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<mods:namePart>Liu, Weixin</mods:namePart>
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<taxonomicName LSID="E483ACABE7A9F291D70E273157526E29" class="Diplopoda" family="Glomeridae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Glomerida" pageId="22" pageNumber="79" phylum="Arthropoda" rank="species" species="maocun">
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<pageBreakToken id="98983050CB455E7998B759D162268A53" pageId="22" pageNumber="79" start="start">Glyphiulus</pageBreakToken>
maocun
</taxonomicName>
<taxonomicNameLabel pageId="22" pageNumber="79">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Trichopeltis liangfengdong sp. nov. from Liangfeng Cave B Pacidesmus trifidus Golovatch &amp; Geoffroy, 2014 from Maomaotou Cave C Glyphiulus maocun sp. nov. from Liangfeng Cave." figureDoi="10.3897/subtbiol.30.35559.figure4" httpUri="https://binary.pensoft.net/fig/314552" pageId="22" pageNumber="79">Figs 4C</figureCitation>
<taxonomicNameLabel id="DED0BB0DFA32CDB95DDCBA707F025E1A" pageId="22" pageNumber="79">sp. nov.</taxonomicNameLabel>
<figureCitation id="E5F303CB8895223FC4C3965307FB3146" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Trichopeltis liangfengdong sp. nov. from Liangfeng Cave B Pacidesmus trifidus Golovatch &amp; Geoffroy, 2014 from Maomaotou Cave C Glyphiulus maocun sp. nov. from Liangfeng Cave." figureDoi="10.3897/subtbiol.30.35559.figure4" httpUri="https://binary.pensoft.net/fig/314552" pageId="22" pageNumber="79">Figs 4C</figureCitation>
,
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="22" pageNumber="79">18</figureCitation>
<figureCitation id="2D5026FE2D26BB25F6EE89EE408A906B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="22" pageNumber="79">18</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="22" pageNumber="79">19</figureCitation>
<figureCitation id="1C00D62F3E9AEC7B102B42A30B4BAF55" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="22" pageNumber="79">19</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="22" pageNumber="79">20</figureCitation>
<figureCitation id="99CFAA5E342FE40D90C323B45CE43764" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="22" pageNumber="79">20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="79" type="type material">
<paragraph pageId="22" pageNumber="79">Type material.</paragraph>
<paragraph pageId="22" pageNumber="79">
<subSubSection id="9C817B89293BA0B11B841906BB9F3EA1" pageId="22" pageNumber="79" type="type material">
<paragraph id="DA1076DD15F8E0F7D3028F6EC445FB74" pageId="22" pageNumber="79">Type material.</paragraph>
<paragraph id="33C248CD90293A072289C0D1158508EC" pageId="22" pageNumber="79">
Holotype male, China, Guangxi Zhuang Autonomous Region, Lingchuan County, Maocun Village, Liangfeng Cave [
<geoCoordinate degrees="25" direction="north" minutes="12" orientation="latitude" precision="1" seconds="34.86" value="25.209684">25°12'34.86&quot;N</geoCoordinate>
<geoCoordinate id="D4DF4FD2035A5A76E69460884986B329" degrees="25" direction="north" minutes="12" orientation="latitude" precision="1" seconds="34.86" value="25.209684">25°12'34.86&quot;N</geoCoordinate>
,
<geoCoordinate degrees="110" direction="east" minutes="31" orientation="longitude" precision="1" seconds="56.8" value="110.53245">110°31'56.8&quot;E</geoCoordinate>
<geoCoordinate id="AE3181DF5E830A356A7A7F55A3F9FBD3" degrees="110" direction="east" minutes="31" orientation="longitude" precision="1" seconds="56.8" value="110.53245">110°31'56.8&quot;E</geoCoordinate>
], 184 m el., deep zone, direct intuitive search, 18 November 2016, J.J. Wynne. Paratypes, 4 females, 4 juv. (SCAU), same data as holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="79" type="etymology">
<paragraph pageId="22" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="22" pageNumber="79">
<subSubSection id="8BE80E38FD9574D119DDEC79622DC417" pageId="22" pageNumber="79" type="etymology">
<paragraph id="2D1ED206484D9F7460CDE9A1AE36A67F" pageId="22" pageNumber="79">Etymology.</paragraph>
<paragraph id="C70BA3F2536408459B0310EAE82FCD47" pageId="22" pageNumber="79">
The species name,
<emphasis italics="true" pageId="22" pageNumber="79">maocun</emphasis>
<emphasis id="076AA7830F6F00683B07262441B97458" italics="true" pageId="22" pageNumber="79">maocun</emphasis>
, is used as a noun in apposition and is the name of the village nearest to the type locality.
