From e962b1e2e98b80b461ea2d5fec684e5f3244fb77 Mon Sep 17 00:00:00 2001 From: ggserver <ggserver@srv1.plazi.org> Date: Thu, 3 Oct 2024 23:43:41 +0000 Subject: [PATCH] Add updates up until 2024-10-03 23:36:39 --- .../4C/03964C6FFF88673DFF5A12FA840FFE1D.xml | 107 ++++++------ .../4C/03964C6FFF9F673BFF72149C8526F9CF.xml | 161 +++++++++--------- 2 files changed, 137 insertions(+), 131 deletions(-) diff --git a/data/03/96/4C/03964C6FFF88673DFF5A12FA840FFE1D.xml b/data/03/96/4C/03964C6FFF88673DFF5A12FA840FFE1D.xml index abaa29b7dbe..6bae05626cc 100644 --- a/data/03/96/4C/03964C6FFF88673DFF5A12FA840FFE1D.xml +++ b/data/03/96/4C/03964C6FFF88673DFF5A12FA840FFE1D.xml @@ -1,51 +1,54 @@ -<document id="8B4ABB0AC44A86DF65A5900DFCA6B5DE" ID-DOI="10.5070/P9401362375" ID-ISSN="0031-0298" ID-ZooBank="EE96562A-AEB9-4E4C-AAAF-036F41D8D3EF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1712167387144" checkinUser="felipe" docAuthor="Kiernan, Caitlin R. & Ebersole, Jun A." docDate="2023" docId="03964C6FFF88673DFF5A12FA840FFE1D" docLanguage="en" docName="PaleoBios.40.13.1-28.pdf" docOrigin="PaleoBios 40 (13)" docSource="http://dx.doi.org/10.5070/p9401362375" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.3:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="3" docTitle="Ectenosaurus shannoni Kiernan & Ebersole 2023, SP. 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pageId="17" pageNumber="16"> +<treatment id="03964C6FFF88673DFF5A12FA840FFE1D" ID-DOI="http://doi.org/10.5281/zenodo.13888198" ID-Zenodo-Dep="13888198" LSID="urn:lsid:zoobank.org:act:4D791F56-9245-41CF-9290-2A0AA0F7C6DA" httpUri="http://treatment.plazi.org/id/03964C6FFF88673DFF5A12FA840FFE1D" lastPageId="23" lastPageNumber="21" pageId="17" pageNumber="16"> <subSubSection id="C325AEF2FF88673BFF5A12FA85B4F904" box="[246,667,1691,1719]" pageId="17" pageNumber="16" type="nomenclature"> <paragraph id="8B80FD79FF88673BFF5A12FA85B4F904" blockId="17.[246,667,1691,1753]" box="[246,667,1691,1719]" pageId="17" pageNumber="16"> <heading id="D0C84A15FF88673BFF5A12FA85B4F904" allCaps="true" box="[246,667,1691,1719]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="16" reason="2"> @@ -58,7 +61,7 @@ </subSubSection> <subSubSection id="C325AEF2FF88673BFE2A12A08509F8D2" pageId="17" pageNumber="16" type="description"> <paragraph id="8B80FD79FF88673BFE2A12A08525F96B" blockId="17.[246,667,1691,1753]" box="[390,522,1729,1753]" pageId="17" pageNumber="16"> -<figureCitation id="1304E1FCFF88673BFE2A12A08525F96B" box="[390,522,1729,1753]" captionStart-0="Figure 11" captionStart-1="Figure 12" captionStart-2="Figure 13" captionStartId-0="19.[112,183,1882,1904]" captionStartId-1="21.[112,183,748,770]" captionStartId-2="22.[111,186,1613,1637]" captionTargetBox-0="[170,1477,207,1856]" captionTargetBox-1="[140,1486,216,722]" captionTargetBox-2="[155,1477,208,1581]" captionTargetId-0="figure-71@19.[143,1499,192,1870]" captionTargetId-1="figure-544@21.[117,1515,189,736]" captionTargetId-2="figure-115@22.[128,1499,191,1601]" captionTargetPageId-0="19" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." captionText-1="Figure 12: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–D. Left quadrate fragment with suprastapedial-infrastapedial contact in posterior (A), anterior/interior (B), lateral (C), and medial (D) views. Arrow indicates interior flange of infrastapedial process. Scale bar=2 cm." captionText-2="Figure 13: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–C. Right splenial in lateral (A), medial (B), and posterior (C) views. D, E. Right coronoid in lateral (D) and medial (E) views. F. Laser scan of cast of right coronoid of ALMNH:Paleo:5452 in posteromedial view. G. Cervical vertebra in ventral view. H. Caudal vertebra in ventral view. Scale bars for A–F=5 cm. Scale bar for G=4 cm." pageId="17" pageNumber="16">FIGS. 11–13</figureCitation> +<figureCitation id="1304E1FCFF88673BFE2A12A08525F96B" box="[390,522,1729,1753]" captionStart-0="Figure 11" captionStart-1="Figure 12" captionStart-2="Figure 13" captionStartId-0="19.[112,183,1882,1904]" captionStartId-1="21.[112,183,748,770]" captionStartId-2="22.[111,186,1613,1637]" captionTargetBox-0="[170,1477,207,1856]" captionTargetBox-1="[140,1486,216,722]" captionTargetBox-2="[155,1477,208,1581]" captionTargetId-0="figure-71@19.[143,1499,192,1870]" captionTargetId-1="figure-544@21.[117,1515,189,736]" captionTargetId-2="figure-115@22.[128,1499,191,1601]" captionTargetPageId-0="19" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." captionText-1="Figure 12: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–D. Left quadrate fragment with suprastapedial-infrastapedial contact in posterior (A), anterior/interior (B), lateral (C), and medial (D) views. Arrow indicates interior flange of infrastapedial process. Scale bar=2 cm." captionText-2="Figure 13: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–C. Right splenial in lateral (A), medial (B), and posterior (C) views. D, E. Right coronoid in lateral (D) and medial (E) views. F. Laser scan of cast of right coronoid of ALMNH:Paleo:5452 in posteromedial view. G. Cervical vertebra in ventral view. H. Caudal vertebra in ventral view. Scale bars for A–F=5 cm. Scale bar for G=4 cm." figureDoi-0="http://doi.org/10.5281/zenodo.13888194" figureDoi-1="http://doi.org/10.5281/zenodo.13888196" figureDoi-2="http://doi.org/10.5281/zenodo.13888200" httpUri-0="https://zenodo.org/record/13888194/files/figure.png" httpUri-1="https://zenodo.org/record/13888196/files/figure.png" httpUri-2="https://zenodo.org/record/13888200/files/figure.png" pageId="17" pageNumber="16">FIGS. 11–13</figureCitation> </paragraph> <paragraph id="8B80FD79FF88673BFF3C1293853DF882" blockId="17.[112,799,1778,1840]" pageId="17" pageNumber="16"> <emphasis id="B94B216BFF88673BFF3C1293864CF8BE" bold="true" box="[144,355,1778,1804]" pageId="17" pageNumber="16"> @@ -175,7 +178,7 @@ from erosional gullies within the lower 21 meters of the unnamed member of the , <location id="8EE0ABA2FF8B6738FFDC16268661FDD2" LSID="urn:lsid:plazi:treatment:03964C6FFF88673DFF5A12FA840FFE1D:8EE0ABA2FF8B6738FFDC16268661FDD2" box="[112,334,582,609]" country="United States of America" county="Greene County" municipality="Arcola Limestone Member" name="Upper Cretaceous" pageId="18" pageNumber="17" stateProvince="Alabama">Upper Cretaceous</location> , -<figureCitation id="1304E1FCFF8B6738FEFB16278682FDD2" box="[343,429,582,608]" captionStart="Figure 2" captionStartId="7.[110,181,1899,1921]" captionTargetBox="[202,1418,204,1874]" captionTargetId="figure-64@7.[155,1457,192,1894]" captionTargetPageId="7" captionText="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="18" pageNumber="17">Fig. 2B</figureCitation> +<figureCitation id="1304E1FCFF8B6738FEFB16278682FDD2" box="[343,429,582,608]" captionStart="Figure 2" captionStartId="7.[110,181,1899,1921]" captionTargetBox="[202,1418,204,1874]" captionTargetId="figure-64@7.[155,1457,192,1894]" captionTargetPageId="7" captionText="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888170" httpUri="https://zenodo.org/record/13888170/files/figure.png" pageId="18" pageNumber="17">Fig. 2B</figureCitation> ), located in the NE 1/4, SE 1/4 of <collectorName id="26CA98AFFF8B6738FF2116088793FD31" box="[141,188,617,643]" pageId="18" pageNumber="17">Sec.</collectorName> 30, @@ -189,7 +192,7 @@ E., , <collectingCountry id="F328BDE9FF8B6738FD1D160885CBFD31" box="[689,740,617,643]" name="United States of America" pageId="18" pageNumber="17">USA</collectingCountry> ( -<figureCitation id="1304E1FCFF8B6738FD5E160887B9FD15" captionStart="Figure 1" captionStartId="4.[112,183,1509,1531]" captionTargetBox="[244,1389,216,1482]" captionTargetId="figure-192@4.[225,1403,191,1493]" captionTargetPageId="4" captionText="Figure 1. Geography and Stratigraphy. A. Generalized map of the United States during the Santonian Stage of the Cretaceous (~85 mya) with localities for YPM VP4673 (1) and ALMNH:Paleo:4673 (2). B. Generalized correlation chart of Coniacian to Campanian surface stratigraphy for Kansas and Alabama, USA. Base map data modified from Blakey (2014). Stratigraphic chart modified from Szabo et al. (1988) and Everhart (2014)." pageId="18" pageNumber="17">Fig. 1A</figureCitation> +<figureCitation id="1304E1FCFF8B6738FD5E160887B9FD15" captionStart="Figure 1" captionStartId="4.[112,183,1509,1531]" captionTargetBox="[244,1389,216,1482]" captionTargetId="figure-192@4.[225,1403,191,1493]" captionTargetPageId="4" captionText="Figure 1. Geography and Stratigraphy. A. Generalized map of the United States during the Santonian Stage of the Cretaceous (~85 mya) with localities for YPM VP4673 (1) and ALMNH:Paleo:4673 (2). B. Generalized correlation chart of Coniacian to Campanian surface stratigraphy for Kansas and Alabama, USA. Base map data modified from Blakey (2014). Stratigraphic chart modified from Szabo et al. (1988) and Everhart (2014)." figureDoi="http://doi.org/10.5281/zenodo.13888168" httpUri="https://zenodo.org/record/13888168/files/figure.png" pageId="18" pageNumber="17">Fig. 1A</figureCitation> ). <location id="8EE0ABA2FF8B6738FF0616ED87F5FD14" LSID="urn:lsid:plazi:treatment:03964C6FFF88673DFF5A12FA840FFE1D:8EE0ABA2FF8B6738FF0616ED87F5FD14" box="[170,218,652,678]" country="United States of America" county="Greene County" municipality="Arcola Limestone Member" name="The" pageId="18" pageNumber="17" stateProvince="Alabama">The</location> locality has since been reclaimed for farmland and is no longer available for study. @@ -250,7 +253,7 @@ specimen during graduate work at the University of The frontal of <collectionCode id="ED2E65BCFF8B6738FE7511288561FAD6" box="[473,590,1353,1380]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="18" pageNumber="17" type="Museum">ALMNH</collectionCode> :Paleo:5452 ( -<figureCitation id="1304E1FCFF8B6738FD42112887EDFA35" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="18" pageNumber="17">Fig. 11A, B</figureCitation> +<figureCitation id="1304E1FCFF8B6738FD42112887EDFA35" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888194" httpUri="https://zenodo.org/record/13888194/files/figure.png" pageId="18" pageNumber="17">Fig. 11A, B</figureCitation> ) is essentially complete, but suffered some minor subaerial weathering prior to collection, particularly on its dorsal side. In overall appearance, the frontal is strikingly similar to that of <emphasis id="B94B216BFF8B6738FE2811B485EDFA43" box="[388,706,1493,1521]" italics="true" pageId="18" pageNumber="17"> <taxonomicName id="4C3F86FAFF8B6738FE2811B48592FA43" baseAuthorityName="Merriam" baseAuthorityYear="1894" box="[388,701,1493,1521]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="17" phylum="Chordata" rank="species" species="clidastoides">Ectenosaurus clidastoides</taxonomicName> @@ -280,7 +283,7 @@ and and <collectionCode id="ED2E65BCFF8B6738FD27129185EBF8B8" box="[651,708,1776,1802]" country="South Korea" httpUri="http://grbio.org/cool/sv00-20xv" name="Sangmiung University" pageId="18" pageNumber="17">SMU</collectionCode> 76350. Instead, the anteriormost portion of the dorsal surface ( -<figureCitation id="1304E1FCFF8B6738FFD5135787CDF8E2" box="[121,226,1846,1872]" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="18" pageNumber="17">Fig. 11A</figureCitation> +<figureCitation id="1304E1FCFF8B6738FFD5135787CDF8E2" box="[121,226,1846,1872]" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888194" httpUri="https://zenodo.org/record/13888194/files/figure.png" pageId="18" pageNumber="17">Fig. 11A</figureCitation> ) of the frontal is transversely arched or domed anteriorly, a trait absent in other specimens referred to the genus. This arching gives way to a generally planar dorsal surface, which thickens parasagittaly, then thins abruptly at the antorbital margins, and forms a broad, shallow sulcus posteriorly. Tiny foramina are present across the entire dorsal surface. The posterior sulcus deepens and expands to form a shallow depression that would have met the dorsal parietal table, forming the mesokinetic axis of the skull, movement along which may have been inhibited by a series of short vertical keels and overlapping flanges arising from the posterior margin of the frontal and the anterior margin of the parietal. This mode of contact appears to have been similar to, but simpler and less restrictive than, that seen in <emphasis id="B94B216BFF8B6738FA2E162484A0FD36" italics="true" pageId="18" pageNumber="17"> <taxonomicName id="4C3F86FAFF8B6738FA2E162484A5FD36" baseAuthorityName="Merriam" baseAuthorityYear="1894" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="17" phylum="Chordata" rank="species" species="clidastoides">E. clidastoides</taxonomicName> @@ -338,7 +341,7 @@ frontal being almost completely obscured beneath the left mandible. In : <collectorName id="26CA98AFFF8B6738FAC2119D829EF9A5" box="[1390,1457,1532,1559]" pageId="18" pageNumber="17">Paleo</collectorName> :5452 ( -<figureCitation id="1304E1FCFF8B6738FCE51241849CF988" box="[841,947,1568,1594]" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="18" pageNumber="17">Fig. 11B</figureCitation> +<figureCitation id="1304E1FCFF8B6738FCE51241849CF988" box="[841,947,1568,1594]" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888194" httpUri="https://zenodo.org/record/13888194/files/figure.png" pageId="18" pageNumber="17">Fig. 11B</figureCitation> ), the frontal is bisected ventrally by the olfactory canal, which is bordered on either side by prominent descending processes that run roughly parallel before converging abruptly, then immediately diverging where the canal opens to accommodate the olfactory bulb. <collectorName id="26CA98AFFF8B6738FCEC12AE8459F95B" box="[832,886,1743,1769]" pageId="18" pageNumber="17">This</collectorName> forms a roughly X-shaped configuration, with the descending processes thickest where they almost meet at the crux of the @@ -384,7 +387,7 @@ or </taxonomicName> and the ventral separating ridge appears to be absent. </paragraph> -<caption id="DF40ADF1FF8A6739FFDC133B8674F810" pageId="19" pageNumber="18" startId="19.