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<document id="03E33B9626680E4D5CE9CFE834AEBDAA" ID-DOI="10.3897/zookeys.1219.130895" ID-ZooBank="2B55BA1C-2E86-4471-997B-624F508870A7" ID-publisher-id="130895" URI-arpha="FFAFBB0D-A51F-5116-A7BA-F8E127183D47" URI-zoobank="http://zoobank.org/2B55BA1C-2E86-4471-997B-624F508870A7" article-type="research-article" checkinTime="1733228135350" checkinUser="pensoft" docAuthor="Khalaji-Pirbalouty, Valiallah &amp; Al-Kandari, Manal Abdulrahman" docDate="2024" docId="FF97B31CFD9456E7BB8576D95DF7F205" docLanguage="en" docName="ZooKeys 1219: 233-248" docOrigin="ZooKeys 1219" docSource="https://zookeys.pensoft.net/article/130895/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Heterosignum behbehanii Khalaji-Pirbalouty &amp; Al-Kandari, 2024, sp. nov." docType="treatment" docVersion="1" dtd-version="3.0" lastPageNumber="248" masterDocId="FFAFBB0DA51F5116A7BAF8E127183D47" masterDocTitle="The first report of two new janiroid isopod species (Asellota, Janiroidea) from the western Indian Ocean" masterLastPageNumber="248" masterPageNumber="233" pageNumber="233" updateTime="1733228264115" updateUser="pensoft">
<mods:mods id="F89C055795DB2235B573C7C80637ABCC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8123A54A0F8FF968A35250C3BD3BD1AA">
<mods:title id="0B87CCE371FB9306A0B113959BA855DC">The first report of two new janiroid isopod species (Asellota, Janiroidea) from the western Indian Ocean</mods:title>
</mods:titleInfo>
<mods:name id="66B0B2AA120B5AF7358B8646161FBAB6" type="personal">
<mods:role id="15A78292B648F5242FB3132938509D65">
<mods:roleTerm id="A5618D338C30038FD47E440180695B76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="280E1D23DABC557515AFCACBCC379BF6">Khalaji-Pirbalouty, Valiallah</mods:namePart>
<mods:nameIdentifier id="903FBFF4BE8ADDAF9E944BBFD0166DA6" type="ORCID">0000-0002-0892-7463</mods:nameIdentifier>
<mods:affiliation id="E5653E8FAD3186FD8181DE70D4C7A7EA">Department of Biology, Faculty of Basic Science, Shahrekord University, Shahrekord, Iran</mods:affiliation>
</mods:name>
<mods:name id="327B7703D6A47CFB6905562B26BD806C" type="personal">
<mods:role id="9EDFC7EDBCB8C8DD739983F7B699D3A3">
<mods:roleTerm id="58A2C416822A313E4B9CCED9E7C372B0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6E572FB83C12495A0D8B13F6F63E542">Al-Kandari, Manal Abdulrahman</mods:namePart>
<mods:nameIdentifier id="81FB793DC75B1629E3EFE4B173337276" type="ORCID">0000-0003-0073-7929</mods:nameIdentifier>
<mods:affiliation id="FE6D224FAE161D87075F472CCCE7EFE2">Coastal and Marine Research Program, Environment and Life Sciences Research Center, Kuwait Institute of Scientific Research, Hamad Al-Mubarak Street, Building 900004, Area 1, Raas Salmiya, Kuwait</mods:affiliation>
</mods:name>
<mods:typeOfResource id="2F1D6AC51FCFD8DC1507CFC5741CCA3E">text</mods:typeOfResource>
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<mods:title id="7BCE8BE36154B3FBA9A2FBB852F8478F">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="69AE8485CA8E24BD762F9440811B349B">
<mods:date id="9E7C3A1538170B0DEA9B9E95A1466820">2024</mods:date>
<mods:detail id="BF076956665EF5C376FCBF5BF20A5B31" type="pubDate">
<mods:number id="3000034CACA11A5946A482C94157D5D4">2024-12-03</mods:number>
</mods:detail>
<mods:detail id="2198E971FE51C7A8688DF47697F84EA6" type="volume">
<mods:number id="2725276D375381BBD4C27E5E49A6EA5A">1219</mods:number>
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<mods:classification id="2B445D8971D59C6F35BD37758250FBB5">journal article</mods:classification>
<mods:identifier id="DB3FF98D8947E442F9F61EFE0659EDBA" type="DOI">10.3897/zookeys.1219.130895</mods:identifier>
<mods:identifier id="FF230B76B5BC5A16DE735D4AB5A3A218" type="ZooBank">2B55BA1C-2E86-4471-997B-624F508870A7</mods:identifier>
</mods:mods>
<treatment id="FF97B31CFD9456E7BB8576D95DF7F205" ID-arpha="FF97B31C-FD94-56E7-BB85-76D95DF7F205" ID-zoobank="https://zoobank.org/38B4B177-F22A-42D0-A139-61D9D21B8F38" LSID="urn:lsid:plazi:treatment:FF97B31CFD9456E7BB8576D95DF7F205" LSID-ZBK="urn:lsid:zoobank.org:act:38B4B177-F22A-42D0-A139-61D9D21B8F38" httpUri="http://treatment.plazi.org/id/FF97B31CFD9456E7BB8576D95DF7F205">
<subSubSection id="F54D5DD19012E8D8287D894B3794163C" type="nomenclature">
<paragraph id="3113EBB3BC12B16AFFAAE702C4CFC8A0">
<taxonomicName id="BF17DA283E638A20437F1CE305E0F1C8" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2024" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="behbehanii" status="sp. nov.">