</paragraph>
</subSubSection>
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<paragraph pageId="22" pageNumber="79">Diagnosis.</paragraph>
<paragraph lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79">
<subSubSection id="F2FF6E4F341A5F1318E63E5DFE6FC97B" lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79" type="diagnosis">
<paragraph id="10DD7456DFC4DD2BDA32154A29506F0E" pageId="22" pageNumber="79">Diagnosis.</paragraph>
<paragraph id="3354ACA48AF9C1558816B763B6865BC4" lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79">
This new species is similar to
<taxonomicName genus="G." lsidName="G. melanoporus" pageId="22" pageNumber="79" rank="species" species="melanoporus">
<emphasis italics="true" pageId="22" pageNumber="79">G. melanoporus</emphasis>
<taxonomicName id="145F9916CD295F81500B78FC601EDC76" lsidName="G. melanoporus" pageId="22" pageNumber="79" rank="species" species="melanoporus">
<emphasis id="8F7AB73310CF0598249B39CBF2A0605F" italics="true" pageId="22" pageNumber="79">G. melanoporus</emphasis>
</taxonomicName>
(from caves in Guilin) in showing carinotaxy formula of collum
<normalizedToken originalValue="IIV">I-IV</normalizedToken>
<normalizedToken id="637CC1D13B5D211440C50B383AD91123" originalValue="IIV">I-IV</normalizedToken>
+5c+6a+pc+ma. However,
<taxonomicName genus="G." lsidName="G. maocun" pageId="22" pageNumber="79" rank="species" species="maocun">
<emphasis italics="true" pageId="22" pageNumber="79">G. maocun</emphasis>
<taxonomicName id="06ACC0E22F4E03F86D4135780026E6D2" lsidName="G. maocun" pageId="22" pageNumber="79" rank="species" species="maocun">
<emphasis id="14E288A2B0CA7860ADAA937B3795B492" italics="true" pageId="22" pageNumber="79">G. maocun</emphasis>
</taxonomicName>
sp. nov. is distinct from
<taxonomicName genus="G." lsidName="G. melanoporus" pageId="22" pageNumber="79" rank="species" species="melanoporus">
<emphasis italics="true" pageId="22" pageNumber="79">G. melanoporus</emphasis>
<taxonomicName id="C4CD775DFB7DF66A6D4D6E0A1CE644B4" lsidName="G. melanoporus" pageId="22" pageNumber="79" rank="species" species="melanoporus">
<emphasis id="7072C62FCE556AD4FB800CF12373DAEA" italics="true" pageId="22" pageNumber="79">G. melanoporus</emphasis>
</taxonomicName>
by (1) carinotaxy formula of metaterga 2/2+I/
<pageBreakToken pageId="23" pageNumber="80" start="start">i</pageBreakToken>
<pageBreakToken id="2571A3367BFC1FD0DE6790B47DB9AA6A" pageId="23" pageNumber="80" start="start">i</pageBreakToken>
+3/3; (2) anterior gonopod with a conspicuous, high, curved downward process on coxosternum (
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<figureCitation id="9DBACCEBD5E6D946DA464D1355579867" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="23" pageNumber="80">Figs 19F</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="23" pageNumber="80">20A</figureCitation>
<figureCitation id="175D4CB107404DBD1E11E0F0FE33B756" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="23" pageNumber="80">20A</figureCitation>
); (3) posterior gonopod tip branch plumose flagellum (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="23" pageNumber="80">
Figs 19
<normalizedToken originalValue="GI">G-I</normalizedToken>
<figureCitation id="DB952167D08CCA00702ECD85127C08E1" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="23" pageNumber="80">
Figs 19
<normalizedToken id="03991747775870D0E1425367396A3BDC" originalValue="GI">G-I</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="23" pageNumber="80">20B, C</figureCitation>
<figureCitation id="61DC7656C42A093399E6BC52CEC7ED63" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="23" pageNumber="80">20B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="81" type="description">
<paragraph pageId="24" pageNumber="81">
<pageBreakToken pageId="24" pageNumber="81" start="start">Description</pageBreakToken>
<subSubSection id="270C710C335C0EEA64880F540ABFECE9" pageId="24" pageNumber="81" type="description">
<paragraph id="87E7AC4A837591FD05B579A1B3294666" pageId="24" pageNumber="81">
<pageBreakToken id="FCD41BBFB1F6E2698969317DAEA9E9E0" pageId="24" pageNumber="81" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="24" pageNumber="81">
<paragraph id="3EA693ADAD600620ED06C2A2AC2027B0" pageId="24" pageNumber="81">
Based on the type specimens. Lengths of both sexes
<emphasis italics="true" pageId="24" pageNumber="81">ca</emphasis>
<emphasis id="4899B060FB4A9580F9889639B2117393" italics="true" pageId="24" pageNumber="81">ca</emphasis>
26-32 mm, mid-body rings round in cross-section, their widths and heights similar, 1.2-1.8 mm.