[112,183,1882,1904]" targetBox="[170,1477,207,1856]" targetPageId="19" targetType="figure"> +<caption id="DF40ADF1FF8A6739FFDC133B8674F810" ID-DOI="http://doi.org/10.5281/zenodo.13888194" ID-Zenodo-Dep="13888194" httpUri="https://zenodo.org/record/13888194/files/figure.png" pageId="19" pageNumber="18" startId="19.[112,183,1882,1904]" targetBox="[170,1477,207,1856]" targetPageId="19" targetType="figure"> <paragraph id="8B80FD79FF8A6739FFDC133B8674F810" blockId="19.[112,1498,1881,1955]" pageId="19" pageNumber="18"> <emphasis id="B94B216BFF8A6739FFDC133B87F7F8DD" bold="true" box="[112,216,1882,1904]" pageId="19" pageNumber="18">Figure 11</emphasis> . @@ -418,9 +421,9 @@ Frontal in dorsal ( Only the incomplete anterior half of the parietal is preserved on <collectionCode id="ED2E65BCFF8D673EFE05178D8536FBB5" box="[425,537,1004,1031]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="20" pageNumber="19" type="Museum">ALMNH</collectionCode> :Paleo:5452 ( -<figureCitation id="1304E1FCFF8D673EFD1D178D87A7FB98" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="20" pageNumber="19">Fig. 11C, D</figureCitation> +<figureCitation id="1304E1FCFF8D673EFD1D178D87A7FB98" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888194" httpUri="https://zenodo.org/record/13888194/files/figure.png" pageId="20" pageNumber="19">Fig. 11C, D</figureCitation> ). Prior to collection, the parietal split horizontally into dorsal and ventral halves that were subsequently glued together during preparation. The dorsal parietal table ( -<figureCitation id="1304E1FCFF8D673EFF111018860BFB21" box="[189,292,1145,1171]" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="20" pageNumber="19">Fig. 11C</figureCitation> +<figureCitation id="1304E1FCFF8D673EFF111018860BFB21" box="[189,292,1145,1171]" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888194" httpUri="https://zenodo.org/record/13888194/files/figure.png" pageId="20" pageNumber="19">Fig. 11C</figureCitation> ) can be divided into two regions: a broad anterior subrhomboidal shelf that merges smoothly into an elongate posterior shaft with rounded, parallel sides, which would have branched into the missing suspensorial rami. There is no evidence of a parasagittal ridge. A large subcircular parietal foramen, very slightly longer than wide, is centered on the dorsal table, and set back from the frontoparietal suture by a distance roughly equal to twice its diameter. Dorsally, the position and shape of the foramen compares well with that of both <collectionCode id="ED2E65BCFF8D673EFFDC11B88795FA41" box="[112,186,1497,1523]" country="0" httpUri="http://grbio.org/cool/5bc7-5n0c" name="Fort Hays Sternberg Museum" pageId="20" pageNumber="19">FHSM</collectionCode> <collectionCode id="ED2E65BCFF8D673EFF6D11B887DDFA41" box="[193,242,1497,1523]" pageId="20" pageNumber="19">VP-</collectionCode> @@ -439,7 +442,7 @@ Most of the lateral descending process (descensus processus parietalis of </paragraph> <paragraph id="8B80FD79FF8D673EFCCC154D8271FCC8" blockId="20.[832,1521,195,1944]" pageId="20" pageNumber="19"> The ventral surface of the parietal ( -<figureCitation id="1304E1FCFF8D673EFABE154D8253FEF5" box="[1298,1404,300,327]" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="20" pageNumber="19">Fig. 11D</figureCitation> +<figureCitation id="1304E1FCFF8D673EFABE154D8253FEF5" box="[1298,1404,300,327]" captionStart="Figure 11" captionStartId="19.[112,183,1882,1904]" captionTargetBox="[170,1477,207,1856]" captionTargetId="figure-71@19.[143,1499,192,1870]" captionTargetPageId="19" captionText="Figure 11. Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A, B. Frontal in dorsal (A) and ventral (B) views. C, D. Parietal in dorsal (C) and ventral (D) views. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888194" httpUri="https://zenodo.org/record/13888194/files/figure.png" pageId="20" pageNumber="19">Fig. 11D</figureCitation> ) is dominated by a large elliptical foramen, at least twice as long as wide, in contrast with its circular dorsal opening. It is unclear whether this dorsoventral variance is an artifact of plastic deformation or accurately reflects the anatomy of <emphasis id="B94B216BFF8D673EFCCF15BA84D2FE45" box="[867,1021,475,503]" italics="true" pageId="20" pageNumber="19"> <taxonomicName id="4C3F86FAFF8D673EFCCF15BA84D6FE45" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[867,1017,475,503]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="19" phylum="Chordata" rank="species" species="shannoni">E. shannoni</taxonomicName> @@ -454,7 +457,7 @@ as the ventral triangular eminence, present in halisaurines and many russellosau <paragraph id="8B80FD79FF8D673FFCCC17E0860AFBFB" blockId="20.[832,1521,195,1944]" lastBlockId="21.[112,801,859,1941]" lastPageId="21" lastPageNumber="20" pageId="20" pageNumber="19"> <emphasis id="B94B216BFF8D673EFCCC17E084D0FC2F" bold="true" box="[864,1023,897,925]" italics="true" pageId="20" pageNumber="19">Quadrate—</emphasis> Only a small portion of the left quadrate was collected ( -<figureCitation id="1304E1FCFF8D673EFB9317C783F9FC72" box="[1087,1238,934,960]" captionStart="Figure 12" captionStartId="21.[112,183,748,770]" captionTargetBox="[140,1486,216,722]" captionTargetId="figure-544@21.[117,1515,189,736]" captionTargetPageId="21" captionText="Figure 12: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–D. Left quadrate fragment with suprastapedial-infrastapedial contact in posterior (A), anterior/interior (B), lateral (C), and medial (D) views. Arrow indicates interior flange of infrastapedial process. Scale bar=2 cm." pageId="20" pageNumber="19">Fig. 12A‒D</figureCitation> +<figureCitation id="1304E1FCFF8D673EFB9317C783F9FC72" box="[1087,1238,934,960]" captionStart="Figure 12" captionStartId="21.[112,183,748,770]" captionTargetBox="[140,1486,216,722]" captionTargetId="figure-544@21.[117,1515,189,736]" captionTargetPageId="21" captionText="Figure 12: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–D. Left quadrate fragment with suprastapedial-infrastapedial contact in posterior (A), anterior/interior (B), lateral (C), and medial (D) views. Arrow indicates interior flange of infrastapedial process. Scale bar=2 cm." figureDoi="http://doi.org/10.5281/zenodo.13888196" httpUri="https://zenodo.org/record/13888196/files/figure.png" pageId="20" pageNumber="19">Fig. 12A‒D</figureCitation> ), but it preserves the contact and fusion of the suprastapedial process with the infrastapedial and the presence of a dorsomedially directed flange from the infrastapedial that overlaps the expanded dorsodistal end of the suprastapedial, a trait already mentioned in the description of <collectionCode id="ED2E65BCFF8D673EFAE5103682ADFBC3" box="[1353,1410,1111,1137]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="20" pageNumber="19" type="Museum">YPM</collectionCode> VP4673 as unknown in any mosasaur other than @@ -480,7 +483,7 @@ or In <collectionCode id="ED2E65BCFF8D673EFB5F110E824BFA38" box="[1267,1380,1391,1418]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="20" pageNumber="19" type="Museum">ALMNH</collectionCode> :Paleo:5452, the infrastapedial process forms a delicate cheliform structure that not only overlaps the suprastapedial posterodorsally, but ventrally as well, embracing it in a sort of pincer grip while also creating a cradle that abuts the suprastapedial terminus. Complete fusion between the two processes can only be observed ventromedially ( -<figureCitation id="1304E1FCFF8D673EFCE512078495F932" box="[841,954,1638,1664]" captionStart="Figure 12" captionStartId="21.[112,183,748,770]" captionTargetBox="[140,1486,216,722]" captionTargetId="figure-544@21.[117,1515,189,736]" captionTargetPageId="21" captionText="Figure 12: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–D. Left quadrate fragment with suprastapedial-infrastapedial contact in posterior (A), anterior/interior (B), lateral (C), and medial (D) views. Arrow indicates interior flange of infrastapedial process. Scale bar=2 cm." pageId="20" pageNumber="19">Fig. 12B</figureCitation> +<figureCitation id="1304E1FCFF8D673EFCE512078495F932" box="[841,954,1638,1664]" captionStart="Figure 12" captionStartId="21.[112,183,748,770]" captionTargetBox="[140,1486,216,722]" captionTargetId="figure-544@21.[117,1515,189,736]" captionTargetPageId="21" captionText="Figure 12: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–D. Left quadrate fragment with suprastapedial-infrastapedial contact in posterior (A), anterior/interior (B), lateral (C), and medial (D) views. Arrow indicates interior flange of infrastapedial process. Scale bar=2 cm." figureDoi="http://doi.org/10.5281/zenodo.13888196" httpUri="https://zenodo.org/record/13888196/files/figure.png" pageId="20" pageNumber="19">Fig. 12B</figureCitation> ). While the full extent of fusion is unclear,, the processes are less extensively fused than in <emphasis id="B94B216BFF8D673EFA0612E98494F975" italics="true" pageId="20" pageNumber="19"> <taxonomicName id="4C3F86FAFF8D673EFA0612E98498F975" baseAuthorityName="Merriam" baseAuthorityYear="1894" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="20" pageNumber="19" phylum="Chordata" rank="species" species="clidastoides">E. clidastoides</taxonomicName> @@ -489,7 +492,7 @@ In The infrastapedial process of <collectionCode id="ED2E65BCFF8D673EFAE412CC82ADF975" box="[1352,1410,1709,1735]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="20" pageNumber="19" type="Museum">YPM</collectionCode> VP4673 ( -<figureCitation id="1304E1FCFF8D673EFCE512B1848EF958" box="[841,929,1744,1770]" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." pageId="20" pageNumber="19">Fig. 3B</figureCitation> +<figureCitation id="1304E1FCFF8D673EFCE512B1848EF958" box="[841,929,1744,1770]" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888172" httpUri="https://zenodo.org/record/13888172/files/figure.png" pageId="20" pageNumber="19">Fig. 3B</figureCitation> ) may have enfolded the suprastapedial in much the same way, but preservation makes this difficult to determine with any certainty; fusion appears to have been more extensive in the Niobrara Chalk specimen. In neither <collectionCode id="ED2E65BCFF8D673EFC08133B833BF8C7" box="[932,1044,1882,1909]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="20" pageNumber="19" type="Museum">ALMNH</collectionCode> :Paleo:5452 nor @@ -506,7 +509,7 @@ or . </emphasis> </paragraph> -<caption id="DF40ADF1FF8C673FFFDC168D87F8FC86" pageId="21" pageNumber="20" startId="21.[112,183,748,770]" targetBox="[140,1486,216,722]" targetPageId="21" targetType="figure"> +<caption id="DF40ADF1FF8C673FFFDC168D87F8FC86" ID-DOI="http://doi.org/10.5281/zenodo.13888196" ID-Zenodo-Dep="13888196" httpUri="https://zenodo.org/record/13888196/files/figure.png" pageId="21" pageNumber="20" startId="21.[112,183,748,770]" targetBox="[140,1486,216,722]" targetPageId="21" targetType="figure"> <paragraph id="8B80FD79FF8C673FFFDC168D87F8FC86" blockId="21.[112,1492,747,821]" pageId="21" pageNumber="20"> <emphasis id="B94B216BFF8C673FFFDC168D87CFFCB3" bold="true" box="[112,224,748,770]" pageId="21" pageNumber="20">Figure 12:</emphasis> <taxonomicName id="4C3F86FAFF8C673FFF49168A86E3FCB0" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[229,460,747,770]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="21" pageNumber="20" phylum="Chordata" rank="species" species="shannoni" status="sp. nov."> @@ -555,7 +558,7 @@ VP4673, it would still have been more complex than the ball-and-socket condyle/c The majority of the right coronoid (Fig. 13D‒F) is preserved, though it should be noted that some portions have been heavily restored with plaster by G.L. Bell, Jr. (M.J. Polcyn <emphasis id="B94B216BFF8C673FFB1E173882C0FCC7" box="[1202,1519,857,885]" italics="true" pageId="21" pageNumber="20">personal communication</emphasis> 2022). Fortunately, the restored portions are unpainted and easily discernible from the fossil itself. As in most russellosaurines, the coronoid is a short, saddle-shaped wedge of bone straddling the anterior rim of the surangular and forming a posteromedial sutural contact with a buttress on the surangular ( -<figureCitation id="1304E1FCFF8C673FFB14104F826CFBFB" box="[1208,1347,1070,1097]" captionStart="Figure 13" captionStartId="22.