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<document id="205F57585CE945777E11F5435822439D" ID-CLB-Dataset="301818" ID-DOI="10.5281/zenodo.13272818" ID-GBIF-Dataset="25561bf2-ef51-4d75-a837-3789f9602786" ID-ISSN="2295-0214" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1723146878667" checkinUser="felipe" docAuthor="Vermeersch, Xavier H. C." docDate="2001" docId="30188791FFD72804FE41FCC885E8E183" docLanguage="en" docName="BelgJEntomol.100.1-18.pdf" docOrigin="Belgian Journal of Entomology 100" docSource="http://dx.doi.org/10.5281/zenodo.13272818" docStyle="DocumentStyle:1C24E9253A34C90F9A25106B2B38F3D3.1:BelgJEntomol.2015-2021.journal_article.type1" docStyleId="1C24E9253A34C90F9A25106B2B38F3D3" docStyleName="BelgJEntomol.2015-2021.journal_article.type1" docStyleVersion="1" docTitle="Titanodula fruhstorferi Vermeersch 2001, comb. nov." docType="treatment" docVersion="2" lastPageNumber="18" masterDocId="CC21FFE9FFDC280BFFB1FF858655E17B" masterDocTitle="Euchomenella adwinae sp. nov., a small stick mantis from southern Central Vietnam (Mantodea: Mantidae: Deroplatyinae: Euchomenellini)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1724918174485" updateUser="ExternalLinkService">
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<document id="E4598E253DBFAC7149500F4DECB78118" ID-DOI="10.5281/zenodo.13587531" ID-ISSN="2295-0214" ID-Zenodo-Dep="13587531" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1723146878667" checkinUser="felipe" docAuthor="Vermeersch, Xavier H. C." docDate="2020" docId="30188791FFD72804FE41FCC885E8E183" docLanguage="en" docName="BelgJEntomol.100.1-18.pdf" docOrigin="Belgian Journal of Entomology 100" docStyle="DocumentStyle:1C24E9253A34C90F9A25106B2B38F3D3.1:BelgJEntomol.2015-2021.journal_article.type1" docStyleId="1C24E9253A34C90F9A25106B2B38F3D3" docStyleName="BelgJEntomol.2015-2021.journal_article.type1" docStyleVersion="1" docTitle="Titanodula fruhstorferi Vermeersch 2001, comb. nov." docType="treatment" docVersion="5" lastPageNumber="18" masterDocId="CC21FFE9FFDC280BFFB1FF858655E17B" masterDocTitle="Titanodula gen. nov., a new genus of giant Oriental praying mantises (Mantodea: Mantidae: Hierodulinae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="1" updateTime="1725003596921" updateUser="ExternalLinkService">
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<mods:titleInfo id="EE873FADBF2D4FE9D18EBA182B1877D6">
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<mods:titleInfo id="785740081ACDE7F2FEEC0E0F84203036">
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<mods:title id="50AF2DF719F155528C58A00943F5300A">Euchomenella adwinae sp. nov., a small stick mantis from southern Central Vietnam (Mantodea: Mantidae: Deroplatyinae: Euchomenellini)</mods:title>
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<mods:title id="E162DACDBFE9BEBE2490F7B390FD068A">Titanodula gen. nov., a new genus of giant Oriental praying mantises (Mantodea: Mantidae: Hierodulinae)</mods:title>
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<mods:namePart id="8C03B3DF24707F3CDED581FACC426908">Vermeersch, Xavier H. C.</mods:namePart>
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<mods:namePart id="C652A10F2558496BA704B71B1B72A8C9">Vermeersch, Xavier H. C.</mods:namePart>
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<mods:affiliation id="0AB9798044A7F23D1E7C60EC84AAD28D">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautierstreet 29, B- 1000 Brussels, Belgium</mods:affiliation>
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<mods:nameIdentifier id="E9FD061BEA82B17758DE703478C734E0" type="email">xavier.vermeersch@outlook.com</mods:nameIdentifier>
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<mods:title id="50A947855BE3C592681D7468805280F3">Belgian Journal of Entomology</mods:title>
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<mods:title id="168D6E7DEA2FB194C16210A050A782D0">Belgian Journal of Entomology</mods:title>
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<mods:date id="F4EDD7A0BA6DEF0D0A7351A7D2CF57EB">2001</mods:date>
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<mods:number id="6E629399CD9E45D508731D136E026A3D">2020-07-28</mods:number>
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<treatment id="30188791FFD72804FE41FCC885E8E183" ID-DOI="http://doi.org/10.5281/zenodo.13449993" ID-Zenodo-Dep="13449993" LSID="urn:lsid:plazi:treatment:30188791FFD72804FE41FCC885E8E183" httpUri="http://treatment.plazi.org/id/30188791FFD72804FE41FCC885E8E183" lastPageId="15" pageId="11">
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<treatment id="30188791FFD72804FE41FCC885E8E183" ID-DOI="http://doi.org/10.5281/zenodo.13449993" ID-GBIF-Taxon="239708203" ID-Zenodo-Dep="13449993" LSID="urn:lsid:plazi:treatment:30188791FFD72804FE41FCC885E8E183" httpUri="http://treatment.plazi.org/id/30188791FFD72804FE41FCC885E8E183" lastPageId="15" pageId="11">
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<subSubSection id="F0AB650CFFD72800FE41FCC88208E213" box="[496,1117,845,872]" pageId="11" type="nomenclature">
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<subSubSection id="F0AB650CFFD72800FE41FCC88208E213" box="[496,1117,845,872]" pageId="11" type="nomenclature">
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<paragraph id="B80E3687FFD72800FE41FCC88208E213" blockId="11.[496,1117,845,907]" box="[496,1117,845,872]" pageId="11">
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<paragraph id="B80E3687FFD72800FE41FCC88208E213" blockId="11.[496,1117,845,907]" box="[496,1117,845,872]" pageId="11">
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<emphasis id="8AC5EA95FFD72800FE41FCC88208E213" bold="true" box="[496,1117,845,872]" pageId="11">
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@ -49,7 +51,7 @@
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<subSubSection id="F0AB650CFFD72800FD40FCF4853AE2F0" box="[753,879,881,907]" pageId="11" type="description">
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<subSubSection id="F0AB650CFFD72800FD40FCF4853AE2F0" box="[753,879,881,907]" pageId="11" type="description">
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<paragraph id="B80E3687FFD72800FD40FCF4853AE2F0" blockId="11.[496,1117,845,907]" box="[753,879,881,907]" pageId="11">
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<paragraph id="B80E3687FFD72800FD40FCF4853AE2F0" blockId="11.[496,1117,845,907]" box="[753,879,881,907]" pageId="11">
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(
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(
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<figureCitation id="208A2A02FFD72800FD48FCF48530E2F0" box="[761,869,881,907]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="11.[247,284,1721,1743]" captionStartId-1="12.[247,284,1798,1820]" captionStartId-2="14.[247,284,1835,1857]" captionTargetBox-0="[251,1383,1100,1695]" captionTargetBox-1="[354,1288,185,1775]" captionTargetBox-2="[298,1335,180,1812]" captionTargetId-0="figure-396@11.[247,1385,1099,1700]" captionTargetId-1="figure-74@12.[330,1303,178,1778]" captionTargetId-2="figure-60@14.[296,1336,178,1814]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Fig. 4. Titanodula fruhstorferi comb. nov., additional specimen ♂ [AS1 – Tam Dao N.P.] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. Titanodula fruhstorferi comb. nov., holotype ♀ (NHMW). E, habitus, dorsal view. F, habitus, ventral view. G, detail of head and prothoracic legs. H, labels. © A−D, Xavier Vermeersch and E−H, Harald Bruckner)" captionText-1="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." captionText-2="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." pageId="11">Figs 4−6</figureCitation>
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<figureCitation id="208A2A02FFD72800FD48FCF48530E2F0" box="[761,869,881,907]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="11.[247,284,1721,1743]" captionStartId-1="12.[247,284,1798,1820]" captionStartId-2="14.[247,284,1835,1857]" captionTargetBox-0="[251,1383,1100,1695]" captionTargetBox-1="[354,1288,185,1775]" captionTargetBox-2="[298,1335,180,1812]" captionTargetId-0="figure-396@11.[247,1385,1099,1700]" captionTargetId-1="figure-74@12.[330,1303,178,1778]" captionTargetId-2="figure-60@14.[296,1336,178,1814]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Fig. 4. Titanodula fruhstorferi comb. nov., additional specimen ♂ [AS1 – Tam Dao N.P.] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. Titanodula fruhstorferi comb. nov., holotype ♀ (NHMW). E, habitus, dorsal view. F, habitus, ventral view. G, detail of head and prothoracic legs. H, labels. © A−D, Xavier Vermeersch and E−H, Harald Bruckner)" captionText-1="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." captionText-2="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." figureDoi-0="http://doi.org/10.5281/zenodo.13587551" figureDoi-1="http://doi.org/10.5281/zenodo.13587555" figureDoi-2="http://doi.org/10.5281/zenodo.13587559" httpUri-0="https://zenodo.org/record/13587551/files/figure.png" httpUri-1="https://zenodo.org/record/13587555/files/figure.png" httpUri-2="https://zenodo.org/record/13587559/files/figure.png" pageId="11">Figs 4−6</figureCitation>
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)
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)
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</paragraph>
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</paragraph>
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</subSubSection>
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</subSubSection>
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@ -74,7 +76,7 @@ in external appearance although profemoral AvS 1, 2, 10 and 15 and the basal spo
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However, both species can be readily differentiated by the unique morphology of male genitalia.
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However, both species can be readily differentiated by the unique morphology of male genitalia.
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</paragraph>
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</paragraph>
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</subSubSection>
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</subSubSection>
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<caption id="ECCE660FFFD72800FF46F93C823BE652" ID-DOI="http://doi.org/10.5281/zenodo.13587551" ID-Zenodo-Dep="13587551" httpUri="https://zenodo.org/record/13587551/files/figure.png" pageId="11" startId="11.[247,284,1721,1743]" targetBox="[251,1383,1100,1695]" targetPageId="11" targetType="figure">
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<paragraph id="B80E3687FFD72800FF46F93C823BE652" blockId="11.[247,1386,1721,1833]" pageId="11">
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<paragraph id="B80E3687FFD72800FF46F93C823BE652" blockId="11.[247,1386,1721,1833]" pageId="11">
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Fig. 4.
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Fig. 4.
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<taxonomicName id="7FB14D04FFD72800FEF0F93C847EE7B5" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[321,555,1721,1742]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="11" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
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<taxonomicName id="7FB14D04FFD72800FEF0F93C847EE7B5" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[321,555,1721,1742]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="11" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
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@ -89,7 +91,7 @@ Fig. 4.
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, holotype ♀ (NHMW). E, habitus, dorsal view. F, habitus, ventral view. G, detail of head and prothoracic legs. H, labels. © A−D, Xavier Vermeersch and E−H, Harald Bruckner)
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, holotype ♀ (NHMW). E, habitus, dorsal view. F, habitus, ventral view. G, detail of head and prothoracic legs. H, labels. © A−D, Xavier Vermeersch and E−H, Harald Bruckner)
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</paragraph>
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</paragraph>
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</caption>
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</caption>
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<caption id="ECCE660FFFD02807FF46F88382A6E60D" pageId="12" startId="12.[247,284,1798,1820]" targetBox="[354,1288,185,1775]" targetPageId="12" targetType="figure">
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<caption id="ECCE660FFFD02807FF46F88382A6E60D" ID-DOI="http://doi.org/10.5281/zenodo.13587555" ID-Zenodo-Dep="13587555" httpUri="https://zenodo.org/record/13587555/files/figure.png" pageId="12" startId="12.[247,284,1798,1820]" targetBox="[354,1288,185,1775]" targetPageId="12" targetType="figure">
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<paragraph id="B80E3687FFD02807FF46F88382A6E60D" blockId="12.[247,1385,1798,1910]" pageId="12">
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<paragraph id="B80E3687FFD02807FF46F88382A6E60D" blockId="12.[247,1385,1798,1910]" pageId="12">
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Fig. 5.