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Coloration</emphasis>
<emphasis id="1EFACCED4952D74852DFAE9F916C53C1" bold="true" italics="true" pageId="24" pageNumber="81">Coloration</emphasis>
: yellow-brown to brownish in alcohol. Ommatidia blackish (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18A</figureCitation>
<figureCitation id="ADD1A66AEBD112C705D2151D6B88E7B6" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18A</figureCitation>
).
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Body</emphasis>
<emphasis id="11588B224E3907A31D83FA107B0AB2EF" bold="true" italics="true" pageId="24" pageNumber="81">Body</emphasis>
: with 46-54 podous rings + 2-1 apodous one(s) + telson.
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Head</emphasis>
<emphasis id="5915E89A13B6657B28A07EBE66816FAC" bold="true" italics="true" pageId="24" pageNumber="81">Head</emphasis>
: clypeus with 4 teeth anteromedially. Each eye patch with about 8-12 ommatidium arranged in three irregular vertical rows (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Figs 18A</figureCitation>
<figureCitation id="A477D7CCFA94F32554F22C0026D17A63" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Figs 18A</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">19A</figureCitation>
<figureCitation id="9BC5DECCE34794E454D46C537E511222" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">19A</figureCitation>
). Antennae long, antennomere 7 with four short apical cones (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18A, B</figureCitation>
<figureCitation id="8A7271AFB16F343903D4CDB9401FC865" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18A, B</figureCitation>
). Gnathochilarium usually, with a separate promentum (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19B</figureCitation>
<figureCitation id="9F864F553EDDE2B05D849052A308E165" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19B</figureCitation>
).
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Exoskeleton</emphasis>
<emphasis id="AE544FA7DC2A907C7CE96CF7264C2A65" bold="true" italics="true" pageId="24" pageNumber="81">Exoskeleton</emphasis>
: Carinotaxy formula of collum
<normalizedToken originalValue="IIV">I-IV</normalizedToken>
<normalizedToken id="567E52501C555CF6289BDF0F78E48D9A" originalValue="IIV">I-IV</normalizedToken>
+5c+6a+pc+ma, anterior margin crests lower (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19A</figureCitation>
<figureCitation id="FB5B1AF19C5FE9B55317FC60BE582471" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19A</figureCitation>
). Subsequent metaterga strongly crested, carinotaxy formula 2/2+I/i+3/3. Ozoporiferous tubercles round (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18</figureCitation>
<figureCitation id="997A4E7B18980EE045ED161B706E082A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18</figureCitation>
). Prozonae delicately alveolate; strictures and metazonae fine longitudinal striations. Rings 2 and 3 with long pleural flaps. Epiproct simple, with a short, low, rounded tubercle medially. Paraprocts rather regularly convex. Hypoproct broadly emarginated (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18D</figureCitation>
<figureCitation id="8E50057C9A70CF2C1AF3D768CCA2E3F7" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18D</figureCitation>
). ♂ legs 1 very strongly reduced, represented only by a sternum devoid of any median or paramedian structures but carrying 1+1 strongly separated prongs both curved anteriad and bearing several strong setae (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18B, C</figureCitation>
<figureCitation id="E5321FE71D3C9CE616E51AF339A23A34" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18B, C</figureCitation>
). ♂ legs 2 with large coxae; penes oblong, each with two strong setae distolaterally (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19D</figureCitation>
<figureCitation id="A75C35E7E5D8FEA10390783C68DC036B" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19D</figureCitation>
). ♂ legs 3 with slender and elongate coxae (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19E</figureCitation>
<figureCitation id="8BF4DCB3F0CE0F271E1490AAA6D67A7F" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19E</figureCitation>
). Legs slender, about 1.2 times as long as mid-body ring height. Claw simple, without any spine at base.
