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<mods:title>Cave millipede diversity with the description of six new species from Guangxi, China</mods:title>
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<mods:title id="BEB0DA9BDC963214AE932795982EDEAB">Cave millipede diversity with the description of six new species from Guangxi, China</mods:title>
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<mods:namePart>Liu, Weixin</mods:namePart>
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<treatment ID-GBIF-Taxon="158498357" LSID="urn:lsid:plazi:treatment:5126B5BADC2E8B7EA71E75AB48C43E1E" httpUri="http://treatment.plazi.org/id/5126B5BADC2E8B7EA71E75AB48C43E1E" lastPageId="21" lastPageNumber="78" pageId="20" pageNumber="77">
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<taxonomicName LSID="5126B5BADC2E8B7EA71E75AB48C43E1E" authority="Zhang & Li, 1981" authorityName="Zhang & Li" authorityYear="1981" class="Diplopoda" family="Glomeridae" genus="Bilingulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bilingulus sinicus" order="Glomerida" pageId="20" pageNumber="77" phylum="Arthropoda" rank="species" species="sinicus">Bilingulus sinicus Zhang & Li, 1981</taxonomicName>
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<treatment id="5126B5BADC2E8B7EA71E75AB48C43E1E" ID-GBIF-Taxon="158498357" LSID="urn:lsid:plazi:treatment:5126B5BADC2E8B7EA71E75AB48C43E1E" httpUri="http://treatment.plazi.org/id/5126B5BADC2E8B7EA71E75AB48C43E1E" lastPageId="21" lastPageNumber="78" pageId="20" pageNumber="77" scope_class="Diplopoda" scope_family="Pericambalidae" scope_order="Spirostreptida">
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<subSubSection id="71FB40B12BBCB66ED47DA7769BDF5E9E" pageId="20" pageNumber="77" type="nomenclature">
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<paragraph id="1A7680CF3B98566E85453A8418C256E5" pageId="20" pageNumber="77">
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<taxonomicName id="816AADB489E2546EC1D724A80D8C34F9" LSID="5126B5BADC2E8B7EA71E75AB48C43E1E" authority="Zhang & Li, 1981" authorityName="Zhang & Li" authorityYear="1981" class="Diplopoda" family="Glomeridae" genus="Bilingulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bilingulus sinicus" order="Glomerida" pageId="20" pageNumber="77" phylum="Arthropoda" rank="species" species="sinicus">Bilingulus sinicus Zhang & Li, 1981</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="21" lastPageNumber="78" pageId="20" pageNumber="77" type="materials_examined">
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<paragraph pageId="20" pageNumber="77">Material examined.</paragraph>
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<paragraph lastPageId="21" lastPageNumber="78" pageId="20" pageNumber="77">
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<subSubSection id="ABB7D33A8C8B3722518C470FBCEEBBE8" lastPageId="21" lastPageNumber="78" pageId="20" pageNumber="77" type="materials_examined">
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<paragraph id="21D909DE282D3DFF4EA7AC57F50AA887" pageId="20" pageNumber="77">Material examined.</paragraph>
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<paragraph id="FD96989A7D3F64D5832DA80FE444E661" lastPageId="21" lastPageNumber="78" pageId="20" pageNumber="77">
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3 males, 3 females, 5 juv. (SCAU), China, Guangxi Zhuang Autonomous Region, Yangshou County, Shangshuiyan Cave [
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<geoCoordinate degrees="24" direction="north" minutes="57" orientation="latitude" precision="1" seconds="43.6" value="24.962112">24°57'43.6"N</geoCoordinate>
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<geoCoordinate id="97926EBDF7C114926F6B13DB20A54D20" degrees="24" direction="north" minutes="57" orientation="latitude" precision="1" seconds="43.6" value="24.962112">24°57'43.6"N</geoCoordinate>
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,
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<geoCoordinate degrees="110" direction="east" minutes="20" orientation="longitude" precision="1" seconds="37.21" value="110.34367">110°20'37.21"E</geoCoordinate>
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<geoCoordinate id="C1AC16A7DB49B6CB8C0BB33D1D2FFD0B" degrees="110" direction="east" minutes="20" orientation="longitude" precision="1" seconds="37.21" value="110.34367">110°20'37.21"E</geoCoordinate>
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], 191 m el., deep zone, cartography station #19, direct intuitive
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<pageBreakToken pageId="21" pageNumber="78" start="start">search</pageBreakToken>
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<pageBreakToken id="B029084E25C7103B510321A4A714437A" pageId="21" pageNumber="78" start="start">search</pageBreakToken>
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, 17 November 2016, J.J. Wynne leg. 4 females (SCAU), China, Guangxi Zhuang Autonomous Region, Xiufeng District, Maomaotou Cave [
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<geoCoordinate degrees="25" direction="north" minutes="18" orientation="latitude" precision="1" seconds="46.12" value="25.31281">25°18'46.12"N</geoCoordinate>
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<geoCoordinate id="1BE02AAD04D54352179FD8CAF456D49D" degrees="25" direction="north" minutes="18" orientation="latitude" precision="1" seconds="46.12" value="25.31281">25°18'46.12"N</geoCoordinate>
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,
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<geoCoordinate degrees="110" direction="east" minutes="16" orientation="longitude" precision="1" seconds="12.64" value="110.27018">110°16'12.64"E</geoCoordinate>
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<geoCoordinate id="F82BF8EA45DD730D4C5F4CD9DA6A1DF8" degrees="110" direction="east" minutes="16" orientation="longitude" precision="1" seconds="12.64" value="110.27018">110°16'12.64"E</geoCoordinate>