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Fig. 5.
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<taxonomicName id="7FB14D04FFD02807FEF1F883847DE660" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[320,552,1798,1819]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="12" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
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<taxonomicName id="7FB14D04FFD02807FEF1F883847DE660" authorityName="Vermeersch" authorityYear="2001" baseAuthorityName="Werner" baseAuthorityYear="1916" box="[320,552,1798,1819]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="12" phylum="Arthropoda" rank="species" species="fruhstorferi" status="comb. nov.">
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@ -106,7 +108,7 @@ Fig. 5.
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</caption>
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</caption>
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<subSubSection id="F0AB650CFFD12806FF46FF3E8285E18C" pageId="13" type="materials_examined">
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<subSubSection id="F0AB650CFFD12806FF46FF3E8285E18C" pageId="13" type="materials_examined">
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<paragraph id="B80E3687FFD12806FF46FF3E8285E18C" blockId="13.[247,1386,183,248]" pageId="13">
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<paragraph id="B80E3687FFD12806FF46FF3E8285E18C" blockId="13.[247,1386,183,248]" pageId="13">
|
||||||
<materialsCitation id="08D93CDAFFD12806FF46FF3E8285E18C" collectionCode="MATERIAL, TYPE, NHMW" collectorName="H. Fruhstorfer" country="Vietnam" location="Vietnam" pageId="13" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
<materialsCitation id="08D93CDAFFD12806FF46FF3E8285E18C" ID-GBIF-Occurrence="4926423303" collectionCode="MATERIAL, TYPE, NHMW" collectorName="H. Fruhstorfer" country="Vietnam" location="Vietnam" pageId="13" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||||||
<smallCapsWord id="BEE8A05BFFD12806FF46FF3E872DE1A8" baselines="206,206" box="[247,376,187,213]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Material" pageId="13">
|
<smallCapsWord id="BEE8A05BFFD12806FF46FF3E872DE1A8" baselines="206,206" box="[247,376,187,213]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Material" pageId="13">
|
||||||
<collectionCode id="DEA0AE42FFD12806FF46FF3E872DE1A8" box="[247,376,187,213]" pageId="13">MATERIAL</collectionCode>
|
<collectionCode id="DEA0AE42FFD12806FF46FF3E872DE1A8" box="[247,376,187,213]" pageId="13">MATERIAL</collectionCode>
|
||||||
</smallCapsWord>
|
</smallCapsWord>
|
||||||
|
|
@ -124,7 +126,7 @@ Fig. 5.
|
||||||
<specimenCount id="AEB7FD0EFFD12806FAE2FF32833FE1AF" box="[1363,1386,183,212]" count="1" pageId="13" type="female">♀</specimenCount>
|
<specimenCount id="AEB7FD0EFFD12806FAE2FF32833FE1AF" box="[1363,1386,183,212]" count="1" pageId="13" type="female">♀</specimenCount>
|
||||||
</emphasis>
|
</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD12806FEB1FF588712E18C" box="[256,327,221,247]" captionStart="Fig" captionStartId="11.[247,284,1721,1743]" captionTargetBox="[251,1383,1100,1695]" captionTargetId="figure-396@11.[247,1385,1099,1700]" captionTargetPageId="11" captionText="Fig. 4. Titanodula fruhstorferi comb. nov., additional specimen ♂ [AS1 – Tam Dao N.P.] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. Titanodula fruhstorferi comb. nov., holotype ♀ (NHMW). E, habitus, dorsal view. F, habitus, ventral view. G, detail of head and prothoracic legs. H, labels. © A−D, Xavier Vermeersch and E−H, Harald Bruckner)" pageId="13">Fig. 4</figureCitation>
|
<figureCitation id="208A2A02FFD12806FEB1FF588712E18C" box="[256,327,221,247]" captionStart="Fig" captionStartId="11.[247,284,1721,1743]" captionTargetBox="[251,1383,1100,1695]" captionTargetId="figure-396@11.[247,1385,1099,1700]" captionTargetPageId="11" captionText="Fig. 4. Titanodula fruhstorferi comb. nov., additional specimen ♂ [AS1 – Tam Dao N.P.] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. Titanodula fruhstorferi comb. nov., holotype ♀ (NHMW). E, habitus, dorsal view. F, habitus, ventral view. G, detail of head and prothoracic legs. H, labels. © A−D, Xavier Vermeersch and E−H, Harald Bruckner)" figureDoi="http://doi.org/10.5281/zenodo.13587551" httpUri="https://zenodo.org/record/13587551/files/figure.png" pageId="13">Fig. 4</figureCitation>
|
||||||
E−F):
|
E−F):
|
||||||
<collectingCountry id="C0A67617FFD12806FE2CFF58845DE18C" box="[413,520,221,247]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
|
<collectingCountry id="C0A67617FFD12806FE2CFF58845DE18C" box="[413,520,221,247]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
|
||||||
, Tonkin, Than Moi, VI−VII, leg.
|
, Tonkin, Than Moi, VI−VII, leg.
|
||||||
|
|
@ -159,7 +161,7 @@ damaged, abdomen and metathoracic legs are missing.
|
||||||
): (
|
): (
|
||||||
<specimenCount id="AEB7FD0EFFD12806FD65FEB884AEE02C" box="[724,763,317,343]" count="1" pageId="13" type="male">1♂</specimenCount>
|
<specimenCount id="AEB7FD0EFFD12806FD65FEB884AEE02C" box="[724,763,317,343]" count="1" pageId="13" type="male">1♂</specimenCount>
|
||||||
):
|
):
|
||||||
<materialsCitation id="08D93CDAFFD12806FCA6FEB98517E0E6" collectingDate="1833-01" collectionCode="R" collectorName="Tam Dao" country="Vietnam" latitude="22.2" location="Cham Chu" longitude="105.1" pageId="13" specimenCount="1" specimenCount-male="3">
|
<materialsCitation id="08D93CDAFFD12806FCA6FEB98517E0E6" ID-GBIF-Occurrence="4926423306" collectingDate="1833-01" collectionCode="R" collectorName="Tam Dao" country="Vietnam" latitude="22.2" location="Cham Chu" longitude="105.1" pageId="13" specimenCount="1" specimenCount-male="3">
|
||||||
<collectingCountry id="C0A67617FFD12806FCA6FEB985D7E02D" box="[791,898,316,342]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
|
<collectingCountry id="C0A67617FFD12806FCA6FEB985D7E02D" box="[791,898,316,342]" name="Vietnam" pageId="13">Vietnam</collectingCountry>
|
||||||
,
|
,
|
||||||
<location id="BD6E605CFFD12806FC22FEB9824CE02D" LSID="urn:lsid:plazi:treatment:30188791FFD72804FE41FCC885E8E183:BD6E605CFFD12806FC22FEB9824CE02D" box="[915,1049,316,342]" country="Vietnam" name="Cham Chu" pageId="13">Cham Chu</location>
|
<location id="BD6E605CFFD12806FC22FEB9824CE02D" LSID="urn:lsid:plazi:treatment:30188791FFD72804FE41FCC885E8E183:BD6E605CFFD12806FC22FEB9824CE02D" box="[915,1049,316,342]" country="Vietnam" name="Cham Chu" pageId="13">Cham Chu</location>
|
||||||
|
|
@ -181,7 +183,7 @@ N.
|
||||||
<collectionCode id="DEA0AE42FFD12806FD67FE068560E0E6" box="[726,821,387,413]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="13" type="Museum">RBINS</collectionCode>
|
<collectionCode id="DEA0AE42FFD12806FD67FE068560E0E6" box="[726,821,387,413]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="13" type="Museum">RBINS</collectionCode>
|
||||||
).
|
).
|
||||||
</materialsCitation>
|
</materialsCitation>
|
||||||
<materialsCitation id="08D93CDAFFD12806FCFFFE06853DE098" collectingDate="1833-01" collectionCode="R" collectorName="Tam Dao" country="Vietnam" latitude="21.516666" location="Cham Chu" longitude="105.55" pageId="13" specimenCount="1" specimenCount-male="3">
|
<materialsCitation id="08D93CDAFFD12806FCFFFE06853DE098" ID-GBIF-Occurrence="4926423308" collectingDate="1833-01" collectionCode="R" collectorName="Tam Dao" country="Vietnam" latitude="21.516666" location="Cham Chu" longitude="105.55" pageId="13" specimenCount="1" specimenCount-male="3">
|
||||||
(
|
(
|
||||||
<specimenCount id="AEB7FD0EFFD12806FCE9FE0685CCE0E6" box="[856,921,387,413]" count="3" pageId="13" type="male">3♂♂</specimenCount>
|
<specimenCount id="AEB7FD0EFFD12806FCE9FE0685CCE0E6" box="[856,921,387,413]" count="3" pageId="13" type="male">3♂♂</specimenCount>
|
||||||
):
|
):
|
||||||
|
|
@ -202,7 +204,7 @@ N.P.,
|
||||||
<collectionCode id="DEA0AE42FFD12806FD4DFE4C850EE098" box="[764,859,457,483]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="13" type="Museum">RBINS</collectionCode>
|
<collectionCode id="DEA0AE42FFD12806FD4DFE4C850EE098" box="[764,859,457,483]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="13" type="Museum">RBINS</collectionCode>
|
||||||
).
|
).
|
||||||
</materialsCitation>
|
</materialsCitation>
|
||||||
<materialsCitation id="08D93CDAFFD12806FCCAFE4C826DE351" collectingDate="1833-01" collectionCode="R" collectorName="Tam Dao" country="Vietnam" latitude="20.316668" location="Cham Chu" longitude="105.60833" pageId="13" specimenCount="1" specimenCount-female="1" specimenCount-male="3">
|
<materialsCitation id="08D93CDAFFD12806FCCAFE4C826DE351" ID-GBIF-Occurrence="4926423304" collectingDate="1833-01" collectionCode="R" collectorName="Tam Dao" country="Vietnam" latitude="20.316668" location="Cham Chu" longitude="105.60833" pageId="13" specimenCount="1" specimenCount-female="1" specimenCount-male="3">
|
||||||
(
|
(
|
||||||
<specimenCount id="AEB7FD0EFFD12806FC34FE4C8583E098" box="[901,982,457,483]" count="10" pageId="13" type="male">10♂♂</specimenCount>
|
<specimenCount id="AEB7FD0EFFD12806FC34FE4C8583E098" box="[901,982,457,483]" count="10" pageId="13" type="male">10♂♂</specimenCount>
|
||||||
):
|
):
|
||||||
|
|
@ -223,7 +225,7 @@ N.P.,
|
||||||
<collectionCode id="DEA0AE42FFD12806FC7DFD8A827DE352" box="[972,1064,527,553]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="13" type="Museum">RBINS</collectionCode>
|
<collectionCode id="DEA0AE42FFD12806FC7DFD8A827DE352" box="[972,1064,527,553]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="13" type="Museum">RBINS</collectionCode>
|
||||||
).
|
).
|
||||||
</materialsCitation>
|
</materialsCitation>
|
||||||
<materialsCitation id="08D93CDAFFD12806FB8EFD8A84C4E314" collectingDate="1833-01" collectingDateMax="2013-07-11" collectingDateMin="2013-07-07" collectionCode="RBINS" collectorName="J. Bresseel" country="Vietnam" latitude="21.18611" location="Cham Chu" longLatPrecision="21" longitude="106.72389" pageId="13" specimenCount="1" specimenCount-female="1" specimenCount-male="3">
|
<materialsCitation id="08D93CDAFFD12806FB8EFD8A84C4E314" ID-GBIF-Occurrence="4926423301" collectingDate="1833-01" collectingDateMax="2013-07-11" collectingDateMin="2013-07-07" collectionCode="RBINS" collectorName="J. Bresseel" country="Vietnam" latitude="21.18611" location="Cham Chu" longLatPrecision="21" longitude="106.72389" pageId="13" specimenCount="1" specimenCount-female="1" specimenCount-male="3">
|
||||||
(
|
(
|
||||||
<specimenCount id="AEB7FD0EFFD12806FBFAFD8A8225E352" box="[1099,1136,524,553]" count="1" pageId="13" type="female">
|
<specimenCount id="AEB7FD0EFFD12806FBFAFD8A8225E352" box="[1099,1136,524,553]" count="1" pageId="13" type="female">
|
||||||
1
|
1
|
||||||
|
|
@ -262,7 +264,7 @@ N.