<emphasis id="7B7651E82575A602ACE9D9AF21628E66" italics="true">Heterosignum behbehanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D1D304377E5A6A48A56A1C2AEC4688C9" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="311F3E14B14AEA6F8A79D65CF340B5D0" type="description">
<paragraph id="FAA36F009C9639FC181DE7F4600FAD48">
<figureCitation id="9C3E156581B14EB5A39196D203D8C374" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">Figs 1</figureCitation>
,
<figureCitation id="C8FD2C8CDE53FF47E481AF2CBEDA16EF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pereopod 1 B pereopod 2 C pereopod 5 D pereopod 7." figureDoi="10.3897/zookeys.1219.130895.figure2" httpUri="https://binary.pensoft.net/fig/1189295">2</figureCitation>
,
<figureCitation id="745817336410DCCA8D084D367AAE6DD8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pleopods 1 B pleopod 2 C pleopod 3 D penial processes E female operculum." figureDoi="10.3897/zookeys.1219.130895.figure3" httpUri="https://binary.pensoft.net/fig/1189296">3</figureCitation>
,
<figureCitation id="ECE4C5AB49DA5C90085A2B8AA44CBA28" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Heterosignum behbehanii sp. nov. Al-Khiran, Kuwait (28 ° 38.484 ' N, 48 ° 23.287 ' E) A male B female." figureDoi="10.3897/zookeys.1219.130895.figure4" httpUri="https://binary.pensoft.net/fig/1189297">4</figureCitation>
,
<figureCitation id="A8D369062D506CDBE44B2EF62F362943" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs A, B Heterosignum behbehanii sp. nov. (Al-Khiran, Kuwait: 28 ° 38.484 ' N, 48 ° 23.287 ' E) A female habitus B pleotelson, ventral view C Uromunna alyamanii sp. nov. (Al-Nuwaseeb, Kuwait: 28 ° 34.792 ' N, 48 ° 24.078 ' E), female pleotelson." figureDoi="10.3897/zookeys.1219.130895.figure5" httpUri="https://binary.pensoft.net/fig/1189298">5 A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXLAC" type="etymology">
<paragraph id="199B8DE6573EE80A3ECA34E498021B8C">
<heading id="28122573923C6B2FAAFDC17E6B540EFF" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="37F155E1E54876C25A6EDF491524DB19">
The species is named in honour of Dr Abdulmanaf Behbehani, a respected figure who taught Marine Biology and Ecology at
<collectingCountry id="0810664AC007900D5DD41B837E5625CF" name="Kuwait">Kuwait</collectingCountry>
University and researched Kuwait's marine environment and Kuwait's intertidal macrofauna for over four decades.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3LAC" type="type material">
<paragraph id="CC79CC43A806BD9E8B2450F9844C4B81">
<heading id="6C62FF6894F1996EAF66D135DB8965AC" reason="title">Type material.</heading>
</paragraph>
<paragraph id="980D32DCD957BAAE328968FA5D6FED6A">
<materialsCitation id="0B4396E1AA70A1CE596ABD166A95C4D3" collectingDate="2014-10-28" collectionCode="V" collectorName="Grintsov" country="Kuwait" latitude="29.360518" location="Al-Salam Beach" longLatPrecision="1" longitude="47.9534" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="411B37D8FFC46CEE3404BED71B5E31E7" bold="true" italics="true">
<typeStatus id="54A34EC8CD6F7296309D574B9217BE85" type="holotype">Holotype</typeStatus>
</emphasis>
: •
<specimenCount id="4982166DB41314F8A7DAE1A9F7B5ECAD" count="1" type="male">♂</specimenCount>
,
<quantity id="1F763C4B9893E139CE295E873269176C" metricMagnitude="-3" metricUnit="m" metricValue="1.12" unit="mm" value="1.12">1.12 mm</quantity>
(ZMH-K- 64934),
<collectingCountry id="C7A589B0BD7BA97F55F506401B2AB5D4" name="Kuwait">KUWAIT</collectingCountry>
,
<location id="9CA8E6234CB1C21ADD839009410BFF80" LSID="urn:lsid:plazi:treatment:FF97B31CFD9456E7BB8576D95DF7F205:9CA8E6234CB1C21ADD839009410BFF80" country="Kuwait" latitude="29.360518" longLatPrecision="1" longitude="47.9534" name="Al-Salam Beach">Al-Salam Beach</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKMAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[47.953400,29.360517]}">
<geoCoordinate id="49C0D5A41DE9F83ABB6F8378F39C0B95" degrees="29" direction="north" minutes="21.631" orientation="latitude" precision="1" value="29.360518">29 ° 21.631 ' N</geoCoordinate>
,
<geoCoordinate id="8715F4283D0644F1460FA7EBE57DAD20" degrees="47" direction="east" minutes="57.204" orientation="longitude" precision="1" value="47.9534">47 ° 57.204 ' E</geoCoordinate>
</named-content>
;
<date id="E3165603572E6A5DFFA6A406AF2E3D33" value="2014-10-28">
<collectingDate id="4D496FFDD4EF0244BE13A704B32D8036" value="2014-10-28">28. X. 2014</collectingDate>
</date>
;
<collectionCode id="C14DBF2B7EE5A7A974B5465A9449F388" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.
<collectorName id="DBFB67F4F612A57BEB9FA353C82E86B3">Grintsov</collectorName>
leg.