[111,186,1613,1637]" captionTargetBox="[155,1477,208,1581]" captionTargetId="figure-115@22.[128,1499,191,1601]" captionTargetPageId="22" captionText="Figure 13: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–C. Right splenial in lateral (A), medial (B), and posterior (C) views. D, E. Right coronoid in lateral (D) and medial (E) views. F. Laser scan of cast of right coronoid of ALMNH:Paleo:5452 in posteromedial view. G. Cervical vertebra in ventral view. H. Caudal vertebra in ventral view. Scale bars for A–F=5 cm. Scale bar for G=4 cm." pageId="21" pageNumber="20">Fig. 13E, F</figureCitation> +<figureCitation id="1304E1FCFF8C673FFB14104F826CFBFB" box="[1208,1347,1070,1097]" captionStart="Figure 13" captionStartId="22.[111,186,1613,1637]" captionTargetBox="[155,1477,208,1581]" captionTargetId="figure-115@22.[128,1499,191,1601]" captionTargetPageId="22" captionText="Figure 13: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–C. Right splenial in lateral (A), medial (B), and posterior (C) views. D, E. Right coronoid in lateral (D) and medial (E) views. F. Laser scan of cast of right coronoid of ALMNH:Paleo:5452 in posteromedial view. G. Cervical vertebra in ventral view. H. Caudal vertebra in ventral view. Scale bars for A–F=5 cm. Scale bar for G=4 cm." figureDoi="http://doi.org/10.5281/zenodo.13888200" httpUri="https://zenodo.org/record/13888200/files/figure.png" pageId="21" pageNumber="20">Fig. 13E, F</figureCitation> ). This sutural contact is more complex in <collectionCode id="ED2E65BCFF8C673FFB3510308226FBDE" box="[1177,1289,1105,1132]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="21" pageNumber="20" type="Museum">ALMNH</collectionCode> :Paleo:5452 than in @@ -605,7 +608,7 @@ and is more dorsally located. In <collectionCode id="ED2E65BCFF8F673CFEB0129C867AF8A5" box="[284,341,1789,1815]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="22" pageNumber="21" type="Museum">YPM</collectionCode> VP4673, are especially prominent on the interior surface of the medial descending wing. </paragraph> -<caption id="DF40ADF1FF8F673CFFC3122C87B9F903" pageId="22" pageNumber="21" startId="22.[111,186,1613,1637]" targetBox="[155,1477,208,1581]" targetPageId="22" targetType="figure"> +<caption id="DF40ADF1FF8F673CFFC3122C87B9F903" ID-DOI="http://doi.org/10.5281/zenodo.13888200" ID-Zenodo-Dep="13888200" httpUri="https://zenodo.org/record/13888200/files/figure.png" pageId="22" pageNumber="21" startId="22.[111,186,1613,1637]" targetBox="[155,1477,208,1581]" targetPageId="22" targetType="figure"> <paragraph id="8B80FD79FF8F673CFFC3122C87B9F903" blockId="22.[111,1513,1613,1713]" pageId="22" pageNumber="21"> <emphasis id="B94B216BFF8F673CFFC3122C87CBF9D7" bold="true" box="[111,228,1613,1637]" pageId="22" pageNumber="21">Figure 13:</emphasis> <taxonomicName id="4C3F86FAFF8F673CFF46122E86F1F9D5" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[234,478,1615,1639]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="22" pageNumber="21" phylum="Chordata" rank="species" species="shannoni" status="sp. nov."> @@ -640,7 +643,7 @@ VP4673, are especially prominent on the interior surface of the medial descendin A total of twenty-seven vertebrae are preserved with <collectionCode id="ED2E65BCFF8F673CFEA91307865AF833" box="[261,373,1894,1921]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="22" pageNumber="21" type="Museum">ALMNH</collectionCode> :Paleo:5452, including five cervicals, thirteen dorsals, five pygals, and four caudals. Most of the vertebrae consist of badly weathered centra lacking most of the cortical surface, neural arches, synapophyses. zygopophyses, and transverse processes. The best cervical vertebra ( -<figureCitation id="1304E1FCFF8F673CFC711323836CF8EE" box="[989,1091,1858,1884]" captionStart="Figure 13" captionStartId="22.[111,186,1613,1637]" captionTargetBox="[155,1477,208,1581]" captionTargetId="figure-115@22.[128,1499,191,1601]" captionTargetPageId="22" captionText="Figure 13: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–C. Right splenial in lateral (A), medial (B), and posterior (C) views. D, E. Right coronoid in lateral (D) and medial (E) views. F. Laser scan of cast of right coronoid of ALMNH:Paleo:5452 in posteromedial view. G. Cervical vertebra in ventral view. H. Caudal vertebra in ventral view. Scale bars for A–F=5 cm. Scale bar for G=4 cm." pageId="22" pageNumber="21">Fig. 13F</figureCitation> +<figureCitation id="1304E1FCFF8F673CFC711323836CF8EE" box="[989,1091,1858,1884]" captionStart="Figure 13" captionStartId="22.[111,186,1613,1637]" captionTargetBox="[155,1477,208,1581]" captionTargetId="figure-115@22.[128,1499,191,1601]" captionTargetPageId="22" captionText="Figure 13: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–C. Right splenial in lateral (A), medial (B), and posterior (C) views. D, E. Right coronoid in lateral (D) and medial (E) views. F. Laser scan of cast of right coronoid of ALMNH:Paleo:5452 in posteromedial view. G. Cervical vertebra in ventral view. H. Caudal vertebra in ventral view. Scale bars for A–F=5 cm. Scale bar for G=4 cm." figureDoi="http://doi.org/10.5281/zenodo.13888200" httpUri="https://zenodo.org/record/13888200/files/figure.png" pageId="22" pageNumber="21">Fig. 13F</figureCitation> ) preserves a distinct anteroventral ridge extending from the anterior rim of an elliptical hypapophyseal peduncle to the ventral rim of the cotyle. In lateral profile, this ridge forms a shallowly concave arc, as seen in <collectionCode id="ED2E65BCFF8E673DFF5C14878606FEB2" box="[240,297,230,256]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="23" pageNumber="22" type="Museum">YPM</collectionCode> VP4673 and other species of @@ -649,7 +652,7 @@ VP4673 and other species of . </emphasis> The articular surfaces of the pygals are sub-square in anteroposterior view. Only one of the caudles is well preserved ( -<figureCitation id="1304E1FCFF8E673DFF50152E8648FEDB" box="[252,359,335,361]" captionStart="Figure 13" captionStartId="22.[111,186,1613,1637]" captionTargetBox="[155,1477,208,1581]" captionTargetId="figure-115@22.[128,1499,191,1601]" captionTargetPageId="22" captionText="Figure 13: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–C. Right splenial in lateral (A), medial (B), and posterior (C) views. D, E. Right coronoid in lateral (D) and medial (E) views. F. Laser scan of cast of right coronoid of ALMNH:Paleo:5452 in posteromedial view. G. Cervical vertebra in ventral view. H. Caudal vertebra in ventral view. Scale bars for A–F=5 cm. Scale bar for G=4 cm." pageId="23" pageNumber="22">Fig. 13G</figureCitation> +<figureCitation id="1304E1FCFF8E673DFF50152E8648FEDB" box="[252,359,335,361]" captionStart="Figure 13" captionStartId="22.[111,186,1613,1637]" captionTargetBox="[155,1477,208,1581]" captionTargetId="figure-115@22.[128,1499,191,1601]" captionTargetPageId="22" captionText="Figure 13: Ectenosaurus shannoni sp. nov., ALMNH:Paleo:5452. A–C. Right splenial in lateral (A), medial (B), and posterior (C) views. D, E. Right coronoid in lateral (D) and medial (E) views. F. Laser scan of cast of right coronoid of ALMNH:Paleo:5452 in posteromedial view. G. Cervical vertebra in ventral view. H. Caudal vertebra in ventral view. Scale bars for A–F=5 cm. Scale bar for G=4 cm." figureDoi="http://doi.org/10.5281/zenodo.13888200" httpUri="https://zenodo.org/record/13888200/files/figure.png" pageId="23" pageNumber="22">Fig. 13G</figureCitation> ); it is subcircular in anteroposterior profile and preserves two rounded, posteroventrally oriented peduncles for articulation with the haemal arch. </paragraph> </subSubSection> diff --git a/data/03/96/4C/03964C6FFF9F673BFF72149C8526F9CF.xml b/data/03/96/4C/03964C6FFF9F673BFF72149C8526F9CF.xml index 08f1268fa00..458f1891bad 100644 --- a/data/03/96/4C/03964C6FFF9F673BFF72149C8526F9CF.xml +++ b/data/03/96/4C/03964C6FFF9F673BFF72149C8526F9CF.xml @@ -1,51 +1,54 @@ -<document id="8C0A6CAB8C2B38655DC5DEC5902B635D" ID-DOI="10.5070/P9401362375" ID-ISSN="0031-0298" ID-ZooBank="EE96562A-AEB9-4E4C-AAAF-036F41D8D3EF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1712167387144" checkinUser="felipe" docAuthor="Kiernan, Caitlin R. & Ebersole, Jun A." docDate="2023" docId="03964C6FFF9F673BFF72149C8526F9CF" docLanguage="en" docName="PaleoBios.40.13.1-28.pdf" docOrigin="PaleoBios 40 (13)" docSource="http://dx.doi.org/10.5070/p9401362375" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.3:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="3" docTitle="Ectenosaurus tlemonectes Kiernan & Ebersole 2023, SP. 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pageId="6" pageNumber="5"> +<treatment id="03964C6FFF9F673BFF72149C8526F9CF" ID-DOI="http://doi.org/10.5281/zenodo.13888192" ID-Zenodo-Dep="13888192" LSID="urn:lsid:zoobank.org:act:E2E61DA4-E21D-4643-A0B6-A8C04418A062" httpUri="http://treatment.plazi.org/id/03964C6FFF9F673BFF72149C8526F9CF" lastPageId="17" lastPageNumber="16" pageId="6" pageNumber="5"> <subSubSection id="C325AEF2FF9F672CFF72149C859BFEAB" box="[222,692,253,281]" pageId="6" pageNumber="5" type="nomenclature"> <paragraph id="8B80FD79FF9F672CFF72149C859BFEAB" blockId="6.[222,692,253,316]" box="[222,692,253,281]" pageId="6" pageNumber="5"> <heading id="D0C84A15FF9F672CFF72149C859BFEAB" allCaps="true" box="[222,692,253,281]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="5" reason="2"> @@ -58,7 +61,7 @@ </subSubSection> <subSubSection id="C325AEF2FF9F672CFE211545852DFE20" pageId="6" pageNumber="5" type="description"> <paragraph id="8B80FD79FF9F672CFE211545852CFE8E" blockId="6.[222,692,253,316]" box="[397,515,292,316]" pageId="6" pageNumber="5"> -<figureCitation id="1304E1FCFF9F672CFE211545852CFE8E" box="[397,515,292,316]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStart-6="Figure 8" captionStart-7="Figure 9" captionStart-8="Figure 10" captionStartId-0="7.[110,181,1899,1921]" captionStartId-1="9.[112,183,1614,1636]" captionStartId-2="10.[112,183,1615,1637]" captionStartId-3="11.[112,183,1463,1485]" captionStartId-4="12.[113,184,1350,1372]" captionStartId-5="13.[112,183,1899,1921]" captionStartId-6="14.[112,183,1666,1688]" captionStartId-7="16.[112,183,1546,1568]" captionStartId-8="17.[828,899,695,717]" captionTargetBox-0="[202,1418,204,1874]" captionTargetBox-1="[172,1437,217,1589]" captionTargetBox-2="[149,1487,223,1589]" captionTargetBox-3="[178,1453,216,1431]" captionTargetBox-4="[142,1483,218,1315]" captionTargetBox-5="[245,1374,215,1847]" captionTargetBox-6="[177,1450,211,1623]" captionTargetBox-7="[161,1466,208,1518]" captionTargetBox-8="[848,1509,219,666]" captionTargetId-0="figure-64@7.[155,1457,192,1894]" captionTargetId-1="figure-185@9.[125,1482,192,1603]" captionTargetId-2="figure-185@10.[117,1510,190,1605]" captionTargetId-3="figure-261@11.[128,1499,191,1445]" captionTargetId-4="figure-254@12.[112,1519,191,1332]" captionTargetId-5="figure-15@13.[171,1448,192,1872]" captionTargetId-6="figure-106@14.[117,1513,192,1649]" captionTargetId-7="figure-159@16.[141,1485,191,1531]" captionTargetId-8="figure-677@17.[831,1520,192,685]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionTargetPageId-6="14" captionTargetPageId-7="16" captionTargetPageId-8="17" captionText-0="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." captionText-1="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." captionText-2="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." captionText-3="Figure 5. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Dentaries in left lateral (A), left medial (B), left dorsal (C), right lateral (D), and right medial (E) views. Scale bars=10 cm." captionText-4="Figure 6. Ectenosaurus tlemonectes sp. nov., YPM VP 4673, A–D. Detail of pathologic and non-pathologic anterior terminus of left dentary in left lateral (nonpathologic) (A), right lateral (pathologic) (B), right medial (pathologic) (C), and right dorsal (pathologic) (D) views. Scale bars for A–C=5 cm. Scale bar for D=3 cm." captionText-5="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." captionText-6="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." captionText-7="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." captionText-8="Figure 10. Ectenosaurus tlemonectes sp. nov., YPM VP4673. Cervical vertebra in left lateral view. Arrow indicates ventral ridge connecting the hypapophyseal peduncle and cotyle. Scale bar=5 cm." pageId="6" pageNumber="5">FIGS. 2–10</figureCitation> +<figureCitation id="1304E1FCFF9F672CFE211545852CFE8E" box="[397,515,292,316]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStart-6="Figure 8" captionStart-7="Figure 9" captionStart-8="Figure 10" captionStartId-0="7.[110,181,1899,1921]" captionStartId-1="9.