|
||||||
<paragraph id="B80E3687FFD12806FF46FD038207E209" blockId="13.[247,1386,645,882]" pageId="13">
|
<paragraph id="B80E3687FFD12806FF46FD038207E209" blockId="13.[247,1386,645,882]" pageId="13">
|
||||||
<smallCapsWord id="BEE8A05BFFD12806FF46FD0387E1E3E5" baselines="665,665" box="[247,436,646,672]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Redescription" pageId="13">REDESCRIPTION</smallCapsWord>
|
<smallCapsWord id="BEE8A05BFFD12806FF46FD0387E1E3E5" baselines="665,665" box="[247,436,646,672]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Redescription" pageId="13">REDESCRIPTION</smallCapsWord>
|
||||||
. Measurements (see
|
. Measurements (see
|
||||||
<tableCitation id="F533033CFFD12806FD04FD008515E3E4" box="[693,832,645,671]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="7.[247,304,421,443]" captionStartId-1="9.[247,304,316,338]" captionText-0="Table 1. Detailed measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. From left to right: T. attenboroughi holotype ♂, T. attenboroughi paratype ♀, T. fruhstorferi male, additional specimen 1, T. fruhstorferi female, additional specimen 2." captionText-1="Table 2. Measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. to assess intraspecific variability." pageId="13">Tables 1−2</tableCitation>
|
<tableCitation id="F533033CFFD12806FD04FD008515E3E4" box="[693,832,645,671]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="7.[247,304,421,443]" captionStartId-1="9.[247,304,316,338]" captionTargetId-0="graphics-141@7.[282,1350,516,1675]" captionTargetId-1="graphics-59@9.[355,1278,381,1304]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="Table 1. Detailed measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. From left to right: T. attenboroughi holotype ♂, T. attenboroughi paratype ♀, T. fruhstorferi male, additional specimen 1, T. fruhstorferi female, additional specimen 2." captionText-1="Table 2. Measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. to assess intraspecific variability." httpUri-0="http://table.plazi.org/id/ECCE660FFFDB280CFF46FE208568E08C" httpUri-1="http://table.plazi.org/id/ECCE660FFFD52802FF46FEB9847EE00B" pageId="13" tableUuid-0="ECCE660FFFDB280CFF46FE208568E08C" tableUuid-1="ECCE660FFFD52802FF46FEB9847EE00B">Tables 1−2</tableCitation>
|
||||||
). Very large and robust praying mantis. Head triangular, antennae filiform. Long but robust pronotum, with smooth dorsal surface. Pronotum very finely denticulate along the margins of the prozone in female, without denticulations in the metazone, entirely smooth edges in males. Spinal formula: F = 4DS/15AvS/4PvS; T = 14AvS/10PvS. Ratios
|
). Very large and robust praying mantis. Head triangular, antennae filiform. Long but robust pronotum, with smooth dorsal surface. Pronotum very finely denticulate along the margins of the prozone in female, without denticulations in the metazone, entirely smooth edges in males. Spinal formula: F = 4DS/15AvS/4PvS; T = 14AvS/10PvS. Ratios
|
||||||
<specimenCount id="AEB7FD0EFFD12806FDB7FC978475E257" box="[518,544,786,812]" count="1" pageId="13" type="male">♂</specimenCount>
|
<specimenCount id="AEB7FD0EFFD12806FDB7FC978475E257" box="[518,544,786,812]" count="1" pageId="13" type="male">♂</specimenCount>
|
||||||
: MzL/PzL: 3.1;
|
: MzL/PzL: 3.1;
|
||||||
|
|
@ -278,7 +280,7 @@ anteroventral spine. Protarsus (all segments) entirely black on the anterior sid
|
||||||
<paragraph id="B80E3687FFD12806FF46FC0287D5E761" blockId="13.[247,1386,903,1562]" pageId="13">
|
<paragraph id="B80E3687FFD12806FF46FC0287D5E761" blockId="13.[247,1386,903,1562]" pageId="13">
|
||||||
<emphasis id="8AC5EA95FFD12806FF46FC0287FCE2DA" box="[247,425,903,929]" italics="true" pageId="13">Male genitalia</emphasis>
|
<emphasis id="8AC5EA95FFD12806FF46FC0287FCE2DA" box="[247,425,903,929]" italics="true" pageId="13">Male genitalia</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD12806FE0BFC028471E2D9" box="[442,548,903,930]" captionStart="Fig" captionStartId="12.[247,284,1798,1820]" captionTargetBox="[354,1288,185,1775]" captionTargetId="figure-74@12.[330,1303,178,1778]" captionTargetPageId="12" captionText="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." pageId="13">Fig. 5 A</figureCitation>
|
<figureCitation id="208A2A02FFD12806FE0BFC028471E2D9" box="[442,548,903,930]" captionStart="Fig" captionStartId="12.[247,284,1798,1820]" captionTargetBox="[354,1288,185,1775]" captionTargetId="figure-74@12.[330,1303,178,1778]" captionTargetPageId="12" captionText="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." figureDoi="http://doi.org/10.5281/zenodo.13587555" httpUri="https://zenodo.org/record/13587555/files/figure.png" pageId="13">Fig. 5 A</figureCitation>
|
||||||
): Ventral phallomere of left phallic complex elongated, much longer than broad, with two short and strongly sclerotised distal processes (sdp) of about the same length with acute apex. Median process (sdpm) bend at the middle and pointing sideways, lateral process (sdpl) much more strongly bend and pointing upwards, with much broader base than sdpm. Apical process of left phallomere (paa) complex in shape, with sclerotised apex and sclerotizations near the base, but membranous in the middle. Apex abruptly projecting dorsally and to the left, the apex sharply curved in a 90° hook with narrower but rounded tip. Phalloid apophysis (afa) with two sclerotised processes, anterior process (aafa) small and flat, tubercle-shaped, without additional projections or extensions. Posterior process (pafa) spear-shaped, with very strongly developed base, bulging anteriorly, strongly curving towards an almost straight, very long and heavily sclerotised structure, projecting straight posteriad. Membranous lobe loa rounded and small. Genital lobe goa membranous, sometimes well-developed anteriorly but in some specimens also completely collapsed, and much weaker and wrinkled in appearance posteriorly, running almost parallel to pafa posteriad. Right phallomere with sclerite R1A more or less triangular. Sclerite R3 shovel-shaped, very wide at its anterior end. Ventral process (pia) triangular with many longitudinal grooves on its surface, sclerified ventral process (pva) strongly sclerotized, finger-shaped, ending in a blunt hook. Coxosternite IX (subgenital plate) with small black spikes on the right lateral edge, sometimes also in the middle of the distal edge.
|
): Ventral phallomere of left phallic complex elongated, much longer than broad, with two short and strongly sclerotised distal processes (sdp) of about the same length with acute apex. Median process (sdpm) bend at the middle and pointing sideways, lateral process (sdpl) much more strongly bend and pointing upwards, with much broader base than sdpm. Apical process of left phallomere (paa) complex in shape, with sclerotised apex and sclerotizations near the base, but membranous in the middle. Apex abruptly projecting dorsally and to the left, the apex sharply curved in a 90° hook with narrower but rounded tip. Phalloid apophysis (afa) with two sclerotised processes, anterior process (aafa) small and flat, tubercle-shaped, without additional projections or extensions. Posterior process (pafa) spear-shaped, with very strongly developed base, bulging anteriorly, strongly curving towards an almost straight, very long and heavily sclerotised structure, projecting straight posteriad. Membranous lobe loa rounded and small. Genital lobe goa membranous, sometimes well-developed anteriorly but in some specimens also completely collapsed, and much weaker and wrinkled in appearance posteriorly, running almost parallel to pafa posteriad. Right phallomere with sclerite R1A more or less triangular. Sclerite R3 shovel-shaped, very wide at its anterior end. Ventral process (pia) triangular with many longitudinal grooves on its surface, sclerified ventral process (pva) strongly sclerotized, finger-shaped, ending in a blunt hook. Coxosternite IX (subgenital plate) with small black spikes on the right lateral edge, sometimes also in the middle of the distal edge.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
</subSubSection>
|
</subSubSection>
|
||||||
|
|
@ -299,7 +301,7 @@ specialist.
|
||||||
</taxonomicName>
|
</taxonomicName>
|
||||||
<taxonomicNameLabel id="91F657EEFFD12806FDC5F9208554E7BB" box="[628,769,1701,1728]" pageId="13" rank="species">comb. nov.</taxonomicNameLabel>
|
<taxonomicNameLabel id="91F657EEFFD12806FDC5F9208554E7BB" box="[628,769,1701,1728]" pageId="13" rank="species">comb. nov.</taxonomicNameLabel>
|
||||||
inhabits the closed canopy forests of the North Vietnamese lowlands (
|
inhabits the closed canopy forests of the North Vietnamese lowlands (
|
||||||
<figureCitation id="208A2A02FFD12806FDA3F94D8409E799" box="[530,604,1736,1762]" captionStart="Fig" captionStartId="14.[247,284,1835,1857]" captionTargetBox="[298,1335,180,1812]" captionTargetId="figure-60@14.[296,1336,178,1814]" captionText="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." pageId="13">Fig. 6</figureCitation>
|
<figureCitation id="208A2A02FFD12806FDA3F94D8409E799" box="[530,604,1736,1762]" captionStart="Fig" captionStartId="14.[247,284,1835,1857]" captionTargetBox="[298,1335,180,1812]" captionTargetId="figure-60@14.[296,1336,178,1814]" captionText="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." figureDoi="http://doi.org/10.5281/zenodo.13587559" httpUri="https://zenodo.org/record/13587559/files/figure.png" pageId="13">Fig. 6</figureCitation>
|
||||||
). Forests in this area are highly threatened due to a high rate of deforestation and suitable habitats are strongly fragmented with large gaps between them. More distribution data is needed to assess its conservation status.
|
). Forests in this area are highly threatened due to a high rate of deforestation and suitable habitats are strongly fragmented with large gaps between them. More distribution data is needed to assess its conservation status.