</materialsCitation>
<materialsCitation id="998F5E1B7F78E29168F01E0E29BDF263" collectingDate="2014-10-28" collectionCode="V" collectorName="Grintsov" country="Kuwait" latitude="29.360518" location="Al-Salam Beach" longLatPrecision="1" longitude="47.9534" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="CBE190B84830B49769E503D092E43343" bold="true" italics="true">
<typeStatus id="09B4406AF39AE3A3DC31F76A6C247A6E" type="paratype">Paratypes</typeStatus>
</emphasis>
: •
<specimenCount id="73B5710DD6AA14EFA4D0D6C842A6931F" count="1" type="female">1 ♀</specimenCount>
,
<quantity id="064563AEEF6FB91402D5056ED378324F" metricMagnitude="-3" metricUnit="m" metricValue="1.34" unit="mm" value="1.34">1.34 mm</quantity>
(ZMH-K- 64935), same data as the holotype
</materialsCitation>
•
<materialsCitation id="6719D1C5F2603DDDCC125B0369356B92" collectingDate="2014-11-11" collectionCode="V" collectorName="Grintsov" country="Kuwait" latitude="28.679634" location="Umm Al-Maradim Island" longLatPrecision="1" longitude="48.65345" specimenCount="5" specimenCount-female="4" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D492D6AA2458BE9D3D5745B6A1D8F9B1" count="1" type="male">1 ♂</specimenCount>
<quantity id="BF3E75AF9AA6A8BDE21CEB11B8D0D567" metricMagnitude="-3" metricUnit="m" metricValue="1.1" unit="mm" value="1.1">1.1 mm</quantity>
;
<specimenCount id="2D602D34C0B335FFD0C2D6537C1B32E4" count="4" type="female">4 ♀♀</specimenCount>
up to
<quantity id="03E06E2B5F0B807CB5C9F19ED6FFA1F7" metricMagnitude="-3" metricUnit="m" metricValue="1.25" unit="mm" value="1.25">1.25 mm</quantity>
(ZMH-K- 64936),
<location id="8698C016C91EBC9A9E00CD36E9086ED2" LSID="urn:lsid:plazi:treatment:FF97B31CFD9456E7BB8576D95DF7F205:8698C016C91EBC9A9E00CD36E9086ED2" country="Kuwait" latitude="28.679634" longLatPrecision="1" longitude="48.65345" name="Umm Al-Maradim Island">Umm Al-Maradim Island</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EVMAC" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[48.653450,28.679633]}">
<geoCoordinate id="DCA8AC5964CC3E524F81CADDD546C3C9" degrees="28" direction="north" minutes="40.778" orientation="latitude" precision="1" value="28.679634">28 ° 40.778 ' N</geoCoordinate>
,
<geoCoordinate id="8FDCD4ABABE3438E80F1486F0C9254E4" degrees="48" direction="east" minutes="39.207" orientation="longitude" precision="1" value="48.65345">48 ° 39.207 ' E</geoCoordinate>
</named-content>
;
<date id="1B83773BF8DE3793F56914D17D76D116" value="2014-11-11">
<collectingDate id="27675F52D0BE12091612926AE59C80D3" value="2014-11-11">11. XI. 2014</collectingDate>
</date>
;
<collectionCode id="0BAF9C9551EEDC38F0DE24B82B46D248" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
.
<collectorName id="F1F1655226E6199FA9D8803C1ECA0B28">Grintsov</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1MAC" type="diagnosis">
<paragraph id="34735E5143F6D16999CEC54A9CC404DF">
<heading id="6EB7C7CEC0582669B55865A9A3D4CDAA" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="B6AACD0C5885EF42266724C0F9ABFF99">Eyes with 3 ommatidia; eyestalks with medium length, reaching proximal half of peduncular article 2 of antenna 2. Pereonites 2–7 with a pair of relatively short single lateral spine-like processes. The pleotelson has a relatively short anterior neck, length about 0.1 pleotelson length, lateral margins with 6 denticles each. Male pleopod 2 with an elongate, downwardly curved and distally pointed apex endopodal stylet.</paragraph>
</subSubSection>
<subSubSection id="SECID0E6MAC" type="description">
<paragraph id="8E1A3A5F864F99DF0768A113B41DBD09">
<heading id="740018A596934D43A585B01C38A3AB0D" reason="title">Description.</heading>
</paragraph>
<paragraph id="0AB442C7450920044687AFF46440DD54">
<emphasis id="5C434096F0B0C89183E35BDFC3C3DF44" bold="true">Male</emphasis>
: Body (Fig.
<figureCitation id="57C04B50095188E7E5032CEE98C85B4D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 B</figureCitation>
) length
<quantity id="D11D2BB1B6A5A07D501B5866BE25B11A" metricMagnitude="-3" metricUnit="m" metricValue="1.28" unit="mm" value="1.28">1.28 mm</quantity>
; about 2 times as long as wide, widest at pereonite 3.
</paragraph>
<caption id="A217C6FB8D18C16268D9E55655CBC97E" ID-DOI="10.3897/zookeys.1219.130895.figure1" ID-arpha="63B06437-8882-5EBF-B6FC-862FFCE48FCA" httpUri="https://binary.pensoft.net/fig/1189294" startId="F1">
<paragraph id="0BE461E9BCEB8D345FEDD0CA6DF3CDCF">
<label id="CFC7CA2D89C35619EA45DD4C4C36A827">Figure 1.</label>
</paragraph>
<paragraph id="AE082EAABA32DB4D12FADBAA21146BE8">
<taxonomicName id="99BF850E52EB480447F1826942783166" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2024" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="behbehanii" status="sp. nov.">
<emphasis id="D7DCAB2D8805C0E3A0C87682F277E295" italics="true">Heterosignum behbehanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0EB8235D473209D8B21B9C88627E9BDE" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="C386B9BAA60EC7C71BD0A8E7A40FFF87" bold="true">A</emphasis>
female habitus
<emphasis id="7AF5FCD0B4E28FFE2FAC0DF4860A984F" bold="true">B – J</emphasis>
male holotype (ZMH-K- 64934)
<emphasis id="6B5C9D469D916C09F091004DFFF9D3F3" bold="true">B</emphasis>
habitus
<emphasis id="18C66D6E81DCC84318F41D85634FE3FD" bold="true">C</emphasis>
antennula
<emphasis id="8A090C510E275D948BF4F19D4C426123" bold="true">D</emphasis>
antenna
<emphasis id="5398B27C7B67962A4961EECDFD79CD9D" bold="true">E</emphasis>
mandible
<emphasis id="E4C2CBEFCFB0827166567D7B74B3052B" bold="true">F</emphasis>
maxillula
<emphasis id="F3605B1FA7E51DACC2C20AF6FFAB74BF" bold="true">G</emphasis>
maxilla
<emphasis id="49D2B5861557D3E9C90441096E3A8169" bold="true">H</emphasis>
maxilliped
<emphasis id="93B8A9CBC73727FC507CFBC85AEB57F4" bold="true">I</emphasis>
uropod
<emphasis id="A7A9D7228A7197A61B97901732693E5F" bold="true">J</emphasis>
pleotelson lateral margin, ventral view.