[112,183,1614,1636]" captionStartId-2="10.[112,183,1615,1637]" captionStartId-3="11.[112,183,1463,1485]" captionStartId-4="12.[113,184,1350,1372]" captionStartId-5="13.[112,183,1899,1921]" captionStartId-6="14.[112,183,1666,1688]" captionStartId-7="16.[112,183,1546,1568]" captionStartId-8="17.[828,899,695,717]" captionTargetBox-0="[202,1418,204,1874]" captionTargetBox-1="[172,1437,217,1589]" captionTargetBox-2="[149,1487,223,1589]" captionTargetBox-3="[178,1453,216,1431]" captionTargetBox-4="[142,1483,218,1315]" captionTargetBox-5="[245,1374,215,1847]" captionTargetBox-6="[177,1450,211,1623]" captionTargetBox-7="[161,1466,208,1518]" captionTargetBox-8="[848,1509,219,666]" captionTargetId-0="figure-64@7.[155,1457,192,1894]" captionTargetId-1="figure-185@9.[125,1482,192,1603]" captionTargetId-2="figure-185@10.[117,1510,190,1605]" captionTargetId-3="figure-261@11.[128,1499,191,1445]" captionTargetId-4="figure-254@12.[112,1519,191,1332]" captionTargetId-5="figure-15@13.[171,1448,192,1872]" captionTargetId-6="figure-106@14.[117,1513,192,1649]" captionTargetId-7="figure-159@16.[141,1485,191,1531]" captionTargetId-8="figure-677@17.[831,1520,192,685]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionTargetPageId-6="14" captionTargetPageId-7="16" captionTargetPageId-8="17" captionText-0="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." captionText-1="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." captionText-2="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." captionText-3="Figure 5. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Dentaries in left lateral (A), left medial (B), left dorsal (C), right lateral (D), and right medial (E) views. Scale bars=10 cm." captionText-4="Figure 6. Ectenosaurus tlemonectes sp. nov., YPM VP 4673, A–D. Detail of pathologic and non-pathologic anterior terminus of left dentary in left lateral (nonpathologic) (A), right lateral (pathologic) (B), right medial (pathologic) (C), and right dorsal (pathologic) (D) views. Scale bars for A–C=5 cm. Scale bar for D=3 cm." captionText-5="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." captionText-6="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." captionText-7="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." captionText-8="Figure 10. Ectenosaurus tlemonectes sp. nov., YPM VP4673. Cervical vertebra in left lateral view. Arrow indicates ventral ridge connecting the hypapophyseal peduncle and cotyle. Scale bar=5 cm." figureDoi-0="http://doi.org/10.5281/zenodo.13888170" figureDoi-1="http://doi.org/10.5281/zenodo.13888172" figureDoi-2="http://doi.org/10.5281/zenodo.13888174" figureDoi-3="http://doi.org/10.5281/zenodo.13888176" figureDoi-4="http://doi.org/10.5281/zenodo.13888178" figureDoi-5="http://doi.org/10.5281/zenodo.13888182" figureDoi-6="http://doi.org/10.5281/zenodo.13888184" figureDoi-7="http://doi.org/10.5281/zenodo.13888188" figureDoi-8="http://doi.org/10.5281/zenodo.13888190" httpUri-0="https://zenodo.org/record/13888170/files/figure.png" httpUri-1="https://zenodo.org/record/13888172/files/figure.png" httpUri-2="https://zenodo.org/record/13888174/files/figure.png" httpUri-3="https://zenodo.org/record/13888176/files/figure.png" httpUri-4="https://zenodo.org/record/13888178/files/figure.png" httpUri-5="https://zenodo.org/record/13888182/files/figure.png" httpUri-6="https://zenodo.org/record/13888184/files/figure.png" httpUri-7="https://zenodo.org/record/13888188/files/figure.png" httpUri-8="https://zenodo.org/record/13888190/files/figure.png" pageId="6" pageNumber="5">FIGS. 2–10</figureCitation> </paragraph> <paragraph id="8B80FD79FF9F672CFF3C1534852DFE20" blockId="6.[112,800,341,402]" pageId="6" pageNumber="5"> <emphasis id="B94B216BFF9F672CFF3C1534864DFEDD" bold="true" box="[144,354,341,367]" pageId="6" pageNumber="5"> @@ -186,13 +189,13 @@ of the ( <collectingMunicipality id="6BE46703FF9F672CFB79169784FFFC81" pageId="6" pageNumber="5">upper Coniacian-lower Campanian</collectingMunicipality> ; -<figureCitation id="1304E1FCFF9F672CFC761778831FFC81" box="[986,1072,793,819]" captionStart="Figure 1" captionStartId="4.[112,183,1509,1531]" captionTargetBox="[244,1389,216,1482]" captionTargetId="figure-192@4.[225,1403,191,1493]" captionTargetPageId="4" captionText="Figure 1. Geography and Stratigraphy. A. Generalized map of the United States during the Santonian Stage of the Cretaceous (~85 mya) with localities for YPM VP4673 (1) and ALMNH:Paleo:4673 (2). B. Generalized correlation chart of Coniacian to Campanian surface stratigraphy for Kansas and Alabama, USA. Base map data modified from Blakey (2014). Stratigraphic chart modified from Szabo et al. (1988) and Everhart (2014)." pageId="6" pageNumber="5">Fig. 1B</figureCitation> +<figureCitation id="1304E1FCFF9F672CFC761778831FFC81" box="[986,1072,793,819]" captionStart="Figure 1" captionStartId="4.[112,183,1509,1531]" captionTargetBox="[244,1389,216,1482]" captionTargetId="figure-192@4.[225,1403,191,1493]" captionTargetPageId="4" captionText="Figure 1. Geography and Stratigraphy. A. Generalized map of the United States during the Santonian Stage of the Cretaceous (~85 mya) with localities for YPM VP4673 (1) and ALMNH:Paleo:4673 (2). B. Generalized correlation chart of Coniacian to Campanian surface stratigraphy for Kansas and Alabama, USA. Base map data modified from Blakey (2014). Stratigraphic chart modified from Szabo et al. (1988) and Everhart (2014)." figureDoi="http://doi.org/10.5281/zenodo.13888168" httpUri="https://zenodo.org/record/13888168/files/figure.png" pageId="6" pageNumber="5">Fig. 1B</figureCitation> ), <collectingCounty id="62E185F5FF9F672CFBE9177883CDFC81" box="[1093,1250,793,819]" pageId="6" pageNumber="5">Gove County</collectingCounty> , <collectingRegion id="49FB339BFF9F672CFB47177B826BFC86" box="[1259,1348,794,820]" country="United States of America" name="Kansas" pageId="6" pageNumber="5">Kansas</collectingRegion> ( -<figureCitation id="1304E1FCFF9F672CFAFE17788282FC81" box="[1362,1453,793,819]" captionStart="Figure 1" captionStartId="4.[112,183,1509,1531]" captionTargetBox="[244,1389,216,1482]" captionTargetId="figure-192@4.[225,1403,191,1493]" captionTargetPageId="4" captionText="Figure 1. Geography and Stratigraphy. A. Generalized map of the United States during the Santonian Stage of the Cretaceous (~85 mya) with localities for YPM VP4673 (1) and ALMNH:Paleo:4673 (2). B. Generalized correlation chart of Coniacian to Campanian surface stratigraphy for Kansas and Alabama, USA. Base map data modified from Blakey (2014). Stratigraphic chart modified from Szabo et al. (1988) and Everhart (2014)." pageId="6" pageNumber="5">Fig. 1A</figureCitation> +<figureCitation id="1304E1FCFF9F672CFAFE17788282FC81" box="[1362,1453,793,819]" captionStart="Figure 1" captionStartId="4.[112,183,1509,1531]" captionTargetBox="[244,1389,216,1482]" captionTargetId="figure-192@4.[225,1403,191,1493]" captionTargetPageId="4" captionText="Figure 1. Geography and Stratigraphy. A. Generalized map of the United States during the Santonian Stage of the Cretaceous (~85 mya) with localities for YPM VP4673 (1) and ALMNH:Paleo:4673 (2). B. Generalized correlation chart of Coniacian to Campanian surface stratigraphy for Kansas and Alabama, USA. Base map data modified from Blakey (2014). Stratigraphic chart modified from Szabo et al. (1988) and Everhart (2014)." figureDoi="http://doi.org/10.5281/zenodo.13888168" httpUri="https://zenodo.org/record/13888168/files/figure.png" pageId="6" pageNumber="5">Fig. 1A</figureCitation> ). <location id="8EE0ABA2FF9F672CFA6C177882DFFC81" LSID="urn:lsid:plazi:treatment:03964C6FFF9F673BFF72149C8526F9CF:8EE0ABA2FF9F672CFA6C177882DFFC81" box="[1472,1520,793,819]" county="Gove County" municipality="upper Coniacian-lower Campanian" name="The" pageId="6" pageNumber="5" stateProvince="Kansas">The</location> precise locality and stratigraphic level within the @@ -213,7 +216,7 @@ Species name from Τλήμων [tlḗmōn], Classical Greek for "patient&qu <paragraph id="8B80FD79FF9F6722FCCC1014840EFDD2" blockId="6.[832,1520,1141,1943]" lastBlockId="8.[112,801,195,1946]" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5"> <emphasis id="B94B216BFF9F672CFCCC101484F1FB23" bold="true" box="[864,990,1141,1169]" italics="true" pageId="6" pageNumber="5">Frontal—</emphasis> The frontal ( -<figureCitation id="1304E1FCFF9F672CFBD9101783C5FB23" box="[1141,1258,1142,1169]" captionStart="Figure 2" captionStartId="7.[110,181,1899,1921]" captionTargetBox="[202,1418,204,1874]" captionTargetId="figure-64@7.[155,1457,192,1894]" captionTargetPageId="7" captionText="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="6" pageNumber="5">Fig. 2A, B</figureCitation> +<figureCitation id="1304E1FCFF9F672CFBD9101783C5FB23" box="[1141,1258,1142,1169]" captionStart="Figure 2" captionStartId="7.[110,181,1899,1921]" captionTargetBox="[202,1418,204,1874]" captionTargetId="figure-64@7.[155,1457,192,1894]" captionTargetPageId="7" captionText="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888170" httpUri="https://zenodo.org/record/13888170/files/figure.png" pageId="6" pageNumber="5">Fig. 2A, B</figureCitation> ) is an elongate, triangular bone with very slight antoribital bulges and short, roughly triangular posterolateral alae. The posterior quarter of the frontal is weathered and broken into three fragments. As preserved, the midline length is ~ <quantity id="4CC7509CFF9F672CFA161162845DFAF3" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="5" unit="mm" value="200.0">200 mm</quantity> , with a maximum width of @@ -221,7 +224,7 @@ The frontal ( and an interorbital width of <quantity id="4CC7509CFF9F672CFBA011288348FAD6" box="[1036,1127,1353,1380]" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="6" pageNumber="5" unit="mm" value="68.0">68 mm</quantity> . A very well-developed median dorsal ridge ( -<figureCitation id="1304E1FCFF9F672CFC4A110D836DFA34" box="[998,1090,1388,1414]" captionStart="Figure 2" captionStartId="7.[110,181,1899,1921]" captionTargetBox="[202,1418,204,1874]" captionTargetId="figure-64@7.[155,1457,192,1894]" captionTargetPageId="7" captionText="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="6" pageNumber="5">Fig. 2A</figureCitation> +<figureCitation id="1304E1FCFF9F672CFC4A110D836DFA34" box="[998,1090,1388,1414]" captionStart="Figure 2" captionStartId="7.[110,181,1899,1921]" captionTargetBox="[202,1418,204,1874]" captionTargetId="figure-64@7.[155,1457,192,1894]" captionTargetPageId="7" captionText="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888170" httpUri="https://zenodo.org/record/13888170/files/figure.png" pageId="6" pageNumber="5">Fig. 2A</figureCitation> ) extends at least three-quarters of the frontal's length (~ <quantity id="4CC7509CFF9F672CFBF411F183CBFA18" box="[1112,1252,1424,1450]" metricMagnitude="-1" metricUnit="m" metricValue="1.328" pageId="6" pageNumber="5" unit="mm" value="132.8">132.8 mm</quantity> ), widening markedly posteriorly (from @@ -229,7 +232,7 @@ and an interorbital width of to <quantity id="4CC7509CFF9F672CFB0C11D2822EFA7F" box="[1184,1281,1459,1485]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="5" unit="mm" value="8.6">8.6 mm</quantity> ). Ventrally ( -<figureCitation id="1304E1FCFF9F672CFA3F11D2847BFA42" captionStart="Figure 2" captionStartId="7.[110,181,1899,1921]" captionTargetBox="[202,1418,204,1874]" captionTargetId="figure-64@7.[155,1457,192,1894]" captionTargetPageId="7" captionText="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." pageId="6" pageNumber="5">Fig. 2B, C</figureCitation> +<figureCitation id="1304E1FCFF9F672CFA3F11D2847BFA42" captionStart="Figure 2" captionStartId="7.[110,181,1899,1921]" captionTargetBox="[202,1418,204,1874]" captionTargetId="figure-64@7.[155,1457,192,1894]" captionTargetPageId="7" captionText="Figure 2. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A-C. Frontal in dorsal (A) and ventral (B) views. C. Detail of olfactory canal. Abbreviations: POF, excavation for postorbitofrontal; PRF, excavation for prefrontal; VSR, ventral separation ridge. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888170" httpUri="https://zenodo.org/record/13888170/files/figure.png" pageId="6" pageNumber="5">Fig. 2B, C</figureCitation> ), a pair of broad, rounded, and roughly violin-shaped descending processes (the crista cranii frontalis of some authors) bracket the olfactory tract, almost completely enclosing it for nearly half its length ( <quantity id="4CC7509CFF9F672CFAB6125E82A0F9E8" box="[1306,1423,1599,1626]" metricMagnitude="-2" metricUnit="m" metricValue="5.93" pageId="6" pageNumber="5" unit="mm" value="59.3">59.3 mm</quantity> ) before opening anteromedially into a flat amygdaliform sulcus to accommodate the olfactory lobes. Though neither the prefrontals nor postorbitofrontals are preserved, articulation between these elements and the frontal is evident from well-defined, striated depressions on the ventral surface of the frontal. These articular surfaces are completely separated from one another by a wedge or ridge (the ventral separation ridge of @@ -266,7 +269,7 @@ Mosasauridae ( </taxonomicName> . The contact with the parietal is roughly transverse and possesses no prominent posteromedial projections onto the parietal. The parietal is not preserved, and it is unclear whether, and to what degree, it may have invaded/overlapped the frontal anteromedially. </paragraph> -<caption id="DF40ADF1FF9E672DFFC2130A82B1F82B" pageId="7" pageNumber="6" startId="7.