|
||||||
</paragraph>
|
</paragraph>
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@ -1,40 +1,42 @@
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<figureCitation id="208A2A02FFD8280FFD6BFAE0851DE404" box="[730,840,1381,1407]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[247,284,1261,1283]" captionStartId-1="8.[247,284,1459,1481]" captionStartId-2="10.[247,284,1431,1453]" captionTargetBox-0="[254,1376,536,1218]" captionTargetBox-1="[256,1378,887,1429]" captionTargetBox-2="[247,1385,273,1410]" captionTargetId-0="figure-400@6.[247,1385,529,1227]" captionTargetId-1="figure-458@8.[247,1385,870,1438]" captionTargetId-2="figure-209@10.[247,1385,273,1410]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-1="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-2="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="4">Figs 1−3</figureCitation>
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<figureCitation id="208A2A02FFD8280FFD6BFAE0851DE404" box="[730,840,1381,1407]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[247,284,1261,1283]" captionStartId-1="8.[247,284,1459,1481]" captionStartId-2="10.[247,284,1431,1453]" captionTargetBox-0="[254,1376,536,1218]" captionTargetBox-1="[256,1378,887,1429]" captionTargetBox-2="[247,1385,273,1410]" captionTargetId-0="figure-400@6.[247,1385,529,1227]" captionTargetId-1="figure-458@8.[247,1385,870,1438]" captionTargetId-2="figure-209@10.[247,1385,273,1410]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-1="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-2="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." figureDoi-0="http://doi.org/10.5281/zenodo.13587537" figureDoi-1="http://doi.org/10.5281/zenodo.13587541" figureDoi-2="http://doi.org/10.5281/zenodo.13587547" httpUri-0="https://zenodo.org/record/13587537/files/figure.png" httpUri-1="https://zenodo.org/record/13587541/files/figure.png" httpUri-2="https://zenodo.org/record/13587547/files/figure.png" pageId="4">Figs 1−3</figureCitation>
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||||||
,
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,
|
||||||
<figureCitation id="208A2A02FFD8280FFCE4FAE085D0E4FB" box="[853,901,1381,1408]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[247,284,1798,1820]" captionStartId-1="14.[247,284,1835,1857]" captionTargetBox-0="[354,1288,185,1775]" captionTargetBox-1="[298,1335,180,1812]" captionTargetId-0="figure-74@12.[330,1303,178,1778]" captionTargetId-1="figure-60@14.[296,1336,178,1814]" captionTargetPageId-0="12" captionText-0="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." captionText-1="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." pageId="4">5−6</figureCitation>
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<figureCitation id="208A2A02FFD8280FFCE4FAE085D0E4FB" box="[853,901,1381,1408]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[247,284,1798,1820]" captionStartId-1="14.[247,284,1835,1857]" captionTargetBox-0="[354,1288,185,1775]" captionTargetBox-1="[298,1335,180,1812]" captionTargetId-0="figure-74@12.[330,1303,178,1778]" captionTargetId-1="figure-60@14.[296,1336,178,1814]" captionTargetPageId-0="12" captionText-0="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." captionText-1="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." figureDoi-0="http://doi.org/10.5281/zenodo.13587555" figureDoi-1="http://doi.org/10.5281/zenodo.13587559" httpUri-0="https://zenodo.org/record/13587555/files/figure.png" httpUri-1="https://zenodo.org/record/13587559/files/figure.png" pageId="4">5−6</figureCitation>
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)
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)
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<mods:title id="8ABD3005D3B4BE5762A47813F6B2BAE7">Euchomenella adwinae sp. nov., a small stick mantis from southern Central Vietnam (Mantodea: Mantidae: Deroplatyinae: Euchomenellini)</mods:title>
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<mods:title id="8FB1631D9FA5F2EB17BD208024799F28">Titanodula gen. nov., a new genus of giant Oriental praying mantises (Mantodea: Mantidae: Hierodulinae)</mods:title>
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<mods:namePart id="BED4A6EA490902B0406950D5C3727C03">Vermeersch, Xavier H. C.</mods:namePart>
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<mods:affiliation id="AA254FC058FED9FD69961218608F4D45">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautierstreet 29, B- 1000 Brussels, Belgium</mods:affiliation>
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<mods:nameIdentifier id="3A8B72155753D044E4C94AE378C09A19" type="email">xavier.vermeersch@outlook.com</mods:nameIdentifier>
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<mods:title id="AC8DD29997A92B7E296E6C8B4398283F">Belgian Journal of Entomology</mods:title>
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<mods:title id="5E0A4CD1F461B05724FDA8308B3DC322">Belgian Journal of Entomology</mods:title>
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<treatment id="30188791FFD92800FDEAFAD482DCE250" ID-DOI="http://doi.org/10.5281/zenodo.13449985" ID-Zenodo-Dep="13449985" LSID="urn:lsid:plazi:treatment:30188791FFD92800FDEAFAD482DCE250" httpUri="http://treatment.plazi.org/id/30188791FFD92800FDEAFAD482DCE250" lastPageId="11" pageId="5">
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<treatment id="30188791FFD92800FDEAFAD482DCE250" ID-DOI="http://doi.org/10.5281/zenodo.13449985" ID-GBIF-Taxon="239708202" ID-Zenodo-Dep="13449985" LSID="urn:lsid:zoobank.org:act:4CEFD21C-CB3F-4FB8-A35E-93B0966A5C63" httpUri="http://treatment.plazi.org/id/30188791FFD92800FDEAFAD482DCE250" lastPageId="11" pageId="5">
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<subSubSection id="F0AB650CFFD9280EFDEAFAD48251E417" box="[603,1028,1361,1388]" pageId="5" type="nomenclature">
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<subSubSection id="F0AB650CFFD9280EFDEAFAD48251E417" box="[603,1028,1361,1388]" pageId="5" type="nomenclature">
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||||||
<paragraph id="B80E3687FFD9280EFDEAFAD48251E417" blockId="5.[445,1187,1361,1459]" box="[603,1028,1361,1388]" pageId="5">
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<paragraph id="B80E3687FFD9280EFDEAFAD48251E417" blockId="5.[445,1187,1361,1459]" box="[603,1028,1361,1388]" pageId="5">
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||||||
<emphasis id="8AC5EA95FFD9280EFDEAFAD48251E417" bold="true" box="[603,1028,1361,1388]" pageId="5">
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<emphasis id="8AC5EA95FFD9280EFDEAFAD48251E417" bold="true" box="[603,1028,1361,1388]" pageId="5">
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@ -54,9 +56,9 @@ urn:lsid:zoobank.org:act:
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<paragraph id="B80E3687FFD9280EFD63FA1D85DBE4C8" blockId="5.[445,1187,1361,1459]" box="[722,910,1432,1459]" pageId="5">
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<paragraph id="B80E3687FFD9280EFD63FA1D85DBE4C8" blockId="5.[445,1187,1361,1459]" box="[722,910,1432,1459]" pageId="5">
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||||||
(
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(
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||||||
<figureCitation id="208A2A02FFD9280EFD6BFA1D851DE4C9" box="[730,840,1432,1458]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[247,284,1261,1283]" captionStartId-1="8.[247,284,1459,1481]" captionStartId-2="10.[247,284,1431,1453]" captionTargetBox-0="[254,1376,536,1218]" captionTargetBox-1="[256,1378,887,1429]" captionTargetBox-2="[247,1385,273,1410]" captionTargetId-0="figure-400@6.[247,1385,529,1227]" captionTargetId-1="figure-458@8.[247,1385,870,1438]" captionTargetId-2="figure-209@10.[247,1385,273,1410]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-1="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-2="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="5">Figs 1−3</figureCitation>
|
<figureCitation id="208A2A02FFD9280EFD6BFA1D851DE4C9" box="[730,840,1432,1458]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="6.[247,284,1261,1283]" captionStartId-1="8.[247,284,1459,1481]" captionStartId-2="10.[247,284,1431,1453]" captionTargetBox-0="[254,1376,536,1218]" captionTargetBox-1="[256,1378,887,1429]" captionTargetBox-2="[247,1385,273,1410]" captionTargetId-0="figure-400@6.[247,1385,529,1227]" captionTargetId-1="figure-458@8.[247,1385,870,1438]" captionTargetId-2="figure-209@10.[247,1385,273,1410]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-1="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." captionText-2="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." figureDoi-0="http://doi.org/10.5281/zenodo.13587537" figureDoi-1="http://doi.org/10.5281/zenodo.13587541" figureDoi-2="http://doi.org/10.5281/zenodo.13587547" httpUri-0="https://zenodo.org/record/13587537/files/figure.png" httpUri-1="https://zenodo.org/record/13587541/files/figure.png" httpUri-2="https://zenodo.org/record/13587547/files/figure.png" pageId="5">Figs 1−3</figureCitation>
|
||||||
,
|
,
|
||||||
<figureCitation id="208A2A02FFD9280EFCE4FA1D85D0E4C8" box="[853,901,1432,1459]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[247,284,1798,1820]" captionStartId-1="14.[247,284,1835,1857]" captionTargetBox-0="[354,1288,185,1775]" captionTargetBox-1="[298,1335,180,1812]" captionTargetId-0="figure-74@12.[330,1303,178,1778]" captionTargetId-1="figure-60@14.[296,1336,178,1814]" captionTargetPageId-0="12" captionText-0="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." captionText-1="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." pageId="5">5−6</figureCitation>
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<figureCitation id="208A2A02FFD9280EFCE4FA1D85D0E4C8" box="[853,901,1432,1459]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="12.[247,284,1798,1820]" captionStartId-1="14.[247,284,1835,1857]" captionTargetBox-0="[354,1288,185,1775]" captionTargetBox-1="[298,1335,180,1812]" captionTargetId-0="figure-74@12.[330,1303,178,1778]" captionTargetId-1="figure-60@14.[296,1336,178,1814]" captionTargetPageId-0="12" captionText-0="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." captionText-1="Fig. 6. Distribution map of Titanodula fruhstorferi comb. nov. (▲) and Titanodula attenboroughi sp. nov. (●) in Vietnam. From top to bottom: T. fruhstorferi; Cham Chu N.R., Tam Dao N.P., Tay Yen Tu N.R., Cuc Phuong N.P. T. attenboroughi; Da Krong N.R., Kon Chu Rang N.R." figureDoi-0="http://doi.org/10.5281/zenodo.13587555" figureDoi-1="http://doi.org/10.5281/zenodo.13587559" httpUri-0="https://zenodo.org/record/13587555/files/figure.png" httpUri-1="https://zenodo.org/record/13587559/files/figure.png" pageId="5">5−6</figureCitation>
|
||||||
)
|
)
|
||||||
</paragraph>
|
</paragraph>
|
||||||
</subSubSection>
|
</subSubSection>
|
||||||
|
|
@ -78,7 +80,7 @@ anteroventral spine. Protarsus (all segments) entirely black on the anterior sid
|
||||||
</subSubSection>
|
</subSubSection>
|
||||||
<subSubSection id="F0AB650CFFDA280DFF46FF3E836BE081" pageId="6" type="materials_examined">
|
<subSubSection id="F0AB650CFFDA280DFF46FF3E836BE081" pageId="6" type="materials_examined">
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||||||
<paragraph id="B80E3687FFDA280DFF46FF3E833CE061" blockId="6.[247,1386,183,283]" pageId="6">
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<paragraph id="B80E3687FFDA280DFF46FF3E833CE061" blockId="6.[247,1386,183,283]" pageId="6">
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||||||
<materialsCitation id="08D93CDAFFDA280DFF46FF3E833CE061" collectingDate="1840-01" collectingDateMax="2018-07-20" collectingDateMin="1840-01" collectionCode="RBINS" collectorName="J. Constant & J. Bresseel" country="Vietnam" latitude="14.474444" location="Gia Lai Province" longLatPrecision="21" longitude="108.54083" pageId="6" specimenCount="1" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="holotype">
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<materialsCitation id="08D93CDAFFDA280DFF46FF3E833CE061" ID-GBIF-Occurrence="4926423307" collectingDate="1840-01" collectingDateMax="2018-07-20" collectingDateMin="1840-01" collectionCode="RBINS" collectorName="J. Constant & J. Bresseel" country="Vietnam" latitude="14.474444" location="Gia Lai Province" longLatPrecision="21" longitude="108.54083" pageId="6" specimenCount="1" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="holotype">
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<collectionCode id="DEA0AE42FFDA280DFF46FF3E8763E1A8" box="[247,310,187,213]" pageId="6">TYPE</collectionCode>
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<collectionCode id="DEA0AE42FFDA280DFF46FF3E8763E1A8" box="[247,310,187,213]" pageId="6">TYPE</collectionCode>
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</smallCapsWord>
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||||||
|
|
@ -91,9 +93,9 @@ anteroventral spine. Protarsus (all segments) entirely black on the anterior sid
|
||||||
<specimenCount id="AEB7FD0EFFDA280DFDF9FF32840AE1AF" box="[584,607,183,212]" count="1" pageId="6" type="male">♂</specimenCount>
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<specimenCount id="AEB7FD0EFFDA280DFDF9FF32840AE1AF" box="[584,607,183,212]" count="1" pageId="6" type="male">♂</specimenCount>
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||||||
</emphasis>
|
</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFDA280DFDC0FF3F8494E1AF" box="[625,705,186,212]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Figs 1</figureCitation>
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<figureCitation id="208A2A02FFDA280DFDC0FF3F8494E1AF" box="[625,705,186,212]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="6">Figs 1</figureCitation>
|
||||||
,
|
,
|
||||||
<figureCitation id="208A2A02FFDA280DFD62FF3F84B7E1AF" box="[723,738,186,212]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="6">3</figureCitation>
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<figureCitation id="208A2A02FFDA280DFD62FF3F84B7E1AF" box="[723,738,186,212]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." figureDoi="http://doi.org/10.5281/zenodo.13587547" httpUri="https://zenodo.org/record/13587547/files/figure.png" pageId="6">3</figureCitation>
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||||||
A−C):
|
A−C):
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||||||
<collectingCountry id="C0A67617FFDA280DFCF4FF3F85E5E1AF" box="[837,944,186,212]" name="Vietnam" pageId="6">Vietnam</collectingCountry>
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<collectingCountry id="C0A67617FFDA280DFCF4FF3F85E5E1AF" box="[837,944,186,212]" name="Vietnam" pageId="6">Vietnam</collectingCountry>
|
||||||
,
|
,
|
||||||
|
|
@ -141,9 +143,9 @@ prep.