</paragraph>
</caption>
<paragraph id="4079044CBD780C5653BBFD5D4026E228">
<emphasis id="89A47071D9F8BE2CC1D2BC9521B3FDED" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="8F7C531F55678B992347EE30D5172C92" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 B</figureCitation>
) 0.36 times as long as wide (including eyestalks); frontal margin convex, having pair of weak preocular lobes.
<emphasis id="6A74791FFB6D3D0AB562F1A86DB7A12F" bold="true" italics="true">Eyestalks</emphasis>
(Figs
<figureCitation id="C46B97176215CA683A7153F12009D0CE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 B</figureCitation>
,
<figureCitation id="1CA7BE7F3EDEFD88BDB98615EA6F4650" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Heterosignum behbehanii sp. nov. Al-Khiran, Kuwait (28 ° 38.484 ' N, 48 ° 23.287 ' E) A male B female." figureDoi="10.3897/zookeys.1219.130895.figure4" httpUri="https://binary.pensoft.net/fig/1189297">4 A</figureCitation>
) arising from posterior half of head, reaching proximal half of peduncular article 2 of antenna 2, 0.62 length of head, eyes with 3 ommatidia.
</paragraph>
<paragraph id="6557A814D1FDDB5281D447532C4A4EC0">
<emphasis id="87895D4CDFD0BA65AA9400C7E79FDFB3" bold="true" italics="true">Pereon</emphasis>
(Figs
<figureCitation id="6563B3323F8BAA8D9AE8CCF737F80A2D" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 B</figureCitation>
,
<figureCitation id="5166B7DC5DE13A3123C0D79EADBBEA4C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Heterosignum behbehanii sp. nov. Al-Khiran, Kuwait (28 ° 38.484 ' N, 48 ° 23.287 ' E) A male B female." figureDoi="10.3897/zookeys.1219.130895.figure4" httpUri="https://binary.pensoft.net/fig/1189297">4 A</figureCitation>
,
<figureCitation id="A2AF415AA3B788327AD2BD775B999EB2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs A, B Heterosignum behbehanii sp. nov. (Al-Khiran, Kuwait: 28 ° 38.484 ' N, 48 ° 23.287 ' E) A female habitus B pleotelson, ventral view C Uromunna alyamanii sp. nov. (Al-Nuwaseeb, Kuwait: 28 ° 34.792 ' N, 48 ° 24.078 ' E), female pleotelson." figureDoi="10.3897/zookeys.1219.130895.figure5" httpUri="https://binary.pensoft.net/fig/1189298">5 A</figureCitation>
) dorsal surface smooth;
<emphasis id="E2F5BBB888DD74894C7B15C6DC46F302" bold="true" italics="true">pereonite 1</emphasis>
laterally rounded, without lateral spine-like processes;
<emphasis id="062797E88620523818F063A065B530DA" bold="true" italics="true">pereonites</emphasis>
2–7 with a pair of short single lateral spine-like processes;
<emphasis id="59302867CAAC755BAC91C033877B15D3" bold="true" italics="true">pereonites</emphasis>
1–3 increasing in width and
<emphasis id="A39F294BC783EACF07A0514EAD02E874" bold="true" italics="true">pereonites</emphasis>
4–7 decreasing in width.
<emphasis id="D6BF47108AF3F57B9C0AD3F72C0C69D5" bold="true" italics="true">Pleonite 1</emphasis>
small, 0.26 times as long as wide, with pair of fine dorsal setae.
</paragraph>
<paragraph id="AB6DFB9DD6426EE205E7CF7E11B0283A">
<emphasis id="EE70A09CC9A291F78D318CB2E94BFFCE" bold="true" italics="true">Pleotelson</emphasis>
(Fig.
<figureCitation id="512765F70A96069223BB2514CF0BEA0B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 B, J</figureCitation>
) about 1.3 times longer than wide, anterior part narrow, about 0.13 times of total length of pleotelson, posterior part bulbous, tapering posteriorly, lateral margins each with 6 denticles, posterior margin rounded, ventral margins with a raw of small submarginal simple setae.
<emphasis id="38EAE6791FDC3DFA8E3DFAF48BA1CE93" bold="true" italics="true">Uropod</emphasis>
(Fig.
<figureCitation id="7A3BA11A07F00ACFEF6ED4FF3BAEE2CC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 I</figureCitation>
) cylindrical, endopod with 2 lateral and 6 apical setae; exopod about 0.3 length of endopod, with 3 apical setae.
</paragraph>
<paragraph id="C0820C343EC07528745BD664E1BCF52A">
<emphasis id="ED52E3557E0E47629958FCEE9EF64130" bold="true" italics="true">Antennula</emphasis>
(Fig.
<figureCitation id="31F48C7DBD707DCA04C7EF5A2E373B35" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 C</figureCitation>
) peduncular article 1 shorter than article 2; article 2 with 1 simple and 3 small sensory palmate setae distodorsally; article 3 about 0.5 length of article 2, with 1 simple seta distally; flagellum with three articles, article 1 shortest, with 1 distoventral seta; article 3 subequal to article 2, distally with 1 long, 2 short setae and 1 aesthetasc.
</paragraph>
<paragraph id="C959DFFB9C536DF904D95978B5A709FB">
<emphasis id="BA15C9A823B9250B91DD2E889802CD4D" bold="true" italics="true">Antenna</emphasis>
(Fig.