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Anterior gonopods</emphasis>
<emphasis id="850515A1ABE7729F6C4B9046B42A64AC" bold="true" italics="true" pageId="24" pageNumber="81">Anterior gonopods</emphasis>
: (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Figs 19F</figureCitation>
<figureCitation id="D10EBC5D1977DE6C4E244DA00499E9E1" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Figs 19F</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="24" pageNumber="81">20A</figureCitation>
<figureCitation id="6C99DCF28ADEEC5A8068A8A118B7A1AE" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="24" pageNumber="81">20A</figureCitation>
) with a conspicuous, high, lobe-shaped, curved downward process on coxosternum, which is much higher than telopdite; telopodite slightly coiled, 1-segmented, lateral in position, with 2 or 3 strong apical setae.
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Posterior gonopods</emphasis>
<emphasis id="D56E4B254BA5E92A8345A03ACC555FE9" bold="true" italics="true" pageId="24" pageNumber="81">Posterior gonopods</emphasis>
: (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">
Figs 19
<normalizedToken originalValue="GI">G-I</normalizedToken>
<figureCitation id="BDABF6A87A1B337E264D8F89DA0DF9C4" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">
Figs 19
<normalizedToken id="30E561C2C1D05AED691B873D541AAE12" originalValue="GI">G-I</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="24" pageNumber="81">20B, C</figureCitation>
<figureCitation id="B3E5EBAC46BA44D2A7874D80821D0DEC" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="24" pageNumber="81">20B, C</figureCitation>
) laterally with a low, lateral lobe carrying a dozen of long setae, tip branch very slender and high, plumose flagellum.
</paragraph>
<caption doi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81" start="Figure 18" startId="F18">
<paragraph pageId="24" pageNumber="81">
<caption id="80D4492052CE587AE21BBDBF4372B8ED" doi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81" start="Figure 18" startId="F18">
<paragraph id="27563147FD1628C511BF4E1B69CB6882" pageId="24" pageNumber="81">
Figure 18.
<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Spirostreptida" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="maocun">
<emphasis italics="true" pageId="24" pageNumber="81">Glyphiulus maocun</emphasis>
<taxonomicName id="0BCA5997C966B5881A3BEFF03AB2161A" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Spirostreptida" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="maocun">
<emphasis id="F27FA142A193B649C58A06A4E0CA1237" italics="true" pageId="24" pageNumber="81">Glyphiulus maocun</emphasis>
</taxonomicName>
sp. nov., holotype.
<emphasis bold="true" pageId="24" pageNumber="81">A, B</emphasis>
<emphasis id="8DCF1023088D7AE043C72BC426521555" bold="true" pageId="24" pageNumber="81">A, B</emphasis>
Anterior part of body, sublateral views
<emphasis bold="true" pageId="24" pageNumber="81">C</emphasis>
<emphasis id="5B250195F5902646A23B0791F56EDD44" bold="true" pageId="24" pageNumber="81">C</emphasis>
mid-body rings, dorsal view
<emphasis bold="true" pageId="24" pageNumber="81">D</emphasis>
<emphasis id="F98EB333B4E195C0E9B1C8A8F2138C11" bold="true" pageId="24" pageNumber="81">D</emphasis>
posterior part of body, ventral view.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81" start="Figure 19" startId="F19">
<paragraph pageId="24" pageNumber="81">
<caption id="C7209A9124ACDF2B548AF3B0A9B89B82" doi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81" start="Figure 19" startId="F19">
<paragraph id="A2637C0639FBE7BF40AD4B44CDE81040" pageId="24" pageNumber="81">
Figure 19.
<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Spirostreptida" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="maocun">
<emphasis italics="true" pageId="24" pageNumber="81">Glyphiulus maocun</emphasis>
<taxonomicName id="81CE35B1E4541DD3C061121E54BC59BD" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Spirostreptida" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="maocun">
<emphasis id="B648028594831A9A973B37CD2CE9BE80" italics="true" pageId="24" pageNumber="81">Glyphiulus maocun</emphasis>
</taxonomicName>
sp. nov., holotype.