[110,181,1899,1921]" targetBox="[202,1418,204,1874]" targetPageId="7" targetType="figure"> +<caption id="DF40ADF1FF9E672DFFC2130A82B1F82B" ID-DOI="http://doi.org/10.5281/zenodo.13888170" ID-Zenodo-Dep="13888170" httpUri="https://zenodo.org/record/13888170/files/figure.png" pageId="7" pageNumber="6" startId="7.[110,181,1899,1921]" targetBox="[202,1418,204,1874]" targetPageId="7" targetType="figure"> <paragraph id="8B80FD79FF9E672DFFC2130A82B1F82B" blockId="7.[110,1509,1899,1946]" pageId="7" pageNumber="6"> <emphasis id="B94B216BFF9E672DFFC2130A87E1F833" bold="true" box="[110,206,1899,1921]" pageId="7" pageNumber="6">Figure 2.</emphasis> <taxonomicName id="4C3F86FAFF9E672DFF7F130A86F9F830" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[211,470,1899,1922]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="6" phylum="Chordata" rank="species" species="tlemonectes" status="sp. nov."> @@ -293,7 +296,7 @@ Mosasauridae ( <paragraph id="8B80FD79FF916722FF3C1606857FFA38" blockId="8.[112,801,195,1946]" pageId="8" pageNumber="7"> <emphasis id="B94B216BFF916722FF3C1606853EFD31" bold="true" box="[144,529,615,644]" italics="true" pageId="8" pageNumber="7">Quadrates and Squamosals—</emphasis> The left quadrate ( -<figureCitation id="1304E1FCFF916722FD5D16088799FD15" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." pageId="8" pageNumber="7">Fig. 3A‒C</figureCitation> +<figureCitation id="1304E1FCFF916722FD5D16088799FD15" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888172" httpUri="https://zenodo.org/record/13888172/files/figure.png" pageId="8" pageNumber="7">Fig. 3A‒C</figureCitation> ) is reasonably complete but lacks most of the tympanic ala and portions of the anterior quadratic shaft. The suprastapedial process is medially deflected and is large and elongate, as is typical of plioplatecarpines, measuring ~60% of the quadrate height. The borders of the suprastapedial process are straight medially, slightly concave laterally, and expand distally to form a bulbous, irregularly rounded extremity that is fused with the infrastapedial process. In the case of <taxonomicName id="4C3F86FAFF916722FDE917C485C6FC73" authorityName="Russell" authorityYear="1967" box="[581,745,933,961]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="7" phylum="Chordata" rank="genus"> <emphasis id="B94B216BFF916722FDE917C485C6FC73" box="[581,745,933,961]" italics="true" pageId="8" pageNumber="7">Ectenosaurus</emphasis> @@ -335,11 +338,11 @@ The suprastapedial pit is severely distorted anteroposteriorly, obscuring its or </paragraph> <paragraph id="8B80FD79FF916722FF3C135882C3FE03" blockId="8.[112,801,195,1946]" lastBlockId="8.[832,1521,195,1946]" pageId="8" pageNumber="7"> The posterior end of the right squamosal ( -<figureCitation id="1304E1FCFF916722FD3013588437F8E1" box="[668,792,1849,1875]" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." pageId="8" pageNumber="7">Fig. 3D‒F</figureCitation> +<figureCitation id="1304E1FCFF916722FD3013588437F8E1" box="[668,792,1849,1875]" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888172" httpUri="https://zenodo.org/record/13888172/files/figure.png" pageId="8" pageNumber="7">Fig. 3D‒F</figureCitation> ) is preserved and in good condition. The medial face ( -<figureCitation id="1304E1FCFF916722FFD513E187FEF828" box="[121,209,1920,1946]" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." pageId="8" pageNumber="7">Fig. 3E</figureCitation> +<figureCitation id="1304E1FCFF916722FFD513E187FEF828" box="[121,209,1920,1946]" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888172" httpUri="https://zenodo.org/record/13888172/files/figure.png" pageId="8" pageNumber="7">Fig. 3E</figureCitation> ) is marked by an elaborate series of ridges and troughs that would have received corresponding surfaces on the supratemporal. Numerous large foramina are also present. The ventroposterior fossa for articulation with the quadrate ( -<figureCitation id="1304E1FCFF916722FC5D154D8361FEF5" box="[1009,1102,300,327]" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." pageId="8" pageNumber="7">Fig. 3D</figureCitation> +<figureCitation id="1304E1FCFF916722FC5D154D8361FEF5" box="[1009,1102,300,327]" captionStart="Figure 3" captionStartId="9.[112,183,1614,1636]" captionTargetBox="[172,1437,217,1589]" captionTargetId="figure-185@9.[125,1482,192,1603]" captionTargetPageId="9" captionText="Figure 3. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–C. Left quadrate in medial (A), anterior (B), and posterior (C) views. D–F. Left squamosal in ventral (D), lateral (E), and medial (F) views. Arrow indicates suprastapedial/infrastapedial contact. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888172" httpUri="https://zenodo.org/record/13888172/files/figure.png" pageId="8" pageNumber="7">Fig. 3D</figureCitation> ) forms a rough isosceles triangle and is shallowly concave. Anterodorsally there is a small but distinct knob marking the posterior terminus of the groove for insertion of the postorbitofrontal processes. </paragraph> <paragraph id="8B80FD79FF916722FCCC15D98322FB98" blockId="8.[832,1521,195,1946]" pageId="8" pageNumber="7"> @@ -347,18 +350,18 @@ The posterior end of the right squamosal ( The premaxilla and the majority of both maxillae and dentaries are present in <collectionCode id="ED2E65BCFF916722FA1B15BC82DFFE45" box="[1463,1520,477,503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="8" pageNumber="7" type="Museum">YPM</collectionCode> VP4673. The premaxilla ( -<figureCitation id="1304E1FCFF916722FB291661822BFDA8" box="[1157,1284,512,538]" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." pageId="8" pageNumber="7">Fig. 4A‒C</figureCitation> +<figureCitation id="1304E1FCFF916722FB291661822BFDA8" box="[1157,1284,512,538]" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." figureDoi="http://doi.org/10.5281/zenodo.13888174" httpUri="https://zenodo.org/record/13888174/files/figure.png" pageId="8" pageNumber="7">Fig. 4A‒C</figureCitation> ) is well preserved and complete except for the posteriormost portion of the internarial bar. There is a very short, spatulate edentulous rostrum (5.0 mm). Dorsally, the premaxilla is planar and bears a small median keel or crest ( <quantity id="4CC7509CFF916722FA8E16EC8253FD15" box="[1314,1404,653,679]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="7" unit="mm" value="19.0">19 mm</quantity> ) near the base of the internarial bar that is subtriangular in lateral profile ( -<figureCitation id="1304E1FCFF916722FC0816B2832FFD5F" box="[932,1024,723,749]" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." pageId="8" pageNumber="7">Fig. 4C</figureCitation> +<figureCitation id="1304E1FCFF916722FC0816B2832FFD5F" box="[932,1024,723,749]" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." figureDoi="http://doi.org/10.5281/zenodo.13888174" httpUri="https://zenodo.org/record/13888174/files/figure.png" pageId="8" pageNumber="7">Fig. 4C</figureCitation> ). This structure has not been observed in any other specimen of <emphasis id="B94B216BFF916722FB2016948218FCA3" box="[1164,1335,757,785]" italics="true" pageId="8" pageNumber="7"> <taxonomicName id="4C3F86FAFF916722FB201694821CFCA3" authorityName="Russell" authorityYear="1967" box="[1164,1331,757,785]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="7" phylum="Chordata" rank="genus">Ectenosaurus</taxonomicName> . </emphasis> The posterior terminus of the premaxillary-maxillary suture occurs at the deepest anterior portion of the maxilla ( -<figureCitation id="1304E1FCFF916722FACB175D82CDFCE5" box="[1383,1506,828,855]" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." pageId="8" pageNumber="7">Fig. 4D, E</figureCitation> +<figureCitation id="1304E1FCFF916722FACB175D82CDFCE5" box="[1383,1506,828,855]" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." figureDoi="http://doi.org/10.5281/zenodo.13888174" httpUri="https://zenodo.org/record/13888174/files/figure.png" pageId="8" pageNumber="7">Fig. 4D, E</figureCitation> ), at a position even with the fourth maxillary tooth. The internarial bar narrows abruptly behind the premaxillomaxillary suture (the "step-wise constriction" described by <bibRefCitation id="EFAE8088FF916722FCC417A8834BFC51" author="Willman, A. J. & T. Konishi & M. W. Caldwell" box="[872,1124,969,996]" pageId="8" pageNumber="7" pagination="741 - 755" refId="ref18000" refString="Willman, A. J., T. Konishi, and M. W. Caldwell. 2021. A new species of Ectenosaurus (Mosasauridae: Plioplatecarpinae) from western Kansas, USA, reveals a novels suite of osteological characters for the genus. Canadian Journal of Earth Science 58 (9): 741 - 755. [https: // doi. org / 10.1139 / cjes- 2020 - 0175]." type="journal article" year="2021">Willman et al. 2021</bibRefCitation> ), is triangular in cross section, and expands posteriorly to receive the anterior process from the frontal. @@ -382,7 +385,7 @@ VP4673, a distance of 3.5 tooth bases, which is slightly longer than that of </paragraph> <paragraph id="8B80FD79FF916723FCCC11B885C3F887" blockId="8.[832,1521,195,1946]" lastBlockId="9.[112,800,1713,1951]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7"> The dentaries ( -<figureCitation id="1304E1FCFF916722FBB111B883B5FA46" box="[1053,1178,1497,1524]" captionStart="Figure 5" captionStartId="11.[112,183,1463,1485]" captionTargetBox="[178,1453,216,1431]" captionTargetId="figure-261@11.[128,1499,191,1445]" captionTargetPageId="11" captionText="Figure 5. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Dentaries in left lateral (A), left medial (B), left dorsal (C), right lateral (D), and right medial (E) views. Scale bars=10 cm." pageId="8" pageNumber="7">Fig. 5A‒E</figureCitation> +<figureCitation id="1304E1FCFF916722FBB111B883B5FA46" box="[1053,1178,1497,1524]" captionStart="Figure 5" captionStartId="11.[112,183,1463,1485]" captionTargetBox="[178,1453,216,1431]" captionTargetId="figure-261@11.[128,1499,191,1445]" captionTargetPageId="11" captionText="Figure 5. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Dentaries in left lateral (A), left medial (B), left dorsal (C), right lateral (D), and right medial (E) views. Scale bars=10 cm." figureDoi="http://doi.org/10.5281/zenodo.13888176" httpUri="https://zenodo.org/record/13888176/files/figure.png" pageId="8" pageNumber="7">Fig. 5A‒E</figureCitation> ) are gracile, delicately constructed bones, more so than in either <taxonomicName id="4C3F86FAFF916722FAEB119A84FBF988" baseAuthorityName="Merriam" baseAuthorityYear="1894" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="7" phylum="Chordata" rank="species" species="clidastoides"> <emphasis id="B94B216BFF916722FAEB119A84FBF988" italics="true" pageId="8" pageNumber="7">Ectenosaurus clidastoides</emphasis> @@ -413,7 +416,7 @@ is incipient in </emphasis> where the dental formula is 2=~12=13, nearer to the formula seen in most plioplatecarpines (2=12=12). </paragraph> -<caption id="DF40ADF1FF906723FFDC122F82F8F9CE" pageId="9" pageNumber="8" startId="9.[112,183,1614,1636]" targetBox="[172,1437,217,1589]" targetPageId="9" targetType="figure"> +<caption id="DF40ADF1FF906723FFDC122F82F8F9CE" ID-DOI="http://doi.org/10.5281/zenodo.13888172" ID-Zenodo-Dep="13888172" httpUri="https://zenodo.org/record/13888172/files/figure.png" pageId="9" pageNumber="8" startId="9.[112,183,1614,1636]" targetBox="[172,1437,217,1589]" targetPageId="9" targetType="figure"> <paragraph id="8B80FD79FF906723FFDC122F82F8F9CE" blockId="9.[112,1495,1613,1660]" pageId="9" pageNumber="8"> <emphasis id="B94B216BFF906723FFDC122F87FFF9D1" bold="true" box="[112,208,1614,1636]" pageId="9" pageNumber="8">Figure 3.</emphasis> <taxonomicName id="4C3F86FAFF906723FF79122C86F7F9D6" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[213,472,1613,1636]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="8" phylum="Chordata" rank="species" species="tlemonectes" status="sp. nov."> @@ -468,7 +471,7 @@ explained this as a case convergent evolution, a conclusion with which we concur </paragraph> <paragraph id="8B80FD79FF906720FCCC13E58548F82D" blockId="9.[832,1520,1711,1951]" lastBlockId="10.[112,800,1713,1951]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8"> A small edentulous prow is present on the dentary, which on the right is severely deformed, probably from a healed injury ( -<figureCitation id="1304E1FCFF936720FE9D12B58659F95D" box="[305,374,1748,1775]" captionStart="Figure 6" captionStartId="12.[113,184,1350,1372]" captionTargetBox="[142,1483,218,1315]" captionTargetId="figure-254@12.