<figureCitation id="B1241BE9AA00CDA82806A8FF063D8ABB" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 D</figureCitation>
) with peduncle articles 1 and 2 combined about 0.8 length of article 3; peduncle article 3 with convex proximal protrusion bearing single seta, with 1 simple seta medially and 1 simple seta distally; article 4 about half length of article 3, with 2 distolateral setae.
</paragraph>
<paragraph id="3C365950B85B2D4B828DCADE643974C7">
<emphasis id="7FA54D5181396C0332D2BC1C1C2376CA" bold="true" italics="true">Mandible</emphasis>
(Fig.
<figureCitation id="65EE7386A411590004414DF74ED557FF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 E</figureCitation>
) incisor process with 5 and
<emphasis id="F2529D53830A711CFBC4094E8B301175" bold="true" italics="true">lacinia mobilis</emphasis>
with 4 cusps; spine row with 4 robust serrate setae.
</paragraph>
<paragraph id="CD11D83A3B3D5C4823DECE6E358DF29D">
<emphasis id="0A3887EE661EA0E546DC8C8EBCBB33EB" bold="true" italics="true">Maxillula</emphasis>
(Fig.
<figureCitation id="B72CF813E09FB165060F179FD6ED0DB5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 F</figureCitation>
): lateral lobe with 9 robust apical setae; medial lobe with 4 robust serrate setae on distal margin.
</paragraph>
<paragraph id="E79A030D5064AD3E0BA1781CC6603250">
<emphasis id="290AFA38E0012B674F68C5986B10EDEC" bold="true" italics="true">Maxilla</emphasis>
(Fig.
<figureCitation id="875A16C292F01DBEA982BDA204AB9F89" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 G</figureCitation>
) lateral and mesial lobes, each with 4 pectinate robust apical setae; medial lobe with 9 robust setae along distal margin, some of them serrated.
</paragraph>
<paragraph id="1C9967656A51B11339351B1994D7CEF7">
<emphasis id="EE8C707EE9BE1BED7538893D855D07B0" bold="true" italics="true">Maxilliped</emphasis>
(Fig.
<figureCitation id="85200356365DE63260DBC3F00F916F14" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 H</figureCitation>
) endite distal margin with 1–2 fan-shaped setae, 3 serrate setae, and 1 sub marginal seta medially, mesial margin with 2 coupling hooks; palp article 1 with 1 short seta laterally; article 2 shorter than article 3 with 2 setae medially; article 3 with 3 setae medially and 1 seta laterally; article 4 subequal in length to article 3, with 3 setae distomedially; article 5 narrowest, with 5 setae distally.
</paragraph>
<paragraph id="38127FA8BAA1546CD75691274547B7BC">
<emphasis id="2523AFA10B6AD8C3C2FC66B812AEB065" bold="true" italics="true">Pereopod 1</emphasis>
(Fig.
<figureCitation id="6553DA093B76E66AE8734C6A10BDC8DE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pereopod 1 B pereopod 2 C pereopod 5 D pereopod 7." figureDoi="10.3897/zookeys.1219.130895.figure2" httpUri="https://binary.pensoft.net/fig/1189295">2 A</figureCitation>
) shorter than
<emphasis id="F349BA79F8736DC3276A7BED2240EA2E" bold="true">
<emphasis id="2E0CBFB59AF39A0D91F935F345BA7501" bold="true" italics="true">pereopods 2</emphasis>
–
<emphasis id="6B8D6306E2CF12AD0F16C35BD7B38394" bold="true" italics="true">7</emphasis>
</emphasis>
: basis 3.6 times as long as greatest width, superior margin with 1 and inferior margin with 2 small setae; ischium 0.5 times as long as basis, with 2 long setae distally; merus shorter than carpus, superior margin with 1 robust seta medially, with 2 robust setae on inferodistal corner; carpus inferior margin with 4 denticles, 3 robust setae and 1 submarginal simple seta, superior margin with 1 simple seta on distal corner; propodus longer than carpus, with 3 robust setae on inferior margin; dactylus with 1 subapical simple seta, unguis subequal in length to dactylus, accessory claw length 0.6 unguis length.
</paragraph>
<caption id="3FECCDE8F6EF2A4960436E92AC3E6C95" ID-DOI="10.3897/zookeys.1219.130895.figure2" ID-arpha="13D09B6D-E577-5C78-933F-08DF07B462D4" httpUri="https://binary.pensoft.net/fig/1189295" startId="F2">
<paragraph id="FCCEF236EB73F680F67F25F2E9D4C373">
<label id="1BFA3FD0E5A26CB52B2BD5BBFE666C71">Figure 2.</label>
</paragraph>
<paragraph id="B602C7258428E911D7276FA1F9BAA71E">
<taxonomicName id="AEE7E00376355A8193F55CBE6604964C" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2024" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="behbehanii" status="sp. nov.">
<emphasis id="9C996FEE652636A2A33BDCEA95EB63DF" italics="true">Heterosignum behbehanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00AE9172007BBDD9BB4560CC15C2A4D7" rank="species">sp. nov.</taxonomicNameLabel>
(ZMH-K- 64934), male holotype
<emphasis id="88107A50A13EEF3CE5872B6C48B9DA4C" bold="true">A</emphasis>
pereopod 1
<emphasis id="4432E95731466FFF439DBFC14E742449" bold="true">B</emphasis>
pereopod 2
<emphasis id="52E6636478DC838B28250368057DCE1D" bold="true">C</emphasis>
pereopod 5
<emphasis id="9613A8AAEE5C41E322C449FCDAEB9198" bold="true">D</emphasis>
pereopod 7.
</paragraph>
</caption>
<paragraph id="C117D31D2B70F7879A9588774612575D">
<emphasis id="2F0939FE53064F8F8BDC700E0B8B3207" bold="true" italics="true">Pereopod 2</emphasis>
(Fig.