|
||||||
):
|
):
|
||||||
<specimenCount id="AEB7FD0EFFDA280DFD9DFEB5843CE031" box="[556,617,304,330]" count="2" pageId="6" type="female">2♀♀</specimenCount>
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<specimenCount id="AEB7FD0EFFDA280DFD9DFEB5843CE031" box="[556,617,304,330]" count="2" pageId="6" type="female">2♀♀</specimenCount>
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||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFDA280DFDCFFEB58498E031" box="[638,717,304,330]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Fig. 2</figureCitation>
|
<figureCitation id="208A2A02FFDA280DFDCFFEB58498E031" box="[638,717,304,330]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587541" httpUri="https://zenodo.org/record/13587541/files/figure.png" pageId="6">Fig. 2</figureCitation>
|
||||||
): same data as holotype, Night Collecting.
|
): same data as holotype, Night Collecting.
|
||||||
<materialsCitation id="08D93CDAFFDA280DFB45FEB585B9E0C8" collectingDate="2019-05-27" collectingDateMax="2019-05-28" collectingDateMin="2019-05-27" collectionCode="UniFI" collectorName="L. Bartolozzi & S. Bambi & A. Bandinelli" country="Vietnam" elevation="1110" latitude="14.520278" location="Gia Lai Province" longLatPrecision="21" longitude="108.47" pageId="6" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="paratype">
|
<materialsCitation id="08D93CDAFFDA280DFB45FEB585B9E0C8" ID-GBIF-Occurrence="4926423302" collectingDate="2019-05-27" collectingDateMax="2019-05-28" collectingDateMin="2019-05-27" collectionCode="UniFI" collectorName="L. Bartolozzi & S. Bambi & A. Bandinelli" country="Vietnam" elevation="1110" latitude="14.520278" location="Gia Lai Province" longLatPrecision="21" longitude="108.47" pageId="6" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="paratype">
|
||||||
<typeStatus id="670A8825FFDA280DFB45FEB5833CE031" box="[1268,1385,304,330]" pageId="6" type="paratype">Paratypes</typeStatus>
|
<typeStatus id="670A8825FFDA280DFB45FEB5833CE031" box="[1268,1385,304,330]" pageId="6" type="paratype">Paratypes</typeStatus>
|
||||||
(
|
(
|
||||||
<specimenCount id="AEB7FD0EFFDA280DFEB0FED68717E016" box="[257,322,339,365]" count="2" pageId="6" type="male">2♂♂</specimenCount>
|
<specimenCount id="AEB7FD0EFFDA280DFEB0FED68717E016" box="[257,322,339,365]" count="2" pageId="6" type="male">2♂♂</specimenCount>
|
||||||
|
|
@ -186,7 +188,7 @@ asl
|
||||||
<specimenCount id="AEB7FD0EFFDA280DFC1CFE1C85B9E0C8" box="[941,1004,409,435]" count="4" pageId="6" type="female">4♀♀</specimenCount>
|
<specimenCount id="AEB7FD0EFFDA280DFC1CFE1C85B9E0C8" box="[941,1004,409,435]" count="4" pageId="6" type="female">4♀♀</specimenCount>
|
||||||
</materialsCitation>
|
</materialsCitation>
|
||||||
:
|
:
|
||||||
<materialsCitation id="08D93CDAFFDA280DFC4BFE1C836FE081" collectingDate="1910-01" collectingDateMax="2011-07-10" collectingDateMin="1910-01" collectionCode="R, RBINS" collectorName="J. Constant & J. Bresseel" country="Vietnam" latitude="16.616667" location="Quang Tri Province" longLatPrecision="1281" longitude="106.78333" pageId="6" specimenCount="1" stateProvince="Quang Tri" typeStatus="paratype">
|
<materialsCitation id="08D93CDAFFDA280DFC4BFE1C836FE081" ID-GBIF-Occurrence="4926423305" collectingDate="1910-01" collectingDateMax="2011-07-10" collectingDateMin="1910-01" collectionCode="R, RBINS" collectorName="J. Constant & J. Bresseel" country="Vietnam" latitude="16.616667" location="Quang Tri Province" longLatPrecision="1281" longitude="106.78333" pageId="6" specimenCount="1" stateProvince="Quang Tri" typeStatus="paratype">
|
||||||
<collectingCountry id="C0A67617FFDA280DFC4BFE1C8230E0C8" box="[1018,1125,409,435]" name="Vietnam" pageId="6">Vietnam</collectingCountry>
|
<collectingCountry id="C0A67617FFDA280DFC4BFE1C8230E0C8" box="[1018,1125,409,435]" name="Vietnam" pageId="6">Vietnam</collectingCountry>
|
||||||
,
|
,
|
||||||
<collectingRegion id="7A75F865FFDA280DFBC0FE1C8331E0C8" box="[1137,1380,409,435]" country="Vietnam" name="Quang Tri" pageId="6">Quang Tri Province</collectingRegion>
|
<collectingRegion id="7A75F865FFDA280DFBC0FE1C8331E0C8" box="[1137,1380,409,435]" country="Vietnam" name="Quang Tri" pageId="6">Quang Tri Province</collectingRegion>
|
||||||
|
|
@ -224,7 +226,7 @@ prep.
|
||||||
.
|
.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
</subSubSection>
|
</subSubSection>
|
||||||
<caption id="ECCE660FFFDA280DFF46FB688254E45A" pageId="6" startId="6.[247,284,1261,1283]" targetBox="[254,1376,536,1218]" targetPageId="6" targetType="figure">
|
<caption id="ECCE660FFFDA280DFF46FB688254E45A" ID-DOI="http://doi.org/10.5281/zenodo.13587537" ID-Zenodo-Dep="13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="6" startId="6.[247,284,1261,1283]" targetBox="[254,1376,536,1218]" targetPageId="6" targetType="figure">
|
||||||
<paragraph id="B80E3687FFDA280DFF46FB688254E45A" blockId="6.[247,1386,1261,1313]" pageId="6">
|
<paragraph id="B80E3687FFDA280DFF46FB688254E45A" blockId="6.[247,1386,1261,1313]" pageId="6">
|
||||||
Fig. 1.
|
Fig. 1.
|
||||||
<taxonomicName id="7FB14D04FFDA280DFE8FFB688415E479" authorityName="Vermeersch" authorityYear="2001" box="[318,576,1261,1282]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="6" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
<taxonomicName id="7FB14D04FFDA280DFE8FFB688415E479" authorityName="Vermeersch" authorityYear="2001" box="[318,576,1261,1282]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="6" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
||||||
|
|
@ -240,15 +242,15 @@ Fig. 1.
|
||||||
.
|
.
|
||||||
<smallCapsWord id="BEE8A05BFFDA280DFE19FAC687A4E420" baselines="1366,1366" box="[424,497,1347,1373]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Male" pageId="6">MALE</smallCapsWord>
|
<smallCapsWord id="BEE8A05BFFDA280DFE19FAC687A4E420" baselines="1366,1366" box="[424,497,1347,1373]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Male" pageId="6">MALE</smallCapsWord>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFDA280DFDB4FAC78401E427" box="[517,596,1346,1372]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Figs 1</figureCitation>
|
<figureCitation id="208A2A02FFDA280DFDB4FAC78401E427" box="[517,596,1346,1372]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="6">Figs 1</figureCitation>
|
||||||
,
|
,
|
||||||
<figureCitation id="208A2A02FFDA280DFDD7FAC78420E427" box="[614,629,1346,1372]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="6">3</figureCitation>
|
<figureCitation id="208A2A02FFDA280DFDD7FAC78420E427" box="[614,629,1346,1372]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." figureDoi="http://doi.org/10.5281/zenodo.13587547" httpUri="https://zenodo.org/record/13587547/files/figure.png" pageId="6">3</figureCitation>
|
||||||
A−C). Measurements (see
|
A−C). Measurements (see
|
||||||
<tableCitation id="F533033CFFDA280DFC7DFAC7820DE427" box="[972,1112,1346,1372]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="7.[247,304,421,443]" captionStartId-1="9.[247,304,316,338]" captionText-0="Table 1. Detailed measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. From left to right: T. attenboroughi holotype ♂, T. attenboroughi paratype ♀, T. fruhstorferi male, additional specimen 1, T. fruhstorferi female, additional specimen 2." captionText-1="Table 2. Measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. to assess intraspecific variability." pageId="6">Tables 1−2</tableCitation>
|
<tableCitation id="F533033CFFDA280DFC7DFAC7820DE427" box="[972,1112,1346,1372]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="7.[247,304,421,443]" captionStartId-1="9.[247,304,316,338]" captionTargetId-0="graphics-141@7.[282,1350,516,1675]" captionTargetId-1="graphics-59@9.[355,1278,381,1304]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="Table 1. Detailed measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. From left to right: T. attenboroughi holotype ♂, T. attenboroughi paratype ♀, T. fruhstorferi male, additional specimen 1, T. fruhstorferi female, additional specimen 2." captionText-1="Table 2. Measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. to assess intraspecific variability." httpUri-0="http://table.plazi.org/id/ECCE660FFFDB280CFF46FE208568E08C" httpUri-1="http://table.plazi.org/id/ECCE660FFFD52802FF46FEB9847EE00B" pageId="6" tableUuid-0="ECCE660FFFDB280CFF46FE208568E08C" tableUuid-1="ECCE660FFFD52802FF46FEB9847EE00B">Tables 1−2</tableCitation>
|
||||||
). Colouration (
|
). Colouration (
|
||||||
<figureCitation id="208A2A02FFDA280DFAA3FAC78337E427" box="[1298,1378,1346,1372]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Figs 1</figureCitation>
|
<figureCitation id="208A2A02FFDA280DFAA3FAC78337E427" box="[1298,1378,1346,1372]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="6">Figs 1</figureCitation>
|
||||||
,
|
,
|
||||||
<figureCitation id="208A2A02FFDA280DFF46FAE38751E4FB" box="[247,260,1382,1408]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="6">3</figureCitation>
|
<figureCitation id="208A2A02FFDA280DFF46FAE38751E4FB" box="[247,260,1382,1408]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." figureDoi="http://doi.org/10.5281/zenodo.13587547" httpUri="https://zenodo.org/record/13587547/files/figure.png" pageId="6">3</figureCitation>
|
||||||
A−C): All known male specimens of
|
A−C): All known male specimens of
|
||||||
<taxonomicName id="7FB14D04FFDA280DFD53FAE385E5E404" authorityName="Vermeersch" authorityYear="2001" box="[738,944,1381,1408]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="6" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
<taxonomicName id="7FB14D04FFDA280DFD53FAE385E5E404" authorityName="Vermeersch" authorityYear="2001" box="[738,944,1381,1408]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="6" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
||||||
<emphasis id="8AC5EA95FFDA280DFD53FAE384A6E404" box="[738,755,1382,1407]" italics="true" pageId="6">T</emphasis>
|
<emphasis id="8AC5EA95FFDA280DFD53FAE384A6E404" box="[738,755,1382,1407]" italics="true" pageId="6">T</emphasis>
|
||||||
|
|
@ -265,18 +267,18 @@ feature a uniformly green colouration without any variations in darkness or tone
|
||||||
<paragraph id="B80E3687FFDA280DFF46F90E830BE62E" blockId="6.[247,1386,1675,1877]" pageId="6">
|
<paragraph id="B80E3687FFDA280DFF46F90E830BE62E" blockId="6.[247,1386,1675,1877]" pageId="6">
|
||||||
<emphasis id="8AC5EA95FFDA280DFF46F90E876DE7DE" box="[247,312,1675,1701]" italics="true" pageId="6">Head</emphasis>
|
<emphasis id="8AC5EA95FFDA280DFF46F90E876DE7DE" box="[247,312,1675,1701]" italics="true" pageId="6">Head</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFDA280DFEFAF90E87E9E7DE" box="[331,444,1675,1701]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="6">Figs 1 C</figureCitation>
|
<figureCitation id="208A2A02FFDA280DFEFAF90E87E9E7DE" box="[331,444,1675,1701]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="6">Figs 1 C</figureCitation>
|
||||||
;
|
;
|
||||||
<figureCitation id="208A2A02FFDA280DFE7AF90E878FE7DE" box="[459,474,1675,1701]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="6">3</figureCitation>
|
<figureCitation id="208A2A02FFDA280DFE7AF90E878FE7DE" box="[459,474,1675,1701]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." figureDoi="http://doi.org/10.5281/zenodo.13587547" httpUri="https://zenodo.org/record/13587547/files/figure.png" pageId="6">3</figureCitation>
|
||||||
A−C): Wider than long with large, rounded compound eyes. Vertex flat. Ocellar tubercle not visible, in same plane as frons, without protruding ridges. Ocelli small, all about same size. Lower frons transverse with external margins posteriorly and laterally, no margin anteriorly, with two faintly defined vertical ridges internally that run from posterior end towards middle where they are abruptly interrupted. Two more clearly defined tubercules are placed in their extension and border anterior part of lower frons. Clypeus and labrum smooth.