[112,1519,191,1332]" captionTargetPageId="12" captionText="Figure 6. Ectenosaurus tlemonectes sp. nov., YPM VP 4673, A–D. Detail of pathologic and non-pathologic anterior terminus of left dentary in left lateral (nonpathologic) (A), right lateral (pathologic) (B), right medial (pathologic) (C), and right dorsal (pathologic) (D) views. Scale bars for A–C=5 cm. Scale bar for D=3 cm." pageId="10" pageNumber="9">Fig. 6</figureCitation> +<figureCitation id="1304E1FCFF936720FE9D12B58659F95D" box="[305,374,1748,1775]" captionStart="Figure 6" captionStartId="12.[113,184,1350,1372]" captionTargetBox="[142,1483,218,1315]" captionTargetId="figure-254@12.[112,1519,191,1332]" captionTargetPageId="12" captionText="Figure 6. Ectenosaurus tlemonectes sp. nov., YPM VP 4673, A–D. Detail of pathologic and non-pathologic anterior terminus of left dentary in left lateral (nonpathologic) (A), right lateral (pathologic) (B), right medial (pathologic) (C), and right dorsal (pathologic) (D) views. Scale bars for A–C=5 cm. Scale bar for D=3 cm." figureDoi="http://doi.org/10.5281/zenodo.13888178" httpUri="https://zenodo.org/record/13888178/files/figure.png" pageId="10" pageNumber="9">Fig. 6</figureCitation> ); the deformation involves the first dentary tooth. A very large and elongate, longitudinally oriented foramen is present below the tenth tooth position, a character also present in <emphasis id="B94B216BFF936720FE51135D85E4F8EA" box="[509,715,1852,1880]" italics="true" pageId="10" pageNumber="9"> <taxonomicName id="4C3F86FAFF936720FE51135D85E9F8EA" box="[509,710,1852,1880]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="9" phylum="Chordata" rank="species" species="everhartorum">E. everhartorum</taxonomicName> @@ -478,7 +481,7 @@ where the foramen occurs between the tenth and eleventh dentary teeth ( <bibRefCitation id="EFAE8088FF936720FE8713E58579F82C" author="Willman, A. J. & T. Konishi & M. W. Caldwell" box="[299,598,1924,1951]" pageId="10" pageNumber="9" pagination="741 - 755" refId="ref18000" refString="Willman, A. J., T. Konishi, and M. W. Caldwell. 2021. A new species of Ectenosaurus (Mosasauridae: Plioplatecarpinae) from western Kansas, USA, reveals a novels suite of osteological characters for the genus. Canadian Journal of Earth Science 58 (9): 741 - 755. [https: // doi. org / 10.1139 / cjes- 2020 - 0175]." type="journal article" year="2021">Willman et al. 2021:746</bibRefCitation> ). </paragraph> -<caption id="DF40ADF1FF936720FFDC122E8418F9CC" pageId="10" pageNumber="9" startId="10.[112,183,1615,1637]" targetBox="[149,1487,223,1589]" targetPageId="10" targetType="figure"> +<caption id="DF40ADF1FF936720FFDC122E8418F9CC" ID-DOI="http://doi.org/10.5281/zenodo.13888174" ID-Zenodo-Dep="13888174" httpUri="https://zenodo.org/record/13888174/files/figure.png" pageId="10" pageNumber="9" startId="10.[112,183,1615,1637]" targetBox="[149,1487,223,1589]" targetPageId="10" targetType="figure"> <paragraph id="8B80FD79FF936720FFDC122E8418F9CC" blockId="10.[112,1506,1615,1663]" pageId="10" pageNumber="9"> <emphasis id="B94B216BFF936720FFDC122E87FFF9D7" bold="true" box="[112,208,1615,1637]" pageId="10" pageNumber="9">Figure 4.</emphasis> <taxonomicName id="4C3F86FAFF936720FF79122E86F7F9D4" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[213,472,1615,1638]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="10" pageNumber="9" phylum="Chordata" rank="species" species="tlemonectes" status="sp. nov."> @@ -513,14 +516,14 @@ A fragment of the left pterygoid is present, preserving the bases of seven teeth <paragraph id="8B80FD79FF936721FCCC13E382DFF98F" blockId="10.[832,1520,1713,1950]" lastBlockId="11.[832,1520,1571,1949]" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9"> <emphasis id="B94B216BFF936720FCCC13E384FFF82C" bold="true" box="[864,976,1922,1950]" italics="true" pageId="10" pageNumber="9">Jugals—</emphasis> Portions of both jugals are preserved ( -<figureCitation id="1304E1FCFF936721FA6D13E28799F98F" captionStart="Figure 7" captionStartId="13.[112,183,1899,1921]" captionTargetBox="[245,1374,215,1847]" captionTargetId="figure-15@13.[171,1448,192,1872]" captionTargetPageId="13" captionText="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">Fig. 7C‒F</figureCitation> +<figureCitation id="1304E1FCFF936721FA6D13E28799F98F" captionStart="Figure 7" captionStartId="13.[112,183,1899,1921]" captionTargetBox="[245,1374,215,1847]" captionTargetId="figure-15@13.[171,1448,192,1872]" captionTargetPageId="13" captionText="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." figureDoi="http://doi.org/10.5281/zenodo.13888182" httpUri="https://zenodo.org/record/13888182/files/figure.png" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">Fig. 7C‒F</figureCitation> ). Posteroventral processes are present on both, though a considerable degree of asymmetry exists between the two elements. The process is small on the right jugal ( -<figureCitation id="1304E1FCFF926721FF1612ED861EF915" box="[186,305,1676,1703]" captionStart="Figure 7" captionStartId="13.[112,183,1899,1921]" captionTargetBox="[245,1374,215,1847]" captionTargetId="figure-15@13.[171,1448,192,1872]" captionTargetPageId="13" captionText="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." pageId="11" pageNumber="10">Fig. 7C, D</figureCitation> +<figureCitation id="1304E1FCFF926721FF1612ED861EF915" box="[186,305,1676,1703]" captionStart="Figure 7" captionStartId="13.[112,183,1899,1921]" captionTargetBox="[245,1374,215,1847]" captionTargetId="figure-15@13.[171,1448,192,1872]" captionTargetPageId="13" captionText="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." figureDoi="http://doi.org/10.5281/zenodo.13888182" httpUri="https://zenodo.org/record/13888182/files/figure.png" pageId="11" pageNumber="10">Fig. 7C, D</figureCitation> ), but far more developed on the left ( -<figureCitation id="1304E1FCFF926721FD5D12ED879DF978" captionStart="Figure 7" captionStartId="13.[112,183,1899,1921]" captionTargetBox="[245,1374,215,1847]" captionTargetId="figure-15@13.[171,1448,192,1872]" captionTargetPageId="13" captionText="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." pageId="11" pageNumber="10">Fig. 7E, F</figureCitation> +<figureCitation id="1304E1FCFF926721FD5D12ED879DF978" captionStart="Figure 7" captionStartId="13.[112,183,1899,1921]" captionTargetBox="[245,1374,215,1847]" captionTargetId="figure-15@13.[171,1448,192,1872]" captionTargetPageId="13" captionText="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." figureDoi="http://doi.org/10.5281/zenodo.13888182" httpUri="https://zenodo.org/record/13888182/files/figure.png" pageId="11" pageNumber="10">Fig. 7E, F</figureCitation> ), where it forms a scalloped posterior border for the vertical ramus. While we suspect this asymmetry is the result of pathology, additional examples of the species will be needed to determine which jugal exhibits the abnormality. A deep triangular depression on the lateral surface of the vertical ramus marks the point of articulation with the postorbitofrontal. The vertical ramus is short, less than 50% of the length of the horizontal ramus. </paragraph> -<caption id="DF40ADF1FF926721FFDC11D6856EFA54" pageId="11" pageNumber="10" startId="11.[112,183,1463,1485]" targetBox="[178,1453,216,1431]" targetPageId="11" targetType="figure"> +<caption id="DF40ADF1FF926721FFDC11D6856EFA54" ID-DOI="http://doi.org/10.5281/zenodo.13888176" ID-Zenodo-Dep="13888176" httpUri="https://zenodo.org/record/13888176/files/figure.png" pageId="11" pageNumber="10" startId="11.[112,183,1463,1485]" targetBox="[178,1453,216,1431]" targetPageId="11" targetType="figure"> <paragraph id="8B80FD79FF926721FFDC11D6856EFA54" blockId="11.[112,1517,1463,1511]" pageId="11" pageNumber="10"> <emphasis id="B94B216BFF926721FFDC11D687E0FA7F" bold="true" box="[112,207,1463,1485]" pageId="11" pageNumber="10">Figure 5.</emphasis> <taxonomicName id="4C3F86FAFF926721FF7911D686F8FA7C" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[213,471,1463,1486]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="11" pageNumber="10" phylum="Chordata" rank="species" species="tlemonectes" status="sp. nov."> @@ -547,7 +550,7 @@ Portions of both jugals are preserved ( In most russellosaurines, the coronoid is a short, saddle-shaped wedge of bone straddling the anterodorsal margin of the surangular, which it encloses to varying degrees between simple medial and lateral descending processes or wings that reach the prearticular. In the plioplatecarpines, these wings are usually short, compared with those of mosasaurines. In <collectionCode id="ED2E65BCFF926721FAEB137B82AFF886" box="[1351,1408,1818,1844]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="11" pageNumber="10" type="Museum">YPM</collectionCode> VP4673, only the right coronoid ( -<figureCitation id="1304E1FCFF926721FBD8135D83DFF8E5" box="[1140,1264,1852,1879]" captionStart="Figure 7" captionStartId="13.[112,183,1899,1921]" captionTargetBox="[245,1374,215,1847]" captionTargetId="figure-15@13.[171,1448,192,1872]" captionTargetPageId="13" captionText="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." pageId="11" pageNumber="10">Fig. 7A, B</figureCitation> +<figureCitation id="1304E1FCFF926721FBD8135D83DFF8E5" box="[1140,1264,1852,1879]" captionStart="Figure 7" captionStartId="13.[112,183,1899,1921]" captionTargetBox="[245,1374,215,1847]" captionTargetId="figure-15@13.[171,1448,192,1872]" captionTargetPageId="13" captionText="Figure 7. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Right coronoid in lateral (A) and medial (B) views. C, D. Left jugal in lateral (C) and medial (D) views. E, F. Right jugal in lateral (E) and medial (F) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm." figureDoi="http://doi.org/10.5281/zenodo.13888182" httpUri="https://zenodo.org/record/13888182/files/figure.png" pageId="11" pageNumber="10">Fig. 7A, B</figureCitation> ) is preserved and is slightly crushed lateromedially.As with most russellosaurines, the coronoid is a slender, short, selliform element divided into lateral and medial wings which straddle the anterodorsal surface of the surangular. The coronoid dorsal margin is concave, and the ascending posterior process was well-developed, though the apex of the crest is missing. The stout medial wing would have overlapped the dorsomedial margin of the surangular, and the interior surface of this wing is highly vascularized, marked by numerous foramina. In contrast, the anterolateral wing is very short, and hardly more than a low, diagonal ridge. The shallow anteromedial bifurcation ("cleft" of <bibRefCitation id="EFAE8088FF956726FFDC1378866EF886" author="Russell, D. A." box="[112,321,1817,1844]" pageId="12" pageNumber="11" pagination="1 - 241" refId="ref16943" refString="Russell, D. A. 1967. Systematics and morphology of American mosasaurs (Reptilia, Sauria). Bulletin of the Peabody Museum of Natural History 23: 1 - 241." type="journal article" year="1967">Russell 1967:53</bibRefCitation> ) is long and well-defined, extending more than a third of the bone's length. We will discuss the coronoid of @@ -558,7 +561,7 @@ further in our description of <collectionCode id="ED2E65BCFF956726FF2313E3862EF82F" box="[143,257,1922,1949]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="12" pageNumber="11" type="Museum">ALMNH</collectionCode> :Paleo:5452. </paragraph> -<caption id="DF40ADF1FF956726FFDD11278566FA3C" pageId="12" pageNumber="11" startId="12.[113,184,1350,1372]" targetBox="[142,1483,218,1315]" targetPageId="12" targetType="figure"> +<caption id="DF40ADF1FF956726FFDD11278566FA3C" ID-DOI="http://doi.org/10.5281/zenodo.13888178" ID-Zenodo-Dep="13888178" httpUri="https://zenodo.org/record/13888178/files/figure.png" pageId="12" pageNumber="11" startId="12.[113,184,1350,1372]" targetBox="[142,1483,218,1315]" targetPageId="12" targetType="figure"> <paragraph id="8B80FD79FF956726FFDD11278566FA3C" blockId="12.[112,1505,1349,1423]" pageId="12" pageNumber="11"> <emphasis id="B94B216BFF956726FFDD112787FFFAE9" bold="true" box="[113,208,1350,1372]" pageId="12" pageNumber="11">Figure 6.</emphasis> <taxonomicName id="4C3F86FAFF956726FF7A112486F6FAEE" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[214,473,1349,1372]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="11" phylum="Chordata" rank="species" species="tlemonectes" status="sp. nov."> @@ -581,13 +584,13 @@ further in our description of <paragraph id="8B80FD79FF956725FCCC11D687D0FEB2" blockId="12.[832,1520,1463,1949]" lastBlockId="15.