<figureCitation id="FAC490B75F27F058DFC39C5142E35C90" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pereopod 1 B pereopod 2 C pereopod 5 D pereopod 7." figureDoi="10.3897/zookeys.1219.130895.figure2" httpUri="https://binary.pensoft.net/fig/1189295">2 B</figureCitation>
) basis about 3.5 times as long as greatest width; ischium about 0.9 times as long as basis, with 1 long seta on inferior margin; merus about 0.5 times as long as carpus, inferior margin with 2 and superior margin with 1 long simple setae; carpus and propodus subequal in length, carpus inferior margin with 3 robust setae, superior margin with 1 seta medially and 2 simple long setae on distal corner; propodus inferior margin with 3 robust setae and 1 long simple seta, superior margin with 3 long simple setae; dactylus with 2 subapical simple setae, a small accessory claw, unguis 1.5 times as long as dactylus.
</paragraph>
<paragraph id="7FC8A355AA52E296FA26B4A07F3D2B91">
<emphasis id="0C160258007F2C24D6F5A6652150E661" bold="true" italics="true">Pereopod 5</emphasis>
(Fig.
<figureCitation id="9FCA5F72F5E6F90A6855EFF387563E90" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pereopod 1 B pereopod 2 C pereopod 5 D pereopod 7." figureDoi="10.3897/zookeys.1219.130895.figure2" httpUri="https://binary.pensoft.net/fig/1189295">2 C</figureCitation>
) basis about 3.8 times as long as greatest width; ischium about 0.92 times as long as basis, with 1 long seta on inferior margin and 2 long setae on superior margin; merus about 0.46 times as long as carpus, inferior and superior margin with 2 simple setae each; carpus shorter than propodus, with 2 robust setae on inferior margin and 1 simple seta on superior margin medially; propodus inferior margin with 3 robust setae, superior margin with 3 simple setae; dactylus with 2 subapical simple setae; unguis 1.4 times as long as dactylus.
</paragraph>
<paragraph id="32F7356A9C2B171D818D157B53579124">
<emphasis id="09EBC941F54AFE73F6D952F25002E664" bold="true" italics="true">Pereopod 7</emphasis>
(Fig.
<figureCitation id="0AE07046C4403C3A8C2980C5B9379D52" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pereopod 1 B pereopod 2 C pereopod 5 D pereopod 7." figureDoi="10.3897/zookeys.1219.130895.figure2" httpUri="https://binary.pensoft.net/fig/1189295">2 D</figureCitation>
) basis about 3.8 times as long as greatest width; ischium about 0.85 times as long as basis, with 1 long seta on inferior margin; merus about 0.47 times as long as carpus, inferior and superior margins with 1 long simple seta; carpus shorter than propodus, with 2 robust setae on inferior margin; propodus inferior margin with 2 robust setae, superior margin with 3 simple setae; unguis 1.5 times as long as dactylus.
</paragraph>
<paragraph id="FB9E52E08C155EBFA37BE41F60E0E9DD">
<emphasis id="AFA732FDD3E28956DE3C879938A8D887" bold="true" italics="true">Pleopod 1</emphasis>
(Fig.
<figureCitation id="F9D012685F2C95268799AB800BDE4E64" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pleopods 1 B pleopod 2 C pleopod 3 D penial processes E female operculum." figureDoi="10.3897/zookeys.1219.130895.figure3" httpUri="https://binary.pensoft.net/fig/1189296">3 A</figureCitation>
) distally separate, with 2–4 apical setae; short lateral projections present, each projection with 1–2 setae.
</paragraph>
<caption id="92483E027A2734731EB96EBA1C7F22B8" ID-DOI="10.3897/zookeys.1219.130895.figure3" ID-arpha="4EBD40FC-000B-5986-8583-9708CB1B49AC" httpUri="https://binary.pensoft.net/fig/1189296" startId="F3">
<paragraph id="91134F866E9ED34DB2D496E76742CF13">
<label id="494BA72DDFEBD89017EF2F471181A321">Figure 3.</label>
</paragraph>
<paragraph id="FBE2F2552746D670BACAE0F1749C508A">
<taxonomicName id="6875381306D93A828BE7559048EAA9AB" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2024" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="behbehanii" status="sp. nov.">
<emphasis id="98B3949EF8197C4F24BEA9ED80070F05" italics="true">Heterosignum behbehanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7B89E157B883A96569AA7FA7CC25301B" rank="species">sp. nov.</taxonomicNameLabel>
(ZMH-K- 64934), male holotype
<emphasis id="DA01F3FE86146DF038F94A1F080CBE7F" bold="true">A</emphasis>
pleopods 1
<emphasis id="58FA64684C5F2BA7AFD9E0B4A2F73A32" bold="true">B</emphasis>
pleopod 2
<emphasis id="7405B53FE8FE8E8BE458210E68E9A169" bold="true">C</emphasis>
pleopod 3
<emphasis id="5CA04FFD55F663A6668897ADF6A4DF21" bold="true">D</emphasis>
penial processes
<emphasis id="71EE19D79B77A27B3895E742B9ABCAA2" bold="true">E</emphasis>
female operculum.
</paragraph>
</caption>
<paragraph id="9BE96CF416FC6999DD95D59CC3959F19">
<emphasis id="5DF09540641C242008D8C03D2246F190" bold="true" italics="true">Pleopod 2</emphasis>
(Fig.
<figureCitation id="87DF320D301E6E335886A36CAC5D4944" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pleopods 1 B pleopod 2 C pleopod 3 D penial processes E female operculum." figureDoi="10.3897/zookeys.1219.130895.figure3" httpUri="https://binary.pensoft.net/fig/1189296">3 B</figureCitation>
) protopod length 2.4 width, with 16 simple marginal setae; second article of endopod elongate, downwardly curved, proximally swollen, becoming narrower until distal pointed apex, about 2.4 as long as protopod, without any ornamentation.