|
A−C): Wider than long with large, rounded compound eyes. Vertex flat. Ocellar tubercle not visible, in same plane as frons, without protruding ridges. Ocelli small, all about same size. Lower frons transverse with external margins posteriorly and laterally, no margin anteriorly, with two faintly defined vertical ridges internally that run from posterior end towards middle where they are abruptly interrupted. Two more clearly defined tubercules are placed in their extension and border anterior part of lower frons. Clypeus and labrum smooth.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<paragraph id="B80E3687FFDB280CFF46FF3F8315E0FF" blockId="7.[247,1386,186,388]" pageId="7">
|
<paragraph id="B80E3687FFDB280CFF46FF3F8315E0FF" blockId="7.[247,1386,186,388]" pageId="7">
|
||||||
<emphasis id="8AC5EA95FFDB280CFF46FF3F871BE1AF" box="[247,334,186,212]" italics="true" pageId="7">Thorax</emphasis>
|
<emphasis id="8AC5EA95FFDB280CFF46FF3F871BE1AF" box="[247,334,186,212]" italics="true" pageId="7">Thorax</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFDB280CFED1FF3F87FAE1AF" box="[352,431,186,212]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="7">Figs 1</figureCitation>
|
<figureCitation id="208A2A02FFDB280CFED1FF3F87FAE1AF" box="[352,431,186,212]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="7">Figs 1</figureCitation>
|
||||||
A−B; 3 A−C): Pronotum long, straight and robust. Margins smooth, without denticulations or projections. Dorsal surface also entirely smooth, with a darker spot at each side laterally adjacent to the posterior margin of pronotum. Largest width before one third of pronotum, narrowest width located after middle of pronotum. Cervix with strongly sclerotized lateral and intercervical sclerites, merged in the middle. Postcervical plate and posterior ventral part of pronotum entirely smooth. Ventral lateral margins concolour with rest of pronotum.
|
A−B; 3 A−C): Pronotum long, straight and robust. Margins smooth, without denticulations or projections. Dorsal surface also entirely smooth, with a darker spot at each side laterally adjacent to the posterior margin of pronotum. Largest width before one third of pronotum, narrowest width located after middle of pronotum. Cervix with strongly sclerotized lateral and intercervical sclerites, merged in the middle. Postcervical plate and posterior ventral part of pronotum entirely smooth. Ventral lateral margins concolour with rest of pronotum.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<caption id="ECCE660FFFDB280CFF46FE208568E08C" pageId="7" startId="7.[247,304,421,443]" targetType="table">
|
<caption id="ECCE660FFFDB280CFF46FE208568E08C" ID-Table-UUID="ECCE660FFFDB280CFF46FE208568E08C" httpUri="http://table.plazi.org/id/ECCE660FFFDB280CFF46FE208568E08C" pageId="7" startId="7.[247,304,421,443]" targetBox="[332,1312,525,1661]" targetIsTable="true" targetPageId="7" targetType="table">
|
||||||
<paragraph id="B80E3687FFDB280CFF46FE208568E08C" blockId="7.[247,1386,421,504]" pageId="7">
|
<paragraph id="B80E3687FFDB280CFF46FE208568E08C" blockId="7.[247,1386,421,504]" pageId="7">
|
||||||
Table 1. Detailed measurements [mm] of
|
Table 1. Detailed measurements [mm] of
|
||||||
<taxonomicName id="7FB14D04FFDB280CFD17FE2085FFE0C1" authorityName="Vermeersch" authorityYear="2001" box="[678,938,421,442]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
<taxonomicName id="7FB14D04FFDB280CFD17FE2085FFE0C1" authorityName="Vermeersch" authorityYear="2001" box="[678,938,421,442]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="7" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
||||||
|
|
@ -523,7 +525,7 @@ AS
|
||||||
<paragraph id="B80E3687FFDB2803FF46F946845EE237" blockId="7.[247,1385,1731,1934]" lastBlockId="8.[247,1386,180,845]" lastPageId="8" pageId="7">
|
<paragraph id="B80E3687FFDB2803FF46F946845EE237" blockId="7.[247,1385,1731,1934]" lastBlockId="8.[247,1386,180,845]" lastPageId="8" pageId="7">
|
||||||
<emphasis id="8AC5EA95FFDB280CFF46F9468793E7A6" box="[247,454,1731,1757]" italics="true" pageId="7">Prothoracic legs</emphasis>
|
<emphasis id="8AC5EA95FFDB280CFF46F9468793E7A6" box="[247,454,1731,1757]" italics="true" pageId="7">Prothoracic legs</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFDB280CFE69F9468472E7A5" box="[472,551,1731,1758]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="7">Figs 1</figureCitation>
|
<figureCitation id="208A2A02FFDB280CFE69F9468472E7A5" box="[472,551,1731,1758]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="7">Figs 1</figureCitation>
|
||||||
A−B, D; 3 A−C): Coxa with 8−10 very small more or less blunt spinules of more or less equal size with same colour as coxa. Dorsal and ventral coxal lobes rounded, equal in length and adjacent, both lobes equally as broad. Black spot at proximal edge of ventral coxal lobe. Trochanter with black spot at anterior tip, adjacent matching black spot of same size on anterior side of femur. Femur with gently convex but straight dorsal margin, surface entirely smooth. Femoral brush ellipse-shaped, starting from 12
|
A−B, D; 3 A−C): Coxa with 8−10 very small more or less blunt spinules of more or less equal size with same colour as coxa. Dorsal and ventral coxal lobes rounded, equal in length and adjacent, both lobes equally as broad. Black spot at proximal edge of ventral coxal lobe. Trochanter with black spot at anterior tip, adjacent matching black spot of same size on anterior side of femur. Femur with gently convex but straight dorsal margin, surface entirely smooth. Femoral brush ellipse-shaped, starting from 12
|
||||||
<superScript id="4FC49BCFFFDB280CFBCFF8E882DBE6FB" attach="left" box="[1150,1166,1901,1920]" fontSize="8" pageId="7">th</superScript>
|
<superScript id="4FC49BCFFFDB280CFBCFF8E882DBE6FB" attach="left" box="[1150,1166,1901,1920]" fontSize="8" pageId="7">th</superScript>
|
||||||
or 13
|
or 13
|
||||||
|
|
@ -532,7 +534,7 @@ AvS and ending just a little distally of 15
|
||||||
<superScript id="4FC49BCFFFD42803FDC5FF3184D6E1BC" attach="left" box="[628,643,180,199]" fontSize="8" pageId="8">th</superScript>
|
<superScript id="4FC49BCFFFD42803FDC5FF3184D6E1BC" attach="left" box="[628,643,180,199]" fontSize="8" pageId="8">th</superScript>
|
||||||
AvS. Genicular lobes large and rounded, with minute, almost invisible genicular spur. Colouration of prothoracic legs green posteriorly, yellow anteriorly. Tibial spine groove proximally placed from middle of femur. AvS 1, 2, 10 and 15 black, black only along the distal and proximal edges or strongly apically infuscate (variable colouration between specimens) with black spot at base of AvS 2, 10 and 15 that slightly extends on anterior surface of femur; other AvS concolour with femur, only apically infuscate; AvS 10 and 12 large, but with 2 much smaller spines of equal size between them. Large and medium spines alternate between AvS 1 and 12, first spine being a medium one; medium spines subtly inclined towards anterior side of femur. PvS 4 smaller than others, PvS 1, 2 and 3 about equal in size; all concolour with femur and slightly apically infuscate. DS 1 and 4 more or less equal in size, DS 2 slightly longer than DS1, DS 3 longest and about 2x size of DS 2; DS 1 and 3 entirely black on anterior side (note: in some specimens only DS1 entirely black and DS3 apically infuscate), apically infuscate on posterior side, others apically infuscate on both sides. DS1 with black spot at base on anterior side. Protibiae smooth with longitudinal groove across entire length of posterior side. Tibial AvS gradually elongated towards tibial spur, all concolour to tibia but apically infuscate. PvS slightly more procumbent, spines gradually longer towards distal end. Tibial spur apically infuscate. Protarsi apically darkened posteriorly, entirely black anteriorly; first tarsomere longer than others combined. Spinal formula: F = 4DS/15AvS/4PvS; T = 13−14AvS/10PvS.
|
AvS. Genicular lobes large and rounded, with minute, almost invisible genicular spur. Colouration of prothoracic legs green posteriorly, yellow anteriorly. Tibial spine groove proximally placed from middle of femur. AvS 1, 2, 10 and 15 black, black only along the distal and proximal edges or strongly apically infuscate (variable colouration between specimens) with black spot at base of AvS 2, 10 and 15 that slightly extends on anterior surface of femur; other AvS concolour with femur, only apically infuscate; AvS 10 and 12 large, but with 2 much smaller spines of equal size between them. Large and medium spines alternate between AvS 1 and 12, first spine being a medium one; medium spines subtly inclined towards anterior side of femur. PvS 4 smaller than others, PvS 1, 2 and 3 about equal in size; all concolour with femur and slightly apically infuscate. DS 1 and 4 more or less equal in size, DS 2 slightly longer than DS1, DS 3 longest and about 2x size of DS 2; DS 1 and 3 entirely black on anterior side (note: in some specimens only DS1 entirely black and DS3 apically infuscate), apically infuscate on posterior side, others apically infuscate on both sides. DS1 with black spot at base on anterior side. Protibiae smooth with longitudinal groove across entire length of posterior side. Tibial AvS gradually elongated towards tibial spur, all concolour to tibia but apically infuscate. PvS slightly more procumbent, spines gradually longer towards distal end. Tibial spur apically infuscate. Protarsi apically darkened posteriorly, entirely black anteriorly; first tarsomere longer than others combined. Spinal formula: F = 4DS/15AvS/4PvS; T = 13−14AvS/10PvS.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<caption id="ECCE660FFFD42803FF46FA36826AE49C" pageId="8" startId="8.[247,284,1459,1481]" targetBox="[256,1378,887,1429]" targetPageId="8" targetType="figure">
|
<caption id="ECCE660FFFD42803FF46FA36826AE49C" ID-DOI="http://doi.org/10.5281/zenodo.13587541" ID-Zenodo-Dep="13587541" httpUri="https://zenodo.org/record/13587541/files/figure.png" pageId="8" startId="8.[247,284,1459,1481]" targetBox="[256,1378,887,1429]" targetPageId="8" targetType="figure">
|
||||||
<paragraph id="B80E3687FFD42803FF46FA36826AE49C" blockId="8.[247,1385,1459,1511]" pageId="8">
|
<paragraph id="B80E3687FFD42803FF46FA36826AE49C" blockId="8.[247,1385,1459,1511]" pageId="8">
|
||||||
Fig. 2.