[112,801,194,1946]" lastPageId="15" lastPageNumber="14" pageId="12" pageNumber="11"> <emphasis id="B94B216BFF956726FCCC11D683F2FA61" bold="true" box="[864,1245,1463,1491]" italics="true" pageId="12" pageNumber="11">Articular and Surangular—</emphasis> The majority of both the left and right surangular-articular units are preserved ( -<figureCitation id="1304E1FCFF956726FC0E119E84C8F9AB" box="[930,999,1535,1561]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." pageId="12" pageNumber="11">Fig. 8</figureCitation> +<figureCitation id="1304E1FCFF956726FC0E119E84C8F9AB" box="[930,999,1535,1561]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888184" httpUri="https://zenodo.org/record/13888184/files/figure.png" pageId="12" pageNumber="11">Fig. 8</figureCitation> ), though the left is the most complete. The dorsal margin of the surangular is slightly concave and broadens anteriorly as it approaches the contact with the coronoid, expanding from a height of about 3–~ <quantity id="4CC7509CFF956726FA7312088447F915" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="11" unit="cm" value="9.0">9 cm</quantity> . The surangular-articular suture is obvious on the lateral side of the left mandible ( -<figureCitation id="1304E1FCFF956726FB7012CE8254F978" box="[1244,1403,1711,1738]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." pageId="12" pageNumber="11">Fig. 8A, B, D</figureCitation> +<figureCitation id="1304E1FCFF956726FB7012CE8254F978" box="[1244,1403,1711,1738]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888184" httpUri="https://zenodo.org/record/13888184/files/figure.png" pageId="12" pageNumber="11">Fig. 8A, B, D</figureCitation> ), curving beneath the glenoid fossa before running more or less straight beneath the coronoid to the intramadibular joint. The right and left articulars are both missing their posteroventral corners, but the ventral margins appear to have been generally straight. Most of the glenoid fossa is located on the articular. The posterodorsal tongue of the surangular terminates abruptly in a rugose, highly vascularized vertical face that forms the anterior wall of the glenoid fossa ( -<figureCitation id="1304E1FCFF976724FEDE132086E3F8E9" box="[370,460,1857,1883]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." pageId="14" pageNumber="13">Fig. 8C</figureCitation> +<figureCitation id="1304E1FCFF976724FEDE132086E3F8E9" box="[370,460,1857,1883]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888184" httpUri="https://zenodo.org/record/13888184/files/figure.png" pageId="14" pageNumber="13">Fig. 8C</figureCitation> ). <bibRefCitation id="EFAE8088FF976724FE4C13208434F8E9" author="Willman, A. J. & T. Konishi & M. W. Caldwell" box="[480,795,1857,1884]" pageId="14" pageNumber="13" pagination="741 - 755" refId="ref18000" refString="Willman, A. J., T. Konishi, and M. W. Caldwell. 2021. A new species of Ectenosaurus (Mosasauridae: Plioplatecarpinae) from western Kansas, USA, reveals a novels suite of osteological characters for the genus. Canadian Journal of Earth Science 58 (9): 741 - 755. [https: // doi. org / 10.1139 / cjes- 2020 - 0175]." type="journal article" year="2021">Willman et al. (2021:747</bibRefCitation> ; fig. 12) stressed the presence of a "distinct notch on the dorsal surface...just prior to the glenoid fossae" in @@ -600,14 +603,14 @@ The majority of both the left and right surangular-articular units are preserved and <collectionCode id="ED2E65BCFF976724FB4C129A8236F8A7" box="[1248,1305,1787,1813]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="14" pageNumber="13" type="Museum">YPM</collectionCode> VP4673 ( -<figureCitation id="1304E1FCFF976724FA3C129A82C8F8A7" box="[1424,1511,1787,1813]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." pageId="14" pageNumber="13">Fig. 8C</figureCitation> +<figureCitation id="1304E1FCFF976724FA3C129A82C8F8A7" box="[1424,1511,1787,1813]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888184" httpUri="https://zenodo.org/record/13888184/files/figure.png" pageId="14" pageNumber="13">Fig. 8C</figureCitation> ) as a character "absent in all other mosasauroids" and therefore considered diagnostic of the genus by those authors. In <collectionCode id="ED2E65BCFF976724FC7E13048324F8CD" box="[978,1035,1893,1919]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="14" pageNumber="13" type="Museum">YPM</collectionCode> VP4673, portions of the glenoid fossa are bordered dorsally and medially on both the angular and surangular by a thick, raised rim with rounded edges ( -<figureCitation id="1304E1FCFF966725FFD5148487DDFEB2" box="[121,242,229,256]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." pageId="15" pageNumber="14">Fig. 8B, C</figureCitation> +<figureCitation id="1304E1FCFF966725FFD5148487DDFEB2" box="[121,242,229,256]" captionStart="Figure 8" captionStartId="14.[112,183,1666,1688]" captionTargetBox="[177,1450,211,1623]" captionTargetId="figure-106@14.[117,1513,192,1649]" captionTargetPageId="14" captionText="Figure 8. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A, B. Left surangular-articular in lateral (A) and medial (B) views. C. Close-up of glenoid fossa in lateral view. D. Close-up of glenoid fossa in medial view. E. Close-up of glenoid fossa in dorsal view. Arrow indicates postglenoid notch. Scale bars=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888184" httpUri="https://zenodo.org/record/13888184/files/figure.png" pageId="15" pageNumber="14">Fig. 8B, C</figureCitation> ). </paragraph> -<caption id="DF40ADF1FF946727FFDC130A82B0F828" pageId="13" pageNumber="12" startId="13.[112,183,1899,1921]" targetBox="[245,1374,215,1847]" targetPageId="13" targetType="figure"> +<caption id="DF40ADF1FF946727FFDC130A82B0F828" ID-DOI="http://doi.org/10.5281/zenodo.13888182" ID-Zenodo-Dep="13888182" httpUri="https://zenodo.org/record/13888182/files/figure.png" pageId="13" pageNumber="12" startId="13.[112,183,1899,1921]" targetBox="[245,1374,215,1847]" targetPageId="13" targetType="figure"> <paragraph id="8B80FD79FF946727FFDC130A82B0F828" blockId="13.[112,1489,1899,1947]" pageId="13" pageNumber="12"> <emphasis id="B94B216BFF946727FFDC130A87FFF833" bold="true" box="[112,208,1899,1921]" pageId="13" pageNumber="12">Figure 7.</emphasis> <taxonomicName id="4C3F86FAFF946727FF79130A86F7F830" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[213,472,1899,1922]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="12" phylum="Chordata" rank="species" species="tlemonectes" status="sp. nov."> @@ -635,7 +638,7 @@ VP4673, portions of the glenoid fossa are bordered dorsally and medially on both ) views. Scale bars for A, B=5 cm. Scale bar for C–F=2 cm. </paragraph> </caption> -<caption id="DF40ADF1FF976724FFDC12E3869FF978" pageId="14" pageNumber="13" startId="14.[112,183,1666,1688]" targetBox="[177,1450,211,1623]" targetPageId="14" targetType="figure"> +<caption id="DF40ADF1FF976724FFDC12E3869FF978" ID-DOI="http://doi.org/10.5281/zenodo.13888184" ID-Zenodo-Dep="13888184" httpUri="https://zenodo.org/record/13888184/files/figure.png" pageId="14" pageNumber="13" startId="14.[112,183,1666,1688]" targetBox="[177,1450,211,1623]" targetPageId="14" targetType="figure"> <paragraph id="8B80FD79FF976724FFDC12E3869FF978" blockId="14.[112,1518,1665,1739]" pageId="14" pageNumber="13"> <emphasis id="B94B216BFF976724FFDC12E387FFF925" bold="true" box="[112,208,1666,1688]" pageId="14" pageNumber="13">Figure 8.</emphasis> <taxonomicName id="4C3F86FAFF976724FF7912E086F7F92A" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[213,472,1665,1688]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="13" phylum="Chordata" rank="species" species="tlemonectes" status="sp. nov."> @@ -660,7 +663,7 @@ VP4673, portions of the glenoid fossa are bordered dorsally and medially on both <paragraph id="8B80FD79FF966725FF3C156685F7FC51" blockId="15.[112,801,194,1946]" pageId="15" pageNumber="14"> <emphasis id="B94B216BFF966725FF3C156686BCFE91" bold="true" box="[144,403,263,291]" italics="true" pageId="15" pageNumber="14">Splenial-Angular—</emphasis> Portions of both splenials and angulars are present ( -<figureCitation id="1304E1FCFF966725FED7154D86DBFEF4" box="[379,500,300,326]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." pageId="15" pageNumber="14">Fig. 9A–L</figureCitation> +<figureCitation id="1304E1FCFF966725FED7154D86DBFEF4" box="[379,500,300,326]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." figureDoi="http://doi.org/10.5281/zenodo.13888188" httpUri="https://zenodo.org/record/13888188/files/figure.png" pageId="15" pageNumber="14">Fig. 9A–L</figureCitation> ), including the articulating faces for all four elements. <bibRefCitation id="EFAE8088FF966725FE4F152E840FFED8" author="Willman, A. J. & T. Konishi & M. W. Caldwell" box="[483,800,335,362]" pageId="15" pageNumber="14" pagination="741 - 755" refId="ref18000" refString="Willman, A. J., T. Konishi, and M. W. Caldwell. 2021. A new species of Ectenosaurus (Mosasauridae: Plioplatecarpinae) from western Kansas, USA, reveals a novels suite of osteological characters for the genus. Canadian Journal of Earth Science 58 (9): 741 - 755. [https: // doi. org / 10.1139 / cjes- 2020 - 0175]." type="journal article" year="2021">Willman et al. (2021:747)</bibRefCitation> described the posterior surface of the splenials in @@ -674,16 +677,16 @@ and as possessing a "robust, dorsomedially expanding flange" giving an "overall square shape, in which they are almost equidimensional in height and width." Neither splenial of <collectionCode id="ED2E65BCFF966725FDFD166185A5FDA8" box="[593,650,512,538]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="15" pageNumber="14" type="Museum">YPM</collectionCode> VP4673 has a complete articulating surface, and the left is slightly crushed lateromedially, but both have curving margins and appear to have been more rounded than square ( -<figureCitation id="1304E1FCFF966725FFD516ED87E5FD14" box="[121,202,652,678]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." pageId="15" pageNumber="14">Fig. 9J</figureCitation> +<figureCitation id="1304E1FCFF966725FFD516ED87E5FD14" box="[121,202,652,678]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." figureDoi="http://doi.org/10.5281/zenodo.13888188" httpUri="https://zenodo.org/record/13888188/files/figure.png" pageId="15" pageNumber="14">Fig. 9J</figureCitation> ). Therefore, we do not consider a splenial with a squared articulating surface as diagnostic of the genus <emphasis id="B94B216BFF966725FD5616CF87D2FD5F" italics="true" pageId="15" pageNumber="14"> <taxonomicName id="4C3F86FAFF966725FD5616CF87D6FD5F" authorityName="Russell" authorityYear="1967" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="14" phylum="Chordata" rank="genus">Ectenosaurus</taxonomicName> . </emphasis> Only the anterior termini of the angulars are preserved ( -<figureCitation id="1304E1FCFF966725FF5016948653FCA2" box="[252,380,757,784]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." pageId="15" pageNumber="14">Fig. 9A‒H</figureCitation> +<figureCitation id="1304E1FCFF966725FF5016948653FCA2" box="[252,380,757,784]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." figureDoi="http://doi.org/10.5281/zenodo.13888188" httpUri="https://zenodo.org/record/13888188/files/figure.png" pageId="15" pageNumber="14">Fig. 9A‒H</figureCitation> ). While the left is less crushed, the right includes enough of the lateral wing that the surface for its contact with the surangular is preserved ( -<figureCitation id="1304E1FCFF966725FD16175D843CFCE4" box="[698,787,828,854]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." pageId="15" pageNumber="14">Fig. 9C</figureCitation> +<figureCitation id="1304E1FCFF966725FD16175D843CFCE4" box="[698,787,828,854]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." figureDoi="http://doi.org/10.5281/zenodo.13888188" httpUri="https://zenodo.org/record/13888188/files/figure.png" pageId="15" pageNumber="14">Fig. 9C</figureCitation> ). This surface is marked by pronounced horizontal striations and, unlike the condition present in <taxonomicName id="4C3F86FAFF966725FDC317E08435FC2F" baseAuthorityName="Merriam" baseAuthorityYear="1894" box="[623,794,897,925]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="14" phylum="Chordata" rank="species" species="clidastoides"> <emphasis id="B94B216BFF966725FDC317E08435FC2F" box="[623,794,897,925]" italics="true" pageId="15" pageNumber="14">E. clidastoides</emphasis> @@ -708,16 +711,16 @@ The splenio-articular joint in mosasaurs is usually described as a simple ball-a , for example, described it simply as "the splenial is the receiving part, the angular the received part." In <collectionCode id="ED2E65BCFF966725FED01082869AFB4F" box="[380,437,1251,1277]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="15" pageNumber="14" type="Museum">YPM</collectionCode> VP4673, the splenio-angular joint is more complex ( -<figureCitation id="1304E1FCFF966725FE22116486C8FA92" box="[398,487,1285,1312]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." pageId="15" pageNumber="14">Fig. 9L</figureCitation> +<figureCitation id="1304E1FCFF966725FE22116486C8FA92" box="[398,487,1285,1312]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." figureDoi="http://doi.org/10.5281/zenodo.13888188" httpUri="https://zenodo.org/record/13888188/files/figure.png" pageId="15" pageNumber="14">Fig. 9L</figureCitation> ), with each element contributing to the cotyle/condyle arrangement in a contact consisting of a series of more or less vertically oriented ridges and sulci. A deep median sulcus is present on the angular, for example, receiving a matching ridge on the splenial, while strong medial and lateral ridges have matching splenial sulci. While it is possible to deduce the general nature of the splenio-angular joint in <collectionCode id="ED2E65BCFF966725FD4B119D840FF9A4" box="[743,800,1532,1558]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="15" pageNumber="14" type="Museum">YPM</collectionCode> VP4673, deformation has altered the details of the articular faces of the angulars ( -<figureCitation id="1304E1FCFF966725FE3212238539F9EE" box="[414,534,1602,1628]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." pageId="15" pageNumber="14">Fig. 9B, G</figureCitation> +<figureCitation id="1304E1FCFF966725FE3212238539F9EE" box="[414,534,1602,1628]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." figureDoi="http://doi.org/10.5281/zenodo.13888188" httpUri="https://zenodo.org/record/13888188/files/figure.png" pageId="15" pageNumber="14">Fig. 9B, G</figureCitation> ). The elements do not match, and it is unclear which most closely approximates the original morphology. </paragraph> <paragraph id="8B80FD79FF966725FF3C12CD869AF8C4" blockId="15.