<emphasis bold="true" pageId="24" pageNumber="81">A</emphasis>
<emphasis id="5F3959891A60B015A72FF2162F9FD332" bold="true" pageId="24" pageNumber="81">A</emphasis>
Head and collum, dorsal view
<emphasis bold="true" pageId="24" pageNumber="81">B</emphasis>
<emphasis id="BB65C1F18CC5EF67AB4A45CA652118C6" bold="true" pageId="24" pageNumber="81">B</emphasis>
head, collum, and legs 1, ventral view
<emphasis bold="true" pageId="24" pageNumber="81">C</emphasis>
<emphasis id="AF02A599A425399E826D9C27049A7848" bold="true" pageId="24" pageNumber="81">C</emphasis>
legs 1, anterior view
<emphasis bold="true" pageId="24" pageNumber="81">D</emphasis>
<emphasis id="FB8F55333258CBF5955537FBFDC5015D" bold="true" pageId="24" pageNumber="81">D</emphasis>
leg 2, caudal view
<emphasis bold="true" pageId="24" pageNumber="81">E</emphasis>
<emphasis id="05F8BC43AA67DC8EDE8858266C2DB948" bold="true" pageId="24" pageNumber="81">E</emphasis>
legs 3, frontal view
<emphasis bold="true" pageId="24" pageNumber="81">F</emphasis>
<emphasis id="9F564D4D3962079CA3489C51315246BA" bold="true" pageId="24" pageNumber="81">F</emphasis>
anterior gonopods, caudal view
<emphasis bold="true" pageId="24" pageNumber="81">G</emphasis>
<emphasis id="C255AD5EFA19AC2191AC4D73A23479AC" bold="true" pageId="24" pageNumber="81">G</emphasis>
posterior gonopods, anterior view
<emphasis bold="true" pageId="24" pageNumber="81">H</emphasis>
<emphasis id="B64477C3C4B32120D4DE0310E93CB839" bold="true" pageId="24" pageNumber="81">H</emphasis>
posterior gonopods, caudal view.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="24" pageNumber="81" start="Figure 20" startId="F20">
<paragraph pageId="24" pageNumber="81">
<caption id="48B9B3125B2EEC5FF0765E3DFD94997F" doi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="24" pageNumber="81" start="Figure 20" startId="F20">
<paragraph id="0EEED072F3C463A97404D37250D33780" pageId="24" pageNumber="81">
Figure 20.
<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Spirostreptida" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="maocun">
<emphasis italics="true" pageId="24" pageNumber="81">Glyphiulus maocun</emphasis>
<taxonomicName id="F6F3566360DBF6B8131A28BA35A214B2" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Spirostreptida" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="maocun">
<emphasis id="DFAD4E2762306190D7B4BC10941E010E" italics="true" pageId="24" pageNumber="81">Glyphiulus maocun</emphasis>
</taxonomicName>
sp. nov., holotype.
<emphasis bold="true" pageId="24" pageNumber="81">A</emphasis>
<emphasis id="C708C9AD6C0140C3554884430AE57F16" bold="true" pageId="24" pageNumber="81">A</emphasis>
Anterior gonopods, caudal view
<emphasis bold="true" pageId="24" pageNumber="81">B</emphasis>
<emphasis id="45CC29EE480000CB79629894899D01F4" bold="true" pageId="24" pageNumber="81">B</emphasis>
posterior gonopods, anterior view
<emphasis bold="true" pageId="24" pageNumber="81">C</emphasis>
<emphasis id="83A2F9BB93A44D8C2407C5B7F25386B9" bold="true" pageId="24" pageNumber="81">C</emphasis>
posterior gonopods, caudal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="24" pageNumber="81" type="habitat">
<paragraph pageId="24" pageNumber="81">Habitat.</paragraph>
<paragraph pageId="24" pageNumber="81">Specimens were collected within or at proximity to flood detritus.</paragraph>
<subSubSection id="C1236ED3E9CBF205BDE3617200852675" pageId="24" pageNumber="81" type="habitat">
<paragraph id="BF390E0A1D1E5AB8BE8E827C1A1596D4" pageId="24" pageNumber="81">Habitat.</paragraph>
<paragraph id="13E1B032A45CEF177879DCF033E41127" pageId="24" pageNumber="81">Specimens were collected within or at proximity to flood detritus.</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="81" type="notes">
<paragraph pageId="24" pageNumber="81">Notes.</paragraph>
<paragraph pageId="24" pageNumber="81">Specimens exhibited no characteristics suggestive of cave adaptation. It had a pigmented cuticle and well-developed blackish ommatidia. We consider this species to be troglophillic within Liangfeng Cave. Subsequently, this species may have a larger, more regional distribution.</paragraph>
<subSubSection id="6182E459EF85D872F802E08557D75371" pageId="24" pageNumber="81" type="notes">
<paragraph id="61F78839A6F59F6506A01186ED374483" pageId="24" pageNumber="81">Notes.</paragraph>
<paragraph id="F62BAFF8155348D0A346D591247E0409" pageId="24" pageNumber="81">Specimens exhibited no characteristics suggestive of cave adaptation. It had a pigmented cuticle and well-developed blackish ommatidia. We consider this species to be troglophillic within Liangfeng Cave. Subsequently, this species may have a larger, more regional distribution.</paragraph>
</subSubSection>
</treatment>
</document>