|
Fig. 2.
|
||||||
<taxonomicName id="7FB14D04FFD42803FE8CFA36846AE4B3" authorityName="Vermeersch" authorityYear="2001" box="[317,575,1459,1480]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="8" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
<taxonomicName id="7FB14D04FFD42803FE8CFA36846AE4B3" authorityName="Vermeersch" authorityYear="2001" box="[317,575,1459,1480]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="8" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
||||||
|
|
@ -545,22 +547,22 @@ Fig. 2.
|
||||||
<paragraph id="B80E3687FFD42803FF46F98C8592E7D4" blockId="8.[247,1385,1545,1712]" pageId="8">
|
<paragraph id="B80E3687FFD42803FF46F98C8592E7D4" blockId="8.[247,1385,1545,1712]" pageId="8">
|
||||||
<emphasis id="8AC5EA95FFD42803FF46F98C8409E758" box="[247,604,1545,1571]" italics="true" pageId="8">Meso- and metathoracic legs</emphasis>
|
<emphasis id="8AC5EA95FFD42803FF46F98C8409E758" box="[247,604,1545,1571]" italics="true" pageId="8">Meso- and metathoracic legs</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD42803FDDDF98C84EFE758" box="[620,698,1545,1571]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="8">Figs 1</figureCitation>
|
<figureCitation id="208A2A02FFD42803FDDDF98C84EFE758" box="[620,698,1545,1571]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="8">Figs 1</figureCitation>
|
||||||
A−B; 3 A−C): Long and slender, without dilatations or projections. Femora with rounded genicular lobes and a short genicular spur. Tibiae with two apical spurs. Tarsi 5-segmented, green and slightly darkened apically. First tarsomere of mesotarsus shorter than remaining segments combined, first tarsomere of metatarsus about equal in length compared to remaining segments combined.
|
A−B; 3 A−C): Long and slender, without dilatations or projections. Femora with rounded genicular lobes and a short genicular spur. Tibiae with two apical spurs. Tarsi 5-segmented, green and slightly darkened apically. First tarsomere of mesotarsus shorter than remaining segments combined, first tarsomere of metatarsus about equal in length compared to remaining segments combined.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<paragraph id="B80E3687FFD42803FF46F94385D9E633" blockId="8.[247,1386,1733,1864]" pageId="8">
|
<paragraph id="B80E3687FFD42803FF46F94385D9E633" blockId="8.[247,1386,1733,1864]" pageId="8">
|
||||||
<emphasis id="8AC5EA95FFD42803FF46F9438781E7A4" box="[247,468,1733,1759]" italics="true" pageId="8">Tegmina and alae</emphasis>
|
<emphasis id="8AC5EA95FFD42803FF46F9438781E7A4" box="[247,468,1733,1759]" italics="true" pageId="8">Tegmina and alae</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD42803FE55F9408467E7A4" box="[484,562,1733,1759]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="8">Figs 1</figureCitation>
|
<figureCitation id="208A2A02FFD42803FE55F9408467E7A4" box="[484,562,1733,1759]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="8">Figs 1</figureCitation>
|
||||||
A−B; 3 A−C): Tegmina uniformly green on costal area, hyaline in discoidal area, longer than tip of abdomen when folded in rest. Veins green, stigma white with strong hue of yellowish-green. Stigma located anteriorly from middle of tegmen. Alae fully developed, hyaline with only greenish anterior margin.
|
A−B; 3 A−C): Tegmina uniformly green on costal area, hyaline in discoidal area, longer than tip of abdomen when folded in rest. Veins green, stigma white with strong hue of yellowish-green. Stigma located anteriorly from middle of tegmen. Alae fully developed, hyaline with only greenish anterior margin.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<paragraph id="B80E3687FFD52802FF46FF3F845EE061" blockId="9.[247,1386,186,283]" pageId="9">
|
<paragraph id="B80E3687FFD52802FF46FF3F845EE061" blockId="9.[247,1386,186,283]" pageId="9">
|
||||||
<emphasis id="8AC5EA95FFD52802FF46FF3F873CE1AF" box="[247,361,186,212]" italics="true" pageId="9">Abdomen</emphasis>
|
<emphasis id="8AC5EA95FFD52802FF46FF3F873CE1AF" box="[247,361,186,212]" italics="true" pageId="9">Abdomen</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD52802FEC8FF3F8794E1AF" box="[377,449,186,212]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="9">Fig. 1</figureCitation>
|
<figureCitation id="208A2A02FFD52802FEC8FF3F8794E1AF" box="[377,449,186,212]" captionStart="Fig" captionStartId="6.[247,284,1261,1283]" captionTargetBox="[254,1376,536,1218]" captionTargetId="figure-400@6.[247,1385,529,1227]" captionTargetPageId="6" captionText="Fig. 1. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587537" httpUri="https://zenodo.org/record/13587537/files/figure.png" pageId="9">Fig. 1</figureCitation>
|
||||||
A−B): Fusiform, uniformly green ventrally, yellowish-green dorsally. Cerci setose, not flattened, with 22−23 segments, proximal segments being merged together and difficult to distinguish.
|
A−B): Fusiform, uniformly green ventrally, yellowish-green dorsally. Cerci setose, not flattened, with 22−23 segments, proximal segments being merged together and difficult to distinguish.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<caption id="ECCE660FFFD52802FF46FEB9847EE00B" pageId="9" startId="9.[247,304,316,338]" targetType="table">
|
<caption id="ECCE660FFFD52802FF46FEB9847EE00B" ID-Table-UUID="ECCE660FFFD52802FF46FEB9847EE00B" httpUri="http://table.plazi.org/id/ECCE660FFFD52802FF46FEB9847EE00B" pageId="9" startId="9.[247,304,316,338]" targetBox="[404,1262,388,1290]" targetIsTable="true" targetPageId="9" targetType="table">
|
||||||
<paragraph id="B80E3687FFD52802FF46FEB9847EE00B" blockId="9.[247,1385,316,368]" pageId="9">
|
<paragraph id="B80E3687FFD52802FF46FEB9847EE00B" blockId="9.[247,1385,316,368]" pageId="9">
|
||||||
Table 2. Measurements [mm] of
|
Table 2. Measurements [mm] of
|
||||||
<taxonomicName id="7FB14D04FFD52802FDFEFEB98500E02A" authorityName="Vermeersch" authorityYear="2001" box="[591,853,316,337]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
<taxonomicName id="7FB14D04FFD52802FDFEFEB98500E02A" authorityName="Vermeersch" authorityYear="2001" box="[591,853,316,337]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="9" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
|
||||||
|
|
@ -779,10 +781,10 @@ to assess intraspecific variability.
|
||||||
<paragraph id="B80E3687FFD52801FF46FAD582E1E18C" blockId="9.[247,1386,1360,1878]" lastBlockId="10.[247,1385,186,248]" lastPageId="10" pageId="9">
|
<paragraph id="B80E3687FFD52801FF46FAD582E1E18C" blockId="9.[247,1386,1360,1878]" lastBlockId="10.[247,1385,186,248]" lastPageId="10" pageId="9">
|
||||||
<emphasis id="8AC5EA95FFD52802FF46FAD587F3E411" box="[247,422,1360,1386]" italics="true" pageId="9">Male genitalia</emphasis>
|
<emphasis id="8AC5EA95FFD52802FF46FAD587F3E411" box="[247,422,1360,1386]" italics="true" pageId="9">Male genitalia</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD52802FE04FAD587A9E410" box="[437,508,1360,1387]" captionStart="Fig" captionStartId="12.[247,284,1798,1820]" captionTargetBox="[354,1288,185,1775]" captionTargetId="figure-74@12.[330,1303,178,1778]" captionTargetPageId="12" captionText="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." pageId="9">Fig. 5</figureCitation>
|
<figureCitation id="208A2A02FFD52802FE04FAD587A9E410" box="[437,508,1360,1387]" captionStart="Fig" captionStartId="12.[247,284,1798,1820]" captionTargetBox="[354,1288,185,1775]" captionTargetId="figure-74@12.[330,1303,178,1778]" captionTargetPageId="12" captionText="Fig. 5. Titanodula fruhstorferi comb. nov., additional specimen ♂ from Cuc Phuong N.P. (RBINS). A, Genitalia prep. Vermeersch Nr. GEN1839, left phallic complex, dorsal view. Titanodula attenboroughi sp. nov., holotype ♂ (RBINS). B, genitalia prep. Vermeersch Nr. GEN1840, left phallic complex, dorsal view. C, right phallic complex, ventral view. D, coxosternite IX (subgenital plate), dorsal view. Drawings © Mado Berthet." figureDoi="http://doi.org/10.5281/zenodo.13587555" httpUri="https://zenodo.org/record/13587555/files/figure.png" pageId="9">Fig. 5</figureCitation>
|
||||||
B−D): Ventral phallomere of left phallic complex elongated, much longer than broad, with two short and strongly sclerotised distal processes (sdp) of about the same length with acute apex. Median process (sdpm) bend at the middle and pointing sideways, lateral process (sdpl) much more strongly bend and pointing upwards, with broader base than sdpm. Apical process of left phallomere (paa) complex in shape, with sclerotised apex and sclerotizations near the base, but membranous in the middle. Apex abruptly projecting dorsally and to the left, the apex sharply curved in a 90° hook with narrower but rounded tip. Phalloid apophysis (afa) with two sclerotised processes, anterior process (aafa) small and tubercle-shaped, broad and dome-like at the base with a smaller rounded projection on top of it, located posteriorly from the middle. Posterior process (pafa) spear-shaped, almost straight from the base onwards, long and heavily sclerotised, projecting straight or slightly diagonally posteriad. Membranous lobe loa rounded and small. Genital lobe goa strongly developed, extending laterally over its whole length. Right phallomere with sclerite R1A more or less triangular. Sclerite R3 shovel-shaped, very wide at its anterior end. Ventral process (pia) triangular with many longitudinal grooves on its surface, sclerified ventral process (pva) strongly sclerotized, finger-shaped, ending in a blunt hook. Coxosternite IX (subgenital plate) with small black spikes on the right lateral edge, sometimes also in the middle of the distal edge.
|
B−D): Ventral phallomere of left phallic complex elongated, much longer than broad, with two short and strongly sclerotised distal processes (sdp) of about the same length with acute apex. Median process (sdpm) bend at the middle and pointing sideways, lateral process (sdpl) much more strongly bend and pointing upwards, with broader base than sdpm. Apical process of left phallomere (paa) complex in shape, with sclerotised apex and sclerotizations near the base, but membranous in the middle. Apex abruptly projecting dorsally and to the left, the apex sharply curved in a 90° hook with narrower but rounded tip. Phalloid apophysis (afa) with two sclerotised processes, anterior process (aafa) small and tubercle-shaped, broad and dome-like at the base with a smaller rounded projection on top of it, located posteriorly from the middle. Posterior process (pafa) spear-shaped, almost straight from the base onwards, long and heavily sclerotised, projecting straight or slightly diagonally posteriad. Membranous lobe loa rounded and small. Genital lobe goa strongly developed, extending laterally over its whole length. Right phallomere with sclerite R1A more or less triangular. Sclerite R3 shovel-shaped, very wide at its anterior end. Ventral process (pia) triangular with many longitudinal grooves on its surface, sclerified ventral process (pva) strongly sclerotized, finger-shaped, ending in a blunt hook. Coxosternite IX (subgenital plate) with small black spikes on the right lateral edge, sometimes also in the middle of the distal edge.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<caption id="ECCE660FFFD62801FF46FA128482E4B1" pageId="10" startId="10.[247,284,1431,1453]" targetBox="[247,1385,273,1410]" targetPageId="10" targetType="figure">
|
<caption id="ECCE660FFFD62801FF46FA128482E4B1" ID-DOI="http://doi.org/10.5281/zenodo.13587547" ID-Zenodo-Dep="13587547" httpUri="https://zenodo.org/record/13587547/files/figure.png" pageId="10" startId="10.[247,284,1431,1453]" targetBox="[247,1385,273,1410]" targetPageId="10" targetType="figure">
|
||||||
<paragraph id="B80E3687FFD62801FF46FA128482E4B1" blockId="10.[247,1385,1431,1483]" pageId="10">
|
<paragraph id="B80E3687FFD62801FF46FA128482E4B1" blockId="10.[247,1385,1431,1483]" pageId="10">
|
||||||
Fig. 3.
