</caption>
<caption id="108F35F266853275CCBBA6DB6E14C3CB" ID-DOI="10.3897/zse.100.118522.figure5" ID-arpha="1165D1D9-977D-5FE0-AF58-979A3888CFE9" httpUri="https://binary.pensoft.net/fig/1156332" startId="F5">
<paragraph id="9E66FD02EE93BFFD92712F519FF04185">
<label id="441E8CDE83418FEF68A25E5CFA0ABDDA">Figure 5.</label>
</paragraph>
<paragraph id="4B1910971C6E68D5C13D679F806B9D73">
Photo comparison of a nearly 150 - year-old preserved syntype of
<taxonomicName id="CBCFCD7E0B3CF8F378F8A5C04FD69E74" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="3D27CC16857A16436246383D1FE19629" italics="true">Chaetostoma branickii</emphasis>
</taxonomicName>
(NMW 47270-71, photos by Mark Sabaj &amp; Kyle Luckenbill, copyright NMW) with a live specimen collected in 2018 from the Marañon River, Cajamarca Province, Peru (ROM 109213, photos by NKL). Scale bars: 1 cm.
</paragraph>
</caption>
<paragraph id="D68D7CDD24ADBDC930CA0277F9A6D709">
<taxonomicName id="23304D6AD2D809B39E69977DA24D0976" authority="Steindachner 1882" authorityName="Steindachner" authorityYear="1882" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="taczanowskii">
<emphasis id="C0BA7B571D9D4EE6F5FC4FFDAC6DDDB7" italics="true">Chaetostoma taczanowskii</emphasis>
Steindachner 1882
</taxonomicName>
is another taxonomically ambiguous species from northern
<collectingCountry id="BD48A8B201E6A033F56DD921C7C7B6B1" name="Peru">Peru</collectingCountry>
, described from the Huallaga River Basin, a southern tributary of the Marañon.
<typeStatus id="AF4947734377308AEABF37B088AB0952">Type</typeStatus>
images of this species suggest that it is also closely related to
<taxonomicName id="90985AE0C067157007725E38A54C9307" family="Loricariidae" genus="Chaetostoma" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="branickii">
<emphasis id="7FB4F2E0403946CF8B5C0CDB704A517B" italics="true">Chaetostoma branickii</emphasis>
</taxonomicName>
, if not another junior synonym, but a robust evaluation of the status of this species must await availability of fresh topotypic specimens and tissues.
</paragraph>
</subSubSection>
</treatment>
</document>