</paragraph>
<paragraph id="AD6879AAF1861596322F71B8D3A1BAA2">
<emphasis id="51BB690C5ED9C7461F3C4D1301E36178" bold="true" italics="true">Pleopod 3</emphasis>
(Fig.
<figureCitation id="42F87D8936786FC6EF350FD4FD32CD8B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pleopods 1 B pleopod 2 C pleopod 3 D penial processes E female operculum." figureDoi="10.3897/zookeys.1219.130895.figure3" httpUri="https://binary.pensoft.net/fig/1189296">3 C</figureCitation>
), endopod with 2 long plumose setae apically and 1 long plumose seta laterally; exopod composed of two articles, narrower than endopod; article 2 with 1 terminal seta.
</paragraph>
<paragraph id="3D1C0CE2D2B0FB344DFB280A1A05D623">
<emphasis id="54B4FA3BCAF50140CD5DA0A5EDCDE189" bold="true" italics="true">Penes</emphasis>
(Fig.
<figureCitation id="30474517A070A6738F98CE8EF71B7B91" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pleopods 1 B pleopod 2 C pleopod 3 D penial processes E female operculum." figureDoi="10.3897/zookeys.1219.130895.figure3" httpUri="https://binary.pensoft.net/fig/1189296">3 D</figureCitation>
) fused along basal part, well separated at distal part, tapering to narrowly rounded apices.
</paragraph>
<paragraph id="2669D4C2763EF25EC3DD2BE25DADBE00">
<emphasis id="042D839662B7E74A80BBA2ADC9503A19" bold="true">Female</emphasis>
(Figs
<figureCitation id="A6C7209D240C11AEF4F162640B1D66F4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Heterosignum behbehanii sp. nov. A female habitus B – J male holotype (ZMH-K- 64934) B habitus C antennula D antenna E mandible F maxillula G maxilla H maxilliped I uropod J pleotelson lateral margin, ventral view." figureDoi="10.3897/zookeys.1219.130895.figure1" httpUri="https://binary.pensoft.net/fig/1189294">1 A</figureCitation>
,
<figureCitation id="8D20212632D1B58694D3422AEAB85791" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Heterosignum behbehanii sp. nov. Al-Khiran, Kuwait (28 ° 38.484 ' N, 48 ° 23.287 ' E) A male B female." figureDoi="10.3897/zookeys.1219.130895.figure4" httpUri="https://binary.pensoft.net/fig/1189297">4 B</figureCitation>
,
<figureCitation id="EE3530F770806BD4AF7316932F943D5F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Scanning electron micrographs A, B Heterosignum behbehanii sp. nov. (Al-Khiran, Kuwait: 28 ° 38.484 ' N, 48 ° 23.287 ' E) A female habitus B pleotelson, ventral view C Uromunna alyamanii sp. nov. (Al-Nuwaseeb, Kuwait: 28 ° 34.792 ' N, 48 ° 24.078 ' E), female pleotelson." figureDoi="10.3897/zookeys.1219.130895.figure5" httpUri="https://binary.pensoft.net/fig/1189298">5 A, B</figureCitation>
). Similar to males in the morphology of all pereonal appendages. Body broader than in males, about 1.8 times as long as wide; widest at pereonite 3. The largest female length is
<quantity id="945EFD4723DAB5E36DFF0098F45E2E88" metricMagnitude="-3" metricUnit="m" metricValue="1.34" unit="mm" value="1.34">1.34 mm</quantity>
.
<emphasis id="2C1DD8337CFC26EC69B10FF29D627127" bold="true" italics="true">Operculum</emphasis>
(Fig.
<figureCitation id="0D4C0A979BBD869A1561F46CF699AB06" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Heterosignum behbehanii sp. nov. (ZMH-K- 64934), male holotype A pleopods 1 B pleopod 2 C pleopod 3 D penial processes E female operculum." figureDoi="10.3897/zookeys.1219.130895.figure3" httpUri="https://binary.pensoft.net/fig/1189296">3 E</figureCitation>
) ovate, 1.2 times as long as broad, without any marginal setae.
</paragraph>
<caption id="A56715F3B27748223486D0CE81104284" ID-DOI="10.3897/zookeys.1219.130895.figure4" ID-arpha="7EB354A0-231C-5A02-A817-62B4C3132BF7" httpUri="https://binary.pensoft.net/fig/1189297" startId="F4">
<paragraph id="291B8E676457257D4C9C4BF91104ACF0">
<label id="DF53B6C2EF32936387D539A1FD291DBF">Figure 4.</label>
</paragraph>
<paragraph id="48FF0B96927CFD0520A9B8F4415E22E7">
<taxonomicName id="775A26E6FC4A736CAF619D3888FCB530" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2024" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="behbehanii" status="sp. nov.">
<emphasis id="155722D2B056693268A5D810F5E8EDC9" italics="true">Heterosignum behbehanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3E881CAF33821806F8724D9A0DB08B85" rank="species">sp. nov.</taxonomicNameLabel>
Al-Khiran, Kuwait (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1JAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[48.388117,28.641400]}">28 ° 38.484 ' N, 48 ° 23.287 ' E</named-content>
)
<emphasis id="F0C3E8FC6BE0F864581E4E2DCE60B988" bold="true">A</emphasis>
male
<emphasis id="97A96EF1666E32449048A5F1EB7CE707" bold="true">B</emphasis>
female.