[112,801,194,1946]" pageId="15" pageNumber="14"> An anteroventral rugosity on the right angular appears to be pathologic ( -<figureCitation id="1304E1FCFF966725FEEF12AE86B4F95B" box="[323,411,1743,1769]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." pageId="15" pageNumber="14">Fig. 9E</figureCitation> +<figureCitation id="1304E1FCFF966725FEEF12AE86B4F95B" box="[323,411,1743,1769]" captionStart="Figure 9" captionStartId="16.[112,183,1546,1568]" captionTargetBox="[161,1466,208,1518]" captionTargetId="figure-159@16.[141,1485,191,1531]" captionTargetPageId="16" captionText="Figure 9. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Left angular in lateral (A), medial (B), anterior (C), dorsal (D), and ventral (E) views. F–H. Right angular in lateral (F), anterior (G), and medial (H) views. I–K. Right splenial in lateral (I), anterior (J), and dorsal (K) view. L. Articulated right splenial and angular. Arrows point to pathologic ventral rugosity on left angular. Abbreviations: SAS, surangular articulation surface. Scale bars for A–E, I–K, L=5 cm. Scale bars for F–H=3 cm." figureDoi="http://doi.org/10.5281/zenodo.13888188" httpUri="https://zenodo.org/record/13888188/files/figure.png" pageId="15" pageNumber="14">Fig. 9E</figureCitation> ), as it is absent on the left angular. Considering the severe damage to the anterior end of the right dentary, it is possible this pathology resulted from the same trauma, produced by torsion inflicted on the intramandibular joint. </paragraph> <paragraph id="8B80FD79FF966725FF3C131F82DFFDAB" blockId="15.[112,801,194,1946]" lastBlockId="15.[832,1521,194,1945]" pageId="15" pageNumber="14"> @@ -735,13 +738,13 @@ VP4673 will be the focus of a future collaborative study between the senior auth Almost all the marginal teeth of YP VP4673 have broken free of the bone and been lost during post-exposure weathering and/or during collection of the specimen. The few teeth that remain <emphasis id="B94B216BFF966725FADD16EB8290FD14" box="[1393,1471,650,678]" italics="true" pageId="15" pageNumber="14">in situ</emphasis> are incomplete and missing their apices, though the apices can be observed for several replacement teeth. Possible pterygoid teeth are also present, but none are attached to the preserved fragments of the pterygoids. Preserved marginal teeth include the third tooth in the left dentary ( -<figureCitation id="1304E1FCFF966725FCE5173E84EBFCC8" box="[841,964,863,890]" captionStart="Figure 5" captionStartId="11.[112,183,1463,1485]" captionTargetBox="[178,1453,216,1431]" captionTargetId="figure-261@11.[128,1499,191,1445]" captionTargetPageId="11" captionText="Figure 5. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Dentaries in left lateral (A), left medial (B), left dorsal (C), right lateral (D), and right medial (E) views. Scale bars=10 cm." pageId="15" pageNumber="14">Fig. 5A, B</figureCitation> +<figureCitation id="1304E1FCFF966725FCE5173E84EBFCC8" box="[841,964,863,890]" captionStart="Figure 5" captionStartId="11.[112,183,1463,1485]" captionTargetBox="[178,1453,216,1431]" captionTargetId="figure-261@11.[128,1499,191,1445]" captionTargetPageId="11" captionText="Figure 5. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Dentaries in left lateral (A), left medial (B), left dorsal (C), right lateral (D), and right medial (E) views. Scale bars=10 cm." figureDoi="http://doi.org/10.5281/zenodo.13888176" httpUri="https://zenodo.org/record/13888176/files/figure.png" pageId="15" pageNumber="14">Fig. 5A, B</figureCitation> ), the third, fourth, fifth, and seventh teeth in the right dentary ( -<figureCitation id="1304E1FCFF966725FB8117E38383FC2F" box="[1069,1196,898,925]" captionStart="Figure 5" captionStartId="11.[112,183,1463,1485]" captionTargetBox="[178,1453,216,1431]" captionTargetId="figure-261@11.[128,1499,191,1445]" captionTargetPageId="11" captionText="Figure 5. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Dentaries in left lateral (A), left medial (B), left dorsal (C), right lateral (D), and right medial (E) views. Scale bars=10 cm." pageId="15" pageNumber="14">Fig. 5D, E</figureCitation> +<figureCitation id="1304E1FCFF966725FB8117E38383FC2F" box="[1069,1196,898,925]" captionStart="Figure 5" captionStartId="11.[112,183,1463,1485]" captionTargetBox="[178,1453,216,1431]" captionTargetId="figure-261@11.[128,1499,191,1445]" captionTargetPageId="11" captionText="Figure 5. Ectenosaurus tlemonectes sp. nov., YPM VP4673. A–E. Dentaries in left lateral (A), left medial (B), left dorsal (C), right lateral (D), and right medial (E) views. Scale bars=10 cm." figureDoi="http://doi.org/10.5281/zenodo.13888176" httpUri="https://zenodo.org/record/13888176/files/figure.png" pageId="15" pageNumber="14">Fig. 5D, E</figureCitation> ), the fifth right maxillary tooth (with replacement tooth) ( -<figureCitation id="1304E1FCFF966725FB4317C4825BFC72" box="[1263,1396,933,960]" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." pageId="15" pageNumber="14">Fig. 4D, E</figureCitation> +<figureCitation id="1304E1FCFF966725FB4317C4825BFC72" box="[1263,1396,933,960]" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." figureDoi="http://doi.org/10.5281/zenodo.13888174" httpUri="https://zenodo.org/record/13888174/files/figure.png" pageId="15" pageNumber="14">Fig. 4D, E</figureCitation> ), and the second left maxillary tooth (not figured). None of the four premaxillary teeth are preserved, but the bases indicate they may have been slightly prognathous ( -<figureCitation id="1304E1FCFF966725FA6D106E84ACFBFF" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." pageId="15" pageNumber="14">Fig. 4A, B</figureCitation> +<figureCitation id="1304E1FCFF966725FA6D106E84ACFBFF" captionStart="Figure 4" captionStartId="10.[112,183,1615,1637]" captionTargetBox="[149,1487,223,1589]" captionTargetId="figure-185@10.[117,1510,190,1605]" captionTargetPageId="10" captionText="Figure 4. Ectenosaurus tlemonectes sp. nov., YPM VP4673, A–C. Premaxilla in dorsal (A) and ventral (B) views. C. Detail of dorsal ridge. D, E. Right maxilla in lateral (D) and medial (E) views. Scale bars=10 cm." figureDoi="http://doi.org/10.5281/zenodo.13888174" httpUri="https://zenodo.org/record/13888174/files/figure.png" pageId="15" pageNumber="14">Fig. 4A, B</figureCitation> ). The bony tooth base is significantly taller in <collectionCode id="ED2E65BCFF966725FA1B105282DFFBFF" box="[1463,1520,1075,1101]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34880" lsid="urn:lsid:biocol.org:col:34880" name="Peabody Museum of Natural History" pageId="15" pageNumber="14" type="Museum">YPM</collectionCode> VP4673 than in @@ -817,7 +820,7 @@ VP4673 are posteromedially recurved, as in </emphasis> A number of loose teeth believed to have broken off the pterygoids are more elliptical in cross section, lack almost all ornamentation, but have strongly developed carinae. </paragraph> -<caption id="DF40ADF1FF89673AFFDC126B84AEF9DE" pageId="16" pageNumber="15" startId="16.[112,183,1546,1568]" targetBox="[161,1466,208,1518]" targetPageId="16" targetType="figure"> +<caption id="DF40ADF1FF89673AFFDC126B84AEF9DE" ID-DOI="http://doi.org/10.5281/zenodo.13888188" ID-Zenodo-Dep="13888188" httpUri="https://zenodo.org/record/13888188/files/figure.png" pageId="16" pageNumber="15" startId="16.[112,183,1546,1568]" targetBox="[161,1466,208,1518]" targetPageId="16" targetType="figure"> <paragraph id="8B80FD79FF89673AFFDC126B84AEF9DE" blockId="16.[112,1516,1545,1644]" pageId="16" pageNumber="15"> <emphasis id="B94B216BFF89673AFFDC126B87FFF9AD" bold="true" box="[112,208,1546,1568]" pageId="16" pageNumber="15">Figure 9.</emphasis> <taxonomicName id="4C3F86FAFF89673AFF7A126886F7F992" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[214,472,1545,1568]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="15" phylum="Chordata" rank="species" species="tlemonectes" status="sp. nov."> @@ -862,7 +865,7 @@ A number of loose teeth believed to have broken off the pterygoids are more elli <paragraph id="8B80FD79FF88673BFF3C152F851EFCC8" blockId="17.[112,800,193,890]" pageId="17" pageNumber="16"> <emphasis id="B94B216BFF88673BFF3C152F861BFED8" bold="true" box="[144,308,334,362]" italics="true" pageId="17" pageNumber="16">Vertebrae—</emphasis> Vertebrae are the sole preserved elements of the postcranial skeleton, though only a few cervical (including the axis) and dorsal vertebrae were collected and so the vertebral formula is unknown. The vertebrae that are present have all suffered varying degrees of plastic deformation. Where the cotylar and condylar surfaces are preserved, the interarcticular surfaces appear to have been broadly elliptical. The two cervicals have strongly developed, posteriorly-inclined hypapophyses. The hypapophyses are joined to the ventral rim of the condyle by a narrow, gently concave ridge ( -<figureCitation id="1304E1FCFF88673BFF6916B2860EFD5F" box="[197,289,723,749]" captionStart="Figure 10" captionStartId="17.[828,899,695,717]" captionTargetBox="[848,1509,219,666]" captionTargetId="figure-677@17.[831,1520,192,685]" captionTargetPageId="17" captionText="Figure 10. Ectenosaurus tlemonectes sp. nov., YPM VP4673. Cervical vertebra in left lateral view. Arrow indicates ventral ridge connecting the hypapophyseal peduncle and cotyle. Scale bar=5 cm." pageId="17" pageNumber="16">Fig. 10</figureCitation> +<figureCitation id="1304E1FCFF88673BFF6916B2860EFD5F" box="[197,289,723,749]" captionStart="Figure 10" captionStartId="17.[828,899,695,717]" captionTargetBox="[848,1509,219,666]" captionTargetId="figure-677@17.[831,1520,192,685]" captionTargetPageId="17" captionText="Figure 10. Ectenosaurus tlemonectes sp. nov., YPM VP4673. Cervical vertebra in left lateral view. Arrow indicates ventral ridge connecting the hypapophyseal peduncle and cotyle. Scale bar=5 cm." figureDoi="http://doi.org/10.5281/zenodo.13888190" httpUri="https://zenodo.org/record/13888190/files/figure.png" pageId="17" pageNumber="16">Fig. 10</figureCitation> ). <bibRefCitation id="EFAE8088FF88673BFE9716B2857EFD5F" author="Willman, A. J. & T. Konishi & M. W. Caldwell" box="[315,593,723,749]" pageId="17" pageNumber="16" pagination="741 - 755" refId="ref18000" refString="Willman, A. J., T. Konishi, and M. W. Caldwell. 2021. A new species of Ectenosaurus (Mosasauridae: Plioplatecarpinae) from western Kansas, USA, reveals a novels suite of osteological characters for the genus. Canadian Journal of Earth Science 58 (9): 741 - 755. [https: // doi. org / 10.1139 / cjes- 2020 - 0175]." type="journal article" year="2021">Willman et al. (2021)</bibRefCitation> stated that this ridge is unknown in all other plioplatecarpines except @@ -875,7 +878,7 @@ and </taxonomicName> . The hypapophyseal peduncles are slightly crushed but appear to have been elliptical in life. </paragraph> -<caption id="DF40ADF1FF88673BFC9016D682E4FCB2" pageId="17" pageNumber="16" startId="17.[828,899,695,717]" targetBox="[848,1509,219,666]" targetPageId="17" targetType="figure"> +<caption id="DF40ADF1FF88673BFC9016D682E4FCB2" ID-DOI="http://doi.org/10.5281/zenodo.13888190" ID-Zenodo-Dep="13888190" httpUri="https://zenodo.org/record/13888190/files/figure.png" pageId="17" pageNumber="16" startId="17.[828,899,695,717]" targetBox="[848,1509,219,666]" targetPageId="17" targetType="figure"> <paragraph id="8B80FD79FF88673BFC9016D682E4FCB2" blockId="17.[828,1500,695,768]" pageId="17" pageNumber="16"> <emphasis id="B94B216BFF88673BFC9016D68485FD7F" bold="true" box="[828,938,695,717]" pageId="17" pageNumber="16">Figure 10.</emphasis> <taxonomicName id="4C3F86FAFF88673BFC1D16D6839CFD7C" authority="Kiernan & Ebersole, 2023" authorityName="Kiernan & Ebersole" authorityYear="2023" box="[945,1203,695,718]" family="Mosasauridae" genus="Ectenosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="16" phylum="Chordata" rank="species" species="tlemonectes" status="sp. nov.">