|
Fig. 3.
|
||||||
<taxonomicName id="7FB14D04FFD62801FEF1FA128416E4D7" authorityName="Vermeersch" authorityYear="2001" box="[320,579,1431,1452]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="10" phylum="Arthropoda" rank="species" species="attenboroughi" status="gen. nov. et sp. nov.">
|
<taxonomicName id="7FB14D04FFD62801FEF1FA128416E4D7" authorityName="Vermeersch" authorityYear="2001" box="[320,579,1431,1452]" class="Insecta" family="Mantidae" genus="Titanodula" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Mantodea" pageId="10" phylum="Arthropoda" rank="species" species="attenboroughi" status="gen. nov. et sp. nov.">
|
||||||
|
|
@ -797,37 +799,37 @@ Fig. 3.
|
||||||
<paragraph id="B80E3687FFD62801FF46FA6887D3E752" blockId="10.[247,1385,1516,1577]" pageId="10">
|
<paragraph id="B80E3687FFD62801FF46FA6887D3E752" blockId="10.[247,1385,1516,1577]" pageId="10">
|
||||||
<smallCapsWord id="BEE8A05BFFD62801FF46FA68870FE77E" baselines="1536,1536" box="[247,346,1517,1543]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Female" pageId="10">FEMALE</smallCapsWord>
|
<smallCapsWord id="BEE8A05BFFD62801FF46FA68870FE77E" baselines="1536,1536" box="[247,346,1517,1543]" lowerCaseFontSize="9" mainFontSize="11" normCase="title" normString="Female" pageId="10">FEMALE</smallCapsWord>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD62801FEDFFA6987EAE77D" box="[366,447,1516,1542]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="10">Figs 2</figureCitation>
|
<figureCitation id="208A2A02FFD62801FEDFFA6987EAE77D" box="[366,447,1516,1542]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587541" httpUri="https://zenodo.org/record/13587541/files/figure.png" pageId="10">Figs 2</figureCitation>
|
||||||
,
|
,
|
||||||
<figureCitation id="208A2A02FFD62801FE61FA69878AE77D" box="[464,479,1516,1542]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="10">3</figureCitation>
|
<figureCitation id="208A2A02FFD62801FE61FA69878AE77D" box="[464,479,1516,1542]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." figureDoi="http://doi.org/10.5281/zenodo.13587547" httpUri="https://zenodo.org/record/13587547/files/figure.png" pageId="10">3</figureCitation>
|
||||||
D−E). Measurements (see
|
D−E). Measurements (see
|
||||||
<tableCitation id="F533033CFFD62801FC86FA698590E77D" box="[823,965,1516,1542]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="7.[247,304,421,443]" captionStartId-1="9.[247,304,316,338]" captionText-0="Table 1. Detailed measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. From left to right: T. attenboroughi holotype ♂, T. attenboroughi paratype ♀, T. fruhstorferi male, additional specimen 1, T. fruhstorferi female, additional specimen 2." captionText-1="Table 2. Measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. to assess intraspecific variability." pageId="10">Tables 1−2</tableCitation>
|
<tableCitation id="F533033CFFD62801FC86FA698590E77D" box="[823,965,1516,1542]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="7.[247,304,421,443]" captionStartId-1="9.[247,304,316,338]" captionTargetId-0="graphics-141@7.[282,1350,516,1675]" captionTargetId-1="graphics-59@9.[355,1278,381,1304]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="Table 1. Detailed measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. From left to right: T. attenboroughi holotype ♂, T. attenboroughi paratype ♀, T. fruhstorferi male, additional specimen 1, T. fruhstorferi female, additional specimen 2." captionText-1="Table 2. Measurements [mm] of Titanodula attenboroughi sp. nov. and Titanodula fruhstorferi comb. nov. to assess intraspecific variability." httpUri-0="http://table.plazi.org/id/ECCE660FFFDB280CFF46FE208568E08C" httpUri-1="http://table.plazi.org/id/ECCE660FFFD52802FF46FEB9847EE00B" pageId="10" tableUuid-0="ECCE660FFFDB280CFF46FE208568E08C" tableUuid-1="ECCE660FFFD52802FF46FEB9847EE00B">Tables 1−2</tableCitation>
|
||||||
). Similar to male, with following differences:
|
). Similar to male, with following differences:
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<paragraph id="B80E3687FFD62801FF46F9BA8593E707" blockId="10.[247,1385,1599,1660]" pageId="10">
|
<paragraph id="B80E3687FFD62801FF46F9BA8593E707" blockId="10.[247,1385,1599,1660]" pageId="10">
|
||||||
<emphasis id="8AC5EA95FFD62801FF46F9BA871BE722" box="[247,334,1599,1625]" italics="true" pageId="10">Thorax</emphasis>
|
<emphasis id="8AC5EA95FFD62801FF46F9BA871BE722" box="[247,334,1599,1625]" italics="true" pageId="10">Thorax</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD62801FEEEF9BA87F9E721" box="[351,428,1599,1626]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="10">Figs 2</figureCitation>
|
<figureCitation id="208A2A02FFD62801FEEEF9BA87F9E721" box="[351,428,1599,1626]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587541" httpUri="https://zenodo.org/record/13587541/files/figure.png" pageId="10">Figs 2</figureCitation>
|
||||||
A−B, 3 D−E): Margins of pronotum finely denticulated anteriorly, fading out after pronotal supracoxal dilatation, absent in posterior end.
|
A−B, 3 D−E): Margins of pronotum finely denticulated anteriorly, fading out after pronotal supracoxal dilatation, absent in posterior end.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<paragraph id="B80E3687FFD62801FF46F91784CEE789" blockId="10.[247,1385,1682,1779]" pageId="10">
|
<paragraph id="B80E3687FFD62801FF46F91784CEE789" blockId="10.[247,1385,1682,1779]" pageId="10">
|
||||||
<emphasis id="8AC5EA95FFD62801FF46F917879CE7D7" box="[247,457,1682,1708]" italics="true" pageId="10">Prothoracic legs</emphasis>
|
<emphasis id="8AC5EA95FFD62801FF46F917879CE7D7" box="[247,457,1682,1708]" italics="true" pageId="10">Prothoracic legs</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD62801FE6CF9178465E7D7" box="[477,560,1682,1708]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="10">Figs 2</figureCitation>
|
<figureCitation id="208A2A02FFD62801FE6CF9178465E7D7" box="[477,560,1682,1708]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587541" httpUri="https://zenodo.org/record/13587541/files/figure.png" pageId="10">Figs 2</figureCitation>
|
||||||
A−B, D, 3 D−E): Coxae with 8−10 small sharp and regularly interspaced spinules of more or less equal size with same colour as coxa, spinules larger and more pronounced than in the male.
|
A−B, D, 3 D−E): Coxae with 8−10 small sharp and regularly interspaced spinules of more or less equal size with same colour as coxa, spinules larger and more pronounced than in the male.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<paragraph id="B80E3687FFD62801FF46F88C84A2E6F0" blockId="10.[247,1385,1800,1932]" pageId="10">
|
<paragraph id="B80E3687FFD62801FF46F88C84A2E6F0" blockId="10.[247,1385,1800,1932]" pageId="10">
|
||||||
<emphasis id="8AC5EA95FFD62801FF46F88C879AE659" box="[247,463,1800,1826]" italics="true" pageId="10">Tegmina and alae</emphasis>
|
<emphasis id="8AC5EA95FFD62801FF46F88C879AE659" box="[247,463,1800,1826]" italics="true" pageId="10">Tegmina and alae</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD62801FE6DF88D8472E659" box="[476,551,1800,1826]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="10">Figs 2</figureCitation>
|
<figureCitation id="208A2A02FFD62801FE6DF88D8472E659" box="[476,551,1800,1826]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587541" httpUri="https://zenodo.org/record/13587541/files/figure.png" pageId="10">Figs 2</figureCitation>
|
||||||
A−B, 3 D−E): Tegmina uniformly green in costal area, partly hyaline in discoidal area, being green anteriorly, then quickly becoming hyaline in a gradient posteriorly. Tegmina reaching tip of abdomen when folded in rest. Alae fully developed, hyaline with faintly green anterior margin.
|
A−B, 3 D−E): Tegmina uniformly green in costal area, partly hyaline in discoidal area, being green anteriorly, then quickly becoming hyaline in a gradient posteriorly. Tegmina reaching tip of abdomen when folded in rest. Alae fully developed, hyaline with faintly green anterior margin.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<paragraph id="B80E3687FFD72800FF46FF3F8218E18C" blockId="11.[247,1385,186,248]" pageId="11">
|
<paragraph id="B80E3687FFD72800FF46FF3F8218E18C" blockId="11.[247,1385,186,248]" pageId="11">
|
||||||
<emphasis id="8AC5EA95FFD72800FF46FF3F873CE1AF" box="[247,361,186,212]" italics="true" pageId="11">Abdomen</emphasis>
|
<emphasis id="8AC5EA95FFD72800FF46FF3F873CE1AF" box="[247,361,186,212]" italics="true" pageId="11">Abdomen</emphasis>
|
||||||
(
|
(
|
||||||
<figureCitation id="208A2A02FFD72800FEC8FF3F87B0E1AF" box="[377,485,186,212]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." pageId="11">Figs 2 A</figureCitation>
|
<figureCitation id="208A2A02FFD72800FEC8FF3F87B0E1AF" box="[377,485,186,212]" captionStart="Fig" captionStartId="8.[247,284,1459,1481]" captionTargetBox="[256,1378,887,1429]" captionTargetId="figure-458@8.[247,1385,870,1438]" captionTargetPageId="8" captionText="Fig. 2. Titanodula attenboroughi sp. nov., paratype ♀, [PT1] (RBINS). A, habitus, dorsal view. B, habitus, ventral view. C, head, frontal view. D, left prothoracic leg, anterior view. C, D not to scale." figureDoi="http://doi.org/10.5281/zenodo.13587541" httpUri="https://zenodo.org/record/13587541/files/figure.png" pageId="11">Figs 2 A</figureCitation>
|
||||||
,
|
,
|
||||||
<figureCitation id="208A2A02FFD72800FE43FF3F8476E1AF" box="[498,547,186,212]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." pageId="11">3 D</figureCitation>
|
<figureCitation id="208A2A02FFD72800FE43FF3F8476E1AF" box="[498,547,186,212]" captionStart="Fig" captionStartId="10.[247,284,1431,1453]" captionTargetBox="[247,1385,273,1410]" captionTargetId="figure-209@10.[247,1385,273,1410]" captionTargetPageId="10" captionText="Fig. 3. Titanodula attenboroughi gen. nov. et sp. nov., live photographs of holotype ♂ (A, B, C) and paratype ♀ [PT1] (D, E) at Kon Chu Rang N.R., VII. 2018." figureDoi="http://doi.org/10.5281/zenodo.13587547" httpUri="https://zenodo.org/record/13587547/files/figure.png" pageId="11">3 D</figureCitation>
|
||||||
): Elongated ovaliform, much broader than in male, uniformly green ventrally, bright yellowish dorsally with a hint of green near the edges.
|
): Elongated ovaliform, much broader than in male, uniformly green ventrally, bright yellowish dorsally with a hint of green near the edges.
|
||||||
</paragraph>
|
</paragraph>
|
||||||
<paragraph id="B80E3687FFD72800FF46FE88840FE0AD" blockId="11.[247,1385,269,470]" pageId="11">
|
<paragraph id="B80E3687FFD72800FF46FE88840FE0AD" blockId="11.[247,1385,269,470]" pageId="11">
|
||||||
|
|
|
||||||
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Reference in a new issue