</paragraph>
</caption>
<caption id="87FB873D2064775A3899D4D7B1EAF721" ID-DOI="10.3897/zookeys.1219.130895.figure5" ID-arpha="FD521A0F-A2B2-54ED-B795-7EBFFA008182" httpUri="https://binary.pensoft.net/fig/1189298" startId="F5">
<paragraph id="3C60E0694553097E53830F313C168276">
<label id="CFED8FC1B093B65BE2C3564A67518DD6">Figure 5.</label>
</paragraph>
<paragraph id="17419B3AAF47AF022E9A2154180A33AC">
Scanning electron micrographs
<emphasis id="5684C72B9774DE23E77D90B2465098CE" bold="true">A, B</emphasis>
<taxonomicName id="7B97DB59E405311DCB9B969702BECD1C" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2024" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="behbehanii" status="sp. nov.">
<emphasis id="F8BC062575CAD3AC5136A5EE0C0CF0D0" italics="true">Heterosignum behbehanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F22F9B5BE504A09F042A15C8328A1FC6" rank="species">sp. nov.</taxonomicNameLabel>
(Al-Khiran, Kuwait:
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EALAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[48.388117,28.641400]}">28 ° 38.484 ' N, 48 ° 23.287 ' E</named-content>
)
<emphasis id="CD4EE4F9053B6000FB2F98B5975A6B31" bold="true">A</emphasis>
female habitus
<emphasis id="F83AE181816FA82851FD78430792E17B" bold="true">B</emphasis>
pleotelson, ventral view
<emphasis id="BDB28B324623AFD13030FB9FE14272C0" bold="true">C</emphasis>
<taxonomicName id="4842FE3028C2917E17EA3EC8AF24C911" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2024" class="Malacostraca" family="Munnidae" genus="Uromunna" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="alyamanii" status="sp. nov.">
<emphasis id="76E4B9FEBDB607DAB0DE1DA3B32B64F0" italics="true">Uromunna alyamanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="232FF66B08033BDE0327AD8543D390BB" rank="species">sp. nov.</taxonomicNameLabel>
(Al-Nuwaseeb, Kuwait:
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EZLAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[48.401300,28.579867]}">28 ° 34.792 ' N, 48 ° 24.078 ' E</named-content>
), female pleotelson.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EDMAE" type="remarks">
<paragraph id="79019AF689BDC2E50133E02A47D019AF">
<heading id="3D6274C0EA75F665F3F639BE2E478E2C" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="8BE7A057683C1D97F3C2A212F1F20294">
<taxonomicName id="4EF28077E3F8DF98031ABFC61C096241" authorityName="Khalaji-Pirbalouty &amp; Al-Kandari" authorityYear="2024" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="behbehanii" status="sp. nov.">
<emphasis id="20883312E0B787A3C4FE5680E7060EDF" italics="true">Heterosignum behbehanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B788280413BB216647CF7149CEE6FCD4" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="24E50123CD6AFE95420801B8672D6127" authority="Shimomura &amp; Mawatari, 2002" authorityName="Shimomura &amp; Mawatari" authorityYear="2002" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="915DF4ED20BD439B3834199994A2B87E" italics="true">H. elegans</emphasis>
<bibRefCitation id="2D8A77768CD251735797C1A512564A63" DOI="10.1080/00222930110088734" author="Shimomura M &amp; Mawatari SF" journalOrPublisher="Journal of Natural History" pagination="2103-2126" refId="B14" refString="Shimomura M, Mawatari SF (2002) A revision of Heterosignum (Asellota, Isopoda, Paramunnidae) with descriptions of three new species. Journal of Natural History 36: 2103 - 2126. DOI: https: // doi. org / 10.1080 / 00222930110088734" title="A revision of Heterosignum (Asellota, Isopoda, Paramunnidae) with descriptions of three new species." volume="36" year="2002">Shimomura &amp; Mawatari, 2002</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F142CB56C129650FE394353001E7BBC0" name="Japan">Japan</collectingCountry>
, as both of these species have lateral spine-like processes on pereonites 2–7, a pleotelson with 6 denticles on its lateral margins of the posterior part, and pleopod 2 with a slender very long, curved second article of the endopod. However, the new species is distinguished from
<taxonomicName id="74F11EFB5EA1E9E4E835FDA6FA1AC4CC" authorityName="Shimomura &amp; Mawatari" authorityYear="2002" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="A42DCC999B7DA7BD94C5D445FD2A5BEE" italics="true">H. elegans</emphasis>
</taxonomicName>
in having its notably shorter lateral spine-like processes on pereonites 2–7, a smaller anterior part of the pleotelson (about 0.1 vs. 0.4 times the total length of pleotelson), and the first pleopod being distally separate instead of distally connected.
</paragraph>
<paragraph id="DD7FD864F500CB13C8E6B92B81B91650">
The new species differs from
<taxonomicName id="13232BA65D23291F66A33E88AA1CDD1D" authority="(Kensley, 1976)" baseAuthorityName="Kensley" baseAuthorityYear="1976" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="208DAE4C8E71A1A3E6135EB61A8072E1" italics="true">Heterosignum unicornis</emphasis>
(
<bibRefCitation id="8C14003A7AF5F1D43E803C1BE69695E0" author="Kensley B" journalOrPublisher="Annals of the South African Museum" pagination="261-323" refId="B7" refString="Kensley B (1976) Isopodan and Tanaidacean Crustacea from the St Paul and Amsterdam Islands, Southern Indian Ocean. Annals of the South African Museum 69: 261 - 323." title="Isopodan and Tanaidacean Crustacea from the St Paul and Amsterdam Islands, Southern Indian Ocean." volume="69" year="1976">Kensley, 1976</bibRefCitation>
)
</taxonomicName>
, the only species reported from the Indian Ocean (southern Indian Ocean, Amsterdam Island).
<taxonomicName id="5D191F9E30B5897A8F639E65C97A2004" baseAuthorityName="Kensley" baseAuthorityYear="1976" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" phylum="Arthropoda" rank="species" species="unicornis">
<emphasis id="2973A02B4CE0B6191E76926B2E47FA5A" italics="true">Heterosignum unicornis</emphasis>
</taxonomicName>
has a stout mid-dorsal spine (horn) on the first pereonite, lacking lateral spine-like processes on pereonites 5–7, and a pleotelson with 4 denticles on its lateral margins.
</paragraph>
</subSubSection>
</treatment>
</document>