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], 225 m el., deep zone, direct intuitive search, 15 November 2016, J.J. Wynne leg. 1 juv. (SCAU), China, Guangxi Zhuang Autonomous Region, Lingchuan County, Liangfeng Cave [
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<geoCoordinate degrees="25" direction="north" minutes="12" orientation="latitude" precision="1" seconds="34.86" value="25.209684">25°12'34.86"N</geoCoordinate>
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<geoCoordinate id="2FB4318B75E7C11CF2EDCD7095A723EC" degrees="25" direction="north" minutes="12" orientation="latitude" precision="1" seconds="34.86" value="25.209684">25°12'34.86"N</geoCoordinate>
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,
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<geoCoordinate degrees="110" direction="east" minutes="31" orientation="longitude" precision="1" seconds="56.8" value="110.53245">110°31'56.8"E</geoCoordinate>
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<geoCoordinate id="1418EC7230A19C35495ACDE79B2A7B6E" degrees="110" direction="east" minutes="31" orientation="longitude" precision="1" seconds="56.8" value="110.53245">110°31'56.8"E</geoCoordinate>
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], 184 m el., deep zone, direct intuitive search, 18 November 16, J.J. Wynne leg.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="78" type="notes">
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<paragraph pageId="21" pageNumber="78">Notes.</paragraph>
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<paragraph pageId="21" pageNumber="78">
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<subSubSection id="CF4E42767DE2BCFD200753641D368CD0" pageId="21" pageNumber="78" type="notes">
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<paragraph id="7ECD0BF17D0977665969999D58E8AB36" pageId="21" pageNumber="78">Notes.</paragraph>
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<paragraph id="6A6DF28FB986E74C90FFD91E1E817A90" pageId="21" pageNumber="78">
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This species lacked characters suggestive of troglomorphy. It had a pigmented cuticle and ommatidia were present. Thus,
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<taxonomicName genus="B." lsidName="B. sinicus" pageId="21" pageNumber="78" rank="species" species="sinicus">
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<emphasis italics="true" pageId="21" pageNumber="78">B. sinicus</emphasis>
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<taxonomicName id="AD293950A6DD2F0F5F473BD56B376687" lsidName="B. sinicus" pageId="21" pageNumber="78" rank="species" species="sinicus">
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<emphasis id="DE936ABD58F6F8A1E6E56FBC298E9CAD" italics="true" pageId="21" pageNumber="78">B. sinicus</emphasis>
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</taxonomicName>
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is considered a troglophile. It was first described by
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<bibRefCitation author="Zhang, CZ" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="32" pageNumber="89" pagination="373 - 377" refId="B83" refString="Zhang, CZ, Li, ZY, 1981. Bilinguidae family nov. of Diplopoda in karst landform of south China. . Acta Zootaxonomica Sinica 6: 373 - 377" title="Bilinguidae family nov. of Diplopoda in karst landform of south China." volume="6" year="1981">Zhang and Li (1981)</bibRefCitation>
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<bibRefCitation id="2A3D2889D60CBCB48FC7CB2F866BF3B0" author="Zhang, CZ" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="32" pageNumber="89" pagination="373 - 377" refId="B83" refString="Zhang, CZ, Li, ZY, 1981. Bilinguidae family nov. of Diplopoda in karst landform of south China. . Acta Zootaxonomica Sinica 6: 373 - 377" title="Bilinguidae family nov. of Diplopoda in karst landform of south China." volume="6" year="1981">Zhang and Li (1981)</bibRefCitation>
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from a cave in Guilin; no cave name or coordinate data were provided. While we detected this species in three different Guilin caves,
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<normalizedToken originalValue="we’re">we're</normalizedToken>
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<normalizedToken id="261E1721FC3ABFB10B90879727EE041E" originalValue="we’re">we're</normalizedToken>
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unable to determine a distance from the type locality. The caves where we detected
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<taxonomicName genus="P." lsidName="P. trifidus" pageId="21" pageNumber="78" rank="species" species="trifidus">
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<emphasis italics="true" pageId="21" pageNumber="78">P. trifidus</emphasis>
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<taxonomicName id="3B529504242FFC6AC3866787C42FB3D2" lsidName="P. trifidus" pageId="21" pageNumber="78" rank="species" species="trifidus">
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<emphasis id="B5F95972181EF90CB6B5BAC3ED914996" italics="true" pageId="21" pageNumber="78">P. trifidus</emphasis>
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</taxonomicName>
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is separated by an average distance of 34.1 km. Maomaotou Cave occurs to the north-northwest and is 45 km from Liangfeng Cave and 40.4 km to the north of Shangshuiyan Cave. Based upon its epigean characteristics and that this species is regionally distributed on the landscape, we consider it a troglophile.
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</paragraph>
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